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FIELDIANA: BOTANY

A Continuation of the

BOTANICAL SERIES

of

FIELD MUSEUM OF NATURAL HISTORY

VOLUME 31

The Library of the

JAN 1 8 1979

FIELD MUSEUM OF NATURAL HISTORY

CHICAGO, U. S. A.

520.5

-F5

vi 31

TABLE OF CONTENTS

PAGE

1. Two New Species of Palms from Nicaragua. By S. F. Glassman ... 1

2. Tropical American Plants, VI. By Louis O. Williams 11

3. Agriculture, Tehuacan Valley. By C. Earle Smith, Jr 49

4. Flora, Tehuacan Valley. By C. Earle Smith, Jr 101

5. Preliminary Studies in the Palm Genus Syagrus Mart, and Its Allies.

By S. F. Glassman 145

6. Tropical American Plants, VII. By Louis 0. Williams 165

7. Supplement to Orchids of Guatemala. By Donovan S. Correll .... 175

8. Preliminary Notes on Scrophulariaceae of Peru. By Gabriel Edwin . . 223

9. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 233

10. Tropical American Plants, VIII. By Louis O. Williams 247

11. Notes on the Flora of Costa Rica, I. By William C. Burger 273

12. A New Eurystyles from Nicaragua. By Alfonso H. Heller 279

13. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 285

14. A Revision of the Family Geastraceae. By Patricio Ponce de Leon . . 303

15. Studies in American Plants. By Dorothy N. Gibson 353

16. Two New Nicaraguan Juglandaceae. By Antonio Molino R 357

17. Studies in the Palm Genus Syagrus Mart. By S. F. Glassman .... 363

18. Tropical American Plants, IX. By Louis O. Williams 401

--7

PRELIMINARY STUDIES IN THE PALM GENUS

SYAGRUS MART. AND ITS ALLIES

S. F. GLASSMAN

mUMARTIfTHE

TROPICAL AMERICAN PLANTS, VII

LOUIS 0. WILLIAMS

ft waian or ILLINOIS

SUPPLEMENT TO ORCHIDS OF GUATEMALA

AND BRITISH HONDURAS

DONOVAN S. CORRELL

I

oo

CM

FIELDIANA: BOTANY

VOLUME 31, NUMBERS 5, 6 AND 7

Published by

CHICAGO NATURAL HISTORY MUSEUM

DECEMBER 22, 1965

BIOLOGY

101 BURRILL HALJJ

SUPPLEMENT TO

ORCHIDS OF GUATEMALA

AND BRITISH HONDURAS

DONOVAN S. CORRELL

Principal Botanist, Texas Research Foundation

FIELDIANA: BOTANY

VOLUME 31, NUMBER 7

Published by

CHICAGO NATURAL HISTORY MUSEUM

DECEMBER 22, 1965

PUBLICATION 999

The author wishes to acknowledge the assistance of

Mr. Charles Schweinfurth during the preparation of this paper.

Library of Congress Catalog Card Number: 65-291 73

PRINTED IN THE UNITED STATES OF AMERICA

BY CHICAGO NATURAL HISTORY MUSEUM PRESS

Supplement to the Orchids of Guatemala

(and British Honduras)

Since the appearance of Oakes Ames' and my work on the orchids

of Guatemala (including British Honduras), published by Chicago

Natural History Museum (1952-1953, Fieldiana: Botany, 26, Nos.

1 and 2), important collections of several individuals have been

made available to us. These are notably those of Percy H. Gentle

from British Honduras, and C. L. Lundell and Elias Contreras from

the Department of Pete"n, Guatemala, deposited in the Lundell Her-

barium, Chicago Natural History Museum, and elsewhere. In addi-

tion to these, there are some specimens of various collectors in the

United States National Herbarium which were inadvertently omitted

in the original publication.

In view of the continued, if not increasing, interest in the orchids

of this part of Latin America, it was thought advisable to make avail-

able this additional information. In citing supplementary distri-

bution data for each taxon, page reference is made to where it is

described and treated in Orchids of Guatemala. In the case of new

additions to the flora, a brief description is included of the taxon, and

the salient characteristics that distinguish it from all other presently

known orchid taxa in Guatemala and British Honduras are given.

Orchids of Guatemala included a total of 527 species or 552 specific

and infraspecific taxa in 89 genera. The present publication brings

to 542 species or 567 specific and infraspecific taxa in 90 genera the

number now known to occur in Guatemala and British Honduras.

One new species, Pleurothallis tikalensis, and one new combina-

tion, Scaphyglottis major, are proposed. The genus Warrea is defi-

nitely new to our flora and apparently the genus Epistephium should

be included. The species new to our flora are Habenaria Leprieurii,

Vanilla Hartii, Elleanthus caricoides, Spiranthes pseudogoodyerioides,

Pleurothallis gelida, P. ovatilabia, Epidendrum cystosum, E. equitanti-

folium, E. sculptum, Bletia tenuifolia, Warrea costaricensis, Coryanthes

maculata, lonopsis satyrioides and Campylocentrum hondurense.

177

178 FIELDIANA: BOTANY, VOLUME 31

ENUMERATION OF SPECIES

Habenaria bractescens Lindl.; Ames & Correll, p. 28, fig. 4.

Guatemala. — Pete"n: Dos Lagunas, Contreras 1771.

Habenaria clypeata Lindl. ; Ames & Correll, p. 28, fig. 5.

British Honduras. — Toledo: Monkey River, between Monkey

River and Cockscomb, Gentle 4229A.

Habenaria distans Griseb. ; Ames & Correll, p. 32, fig. 6.

Guatemala. — Pete"n: Dolores, between kms. 86/87, east of Macha-

quila Road, Contreras 2976.

Habenaria Leprieurii Reichb. f. Linnaea 19: 376. 1846; Schultes,

Native Orchids of Trinidad and Tobago 28. 1960. H. mesodactyla

Griseb. Fl. Brit. W. Ind. Isl. 644. 1864; Ames, Orch. 4: 211, pi. 70.

1910; Williams, Ceiba 5(1): 14. 1956. H. Cruegeri Cogn. in Urban,

Symb. Ant. 6: 303. 1910.

On sandy open pine ridges and uplands. Uncommon in British

Honduras, Panama, Trinidad, and northern South America.

Plant slender, rigidly erect, up to 5 dm. tall, with one or more tuberoids at

base. Leaves linear-setaceous, distant. Flowers greenish yellow, distantly placed

in a 1- to 7-flowered spike. Bracts ovate-lanceolate to lanceolate, acuminate, 1-

1.5 cm. long, exceeded by the incurved conspicuously ribbed pedicellate ovary.

Sepals often with prominently ciliate margins, 5-6 mm. long; dorsal sepal broadly

ovate-elliptic, obtuse, concave, about 3 mm. wide; lateral sepals obliquely ovate-

lanceolate, acuminate, about 2 mm. wide below middle. Petals bipartite; anterior

lobe setaceous, about 5 mm. long, usually much exceeding the linear posterior lobe.

Lip tripartite, about 6 mm. long, the base about 2 mm. wide; lateral lobes linear to

setaceous, curved, mostly somewhat exceeding the linear pendulous mid-lobe.

This species was not included in Ames and Correll (1952), but

Williams (1956) reported it from British Honduras. It is quite pos-

sible that our plant should be at least varietally segregated from typ-

ical H. Leprieurii. It differs from the island plant in having larger

flowers with the lateral lobes of the lip exceeding the mid-lobe and

the anterior lobe of the petals exceeding the posterior lobe. Also the

sepals are often conspicuously cellular-ciliate, a characteristic not

noted in typical H. Leprieurii. The flowers of our plants agree per-

fectly with the illustration of H. mesodactyla in Ames, but it does not

agree with any of the descriptions published of this plant, including

the latest by Schultes.

British Honduras. — El Cayo: Mountain Pine Ridge, San Augus-

tin, Lundell 6712. — Stann Creek: Commerce Bight Pine Ridge, Gentle

CORRELL: ORCHIDS OF GUATEMALA 179

8291, 8361.— Toledo: near Jenkins Creek, north of Monkey River,

Gentle 4097. Punta Gorda-San Antonio Road, Gentle 6597.

Habenaria novemfida Lindl. ; Ames & Correll, p. 38, fig. 5.

British Honduras. — Toledo: in sacatal near Machaca, Gentle 6898.

Habenaria pauciflora (Lindl.) Reichb. f.; Ames & Correll, p. 39.

British Honduras. — Toledo: near Jenkins Creek, north of Monkey

River, Gentle 4036.

Habenaria repens Nutt. ; Ames & Correll, p. 42, figs. 5, 9.

Guatemala. — Solola: Panajachel, swamp along Lake Atitlan,

0. Degener & I. Degener 558. — Pete"n: Tikal National Park, Aguada

Terminos, Lundell 16609.

British Honduras. — Stann Creek: Commerce Bight Pine Ridge,

Gentle 8336.— Toledo: 12 Miles, Punta Gorda-San Antonio Road,

Gentle 6596.

Habenaria strictissima var. odontopetala (Reichb. f.) L.

Wms.; Ames & Correll, p. 46, figs. 5, 10.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa FC", Ar-

royo Corriental, on Aguada Terminos Road, Contreras 1934.

Vanilla Hartii Rolfe Kew Bull. (1899) 133; Porteres in Bouriquet

Le Vanillier et la Vanille (Encycl. Biol. XLVI) 250, fig. 113. 1954;

Schultes Native Orchids of Trinidad and Tobago 39. 1960.

Climbing on trees in moist forests at low elevations, commonly on

ridges. Uncommon in Trinidad, Tobago, British Honduras, Guate-

mala, Honduras (Dept. Atlantida, Standley 52824), Costa Rica (Prov.

Limon, Standley & Valeria 48905), and Panama (Prov. Panama,

Standley 29353).

Stem slender. Leaves with a short twisted petiole up to 1 cm. long; blade

fleshy but thinnish, elliptic-lanceolate to ovate-lanceolate or oblong-elliptic, more

or less abruptly acuminate, up to 11 cm. long and 3.5 cm. wide, usually smaller,

about as long as the internodes. Inflorescence axillary, compactly few-flowered, the

floral axis 2 cm. or less long. Floral bracts broadly ovate, rarely more than 5 mm.

long. Flowers white. Sepals and petals similar, very narrowly linear-oblanceolate,

narrowly obtuse at apex, up to about 4 cm. long and 7 cm. wide above the middle.

Lip attached to the column almost to its apex to form a tube, with the tube slightly

inflated at base, about 4 cm. long from its attachment at base of column to the

crenulate-erose apical lamina, about 1.5 cm. wide across the free apical portion

when spread out; disk with veins somewhat thickened toward apex of lip, with a

small tuft of retrorse centrally placed plates arising about 12 mm. below apex of

lip. Fruit short, 5-8 cm. long.

180 FIELDIANA: BOTANY, VOLUME 31

This species, which until now was thought to be endemic to Trini-

dad and Tobago, is new to Central America. The Central American

plants have, on the whole, larger leaves than those described for the

island plants by both Porteres and Schultes, but Schweinfurth and I

can find no clear distinction for separating the plants from these

two regions. It may be distinguished from all other vanilla-species

in our area not only by the floral difference and the long-acuminate

leaves but by the extremely abbreviated floral axis.

Guatemala. — Pete"n: Dolores, on Rio Mopan trail, Contreras 3063.

British Honduras. — Toledo: Bolo Camp, upper reach, Golden

Stream, Gentle 4560. Between Orange Point and Moho River, Gen-

tle 7673.

Vanilla Pfaviana Reichb. f.; Ames & Correll, p. 55, fig. 15.

British Honduras. — Toledo: near river beyond Columbia, Gentle

6152. Near ocotal pine ridge, 1-3 miles in trail from 7 miles Punta

Gorda-San Antonio Road, Gentle 6957, 7078. Feeders Road leading

to Big Fall, Gentle 7023. Near Columbia, Gentle 7108.

Vanilla planifolia Andrews; Ames & Correll, p. 57, fig. 16.

Guatemala. — Pete"n: Tikal National Park, Tikal, Contreras 1841

(narrow-leaved form) . About 2 km. south of Dolores, Contreras 2239.

Tikal National Park, Bajo de Santa Fe\ Arroyo Corriental, on Aguada

Terminos road, Lundell 15818, 15940.

British Honduras. — Toledo: Bolo Camp, upper reach of Golden

Stream, Gentle 4521. Canada Hill-Alta Visa Road, Gentle 8054.

The following collections have their inflorescences terminating a

leafy branch or a mainstem. This is not a common occurrence, at

least not in herbarium specimens, and it might be of some significance

in this species.

British Honduras. — Toledo: Near Jacinto Hills, Gentle 5106. Feed-

ers Road, 14 Miles, San Antonio-Punta Gorda Road, Gentle 6721.

Beyond San Antonio, Gentle 7557.

Vanilla pompona Schiede; Ames & Correll, p. 59.

Guatemala. — Pete*n: Dolores, about 800 m. south of the village on

the Machaquila Road, Contreras 2333. Dolores, on Rio Mopan trail,

Contreras 3063. Canchacan, Lundell 16457.

Epistephium sp. An unidentifiable specimen from British Hon-

duras which I believe belongs to this genus is in the Ames Orchid

CORRELL: ORCHIDS OF GUATEMALA 181

Herbarium at Harvard University. If correct, this would be its only

known occurrence in North America.

Elleanthus caricoides Nash, Bull. Torr. Bot. Club 34: 119, pi. 7,

1907.

Epiphytic on trees in moist forests and groves. Uncommon in

British Honduras, Nicaragua and Costa Rica.

Plant densely tufted, essentially smooth and glabrous, up to 4 dm. tall. Stems

slender, numerous, fasciculate, terete, about 2 mm. in diameter, sparsely spotted

with purple. Leaves 5 to 7; sheaths grooved, spotted with purple, much shorter

than the blades; blades flat, shining, 7-nerved, erect, somewhat twisted and re-

curved above base, linear to linear-lanceolate, long-acuminate at apex, narrowed

and sheathing at base, 1-2 dm. long, about 7 mm. wide, reduced above. Inflores-

cence congested and subcapitate, 2-3 cm. long, on a peduncle less than 1.5 cm.

long. Floral bracts scarious, ovate-lanceolate, long-acuminate, greenish, 2 to 3

times as long as the white flowers, nearly erect and recurved above the middle.

Flowers and ovary appressed-pubescent with black-brown hairs, about 1 cm. long,

gibbous on one side. Lateral sepals oblong to oblong-elliptic, rather abruptly con-

tracted above into a laterally compressed and keeled apex, 7-8 mm. long, about

3 mm. wide. Dorsal sepal 5-6 mm. long and about 2 mm. wide. Petals linear-

oblong, somewhat dilated at the rounded and apiculate apex, 6-7 mm. long, 1.5-

2 mm. wide. Lip nearly campanulate, completely surrounding the column so that

its margins meet, 7-8 mm. long, when spread out 10-12 mm. wide at the truncate

and sparingly ciliate apex, 5-6 mm. wide at the base; disk with a 2-ridged crest just

below the middle, the 2 oblong appendages near the base of the lip oblong and 1.5-2

mm. long and 1 mm. wide. Column 7-8 mm. long, 4-toothed at the broadened apex.

