A Stereo-Atlas of Ostracod Shells

edited by I. Boomer, D. J. Horne, A. R. Lord and D. J. Siveter

Volume 23, Parts 1 and 2; 1996

Published under the aegis of the British Micropalaeontological Society, London

ISSN 0952-7451

Editors

Dr Ian Boomer, School of Environmental Sciences, University of East Anglia, Norwich NR4 7TJ. Tel: +1603

593123; Fax: +1603 507719; Email: i.boomer@uea.ac.uk.

Dr David J. Horne, School of Earth Sciences, University of Greenwich, Chatham Maritime, Kent ME4 4AW. Tel:

+ 181 331 9841; Fax: + 181 331 9805; Email: d.j.horne@greenwich.ac.uk.

Professor Alan R. Lord, Department of Geological Sciences, University College London, Gower Street, London

WC1E 6BT. Tel: + 171 380 7131; Fax: + 171 388 7614; Email: dean.maps@ucl.ac.uk.

Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH. Tel: + 116 523925; Fax: + 116

523918; Email: djs@leicester.ac.uk.

Editorial Board

Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, F-33321 Begles, France.

Dr M.A. Ayress, Department of Geology, The Australian National University, G.P.O. Box 4, Canberra, ACT

2601, Australia.

Dr R.E.L. Schallreuter, Geologisches-Palaontologisches Institut, Universitat Hamburg, Bundesstrasse 55, D-20146

Hamburg, Germany.

Professor N. Ikeya, Institute of Geosciences, Shizuoka University, Shizuoka 422, Japan.

Subscriptions

Subscriptions should be sent to Prof. Alan Lord at the above address. North American subscribers may, if they

wish, send their subscriptions to Dr Mary McGann, M.S. 915, USGS, 345 Middlefield Road, Menlo Park,

California, 94025 USA.

Officers of the British Micropalaeontological Society

Chairman: Prof. R.J. Aldridge, Department of Geology, University of Leicester, Leicester, LEI 7RH.

Secretary: Mrs S.L. Matthews, c/o Department of Geological Sciences, University College London, Gower Street,

London WC1E 6BT.

Treasurer: Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GG.

Membership Treasurer: Dr L.T. Gallagher, Network Stratigraphic Consulting Ltd., Unit 57, The Enterprise

Centre, Cranborne Road, Potters Bar, Hertfordshire EN6 3DQ.

Editor, Journal of Micropalaeontology : Professor J.W. Murray, Department of Geology, Southampton

Oceanography Centre, Empress Dock, Southampton S014 3ZH.

Editor, Newsletter of Micropalaeontology: Dr A.J. Powell, Millenia Ltd., Unit 3, Weyside Park, Newman Lane,

Alton, Hampshire GU34 2PJ.

Calcareous Nannofossil Group: Chairman - Dr J.R. Young; Secretary - Mr. M. Hampton.

Conodont Group: Chairman - Dr I.J. Sansom; Secretary - Dr C.G. Miller.

Foraminifera Group: Chairman - Dr M.A. Kaminski; Secretary - Mr M.D. Bidgood.

Ostracod Group: - Chairman - Dr D.J. Horne; Secretary - Dr M. Williams.

Palynology Group: Chairman - Dr D.W. Jolley; Secretary - Mr D. McLean.

Instructions to Authors

Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. All

contributions submitted for possible publication in A Stereo-Atlas of Ostracod Shells are peer-reviewed by an

appropriate international specialist. “Instructions to Authors” and plate blanks for mounting photographs may be

obtained from any Editor. Manuscripts should be submitted together with a copy of the text on disk (MS WORD,

or ASCII), to Dr Ian Boomer.

The front cover shows two specimens of Cytheropteron bronwynae Joy & Clark, 1977 from a Recent sample on

the Morris Jesup Rise, Arctic Ocean (lat. 85° 19.4'N, long. 14° O'W). Upper specimen, RV of a male carapace,

dorsal view, BMNH no. 1995.1281), lower specimen, RV, external lateral view, BMNH no. 1995.1288). This

species was illustrated in Stereo-Atlas of Ostracod Shells, 22, 41-44.

A Stereo-Atlas of Ostracod Shells

edited by I. Boomer, D. J. Horne, A. R. Lord, D. J. Siveter

Volume 23, 1996

Published under the aegis of the British Micropalaeontological Society, London

Stereo-Atlas of Ostracod Shells 23, ii

Contents

1 On Swainocythere miniscula (Ruggieri); by C.P. Dickson. 1

2 On Pellucistoma punctata Ayress sp. nov.; by M. Ayress. 5

3 On Hemicytherura fulva McKenzie, Reyment & Reyment; by K.G. McKenzie,

R.A. Reyment & E.R. Reyment. 9

4 On Eucytherura loenensis sp. nov.; by I. Boomer. 13

5 On Scepticocythereis sanctivincentis sp. nov.; by S. Majoran. 17

6 On Schizocythere inexpecta sp. nov.; by S. Majoran. 21

7 On Echinocythereis leckwycki sp. nov.; by K. Wouters. 25

8 On Orionina caboverdensis sp. nov.; by K. Wouters. 29

9 On Darwinula incae (Delachaux); by G. Rossetti, K. Martens & P. Mourguiart. 35

10 On Wenlockiella phaseola (Jones); by L.E. Petersen & R.F. Lundin. 41

11 On Cytherellina elegans (Jones); by L.E. Petersen & R.F. Lundin. 45

12 On Cytherellina ruperti sp. nov.; by L.E. Petersen & R.F. Lundin. 49

13 On Ogmoconcha contractula Triebel; by I. Boomer & T. Jellinek. 53

14 On Eucypris virens (Jurine); by R. Smith & K. Martens. 61

15 On Baltonotella kuckersiana (Bonnema); by R.E.L. Schallreuter. 69

16 On Karinutatia ren Schallreuter; by R.E.L. Schallreuter. 73

17 On Soanella ova/is (Ivanova); by R.E.L. Schallreuter. 77

18 On Valentella costata (Ivanova); by R.E.L. Schallreuter. 81

19 On Trapezilites minimus (Kummerow); by I.C.U. Hinz-Schallreuter. 85

20 On Falites fala Muller; by I.C.U. Hinz-Schallreuter. 89

21 On Cytheropteron kempfi nom. nov.; by I. Boomer. 95

22 Index for volume 23 (1996). 96

Stereo-Atlas of Ostracod Shells 23 (1) 1-4 (1996)

595.337.14 (119.4) (261.27 : 162.006.54) : 551.351

Swainocythere miniscula (1 of 4)

ON SWAINOCYTHERE MINISCULA RUGGIERI

by Carol P. Dickson

(School of Geography, Kingston University, Surrey)

Swainocythere miniscula Ruggieri, 1976

1975 Cytheropteron? minisculum sp. nov. G. Ruggieri, A. Unti, M. Unti, and M.A. Moroni, Soc. Geol. Ital., Boll., 94: 1654.

1981 Swainocythere chejudoensis sp. nov. K. Ishizaki. Tohoku Univ., Sci. Rep., 2nd ser. (Geol.), 51 (1-2): 37-65.

Type Specimens:

Type locality:

Figured specimens:

Believed to be deposited in the personal collection of Ruggieri (Palermo, Italy); figured holotype no.

O.C.R. SI. 2602a female carapace:

Figured paratype no O.C.R. SI. 2602b male carapace; unfigured paratype no. O.C.R. SI. 2602c.

Lower Pleistocene glauconitic clayey sand with Pecten shells at the base of calcarenites in a cave opening

at Piano Messina, between Campobello and Mazara, SSW of kilometre post 63 of S.S. 115, westernmost

Sicily, Italy (latitude 37° 32' N and longitude 12° 40' E).

Deposited in the collections of the British Geological Survey, Keyworth, England. No. BGS 89/15-1.1-1

(9 LV: PI. 23, 2, fig. 1) from 1.05-1.15 m down core; no. BGS 89/15-1.1-2 (9 RV: PI. 23, 2, figs. 2, 4-5)

from 1.05-1.15 m down core; no. BGS 89/15-1.1-3 (o* LV: PI. 23, 2, fig. 3) from 1.05-1. 15m down core;

no. BGS 89/15-1.1-4 (9 PI. 23, 4, figs. 1, 3-4) from 1.05-1.15 m down core; no. BGS 89/15-2.85-5 (9 RV:

PI. 23, 4, fig. 2) from 2. 8-2.9 m down core; no. BGS 89/15-5.85-6 (A-l RV: PL 23, 4, fig. 5) from

5. 8-5. 9 m down core. All specimens from the Holocene sediments of BH 89/15 (latitude 54° 02.208' N,

longitude 5° 20.645' W, from the northern Irish Sea, present day water depth 92 m).

Explanation of Plate 23, 2

Fig. 1, 9 LV, ext. lat. BGS 89/15-1.1-1, 241 pm long); figs. 2, 4-5, 9 RV, int. lat. BGS 89/15-1.1-2, 249 pm long; fig. 2, anterior hinge;

fig. 4, hinge; fig. 5, posterior hinge; fig. 3, o’ LV, ext. lat. BGS 89/15-1.1-3, 238 pm long).

Scale A (100 pm; x200), figs. 1, 3; scale B (20 pm; x600), figs. 2, 5; scale C (100 pm\ x300), fig. 4.

Stereo-Atlas of Ostracod Shells 23, 3

Swainocythere miniscula (3 of 4)

Diagnosis:

Remarks:

Distribution:

A ckno wledgement:

Carapace small; sub-trapezoid in outline; dorsal and ventral margins sinuous, anterior margin rounded

and posterior margin tapering to a short caudal process. Greatest height anterior of mid-length. Marginal

ridge runs sub-parallel to the margins and provides a distinct shoulder. Small punctae cover the surface

and extend in rows sub-parallel to the margins, the rows of puncta lie between gentle ridges. Hinge modi-

fied antimerodont, the right valve terminal teeth are fairly large, the median element is smooth centrally

but becomes loculate posteriorly and only very weakly so anteriorly, the left valve has complementary

elements.

Ruggieri et al. (op. cit., 1975) tentatively assigned this species to the genus Cytheropteron Sars, 1866.

Ishizaki (1981 Tohoko Univ. Sci. Rep., 2nd ser. (Geol.), 51 (1-2), 37-65) studying ostracods from the

East China Sea, established the genus Swainocythere with the type species S. chejudoensis Ishizaki, 1981.

Cytheropteron and Swainocythere are similar in the character of the hinge structure and in the arrangement

of the radial pore canals although the two genera can be distinguished in general appearance,

Swainocythere being more elongate and lacking an alar process (Ishizaki, 1981). The species S. chejudoensis

Ishizaki, 1981 may be conspecific with Cytheropteron? minisculum Ruggieri, 1976. The East China Sea

specimens differ slightly from the type specimens of S. minisculum and those figured herein, in that the

surface ornament is more strongly ridged and the hinge structure is less modified, as in the right valve the

anterior and posterior teeth are smaller and the median element is more strongly loculate throughout.

