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FIELDIANA: BOTANY
A Continuation of the
BOTANICAL SERIES
of
FIELD MUSEUM OF NATURAL HISTORY
VOLUME 31
The Library of the
JAN 1 8 1979
FIELD MUSEUM OF NATURAL HISTORY
CHICAGO, U. S. A.
520.5
•FB
v-31
TABLE OF CONTENTS
PAGE
1. Two New Species of Palms from Nicaragua. By S. F. Glassman ... 1
2. Tropical American Plants, VI. By Louis O. Williams 11
3. Agriculture, Tehuacan Valley. By C. Earle Smith, Jr 49
4. Flora, Tehuacan Valley. By C. Earle Smith, Jr 101
5. Preliminary Studies in the Palm Genus Syagrus Mart, and Its Allies.
By S. F. Glassman 145
6. Tropical American Plants, VII. By Louis 0. Williams 165
7. Supplement to Orchids of Guatemala. By Donovan S. Correll . . . .175
8. Preliminary Notes on Scrophulariaceae of Peru. By Gabriel Edwin . . 223
9. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 233
10. Tropical American Plants, VIII. By Louis O. Williams 247
11. Notes on the Flora of Costa Rica, I. By William C. Burger 273
12. A New Eurystyles from Nicaragua. By Alfonso H. Heller 279
13. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 285
14. A Revision of the Family Geastraceae. By Patricio Ponce de Leon . . 303
15. Studies in American Plants. By Dorothy N. Gibson 353
16. Two New Nicaraguan Juglandaceae. By Antonio Molino R 357
17. Studies in the Palm Genus Syagrus Mart. By S. F. Glassman .... 363
18. Tropical American Plants, IX. By Louis O. Williams 401
STUDIES IN AMERICAN PLANTS
DOROTHY N. GIBSON
TWO NEW NICARAGUAN JUGLANDACEAE
ANTONIO MOLINA R.
STUDIES IN THE PALM GENUS SYAGRUS MART.
S. F. GLASSMAN
TROPICAL AMERICAN PLANTS, IX
LOUIS 0. WILLIAMS
FIELDIANA: BOTANY
VOLUME 31 NUMBERS 15, 16, 17, 18
Published by
FIELD MUSEUM OF NATURAL HISTORY
MAY 23, 1968
JUL 11 1368
STUDIES IN AMERICAN PLANTS
DOROTHY N. GIBSON
Custodian of the Herbarium
FIELDIANA: BOTANY
VOLUME 31, NUMBER 15
Published by
FIELD MUSEUM OF NATURAL HISTORY
MAY 23, 1968
PUBLICATION 1045
Library of Congress Catalog Card Number: 68-26374
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
Studies In American Plants
DOROTHY N. GIBSON
During recent studies of the Hydrophyllaceae and Polemoniaceae
for the Floras of Guatemala and Peru, it became clear that several
concepts of species within certain highly variable genera of these fam-
ilies were incorrect, due probably to the limited amount of compara-
tive material available to earlier workers. Dr. Standley was becom-
ing aware of this situation as early as 1945 when he occasionally
noted in manuscript that certain species "might prove to be only
varieties." Since then, additional collections, by providing enough
material to show the wide range of variability in these plants, have
brought us to the conclusion that some of these minor differences are
not constant and therefore do not merit specific rank.
The notes in this paper were made possible by study of collections
borrowed from the United States National Herbarium, New York
Botanical Garden and Gray Herbarium of Harvard University, as
well as those on deposit in the herbarium of the Field Museum, and
I wish to express my thanks to the curators of those herbaria.
HYDROPHYLLACEAE
Wigandia urens var. caracasana (HBK.) comb. nov. Wigandia
caracasana HBK. Nova Gen. & Sp. 3: 128. 1819. W. macrophylla
Cham. & Schlecht. Linnaea 6: 382. 1831. W. scorpioides Choisy,
Ann. Sc. Nat. Se*r. 1, 30: 249. 1833. W. caracasana var. macrophylla
(Cham. & Schlecht.) Brand in Engler, Pflanzenr. IV, 251: 136. 1913.
W. caracasana var. calycina Brand, I.e. W. kunlhii var. intermedia
Brand, I.e. 137. W. kunthii var. viscosa (Donn.-Sm.) Brand, I.e.
W. caracasana var. viscosa (Donn.-Sm.) Macbride, Contr. Gray Herb.
New Ser. 49:42. 1917.
Mexico to Costa Rica; Colombia and Venezuela.
Differs from W. urens only in its lack of bristles on leaves and
stems. The leaves usually appear velvety, without bristles on the
midvein. The calyx lobes and capsule, however, may be hispid to
bristly-hirsute, as in W. urens. Earlier concepts of specific and vari-
353
354 FIELDIANA: BOTANY, VOLUME 31
etal differences were apparently based on such tenuous characters as
vesture (highly variable in the genus) and style length (the styles of
both the typical species and the variety elongate markedly after
an thesis).
POLEMONIACEAE
Cobaea lutea f. villosa (Standl.) comb. nov. C. villosa Standl.
Contr. U. S. Nat. Herb. 17: 454. 1914.
Differs from C. lutea only by villous nodes and by occasional vil-
losity and/or puberulence of petioles.
Known only from El Salvador.
Cobaea lutea f. viorna (Standl.) comb. nov. C. viorna Standl.
Contr. U. S. Nat. Herb. 17: 453. 1914.
Differs from C. lutea only in the absence of long cilia on sepals
and leaf bases. Standley, in manuscript, placed his C. viorna in
synonymy with C. lutea.
Known only from Guatemala.
Cobaea pachysepala f. tomentulosa (Standl.) comb. nov.
C. tomentulosa Standl. Contr. U. S. Nat. Herb. 17: 457. 1914.
Differs from C. pachysepala only in more dense vesture: the stems
may be obscurely tomentulose to densely villous, the nodes often
densely white-villous, the petiolules short-villous to wooly, the leaf-
lets sparsely pubescent to villous beneath, especially along veins, the
sepals usually densely tomentulose outside as well as along inside
margins, and the corollas may have more scattered puberulence on
the lobes.
Although Standley, in his original descriptions, states that there
are eight or nine seeds in each cell of the capsule of C. pachysepala
and four to six seeds in each cell of his C. tomentulosa, I found only
six or seven seeds in each locule of both C. pachysepala and C. tomen-
tulosa. Further, Standley, in manuscript, says of C. tomentulosa:
"Closely related to C. pachysepala, of which it may be only a form
or variety."
Publications 1045, 1046, 1047, and 1048
UNIVERSITY OF ILLINOIS-URBANA