A Stereo-Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter,
and J. E. Whittaker
Published under the aegis of the British Micropalaeontological Society, London
ISSN 0952-7451
Editors
Dr J. Athersuch, Exploration Technology Branch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Dr D.J. Horne, School of Earth Sciences, Thames Polytechnic, Walburgh House, Bigland Street,
London El 2NG.
Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.
Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.
Editorial Board
Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.
Dr P. de Deckker, Department of Geology, Australian National University, G.P.O. Box 4, Canberra,
ACT 2601, Australia.
Dr W. Hansch, Ernst-Moritz-Arndt Universitat, Sektion Geologische Wissenschaften, F.L.-Jahnstr. 17a,
2200 Greifswald, Germany.
Prof. R. Lundin, Department of Geology, Arizona State University, Tempe, Arizona 85287-1404, U.S.A.
Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55,
D 2000 Hamburg 13, Germany.
Prof. N. Ikeya, Institute of Geosciences, Shizuoka University, Shizuoka 422, Japan.
Officers of the British Micropalaeontological Society
Chairman Professor M.B. Hart, Department of Geological Sciences, Polytechnic South West, Drake Circus,
Plymouth, Devon PL4 8AA.
Secretary Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.
Assistant Treasurer Dr I.P. Wilkinson, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Journal Editor Dr M.C. Keen, Department of Geology, The University, Glasgow G12 8QQ.
Newsletter Editor Dr D.J. Shipp, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Conodont Group Chairman Dr H.A. Armstrong, Department of Geological Sciences, University of Durham,
Science Laboratories, South Road, Durham DH1 3LE.
Conodont Group Secretary Mr M.T. Dean, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Foraminifera Group Chairman Dr H.W. Bailey, Paleo Services Ltd., Unit 15, Paramount Industrial Estate,
Sandown Road, Watford WD2 4XA.
Foraminifera Group Secretary Dr F.M.D. Lowry, School of Earth Sciences, Thames Polytechnic,
Walburgh House, Bigland Street, London El 2NG.
Ostracod Group Chairman Dr J. Athersuch, BP Research, Sunbury Research Centre, Chertsey Road,
Sunbury-on-Thames, Middlesex TW16 7LN.
Ostracod Group Secretary Dr I.D. Boomer, School of Environmental Sciences, University of East Anglia,
Norwich NR4 7TJ.
Palynology Group Chair Dr R.J. Davey, The Robertson Group pic, Ty’n-y-Coed, Llanrhos, Llandudno,
Gwynedd LL30 ISA.
Palynology Group Secretary Dr A. McNestry, British Geological Survey, Keyworth, Nottingham NG12 5GG.
Calcareous Nannofossil Group Chairman Dr L.T. Gallagher, Paleo Services, Unit 15, Paramount Industrial
Estate, Sandown Road, Watford WD2 4XA.
Calcareous Nannofossil Group Secretary Dr N.M. Hine, British Geological Survey, Keyworth, Nottingham
NG12 5GG.
Instructions to Authors
Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. Format
should follow the style set by the papers in this issue. Descriptive matter apart from illustrations should be
cut to a minimum; preferably each plate should be accompanied by only one page of text. Blanks to aid in
mounting figures for plates may be obtained from any one of the Editors or Editorial Board. Completed
papers should be sent to one of the Editors. All contributions submitted for possible publication in the Stereo-
Atlas of Ostracod Shells are reviewed by an appropriate international specialist.
The front cover shows a male left valve and appendages, internal view, of Linmocythere borisi borisi Martens,
1990. Paratypc, K.B.I.N., Brussels, OC.1406. From Lake Abijata, Ethiopia. Photographed by K. Martens and
J. Cillis.
A Stereo-Atlas of Ostracod Shells
edited by J. Athersuch, D. J. Horne, D. J. Siveter,
and J. E. Whittaker
Volume 18, 1991
Part 1 (pp. 1-68); 31st July, 1991
Part 2 (pp. 69-137); 31st December, 1991
Published under the aegis of the British Micropalaeontological Society, London
Stereo-Atlas of Ostracod Shells 18, ii
Contents
Contents
1 On Orcofabella testata (Gailite); by D.J. Siveter & L. Sarv. 1
2 On Limbinariella macroreticulata Sarv; by D.J. Siveter & L. Sarv. 5
3 On Vertzavella costata (Neckaja); by DJ. Siveter & L. Sarv. 9
4 On Lomatopisthia simplex (Harris); by M. Williams. 13
5 On Lomatopisthia auricula (Harris); by M. Williams. 17
6 On Cleithranchiste paulusi Becker; by G. Becker. 21
7 On Bufanchiste sotoi Becker; by G. Becker. 25
8 On Youngiella rectidorsalis (Jones & Kirkby); by C.P. Dewey & J.E. Coker. 29
9 On Ektyphocythere bizoni Ainsworth; by I. Boomer & N.R. Ainsworth. 33
10 On Frambocythere tumiensis (Helmdach) ferreri Colin; by J.-P. Cohn. 37
11 On Valdoniella mackenziei Babinot; by J.-F. Babinot. 41
12 On Hemingwayella pumilio (Brady); by R.C. Whatley & C.A. Maybury. 45
13 On Cytheromorpha fuscata (Brady); by I. Boomer & D.J. Horne. 49
14 On Vitjasiella ferox (Hornibrook); by MA. Ayress. 57
15 On Puncia levis Herrig; by E.R. Herrig. 61
16 On Capricambria comucopiae Hinz gen. et sp. nov.; by I.C.U. Hinz. 65
17 On Ulopsis ulula Hinz gen. et sp. nov.; by I.C.U. Hinz. 69
18 On Mammoides dorsospinosus Sohn; by C.P. Dewey & T.M. Puckett. 73
19 On Qujingsia nonaculeata Hansch & Wang gen. et sp. nov.; by W. Hansch &
Wang Shang-qi. 77
20 On Daleiella corbuloides (Jones & Holl); by R.F. Lundin & L.E. Petersen. 81
21 On ViUozona villosa (Griindel); by G. Becker & D. Weyer. 85
22 On Huntonella bransoni Lundin; by D.J. Siveter & R.F. Lundin. 89
23 On Paracathaycythere costaereticulata Whatley & Zhao gen. et sp. nov.; by R.C. Whatley &
Zhao Quanhong. 93
24 On Paracathaycythere scabra Zhao & Whatley gen. et sp. nov.; by Zhao Quanhong &
R.C. Whatley. 97
25 On Cocoonocythere sinensis Zhao; by Zhao Quanhong & R.C. Whatley. 101
26 On Polydontoconcha hyperdonta Zhao & Whatley gen. et sp. nov.; by Zhao Quanhong &
R.C. Whatley. 105
27 On Palmoconcha ruggierii Maybury sp. nov.; by CA. Maybury. 109
28 On Ektyphocythere cookiana (Anderson); by I. Boomer. 113
29 On Tethysobuntonia govoroffi Colin & Babinot gen. et sp. nov.; by J.-P. Colin &
J.-F. Babinot. 117
30 On Ogmoconchella martini (Anderson); by I. Boomer. 121
31 On Frostiella groenvalliana Martinsson; by W. Hansch, D.J. Siveter & C.G. Miller. 125
32 Index for Volume 18, (1991). 135
Stereo-Atlas of Ostracod Shells 18 (17) 69-72 (1991) Ulopsis ulula (1 of 4)
595.33.330 (113.23) (943 : 163.139.22): 551.35 + 552.64
ON ULOPSIS ULULA HINZ gen. et. sp. nov.
by Ingelore C.U. Hinz
(University of Bonn, Germany)
Derivation of name:
Diagnosis:
Remarks:
Genus ULOPSIS gen. nov.
Type-species: Ulopsis ulula sp. nov.
Latin ulula, owl, and -opsis, like; gender feminine.
Hesslandonid with nearly equivalved, subamplete to slightly postplete carapace. Valve outline almost semi-
circular; “hinge line’’ straight, except near cardinal corners. Carapace very inflated. Maximum length above
mid-height. Interdorsum (= dorsum and dorsal bar of Muller 1964, 1982 respectively) fairly broad, apparently
demarcated by simple bends or breaks (“hinges”), forms acroidal processes. Free margin equally convex, free
marginal area flattened and distinctly set off from inflated rest of valve. Large oval lobe just in front of mid-
dorsal area, separated from much smaller, curved anterodorsal lobe by shallow sulcus. Inclined, elongate lobe
occurs behind approximately mid-dorsal, triangular depression. Posterodosal area has a subdued node, below
which another, much weaker node may be developed.
In larger instars the oval, anterior lobe is laterally inflated to beyond the lower half of the valve, whereas the
oblique, elongate lobe remains relatively small. The smooth outer surface of the valve may be crumpled, thus
indicating a certain flexibility of the shell.
With its distinct interdorsum Ulopsis belongs to hesslandonids that are characterised by a “double hinge”
(Muller, K.,NeusJb. Geol. Paldont . Abh. ,121, 1964; Muller, K .in: Bate, R. H . et al. , Fossil and Recent Ostracods,
1982, Ellis Horwood, Chichester). It is uncertain whether or not Ulopsis has adont (nullidont) hinges or only
bends in the shell. Carapaces with dorsal bends indicate a fairly flexible shell material; however, a distinct
separation into valves results from increased mineralisation which presumably made the shell more brittle. Both
Explanation of Plate 18, 70
Figs. 1, 2, car. (holotype, CPC 23/S4, 0.80 mm long); fig. 1, ext. It. lat.; fig. 2, ext. dors. Fig. 3, incomplete car. (CPC 23/S3, 1.34 mm
long), ext. dors. Scale A (100 pm\ x75), figs. 1, 2; scale B (250//m; x60), fig. 3.
Stereo-Atlas of Ostracod Shells 18, 71
Ulopsis ulula (3 of 4)
developments are considered not to differ fundamentally from each other; they may be due to evolutionary
processes or may represent ecological adaptations. In any event the presence of a broad interdorsum seems to
be quite a primitive character, possibly resulting from a minor lateral compression of the ostracod body at an
early stage of evolution. It is presently uncertain whether hesslandonids are intermediate in the development
from univalved dabashanellids to “normal” bivalved forms or whether they represent a special off-shoot in the
early evolution of ostracods.
In its basic lobation, having antero- and posterodorsal lobes, Ulopsis is comparable to the Palaeozoic
Binodicopa. There is also superficial similarity to Kunmingelloides Shu, 1990 ( Cambrian and Lower Ordovician
Bradoriida from Zhejiang, Hunan and Shaanxi Provices , 54, Northwest Univ. Press), whose distinct lobes,
located at either end of the shell, might be homologous to the small lobes of Ulopsis. However, Ulopsis is
distinguished by its fairly broad interdorsum, the presence of acroidal processes and by its shape and lobal
arrangements.
Soft integument preservation in phosphatocopid ostracods is only known in specimens from Sweden (e.g.
Muller 1979, Lethaia, 12, 1-27; 1982) and Great Britain (Hinz 1987, Palaeontographica, 198-A, 59). However,
one dorsally broken specimen of Ulopsis (PI. 18, 70, fig. 3) exposes some relics of its original body. Proper
appendages are not recognizable; instead, phosphatized, now crumpled tissue that lined or filled the integument
roughly reflects its original position. These structures occur together with hypha-like threads that are quite
common in body cavities. These separate phenomena should not be confused.
Derivation of name:
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Distribution:
Ulopsis ulula sp. nov.
Latin ulula , an owl; fancied resemblance of valve in lateral view.
Bureau of Mineral Resources, Canberra, no. CPC 23/S4; carapace.
Rogers Ridge, Queensland, Australia (lat. 21°45,4'S, long. 139°58,8'E); phosphorite deposits of the Duchess
Region, Triplagnostus gibbus Zone, middle Cambrian.
Bur. Min. Res. nos. CPC 23/S4 (holotype, car.: PI. 18, 70, figs. 1, 2), CPC 23/S3 (incomplete car.: PI. 18, 70,
fig. 3), CPC 23/S5 (crumpled RV: PI. 18, 72, fig. 1), CPC 23/S7 (car.: PI. 18, 72, fig. 2) and CPC 23/S6 (car.:
PI. 18, 72, fig. 3). All from the type locality.
As for the genus. Ulopsis is currently monotypic.
Known only from type locality.
Explanation of Plate 18, 72
Fig. 1, crumpled RV (CPC 23/S5, 1.29 mm long), ext. lat.; Fig. 2, car. (CPC 23/S7, 1.37 mm long), ext. vent.; Fig. 3 car. (CPC 23/S6,
1.22 mm long), ext. ant. Scale A (250 /rm; x50), fig. 1; scale B (100 gm; x70), fig. 2; scale C (250 /tm; x60), fig. 3.
Stereo-Atlas of Ostracod Shells 18, 70
Stereo-Atlas of Ostracod Shells 18, 72
Ulopsis ulula (2 of 4)
Ulopsis ulula (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (17) 69-72 (1991) Ulopsis ulula (1 of 4)
595.33.330 (113.23) (943 : 163.139.22) : 551.35 + 552.64
ON ULOPSIS ULULA HINZ gen. et. sp. nov.
by Ingelore C.U. Hinz
(University of Bonn, Germany)
Derivation of name:
Diagnosis:
Remarks:
Genus ULOPSIS gen. nov.
Type-species: Ulopsis ulula sp. nov.
Latin ulula, owl, and -opsis, like; gender feminine.
Hesslandonid with nearly equivalved, subamplete to slightly postplete carapace. Valve outline almost semi-
circular; “hinge line’’ straight, except near cardinal corners. Carapace very inflated. Maximum length above
mid-height. Interdorsum (= dorsum and dorsal bar of Muller 1964, 1982 respectively) fairly broad, apparently
demarcated by simple bends or breaks (“hinges”), forms acroidal processes. Free margin equally convex, free
marginal area flattened and distinctly set off from inflated rest of valve. Large oval lobe just in front of mid-
dorsal area, separated from much smaller, curved anterodorsal lobe by shallow sulcus. Inclined, elongate lobe
occurs behind approximately mid-dorsal, triangular depression. Posterodosal area has a subdued node, below
which another, much weaker node may be developed.
In larger instars the oval, anterior lobe is laterally inflated to beyond the lower half of the valve, whereas the
oblique, elongate lobe remains relatively small. The smooth outer surface of the valve may be crumpled, thus
indicating a certain flexibility of the shell.