Other species of this genus include Swainocythere nanseni (Joy and Clark, 1981) (Correge et al.,

Stereo-Atlas Ostracod Shells, 19, 107-110, 1992) and two other undescribed species from the Arctic and

South Pacific waters; which underlines the circum-polar nature of this genus. The difference between this

species and S. nanseni (Joy and Clark, 1981) are described in Correge et al. (1992, op. cit.).

Lower Pleistocene of Italy (Ruggieri, 1975); Holocene to Recent of the Celtic and Irish Seas (herein) at

depths of greater than 80 m.

I would like to thank Robin Whatley for reviewing the manuscript.

Explanation of Plate 23, 4

Fig. 1, 3-4, 9 LV, int. lat. (BGS 89/15-1.1-4, 250 long); fig 1, posterior hinge; fig 3, hinge; fig. 4, anterior hinge; fig. 2, 9 RV, ext.

lat. (BGS 89/15-2.85-5, 223 pm long); fig. 5, A-l RV, ext. lat. (BGS 89/15-5.85.6, 219 pm long).

Scale A (20 pm\ x600), figs. 1, 4; scale B (100 pm; x200), fig. 2, 3; scale C (100 ^m; x230), fig. 5.

Swainocythere miniscula (4 of 4)

Stereo-Atlas of Ostracod Shells 23, 2

Swainocythere miniscula (2 of 4)

Stereo-Atlas of Ostracod Shells 23, 4

Stereo-Atlas of Ostracod Shells 23 (2) 5-8 (1996)

595.337.14 (119.4) (265.7 : 163.159.59) : 551 .352

Pellucistoma punctata (1 of 4)

ON PELLUCISTOMA PUNCTATA AYRESS sp. nov.

by Michael A. Ayress

(Department of Geology, Australian National University, Canberra, Australia)

Pellucistoma punctata sp. nov.

Ho lo type:

Type locality:

Derivation of name:

Figured specimens:

Diagnosis:

National Museum of Victoria, Melbourne, Australia, no. P197956, Male, LV.

Tasman Sea, Institute of Oceanographic Sciences, Sydney core 1/86 6GC3, 30-31 cm, West Lord

Howe Rise, water depth 1540 m. Latitude 32° 58.8' S, longitude 159° 59.9' E. Holocene

foraminiferal ooze.

Referring to the punctate ornament of this species.

National Museum of Victoria, Melbourne, Australia, nos. P197956 (holotype, LV-: PL 23, 2

figs. 1, 4; PI. 23, 4, fig. 1), P197957 (paratype, RV: PI. 23, 2, figs. 2, 5; PI. 23, 4, fig. 2), P197958

(incomplete paratype, RV: PI. 23, 2, fig. 3). The holotype and paratype (P197957 are from the type

locality and horizon. The paratype (P197958) is from the Lord Howe Rise, Tasman Sea (water

depth 1340 m) Late Pleistocene sample at 97.5 cm in core Sonne 36-61.

A species of Pellucistoma with a finely punctate carapace, subrectangular in lateral view. Hinge in

the left valve supported by an antislip tooth at both ends. Sexual dimorphism not apparent in

available material.

Explanation of Plate 23, 6

Figs. 1, 4, LV (holotype, P197956, 420 pm long): fig. 1, ext. lat.; fig. 4, ant. anti-slip tooth. Figs. 2, 5, RV (P197957, 420 pm long):

fig. 2, ext. dors.; fig. 5, ext. lat. Fig. 3, RV, subcentral muscle scars, (P197958, 300 pm incomplete length). Scale A (100 //m; xl45),

figs. 1, 2, 5; Scale B (50^m, x420), fig. 3; Scale C (20 pm, x800), fig. 4.

Stereo- Atlas of Ostracod Shells 23, 7

Pellucistoma punctata (3 of 4)

Remarks:

Distribution:

A cknowledgemen t:

In almost all aspects this species is identical to Pellucistoma coombsi Ayress, 1990 {New Zealand

Nat. Sci. 17, 68) known from the Late Eocene to Recent of New Zealand, and has probably

descended from that species. Pellucistoma punctata differs from P. coombsi most notably in its

punctate surface, the surface of the latter species being smooth. Carapace ornament appears to be

a rare feature of the genus hitherto recorded, to the authors knowledge, only in the Tertiary of the

Caribbean, e.g. Pellucistoma? spurium Bold (Bold, Bull. Am. Pal., 94 (329), 66, pi. 12, figs 9-12,

1988). The discovery of P. punctata is the first record of the genus from bathyal depths. Compari-

sons between the central American Pellucistoma and the Indo-Pacific Javanella suggest that the

two genera are synonymous separated only on geographical grounds (Howe and McKenzie, 1989

(N. Terr. Mus. Arts & Sci., monograph 3, 50p); Ayress, 1990 op. cit.).

Late Quaternary to Recent of the Tasman and Coral Seas: sample at 30 cm (Holocene) and 125 cm

(Late Pleistocene) in Ocean Sciences Institute, University of Sydney core 1/86 6GC3 (water depth

1540 m); sample at 97.5 cm (Late Pleistocene) in core Sonne 36-61 (water depth 1340 m); James

Cook University surface sediment grab sample 590/9 (water depth 1242 m); and sample at 35 cm

(Holocene) in core V24-160 (water depth 1007 m).

The staff of the Electron Microscope Unit (ANU) are thanked for their assistance.

Explanation of Plate 23, 8

Fig. 1, LV, int. lat. (P197956, 420 pm long). Fig. 2, RV, int. lat. (P197957, 420 pm long).

Scale A (100 pm; x225), figs. 1, 2.

Pellucistoma punctata (2 of 4)

Stereo-Atlas of Ostracod Shells 23, 6

Stereo-Allas of Ostracod Shells 23 (3) 9-12 (1996)

595.337.14 (118.14) (94 : 163.142.39) : 551.35)

Hemicytherura fulva (1 of 4)

ON HEMICYTHERURA FULVA McKENZIE, REYMENT & REYMENT

by Ken G. McKenzie1, Richard A. Reyment2 and Eva R. Reyment2

(Geology School, University of Melbourne, Victoria, Australia,

2 Institute of Earth Sciences, University of Uppsala, Uppsala, Sweden)

Hemicytherura fulva McKenzie, Reyment and Reyment, 1993

1993 Hemicytherura fulva K.G. McKenzie, , R.A. Reyment and E.R. Reyment, Revta Esp. Paleontol., 8, 97, PI. 4, figs. 18-20.

Holotype: Palaeontological Museum, Institute of Earth Sciences, University of Uppsala, Sweden, PMAus441,

male carapace, SEM stub Vic-2 (Kl).

Type Locality : Browns Creek, Victoria, southeastern Australia, long. 142.15° E, lat. 38.18° S. Late Eocene.

Figured specimens: Palaeontological Museum, Institute of Earth Sciences, University of Uppsala. PMAus441 (holotype,

male car.: PI. 23, 10, fig. 1), PMAus441-l (female car.: PI. 23, 10, fig. 2), PMAus441-2 (car.: pi.

23, 10, fig. 3), PMAus443 (paratype, female? LV: PI. 23, 12, fig. 1), PMAus442 (paratype, female

car.: PI. 23, 12, fig. 2), PMAus441-3 (car.: PI. 23, 12, fig. 3). All specimens from the type locality,

Eocene, Victoria.

Explanation of Plate 23, 10

Fig. 1. Male car., It. lat. (holotype, PMAus441, 338/rni long). Fig. 2. Female car., rt. Lat. oblique (PMAus441-l, 361 pm long).

Fig. 3. Adult car., vent. (PMAus441-2, 350/zm long).

(Scale = 50 pm; xl80).

Stereo-Atlas of Ostracod Shells 23, 1 1

Hemicytherura fulva (3 of 4)

Diagnosis:

Remarks:

Distribution:

Small, oval in lateral view, subcaudate; ornament consists of a longitudinal median rib and a marginal

ridge that rims each valve almost continuously; diagonal ribs, often bearing finer reticulation, cross

the lateral surfaces. Polymorphic, one morph having lace-like costulations (“laced”), the other with

greatly subdued ornament (“effaced”), the latter being the most frequently recorded morph is

defined in the main diagnosis. Dorsum convex; anteroventral margin subacuminate, bearing a few

marginal denticles; venter almost straight (and partly overlapped by the ventromarginal ridge in

lateral view); posterior cauda form a depressed platform behind the posteromarginal ridge. Greatest

height medial; subhastate in dorsal view, broadest posteromedially. Muscle scars and hinge typical

of the genus. Sexual dimorphism weakly developed with females slightly larger than males.

The ornament of this species is unlike that of any previously reported Australian Hemicytherura.

Polymorphism in ostracods may involve variations in size, shape, ornament, or a combination of all

three. It is most commonly due to ecophenotypy, but genuine cases of evolutionary polymorphism

also occur, polymorphism om H. fulva is well developed and has recently been studied by geometric

morphometries (Reyment, R.A., Revta esp. Paleontol. 8, 125-131, 1993). The “laced” morph

encompasses adults that retain, in part, a larval aspect.

At Browns Creek, H. fulva ranges from 7 m below the contact of the Browns Creek Clay with the

Johanna River Greensand Member, all Eocene.

Castle Cove and Browns Creek, Victoria, southeastern Australia; Middle? to Late Eocene.

Explanation of Plate 23, 12

Fig. 1. Female? LV. int. detail of hinge and muscle scars (paratype, PMAAus443, 356^m long). Fig. 2. Female car., dors, (paratype,

PMAus442, 361 pm long). Fig. 3. Male car., It. Lat. (“laced” morph), (PMAus441-3, 334 gm long).

(Scale = 50//m; xl80).

Hemicytherura fulva (2 of 4)

Hemicytherura fulva (4 of 4)

Stereo-Atlas of Ostracod Shells 23, 10

Stereo-Atlas of Ostracod Shells 23, 12

Stereo-Atlas of Ostracod Shells 23 (4) 13-16 (1996)

595.337.14 (118.15) (265 : 162.180.18) : 551.352 + 552.54

Eucytherura loenensis (1 of 4)

ON EUCYTHERURA LOENENSIS BOOMER sp. nov.

by Ian Boomer

(School of Environmental Sciences, University of East Anglia, Norwich, England, U.K.)

Eucytherura loenensis sp. nov.

Holotype:

Type locality:

Derivation of name:

Figured specimens:

The Natural History Museum, London [BMNH] no. OS 14896; adult RV. [Paratype: no. OS

14897].

Lo En Guyot, Central Pacific Ocean, Ocean Drilling Program, Leg 144, Site 872C, (10° 5.8' N,

162° 51.9' W) Core 16, core-catcher (0-8 cm); Upper Oligocene.

With reference to the type locality Lo En Guyot, Central Pacifc Ocean.

The Natural History Museum, London [BMNH] nos. OS 148896 (holotype, LV: PI. 23, 14, figs.

1, 2, 4, PI. 23, 16, figs. 1, 2, 5), OS 14897 (paratype, RV: PI. 23, 14, fig. 3, PI. 23, 16, figs. 3, 4).).

Both holotype and paratype from type level and locality.

Explanation of Plate 23, 14

Fig. 1, 2, 4, RV (holotype, OS 14896, 290 /um long): fig. 1 ext. lat.; fig. 2 dors.; fig. 4 int. lat. Fig. 3, RV, dors, (paratype. OS 14897,

290 //m long).

