THOMAS LINCOLN
CASEY
LIBRARY
1925
Ps
3
7
bic
= +
rs a
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in =
TRANSACTIONS
OF THE
ENTOMOLOGICAL SOCIETY
OF
LONDON.
Page 104, for
Page 104, for
Page 105, for
Page 107, for
Page 109, for
Page 110, for
Page 116, for
Page 116, for
Page 167, for
Page 180, for
Page 181, for
Page 181, for
Page 221, for
Page 223, for
Page 249, for
Page 273, for
Page 310, for
Page 510, for
Page xxxvii, for
ERRATA.
TRANSACTIONS.
Wotogenta
subtesselata
prurvosus
read Votogonia.
read subtessellata.
read = pruinosus.
crassiscornis read crass?cornts.
Droylus read Dorylus.
V. armata read SS. armata.
(Estuans read stuans.
read Apis.
Gynandroph halna read Gynandrophthalma,
Metotoplax read Metopoplax.
P. strichnocera read WD, strichnocera.
Megalocerea read Megalocerwa.
sp.nov. vead gen. nov.
Evichoma read Evirchoma,
Hybocompa read Hybocampa.
Lyceenide read Lycenide.
T. binghami read J. binghamt,
Ur ys read Urowxys,
Aphis
PROCEEDINGS.
Fuliginose read fuliginosa.
Page xxxvii, for Poncra read Ponera.
Page xlvi, for Mrs. Nicholls read Mrs. Nicholl.
Page xvii, for (orthocladius) read (Orthocladius).
Page lvi, for 16 ceratinia read 16 Ceratinia.
Page lvi, for 4ceratinia read 4 Ceratinia.
Page lviii, for JANSEN read JANSON.
Page lix, for Anisolobis read Anisolabis.
Page lix, for Huntingford read Huntingfield
THE
TRANSACTIONS
OF THE
ENTOMOLOGICAL SOCIETY
LOIN DON
I (Ma IRIs Peis ee la:
Os.
LONDON:
PRINTED FOR THE SOCIETY BY RICHARD CLAY AND SONS, LIMITED,
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SOLD AT THE SOCIETY’S ROOMS, 11, CHANDOS STREET
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AND BY LONGMANS, GREEN, AND CO,
PATERNOSTER ROW, E.C. ; AND NEW YORK.
1903-1904,
DATES OF PUBLICATION IN PARTS. .
Part I. (Trans., pp. 1-140, Proc., i-xii) was published 29th April, 1903.
tC. ee 141-238, ,, xXlli-xxiv) » gndJune, ,,
eit. (. F 239-498, ,, XXV—xxxil) » odth Oct., 2
»1V-( 4» 499-575, ,, xxxiii-Ixiv) . 4thDec, ,,
» We (———_————__ ,,__ I xv-ellxi) . 16th Mar., 1904.
ENTOMOLOGICAL SOCIETY OF LONDON.
FOUNDED, 1833.
INCORPORATED BY RoyaLt CHARTER, 1885.
OFFICERS and COUNCIL for the Session 1903-1904.
President.
Pror. EDWARD B. POULTON, M.A., D.Sc., F.R.S.
Vice-Presidents.
THe Rev. CANON FOWLER, M.A., D.Sc., F.LS.
Pror. RAPHAEL MELDOLA, F.R.S., F.C.S.
Dr. DAVID SHARP, M.A., F.R.S., F.LS.
Treasurer.
ROBERT McLACHLAN, F.R.S., F.LS.
Secretaries.
HERBERT GOSS, F.L:S.
HENRY ROWLAND-BROWN, M.A.
Wibrarian.
GEORGE CHARLES CHAMPION, F.ZS.
Otber Members of Council.
CotonEL CHARLES T. BINGHAM, F.ZS.
MALCOLM BURR, B.A., F.LS., F.Z.S.
Dr. THOMAS ALGERNON CHAPMAN, M.D., F.Z.S.
ARTHUR JOHN CHITTY, M.A.
HAMILTON H.C. J. DRUCE, F.Z.S.
Pror. LOUIS COMPTON MIALL, F.R.S.
THe Rev. FRANCIS DAVID MORICE, M.A.
CoLtoneEL CHARLES SWINHOE, M.A, F.L.S., F.Z.S.
CotonEL JOHN W. YERBURY, R.A., F.ZS.
Resident Librarian.
WILLIAM R. HALL.
(ay
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ENTOMOLOGICAL SOCIETY O# LONDON.
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CONT ENTS.
PAGE
Errata ae ii
Explanation of the slefion viil
List of Fellows... ; ix
Additions to the Library XXliL
MEMOIRS.
I. A further contribution to our knowledge of African Phyto-
phagous Coleoptera, Part II. By Martin Jacopy, F.E.S.... 1
II. On the life history of Drilus flavescens, Rossi. By Lioner R.
CrawsHay, M.A. Oxon. Communicated by CHARLES OWEN
WarteruHOUSE, F.E.S. ase a aes eo a0 ld)
III. On the genus Dedlemera, Hiibner. By CoLoneL CHARLES
SWINHOE, M.A., F.L.S., IAG) és <0 08
IV. Some notes on the habits of WManophyes ing irene! as
observed in Central Spain. By GrorGE CHARLES CHAMPION,
F.Z.S., and Dr. THomas A. Cuapman, M.D., F.Z.8., with a
description of the larva and pupa by Dr. T. A.CHapman, M.D. 87
V. The <Aculeate Hymenoptera of Barrackpore, Spaces By
GEORGE ALEXANDER JAMES ROTHNEY, F.E.S.. 93
VI. Descriptions of nineteen new species of Lar ae (Ouener us end
Apidz from Barrackpore. By PETER CAMERON. Communi-
cated by GrORGE ALEXANDER JAMES ROTHNEY, F.E.S. 117
VIL. Notes on the nests of Bees of the genus Trigona. By CHARLES
OwrEN WATERHOUSE, F.E.S. nea : . 133
VIII. A remarkable new Lepidopterous Insect on alalandl By
Srr Grorce F. Hampson, Bart., B.A., F.Z.S. 5 Ms}?
IX. On Lepidoptera from the White ale! eollected by Mr. W. L.8
Loat, F.Z.8.; together with further Notes on Mee
Dimorphism in Butterflies. By Dr. Freperick A. DIxeEy,
M.A., M.D., Fellow of Wadham College, Oxford ... 141
X. An Entomological Excursion to Bejar, Central eeu By
GEORGE CHARLES CHAMPION, F.Z.S.... 207 . 165
XI. An Account of a Collection of Rhopalocera made on the
Anambara Creek in Nigeria, West Africa. By Percy I.
Latuy, F.Z.8., F.E.S. 183
XII. Rigi iae aculeata, collected i. the Rev. AueeED E.
Eaton, M.A., in Madeira and Tenerife, in the spring of 1902,
including notes on species taken by the late T. VERNON
Wottaston and F. A. BreLiamy. ah EpWwARD SAUNDERS,
F.R.S., F.L.S., ete. . 207
XIII. Deceripuars of twelve new genera aad species Gf We acernonil
(Herestarchint and Amblypygt), and three species of Ampulex
from the Khasia Hills, India. By Prrer Cameron. Com-
municated by GrorGe ALEXANDER JAMES RoTHNEY, F.E.S. 219
XIV. On a collection of Le ae een a from Arctic America. at HENRY
Joun Etwes, F.R.S., ¥.L.8., ete. ee . 239
XV. A contribution to the life iisbae of Orina (Chr eden loa) tr eh,
Fabr., var. smaragdina, Weise. By Dr. THomas ALGERNON
CHAPMAN, M.D., F.Z.S. 245
( viii )
PAGE
XVI. The Butterflies of Chile. By Henry Joun Etwes, F.R.S.,
F.L.S., ete. : ss a6 Ao ASE 50 ..- 263
XVII. On the genus ‘Theodosia and other Eastern Goliathidcs, with
descriptions of some new species By, OLIvER E. JANSON,
F.E.S. Py ae a3 ss o0 on ... 303
XVIII. Experiments in 1893, 1894, and 1896 upon the colour-relation
between lepidopterous larvee and their surroundings, and
especially the effect of lichen-covered bark upon Odontopera
bidentata, Gastropacha quercifolia, etc. By Professor
Epwarp B. Poutton, D.Sc., M.A., LL.D., F.R.S., etce.,
Hope Professor of Zoology in the University of Oxford,
Fellow of Jesus College, Oxford 900 ee . oll
XIX. A Revision of the Old World Lymantriidz in the National
Collection. a COLONEL CHARLES SwInHOE, M.A., F.L.S.,
eles =. > oo
XX. On the antennz of ee Gresca ees adee _) By
AMBROSE QUAIL, F.E.S. ... Ae ah ee Ee . 499
XXI. On the Laparostict Lamellicorn Galsepters of Grenada Me
St. Vincent (W. Indies). By Girpert J. Arrow, F.E.S8. 509
XXII. Note on the habits of Chtronomus (Orthocladius) sordidellus.
By THomas Harotp Taytor, M.A., F.E.S. ... ap .- O21
XXIII. Additions to the Rhynchotal fauna of Central America. By
Weel DISTANT hb sHiS.. ures ea a sce 230 wee O20
XXIV. Notes on some Central and South American Eryctnide, with
descriptions of new species. By Freprerick Du CANE
Gopman, D.C.L., F.R.S., etc. ... oes aa “ss Sen PAY)
XXYV. Supplementary Note to a Paper entitled Hymenoptera aculeata,
collected by the Rev. Arrrep E, Eaton, M.A., in Madeira
and Tenerife, in the aes of 1902. By Epwarp SAUNDERS,
BRS, ELS: : re sas eee Apo GIL
XXVI. Protective Gbieation in its eeaon to Mimicry, Common
Warning Colours, and Sexual Selection. By Assorr H.
TuHayEerR. Communicated by Professor Epwarp B. Poutton,
M.A., D.Sce., F-R.S. 36C ee 2 nbd 500 w. 593
XXVII. A brief discussion of A. H. THayeEr’s suggestions as to the
meaving of colour and pattern in insect bionomics. By
Professor Epwarp B. Pouttron, M.A., D.Sc., F.R.S. Boo ek)
Proceedings for 1903 Fas 2 se ae $53 oe: .. 1—Ixvill
Annual Meeting ... see tte ons a a sae “oe Ixviii
President’s Address sd wee ise ae es sh des xxiii
Index ae sea He oe oa A aes oH wee exvil
EXPLANATION OF THE PLATES.
PlatesI XII. See pages 39—51 Plates X & XI. See pages 245—261
Plates III & IV. - 58—84 Plates XII to XV. ,, 268—801
Plate V. 3 87—182 Plates XVI to ) 311374
Plate VI. | 138186 KV f 2 Ke
Plate VII. Me 141—163 Plate XIX. 5 499—508
Plate VIII. . 183—206 Plates XX to ) 529.550
Plate IX. a 239—243 LOCH J 2 a
Wist of Fellows
OF THE
ENTOMOLOGICAL SOCIETY OF LONDON.
Date of HONORARY FELLOWS.
Election.
1900 AvRIVILLIUS, Professor Christopher, Stockholm.
1900 Braver, Professor Friedrich Moritz, Mayerhofyasse 6, Viennu.
1901 Fapre, J. H., Sérignan, Vaucluse, France.
1894 Foret, Professor Auguste, M.D., Chigny, prés Morges, Switzerland.
1898 Grassi, Professor Battista, The University, Rome.
1884 OsTEN SACKEN, Baron C. R., Bunsenstrasse 8, Heidelberg.
1884 PackarD, Dr. Alpheus S., Providence, Rhode Island, U.S.A.
1872 Saussure, Henri F. de, Tertasse 2, Geneva.
1895 ScuppER, Samuel Hubbard, Cambridge, Mass., U.S.A.
1885 SNELLEN, Pieter Carl T., Rotterdam.
1893 WatrENwyL, Hofrath Dr. Carl Brunner Von, Lerchenfeldstrasse 28,
Vienna.
1898 Weismann, Dr. August, Frevburg, Baden.
FELLOWS.
Marked + have compounded for their Annual Subscriptions.
Date of
Election.
1901 + Aparr, Sir Frederick E. 8., Bart., Flixton Hall, Bungay.
1877 Apams, Frederick Charlstrom, F.Z.S., 50, Ashley-gardens, Victoria-
street, S.W.
1877 Apams, Herbert J., Roseneath, London-road, Enfield, N.
1902 ADKIN, Benaiah Whitley, Trenoweth, Hope-park, Bromley, Kent.
1885 ADKIN, Robert, Welljeld, Lingards-road, Lewisham, 8.E.
1899 ANnpDREws, Henry W., Shirley, Welling, S.O., Kent.
1901 Awnine, William, 39, Lime Street, E.C.
1899 + ARRow, Gilbert J., 87, Union-grove, Clapham, S.W.; and British
Museum (Natural History), Cromwell-road, S.W.
1886 ArmorE, E. A., 48, High-street, King’s Lynn.
1850 f AveBuRY, The Right Honble. Lord, D.C.L., F.R.S., F.L.S., F.G.S.,
etc., High Elms, Farnborough, Kent.
1901 Bacor, Arthur W., 154 Lower Clapton-road, N.E.
1903 Batpock, G. R., 71, Hertford-road, Lower Edmonton, N.
1886 Bankes, Eustace R., M.A., Norden, Corfe Castle, Wareham.
1890
1886
1895
1887
1902
1884
(2a)
Barcuay, Francis H., F.G.S., The Warren, Cromer.
BarGaGul, Marchese Piero, Piazza S. Maria, Palazzo Tempi No. 1,
Florence, Italy.
BarkER, Cecil W., Rownham, Malvern, Natal, South Africa.
Barker, H. W., 147, Gordon-road, Peckham, S.E.
BaRRAvD, Philip J., Bushey Heath, Watford.
Barrett, Charles Golding, Tremont, Peckham Rye, 8.E.
1894 — Bateson, William, M.A., F.R.S., Fellow of St. John’s College,
Cambridge, Merton House, Grantchester, Cambridge.
1896 | Beare, Prof. T. Hudson, B.Sc, F.R.S.E., 10 Regent Terrace,
Edinburgh,
1851 + Beaumont, Alfred, Gosfield, Halstead, Essex.
1899
1903
1897
1885
1895
1880
1895
1891
BEDWELL, Ernest C., Elinlea, Clevedon-roud, Norbiton, Surrey.
Bewi-Marzey, H. W., 50, Winder-street, Buyside, Durham, Natal.
Bennett, W. H., 15, Wellington-place, Hastings.
BretHune-Baker, George T., F.L.S., 19, Clarendon-road, Edgbaston,
Birmingham.
Bevan, Lieutenant H. G. R., R.N., H.M.S. “ Doris,” Channel
Squadron.
BIGNELL, George Carter, The Ierns, Homepark-road, Saltash.
Bryenam, Lieut.-Col. Charles T., F.Z.8., Bombay Staff Corps,
6 Gwendwr-road, West Kensington, 8.W.
BuaBer, W. H., F.L.S., 12, Great Castle-street, Regent-street, W.
1894 | BuackBuRNE- Maze, W. P., Shaw House, Newbury.
1889
1885
1886
1903
1891
1876
1875
1902
1892
1888
BuanprorpD, Walter F. H., M.A., F.Z.S., 12, Arundel Gardens,
Ladbroke-qrove, W.
Buatuwayt, Lieut.-Col. Linley, F.L.8., Eagle House, Batheaston,
Bath.
BLOOMFIELD, The Rev. Edwin Newson, M.A., Guestling Rectory,
Hastings. ;
Bogus, W. A., Wilts and Dorset Bank, Shepton Mallet
Boorn, George A., Fern Hill, Grange-over-Sands, Carnforth.
Borre, Alfred Preudhomme de, Villa la Fauvette, Petit Saconnex,
Geneva.
Borrer, Win, F.GS., Pakyns Manor House, Hurstpierpoint,
Hassocks, R1S.O., Sussex.
Bostock, E. D., Holly House, Stone, Staffs.
BouskE.L, Frank, Market Bosworth, Nuneaton.
Bower, Benjamin A., Langley, Willow Grove, Chislehurst.
1894 + Bowuss, E. Augustus, M.A., Myddelton House, Waltham Cross.
1852 + Boyp, Thos., Woodvale Lodge, South Norwood Hill, 8.E.
1893
1877
1870
1894
BraBant, Edouard, Chateaw de Morenchies, par Cambrai (Nord),
France.
Briaas, Charles Adolphus, Rock House, Lynmouth, R.S.O., N. Devon.
Briaes, Thomas Henry, M.A., Rock House, Lynmouth, RS.O., N.
Devon.
Bricut, Percy M., Chumar, Lansdowne-road, Bournemouth,
ot a)
1897 BricHtTweEn, Mrs. E., The Grove, Great Stanmore.
1890 Bristow, B. A., The Cottage, Stoke D’ Abernon, Cobham, Surrey.
1878 Broun, Capt. Thomas, Drury, Auckland, New Zealand.
1902 Brovuauton, Captain T. Delves, R.E., Alderney.
1886 Brown, John, 5, King’s Parade, Cambridge.
1892 Browne, Major Clement Alfred Righy, R.E., Lahore, India.
1898 | BucHan-Heprsurn, Sir Archibald, Bart., J.P., D.L., Smeaton-
Hepburn, Prestonkirk.
1883 Bucxron, George Bowdler, F.R.S., F.L.S., Weycombe, Haslemere,
S.0., Surrey.
1902 Buuurr, Arthur Percival, Wellington, New Zealand.
1896 | Burr, Malcolm, B.A., F.LS., F.Z.S., Royal Societies Club, St.
James's, 8.W.
1868 , Butter, Arthur G., Ph.D., F.LS., F.Z.8., The Lilies, Penge-road,
Beckenham.
1883 Burier, Edward Albert, B.A., B.Sc., 53, Tollington Purk, N.
1902. BurieEr, William E., Hayling House, Oxford-road, Reading.
1886 CaLveRT, Win. Bartlett, Liceo de Quillota, Quillota, Chili.
1902. Cameron, Malcolm, M.B., R.N., H.M.S. “ Harrier,” Mediterranean
Station.
1885 CampBELL, Francis: Maule, F.LS8., F.Z.S., &¢., Brynllwydwyn,
Machynlleth, Montgomeryshire.
1898 CANDEZE, Léon, 64, Rue de ? Ouest, Liege.
1880 CanspaLE, W. D., Sunny Bank, South Norwood, S.E.
1889 Cant, A., c/o Fredk. DuCane Godman, Esq., F.R.S., 10, Chandos-
street, Cavendish-square, W.
1890 Capper, Samuel James (President of the Lancashire and Cheshire
Entomological Society), Huyton Park, Liverpool.
1894 Caraccrtoto, H., H.M. Customs, Port of Spain, Trinidad, British
West Indies.
1892 CARPENTER, The Honble. Mrs. Beatrice, Avplin, Northallerton.
1895 CARPENTER, G. H., B.Se., Museum of Science and Art, Dublin.
1898 CaRPENTER, J. H., Riverdale, Leatherhead.
1868 CaARRINGTON, Charles, Hailey Hall, Hertford.
1890 Carrer, George Wm., M.A., F.L.S., Clif End House, Scarboro’.
1895 Carter, Sir Gilbert, K.C.M.G., 43, Charing Cross, W.C.; and
Government House, Nassau, Bahamas.
1900 Carter, J. W., 25, Glenholme-road, Manningham, Bradford.
1900 Cassat, R. T., M.R.C.S., Ballaugh, Isle of Man.
1903 Carrne, John Rowland, 59 and 61, Chancery Lane, E.C.
1889 + Cave, Charles J. T., Binstead, Cambridge. °
1900 CHAMBERLAIN, Neville, Highbury, Moor Green, Birmingham.
1871 CHampion, George C., F.Z.S., Liprartan, Heatherside, Horsell,
Woking ; and 10, Chandos-street, Cavendish-square, W.
1891 CHAPMAN, Thomas Algernon, M.D., F.Z.S8., Vick-PRESIDENT, Betula,
Reigate.
Cites)
1902 CHARNLEY, James Roland, Howick House, Howick, nr. Preston,
Lancashire.
1890 CHaATTERTON, Frederick J.S.,5, Camden Studios, Camden-street, N.W.
1897 CHAWNER, Miss Ethel F., Forest Bank, Lyndhurst, R.S.O., Hants.
1898 CHAWNER, Lawrence C., Forest Bank, tao R.S.O., ene
1902 CHEESMAN, E. M., c/o J. Garson, 63, Railway-street, Durban, Natal.
1891 + Carrry, Arthur John, M.A., 27, Hereford-square, 8S.W.; and Hunt-
ingfield, Faversham, ieee
1890 CHortEy, Mrs. H. S8., MJoorville Cottage, Bur ie in- Wharfedale,
Leeds.
1889 CuHristy, William M., M.A., F.L.S., Watergate, Emsworth.
1886 + CLaRK, John Adolphus, 57, Weston Park, Crouch End, N.
1867 CLaRKE, Alex. Henry, 109, Warwick-road, Earls Court, S.W.
1886 CLARKE, Charles Baron, M.A., F.RS, F.LS, F.GS., 13, Kew
Gardens-road, Kew, S.W.
1891 OLaRKE, Henry Shortridge, 2, Osborne-terrace, Douglas, Isle of Man.
1873 Coxe, William, F.L.S., Springfield, Buckhurst Hill, Essex.
1899 Coin, James E., Sussex Lodge, Newmarket.
1901 Connotp, Edward, 7, Magdalen Terrace, St. Leonards-on-Sea.
1900 Corton, Dr. John, 126, Prescot-road, St. Helens.
1892 Cowan, Thomas William, F.L.S., F.GS., F.R.MS., Pinehurst,
Pacific Grove, California.
1886 CoweELL, Peter (Librarian of the Liverpool Free Public Library),
William Brown-street, Liverpool.
1867 Cox, Herbert Ed., c/o Mrs. Eve, 125, Harley-street, W.
1895 CRABTREE, Benjamin Hill, The Oaklands, Levenshulme, Manchester.
1888 CrecoEr, J. P., Tredinick, Mayow-road, Sydenham, S.E.
1890 CREWE, Sir ieee Harpur, Bart., Ci tlke Abbey, Derbyshire.
1880 + Crisp, Frank, LL.B., B.A., J.P., ieeeaee L.S., 17, Throgmorton-
avenue, EC, and Heir Park ‘Henley on- Thames.
1902 CrutTtwELLt, The Rev. Canon Charles Thomas, M.A., Eivelme
Rectory, Wallingford.
1901 Dapp, Edward Martin, Bismarckstrasse 1, Charlottenburg, Germany.
1873 Dats, C. W., Glanville’s Wootton, Sherborne, Dorset.
1900 DateisH, Andrew Adie, 21, Prince’s-street, Glasgow.
1887 Dattry, The Rev. Thomas W., M.A., F.L.S., Madeley Vicarage,
Newcastle, Staffordshire.
1886 Dannatt, Walter, Donnington, 75, Vanbrugh Parl:, Blackheath, S.¥E.
1903 Day, F. H., 27, Currock-road, Carlisle.
1898 Day, G. O., Parr’s Bank-house, Knutsford.
1875 Distant, Wm. Lucas, Steine House, Selhurst-road, South Norwood,S.E.
1887 Drxey, Frederick Augustus, M.A., M.D., Fellow and Bursar of
Wadham College, VicE-PRESIDENT, Wadham College, Oxford.
1898 Drxon, G. B., St. Peter’s-road, Leicester.
1895 Dosson, H. T., Ivy House, Acacia Grove, New Malden, S.0., Surrey.
1903 Dotuman, J. C., Hove House, Newton-grove, Bedford-park, W.
1891
1885
1873
1845
1899
1884
1867
1900
1894
1883
1890
1865
1902
1886
1884
1900
1900
1886
1903
1878
1886
1903
1899
1890
1900
1861
1886
1889
1898
1878
1900
1874
1886
1900
1898
( xiii )
DonistHoRPE, Horace St. John K., F.Z.S., 58, Kensington-mansions,
South Kensington, S.W.
Donovan, Major Charles, M.D., R.A.M.C., c/o Messrs. P, Macfadyen
& Co., Winchester House, Old Broad-street, E.C.
Dorta, Marchese Giacomo, Strada Nuova, Genoa.
Dovatas, John Wm., 61, Craven Park, Harlesden, N.W.
Drewitt, Frederic G. Dawtrey, M.A., M.D., F.R.C.P., F.ZS.,
14, Palace Gardens-terrace, Kensington, W.
Druce, Hamilton H. C. J., F.Z.S., 43, Circus-road, St. John’s
Wood, N.W.
Druce, Herbert, F.L.S., F.Z.S8., 43, Circus-road, St. John’s Wood,
NCW:
Drory, W. D., Rocquaine, West Hill Park, Woking.
DupcGeon, G. C., Holia, Kangra Valley, P.O. Palimpur, Punjab,
India.
Durrant, John Hartley, The Cottage, Merton Hall, Thetford.
Eastwoop, John Edmund, Enton Lodge, Witley, Godalming.
Eaton, The Rev. Alfred Edwin, M.A., Woodlands, Seaton, Devon.
EpDEtstEN, Hubert M., The Elms, Forty Hill, Enfield, Middlesex.
Epwarps, James, Colesborne, Cheltenham.
Epwarps, Stanley, F.L.S., F.Z.S., 15, St. Germans-place, Black-
heath, S.E.
Extiort, E. A., 41, Holland Park, W.
Exuis, H. Willoughby, Knowle, Birmingham.
Ettis, John W., M.B., L.R.C.P., 18, Rodney-street, Liverpool.
ELTRINGHAM, Harry, Eastgarth, Westoe, South Shields.
Etwes, Henry John, J.P., F.RS. F.LS., F.Z.S., Colesborne,
Cheltenham.
Enock, Frederick, F.L.S., 13, Tufnell Park-road, Holloway, N.
ETHERIDGE, Robert, Curator, Australian Museum, Sydney, N.S.W.
FaRMBOROUGH, Percy W., F.Z.S., Lower Edmonton, N.
Farn, Albert Brydges, Mount Nod, Greenhithe, Kent ; and Medical
Department, Local Government Board, Whitehall, S.W.
Fettuam, H. L. L., P. 0. Box, 46, Johannesburg, Transvaal.
FENN, Charles, Hversden House, Burnt Ash Hill, Lee, 8.E,
Fenwick, Nicolas Percival, 7’e Gables, New-road, Esher.
FERNALD, Prof. C. H., Amherst, Mass., U.S.A.
Finer, F. E., 122, Stockwell Park-road, Brixton, 8.W.
Finzi, John A., 53, Hamilton-terrace, N.W.
Firts, J. Digby, F.L.S., Boys’ Modern School, Leeds.
Fitcu, Edward A., F.L.S., Brick House, Maldon.
Fircu, Frederick, Hadleigh House, Highbury New Park, N.
FLeMyNnG, The Rev. W. Westropp, M.A., Coolfin, Portlaw, Co.
Waterford.
FLetcaer, T. B., R.N., St. Catherines, Salishbury-road, Wimbledon.
(bake ))
1883 + FLetcHer, William Holland B., M.A., Aldwick Manor, Bognor.
1885 FoxKeEr, A. J. F., Zierikzee, Zeeland, Netherlands.
1900 FouxnKss, P. Hedworth, B.Sc., Harper-Adams Agricultiral College,
Newport, Salop.
1898 Fountaine, Miss Margaret, 7, Lansdowne-place, Bath.
1880 FowtrerR, The Rey. Canon, M.A., D.Sc, F.LS., Rotherfield
Peppard Rectory, Henley-on-Thames.
1883. FREEMAN, Francis Ford, Abbotsfield, Tavistock.
1896 FREKE, Percy Evans, Southpoint, Limes-road, Folkestone.
1888 FRemutn, H. Stuart, M.R.C.S., L.R.C.P., Mereworth, Maidstone.
1903 FReNcH, Charles, F.L.S., Government Entomologist, Victoria,
Australia.
1891 FRowawk, F. W., Rose Cottage, Hockley, S.O., Essex.
1855 Fry, Alexander, F.L.S., Vhornhill House, Dulwich Wood Park,
Norwood, 8.E.
1900° Fryer, H. Fortescue, The Priory, Chatteris, Cambs.
1884 Funnier, The Rev. Alfred, M.A., The Lodge, 7, Sydenham-hill,
Sydenham, S.E.
1898 Fuiier, Claude, Government Entomologist, Pietermaritzburg, Natal.
1887 GanHan, Charles Joseph, M.A., Whyola, Lonsdale-road, Bedford
Park, W.; and British Museum (Natural History), Cromovell-
road, S.W.
1892 GarRDE, Philip de la, R.N., H.M.S. “ Pegasus,” Mediterranean.
1890 GarRDNER, John, 6, Friars-gate, Hartlepool.
1901 | GARDNER, Willoughby, F.L.S., Deganwy, N. Wales.
1899 GAYNER, Francis, 20, Q@ueen-square, W.C.
1899 GeELDART, William Martin, M.A., Trinity College, Oxford.
1902 GiLLaNDERS, A. T., Park Cottage, Alnwick.
1865 | GopMan, Frederick Du Cane, D.C.L., F.R.S., F.L.S., F.Z.S., South
Lodge, Lower Beeding, Horsham; 7, Carlos-place, Grosvenor-
square; and 10, Chandos-street, Cavendish-square, W.
1890 GonLpruwalrt, Oliver C., 5, Queen’s-road, South Norwood, S.E.
1886 f GoopRicH, Captain Arthur Mainwaring, Lennox Lodge, Malvern
Link, Malvern.
1898 Gorpon, J. G. McH., Corsemalzie, Whauphill, R.S.O., Wigtownshire.
1898 Gorpon, R. 8. G. McH., Corsemalzie, Whauphill, R.S.O., Wigtown-
shire.
1855 GoruaMm, The Rev. Henry Stephen, F.Z.8., The Chestnuts, Shirley
Warren, Southampton.
1874 Goss, Herbert, F.L.S., Secretary, The Avenue, Surbiton-hill,
Surrey.
1886 GREEN, A. P., c/o 8. Green, Esq., 1, Gordon-place, Kensington, W.
1891 | GreEN, E. Ernest, Government Entomologist, Royal Botanic
Gardens, Peradeniya, Ceylon.
1850 GREENE, The Rey. Joseph, M.A., Rostrevor, Clifton, Bristol.
1898 GREENSHIELDS, Alexander, 38, Blenheim-gardens, Willesden, N.W.
1899
( xv )
GreEeNwoop, Edgar, Bellevue, Riffel-road, Willesden Green, N.W.
1893 + Gremnwoop, Henry Powys, F.L.S., Sandhill Lodge, Fordingbridge,
1888
1894
1900
1869
1899
1897
Salisbury.
GriFFitus, G. C., F.Z.S., 48, Caledonian-place, Clifton, Bristol.
GrimsHaw, Percy H., Natural History Department, Museum of
Science and Art, Edinburgh.
Groom, Prof. Perey, M.A., F.L.S., Royal Indian Engineering
College, Cooper's Hill, Staines.
GrosE-SmitH, Henley, J.P., B.A., F.Z.8.,5, Bryanston-square, Hyde
Park, W.
GunninG, Montague, Narborough, Leicester.
Haaug, Henry, 2, First-place, Brooklyn, U.S.A.
1890 + Haut, A. E., Norbury, Pitsmoor, Sheffield.
1885
1898
1891
1891
1903
Hat, Thomas William, Stanhope, The Crescent, Croydon,
Hamuyn- Harris, R., D.Se., F.Z.8., F.R.M.S., Toowoomba Grammar
School, Queensland, Australia.
Hampson, Sir George Francis, Bart., B.A., F.Z.S., 62, Stanhope-
gardens, S.W.
Hansury, Frederick J., F.L.8., Stainforth House, Upper Clapton,
N.E.
Hare, E. J., 8, Hillsborough-road, East Dulwich, 8.E.
1897 | Harrison, Albert, F.LS., F.C.8., Delamere, Grove-road, South
1889
1892
1881
1903
1898
1888
Woodford, Essex.
Harrison, John, 7, Gawlber-road, Burnsley.
HeApD.Ly, Charles Burnard, Two Elms, Alewandra-road, Stoneygate,
Leicester.
Henry, George, 38, Wellington-square, Hastings.
Herron, William, Horticultural College, Swanley, Kent.
Heron, Francis A., B.A., British Museum (Natural History),
Cromwell-road, 8.W.
Hiaes, Martin Stanger, F.C.S., F.G.8., Mine Office, Venterskroon,
Transvaal.
1876 | Hittman, Thomas Stanton, Hastgate-street, Lewes.
1888
1902
1887
1898
1897
1901
Hopson, The Rev. J. H., B.A., B.D., Harefield, Ansdell-road, Lythum.
Hor, R. 8., Indian Forest Service, c/o Messrs. King, King and
Co., Bombay.
Houianpd, The Rev. W. J., D.D., Ph.D., 5th Avenue, Pittsburg,
ent. OS. Al.
Hotman-Hunt, C. B., Aneimandi Esp., Munaar, P.O., Travancore,
S. India.
Horne, Arthur, 58, Gladstone-place, Aberdeen.
Hopson, Montagu F., L.D.S., R.C.S.Eng., F.L.S., 30, Thurlow-road,
Rosslyn Hill, N.W.
1876 | Horniman, Fredk. John, M.P., F.LS., F.Z.S., &e., Surrey Mount,
Forest Hill, S.E.
1903 Hovuenton, J. T., 1, Portland-place, Worksop.
(xvi. a)
1900 Howes, George H., Box 180, Dunedin, New Zealand.
1865 Hupp, A. E., Clinton, Pembroke-road, Clifton, Bristol.
1888 Hupson, George Vernon, The Post Office, Wellington, New Zealand,
1902. Hurron, Captain Frederick W., F.R.S., Director of the Canterbury
Museum, Christchurch, New Zealand.
1897 ImaceE, Selwyn, M.A., 20, Fifzroy-street, Fitzroy-square, W.
1893 Irpy, Lieutenant-Colonel Leonard Howard Loyd, F.L.S., F.ZS.,
14, Cornwall-terrace, Regent's Park, N.W.
1891 IsaBELL, The Rey. John, Sunnycroft, St. Sennen, R.S.O., Cornwall.
1886 Jacopy, Martin, 7, Hemstall-road, West Hampstead, N.W.
1869 Janson, Oliver E., Cestria, Claremont-road, Highgate, N.; and 44,
Great Russell-street, Bloomsbury, W.C.
1898 Janson, Oliver J., Cestria, Claremont-road, Highgate, N.
1886 JENNER, James Herbert Augustus, 209, School Hill, Lewes.
1899 Jenntnes, F. B., 152, Silver-street, Upper Edmonton, N.
1886 Joun, Evan, Llantrisant, R S.O., Glamorganshire.
1889 Jounson, The Rev. W. F., M.A., Acton Rectory, Poyntz Pass,
Co. Armagh.
1888 Jonss, Albert H., Shrublands, Eltham.
1894 + Jorpan, Dr. K., The Museum, Tring.
1902 Joy, Norman H., M.R.C.S., L.R.C.P., Bradfield, Reading.
1884 Kane, W. F. de Vismes, M.A., M.R.I.A., Drumleaske House,
Monaghan.
1884 Kappet, A. W., F.L.S., Hilden, 18, Sutton Court-road, Chiswick, W.
1876 F Kay, John Dunning, Leeds.
1896 | Kaye, William James, Caracas, Ditton Hill, Surbiton.
1884 Keays, Lovell.
1902 Kemp, Stanley W., Trinity College, Dublin.
1890 Kenrick, G. H., Whetstone, Somerset-road, Edgbaston, Birmingham.
1898 KerrsHaw,J.A., Morton Banks, Lewisham-road, Windsor, Melbourne
Victoria.
1901 KerrsHaw, John C. W., c/o F. W. Styan, Esq., Shanghai, China.
1900 Keys, James H., Morwell, Freedom-villas, Lipson-road, Plymouth.
1889 Kune, J. J. F. X., Lecturer on Economie Entomology at the West of
Scotland Agricultural College, 1, Athole Gardens-terrace, Kelvin-
side, Glasgow.
1861 Kuirsy, William F., F.L.S., Wilden, 18, Sutton Court-road, Chiswick,W.
1893 KirRKALDY, George Willis, Board of Agriculture, Division of
Entomology, Honolulu, Hawaii.
1889 KLAPALEK, Professor Franz, Karlin 263, Prague, Bohemia.
1887 + KLEIN, Sydney T., F.L.S.,F.R.A.S., Hatherlow, Raglan-road, Reigate.
1876 Kraatz, Dr. G., 28, Link-strasse, Berlin.
1901 Lane, E. W., Parkholme, 40, Fletching-road, Clapton, N.E.
1868 Lane, Colonel A. M., R.E., Box Grove Lodge, Guildford,
( xvii )
1960 Lane, The Rev. H. C., M.D., All Saints’ Vicarage, Southend-on-Sea,
1901 Laruy, Percy I., Lynton Villa, Sydney-road, Enfield.
1895 Larter, Oswald H., M.A., Charterhouse, Godalming.
1899 Lea, Arthur M., Government Entomologist, Hobart, Tasmania.
1900 Lerroy, H. Maxwell, B.A., Court, Crondall, Hants.
1901 Leicu, George F., corner of Sydenham and Essenwood-roads, Durban,
Natal.
1883 LeMmany, Fredk. Charles, Blackfriars House, Plymouth.
1892 Lesuiez, J. H., Bryn Glas, 33, Streathbourne-road, Upper Tooting, S.W.
1898 LetupripcEe, Ambrose G., Knowle, Dunster, Taunton.
1903 Levert, The Rev. Thomas Prinsep, Frenchgate, Richmond, Yorks.
1898 Lewis, E. J., F.L.8., Hadingford, Yalding, Kent.
1876 Lewis, George, F.L.S., 87, Frant-road, Tunbridge Wells.
1902 Lewis, J. H., Ophir, Otago, New Zealand.
1892 Liearroot, R. M., Bree-st., Cape Town, Cape of Good Hope.
1903 Lirrier, Frank M., Althorne, High-street, Launceston, Tasmania.
1865 | LuEwetyy, Sir John Talbot Dillwyn, Bart, M.A. F.LS.,
Penllergare, Swansea.
1881 + Luoyp, Alfred, F.C.S., The Dome, Bognor.
1885 + Luoyp, Robert Wylie, St. Cuthberts, Thurleigh-road, Balham, S.W.
1903 LorrHousg, Thomas Ashton, The Croft, Linthorpe, Middlesbrough. -
1899 Lounsspury, Charles P., B.Sc., Government Entomologist, Cape
Town, S. Africa.
1894 Lows, The Rev. Frank E., M.A., St. Stephen’s Vicarage, Guernsey.
1893 Lower, Oswald B., St. Oswald’s, Bartley-crescent, Wayville, South
Australia.
1901 Lower, Rupert 8., Bantley Crescent, Wayville, South Australia.
1898 Lucas, William John, B.A., 28, Knights Park, Kingston-on- Thames.
1880 Lupron, Henry, Lyndhurst, North Grange-road, Headingley, Leeds.
1903 Lyett, G., Junr., Gisborne, Victoria, Australia.
1901 Lyman, Henry H., M.A., F.R.G.S., 74, McTavish-street, Montreal,
Canada.
1902 Macponatp, George B. Douglas, M.B.
1887 M‘Doveatt, James Thomas, Dunolly, Morden-road, Blackheath, S.E.
1901 McGrecor, T. M., 48, Glasgow-road, Perth.
1888 Mackinnon, P. W., Lynndale, Mussoorie, N.W.P., India.
1900 Macxkwoop, The Hon. F. M., M.L.C., Colombo, Ceylon.
1858 McLacuuan, Robert, F.R.S., F.L.S8., F.Z.S., TREASURER, Westview,
23, Clarendon-road, Lewisham, S.E.
i898 Mappison, T., South Bailey, Durham.
1899 + Mary, Hugh, B.Sc., Almondale, Buckingham-road, South Woodford,
N.E.
1887 ManpeErs, Major Neville, R.A.M.C., c/o Sir Charles McGrigor,
Bart., and Co., 25, Charles-street, St. James’s-square, S.W.
1892 Manssripex, William, 27, Elmbank-road, Sefton-park, Liverpool.
1894 + MarsHatt, Alick, Auchinraith, Bexley, S.O., Kent. ;
( xvii)
1895 MarsHatt, G. A. K., P.O. Box 149, Salisbury, Mashonaland,
S. Africa.
1896 MarsHat., P., M.A., B.Sc, F.G.S., University School of Mines,
Dunedin, New Zealand.
1856 + MarsHA.L, William, Auchinraith, Bexley, S.O., Kent.
1897 Martineau, Alfred H., Solihull, Birmingham.
1895 Massy, Herbert, Zvy-Lea, Burnage, Withington, Manchester.
1865 Maruew, Gervase F., F.L.S., F.Z.S., F.R.G.S., Paymaster-in-chief,
R.N., Lee House, Dovercourt, Harwich.
1887 MatrHews, Coryndon, Stentaway, Plymstock, Plymouth.
1899 May, Harry Haden, 6, Citadel Terrace, Plymouth.
1872 + Metpoua, Professor Raphael, F.R.S., F.C.S., 6, Brunswick-
square, W.C.
1885 Metviti, James Cosmo, M.A., F.L.S., 36, George-street, Manchester.
1887 MERRIFIELD, Frederic, 24, Vernon-terrace, Brighton.
1888 Meyer-Darcts, G., c/o Sogin and Meyer, Wohlen, Switzerland.
1880 Meyrick, Edward, B.A., F.Z.S., Elmswood, Marlborough.
1894 Murat, Professor Louis Compton, F.R.S., 1, Richmond Mount,
Headingley, Leeds.
1883 Miss, W. H., The New Club, Calcutta.
1896 Moserty, J. C., M.A., 9, Rockstone-place, Southampton.
1879 Monrerro, Dr. Antonio Augusto de Carvalho, 70, Rua do Alecrinar,
Lisbon.
1902 Monteomery, Arthur Meadows, 83, Osborne-road, Forest Gate, E.
1853 Moors, Frederic, D.Sc., A.L.S., F.Z.S., 17, Maple-road, Penge, S.E.
1899 Moors, Harry, 12, Lower-road, Rotherhithe.
1886 Morean, A. C. F., F.LS., 24, Leinster-square, W.
1889 | Morice, The Rev. F. D., M.A., Fellow of Queen’s College, Oxford,
VicE-PRESIDENT, Brunswick, Mount Hermon, Woking.
1895 + Morey, Claude, Jpswich.
1893 Morroyn, Kenneth J., 13, Blackford-road, Edinburgh.
1900 Moser, Julius, 90, Bulow-strasse, Berlin.
1882 Mostey, 8. L., Beaumont Park, Huddersfield.
1898 Mousey, H., Burnfoot, Buxton.
1901 Murr, Frederick, EH. and 8S. African Telegraph Co., Mozambique.
1869 + Méiuer, Albert, F.R.G.S., c/o Herr A. Miiller-Mechel, Gren-
zacherstrasse, 60, Basle, Switzerland.
1872 + Murray, Lieut.-Col. H., 43, Cromwell Houses, Cromwell-road, S.W.
1903 Neavg, 8. A., B.A., Magdalen College, Oxford.
1896 NesHAwm, Robert, Utrecht House, Queen’s-road, Clapham Park, S.W.
1889 Nervinson, Basil George, M.A., F.ZS., 3, Tedworth-square,
Chelsea, S.W.
1901 Nevinson, E.G.B., 5, Bentinck-terruce, Regent’s Park, N.W.
1890 Newstead, R., The Museum, Chester.
1900 NicHox1, Mrs. M. Dela B., Merthyr Mawr, Bridgend, Glamorganshire.
1886 NicHotson, William E., School Hill, Lewes.
(pcb 1)
1893 NonrrieD, A. F., Rakonitz, Bohemia.
1886 Norris, Herbert E., 15, Market-place, Cirencester.
1878 Norrrmes, Thomas, Ashford, Kent.
1895 Nurse, Major C. G., Indian Staff Corps, Quetta, Baluchistan, India.
1869 OpsrrtHiir, Charles, Rennes (Ille et Vilaine), France.
1877 OsertHir, René, Rennes (Ille et Vilaine), France.
1893 + OcLE, Bertram S., Steeple Aston, Oafordshire.
1893. Outver, John Baxter, Elmleigh, Elm-row, Hampstead, N.W.
1873 Ouivier, Ernest, Ramillons, prés Moulins (Allier), France.
1895 Pacx, Herbert E., Bertrose, Gellatly-road, St. Catherine’s Park, 5.E.
-1898 PauuisER, H. G., Holmwood, Addlestone, Surrey.
1901 Prat, Henry Woolner, Indian Museum, Calcutta.
1883 PséRINGUEY, Louis, South African Museum, Cape Town, South Africa.
1903 + Perkins, R. C. L., B.A., Board of Agriculture, Division of Ento-
mology, Honolulu, Hawaii.
1879 PrRKINS, Vincent Robt., Wotton-under- Edge.
1900 Pures, The Rev. W. J. Leigh, 7'he Cottage, Parkwood-road,
Tavistock.
1897 Purnies, Hubert C., M.R.C.S., M. and L.8.A., 262, Gloucester-terrace,
Hyde-park, W.
1903 Parutrs, Montagu A., F.R.G.S., F.Z.S., 22, Petherton-road, Canon-
bury, N. ;
1901 Pickert, C. P., 99, Dawlish-road, Leyton, Essex.
1891 Prerce, Frank Nelson, 1, The Elms, Dingle, Liverpool.
1901 Prrrarp, Albert, Felden, Bowmoor, Hemel Hempstead.
1903 PrincuER, Colonel Jesse George, I.M.S., F.R.C.S., 133, Gloucester-
road, Kensington, W.
1885 Pott, J. R. H. Neerwort van de, Drishergen, Netherlands.
1870 + Porritt, Geo. T., F.L.S., Mayfield, Edgerton, Huddersfield.
1884 + Poutton, Professor Edward B., M.A., D.Sc., F.R.S., F.L.S., F.G.S.,
F.Z.S., Hope Professor of Zoology in the University of Oxford,
PRESIDENT, Wykeham House, Banbury-road, Oxford.
1851 Preston, The Rev. Thomas Arthur, M.A., F.L.8., Thurcaston
Rectory, Leicester.
1878 Prick, David, 48, West-street, Horsham.
1893 Prout, Louis Beethoven, 246, Richmond-road, Dalston, N.E.
1898 QuatiL, Ambrose, 15, Stamford-hiil, N.
1900 Ratnsow, William J., The Australian Museum, Sydiey, NS.W.
1874 Reep, E. C., Director del Museo de Concepcion, Concepcion, Chile.
1900 Retp, Percy Charles, Feering Bury, Kelvedon, Essex.
1893 Rep, Captain Savile G., late R.E., Zhe Elms, Yalding, Maidstone,
1891 Rerp, William, St. Andrews-road, Rondebosch, Cape Town, South
Africa.
1898 Re.toy, R. H., c/o Perkins and Co., Ltd., Brisbane, Queensland.
(==)
1890 RENDLESHAM, The Right Honble. Lord, Rendlesham Hall, Wood-
bridge.
1898 Reuter, Professor Enzio, Helsingfors, Finland.
1894 Rriprne, William Steer, B.A., M.D., Buckerell Lodge, Honiton.
1853 Ripon, The Most Honble. the Marquis of, K.G., D.C.L., F.R.S., F.L.S.,
ete., 9, Chelsea Embankment, S.W.
1892 Roprnson, Sydney C., Goldsmiths’ Hall, F.C.
1869 + Roprnson-Dovueras, William Douglas, M.A., F.LS.,. F.R.G.S.,
Orchardton, Castle Douglas.
1899 Rosson, John Emmerson, 15, Northgate, Hartlepool.
1886 Ross, Arthur J., 37, Church Crescent, Muswell Hill, N.
1868 Rotrunery, George Alexander James, Pembury, Tudor-road, Upper
Norwood, 8.E.
1894 + Rorascainp, The Honble. Nathaniel Charles, M.A., F.L.S., F.Z.S.,
148, Piccadilly, W. ; and Tring Park, Tring.
1888 + Roruscuitp, The Honble. Walter, D.Sc., M.P., F.LS., F.Z.8,, 148,
Piccadilly, W.; and Tring Park, Tring.
1890 RovutiepeGE, G. B., Tarn Lodge, Heads Nook, Carlisle.
1887 RownaNnp-Brown, Henry, M.A., Secretary, Oxhey-grove, Harrow
Weald.
1903 Rowtanps, Osbert William, Lickey Grange, nr. Bromsgrove.
1898 RussEtL, A., The Limes, Southend, Catford, 8.E.
1892 RussEtt, 8. G. C., 19, Lombard-street, E.C.
1899 Rywes, William E., B.A., 11, Waverley Mount, Nottingham.
1865 + SaunpERS, Edward, F.R.S.,F.L.8.,S4 Ain’s, Mount Hermon, Woking.
1861 | SaunpERS, G. S., F.LS., 20, Dents-road, Wandsworth Common,
S.W.
1886 SaunpERs, Prof. Wm., Central Experimental Farm, Ottawa, Canada.
1901 ScwHaus, W., F.Z.8S., Trentham House, Twickenham.
1881 Scoxnutck, A. J., The Hazels, Babington-road, Streatham.
1864 SempER, George, Klopstock-strasse 23, Altona, Hlbe, Germany.
1862 SwHarp, David, M.A., M.B., F.R.S., F.L.S., F.Z.S., Hawthorndene,
Hills-road, Cambridge ; and University Museum of Zoology and
Comparative Anatomy, Cambridge.
1902 SHarp, W. E., 9, Queen’s-road, South Norwood, S.E.
1883 SHaw, A. Eland, M.R.C.S., Overdale, Laxey, Isle of Man.
1901 SHELFORD, R., M.A., C.M.Z.S., The Musewm, Sarawak, Borneo.
1883 + SHELLEY, Capt. George Ernest, F.G.S., F.Z.S., 39, Eyerton-gardens,
S.W.
1900 + SHEPHEARD-Watwyy, H. W., M.A., Dalwhinnie, Kenley, Surrey.
1887 SricuH, Alfred, Corney House, Chiswick, W.
1901 SxkertcuHty, Ethelbert Forbes, c/o ‘Penang Gazette, Penang, Straits
Settlements.
1902 SLapEN, Frederick William Lambart, 2, Sydney-road, Walmer,
Deal.
1902 Storer, Gerard Orby, Westrop House, Highworth, Wilts.
1901
1901
1895
1898
1885
1897
1889
1898
1890
1897
1898
1889
1896
1900
1895
1903
1882
1884
1894
1876
1893
1892
1903
1901
1892
1897
(xx)
Situ, Arthur, 16, Hdward-street, Grimsby.
Smitu, W. G., Rosebank, Brecknock-road, Knowle, Bristol.
Samira, W. W., Ashburton, Canterbury, New Zealand.
Sopp, Erasmus John Burgess, F.R.Met.S., 104, Liverpool-road,
Birkdale, Lancashire.
SourtH, Richard, 96, Drakefield-road, Upper Tooting, 8.W.
SparKE, E. G. J., B.A., 1, Christchurch-Villas, Tooting Bec-road,
S.W.
STANDEN, Richard 8., F.L.S., Townlands, Lindfield, Sussex.
Srares, C. L. B., M.R.C.S., L.R.C.P., The Infirmary, Wandsworth,
S.W.
Srearns, A. E., 99, Gloucester-terrace, Hyde-park, W.
Sreppine, E. P., Indian Forest Service, c/o King, Hamilton and
Co., Calcutta.
STEBBING, Henry, 134, Westbowrne-grove, W.
Srraton, C. R., F.R.C.S., West Lodge, Wilton, Salisbury.
STRICKLAND, T. A. Gerald, 21, Kensington Gate, W.
Strupp, BE. A. C., Downton, near Salisbury.
Stupp, E. F., M.A., B.C.L., Oxton, Exeter.
Swag, Harold, M.B., Arawa House, Rotorua, New Zealand.
Swanzy, Francis, Stanley House, Granville-road, Sevenoaks.
SWINHOE, Colonel Charles, M.A., F.L.S., F.Z.S., 7, Gloucester Walk,
Kensington, W.
SwinHog, Ernest, 37, Addison-gardens, Kensington, W.
Swinton, A. H., c/o General Callender, Vineyard, Totnes.
Tayior, Charles B., Rae-street, Rae Town, Kingston, Jamaica.
Taytor, The Rev. George W., F.R.S. (Canada), St. Mutthew’s
Rectory, Wellington, British Columbia.
Taytor, Thomas Harold, M.A., Yorkshire College, Leeds.
THompson, Matthew Lawson, 2, Thornclitf Villas, Saltburn-by-the-
Sea.
THoRNLEY, The Rev. A., M.A., F.L.S., South Leverton Vicarage,
Lincoln.
Tomuin, B., M.A., Estyn, Chester.
1859 + TrimEN, Roland, M.A., F.R.S., F.L.S., 26, Campden-grove, Campden
1895
1897
1898
1893
1894
1886
1893
Hill, Kensington, W.
TuUNALEY, Henry, 30, Pairmont-road, Brixton Hill, S.W.
TunstaLL, Wilmot, Brook House, Meltham, Huddersfield.
Turner, A. J., M.D., Wickham Terrace, Brisbaue, Australia.
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TuRNER, Thomas, Cullompton, Devon.
Turt, James W., Rayleigh Villa, Westcomble Hill, S.E.
Uric, Frederick William, Port of Spain, Trinidad, British West
Indies.
1900
1866
1897
1895
1901
1899
1897
1870
1878
(= xxii")
Urwick, W. F., 34, Great Tower-street, E.C.
VERRALL, George Henry, Sussex Lodge, Newmarket.
Vice, William A., M.B., 19, Belvoir-street, Leicester.
WacHER, Sidney, F.R.C.S., Dane John, Canterbury.
WappinetTon, John, Park Holme, Harehill-avenue, Leeds.
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WALKER, James J., R.N., F.LS., c/o Admiralty, Whitehall, S.W.
1863 , Wattace, Alfred Russel, D.C.L., Oxon., F.R.S., F.LS., F.Z.S.,
Broadstone, Wimborne, Dorset.
1866 | WaLsINGHAM, The Right Honble. Lord, M.A., LL.D., F.R.S., F.LS.,
1886
1869
1901
1900
1893
F.Z.S., High Steward of the University of Cambridge, . Merton
Hall, Thetford ; and 66a, Eaton-square, S.W.
WaRREN, Win, M.A., 61, Wilton-avenue, Chiswick-lane, W.
WatTERHOUSE, Charles O., Ingleside, Avenue-gardens, Acten, W. ;
and British Museum (Natural History), Cromwell-road, S.W.
WATERHOUSE, Gustavus A., B.Se., F.C.S., Royal Mint, Sydney, New
South Wales, Australia.
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shire.
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1876 | WESTERN, E. Young, 36, Lancaster Gute, Hyde Park, W.
1886
1884
1903
1896
1894
1894
1900
1881
1901
1899
1891
1888
1892
WHEELER, Francis D., M.A., LL.D., Paragon House School, Norwich.
Wuire, William, 75, Thurlow Park-road, West Dulwich, S.E.
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British East Africa.
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Common, S.W.
WoopForDkE, F. C., Market Drayton.
Woo ttey, H. 8., 7, Park-row, Greenwich, S.E.; and P. 0. Box
1047, Waterbury, Conn., U.S.A.
Wrovueuton, R. C., Inspector General of Forests, Indian Forest
Service, c/o Army and Navy Co-operative Society, Ltd., 105,
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Yersury, Colonel John W., late R.A., F.Z.S., Army and Navy
Club, Pall Mall, 8.W.
Youpa.e, William Henry, F.R.M.S., Daltonleigh, Cockermouth.
(Texte *)
ADDITIONS TO THE LIBRARY
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AvSsTEN (E. E.). A Monograph of the Tsetse-flies. Svo, London, 1903.
[Published by the Trustees of the British Museum. ]
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Banxs (Nathan). A List of Spiders collected in Arizona by Messrs.
Schwarz and Barber during the Summer of 1901.
[Proc. U. S. Nat. Mus., Vol. XXV.]
The Smithsonian Institution.
BarGacwi (P.). Adolfo Targioni-Tozzeti (Obituary notice of).
[Bull. Soc. Ent. Ital., 1902. ]
The Society.
Barrett (C. G.). The Lepidoptera of the British Islands. Vol. VIII.
8vo, London, 1902.
The Publishers.
BETHUNE-BakER (G. T.). A Revision of the Amblypodia Group of Butter-
flies of the Family Lycaenidae.
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The Author.
Borxeat (H.). Notes sur quelques Lucanides.
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BouskeEtt (F.). Three Weeks in South Kerry, June 1902.
[Irish Naturalist., Vol. XII, 1903.]
H. St. J. Donisthorpe.
Burr (Malcolm). Orthoptera.
[Wytsman’s Genera Insectorum.] 4to, Bruxelles, 1903.
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CatvEert (P. P.). [See Gopman (F. D.). Biologia Centrali-Americana. ]
CaMBRIDGE (F. O. Pickard). [See Gopman (F. D.). Biologia Centrali-
Americana. |
CARPENTER © a ). Injurious Insects observed in Ireland during the Year
(Roy. Dublin Soc., Economic Proc. 1902. ] By Exchange.
CaupDELt (A. N.). Notes on Orthoptera from Colorado, New Mexico, Arizona,
He Texas, with descriptions of new species.
[Proc. U. 8. N. Mus., Vol. XXVI.] 8vo, Washington, 1903.
The Phasmidz or Walking-sticks of the United States.
[Proc. U. S. N. Mus. eV ol. XXVI .] 8vo, Washington, 1901.
The Smithsonian Institution.
Cuampron (G.C.). [See Gopman (F. D.). Biologia Centrali-Americana. ]
Coqurttett (D. W.). New Diptera from North America.
[Proc. U. S. Nat. Mus., Vol. XXV.]
(ayy)
Distant (W. L.). Insecta Transvaaliensia. Parts I—IV. 4to, London,
1900—1903. Purchased.
Dyar (H. G.). List of North American Lepidoptera, and Key to the
Literature of this order of Insects.
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Descriptions of the Larve of some Moths from Colorado.
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The Sinithsonian Institution.
FarrMaire (L.). Coléoptéres. d H
[Histoire Naturelle de la France, 8° partie.] S8vo, Paris, 1902.
The Editors.
Fisker (W. F.). The Parasites of the American Tent Caterpillar.
[N. Hampshire Coll. Agric. Exp. Stn. Techn. Bull. 6. 1903. |
The Author.
Forpes (8. A.). Report (22nd) of the State Entomologist on the Noxious
and Beneficial Insects of the State of Illinois. S8vo, Champaign,
1903. The Author.
Foret (A.). Les Formicides de l’Empire des Indes et de Ceylan.
(Journ. Bombay N. H. Soc., Vol. XIV. |
Fourmis nouvelles d’ Australie.
[Revue Suisse Zool.] 8vo, Genéve, 1902.
Faune Myrmécologique des Noyers dans le Canton de Vaud. Svo,
Lausanne, 1903.
Recherches biologiques récentes de Malle Adéle Fielde sur les Fourmis.
(Bull. Soc. Vaud Sc. Nat., XX XIX, 146.] 8vo, Lausanne, 1903.
Mélanges Entomologiques, biologiques et autres.
[Ann. Soc. Ent. Belgique, T. XLVII.] 8vo, Bruxelles, 1903.
Les Fourmis des iles Andamans et Nicobares.
[Rev. Suisse Zool., Tome II.] 8vo, Genéve, 1903. ‘
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Fow er (W. W.). [See Gopman (F. D.). Biologia Centrali-Americana. |
Gopman (F. Ducane). Biologia Centrali-Americana. Parts CLXXVI—
CLXXX. The Editor.
Arachnida Araneidea, by F. O. Pickard, Cambridge.
Diplopoda Araneidea, by R. I. Pocock.
Insecta, by P. P. Calvert, G. C. Champion, W. W. Fowler, F. M.
van der Wulp, and 8. W. Williston.
Guturikz (J. E.). The Collembola of Minnesota. 8vo, Minneapolis, 1903.
The Editor.
Hasert (J. N.). [See Jounson (W. F.).]
Hampson (Sir George, Bt.). Catalogue of the Lepidoptera Phalzenz in the
British Museum, Vol. IV. Text and Plates. 8vo, London, 1903.
By Exchange.
Herz (Otto). Verzeichniss der auf der Mammuth Expedition gesammelten
Lepidopteren.
[Ann. Mus. Zool. Acad. Imp., T. VIII.] 8vo, St. Petersburg, 1903.
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The Smithsonian Institution.
Hine (J. 8.). Tabanidze of Ohio.
[Ohio Acad. Sci., Sp. Papers, No. 5.] 8vo, Columbus, 1903.
The Author,
(Sexxy)
‘*Hopre Reports,” edited by Prof. E. B. Poulton, Vol. III, 1902. 8vo,
Oxford, 1903.
edited by Prof. E. B. Poulton, Vol. IV, 1900—03. 8vo, Oxford,
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Hovusert (C.). Les insectes ennemis des livres. 8vo, Paris, 1903.
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ILtipGk (R.) and Quarn (A.). Australian Wood-boring Cosside.
ie Roy. Soc. Queensland, 1903. | The Authors.
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Jacoby (M.). Descriptions of the New Genera and Species of Phytophagovs
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Jounson (W. W.) and Harperr (J. N.). <A List of the Beetles of Ireland.
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Kirpy (W, F.). A Handbook to the Order Lepidoptera, Vols. I—IV.
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LAMEERE (A.). Edmond de Selys Longchamps (Obituary notice).
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Marcuar (P.). Les Tarsonemus des Graminées,
[Bull. Soc. Ent. France, 1902. ]
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{ Bull. Brooklyn Inst. Arts and Sci., 1902. ] The Institute.
Morpivitxo (A.). Zur Biologie und Morphologie der Pflanzenliuse (Fam.
Aphididx, Pass.)
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Morey (Claude). The Ichneumons of Great Britain. 8vo, Plymouth,
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II. [Ray Society.] 8vo, 1903. Purchased.
( xxvi )
OsTEN-SackEN (C. R.). Record of My Life Work in Entomology. Svo,
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Packarp (A. 8.). Obituary Notice of Alpheus Hyatt.
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Hints on the Classification of the Arthropoda; the Group a
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The Author.
Perincuey (L.). Descriptive Catalogue of the Coleoptera of South Africa.
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[Trans. 8. African Phil. Soc., Vol. XII.] 8vo, London, 1902.
The Author.
Perkins (R. C. L.). The Leaf-Hopper of the Sugar Cane. 8vo, Honolulu,
1903. The Author.
Pocock (R. I.). [See Gopman (F. D.). Biologia Centrali-Americana. |
Quart (A.). [See IttipGe (R.).]
Ropztanko (W. N.). Some Observations on Parmene rediella, Clerck.
(Tortricina, Lepidoptera.) Svo, Moscow, 1903.
The Author.
RoruscHiLD (Hon. Walter) and JorDaAn (Karl), A Revision of the Lepido-
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[Novitates Zoologicae, Vol. IX, Supplement.
The Hon, Walter Rothschild,
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Saitu (J. B.). Contributions toward a Monograph of the Lepidopterous
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WateEruousE (G. A.). Catalogue of the Rhopalocera of Australia.
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FoR THE YEAR 1903.
I. A further contribution to our knowledge of African
Phytophagous Coleoptera, Part II By Marrin
JACOBY, F.E.S.
[Read October 15th, 1902. ]
THIS paper is a continuation of the one published in
these Transactions in 1901. It deals with the groups
Halticinze and Galerucine.
HALTICINE.
Haltica malvernensis, sp. 0.
Metallic green with brassy reflection, the basal three joints of the
antenne more or less fulvous, thorax impunctate, the basal sulcus
sinuate, extending to the sides, elytra strongly and closely punctured
in semiregular rows, the interstices slightly rugose, tibise rather
broad and deeply sulcate.
Length 4 millim.
Of a brassy green colour, the head impunctate, the frontal tuber-
cles very strongly raised, the carina very narrow, acutely raised and
elongate, labrum and palpi blackish, antenne extending rather
beyond the middle of the elytra, black, the lower three joints more
or less fulvous, the basal joint stained with w#neous above, third
joint one-half shorter than the fourth ; thorax about one-half broader
than long, distinctly narrowed anteriorly, the sides very feebly
TRANS. ENT. SOC, LOND. 1903.—PART I. (APRIL) 1
2 Mr. M. Jacoby on
rounded, the anterior angles thickened, obtuse, the dise convex and
shining, impunctate, the basal sulcus sinuate, not foveolate at the
sides but extending to the latter and placed at a proportionately
broad distance from the basal margin, scutellum blackish ; elytra
with very closely approached irregular rows of distinct punctures
which are getting smaller towards the apex, the interstices, especially
at the sides, rather uneven with some faint traces of longitudinal
costee below the shoulders, the latter prominent, the tibize of the male
rather strongly widened anteriorly and suleate, the male organ
slender, slightly narrowed at the middle, the anterior margin nearly
truncate, with a small central point, the upper surface suleate at
each side near the apex.
