A Stereo -At las of Ostracod Shells
edited by R.H. Bate, J. W. Neale, Lesley M. Sheppard
and David J. Siveter
Volume 6 , Part 2 ; 29th November 1979
Published by The British Micropalaeontological Society
in association with Robertson Research International Ltd., Llandudno, Wales
Editors
Dr. R.H. Bate, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.
Prof. J. W. Neale, Department of Geology, The University, Hull HU6 7RH.
Ms. Lesley M. Sheppard, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.
Dr. David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.
Editorial Board
Dr. Richard H. Benson, Smithsonian Institution, Washington, D.C., 20560. U.S.A.
Dr. Alwine Bertels, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires,
Argentina.
Dr. K. Ishizaki, Institute of Geology and Paleontology, Tohoku University, Sendai, Japan.
Dr. C.W. Haskins, Robertson Research International Limited, ‘Ty’n-y-Coed’, Llanrhos, Llandudno,
N. Wales, LL30 ISA.
Dr. P.J. Jones, Bureau of Mineral Resources, P.O. Box 378, Canberra City, A.C.T 2601 , Australia.
Prof. Dr. E. Kempf, Geologisches Institut der Universitat Koln, Ziilpicher Strasse 49, D-5 Koln 1,
German Federal Republic.
Dr. H.J. Oertli, S.N.P.A., Centre de Recherches, 64001 Pau, France.
Prof. G. Ruggieri, Instituto e Museo di Geologia dell’Universita di Palermo, Corso Tukory, 131,
90134 Palermo, Italy.
Mr. P.F. Sherrington, Petro-Canada, P.O. Box 2844, Calgary, Alberta T2P 2M7, Canada.
Instructions to Authors
Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited.
Full instructions may be obtained on request from any one of the Editors or Editorial Board. Format
should follow the style set by the majority of papers in this issue. Descriptive matter apart from
illustrations should be cut to a minimum; preferably each plate should be accompanied by one page
of text only. Blanks to aid in mounting figures for plates may be obtained from the Editors.
Completed papers should be sent to Ms. L.M. Sheppard, Department of Palaeontology, British
Museum (Natural History), Cromwell Road, London SW7 5BD.
Acknowledgments
This Volume of the Stereo-Atlas has been aided by generous financial support from Robertson
Research International Limited.
Stereo -viewing for users of the Atlas
In order to obtain maximum information and benefit from the use of the Stereo-Atlas it is essential
that the user view the micrographs stereoscopically. Small pocket-sized stereo-viewers are most
suitable for this purpose. Two suppliers are:
C.F. Casella & Co. Ltd., Regent House, Britannia Walk, London N 1 7ND, and
Air Photo Supply Corpn., 158 South Station, Yonkers, New York 10705. U.S.A.
The front cover shows a female (external & internal views) of Cativella bensoni Neale
Plates reproduced by Torr of Silverstone, Northants., England.
-+t~r-
--M
Stereo-Atlas of Ostracod Shells 6(14)71 - 74 (1979)
595.336.13 (1 13.312) (438:161.015.54) : 551.35 + 552.55
Brevibolbina dornbuschi (1 of 4)
ON BREVIBOLBINA DORNBUSCHI SCHALLREUTER
by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)
Brevibolbina dornbuschi Schallreuter, 1964
1964 Brevibolbina dornbuschi sp. nov. R. E. L. Schallreuter, Ber. geol. Ges. D. D. R., 9,381, 442, pi. 9, fig. 2.
1973 Brevibolbina dornbuschi Schallreuter; R. E. L. Schallreuter, Palaeontographica (A), 144 (1/3), 74, 75, tab. 5, pi. 16,
figs. 1, 2, pi. 20, figs, 7, 8 (q.v. for further synonymy).
1973 Brevibolbina dornbuschi Schallreuter; W. Neben & H. H. Krueger, Staringia 2 (Bijvoegsel van Grondboor en hamer, 6),
pi. 90, fig. 2 (= Schallreuter, op. cit . , pi. 16, fig. 1 ).
1976 Brevibolbina dornbuschi Schallreuter; R. E. L. Schallreuter , Palaeontographica (A), 153 (4/6), 164.
Holotype: Department of Geological Sciences, University of Greifswald, German Democratic Republic, no. 2/2
(Os 169), 9 LV.
Type locality: Beach at Dornbusch, Isle of Hiddensee (Baltic Sea); lat. 54° 36' N, long. 13° 7' E. Backsteinkalk erratic
boulder ( 1 B 1 3 Type, no. 1 B 1 3), Middle Ordovician.
Explanation of Plate 6, 72
Fig. 1 , 9 LV, ext. lat. (GPIH 2207, ISO pm long); figs. 2 - 4, 9 LV (GPIH 2208, 720 pm long): fig. 2, int. ant. obi.; fig. 3, ext.
vent.; fig. 4, int. lat.
Scale A (250 pm\ x 90), figs. 1,2,4; scale B (250 pm\ x 95), fig. 3.
=A
4— b
44-
-44-4
"4--*-
Stereo-Atlas of Ostracod Shells 6, 73
Brevibolbina dornbuschi (3 of 4)
Figured specimens: Geologisch-Palaontologisches Institut, University of Hamburg (GPIH), nos. 2207 (9 LV: PI. 6, 72, fig. 1),
2208 (9 LV: PI. 6, 72, figs. 2 - 4), 2209 (juv. tecnomorphic LV: PI. 6, 74, fig. 1), 2210 (juv. tecnomorphic
LV : PI. 6, 74, fig. 2) and 221 1 (juv. tecnomorphic RV: PI. 6, 74, fig. 3).A11 the figured specimens are from
Backsteinkalk erratic boulder no. Ho - 2 (for further data see Schallreuter 1976, op. cit., 164); Middle
Ordovician of Baltoscandia.
Diagnosis: Adults 0.65 - 0.80mm long. S2 distinct, sinuous, open at both ends, behind the relatively small, conical
preadductorial node is more or less pit-like, below the node appears as a shallow sulcal depression.
Posteroventral lobe distinct, oblong, elevated projected posteriorly as a lobal spine (Stachel) and near its
termination bearing a hollow, ornamental spine (dorn). Dolon occupies the antero- and centroventral
regions of females, terminating posteriorly adjacent to the base of the posteroventral lobe; between dolon
and posteroventral lobe there is an open U-shaped laterovelar furrow. Surface reticulogranulose with some
tubercles.
Remarks: The ornamental spine on the posteroventral lobal spine was not mentioned in the previous detailed
description of the species (Schallreuter 1973).
Distribution: Lower part of the Keila Stage (D2o:), Isle of Dago (= Hiiumaa; Estonia), Baltic Sea. Backsteinkalk erratic
boulders of N Germany; Sandoflint (= Baltic Backsteinkalk) of the Isle of Gotland, Baltic Sea (1B3,
1 B13, 14B2 Types).
Explanation of Plate 6, 74
Fig. 1 , tecnomorphic juv. LV, ext. lat. (GPIH 2209, 6 1 3 pm long); fig. 2, tecnomorphic juv. LV, ext. lat. (GPIH 2210, 5 1 5 pm
long); fig. 3, tecnomorphic juv. RV, ext. lat. (GPIH 2211, 488 pm long).
Scale A (100 pm \ x 125), fig. 1 ; scale B (100 pm \ x 110), fig. 2; scale C ( 1 00 pm \ x 148), fig. 3.
-44-
{--1-4—
Stereo-Atlas of Ostracod Shells 6 ( 1 5) 75 - 78 (1979)
595.336.13 (1 13.312) (438:161.015.54) : 551.35 + 552.55
Homeokiesowia frigida (1 of 4)
ON HOMEOKIESOWIA FRIGIDA (SARV)
by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)
Derivation of name:
Diagnosis:
Genus HOMEOKIESOWIA gen.nov.
Type-species: Kiesowia frigida Sarv, 1959
Referring to the homeomorphy with Kiesowia Ulrich & Bassler, 1908. Gender, feminine.
A medium-sized, quadrilobate genus of Tallinnellinae. Lobes dissolved into single nodes: three nodes in
front of S2 and four behind; node occupying site of ventral part of L3 (= “posteroventral lobe”)
strongest; posteroventral-most node (= L4 vent.) smallest, is sometimes (especially in larvae) represented
only as a tubercle, and can be fused with the posterodorsal node (L4 dors.). At the dorsal border above
LI and L2 two small node- or spine-like tubercles may occur. Marginal sculpture represented as a row of
spines.
Remarks: The genus differs from other genera of Tallinellinae by having lobes dissolved into single nodes. The
homeomorph Kiesowia Ulrich & Bassler, 1908 belongs to the Sigmoopsinae because it develops a histium
(see Schallreuter, Stereo-Atlas of Ostracod Shells 6 (16) 79 - 86, 1979). The only additional taxon
referred to Homeokiesowia is Kiesowia pernodosa Opik (Ann. Naturalists Soc. Tartu Univ. 43 (1/2), 95,
1937 ;Publ. Geol. Inst. Univ. Tartu 50, 31, 1937), a poorly known species.
Distribution: The Middle Ordovician (Viru Series) of Baltoscandia.
Explanation of Plate 6, 76
Figs. 1-3,9 RV (GPIH 2023a, 1110 pm long): fig. 1 , ext. vent, obi; fig. 2, ext. ant. obi.; fig. 3, ext. lat.
Scale A (250 pm\ x 72), figs. 1,2; scale B (250 pm', x 79), fig. 3.
r4-F-
-+4-r
I
Stereo-Atlas of Ostracod Shells 6, 77
Homeokiesowia frigida (3 of 4)
Homeokiesowia frigida (Sarv, 1959)
1959 Kiesowia frigida sp. nov. L. I. Sarv, Eesti NSV Tead. Akad. Geol. Inst, uurimused , 4, 79, pi. 12, figs. 7, 8.
1976 Kiesowia frigida Sarv; R. E. L. Schallreuter, Palaeontographica (A), 153 (4/6), 164, 176 - 178, pi. 34 (1), figs. 5 - 10,
tab 4 (q.v. for further synonymy).
1976 Kiesowia frigida (Sarv); N. SidaraviCiene, Sov. Geol. 1976 (8), 54, tab. 1 (49).
Holotype: Geological Museum of the Academy of Science of the Estonian SSR, Tartu, no. Os 2201, 9 LV.
Type locality : Rakvere (=Wesenberg), Estonia; approx, lat. 59°21'N, long. 26°22'E. Laagri substage of Keila Stage
(D2a), upper Middle Ordovician.
Figured specimens: Geologisch-Palaontologisches Institut, University of Hamburg (GPIH), nos. 2023a (Schallreuter, op. cit.,
tab. 4, no. 3; 9 RV: PL 6, 76, figs. 1 - 3; PI. 6, 78, fig. 1) and 2023b (loc. cit. no. 5 ; tecnomorphic RV: PL
6, 78, figs. 2 - 3). Both from the village of Klein-Horst, Pomerania, Poland; lat. 54°6'N, long. 15°5'E;
Backsteinkalk erratic boulder no. Ho-2 (Schallreuter, op. cit., 164); Middle Ordovician; coll, by the
author, 1971 .
Diagnosis: Adult female valves 1.05 - 1.45mm long. L4 divided into two nodes. Dolon extends from anterocentral
region to the centro-posteroventral region.
Remarks: Homeokiesowia pernodosa is very similar to H. frigida but seems to be larger and according to Opik (op.
cit.) its two posterior nodes (L4) are fused.
Distribution: Idavere (C3) to Keilo (D2) stages of Estonia and Lithuaniana and corresponding beds of Podolia.
Backsteinkalk erratic boulders of N Germany and of the Isle of Gotland, Baltic Sea (Baltic Backsteinkalk,
types 14B2 and 1B13).
Explanation of Plate 6, 78
Fig. 1 , 9 RV, ext. vent. (GPIH 2023a); figs. 2, 3, tecnomorphic RV (GPIH 2023b, 970 ,um long); fig. 2, ext. vent.; fig. 3, ext. lat.
Scale A (250 pm\ x 79), fig. 1 ; scale B (250 pm ; x 90), figs. 2,3.