With its distinct interdorsum Ulopsis belongs to hesslandonids that are characterised by a “double hinge”
(Muller, K.,Neus Jb. Geol. Palaont.Abh., 121, 1964; Muller, K. in: Bate, R. H . et al. , Fossil and Recent Ostracods ,
1982, Ellis Horwood, Chichester). It is uncertain whether or not Ulopsis has adont (nullidont) hinges or only
bends in the shell. Carapaces with dorsal bends indicate a fairly flexible shell material; however, a distinct
separation into valves results from increased mineralisation which presumably made the shell more brittle. Both
Explanation of Plate 18, 70
Figs. 1, 2, car. (holotype, CPC 23/S4, 0.80 mm long); fig. 1, ext. It. lat.; fig. 2, ext. dors. Fig. 3, incomplete car. (CPC 23/S3, 1.34 mm
long), ext. dors. Scale A (100 /rm; x75), figs. 1, 2; scale B (250 pm; x60), fig. 3.
Stereo-Atlas of Ostracod Shells 18, 71
Ulopsis ulula (3 of 4)
developments are considered not to differ fundamentally from each other; they may be due to evolutionary
processes or may represent ecological adaptations. In any event the presence of a broad interdorsum seems to
be quite a primitive character, possibly resulting from a minor lateral compression of the ostracod body at an
early stage of evolution. It is presently uncertain whether hesslandonids are intermediate in the development
from univalved dabashanellids to “normal” bivalved forms or whether they represent a special off-shoot in the
early evolution of ostracods.
In its basic lobation, having antero- and posterodorsal lobes, Ulopsis is comparable to the Palaeozoic
Binodicopa. There is also superficial similarity to Kunmingelloides Shu, 1990 ( Cambrian and Lower Ordovician
Bradoriida from Zhejiang, Hunan and Shaanxi Provices, 54, Northwest Univ. Press), whose distinct lobes,
located at either end of the shell, might be homologous to the small lobes of Ulopsis. However, Ulopsis is
distinguished by its fairly broad interdorsum, the presence of acroidal processes and by its shape and lobal
arrangements.
Soft integument preservation in phosphatocopid ostracods is only known in specimens from Sweden (e.g.
Muller 1979, Lethaia, 12, 1-27; 1982) and Great Britain (Hinz 1987, Palaeontographica, 198-A, 59). However,
one dorsally broken specimen of Ulopsis (PL 18, 70, fig. 3) exposes some relics of its original body. Proper
appendages are not recognizable; instead, phosphatized, now crumpled tissue that lined or filled the integument
roughly reflects its original position. These structures occur together with hypha-like threads that are quite
common in body cavities. These separate phenomena should not be confused.
Derivation of name:
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Distribution:
Ulopsis ulula sp. nov.
Latin ulula, an owl; fancied resemblance of valve in lateral view.
Bureau of Mineral Resources, Canberra, no. CPC 23/S4; carapace.
Rogers Ridge, Queensland, Australia (lat. 21° 45,4' S, long. 139° 58,8' E); phosphorite deposits of the Duchess
Region, Triplagnostus gibbus Zone, middle Cambrian.
Bur. Min. Res. nos. CPC 23/S4 (holotype, car.: PI. 18, 70, figs. 1, 2), CPC 23/S3 (incomplete car.: PI. 18, 70,
fig. 3), CPC 23/S5 (crumpled RV: PI. 18, 72, fig. 1), CPC 23/S7 (car.: PL 18, 72, fig. 2) and CPC 23/S6 (car.:
PL 18, 72, fig. 3). All from the type locality.
As for the genus. Ulopsis is currently monotypic.
Known only from type locality.
Explanation of Plate 18, 72
Fig. 1, crumpled RV (CPC 23/S5, 1.29 mm long), ext. lat.; Fig. 2, car. (CPC 23/S7, 1.37 mm long), ext. vent.; Fig. 3 car. (CPC 23/S6,
1.22 mm long), ext. ant. Scale A (250/^m; x50), fig. 1; scale B (100 //m; x70), fig. 2; scale C (250//m; x60), fig. 3.
Stereo-Atlas of Ostracod Shells 18, 70
Ulopsis ulula (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 72
Ulopsis ulula (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (18) 73-76 (1991)
595.336 (113.51) (761 : 162.88.34 + 87.34): 551.351 + 552.52 + 552.54
Mammoides dorsospinosus (1 of 4)
ON MAMMOIDES DORSOSPINOSUS SOHN
by Christopher P. Dewey & T. Mark Puckett
( Mississippi State University & Alabama Geological Survey, U.S.A.)
Mammoides dorsospinosus Sohn, 1961
1961 Mammoides dorsospinosa (sic) sp. nov., I. G. Sohn, Prof. Pap. U.S. geol. Surv., 330-B, 114, pi. 7, figs. 15, 16.
Ho lo type:
Type locality:
Figured specimens:
United States National Museum, Washington, U.S.A. , no. USNM 119818; crushed? juvenile
carapace.
[Paratypes, no. USNM 119819, 15 specimens].
USNM locality no. 1086, Brownwood Shale, Canyon Group, Missourian, Upper Pennsylvanian,
Carboniferous; central Texas, U.S.A. Collected by A.R. Loeblich. Loeblich’s field notes have been
lost and an exact type locality is unknown (Sohn, pers. comm.).
Dunn-Seiler Museum of Geology, Mississippi State University, U.S.A., nos. 3341-3a (RV: PI. 18,
76, fig. 2), 3341-3b (RV: PI. 18, 76, fig. 1), 3341-3c (RV: PI. 18, 74, fig. 4), 3341-3d (LV: PI. 18, 74,
fig. 3), 3341-3e (RV: PI. 18, 74, figs. 1, 2).
3341-3a, 3b and 3e, from Dry Creek Quarry, N of Trussville, Alabama, U.S.A., lat. 33° 37' 30"N,
long. 86° 37' 30" W; grey shale with abundant goniatites and other macrofauna; 3341-3a and 3b from
2.9 m and 3341-3e from 2.2 m above base of unit. Nos. 3341-3c and 3d from Henson Creek, S of
Barton, Alabama, lat. 34°40'N, long. 87° 52' 28" W; thin bioclastic shaley limestone horizon with
bryozoans, crinoid debris, brachiopods and molluscs, 3.95 m above base of section. All from Pride
Mountain Formation, Chesterian, Mississippian, Carboniferous.
Explanation of Plate 18, 74
Figs. 1, 2, adult RV (3341-3e, 0.85 mm long): fig. 1. ext. lat.; fig. 2. int. lat. Fig. 3, adult LV (3341-3d, 0.9 mm long): int. lat. Fig. 4,
adult RV (3341-3c, 0.95 mm long): int. lat.
Scale A (250 /im; x60), figs. 1-4.
Stereo-Atlas of Ostracod Shells 18, 75
Mammoides dorsospinosus (3 of 4)
Diagnosis:
Remarks:
Distribution:
A ckn o w led gem en t:
Thick, tumid, semicircular, bilobate Mammoides. Dorsal margin straight, cardinal angles obtuse.
Posterodorsal corner slightly rounded, anterodorsal corner recurved. Ends evenly rounded,
maximum curvature of anterior end slightly below midheight; posterior with greater ventral swing,
maximum curvature at midheight. Shallow S2 at midlength between posteriorly curved spines of L2
and L3. Ventral lobe confluent with low, ridge-like LI, terminates below L3. Posterior spine at or
just above midheight. Surface reticulate. Inner lamella wide, narrows slightly to posterodorsal
corner. Tongue and groove hinge.
Three species of Mammoides Bradfield, 1935 possess a posterior spine at or above midheight: M.
dorsospinosus Sohn, 1961, M. longispina Green, 1963 and M. bouckaerti Bless & Massa, 1982. Sohn
described M. dorsospinosus from crushed specimens collected from the Upper Pennsylvanian of
central Texas, U.S.A. and Green (Bull. Res. Coun. Alb., 11, 72-75, 1963) described M. longispina
from the Lower Mississippian of Alberta, Canada. The difference between these species is the extreme
development of the dorsal and posterior spines in M. longispina. M. bouckaerti, from the Upper
Visean of the Rhadames Basin, Libya (Bless & Massa, Revue Inst. fr. Petrole, 37, no. 1, 26), differs
from both of the other species by its much more circular outline and more closely spaced dorsal spines.
The material described herein shows a wide inner lamella, which has not been described previously
for the genus. Shape variations in M. dorsospinosus from Alabama may be observed in length :
height ratio and the development of the ventral lobe. Shape variations in Mammoides have also been
noted by Green (1963) through a 400 m interval of the Banff Formation in Alberta and also within
individual samples, therefore making it unlikely that the variants represent sexual dimorphs.
Brownwood Shale, Canyon Group, Missourian, Upper Pennsylvanian, Central Texas; Pride
Mountain Formation, Chesterian, Mississippian, Black Warrior Basin, Alabama, U.S.A.
We acknowledge the financial support given by the Donors of the Petroleum Research Fund
administered by the American Chemical Society; the Mississippi Mineral Resources Institute and
Mississippi State University.
Explanation of Plate 18, 76
Fig. 1, adult RV (3341-3b, 0.91 mm long): ext. lat. Fig. 2, adult LV (3341-3a, 0.9 mm long): ext. lat. Fig. 3, adult LV (specimen lost,
0.95 mm long): int. lat.
Scale A (250 yum; x70), figs. 1, 2; scale B (250 pm; x60), fig. 3.
Mammoides dorsospinosus (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 74
‘
Stereo-Atlas of Ostracod Shells 18 (19) 77-80 (1991) Qujingsia nonaculeata (1 of 4)
595.336.1 1 (113.333) (510 : 161.103.25): 551.351
ON QUJINGSIA NONACULEATA HANSCH & WANG gen. et sp. nov.
by Wolfgang Hansch & Wang Shang-qi
(University of Greifswald, Germany
& Nanjing Institute of Geology & Palaeontology, People’s Republic of China)
Derivation of name:
Diagnosis:
Remarks:
Genus QUJINGSIA gen. nov.
Type-species: Qujingsia nonaculeata sp. nov.
After Qujing, name for the district of the type locality.
Small Beyrichiidae with a fused lobation and a low, narrow, flange-like uninterrupted velar
structure extending between cardinal corners in both dimorphs. Crumina assimulated within the
lobal area, without any distinct striate ornament ventrally. No zygal arch.
This genus shows similarity to the Baltoscandian Bingeria, an atypical beyrichiine genus, but
differs especially in the shape of the crumina and in the lack of striate subcruminal ornament (cf.
Martinsson, A., 1962, Bull. geol. Instn Univ. Uppsala , 41). Furthermore, Qujingsia lacks a distinct
zygal arch. Qujingsia can be distinguished from the several subgenera of Beyrichia by the cuspidal
morphology of its syllobium, its cruminal morphology and the lack of distinct subcruminal (striate)
ornament and by its very poor development of lobal ornamental features. The flange-like reduction
of the velum in Qujingsia indicates relationships to the wellerielliide genera which lack traces of a
velar structure (cf. Abushik, A.F. in: Abushik et al., Palaeozoic ostracodes from key sections in
the European part of the U.S.S.R., 7-133, 1971, Nauka, Moscow).
Explanation of Plate 18, 78
Figs. 1, 3, 9 car. (holotype, NIGPAS 115620, 820 /tm long): fig. 1, LV, ext. lat.; fig. 3, RV, ext. lat. Fig. 2, 9 car., ext. vent. (NIGPAS
115622, 880 pm long). Fig. 4, 9 car., ext. dors. (NIGPAS 115624, 850/rm long). Fig. 5, o * car., LV, ext. lat. (NIGPAS 115621,
890 /urn long).
Scale A (200 gm; x65), figs. 1, 3; scale B (200 /rm; x60), figs. 2, 4, 5.
Stereo-Atlas of Ostracod Shells 18, 79
Qujingsia nonaculeata (3 of 4)
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Qujingsia nonaculeata sp. nov.
Nanjing Institute of Geology and Palaeontology, Academia Sinica, People’s Republic of China,
no. NIGPAS 115620, 9 carapace.
Liaojiao Mountain, Qujing district, Yunnan Province, People’s Republic of China, approx, lat.
25°50'N, long. 103° 7' E, Miaokao Formation, Ludlow to Pridoli Series, Upper Silurian.
Latin non and aculeatus, prickly; because of the relatively smooth valve surface.
Nanjing Institute of Geology and Palaeontology, People’s Republic of China, nos. NIGPAS
115620 (holotype, 9 car.: PI. 18, 78, figs. 1, 3), NIGPAS 115621 (cr car.: PI. 18, 78, fig. 5; PI. 18,
80, fig. 4), NIGPAS 115622 (9 car.: PI. 18, 78, fig. 2; PI. 18, 80, fig. 3), NIGPAS 115623 (tecno-
morphic car.: PI. 18, 80, figs. 1, 2), NIGPAS 115624 (9 car.: PI. 18, 78, fig. 4) and NIGPAS
115625 (9 car.: PI. 18, 80, figs. 5-7). All from sample YQM-33 of the type locality.
The crumina is ellipsoid-like, not reaching the anterior end of the valve. On the syllobium and
sometimes also very weakly developed on the anterior lobe a faint, curved cuspidal ridge is
discernible.
In tecnomorphs the preadductorial node in front of a sometimes pit-like adductorial sulcus is
mostly completely fused with the very weakly developed anterior lobe. The valve surface is
generally smooth; however, in some individuals a very sparse reticulation/reticulostriation or
punctation can be traced, especially on the crumina. The velar edge is gently curved along the base
of the crumina and is nearly parallel to the marginal structure.
Known only from the type area. Kuanti and Miaokao formations, Ludlow to Pridoli Series, Upper
Silurian.
Explanation of Plate 18, 80
Figs. 1, 2, tecnomorphic car. (NIGPAS 115623, 740 //m long): fig. 1, RV, ext. lat.; fig. 2, ext. vent. Fig. 3, 9 car., detail of ventral side
(NIGPAS 115622). Fig. 4, o- car., RV, ext. lat. (NIGPAS 115621). Figs. 5-7, 9 car. (NIGPAS 115625, 880 jum long): fig. 5, RV,
detail of adductorial sulcus; fig. 6, RV, ext. lat.; fig. 7, RV, detail of crumina.
Scale A (150//m; x75), figs. 1, 2; scale B (50/tm; x240), figs. 3, 5, 7; scale C (200 //m; x60), figs. 4, 6.
Stereo-Atlas of Ostracod Shells 18, 78 Qujingsia nonaculeata (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (20) 81-84 (1991) Daleiella corbuloides (1 of 4)
595.337.21 (113.331) (420 : 162.003.52): 55.351 + 552.54
ON DALEIELLA CORBULOIDES (JONES & HOLL)
by Robert F. Lundin & Lee E. Petersen
(Arizona State University, Tempe & Anardarko Petroleum Corporation, Houston, U.S.A.)
