A Stereo-Atlas of Ostracod Shells

edited by J. Athersuch, D. J. Horne, D. J. Siveter,

and J. E. Whittaker

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Published under the aegis of the British Micropalaeontological Society, London

ISSN 0952-7451

Editors

Dr J. Athersuch, StrataData Ltd., 16 Ottershaw Park, Ottershaw, Surrey KT16 OGQ.

Dr D.J. Horne, School of Earth Sciences, University of Greenwich, Walburgh House, Bigland Street,

London El 2NG.

Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.

Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road,

London SW7 5BD.

Editorial Board

Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France.

Dr M.A. Ayress, Department of Geology, Australian National University, G.P.O. Box 4, Canberra,

ACT 2601, Australia.

Dr W. Hansch, Emst-Moritz-Amdt Universitat, Sektion Geologische Wissenschaften, F.L.-Jahnstr. 17a,

2200 Greifswald, Germany.

Prof. R. Lundin, Department of Geology, Arizona State University, Tempe, Arizona 85287-1404, U.S.A.

Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55,

D 2000 Hamburg 13, Germany.

Prof. N. Ikeya, Institute of Geosciences, Shizuoka University, Shizuoka 422, Japan.

Officers of the British Micropalaeontological Society

Chairman Professor A.R. Lord, Department of Geological Sciences, University College London, Gower

Street, London WC1E 6BT.

Secretary Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.

Treasurer Dr I.P. Wilkinson, British Geological Survey, Keyworth, Nottingham NG12 5GG.

Journal Editor Dr M.C. Keen, Department of Geology, The University, Glasgow G12 8QQ.

Newsletter Editor Dr A.J. Powell, Millenia Ltd., Unit 3, Weyside Park, Newman Lane, Alton, Hampshire

GU34 2PJ.

Conodont Group Chairman Dr J.J. Stone, Department of Geology, Trinity College, Dublin 2, Ireland.

Conodont Group Secretary Dr S.J. Tull, Cambridge Arctic Shelf Programme, West Building, Gravel Hill,

Huntington Road, Cambridge CB3 0DJ.

Foraminifera Group Chairman Dr M.D. Simmons, BP Exploration Operating Company Ltd., 4/5 Long

Walk, Stockley Park, Uxbridge, Middlesex UB11 IBP.

Foraminifera Group Secretary Dr S.R. Packer, Millenia Ltd., Unit 3, Weyside Park, Newman Lane,

Alton, Hampshire GU34 2PJ.

Ostracod Group Chairman Dr N.R. Ainsworth, Paleo Services Ltd., Unit 15, Paramount Industrial Estate,

Sandown Road, Watford WD2 4XA.

Ostracod Group Secretary Dr I.D. Boomer, Institute of Earth Studies, The University of Wales, Penglais,

Aberystwyth, Dyfed SY23 3DB.

Palynology Group Chair Professor D.J. Batten, Institute of Earth Studies, The University of Wales,

Penglais, Aberystwyth, Dyfed SY23 3DB.

Palynology Group Secretary Dr A. McNestry, British Geological Survey, Keyworth, Nottingham

NG12 5GG.

Calcareous Nannofossil Group Chairman Dr L.T. Gallagher, Paleo Services, Unit 15, Paramount

Industrial Estate, Sandown Road, Watford WD2 4XA.

Calcareous Nannofossil Group Secretary Dr N.M. Hine, British Geological Survey, Keyworth,

Nottingham NG12 5GG.

Instructions to Authors

Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. Format

should follow the style set by the papers in this issue. Descriptive matter apart from illustrations should

be cut to a minimum; preferably each plate should be accompanied by only one page of text. Blanks to

aid in mounting figures for plates may be obtained from any one of the Editors or Editorial Board.

Completed papers should be sent to one of the Editors. All contributions submitted for possible publication

in the Stereo-Atlas of Ostracod Shells are reviewed by an appropriate international specialist.

The front cover shows a male left valve (upper) and a female right valve (lower) of Eurybolbina bispinata

(Harris, 1957) from the middle Ordovician Edinburg Formation of Virginia, U.S.A. British Museum

(Natural History), nos. OS14028 and OS13536 respectively. Photographed by M. Williams and C. Giles

Miller.

Stereo-Atlas of Ostracod Shells 20 (1) 1-4 (1993) Cytheromorpha diamphidia (1 of 4)

595.337.14 (118.22) (420 : 162.006.50): 551.35

ON CYTHEROMORPHA DIAMPHIDIA MAYBURY sp. nov.

by Caroline A. Maybury

(Institute of Earth Studies, University of Wales, Aberystwyth)

Cytheromorpha diamphidia sp. nov.

The Natural History Museum, London [BMNH] no. OS 14217; 9 RV.

[Paratypes nos. OS 14218-OS 14221],

Sample no. 1, Vicarage Pit, St. Erth, Cornwall, England (5°26'W, 50° 10' N; Nat. Grid Ref. SW

556352); Upper Pliocene.

Greek, 8tap(pt5to<; — diamphidios — utterly, entirely different; referring to the marked dimorphism

of this species.

The Natural History Museum, London [BMNH] nos. OS 14217 (holotype, 9 RV: PI. 20, 2, fig. 1),

OS 14218 (paratype, 9 LV: PI. 20, 2, figs. 2, 3), OS 14219 (paratype, o* RV: PI. 20, 2, fig. 4; PI.

20, 4, fig. 1), OS 14220 (paratype, O' LV: PI. 20, 4, figs. 2, 3), OS 14221 (paratype, 9 LV: PI. 20,

4, fig. 4). All paratypes are from the same sample as the holotype, with the exception of paratype

OS 14220 which is from sample no. 29 (Blue Clay), from the type locality. See C.A. Maybury, Tax-

onomy, Pa/aeoeco/ogy and Biostratigraphy of Pliocene Benthonic Ostracoda from St. Erth and

NW France, unpub. PhD thesis, Univ. Wales, 1, 3-6, 1985 for further sample details.

A medium-sized, strongly dimorphic Cytheromorpha. Female subovate in lateral view and male

subreniform. Ornament of reticulae of varying shape, but predominantly subcircular in outline.

Holotype:

Type locality:

Derivation of name:

Figured specimens:

Diagnosis:

Explanation of Plate 20, 2

Fig. 1, 9 RV, ext. lat. (OS 14217, 650 pm long); Figs. 2, 3, 9 LV, dors, and ext. lat. (OS 14218, 660 pm long); Fig. 4, o' RV, ext. lat.

(OS 14219, 730 pm long).

Scale A (200 pm; x91), figs. 1-3.

Stereo-Atlas of Ostracod Shells 20, 3 Cytheromorpha diamphidia (3 of 4)

Reticulae are ordered concentrically peripherally; but centrally and dorsomedianly they are more

irregularly disposed. Posterior marginal rim narrow. Male valves are unusual in that their

ornament is not precisely complementary to that of the female valves: the males having a somewhat

punctate appearance medianly.

Remarks: This species is similar to Cytheromorpha macchesneyi (Brady & Crosskey) ( Geol . Mag., 8(80), 63,

pi. 2, figs. 1, 2, 1871), in overall shape and ornament. Like C. diamphidia, the ornament of C.

macchesneyi differs between sexes: males possessing more rounded and numerous punctae than

females. The two species differ in size (the type material of C. macchesneyi measures: length —

500 pm). The valves of the latter are also more tapered posteriorly and possess a series of

longitudinal furrows parallel with their free margins and a distinct, smooth area ventromedianly.

In addition, oral incurvature is better developed in the right valves of C. macchesneyi. In contrast

to the new species, C. macchesneyi is a distinct cold water indicator. It has not been recorded south

of latitude 62.46° N (see T.M. Cronin et al., U.S. Geol. Surv., Open File Report, 91-355, 51pp,

1992 for detailed distribution data). Subsequent to the publication of this report it has also been

described by Hartmann (Mitt. hamb. zoo/. Mus. Inst., 89, 185-186, pi. 1, figs. 5-10; pi. 2, fig. 1,

text-figs. 1-11, 1992) from Recent and subfossil material from N Spitzbergen. C. macchesneyi has

a general range from Quaternary to Recent.

Distribution: Upper Pliocene deposits of St. Erth, Cornwall, England (sample nos. 1-4, 7, 10-12, 14, 16, 18, 21,

23, 25-29) and Upper Pliocene (Redonian) deposits of Apigne (Le Temple du Cerisier), NW

France. See C. Maybury (op. cit.) and J.-P. Margerel, Les Foraminiferes du Redonien.

Systematique, Repartition stratigraphique, Paleoecologie, Nantes, 1, 8-26, 1968 for details of the

British and French samples, respectively.

Explanation of Plate 20, 4

Fig. 1, o* RV, int. lat. (OS 14219, 730 pm long); Figs. 2, 3, o' LV, (OS 14220, 690 ,um long): fig. 2, ant. hinge element; fig. 3, post, hinge

element; Fig. 4, 9 LV, muse. sc. (OS 14221, 620 pm long).

Scale A (200 //m; x91), fig. 1; scale B (100 /im; x220), figs. 2, 3; scale C (40 pm\ x512), fig. 4.

Cytheromorpha diamphidia (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 2

Stereo-Atlas of Ostracod Shells 20, 4

Cytheromorpha diamphidia (4 of 4)

Stereo-Atlas of Ostracod Shells 20 (2) 5-8 (1993) Semicytherura paraclausi (1 of 4)

595.337.14 (118.22) (420 : 162.006.50): 551.35

ON SEMICYTHERURA PARACLAUSI MAYBURY sp. nov.

by Caroline A. Maybury

(Institute of Earth Studies, University of Wales, Aberystwyth)

Holotype:

Type locality:

Derivation of name:

Figured specimens:

Semicytherura paraclausi sp. nov.

The Natural History Museum, London [BMNH] no. OS 14222; 9 RV.

[Paratypes nos. OS 14223-14225].

Sample no. 7, Vicarage Pit, St. Erth, Cornwall, England (50°26'W, 50° 10' N; Nat. Grid Ref. SW

556352); Upper Pliocene.

Latin, referring to the similarity of the new species to Semicytherura clausi (Brady, 1880) (Rep.

scient. Results. Voy. Challenger, Zoology, 1(3), 134, pi. 32, figs. 8a-d).

The Natural History Museum, London [BMNH] nos. OS 14223 (paratype, 9 LV: PI. 20, 6, fig. 1),

OS 14222 (holotype, 9 RV: PI. 20, 6, fig. 2), OS 14224 (paratype, cr LV: PI. 20, 6, fig. 3), OS

14225 (paratype, o * RV: PI. 20, 8, fig. 1), OS 14226 (paratype, cr LV: PI. 20, 8, fig. 2), OS 14228

(9 RV: PI. 20, 8, fig. 3), OS 14227 (9 LV: PI. 20, 8, fig. 4), OS 14229 (9 LV: PI. 20, 8, fig. 5).

