A Stereo -Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter,
and J. E. Whittaker
Volume 17, Part 1; 31st July, 1990
Published by the British Micropalaeontological Society, London
ISSN 0952-7451
Editors
Dr J. Athersuch, Exploration Technology Branch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Dr D.J. Horne, School of Earth Sciences, Thames Polytechnic, Walburgh House, Bigland Street,
London El 2NG.
Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.
Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.
Editorial Board
Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.
Dr P. Deckker, Department of Geology, Australian National University, G.P.O. Box 4, Canberra, ACT 2601,
Australia.
Dr W. Hansch, Ernst-Moritz-Arndt Universitat, Sektion Geologische Wissenschaften, F.L.-Jahnstr. 17a,
2200 Greifswald, German Democratic Republic.
Prof. R. Lundin, Department of Geology, Arizona State University, Tempe, Arizona 85287-1404, U.SA.
Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55,
D 2000 Hamburg 13, German Federal Republic.
Dr Zhao Yuhong, Nanjing Institute of Geology and Palaeontology, Academia Sinica, Chi-Ming-Ssu, Nanjing,
People’s Republic of China.
Officers of the British Micropalaeontological Society
Chairman Professor M.B. Hart, Department of Geological Sciences, Polytechnic South West, Drake Circus,
Plymouth, Devon PL4 8AA.
Secretary Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.
Journal Editor Dr M.C. Keen, Department of Geology, The University, Glasgow G12 8QQ.
Newsletter Editor Dr D.J. Shipp, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Conodont Group Chairman Dr H.A. Armstrong, Department of Geology, The University,
Newcastle upon Tyne, NE1 7RU.
Conodont Group Secretary Mr M.T. Dean, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Foraminifera Group Chairman Dr H.W. Bailey, Paleo Services Ltd., Unit 15, Paramount Industrial Estate,
Sandown Road, Watford WD2 4XA.
Foraminifera Group Secretary Dr F.M.D. Lowry, School of Earth Sciences, Thames Polytechnic,
Walburgh House, Bigland Street, London El 2NG.
Ostracod Group Chairman Dr J. Athersuch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Ostracod Group Secretary Dr I.D. Boomer, School of Environmental Sciences, University of East Anglia,
Norwich NR4 7TJ.
Palynology Group Chair Dr R.J. Davey, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Palynology Group Secretary Dr A. McNestry, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Calcareous Nannofossil Group Chairman Dr M.K.E. Cooper, ECL Stratigraphic Services Ltd., Chancellor
Court, Surrey Research Park, Guildford GU2 5YL.
Calcareous Nannofossil Group Secretary Dr P.R. Bown, Department of Geology (Micropalaeontology),
University College, Gower Street, London WC1E 6BT.
Instructions to Authors
Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. Format
should follow the style set by the papers in this issue. Descriptive matter apart from illustrations should be
cut to a minimum; preferably each plate should be accompanied by only one page of text. Blanks to aid in
mounting figures for plates may be obtained from any one of the Editors or Editorial Board. Completed
papers should be sent to Dr David J. Siveter.
The front cover shows a male carapace (left side) of Callistocythere murrayi Whittaker from Mother Siller’s
Channel, Christchurch Harbour, Southern England; in brackish water. Photographed by J.E. Whittaker,
British Museum (Natural History).
Printed by BPCC Blackpool Ltd . Stanley Road. Blackpool FY1 4QN
Member of BPCC Ltd
Stereo-Atlas of Ostracod Shells 17 (1) 1-4 (1990) Welchella foveata (1 of 4)
595.336.21 (113.51) (761 : 162.087.33) : 551.351 + 552.52
ON WELCHELLA LOVEATA DEWEY & PUCKETT gen. et sp. nov.
by Christopher P. Dewey & T. Mark Puckett
(Mississippi State University & Alabama Geological Survey, U.S.A.)
Genus WELCHELLA gen. nov.
Type-species: Welchella foveata sp. nov.
After Stewart W. Welch, who named the Pride Mountain Formation. Gender (diminutive -ella), feminine.
Medium sized geisinid, semicircular, equivalve carapace with straight dorsum and very subdued bilobation.
Wide inner lamella but lacking vestibulae. Dimorphic, females postplete, males amplete.
Welchella is a geisinid (Superfamily Kloedenellacea) by virtue of the bilobation, the straight dorsum with
tongue and groove hinge and the inner lamella. Welchella differs from Knoxiella Egorov, 1950 ( Trudy vses.
neft. nauchno-issled. geol. -razv. Inst.) by the postplete nature of the females and the very wide inner lamella.
In Knoxiella, dimorphism is expressed in a posterior swelling of the female and the inner lamella is narrow.
Welchella foveata sp. nov.
Dunn-Seiler Museum of Geology, Mississippi State University, no. 3341-la; cf carapace.
[Paratypes nos. 3341-lb to le; 1 $ carapace and 3 $ valves.]
Section in Dry Creek Quarry, N of Trussville, Alabama, U.S.A. ; Sec. 14 T16S R1W; lat. 33°37' 30"N, long.
86° 37' 30" W. Basal Pride Mountain Formation, Chesterian, Mississippian, Carboniferous; 2.2m above the
base of the formation in black fossiliferous shale, marine.
Latin foveatus -a, -um, pitted; referring to the surface ornament.
Dunn-Seiler Museum of Geology, Mississippi State University, nos. 3341-la (holotype, 9 car.: PI. 17, 2,
figs. 1-3), 3341-lb (paratype, $ RV: PI. 17, 4, fig. 4), 3341-lc (paratype, $ car.: PI. 17, 2, figs. 4-6),
3341-ld (paratype, $ LV: PI. 17, 4, fig. 3), 3341-le (paratype, 9 LV: PI. 17, 4, fig. 1), 3341-lf (broken
valve material: PI. 17, 4, figs. 2, 5, 6). All from the type locality; black shale with abundant goniatites,
bryozoans, brachiopods, bivalves and gastropods. From 2.2 m (nos. 3341-la, 3341-le) and 2.9 m (nos.
3341-lb, 3341-lc, 3341-ld and 3341-lf) above the base of the formation.
Explanation of Plate 17, 2
Figs. 1-3, cf car. (holotype, 3341-la, 775 pm long): fig. 1, LV ext. lat.; fig. 2, dors.; fig. 3, RV ext. lat. Figs. 4-6, 9 car. (paratype,
3341-lc, 115pm long): fig. 4, LV ext. lat.; fig. 5, RV ext. lat.; fig. 6, dors. Scale A (100/u.m; x72), figs. 1-6.
Derivation of name:
Diagnosis:
Remarks:
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Stereo-Atlas of Ostracod Shells 17, 3 Welchella foveata (3 of 4)
Diagnosis: Medium sized (Text-fig. 1), equivalved carapace with straight dorsum. Semicircular in lateral outline,
fusiform in dorsal view. Cardinal angles obtuse, distinct. Bilobation subdued, marked only by shallow,
comma-shaped S2 anterior of mid-length and above mid-height. Ornamentation consists of pits over lateral
surface; intensity of pitting fades to valve margins. Hinge simple, tongue and groove, adont. Wide inner
lamella, narrows to cardinal angles, widest anteriorly. Vestibulae absent, inner and outer lamella fused.
Dimorphic, males amplete, females postplete and slightly broader in posterior.
Welchella is only known from a single species.
Pride Moutain Formation, Chesterian, Mississippian of the Black Warrior Basin, Alabama, U.S.A. Samples
collected 2.2 m and 2.9 m above the top of the Tuscumbia Limestone.
We acknowledge the financial support given by the Donors of the Petroleum Research Fund administered by
the American Chemical Society; the Mississippi Mineral Resources Institute and Mississippi State University.
Remarks:
Distribution :
Acknowledgement :
700
Text-fig. 1. Ontogeny of W. foveata from Dry Creek -
Quarry at 2.2 m and 2.9 m above the base of the 2
Pride Mountain Formation.
100
DC Q 2 2
■ • ••
- •••
••
- • ••
••••
• •••
rr prrrp n r j i i i | i i i p it
Length (tim)
700 ■
• ■ •
. •••
••••
• ■ •
• ■
••
200
M I | I II | I I I | I I I | I '
Length (ttm)
900
Explanation of Plate 17, 4
Fig. 1, 9 LV, ext. lat. (paratype, 3341-le, 725 /i,m long); fig. 2, ant. valve wall and inner lamella (27 p m thick, broken valve material,
3341-lf); fig. 3, 9 LV, int. lat. (paratype 3341-ld, 725 pm long); fig. 4, 9 RV, int. lat. (paratype 3341-lb, 725 pm long); figs. 5,
6, post, valve wall (25pm thick, broken valve material, 3341-lf).
Scale A (100 pm\ x72), figs. 1, 3, 4; scale B (100 /urn; xl80), figs. 2, 5; scale C (10 /urn; x2000), fig. 6.
Welchella foveata (2 of 4)
Stereo- Atlas of Ostracod Shells 17, 2
-Stereo-Atlas of Ostracod Shells 17, 4
Welchella foveata (4 of 4)
Stereo-Atlas of Ostracod Shells 17 (2) 5-8 (1990)
595.336.21 (113.51) (761: 162.088.34 + 773 : 162.089.37) : 551 .351 + 552.52
Glyptopleura henbesti (1 of 4)
ON GLYPTOPLEURA HENBESTI CRONEIS & GUTKE
by Christopher P. Dewey
(Mississippi State University, Mississippi, U.S.A.)
Glyptopleura henbesti Croneis & Gutke, 1939
1939 Glyptopleura henbesti n. sp. C. Croneis & R. L. Gutke, J. scient. Labs Denison Univ., 34, 51, 52, pi. 2, figs. 7, 8.
1939 Glyptopleura hendricksi n. sp. C. Croneis & R. L. Gutke, Ibid. , 34, 52, 3, pi. 2, figs. 5, 6.
1941 Glyptopleura henbesti Croneis & Gutke; C. L. Cooper, Rep. Invest. III. St. geol. Surv., no. 77, 40, 41, pi. 7, figs. 9-11.
Holotype: Field Museum of Natural History, Chicago, U.S.A., no. UC 45169; female carapace.
Type locality : Locality no. .0526.42; greenish-grey non-laminated, fossiliferous clay-shale; W side of road, at base of
Melcher Hill, Sec. 26, T12S R7E, N of Shetlerville, Hardin County, Illinois, U.S.A. ; lat. 37° 26' 29" N, long.
88°24'21"W. Renault Formation, Chesterian, Mississippian, Carboniferous.
Figured specimens: Field Museum of Natural History, U.S.A., nos. UC 45169 (holotype, $ car.: PI. 17, 8, figs. 2, 3), UC 51725
(paratype, 9 car.: PI. 17, 6, figs. 1, 2), UC 45170 (cf car. [holotype of G. hendricksi ]: PI. 17, 8, figs. 1, 2).
Illinois State Geological Survey, U.S.A., no. ISGS 45P131 ($ car.: PI. 17, 8, fig. 5). Dunn-Seiler Museum of
Geology, Mississippi State University, U.S.A., no. 3341-2 (9 LV: PI. 17, 6, figs. 3, 4).
Nos. UC 45169 and 51725 are from the type locality. UC 45170 is from locality no. .0526.41, N of
Shetlerville, Illinois; lat. 37° 26' 23" N, long. 88° 24' 21" W. ISGS 45P131 is from Sec. 11, T12S, R7E, S of
Eichorn, Hardin County, Illinois; lat. 37°29'20"N, long. 88°24'20"W. All from Renault Formation,
Chesterian, Mississippian, Carboniferous. 3341-2 is from light brown fossiliferous mudstone, county highway
37, Sec. 31, T5S R10W, Colbert County, Alabama; lat. 34° 34' 12"N, long. 87° 37' 28" W; Bangor Limestone
Formation, Chesterian. Mississippian, Carboniferous.
Diagnosis: Medium-sized, subquadrate, bilobate distinct S2 at mid-length, ends evenly rounded, left valve larger than
right. Dorsum straight, cardinal angles obtuse, distinct. Dorsal aspect cuneate, posterior acuminate, anterior
blunt, maximum width in posterior. Lateral surface has six, major, striate costae (Text-fig. 1) subparallel to
Explanation of Plate 17, 6
Figs. 1, 2, 9 car. (paratype, UC 51725, 840 /am long): fig. 1, ext. vent.; fig. 2, RV, ext. lat. Figs. 3, 4, 9 LV (3341-2, 900 /um long):
fig. 3, lat. ext. anterodors. ; fig. 4, ext. lat. Scale A (100 ^im; x65), figs. 1, 2, 4; scale B (50 /um; x295), fig. 3.
Stereo-Atlas of Ostracod Shells 17, 7 Gyptopleura henbesti (3 of 4)
long axis. Costa 1 is marginal and fades posteriorly. Costae 2 and 6 form “U”, closed anteriorly but with short
anterodorsal extension “a”; costa 3 not connected to “U”; costae 4 and 5 extend posteriorly from loop of
“U”. Two minor costae (a and b) at 90° in anterodorsal region; “b” posterior to “a”, subparallel to costa 2.