Homeokiesowia frigida (4 of 4)
Stereo-Atlas of Ostracod Shells 6, 76
Stereo-Atlas of Ostracod Shells 6, 78
Homeokiesowia frigida (2 of 4)
"T
H-
Stereo-Atlas of Ostracod Shells 6 (16) 79 - 86 (1979) Kiesowia dissecta (1 of 8)
595.336.13 (1 13.313) (486:161.018.57 + 492.71 : 161.008.54) : 551.35 + 552.55
ON KIESOWIA (KIESOWIA) DISSECTA (KRAUSE)
by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)
Genus KIESOWIA Ulrich & Bassler, 1908
Type-species (by original designation): Beyrichia dissecta Krause, 1892
Diagnosis: A medium-sized to large genus of Sigmoopsinae. Outline almost amplete — slightly preplete. Quadrilobate.
S2 strongest sulcus, long and sigmoidal; S3 curved, SI weakest sulcus, sometimes almost obselete. Lobes
relatively flat; LI sometimes has a dorsal, posteriorly directed spine. Male velum as an entire, more or less
distinct, broadly rounded elevation (“Wulst”) covered with spines or granules. Anteriorly and ventrally
the female velum forms a restricted, flange-like, only weakly convex dolon. Histium ridge-like, narrow to
broad, short to long, present only in females. Often no histial (supravelar) antrum. Marginal sculptures
formed by row of spines.
Remarks: Kiesowia does not belong to the Tallinnellinae (cf. Schallreuter, Palaeontographica (A), 153 (4/6), 175,
1976); the occurrence of a rudimentary histium in the type-species indicates an assignment to the
Sigmoopsinae. Further, the lobation of Kiesowia indicates a closer relationship to the Sigmoopsines
Carinobolbina Henningsmoen (Norsk geol. Tidsskr, 31, 205, 1953) and Pseudotallinnella Sarv ( Ecsti NSV
Tead. Akad. Geol. Inst, uurimused 4, 139, 1959) rather than to Sigmoopsis Henningsmoen (204. 1953) or
Severobolbina Schallreuter (Geol. For. Stockh. Fork. 96 (3 = 558), 278, 1974; = Severella Schallreuter,
Ber. geol. Ges. D. D. R. 9 (3), 395, 1964). Carinobolbina and Pseudotallinnella are, therefore, here
considered as subgenera of Kiesowia. The resemblance between Pseudotallinnella and Carinobolbina was, in
Explanation of Plate 6, 80
Figs. 1-4,9 RV (GPIH 2194, 1580 pm long): fig. 1 , ext. lat.; fig. 2, ext. vent.; fig. 3, ext. ant. obi.; fig. 4, ext. post. obi.
Scale A(250/um;x51), figs. 1 , 2; scale B (250 pm; x 49), fig. 3 ; scale C (250 pm; x 41), fig. 4.
1 — U4--
I Hi-
Stereo-Atlas of Ostracod Shells 6, 81 Kiesowia dissecta (3 of 8)
fact, emphasised by Jaanusson (Bull. geol. Instn. Univ. Upsala 43 (6/8), 7, 1967). Sarv (op. cit ., 140)
considered that Pseudotallinnella was a probable descendant of Carinobolbina, and he also pointed out the
similarity between Pseudotallinnella (P. regalis) and Kiesowia. Tire main difference between (K.)
Carinobolbina and (K.) Pseudotallinnella is the same as that between the subgenera Sigmoopsis (S.) and S.
(Sigmoopsoides) Schallreuter (Ber. geol. Ges. D. D. R. 2, 87, 1964): the histium may or may not become
confluent anteriorly with the velum.
Distribution: Middle Ordovician — Silurian of Baltoscandia.
Subgenus KIESOWIA Ulrich & Bassler, 1908
Diagnosis: Kiesowia species with lobes dissolved into broad nodes. Histium and velum join anteriorly. Histium
sometimes only rudimentary or is lacking.
Remarks: In both the subgenera K. (Carinobolbina) Henningsmoen, 1953 and K. Pseudotallinnella Sarv, 1959 the
lobes are not dissolved into single nodes. In K. (Kiesowia) and K. Pseudotallinnella the histium is
confluent with the velum, whereas in K. (Carinobolbina) it is not.
Kiesowia (Kiesowia) dissecta (Krause, 1892)
1892 Beyrichia dissecta n. sp. A. Krause, Z. Deutsch. geol. Ges. 44 (3), 391, 392, 398, 399, pi. 21, fig. 3.
1954 Kiesowia dissecta (Krause); G. Henningsmoen, Norsk geol. Tiddskr. 33 (1/2), 78, 79, 80 (pars), pi. 2, figs. 1 - 3; non
80 (pars), pi. 2, fig. 5 (= ? Hithis leviconvexus Schallreuter, Geologie 16 (5), 621, 1967).
1956 Kiesowia septenaria sp. n. K. Stumbur, Tartu Riikliku Ulik. Toim. 42, 188, 189, pi. 2, fig. 1 (- K. dissecta according to
Sarv, Eesti NSV Tead. Akad. Geol. Inst, uurimused 9, 95, 109, 1962).
1956 Kiesowia dissecta (Krause); K. Stumbur, Ibid., 189.
1960 Kiesowia septenaria Stum ; V. S. Krandijevskij , Eesti NSV Tead. Akad. Geol. Inst, uurimused 5, 175.
1 962 Kiesowia dissecta (Krause); L. I. Sarv, Eesti NS V Tead. Akad. Geol. Inst, uurimused 9, 95, 97, 98, 1 09, 1 1 0, pi. 4, fig. 9.
j-j— h
Explanation of Plate 6, 82
Figs. 1, 2, 6 LV (GPIH 2195, 1490 pm long): fig. 1, ext. lat. ; fig. 2, ext. vent. obl.;fig. 3, fragmentary 9 RV, int. lat., marginal
sculpture in centroventral region (GPIH 2196); fig. 4, incomplete 9 RV, ext. posterovent. obi., detail showing surface
ornamentation (GPIH 2197).
Scale A (250 pm ; x 53), figs. 1 , 2; scale B (100 ^m; x 105), fig. 3; scale C (100 pm ; x 155), fig. 4.
Kiesowia dissecta (4 of 8)
Stereo-Atlas of Ostracod Shells 6, 82
Stereo-Atlas of Ostracod Shells 6, 83
Kiesowia dissecta (5 of 8)
Type locality:
Figured specimens:
Lectotype: (designated by Henningsmoen 1954, 79). Unnumbered specimen, Palaontologisches Museum, Museum
fur Naturkunde, Humboldt University, Berlin; a presumed 6 RV (Krause, op. cit ., pi. 21 , fig. 3).
Upper Ordovician erratic boulder from Miiggelheim, Berlin.
Geologisch-Palaontologisches Institut, University of Hamburg (GPIH), nos. 2194 (9 RV: PI. 6, 80, figs.
1 - 4), 2195 (6 LV: PI. 6, 82, figs. 1, 2), 2196 (fragmentary 9 RV: PI. 6, 82, fig. 3), 2197 (incomplete 9
RV: PI. 6, 82, fig. 4; PI. 6, 86, fig. 6), 2198 (juv. LV: PI. 6, 84, fig. 1), 2199 (6 LV: PI. 6, 84, figs. 2, 3),
2200, (9 RV: PI. 6, 84, fig. 4; PI. 6, 86, figs. 1 - 4) and 2201 (9 RV: PI. 6, 84, fig. 5; PI. 6, 86, fig. 5).
Specimens 2196, 2197 and 2199 are from Ojlemyrflint (Upper Ordovician) erratic boulders nos. Sy 2, Sy
35 and Sy 1 respectively, of the Kaolinsand (Pliocene — Pleistocene), near Braderup, Isle of Sylt
N Frisian Is., N Sea); lat. 54°56'N, long. 8°21'E; coll, by Ulrich von Hacht, 1976. The other specimens
are from Ojlemyrflint (Upper Ordovician) erratic boulder no. G30 from the Isle of Gotland (Baltic Sea),
beach opposite the Isle of Lilia Karlso, lat. 57° 18' N, long. 18° 8' E; coll, by the author, 1976.
Diagnosis: Females 1.43 - 2.30 mm long. Histium rudimentary: a short, anteroventral ridge, not connected with
velum, sometimes absent.
Remarks: Pseudotallinnella regalis sensu Sarv (Eesti NSV Tead. Akad. Geol. Inst, uurimused 4, pi. 21 , fig. 17, 1959)
belongs to K. (Kiesowia) and is possibly a new species. Its lobes are dissolved into nodes, a small velar
flange seems to be present anteriorly and ventrally and above the flange there is a histial ridge which is
longer than in Kiesowia dissecta and which seems to be connected with the velum somewhat below the
anterior cardinal corner.
Explanation of Plate 6, 84
Fig. 1, juv. LV, ext. lat. (GPIH 2198, 1300 pm long); figs. 2, 3, 6 LV (GPIH 2199, 1460 ^m long): fig. 2, ext. lat ; fig. 3, ext.
vent, obi.; fig. 4, 9 RV, ext. post. (GPIH 2200); fig. 5, 9 RV, ext. ant. obi. (GPIH 2201).
Scale A (250 pm\ x 6 1 ), fig. 1 ; scale B(250lum;x^5), figs. 2,3; scale C(250lum;x56), figs. 4,5.
-tt-t-
— ! — ! — i —
Stereo-Atlas of Ostracod Shells 6,85
Kiesowia dissecta (7 of 8)
Remarks: The adult valves of K. dissecta from Norway (Henningsmoen 1954) are about 1.90mm long, Sarv (1962)
(contd) mentions an adult length of 2.15mm for Estonian material and the lectotype is 2.20 or 2.30mm long
(Krause 1892; Helmdach 1977). The specimens figured herein possibly represent a new, smaller subspecies.
Kiesowia dissecta is similar to, and can occur together with Hithis leviconvexus Schallreuter,
1967. H. leviconvexus is smaller, it lacks a histium in females (also lacking in some females of K. dissecta:
PI. 6, 86, fig. 5) and its dolon terminates posteriorly below L3 whereas in K. dissecta it reaches below L4.
The inner antral fence (marginal sculpture) in H. leviconvexus is formed by a broad flange and in
K. dissecta by a row of spines. The nodes in H. leviconvexus are more typically node-like (smaller, higher
and more rounded) but in K. dissecta they are more lobe-like (broad and flattened). The nodes in front of
the preadduct orial node of K. dissecta are mostly more or less fused, not distinct as in H. leviconvexus.
According to Henningsmoen (1954, 80) Beyrichia mamillosa Krause, 1892 agrees rather well with larval
forms of K. dissecta and may be synonymous. His figured juvenile of K. dissecta (Henningsmoen, pi. 2,
fig. 5) is, in my opinion, possibly conspecific with Hithis leviconvexus (see above), which resembles the
holotype of B. mamillosa Krause, 1892 (op. cit.. pi. 22, fig. 14) more than K. dissecta. B. mamillosa is
more probably a senior synonym of H. leviconvexus.
Distribution: Porkuni Stage (F2), Ordovician and Tamsalu Stage (G2), Silurian of Estonia. Stages 5a, 5b, Oslo region,
Norway. Erratic boulders of Germany; Ojlemyrflint (Upper Ordovician) erratic boulders of the Isle of
Gotland (Baltic Sea) and of the Kaolinsand (Pliocene — Pleistocene) of the Isle of Sylt (N Frisian Is.,
N Sea).
Explanation of Plate 6, 86
Figs. 1-4,9 RV (GPIH 2200, 1430 pm long): fig. 1 , ext. lat.; fig. 2, ext. vent.; fig. 3, ext. vent., detail of velum; fig. 4, ext. lat.,
anterodorsal spine; fig. 5, 9 RV without histium, ext. lat. (GPIH 2201, 1460 pm long); fig. 6, incomplete 9 RV, ext. post. obi.
(GPIH 2197, 1060 broad).
Scale A (250 pm; x 54), figs. 1,5; scale B (250 pm; x 38), figs. 2,6; scale C (100 ^m: x 200), fig. 3; scale D (100 pm ; x 170),
fig. 4.
Stereo-Atlas of Ostracod Shells 6, 86
Kiesowia dissecta (8 of 8)
Kiesowia dissecta (6 of 8)
Stereo-Atlas of Ostracod Shells 6, 84
Stereo-Atlas of Ostracod Shells 6 (17) 87 - 90 (1979) Pyxion posterobicarinatum (1 of 4)
595.336.22 (1 13. 3 13) (492. 71: 161. 008. 54) : 551.35 + 552.55
ONPYXION POSTEROBICARINATUM SCHALLREUTER sp. nov.
by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)
Pyxion posterobicarinatum sp. nov.