This species, which was not included in Ames and Correll (1952),

is new to British Honduras. It may be distinguished from all other

species of Elleanthus (cf . Ames and Correll, p. 60) found in our region,

especially E. poiformis Schltr., by its narrowly linear-lanceolate leaves

and the tight subcapitate inflorescence with long-attenuate floral

bracts that greatly exceed the flowers.

British Honduras. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4422. On hill top, Balsam Hill, Edwards

Road, beyond Columbia, Gentle 6329.

Elleanthus linifolius Presl; Ames & Correll, p. 62.

Guatemala.— Pete" n: Santo Toribio, north of road, Contreras 2753.

Dolores, on Machaquila Road, between kms. 85-87, Contreras 3021,

3135.

British Honduras. — Stann Creek: Dry Creek, Humming Bird

Highway, Gentle 8399.— Toledo: Jacinto Creek, Rio Grande, Gentle

4924. Punta Gorda-Seven Hills Road, ascending the Rio Grande,

182 FIELDIANA: BOTANY, VOLUME 31

Gentle 5003. Balsam Hill, Edwards Road, Gentle 6331. On hill slope

beyond Pate's Camp, Edwards Road beyond Columbia, Gentle 7159.

Sobralia decora Batem.; Ames & Correll, p. 64, figs. 18, 19.

Guatemala. — Alta Verapaz, Cacao, Finca Trece Aguas, G. P. Goll

48.— Pet<§n: Tikal National Park, Bajo de Santa Fe", Contreras 452.

On Remate Road, Tikal National Park, Contreras 1494. Dolores,

km. 82, west of Machaquila Road, Contreras 2728. Dolores, on Rio

Mopan trail, Contreras 3058. On Remate Road, Tikal National

Park, Lundell 16891.

British Honduras. — Stann Creek: Stann Creek -Sittee Road, Gen-

tle 7927. Chavarrias Road, Resemederes, across Columbia River,

Gentle 6279. Near San Antonio, Gentle 7535.

Sobralia f ragrans Lindl. ; Ames & Correll, p. 66.

Guatemala. — Pete"n: Dolores, kms. 83-86 on Machaquila Road,

Contreras 2107, 2938. Dolores, on Rio Mopan trail, about 3 km. east

of village, Contreras 2836.

British Honduras. — Stann Creek: Eve's Pine Ridge, Gentle 8508.

—Toledo: Jacinto Creek, Rio Grande, Gentle 4925. Near Jacinto

Creek, 9 Miles, Punta Gorda-San Antonio Road, Gentle 4962. Chava-

rrias Road, Resemederes, across Columbia River, Gentle 6259. Near

Mafridyle Lagoon, Gentle 7703.

Sobralia macrantha Lindl.; Ames & Correll, p. 68.

Guatemala. — Quiche", Chimique, Heyde & Lux 3515.

Sobralia mucronata Ames & C. Schweinf.; Ames & Correll,

p. 69.

British Honduras. — Toledo: near Carmelita Camp, Edwards Road

beyond Columbia, Gentle 7318. Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4314. Mira Mar Hill, Edwards Road,

beyond Columbia, Gentle 6404.

Prescottia stachyodes (Sw.) Lindl. ; Ames & Correll, p. 76.

Guatemala. — San Marcos: San Lorenzo, W. C. Shannon 449.

British Honduras. — Stann Creek: 20 miles Section Stann Creek

Valley, Gentle 9275.— Toledo: Cero Hill, Gentle 6991.

Spiranthes acaulis var. assurgens (Reichb. f.) Correll; Ames &

Correll, p. 100.

Guatemala. — Pete"n: Lacandon, Contreras 3462.

CORRELL: ORCHIDS OF GUATEMALA 183

Spiranthes adnata (Spreng.) Benth. ex Fawc.; Ames & Correll,

p. 100.

Although this species was included by Ames and Correll on the

basis of its occurrence in British Honduras, this is the first collection

that has been seen from Guatemala.

Guatemala. — Pete"n, Dolores, on Machaquila Road, between kms.

83-84, Contreras 2168.

Spiranthes costaricensis Reichb. f.; Ames & Correll, p. 106,

fig. 28.

Guatemala.— Pete'n: Tikal National Park, Tikal, Ibarra 54. Tikal

National Park, Tikal, Lundell 15636, 16796. On trail to Aguada La

Aurora, Tikal National Park, Lundell 15686. Aguada Las Cucas,

Tikal National Park, Lundell 16898.

British Honduras. — Toledo: On hill slope beyond Central Camp,

Edwards Road beyond Columbia, Gentle 7304.

Spiranthes orchioides (Sw.) A. Rich.; Ames & Correll, p. 118,

fig. 31.

Guatemala. — Izabal: Cristina, open grassy plain, S.F.Blake 7616.

British Honduras. — El Cayo: Road to Resthouse, Augustine,

Mountain Pine Ridge, D. R. Hunt 404. — Stann Creek: Broken Pine

Ridge, Stann Creek-Sittee road, Gentle 7931, 7950.— Toledo: Con-

demned Branch Pine Ridge, Gentle 5356, 5357. Beyond Columbia,

Gentle 6140. Between Punta Gorda-San Antonio Road and Moho

River, Gentle 6753. Cero Road, Gentle 7064.

Spiranthes pseudogoodyerioides L. Wms. Bot. Mus. Leafl.

Harvard Univ. 12(7): 231. PI. 33. 1946.

On rocks in shady moist places. Rare in Mexico (San Luis Potosi)

and Guatemala.

Small terrestrial plants, scapose, erect, up to 2 dm. tall. Leaves basal, petio-

late; lamina elliptic to ovate-elliptic, acute to somewhat acuminate, 5- to 7-nerved,

submembranous, 2-8 cm. long, 1.5-4 cm. wide; petiole slender, shorter than the

lamina. Scape pubescent, covered with scarious lanceolate acuminate bracts, the

raceme spicate and compact with small flowers. Sepals greenish, about 2 mm. long

and 1.2 mm. wide; dorsal sepal triangular-lanceolate, cochleate, 1-nerved, dorsally

pubescent; lateral sepals obliquely broadly lanceolate, acute, 3-nerved, dorsally pu-

bescent. Petals grayish red, elliptic, acute, arcuate, 1-nerved, about 1.5 mm. long

and 0.5 mm. wide. Lip whitish with the posterior half orange-color, triangular-

sagittate, somewhat fleshy, 1-nerved, 1.5-1.8 mm. long, 1-1.5 mm. wide. Column

about 1.2 mm. long; anther erect, ovate. Ovary densely pubescent.

184 FIELDIANA: BOTANY, VOLUME 31

This species is new to Guatemala. It is distinguished by its small

triangular-sagittate lip.

Guatemala. — Pete"n: In ramonal covering the ruins, Tikal National

Park, Contreras s.n.

Spiranthes rosulata (W. Baxt.) Lindl; Ames & Correll, p. 127.

Guatemala. — Santa Rosa, Cenaguilla, Heyde & Lux 3853.

Spiranthes seminuda Schltr. ; Ames & Correll, p. 130.

Guatemala.— Pet^n: Tikal National Park, Contreras 608, 1839,

2139; Ibarra 5Z;Lundell 15495, 15560, 16562, 16600, 16827, 16887.

Spiranthes tortilis (Sw.) L. C. Rich.; Ames & Correll, p. 134.

British Honduras. — El Cayo: Mountain Pine Ridge, near summit

of Baldy Beacon, 20 miles east northeast of Augustine, 3400 ft.,

D. R. Hunt 220.

Erythrodes purpurea Ames; Ames & Correll, p. 144.

Guatemala. — Pete*n: In high forest, south of Lacandon, Contreras

3461.

Erythrodes querceticola (Lindl.) Ames; Ames & Correll, p. 145,

fig. 39.

Guatemala. — Alta Verapaz: Cacao, Trece Aguas, F. L. Lewton

326. — Pete"n: Tikal National Park, Tikal, growing on top of Tem-

ple IV, Contreras 1801, 1802. Tikal National Park, Tikal, in ramonal

covering the ruins, Contreras 350, 1889;Lundell 15481, 15635, 16888.

British Honduras. — Stann Creek: S. J. Record s.n. — Toledo: Be-

yond Central Camp, Edwards Road beyond Columbia, Gentle 7264.

Tropidia Polystachya (Sw.) Ames; Ames & Correll, p. 151,

fig. 41.

This species was included in the Orchids of Guatemala, based on a

report of its occurrence there by Schlechter. The specimens cited

here are the first we have seen for Guatemala.

Guatemala. — Alta Verapaz: Cubilqiiitz, Tiirckheim 7999. — Pe-

te"n: Tikal National Park, Tikal, in ramonal covering the ruins, Con-

treras 145. Tikal National Park, in ramonal at base of Temple III,

Contreras 302.

Corymborchis forcipigera (Reichb. f.) L. Wms.; Ames &

Correll, p. 153, fig. 42.

Guatemala. — Pete"n: Dolores, between km. 84 and km. 85, east

of Machaquila Road, Contreras 2812.

CORRELL: ORCHIDS OF GUATEMALA 185

Stelis aemula Schltr. Beih. Bot. Centralbl. 36, Abt. 2: 385. 1918.

S. perplexa Ames; Ames & Correll, p. 176.

Guatemala. — Pete"n: 4-5 kms. south southeast of Dolores, Con-

treras 2486, 3030.

Stelis ciliaris Lindl.; Ames & Correll, p. 160, figs. 46-48.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 199, 404, 1976; Lundell 16195, 16882. Tikal

National Park, Bajo de Santa Fe", Contreras 585. Tikal National

Park, northeast of Tikal, Contreras 607. Remate, on Fallabon Road,

Contreras 963. Tikal National Park, Tikal, on top of temple in "H"

Group, Contreras 1042. Tikal National Park, Bajo de Santa Fe",

Arroyo Corriental, on Aguada Terminos road, Lundell 15796.

Stelis gracilis Ames; Ames & Correll, p. 164, figs. 52, 53.

Guatemala. — Pete"n: On old Machaquila Road, 800 m. south of

Dolores, Contreras 2340.

Stelis Johnsonii Ames; Ames & Correll, p. 169, fig. 56.

British Honduras. — Toledo: Jacinto Creek, Rio Grande, Gentle

4938. Moffredye Creek, San Antonio, Gentle 5083.

Stelis purpurascens A. Rich. & Gal.; Ames & Correll, p. 177,

fig. 62.

Guatemala. — Quiche"; near Iscopilas, F. C. Lehmann 1573.

Stelis rubens var. oxypetala (Schltr.) Ames; Ames & Correll,

p. 180, fig. 64.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 140, 251, 347; Lundell 15460. Tikal National

Park, northeast of Tikal, Contreras 1486. About 400 m. south of

Dolores, Contreras 2837, 2839. Dolores, between kms. 86-87, east

of Machaquila Road, Contreras 2975, 2981.

British Honduras. — Stann Creek: North Branch Section, Hum-

ming Bird Highway, Gentle 9223. Humming Bird Highway, beyond

Middlesex, Gentle 3046, 7971.— Toledo: Between Monkey River and

Cockscomb, Gentle 4261. Near Columbia, Gentle 6092.

Masdevallia linear! folia Ames; Ames & Correll, p. 188.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering ruins, Contreras 155. Tikal National Park, Tikal, growing on

top of Temple IV, Contreras 166A. Baja de Santa FC", Tikal National

Park, Contreras 562.

186 FIELDIANA: BOTANY, VOLUME 31

British Honduras. — Toledo: On hill slope near Pate's Camp, Ed-

wards Road, beyond Columbia, Gentle 7130. Between Machaca and

Rio Grande, Gentle 6936. Mafridyle Creek, Gentle 7608.

Masdevallia tubuliflora Ames; Ames & Correll, p. 191, fig. 68.

British Honduras. — Toledo: between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4417.

Lepanthes inaequalis Schltr.; Ames & Correll, p. 200.

Guatemala. — Chimaltenango: Near Tecpam, A.F. Skutch 520.

Lepanthes samacensis Ames; Ames & Correll, p. 202.

Guatemala. — Alta Verapaz: in mountain woods near Samac,

Turckheim 47.

Lepanthes turialvae Reichb. f.; Ames & Correll, p. 204.

Guatemala. — Pete*n: 2-3 kms. south of Dolores, Contreras 2355,

2614, 3143. Dolores, between km. 84-87, east of Machaquila Road,

Contreras 2809, 2977.

British Honduras. — Toledo: 1^ miles in from 7 Miles, San An-

tonio-Punta Gorda Road, Gentle 7083. Mafridyle Creek, Gentle 7607.

Near Jacinto Creek, 9 Miles, Punta Gorda-San Antonio Road, Gentle

4958.

Pleurothallis abjecta Ames; Ames & Correll, p. 210, fig. 75.

Guatemala. — Pete"n: Dolores, bordering the village clearing, Con-

treras 2201.

Pleurothallis Blaisdellii S. Wats.; Ames & Correll, p. 212, fig. 71.

British Honduras. — El Cayo: Humming Bird Highway, 39 miles

section, Gentle 9140. — Toledo: Cow Pen, near Monkey River, Gentle

4130. Near Jacinto Creek, 9 Miles, Punta Gorda-San Antonio Road,

Gentle 4956. Edwards Road beyond Columbia, Gentle 6258. Between

Orange Point and Moho River, Gentle 7628.

Pleurothallis Brighamii S. Wats.; Ames & Correll, p. 213,

fig. 72.

Guatemala. — Alta Verapaz : Cacao, Trece Aguas,F. L. Lewton 271.

— Pete'n: Tikal National Park, Tikal, in ramonal covering the ruins,

Contreras 279, 3238; Lundell 15454, 16196, 16855. Tikal National

Park, Bajo de Santa Fe\ Contreras 564. Tikal National Park, on top

of temple in "H" group, Contreras 1041. Tikal National Park, west

of Isla de los Pavos, Bajo de Santa Fe", Contreras 1335. Tikal National

Park, on Remate road south of Tikal, Lundell 15562.

CORRELL: ORCHIDS OF GUATEMALA 187

British Honduras.— El Cayo: Valentin, Lundell 6186, 6342.—

Toledo: Between Monkey River and Cockscomb, Gentle 4229.

Pleurothallis ciliaris (Lindl.) L. Wms.; Ames & Correll,

p. 219.

Guatemala. — Pete"n : Tikal National Park, Isla de los Pavos, Con-

treras 1321. Tikal National Park, Tikal, in ramonal covering the

ruins, Contreras 3239.

Pleurothallis compacta (Ames) Ames & C. Schweinf.; Ames &

Correll, p. 223.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Lundell 15433. Tikal National Park, on Remate

Road, Lundell 15558, 15563, 17122.

British Honduras. — Toledo: Edwards Road, beyond Columbia,

Gentle 6139, 6286-A. Mafridyle Creek, Gentle 7609.