Third minor costa (“c”) ventral to costa 6. Costae do not extend beyond the posterior lobe; costae 2 and 4 end
as short spines. Surface also reticulate and papillate. Posterior free margin denticulate. Dimorphic, females
wider posteriorly.
Remarks: Intraspecific variation of the multicostate pattern in the genus Glyptopleura Girty, 1910, is common.
Variations in the costae can occur on either valve of a single carapace (I. G. Sohn, Prof. Pap. U.S. geol.
Surv., 606F, 47, 1969), or between the sexes, adults of the same sex or instars and adults. Normally the
number and general position of the costae are constant within a species; however the length, exact position
and anastomoses of costae may be variable. G. hendricksi is placed in synonymy with G. henbesti because the
variation in the posterior portion of the carapace (PI. 17, 8, fig.
2) is a dimorphic character and the only costa in G. hendricksi
to show any significant difference is the marginal costa, which
is formed from separate dorsal and ventral costae that fade as
they pass each other and wrap around the anterior margin.
Distribution: Illinois Basin, Illinois and Black Warrior Basin, Alabama;
Chesterian, Mississippian, Carboniferous.
Acknowledgement : I acknowledge the financial support given by the Donors of the
Petroleum Research Fund administered by the American
Chemical Society; the Mississippi Mineral Resources Institute
and Mississippi State University.
6
Text-fig. 1. Costae, left valve of G. henbesti.
Explanation of Plate 17, 8
Figs. 1, 2, 9 car- (UC 45170, 840 pm long): fig. 1, LV, lat. ext. ; fig. 2, ext. dors. Figs. 3, 4, 9 car. (holotype, 45169, 840 yum long): fig.
3, LV, ext. lat.; fig. 4, ext. dors. Fig. 5, 9 car. (ISGS 45P131, 840 ^im long), LV, ext. lat. Scale A (100 /xm; x65), figs. 1-5.
b S 2
Stereo-Atlas of Ostracod Shells 17, 6
Glyptopleura henbesti (2 of 4)
Stereo-Atlas of Ostracod Shells 17, 8
Glyptopleura henbesti (4 of 4)
Stereo-Atlas of Ostracod Shells 17 (3) 9-12 (1990)
595.336.13 (113.312) (766: 162.097.34) 551.351 + 552.54
Winchellatia longispina (1 of 4)
ON WINCHELLATIA LONGISPINA KAY
by Mark Williams
( University of Leicester, England)
Genus WINCHELLATIA Kay, 1940
Type-species (by original designation): Winchellatia longispina Kay, 1940
Diagnosis: Unisulcate Glossomorphitinid ; S2 sigmoidal, beginning slightly dorsal of dimunitive preadductorial node,
continued ventrally and defining anterior and ventral termination of posteroventral lobe and confluent
ventrally with distinct laterovelar furrow. Dimorphic in some species. Tecnomorphic velum a narrow flange
beginning posteroventrally, continues to near anterocardinal corner, much wider ventrally in heteromorph.
Posteroventral lobe inflated with a distinct spine.
Remarks: Winchellatia is most similar to Collibolbina Schallreuter (1967, Neues Jb. Geol. Palaont. Mh. 7, 431-446)
differing only in having a row of spines admarginally and by having S2 continuous ventrally with the
laterovelar furrow. Schallreuter (pers. comm.) considers that Collibolbina may be a subgenus of Winchellatia.
The type-species of Acronotella, A. schilderi Ulrich & Bassler ( Maryland Geol. Surv., Silurian Volume,
298, text-figs. 25-27, 1923), is very similar to W. longispina (see also Treatise of Invertebrate Paleontology
part Q, fig. 186, la, b, 1961) in having a strong laterally projecting spine on the posteroventral lobe, a strong
S2, and a similar outline. Acronotella appears to differ from Winchellatia by the lack of the confluence of S2
with the strong laterovelar furrow and by having strong anterocardinal spines.
At present Winchellatia includes a large number of species showing a wide range of morphology. W.
minnesotensis Kay (1940, op. cit. 225, pi. 32, figs. 13-19), W. lansigensis Kay (1940, op. cit., 254, pi. 32, figs.
6-8) and W. nahanniensis Copeland (1982, Bull. geol. Surv. Can., 347, 16 pi. 3, figs. 16, 25-30) are not
known to de dimorphic and are here questionably assigned to Winchellatia. W. ? bullata Pribyl (Sb. nar. mus.
Praze, 33,64, text-fig. 2, fig. 9, pi. 12, fig. 4, 1977) is non-dimorphic, very large, has no spines or marginal
tubercles and almost certainly belongs in a separate genus. Several Devonian species have also been assigned
to Winchellatia by Kesling & Tabor (/. Paleont., 26, 761-763, 1952; Contr. Mus. Paleont. Univ. Mich., 10,
Explanation of Plate 17, 10
Fig. 1-3, $ LV. (OS 13471, 1 mm long): fig. 1 , ext. lat. ; fig. 2, ext. lat. obi.; fig. 3, ext. vent. Fig. 4, 9 RV ext. lat. (OS 13472, 1.02mm
long).
Stereo-Atlas of Ostracod Shells 17, 11 Winchellatia longispina (3 of 4)
83-100, 1953); the lobation and sulcation of these species is quite unlike that of Winchellatia. Other features
which characterise species of Winchellatia include the bending of the velum towards the margin posteriorly
and the presence of an admarginal ridge surmounted by tubercles on both valves. This ridge can be seen to
migrate towards the velum anteroventrally, a feature also described in Collibolbina (Schallreuter, 1967).
1940
1957
1957
1965
Winchellatia longispina Kay, 1940
Winchellatia longispina n. sp. G. M. Kay, J. Paleont., 10, 235, pi. 32, figs. 1-5.
Winchellatia longispina Kay; R. W. Harris, Bull. Okla geol. Surv., 75, 220, pi. 9, figs. 9a-d, 10, lla-d.
Winchellatia cornuta n. sp. R. W. Harris, Bull. Okla geol. Surv., 75, 220, pi. 9, figs. 4a, b.
Winchellatia longispina Kay; M. J. Copeland, Bull. geol. Surv. Can., 127, 20, pi. 10, fig. 6.
Holotype: University of Columbia, U.S.A., Department of Geology, no. 275861; § left valve.
Gutenburg Member, Decorah Formation, middle Ordovician, Church, Iowa, U.S.A. (see Kay, 1940).
British Museum (Nat. Hist.) nos. OS 13471 (9 LV: PI. 17, 10, figs. 1-3; PI. 17, 12, fig. 2),
OS 13472 (9 RV: PI. 17, 10, fig. 4; PI. 17, 12, fig. 1), OS 13479 (c? RV: PI. 17, 12, fig. 3), OS 13473 (c? LV:
PI. 17, 12, fig. 4), OS 13474 (9 RV: PI. 17, 12, fig. 5). All figured specimens from the Pooleville Member,
Bromide Formation, Rock Crossing, in the Criner Hills (see Harris, 1957), Oklahoma, U.S.A. ;
approximately latitude 34°08'N, longitude 97°10'W.
Winchellatia species with an elongate, posteroventrally directed spine on the posteroventral lobe. Subvelar
field with marginal finely tuberculate ridge present on both valves.
The holotype of Winchellatia cornuta Harris, 1957 (Museum of Comparative Zoology, Harvard University,
U.S.A., no. 4615) is a badly abraded specimen of W. longispina. Juveniles have the strong marginal ridge of
adults, but the velum is weaker, being developed only as a right angled bend. The preadductorial node and S2
are also weaker in juveniles. The subelar field is smooth in all of the heteromorphic (9) specimens examined,
but in tecnomorphs it is distinctly reticulate.
Middle Ordovician of the U.S.A.: Gutenburg Member, Decorah Formation, Iowa and the Bromide
Formation, Oklahoma.
Drs. R. E. L. Schallreuter (Hamburg) and D. J. Siveter (Leicester), for discussion.
Type locality :
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Ackno wledgemen t :
Explanation of Plate 17, 12
Fig. 1, 9 RV, ext. ant. (OS 13472, 1.02mm long); fig. 2, 9 LV, ext. ant. (OS 13471, 1mm long); fig. 3, <d RV, ext. lat. (OS 13479,
1mm long); fig. 4, <d LV, int. lat. (OS 13473, 1mm long); fig. 5, 9 RV, close up of ant. vent. (OS 13474, fragment).
Scale A (200/xm; x66), figs. 1-4; scale B (100 gm; x75), fig. 5.
Stereo-Atlas of Ostracod Shells 17, 10
Winchellatia longispina (2 of 4)
Winchellatia longispina (4 of 4)
Stereo-Atlas of Ostracod Shells 17, 12
Stereo-Atlas of Ostracod Shells 17 (4) 13-18(1990) Eridoconcha simpsoni (1 of6)
595.336 (113.312) (766 : 162.097.34) 551.351 + 552.52
ON ERIDOCONCHA SIMPSONI HARRIS
by Mark Williams & Peter J. Jones
( University of Leicester, England & Bureau of Mineral Resources, Canberra, Australia)
Eridoconcha simpsoni Harris, 1931
1931 Eridoconcha simpsoni n. sp. R. W. Harris, Bull. Okla geol. Surv., 55, 90, pi. 14, figs, la, b, pi. 11. figs. la-d.
1934 Eridoconcha simpsoni Harris; R. S, Bassler & B. Kellett, Spec. Pap. geol. Soc. Am.. 1, 310.
1951 Eridoconcha simpsoni Harris; J. E. Keenan, J. Paleont., 25, 565.
1951 Cryptophyllus simpsoni (Harris); S. A. Levinson, J. Paleont., 25, 558.
1957 Cryptophyllus simpsoni (Harris); R. W. Harris, Bull. Okla geol. Surv., 75, 183, pi. 5, figs. 12a, 13a, b, 14a, b, 15a, b.
1961 Aberroconcha? simpsoni (Harris); F. J. Adamczak, Acta palaeont. pi, 6, 73.
1962 Cryptophyllus simpsoni (Harris); P. J. Jones, Bull. Bur. Miner. Resour. Geol. Geophys. Aust., 62-3, 19.
1968 Eridoconcha? simpsoni Harris; R. E. L. Schallreuter, Palaeont. Z. , 42, 109.
Holotype:
Type locality:
Figured specimens:
Museum of Comparative Zoology, Harvard University, U.S.A., no. 7447; carapace (ninth
lamellae damaged).
From C. E. Decker's ‘Zone 16’ (see Harris, 1957), Bromide Formation, Simpson Group, middle
Ordovician; about 400m W of U.S. Highway 77 (sec. 25 T. 2s, R.l E), Arbuckle Mountains,
Oklahoma, U.S. A.; approximately lat. 34°25'N., long. 97°08'W.
Museum of Comparative Zoology, Harvard University, U.S. A., no.7447 (car.; PI. 17, 16, fig. 1).
Commonwealth Palaeontological Collections, Canberra, Australia nos. CPC 28741 (LV; PI. 17,
16, fig. 4) and CPC 28472 (RV: PI. 17, 16, fig. 5). British Museum (Nat. Hist.) nos. OS 13477 (LV:
PI. 17, 14, figs. 1-5; PI. 17, 16, fig. 1), OS 13476 (RV: PI. 17, 16, fig. 3), OS 13475 (LV: PI. 17, 16,
fig. 6).
Explanation of Plate 17, 14
Figs. 1-5, LV (OS 13477, 0.77mm long): fig. 1, ext. lat.; fig. 2, ext. lat obi.; fig. 3, dors obi.; fig. 4, vent.; fig. 5, ant.
Scale A (200 /nmm; x76), figs. 1-5.
Stereo-Atlas of Ostracod Shells 17, 15 Eridoconcha simpsoni (3 of 6)
All of the figured specimens come from the Bromide Formation. MCZ 7447 is from the type
horizon and locality. CPC 28741 and 28742 are from Decker’s ‘Zone 35’, Mountain Lake Member,
Highway 99 road section (see Harris, 1957), collected by V. Jaanusson (1959). OS 13475 to 13577
are from the top bed of the Mountain Lake Member, Bromide Formation, North Interstate 35
locality (see Fay & Grafham, Univ. Kans. paleont. Contr., Monograph, 1, 14, 1982).
Diagnosis: Species of Eridoconcha with naupliconch having an elongated posteriorly directed spine.
Maximum of nine lamellae, each delimited by wide “U”-shaped grooves. Greatest valve width
ventral of umbo at second to third lamellae. Final lamella preplete in lateral outline. Internal
adductorial sulcament strongly developed.
Remarks: Levinson (1951) assigned E. simpsoni to his new genus Cryptophyllus. However, the adventral
structures and deep “U”-shaped grooves (Text-fig. 1) between the lamellae in E. simpsoni clearly
distinguishes it from the type-species of Cryptophyllus , C. oboloides (Ulrich & Bassler, 1923).
E. rugosa, the type-species of Eridoconcha Ulrich & Bassler, 1923, has recently been
examined by Warshauer & Berdan {Prof. Pap. U.S. geol. Surv. , 1066-H, 1982), and by P. J. Jones.