All paratypes are from the same sample as the holotype, with the exception of paratype OS 14226

which is from sample no. 1, but from the type locality and horizon. See C.A. Maybury,

Taxonomy, Palaeoecology and Biostratigraphy of Pliocene Benthonic Ostracoda from St. Erth

and NW France, unpub. PhD thesis, Univ. Wales, 1, 3-6, 1985 for further sample details.

Explanation of Plate 20, 6

Fig. 1, 9 LV, ext. lat. (OS 14223, 530/im long); Fig. 2, 9 RV, ext. lat. (OS 14222, 560 //m long); Fig. 3, cr LV, ext. lat. (OS 14224,

600 pm long).

Scale A (200 pm; X110), figs. 1-3.

Stereo-Atlas of Ostracod Shells 20, 7 Semicytherura paraclausi (3 of 4)

Diagnosis: A small to medium sized Semicytherura with prominent keel-like alar process and posteroventral

concavity. Ornament of reticulae, punctae and micropunctae with 3 horizontal, anteromarginal

ridges. Pore conuli more numerous in the anterior half of the valve.

Remarks: The new species closely resembles Semicytherura clausi (Brady, op. cit.), a Recent species dredged

from Simon’s Bay, S Africa from depths of 15 to 20 fms and reported also from the coast of New

Zealand (N. de B. Hornibrook, Pal. Bull. N.Z. Geol. Survey, 18, 51, pi. 15, figs. 242-244, 1952).

The two species are similar in shape, size and gross ornamental configuration; but S. paraclausi

differs from S. clausi in its more variable ornament and in its possession of micropunctae and three

lateral ridges which terminate at the anterior margin. C. clausi has only two ridges.

Semicytherura reticulata Blondeau, 1971 (M.-A. Blondeau, pub. PhD thesis, Univ. Nantes, 86,

pi. 9, figs. 11-14), a Lutetian (Middle Eocene) species from Campbon, France is also similar and

perhaps ancestral to the new species. It differs from S. paraclausi in size (the holotype of S.

reticulata measures: length 370/ym and height 210/rm) and in the less pronounced development of

its alar process. It also bears a prominent, trifurcating lateral rib; which is absent in S. paraclausi.

Ornamental variants of S. paraclausi are illustrated in PI. 20, 8, figs. 3-5. These are less

reticulate than S. paraclausi sensu strictu and are smaller in size; characters which might could be

interpreted as juvenile, were not both forms represented by complete suits of instars and given the

adult development of the hinge and inner lamella as shown in PI. 20, 8, fig. 5.

Distribution: Upper Pliocene deposits of St. Erth, Cornwall, England (sample nos. 1-4, 7, 10-16, 18, 21-23,

25-29) and Upper Pliocene (Redonian) deposits of Apigne (Gite d’Apigne, Borehole II, Le Temple

du Cerisier), Beugnon (sample no. 1), Le Bosq d’Aubigny, Le Bosq d’Aubigny (Manche) and

Palluau II (670 cm), NW France. See C. Maybury (op cit.) and J.-P. Margerel, Les Foraminiferes

du Redonien. Systematique, Repartition stratigraphique, Paleoecologie, Nantes, 1, 8-26, 1968 for

details of British and French samples, respectively.

Explanation of Plate 20, 8

Fig. 1. o- RV, ext. lat. (OS 14225, 600 /un long); Fig. 2, or LV, int. lat. (OS 14226, 620 pm long); Fig. 3, 9 RV, ext. lat. (OS 14228,

460 /um long); Fig. 4, 9 LV, ext. lat. (OS 14227, 450/^01 long); Fig. 5, 9 LV, int. lat. (OS 14229, 450//m long).

Scale A (200 pm\ xllO), figs. 1-5.

Stereo-Atlas of Ostracod Shells 20, 6

Semicytherura paraclausi (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 8

Semicytherura paraclausi (4 of 4)

Stereo-Atlas of Ostracod Shells 20 (3) 9-12 (1993) Kiltsiella rosensteinae (1 of 4)

336.11 (113.331) (47: 161.023.58): 552.54 + 551.351

ON KILTSIELLA ROSENSTEINAE (SARV)

by David J. Siveter & Lembit I. Sarv

(University of Leicester, England & Institute of Geology, Estonian Academy of Sciences, Tallinn, Estonia)

Genus KILTSIELLA Sarv, 1968

Type-species: (by original designation): Craspedobolbina? rosensteini Sarv, 1962

Diagnosis: Finely reticulate, essentially non-lobate Zygobolbinae (?) with a diminutive slit-like adductorial sulcus. Lobel area

forms a low, gently curved profile above the hinge line. Velum flange-like, of more or less constant width between

cardinal corners in tecnomorphs. Cruminae discrete, unornamented, elongate sausage shaped, from mid-anterior

to just behind mid-venter; overhangs ventral margin in lateral and ventral views for most of its length; has velum

attached simply, both anteriorly and posteriorly. At its mid-length the crumina possibly breaks through the valve

margin. Torus lacking in both dimorphs.

Remarks: Although Sarv (1968 op. cit.) assigned Kiltsiella to the Zygobolbinae Ulrich & Bassler, 1923 (Beyrichiacea,

Palaeocopa), its subfamilial/familial taxonomic assignment is equivocal. Only one female specimen of Kilsiella

is known (PI. 20, 10, figs. 1-5), the subcruminal morphology of which seems to be broken away in the regions

of the valve margin adjacent to the ventral and anterior ends of the crumina. Alternatively, this arrangement

could be the true morphology of the species; if so, it would represent a narrowing and loss of a strip of the valve

margin by virtue of the (diagnostically zygobolbinine) encroachment of the adjacent (ventral and anterior) parts

of the crumina. However, no clear dolonoid scar ( sensu Martinsson, Bull. geol. Instn Univ. Uppsala., 41, 1962),

in the form either of a fold or fissure, is apparent in this subcruminal region in the female in question, thus

suggesting that damage is the more likely explanation for the observed morphology.

In lateral view the lobal, cruminal and velar morphology and ornament of Kiltsiella is very similar to the

beyrichiacean craspedolbolbinine Clintiella Martinsson, 1962. Tecnomorphs of Kiltsiella also have much in

Explanation of Plate 20, 10

Figs. 1-5, 9LV (Os 5163, 1050/im long); fig. 1, post.; fig. 2, ext. lat.; fig. 3, syllobial ornament; fig. 4, int. obi. vent.; fig. 5, vent.

Scale A (200 //m; x48), figs. 1, 2, 4, 5; scale B (50 pm\ xl45), fig. 3.

Stereo- Atlas of Ostracod Shells 20, 12

Kiltsiella rosensteinae (3 of 4)

common with beyrichiacean treposellid tecnomorphs belonging to genera such as Garniella and especially

Retisacculus (both Martinsson 1962). Kiltsiella differs from both craspedobolbinines and treposellids in apparently

lacking traces of the dolonoid closing mechanism of the crumina (i.e. dolonoid scars or “treposelline bridges”

respectively).

Kiltsiella rosensteinae (Sarv, 1962)

1962 Craspedobolbina? rosensteini (sic) sp. nov. L.I. Sarv, Eesti NSV Tead. Akad. Geol. Inst. Uurim., 9, 126, pi. 7, figs. 5-9.

1968 Kiltsiella rosensteinae (Sarv); L.I. Sarv, Ostracode families Craspedobolbinidae, Beyrichiidae and Primitiopsidae in the Silurian of Estonia. Eesti

NSV Tead. Akad. Geol. Inst. Tallinn, 30, pi. 3, figs. 7-9.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

Distribution:

A cknowledgement:

Institute of Geology, Estonian Academy of Sciences, Tallinn, no. Os 5162; male left valve.

Old quarry near Kiltsi, about 3 km SW of Haapsalu, approx, lat. 58°58'N, long, 23°32'E, NW Estonia. Juuru

regional “stage” (G1-2), lower part of the Llandovery Series, Silurian.

Institute of Geology, Estonian Academy of Sciences, Tallinn, nos. Os 5162 (holotype, a LV: PI. 20, 12, figs. 1-4),

Os 5163 (9 LV; pi. 20, 10, figs. 1-5) and Os 5172 (tecnomorph LV: PI. 20, 12, figs. 5, 6). All topotye specimens.

As for the genus (monotypic).

Specimens Os 5162 and Os 5163 are the originals of Sarv 1962, pi. 7, figs. 8 and 9 respectively. The original of

Sarv 1962, pi. 7, figs. 5-7 (Os 5164: small tecnomorphic carapace) is now lost.

As noted by Sethi (in: Jaanusson, V. et al. [eds.], Sver. geol. Unders. Afh., ser. C, 272, 162), the smooth/finely

punctate Kiltsiella sarvi Copeland (Bull. geol. Surv. Can., 241, 23, 1974) has a much less well defined crumina

than K. rosensteinae and should be assigned to another genus.

K. rosensteinae is the only known beyricheacean ostracod from the type locality, where it associates are mostly

unstudied podocopids including Medianella cf. lubricei (Stumber). Bolbiprimitial tamsaluensis Sarv, 1962 and

A it ilia senecta Sarv, 1968 are two beyrichiaceans which occur at a similar horizon to K. rosensteinae, but on the

island of Huumaa in northern Estonia.

Known only from the type locality; about 15 specimens, collected by E. Rosenstein in about 1938.

Support from the NATO collaborative research programme is gratefully acknowledged.

Explanation of Plate 20, 12

Figs. 1-4, cr LV (holotype, Os 5162, 1030/rm long): fig. 1, post.; fig. 2, ext. lat.; fig. 3, ant.; fig. 4, vent. Figs. 5, 6, tecnomorphic LV

(Os 5172, 800 /2m long): fig. 5, ext. lat.; fig. 6, ant.

Scale A (200 pm; x48), figs. 1-4; scale B (200 pm; x57), figs. 5, 6.

Stereo-Atlas of Ostracod Shells 20, 10

Kiltsiella rosensteinae (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 12

Kiltsiella rosensteinae (4 of 4)

Stereo-Atlas of Ostraeod Shells 20 (4) 13-16 (1993) Sulcella huecoensis (1 of 4)

595.337.3 (113.61) (789 : 162.003.32): 551.351 + 552.52

ON SULCELLA HUECOENSIS DEWEY & KOHN sp. nov.

by Christopher P. Dewey & Peter Kohn

(Mississippi State University, Mississippi, U.S.A.)

Holotype:

Type locality:

Derivation of name:

Figured specimens:

Sulcella huecoensis sp. nov.

Dunn Seiler Museum of Geology, Mississippi State University, U.S.A., no. 3341-9a; adult female

carapace.

[Paratypes nos. 3341-9b, 3341-9c, 3341-9(1; one adult female carapace, one female right valve

and one juvenile carapace].

Unnamed canyon. Sec. 30, T22S, R1E, Dona Ana County, Picacho Mountain Quadrangle, New

Mexico, U.S.A. ; lat. 32°21'51"N, long. 106°52'44"W. Upper part of the Hueco Formation,

Wolfcampian, Lower Permian; 100.84 m above the base of the measured section, in olive-grey

shale. Shale contains spirorbid worms, fenestrate bryozoans, crinoid columnals, small rostroconchs

and endothyrid foraminifera. Shallow marine.