Holotype: Geologisch-Palaontologisches Institut, University of Hamburg, no. 2202, RV.
Type locality: Middle Ordovician Hornstein erratic boulder no. Sy 52 of the Kaolinsand (Pliocene — Pleistocene), near
Braderup, Isle of Sylt (N Frisian Is, N Sea), Germany; lat. 54° 56' N, long. 8° 21 ' E.
Derivation of name: With reference to the two carinae in the posterior part of the valve.
Figured specimens: Geologisch-Palaontologisches Institut, University of Hamburg (GPIH), no. 2202 (holotype, RV: PI. 6,
88, figs. 1 - 4), 2203 (LV: PI. 6, 90, figs. 1,2), 2204 (juv. RV: pi. 6, 88, fig. 5; PI. 6, 90, fig. 3), 2205
(RV: PI. 6, 90, fig. 4) and 2206 (LV: pi. 6, 90, fig. 5). All the figured specimens are from Hornstein
erratic boulder no. Sy 52 (see type locality for details); coll, by Ulrich von Hacht, 1978.
Explanation of Plate 6. 88
I-4-.
I — (- +
Figs. 1 - 4, RV (holotype, GPIH 2202, 560 pm long); fig. 1, ext. lat.; fig. 2, ext. vent, obi.; fig. 3, ext. dors, obi.; fig. 4, ext.
vent; fig. 5, juv. RV, ext. vent. (GPIH 2204, 482 pm long).
Scale A (1 00 ^m; x 1 25), figs. 1 - 3 ; scale B (100 pm; x 115), figs. 4,5.
4-f -
-H-r
i i
44—
i i
i i
• i.4..
Stereo-Atlas of Ostracod Shells 6, 89
Pyxion posterobicarinatum (3 of 4)
Diagnosis: Length of valves up to 0.59mm. Length/height ratio without the sculptures dorsal of hinge-line is high
(1.45 to 1.51); with dorsal sculptures, is very high (<1.45). Outline postplete but with forward swing
because anterior cardinal corner considerably larger than posterior cardinal corner; dorsum epicline. Short,
faint sulcal depression dorsal, slightly in front of mid length; two faint more or less confluent nodes
anteroventrally in front of the depression. Anteriorly the indistinct, broadly elevated, ventral carina
gradually merges into the lateral surface, posteroventrally it is more ridge-like, converging with the free
margin in dorsal direction and lacking in the posterodorsal region. Marginal surface confluent with ventral
elevation is convex and in lateral view overhangs the free border; marginal surface confluent with postero-
ventral ridge is concave and does not protrude over the free margin. A second, rounded carinal ridge
occurs posterodorsally. Surface appears to be smooth.
Remarks: The contact conditions of the carapace of Pyxion are here described for the first time (cf. Jaanusson, Bull.
geol. Instn. Univ. Upsala 37 (3/4) 427, 1957). Internal views of P. posterobicarinatum (PI. 6, 90, figs. 4, 5)
show that the left valve overlaps the right valve at the contact margin: the left valve bears an outer list and
an entire, uninterrupted, inner semi-groove; conditions are reversed in the right valve.
The type-species of Pyxion, P. carinatum (Hadding, 1913), possesses an anterodorsal oblong
knob, its crest forming the highest point of the valve. Behind the adductor scar it has a broad, low
inflation extending posteriorly almost to the carinal ridge (Jaanusson, op. cit ., 428).
•1 — 1-4
Distribution: Lower Upper Viruan Hornstein erratic boulders of the Kaolinsand (Pliocene — Pleistocene) near Braderup,
Isle of Sylt (N Frisian Is, N Sea), Germany.
Explanation of Plate 6, 90
Figs. 1,2, LV (GPIH 2203, 585 pm long): fig. 1 , ext. lat.; fig. 2, ext. vent, obi.; fig. 3, juv. RV, ext. lat. (GPIH 2204); fig. 4, RV,
int. lat. (GPIH 2204); fig. 4, R V, int. lat. (GPIH 2205 ,564 pm long); fig. 5 , LV, int. lat. (GPIH 2206, 572 pm long).
Scale A ( 100 pm; x 110), figs. 1,3; scale B (250 ^m; x 70), fig. 2; scale C (250 ,um; x 90), figs. 4, 5.
44-
4-44 +•*.;
Stereo-Atlas of Ostracod Shells 6 ( 1 8) 9 1 -98 (1979) Argenticytheretta punctata (1 of 8)
595.337.14 (1 18.15) (83:164.072.5 3) : 55 1.35 1
ON ARGENTICYTHERETTA (ARGENTICYTHERETTA) PUNCTATA
ROSE sp. nov.
by John F. Rose
(BP Research Centre , Sunbury-on-Thames, England)
Argenticytheretta (Argenticytheretta) punctata sp. nov.
Holotype: University of Hull coll. no. HU220.T20, d RV.
Type locality : Laguna Blanca No. 1 well between 1527 and 15285m, N of Peninsula Brunswick, Magallanes Province,
Chile; approx, lat. 52° 16' S, long 71° 07' W. Grey siltstone, with an abundant, largely foraminiferal
fauna, suggesting a cool water, inner sub-littoral environment; Loreto Formation, uppemost Oligocene,
Miradorian Stage.
Derivation of name: A reference to the punctate ornamentation.
Explanation of Plate 6, 92
Fig. 1 , 9 car., ext. rt. lat. (paratype, 700 pm long); fig. 2,9 car., ext. It. lat. (HU.220.T.2 1 .1 , 730 pm long).
Scale A (250 ^m; x 100), figs. 1,2.
4-tt
44
4-f
44
) H-t-
■44— 4-
Stereo-Atlas of Ostracod Shells 6, 93
Argenticytheretta punctata (3 of 8)
Figured specimens: University of Hull coll. nos. HU.220.T.21.1 (9 car., PL 6, 92, fig. 2), HU.217.T.10 (d car., PL 6, 94, fig. 1),
HU.220.T.21.2 (d car., PL 6, 94, fig. 2), HU217.T.11 (9 car., PL 6, 96, fig. 1), HU.220.20 (d RV, pi. 6,
98, figs. 1 - 3), Paratype, to be lodged with Empressa Nacional del Petroleo at Puncta Arenas, Chile (9 car.
PL 6, 92, fig. 1 and PL 6, 96, fig. 2). The figured specimens were obtained from three samples.
HU.220.T 21 .1 ., HU.220.T.21.2 and HU.217.T.1 1 are from Ciaike No. 1 well (approx, lat. 52° 9' S, long.
70° 3' W) between 1648 and 1651m, Late Oligocene, Miradorian stage. HU.217.T.10 is from Ciaike No. 1
well between 1666 and 1669m, Late Oligocene, Miradorian stage. HU.220.T.20 and the paratype are from
the type locality and level.
Diagnosis: A species with a punctate ornamentation, in which the punctation becomes strongest in the central and
centrodorsal portions of the valves. Fine longitudinal ribs are developed dorsally in both valves and ventro-
laterally in the right valve.
I
Explanation of Plate 6, 94
Fig. 1 , d car., ext. rt. lat. (HU.217.T.10, 750 pm long); fig. 2,d car., ext. It. lat. (HU.220.T.2 1 .2, 750 pm long).
Scale A (250 pm\ x 100), figs. 1,2.
r-T-T
I I I
Stereo-Atlas of Ostracod Shells 6, 92
Argenticytheretta punctata ( 2 of 8)
— M —
Stereo-Atlas of Ostracod Shells 6, 95
Text-fig. 1. Internal view of holotype, HU.220.T.20., x 130
(approx.) showing marginal pore canals.
Argenticytheretta punctata (5 of 8)
Explanation of Plate 6, 96
Fig. 1 , 9 car., ext. dors. (HU.217.T.1 1, 700 long); fig. 2, 9 car., ant. dors. obi. (paratype, 700 //m long).
Scale A (250 pm; x 100), figs. 1,2.
i
i
-4—
-44-t-
Stereo-Atlas of Ostracod Shells 6, 97
Argenticytheretta punctata (7 of 8)
Remarks: A. (A.) punctata resembles the type species, A. (A.) miocenica Rossi de Garcia, 1969 and Morphotype A
of A. (A.) fuegoensis Rose,1975 ( Stereo-Atlas of Ostracod Shells 2, 199 - 202, 1975) but is smaller in size,
88 being only 0.75mm long compared with 88 of A. (A.) fuegoensis which are 0.87mm. A. (A.) fuegoensis
and A. (A.) miocenica also lack any longitudinal ribs.
Like some other species of Argenticytheretta from Southern Chile, [A. (A.) patagoniensis Rose,
1975, A (A.) gonzalezi Rose, 1975 and A. (A.) riescoensis Rose, 1975 (Stereo-Atlas of Ostracod Shells 2,
195 - 198, 1975)]>1. (A .) punctata has a posterior snap-knob (PI. 6, 98, fig. 3 ). There is also a possibility
that this species possesses a ventral snap-knob in the form of a multi-dentate bar below the selvage, as is
the case with many other yet undescribed members of the genus, but on the only single valve found of
this species, this structure is obscured by adherent sediment.
Distribution: The thirteen specimens found come from four localities (text -fig. 2), Laguna Blanca No. 1 well in two
samples between 1518 and 1528.5m, Ciaike No. 1 well in six samples between 1531 and 1669m,Rio del.
Oro No. 1 well in one sample from 978 to 981m, and a surface sample, F.304 from the north coast of
Bahia Inutil (approx, lat. 53° 25 'S, long. 69° 35'W). All are of Late Oligocene age (Miradorian Stage) and
are from deposits with a probable inner sub-littoral origin.
The genus is very widespread in the southern half of South America and makes up a significant
part of the Tertiary ostracod fauna.
Explanation of Plate 6, 98
Figs. 1 - 3, ( holotype , HU.220.T.20, 750 pm long)-, fig. 1 , 8 int. rt. lat.; fig. 2, int. muse, sc.; fig. 3, post, snap-knob.
Scale A (250 pm ; x 100), fig. 1 ; scale B (50 pm; x 400), fig. 2 ; scale C (50 ^m; x 300), fig. 3.
Stereo-Atlas of Ostracod Shells 6, 96
Stereo-Atlas of Ostracod Shells 6, 98
Argenticytheretta punctata (8 of 8)
Argenticytheretta punctata (6 of 8)
+ -f-r
Stereo-Atlas of Ostracod Shells 6 (19) 99 - 106 (1979) Ghardaglaia ambigua (1 of 8)
595.337.12 (1 19.9) (267.161.081.06) : 5 5 1 .35 + 55 1 .3 14 (26.03) : 55 2.51
ON GHARDAGLAIA AMBIGUA NEALE sp. nov.
by John W. Neale
( University of Hull, England)
Ghardaglaia ambigua sp. nov.
University of Hull coll. nos. HU.261 .R.3a, b, 6 car. (a = valves, b = soft parts).
Gonalabba, Yala National Park, Sri Lanka, lat. 6° 20' 58" N, long. 81° 30' 0" E. Recent, in very saline
conditions on a sand bottom.
Latin, meaning ambiguous, referring to the mixture of characters of both Ghardaglaia and Aglaiella.
University of Hull coll. nos. HU.261 ,R.3a (<J car; RV: PL 6, 100, fig. 1 ;LV: PL 6, 102, fig. 1), HU.261 R.3b
(<3 car; Zenker’s organ: PL 6, 106, fig. 1; Hemipene: PL 6, 106, fig. 3), HU.261 ,R. 2a (9 car; RV: PL 6,
100, fig. 2; LV: PL 6, 102, fig. 2), HU.261 .R.la (9 car; RV: PL 6, 104, fig. 1 ; LV: PI. 6, 104, fig. 2, PL 6,
106, fig. 2), HU.261.R.lb (9 furcal attachment: text-fig. 1), HU.261.R.4b (9 limb 5: text-fig. 1),
HU.261.R.4a (9 car.; RV: text -fig. 1).
Holotype:
Type locality:
Derivation of name:
Figured specimens:
Explanation of Plate 6, 100
Fig. 1 , d RV, ext. lat. (holotype, HU.261 .R. 3a, 7 1 2 pm long); fig. 2, 9 RV, ext. lat. (paratype, HU.261 .R. 2a, 7 1 2 pm long).