It possesses up to four lamellae, each with a distinct adventral structure (a swollen rib) situated
some distance from its free margin, a character diagnostic of the genus as a whole, and recognised
in E. simpsoni (Text-fig. 1). E. simpsoni is readily distinguished from E. rugosa by its consistently
greater number of lamellae.
From his figures it is clear that Harris (1957, pi. 5, figs. 14a, b, 15a, b, 12) considered E.
simpsoni to be dimorphic, with males more elongate than females. No such dimorphism has been
recognised by us. Based on specimens recovered from several samples, E. simpsoni shows quite
wide adult size variation (Text-figs. 2, 3). Wide intraspecific variation of the carapace outline is
also a noted feature (see above) of the species.
Explanation of Plate 17, 16
Fig. 1, LV, post. (OS 13477, 0.77mm); fig. 2, car., ext. Lt. lat. (MCZ 7447, 0.6mm long); fig. 3, RV, int. lat. (OS 13476, 0.73mm
long); fig. 4, LV, detail int. vent, showing contact groove (CPC 28741, 0.63mm); fig. 5, RV, detail int. vent. (CPC 28742,
0.55 mm); fig. 6, int. lat. (OS 13475, 0.75 mm).
Scale A (200 /xm; x 76), fig. 1; scale B (100 /i,m; x 95), fig. 2; scale C (150 /u.m; x 72), figs. 3, 6; scale D (100 /am; x 81), fig. 4; scale E
(100Mm; X87), fig. 5.
Stereo-Atlas of Ostracod Shells 17, 14
Eridoconcha simpsoni (2 of 6)
Eridoconcha simpsoni (4 of 6)
Stereo-Atlas of Ostracod Shells 17, 16
Stereo-Atlas of Ostracod Shells 17, 17 Eridoconcha simpsoni (5 of 6)
A marked contact groove exists in the last lamella of the left valve, which receives a
rudimentary contact list of the right valve (PI. 17, 16, figs. 4,5). Carapaces display left over right
valve overlap. The right valve hinge appears to have a distinct groove; there is a ridge in the
opposing position in the left valve.
E. simpsoni occurs in the Bromide Formation together with other eridostracans (C. gibbosum
Harris, 1957 and C. nuculopsis Harris, 1957), where they characterise the deeper water platform
biofacies, as they do in other N. American sequences (see M. J. Copeland, Bull. geol. Surv. Can. ,
127, 1982). It is readily distinguished from these species on the basis of the surface relief of the
lamellae.
Distribution: E. simpsoni characterises marine offshore shelf facies in the middle Ordovician Bromide
Formation, Oklahoma, U.S.A.
Text-fig. 1. Schematic cross-section of the adventral ridges of the 7th, 8th and 9th lamellae of E. simpsoni.
Stereo-Atlas of Ostracod Shells 17, 18 Eridoconcha simpsoni (6 of 6)
Text-fig. 2. Size dispersion of 17 adult left valves of
E. simpsoni from three samples in the Mountain Lake
Member, Bromide Formation, Oklahoma, U.S.A.
Text-fig. 3. Size dispersion of 21 adult right valves of
E. simpsoni from three samples in the Mountain Lake
Member, Bromide Formation, Oklahoma, U.S.A.
Stereo-Atlas of Ostracod Shells 17 (5) 19-22 (1990) Cypridea unicostata chinensis (1 of 4)
595.337.12 (116.312) (510 : 161.107.40) : 551.312
ON CYPRIDEA UNICOSTATA GALEEVA CHINENSIS NEALE & SU subsp. nov.
by John W. Neale & Su Deying
(University of Hull, England & Institute of Geology, Chinese Academy of Geological Sciences, Beijing, China)
Cypridea unicostata Galeeva chinensis subsp. nov.
1980 Cypridea unicostata Galeeva; Su Deying, Li Yougui, Pang Qiqing & Chen Sue, The Mesozoic strata and paleontology in
Shanganning Basin, part 2. Fossil Ostracoda, Geological Publishing House, Beijing, 77, pi. 116, figs. 2a-f.
Holotype: Institute of Geology, Chinese Academy of Geological Sciences, Beijing, China, coll. no. 10.2;
carapace.
Type locality: Jingchuan Formation, Zhidan Group, Wujiamiao (=Wujia or Wuyia Temple), Shanganning
Basin, lat. 40°23'N, long. 107° 49' E. Hauterivian-Barremian, Early Cretaceous.
Derivation of name: From its wide occurrence in China.
Fig. 1, car., ext. rt. lat. (10.1, 985 gm long); fig. 2, car., ext. dors. (10.4, 1010 /urn long); fig. 3, car., ext. It. lat. (holotype, 10.2, 990 /urn
long).
Scale A (200 pm; x66), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 21 Cypridea unicostata chinensis (3 of 4)
Fig. 1, LV, int. lat. (10.3, 935 ^tm long); fig. 2, car., ext. vent. (10.6, 950 yum long); fig. 3, car., ext. rt. lat. (10.5, 890 yum long).
Scale A (200 /Am; x70), figs. 1-3.
Stereo- Atlas of Ostracod Shells 17, 20
Cypridea unicostata chinensis (2 of 4)
Cypridea unicostata chinensis (4 of 4)
Stereo-Atlas of Ostracod Shells 17, 22
Stereo- Atlas of Ostracod Shells 17 (6) 23-30 (1990) Sunliavia tumida (1 of 8)
595.337.12 (116.312/313) (510 : 161.125.44) : 551.312
ON SUNLIAVIA TUMIDA SOU
by Su Deying & John W. Neale
(Institute of Geology, Chinese Academy of Geological Sciences, Beijing, China &
University of Hull, England)
Genus SUNLIAVIA Sou, 1959
Type-species (original designation): Sunliavia tumida Sou, 1959
1959 Sunliavia gen. nov. Sou Zhixi, in M. A. Netchaeva, Liu Zhongyun, Su Deying, Sou Zhixi, Tian Guizhen & Tsao Lianbi, Lower
Cretaceous Ostracoda from the Songliao Basin, Geol. Publ. House, Beijing, 48.
1974 Sunliavia Sou; Hao Yichun, Su Deying, Li Yougui, Ruan Peihua & Yang Fengtian, Cretaceous -Tertiary Ostracoda from the
Songliao Basin, Geological Press, Beijing, 80.
Diagnosis: Carapace tumid, sub-triangular in shape, dorsal margin strongly arched, with strong over-reach of
right valve over left, with this accentuated at cardinal angles. Internal details poorly known.
Remarks: Extensive dorsal over-reach of one valve by the other is seen in Cyprinotus Brady, 1886 (7. Linn.
Soc., 19, 301) but here the right valve is the larger and the nature of the over-reach as well as the
shape of the carapace is quite different. Sunliavia is closest to Limnocypridea Lubimova, 1956 (in
Mandelstam et al., Trudy vses. nauchno-issled. geol. Inst., n.s., 12, 106), from the Early
Cretaceous Dzunbainsk Formation of Mongolia. In the latter the left valve also strongly overlaps
the right but the overlap along the anterior and posterior margins is more pronounced, the valves
Explanation of Plate 17, 24
Figs. 1-3, d" car. (holotype, 184, 1145 /am long): fig. 1, ext. rt. lat.; fig. 2, ext. dors.; fig. 3, ext. It. lat.
Scale A (200 gm; x57), figs. 1-3.
Stereo- Atlas of Ostracod Shells 17, 25 Sunliavia tumida (3 of 8)
are subrectangular with a relatively straight dorsal margin, and in dorsal view the carapace is much
less tumid. Sunliavia, in contrast, shows a much more arched margin in the right valve with
consequent accentuation of over-reach by the left valve at the cardinal angles, the shell is more
rounded-triangular than subrectangular in side view and is far broader proportionally in dorsal
view. Lubimova (op. cit.) placed Limnocypridea in the Cyprididae, subfamily Cyprideinae, and it
is here that we are placing Sunliavia until further information, particularly concerning its internal
features, becomes known.
Sunliavia tumida Sou, 1959
1959 Sunliavia tumida sp. nov. Sou Zhixi, in M. A. Netchaeva, Liu Zhongyun, Su Deying, Sou Zhixi, Tian Guizhen & Tsao Lianbi,
Lower Cretaceous Ostracoda from the Songliao Basin, Beijing, 48, pi. 15, figs. 4a-d.
1974 Sunliavia tumida Sou ; Hao Yichun, Su Deying, Li Yougui, Ruan Peihua & Yang Fengtian, Cretaceous -Tertiary Ostracoda from
the Songliao Basin, Beijing, 80, pi. 30, figs. la-e.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Institute of Geology, Chinese Academy of Geological Sciences, Beijing, no. 184; d" carapace.
Yaojia Formation, Dekhoi (lat. 44°31'N, long. 125° 40' E), Songliao Basin, China (see Text-
fig. 1); Aptian/ Albian.
Institute of Geology, Chinese Academy of Geological Sciences, Beijing, nos. 184 (holotype, C?
car.: PI. 17, 24, figs 1-3), 10.33 ($ car.: PI. 17, 26, fig. 1), 10.38 (cf car. juv.: PI. 17, 26, fig. 2; PI.
17, 28, fig. 2), 10.39 (9 car. juv.: PI. 17, 26, fig. 3; PI. 17, 30, fig. 3), 10.34 (9 car.: PI. 17, 28, fig.
1; PI. 17, 30, fig. 1), 10.37 (9 LV: PI. 17, 28, fig. 3; PI. 17, 30, fig. 2). All from the type locality and
horizon.
Carapace sub-triangular in lateral view, greatest height anteriorly or centrally, greatest length
ventrally, dorsal margin strongly arched; in dorsal view, tumid. Left valve rather larger than right
Explanation of Plate 17, 26
Fig. 1, 9 car., ext. It. lat. (10.33, 1040 ;u,m long); fig. 2, c? juv. car., ext. dors. (10.38, 950 gm long); fig. 3, 9 juv. car., ext. rt. lat.
(10.39, 950 n.m long).
Scale A (200 /xm; x64), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 24
Sunliavia tumida (2 of 8)
Stereo- Atlas of Ostracod Shells 17, 26
Sunliavia tumida (4 of 8)
Stereo-Atlas of Ostracod Shells 17, 27
Sunliavia tumida (5 of 8)
Cypridea unicostato
chmensis
Cretaceous outcrops
Songliao Basin ■
A Sunliavia tumida
Text-fig. 1: Map and stratigraphic section showing, respectively, the type locality and horizon of Sunliavia tumida. Map also shows
type locality of Cypridea unicostata chinensis (see J. W. Neale & Su Deying, Sterero-Atlas Ostracod Shells, 17, 19-22, 1990).
Explanation of Plate 17, 28
Fig. 1, $ car., ext. rt. lat. (10.34, 1025 yam long); fig. 2, cf juv. car., ext. rt. lat. (10.38, 950 /am long); fig. 3 9 LV, ext. lat. (10.37,
1055 /u,m long). Scale A (200 /am; x64), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 29 Sunliavia tumida (7 of 8)
valve, over-reach accentuated at dorsal cardinal angles. Adults large (>lmm), surface of valves
with delicate reticulum.
Remarks: The most obvious feature of this species is the very strong over-reach of the right valve by the left
in the dorsal region (PI. 17, 24, fig. 1; PI. 17, 28, fig. 1) and the delicate reticulate ornament seen in
well preserved specimens (PI. 17, 28, figs. 2, 3). The forms which are narrower posteriorly and
have a ventral bulge are provisionally interpreted as males (PI. 17, 24, figs. 1-3), while the forms
which maintain the height more posteriorly are regarded as females (PI. 17, 28, figs. 1, 3). The
penultimate instars also appear to show this dimorphism, with presumed males (PI. 17, 28, fig. 2)
and females (PI. 17, 26, fig. 3). All material is in the form of carapaces except for one single valve
(PI. 17, 28, fig. 3 ; PI. 17, 30, fig. 2). Careful removal of the infilling material reveals that it contains
five valves stacked one inside the other. The two larger ones are left valves of an adult and the
penultimate instar, the three smaller ones are juvenile right valves. The adult valves show that the
larger left valve accommodated the smaller right valve by means of a simple tongue and groove
arrangement all around the shell with some expansion of the groove at the extremities of the dorsal
margin. The smaller right valves, with the positive elements, show this expansion clearly in the
form of simple elongate tooth plates. Unfortunately it has not been possible to determine the
nature of the muscle scar pattern. Some possible traces are seen in PI. 17, 28, fig. 2 but are
indistinct.
Distribution: Yaojia Formation, lower part of the Songhuajiang Group ( Aptian/ Albian), Songliao Basin (see
Text-fig. 1), associated with Cypridea acclinia, C. ( M ). dekhoinensis , C. tenuis, C. lunata and
Ziziphocypris simakovi which form part of Assemblage 4 of the Mesozoic non -marine Ostracoda
of China (see Hao Yichun et al., 1974, op. cit.).
Acknowledgement: We wish to express our thanks to the W. C. Wong Foundation who kindly provided a Royal
Society Fellowship for Dr Su to study in Hull.