Pleurothallis gelida Lindl. Bot. Reg. 27: Misc. p. 91. 1841;

Correll, Native Orchids of N. Amer. 250, pi. 98. 1950.

Epiphytic on trees and on rocks in forests. Widespread from

Florida and the West Indies to Mexico, through Central America to

northern South America.

Plant erect, cespitose, up to 6 dm. tall, usually much smaller. Roots from a

short creeping rhizome, slender, clustered. Secondary stems several, approximate,

glabrous, clothed with one or more tubular membranous sheaths that are 6-8 cm.

long, bearing a solitary leaf at the summit. Leaf ovate-elliptic to oblong-elliptic

or narrowly linear-oblong, obtuse to subacute, usually keeled through the center

beneath, 7-25 cm. long (including the petiole), 1.5-7 cm. wide, coriaceous. Racemes

1 to 15, clustered in the axil of the leaf, slender, many-flowered, 6-28 cm. long,

about 2 cm. in diameter, enclosed at the base by a short membranous spathe less

than 2 cm. long. Floral bracts minute, tubular, apiculate, scarious and translucent,

2-3 mm. long. Flowers pale yellow to greenish yellow, 2-ranked, with the slender

arching pedicellate ovaries 4-5 mm. long. Dorsal sepal oblong-elliptic to broadly

lanceolate, strongly concave and dorsally keeled at the base, obtuse to acute and

somewhat reflexed at the apex, with the inner surface pilose, 5-8 mm. long, about

3 mm. wide. Lateral sepals coherent at the base and forming a cup, narrowly ob-

long to elliptic, obtuse, dorsally keeled, pilose on the inner surface, 5-7 mm. long,

about 2.5 mm. wide near the base. Petals whitish, obliquely oblong-obovate, either

acute, broadly rounded, retuse or tridentate at the apex, 3-4 mm. long, about

2.5 mm. wide. Lip small, obscurely or obsoletely 3-lobed, narrowly cuneate or

sometimes pandurate in outline (rarely broadest below the middle), broadly trun-

cate and more or less retuse at the apex, arcuate-recurved in natural position, usu-

ally with a pair of minute lateral lobes or teeth; disc with a pair of erect keels near

the middle. Column short, with 2 minute teeth on each side at the apex. Capsule

ellipsoid, strongly 3-angled and triangular in cross section, light brown, 5-10 mm.

long, 3-5 mm. in diameter.

188 FIELDIANA: BOTANY, VOLUME 31

This species was not included in Ames and Correll (1952). The

specimens cited below are the first that have been seen from our re-

gion. The numerous rather large distinctive flowers are characteristic.

Guatemala. — Pete'n: On Rio Mopan Trail, about 2 km. southeast

of Dolores, Contreras 2517.

British Honduras/ — Toledo: Beyond Carmelita Camp, Edwards

Road beyond Columbia, Gentle 7215.

Pleurothallis Grobyi Batem. ex Lindl.; Ames & Correll, p. 230,

fig. 72.

Guatemala. — Pete'n: Remate, on Fallabon Road, Contreras 962,

972. Remate, west of the village, Contreras 992. Tikal National

Park, Tikal, on top of Temple IV, Contreras 1057. Tikal National

Park, Bajo de Santa Fe\ Arroyo Corriental, on Aguada Terminos

road, Contreras 1070. 2 km. south of Dolores, Contreras 2241. Tikal

National Park, in ramonal covering the ruins, Contreras 3235; Lun-

dell 15459, 16194. About 2 km. south of Lacandon, Contreras 3527.

Tikal National Park, in growth bordering airfield, Lundell 15951.

Tikal National Park, Tikal, Aguada Naranjal, Lundell 16166. Pop-

tun, about 2 km. east of village Lundell 16417.

British Honduras. — Toledo: Temash River, Gentle 5209. Near

Agua Caliente, near San Antonio, Gentle 5337. Edwards Road be-

yond Columbia, Gentle 6288 p.p., 7319. Big Hill, Gentle 7010. Joe

Taylor Creek, Gentle 7409, 7767. Between Orange Point and Moho

River, Gentle 7667. Condemned Branch Pine Ridge, Gentle 7758.

Pleurothallis hastata Ames; Ames & Correll, p. 231, fig. 75.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, Gentle 4298 p.p. Edwards Road beyond Columbia, Gentle

6288 p.p.

Pleurothallis hondurensis Ames; Ames & Correll, p. 234, fig. 78.

Guatemala. — Pete'n: Lacandon, Contreras 3292.

British Honduras. — Toledo: Cattle Pass, Swasey Branch, Mon-

key River, Gentle 4447. On hill slope near San Antonio, Gentle 6651.

Between Punta Gorda and Joe Taylor Creek, Gentle 6793. Feeder

Road leading to Big Fall, Gentle 7044. Mafridyle Creek area, Gentle

7614. Near Mafridyle Lagoon, Gentle 7690.

Pleurothallis lancilabris var. oxyglossa (Schltr.) C. Schweinf.;

Ames & Correll, p. 236.

CORRELL: ORCHIDS OF GUATEMALA 189

Guatemala. — Pete"n: 2-3 miles south of Dolores, Contrems 2354,

3142.

British Honduras. — Stann Creek: Eve's Pine Ridge, Gentle 8504.

Edwards Road beyond Columbia, Gentle 6207. Between Orange Point

and Moho River, Gentle 7653.

Pleurothallis Lewisae Ames; Ames & Correll, p. 238, fig. 76.

This species is new to British Honduras.

Guatemala. — Pete"n : Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 2005.

British Honduras. — Toledo: Punta Gorda-Seven Hills Road, as-

cending Rio Grande, Gentle 5007. Edwards Road beyond Columbia,

Gentle 6285. Mafridyle Creek area, Gentle 7616.

Pleurothallis Matudiana C. Schweinf.; Ames & Correll, p. 238.

This species is new to British Honduras.

British Honduras. — Toledo: Monday Morning Hill, Edwards

Road beyond Columbia, Gentle 6429.

Pleurothallis ovatilabia Ames & C. Schweinf., Sched. Orch. 10:

33. 1930.

In rain forests below 1500 m. alt. Uncommon in Guatemala,

Honduras, Costa Rica and Panama.

Plant small for the genus, 5.5-7 cm. tall, cespitose. Roots filiform, flexuous,

glabrous. Stems very short, 4-6 mm. long, entirely concealed and surpassed by

several loose scarious imbricating sheaths. Leaf oblanceolate-spatulate, gradually

narrowed to a rather long petiole; blade 2-4 cm. long, up to 6 mm. wide, minutely

bilobed and mucronate at the rounded apex, chartaceous, the mid-nerve sulcate

above and prominently carinate beneath; petiole 1-2 cm. long, slender, jointed

near the middle. Inflorescence solitary, subequaling or surpassing the leaf, 4.7-

5.5 cm. long. Peduncle up to the raceme 2.4-3.5 cm. long, filiform, with 2 or 3

scarious infundibuliform sheaths. Raceme short, often flexuous. Flowers very

small. Sepals ovate, 1.7-1.8 mm. long, acute to subacute, strongly cucullate,

1-nerved, dorsally keeled. Petals linear-lanceolate, falcate, 1.6-1.7 mm. long, sub-

acute, 1-nerved. Lip simple, ovate, 1.6-1.8 mm. long, subacute, densely papillose

on the inner surface. Column short, stout.

This species is new to Guatemala. It differs from its nearest ally,

P. stenostachya, in having solitary inflorescences that are relatively

elongate and even sometimes surpass the leaf. Its larger leaves,

shorter pedicels and ovate, acute perianth-segments distinguish it

from P. lancilabris, whose segments are lanceolate and acuminate.

Guatemala. — Pete"n: Canchacan, in high rain forest of southeast-

ern Pete"n, Lundell 16315.

190 FIELDIANA: BOTANY, VOLUME 31

Pleurothallis pansamalae Schltr.; Ames & Correll, p. 243.

British Honduras. — Toledo: Hill slope beyond Union Camp, Ed-

wards Road beyond Columbia, Gentle 6483.

Pleurothallis sertularioides (Sw.) Spreng.; Ames & Correll,

p. 252.

This species is new to British Honduras.

British Honduras. — Toledo: Near Jacinto Creek, Punta Gorda-

San Antonio Road, Gentle 4974. Between Condemned Branch Pine

Ridge and Moffredye Lagoon, Gentle 6043. Edwards Road beyond

Columbia, Gentle 6227, 6250, 6302. Near Orange Point, Gentle 7460.

Near Mafridyle Creek, near San Antonio, Gentle 7560. Condemned

Branch Pine Ridge, Gentle 7761.

Pleurothallis setosa C. Schweinf.; Ames & Correll, p. 253.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, Gentle 4298 p.p. Edwards Road beyond Columbia, Gentle

6288 p.p.

Pleurothallis stenostachya Reichb. f.; Ames & Correll, p. 254,

fig. 78.

Guatemala. — Pete"n: Tikal National Park, Tikal, on top of Tem-

ple IV, Contreras 165. Tikal National Park, Tikal, on ramonal cov-

ering the ruins, Contreras 445; Lundell 16879, 17084. Aguada El

Cartucho, 8-11 km. west of Dos Lagunas, Contreras 1594, 1746.

Tikal National Park, Tikal, on top of Temple V, Contreras 3229.

Lacandon, Contreras 3325. Tikal National Park, on Remate road

south of Tikal, Lundell 15555. Tikal National Park, Tikal, on Agu-

ada Pucte trail, Lundell 16825.

British Honduras. — El Cayo: On hill slope, 41 Miles Section,

Humming Bird Highway, Gentle 9153. Near Jacinto Creek, 9 Miles,

Punta Gorda-San Antonio Road, Gentle 4957. Near San Antonio,

Gentle 5439. Edwards Road, beyond Columbia, Gentle 6135, 6287.

Joe Taylor Creek, Gentle 7412. Between Orange Point and Moho

River, Gentle 7626. Beyond Mafridyle Lagoon, Gentle 7713.

Pleurothallis tikalensis Correll & C. Schweinfurth, sp. nov.

Fig. 52.

Herba mediocris, caespitosa. Radices fibrosae, numerosae. Caules vaginis

pluribus laxis omnino celati. Folium erectum, oblongum vel ovato-oblongum,

acutum, basi sessili rotundatum vel subcuneatum. Inflorescentia solitaria, ra-

cemosa, quam folium multo brevior, dense multiflora, vagina scariosa conduplicata

sustentata. Flores perparvi, bilabiati. Sepalum dorsale lineari-lanceolatum, acu-

FIG. 52. Pleurothallis tikalensis. Plant (X 1A)', 1, sepals, spread out, and

column ( X 5) ; 2, lip, spread out ( X 5); 3, petal ( X 5). Drawn by Vivien Frazier.

191

192 FIELDIANA: BOTANY, VOLUME 31

turn vel acuminatum. Sepala lateralia aequilonga, in laminam late ovatam biden-

tatam connata. Petala minuta, rhombico-spathulata, parte superiore irregulariter

lacerato-dentata. Labellum simplex, expansum ovatum, apice rotandatum, basi

cuneatum, prope basim utrinque triangulari-dilatatum. Columna brevis, in pedem

conspicuum productum.

Plant medium-sized, cespitose, about 15 cm. high. Roots fibrous, flexuous,

numerous. Stem 4-4.5 cm. high, entirely concealed by several loose scarious eva-

nescent sheaths. Leaf solitary, erect, oblong or ovate-oblong, acute, rounded or

subcuneate at the sessile base, up to 11 cm. long and 2.7 cm. wide, subcoriaceous.

Inflorescence solitary, racemose, erect, much shorter than the leaf, about 3.5-4.5

cm. high, densely many-flowered to the base, subtended by a scarious conduplicate

sheath up to 1.4 cm. long. Floral bracts minute, infundibuliform, scarious. Flowers

very small, bilabiate. Dorsal sepal linear-lanceolate, acute or acuminate, 3-nerved,

about 9.2 mm. long and 2.1 mm. wide. Lateral sepals connate into a broadly ovate

bifid lamina about 9.2 mm. long and 7.2 mm. wide when expanded, 6-nerved, with

the tips acute and mucronate. Petals very small, rhombic-spatulate from a nar-

row basal third, 1-nerved, the apical portion oblique, acuminate and irregularly

lacerate-dentate, about 4.2 mm. long and 3 mm. wide above. Lip simple, carnose,

longitudinally concave with the lower sides erect, ovate when expanded, with the

cuneate base gradually dilated on each side into an obtusely triangular point and

with a pair of indistinct fleshy ridges on each side near the middle, obtuse, minutely

denticulate on the anterior margins, about 5 mm. long and 2.1 mm. wide in natural

position. Column small, about 3 mm. high, gradually wing-dilated toward the

subtruncate apex, produced into a prominent fleshy foot.

This species is allied to Pleurothallis angustisepala Ames & Cor-

rell, but it has dissimilar petals and lip. The vegetative appearance

recalls P. pardipes Reichb. f.

Guatemala. — Pete*n: Tikal National Park, Tikal, bordering Aguado

Seca, epiphyte on tree trunk, flowers green and maroon, October 14,

1959, Elias Contreras 280 (type in Lundell Herbarium).

Pleurothallis tribuloides (Sw.) Lindl.; Ames & Correll, p. 256.

Guatemala. — Pete"n: About 400 m. south of Dolores, Contreras

2838. About 5 km. south southeast of Dolores, Contreras 3031.

British Honduras. — Toledo: Beyond San Antonio, Gentle 6587.

Joe Taylor Creek, Gentle 7407, 7777.

Pleurothallis yucatanensis Ames & C. Schweinf. ; Ames &

Correll, p. 260.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 156.

British Honduras. — Toledo: Punta-Gorda Seven Hills Road, as-

cending Rio Grande, Gentle 4994.

CORRELL: ORCHIDS OF GUATEMALA 193

Malaxis Parthonii Morren; Ames & Correll, p. 272.

Guatemala. — Pete"n: Tikal National Park, Tikal, Contreras 1840.

Santo Toribio, Contreras 2749. East of Dolores, Contreras 3092. Tikal

National Park, on Remate Road, Lundell 17097. Tikal National

Park, on Uaxactun, Lundell 17131. — Santa Rosa: Cenaguilla, Heyde

& Lux 3866.

Epidendrum alatum Batem.; Ames & Correll, p. 300.

Guatemala. — Pete"n: Tikal National Park, Tikal, on top of Tem-

ple III, Contreras 921. Tikal National Park, Bajo de Santa F£, Con-

treras 1262.

British Honduras. — Toledo: Rio Grande, Gentle 4695. Near Ja-

cinto Creek, Gentle 5571, 5578. Between Punta Gorda and Moho

River, Gentle 6779. Cero Hill, Gentle 7059.— Indefinite: Mountain

Pine Ridge, H. H. Bartlett 11864.

Epidendrum anceps Jacq.; Ames & Correll, p. 303, fig. 88.

Guatemala. — Pete"n: 1 km. west of Dolores, Contreras 2086. About

700 m. south of Dolores on old Machaquila Road, Contreras 2247.