Hab. Malvern, NATAL (C. Barker) ; also Sedhion, WEST
AFRICA (C. Alluand).
The general colour of this species, of which I have
eight specimens before me, is not blue as is mostly the
case in this genus, but a brassy green, and the punctuation
of the elytra is proportionately strong and very close, and
the tibize (at least in the male) are more dilated and more
strongly sulcate than is usually the case, they may almost
be called triangularly compressed.
Haltica pyritosa, Krichs.
It is at present almost impossible to refer with certainty
any of the numerous species of Haltica from different
parts of Africa which are before me, to Erichson’s species
without the type for comparison ; I look, however, upon
specimens from Mozambique, Zambesi R., Mashonaland,
and West Africa as representing Erichson’s species. ‘The
author gives the colour as “neous”; all the specimens
before me are cupreous above, and the elytral punctuation
is comparatively strong and arranged in nearly regular
and closely- approached rows; the “male organ does not
differ from that of several other closely- -allied or perhaps
identical species, but all the specimens from the above
localities are decidedly of more elongate and_ parallel
shape than those from South Africa before me, and I am
able to separate them also by the sculpturing of the
elytra.
Haltica cuprea, Jac.
This species was described by me in the Trans. Ent.
Soc. London, 1895, founded on specimens of entirely
African Phytophagous Coleoptera. 3
cupreous colour above, and in which the elytral punctu-
ation is extremely fine and irregular as_ well as close;
numerous specimens since received from Natal differ in
having the thorax of a bright metallic brassy colour, but I
am unable to find any other differences of importance, nor
in that of the male organ; the latter is elongate and
parallel, the apex rounded with a small projecting point at
the middle, the upper surface below the anterior cavity is
finely transversely wrinkled ; the size of the insects differs
rather considerably, some being much larger than others,
but all are more oblong and convex than the species I
refer to H. pyritosa.
Haltica cyanicollis, Jac.
The description of this species was published at the
same time and in the same Journal as the preceding one,
but whether the insect is really specifically distinct or
only represents a local form I am unable to say, The size
is larger than the largest specimens of HH. cuprea, the
thorax is metallic blue or greenish, and the basal sulcus
is placed at a longer distance from the basal margin than
is the case in the allied species; other differences consist
in the strongly-punctured space in front of the eyes and
the more distinctly-punctured elytra.
Dibolia thoraciea, sp. n.
Black, the basal joints of the antenne and the legs fulvous, head
and thorax dark greenish, minutely transversely wrinkled, elytra
dark bluish, microscopically punctured, posterior femora black at
the apex.
Leneth 23-3 millim.
Head greenish, minutely granulate or wrinkled, eyes elongate,
occupying the entire sides of the head, frontal elevations absent,
clypeus broad, antennze not extending to the middle of the elytra,
the lower fou joints flavous, the others black, basal joint elongate,
the following three nearly equal, terminal joints short; thorax
nearly three times broader than long, the sides obliquely narrowed
anteriorly, the anterior angles thickened, the entire surface covered
with very fine transverse wrinkles, greenish, basal margin broadly
and evenly rounded, scutellum small, black ; elytra strongly convex,
evenly rounded and widened towards the middle, bluish, with some
extremely fine punctures, only visible under a strong lens, legs
flavous, the apex of the posterior femora black, posterior tibie
4 Mr. M. Jacoby on
widened near the apex and strongly sulcate with a broad emarginate
spur.
Hab. Frere, NATAL.
The colour of the thorax which differs from that of the
elytra, and the peculiar finely-wrinkled surface form the
principal characteristic characters of this species, of which
two exactly similar specimens are contamed m my
collection.
Dibolia abdominalis, sp. n.
Piceous below, the basal joints of the antennze and the abdomen
more or less testaceous, above obscure ceneous, eyes closely approached,
upper surface finely or scarcely perceptibly punctured, the tibize and
tarsi obscure flavons.
Length 3 millim.
Ovate and very convex, the head impunctate, greenish-zneous,
the eyes very large and closely approached at the vertex, the inter-
mediate space smaller than their diameter, antennz flavous, the apical
joints darker, second and third joint short, equal, the fourth the
longest ; thorax very short, three times broader than long, the sides
straight, the anterior angles thickened, the median lobe somewhat
pointed, the disc minutely transversely wrinkled, without punctures,
elytra very convex, widened towards the middle, minutely punctured
and obsoletely longitudinally sulcate, legs flavous, the posterior
femora and the breast piceous, abdomen partly flavous.
Hab, Grahamstown and Dunbrody, S. Arrica (Rev.
O Neil).
The punctuation in this species seems to vary from
extreme fineness to absence of punctures, but I am not
able to separate the specimens on account of any other
differences ; in most of them, the elytral punctuation is
very fine and closely placed, but in one specimen no trace
of punctures can be seen and the interstices are obsoletely
sulcate; in all, the eyes are approached at the vertex (in
one, probably the male, much more closely so), but the
space separating them gradually widens.
Dibolia bimaculata, Jac. (Proc. Zool, Soc., 1900).
Whether this species is identical with D. maculata, Har.
(Coleopter. Hefte, xvi. p. 232), or a variety of it I am not
able to say without seeing the type, but Harold describes
-
African Phytophagous Coleoptera. 5
the insect as obscure zwneous with the elytra blue pos-
teriorly only, and with the interstices of the elytral punc-
tured striz likewise punctured. In D. bimaculata the
first five joints of the antenne (not four) are flavous, the
entire upper surface is metallic-blue, and the elytra are
punctured in closely-approached rows of rather irregular
shape without extra punctures in the interstices. In other
respects Harold’s description is too short and gives no
other details.
Dibolia affinis, sp. n.
Blackish-blue, the antennae and legs black, thorax closely and
sirongly punctured, elytra very finely punctate-striate, each with a
large fulvous spot near the apex.
Length 3 millim.
Of elongate, subcylindrical shape, the head finely and sparingly
punctured, dark blue, eyes widely s2parated, the intermediate space
with a transverse groove, antenne black, the second, third and
fourth joints of nearly equal length, fifth slightly longer, four terminal
joints thickened ; thorax very strongly transverse, the sides rounded,
narrowly margined, the disc strongly, evenly and closely punc-
tured, the interstices slightly aciculate ; scutellum smooth ; elytra
subeylindrical, slightly narrowed posteriorly, finely punctate-striate,
the strie not very distinct, and irregular, each elytren with a bright
fulvous, nearly round spot near the apex ; legs black, the tibial spur
fulvous.
Hab. Bothaville, ORANGE FREE STATE (Dr. Brauns).
Again closely allied in coloration to the preceding species
but differing in its larger size, the entirely. black antenne
and legs, the strong punctuation of the thorax and the
impunctate elytral interstices. I received a_ single
‘specimen from Dr. Brauns in 8. Africa.
espera maculicollis, sp. 0.
Pale testaceous, finely pubescent, antennz fuscous, thorax finely
rugosely punctured, the sides with a fuscous stripe, elytra microsco-
pically punctured, the lateral inargins narrowly piceous, the breast,
the apex of the posterior femora and the posterior tibia and tarsi
piceous.
Length 2-3 millim.
Head finely rugose, obscure fulvous, frontal tubercles distinet but
rather flat, antenne extending beyond the apex of the elytra, fuscous,
the base of each joint pale, second joint small, third shorter than
6 Mr. M. Jacoby on
the fourth, rather widened as well as the following three joints,
terminal joints more elongate and slender; thorax nearly twice as
broad as long, the sides straight, the surface rather depressed, very
obsoletely transversely suleate at the middle, finely punctured and
pubescent, rather darker than tbe elytra, the sides with a narrow
fuscous stripe, scutellum piceous ; elytra pale testaceous, clothed
with pale fulvous pubescence, the interstices minutely punctured,
the lateral margins narrowly black, this colour however not quite
extending to the apex ; legs rather slender, tibize mucronate, the
metatarsus of the posterior legs as long as the following joints
together, claws appendiculate, anterior cavities open.
Hab. Malvern, Natat (C. Barker).
In the specimen, which I look upon as the female, the
antennz are more slender, without widened intermediate
joints, and the general size of the insect is larger.
Hespera pallida, sp. ua.
Pale testaceous, the antenne fuscous, the base of each joint pale,
upper surface pubescent, thorax transverse, minutely punctured and
wrinkled, elytra extremely finely punctured.
Length 3 millim.
Head scarcely perceptibly punctured, frontal tubercles small and
obsolete, antenne extending beyond the apex of the elytra, fuscous,
the base of each joint pale, the third joint slightly shorter than the
fourth, all the rest elongate and slender ; thorax twice as broad as
long, the sides straight, the anterior angles slightly tuberculate, the
surface minutely punctured and wrinkled and finely pubescent ;
elytra with a shallow depression below the base, sculptured like the
thorax, under-side and the legs pale flavous,
Hab, Grahamstown, Pt. Alfred, S. AFRICA (Licv. O'Neil).
This species scarcely differs from the preceding in
any other point except the coloration, in having no black
markings of any kind; there are, however, four specimens
before me which all agree with each other, and it is there-
fore unlikely that the insect is only a pale variety of HZ.
maculicollis.
Lutornus picturatus, sp. 0.
Black, thorax testaceous, with several small black spots, strongly
and remotely punctured, elytra very closely and finely punetured,
black, a round spot near the scutellum, a transverse band at the
middle, another semicircular band near the apex and the Jateral
margins, testaceous,
African Phytophagous Coleoptera. 7
Length 5-6 millim.
Head strongly punctured near the eyes, black, shining, with a
deep, concave groove between the eyes, clypeus very broad, convex,
antennee short, black, the second and fourth joint small and equal,
the third elongate, the others transversely widened ; thorax more
than twice as broad as long, the sides feebly rounded, broadly
flattened, the anterior angles thickened but not produced, basal
margin sinuate, preceded by a shallow transverse sulcus, the surface
irregularly and rather deeply punctured, testaceous, with two small
black spots at the middle near the anterior margin ; another more
indistinct spot is placed at the sides near the base; scutellum black ;
elytra very closely and distinctly punctured, the black colour
interrupted by a round spot near the scutellum, a transverse regular
band at the middle not quite extending to the suture and another
V-shaped band near the apex of each elytron, under-side and the legs
black, posterior femora strongly widened, their tibizee with a small
tooth near the apex, claw joint strongly inflated, apical segments
of the abdomen and the prosternum more or less testaceous.
Hab. Salisbury, MASHONALAND (G. Marshall).
Longitarsus barkert, sp. 0.
Apterous, ovate, black, shining, the antenne, the anterior and
intermediate legs and the posterior tibiee and tarsi flavous, thorax
impunetate, elytra very finely punctate-striate anteriorly, the
punctures obsolete near the apex.
Length 2 millim.
Head impunctate, the frontal elevations small and transverse,
bounded by a straight groove behind, the carina acute, antennze
extending beyond the middle of the elytra, flavous, the terminal
joints slightly fuscous at their apex, basal joint thick and sub-
eylindrical, the following three of equal length, fifth joint much
longer, apical ones slightly thickened ; thorax subquadrate, the sides
straight, the anterior angles strongly oblique, forming a slight tubercle
before the middle, the dise rather convex, impunctate, black, shining,
scutellum strongly transverse ; elytra ovate, convex, distinctly widened
at the middle, the apex of each rounded, the punctuation very fine and
arranged in irregular rows anteriorly, irregular and scarcely per-
ceptible near the apex ; under-side and the posterior femora black,
the rest of the legs flavous, metatarsus of the posterior legs as long as
the following joints together.
Hab. Malvern, Nata (C. Barker).
Smaller than Z. nigritula, the antenne entirely flavous,
8 Mr. M. Jacoby on
the head differently sculptured, and the elytra more
distinctly punctured in rows.
Longitarsus apricupes, sp. 0.
Apterous, ovate, convex, black, the basal joints of the antenna,
the apex of the tibize and the tarsi, fulvous, thorax impunctate,
elytra strongly punctured in irregular double rows.
Length 2 millim.
Head inipunctate, black, shining, obliquely grooved above the
eyes, frontal elevations indistinct, carina long and distinct, antennz
rather short, the lower five or six joints flavous, the rest black, the
basal joint black above, the second and the following two joints
equal in length, the fifth, longer ; thorax transversely subquadrate,
convex, the sides straight, the anterior angles oblique, the surface
impunetate ; elytra gradually rounded and widened towards the
apex, strongly convex, the shoulders not prominent, the punctuation
comparatively strong, arranged in irregular double rows which get
gradually finer towards the apex, the lateral margins accompanied by
a distinct single row of punctures, under-side and the legs black,
the extreme apex of the tibia and the ta‘si fulvous, the metatarsus
of the posterior legs as long as the following joints together,
Hab, Dunbrody, Cape CoLony (Rev. O' Nei/).
The shape of this species differs from many others of
this genus in the posteriorly-widened elytra, which attain
the greatest width at the apex instead of the middle; the
sculpture of the same parts and the colour of the legs are
other distinguishing features of this species.
Longitarsus dunbrodensis, sp. 0.
Apterous, elongate and narrow, dark greenish-neous, the basal
joints of the antennw, the base of the femora and the tibice and tarsi
fulvous, head and thorax minutely granulate and punctured, elytra
truncate at the apex, sculptured like the thorax.
Length 2 millim.
Head rather broad, without any frontal elevations, dark greenish,
lower portion of the face obscure fulvous, antennee extending beyond
the middle of the elytra, the lower six joints fulvous, the others
black, the second and third joint equal in length, the terminal joints
distinctly thickened, subquadrate ; thorax scarcely one-half broader
than long, subquadrate, the sides straight, or nearly so, the anterior
angles slightly obliquely thickened, the dise rather flat, extremely
minutely and closely punctured, not very shining, seutellum broad,
African Phytophagous Coleoptera. 9
bluish black, impunctate, elytra slightly widened towards the middle,
the apex of each almost truncate, the punctuation similar to that of
the thorax ; posterior femora eeneous at the apex, the base fulvous as
well as the other legs and the posterior tibize, the first joint of the
posterior tarsi as long as the following joints together.
Hab. Dunbrody, CAPE CoLony (Lev, O'Nei/).
This species is of a more peculiar shape and depressed
appearance than any of its allies from the same country ;
the absence of any frontal tubercles, the extremely fine
punctuation of the upper surface, the truncate elytra and
comparatively short metatarsus will assist further in its
recognition.
Aphihona barkert, sp. 1.
Ovate, black, thorax transverse, closely punctured, elytva very
closely, strongly and irregularly punctate, posterior femora short
and very thick, their tibice with a strong spur.
Length 3 millim.
Head broad, nearly impunetate, shining, the frontal elevations
broadly transverse, bounded behind by deep grooves, the carina
short, antenne extending beyond the middle of the elytra, black, the
second and third joint short, more or less flavous, the fourth and
fifth as long as the preceding two joints together, the following
slightly shorter ; thorax transversely subquadrate, twice as broad as
long, the sides feebly rounded, the anterior angles thickened and
somewhat oblique, the posterior ones rather obtuse, the sides with
a narrow margin, the surface rather convex, closely and strongly
punctured, the punctures irregularly placed, somewhat more closely
at the sides than at the middle, scutellum broad, triangular, im-
punctate, elytra much wider at the base than the thorax, much more
strongly and more closely punctured than the latter, under-side and
the legs black, the posterior femora strongly thickened, ovate, the
first joint of the posterior tarsi as long as the following two joints
together.
Hab. Malvern, NATAL (C. Barker)
This species is well distinguished by its entirely deep
black and shining colour and the strong and close elytral
punctuation; the structural characters all agree with
Aphthona; the anterior tibie have a very small spine, the
claws are appendiculate and the anterior coxal cavities are
open; the prosternum is very narrow. Iam not able to
say anything with regard to the male organ as I seem to
have only female specimens before me.
10 Mr. M. Jacoby on
Aphthona similis, sp. n.
Black, shining, the lower four joints of the antennz flavous, thorax
transverse, finely and sub-remotely punctured, elytra more strongly
punctured in closely approximate rows.
Length 3 millim.
Head impunetate, shining, black, the frontal tubercles obliquely
transverse, the carina acutely raised, antennae extending to about
the middle of the elytra, black, the second to the sixth joint flavous,
the second and third joints short, equal, terminal joints slightly
thickened and shortened ; thorax twice as broad as long, with the sides
feebly rounded, the anterior angles thickened and slightly obliquely
produced, and the surface very distinctly but not very closely
punctured; scutellum broad, impunctate, elytra scarcely more strongly
punctured than the thorax, the punctures arranged in closely approxi-
mate rows, distinct to the apex, under-side and the legs black, the
first joint of the posterior tarsi as long as the following three joints
together.
Hab. Malvern, Nata (C. Barker).
At first sight this species exactly resembles A. barkeri,
but it is quite «distinct, the thorax has not the narrow
Jateral margin of the allied species, and the sculpturing of
the upper parts is different and much less strong, the
antenne have the second and the following five joints
flavous, and the metatarsus of the posterior legs is elongate.
I have received three specimens from Mr. Barker.
Aphthona senegalensis, sp. n.
Flavous, shining, the apical joints of the antenne and the posterior
femora piceous, head and thorax impuncetate, the sides of the latter
straight, elytra not perceptibly punctured.
Length 33 millim.
Of broadly ovate shape, the head impunctate, the frontal tubercles
small but broad, deeply grooved behind, the clypeus very broad, the
space between the antennze but little constricted, the labrum and
palpi obscure piceous, antennae not extending to the middle of the
elytra, piceous, the lower four joints flavous, third and fourth joints
equal, not much longer than the second one, the apical joints slightly
thicker and scarcely shorter, thorax about one-half broader than
long, the sides straight, the anterior angles oblique, the surface
impunctate with an obsolete longitudinal groove near the lateral
margins, elytra smooth and impunctate, flavous, under-side of the
latter colour, the posterior femora piceous, their tibize strongly widened
African Phytophagous Coleoptera. 11
towards the apex, with a strong spur placed at the outer edge, the
first joint of the posterior tarsi as long as the following three joints
together ; abdomen finely pubescent.
Hab. SENEGAL.
A comparatively robust and large species, distinguished
by the straight lateral margins of the thorax and the
impunetate upper surface; the prosternum is extremely
n-rrow, and the legs with the exception of the posterior
femora are of entirely flavous colour,
Aphthona bimaculata, sp. n.
Head and the under-side obscure piceous, the basal joints of the
antenne and the thorax pale fulvous, the latter finely punctured,
elytra pale flavous, very closely and distinctly punciured, the
suture, the sides and an elongate spot below the middle, piceous.
Length 3 millim.
Head impunctate, the vertex nearly black, the lower portion fulvous,
frontal tubercles obsolete, clypeus thickened, very broad at the base,
pale fulvous, antennse short, not extending to the middle of the
elytra, black, the lower four joints fulvous, the second joint thicker
but not-shorter than the third one, terminal joints slightly thickened,
about one-half longer than broad ; thorax transverse, twice as broad
as long, the sides very feebly rounded, anterior angles slightly
oblique, the dise with a very obsolete transverse groove near the
base, remotely and finely punctured, the punctures rather shallow,
the surface pale fulvous, stained with some very obsolete darker
spots, scutellum piceous ; elytra pale flavous, very closely and finely
punctured with an obsolete short longitudinal costa below the
shoulders, the suture and lateral margins narrowly piceous, each
elytron with another short piceous longitudinal stripe below the
middle, under-side and the posterior femora piceous, tibiee and tarsi
fulvous, the first named widened towards the apex, armed with a
distinct spine ; the spine of the posterior tibize is placed at_the
outer margin ; prosternum narrow, coxal cavities open.
Hab. Malvern, Nata (C. Barker).
It will not be difficult to distinguish this proportionately
large and well-marked species, of ‘which I received a single
specimen from Mr. Barker.
Podagrica vmpressipennis, sp. 0.
Ovately rounded, very convex, black, the basal joints of the
antenne, the head, thorax and legs fulvous, thorax transverse,
impunctate, elytra extremely finely and irregularly punctured with
12 Mr. M. Jacoby on
a short transverse depression below the shoulders impressed with some
deeper punctures.
Length 45 millim.
Of the shape of a species of Sphaeroderma ; the head impunctate,
with a transverse groove between the eyes, frontal elevations small,
clypeus rather widened between the antenne, palpi thickened, the
apical joint conical, antennie filiform, the lower six and part of the
seventh joint pale fulvous, the rest black, second and third joint
short, equal, the following elongate ; thorax twice as broad as long,
the sides straight at the base, rounded from the middle to the apex,
the anterior angles blunt, the median lobe of the posterior margin
nearly straight at the middle, each side of the same margin provided
with a small notch, the surface entirely impunetate, reddish-fulvous,
seutellum black, elytra wider at the base than the thorax, strongly
convex, finely punctured in irregular rows, with a short but deep
transverse depression below the shoulders within which the com-
mencement of deeper rows of punctures are placed, apex of the
elytra nearly impunctate, legs rather robust, fulvous, the first joint
of the posterior tarsi as long as the following two joints together,
anterior coxal cavities closed, prosternum elongate and narrow.
Hab. Malvern, NATAL (C. Barker).
This species has entirely the shape and coloration of
P. indica, Fab., but differs in the short second and third
joints of the antenne and the deeply punctured elytral
depression. I received a single specimen from Mr.
Barker; another one in my collection has only the first
four joints of the antennze fulvous but agrees in all other
respects.
Crepidodera carinipennis, Sp. 1.
Fulvous, the outer joints of the antennie black, thorax with
very deep and broad suleus, finely punctured, elytra very closely
and rather strongly punctate-striate, metallic dark green, with an
acute ridve from the shoulders downwards.
Mus. Antenne long and robust, the basal joint strongly thickened.
Fem. Antenne shorter and thinner.
Var. Thorax and elytra dark blue.
Length 4 millim.
Elongate and parallel, the head finely wrinkled at the vertex, the
frontal elevations very strongly raised, subquadrate, clypeus triangu-
larly raised, antennie black, the lower four joints fulvons, the basal
joint strongly thickened and subquadrate in the male insect, fourth
African Phytophagous Coleoptera. 15
joint very slightly longer than the third ; thorax one-half broader
than long, the sides straight, the anterior angles obliquely thickened
and produced outwards, the surface convex, finely and sparingly
punctured, the basal sulcus very broad, bounded at the sides by an
acute, sightly-curved ridge, the suleus with some deeper punctures
anteriorly, scutellum black; elytra wider at the base than the
thorax, the base slightly raised, the disc very closely punctured in
rows, bluish green, the shoulders prominent and joined by an acute
ridge which extends nearly to the apex ; this ridge is preceded by a
more feebly raised one near the apex, and the space between the outer
ridge and the lateral margin is concave ; legs robust, fulvous, all the
femora thickened, the intermediate tibiw dilated at the apex.
Hab. Drakensberg, Nata (@. Marshall); Malvern,
NATAL (C. Barker).
This is rather an aberrant species, the male of which
resembles much in shape and colour the Central American
genus Plectotetra especially in the structure of the antenne.
The sculpturing of the elytra will at once distinguish
the species. The single specimen sent by Mr. Barker
only differs in the entirely blue upper surface and the
less strongly developed elytral costze.
Crepidodera marshall, sp. n.
Obscure piceous below, the basal joints of the antenna, the head
(the vertex excepted) thorax and the legs fulvous, elytra very
strongly and closely punctate-striate, metallic green.
Length 3 millim.
Head rather elongate, impunctate, the vertex metallic green,
frontal elevations broad and strongly raised, carina acute, antenne
extending beyond the middle of the elytra, black, the lower four
joints fulvons, the third joint one-half longer than the second one ;
thorax scarcely twice as broad as long, rather convex, the sidesrounded,
narrowly margined, anterior angles thickened, the base with a deep
transverse groove which does not extend to the lateral margins, the
entire surface impunctate, scutellum black; elytra slightly wider at
the base than the thorax, the shoulders rather prominent, the base
slightly raised, the punctuation very strong and closely arranged in
not very regular double rows here and there, finer at the apex,
legs and the extreme apex of the abdomen more or less fulvous ;
posterior tibia with a small spine ; prosternum rather narrow
Hab. Drakensberg, Nata (G. Marshall).
14 Mr. M. Jacoby on
The general coloration of this little species agrees with
that of many of its allies, but the metallic-green vertex
of the head and the strong and close elytral punctuation
distinguish the species from any of its African congeners.
Crepidodera uniforms, sp. 0.
Flavous, the apical joint of the antenne infuscate, thorax trans-
verse, impunctate, the basal sulcus deep, elytra strongly punctate-
striate, the interstices finely punctured, flavous, the sutural margin
narrowly obscure piceous.
Length 35 millim.
Head impunctate, of darker colour than the re-t of the upper
surface, obliquely grooved between the eyes, without any frontal
tubercles, the clypeus broad and flat, impunctate, antennee about
half the length of the body, flavous, the apical joint fuscous, the
second one about one-half shorter than the third, terminal joints
slightly thickened ; thorax twice as broad as long, narrower in
front than at the base, the sides rounded, the angles distinct but
not produced, the surface impunctate, with a deep transverse sulcus
at the base, bounded at the sides by a perpendicular groove which
extends upwards a little way beyond the sulcus, scutellum triangular,
flavous ; elytra not wider at the base than the thorax, rather strongly
punctate-striate, the punctures much finer at the apex, the interstices
extremely finely punctate, only visible under a strong lens, the
sutural margins narrowly piceous, under-side and the legs flavous,
the metatarsus of the posterior legs as long as the following three
joints together.
Hab. Malvern, Nata (C. Barker’).
I have received two specimens of this species which
will be easily recognized by the general flavous coloration,
the dark elytral suture, and the absence of any frontal
elevations.
Lypnea africana, sp. n.
Elongate, flavous, the outer joints of the antenne and the tibia
and tarsi black, thorax transverse, nearly impunctate, the base with
a transverse sulcus, elytra finely punctate-striate,
Length 5 millim.
Head impunctate, the frontal elevations nearly contiguous, trans-
verse, antenne filiform, black, the lower two joints flavous, third
joint one-half longer than the second one, terminal joint more
slender and elongate ; thorax about one-half broader than long, the
sides very feebly rounded, the anterior angles oblique, the base with
African Phytophagous Coleoptera. 15
a shallow transverse sulcus, bounded at the sides by a short perpen-
dicular groove, the surface with a few minute punctures, shining,
flavous ; elytra slightly wider at the shoulders than the thorax,
elongate, subcylindrical and parallel, finely punctate-striate, the
interstices flat and impunctate ; under-side and the femora flavous,
antertor and intermediate tibise unarmed, posterior ones with a
minute tooth, the first joint of the posterior tarsi as long as the
following joints together, claws appendiculate, anterior coxal cavities
open.
Hab. Umtali, MASHONALAND (G. Marshall).
I can find no characters of any importance to separate
this species generically from Baly’s Eastern genus Lypnea,
with which it has not only the structural characters in
common but also the coloration, except that of the tibiz
and tarsi; the elytral punctuation also is very fine, not
strong. Poéphila, Weise, differs in_ the finely pubescent
eyes, ‘the quadrate thorax, and the shape of its sulcus and
in the longer metatarsus; the present is the first African
representative of this genus,
Livolia, gen, n.
Body narrowly elongate, glabrous, head broad, the frontal tubercles
obsolete, eyes small, antennie filiform, the terminal joints slightly
thickened, thorax subquadrate, constricted at the base, the surface
with a transverse groove near the base, extending to the lateral
margins, elytra wider at the base than the thorax, punctate-striate,
posterior femora very moderately thickened, their tibiae with a
small spine, the metatarsus scarcely as long as the following two
joints together, claws appendiculate, prosternum narrow between the
coxee, the anterior cotyloid cavities closed.
This genus is proposed for a very small species of
Halticide, having the appearance of a species of Corticaria
and affinities with Crepidodera; the very obsolete frontal
tubercles of the head, the thoracic sulcus which is placed
close to the basal margin and extends to the sides, as well
as the but slightly thickened posterior femora and un-
armed anterior tibiz separate the genus from the last-
named one and those allied to it; it seems a transitional
form between the Halticine and Galerucine.
Livolia sulcicollis, sp. n.
Fulvous, the breast and abdomen black, thorax strongly and
16 Mr. M. Jacoby on
remotely punctured, elytra finely punctate-striate, the interstices
impunctate.
Length 2 millim.
Head broad, impunctate, fulvous, shining, the eyes small, widely
separated, clypeus wide at the base, palpi thin and slender, antenne
half the length of the insect, fulvous, the basal joint strongly and
suddenly thickened at the apex, the second joint longer than the
third, the latter and the fourth equal, the terminal joints thickened,
thorax about one-third broader than long, the sides rounded and
widened at the middle, the base distinctly narrower than the anterior
portion, the angles acute, the basal suleus narrow but deep, the dise
rather convex, strongly but remotely punctured, scutellum broad,
impunetate ; elytra wider at the base than the thorax, rather
depressed, finely punctate-striate, the punctures closely placed,
elytral epipleure broad ; under-side black, legs fulvous, tibize shghtly
widened towards the apex, non-sulcate, the four anterior femora
searcely less widened than the posterior ones, the first joint of the
anterior tarsi of the male, dilated.
Hab. Salisbury, MASHONALAND (G, Marshall).
I have received three specimens of this curious little
species from Mr. Marshall.
Weiseana barkert, sp. 0.
Under-side nearly black, upper-side obscure testaceous, finely pube-
scent, terminal joints of the antenne and the head fuscous or piceous,
thorax with three piceous spots (more or less connected), elytra
opaque, narrowly margined with black, legs testaceous, tarsi piceous.
Length 33-4 millim.
Head rugose, the vertex blackish, the clypeus strongly triangularly
convex, flavous, labrum piceous, margined with testaceous, antennze
robust, blackish, the lower three or four joints testaceous at the
base, third joint slightly longer than the fourth, the folowing joints
thickened ; thorax rather more than twice as broad as long, the
sides nearly straight, the angles obtuse, the disc with three obsolete
depressions, rugosely punctured, the interstices minutely granulate,
the surface with a central marking and two lateral black markings,
sometimes connected in shape of /\ ; scutellum fuscous; elytra clothed
with very fine silky pubescence, opaque, very finely transversely
wrinkled, without distinct punctuation, obscure testaceous, the sutural
and lateral margins narrowly black ; legs testaceous, the femora
stained with fuscous as well as the apex of the tibiz and the tarsi,
claws with a basal tooth.
African Phytophagous Coleoptera. iki
Hab. Malvern, Natau (C. Barker).
I received four specimens from Mr, Barker.
GALERUCIN A.
Genus Diacantha, Chev.
I cannot agree with Weise in his definition of this and
the allied genera (Deutsche Ent. Zeitsch. 1901, p. 274).
He has evidently overlooked the type D. bispinosa, Oliv.
Chevrolat in his diagnosis of the genus in D’Orbigny,
Dict. Univ. d’Hist. Naturelle, p. 718, iv, quotes 12 species
mentioned by Dejean in his catalogue. These are—D.
picea, Fab.; spinosa, Oliv.; (bispinosa) festiva, Dalm. ;
unifasciata, Oliv.; 11 punctata, De}. ; Dreget (6 pustulata
= tricincta, Chev.) ; fuscitarsis, gloriosa, testudinaria, De}. ;
unipunctata, Chev., et generosa, De}.
Of all these only D. bispinosa and D. dreger belong to
Dacantha as defined by Chapuis, and the first named
must undoubtedly be taken for the type. In this species
the claws are bifid and the male has two elytral tubercles
at the base; most of the other species named above belong
to entirely different genera, thus picea is the genus
Stenoplatys, Baly, 11 punctata, De}j., is an Awlacophora, ete.
Harold gives D. bidentuta, Fab., as the type of the genus
Diacantha, but as this is not one of the species mentioned
either by Chevrolat or Dejean, it cannot be looked upon
as the type. Chapuis was therefore perfectly right when
he established the genus Hyperacantha, in which the
claws are appendiculate, and his genus must be accepted.
Weise has not said which species he looks upon as the
type, and has not mentioned the only true Diacantha
of Chevrolat’s list D. bispinosa. A typical specimen of
this species is contained in the British Museum, and J am
indebted to Mr. Gahan for his examination of the specim:n
and the other details given above concerning the genus.
Hyperacantha bituberculata, Fab.
According to Mr. Gahan the specimen contained in the
British Museum is certainly the typical bitwberculata, Fab.,
and ramed so by Dejean himself and also by Chevrolat.
Weise expresses a doubt that I had this species before
me when I remarked on it in Novitates, 1894, and gives
the last abdominal segment of the 2 with three deep
TRANS. ENT. SOC. LOND. 1903. PARTI. (APRIL) 2
,
18 Mr. M. Jacoby on
incisions; this is a mistake, as in HZ. bitwherculata this part
is ce very slizhtly concave, and Weise’s insect must
represent another species. In the British Museum speci-
men the elytral fulvous margin is only just visible.
Hyperacantha abdominalis, Duy., C. Rend. Belg. Entom.
Soc. (1891, July).
Hyperacantha abdominalis, Jac., the Entomologist (1891,
May).
On account of my name having the priority of two
months, I alter that of Duvivier’s to H. duvivieri.
Hyperacantha abdominalis, Jac.
In the Entomologist for 1891, I have described this
species from specimens obtained at the Transvaal and
other localities in Africa. The typical form has a reddish
fulvous upper surface, the elytra are narrowly margined
with black, and have a transverse black band at the middle,
the under-side is black with the exception of the flavous
last segment. I have lately received other specimens
from the Umkomaas Mountains in Natal, obtained by Mr.
Guy Marshall, which Iam quite unable to separate from
the type structurally, although one specimen only resembles
it in the coloration of the upper surface ; in this specimen,
however, the under-side is entirely flavous; the others
represent the following varieties.
Var. (a) Elytra margined with black at the anterior half only,
the transverse band narrower, not extending to the suture, under-
side flavous, general size much smaller.
Var. (b) Elytra with a small black stripe at the lateral margins
from the base to the shoulders and a small spot at the middle of
each elytron, under-side and the femora black.
Var. (c) Upper- and under-sides entirely flavous.
In all these varieties the tibia have the apex black
and the tarsi are entirely of that colour as is the case in
the type. The insect seems to be subject to a very great
amount of variation in regard to colour and also to size;
the general shape is broadly ovate and convex. I cannot
find any differences in the antenne or the sculpture.
Hyperacantha apicipes, sp. 0.
Broadly oblong, testaceous, the antennie (the basal joints excepted)
the breast, the apex of the tibie and the tarsi black, thorax strongly
African Phytophagous Coleoptera. 19
transverse and sulcate, impunctate, elytra finely and closely punc-
tured, black.
Length 6 millim.
Of broadly ovate and convex shape, the head impunctate, testace-
ous, strongly transversely grooved between the eyes, the carina
narrowly acute, palpi piceous, antenne slender, the third and
following joints elongate and nearly equal, terminal joints more
slender, the basal two testaceous, the rest, black ; thorax twice as
broad as long, the sides nearly straight, the anterior angles slightly
produced outwards but blunt, the disc impunctate, with a deep and
straight transverse sulcus near the middle, pale testaceous, scutellum
black ; elytra very convex, widened towards the middle, extremely
finely and rather closely punctured, black and shining ; under-side
and the legs testaceous, the breast, the apex of the tibice and the tarsi
‘black, the tibize all mucronate, the claws appendiculate ; the last
abdominal segment of the male trilobate, the median lobe with a
deep fovea.
Hab, Ulundi, Navan (@, Marshall).
A species of broadly ovate shape of which I received
two specimens, one of which showing traces of paler spots
on the elytra, probably due to immaturity.
Hyperacantha militaris, sp. n.
Testaceous or fulvous, the terminal joints of the antennie, the
apex of the tibie and the tarsi black, thorax transverse, impunctate,
deeply sulcate at the sides, elytra impunctate, with basal depression,
black, the basal third portion fulvous.
Length 6 millim.
Head impunctate, the vertex fulvous, the lower portion generally
paler in colour, frontal elevations transverse, carina acute, antenne
black, the lower three joints more or less testaceous, the third joint
slightly longer than the fourth ; thorax twice as broad as long, the
sides feebly rounded anteriorly, slightly constricted at the base, the
dise impunctate, with a rather deep transverse sulcus at each side,
scutellum fulvous or testaceous, elytra widened posteriorly, the
lateral margins flattened and widened below the shoulders, the base
with a distinct transverse depression, fulvous or testaceous, this
colour occupying the basal third portion, the rest of the surface
black ; under-side, the femora and the base of the tibize testaceous,
their lower portion and the tarsi black, the latter appendiculate, all
the tibize mucronate.
Hiab. Dar es Salaam, East Arrica; also ZANZIBAR
and DELAGOA Bay.
20 Mr. M. Jacoby on
This is one of the smaller species which is well dis-
tinguished by its coloration ; in some specimens, however,
the black elytral portion includes a larger or smaller spot
or space of the fulvous ground-colour; in the male, the
last abdominal segment has the median lobe of transversely
subquadrate and flattened shape.
Luperodes nigrotibialis, sp. 0.
Oblong, very convex, the antenna, tibia and tarsi and the under-
side black, upperside testaceous, thorax subquadrate, impunctate like
the elytra.
Length 5 millim.
Head impunctate, pale fulvous, shining, frontal tubercles strongly
raised, trigonate, carina broad, convex, labrum black, antennae ex-
tending below the middle of the elytra, black, the base of each
joint more or less fulvous, second one one-third lower than the
third, the latter about half the length of the fourth joint ; thorax
scarcely broader than long, convex, the sides sightly rounded before
the middle, the anterior angles thickened, the surface entirely im-
punctate, testaceous; elytra scarcely wider at the base than the
thorax, very slightly widened below the middle, impunctate, their
epipleure continued below the middle ; under-side black, the femora
flavous, tibize with a long spine, black like the tarsi, the metatarsus
of the posterior legs rather longer than the following joints together,
anterior coxal cavities open.
Hab. Grahamstown, 8. AFrrica (Rev. O'Neil).
Of less than half the size of Z. sulphuripennis, Jac., and
with black antenne tibiz and tarsi, but otherwise similar
to the last-named species.
Luperus dunbrodyensis, sp. 0.
Narrowly elongate, subdepressed, flavous, thorax subquadrate,
nearly impunctate, elytra extremely minutely punctured, rather
flattened.
Length 24-3 millim.
Head impunctate, the vertex more or less obscure piceous, shining,
frontal tubercles strongly raised, transverse, carina small but distinct,
antennie entirely flavous, extending below the middle of the elytra,
the third joint one-half longer than the second one ; thorax one-
half broader than long, the sides slightly rounded at the middle,
with a very narrow margin, anterior angles slightly thickened, not
produced, the disc only perceptibly punctured when seen under a
African Phytophagous Coleoptera. 21
strong lens, the sides with an obsolete fovea, scutellum flavous ;
elytra somewhat flattened, very finely but distinctly punctured, the
suture slightly infuscate, legs flavous, all the tibive mucronate, the
metatarsus of the posterior legs as long as the following joints
together,
Hab. Dunbrody, CAPE CoLony (fev. O'Nei/).
Of this species I received a male and female specimen
from the Rev. O'Neil, taken “in cop.” ; the first-named sex
is smaller than the female, the under-side is stained with
piceous, and the last segment of the abdomen is rather
deeply suicate. The species is very closely allied to Z,
weiser, Jac., and LZ. verticalis, Jac., but differs from both in
being of narrower, smaller, and more depressed shape, and
in the entirely flavous antennz ; the sides of the thorax in
L. weiset are also more strongly constricted at the base,
and the disc is more convex and without any depressions.
Luperus tugelaensis, sp. D.
Black, the antennee very long, their basal three joints, the thorax
and the anterior legs, flavous, thorax impunctate, elytra minutely
punctured, flavous, narrowly margined with black.
Length 4 millim.
Narrow and elongate, the head black, impunctate, the clypeus,
labrum and the palpi flavous, frontal elevations strongly raised,
broad, antennie extending to the apex of the elytra, black, the lower
three joints and the apex of the fourth and fifth joint flavous, the
third longer than the second but much shorter than the fourth joint ;
thorax scarcely broader than long, the sides straight at the base,
slightly rounded before the middle, the angles distinct, the surface
impunctate, flavous, scutelluin black ; elytra slightly wider at the
base than the thorax, very finely and closely punctured, margined
with black, the suture more broadly so ; under-side and the legs black,
the anterior ones flavous, all the tibie mucronate, the metatarsus of
the posterior legs as long as the following joints together.
Hab. Upper Tugela, Nata (C. Barker).
Of this distinctly-marked species I received a single
specimen (4) of Mr. C. Barker; the protruding apex “of
the penis is straight and acutely pointed.
Luperus malvernensis, sp. n.
Below black, above testaceous, clypeus, thorax and the femora
flavous, antenne piceous, thorax subquadrate, with a shallow lateral
yap Mr. M. Jacoby on
fovea, minutely punctured, elytra very closely and finely punctured,
the suture narrowly fuscous.
Length 3-4 millim.
Head black at the vertex, impunctate and shining, the frontal
tubercles transverse, strongly raised, carina acute, flavous lke the
elypeus, labrum and palpi black, antennie rather long and slender,
piceous, the third but slightly longer than the second but much
shorter than the fourth ; thorax but little broader than long, flavous,
very minutely g ssa es and punctured, with an obsolete depression
at each side, lateral mareins feebly rounded at the middle, anterior
angles thickened, sentellum black ; elytra wider at the base than
the thorax, very feebly transversely depressed below the base, ex-
tremely closely and finely punctured, obscure testaceous (sometimes
fuscous) with the suture narrowly piceous or fuscous ; under-side
black, the femora flavous, metatarsus of the posterior legs as long
as the following joints together.
Hab. Malvern, Natau (C. Barker); Dunbrody, CAPE
CoLony (fev. O'Neil),
This Luperws may easily be mistaken for LZ. cnconspiewus,
Jac., from Mashonaland, with which it has the same colora-
tion in common, but in that species the vertex of the
head is finely eranulate and not shining, the clypeus is
black and very “broad as well as the frontal elevations,
the thorax is more transversely shaped, and the legs are
generally entirely dark coloured.
Apophylia, Chev. (nee Chapuis).
This genus has been entirely mistaken by Chapuis,
Allard, and myself. The description given by Chapuis
of Apophylia does not apply to the type at all; the latter
is the .A. chloroptera, Thoms., who is the first who char-
acterized the genus, and that insect belongs to the genus
known at present as Malaxia, Fairm. = G Lyptolus, Jac. This
latter name, therefore, cannot stand,.and all the species
placed in it must be placed in Apophylia, A. tricolor, Fab.,
and others must receive another generic name as they do
not belong to Apophylia. It is difficult to know what
insect Chapuis had before him when he drew up _ his
description of the genus; he gives as the type A. smarag-
dina, De}. (a catalooue name), and says that the anterior
and intermediate tibixe are mucronate; but this isan error,
African Phytophagous Coleoptera. 23
as A. smaragdina, Dej., undoubtedly is also a Malaaia in
which the tibize are all unarmed. I have seen specimens,
named by Chevrolat A. smaragdina, which certainly re-
present the last-named genus. This is also the case with
A, murina, Gerst., and A. nobilitata, Gerst., of which a
figure is given and which belong to Malaxia. I am
indebted to Mr. Gahan for his assistance in clearing up
the above synonyms.
Pseudapophylia, gen. n.
Elongate, antennz filiform, thorax transverse and short, with an
obsolete transverse depression, scutellum broad ; elytra elongate and
parallel, finely rugose, their epipleurze broad anteriorly, indistinct
below the middle; legs robust, the anterior and intermediate tibice
with a small spine, the posterior ones unarmed, their metatarsus as
long as the following joints together, claws appendiculate, prosternum
invisible between the coxe, the anterior cavities open.
Type. P. smaragdipennis, Jac. (sub Apophylia).
I am obliged to place this species in a special genus, as
it has nothing to do with Apophylia (now Palewophylia),
under which name I described it (Trans. Ent. Soc. 1888) ;
the thorax differs quite in shape, being transverse and
short as well as obsoletely sulcate. It was perhaps this
species which Chapuis referred to Apophylia.
Paleophylia, gen. n.
Antenne filiform, the second to the fourth joint gradually
lengthened, eyes entire, thorax transverse, the sides, the posterior
angles and margin rounded, the surface rather convex, without de-
pression, elytra generally metallic, finely rugosely punctured, their
epipleurze very broad, gradually narrowed towards the apex, the
anterior and the intermediate tibiz mucronate, the posterior ones
unarmed, the metatarsus of the posterior legs as long as the follow-
ing two joints together, claws appendiculate ; prosternum invisible,
anterior coxal cavities open.
Type. LP. tricolor, Fab.
In this genus will have to be placed, besides the type
(the oldest described species) the Apophylia tricolor of
Gemminger’s Catal., the following species, all described
under Apophylia—P. berret, Alld.; P. maculicollis, Alld.; P.
bipunctata, Alld.; P. viridiniteus, Alld.; P. nigritarsis, Jac,
24 Mr. M. Jacoby on
Whether the other species described by Allard belong to
the same genus, is doubtful. Of those mentioned above,
however, I was able to compare the types, so that there
is no doubt about it. A. marginata, Jac., has, as 1 now
find, entirely unarmed tibie and elytral epiplearee which
are absent below the middle; this insect must therefore
find another place, although the general appearance and
structure is that of the genus here proposed.
Paleophylia granilosa, sp. v.
Metallie green below, the antenne and legs fulvous, above less
shining, minutely granulate, thorax subquadrate, the extreme lateral
margins fulvous, elytra minutely punctured and granulate.
Length 5 millim.
Head entirely impunctate, the frontal tubercles trigonate, strongly
raised and shining, carina short but distinct, labrum flavous, antennwe
extending beyond the middle of the elytra, fulvous, the second joint
half the length of the third, fourth and following joints slightly
longer than the third; thorax scarcely one-half broader than long,
the sides rounded, the angles obtuse, the surface rather convex,
without impressions, bright green, minutely granulate, scutellum
impunctate ; elytra parallel, sculptured like the thorax but with some
minute punctures ; underside more shining, green, the legs fulvous,
the anterior and intermediate tibize with a small spine, posterior ones
unarmed, the first joint of their tarsi as long as the following two
joints together, anterior coxal cavities open.
Hab. Upper Tongaat, Upper Tugela, Navan (C. Barker).
Of this species I received several specimens which may
be known by the silky green and finely granulate upper
surface, and by the narrow flavous margin of the thorax ;
the last segment of the abdomen in the male has a longi-
tudinal suleus of fulvous colour; the antennze in the
female are shorter, and the abdominal sulcus is absent.
I cannot identify this species with A. Duvivieri, Alld.
(Comptes-rendus Ent. Soc. Belg. 1889), although it is
evidently closely allied, but Allard makes no mention of
the flavous labrum nor similarly-coloured thoracic margins,
nor does lie mention the fine elytral punctuation,
Palxophylia semirugosa, sp. n.
Metallic green, the labrum, antenne and the legs flavous, thorax
finely granulate, the sides broadly flavous, elytra finely punctured
and transversely wrinkled.
bro
or
African Phytophagous Coleoptera.
Length 6 millim.
Of exactly similar coloration to A. granulosa but larger, the
frontal elevations transverse, the antenn extending to the apex of
the elytra, the second and following joints gradually elongate ;
thorax one-half broader than long, the sides strongly rounded, the
dise minutely granulate, with some extremely fine punctures, bright
green, the sides broadly flavous, this colour narrowed posteriorly ;
near the middle of the anterior margin a shght depression is visible ;
elytra closely and finely punctured, the interstices finely transversely
rugose, legs flavous, the first joint of the anterior tarsi of the
male dilated, the last segment of the abdomen deeply triangularly
emarginate.
Hab. Upper Tongaat, Navan (C. Barker).
Of this species I received a single specimen from Mr.
Barker ; the broad flavous margin of the thorax and the
sculpture of the elytra will at once distinguish it from
A. granulosa, which is also of general smaller size.
Malacosoma, Chev
Although this name has been in use for a genus of
Galerucine ever since 1846, it is high time to change it,
since it has long before that time been employed by
Hiibner for a genus of Lepidoptera. I am _ therefore
reluctantly obliged to change it to Hxosoma.
Exosoma barhkert, sp. 0.
Under-side black or piceous, upper-side flavous, thorax trans-
versely subquadrate, impunctate, elytra microscopically punctured.
Length 4-5 millim.
Head impunetate, frontal elevations transverse, nearly contiguous,
carina acutely raised, labrum and palpi flavous, antennee in the male
long and slender, extending beyond the middle of the elytra, flavous,
the terminal joints slightly darkened, third joint twice as long as the
second one, fourth joint distinctly longer than the third ; thorax
about one-half broader than long, of equal width, the sides rounded
at the middle, the anterior angles thickened but not produced, the
dise rather convex, not perceptibly punctured, scutellum triangular,
flavous, elytra slightly wider at the base than the thorax, closely
and extremely finely punctured when seen under a very strong lens ;
breast and abdomen nearly black, legs flavous, the first joint of the
anterior tarsi in the male dilated, that of the posterior legs nearly as
long as the following joints together, ‘all the tibie with a small
spine.
26 Mr. M. Jacoby on
Hab. Malvern, NATAL (C. Barker).
At once to be distinguished from other nearly similarly-
coloured species by the black under-side in connection
with the flavous head; the female insect is larger, the
antenn have the basal joints flavous only, and the tarsi
are more or less infuscate; the male organ has a very
elongate and pointed apex with the lateral margins acute
and subangulate, and a slender filiform process is placed at
the anterior cavity.
Monocida thoracica, sp. n.
Upper-side black, the clypeus, antennz, femora and apex of the
tibiz testaceous ; thorax minutely punctured, with a small fovea
at each side; elytra extremely closely and finely punctured, the
extreme suture piceous.
Var. Head, antenne, the elytra and the tibiz and tarsi, black.
Length 4 millim.
Head impunctate, the vertex black, frontal tubercles transverse,
strongly raised, bounded behind by a deep, transverse groove,
clypeus flavous, labrum piceous, antenn slender and filiform,
testaceous, the third joint one-half longer than the second one, the
following joints more elongate; thorax subquadrate, one-half
broader than long, the sides feebly rounded, the angles thickened,
the surface with a shallow fovea at each side, very minutely
granulate and punctured, scutellum black; elytra with a very
shallow depression below the base, very finely but more distinctly
punctured than the thorax, their epipleure broad, continued below
the middle, under-side black, legs pale fulvous, the base of the
tibiz black.
Hab. Lower Tugela, Nata (C. Bavier).
Closely allied to I. inornata, Jac., but differmg in the
following points:—the head is black at the vertex, not
fulvous, the frontal tubercles are not subquadrate but
strongly transverse, the thorax has a lateral fovea and the
elytra a shallow basal depression of which there is no
trace in I. inornata.
Sardoides transvalensis, sp. n.
Metallic green, the lower part of the face, the antenne and legs
fulvous, thorax subquadrate, rugosely punctured, elytra elongate, of
similar sculpture, abdomen piceous.
Length 6 millim.
African Phytophagous Coleoptera. a |
Head coarsely punctured and slightly wrinkled, frontal elevations
strongly raised, subquadrate, divided by a deep central groove,
clypeus and labrum flavous, antennze fulvous, the terminal three
joints black, fourth joint longer than the third and the longest ;
thorax about one-half broader than long, slightly narrowed
anteriorly and posteriorly, the angles tuberculate, the surface rugose
and strongly punctured, with three obsolete small fovez, scutellum
black ; elytra much wider at the base than the thorax, very closely
punctured and finely transversely rugose throughout, their epipleurz
broad and concave anteriorly, disappearing below the middle, breast
metallic green, abdomen piceous, legs fulvous, the tibie unarmed,
the first joint of the posterior tarsi as long as the following joints
together, claws appendiculate, anterior coxal cavities open.
Hab, TRANSVAAL.
There are some slight structural differences to be
noticed in this species in comparing it with the type,
S. viridicollis, Jac.; in the latter the thorax is deeply
bifoveolate and narrower at the base, and the elytral
epipleurez, although very narrow below the base, are
visible; in the present species they are absent, but as all
these differences are but one of degree, and the unarmed
tibiz and open cavities agree with the generical characters,
it would not be wise to separate the insect, of which I
possess three specimens. In the male, which may be
known by the dilated first joint of the anterior tarsi, the
last abdominal segment is rather deeply foveolate and
slightly emarginate at the apex with thickened sides.
Megalognatha, Baly.
The type of this genus is JM. elegans, Baly, which has
been described from apparently female specimens, now in
the British Museum. In the male insect the terminal
joints are considerably thickened, much more so than in
the other sex. Apophylia elegantula, Jac. (the Entomo-
logist, 1591), is identical with this species, and must be
omitted altogether. Baly’s description of his Jf elegans
differs somewhat from my specimens, as the lower part of
the face is not “nigro-piceous” but fulvcus, nor does Baly
mention the narrow fulvous lateral elytral margins.
Megalognatha bohemani, Baly (Cneorane foveicollis, Jac.).
This is another species which scarcely fits into the
genus, as the thorax is transverse and not subquadrate,
28 Mr. M. Jacoby on
although there is a fovea as in most species of the genus.
I have described this in the Entomologist, 1891, under the
above name, as Baly described his species from meee
specimens, in which the antenne are but little dilated ;
the male they are considerably so, and the terminal canis
are fulvous. As Baly’s name is the oldest, mine cannot
stand, although Cneorane is as good a genus for the species
as that of Baly’s.
Megalognatha hirticollis, sp. n.
Black, the abdomen flavous; thorax finely rugose and pubescent,
foveolate anteriorly, elytra closely punctured and finely wrinkled.
Mas. Antenne very long, the seventh joint dilated anteriorly,
hatchet shaped, thorax with a small triangular fovea.
Fem. Antenne shorter and simple, thorax of nearly similar
structure.
Length 8 millim.
Head impunctate, the frontal elevations very strongly raised,
narrowly oblong, antenne nearly extending to the apex of the
elytra, black, the third joint rather longer and more slender than
the following three joints, the seventh more strongly dilated at the
apex than the rest in the male; thorax as long as broad, all the
margins nearly straight, the anterior margin thickened at the middle
and followed by a small triangular fovea, rest of the surface finely
Tugose and pubescent ; elytra sculptured like the thorax but with-
out the pubescence, the sutural margin accompanied by another
ridge near the apex, under-side and the legs black, abdomen flavous,
the last segment more or less black.
Hab. Salisbury, MASHONALAND, on Zizyphus (G4.
Marshall).
From JL, rufiventris, Baly, Ml. abyssinica, Jac., and J,
imbecilla, Weise, this species may at once be distinguished
by the pubescent thorax and its sculpture.
Megalognatha granulicollis, sp. 1.
Head, antennz, the breast and legs black, thorax rufous, minutely
punctured and granulate, elytra flavous, convex posteriorly, very
finely punctured with a small piceous spot below the middle,
abdomen flavous,
Mas. Thorax with a shallow triangular depression, minutely
punctured and granulate.
Fem. Thorax coarsely rugose with a.very deep triangular excava-
tion.
African Phytophagous Coleoptera. 29
Length 6 millim.
Mas. Head black, impunctate, labrum flavous, antenne black,
the sixth to the ninth joint dilated, the eighth with a flattened
appendage below, the preceding two joints hollowed at the sides ;
thorax not broader than long, of usual shape with an acute but
shallow triangular depression before the middle and a shallow
transverse sulcus in front of the depression, the surface very
minutely punctured and granulate, scutellum piceous, elytra strongly
convex at the posterior portion, constricted at the sides, extremely
finely and closely punctured ; legs and the breast black.
V. Reenen’s Pass, Malvern, NataL (C. Barker).
In spite of the great resemblance of this species to J.
suturalis, Baly, and M. bipunctata, Jac., 1 cannot identify it
with either of them, as the thorax is quite differently
sculptured and punctured in both sexes, and the head is
black. In the first-named species the thorax is strongly
punctured and shining and without the triangular groove.
M. bipunctata has a fulvous head and differently-punctured
thorax. The thorax in the female has the depression
very deep, bounded at the sides by strongly-raised ridges
of perfectly triangular shape, and there are also some raised
feeble smooth lines visible at each side ; the elytral spot in
the specimen before me is just indicated and the suture
in both sexes has the usual ridge at the posterior portion.
Megalognatha natalensis, sp. 0.
Head, antenne and the breast black, thorax pale fulvous, strongly
punctured, deeply transversely sulcate anteriorly ; elytra flavous,
finely and closely punctured, strongly depressed below the hase.
Length 6 millim.
Fem. Head black, impunctate, antennee with the apical joints
widened, the third and fourth joints more slender, equal ; thorax
subquadrate, narrowed at the base, the surface strongly and deeply
punctured, deeply transversely sulcate anteriorly and to a less
extent near the base, the anterior edge of the anterior sulcus strongly
raised, scutellum piceous and pubescent at the base ; elytra rather
deeply depressed below the base, very finely and closely punctured,
legs obscure testaceous, stained with fuscous and pubescent, abdomen
flavous.
Hab. Frere, NATAL.
Of this species I only know a single female specimen,
like M. melanocephala the head is black, but the thorax
has no tubercles, is less strongly punctured, and altogether
30 Mr. M. Jacoby on
of different sculpture; as in the allied species, the elytral
suture has a narrow ridge at the posterior portion.
Megalognatha niyrofasciata, sp. n.
Head and thorax rufous, antennze, the breast and the legs black,
thorax closely punctured, triangularly depressed anteriorly, elytra
testaceous, finely and closely punctured, the suture narrowly and a
broader lateral band black, abdomen flavous.
Length 5 millim.
Mas. Head impunetate, rufous, the frontal elevations highly
raised, trigonate, antennie black, the seventh joint widened at the
apex, the eighth shorter, triangularly produced into a sharp point at
the middle, thorax as long as broad, the sides straight at the base,
-the disc rather closely and distinctly punctured, with a triangular
rather deep depression below the anterior margin, the latter
thickened, but not acutely margined, scutellum piceous ;— elytra
searcely depressed below the base, finely and very closely punctured,
testaceous, the suture narrowly piceous, the sides with a broad
black band which does not quite extend to the apex ; legs and breast
black.
TTab. CAPE (my collection).
M. bicostata, Alld., is evidently a closely-allied species,
but is described with fulvous legs and antennz and with
lateral elytral costs, not with a broad black lateral band.
The female is unknown to me.
Megalognatha melanocephala, sp. n.
Testaceous, the head, antenne, the breast and the legs black ;
thorax subquadrate, elytra finely and closely punctured,
Mas. Thorax impunetate, the anterior margin thickened, the
dise obsoletely depressed at the sides and the middle.
Fem. Thorax rufous, strongly punctured, deeply transversely
suleate anteriorly, the suleus bounded in front by a strongly-raised
ridge, the dise with three round tubercles.
Length 7 millim.
¢. Head longer than broad, impunctate, black, shining, frontal
tubercles strongly raised, carina acute, antennie black, the third and
fourth joints equal, the following three joints shorter and wider, the
rest more slender and elongate ; thorax scarcely broader than long,
the sides slightly narrowed at the base, the surface impunctate,
testaceous, the anterior portion slightly transversely thickened and
followed by a lateral and median very feeble depression, seutellum
pubescent, piceous; elytra distinctly transversely depressed below
African Phytophagous Coleoptera, 31
the base, very closely and somewhat rugosely punctured ; the breast
and legs black, closely pubescent, the abdomen testaceous, the
anterior margin of the last segment rather deeply concave.
2. Antenne simple, the terminal joints slightly widened ; thorax
rufous, strongly punctured, the dise with three blunt tubercles and
preceded by a deep transverse sulcus which is anteriorly limited by a
strongly-raised ridge.
Hab. TRANSVAAL.
The black head, testaceous colour, the transverse
depression of the elytra and their fine punctuation will
distinguish this species from Jf. rujficollis, Alld., and the
other species.
Pseudolognatha, gen. n.
Body elongate and parallel, head not longer than broad, frontal
elevations transverse, antenne filiform, the second joint small, the
others nearly equal, of normal structure, thorax transverse, sub-
quadrate, the surface more or less foveolate, elytra irregularly
punctured and granulate, tibiz unarmed, metatarsus of the posterior
legs as long as the following joints together ; claws appendiculate,
the anterior coxal cavities open.
Type. LP. immaculata, Jac. (sub Megalognatha).
In comparing this insect with a species of Megalognatha
and the type JZ elegans, Baly, to which it is generically
closely allied, there will be found differences sufficient to
justify the separation of the two genera. In the present
genus, the head is broad, not produced or longer than
broad as in Megalognatha, the antenne in the male have
no abnormal joints, and the thorax is distinctly transverse ;
there is also no thickening of the suture near the apex of
the elytra as is nearly always the case in the allied genus,
nor are the elytra widened and convex at the posterior
portion, but the short and deflexed head will form the
principal character of separation between the two genera.
Pseudolognatha salisburiensis, sp. u.
Fulvous, the apical three joits of the antennc fuscous, thorax
transversely subquadrate, impunctate, elytra metallic blue, extremely
minutely granulate and microscopically punctured,
Length 4 millim.
Head impunctate, fulvous, frontal elevations transverse, narrow,
labrum black, antennz about half the length of the body, fulvous,
32 Mr. M. Jacoby on
the terminal three joints blackish, the intermediate joints shghtly
triangularly widened, of equal length, the second one short, thorax”
one-half broader than long or rather broader, the sides feebly
rounded, the anterior angles more distinct than the posterior ones,
the surface with a very short and obsolete depression anteriorly and
a slightly more distinct one at each side, entirely impunctate,
fulvous ; scutellum of the latter colour ; elytra with a distinct
depression below the base, metallic blue, minutely punetured and
extremely finely granulate, under-side and the legs fulvous.
Hah, Salisbury, MASHONALAND (@. Marshall).
The above description applies to what seems to be the
male insect; in the female, the thorax has a deep, trans-
verse fovea near the anterior margin and the antenne are
somewhat thinner, otherwise there is no difference.
Hurycycla, gen. n.
Body elongate, glabrous, antenne filiform ; thorax transverse,
broader than the head and as broad as the elytra, narrowed anteriorly,
elytra metallic, irregularly punctured, epipleure prolonged, pos-
teriorly, tibizw unarmed, the metatarsus of the posterior legs as long
as the following joints together, claws appendiculate, anterior coxal
cavities open,
Type. L. balyi, Jac. (sub Megalognatha).
It is impossible to leave this species in Megalognatha, as
the thorax is of totally different shape without any
depressions, strongly transverse and narrowed anteriorly ;
in other respects the generic characters agree; I know of
no other African genus having a similar-shaped thorax.
Hemixantha flavicornis, sp. 0.
Black below, upper-side testaceous, the antennze very long, flavous,
thorax subquadrately transverse, impunctate, elytra subcylindrical,
extremely minutely punctured, tibiz unarmed.
Length 45 millim.
Elongate and parallel, the head impunctate, the frontal elevations
strongly raised, transverse, clypeus triangularly convex, antennie
nearly as long as the body, flavous, the apical joint stained with fuscous,
the third twice as long as the second, but shorter than the fourth
joint ; thorax one-half broader than long, the sides rather strongly
rounded at the middle, the anterior angles obliquely thickened and
slightly produced, posterior margin sinuate, oblique at the angles,
the dise with some minute punctures, only visible under a strong
African Phytophagous Coleoptera. 33
lens, seutellum smooth ; elytra subeylindrical and parallel, extremely
closely and finely punctured, under-side black, legs testaceous, the
metatarsus longer than the following joints together ; anterior
cavities closed.
Hab. Malvern, Nata (C. Barker).
A comparatively small species, allied to H. piceipes, Jac.,
likewise from Natal, but smaller, with entirely flavous and
much more slender and longer antenni, the legs differently
coloured and the metatarsus longer. I received two
specimens from Mr. Barker.
Platyxantha varicornis, sp. 0.
Pale testaceous, the head, thorax and the femora darker, the basal
and the apical two or three joints of the antennie black, thorax bifo-
veolate, impunctate ; elytra extremely finely punctured, tibiwe and
tarsi black.
Length 5 millim.
Head as broad as long, the vertex piccous or pale fulvous, unpunetate,
frontal elevations broadly transverse, divided by a short groove which
extends a little way upwards, clypeus acutely carinate, palpi
thickened, antennw nearly extending to the apex of the elytra, the
basal joint or two joints black, the following six fulvous, the apical
joints black, second joint very small, the rest nearly equal in length ;
thorax subquadrate, scarcely one-half broader than long, slightly con-
stricted at the base, the angles slightly oblique; the disc bifoveolate,
impunctate, pale fulvous ; elytra pale testaceous, extremely finely
but not very closely punctured, femora, the base of the tibice and the
abdomen pale fulvous, the breast, the lower part of the tibiae and the
tarsi black.
Hab. Malvern, NATAL (C. Barker).
In one specimen ( 2 ) of this species the extreme vertex
of the head and the greater part of the breast is blackish,
in the other these parts are fulvous ; the coloration of the
antenne and that of the legs are the principal characters
of distinction in this species; the tibiz are, as usual,
unarmed and the metatarsus of the posterior legs is as
long as the following joints together.