Explanation of Plate 17, 30
Fig. 1, 9 car., ext. dors. (10.34, 1025 gm long); fig. 2, 9 LV, int. lat. (10.37, 1055 pm long); fig. 3, 9 juv. car., ext. vent. (10.39,
950 /x.m long). Scale A (200 /u,m; x64), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 28
Sunliavia tumida (6 of 8)
Stereo-Atlas of Ostracod Shells 17, 30
Sunliavia tumida (8 of 8)
Stereo-Atlas of Ostracod Shells 17 (7) 31-40 (1990) Theriosynoecum conopium (1 of 10)
595.337.14 (116.222) (411 : 162.007.57) : 551.312 + 552.52
ON THERIOSYNOECUM CONOPIUM WAKEFIELD & ATHERSUCH sp. nov
Fig. 1, cf car., ant. (holotype, OS 13463, 1218/i.m long); fig. 2, cf car., post. (OS 13463); fig. 3, cf car., ext. lat. (OS 13463); fig. 4, cf
car., ext. lat. (OS 13463); fig. 5, cf car., dors. (OS 13463); fig. 6, cf car., vent. (OS 13463); fig. 7, cf car., ornament (OS 13463).
Scale A (200 /u,m; x38), figs. 1-6; scale B (100 fim; x 120), fig. 7.
Stereo-Atlas of Ostracod Shells 17, 33 , Theriosynoecum conopium (3 of 10)
Fig. 1, 9 car., ant. (paratype, OS 13464, 1045 pm long); fig. 2, $ car., post, (paratype, OS 13465, 1082 pm long); fig. 3, 9 car., ext.
lat. (OS 13464); fig. 4, 9 car., ext. lat. (OS 13465); fig. 5, 9 car., vent. (OS 13464); fig. 6, 9 car., dors. (OS 13465); fig. 7, 9 car.,
vent, oranment (OS 13464); fig. 8, 9 car., dors, ornament (OS 13465).
Scale A (200 /u.m; x44), figs. 1, 3, 5; scale B (200 /urn; x43), figs. 2, 4, 6; scale C (50 /urn; x200), fig. 7; scale D (100 /urn; x 140),
fig. 8.
Stereo-Atlas of Ostracod Shells 17, 34 Theriosynoecum conopium (4 of 10)
Stereo- Atlas of Ostracod Shells 17, 35
Theriosynoecum conopium (5 of 10)
Text-fig. 1. Ontogeny of a
topotype populations of T.
conopium , showing sexual
dimorphism.
Text-fig. 2. Sexual dimorph-
ism of a topotype popula-
tion of T. conopium.
Explanation of Plate 17, 36
Fig. 1, A-l car., ext. lat. (paratype, OS 13466, 709 gm long); fig. 2, A-l car., dors. (OS 13466); fig. 3, A-l car., tubercle (OS 13466);
fig. 4, A-l car., anterior margin pore conuli (OS 13466); fig. 5, A-l RV, int. lat. (OS 13478, 700 pm long); fig. 6, A-l, oblique
int. ant. (OS 13478).
Scale A (100 gm; x 65), figs. 1, 2; scale B (10 gm; x 650), fig. 3; scale C (10/Ltm; x 550), fig. 4; scale D (100 gm; x 74), fig. 5; scale E
(100 /i.m; x90), fig. 6.
Stereo-Atlas of Ostracod Shells 17, 37 Theriosynoecum conopium (7 of 10)
1, 205-212, 1973), differs from T. conopium in having a strong longitudinal ridge ventro-laterally.
The muscle scars are typical of the genus (Text-fig. 5).
Distribution: Known only from the Duntulm and Kimaluag formations, Great Estuarine Group, Trotternish,
Skye; Laig Gorge, Eigg; and Camas Mor, Muck, Scotland.
A presumed low salinity species found in association with Darwinula and an undescribed
species of conchostracan (J. D. Hudson pers. comm.). The Kilmaluag Formation has been
intepreted as having been deposited in shallow, freshwater lagoons (Andrews, 1985). Ostracod
valves in the lagoons were constantly agitated by shoreline waves resulting in imbricate stacks (cup
in cup) of several instars; of the type described by Guernet & Lethiers (Bull. Soc. geol. Fr., 8,
557-588, 1989).
Acknowledgement: M. I. Wakefield thanks NERC and BP for CASE studentship support. Drs. R. G. Clements and
David J. Siveter are thanked for their constructive comments.
Text-fig. 4. Differences in posterior
outline of the named Theriosy-
noecum species. (British Museum
(Nat. Hist.) collection numbers).
lo 2284
10 2285
T. anglica
C C "
10 2275 OS 13464
c c -
10 2277 OS 13463
Juveniles
lo 2280 OS 13466
T. ancasterensis T. conopium
CO
CO
o o
100 pm
Text-fig. 5. Muscle scars of
T. conopium 9 LV.
Explanation of Plate 17, 38
Fig. 1, A- 2 car., ext. lat. (paratype, OS 13467, 655 pm long); fig. 2, A- 2 car., ornament and sieve pores (OS 13467); fig. 3, A-l LV,
int. lat. “cup-in-cup” valves (paratype, OS 13468, 909 pm long); fig. 4, A-l LV, int. lat. hinge (OS 13468); fig. 5, A-l RV, ext.
lat. (paratype, OS 13469, 709 gm long); fig. 6, $ car., ext. lat. (paratype, OS 13470, 1127 /u.m long); fig. 7, $ car., post, cardinal
angle (OS 13470).
Scale A (100 gm; x70), fig. 1; scale B (10 /xm; x 500), fig. 2; scale C (100 /urn; x50), fig. 3; scale D (50 pm; x 115), fig. 4; scale E
(lOOpun; x65), fig. 5; scale F (200/a.m; x40), fig. 6; scale G (50/i.m; x 160), fig. 7.
Stereo- Atlas of Ostracod Shells 17, 36
Theriosynoecum conopium (6 of 10)
Stereo-Atlas of Ostracod Shells 17, 38
Theriosynoecum conopium (8 of 10)
Text-fig. 6. (see also Text-fig. 7.) Position of tubercles (centred on pores) and pore conuli for species of Theriosynoecum. Because of
the variable quality of photographic reproduction, some pores may not have been recognised in published papers. A best fit
position for the pores in species of Theriosynoecum is also given.
Those species indicated (*) were originally described under Bisulcocypris. References from which Text-figs. 5 and 6 are drawn:
T. fittoni I. G. Sohn & F. W. Anderson, Palaeontology , 7, 78-84, 1964.
T. anglica and T. ancasterensis R. H. Bate, Bull. Br. Mus. nat. Hist. (Geol.), 14, 29-33, pis. 2 and 3, 1967.
T. wyomingense P. C. Sylvester-Bradley, Stereo-Atlas Ostracod Shells , 1, 205-212, 1973.
T. kirtlingtonense R. H. Bate, Stereo-Atlas Ostracod Shells, 1, 221-228, 1973.
T. bathonicum P. C. Sylvester-Bradley, Stereo-Atlas Ostracod Shells, 1, 229-236, 1973.
Remaining species- I. D. Pinto & Y. T. Sanguinetti, Publishes Esp. Esc. Geol. P. Alegre, 4, 1-165, 17 pis. 1962.
Stereo-Atlas of Ostracod Shells 17, 40
Theriosynoecum conopium (10 of 10)
T. alleni
T. roberti cornata
T. pahasapensis
T. forbesi forbesi
T. verrucosa
T. wyomingense
T. praetuberculata
Position of pores
for the genus
Text-fig. 7. (see also Text-fig. 6). Position of tubercles (centred on pores) and pore conuli for species of Theriosynoecum. Because of
the variability of photographic reproduction in published papers, some pores may not have been recognised. A best fit position
for the pores in species of Theriosynoecum is also given.
For references see explanation for Text-fig. 5.
Stereo-Atlas of Ostracod Shells 17 (8) 41-44 (1990) Darwinula incurva (1 of 4)
595.337.11 (116.222) (411 : 162.006.57 + 420.162.001.53) : 551.312.4 + 552.52
ON DARWINULA INCURVA BATE
by Matthew I. Wakefield
(University of Leicester, England)
Darwinula incurva Bate, 1967
1965 Darwinula sp. A; R. H. Bate, Palaeontology, 8, 751, pi. 109, figs. 1-4.
1967 Darwinula incurva sp. nov. R. H. Bate, Bull. Br. Mus. nat. Hist. (Geol.), 14, 28-29, pi. 1, figs. 7-12.
Holotype: British Museum (Nat. Hist.), no. Io 2259; carapace.
[Paratypes: nos. Io 2260-74.]
Type locality: Admix Refractories Clay Pit, King’s Cliffe, Northamptonshire, England; National Grid
Reference: TL 012,966; lat. 52°38'N, long. 0°31'W. Bed 7 (of P. C. Sylvester-Bradley, D. F.
Merriam & C. J. Aslin, Geol. Ass. Guide, Uppingham Area, University of Leicester, 27,
1968), Upper Estuarine Series, Bathonian, middle Jurassic.
Figured specimens: British Museum (Nat. Hist.), nos. Io 2259 (holotype, car.: PI. 17, 42, figs. 3, 4), Io 2262 (paratype,
RV: PI. 17, 44, fig. 3), OS 13461 (LV: PI. 17, 42, figs. 1, 2), OS 13460 (RV: PI. 17, 42, figs. 5, 6),
OS 13462 (RV: PI. 17, 44, fig. 1) and OS 13459 (car.: PI. 17. 44, fig. 2). Io 2259 and Io 2262 are
from the type locality. OS 13459-OS 13462 are from the middle Jurassic, Great Estuarine Group,
Laig Gorge, Eigg, lat. 6° 08' W, long. 56° 55' N. OS 13459 is from the top 5 cm of Bed 1, Kilmaluag
Formation. OS 13460 and OS 13461 are from 20cm above the base of Bed 21, Duntulm formation.
Explanation of Plate 17, 42
Fig. 1, LV, ext. lat. (OS 13461, 836 /am long); fig. 2, LV, vent. (OS 13461); fig. 3, car. ext. lat. (holotype, Io 2259, 1009 /am long); fig.
4, car. vent. (Io 2259); fig. 5, RV, ext. lat. (OS 13460, 818 /am long); fig. 6, RV, vent. (OS 13460).
Scale A (200 jum; x73), figs. 1, 2, 5, 6; scale B (200 /am; x60), figs. 3, 4.
Stereo- Atlas of Ostracod Shells 17, 43 Darwinula incurva (3 of 4)
Diagnosis:
Remarks:
Distribution:
Acknowledgement:
OS 13462 is from 65 cm above the base of Bed 5, Kilmaluag Formation of J. E. Andrews ( Aspects
of Sedimentary Facies and Diagenesis in Limestone - Shale Formations of the Middle Jurassic Great
Estuarine Group, Inner Hebrides , Unpubl. PhD Thesis, University of Leicester, 1984).
Darwinula with convex dorsum and ventral inflexure anterior to mid-line. Left valve larger than
and overlapping right valve, with maximum overlap at ventral inflexure. Adductor muscle-scar
rosette has 11 segments.
The adductor muscle-scar of D. incurva occurs immediately above the ventral inflexure and is
moderately variable. Occasionally only 10 segments are present in the muscle-scar; however, the
basal segment clearly shows a dividing line which in other specimens results in the occurrence of
the more typical 11 segment muscle-scar morphology. Further sub-divisions are observed in the 2
segments immediately anterior to the basal segments (PI. 17, 44, figs. 1-3).
D. incurva is similar to Darwinula tubiformis Ljubimova (P. S. Ljubimova, Trudy vses. neft.
nauchno-issled. geol.-razv. Inst., n.s., 93, 119, 120, 1956) but is less inflated posteriorly, the LV
ventral overlap is more prominent and the inflexure is closer to the anterior margin.
A freshwater species, found in association with the ostracods of the genus Theriosynoecum. Found
in the Upper Estuarine Series, King’s Cliffe, Northamptonshire, England and the Duntulm and
Kilmaluag formations, Great Estuarine Group, Eigg, Scotland (Wakefield in prep.).
NERC & BP for CASE studentship support and David J. Siveter for helpful comments.
Explanation of Plate 17, 44
Fig. 1, damaged car. showing int. mould of muscle-scar of RV (OS 13462, 836 /am long); fig. 2, RV, int. mould of muscle-scar
(OS 13459, 764 /am long); fig. 3, RV, int. muscle-scar (Io 2262, damaged valve).