Epidendrum aromaticum Batem.; Ames & Correll, p. 306.

Guatemala. — Quiche^ Sacabaja, Heyde & Lux 3512.

Epidendrum atropurpureum Willd.; Ames & Correll, p. 307,

fig. 89.

A rather rare species in Mexico (Chiapas, Mt. Ovando, near Es-

cuintla, Matuda 6092) becoming frequent southward in northern

South America.

Guatemala. — Quezaltenango: San Carlos Miramar, Tonduz &

Rojas 180.

Epidendrum Boothii (Lindl.) L. Wms.; Ames & Correll, p. 309.

Guatemala. — Pete'n: Tikal National Park, Tikal, on Temple of

the Inscriptions, Contreras 77. Tikal National Park, Tikal, in ra-

monal covering the ruins, Contreras 144. Tikal National Park, on

Remate Road bordering Aguada El Charco, Contreras 1408. Santo

Toribio, Contreras 2679.

British Honduras. — Toledo: Rio Grande, Gentle 4741. Near Con-

demn Branch Pine Ridge, Gentle 5381. Near San Antonio, Gentle

5447. Crique Queva, Gentle 6020. Edwards Road beyond Colum-

bia, Gentle 6245. Between Punta Gorda-San Antonio Road and

Moho River, Gentle 6750. Near Orange Point, Gentle 7477. Be-

tween Punta Gorda and Paslow, Gentle 7809.

194 FIELDIANA: BOTANY, VOLUME 31

Epidendrum bractescens LindL; Ames & Correll, p. 311.

Guatemala. — Pete'n: Tikal National Park, Bajo de Santa FC", Con-

treras 549. Remate, on Tikal Road, Contreras 824. On Santa Ana

trail, about 7 km. east southeast of Lake Pete'n, Contreras 2032. Tikal

National Park, Bajo de Santa F£, Arroyo Corriental, on Aguada Ter-

minos road, Lundell 15876.

British Honduras. — Toledo: Bolo Camp, upper reach of Golden

Stream, Gentle 4547. Near Condemn Branch Pine Ridge, Gentle 5290.

Near Columbia, Gentle 6082. Chavarrias Road, Resemederes, across

Columbia River, Gentle 6284.

Epidendrum ciliare L.; Ames & Correll, p. 318, fig. 91.

Guatemala. — Pete'n: About 3 km. south of Dolores, Contreras

3146. In forest bordering Dolores, Contreras 3190.

Epidendrum Clowesii Batem. ex Lindl.; Ames & Correll, p. 320.

Guatemala. — Suchitepe"quez, vicinity of Mazatenango, W. R.

Maxon & R. Hay 3619.

Epidendrum cobanense Ames & Schltr.; Ames & Correll, p. 322,

fig. 79.

Guatemala. — Santa Rosa: Santa Rosa, Heyde & Lux 4605.

Epidendrum cochlea turn L.; Ames & Correll, p. 323, fig. 92.

Guatemala.- — Pete'n: Tikal National Park, Tikal, growing on Tem-

ple of the Inscriptions, Contreras 515. West of Remate, bordering

lake, Contreras 983. About 2 km. 500 m. south of Dolores, Contreras

2357. Santo Toribio, Contreras 2691. Lacandon, Contreras 3463.

Tikal National Park, Tikal, in Aguada Corriental, Lundell 15497.

Tikal National Park, Tikal, in ramonal covering the ruins, Lundell

15501. Tikal National Park, Bajo de Santa F4, along Rio Holmul,

Lundell 15591.

British Honduras. — Corozal: Edge of New River, Pueblo Nuevo,

Gentle 469. — El Cayo: Cave Branch Section, Humming Bird High-

way, Gentle 9059.— Toledo: Rio Grande, Gentle 4717. Near Mof-

fredye Creek, near San Antonio, Gentle 5330. Columbia Road, Gen-

tle 6126. Between Orange Point and Moho River, Gentle 7627.

Beyond Mafridyle Lagoon, Gentle 7712.

Epidendrum condylochilum Lehm. & Kranzl.; Ames & Cor-

rell, p. 325.

Guatemala. — Guatemala: Fiscal, C. C. Deam 6198. — Pete'n: Re-

mate, Contreras 847, 980.

CORRELL: ORCHIDS OF GUATEMALA 195

Epidendrum cristatum Ruiz & Pavon; Ames & Correll, p. 327.

Guatemala. — Pete"n: Tikal National Park, Tikal, growing on top

of temple in "H" Group, Contreras 286, 1043.

British Honduras. — Toledo: Beyond Central Camp, Edwards

Road beyond Columbia, Gentle 7342.

Epidendrum cystosum Ames Bot. Mus. Leafl. Harvard Univ.

2: 106, t. 1934.

Epiphytic in forests, up to about 1500 m. alt. Rare in British

Honduras and Honduras.

Plant about 13 cm. tall. Roots coarsely fibrous, smooth, longitudinally sulcate,

whitish, spreading. Stems cespitose, erect, 4.5-5.5 cm. long, bearing about 6 leaves,

complanate. Leaves 4.5-11 cm. long, 3.5-6 mm. wide, obliquely ascending, the

uppermost one exceeding the tip of the raceme, chartaceous when dry, narrowly

oblong or linear-lanceolate, tapering gradually to the acute to acuminate-attenuate

apex, narrowed toward the base and jointed to the tubular complanate sheaths that

are about 15 mm. long, and 2-3 mm. in greatest diameter. Peduncle terminal, to

the tip of the rachis about 4.5 cm. long, few-flowered, usually one flower open at a

time with the succeeding flower in bud. Peduncular sheaths about 3, closely ap-

pressed, imbricating, complanate, acute, the tip of the uppermost one extending

beyond the lowermost floral bract, the lowermost sheath about 2 cm. long. Rachis

of the raceme 4-7 mm. long. Bracts of the raceme infundibuliform, about 2.5 mm.

long, acute, crowded. Lateral sepals lanceolate, 9 mm. long, 3 mm. wide, obliquely

porrect, very slightly coherent at the base, acuminate, somewhat conduplicate at

the acutish apex, 3-nerved with the mid-nerve carinate on the outer surface. Dor-

sal sepal similar to lateral sepals, 3-nerved, reflexed, at the base lightly coherent

with the lateral sepals. Petals linear, 8.5 mm. long, spreading, tapering gradually

to an acute tip, 1-nerved. Lip about 8 mm. long, adnate below the middle to the

column, free portion 5 mm. long and 5 mm. wide, elliptical to suborbicular, acute,

very fleshy, rigid, somewhat deflexed, lightly concave with a conspicuous elongated

callus extending along the mid-nerve which is very prominent from the middle

almost to the apex, this callus being joined on each side by an elongated much

thickened simple or branching costa and with a free intramarginal costa, simple or

branched on each side extending from the base of the free portion almost to the

apex. Column 3.5 mm. long, completely adnate to the lip, minutely denticulate

on the margin of the clinandrium. Pedicellate ovary 11 mm. long, with a pouchlike

swelling where the ovary joins the base of the lip.

This species is new to our region. It differs from the distantly

related E. physodes in its obliquely ascending chartaceous leaves,

shorter stem, and differently shaped lip.

British Honduras. — Toledo: Between Orange Point and Moho

River, Gentle 7660, 7658, 7659. Jacinto Creek, Rio Grande, Gentle

4898. Punta Gorda-Seven Hills Road, ascending Rio Grande, Gen-

tle 4995.

196 FIELDIANA: BOTANY, VOLUME 31

Epidendrum difforme Jacq.; Ames & Correll, p. 328, fig. 93.

Guatemala. — Pete"n: Dolores, between kms. 81 and 82, Contreras

2541. 400 m.-2 km. south of Dolores, Contreras 2620, 2859.— Saca-

tepe"quez: Villa Nueva, Heyde & Lux 4608.

British Honduras. — El Cayo: In advanced forest, limestone val-

ley, Lundell 6424.

Epidendrum diffusum Sw.; Ames & Correll, p. 331.

Guatemala. — Pete"n: Tikal National Park, Tikal, growing on top

of Temple III, Temple IV and temple in "H" Group, Contreras 99,

160, 488. Tikal National Park, Tikal, in ramonal covering the ruins,

Contreras 491; Lundell 15438. Tikal National Park, Bajo de Santa

Fe", Contreras 541. Tikal National Park, Aguada Naranjal, Contre-

ras 1497.

Epidendrum equitantifolium Ames Sched. Orch. 4: 39. 1923.

Epiphytic on ridges and hills in forests at low elevations, up to

about 1000 m. alt. Uncommon in Mexico and in Central America

south to Panama.

Epiphytic or pendent cespitose plants up to about 4 dm. tall or long. Stems

slender, leafy, ancipitous. Leaves 8-27 cm. long, 4-12 mm. wide, equitant, ancipi-

tous, linear-lanceolate, acute to acuminate, fleshy. Inflorescence borne on an an-

cipitous winged peduncle up to 25 cm. long and simulating a leaf, 1- to few-flowered,

subcapitate, subtended by a reduced leaf-like bract. Flowers small, dull-colored.

Sepals 12-17 mm. long, 1.5-2.5 mm. wide, linear to linear-oblong or elliptic, acute

to obtuse. Petals 8-13 mm. long, 1-2 mm. wide, linear-oblanceolate to linear-

elliptic. Lip unguiculate; the claw adnate to the column for about half its length;

lamina 7-9 mm. long, 5-6 mm. wide, 3-lobed, provided with a callus on the claw

at the base; lateral lobes semiorbicular; terminal lobe lanceolate to triangular-

lanceolate; disc concave or cochleate.

This species was not included in Ames and Correll (1952-1953).

It is included here on the basis of its occurrence in British Honduras.

British Honduras. — Toledo: On tree, in high ridge, near Union

Camp, Edwards Road, about 25 miles beyond Columbia, Gentle 6497.

Epidendrum ibaguense H.B.K.; Ames & Correll, p. 333.

Guatemala. — Alta Verapaz: Along National Route No. 5, about

1 mile south of Coban, R. M. King 3307. — Pete"n: Dolores, on Macha-

quila Road, between kms. 85-86, Contreras 3131.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, Gentle 4282. Between Rancho and Cockscomb, Gentle 4346.

CORRELL: ORCHIDS OF GUATEMALA 197

Epidendrum imatophyllum Lindl.; Ames & Correll, p. 335.

Guatemala.— Pet^n: Uaxactun, H. H. Bartlett 12669. On Falla-

bon Road, about 9 km. east of Remate, Contreras 892. Lacandon,

Contreras 3315. Lacandon, bordering Sibal, Contreras 3445. Tikal

National Park, Tikal, in ramonal covering the ruins, Lundell 15922,

16880. Tikal National Park, Tikal, bordering airfield, Lundell 15958.

British Honduras. — Toledo: Botan Creek, Rio Grande, Gentle

4622. Temash River, Gentle 5218. Near Jacinto Creek, Gentle 5566.

Epidendrum ionophlebium Reichb. f . ; Ames & Correll, p. 336.

Guatemala. — Escuintla: Escuintla, J. D. Smith 2248. — Suchite-

pe"quez: Mazatenango, Rio Sis, J. D. Smith 2644.

Epidendrum isomerum Schltr. ; Ames & Correll, p. 338.

Guatemala. — Pete*n: Dolores, Contreras 2212. About 400 m. south

of Dolores, Contreras 2843.

British Honduras. — El Cayo: Humming Bird Highway, Gentle

8586. 41 Miles Section, Belize-Cayo Road, Gentle 9680.— Stann

Creek: Humming Bird Highway, Gentle 8653. — Toledo: Edwards

Road beyond Columbia, Gentle 6384, 6434, 7222. Near San An-

tonio, Gentle 7540.

Epidendrum neurosum Ames; Ames & Correll, p. 345.

This species is new to British Honduras.

British Honduras. — Toledo: On tree, in high ridge, near Union

Camp, Edwards Road beyond Columbia, Gentle 6495.

Epidendrum nocturnum Jacq.; Ames & Correll, p. 346, fig. 96.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa Pe", Con-

treras 576, 1283. Tikal National Park, Arroyo Corriental, between

Aguada Naranjal and Agua Perdida, Contreras 1904. Tikal National

Park, Bajo de Santa Fe", Arroyo Corriental, on Aguada Terminos

Road, Lundell 16497.

British Honduras. — Stann Creek: Stann Creek Valley, Mt. Cow

Ridge, Gentle 3561. — Toledo: Near Jenkins Creek, north of Monkey

River, Gentle 4082. Rio Grande, Gentle 4846. Near Jacinto Creek,

9 Miles, Punta Gorda-San Antonio Road, Gentle 4970. Near San

Antonio, Gentle 5507. Near Columbia, Gentle 6078. Chavarrias

Road, across Columbia River, Gentle 6282. Beyond Pate's Camp,

Edwards Road beyond Columbia, Gentle 7129. Lower Joe Taylor

Creek, Gentle 7498. Mafridyle Creek near San Antonio, Gentle 7563.

198 FIELDIANA: BOTANY, VOLUME 31

Epidendrum oncidioides Lindl.; Ames & Correll, p. 350.

Guatemala. — Pete'n: 5-6 km. east of Dos Lagunas, Contreras 1732,

1753.

British Honduras. — Belize: Colonel English pine ridge, Belize-

Cayo Road, Gentle 9740. — Toledo: Between Monkey River and

Cockscomb, Gentle 4235.

Epidendrum paniculatum Ruiz & Pavon; Ames & Correll,

p. 352.

Guatemala. — Pete'n: Bordering Bajo de Santa Fe\ east of Tikal

National Park, Contreras 1446.

British Honduras. — Toledo: near Mafridyle Creek, Gentle 7566.

Epidendrum Parkinsonianum Hook.; Ames & Correll, p. 354.

Guatemala. — Quiche" :F. C.Lehmann 1570.

Epidendrum pentotis Reichb. f.; Ames & Correll, p. 355.

This species is new to British Honduras.

British Honduras. — Toledo: On tree, in high ridge, at base of hill

near Union Camp, Edwards Road beyond Columbia, Gentle 6475.

Epidendrum physodes Reichb. f.; Ames & Correll, p. 356.

This species is new to British Honduras.

Guatemala. — Pete'n: Tikal National Park, about 9 km. northeast

of Tikal, Contreras 1480. Julec, between Santo Toribio and Santa

Ana, Contreras 2661. Canchacan, Lundell 16454.

British Honduras. — Toledo: On tree, in Give and Take Ridge,

Rio Grande, Gentle 4878.

Epidendrum polyanthum Lindl.; Ames & Correll, p. 357.

Guatemala. — Pete'n: Tikal National Park, Aguada San Geronimo,

Contreras 1502. Dos Lagunas, on Uaxactun trail, Contreras 1567.

Tikal National Park, on Remate Road, Lundell 16130. — Santa Rosa:

Estanzuela, Heyde & Lux 4275.

British Honduras. — Toledo: Rio Grande, Gentle 4745.

Epidendrum pygmaeum Hook.; Ames & Correll, p. 361, fig. 98.