Genus DALEIELLA Boucek, 1937
Type-species (by original designation): Cythere corbuloides Jones & Holl, 1869
Diagnosis: Large, strongly inequivalved Pachydomellidae(?) with a perimarginal carina on the posteroventral, extending to the ventral
surface and even to the anteroventral surface in some species, part of the admarginal surface of one or both valves. Overreach
of the right by the left valve strong around entire periphery of the right valve, but especially strong dorsally and posteroventrally.
Hinge parallel to longitudinal axis of valve. Shell wall without tubules or pore canals. Carapace width greater than height.
Dimorphic)?) by posteriorward displacement of maximum width in heteromorph (presumed female).
Remarks: This genus is similar in general appearance to pachydomellids but the combination of characters including the perimarginal
carinae, strong overreach, great carapace width, orientation of the hinge and absence of tubules and pore canals distinguish it
from the most similar genera Microcheilinella Geis, 1933, Tubulibairdia Swartz, 1936 and Newsomites Morris & Hill, 1952.
The genus is represented by the type-species, D. corbuloides (Jones & Holl, 1869), from the Silurian of Great Britain and
Podolia and the following species from the Silurian of Podolia and/or the East Baltic area: D. acutifinis Neckaja, 1960, D.
acutifiniformis = M. acutifiniformis Abushik, 1979, and D. ianica Neckaja, 1960. The existence of the diagnostic generic charac-
ters in D. triangularis Boucek, 1937 have not been verified and D. americana Morris & Hill, 1952 is most probably a Krausella
Ulrich, 1894. D. ? canadensis Copeland, 1962, D. caleyi Copeland, 1973 and D. deubeli Zagora, 1967 have little in common with
the type-species and certainly do not belong to Daleiella.
The placement of Daleiella and other similar genera without tubules in the Pachydomellidae Berdan & Sohn, 1961 is open to
question and would necessitate redefinition of that family. Discussion of suprageneric systematics of this large group of superfi-
cially similar genera is beyond the scope of this paper. It is clear, however, that if the presence of tubules is considered to be of
less than familial significance, Daleiella can be placed with Tubulibairdia, Microcheilinella, Newsomites and others in one family
of the Bairdiacea.
Explanation of Plate 18, 82
Figs. 1-3, car. (ASU X-128, 959 pm long): fig. 1, ext. rt. lat.; fig. 2, ext. post.; fig. 3, ext. dors. Fig. 4, car., ext. It. lat. (ASU X-129,
1071 pm long).
Scale A (200 pm; x53), figs. 1-3; scale B (200 pm; x48), fig. 4.
Stereo-Atlas of Ostracod Shells 18, 83
Daleiella corbuloides (3 of 4)
Daleiella corbuloides (Jones & Holl, 1869)
1869 Cythere corbuloides sp. nov. T.R. Jones & H.B. Holl, Ann. Mag. nat. Hist., (4), 3, 211, pi. 15, figs. 4, 5.
1887 Xestoleberis corbuloides (Jones & Holl); T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 410.
1892 Xestoleberis corbuloides (Jones & Holl); J. Smith, Trans, nat. Hist. Soc. Glasg., 3, 158.
1934 Microcheilinella corbuloides (Jones & Holl); R. S. Bassler & B. Kellett, Spec. Pap. geol. Soc. Am., 1, 412.
1937 Daleiella corbuloides (Jones & Holl); B. Boucek, Mem. Soc. r. Sci. Boheme, 1936(2), 7.
1952 Daleiella corbuloides (Jones & Holl); R. W. Morris & B. W. Hill, Bull. Am. Paleont., 34, 13.
1960 Tubulibairdia? corbuloides (Jones & Holl); I.G. Sohn, Prof. Pap. U.S. geol. Surv., 330-A, 75.
1961 Daleiella corbuloides (Jones & Holl); R.H. Shaver, Treatise on Invertebrate Paleontology, Q, 385, fig. 310A, 5.
1979 Microcheilinella mukschensis sp. nov. A.F. Abushik, Ezheg. uses paleont. Obshch., 22, 51, pi. 3, figs. 1, 2.
1991 Daliella [s/c] corbuloides (Jones & Holl); R.F. Lundin, L.E. Petersen & D.J. Siveter, /. Micropalaeont., 9 (2 for 1990), 179, pi. 1, figs. 5, 6.
Lectotype:
Type locality:
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Acknowledgements:
Designated herein. British Museum (Nat. Hist.) no. 12058; adult carapace. Jones & Holl 1869, pi. 15, figs. 4a-e.
“Croft’s Quarry,” 0.5 km W of Malvern, Hereford & Worcester, England; approximately Nat. Grid Ref. SO 757 464, lat.
52°08'N, long. 2°18'W. Much Wenlock Limestone Formation, Wenlock Series, Silurian.
Department of Geology, Arizona State University (ASU), nos X-128 (PI. 18, 82, figs. 1-3), X-129 (car.: PI. 18, 82, fig. 4), X-195
(car.: PI. 18, 84, fig. 6), X-196 (car.: PI. 18, 84, fig. 5). British Museum (Nat. Hist.), no. 12058 (lectotype, car.: PI. 18, 84, figs. 1-4).
The lectotype and ASU X-195 are topotypes. ASU X-128 and X-129 are from Harley Hill (A458) road cutting, 1.2 km NW of
Much Wenlock, Shopshire, England (Nat. Grid Ref. SJ 6103 0036) and ASU X-196 is from an exposure along path just above
old railway track, S side of River Severn and about 400 m W of Browers Brook, Benthall Edge, Shropshire (Nat. Grid Ref. SJ
6635 0355); all three from Farley Member, Coalbrookdale Formation, Wenlock Series.
Daleiella with a short perimarginal carina on the posteroventral admarginal surface of both valves and on the anteroventral
admarginal surface of the right valve. Surface smooth.
Possible dimorphism in this species is suggested by differences in the position of greatest width (compare PI. 18, 84, figs. 5, 6).
Data are not available at this time to demonstrate if this difference is due to normal variation or dimorphism because large single-
sample populations have not been found.
Comparison of Abushik’s (1979) figures and specimens she has supplied, with the British materials described here indicate that
D. corbuloides is conspecific with Microcheilinella mukschensis Abushik, 1979. This extends the known geographic distribution
of D. corbuloides, which heretofore was known only from the Welsh Borderland and English West Midlands (Lundin et al. 1991).
Known from many samples of Late Wenlock, Homerian, age and from one sample (locality no. 59 of Lundin et al. 1991) of
Ludlow, early Gorstian, age in the Welsh Borderland and English West Midlands, and from Late Wenlock stata of Podolia.
We gratefully acknowledge the support of NATO, the National Science Foundation (Grant No. EAR-8200816) and the College
of Liberal Arts and Sciences, Arizona State University.
Explanation of Plate 18, 84
Figs. 1-4, car. (lectotype, BMNHI2058, 1050/rm long): fig. 1, ext. rt. lat.; fig. 2, ext. vent.; fig. 3, ext. dors.; fig. 4, ext. It. lat. Fig. 5,
car., ext. vent. (ASU X-196, 1053 pm long). Fig. 6, car., ext. vent. (ASU X-195, 1053 pm long).
Scale A (200 pm; x48), figs. 1-4; scale B (200 pm; x49), figs. 5, 6.
Stereo-Atlas of Ostracod Shells 18, 84
Daleiella corbuloides (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 82
Daleiella corbuloides (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (21) 85-88 (1991) Villozona villosa (1 of 4)
595.336.14 (113.51) (430 : 161 .01 1 .50): 551.351 + 552.54
ON VILLOZONA VILLOSA (GRUNDEL)
by Gerhard Becker & Dieter Weyer
(University of Frankfurt-am-Main & Kulturhistorisches Museum, Magdeburg, Germany)
Genus VILLOZONA Grundel, 1965
Type-species (by original designation): Amphissites (Ectodemites) villosus Grundel, 1961
Diagnosis: Non-lobate kirkbyid genus with (principally) evenly convex lateral surface. Inner carina can be
present or absent; incomplete, weak dorsal ridge may be developed, but no distinct dorsum present.
Carapace surface smooth or finely reticulate to irregularly striate. Adductorial pit defined or lacking.
Distribution: Europe, Asia, N Africa; Lower Devonian-Lower Carboniferous.
Villozona villosa (Grundel, 1961)
1961 Amphissites (Ectodemites) villosus sp. nov. J. Grundel, Freiberger-ForschHft. , Clll, 87, 88, pi. 2, figs. 3-6.
1965 Villozona villosa (Grundel); J. Grundel, Freiberger-ForschHft., 082, 60.
1987 Villozona villosa (Grundel); G. Becker, Palaeontographica, A200, 60 (q.v. for full synonymy).
Holotype: Geological Institute, “Bergakademie Freiberg, Sachsen”, Germany, no. 21/21; an adult, silicified
left valve.
Type locality: Quarry “Pfaffenberg NE”, near Obernitz village, 3 km SSE Saalfeld, E Thiiringisches Schieferge-
birge, Germany; lat. 50° 38' N, long. 1 1 ° 24' W. Nodule bearing cephalopod limestones; Gattendorfia
stage, Lower Carboniferous. Pelagic facies, ostracod fauna of Thuringian ecotype.
Explanation of Plate 18, 86
Figs. 1, 2, adult LV (topotype, SMF Xe 15144, 1500 pm long): fig. 1, ext. lat.; fig. 2, detail of ext. lat. surface. Fig. 3, adult LV, dors.
obi. (SMF Xe 15145, 1380 /um long). Fig. 4, juv. LV, ext. lat. (topotype, SMF Xe 15146, 1150 pm long).
Scale A (300 pm; x50), figs. 1, 3, 4; scale B (100 /tm; x87), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 87
Villozona villosa (3 of 4)
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Forschungs-Institut Senckenberg, Frankfurt am Main (SMF), Federal Republic of Germany, nos.
SMF Xe 15144 (adult LV: PI. 18, 86, figs. 1, 2), SMF Xe 15145 (adult LV: PI. 18, 86, fig. 3; PI.
18, 88, fig. 6), SMF Xe 15146 Guv. LV: PI. 18, 86, fig. 4), SMF Xe 15147 Guv. car.; PI. 18, 88, fig.
1), SMF Xe 15148 Guv. RV: PI. 18, 88, fig. 2), SMF Xe 15149 Guv. LV: PI. 18, 88, fig. 3), SMF
Xe 15150 (adult LV: PI. 18, 88, fig. 4), SMF Xe 15151 Guv. car.: PI. 18, 88, fig. 5).
All except one of the figured specimens are topotype material; specimen SMF Xe 15145 is from
Quarry “Pfaffenberg SE”, Gattendorfia stage, Lower Carboniferous.
Villozona species with narrow to wide inner carina. Adductor spot obscure.
Two subspecies are distinguished by means of the inner carina. The typical V. villosa villosa
(Grundel, 1961), is characterized by a very wide inner carina and an ornamented carapace surface;
it occurs in the early Lower Carboniferous. All of the figured specimens belong to this subspecies.
Villozona villosa praecursor Bartzsch & Weyer, 1980, (Abh. Ber. Naturk. Vorgesch., 12(2), 43,
figs. 5.1-5), which is abundant in the upper Famennian (do V-VI; Upper Devonian) of Europe and
N Africa, has a narrow inner carina and a smooth carapace surface.
The late Palaeozoic to early Triassic Carinaknightina Sohn, 1970 shows a distinct dorsal carina.
Devonian and Carboniferous “ Kummerowia ” species (in litt.) belong to the “ Villozona line”
(Weyer, D. & Becker, G., Senckenberg leth., 71, 221, 1991).
V. villosa (sensu lato) is considered to have been a nectobenthic species. The dorsoterminal spines
(of juveniles) are thought to be biotope indicative features characteristic of low-energy palaeo-
environments.
V. v. villosa occurs in Central Europe; Gattendorfia stage. Lower Carboniferous.
Explanation of Plate 18, 88
Fig. 1, juv. car., dors, (topotype, SMF Xe 15147, 1120 pm long). Fig. 2, juv. RV, vent, (topotype, SMF Xe 15148, 1150/tm long).
Fig. 3, juv. LV, int. lat. (topotype, SMF Xe 15149, 1150//m long). Fig. 4, adult LV, post, (topotype, SMF Xe 15150, 1450/tm long).
Fig. 5, juv. car., vent, (topotype, SMF Xe 15151, 1270//m long). Fig. 6, adult LV, vent. (SMF Xe 15145, 1380 /tm long).
Scale (300 //m; x50), figs. 1-6.
Stereo-Atlas of Ostracod Shells 18, 86
Villozona villosa (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 88 Villozona villosa (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (22) 89-92 (1991) Huntonella bransoni (1 of 4)
595.336.11 (113.42) (766:162.097.34): 551.351 + 552.54
ON HUNTONELLA BRANSONI LUNDIN
by David J. Siveter & Robert F. Lundin
(University of Leicester, England & Arizona State University, Tempe, U.S.A.)
Genus HUNTONELLA Lundin, 1968
Type-species (by original designation): Huntonella bransoni Lundin, 1968
Diagnosis: Tuberculate Amphitoxotidinae in which both the velar edge and the torus cross the crumina without
any deflection or interruption. Velum wide, continues precruminally, is abruptly restricted postero-
ventrally, has a prominent border crest. Syllobium, preadductorial node and anterior lobe are well
differentiated, have lowly elevated connections, lack cusps.
Remarks: An entire and unmodified velar edge across the crumina is also a characteristic of the amphitoxo-
tidines Dibolbina Ulrich & Bassler, 1923 (Md geol. Surv., Silurian volume), Berolinella Martinsson,
1962 (Bull. geol. Instn Univ. Uppsala, 41) and Tropidotoxotis Siveter, 1980 (Palaeontogr. Soc.
[Monogr.], 133, (No. 556), for 1979). Huntonella differs from Dibolbina in having no basal crest,
crista or entire velum; from Berolinella (see Hansch, W. & Siveter, D.J., Stereo-Atlas Ostracod
Shells, 16, 106-111, 1989) in lacking a basal crest and in having a border crest; from Tropidotoxotis
in having a torus and a border crest; and from all three genera in details of lobal morphology and
ornament. Furthermore, the cruminal part of the velar edge in Huntonella is particularly wide in
lateral view.