Scale A (50 /am; x395), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 44 Darwinula incurva (2 of 4)
Stereo- Atlas of Ostracod Shells 17 (9) 45-52 (1990) Londinia kiesowi (1 of 8)
336.11 (113.333) (430.2 : 161.013.54 + 014.54 + 438 : 161.015.54 + 485) : 551.351 + 552.54
ON LONDINIA KIESOWI (KRAUSE)
by Wolfgang Hansch & David J. Siveter
(University of Greifswald, German Democratic Republic & University of Leicester, England)
Londinia kiesowi (Krause, 1891)
1891 Kloedenia kiesowi sp. nov. A. Krause, Z. dt. geol. Ges., 43 (2), 506-507, 518-519, pi. 32, figs. 12, 13.
1963 Londinia kiesowi (Krause); A. Martinsson, Bull. geol. Instn Univ. Uppsala, 42, 1, 20-24, 28, figs. 7A, 9, 10, 13B.
1977 Londinia kiesowi (Krause); A. Martinsson, The Silurian- Devonian Boundary, IUGS Series A, no. 5, 46, 328.
1986 Londinia kiesowi (Krause) ; R. E. L. Schallreuter, Mitt. geol. -palaont. Inst. Univ. Hamburg, 61, 200-201, pi. 2, fig. 1, pi. 6, fig.
8a (q.v. for full synonymy).
1989 Londonia kiesowi (Krause) ; D. J. Siveter, in : M. G. Bassett & C. H. Holland (Eds.), A global standard for the Silurian System,
263, fig. 168H, Nat. Mus. Wales, Geol. Series, no. 9, Cardiff.
Lectotype: Museum fur Naturkunde Berlin, German Democratic Republic (GDR), no. M.B.O. 143; 9 LV.
Krause, 1891, pi. 32, fig. 13; Martinsson, 1963, 22, fig. 10A.
[Paratypes: Museum fur Naturkunde Berlin, M.B.O. 144, tecnomorphic LV; Krause, 1891, pi.
32, fig. 12 and Martinsson, 1963, 22, fig. 10B. M.B.O. 145, $ RV; Martinsson, 1963, 19, fig. 7A,
28, fig. 13B. British Museum (Nat. Hist.), I 6022, 9 RV.]
Type locality: Erratic boulder no. 549 of Krause, 1891. From ‘Klein -Horst’, Poland, lat. 54°6'N, long.
15° 4.5' E.
Explanation of Plate 17, 46
Figs. 1, 2, 9 LV (SGWG 83/13, 2880 yum long); fig.l, ext. lat.; fig. 2, ext. ant. Figs. 3, 4, 9 LV (SGWG 83/14, 2930 long): fig. 3,
ext. ant.; fig. 4, ext. lat.
Scale A (500 /xm; x23), figs. 1-4.
Stereo- Atlas of Ostracod Shells 17, 47 Londinia kiesowi (3 of 8)
Figured specimens: Sektion Geologische Wissenschaften der E.-M.-Arndt-Universitat Greifswald (SGWG), GDR,
nos. SGWG 83/13 (9 LV: PI. 17, 46, figs. 1, 2; PI. 17, 48, fig. 6) and SGWG 83/15 (cf RV; PI. 17,
48, figs. 1-3), from erratic boulder no. Bey. A15, Koserow, Isle of Usedom, GDR; lat. 54° 03' N,
long. 14° 0' E. SGWG 83/14 (9 LV: PI. 17, 46, figs. 3, 4; PI. 17, 48, figs. 4, 5), SGWG 83/17 (9 LV:
PI. 17, 50, figs. 4, 5, 7), SGWG 83/18 (9 LV: PI. 17, 50, fig. 6) and SGWG 83/19 (9 RV: PI. 17, 52,
figs. 6, 7), from erratic boulder no. Bey. D54, Gager, Isle of Ruegen, GDR; approx, lat. 54° 17' N,
long. 13° 45' E. SGWG 83/16 (cf LV: PI. 17, 48, fig. 7; PI. 17, 50, figs. 1-3), from erratic boulder
no. Bey. D49, Gager, Isle of Ruegen, GDR. British Museum (Nat. Hist.), no. I 6022 (9 RV: PI.
17, 52, figs. 1-3), from erratic boulder no. 549 of Krause (1891, op. cit.). Paleont. Mus. Univ.
Lund, no. LO 2182t (9 LV: PI. 17, 52, figs. 4, 5; figured J. C. Moberg & K. A. Gronwall, Acta
Univ. lund., N.F., 5 (1), pi. 4, fig. 17, 1909), from (Oved-Ramsasa) bed 4 of Moberg & Gronwall
(1909, op. cit.; = Oved Sandstone Formation, Oved-Ramsasa Group, of Jeppsson & Laufeld,
Sver. geol. Unders. Afn., Ser. Ca, 58, 1987), Scane, S Sweden. All specimens are from the Pridoli
Series, Silurian.
Diagnosis: Londinia species with symmetrically arranged lobes on both sides of the adductorial sulcus which
hardly, if at all, protrude over the dorsal margin. Generally lobes do not show distinct lateral
surface facets or reticulation except on the cuspidal part of the preadductorial lobe and anterior
lobule of the syllobium. Adductorial sulcus does not reach below valve mid-height.
Remarks: A detailed description of L. kiesowi based on the original erratic boulder material from Krause
(1891, op cit.) was given by Martinsson (1963, op. cit.). Our investigations of material from the
Leba 1 borehole in the Peribaltic area of Poland (see Martinsson, Geol. For. Stockh. Forh., 86,
(2), 138, figs. 7A-C, 1964), erratic boulder material from the T. R. Jones collection (British
Museum (Nat. Hist.)), London, the figured types of Moberg & Gronwall (1909, 64-66 pars, 81,
Explanation of Plate 17, 48
Figs. 1-3, cf RV ( SGWG 83/15, 2730 /j,m long): fig. 1, ext. lat. ; fig. 2, ext. vent. ; fig. 3, ext. ant. Figs. 4, 5, 9 LV (SGWG 83/14): fig.
4, ext. vent. ; fig. 5, detail of cuspidal parts of preadductorial lobe and ant. syllobial lobule. Fig. 6, 9 LV, detail of cuspidal parts of
preadductorial lobe and ant. syllobial lobule (SGWG 83/13). Fig. 7, cf LV, ext. post. (SGWG 83/16, 3080 )j.m long).
Scale A (500 pun; x23), figs. 1-4; scale B (215 /u.m; x40), figs. 5, 6; scale C (375 /xm; x24), fig. 7.
Londinia kiesowi (2 of 8)
Stereo-Atlas of Ostracod Shells 17, 46
Stereo-Atlas of Ostracod Shells 17, 48
Londinia kiesowi (4 of 8)
Stereo-Atlas of Ostracod Shells 17, 49 Londinia kiesowi (5 of 8)
86, pi. 4, figs. 16, 17; cf. Martinsson, 1963, 23-24) and newly collected Baltic erratic boulder
specimens indicate a bigger morphological variability of this species than that noted by Martinsson
(1963, op. cit.).
Variation affects especially the shape and the size of the crumina, the nature of the cuspidal
part of the two main lobes, and the morphology of the adductorial sulcus. The crumina extends
either from a point almost vertically below the anterior end of the valve or the anterior limit of the
anterior lobe to about the centre of the anterior lobule of the syllobium. The width of the
subcruminal fingerprint-like striation and the distinctness of the dorsal delimination of the
crumina from the domicilial part of the valve varies with cruminal size. In some specimens the
preadductorial lobe and the anterior lobule of the syllobium are higher and show well developed
cuspidal facets. In such specimens a fine, dorsal, striate or sometimes reticulate lobal field is visible
and is bounded by two weakly swollen bends as in other kloedeniine species such as Klodenia
leptosoma Martinsson, 1963. Such specimens also display a syllobial anterior lobule which is less
angular dorsally than its equivalent preadductorial lobe (cf. PI. 17, 50, figs. 6, 7). In tecnomorphs
differences in lobal faceting and ornament are less pronounced than in females. The adductorial
sulcus becomes deeper and longer during ontogeny. One heteromorphic (9) valve found together
with other ‘typical’ specimens of L. kiesowi shows an extremely indistinct lobal and cruminal
development (cf. PI. 17, 52, figs. 6, 7). Unfortunately there is no more material available but
probably it is currently necessary to regard it as an extreme variant.
L. kiesowi differs from the type-species of Londinia ( L . reticulifera Martinsson, 1963) mainly
by its shorter adductorial sulcus in adult specimens, by the lack of lobal reticulation and by the
limitation of the lobes to generally below the dorsal margin. Martinsson (1963, 26-28, op. cit.)
mentioned that the typical features of L. reticulifera becorhe obvious only in the last moult stage.
For this reason a species assignment of Londinia specimens based only on tecnomorphic specimens
Explanation of Plate 17, 50
Figs. 1-3, cf LV (SGWG 83/16): fig. 1, ext. lat. ; fig. 2, ext. vent.; fig. 3, detail of cuspidal part of preadductorial lobe. Figs. 4, 5, 7, 9
LV (SGWG 83/17, approx. 2900 /a,m long): fig. 4, ext. lat. ; fig. 5, detail of ventral side of crumina; fig. 7, detail of cuspidal part of
preadductorial lobe. Fig. 6, 9 LV, detail of cuspidal part of ant. syllobial lobule (SGWG 83/18, 3090 /um long).
Scale A (500;u,m; x 20), figs. 1, 2, 4; scale B (40 /xm ; x 240), fig. 3; scale C (150 gm; x 60), fig. 5; scale D (100 /a.m; x 90), figs. 6, 7.
Stereo-Atlas of Ostracod shells 17, 51 Londinia kiesowi (7 of 8)
is difficult. The taxonomic position of the Londinia tecnomorphs occurring together with
Poloniella (Hoia) hieroglyphica (Krause, 1891) (Schallreuter, 1986, op. cit.) is, therefore
uncertain. However, if that association should be confirmed by finds of adults of L. kiesowi ,
together with well preserved P. (H.) hieroglyphica specimens, the latter species would not be
suitable as an index species for the upper most ostracode association of the East Baltic area (cf. L.
Sarv in: D. Kaljo & E. Klaamann (Eds.), Ecostratigraphy of the East Baltic Silurian, 77 -78,
Tallinn (Valgus), 1982).
Martinsson ( Geol . For. Stockh. Forh . , 89, 379, 1967) considered that L. kiesowi is conspecific
with L. arisaigensis Copeland (Bull. geol. Surv. Can., 117, 1964) from Nova Scotia, Canada.
However, particularly the shape and size of the lobes seem to be different between these species
(cf. Schallreuter, 1986, op. cit.). Nevertheless, it is obvious that these species are closely related
and because of their limited stratigraphic distribution at the base of the Pridoli Series they are, like
Frostiella groenvalliana Martinsson, 1963, important species for regional correlation between
Canada, Great Britain and the Baltic area at that horizon (cf. Martinsson, 1977 ; Siveter, 1989, op.
cit.). It is also possible that the British species described under Londinia by R. W. L. Shaw (Geol.
For. Stockh. Forh., 91, 1969) may hide conspecific material, but this requires further study.
Distribution: Late Silurian (Pridoli Series), Baltascandia region.
Sweden: Scania, Upper Oved-Ramsasa Beds 3-4 sensu Martinsson (1967, op. cit.).
Peribaltic area of Poland: Leba 1 borehole, ‘intermediate fauna’, 686.15m - 703.60m
(Martinsson, 1964, op. cit.); Leba 2 borehole, post-Ludlow, Neobeyrichia incerta Zone (B.
Zbikowska, Acta geol. pol., 23, 628-629, tab. 2, 1973).
Erratic boulders, Beyrichienkalk type B and ‘Red Beyrichienkalk’ sensu W. Hansch (Lethaia,
18, 274, tab. 1, 1985); erratic boulder no. Sy58, Isle of Sylt (Schallreuter, 1986, op. cit.).
Explanation of Plate 17, 52
Figs. 1-3, 9 RV (paratype, I 6022, 2700 /i.m long): fig. 1, ext. post.; fig. 2, ext. lat.; fig. 3, ext. vent. Figs. 4, 5, 9 LV (LO 2182t,
2560 ixm long): fig. 4, ext. lat. ; fig. 5, ext. vent. Figs. 6, 7, 9 RV (SGWG 83/19, 2580 /urn long): fig. 6, ext. lat. ; fig. 7, ext. vent.
Scale A (375 ^m; x24), fig. 1; scale B (500 gm; x21), figs. 2-7.
1
Stereo-Atlas of Ostracod Shells 17 (10) 53-60 (1990) Hemsiella maccoyiana (1 of 8)
336.1.11 (113.333) (430.2 : 161.013.54 + 014.52 + 438 : 161.017.51) : 551.351 + 552.54
ON HEMSIELLA MACCOYIANA (JONES)
by David J. Siveter & Wolfgang Hansch
(University of Leicester, England & University of Greifswald, German Democratic Republic)
Hemsiella maccoyiana (Jones, 1855)
1855 Beyrichia maccoyiana nov. sp. T. R. Jones, Ann. Mag. nat. Hist., ser, 2, 16, 88, pi. 5, fig. 14.
1855 Beyrichia dalmaniana nov. sp. T. R. Jones, Ibid. , 88, pi. 5, fig. 13.
1862 Beyrichia maccoyiana Jones; E. Boll. Arch. Ver. Freunde Naturg. Mecklenb., 16 (7), 134, pi. 1, fig. 9.
1862 Beyrichia dalmaniana Jones; E. Boll, Ibid., 127, pi. 1, fig. 15.
1909 Beyrichia maccoyiana Jones; J. Ch. Moberg & K. A. Gronwall, Acta Univ. lund. , N.F., 5 (1), 58, 81, 86, pi. 4, fig. 8, pi. 6, fig. 4; non fig. 5
(= Hemsiella hians (Boll, 1856)).