Monolepta punctipeninis, sp. n.
Under-side black, the basal joints of the antennze, the head, thorax
and legs flavous, head and thorax distinctly punctured, elytra dark
blue, strongly and closely punctured.
Var. Elytra testaceous. ;
TRANS, ENT. SOC. LOND. 1903.—PART I. (APRIL) 3
34 Mr. M. Jacoby on
Length 3 millim.
Head obscure fulvous, distinctly but not closely punetured,
minutely granulose, frontal elevations broad and flattened, carina not
very acute, labrum flavous, palpi piceous, antenné long and slender,
extending to the middle of the elytra, black, the lower four joints
flavous, basal joint long and slender, the second, third and fourth,
gradually lengthened ; thorax twice as broad as long, the sides
straight, narrowed anteriorly, anterior angles slightly oblique, the
surface very closely and comparatively strongly punctured, scutellum
small, black ; elytra not wider at the base than the thorax, much
more strongly punctured than the latter, the punctures arranged in
very close irregular rows, epipleur indistinct below the middle,
breast and abdomen black, legs flavous, the metatarsus very
elongate.
Hab. Malvern, NATAL (C. Barker).
This Monolepta may be known by the strong and close
punctuation of the elytra, which differs in that respect
trom any other species of the genus or those placed in
Candezea which I am acquainted with. I received two
blue and two flavous specimens in regard to the elytra,
but these agree in all other respects except colour.
Monelepta cerulea, sp. n.
Under-side black, upper-side metallic dark blue, the basal joints of
the antennie and the legs fulvous ; thorax subquadrate, impunctate,
elytra very closely punctured, the interstices likewise very finely
punctate and wrinkled.
Length 4 millim.
Oblong-ovate, the head impunctate, frontal tubercles transverse,
narrow, elypeus broad and thickened, blackish, the space below the
eyes at the sides fulvous, antenne extending to about the middle of
the elytra, the lower three or four joints fulvous, the rest black,
third joint one-half longer than the second but shorter than the
fourth joint; thorax about one-half broader than long, the sides
feebly rounded, posterior margin nearly straight, the anterior angles
thickened, the dise convex, impunctate, metallic dark blue, seutellum
black ; elytra strongly convex, their epipleura absent below the
middle, the punctuation very fine and close, slightly arranged in
rows, the interstices everywhere very minutely punctured and
wrinkled ; below black, legs fulvous, the first joint of the posterior
tarsi as nan as the follows joints together.
Hab. Dunbrody, Carr CoLony (Rev. O'Neil).
The metatarsus of the posterior legs in this species is
African Phytophagous Coleoptera. 35
shorter than is generally the case in this genus, with which
it agrees in all other characters; the unifor mly blue upper
surface and the sculpture of the elytra will comparatively
easily distinguish this species, of which two specimens were
forwarded by the Rev. O'Neil.
Candezxea (Iphidea) capensis, Baly.
In a publication by the late Miss Ormerod (Observat.
on African injur. Insects, 1889, p. 34), Baly has described
the above insect and placed it in Jphidea, a genus he him-
self had already withdrawn and considered identical with
Luperodes in 1873 (Trans. Ent. Soc., 1, p. 187). Miss
Ormerod some time ago kindly presented me with the
type, and on examination I find the anterior coxal cavities
closed and all the other characters identical with Candezea.
The species agrees in coloration with several others from
Africa ; it is testaceous above, black below, the thorax is
narrowed in front, transverse and impunctate, the scutel-
lum is black, and the elytra are finely and closely punc-
tured with the extreme sutural and lateral margins piceous.
I have also received this insect from Grahamstown where
the type was obtained, as well as from Malvern, Natal.
C. nigrosuturalis, Jac., is another closely-allied species, but
is amuch more narrowly-shaped insect, the antenne are
longer, and the elytra are distinctly margined with black.
Candezea braunsi, sp. 0.
Under-side black, upper-side testaceous, antennze fuscous, legs
fulvous, thorax transverse with three small fovez, elytra extremely
closely and finely punctured, their epipleurz continued to the apex.
Length 7 millim.
Very convex and elongate, the vertex fulvous, impunctate, frontal
elevations trigonate, small, carina acute, lower portion of face testace-
ous, antennz long and slender, the lower three jointsand the base of
the fourth and fifth, flavous, the rest fuscous, third joint twice as long
as the second, fourth joint longer than the preceding one ; thorax
twice as broad as long, the sides and the posterior margin rounded,
the angles distinct, the disc convex, finely granulate, with three very
small foveze, placed triangularly, scutellum rather small, testaceous ;
elytra wider at the base than the thorax, extremely closely impressed
with small piceous punctures, under-side black, closely covered with
grey pubescence, legs robust, fulvous, the coxee and the extreme base
36 Mr. M. Jacoby on
of each joint black, the metatarsus of the posterior legs much longer
than the following joints together.
Hab. AuGoa Bay (Dr. Brauwns).
Of nearly similar shape and coloration to C. hamatura,
Fairm., but the sculpture of the head quite different, the
labrum testaceous, not black, the antenne differently
coloured as well as the scutellum, and the elytra more
distinctly punctured ; from C. j/laveola, Gerst., the species
may be at once distinguished by the black under-side.
Candezea scutellata, sp. 1.
Under-side black, the basal joints of the antennie and the head and
thorax pale fulvous, impunctate, seutellum black, elytra testaceous,
extremely minutely punctured, tarsi and the posterior femora more
or less black.
Length 7 millim.
Again closely allied to CL Braunsi and C. hematiura, but with the
frontal elevations scarcely raised and the elypeus very broad and
flat, without carina, the antennie rather shorter, black, the lower three
or four joints flavous ; the thorax twice as broad as long, slightly
narrowed anteriorly, the sides nearly straight, the surface entirely
impunctate, without depressions, scutellum black ; elytra somewhat
narrowed posteriorly, their punctuation scarcely perceptible, of paler
coloration than the thorax, legs fulvous, the posterior femora and the
tarsi black, as well as the under-side ; metatarsus very elongate.
Hab, AuGoA Bay, Care (Dr. Brauns).
Of the two specimens, kindly sent by Dr. Brauns, one
has the tarsi almost concolorous with the legs; the
sculpture of the head and the black scutellum will
distinguish this species from its allies.
Candezea infuscata, sp. n.
Bluish-black, the basal joints of the antenni and the thorax flavous,
the latter very finely and obsoletely punctured, elytra bluish-black,
very closely and distinctly punctured in irregular rows, legs flavous,
the femora and the tarsi more or less fuscous.
Length 3 millim.
Head blackish, impunctate, the frontal elevations strongly raised,
transverse, clypeus highly convex between the antennze, the latter of
about half the length of the body, black, the lower three or four joints
flavous, the third joint about one-half longer than the second ; thorax
about twice as broad as long, the sides very feebly rounded, the
anterior angles slightly obliquely thickened, the surface very
or
African Piytophagous Coleoptera. 37
minutely, closely and obsoletely punctured, flavous; elytra very
closely and much more strongly punctured than the thorax, the inter-
stices very slightly wrinkled, bluish-black ; under-side of the latter
colour, the legs flavous, the femora and the tarsi obscure fuscous to a
greater or smaller degree ; the metatarsus of the posterior legs as long
as the following three joints together.
Hah. Malvern, NATAL.
Closely allied to C. nigroerrulea, Jac., and nearly
similarly coloured, but the thorax more obliquely narrowed
and the sides quite straight in the last-named species, the
elytral punctures finer, and the legs entirely flavous.
Buphonella, gen, n.
Body narrowly elongate, pubescent above, antenne subfiliform,
eyes entire ; thorax transversely subquadrate, finely rugose like the
elytra, the latter with the lateral margins deflexed, their epipleuree
extremely narrow; anterior cox raised, closely approximate, legs
slender, the tibizs unarmed, the metatarsus of the posterior legs as
long as the three following joints together, claws bifid at the extreme
tip only, prosternum invisible between the coxze, the anterior cavities
closed.
At first sight, this genus here proposed bears a greater
resemblance to the Hwmolpidx, and more especially to the
group Leprotine than to that of the Galerucidx, as the
antennze are more widely apart than is the rule in this last-
named tribe ; these organs are, however, inserted in front
and a little below the eyes; there are the usual frontal
elevations, and most important of all, the anterior coxze are
almost united, and hide the prosternum ;. they resemble in
that respect the genus Buphonida, Baly, which has likewise
closed coxal cavities and bifid claws, but the thorax in that
genus is transversely sulcate and of different shape; the
claws in the present genus are also rather peculiar, as cach
claw has the division united and bifid to a small extent at
the tip only. MJombasica, Fairm., seems also more nearly
allied to Buphonella than to any other genus of Calerucide.
Buphonella elongata, sp. n.
Narrowly elongate and parallel, black, finely pubescent above, the
basal joint of the antenne and the femora fulvous, thorax subquadrate,
very finely rugose, elytra still more finely sculptured, clothed with
short grey pubescence ; tibize unarmed.
Length 5 millim,
38 Mr. M. Jacoby on African Phiytophagous Coleoptera.
Head broad, closely rugose, the middle. with a narrow, elongate,
smooth raised space, eyes rounded, entire, frontal tubercles trigonates
smooth and tubereulate, clypeus transverse, antenne about two-thirds
the length of the body, black, the basal joint fulvous, the third joint
more than twice as long as the second, slightly shorter than the fourth,
the following joints slightly thickened, terminal ones thinner ; thorax
transversely subquadrate, one-half broader than long, the base shghtly
constricted, the anterior angles’ obtuse, thickened, the surfaee seulp-
tured like the head, the rugosities partly confluent, scutellum broader
than long, finely rugose ; elytra opaque like the rest of the upper
surface, extremely finely rugose and pubescent, body below and the
tibise and tarsi black, the femora fulvous, the last abdominal segment
triangularly emarginate (9 ).
Hab. Salisbury, MASHONALAND (G. Marshall),
Two exactly similar specimens were obtained by Mr.
Marshall.
Lrgana fulvipes, sp. n.
Under-side black, upper-side dark metallic blue, basal joints of the
antenne and the legs fulvous, thorax transverse, very minutely
punctured, elytra closely and strongly punctured.
Length 43 millim.
Head broad and short, impunctate, deeply transversely grooved,
frontal elevations strongly convex, trigonate, clypeus very broad and
swollen, labrum and palpi blackish, the penultimate joint of the latter
dilated, antenne extending to half the length of the body, black, the
lower four joints flavous, third jomt longer than the second, the
intermediate ones slightly widened; thorax more than one-half
broader than long, the sides strongly rounded, the posterior margin
to a less extent, the surface convex, very finely and closely punctured,
scutellum black, impunctate, elytra slightly wider at the base ‘than
the thorax, posteriorly widened, extremely closely and more strongly
punctured than the thorax, the interstices finely wrinkled and
furnished with still smaller punctures ; legs fulvous, all the tibie
mucronate, the first joint of the posterior tarsi as long as the following
joints together, claws appendiculate, anterior coxal cavities closed,
Hab. Dunbrody, CAPE Cotony (fev. O'Neil).
A typical species, and distinguished by its system of
coloration.
II. On the Life History of Drilus flavescens, Rossi. By
LIoNEL R. CrawsHay, M.A. Oxon. Communicated
by CHARLES OWEN WATERHOUSE, F.E.S.
[Read November 5th, 1902. ]
PuaTes I anp II.
THE following notes on Drilus flavescens are collected
from observations made during the past three years. In
July 1900 I first found the larva on the Downs, near
Seaford, Sussex, and in this year reared five larvae (all
females), four of them emerging in the following spring,
and the fifth continuing its growth for another summer.
In 1901 I collected several more larvee from the same
locality, and from these I obtained in the spring of the
present year (1902) a single male, and afew females, the
remainder reappearing as larvee.
In spots where snails—/felicella itala and Helicella
virgata especially—cover the ground in immense numbers,
it is not surprising that the larva thrives on its food-
supply, and it may often be seen during the summer
months, running hastily over the ground in search of food.
Before passing to its life history, the form of the larva
deserves some notice.
The larva, which is narrowed in front and much widened
behind, has the upper surface of the abdomen rather
thickly covered with coarse hairs of a bright burnt-sienna
colour, springing chiefly from four longitudinal rows of
fleshy processes, the processes increasing in length towards
the posterior. The head is reddish-brown, flat above,
with strong sharp mandibles, curving upwards and crossing
one another above the labrum; the eyes consist of a single
ocellus on either side. The antenne (Plate I, fig. 1;
Plate II, fig. 1) are two-jointed with a supplement to the
second joint, and can be partially extended or withdrawn
by the larva by means of a collapsible membranous tube
which carries the first joint, the latter being thrust forward
or partially withdrawn within the tube by a muscle which
passes up the centre of it to the apex of the first joint.
The second joint is somewhat flattened and bears at its
TRANS. ENT. SOC. LOND. 1903 —PART I. (APRIL)
40 Mr. L. R. Crawshay on
apex—on the inner margin a two-jointed supplementary
process terminated by a long seta, and on the outer margin
a smaller colourless process. The thoracic segments have
a reddish-brown corneous covering above—marked with
dark patches—with a few hairs. On each of the abdominal
segments to the penultimate one the soft white body of
the larva is protected above by a dark brown dorsal shield,
sparsely hairy, bearing a bristly process on each margin,
and by a pair of similar processes outside these, arising
from the sides. The last segment has a single pair of
larger bristly processes extending behind, with a spine at
the apex of each, completing a covering which doubtless
goes far to protect the larva from attack during its occu-
pation of the snail-shell. The spiracles (Plate I, fig. 2)
lie in an uncovered space between the dorsal shields and
lateral processes, and project from small horny encase-
ments which lie along the surface of the segments. A
fleshy nipple on the under-side of the last segment aids
the progression of the larva, and enables it to cling firmly
to the surface of the snail-shell. It moves rapidly about
the outside of the shell, aided also by a pair of small
colourless pulvilli, attached by slender stalks to the base
of the tarsal claw (Plate I, fig. 3). ;
The young larva, which is hatched about the middle
of July, feeds till September, and then hibernates in the
shell it has last occupied, attaining only a small size in
the first year. This shell is evacuated in the following
spring, and after this the larva may continue feeding at
regular intervals till September. But owing to the exist-
ence of a distinct, inactive form for hibernation, and the
fact that this may be assumed at any time of the summer,
it is equally liable to pass nearly the whole of the year in
a dormant state in the shell. Doubtless the larva often
becomes full-fed in the year after hatching, but it probably
more often feeds fora third summer. The full-fed larva
then changes into a second hibernating form, pupating
in the following spring, shortly before the appearance of
the imago in May or June of the third or fourth year,
as the case may be.
The snails which chiefly constitute its food in the
locality referred to are Helicella itala, Linné; Helicella
virgata, Da Costa; and Helicella caperata, Montagu; but
the larva has shown the same readiness to attack any
other species that I have hitherto tried, including :—
The Life History of Drilus flavescens. 4]
Helicella cantiana, Montagu ; Hygremia rufescens, Pennant ;
Vitrea cellaria, Miiller; Helix nemcralis, Linné; and even
the largest examples of Helix aspersa, Miiller; passing
indifferently from one to another in the course of its
growth.
When a snail is found, the larva raises itself at once
on to the shell and examines it carefully, assuring itself
of the presence of its occupant. If unsuited to the size
of the larva, the snail is left and the search resumed. If
it is satisfactory, the larva (more especially in its earlier
stages at least) then proceeds to sound the surroundings,
reaching out to its full length and ranging about on all
sides above and below, but retaining all the while its hold
on the shell by means of the anal clasping-organ. This
is done with a view to secreting the snail before attacking
it. Ifthe position is found to afford sufficient cover, the
snail is soon pushed or dragged into it, till out of sight;
otherwise, if it is too much exposed (as on a roadway, for
example), the larva endeavours to remove it to a better
one. This removal of the snail to a place of hiding may
prove a difficult matter, and remarkable perseverance is
often shown in the endeavour to accomplish it. The larva
takes a firm anal grip on the outer surface of the shell,
and bending over the side, plants its head on the ground
beneath; then, taking hold of the ground with its man-
dibles, it proceeds to “punt” the snail backwards over
the ground, so far as possible in a straight line. Where
an obstruction occurs, in the endeavour to pass it, the
larva will often raise the snail entirely into the air and
throw it forward bodily, but never losing its hold upon the
shell. Ifin the meantime the snail appears and attempts
to crawl away it is attacked with the mandibles and
driven back. From time to time the larva returns to
the summit of the shell to try the surroundings as before,
afterwards going back to its task, which, if no cover should
be found, may often be continued for an hour, or, as I
have once noticed, for as much as three hours before the
snail is attacked. It is advisable, when feeding the larva,
to provide some loosely placed moss, into which it will
soon carry the snail out of sight. In the case of larger
larvee (about 14 mm. and over) the snail is generally
attacked very soon, without any attempt to remove it,
however exposed the position may be. Larger snails (and
I refer especially to H. cantiana) seem instinctively to
42 Mr. L. R. Crawshay on
know their danger when this horrible parasite has attached
itself to their shells, and will do their utmost to dislodge
it by making rapid contortions of their shells around them
as they crawl, even to the extent of overbalancing them-
selves, and while the larva perhaps lies motionless on the
shell. It is surprising that at such times the larva retains
its hold or escapes injury, but it quickly takes up a position
near the orifice and makes a ferocious onslaught with its
mandibles, which probably soon overpowers its wretched
victim ; I have, however, more than once seen a large
H. cantiana throw off its assailant and escape.
When the larva enters the shell it lies inside, against
the outer wall, and gradually absorbs the moisture, finally
consuming the whole or the greater part of the body of
the molluse. After a period varying from eight to sixteen
days the shell is thoroughly cleaned out, preparatory to
a succeeding period of inactivity and moulting. The
larva may then be heard scraping with its mandibles
within the shell, and may be seen making a number of
journeys backwards, from the centre of the spire to the
orifice, to eject from the shell the accumulated and wneon-
sumed matter. The hairy processes seem to aid the work,
as a brush, while the larva shuffles backwards down the
shell. As the terminal segment reaches the orifice, the
anal clasper grips hold of the edge, and, by an undulating
movement of the body, the slimy matter 1s ejected from
the shell. I once found a very large larva thus engaged
at 10.30 p.m. (having already been working for perhaps
half-an-hour). I then watched it for an hour and a half,
while it made nine journeys down to the orifice, mostly
at intervals of three to five minutes, and with generally
about twenty to twenty-five of these undulating move-
ments at the end of a journey. After this J] ceased
observing it, but I have found that with varying intervals
the work may extend over the greater part of a day. In
this way the interior of the shell is at length thoroughly
cleaned out and made fit for habitation. The larva, which
has become gorged with food almost beyond recognition
(the dark scuta appearing as isolated patches on the
distended white body), then thrusts itself up, with its
head towards the centre of the spire, and prepares for
moulting, which takes place about four or five weeks from
the time of entering the shell, the skin being then cast
towards the orifice, “where it sometimes closely stops up
The Life History of Drilus flavescens. 43
the entrance. While the summer lasts, the undeveloped
larva, as a rule, evacuates the shell about forty days after
entering it, and goes off in search of another. Excluding
the shorter feeding interval (July to September) in the
year of hatching, it may thus devour as many as three,
or perhaps four snails in the course of the summer,
moulting once in each case before leaving the shell. But
it often ceases feeding as early as July, and in one example,
at least, under notice this year, the first shell entered by
the larva on June 2nd, after hibernation, was not left after-
wards. In such cases the larva, on moulting, reassumes
the winter form (which is referred to below), and the
growth is again suspended until the following year, From
examples under observation, this early return to the winter
form seems to be caused by a period of low temperature
preceding the time of moulting. An instance of it occurs
in the following incomplete record of a remarkable larva:
1900, July 8th: Larva (length about 8 mm.) entered
HT, virgata (greatest diameter 8
mm.) ;
1901, May 30th: evacuated this shell (length about
10 mm.), and entered H. aspersa
(greatest diameter 8 mm.) ;
» duly 10th: left this shell (length about 13 mm.),
and entered //. aspersa (greatest
diameter 28 mm.) ;
, August 18th: left this shell (length 21 mm.), and
entered Hf. aspersa (greatest
diameter 25 mm.);
a », olst: seen to be cleaning out this shell
(length, at the time, fully 30 mm.).
Unfortunately, in the spring of 1902, it died while still
in the winter stage. The larva was doubtless hatched in
July. 1899, and must therefore have become full-fed in
the third year, as seems to be more often the case.
The winter form, into which the undeveloped larva
changes about the middle of September, or often earlier,
as stated, is incapable of feeding or of more than a heavy
grub-like motion, when disturbed. In general outline
it much resembles the ordinary form of the larva, but it
rather perhaps deserves the term “false pupa.” The scuta
are absent, the body being almost entirely soft, of a dingy
whitish colour, and, except on the last three or four
segments, almost hairless. The head is small, rather soft,
44 Mr. L. R. Crawshay on
and pale, with the mouth parts rudimentary, and the
antenne very short and much modified. The legs are
soft and short, with the claws absent and replaced by a
small prominence. The processes on the body are much
smaller and less distinct, with only a few fine hairs, until
the last three or four segments, where they become longer
and rather thickly hairy, but with the hairs shorter than
in the larva. The terminal processes are likewise shorter,
but with the spines long (Plate IJ, fig. 2a). This skin is
cast about the middle of May, and the larva then reappears
from the shell in its ordinary form, continuing its life as
before, until it is full-fed, in the second, or, probably in
most cases, the third summer.
When full-fed, it changes into a second inactive - winter
form, which more nearly approaches the pupa, and which,
like the other, may be assumed early or late in the year,
often as early, at least, as July. Though this is very
similar to the previous one, it differs from it especially
in the much more stumpy form of the antenne and of
the processes on the last three or four segments; the hairs
also on the latter are finer and shorter, and the spines on
the terminal pair are very short (Plate II, fig. 3a).
Points of difference in the head and last segment of
the two forms are figured in the accompanying drawings
beside the corresponding parts in the ordinary form of
the larva (Plate IT).
The two small apical processes in the antenne of the
larva are retained in both winter forms; but what is the
“outer” process in the larva becomes inferior and invisible
from above in the first form, and partly inferior while
visible on the dnne7 side in the second form.
These sluggish winter forms (or “ false-pupz,” if they
may be so termed) appear to possess a strange tenacity
of life. A specimen of the sezond form, lately kept out
for examination, passed a day and a half in a solution
one-third alcohol and two-thirds glycerine; then, two
days afterwards, an hour in turpentine, followed by a day
in alcohol and glycerine. Three days after removal from
the latter, it seemed so little the worse that, after washing
it in alcohol and in water, I put it into an empty shell,
which it entered without difficulty, and it appeared to
have its usual amount of vitality when examined a week
afterwards. Though in a much lesser degree, the same
point is noticeable in the 2 imago, which will live for two
The Life History of Drilus flavescens. 45
or three hours in the same preservative solution, or return
to its dull existence apparently unaffected by an immersion
of an hour and a half in whisky half diluted.
An offensive acrid fluid is emitted from both extremities
of the body by the two winter forms of the larva, when
disturbed ; the active form does not seem to possess this
resource, but rolls itself into a ring.
In Mulsant’s account of the insect (“ Histoire Naturelle
des Coléopteres de France; Mollipennes:” pp. 422 ff),
pupation is said to take place fifteen to eighteen days
before the imago appears. Prior to this, the position
within the shell is reversed, so that the head lies in the
direction of the orifice (in which respect this second winter
form differs from the earlier one). In a single instance
this change of position had already occurred when I ex-
amined the shell on October 8th, but ordinarily it appears
to be made in the spring. At the time of pupation, the
skin is cast far back, near the centre of the spire. The
skin of the pupa is soft and unprotected, and merely
displays rather distinctly the enclosed form of the imago.
When the final change takes place, about the middle
of May, the imago moves forward and occupies the inter-
vening space between the pupa and the larva-skin of the
previous year, lying with its head thrust against the
anterior part of the latter. There are thus, at this point,
four stages of the insect represented in the shell. Here
the imago remains for some days before it is able to leave
the shell. In three female examples observed in the
present year, at least eight days were passed in this way,
and the case of a male, referred to below, was similar.
The 1mago appears at the end of May or beginning of
June, about the same time as the hibernated larva.
The apterous female (Plate I, fig. 4} is elongate and
broader behind, with the abdominal segments sharply
explanate under the spiracles, so that the sides have a
scalloped appearance. It is of a tawny testaceous colour,
marked on each segment, except the last, with a pair of
large dark-brown digitate patches more or less regular in
shape, and thinly covered, especially above, with fine short
rufous hairs. The last segment is terminated by two
processes with a small palpiform appendage at the apex
of each. The supplement to the antenne found in the
larva and retained in both winter forms, reappears very
similarly in the female imago. At the apex of the tenth
46 Mr. L. R. Crawshay on
joint there isa small inner supplementary joint, terminated
by a short seta (which is sometimes scarcely apparent),
and a smaller process at the outer margin. The antennz
of the female are normally composed of ten joints (omitting
the supplement), but the ninth joint is often imperfectly
formed, being sometimes confounded with the preceding
one, so as to be scarcely visible, and sometimes entirely
absent. This deformity may even appear in different
degrees in the two antennze of the same insect.
Among the imagines that emerged this year I obtained
only a single male, and this one happened to occupy one
of two shells which I examined on May 18th. It seemed
inclined to leave the shell, so I removed it, but it proved
to be quite helpless and could only le on its side, in a
curved position, twisting the distended abdomen about
like a pupa. I put it on some moss in a shady place, and
its condition gradually advanced till, seven days afterwards,
it was quite strong. For breeding purposes I had there-
fore to rely on this male alone, and it paired successively
with four females (twice over with one of them) on May
25th, 31st, June 2nd, 4th, and 9th, respectively.
~ In the spring of the previous year a few eggs had been
laid by some infertile females, a week after emerging,
on the moss in which they were kept; but this year,
under similar conditions, the first female that paired
passed six days without laying: there was also reason for
believing that the eggs are naturally secreted. I therefore
cut a piece of fine turf from the Downs, ten inches by six
inches and two inches deep, which I fitted closely into
a shallow deal box, and on this the females were after-
wards kept near an open window. The following is an
account of the movements of these four females which
I reserved, and the attempt to breed from them.
The first ?, as stated, was not put on the turf till six
days after pairing, namely on May 31st. From May 31st
till June 4th she was up about the surface, generally
resting in an exposed position on the short herbage, during
each day, but going down every evening under the grass.
On June 5th she came up in the morning and rested in
the usual attitude on a short blade of grass, with the heavy
body curving under her, and thus remained, never leaving
the position, so far as I know, day or night, until June th,
when she fell off and died without having laid.
A second 2 emerged on May 26th and paired on the
The Life History of Drilus flavescens. 47
turf on June 2nd, at 11.80 am. She continued to move
restlessly about the surface till 3.30 p.m., when she dived
down a hole between the turf and the side of the box, and
did not appear again.
A third emerged on June 2nd and paired on the turf
on June 4th, disappearing on the same day.
A fourth emerged and paired on May 31st, but was not
put on the turf till five hours afterwards. From this date,
this 2, like the first, was about the surface (generally
resting on the herbage) regularly during the daytime, but
always going down under the grass in the evening, until
June 9th. I then tried the % again, and she paired a
second time, disappearing soon afterwards on the same
day.
‘After June 9th none of them appeared again.
Though it is to a great extent the instinct of the @ to
hide herself, she seems, in fact, after emerging, to choose
more often some exposed position on the ‘short herbage,
and to remain clinging to this for hours at a time without
moving. From the moment of pairing she proceeds at
once to make her way down under the grass (the probably
sharing the same instinct), and after separation—7. v. after
about forty mimutes—disappears very shortly, as 1t seems,
into the turf, where she lays her eggs and dies. The
failure of the first of these tour was perhaps due to the
absence of natural conditions in the first week.
On June 20th I examined the turf in search of ova.
The body of one 2 was found resting in a perpendicular
position between the turf and the side of the box. The
body seemed half empty, but I could find no eggs that
had been laid. Probably this was the second 2 mentioned
above.
The bodies of the other two I found in the bottom of
the turf (2. e. two inches deep), each lying with its contents
emptied in a compact heap within a distance of half-an-
inch from it. Doubtless both had died very soon after
laying.
The egg is nearly spherical, about 1 mm. in diameter,
and of a pale yellow colour. Roughly speaking, there
were about thirty to thirty-five eggs in each heap.
It was a mistake to disturb them so soon, but the period
of development proved to be much longer than I expected,
and to leave them as they were, and unobserved, was
unsatisfactory. In an endeavour to preserve natural
48 Mr. L. R. Crawshay on
conditions, the evils of excessive moisture and excessive
dryness crept in, and worked serious injury in the first
week or ten days. On July 2nd the embryo began to
show signs of development, and on July 8th the form of
the young larva was discernible. After this they advanced
rapidly till the first hatched on July 20th, or about six
weeks from the approximate date of laying. But mean-
time the greater number had fallen off in various stages
of development, and only five, out ‘of perhaps sixty-five,
survived, hatching respectively on July 20th, 22nd, 24th,
25th, and 26th. One of these was a cripple and another
was lost by an accident.
At the time of hatching, with the exception of the
conspicuous black ocelli and a few reddish hairs, they were
nearly white, the head and the shields about the segments
attaining their darker colour several hours afterwards.
The length is about 25 mm.
Of the three that remained, two began to feed about
thirty-six hours after hatching; the third clung to its egg-
shell for twenty-four hours, and refused food for three
and a half days. The snails at length given them were
Helicella caperata, and Hygromia rufescens, 3 to 4 mm. in
diameter.
In these three examples (as also the one that was lost)
I noticed most of the instincts of the older larva, including
the feigning of death for as long as fifteen or twenty
minutes at a time. There was no attempt to push the
snail away, but a decided tendency to work underneath
it, perbaps with the same purpose. More than once a
snail that began to appear from the shell was sharply
struck with the mandibles and driven back; in another
case the hardened film quickly thrown by the snail as a
barrier across the orifice was cut away by the larva after
an hour and a half’s work, and the shell was then entered.
The ultimate loss of these larvae was perhaps largely due
to the period of exceptionally cold weather that followed
the time of hatching, and particularly so in the Cotswolds,
where I happened to be staying. The fragmentary
account of them, so far as it goes, may, however, be worth
recording.
Larva (a) hatched on July 20th, and entered the first
shell on July 21st; partly devoured this snail; left it and
entered another on August 2nd, but afterwards became en-
tangled inside the shell, and died without having moulted.
The Life History of Drilus flavescens. 49
Larva ()) hatched on July 22nd, and on July 25th
entered the first shell, in which it became entangled, and
died without moulting.
Larva (e) hatched on July 25th, entered the first shell
on July 26th, and, after partly devouring the occupant,
left it and attacked a second on July 28th; was ejected
some hours afterwards by the latter, and left entangled ;
I released it and put it back on the first shell, which it
at once re-entered, but it afterwards died without having
moulted.
I have nevertheless found the exuvie of the young
larva at about 3 mm. in very small snail-shells (8-4 mm.
in diameter), and it seems more probable that the economy
of the larva in its infancy is the same as in its later growth,
and that under ordinary circumstances the first shell is
not left till after moulting, in the usual manner. It would
thus be able to attain a length of about 4 mm. at the
time of entering the second shell, at which point it is to
be found feeding, about the middle of September, prior
to hibernation. After this, the growth of the larva is
regulated by the size of the snails it happens to meet
with, its length being increased, as I have found in most
cases observed, roughly by about half the greatest diameter
of the snail attacked. The female imago may thus be
no more than 8 mm. in length, or, on the other hand,
probably not far short of 830 mm. On the Downs, larger
snails such as H. cantiana are seldom met with by the
larva, and the average length of nine naturally-reared
females which emerged this year was 10°7 mm. only.
In regard to the life history of the male, Mulsant
mentions the fact that out of one hundred and fifty shells
containing larvae, Desmarest obtained only two males,
while M. Rouget obtained only one male among about
two hundred females. Appended are a few details bearing
on this point.
As already mentioned, all of five larvae which I reared
in 1900 were females.
The shells collected in the autumn of 1901 and early
in 1902 varied from 4 to 12 mm. in their greatest
diameter (i.e. the measurement across the shell to the
outermost point of the orifice). Of those larvae that were
full-fed, nine were females and five were males. All of
the females emerged, but four of the males died as pupx
(the fifth was taken in February ; the others in September).
_TRANS, ENT. SOC, LOND. 1903.—PART I. (APRIL) 4
50 Mr. L. R. Crawshay on
Another shell contained the remains of a male pupa
which had probably failed to emerge in the previous
spring.
The shells containing these six males were :—
Two of them Helicella virgata, Da Costa; 8 and 9 mm.
in diameter respectively.
Four of them /elicella caperata, Montagu, 7, 7, 7, and
S mm. in diameter respectively.
Another shell found at the same time, containing a
male pupa skin, was that of Helicella caperata, 6 mm. in
diameter.
The following table shows the greatest diameter of
thirty-seven winter shells {all naturally occupied by larvee),
and the state of the insect after hibernation :—
2 shells of 4 mm. in diam. produced 2 undeveloped larve.
9)
= ” Dy ” ” 2 ” ”
a) ” 6 5, ” ” es ” ” and 1 or
6 ” 7 ” ” ; 3 ” ” » 36.
6 ” 8 ” ” 4 ” F) » 26.
5 ” 9s, ” ” 4 ” ” Lt
11 ” 10-12 ” 9 2 ” ” ” 9 ce
e
In other words, all full-fed larvee occupying shells of
6—9 mm., and averaging 7} mm., were males, and all those
occupying shells over 9 mm. were females; which result
might, in fact, be expected approximately, considering
that the male imago never perhaps attains the minimum
length of the female.
The females probably outnumber the males very consider-
ably, while the male larva must be sought for in shells of a
small size only. But, apart from these considerations, the
male seems, for some reason, to be more difficult to rear
than the female—possibly owing to the fact that it is more
likely to complete its limited growth early in the year,
and so to be subject to a longer “period of exposure to the
exigencies of weather.
The number of eggs laid by the female may be as many
as three hundred, or even more; but the female, which
is little more than an ege-magazine, varies in size to an
extraordinary degree, and the number of eggs must vary
greatly in proportion. In a female of 19 mm. I found
two hundred and sixty-seven, but in smaller examples
the numbers were far short of this. The two that laid
(presumably 10 mm. and 13 mm. respectively) cannot
either of them have laid more than about forty; the abdo-
The Life History of Drilus flavescens. 51
men of another (11 mm.), nine days after leaving the shell,
contained thirty-three; while in another (8 mm.), seven
days after leaving the shell, there were only four or five
developed ova, and the abdomen could hardly have
contained more than fifteen or twenty.
The following case of cannibalism occurred in the
present year :—
Two larve, “A” and “B,” were due to leave their shells
about the same date, early in July, and both had already
moulted. “A” left its shell first, and escaping unobserved
into the division of “B,’ entered the shell and attacked
and devoured the unfortunate occupant. The unconsumed
remains of “B” were afterwards cast back and left near
the orifice of the shell, and the larva “A” in due time
moulted, evacuating the shell thirty-five days after the
probable date of entering it.
Among a number of shells examined, which have con-
tained larve, I have not noticed any clear case of parasitic
attack on the species. I am indebted to Mr. C. O. Water-
house for his kind assistance in connection with the
subject of this paper.
EXPLANATION OF PLATE I.
Figure 1, Left antenna of larva, partially withdrawn.
2. Spiracle of larva, projecting from horny encasement.
3. Part of intermediate leg of larva.
» 4 Female imago.
EXPLANATION OF PLATE II.
Figure 1. Active form of larva; part of head, showing antennze
and mandibles (antennze extended),
2. Winter form of undeveloped larva; part of head, showing
antenn and mandibles.
3. Winter form of full-fed larva; part of head, showing
antennee and mandibles,
», la. Active form of larva ; last segment.
2a. Winter form of undeveloped larva ; last segment.
oe oa: ES » 9) full-fed larva ; last segment.
+P)
Ill. On the Genus Deilemera, Hiibner. By CoLONEL
CHARLES SWINHOE, M.A., F.L.S., F.Z.S.
[Read Feb. 4th, 1903.]
_Puates IIT anp IV.
THE genus Nyctemera was erected by Hiibner in the
Verzeichniss, p. 178, and diagnosed as “ Die Fliigel weiss
und weissfleckig schwarz gesaiumt.” Under this genus
Hiibner put seven species in the following order :—
hesperia, Cram., Pap. Exot., iui, pl. 251, f. A. B. (1780).
cajras Orin Wl Nx. Hat.ai1, pl..p, fF 780):
nerina, Drury, l.c., f. 2.
coleta, Cram., ].c., iv, pl. 368, f. D. H. (1782).
jamal, Drury, le. u, pl. 1 £3 A773):
atralba, Hiibner = tripunctaria, Linn., Syst. Nat., i, p.
523 (1758).
lacticinia, Cram., |. c., 11, pl. 128, f. E. (1779).
Walker in the 2nd vol. of his Catalogue, p. 402, erected
the genus Ofroeda, placing therein in the following order :—
hesperia, Cram., |. ¢.,
occidentis, Walker, 11, 403,
vesperina, Walker, 1, 403,
caffra, Drury, 1. c.,
nerina, Drury, |. c.,
leaving coleta, Cram., as the type of Hiibner’s genus Vycte-
mera; and this seems to have been followed ever since,
except that Hampson makes /acficinia, the seventh in the
list of Hubner’s named species, the type of the genus.
No author in erecting a new genus can be justified in
removing the first three species, including of course the
type, of a genus already in existence, yet this is what
Walker did.
It is not only that hesperia, being the first species in the
seven under the genus Nyctemera, is thus made the type
of the genus, but Hiibner’s diagnosis, “ wings white with
white-spotted black borders,” is a good description of
hesperia, and not of any one of Walker’s so-called Nyctemeras;
consequently as hesperiais the type of the genus Nyctemera
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL)
54 Colonel C. Swinhoe on
Walker’s genus Otvoeda becomes asynonym, and as hesperia
is a Lymantrid, another name must be found for the species
hitherto erroneously referred to WV; ye temera.
This is not far to seek ; Hiibner’s next genus Deilemera
contains a single species only, evergisia, Cram. .. and about
this type there can be no mistake.
For some of the other species in this genus five genera
have been erected by different authors. I have had one
or two examples of species in each genus denuded of their
wing scales, and after carefully working through the species
have come to the conclusion that structurally there is no
essential difference* between them, and that the genus
Deilemera, Hiibner, must include them all.
The scheme of neuration of the fore-wings is practically
the same throughout; the length of the areole, by which
Hampson separates Deilemera from the rest of the species,
appears to be as variable as its breadth, in fact both its size
and shape vary not only in individuals ‘of the same species,
but in the right and left wing of the same individual.
In the hind-wings veins 6 and 7 are generally stalked ;
in the few instances where 6 and 7 arise from the cell, this
difference is found to be correlated with a difference in the
structure of the antenne and palpi, and may be legitimately
employed for the purposes of sub-division ; thus the species
included under Pitasi/a, Moore, and Atasea, Swinhoe, which
have veins 6 and 7 arising from the cell, agree in also
having the palpi much shorter than the others, Atasca
being separated from Pitasi/a by the antenne in both
sexes being simply pubescent, with a short bristle from
each joint, whereas in Pitasila they are bipectinate in both
sexes, the pectinations being short.
Tn the large majority of cases, those with veins 6 and 7
of the hind-wings stalked, with longer palpi and more fully
pectinated antennie, secondary sexual characters are avail-
able for sub-division; thus a few species with evergista,
Cram., the type of the genus, have in the male the inner
margin of the fore-wings strongly convex, and the inner
margin of hind-wings amplified and containing a fold and
furrow, the outer margin produced and straight from the
anal angle, not rounded, so that the hind-wing becomes
oblong im shape.
Again in coleta, Craim., the fore tibia in the male bears a
long pencil of hairs, and the species is further separated by
* Except in coleta, where vein 11 is short stalked with 9 and 10.
IU
The Genus Deilemera. 55
a slight difference in neuration, vein 11 of fore-wings being
stalked with 9 and 10.
A third sub-division of Section IL is necessary for the
African species, including lewconoé, Hoptter; in this the
hind-legs of the male sutfer great modification, the coxe
are greatly elongated, the femora still more so, the tibiz
are all but obsolete and without spurs, while the tarsi are
much attenuated and almost equal to the femora in length,
and are accompanied by a tuft of hairs as long as them-
selves, from the femoro-tibial joint.
The fourth sub-division contains species without second-
ary sexual characteristics, and will include all the species
heretofore placed under TZvripheromera, Zonosoma, and
Leptosoma, amongst which there is no ditference whatever,
except in the pattern and sometimes as in cenis in the
length of the wings.
As might be expected with moths of weak flight, nearly
every island contains its own peculiar form; this of course
does not of a necessity make each a good species, as in
baulus, Boisd.= mundipicta, Walker, forms of which are
found in nearly all the islands (probably in all), from the
Philippines to the Solomon Group, and although one can
generally trace some small difference in each, the differences
are too slight for any attempt to make any distinctions.
The Family NYCTEMERID stood for many years by
itself, next to the CALLIDULIDA, where Kirby puts it
in his catalogue of Heterocera. Meyrick, in Proc. Linn. Soc.
N.S.W. 1886, p. 687, put the members of this group into
the Family HYPSIDA = AGANAIDA, and though
Hampson, in his Moths of India, vol. 11, put them into
the ARCTIIDA, they have now been put in the National
Collection with the Aganaidz, which is evidently their
proper place, and they form a sub-division of that Family.
KEY TO THE SUB-DIVISIONS OF Deilemera.
Section I, Veins 6 and 7 of hind-wings not stalked, palpi short.
A. Antenne not pectimated. -. . . . .. . (Atasca)
p. Antenne shortly pectinated. . . . . . (Pitasild)
Section II. Veins 6 and 7 of hind-wings stalked, palpi longer,
antenne with long pectinations.
A, Both wings of male abnormally developed. (Deilemera)
B. Male with long tuft of hair to fore tibia. (coleta group)
c. Male with abnormal hind-legs. . . . . (lewconoé group)
D. Without secondary sexual characters. (Tripheromera* )
* Leptosoma and Zoiosoma being praeoecupied.
56 Colonel C. Swinhoe on
Genus DEtLeMERA, Hiibner, Verz. Schmett, p. 178 (1815).
Leptosoma, Boisd., Voy de l Astrolabe Lep., p. 197 (1832)
(pracoce. ).
Pitasila, Moore, P.Z. 8., 1877, p. 599.
Zonosoma, Butler, Ill. Het. B. M., v, p. 44 (1881) (praeocc.).
Tripheromera, Butler, l.c., p. 45.
Tristania, Kirby, Cat. Het., i, p. 423 (1892).
Atasca, Swinhoe, Cat. Het. Mus. Oxon., i, p. 139 (1892).
SECTION 1.
Veins 6 and 7 of hind-wings not stalked, palpi short.
A, ANTENNA NOT PECTINATED. . . . . (Atascc)
DEILEMERA PELLEX.
Phalena (Noctua) pellex, Linn., Syst. Nat., x, p. 530 (1758).
Leptosoma pellex, Aurivillius, Ree. Crit. in Sv. Ak. Handl.,
xix (5), p. 161, pl. 1, £ 5 (1882).
Atasca pellex, Swinhoe, |. c.
Nyctemera artemis, Walker, 1, 394 (1854).
Nyctemera separata, Walker, xxxi, 204 (1864).
Sixteen examples from Batjan, Ceram, Aru, N. Guinea,
Fergusson Isl., and N. Ireland. Type separate from Gilolo
is in Mus. Oxon.
DEILEMERA ALBIPUNCTA. (PI. III, f. 8.)
Deilemera albipuncta, Druce, P. Z.S., 1888, p. 573.
x . Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 164.
Two examples from Guadaleanar Island (J/eck). A
very distinct species.
DEILEMERA SIGNATA.
Deilemera signata, Butler, P. Z.8., 1878, p. 586.
Nyctemera signata, Pag., Jahrb. Nass. Ver. Naturk., 1891,
p. 152.
The type female from Darnley Island; it may be ar
extreme form of pe//ex, but it is impossible to say, without
seeing more examples.
DEILEMERA EGROTA.
Leptosoma xgrotum, Swinhoe, Cat. Het. Mus. Oxon., 1,
p- 145, pl. v, f. 15 (1892).
The Genus Deilemera. al
Nyctemera xgrotwm, Pag., Jahrb. Nass. Ver. Naturk., 1901,
QF
p: 135.
Two females from Queensland and one male from S.E.
Australia. Type from N.S. Wales in Mus. Oxon.
DEILEMERA SIMPLEX.
Nyctemera simplex, Walker, xxxi, 207 (1864).
Atasea simplex, Swinhoe, Cat. Het. Mus. Oxon., i, p. 140,
pl. 5, £ 18 (1892).
Nyctemera dovix, Oberth., Ann. Mus. Genov., xv, pl. 4, f. 2
(1883).
The type from N. Guinea is in Mus. Oxon. Oberthiir’s
type came from the same locality.
It is not in the B. M.
DEILEMERA QUADRIPLAGA.
Nyctemera quadriplaga, Walker, xxxi, 207 (1864).
Atasca quadriplaga, Swinhoc, Cat. Het. Mus. Oxon., 1,
p. 140, plo, & 991892).
The type from New Guinea is in Mus. Oxon.
It is not in the B. M.
B. ANTENNE SHORTLY PECTINATED. . . (P2tasilw)
DEILEMERA VARIANS.
Nyctemera varians, Walker, 11, 400 (1854).
Pitasila varians, Butler, Ill. Het. B. M., v, p. 46, pl. 88,
f. 4 (1881).
Pitasila moolaica, Moore, P. Z.8., 1878, p. 847, pl. 53, f. 10.
Fourteen examples from Kashmir, Sikkim, Cachar,
Karen Hills, Toungyen and Tenasserim, including both
types.
DEILEMERA INCONSTANS.
Pitasila inconstans, Butler, P. Z.S., 1880, p. 672.
5 Es Swinhoe, Cat. Het. Mus. Oxon.,i, p. 139.
(1892).
Six examples from Formosa, including the type, one
from Carniguen, and one from Tizard Bank, China.
DEILEMERA SEMPERT, nov.
d. Palpi black, face, frons, head and fore part of thorax ochreous,
a black spot on the frons, one on top of head, two on the collar, two
58 Colonel C. Swinhoe on
on each patagia, two transverse black bands on the thorax in front,
and a black spot behind, lower half of thorax and abdomen white, the
latter with a dorsal row of black spots, one on each segment, and
lateral double rows of black dots, anal tuft ochreous ; fore-wings
black, with white markings, much as in P. disrupta, Butler, from the
Solomon Islands, but there is an additional band of three white spots
between the basal streak and the discal band, the latter is entire and
is composed of five large spots: the hind-wings are white with a
black marginal band containing two apical spots, and another a little
below the middle, as in disrupte.
Expanse of wings 2,'; inches.
Hab, PALAWAN (Doherty).
One example; there are several from the Philippines
in Mr. Herbert Druce’s coll. It is the insect figured by
Herr Semper as Pitasila leucospilota, Moore, in his great
work on the Phil. Schmett, Heterocera, pl. 59, f. 6;
though of course allied to that species it is absolutely
distinct from it.
DEILEMERA BIJUNCTELLA.
Nyctemera bijunetella, Walker, xxxv, 1880 (1866).
Pitasila bijunctella, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 139
(1892),
Leptosoma maculosum, Felder (nec Walker), Reise Nov.
Lep., pl. 1038, f. 2 (1869).
The type example from the Philippines.
DEILEMERA BURICA,
Nyctemera burica, Holland, Nov. Zool., vu, p. 560 (1900).
be » Lag., Jahrb. Nass. Ver. Naturk., 1901,
p. 156.
One example from Sula Mangoli.
The type came from Buru.
DEILEMERA DISTINCTA, nov. (Pl. IV, f. 4.)
d ¢@. White, head, thorax and tip of abdomen tinged with
ochreous, last joint of palpi brown, one black spot on frons, two
on collar and five on thorax, and a duplex row of black dots on the
sides of the pure white abdomen ; wings with pale black bands, very
much thinner in the female than in the male ; on the fore-wings the
basal two-thirds of the subeostal vein is broadly black and is joined
to a spot on the costa before its end, and connected with the hinder
The Genus Deilemera. 59
margin by two bands, forming a circle ; there is a spot on the costa
near the base, another on the hinder margin beyond the middle,
submarginal and marginal irregular bands, the former almost con-
nected by a streak with a spot on the costa above the end of the cell ;
hind-wings with an outer marginal band, which is disconnected in its
middle and has an apical white spot in the male, and is disconnected
both at the middle and near the apex in the female.
Expanse of wings 2 inches.
Hab, SANGIR (Doherty).
Two males and one female.
DEILEMERA OROYA, nov. (PI. LV, f. 8.)
Q. Palpi with the second and third joints black, first joint white,
antennze black ; head, body, legs and wings white, a tinge of ochreous
on face, head, shoulders, and tip of abdomen ; a black spot on the
frons, one on the head, two on the collar, five on the thorax, a black
dot on each segment of the abdomen on the sides, and a black line
below them divided by the segments : wings with the bands and
spots pale black ; fore-wings with a spot on costa before the middle,
one at upper end of cell, a submarginal irregular band, ending at
the hinder angle and joined to the outer margin below the apex,
forming a large subapical white spot or space, a thickening on the
basal half of subcostal vein, a slight thickening at the origin of vein
2 and another on the discoidal vein; hind-wings with an outer
marginal band, containing two submarginal spots, subapical and
medial.
Expanse of wings 1 8, inches.
Hab. SULA Best (Voherty).
Two examples.
DEILEMERA ABRAXOIDES.
Nyctemera abraxoides, Walker, Journ. Linn. Soc. Lond., vi,
p. 93 (1862).
Pitasila abravoides, Swinhoe, Cat. Het. Mus. Oxon., i,
p. 139 (1892).
Two examples from Borneo, including the type.
DEILEMERA AMOSA, nov. (Pl. IV, f.6.)
3 2. Palpi, head and thorax yellow, a black spot between the
antennee, another behind it, four on the collar and four on the thorax,
abdomen white, anal tuft yellow ; fore-wings pale chocolate-erey, a
white spot before the middle divided by the median vein, a longer
60 Colonel C. Swinhoe on
white spot beneath it in the hinder margin, a broad discal white
band, from the costa a little beyond the middle, widening in its
centre and attenuated towards the hinder margin, three white sub-
marginal spots as in Atasca pellex, Linn. ; hind-wings white with a
broad chocolate-grey even band, containing two submarginal white
spots, one subapical, the other beyond the middle.
Expanse of wings 13 inches.
Hab. DAMMA ISLAND.
DEILEMERA DISRUPTA.
Pitasila disrupta, Butler, Ann. Mag. N. H. (5), xix, p. 223
(1887).
Six examples from the Duke of York Island, Kiriwini
and Alu, including the type.
DEILEMERA SELECTA.
Nyctemera selecta, Walker, 1, 399 (1854).
~ » Lag., Jahrb. Nass. Ver. Naturk., 1901,
p. 156. .
Six examples from Alu, and the type without locality.
DEILEMERA VARIOLOSA.
Nyctemera variolosa, Felder, Reise Nov. Lep., pl. 129, f.
15 (1875).
Pitasila variolosa, Cotes and Swinhoe, Cat. Het., p. 80
(1887).
Two examples from the Andaman Islands and one from
the Nicobars.
DEILEMERA LEUCOSPILOTA.
Nyctemera leucospilota, Moore, P. Z.8., 1887, p. 509, pl. 58,
tee
Pitasila leucospilota, Kirby, Cat., p. 422 (1892).
Hight examples from the Andaman Islands, including
the type.
This form and disrupia, Pagenstecher sinks as synonyms
to selecta (p. 156): Sir George Hampson sinks abraxoides,
byunetella, and disrupta, omittiug inconstans. I cannot
agree with either. Semper reared some from larve, but there
is nothing to show that the eggs of one female produced
two forms: dijunctella and inconstans are nearest to each
The Genus Deilemera. 61
other in pattern, and may be local forms of the same thing,
abraxoides seems to be very distinct: the members of each
species shown above do not vary, there is not one inter-
mediate form amongst them, and therefore unless proved
otherwise by breeding they must be kept distinct.
DEILEMERA GUTTULOSA.
Nyctemera guttulosa, Walker, xxxi, 201 (1864).
Pitasila guttulosa, Swinhoe, Cat. Het. Mus. Oxon., i, p. 138
(1892),
Nyctemera abrazxata, Snellen, Tijd. v. Ent., xxii, p. 73, pl. 6,
f. 6 (1879).
One example (the type) from Celebes; there are two
from the same locality in Mus. Oxon.
DEILEMERA SPECULARIS.
Nyctemera specularis, Walker, vii, 1665 (1856).
Pitasila specularis, Swinhoe, Cat. Het. Mus. Oxon., i, p. 139
(1892).
Two examples including the type from Ceram; there are
two also from the same locality in Mus. Oxon.
SECTION II.
Veins 6 and 7 of hind-wings stalked, palpi longer, antennze with
long pectinations.
A, BOTH WINGS OF MALE ABNORMALLY DEVELOPED, (Deilemera)
DEILEMERA EVERGISTA.
Phalena (Geometra) evergista, Cram., Pap. Exot., iv, p. 155,
pl. 369, f. E (1781).
Deilemera evergista, Swinhoe, Cat. Het. Mus. Oxon.,1, p. 147
(1892).
Two examples from Amboina.
DEILEMERA UNIPLAGA, nov. (PI. III, f. 2.)
é- Palpi with first joint ochreous, second and third joints black,
ochreous inside and with ochreous tips; antenne black ; head,
thorax and abdomen dark ochreous with black bands and spots, a
spot on the frons, one behind base of antenne, four on the collar,
three on the thorax, and a band on each segment of the abdomen ;
fore-wings blackish-brown with a large white patch in the disc,
extending from below vein 3 across the end of the cell to the costa
where it is narrowest ; hind-wings white with a broad blackish-brown
62 Colonel C. Swinhoe on
marginal border, which suddenly becomes diffuse and ends after
rounding the angle: on the under-side the wings are as on the upper-
side, quite as dark, the veins in the fore-wings pale and a subapical
white dot.
Expanse of wings 2;'; inches.
Hab, Frerausson Ist. (A. S. Meek).
Two examples; wings shaped as in D. evergista, Cram.
Pagenstecher in Jahrb, Nass. Ver. Naturk., 1901, p. 165,
sinks mutabilis, Walker, from Ternate (noted in Cat. Het.
Mus. Oxon., 1, p. 147), to evergista, Cram., but this cannot
hold, the white patch in the disc of fore-wing inclines
inwards, whereas in evergista it inclines outwards and is
altogether of a different nature.
DEILEMERA INTERCISA.
Nyctemera wntercisa, Walker, xxxi, 205 (1864).
Deilemera intercisa, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 147
(1892).
The type from Amboina is in Mus, Oxon. It is not in
the B. M.
DEILEMERA MACULATA.
Nyctemera maculata, Walker, 11, 396 (1854).
. a Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 1682, 1.0:
Nyctemera noviespunctatum, Voll., Tid. v. Dierk., 1, p. 42
(1863).
One example (the type) without locality, from Children’s
coll., probably Java.
DEILEMERA ZRES.
Leptosoma exes, Boisd., Voy. de lAstrolabe Lep., p. 198
(1832).
Nyctemera mutabilis, var., Walker, xxxi, 206 (1864).
Deilemera mutabilis, Swinhoe (note), Cat. Het. Mus. Oxon.,
i, p. 147 (1892).
Walker’s type from Ternate and one example from
Buru.
DEILEMERA LEUCTRA, nov. (PI. IV, f. 5.)
49. Palpi with the first joint yellow, second and third black,
head, thorax and abdomen ochreous yellow, with black spots and
The Genus Deilemera. 63
bands, a spot on the frons, one behind the base of the antenne, four
on the collar, five on the thorax and broad segmental bands on the
abdomen ; fore-wings black, witha large white patch (as in D. miilleri,
Voll.) at the base, a larger white patch covering the whole of the dise,
extending from near the costa to near the hinder margin, its inner
margin recurved, its outer margin outwardly dentated, its lower side
having a small square projection, two submarginal white spots, one
a little above the middle, the other larger and subapical ; hind-wings
white with a broad black band, with two white spots as in the fore-
wings, the band complete in the female, but in the male it terminates
abruptly at vein 2, and is followed by two black spots.
Expanse of wings 2,1; inches.
Hab. SANGIR (Doherty); TALAUT (Doherty).
Allied to D. miilleri, Voll, from Borneo, which is very
distinct from D. cavissima, Swinhoe, from Assam. Pagen-
stecher on p. 164 puts them together ; though the pattern
of the wings is somewhat similar the shape of the two
insects is very different, carissima having long narrow
wings and in the male an excavation in the hind-wings
a little before the anal angle, making the angle produced
hindwards.
DEILEMERA GERRA, nov. (PI. IV, f. 1.)
¢ 2. Palpi, head and body as in D. leuctra ; fore-wings blackish-
brown, a white basal patch narrower than in Jevctra, a narrow discal
white patch with a round excavation on its inner side, toothed on its
outer side, but with no square projection hind-wards ; hind-wings
white with a broad blackish-brown band with the white spots as in
leuctra, but without the two black spots in the male in continuation
of the marginal band.
Expanse of wings 2 inches.
Hab, TALAvT (Doherty).
The white bands on the fore-wings of the female are
rather broader than those in the male.
DEILEMERA MULLERI.
Leptosoma miilleri, Voll., Tijd. v. Dierk, i, p. 41 (1863).
Deilemera miilleri, Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 164.
Two males and three females from Borneo.
64 Colonel C. Swinhoe on
DEILEMERA CARISSIMA.
Deilemera carissima, Swinhoe, Trans. Ent. Soc., Lond., 1891,
p. 447, pl. 19, f. 1.
Deilemera carissina, Hampson, Moths, India, ii, p. 46
(1894). -
One male and one female (types) from the Khasia Hills;
T have also a pair from the same locality in my museum.
DEILEMERA ZERENOIDES.
Tripheromera zerenoides, Butler, Ann. Mag. N. H. (5), viii,
p. 380 (1881).
One female (the type) from Sumatyra.
It more nearly resembles carisstma, Swinhoe, than
anything else.
DEILEMERA ARCTATA.
Nyetemera arctata, Walker, vii, 1664 (1856).
Deilemera arctuta, Hampson, Moths, India, ui, p. 45, f. 21 2
(1894).
Nyctemera maculosa, Walker, xxxi, 198 (1864).
Five examples from Sikkim, Assam, and Burma; the
type from Cherra Punji in the Khasia Hills, Assam, is in
Mus. Oxon.
B. MALE WITH LONG TUFT OF HAIR TO FORE TIBI&.
(coleta group)
DEILEMERA COLETA.
Phalena coleta, Cram., Pap. Exot., iv, pl. 368, f. H-(1781).
Nyctemera coleta, Hiibner, Verz. Schmett., p. 178 (1818).
_ _ Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 142.
Sixteen examples from Tenasserim, Singapore, Sumatra,
Borneo, Java, Amboina and Ceram.
DEILEMERA NIGROVENOSA.
Nyctemera nigrovenosa, Moore, P. Z. S., 1879, p. 394.
Swinhoe, Cat. Het. Mus. Oxon., 1, p.
>
141 (1892).
The Genus Deilemera. 65
Seven examples from Ceylon, including the type;
Pagenstecher following Hampson has put it as a synonym
to coleta.
DEILEMERA MELANEURA.
Leptosoma melaneura, Butler, Ann. Mag. N. H. (5), xii, p.
160 (1888).
2 “5 Weymer, Stett. Ent. Zeit., xlvi, p.
974 (1885).
The type example from Nias.
In my collection are two from Nias and some in Mr.
Herbert Druce’s collection ; it seems to be confined to the
Island and is quite distinct from either coleta or nigrovenosa.
C, MALE WITH ABNORMAL HIND LEGS. . (leucono’ group)
DEILEMERA LEUCONOE.
Nyctemera leuconoé, Hopfter, Berl. Mon. Acad., p. 422
(1857).
Leptosoma leuconoé, Butler, P. Z.8., 1898, p. 419.
Three examples from Abyssinia, one from Foda, four
labelled E. Africa, two from Nyassa Land, two from Zomba,
one from Delagoa Bay, and six from Natal.
DEILEMERA INSULARIS,
Leptosoma insulare, Boisd., Faun. Madag., p. 84, pl. 12, f 1
(1833). ;
Three examples from Madagascar.
DEILEMERA CONSORS.
Leptosoma consors, Butler, Ann. Mag. N. H. (5), xv, p. 192
GUSTO):
The type from the Island of Johanna.
DEILEMERA FALLAX.
Nyctemera fallax, Holland, Ent. News. and Proc. Ent. Soc.
Ac. Nat. Sci. Phil., iv, p. 59 (1893).
F Pag., Jahrb. Nass. Ver. Naturk., 1901, p.
147.
One example from Ogove River ; the type came from the
same locality and is in coll. Holland.
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL) 5
66 Colonel C. Swinhoe on
DEILEMERA PERSPICUA.
Nyctemera perspicua, Walker, 11, 898 (1854).
Ten examples from Sierra Leone, Old Calabar, and the
Congo, including the type.
DEILEMERA BIFORMIS.
Nychthemera biformis, Mab., Bull. Soc. Zool. Fr., ii, p. 87
(1878).
Nyctemera biformis, Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 149.
Nyctemera mabillei, Butler, Ent. Mo, Mag., xix, p. 57
(1882).
Two males and one female (Butler's type) from
Madagascar.
In Ent. Mo. Mag., xix, p. 57, Butler says, that as they
have received in the B. M. a black female from Madagascar
almost exactly like the male, the white insect described
by Mabille cannot be the female of biformis. J have care-
fully examined thisso-called black female ; it is not a female,
but a male with frenulum and retinaculum complete,
and I cannot therefore see why this white female should
not be the female of Jiformis, and it must stand as such
until proved otherwise.
DEILEMERA APICALIS.
Nyctemera apicalis, Walker, 1, 395 (1854).
- 2 Waterhouse, Aid, pl. 178, f. 4 (1880).
. A Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 145.
One example (type) from W. Africa, and one from
Ashanti.
Kirby put tricolor, Felder, as a var. of this species,
and Pagenstecher does the same; but judging from
Felder’s figure I do not believe this to be correct, apicalis
having a very broad discal pure white band.
DEILEMERA RESTRICTA.
Leptosoma restrictum, Butler, P. Z.S., 1894, p. 585.
Nyctemera restrictum, Pag. Jahrb. Nass. Ver. Naturk.,
1901, p. 146.
Four examples from E. Africa, including the type.
The Genus Deilemera. 67
DEILEMERA ANTINORILI.
Nyetemera antinorii, Oberthiir, Ann. Mus. Genova, xv,
po les; plot; fT (sso):
Two from Sierra Leone, two from Mongo-ma-Lobak,
nine from Old Calabar, three from the River Nigra, and
one from the Cameroons.
DEILEMERA ACRAINA.
Nyctemera acreina, Druce, P. Z. 8., 1882, p. 780.
Type, Calabar in coll. Druce.
It is not in the B. M.
DEILEMERA CHROMIS.
Nyctemera chromis, Druce, P. Z. S., 1882, p. 780, pl. 65, f. 2.
The type from W. Africa is in coll. Druce.
It is not in the B. M.
DEILEMERA FULLERL.
Nyctemera fulleri, Drace, Ent. Mo. Mag., xx, p. 157 (1883).
The type from the Cameroons is in coll. Druce.
It is not in the B. M.
D. WITHOUT SECONDARY SEXUAL CHARACTERS. — (7'ripheromera)
DEILEMERA PLAGIFERA.
Nyctemera plagifera, Walker, u, 400 (1854).
Tripheromera plagifera, Butler, Il. Het. B. M., v, p. 45,
pl. 88, f. 3 (1881).
Twenty-five examp'es from W. China, Loochoo, Hong
Kong, Nepaul, Darjiling, Assam, Silhet (type), and the
Nilgiri Hills.
DEILEMERA LUDEKINGII.
Leptosoma ludekingit, Voll., Tijd. v. Dierk., 1, p. 49 (1863).
F Snellen, Tijd. v. Ent., xlii, p. 108,
pl os b. t h (1S99)
> ys Pag., Jahrb. Nass. Ver. Naturk.,
Osea pleat, Lill,
One example from Sumatra. Pagenstecher’s figure is
better than Snellen’s, Pagenstecher puts zerenoides,
68 Colonel C. Swinhoe on
Butler, as a synonym, but that species belongs to a dif-
ferent section of the Family.
DEILEMERA CENIS.
Phalena cenis, Cram., Pap. Exot., 1, pl. 147, f. E (1777).
Nyctemera interlectum, Walker, 11,400 (1854).
Fifteen examples from Loo Choo, Shan States, Kashmir,
Dera Dhun, Chin Hills, Jawai Hills, Darjiling, Sikkim,
Assam, and Walker's type from Silhet.
DEILEMERA ANNULATA.