Scale A (100 /rm; x225), fig. 1; scale B (100 /um; x205), figs. 2, 3; scale C (100 gm; x235), fig. 4.

Stereo-Atlas of Ostracod Shells 23, 15 Eucytherura loenensis (3 of 4)

Diagnosis: A small, reticulate species of Eucytherura with distinct anterodorsal, posterodorsal and ventral

flanges. The carapace is subquadrate in lateral view, dorsal, and ventral margins converge slightly

posteriorly. Anterior margin broadly rounded, posterior margin subtriangular. Greatest height at

anterior cardinal angle, greatest width just above midheight. The anterodorsal region bears two

short subparallel flanges with a third present posterodorsally. The lateral and dorsal reticulae are

regular and subrectangular, the posteroventral reticulae are raised to form a box-like structure.

Ventral margin bears microreticulation with fine longitudinal ribs. The species is apparently blind.

The inner lamella is broad, no vestibule observed, the inner margin of the posterior lamella is

notable as it bears a forward projecting flexure (PI. 23, 14, fig. 4; PI. 23, 16, figs. 4, 5). Hinge

antimerodont with coarse crenulae on median element (PI. 23, 16, fig. 2). Muscle scars not

observed.

Remarks: The species bears similarities to a number of taxa described from the Upper Cretaceous of Western

Australia ( E . antipodum, Neale, J.W. Spec. Pap. Palaeont. 16, 1-82, 1975), Cainozoic of the

North Atlantic ( E . pseudoantipodum Coles. G. and Whatley, R., Revta esp. Micropaleont., 21,

81-124, 1989) and the Cainozoic of the SW Pacific and Indian Ocean ( E . aff. antipodum, E.

pseudoantipodum, Ayress, M.A. et al., Rec. Aust. Mus., 47, 203-223, 1995). The present species

differs from the aforementioned taxa in possessing square, open reticulae, a marked posterodorsal

crescentic rib and in the forward flexure of the posterior lamella.

Distribution: Known only from the type locality.

Explanation of Plate 23, 16

Figs 1, 2, 5, LV (holotype, OS 14896, 290 /rm long): fig. 1, vent.; fig. 2, int. hinge detail; fig. 5, int. detail of posterior lamella.

Figs. 3, 4, RV (paratype, OS 14897, 290 um long): fig. 3, ext. lat.; fig. 4, int. lat.

Scale A (100 gm; x200), figs 1, 3, 4; scale B (50 /mi; x370), fig. 2; scale C (10 /um; X625), fig. 5.

Stereo-Atlas of Ostracod Shells 23, 14

Eucytherura loenensis (2 of 4)

Stereo-Atlas of Ostracod Shells 23 (5) 17-20 (1996)

595.337.14 (118.14) (94 : 163.138.36) : 551.35

Scepticocythereis sanctivincentis (1 of 4)

ON SCEPTICOCYTHEREIS SANCTIVINCENTIS MAJORAN sp. nov.

by Stefan Majoran

(Department of Marine Geology, Goteborg University, Sweden)

Scepticocythereis sanctivincentis sp. nov.

1993 “ Cythereis ” sp., K.G. McKenzie, R.A. Reyment and E.R. Reyment, Revta esp. Paleont., 8, 106, pi. 6, fig. 9.

Ho l o type:

Type locality:

Derivation of name:

Figured specimens:

South Australian Museum, Adelaide, Australia no. SAM P35501; Female LV.

Type section of the Blanche Point Formation, near Willunga, South Australia (lat. 35° 15' S, long.

138° 24' E). Late Eocene, Priabonian. Holotype collected 1.3 m above the base of the Gull Rock

Member (dated by planktonic foraminifera as PI 6, see McGowran et al., 1992 in: D.R. Prothero

and W.A. Berggren (Eds.), Eocene-Oligocene Climatic and Biotic Evolution , Princeton University

Press, 178-201).

After the provenance of the holotype in the St. Vincent Basin, South Australia.

South Australian Museum, Adelaide, Australia, nos SAM P35501 (holotype, female LV: PI. 23,

18, fig. 1), SAM P35502 (female car.: PI. 23, 18, fig. 2), SAM P35503 (male RV: PI. 23, 18,

fig. 3), SAM P35504 (male RV: PI. 23, 20, fig. 1), SAM P35505 (female car.: PI. 23, 20, fig. 2),

SAM P35506 (male LV: PI. 23, 20, fig. 3), All specimens from the Gull Rock Member of the

Blanche Point Formation.

Explanation of Plate 23, 18

Fig. 1, Female LV, ext. lat. (holotype, SAM P35501, 1000 pm long). Fig. 2, Female car., ext. dors. (SAM P35502, 990 pm long).

Fig. 3, Male RV, ext. lat. (SAM P35503, 1150/im long).

Scale A (200 /mi; x70), figs. 1-3.

Stereo-Atlas of Ostracod Shells 23, 19 Scepticocythereis sanctivincentis (3 of 4)

Diagnosis: A species of Scepticocythereis with a conspicuously inflated anterior margin being particularly

prominent along its ventral part. Carapace subrectangular in lateral view, subhastate in dorsal

view. Inequivalved as the left valve overreaches the right antero- and posterodorsally. Entire

surface reticulate and characterised by small inwards facing mural spines extending into the fossae.

Fossae more elongate in the marginal regions compared to the central region. Reticulate surface

disrupted by a spiny posteroventral projection. Ventral section of anterior and posterior margins

bears stout spines. Eye-tubercle large. Sexual dimorphism prominent. Right hinge with a crenulate

anterior tooth, faintly crenulate posteromedian furrow and a bilobate posterior tooth; left hinge

complementary. Muscle-scars with a V-shaped frontal scare and four adductors in a subvertical

series.

Remarks: The external and internal morphology of the new species is very similar to the Upper Cretaceous

type-species S. ornata (see Bate, R.H., Spec. Pap. Palaeont, 10, 67-70, pi. 26, figs. 1-8, pi. 27,

figs. 11-12, text-figs. 37A-F, 1972; and Neale, J.W. Spec. Pap. Palaeont, 16, 61-62, pi. 2, fig. 10,

pi. 21, fig. 4, text-figs. 12d, f, h., 1975) from Western Australia. This only concerns the shape of

the frontal muscle-scar. It is clearly V-shaped in the present material but was described as “oval or

slightly crescentic” (see Bate, 1972, op. cit .; and Neale., 1975, op. cit.). Majoran ( GFF 117, 80

appendix, 1995; Revta esp. Paleont., 11, 33-34, appendices, 1996) provisionally named this species

Cletocythereis sp.

Distribution: Presently known from the Tortachilla Limestone and the Tuketja, Gull Rock and Perkana

Members of the Blanche Point Formation, South Australia (Late Eocene, Priabonian, Zones

P15-P16, see McGowran et al., 1992 in: D.R. Protero and W.A. Berggren (Eds.), op. cit.). Also,

from the Browns Creek Clays at Browns Creek, Victoria (Late Eocene, see McKenzie et al., 1993,

op. cit.).

Explanation of Plate 23, 20

Fig. 1, Male RV, int. lat. (SAM P35504, 1080 pm long). Fig. 2, Female car., ext. vent. (SAM P35505, 970 pm long). Fig. 3, Male LV,

int. lat. (SAM P35506, 1060 pm long).

Scale A (200 pm: x70), figs, 1-3.

Stereo-Atlas of Ostracod Shells 23, 18

Scepticocythereis sanctivincentis (2 of 4)

Scepticocythereis sanctivincentis (4 of 4)

Stereo-Atlas of Ostracod Shells 23, 20

Stereo-Atlas of Ostracod Shells 23 (6) 21-24 (1996)

595.337.14 (118.15) (94 : 163.144.39) : 551.35

Schizocythere inexpecta (1 of 4)

ON SCHIZOCYTHERE INEXPECTA McKENZIE, REYMENT & REYMENT

by Stefan Majoran

(Department of Marine Geology, Goteborg University, Sweden)

Schizocythere inexpecta McKenzie, Reyment and Reyment, 1991

1991 Schizocythere inexpecta sp. nov., K.G. McKenzie, R.A. Reyment and E.R. Reyment, Revta. esp. Paleont., 6, 149, pi. 3, fig. 13.

Holotype: Institute of Earth Sciences, Palaeontology, Uppsala Univeristy, Uppsala, Sweden, no. PAM. Au. 234;

Male LV.

Type locality: Bells Headland, Victoria, Australia; approx, lat. 38° 24' S, long. 144° 6' E. Dated as Lower

Oligocene (Janjukian) by McKenzie et al. (op. cit.).

Figured specimens: Department of Marine Geology, Goteborg University, Sweden, nos. DMGUG.Au.130 (male LV:

PI. 23, 22, fig. 1), DMGUG.Au.131 (female LV: PI. 23, 22, fig. 2). DMGUG.Au.132 (male RV: PI.

23, 22, fig. 3), DMGUG.Au.133 (male RV: PI. 23, 24, fig. 1), DMGUG.Au.134 (female car.: PI. 23,

24, fig. 2), DMGUG.Au.135 (female LV: PI. 23, 24, Fig. 3). All specimens from the Ruwarung

Member of the Port Willunga Formation, Lower Oligocene.

Explanation of Plate 23, 22

Fig. 1, Male LV, ext. lat. (DMGUG.Au.130, 540 pm long). Fig. 2, Female LV, ext. lat. (DMGUG.Au.131, 520 pm long). Fig. 3, Male

RV, ext. lat. (DMGUG. Au.132, 560 pm long).

Scale A (100 pm; xl20), figs. 1-3.

Stereo-Atlas of Ostracod Shells 23, 23 Schizocythere inexpecta (3 of 4)

Diagnosis: Carapace subrectangular in lateral view, subovate in dorsal view with subacuminate posterior

margin. Caudal process slightly above mid-height. Ornament evenly reticulate with strongly

developed muri and rather deep rounded fossae which are largest in the ventral region. Dorsal ridge

almost completely suppressed, ventral ridge more distinctly developed but relatively thin. Sexual

dimorphism prominent. The presumed males are somewhat larger in size than the females displaying

a more distinct posterodorsal disruption of the ornament and a more distinct projection of the

posterior cardinal angle in the left valve. Two posteroventral projections of the surface ornament are

discernible in both valves, the ventral one being more strongly developed in male right valves.

Internal features as for genus.

Remarks: This species is common in the Port Willunga Formation, South Australia, and was identified after

examining the types of Schizocythere inexpecta McKenzie, Reyment and Reyment, which was the

first record of Schizocythere in Australia. McKenzie et al. (op. cit.) only illustrated an external view

of the holotype. The following observations are added to the type description. The more numerous

representatives of the Port Willunga Formation exhibit sexual dimorphism, referred to above. The

posteroventral projections differentiate this species from other members of the genus. Although

McKenzie et. al. (op. cit.) believed the holotype to be female, in the present authors opinion the

specimen is male.

Distribution: Presently known from Bells Headland, Victoria (Janjukian, Lower Oligocene) and from the Aldinga

and Ruwarung Members of the Port Willunga Formation, South Australia (Zones P18-P21,

Rupelian, Lower Oligocene, McGowran et al., in: D.R. Prothero and W.A. Berggren (Eds.),

Eocene-Oligocene Climatic and Biotic Evolution, Princeton University Press, 178-201, 1992).