Explanation of Plate 18, 90
Figs. 1-3, 9 LV (OU 5923, 1575 /an long): fig. 1, ext. lat.; fig. 2, ant.; fig. 3, vent. Figs. 4-6, tecnomorphic RV (OU 5922c, 1200 /im
long): fig. 4, vent.; fig. 5, ant.; fig. 6, ext. lat.
Scale A (300 /^m; x30), figs. 1-3; scale B (300 /tm; x40), figs. 4-6.
Stereo-Atlas of Ostracod Shells 18, 91
Huntonella bransoni (3 of 4)
Distribution: Henryhouse Formation, Oklahoma (Lundin, Bull. Okla geol. Surv., 108, 1965; 1968 op. cit.);
Ludlow/Pfidoli series, Silurian. Haragan Formation, Oklahoma (Lundin 1968) and Rockhouse and
Birdsong formations, western Tennessee (Lundin & Petersen 1974; Petersen & Lundin, Okla geol.
Surv., in press)-, Gedinnian, Devonian.
Huntonella bransoni Lundin, 1968
1968 Huntonella bransoni n. sp. R.F. Lundin, Bull. Okla. geol. Surv., 116, 22, pi. 1, figs. la-k.
1974 Huntonella bransoni Lundin; R.F. Lundin & L.E. Petersen, J. Paleont., 48, 242, pi. 1, figs. 11-13.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Remarks:
Distribution:
Ackno w led gem ent:
Oklahoma University, Norman, no. OU 5921; female right valve.
Near old Hunton townsite, Coal County, Oklahoma; approx, lat. 34° 30' N, long. 96° 30' W; 166 feet
above base of Haragan Formation, Devonian.
Oklahoma University, nos. OU 5921 (holotype, 9 RV: PI. 18, 92, figs. 1-3), OU 5922b (tecno-
morphic RV: PI. 18, 92, figs. 4, 5), OU 5922c (tecnomorphic RV: PI. 18, 90, figs. 4-6), OU 5923 (9
LV: PI. 18, 90, figs. 1-3). All from between 155 and 212 feet above base of the Haragan Formation
at the type section, Oklahoma.
Huntonella species with an abruptly restricted velum posteroventrally.
H. bransoni differs from Huntonella fittsi (Roth, 1929), from the Henryhouse Formation of
Oklahoma, in details of velar and lobal morphology.
Haragan Formation, Oklahoma (Lundin 1968) and Rockhouse and Birdsong formations, western
Tennessee (Lundin & Petersen 1974; Petersen & Lundin in press)-, Gedinnian, Devonian.
The authors wish to thank NATO for its support for their collaborative research programme.
Explanation of Plate 18, 92
Figs. 1-3, 9 RV (holotype, OU 5921, 1500 pm long): fig. 1, ant.; fig. 2, ext. lat.; fig. 3, ornament on crumina. Figs. 4, 5, tecnomorphic
RV (OU 5922b, 975 /rm): fig. 4, ant.; fig. 5, ext. lat.
Scale A (300 pm\ x35), figs. 1, 2; scale B (50 pm; xl60), fig. 3; scale C (300 /^m; x50), figs. 4, 5.
Huntonella bransoni (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 92
Stereo- Atlas of Ostracod Shells 18, 90
Huntonella bransoni (4 of 4)
Stereo- Atlas of Ostracod Shells 18 (23) 93-96 (1991) Paracathaycythere costaereticulata (1 of 4)
595.337.14 (119.9) (510 : 161 . 1 19.34): 551.351
ON PARACA THA YCYTHERE COSTAERETICULA TA
WHATLEY & ZHAO gen. et sp. nov.
by Robin C. Whatley & Zhao Quanhong
(University College of Wales, Aberystwyth, UK & Tongji University, Shanghai, China)
Genus PARACATHA YCYTHERE gen. nov.
Type species: Paracathaycythere costaereticulata sp. nov.
Derivation of name: Gr. napa = near + Cathaycy there.
Diagnosis: Medium-sized; thin-shelled, elongate and subrectangular in lateral view, greatest height at blunt anterior cardinal angle
and greatest length subventrally; dorsal margin straight, slightly oblique; ventral margin with slight oral concavity;
anterior margin broadly rounded; posterior margin with truncated posterodorsal slope and narrowly rounded postero-
ventral slope; posterior cardinal angle distinct. LV slightly overlaps RV at anterior cardinal angle and along posterdorsal
slope of posterior margin. Surface reticulate and costate or tuberculate/papillate with costae, always with prominent rib
extending from the ocular region diagonally towards the anteroventral margin. Sub-central tubercle and its surrounding
sulcus feebly developed, best seen interiorly; anterior margin finely or moderately denticulate; coarser denticules line the
posteroventral margin; mid-posterior margin with distinct flange. Eye tubercle weakly present or indistinct. Sexual
dimorphism distinct, male more slender than female. Hinge weak, modified amphidont. In RV the terminal short
elements are slightly denticulate teeth, the anteromedian locule is shallow while the posteromedian groove is locellate
and open antero-ventrally; in LV each terminal socket has a rounded anti-slip toothlet ventrally, the club-like antero-
median element is slightly crenulate with a cusp at its anteriormost end. Inner lamella wide and avestibulate; radial pore
canals long, fine, simple and few; selvage peripheral except at posterior margin where it is directed inwards away from
the outer margin. Adductor scars small, in a vertical row of 4 closely-packed scars; frontal scar single, heart-shaped.
Explanation of Plate 18, 94
Fig. 1, 9 car., ext. lat. (paratype, OS 13743, 554 pm long); Fig. 2, o’ RV, ext. lat. (paratype, OS 13744, 585 pm long); Fig. 3, o’ car.,
ext. lat. (paratype, OS 13745, 585 pm long).
Scale A (200 /rm; x98); figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 95
Paracathaycythere costaereticulata (3 of 4)
Remarks: Paracathaycythere is somewhat similar to Cathaycythere Whatley & Zhao, 1987, Neosinocythere Huang, 1985,
Sinocythere Hou, 1982, and Spinileberis Hanai, 1961 in many features of carapace morphology. All share the widely
flared anterior margin, the very wide avestibulate anterior inner lamella and the small ventral tooth situated within the
posterior terminal socket of the LV hinge. Paracathaycythere is thinner-shelled than the other genera and also has a
much less strongly developed amphidont hinge. The genus is probably related to Hemikrithe Van den Bold, 1950 which
has a similar shape but lacks the same details of hingement. The ventral tooth in the terminal hinge element of the LV
is very similar to the “auriline” tooth of Aurila Pokorny, 1955 and its allies within the Hemicytheridae. That this is an
entirely coincidental resemblance is evidenced by other carapace characters, such as the muscle scars which clearly place
this genus in the Trachyleberididae although, by virtue of its similarity to Cathaycythere , Sinocythere and Spinileberis
it probably belongs to the Sinocytherinae Huang, 1985. Hemikrithe may also belong to this subfamily.
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Diagnosis:
Distribution:
Paracathaycythere costaereticulata sp. nov.
British Museum (Nat. Hist.), no. OS 13742, o’ car., subsequently split into RV and LV.
[Paratypes, British Museum (Nat. Hist.), nos. OS 13743-6].
Lianyungang Harbour, Jiangsu Province, on the Yellow Sea cost, approx, lat. 34°44'N, long. 1 19°23'E; silty fine sand,
intertidal zone, Recent.
L. costae = ribs + reticulata , referring to the nature of ribs and reticulae which constitute the ornament of this species.
British Museum (Nat. Hist.) nos. OS 13743 (paratype, 9 car.: PI. 18, 94, fig. 1), OS 13744 (paratype, o’ RV: PI. 18, 94,
fig. 2), OS 13745 (paratype, o’ car.: PI. 18, 94, fig. 3), OS 13746 (paratype, o’ car.: PI. 18, 96, fig. 1), OS 13742
(holotype, o"LV: PI. 18, 96, fig. 2), OS 13742 (holotype, o’ RV: PI. 18, 96, fig. 3); all are from the type locality.
As for genus but with ornament of ribs and reticulae. Apart from the oblique ocular rib characteristic of the genus, a
diagonal median rib extends across the carapace from the posterodorsal to anteroventral, being interrupted by the weak
subcentral tubercle. A ventrolateral rib extends approximately parallel to the ventral margin to its anterior union with
the median rib. A short rib extends towards the median rib from the posterodorsal loop. Intercostal area irregularly
reticulate; solae of reticulae finely punctate.
Pleistocene to Recent, eastern China. Living specimens are found in the littoral zone along Chinese coasts from approx,
lat. 18° to 35°N, in salinity of 32 to 35%0, and on silt to medium sand.
Explanation of Plate 18, 96
Fig. 1, o* car. dors, (paratype, OS 13746, 585 pm long); Fig. 2, cr LV, int. lat. (holotype, OS 13742, 615yum long); Fig. 3, o* RV, int.
lat. (holotype, OS 13742, 615 pm long).
Scale A (200 pm\ x98); figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 94
Paracathaycythere costaereticulata (4 of 4)
Paracathaycythere costaereticulata (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 96
Stereo-Atlas of Ostracod Shells 18 (24) 97-100 (1991) Paracathaycythere scabra (1 of 4)
595.337.14 (119.1) (510 : 161.1 14.20): 552.51 + 52
ON PARACATHAYCYTHERE SCABRA ZHAO & WHATLEY sp. nov.
by Zhao Quanhong & Robin C. Whatley
(Tongji University, Shanghai, China & University College of Wales, Aberystwyth, UK)
Paracathaycythere scabra sp. nov.
British Museum (Nat. Hist.) no. OS 13747, 9 right valve.
[Paratypes: British Museum (Nat. Hist.) nos. OS 13748-51].
Shelf hole ZQ3, lat. 20° 58.05' N, long. 114° 30.00' E, water depth 89 m, off Guangdong Province,
northern part of the South China Sea; core samples from hole depths of 54.0-54.1, 92.1-92.2 and
96.3-96.9 m, silt and fine sand, mid-Pleistocene.
L. scabra = rough, with reference to the roughly tuberculate surface ornament of this species.
British Museum (Nat. Hist) no. OS 13747 (holotype, 9 RV: PI. 18, 98, fig. 1), OS 13748 (paratype,
9 LV: PI. 18, 98, fig. 2), OS 13749 (paratype, o* LV: PI. 18, 98, fig. 3), OS 13750 (paratype, 9 LV:
PI. 18, 100, fig. 1), OS 13751 (parataype, o* RV: PI. 18, 100, fig. 2); all are from the type locality.
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Explanation of Plate 18, 98
Fig. 1, 9 RV, ext. lat. (holotype, OS 13747, 533 nm long); Fig. 2, 9 LV, ext. lat. (paratype, OS 13748, 514/rm long); Fig. 3, o* LV,
ext. lat. (paratype, OS 13749, 562 /rm long).
Scale A (100 /xm; xl05), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 99
Paracathaycythere scabra (3 of 4)
Diagnosis: As for the genus (see R. C. Whatley & Zhao Quanhong, Stereo-Atlas Ostracod Shells 18, 93-96,
1991) but with an ornament of dense, rather irregularly distributed tubercles and papillae. Some of
these are conical, others are spatulate and others, particularly dorsally are castellate; most, if not
all, are penetrated by a normal pore. A strong diagonal ocular rib is formed in front by aligned
tubercles.
Distribution: Only known from the type locality.
Explanation of Plate 18, 100
Fig. 1, 9 LV, int. lat. (parataype, OS 13750, 545 //m long); Fig. 2, o ■ RV, int. lat. (paratype, OS 13751, 562 /im long).
Scale A (100 /^m; xl73), figs. 1, 2.
Stereo-Atlas of Ostracod Shells 18, 98
Paracathaycythere scabra (2 of 4)
Paracathaycythere scabra (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 100
Stereo-Atlas of Ostracod Shells 18 (25) 101-104 (1991) Cocoonocythere sinensis (1 of 4)
595.337.14 (119.9+ 119.4) (510 : 161.121.29+ 161.1 19.33): 551.313.1 +552.52
ON COCOONOCYTHERE SINENSIS ZHAO
by Zhao Quanhong & Robin C. Whatley
(Tongji University, Shanghai, China & University College of Wales, Aberystwyth, UK)
Genus COCOONOCYTHERE Zhao, 1984
Type-species: Cocoonocythere sinensis Zhao, 1984
1984 Cocoonocythere gen. nov. Zhao Quanhong, Mar. Geol. & Quatern. Geol., 4, 53.
Diagnosis: Small; ovate in lateral view with greatest height posteromedianly and greatest length at mid-height;
dorsal margin slightly arched; ventral margin parallel and gently concave medially; anterior margin
broadly rounded; posterior margin bluntly rounded in female and narrowly rounded in male. Cara-
pace inflated, ovate in dorsal view with convex lateral margins, narrowly rounded anterior and
bluntly rounded posterior ends. Sexual dimorphism distinct, male slimmer than female. External
surface smooth and internal surface finely and densely pitted. Hinge lophodont: terminal teeth in
right valve small and simple, long median groove smooth and open ventrally. Inner lamella very
narrow, avestibulate; radial pore canals short, straight and simple, moderate in number. Adductor
scars large, in a vertical row of 4 with a large V-shaped frontal scar.
Remarks: Cocoonocythere is somewhat similar in its small size, hinge and muscle scars, to Microcythere G. W.
Muller, 1984, but differs in its carapace being rounded anteriorly and posteriorly and in its very narrow
inner lamella. Given its large V-shaped frontal scar and narrow inner lamella it is readily distinguished
Explanation of Plate 18, 102
Fig. 1, 9 RV, ext. lat. (OS 13711, 339 pm long); Fig. 2, o • LV, ext. lat. (OS 13712, 342 /ym long); Fig. 3, 9 LV, ext. lat. (OS 13713,
342 long).
Scale A (100 pm; xl80); figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 103
Cocoonocythere sinensis (3 of 4)
from other smooth genera common in the coastal zone such as Cobanocythere Hartmann, 1959,
Cytherois G. W. Muller, 1884, Paracytherois G. W. Muller, 1894 and Paracytheroma Juday, 1907.
Cocoonocythere sinensis Zhao, 1984
1982 Gen. et sp. 2, Hou et al., in Hou Youtang et at., Cretaceous-Quaternary ostracode fauna from Jiangsu, Geol. Publ. House
(Beijing), 245, 246, pi. 88, figs. 21-23.
1984 Cocoonocythere sinensis gen. et sp. nov. Zhao Quanhong, Mar. Geol. & Quatern. Geol., 4(1), 53, text-fig. 4; pi. 2, figs. 15-21.