After the Hueco Formation, being the stratigraphic unit in which this species was first recorded.

Dunn-Seiler Museum of Geology, Mississippi State University, U.S.A., nos. 3341-9a (holotype

adult car.: PI. 20, 14, figs. 1-4), 3341-9b (adult car.: PI. 20, 16, figs. 5-7), 3341-9c (9 RV: PI. 20,

16, fig. 4), 3341-9d (juv. car.: PI. 20, 16, figs. 1-3). All from olive-grey fossiliferous shale, side of

canyon wall, at the type locality.

Explanation of Plate 20, 14

Figs. 1-4, adult car. (holotype, 3341-9a, 0.72 mm long): fig. 1, RV ext. lat.; fig. 2, ext. dors.; fig. 3, ext. vent.; fig. 4, LV ext. lat.

Scale A (250 /urn; x72), figs. 1-4.

Stereo-Atlas of Ostraeod Shells 20, 15

Sulcella huecoensis (3 of 4)

Diagnosis:

Remarks:

Distribution:

A ckn o w ledge men t:

Small, elongate, ellipsoidal carapace. Dorsal margin broadly arched, gently sloping towards

anterior. Anterior end evenly rounded, posterior margin bluntly pointed. Posteroventral border

slopes steeply from just above midheight to ventral margin. Slight concavity in midventral outline

of right valve. Right valve overlaps left, overlap conspicuous around entire margin. Greatest

overlap along ventral margin. Internal contact groove around free margin of right valve. Hinge

stragulate. Small, shallow, pit-like sulcus located just above midheight and anterior of midlength.

Dorsal margin of right valve pinched above and anterior of sulcus, most pronounced in females.

Dimorphic; females cuneate in dorsal aspect, maximum width posterior, with interior limen.

Surface ornament consists of small-celled, thin-walled reticulation.

According to Benson et al. ( Treatise on Invertebrate Paleontology, Geol. Soc. Amer. & Kansas

Univ. Press, Q370, 1961), the cavellinid genus Sulcella Coryell & Sample, 1932 only occurs in

Carboniferous strata. The presence of Sulcella huecoensis in the Hueco Formation of New Mexico

therefore extends the range of the genus into the Lower Permian. Of congeneric taxa only Sulcella

cf. indistincta (Tschigova, 1958) sertsu Robinson ( Geol . J. (Special Issue) 8, 134, pi. 3, figs. 6a-b,

1978), from the Tournaisian, Lower Carboniferous of England, is known to have a reticulate

surface. S. huecoensis differs from S. cf. indistincta in lateral outline. In particular, the outline of

S. huecoensis has a greater length to height ratio than 5. cf. indistincta, is more rounded and less

quadrate, has a more pronounced posterior acumination and possesses a medial concavity in the

ventral margin of the right valve.

U.S.A.; Hueco Formation, Wolfcampian Series, Lower Permian.

CD acknowledges the financial support given by the Donors of the Petroleum Research Fund

administered by the American Chemical Society.

Explanation of Plate 20, 16

Figs. 1-3, juv. car. (paratype, 3341-9d, 0.62 mm long): fig. 1, ext. dors.; fig. 2, LV ext. lat.; fig. 3, RV ext. lat. Fig. 4, 9 RV (paratype

3341-9c, 0.66 mm long): RV int. (note post, limen). Figs. 5-7 adult car. (paratype 3341-9b, 0.70 mm long): fig. 5, ext. dors.; fig. 6,

LV ext. lat.; fig. 7, RV ext. lat.

Scale A (250/rm; x72), figs. 1-7.

Stereo-Atlas of Ostracod Shells 20, 14

Stereo-Atlas of Ostracod Shells 20, 16

Sulcella huecoensis (4 of 4)

Sulcella huecoensis (2 of 4)

Stereo-Atlas of Ostracod Shells 20 (5) 17-24 (1993) Nipponocythere colalongoae (1 of 8)

595.337.14 (119) (265.7 : 163.141.39): 551.353 + 552.52

ON NIPPONOCYTHERE COLALONGOAE (CIAMPO)

by Victoria Drapala & Michael A. Ayress

(Department of Geology, Australian National University, Canberra)

Nipponocythere colalongoae (Ciampo, 1986)

1976 Nipponocythere sp. K. Ishizaki & F.J. Gunther, Sci. Rep. Tohoku Univ. 2nd Ser. (Geol.), 46, 25, pi. 9, fig. 15; pi. 10, figs.

15-17.

1986 Flexuocythere colalongoae n. sp. G. Ciampo, Boll. Soc. paleont. ital., 24, 85, pi. 6, figs. 4-6, pi. 18, fig. 5.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Department of Earth Sciences, University of Naples, Italy; COC n. 412; male RV.

Rio Mazzapiedi section, Piedmont, Italy; Globorotalia mediterranea subzone, Messinian, Upper

Miocene.

National Museum of Victoria (Australia) nos. P 197916 (o* car. (disarticulated into LV & RV): PI.

20, 18, figs. 1, 2; PI. 20, 20, fig. 2, PI. 20, 24, fig. 2), P 197917 (9 RV: PI. 20, 18, fig. 3; PI. 20,

20, fig. 1; PI. 20, 22, figs. 1, 2), P 197918 (o* LV: PI. 20, 20, fig. 3; PI. 20, 24, fig. 1; text-fig. 1).

All from Eltanin core PC55-6; Otway Basin, off Victoria, Australia (lat. 38°51'S, long.

141° 03' E); Late Quaternary foraminiferal-nannofossil ooze, water depth 2346 m. Specimen P

197916 is from interval 29-30 cm, P 197917 is from 39-40 cm and P 197918 is from 99-100 cm.

A moderately well-inflated, subtriangular to sub-rectangular species of Nipponocythere with fine

punctation mainly confined to the centre of the valve and very fine reticulation around the margins;

smooth elsewhere. Short posterodorsal ridge reaches above hinge line. Females higher than males.

Explanation of Plate 20, 18

Fig. 1, o* car., ext. lat. (P 197916, 400 /rm long); Fig. 2, O’ car. dors. (P 197916, 400 pm long); Fig. 3, 9 RV, ext. lat. (P 197917, 400 pm

long).

Scale A (100 pm; xl40), figs. 1-3.

Stereo-Atlas of Ostracod Shells 20, 19 Nipponocythere colalongoae (3 of 8)

Remarks: Nipponocythere is closest to Heinia Bold, 1985 (J. Paleont., 59, 1). These genera share the same

modified gongylodont hinge type, both having asymmetrical posterior elements. A comparison of

the type species of both genera show distinct differences in shape; Nipponocythere bicarinata

(Brady, 1880) {Rep. scient. Results Voy. Challenger, (Zoology), 1(3), 70, pi. 16, figs. 6a-d) is ovate

in lateral outline and the inflation is relatively even, whereas Heinia howei has a more rectangular

outline and strongly compressed marginal regions. However, the outline of some other species

included in Nipponocythere is very similar to that of certain other species assigned to Heinia e.g.

a rectangular outline in Nipponocythere nagaseae Ishizaki & Gunther, 1976 (Sci. Rep. Tohoku

Univ., 2nd Ser. (Geol.), 46, 24) and Heinia caudata (Bold, 1966) ( Verb . K. ned. Akad. Wet., (1),

23, 20); or a subtriangular shape in Nipponocythere sp. Tabuki, 1986 (Bull. Coll. Ed., Univ.

Ryukyus, 29, 105) and Heinia sp. aff. H. howei Bold (J. Paleont., 59, 6).

After examination of the type specimens of Heinia howei and a consideration of all published

species of both genera, we conclude that Heinia is perhaps separable on the basis of the enlarged

upper two adductor muscle scars (see Ayress & Correge, Stereo-Atlas Ostracod Shells, 20, 25-28,

1993), it usually lacks a ventral ridge and the reticulation is usually more strongly developed.

Several species of Nipponocythere, including N. colalongoae, have an elongated dorsomedian

adductor muscle scar, which may represent an intermediate stage in development towards the

Heinia muscle scar pattern. However, these differences are probably insufficient to separate at

generic level and we therefore consider Heinia to be a junior synonym of Nipponocythere.

Loxoconchidea Bonaduce, Ciampo & Masoli, 1975 (Pubbl. Staz. zool. Napoli, 40, 112) also has

the same hinge type and is presumably closely related. It differs from Nipponocythere in having a

symmetrically convex posterior.

Explanation of Plate 20, 20

Fig. 1, 9 RV, int. lat. (P 197917, 400 /um long); Fig. 2, o’ car., dissarticulated LV int. lat. (P 197916, 400 pm long); Fig. 3 o ■ LV,

adductor muse, scar detail (P 197918, 400 pm long).

Scale A (100 gm; xl40), figs. 1, 2; scale B (20 x650), fig. 3.

Stereo-Atlas of Ostracod Shells 20, 18

Nipponocythere colalongoae (2 of 8)

Stereo-Atlas of Ostracod Shells 20, 20

Nipponocythere colalongoae (4 of 8)

Stereo-Atlas of Ostracod Shells 20, 21 Nipponocythere colalongoae (5 of 8)

Although details of the muscle scars have not been described or illustrated, the type species of

Flexuocythere Ciampo, 1986 {Boll. Soc. paleont. ital., 24, 85), “ Buntonia ” parva Colalongo

& Pasini, 1980 {Boll. Soc. paleont. ital., 19, 68), is considered by us to belong to Heinia

(= Nipponocythere).

Nipponocythere sp. Tabuki, 1986 {Bull. Coll. Ed., Univ. Ryukyus, 29, 105, pi. 19, figs. 14, 15;

text-fig. 19-7) is similar to N. colalongoae but can be distinguished by its slightly more triangular

outline, its narrower posterior and slightly stronger ornamentation. The female specimen of N.

nagaseae Ishizaki & Gunther {op. cit.), is also very similar in shape to N. colalongoae. However,

the male of N. nagaseae is much more elongate and also differs in having more extensive

ornamentation.

Distribution: Previous records of this species are from bathyal sediments (1404 m) of the Gulf of Panama

(Ishizaki & Gunther, op. cit.) and from the Pleistocene of Italy (Ciampo, op. cit.). We have found

it in Late Quaternary sediments from deep-sea cores surrounding Australia: off Victoria, Eltanin

core PC55-6, 38°51.2'S, 141°33.8'E, water depth 2346m; Tasman Sea, BMR core 71 GC044,

29° 30.90'S, 153° 54.79', water depth 1298 m; Timor Sea ODP Site 262, 10° 52. 19' S, 123° 50.78' E,

present water depth 2298 m.

Acknowledgements: We would like to thank the Electron Microscope Unit (ANU) for their assistance and use of their

scanning electron microscopes.

Explanation of Plate 20, 22

Figs. 1, 2, 9 RV (P 197917, 400 /vm long): fig. 1, ant. hinge detail; fig. 2, post, hinge detail.

Scale A (20 /tm; x800), figs. 1, 2.