Explanation of Plate 23, 24

Fig. 1, Male RV, int. lat. (DMGUG. Au. 133, 560 //m long). Fig. 2, Female car., ext. dors. DMGUG. Au. 134, 500 /rm long). Fig. 3,

Female LV, int. lat. (DMGUG. Au. 135, 500 pm long).

Scale A (110/im; Xl20), figs. 1-3.

Stereo-Atlas of Ostracod Shells 23 (7) 25-28 (1996)

595.337.14 (118.21) (493 : 161 .004.51) : 551 .35

Echinocythereis leckwijcki (1 of 4)

ON ECHINOCYTHEREIS LECKWIJCKI WOUTERS sp. nov.

by Karel Wouters

(Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, Belgium)

Echinocythereis leckwijcki sp. nov.

1978 Cythere Wetherelli, Jones; G.S. Brady, Trans, zool. Soc. Lond., 10, 390, pi. 64, figs. 7a-7d ( non Jones, 1856).

1918 Cythereis Wetherellii Brady; W.N. Kuiper, Oligocane und Miocane Ostracoden aus den Niederlanden, 66-67, pi. 3, figs. 28a-28c (non Jones,

1856).

1981 Echinocythereis variolata (Egger, 1859) s.l . ; H. Uffenorde, Palaeontographica, A172, 156, pi. 2, figs. 13, 16 (non Egger, 1859).

Holotype:

Type locality:

Figured specimens:

Derivation of name:

Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, Belgium, no. TCTI 6265; female LV

[Paratypes nos. TCTI 6266-6270, 6291, 6292].

S.W. Antwerp, Belgium, near the entrance to the railroad tunnel under the River Scheldt (long.

4° 22' 35" E, lat. 51° 12' 14" N). Edegem Sand Member, Lower Miocene.

Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels (KBIN) nos. TCTI 6265 (holotype,

female LV: PI. 23, 26, fig. 1), TCTI 6266 (paratype, female RV: PI. 23, 26, fig. 2), TCTI 6267 (paratype,

male LV; PL 23, 26, fig. 3), TCTI 6268 (paratype, male RV: PI. 23, 26, fig. 4), TCTI 6269 (paratype,

female LV: PL 23, 28, fig. 1), TCTI 6270 (paratype, male RV: PL 23, 28, fig. 2), TCTI 6291(paratype,

female car.: PL 23, 28, fig. 3). All specimens from type locality and horizon.

In honour of the late Prof. W.P. Van Leckwijck (1902-1975).

Explanation of Plate 23, 26

Fig. 1, female LV, ext. lat. (holotype, TCTI 6265, 1042 pm long). Fig. 2, female RV, ext. lat. (paratype, TCTI 6266, 1090 pm long).

Fig. 3, male LV, ext. lat. (paratype, TCTI 6267, 1060 gm long). Fig. 4, male RV, ext. lat. (paratype, TCTI 6268, 1087 pm long).

Scale A (250 /^m; x56) figs. 1-54.

Stereo-Atlas of Ostracod Shells 23, 27 Echinocythereis leckwijcki (3 of 4)

Diagnosis: A species of Echinocythereis with inflated valves bearing faint polygonal reticulation; muri bearing very

short, blunt spines; anterolateral region unornamented but bearing a weak submarginal rib.

Remarks: Echinocythereis leckwijcki sp. nov. resembles E. subcornuta (Lienenklaus, 1900). The latter species,

however, is distinguished by a ventro-lateral ridge consisting of small denticles and a stout postero-lateral

spine. Furthermore, the weakly developed submarginal ridge is set with small spines. E. variolata (Egger,

1858) differs from the new species in being more oblong in lateral view, and in having only weakly

inflated valves in dorsal view. The muri of the reticulum are markedly thicker. Although there is an

overall resemblance. E. variolata and E. leckwijcki sp. nov. ( = Cythere wetherelli Jones sensu Brady,

non Jones) are not synonymous as supposed by Witt (1967, Geol. Bavar., 57: 41-42). Eagar (1965, Rev.

Micropaleont., 8: 20) assigned Cythere wetherelli to Cytheropteron ( Eocytheropteron ), followed by

Haskins (1970), Rev. Micropaleont., 13, 18, pi. 1, figs. 35-42). Species such as E. scabra (Muenster,

1830), E. scabrella (Lienenklaus, 1900), E. lima (Reuss, 1850) and E. reticulatissima Eagar, 1965, from

the Tertiary of Europe, are more strongly ornamented. E. semireticulata (Haskins, 1971) is similar to the

new species in shape, size and type of ornamentation but differs in that it possesses ventral and

ventrolateral longitudinal striae. E. jacksonensis (Howe and Chambers, 1935) (see Hazel, J.E., Mumma,

M.D. and Huff, W.J., Trans. Gulf Coast Ass. Geol. Soc., 30, pi. 1, fig. 10, 1980) from the Lower

Oligocene of Mississippi and Alabama, is similar to the present species but differs by the presence of rows

of spinules along the anteromarginal zone.

Distribution: Belgium: Lower and Middle Miocene (Edegem Sand Member at Antwerp, Lier, Terhagen, Wilrijk;

Antwerpen Sand Member at Borderhout and Zonderschot Sand Member at Heist-op-den-Berg). The

Netherlands: Middle Miocene (Liessel and Sevenum, Kuiper, 1918) and in the well at Haamstede (coll.

Noordermeer). Germany: Upper Oligocene to Middle Miocene (several localities, Uffenorde, 1981).

Explanation of Plate 23, 28

Fig. 1, female LV, int. lat. (paratype, TCTI 6269, 1062 pm long). Fig. 2, male RV, int. lat. (paratype TCTI 6270, 1067 jum long).

Fig. 3, female car., dors, (paratype, TCTI 6291, 1057/rm long).

Scale A (250//m; x56); figs. 1-3.

Stereo-Atlas of Ostracod Shells 23, 26

Echinocythereis leckwijcki (2 of 4)

Stereo-Atlas of Ostracod Shells 23, 28

Echinocythereis leckwijcki (4 of 4)

Stereo-Atlas of Ostracod Shells 23 (8) 29-34 (1996)

595.337.14 (119.9) (261.1 : 162.019.25) : 551.351 + 552.51

Orionina caboverdensis (1 of 6)

ON ORIONINA CABOVERDENSIS WOUTERS sp. nov.

by Karel Wouters

(Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, Belgium)

Orionina caboverdensis sp. nov.

1869 Cythere finmarchica (G.O. Sars); G.S. Brady, in De Folin and Perier, Fonds de la mer, p. 138 ( non Sars, 1866).

1892 Cythere finmarchica (G.O. Sars); G.S. Brady and A.M. Norman, Sc. Trans. Roy. Dublin Soc., 4, ser. 2, p. 163.

Holotype:

Type locality:

Derivation of name:

Figured specimens:

Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, (KBIN) no. OC 1779, dissected male. (Paratypes

nos. OC 1780-OC 1790; 7 dissected specimens, 75 adults and 15 juveniles preserved in alcohol.)

Cape Verde Islands (Atlantic Ocean), Sao Vicente, Baia das Gatas, intertidal, on sand between boulders (collected by

T. Backeljau, February 5th, 1996).

With reference to the type locality.

Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels, nos. OC 1779 (holotype, male LV: PI. 23, 30,

fig. 1, Text-figs, la, b, 2f), OC 1780 (paratype, female LV: PI. 23, 30, fig. 2; female RV: PI. 23, 30, fig. 3, Text-fig.

lc), OC 1784 (paratype, female RV: PI. 23, 32, fig. 1; female LV: PI. 23, 32, fig. 2), OC 1787 (paratype, female car.:

PI. 23, 32, fig. 3), OC1788 (paratype, male car.: PI. 23, 32, fig. 4), OC 1789 (paratype, female LV, Text-fig. la),

OC 1782 (paratype, male appendages, dissection: Text-figs. 2a,b,e), OC1783 (paratype, male appendages, dissection:

Text-fig. 2d), OC 1785 (paratype, female appendages, dissection: Text-figs. la,b,d, 2c), OC 1786 (paratype, female

appendages, dissection: Text-figs. le,f). All specimens from the type locality.

Explanation of Plate 23, 30

Fig. 1, o ■ LV, ext. lat. (holotype, OC 1779, 500 pm long). Fig. 2, 9 LV, ext. lat. (paratype, OC 1780, 510/rm long). Fig. 3, 9 RV, ext.

lat. (paratype, OC1780, 510/tm long).

Scale A (100 /tm; xllO), figs. 1, 2, 3.

Stereo-Atlas of Ostracod Shells 23, 31

Orionina caboverdensis (3 of 6)

Diagnosis:

Remarks:

Distribution:

Acknowledgement:

A species of Orionina with weak ornamentation. Of the three longitudinal ridges, so prominent in the type species, only

two are recognized in the present species. The posterior transverse rib is weakly developed, recticulation very faint with

approximately ten knob-like protuberences in the antero-ventral area. Two frontal muscle scars are recorded, the ventral

one sutured; sexual dimorphism pronounced.

Most species of the genus Orionina are characterized by a number of strong ribs (in some species up to seven) with a

well developed reticulation pattern. Bold (J. Paleontol., 2>1, 33-50, 1963), has noted that there is a wide range of

variation, and that in some species only three ribs or weak striations and pitting are present. Such is the case in Orionina

caboverdensis sp. nov., where the reticulation and the ridges are much reduced. Because of its weak ornamentation,

Orionina fragilis Bold, 1963 from the Upper Miocene of Trinidad, is similar to the present species, however, the latter

species differs, in possessing a more elongate lateral outline with less well developed longitudinal ridges and a more

prominent posterior ridge. Although it is generally accepted that Orionina species have three frontal scars, Orionina

caboverdensis sp. nov. has only two, the ventral frontal scar being sutured. In some specimens this suture is very

indistinct whereas in others it is clearly visible, and can give the impression that three scars are present.

The new species may be confused with Finmarchinella finmarchica (Sars, 1866) because of the strking external

similarity between the two. However, observations on the morphology of the inner lamella (with pillar structures) and

the nature of the marginal pore canals clearly identified the new species as belonging to the genus Orionina. Hazel ( Geol .

Surv. Prof. Pap., 564, p. 19, 1967) stressed that records of F. finmarchica in the Cape Verde Islands were inconsistent

with other occurrences of the genus and, thus considered them to be misidentified, a point of view which was followed

by Neale (Bull. Br. Mus. nat. Hist. Zool., 27: p. 85, 1974).

The new species is known only from the type locality, having been previously recorded from St. Vincent (Cape Verde

Islands) by Brady (1869), op. cit.) and Brady and Norman (1892, op. cit.) as Cythere finmarchica. At the type locatity

the species occurs together with living populations of Kotoracythere inconspicua (Brady, 1880) and Keija demissa

(Brady, 1868). This is the first record of the genus Orionina in the Eastern Atlantic. The new species may well be

endemic to the islands, since it was not found by Witte (Verb. Kon. Ned. Akad. Wet. Afd. Natuurk., 39: 1-84, 1993)

in his extennsive study on ostracods from Senegal and the Gambia (620 km east of the Cape Verde Islands).