1985 Cocoonocythere sinensis Zhao; Zhao Quanhong, Acta oceanol. sin., 7(2), 196-199, pi. 1, fig. 15.
1985 Cocoonocythere sinensis Zhao; Wang et al., in Wang Pinxian et al., Marine Micropaleontology of China, China Ocean Press
& Springer-Verlag, pi. 30, fig. 12.
Holotype:
Type locality:
Figured specimens:
Diagnosis:
Distribution:
Department of Marine Geology, Tongji University, Shanghai, China; no. T6204, 9 carapace (not
figured herein).
Xiangshangang Bay, Xidian, Zhejiang Province, China, approx, lat. 29°22'N, long. 121°27'E;
supratidal pool, grey silt. Recent.
British Museum (Nat. Hist.) nos. OS 13711 (9 RV: PI. 18, 102, fig. 1), OS 13712 (o* LV: PI. 18, 102,
fig. 2), OS 13713 (9 LV: PI. 18, 102, fig. 3), OS 13714 (9 car.: PI. 18, 104, fig. 1), OS 13715 (9 LV:
PI. 18, 104, fig. 2), OS 13716 (9 RV: PI. 18, 104, fig. 3). No. OS 13714 is from the type locality and
others are from Jianhu County, Jiangsu province, East China, approx, lat. 33°26'E, long.
119°46'N, Holocene, grey mud.
As for the genus.
Pleistocene to Recent, Jiangsu and Zhejiang Provinces, East China. Living specimens are found in
bottom samples of mud or silt of supratidal pools, channels, marshes and littoral flats, with water
salinity ranging from 5 to 32%0.
Explanation of Plate 18, 104
Fig. 1, 9 car. ext. dors. (OS 13714, 328 pm long); Fig. 2, 9 LV, int. lat. (OS 13715, 339 pm long); Fig. 3, 9 RV, int. lat. (OS 13716,
339 pm long).
Scale A (100 /rm; xl80); figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 102
Cocoonocythere sinensis (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 104
Cocoonocythere sinensis (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (26) 105-108 (1991) Polydontoconcha hyperdonta (1 of 4)
595.337.14 (119.1) (510 : 161. 1 14.20): 551.35 + 552.52
ON POLYDONTOCONCHA HYPERDONTA
ZHAO & WHATLEY gen. et sp. nov.
by Zhao Quanhong & Robin C. Whatley
(Tongji University, Shanghai, China & University College of Wales, Aberystwyth, UK)
Genus POLYDONTOCONCHA gen. nov.
Type-species: Polydontoconcha hyperdonta sp. nov.
Derivation of name: Gr. nokkrj = many + Sovra = teeth + Kovyrj = shell.
Diagnosis: Carapace resembling that of Palmoconcha Swain & Gilby, 1974. Elongate to ovate in lateral view;
dorsal cardinal angles distinct, especially the posterior; dorsal margin straight; ventral margin
subparallel but slightly concave anteromedianly and convex posteromedianly; anterior margin
obliquely rounded; posterior margin with short caudal process medianly and truncated above. Eye
spot indistinct. Surface with fine punctae medianly and weak parallel concentric ribs peripherally.
Sexual dimorphism distinct with male more elongate than female. Hinge modified gongylodont: in
the LV the anterior terminal element comprises 3 prominent cubic teeth; median element a strongly
locellate groove; posterior terminal element a reniform socket enclosing an elongate tooth. RV
hinge with complementary structures; anterior elongate socket with 3 deeper and ventrally open
loculi. Inner lamella of moderate width with vestibulae at each end; radial pore canals few, short,
straight and simple. Adductor scars in a semicrescentic row of 4; frontal scar single, heart-shaped.
Explanation of Plate 18, 106
Fig. 1, cr RV, ext. lat. (paratype, OS 13718, 536 pm long); Fig. 2, 9 RV, ext. lat. (paratype, OS 13719, 505 pm long); Fig. 3, 9 car.,
dors, (paratype, OS 13720, 559 pm long).
Scale A (200 y«m; xllO), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 107
Polydontoconcha hyperdonta (3 of 4)
Remarks: This genus clearly belongs to the family Loxoconchidae on the basis of its modified gongylodont
hinge and “loxoconchid” shape. It differs from Palmoconcha in its additional anterior terminal
teeth and locellate median element in the LV. The genus is monotypic.
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Diagnosis:
Distribution:
Polydontoconcha hyperdonta sp. nov.
British Museum (Nat. Hist.) no. OS 13717, 9 left valve.
[Paratypes: British Museum (Nat. Hist.) nos. OS 13718-22].
Shelf hole ZQ3, lat. 20° 58.05' N, long. 1 14° 30.00' E, water depth 89 m, off Guangdong Province,
northern part of the South China Sea. Core samples from hole depth 96.0-96.9 m, grey silt, mid-
Pleistocene.
Gr. V7 zep = beyond or very + <5ovra = teeth, because of the large number of teeth in the hinge of
this species.
British Museum (Nat. Hist.) nos. OS 13718 (paratype, a RV: PI. 18, 106, fig. 1), OS 13719
(paratype, 9 RV: PI. 18, 106, fig. 2), OS 13720 (paratype, 9 car.: PI. 18, 106, fig. 3), OS 13717
(holotype, 9 LV: PI. 18, 108, fig. 1), OS 13721 (paratype, 9 RV: PI. 18, 108, fig. 2), OS 13722
(paratype, o* LV: PI. 18, 108, fig. 3); all are from the type locality.
As for the genus.
Pleistocene to Recent, northern shelf of the South China Sea. Recent specimens were recovered
from two bottom samples of the South China Sea off Guangdong Province, water depth
180-220 m, fine sand.
Explanation of Plate 18, 108
Fig. 1, 9 LV, int. lat. (holotype, OS 13717, 514/zm long); Fig. 2, 9 RV, int. lat. (paratype, OS 13721, 555 pm long); Fig. 3, o' LV,
ext. lat. (paratype, OS 13722, 550/rm long).
Scale A (200 //m; xllO), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 108
Polydontoconcha hyperdonta (4 of 4)
Stereo-Atlas of Ostracod Shells 18, 106
Polydontoconcha hyperdonta (2 of 4)
Stereo-Atlas of Ostracod Shells 18 (27) 109-112 (1991) Palmoconcha ruggierii (1 of 4)
595.337.14 (118.22) (44 : 162.002.49 + 420 : 006.50): 552.51
ON PALMOCONCHA RUGGIERII MAYBURY sp. nov.
by Caroline A. Maybury
(University College Wales, Aberystwyth, UK)
Holotype :
Type locality:
Derivation of name:
Figured specimens:
Palmoconcha ruggierii sp. nov.
British Museum (Natural History), London, OS 13755; 9 LV.
[Paratypes nos. OS 13756-OS 13760].
Light grey, fine to medium grained sand, Le Bosq d’Aubigny (Manche), near St. Lo (approx, lat.
49°07'N, long. 01°05'W), NW France; Upper Pliocene, Redonian.
In honour of Professor G. Ruggieri, in recognition of his extensive research on Tertiary to Recent,
Mediterranean Ostracoda.
British Museum (Natural History) nos. OS 13755 (holotype, 9 LV: PL 18, 110, fig. 1), OS 13756
(paratype, 9 RV: PL 18, 110, fig. 2), OS 13757 (paratype, o* LV: PL 18, 110, fig. 3), OS 13758
(paratype, o* RV: PL 18, 112, fig. 1), OS 13759 (paratype, a RV: PL 18, 112, fig. 2), OS 13760
(paratype, cr LV: PL 18, 1 12, fig. 3). Paratypes OS 13756-OS 13758 are from the type locality, but
from a different sample; paratypes OS 13759 and OS 13760 are from a mixed sample (no. 7),
Vicarage Pit, St. Erth, Cornwall, England. See J.-P. Margerel, Les Foraminiferes du Redonien,
Systematique, Repartition stratigraphique, Paleoecologie, Nantes, 1, 8-26, 1968 and C.A.
Maybury, Taxonomy, Palaeoecology and Biostratigraphy of Pliocene Benthonic Ostracoda from
St. Erth and NW France, unpub. PhD thesis, Univ. Wales, 1, 3-6, 1985 for further details of the
French and British samples, respectively.
Explanation of Plate 18, 110
Fig. 1, 9 LV, ext. lat. (OS 13755, 480 pm long); Fig. 2, 9 RV, ext. lat. (OS 13756, 500 qm long); Fig. 3, cr LV, ext. lat. (OS 13757,
470 qm long).
Scale A (100 /^m; xl36), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 111 Palmoconcha ruggierii (3 of 4)
Diagnosis: A small, subelliptical species of Palmoconcha. Anterior margin asymmetrically rounded; posterior
margin almost caudate; dorsal margin arcuate in 9 LV, oblique in 9 RV and o * LV and slightly
concave medianly in o* RV. Ventral margin curved anteriorly and posteriorly, with very slight oral
incurvature. Smooth marginal rim narrowest orally. Lateral surface punctate with punctae
becoming smaller and more densely distributed peripherally. Pore conuli prominent just at the
anterior and posterior edges of the tumid region of each valve. Eye spot large and smooth; selvage
blade-like orally. Hinge and muscle scars typical of the genus.
Remarks: Ruggieri (pers. comm.) has compared the present material with a similar species: Palmoconcha
subrugosa (Ruggieri) (G. Ruggieri, Boll. Soc. paleont. ital., 15(2), 184, 1976) and concludes that
the two are different, especially with respect to their outlines; P. subrugosa being subrectangular
and P. ruggierii, subelliptical . The present species is also more inflated anteriorly and smaller:
males of P. subrugosa are 570-580 pm in length and the female 565 pm (see Loxoconcha aff. L.
agilis Ruggieri, 1967 (G. Bonaduce, G. Ciampo & M. Masoli, Pubbl. Staz. zool. Napoli, 40, 102,
1975) which has been assigned to P. subrugosa by Ruggieri {op. cit.)). Specimens of P. subrugosa
from the Italian Quaternary kindly sent by Ruggieri have enabled me to concur with this opinion.
Ruggieri notes that the males of P. ruggierii are similar to an unpublished species of Palmoconcha
from the late Miocene of Tunisia. Again the difference between the two is in the outline: the
unpublished species is higher anteriorly than P. ruggierii.
Aruta, 1966 (Riv. Miner. Sicil., 17, 203, pi. 1, fig. 8) described a Loxoconcha ruggierii from the
late Miocene of Sicily; this is a true Loxocorniculum, highly reticulate with posterodorsal and
posteroventral protuberances.
Distribution: In addition to its occurrence in two samples from the type locality, this species has also been found
in Redonian (Upper Pliocene) deposits from Apigne (Le Temple du Cerisier) and Palluau II, NW
France and late Pliocene deposits from St. Erth, England (sample nos. 1-3, 7, 10, 12-14, 16-17,
21, 23, 25-29).
Explanation of Plate 18, 112
Fig. 1, cr RV, ext. lat. (OS 13758, 480 qm long); Fig. 2, cr RV, int. lat. (OS 13759, 480 qm long); Fig. 3, O’ LV, int. muse. sc. (OS 13760,
470 //m long).
Scale A (100 /um\ X136), figs. 1, 2; scale B (10/vm; x640), fig. 3.
Palmoconcha ruggierii (4 of 4)
Palmoconcha ruggierii (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 112
Stereo-Atlas of Ostracod Shells 18, 110
Stereo-Atlas of Ostracod Shells 18 (28) 113-116 (1991) Ektyphocythere cookiana (1 of 4)
595.337.14 (116.14) (420 : 162.003.51 + 003.53): 551.351 +552.52
ON EKTYPHOCYTHERE COOKIANA (ANDERSON)
by Ian Boomer
(University of East Anglia, Norwich)
Ektyphocythere cookiana (Anderson, 1964)
1964 Klinglerella? cookiana sp. nov. F.W. Anderson, Bull. geol. Surv. Gt Br., 21, 143, pi. 9, figs. 16, 17, pi. 15, fig. 122.
Holotype: British Geological Survey (Keyworth) GSM Mik (j) 276001. 9 carapace.
Type locality: Plattlane Borehole, Whixall, Shropshire (Grid Ref. SJ 5140 3645). Westbury Formation, “Lower
Rhaetic’’, at a depth of 243 '0" to 243' 6".
Figured specimens: British Geological Survey (Keyworth) no. GSM Mik (j) 276001 (holotype, car.: PI. 18, 1 14, fig. 3).
Bristol City Museum and Art Gallery (BRSMG) no. Cel7020 (LV: PI. 18, 114, fig.l), Cel7021 (LV:
PL 18, 114, fig. 2), Cel7022 (car.: PI. 18, 114, fig. 4), Cel7023 (RV: PI. 18, 116, figs. 1, 2), Cel7024
(car.: PI. 18, 116, figs. 3, 4). All specimens (apart from holotype) are from uppermost bed of the
Westbury Formation, Penarth Group at Hampstead Farm Quarry, Avon (Grid Ref. ST 726 839)
( sensu D.T. Donovan et al.. Palaeontology, 32, 231, 1989); collected by M. T. Curtis, to whom
thanks are due for making them available to the author.
Explanation of Plate 18, 114
Fig. 1, LV, ext. lat. (BRSMG Cel7020, 500 pm long); Fig. 2, LV, int. lat. (BRSMG Cel7021, 462 pm long); Fig. 3, car., ext. It. lat.
(holotype, GSM Mik (j) 276001, 487 pm long); Fig. 4, car., ext. rt. lat. (BRSMG Cel7022, 538//m long).
Scale A (100 //m; X100), figs. 1-4.
Stereo-Atlas of Ostracod Shells 18, 115
Ektyphocythere cookiana (3 of 4)
Diagnosis:
Remarks:
Distribution:
Carapace small (LV > RV), inflated ventro-laterally, outline sub-oval in lateral view tapering to a
narrowly rounded posterior margin. Anterior margin well rounded. Both anterior and posterior
margins compressed, the former bearing 6-8 and the latter 3-4 short radial ribs. Greatest height in
front third of valve. External ornamentation comprises coarse longitudinal ribs on ventral and
ventro-lateral surfaces, lateral surfaces possess well developed coarse anastomosing ribs, forming a
weak reticulation. Hinge antimerodont, adductor muscle scars not visible, inner lamellae of
moderate width anteriorly and posteriorly, no vestibula observed, marginal pore canals appear few
and simple although indistinct. Sexual dimorphism not apparent although the holotype would
appear to be somewhat smaller than the material from Avon.