Leptosoma annulatum, Boisd., Voy. de Astrolabe Lep., p.
MES alloaay ih, 2) (iets 74)
Nyctemera annulata, Pag., Jahrb. Nass. Ver. Naturk., 1901,
pe
Nyctemera doubledayi, Walker, 11,392 (1854).
Fourteen examples from New Zealand including Walker's
type.
DEILEMERA AMICA.
Aglages amicus, White in Grey’s Journ. Exped. Austral.,
ii, p. 482 (1841).
Nyctemera amica, Meyrick, Proc. Linn. Soc. N.S.W. (2), 1,
p. 760 (1886).
Nyctemera annulata, Walker (nec Boisd.), 11, 391 (1854).
Leptosoma plagiatum, Guen., Ent. Mo. Mag., v, p. 2 (1868).
Nyetemera conica, Pag., Jahrb. Nass. Ver. Naturk., 1901,
(oe
Eleven examples from $.E. Australia; Pagenstecher’s
conica is evidently a misprint for @mzcc.
DEILEMERA SECUNDARIA.
Leptosoma secundariwm, Lucas, Proc. Linn. Soc. N.S.W.,
vi, p. 280 (1891).
Four examples from Queensland, ditfering nearly as
much from amica as amica does from annulata.
DEILEMERA LATEMARGINATA. (PI. III, fi 4.)
Nyctemera latemarginata, Pag., Jahrb. Nass. Ver. Naturk.,
1901;-p) 14:
One example from N. Guinea.
The Genus Deilemera. 69
It has only two spots on the fore-wings, and is therefore
not absolutely identical with Pagenstecher’s type, but I
am convinced it must belong to the same species.
DEILEMERA DISTINCTA.
Nyctemera distincta, Walker, 11, 892 (1854).
-f : Pag., Jahrb. Nass. Ver. Naturk., 1901,
PaclO SH ole 2ecteele
Five examples from Java including the type. Pagen-
stecher’s figure does not well represent the typical form ;
the type has hardly an indication of the white discal band
of fore-wings, and in none of the examples in this Museum
is the band plainly shown, and the yellow streaks are very
faintly indicated.
DEILEMERA Poplya, nov. (PI. III, figs. 3 and 7.)
¢ Q. First joint of palpi bright ochreous, second and third black, a
large black spot on frons, one behind the base of antennz, twoon collar,
one on each shoulder, three black longitudinal stripes on the thorax,
all the spots and stripes edged with bright ochreous ; fore-wings dark
blackish-brown above and below, a broad white discal band on the
under-side, from vein 2 to the costa; vein 1, the median and sub-costal
veins white ; on the upper-side the broad white band can be very dis-
tinctly seen, and all the veins on the basal third are yellow ; the hind-
wings are white with a broad blackish-brown border and with some
brown suffusion at the base; cilia whitish ; abdomen grey with
white segmental bands and gold tip.
Expanse of wings 1;%5 inches.
Hab. JAVA, six examples.
On the upper-side it is something lke distincta, but
the band on the hind-wings is much narrower, and the
abdomen is whiter, pure white beneath, with a double
row of black spots on each side ; underneath the markings
of fore-wings are very distinctive.
DEILEMERA TRITA.
Nyctemera trita, Walker, u, 394 (1854).
rr 7 Moore, Cat; ep. EAC. ix) p. 331, pl. 8a,
f. 9 (1858).
Leptosoma tritum, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 148
(1892).
Nine examples from Java, including the type.
70 Colonel ©. Swinhoe on
DEILEMERA SUBVELATA.
Nyctemera subvelata, Walker, xxx1, 200 (1864).
Leptosoma subvelatwm, Swinhoe, Cat. Het. Mus. Oxon., 1,
p. 144 (1892).
Leptosoma infuscata, Hoptier, Stett. Ent. Zeit., xxxv, p. 44
(1874).
Nyctemera infuscata, Pag., Jahrb. Nass. Ver. Naturk., 1901,
p: 129.
Three examples from Celebes, including Walker’s type.
I have examples in my own Museum from Celebes that
have been identified as in/uscata by Pagenstecher.
DEILEMERA CLATHRATA.
Leptosoma clathratum, Voll., Tijd. v. Dierk., 1, p. 48 (1863).
Nyctemera clathratum, Pag. Jahrb. Nass. Ver. Naturk.,
TOOM os USC ipl 2a 2s
One male from Ceram which seems identical with
Pagenstecher’s figure 1s in my coll.
lt is not in the B. M.
D&EILEMERA ASSIMILE.
Leptosoma assinile, Voll.. Tijd. v. Dierk., 1, p. 89 (1863).
Nyetemera assimile, Pag., Jahrb. Nass. Ver. Naturk., 1901,
pe LOS ple 2.
Leptosoma confusum, Swinhoe, Cat. Het. Mus. Oxon., 1, p.
144 (1892)
One example from Java. The types of confusum from
Java are in Mus. Oxon.
DEILEMERA ALTERNATA.
Nyctemera alternata, Walker, xxxv, 1879 (1866).
* - Semper, Phil. Sclimett., p. 495, pl. 58,
Peo Los S99):
The type example from the Philippines.
DEILEMERA RADIATA.
Nyctemera radiata, Walker, vu, 1664 (1856).
7 Semper, Phil. Schmett., p. 494, pl. 58,
f, 8, 2 (1899).
Leptosoma radiata, Swinhoe, Cat. Het. Mus. Oxon., i, p. 142,
luo, f LOEIS02);
The Genus Deilemera. 71
Two females from the Philippines.
The type from Manilla is in Mus. Oxon.
DEILEMERA VELANS.
Nyctemera velans, Walker, xxxi, 200 (1864).
Leptosoma velans, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 144,
pl. 5,.f£ 7 (1892).
Nyctemera velans, Pag., Jahrb. Nass. Ver. Naturk., 1901, p.
131.
The type from Celebes is in Mus. Oxon.
It is not in the B. M.
DEILEMERA LOMBOKIANA, Nov.
¢ 2. Palpi with first joint ochreous, second and third black, frons
and head pale ochreous, nearly white, a black spot on frons, one on
head and two on collar, all large ; thorax and abdomen dark brown,
the former with longitudinal pale yellow lines, the latter with white
hairs and indistinct whitish segmental bands and ochreous tip ;
wings of auniform dark brown ; fore-wings with veins yellowish on
basal third, an indistinct discal band of six inconspicuous small
spots, the last two the largest, the others uniform in size and very
small, all of them suffused more or less with brown ; hind-wings
with a small central space whitish.
Expanse of wings 1,5, inches.
Hab. LOMBOCK, one pair.
Received from a Continental dealer as NV. lombokiana,
Fruhstorfer; but I can find no reference: it is allied to
assimile, Voll., and propria, Swinhoe.
DEILEMERA PROPRIA.
Leptosoma proprium, Swinhoe, Cat. Het. Mus. Oxon., 1,
p. 144, pl. 5, f. 12 (1892).
Nyctemera proprium, Semper, Phil. Schmett., p. 495, pl. 58,
fi 11 3; 12; 13; 14 2 899).
The types f 9 from the Philippines are in Mus. Oxon.
Semper records it from the Babuyan Islands, Bohol,
Camotes, and East Mindanao. It is not in the B. M.
DEILEMERA QUADRIGUTTATA.
Leptosoma quadriguttatwm, Voll., Tijd. v. Dierk., 1, p. 40
(1863).
Nyctemera quadriguttatum, Pag., Jahrb. Nass. Ver. Naturk.,
1901, p. 114, pl. 2, f. 9.
~T
SJ)
2 Colonel C. Swinhoe on
One example from Java.
Not the same as hondekwm, Swinhoe, as Pagenstecher
says: that species being more nearly allied to distincta,
Walker.
DEILEMERA SEXMACULATA,
Leptosoma seemaculatum, Butler, Ann. Mag. N. H. (5), xix,
222 (1887).
Nyctemera aolensis, Druce, P. Z.8., 1888, p. 573.
Two males and four females, Alu, Solomon Islands,
including Butler’s type ; Druce’s type came from Guadal-
canar, Solomon Islands; there is much less white in the
females than in the males, especially on the hind-wings.
DEILEMERA EXTENDENS.
Nyetemera extendens, Walker, vii, 1666 (1856).
. Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. L2G plo eel
Nyctemera horites, Druce, P. Z.8., 1888, p. 573.
Walker’s type from New Hebrides and nine examples
from Alu, Solomon Islands; Druce’s type, a female, also
came from the Solomon Islands. I have it in my coll.
from Treasury Isl. and New Georgia.
DEILEMERA SYRNIA, nov. (PI. III, f. 6.)
2. Palpi with the first joint dull orange, second and third joints
black, large deep black spots edged with pale orange, filling up all
the spaces, one on the frons, one behind the base of antenne, two on
the collar, one on each shoulder, and three longitudinal stripes
similarly edged filling up the entire surface of the thorax ; abdomen
black, tip ochreous, segmental bands thin and white above, broad
and ochreous beneath ; fore-wings deep black, vein 1 and the median
vein slightly ochreous towards the base, a discal white band, divided
by the veins into seven spots, much indented, almost dislocated at
the lower end of the cell, the upper spot near the costa very small,
the fourth and seventh much smaller than the rest, the seventh just
below vein 2; hind-wings with a broad deep black marginal band,
scooped inwards from veins 2 to 5, extending along the costa and
partly up the abdominal margin.
Expanse of wings 1,8, inches.
Hab, FERGUSSON ISLAND (Meek).
Two examples.
The Genus Deilemera. ie
DEILEMERA KAPAURENSIS, nov. (PI. IIT, f. 1.)
Q. Allied to syrnia, fore-wings not quite so deep black, all the
veins in the basal half whitish, the discal band similarly shaped, but
broader, the subcostal spot as large as the lowest spot, the band
extending below vein 1 forming eight spots, a white stripe in the
basal two-thirds of the hinder margin ; hind-wings as in syrnia, but
the costal band with a short production hindwards in its centre ; the
abdomen also differs in being ochreous below, with black segmental
macular bands.
Expanse of wings 1,5, inches.
Hab. Kapaur, N. Guinea (Doherty).
One example.
DEILEMERA DRUCEI, nov.
¢ 9. Palpi black, ochreous beneath and at the base, face and head
ochreous, but with the spots so large as to make them look black,
thorax black with ochreous longitudinal stripes, abdomen black ; with
ochreous segmental bands and ochreous tip ; fore-wings deep black, in
the male there is a white mark below end of cell which looks like the
termination of a stripe from the base, this is wanting in the female ;
a white discal band, not macular, fairly uniform in width, curved in-
wards, with two indentations on its inner side ; on the basal third the
veins are s'ightly ochreous, and there is a white streak on the hinder
margin, from the base to two-thirds its length ; hind-wings white
with the usual black marginal band, without any spots.
Expanse of wings 1,°; inches.
Hab, N. AUSTRALIA, types in Mus. Druce ; not in B. M.
The only species in the Family I have yet seen with the
discal band of fore-wings curving inwards.
DEILEMERA DISPAR, nov.
¢. Palpi with first and second joints ochreous, last joint black,
frons, head and collar ochreous, frons with one black spot, head with
a large black spot, and the collar with two, oecupying nearly the
whole space, thorax and abdomen black, the former with ochreous
longitudinal stripes, the latter with segmental white bands, which go
completely round the abdomen, are ochreous in the last two segments,
and there is an ochreous stripe along each side containing black spots ;
tip orange ; fore-wings black, veins without markings, the discal
white band broad, extending almost to the hinder margin near the
angle, divided by the veins into seven spots, much as in D. alwensis,
74 Colonel C. Swinhoe on
Butler, but closer together and more compact ; hind-wings white,
with the usual black band on the margins.
Expanse of wings 1,§, inches.
Hab, Rosset IsLtaAnp, Louisiade Group.
Its black colour, without any vein markings, separates it
from the mundipicta group, which it otherwise somewhat
resembles.
DEILEMERA LACTICINIA.
Phalena (Geometra) lacticinia, Cram., Pap. Exot., ii, pl. 128,
ee DOW iCO
Leptosomna lacticinia, Swinhoe, Cat. Het. Mus. Oxon., 1,
p- 141 (1892).
Fifteen examples from Hong Kong, Canara, Cachar,
Malabar, Nilgiri Hills, Ceylon, and Penang; the Penang
examples have the discal band of the fore-wings rather
wide, but seem otherwise identically the same as those
from India.
DEILEMERA SIMULATRIX.
Nyctemera simulatrix, Walker, xxxi, 198 (1864).
Leptosoma simulatriz, Swinhoe, Cat. Het. Mus. Oxon., 1,
p. 143 (1892).
One example from Celebes. The type from Celebes
is in Mus. Oxon.
DEILEMERA NIGROVENA, nov. (Pl. IV, fig. 2.)
Q. First two joints of palpi dull orange-yellow, last joint black,
head and thorax orange-yellow, one black spot on the frons, one
behind the base of antennze, two black bands on the collar, a spot on
each shoulder, and three longitudinal bands on the thorax ; abdomen
white above with black segmental bands, below slightly yellow
tinged, with the bands macular ; fore-wings black, the veins on the
basal half white, discal band composed of eight spots as in mundipicta,
but well separated from each other by the veins, except the spot at
the end of the cell, which is joined to the spot outside the end ;
hind-wings white, with the usual broad marginal band, with all the
veins black ; veins 1, la, the median, and subcostal veins with a thin
band of black suffusion on each side.
Expanse of wings 2,°, inches,
Hab, SoutH CELEBES (Pruhstorfer).
One example.
The Genus Deilemera.
SY
OU
DEILEMERA KALA.
Leptosoma kala, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 143,
pl. 5, f. 8 (1892):
The type from Ké Island is in Mus. Oxon.
It is not in the B. M.
DEILEMERA BAULUS.
Leptosoma baulus, Boisd., Voy. de Astrolabe Lep., p. 200
(18382).
Nyctemera mundipicta, Walker, Saran Linn. Soc. Lond.,
il, p. 184 (1859).
Leptosoma mundipicta, Swinhoe, Cat. Het. Mus. Oxon., 1
p. 141, pl. 5, f. 14 (1892).
Nyetemera herklotsit, Voll., Tijd. v. Dierk., 1, p. 39 (1863
" fr Pag., Jahrb. Nass. Ver. Naturk., L901,
joe LST OL ai,
Thirty-five examples from New Britain, New Ireland,
Borneo, Singapore, Ternate, Amboina, Trobriand Island,
Fergusson Island, Sumatra, Java and Ceram; in the Oxford
Museum it is also from Flores, Gilolo, New Guinea,
Mysol, and New Caledonia. It seems to have a very wide
range, and is variable as to the width of the discal white
wa arlhire band of the fore-wings and the marginal band of
hind-wings. There are in the B. M. ten examples from
Java, some identical with Pagenstecher’s figure of herh/otsi,
and none of them separable from baulus, and also many
examples from Buru (where Boisduval’s type came from)
are in the Tring Museum, some of which I have examined.
DEILEMERA FASCIATA.
Nyctemera fasciata, Walker, vu, 1665 (1856).
Nyetemera latistriga, Snellen (nec Walker), Tijd. v. Ent.,
Oth tO, Ceol Gp a, 5) CUS)
. x Pag., Jahrb. Nass. Ver. Naturk.,
1901, p. 120.
Nyetemera latistriya, var. fasciata, Rober, Tijd. v. Ent.,
LOSING ]on Bd (SSH),
Nyctemera tertiana, Meyrick, Ent. Mo..Mag., xxii, p. 15
(1886).
~ . Fags lec, ps Lan
The type from Aneitum, and six examples from Fiji
76 Colonel ©. Swinhoe on
and Adelaide: it has been recorded from Celebes, Ceram,
Aru, N. Guinea, Flores, Timorlaut, Ulasser, Shortland
Island, and Solomon Islands; it 1s very doubtfully distinct
from baulus.
DEILEMERA INTEGRA. (PI. III, f. 5.)
Nyctemera integra, Walker, xxxv, 1879 (1866).
Leptosoma integra, Kirby, Ann. Mag. N. H. (6), ii, p. 187
(1889).
Leptosoma imundipicta, Swinhoe (part), Cat. Het. Mus.
Oxon ip. alee otal
(1892).
Semper, Phil. Schmett., p. 498,
pl. 50, f. 4 2 (1899).
One female example (the type) from the Philippines,
one from Alu, Solomon Islands, and two from the Caroline
Islands.
This species is very similar in some respects to avwndi-
preta, Walker, but it is blacker, and the spots forming the
discal band of the fore-wings are closer together and more
compact, the veins dividing them being white, not black ;
the latter have the abdomen yellower, but are otherwise
identical. The figure represents one of the Caroline Island
examples.
DEILEMERA ALUENSIS.
Nyctemera aluensis, Butler, Ann. Mag. N. H. (5), xix, p. 222
(1887).
, Pag., Jahrb. Nass. Ver. Naturk., 1901,
peo.
Two male and three female examples from Alu,
Solomon Islands, including the types.
5
DEILEMERA KINAGANANGA, nov.
gd @. Palpi with first joint ochreous, last two black ; frons, lead
and thorax ochreous, the first with one spot, one on the head, two on
the collar ; thorax with longitudinal black stripes, and a black spot at
the base ; abdomen white with black segmental bands above, three
rows of black spots on each side ; wings very much as in D, baulus,
3o0isd, = mundipicta, Walker, except that the hinder margin of
fore-wings has a white stripe from base to below the discal band, and
A . Land
The Genus Deilemera. mm
a prominent white streak in the interno-median interspace, also from
the base to near the discal band; the black band on the hind-wings
is as narrow as it 1s in lacticinia, Cram.
Expanse of wings 1,°, inches.
Hab. KINAGANANG, New Britain.
DEILEMERA KONDEKA.
Leptosoma kondekum, Swinhoe, Cat. Het. Mus. Oxon., 1,
p. 144 (1892).
One example from Java. Types from Java in Mus.
Oxon.
DEILEMERA ILLUSTRIS, nov.
¢ @. Last joint of palpi black, first and second joints, the head
and collar bright ochreous, a black spot on the frons, one on top of
head and four on the collar ; thorax and abdomen white, tinged with
yellow in parts, the last two joints of abdomen bright ochreous ;
thorax with three longitudinal black bands, abdomen with a black
band on each segment ; fore-wings brown, blackish on the outer half,
pale and tinged with green on the basal half; a white streak on
hinder margin for two-thirds its length, the veins on the basal half
white, a white streak through the middle of the cell, another white
streak much broader in the interno-median interspace from the base
to the discal band, where it runs into and forms the lowest spot of
the six that form the band; the second spot is large and kidney-
shaped, the first and third are the smallest, the fourth longer, the
fifth longest ; hind-wings with a black outer border, as in mundipicta.
Expanse of wings 2 inches.
Hab, Atv, Solomon Islands.
Two pairs, described by Dr. Butler as a/uensis, var., but
they are certainly quite distinct from that species.
DEILEMERA NISA, nov. (PI. IV, f. 7.)
6 ¢@. Palpi with the first joint ochreous, second and third black,
head and collar ochreous, a black spot on the frons, one on top of head,
two on the collar, one on each shoulder ; thorax and abdomen white,
the hind part of the former and the last two segments of the latter
ochreous ; three longitudinal black bands on thorax, and a black spot
on the yellow part ; abdomen with black segmental bands above, and
a double row of black spots on each side; fore-wings black, veins on
the basal half pure white, a white streak throughout the middle of
78 Colonel C, Swinhoe on
the cell to the diseal band, a much broader and prominent white
streak in the interno-median interspace, also from base to discal band,
the latter composed of white spots much as in bawlus, but broader
than usual, the large spot second from the costa nearly round, hinder
marginal border with a white streak for two-thirds its length ;
hind-wings white with marginal black band as in bavlus.
Expanse of wings 1,3, inches.
Hab. SANGIR (Doherty).
Male and female, the types, and one male marked
Celebes.
It somewhat resembles a@/wensis, Butler.
DEILEMERA OBTUSA.
Nyctemera obtusa, Walker, vu, 1666 (1856).
- ie Pag., Jahrb. Nass. Ver. Naturk., 1901,
joe dhs
Three examples from Celebes, including the type.
In Kiikenthal’s Reise (Abhandl. Senckenbergschen
Gesellschaft), p. 440, pl. 18, f 4, Pagenstecher described
and figured NV. consobiina, Hopffer. In Jahrb. Nass. Ver.
1901, p. 115, he says that this was a mistake, and that the
description and figure represent 1. obtusa, Walker; but this
also is an error, the figure being very unlike Walker’s
species, and representing a species with which I am not
acquainted.
DEILEMERA LATISTRIGA.
Nyctemera latistriga, Walker, 11, 397 (1854).
Leptosoma latistriga, Butler, Hl. Het. B. M., v., p. 44, pl.
88, f. 1 (1881).
Nyctemera inconstans, Voll, Tijd. v. Dierk., 1, p. 47 (1863).
Nyctemera consobrina, Hopter, Stett. Ent. Zeit., 1874,
p. 45.
Nyctemera arcuatum, Swinhoe (nec Voll.), Cat. Het. Mus.
Oxon., i, p. 142 (1892).
Twenty-two examples from Borneo, Silhet, Ceylon,
Burma, Sumatra, Java, Pulo Laut, and the type from
Maulmein.
Arcuatum, Voll., was wrongly identified in the B. M., and
this led me to sink itas asynonym to datistriga in Cat. Het.
Mus. Oxon. Snellen in Tijd. v. Ent., xli, p. 26, pl. 1, £3, 4,
5 (1898), says it belongs to the Chalcosiide ; Pagenstecher
(p. 114) makes consobrina, Hopfter, a distinct species ; he
The Genus Deilemera. 79
does not say he has seen Hopffer’s type, but the description
exactly fits /atistriga: “Der Raum zwischen mediana und
submediana ist auf der innern Fliigelhalfte durch eine
spindelformige oder ein langgezogenes Dreieck bildende
Liingsbinde ausgefiillt die aber die weisse Querbinde nicht
erreicht.” If this does not describe latistviga it describes
something very near it; the elongated white triangle in
the interno-median interspace together with the dese ription
of the discal band of fore-wings is peculiarly distinctive to
latistriga.
DEILEMERA ACCEPTA.
Leptosoma acceptum, Swinhoe, Cat. Het. Mus. Oxon., 1, p.
143, pl. byt. 3.1892).
Pag., Kiikenthal’s Reise in Abh,
Senckenb. Naturf, Ges. ., 1897, p. 440.
The type from Celebes, a female, is in Mus. Oxon.
I have in my collection a male from Flores (Everett),
which seems to me to be identical.
It is not in the B. M.
DEILEMERA SONTICA.
Leptosoma sonticum, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 142
(1892).
Nyctemera sonticum, Semper, Phil. Schmett., p. 494, pl.
f. 5 2 (1898).
The types ? 2 from the Philippines are in Mus. Oxon. ;
it is recorded by Semper from Luzon, Mindoro and Bohol ;
I have in my collection a male from Mindoro and a female
from Palawan.
It is not in the B. M.
DEILEMERA ABSURDA.
Leptosoma absurdum, Swinhoe, Cat. Het. Mus. Oxon., i, p.
143 (1892).
Nyctemera absurdum, Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 135.
The male type, from Salawatti Island, is im Mus. Oxon.;
it is not in the B. M.
SO Colonel C. Swinhoe on
DEILEMERA LUCTUOSA.
Leptosoma luctuosum, Voll., Tijd. v. Dierk., 1, p. 42 (1863).
Swinhoe, Cat. Het. Mus. Oxons 1p:
146 (1892).
Nyctemera crescens, Walker, xxxi, 204 (1864).
? >
One example from Morova, one from Batjan, and one
from Sula; the type of evescens from Morty is in Mus.
Oxon.
DEILEMERA GALBANA.
Leptosoma galbanum, Swinhoe, Cat. Het. Mus. Oxon., 1, p.
146 (1892).
Nyctemera galbanum, Semper, Phil. Schmett., p. 429, pl. 58,
heel 899):
The types ¢ 2 from the Philippines are in Mus. Oxon.
Semper records it from Luzon.
It is not in the B. M.
DEILEMERA ONETHA.
Leptosoma onetha, Swinhoe, Ann. Mag. N. H. (7), vu, p.
466 (1901).
The type example from New Britain.
DEILEMERA TRIPUNCTARIA.
Phalena tripunctaria, Linn., Syst. Nat., x, p. 523 (1758).
Phalena petulea, Sparrm. Ameen. Acad., xii, p. 500 (1769).
Nyetemera atralba, Hiibn., Verz. Schmett., p- 178 (1818).
Nyctemera suimatrensis, Heylarts, Compt. Rend. Soc. Ent.
Belg., xxxiv, p. xvii (1890).
Pag., Jahrb. Nass. Ver. Naturk.,
1901, p= 139s pl 2.1..6:
Eleven examples from Penang, Malacca, Lankawi, Singa-
pore. Sumatra, Borneo, and Siam.
The example from Sumatra in the National Collection is
identical with Pagenstecher’s figure of swmatrensis and is
not separable from the other examples of tr7punctaria.
The Genus Deilemera. 81
DEILEMERA CELSA.
Nyetemera celsa, Walker, xxxi, 199 (1864).
Leptosoma celsum, Swinhoe, Cat. Het. Mus. Oxon., 1, p. 144
(1892).
Four examples from Hong Kong and Foo-Chow; the
type from Siam is in Mus. Oxon.; it is the extreme
Eastern form of tripunctaria.
DEILEMERA PICATA.
Secusto picatus, Butler, Ann. Mag. N. H. (5), vii, p. 380
(1881).
The type example from Sumatra.
This is much hke regularis, but it is I think distinct;
the type example is that of a female with a white streak in
the interno-median interspace on fore-wimgs, and the discal
band is much broader ; regu/aris, of which there are both
sexes in the National Collection, has no sign whatever of
a discal streak and is well figured by Pagenstecher.
DEILEMERA REGULARIS.
Leptosoma regularis, Snellen, Mid. Sum. Lep., p. 384
(1880).
Nyetemera regularis, Pag.,
Pp 1395 ple ra 5:
Three examples from Sumatra.
Jahrb. Nass. Ver. Naturk., 1901,
DEILEMERA KINABALINA.
Nyctemera kinabalina, Snellen, Tijd. v. Ent., xlii, p. 110, pl.
5 B, f. 2 (1899).
One female example marked Borneo; it is considerably
larger then Snellen’s figure of the male, but is undoubtedly
the same species,
DEILEMERA HARCA.
Leptosoma harca, Swinhoe, Ann. Mag. N. H. (6), xii, p. 215
(1893).
Nyctemera herce (sic), .Holland (sic), Pag., Jahrb. Nass.
Ver. Naturk., 1901, p. 138.
Nine examples from Malacca, including the type.
There are some curious printer’s errors in Pagenstecher’s
paper. 2g
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL) 6
82 Colonel C. Swinhoe on
DEILEMERA OVADA, Nov.
2. Palpi white, third joint and top of second black ; frons, thorax
and abdomen white, slightly tinged in parts with yellow, a black
spot on frons, one on head, two on collar and one on each shoulder ;
thorax with brown longitudinal stripes, abdomen with no markings
above, a double row of brown stripes on each side, tip ochreous; fore-
wings pale bronzy-brown, a white spot on the costa with an indistinct
small streak below it, indicating the commencement of the usual discal
band, no other markings above ; below, the discal band is more
apparent, consisting of two large wedge-shaped spots disconnected
from each other; hind-wings white, with the usual marginal band,
of the same colour as the fore-wings.
Expanse of wings 15 inches.
Hab. WAINGAPO, Sumba Isls.
I have also a female from the same locality in my coll. ;
it is allied to nothing I know of.
DEILEMERA OPTATA. (PI. IV, f. 3.)
¢. First joint of palpi ochreous, second and third brown, frons
pure white with a black spot, top of head and collar ochreous, the
former with one black spot, the latter with two ; thorax white with
three longitudinal black stripes, abdomen white with grey dorsal
spots and some blackish spots on the sides towards the tip, which
is ochreous ; fore-wings pale greyish-fawn colour, a white stripe in
the interno-median interspace from base to near hinder angle, the
diseal band white, divided by the fawn-coloured veins and all run-
ning on to the margin, with the exception of the costal spot above
end of cell which is small ; hind-wings white with short fawn-coloured
indistinct streaks in the interspaces near the outer margin; under-side
same as above but with the fawn-coloured parts darker.
Expanse of wings 1;‘s inches.
Hab. SUMATRA, one example.
DEILEMERA AMPLIFICATA.
Tanada (?) amplificata, Walker, xxxii, 377 (1865).
Tripheromera amplificata, Kirby, Cat. Moths, 1, p. 423
(1892).
Nyctemera cydippe, Weymer, Stett. Ent. Zeit., xlvi, p. 274,
pl. 2, f. 8 (1885).
= es Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 134.
‘Three examples from Nias and the type without locality ;
The Genus Deilemera. 83
seems to be contined to the Island of Nias; there are a
number in Mr. Herbert Druce’s collection and in mine.
DEILEMERA ALBA.
Nyctemera alba, Pag., Jahrb. Nass. Ver. Naturk., 1901, p.
135.
Three examples from Viti and Samoa.
The following species, described as such, do not belong
to this Genus or Family :—
Nyectemera (2) vagata, Walker, xxxi, 208 (1864), N.
Australia.
Is a Noctuid near the genus Ipana, Jordan, Nov. Zool.,
iii, p. 54 (1896).
Nyctemera arcuatwm, Voll, Tijd. v. Dierkunde, 1, p. 45
(1863), Lombok.
This is said to be a Chalcosia, vide Snellen, Tijd. v. Ent.,
xli, p. 26, pl. i, f. 3 (1898) ; it was wrongly identified in the
B. M. collection.
Nyctemera (2) contructa, Walker, xxxi, 208 (1864), Mysol.
This is a Boarmid belonging to the genus Bursada.
The following species are not in the National Collection,
and are unknown to me :—
Nyctemera confiuens, Felder, Sitz. Ac. Wien., xliii, p. 38
(1861), Amboina.
Nyctemera aurwillu, Pag., Jahrb. Nass. Ver. Naturk., 1901,
p. 154 (note)=W. fasciata, Aurivillius, Ent. Tidskrift,
1897, p. 220, text f. 5 (praeocc.), Nyasaland.
Nyctemera melaneura, var. melas, Rober, Tijd. v. Ent., 1891,
p. 326, Ceram.
Nyctemera mesolychna, Meyrick, Trans. Ent. Soc. Lond.,
1889, p. 466, N. Guinea.
Nyctemera pagenstecheri, Pag., Jahrb. Nass. Ver. Naturk.,
1898, p. 199, Lombok.
Nyctemera quaternarium, Pag., Lep. Fauna, Bismark-Arch.,
li, p. 54, pl. 2, f 29 (1900), N. Ireland.
84 Colonel C. Swinhoe on
Leptosoma pallens, Voll., Tijd. v. Dierk., i, p. 45 (1863),
Java.
Nyctemera apensis, Semper, Phil. Schmett., p. 496, pl. 55
f. 9, Mindanao.
Leptosoma fuscipenne, Wallgn., Wien. Ent. Mon., iv., p. 161
(1860), Caffraria.
Leptosoma tricolor, Felder, Reise Nov. Lep., pl. 103, f. 3
(1874), Africa.
Nyctemera rasana, Mab., Ann. Soc. Ent. Fr., 1879, p. 304,
Madagascar.
Nyctemera vollenhovii, Snellen, Tijd. v. Ent., xxxiii, p. 276,
pl ii, f. 4 (1890), Tanah Djampea.
Nyetemera tenuifascia, Snellen, Tijd. v. Ent. xh, p. 26, pl.
, ft. 2 (1898), Lombok.
Nye ae ‘a dentifascia, Snellen, |. ¢., p. 24, f. 1, Sumatra.
Leptosoma anthracinum, Voll., Tijd. v. Dierk, i, p. 38 (1863),
Java.
Nyctemera tritoides, Heylarts, Compt. Rend. Soc. Ent. Belg.,
XXXIV, p. xvii (1890), Sumatra.
Leptosoma macklotti, Voll., Tijd. v. Dierk., 1, p. 18 (1868),
=N. latifascia, Hopffer (teste Pag.), Stett. Ent. Zeit.,
1874, p. 45, recorded from Ceram, Flores, Amboina,
and Celebes and is probably some well-known species,
but I am unable to identify it.
Nyctemera menes, Felder, Sitz. Ac.,Wiss. Wien., xlii, p. 38
(1861), Amboina.
Leptosoma agagles, Boisd., Voy. de l Astrolabe Lep., p. 198
(1832), Amboina.
Deilemera uniformis, Plotz, Stett. Ent. Zeit., xl, p. 38 (1880),
W. Africa.
Leptosoma flavescens, Voll., Tijd. v. Dierk., 1, p. 64 (1863),
Sumatra.
Leptosoma leucostigma, Voll., 1. ¢., p. 44, Java.
= Leptosoma nubecula, Voll. (teste Rao.) te. pao:
Nuctemera McKieana, Lucas, Proc. Roy. Soe, Queensland,
x1, p. 60 (1898).
Tne Genus Deilemera.
EXPLANATION OF PLATE III.
Fie. 1. Deilemera kapaurensis, Swinhve.
2. z
3 ’
4. -
5. 9
6. »
Z
(. ”
8, *
uniplaga, Swinhoe.
popiya, Swinhoe.
lutemarginata, Pag.
integra, Walker.
syrmia, Swinhoe,
popiya, Swinhoe (under-side).
albipuncta, Druce.
EXPLANATION OF PLATE IV.
Fia.
. Deilemera gerra, Swinhoe.
1
2?
Z. ”
3. ”
4. ”
v0. ”
6. ”
=
(. ”
8.
nigrovena, Swinhoe.
optata, Swinhoe.
distincta, Swinhoe.
leuctra, Swinhoe.
amosa, Swinhoe.
nisa, Swinhoe.
oroya, Swinhoe,
55
IV. Some notes on the habits of Nanophyes durieui, Lucas,
as observed in Central Spain. By GEORGE CHARLES
CHAMPION, F.Z.S., and Dr. THomas A. CHAPMAN,
M.D., F.Z.S., with a description of the larva and
pupa by Dr. T. A. CHApman, M.D.
[Read Noy. 19th, 1902,]
BEATE a,
DURING a recent visit to Bejar, Central Spain, June 26th—
July 17th, 1902, we noticed numerous large fleshy excre-
scences or galls on the stems of a Coty/edon, a plant growing
abundantly between the crevices of the stone walls in the
outskirts of the town. Many of these galls, on examin-
ation, were found to contain specimens of a Nanophyes,
subsequently identified as V. duriewr, Lucas, in the larval,
pupal,and imaginal condition. The beetles, at the beginning
of July, were mostly immature, and we therefore contented
ourselves by bringing home a supply of the galls, from
which, during August and September, quite two hundred
examples have emerged, accompanied, in September, by a
few Apion sedi, Germ. In the accompanying plate an
illustration is given of the perfect insect, the larva, pupa, and
gall, the beetle only having been previously figured by
Lucas, who, in his description of the species, from a few
specimens found at Oran, Algeria, merely states that the
insect passes its metamorphosis in swellings on the stems
of Cotyledon (Umbilicus) horizontalis. Our observation,
therefore, is not new, still it is interesting as confirming
the statements of MM. Lucas and Durieu, and will no
doubt induce entomologists to search for the insect in
other European localities. . duriewi is recorded from
Southern Spain in Heyden, Reitter and Weise’s “ Catalogus
Coleopterorum Europe ” (1891), but on what authority we
know not, as MM. Brisout and Xambeu, both of whom
have described or noticed the species, simply refer to
Lucas’ work. M. Xambeu (Le Naturaliste, 1901, pp. 224,
225) has recently described the larva and pupa of three
species of the genus: J. lythri, Fabr., on Lythrum salicaria ;
N. telephii, Bedel, on Sedum telephiwum, an insect that. is
TRANS. ENT. SOC, LOND. 1903.—PART I. (APRIL)
88 Mr. G. C. Champion’s Notes on
not unlikely to occur in England, if its food-plant was
searched in various places; and V. hemisphxricum, Oliv.,
on Lythrum hyssopifolium. He also notes (loc. cit.) the
larva of WV. tamarisci, Gyll., as attacking Tamarix, that of
NV. siculus, Boh., on Erica scoparia, and that of V. durieut,
Lue., on Cotyledon (Umlilicus) pendulinus. The habits of
JV. telephii appear to be very similar to those of V. duriewi,
both species making galls on the stems of the plants a little
above the roots. Apion sedi has not, we believe, been
previously recorded as attacking Cotyledon, though it is
known to pass its metamorphoses in the stems of Sedum.
The galls formed by .V. duriewi are very conspicuous, owing
to their large size. They are somewhat kidney -shaped,
several of them being often clustered together, green in
colour, more or less streaked with reddish or purple, and
marked here and there with a minute scar, showing the
original punctures made by the parent insect in the stem
of the plant. The beetle is attacked by a small Chalcid,
of which a number of specimens emerged from the
galls.
The larva of Nanophyes duriewi is a footless maggot very
like many other weevil larve. Its length is, or rather
would be if straightened out, about 3:0 mm., but as it lies
curled up into an arc of about 100 degrees, its actual
measure when full-grown is a little over 200 mm. The
thoracic segments are decidedly thicker than the others,
and here the diameter of the larva approaches 1:00. In the
preserved specimens examined to find them, no spiracles
can be detected, if they exist they are very small and have
no coloured chitinous surroundings. The larva is white or
colourless, except the jaws and some other chitinous portions
in connection with the mouth parts. There is a very
definite ventral prominence of each of the thoracic segments
representing the true legs, but no actual fore-legs exist; at
what may be supposed to be the site of each, is a rather
stronger hair than exists anywhere else on the larva, but
there is nothing to show whether this surviving hair corre-
sponds to one that naturally (7.¢. where the fore-legs are
present) exists at the base of the fore-legs, or is one of those
arming some joint of the fore-leg itself, The marginal
flange is well developed in three definite prominences on
each segment, a lower one that is almost ventral, rather
flat and well delimited, a median one, full and rounded and
almost continuous with the next above, which forms the
the habits of Nanophyes duriew. 89
lower part of the dorsal area, but is marked off by a
longitudinal depression from the portion next above it.
This carries at least on the hinder abdominal segment a
minute colourless bristle. The sulci above this are rather
complicated; they mark off an anterior and posterior subseg-
ment, of which the anterior is dorsally the higher, each
carries a minute bristle, the anterior the larger—one or
two more are present on the prothorax. The interseg-
mental sulcus branches downwards and there is also a
small eminence, at the anterior margin of the segment
immediately above the flange. The 10th abdominal
segment cannot be seen, but is probably present, retracted.
The hairs or bristles made out are very small and very
colourless, and cannot be detected without considerable
search, so that it is possible that others exist; of these
others the only ones supposed to have been observed are
abdominal ones continuing the thoracic pedal series.
The head is comparatively large and well developed,
rounded. Looked at from the front there is a median
suture, losing itself below, without definitely ending in
sutures marking off a clypeus. The head is faintly if at
all tinted, but the clypeal region has some faint fuscous
clouds. The front ends below ina brown chitinous margin,
arched upward centrally over the labrum, and again
laterally into rather more than semicircles round the
antennal origins. In the area projecting between the
Jabrum and antenna, a short bristle arises, and another,
still in the clypeal region, higher up and nearer the middle
line. Marginally (as seen from in front) are three bristles
on each side at about equal intervals, the lower and smaller
behind the antenna, and the highest one about half-way
to the vertex.
The antenne have a very large colourless circular basal
region, bounded by the chitinous semicircle already alluded
to above, a less marked chitinous margin behind, and the
bases of the jaw in front. The antenna itself, arising im
the centre of this area, is very minute and slender, and
deflexed, and appears to consist of a large basal and a
minute terminal joint. The jaws are large and strong,
broad at base, about half that width at their cutting
margin, which consists of two sharp teeth below (as seen
from front), separated from another above by a smooth
hollowed margin.
The maxilla is full and fleshy, has two fine bristles
90) Mr. G. C. Champion’s Wotes on
externally and in front a palpus rising from the middle
of a clear area surrounded by a brown chitinous circle, and
apparently of two joints, below this and more central 1
the labrum formed of two round processes each having
in front a minute palpus, looking (in front view) like two
small concentric circles. Below this the gular region is
swollen and carries two bristles on either side.
The pupa of Nanophyes duriewt has much the appearance
and outline of the larva, is rather shorter and wider, and
is absolutely without any hard, coloured, or chitinous parts.
Whilst the dorsum has much the same curve as in the larva,
the ventral aspect is nearly straight, the (larval) concavity
being filled by the various projections of the appendages.
The head is deflexed, and the rostrum extends downwards
ventrally, to about the-middle (antero-posteriorly) of the
pupa, the anterior end of the pupa being the front margin
of the prothorax. From each side of the rostrum extend out-
wards, first at its base the antenna, then the first femur,
and against it the tibia completely flexed, then similarly
the second femur and tibia, the tarsi (Ast and 2nd) lie
close against the rostrum, the end of the second being
slightly beyond it. The femoro-tibial articulation of these
legs hes outside the elytra cases, which however, with the
wing cases directed backwards, ventrally, and finally in-
wards cover the third legs, except the femoro-tibial articula-
tion which just projects ; dorsally from beneath them. The
wings nearly meet in the middle just beyond the end of
the rostrum.
On the prothorax are a series of bristles, remarkable in
being perched each on the summit of a conical projection,
giving an armed appearance to the front of the pupa.
These appear to be arranged as a row towards the anterior
border and another towards the po terior border of the
segment, two on either side in the front and four in the
back row, but the marginal two of the latter are not so
directly placed in the row as to prevent their position
being otherwise described.
The antennz have a thick basal piece lying agaist the
head or rostrum, from the anterior (position if the head
were extended) end of this the flagellar joints extend
outwards as above noted, to the number of 10 Corsldaye:
each being larger than the preceding. The joints are
somewhat angular, and at about the fourth or filth the angles
appear as definite projections; on the fifth from the
the habits of Nanophyes duriewi. ol
extremity these appear as one or two very definite mammil-
lary projections, and on the three next they form on each a
ring of seven or eight sharp projecting angular points,
with picturesque effect, reminding one strongly of the
similarly studded clubs ‘and chained balls anciently used
as weapons. Very similar conical projections occur on the
tibize, to the number of three or four on each, and a hair
at the exposed extremity of the first and second femora.
Two pairs of conical points are present on the front of the
head (or base of rostrum) basal to the antennal origin.
Some very minute hairs are with difficulty seen on the
abdominal segments. There is a rather larger conical
eminence or two between the projecting elytral bases on
the dorsal centre of the mesothorax.
EXPLANATION OF PLATE V.
Fra. 1. Nanophyes duriewi, larva, x 15.
la. 5 55 », head, from in front.
2. % 5 pupa, x 16.
3. - 5 imago, x 12.
3a. ss 5 hind tarsus.
3b. \ a antenna.
4,5. The galls made by the beetle on the stems of
Cotyledon, natural size. N.B.—These figures
do not show the minute scars that suggest
that each egg is separately laid.
( 93)
V. The Aculeate Hymenoptera of Barrackpore, Bengal.
By GrorGE ALEXANDER JAMES RorTHney, F.E.S.
[Read March 4th, 1903.]
As it is a somewhat uncommon feature for a definite
limited area within the tropics to be worked continuously
for a number of years, I have ventured to hope that a
thoroughly representative list of the Acu/eate Hymenoptera
collected by me inthecantonment of Barrackpore may prove
of some interest to the Society, as this order has of late
become so much more popular, and the Indian and Far
Eastern fauna in particular has attracted an unusual amount
of attention.
My list represents the result of steady collecting from
1872 to 1886, and a flying visit in 1893.
Barrackpore, Bengal, is situated on the east bank of the
Hooghly ; lat. 22° 40° 30”, long. 88° 22’ 45” and 88°
30’. Area, forty-two square miles, and about sixteen miles
north of Calcutta. It is practically contained within a
bend of the river, and includes Monirampore, Pulta, Ischa-
pur Park, Tittaghur, Neilgunge, but above all the beautiful
Barrackpore Park, one of the most charming spots in Lower
Bengal, and one which offers many attractions to the
entomologist.
There are practically three seasons in the year—the
“cold weather” lasting from the end of October to the
beginning of March, the “hot weather” from the middle
of March to the middle of June, when the “rains” set in
and last till October. Very few Hymenoptera (except ants)
are about in the cold season. The hot and rainy seasons
are the best for the collector, and the choicest time of all,
a bright little interval in October, just as the rains are
clearing up, and before the mornings and evenings show
signs of the coming cold weather. The sun then shines
out with beautiful brightness but without the severity of
the hot weather, vegetation is unsoiled by dust, and there
is everywhere a splendid burst and buzz of insect life.
The most attractive spots for collecting are undoubtedly
Barrackpore Park and the jungly ground round the Pulta
powder magazine, for curiously, Ischapur Park and Sam-
nagar (with its extensive earthworks, the ruins of an old
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL)
94 Mr. G, A. J. Rethney on
fort), Neilgunge and Tittaghur, though apparently possess-
ing equally attractive features, never proved as rich in the
number and variety of species, but Monirampore water-
works, with its heaps of white sand, used for filtering
purposes, was always a sure find for the Muti//idz, and the
only spot where they could be looked for with confidence.
The flowers most frequented in the Park are Duranta
plumieri, Quisqualis indica, Poinciana pulcherrima, Mus-
senda macrophylla, Thevetia nereifolia, and a tree with
a small white hanging blossom (probably a variety of
Duranta). At Pulta, a species of Pulicaria which grew in
quantities was very attractive to a variety of species; and
a low jungle shrub with a shiny leaf and greeny-white
flowers, not unlike the privet (but the name* of which
I could never find out), was a still greater favourite, and
especially with the larger species of Hymenoptera. The
Castor-oil plant, Ricinus communis, wasalso much frequented.
The following is a summary of the collection :—
ANALYSIS OF SPECIES.
Species. Percentage. Species. Percentage.
Chrysidide. 13 2-7 13 2°78
Formicide. aT =
Doryline. 3 “64
Ponerine. 15 3°20
Miurmicine. 44 9°40
Dolichoderine. 5 1°07
Camponotine. 25 5°34 92 19°65
Fossores. =3
Mutillide. 54 11°54
Thynnide. 2 “43
Scoliide. 18 3°84
Pompilide. 48 - 10°26
Sphegide. 128 27°35 250 53-42
Diloptera. =
Eumenide. 27 577
Vespidz. 13 2°78 40 8°55
Anthophila. ==
Colletide. 4 "85
Apide. 69 14°75 73 15°60
Total 468
* Since writing this, Col. Bingham has very kindly named this
plant for me at the Museum Herbarium, Glycosmis pentaphylla, Corr.
I estimate that 407 of the species of the Fossores and Apide were
captured at the flowers of this jungle plant.
The Aculeate Hymenoptera of Barrackpore. 95
From this it will be seen that the Fossoves are much
the strongest group, though it is only fair to state that the
number of species of Mutilide, which help to swell the
total, is out of proportion to the specimens, and they have,
too, been much more:closely collected than other families,
for there is always something irresistibly attractive to the
collector about Mutilla.
Tiphia and Myzine, on the other hand, are very poorly
represented, aud both appear to affect northern, or at
least mountain, localities rather than the plains.
The Scoliide, Pompilide, and Sphegide are all well
represented, both in numbers and species, many being
exceedingly common and familiar objects of the bungalow
and compound, as also are several species of Humenide and
Vespide.
The Apidex are the poorest both in species and numbers
in the Barrackpore district (they distinctly strengthen as
you go up-country to the North-West Provinces at ; Allaha-
bad, ‘and still more so at the Mussoorie Hills) ; and with the
exception of Yy/locopa, a few species of Megachile, Crocisa,
and Anthophora, there are no species sutticiently common
to attract general attention; and even with Aylocopa it
is more their great size, noisy booming ways, and the
brilliant effulgence of their wings rather than their actual
numbers which makes them so conspicuous.
In the Formicide, Camponotus, cophylla, Prenolepis,
MNacamma, Solenopsis, Cremastogaster, and Sima are most
in evidence, the three commonest species being Camponotus
compressus, Solenopsis geminata, and Simarufonigra. Pheidole
curiously is not so strongly represented as itis in the Bombay
and Madras Presidencies. Species of other genera are either
local and uncertain in their appearance, or really uncommon
and rare.
I never came across any Jchnewmons in Barrackpore.
No doubt they are to be found if carefully sought for,
fut they must, at least, be exceedingly rare. On the other
hand, in hill-stations like Mussoorie and Darjeeling, they are
fairly common.
Several species of Chrysididz are not uncommon.
I am greatly indebted to Colonel C. T. Bingham, for
his kindness in revising my list of Formicide, and to Mr,
Peter Cameron for revising the Fossores and Apide, ete.
With very few exceptions, the types of all Mr, Cameron’s
Species are in my collection.
96 Mr. G. A. J. Rothney on
CATALOGUE OF SPECIES.*
CHRYSIDID&.
Chrysis, Lin.
Perfecta, Cam. Rare.
Furiosa, Cam. Rare.
Mendicalis, Cam. Rare.
Cotesi, Dubuy. Common.
Disparilis, Cam. Rare.
Oculata, Fab. Common.
Principalis, Sm. Rare.
Oricntalis, Guer. Rare.
Stilbuim, Fab.
Splendidum, Fab, Type, common.
ei var. Ainethystinum, Fab.
Parnopes, Latr.
Viridis, Bé. Rare.
Hedychrum, Latr.
Lugubre, Cam. Rare.
Flammulatum, Dubuysson. Rare.
Family FORMICID.
Subfamily DORYVLINA.
Dorylus orientalis, Westw. f and 9% common.
Anictus pachycerus, Smith. Rare.
Ainictus brevicornis, Mayr. Rare.
Subfamily PONERIN AE.
Anochetus punctiventris, Mayr. Rare; found under
bricks.
Bothroponera tesserinoda, Mayr. Common.
Platythyrea victoriv, Forel. Rare
Diacamma vagans, Smith. Common.
Diacamma sculptum, Jerdon. Rare.
* The number of Aculeata (other than ants) recorded from British
India in Colonel Bingham’s work published in 1897 is 995, but a
considerable number of species have been described since.
The Aculeate Hymenoptera of Barrackpore. 97
Col. Bingham writes:—“I have carefully compared
specimens in your collection with Smith’s type of D.
versicolor, D. versicolor is not an Indian species. The
larger form of Diacamma found in India is, according to
my ideas, J. sew/ptwin.
Brachyponera jerdoni, Forel. Rare.
Ponera confinis, Roger, var. Aitkeni, Forel.
Lobopelta ocellifera, Roger. Rare.
chinensis, Mayr. Common,
minchini, Forel. Rare.
punctiventris, Mayr. Rare.
diminuta, Smith. Rare.
kittelit, Mayr. Rare.
Lioponera longitarsus, Mayr. Rare.
Stigmatomma rothneyi, Forel.= Amblyopone vothneyi.
Rare.
Subfamily J7V RIVICIN AZ,
Cataulacus taprobane, Smith. Rare.
Meranoplus bicolor, Guer. Common.
Triglyphotrix striatidens, Emery. Rare.
Folcomyrmex scabriceps, Mayr. Common,
Myrmicaria brunnea, Saunders. Common.
Col. Bingham writes me :—“ JZ. fodiens, Jerdon = AZ,
brunnea, Saunders; latter has priority. JL. subcarinata,
Smith, is a local variety.”
Tetramorium obeswm, Andre. Rare.
smitht, Mayr. Rare.
simillimwim, Smith. Rare.
denticulatwm, Forel. Rare.
»
In Col. Bingham’s opinion the latter is only a var. of
simillimum.
Monomorium salomonis, Linn. Common.
vastator, Smith = JZ. destructor, Jerdon.
Common.
pharaonis, Linn. Common,
atomus, Forel. Rare.
latinoda, Mayr. Common.
orientale, Mayr. Rare.
3 floricola, Jerdon=M. speculare, Mayr. Col.
Bingham writes :—“Jerdon’s name has priority.” Rare.
TRANS. ENT. SOC. LOND. 1908.—PART I. (APRIL) 7
98 Mr. G, A. J. Rothney on
Cardiocondyla nuda, Mayr. Rare.
Solenopsis geminata, Fabr. Very common. S. armata is
only a var. A very variable species.
Pheidologeton diversus, Jerdon = P. ocellifera, Mayr.
Col. Bingham writes :—“ Jerdon’s name has _ priority.”
Rare.
Pheidologeton afinis, Jerdon =P. laboriosus, Smith, Col.
Bingham writes :—“Jerdon’s name has priority.”, Common,
Pheidole latinoda, Roger. Common.
» vrhombinoda, Mayr. Common.
striativentris, Mayr. Rare.
indica, Mayr. Common.
» parva, Mayr. Rare.
» javana, Mayr. Rare.
», watsoni, Forel. Rare.
», spathifera, Forel. Rare.
» smythiest, Forel, var. bengalensis, Forel. Rare.
Cremastogaster subnuda, Mayr. Common.
e rabula, Forel. Rare.
. rogenhoffert, Mayr. Common.
_ rothneyt, Mayr. Common.
contemta, Mayr. Common.
% aberrans, Forel. Rare.
6 minchint, Forel. in lit. Rare.
Lophomyrmex 4-spinosus, Jerdon. Rare.
Sima rufonigra, Jerdon. Very common.
, nigra, Jerdon. Common.
,, allaborans, Walker = S. compressa, Roger. Col.
Bingham writes:—“ Walker’s name has priority.”
Rare.
Atopomyrmex ceylonicus, Emery, two, §.
Oligomyrmex bengalensis, Forel. Rare.
2 rothneyt, Forel. Rare.
Leptothorax taylori, Forel. Rare.
Subfamily DOLICHODERIN A.
Technomyrmex albipes, Smith. Rare.
Iridomyrmex anceps, Roger = J. excisus, Mayr. Col.
e 9; 5 >
Bingham writes :—* Roger’s name has priority.” Rare.
is) 5
Bothriomyrmex wroughtoni, Forel. Rare.
Tapinoma melanoeephalum, Fabr. Common.
The Aculeate Hymenoptera of Barrackpore, 99
Dolichoderus taprobane, Smith = D. gracilipes, Mayr.
Col. Bingham writes :—‘Smith’s name has priority. JD.
“Je OF . . 9 . ”
gracilipes is only a variety of Smith’s insect.” Common.
Subfamily C4 AZPONOTIN AL.
Camponotus compressus, Fabr. Very common.
mitis, Smith. Col. Bingham writes :—*“ Var.
Juscithorax grades into genuine itis
when you get a big enough series.”
Common,
arrogans, Smith = C. junctus, Forel. Col.
Bingham writes :—“ It stands thus in my
book. I have compared your specimens
with the type.” Rare.
paria, Emery. Common.
sericeus, Fabr. Var. opaceventris, Mayr.
Common.
- taylori, Forel. Rare.
Colobopsis rothneyt, Forel. Rare.
Polyrhachis levissima, Smith. Common.
bicolor, Smith. Rare.
argentea, Mayr.
acasta, Smith. + Rare.
iba, Shatin |)
¥ spunagerd, ad Onion
simplex, Mayr.
‘ thrinax, Roger. Rare.
tubericeps, Forel. Rare.
CEcoph ylla smaragdina, Fabr. Common.
Prenolepis bengalensis, Forel. Common.
longicormis, Latr. Common,
andica, Forel. Rare.
z yerburyt, Forel. Rare.
Plagiolepis longipes, Serdon, Common.
» vothneyt, Forel. Rare.
Acantholepis frauenfeldi, Mayr. Common.
bipartita, Smith. Rare.
capensis, Mayr. Common.
9)
+P]
>
>
100
Mr, G, A. J. Rothney on
Tribe FOSSORES.
Family MUTILLID.
Mutilla sexmaculata, Swed. For # see Cameron, Ann.
and Mag. Nat. Hist., iv (7), 61. Common.
A-carinata, Cam. f Rare.
cona, Cam. & Rare.
interrupta, Lep. Common.
perverst, Cam. ff Rare.
dilecta, Cam. ¢. Rare.
discreta, Cam. 2 Rare.
sabellica, Cam. f Rare.
labiena, Cam. 2 Rare.
Junebrana, Cam. f Rare.
analis, Lep. Rare.
serena, Cam. f Rare.
cleonyma, Cam. 2 Rare.
maculicornis, Cam. f Rare.
poesia, Cam. gf Rare.
phenna,Cam. ff Rare.
idyia, Cam. f Rare.
ludovica, Cam. Rare.
illa, Cam. 2 Rare.
erzia, Cam. f Rare.
lena,Cam. f¢ Rare.
fianna,Cam. 2 Rare.
cara,Cam. @ Rare.
selma, Cam. f¢ Rare.
Jjortinata, Cam. f Rare.
bidens, Cam. f Rare.
wdipus, Cam. &¢ Cf Cameron, Manchr. Mem.,
1898, No. 11, p. 6. One specimen.
interrupta, Oliv. Rare.
Juneraria, Sm. Common.
argentipes, Sm. Rare.
aureo-rubra, Rad. = egregia, Sauss. Rare.
pulcherina, Sm. Rare.
dryta, Cam. -¢ ~ Rare.
compactilis, Cam. 2 Rare.
martialis, Cam. 9 Rare.
pamphia, Cam. 92 Rare.
marcia, Cam. 2 Rare.
The Aculeate Hymenoptera of Barrachpore.
Mutilla gnoma, Cam. 2 Rare.
» emancipata,Cam. 2 Rare.
parthenia, Cam. 2 Rare.
» lethargia,Cam. @ Rare.
» o-maculata,Cam. 2 Rare.
valida, Cam. § Rare.
, leta,Cam. § Rare.
ariel, Cam. 2 Rare.
durga, Bing. = dives, Cam. Common,
peregrina, Cam. & Rare.
cotesii, Cam. YQ Rare.
rothneyt, Cam. § Rare.
redacta, Cam. 2 Rare.
agelia, Cam. § Rare.
mithila, Cam. 2 Rare.
» ocellata, Sauss. Common.
Family THYNNID.
Methoca licolor, Cam. Rare.
orientalis, Smith. Rare.
)
Family SCOLITIDA.
Tiphia brevipennis, Cam. Rare
Plesia (Myzine) dimidiata, Guer. Rare.
bengalensis, Cam. Rare.
» petiolata, Sm. Rare.
Scolia cupitata, Guer. Common.
rubiginosa, Fab. Rare.
humeralis, Sauss. Rare.
redtenbachert, Sauss. Rare.
» aureipennis, Lep. Common.
quadripustulata. Common.
indica, Sauss. Rare.
Elis mai -ginella, Klug. Common.
» thoracica, Fab. "Common.
,» anniulata, Fab. Common.
hirsuta, Sauss. Rare.
brevipennis, Cam. Rare.
Liacos analis, Fab. Rare.
s fulvopicta, Cam. Rare.
>
101
102 Mr. G. A. J. Rothney on
Family POMPILID.
Mr. Cameron informs me that the name Pompilus is
pre-occupied, and that its use cannot be retained in
Hymenoptera. 1 have, however, retained the name here,
and in the sense in which it is used by Col. Bingham.
The family must hereafter be called Ceropalide.
Macromeris violucea, Lep. Common.
Pompilus analis, Fab. Common.
™ zeus, Cam. Rare.
ee ariadne, Cam. Rare.
5 veflecus, 5m. Rare.
7 hecate, Cam. Rare.
" incognitus, Cam. Rare.
a lascivus, Cam. Rare.
a maculipes, Sm. Rare.
. pedalis, Cam. Rare.
“ canifrons, Sm. Rare.
, simillimus, Sm. Rare.
” hero, Cam. Rare.
. rothneyi, Cam. One specimen.
' vivax, Cam. Rare.
A unifasciatus, Sm. Very common.
orientalis, Cam. Rare.
, vagabundus, 5m. Rare.
a incognitus, Cam. Rare.
. vischnu, Cam. Common.
7 cellularis, Cam. Rare.
cf implactibilis, Cam. Rare.
. perturbans, Cam. Rare.
. edilis, Cam. Rare.
Planiceps orientalis, Cam. Rare.
Aporus bengalensis, Cam. Rare.
» cotesii, Cam. Rare.
Pseudagenia deceptriz, Sm. Rare.
a ariel, Cam. Rare.
. pedunculata, Sm. Rare.
cxerulea, Sm. Rare.
blanda, Guer. Rare.
4 Jestinata, Sm. Rare.
z veda, Cam. Rare.
- tincta, Sm. Rare.
- morna, Cam. Rare.
The Aculeate Hymenoptera of Barrackpore. 108
Pseudagenia mutua, Cam. Rare.
Salius flavus, Fab. Very common.
» Jfulvipennis, Fab. Common.
» Otpartitus, Sm. Rare.
» Julgidipennis, Sauss. Rare.
» ~widipennis, Sm. Rare.
» peregrinus, Sm. Rare.
, rothneyi, Cam. One specimen.
, excellus, Cam.=atropus, Sm., 1875, non Smith,
1855. Cf. Cameron, Manr. Mem., 1891,
443. Rare.
,» madraspatanus,Sm. Rare.
» mirandus, Cam. Common.
» electus,Cam. Rare.
Family SPHEGIDA.
Astata agilis, Sm. Rare.
» nigricans, Cam. Rare,
Lyroda argenteofacialis, Cam. Rare.
Piagetia ruficornis, Cam, Rare.
Tachytes monetaria, Sm. A most beautiful species when
alive and glistening in the Indian sun.
Common.
modesta, Sm. Common.
, ornatipes, Cam. Rare.
. rothneyt, Cam. Mr. Cameron writes me that
this is a different species from that named
for him, sinensis, Sm., by the late Mr. F.
Smith. It is rare.
7 hrevipennis, Cam. Rare.
» maculitarsis, Cam. Rare.
» vieina, Cam. Rare.
i tarsata, Sm. Rare.
Tachysphea varihivta, Cam., i litt. Rare.
: argyreus, Sm. Rare.
A: puncticeps, Cam., in litt. Rare.
» + awriceps,Cam. Rare.
< striolata, Cam., in litt. Rare.
Larva simillima, Sm. Common.
» testphone,Sm. Rare.
» » wridipennis,Cam. Rare.
» longicornis, Cam. Rare.
104 Mr. G. A. J. Rothney on
Larva rufipes, Sm. Rare
» erythrogaster, Cam. Rare.
» nigriventris, Cam. Rare.
Leptolarra flavinerva, Cam. Rare.
longitarsis, Cam, Rare.
i reticulata, Cam. Rare.
Spanolarra rufitarsis, Cam. Rare.
Cenolarra appendiculata, Cam. Rare.
Notogenia anthracina, Cam., in litt. Rare.
2)
- picipes, Cam., in litt. Rare.
is Juscistigma, Cam., in litt. Rare.
» pygmexa, Cam., in litt. Rare.
m1 varipilosa, Cam., in litt. Rare.
zs intermedia, Cam., in litt. Rare.
3 fuscipennis, Cam. Rare.
< bengalensis, Cam., in litt. Rare.
a striaticollis, Cam., in litt. Rare.
f pilosa, Cam., in litt. Rare.
” pared, Cam., mn litt. Rare.
fe piliventris, Cam., in litt. Rare.
. indica, Cam., in litt. Rare.
. basalis, Cam. Rare.
op subtesselata, Sm. Common.
. jaculatriz, Sm. Common.
55 erythropoda,Cam,. Rare.
Tiris auratus, Fab. Common. A beautiful species when
seen in the sun.
Gastroscricus rothneyt, Cam. ‘Three specimens.
3 binghami, Cam. One specimen.
Miscophus rothneyi, Bing. One specimen.
Pison suspiciosus, Sia. Rare.
», appendiculatus, Cam. Rare.
» erassicornis, Cam. Rare.
5 rugosus, Sr. Rare.
» orrentalis, Cam. Rare.
Parapison 9 cothneyi, Cam. One specimen.
Anunophila atripes, Sm, Common.
- dimidiata, Sm. Rare.
- nigripes, Sm. Rare.
- punctata, Sm. Rare.
- basalis, Sm. Common.
= erythrocephala, Fab. Rare.
Seeliphron madraspatanum, Fab, Very common.
x coromandelicum, Lep. Common.
The Aculeate Hymenoptera of Barrackpore. 105
Sceliphron violaceum,
Fab, One of the very commonest
species of Barrackpore.
Sphex lobatus, Fab.
» pruosus, Ger. =
Car eae ie
Very common.
= vicinus Bing., var. LRothneyt,
Cameron, Man. Memoirs, 1898, 24.
Rare.
,, luteipennis, Mose. Common.
Cf. Cameron, l.c. Rare.
» wmbrosus, Christ. Common.
aurulentus, Fab. Common.
Ampulex compressa, Fab. Very common.
Rhinopsis ruficornis, Cam. Rare.
Trirhogma cxrulea, Westw. Rare.
Dolichurus clavipes, Cam, One specimen.
Cemonus fuscipennis, Cam. One specimen.
Passaloecus reticulatus, Cam. One specimen.
Nysson rugosus, Cam. Rare.
cinerascens, Dom. =xanthopterus, Cam.
and Bing.