Guatemala. — Pete'n: About 400 m. south of Dolores, Contreras

2860. Dolores, on Machaquila Road, between kms. 85-86, Contre-

ras 3132.

British Honduras. — El Cayo: 31 Miles Section, Humming Bird

Highway, Gentle 8922. — Stann Creek: Dry Creek, Humming

CORRELL: ORCHIDS OF GUATEMALA 199

Bird Highway, Gentle 8398. — Toledo: Between Monkey River and

Cockscomb, Gentle 4273. Edwards Road beyond Columbia, Gentle

6368, 6454, 7219.

Epidendrum radiatum Lindl.;Ames & Correll, p. 362, fig. 99.

Guatemala.- — Guatemala: San Rafael, F. C. Lehmann 1848.—

Pete"n: Tikal National Park, Tikal, growing on top of Temple IV,

Contreras 127. Tikal National Park, Tikal, in ramonal covering the

ruins, Contreras 138, 490; Lundell 17176. Tikal National Park, Bajo

de Santa Fa", Contreras 546, 602. About 3 km. north northwest of

Dolores, Contreras 3080. Uaxactun, south of ruins in ramonal, Lun-

dell 16635.

Epidendrum rigidum Jacq.; Ames & Correll, p. 369, fig. 100.

Guatemala.- — Pete"n: Tikal National Park, Tikal, on top of tem-

ple in "H" Group and Temple IV, Contreras 372, 1054, 1187. Tikal

National Park, Bajo de Santa Fa", Contreras 599. Tikal National

Park, Tikal, in ramonal covering the ruins, Contreras 1199, 1211,

1985; Lundell 15437, 16084, 16176. About 2^ km. south of Do-

lores, Contreras 2356. About 2 km. east of Poptun, Lundell 16418.

About 4 km. south southeast of Dolores, Contreras 2481. Dolores,

Contreras 3004.

British Honduras. — El Cayo: 47 Miles Section, Humming Bird

Highway, Gentle 8853. 32 Miles Section, Humming Bird Highway,

Gentle 9163. 41 Miles Section, Belize-Cayo Road, Gentle 9679.—

Toledo: Botan Creek, Rio Grande, Gentle 4644. Bank of Temash

River, Gentle 4655. Crique Queva, Gentle 6027. Edwards Road be-

yond Columbia, Gentle 6299, 6379. Feeders Road, 14 Miles, San

Antonio-Punta Gorda Road, Gentle 6698. Joe Taylor Creek, Gen-

tle 7496.

Epidendrum sculptum Reichb. f. Bonpl. 2: 89. 1854. Williams

in Woodson & Schery, Fl. of Panama (Ann. Mo. Bot. Gard. 33: 306

(374) 1946).

Epiphytic in hills and low mountain forests. Infrequent in Brit-

ish Honduras, Honduras, Costa Rica, Panama and northern South

America.

Plants repent or pendent, up to about 5 dm. tall or long, usually much smaller.

Stems simple or branched, densely leaved, covered with the chartaceous leaf-sheaths

or becoming naked with age. Leaves spreading horizontally, 2-4.5 cm. long, 1-

1.8 cm. wide, oblong to oblong-lanceolate, obtuse, retuse, coriaceous. Inflorescence

with 1 to 3, usually 2, terminal flowers that are subtended by spathaceous bracts.

Flowers leathery, green or yellowish green. Dorsal sepal 1-1.4 cm. long and 2.5-

200 FIELDIANA: BOTANY, VOLUME 31

3 mm. wide, lanceolate to oblong-lanceolate, obtuse. Lateral sepals 1-1.5 cm. long,

about 3 mm. wide, oblong-lanceolate, oblique, acute. Petals 9-14 mm. long, 2-3

mm. wide, linear-elliptic to narrowly oblanceolate, oblique, obtuse. Lip unguicu-

late; claw adnate to the column; lamina 9-13 mm. long, 5.5-7 mm. wide, ovate-

lanceolate, acute, 3-lobed; lateral lobes small, rounded, reaching to about the middle

of the lip; mid-lobe lanceolate; disc provided with two callus-like thickenings at the

base on the lateral lobes and with two longitudinal ridges along the median line

and reaching to about the middle of the lip.

This species was not included in Ames and Correll (1952-1953).

It is included here on the basis of its occurrence in British Honduras.

British Honduras. — Toledo: Punta Gorda-Seven Hills Road, as-

cending Rio Grande, Gentle 5029. Near San Antonio, Gentle 5431,

7536. Between Machaca and Rio Grande, Gentle 6937.

Epidendrum selligerum Batem. ex Lindl.; Ames & Correll,

p. 374.

Guatemala. — Quiche": Sacabaja, Heyde & Lux 3508.

Epidendrum Stamfordianum Batem.; Ames & Correll, p. 380,

fig. 105.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Lundell 15566, 16881. Tikal National Park, Dos

Aguadas, Lundell 15600.

Epidendrum stenopetalum Hook.; Ames & Correll, p. 382.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 1984, 345.

British Honduras. — Toledo: Bolo Camp, upper reach of Golden

Stream, Gentle 4503. Jacinto Creek, Rio Grande, Gentle 4937. Punta

Gorda-Seven Hills Road, ascending Rio Grande, Gentle 5035. Near

Moffredye Lagoon, Gentle 5299. Near San Antonio, Gentle 5424,

5485. Edwards Road beyond Columbia, Gentle 6178, 6382, 7013.

Epidendrum strobiliferum Reichb. f.; Ames & Correll, p. 382,

fig. 106.

British Honduras. — Toledo: Columbia Road, Gentle 6130. Lower

Joe Taylor Creek, Gentle 7502. Between Orange Point and Moho

River, Gentle 7654.

Epidendrum varicosum Batem. ex Lindl.; Ames & Correll,

p. 389.

Guatemala. — Santa Rosa: Santa Rosa, Heyde & Lux 4637.

CORRELL: ORCHIDS OF GUATEMALA 201

Diacrium bilamellatum (Reichb. f.) Hemsl.; Ames & Correll,

p. 405, fig. 108.

British Honduras. — Toledo: Near Condemn Branch Creek, Gen-

tle 5553. Between Punta Gorda and Joe Taylor Creek, Gentle 6794.

Feeder Road leading to Big Fall, Gentle 7048. Mafridyle Creek,

Gentle 7590. Between Condemned Branch Pine Ridge and Mafridyle

Lagoon, Gentle 7685.

Cattleya aurantiaca (Batem. ex Lindl.) P. N. Don; Ames &

Correll, p. 407.

Guatemala. — Amatitlan: Lake Amatitlan, W. A. Kellerman 4544.

—Guatemala: El Naranjo, G. Salas 206. Guatemala, J. D. Smith

2252. — Sacatepe"quez : Villa Nueva, Heyde & Lux 4598.

Cattleya Skinneri Batem.; Ames & Correll, p. 412, fig. 111.

British Honduras. — El Cayo: San Agustin, on granite banks of

Rio Frio,Lundell 6701.

Laelia Digbyana Benth.; Ames & Correll, p. 415, fig. 112.

This species was included in the Orchids of Guatemala since it is

known from Mexico (Yucatan) and British Honduras, southward.

The specimens cited here are the first we have seen for Guatemala.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa F4, Con-

treras 457, 556. About 10 km. west of Dos Lagunas, Contreras 1769.

Laelia tibicinis (Batem. ex Lindl.) L. Wms.; Ames & Correll,

p. 419.

British Honduras. — Belize: Turneffe Island, Calabash Section,

F. E. Engler 42-24.— Toledo: Botan Creek, Rio Grande, Gentle 4641,

4642, 4643. Temash River, Gentle 4656. Along bank of Jacinto

Creek, Gentle 5575, 5576.

Brassavola cucullata (L.) R. Br.; Ames & Correll, p. 422.

Guatemala. — Pete"n: West of Remate, bordering the lake, Con-

treras 982, 1028.

Brassavola nodosa (L.) Lindl.; Ames & Correll, p. 423, fig. 114.

British Honduras. — Stann Creek: Stann Creek-Sittee Road, Gen-

tle 7987.— Toledo: near Columbia, Gentle 6391.

Meiracy Ilium trinasutum Reichb. f.; Ames & Correll, p. 425,

fig. 115.

Guatemala.— Amatitlan: Lake Amatitlan, W. A. Kellerman 5035.

202 FIELDIANA: BOTANY, VOLUME 31

Scaphyglottis Behrii (Reichb. f.) Benth. & Hook.; Ames & Cor-

rell, p. 430, fig. 116.

British Honduras. — El Cayo: Vaca, Gentle 2547. Humming Bird

Highway, Gentle 8596, 9067.— Stann Creek: Stann Creek -Sittee Road,

Gentle 7928. — Toledo: Between Rancho Chico and Cockscomb, Mon-

key River, Gentle 4340. Beyond Columbia, Gentle 6165. Near Cen-

tral Camp, Edwards Road beyond Columbia, Gentle 7282. Mafridyle

Creek, Gentle 7592.

Scaphyglottis cuneata Schltr.; Ames & Correll, p. 434.

Guatemala. — Pete"n: Tikal National Park, Tikal, growing on top

of Temple IV, Contreras 1794. Tikal National Park, on Remate

Road, Lundell 17121.

British Honduras. — Stann Creek: Between Silkgrass and Sapon

Road, Gentle 8110. Eve's Pine Ridge, Gentle 8505.— Toledo: Cow

Pen, near Monkey River, Gentle 4077. Between Monkey River and

Cockscomb, Gentle 4295. Near San Antonio, Gentle 5470. Columbia

Road, Gentle 6110. Near Big Falls, Feeders Road, 14 Miles, San An-

tonio-Pun ta Gorda Road, Gentle 6735. Mafridyle Creek, Gentle 7610.

Scaphyglottis Lindeniana (A. Rich. & Gal.) L. Wms.; Ames &

Correll, p. 436.

British Honduras. — Toledo: Beyond Union Camp, Edwards Road,

beyond Columbia, Gentle 6517.

Scaphyglottis livida (Lindl.) Schltr.; Ames & Correll, p. 438.

Guatemala. — Pete"n: About 3 km. south of Dolores, Contreras

3145.

Scaphyglottis major (C. Schweinf.) Correll, comb. nov.

Scaphyglottis Wercklei Schltr. var. major C. Schweinf. Bot. Mus.

Leafl. Harvard Univ. 4: 117. 1937. S. cuneata var. major (C.

Schweinf.) C. Schweinf. in Correll, Lloydia 10: 212. 1947; Ames &

Correll, Orchids of Guatemala (Fieldiana: Botany, 26 (2): 435. 1953).

This species differs in several respects from S. cuneata to which it

was referred in our work on the orchids of Guatemala. Although the

lips are quite similar in shape, the lip of this species has no callus

whatsoever on its disc and the apex of the lip is rather prominently

lobulate. Also, the plants, especially the leaves, are much larger than

in that species. No plants that could be considered as intermediate

have been seen.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa EC",

Arroyo Corriental, on Aguada Terminos road, Contreras 334. Tikal

CORRELL: ORCHIDS OF GUATEMALA 203

National Park, Tikal, growing on top of temple in "H" Group, Con-

treras 383, 1822.

Scaphyglottis minutiflora Ames & Correll, p. 440, fig. 118.

In addition to being known from Guatemala, British Honduras,

and Panama (P. H. Allen 2258), specimens in the United States

National Herbarium have been seen from Honduras (Dept. Cortes,

Santa Cruz de Yozoa, J. B. Edwards 618) and Costa Rica (Prov.

Cartago, vie. of Pejivalle, P. C. Standley & J. Valerio 47251 bis).

Guatemala. — Alta Verapaz: Cubilqiiitz, Tuerckheim II 526, II

1062, 8758. Secanquim, near Finca Sepacuite', 0. F. Cook & R. F.

Griggs 267. Cacao, Trece Aguas, F. L. Lewton 268. — Pete'n: Dolores,

between kms. 85-87, west of Machaquila Road, Contreras 2941, 2974,

3141.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, Gentle 4265. Near Jacinto Creek, Gentle 4984. Chavarrias

Road, Resemederes, across Columbia River, Gentle 6281. Edwards

Road beyond Columbia, Gentle 6408, 6533.

Ponera stria ta Lindl.; Ames & Correll, p. 445, fig. 119.

Guatemala. — Pete'n: Tikal National Park, Tikal, growing on top

of Temple III and temple in "H" Group, Contreras 487, 1783. Tikal

National Park, Bajo de Santa FC", Contreras 536. Tikal National

Park, growing on top of Temple I, Lundell 15572.

British Honduras. — Toledo: Punta Gorda-Seven Hills Road, as-

cending Rio Grande, Gentle 5012. Beyond Carmelita Camp, Edwards

Road beyond Columbia, Gentle 7213.

Jacquiniella globosa (Jacq.) Schltr.; Ames & Correll, p. 448,

fig. 120.

This species is new to British Honduras.

Guatemala. — Pete'n: Dolores, bordering the village clearing, Con-

treras 2202. About 4 km. south southeast of Dolores, Contreras 2488.

On Rio Mopan Trail, about 3 km. southeast of Dolores, Contreras

2567. About 3 km. south of Dolores, Contreras 3144.

British Honduras. — Toledo: Near Big Falls, Feeders Road, 14

Miles, San Antonio-Punta Gorda Road, Gentle 6743. Beyond Co-

lumbia, Gentle 6134, 6286-B.

Isochilus linearis (Jacq.) R. Br.; Ames & Correll, p. 450.

Guatemala. — Baja Verapaz: near Santa Rosa, W. R. Maxon &

R. Hay 3374.— Santa Rosa: Santa Rosa, Heyde & Lux 3488.

204 FIELD IANA: BOTANY, VOLUME 31

Isochilus linearis var. carnosiflorus (Lindl.) Correll; Ames &

Correll, p. 451.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa Fe", Con-

treras 458, 586. About 10 km. west-northwest of Dos Lagunas, Contre-

ras 1712. Tikal National Park, in ramonal covering the ruins,

Contreras 1823. Tikal National Park, Bajo del Hormiguero, 3 leagues

south of Tikal on old Remate Road, Lundell 15536. Tikal National

Park, Bajo de Santa Fe, bordering Aguada Pucte, Lundell 17011.

Tikal National Park, Bajo del Hormiguero, Lundell 17191.

British Honduras. — Toledo: Punta Gorda-Seven Hills Road, as-

cending Rio Grande, Gentle 5011. Temash River, Gentle 5205. Be-

yond Columbia, Gentle 6131, 6412. Near San Antonio, Gentle 7541.

Mafridyle Creek, Gentle 7567.

Arpophyllum spicatum La Llave & Lex.; Ames & Correll,

p. 457.

This species is new to British Honduras.

Guatemala. — Sacatep4quez : Embaulada, Heyde & Lux 4597.

British Honduras. — Toledo: On tree, in high ridge, on hill top,

beyond Manga Camp, Edwards Road beyond Columbia, Gentle 6508.