I wish to thank my colleague Dr. Thierry Backeljau (Koninklijk Belgisch Instituut voor Natuurwetenschappen, Brussels)

who collected the material.

Explanation of Plate 23, 32

Fig. 1, 9 RV, int. lat. (paratype, OC 1784, 530 pm long). Fig. 2, 9 LV, int. lat. (paratype, OC 1784, 530/tm long). Fig. 3, 9 car. dors.

(paratype, OC 1787, 540 pm long). Fig. 4, crcar. dors, (paratype, OC 1788, 520 pm long).

Scale A (100 pm; xllO), figs. 1, 2, 3.

rfit^n

...i<

Orionina caboverdensis (2 of 6)

Stereo-Atlas of Ostracod Shells 23, 32

Orionina caboverdensis (4 of 6)

Stereo-Atlas of Ostracod Shells 23, 30

Stereo-Atlas of Ostracod Shells 23, 33

Orionina caboverdensis (5 of 6)

Text-fig. la. 9 LV int. lat. (paratype, OC 1789, 525 //m long); lb, cr antenna (holotype, OC 1779); lc, 9 antenna (paratype, OC 1780); Id, o' antennule

(holotype, OC 1779); le, 9 mandibule (paratype, OC 1786). If, 9 maxillule (paratype, OC1786). Scales, fig. la: 200 /rm, figs, lb-f: 50 /urn.

Stereo-Atlas of Ostracod Shells 23, 34 Orionina caboverdensis (6 of 6)

Text-fig. 2a. o' 2nd leg (paratype, OC 1782), 2b, cr 3rd leg (paratype, OC 1782); 2c, 9 3rd leg and abdominal extremity (paratype, OC 1785);

2d, a 1st leg (paratype, OC 1783); 2e, cr brush-like organ (paratype, OC 1782); 2f crcopulatory organ (holotype, OC1779). All scales 50 fxm.

Stereo-Atlas of Ostracod Shells 23 (9) 35-40 (1996)

595.337.13 (119.9) (84.164 : 069.17) : 551 .312

Darwinula incae (1 of 6)

ON DARWINULA INCAE DELACHAUX

by Giampaolo Rossetti, Koen Martens and Philippe Mourguiart

(Department of Environmental Sciences, University of Parma, 43100 Parma, Italy,

Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium and

ORSTOM, 213 rue La Fayette, 75480 Paris, France)

Darwinula incae Delachaux

1928 Darwinula incae sp. nov., T. Delachaux, Bull. Soc. neuch. Sc. nat., 1, 54-56, pi. 5, figs. 28-39.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Repository unknown. One adult decalcified female with the following measurements: L = 0.87 mm, H = 0.4 mm. See

orginal description by Delachaux ( op . cit.).

Lake Huaron (Region of Huancavelica, Department of Junin, Peru) (lat. 13° 23' S, long. 72° 15' W).

Royal Belgian Institute of Natural Sciences (Brussels, Belgium), Ostracod Collection, nos. OC 1791 (9 car.: PI. 23, 36.

fig. 1), OC 1792 (9 car.: PI. 23, 36, fig. 2), OC 1793 (9 car.: PI. 23, 36, fig. 3), OC 1794 ( 9 RV and LV + appendages:

PI. 23, 38, figs. 1-4; Text-figs. 1A, IB, ID, 2A, 2C), OC 1795 (9 appendages: Text-figs. 1C, 2D, 2E), OC 1796

(9 appendages: Text-figs. IE, 2B).

All specimens collected on July 7, 1995 from Laguna “Guaqui,” Bolivia (3810 m a.s.l., approx, lat. 16° 30' S, long.

68° 48' W). Shallow pool and canal in largely dry Laguna, turbid, many algae, c. 10 cm deep, c. 150 x 50 m large; water

temperature =15.2 °C, pH = 9.3, conductivity = 767 /rS/cm, dissolved oxygen concentration = 9.2 mg/1.

Large-sized Darwinulid. Valves unequal, left overlapping the right on all sides. Shell with smooth surface. Seen dorsally,

carapace ovoid in outline, posterior extremity broadly rounded, anterior rather convex. In lateral view, both ends

rounded, anterior narrower than posterior; ventral margin almost straight, dorsal broadly arched. Central muscle scars

consisting of 13-14 small spots arranged in a nearly circular rosette. Postero-ventral keel on right valve and internal teeth

in left valve absent.

Explanation of Plate 23, 36

Fig. 1, female car., dors. (OC 1791, 804 pm long); fig. 2, female car., lat. (OC 1792, 788 pm long); fig. 3, female car., vent. (OC 1793,

772 pm long).

Scale A (200 pm; xl20), figs. 1-3.

Stereo-Atlas of Ostracod Shells 23, 37 Darwinula incae (3 of 6)

Exopodite of antenna with bristles a and b (text-fig. 1A) of equal length, reaching the distal extremity of the third

article of the endopodite. Second segment of mandibular palp bearing in distal position two long setae (y and z) and two

shorter setae ( w and y), the latter not exceeding the next segment (text-fig. 2A); distal margin of third segment with 5

claws of different lengths, two external setae (a and b ) and one smaller internal setae c, the latter half as long as the former

two (text-fig. 2A). Second segment of the maxillar endopodite with only one setae a (text-fig. 2B). Furcae present as two

long setae. Male unknown.

Remarks: The original description of Darwinula incae was based on only one decalified specimen. This implies an unsatisfactory

representation of the valve shape; moreover, a complete description of the appendages is lacking. No further descriptions

of D. incae are available up to the present. Darwinula specimens from Laguna Guaqui are attributed to D. incae because

of the great similarity with the original description in several diagnostic details of the appendages, which are not shared

with any other living species of this genus. Our specimens are smaller than the holotype, but still larger than any other

Recent Darwinula species. There are 25 described Recent species in the genus Darwinula , which can roughly be divided

into two main groups: the D. stevensoni group, with RV overlapping LV on all sides, and the D. africana group, with

LV overlapping the RV. This subdivision is not absolute, as the position of at least D. serricaudata, with stevensoni-Mke

morphology, but with LV overlapping RV, remains at present ill understood. The second lineage can effectively be

divided again into several sublineages (see D. Danielopol, Bijdr. Dierk. 50(2), 243-291, 1980), but these groupings will

be reassessed elsewhere (Rossetti and Martens, in prep.). Darwinula incae clearly belongs in the second lineage, but due

to its exceptionally large size takes a rather isolated position within this group. Apart from size and number of muscle

scars, D. incae is in general easily distinguishable by its morphology from the other living representatives of Darwinula

recorded from central and South America. The ill-described D. managuensis Swain and Gilby, 1965 presents compressed

valves and a nearly ovoid left valve; D. africana brasiliensis Pinto and Kotzian, 1961 and D. pagliolii Pinto and Kotzian,

1961 are characterized by an internal postero-ventral tooth in the left valve and by a small keel at the postero-ventral

corner of the right valve, respectively; D. serricaudata espinosa Pinta and Kotzian, 1961 has an elongated shell in lateral

view; in D. auraucana Loftier, 1961 right valve overlaps the left; D. dicastrii Loffler, 1966 is quite compressed in lateral

view, with anterior and posterior margins slightly rounded. Not so clearly distinguishable from D. incae is D. setosa

Daday, 1902. Possibly, D. setosa will in time have to be considered a senior synonym of D. incae, but the inadequate

original description of the former and the fact that its type material consists of one crushed female only do not allow a

decision to date.

Explanation of Plate 23, 38

Fig. 1, female LV, int. lat. (OC 1794, 773 pm long); fig. 2, female RV, int. lat. (OC 1794, 741 pm long); fig. 3; female LV, int. lat.,

detail of adductor muse. sc. (OC 1794); fig. 4, female RV, int. lat., detail of adductor muse. sc. (OC 1794).

Scale A (200 pm\ xl20), figs. 1, 2; scale B (100 pm; x503), figs. 3, 4.

Stereo-Atlas of Ostracod Shells 23, 39 Darwinula incae (5 of 6)

Distribution: Recent, freshwater: Peruvian and Bolivian Altiplano.

Acknowledgements: We gratefully acknowledge support from the E.U. Human Capital and Mobility Program (contract ERBCHRXCT/

93/0253). J. Cillis and C. Behen (Brussels, Belgium) offered technical assistance with the SEM images and with the line

drawings respectively.

Text-fig. 1. Appendages: A, A1 (OC 1794); B, A2 (OC 1794); C, upper lip (OC 1795); D, Md-masticatory process (OC 1794); E, Mxl-palp (OC 1796).

Scale bar = 54 //m for 1A,B; 106 /urn for 1C; 43/rm for 1D,E.

Stereo-Atlas of Ostracod Shells 23, 40

Darwinula incae (6 of 6)

Text-fig. 2. Appendages: A, Md-palp, 2nd and 3rd segment (OC 1794); B, M x 2-endopodite (OC 1796); C, T1 (OC 1794); D, T2 (OC 1795); E, furcae

(OC 1795). Scale bar = 54 nm for 2A-E.

Stereo-Atlas of Ostracod Shells 23 (10) 41-44 (1996)

595.337.23 (113.331) (420: 162.003.52) : 551.351

Wenlockiella phaseola (1 of 4)

ON WENLOCKIELLA PHASEOLA (JONES)

by Lee E. Petersen and Robert F. Lundin

(Anadarko Petroleum Corporation, Houston, U.S.A. and

Arizona State University, Tempe, U.S.A.)

Wenlockiella phaseola (Jones, 1887)

1887 Bythocypris phaseolus sp. nov. T.R. Jones, Ann. mag. nat. Hist., (5), 19, 189, pi. 7, figs 11a, 12b.

non 1923 Bythocypris phaseolus Jones; E.O. Ulrich and R.S. Bassler, Maryland Geol. Surv., Silurian, 702, pi. 63, figs. 5, 6.

1934 Bythocypris phaseolus Jones; R.S. Bassler and B. Kellett, Geol. Soc. Am. Sp. Pap. 1, 230.

1991 “Bairdiocypris” phaseolus (Jones); R.F. Lundin, L.E. Petersen & D.J. Siveter, J. Micropalaeontol., 9 (part 2 for 1990), pi. 2, fig. 11.

Lectotype: We here designate as the lectotype the specimen on Natural History Museum (NHM), London,

England, slide no. IN 52405 (ex. 1 1919) which contains one carapace of an immature specimen that

agrees well with the specimen illustrated by Jones (1887, op. cit., figures 12a, 12b).

Type locality: Buildwas, Shropshire, England; Vine collection XII, bed no. 38, Buildwas Formation, Wenlock

Series, Silurian. G.R. Vine (1887, Proc. Yorks, geol. polytech. Soc., 9, 224-248) records bed no.

38 as from “above Buildwas Bridge’’.

Figured specimens: Department of Geology, Arizona State University (ASU), nos. X-147 (car.: PI. 23, 44, figs. 5, 6),

X-277 (car.: PI. 23, 42, figs. 1-4), X-278 (car.: PI. 23, 44, figs. 1-4) and X-279 (car.: PI. 23, 42,

fig. 7. Natural History Museum, London, no. IN 52405 (lectotype, car.: PI. 23, 42, figs. 5, 6).