» erythropoda, Cam. Rare.
Alyson annulipes, Cam, Rare.
Midineis orientalis, Cam. One specimen.
Gorytes amatorius, Sm. Rare.
» pretus, Sm.
Rare.
Stizus calopteryx, Handl. Rare.
» blandinus, Sm, Rare.
» Tufescens, Guer. Rare.
» melleus, Sm. Rare.
» vreversus, Sm. Common.
lateralis, Cam. Rare.
Boney lunata, Fab. Rare.
. trepanda, Dbm. Common,
» buddha, Handl. Common.
OT: ientals as, Handl. Rare.
» pinguis, Handl. Rare.
» mdica, Fab.
Rare.
Philanthus depredator, Sm. 'Two specimens.
Cerceris rothneyt, Cam. One specimen.
» orientalis, Sm. Rare.
, humbertiana,
Sauss. Common.
» pretiventris, Dom. Rare.
» flavopicta, Sm. Rare.
» wmstabilis,Sm. Very common,
» pulchra, Cam. Common.
» vischnu, Cam.
Rare.
106 Mr. G, A. J. Rothney on
Cerceris viligans, Sm. Very common.
» btristis, Cam. Common.
» pentadonta,Cam. Rare.
Trypoxylon pileatum, Sm. Rare.
7 canaliculatum, Cam. Rare.
buddha, Cam, Rare.
Fs bicolor, Sm. Rare.
F geniculatwm, Cam. Rare.
- cognatum, Cam. Rare.
i pygmxum, Cam. One specimen.
Oxybelus squamosus, Sm. Bare.
¥ flavipes, Cam. Rare.
- Julvopilosus, Cam. Rare.
Pa canescens, Cam. Rare,
- robustus, Cam. Rare.
Crabro buddha, Cam. Rare.
» orrentalis,Cam. Rare.
, odontophorus, Cam. Rare.
» ardens,Cam. Rare.
, argentatus, Lep. Rare.
» Gellus,Cam. Rare.
» nanus,Cam. Rare.
» nitidus, Cam. Rare.
Tribe DIPLOPTERA.
Family EUMENIDA.
Humenes quadrispinosa, Sauss. Rare.
architectus, Sm. Common.
. punctata, Sauss. Rare.
. buddha, Cam. Rare.
» petiolata, Fab. Very common.
escurtens, Fab. Very common.
is conica, Fab. One of the very commonest
species.
5 flavopicta, Blanch. Common.
* arcuata, Fab. Rare.
. ceylonicus, Sauss. Rare.
Rhynchium hemorrhoidale, Fab. Very common.
* brunneum, Fab. Common.
r abdominale, ig. Common.
A rugulatum, Cam. Rare.
nitidulum, Fab. Common.
argentatum, Fab. Very common.
The Aculeate Hymenoptera of Barrackpore. 107
Rhynchium clypeatum, Cam, Rare.
a)
bengalense, Sauss. Common.
basimacula, Cam. ‘This species is easily
separated from &. flavomarginatum, Sm.
(which does not occur in Bengal), by the
different form of the face and clypeus.
Cf. Cameron, Ann. and Mag. Nat, Hist.,
Dec., 1900, p. 532. Rare.
Odynerus hindostanus, Cam. Rare.
Fs antoni, Cam. Rare.
. sicheli, Sauss. Common.
ie punctum, Fab. Very common.
x difinis, Sauss. Rare.
“ bipustulatus, Sauss. Common.
= ovalis, Sauss. Common.
> sibilans, Cam., in litt. Rare.
Family VESPIDZ.
Learva quadrimaculata, Cam. Rare.
» jserruginea, Fab. Common.
» artifex, Sauss. Rare.
Polistes rothneyi, Cam. One specimen only.
» rufolineatus, Cam. Rare. _
» hebreus, Fab. The commonest
Barrackpore.
» saggitarius, Sauss. Rare.
, stigma, Fab. Very common.
» ngritarsis, Cam. Rare.
Vespa cincta, Fab. Very common.
» affinis, Fab. Common.
» bvasalis, Sin. Rare.
» orrtentalis, Lin. Rare.
Tribe ANTHOPHILA.
Family COLLETIDA.
Prosopis feai, Vach. Rare.
» strenua, Cam. Rare.
» adsoluta, Cam. Rare.
» bellicosa, Cam. Rare.
Family APIDA.
Sphecodes crassiscornis, Sm. Rare.
, propinquus,Sm. Rare.
108 Mr. G. A. J. Rothney on
Halictus itinerans, Cam., in litt. Rare.
ciris, Cam. Rare.
interstitialis, Cam., in litt. Rare.
zonatulus,Cam. Rare.
bengalensis, Cam., in litt. Rare.
Nomia elliotii, 8m. Common.
westur ood, Grib. Rare.
curvipes, Fab. Common.
thoracica, Sm. Common.
clypeata, Sm. Rare.
oxybeloides, Sm. Common.
scutellata,Sm. Rare.
albofimbriata,Cam. Rare.
aureobalteata, Cam. Rare.
argentcobalteata, Cam. Rare.
frederici, Cann. Rare.
lammellata, Cam. Rare.
zebrata, Cam. Rare.
purpureolineata, Cam. Rare.
» latipes,Cam. Rare. -
Steganomus nodicornis, Sm. Common.
Nomada adusta, Sm. Rare.
advena, Sm. Rare.
, ~ ceylonica; Cam. Rare.
Megachile bicolor, Fab. Very common.
lanata, Fab. Very common.
umbripennis, Sm. Rare.
femorata, Sm. Common.
nigricans, Cam. Rare.
disjuneta, Fab. Very common.
ip albifrons, Sm. Rare.
Anthidium rasorum, Sm. Rare.
Parevaspis carbonaria, Sm. Common,
Stelis parvula, Cam. Rare.
» jflavomaculata, Cam. Rare.
Ceratina viridissima, Dall. Common.
_ hieroglyphica, Sm. Common.
Allodape marginata, Sin. Rare.
Celioxzys sexmaculata,Cam. Rare.
argentifrons, Sm. Common.
basalis, Sm. Common.
cuncata, Sm. Common.
confusa, Sm. Rare.
discipiens, Spin. Rare.
3?
>
23
>?
33
2
2:
)
The Aculeate Hymenoptera of Barrackpore. 109
Celioxys fuscipennis, Sm. Rare.
Crocisa emarginata, Lep. Common.
» histrio, Fab. Common.
Anthophora cincta, Lin. Common.
zonata, Lin. Very common.
Jallaz, Sm. Common.
violacea, Lep. Very common.
- bicincta, Fab. Rare.
Xylocopa fenestrata, Fab. Common.
tenuiscapa, Westw. Common.
latipes, Drury. Common.
ewstuans, Lin. Common.
collaris, Lep. Rare.
amethystina, Fab. Rare.
auripennis, Lep. Common.
bryorum, Fab. Rare.
dissimilis, Lep. Rare.
acutipennis, Sm. Rare.
verticalis, Lep. Rare.
Apis indica, Fab. Common.
» jlorea, Fab. Common.
Melipona bengalensis, Cam. Rare.
iridipennis, Sm. Rare
29
33
NOTES ON CONSPICUOUS BARRACKPORE
SPECIES.
FORMICID.
Droylus orientalis, Westw.—Nests in the earth, under
bricks, stones, or in masonry. The {¢ comes into your
bungalows at night, attracted by light, generally at the
end of the cold “weather, February or early March.
Diacamma vagans, Smith—Not uncommon; nests in
walls or other brickwork, under bricks or stones; not
populous; pungent sting; a most intelligent species.
Brachyponera jerdoni, Forel.—Found in some numbers
in December amongst the débris of stacks of wood ; also
in the Botanical Gardens under the same conditions.
Lobopelta diminuta, Smith.—Rare in Barrackpore (but
common in the Botanical Gardens, Calcutta), marches in
long lines two deep.
110 Mr. G. A. J. Rothney on
Meranoplus bicolor, Gaér—Common in the Park; nests
in the earth, particularly the bank and ditch separating it
from the trunk-road. When built in exposed places, the
entrances to the nests are often found covered with the
little pik and blue flowers of some weed. Sexes end of
May.
Holecomyrmex scabriceps, Mayr—The Harvesting Ant of
the Barrackpore district. Common in the Park. Swarms
early in June; nests in roads or hard sun-baked earth, but
always on or near grass land; easily found by the mounds
of seed husks piled up round the entrances. Aunts difficult _
to find after the rains commence.
Myrmicaria brunnea, Saunders.— Nests or colonies found
round the stems of the great banyan tree in the Park by
excavating large fosses, the earth being heaped up in
mounds like regular fortifications. Swarms July 7th-L0th
(Note in the Proceedings Entom. Soc. February 24th,
1892, under the name J/. subear inata, Kmery).
Solenopsis geminata, Fab.; V. armata, Forel—The Red
Ant. Very common, nests in the earth, under bricks or
stones, or almost anywhere. Swarms several times from
March to October.
Pheidologeton laboriosus, Smith = P. affinis, Jerdon.—Not
uncommon in the Park: nests under bricks, stones, flower-
pots, and rock work; occasionally found on the march
changing quarters, when they construct elaborate covered
ways, in which the huge-headed workers-major take an
important and highly-specialized part.
Pheidole rhombinoda, Mayr-—Nests found in the Park
where the grass has been worn bare ; the ants cover these
patches round the entrance to the nests with the leaflets
of a species of mimosa; this covering is arranged in the
form of a circle, and is apparently for protection from the sun.
Sima rufonigra, Jerdon.—Very common ; nests in trees,
particularly fruit-trees like the mango, baél and lychee,
and a species of fig in the Park had its fruit completely
riddled by this ant. Winged ? found in May; is armed
with the most poisonous sting of any Aculeate with which
I am acquainted. Nests frequented by a species of spider,
Salticus; also by Ampulex (Lhinopsis) ruficornis, Cam.,
which very closely mimics the ant, and which I have never
found in any other situation.
Sima nigra, Jerdon.—Not uncommon in trees; is also
mimicked by a Sa/ticus.
The Aculeate Hymenoptera of Barrackpore. 111
Camponotus compressus, Fab.—The Black Ant. Very
common; nests in the earth; swarms in May or early
June.
Polyrhachis levissima, Smith—Nests in the decayed
wood of trees. Swarms June 15th to July 7th.
Polyrhachis thrinaxz, Roger—Nest formed by binding
together one or two leaves with a few silken threads;
very small communities.
Polyrhachis simplex, Mayr.; P. spinigera, Mayr—Nests
formed by web-work binding together a few twigs of a spiny
shrub.
Beophylla smaragdina, Fab.—Common in the Park and
along the trunk-road; nests in trees; sexes in the rains.
The name gives a wrong impression of the colour, the
2, which is seldom seen, alone answering the description ; ;
the workers, which may be seen in thousands, being a
brilliant gamboge yellow.
In « Notes on Indian Ants,” Trans. Entom. Soc. 1889,
IIL, and 1895, II., I have described the habits of many of
the Barrackpore species more fully.
FOSSORES.
Mutilla.—The species of this genus are never found in
any numbers (except perhaps the males of sexzmaculata) ;
single specimens may be picked up anywhere, but the only
spot where you could start out collecting with some
certainty of finding them was Monirampore, where the
heaps of white sand used for the filter-beds of the water-
works (and which in carting was scattered about the paths
and roads leading to the grounds) formed a great attraction
for these sun- and sand-loving insects. The waterworks
are quite of modern date, and it seems véry curious that
imported sand should in a few years have proved an
attraction sufficient to establish a known locality or resort.
How can the slow-moving 2 Mutil/a discover that sand is
to be found within the restricted area (a few acres) of the
Monirampore compound ?
The courting of the sexes of sexmacu/ata may occasionally
be watched on the leaf of a tree or shrub in some secluded
spot, and in its phases is almost ludicrous. The ¢ will
caress, coax, and pet his lady-love for hours together (the
antennz being the chief means of conveying or expressing
t=)
his feelings) ; at last, losing all patience, he will take her
112 Mr. G. A. J. Rothney on
up in his mandibles and shake her, then perhaps feeling
ashamed of himself, will redouble his caresses and begin
the courting over again. I have several times captured
them in their marriage flight.
Scolia.—Common in Barrackpore Park in the rains;
they can often be seen flying along the grass at the edge
of the roads, the beautiful iridescence of their wings
flashing in the sun.
Scolia indica, Sauss., and S. rwhiginosa, Fab.—Found in
the jungle at Pulta. There would come a boom of some
large insect, a flash of black and red, and it had dis-
appeared with a thud in the low thick scrub, from which
it would take much careful beating to rouse and secure.
Elis marginella, Klag.—Was very common about Pulta
in the early rains, preferring shade to sun. I never found
it in the Park.
Sceliphron violaceum, Fab—One of the commonest
species in India, frequenting the verandahs of our
bungalows, and building its little mud-cells on the walls.
Sceliphron madrasapatanum, Fab.—Not so common as
violaceum, but sufficiently so to attract the attention of
any observer.
Sphex lobatus, Fab.—This is one of the most strikingly
handsome of all the Indian Aculeates. It is common
throughout the hot weather and rains, and makes its nests
in the hard ground (being especially fond of bare patches
in the grass land of the Park); it provisions its nests with
a store of field crickets. Before taking a prisoner in, it will
go in and out many times to see that all is secure, and if
during this process you remove the cricket a few yards or .
so from the entrance, it will display great intelligence in
finding it by working round and round in gradually in-
creasing circles till the circumference at last crosses the
cricket, which is always 1n too stupefied a state to escape.
When the females settle on the ground they come with a
clash and a clatter like the clank of cavalry, and in the
brilhant sunlight suggest ideas of fifteenth-century knights
blazing in plate-armour. The males are very fond of the
flowers of the castor-oil plant.
Sphex wmbrosus, Christ; S. awrulentus, Fab.; S. luter-
pennis, Mose.—These three species are fairly common about
the jungly ground round Pulta, particularly so at the close
of the rains.
Ampulex compressa, Fab—This is a common species,
The Aculeate Hymenoptera of Barrackpore. 118
and is very often met with sunning itself on the trunks
of peepul trees. I have found it occasionally in this
situation busily engaged with the workers of Sima rufo-
nigra, picking them up with its mandibles and tossing
them off the tree, but with a sporting air rather than any
serious or deadly purpose, for the ants were never in the
least injured.
Rhinopsis ruficornis, Cam.—A rare species, only found
frequenting the nests of Sima rufonigra, which it very
closely mimics. For some years I had visited a particular
nest of Stima rufonigra in the hope of finding the male,
and at last I was rewarded with what at first sight looked
like an undoubted g. I eagerly captured it; it was not
an ant at all, but something even more interesting, a clever
case of mimicry by a sand-wasp. I also found Lhinopsis
at a second nest, and Mr. Wroughton at a later date, on
discovering Suma rufonigra in the Konkan, found its
understudy in a similar species, Riinopsis constancex, Cam.,
in considerable numbers, a most convincing proof that this
mimicry is genuine, and in no way accidental. The double
mimicry by Lhinopsis and Salticus of a fierce and exception-
ally armed ant appears to me to be specially interesting.
Gastrosericus rothneyt, Cam.; G. binghami, Cam.; Nysson
rugosus, Cam.; NV. erythropoda, Cam.; Gorytes amatorius,
Smith; G. pictus, Smith, were all taken in a bank of fine
earth in a little lane near Pulta, so shaded with dense
jungle that at noon in the hottest weather it was like
twilight and almost cool.
Bembex lunata, Fab,—A rare species. A small colony
found in the sandy bank of a nullah at Pulta.
Bembex trepanda, Dbm.—Common in the rains in the
low jungle about Pulta.
Bembex buddha, Handl.—Found in the hot weather on
sandy patches of ground about Pulta.
Cerceris—Found most commonly during the rains on
flowers. A nullah between Barrackpore racecourse and
Pulta overgrown with jungle, vegetation, and flowers was
especially rich in this genus.
Eumenes conica, Fab.—One of the commonest and most
familiar of Indian Aculeates, building its nests in verandahs
and any other convenient spot, provisioning them with an
abundant store of caterpillars, mostly geometers.
Lhynchiwm basimacula, Cam.—A rare species, but one
that lends a zest to collecting, as you can never be quite
TRANS. ENT. SOC. LOND. 1903.—PART 1. (APRIL) 8
114 Mr. G. A. J. Rothney on
certain when on the wing if one of the common species
may not turn out to be basimacula, with its broad white
band. On one occasion basimacula made its nest in
the turret-staircase leading to the roof of my bungalow.
The cells were very strongly formed and covered with a
particularly sticky kind of cement like shell-lac.
Polistes hebreus, Fab.—The commonest wasp in Barrack-
pore, and generally known as the Yellow wasp. It will
take up its position in a verandah, outhouse, or other
buildizg, and go on piling up its nest on nests year after
year with a persistence which will take no denial. It
varies greatly in size and colour, some specimens being
pinched and starved in appearance and of a dirty grey-
green colour, while others are really fine insects with a
brilliant yellow colouring like fresh gamboge. Season and
environment may have something to do with this varia-
tion. A most striking instance of this was a colony
formed in a little white dome-shaped building which
protected a monument to one of the former worthies of
the Calcutta Botanical Gardens. It stands in one of the
main roads of the gardens, without a particle of shelter or
shade from the blazing sun, and here hebrwus built its
nests year after year, the wasps developing to such a size
and of such an intense yellow as almost to suggest a new
variety.
Vespa cincta, Fab—This species may be taken as the
Bengal hornet; it seldom frequents houses, but is common
enough in trees, shrubs, and old out-buildings. For many
years a large nest was established in the Chirya Khana
(aviary) in the Park. Another, a very curious nest, was
built in a large square terra-cotta flower-pot in the Park.
A tree of some sort had died, leaving a bare stem about
four feet high, and up this the nest was built and added
to year after year. I hada great ambition-to secure this
nest for the British Museum, but the great difficulty
of packing it for sea-transport compelled me to give up
the idea. This species is very fond of frequenting the
date palm when cut by the natives for collecting toddy.
They will settle round the stems in thousands to feast on
the exuding juice. It is curious that at such times the
little grey Indian squirrel (Sciwrus palmarwm) will come
and clear the hornets out with its paws, and take its fill of
the toddy without being molested in any way, and yet
it is not an unknown incident for cincfa to attack even
The Aculeate Hymenoptera of Barrackpore. 115
elephants, as the following cutting from The Hinpress of
India will show :-—
“To walk about Gaur is impossible; there are too many
swamps and jungles; the only way to get about is on
elephants, and even on these one’s progress is sometimes
checked, as happened when we were trying to force our
way through the jungle to the Dakhil Gate. We were in
single file, the Lieutenant-Governor and Mr. Hillow, the
Rajah of Mymensing’s agent, leading, when suddenly orders
were passed back to right-about-face, much shouting took
place, and we found that the whole line of elephants had
been routed by a hornets’ nest, an enemy which is by no
means to be despised.”—Himpress, Aug., 1899 (Sir John
Woodburn’s Tour).
Vespa orientalis, Lin.—Only once have I found this
species in the Barrackpore district. A native sweet-seller
set up astall one day under a peepul tree near the church,
and with it appeared orientalis. After a few weeks sweet-
seller and wasp disappeared and were not seen again.
Although so rare at Barrackpore, this is about the com-
monest wasp up-country, and even at Burdwan, fifty miles
from Barrackpore, it can be found in hundreds in the
native bazar.
APID.
Nomia curvipes, Fab.—Occurred occasionally in the Park,
but more commonly in a grassy lane between the Barrack-
pore Maidan and Pulta. It likes the long “dudh” grass
which grows in damp ditches along hedgerows. At Nis-
chindipore Nuddea it appeared in sufficient numbers to
be quite a feature, and the beautiful gold-brown colouring
and bright golden bands glittering in the sun as the bees
flitted to and fro against the background of emerald
green was an entomological pleasure not readily forgotten.
Steganomus nodicornis, Smith—Somewhat rare in the
Park, its chief habitat being Pulta, where it is common in
the hot weather and rains. Itis a charming little bee and
has a quiet weird little flight of its own, which is very
puzzling till you get accustomed to it. The little white
flowers of a species of Pulicaria are much frequented, ard
it has a habit of settling drawn up in a little compact ball
on the stem beneath the flowér, when it is almost impossible
to discover it. I have been out collecting with a friend
a whole day where tliis bee was fairly common without
116 The Aculeate Hymenoptera of Barrackpore.
his capturing a single specimen until initiated in their
ways.
Nomada adusta, Smith.—You could always find one or
two specimens of this little Nomada flitting about and
settling on blades of grass in the shade of the big trees in
Barrackpore Park, near the Vice-regal kitchens.
Parevaspis carbonaria, Smith.—Common in Barrackpore
Park on the blossom of Duranta plumieri and other
flowering shrubs and trees.
Megachile bicolor, Fab.; MW. lanata, Fab. ; M. disjuncta,
Fab.—Three of the commonest bees in the district, and
may be seen at any time anywhere, busy with their leaf-
cutting operations.
Crovisee emarginata, Lep.; CU. histrio, Fab.—Common in
the Park on the flowers of various shrubs and trees,
particularly the blossom of Duranta plumieri.
Anthophora ciuta, Lin.; A. bicincta, Fab—Common
throughout the hot weather and rains about Pulta,
frequenting the flowers of a plant much like our own
dead-nettle, which is so attractive to our species in
England ; indeed, I came across this plant at Pulta one
day before I had seen the bee. I felt Anthophora should
be near, and sure enough in a few minutes Uicincta and
cincta appeared. My delight was great, for I had not
been long in India and it reminded me of home.
Xylocopa.—Generally common, but more particularly so
in both Barrackpore and Ischapur Parks. They form their
colonies in the half-decayed trunks of trees, and can be
found nearly the whole year round at the flowers of every
tree, shrub, or plant. The yellow blossoms of Thevetia
nereifolia are a constant attraction to them, also Argyretu
nervosa (elephant creeper) and the always favoured
Duranta plumieri.
Themales of Zstwans have a curious habit of frequenting
trees of the china-box which grow in the Park and also
in the deserted gardens of some ruins at Pulta; they circle
round and round the trees high up and well out ‘of reach’ for
hours together, but never settling or resting for a second.
Aphis indica, Fab.; A. florea, Fab.—Common in the
Park and in one’s gardens, but not to the same extent as
our own honey bee is at home. The fine bee, Aphis
dorsata, which is so common “ up-country ” in the North-
West Provinces, Oudh and the Punjaub, I have never
met with in the Barrackpore or Calcutta districts.
VI. Descriptions of nineteen new species of Larride,
Odynerus and Apide from Barrackpore. By
PereR CAMERON. Communicated by GEORGE
ALEXANDER JAMES ROTHNEY, F.E.S.
{Read March 4th, 1903.]
LARRIDES.
VerY little attention has been paid to the smaller
Indian Larride. In Colonel Bingham’s work only one
species of 7 mm. is described. Hence, it is not surprising
that many small species should be undescribed.
Notogonia anthracina, un. s.
Nigra, alis fusco-hyalinis, nervis fuscis, cellula cubitali 1* longiore
quam 2*, metanoto reticulato, 9. Long. 7 mm.
Hab, BARRACKPORE (Ltothney).
Scape of antenne brownish in the middle beneath, shining ; the
front closely punctured and with a shallow, narrow furrow in the
middle ; the vertex alutaceous; the ocellus has a short, narrow,
conical point in front, and is longer than broad, the furrow in front
of it is wide and deep ; the curved furrow on the vertex is wide and
shallow. Pro- and mesonotum closely punctured. Median segment
irregularly reticulated ; the basal two-thirds of the apex irregularly
transversely striated ; the apex with a few longitudinal striz in the
middle ; the furrow is wide and deep and does not reach beyond
the middle. Propleurw irregularly striated ; mesopleure below
the furrow strongly and closely punctured ; above it aciculated.
The sternum between the middle cox is closely longitudinally
striated ; the furrow is deep; the sternal process is stoutly keeled
down the middle and on the sides; the apex with a small incision in
the middle, the sides oblique. The apical abscissa of the radius is
oblique, large ; the first cubital cellule above is shortly, but distinctly,
longer than the second ; the first transverse cubital nervure is angled
from shertly below the middle, where it is bullated; the seeond
recurrent nervure is received shortly behind the middle ; the two
recurrent nervures are separated by not much more than half the
length of the top of the first cubital cellule. Abdomen pruinose ;
the middle ventral segments covered with longish black stiff hair.
TRANS. ENT. SOC. LOND. 1908.—PART I. (APRIL)
118 Mr. P. Cameron on
Notogonia intermedia, n. s.
Long. 7 mm. ?
Hab. BARRACKPORE (Rothney).
Is closely related to V. anthracina, from which it may be
known by the top of the first cubital cellule being longer
compared with the second; by its curved form; by the
apical abscissze being broader and rounder; and by the
recurrent nervures being nearer each other, being separated
by not much more than half the length of the top of the
second cubital cellule.
Scape of antenn brownish in the middle beneath, the lower part
of the front of the face thickly covered with silvery pubescence ; the
front and vertex alutaceons, the latter closely punctured behind the
keel ; the ocellus circular ; a narrow, pointed piece at the apex ; the
furrow in front of it is wide and deep ; the raised part has a shallow,
indistinct furrow in the middle. Palpi testaceous ; pro- and meso-
notum closely and distinctly punctured ; the metanotum irregularly
reticulated at the base, the apex irregularly transversely striated ;
the apical slope has a deep furrow to shortly beyond the middle; on
either side of this it is obscurely striated; the apex has a few
irregular longitudinal striw. The propleure have some irregular
striw ; the mesopleure closely and distinctly punctured below the
furrow ; the metapleure longitudinally striated, closely below, more
widely above. Sternal process keeled down the centre, the keel
widest at the base; the apical incision small, the sides slightly
oblique. The apical abscissa of the radius is rounded below ; the
first cubital cellule is more than one-half longer than the second, the
recurrent nervures are received close together at the apex of the basal
third of the cellule ; legs and abdomen pruinose.
In this species the recurrent nervures are more closely
united than usual.
Notogonia piliventris, n. 8.
Nigra, dense argenteo pilosa ; alis hyalinis, nervis fuscis ; metanoto
reticulato ¢@.
Long. fere 5 mm.
Hab. BARRACKPORE (Ltothney).
Scape of antennze covered with a silvery pile, the middle brownish
beneath, the flagellum opaque. Head opaque, closely and distinctly
punctured, the vertex behind distinctly raised, triangularly incised
in the middle, the edges rounded ; the front ocellus has a minute
Larride, Odynerus and Apide from Parrachkpore. 119
keel in the middle before and behind, the furrow in front of it is
deep, narrow ; the antennal fovee are large, smooth and shining ;
the lower orbits, face and oral region are thickly covered with silvery
pubescence. Mandibles broadly rufous in the middle; the palpi
dark testaceous. Pro- and mesonotum closely punctured, covered
with a pale down; the median segment is closely transversely
reticulated ; the apex is irregularly transversely striated ; and is
thickly covered with silvery hair. Propleura closely and distinctly
punctured ; below marked with enrved stria ; mesopleurwe almost
smooth above the furrow, below closely punctured ; metapleure
closely striated. The mesosternum behind is stoutly keeled round
the edges behind, and has a narrow keel down the middle ; the meta-
sternal process is hollowed, and is slightly roundly incised in the
middle at the apex. Legs pruinose, the spines blackish. Radial
cellule short, wide, the apical abscissa of the radius is oblique, and
makes an angle with the lower one, which has also an oblique slope ;
the first cubital cellule at the top is almost equal in length to the
second ; the second recurrent nervure is received shortly beyond the
middle, the first half-way between it and the first transverse cubital
nervure, which is straight, oblique, and has a large bulla shortly
below the middle. Abdomen pruinose ; the pile on the pygidium
silvery ; the third and following ventral segments are sparsely
covered with long black hair.
Notogonia parva, ua. s.
Long. 5mm. 2.
Hab. BARRACKPORE (Lothney).
In size this species agrees with NV. piliventris, but is
readily separated from it by the alar neuration.
Antenne stout, the keel on the scape fuscous ; the scape is almost
bare. The front and oral region thickly covered with bright silvery
pubescence, the front and vertex are strongly alutaceous ; the furrow
below the ocellus is deep, the ocellus projects triangularly behind ;
the raised portion of the vertex has a narrow furrow down the
middle ; behind, uniting the eyes, is a ——-shaped furrow. Man-
dibles piceous at the apex. Palpi dark testaceous. Pro- and meso-
notum closely and distinetly punctured. The basal portion of the
median segment is transversely striated; the striae distinctly
separated, and at the base they are irregularly joined by longitudi-
nal ones ; the apical slope has a few transverse striw ; the furrow is
of equal width and extends to near the apex. Propleure strongly
aciculated, and is for the greater part striated. Except above the
furrow the mesopleure are distinctly and uniformly punctured,
120 Mr, P. Cameron on
Metapleuree closely and distinctly striated ; near the base is a
perpendicular furrow, divided above the middle by a fovea. The
metasternal process is slightly and gradually widened towards the
apex, which is incised in the middle, but not widely ; there is an
interrupted keel in the middle. The radial cellule is wide ; the
apical abscissa of the radius is rounded below ; the first cubital
cellule is, if anything, longer than the second ; the first transverse
cubital nervure is largely bullated below the middle and _ less
distinctly at the top; the upper part has an oblique slope ; the
second recurrent nervure is received distinetly behind the middle ;
the two are separated by about one-half the length of the top of the
first cubital cellule.
Abdomen pruinose, the ventral surface is covered with long
black hair; the petiole below is closely strongly longitudinally
aciculated.
Characteristic of this species is the shape of the radius, the
apical abscissa of which does not form, on the lower side, a sharp
angle, but is broadly rounded.
Notogonia indica, n. 8.
Long. 5 mmm.
Hab, BARRACKPORE (Lothney).
Comes near to N. parva; may be known from it by the
apical abscissa of the radius not being rounded on the
lower side, by the upper (and larger) part of the first
transverse cubital nervure being more distinctly angled,
by the first cubital cellule being slightly, but distinctly,
shorter than the second; by the striation on the meta-
pleuree being closer; and the first recurrent nervure is
more distinctly roundly curved.
Vertex and front alutaceous ; the lower part of the front and the
oral region densely covered with silvery pubescence. The ocellus is
broader than long, rounded at the sides, and not produced in front
or behind; the part behind it is distinctly raised and furrowed
down the middle, the curved furrow behind this is distinct, and is
produced backwards in the middle, but not very deeply. Palpi
dark testaceous. Pro- and mesonotum closely and minutely
punctured. The base of the median segment reticulated, more
strongly at the base than at the apex ; the apical slope is transversely
striated ; the striw rather widely separated ; the middle is deeply
furrowed. Propleure irregularly striated. Mesopleuree, below the
furrow, strongly and distinctly punctured; metapleure closely
longitudinally and somewhat irregularly striated. Wings hyaline,
Larvridx, Odynerus and Apide fron Darrackpore. 121
the stigma and nervures fuscous ; the apical abscissa of the radius
oblique, straight ; the first cubital cellule is slightly but distinctly
shorter than the second ; the upper two-thirds of the first transverse
cubital nervure has a distinct curve, and is slightly rounded ; the
second recurrent nervure is received near the basal third of the
cellule, the space separating it from the first is less than the length
of the first cubital cellule above. Abdomen shining, the segments
banded with silvery pubescence ; the pile on the pygidinm dense
and silvery.
Notogonia striaticollis, n. s.
Nigra, dense pruinosa ; metanoto reticulato ; metapleuris striolatis ;
alis hyalinis, nervis nigris @.
Long. 7 mm.
Hab. BARRACKPORE (Lothney).
Scape of antenne aciculated, the sides covered with silvery
pubescence ; beneath it is sharply keeled laterally, the flagellum is
covered with a pale pile. Front and vertex closely minutely
punctured ; the front thickly covered with minute silvery pubes-
cence ; the face and clypeus covered thickly with silvery pubes-
cence; the clypeus bears also some long hairs. The ocellus is
sharply triangularly produced on the lower side ; below it is a wide
and deep furrow. Mandibles broadly rufous on the apex. The
eyes at the top are separated by the length of the third antennal
joint. Pronotum alutaceous, behind thickly covered with silvery
pubescence ; the mesonotum is closely and distinctly punctured all
over, and covered with a minute fuscous pile ; the scutellum is
more shining, and is dictinctly punctured, but the punctures are
not so closely pressed together as they are on the mesonotum., The
basal part of the median segment is closely and distinctly reticulated
in the middle, the sides transversely striated ; the apical slope is
closely irregularly transversely striated; the central furrow is
narrow. The basal half of the propleurse is covered with stout
curved strize ; mesopleuree closely and distinctly punctured ; the
metapleure closely obliquely striated. The metasternal process is
rather deep, and is stoutly keeled down the middle. Lees thickly
pruinose ; the tarsal and tibial spines black. The first cubital
cellule above is shortly, but distinctly, longer than the second ; the
first transverse cubital nervure is obliquely sloped at top and
bottom ; both the recurrent nervures are received behind the middle
of the cellule, and are close together. The basal three segments of
the abdomen have broad pruinose bands; the pyvidium is thickly
covered with dark silvery, almost golden, pubescence.
122 Mr. P. Cameron on
Notogonia varipilosa, 0. s.
Nigra, thorace pallide fulvo pilosa, metathorace alutaceo ; alis
tof b) 5) ’
hyalinis, apice fumatis <¢.
e ’
Long. 7 mm.
fab. BARRACKPORE (Lothney).
Antenne stout, longish; the scape densely covered with silvery
pubescence. Front and face densely covered with silvery pubes-
cence ; the vertex alutaceous, the ocellus triangularly produced in
front, rounded behind.
The apices of the mandibles are rufous, the base thickly covered
with silvery pubescence. Palpi dark testaceous. The upper side of
the thorax is thickly covered with dark, somewhat fulvous, pubes-
cence ; the pubescence on the pleure is longer, brighter, and more
silvery in tint. ‘The pubescence is thicker on the median segment,
which is alutaceous, and is only indistinctly striated near the apex ;
its apical furrow is distinct, narrow, of equal width throughout, and
extends from the base to the apex. The sternal process is broad ;
the lateral keels are stout ; there is none in the centre ; the apical
incision is narrow, twice longer than broad, of equal width through-
out, and rounded at the base. Legs thickly pruinose ; the spines
black. Wings hyaline, the apex slightly, but distinctly, smoky ;
the two cubital cellules are equal in length above ; the upper part
of the first transverse cubital nervure has a more sharply oblique
slope than the lower part; the second recurrent nervure is
received, if anything, behind the middle of the cellule. The apices
of the abdominal segments are broadly covered with silvery pile.
Notogonia fuscistigma, n. s.
Nigra, thorace supra dense fulvo pilosa; alis fere hyalinis, stigmate
fusco, nervis nigris, cellula cubitali 2* longiore quam 1* @.
Long. 7 mm.
Hab, BARRACKPORE (Lothney).
Comes near to V. varipilosa, with which it agrees in the
colour of the pubescence, but the latter is stouter built, |
and has the first cubital cellule as long as the second,
whereas in the present species it is perceptibly shorter.
Front and vertex alutaceous ; the ocellus is triangularly produced
in front ; the clypens is distinctly punctured ; the pile on the lower
part of the front has a slight golden tint. Mandibles at the base
Larride, Odynerus and Apide from Barrackpore. 125
thickly covered with silvery pubescence ; from the incision they
are, for the greater part, rufous. The pro-, meso-, and base of meta-
notum are covered densely with fulvous pubescence, the pubescence on
the pleurse and the apex of the metanotum silvery. Pro-, meso-, and
base of metanotum alutaceous; there is a narrow furrow on the
basal two-thirds of the metanotuim ; its apical slope is obscurely
transversely striated and has a narrow deep furrow in the middle
extending to the apex. The mesopleural furrow is distinct ; the
metapleurze closely obscurely obliquely striated. The sternal
process is large; the basal third has an oblique slope and is deeply
furrowed in the middle ; the apical laps are rounded at the apex ;
the incision is short. Legs densely covered with silvery pubescence.
The first transverse cubital nervure is largely bullated at the top
and near the bottom; above the lower bulla it has an oblique
rounded slope ; the first cubital cellule at the top is half the length
of the second ; the two recurrent nervures are separated by the
. length of the second cellule. Abdomen pruinose. Tegulee brown.
N. bengalensis, ere described, comes near to the present
species, but it is a more slenderly-built insect; it may be
known from it by the sternal keel not being depressed at
the base and keeled down the middle; and by the second
recurrent nervure being more sharply angled in the middle,
the curve not being so broadly rounded.
Notogonia bengalensis, n. s.
Nigra, thorace dense, fulvo piloso ; alis hyalinis, stigmate fusco ;
cellula cubitali 22 duplo longiore quam 1* 9.
Long. 6 mm.
Hab. BARRACKPORE (Lothney).
Comes near to N. vavipi/osa, but is smaller, is more
slenderly built, and is readily known from it by the shorter
second cubital cellule, it being with varipilosa equal in
length to the first.
Antenne stout, covered with a pale pile; the scape shining,
keeled on the under-side. Head alutaceous; the lower part of the
front and the oral region thickly covered with silvery pubescence.
Mandibles dark piceous in the middle. Palpi black, thickly covered
with white pubescence. Pro-, meso-, and base of metanotum covered
with fulvous; the apex of the metanotum and the pleure with
bright, silvery pubescence. There is a shallow broad furrow in the
centre of the metanotum; the base of the metanotum enarsely
124 Mr. P. Cameron on
alutaceous, and with a keel in the middle; the apical slope is thickly
covered with silvery pubescence, is obscurely transversely striated,
and has a narrow furrow in the middle which extends to near the
apex. Pleuré alutaceous. Sternal process large, keeled down the
middle, the keel much stronger on the base ; the apex in the middle
incised. Legs thickly pruinose, the spines black. Wings hyaline,
slightly infuscated at the base ; the first cubital cellule is half the
length of the second ; the first transverse cubital nervure is roundly
curved, the second recurrent nervure is received shortly behind the
middle of the cellule. Abdomen pruinose.
Notogonia pygmed, n. s.
Nigra, dense argenteo pilosa, alis hyalinis, apice nervisque: fuscis,
cellula cubitali 1™* duplo longiore quam 2* 9.
Long. 5 mm.
Hab, BARRACKPORE (fothncy).
Seape of antennz bare, brownish in the middle beneath. The
face and the lower part of the front thickly covered with silvery
pubescence ; the ocellus round ; the furrow below it wide and not
very deep; the transverse furrow behind it is wide, shallow, its sides
slightly oblique ; in its middle is a shining longitudinal furrow ;
mandibles broadly rufous at the apex; the base thickly covered with
silvery pubescence. Palpi dark testaceous. Pro- and mesonotum
thickly covered with sericeous pubescence ; that on the former being
brighter in tint than on the latter, which is alutaceous, and has a
broad shallow furrow down the middle. The basal part of the
median segment is alutaceous, neither striated nor reticulated ; the
apex is deeply, but not widely, furrowed down the middle. Pleura:
alutaceous, without any striations. The metasternal process is wide,
is slightly incised in the middle at the apex, and has an obscure keel
down the middle. The first cubital cellule at the top is one-half the
length of the second, the first transverse cubital nervure has a
gradually rounded slope to near the top; the second recurrent
nervure is received shortly behind the middle of the cellule, and is
sharply angled in the middle; the two are separated by slightly
more than the leneth of the top of the first cubital cellule ; the apical
abscissa of the radius is straight, not oblique, and forms an acute
angle with the lower part. Abdomen pruinose, especially on the
apices of the segments, Leys thickly pruinose ; the spines and
ealearia black ; the apices of the anterior tarsi testaceous. .Tegulie
rufo-testaceous. :
Larridx, Odynerus and Apide from Barrackpore. 125
Notogonia picipes, Vr. 8.
Nigra, femoribus posticis piceis ; alis hyalinis, stigmate fusco ?.
Long. 7 mm.
Hab. BARRACKPORE (Lothney).
Comes near to NV. suwhtessellata, but is not much more
than half the length of the smaller examples of the latter ;
the hinder femora, instead of being bright red, are only
piceous-red above and almost black below ; the apex of the
inedian segment is not so widely excavated, and not at all
below the middle ; and the sternal process is not keeled
down the middle, and not obliquely raised towards the
apex.
Mandibles broadly rufous towards the apex. Palpi testaceous.
Front and face thickly covered with silvery pubescence. Ocellus
irregularly round. Thorax alutaceous; covered with a silvery pile,
the middle of the metapleure irregularly striated at the base ; the
apical slope is indistinctly transversely striated, its upper half is
furrowed in the middle ; the sides of the furrow have an oblique
slope. The sternal process is wide, and is not keeled in the middle ;
its apical incision is small. Wings clear hyaline, the stigma fuscous,
the apical abscissa of the radius is rounded below ; the first cubital
cellule at the top is about one-third shorter than the second, the
second recurrent nervure is received shortly, but distinctly, behind
the middle of the cellule. Legs black, pruinose; the hinder femora
for the greater part above reddish-piceous ; the hinder tibia have a
piceous tint; spines and calearia black. Abdomen pruinose, the
apical segment is distinctly and roundly incised.
Notogonia pilosa, 0. s.
Nigra, dense albo pilosa ; alis fusco-hyalinis, stigmate nervisque
testaceis ¢.
Long. 12 mm.
Hab. BARRACKPORE (Leothney).
Front and vertex strongly and closely punctured, thickly covered
with long fuscous hair; the lower orbits, face and clypeus thickly
with longer silvery pubescence ; the projecting apex of the clypeus
smooth, shining and bare. The anterior ocellus is minute, glassy,
slightly pointed in front; in the middle of the front is a wide and
shallow longitudinal furrow ; the depression on the vertex is wide,
deep, transverse, and narrower at the sides. Mandibles black, at the
126 Mr. P. Cameron on
base thickly covered with silvery pubescence, Palpi thickly covered
with silvery pubescence ; the apical joints testaceous. Pro- and
mesonotum strongly and closely punctured and thickly covered with
short white pubescence ; the scutellum is closely punctured like the
mesonotum ; the post-secutellum has the punctures smaller and closer.
Median segment thickly covered with white pubescence, closely
rugosely punctured ; on the apex the punctation is stronger ; its
sides are transversely striated ; the central furrow is distinct in the
middle. Propleure closely punctured, the punctures more closely
pressed together behind ; in the middle is a wide, shallow, oblique
furrow; the tubercles are thickly fringed behind with grey
pubescence ; the perpendicular furrow is crenulated, deep and moder-
ately wide ; the longitudinal one is narrower and not so deep. Meta-
pleure punctured, slightly so at the base ; the lower part.at the base
and middle striated. Pro- and the basal part of the mesosternum
keeled down the middle ; the metasternal process closely punc-
tured and thickly covered with long white hair; the apical lobes
rounded ; coxee and femora thickly covered with white hair ; the
tibiz more thickly and shortly with white pubescence, their spines
pale rufous ; the tarsi thickly pruinose, their spines rufous. Wings
fusco-hyaline, the neryures pallid fuscous; the first cubital cellule
above is slightly shorter than the second, the first transverse cubital
nervure is oblique and bulges out slightly backwards in the middle ;
both the recurrent nervures are received shortly behind the middle
of the cellule; the second recurrent nervure has a broadly rounded
curve. Abdomen thickly pruinose, especially on the apices of the
segments ; the basal ventral segment is rufous.
Comes near to JV, laboriosa and NV. jaculatriz, but is quite
distinct from either.
Tachysphex striolata, uv. s.
Nigra, tegulis testaceis ; alis clare hyalinis ; nervis fuscis ; segmento
mediali striolato 9.
Long. 8 mm.
Hab, BARRACKPORE (Lothney).
Head opaque, closely and distinctly punctured ; the face and sides
of the clypeus thickly covered with silvery pubescence ; the front
and vertex sparsely haired. The deformed hinder ocelli are oblique,
shining, the anterior is broader than long ; the ocellar region is raised
and has a shallow furrow down the middle between the ocelli;
behind them is a deep semi-circular furrow, behind which the vertex
Larridx, Odynerus and Apide from Barrackpore. 127
is raised ; behind the antenne are two distinct oblique shining
tubercles. The apex of the clypeus is shining, smooth, bare ; its
extreme apex depressed. Mandibles broadly rufous in the middle,
their base thickly covered with silvery pubescence. Pro- and meso-
notum closely punctured, the former thickly covered with white
pubescence. Scutellum less strongly punctured than the mesonotum.
Median segment at the base closely obliquely striated ; in the middle
with curved transverse striae; the apex is closely transversely
striated. Propleure shining, aciculated above. Mesopleur closely
punctured ; at the base is a wide perpendicular furrow which is
striated above. The apex of the metanotum is longitudinally closely
striated, the base above is sparsely, below closely punctured. Meta-
sternal area with raised sides ; the central keel is broad and does not
quite reach the base, Legs black, covered with a white down; the
tibial and tarsal spines are white ; the calcaria pale testaceous ; wings
clear hyaline, iridescent ; the stigma and nervures dark fuscous ; the
first cubital cellule above is about one-fourth longer than the second,
the first transverse cubital nervure is oblique and is slightly elbowed
on the lower side; the second recurrent nervure is received shortly
beyond the middle. Abdomen shining, the apices of the segments
pruinose ; the pygidium shining, its apex rufous,
Comes near to 7’. bituberculata, but that species is larger ;
its wings have a distinct yellowish tinge and are fuscous at
the apex, otherwise may easily be known by the median
segment being reticulated ; with the striz stouter and not
oblique.
Tachysphex puncticeps, n. s.
Nigra, capite thoraceque dense punctatis, metanoto reticulato,
apice striato ; alis hyalinis, nervis fuscis ?.
Long. 7 mm.
Hab. BARRACKPORE (Lothucey).
Scape of antennae smooth and shining, the sides bearing a pale
microscopic pile; the flagellum opaque. Vertex closely and dis-
tinctly punctured, the punctures distinetly separated ; behind the
ocelli is a large depression, which is obliquely narrowed behind, the
apex is deeper and shining, the front is closely rugosely punctured.
The antennal tubercles ave smooth and shining, the cheeks, face
and clypeus are thickly covered with silvery pubescence, the cly-
peus is distinctly depressed, smooth, bare and shining. Mandibles
broadly rufous in the middle ; the palpi dark testaceous. The eyes
at the top are separated by slightly more than the length of the third
128 Mr. P. Cameron on
antennal joint. Mesonotum closely and distinctly punctured and
thickly covered with a short pale down. Scutellum shining, only
very minutely punctured. Median segment closely reticulated ; the
apex strongly and closely transversely striated ; the furrow is wide
and deep and does not reach the middle. Propleurze smooth ; meso-
pleure closely punctured ; the metapleure, closely, obliquely striated.
Prosternum furrowed rather widely in the middle. Metasternal pro-
cess not defined. Legs pruinose, the tibial and tarsal spines are
white. Wings clear hyaline, the nervures fuscous, the first cubital
cellule is shortly, but distinctly, longer than the second ; the upper
(and larger) part of the first transverse cubital nervure has a distinct,
oblique slope ; the second recurrent neryure is received in the middle
of the cellule. The basal three segments of the abdomen are fringed
with silvery pubescence,
Tachysphex varihirta, n. s.
Nigra, dense argenteo pilosa, basi metanoti rugoso, apice striolata ;
alis hyalinis, cellula cubitali 1* duplo longiore quam 2*; nervis
stigmateque nigris ¢.
Long. fere 6 mm.
Hab. BARRACKPORE (Rothney).
Front and vertex closely punctured, the front more strongly than
the vertex ; the lower part of the front closely covered with silvery
pubescence, the inner orbits in the middle with pale golden pubes-
cence; the face and clypeus thickly covered with silvery pubes-
cence ; its middle with a slight rounded incision ; the labrum smooth
and shining. Base of mandibles thickly. covered with silvery pubes-
cence, the middle rufous. Palpi dark testaceous. The hinder part of the
head is thickly covered with silvery pubescence. Pro- and mesonotum
covered with silvery pubescence which is very thick and long near the
tegule ; mesonotum closely and distinctly punctured ; the scutellum
has the punctures more widely separated. Median segment closely
rugosely longitudinally rugose ; in the middle, at the base, are some
irregularly waved strie and it is thickly covered with white pubes-
cence ; the apical slope is closely transversely striated. Propleurce
strongly punctured above, below smooth and shining ; mesopleuree
closely, but not very strongly, punctured ; the lower part thickly
covered with silvery pubescence ; metapleure closely striated, the
striz more widely separated at the base. Mesosternum closely punc-
tured, shining ; the metasternal area closely punctured, the apex in-
cised in the middle ; the raised sides stout, the middle indistinctly
keeled. Legs thickly covered with silvery pubescence ; the four
Larride, Odynerus and Apide from Barrackpore. 129
anterior calcaria pale testaceous. The first cubital cellule at the top is
somewhat more than twice the length of the second, the first trans-
verse cubital nervure is oblique, straight ; the second recurrent nervure
is received almost in the middle of the cellule. Abdomen shorter than
the thorax, the basal three segments broadly banded with silvery
pubescence on the apices; pygidium smooth and shining.
Odynerus sibilans, n. Ss.
Niger, flavo maculato ; clypeo flavo, nigro maculato, flagello an-
tennarum subtus rufo ; pedibus flavis, coxis, trochanteribus femori-
busque posticis nigris; alis hyalinis, apice violaceis 9.
Long. 8 mm.
Hab. BARRACKPORE (Lothney).
Antenne black, the scape yellow, the flagellum brownish beneath ;
head black, the eclypeus, except a triangular black mark in the centre
at the apex, the lower inner orbits, the eye incision, an irregular broad
line across the vertex, dilated downwards in the middle and broadly
incised above, and all united together and the upper half of the outer
orbits, yellow. Frontand vertex strongly rugosely punctured, the centre
of the front between the antennz shining and minutely punctured.
Clypeus broad above, rounded there, obliquely narrowed towards the
apex, sparsely punctured, the punctures rather large and deep, the
apex with a broad shallow incision. Mandibles rufous, yellow at the
base, the inner side black. Thorax black, closely and rather strongly
punctured ; the pronotum broadly above, a round spot below the
tegul, scutellum, post-scutellum and the sides of the median segment
broadly yellow. Median segment rugosely punctured, the middle
stoutly keeled and transversely striated ; from the keel near the
top, a transverse, stout, slightly curved keel runs from either side.
Pleure closely punctured, the base of the meso- and metapleure
smooth. Legs yellow, the fore coxe, the others behind, the trochanters
base of anterior femora and the posterior entirely, black. Wings
hyaline, infuscated along the fore margin; the apex violaceous, the
second cubital cellule is much narrower above, being not one-fourth
of the length of the third. Abdomen black, the apex of the petiole, its
sides broadly, a large mark on the sides of the second segment, its apex
broadly, the sides narrowly, the apices of the third and fourth and
the fifth and sixth in the middle, yellow. The ventral segments are
broadly yellow.
There is an indistinct keel, more distinct on the sides
than in the middle, on the petiole behind the yellow mark.
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL) 9
130 Mr. P. Cameron on
APID &.
Halictus itinerans, n. s.
Black, thickly covered with longish white hair; the sides and apex
of the median segment closely and distinctly reticulated; the
apices of the segments depressed and less strongly punctured than
the base ¢.
Long. 7-8 mm.
Hab, BARRACKPORE (LRothney).
Antenne stout, the flagellum bare, the scape covered with longish
white hair. Vertex shining, sparsely and minutely punctured ; the
front is closely, somewhat strongly and uniformly punctured ; its
lower side, the face and the vertex are thickly covered with white
pubescence. Clypeus strongly, but not very closely, punctured, its
apex transverse. Mesonotum distinctly, but not very strongly punc-
tured and with a narrow furrow on either side towards the apex.
Scutellum shining, punctured. The area on the median segment is
large and extends to the top of the apical slope ; it is closely, strongly,
longitudinally, punctured ; the apical slope and the metapleure are
closely reticulated ; the pleurze more closely reticulated, with the
reticulations less clearly defined than on the apex. Mesopleure
rugosely punctured. The apical slope of the median segment is
distinctly margined and is furrowed down the middle. Wings clear
hyaline, the stigma dark fuscous, the nervures black ; the first re-
current nervure is received quite close to the second transverse
cubital. Legs black, rather thickly covered with white hair ; the hair
on the under-side of the tarsi and their spines are rufous; the spurs
are pale rufous. Abdomen closely and distinctly punctured, except
on the apices of the segments which are depressed ; the base of the
dorsal segments are covered with white hair; the ventral surface is
thickly covered with longer white hair. The base of the last ventral
segment is distinctly raised, the raised part forming a semi-circle ; the
apex of the segment is transverse. Tegule black.
Comes near to H. ciris and H. vishnu, but cannot well
be confounded with either.
Halictus interstitialis, n. s.
Black, the apices of the tarsi testaceous, the mesonotum and the
scutellum reticulated closely, the base of the median segment more
widely and less regularly reticulated, the abdomen impunctate, the
base of the second segment with a broad band of grey pubescence ;
the front distinctly keeled 2.
Long. 6 mm,
Larride, Odynerus and Apide from Barrackpore. 131
Hab. BARRACKPORE (Lothney).
Antenne black, the apical half of the flagellum brownish, black
above. The clypeus is strongly, regularly but not very closely,
punctured—the face is less strongly punctured—the front and vertex
are coarsely alutaceous ; the front has a narrow distinct keel which
extends from the ocelli to the base of the antenne ; the front, face and
elypeus are covered with white pubescence ; the vertex more sparsely
with longish pale hair. The reticulation on the mesonotum becomes
stronger towards the apex ; the scutellum is less strongly reticulated
at the sides ; the post-scutellar region is thickly covered with white
hair. The apex of the pronotum is raised laterally at the base into
a sharp plate-like projection, which above is thickly covered with
white pubescence. Mesopleure obscurely reticulated. The base of
the median segment is irregularly reticulated, the longitudinal striz
are regular ; the transverse ones much more irregular and more or less
broken, the sides and the apical slope are thickly covered with white
pubescence, the hair on the sides is much longer than elsewhere.
The hair on the tibie and tarsi are fulvous, on the femora, sparser,
longer and white, the hair is fringed ; the outer spur on the hinder
tibia is armed with stout spines. Wings clear hyaline, the stigma
and nervures testaceous, the first recurrent nervure is interstitial.
Abdomen smooth, shining, impunctate ; the base of the second segment
has a band of white, depressed pubescence at the base ; the other
segments are broadly, but not thickly, fringed, with white pubescence
on the apex, the anal fimbra is rufous, the ventral surface is covered
rather thickly with long white soft hair.
The hinder tibiz and tarsiare distinctly rufous, this being also the
case with the femora beneath ; the tegul are rufo-piceous ; there are
no transverse furrows on the abdominal segments.
Comes near to H. /iodomus, Vachal.
Halictus bengalensis, un. s.
Long. 7 mm.
Hab. BARRACKPORE (Lothiey).
This species comes near to H. ceylonicus, but it wants
the blue tint on the head and thorax; the base of the
median segment is strongly marked with stout twisted
keels, the clypeus is more strongly and distinctly punctured
all over, and there is a distinct transverse furrow on the
basal two abdominal segments.
Antenne black, brownish beneath towards the apex. Front
closely and uniformly punctured, the vertex is less closely and
132 Larridx, Odynerus and Apide from Barrackpore.
distinctly punctured especially near the ocelli. The cheeks are
thickly covered with white pubescence ; the face is sparsely and
minutely punctured ; the clypeus is much more strongly and dis-
tinctly punctured except at the apex. The punctures are large and
elongate, the centre at the apex is narrowly depressed. The labrum
projects slightly in the centre, it is closely and coarsely aciculated
with the sides smooth. Mesonotum shining, closely punctured,
more closely punctured at the sides than in the middle. There is a
distinct basal central furrow which extends beyond the middle and
a shorter one on the sides. Scutellum smooth, the sides and apex
punctured. Post-scutellar region covered closely with white pubes-
cence. The base of the median segment is covered with stout
irregularly twisted striz, which are more oblique laterally than in
the centre. The apical slope is smooth, shining, and is distinctly
bordered round the edges; its top in the centre is slightly bent
downwards with the edges oblique ; in the centre of the apical half
is a deep, clearly defined, furrow. The lower part of the propleurze
is irregularly striated ; below the middle at the base is an oblique
furrow. Mesopleure thickly covered with white hair. Wings
hyaline, the stigma testaceous, the nervures of a darker testaceous
colour ; the second recurrent nervure is interstitial. Legs black,
the hair white ; on the under-side of the tarsi fulvous, the calcaria
testaceous ; the spines on the hinder calcaria are long at the baze
and become gradually shorter towards the apex. Abdomen smooth,
shining and impunctate ; the segments at the base with a band of
white depressed pubescence. The anal fimbra is brownish and
broad ; the segment is brownish and the hair covering on it is long
and fuscous.
The eyes converge slightly below, there is a distinct keel on the
lower part of the front, the metathoracic area is not defined at the
apex by a keel or distinct margin ; there is a distinct transverse
furrow beyond the middle of the basal two abdominal segments ;
the apices of the tarsi are rufous. The pubescent band on the base
of the second segment is broader than it is on the other segments.
Comes near to H. albesens, which may be known from
it by the front not being carinate. It is related to H.
carinifrons, Cam.; that is a smaller, more slenderly-
built insect; the striated area reaches to the top of the
apical slope and the sides at the top are rounded, not
straight and oblique; the basal two segments of the
abdomen, too, want the transverse furrows. The calcaria
are more distinctly spined than usual.
( “33 -*)
VII. Notes on the nests of Bees of the genus Trigona. By
CHARLES OWEN WATERHOUSE, F.E.S.
[Read March 4th, 1903.]
Priate V1.
FRoM time to time portions of the resinous masses formed
by species of Zrigona have been received by our National
Museum, and I believe have been exhibited at meetings
of this Society ; but so far as I know no complete nest has
ever been shown. When my friend Mr. Ridley, of the
Royal Botanic Gardens, Singapore, was last in England, I
asked him to endeavour to procure me one. This is
attended with some difficulty, as the nests are generally
built in hollow trees away in the forest, and usually at a
considerable height from the ground. Soon after his
return to Singapore, Mr. Ridley was fortunate enough to
see a large nest of 7. collina from Malacca being exhibited
at an Agricultural Show, and he very kindly purchased it
and sent it home tome. The bees were still alive in it
when it arrived. As soon as they ceased to come out, I
had the great mass of resin, which weighed 40 lbs., sawn
in half. Roughly speaking the nest is 24 inches long,
and 9 inches wide. The resinous parts, about 8 inches at
the top, and nearly the same at the bottom, have numerous
irregular galleries and cavities. Some of these cavities
are empty, but many of them are filled with pollen. This
was soft and spongy when the nest arrived; now it is
very hard, but the appearance is the same. The central
part of the block is occupied by what one may call the
nest proper. This is about 7 inches by 6. It consists of
innumerable galleries and chambers which are quite
irregular, separated by yellowish waxy partitions that are
not thicker than stout paper. In these partitions are
seen the oval cells containing immature bees.
The cells are 8 millimetres long, and about 4} broad.
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL)
134 Mr. C. O. Waterhouse on
They are placed quite irregularly as to position and
ee Sl ieee, eens
direction, sometimes in groups, sometimes isolated.
(Fig. 1.)
Although the Eastern 7vrigone are numerous and we
have a fair number in the British Museum, all the
specimens are workers, and only workers appear to have
been described. I was very glad to find some males in
the nest described.
Trigona collina, 2.
The male so closely resembles the worker that it might
easily escape observation. The antenne are a trifle longer,
and more slender, with the second joint of the flagellum
very short and strongly transverse. The face is relatively
narrower ; the clypeus with its limits ill-defined, and with
only the front margin obscure ferruginous. The posterior
tibiz are nearly of the same form as in the worker, but
the metatarsus, instead of being very broad and inclined
to be triangular, is parallel and as long as the following
joints of the tarsus taken together. The abdomen instead
of being obtusely acuminate and pubescent at the apex,
is almost truncate and beset with long, stiff, black hairs.
On the under-side it is longitudinally concave, and
generally more or less testaceous. The genitalia are
nearly always more or less visible.
The entrance to the nest of 7. col/ina is by means of a
long resinous tube, that stands out from the trunk of the
tree. The only complete one which has reached me and
which was exhibited at this Society in March 1900, is
14 inches in length, with a somewhat spoon-shaped
prolongation at the entrance.
The Nests of Bees of the genus Trigona. 135
Trigona ruficormis, Sinith.*
When the nest above described was received, Mr. Ridley
also sent a nest of Zrigona ruficorinis, Smith, one of the
smallest bees known. This was in a post in the Botanic
Gardens at Singapore, in a cavity evidently excavated by
Termites. The nest from which the type of this species
was described (Tr. Zool. Soc., vu, p. 185) was in a similar
excavation.
The nest is 8 inches long, in a cylindrical cavity of
about 24 inches diameter. There are various irregular
galleries or means of communication formed of black
waxy substance. In the lower part for about 4 inches
the space is occupied by closely-placed irregular closed
chambers, generally about one quarter of an inch in length,
formed of very thin, brownish wax. (Fig. 2.) These
chambers were nearly all filled with honey; two or three
with pollen. In the upper part of the nest are the brood
cells. These are oval, about 3; mm. long, and nearly
3 mm. broad, formed of brownish-yellow wax, either
isolated or massed together. (Fig. 3.) Many of them are
on a short peduncle. Most of them were filled with
pollen, but many contained immature bees. Those con-
taining the bees were of a lighter, more testaceous colour.
This difference is no doubt due to the pollen having been
consumed, so that they are dry.
Among the bees (which were alive when the nest
arrived) there were males and workers. They are
extremely alike, but nearly all the workers have the
abdomen entirely yellow; a few, however, have the apex
* If the genus Trigona is united with Melipona as is done by
some authors, the name smithii, proposed for it by Col. Bingham
(Fauna Indica) will have to be adopted on account of the older
Melipona ruficornis, Latr.
136 The Nests of Bees of the genus Trigona.
dusky. Possibly this is due to staining. All the males
have it fuscous at the apex, with the base clear yellow.
The apices of the genitalia are nearly always visible. The
form of the posterior legs is almost identical in the male
and worker, but the hairs forming the fringe on the upper
margin of the tibiz in the worker are white and brush-
like at the tip, whereas in the male they are nearly all
simple.
The type specimens of this species, which are from N.W.
India, have the antennz rusty-red. All the males from
the nest above described have them dark-brown, nearly
black. In the workers the scape is clear yellow, and the
flagellum hght-brown,
EXPLANATION OF PLATE VI,
Nest of Trigona collina, about one-fourth natural size.
P, P, chambers filled with pollen.
R, R, solid resin.
© agye
VU A remarkable new Lepidopterous Insect from Zulu-
land. By Sir GeorcE F, Hampson, Bart., B.A.,
F.ZS.
[Read April 1st, 1903.]
Genus APOPROGONES, nov.
Proboscis fully developed ; palpi porrect extending about the
length of head, the 1st and 2nd joints fringed with long hair below,
the 3rd rather long, smoothly scaled ; frons smooth with large tuft
of woolly hair; eyes widely separated on vertex of head, large,
round, naked, overhung by long cilia; ocelli absent ; antenne
simple, dilated into a club before the terminal hook ; vertex of
head and thorax thickly clothed with rough scales mixed with some
hair; pectus with long hair ; tibize smoothly scaled, the mid-tibiz
with one, the hind-tibize with two pairs of moderate spurs ; abdomen
with some rough scales and hair on dorsum especially towards base.
Apoprogones hesperistis, 3 4
Fore-wing broad with the costa arched towards apex which is
rounded, the termen evenly curved ; vein la forming a fork with
1b; le absent ; 2 from two-thirds length of cell; 3 from before
angle ; 5 from above middle of discocellulars ; 6 from upper angle ;
7, 8,9, 10 strongly stalked from just before upper angle, 10 from long
beyond 7; 11 from cell, straight; retinaculum a corneous bar
from subcostal nervure ; frenulum strong. Hind-wing with the
costa moderately arched; the termen excised from below apex to
vein 4, then crenulate, the inner margin long ; the inner area narrow,
vein la not extending to tornus ; le absent ; vein 2 from near angle
of cell; 3,4 from angle ; 5 from above middle of discocellulars ;
6,7 from upper angle ; 8 bent down and touching cell near base,
then widely separated.
TRANS. ENT. SOC. LOND. 1903.—PART I. (APRIL)
138 Sir G. F, Hampson on
Apoprogoncs hesperistis, n. sp.
¢. Head, thorax and abdomen black with a few grey scales and
hairs ; pectus with the hair mostly grey ; tarsi with white rings. Fore-
wing dark leaden grey irrorated and striated with black, forming
very ill-defined antemedial, medial and two postmedial waved lines ;
a small pink spot above base of vein 1; an ill-defined white bar
beyond the cell between veins 6 and 4, and a small triangular post-
medial spot above vein 6 ; a subterminal series of white points in
the interspaces with a small rufous spot before the one above vein 4.