Stereo-Atlas of Ostracod Shells 20, 23 Nipponocythere colalongoae (7 of 8)

Text-fig. 1, Internal features observed through transmitted light. Male LV (P 197918, 400 pm long).

Explanation of Plate 20, 24

Fig. 1, cr LV, post, hinge detail (P 197918, 400 pm long); Fig. 2, cr car., dissarticulated LV, ant. hinge detail (P 197916,

400 pm long).

Scale A (20 x750), fig. 1; scale B (20 //m; X1500), fig. 2.

Stereo-Atlas of Ostracod Shells 20, 22

Nipponocythere colalongoae (6 of 8)

Stereo-Atlas of Ostracod Shells 20, 24

Nipponocythere colalongoae (8 of 8)

Stereo-Atlas of Ostracod Shells 20 (6) 25-28 (1993) Nipponocythere cuneata (1 of 4)

595.337.14 (119.1) (265.7 : 163.146.25): 551.352 + 552.52

ON NIPPONOCYTHERE CUNEATA AYRESS & CORREGE sp. nov.

by Michael A. Ayress & Thierry Correge

(Department of Geology, Australian National University, Canberra)

Nipponocythere cuneata sp. nov.

1985 Heinia sp. aff. H. howei Bold, J. Paleont., 59, 6, figs. 6.6, 6.8, 6.9.

Holotype:

Type locality:

Derivation of name:

Figured specimens:

Diagnosis:

Remarks:

National Museum of Victoria, Melbourne, Australia, no. P 197927.

Coral Sea, ODP Site 822, core 3, section 5, interval 60-62 cm, water depth 955 m. Latitude

16° 25.379' S, longitude 146° 12.904' E. Late Pleistocene foraminiferal ooze.

From Latin, cuneatus, wedge-shaped; referring to the outline in lateral view.

National Museum of Victoria, Melbourne, Australia, nos. P 197927 (holotype, LV: PI. 20, 26, figs.

1,3, PI. 20, 28, fig. 2), P 197928 (paratype, RV: PI. 20, 26, fig. 2, PI. 20, 28, figs. 1, 3); both from

the type locality.

A wedge-shaped species-of Nipponocythere with dense reticulation, in which the horizontal muri

predominate and secondary punctation anteriorly. A short dorsal rib overreaches dorsal margin

posterodorsally. Caudal process well developed subventrally. Sexual dimorphism not apparent.

The type specimens of Heinia howei Bold, 1985, the type species of Heinia, have been examined

by us and display internal features identical to those of N. cuneata. In addition to Bold’s

observations on Heinia, the asymmetrical posterior hinge elements, in which the tooth in the RV

Explanation of Plate 20, 26

Fig. 1, LV, ext. lat. (P 197927, 353 pm long); Fig. 2, RV, ext. lat. (P 197928, 353 pm long); Fig. 3, LV, subcentral muse, scars

(P 197927, 353 //m long).

Scale A (100 ^m; xl70), figs. 1, 2; scale B (lO/zm; X1000), fig. 3.

Stereo-Atlas of Ostracod Shells 20, 27 Nipponocythere cuneata (3 of 4)

is positioned in the anterior half of the socket and the enlarged upper two adductor muscle scars

appear also to be characteristic of Heinia. The former feature is shared with Nipponocythere

Ishizaki (1971, Sci. Rept. Tohoku Univ., 2nd Ser. (Geol.), 43, 88) and only the latter feature

appears to be unique to Heinia. We regard this sole feature insufficient to separate the two genera

and thus we consider Heinia to be a junior synonym of Nipponocythere (for further discussion of

affinities see Drapala & Ayress, Stereo-Atlas Ostracod Shells, 20, 17, 1993). The dorsal surface of

the posterior terminal hinge element of N. howei and N. cuneata sometimes has weak lobation but

this is never developed as strongly as it is in Kuiperiana (considered here to be the senior synonym

of Myrena Neale 1967, Scient. Rep. Br. Antarct. Surv., 58, 19).

TV. cuneata differs from TV. howei mostly in its more triangular lateral outline and stronger

ornament in the posterior half of the carapace. A closely similar species, TV. parva (Colalongo &

Pasini, 1980) (Boll. Soc. paleont. ital., 19, 68, pi. 21, fig. 9) from the Pleistocene of Calabria, Italy,

differs in its more rectangular shape and weaker ornament. TV. caudata (Bold, 1985) is also some-

what similar but that species is more rectangular and its reticulum is more regularly developed

lacking secondary punctation. Other species recorded by Bold, 1985 (op. cit.) and left in open

nomenclature, differ mainly in detail of the surface ornament.

Distribution: Bold (1985) recorded this species in the Pliocene of the Caribbean and Gulf of Mexico. We have

encounted it in the late Pliocene of ODP Site 815, water depth 465.5 m and in the late Pleistocene

of ODP Site 822, water depth 955 m, both in the Coral Sea, SW Pacific.

Acknowledgements: We would like to thank the staff of the Electron Microscope Unit (ANU) for their technical

assistance and Professor Whatley for critically reviewing the manuscript.

Explanation of Plate 20, 28

Fig. 1, RV, int. lat. (P 197928, 353 pm long); Figs. 2, 3, LV (P 197927, 353 pm long): fig. 2, int. lat.; fig. 3, post, hinge.

Scale A (100//m; xl70), figs. 1, 2; scale B (10/ym; X1200), fig. 3.

Nipponocythere cuneata (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 28

Nipponocythere cuneata (4 of 4)

Stereo-Atlas of Ostracod Shells 20, 26

Stereo-Atlas of Ostracod Shells 20 (7) 29-32 (1993) Kuiperiana dryppa (1 of 4)

595.337.14 (119.1) (265.7 : 163.138.34+ 141.39): 551.352 + 552.52

ON KUIPERIANA DRYPPA (WHATLEY & COLES)

by Michael A. Ayress & Victoria Drapala

(Department of Geology, Australian National University, Canberra)

Kuiperiana dryppa (Whatley & Coles, 1987)

1987 Heinia dryppa sp. nov. R.C. Whatley & G. Coles, Revta esp. Micropaleont., 19, 75, pi. 4, figs. 20-23.

1988 Palmoconchal sp. 2 R.C. Whatley & M.A. Ayress, in Hanai, T. (et al.) (Eds.), Evolutionary Biology of Ostracoda, its

Fundamentals and Applications, Kodansha, Tokyo, etc., 742, p. 2, figs. 10a, b.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

The Natural History Museum, London [BMNH] no. OS 12495; 9 RV.

DSDP Site 607 (lat. 41 ° 00.07' N, long. 32° 47.44' W); core. Water depth 3427 m. Late Pliocene (NN

Zone 16).

National Museum of Victoria (Australia) nos. P 197919 (adult RV: PL. 20, 30, fig. 1) and P 197920

(adult RV: PI. 20, 32, figs. 1, 2, 3) from intervals 35-36 cm and 56-57 cm respectively of BMR core

67 GC03, lat. 37° 33.0'S, long. 138° 35.0' E; P 197921 (adult LV: PI. 20, 30, figs. 2, 3) from interval

24-25 cm of Eltanin core PC55-6, lat. 38°51.2'S, long. 141°03.8'E.

A species of Kuiperiana with wide compressed margins and ornament of reticulae and secondary

punctae in which the horizontal muri predominate. Caudal process moderately well developed.

Explanation of Plate 20, 30

Fig. 1, adult RV, ext. lat. (P 19719, 440 pm long); Figs. 2, 3, adult LV (P 197921, 450 pm long): fig. 2, int. lat.; fig. 3, ant. hinge detail.

Scale (100 pm; xl30), figs. 1, 3; scale B (20 pm\ x600), fig. 3.

Stereo-Atlas of Ostracod Shells 20, 31

Kuiperiana dryppa (3 of 4)

Remarks:

Distribution:

A ckn o w ledge men ts:

The well preserved specimens of this species, which we have encountered in deep-sea cores adjacent

to Australia, show clearly the detail of internal features important for a consideration of the generic

placement of this species. In previous works K. dryppa has been placed in Heinia and Palmoconcha.

However, we show here that the hingement of this species is inconsistent with those genera. Both

lack the strongly denticulate dorsal border of the posterior tooth in the right valve, and on this basis

it clearly belongs to Kuiperiana. Heinia (= Nipponocy there, see Drapala & Ayress, Stereo-Atlas

Ostracod Shells, 20, 17-24, 1993) also differs significantly in its ventral caudal process, in its

enlarged upper two adductor scars (see Ayress & Correge, Stereo-Atlas Ostracod Shells, 20, 25-28,

1993) and in its discontinuous selvage.

This species is distributed worldwide in the deep-sea. We have recorded it in Pleistocene cores in the

eastern Indian Ocean and Tasman Sea over a depth range of 1321 m to 2346 m.

We would like to thank the Electron Microscope Unit (ANU) for their assistance and use of their

scanning electron microscopes.

Explanation of Plate 20, 32

Figs. 1-3, adult RV (P 197920, 430 pm long), fig. 1, int. lat.; fig. 2, post, hinge detail; fig. 3, adductor muse, scar detail.

Scale A (100 pm; xl40), fig. 1; scale B (20 pm; x600), fig. 2; scale C (20 pm; x550), fig. 3.

Stereo-Atlas of Ostracod Shells 20, 30

Kuiperiana dryppa (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 32

Kuiperiana dryppa (4 of 4)

Stereo-Atlas of Ostracod Shells 20 (8) 33-36 (1993) Aboilia blessi (1 of 4)

595.337.1 (113.31) (71 : 162.047.50): 551.351 + 552.55

ON ABOILIA BLESSI BECKER & ADAMCZAK gen. et sp. nov.

by Gerhard Becker & Franciszek F. Adamczak

(University of Frankfurt am Main, Germany & University of Stockholm, Sweden)

Genus ABOILIA gen. nov.

Type-species: Aboilia blessi sp. nov.

Play on the English term “a boil”; referring to the fanciful resemblance of the dorsal projection of the

small valve. Gender, feminine.

Strongly inequivalved podocopine genus with right valve larger and having a distinct bow-shaped

projection (ventriculum) overlapping the left valve ventrally. Left valve provided with a pronounced,

poster odor sally situated ridge-like swelling, occasionally overreaches the hinge margin. Dorsal margin

convex (as seen in lateral aspect), whereas ventral margin is concave. Right hinge nearly tripartite in

appearance.

The bow-shaped projection, the almost tripartite hinge situated in a depression, the carapace outline, and

the protruding end margins of the right valve of A. blessi indicate pachydomelmellid affinities (see

Adamczak, F.J., Senckenberg. leth., 57, 332, 1976). However, the hitherto known representatives of this

family characteristically show left-over-right overlap; thus, a reversed overlap for the early podocopines

is reported here for the first time, from Aboilia.

The distinct valve asymmetry of Aboilia is not exceptional for the podocopines, several representatives

of which ( Pseudorayella Neckaja, 1960, Bairdiocypris Kegel, 1932, Pachydomella Ulrich, 1891) show this

characteristic. However, the reason for this phenomenon may only be speculated upon; possibly it has

Derivation of name:

Diagnosis:

Remarks:

Explanation of Plate 20, 34

Figs. 1-3, adult car. (holotype, RGM 414.009, 580 pm long): fig. 1, It. lat.; fig. 2, vent, obi.; fig. 3, post. Figs. 4, 5, adult car. (paratype,

RGM 414.010, 640 pm long): fig. 4, It. lat.; fig. 5, vent. obi.