This species represents one of the earliest representatives of the genus Ektyphocythere Bate, 1963
C Bull . Br. Mus. nat. Hist. (Geol.) 8, 213). It appears to have a limited geographical and stratigraphical
distribution.
Rhaetian (late Triassic) of the English Midlands (Anderson, 1964) and Avon, SW England (this
study).
Explanation of Plate 18, 116
Fig. 1, RV, int. lat. (BRSMG Cel7023, 526pm long); Fig. 2, RV, detail of hingement (BRSMG Cel7023); Fig. 3, car., ext. vent.
(BRSMG Cel7024, 551 pm long); Fig. 4, car., ext. dors. (BRSMG Cel7024).
Scale A (100 ,«m; X100), fig. 1; scale B (100 pm\ xl45); scale C (100 pm; x95), figs. 3, 4.
Stereo-Atlas of Ostracod Shells 18, 114
Ektyphocythere cookiana (2 of 4)
Stereo-Atlas of Ostracod Shells 18, 116
Ektyphocythere cookiana (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (29) 117-120 (1991) Tethysobuntonia govoroffi (1 of 4)
595.337.14 (116.333.3) (662.6 : 162.016. 1 1): 551.35
ON TETHYSOBUNTONIA GOVOROFFI COLIN & BABINOT gen. et sp. nov.
by Jean-Paul Colin & Jean-Frangois Babinot
(Esso Rep, Begles & Universite de Provence, Marseille, France)
Genus TETHYSOBUNTONIA gen. nov.
Type-species: Tethysobuntonia govoroffi sp. nov.
Tethyan representative of the subfamily Buntoniinae.
A genus of the sub-family Buntoniinae with a distinct sub-central tubercle surrounded by a sub-
circular depression. Small denticles occur along the anterior margin. Ventral margin typically
depressed into selvage area, especially on the right valve. Carapace surface smooth or irregularly
pitted. Hinge amphidont (heterodont). Marginal zone moderately wide, without vestibulum. Very
pronounced sexual dimorphism with males being much larger and more elongate than females.
Tethysobuntonia differs from other representatives of the sub-family Buntoniinae as defined by
Apostolescu ( Revue Inst. fr. Petrole, 16(7-8), 1961), by the presence of a well developed sub-
central tubercle.
Tethysobuntonia govoroffi sp. nov.
1956 Eobuntonia? curta n. sp. A.S. Sayyab, Cretaceous Ostracoda from the Persian Gulf Area, Unpubl. Thesis, College State Univ.
Iowa, 109-111, text-fig. 2N, pi. 5, figs. 5, 11, 21.
1973 Buntonia sp. B 816, Y. Bellion, P. Donze & R. Guiraud, Pubis Serv. Carte geol. Alger., 44, 20, pi. 5, figs. 6-11.
1988 “ Eobuntonia cf. curta'’ Sayyab; J. Athersuch, in T. Hanai, N. Ikeya & K. Ishizaki (eds.), Evolutionary Biology of Ostracoda,
Elsevier, Amsterdam, 1189, 1191, 1197, pi. 1, fig. 17 only (non fig. 18).
Derivation of name:
Diagnosis:
Remarks:
Explanation of Plate 18, 118
Fig. 1, o* LV, ext. lat. (paratype, PCA 1/1, 559 pm long); Fig. 2, o* LV, int. lat. (paratype, PCA 1/2, 593 /rm long); Fig. 3, cr car. ext.
rt. lat. (holotype, HCA 1, 602 pm long).
Scale A (100 //m; x93), figs. 1-3.
Stereo-Atlas of Ostracod Shells 18, 119
Tethysobuntonia govoroffi (3 of 4)
Holotype:
Type locality:
Figured specimens:
Derivation of name:
Diagnosis:
Distribution:
A ckn o wledgemen ts:
Centre de Sedimentologie-Paleontologie, Universite de Provence, Centre Saint-Charles, Marseille,
no. HCA 1, cr carapace.
[Paratypes; nos. PCA 1/1-PCA 1/7; 3 carapaces and 4 valves].
Aschia-Tinamou water well, 45 km WSW of South Termit Massif, eastern Niger Republic, lat.
11°05'N, long. 15°27'W (H. Faure, Mem. Bur. Rech. geol. minier., 47, 1966; J. Greigert & R.
Pougnet, Ibid., 48, 1967). Upper part of the Aschia-Tinamou Formation, Campanian, Upper
Cretaceous.
Centre de Sedimentologie-Paleontologie, Universite de Provence, Centre Saint-Charles, Marseille,
coll. nos. PCA 1/1 (paratype, o' LV: PI. 18, 118, fig. 1), PCA 1/2 (paratype, o' LV: PI. 18, 118,
fig. 2), HCA 1 (holotype, o’ car.: PI. 18, 118, fig. 3), PCA 1/3 (paratype, 9 car.: PL 18, 120,
fig. 1), PCA 1/4 (paratype, 9 car.: PL 18, 120, fig. 2), PCA 1/5 (paratype, cr car.: PI. 18, 120,
fig. 3), PCA 1/6 (paratype, 9 LV: PI. 18, 120, fig. 4), PCA 1/7 (paratype, 9 LV: PL 18, 120,
fig. 5). All from the type locality.
In honour of Dr. N. Govoroff (Esso Exploration West Africa, Libreville, Gabon) who collected
and sent us the samples.
As for the genus. The genus Tethysobuntonia is currently monotypic.
Known from the Campanian of the Niger Republic (outcrops and subsurface), the Middle
Campanian to Early Maastrichtian of Algeria (Bellion et al. 1973, op. cit.). Upper Cretaceous of
Saudi Arabia (Sayyab, 1956, op. cit.), Campanian-Early Maastrichtian of Oman, Coniacian?-
Campanian of Iraq and Coniacian-Santonian of Ethiopia (Athersuch, 1988, op. cit.).
We sincerely thank Drs. N. Govoroff (Libreville, Gabon) for sending us the material to study,
P. Donze (Lyon, France) for the loan of the Algerian material and J. Athersuch (British Petroleum
pic) for access to Sayyab’s thesis.
Explanation of Plate 18, 120
Fig. 1, 9 car., ext. rt. lat. (paratype, PCA 1/3, 473 pm long); Fig. 2, 9 car., ext. vent, (paratype, PCA 1/4, 452 pm long); Fig. 3. cr
car., ext. dors, (paratype, PCA 1/5, 559 pm long); Fig. 4, 9 LV, int. lat. (paratype, PCA 1/6, 441 long); Fig. 5, 9 LV, ext. lat.
(paratype, PCA 1/7, 441 //m long).
Scale A (100 pm\ x93), figs. 1-5.
Stereo-Atlas of Ostracod Shells 18, 120
Stereo-Atlas of Ostracod Shells 18, 118
Tethysobuntonia govoroffi (2 of 4)
Tethysobuntonia govoroffi (4 of 4)
Stereo-Atlas of Ostracod Shells 18 (30) 121-124 (1991) Ogmoconchella martini (1 of 4)
595.337.2 (116.14) (420 : 162.003.51 + 003.53): 551.351 + 552.52
ON OGMOCONCHELLA MARTINI (ANDERSON)
by Ian Boomer
(University of East Anglia, Norwich)
Ogmoconchella martini (Anderson, 1964)
1951 Ostracode 800, C.A. Wicher, Erdol u. Kohle, 4, 759, pi. 1, figs. 12, 13.
1964 Hungarella martini sp. nov., F.W. Anderson, Bull. geol. Surv. Gt. Br., 21, 147, pi. 13, figs. 83-89.
1969 Healdia? tenuivirgata sp. nov., H.J. Will, Beih. geol. Jb., 54, 52, pi. 1, figs. 2a-d.
1980 Hungarella? reticulata sp. nov., E. Kristan-Tollmann, Mitt. ost. geol. Ges., 73, 197, pi. 11, figs. 12-15, pi. 12, figs. 15, 16.
Holotype: British Geological Survey (Keyworth) GSM Mik (j) 280001, 9 carapace.
Type locality: Plattlane Borehole, Whixall, Shropshire (Grid Ref. SJ 5140 3645). Westbury Formation, “Lower
Rhaetic” at a depth of 243' 0" to 243' 6".
Figured specimens: British Geological Survey (Keyworth) GSM Mik (j) 280001 (holotype, 9 car.: PI. 18, 122, figs. 2, 3,
PI. 18, 124, fig. 1); Bristol City Museum and Art Gallery (BRSMG) nos. Cel7025 (9 LV: PI. 18,
122, fig. 1), Cel 7026 (9 LV: PI. 18, 124, fig. 2), Cel7027 (9 car.: PI. 18, 124, fig. 3), Cel7028
(o* car.: PL 18, 124, fig. 4). All specimens (apart from holotype) are from uppermost bed of the
Westbury Formation, Penarth Group at Hampstead Farm Quarry, Avon (Grid Ref. ST 726 839)
(sensu D.T. Donovan et al.. Palaeontology, 32, 231, 1989); collected by M. T. Curtis, to whom
thanks are due for making them available to the author.
Explanation of Plate 18, 122
Fig. 1, 9 LV, ext. lat. (BRSMG Cel7025, 513 um long); Figs. 2, 3, 9 car. (holotype, GSM Mik (j) 280001, 538 fu m long), fig. 2, It. lat.,
fig. 3, rt. lat.
Scale A (100 //m; xlOO), figs. 1, 3; scale B (100 /rm; xl05), fig. 2.
Stereo-Atlas of Ostracod Shells 18, 123
Ogmoconchella martini (3 of 4)
Diagnosis:
Remarks:
Distribution:
Distinguished from similar, contemporary metacopine Ostracoda by the presence of short rounded
process at mid-height on the posterior of the larger left valve. Overlap entire (LV > RV), strong in
lateral view particularly along posterior and ventral margins. Greatest height behind mid-length,
anterior extremity below mid-height in both valves especially in more tumid left valve. External
lateral surfaces possess fine “fingerprint” sculpture arranged concentrically around postero-mid
valve area. Muscle scar pattern not observed. Hinge consists of deep contact groove in left valve
finely crenulated along dorsal margin and without distinct terminal widenings.
In the original description of this species, Anderson {op. cit.) commented on the presence of the fine
ornament as distinguishing it from other species of the genus. However, the current author has also
recorded such ornament, although often much reduced, in Ogmoconchella aspinata (Drexler, 1958),
Ogmoconchella aequalis { Herrig, 1969), Ogmoconchella adenticulata (Pietrzenuk, 1964), Ogmocon-
chella mouhersensis (Apostolescu, 1961) and Ogmoconchella propinqua Malz, 1971. The presence
of such ornament will be strongly dependent upon the state of preservation. This feature cannot,
therefore, be meaningfully described as characteristic of this or any other metacopine species.
Recorded from the Rhaetian (late Triassic) deposits of Germany (Wicher, 1951; Will, 1969), Great
Britain (Anderson. 1964; this work) and Iran (Kristan-Tollmann, 1980).
Explanation of Plate 18, 124
Fig. 1, 9 car. (holotype, GSM Mik (j) 280001), detail of posterior process and ornament; Fig. 2, 9 LV, int. lat. (BRSMG Cel7026,
526 qm long); Fig. 3, o* car., ext. dors. (BRSMG Cel7027, 513 /rm long); Fig. 4, 9 car., ext. dors. (BRSMG Cel7028, 487 /rm long).
Scale A (50/rm; x200), fig. 1; scale B (100 /rm; xlOO), figs. 2, 4; scale C (100 /rm; xl05), fig. 3.
Ogmoconchella martini (4 of 4)
Ogmoconchella martini (2 of 4)
Stereo- Atlas of Ostracod Shells 18, 122
Stereo-Atlas of Ostracod Shells 18, 124
Stereo-Atlas of Ostracod Shells 18 (31) 125-134 (1991) Frostiella groenvalliana (1 of 10)
595.336.11 (113.333) (420 : 162.003.52 + 430 : 161.013.52/012.54 + 438.161 .017.54 + 485 : 161.013.55): 551.351 + 552.51 +552.54
ON FROSTIELLA GROENVALLIANA MARTINSSON
by Wolfgang Hansch, David J. Siveter & C. Giles Miller
(University of Greifswald, Germany & University of Leicester, England)
Genus FROSTIELLA Martinsson, 1963
Type-species (by original designation): Frostiella groenvalliana Martinsson, 1963
Diagnosis: Kloedeniinae with crumina strongly assimilated with the domicilum; crumina with a narrow, striate,
and somewhat swollen field between the distinct velar bend and the marginal structure. Syllobium
with protruding cusp (after Martinsson 1963).
Remarks: This beyrichiacean genus also currently includes F. pliculata Martinsson, 1963; F. cornuta
Martinsson, 1965; F. loodensis Sarv, 1968; F. bicristata Shaw, 1969; and/7, modesta Abushik, 1971.
Frostiella is traditionally regarded as of Upper Silurian Pridoli Series (cf. Martinsson 1963) age but
may possibly also occur in the top of the Ludlow Series (e.g. see Siveter 1989 and below).
Frostiella groenvalliana Martinsson, 1963
1897 Kloedenia wilckensiana Jones; K.A. Gronwall, Sver. geol. Unders. Afh., ser. C, 170 (passim).
1909 Kloedenia wilckensiana Jones et var. plicata Jones; J.Ch. Moberg & K.A. Gronwall, Acta Univ. lund., ser. 2, 5, 1, 66-67,
pi. 6, figs. 6, 7.
1963 Frostiella groenvalliana n. sp., A. Martinsson, Bull. geol. Instn Univ. Uppsala , 42(2), 29-33, figs. 7C, 8, 14, 15A, 15B, 16A,
16B, 17A-F.
1963 Frostiella cf. groenvalliana ; A. Martinsson, Ibid., 34, figs. 18A-D.
Explanation of Plate 18, 126
Figs. 1-3, 9 LV (LO 2183T, 2440 /rm long): fig. 1, ext. lat.; fig. 2, ext. vent.; fig. 3, detail of preadductorial lobe. Figs. 4-6, cr LV
(LO 2184T, 2400 ,um long): fig. 4, detail of preadductorial lobe; fig. 5, ext. lat.; fig. 6 ext. vent.
Scale A (500 jum; x23), figs. 1, 2; scale B (150 /rm; x60), fig. 3; scale C (125 //m; x72), fig. 4; scale D (500 /um; x24), figs. 5, 6.
Stereo-Atlas of Ostracod Shells 18, 127 Frostiella groenvalliana (3 of 10)
1963 Frostiella aff. groenvalliana ; A. Martinsson, Ibid. , 34.
1964 Frostiella lebiensis n. sp., A. Martinsson, Geol. For. Stockh. Fork., 86(2), 139-142, 155-156, 158, figs. 8A-D, text-fig. 15.