Scale A (100 ,um; x 132), figs. 1 , 2.
4-F-
4-r-
44-
■44-*-
Stereo-Atlas of Ostracod Shells 6,101
Ghardaglaia ambigua (3 of 8)
Diagnosis: Smooth, elongate, sub-reniform valves; large vestibules with branched marginal pore canals anteriorly, less
so posteriorly. Small, simple pore canals. Fresh material with diffuse sepia-coloured pattern, black eye and
Zenker’s Organ with 7 rosettes.
Remarks: This is referred to Hatmann’s genus Ghardaglaia (hitherto known only from the Red Sea) because of the
characteristic setal ‘brush’ developed at the tip of Limb 7. The antennule, however, has 6 segments
(characteristic of Aglaiella ) not 5 (Ghardaglaia) but the proportions are more in accordance with a division
of segment 2 of the Ghardaglaia type species (Hartmann, Kieler Meeresforschungen 20,1964) than the
proportions seen in the type of Aglaiella as figured by Daday 1910 (Sber. ost. Akad. Mss., 119).
Distribution: G. ambigua is known from four collections made in the Yala National Park, Sri Lanka on the S E coast
of the island viz. Gonalabba, Yala 1 , Yala 2 and Mahisilawa (text -fig. 3). The first three yielded only this
species. The collection at Mahasilawa yielded also Indiacypris luxata (Brady 1886 ), Strandesia elongata
Hartmann 1964 and ? Cytheridea pusilla Brady 1886 which suggests a fauna of mixed salinity tolerances.
Salinity at Gonalabba was high (44 - 72 %o) with pH 7.5 . At Yala 1 salinity was 15 - 30 %o with pH 7.5 -8.5;
in both cases the bottom was sandy.
NJi. Gonalabba is also occasionally spelled Gonalabbe and Mahasilawa — Mahaseelawa.
I am grateful to John Benzie and the Aberdeen University Expedition of 1978 who kindly
provided the material on which this paper is based.
Explanation of Plate 6, 1 02
Fig 1 , 6 LV, ext. lat . (holotype, HU.261 .R.3a, 7 1 2 pm long); fig. 2, 9 LV, ext . lat. (paratype, HU.261 .R.2a, 120 pm long).
Scale A ( 1 00 pm ; x 132), figs. 1,2.
Ghardaglaia ambigua (4 of 8)
Stereo-Atlas of Ostracod Shells 6, 102
Stereo-Atlas of Ostracod Shells 6, 100
Ghardaglaia ambigua (2 of 8)
-J — I- -} —
Stereo-Atlas of Osfracod Shells 6, 103
Ghardaglaia ambigua (5 of 8)
Explanation of Plate 6, 104
Fig. 1 , 9 RV, int. lat. (paratype, HU.261.R.la, 744 long); fig. 2, 9 LV, int. lat. (paratype, HU. 261. R. la, 752 /um long).
Scale A (1 00 /n m; x 1 36), fig. 1 ; scale B ( 1 00 ; x 1 26), fig. 2.
-4-
-H — i-
-4-
-L4-
-44-
Stereo-Atlas of Ostracod Shells 6, 105
Ghardaglaia ambigua (7 of 8)
$ RV External HU. 261. R. 4a
Text -fig. 2.
Text -fig. 3.
Explanation of Plate 6, 106
Fig. 1, 6 Zenker’s Organ (holotype, HU. 261 ,R. 3b); fig. 2, 9 LV, int. muse. sc. (paratype, HU. 261 .R. la); fig. 3, 6 hemipene
(holotype, HU. 261 ,R.3b).
Scale A (5 0 ^m; x 424), fig. 1 ; scale B (25 /tm; x 625), fig. 2; scale C (50 /tm; x 487), fig. 3.
f+-
-h-P- f-
-i-T-r-
i
i
I
Stereo-Atlas of Ostracod Shells 6, 1 06
Ghardaglaia ambigua (8 of 8)
Stereo-Atlas of Ostracod Shells 6, 104
Ghardaglaia ambigua (6 of 8)
-1-4-
I I
Stereo -At las of Ostracod Shells 6 (20) 107 - 1 12 (1979)
595.337.14 (1 19.9) (267.35:161.024.54) : 55 1.351
Paijenborchellina venosa (1 of 6)
PA I JEN BO R C H ELLIN A VENOSA GURNEY sp. nov.
by Ann R. Gurney
(British Museum [Natural History] , London)
Paijenborchellina venosa sp. nov.
1971 Paijenborchellina sp. R. H. Bate, Bull. Cent. Rech. Pau, 5 , 245, 248, 250, 252 - 253, pi. 1 , fig. a.
1978 Paijenborchellina sp. S. P. Jain, Bull. Ind. Geol. Assoc., 1 1 (2), 1 28, pi. 5 , figs. FI - 3.
Holotype: Brit. Must. (Nat. Hist.) no. 1979.199, 9 car.
[Paratypes: 1979.200 -208] .
Type locality : Nearshore, shelf, sample 62222, between Abu Dhabi and Sadiyat Islands, Arabian Gulf; lat. 24° 32' N,
long. 54° 27' E (text -fig. 2).
Derivation of name: Referring to vein like ridges on the reticulate carapace.
Explanation of Plate 6, 108
Fig. 1 , d car., ext. rt. lat. (paratype, 1979.202, 603 pm long); fig. 2, d car., ext. It. lat. (paratype, 1979.200, 595 pm long); fig.
3, 9 car., ext. It. lat. (holotype, 1979.199, 629 pm long); fig. 4, 9 car., ext. rt. lat. (holotype, 1979.199, 629 pm long); fig. 5,
9 LV, int. lat. hinge (paratype, 1979.205, 5 1 8 pm long); fig. 6, 9 LV, int. lat. (paratype, 1979.205, 5 18 pm long); fig. 7, 6 car.,
ext. It. lat. (paratype, 1979.201 , 5 10 long); fig. 8, 9 car., tubercle (paratype, 1979.207, 5 18 pm long).
Scale A(150(um;x67), figs. 1,2,3, 4; scale B(75,um;xl23),fig.5; scale C(150Jttm;x81), fig. 6; scale D (1 30 pm\ x 88), fig.
7; scale E(20lum;x373), fig. 8 .
r4-f-
-4-{~
I
-ft-
-44-r
I
rr
-44-+-
Stereo-Atlas of Ostracod Shells 6, 109
Paijenborchellina venosa (3 of 6)
Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1979.199 (holotype, 9 car.; LV: PI. 6, 108, fig. 3; RV: PI. 6, 108, fig. 4),
1979.200 (d car.; LV: PI. 6, 108, fig. 2), 1979.201 (d car.; LV: PI. 6, 108, fig. 7), 1979.202 (d car.; RV: PI.
6, 108, fig. 1), 1979.203 (9 car.; dorsal: PI. 6, 110, fig. 1), 1979.204 (9 car.; vent: PI. 6, 110 fig. 2),
1979.205 (9 car.; RV internal: PI. 6, 1 10, fig. 3), 1979.206 (9 car.; LV internal: PI. 6, 108, fig. 6; hinge:
PI. 6, 108, fig. 5), 1979.207 (9 car.; tubercle: PI. 6, 108, fig. 8), 1979.208 (d car.; marginal pore canals:
text -fig. 1). 1979.199 - 201, 203 - 208 are from the nearshore shelf and 1979.202 from the lagoonal
channel, Abu Dhabi Lagoon, Arabian Gulf. All specimens described here were obtained from samples
collected by Dr. G. Evans, Imperial College of Science, London.
Diagnosis: Paijenborchellina having reticulate ornament with pitting within the reticulae and superimposed vein-like
ridges. Dorsal margin with saddle shaped depression anterodorsally. External muscle scar swelling
sometimes tuberculate (PI. 6,108, fig. 8). Two tubercles may be present in posterior half.
Remarks: As for other species of Paijenborchellina the male dimorph is more elongate than the female. Internally
P. venosa has 13 long straight anterior marginal pore canals (text -fig. 1) similar to P. alata Gurney, 1979,
where 12 have been recorded. P. ijuensis Reyment, 1959, from Lower Eocene, Nigeria (Stockh. Contr.
Geol. 3 (7): 139 - 143) and P. sp. Reyment, 1963, from beach sand, Gabon (Ann. Mag. nat. Hist., ser. 13,
6,271 - 272) are figured showing less than 10 canals.
Distribution: In addition to Abu Dhabi lagoon P. venosa is also known to occur in beach sediments, Mandi, Kutch,
Arabian Sea (Jain 1978).
Explanation of Plate 6,110
Fig. 1,9 car., ext. vent, (paratype, 1979.203 [specimen broken ] ); fig. 2, 9 car., ext. dors, (paratype, 1979.204, 612 pm long);
fig. 3,9 RV, int. lat. (paratype, 1979.205, 5 1 8 pm long).
Scale A(150(um;x98), figs. 1,2; scale B ( 150 //m; x 115), fig. 3.
+ ->-r
Stereo-Atlas of Ostracod Shells 6,111 Paijenborchellina venosa (5 of 6)
Ecology: The reticulate P. venosa inhabits the more turbulent waters of the lagoonal channel and nearshore shelf,
while the smooth, alate P. alata Gurney,1979 inhabits the calmer back waters (text -fig. 2). A specimen of
P. alata, found at the exit of the lagoon channel is considered to have been brought out of the lagoon and
deposited there by tidal currents. The reticulate P. venosa is better adapted to a life in the more turbulent
waters of the Gulf and lagoon channel than the smooth shelled P. alata. The two species do not overlap in
their ecological distribution and both appear to be ideally adapted to life in the conditions where they are
now found. Unfortunately no living specimens have been obtained but both species are considered to have
lived in the areas where they occur with the exception of the single specimen of P. alata found outside the
lagoon.
Corrigendum: While the present paper was in press Dr. A. J. Keij drew my attention to a paper by Griindel (Z. Geol.
Mss., 9, 1295, 1976) in which he erected Gibboborchella for those non sulcate species previously placed
in Paijenborchellina Kuznetsova, 1961 . Unfortunately it was not possible to amend the genus designation
of this paper and a corrigendum has had to be inserted. P. venosa sp. nov. (present paper) and P. alata
Gurney ( Stereo Atlas of Ostracod Shells, 6, 27 - 30, 1979) should now be assigned to Gibboborchella
together with those already assigned to the genus by Griindel.
I
I I I
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i 1 1
i i i
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I > I
Stereo-Atlas of Ostracod Shells 6,112
I
1 i !
! ! I
44-|-
I I >
I I I
• I I
> I I
1 I I
- — ttT
! ! 1
Paijenborchellina venosa (6 of 6)
Text-fig. 1 . Anterior ( a ) and posterior (b ) radial pore canals
6 RV (paratype, 1979.208).
Text-fig. 2. Distribution of P. venosa (•) and P. alata (▼).
Holotype locations ringed.
1
+4
-b-f '
4'
4-|-b
r— r-r
Stereo-Atlas of Ostracod Shells 6 (2 1) 113-116 (1979) Monoceratina scrobiculata (1 of 4)
595.337.14(116.21 + 1 1 6.22 + 1 16.23 1 ) (4: 1 6 1 .007.57 + 003.58 + 002.5 1 +002.52 + 00 1 .49 + 1 62.009.48 + 007.47 + 007.53)
551.351 + 552.52
ON MONOCERATINA SCROBICULATA TRIEBEL & BARTENSTEIN
by Lesley M. Sheppard
(British Museum [ Natural History] , London )
Monoceratina scrobiculata Triebel & Bartenstein, 1938
1938 Monoceratina scrobiculata n. sp. E. Triebel & Bartenstein, Senckenbergiana, 20 (6), 508, pi. 1 figs. 4a, b, pi. 2 fig. 6.
1959 Monoceratina scrobiculata Triebel & Bartenstein; H. J. Oertli, Denkschriften der Schweiz. Naturf. Gesellschaft, 83,
26, pi. 4 figs. 92 - 95.
1960 Monoceratina cf. scrobiculata Triebel & Bartenstein; G. F. Lutze, Geol. Jb., 77, 433, pi. 37 fig. 7.
non 1969 Monoceratina cf. scrobiculata Triebel & Bartenstein; F. Depeche, Bull. Centre Rech. Pau - SNPA, 3 (2), pi. 2 fig. 9.
