A Stereo-Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter,
and J. E. Whittaker
A fa
Published under the aegis of the British Micropalaeontological Society, London
ISSN 0952-7451
Editors
Dr J. Athersuch, Exploration Technology Branch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Dr D.J. Horne, School of Earth Sciences, Thames Polytechnic, Walburgh House, Bigland Street,
London El 2NG.
Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.
Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.
Editorial Board
Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.
Dr P. de Deckker, Department of Geology, Australian National University, G.P.O. Box 4, Canberra,
ACT 2601, Australia.
Dr W. Hansch, Ernst-Moritz-Arndt Universitat, Sektion Geologische Wissenschaften, F.L.-Jahnstr. 17a,
2200 Greifswald, Germany.
Prof. R. Lundin, Department of Geology, Arizona State University, Tempe, Arizona 85287-1404, U.S.A.
Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55,
D 2000 Hamburg 13, Germany.
Prof. N. Ikeya, Institute of Geosciences, Shizuoka University, Shizuoka 422, Japan.
Officers of the British Micropalaeontological Society
Chairman Professor M.B. Hart, Department of Geological Sciences, Polytechnic South West, Drake Circus,
Plymouth, Devon PL4 8AA.
Secretary Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.
Assistant Treasurer Dr I.P. Wilkinson, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Journal Editor Dr M.C. Keen, Department of Geology, The University, Glasgow G12 8QQ.
Newsletter Editor Dr D.J. Shipp, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Conodont Group Chairman Dr H.A. Armstrong, Department of Geological Sciences, University of Durham,
Science Laboratories, South Road, Durham DH1 3LE.
Conodont Group Secretary Mr M.T. Dean, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Foraminifera Group Chairman Dr H.W. Bailey, Paleo Services Ltd., Unit 15, Paramount Industrial Estate,
Sandown Road, Watford WD2 4XA.
Foraminifera Group Secretary Dr F.M.D. Lowry, School of Earth Sciences, Thames Polytechnic,
Walburgh House, Bigland Street, London El 2NG.
Ostracod Group Chairman Dr J. Athersuch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Ostracod Group Secretary Dr I.D. Boomer, School of Environmental Sciences, University of East Anglia,
Norwich NR4 7TJ.
Palynology Group Chair Dr R.J. Davey, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Palynology Group Secretary Dr A. McNestry, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Calcareous Nannofossil Group Chairman Dr L.T. Gallagher, Paleo Services, Unit 15, Paramount Industrial
Estate, Sandown Road, Watford WD2 4XA.
Calcareous Nannofossil Group Secretary Dr N.M. Hine, British Geological Survey, Keyworth, Nottingham
NG12 5GG.
Instructions to Authors
Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. Format
should follow the style set by the papers in this issue. Descriptive matter apart from illustrations should be
cut to a minimum; preferably each plate should be accompanied by only one page of text. Blanks to aid in
mounting figures for plates may be obtained from any one of the Editors or Editorial Board. Completed
papers should be sent to one of the Editors. All contributions submitted for possible publication in the Stereo-
Atlas of Ostracod Shells are reviewed by an appropriate international specialist.
The front cover shows a male left valve and appendages, internal view, of Linmocythere borisi borisi Martens,
1990. Paratype, K.B.I.N., Brussels, OC.1406. From Lake Abijata, Ethiopia. Photographed by K. Martens and
J. Cillis.
Stereo-Atlas of Ostracod Shells 18 (1) 1-4 (1991) Orcofabella testata (1 of 4)
595.336.16 (113.333) (47 : 161.022.58) : 551.351 + 552.54
ON ORCOFABELLA TESTATA (GAILITE)
by David J. Siveter & Lembit Sarv
(University of Leicester, England & Institute of Geology, Tallinn, Estonia)
Genus ORCOFABELLA Gailite, 1967
Type-species (by original designation): Orcus testatus Gailite, 1966.
1966 Orcus gen. nov. L. Gailite, Palaeontology & Stratigraphy of the Baltic & Byelorussia, Mintis, Vilnius, 1, (6), 109.
1967 Orcofabella nom. nov. L. Gailite, Geol. For. Stockh., Forh., 89, 387 (pro Orcus Gailite, 1966, non Orcus Mulsant, 1850; nec Uljanin,
1870; nec Needham, 1897).
Diagnosis: Primitiopsacea with a curved, reticulate lateral valve surface and a prominent adductorial pit.
Reticulation generally coarse, is delimited laterally by a weak ridge, typically contains elongate fossae
dorsally adjacent to the adductorial pit. Velum occurs as a narrow, rounded ridge forming the valve
margin in lateral view, continuous posteriorly as the dolon in females. Dolon normally open; can be
ornamented with fine ridges.
Remarks: Clavofabella Martinsson, 1955, Primitiopsis Jones, 1887 and Limbinariella Sarv, 1968 are other Silurian
reticulate primitiopsaceans. As represented by the type-species, Orcofabella differs from Limbinariella
(see Siveter, D.J. & Sarv, L., Stereo-Atlas Ostracod Shells, 18, 5-8, 1991) in having a curved valve
surface, a more incurved and ornamented dolon and a different style of reticulation involving the
occurrence of elongate fossae adjacent to a more discretely demarcated adductorial pit; the nature of
the velum also differs between these genera. Orcofabella is distinguished from both Primitiopsis and
Clavofabella by its typically coarser ornament, which is demarcated laterally by a more or less
continuous ridge. Furthermore, the type-species of Primitiopsis has a closed dolon and a perimarginal
ridge. Perimarginal structures have not yet been documented from Orcofabella; none are discernable in
our material (PI. 18, 2, fig. 6).
Explanation of Plate 18, 2
Figs. 1-6, 9 RV (OS 5501, 1050/xm long): fig. 1, ant.; fig. 2, ext. lat.; fig. 3, post.; fig. 4, vent, obi.; fig. 5, vent.; fig. 6, int. lat.
Scale A (200 /u,m; x 48), figs. 1-6.
Stereo- Atlas of Ostracod Shells 18,3
Orcofabella testata (3 of 4)
Orcofabella testata (Gailite, 1966)
1966 Orcus testatus sp. nov. L. Gailite, Palaeontology & Stratigraphy of the Baltic & Byelorussia, Mintis, Vilnius, 1 (6), 110. pi. 2, figs. la-c.
1967 Orcus testatus Gailite; L. Gailite in L. Gailite et al., The Stratigraphy, fauna and conditions of formation of Silurian rocks in the central
part of the Baltic region, Zinatne, Riga, 104, pi. 3, figs. 5a-e.
1967 Orcofabella testata nom. nov.; L. Gailite, Geol For. Stockh. Forh., 89, 387.
1968 Orcofabella testata (Gailite); L. Sarv, Ostracode families Craspedobolbiniidae, Beyrichiidae and Primitiopsidae in the Silurian of
Estonia, Valgus, Tallinn, 76, pi. 26, figs. 1-4.
Holotype: Formerly in the All-Union Marine Scientific Producing Enterprise “Sojuzmorinzhgeologia”, Riga and is
now in the Nature Museum, Riga, Latvia, no. Os 31/121; 9 carapace.
Type locality : At 419m depth in the Piltene 1 borehole, W Latvia, Jura Formation; = Ohesaare regional ‘stage’,
Pndoli Series, Silurian.
Diagnosis: Orcofabella species with lateral reticulation coarse overall, contains several obliquely elongate fossae
above the adductorial pit, shows a few areas of sparsely developed second order reticulation. External
surface of dolon has anastomising ridges.
Figured specimens: Institute of Geology, Tallinn, Estonia, nos. Os 5501 ($ RV: PI. 18, 2, figs. 1-6), Os 5502 (9 RV: PL 18,
4, figs. 1-3), Os 5503 (tecnomorphic RV: PI. 18, 4, figs. 4—6). All originals of Sarv, 1968 (pi. 26, figs.
1-4; from cliff section near Ohesaare, Saaremaa Island, Estonia (lat. 58°12'N, long. 22°30'E); Ohesaare
(K4) regional ‘stage’, Pndoli Series, Silurian.
Remarks: Of the described Orcofabella species O. levireticulata Schallreuter (Mitt, geol.-palaont. Inst. Univ.
Hamburg , 61, 1986) is known from erratics (Pndoli age) in Germany and O. arguta Gailite, 1966, O.
araneosa (Gailite, 1966), O. obscura Sarv, 1968 and the type-species are recorded almost exclusively
from the East Baltic. O. arguta is closest to O. testata but differs most obviously in ornament.
Orcofabella also probably occurs in Podolia, USSR (A.F. Abushik, unpublished).
Distribution: Silurian. East Baltic. Overall stratigraphic range: Kaugatuma (K3b) and Ohesaare (K4) regional ‘stages’
Pndoli Series. Saaremaa Island, Estonia: cliff near Ohesaare (Sarv, 1968). Latvia: Piltene 1 (Gailite,
1967) and Piltene 31 and Kolka 4 boreholes (Gailite, L. in: Phanerozoic Stratigraphy of the East Baltic,
Zinatne, Riga, 1978). Lithuania: boreholes no. 89 and 112 (Sidaraviciene, N. in: Kaljo, D. & Klaaman,
E. (eds.). Theory & Practice of Ecostratigraphy , Valgus, Tallinn, 1986).
Explanation of Plate 18, 4
Figs. 1-3, 9 RV (Os 5502, 1100/xm long): fig. 1, ant; fig. 2, ext. lat.; fig. 3, vent. Figs. 4-6, tecnomorphic RV (Os 5503, 880 /xm long): fig. 4,
vent.: fig. 5, ext. lat.; fig. 6, post. Scale A (200 /^m; x46), figs. 1-3; scale B (200 /am; x56), figs. 4-6.
Orcofabella testata (2 of 4)
Stereo- Atlas of Ostracod Shells 18, 4
Stereo-Atlas of Ostracod Shells 18, 2
Orcofabella testata (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (2) 5-8 (1991) Limbinariella macroreticulata (1 of 4)
595.336.16 (113.333) (47 : 161.022.58) : 551.351 + 552.54
ON LIMBINARIELLA MACRORETICULATA SARV
by David J. Siveter & Lembit Sarv
( University of Leicester, England & Institute of Geology, Tallinn, Estonia)
Genus LIMBINARIELLA Sarv, 1968
Type-species (by original designation): Limbinariella macroreticulata Sarv, 1968.
Coarsely reticulate Primitiopsacea. Valve lateral surfaces flat, bounded by a narrow ridge. Adductorial sulcus
large, distinct, pit-like, connecting dorsally with a large fossa in the reticulation. Posterior dolon with open
antrum. In female right valves velar ridge occurs ventrally and is confluent posteroventrally with the dolon;
velum absent in tecnomorphs and ventrally in female left valves.
The left/right valve variation in the development of the velum of Limbinariella (see ‘Diagnosis’) has been
confirmed in topotype material of both Estonian members of the genus, the type-species and L. malornata
Sarv, 1968. A presence/absence of the (non-dolonal part of the) velum between right and left valves is a
feature also known from another Baltic primitiopsacean, Venzavella costata (Neckaja, 1960) (see Siveter, D.
J. & Sarv, L, Stereo-Atlas Ostracod Shells, 18, 9-12, 1991), but in that type-species the phenonemon
characterises both males and females. The dolonal morphology and flat valve surface bordered by a ridge is
also common to both genera; thus, in many essential respects they are similar and may be more closely related
than previously supposed.
Limbinariella differs from Venzavella in ornament and morphology of the adductorial sulcus.