Polystachya cerea Lindl.; Ames & Correll, p. 466, fig. 126.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa Fe",

Arroyo Corriental, on Aguada Terminos road, bordering Aguada

Pucte, Contreras 267. On Rio Mopan trail, about 3 km. east of

Dolores, Contreras 2826.

British Honduras. — Toledo: Near Mafridyle Lagoon, Gentle 7823.

—Indefinite: Honey Camp, Lundell 664.

Polystachya clavata Lindl.; Ames & Correll, p. 468.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 203. Tikal National Park, Bajo de Santa

F£, Contreras 267 A, 548. Tikal National Park, Bajo del Hormiguero,

3 leagues south of Tikal on old Remate road, Lundell 15568.

British Honduras.— El Cayo: Lundell 6913.— Stann Creek: 19

Mile Section, Stann Creek Valley, Gentle 9340.— Toledo: Near Cow

Pen, Monkey River, Gentle 4119. Rio Grande, Gentle 4790, 4877.

Near Jacinto Creek, 9 Miles, Punta Gorda-San Antonio Road, Gen-

tle 4969. Near San Antonio, Gentle 5432, 7529, 7795. Between Con-

demned Branch Pine Ridge and Moffredye Lagoon, Gentle 6059.

Chavarrias Road, Resemederes, across Columbia River, Gentle 6280,

6283. Middle Camp, Edwards Road beyond Columbia, Gentle 6411

CORRELL: ORCHIDS OF GUATEMALA 205

p.p. Between Punta Gorda and Moho River, Gentle 6799. Near

Mafridyle Creek, Gentle 7593. — Indefinite: 2 miles south of Maskall,

H. O'Neill 8342.

Polystachya masayensis Reichb. f.; Ames & Correll, p. 470.

This species is new to British Honduras.

British Honduras.— Toledo: On tree, in high ridge, on hill top,

Middle Camp, Edwards Road beyond Columbia, Gentle 6411 p.p.

Galeandra Baueri Lindl. ; Ames & Correll, p. 472, fig. 128.

British Honduras. — Toledo: Rio Grande, Gentle 4841. — Indefinite:

On pines in open places, W. A. Schipp 590.

Bletia purpurea (Lam.) DC.; Ames & Correll, p. 486, fig. 133.

Guatemala. — Alta Verapaz: Between Cajabon and Lanquin, G. P.

Goll 119.— Guatemala: Pinula, J. D. Smith 2251.— Pet^n: West of

Dolores, Contreras 2128. — Quiche": Sacabaja, Heyde & Lux 3513, 4607.

British Honduras. — Belize: Colonel English Pine Ridge, Belize-

Cayon Road, Gentle 9518, 9717.— El Cayo: 37 Miles Section, Hum-

ming Bird Highway, Gentle 8962. — Toledo: Temash River, Gentle

4661. Condemn Branch Pine Ridge, Gentle 5307. Edwards Road

beyond Columbia, Gentle 6214.

Bletia reflexa Lindl. ; Ames & Correll, p. 488.

Guatemala. — Sacatepe"quez: Volcan de Agua, W. C. Shannon 3669.

Bletia tenuifolia Ames & C. Schweinf., Bot. Mus. Leafl., Har-

vard Univ. No. 10: 6, f. 1933.

Terrestrial in forests of ridges. Rare in Mexico and British Hon-

duras.

Plant scapose, up to about 5 dm. tall, with coarse flexuous fibrous roots and a

subglobose rugose shining corm about 1.2 cm. long. Leaves 4 or 5 in a cluster aris-

ing from base of plant, very narrowly linear, articulated to convolute sheaths that

are in turn loosely enveloped by several imbricating nervose scarious leaf-bearing

sheaths; leaf-blades 15-34 cm. long, up to 3 mm. wide, complicate-acuminate,

3-nerved, erect-spreading. Scape somewhat surpassing the leaves, slender, con-

cealed at base by 3 short imbricating nervose sheaths and above by only 3 remote

appressed tubular sheaths. Rachis commonly subpaniculate with a single basal

branch, up to 15 cm. long. Floral bracts lanceolate, long-acuminate, scarious, the

lowermost about 5.5 mm. long. Raceme loose, with the flowers 1 cm. or more dis-

tant. Dorsal sepal elliptic-lanceolate, acute, 1 cm. long, 4 mm. wide, 5-nerved.

Petals asymmetrically elliptic, narrowed to an obtuse to subacute apex, about

9 mm. long and 4.5 mm. wide, the margins of the upper half irregular, 3-nerved

with the side nerves branching. Lip ovate in outline, 3-lobed near the apex, articu-

lated to the short column-foot, 8 mm. long and about 6 mm. wide, broadly cuneate

206 FIELDIANA: BOTANY, VOLUME 31

at base; lateral lobes semiovate, 6.2 mm. long, lobulate at their rounded apex; mid-

lobe much smaller than the lateral lobes, suborbicular-quadrate, about 2 mm. long

and wide, shallowly retuse and apiculate, the margins strongly undulate and lobu-

late; disc traversed by 5 parallel keels that are highest in the middle of the lip and

become undulate near their apex; the 3 middle keels extend about to the middle

of the mid-lobe, the lateral keels stop below the base of the mid-lobe. Column

arcuate, 7.5 mm. long.

This species is new to our region.

British Honduras. — Belize: Wooded island, Colonel English Pine

Ridge, Belize-Cayo Road, Gentle 9736.

Chysis aurea Lindl.; Ames & Correll, p. 491, fig. 135.

British Honduras. — Toledo: Beyond Central Camp, Edwards

Road beyond Columbia, Gentle 7401.

Bulbophyllum pachyrhachis (A. Rich.) Griseb.; Ames & Cor-

rell, p. 496, figs. 137, 138.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa F4,

Arroyo Corriental, on Aguada Terminos road, Contreras 1935; Lun-

dell 16570.

British Honduras. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4339. Edwards Road beyond Columbia,

Gentle 6377, 7301.

Eulophia alta (L.) Fawc. & Rendle; Ames & Correll, p. 499,

fig. 139.

British Honduras. — Stann Creek: 17 Miles Section, Stann Creek

Valley, Gentle 9262. Carib Switch near Stann Creek River, Gentle

7841. — Toledo: Condemn Branch Pine Ridge, Gentle 5143. Near and

beyond Columbia, Gentle 6075, 6151. Near San Antonio, Gentle 7577.

Cyrtopodium punctatum (L.) Lindl.; Ames & Correll, p. 501,

fig. 140.

Guatemala. — Alta Verapaz: Road from Secanquim to Sepacuite",

0. F. Cook & R. F. Griggs 368. Cacao, Finca Trece Aguas, G. P. Goll

105, F. L. Lewton 306.

WARREA Lindl.

Erect terrestrial highland herbs with lanceolate acute to acuminate plicate

strongly veined leaves. Leaves contracted at the base into a short stout sheath-

ing petiole, either with or without an enclosed short cylindric tapering pseudobulb.

Inflorescence an erect leafless raceme produced from the axil of the lowest basal

petiolar bract. Flowers relatively large and conspicuous. Sepals subequal, con-

CORRELL: ORCHIDS OF GUATEMALA 207

cave, the bases of the lateral sepals oblique and adnate to the column-foot. Petals

subequal to the sepals but usually somewhat wider. Lip subentire, the lateral

margins erect and somewhat recurved; base of lip adnate to the column-foot; apex

of lip spreading, emarginate, bilobed, obscurely apiculate or entire; disc lamellate.

Column elongate, semiterete, with a short basal foot. Anther terminal, operculate,

incumbent; pollinia 2 to 4, broadly semiglobose, waxy.

A small complex genus of about 6 species that occur from Guate-

mala south to northern South America.

Warrea costaricensis Schltr., Repert. Sp. Nov. 16: 446. 1920.

Figure 53.

Terrestrial or semi-epiphytic in high forests, up to about 1200 m.

alt. Uncommon in Guatemala, Costa Rica and Panama.

Plants erect, up to about 7 dm. tall. Leaves broadly elliptic-lanceolate, long-

acuminate, 25-60 cm. long, up to 12 cm. wide, contracted at base into a short stout

sheathing petiole. Inflorescence erect racemes of 6 or more large flowers, equal to

or somewhat exceeding the leaves, produced from the sheathing base of a leafy peti-

ole. Flowers large, with pedicellate ovaries up to 3 cm. long, reddish brown to deep

bronze-red, the lip lighter with reddish-bronze markings. Floral bracts ovate to

elliptic-lanceolate, about 1.5 cm. long. Dorsal sepal ovate-oblong, obtuse, concave,

3.2-3.5 cm. long, about 1.5 cm. wide. Lateral sepals subequal to the dorsal sepal,

the bases oblique to form a short rounded mentum with the column-foot. Petals

obovate, obtuse, oblique, with undulate-pleated margins, 2.8-3 cm. long, 1.3-1.5

cm. wide. Lip fleshy-thickened, 2.5-3.5 cm. long, 2.5-3 cm. wide, suborbicular to

broadly elliptic when spread out, the margins coarsely undulate-crenate; disc with

a narrow basal callus about 1.5 cm. long. Column slender, arcuate, about 2.5 cm.

long, produced at the base into a foot.

This genus and species is new to Guatemala.

Guatemala. — Pete"n: In high forest on Rio Mopan trail, about

5 km. east of Dolores, Contreras 3191.

Govenia superba (La Llave & Lex.) Lindl. ex Lodd.; Ames &

Correll, p. 505, fig. 141.

Guatemala. — Jalapa: Laguna de Ayarza, Heyde & Lux 3851.

Govenia utriculata (Sw.) Lindl.; Ames & Correll, p. 507.

Guatemala. — Sacatepe"quez : Volcan de Agua, T. Shannon 3668;

L. 0. Williams and A. Molina R. 13194.

Mormodes buccinator Lindl.; Ames & Correll, p. 509.

British Honduras. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4325.

Catasetum integerrimum Hook.; Ames & Correll, p. 516,

fig. 144.

FIG. 53. Warrea costaricensis. Plant (X 1/6); flower, spread out (X 1).

Drawn by Vivien Frazier.

208

CORRELL: ORCHIDS OF GUATEMALA 209

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Lundell 16241.

British Honduras. — Toledo: Near Rio Grande, Gentle 4872. Crique

Queva, Gentle 6021. Beyond San Antonio, Gentle 6586. Near Orange

Point, Gentle 7445. Lower Joe Taylor Creek, Gentle 7497.

Cycnoches ventricosum Batem.; Ames & Correll, p. 519.

Guatemala. — Pete"n: Tikal National Park, on Uaxactun trail,

Contreras 1229. Tikal National Park, about 11 km. northeast of

Tikal, Contreras 1918. On Santo Toribio Trail, about 3 km. north-

west of Dolores, Contreras 2430.

Stanhopea ecornuta Lem.; Ames & Correll, p. 531, fig. 150.

Guatemala. — Alta Verapaz: Cubilqiiitz, Turckheim 7996.

Stanhopea oculata (Lodd.) Lindl.; Ames & Correll, p. 532.

Guatemala. — Pete"n: Santo Toribio, Contreras 3009. Tikal Na-

tional Park, in ramonal covering the ruins, Contreras 1228. Tikal

National Park, on Uaxactun trail, Lundell 16521.

Gongora macula ta Lindl.; Ames & Correll, p. 540, fig. 155.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 610, 1844; Lundell 15565. Dolores, km. 84

of road, Contreras 2500. Lacandon, Contreras 3433. Tikal National

Park, Aguada Las Cucas, J. A. Ibarra 71; Lundell 16896.

British Honduras. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4319. Beyond Central Camp, Edwards

Road beyond Columbia, Gentle 7362.

Coryanthes macula ta Hook. Bot. Mag. t. 3102. 1831; Allen,

Fl. Panama (Ann. Mo. Bot. Gard. 36: 67 (403), fig. 173. 1949).

Epiphytic on trees in forests on high ridges and hills at low ele-

vations. Rare in British Honduras, Panama and northern South

America.

Plants erect, with elongate subcylindric tapering and strongly ridged bifoli-

ate pseudobulbs, 6.5-15 cm. long and 2.5-4 cm. wide. Leaves 2, at apex of pseu-

dobulb, firm, lanceolate, acute to acuminate, plicate, strongly veined, 30-60 cm.

long, 4-10 cm. wide. Inflorescence from base of pseudobulb, a slender, elongate

raceme 3-5 dm. long, with the peduncle, rachis and pedicellate ovaries glandular-

scurfy. Flowers usually 2 to 4, of variable color, large, subtended by membranous

bracts up to 3 cm. long. Sepals membranous, free, clear yellow with a few purple

spots, pale purple or reddish brown. Dorsal sepal suborbicular-ovate, broadly

acute and minutely apiculate, the apical half strongly reflexed, the lateral margins

revolute, 2-2.5 cm. long, 3-3.5 cm. wide when spread out. Lateral sepals very

strongly reflexed, from a broadly lobulate base, obliquely subfalcate or the pair

210 FIELDIANA: BOTANY, VOLUME 31

(when spread out) dolabriform, 5-6.5 cm. long, 2.5-4 cm. wide. Petals membra-

nous, usually colored similarly to the sepals, ligular-lanceolate, the apices obliquely

subacute, the margins undulate, 3-3.5 cm. long, 6-10 mm. wide. Lip very fleshy,

complexly 4-parted, clear waxy yellow, pale purplish spotted with dark red or the

hypochile yellow, the mesochile and epichile reddish brown on the outer surfaces,

the inner epichile white spotted red-brown: claw semiterete, continuous with the

base of the column; hypochile broadly galeate, glabrous, the apex obtuse to sub-

acute, in profile 2-2.5 cm. long and 2-2.5 cm. wide; mesochile canaliculate, gla-

brous, the basal half covered by the hypochile, in profile 2.5-3 cm. long and 8-15

mm. wide; epichile deeply cucullate or galeate, broadly obtuse, seen in profile 3-

3.5 cm. long and 2.5-3.5 cm. wide, in fresh material resembling a waxy cup or

bucket, prolonged below the apex of the column into a 3-cornute spout or channel.

Column subterete, without a foot, near the base with 2 broad lateral subfalcate or

obliquely auriculate glands, 8-10 mm. long, 5-10 mm. wide, the apex strongly re-

flexed and shortly 2-alate on either side of the clinandrium.

This species is new to our region.

British Honduras. — Toledo: On tree, in high ridge, on hill top

near Central Camp, Edwards Road beyond Columbia, Gentle 7269.

Lycaste aroma tica (Graham) Lindl.; Ames & Correll, p. 550.

Guatemala. — Alta Verapaz: Cajval, bank of Gahaton River, H.

Pittier 210.

Lycaste cruenta Lindl.; Ames & Correll, p. 552.

Guatemala. — Amatitlan, Las Cachiflanas, V. de Pacaya, Tonduz

407.

Lycaste tricolor (Kl.) Reichb. f. ; Ames & Correll, p. 554, fig. 158.

Guatemala. — Alta Verapaz: Sepacuite", 0. F. Cook 118. Cubil-

giiitz, Turckheim II 1909.

Maxillaria alba (Hook.) Lindl.; Ames & Correll, p. 563.