1957 Beyrichia maccoyiana sulcata Reuter; R. V. Kesling, Contr. Mus. Paleont. Univ. Mich., 14 (6), pi. 4, figs. 7-9.
1957 Beyrichia maccoyiana sulcata Reuter; R. V. Kesling & K. J. Rogers, J. Paleont., 31 (5), 1002-1003, pi. 128, figs. 1, 2, pi. 129, figs. 12-14.
1962 Hemsiella maccoyiana (Jones); A. Martinsson, Bull. geol. Instn Univ. Uppsala, 41, 16-17, 221, 223, fig. 2B.
1962 Hemsiella dalmaniana (Jones); A. Martinsson, Ibid., 16-17, 221, 223, fig. 2C.
1962 Beyrichia (Neobey richia) maccoyiana var. sulcata Reuter; M. J. Copeland, Palaeontology , 3, 99, pi. 23, figs. 14-16.
1964 Hemsiella cf. maccoyiana (Jones); A. Martinsson, Geol. For. Stockh. Forh., 86 (2), 133-135, figs. 4a-c.
1964 Hemsiella maccoyiana (Jones); A. Martinsson, Ibid., 86, 241-242, fig. 2F.
1964 Hemsiella maccoyiana sulcata Reuter; M. J. Copeland, Bull. geol. Surv. Can., 117, 8-9, pi. 1, fig. 3.
1965 Hemsiella dalmaniana (Jones); L. Gailite, Latv. PSR zinat. Akad. Vest., 2 (211), 68-70.
1966 Hemsiella maccoyiana (Jones) (= dalmaniana ) ; D. Kaljo & L. Sarv, Eesti NSV Tead. Akad. Toim. , ser. Tekhn. Fiz.-Mat. 2, 279, tab. 1.
1967 Hemsiella maccoyiana (Jones); A. Martinsson, Geol. For. Stockh. Forh., 89 (3), 375-377.
1967 Hemsiella dalmaniana (Jones), 1855; L. Gailite, in: L. Gailite, M. Rybnikowa & R. Ulste, Stratigrafija, fauna i uslovija obrazovanija
silurijskich porod srednej Pribaltiki, 121-122, pi. 8, figs. 2a-e, Riga (Zinatne).
Explanation of Plate 17, 54
Figs. 1-5, 9 car. (SGWG 83/7, 2020 /am long): fig. 1, LV ext. lat. ; fig. 2, ext. vent. ; fig. 3, RV ext. lat., fig. 4, cusp of ant. lobe of LV; fig. 5, ext. RV,
vent. obi. Scale A (375 pm: x26), figs. 1-3, 5; scale B (75 /am; x 120), fig. 4.
Stereo-Atlas of Ostracod Shells 17, 55 Hemsiella maccoyiana (3 of 8)
1968 Hemsiella cf. maccoyiana (Jones); L. Sarv, Ostrakody Craspedobolbinidae, Beyrichiidae i Primitiopsidae silura Estonii, 24, pi. 7, figs. 1-4,
tabs. 2, 3, Tallinn (Valgus).
1970 Hemsiella cf. maccoyiana (Jones); L. Sarv, in: D. Kaljo (Ed.), Silur Estonii, 158, 169-170, 299 Tallinn (Valgus).
1970 Hemsiella dalmaniana (Jones); A. Pranskevicius, Dokl. Akad. Nauk SSSR, 192 (6), 85.
1971 Hemsiella maccoyiana (Jones); R. W. L. Shaw, Palaeontology, 14, 597, pi. 109, figs. 1-4.
1971 Hemsiella cf. maccoyiana (Jones); L. Sarv, Eesti NSV Tead. Akad. Toim., Khim. Geol., 20 (4), 353, 355, tab. 2.
1972 Hemsiella dalmaniana (Jones); A. Pranskevicius, Geol. For. Stockh. Forh., 94, 439.
1972 Hemsiella dalmaniana (Jones); A. Pranskevicius, Trudy LitNIGRI, 15, 20, 35, 77, tabs. 1, 2, fig. 11 (log), pi. 8, figs. 1-3.
1972 Hemsiella dalmaniana (Jones); L. Gailite, Eesti NSV Tead. Akad. Toim., Khim. Geol., 21, 352-353.
1973 Hemsiella dalmaniana (Jones); B. Zbikowska, Acta geol. pot, 23, 610, 614, 623, tab. 2, pi. 4, figs. 1, 2.
1974 Hemsiella dalmaniana (Jones); E. Tomczykowa & E. Witwicka, Biul. Inst, geol., 276, 59, 61, 68, figs. 2, 3 (logs).
1974 Hemsiella dalmaniana (Jones); B. Zbikowska, Bull. Acad. pol. Sci. Ser. Sci. Terre, 22 (1), 47.
1975 Hemseilla dalmaniana (Jones); A. Pranskevicius, Geol. For. Stockh. Forh., 97, 53-54.
1977 Hemsiella cf. maccoyiana (Jones); L. Sarv, in: D. Kaljo (Ed.), Fazii i fauna silura Pribaltiki, 161, 164, 169, 173-175, tabs. 1-7.
1977 Hemsiella maccoyiana sulcata (Reuter); M. J. Copeland & J. M. Berdan, Geol. Surv. Pap. Can., 77-1B, pi. 2, 3, figs. 9, 10.
1978 Hemsiella dalmaniana (Jones); T. I. Moiseeva, in: Stratigraficeskie i paleontologiceskie issledovanija v Belorussii, 59, 178, tab. 7.
1978 Hemsiella dalmaniana (Jones); L. Gailite, in : Stratigrafija fanerozoja Pribaltiki. Paleontologiceskie kompleksy, stroenie i sostav otlozenie,
16, 18-20, Riga (Zinatne).
1982 Hemsiella maccoyiana (Jones); L. Sarv, in: D. Kaljo & E. Klaamann (Eds.), Ecostratigraphy of the East Baltic Silurian, 74-75, Tallinn
(Valgus).
1985 Hemsiella maccoyiana (Jones); W. Hansch, Lethaia, 18, 275, 277, tab. 1.
1985 Hemsiella dalmaniana (Jones); W. Hansch, Ibid., 275, tab. 1.
1986 Hemsiella maccoyiana (Jones); L. Gailite, in: D. Kaljo & E. Klaamann (Eds.), Teorija i onyt ecostratigrafija, 113, Tallinn (Valgus).
1986 Hemsiella dalmaniana (Jones); I. Sidaraviciene, Ibid., 119, 124.
1988 Hemsiella maccoyiana (Jones); W. Hansch, Neues Jb. Geol. Palaont. Mh., 1988, 481-482, 491.
1988 Hemsiella dalmaniana (Jones); W. Hansch, Ibid.
1989 Hemsiella maccoyiana (Jones) ; D. J. Siveter, in : M. G. Bassett & C. H. Holland (Eds.), A global standard for the Silurian System, 263,
fig. 168G, Nat. Mus. Wales Geol. Ser., No. 9, Cardiff.
Explanation of Plate 17, 56
Figs. 1-7, cf car. (SGWG 83/8, 21 10 /am): fig. 1, LV syllobial cusp; fig. 2, detail of velum of LV ; fig. 3, LV ext. lat. ; fig. 4, ext. vent. ; fig. 5, RV ext.
lat.; figs. 6, 7, RV syllobial cusp. Fig. 8, 9 car., LV syllobial cusp (SGWG 83/7).
Scale A (75 /am; x 125), fig. 1; scale B (30 /u.m; X 300), fig. 2; scale C (375 /am; x 26), figs. 3-5; scale D (50 /am; x 180), fig. 6; scale E (15 /am; x 600),
fig. 7; scale F (75 /am; X90), fig. 8.
Stereo-Atlas of Ostracod Shells 17, 56 Hemsiella maccoyiana (4 of 8)
Type locality:
Figured specimens:
Stereo-Atlas of Ostracod Shells 17, 57 Hemsiella maccoyiana (5 of 8)
Lectotype: British Museum (Nat. Hist.), I 6953; tecnomorphic RV. Martinsson, 1962, 16, fig. 2B ; Jones, 1855, pi. 5, fig.
14. [Paratypes: I 7019, 9 RV (Martinsson, 1962, 16, fig. 2C); I 7018, 9 LV (Jones, 1855, pi. 5, fig. 13).]
Erratic boulder no. 2 of Jones, 1855, near Breslau (Wroclaw), Poland; approx, lat. 51°5'N, long. 17°E.
Sektion Geologische Wissenschaften der E.-M.-Arndt-Universitat Greifswald, GDR, nos. SGWG
83/7 (9 car.: PI. 17, 54, figs. 1-5; PI. 17, 56, fig. 8), SGWG 83/8 (cf car.: PI. 17, 56, figs. 1-7) and SGWG
83/10 (cf car.: PI. 17, 58, figs. 4, 7), from erratic boulder no. Bey. BIO, Sellin, Isle of Ruegen, GDR; lat
54° 23' N, long. 13°41' E. SGWG 83/9 (9 RV: PI. 17, 58, figs. 1-3; PI. 17, 60, fig. 4), SGWG 83/11 (9 LV: PI.
17, 58, figs. 5, 6; PI. 17, 60, fig. 3) and SGWG 83/12 (tecnomorphic LV: PI. 17, 60, figs. 1, 2) from erratic
boulder no. Bey B16, Oderberg-Bralitz, GDR; lat. 52°52'N, long. 14°3'N. British Museum (Nat. Hist.),
nos. I 7019 (paratype, 9 RV: PI. 17, 60, figs. 5, 6) and I 7018 (paratype, 9 LV: PI. 17, 60, fig. 9), both from
erratic boulder no. 3 of Jones, 1855, near Breslau (Wroclaw), Poland ; I 6953 (lectotype, tecnomorphic RV:
PI. 17, 60, figs. 7, 8), from erratic boulder no. 2 of Jones, 1855, near Breslau (Wroclaw), Poland.
Hemsiella species in which the syllobium is more or less dissected by a dorsal depression in both sexes. Right
valves show a distinct depression below the rounded, protuding cuspidal part of syllobium, in left valves this
depression is shallower and the cusp not so protusive over the dorsal margin. Crumina shows a wide,
depressed unornamented zone lateral to velar ridge, and is reticulostriate between the velar ridge and valve
margin. Lobes and lateral surface of crumina are either reticulate, reticulostriate, weakly punctate or smooth.
Hemsiella is a well defined genus but one containing many taxa showing wide and often overlapping
intraspecific variation. For example, Hansch (1988) considered that H. hians (Boll, 1856), H. elegans (Boll,
1862) and the type-species H. loensis Martinsson, 1962, were synonymous. Furthermore, Martinsson (e.g.
1962, 223; 1964, 134) noted the wide intraspecific characteristics of H. dalmaniana and H. maccoyiana ,
species for which he erected lectotypes. Both species names have been used in the literature but we prefer to
adopt H. maccoyiana because of the more extensive recent use. H. maccoyiana mclearni Copeland, 1964 and
H. latviensis Gailite, 1967 also are very similar to H. maccoyiana. As noted by Hansch (1988, 481) there are
three groups of Hemsiella species, characterised by H. hians (Boll, 1856), H. maccoyiana (Jones, 1855) and
H. pulchricruminata Martinsson, 1862.
Gailite (1967) was the first to note an assymmetry in the syllobial morphology between right and left
valves of ‘ Hemsiella dalmaniana' specimens from the Piltene 1 borehole in Latvia. Usually the dorsally
depressed, unornamented area is larger in the left valve. Furthermore, the anterior lobe of the left valve is
Explanation of Plate 17, 58
Figs. 1-3, 9 RV (SGWG 83/9, 1640 /um long): fig. 1, ext. lat. ; fig. 2, ext. vent. ; fig. 3, ext. ant. Figs. 4, 7, c f car. (SGWG 83/10, 2360 gm long): fig. 4.
vent, detail; fig. 7, ext. ant. Figs. 5, 6, 9 LV (SGWG 83/11, 2000/u.m long), fig. 5, detail of ventral side of crumina; fig. 6, ext. post.
Scale A (300 /urn; x30), figs. 1-3, 6, 7; scale B (150 /urn; x60), fig. 4; scale C (150/xm; x70), fig. 5.
Diagnosis:
Remarks:
Stereo-Atlas of Ostracod Shells 17, 59 Hemsiella maccoyiana (7 of 8)
typically more prominent than in the right valve. Sometimes the ornament, especially that of the syllobium of
both sexes and of the crumina, is effaced. The denticles of the velar border crest and the tubulosity of the
velum are only visible in tecnomorphs, which also show the typical anteroventral undulation of the velum and
a weak toric ridge which in some tecnomorphs (right valve only) appears to be divided into two (PI. 17, 56,
fig. 4). There is no velum in front of the crumina. In small tecnomorphs the preadductorial node is better
connected with the anterior lobe and the syllobium than in adults.