Limbinariella differs from Limbinaria Swartz (in Swartz, F. M. & Whitmore, F. C., J. Paleont., 30, 1054,
1956), from the Silurian of the U.S.A., in valve profile and dolonal morphology.
Preparation of the antral region of the material of L. macroreticulata is extremely difficult. So far no
perimarginal structures have been observed.
Explanation of Plate 18, 6
Figs. 1, 2, 9 RV (OS 13705, 700 /xm long): fig. 1, ext. lat. ; fig. 2, vent. Figs. 3-5, cf RV (OS 13708, 625 /xm long): fig. 3, ext. lat. ; fig. 4. vent.; fig. 5,
post. Figs. 6-8, $ LV (OS 13706, 700/xm long): fig. 6, post.; fig. 7, ext. lat.; fig. 8. vent.
Scale A (200^m; x67), figs. 1, 2, 6-8; scale B (200/am; x73), figs. 3-5.
Stereo-Atlas of Ostracod Shells 18, 7 Limbinariella macroreticulata (3 of 4)
Diagnosis:
Remarks:
Limbinariella macroreticulata Sarv, 1968
1968 Limbinariella macroreticulata gen. et sp. nov. L. Sarv, Ostracode families Craspedobolbinidae, Beyrichiidae and Primitiopsidae in the Silurian of
Estonia , Valgus, Tallinn, 71, pi. 15, figs. 1-4.
Holotype:
Type locality:
Ligured specimens:
Remarks:
Distribution:
Institute of Geology, Academy of Sciences, Tallinn, Estonia, no. OS 5541; $ left valve.
Unimae, 6km N of Kuressaare (formerly known as Kingissepp), Saaremaa Island, Estonia. Paadla regional
‘stage’ (K^), Ludlow Series, Silurian.
British Museum (Nat. Hist.), nos. OS 13705 ($ RV: PI. 18, 6, figs. 1, 2), OS 13708 (cf RV: PI. 18, 6, figs.
3-5), OS 13706 ($ LV: PI. 18, 6, figs. 6-8), OS 13709 (9 RV: PI. 18, 8, figs. 1-3),
OS 13707 (cf LV: PI. 18, 8, figs. 4-6), OS 13710 (tecnomorphic LV: PI. 18, 8, figs. 7, 8). All topotype
(approximately lat. 58°12'N; long. 22°30'E); collected Sarv, 1959.
L. macroreticulata differs from the upper Ludlow L. malornata mainly by its coarser ornament. Limbinariella
semiplicata Schallreuter (Mitt, geol.-palaont. Inst. LJniv. Hamburg , 61, 204, 1986), from erratic boulders
(Pndoli Series, Isle of Sylt, Germany), differs in valve shape and in the size of its dolon and adductorial
sulcus.
Limbinariella cf. macroreticulata has been recorded from the Isakovsky beds of Podolia, USSR, which
are correlatives of the Kuressaare regional ‘stage’ of Estonia (Abushik, A. F. et al ., Lethaia, 18, 139, 143,
1985; Koren, T. N. et al., in Holland, C. H. & Bassett, M. G., A global standard for the Silurian System, Nat.
Mus. Wales Geol. Ser. no. 9, Cardiff, 1989).
Silurian. East Baltic and possibly Podolia, USSR. Overall stratigraphic range: uppermost part of Paadla (K2)
and Kuressaare (K3a) regional ‘stages’, Ludlow Series. Saaremaa Island, Estonia: the type locality (Unimae)
and locality Kuressaare (Sarv, 1968); and the Kaugatuma (unpublished information) and Ohesaare boreholes
(Sarv, 1968 and Eesti NSV Tead. Akad. Toim., (Keemia, Geol.), 20, 1971). Latvia: Kolka 54 borehole (Sarv,
L. in: Kaljo, D. (Ed.), Lacies & Launa of the Baltic Silurian, Acad. Sci. Estonian S.S.R. Tallinn, 1977).
Lithuania: Virbalis and Kunkojai boreholes (Sarv, 1977).
Explanation of Plate 18, 8
Figs. 1-3, $ RV (OS 13709, 700/u.m long): fig. 1, ext. lat. ; fig. 2, vent. ; fig. 3, obi. vent. Figs 4-6, cf LV (OS 13707, 680 /xm long): fig. 4, obi. vent. ; fig.
5, ext. lat.; fig. 6, vent. Figs. 7, 8, tecnomorphic LV (OS 13710, 580/am long): fig. 7, ext. lat.; fig. 8, vent.
Scale A (200/am; x65), figs. 1-6; scale B (100/am; x70), figs. 7, 8.
Stereo- Atlas of Ostracod Shells 18, 6
Limbinariella macroreticulata (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 8
Limbinariella macroreticulata (4 of 4)
\
Venzavella costata ( 1 of 4)
Stereo-Atlas of Ostracod Shells 18 (3) 9-12 (1991)
595.336.16 (113.333) (47 : 161.022.58) : 551.351 + 552.54
ON VENZAVELLA COSTATA (NECKAJA)
by David J. Siveter & Lembit Sarv
(University of Leicester, England & Institute of Geology, Tallinn, Estonia)
Genus VENZAVELLA Gailite, 1967
Type-species (by original designation): Limbinaria costata Neckaja, 1960
Diagnosis: Primitiopsacea with a velar ridge in females and tecnomorphs. Females have posterior dolon with open
antrum. Rounded adductorial pit. Ornament: 2-20 oblique ridges (striae). Perimarginal structure
(ridge) present. Right valve larger.
Remarks: A perimarginal ridge is known only in the type-species. Venzavella differs from Limbinariella Sarv. 1968
(see Siveter D. J. & Sarv, L., Stereo-Atlas Ostracod Shells, 18, 5-8, 1991) chiefly in ornament and
morphology of the adductorial sulcus. The general nature of the adventral structures and dimorphism is
essentially similar in both genera.
Five Venzavella species are known, principally from East Baltic sequences. Venzavella also occurs
in Baltic erratics. (Schallreuter, R. E. L., Mitt, geol.-paldont. Inst. Univ. Hamburg, 61, 1986). V.
germana Sarv, 1968 (lower Wenlock Jaani regional ‘stage’, Estonia) is the oldest species. V. multicostata
(Neckaja, 1960) [= V. loriei (Bonnena, 1910), see Schallreuter, 1986], V. subcostata Gailite, 1967 and
the type-species are all from the Pfidoh Series (Kaugatuma and Ohesaare regional ‘stages'). V. dicostata
(Gailite) ( Palaeontology & Stratigraphy of the Baltic and Byelorussia, Mintis, Vilnius, 1 (6), 1966) is
known only from the Ohesaare level.
Venzavella costata (Neckaja, 1960)
1960 Limbinaria costata Neckaja sp. nov. A. 1. Neckaja, in: Novye vidy drevnikh rastenii i bespozvonochnykh , Moscow, 2, 316,
pi. 61, figs. 7, 8.
1967 Venzavella costata (Neckaja); L. Gailite, in: L. Gailite etai, The Stratigraphy, fauna and conditions of formation of Silurian rocks in the
central part of the Baltic Region, Zinatne, Riga, 102, pi. 2, figs. 6a. b.
Explanation of Plate 18, 10
Figs. 1-4, 6, 2 LV (OS 13702, 1000 pm long): fig. 1, post.; fig. 2, ext. lat. ; fig. 3, vent.; fig. 4, int. lat. ; fig. 6, posterolat. obi. Fig. 5, RV ext.
lat. (holotype, 60/157, 980 p.m long). Fig. 7, 2 RV, ext. lat. (Os 5562, 1010/u.m long).
Scale A (200/u.m; x50), figs. 1-5, 7; scale B (200 /rm; X70), fig. 6.
Stereo-Atlas of Ostracod Shells 18, 11 Venzavella costata (3 of 4)
1968 Venzavella costata (Neckaja); L. Sarv, Ostracode families Craspedobolbinidae, Beyrichiidae and Primiliopsidae in the Silurian of
Estonia, Valgus, Tallinn, 78, pi. 28, figs. 3-14.
Holotype:
Type locality:
Diagnosis:
Figured specimens:
Remarks:
Distribution:
All-Union Petroleum Scientific Research Geological Institute (VNIGRI), Lenningrad, no. 60/157; cf
right valve.
Cliff near Kaugatuma, Saaremaa, Estonia; Kaugatuma regional ‘stage’ (K3b), Pffdolf Series, Silurian.
Species of Venzavella with 6-7 main ridges on each valve. Females have perimarginal ridge and
relatively wide dolon. Lobate area bounded by fine ridge weakly bipartite above hinge line. Velar ridge
along ventral part of valve in right valve of tecnomorphs and females, is typically absent in left valves.
Institute of Geology, Estonian Academy of Sciences, Tallinn, nos. Os 5562 (2 RV: PI. 18, 10,
fig. 7), Os 5563 (2 car.: PI. 18, 12, figs. 3, 4), Os 5569 (cf car.: PI. 18, 12, figs. 1, 2); originals Sarv, 1968.
VNIGRI, Lenningrad, no. 60/157 (holotype, cf RV: PI. 18, 10, fig. 5). British Museum (Nat. Hist.),
nos. OS 13702 (2 LV: PI. 18, 10, figs. 1-4, 6), OS 13703 (2 RV: PI. 18, 12, fig. 6), OS 13704
(tecnomorphic car.: PI. 18, 12, fig. 5). All topotypes; approximately lat. 58°12'N, long. 22°30'E.
The perimarginal ridge in females has only been seen with certainty in left valves (PI. 18, 10, figs. 4, 6).
The (ventral) velar ridge is normally found only in the right valve (PI. 18, 12, figs. 2, 3); however,
sometimes it appears also to be (relatively weakly) developed in left valves. V. costata differs from V.
germana chiefly in having a much wider dolon. V. costata is most easily distinguished from its
congeneric, coeval species by its number of ornamental ridges.
Silurian, East Baltic. Overall stratigraphic range: Kaugatuma (K3b) and Ohesaare (K4) regional ‘stages’,
Pndoli Series. Saaremaa, Estonia: type locality and localities Ohesaare, Aigu and Venekula (Sarv,
1968) and the Kaugatuma borehole (unpublished information). Latvia: Piltene 1 (Gailite, 1967), Piltene
32 (Sarv, L. in: Kaljo, D. (ed.). Facies & Fauna of the Baltic Silurian , Acad. Sci. Estonian S. S. R.,
Tallinn, 1977) and Kolka 4 (Gailite, L. in: Phanerozoic Stratigraphy of the East Baltic , Zinatne, Riga,
1978) boreholes. Lithuania: Virbalis borehole (Gailite, 1967) and boreholes nos. 87, 89 and 98
(Sidaraviciene, N. in: Kaljo. D., & Klaaman, E. (eds.), Theory & Practice of Ecostratigraphv, Valgus,
Tallinn, 1986).
Explanation of Plate 18, 12
Figs. 1, 2, cf car. (Os 5569, 970 pm long): fig. 1, ext. lat. ; fig. 2, vent. Figs. 3, 4, 2 car. (Os 5563, 960 /urn long); fig. 3, vent.; fig. 4, post. Fig. 5,
tecnormorphic car., ext. lat. (OS 13704, 900 /xm long). Fig. 6, 2 RV, ext. lat. (OS 13703, 1050 /um long).
Scale A (200 /xm; x50), figs. 1-6.