Hind-wing black-brown, the costal area whitish towards base, the
inner area clothed with large rough leaden-blue scales and crossed by
numerous indistinct waved black lines with very ill-defined pink
striz between them towards tornus; an oblique white bar just
beyond the cell and traces of the white postmedial band of under-side ;
cilia of both wings chequered black and white. Under-side of fore-
wing with the basal area pale yellow slightly irrorated with brown ;
a large elliptical black discoidal spot with a white bar beyond it ; a
postmedial white band straight from costa to vein 4, then incurved ;
hind-wing with the basal area pale yellow thickly irrorated with
black-brown ; an elliptical black discoidal spot with oblique white
bar beyond it ; a postmedial white band with waved outer edge, bent
inwards and narrower towards costa ; the inner area whitish to the
postmedial line, interrupted below end of cell and before the
postmedial band.
Hab. ZULULAND, Eshowe 1500 feet, very thickly-wooded
country (Mrs. W. M. Mercer). Exp. 46 mill.
Type in Brit. Mus. A good deal rubbed.
The genusmust be assigned to the family EUSCHEMONID
represented by the single species Huschemon rafilesia,
Westw., from Australia, that however isa typical Hesperid,
except in having a strong frenulum and retinaculum,
whilst the present genus has veins 7, 8, 9, 10 of the fore-
wings stalked, in Luschemon and all HESPERIAD# all the
subcostal ves arising from the cell. -Apoprogones must
therefore be considered a development from the Lepido-
pterous stock at the point where the butterflies branched
off.
In what part of the world the butterflies first arose
we cannot of course say, but all the surviving forms at
all closely related to their point of origin belong to the
scattered remnants of the old Antarctic fauna; we have
first the Castniade found in the Neotropical region and
A remarkable new Lepidopterous Insect. 139
again in Australia, then the Neocustniadx# represented by
Tascina orientalis, Westw., from Singapore, Neocastnia
niceviller, Hmpsn., from Tenasserim, and NV. inetallica, Pag.,
from Palawan, next the Huschemonide represented by
Huschemon rapiesia from Australia, and finally Apoprogones
from Zululand. From this Antarctic fauna probably is
derived the greater part of the Australian fauna, a few
scattered representatives in Christmas Jsland, the Nicobars,
Singapore, the Malay Peninsula, Borneo and Palawan,
some in the Ceylonese subregion and Madagascar, a few
in South Africa, and it seems likely that the very specialized
and characteristic Neotropical fauna is derived from the
same stock.
APRIL 29, 1903.
( 141 )
IX. On Lepidoptera from the White Nile, collected by Myr.
W. L. S. Loat, F.Z.8.; together with further notes
on Seasonal Dimorphism wm Butterflies. By Dr.
FREDERICK A. Dixty, M.A., M.D., Fellow of Wad-
ham College, Oxford.
[Read March 18th, 1903. ]
PAE Vell
Mr. W. L. 8. Loar, during his tenure of office as super-
intendent of the Nile Fish Survey under the Egyptian
Government, spent more than three years in a scientific
investigation of the waters of the Nile and its tributaries.
Though his biological activities were mainly engaged in
other directions, he took several opportunities of collecting
butterflies; and the specimens captured by him in April
1901, and in January and March 1902, are in many
respects of great interest. They are generally in excellent
condition, and are accompanied by ample data; but the
collection (which has been liberally presented by Mr. Loat
to the Hope Museum at Oxford) derives what is perhaps
its greatest value from the fact that the specimens are
marked in such a manner as to allow all those caught at
one time and in one spot to be grouped together. The
captor informs me that on these occasions no voluntary
selection was made on his part; that he tried, in fact, to
catch all he saw; so that each of his“ bags” may be taken
as a fair sample of the butterfly life that was at that
particular time on the wing.
The localities of Mr. Loat’s captures are as follows :—
J. Waite NILE; GHARB-EL-AISH, near KAKA; about
11° N. Lat.
II. WuirE NILE; near KAKA; about 10° 30’ N. Lat.
III. BAuR-EL-GEBEL (Nile); near MANGALA; about 5°
ING Wat:
IV. BAHR-EL-GEBEL (Nile); GonpoKoRO; 4° 43’ N. Lat.
Several of the species obtained from all these localities
are well known to be seasonally dimorphic. As will be
seen later, the present collection contains a preponderance
TRANS. ENT. SOC. LOND. 1903.—PART II. (JUNE) 10
142 oe, Dixey on
of “dry-season” forms. The large proportion of Pierines
to the whole number is remarkable ; as also is the general
resemblance borne by the whole assemblage to the butter-
fly fauna of Aden. It was remarked by Dr. Butler
Proc. Zool. Soc., 1901, p. 25, that the collection made by
Captain Dunn on the Bahr-el-Zeraf (White Nile) had a
very Aden-like aspect, and in particular that it contained
all the forms of Limnas chrysippus, Linn., just as they occur
at Aden. In Somaliland, as is well know n, the prevailing
form of L. chrysippus is L. klugit, Butl., w itha sprinkling of
its modification LZ. dorippus, Klug; in other respects the
present collection recalls the Somaliland fauna almost as
distinctly as it does that of Aden.
It will be neticed that Mr. Loat’s insect-collecting was
limited to a very few occasions, and to only three months
in the year. This will no doubt account for the absence
from his collection of a few species which occur in that of
Captain Dunn (/oc, cit.). Among such are Precis bodpis,
Trim., Adella phalantha, Drury, and especially Hypolimnas
misippus, Linn. There is no new species among Mr, Loat’s
captures, but they include the male of Pinacopterys
venatus, Butl., of which the female only has hitherto been
known to science,
Subjoined is an account of the places and dates of
capture, as carefully recorded by Mr. Loat, together with a
list of the specimens taken on each occasion. It has been
thought worth while to preserve his details, even to the
time of day at which the captures were made.
J. WuireE NILE; GHARB-EL-AISH, near KAKA; about
die IN] Lat:
1901, April 13
A, Between 1 and 2 p.m.
PIERIN &.
TERACOLUS GLYCERA, Butl.
Teracolus glycera, Butl., Proc. Zool. Soc., 1876, p. 144; Aun.
Mag. Nat. Hist., 1897, vol. 11, p. 461; Proc. Zool. Soc.,
L901, p. 25.
424,39. This, as Dr. Butler remarks, is a form of the
T. antigone group. It is barely, if at all, distinguishable
from 7’, evagore, Klug (7. saxeus, Swinh.). See below, p. 161.
Lepidoptera from the White Nile. 145
HESPERIID A.
GEGENES NOSTRODAMUS, Fabr.
1 @; with sharper wings and paler than Mr, Bennett's
Socotran example (also a ?).*
ZL. Between 5 and 5 p.m.
DANAIN A.
LIMNAS CHRySIPPUS, Linn.
L. chrysippus, Linn., 4 23 alcuppoides, Moore, 2 f; alcippus,
Cram., 12; klugit, Butl, 18; dorippus, Klug, 1 &.
These are of the colourmg usual in examples from the
northern districts of the “East African” subregion.
The chrysippus are duller and darker in hue than speci-
mens from Socotra, and the subapical white spots are
less discrete. The specimen of A/ugii is somewhat paler
than the average of that form. It is remarkable that
all five variations of chrysippus are represented in these
nine examples taken at the same time and in the same
place.+
NYMPHALIN &.
BYBLIA ILITHYIA, Drury.
12; intermediate between “wet” and “ dry.”
PRECIS CEBRENE, Trim.
1 9; wet.
PIERIN A.
TERACOLUS GLYCERA, Butl.
(a)
BELENOIS MESENTINA, Cram.
26,0 23 wet.
LYCAININ Al.
CHILADES TROCHILUS, Freyer.
LAS
TARUCUS THEOPHRASTUS, F'abr.
1 2; rather small and dark.
* Proc. Zool. Soc., 1898, p. 382.
+ On the varying forms of Limnas chirysippus, Linn., with their
distribution, see Poulton in Trans. Ent. Soc. Lond., 1902, pp. 473-
482, ibique citata; to which may be added a note by the present
writer in Proc. Zool. Soc., 1900, pp. 10, 11.
144 Dr. F. A. Dixey on
PoLYOMMATUS B&TICUS, Linn.
Ze eeomal
HESPERITD#®
GEGENES NOSTRODAMUS, Fabr.
Le,
April 14.
Between 9 and 10 a.m.
DANAINA.
LIMNAS CHRYSIPPUS, Linn.
L, klugw, Butl,-1 ¢.
NYMPHALIN&.
VANESSA CARDUI, Linn.
lie o..
PIERINE.
TERACOLUS GLYCERA, Butl.
lee Se:
BELENOIS MESENTINA, Crain.
1 f; wet.
LYCAININ A
TARUCUS THEOPHRASTUS, Fabr.
Ia
TARUCUS TELICANUS, Lan
or
1 #3; rather small.
O
POLYOMMATUS B&TICUS, Linn.
1 9.
HESPERIIDA.
GEGENES NOSTRODAMUS, Fabr.
9 ft
ie
At the time of the two days’ collecting above recorded,
the dry weather was breaking up. ‘Thunder was heard in
the distance, and the rains were just about arriving. The
next batch of butterflies was caught at a point on the
Nile a little further south, and a week later in the season.
Lepidoptera from the White Nile. 145
II. Waite NILE; near Kaka; 10° 30’ N. Lat.
1901, April 21.
A. Between 9 and 11 a.m.
DANAINA.
LIMNAS cHRystPpus, Linn.
L. alcippoides, Moore, 2 2; alcippus, Cram., 1 23 hlugii,
Buttle
NYMPHALIN.
PRECIS CEBRENE, Trim.
1 f; wet.
PIERINA.
TERACOLUS GLYCERA, Butl.
lee .
TERACOLUS DATRA, Klue.
3 4,22; all somewhat hghtly marked.
) ) 5 J
TERACOLUS LEO, Butl.
1 f, “dry,” but with no pink shade beneath; 1 9, white,
with very slight orange flush. Inclined towards “ wet.”
TERACOLUS PHISADIA, Godt..
12: with orange flush on upper surface of fore-wing,
and decided reddish shade beneath.
TERACOLUS PROTOMEDIA, Klug.
22
BELENOIS GIDICA, Godt.
Northern form (2. abyssinica, Luc.), 2 2, 3 2. The males
tend to the wet form; one has the wings very sharply
pointed. Of the females, one is wet, one intermediate,
and the remaining one verges towards dry.
BELENOIS MESENTINA, Cram.
lore wel:
LYCANINA.
CHILADES TROCHILUS, Freyer.
oR
POLYOMMATUS B&TICUS, Linn.
29.
146 Dr. F. A. Dixey on
HESPERIIDA.
CHAPRA MATHIAS, Fabr.
pea
PE. Between 3 and 5 p.m.
DANAINA.
LIMNAS CHRYSIPPUS, Linn.
L. chrysippus, Linn. 12; aleippus, Cram., 3 2%; klugit,
Butoh.
PIERIN A.
TERACOLUS PHLEGYAS, Butl.
1f,12 (yellow); both wet.
TERACOLUS EUPOMPE, Klug.
1¢,12. Both of these are “dry” in character, but the
male is without the pinkish shade beneath, and the female
only has it to a moderate extent. Mr. Loat describes the
flight of the male m this and the preceding species as
er wil laeaes
TERACOLUS GLYCERA, Butl.
ike
Oo,» —-4-.
TERACOLUS DAIRA, Klug.
Abe
TERACOLUS LEO, Butl.
1 2; yellow, with orange flush. Under-side pinkish, as in
the dry-season form.
TERACOLUS PLEIONE, Klug.
1; intermediate, verging towards wet.
TERACOLUS PHISADIA, Godt.
5¢, 19. Three of the males are full wet-season
forms, the other two show an infusion of flesh-colour in
the yellow of the under-side. The female is less reddish
beneath than the specimen of the same sex taken earlier
on the same day.
* Of. Mr. G. A. K. Marshall’s observations on the flight of “ purple-
tips.’—Trans. Ent. Soc. Lond., 1902, pp. 354, 371; ‘see also Trimen,
“ South-African Butterflies,” vol. 111, 1889, p. 107.
Lepidoptera from the White Nile. 147
TERACOLUS AMELIA, Luc.
1 9. Dry-season; no basal duskiness on the upper
surface,
TERACOLUS PROTOMEDIA, Klug.
D)
2) 2.
BELENOIS GIDICA, Godt.
Northern form (B, abyssinica, Luc.). 1 2%, wet; 2 2,
dry.
BELENOIS MESENTINA, Cram.
2 ¢,2 9; all wet.
III. BAwr-EL-GEBEL (Nile); near MANGALA; about 5°
N. Lat. More or less open wooded country,
with scrub and dried grass in places; near the
river.
1902, January 8.
Between 3.30 and 5 p.m.
PIERIN &.
TERACOLUS EUPOMPE, Klug.
12%, 12, intermediate; 1 9, dry. All these have more
or less basal duskiness.
TERACOLUS EVARNE, Klug.
aes lediye
TERACOLUS EPIGONE, Feld.
T. microcale, Butl. See Ann. May. Nat. Hist., 1897, i,
p. 472.
32; all dry.
TERACOLUS GLYCERA, Butl.
he ACS,
TERACOLUS DATRA, Klug.
Zee O.
BELENOIS SEVERINA, Cram.
Form boguensis, Feld. 1 2, intermediate.
148 Dr. F, A. Dixey on
January 9.
Between 9 and 11 a.m.
PIERINA.
TERACOLUS EUPOMPE, Klug.
24,19, intermediate ; 19 dry.
TERACOLUS EVARNE, Klug.
192,39; all dry. The dry-season character especially
well-marked in the females.
TERACOLUS EPIGONE, Feld.
te 3 pen alldny,.
TERACOLUS GLYCERA, Butl.
TE gai ee
TERACOLUS DAIRA, Klug.
eels
BELENOIS SEVERINA, Cram.
1%, wet. Form boguensis, Feld. 32,19 (yellow); all
dry.
BELENOIS MESENTINA, Cram.
22,12, wet; 12, intermediate.
ERONIA CLEODORA, Hiibn.
1 g, wet.
LYCANINA.
TARUCUS THEOPHRASTUS, Fabr.
1g, 12. The male corresponds very closely with
specimens in Coll. Hope from Syria. The female, which is
of the blue form, comes very near to 7. sybaris, Hopff.
AZANUS JESOUS, Guér.
1h
LYCENESTHES AMARAH, Gueér.
ia
In a letter to Prof. Poulton, which he kindly permits
me to quote, Mr. Loat describes the present collecting-
ground as follows :—‘“ The country was flat, ground hard
and dry, very little undergrowth, a few large trees, open
bare patches of ground, a good deal of thorny scrub in
Lepidoptera from the White Nile. 149
clumps, and every here and there tufts of dried grass.
The thorny scrub was mostly composed of a low-growing
shrub bearing a small whitish flower with a slight nonde-
script scent; this seemed rather attractive to the whites
and their allies.” The great preponderance of Pierimes in
this two-days collection at Mangala will be noticed.
The remaining captures were all made at Gondokoro or
in its immediate vicinity. With respect to them Mr. Loat
writes as follows:—“The series caught at Gondokoro
[Jan. 12, 16] were collected on a strip of land cleared of
elephant-grass and weeds, and turned into a kind of garden
with sweet potatoes, etc., growing in it, and a few wild
flowers about, close to the edge of the river. The Z.
chrysippus, with hardly an exception, and also the Acraas
were obtained on, or close to, the same strip of ground,
some of the remainder [Jan. 13, March 8] were taken on
ground like that described at Mangala. The weather on
January 8, 9, 12, 13, was dry, sunny and warm. The
rainy season generally begins [at Gondokoro] about the
15th of March, but this year (1902) it commenced about
one month earlier; that is to say, we had occasional
showers and squalls, with long intervals of fine weather.
About the end of March the rains start with a certain
amount of regularity, and last off and on till about October.”
In the neighbourhood of Kaka, 6° further north, the rains
are later. As was stated above, at Gharb-el-Aish, on April
13 (1901), they were only just beginning.
IV. BAHR-EL-GEBEL (Nile); Gonpokoro; 4° 43’ N.
Lat. Strip of ground by the river.
1902, January 12.
Between 3 and 4 p.m.
DANAIN.
LIMNAS CHRYSIPPUS, Linn.
LL, chrysippus, Linn., 4 2 (one with slight white powdering
round gland-patch) ; a/cippoides, Moore, 1 2; aleippus,
Cram., 2 ¢; Alugit, Butl., 2 3; dorippus, Klug, 1 &.
ACRAIN A.
ACRHA VINIDIA, Hewits.
319,19. “Very numerous; takes a long time to die
when put in the killing-bottle.”—W. L. S. L.
150 Dr. F. A. Dixey on
NYMPHALIN&.
NEPTIS AGATHA, Cram.
Ege
PIERIN A.
TERIAS BRIGITTA, Cram.
1 @, wet. The occurrence in January of this well-
marked wet-season form is remarkable.
PAPILIONINA.
PAPILIO PYLADES, Cram.
19. “Rare.”—W. LS. L.
LYCANIN A.
POLYOMMATUS Ba&tIcusS, Linn.
12.
V. GonpdoKoro and neighbourhood. Ground as at
MANGALA.
January 13,
Between 3 and 3.30 p.m.
DANAINA,
LIMNAS CHRYSIPPUS, Linn.
L. chrysippus, Linn., 2¢; klugu, Butl, 1 g and 1 &.
»)
Mostly on the strip of ground by the river.
ACRAINA.
ACREA VINIDIA, Hewits.
3%. On the strip of ground by the river.
PIERIN A.
TERIAS SENEGALENSIS, Boisd.
1 7, dry or intermediate.
TERACOLUS EUPOMPE, Klug.
2 ¢,1 2; all somewhat “dry.
+B)
TERACOLUS GLYCERA, Butl.
have
BELENOIS GIDICA, Godt.
Northern form. 1 §, dry.
Lepidoptera from the White Nile. 151
BELENOIS MESENTINA, Cram.
IL oliae
LYCANIN &.
TARUCUS THEOPHRASTUS, Fabr.
1 Re
Jan. 16.
A. Between 10 and 11 a.m.
DANAIN/.
LIMNAS CHRYSIPPUS, Linn.
L. chrysippus, Linn., 8 2, of which two are small, and
several have a faint white powdering on the hind-
wing ; wleippus, Cram. 1 $; klugit, ButL, 3 2, one
with faint white powdering on hind-wing.
B. Between 11 and 12 a.m.
DANAINZ.
LIMNAS CHRYSIPPUS, Linn.
L. chrysippus, Linn., 3 2,19; aleippus, Cram., 1 2; klagr,
Butl., 1 Zand 1 , the latter with a slight powdering
of white on the hind-wings; dorippus, Klug., 1 %.
ACRAIN A.
ACRA&A ENCEDON, Linn.
1 ¢. This specimen is intermediate between typical
A. encedon and the form A. daira, Godm. & Salv. The
apical area is dusky, but the subapical bar is scarcely
paler than the general ground-colour. On the various
forms of A. encedon, and their relation with the correspond-
ing forms of L. chrysippus, see Poulton and Marshall in
Trans. Ent. Soc. Lond., 1902, pp. 479-484, etc., Plates
XLV, EV):
NYMPHALIN A.
PRECIS CLELIA, Cram.
1 f; rather small, intensely coloured; no trace of ocelli
on upper surface ; under-side looks “ dry.”
NEPTIS AGATHA, Cram.
ae
PIERIN A.
TERIAS BRIGITTA, Cram.
1 2; wet or intermediate.
152 Dr. F. A. Dixey on
TERIAS SENEGALENSIS, Boisd.
2 ¢, dry, one small; 1 2 dry or intermediate, also
small.
TERACOLUS EUPOMPE, Klug.
1 9, rather worn; dry or intermediate.
BELENOIS GIDICA, Godt.
Northern form. 1 ¢, dry or intermediate.
BELENOIS SEVERINA, Cram.
1 g, dry or intermediate.
All the captures on Jan. 16 were made in the “garden
strip,” on a patch of ground 50 yards square.
Jan. 18.
PIERINE.
TERACOLUS PROTOMEDIA, Klug.
1 gf and 1 Q, paired.
March 8.
Between 9.30 and 11.80 a.m.
DANAINA.
LIMNAS CHRYSIPPUS, Linn.
L, klugit, Butl., 1 &.
PIERIN A.
TERACOLUS EUPOMPE, Klug.
5 ,1; intermediate and dry.
Oo
TERACOLUS EVARNE, Klug.
15 ¢,12; all dry, most of them markedly so.
Om
TERACOLUS GLYCERA, Butl.
Dae
LEUCERONIA BUQUETII, Boisd.
12)
PINACOPTERYX VENATUS, But.
1 ¢. This interesting Pinacopleryx is represented in
the National Collection by only two specimens, both
females. The first of these is the type, described and
figured by Dr. Butler in Trans, Ent. Soc. Lond., 1871, p.
Lepidoptera from the White Nile. 153
169; Pl. VII, fig. 7. It was collected on the White Nile
by Petherick. The second is Captain Dunn’s, and was
captured in 1900 on the Bahr-el-Zeraf (Giraffe River,
White Nile). The former is much the more heavily
marked. Mr. Loat’s male corresponds rather with the
second specimen, which has more sharply-pointed and
narrower wings than the type, and probably belongs to
the dry-season phase of the species. The male appears to
be hitherto undescribed.
Pinacopteryx venatus, Butl.—Male.—General aspect somewhat like
that of P. Liliana, Grose Smith ; but smaller, and with fore-wings
narrower and more sharply pointed.
Exp. al. 40 mm.
Upper surface :—Wings white, Fore-wings with inner third of costa
and basal half of cell greyish ; a marginal black point at the
extremity of the third median branch, and marginal black spots at
the extremities of the first and second median, the two radial
branches, and the third subcostal. These spots increase in size from
behind forwards, and those belonging to the first radial and the
subcostal branches become fused, together with a costal spot, into a
dark apical patch, in which however the separate constituents are
still visible. A thin dark costal streak unites the apical black patch
with the basal grey. In the hind-wing, the marginal extremity of
each vein or branch, except the internal, is marked by a small black
spot ; the spots belonging to the second subcostal, radial, and second
and third median, are somewhat linear, being elongated in the
direction of the vein. Lower surface :—Fore-wings white, slightly
greyish along costa and towards base; a small roundish black spot
on lower disco-cellular venule. A fuscous mark, wedge-shaped with
the base uppermost, passes downwards from the second subcostal
near its termination to the space between the first and second radial
veins, crossing the common trunk of the third and fourth subcostals ;
and a large roundish dark spot is situated between the second and
third median branches, about half-way between cell and margin.
All the veins and their branches, except the submedian, are beset
near the margin with a shght powdering of fuscous scales, which at
the extremity of each vein or branch become collected into a more
or less definite spot. Hind-wings creamy white towards base and
costa, elsewhere white like the fore-wings ; costa edged with pale
yellow. A dark oval spot, several times larger than the discoidal
spot on the fore-wing, occupies the anterior half of the lower disco-
cellular vein, the latter forming its long axis. There is a large
fuscous spot on the costa at the termination of the costal vein,
154 Dr, F. A. Dixey on
continued posteriorly by another similar spot in the interspace
between the subcostals. From the posterior extremity of the latter
spot a fuscous band passes across the wing as far as the interspace
between the median and submedian veins, half-way across which it
terminates. This band is of irregular width and runs generally
parallel with the margin of the wing, about half-way between the
cell and border. The veins are more richly powdered with fuscous
seales than in the fore-wing, and the marginal spots. are more
preminent, all but that on the submedian being linear, like most of
those on the upper surface. The fuscous powdering is more strongly
marked on the third median branch than elsewhere, and is continued
inwards along two-thirds of the posterior area of the cell as a definite
dark streak, There is a rudimentary light fuscous spot in the
interspace between the costal vein and the cell, a little internal to
the origin of the first subcostal.
In the Hope Collection, Oxford. This form appears to
be quite distinct from P. simana, Hoptt., and P. Liliana,
Grose Smith. On the whole it most resembles the dry-
season phase of the latter; but the absence of veining on
the upper surface, and the much paler costal margin in
P. venatus f , together with the well-developed apical dark
patch and the dark marginal spots, seem to be distinctive.
BELENOIS SEVERINA, Cram.
cnya
A few moths were included in Mr. Loat’s collection.
They are as follows :—
SYNTOMID.
SYNTOMIS, sp.
LYMANTRIID.
DASYCHIRA ACRISIA, Plotz.
PYRALIDA.
SURATTHA, sp.
The above were all taken by Captain Bell on board a
White Nile steamer between 5° and 6° 30’ N. Lat., Feb.—
March, 1902.
ARCTIIDA.
DEIOPEIA PULCHELLA, Linn.
29; near Kaka, April 21, 1901.
GEOMETRIDA‘.
CZNINA AURIVINA, Butl.
Gondokoro, Jan. 14, 1902.
Lepidoptera from the White Nile. 155
FURTHER NOTES ON SEASONAL DIMORPHISM, SUGGESTED
BY THE ABOVE COLLECTION.
It will have been noticed that Mr. Loat’s specimens fall
roughly into three series; the first (A) consisting of the
butterflies captured near Kaka from April 13 to April 21,
1901; the second (£) comprising those collected at
Mangala and Gondokoro from Jan. 8 to Jan. 18, 1902;
and the third (C) being the final batch from Gondokoro
caught on March 8, 1902. From the accounts that have
been given above of the meteorological conditions prevalent
at these periods in the several localities, we should expect
all three series to show a preponderance of dry-season
forms, though some specimens in series 4 might exhibit
the influence of the early rains. The facts are well in
accordance with this expectation, but it will be seen that
series &, though belonging in point of time to the height
of the dry season, affords examples of the statement that
“in many cases where the existence of seasonal modifica-
tion has been reasonably presumed, or even actually
demonstrated, the seasonal relation is far from being
rigidly fixed.” *
Thus, the two January specimens of Z'erias brigitta are
both wet-season forms; and the same series (/) contains
several wet-season examples of Belenois mesentina and one
of B. severina. But the most curious instance of apparent
seasonal irregularity occurs in the case of Teracolus daira.
All the specimens of this form caught at Mangala on Jan.
8 and 9 are heavily marked on the upper surface, and
would certainly be pronounced at once by most authorities
to belong to the wet season. Those on the other hand
taken near Kaka on April 21, when the rains had well
started, are lightly marked, and bear all the appearance of
dry-season examples. Facts of this kind help to emphasise
the need that exists for still fuller and more accurate data
than we at present possess, if the problems of seasonal
dimorphism are to be satisfactorily unravelled.
Persistence of dry-season coloration. in the females of
seasonally dimorphic species—Myr. G. A. K. Marshall has
lately drawn attention to the fact that in the genus Acrwa
“where the summer males exhibit any particular brilliancy,
as petra, atolmis, or nohara, it is always compensated for
by an exceptional dulness on the part of their respective
* Trans, Ent. Soc. Lond., 1902, p. 193.
156 Dr. F. A. Dixey on
females.” * This dulness of coloration in the wet-season
females mentioned is with some hesitation imterpreted by
Mr. Marshall as being protective in its object. The present
is perhaps a fitting opportunity for pointing out that the case
of these three Acrwas seems to bear some relation to a far-
reaching principle which has met with less notice than it
deserves, and as to the significance of which no suggestion
has hitherto been made. The principle I refer to is
this :—that the dry-season garb of a seasonally dimorphic
butterfly, at least as regards its under-surface, 1s often far
better marked and more persistent in the female than in
the male. This is obviously of interest in view of Professor
Poulton’s interpretation of the cryptic character of dry-
season and desert forms. It would accord with all that
we know as to the special importance attaching to the
life of the female, and the means that are taken for
preserving it, that the more efficient mode of protection,
such as on Professor Poulton’s hypothesis the dry-season
colouring must be, should be more completely and persist-
ently adopted by the sex whose safety from enemies is of
such vital moment to the species. The interest of the
point perhaps justifies a slight digression, and I propose to
give here a few instances which will serve to support the
above ceneralization.
NXanthidia nicippe, Cram. North and Central America.
The wet-season female retains on the under-surface a
tinge of the dry-season purple
Ivias pyrene, Linn, India, etc. Here also the wet-
season female usually retaims the dry-season mottling.
Tvias marianne, Cram. India. The wet-season female
is nearly always brown beneath, as are both sexes in the
dry season. The under-side of the wet-season male is
yellow.
Catopsilia pomona, Fabr. Oriental and Australian
Region. In the wet-season form (C. crocale, Cram.)} the
female often retains in some degree the dry-season
* Trans. Ent. Soc, Lond., 1902, pp. 433, 434. It is hardly neces-
sary to remark that the peculiar need for protection experienced by
the female sex was first pointed out by Mr. Wallace. Some of the
provisions towards this end were recoguized by him as cryptic (as
in many birds) ; others were supposed to be heendapGesinane It
is now known that synaposematism may also play an important part
in the special protection enjoyed by female insects. See Trans, Ent.
Soc. Lond., 1902, pp. 466, 467, ibique cit.
+ Ibid., pp. 431-433, ete. ser lbad; spat OO:
Lepidoptera from the White Nile. 157
ocellation and other characters, though there is an extreme
erocale-form in which they are lost. The “dry-season ”
form C. catilla, Cram., which is probably strongly cryptic,
belongs solely to the female sex.
Pyrisitia proterpia, Fabr. Central and South America.
This is a wet-season form of which there is every reason
to suppose that P. gundlachia, Poey, is the dry-season
modification. The seasonal changes of this species aftord
an interesting parallel with those in the genus Precis
which have lately been so completely dealt with by Mr.
Marshall and Professor Poulton.* The cryptic under-side
of the dry phase is rendered still more leaf-like by the
uncination of the fore-wing and the prolongation of the
hind-wing into a tail-like process, as in Precis archesia and
P. antilope. This applies to both sexes, but is better marked
in the female, as can be seen in the specimens figured (PI.
VII, figs. 1-4). In the wet season both sexes have lost their
leaf-like contour, but the female remains of a duller hue than
the male. A somewhat similar case is that of Zeracolus
auxo, Luc., both sexes of which in the dry-season form (7
topha, Waller., or 7. keiskamma, Trim.) often show a slight
uncination in the fore-wing, though in this species “ tails”
are not developed. The cryptic colouring of the dry-season
under-side is to some extent retained by the wet-season
female (Pl. VII, figs. 5-8).
This latter is a common feature in the African and
Indian genus Teracolus. T. phlegyas, Butl., T. ione, Godt.,
T. regina, Trim., 7. danae, Fabr., 7. eucharis, Fabr., 7. ercs,
Klug, are all cases in point, for in each of them the wet-
season female shows beneath at least a trace of the
characteristic dry-season tinge. Even where this does not
occur, the under-side of the female in the wet season is
usually more cryptic than that of the male, as may be
seen, ¢.g., in 7. omphale, Godt., and 7. achine, Cram.
In Teracolus phisadia, Godt., and 7. puellaris, Butl. (Pl.
VII, figs. 9-12, 13-16) the principle receives perhaps its
highest development. The female of the latter species
retains its cryptic sand-coloured under-side at all seasons,
the under-side of the male in the wet season being bright
yellow. In the former species the female is always, on the
under-side, a cryptic, sand-coloured, ‘“ dry-season” form ;
the male in the dry season may be similarly cryptic, or
* See Trans. Ent. Soc. Lond., 1902, pp. 424 et sequ., P]. XII, XIII.
TRANS, ENT. SOC. LOND. 1903.—PART II. (JUNE) 11
158 Dr. F. A. Dixey on
may possess, as it always does in the wet season, a
bright yellow under-surface, hke that of 7. puellaris.
Simultaneous occurrence of diverse seasonal forms.—
Attention has frequently been called to the fact that at
Aden, and probably in other arid districts, “dry,” “ wet”
and “intermediate” forms may all be found on the wing
together. Colonel Yerbury remarks with reference to
Aden that “seasonal dimorphism does not seem to occur
to any extent in the neighbourhood; though it may
possibly do so in the case of Teracolus calais and dynamene.” *
We may take this to mean, not necessarily that the
different phases usually associated with different times of
ear are never found at Aden (for the occurrence of some
of them at that spot is well attested), but that they do not
there undergo, as in many places, a regular alternation in
correspondence with the change of season. On the excep-
tional case of 7. calais Colonel Yerbury remarks further
as follows :—“The year 1883 was very wet, heavy rain
having fallen in May, consequently in July a large number
of Butterflies appeared—among others, a very brightly-
coloured form of 7’ calais (all, I believe, females however):
this may point to 7. calais ‘being the rainy-season form
and 7’, dynamene the dry. I never ey with this unusually
brightly-coloured form in after years.”
It may be noted in this record that at least a month
must have elapsed between the heavy rain and its supheree
effect on the numbers and aspect of the butterfly fauna ;
this seems to point (like the facts recounted by Poulton
for the genus Precis +) to the larval being the susceptible
stage. ‘On the other hand, the effect of rain may in some
instances be less remote, as appears from another state-
ment by Colonel Yerbury,t as follows :—‘“ Few passen-
gers (for the matter of that, no great number of the
residents) have any idea of the effect on ‘the barren rocks
of Aden’ of a few heavy showers; how almost immediately,
as if by magic, vegetation springs up in every ravine and
watercourse, accompanied by a tolerably abundant insect
fauna.” In the discussion that followed the reading of
the author’s paper on “ Seasonal Dimorphism ” (Proc. Ent.
Soc. Lond., March 19, 1902), Colonel Yerbury further ob-
served that “a temporary rainfall in a dry season in dry
* Proc. Zool. Soc., 1896, p. 257.
+ Trans. Ent. Soc, Lond., 1902, p. 457, etc.
t Journal Bombay Nat. Hist. Soc., vol. vii, 1892, p. 208.
Lepidoptera from the White Nile. 159
places had a marvellous effect in producing intermediate
and wet-season forms.” That the meteorological conditions
prevailing at or about the time of emergence may in some
cases influence the aspect of a brood appears also from
many experiments of Mr, Merrifield, especially those with
Selenia tetralunaria, Hufn., by which it was conclusively
proved that for certain effects of seasonal coloration “ the
later days of the pupal period were especially important.”*
It is worthy of notice that the rule which obtains in Precis,
as to the superiority in size of the dry-season form, is not
of universal application. Mr. Marshall rightly points out +
that Mr. Barker’s statement as to the generally smaller
size of dry-season forms is too sweeping; but there can be
no doubt that im many instances the statement in question
holds good. This is perhaps especially the case among the
Picrinze, concerning the Indian species of which group
Captain Watson says:—“In all genera the dry-season
forms are as a rule smaller than the rainy-season forms.” ¢
In other instances there appears to be no constant ditterence.
The superiority in bulk of the dry-season form in
certain species of Precis, resting as it does upon the result
of a careful series of weighings of the two forms by Pyo-
fessor Poulton, is quite beyond doubt; but it may be well
to remember that in other instances a difference 1n size may
sometimes be more apparent than real. This may possibly
be the case with the broods mentioned by Mr. Merrifield
in Trans. Ent. Soc. Lond., 1892, pp. 40, 41, on which,
together with a similar experience of Weismann’s, he
bases a guarded opinion that both size and shape may
be individually altered during the pupal state. This, is
a point that no doubt calls for further investigation, but
in the meantime it will probably be allowed that, whatever
may be the case with Precis, there is reason to believe
that the seasonal aspect is not in all instances deter-
mined before the assumption of the pupal condition. It
is, as has just been remarked, by no means certain that
the differences in size noticed by Mr. Merrifield were as
real as those in Precis, but, whether they were so or not,
they could not under the circumstances have originated
in the larval state.§
* Trans. Ent. Soc. Lond., 1891, pp. 155-167.
Tt Lbid., 1896, p. 551 ; 1895, p. 413.
{ Journal Bombay Nat. Hist. Soc., vol. viii, 1894, p. 492.
§ In considering the case of Precis it should not be forgotten that
160 Dr. F. A. Dixey on
The foregoing may perhaps help us towards an explan-
ation of the well-attested facts relating to the simultaneous
occurrence of seasonal forms in generally dry localities
like Aden. A feature in Colonel Yerbury’s graphic
description of a temporary rainfall and its effects is the
rapid springing-up of vegetation and the accompanying
increase of insect life, As Professor Poulton has shown,
these are the exact conditions which allow of the assump-
tion of aposematic colouring and habits in exchange for
those of a cryptic character. Now if we assume, as it
seems from Colonel Yerbury’s observations we may, that
many of the Aden species are in a condition to respond
almost immediately to a sudden access of moisture, the
occurrence of the more conspicuous “wet-season’’ con-
temporaneously with the cryptic “dry-season” forms
receives some explanation. Where there is a regular
alternation of long periods of drought and humidity, the
seasonal phases of “the insect fauna fall into a correspond-
ing regularity of succession; but where, as at Aden, a
general state of aridity is liable to be occasionally dis-
turbed by heavy rainfalls of a temporary character, the
intermittent meteorological conditions are apt, we may
suppose, to be reflected in a similar intermixture of apose-
matic and cryptic forms of insect life. It would not be
difficult for residents in such localities to test the
suggestion here offered.*
Note on Teracolus daira and 'T. evagore, Klug.—Dr.
Butler, in his “ Revision of the Genus Zeracolus” (Ann.
Mag. Nat. Hist., 1897), distinguishes Zeracolus yerburit,
the usual rule as to the superiority in weight of the seswmus form
did not obtain in the instance of Mr. Marshall’s P. sesamus and P.
natalensis bred from two eggs laid by the same mother, The weights
as determined by Professor Poulton (Trans. Ent. Soc. Lond., 1902
p. 451) show that in this case the natalensis form was the heavier of
the two. Whatever then may have been the influence which caused
the diversity between the two offspring, it did not find expression
in any increased bulk of the sesamus larva. It should also be borne
in mind that the larval conditions of the first dry- or wet-season
brood will probably differ from those of the second and subsequent
broods (should there be more than one) of the same season,
* See Mr. G. A. K. Marshall’s account of the simultaneous flight
of different seasonal phases during an abnormal season in Mashona-
Jand (Ann. Mag. Nat, Hist., 1901, vol. ii, p. 402), and compare the
discussion of the succession of seasonal phases in Precis by Pro-
fessor Poulton and Mr. Marshall in Trans. Ent. Soc. Lond., 1902, pp.
443-449,
Lepidoptera from the White Nile. 161
Swinh., from 7. daira, Klug. Specimens of 7. yerburii,
however, collected at Lahej, Arabia, by Captain Nurse and
determined by Colonel Swinhoe, who presented them to
the Hope Collection, correspond closely with 7. daira as
figured by Klug. .The locality given by Klug for 7. daira
being “ Arabia felix,” it seems improbable that 7. yerburi
can be more than a synonym for 7. daira, though Mr.
G. A. K. Marshall and Professor Aurivillius agree with Dr.
Butler in keeping them distinct. 7”. evagore, * Klug, which
is no doubt identical with 7. sazews, Swinh., is regarded
by Butler as the dry-season form of 7. yerburii. Whether
7. glycera, Butl., be considered as a species, or as a mere
local race of 7. antigone, there is no doubt that it is barely,
or perhaps not at all, separable from 7. evagore. Hence,
int JA evagore ae the dry- -season form of the Arabian T.
yerburia (i.e. Z. daira), T. glycera should be the dry-season
phase of the s form of 7 daira occur’ ing in the same locality
with itself, viz. on the White Nile. When, however,
Captain Dunn’s collection arrived from the Bahr-el-Zeraf
branch of that river, it was found to contain 7. glycera
in different seasonal phases, all of which were quite distinct
from the specimens of 7’. daira captured in the same
district. In dealing with Captain Dunn’s collection Dr.
Butler * makes no attempt to unite these two forms, and
the examination of Mr, Loat’s specimens convinces me that
he is right in keeping them distinct. But this seems to
carry with it a similar conclusion with regard to the Arabian
TZ’. evagore, which, if the foregoing be correct, cannot be con-
sidered as the dry-season form of “ 7 yerburii” or 7. daira.
The only piece of evidence that seems to make against this
view is the supposed breeding of 7. yerburti and 7’. evagore
by Captain Nurse from similar larvae, Since writing the
note in Trans. Ent. Soc. Lond., 1902, p. 195, I have been
strongly disposed to think that the inference drawn from
Captain Nurse’s account needed confirmation, and I was
accordingly quite prepared to agree with Colonel Yerbury
when I found him urging in Miss Sharpe’s very useful
“Monograph of the Genus Zeracolus” (p. 137) that ‘too
much stress should not be laid on the fact that Captain
Nurse bred a specimen of 7’ evagore from a caterpillar
taken with the larvee of 7. yerburit. ... 7. evagore and
T. yerburw may be seasonal forms [of each other], but
* Proc. Zool. Soc., 1901, pp. 25, 26.
162 Dr. F. A. Dixey on
at present the fact is not proved.” Mr, Loat’s collection
appears to me to bring positive evidence against the view
ot their identity, and from the above considerations I think
there can be little doubt that Mr. G. A. K. Marshall is
right in entirely dissociating 7’. evagore in all its forms
(including 7. nowna, Lue.) from 7. yerburii (i.e. 7. daira).
I am indebted to Professor Poulton, F.R.S., for the
opportunity of working out Mr. Loat’s interesting col-
lection, and also for the Plate which accompanies this
paper.
EXPLANATION OF PLATE VII.
Fic. 1. Pyrisitia proterpia, Fabr., wet-season form, male.
oy tee - . ee i female.
» 93. P. proterpia, dry-season form (P. gundlachia, Poey), male,
sy oF female.
The wet-season 9 (Fig. 2) is less brightly coloured than the
corresponding ¢ (Fig. 1).
In the dry season both sexes adopt cryptic colouring on
the under-side, accompanied by leaf-like shaping of the
fore- and hind-wing, more pronounced in the 2 (Fig. 4)
than in the ¢ (Fig. 3).
Fic. 5, Teracolus auxo, Lucas, wet-season form, male.
59 Ah 5 oe “A 3 female.
» v7. T.auxo, dry-season form (2. topha, Wallgrn, = 7. keiskamma,
Trimen), male,
2 «
yy eh os "5 female.
The wet-season 9 (Fig. 6) retains some of the colouring of
the dry-season form.
Both sexes in the dry season (Figs. 7 and 8) show slight
uncination of the fore-wing.
The specimens represented in Figs, 6 and 8 were bred by
Mr. Mansel Weale, and belong to the series referred to in
Trans. Ent. Soc., Lond., 1902, p. 201.
Fria.
8).
10.
1k,
12,
ales
14,
15.
16.
Lepidoptera from the White Nile. 165
Teracolus phisadia, Godt., wet-season form, male.
im 35 3 5 female.
T’. phisadia, dry-season form, male.
Po * female.
The under-side of the hind-wing in the g (Figs. 9 and 11)
is generally bright yellow, especially in the wet season.
The under-side of the hind-wing in the ? (Figs. 10 and 12)
is sand-coloured at all seasons.
Teracolus puellaris, Butl., wet-season form, male.
Fy 9 . Ar female.
T. puellaris, dry-season form (7. ochreipennis, Butl., = 7.
vorus, Swinh.), male,
<5 ” female,
The under-side of the ¢ is bright yellow in the avet season
(Fig. 13), sand-coloured in the dry (Fig. 15).
The under-side of the @ is sand-coloured at all seasons.
See pp. 157, 158 above. In all the figures, the under-side
alone is represented.
It should be borne in mind that in the absence of colour it
is impossible to represent the tone-values with complete
aceuracy. The difference, ¢.g., between the clear yellow
of Figs. 9, 18, and the sandy tint of Figs. 10, 14, is far
more conspicuous in nature than in the Plate.
X. An Entomological Excursion to Bejar, Central Spain
By GEORGE CHARLES CHAMPION, F.ZS.
[Read March 18th, 1903. ]
In our Transactions for last year (pp. 115-129) I gave a
short account of a journey made by Dr. T. A. Chapmar
and myself in 1901 to Cuenca and other places in Central
Spain, with a list of the principal Coleoptera and Hemi-
ptera met with. In 1902 we made another excursion to
Central Spain, our objective point on this occasion being
the Sierra de Bejar, at the extreme southern extremity of
the large province of Leon, about forty miles south of
Salamanca, The locality was not a new one, entomologic-
ally, Dr. G, Seidlitz having spent a few days in the district
in 1865, at the same time of year, and published some
notes on the species of beetles he met with (cf. Berl. ent.
Zeitschr. 1867, pp. 167-178).* » Leaving London on the
morning of June 21st, we reached San Sebastian the
following afternoon, and after spending a few days at this
attractive seaside resort on the north coast, proceeded on
our way to Bejar, arriving at 8 am. on June 26th, passing
over the dreary plains ‘of Castile and Leon during the
night and early hours of the morning, but little of the
country being visible after daylight, owing to fog. Making
our head-quarters at the Fonda del Nuevo Siglo, we
remained at Bejar till July i7th. From this place we
made numerous excursions, chiefly to the summit of the
Sierra, about 8000 feet, the small omnibus running daily
to Candelario in the early morning and evening helping
us each way, and to Bajos, through the “ puerto,’ or gap
in the range separating Leon from Estremadura, whence
there is rough track back to Bejar over the mountains by
La Garganta. The town of Bejar, which is situate at an
elevation of about 3300 feet, is built on a narrow rocky
ridge just under the mountains sloping down to it on the
southern side, and on the north side the river Cuerpo del
* Dr. Chapman has already published some notes on the bap
doptera observed during our journey (Proce. Ent. Soc. London, 1902,
pp. XXxv-xxxvil ; Ent. ‘Record, 1903, pp. 14-16 et seq.).
TRANS. ENT. $0C, LOND. 1903.—PART II. (JUNE)
166 Mr. G. C. Champion on
Hombre runs in a deep hollow, beyond which the arid
adjacent slopes are mostly occupied by terraced vineyards.
The hillsides were very gay on our arrival with one of the
various species of “broom” characteristic of the district,
viz. Genista florida, this being in full flower at the end of
June. On the higher parts of the Sierra this plant was
replaced by Cytisus purgans, the dense masses of its yellow
flowers being visible from a long distance. On the slopes
near the town, the vegetation chiefly consisted of chestnut
and small deciduous oaks, of which latter there were woods
atan elevation of quite 5500 feet, near La Garganta, as well
as low down on the other side of the river near San Gil;
and amongst the broom there was a great deal of arbores-
cent heath, lavender, and other aromatic plants, etc. The
whole of the Bejar region visited seemed to be of granite
formation, and no pines or other Coniferse were seen.
From the summit of the Sierra de Bejar, on which there
were large patches of snow up to the beginning of July, a
magnificent view was obtained of the ‘higher Sierra de
Gred dos, with its numerous jagged, snowy peaks, this
range extending far to the eastward towards the Guadar-
rama, becoming gradually lower in that direction; the
Sierra de Gata,* and some of the mountains of Portugal,
none of these appearing to be much over 6000 feet; the
plains of Leon and Castile, and a portion of those of
Estremadura, ete. The abundance and variety of “ brooms”
and the vast number of insects to be found on them when
in flower, especially on Genista florida, was to us the most
interesting feature of the district, nothing of the kind
having been noticed by us at Cuenca, ete., the previous
year. On the higher parts of the Sierra many beetles
were to be found, such as Asida castellana, Dorcadion
dejeant (the only species of the genus met with during
the entire trip), Corymbites of several species, including
C. hemapterus, a very elongate Athous, Otirrhynchus
dentipes, two Nebrivw, Cymindis of several species, Amara
ooptera, Trechus pandeller, a Bembidium of the B. glaciale-
eroup, Pierostichus azare, four species of Zabrus, cluding
Z. constrictus, Byrrhus depilis, Olocrates, Heliopates, Ten-
tyria, Timarcha, etc., many of them in plenty. A little
lower down Chienius dives ran about on the grassy slopes,
* Dr. Seidlitz visited Alberca and the Pena de la Francia in this
district in 1865. - We had intended also to go there, but deferred the —
journey till too late.
An Entomological Exewrsion to Bejar, Central Spain, 167
or among the stones, often with Dorcadion dejeani ; and
Platycerus spinifer was taken on the wing amongst Cytisus
purgans, this plant not ascending much over 7000 feet.
On its flowers, or on those of Genista horrida, a Zonabris
occurred in abundance, with Lytta vesicatoria, Cryptoce-
phalus koyi, Gynandroph halma amabilis, and a striped
Rhagonycha. On the grass-stems thereabouts Barypithes
sulcifrons, Polydrusus affinis, a Dasytes, a large Henicopus
—the ¢ with long black hair, and the @ with greyish-
white hair—Meloe mayalis, etc., were met with. Aphodius
bonvouloiri, too, was very common at about this elevation.
The tracks made by the goats amongst the dense growth
of Cytisus seemed to be the favourite haunts of Asida, Ten-
tyria, Olocrates, various Coprophaga, Histeridee, etc., as many
of the specimens captured had evidently been trodden
upon and injured by these animals. On the lower sum-
mits, especially in the walled-in enclosures where the
goats are driven in to rest, lots of common Coprophaga
were noticed—Aleuchus, Gymnopleurus, Sisyphus, Geotrupes,
etc. as well as Scawrus, Pimelia, and other Tenebrionids,
Histeridee, etc. In some of the open grassy places in or
near these enclosures the males of a Chasmatopterus flew
in the greatest profusion in the sunshine, close to the
ground, looking like bees on the wing, and they could be
caught in any quantity with a butterfly-net, paired speci-
mens frequenting flowers later in the day. In an oak
wood above La Garganta, in which there were many
“brooms,” Cardiophorus signatus and other members of
the genus, Lebia trimaculata, Strophosomus erinaceus,
various Cryptocephali, Clythra, Agrilus biguttatus, Melano-
tus, Athous, Brachyderes, Polydrusus, Phyllobius, and other
species were taken, some of them in plenty. On the
hillsides near the town of Bejar the Genista florida, while
in flower, produced a great variety of beetles, as Luperus
nigrofasciatus (in the greatest profusion), Cneorrhinus, sp.
(an abundance, but all females), Sciaphilus carinula, Apion
argentatum and others of the genus, Sitones, Polydrusus,
Melanotus, Cardiophorus, Coryimbites, Rhagonycha, Dasytes,
Haplocnemus, Danacea, Malachius, Malthinus, Ebeus,
Amauronia, Helops, Agrilus, Corebus, Cryptocephalus,
Zonabris, Lebia, etc. ; Hlater preustus, var. aurilegulus (an
plenty), a large Lachnea, various Cryptocephali, Athous,
etc, occurred on oak; a brassy Ha/tica on heath, ete.
In the open places on these slopes Corymbites gougelete
168 Mr. G. C. Champion on
was frequently taken on the wing, or found on grass-
stems; and Cerocoma schrebert was locally common on
flowers. In an oak wood on the dry slopes near San Gil,
in which there were many small flowering plants, a number
of interesting insects were captured, especially towards
evening, as Corebus fasciatus (rarely, on oak, and also
found in an ants’ nest under a stone), Anthaxia hungarica
and others of the genus, Serica mutata, Lagria rubida Gn
abundance, but apparently all males), a fine Pachytychius
(P. sparsutus) on Cytisus welwitschi, P. scabricollis, Smi-
cronyx, Orchestes, Cryptocephalus globicollis, etc., Pachy-
brachys, various Clythrids, ete. On the slopes between
this wood and the river a species of the remarkable
Neuropterous genus Nemoptera flew freely towards even-
ing. The river itself, or a small affluent of it, produced
many Hydradephaga, as Bidessus minutissimus, Deronectes
cartnatus, D. bicostatus, Hydroporus flavipes, H. bilineatus,
HH. lepidus, Agabus heydeni, A. brunneus, A, didymus, A.
paludosus, ete, also Gyrinus urinator, Orectochilus, and
various Hydrophilide. On the walls of the town, as
already recorded,* we noticed various galls on the stems
of a species of Cotyledon, a plant growing very commonly
in such situations, and these were found to contain speci-
mens of Nanophyes durieut in all its stages, a species not
mentioned by Dr. Seidlitz in his list of Bejar insects, and
originally described from Algeria.
Leaving Bejar on July 17th, we went on by the dili-
gence to Piedrahita, about five hours’ journey, in the heat
of the day, where we remained till the 28rd, our road
beyond the “puerto” being fairly level and extending
through a somewhat arid country, where the ground was
everywhere cultivated, even between the very old scattered
olive-like, evergreen oaks ( Quercus gramuntia) to be
seen In many places, the river Tormes being crossed by
an ancient bridge at about half-way. At Piedrahita the
best collecting-ground was the slope and summit of the
adjacent Sierra, about 5500 feet elevation, which runs
parallel with the Gredos, and is separated from it by large
elevated tracts and valleys, affording extensive pasturage
for cattle in the summer months. Cyfisus purgans grew
im abundance in these places (the old stems apparently
being in much demand for fuel), extending down the
slopes for about five hundred feet. Below this again was
* Trans. Ent. Soc. Lond. 1903, pp. 87-91.
An Entomological Excursion to Bejar, Central Spain. 169
a belt of oak wood. On the summit of the Sierra a few
Dorcadion dejeant were seen, but all too worn to take;
also Chlenius dives, Amara oopte ra, various Cymindis and
Harpalus, Asida castellana, Steropus guiliani, a stoall
Olocrates, Zabrus, etc. Here, on Cytisus purgans,a curious
Cneorrhinus was met with: the ¢ of elongate shape and
with greyish scales, the 2 much larger, pyritorm, and with
green scales, the female in abundance, and the two sexes
often in company ; and with these occurred a few females
of the species found at Bejar on Genista florida. On the
stems of the Cytisus the local Cryptocephalus erosus was
often to be seen, but rarely captured, also various Cly-
thrids. In grassy places amongst this plant, on the Sierra,
Acmexodera teniata was to be found in profusion on flowers ;
and wherever there was any dung Aphodius bonvouloiri
could be taken in any quantity, occasionally in company
with the brilliantly-coloured Geotrupes coruscans. A patch
of uncultivated rocky ground near the town of Piedrahita, on
which there were many ancient evergreen oaks, and Genista
cinerea and the usual brooms, Eryngium, ete., produced a
good many interesting insects not seen elsewhere. On
July 19th an excursion was made to El Barco, on the
Tormes, in the valley between the “Bejar” and the
“ Gredos,” whence a rough road extends to the plains of
Estremadura. The long time occupied on the journey,
which was made in a not uncomfortable country cart
(tartana), gave us, however, but a few hours at the place,
our investigations being limited to the ground in the
immediate vicinity of the river. Here there was an
abundance of alder, sallow, and familiar marsh-plants, but
‘very little to be got by beating. Anomala rugatipennis
flew about pretty freely in the sunshine; and on the
flowers or stems of the gigantic Umbelliferze were Cardio-
phorus rufipes, Leptura fontenayi ; (f and §), Clytanthus
trifasciatus, Trichodes octopunciatus, an Agapanthia, and
Cerocoma schrebert, and an abundance of conspicuous
common Hymenoptera and Hemiptera. By the banks
of the Tormes Perileptus, Pxderus, Bledius, etc., were
found, as well as Pseudophleus, Emblethis, and other
Hemiptera.
On July 23rd we left Piedrahita for Avila, by the dili-
gence, a wearisome journey of six hours, remaining in the
last-mentioned place for three days. From this most
interesting old city the high ground was quite inaccessible,
.
170 Mr. G. C. Champion oz
though visible, so we had to content ourselves by collecting
in the immediate vicinity. The season was now too far
advanced for many of the beetles, though Zonabris, Cero-
coma, Clytanthus trifasciatus, and various Clythrids were
to be seen on flowers, and a Zhylacites abounded under
stones, exposed to the full heat of the sun. In the sandy
bed of the nearly-dried-up River Adaja there was, how-
ever, an abundance of interesting insects, including many
species of Bembidium and Philonthus, Aleochara rufipennis,
an undescribed Lathrolium, Tachyusa coarctata, Pederus
rujicollis, Heterocerus, Parnus, Aphodius, ete.
‘On our homeward journey we spent a day or two at
both Burgos and St. Jean de Luz, but the insects noticed
at these places do not call for special comment. At San
Sebastian, however, on June 23rd, we met with Orina rugu-
losa, STP S in some numbers, on a species of Centaurea, a
species that does not appear to have been recorded from
Spain.
The Coleopterous fauna of the Bejar and Piedrahita
districts seems, on the whole, to be very similar to that
of the Guadarrama, yet many genera are represented
by one or more different species, as Dorcadion, Nebria,
Creorrhinus, etc. The total absence of Coniferous trees,
so abundant at La Granja, must, however, make a great
difference, and it is probable therefore that the Guadar-
rama is much the richer of the two. Unfortunately, we
were unable to work the higher main Gredos range,
though rough accommodation could no doubt be obtained
at one or other of the small villages lower down in the
valleys on its northern or southern slope. The genus
Dima, amongst the Elateridz, well known from the
Guadarrama, was not found by us. The Sierra de Bejar,
it may be noted, could be much better worked from
Candelario, than from the more distant town of Bejar.
As regards the Hemiptera-Heteroptera, several of the
species have not previously been recorded from Spain, two
of the Capsidz being new.
The following is a list of the species of Coleoptera and
Hemiptera-Heteroptera taken, so far as at present identi-
fied. To avoid repetition the principal localities are in
most cases abbreviated thus :—S. Bej. = Sierra de Bejar;
Bej. = Bejar; Piedr. = Piedrahita; Av. = Avila.
An Entomological Excursion to Bejar, Central Spain. 171
COLEOPTERA.*
Cicindela hybrida, L., rarely, Av. Carabus melancholicus,
F., and (. latus, Dej., rarely, nodes stones, S. Bej. Nebria
paz, Seidl, one example, and JN. sp. n.?, a few speci-
mens, at the source of a small stream, near the melting
snow, S. Bej.: the types of the former were obtained
in the same locality. Bembidium carpetanum, Sharp
(the Guadarrama form of /. glaciale, Heer, of the Swiss
Alps, has been recently named &. carpetanum, and the
Sierra Nevada form B. montanum, Ramb., by Dr. Sharp),
in plenty, on the summit of the Sierra de Bejar,
near the melting snow. JB. elongatum, De}j., B. paulinoi,
Heyd., B. quadripustulatum, Serv., B. quadriguttatum, F.,
B. octomaculatum, Goeze, B. hispanicum, Dej., in profusion,
and others of the genus, Av., in the sandy bed of the
River Adaja; B. punctulatum, Drap., Av. and on the
banks of the Tormes at El Barco; B. maculatum, Dej.,
El Barco; B. ibericwm, Pioch., Piedr.; B. laterale, Dej., in
plenty, on the banks of streams, Bej., Piedr., Av. TZachys
parvulus, Dej. and T. sexstriatus, Duft., Bej. Trechus
pandellei, Putz. S. Bej. Perileptus areolatus, Creutz.,
banks of the Tormes, El Barco. Platynus marginatus,
L., Av.; P. viridicupreus, Goeze, Be}. Abacetus salzmanni,
Germ., Bej. Steropus ghiliani, Putz., under stones, S. Bej.,
Piedr.; S. sp.?, Piedr. Pacilus crenulatus, Dej., Be}.
Pterostichus nigrita, F., Bej.; P. azarz, Perez, S. Bej.,
four specimens. Amara ooptera, Putz. under stones,
summit of the Sierra de Bejar, a few specimens. Zabrus
constrictus, Graells, abundant, and Z. neglectus, Schaum,
4Z. silphoides, De}., and Z. estrellanus, Heyd., sparingly,
S. Bej. Chlenius dives, Dej., sparingly, and varying
greatly in colour, running in grassy places, near the
summit of the Sierras de Bejar and Piedrahita ; C. velu-
tinus, Duft., banks of the Cuerpo del Hombre, Bej.
Lelia eyanocephala, L., and L. trimaculata, Vill., sparingly,
by beating broom, oak, ete., Bej. Lienychus albonotatus,
Dej., banks of the Tormes, El Barco. Cymindis melano-
cephala, De}. (ruficollis, Graells), in plenty, and others of
the genus more sparingly, S. Bej.; C. variolosa, F., 8. Bej.,
Piedr.
* T am indebted to MM. Bedel, Fauvel, Reitter, and Schilsky, and
Dr. J. Daniel, for assistance in the determination of several of the
species enumerated.
172 Mr. G. C. Champion on
Haliplus spp., Bidessus ininutissimus, Germ., Deronectes
bicostatus, Schaum,* rarely, Hydroporus lepidus, Oliv.,
Hf, flavipes, Oliv., H. varius, Aubé, and A. tessellatus,
Drap., commonly, and various other species of the genus,
Liopterus ruficollis, Schall., Agabus brunneus, F., and A.
didynvus, Oliv., in plenty, and A. paludosus, F., A. heydent,
Wehncke, A. bipustulatus, L., and A. nitidus, F. , sparingly,
Liylius fenestratus, F., ete, in a small affluent of the
Cuerpo del Hombre, Bejar; Deronectes carinatus, Aubé,
not rare, and Dytiscus marginalis, L., in the river itself,
Bej. Gyrinus urinator, Ul. Orectochilus villosus, F.,
Be}.
Paracymus wneus, Germ., Calostoma hispanicum, Wiist.,
and various species of Phith ydrus, [Helochares, Limnebius,
Laccobius, Hydrochus, Helophorus, Hydrena, and Pornus,
Be}.
Aleochara rufipennis, Kr., not rare, in the sandy bed of
the Adaja, Av.; A. brevipennis, Grav., A. fuscipes, F., Be}.
Tachyusa co aretata, Er., Av. Notcthecta levicollis, Rey, in
an ants’ nest, Bej. Philonthus suavis, Bris., in abundance,
and -P. prea. eae and LP. pwnetus, Grav., rarely, 1
the bed of the Adaja, Av.; P. varius, Gyll., var. bimacu-
latus, Grav., P. laminatus, ibe P. politus, L., Be}. Scopeus
didymus, Er, Av. Sunius (Astenus) nh. sp., singly, under
a stone, Bej. Lathrobium n, sp.? ear L. punctatum,
Zett.), banks of the Cuerpo del Hombre, Bej. Pederus
ruficollis, F., common on banks of streams, Av., -Bej. ;
P. fuscipes, Curt., El Barco. Stenus longitarsis, Thoms.,
rarely, S. nanus, Steph., and S. lepriewri, Cussac, in plenty,
banks of stream, Bej. Dledius fracticornis, Payk., Av.;
L. hispidulus, Fairm., banks of the Tormes, El Barco.
Anthobium hispanicum, Bris., A. torquatwn, Marsh., A.
minutum, F., Philorhinum sordidum, Steph., Be}.
Mastigus palpalis, Latr., a few specimens by beating
herbage in damp places, Bej., also at Bafios. Si/pha sp.,
S. Bej. Olibrus bisignatus, Mén., in plenty on thistles, Be}. ;
O. xnescens, Kiist., not rare, Av. Dapsa acuticollis, Reitt.,
one specimen, Bej,, with the Mastic gus. Meligethes fuscus,
Oliv., Bej. Lrontes planatus, L., and Silvanus wiidentatus,
Oliv., under bark of an old cherry-tree, Bafhos. Aira-
philus sp., Be}. Atlagenus trifasciatus, F., common on
* The D. bicostatus of my Cuenca list (Trans. Ent. Soe. Lond.
1902, p. 122) was incorrectly identified ; it is unknown to Dr.
Sharp, and is probably an undescribed species.
An Entomological Excursion to Bean Central Spain. 173
flowers, Bej. Ctesias serra, Be}. Orphilus niger, Rossi,
Be}. Byrrhus depilis, Graells, 8. Bej. Ltinnichus incanus,
Kies., Bej., El Barco. Hister major, L., H. inequalis,
Oliv., H. amplicollis, Er., here and there, S. Bej.; 7.
sinuatus, Ill, common in dung, Bej., Piedr., Av.
Lucanus cervus, L, Dorcus parallelopipedus, L., Be}.
Platycerus spwnifer, Schauf., a few specimens taken on the
wing amongst old plants of Cytisus purgans, high up on
the Sierra de Bejar.
Ateuchus sacer, L, A. laticollis, L., 8. Bej. Sisyphus
scheffert, L., abundant, Bej. Gymnopleurus pilularius, L.,
El Barco; G. flagellatus, F., 8. Bej. Bubas bubalus, Oliv.,
S. Bej. Onitis hungaricus, Herbst, Av. Onthophagus
schrebert, L., common in dung, and other species of the
cenus, Bej. ; 0. verticicornis, Laich., Piedr.; O. furcatus, F.,
Av. Oniticellus flavipes, ¥., Piedr., Av. Aphodius bon-
voulowrt, Harold, in abundance, Sierras de Bejar au
Piedrahita ; A, ferrugineus, Muls., Bej;; A. seroja,, EF;
Piedr. ; A niger, Panz, and others of the genus, Bej. ;
A. varians, Duft., Av. Ammecvus sp., Be}. Pleurophorus
cxsus, Panz., Bej. Geotrupes coruscans, Chevr., Sierra de
Piedr. ; G. levigatus, F., G. hypocrita, Serv., etc., S. Bej.