1967 Frostiella groenvalliana-, A. Martinsson, Geol. For. Stockh. Forh., 89(3), 376, 377, 379, text-figs. 2, 3.
1967 Frostiella lebiensis Martinsson; L. Gailite, in: L. Gailite, et ah, Stratigrafija, fauna i uslovija obrazovania silurijskich porod
sredney Pribaltiki, 143, pi. 10, fig. 13, Riga (Zinatne).
1968 Frostiella groenvalliana Martinsson; L. Sarv, Ostrakody Craspedobolbinidae, Beyrichiidae, Primitiopsidae silura Estonii, 58,
pi. 20, figs. 6-8, Tallinn (Valgus).
1969 Frostiella groenvalliana Martinsson; R.W.L. Shaw, Geol. For. Stockh. Forh., 91(1), 55, 65, 67, 68, 70, figs. 1A-F, text-fig. 8.
1970 Frostiella groenvalliana Martinsson; L. Sarv, in: D. Kaljo (Ed.), Silur Estonii, 159, 169, tab. 18, Tallinn (Valgus).
1971 Frostiella groenvalliana Martinsson; R.W.L. Shaw, Palaeonotology , 14(4), 603, pi. 113, figs. 1-5, 7.
1971 Frostiella groenvalliana Martinsson; L. Sarv, Eesti NSV Tead. Akad. Toim., Khim. Geol., 20(4), 353-355, text-fig. 2.
1974 Frostiella groenvalliana Mart.; L. Gailite & R. Ulste, in: Regional’naja geologija Pribaltiki, 40-41, text-fig. 1, Riga (Zinatne).
1974 Frostiella lebiensis Martinsson; E. Tomczykowa & E. Witwicka, Biul. Inst, geol., 276, 58-59, 63, text-fig. 2.
1976 Frostiella groenvalliana Martinsson; D. Kaljo & L. Sarv, Eesti NSV Tead. Akad. Toim., Khim. Geol., 25(4), 328.
1976 Frostiella lebiensis Martinsson; Ibid. , 326, 328-329.
1977 Frostiella groenvalliana Martinsson; A. Martinsson, in: A. Martinsson (Ed.), The Siluro-Devonian Boundary, IUGS ser. A,
no. 5, 46, 48, 49, 327-329, fig. 1, text-fig. 3.
1977 Frostiella groenvalliana Martinsson; L. Sarv, in: D. Kaljo (Ed.), Fazii i fauna silura Pribaltiki, 166, 7171, 173, 175, ?text-fig. 7,
Tallinn (Valgus).
1977 Frostiella sp. cf. F. groenvalliana Martinsson; M.J. Copeland & J.M. Berdan, Geol. Surv. Pap. Can., 77-1B, pi. 2.3, fig. 19.
1978 Frostiella groenvalliana Martinsson; D.J. Siveter, in: R. Bate & E. Robinson (Eds.), Geol. J., (Special Issue) 8, 68, 86, pi. 9,
figs. 7, 8, tab. 2.
1978 Frostiella groenvalliana Mart.; L. Gailite, in: Stratigrafija fanerozoja Pribaltiki, 13, 15, 18, 19, Riga (Zinatne).
1978 Frostiella groenvalliana Martinsson; D. Kaljo, Eesti NSV Tead. Akad. Toim., Geol., 27(1), 7-9.
1978 Frostiella groenvalliana Martinsson; D.E. White & B.C. Coppack, Bull. geol. Surv. Gt. Br., 62 (for 1977), 30, pi. 1, figs. 10-12.
1982 Frostiella groenvalliana Martinsson; M.G. Bassett et al., Lethaia, 15(1), 8, 15-18, text-fig. 6.
1982 Frostiella groenvalliana Martinsson; I.J. Paskevicius, Geologija, 3, 20, 44, 46, 47, text-fig. 1.
Explanation of Plate 18, 128
Figs. 1-3, 9 RV (SGWG 90/1, approx. 2250/rm long): fig. 1, ext. lat.; fig. 2, ext. vent.; fig. 3, detail of preadductorial lobe. Figs. 4-6,
cr RV (SGWG 90/2, approx. 2100 /rm long): fig. 4, detail of preadductorial lobe; fig. 5, ext. lat.; fig. 6, ext. vent.
Scale A (350 /um; x27), figs. 1, 2, 5, 6; scale B (175 //m; x55), fig. 3; scale C (125 /rm; x70), fig. 4.
Stereo- Atlas of Ostracod Shells 18, 126
Frost iella groenvalliana (2 of 1 0)
Frostiella groenvalliana (4 of 10)
Stereo-Atlas of Ostracod Shells 18, 128
Slereo-Atlas of Ostracod Shells 18, 129
Frostiella groenvalliana (5 of 10)
1982 Frostiella groenvalliana; L. Sarv, in: D. Kaljo & E. Klaamann (Eds.), Ecostratigraphy of the East Baltic Silurian, 75, 76, 78,
Tallinn (Valgus).
1982 F. groenvalliana ; A. Abushik, in: Tezisy dokladov 28th sessii Vsesojuznogo Paleontologiceskogo obscestva, 3, 4, Taskent.
1983 Frostiella groenvalliana Martinsson; J.M. Berdan, in: R.F. Maddocks (Ed.), Applications of Ostracoda, 314, fig. 31, Univ.
Houston.
1983 Frostiella groenvalliana ; D. Kaljo et al., in: Problemy ekologii fauny i flory drevnych basseinov, 48, Moskva.
1984 Frostiella groenvalliana; D. Kaljo et al., in: Stratigrafija i paleontologija drevnesego fanerozoja, 101, Moskva.
1984 Frostiella groenvalliana Martinsson; D.J. Siveter, Spec. Pap. Palaeont., 32, 82.
? 1 985 Frostiella groenvalliana s.l.; A. Abushik et al., Lethaia, 18(2), 139, 142, 143.
1985 Frostiella groenvalliana Martinsson; W. Hansch, Lethaia, 18(4), 274, 277, 278, tab. 1, fig. IF, text-fig. 3.
1986 Frostiella groenvalliana; L. Gailite, in: D. Kaljo & E. Klaamann (Eds.), Teorija i opyt ekostratigrafija. 111, 114, Tallinn
(Valgus).
1986 Frostiella groenvalliana; N.V. Sidaraviciene, Ibid. , 119, tab. 1.
1988 Frostiella groenvalliana Martinsson; D.J. Siveter, The Lower Palaeozoic of the Northern Welsh Borderland and South Wales,
10th Int. Symp. on Ostracoda, Aberystwyth, Field Guide No. 2, 36, text-fig. 7, pi. 2, fig. 2.
1989 Frostiella groenvalliana Martinsson; D.J. Siveter, in: C.H. Holland & M.G. Bassett (Eds.), A global standard for the Silurian
System, 258, 263, text-fig. 167, figs. 168 J, K, Nat. Mus. Wales, Geol. Ser. no. 9, Cardiff.
1989 Frostiella groenvalliana Martinsson; D.J. Siveter et. al., Silurian field excursions. A geotraverse across Wales and the Welsh
Borderland, 45, text-figs. 30, 38, 40, pi. 3, fig. 14, Nat. Mus. Wales Geol. Ser., no. 10, Cardiff.
1990 Frostiella groenvalliana Martinsson; T. Meidla & L. Sarv, in: D. Kaljo & H. Nestor (Eds.), Field Meeting Estonia 1990. An
Excursion Guidebook, pi. 9, fig. 4, tab. 11, Tallinn.
Holotype: Palaeontological Inst. Univ. Lund, Sweden, no. 4084T; 9 left valve.
[Paratypes; Museum Naturkunde, Berlin, nos. MB.O. 174-177 (Krause sample Ringsho 1),
MB.O. 178 & 179 (Krause sample Ringsho 2). Figured Martinsson 1963].
Explanation of Plate 18, 130
Fig. 1, o* LV, detail of uppermost part of preadductorial lobe (LO 2184T). Figs. 2, 3, 9 LV (LO 2183T): fig. 2, detail of ornament on
ventral side of crumina; fig. 3, ext. ant. Figs. 4-6, o* RV (X 2603, 2320 pm long): fig. 4, ext. lat.; fig. 5, ext. vent.; fig. 6, detail of
preadductorial lobe. Fig. 7, cr LV, ext. lat. (MB.O. 178, 2065 pm long).
Scale A (150,um; x60), fig. 1; scale B (50 pm; xl80), fig. 2; scale C (375 pm; x24), fig. 3; scale D (390 pm; x23), figs. 4, 5; scale E
(80 pm; x70), fig. 6; scale F (340 pm; x26); fig. 7.
Stereo-Atlas of Ostracod Shells 18, 131 Frostiella groenvalliana (7 of 10)
Type locality: “Beds 3-4” sensu Gronwall at Ramsasa, Scania, Sweden; lat. 55°33'N, long. 13°53'E; Pridoli
Series, Silurian.
Figured specimens: Palaeontological Institute, University of Lund, Sweden, nos. LO 2183T (9 LV: PI. 18, 126, figs.
1-3; PI. 18, 130, figs. 2, 3); LO 2184T (cr LV: PI. 18, 126, figs. 4, 5, 6; PI. 18, 130, fig. 1). Both
Moberg & Gronwall coll. (1909, pi. 6, figs. 6, 7); “Bed 4” sensu Gronwall, at Ramsasa, Sweden.
Sektion Geologische Wissenschaften der E.-M.-Arndt-Universitat Greifswald, Germany, nos.
SGWG 90/1 (9 RV: PI. 18, 128, figs. 1-3) from erratic boulder no. 549 of Krause, Mtiggelheim,
Berlin, approx, lat. 52° 32' N, long. 13°25'E; SGWG 90/2 (o* RV: PI. 18, 128, figs. 4-6) from erratic
boulder no. Bey. B20, Graal-Miiritz near Rostock, Germany, lat. 54° 15'N, long. 12° 15'E.
Zentrales Geologisches Probenarchiv Bernau bei Berlin, Germany, no. X 2603 (o* RV: PI. 18, 130,
figs. 4-6; = holotype of F. lebiensis Martinsson 1964, figs. 8C, D). From borehole Leba 1, 687.5 m,
Pomerania, Poland; lat. 54°45'N, long. 17°34'E.
Museum fur Naturkunde Berlin, Germany, no MB.O. 178 (o* LV: PI. 18, 130, fig. 7; = paratype
of F. groenvalliana Martinsson, 1963, fig. 16B). From Krause’s sample “Ringsho 2”, Klinta,
Scania, Sweden; lat. 55°51'N, long. 13°30'E.
British Museum (Natural History), London, nos. OS 6618 (9 RV: PI. 18, 132, fig. 4), OS 6619
(tecnomorphic LV: PI. 18, 132, fig. 1), OS 6620 (o* LV: PI. 18, 132, fig. 2), OS 6621 (9 RV: PI. 18,
132, fig. 3), OS 6622 (tecnomorphic RV: PI. 18, 132, fig. 6). All from Platyschisma Shale Member,
Downton Castle Sandstone Formation, Downton Group, c. 1.5 m above the Ludlow Bone Bed on
N side of Ludford Lane, Ludlow, Shropshire, Great Britain (Grid Ref.: SO 5119 7413); coll. D.J.
Siveter. OS 13922 (tecnomorphic LV: PI. 18, fig. 5); loose material, Downton Group, Ludford
Corner excavation, Ludlow (Grid Ref.: SO 5123 7413); coll. C.G. Miller.
Explanation of Plate 18, 132
Fig. 1, tecnomorphic LV (OS 6619, 1780yi/m long): ext. lat. Fig. 2, cr LV (OS 6620, 2025 «m long): preadductorial lobe. Fig. 3, 9 RV
(OS 6621, 2100 pm long): obi. vent. Fig. 4, 9 RV (OS 6618, 2010/rm long): ext. lat. Fig. 5, tecnomorphic LV (OS 13922, 1470^m
long): dors. Fig. 6, tecnomorphic RV (OS 6622, 1370/rm long): ext. lat.
Scale A (400 pm; x30), fig. 1; scale B (200 //m; x40), fig. 2; scale C (400 pm; x27), figs. 3, 4; scale D (300 pm; x35), fig. 5; scale E
(300 pm; x35), fig. 6.
Stereo-Atlas of Ostracod Shells 18, 132
Frostiella groenvalliana (8 of 10)
Stereo-Atlas of Ostracod Shells 18, 130
Frostiella groenvalliana (6 of 10)
Stereo-Atlas of Ostracod Shells 18, 133 Frostiella groenvalliana (9 of 10)
Diagnosis: Frostiella species with well developed lobation and prominent cusps on the anterior lobe and the
anterior lobule of the syllobium. In adults cristal loop on the preadductorial lobe complete, drawn
out in sagitto-dorsal direction or nearly rounded. Valve surface smooth except for the striate
cruminal field and the ornament (reticulostriation/striation/punctation) on lateral facet of the
preadductorial lobe.
Remarks: F. groenvalliana differs from the other Frostiella species particularly by its more distinctly developed
lobal cusps and the characteristic form of its preadductorial lobe, a feature which is also obvious in
juveniles. Martinsson (1977) assumed that F. groenvalliana and F. lebiensis are synonymous. Their
morphological characteristics and stratigraphical ranges are both very similar to each other. Only in
the development of the preadductorial lobe is there a slight difference. In specimens hitherto
described as F. lebiensis there is mostly a more rounded cristal loop (not pointed and somewhat dis-
torted as in “typical” F. groenvalliana specimens) on the preadductorial lobe. Furthermore, it is
probable that the “typical” F. groenvalliana is restricted to the basal Pridoli in Britain (Downton
Group) and the Baltic whereas specimens with a more rounded cristal loop have a somewhat greater
stratigraphical range (and may pass over continuously into the F. cornuta lineage). In the Baltic area
such changes appear to correlate with ecological ranges from shallow water facies ( groenvalliana
specimens) to somewhat deeper water conditions ( lebiensis specimens). This assumption is supported
by the occurrence of the latter in the probably deeper, basinal (outer shelf) areas represented in, for
example, the Leba elevation (Tomczykowa & Witwicka 1974) and the Kaliningrad region Dubovskoe
borehole (Kaljo & Sarv 1976). F. groenvalliana and F. lebiensis are considered as ecophenotypical
intraspecific variants. Compared to the Scanian (shell) material of F. groenvalliana, Welsh Basin
specimens (moulds) have a less ventrally drooping lateral profile to the lateroventral lobal connec-
tion (tecnomorphs) and crumina (females). This and other minor morphological differences are
judged to be of infraspecific significance.