Stereo-Atlas of Ostracod Shells 18, 10
Venzavella costata (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (4) 13-16 (1991) Lomatopisthia simplex (1 of 4)
595.33 (113.312) (766 : 162.097.34) : 551 .351 + 552.52
ON LOMATOPISTHIA SIMPLEX (HARRIS)
by Mark Williams
(University of Leicester, England)
Genus LOMATOPISTHIA Guber & Jaanusson, 1964
Type-species (by original designation): Thomasatia simplex Harris, 1957.
Diagnosis: Small, quadrilobate Lomatopisthidae. Lobes confluent ventrally with connecting lobe. LI and L4
reaching the dorsal margin. L3 a broad lobe not reaching the dorsal margin. L2 often as a discrete node.
L4 continued posteriorly as a ridge, or overhanging the posterior margin. Lobate area bordered by
extralobal area thickened into a marginal ridge. Well developed inner lamella. Heteromorphic carapace
inflated posterior to S2. (Modified from Guber & Jaanusson, 1964).
Discussion: In their diagnosis for the new Family Lomatopisthidae (Superfamily and Suborder uncertain) and
new genus Lomatopisthia Guber & Jaanusson (1964) did not refer to the presence of an inner
lamella. This is present in all species they referred to Lomatopisthia, and also appears to be
present in the lomatopisthid genus Raymondatia Kay, 1934, based on published photographs
(Moore et al. 1961, Treatise on Invertebrate Paleontology, fig. 72. 3e). Other new genera referred to
the Lomatopisthidae by Guber & Jaanusson in 1964 include Bolbopisthia, Dibolbopisthia. and
Phyladopisthia. These appear to lack an inner lamella. Schallreuter (1978) referred his new genus
Europisthia to the Lomatopisthidae, but did not mention the presence of an inner lamella. The
Family Lomatopisthidae may require revision based on the presence or absence of the inner
lamella.
Explanation of Plate 18. 14
Fig. 1 , cf LV, ext. lat. (paratype, MCZ 4641b, 0.60mm long). Figs 2, 5, cf RV (OS 13500. 0.56mm long); fig. 2, ext. lat. ; fig. 5, int. lat. Fig. 3.
Cf RV, vent. (OS 13497, 0.51mm long). Fig. 4, cf LV, vent. (OS 13499, 0.56mm long).
Scale A (100/u.m; x87), figs. 1, 2, 5; scale B (100/u.m; x 102), fig. 3; scale C (100/u.m; 95), fig. 4.
Stereo-Atlas of Ostracod Shells 18, 15
Lomatopisthia simplex (3 of 4)
1957
1964
1982
Type locality:
Figured specimens:
Lomatopisthia simplex (Harris, 1957)
Thomasatia simplex n. sp., R. W. Harris, Bull. Okla. geol. Surv., 75, 245, pi. 8, figs. 15, 17a, b.
Lomatopisthia simplex (Harris); A. L. Guber & V. Jaanusson, Bull. geol. Instn Univ. Uppsala , 43, 27, pi. 3, figs. 5-15. text-fig. 12.
Lomatopisthia simplex (Harris); M. J. Copeland, Bull. geol. Surv. Can.. 347, 10, pi. 8, fig. 5.
Holotype: Museum of Comparative Zoology, Harvard University, U.S.A., no. 4641; a tecnomorphic left valve.
Decker’s Bed 36 (see Harris, 1957), Bromide Formation, Simpson Group. Middle Ordovician: Highway
99 Section, Oklahoma, U.S.A.; approximate latitude 34°35'N, longitude 96°41'W.
Museum of Comparative Zoology, Harvard University, nos. 4641 (holotype, 9 LV: PI. 18. 16, figs.
1-3). 4641b (paratype, cf LV: PL 18, 14, fig. 1). British Museum (Nat. Hist.), London, nos. OS 13500
(Cf RV: PI. 18, 14, figs. 2, 5), OS 13497 (cf LV: PI. 18, 14, fig. 3). OS 13499 (cf RV: PI. 18, 14. fig. 4).
OS 13498 ($ RV: PI. 18, 16, fig. 4). Holotype and paratype specimens from the type locality and
horizon. All other specimens from a single sample in the Mountain Lake Member of the Bromide
Formation at the type locality. Sample provided by Mr. A. Grafham, Geological Enterprises, Ardmore.
Oklahoma, U.S.A.
Lomatopisthia species having rounded lobes with L3 especially broad. Tecnomorph with deep crescentic
S3. Posterior part of heteromorph shows domiciliar inflation and much reduced S3, with L3 bordered by
a distinct dorsal furrow.
L. simplex has the weakest lobation of all Lomatopisthia species except L. varicata (Harris, 1957). It is
most similar to L. rectantulata (Kraft, 1962), differing only by lacking a furrow between L3 and the
connecting lobe, and by having the anterior margin of L2 clearly separate from LI. L. auricula (Harris.
1957) has much stronger lobation (see Williams, M., Stereo-Atlas Ostracod Shells. 18. 17-20. 1991).
A species occuring in open marine sediments. Found in the Middle Ordovician Tulip Creek and
Bromide formations of Oklahoma, and the Middle Ordovician Day Point Formation of New York.
U.S.A. (Copeland, 1982).
Dr David J. Siveter and Mr Matthew Wakefield (University of Leicester) for useful discussion.
Diagnosis:
Discussion :
Occurrence:
Acknowledgement :
Explanation of Plate 18. 16
Figs. 1-3, 9 LV, (holotype, MCZ 4641, 0.54mm long), fig. 1, ext. lat.; fig. 2, ext. lat. obi.: fig. 3, vent. Fig. 4, 9 RV. ext. lat. (OS 13498.
0.56mm long). Scale A (100/um; x85), figs. 1-4.
Stereo-Atlas of Ostracod Shells 18, 16
Lomatopisthia simplex (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 14
Lomatopisthia simplex (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (5) 17-20 ( 1991)
595.33 (113.312) (766: 162.098.37) 551.351+552.54
Lomatopisthia auricula ( 1 of 4)
ON LOMATOPISTHIA AURICULA (HARRIS)
by Mark Williams
(University of Leicester, England)
Lomatopisthia auricula (Harris, 1957)
1957 Thomasatia auricula n. sp. R. W. Harris, Bull. Okla. geol. Surv., 75, 246. pi. 8, figs. 11a, b.
1964 Lomatopisthia auricula (Harris); A. L. Guber & V. Jaanusson, Bull. geol. Instn Univ. Uppsala , 43, 26.
Holotype: Museum of Comparative Zoology, Harvard University, U.S.A., no. 4639; a heteromorphic
carapace.
Type locality: Decker’s bed 3 (see Harris, 1957), Bromide Formation, Simpson Group, Middle Ordovician;
Rock Crossing Section, Criner Hills, Oklahoma, U.S.A.; approximately latitude 37°08'N,
longitude 97°10'W.
Figured specimens: Museum of Comparative Zoology, Harvard University, U.S.A., no. 4639 ($ car.: PI. 18, 18, figs.
1-3). British Museum (Natural History), London, nos. OS 13496 (9 car.: PI. 18, 18, fig. 4). OS
13493 (cf RV: PI. 18, 20, figs. 1, 2), OS 13494 ( 9 RV; PI. 18, 20, fig. 3), OS 13495 (9 LV: PI. 18,
20, fig. 4). Holotype from the type locality and horizon. OS 13494 - 13496 from the Pooleville
Member of the Bromide Formation at the type locality. OS 13493 from a sample in the Mountain
Lake Member of the Bromide Formation, Highway 99 Section, Arbuckle Mountains, Oklahoma.
Diagnosis: Lomatopisthia species with well developed lobation. Heteromorph with a pronounced extralobal
ridge (showing a distinct midventral bend) and two egg-shaped inflations posterior to L4.
Explanation of Plate 18, 18
Figs. 1-3, 9 car. (holotype, MCZ 4639, 0.63mm long): fig. 1. RV, ext. lat. ; fig. 2, RV, ext. lat. obi.: fig. 3, LV, ext. lat. Fig. 4, 9 car.,
vent. (OS 13496, 0.60mm long).
Scale A (100/xm; x85), figs. 1-3; scale B (100/u.m: x87), fig. 4.
Stereo-Atlas of Ostracod Shells 18, 19 Lomatopisthia auricula (3 of 4)
Discussion: An inner lamella is well developed in L. auricula , being wide anteriorly and narrowing rapidly
posteriorly. The same structure has been identified in all congeneric species from Oklahoma,
many of which were described by Harris (1957) under other genera.
L. auricula is markedly dimorphic; L3 is very wide in the tecnomorph, but is considerably
narrower in the heteromorph because of the posterior domiciliar inflation of the carapace. S3 is
strongly developed and crescentic in both dimorphs, whereas in L. simplex (Harris, 1957) it tends
to be considerably reduced or absent in the heteromorph. The marked midventral bend in the
extralobal ridge of the heteromorph of L. auricula is apparently not present in the tecnomorph
(here the extralobal ridge is more weakly developed).
The two-egg shaped inflations posterior of L4 in the heteromorph of L. auricula compare
closely with features in the presumed heteromorph of Saturnites harrisi Levinson, 1961
(Micropaleontology, 7, 362, pi. 1, fig. 6), the type-species for Saturnites Levinson, 1961. The
internal features of the latter species are unknown but the genus may be closely related to
Lomatopisthia.
Assemblages of L. auricula in the Bromide Formation are very much dominated by
heteromorphs. L. auricula has the most strongly developed lobation of all Lomatopisthia species;
for example, see the type-species, L. simplex (Harris, 1957), in Williams, M., Stereo-Atlas
Ostracod Shells, 18, 13-16, 1991.
Distribution: A species occurring in open marine sediments. Only known from the Mountain Lake and
Pooleville members, Bromide Formation, Middle Ordovician Simpson Group of Oklahoma,
U.S.A.
Acknowledgement: Dr. David J. Siveter and Mr. Matthew Wakefield (University of Leicester) for useful discussion.
Explanation of Plate 18, 20
Figs. 1, 2, cf RV (OS 13493, 0.59mm long): fig. 1, ext. lat.; fig. 2, ext. lat. obi. Fig. 3, 9 RV, int. lat. (OS 13494, 0.63mm long). Fig. 4,
9 LV, int. lat. (OS 13495, 0.63mm long).
Scale A (100/j.m; x93), figs. 1, 2; scale B (100/xm; x84), figs. 3, 4.
Stereo-Atlas of Ostracod Shells 18, 18 Lomatopisthia auricula (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (6) 21-24 (1991) Cleithranchiste paulusi (1 of 4)
595.337.23 (113.44) (430 : 161.006.50) : 551.351 + 552.54
ON CLEITHRANCHISTE PAULUSI BECKER
by Gerhard Becker
(University of Frankfurt, Germany )
Genus CLEITHRANCHISTE Becker, 1965
Type-species (by original designation): Cleithranchiste paulusi Becker, 1965.
Diagnosis: Thick-shelled, large, smooth, asymmetrical (left valve distinctly larger than right valve) thlipsurid
genus. Elliptical outline to the carapace; posterior spine(s) only on the right valve. Hinge
hemisolen to tripartite, muscle-scar pattern a healdiid cluster of about 30 scars. No marginal
structures.
Distribution: Central and possibly W Europe; Middle Devonian; Eifelian and possibly the Givetian.