Scale A (100 /mi; xl07), figs. 1-5.

Stereo-Atlas of Ostracod Shells 20, 35 Aboilia blessi (3 of 4)

something to do with the absence, in larger valve of these forms, of an effective stop-structure (which

could arrest possible excessive overlap).

Holotype:

Derivation of name:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

Distribution:

Aboilia blessi sp. nov.

Nationaal Natuurhistorisch Museum (RGM), Leiden, The Netherlands, no. RGM 414.009; an adult

carapace.

In honour of Dr Martin Bless (Heerlen), in recognition of his research on Palaeozoic Ostracoda.

From a seamount (Orphan Knoll, see Ruffman, A. & van Hinte, J.E., Geol. Surv. Pap. Can., 71-23,

407, 1973), in the Labrador sea, approximately 500 km NE of Newfoundland. The material was obtained

from a single dredge (LYNCH 7/11/71 cruise, biological dredge station no. D3-7- 1 1-71) at c.0140 GMT,

May 23rd 1971, at an average position of lat. 50°33'N, long. 46°22'W and an average depth of 1775 m

(see Ruffman, A., Geol. Surv. Canada, open file 2065, 1989). The silicified ostracods come from a single

limestone pebble of middle to Upper Ordovician age.

Nationaal Natuurhistorisch Museum (RGM), Leiden, The Netherlands, nos. RMG 414.009 (adult car.,

holotype: PI. 20, 34, figs. 1-3; PI. 20, 36, fig. 4), RGM 414.010 (adult car., paratype: PI. 20, 34, figs.

4, 5; PI. 20, 36, fig. 2) and RGM 414.011 (adult RV, paratype: PI. 20, 36, figs. 1, 3, 5).

All of the figured specimens are from the type locality.

As for the genus, which is monotypic.

The dorsal swelling is present in both adult and juvenile specimens in our material (13 specimens). In

adult specimens variation of the size of the dorsal swelling occurs; the swelling influences the lateral

outline of the valve if it is particularly pronounced and overreaches the dorsal margin (see PI. 20, figs.

1, 3). This variation may be both intraspecific and a product of variable preservation.

The species is considered to be benthic.

Only known from the type locality; middle to upper Ordovician.

Explanation of Plate 20, 36

Figs. 1,3,5, adult RV (paratype, RGM 414.011, 630 pm long): fig. 1, int. lat.; fig. 3, int. vent, obi., ventriculum; fig. 5, int. dors, obi.,

hinge structure. Fig. 2, adult car., dors, (paratype, RGM 414.010, 640 pm long). Fig. 4, adult car., vent, (holotype, RGM 414.009,

580 pm long).

Scale A (100 pm; xl07), figs. 1-5.

Aboilia blessi (2 of 4)

Aboilia blessi (4 of 4)

Stereo-Atlas of Ostracod Shells 20, 34

Stereo-Atlas of Ostracod Shells 20, 36

Stereo- Atlas of Ostracod Shells 20 (9) 37-40 (1993) Baltonotella elegans (1 of 4)

595.336 (113.312) (766 : 162.097.34): 551.351 + 552.54

ON BALTONOTELLA ELEGANS (HARRIS)

by Mark Williams & Jean Vannier

(University of Leicester, England & Universite Claude Bernard, Lyon, France)

Baltonotella elegans (Harris, 1957)

1957 Macronotella elegans n. sp., R.W. Harris, Bull. Okla. geol. Surv., 75, 184, pi. 4, figs. 12a-c.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Museum of Comparative Zoology, Harvard University, U.S.A., no. MCZ 4574; a juvenile

carapace.

From Decker’s Bed 64 (see Harris 1957, op cit.), Oil Creek Formation, Simpson Group, middle

Ordovician. West Spring Creek locality, Secs. 7 and 8, T2S, R1W, Arbuckle Mountains,

Oklahoma, U.S.A.; approximately lat. 34°30'N, long. 97°20'W.

Museum of Comparative Zoology (MCZ) , Harvard University, U.S.A., no. MCZ 4574 (juv. car.

holotype: PI. 20, 38, figs. 1-4). Natural History Museum, London [BMNH] no. OS 13637 (car.: PI.

20, 40, figs. 1-4). MCZ 4574 is from the type locality and horizon. OS 13637 is from the Oil Creek

Formation, collected approximately 100 m above the base of the Formation at the type locality.

Both valves with coarse puncta concentrically arranged around a central smooth (adductor muscle)

spot. Marginal areas smooth. Left valve with a row of marginally situated minute denticles. Right

valve with an overlap frill demarcated from the ventral valve surface of the right valve by a groove.

Overlap contact straight.

Explanation of Plate 20, 38

Figs. 1-4, juv. car. (holotype, MCZ 4574, 0.64 mm long): fig. 1, LV ext. lat.; fig. 2, RV ext. lat.; fig. 3, dors, obi.; fig. 4, RV ext. lat.

obi.

Scale A (150/rm; x84), figs. 1-4.

Stereo-Atlas of Ostracod Shells 20, 39

Baltonotella elegans (3 of 4)

Remarks:

Distribution:

A ckno wledgements:

Baltonotella elegans clearly differs from Macronotella , which is dimorphic (see Kesling et al.,

Contr. Mus. Paleont. Univ. Mich., 10, 83-100, 1960). B. elegans is very similar to Baltic species of

Baltonotella, particularly to the type species Baltonotella kuckersiana (Bonnema, 1909) (see Sarv

1959, Eesti NSV Tead. Akad. Geol. Instit. Uurim., 4, pi. 32, fig. 17).

Features of B. elegans which are characteristic of all Baltonotella species include the right over

left valve overlap, the straight contact margin, the presence of marginal structures on the left valve,

the overreach of the left valve over the right valve dorsally and the subcircular carapace outline. In

addition, species of Baltonotella are among the most widespread leiocopes (Aparchitidae) in the

Ordovician of N America and Europe.

Adults and juveniles of B. elegans show great difference in the development of ornament, with

punctation being much reduced in adults. Variation of ornament between adults and juveniles

appears to be a common feature of Baltonotella species.

Only in the Oil Creek Formation, Simpson Group, middle Ordovician, Arbuckle Mountains,

Oklahoma, U.S.A.

M. Williams acknowledges support from N.E.R.C., the Alexander von Humboldt Foundation,

Bonn, and the Universite Claude Bernard, Lyon. Dr J.M. Berdan and Mr M.A. Miller are thanked

for loan of specimens.

Explanation of Plate 20, 40

Figs. 1-4, car. (OS 13637, 1.30 mm long): fig. 1, LV ext. lat.; fig. 2, vent.; fig. 3., ant.; fig. 4, close-up of marginal structure (LV = top).

Scale A (200 gm; X39), figs. 1, 2; scale B (200 pm; x42), fig. 3; scale C (500 pm; x 117), fig. 4.

Baltonotella elegans (2 of 4)

Baltonotella elegans (4 of 4)

Stereo-Atlas of Ostracod Shells 20, 38

Stereo-Atlas of Ostracod Shells 20, 40

Stereo-Allas of Ostracod Shells 20 (10) 41-44 (1993) Kayina hybosa (1 of 4)

595.336 (113.312) (776 : 162.096.34): 551.351 + 552.54

ON KA YIN A HYBOSA HARRIS

by Mark Williams & Jean Vannier

(University of Leicester, England & Universite Claude Bernard, Lyon, France)

Diagnosis:

Remarks:

Distribution:

Genus KA YIN A Harris, 1957

Type-species (by original designation): Kayina hybosa Harris, 1957

Postplete to sub-amplete. Valves strongly asymmetric; left valve having a pronounced posterodorsal node, can

extend above dorsal margin. Left valve weakly overlaps right valve. Contact margin straight. No sulcation or

adventral structures.

Schallreuter (NeuesJb. Geol. Palaont., Mh., 11, 690, 1971) assigned Kayina to his leiocope family Jaanussoniidae,

based primarily on valve asymmetry manifested by a posterodorsally situated node on the left valve. However,

Kayina does not conform to the typical morphology of jaanussoniids (Vannier, Lethaia, 23, 103, 1990): it has

a leperditellid outline (its width and height increase posteriorly) and its carapace is far more elongate than typical

jaanussoniids; it shows left over right valve overlap (the opposite is normal in jaanussoniids); its degree of

overlap (Text-fig. 1) is, unusually for leiocopes, very slight. Kayina appears to be more closely related to the non-

leiocope Leperditella (Berdan, Mem. Bur. Mines Mineral Resourc., New Mex., 44, 287, 1988).

Middle Ordovician of U.S.A. and possibly the Baltic region.

Kayina hybosa Harris, 1957

1957 Kayina hybosa n. sp., R.W. Harris, Bull. Okla. geol. Surv., 75, 160, pi. 3, figs, lla-d.

1965 Kayina hybosa Harris; M.J. Copeland, Bull. geol. Surv. Can., 127, 38.

1971 Kayina hybosa Harris; R.E.L. Schallreuter, Neues. Jb. Geol. Palaont. Mh., 11, 254.

1988 Kayina hybosa Harris; J. Vannier, Stereo-Atlas Ostracod Shells, 15, 139.

Explanation of Plate 20, 42

Figs. 1-3, car. (holotype, MCZ 4530a, 1.08 mm long); fig. 1, LV ext. lat.; fig. 2, LV ext. lat. obi.; fig. 3, RV ext. lat.

Scale A (150/rm; x57), figs. 1-3.

Stereo-Atlas of Ostracod Shells 20, 43 Kayina hybosa (3 of 4)

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

Distribution:

A cknowledgements:

Museum of Comparative Zoology, Harvard University, U.S.A. , no. MCZ 4530a; a carapace.

From Decker’s Bed 36 (see Harris 1957), the Bromide Formation, Simpson Group, middle Ordovician. Highway

99 locality, Sec. 11, T1S, R3E, Arbuckle Mountains, Oklahoma, U.S.A.: approximately, lat. 34°35'N, long.

96°41'W.

Museum of Comparative Zoology (MCZ), Harvard University, no. MCZ 4530a (holotype, car.: PI. 20, 42, figs.

1-3, PI. 20, 44, figs. 2, 4); from type horizon and locality. Natural History Museum, London, no. OS 13571 (car.:

PI. 20, 44, figs. 1, 3, 5); from Mountain Lake Member, 38 m below top of Bromide Formation at type locality.

Markedly postplete Kayina with pronounced posterodorsal node on left valve; when fully developed over-

reaching dorsal margin. No marginal structures. Valve surfaces gently convex, smooth.