1970 Monoceratina scrobiculata Triebel & Bartenstein; R. C. Whatley, Bull. Br. Mus. nat. Hist. (Geol.), 19 no. 6, 318, pi.
3, figs. 1 - 7, 9, 10 (q.v. for full synonymy).
Holotype: Senckenberg Museum coll. no. SMF Xe 133a, RV.
Type locality: Kremmeldord, near Bamberg, Baden, S Germany. Dogger alpha, opalinus zone, L. Aalenian, M. Jurassic.
Figured specimens: Senckenberg Museum coll. nos. SMF Xe 133b (LV : PI. 6 1 14, fig. 1),SMF Xe 113c (LV : PI. 6, 1 16, fig. 3).
Brit. Mus. (Nat. Hist.) nos. OS 11705(RV: PI. 6, 1 14, fig. 2; PI. 6, 11 6, fig. 2), OS 11706 (LV: PI. 6, 114,
fig. 3; PI. 6, 116, figs. 4, 5), OS 11707 (LV : PI. 6, 1 16, fig. 1). SMF Xe 113b is from the type locality and
level, SMF Xe 113c is from the type level at Peulendorf, near Bamberg. Baden. S Germany, and
Explanation of Plate 6,114
Fig. 1, LV, ext. lat. (paratype, SMF Xe 113b, 654 p m long); fig. 2, RV, ext. lat. (OS 11705, 646 pm long); fig. 3, LV, ext. lat.
(OS 1 1706, 646 pm long).
Scale A (200 pm\ x 92), fig. 1 ; scale B (200 ^m; x 94), figs. 2,3.
Stereo-Atlas of Ostracod Shells 6, 1 15
Monoceratina scrobiculata (3 of 4)
Figured specimens: OS 11705 - OS 11708 are from Port-en-Bessin, Normandy, France; Lower Fuller’s Earth, Bathonian,
(contd.) Middle Jurassic. The paratypes were kindly lent by Dr H. Malz of the Senckenberg Museum, Frankfurt.
Diagnosis: Spined species of Monoceratina characterised by strong reticulation over entire valve surfaces.
Remarks: This is a very striking ostracod with the surface reticulation comprising large, essentially hexagonal pits.
The medial area is greatly swollen with a deep sulcus forming a distinctive crescentic lobe, the postero-
ventral part bearing the spine which in well preserved specimens is as long as xh valve length and usually
directed posteroventrally. The paratype figured here shows a more even ornament around the valve
edges than the French material but this is considered merely a function of preservation. The hinge is
lophodont with a long median ridge in the left valve fitting into a groove in the right valve; terminal
elements weak.
Distribution: M. scrobiculata ranges from Lower to Upper Jurassic in NW Europe. It has been recorded in the Lower
Bathonian of Normany (listed previously by Depeche, Revue Micropaleont., 15, 4, 214, 1973 from the
Port-en-Bessin section, and by Bizon, Revue Inst, fr., Petrole, 13, 1 , 23, 1958 from Villers-sur-mer); from
the Middle Callovian ( coronatum Zone) to the base of the Upper Oxfordian (plicatilis Zone) at various
localities in England and Scotland (see Whatley, op. cit.); and from the Upper Callovian to Lower
Oxfordian of N France (J. Guyader, Le Jurassique superieur de la baie de la Seine - Etude stratigraphique
et micropaleontologique, unpub. PhD thesis, University of Paris, p. 197, 1968). In NW Germany it has
been recorded from the Upper Bathonian ( aspidoides Zone) to Lower Oxfordian ( miriae Zone) and in
S Germany from the Upper Lias and Lower Aalenian; also from the Upper Oxfordian of Switzerland
(Oertli, op. cit.).
Explanation of Plate 6, 1 16
Fig. 1, LV dors. (OS 11707, 663 pm long); fig. 2, RV dors. (OS 11705, 646 p m long); fig. 3, RV, int. lat. (paratype, SMF Xe
1 13c, 646 pm long); fig. 4, LV, int. lat. (OS 11706, 646 pm long); fig. 5, ornament (OS 1 1706).
Scale A (200 pm-,xl6), fig. 1 ; scale B (200 pm; x 78) figs. 2,3, 4; scale C (50 pm; x 400), fig. 5.
Stereo-Atlas of Ostracod Shells 6, 1 14
Monoceratina scrobiculata (2 of 4)
Stereo-Atlas of Ostracod Shells 6,116
Monoceratina scrobiculata (4 of 4)
I I I
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: !
Stereo- Atlas of Ostracod Shells 6 (22) 1 17 - 124 (1979) Heterocythereis albomaculata { 1 of 8)
i i i 595.337.14 (1 19 + 1 1 9.9) (420: 1 61 .002.55 + 002.50 + 005.5 1 + 45:162.012.44) 551.351
ON HETEROCYTHEREIS ALBOMACULATA (BAIRD)
by John Athersuch & John E. Whittaker
(B.P. Research Centre, Sunbury, and British Museum [ Natural History ] , London)
Genus Heterocythereis Elofson, 1941
Type-species (by original designation): Cy there albomaculata Baird, 1838.
Diagnosis: Valves weakly ornamented; sexual dimorphism pronounced. Hinge amphidont with very strong anterior
tooth; posterior tooth crenulate, median elements smooth. Muscle scars with three frontal scars; only
second from top adductor muscle-scar subdivided; fulcral point present.
Heterocythereis albomaculata (Baird, 1838).
Cy there alba sp. nov. W. Baird, Hist. Berwicksh. Nat. Club, 1, 98, pi. 3, fig. 6. (? juv.) ( nomen dubium).
Cythere albo-maculata sp. nov. W. Baird, Mag. Zool. Bot., 2, 142, pi. 5, fig. 23.
Cythereis (Heterocythereis (albomaculata (Baird); O. Elofson, Zool. Bidr, Upps., 19, 292.
Eucythereis albomaculata (Baird); A.P.C. de Vos, Arch. Zool. exp. gen., 45, 28, pi. 11, figs. 3a - c; pi. 12, figs.
2a - g.
Heterocythereis albomaculata (Baird); C. W. Wagner, Sur les Ostracodes du Quaternaire recent des Pays-Bas et leur
utilization dans Vetude geologique de depots holocenes, Mouton & Co., The Hague, 57, pi. 24, figs. 1-7.
Heterocythereis albomaculata (Baird); G. Bonaduce, G. Ciampo & M. Masoli, Pubbl. Staz. zool. Napoli, 40, 46, pis.
21, figs. 8 - 1 1. (= H. voraginosa Athersuch, 1979).
Explanation of Plate 6,118
! | j Fig. 1 , 9 car., ext. It. lat. (Neotype, 1979.210, 850 /um long); fig. 2, 9 RV, ext. lat. (1979.21 1, 860 pm long); fig. 3, d car., ext.
It. lat. (1979.212, 980 pm long),
j 1! Scale A (250 pm\ x 64), figs. 1 - 3.
-4-jt
71835
1838
1941
1957
1957
non 1975
■t— j-4..
■h— Hi-
St ere o- Atlas of Ostracod Shells 6, 1 19
Neotype: (here designated). Brit. Mus. (Nat. Hist.) no. 1979210; 9 car.
Heterocythereis albomaculata (3 of 8)
Type locality: Berwick-on-Tweed, N E England, approx, lat. 55° 46 'N, long. 02° 00' W; Recent.
Diagnosis: Carapace surface finely punctate with conspicuous normal pores; lineate and often faintly reticulate
ventrally; gently convex in dorsal view, greatest width in middle. Shape of opaque areas and copulatory
appendages diagnostic.
Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1979.210 neotype (9 car.: PI. 6, 1 18, fig. 1), 1979.211 (9 RV: PI. 6, 1 18, fig.
2), 1979.212 (dear.: PI. 6, 118, fig. 3; PI. 6, 122, fig. 2), 1979.213 (9 car.: PI. 6, 120, fig. 1), 1979.214 (d
car.: PI. 6, 120, fig. 2), 1979.215 (9 car.: PI. 6, 120, fig. 3), 1979.216 (d LV: PI. 6, 120, fig. 4;P1. 6, 124,
figs. 1, 3, 4), 1979.217 (juv.-4: PI. 6, 122, fig. 1), 1979.218(9 car.: PI. 6, 122, fig. 3), 1979.219 (d LV:
PI. 6, 122, fig. 4; PI. 6, 124, figs. 2, 5), 1979.220 (d soft-parts: Text-fig. la-d), 1979.275 (9 RV: Text-fig.
2). 1979.210 from algae in an intertidal rock pool at the type locality, collected by J. Athersuch and
J. E. Whittaker during March 1979. 1979.211 - 214, 216, 218, 219, 275, collected by K. Trier and
J. Athersuch from intertidal rock pools at Abereiddy, S W Wales (approx, lat. 51° 56' N, long. 05° 13' W),
during September 1977. 1979.217 from Osmington Mills, Weymouth Bay, S England (approx, lat.
50° 38' N, long. 02° 23' W), collected from green algae in an intertidal rock pool during August 1969 by
J. E. Whittaker. 1979.220 from Shell Bay, Poole, S England (approx, lat. 50° 17 'N, long. 02° 05 ' W),
collected August 1975 by C. P. Palmer, from beach sand. 1979.215 from Rimini, N E Italy (approx, lat.
44° 03' N, long. 12° 34' E), collected by G. Ruggieri, from beach sand. All specimens were collected from
normal marine salinities. All had soft-parts intact.
Explanation of Plate 6, 120
Fig. 1, 9 car., ext. dors. (1979.213, 860 pm long); fig. 2, d car., ext. vent. (1979.214, 980 pm long); fig. 3, 9 car., ext. It. lat.
( 1 979.2 1 5, 780 pm long); fig. 4, d LV, int. lat. ( 1979.2 16, 1 000 pm long).
Scale A (250 pm-, x 64), figs. 1 - 4.
E-l-f
I
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4
• i !
! 1 1
■tl-f-
i
i
i
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i
i
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i
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i
i
i
i
i
Stereo-Atlas of Ostracod Shells 6, 120
Heterocythereis albomaculata (4 of 8)
Stereo-Atlas of Ostracod Shells 6,118
Heterocythereis albomaculata (2 of 8)
-tf-
-!--M —
Stereo-Atlas of Ostracod Shells 6, 121
Heterocythereis albomaculata (5 of 8)
Explanation of Plate 6, 122
Fig. 1, juv. -4 car., ext. It. lat. (1979.217, 310 /un long); fig. 2, 6 car., pores in oral region (1978.212); fig. 3, 9 car., pores in
posterodorsal region (1978.218); fig. 4, 6 LV, int. lat., detail of post. vent, region of RV showing clusters of marginal setae.
(1978.219).
Scale A (250 «m; x 64), fig. 1 ; scale B ( 10 /um; x 750), fig. 2 ; scale C ( 10 pm; x 1 ,000), fig. 3; scale D(25pm;x 500), fig. 4.
r4-F-
-H-
-4-44-
i
i
-L~l—
-44-
i
-(-*■-
-t+-r
Stereo- Atlas of Ostracod Shells 6, 123
Heterocythereis albomaculata (7 of 8)
Remarks:
The types of both Cy there alba and C. albomaculata could not be found in the remaining part of the W.
Baird collection in the Brit. Mus. (Nat. Hist.). Brady {Trans. Linn. Soc. Lond. 26, 403, 1868) suggested
that Baird had applied the name C. alba to a juvenile of C. albomaculata. If C. alba and C. albomaculata
could be shown to be conspecific, the former name would have priority. However, since small instars of
many species of the Hemicytheridae look very similar, and because the name C. alba does not appear to
have been used since 1850 (Baird, The Natural History of the British Entomostraca, Ray Soc., London,
170) we prefer to regard the name as a. nomen dubium. A small instar of Heterocythereis albomaculata is
shown in PI. 6, 122 , fig. 1 for comparison with Baird’s 1838 illustration of C. alba (pi. 3, fig. 6).
Distribution: Recent; a widely distributed phytal species in the littoral and sublittoral waters of N W Europe as far
north as N Norway (Brit. Mus. [Nat. Hist.]
(Wagner 1957, op. cit.).
coll.); also occurs in the Mediterranean. Quaternary; Holland
250 pm
Text-fig. 2. 9 RV, transmitted light to show
opaque areas and radial pore canals (scale
= 250 pm; x 1979.275).