Cleithranchiste paulusi Becker, 1965
1965 Cleithranchiste paulusi sp. nov. G. Becker, Senckenberg. leth., 46, 371, 372, pi. 28, fig. 1, pi. 29, figs. 5, 6.
1969 Cleithranchiste paulusi Becker; G. Becker, Senckenberg. leth., 50, tabs. 2-4.
1969 Cleithranchiste paulusi Becker; H. Groos, Gottinger Arb. Geol. Palaont., 1, 57.
Holotype: Forschungs-Institut Senckenberg, Frankfurt am Main, Germany, no. SMF Xe 5146; an adult
carapace.
Type locality: Road cut behind transformer house, W exit from Soetenich village, Soetenich syncline, Eifel
Explanation of Plate 18, 22
Fig. 1, adult car. rt. lat. (holotype, SMF Xe 5146, 1100 /i.m long). Figs. 2, 3, adult RV (paratype, ex SMF Xe 5149. 1115/i.m long): fig.
2, adductor muscle scar; fig. 3, int. vent. obi.
Scale A (300/i.m; x70), figs. 1, 3; scale B (30/i.m; x230), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 23 Cleithranchiste paulusi (3 of 4)
Type locality
(cont):
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Mountains, Linksrheinisches Schiefergebirge, Germany; lat. 50°30'N, long. 06°38'W. Dark grey
marls with colonies of rugose corals; Rohr Member, Junkerberg Formation, late Eifelian, Middle
Devonian. Neritic facies, ostracod fauna of the Eifelian ecotype.
Forschungs-Institut Senckenberg (SMF), Frankfurt am Main, Germany, nos. SMF Xe 5146 (adult
car., holotype: PI. 18, 22, fig. 1; PI. 18, 24, figs. 2, 3), ex SMF 5149 (adult RV, paratype: PI. 18, 22,
figs. 2, 3; PI. 18, 24, figs. 4, 5), ex SMF Xe 5149 (adult LV, paratype: PI. 18, 24, fig. 1).
All of the figured specimens are topotype material.
Cleithranchiste species with almost symmetrically elliptical outline to the carapace; posterior end a
little more pointed than the anterior one. Right valve with a comparatively strong posteroventral
spine and distinct mid-dorsal depression. Dorsal outline of the carapace symmetrically biconvex.
Contact groove not interrupted mid-ventrally.
Cleithranchiste paulusi Becker, 1965 was originally put into the Family Healdiidae Harlton, 1933
(Healdiacea Harlton, 1933, Metacopina Sylvester-Bradley, 1961) because of its ‘healdiid’ shape,
and into the Subfamily Healdiopsidinae Griindel, 1962 because of its tripartite hinge. Adamczak
(Senckenberg. leth., 57, 360, 1976) demonstrated the tripartite hinge to be characteristic of the
Superfamily Thlipsuracea Ulrich, 1894 (Metacopina Sylvester-Bradley, 1961) and consequently he
assigned Cleithranchiste Becker to the Family Thlipsuridae Ulrich, 1894, and the Subfamily
Bufininae Sohn & Stover, 1961.
C. quasillitilis Adamczak, 1976, from the Eifelian of the Polish Mittelgebirge, is not closely
related to C. paulusi Becker; because of its marginal tubercles, it resembles Bufanchiste Becker,
1989 (see Becker, G., Stereo-Atlas Ostracod Shells , 18, 25-28, 1991). C. paulusi is considered to be
a benthic species.
Rheinisches Schiefergebirge of Germany; Eifelian, Middle Devonian.
Explanation of Plate 18, 24
Fig. 1, adult LV, int. lat. (paratype, ex SMF Xe 5149, 930 /xm long). Figs. 2, 3, adult car. (holotype, SMF Xe 5146, 1100 /im long): fig.
2, dors.; fig. 3, vent. Figs. 4, 5, adult RV (paratype, ex SMF Xe 5149, 1115/i.m long): fig. 4, adductor muscle scar; fig. 5, int. lat.
Scale A (300/xm; x70), figs. 1-5.
Stereo-Atlas of Ostracod Shells 18, 22
Cleithranchiste paulusi (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 24
Cleithranchiste paulusi (4 of 4)
Stereo-Atlas of Ostracod Shells 18(7)25-28(1991) Bufanchiste sotoi (1 of 4)
595.337.23 (113.44) (460 : 162.005.42 + 64) : 351.352 + 552.54
ON BUFANCHISTE SOTOI BECKER
by Gerhard Becker
(University of Frankfurt, Germany)
Genus BUFANCHISTE Becker, 1989
Type-species (by original designation): Bufanchiste sotoi Becker, 1989
Diagnosis: Thick-shelled, large, smooth and asymmetrical (left valve larger than right valve) thlipsurid genus.
Elliptical outline to the carapace. Two spines on each valve, perpendicularly set off from the
surface and situated equidistant from the ends of the valve. Hinge apparently tripartite. Marginal
tubercles present.
Distribution: W Europe and N Africa; late Emsian and possibly the late Eifelian, Devonian.
Bufanchiste sotoi Becker, 1989
1989 Bufanchiste sotoi sp. nov. G. Becker, Palaeontographica, A 209, 154, pi. 7, figs. 6, 7, pi. 10, fig. 9, tab. 2.
Holotype: Forschungs-Institut Senckenberg, Frankfurt am Main, Germany, no. SMF Xe 14395; an adult
carapace.
Type locality: W slope in the valley of the “Arroyo de la Vega”, 2 km W of the hamlet of Polentinos, about
10 km NW of Cervera de Pisuerga, Provincia de Palencia, N Spain; lat. 42°57'N, long. 04°32'W.
Dark grey marls from the top of the “middle” limestone lens; Polentinos Member, Abadia
Formation, late Emsian, Lower Devonian. Pelagic facies sensu lato\ deeper, open marine
environment with ostracod faunas of the Eifelian ecotype.
Explanation of Plate 18, 26
Fig. 1, adult car., ext. rt. lat (holotype, SMF Xe 14395, 1710/u.m long). Figs. 2, 3, adult LV (paratype, ex GPIF Cr 20/7, 1675pun long):
fig. 2, vent., posterior part of valve showing marginal tubercles; fig. 3, int. lat.
Scale A (300 pun; x48), figs. 1, 3; scale B (90 pun; x 100), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 27 Bufanchiste sotoi (3 of 4)
Figured specimens:
Diagnosis :
Remarks:
Distribution:
Forschungs-Institut Senckenberg (SMF), Frankfurt am Main, Germany, nos. SMF Xe 14395 (adult
car., holotype: PI. 18, 26, fig. 1), ex SMF 14397 (adult LV. paratype: PI. 18, 28, fig. 1), ex SMF Xe
14400 (adult RV, paratype: PI. 18, 28, figs. 2, 3). Geologisch-Palaeontologisches Institut (GPIF),
Frankfurt am Main, Germany, no. ex GPIF Cr 20/7 (adult LV, paratype: PI. 18, 26, figs. 2, 3).
All the figured paratypes are from the (possibly) late Eifelian; Jebl Rich, Antiatlas area, SW
Morocco.
Relatively very large Bufanchiste species with both spines about one quarter of carapace length
from the ends of the valve.
Bufanchiste sotoi Becker, 1989 is closely related to Cleithranchiste paulusi Becker, 1965 (see
Becker, G., Stereo-Atlas Ostracod Shells, 18, 21-24, 1991). It is, however, distinguished from this
taxon in having an elongated subelliptical lateral outline, two spines on each valve and a row of
marginal tubercles. According to Adamczak ( Senckenberg . leth., 57, 368, 1976), the latter feature
is characteristic of the Family Bufinidae Sohn & Stover, 1961. Bufinia species are distinctly smaller
than Bufanchiste sotoi ; moreover, their lateral outline is subrectangular to subelliptical and the
spines (or ridges) are situated asymmetrically on end margins of the valves.
Late Emsian, Lower Devonian of the Cantabrian Mountains, N Spain. Also the late Eifelian (?),
Middle Devonian, of the Antiatlas of SW Morocco.
Explanation of Plate 18, 28
Fig. 1, adult LV, vent, (paratype, SMF Xe 14397, 1850 gm long). Figs. 2, 3: adult RV (paratype, ex SMF Xe 14400, 1620 pm long): fig.
2, int. lat. obi.; fig. 3, vent.
Scale A (300 pun; x40), fig. 1; scale B (300 pun; x50), figs. 2, 3.
Bufanchiste sotoi (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 26
Stereo-Atlas of Ostracod Shells 18, 28
Bufanchiste sotoi (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (8) 29-32 (1991)
595.33.336.17 (113.51) (411 : 162.005.55 + 496.5 : 162.088.34): 551.351 + 552.52
Youngiella rectidorsalis (1 of 4)
ON YOUNGIELLA RECTIDORSALIS (JONES & KIRKBY)
by Christopher P. Dewey & Janet E. Coker
(Mississippi State University, Mississippi, U.S.A.)
Genus YOUNGIELLA Jones & Kirkby, 1886
Type-species (by original designation): Youngia rectidorsalis Jones & Kirkby, 1886.
r
1886 Youngia gen. nov. T. R. Jones & W. Kirkby, Proc. Geol. Ass., 9, 515, figs. 5-7.
1895 Youngiella nom. nov. T. R. Jones & W. Kirkby, Q. Jl geol. Soc. Lond , 42, 507 (pro Youngia Jones & Kirkby, 1886; non
Lindstorm, 1885).
Diagnosis: Minute, elongate, subrectangular, smooth youngiellid. Dorsal border straight, hinge taxodont.
Youngiella rectidorsalis (Jones & Kirkby, 1886)
1886a Youngia rectidorsalis n. sp. T. R. Jones & W. Kirkby, Proc. Geol. Assoc., 9, 515, figs. 5-7.
1886b Youngia rectidorsalis (Jones & Kirkby); T. R. Jones & W. Kirkby, Q. Jl geol. Soc. Lond., 42, 507.
1895 Youngiella rectidorsalis (Jones & Kirkby); T. R. Jones & W. Kirkby, Ann. Mag. Nat. Hist., ser. 6, 16, 456, pi. 21, figs. 5a-d.
Lectotype: (Here designated). British Museum (Natural History), London, no. OS 13641 (ex slide I 56).
[Paralectotypes nos. OS 13639, 13640, 13642, 13643, ex slides nos. I 1743 and I 56],
Type locality: Robroyston, about 4 miles NE of Glasgow, Lanarkshire, Scotland; lat. 55°54'N, long. 4°12'W.
Carboniferous Limestone, Visean, Lower Carboniferous.
Ligured specimens: British Museum (Natural History), London, nos. OS 13639, 13640 (ex slide I 1743); OS
13641-13643 (ex slide I 56). OS 13639 (car: PI. 18, 30, fig. 4), OS 13640 (RV: PI. 18. 32, figs. 1, 2).
OS 13641 (car: PI. 18, 30, fig. 3), OS 13642 (car: PI. 18, 30, fig. 2), OS 13643 (car: PI. 18, 30, fig. 1).
Dunn-Seiler Museum of Geology, Mississippi State University, U.S.A. , nos. 3341-4a (LV: PI. 18.
Explanation of Plate 18, 30
Fig. 1, car. ext. dors. (OS 13643, 410/j.m long); fig. 2, car., RV ext. lat. (OS 13642, 400 /xm long); fig. 3 car., LV ext. lat. (lectotype, OS
13641, 400p,m long); fig. 4, car. LV ext. lat. (OS 13639, 410/xm long). Scale A (100/u.m; x 140), figs. 1-4.