The relationship of N American and Baltic species assigned to Kayina needs to be verified. K. hybosa differs

considerably from the Baltic K. subampleta Schallreuter, 1971 by being distinctly more postplete; by having the

dorsal node situated more posteriorly and more distally from the hinge; and by being wider posteriorly. The

coeval (Simpson Group) K? porosa Harris (1957, op cit.)

is poorly known; the holotype (MCZ 4531) is a crushed

carapace with very distorted valve overlap. It appears to

differ from K. hybosa in being punctate and having a

less distinct posterodorsal node.

Only in the Bromide Formation, Oklahoma.

M. Williams gratefully acknowledges support from

N. E.R.C., the Alexander von Humboldt Foundation,

Bonn and the Universite Claude Bernard, Lyon. Dr J.M.

Berdan is thanked for the loan of specimens.

Text-fig. 1 . Carapace of K. hybosa in thin section (specimen FSL 575052; collections at Lyon). A, line of section. /

B, thin section, indicating morphology relative to a theoretical ellipse enclosing the carapace; carapace is markedly C LV Xvy/ rv

narrower ventrally. C, ventral overlap structures.

Explanation of Plate 20, 44

Figs. 1, 3, 5, car. (OS 13571, 1.00 mm long): fig. 1, LV dors, obi.; fig. 3, close-up of ventral surface; fig. 5, post, detail showing dorsal

node. Figs. 2, 4, car. (holotype, MCZ 4530a, 1.08 mm long): fig. 2, dors.; fig. 4, RV ext. lat. obi.

Scale A (150/rm; x63), fig. 1; scale B (150/rm; x57), figs. 2, 4; scale C (150/rm; x76), fig. 3; scale D (100 gm; x93), fig. 5.

Stereo-Atlas of Ostracod Shells 20, 44

Kayina hybosa (4 of 4)

Stereo-Atlas of Ostracod Shells 20, 42

Kayina hybosa (2 of 4)

Stereo-Atlas of Ostracod Shells 20 (11) 45-48 (1993) Punctoschmidtella pauciperforata (1 of 4)

595.336 (113.312) (766 : 162.097.34): 551.351 + 552.54

ON PUNCTOSCHMIDTELLA PAUCIPERLORATA (HARRIS)

by Mark Williams & Jean Vannier

(University of Leicester, England & Universite Claude Bernard, Lyon, France)

Punctoschmidtella pauciperforata (Harris, 1957)

71931 Aparchites perforata n. sp., R.W. Harris, Bull. Okla. geol. Surv., 55, 87, pi. 5, figs. 4a, b.

1957 Paraschmidtella pauciperforata n. sp., R.W. Harris, Bull. Okla. geol. Surv., 75, 173, pi. 4, figs. 15a, b, 16.

1957 Paraschmidtella perforata (Harris); R.W. Harris, Bull. Okla. geol. Surv., 75, 174, pi. 4, figs. 17a, b.

1988 Punctoschmidtella perforata (Harris); J.M. Berdan, Mem. Bur. Mines Mineral Resourc., New Mex., 44, 287.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Museum of Comparative Zoology, Harvard University, U.S.A., no. MCZ 4550; a carapace.

From Decker’s Bed 59 (see Harris 1957), the Oil Creek Formation, Simpson Group, middle Ordovician, West

Spring Creek locality, Secs. 7 and 8, T2S, R1W, Arbuckle Mountains, Oklahoma, U.S.A.; approximately, lat.

34°30'N, long. 97°20'W.

Museum of Comparative Zoology (MCZ), Harvard University, U.S.A. nos. MCZ 4550 (car. holotype: PI. 20,

46, figs. 1, 2, 5; PI. 20, 48, figs. 2, 3) and MCZ 4552 (juv. car.: PI. 20, 48, fig. 1). Universite Claude Bernard

(FSL), Lyon, no. FSL 575051 (RV; PI. 20, 46, figs. 3, 4). MCZ 4550 (holotype) is from the type horizon and

locality. MCZ 4552 is from the type locality, Decker’s Bed 50, Oil Creek Formation. FSL 575051 is from the Oil

Creek Formation, South Interstate 35 locality (see Fay, R.O. et al., Paleont. Contr. Univ. Kans., Monograph,

1, Section 3, 1982), approximately 150 m below the top of the Formation at this locality.

Punctoschmidtella with external smooth adductor muscle spot anteriorly delimited by a shallow sulcal

depression. Inner umbonal surface of the smaller right valve with a low dorsocentral node.

Explanation of Plate 20, 46

Figs. 1, 2, 5, car. (holotype, MCZ 4550, 1.05 mm long): fig. 1, ext. It. lat.; fig. 2, ext. It. lat. obi.; fig. 5, dors. obi. Figs. 3, 4, RV (FSL

575051, 1.17 mm long): fig. 3, int. detail of contact groove; fig. 4, int. detail of hinge.

Scale A (150 //m; x63), figs. 1, 2, 5; scale B (200 /mi; x53), figs. 3, 4.

Stereo-Atlas of Ostracod Shells 20, 47

Punctoschmidtella pauciperforata (3 of 4)

Remarks:

Distribution:

Acknowledgements:

Like many schmidtellids P. pauciperforata has thick valves. The relationship of P. pauciperforata to other species

of Punctoschmidtella is indicated in Text-fig. 1. Features constant in definitive species of Punctoschmidtella

include right over left valve overlap and the presence of a contact groove in the right valve. No adventral

structures are present and sulcation is absent or only weakly developed. The dorsum is invariably umbonate and

the hinge of the left valve has a groove (see also Berdan 1988).

Berdan (1988) assigned Paraschmidtella perforata (Harris, 1931) to her genus Punctoschmidtella, presumably

on the basis of published figures of this species in Harris 1957 (pi. 4, figs. 17a, b = MCZ 4552; herein PI. 20, 48,

fig. 1). The holotype of Aparchites perforata Harris 1931 (pi. 5, figs. 4a, b = MVZ 7446) was figured as being

punctate but is in fact a severely abraded specimen which cannot be clearly identified and is herein considered

a nomen dubium. Although a juvenile, Harris’ 1957 (MCZ 4522) figured specimen referred to P. perforata is

clearly conspecific with Paraschmidtella pauciperforata Harris, 1957 (holotype MCZ 4550; see PI. 20, 46, figs.

1, 2, 5, PI. 20, 48, figs. 2, 3).

Only in the Oil Creek Formation,

Oklahoma.

M. Williams gratefully acknowledges

support from N.E.R.C., the Alex-

ander von Humboldt Foundation,

Bonn and the Universite Claude

Bernard, Lyon. Dr J.M. Berdan is

thanked for the loan of type

specimens.

Text-fig. 1. Carapace features of species

of Punctoschmidtella.

• relative to P pauapedorata

only known from a aingie valve

Explanation of Plate 20, 48

Fig. 1, juv. car., ext. It. lat. (MCZ 4552, 0.72 mm long). Figs. 2, 3, car. (holotype, MCZ 4550, 1.05 mm long): fig. 2, ext. rt. lat.; fig. 3,

ext. rt. lat. obi. Scale A (100 /an; x88), fig. 1; scale B (150 /mi; x63), figs. 2, 3.

Stereo-Atlas of Ostracod Shells 20, 46

Punctoschmidtella pauciperforata (2 of 4)

Stereo-Atlas of Ostracod Shells 20, 48

Punctoschmidtella pauciperforata (4 of 4)

Stereo-Atlas of Ostracod Shells 20 (12) 49-54 (1993) Wenlockiella phillipsiana (1 of 6)

595.337.23 (113.331) (420 : 162.003.52): 551.351 + 552.52

ON WENLOCKIELLA PHILLIPSIANA (JONES & HOLL)

by Robert F. Lundin & Lee E. Petersen

(Arizona State University, Tempe & Anadarko Petroleum Corporation, Houston, U.S.A.)

Genus WENLOCKIELLA gen. nov.

Type-species: Bairdia phillipsiana Jones & Holl, 1869

Derivation of name: For the type Wenlock Series of the Silurian, in which the genus is abundant.

Diagnosis: Smooth Thlipsuracea with subreniform to subtriangular outline in lateral view and subtriangular to ovate transverse

outline. Typically with strong dorsal L/R overreach. Ventral commissure straight in the anterior/posterior direction.

Contact groove in left valve poorly developed or absent; where present, best developed along anteroventral and

posteroventral contact margin. Hinge distinctly inclined to longitudinal axis of carapace.

Remarks: The type-species and similar related species are homeomorphs with the Middle Devonian genus Bairdiocypris Kegel,

1932. The discovery of a calcified inner lamella in Bairdiocypris (Adamczak, F. in: J.W. Neale (Ed.), Taxonomy,

Morphology and Ecology of Recent Ostracoda, Oliver & Boyd, Edinburgh, 93, 1969) and Adamczak’s subsequent

(Senckenberg. leth., 57, 265, 1976) analysis of several species of the same genus demonstrates that the Silurian and

probably the early Devonian forms referred to herein are not members of Bairdiocypris. Wenlockiella differs from

Bairdiocypris in lacking a calcified inner lamella, in having a unipartite hinge and in lacking a bow-shaped projection

(ventriculus). In addition, in most species of Wenlockiella the hinge is distinctly inclined to the longitudinal axis of the

carapace whereas in Bairdiocypris the hinge is typically parallel or subparallel to the longitudinal axis of the carapace.

Wenlockiella is distinguished from Silenis Neckaja, 1958 by details of hingement and by valve relationships along the

dorsum. It is distinguished from Octonaria Jones, 1887, to which it is ancestral (Petersen, L.E. & Lundin, R.F., J.

micropa/aeontol., 6(1), 77, 1987), by lack of shell sculpture.

We place the following species in Wenlockiella: Bairdia phillipsiana Jones & Holl, 1869; Macrocypris crassula Jones,

1887; Bythocypris phaseolus Jones, 1887; and Bythocypris gotlandica Jones, 1889. The following species are placed in

Explanation of Plate 20, 50

Figs. 1-4, car. (ASU X-148, 1429 pm long): fig. 1, ext. post.; fig. 2, ext. dors.; fig. 3, ext. vent.; fig. 4, ext. rt. lat. Figs. 5, 6, car.

(holotype, BMNH I 2066, 1250/rm long): fig. 5, ext. rt. lat.; fig. 6, ext. It. lat.

Scale A (200 pm; x36), figs. 1-4; scale B (200 pm: x40), figs. 5, 6.

Stereo-Atlas of Ostracod Shells 20, 51 Wenlockiella phillipsiana (3 of 6)

Wenlockiella provisionally: Bythocypris transversa Roth, 1929; Bairdiocypris ? sp. A of Lundin, 1965; Bairdiocyprisl

sp. B of Lundin, 1965; and Kuresaaria blackstonensis Berdan & Copeland, 1973.

The calcified inner lamellae reported by Lundin (Bull. Okla. geol. Surv., 108, 58, 1965) for Bairdiocyprisl spp. A and

B need to be verified by thin sections. It is possible that these species possess only thickenings of the shell along the

anteroventral and posteroventral parts of the contact margin. These species and Bythocypris transversa Roth, 1929 have

no bow-shaped projection and probably belong to Wenlockiella, but additional study is required to confirm this. We

conclude from illustrations (Berdan & Copeland (Prof. Pap. U.S. geol. Surv., 825, 35, pi. 12, figs. 13-25, 1973) that

the holotype of K. blackstonensis and all of the paratypes illustrated except the one in figure 22 ( = a Kuresaaria), and

possibly the one in figure 21, probably belong to Wenlockiella.