This species was included in the Orchids of Guatemala based on a

report of its occurrence there by Fawcett and Rendle. The speci-

mens cited here are the first we have seen for both Guatemala and

British Honduras.

Guatemala. — Alta Verapaz: Coban, Turckheim 1216. Cubilqtiitz,

Turckheim 7785.

British Honduras. — El Cayo: Blue Hole Section, Humming Bird

Highway, Gentle 8941.— Toledo: Lower Joe Taylor Creek, Gentle 7493.

Between Rancho Chico and Cockscomb, Monkey River, Gentle 4338.

Middle Camp, Edwards Road beyond Columbia, Gentle 6459.

Maxillaria crassifolia (Lindl.) Reichb. f.; Ames & Correll, p. 568,

fig. 163.

CORRELL: ORCHIDS OF GUATEMALA 211

Guatemala. — Pete"n: About 5 km. south-southeast of Dolores,

Contreras 3032. Tikal National Park, Tikal, in ramonal at base of

Temple V, Lundell 17171.

British Honduras. — Stann Creek: Dry Creek, Humming Bird

Highway, Gentle 8404. — Bolo Camp, upper reach of Golden Stream,

Gentle 4520. Near San Antonio, Gentle 6660. Lower Joe Taylor

Creek, Gentle 7501.

Maxillaria cucullata Lindl.; Ames & Correll, p. 570.

Guatemala. — Baja Verapaz: Santa Rosa, Turckheim 1171.

Maxillaria densa Lindl.; Ames & Correll, p. 572, fig. 164.

Guatemala. — Pete"n: Tikal National Park, Tikal, east of Laguna

Dimick, Contreras 639. Tikal National Park, Bajo de Santa Fe\

Arroyo Corriental, on Aguada Terminos Road, Lundell 16566. —

Quiche" : Sacabaja, Heyde & Lux 3506. — Sacatepe"quez : Santiago, R.

Gomez 886.

British Honduras. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4337. Beyond Union Camp, Edwards

Road beyond Columbia, Gentle 6500.

Maxillaria elatior Reichb. f.; Ames & Correll, p. 575.

This species is new to British Honduras.

British Honduras. — Toledo: Temash River, Gentle 5219. Beyond

Columbia, Gentle 6136. Monday Morning Hill, Edwards Road be-

yond Columbia, Gentle 6414.

Maxillaria Friedrichsthalii Reichb. f.; Ames & Correll, p. 576,

fig. 165.

Guatemala. — Pete"n: Tikal National Park, Tikal, growing on top

of temple in "H" Group, Contreras 369, 370. Tikal National Park,

Bajo de Santa FC", Contreras 453, 558, 559. Dolores, km. 87, east of

Machaquila Road, Contreras 2982. Tikal National Park, on Remate

road south of Tikal, Lundell 15557. — Santa Rosa: Casillas, Heyde &

Lux 4592.

British Honduras. — Toledo: Near San Antonio, Gentle 5515. Co-

lumbia Road, Gentle 6125. Edwards Road beyond Columbia, Gentle

7161, 7255.

Maxillaria pulchra (Schltr.) L. Wms. ex Correll; Ames & Cor-

rell, p. 581.

Guatemala. — Pete"n: About 400 m. south of Dolores, Contreras

2844. Monte Santa Teresa, Lundell 2978. Canchacan, Lundell 16472.

212 FIELDIANA: BOTANY, VOLUME 31

British Honduras. — Toledo: Edwards Road beyond Columbia,

Gentle 6531, 7238.

Maxillaria ringens Reichb. f.; Ames & Correll, p. 583.

Guatemala. — Pete'n: Santo Toribio, Contreras 2756. On Rio Mo-

pan trail, about 3 km. east of Dolores, Contreras 2827. About 400 m.

south of Dolores, Contreras 2846.

British Honduras. — El Cayo: Blue Hole Section, Humming Bird

Highway, Gentle 8883. — Between Monkey River and Cockscomb,

Gentle 4305. Rio Grande, Gentle 4828. Near San Antonio, Gentle

5419. Joe Taylor Creek, Gentle 7427, 7494.

Maxillaria rufescens Lindl.; Ames & Correll, p. 584.

Guatemala. — Pete'n: Tikal National Park, Tikal, on top of Tem-

ple IV, Contreras 387. Tikal National Park, Tikal, in ramonal cover-

ing the ruins, Contreras 443; Lundell 15502, 17085. Tikal National

Park, Bajo de Santa F£, Contreras 574, 577. Tikal National Park, in

tintal on pinal trail, Contreras 606. Aguada El Cartucho, about 8 km.

west of Dos Lagunas, Contreras 1592. About 5 km. east of Dos La-

gunas, Contreras 1752. Tikal National Park, Bajo del Hormiguero,

on old Remate road, Contreras 1958.

British Honduras. — El Cayo: 40 Miles Section, Humming Bird

Highway, Gentle 9012.— Toledo: Near Jacinto Creek, Gentle 4986.

Near Jacinto Hill, Gentle 5131. Middle Camp, Edwards Road be-

yond Columbia, Gentle 6410. Lower Joe Taylor Creek, Gentle 7495.

Maxillaria simulans (Ames & C. Schweinf.) L. Wms., Ceiba

5(4) : 207. 1956. M. purpurea (Spreng.) Ames & Correll, p. 582.

This species is new to British Honduras.

British Honduras/ — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4311. Punta Gorda-Seven Hills Road,

ascending Rio Grande, Gentle 4996. Near Columbia, Gentle 6077.

Edwards Road beyond Columbia, Gentle 6456, 7160.

Maxillaria tenuifolia Lindl.; Ames & Correll, p. 585.

Guatemala. — Alta Verapaz: Road from Secanquim to Sepacuite1,

0. F. Cook & R. F. Griggs 375.— Pete'n: Tikal National Park, Bajo

de Santa F£, Arroyo Corriental, on Aguada Terminos road, Lundell

15797.

British Honduras. — Toledo: Edwards Road beyond Columbia,

near Central Camp, Gentle 7267, 7268.

CORRELL: ORCHIDS OF GUATEMALA 213

Maxillaria uncata Lindl.; Ames & Correll, p. 586.

Guatemala. — Pete"n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 141, 647; Lundell 15434. Tikal National

Park, Bajo de Santa F4, Contreras 340, 557, 561. Tikal National Park,

Tikal, growing on top of temple of "H" Group, Temple III and Tem-

ple IV, Contreras 371, 494, 1796; Lundell 17163. Aguada El Car-

tucho, about 8 miles west of Dos Lagunas, Contreras 1593. About

400 m. south of Dolores, Contreras 2840. Dolores, km. 87, west of

Machaquila Road, Contreras 2983. Dolores, Contreras 2998. 2 km.

east of Poptun, Lundell 16420.

British Honduras. — El Cayo: 49 Miles Section, Humming Bird

Highway, Gentle 9109.— Toledo: Rio Grande, Gentle 4830. Near San

Antonio, Gentle 5475, 7727. Edwards Road beyond Columbia, Gen-

tle 6148, 6290, 6409, 7131, 7214. Joe Taylor Creek, Gentle 7433.

Maxillaria variabilis Batem. ex Lindl.; Ames & Correll, p. 587,

fig. 166.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa FC", Con-

treras 336, 563; Lundell 16564. Tikal National Park, Tikal, on top

of Temple IV, Contreras 388. Tikal National Park, Tikal, growing on

Temple of the Inscriptions, Contreras 510. Tikal National Park,

northeast of Tikal, Contreras 605. Tikal National Park, Tikal, in

ramonal covering the ruins, Contreras 3241; Lundell 15448. Tikal

National Park, on Remate road south of Tikal, Lundell 15556.

British Honduras. — El Cayo: 31 % Miles Section, Humming Bird

Highway, Gentle 8718. Blue Hole Section, Humming Bird Highway,

Gentle 8940. 38 Miles Section, Belize-Cayo Road, Gentle 9359.—

Stann Creek: 28 Miles Section, Humming Bird Highway, Gentle 9309.

North Branch Section, Humming Bird Highway, Gentle 9222. — To-

ledo : Between Monkey River and Cockscomb, Gentle 4274. Edwards

Road beyond Columbia, Gentle 6137, 6413, 7258.

Mormolyca ringens (Lindl.) Schltr.; Ames & Correll, p. 590,

fig. 167.

Guatemala. — Pete*n : Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 444; Lundell 15449, 15500. Tikal National

Park, Bajo de Santa F<§, Lundell 17013.

British Honduras.- — El Cayo: Humming Bird Highway, Gentle

8443.— Toledo: Moffredye Creek, San Antonio, Gentle 5082. Ed-

wards Road beyond Columbia, Gentle 6376, 6496, 7254.

214 FIELDIANA: BOTANY, VOLUME 31

Trigonidium Egertonianum Batem. ex LindL; Ames & Cor-

rell, p. 592, fig. 168.

Guatemala. — Pete'n : Tikal National Park, Bajo de Santa Fe", Con-

treras 550. Dolores, km. 81, on Machaquila Road, Contreras 3112.

Tikal National Park, Tikal, in ramonal covering the ruins, Lundell

15503. — Suchitepe"quez : vicinity of Mazatenango, W. R. Maxon &

R. Hay 3631.

British Honduras. — El Cayo: Humming Bird Highway, Gentle

8644.— Toledo: Swasey Branch, Monkey River, Gentle 3860. Be-

tween Rancho Chico and Cockscomb, Monkey River, Gentle 4405.

Moffredye Creek, San Antonio, Gentle 5081. Edwards Road beyond

Columbia, Gentle 6208. Near Mafridyle Lagoon, Gentle 7702.

lonopsis satyrioides Reichb. f. in Walpers, Ann. Bot. 6: 683.

1863; Schultes, Nat. Orch. Trinidad and Tobago 216. 1960.

At low elevations among ruins. Uncommon in Guatemala, Ja-

maica, Hispaniola, Trinidad and northern South America.

Plant up to about 3 dm. tall, usually much shorter. Pseudobulbs obsolescent.

Leaves fleshy, terete, subulate, falcate, 4-13 cm. long, 2-4 mm. wide. Inflorescence

racemose or laxly paniculate, up to about 15 cm. long, few-flowered; peduncle pur-

ple or greenish purple, with a few distant lanceolate closely appressed sheaths that

are up to 8 mm. long. Floral bracts small, triangular, acute, 2-3.5 mm. long.

Flowers yellowish white, fragrant, about 5 mm. across, the lip and petals occasion-

ally red-striped. Sepals subequal, lanceolate, acute, 7-8 mm. long, 2-2.5 mm. wide;

lateral sepals basally united to form a 2-lobed sac. Petals ovate-oblong, obtuse to

acuminate, marginally undulate, about 7 mm. long and 3 mm. wide. Lip entire,

obovate, narrowly clawed, marginally undulate, obtuse, about 8.5 mm. long; claw

basally with 2 minute bright yellow calluses and apically bilamellate. Column

about 2 mm. long. Capsule ellipsoidal, up to 15 mm. long and 5 mm. in diameter.

This is the first known occurrence of this species in Central Amer-

ica. It may be distinguished from 7. utricularioides in our flora by its

inconspicuous more or less obsolescent pseudobulb, terete, not flat,

leaves, and entire, not 2-lobed, lip.

Guatemala. — Pete'n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Conteras 1975.

lonopsis utricularioides (Sw.) Lindl.; Ames & Correll, p. 597,

fig. 170.

Guatemala. — Alta Verapaz: Cacao, Trece Aguas, F. L. Lewton

275. — Pete'n: Tikal National Park, Tikal, in ramonal covering the

ruins, Contreras 139. Tikal National Park, Isla de Los Pavos, Con-

treras 1277.— Santa Rosa: Casillas, Heyde & Lux 4626.

CORRELL: ORCHIDS OF GUATEMALA 215

British Honduras. — Toledo: Cattle Pass, Swasey Branch, Mon-

key River, Gentle 4448.

Odontoglossum bictoniense (Batem.) Lindl.; Ames & Correll,

p. 608.

Guatemala. — Sacatepe"quez : San Rafael, J. D. Smith 2638.

Odontoglossum convallarioides (Schltr.) Ames & Correll, p. 610.

Guatemala. — Quiche": Chiul, Heyde & Lux 3503.

Odontoglossum Londesboroughianum Reichb. f. Gard. Chron.

n.s. 6: 772. 1876.

Williams, in The Orchidaceae of Mexico (Ceiba 2: 280. 1951), gives

this species as from Guatemala. He did not, however, include it from

Guatemala in his An Enumeration of the Orchidaceae of Central Amer-

ica, British Honduras and Panama (Ceiba, Vol. 5. 1956). Since we

have seen no specimens from either Guatemala or British Honduras

it was, and has been, excluded from our work.

Odontoglossum majale Reichb. f.; Ames & Correll, p. 616.

Guatemala. — Quiche": Chiul, Heyde & Lux 3502.

Odontoglossum pulchellum Batem. ex Lindl.; Ames & Cor-

rell, p. 617.

Guatemala. — Baja Verapaz: Tactic, F. C. Lehmann 1437. — Saca-

tepe"quez: Volcan Fuego, J. D. Smith 2641.

Odontoglossum Rossii Lindl.; Ames & Correll, p. 618.

Guatemala. — Chimaltenango: Tecpam, Montanas de Chichavac,

G. Salas 577.

Brassia caudata (L.) Lindl.; Ames & Correll, p. 624, fig. 176.

Guatemala. — Alta Verapaz: Cubilqiiitz, Turckheim 7992. — Pete"n:

Tikal National Park, Tikal, in ramonal covering the ruins, Contreras

506. Tikal National Park, northeast of Tikal, Contreras 1359. About

400 m. east of km. 78 of road, Dolores, Contreras 2275. Dolores, be-

tween kms. 86-87, east of Machaquila Road, Contreras 2964. Tikal

National Park, on Remate Road south of Tikal, Lundell 16209.

Tikal National Park, on Uaxactun trail, Lundell 16522.

British Honduras. — El Cayo: 49 Miles Section, Humming Bird

Highway, Gentle 9110. — Punta Gorda-Seven Hills Road, ascending

Rio Grande, Gentle 5015. Beyond Manga Camp, Edwards Road be-

yond Columbia, Gentle 6506.

216 FIELDIANA: BOTANY, VOLUME 31

Brassia macula ta R. Br.; Ames & Correll, p. 626.

Guatemala. — Pete"n : Tikal National Park, on Temple V, Lundell

15863.

Brassia verrucosa Lindl.; Ames & Correll, p. 628, fig. 177.

Guatemala. — Alta Verapaz: Pansamala, Turckheim 107.

Oncidium ascendens Lindl. ; Ames & Correll, p. 638.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa Fa", Con-

treras 552. Lake Pete"n Itza, northwest of San Andres, Contreras

3556. Tikal National Park, Bajo del Hormiguero, 3 leagues south of

Tikal on old Remate road, Lundell 15537. Tikal National Park,

Tikal, in ramonal covering the ruins, Lundell 15567, 16693. Uaxac-

tun, Lundell 16636. Tikal National Park, Tikal, on temple of "H"

Group, Lundell 16795.