Distribution: F. groenvalliana is considered generally indicative of early Pridoli Series (Upper Silurian) levels in
an area extending from Podolia to eastern North America (Siveter 1989, 258-263, fig. 164). However,
it should be noted that F. groenvalliana (“lebiensis”) and the key graptolite ( Monograptus parultimus
Jaeger) for the base of the Pridoli in the Czechoslovakian stratotype area do not occur coevally in any
of the relevant sections. M. parultimus and F. groenvalliana (“lebiensis”) occur geographically
together only in the Dubovskoe borehole (Kaliningrad region; Kaljo & Sarv 1976), but at different
Stereo-Atlas of Ostracod Shells 18, 134 Frostiella groenvalliana (10 of 10)
Distribution: horizons (Kuressare horizon and the younger Aigu Member, Kaugatuma horizon respectively). As
(continued) possibly indicated by conodont correlation (Schonlaub, H.P. in: Kriz, J. etal.,Jb. Geol. Bundesanst.
Wein, 129, 1986) the Ludlow-Pridoli boundary may be slightly above the first occurrence of F.
groenvalliana at Ludlow, Britain (i.e. may be in the Downton Group) and parts of the Baltic.
Sweden: Gronwall’s “Bed 3” and “Bed 3-4” at Klinta and Ramsasa, Scania (Martinsson 1963,
1967); = top part of Oved-Ramsasa Group sensu Jeppsson & Laufeld (Sver. geol. Unders. Afh.,
ser. Ca, no. 58, 1987).
Great Britain: Scout Hill Flags, Lake District (Shaw 1971). Downton Castle Sandstone Formation
at Long Mountain (Shaw 1969), Shropshire (Shaw 1969, Siveter 1978, 1988, 1989, Bassett et al.
1982), and English W Midlands ( F . cf. groenvalliana ; Siveter 1989) parts of Welsh Basin. Also (?)
Lakenheath borehole, E England and (?) uppermost Ludlow, Cennen Valley, Wales (see Bassett
et al. 1982, Siveter 1989).
N America: Leighton Formation, Maine, U.S.A. (Martinsson 1967; Copeland & Berdan 1977;
Berdan 1983). Possibly also occurs in Stonehouse Formation, Nova Scotia, Canada (Martinsson
1967).
East Baltic area: Ohessare 1 borehole, Venekjula and Ejgu, Isle of Saaremaa, Estonia; Kaugatuma
Formation (Sarv 1968, 1970, 1971). Piltene 1, 31, 32, Kolka 4, 54, Pavilosta 51, Ventspils 3, Talsy
55 and Ezere boreholes, Latvia; Minija and basal part of Jura formations (Gailite & Ulst 1974,
Gailite 1978, 1986, Sarv 1982). Stoniskiai and Vidukle boreholes, Lithuania; Minija Formation
(Sarv 1982, Paskevicius 1982). Borehole 110 of Arjogal profile, Lithuania; Minija Formation
(Sidaraviciene 1986). Dubovskoe borehole, Kaliningrad Region; Aigu Beds, Kaugatuma Formation
(Kaljo & Sarv 1976).
Poland: Borehole Leba 1 (Martinsson 1964). Several boreholes of the Peribaltic area (cf.
Tomczykowa & Witwicka 1974, 58); lowermost Podlasian.
Erratic boulders: Beyrichienkalk type B and “Red Beyrichienkalk” sensu Hansch (1985);
“Local” limestone boulders of the Hoburg Bank (Martinsson 1967, 1977).
?Podolia, USSR: Dnestr river, between Okopy and Belovtsy, Raskov suite, Skala Horizon
(Abushik et al. 1985, Koren et al. in: Holland & Bassett (Eds.), 1989).
Stereo-Atlas of Ostracod Shells 18 (32) 135-137 (1991)
Index, Volume 18, 1991 (1 of 3)
General Index
Ainsworth, N. R. & Boomer, I., On Ektyphocythere bizoni Ainsworth; 33-36
auricula, Lomatopisthia; 17-20
Ayress, M. A., On Vitjasiella ferox (Hornibrook); 57-60
Babinot, J.-F., On Valdonniella mackenziei ; 41-44
Babinot, J.-F. & Colin, J.-P., On Tethysobuntonia govoroffi Colin & Babinot gen. et sp. nov.; 117-120
Becker, G., On Bufanchiste sotoi Becker; 25-28
Becker, G., On Cleithranchiste paulusi Becker; 21-24
Becker, G. & Weyer, D., On Villozona villosa (Griindel); 85-88
bizoni, Ektyphocythere ; 33-36
Boomer, I., On Ektyphocythere cookiana (Anderson); 113-116
Boomer, I., On Ogmoconchella martini (Anderson); 121-124
Boomer, I. & Ainsworth, N. R., On Ektyphocythere bizoni Ainsworth; 33-36
Boomer, I. & Horne, D. J., On Cytheromorpha fuscata (Brady); 49-56
bransoni, Huntonella ; 89-92
Bufanchiste sotoi Becker; 25-28
Capricambria cornucopiae Hinz gen. et sp. nov.; 65-68
Cleithranchiste paulusi Becker; 21-24
Cocoonocythere sinensis Zhao; 101-104
Coker, J. E. & Dewey, C. P., On Youngiella rectidorsalis (Jones & Kirkby); 29-32
Colin, J.-P. On Frambocythere tumiensis (Helmdach) ferreri Colin; 37-40
Colin, J.-P. & Babinot, J.-F., On Tethysobuntonia govoroffi Colin & Babinot gen. et sp. nov.; 117-120
cookiana, Ektyphocythere', 113-116
corbuloides, Daleiella; 81-84
cornucopiae, Capricambria', 65-68
costaereticulata, Paracathaycythere', 93-96
costata, Venzavella', 9-12
Cytheromorpha fuscata (Brady); 49-56
Daleiella corbuloides (Jones & Holl); 81-84
Dewey, C. P. & Coker, J. E., On Youngiella rectidorsalis (Jones & Kirkby); 29-32
Dewey, C. P. & Puckett, T. M., On Mammoides dorsospinosus Sohn; 73-76
dorsospinosus, Mammoides', 73-76
Ektyphocythere bizoni Ainsworth; 33-36
Ektyphocythere cookiana (Anderson); 113-116
ferox, Vitjasiella', 57-60
Frambocythere tumiensis (Helmdach) ferreri Colin; 37-40
Frostiella groenvalliana Martinsson; 125-134
fuscata, Cytheromorpha', 49-56
govoroffi, Tethysobuntonia', 117-120
groenvalliana, Frostiella', 125-134
Hansch, W., Siveter, D. J. & Miller, C. G., On Frostiella groenvalliana Martinsson; 125-134
Hansch, W. & Wang Shang-qi, On Qujingsia nonaculeata Hasch & Wang gen. et sp. nov.; 77-80
Hemingwayella pumilio (Brady); 45-48
Herrig, E. R., On Puncia levis Herrig; 61-64
Hinz, I. C. U., On Capricambria cornucopiae Hinz gen. et sp. nov.; 65-68
Hinz, I. C. U., On Ulopsis ulula Hinz gen. et sp. nov.; 69-72
Horne, D. J. & Boomer, I., On Cytheromorpha fuscata (Brady); 49-56
Huntonella bransoni Lundin; 89-92
hyperdonta, Polydontoconcha; 105-108
levis, Puncia', 61-64
Limbinariella macroreticulata Sarv; 5-8
Lomatopisthia auricula (Harris); 17—20
Lomatopisthia simplex (Harris); 13-16
Lundin, R. F. & Petersen, L. E., On Daleiella corbuloides (Jones & Holl); 81-84
Lundin, R. F. & Siveter, D. J., On Huntonella bransoni Lundin; 89-92
mackenziei, Valdonniella', 41-44
macroreticulata, Limbinariella', 5-8
Mammoides dorsospinosus Sohn; 73-76
martini, Ogmoconchella', 121-124
Maybury, C. A., On Palmoconcha ruggierii Maybury sp. nov.; 109-112
Maybury, C. A. & Whatley, R. C., On Hemingwayella pumilio (Brady); 45-48
Miller, C. G., Hansch, W. & Siveter, D. J., On Frostiella groenvalliana Martinsson; 125-234
nonaculeata, Qujingsia; 77-80
Ogmoconchella martini (Anderson); 121-124
Orcofabella testata (Gailite); 1-4
Palmoconcha ruggierii Maybury sp. nov.; 109-112
Paracathaycythere costaereticulata Whatley & Zhao gen. et sp. nov.; 93-96
Paracathaycythere scabra Zhao & Whatley sp. nov.; 97-100
paulusi, Cleithranchiste; 21-24
Petersen, L. E. & Lundin, R. F., On Daleiella corbuloides (Jones & Holl); 81-84
Polydontoconcha hyperdonta Zhao & Whatley gen. et sp. nov.; 105-108
Puckett, T. M. & Dewey, C. P., On Mammoides dorsospinosus Sohn; 73-76
pumilio, Hemingwayella; 45-48
Puncia levis Herrig; 61-64
Qujingsia nonaculeata Hansch & Wang gen. et sp. nov.; 77-80
rectidorsalis, Youngiella; 29-32
ruggierii, Palmoconcha; 109-112
Stereo-Atlas of Ostracod Shells 18, 136
Index, Volume 18, 1991 (2 of 3)
Sarv, L. & Siveter, D. J., On Limbinariella macroreticulata Sarv; 5-8
Sarv, L. & Siveter, D. J., On Orcofabella testata (Gailite); 1-4
Sarv, L. & Siveter, D. J., On Venzavella costa ta (Neckaja); 9-12
sea bra, Paracathaycy there; 97-100
simplex, Lomatopisthia ; 13-16
sinensis, Cocoonocythere; 101-104
Siveter, D. J., Hansch, W. & Miller, C. G., On Frostiella groenvalliana Martinsson; 125-134
Siveter, D. J. & Lundin, R. F., On Huntonella bransoni Lundin; 89-92
Siveter, D. J. & Sarv, L., On Limbinariella macroreticulata Sarv; 5-8
Siveter, D. J. & Sarv, L., On Orcofabella testata (Gailite); 1-4
Siveter, D. J. & Sarv, L., On Venzavella costata (Neckaja); 9-12
sotoi, Bufanchiste; 25-28
testata, Orcofabella ; 1-4
Tethysobuntonia govoroffi Colin & Babinot gen. et sp. nov.; 117-120
tumiensis ferreri, Frambocythere; 37-40
Ulopsis ulula Hinz gen. et sp. nov.; 69-72
ulula, Ulopsis; 69-72
Valdonniella mackenziei Babinot; 41-44
Venzavella costata (Neckaja); 9-12
villosa, Villozona; 85-88
Villozona villosa (Griindel); 85-88
Vitjasiella ferox (Hornibrook); 57-60
Wang Shang-qi & Hansch, W., On Qujingsia nonaculeata Hansch & Wang gen. et sp. nov.; 77-80
Weyer, D. & Becker, G., On Villozona villosa (Griindel); 85-88
Whatley, R. C. & Maybury, C. A., On Hemingwayella pumilio (Brady); 45-48
Whatley, R. C. & Zhao Quanhong, On Coccoonocythere sinensis Zhao; 101-104
Whatley, R. C. & Zhao Quanhong, On Paracathaycy there costaereticulata Whatley & Zhao gen. et sp. nov.; 93-96
Whatley, R. C. & Zhao Quanhong, On Paracathaycythere scabra Zhao & Whatley sp. nov.; 97-100
Whatley, R. C. & Zhao Quanhong, On Polydontoconcha hyperdonta Zhao & Whatley gen. et sp. nov.; 105-108
Williams, M., On Lomatopisthia auricula (Harris); 17-20
Williams, M., On Lomatopisthia simplex (Harris); 13-16
Youngiella rectidorsalis (Jones & Kirkby); 29-32
Zhao Quanhong & Whatley, R. C., On Cocoonocythere sinensis Zhao; 101-104
Zhao Quanhong & Whatley, R. C., On Paracathaycythere costaereticulata Whatley & Zhao gen. et sp. nov.; 93-96
Zhao Quanhong & Whatley, R. C., On Paracathaycythere scabra Zhao & Whatley sp. nov.; 97-100
Zhao Quanhong & Whatley, R. C., On Polydontoconcha hyperdonta Zhao & Whatley gen. et sp. nov.; 105-108
Stereo-Atlas of Ostracod Shells 18, 137
Index, Volume 18, 1991 (3 of 3)
Index; Geological Horizon
See 1 (1) 5-22 (1973) for explanation of the Schedules in the Universal Decimal Classification
The editors gratefully acknowledge financial support from
BP International Ltd for the publication of this volume
BPCC Blackpool
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Stereo-Atlas of Ostracod Shells: Vol. 18, Part 2
CONTENTS
Back volumes:
On Ulopsis ulula Hinz gen. et sp. nov.; by I.C.U. Hinz.
On Mammoides dorsospinosus Sohn; by C.P. Dewey & T.M. Puckett.
On Qujingsia nonaculeata Hansch & Wang gen. et sp. nov.; by W. Hansch & Wang
Shang-qi.
On Daleiella corbuloides (Jones & Holl); by R.F. Lundin & L.E. Petersen.
On Villozona villosa (Griindel); by G. Becker & D. Weyer.
On Huntonella bransoni Lundin; by D J. Siveter & R.F. Lundin.
On Paracathaycythere costaereticulata Whatley & Zhao gen. et sp. nov.; by R.C.
Whatley & Zhao Quanhong.
On Paracathaycythere scabra Zhao & Whatley gen. et sp. nov.; by Zhao Quanhong &
R.C. Whatley.
On Cocoonocythere sinensis Zhao; by Zhao Quanhong & R.C. Whatley.
On Polydontoconcha hyperdonta Zhao & Whatley gen. et sp. nov.; by Zhao
Quanhong & R.C. Whatley.
On Palmoconcha mggierii Maybury sp. nov.; by CA. Maybury.
On Ektyphocythere cookiana (Anderson); by I. Boomer.
On Tethysobuntonia govoroffi Colin & Babinot gen. et sp. nov.; by J.-P. Colin & J.-F.
Babinot.
On Ognioconchella martini (Anderson); by I. Boomer.
On Frostiella groenvalliana Martinsson; by W. Hansch, D J. Siveter & C.G. Miller.
Index for Volume 18, (1991).
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