Wenlockiella is known from the Silurian of Great Britain, Gotland, the eastern Baltic area and Podolia. Species provi-

sionally included in the genus are from the upper Silurian and lower Devonian of the North American Midcontinent (Okla-

homa and western Tennessee) and the lower Devonian of northwestern North America (Alaska and the Yukon

Territories).

Wenlockiella phillipsiana (Jones & Holl, 1869)

Bairdia phillipsiana sp. nov. T.R. Jones & H.B. Holl, Ann. Mag. nat. Hist., (4), 3, 213, pi. 14, figs. 7a-c.

Bythocypris phillipsiana var. typica (Jones & Holl); T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 187, pi. 5, figs. 4a-c.

Bythocypris phillipsiana var. major Cones & Holl); T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 187, pi. 5, figs. 3a, b.

Bythocypris phillipsiana (Jones & Holl); E.O. Ulrich & R.S. Bassler, Maryland Geol. Surv., Silurian, 320, fig. 25:2.

Bythocypris phillipsiana (Jones & Holl); E.O. Ulrich & R.S. Bassler, Maryland Geol. Surv., Silurian, 702, pi. 63, fig. 9.

Bythocypris phillipsiana (Jones & Holl); M.J. Copeland, Palaeontology, 3, 101, pi. 23, figs. 21, 22 [$/c], 19, 20.

“Bythocypris” phillipsiana (Jones & Holl); D.M. Hoskins, Bull. Pa topogr. geol. Surv., G36, 97, pi. 7, figs. 19-21 .

Bairdiocypris phillipsiana (Jones & Holl); V.S. Krandijevsky, Fauna ostrakod silurijskich vidkladiv Podillia, Akad. Nauk Ukr. SSR (in Ukranian), 1 14,

pi. 11, figs. 1-4.

Bairdiocypris phillipsianus (Jones & Holl); A.F. Abushik, Ostracoda from Silurian-Lower Devonian key sections of Podolia, in: A.F. Abushik,

E.A. Gusseva& l.E. Zanina (Eds.), Palaeozoic ostracodes from key sections in the European part of the USSR, Moscow Akad. Nauka (in Russian), 1 17,

pi. 42, figs. 3-6.

Bairdiocypris phillipsianus (Jones & Holl); A. A. Pranskevichius, Trans. Lithuanian Scientific-Research Geol. Surv. Inst, (in Russian), 15, 138, pi. 28,

figs. 1 , 2.

“Bairdiocypris” phillipsiana (Jones & Holl); R.F. Lundin, L.E. Petersen & D.J. Siveter, J. micropalaeontol. , 9(2) (for 1990), pi. 2, fig. 13.

Explanation of Plate 20, 52

Figs. 1-3, car. (ASU X-201, 1429 pm long): fig. 1, ext. post.; fig. 2, ext. rt. lat.; fig. 3, ext. It. lat. Figs. 4, 5, car. (ASU X-202, 1128/im

long): fig. 4, ext. vent.; fig. 5, ext. rt. lat.

Scale A (200 pm\ x36), figs. 1-3; scale B (200 gm; x44), figs. 4, 5.

1869

1887

non 1887

1923

non 1923

non 1960

non 1961

1963

1971

1972

1991

Wenlockiella phillipsiana (2 of 6)

Wenlockiella phillipsiana (4 of 6)

Stereo-Atlas of Ostracod Shells 20, 50

Stereo-Atlas of Ostracod Shells 20, 52

Stereo-Atlas of Ostracod Shells 20, 53

Wenlockiella phillipsiana (5 of 6)

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

Distribution:

A cknowledgements:

Natural History Museum, London [BMNH] slide no. I 2066; one carapace that agrees well with the original type figures.

“Croft’s Quarry,” 0.5 km W of Malvern, Hereford & Worcester, England; approximately Nat. Grid. Ref. SO 757464,

lat. 52°08'N, long. 2°18'W. Much Wenlock Limestone Formation, Wenlock Series, Silurian.

Department of Geology, Arizona State University (ASU), nos. X-148 (car.: PI. 20, 50, figs. 1-4), X-201 (car.: PL 20,

52, figs. 1-3), X-202 (car.: PI. 20, 52, figs. 4, 5), X-203 (car.: Text-fig. la) and X-204 (car.: Text-fig. lb). Natural

History Museum, London [BMNH], no. I 2066 (holotype, car.: PI. 20, 50, figs. 5, 6).

ASU X-148 is from the Farley Member of the Coalbrookdale Formation below the east end of Benthall Edge,

Shropshire; lat. 52°37'N, long. 2°29'W. ASU X-201. X-202, X-203, and X-204 are from the Much Wenlock Limestone

Formation at Lincoln Hill near Ironbridge, Shropshire; lat. 52°38'N, long. 2°29'W. All specimens are from the

Homerian, Wenlock Series, Silurian.

Large Wenlockiella with subtriangular lateral and transverse outlines. Dorsum strongly arched and L/R overreach along

hinge line strong. Perimarginal ridge present along anteroventral margin of right valve of many specimens. Adductor

muscle field marked on interior of valves by circular depression posterior to which is a low limen-like ridge.

The species described here is most similar to W. gotlandica (Jones, 1889), from which it can be distingushed by its larger

size, its more angular lateral and transverse outlines and by the distinctly greater L/R overreach along the hinge line. All

of the approximately 1000 British specimens studied are carapaces. Accordingly, the contact margin features and interior

features are known only from thin sections (Text-figs, la, b) and by inference from the interior morphology of the close

relative, W. gotlandica. Maximum width is more posterior on some specimens of the British species than on others

(compare PI. 20, 50, figs. 2, 3 with PI. 20, 52, fig. 4). We have no evidence that this character is dimorphic nor do we

have any evidence from numerous single valves of W. gotlandica that the limen-like ridge is a dimorphic character.

Variation in size and length/height ratio of W. phillipsiana is indicated in Test-fig. lc.

Jones’ (1887, op. cit.) report of a variety of the species, Bythocypris phillipsiana var. major , needs to be checked. We

conclude from the original illustrations of this form that it does not belong to this species and probably does not belong

to Wenlockiella.

Known from late Llandovery to Ludlow strata of Britain and has been reported from rocks of the same age in the eastern

Baltic area and Podolia.

We gratefully acknowledge support from NATO (Grant No. 870445) and the National Science Foundation (Grant No.

EAR-8200816).

Stereo-Atlas of Ostracod Shells 20, 54 Wenlockiella phillipsiana (6 of 6)

Text-fig. 1, Outline drawings from photographs of transverse (fig. la, ASU X-203, posterior view, x63, sample MS 531) and

longitudinal (fig. lb, ASU X-204, ventral view, x37, sample MS 533) thin sections and a scatter diagram (fig. lc) for 48 carapaces

(sample MS 514) from the Farley Member of the Coalbrookdale Formation at Tickwood (cf. locality 44 of Lundin, Petersen &

Si veter, 1991, op. cit.).

Stereo-Atlas of Ostracod Shells 20 (13) 55-58 (1993) Parulrichia diversa (1 of 4)

595.33.12 (113.331) (420 : 162.003.52): 551.351 + 552.52

ON PARULRICHIA DIVERSA (JONES & HOLL)

by David J. Siveter & Robert F. Lundin

(University of Leicester, England & Arizona State University, Tempe, U.S.A.)

Genus PARULRICHIA Schmidt, 1941

Type-species (by original designation): Primitia diversa Jones & Holl, 1986

Diagnosis: Quadrilobate Drepanellacea with weak anterior lobe (LI), node-like L2 and L3, one or both of which may be ventrally

confluent with LI. L4 a posteroventral node to dorsoventrally oriented lobe. SI and S3 weak. S2 oval to arcuate around

dorsal, posterior and rarely ventral side of L2.

Remarks: Scott (1961 , op. cit.) placed Parulrichia in the Richinidae, an assignment based on the erroneous premise that P. diversa is

bilobate; a new familial designation must await study of related forms. Of the taxa which Schmidt ( 1 941 , op cit. ) placed in

Parulrichia we retain only the type-species — the only quadrilobate form — within the genus. The only congeneric species

known to us is P. bispinosa Lundin & Siveter ( Stereo-Atlas Ostracod Shells, 20, 59, 1993), from the Ludlow Series of

Tennessee. Parulrichia inarguta, Parulrichia minuta and Parulrichia ? tabulata (all Neckaja 1966) and Parulrichia

minima Sarv, 1956 belong outside Parulrichia. The species from the Devonian of the U.S.A. which Lundin and Petersen

& Lundin referred to Parulrichia (Bull. Okla. geol. Surv., 116, 1968 and 145, 1992 respectively) belong to a new genus.

Parulrichia diversa (Jones & Holl, 1886)

1886 Primitia cornuta sp. nov., T.R. Jones & H.B. Holl, Arm. Mag. nat. Hist., 17, 411, pi. 14, figs. 12a, 12b, 13.

1886 Primitia diversa sp. nov., T.R. Jones & H.B. Holl, Ann. Mag. nat. Hist., 17, 412, pi. 14, figs. lOa-c.

1941 Parulrichia diversa (Jones & Holl); E.A. Schmidt, Abh. senckenb. naturforsch. Ges., 454, 66.

1961 Primitia diversa Jones & Holl; H.W. Scott, in R.C. Moore & C.W. Pitrat (Eds.), Treatise on Invertebrate Paleontology, Kansas Univ. Press, Q132, figs. 4a-c.

1978 Parulrichia diversa (Jones & Holl); D.J. Siveter, in R. Bate & E. Robinson (Eds.), Geol. J. (Special Issue), 8, 72, pi. 2, figs. 9, 10, tab. 3.

1981 Parulrichia diversa (Jones & Holl); R.J. Aldridge, K.J. Doming & D.J. Siveter, in Neale, J.W. & Brasier, M.D. (Eds.), Microfossils from Recent and Fossil Shelf Seas,

Ellis Horwood, Chichester, 21, pi. 2.1, fig. 16.

1985 Parulrichia diversa (Jones & Holl); J.E. Mabillard & R.J. Aldridge, Palaeontology, 28, 98, text-figs. 8, 9c, 10.

1988 Parulrichia diversa (Jones & Holl); D.J. Siveter, British Micropalaeontological Society Field Guide, 2, 31, text-fig. 7, pi. 2, fig. 19.

Explanation of Plate 20, 56

Figs. 1-3, RV (OS 14162, 1130/rm long): fig. 1, ext. lat.; fig. 2, vent.; fig. 3, post. Figs. 4, 5, juv. RV (OS 14163, 770 pm long): fig.

4, ext. lat.; fig. 5, vent. Figs. 6-8, RV (OS 14166, 1050 pm long): figs. 6, ext. lat.; fig. 7, vent.; fig. 8, post.