:4:,-
Explanation of Plate 6, 124
Fig. 1, 6 LV, int. lat. muse. sc. (1979.216); figs. 2, 5, 6 RV, ant. and post, hinge (1979.219); figs. 3, 4, 6 LV, post, and ant.
hinge (1979.216).
Scale A (25 pm ; x 570), fig. 1 ; scale B (50 pm; x 260), figs. 2-5.
-44-
-rt-4-
-4-44-
-44-4-
I I I
Stereo-Atlas of Ostracod Shells 6, 122
Heterocythereis albomaculata (6 of 8)
Stereo-Atlas of Ostracod Shells 6, 1 24
Heterocythereis albomaculata (8 of 8)
t-fl +-f-r
Stereo-Atlas of Ostracod Shells 6 (23) 125 - 132 (1979) Heterocythereis voraginosa (1 of 8)
595.337.14 (1 19.9) (262.2:161.033.35 + 161.034.35 + 262.3) : 551.351 + 552.51
ON HETEROCYTHEREIS VORAGINOSA ATHERSUCH sp. nov.
by John Athersuch
(B.P. Research Centre, Sunbury-on-Thames, England)
Heterocythereis voraginosa sp. nov.
1975 Heterocythereis albomaculata (Baird); G. Bonaduce, G. Ciampo & M. Masoli, Pubbl. Staz, zool. Napoli, 40, 46, pi. 21,
figs. 8-11. (non Cy there albomaculata Baird 1838).
Holotype: Brit. Mus. (Nat. Hist.) no. 1979.224, 9 car.
[Paratypes: nos. 1976.943; 1979.221 - 223, 225 - 229] .
Type locality: Dhavlos Bay, Cyprus, approx, lat. 35° 25 'N, long. 33° 55' E; beach sand; Recent.
Derivation of name: voraginosus, -a, -um, Latin; covered with pits — alluding to the many obvious sunken sieve-pores.
Diagnosis: Carapace surface with faint reticulum; soli foveolate and perforated by large, sunken sieve-pores; angular
ventrolaterally; truncate anteriorly and posteriorly.
-4-4-
i
Explanation of Plate 6, 126
Fig. 1 , 9 LV, ext. lat. (1979.221, 750 pm long); fig. 2, d LV, ext. lat. (1979.222, 860 pm long); fig. 3, d RV, ext. lat. (1979.223,
850 pm long).
Scale A (250 pm\ x 70), figs. 1-3.
i-i-r-
-44-
-44- 1--
Stereo-Atlas of Ostracod Shells 6, 127
Heterocythereis voraginosa (3 of 8)
Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1979.221 (9 LV: PI. 6, 126, fig. 1); 1979.222 (d LV: PI. 6, 126, fig. 2; PI. 6,
130, fig. 1); 1979.223 (d RV: PI. 6, 126, fig. 3); 1979.224 Holotype (9 car.: PI. 6, 128, fig. 1); 1979.225
(d car.: PI. 6, 128, fig. 2); 1976.943 (d car.: PL 6, 128, fig. 3); 1979.226 (d RV: PI. 6, 130, fig. 2; PI. 6,
132, figs. 2, 5); 1979.227 (d LV: PI. 6, 130, fig. 3; PL 6, 132, figs. 1, 3, 4); 1979.228 (d copulatory
appendage: Text-fig. 1); 1979.229 (d RV: Text-fig. 2). 1976.943 from Yialousa, Cyprus, approx, lat.
35° 31' N, long. 34° 12' E, collected by J. Athersuch during November, 1973; depth 11m, water temp.
21°C, salinity 39%o, pH 8.1, living in sand and algal debris. 1979.221 - 229 from beach sand at the type
locality, collected by P.C. Sylvester-Bradley during Summer, 1972 ; soft-parts intact.
Remarks: Differs from H. albomaculata (Baird)(see Stereo-Atlas of Ostracod Shells 6, 117 - 124, 1979) in having
larger sieve-pores and in being finely reticulate and generally more angular in outline. Opaque area and
male copulatory appendages distinctive.
Distribution: The only previous record is from the N Adriatic Sea (Bonaduce et al., 1975 op. cit.).
Explanation of Plate 6, 1 28
Fig. 1 , 9 car., ext. dors, (holotype, 1979,224, 760 pm long); fig. 2, d car., ext. vent. (1979.225, 860 pm long); fig. 3, d car., ext.
rt. lat. (1976.943, 840 pm long).
Scale A (250 ^m; x 70), figs 1 - 3.
i-r
Stereo-Atlas of Ostracod Shells 6, 126 Heterocythereis voraginosa (2 of 8)
Stereo-Atlas of Ostracod Shells 6, 1 28
Heterocythereis voraginosa (4 of 8)
Hi
4--U4-
Explanation of Plate 6, 130
Fig. 1, d LV, detail of post. vent, region (1979.222); fig. 2, d RV, int. lat. (1979.226, 840 /r m long); fig. 3, d LV, int. lat.
(1979.227, 840 /jm long).
Scale A (25 /um\ x 665), fig. 1 , scale B (250 /tra; x 70), figs. 2, 3.
r4-f
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Stereo-Atlas of Ostracod Shells 6, 131
Heterocythereis voraginosa (7 of 8)
Text-fig. 2. d RV, transmitted light to show opaque area and radial pore canals (scale = 250 ,um; 1979.229).
Explanation of Plate 6, 1 32
Fig. 1 , d LV, int. lat. muse. sc. ( 1979.227); figs. 2, 5, d RV, ant. and post, hinge ( 1979.226); figs. 3, 4, d LV, post, and ant. hinge
(1979.227).
Scale A (25 fxm,x 630), fig. 1 ; scale B(50lum;x 230), figs. 2-5.
Heterocythereis voraginosa (8 of 8)
Stereo-Atlas of Ostracod Shells 6, 132
Heterocythereis voraginosa (6 of 8)
Stereo-Atlas of Ostracod Shells 6, 130
Stereo-Atlas of Ostracod Shells 6 (24) 133 - 140 (1979) Acanthocythereis hystrix (1 of 8)
595.337.14 (1 18.21 - 1 1 9.9) (4: 1 62.0 1 6.49 + 032.35 + 033.34 + 45 + 61 + 262) 551.351 (26.03:24.08.69)
ON ACANTHOCYTHEREIS HYSTRIX (REUSS)
by John Athersuch
(B. P. Research Centre, Sunbury-on-Thames, England)
Acanthocythereis hystrix (Reuss, 1849)
1795 Mytuli scabri vel asperi; A. Soldani {pars), Testaceographiae ac Zoophytographiae parvae et micro scopicae, 1 (3), 218,
pi. 148, fig. Z.
1849 Cypridina hystrix sp. nov. A. E. Reuss, Naturw. Abh. Vienna, 3 (1), 74, pi. 10, figs. 6a - c.
1850 Cythereis senticosa sp. nov. W. Baird, Pro c. zool. Soc. Lond., (18), 256, pi. 18, figs. 16 - 18.
1941 Cythereis? hystrix (Reuss); E. Triebel, Senckenbergiana, 23, pi. 12, fig. 146.
1950 Trachyleberis hystrix (Reuss); G. Ruggieri, Giorn. Geol. ser. 2a, 21, 14 (new combination).
1953 Cythereis hystrix (Reuss); G. Ruggieri, Giorn. Geol. ser. 2, 23, 65, pi. 1, fig. 2.
1962 Trachyleberis (Trachyleberis) hystrix (Reuss);G. Ruggieri, Palaeontogr. ital., 56, 18, pi. 1 1, fig. 21.
1972 Acanthocythereis hystrix (Reuss); W. Sissingh, Utrecht micropaleont. Bull., 6, 92 (new combination).
1975 Acanthocythereis hystrix (Reuss); E. Bremen, The Distribution of Ostracodes in bottom sediments of the Adriatic Sea,
Dissertation, University of Amsterdam, 56, pi. 7, fig. 102.
Lectotype: (here designated). In the Reuss collection, Natural History Museum, Vienna no. 1848. XXXVIII. 23
? 6 RV.
Explanation of Plate 6, 1 34
Fig. 1, 6 LV, ext. lat. (1979.150, 1000 pm long); fig. 2, 6 RV, ext. lat. (1979.151, 951 pm long); fig. 3, 9 LV, ext. lat. (1979.152,
890 pm long).
Scale A (250 pm\ x 63), figs. 1 - 3.
•I H-t-
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H
44-
I I
I I
I I
I I
I I
Stereo-Atlas of Ostracod Shells 6, 135
Acanthocythereis hystrix (3 of 8)
Type locality : Rudelsdorf (Rudoltice), N W Czechoslovakia, approx, lat. 49° 54' N, long. 16° 35' E; Middle Miocene
(Badenian).
Figured specimens: Natural History Museum, Vienna no. 1848.38.23 (lectotype, 6 RV: PI. 6, 136, fig. 1), Brit. Mus. (Nat.
Hist.) nos. 1979.150 (6 LV: PI. 6, 134, fig. 1;P1. 6, 138, fig. 2), 1979.151 (dRV: PI. 6, 134, fig. 2; PI. 6,
138, fig. 1), 1979.152 (9 LV: PI. 6, 134, fig. 3, PI. 6, 138, fig. 3), 1979.153 (9 LV: PI. 6, 136, fig. 2),
1979.154 (9 LV: PI. 6, 136, fig. 3), 1979.155 (9 RV: PI. 6, 136, fig. 4), 1979.156(6 car.; RV: PI. 6,140,
figs. 1 , 2, 4; LV: PI. 6, 140, figs. 3, 5;text-fig. la - d).
Lectotype from the type locality. All other specimens from grab samples collected during 1972
summer cruise of R. R. S. ‘Shackleton’ off the N coast of Cyprus. Samples kindly supplied by Dept, of
Oceanography, University of Liverpool and Dept, of Earth Sciences, University of Leeds. 1979.150 - 155
from approx, lat. 35° 12' N, long. 32° 38' E; water depth 69m. 1979.156 from approx, lat. 34° 42' N,
long. 33° 25' E; water depth 75m.
Diagnosis: Long multifurcate spines borne in consistent pattern conjunctively on surface reticulum and marginally.
Remarks: The list of species in the main part of Reuss’s ostracod collection has been lost for more than fifty years
(pers. comm. Dr. F. Rogl, Natural History Museum, Vienna), and it is not possible, without a thorough
search of the whole collection, to determine if any specimens of Cypridina hystrix are present. However, a
subsidiary collection, received by the Natural History Museum, Vienna from Reuss in 1848, contained a
single complete, but abraded RV of this species. Since this specimen is topotypic and was known to Reuss
at the time of publication of his 1849 paper, it has been selected as the lectotype.
Explanation of Plate 6, 136
Fig. 1, ?d RV, ext lat. (lectotype, 933 pm long); fig. 2, 9 LV, int. lat. (1979.153, 902 pm long); fig. 3, 9 LV, ext. dors.
(1979.154, 902 pm long); fig. 4, 9 RV, ext. dors. (1979.155, 902 pm long).
Scale A (250 pm' x 63), figs. 1 - 4.
T
T'
t “t—r
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I I I
t-t-i — i-H-
Stereo-Atlas of Ostracod Shells 6, 137 Acanthocythereis hystrix (5 of 8)
Remarks: The present species resembles the type species of Acanthocythereis Howe, 1963 (A. araneosa Howe,1963)
(contd.) in having a reticulum bearing conjunctive spines and in having strongly developed eye tubercles and
marginal spines. The double row of ventrolateral spines might indicate affinities to Actinocythereis Puri
1953 which is characterised by lineation of the lateral spines. However, the type species (A. exanthemata
[Ulrich & Bassler, 1904]) lacks a reticulum and has poorly developed marginal spines and small eye
tubercles. Moreover, the spines of Actinocythereis tend to be nodose laterally and denticulate
anteromarginally, unlike those of A. araneosa or A. hystrix. In addition, males of Actinocythereis spp.
typically have distinct assymetry of the valves (see J. E. Hazel, Prof. Pap. U. S. geol. Surv., No. 564, 1967).
On balance, therefore, I prefer to regard the present species as belonging to the genus Acanthocythereis.
Distribution: Recent and sub-Recent: Aegean (Baird (1850); Adriatic (Bremen, op.cit., Bonaduce coll.); Italy (Ruggieri
and Bonaduce colls.); Cyprus (herein); N W. africa (Rosenfeld & Bein , Meteor ForschErgebn. (C), 17:17,
pi. 1, fig. 4, 1978). Miocene, Pliocene and Pleistocene of Italy (Ruggieri 1950, 1962, op. cit., Reuss, op.
cit .) and the Aegean Islands (Sissingh, op. cit.). Miocene of Czechoslovakia (herein) and Austria (Triebel,
op. cit.).