Explanation of Plate 23, 42

Figs. 1-4, car. (ASU X-277, 987 /ym long, 508 pm high): fig. 1, ext. It. lat.; fig. 2, ext. rt. lat. ; fig. 3, ext. dors.; fig. 4, ext. post.

Figs. 5, 6, juv. car. (lectotype, NHM IN 52405, 725 /ym long): fig. 5, ext. rt. lat.; fig. 6, ext. It. lat. Fig. 7, juv. car. (ASU X-279, 677 pm

long): ext. rt. lat.

Scale A (200 /ym; x53), figs. 1-4; scale B (200 /ym; x58), figs. 5, 6; scale C (150/ym; x72), fig. 7.

Stereo-Atlas of Ostracod Shells 23, 43

Wenlockiella phaseola (3 of 4)

Diagnosis:

Remarks:

Distribution:

A cknowledgements:

ASU X-147 and X-279 are from the lower part of the Coalbrookdale Formation at Buildwas

Bridge (Nat. Grid Ref. SJ 6451 0445), Shropshire, England (locality 34 of Lundin et al., op. cit.,

1991). ASU X-277 and X-278 are from the Buildwas Formation at Buildwas Abbey (Nat. Grid Ref.

SJ 643 045), Shropshire, England (locality 37 of Lundin et al., op. cit., 1991). NHM IN 52405 is

from the Buildwas Formation at the type locality. All specimens are from the Sheinwoodian,

Wenlock Series, Silurian, at approximately lat. 52° 39' N, long. 2° 33' W.

Medium-sized Wenlockiella with elongate subreniform lateral outline and subellipsoidal longi-

tudinal and transverse outlines. Dorsum very gently arched and left/right overreach along hinge

line weak. Perimarginal ridge present along anteroventral margin of right valve of well-preserved

adult specimens, absent from juveniles.

The species described here is most similar to W. crassula (Jones, 1887), from which it can be

distinguished by its more elongate lateral outline, its less-arched dorsum and by its weaker

left/right overreach along the hinge structure. All of the approximately 80 specimens studied are

carapaces. The contact margin features and interior features, interpreted from a single longitudinal

thin section, are similar to those of the type-species of the genus (see Lundin and Petersen, Stereo-

Atlas Ostracod Shells, 20(12), 53, text-fig. lb, 1993).

W. phaseola is morphologically more similar to W. crassula than to Wenlockella phillipsiana

(Jones & Holl, 1869) suggesting that it is directly ancestral to W. crassula. However, the possibility

that W. phillipsiana is the direct ancestor of both W. phaseola and W. crassula cannot be ruled out.

W. phaseola is known from late Llandovery and Sheinwoodian strata, Silurian of Britain (Lundin

et al., op. cit., 1991).

We gratefully acknowledge support from the National Science Foundation (Grant No.

EAR-8200816).

Explanation of Plate 23, 44

Figs. 1-4, car. (ASU X-278, 1053/ym long, 526 pm high): fig. 1, ext. It. lat.; fig. 2, ext. rt. lat.; fig. 3, ext. vent.; fig. 4, ext. post.

Figs. 5, 6, car. (ASU X-147, 921 pm long); fig. 5, ext. It. lat.; fig. 6, ext. rt. lat.

Scale A (200 /ym; x49), figs. 1-4; scale B (200 /ym; x57), figs. 5, 6.

Stereo-Atlas of Ostracod Shells 23, 42

Wenlockiella phaseola (2 of 4)

Stereo-Atlas of Ostracod Shells 23, 44

Wenlockiella phaseola (4 of 4)

Stereo- Atlas of Ostracod Shells 23 (11) 45-48 (1996)

595.337.21 (113.331) (420 : 162.003.52) : 551.351

Cytherellina elegans (1 of 4)

ON CYTHERELLINA ELEGANS (JONES)

by Lee E. Petersen and Robert F. Lundin

(Anardarko Petroleum Corporation, Houston, and Arizona State University, Tempe, U.S.A.)

Cytherellina elegans (Jones, 1887)

1887 Macrocypris elegans sp. nov. T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 180, pi. 5, figs. 8a-c.

1887 Macrocypris siliquoides sp. nov. T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 181, pi. 5, figs. 9a-c.

1887 Bythocypris concinna sp. nov. T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 186, pi. 5, figs. 6a-c.

1887 Bythocypris testacella sp. nov. T.R. Jones, Ann. Mag. nat. Hist., (5), 19, 186, pi. 5, figs. 5a-c. pi. 3, figs. 1, 2.

1991 "Cytherellina” elegans (Jones); R.F. Lundin, L.E. Petersen and D.J. Siveter, J. Micropalaeontol. 9 (part 2 for 1990), 179, pi. 1, fig. 8.

Holotype: The Natural History Museum (NHM), London, England, no. 1 1911; adult carapace. This was the only

specimen available to Jones (1887, op. cit.), and it agrees well with his illustration.

Type locality: Vine Collection no. Ill, Bed no. 40, “Buildwas Beds”, as reported by Jones (1887, op. cit.). This Silurian

collection is reported by Vine ( Proc . Yorks, geol. polytech. Soc., 9, 224-248, 1887) to be from the “banks

of the River Severn, above Buildwas Bridge,” making it approximately equivalent to loc. 34 of Lundin

et al., (1991, op. cit.)-, approximately lat. 52° 38' 15" N, long. 2° 31' 30" W (National Grid Ref.

SJ 6451 0445). See comments below under Distribution.

Figured specimens: Department of Geology, Arizona State University, (ASU), nos. X-130 (adult car.: PI. 23, 48, figs. 3-5),

X-312 (adult car.: PI. 23, 48, figs. 1, 2) and X-313 (adult car.: PI. 23, 46, figs. 1-4). Specimen NHM

1 1911 (holotype, adult car.: PI. 23, 46, figs. 5, 6).

Explanation of Plate 23, 46

Figs. 1-4, car. (ASU X-313, 1165 pm long): fig. 1, ext. rt. lat.; fig. 2, ext. It. lat.; fig. 3, ext. vent.; fig. 4, ext. post. Figs. 5, 6, car.

(holotype, NHM 1 1911, 1180//m long): fig. 5, ext. rt. lat; fig. 6, ext. vent.

Scale A (200 pm; x45), figs. 1-4; scale B (200 //m; x43), figs. 5, 6.

Stereo-Atlas of Ostracod Shells 23, 47

Cytherellina elegans (3 of 4)

Diagnosis:

Remarks:

Distribution:

A cknowledgements:

NHM 1 1911 is from the type locality. ASU X-130, X-312 and X-313 are from the Much Wenlock

Limestone Formation at Lincoln Hill (loc. 49 of Lundin, Petersen and Siveter, 1991, op. cit.),

Shropshire; approximately lat. 52° 38' N, long. 2° 29' W (National Grid Ref. SJ 6693 0381); Homerian

Stage, Wenlock Series, Silurian.

Relatively elongate, narrow Cytherellina with a poorly developed posterior straguloid process, and

ventriculus with distinct anterior boundary but indistinct posterior boundary. Left/right overreach weak

but best developed antero- and posterodorsally and midventrally. Surface smooth. Details of hinge and

contact margin unknown.

We have studied the type specimens of Macrocypris siliquoides, Bythocypris concinna and Bythocypris

testacella, all of which were erected by Jones (1887, op. cit.; see synonymy above). We conclude that

these specimens are conspecific with C. elegans. The latter is readily distinguished from Cytherellina

jonesi Petersen and Lundin ( Stereo-Atlas of Ostracod Shells, 23, 49-52, 1996) by its smaller

height/length and width/length ratios.

Known from seven samples of late Wenlock, Homerian, age and from one sample (locality no. 59 of

Lundin, Petersen and Siveter 1991, op. cit.) of Ludlow, early Gorstian, age in the Welsh Borderland.

Jones (1887, op. cit.) reported the holotype and one of his two specimens of Macrocypris siliquoides, here

placed in synonymy with C. elegans, to be from the “Buildwas Beds” and thus of Sheinwoodian, early

Wenlock age. We have examined some thirty-six samples and identified more than 3000 non-palaeocope

ostracode specimens from the Sheinwoodian of the Wenlock type area and have never found a specimen

of C. elegans from that stratigraphic horizon. We conclude that the anomalous occurrences reported by

Jones (1887, op. cit.) are the result of contamination from Homerian strata or collections.

We gratefully acknowledge the support of the College of Liberal Arts and Sciences, Arizona State

University and the help of David J. Siveter in determining geographic and stratigraphic positions of some

of the collections described by T.R. Jones.

Explanation of Plate 23, 48

Figs. 1, 2, car. (ASU X-312, 1109//m long): fig. 1, ext. vent.; fig. 2, ext. rt. lat. Figs. 3-5, car. (ASU X-130, 1165/rm long): fig. 3,

ext. rt. lat.; fig. 4, ext. dors.; fig. 5, ext. It. lat.

Scale A (200 pm; x46), figs. 1, 2; scale B (200 pm; x49), figs. 3-5.

Stereo-Atlas of Ostracod Shells 23, 46

Cytherellina elegans (2 of 4)

Stereo-Atlas of Ostracod Shells 23, 48

Cytherellina elegans (4 of 4)

Stereo-Atlas of Ostracod Shells 23 (12) 49-52 (1996)

595.337.21 (113.333 + 113.331) (420: 162.003.52) : 551.351

Cytherellina ruperti (1 of 4)

ON CYTHERELLINA RUPERTI sp. nov.

by Lee E. Petersen and Robert F. Lundin

(Anadarko Petroleum Corporation, Houston, U.S.A. and

Arizona State University, Tempe, U.S.A.)

Cytherellina ruperti sp. nov.

1869 Cytherellina siliqua (Jones); T.R. Jones and H.B. Holl, Ann. Mag. nat. Hist., (4), 3, 216, pi. 14, fig. 2, ? fig. 5.

1991 “Cytherellina” sp. nov. R.F. Lundin, L.E. Petersen and D.J. Siveter, J. Micropalaeontol., 9 (pt. 2 for 1990), pi. 1, fig. 9.

Holotype:

Type locality:

Derivation of name:

Figured specimens:

The National History Museum (NHM), London, England, OS 14898 (ex., no. 12069); adult carapace.

[Paratypes: Department of Geology, Arizona State University (ASU), nos. X-131, X-280 and X-282],

Chance’s Pitch, a road section about 2 km W of Little Malvern, Hereford and Worcestershire, England;

approximately lat. 52°03'N, long. 2° 18' W. National Grid Ref.: SO747402; Aymestry Limestone, Gorstian

Stage, Ludlow Series, Silurian.

In honour of Professor Thomas Rupert Jones, doyen of British micropalaeontology in the 19th century.

Department of Geology, Arizona State University (ASU), nos. X-131 (paratype, adult car.: PI. 23, 50, figs. 1-4),

X-282 (paratype, adult car.: PI. 23, 50, fig. 5), X-280 (paratype, adult car.: PI. 23, 52, figs. 1-3), and

NHM OS 14898 (holotype, adult car.: PI. 23, 52, figs. 4, 5).