Scale A (200 //m; x46), figs. 1-3, 6-8; scale B (200 pm; x51), figs. 4, 5.

Stereo-Atlas of Ostracod Shells 20, 57

Parulrichia diversa (3 of 4)

Neotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

Distribution:

A cknowledgements:

Of the syntypes of P. diversa (Vine Collection; Natural History Museum, London) the referred specimens XXXVII and

LXIV (Jones & Holl 1886, 412) are extant (= I 1944 [4 valves] and IN 52449 [6 valves] respectively). The lectotype

(Schmidt 1941; = Jones & Holl 1886, pi. 14, fig. 10a, a left valve; Vine coll. XXXVI3) is lost; the labelled slide contains

a broken right valve. Of the other originals of P. diversa ( = Vine Coll. XXXVIi and 2) that of pi. 14, fig. 10b is reported

to be IN 52433 (N.M.H. catalogue) but is absent from the collections and that of pi. 14, fig. 10c is also not present or

even catalogued. We here designate the largest and only complete valve from slide 1 1944 (from the type bed no. 37) as

neotype of P. diversa. Based on its lectotype (here designated; Jones & Holl 1886, pi. 14, figs. 12a, 12b = carapace

I 1942, Vine coll. XXXV) Primitia cornuta is synonymous.

Vine’s “bed” no. 37 = Buildwas Fm., Silurian, upstream from bridge over River Severn at Buildwas, Shropshire,

England (Vine, G.R., Proc. Yorks, geol. polytech. Soc., 9, 233, 1887); lat. 52.39N, 2.33W.

Natural History Museum, London [BMNH], 1 1942 (lectotype of Primitia cornuta, carapace: PI. 20, 58, fig. 4), I 1944

pars (neotype LV: PI. 20, 58, fig. 5), OS 14162 (RV: PI. 20, 56, figs. 1-3), OS 14163 Guv. RV: PI. 20, 56, figs. 4, 5),

OS 14164 (LV: PI. 20, 58, figs. 1-3), OS 14165 (RV.: PI. 20, 58, figs. 6, 7) and OS 14166 (RV: PI. 20, 56, figs. 6-8).

All from Buildwas Fm., Sheinwoodian Stage, Wenlock Ss., Shropshire; lat. 52.36N, long. 2.3W. 1 1942 from near type

locality (“bed” no. 40; see Vine 1887). OS 14162-5 from Harley Brook, near Domas; OS 14162, OS 14163 from loc. 36

of Siveter 1980 (Palaeontogr. Soc. [Monogr.], 1), OS 14164, OS 14165 from c.50 m S of loc. 36 (Nat. Grid Ref. c.SJ 5922

0030). OS 14166 from Buildwas (loc. 35 of Siveter 1980).

Weakly punctate Palulrichia\ posteroventral node-like, to dorsoventrally ridge-like, L4.

L4 shows much intraspecific variation; typically a posteroventral node (PI. 20, 56, figs. 1-5), in all specimens of some

samples it is a well developed dorsoventrally aligned lobe (PI. 20, 58, figs. 1-3, 6, 7). Many single samples contain both

morphotypes; in still other samples one morphotype or the other occurs with an intermediate form having a dorso-

ventrally elongate, but not fully developed,. L4 (PI. 20, 56, figs. 6, 7). P. bispinosa differs in having a posterodorsal

spine, in lacking puncta and in details of lobation and sulcation.

Low to middle part of Sheinwoodian Stage, Wenlock Ss., Buildwas Fm., C. centrifugus & C. murchisoni zones,

Shropshire; localities given above, plus Siveter 1980 loc. 37, “Domas” and “Buildwas” of Lundin et al. (J.

micropalaeontol., 9, 178, 1991) and “Leasows” of Mabillard & Aldridge (1985). Also in basal part of Brinkmarsh Fm.

(basal Wenlock), Tortworth inlier, Avon; and Barr Limestone Mbr. (M. riccartonensis Zone), Coalbrookdale Fm.,

Walsall, W Midlands ( = Siveter 1980, Iocs 12 & 25 respectively).

We thank NATO and Arizona State University for support.

Explanation of Plate 20, 58

Figs. 1-3, LV (OS 14164, 1125/rm long): fig. 1, post.; fig. 2, ext. lat.; fig. 3, vent. Fig. 4, carapace. It. lat. (I 1942, 1240 pm long). Fig.

4, RV ext. lat. (neotype, I 1944 pars', 970 pm long). Figs. 6, 7, RV (OS 14165; 1080 //m long): fig. 6, ext. lat.; fig. 7, vent.

Scale A (200 ^m; x46), figs. 1-3, 6, 7; scale B (200 gm; x42), fig. 4; scale C (200 /rm; x49), fig. 5.

Stereo-Atlas of Ostracod Shells 20, 56

Stereo-Atlas of Ostracod Shells 20, 58

Parulrichia diversa (2 of 4)

Parulrichia diversa (4 of 4)

Stereo-Atlas of Ostracod Shells 20 (14) 59-62 (1993) Parulrichia bispinosa (1 of 4)

595.336 (113.33) (768 : 162.089.35): 551.351 + 552.54

ON PARULRICHIA BISPINOSA LUNDIN & SIVETER sp. nov.

by Robert F. Lundin & David J. Siveter

(Arizona State University, Tempe, U.S.A. & University of Leicester, England)

Holotype:

Type locality:

Derivation of name:

Diagnosis:

Figured specimens:

Remarks:

Parulrichia bispinosa sp. nov.

Department of Geology, Arizona State University (ASU), no. ASU X-104; RV.

[Paratypes: Arizona State University nos. ASU X-101-X-103].

Section P5, glade SE of Decaturville, Perryville Quadrangle, Tennessee, U.S.A. ; lat.

35° 30'49.5"N, long. 88°3'24"W. Holotype from sample P5-9, 15.1m above the base of the

Brownsport Fm, late Ludlow Series, Silurian.

Latin bi, two, and spina thorn, referring to the posterodorsal thorn-like spines.

Parulrichia with node-like L2, L3, and L4. Large posterodorsal spine on each valve above poorly

developed S3. The posteroventral node is spine-like on some specimens. S2 pit-like to slightly

arcuate around posterodorsal side of L2. L2 and L3 ventrally confluent with LI.

Department of Geology, Arizona State University (ASU), nos X-104 (holotype, RV; PI. 20, 60,

figs. 1, 2), X-103 (paratype, LV: PI. 20, 60, figs. 3-5), X-101 (paratype, RV: PI. 20, 62, figs. 1, 2),

X-102 (paratype, juv. LV: PI. 20, 62, figs. 3-5). All specimens from same sample as holotype.

Parulrichia bispinosa is distinguished from the type-species, P. diversa (Jones & Holl, 1886) (see

Siveter & Lundin, Stereo-Atlas Ostracod Shells , 20, 55, 1993), in having a posterodorsal spine on

each valve. In other respects, the two species are quite similar. We consider that P. bispinosa was

Explanation of Plate 20, 60

Figs. 1, 2, RV (holotype, ASU X-104, 695 pm long): fig. 1, ext. lat.; fig. 2, ext. vent. Figs. 3-5, LV (ASU X-103, 658pm long);

fig. 3, ext. dors.; fig. 4, int. lat.; fig. 5. ext. lat. All measurements exclusive of spines and nodes.

Scale A (200 pm; x91), fig. 1; scale B (200 ^m; x88), figs. 2, 3; scale C (200 pm\ x90), figs. 4, 5.

Stereo-Atlas of Ostracod Shells 20, 61 Parulrichia bispinosa (3 of 4)

derived from P. diversa or a close descendant of it. Lundin (Bull. Okla. geol. Surv., 116, 45, 1968)

has discussed the relationships of Parulrichia to a complex of early Devonian species which must

have been derived from P. bispinosa or a close descendant. The occurrence of Parulrichia in

western Tennessee strengthens the link between British and North American midcontinent ostracod

faunas of Silurian age (see also Xystista auricularis and X. graffhami; Stereo-Atlas Ostracod

Shells, 12, 77-80 & 12, 81-84, 1985; by Siveter and Lundin & Siveter respectively). This link,

though generally considered weak, is significant to Silurian ostracod biogeography.

Distribution: Known from four samples from two localities in the Brownsport Fm, Ludlow Series, Silurian, of

Perryville and Olive Hill quadrangles, western Tennessee. The samples range from 12.3 m to

15.1 m above the base of the Brownsport Formation.

Acknowledgements: The authors acknowledge support from NATO for their collaborative research programme. RFL

also acknowledges support of the College of Liberal Arts and Sciences, Arizona State University.

0.4

i0-3

Text-fig. 1. Size dispersion diagram of twenty-two right and left valves of P. bispinosa from sample P5-9, Brownsport Formation,

western Tennessee (see Type locality). Open circle (top right) represents the holotype.

Explanation of Plate 20, 62

Figs. 1, 2, RV (ASU X-101, 611 pm long): fig. 1, ext. lat.; fig. 2, ext. post. Figs. 3-5, juv. LV (ASU X-102, 469 pm long): fig. 3, ext.

dors.; fig. 4, ext. vent.; fig. 5, ext. lat. All measurements exclusive of spines and nodes.

Scale A (200 pm\ x89), fig. 1; scale B (200 pm\ x90), fig. 2; scale C (200 pm\ xl28), figs. 3-5.

_l i Ll

0.5 0.6 0.7

Length, mm

Stereo-Atlas of Ostracod Shells 20, 62

Stereo-Atlas of Ostracod Shells 20, 60

Parulrichia bispinosa (2 of 4)

Parulrichia bispinosa (4 of 4)

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Stereo-Atlas of Ostracod Shells: Vol. 20, Part 1

CONTENTS

On Cytheromorpha diamphidia Maybury sp. nov.; by C.A. Maybury.

On Semicytherura paraclausi Maybury sp. nov.; by C.A. Maybury.

On Kiltsiella rosensteinae (Sarv); by D.J. Siveter & L.I. Sarv.

On Sulcella huecoensis Dewey & Kohn sp. nov.; by C.P. Dewey & P. Kohn.

On Nipponocy there colalongoae (Ciampo); by V. Drapala & M.A. Ayress.

On Nipponocythere cuneata Ayress & Correge sp. nov.; by M.A. Ayress & T.

Correge.

On Kuiperiana dryppa (Whatley & Coles); by M.A. Ayress & V. Drapala.

On Aboilia blessi Becker & Adamczak gen. et sp. nov.; by G. Becker & F.F.

Adamczak.

On Baltonotella elegans (Harris); by M. Williams & J. Vannier.

On Kayina hybosa Harris; by M. Williams & J. Vannier.

On Punctoschmidtella pauciperforata (Harris); by M. Williams & J. Vannier.

On Wenlockiella phillipsiana (Jones & Holl); by R.F. Lundin & L.E. Petersen.

On Parulrichia diversa (Jones & Holl); by D.J. Siveter & R.F. Lundin.

On Parulrichia bispinosa Lundin & Siveter sp. nov.; by R.F. Lundin & D.J. Siveter.

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