Distribution: Canada: Upper Stonehouse Formation, Nova Scotia (Copeland, 1964, Copeland & Berdan, 1977).
Great Britain: Upper Underbarrow Flags, Kirkby Moor Flags and Scout Hill Flags, Cumbria; Whitcliffe
Formation and Downtown Castle Sandstone Formation, Long Mountain region; Whitcliffe Formation,
Ludlow area (Shaw, 1969, 1971; see also Siveter, 1980).
Peribaltic area of Poland: Leba 1 borehole; Beyrichienkalk pebbles in the ‘Zechsteinkonglomerat’
(Martinsson, 1964). Lebork and Wejherowo boreholes; post-Ludlow (Witwicka, 1967). Leba 2, 5 and Debki
3 boreholes; post-Ludlow, Neobeyrichia incerta to Nodibeyrichia tuberculata zones (Zbikowska, 1973).
Miloszewo, Wejherowo, Karwia, Opalina, Salino, Bialogard, Leba and Lebork IG 1 boreholes;
post-Ludlow, Neobeyrichia incerta to Kloedenia wilckensiana zones (Tomczykowa & Witwicka, 1974).
Chojnice borehole; post-Ludlow (Zbikowska, 1974).
East Baltic area, U.S.S.R.: Ohesaare 1 and 2 boreholes, Isle of Saremaa, Estonia; Kaugatuma and
Ohesaare formations (Sarv, 1968, 1971). Piltene 1, 31, 32, Ezere 9, Talcy 55, Pavilosta 51, Kolka 1, 54 and
Stoniskjaj boreholes, Latvia; Minija and Jura formations (Gailite, 1967, 1978). Kunkojaj and Virbalis
boreholes, Kaugatuma and Ohesaare formations (Sarv, 1977); Taurage borehole, Minija Formation
(Pranskevicius, 1972); boreholes 94, 96, 98, 110, 112 and 89 (Minija Formation) and boreholes 89, 98, 110,
112 and possibly 108 (Jura Formation), Arjogalskij profile, Lithuania (Sidaraviciene, 1986). Dubovskoje
borehole, Kaliningrad district; Kaugatuma Formation (Kaljo & Sarv, 1976). Brest basin, Muchavez and
Kustin beds (Moiseeva, 1978).
Sweden: Scania, Klinta Formation (?) and Oved Sandstone sensu Jeppsson & Laufeld (Sver. geol.
Unders. Afh., ser. Ca, 58, 1987); see also Moberg & Gronwall, 1909. Erratic boulders, Beyrichienkalk sensu
Martinsson (1963, 1967, 1977); Beyrichienkalk type B, C and “Red Beyrichienkalk” sensu Hansch (1985).
Explanation of Plate 17, 60
Figs. 1, 2, tecnomorphic LV (SGWG 83/12, 1530 gm long): fig. 1, ext. lat. ; fig. 2, ext. vent. Fig. 3, $ LV, ext. lat. (SGWG 83/11). Fig. 4; 9 RV, detail
of crumina (SGWG 83/9). Figs. 5, 6, 9 RV (paratype, 1 7019, approx. 1620 gm long): fig. 5, ext. lat.; fig. 6, ext. vent. Figs. 7, 8, tecnomorphic RV
(lectotype, I 6953, approx. 1510 /am long): fig. 7, ext. lat.; fig. 8, detail of syllobium. Fig. 9, 9 LV, ext. lat. (paratype, I 7018, 1680 /urn long).
Scale A (300 /um; x30), figs. 1, 2; scale B (375 /urn; x25), fig. 3; scale C (30 /urn; x300), fig. 4; scale D (300 /um; x30), figs. 5-7; scale E (75 gm;
x 120), fig. 8; scale F (300/izm; x27), fig. 9.
Stereo- Atlas of Ostracod Shells 17, 58
Hemsiella maccoyiana (6 of 8)
Stereo-Atlas of Ostracod Shells 17 (11) 61-64 (1990) Cytheropteron glintzboeckeli (1 of 4)
595.337.14 (116.331) (64.161.007.39) : 551.35 + 552.54
ON CYTHEROPTERON GLINTZBOECKELI (DONZE & LEFEVRE)
by Richard Symonds
(Institute of Earth Studies, University College of Wales, Aberystwyth)
Cytheropteron glintzboeckeli (Donze & Lefevre, 1981)
1963 Cytheropteron sp. 2, M. Masoli, Mem. Bur. Rech. geol. minier., 32, 122-123, pi. 1, figs. 4a-4d.
1981 Eocytheropteron glintzboeckeli sp. nov. P. Donze & J. Lefevre, in H. Bismuth et al., Cah. Micropaleont. , 1981-3 (2), 58,
pi. 1, figs. 13-16.
1984 Eocytheropteron glintzboeckeli Donze & Lefevre; H. Bismuth, Bull. Cent. Rech. Explor.-Prod. Elf -Aquitaine, 8, 471-472.
(q.v. for full pre-1984 synonymy).
1985 Eocytheropteron glintzboeckeli Donze & Lefevre; J. Viviere, Les Ostracodes du Cretace superieur (Vraconien a Campanien
basal ) de la region de Tebessa (Algerie du Nord-Est): Stratigraphie, Paleoecologie, Systematique, Univ. P. & M. Curie,
Paris VI, 250, pi. 26, figs. 6, 7.
Holotype: Service geologique de Tunisie (Tunis), no. C-Ce 13; car.
Type locality: Djebel Semmama (central N Tunisia), lat. 39°20'N, long. 7°15'E. Cretaceous (Cenomanian).
Figured specimens: British Museum (Nat. Hist.) specimen nos. OS 13454 (LV: PI. 17, 62, fig. 1), OS 13455 (car: PI.
17, 62, fig. 2), OS 13456 (RV: PI. 17, 62, fig. 3), OS 13457 (RV: PI. 17, 64, figs. 1, 3), OS 13458
(LV: PI. 17, 64, figs. 2, 4). From Cenomanian marls (Marnes dA.it Lamine) exposed by coast road
c. 20 km N of Agadir, Morocco. Sample taken 150 m stratigraphically above prominent Albian
limestones (Calcaires dolomitiques du Kechoula); lat. 30° 33' 30" N, long. 9° 44' 30" W.
Explanation of Plate 17, 62
Fig. 1, LV, ext. lat. (OS 13454, 700 /am long); fig. 2, car., dors. (OS 13455, 710 /u.m long); fig. 3, RV, ext. lat. (OS 13456, 680 /am long).
Scale A (200 /am; x90), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 63 Cytheropteron glintzboeckeli (3 of 4)
Diagnosis: Alate; angular flange extends from anterior around alar process and towards posterior; above
flange ornament has appearance of overlapping plates, beneath flange on flat ventral surface are a
series of sub-parallel ridges; both valves have single spine below caudal process; subrhombic
outline in dorsal view.
Remarks: The specimens illustrated herein have been compared to topotypes (W. Austin, 1988, unpublished
MSc thesis, UCW Aberystwyth) and are conspecific.
The micrographs reveal hitherto undescribed features of the ornament; an alar spine, a line of
small pustules along the alar flange and a very fine “string of pearls” polygonal reticulation over
much of the carapace. The hinge is antimerodont and the adductor muscle scars are arranged in a
column of four with a circular frontal scar about twice the diameter of the small mandibular scar.
Previous authors have placed this species in the genus Eocytheropteron Alexander, 1933.
However the original generic description of Eocytheropteron (C. A. Alexander, J. Paleont . , 7,
181-214, 1933) states that the genus is like Cytheropteron Sars, 1866, in most respects but that the
species within it “do not bear any trace of wing-like lateral expansions”. This species is therefore
more appropriately ascribed to the genus Cytheropteron.
Distribution: Previously recorded from Tunisia, NE Algeria (Viviere, 1985 op. cit.), and, as an unnamed
species, from S Morocco (Masoli, 1963 op. cit.). The stratigraphic range of this species in Morocco
is Vraconian (Late Albian) to Early Cenomanian (acme); this is consistent with previously
assigned stratigraphic ranges.
Explanation of Plate 17, 64
Fig. 1, RV, int. obi. vent. (OS 13457, 700 /am long); fig. 2, LV, int. muscle scars (OS 13458, photo 120 /am long); fig. 3, RV, int. hinge
(OS 13457, photo 210 /am long); fig. 4, LV, int. obi. vent. (OS 13458, 710 /am long).
Scale A (200/am; x 100), figs. 1, 4; scale B (50/am; x410), fig. 2; scale C (100/am; x210), fig. 3.
Stereo-Atlas of Ostracod Shells 17, 62
Cytheropteron glintzboeckeli (2 of 4)
Stereo-Atlas of Ostracod Shells 17 (12) 65-68 (1990) Loxocorniculum grateloupianum (1 of 4)
595.337.14 (118.22) (44 : 162.002.48) : 551 .351 + 552.51 + 552.52
ON LOXOCORNICULUM GRATELOUPIANUM (BOSQUET)
by Caroline A. Maybury
(Institute of Earth Studies, University College of Wales, Aberystwyth)
Loxocorniculum grateloupianum (Bosquet, 1852)
Cy there grateloupiana sp. nov. J. Bosquet, Mem. cour. Acad. r. Sci. Belg., 24, 81, pi. 4, fig. 3.
Loxoconcha grateloupiana (Bosquet); G. S. Brady, Trans, zool. Soc. Lond., 10, 399, pi. 68, figs. 3a-g.
Loxoconcha grateloupiana (Bosquet); W. N. Kuiper, Oligocane und Miocdne Ostracoden aus den Niederlanden,
Gebroeders Hoitsema, Groningen, 23, pi. 1, figs. 6a-c.
Loxoconcha grateloupiana (Bosquet); H. J. Oertli, Schweiz, palaeont. Abh., 74, 68, pi. 8, fig. 210.
Loxoconcha grateloupiana (Bosquet); A. J. Keij, Mem. Inst. r. Sci. nat. Belg., 136, 140, pi. 21, fig. 19; pi. 22, figs. 9-11.
Loxoconcha grateloupiana (Bosquet); J. Moyes, Les Ostracodes du Miocene Aquitain. Essai de paleoecologie
stratigraphique et de paleogeographie, Univ. Bordeaux (Drouillard Impr.), 70, pi. 7, fig. 1.
Sagmatocy there ? grateloupiana (Bosquet); P. Carbonel in: H. J. Oertli (Ed.), Atlas des Ostracodes de France, Bull.
Centres Rech. Explor.-Prod. Elf -Aquitaine, 9, 326, pi. 93, figs. 3-4.
Lectotype: Roy. Inst. Nat. Sci. Belg., Brussels, cf RV (Bosquet Coll., no. 44), designated Keij, op. cit.
Type locality : Leognan, Aquitaine Basin, France; Lower Miocene.
Figured specimens: British Museum (Nat. Hist.) nos. OS 13480 (9 LV : PI. 17, 66, fig. 1), OS 13481 (9 RV: PI. 17, 66. fig. 2). OS
13482 (Cf LV: PI. 17, 66, fig. 3), OS 13483 (cf RV: PI. 17, 68, fig. 1), OS 13484 (cf RV: PI. 17, 68, figs. 2-4).
All specimens are from Apigne, near Rennes (approx, lat. 48°07'N, long. 1°41'W), NW France; Upper
Pliocene, Redonian. OS 13480 and OS 13481 from the grey, marine sands of Apigne Borehole II; OS 13482
and OS 13484 from the shell-rich sand of the cutting at Le Temple du Cerisier and OS 13483 from the yellow,
argillaceous, calcareous sands with high clay content of Gite d’Apigne (see J.-P. Margerel, Les Foraminiferes
du Redonien. Systematique, Repartition stratigraphique, Paleoecologie, Nantes, 1, 8-26, 1968).
Explanation of Plate 17, 66
Fig. 1, 9 LV, ext. lat. (OS 13480, 620 p.m long); fig. 2, 9 RV, ext. lat. (OS 13481, 630 /i.m long); fig. 3, cf LV, ext. lat. (OS 13482,
710 yum long).
Scale A (100 /um; x 100), figs. 1-3.
Stereo-Atlas of Ostracod Shells 17, 67 Loxocorniculum grateloupianum (3 of 4)
Diagnosis: A medium to large (adults 600-750 pm long), coarsely reticulate Loxocorniculum, with a rounded
posteroventral protuberance and just posterior and immediately below this, a smooth, deeply depressed area.
Hinge modified gongylodont with a crenulate median element. Adductor muscle scars a curved row of 4.