NMH OS 14898 is from the type locality. ASU X-131 and X-280 are from the Much Wenlock Limestone

Formation at Croft Farm 0.5 km W of West Malvern, Hereford and Worcestershire; approximately lat.

52°08'N, long. 2° 18' W (National Grid Ref. SO75674650); X-282 is from the Much Wenlock Limestone

Formation at Wren’s Nest, Dudley, West Midlands; approximately lat. 52° 27' N, long. 2° 3' W (National Grid

Ref. S093579199); Homerian Stage, Wenlock Series, Silurian.

Explanation of Plate 23, 50

Figs. 1-4, car. (paratype, ASU X-131, 1109 /tm long): fig. 1, ext. post.; fig. 2, ext. vent.; fig. 3, ext. dors.; fig. 4, ext. rt. lat. Fig. 5,

LV (paratype, ASU X-282, 1128/ym long), int. lat.

Scale A (200 pm, x47), figs. 1-5.

Stereo-Atlas of Ostracod Shells 23, 51 Cytherellina ruperti (3 of 4)

Diagnosis: Cytherellina species with poorly developed adductorial recess, moderately developed ventriculus with distinct

anterior boundary but indistinct posterior boundary. Posterior straguloid process weak; commissure anterior to

hinge straight. Short stop ridge present along ventral contact margin just behind midlength. Surface very finely

striate.

Remarks: In establishing the genus Cytherellina Jones and Holl (1869, op. cit.) emphasized the “undulated contours”

which are present on steinkerns of the type-species, C. siliqua (Jones, 1855). These “undulations” are the

reflection of a large, well-developed adductorial recess on the interior surface of the valves of this species. The

development of the adductorial recess is, however, variable between species as shown by C. ruperti and another

congeneric species from the Hemse Beds (Ludlow Series) of Gotland. The Gotland species has a small distinct

adductorial recess but it is less well-developed than that of the type-species, whereas the adductorial recess of

C. ruperti is very poorly developed. The species described here also differs from the type-species in its much

smaller size, and from the Gotland species in its distinctly less-arched dorsal margin. Two species erected by

A. Pranskevichius {Lithuanian Sci-Res. Geol. Surv. Inst., SSR, Trans. 15, 110, 111, 1972), namely Healdianella

piriformis and Healdianella virbalica, are similar to C. ruperti and certainly belong to Cytherellina. The first of

these species differs from C. ruperti in having a distinctly sinuate ventral margin and more sharply rounded

anterior margin. C. virbalica has a more bluntly rounded anterior margin and less left/right ventral overreach

than C. ruperti.

Jones and Holl (1869, op. cit., pi. 14, fig. 5) illustrated another specimen, from basal upper Ludlow beds near

the type locality, which they concluded to be conspecific with the holotype. We have not seen this specimen but,

on the basis of Jones and Holl’s illustration, we agree with question.

Specimens of C. ruperti are normally abraided. However, the excellent preserved holotype clearly shows that

the species is very finely striate.

Distribution: This species is known from strata of late Wenlock (Homerian), as reported by Lundin, Petersen and Siveter

(1991, op. cit.), through early (and possibly early late) Ludlow, Silurian age of the English West Midlands and

Welsh Borderland.

Explanation of Plate 23, 52

Figs. 1-3, car. (paratype, ASU X-280, 1222 pm long): fig. 1, ext. rt. lat.; fig. 2, ext. It. lat.; fig. 3, ext. vent. Figs. 4, 5, car. (holotype,

NMH OS 14898, 1250 pm long): fig. 4, ext. rt. lat.; fig. 5, ext. vent.

Scale A (200 pm; x43), figs. 1-3; scale B (200 pm; x41), figs. 4, 5.

Cytherellina ruperti (2 of 4)

Cytherellina ruperti (4 of 4)

Stereo-Atlas of Ostracod Shells 23, 52

Stereo-Atlas of Ostracod Shells 23, 50

Stereo-Atlas of Ostracod Shells 23 (13) 53-60 (1996)

595.337.21 (116.212) (430 : 162.009.52) : 551.35 + 552.52

Ogmoconcha contractula (1 of 8)

ON OGMOCONCHA CONTRACTULA TRIEBEL

by Ian Boomer and Thomas Jellinek

(University of East Anglia, Norwich, England and Senckenberg Museum, Frankfurt, Germany)

Genus OGMOCONCHA Triebel, 1941

Type-species (by orginal designation): Ogmoconcha contractula Triebel, 1941

Diagnosis: (Original) A genus of the Healdiidae Harlton, 1933 with the following characteristics. Carapace inflated,

egg-shaped, usually with marginal denticles. Lacking a steeply sloping postero-dorsal margin and without

a vertical flexture in the posterior part of the carapace. (Additional) Carapace heavily calcified, overlap

entire and well marked, central margin broadly convex particularly in the larger left valve. Contact groove

in the left valve is entire. Hinge elements broad and inflated terminally, bearing many fine transverse

crenulae. Adductor muscle pattern consists of a double row of 3-7 central scars surrounded by a single

outer ring of 12-13 smaller scars, a single rounded frontal muscle scar is present (note that the develop-

ment of the muscle scar pattern will be strongly influenced by preservation). Greatest width at or just

behind midlength. External lateral surfaces generally unornamented although a few shallow pits may

occur, small marginal spines common anteriorly and posteroventrally.

Remarks: Much discussion has centred around the validity of this genus and its possible synonymy with Triassic

Hungarella (Mehes, 1911) and Liassic Ogmoconchella Griindel, 1971 (Lord, A.R., Bull. geol. Soc.

Denmark, 21, 319-336, 1972; Malz, H., Senckenberg. leth., 52, 433-455, 1971). The possible synonymy

with Hungarella is unlikely given the large number of scars in the central muscle field of the type

specimen which was figured by Lord (1972, ibid.). The possible synonymy between Ogmoconcha and

Explanation of Plate 23, 54

Fig. RV, ext. lat. (paratype, Xel268, 780 //m long). Figs. 2, 3, car, (holotype, Xel249, 830 /im long): fig. 2, dors.; fig. 3, rt. lat.

Scale A (100 pm\ x70), figs. 1-3.

Stereo-Atlas of Ostracod Shells 23, 55 Ogmoconcha contractula (3 of 8)

Ogmoconchella is considered unproven by the present authors. Although their central muscle scar patterns

are similar, there are a number of features which consistently distinguish these two genera. The position

of greatest height is in front of midlength in Ogmoconcha while it is at or behind midlength in Ogmocon-

chella. The greatest width is at or just behind midlength in Ogmoconcha while it is towards the posterior

in Ogmoconchella. In the former genus the anterior margin is more broadly rounded than the posterior

while in the latter genus the reverse is true. Hingement is always stronger in Ogmoconcha with terminal

widening and well developed crenulations. Externally Ogmoconcha is almost always smooth with only

marginal spines and occasionally a few shallow puncta (as seen in O. hagenowi Drexler, 1958). Ogmocon-

chella, however, may possess a fine “fingerprint” ornament (although this is only observed in well

preserved specimens), posteroventral spines and anteromarginal flanges may also be developed.

Ogmoconcha may be synonymous with some of the Triassic genera described by Kristan-Tollmann from

the European Alps. A comprehensive examination of the genotype material from Kristan-Tollmann’s

collections must be undertaken to establish their validity. In summary, the genus is here considered to be

a valid taxon distinct from both Hungarella and Ogmoconchella.

Ogmoconcha contractula Triebel

1941 Ogmoconcha contractula gen. et sp. nov. E. Triebel, Senckenbergiana , 23, 378, pi. 14, figs, 156-160.

Holotype: Senckenberg Museum, Frankfurt, Germany, no. Xel249, adult carapace. [Paratypes: nos. Xel248,

Xel250, Xel251, Xel267-Xel276].

Type locality: Lias <5, Upper Pliensbachian, Hambiihren Borehole WA2 (depth 495-503 m), Hannover, Germany.

Figured specimens: Senckenberg Museum, Frankfurt, Germany, nos. Xel268 (paratype, RV: PI. 23, 54, fig. 1; pi. 23, 58, figs.

1, 3; PI. 23, 60 figs. 1, 2), Xel249 (holotype, car.: PL 23, 54, figs. 2, 3; PI. 23, 56, fig. 2). Xel248 (para-

type, LV: PL 23, 56, figs. 1, 4; PI. 23, 58, figs. 2, 4; PL 23, 60, fig. 3), Xel274 (paratype, car.: PI. 23, 56,

fig. 3). All specimens from type locality and horizon.

Explanation of Plate 23, 56

Figs. 1, 4, LV (paratype, Xel248, 775 pm long): fig. 1, detail of muse, sc.: fig. 4, ext. lat. Fig. 2, car. It. lat. (holotype Xel249, 830 pm

long). Fig. 3, car. dors, (paratype, Xel274, 830 pm long).

Scale A (20 pm\ x460), fig. 1; scale B (100 pm\ x70), figs. 2-4.

Stereo-Atlas of Ostracod Shells 23, 54

Ogmoconcha contractula (2 of 8)

Stereo-Atlas of Ostracod Shells 23, 57

Ogmoconcha contractula (5 of 8)

Diagnosis:

Remarks:

Distribution:

A species of Ogmoconcha with the following characteristics. Valves bear weak anteromarginal denticles,

lateral surfaces compressed about the muscle scars so that in dorsal view the lateral extremities of the

carapace appear concave (PI. 23. 54, fig. 2). Adductor muscle scar and hinge details as for genus.

Park ( Stereo-Atlas Ostracod Shells , 11, 67-70, 1984) described Ogmoconcha eocontractula from the

Pliensbachian of the southern North Sea Basin. O. eocontractula predates the present species and is

distinguished by its larger size and the different lateral outline of each valve. It is almost certainly ancetral

to O. contractula as it is the only other known liassic species with a laterally compressed carapace. Boomer

(/. Micropalaeont., 9, 205-218, 1991) described O. convexa from the lower Toarcian of the Mochras

Borehole, Wales. That species is somewhat smaller than O. contractula but similar in lateral outline, it

differs, however, in that the lateral faces of the carapaces are rounded and not flattened. Boomer

(J. Micropalaeont., 10, 47-57, 1992) recorded a number of metacopine taxa from the Lower Toarcian of

SW England which are smaller than O. convexa, and possess weakly compressed lateral surfaces.

Known from the Pliensbachian and Lower Toarcian of Northwest Europe.

Explanation of Plate 23, 58

Figs. 1, 3, RV (paratype, Xel268, 780 pm long): fig. 1, ant. hinge detail (arrow points anterior); fig. 3, detail of hinge crenulation. Figs.

2, 4, LV (paratype, Xel248, 775 n m long): (arrow points anterior) fig. 2, ant. hinge detail; fig. 4, post, hinge detail. Scale A

(100 fm\; xl30), fig. 1; scale B (100 /tm; xl45), figs. 2, 4; scale C (25 gm; x450), fig. 3.

Stereo-Atlas of Ostracod Shells 23, 59

Ogmoconcha contractula (7 of 8)

A

0

Text-fig. 1. Details of adductor muscle scar patterns taken from published images. A. Xel268 RV, x200. B. Xe8414 RV, x300. C. Xel248 LV, x300.