Explanation of Plate 6, 140
Fig. 1, d RV, int. lat. muse. sc. (1979.156, 950 pm long); figs. 2, 4, d RV, term, hinge elements (1979.156); figs. 3, 5, d LV,
term, hinge elements (1979.156).
Scale A (25 pm; x 500), fig. 1 ; scale B (25 pm; x 325), figs. 2-5.
Text-fig. 1. Appendages of male (1979.156). a: 1st antenna; b: 2nd antenna; c: copulatory appendage; d: mandible (all x290).
Explanation of Plate 6, 1 38
Fig. 1, d RV, eye tubercle (1979.151); fig. 2, d LV, tip of spine (1979.152); fig. 3, 9 LV, ext. lat., sieve pore and seta
(1979.150).
Scale A(25pm;x 435, fig. 1 ; scale B(25pm;x755), fig. 2 ; scale C (5 pm; x 2100), fig. 3.
Stereo-Atlas of Ostracod Shells 6, 139
Acanthocythereis hystrix (7 of 8)
•y -
Stereo- Atlas of Ostracod Shells 6, 138
Stereo-Atlas of Ostracod Shells 6, 140
Acanthocythereis hystrix (8 of 8)
Acanthocythereis hystrix (6 of 8)
I I I
1,1
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I l I
I I I
i I I
I i I
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I I I
— -
i i |
Stereo-Atlas of Ostracod Shells 6 (25) 141 - 150 (1979) Loxoconcha ovulata (1 of 10)
595.337.14 (1 19 + 1 19.9) (45:161.014.40 + 262.2:161.033.35 + 262.4:161.025.37 + 262.4 : 161.023.37 + 262.4 : 1 61 .026.39
j ii + 262.1 : 161. 014.40 + 262.1:161.007.43 + 262.3) : 551.351 + 552.51
ON LOXOCONCHA OVULATA (COSTA)
by John Athersuch
(B. P. Research Centre, Sunbury-on-Thames, England)
Loxoconcha ovulata (Costa, ‘1863’)
‘1863’ Cytherina ovulata sp. nov. O. G. Costa , Paleontologia del Regno di Napoli, pt. 3, 181, pi. 16, fig. 7.
1864 Cytherina ovulata Costa; O. G. Costa , Atti Accad. Pontan., 8, 177, pi. 16, fig. 7.
1869 Loxoconcha tumida sp. nov. G. S. Brady, Ann. Mag. nat. Hist., ser. 4, 2, 48, pi. 8, figs. 11, 12. (non L. tumida Chapman,
1902).
1965 Loxoconcha ovulata (Costa);P. Ascoli, Arch. Oceanogr. Limnol., 14 (1), 99.
1 968 Loxoconcha tumida Brady ; M. Masoli, Mem. Mus. Trident. Sci. Nat., 17 ( 1), 55, pi. 3, fig. 34 ; pi. 12, figs. 191 - 193.
1971 Loxoconcha ovulata (Costa); P. J. Barbeito-Gonzalez, Mitt. hamb. Zool.Mus., 67, 307, pi. 32, figs, lb, 2b, 3b, 4b.
1972 Loxoconcha ovulata (Costa); H. Uffenorde, Gottinger Arb. Geol. Palaont., 1 3, 85, pi. 9, fig. 4.
1975 Loxoconcha tumida Brady; E. Bremen, The Distribution of Ostracodes in the Bottom Sediments of the Adriatic Sea,
Dissertation, University of Amsterdam, 66, pi. 9, fig. 130.
1 975 Loxoconcha tumida Brady; G. Bonaduce, G. Ciampo & M. Masoli ,Pubbl. Staz. zool. Napoli, 40 ( 1 ), 1 10, pi. 60, figs. 1-7.
-i— 1-4
Explanation of Plate 6, 142
Fig. 1, $ LV, ext. lat. (Hancock Museum specimen, 720 ^m long); fig. 2, 9 RV, ext. lat. (Hancock Museum specimen, 720 pm
long); fig. 3, 6 LV, ext. lat. (1976.1084, 780 pm long).
Scale A (250 /zm; x 80), figs. 1 - 3.
Stereo-Atlas of Ostracod Shells 6, 143
Loxoconcha ovulata (3 of 10)
4-4--
4—
-rt
44-
Lectotype: (not figured herein) In the Costa collection, Institute of Palaeontology, University of Naples; 9 LV.
Type locality: La Starza, Pozzuoli, Italy, approx, lat. 40° 49' N, long. 14° 07'E; Quaternary.
I
I
I
Figured specimens: Hancock Museum specimen (9 car.; LV : PI. 6, 1 42, fig. 1 ; RV : PI. 6, 142, fig. 2); Brit. Mus. (Nat. Hist.) nos.
1976.1084 (d car.; LV: PI. 6, 142, fig. 3;RV: PI. 6, 144, fig. 3 ; PL 6, 150, fig. 1); 1976.1085 (dLV: PI. 6,
144, fig. 1); 1976.1086 (9 car.; RV: PI. 6, 144,fig. 2; PI. 6, 150, figs. 2,4); 1976.1087 (9 LV: PL 6, 148,
fig. 1); 1976.1088 (9 LV: PI. 6, 148, fig. 3); 1976.1089 (d LV: PL 6, 148, fig. 4); 1976.1090 (d LV: PL 6,
150, figs. 3, 5); OS 11421 (d RV: PL 6, 148, fig. 2). The specimen from G. S. Brady’s collection in the
Hancock Museum is from Piraeus, Greece. 1976.1084, 1086 were living at the time of collection.
1976.1084 collected by J. Athersuch during November 1973 off Famagusta, Cyprus, approx, lat. 35°
08' N, long. 33° 57' E; water depth 27m., salinity 39%o in coarse sand.
Dr. G. Bonaduce collected 1976.1085 - 7, 1089 from the Bay of Naples, Italy and OS 11421
from Costa’s type locality at Pozzuoli, Italy. 1976.1088, 1090 are from silt in Morphou Bay, Cyprus,
collected by members of the Fisheries Dept, of the Ministry of Agriculture and Natural Resources, Nicosia,
Cyprus.
Diagnosis: Tumid. Margins, particularly those of the 9, smoothly rounded and lack a flange anteriorly, d copulatory
appendages of L. rhomboidea (Fischer) type, but with long curved, pointed, distal ‘lappet’.
I
I
I
I
I
I
Explanation of Plate 6, 144
Fig. 1, d LV, ext. lat. (1976.1085, 768 pm long); fig. 2, 9 RV, int. lat. (1976.1086, 683 pm long); fig. 3, d RV, int. lat.
(1976.1084, 780 pm long).
Scale A (250 pm\ x 80), figs. 1 - 3.
44
I
Loxoconcha ovulata (4 of 10)
Stereo-Atlas of Ostracod Shells 6, 144
Stereo-Atlas of Ostracod Shells 6, 142
Loxoconcha ovulata (2 of 10)
-I- -I —
Stereo-Atlas of Ostracod Shells 6, 145
Loxoconcha ovulata (5 of 10)
Text-fig. 1. Appendages, a: 2nd. It. antenna;b: 1st. rt. antenna;c. maxilla;d: mandible.
-r4-f ■
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I !
I I 1
I I 1
I I 1
44-4
I I I
I I I
i I I
1 I I
1 * I
.X-L- + -
Stereo-Atlas of Ostracod Shells 6, 146
Loxoconcha ovulata (6 of 10)
/
Stereo-Atlas of Ostracod Shells 6, 147 Loxoconcha ovulata (7 of 10)
Date of publication: Costa’s Paleontologia del Regno di Napoli was published both as a book (in three parts) and in the
volumes of the Atti Accad. Pontan. The two issues are essentially identical, except for minor details such
as page numbering, and it is obvious that the text and plates were pulled from one setting of type. The
most important difference concerns the dates of publication; 1864 for the Atti, but 1857 - 1863 for the
separate issue. On p.144 of the latter is a footnote reference to the compte rendue of the Paris Academy
of Sciences of 29.4.1861. It is certain, therefore, that p. 1 8 1 , where C. ovulata appears, must have been
published later than this date, probably in 1863, according to the title page. It seems likely that the whole
of pp.144 - 202, entitled ‘Capitolo IV, Malacostraci’, is of that date. The date of C. ovulata is, therefore,
given as ‘1863’.
Explanation of Plate 6, 148
Fig. 1, 9 LV, ext. lat. (1976.1087, 732 p m long); fig. 2,6 RV, ext. lat. (OS 11421, 760 pm long); fig. 3, 9 LV, ext. dors.
(1976.1088, 720 long); fig. 4, 6 LV, ext. dors. (1976.1089, 850 pm long).
Scale A (250 ^m; x 80), figs. 1 - 4.
u+.
4
■(--I—
Stereo-Atlas of Ostracod Shells 6, 149
Loxoconcha ovulata (9 of 10)
Remarks: Ascoli (1965, op. cit .) was the first to consider, after examination of Costa’s type collection, in the
University of Naples, that Cytherina ovulata Costa from the Quaternary of Naples (Costa ‘1863’, op. cit.)
was conspecific with Loxoconcha tumida Brady. He used L. ovulata (Costa) as the valid name, although,
under I.C.Z.N. Article 23b as it then stood, he should have considered the name a nomen oblitum and
referred the case to the Commission.
Ruggieri ( Atti Soc. Ital. Sci. nat., 106 (2), 106, 1967) criticised Ascoli’s revival of Costa’s name,
but he, too, did not refer the matter to the I.C.Z.N. Since then usage has been divided. By adopting the
senior synonym herein the Law of Priority is strictly applied. Wouters ( The Ostracodologist, 21, 8, 1974)
considered L. aequalis Terquem, 1878 and Cy there dispar Terquem 1878 as junior synonyms of
L. tumida Brady 1869, and therefore of L. ovulata.
Little is yet known about the ecology of this species. All records are from shallow marine
environments. It is interesting to note that Muller {Fauna Flora Golf. Neapel, 21, 1894) did not record
this species from the Bay of Naples, although recent collections have shown that it occurs there quite
commonly. Males more elongate than females.
Distribution: Recent: N Adriatic, (Uffenorde, op. cit.), Naxos, Greece (Barbeito-Gonzalez, op. cit.). As L. tumida:
Beskika Bay, Turkey and Piraeus, Greece (Brady, op. cit.), Monaco ( British Museum (Nat. Hist.)
collection), Italy and Cyprus (herein), Adriatic, (Masoli, op. cit.). ?Sub-Recent: Italy (herein), Cyprus
(author’s collection), Adriatic (Bremen, op. cit). Quaternary: Pozzuoli, Italy (Costa ‘1863’, op. cit.,
herein).
Explanation of Plate 6, 1 50
Fig. 1,6 RV, int. muse. sc. ( 1976. 1084); figs. 2, 4, 9 R V, terminal hinge elements (1976.1086); figs. 3,5 ,6 LV, terminal hinge
elements (1976.1090, 805 pm long).
Scale A (50 pm; x 418), fig. 1 ; scale B (50 pm\ x 564), figs. 2 - 5.