Stereo- Atlas of Ostracod Shells 18, 31
Youngiella rectidorsalis (3 of 4)
Ligured specimens
(cont):
Diagnosis:
Remarks:
Distribution :
Acknowledgement:
32, figs. 2, 3), 3341-4b (LV: PI. 18, 32, figs. 4, 5). B.M. (N.H.) specimens are from the type
locality: Nos. 3341-4a and 3341-4b are from light-brown fossiliferous mudstone, county highway
55, Sec. 35, T5S R11W, Colbert County, Alabama, U.S.A., lat. 34°34T5"N, long. 87°39T5"W.;
Bangor Limestone Formation, Chesterian, Mississippian, Carboniferous.
Minute, sub-oblong; dorsal and ventral margins straight, parallel; ends evenly rounded, posterior
end has slight ventral swing. Dorsal aspect suboblong, maximum width at midlength. Surface
smooth or faintly reticulate; marginal rims subdued, fade to ventral margin. Hinge taxodont; inner
lamella wide, narrows to posterior.
Jones & Kirkby (1886a) described the new genus Youngia and figured the dorsal and left lateral
views of a carapace and a right valve with taxodont hingement. Later in the same year Jones &
Kirkby, (1886b, 503, 507, 513) provided a generalised locality together with another description of
the genus. Armstrong etal. (1876) (Catalogue of Western Scottish Lossils, Blackie & Son, Glasgow,
45) listed Robroyston as being the only locality from which Y. rectidorsalis had been found. Later,
Jones & Kirkby described new material from the Yoredale ‘Series’ of Dunholme, Yorkshire,
England, changed the name of the genus (Youngia was preoccupied) and refigured the Scottish
material (Jones & Kirkby, 1895, 456, pi. 21, figs. 5a-d). Two slides of Y. rectidorsalis exist in the
British Museum (Nat. Hist.), both of which include specimens from the upper part of the
Carboniferous Limestone of Robroyston, Scotland. One of the slides (I 56) was purchased from J.
Armstrong in 1880 and the other (I 1743) was purchased from W. Kirkby in 1888. The types
designated herein are from these slides. No slides of the Yoredale material have been found.
Specimens of Y. rectidorsalis from the Black Warrior Basin in Alabama, U.S.A., confirm the
presence of a taxodont hinge and calcified inner lamella in this species.
Europe and U.S.A.; Lower Carboniferous.
We wish to acknowledge the financial support given by the Donors of the Petroleum Research
Fund administered by the American Chemical Society.
Explanation of Plate 18, 32
Fig. 1, RV int. lat. (OS 13640, 400 long). Figs. 2, 3, LV (3341-4a, 470/xm long): fig. 2, int. lat; fig. 3, ext. lat. Figs 4, 5, LV
(3341-4b, 405/u.m long): fig. 4, int. post, hinge, fig. 5, int. lat.
Scale A (100)u.m; x 137), fig. 1; scale B (100 /xm; x 130), figs. 2, 3, 5; scale C (20/u.m; x480), fig. 4.
Stereo-Atlas of Ostracod Shells 18, 32
Youngiella rectidorsalis (4 of 4)
Stereo -Atlas of Ostracod Shells 18, 30
Youngiella rectidorsalis (2 of 4)
Stereo-Atlas ofOstracod Shells 18 (9) 33-36 (1991)
595.337.14 (116.213) (261.27 : 162.010.50 + 429
162.004.53) : 551.351 + 552.52
Ektyphocythere bizoni (1 of 4)
ON EKTYPHOCYTHERE BIZONI AINSWORTH
by Ian Boomer & Nigel R. Ainsworth
(University of East Anglia , Norwich & Paleoservices, Watford, England)
Ektyphocythere bizoni Ainsworth, 1986
1986 Ektyphocythere bizoni sp. nov. N. R. Ainsworth, Bull. geol. Surv. lr., 3, 315, pi. 8, figs. 1-4.
Holotype: Trinity College, Dublin no. TCD 27570; 9 carapace.
[Paratypes nos: TCD 27571 - 27574],
Type locality: Fastnet Basin, Deminex Well 56/21-1 (lat. 50°18'54.66"N, long. 09°55T4.06"W). Late Toarcian-
Aalenian.
Figured specimens: Trinity College, Dublin no. TCD 27570 (holotype, 9 car.: PI. 18, 34, figs. 2, 5); British Geological
Survey, Keyworth, Nottingham nos. MPK 6957 (9 car.: PI. 18, 34, fig. 1), MPK 6460 (9 LV: PI.
18, 34, fig. 3), MPK 6954 (cf LV: PI. 18, 34, fig. 4), MPK 6956 (9 car.: PI. 18, 36, fig. 1), MPK
6952(9 RV: PI- 18, 36, fig. 2), MPK 6955 (cf car.: PI. 18, 36, fig. 3), MPK 6953 (cf RV: PI. 18,36,
fig. 4).
All specimens, apart from the holotype, are from the Mochras Borehole, Dyfed, Wales (Grid
Ref. SH 5533 2594); lat. 52°51'00"N, long. 4°06'30"W; Late Toarcian, Dumorteria levesquei Zone
( D . levesquei Subzone), Early Jurassic, at a depth of 605.43 - 606.88 metres.
Diagnosis: Carapace medium sized (550- 650 gm long), sub-triangular. Ornament of open ribbing arranged
in a triangular pattern. Three primary ribs form inverted “V”s above the median line, while two
arcuate/straight ribs occur below it. A further two or three longitudinal ribs can be seen along the
Explanation of Plate 18, 34
Fig. 1, 9 car., ext. dors. (MPK 6957, 564 /run long); figs. 2, 5, 9 car. (holotype, TCD 27570, 580 pm long): fig. 2, ext. It. lat.; fig. 5, ext.
rt. lat.; fig. 3, 9 LV, int. lat. (MPK 6460, 603/011 long); fig. 4, cf LV, ext. lat. (MPK 6954, 603/un long).
Scale A (100 /u,m; x90), figs. 1-5.
Stereo-Atlas of Ostracod Shells 18, 35
Ektyphocythere bizoni (3 of 4)
Diagnosis (cont): ventral and ventro-lateral surfaces. Intercostate regions smooth. Hinge antimerodont. Inner
lamella wide anteriorly, narrow posteriorly, no vestibula present. Anterior marginal pore canals
and muscle scars not observed.
Remarks: A species externally similar to both Ektyphocythere champeauae (Bizon, 1960, Rev. Micropa-
leont., 2, 206, pi. 1, fig. 1; pi. 22, fig. 1) and E. vitilis (Apostolescu et al ., 1961, in R. Mouterde
(Ed.), Mem. Bur. Rech. geol minier., 4, 399, pi. 1, fig. 1), but can be distinguished by the greater
number of primary longitudinal ribs. The specimen figured by Lord (1974) as E. cf. E. champeauae
(Palaeontology, 17, 614, pi. 90, fig. 16), appears similar to the present species although the
longitudinal ribs in his specimen seem more strongly developed. E. furcata (Weinholz) (in N.
Stoermer & E. Weinholz, 1967, Jb. Geol., 1 (for 1965), 548, pi. 2, figs. 19, 20), described from the
Toarcian of S. Germany, is similar to E. bizoni ; however, the former is slightly larger with more
robust ribbing.
In a review of N.W. European Liassic reticulate ostracods, Herrig (1985, Wiss. Z.
Ernst-Moritz-Arndt Univ. Griefswald, 34 (4), 45-50) erected the genera Nudacythere, Ernstella
and Rucholzella. He considered E. vitilis, E. furcata and E. champeauae to belong to Nudacythere ,
with the last-named as type species. The new genera of Herrig were erected on differences in the
development of the ribbing within the genus Ektyphocythere. We believe that since the hingement,
muscle scar patterns and marginal features within “ Ektyphocythere ” species are constant, generic
differentiation based on development of ornament alone is unnecessary.
Distribution: In the type area, poor stratigraphical control precludes an accurate assessment of the species
range. In the Mochras section, E. bizoni first appears near the top of the H. variabilis Zone, M.
Toarcian, becoming an abundant faunal element towards the top of the Lower Jurassic. Due to the
presence of an almost complete Lower Jurassic section at this site, it is assumed that the species
ranges into the Middle Jurassic.
Explanation of Plate 18, 36
Fig. 1, 9 car., ext. rt. lat. (MPK 6956, 551 pm long); fig. 2, 9 RV, int. lat. (MPK 6952, 513/xm long); fig. 3, cf car., ext. rt. lat. (MPK
6955, 577/u.m long); fig. 4, cf RV, int. lat. (MPK 6953, 538pm long).
Scale A (100/i.m) x90), figs. 1, 3; scale B (100/i.m; xl!4), figs. 2, 4.
Ektyphocythere bizoni (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 34
Stereo -Atlas of Ostracod Shells 18, 36
Ektyphocythere bizoni (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (10) 37-40 ( 1991) Frambocythere tumiensis ferreri (1 of 4)
595.337.14 (116.333.3) (460 : 161.004.42) : 551.312.4+552.52
ON FRAMBOCYTHERE TUMIENSIS (HELMDACH) FERRERI COLIN
by Jean-Paul Colin
(Esso Rep, Begles, France)
Frambocythere tumiensis (Helmdach) ferreri Colin, 1980
1971 Bisulcocypris (2 spp.?) A. Liebau, Bull. Cent. Rech. Pau, 5 suppl., 596, pi. 1, figs. 6, 7.
1980 Frambocythere tumiensis ferreri (sic) gen. et. subsp. nov., J.-P. Colin, in J.-P. Colin & D. L. Danielopol, Paleobiol. contin., 11.
16, pi. 8, figs. 1-10.
1980 Frambocythere tumiensis ferreri Colin; J.-F. Babinot, Trav. Lab. Geol. hist. Paleont. Univ. Provence, 10, 232, pi. 46, figs. 5-14.
1985 Frambocythere tumiensis ferreri Colin; J.-F. Babinot, J.-P. Colin & R. Damotte, Bull. Cent. Rech. Explor.-Prod.
Elf- Aquitaine, Mem. 9, 222, 254, pi. 70, figs. 8-14.
Holotype: Author’s collection, no. P 29-1; 9 left valve.
Type locality: Els Miquels de Moror, Lerida Province, Spain (lat. 42°04'10''N, long. 04°30'40"E) (see J. M. Pons,
Publnes. Geol. Univ. Auton. Bare., 3, 1—105 , 1977). Lagoonal facies with charophytes. Late
Maastrichtian (Garumnian).
Figured specimens: Author’s collection, nos. P 29-1 (9 LV: PI. 18, 38, fig. 1), P 29-2 (9 RV: PI. 18, 38. fig. 2), P 29-3
(CT RV: PI. 18, 38, fig. 3), P 29-4 (9 RV: PI. 18, 40, fig. 1), P 29-5 (cf LV: PI. 18, 40, fig. 2), P 29-6
(9 RV: 18, 40, fig. 3).
All are from the type locality.
Explanation of Plate 18, 38
Fig. 1, 9 LV, ext. lat. (holotype, P 29-1, 485 /um long); fig. 2, 9 RV ext, lat. (P 29-2, 476/xm long); fig. 3. Cf RV, ext. lat. (P 29-3,
461/u.m long).