Lhizotrogus pygialis, Muls., Piedr., Av., flying im the
sunshine; &. pineticola, Graells, El Barco. Serica mutata,
Gyll., commonly, by beating herbage in open places in an
oak wood towards evening, ‘Bej. Triodonta aquila, Lap. (2),
Be}. Hymenoplia, sp., 1 plenty, on stems of grass, etc., 5.
Be}j., Piedr. Chasmatopterus villosulus, Ill., males found in
profusion flying over grass, near the ground, looking very
much like Hymenoptera on the wing, paired specimens
found on flowers, later in the day, the black females
comparatively scarce, Bej. and 8. Bej.; C. hispidulus,
Graells, C. hirtulus, Ill, sparingly, Bej. Anomala rugati-
pennis, Graells, not rare, on Hchiwm or flying in the
sunshine, banks of the Tormes, El Barco; A. sp., on the
wing towards evening, Av. Anisoplia betica, Er., Bej.
Hoplia spp. Bej. Cetonia oblonga, Gory, C. morio, F.,
Bej., Piedr. Hpicometis hirta, Poda, El Barco. Leucocelis
stictica, L., Be}.
Anthawia hungarica, Scop., rarely, by sweeping low
plants, Bej.; A. hg ypomelena, Ill., not uncommon on L7ryn-
gum, Piedr.; A. funerula, Ill., A. millefolu, F., Piedr., Be}.
Acmeodera txniata, F., in abundance on Aieraciwm
flowers, amongst the dense growth of Cytisus purgans,
TRANS, ENT. SOC, LOND. 1903.—PART II. (JUNE) 12
174 Mr. G. C. Champion on
Sierra de Piedr.; A. discoidea, F., A. adspersula, Tll., ete.,
Bej. Chrysobothrys affinis, F., one specimen, on an old
stem of Cytisus, 8. Bej. Corabus fasciatus, Vill., rarely,
by beating herbage, also found dead in an ants’ nest
under a stone, San Gil, near Bejar; C. xneicollis, Vill,
Bej.; C. giblicollis, Ill, Piedr., Bej. Agrilus biguttatus,
F., on oak, Bej.; A. cinctus, Oliv., not rare on Cenista
jlorida, Bej., Piedr., El Barco; A. roscidus, Kies., A. solieri,
Cast., A. hyperict, Cr., A. angustulus, Ill., Be}. Aphanis-
ticus elongatus, Villa, and A. emarginatus, F., Bej., by
sweeping. TZ'rachys refleca, Gené, one specimen, by sweep-
ing on the banks of a small stream, Bej., perhaps new to
the Spanish list.
Drasterius bimaculatus, Rossi, Be}. Hlater preustus, ¥.,
var. aurilegulus, Schauf., in numbers, by beating oak, on
the hillside near the town of Bejar; LZ. nigerrimus, Lac.,
one specimen, with the preceding, Bej. Jelanotus tene-
brosus, Er., and M. castanipes, Payk., common, Be}.
Cardiophorus signatus, Oliv., im abundance by beating
broom, etc., in oak woods; C. eqgwiseti, Herbst, C. graellsi,
Cand., etc., with C, signatus, but more sparingly, Bej.; C.
rufipes, Goeze, El Barco. Athous godarti, Muls., and other
species of the genus, not yet determined, Bej. Corymbites
hemapterus, Ill., under stones, on the summit of the Sierra
de Bejar, not rare; C. gowgeleti, Fairm., in numbers on the
hillside close to the town of Bejar, on stems of low
plants, on the wing, etc.; C. kiesenwetteri, Bris. (2), C.
latus, F., C. amplicollis, Germ., C. nivicola, Kies., ete.,
rarely, S. Bej. Stlesis rutilipennis, Ill, Be}.
Hydrocyphon sp.?, in plenty on sallows, Bej. Lampyris
sp., Be}. Telephorus, Rhagonycha, Malthinus, Malachius,
Attalus, Charopus, Hypebeus, Dasytes, spp. undetermined,
Faplocnemus albipilis Kies., ete., Bej. Henicopus, 2 spp.,
S. Bej., Piedr.: (1) (Z. heydeni, Kies.) very large, the ¢
deep black, with entirely black pubescence, the 2 with
ereyish- white hairs, the ¢ in abundance, and the 2
rarely, on grass-stems, etc., on the mountain slopes; (2)
(?H. rugosicollis, Duv.) $ and 2 with grey pubescence.
Danacxa atripes, Graells, Bej. Amawronia elegans, Kies.,
Bej. Trichodes octopunctatus, F., not rare, on Umbelliferze,
El Barco., 7. leucopsideus, Oliv., 7. ammios, F., Bej.
Lasioderma lxve, Ill., on flowers, Bej., Piedr. Xylopertha
sp., Be}.
Tentyria spp., 8. Bej., in plenty, running about on the
An Entomological Excursion to Bejar, Central Spain. 175
slopes, 8. Bej. Dichillus subcostatus, Sol., rarely, under
stones, Bej. Akis acuminata, F., in abundance, under old
straw, under the shelter of a large rock, Av. Scawrus
striatus, F., in numbers, under dung, etc., 8. Bej. Asida
castellana, Graells, in abundance at or near the summit of
the Sierra de Bejar, and rarely on that of Piedrahita,
running about in the sunshine, especially about dung in
the tracks made by goats amongst the Cytisws bushes ; A.
goudott, Sol., rarely, S. Bej., also taken at Burgos, and at
Guadalaviar and Albarracin, in 1901; A. sericea, Oliv., S.
Bej., also taken at Burgos, and at Tragacete and Albarracin,
in 1901. Pimelia sp., a few specimens, on the paths, near
dung, S. Bej. Crypticus kraatzt, Bris. (?), and Oochrotus
unicolor, Lue., rarely, under stones, Be}. Olocrates saxicola,
Muls. (saxeticola, Graells), a few specimens apparently
belonging to this species, found under stones, summit of
the Sierra de Piedrahita; O. foveolatus, Graells, i in abund-
ance, summit of the Sierra de Bejar, running about in
the sunshine, frequently in copula. Dendarus, Heliopates,
Micrositus ulyssiponensis, Germ., Gonocephalum, spp. un-
determined, Bej., Av., ete. Calometopus clypeatus, Germ.,
rarely, under bark of an old cherry-tree, and also under
a stone, Banos, Bej.: recorded by Graells as having been
found in a rotten chestnut-tree at Tornavacas. /elops
laticollis, Kiist. (2), by beating Grenista, etc., not rare, Be}.
Hymenalia rufipes, F., not rare, Bej. Omophlus ruficollis,
F., in abundance everywhere on flowers, Bej., Piedr.; 0.
incertus, Muls., O. picipes, Fabr., rarely, Bej. Lagria lie,
F., common on stone walls, etc., Piedr.. Av.; Z. grenieri,
Bris., Bej.; L. hirta, L., L. atripes, Muls., L. rubida, Graells
(parvula, Perris), Bej., the last mentioned in abundance
by sweeping low plants towards evening, but males only
seen, Bej. Scraptia dubia, Oliv., by beating oak, Be}.
Notoxus trifasciatus, Rossi, Bej., El Barco, Av.; NV. platy-
cerus, Laf., banks of the Tormes, El Barco. <Anthicus
rodriguesi, Latr., common, on the ground beneath low
plants, Bej.; A. quadriguttatus, Rossi, Av.; A. tristis,
Schmidt, Bej. Ochthenomus wunifasciatus, Bon., Bej.
Mordella aculeata, L., Mordellistena spp. Anaspis spp.,
Bej. Silaria brunnipes, Muls., S. trifasciata, Chevr.,
common on flowers, Bej. Hmenadia larvata, Schrank,
one specimen, on a flower, Piedr. Meloe majalis, L.,
common, occurring even high up on the Sierra, Bej.
176 Mr. G. C. Champion on
Cerocoma schrebert, F., locally common, on flowers, Bej.,
Piedr., Av. Lytta vesicatoria, L., in plenty in various
places, numerous males often seen near a single female,
on Cytisus purgans and C. scoparius, Be}j., Piedr., most of
the specimens with a space down the middle of the disc of
each elytron coppery or golden. Zonabris, various species,
some in great abundance on the Cytisus purgans and
Genista florida: Z. quadripunctata, L., Bej., Piedr., Av.;
Z. dejeani, Gyll., Bej.; Z. amori, Graells (2), Bej., El Barco;
Z. geminata, F., Piedr., and others. Coryna enon Gyll.,
Bej. Zonitis preusta, Fr, EH] Barco. Cidemera simplex, L.,
Bej., Piedr.
Otiorrhynchus dentipes, Graells, in plenty, running about
in the sunshine, summits of the Sierras of Bejar and
Piedrahita. Phyllobius sp. (near P. tuberculifer, Chevr.),
not rare by beating herbage, Be}. Polydrusus setifrons,
Duv., P. confluens, Steph. (2), on the Genista, and others
of the genus, Bej., Piedr.; P. (Homapterus) afinis, Chevr.,
very rarely, by sweeping long grass, 8. Bej. Sczaphilus
carinula, Oliv. (cristatus, Graells), mm plenty on Genista
florida, Bej. Barypithes suleifrons, Boh. commonly, by
sweeping long grass, in a damp place on the mountain
slope, S. Be}. Strophosomus erinaceus, Chevr., in numbers,
by beating oak, etc, Bej. Brachyderes incanus, L. i) in
abundance, with the preceding, Bej. Sidones gressorius, ¥F.,
S. regentsteinensis, Herbst, S. erinitus, Herbst, 8S. flavescens,
Marsh., and its var. cinnamomeus, All., and others of the
genus, chiefly on Genista florida, Bej. Cneorrhinus: various
species of the subgen. Afactogenus occurred at Bejar or
Piedrahita, and although these were the most character-
istic and interesting forms of Curculionidze obtained during
the trip, I am quite unable to name them with any
certainty from the published descriptions of Boheman,
Graells, etc. One of these forms, found in abundance on
or near the summit of the Sierra de Piedrahita, on Cytisus
purgans, is very nearly allied to C. pyriformis, Boh.,
and C. dispar, Graells: the female is of large size and
uniformly clothed with green scales; the male, which was
comparatively rare, and, when seen, usually in company
with the female, is narrow and elongate in form, and
clothed with greyish scales, there being always a few
green scales round the eyes, and sometimes several more
scattered about on the elytra or legs. [A very similar
An Entomological Excursion to Bejar, Central Spain. 177
species occurs at La Granja, specimens of which have been
given me by M. Bedel: this insect, too, has a green female *
and a grey male; the male, however, is less elongate than
the Piedrahita examples of the same sex.] Another form,
occurring in abundance in the immediate vicinity of Bejar,
on Genista florida, varying greatly in size and colour, with
dull greyish scales, the prothorax bi- or trivittate, and the
elytra with the margins whitish and the disc often mottled
with lighter or darker colour, may be referable to C. dispar,
var. meleagris, Graells: all the specimens obtained at
Bejar appear to be of the female sex (the smallest and
narrowest examples have been dissected, to make certain),
and a few females taken at Piedrahita, on Cytisus purgans,
seem to belong to the same species, which I also have
from Ferrol, N. Spain. <A third form, represented by a
single female specimen from Bejar, similar in colour to
(and perhaps an extreme form of) the preceding, bas the
eyes more prominent, the prothorax more transverse, and
the punctures of the striz closely placed and rather coarse.
[C. graellsi, Bris., quoted in Reitter’s Catalogue as a
synonym of C. dispar, var. meleagris, it may be noted, is
recorded as having been taken in plenty at the Escorial,
though females only were found; it is unknown to me.
The species of the <Atactogenus-group of Cneorrhinus
evidently require a thorough revision. It is a curious fact
that in two different localities a species has been taken in
abundance, without the males being obtaimed, and that
when specimens of this sex have been noticed, as at
Piedrahita, they were far outnumbered by the females.]
C. spinipes, Perez, sparingly on Cytisus, Bej. Thylacites
turbatus, Gyll., in abundance on the arid _ hillsides,
under stones fully exposed to the great heat of the sun,
Av.; 7. spp. (2), Piedr. Liwus iridis, Oliv., L. ascanit, L.,
L. spartii, Oliv., and LZ. cardui, Oliv., Bej., one or two of
them on Cytisus or Genista. Larinus spp., Antsorrhynchus
spp.,and Siieronyx sp., Be}. Pachytychius sparsutus, Oliv.,
not rare, on the woolly pods of Cytisus welwitschi, Piedr., Be}.,
Batios; P. scubricollis, Rosenh., in plenty by sweeping low
plants in dry places, Bej. Ceuthorrhynchidius wrens, Gyll.,
Be}. Ceuthorrhynchus geographicus, Goeze, C. biscutellatus,
Chevr., C. larvatus, Schultze, C. squamulatus, Bris., C.
melunostictus, Marsh., var. nvurinus, Gyll. C. denticulatus,
* This form should certainly be named OC. pyriformis, the type of
which, from Lusitania, I have recently seen.
178 Mr. G. C. Champion on
Schr., C. gougeleti, Bris., C. eleganiulus, Faust, var. gracilis,
Schultze, Bej. Baris timida, Rossi, Be}. Sibinia spp., Be}.
Rhynchenus quercus, L., L. sparsus, Fahr., R. avellane, Don.,
hk. ilicis, F., Bej. Lhamphus flavicornas, Clairv., Be).
Cionus blattari iw, F., Bey. Nanophyes duriewi, Lue., bred in
abundance from galls on Cotyledon, Bej., and one specimen
swept from herbage, El Barco; WV. lythri, F., in plenty on
Lytirum, Bej.; N. niger, Waltl, NV. rubricus, Rosenh., and
other species of the genus, Bej. Apion elongatissiinum,
Desbr., Bej., on Cytisws welwitschi, in numbers, and sub-
sequently bred from seed-pods of that plant brought home
to England; A. putoni, Bris., and A. argentatum, Gerst.,
more or less plentiful, on Genista florida, Be}. ; A. brevirostre,
Herbst, 4. cyanescens, Gylk, A. alcyonewm, Germ., A.
cantabricum, Desbr., A. kraatzi, Wenck., A. striatum,
Marsh., A. immune, Kirby, A. levigatum, Payk. (sorbe, EF.)
Cf ¢), A. xthiops, Herbst, and others of the genus, Bej.;.
A, schonherri, Boh., Piedr. Hypoborus ficus, Er, Be}.
Leptura fontenayi, Muls., a few specimens on Umbelliferz,
Piedr., El Barco. Strangalia nigra, L., and S. maculata,
Poda, Be}. Stenopterus ater, L., Piedr. Phymatodes testaceus,
L., Bej. Plagionotus arcuatus, L., a few specimens, Be).
Clytanthus trifasciatus, F., common on flowers (Lryngium,
Umbelliferse, etc.), Bej., El Barco, Av.; C. ruficornis, Oliv.,
one specimen, Piedr. Dorcadion dejeani, Chevr., in abund-
ance on or near the summit of the Sierra de Bejar (whence
it has been recorded by Dr. Seidlitz), at the beginning of
July, and a few worn specimens seen later in the month
on the Sierra de Piedrahita; it frequents grassy places,
and runs about in the sunshine. Tetvops preusta, L., Be}.
Agapanthia cardui, L., and others of the genus, Bej., El
Barco. Calamobius filum, Rossi, Be}. Phytecia virescens,
F., etc., Bej.
Donacia thalassina, Germ., Be}.; D. discolor, Panz., Piedr.
Labidostomis lusitanica, Germ., Be}. Lachnea vicina, Lac.,
Bej., Piedr. ; Z. pubescens, Duf., not rare, on Genista cinerea,
Be}., Piedr, LL. sexpunctata, Scop., common, on oak, readily
taking to wing in the sunshine. Zitubwa seamaculata, ¥.,
Bej., El Barco, Av.; 7. biguitata, Oliv., Piedr. Gynan-
drophthalina affinis, Hellw., Piedr. ; G. concolor, F., common,
on Genista florida, ete., Bej., Piedr.; G. amabilis, Lac., on
Genista horrida and Cytisus purgans, on the mountain
slopes, Bej., Piedr. Clythra atraphaaxidis, Pall, Av.; C.
leviuscula, Ratz., on oak, Bey. Coptocephala scopolina, L.,
An Entomological Excursion to Bejar, Central Spain. 179
Av., Piedr. Cryptocephalus bimaculatus, F., Bej., Piedr. ;
C. imperialis, Laich., Bej.; C. sexmaculatus, Oliv., Bej.,
Piedr.; C. bipunctatus, L., Bej.; C. quadripunctatus, Oliv.,
Bej.; C. erosus, Seidl., frequently seen and occasionally
caught, on Cytisus purgans, Piedr., Be}. (originally de-
scribed from Bejar); C. more, L. , Bej.; ; C. crassus, Oliv.,
not rare, Piedr., Bej., one specimen bred from a cocoon
found on Genista « C. koyt, Suffr., the commonest species
of the district, on flowers, etc., the sexes differing consider-
ably in the coloration of the prothorax, and the markings
of the elytra varying in both sexes, Bej., Piedr.; C. globi-
collis, Suffr., not rare on flowers in dry places, the elytra
usually of a coppery or golden colour, Bej.; C. cwrulescens,
Sahlb., Bej.; C. violaceus, Laich., common, on flowers, Bej.,
Piedr.; C. celtibericus, Suffr., occasional, on Cytisus and
Genista ; C. pygmeus, F., El Barco, Bej.; and others of
the genus. Pachybrachys viridissimus, Suttr., Bej., Piedr. ;
P. pteromelas, Graells, rarely, on Genista cinerea, Piedr.; P
sp.(2), Bej. Stylosomus sp., on arborescent heath, Bej.
Limarcha sp. (with femora and tibie partly red), not rare,
near the snow, 8. Bej. Chrysomela americana, L., and
others of the genus, Bej. Phytodecta variabilis, Oliv, and
P. olivacea, Forst., on Cytisus or Genista, Bej. Malacosoma
lusitanicum, L., abundant, on flowers, etc., Bej., Piedr., ete.
Luperus nigrofasciatus, Goeze (cirewmfusus, Marsh.), in
profusion, on Grenista florida; L. lividus, Joann., Be}. ; L.
niger, Goeze, Piedr. Bej. Adimonia tanaceti, L., Be}.
Ibolia timida, Ul. (?), not rare, on Lryngiwm, Piedr., and
others of the genus, Bej. Psylliodes cyanoptera, Il., ete.,
Bej. Chetocnema chlorophana, Duft., Bej. <Apteropeda,
Phyllotreta, Longitarsus, Aphthona, spp. undetermined,
Bej. Haltica sp., in profusion on arborescent heath,
specimens often brassy in colour, Bej. Hispa atra, Te
rarely, Piedr. Cassida spp., Be}.
Epilachna argus, Foure., Be}. Coccinella 14-pustulata,
L., C. variabilis, F., Bej., Piedr. Adalia mutabilis, Seriba,
Be}. Exochomus auritus, Scriba, Bej. Platynaspis villosa,
Foure., Bej. Hyperaspis reppensis, Herbst, Bey. Scymnus
areuatus, Rossi, and others of the genus, Bej.
180 Mr. G. C. Champion on
HEMIPTERA-HETEROPTERA.*
Odontotarsus grammicus, Linn., Be}. Psacasta cerinthe,
Fabr., one specimen, Bej. Hurygaster maura, Linn, £.
nigrocucullata, Goeze, Be}. Graphosoma lineatum, Linn.,
on flowers, Bej., El Barco. Geotomus punctulatus, Costa,
Be}. Brachypelta aterrima, Forst., Piedr. Ochetostethus
nanus, H.-8., Bej. Sehirus dubius, Scop. Bej. — Sezocoris
helferi, Fieb., Be}. dtlia rostrata, Poh., Bej., Piedr. Neotti-
glossa leporina, H.-S8., Bej. Rhaphigaster nebulosa, Poda,
Be}. Prionotylus brevicor mis, M. & R., Be}. Carpocoris
purpuripennis, De G., Be}. Peribalus sphacelatus, Fabr., Be}.
Hurydema festivum, Linn., El Bareo; E. oleracewm, Linn.,
Be}. Zicrona coerulea, Linn., Bey. Verlusia suleicornis,
Fabr., and V. quadrata, Fabr., Be}. Coreus hirticornas,
Fabr., Be}. Pseudophleus falleni, Schill. at the roots of
plants on the banks of the Tormes, El Barco; P. waltli,
H.-8., Be}. Strobilotoma typhecornis, Fabr. , Bej. Centro-
coris spiniger, Fabr., Bej., Av. Micrelytra Jossularum,
Rossi, Be}. Camptopus lateralis, Ger., Piedr., Bej. Stenoce-
phalus agilis, Scop., Be}. Corizus crassicor nis, Linn., C.
hyalinus, Fabr., C. rufus, Schill. C. parumpunctatus, Schill,
Bej. Neides aduncus, Fieb,, in plenty by sweeping low
plants in dry places, Bej., Piedr.; NW. tipularius, Linn., Be}.
Berytus hirticornis, Brullé, 2B. montivagus, Fieb., Be}.
Lygexus pandurus, Scop., L. saxatilis, Scop., L. albomaculatus,
Goeze, Bej.; LZ. superbus, Poll., in abundance under stones,
Bey. Nysius graminicola, Kol., WN. lineatus, Costa, Bej. ;
NV. punctipeniis, H.-8., two specimens, in sandy places on
the banks of the Tormes, El Barco. Cynius melanocephalus,
Fieb., common, Piedr., Bej. Cymodema tabidum, Spin.,
Bej. Lschnorrhynchus geminatus, Fieb., Bej. Geocoris
stculus, Fieb., El Barco. Heteroguster artemisix, Schill.
Be}. Micropla: v interrupta, Fieb., Piedr., Bej.; J. plagiata,
Fieb., Av. Metotoplax ditomoides, Costa, in plenty on low
plants, Bey. Macroplax fasciata, H.-8., Bej. Pterotinetus
staphylinoides, Burm., Bej., Piedr. Plinthious brevipeniis,
Latr. (macropt. form), Bej. Aoploseelis bivirgatus, Costa,
Av., Piedr. Peritrechus sylvestris, Fabr., Piedr. Rhyparo-
chromus sabulicola, Thoms., Bej., also found at Burgos.
* T am indebted to Dr. O. M. Reuter for the determination of
most of the Capsids and Anthocorids, and to Dr. G. Horvath and
Mr. E, Saunders for assistance with the Lygveids, ete.
An Entomological Excursion to Bejar, Central Spain. 181
Trapezonotus arenarius, Linn., T. ullricha, Fieb., Bej.
Aphanus pint, Linn., El Barco ; A. quadratus, Fabr., El
Barco, Be}. Beosus maritimus, Scop., Bej. Lasiocoris
anomalus, Kol., Bey. Lmblethis angustus, Mont., in numbers,
Bej., El Barco. Notochilus andrez, Puton, three specimens,
Bej., probably not previously recorded from Spain. Monan-
thia echii, Wolff, in profusion in all its stages, on Hehiwm,
El Barco, Bey. Phyllontocheila awriculata, Costa, Bej.; P.
cardui, Linn., Bey. Dictyonota fuliginosa, Costa, in numbers,
by sweeping low plants, Bej.; P. strichnocera, Fieb., Be}.
Catoplatus carthusianus, Goeze,in abundance on Eryngium,
Piedr. Hydrometra stagnorum, Linn., Bej. Gerris najas,
De G., River Cuerpo del Hombre, Bej. Harpactor wra-
cundus, Poda, Bej.; H. sanguineus, Fabr., Bej. Coranus
Lgyptius, Fabr. , El Barco. Nabis reuter ianus, Puton, Piedr. ;
N. rugosus, Linn., Bej.; WV. ferus, Linn., Be}. ; J. lativentris,
Boh., Be]. Salda cocksi, Curt., Bej., Piedr.; S. prlosella,
Thoms. (7), in the dried- up bed of the Adaja, Av. Piczostethus
maculipennis, Baer., Be}.; P. terricola, Reut., one specimen
under a stone, Be}. Z'riphieps discolor, Reut., Bey. Pithanus
merkeli, H.-S., Bej. MMegalocerea erratica, Linn., Bej.
Lopus gothicus, Linn., Bej. Homodemus M-flavum, Goeze,
Piedr. Adelphocoris vandalicus, Rossi, Be}. Calocoris
roscomaculatus, De G., Bej.; C. sulphurews, Reut., Av.
Camptobrochis lutescens, Schill., Bey. Brachycoleus triangu-
lars, Goeze, in abundance on Lryngium, and varying
ereatly i in colour, Piedr., Bej. Capsus cordiger, Hahn, and
var. jfastidiosus, Reut.,* C. scutellaris, Fabr., C. ruber,
Linn., Bej. Strongylocoris lewcocephalus, Linn., and 8.
obscurus, Ramb., common, Be}. Cyllocoris histrionicus,
Linn., Bej. Globiceps pictetr, Fieb., Bej., El Barco. Tini-
cephalus discrepans, Fieb., Be}. Conostethus rosews, Fall,
Be}. Dicyphus geniculatus, Fieb., var. disjwnctus, Reut.,*
Bej.; D. hyalinipennis, Klug., et rare, by sweeping low
plants, Bej.; D. pallicornis, Fieb., var., Piedr., Be]. Grypocoris
noualhiert, Reut., Bej. Orthocephalus saltator, Hahn, Be}.
Dimorphocoris lividipennis, Reut.,* Bej., not rare. Hypsi-
tylus prasinus, Fieb., Bej. Platycranus metriorhynchus,
Reut., Piedr. Systedlonotus championi, Reut.* (gf and
2), Bej. Megalocoleus bolivari, Reut. Bej., in great
abundance on a yellow-flowered aromatic composite plant
(? Santolina sp.), the species in lite being of the same
colour as the flowers. Sthenarus ocularis, M. & R., Piedr.,
Be}. Onychumenus decolor, Fall., Bej.
182 An Entomological Excursion to Bejar, Central Spain.
The species or varieties marked with an asterisk are
described in the Entomologist’s Monthly Magazine for
May 1903, pp. 119-121.
The following species of Capsidze may be added to my
previous list (Trans. Ent. Soc. Lond. 1902, p. 129):—
Phytocoris exoletus, Costa, Albarracin; P. femoralis, Fieb.,
Bronchales; P. vittiger, Reut., var., Albarracin.
( 188 )
XI. An Account of a Collection of Rhopalocera made on
the Anambara Creek in Nigeria, West Africa. By
Percy I. Latuy, F.E.S.
[Read February 4th, 1903. ]
PuLaTe VIII.
THE collection of Rhopalocera dealt with in the following
pages is the largest that has yet been made on the Niger
River; in all it contains two hundred and twenty-six
species, twelve of which are undescribed.
Messrs. Godman, Salvin, and Druce, P.Z.S., 1884, pp.
219-229, and Mr. G. H. Carpenter, Proc. R. Dubl. Soc. (2)
8, pp. 804-310, 1895, have written two papers on Lepi-
doptera from this district; these two papers enumerate
seventy-two species, seventeen of which are not in the
collection here described, the names of these I give later.
I am indebted to Mr. Alex. J. Braham, the collector, for
the following information on the district.
The Anambara Creek flows into the Niger River just
above Asaba; the mouth of the creek is about three
hundred miles from Forcados, and it extends (navigably)
about a hundred miles inland. The whole country is flat,
with only a few slight hills about sixty to eighty metres
above sea-level; the banks of the waterways are densely
clothed with forest and undergrowth, away from the water
open bush prevails. The vegetation is chiefly of mush-
room growth, soft sap trees of the “paw-paw” type,
mingled largely with cocoanut and date-palms; ebony
and mahogany are also to be found, and small mangroves
are in profusion; orcbids are rarely seen in flower, in fact
flowers, the majority of which are glaring yellow in colour,
are seldom seen. The rainfall is very heavy, the difference
in the height of the rivers in wet and dry seasons being
as much as sixty feet; in the wet season the whole
country is practically under water, the high forest trees
appearing merely as bushes on its surface.
The season may be divided as follows: middle of
January to May intensely hot and damp, with many fierce
tornadoes; the weather is then cooler till July, when the
TRANS. ENT. SOC. LOND. 1908.—PART II. (JUNE)
184 © MroPietathys
rainy season commences and lasts till the second week in
October; during November and December dry winds
prevail, and all vegetation is dried up, the temperature
at noon rising as high as 110° Fahr. in the shade, and
dropping to 60° Fahr. at night.
I wish to express my thanks to Mr. Francis A. Heron,
of the Natural History Museum, South Kensington, for
his assistance in determining many of the species, and to
Mr. Henley Grose-Smith, who was so kind as to look over
the new Lyczenidee.
The types of all the new species here described are in
the collection of Mr. Herbert J. Adams.
1. Danais chrysippus.
Pap. chrysippus, L., Syst. Nat., ed. 10, p. 471 °(1758).
The variety alcippus, Cram., appears to be the prevalent
form.
2, Danais petiverana.
D. petiverana, D. and H., Gen. D. L., p. 98 (1847).
3. Amauris vashti.
D, vashii, Butl., Cist. Ent. 1, p. 1 (1865).
4. Amauris niavius,
Pap. miavius, L., Syst. Nat., ed. 10, p. 470 (1758).
5. Amauris psyttalea.
A. psyttalea, Plotz, 8. EK. Z. xli, p. 189 (1880).
6. Amauris hecate.
D. hecate, Butl., P. ZS. 1866, p. 44.
7. Elymnias phegea.
Pap. phegea, Fabr., Ent. Syst. ili, 1, p. 1382 (1798).
8. Lymnias banmakoo,
Mel. bammakoo, Westw., Gen. D. L., p. 405 note,
t. 68, f. 3 (1851).
9. Melanitis leda.
Pap. leda, L., Syst. Nat., ed. 10, p. 474 (1758).
10. Gnophodes parneno,
G. parmeno, D. and Hi; Gen. D. Ly t.-6L, £ 21850).
11. Lieyclus phalanthus.
D. phalanthus, Stgr., Exot. Schin., p. 229, t. 82 (1887).
12. Mycalesis tenias.
M. tenias, Hew., Exot. Butt. Mycalesis, t. 8, f. 55, 56
(1865°.
14.
HG:
ie
Account of a Collection of Rhopalocera. 185
3. MMycalesis ignobilis.
M. ignolilis, Butl., Trans. Ent. Soc. Lond., 1870,
p. 124.
Monotrichtis sandace.
Mycalesis sandace, Hew., Exct. Butt. Mycalesis, t. 10,
fGp9( US (0).
. Monotrichtis mailyas.
Mycalesis milyas, Hew., Exot. Butt. Mycalesis, t. 6,
f. 34 (1864).
Monotrichtis safitza.
Mycalesis safitza, Hew., Gen. D. L. i, p. 394 note,
t. 66, f. 3 (1851).
Monotrichtis vulgaris.
Mycalesis vulgaris, Butl., Cat. Satyr., p. 130, t. 3, f. 8
(1868).
. Monotrichtis nebulosa.
Mycalesis nebulosa, Feld., Reise Novar. Lep., p. 502
(1867).
All the specimens belong to the form agraphis, Karsch.
IGE
Monotrichtis dorothea.
Pap. dorothea, Cram., Pap. Exot. i, p. 19, t. 204,
feb Es (799):
pthima doleta.
Y. doleta, Kirby, Pr. R. Dublin Soc. (2) u, p. 336
(1880).
We
. Vpthima simplicia.
Y. simplicia, Butl., An. N. H. (4) xviii, p. 481 (1876).
2. Acrexa admatha.
A, admatha, Hew., Exot. Butt. Acrea, t. 3, f. 16, 17
(1865).
3. Acrea seis.
A, seis, Feisth., Ann. E. Fr. (2) vii, p. 247 (1850).
. ACTRA CAMBNA,
Ancomeng, Wnrur, Wily Mxoty ins, 1) ps i2t. 7, 1 2
(1773).
. Acreva zetes.
Pap. zetes, L., Syst. Nat., ed. 10, p. 487 (1758).
. Acrxa egina.
Pap. egina, Cranf., Pap. Exot. i, p. 128, t. 81, f. C, D
(1775).
186
A
Mr. P. I. Lathy’s
. Acreva abdera.
A. abdera, Hew., Exot. Butt. Acrzea, t. 1, f. 1, 2
CUS52):
. Acrva pseudegina.
A. pseudegina, Westw., Gen. D. L., p. 5381 (1852).
. Acreva terpsichore.
Pap. terpsichore, L., Syst. Nat. ed. 10, p. 466 (1758).
. Acrvxa vinidia.
A, vinidia, Hew., Ent. M. Mag. xi, p. 180 (1874).
. Acrea bonasia.
Pap. bonasia, Fabr., Syst. Ent., p. 464 (1775).
2. Acrea pharsalus,
A. pharsalus, Ward, Ent. M. Mag. viii, p. 81 ee:
. Acrea encedon.
Pap. encedon, L, Syst. Nat., ed. 10, p. 488 (1758).
variable series, including several examples of ab
aleippina, Auriv.
D4,
Acrea pentapolis.
A, pentapolis, Ward, Ent. M. Mag. viii, p. 60 (1871).
A variable series showing specimens intermediate be-
tween the type, and the form described by Oberthiir
under the name of the/estis; this name will therefore have
to sink as a synonym.
35.
Acrea orestia.
A. orestia, Hew., Ent. M. Mag, xi, p. 131 (1874).
. Acrea parrhasia.
Pap. parrhasia, Fabr., Ent. Syst. iii, 1, p. 175 (1893).
Acrea peneleos.
A, pencleos, Ward, Ent. M. Mag. viii, p. 60 (1871).
. Acrea lycoa.
A. lycoa, Godt., Enc. Meth. ix, p. 239 (1819).
9. Acrea jodutta.
Pap. jodutta, Fabr., Ent. Syst. i, 1, p. 175 (1793).
The series includes ? ab. carmentis, D, and H.
40.
Planema epea.
Pap. epea, Cram., Pap. Exot. iii, p. 64, t. 230, f. B, C
(1779).
41.
46.
47,
48,
Account of a Collection of Rhopalocera. 187
Planema consanguinea.
P. consanguinea, Auriv., Ent. Tidskr. xiv, p. 282, f. 8
(1893).
2. Planema camerunica.
P. camerunica, Auriv., Ent. Tidskr. xiv, p. 282, t. 6,
f. 4, 5 (1898).
. Planema umbra.
Pap. wmbra, Dru., Ill. Exot. Ins. ii, p. 23, t. 18, f. 1,
2 (1782).
. Planema vestalis.
Acrea vestalis, Feld., Reise Novar. Lep., p. 369, t. 46,
f. 8, 9 (1865-7).
. Atella phalantha.
Pap. phalantha, Dru., Ill. Exot. Ins. i, p. 41, t. 21,
fl 27a):
Hypanartia delius.
Pap. delius, Dru., Ill. Exot. Ins. 11, p. 18, t. 14, f. 5, 6
(1782).
Pyrameis cardut.
Pap. cardwi, L., Syst. Nat., ed. 10, p. 475 (1758).
Precis madagascariensis,
Junonia madagascariensis, Guén., Vinson, Voy. Mad.
Lep., p. 37 (1864).
. Precis clelia.
Pap. clelia, Cram., Pap. Exot. i, p. 38, t. 21, f E, F
iis):
. Precis sophia.
Pap. sophia, Fabr., Ent. Syst. ii, 1, p. 248 (1793).
. Precis octavia.
Pap. octavia, Cram., Pap. Exot. ii, p. 60, t. 135, f. B, C
. Precis pelarga.
Pap. pelarga, Fabr., Syst. Ent., p. 513 (1775).
. Precis laodice, Cram., Pap. Exot. ii, p. 64, t. 138,
f, G, H. (1777).
. Precis milonia.
P. milonia, Feld., Reise Novar. Lep., p. 403 (1867).
. Precis terea.
Pap. terea, Dru., Tl. Exot. Ins, ii, p. 32, t. 18, f. 3, 4
(1773).
188
63.
64.
Mr. Ee abathys
. Precis chorimene.
Vanessa chorimene, Gueér., Icon. R. Anim., p. 476
(1844).
. Catacroptera cloantha.
Pap. cloantha, Cram., Pap. Exot. iv, p. 93, t. 338,
f, A,B Gd/7sl):
Salamis temora.
S. temora, Feld., Reise Novar. Lep., p. 404 (1867).
. Salamis anacardii.
Pap. anacardu, L., Syst. Nat., ed. 10, p. 467 (1758).
. Hypolimnas misippus.
Pap, misippus, L., Mus. L. Ulr., p. 264 (1764).
. Hypolimnas dubius,
Pap. dubius, Palisot, Ins. Afr. et Amér., p. 238, Lep.
t. 6, f. 2a, 2b (1805).
. Hypoliinnas anthedon.
Diadema anthedon, Doubld., Ann. N. H. xvi, p. 181
(1845).
Kallima rumia.
K. rwmia, Doubld. and Westw., Gen. D. Lep., p. 325,
tola2s 21850).
Kallima eynodoce.
Pap. cymodoce, Cram., Pap. Exot. 11, p. 5, t. 99, f. G, H
CLOKG:
A single male of this rare species.
65.
68.
69.
Hurytela hiarbas.
Pap. hiarbas, Dru., Il. Exot. Ins. 11, p. 17, t. 14, f. 1,
22):
. Hurytela dryope.
Pup. dryope, Cram., Pap. Exot. i, p. 125, t. 78, f. E, F
(1775).
7. Neptidonsis onhione.
1 ']
Pap. ophione, Cram., Pap. Exot. 1, p. 27, t. 114, f. E, F
(eiere.
Ergolis enotrea.
Pap. enotrea, Cram., Pap. Exot. ii, p. 73, t. 236, f.
Agu 19):
Mescxantha ethosea.
Pap. ethosea, Dru., Ill. Exot. Ins. in, p. 51, t. 37, f. 3,
4 (1782).
70.
Account of a Collection of Rhopalocera. 189
Byblia gotzvus.
Pap. gotzius, Herbst, Naturs. Schm. ix, p. 193, t. 258,
f. 3, 4 (1798),
71. Crenis amulia.
76.
ie
78.
19.
80.
81.
85.
Pap. amulia, Cram., Pap. Exot. 11, p. 128, t. 180, f. C
(1777).
. Cyrestis camallus.
Pap. camillus, Fabr., Spec. Ins. u, p. 11 (1781).
. Neptis nemetes,
N. nemetes, Hew., Exot. Butt. Neptis, t. 1, f. 1, 2
(1868).
. Neptis agatha.
Pap. agatha, Stoll, in Cramer’s Pap. Exot. iv, p. 76,
te o2te ts Ay io Olir30):
. Neptis melicerta.
Pap. melicerta, Dru., Ill. Exot. Ins. ui, p. 34, t. 19,
f. 3, 4 1773).
Pseudacrexa lucretia.
Pap. lucretia, Cram., Pap. Fixot. i, p. 71, t. 45, f. C, D
GLAS):
Pseudacrxa boisduvalit.
Diadema boisduvalii, Doubld., Ann. N. H. xvi, p. 180
(1845).
Pseudacrea ewrytus.
Pap. ewrytus, Linn., Syst. Nat., ed. 10, p. 487 (1758),
Pseudacrea striata.
P. striata, Butl., Cist. Ent. i, p. 215 (1874).
Pseudacrva semire.
Pap. semire, Cram., Pap. Exot. iii, p. 3, t. 194, f B, C
Giana):
Pseudoneptis cenobita.
Hesperia cenobita, Fabr., Ent. Syst. ui, 1, p. 247
: (1798).
. Catuna angustata.
Euomma angustatum, Feld., Reise Novar. Lep., p. 425
(1867).
Aterica galene.
Pap. galene, Brown, New Ill. of Zool., p. 94, t. 37
(1776).
TRANS. ENT. SOC. LOND. 1903.—PART I. (JUNE) 18
190 Mr. P. I. Lathy’s
84. Hamanumida dedalus.
Pay. dxdalus, Fabr., Syst. Ent., p. 482 (1775).
5. BKuphedra ruspina.
Romaleosoma ruspina, Hew., Exot. Butt. Romalzo-
soma, t22, 1-16, 4 (S05):
86. Luphedra eleus.
Pap. eleus, Dru., Ill. Exot. Ins. ii, p. 14, t..12, f. 1, 2
(1782).
87. Huphedra cers.
Pap. ceres, Fabr., Syst. Ent., p. 504 (1775).
88. Luphedra nigrocilia, sp. nov. (Plate VIII, fig. 1.)
Upper-side.—Fore-wing blue-black, shot with greenish along costa
and about apex ; a large oval, greenish-yellow patch beyond cell,
between upper discoidal and upper median nervules; a large tri-
angular yellowish-green patch on innermargin. Hind-wing shining
blue, with yellowish-green discoidal patch ; outer margin black,
widely so at apex ; two large black spots near margin between
upper and lower median nervules.
Under-side.—Fore-wing greenish-yellow, inner margin slate colour,
outer margin and apex greenish ; the following black markings: a
spot in cell near base, two larger spots beyond, and a bar at end of
cell, with another bar outside ; a spot between upper and middle
median nervules, and an obliquely placed bar between middle and
lower median nervules ; a submarginal row of large spots increasing
in size towards anal angle. Hind-wing greenish-yellow, deep yellow
at anal angle and inner margin, greenish on outer margin, and with
the following black markings: a curved sub-basal band extending
nearly half-way along costa, a round spot in upper part of cell, and
an oval spot at end of cell; a series of four patches between sub-
costal nervure, and submedian nervule, decreasing in size towards
latter, a submarginal series of large patches, Cilia of both wings
black above and below.
Exp. 78 millim.
Allied to #. ceves, Cram., from which it differs in the
fore-wing having a subapical patch instead of a band, the
shining blue ground colour of hind-wing, and the black
cilia.
89, Huphedra thems.
Najas hilaris themis, Hiibn., Exot. Schm. 1, t. 60
(1806-16).
Account of a Collection of Rhopalocera. 191
90. Huphedra janetta.
fomalevsoma janetta, Butl., P. Z.S. 1871, p. 80.
91. Huphedra aurata.
E. aurata, Carpenter, Proc. R. Dubl. Soe, (2) viii, p. 305
(1895).
92. Huphedra aureofasciata, sp. nov.
Upper-side.—Fore-wing black, costa shot with green; a wide golden-
yellow band extending from costa to below middle median nervule ;
a triangular bluish-green patch on basal half of inner margin, the
centre of this patch golden. Hind-wing shining bluish-green, outer
margin widely black, a faint trace of golden-yellow on disco-
cellulars.
Under-side.—Fore-wing yellowish-green, with inner margin black
below lower median nervule, and the following black markings : two
large spots in basal half of cell, each touching median nervure ; two
round spots in centre of, and a bar at end of cell ; a series of five
patches beyond cell, of which the lowest is the largest, and obliquely
placed, a submarginal series of square patches; outer margin
blackish. Hind-wing golden-yellow, greenish about cell, and along
outer margin; a wide crimson band along costa extending from
base to near apex ; the following black markings: a large spot in
upper part of cell with a small one below it, a bar at end of cell,
four markings beyond cell, of which the upper and lower are nearer
the base than the others ; the upper is the largest, and the lower
much the smallest of this series; a submarginal series of large
patches, the inner edges of which are dentate ; outer margin black.
Cilia of hind-wing greyish between nervules.
Exp. 90 millim.
This is probably a local race of £. crockeri, Butl., from
which it may be distinguished by the bright golden-yellow
band of the fore-wing and the darker cilia.
Only a single female was obtained, but there are several
specimens in the Natural History Museum from Nigeria
which agree with that here described.
93. Huphedra medon.
Pap. medon, L., Cent. Ins., f. 19 (1763).
94. Huphedra harpalyce.
Pap. harpalycec, Cram., Pap. Exot. ii, p. 78, t. 145,
f DPE Gn):
192
95.
Mr. P. I. Lathy’s
-Huphedra losinga.
Romalexosoma losinga, Hew., Exot. Butt. Romalzo-
soma, t. 1, f. 1 (1864).
One male and two females were taken.
96.
Oi
Huryphene theognis.
E. theognis, Hew., Exot. Butt. Euryphene, t. 1, f. 3, 4
(1864).
Huryphene mardania.
Pap. mardania, Fabr., Ent. Syst. ii, 1, p. 249 (1793).
furyphene sophus.
Pap. sophus, Fabr., Ent. Syst. 1, 1, p. 46 (1793).
. Huryphene letitia.
HE. lxtitia, Plotz, 8. F. Z. xli, p. 192 (1880).
. Luryphene tentyrrs.
E. tentyris, Hew., Exot. Butt. Euryphene, t. 5, f. 21,
22 (1866).
. Diestogyna veronieda.
Pup. veronica, Cram., Pap. Exot. iv, p. 73, t. 325, f. C,
D (1789).
2, Huryphura chaleis.
Harma chalcis, Feld., Wien. Ent. Mon. iv, p. 284
(1860).
. Luryphura plautilla.
Huryphene plautilla, Hew., Exot. Butt. Euryphene,
t. 3, f 14, 15 (1865).
One male has both wings fulvous to beyond middle ;
the form claudianus, Druce, is also fulvous, and has
hitherto been regarded as a female aberration; it is
possible that this specimen is the male of claudianus,
and a distinct species.
104.
Cymothoé theobene.
Harma theobene, D. and H., Gen. D. Lep., p. 288,
t. 40, f. 3 (1850).
. Cymothoێ egesta.
Pap. egesta, Cram., Pap. Exot. i, p. 72, t. 46, f. B, C
(1775).
. Cymothoé becker.
Diademi beckeri, H. S., Aus. Schm. Tagf,, f. 81 (1850).
Account of a Collection of Rhopalocera. 193
107. Cymothoé coccinata.
Harma coccinata, Hew., Exot. Butt. Harma, t. 6,
f, 24, 15 (1874).
108. Huptera svrene.
E. sirene, Staud., Iris, iv, p. 100, t. 1, f. 6 (1891).
A single specimen of this extremely rare species.
109. Huxanthe eurinome.
Pap. ewrinome, Cram., Pap. Exot. 1, p. 109, t. 70, f.
A (1775).
An aberration of the female was obtained, in which the
white markings are more extended, and the black ground
along median nervure and nervules powdered with white
scaling.
110. Charaxes brutus.
Pap. brutus, Cram., Pap. Exot. in, p. 82, t. 241, f.
Dp alice)
111. Charaxes epyasius.
C. epyasius, Reiche, in Ferret and Galin. Voy.
Abyss. Ent., p. 469, t. 32, f. 1, 2 (1849).
Only one specimen in the collection, a female, and
unfortunately in very bad condition.
112. Charaxes castor.
Pap. castor, Cram., Pap. Exot. i, p. 61, t. 37, f. C, D
(1775).
113. Charaxes etesipe,
Nymphalis etesipe, Godt., Enc. Méth. ix, p. 355 (1823).
114, Charazxes cynthia.
C. cynthia, Butl., P. Z. 8. 1865, p. 626, t. 36, f. 3.
The two males received agree best with examples from
Accra, inasmuch as the discal band of hind-wing above is
buff yellow, edged with orange rufous, but differs below in
the much wider discal silvery band. The female agrees
best with a specimen from the Cameroons.
115, Charaxes protoclea.
C. protoclea, Feisth, Ann. E. Fr, (2) viii, p. 260 (1850).
116. Charaxes anticlea.
Pap. anticlea, Dru., Ul. Exot. Ins. i, p. 36, t. 27, f.
5, 6 (1782).
Two males only, they belong to the typical form, not
that described by Rothschild under the name of adwsta.
194 Mr. P. I. Lathy’s
117. Charaxes etheocles.
Pap. etheocles, Cram., Pap. Exot. 11, p. 34, t. 119, f.
IDO CUETO
118. Charaxes tiridates.
Pap. tiridates, Cram., Pap. Exot. ii, p. 100, t. 161, f.
A, B (1777).
119. Charaxes numenes.
C. numenes, Hew., Exot. Butt. Charaxes, t. 2, f. 9-11
(1859).
120. Charaxes varanes.
Pap. varanes, Cram., Pap. Exot. ii, p. 100, t. 160, f.
IDE Glenroy
121. Charazxes fulvescens.
Palla varanes, var. fulvescens, Auriv., Ent. Tidskr. xi,
p. 216 (1891).
122. Charaxes laodice.
Pap. laodice, Dru., Ill. Exot. Ins. in, p. 34, t. 26, f 1,
ZaClio2):
123. Charaxes ewpale.
Pap. eupale, Dru., Exot. Ins, ui, p. 7, t. 6, f. 3 (1782).
124. Monuwra zingha.
Pap. zingha, Cram., Pap. Exot. iv, p. 53, t. 315, f.
BCrgi7s0).
125. Libythea labdaca.
L. labdaca, Westw., Gen. D. Lep., p. 413 note, t. 68,
f, 6 (1851).
126. Abisara gerontes.
Pap. gerontes, Fabr., Spec. Ins. u1, p. 117 (1781).
127. Telipna actinotina, sp. nov. (Plate VIII, fig. 2.)
s
Upper-side.—Fore-wing dark brown, basal half brick-red, darker
towards base and along costa, and bordered on outer edge by a
blackish-brown band; two small red patches beyond cell. Hind-
wing semi-transparent brown, veins blackish.
Under-side.—Fore-wing as above, but ground colour paler and with
a yellowish band beyond dark border of basal red. Hind-wing
yellowish-brown, with veins and lines in interspaces blackish ; seven
small black spots at base.
Exp. 48 millim.
Only a single specimen of this very distinct species was
obtained ; its nearest ally appears to be 7’. parva, Kirby,
Account of a Collection of Rhopalocera. 195
from which it may at once be distinguished by its much
greater size. The neuration of hind-wing resembles that,
of the genus Mimacrexa, inasmuch as the subcostal nervules
are on a long stalk, but the presence of the preecostal
nervure serves to separate it from that genus. It is very
curious that this species should bear such a strong super-
ficial resemblance to Actinote, a South American genus of
the Acreeine.
128. Pentila pauli.
P. pault, Staud., Exot. Schm. i, p. 267 (1888).
129. Pentila amenaida. =
P. amenaida, Hew., Exot. Butt. Pentila and Liptena,
tae Qe Os LSS):
130. Pentila radiata, sp. nov. (Plate VIII, fig. 3.
Upper-side.—Fore-wing fulvous, with broad black margin, the
inner edge of which is dentate towards inner margin ; costa blackish ;
two small black spots above and towards end of cell, and a large
round black spot at end of cell. Hind-wing fulvous, paler at inner
margin and widely bordered with black ; a round black spot at end
of cell.
Under-side.—Fore-wing fulvous, with black spot as above, outer
margin narrowly black, marginal black streaks in interspaces, de-
creasing in length towards anal angle. Hind-wing fulvous, outer
margin narrowly black with marginal streaks as in fore-wing, two
very small black spots near base, a large round black spot below costal
nervure not far from base, and a similar but somewhat larger spot at
end of cell.
Exp. 38 millim,
Nearly allied to P. pauli, Ster., from which it differs in
the total absence of discal spots below and in the sub-
costal spot of the hind-wing below being larger than the
cellular spot.
131. Pentila multipunctata, sp. nov. (Plate VIII, fig. 4.)
Upper-side-Fore-wing fulvous, with broad black margin, the
inner edge of which is dentate ; costa blackish ; the following black
markings ; a small spot above and towards end of cell, and a large
round spot at end of cell, three spots near marginal border between
upper median nervule and submedian nervure. Hind-wing fulvous,
with wide black margin, inner edge of which is dentate ; a round
black spot at end of cell and a series of seven black spots near
196 Mr. P. I. Lathy’s
marginal border, the three nearest anal angle being further from the
border than the others.
Under-side.—Fore-wing fulvous, outer margin narrowly black,
with marginal streaks as in the preceding species, and the following
black markings: two pairs and one single spot on costa, a large
round spot at end of cell anda curved discal series of seven spots,
Hind-wing fulvous, outer margin narrowly black, and streaks as in
fore-wing, with the following black markings: a large spot above
and two spots below cell near base, a large spot at end of cell, a
curved discal series of eight spots, of which that next costa is much
the largest.
Exp. 42 millim.
Near to P. pauli, Stgr., from which it may be distin-
guished by the conspicuous discal spots.
132. Pentila abraxas.
Peabracas, Dy sand Gen DP epaat.. iat
(1852).
133. Iimacrea darwinia.
M. darwinia, Butl., Lep. Exot., p. 104, t. 38, f. 8
(1872).
One pair of this fine species.
134, Pseuderesia isca.
Liptena ica, Hew., Exot. Butt. Pentila and Liptena,
t. 2, f. 14-16 (1873).
135. Pseuderesia gordoni. (Plate VITI, fig. 5.)
P. gordoni, Druce, Ann. Mag. Nat. Hist. (7) xi
(Jan. 1903).
136. Pseuderesia cellularis.,
P. cellularis, Kirby, Ann. N. H. (6) vi, p. 262 (Sept.
1890).
137. Citrinophila similis.
C. similis, Kirby, Ann. N. H. (5) xix, p. 366 (1887).
138. Liptena submacula, sp. nov. (Plate VIII, fig. 6.)
Upper-side.—Fore-wing white, costa and outer margin bordered
with dark brown. Hind-wing white, outer margin bordered with
dark brown.
Under-side.—Both wings as above, with exception of an oval black
spot at end of cell of hind-wing.
Exp. 32 millim.
This species may be distinguished from JL. simplicia,
Account of a Collection of Rhopalocera. 197
Moschl., its nearest ally, by the black spot at end of cell of
hind-wing below.
139. Liptena undularis.
L. undularis, Hew., Exot. Butt. Pentila and Liptena,
t. 1, f. 7 (1866).
140. Liptena otlauqa.
L. otlauga, Smith and Kirby, Rhop. Exot. 14, Lyczen.
Afr., p. 46, t. 11, f. 9, 10 (1890).
141. Hresina corynetes.
E. corynetes, Smith and Kirby, Rhop. Exot., Lyceen.
Atiry a Ae te Itt. 7, 8 L820).
142. Aslauga marginata.
Liptena marginata, Plotz, 8. E. Z. xli, p. 204 (1880).
143. Deudorix nomion.
D. nomion, Ster., Iris, iv, p. 158, t. 1, f. 11 (1891).
144. Deudorix eleala.
Hypolycena eleala, Hew., Ml. D. Lep., p. 52, t. 28, f.
25-27 (1865).
145. Deudorix camerona.
D. camerona, Plotz, 8. E.Z. xli, p. 201 (1880).
146. Deudoriz odana.
D. odana, H. H. Druce, Ent. M. Mag. xxiii, p. 204
(1887).
147. Deudorix caliginosa, sp. nov. (Plate VIII, fig. 7.)
Upper-side.—Both wings black, shot with rich purplish-blue.
Hind-wing with inner margin clothed with black hair ; a pale shin-
ing purplish sex mark above cell ; inner margin pale shining grey.
Under-side.—Fore-wing brownish-grey, with the following white
lines inwardly edged with brown: two at end of cell, a double row
beyond cell extending from costa to lower median nervule, and a
submarginal double row, these being obscure at apex ; a large tuft
of black hair from middle of inner margin. Hind-wing brownish-
grey, a pale-edged dark-brown spot in bend of subcostal nervure,
two whitish lines just beyond cell, and some obscure whitish mark-
ings between cell and inner margin, two double rows of whitish
lines as in fore-wing ; a black spot inwardly bordered with red near
inner margin, between middle and lower median nervules; anal
angle produced into a lobe, black-edged with metallic silvery-blue
scales ; lower median nervule produced into a long slender black
tail tipped with white.
Exp. 34 millim.
198 Mr. P. 1. Lathy's
The nearest ally of this species appears to be D. antalus,
Hopff.; its much darker colour at once serves to dis-
tinguish it from that species; the tuft of hair on inner
margin of the fore-wing below is much longer in D. livida
than in D. antalus.
148. Deudorix bimaculata.
Myrina bimaculata, Hew., Trans. Ent. Soc. 1874, p.
353.
149. Deudoria lorisona.
Myrina lorisona, Hew., Ill. D. Lep., p. 37, t. 16, f 48,
49 (1863).
150. Deudoriz antalus.
Dipsas antalus, Hopff., Monatsb, Akad. Wiss. Berlin,
1855, p. 641.
151. Myrina silenus.
Pap, silenus, Fabr., Syst. Ent., p. 531 (1775).
152. Myrina subornata, sp. nov. (Plate VIII, fig. 8.)
Upper-side.—Fore-wing black, a brilliant deep blue patch in cell,
and a large area of the same colour below cell. Hind-wing black,
cell and area below and beyond brilliant deep blue ; a deep crimson
spot at anal angle ; submedian nervure produced into a moderately
long white centred black tail.
Under-side.—Fore-wing dark brownish-grey, with a faint purple
tinge, paler towards inner margin. Hind-wing dark brownish-grey,
with a faint purple tinge; a black bar at end of cell; a curved
discal series of irregular black markings, those next inner margin
being inwardly edged with whitish ; submarginal whitish powdered
black marking extending from inner margin to middle median
nervule ; anal lobe deep crimson inwardly edged with white.
Exp. 38 millim.
This species may be separated from dL dermaptera,
Wer., a South African species, by the discal black mark-
ings of hind-wings below.
153. Oxylides faunus.
Pap. faunus, Dru., Il. Exot. Ins. ui, p. 2, t. 1, f. 4, 5
(17738).
154. Hypolycena hatita.
Avhatita,Hew., Ul,.D> Wep., p: 51, 23, a. 21-24
(1865).
Account of a Collection of Rhopalocera. 199
155. Hypolycena antifaunus.
H. antifaunus, D. and H., Gen. D. Lep., t. 75, f. 1
(1852).
156. Hypolycena lebona.
HT, lebona, Hew., Ill. D. Lep., p. 51 (1865).
157. Hypolycena philippus.
Hesperia philippus, Fabr., Ent. Syst. i, 1, p. 288
(1793).
158. Dapidodigma hymen.
Pap. hymen, Fabr., Syst. Ent., p. 519 (1775).
159. Jolaus adamsi, sp. nov. (Plate VIII, fig. 9.)
Upper-side.—Fore-wing with basal half bright shining blue, apical
half black. Hind-wing bright shining blue, costa and apex narrowly
black, a black line along outer margin ; anal lobe white, a black
spot edged with dull crimson placed above it on inner margin, and a
black spot beyond it on outer margin, and beyond this a faint whitish
mark ; three slender black-centred white tails, that at the anal angle
being much the longest ; cilia whitish,
Under-side.—Fore-wing white, brownish towards outer margin,
blackish along inner margin, a faint brown submarginal line ; a
conspicuous tuft of black hair on inner margin, Hind-wing white,
a faint brown discal line extending from subcostal nervure to middle
median nervule ; a black spot with a red patch above it between
middle and lower median nervules, the discal line continuing
brokenly from it to inner margin; a short faint submarginal red
line from near apex ; a black spot on inner margin above anal lobe,
edged with a triangular red patch ; a little blackish sealing near outer
margin between anal tails ; a narrow black marginal line.
Exp. 42 millim.
Allied to J. laon, Hew., but differs in the absence of
dark borders to wings below.
160. Jolaus ewrisus.
Pap. ewrisus, Cram., Pap. Exot. ii, p. 47, t. 221, f. D,
Ean):
161. Aphneus orcas.
Pap. orcas, Dru., Ill. Exot. Ins. ii, p. 46, t. 34, f. 2, 3
(1782).
162. Aphneus brahamni, sp. nov. (Plate VIII, fig. 10.)
Upper-side.—Fore-wing dark brownish-grey, cell and area beyond
and below densely clothed with purplish-blue sealing ; outer margin
narrowly blackish. Hind-wing dark brownish-grey, cell and lower
200 Mr. P. I. Lathy’s
discal area densely clothed with purplish-blue scaling, outer margin
narrowly blackish, cilia white except at extremities of nervules,
where they are blackish ; a short black tail at extremity of lower
median nervule, and a very long slender black tail at extremity of
submedian nervure, the basal third of this tail is dull red and the
tip white.
Under-side.—Fore-wing brownish-grey, whitish along inner margin,
with the following silver markings all edged with deep dull crimson :
a spot at base of cell with one above it on costa, one crossing centre
of cell and a large one at end of cell, another on costa beyond cell,
and between this and apex two small spots one above the other, a
round spot on discoidals not far from outer margin, a figure-of-eight
spot on median nervules, and an obliquely-placed oval patch below
this, the lower part of this marking being obscure; outer margin
narrowly edged with deep dull crimson. Hind-wing brownish-grey,
with the following silver markings all edged with deep dull crimson :
a spot at base, a small spot in basal half of cell with a large one
above it, a large round spot at end of cell, an oval patch between
this and inner margin, and a small spot on inner margin; an
irregular series of seven discal spots, of which that on the costa is
large and almost round, the next three very small, and the remainder
long and irregularly shaped ; an obscure narrow dark band between
this series and outer margin ; outer margin narrowly edged with
deep dull crimson; anal angle the same colour ; tails as above.
Exp. 48 millim.
Very near to A. argyrocyclus, Holl, from which it differs in
the large circular spot at end of cell in hind-wing below, and
the more regular and complete series of silver spots beyond.
I have named this species after Mr. Braham the collector.
163. Spindasis mozambica.
Aphneus mozambica, Bertolini, Mem. Acad. Bologna
2, p. 177, sep., p. 13 (1851).
164, Awiocerses harpax.
Pap. harpax, Fabr., Synt. Ent., p. 829 (1775).
165. Lycenesthes sylvanus.
Pap. sylvanus, Dru., Ill. Exot, Ins. ii, p. 5, t. 3, f. 2, 3
(1773).
166. Lycenesthes larydas,
Pap. larydas, Cram., Pap. Exot. iii, p. 160, t. 282, f. H
(1780).
167. Lycenesthes flavomaculata.
L. flavomaculata, Smith and Kirby, Rhop. Exot.
xxvi, Lycen. Afr. p. 104, t. 23, £7, 73 (1893).
168.
169.
170.
ale
178.
io
180.
181.
Account of a Collection of Rhopalocera. 201
Cupido falkensteinie.
C. falkensteinii, Dewitz, N. Acta Ac. N. Cur. xli, 2,
p. 204, t. 25, f. 5 (1879).
Cupido lingeus.
Pap. lingeus, Cram., Pap. Exot. iv, p. 176, t. 379, f.
KE alyteslL)
Cupido plinvus.
Hesperia plinius, Fabr., Ent, Syst. 11, 1, p. 284 (1793).
Cupido isis.
Pap. isis, Dru., Ill. Exot. Ins. i, p. 6, t. 3, f. 4, 5
(1773).
. Cupido mirza.
C. mirza, Plotz, 8. F. Z. xli, p. 203 (1880).
. Cupido beticus.
Pap. beticus, L., Syst. Nat. ed. 12, p. 789 (1767).
. Cupido malathana.
Lycena malathana, Boisd., Faune Madag., p. 25
(1833).
. Cupido osiris.
Lycena osiris, Hopft., Monatsh. Akad. Wiss. Berlin,
1855, p. 642.
}. Cupido micyclus.
Pap. micyclus, Cram., Pap. Exot. i, p. 160, t. 282, f.
F, G (1780).
. Cupido punctatus.
C, punctatus, Dewitz, N. Acta Acad. N. Cur. xli, 2,
p. 205, t. 26, f. 15 (1879).
Leptosia medusa.
Pap. medusa, Cram., Pap, Exot. 11, p. 86, t. 150, f. F
(1777).
Leptosia alcesta.
Pap. alcesta, Cram., Pap. Exot. iv, p. 175, t. 379, f. A
(1781).
Mylothris chloris.
Pap. chloris, Fabr., Syst. Ent., p. 473 (1775).
Mylothris sprea,
M. spica, Moschl., Verh. z. b. Ges. Wien, xxxiil, p. 277
(1883).
Mr. P. I. Lathy’s
2. Appias rhodope.
Pap. rhodope, Fabr., Syst. Ent., p. 478 (1775).
. Appias phuola.
Pieris phaola, Doubld., Ann. N. H. (1) xx, p. 63 (1847).
. Appias epaphia.
Pap. epaphia, Cram., Pap. Exot. iti, p. 26, t. 207, f. D,
E (1779).
5, Pieris creona
Pap. creona, Cram., Pap. Exot. i, p. 148, t. 95, £.C-F
(1776).
. Pieris calypso.
Pap. calypso, Dru., Ill. Exot. Ins. u, p. 30, t. 17, f. 3, 4
(1773).
. Pierrs cebron.
P. cebron, Ward, Ent. Mo. Mag. vii, p. 59 (1871).
. Teracolus evippe.
Pap. evippe, L., Syst. Nat. ed. 10, p. 469 (1758).
. Catopsilia florella,
Pap. florella, Fabr., Syst. Ent., p. 479 (1775).
. Lerias brenda.
ZT. brenda, Wand H., Gen. De iep. pi 19, t20, 126
(1847).
. Terias senegalensis.
T. senegalensis, Boisd., Spec. Gén. Lép. i, p. 672
(1836).
. Terias desjardinsi.
NXanthidia desjardinsii, Boisd., Faune Mad., p. 22, t.
2,4. 6 (1833):
. Papilio dardanus.
P. durdanus, Brown, Ill. Zool., p. 52, t. 22 (1776).
Typical female and one example of @ ab. dionysus, D.
and H., were obtained.
194.
NO:
196.
Papilio cynorta.