Loxoconcha ovulata (8 of 10)
Stereo-Atlas of Ostracod Shells 6, 150
Loxoconcha ovulata (10 of 10)
Stereo-Atlas of Ostracod Shells 6, 148
Stereo-Atlas of Ostracod Shells 6 ( 26) 1 5 1 - 1 53 ( 1979)
Index, Volume 6, 1979 (1 of 3)
General Index
Acanthocythereis hystrix (Reuss); 133 - 140
alata, Paijenborchellina; 27 - 30
albomaculata, Heterocythereis; 117-124
ambigua, Ghardaglaia; 99-106
Argenticytheretta (Argenticytheretta) punctata Rose sp. nov.; 91-98
Athersuch, J., On Acanthocythereis hystrix (Reuss); 133 - 140
Athersuch, J., On Heterocythereis voraginosa Athersuch sp. nov.; 125-132
Athersuch, J., On Loxoconcha ovulata (Costa); 141 - 150
Athersuch, J., On Pelecocy there sylvesterbradleyi Athersuch gen. et sp. nov.; 13-20
Athersuch, J. & Gooday, A., On Zabythocypris redunca Athersuch & Gooday sp. nov.; 21-26
Athersuch, J. & Whittaker, J. E., On Heterocythereis albomaculata (Baird); 1 17 - 124
bensoni, Cativella; 5-12
Brevibolbina dornbuschi Schallreuter; 71-74
capra, Caprabolbina; 63 - 66
Caprabolbina capra Schallreuter; 63 - 66
Cativella bensoni Neale; 5-12
dissecta, Kiesowia (Kiesowia); 79 - 86
dornbuschi, Brevibolbina; 71-74
elegans, Galliaecytheridea; 31-34
frigida, Homeokiesowia; 75-78
Fuller, N. & Lord, A., On Galliaecytheridea elegans (Sharapova); 31-34
Fuller, N. & Lord, A., On Galliaecytheridea gorodischensis Fuller & Lord sp. nov.; 35 - 42
Fuller, N. & Lord, A., On Galliaecytheridea miranda (Lyubimova); 43 - 46
Fuller, N. & Lord, A., On Galliaecytheridea volgaensis (Lyubimova); 47 - 50
Fuller, N. & Lord, A., On Mandelstamia ventrocornuta (Sharapova); 51-54
Fuller, N. & Lord, A., On Oligocythereis kostytschevkaensis (Lyubimova); 55 - 62
Galliaecytheridea elegans (Sharapova); 31-34
Galliaecytheridea gorodischensis Fuller & Lord sp. nov.; 35 - 42
Galliaecytheridea miranda (Lyubimova); 43 - 46
Galliaecytheridea volgaensis (Lyubimova); 47 - 50
Ghardaglaia ambigua Neale sp. nov.; 99 - 106
Gooday, A. & Athersuch, J., On Zabythocypris redunca Athersuch & Gooday sp. nov.; 21-26
gorodischensis, Galliaecytheridea; 35 - 42
Gurney, A., On Paijenborchellina alata Gurney sp. nov.; 27 - 30
Gurney, A., On Paijenborchellina venosa Gurney sp. nov.; 107 - 112
hammi, Schuleridea; 1 - 4
Heterocythereis albomaculata (Baird); 117-124
Heterocythereis voraginosa Athersuch sp. nov.; 125-132
Homeokiesowia frigida (Sarv); 75-78
hystrix, Acanthocythereis; 133 - 140
Kiesowia (Kiesowia) dissecta (Krause); 79 - 86
kostytschevkaensis, Oligocythereis; 55-62
Lord, A. & Fuller, N., On Galliaecytheridea elegans (Sharapova); 31-34
Lord, A. & Fuller, N., On Galliaecytheridea gorodischensis Fuller & Lord sp. nov.; 35 - 42
Lord, A. & Fuller, N., On Galliaecytheridea miranda (Lyubimova); 43 - 46
Lord, A. & Fuller, N., On Galliaecytheridea volgaensis (Lyubimova); 47 - 50
Lord, A. & Fuller, N., On Mandelstamia ventrocornuta (Sharapova); 51-54
Lord, A. & Fuller, N., On Oligocythereis kostytschevkaensis (Lyubimova); 55-62
Loxoconcha ovulata (Costa); 141 - 150
Mandelstamia ventrocornuta (Sharapova); 51-54
miranda, Galliaecytheridea; 43 - 46
Monoceratina scrobiculata Triebel & Bartenstein; 1 13 - 116
Neale, J. W., On Cativella bensoni Neale; 5-12
Neale, J. W., On Ghardaglaia ambigua Neale sp. nov.; 99 - 106
Neale, J. W., On Schuleridea hammi (Triebel); 1 - 4
Oligocythereis kostytschevkaensis (Lyubimova); 55 - 62
ovulata, Loxoconcha; 141-150
Paijenborchellina alata Gurney sp. nov.; 27 - 30 «
Paijenborchellina venosa Gurney sp. nov.; 1 07 - 112
Pelecocy there sylvesterbradleyi Athersuch gen. et sp. nov.; 13-20
Platybolbina (Reticulobolbina) spongiosoreticulata Schallreuter; 67 - 70
posterobicarinatum , Pyxion; 87 - 90
punctata, Argenticytheretta (Argenticytheretta); 91-98
Pyxion posterobicarinatum Schallreuter sp. nov.; 87 - 90
Stereo-Atlas of Ostracod Shells 4,152
Index, Volume 6, 1979 (2 of 3)
redunca, Zabythocypris; 21-26
Rose, J. F., On Argenticytheretta (Argenticytheretta) punctata Rose sp. nov.; 91-98
Schallreuter, R. E. L., On Brevibolbina dornbuschi Schallreuter; 71-74
Schallreuter, R. E. L., On Caprabolbina capra Schallreuter; 63 - 66
Schallreuter, R. E. L., On Homeokiesowia frigida (Sarv); 75-78
Schallreuter, R. E. L., On Kiesowia (Kiesowia) dissecta (Krause); 79 - 86
Schallreuter, R. E. L., On Platybolbina (Reticulobolbina) spongiosoreticulata Schallreuter; 67 - 70
Schallreuter, R. E. L., On Pyxion posterobicarinatum Schallreuter sp. nov.; 87 - 90
Schuleridea hammi (Triebel); 1 - 4
scrobiculata, Monoceratina; 1 1 3 - 116
Sheppard, L. M., On Monoceratina scrobiculata Triebel & Bartenstein; 1 13 - 116
spongiosoreticulata, Platybolbina (Reticulobolbina) ; 67 - 70
sylvesterbradleyi, Pelecocy there; 13-20
venosa, Paijenborchellina; 107 - 112
ventrocornuta, Mandelstamia; 51-54
volgaensis, Galliaecytheridea; 47 - 50
voraginosa, Heterocythereis; 125-132
Whittaker, J. E. & Athersuch, J., On Heterocythereis albomaculata (Baird); 1 17 - 124
Zabythocypris redunca Athersuch & Gooday sp. nov.; 21-26
Paijenborchellina venosa; 107-112
Pelecocythere sylvesterbradleyi; 13-20
Zabythocypris redunca; 21-26
Index; Geographical Location
See 1 (2) 5 - 22 (1973) for explanation of the Schedules in the Universal Decimal Classification
Cativella bensoni; 5-12
Stereo-Atlas of Ostracod Shells 4,153
Index, Volume 6, 1979 (3 of 3)
Stereo-Atlas of Ostracod Shells 6 ( 27) 154 (1979) Pterygocythereis jonesii (1 of 1 )
595.337.14 (1 19.4 + 1 19.9) (4) : 551.351
Addendum to
ON PTERYGOCYTHEREIS JONESII (BAIRD)
by John Athersuch
(B P Research Centre, Sunbury, England)
Editorial comment: Owing to a shortage of space in A Stereo-Atlas of Ostracod Shells 5, 9 - 16, 1978, the generic diagnosis
was omitted. It is, therefore, presented here in order to complete the aforementioned paper.
Genus PTERYGOCYTHEREIS Blake, 1933
1933 Cythereis (Pterygocythereis) Blake; in: Biological Survey of the Mount Desert Region, Wistar Institute of Anatomy and
Biology, Philadelphia, p. 239.
Type-species (by original designation): Cythereis jonesii Baird, 1850.
Diagnosis: Carapace quadrate with prominent, spinose, ventro-lateral alae terminating in a strong spine. Anterior and
posterior margins spinose, carinate or both. Eye tubercles conspicuous. Muscle scar pattern and hinge
typical for Trachyleberidinae.
A Stereo -Atlas of Ostracod Shells
edited by R. H. Bate, J. W. Neale, Lesley M. Sheppard
and David J. Siveter
Volume 6, 1979
Part 1 (pp. 1 - 70) ; 29th June, 1979
Part 2 (pp. 77 - 154) ; 29th November, 1979
Published by The British Micropalaeontological Society
in association with Robertson Research International Ltd., Llandudno, Wales
Stereo-Atlas of Ostracod Shells 6, ii
Contents
Contents
1 On Schuleridea hammi (Triebel); by J. W. Neale 1
2 On Cativella bensoni Neale; by J. W. Neale 5
3 On Pelecocythere sylvesterbradleyi Athersuch gen. et sp. nov.; by J. Athersuch 13
4 On Zabythocypris redunca Athersuch & Gooday sp. nov.; by J. Athersuch & A. Gooday 21
5 On Paijenborchellina alata Gurney sp. nov.; by A. Gurney 27
6 On Galliaecytheridea elegans (Sharapova); by N. Fuller & A. Lord 31
7 On Galliaecytheridea gorodischensis Fuller & Lord sp. nov.; by N. Fuller & A. Lord 35
8 On Galliaecytheridea miranda (Lyubimova); by N. Fuller & A. Lord 43
9 On Galliaecytheridea volgaensis (Lyubimova); by N. Fuller & A. Lord 47
10 On Mandelstamia ventrocornuta (Sharapova); by N. Fuller & A. Lord 51
1 1 On Oligocythereis kostytschevkaensis (Lyubimova); by N. Fuller & A. Lord 55
12 On Caprabolbina capra Schallreuter; by R. E. L. Schallreuter 63
13 On Platybolbina (Reticulobolbina) spongiosoreticulata Schallreuter; by R. E. L. Schallreuter 67
14 On Brevibolbina dornbuschi Schallreuter; by R. E. L. Schallreuter 71
15 On Homeokiesowia frigida (Sarv);by R. E. L. Schallreuter 75
1 6 On Kiesowia (Kiesowia) dissecta (Krause); by R. E. L. Schallreuter 79
17 On Pyxion posterobicarinatum Schallreuter sp. nov.; by R. E. L. Schallreuter 87
18 On Argenticytheretta (Argenticytheretta) punctata Rose sp. nov.; by J. F. Rose 91
19 On Ghardaglaia ambigua Neale sp. nov.; by J. W. Neale 99
20 On Paijenborchellina venosa Gurney sp. nov.; by A. R. Gurney 107
21 On Monoceratina scrobiculata Triebel & Bartenstein;by L. M. Sheppard 1 13
22 On Heterocythereis albomaculata (Baird); by J. Athersuch & J. E. Whittaker 117
23 On Heterocythereis voraginosa Athersuch sp. nov.; by J. athersuch 125
24 On Acanthocythereis hystrix (Reuss); by J. Athersuch 133
25 On Loxoconcha ovulata (Costa); by J. Athersuch 141
26 Index for Volume 6, 1979 151
27 Addendum to On Pterygocythereis jonesii (Baird); by J. Athersuch 154
Stereo -Atlas of Ostracod Shells: Vol.6, Part 2
CONTENTS
6 (14) 71 - 74 On Brevibolbina dornbuschi Schallreuter; by R. E. L. Schallreuter (£1.10)
6 (1 5) 75 - 78 On Homeokiesowia frigida (Sarv); by R. E. L. Schallreuter (£1.10)
6 (16) 79 - 86 On Kiesowia (Kiesowia) dissecta (Krause); by R. E. L. Shallreuter (£2.15)
6 (17) 87 - 90 On Pyxion posterobicarinatum Schallreuter sp. nov.; by R. E. L. Schallreuter (£1.10)
6 (18) 91 - 98 On Argenticytheretta ( Argenticytheretta) punctata Rose sp. nov.;by J. F. Rose (£2.15)
6 (19) 99 - 106 On Ghardaglaia ambigua Neale sp. nov.; by J. W. Neale (£2.15)
6 (20) 107-112 On Paijenborchellina venosa Gurney sp. nov.; by A. R. Gurney (£1 .65)
6 (21) 1 13 - 1 16 On Monoceratina scrobiculata Triebel & Bartenstein; by L. M. Sheppard (£1 .10)
6 (22) 117 -124 On Heterocythereis albomaculata (Baird); by J. Athersuch & J. E. Whittaker (£2.15)
6 (23) 125 - 132 On Heterocythereis voraginosa Athersuch sp. nov.; by J. Athersuch (£2.15)
6 (24) 133 - 140 On Acanthocythereis hystrix (Reuss);by J. Athersuch (£2.15)
6 (25) 141 - 150 On Loxoconcha ovulata (Costa); by J. Athersuch (£2.50)
6 (26) 151 - 153 Index for Volume 6, 1979 (£0.60)
6 (27) 1 54 Addendum to On Ptery’gocythereis jonesii (Baird); by J. Athersuch (£0.30)
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