Remarks: Oertli (pers. comm.) has compared the present material with specimens of grateloupianum from the type
region and has confirmed the two are the same. The generic assignment of Bosquet’s species, however, has
proved problematical. Its gongylodont ( s.l .) hinge undoubtedly led Oertli (1956, op. cit.), Keij (1957, op. cit.)
and Moyes (1965, op. cit.) to refer the species to Loxoconcha, but other authors, notably Bassiouni
(Roemeriana, 3, 65, 1962), Neale ( Scient . Rep. Br. Antarct. Surv., 58, 19, 1967) and Carbonel in Oertli (1985,
op. cit.) have assigned it to Kuiperiana, Myrena and Sagmatocytherel respectively because of the somewhat
aberrant nature of the gongylodont hinge and the fact that the species’ ornament and outline are not typical of
Loxoconcha. I, however, regard Myrena (Neale, 1967, op. cit.) as a junior synonym of Kuiperiana Bassiouni
(1962, op. cit.), a genus that is chartacterised by a gongylodont hinge with a smooth to weakly crenulate
median element and a posterior curved tooth in the RV with a frill -like dorsal surface. Furthermore, it is not
like Sagmatocythere Athersuch ( Stereo-Atlas of Ostracod Shells, 3, 117-124, 1976) because the latter has a
gongylodont hinge with a smooth median element and a pronounced posterior curved tooth, which are
lacking in grateloupianum (see PI. 17, 68, figs. 2-4). I have, instead, assigned the species to Loxocorniculum
Benson & Coleman ( Paleont . Contr. Univ. Kans., 31, 38, 1963) on account of its lateral outline, ornament
and hinge structure. Although grateloupianum does not possess the posterodorsal protuberance of typical
Loxocorniculum, this may be observed in a closely related species, L. micrograteloupianum Maybury
(Stereo-Atlas of Ostracod Shells, 17, 69-72, 1990). In addition, the type species of Loxocorniculum, Cythere
fischeri Brady (in L. De Folin & L. Perier (Eds.), Les Fonds de la Mer, Savy, Paris, 4(1), 154, pi. 18, figs.
15-16, 1869), has a pronounced, posteroventral protuberance adjacent and dorsal to a broad posteroventral
depression, which is present in grateloupianum.
Brady (1878, op. cit.) and Kuiper’s (1918, op. cit.) material appears to consist of several discrete species
none of which is conspecific with Bosquet’s taxon.
Distribution: Middle Oligocene (Rupelian) and Miocene (Aquitanian-Helvetian) of France (Keij, 1957, op. cit.-, Oertli,
1956, op. cit. and Moyes, 1965, op. cit.). I have found it at several Upper Pliocene (Redonian) localities in
NW France: Palluau I and Apigne II boreholes, Gite d’Apigne and Le Temple du Cerisier (herein).
Explanation of Plate 17, 68
Fig. 1, cf RV, ext. lat. (OS 13483, 680 pm long); figs. 2-4, cf RV (OS 13484, 660 gm long): fig. 2, int. lat.; fig. 3, ant. hinge element;
fig. 4, post, hinge element.
Scale A (100/j.m; x 100), figs. 1, 2; scale B (40/u.m; x263), figs. 3, 4.
1852
non 1878
non 1918
1956
1957
1965
1985
Stereo-Atlas of Ostracod Shells 17, 66
Loxocorniculum grateloupianum (2 of 4)
Stereo-Atlas of Ostracod Shells 17, 68
Loxocorniculum grateloupianum (4 of 4)
Stereo-Atlas of Ostracod Shells 17 (13) 69-72 (1990) Loxocorniculum micrograteloupianum (1 of 4)
595.337.14 (118.22) (44 : 162.002.48) : 551.351 + 552.51
ON LOXOCORNICULUM MICROGRATELOUPIANUM MAYBURY sp. nov.
by Caroline A. Maybury
( Institute of Earth Studies, University College of Wales, Aberystwyth)
Loxocorniculum micrograteloupianum sp. nov.
Holotype: British Museum (Nat. Hist.) no. OS 13485, 9 LV.
[Paratypes: British Museum (Nat. Hist.) nos. OS 13486-13489.
Type locality:
Derivation of name:
Figured specimens:
Fine glauconitic, grey sand between 26.7-32.5 m, Apigne (Borehole II), SW of Rennes (approx,
lat. 48°07'N, long. 14°41'W), NW France; Upper Pliocene, Redonian.
With reference to the fact that, while resembling Loxocorniculum grateloupianum (Bosquet, 1852)
(see C. A. Maybury, Stereo-Atlas of Ostracod Shells, 17, 65-68, 1990), the present species is
notably smaller.
British Museum (Nat. Hist.) nos. OS 13485 (holotype, 9 LV: PI. 17, 70, fig. 1), OS 13486
(paratype, 9 LV: PI. 17, 70, fig. 2), OS 13487 (paratype, cf LV: PI. 17, 70, fig. 3), OS 13488
(paratype, cf RV: PI. 17, 72, fig. 1), OS 13489 (paratype, 9 RV: PI. 17, 72, figs. 2, 3, 5).
Specimens OS 13489 and OS 13488 from the type locality and horizon; specimens OS 13486 and OS
13487 from the shell-rich sand of Le Temple du Cerisier, SW of Rennes, NW France; Upper
Pliocene, Redonian (see J.-P. Margerel, Les Foraminiferes du Redonien. Systematique, Repartition
stratigraphique, Paleoecologie, Nantes, 1, 8-26, 1968 for further sample details).
Explanation of Plate 17, 70
Fig. 1, 9 LV, ext. lat. (holotype, OS 13485, 550 gm long); fig. 2, 9 RV, ext. lat. (paratype, OS 13486, 530 gm long); fig. 3, cf LV, ext.
lat. (paratype, OS 13487, 600 p.m long).
Scale A (200/u.m; x 109), figs. 1-3.
Stereo- Atlas of Ostracod Shells 17, 71
Loxocorniculum micrograteloupianum (3 of 4)
Diagnosis: A medium-sized (adults 520-600 gm long), dimorphic, alate Loxocorniculum with a rounded
anterior margin and asymmetrically rounded posterior, possessing a subdorsal caudal process
which is best developed in cf LV. The dorsal margin is straight to convex in females, oblique in cf
LV and slightly concave medianly in cf RV. Ventral margin curved. Posterodorsal protuberance
best developed in males. Ornament very coarsely reticulate with horizontal muri dominant and
costate. Posteroventral incision well developed and smooth; eye tubercle small and ridge-like.
Hinge similar to that of L. grateloupianum (see Maybury, 1990, op. cit.). Adductor muscle scars an
oblique row of 4 with a ‘c’-shaped frontal scar open dorsally, a narrow crescentic fulcral point and
1 (perhaps 2?) elongate mandibular scars.
Remarks: L. micrograteloupianum resembles L. grateloupianum in its coarse reticulation, smooth,
pronounced posteroventral incisure and similar hinge structure (see Maybury, 1990, op. cit.). The
two species differ in that L. micrograteloupianum possesses a posterodorsal protuberance,
ridge-like, horizontal muri which are almost costate and a ridge-like eye tubercle and whereas, in
L. grateluopianum the adductor muscle scars form a curved row, in L. micrograteloupianum they
form an oblique row. The size difference between the new species and L. grategloupianum has
already been emphasized.
Distribution: L. micrograteloupianum has been recovered from the Upper Pliocene, Redonian deposits of
Apigne (Borehole II, Le Temple du Cerisier), Beugnon (Sample No. 3) and L’Aubier ; NW France
(see Margerel (1968, op. cit.) for geographical, stratigraphical and sample details).
Explanation of Plate 17, 72
Fig. 1, 9 RV, ext. lat. (paratype, OS 13488, 590 pm long); figs. 2-4, 9 RV (paratype, OS 13489, 560 pm long): fig. 2, int. lat.; fig. 3,
ant. hinge element; fig. 4, post, hinge element.
Scale A (200/i.m; x 109), figs. 1, 2; scale B (40/u.m; x273), figs. 3, 4.
Stereo-Atlas of Ostracod Shells 17, 70
Loxocorniculum micrograteloupianum (2 of 4)
Loxocorniculum micrograteloupianum (4 of 4)
Stereo- Atlas of Ostracod Shells 17, 72
Stereo-Atlas of Ostracod Shells 17 (14) 73-76 (1990)
595.336.13 (113.312) (766 : 162.097.34) : 551.351 + 552.54
Bromidella papillata (1 of 4)
ON BROMIDELLA PAPILLATA (HARRIS)
by C. Giles Miller, Mark Williams & Matthew I. Wakefield
(University of Leicester, England)
1957
Eurychilina papillata n. sp. R. W. Harris, Bull. Okla geol. Surv., 55, 232, pi. 7, figs. 9, lOa-b.
Museum of Compatative Zoology, Harvard University (MCZ), U.S.A., no. 4626; male carapace.
From zone 3 (see Harris ,1957 ' op. cit.). Bromide Formation, Simpson Group, middle Ordovician,
laldeTs"' !N.eCgimde'9760; WS-’ RIE'’ HU'S’ °klah°ma' U S A';
Harvafd Museum of CDmparatire Zoology, U.S. A. nos. MCZ 4626; (holotype, cf car. : PI. 17 76
,gs. 1-3) MCZ 4626A; (? RV: PI. 17, 74. figs. 1-4, PI. 17, 76, fig. 4). Both figured spec, mens are
from the type locality in the Bromide Formation.
Weakly bisulcate; SI shallow and directed towards the anterocentral margin, S2 well defined and
deepest slightly ventral of pre-adductorial node. Velum an entire well developed flange in both
dimorphs, surmounted by closely spaced small tubercles. Both valves with well developed
tuberculate admarginal ridge, absent in area of dolon in heteromorph.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Figs. 1-5, $ RV
Scale A (200 /um;
Explanation of Plate 17, 74
(MCZ 4626 A, 1.7mm long); fig. 1, ext. lat.; fig. 2, ext. lat. obi.; fig 3
x 35), figs. 1-4. S' ’
ext. vent.; fig. 4, int. lat.
Stereo-Atlas of Ostracod Shells 17, 75
Bromidella papillata (3 of 4)
Remarks:
Distribution :
Acknowledgements :
Eurychilina ^ ^ dlm°rphic structures of B • papillata clearly distinguish it from
B- papillata most closely resembles Bromidella mattea (Kraft) (see Mem. geol Socl Am 86
pi. 10, figs. 9-14, 1962) particularly with respect to heteromorph dolonal morphology. B. papillata
appears to differ from B. mattea by having a less well developed preadductorial node and by
u1 jg.the ve um surmount the dolon, whereas in the latter species the velum is expanded to form
the dolon.
.. The heteromorph dolon of B. papillata is also very similar to that of B. reticulata (see
M. Williams & D. J Siveter, Stereo-Atlas Ostracod Shells , 16, 1, 1989), but the heteromorph
shows no evidence of the histial structure developed anteriorly in B. reticulata.
dorsal of the adductural sulcus can be observed in Harris’ paratype specimen
(MCA 4626A), this feature has not been observed in any other conspecific tecnomorph or
heteromorph specimens.
B papillata occurs in the deeper water mid-shelf facies of the Bromide Formation (see
M. Longman, Univ. Kans. paleont. Contr., Monograph 1, 1982).
B. papillata is known from the Pooleville and Mountain Lake members of the Bromide Formation
middle Ordovician, Arbuckle Mountains and Criner Hills, Oklahoma, U.S. A.
Dr D. J. Siveter (University of Leicester) for helpful discussion, and Dr J. Berdan (Smithsonian
Institution, Washington) for loan of the types.
„ , Explanation of Plate 17, 76
Figs. 1-3, cf car. (holotype MCZ 4626, 1.6mm long); fig. 1, ext. lat. ; fig. 2, ext. lat. obi
detail of heteromorph dolon (MCZ 4626A, 1.7 mm long).
Scale A (200 ^.m; x37), figs. 1-3; scale B (200 /tm; x90), fig. 4.
; fig. 3, vent. Fig. 4, $ RV,
anterior close-
up
Stereo-Atlas of Ostracod Shells 17, 74
Bromidella papillata (2 of 4)
Stereo-Atlas of Ostracod Shells 17, 76
Bromidella papillata (4 of 4)
BPCC Blackpool
COLOUR PRINTERS
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Stereo-Atlas of Ostracod Shells: Vol. 17, Part 1
CONTENTS
On Welchella foveata Dewey & Puckett gen. et sp. nov.; by C.P. Dewey & T.M. Puckett.
On Glyptopleura henbesti Croneis & Gutke; by C.P. Dewey.
On Winchellatia longispina Kay; by M. Williams.
On Eridoconcha simpsoni Harris; by M. Willliams & P.J. Jones.
On Cypridea unicostata Galeeva chinensis Neale & Su subsp. nov.; by J.W. Neale &
Su Deying.
On Sunliavia tumida Sou; by Su Deying & J.W. Neale.
On Theriosynoecum conopium Wakefield & Athersuch sp. nov.; by M.I. Wakefield &
J. Athersuch.
On Darwinula incun’a Bate; by M.I. Wakefield.
On Londinia kiesowi (Krause); by W. Hansch & D.J. Siveter.
On Hemsiella niaccoyiana (Jones); by D.J. Siveter & W. Hansch.
On Cytheropteron glintzboeckeli (Donze & Lefevre); by R. Symonds.
On Loxocomiculum grateloupiamim (Bosquet); by CA.. Maybury.
On Loxocomiculum micrograteloupianum sp. nov.; by C.A. Maybury.
On Bromidella papillata (Harris); by C.G. Miller, M. Williams & M.I. Wakefield.
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