Scale A (100/a.m; X 140), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 39 Frambocythere tumiensis ferreri (3 of 4)
Remarks:
Distribution :
Acknowledgements :
Frambocythere was erected by Colin (in Colin & Danielopol, 1980, op. cit., 15) with Bisulcocypris
tumiensis tumiensis Helmdach as type species. The genus, a member of the Timiriaseviinae, is
characterised by its small size, the presence of two subvertical sulci, pustulose ornamentation
(“raspberry”-type, hence the name), pronounced sexual dimorphism (females with a well
developed brood pouch) and a right valve larger than the left (i.e. inverse).
F. tumiensis ferreri differs from the other subspecies F. tumiensis tumiensis (Helmdach) and F.
tumiensis aepleri (Helmdach) (F. F. Helmdach, Berl. geowiss. Abh., (A), 3, 71-78, 1978), from the
Maastrichtian of N Spain, and F. tumiensis (Helmdach) ludi Tambareau (Y. Tambareau, Rev.
Micropaleont., 27, 145-148, 1984), from the Montian of Belgium, essentially by its unornamented
anterior half. Other characters are typical of the genus.
Late Maastrichtian (early Garumnian) of northern Spain (Pons, 1977, op. cit. ; Colin &
Danielopol, 1980, op. cit.) and of southern France (early Garumnian and Rognacian) (Babinot,
1980, op. cit.; 1986, Bull. Sue. linn. Provence, 38; Babinot et al., 1985, op. cit.; Bilotte et al., 1983,
Geol. mediterr., 10).
Dr. J.-F. Babinot (University of Marseille) is sincerely thanked for providing the SEM
micrographs.
Explanation of Plate 18, 40
Fig. 1, 9 RV, int. lat. (P 29-4, 470/um long); fig. 2, cf LV, int. lat. (P 29-5, 453^tm long); fig. 3, 9 RV, dors. (P 29-6, 484 /u.m long).
Scale A (100/u.m) x 140), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 38
Frambocythere tumiensis ferreri (2 of 4)
Frambocythere tumiensis ferreri (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 40
Stereo-Atlas of Ostracod Shells 18 (1 1) 41-44 ( 1991) Valdonniella mackenziei (1 of 4)
595.337.12 (116.333.3) (44 : 161.005.43) : 551.312.4 + 552.51 + 552.57
ON VALDONNIELLA MACKENZIEI BABINOT
by Jean-Franqois Babinot
(Universite de Provence, Centre Saint- Charles, Marseille, France)
VALDONNIELLA Babinot, 1980
Type-species (by original designation): Valdonniella mackenziei Babinot, 1980
Diagnosis: Carapace elongated, slightly arched in lateral view; left valve overlaps right in the anterior half of
the dorsal margin and medioventrally; a smaller overlap particularly evident at the posterior
cardinal angle. Greatest height in front of mid-point; anterior of this, dorsal margin of RV
concave, posteriorly convex. Carapace regularly inflated in dorsal and ventral view; ornamenta-
tion mostly smooth. Hinge merodont (lophodont) with smooth ridge-like anterior tooth. Central
muscle scars: 2 dorsal subcircular scars, 3 more-or-less connected scars in a horizontal line and 1
small scar below. Marginal zones with wide anterior and posterior vestibulum, marginal
pore-canals numerous, straight; selvage strong, peripheral. Sexual dimorphism inconspicuous.
Remarks: This genus displays several distinctive characters including the configuration of the central muscle
scars (Babinot, 1980, 239, text-fig. 8) and a pronounced anterio-dorsal overlap of the right valve.
Valdonniella shows some similarities with the Candonidae, particularly the lateral outline and
marginal zones. However, Candona Baird has a row of 5 subcircular scars, while Candonopsis
Vavra has 4 circular scars, another scar elongated below and 2 small accessory scars; both have an
adont hinge.
Explanation of Plate 18, 42
Fig. 1, LV, int. lat. (PVF 6/11, 610/tun long); fig. 2, car., ext. lat. (PVF 6/12, 620 /im long); fig. 3, RV, int. lat. (holotype, HVF 6,
610/xm long). Scale A (200 /u,m; x96), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 43
Valdonniella mackenziei (3 of 4)
Valdonniella mackenziei Babinot, 1980
1980 Valdonniella mackenziei n. gen., n. sp., J.-F. Babinot, Trav. Lab. Geol. hist. Paleont. Univ. Provence, 10, 240, pi. 47, figs.
14-16, pi. 48, figs. 1-8.
1985 Valdonniella mackenziei Babinot; J.-F. Babinot etal., Bull. Cent. Rech. Explor.-Prod. Elf -Aquitaine, Mem. 9, 222, 252. pi. 69,
figs. 7-11.
1987 Valdonniella mackenziei Babinot; J.-F. Babinot, Geol. mediterr., 14, 3, pi. 2, fig. 22.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Distribution:
Universite de Provence, Centre Saint-Charles (Centre de Sedimentologie et Paleontologie) no.
HVF 6; RV.
Les Ferrages, near La Fare-Les-Oliviers, Bouches-du-Rhone, SE France; approx, lat. 43°33'N.
long. 05°15'E. Valdonnian, late Cretaceous. In grey marls with lignitic horizons, molluscs,
gastropods and charophytes.
Universite de Provence, Centre Saint-Charles nos. PVF 6/11 (paratype, LV: PI. 18, 42, fig. 1),
PVF 6/12 (paratype, car.: PI. 18, 42, fig. 2), HVF 6 (holotype, RV: PI. 18, 42, fig. 3), PVF 6/13
(paratype, car.: PI. 18, 44, fig. 2), PVF 6/16 (paratype, car.: PI. 18, 44, fig. 1), PVF 6/17 (paratype,
car.: PI. 18, 44, fig. 3).
All specimens from type locality.
As for the genus; Valdonniella is currently monotypic.
Only known from oligohaline to freshwater deposits in the Valdonnian of southeastern France.
The age of the “Valdonnian” is in debate: magnetostratigraphic studies show it to be latest
Santonian, whereas previously it had been regarded as early Campanian.
Explanation of Plate 18, 44
Fig. 1, car., dors. (PVF 6/16, 620 /xm long); fig. 2, car., ext. lat. (PVF 6/13, 610/xm long); fig. 3, car., vent. (PVF 6/17, 610/u.m long).
Scale A (200/u,m) x96), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 42
Valdonniella mackenziei (2 of 4)
Valdonniella mackenziei (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 44
Stereo-Atlas of Ostracod Shells 18 (12) 45-48 (1991) Hemingwayella pumilio (1 of 4)
595.337.14 (119.9) (261.6 : 164.070.53) : 551.351
ON HEMINGWAYELLA PUMILIO (BRADY)
by Robin C. Whatley & Caroline A. Maybury
(Institute of Earth Studies, University College of Wales, Aberystwyth, U.K.)
Hemingwayella pumilio (Brady, 1880)
1880 Bythocythere pumilio sp. nov., G. S. Brady, Rep. scient. Results Voy. Challenger, (Zool.), 1 (3), 142, pi. 33, figs. 4a-d.
1976 Bythocythere pumilio Brady; H. S. Puri & N. C. Hulings, Bull. Br. Mus. nat. Hist. (Zool.), 29, 309, pi. 22, figs. 6-8.
Lectotype: British Museum (Nat. Hist.) no. 81.5.52; carapace. Designated by Puri & Hulings (1976, op. cit.).
Challenger Stn. 149, Balfour Bay, Kerguelen Island (lat. 49°08'S, long. 70°12'W). Depth 20-50
fathoms, in mud. Collected January 1874.
Kansas University Museum, Institute of Paleontology, Lawrence, Kansas, U.S.A. nos. KUMIP
1,084,564 (car.: PI. 18, 46, fig. 1), KUMIP 1,084,565 (car.: PI. 18, 46, fig. 2), KUMIP 1,084,566
(car., subsequently disarticulated: RV - PI. 18, 46, fig. 3; PI. 18, 48, figs. 3, 4; LV - PI. 18, 48, figs.
5, 6), KUMIP 1,084,567 (car.: PI. 18, 48, fig. 1), KUMIP 1,084,568 (car.: PI. 18, 48, fig. 2). All
specimens from Magellan Straits (lat. 52°37.3'S, long. 69°35.8'W), depth 9 m.
Carapace small (adults 450-500 pm in length), sub-rectangular, tumid, widest ventrally. Ornament
reticulate with elongate cribrose fossae, the long axes of which are vertical or slightly oblique;
median sulcus present, below and anterior to which is an inflated triangular area with 4 horizontal
muri; a strong, smooth alar ridge extends from anterior margin of valve to blunt process
posteroventrally (this ridge delimits rounded and inflated lateral surface from flattened venter); a
Type locality:
Figured specimens:
Diagnosis:
Explanation of Plate 18, 46
Fig. 1, car., ext. lat. (KUMIP 1,084,564, 500 /^m long); fig. 2, car., ext. lat. (KUMIP 1,084,565, 500^im long); fig. 3, RV, int. lat.
(KUMIP 1,084,566, 480/xm long).
Scale A (100/u.m; x 130), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 47 Hemingwayella pumilio (3 of 4)
Diagnosis (cont): distal rib extends subparallel to margin from eye tubercle, diverging before curving back to it at
posterior cardinal angle. Calcareous inner lamella wide, particularly anteriorly where there is a
shallow vestibulum. Hinge of RV with smooth, single teeth terminally separated by a long,
strongly locellate groove. Four adductor muscle scars, frontal scars not seen.
Remarks: The material from the Magellan Straits and the Atlantic coast of Patagonia is identical with the
lectotype of Brady’s species from Kerguelen Island (Dr. J. E. Whittaker, pers. comm.).
The genus Hemingwayella (J. W. Neale, Spec. Pap. Palaeont., 16, 30, pi. 13, figs. 8, 9; pi. 20,
figs. 3-6, text-figs. 5c, d, f, 1975) was first described from the Santonian, Upper Cretaceous of
Western Australia. All species of the genus possess the characteristic inflated, triangular area
antero-ventral of the median sulcus. The genus appears to be rare today and H. pumilio and an
undescribed species known to us from the Falkland Islands are possibly the only living
representatives. Such species as ? Eucytherura amfibola Barbieto-Gonzalez (1971, Mitt. hamb.
zool. Mus. Inst., 67, 301, pi. 27, figs, la, 2a, 3a) from the Mediterranean are only superficially
similar to Hemingwayella.
The oldest species of the genus known to the authors is H. aranea (Valicenti & Stephens,
1984) ( Revta . esp. Micropaleont., 16, 187, pi. 4, figs. 8-10; pi. 5, figs. 1-5) from the Valanginian of
the Alyoa Basin, South Africa.
Distribution: Recent of Kerguelen Island, S Indian Ocean (Brady, 1880). the Magellan Straits (9-25 m depth)
(Kaesler et al ., 1979, Proc. Vllth Int. Symp. Ostracodes, Serbian Geol. Soc., 239; and herein), and
the Argentinian coast from approx. 36° to 53°S in the intertidal zone and in offshore sediments
down to 131m water depth (Toy, 1985; Chadwick, 1986, Unpubl. M.Sc. theses, LIniv. Wales).
Explanation of Plate 18, 48
Fig. 1, car., dors. (KUMIP 1,084,567, 500/u.m long); fig. 2, car. vent. (KUMIP 1,084,568, 500/am long). Figs. 3, 4, RV (KUMIP
1,084,566, 480 pm long): fig. 3, ant. hinge element; fig. 4, post, hinge element. Figs. 5, 6, LV (KUMIP 1,084,566, 480 /u.m long):
fig. 5, ant. hinge element; fig. 6, post, hinge element.
Scale A (100/u.m; x 130), figs. 1, 2; scale B (20/xm; x540), figs. 3-6.