British Honduras. — Stann Creek: Silkgrass Road, Gentle 8145.—

Toledo: Beyond Columbia, Gentle 6233. Hutes Creek, Edwards Road

beyond Columbia, Gentle 7198.

Oncidium brachyandrum Lindl.; Ames & Correll, p. 641.

Guatemala. — Santa Rosa: Zamorora, Heyde & Lux 4611. — near

Salama, F. C. Lehmann 1455.

Oncidium carthagenense (Jacq.) Sw.; Ames & Correll, p. 642,

fig. 181.

Guatemala. — Pete'n: Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Lundell 16148.

British Honduras. — Indefinite: Tower Hill Estate, J. S. Karling 50.

Oncidium Cavendishianum Batem.; Ames & Correll, p. 644.

Guatemala. — Alta Verapaz: Turckheim 1400. — Santa Rosa: Chi-

apas, Heyde & Lux 4602.

Oncidium crista-galli Reichb. f.; Ames & Correll, p. 645, fig. 182.

Guatemala. — Alta Verapaz : Finca Mocca, H. Johnson 42.

Oncidium ensatum Lindl. ; Ames & Correll, p. 647.

Guatemala. — Alta Verapaz: Between Cajabon and Lanquin,G. P.

Goll 135.

British Honduras.— Toledo: Condemn Branch Pine Ridge, Gentle

5147. Near San Antonio, Gentle 5505. 12 Miles, San Antonio-Punta

Gorda Road, Gentle 6671. Mafridyle Creek, Gentle 7580. San An-

tonio Road and Moho River, between 17 Miles, Gentle 7597.

CORRELL: ORCHIDS OF GUATEMALA 217

Oncidium luridum Lindl.; Ames & Correll, p. 650, figs. 183, 184.

Guatemala. — Pete"n: Lacandon, Contreras 3352. Tikal National

Park, on Remate road south of Tikal, Lundell 15554. — Sacatepe"quez :

Volcan Fuego, J. D. Smith 2642.

British Honduras. — El Cayo: Humming Bird Highway, Gentle

8642. — Toledo: Between Rancho Chico and Cockscomb, Monkey

River, Gentle 4396.

Oncidium pusillum (L.) Reichb. f.; Ames & Correll, p. 663,

fig. 187.

Guatemala.- — Alta Verapaz: El Cacao, Finca Trece Aguas, G. P.

Goll 104; F. L. Lewton 293.— Escuintla: San Juan Mixtam, J. D.

Smith 2259.— Pet^n: Tikal National Park, Tikal, bordering airfield,

Contreras 632. Tikal National Park, Isla de Los Pavos, Contreras

1319. About 10 km. west of Dos Lagunas, Contreras 1764. Tikal

National Park, Bajo del Hormiguero, on old Remate road, Contreras

1886; Lundell 15524. Bordering Rio Ixcol, 1 km. east of Dolores,

Contreras 2404; Dolores, bordering the village, Contreras 3046. Tikal

National Park, Tikal, in ramonal covering the ruins, Lundell 15835,

17168. Tikal National Park, Aguada Pucte, Lundell 16815.

British Honduras. — Stann Creek: Eve's Pine Ridge, Gentle 8485.

—Toledo: Bolo Camp, upper reach of Golden Stream, Gentle 4498.

Oncidium sphacelatum Lindl.; Ames & Correll, p. 666.

Guatemala. — Alta Verapaz : El Cacao, Finca Trece Aguas, G. P.

Goll 106; F. L. Lewton 305.— Pete"n: Remate, on Tikal Road, Con-

treras 817. Dolores, on Machaquila Road, Contreras 2159. Tikal

National Park, Bajo de Santa FC", Arroyo Corriental, on Aguada Ter-

minos road, Lundell 15793. Tikal National Park, Tikal, in ramonal

covering the ruins, Lundell 16690.

British Honduras.— El Cayo: H. H. Bartlett 12946.— Toledo: Bolo

Camp, upper reach of Golden Stream, Gentle 4504. Edwards Road

beyond Columbia, Gentle 6484, 7201.

Leochilus Johnstonii Ames & Correll, p. 674, fig. 188.

Guatemala. — Pete"n : Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Lundell 17086.

Leochilus labiatus (Sw.) 0. Ktze.; Ames & Correll, p. 676.

Guatemala. — Alta Verapaz: Near Finca Sepacuite", 0. F. Cook &

R. F. Griggs 24.

218 FIELDIANA: BOTANY, VOLUME 31

Leochilus scriptus (Scheidw.) Reichb. f.; Ames & Correll, p. 678.

Guatemala. — Pete"n: Tikal National Park, on Remate Road, south

of Tikal, Contreras 625. Tikal National Park, Tikal, in ramonal cov-

ering the ruins, Contreras 1978, 3248; Lundell 16550.

Lockhartia hercodonta Reichb. f. ex Kranzl.; Ames & Correll,

p. 682.

This species is new to British Honduras.

British Honduras. — Toledo: In high ridge, on hill top beyond

Central Camp, Edwards Road beyond Columbia, Gentle 7363. The

following are sterile but probably belong here. — Toledo: Edwards

Road beyond Columbia, Gentle 6469, 7321.

Ornithocephalus bicornis Lindl.; Ames & Correll, p. 685,

fig. 191.

This species is new to British Honduras.

Guatemala. — Pete"n: About 1 km. 700 m. south of Lacandon, Con-

treras 3501.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, in mountain cabbage ridge, Gentle 4267.

Ornithocephalus inflexus Lindl.; Ames & Correll, p. 686.

Guatemala. — Pete"n: Tikal National Park, Tikal, Aguada Seca,

Contreras 1062. Tikal National Park, Tikal, in ramonal covering the

ruins, Contreras 1086; Lundell 15938. Tikal National Park, Bajo de

Santa Fe", between Isla de Los Pavos and Aguada Lagarto, Contreras

1278. Dolores, bordering the village clearing, Contreras 2203. Do-

lores, about 400 m. west of km. 82 on Machaquila Road, Contreras

2328. On Santo Toribio trail northwest of Dolores, Contreras 2383,

2429. Dolores, bordering Rio Ixcol, Contreras 2416. About 4 km.

south -southeast of Dolores, Contreras 2487. On Rio Mopan Trail,

5 km. southeast of Dolores, Contreras 2568.

British Honduras. — El Cayo: 49 Miles Section, Humming Bird

Highway, Gentle 9108.— Toledo: Feeder Road near Big Fall, Gentle

7027.

Ornithocephalus iridifolius Reichb. f.; Ames & Correll, p. 687.

Guatemala. — Santa Rosa: Naranjo, Heyde & Lux 4618.

Notylia Barker! Lindl.; Ames & Correll, p. 689.

Guatemala. — Alta Verapaz: El Cacao, Finca Trece Aguas, G. P.

Goll 72; F. L. Lewton 276.— Pete'n: Tikal National Park, Bajo de

CORRELL: ORCHIDS OF GUATEMALA 219

Santa Fe", Contreras 31, 1279; Lundell 15592. Bordering clearing,

Dolores, Contreras 2204. Lacandon, bordering Sibal, Contreras 3454.

Tikal National Park, on Remate road south of Tikal, Lundell 15564,

16868. — Santa Rosa: Cuajiniquilapa, Heyde & Lux 6245.

British Honduras. — El Cayo: Humming Bird Highway, Gentle

8590.— Toledo: Jenkins Creek, Monkey River, Gentle 4425. Punta

Gorda-Seven Hills Road, ascending Rio Grande, Gentle 5002, 5010.

Temash River, Gentle 5228. Near San Antonio, Gentle 5438. Near

Condemn Branch Creek, Gentle 5550. Columbia Road, Gentle 6111,

6112, 6115. Edwards Road beyond Columbia, Gentle 6204, 6390.

Between Punta Gorda and Joe Taylor Creek, Gentle 6795. Between

Orange Point and Moho River, Gentle 7662.

Cryptarrhena guatemalensis Schltr.; Ames & Correll, p. 692.

This species is new to British Honduras.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, in mountain cabbage ridge, Gentle 4309. On tree, in high ridge

on hill slope beyond Union Camp, Edwards Road beyond Columbia,

Gentle 6501.

Cryptarrhena lunata R. Br.; Ames & Correll, p. 694, fig. 193.

British Honduras. — Toledo: Between Monkey River and Cocks-

comb, Gentle 4266.

Dichaea hystricina Reichb. f.; Ames & Correll, p. 699.

Guatemala. — Alta Verapaz : Near Finca Sepacuite", 0. F. Cook &

R. F. Griggs 439.

Dichaea muricata (Sw.) Lindl.; Ames & Correll, p. 702.

Guatemala. — Alta Verapaz : El Cacao, Finca Trece Aguas, G. P.

Goll 17, 73 ;F. L. Lewton 263.— Pet<§n: About 2 km. south of Dolores,

Contreras 2242.

Dichaea muricata var. neglecta (Schltr.) Kranzl.; Ames &

Correll, p. 703.

This variety is new to British Honduras.

British Honduras. — El Cayo: Chalillo Crossing-Cohune Ridge

trail, Lundell 6552. — Toledo: Give And Take Ridge, Rio Grande,

Gentle 4879.

Dichaea panamensis Lindl.; Ames & Correll, p. 703, fig. 196.

Guatemala. — Alta Verapaz: El Cacao, Finca Trece Aguas, F. L.

Lewton 272.

220 FIELDIANA: BOTANY, VOLUME 31

British Honduras. — El Cayo: 34 Miles Section, Humming Bird

Highway, Gentle 9183. — Toledo: Between Rancho Chico and Cocks-

comb, Monkey River, Gentle 4395. Between Condemned Branch

Pine Ridge and Moffredye Lagoon, Gentle 6032. Balsam Hill, Ed-

wards Road beyond Columbia, Gentle 6322. Near San Antonio,

Gentle 6632. Feeders Road, near Big Fall, Gentle 7026. Between

Orange Point and Moho River, Gentle 7674.

Campylocentrum fasciola (Lindl.) Cogn.; Ames & Correll,

p. 710.

Guatemala. — Pete"n: Tikal National Park, Bajo de Santa F6", Con-

treras 28, 580. Tikal National Park, Tikal, in ramonal covering the

ruins, Contreras 142, 1102, 1149. Tikal National Park, Isla de Los

Pavos, Contreras 1280. Tikal National Park, growing on top of Tem-

ple V, Contreras 1425. Tikal National Park, Bajo del Hormiguero,

on old Remate Road, Contreras 1960.

British Honduras. — Toledo: Moffredye Creek, San Antonio, Gen-

tle 5077. 17 Miles, San Antonio Road and Moho River, Gentle 7601.

Between Orange Point and Moho River, Gentle 7656.

Campylocentrum hondurense Ames, Sched. Orch. 5: 37, f.

1923.

Epiphytic in forests at low elevations. Rare in British Honduras

and Honduras.

Roots fibrous, up to 15 cm. long, spreading, longitudinally sulcate. Stems up

to 8 mm. long, complanate, concealed by the sheathing imbricating bases of the

leaves, erect or ascending. Leaves about 7, distichous, up to 2.7 cm. long, 3.5-

5 mm. wide, broadest above the middle, oblong, ligulate, articulated with a con-

duplicate base which is 2.5 mm. long, unequally bilobed at the tip, mid-vein prom-

inent beneath. Peduncles with the raceme 1.4-2.7 cm. long, arising from the axil

of the leaf -sheath. Raceme 1-2 cm. long. Bracts of the inflorescence 1.7 mm. long,

exceeding the pedicels of the flowers, rigid, triangular, acute, concave. Flowers

about 10, white, distichous, 3 mm. apart on the glandular-scabridulous rachis.

Dorsal sepal 2 mm. long, triangular, subacute, 1-nerved. Lateral sepals 2 mm.

long, about 1 mm. wide at the base, triangular, acute, 1-nerved, carinate along the

middle on the outer surface, connivent. Petals 1.7 mm. long, linear-oblong, 1-

nerved, acute, membranous, parallel with the dorsal sepal. Lip scarcely 2 mm.

long, a little less than 1 mm. wide near the base, triangular-trulliform to lanceolate,

acute, membranous, produced at the base into a slender spur, ecallose. Spur 3 mm.

long, terete, lightly sigmoid, attenuated at the ends, obtuse, more or less porrect.

Column stout, abbreviated. Capsule 5 mm. long, ellipsoidal, 6-angled, sparsely

glandulose.

This species is new to our region. It is rather distinct with its

characteristic elongated spur and abbreviated stem.

CORRELL: ORCHIDS OF GUATEMALA 221

British Honduras. — Toledo: Governor Creek, Monkey River, on

tree in high ridge, Gentle 4472.

Campylocentrum micranthum (Lindl.) Rolfe; Ames & Cor-

rell, p. 711, fig. 197.

Guatemala. — Alta Verapaz: El Cacao, Finca Trece Aguas, G. P.

Goll 71. — Pete*n: Tikal National Park, on old Remate Road, Contreras

626, 1967, 1995. Tikal National Park, Tikal, bordering airfield, Con-

treras 631. Tikal National Park, Tikal, on top of Temple IV,

Contreras 1186. Tikal National Park, Isla de Los Pavos, Contreras

1303; Lundell 16740. Tikal National Park, Tikal, in ramonal cover-

ing the ruins, Contreras 1979; Lundell 16095, 16574. Bordering vil-

lage clearing, Dolores, Contreras 2210, 3184. On tree in Dolores,

Contreras 3001. On Santo Toribio Trail, Dolores, Contreras 2393.

About 200 m. from Santo Toribio, Contreras 2762. Tikal National

Park, on road to Aguada Naranjal, Lundell 16890. Lacandon, in low

forest bordering Sibal, Contreras 3431.

British Honduras. — Toledo: Moffredye Creek, near San Antonio,

Gentle 5049. Near Central Camp, Edwards Road beyond Columbia,

Gentle 7311.

Campylocentrum Schiedei (Reichb. f.) Benth. ex Hemsl. ; Ames

& Correll, p. 712.

Guatemala. — Escuintla: Conception, J. D. Smith 2246.

EXCLUDED SPECIES

Williams (in Ceiba, vol. 5, Nos. 1-4. 1956) cites the following as

being within our region. I have, however, seen no material of any

of these taxa: Sobralia amparoae Schltr. (p. 22, British Honduras);

Spiranthes graminea Lindl. (p. 36, Guatemala and British Hondu-

ras) ; Spiranthes sceptrodes Reichb. f . (p. 41, Guatemala) ; Corymbor-

chis flava (Sw.) 0. Ktze. (p. 49, British Honduras); Malaxisfastigiata

(Reichb. f.) 0. Ktze. (p. 101, Guatemala); Liparis fantastica var.

cordiformis (C. Schweinf.) L. Wms. (p. 105, Guatemala) ; Maxil-

laria maleolens Schltr. (p. 203, Guatemala) ; Trichopilia marginata

Henfrey (p. 216, Guatemala).

Publications 997, 998 and 999

UNIVERSITY OF ILLINOIS-URBANA