D. Xel269 LV, x250. E. Xel251, LV, x250 (all magnifications approximate).

Explanation of Plate 23, 60

Figs. 1, 2, RV (paratype, Xel268, 780 /tm long): fig. 1, post, hinge detail (arrow points anterior); fig. 2, int. lat. Fig. 3, LV int. lat.

(paratype, Xel248, 775 //m long).

Scale A (50 gm; X205), fig. 1; scale B (100 gm; x71), figs. 2, 3..

Stereo-Atlas of Ostracod Shells 23 (14) 61-68 (1996)

595.337.12 (119.9) (420 : 162.001.52) : 551 .312

Eucypris virens (1 of 8)

ON EUCYPRIS VIRENS (JURINE)

by Robin Smith and Koen Martens

(Department of Geology, University of Leicester, U.K. and

Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium)

Genus Eucypris Vavra, 1891

1891 Eucypris gen. nov. W. Vavra, Arch. Naturwissensch. Landesdurchforsch. Boehmen, 8(3), 82.

Diagnosis: Medium-sized (1-2.5 mm long) genus of the Eucypridinae Bronstein, 1947. Carapace shape elliptical, rounded in lateral view;

valves with external (not marginal) tubercles carrying hairs (“porenwarzen”) anteriorly; not flattened ventrally, ventral margin

sinuously curved in anterior third. Calcified inner lamella broad in both valves, anterior vestibulum approximately 12 times as

wide as the fused zone; selvages (if present) marginal, frontal inner lists possible, but always submarginal. M x 1 with second palp

segment cylindrical and curved (length approx, twice the basal width). Gamma-seta on Md-palp approx, ten times as long as basal

width. Seta dl on T1 approx, three times as long as d2.

Remarks: Martens (Arch. Hydrobiol., Suppl., 83, 227-251, 1989) characterized the tribe Eucypridini in the subfamily Eucypridinae

Bronstein, 1947, and retained four genera: Eucypris Vavra, 1891, Prionocypris Brady and Norman, 1896, Tonnacypris Diebel

and Pietrzeniuk, 1975 and Tranjancypris Martens, 1989. Martens et at. (Zool. Middle East, 7, 95-114, 1992) added the genus

Eucyprinotus Sywula, 1972 to the tribe. All of these genera are united by the presence of a “c”-seta on the Mx2; they can be

separated from each other by the outline of the valve margin and the length ratio of setae dl and d2 on Tl. The genus Eucypris

is chacterised by its wide calcified inner lamella, the submarginal inner lists and the absence of selvages, the cylindrical second

palp on the MX 1 and the length of seta dl compared to d2.

Explanation of Plate 23, 62

Fig. 1 . 9 LV, ext. lat. (OC 2002, 1520 /um long); fig. 2, 9 RV, ext. lat. (OC 2002, 1480 pm long); fig. 3, 9 car. ventr. (OC 2004, 1530 pm

long); fig. 4, 9 car. dors. (OC2004, 1530/rm long).

Scale a (500 /rm; x40), figs. 1-4.

Stereo- Atlas of Ostracod Shells 23, 63

Eucypris virens (3 of 8)

Eucypris virens (Jurine, 1820)

1820 Monoculus virens sp. nov. L. Jurine. Histoire des monocles qui se trouvent aux environs de Geneve, 174, pi. 18, figs. 15-16. Geneve/Paris.

1825 Cypris virens (Jurine); A.-G. Desmarest, Considerations generates sur la classe des crustaces, et description des especes de ces animaux, qui vivent dans la mer, sur

les cotes, ou dans les eaux douces de la France, 384, Paris.

1891 Cypris (Eucypris) virens (Jurine); W. Vavra, Arch. Naturwissensch. Landesdurchforsch. Boehmen, 8(3), 102.

1900 Eucypris virens (Jurine); E. von Daday, Ostracoda Hungaria, 143, Budapest.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

A ckno wledgements:

No type specimens are believed to exist.

Not known precisely; in the surroundings of Geneva, Switzerland.

Royal Belgian Institute of Natural Sciences (Brussels, Belgium), Ostracod Collection, nos. OC2002 (9 LV and RV: PI. 23, 62,

figs. 1, 2; PI. 23, 64, figs. 1, 4; Text-figs la-c, e; Text-figs. 2b-j), OC 2003(9: Text-fig. Id; Text-fig. 2a), OC2004 (9 car: PI.

23, 62, figs. 3, 4; PI. 23, 64, fig. 2), OC 2005 (9 LV and RV: PI. 23, 66, figs. 1-4; PI. 23, 68, figs. 3, 4), OC 2006 (9 RV: PI. 23,

68, figs. 1, 2). All specimens collected on 2/3/96 from a shallow (<20cm) temporary pool in Ketton Quarry, Lincolnshire,

England (lat. 52° 38' N, long. 0° 33' W), pH 8.3, temp. 9 °C.

Adult shell 1.6-2. 3 mm long, colour green in living specimens. Viewed dorsally the carapace is more pointed anteriorly than

posteriorly but lacks compressed or flattened extremities; greatest width behind midlength.

The taxonomy of the Genus Eucypris s.s. is confused: intraspecific variability is high (four subspecies have been described in

E. virens — all of these fit into the variability range of the species) and species are distinguished from each other on the shape and

size of the carapace, length ratio of furcal claws and ramus, etc., not on anatomical differences. A revision of the species of

Eucypris is urgently required (Martens and Baltanas, in prep.); the present redescription of the type species is intended as a primer

to this work. Most European populations are parthenogentic, and only females are described herein. Sexual populations are

known, however, from North Africa, Spain and Sicily; descriptions of males and females from sexual populations will be

described in a future paper. E. virens is widespread and common in Europe, usually being found in temporary freshwater ponds,

a review of the ecology, distribution and reproduction of this species was recently provided by A. Baltanas (in: D.J. Horne and

K. Martens (eds), The Evolutionary Ecology of Reproductive Modes in Non-marine Ostracoda, Greenwich University Press,

9-16, 1994).

This work has been supported by the E.U. Human Capital and Mobility Programe (contract ERBCHRXCT/93/0253). We thank

C. Behen (Brussels) for technical assistance with the line drawings.

Explanation of Plate 23, 64

Fig. 1, 9 RV, ext. lat., detail of anterior region (OC2002); fig. 2, 9 car. post. (OC2004, 1530 /tm long); fig. 3, 9 car. ant. (specimen

lost); fig. 4, 9 RV, ext. lat., detail of anterior margin (OC2002).

Scale A (50/um; xl80), fig. 1; scale b (500 pm\ x45), figs. 2, 3; scale C (20 pm\ x600), fig. 4.

Eucypris virens (4 of 8)

Stereo-Atlas of Ostracod Shells 23, 62

Eucypris virens (2 of 8)

Stereo-Atlas of Ostracod Shells 23, 64

Stereo-Atlas of Ostracod Shells 23 (14) 61-68 (1996)

595.337.12 (119.9) (420 : 162.001.52) : 551 .312

Eucypris virens (1 of 8)

ON EUCYPRIS VIRENS (JURINE)

by Robin Smith and Koen Martens

(Department of Geology, University of Leicester, U.K. and

Royal Belgian Institute of Natural Sciences, Vautierstraat 29, 1000 Brussels, Belgium)

Genus Eucypris Vavra, 1891

1891 Eucypris gen. nov. W. Vavra, Arch. Naturwissensch. Landesdurchforsch. Boehmen, 8(3), 82.

Diagnosis: Medium-sized (1-2.5 mm long) genus of the Eucypridinae Bronstein, 1947. Carapace shape elliptical, rounded in lateral view;

valves with external (not marginal) tubercles carrying hairs (“porenwarzen”) anteriorly; not flattened ventrally, ventral margin

sinuously curved in anterior third. Calcified inner lamella broad in both valves, anterior vestibulum approximately 12 times as

wide as the fused zone; selvages (if present) marginal, frontal inner lists possible, but always submarginal. M x 1 with second palp

segment cylindrical and curved (length approx, twice the basal width). Gamma-seta on Md-palp approx, ten times as long as basal

width. Seta dl on T1 approx, three times as long as d2.

Remarks: Martens ( Arch . Hydrobiol., Suppl., 83, 227-251, 1989) characterized the tribe Eucypridini in the subfamily Eucypridinae

Bronstein, 1947, and retained four genera: Eucypris Vavra, 1891, Prionocypris Brady and Norman, 1896, Tonnacypris Diebel

and Pietrzeniuk, 1975 and Tranjancypris Martens, 1989. Martens et at. (Zool. Middle East, 7, 95-114, 1992) added the genus

Eucyprinotus Sywula, 1972 to the tribe. All of these genera are united by the presence of a “c”-seta on the M x 2; they can be

separated from each other by the outline of the valve margin and the length ratio of setae dl and d2 on Tl. The genus Eucypris

is chacterised by its wide calcified inner lamella, the submarginal inner lists and the absence of selvages, the cylindrical second

palp on the Mx 1 and the length of seta dl compared to d2.

Explanation of Plate 23, 62

Fig. 1. 9 LV, ext. lat. (OC 2002, 1520 pm long); fig. 2, 9 RV, ext. lat. (OC 2002, 1480 /tm long); fig. 3, 9 car. ventr. (OC 2004, 1530 pm

long); fig. 4, 9 car. dors. (OC2004, 1530/rm long).

Scale a (500 pm; x40), figs. 1-4.

Stereo-Atlas of Ostracod Shells 23, 63

Eucypris virens (3 of 8)

Eucypris virens (Jurine, 1820)

1820 Monoculus virens sp. nov. L. Jurine. Histoire des monocles qui se trouvent aux environs de Geneve , 174, pi. 18, figs. 15-16. Geneve/Paris.

1825 Cypris virens (Jurine); A.-G. Desmarest, Considerations generates sur la classe des crustaces, et description des especes de ces animaux, qui vivent dans la mer, sur

les cotes, ou dans les eaux douces de la France, 384, Paris.

1891 Cypris (Eucypris) virens (Jurine); W. Vavra, Arch. Naturwissensch. Landesdurchforsch. Boehmen, 8(3), 102.

1900 Eucypris virens (Jurine); E. von Daday, Ostracoda Hungaria, 143, Budapest.

Holotype:

Type locality:

Figured specimens:

Diagnosis:

Remarks:

A ckn o wledgemen ts:

No type specimens are believed to exist.

Not known precisely; in the surroundings of Geneva, Switzerland.

Royal Belgian Institute of Natural Sciences (Brussels, Belgium), Ostracod Collection, nos. OC 2002 (9 LV and RV: PI. 23, 62,

figs. 1, 2; PI. 23, 64, figs. 1, 4; Text-figs la-c, e; Text-figs. 2b— j), OC2003(9: Text-fig. Id; Text-fig. 2a), OC2004 (9 car: PI.

23, 62, figs. 3, 4; PI. 23, 64, fig. 2), OC2005 (9 LV and RV: PI. 23, 66, figs. 1-4; PI. 23, 68, figs. 3, 4), OC2006 (9 RV: PI. 23,