Hemingwayella pumilio (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 46
Stereo-Atlas of Ostracod Shells 18, 48
Hemingwayella pumilio (4 of 4)
Stereo- Atlas of Ostracod Shells 18 (13) 49-56 (1991) Cytheromorpha fuscata (1 of 8)
595.337.14 (119.9) (493 : 161.004.51+420 : 161.001.52) : 551.312.3 + 551.313.1
ON CYTHEROMORPHA FUSCATA (BRADY)
by Ian Boomer & David J. Horne
(University of East Anglia , Norwich & Thames Polytechnic, London)
Genus CYTHEROMORPHA Hirschmann, 1909
Type species (subsequent designation by Sars, 1925): Cythere fuscata Brady, 1869
( = Cytheromorpha albula Hirschmann, 1909).
1909 Cytheromorpha gen. nov., N. Hirschmann, Meddn Soc. Fauna Flora fenn., 35, 290-292.
Diagnosis: Carapace small, medium or large (300-750/xm long), subquadrate to subreniform in lateral view;
broadly rounded anterior margin with a narrow marginal rim; dorsal and ventral margins
converging posteriorly to a truncate posterior margin. Evenly inflated in dorsal view, tapering
anteriorly and somewhat truncated posteriorly. Pitted or reticulate; sometimes with a post-
eroventral alar protuberance in each valve. Dimorphic, male more elongate than female. Hinge
gongylodont, with two approximately equal-sized posterior teeth in the LV and a smooth median
element. Marginal zone relatively broad with a conspicuous anterior vestibulum; marginal pore
canals simple, few (10-20 anteriorly). Four adductor muscle-scars in a vertical row, frontal scar
tick-shaped, sometimes with a small, round scar above and in front; fulcral point prominent.
Antennula with six stout, articulated podomeres bearing strong chelate setae. Antennal
endopodite with four podomeres and two terminal chelate setae; exopodite (spinneret seta)
two-jointed. Branchial plate on mandible palp with 2-4 setae. Branchial plate on maxillula without
any reflexed setae. Legs slender; setal formulae: (1 + 1:2: 1), ( 1 + 1 : 1 ; 1 ) , 1 +1 : 1: 1). Furca with two
setae.
Explanation of Plate 18, 50
Figs. 1, 3, cf (lectotype, 1.58.27, 680 /xm long): fig. 1, LV ext. lat. ; fig. 3, RV ext. lat; fig. 2, 9 car. 1- lat. (paralectotype, 1.58.28,
540jU.ni long). Scale A (100/u.m; X 100), figs. 1-3.
Stereo- Atlas of Ostracod Shells 18, 51
Cytheromorpha fuscata (3 of 8)
Remarks: Cytheromorpha is externally similar to and often found in association with the genus Leptocy there,
which differs in having an entomodont hinge and branching marginal pore canals. For detailed
discussion of the taxonomy, ecology and distribution of the genus, see J. W. Neale & L. D.
Delorme, Revta esp. Micropaleont., 17, 41-64, 1985.
Cytheromorpha fuscata (Brady, 1869)
1869 Cythere fuscata sp. nov. G. S. Brady, Ann. Mag. nat. Hist., (ser. 4), 3, 47, pi. 7, figs. 5-8.
1869 Cythere drammensis sp. nov. G. O. Sars, Under spgelser over Christianiafjordens Dybvandsfauna, J. Dahl, Christiania, 56.
1909 Cytheromorpha albula sp. nov. N. Hirschmann, Meddn Soc. Fauna Flora fenn., 35, 290-292, figs. 7-8.
1925 Cytheromorhpa fuscata (Brady); G. O. Sars, An account of the Crustacea of Norway, 9, Ostracoda, parts 11-12, 177-178, pi. 81.
Lectotype: Here designated: Hancock Museum, Newcastle-upon-Tyne, no. 1.58.27; cf carapace (separated
into RV and LV).
(Paralectotype: no. 1.58.28; 9 carapace).
River Scheldt, Belgium (approx, lat. 51°15'N, long. 4°20'E). Recent, brackish/freshwater.
Hancock Museum nos. 1.58.27 (lectotype, cf; LV: PI. 18, 50, fig. 1; PI. 18, 52, fig. 3; RV: PI. 18,
50, fig. 3; PI. 18, 52, fig. 1), 1.58.28 (paralectotype, 9 car.: PI- 18, 50, fig. 2; PI. 18, 52, figs. 2, 4).
British Museum (Nat. Hist.) nos. 1991.3 (cf RV: PI. 18, 54, fig. 2), 1991.4 (juv.-l 9 car-: PI- 18,
54, fig. 4), 1991.5 (cf LV: PI. 18, 54, fig. 6), 1991.6 (9 RV; PI. 18, 54, fig. 1), 1991.7 (juv.-l cf
car.: PI. 18, 54, fig. 3), 1991.8 ($ LV: PI. 18, 54, fig. 5), 1991.9 (9 RV: PI. 18, 56, figs. 1, 5),
1991.10 (cf LV: PI. 18, 56, figs. 3, 6), 1991.11 (9 car.: PI. 18, 56, fig. 2), 1991.12 (cf car.: PI. 18,
56, fig. 4), 1991.194 (cf appendages: Text-fig. 2). The holotype and paratype were taken from
slide no. 2.04.02 in the G. S. Brady Collection at the Hancock Museum, Newcastle-upon-Tyne, on
which two cf carapaces still remain. A further 10 9 and 5 cf syntypic carapaces are on faunal slide
no. 2.11.19. Both are from the type locality. The Brit. Mus. (Nat. Hist.) specimens were all
collected alive in 1990 by Ian Boomer from just below water level on the river bank between
Heigham Sound and Martham Broad, Norfolk (approx, lat. 52°43'N, long. 01°36'E, Nat. Grid
Ref. TG 4395 1960); freshwater with brackish incursions on spring tides.
Type locality:
Figured specimens:
Explanation of Plate 18, 52
Figs. 1, 3, Cf (lectotype, 1.58.27, 680 /xm long): fig. 1, RV int. lat.; fig. 3, LV int. lat; figs. 2, 4, 9 car. (paralectotype, 1.58.28, 540 /um
long): fig. 2, vent.; fig. 4, dors. Scale A (100/um; X 100), figs. l^t.
Stereo-Atlas of Ostracod Shells 18, 53
Cytheromorpha fuscata (5 of 8)
Diagnosis:
Remarks:
Distribution :
Carapace medium to large (480-750pm long), strongly pitted with small, rounded fossae. 8-10
anterior marginal pore canals. Sexual dimorphism very conspicuous: male longer and more
inflated posteriorly than female; female with a small, knob-like posteroventral alar protuberance
in each valve, male with a compressed, smooth area behind a weak swelling in the same position.
Although extant populations were recorded from East Anglia in the last century (G. S. Brady &
D. Robertson, Ann. Mag. nat. Hist. (ser. 4), 6, 1-33, 1870), none of the specimens remaining in
Brady’s collection contain any appendages and the species has not subsequently been reported
living in Britain until now. We have found it alive in only one locality, although valves and
carapaces have been obtained in brackish/estuarine locations on the Rivers Yare, Bure and
Waveney.
Pleistocene to Recent. Fresh to brackish water (0.5-20%c) in Europe, Scandinavia, Canada and the
U.S.A. For further details of distribution and ecology see Neale & Delorme (op. cit.).
Text-figure 1: cf RV int. lat., seen in transmitted light (based on study of several
specimens). Scale bar = 100pm.
Explanation of Plate 18, 54
Fig. 1, 9 RV ext. lat. (1991.6, 551pm long); fig. 2, cf RV ext. lat (1991.3, 647 pm long); fig. 3, juv.-l cf car. rt. lat. (1991.7, 514pm
long); fig. 4, juv.-l $ car. rt. lat. (1991.4, 444 /xm long); fig. 5. 9 LV ext. lat. (1991.8, 551pm long); fig. 6, cf LV ext. lat. (1991.5,
667 pm long).
Scale A (100pm; x70), figs. 1-6.
Stereo-Atlas of Ostracod Shells 18, 55
Cytheromorpha fuscata (7 of 8)
Text-figure 2: cf appendages (1991.194); a, antennula; b, antenna; c, mandible; d, maxillula; e, first leg; f, second leg; g, third leg;
h, copulatory appendage. Scale bar = 100p.m.
Explanation of Plate 18, 56
Figs. 1, 5, 9 RV (1991.9, 551pm long): fig. 1, int. lat.; fig. 5, hinge; fig. 2, 9 car. dors (1991.11, 538pm long); figs. 3, 6, cf LV
(1991.10, 667pm long): fig. 3, int. lat.; fig. 6, hinge; fig. 4, cf car. dors. (1991.12, 667pm long).
Scale A (100pm; x85), figs. 1, 3; scale B (100pm; x70), figs. 2, 4; scale C (50pm; x 180), figs. 5, 6.
Cytheromorpha fuscata (6 of 8)
Stereo- Atlas of Ostracod Shells 18, 54
Stereo-Atlas of Ostracod Shells 18, 56
Cytheromorpha fuscata (8 of 8)
Stereo-Atlas of Ostracod Shells 18 (14) 57-60 (1991) On Vitjasiella ferox (1 of 4)
595.337.14 (119.9) (265.7 : 163.171.45) : 551.351
ON VITJASIELLA FEROX (HORNIBROOK)
by Michael A. Ayress
( Department of Geology, University of Otago, Dunedin, New Zealand
(Present address: Department of Geology, The Australian National University, Canberra))
Vitjasiella ferox (Hornibrook, 1953)
1953 Bythocythere ferox sp. nov., N. de B. Hornibrook, Trans. R. Soc. N.Z., 81, 307, text-fig. 2.1.
Holotype: Micropalaeontology Section, DSIR Geology & Geophysics, Lower Hutt, New Zealand no.
TOl 121/1; RV.
Type locality: Road cutting at Pukeuri, N E Otago, South Island, New Zealand; approx, lat. 45°03'S, long.
171°02'E. Originally referred to the “Awamoan” and dated as late Oligocene by Hornibrook {op.
cit .); now placed in the Altonian Stage and dated as early Miocene.
Figured specimens: Geology Museum, University of Otago, Dunedin, New Zealand nos. OU 39975 (LV: PI. 18, 58,
figs. 1, 3, 4; PI. 18, 60, fig. 3), OU 39976 (RV: PI. 18, 58, figs. 2, 5; PI. 18, 60, figs. 1, 2; text-fig. 1).
From off Oamaru, east coast of South Island, New Zealand, approx, lat. 45°06'S, long. 171°05'E:
Recent, from 68m depth.
Diagnosis: A species of Vitjasiella with prominent clavate spines bordering anterior and postero-ventral
margins and along extremity of ventro-lateral inflation. Hinge merodont with lobate anterior and
posterior terminal elements and a median element slightly expanded and crenulate distally.
Explanation of Plate 18, 58
Figs. 1 , 3, 4, LV (OU 39975, 950 yarn long): fig. 1, ext. lat.; fig. 3, ext. dors.; fig. 4, int. lat. Figs. 2, 5, RV (OU 39976, 970yu.m long): fig.
2, ext. lat.; fig. 5, int. lat.
Scale A (200/u.m; xllO), figs. 1-5.
Stereo-Atlas of Ostracod Shells 18, 59 Vitjasiella ferox (3 of 4)
Remarks: Hornibrook (1953, op. cit.) illustrated, by camera lucida drawing, only an external view of a right
valve. The hinge, in addition to Hornibrook’s observations, has a bilobate posterior terminal
element and a smaller arcuate anterior terminal element, also the median element is slightly
crenulate distally. These features, together with the spinose valve borders, serve to distinguish this
species from the type species Vitjasiella belyaevi Schornikov (1976, Abh. Verh. naturw. Ver.
Hamburg (n.s.), 18/19 suppl., 252, text-figs. 3-5), which possesses a smooth lophodont hinge. A
similar hinge-type to that of V. ferox is also present in Vitjasiella fenestrata (Brady) (1880, Rep.
scient. Results Voy. Challenger (Zool.) 1 (3), 139, pi. 34, fig. 6 = Cytheropteron fenestratum Brady;
see also H. S. Puri & N. C. Hulings, 1976, Bull. Br. Mus. nat. Hist. (Zool.), 29, 306, pi. 23, fig. 18,
pi. 24, figs. 1-6).
Recent specimens, as illustrated here, differ from the fossil type specimen very slightly in their
more pointed posterior outline and also lack spines on the short dorsal ridge. A third form of V.
ferox , which possesses two rows of ventro-lateral spines, occurs in the Waitakian (latest Oligocene
- earliest Miocene) to Altonian (early Miocene) stages of New Zealand.
Distribution: This Recent record extends the Runangan (latest Eocene) to Castlecliffian (late Pliocene) range of
V. ferox reported by Hornibrook (1953).
Text-fig. 1, RV, internal features observed in transmitted light (OU
39976, 970 yum long). Scale = 100 yum.
Explanation of Plate 18, 60
Figs. 1, 2, RV (OU 39976), anterior and posterior hinge detail, respectively. Fig. 3, LV (OU 39975), posterior hinge detail.
Scale A (50/zm; x590), fig. 1: scale B (50yum; x440); scale C (50yum; x500), fig. 3.
Stereo-Atlas of Ostracod Shells 18, 58
Vitjasiella ferox (2 of 4)
Stereo -Atlas of Ostracod Shells 18, 60
Vitjasiella ferox (4 of 4)
Stereo-Atlas of Ostracod Shells 18 ( 15) 61-64 ( 1991)
595.337.3 (116.33) (430 : 161.013.54) : 551.351+552.55
ON PUNCIA LEVIS HERRIG
by Ekkehard R. Herrig
( University of Greifswald, Germany)
Puncia levis ( 1 of 4)
Puncia levis Herrig, 1988
1988 Puncia levis sp. nov., E. Herrig, Geschiebekde. Aktuell, 4, 34, figs. 1, 2.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Sektion Geologische Wissenschaften, Universitat Greifswald no. SGWG 9287/1; left valve.
[Paratypes: SGWG 28290/1, left valve; SGWG 28290/2, right valve],
Fahrnitz beach, coast of Jasmund, Island of Riigen (Baltic Sea), Germany; lat. 54°33'N, long.
13°40'E. Flint erratic boulder. Late Maastrichtian.
Sektion Geologische Wissenschaften, Universitat Greifswald nos (SGWG) 28290/1 (LV: PI. 18,
62, figs. 1, 2; PI. 18, 64, fig. 2) and 28290/2 (RV: PI. 18, 64, fig. 1).
Both paratypes from flint erratic boulders of Upper Cretaceous age from Germany. No.
SGWG 28290/1 is from the type locality; 28290/2 is from the beach at Vierow, Greifswald Bay
(Baltic Sea); lat. 54°08'N. long. 13°35'E.
Adult valves elongate, 410-460 gm long. Very weak sulcus in central part of valve, otherwise valve
lateral surface is gently curved. Velum-like adventral ridge extends between cardinal corners, is
best developed and is of reasonable width below mid height; its upper surface has a ridge with tiny,
closely spaced processes. Valve lateral surfaces are finely reticulate to punctate.
Explanation of Plate 18, 62
Figs. 1, 2, LV (paratype, SGWG 28290/1, 410/u.m long): fig. 1, ext. lat.; fig. 2, int. lat.
Scale A (100 ^un; x200), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 18, 63 Puncia levis (3 of 4)
Remarks: Detailed study of the holotype and newly-found additional material of P. levis, here illustrated,
reveals fine, dense ornament on all lateral parts of the valve except the velum. Reticulation is
clearly s.een (PI. 18, 62, fig. 1); a tendency towards developing punctation (PI. 18, 64, fig. 1) may
reflect factors of preservation.
This species is similar to P. goodwoodensis Hornibrook (N. de B. Hornibrook, Micropaleon-
tology, 9, 319, text-figs. 1, 2, 1963) from the Lower Miocene of New Zealand, but differs in the
presence of a ridge at the base of its velum. Puncia Hornibrook (1949, Trans. R. Soc. N.Z., 77,
470) is type genus of the Punciidae Hornibrook, 1949. Like the punciid Manawa Hornibrook
(1949, ibid., 470), for which soft-parts have recently been described in detail (K. M. Swanson,
Cour. Forschlnst. Senckenburg, 113, 11-20, 235-249, 1989), Puncia probably belongs to platycope
stock.
Distribution: Upper Cretaceous of the north German - central Baltic area; flint erratice boulders originally from
late Maastrichtian chalk of the Danish-Polish furrow.
Explanation of Plate 18, 64
Fig. 1, RV, ext. lat. (paratype, SGWG 28290/2, 450 jum long); fig. 2, LV, ext. lat., detail showing pitted surface (paratype, SGWG
28290/1).
Scale A (100/u.m; x200), fig. 1; scale B (lO/un; x 1250), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 62
Puncia levis (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 64
Puncia levis (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (16) 65-68 (1991) Capricambria cornucopiae (1 of 4)
595.330 (113.23) (943 : 163.139.22) : 551.35 + 552.64
ON CAPRICAMBRIA CORNUCOPIAE GEN. ET SP. NOV.
by Ingelore C. U. Hinz
(University of Bonn, Germany)
Genus CAPRICAMBRIA gen. nov.
Type species: Capricambria cornucopiae sp. nov.
From Latin capra , a goat + Cambria ; referring to the large, lateral horn-like spines and the
Cambrian occurrence, respectively. Gender, feminine.
Bradoriid with carapace subamplete, straight hinge line and flattened free marginal area. Valves
symmetrical; each with two large, cornutiform spines and a steeply elevated comarginal ridge
which parallels valve margin. Outer surface of valve reticulate.
Capricambria cornucopiae sp. nov.
Institut fur Palaontologie, University of Bonn, Germany, no. UB 209; carapace.
lkm north of Mt. Murray, Queensland, Australia (lat. 21°48.8'S, long. 139°58.5'E); phosphorite
deposits of the Duchess Region; T. gibbus Zone, Middle Cambrian.
From Latin cornu copiae , the goat’s horn or horn of plenty; alluding to the horn-like lateral spines.
Used as a noun in apposition.
University of Bonn, Germany, no. UB 209 (holotype, car.: PI. 18, 66, figs. 1-3, PI. 18, 68, figs. 1,
2). From the type locality.
Explanation of Plate 18, 66
Figs. 1-3, crumpled car. (holotype, UB 209, 670yu.m long): fig. 1, ext. lt.(?) lat.; fig. 2, ext. ant.; fig. 3, ext. post.
Scale A (100/u.m; x 105), figs. 1, 3; scale B (100/u.m; x 120), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 67 Capricambria cornucopiae (3 of 4)
Diagnosis: Carapace equivalved and almost semicircular in outline; amplete. Dorsal corners form
approximate right angles. Hinge line straight and simple, with distinctly separate valves except for
a very short portion at either end; dorsum rather narrow. Maximum length of valve at about mid
height. Area along free margin flattened and set off from rest of valve by a steep ridge that runs
from anterodorsal to posteroventral region; ridge gradually decreases in height and terminates
close to a well developed vertical spine. Adjacent to the ridge and at about the same height, a
smaller, anterior spine is developed. Except for the spines and ridge the valve’s outer surface is
reticulate, consisting of irregular polygons. Marginally, corners of polygons may be the sites of
minute nodes. Spines show irregular annulations (possibly caused by shrinkage?).
Remarks: Among Cambrian bradoriids, the presence of well developed spines is rather rare. If spines are
developed at all, they are generally situated in the dorsal or ventral area. The genus Monasterium
Fleming (1973, Pubis geol. Surv. Qd., 356, 8), for example, has a long anterodorsal spine on either
valve, but because of its delicate nature it is usually broken. The subgenus Kunmingella
( Spinokunmingella ) Huo & Shu (1985, Cambrian Bradoriida of South China , Northwest Univ.
Publ. House, Xian, 113) is supposed to have a well developed posteriorly directed ventral spine.
Another character of Capricambria is its reticulate outer surface, a feature which it has in
common with, for example, Monasterium and Zepaera Fleming (1973, op. cit.), Polycostalis Shu
(1990, Cambrian and Lower Ordovician Bradoriida from Zhejiang, Hunan and Shaanxi Provinces ,
Chinese Univ. of Geology, Beijing, 63) and Flemingopsis Jones & McKenzie (1981, Alcheringa, 5,
310). A common character between Flemingopsis and Capricambria is the flattened free marginal
area and the highly convex rest of the valve. But, in contrast to the more rounded elevation in
Flemingopsis, Capricambria has a steep ridge.
The virtually unbroken, crumpled nature of the carapace of Capricambria might argue for a
flexible, at most slightly mineralised wall substance. Similar preservation has been observed in a
great many bradoriid specimens, particularly of the genus Monasterium. Due to the crumpled
condition of the carapace, the suggested orientation in Capricambria is somewhat questionable.
Distribution: Known only from the type locality.
Explanation of Plate 18, 68
Figs. 1, 2, crumpled car. (holotype, UB 209): fig. 1, ext. vent.; fig. 2, ext. dors. Scale A (100/xm; x 140), figs. 1, 2.
Derivation of name:
Diagnosis:
Holotype:
Type locality:
Derivation of name:
Figured specimen :
Stereo-Atlas of Ostracod Shells 18, 66
Capricambria cornucopiae (2 of 4)
BPCC Blackpool
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Stereo-Atlas of Ostracod Shells: Vol. 18, Part 1
CONTENTS
Orcofabella testata (Gailite); by D.J. Siveter & L. Sarv.
Limbinariella macroreticulata Sarv; by D.J. Siveter & L. Sarv.
Venzavella costata (Neckaja); by D.J. Siveter & L. Sarv.
Lomatopisthia simplex (Harris); by M. Willliams.
Lomatopisthia auricula (Harris); by M. Willliams.
Cleithranchiste paulusi Becker; by G. Becker.
Bufanchiste sotoi Becker; by G. Becker.
Youngiella rectidorsalis (Jones & Kirkby); by C.P. Dewey & J.E. Coker.
Ektyphocythere bizoni Ainsworth; by I. Boomer & N.R. Ainsworth.
Frambocy there tumiensis (Helmdach) ferreri Colin; by J.-P. Colin.
Valdonniella mackenziei Babinot; by J.-F. Babinot.
Hemingwayella pumilio (Brady); by R.C. Whatley & C.A. Maybury.
Cytheromorpha fuscata (Brady); by I. Boomer & D.J. Horne.
Vitjasiella ferox (Hornibrook); by M.A. Ayress.
Puncia levis Herrig; by E.R. Herrig.
Capricambria comucopiae Hinz gen et sp. nov.; by I.C.U. Hinz.
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