P. cynorta, Fabr., Ent. Syst. i, 1, p. 37 (1798).
Papilio nireus.
P. nireus, L., Syst. Nat. ed. 10, p. 464 (1758).
Papilio demodocus.
P. demodocus, Esp., Aus. Schm., p. 205, t. 51, fi 1
(1798).
Account of a Collection af Rhopalocera. 203
197. Papilio menestheus.
P. menestheus, Dru., Ill. Exot. Ins, 1, p. 15, t. 9, f. 1,
2 (1773).
198. Papilio ridleyanus.
P. ridleyanus, White, Ann. N. H. xi, p. 262, fig.
(1843).
A good series, including an interesting aberration in
which area above subcostal nervure is entirely white and
without black markings with the exception of marginal
border; the nervules “and interspaces, excepting those
Reena median nervules, are powdered with white scales ;
the inner margin is also much whiter than in typical
specimens ; these remarks apply only to the hind-wing, the
fore-wing being normal.
199. Papilio pylades.
P. pylades, Fabr., Ent. Syst. ii, 1, p. 34 (1793).
200. Papilio leonidas.
P. leonidas, Fabr., Ent. Syst, i, 1, p. 35 (1793).
A large number of males, but only one female.
Z01. Papilio policenes.
P. policenes, Cram., Pap. Exot. i, p. 61, t. 37, f. A, B
(1775).
202. Sarangesa synestalinenus.
Antigonus synestalmenus, Karsch, Berl. Ent. Zeit.
XXXVili, p. 263, t. 6, f 8 (1898).
203. Tagiades flesus.
Pap. flesus, Fabr., Spec. Ins. ii, p. 185 (1871).
204, Hagris denuba,
Antigonus denuba, Plotz, 8. KE. Z. xl, p. 361 (1869).
205. Abantis elegantula.
A. elegantula, Mab., Ann. Soc. Ent. France, 1890,
p. 32.
206. Hesperia ploctzi.
Hi. ploctzi, Auriv., Ent, Tidsk., 1891, p. 277.
207. Gorgyra mocquerysit.
G. mocquerysti, Holl, P. ZS, 1896, p. 33, t. 5,
f 10.
208. Oxypalpus fulvus, sp. nov. (Plate VIII, fig. 11.)
204 Mr. P. I. Lathy’s
Upper-side.—Fore-wing deep fulvous, clouded with dark brown at
base ; apex and outer margin narrowly edged with dark brown; a
blackish line along submedian nervure, and an oblong black patch
beyond cell. Hind-wing deep fulvous ; costa, a narrow outer marginal
line, and an oblong patch beyond cell black, inner margin blackish ;
the veins of both wings black towards extremities, cilia deep fulvous.
Under-side.—Fore-wing deep fulvous, a large black patch on basal
third of inner margin; a short black fascia, above - submedian
nervure, from this patch ; a narrow black marginal line. Hind-wing
deep fulvous, with a narrow black marginal line. Antennz beneath
fulvous.
Exp. 28 millim.
Allied to O. ignita, Mab., but exhibits many points of
difference, the chief of which are the smaller area of black
on the fore-wing above, the exterior part of inner margin of
fore-wing below being fulvous, and the antenne fulvous
beneath.
209. Oxypalpus annulifer.
O. annulifer, Holl. Ann. N. H. Oct. 1892, p. 298.
210. Osmodes distincta.
O. distincta, Holl., P. Z. 8., 1896, p. 43, t. 4, f. 16.
211. Hypoleucis ophiusa.
Hesperia ophiusa, Hew., Trans. Ent. Soc. Lond. (3) i,
p. 497 (1866).
212. Cyclopides abjecta.
C. aljecta, Snell., Tid. voor Ent. 1872, p. 52, t. 2,
eur I Ky
213. Chapra mathias.
Hesperia mathias, Fabr.. Ent. Syst. Suppl, p. 438
(1798).
214. Parnara micans.
Py micans, Holl, RP. Z.S:, 1896) p. 63st. 3, 1.19,
215. Baoris alberta.
B. alberit, Holl., P. Z. S., 1896,-p. 67, t. 2, f. 21.
216. Baoris ogrugana, sp. nov. (Plate VIII, fig. 12.)
Upper-side.—Fore-wing dark brown, with the following hyaline
markings : two spots at end of cell, three very small subapical spots,
a series of four discal spots of which the two centre ones are much
the largest, and the upper one is much nearer the outer margin than
Account of a Collection of Rhopalocera. 205
the others. Hind-wing dark brown, with a hyaline spot in cell, and
two very small discal hyaline spots.
Under-side.—Fore-wing dark brown, a short pale submarginal line
from apex, hyaline markings as above, the lowest of the discal series
being bordered outwardly by an oblong whitish patch. Hind-wing
dark brown, with obscure lilac-grey markings on discoidals and near
outer margin ; hyaline markings as above.
Exp. 32 millim,
Near P. wnistriga, Holl., but differs in possessing three
subapical spots, the upper spot of the discal series being
much nearer the outer margin.
217. Baoris netopha.
Hesperia netopha, Hew., Ann. N. H. (5) 1, p. 345
(1878).
218. Platylesches chameleon.
Pamphila chameleon, Mab., C. R. Soc. Ent. Belg. xxxv,
pa ee9 Gisgih):
219. Pardaleodes incerta.
Pamphila incerta, Snell., Tijd. voor Ent. 1872, p. 29,
t. 10, f. 10-12.
220. Pardaleodes reichenowt.
Plastingia reichenowi, Plotz, 8. EH. Z. xl, p. 357 (1879).
221. Pteroteinon laufella.
Hesperia laufella, Hew., Exot. Butt. Hesp., t. 2, f.
28-30 (1867).
222. Ploetzia cerymrea.
Hesperia cerymica, Hew., Exot. Butt. Hesp., t. 2, f.
20, 21 (1867).
223. Ploetzia nobilior.
Panobvilior, Holl P. Zas: 1890; pagouts 9,0. :2.
224. Rhopalocampta forestan.
Pap. forestan, Cram., Pap. Exot. iv, t. 391, fi EH, F
(1782).
225. Rhopalocampta chalybe.
Ismene chalybe, Westw., D.and H. Gen. D. Lep., t. 79,
f. 2 (1852).
226, Lhopalocampta iphis.
Pap. iphis, Dru., Ill, Exot. Ins. ii, t. 15, f. 3, 4 (1773).
TRANS. ENT. SOC. LOND. 1908.—PART Il. (JUNE) 14
206 Account of a Collection of Rhopalocera.
The following is a list of the species not obtained by
Mr. Braham, but which are recorded in the two previously-
mentioned papers :—
Monotrichtis desolata, Butl.
“pthima itonia, Hew.
Acrea cecilia, Fabr.
Atella columbina, Cram.
Precis cebrene, Trim.
Euryphene phranza, Hew.
Charaxes achemenes, Feld.
Lachnocnema bibulus, Fabr.
Zeritis neriene, Boisd.
Deudoriz nomenia, Hew.
Lycenesthes princeps, Butl.*
Cupido hippocrates, Fabr.
» lysimon, Hiibn.
Terias brigitta, Cram.
Eronia argia, Fabr.
Celenorrhinus galenus, Fabr.
Parnara borbonica, Boisd.
* It is probable that this species was not correctly identified, the
type being from Abyssinia.
EXPLANATION OF PuaTteE VIII.
Fig. 1. Euphedra nigrocilia, sp. nov.
2. Telipna actinotina, sp. nov.
3. Pentila radiata, sp. nov.
4. » multipunctata, sp. nov.
5. Pseuderesia gordon, Druce.
6. Liptena submacula, sp. nov.
7. Deudorix caliginosa, sp. nov.
8. Myrina subornata, sp. nov.
9. Jolaus adamst, sp. nov.
10. Aphneus brahami, sp. nov.
11. Oxypalpus fulvus, sp. nov.
12. Baoris ogrugana, sp. Nov.
XII. Hymenoptera aculeata, collected by the Rev. ALFRED
EK. Earon, M.A., in Madeira and Tenerife, in the
spring of 1902, including notes on species taken
by the late T. VERNON WoLLASTON and F. A.
BELLAMY. By Epwarp Saunpers, F.RS.,
F.L.S., ete.
[Read March 18th, 1903.]
In the little collection enumerated below, made by Mr.
Eaton in the above islands, there are several species of
interest. I have described four which appear to be new,
and besides these two or three will probably prove so when
more material comes to hand. A curious variety of Bombus
terrestris, L., is worthy of special notice, it is quite black
with the exception of the apex of the abdomen which is
snowy white. Mr. Eaton captured three males and two
workers, and remarks that the species is common in
Tenerife, and as he sent no ordinarily-coloured specimens, I
presume that this is the usual form in the island. Brullé
does not mention ¢errestris as a Canary Island species,
soroensis being the only representative of the genus which
he records, and I cannot help having a suspicion that he
has mistaken this black var. of terrestris for the similarly
coloured form of soroensis which is well known on the
Continent. So little is known of the Hymenoptera of
these islands that I thought the present communication
might be acceptable to the Society.
Since this paper was read, I have had placed in my
hands by Prof. Poulton two collections from the Hope
Department of the University of Oxford—one from Madeira
made in 1847 and following years by the late T. V.
Wollaston, and one made last year at Tenerife by Mr.
F. A. Bellamy ; as these contain several species not in-
cluded in my paper, I have, at Prof. Poulton’s suggestion,
incorporated them.
Wollaston’s collection was purchased in 1861 by the Rev.
F. W. Hope, and presented to the Oxford University
Museum. The specimens are of course old and more or less
faded, the smaller ones are often gummed on cards, which
TRANS. ENT. SOC. LOND, 1903.—PART II. (JUNE)
208 Mr. Edward Saunders on Hymenoptera aculeata
renders determination doubtful, and as it would be dan-
gerous to remove specimens of this age I have not at-
tempted to found any new species on them. At the same
time there is a bronzy Halictus which I think will prove
to be new, and also a 2 Prosopis entirely black with. the
exception of the yellow facial spots; it is larger than the
species described here and stands in the collection with a
label under it in F. Smith’s handwriting—n. sp.? allied to
signata. It is represented by a single specimen only and
is not referable to any species I know.
In spite of the poor condition of Wollaston’s specimens
it is important to record, as far as possible, the forms
which existed over half a century ago in an island so
liable to accidental immigration as Madeira.
Professor Forel has kindly examined the ants of the
collection and verified those already determined by Mr.
F. Smith. I am indebted also to Mons. J. Vachal, who
has recently examined Brullé’s Canary Island types in the
Paris Museum, for confirming, and in some cases correcting
my identifications of the Halicti.
In Mr. Bellamy’s collection is a fine series of the dark
form of terrestris mentioned above, including six females, all
coloured exactly like those met with by Mr. Eaton. As
all the specimens of both collections came from Tenerife,
it would be interesting to ascertain if the species has the
same coloration in the neighbouring islands.
In the following list Mr. Bellamy’s captures are indicated
by his name, the late T, V. Wollaston’s by his initials,
while all specimens not specially indicated were taken by
Mr, Eaton.
CAMPONOTUS RUFOGLAUCUS, Jerd., r. micans, Ny].
3 7. Orotava, Tenerife. F, A. Bellamy, 31. iii.
COs ven:
LASIUS NIGER, L.
229. Many of each. Madeira. T. V. W.
TAPINOMA ERRATICUM, Ltr.
$28. Several of each. Madeira. T. V. W.
PLAGIOLEPIS PYGMHA, Ltr.
2%. Numerous. Madeira. T. V. W.
collected in Madeira and Tenerife 209
PONERA CONTRACTA, Ltr.
On? O08 /Madeiras i We
TETRAMORIUM SIMILLIMUM, Nyl.
$2. Madeira, T. V. W.
LEPTOTHORAX UNIFASCIATA, Ltr.
S4, Madeira. T. V. W.
MoNoMORIUM SALOMONIS, r. subopacum, Smith.
2%. Numerous. Madeira, T. V. W.
MoNOMORIUM CARBONARIUM, Smith.
22045 52 Madeira, 1. V.. W.
PHEIDOLE MEGACEPHALA, Fab.
98. Several. Madeira. T. V. W.
TACHYSPHEX SIMONYI, Kohl.
221 2. Orotava, Tenerife; on the sea-shore, 20
and) 2111, 02,
MISCOPHUS EATONI, n. sp.
Niger nitidus subeneo-micans, facie punctatissima, mandibulis
testaceis, apicibus piceis ; alis hyalinis, margine apicali late infus-
cata, propodeo transverse rugoso, medio canaliculato, lateribus
oblique striatis, abdomine nigro, punctato et microscopice ruguloso,
segmentorum apicibus subpiceis, pedibus nigris.
? Black, shining, head and thorax in certain lights with a very
slight bronzy tinge, face very slightly convex, so closely punctured as
to be almost dull, a very fine smooth line extends from between the
antennew to about halfway between their insertion and the anterior
ocellus. Clypeus more shining than the rest of the face, slightly
raised down the centre, its anterior margin reflexed and rounded in
the centre. Mandibles testaceous darker at the base and apex. 3rd
joint of the antennee slightly longer than the following ones, vertex
shining, less closely punctured than the face; ocelli equidistant
from each other, cheeks posteriorly with a few short glittering pale
golden hairs, thorax shining and punctured like the vertex of the
head, wings hyaline, with a well-defined darker apical band, not quite
extending to the cell nerves, area of the petiolated cell of much less
extent than that of the radial, mesopleure distinctly punctured,
legs entirely black, propodeum shining transversely and somewhat
diagonally rugose, with a well-defined central sulcature, its sides
210 Mr. Edward Saunders on Hymenoptera aculeata
diagonally strigose, and clothed with very short inconspicuous pale
golden hairs, abdomen black, finely punctured and microscopically
rugose between the punctures, posterior margins of the segments
slightly piceous, and with a very insignificant fringe of white hairs
at the sides, beneath finely and closely punctured, the posterior
margins with a few long exserted hairs,
Long. 5 mm.
Of this very distinct species, one specimen was taken at
Orotava, Tenerife, on waste ground near the road beyond
the Cemetery gate—21. in. 02.
AMMOPHILA (Psammophila) TYDEI, Guill.
@ 1. Laguna. Hill-side east of the city, about 2000
feet altitude, 15. 11. 02.
Orotava. “ Common, provisions its nest with the
larva of a Noctuid moth.”
AMMOPHILA (Psammophila) HIRSUTA, Scop.
¢222. T. V. W. These were named by F. Smith,
“ Madere, Dhb.,” but are I think referable to
the var. of hirsuta, which has pale hairs on
the propodeum; I have taken similar speci-
mens in Jersey and at St, Briac in Brittany.
SCELIPHRON TUBIFEX, Latr.
g 12 3. Madeira. T. V. W. All these have the
pedicel cf the abdomen black.
OXYBELUS TEGULARIS, sp. n.
Niger, abdomine flavomaculato, propodei mucrone apice dilatato
emarginato, tegulis nervisque basalibus alarum lete testaceis, pedibus
nigris, testaceo flavoque variegatis.
This species resembles /atvo, Dahlb., in the widened
emarginate propodeal spine, but differs widely from it in
other characters and is specially recognizable by the
bright testaceous tegule.
Black, without any bronzy tinge, head and thorax densely,
rugosely and closely punctured, mandibles red in the centre,
flagellum of the antennz red towards the apex, clypeus tridentate at
the apex in both sexes, with a strong central keel in the ¢ which
projects beyond the anterior margin on to the central tooth, and a
short angular keel in the 2 abbreviated before it reaches the anterior
margin, which is smooth and shining ; face clothed with glittering
collected in Madeira and Tenerife. 211
silvery hairs, vertex rather densely with short greyish-white ones,
those on the cheeks behind the eyes short and silvery.* Mesonotum
clothed with short greyish hairs with a narrow central keel posteriorly,
extending on to the scutellum, tegule and bases of wing-nerves
bright testaceous, rest of neuration brown—postscutellum longitudi-
nally strigose, lamellee pale ; legs black, anterior tibize and tarsi
except the former posteriorly, in both sexes, and all the tarsi in the
dé, testaceous, bases of the intermediate and posterior tibiz flavous,
the pale colour more extensive in the @; calcaria pale, intermediate
and posterior tarsi black in the ? , posterior metatarsi piceous in the ¢,
propodeal spine short, widened and emarginate at the apex, central
area shining, more or less diagonally rugose on each side, lateral
areas dull, with irregular transverse rugosities, sides shining, trans-
versely strigose ; abdomen more shining in the ? than in the 4,
strongly punctured, clothed with grey adpressed pubescence, especi-
ally in the ¢, puncturation of the basal segment slightly larger
and less close than on the following, ¢ with a narrow trans-
verse yellow spot on each side of the apical margin of the first 5
segments, these become longer and more band-like as the segments
approach the apex, in the @ the spots are paler and only occur on
the first 2 or 3 segments; in the @ the 3rd, 4th, 5th and 6th
segments have a short testaceous spine-like tooth at the posterior
angle,
Long. 5—7 mm.
Orotava, Tenerife, several $ and §, 20. iu. 02.
VESPA GERMANICA, Fab.
5. Monte Funchal, Madeira, above the Church, at
about 2000 feet altitude, visiting Scrophu-
laria, two or three examples at the same
individual plant, 4. 111. 02.
& 1. Funchal, altitude 3000 feet (EK. 8S. Goodrich,
in Hope Collection), 3.1. 01.
oP2) 6/2. Mademas >: TV. W.
% 6. Orotava, Tenerife, March and April 1902.
F. A. Bellamy.
The black on the abdomen of some of Mr.
Bellamy’s specimens is very extensive, quite
as much so as in dark examples of Vespa
vulgaris.
PoLISTES GALLICUS, L.
23. Madeira, .-V. W,
212 Mr. Edward Saunders on Hymenoptera aculeata
ODYNERUS CRUENTATUS, Brullé.
ef and ®. Orotava, Tenerife, westward of the port.
ODYNERUS H&MATODES, Brulleé. :
? 1. La Laguna, Tenerife. Hillside to left of main
road to Tejina along a terrace of volcanic rock.
Altitude about 2200 feet, 15. in. 02.
® 1. Sauzal. Altitude about 900 feet, 4. iv. 02.
¢ 1 22; Madeira, “Va;
PROSOPIS, sp. ?
Madeira. T. V. W.
Carded and marked in collection as n. sp. allied to
signata.
PROSOPIS ATRA, 0. Sp.
Omnino nigra, facie maris flava, antennarum utroque sexu
flagello subtus testaceo exceptis, genis brevissimis, abdominis seg-
mento basali remote punctato, ruguloso, in femina microscopice,
apice linea laterali pilorum albidorum ornato.
Entirely black in the 9 ; ¢ with the face and a spot on the
anterior side of the front tibize yellow, flagellum of the antenni in
both sexes more or less testaceous beneath, wings hyaline, the
nervures brownish-black.
¢ Head and thorax and first joint of the antennz clothed with
long pale hairs, face yellow, of the same colour as in communis, the
colour not or rarely extending above the insertion of the antenne,
elypeus black at the apex, mandibles black, antennee with the first
joint curved but scarcely dilated, cheeks linear, vertex strongly and
closely punctured, thorax strongly punctured, but less closely than
the head, the intervals microscopically rugose, propodeal area
reticulate, abdomen somewhat dull, rugulose, punctured, punctura-
tion of the first segment fine and remote, that of the others,
closer and less definite, apex of the lst segment with a very
ill-defined fringe of white hairs at the sides, and 8th ventral
segment formed much as in pictipes, Nyl., but the apical wings of
the 7th are narrower and longer, and are twisted downwards at
an angle of about 60 degrees from the horizontal basal plates, and
are in this respect quite unlike those of any species I know—armature
with the stipites more or less pointed, with very long apical hairs.
Sagittz wide, paler, and slightly longer than the stipites, each
gradually narrowing to a rather blunt apex.
? punctured and sculptured much as the ¢, but the abdomen more
collected in Madeira and Tenerife. 213
shining, with quite microscopic rugulosities on the basal segment as
in genalis, face short, cheeks linear, orbital furrows much produced
posteriorly, slightly curved at their termination near the base of the
posterior ocelli, propodeal area smoother than in the 4, the reticu-
lations almost obsolete except at the base, basal segment of abdomen
with a distinct very narrow line of snowy-white pubescence ;
calcaria pale.
Long. 5 mm.
Orotava, visiting Prankenia, f 3, 21. i. 02.
Laguna, Tenerife, 2 1. 2 or 3 kilom from the city upon
the ridge to the left of the Tejina road. Altitude 2100—
2500 feet.
Sauzal, #1. Altitude about 900 feet.
HALICTUS, sp. ?
gf and 9. Madeira. T. V. W.
This pair is named quadristrigatus in the collection,
but neither @ nor 2 is I think referable to that species,
and in this opinion Mons. Vachal agrees; whether they
are f and ? of one species is also doubtful; the ¢ has the
clypeus entirely black; the 2 is in such bad condition that
it is impossible to be sure of its identity, and I hesitate to
describe a new species on a single @.
HALICTUS, sp. ?
2 3. Laguna, 15. ii. 02.
Closely allied to, if not a variety of H. scabiosx, Rossi,
but without knowing the other sex, I am afraid to deter-
mine it for certain, as there are already several closely
allied forms described in the group whose specific value
is very doubtful; Brullé (Webb-Berthelot, Hist. Can. 11,
p. 87) records scabiose as occurring in the islands, but
remarks that he has only seen one ¢@, which differs from
scabiose in having the antenne beneath entirely yellow;
this makes me suspect that Brullé’s ¢ and these ?s may
belong to a distinct Canary Island species.
HALICTUS, sp. (?).
Q Madeira. T. V. W.
In Wollaston’s collection there is a single 2 under the
name zebrus, Walck., but it is in such a condition as to
render identification practically impossible.
214 Mr. Edward Saunders on Hymenoptera aculeata
HALICTUS VILLOSULUS, Kirb.
9 1. Monte Funchal (Madeira). Sheltering on a
flower of Vinca Major, 27. ii. 02.
92. Monte Funchal (Madeira). Sheltering on
Crepis and Taraxacum officinale, 4, 11. 02.
2 4, Laguna (Tenerife), 15. ii. 02.
9 2. Madeira, T. V. W.
These females do not quite agree in all points with
British examples of the above species, but I see no dis-
tinctive character to rely upon, so without seeing the ¢ I
think it is wiser to treat them as belonging to it. Mons.
Vachal, to whom I submitted a specimen, returned it to me
as villosulus ?
HALICTUS ALCEDO, Vachal.
g@ 1. Laguna, Tenerife. — Hillside, east of the city,
about 2000 feet altitude.
HALICTUS LETUS, Brullé.
2 1. Orotava, Tenerife, at la Cabezas, 24. 11. 02.
HALICTUS, n. sp.
2 1. Monte Funchal, Madeira, on Sonchus oleraceus,
1500 feet altitude, 27. ii. 02.
Allied to Moric, F., but more strongly punctured. In
the Wollaston collection there are two ¢ and two @ of
I believe the same species, but they are on cards and
therefore no satisfactory description can be made from
them; the males are peculiar in having very large heads
and unusually ovate bodies.
HALICTUS ViIRIDIS, Brullé.
9 4. Orotava, Tenerife, on Oxzalis by the sea-shore,
20 and 25sin. 02:
® 2. Laguna, Tenerife, 25. ii, 21-7. iv. 02. F. A.
Bellamy.
I am indebted to Mons. Vachal for the determination of
this species, as I had described it asnew. I was misled by
Brullé’s remark : “ Elle est revetue d’un duvet blanchatre
non seulement sur tout le corp mais aussi sur les pattes ”—
in the specimens I have seen this rubbed off, but traces of
it are left on portions of the abdomen; also he gives the
collected in Madeira and Tenerife. 215
size as that of swbawrata, Rossi, whereas it is almost as
large as cylindricus, F., but as Mons. Vachal saw Brullé’s
types in Paris recently there can be no doubt that his
identification is correct.
HALICTUS ARCTIFRONS, n. sp.
Niger, abdominis segmento basali apice, segmentis 2, 3 et basi
segmenti 4, testaceis, calcaribus pallidis, 3-spinosis, abdomine crebre
punctato.
? closely allied to H. angustifrons, Vachal (Bull. Soc. Ent. Fr.,
1892, p. xxii), to which at first I was inclined to refer it, but on
communicating a specimen to the author he very kindly points out
that it differs from his species in being stouter and having the
posterior calcaria tridentate whereas in angustifrons they are 4-5
pectinate, and also in having the 2nd transverse cubital nervure
united with the Ist recurrent, whereas in angustifrons the 1st recur-
rent is received into the 2nd cubital cell at about a quarter of the
length of the cell from its apex. In colour it seems to agree well
with angustifrons, the head and thorax are black and the abdomen
black at the base and apex and testaceous red, from the middle of
the basal segment to the middle of the 4th. The eyes converge
rather rapidly so as to give the face rather an elongate triangular
aspect, the head and thorax are closely punctured and clothed with
whitish hairs, and there is a line of dense white pubescence in the
anterior depression of the pronotum and also in the depression
bordering the mesonotum, postscutellum also densely clothed with
whitish pubescence, propodeum with its basal area shallow and
semi-circularly bounded, finely clathrate at the base, shining at the
apex on the brow ; abdomen closely punctured, the puncturation of
the basal segment slightly less close than that of the 2nd, legs black
clothed with whitish hairs, apex of posterior metatarsi with a tuft
of golden bristly hairs.
Long, 5 mm,
Orotava by the sea-shore, visiting Frankenia, 20 and 23.
ili. 02.
ANDRENA BIMACULATA, Kirby ? var.
g and ¢. Madeira. T.V.W. In too bad condi-
tion to determine for certain.
ANDRENA BIPARTITA, Brullé.
9 2. Orotava, Tenerife, 21. iii. 02.
216 Mr. Edward Saunders on Hymenoptera aculeata
ANDRENA MINUTULA, Kirb.
ft 2. Monte Funchal, Madeira, 6.11.02. At about
1100 feet altitude.
g 122. Madeira, IV. W-
These are I believe referable to minutula, but belong to
the form which has the mesonotum rugulose and with
very distant, shallow punctures. We have similarly sculp-
tured specimens in England, but those I possess belong to
the second brood, whereas the males of this collection have
the long-haired face of the first brood.
OSMIA LATREILLEI, Spin.
f2 91. Laguna, Tenerife, 15—17. iii, 02. At
altitudes of 2000—2200 feet.
$424. Madeira. T. V. W.
OsMIA SUBMICANS, Mor.
g 2. Laguna, Tenerife, 16. 11. 02. “ Hill north of
the city, at about 2150 feet altitude and under.”
MEGACHILE APICALIS, Spin.
g¢ 2. Orotava, Tenerife, 21. iii. 02.
MEGACHILE, sp. ?
¢ 1, Madeira. T. V. W.
In bad condition, but apparently closely allied to versi-
color, Smith, with similar pale apical joints to the tarsi.
PODALIRIUS QUADRIFASCIATUS, Villers.
a6 9 3. Madeira, To V. W.
Of the usual Madeiran type with fulvous bands = var.
Maderx, Sichel.
PODALIRIUS CANARIENSIS, N. Sp.
Niger clypeo maris triangulariter albosignato, facie albido-hirta
in lateribus pilis nigris intermixtis. Thorace albido-hirta fascia
interalari nigré abdominis segmento primo albido-hirta, reliquis
nigrohirtis fasciis interruptis apicalibus ornatis. ¢ metatarsis
intermediis ciliis nigris postice dense vestitis.
Although so like atroalba, Lep., as to be scarcely dis-
tinguishable in the §, the f is abundantly distinct in
collected in Madeira and Tenerife. 217
coloration and especially in the disposition of the black
hairs on the intermediate metatarsi; these are arranged
much as in balearicus, Freise, all being on the posterior
side of the joint.
¢ and @ very similar in coloration; black, the ¢ having a
triangular spot on the clypeus, sometimes a more or less extensive
spot on the labrum and a line on the front of the scape yellowish-
white ; in both sexes the hairs of the face are white intermixed with
black. The face in the ? is very broad, and the cheeks between the
eyes and mandibles very short, labrum clothed with silvery hairs.
The thorax is clothed with grey and black hairs intermixed above,
with nearly black hairs on the sides beneath the wings and with paler
hairs round the propodeum, abdomen clothed with greyish-white
hairs on the basal segment, with black hairs on the others, the 2nd,
3rd and 4th segments with a band of snowy-white rather long
hairs at the apex, slightly narrowed at the sides and broken in the
centre, legs clothed with black hairs, the intermediate and posterior
tibiz in both the ¢ and 9 with bright silvery hairs exteriorly and
the metatarsi of the same legs in the ¢ with silvery hairs at the
apex, intermediate metatarsi with a dense fan of black hairs on its
posterior side, in the 2 the silvery hairs of the scopz have a some-
what golden tinge.
Long. 12-13 mm.
La Laguna, Tenerife. Hill-side on left-hand side of main
road to Tejuna, nesting in cavities of volcanic rock, altitude
about 2200 feet, males only.
The 2 which I associate with this ¢ is from the collec-
tion of Mr. Morice taken in Tenerife, it very closely
resembles that sex of atroalba, but the face is wider, its
hairs are mixed with black, whereas in atroalba they are
all white. The hairs of the labrum are silvery and not
dull grey, and the bands of the abdomen are of much
longer hairs.
BompBus HorroRuUM, Linn.
2 1. Monte Funchal, Madeira, 17. 11. 02.
“In garden of Belmonte Hotel, visiting Azalea: nest
subterranean, common between 1450—1700 feet.” Differs
only from the ordinary type in having the hairs of the
apical segments of a brownish tinge, and the hairs of the
scope more or less reddish.
fg 223. Madeira. T.V.W. Similar to the above
in coloration.
218 Mr. Edward Saunders on Hymenoptera aculeata.
BoMBUS TERRESTRIS, Linn.
f 3 92. Laguna, Tenerife, 14—16.ii1. 02. Allti-
tude 1900—2200 feet. “A common species.”
Numerous #9 and 9 various localities in Tenerife.
F. A. Bellamy. March and April 1902.
All the specimens are entirely black, with only the apex
of the abdomen white. A very unusual variety of this
species ; the ¢ armature, however, is like that of normal
specimens.
( 219 )
XIII. Descriptions of Twelve New Genera and Species of
Ichneumonidz (Heresiarchini and Amblypygi) and
three species of Ampulex from the Khasia Hills,
India. By PETER CAMERON, communicated by
GEORGE ALEXANDER JAMES ROTHNEY, F.ES.
[Read May 6th, 1903.]
spate
Caspipina, gen. nov.
Mandibles curved, sickle-shaped, unidentate. Apex of clypeus
transverse ; not separated from the face ; the fovex distinct. Occi-
put sharply margined. Scutellum flat; its sides on the basal half
keeled. Median segment completely areolated ; the areola, if any-
thing, broader than long and open at the base. Areolet 5-angled ;
narrowed at the top, the transverse basal nervure interstitial. Legs
stout ; the tarsi spinose. Antenne compressed and somewhat
dilated beyond the middle. The apex of the median segment has
an oblique, straight, not rounded slope. The inner orbits are sharply
margined. The recurrent nervure is roundly curved outwardly and
bears the stump of a nervure in the middle.
Caspipina ferruginea, sp. nov.
Ferruginous ; the edge of the pronotum, the lower edge of the
propleure, the tubercles, the mesopleurze broadly in the middle at
the base and the centre of the metapleure, yellow ; the edges of the
mesonotum, the space at the sides of the scutellums, the apex of the
median segment—the black projecting upwards along the keels—a
band shortly below the middle of the propleure, the base, top,
and apex of the mesopleure, and the base and lower-side of the
metapleure, black. Legs ferruginous, the fore cox yellowish ; the
tarsi thickly covered with short, stiff hair ; the incision on the base
of the fore tarsi wide and deep. The basal sixteen joints of the
antenne ferruginous, the rest black. Wings fusco-hyaline, the
nervures and stigma black. 9°.
Length, 13 mm.
Hab. Kuasta Hints. Coll. Rothney.
Head shining, the face and clypeus closely, but not strongly,
punctured and sparsely covered with short pale pubescence ; the
TRANS, ENT. SOC. LOND. 1903.—PART Il. (JUNE)
220 Mr. P. Cameron on Twelve New Genera and
front and vertex closely punctured, the inner and outer orbits
obscure yellow. Mandibles with a yellowish tinge at the base, the
apex black. Mesonotum closely and finely punctured, as are also
the scutellums. Median segment closely, rugosely punctured, the
punctures running into striae on the apex and on the spiracular
aree. Propleure shagreened, the meso- and metapleure closely
punctured. Prosternum stoutly keeled in the middle ; its apex and
the base of the meso- black. The base of the Ist abdominal
segment and of the 2nd, black; the 4th and the following
segments more or less black ; the post-petiole is raised in the middle
and separated from the sides ; its middle slightly depressed, closely
punctured and finely longitudinally striated, except at the apex ; the
gastroceli are stoutly striated.
ENCHISIADES, gen. nov.
Mandibles curved, ending in one tooth; the apex gradually
narrowed, rounded. Face flat. Labrum projecting. Clypeus not
separated from the face, its apex transverse. Scutellum roundly
convex, not much raised, its sides distinctly keeled. Post-scutellum
bifoveate at the base ; its sides not keeled. Median segment with a
gradually rounded slope, completely areolated, its areola longer than
broad, rounded behind, its apex bulging backwards, and with the
keel thin. Areolet 5-angled, narrowed above, the nervures almost
touching there. Post-petiole broadly dilated, distinctly separated
from the petiole. Legs stout; the tarsi long, spinose. Antenne
long, slender, slightly but distinctly, dilated towards the apex.
The hinder coxe are larger than usual ; the head is only slightly
developed behind the eyes ; the occiput is broadly rounded inwardly ;
its lower edge narrowly, but distinctly, keeled.
ENCHISIADES RUFIPES, sp. nov.
Black ; the face, clypeus, labrum, mandibles, orbits, the edge of
the pronotum narrowly, the line not extending to the apex, the
lower edge of the propleure slightly more broadly, the scutellar
keels, the sides of the scutellum, post-scutellum, the lateral region
of the apical slope of the metanotum, the tubercles, a line on the
inner- side of the mesopleure, commencing above the middle at the
base, where it is narrowed along the basal half and more broadly
on the lower half, where it is obliquely turned upwards ; two marks
under the hind-wings anda small mark above the middle of the
hinder coxe, lemon-yellow. Legs rufous ; the four front coxe and
trochanters and the four apical joints of the hinder tarsi yellowish-
white, the hinder coxie black ; the apex and an oblique line on the
Species of Ichneumonide from India. 221
middle above, lemon-yellow ; the apex of the hinder femora, the
base of the hinder tibie, their apical third and the metatarsus, black ;
the tarsi thickly spinose. Abdomen black ; an oblique mark on
either side of the post-scutellum, a large mark, broader than long, on
the sides of the 2nd, 8rd, and 4th segments with their apices
narrowly in the middle, lemon-yellow. Wings hyaline, the nervures
and stigma black. @.
Length, 11-12 mm.
Hab, Kuasta Hitis. Coll. Rothney.
Antenne black, the 10th to 17th joints beneath white ; the
scape minutely punctured and covered with black down. Face
and clypeus closely and uniformly punctured, the vertex closely and
minutely punctured, in the centre below closely transversely striated ;
the front smooth and shining. In the centre of the face is a large
bell-shaped black mark, its lower edges uniting with the clypeal
fove. Mandibles yellow, black at the apex ; the palpi pallid yellow.
Mesonotum opaque, granular, more distinctly punctured along the
sides and covered with a pale down. Scutellum with the basal half
indistinctly, the apical closely and distinctly punctured ; it is thickly
covered with pale hair; the basal keels are stout and extend to
shortly beyond the middle. Basal half of post-seutellum punc-
tured ; the two foveze large and deep; the depression at its sides
has five stout longitudinal keels. Metonotum closely and distinctly
punctured ; beyond the basal arew more or less transversely striated.
Pleure closely punctured ; the pro- at the base closely and finely
striated.
Darachosia, sp. nov.
Mandibles with a long, curved apical tooth and a short blunt one
in the middle. Antenne as long as the body, dilated and compressed
before the apex. Labrum projecting. Scutellum flat, its sides keeled.
Median segment completely areolated, its spiracles linear ; the areola
twice longer than broad, rounded behind, its apex indented by the
posterior median area. Legs long, the tarsi longer than usual ; the
anterior and hinder twice the length of their tibiz. Areolet 5-angled,
narrowed above. Abdomen with eight doreal segments; the apex
moderately blunt; the last segment short, about one-half the length
of the penultimate ; the ventral keel is distinct on the 3rd and 4th,
indistinct on the 5th segment ; the petiole is long and slender; the
sheaths of the ovipositor project largely.
The occiput is transverse ; the eyes large, sharply margined on the
inner-side ; the pronotum at the base is broadly raised ; the petiole
is long and slender ; it is not abruptly dilated at the apex.
TRANS. ENT, SOC, LOND. 1903.—PART I. (JUNE) 15
922 Mr. P. Cameron on Twelve New Genera and
A. distinct genus easily known by the form of the
mandibles, by the long antennz dilated before the apex,
and by the very long spined tarsi.
This genus, as also do Hvirchoma, Sycaonia, and Legnatia,
as regards the mandibles, forms a transition between the
Amblypygi aud the Heresiarchini, there being a short sub-
apical tooth, which is indistinct and more widely separated
than it is with the former group, while in the latter the
mandibles become gradually narrowed.
Darachosia fulvipes, sp. nov.
Black ; the face, except for two short black lines in the middle,
the clypeus, the eye orbits—the inner on the upper-side narrowly,
the outer more broadly—the malar space, a line on the pronotum, two
lines on the mesonotum, in the middle, one opposite the tegul, the
seutellum, except in the middle—the black central mark narrowed
and rounded, its apex transverse—the post-scutellum, the sides of the
apical slope of the median segment, the yellow extending laterally
on to the spiracular area and at the apex on to the pleure ; the base
of the pronotum, a line below the middle of the propleure, the meso-
pleurz from shortly below the middle, the tubercles and the apex of
the mesopleure narrowly, pale yellow. Legs fulvous, the four front
coxe and trochanters pallid yellow; the femora lined with black
above, the front tarsi infuscated ; the hinder cox black; the apex
and the middle broadly above on the apical half, pale yellow ; the
trochanters, more than the apical third of the femora, and the apex
of the tibia, black ; the hinder tarsi pale yellowish. Abdomen black ;
the apices of all the segments lined with yellow, the apical three
segments more narrowly than the others. Wings hyaline, their base
with a slight fulvous tinge ; the stigma testaceous, the apical nervures
of a darker testaceous colour, the basal black; the areolet much
narrowed above, the nervures almost touching there. 9.
Length, 15 mm,
Hab. KHASIA Hitis. Coll. Rothney.
Antenne as long as the body, the scape below and joints 11-17
white ; the thickened joints brownish beneath; the scape closely
punctured and thickly covered with short white hair. Face closely
punctured ; it and the clypeus are covered with white pubescence.
The lower part of the front is smooth and shining, the upper
transversely striated, punctured laterally. Mesonotum closely and
uniformly punctured ; the scutellum is closely and more strongly
punctured ; its keels yellow. Median segment, except at the base in
Specves of Ichneumonide from India. 223
the middle, closely punctured ; the areola closely punctured on the
apical half; the basal smooth and shining and furrowed laterally.
Pleure closely punctured; the pro- striated behind; the meta-
more closely and strongly than the meso-. Mesosternum closely
punctured ; its basal slope stoutly transversely striated in the middle
and bearing two transverse yellow marks; on the apex, near the
middle coxe, are two small oblique marks. The petiole closely and
somewhat strongly punctured ; its sides, on the apical half, striated ;
the 2nd, 3rd and 4th segments closely and uniformly punctured ; the
gastroceeli smooth, striated round the edges; the space between
them closely, longitudinally striated.
Hvirchoma, gen. nov.
Mandibles curved, with one longish apical and a short subapical
tooth, not separated behind. Apex of clypeus broad, transverse,
above separated from the face by a wide suture, which is deep on the
sides, shallow and wider on the top. Labrum projecting. Occiput
margined, Scutellum margined laterally at the base, large, convex,
gradually rounded at the base and apex. Median segment completely
areolated ; the areola longer than broad, rounded at the base, of
almost equal width throughout ; the spiracles linear, curved. <Areolet
5-angled, much narrowed at the top, the nervures almost touching
these ; the transverse basal nervure not interstitial, Abdomen with
seven dorsal segments, the last large, not retracted, as long as the
preceding, bluntly pointed ; the ventral keel is on the 2nd, 3rd and
4th segments ; the ovipositor projects, and is as long as the apical two
seginents united. Tarsi spinose.
Lvichoma pallidimaculata, sp. nov.
Scape of antenne rufous, closely, but not strongly, punctured ; the
flagellum stout, the 8th to 20th joints white, tinged with rufous, the
rest black, more or less brownish. Face and clypeus strongly and
closely punctured, the apex and sides of the latter smooth ; the face
yellow, broadly rufous in the middle; the clypeus broadly rufous in
the middle, the sides paler, and there is a darker band between.
Orbits yellow, broadly below; the vertex dark rufous, closely
punctured ; the front blackish, smooth. Apex of labrum fringed
with long, fulvous hair, Mandibles rufous, the teeth black. Palpi
yellow. Thorax dark rufous, the lower part of the pro-, the lower
half of the mesopleure, the scutellums and the sides of the median
segment at the apex, yellow, suffused with rufous. Mesonotum dark
rufous, very closely punctured. Scutellum strongly longitudinally
punctured, almost striated in the middle; the base black, the rest
yellowish, running into rufous. Post-scutellum yellow, finely
224 Mr. P. Cameron on Twelve New Genera and
longitudinally striated. Median segment with the central three
ares smooth ; the basal central wider than long, narrowed towards
the apex ; the basal lateral bear shallow, widely separated punctures ;
the posterior median strongly transversely striated ; the spiracular °
at and behind the spiracles, almost impunctate, the rest obliquely
striated, the striz becoming stronger and more widely separated
towards the apex, especially behind the teeth, which are large,
keeled on the under-side, and united with the lateral keels. Pro-
pleure finely punctured above. Mesopleure yellow, black above,
under the tubercles and halfway down the middle, the lower part of
the base behind the keel black; the lower part is strongly longi-
tudinally striated, the upper strize being much shorter than the
lower. Metapleurew below the keel black, except at the base above,
and the apex obliquely on the lower-side; the yellow part is narrow
at the top, becoming wider at the bottom. Legs dark rufous; the
femora darker ; the four front cox and trochanters yellowish-white ;
the hinder broadly dark rufous at the base in front ; behind yellow,
except for a large mark on the base below ; the basal joint of the
hinder trochanters dark rufous ; the hinder tarsi thickly spined.
Petiole black, its apex yellow; the 2nd and 38rd segments black,
impunctate ; their apices broadly rufous ; the other segments for
the greater part dark rufous, their apices yellowish. The ovipositor
sheaths pilose, largely projecting, rufous in the middle.
Sycaonia, gen. nov.
Mandibles with one short upper tooth, distinctly projecting be-
yond the lower, which hardly projects and is bluntly rounded.
Face distinctly and broadly projecting in the middle, the projecting
part bordered laterally by a shallow furrow, which is wider and
more distinct below. Clypeus not separated from the face. Occiput
margined. Antenne stout, thickened beyond the middle. Scutellum
flat, only slightly roundly convex ; its sides not margined. Post-
scutellum with a depression on either side at the base and bordered
on the outer-side by a sharp keel. Median segment regularly
areolated ; its sides with a blunt tooth ; its spiracles curved, linear ;
areola separated by its own length from the base of the segment ;
it is longer than broad, slightly narrowed towards the apex. Areolet
5-angled, narrowed above. Legs stout, the tarsi spinose. Abdomen
with seven segments ; the last large all round ; if anything longer
than the penultimate ; its apex somewhat bluntly pointed.
Sycaonia rufo-facies, sp. nov.
Black ; the orbits all round narrowly, the apex of the petiole and
a mark, about three times broader than long and rounded on the
Species of Ichneumonide from India. 25
inner-side, on the sides of the 2nd abdominal segment at the apex,
yellow. Legs rufous, the four front coxee and trochanters, the tibiz
to beyond the middle—about the basal two-thirds—pale yellow ;
the hinder cox on the sides and beneath, the hinder trochanters,
apex of femora, their base more narrowly and the apex of the tibie
black ; the tarsi are covered thickly with stiff spines, The wings
have a slight, but distinct, fuscous tinge ; the stigma and nervures
are fuscous ; the 2nd transverse cubital nervure is faint. ?.
Length, 11 mm.
Hab. Kuasta Hints. Coll. Rothney.
Antenne with the 10th to 19th joints white except above ; the
scape rufous beneath, Face and clypeus strongly and closely
punctured and thickly covered with white pubescence. Front above
and elypeus closely and distinctly punctured. Mesonotum closely
punctured and thickly covered with short black hair. Scutellum
slightly roundly convex, sparsely punctured and thickly covered
with longish pale hair. The base of the post-scutellum is obliquely
depressed ; its apex has an oblique straight slope. Areola smooth
and shining ; the posterior median area closely transversely striated ;
the other ares closely and distinctly punctured; the spiracular
transversely striated beyond the stigma; the stricee on the apical
lateral areze are coarser, more irregular and more widely separated.
Pro- and mesopleure closely and minutely punctured, the apex and
middle of the former closely and the lower half of the latter, more
widely and strongly striated. The metapleure, if anything, are
more strongly punctured and with the apical half obliquely striated.
Petiole smooth above, except on the sides of the post-petiole, which
are punctured ; the gastrocceli shallow, closely striated near the
apex, which is testaceous ; the sides are closely striated ; the apices
of the other segments are narrowly testaceous, this being the case
also with the 2nd, 3rd and 4th ventral segments.
Legnatia, gen. nov.
Mandibles with one long, curved upper and a short blunt sub-
apical tooth. Abdomen with eight dorsal segments ; the cerci large ;
the apical segment blunt, small; the ventral keel on the 2nd and
3rd segments only; the ovipositor largely projecting. Antenne
longish, dilated and compressed beyond the middle. Labrum hidden.
Scutellum flat ; its sides keeled. Median segment completely areo-
lated ; it is longish and has a gradually rounded slope ; its areola
widely separated from the base of the segment, which is not depressed ;
it is twice longer than wide; the spiracles linear, curved. Areolet
226 Mr. P. Cameron on Twelve New Genera and
5-angled, narrowed above. Legs longish; the tarsi long, the hinder
almost twice the length of the tibize ; the apices of the joints spined.
The post-petiole becomes gradually wider from the base to the
apex and is not clearly separated ; the gastrocceli large and deep ;
the head is not largely developed behind the eyes, and becomes
obliquely wider below before the middle; the eyes are large and
are sharply margined on the inner-side.
A genus easily known by the long antenne dilated
beyond the middle, by the curved mandibles with long
apical tooth, by the long spined tarsi and by the abdomen
having eight segments.
Legnatia fulvipes, sp. nov.
Black; the face, clypeus, the inner orbits, the outer—narrowly
above, entirely below—the lower part of the prothorax, the edge of
the pronotum, two lines on the basal two-thirds of the mesonotum,
the scutellum, except a narrow line down the middle, a squarish
mark on either side of the base of the metanotum, its lateral areze
entirely, the lower half of the mesopleuree, a mark under the hinder-
wings, the metapleure from shortly behind the middle, where they
are obliquely rounded, a small mark at the spiracles, the sides and
apex of the post-petiole, the apices of the 2nd and 3rd segments
broadly laterally, more narrowly in the centre, a line on the apex
of the 4th, interrupted in the middle, a small round spot on either
side of the middle of the 5th, the 6th on the apical half of the middle,
the whole of the 7th, and the 2nd, 3rd and 4th ventral segments for
the greater part, pale yellow. Legs fulvous; the four front cox
and trochanters and the hinder coxee, pale yellow ; the apex of the
latter on the outer-side, the apex of the hinder femora, of the hinder
tibie and the base of the metatarsus, black. Wings hyaline, the
stigma pale, the nervures black. @.
Length, 11 mm,
Hab. Kuasta Huts. Coll. Rothney.
Antenne black, as long as the body, the scape beneath and the
9th to 16th joints white beneath. Face and clypeus closely punctured
all over and sparsely covered with pale pubescence ; the vertex more
sparsely punctured, more closely and distinctly at the ocelli than
below ; the black band on the vertex and front is contracted on the
top of the latter. Mesonotum closely punctured; the scutellum
much more sparsely punctured ; the post-scutellum smooth. Median
segment closely, rugosely punctured ; the areola is shagreened and
is furrowed round the sides and base; the posterior median area is
Species of Ichneumonide from India. 227
strongly, if somewhat irregularly, transversely striated, as are also
the posterior intermedian. The lower half of the propleure longi-
tudinally striated, the upper smooth. Meso- and metapleure closely
punctured.
Hutanyacra, gen. nov.
d. Apex of abdomen bluntly pointed, with eight segments, the
8th produced in the middle into a long narrow process, which is
three times as long as the basal portion of the segment; the ventral
fold is only distinct on the 2nd and 8rd segments. Face flat, not
separated from the clypeus. Clypeal fovee large, deep. Mandibles
with one long upper and a blunt lower tooth which is not clearly
separated behind. Apex of clypeus transverse, its sides rounded.
Labrum hidden. Occiput sharply margined. Scutellum roundly
convex. Median segment areolated ; the areola about as broad as
long, and with its apex rounded. Spiracles elongated. Legs stout,
of moderate length; the tarsi spinose. Antennx serrate, shorter
than the body, distinctly tapering towards the body.
The edigus is much larger than usual ; it appears to form one
solid piece, united below at the apex and with a broad and deep
furrow on the basal two-thirds ; at the base of this is a large tubercle,
on either side under the base of the projection on the apical segment ;
the apex, looked at from the sides, is bluntly rounded on the lower-
side, which projects more than onthe upper. The 8th dorsal segment
is small and bears longish cerci. Gastrocceli large, deep.
Belongs, by the bluntly-pointed apex of the abdomen
and by the ventral fold being on the 2nd and 3rd segments
only, to the Amblypygi. In Ashmead’s arrangement the
genus comes near Psewdamblyteles. The 2 I do not know,
but the peculiar structure of the 8th ventral segment, the
very large edigus and the large cerci make the % repre-
sentatives of the genus easy of recognition.
Hutanyacra pallidicoxis, sp. nov.
Black ; the face, clypeus, inner orbits, a small spot on the base of
the mandibles, the upper edge of the pronotum, the scutellums,
tubercles, tegule, the apex of the post-petiole narrowly, its sides
more broadly, a large mark on the sides of the 2nd and 3rd segments
on the apical half and narrowed on the inner side, the apex of the
6th segment to near the middle and the greater part of the 7th
segment, pale yellow. Legs pale yellow, the hinder coxx, the
hinder femora, except at the base and the apical two-thirds of the
last joint of the tarsi, black. Wings hyaline, with a slight fulvous
99283 Mr. P. Cameron on Twelve New Genera and
tinge ; the cubitus before the areolet and the 2nd transverse cubital
nervure are largely bullated. ?.
Length, 13 mm.
Hab. Kuasta Hitts. Coll. Rothney.
Antenne not much longer than the abdomen, distinctly tapering
towards the apex, the flagellum densely pilose, the apical joints
serrate ; black, brownish beneath ; the scape beneath, the 2nd and
the 7th to 14th joints white. Face and clypeus strongly and closely
punctured, and thickly covered with short white pubescence ; in
the centre of the face is a line which is irregularly dilated in the
middle; the front and vertex are strongly punctured ; the outer
orbits are less strongly and more closely punctured. Palpi pale
testaceous. Mesonotum closely punctured and thickly covered with
short silvery pubescence ; the scutellum is not so closely punctured
as the mesonotum, and is thickly covered with white pubescence ;
its apex is distinctly narrowed. Median segment closely rugosely
punctured ; the sides at the apex obliquely reticulated, more closely
behind than below. Pleure closely and distinctly punctured, as is
also the breast, which is deeply furrowed down the middle. Post-
petiole closely longitudinally striated ; the 2nd to 4th segments
closely punctured ; the gastrocceli stoutly striated.
Ancyra, gen. nov.
Areolet large, not much narrowed above, 5-angled. Clypeus not
separated from the face, its apex transverse. Labrum projecting.
Mandibles bidentate, the upper tooth not much longer than the
lower. Scutellum roundly convex, its base and apex with an oblique
slope. Median segment regularly areolated ; the areola about twice
longer than broad, transverse at the apex, the sides slightly angularly
produced in the middle. Metathoracic spiracles linear. Legs longish,
not very stout; the tarsi spinose. Abdomen longish, with seven seg-
ments, the last large ; the ventral fold on the 2nd and 3rd segments
only. Antenne short, not much longer than the abdomen, the
joints serrate beyond the middle; the apex distinctly narrowed.
Eyes large, parallel, their lower part reaching below the clypeal
foveze.
The stump of a nervure on the outer-side of the
recurrent nervure is longer than usual; there is none on
the disco-cubital; the transverse basal nervure is inter-
stitial. May be known by the large, wide areolet, by the
long, slender legs, by the large eyes which reach close to
the base of the mandibles, by the pyramidal scutellum
and by the long abdomen.
Species of Ichneumonide from India. 229
Ancyra flavomaculata, sp. nov.
Pallid yellow ; the hollowed lower part of the front, the ocellar
region, the vertex behind, an oblique mark behind and united to
the eyes near the top, the occiput, the mesonotum, except two yellow
lines down the middle, the parts at the sides of the scutellums, the
base, apex and central areee of the metanotum, a line in the centre
of the propleurz on the basal two-thirds, the base, upper part and
apex of the mesonotum, and two marks on the sides of the meso-
notum—broad at the apex, gradually narrowed behind—and the
greater part of the abdomen, black. Legs pallid yellow like the
thorax ; the four front coxee above and more or less below, the
four front trochanters more or less above, the base of the four
anterior femora, the hinder cox below at the base, the base of the
hinder femora to near the middle; the four anterior and the base
and apex of the hinder tarsi, the four front tibiz at the base behind
and the hinder at the base all round, black. Wings smoky-viola-
ceous ; the stigma and nervures black. Abdomen black ; the apical
half of the petiole, the apical half of the 2nd and 8rd segments,
except in the middle, and a small mark on the side of the 4th
segment, pallid yellow. ¢
Length, 19-20 mm.
Hab. Kuasta Hints. Coll. Rothney.
Antenne not much longer than the abdomen ; the scape pallid
yellow, black above ; the basal six joints entirely pallid yellow, the
7th to 13th yellow, marked with black above; the others black ;
serrate, distinctly tapering towards the apex. Face and clypeus
shining, punctured all over, but not very closely or strongly and
almost glabrous; the vertex is more closely punctured. Mandibles
yellow, their teeth black. Mesonotum closely punctured. Median
segment rather strongly punctured ; the areola roughly aciculated ;
on the sides are a few, oblique short keels, and on the apex in the
middle a central one ; except at the base the posterior median area
is closely, transversely striated ; on the smooth base are two longi-
tudinal keels, Pleuree closely punctured. Petiole closely punctured
at the base, the rest smooth. Gastrocceli deep and bearing three
stout keels. The black line on the 2nd abdominal segment is broad
at the base, narrowed at the apex ; that on the 3rd is much broader
and also gradually narrowed towards the apex ; the apex of the
segment is black.
Losgna, gen. nov.
Antenne compressed and dilated beyond the middle. Mandibles
bidentate, the upper tooth much larger than the lower. Scutellum
230 Mr. P. Cameron on Twelve New Genera and
large, quite flat, its sides sharply keeled. Apex of clypeus broadly
transverse. Labrum projecting. Head largely projecting behind
the eyes, at the top obliquely narrowed. Eyes distinctly margined
on the inner-side. Median segment completely areolated ; the areola
large, longer than broad, its base roundly dilated ; the sides of the
segment bluntly toothed ; the keel below leaf-like. Legs normal ;
the tarsi long, spinose. Abdomen with seven segments, the last
bluntly pointed, and as long as the penultimate ; the ventral keel
on the 2nd and 8rd segments only ; the ovipositor largely projects.
The head behind is deeply and roundly incised; above it is
depressed ; the occiput is not distinctly margined. In addition to
the longish spines on the tarsi, their joints bear shorter spines, and
are also thickly covered with stiff hairs. The ocelli are placed on
the edge of the occiput.
Belongs to the Amblypygi. May be known from
Amblyteles by the very flat, keeled scutellum, by the
dilated antennz, by the spined tarsi, and by the abdomen
having seven segments.
Losgna forticeps, sp. nov.
Black ; the outer orbits broadly below, the inner narrowly above ;
the face, clypeus, labrum, palpi and mandibles pallid yellow; a line
on the hinder half of the pronotum, its base, the sides and apex of
the scutellum, the scutellar keels, the sides and apex of the apical
slope of the median segment, the lower half of the mesopleure and a
mark behind the metathoracic spiracles, white. The four anterior legs
pale fulvous, the coxie and trochanters yellow; the hinder coxe,
trochantersand femora pale fulvous, marked with black above, the tibize
and tarsi black. Abdomen black, the base of the petiole broadly, an
oblique mark on the sides at the apex, the sides of the 2nd and 3rd
segments on the apical half, an oblique mark on the sides of the 4th
and 5th—all extending backwards at the base ; the apex of the 6th
ali round and the 7th entirely, pallid yellow, as is also the whole
ventral surface. Wings hyaline, the nervures and stigma black. 9.
Length, 15 mm.
Hab. KHasta Hitis. Coll. Rothney.
Antennz black, the scape and joints 11-20 white beneath. Face
broadly dilated in the middle, punctured ; the clypeus with scattered -
punctures. Front smooth and shining, broadly excavated laterally,
the ocelli in front surrounded by a semicircular furrow. Mesonotum
strongly and closely punctured and with two shallow, wide, longitu-
dinal furrows in the centre. The flat scutellum bears large, deep,
Species of Ichneumonide from India. 231
clearly separated punctures in the middle. Areola stoutly, closely,
mostly transversely, irregularly reticulated ; the posterior median
area closely transversely rugose. The hinder part of the propleure
stoutly, closely obliquely striated. Mesopleure closely, but not
strongly and the meta- closely, strongly, and uniformly punctured,
Pramha, gen, nov.
Metethoracic spiracles small, oval. Antennz thickened and com-
pressed beyond the middle ; annulated with white ; the basal joints
of the flagellum elongated. Clypeus separated from the face ; its
apex broadly rounded. Mandibles with a large apical and a small
subapical tooth. Labrum hidden. Areolet 5-angled ; the transverse
basal nervure interstitial. Abdomen with seven distinct segments ; its
apex blunt ; the ventral fold extends to the apex of the 4th segment,
the ovipositor large, projecting. Legs moderately stout, the penulti-
mate tarsal joint spinose.
Belongs to the Amblypygi. The scutellum is flat and
not keeled, the post-scutellum bifoveate; the face flat,
clypeal foveze large and deep; the occiput margined; the
temples not much developed ; the petiole as in [chnewmon ;
the gastrocceli large, shallow ; the transverse basal nervure
interstitial ; the last ventral segment is large; the cubital
nervure at the base is not angled, but broadly rounded.
The areola is wider than long, transverse in the middle with
the sides rounded at the base ; the apex rounded inwardly.
Pramha mandibularis, sp. nov.
Black, a line on the pronotum, broadest behind, the tecule,
tubercles, scutellums, a band on the base of the propleuree, yellow ;
the sides of the petiole broadly, the apex more narrowly, rufous ;
the 2nd, 3rd and 4th abdominal segments rufous and yellow at the
base and apex, the others yellow at the apex. The four front legs
fulvous, the coxe and trochanters yellow, the hinder pair rufous ;
the coxee fulvous-yellow behind, the tibiee of a much darker rufous
colour, the tarsi fulvous. Wings hyaline, the nervures fuscous, paler
towards the apex ; the lower part of the stigma pale testaceous. 9.
Length, 8 mm.
Hab. Kuasta Hints. Coll. Rothney.
The scape and the 10th to 16th joints of the antennz white, the 2nd
and the apices of the 3rd and-4th joints brownish. Face strongly
punctured, the clypeus less strongly and closely punctured and
covered with short white pubescence. The base of the mandibles
932 Mr. P. Cameron on Twelve New Genera and
broadly and the palpi white. Front and vertex closely, strongly and
uniformly punctured and sparsely covered with short pale hair.
Mesonotum closely and finely punctured and thickly covered with
short fuscous hair. Secutellum with a black mark, rounded above on
the apex; the basal depression wide, deep and stoutly keeled
laterally. Median segment closely punctured all over and, except at
base, thickly covered with long fuscous hair ; the basal area is wider
than long and has oblique sides; the posterior median area slightly
hollowed and not clearly limited, the keels not being very distinct;
it is closely transversely striated. The upper part of the propleure
weakly punctured, the apex below stoutly transversely striated.
Mesopleure closely and strongly punctured ; in the middle above
is an oblique shining furrow. Mesosternal furrow wide, deep and
with some stout keels in the middle. Metapleurz closely punctured ;
the lower half of the basal depression with three stout keels. Apex
of petiole aciculated and marked with large, closely separated, punc-
tures ; the basal third of the 2nd segment is closely and strongly
striated ; the gastrocceli are represented by three shallow depressioxs,
the space between them being finely striated.
Benyllus, gen. nov.
Median segment completely areolated, its apex spined ; the posterior
median area largely projecting into the areola ; its spiracles linear,
curved. Scutellum convex, rounded, its sides sharply margined.
Mandibles unequally toothed, the upper much longer than the
lower. Clypeus not separated from the face. Occiput not margined.
Wing areolet 5-angled, narrowed above ; the transverse basal nervure
interstitial. Legs stout, the tarsi long, spinose. Abdomen with
seven segments ; the apex of the petiole curved ; the gastrocceli almost
obsolete ; the ovipositor largely projecting ; the 7th segment is nearly
as long as the 6th; the ventral keel stout, extending to the apex
of the 5th segment—to the base of the ovipositor. Antenne stout.
A genus of <Amblypygi characterized by the stout,
unequally-toothed mandibles, the stoutly-keeled scutellums,
the spined median segment, and by the posterior median
segment area largely projecting into the areola.
Benyllus rufus, sp. nov.
Rufus, nigro maculato, facie, clypeo orbitisque oculorum flavis,
mesonoto nigro, alis hyalinis, stigmate fusco. 9,
Length, 10 mm.
Hab, Kuasta Hinus. Coll. Rothney.
Species of Ichneumonide from India. 233
Head black ; the face, clypeus, the inner orbits and the outer more
broadly, except above, lemon-yellow ; the palpi of a paler yellow ;
the face and clypeus, except at the sides, punctured and covered with
pale pubescence ; the front smooth and shining, depressed, except at
the tap, where there is, in the middle, a raised punctured band ; the
vertex is closely and distinctly punctured ; a furrow runs down from
the lower ocellus. Mandibles yellowish, black at the apex. Thorax
rufous; the mesonotum and the sides of the median segment at the
base, black. Mesonotum and scutellum closely punctured ; the post-
scutellum irregularly punctured and striated. The basal areze of the
median segment are closely punctured ; the areola obliquely, stoutly
striated on the sides ; the posterior median punctured all over, the
punctures clearly separated, but not deep; the outer arex irregularly
punctured on the upper half, on the lower transversely striated ; the
spinesare large. Propleurz smooth and yellowish in tint; the meso-
metapleurz and mesosternum closely and distinctly punctured ; the
mesosternal furrow striated. Abdomen of a slightly darker rufous
colour than the thorax ; a mark on the petiole—half on the petiole
and half on the base of the post-petiole—and the basal two-thirds of
the 2nd and 3rd segments, black ; the post-petiole and 2nd segment
closely and distinctly, the 3rd less distinctly punctured ; the sheath
of the ovipositor blackish. Legs coloured like the body ; the four
front coxz and trochanters pale yellow.
Buathra, gen. nov.
¢. Antenne long, stout, distinctly tapering towards the apex ;
the 1st joint of the flagellum distinctly longer than the 2nd. Eyes
prominent, parallel. Front and vertex deeply excavated. Clypeus
small, its apex rounded. Mandibles stout, with two stout, blunt,
subequal teeth. Parapsidal furrows, deep, distinct. Scutellum not
much raised, not keeled. Median segment rugosely punctured,
without areze, but with two transverse keels, bent backwards in the
middle ; at the sides these are united by a keel which ends in a
tooth, which is blunt, but distinct. Areolet almost square, hardly
narrowed in front; the disco-cubital nervure has a stump of a
nervure ; the transverse basal nervure is interstitial. Petiole
slender, the spiracles at the base of the post-petiole, which is not
much dilated. Gastrocceli indistinct. Metathoracic spiracles large,
oblique, linear. Clypeus roundly projecting.
I have only a ¢ of this genus, and am not sure about:
its exact systematic position. In the distinct parapsidal
furrows and in the squarish areolet it agrees with the
Cryptina, as it does also in the paucity of arez on the
234 Mr. P. Cameron on Twelve New Genera and
median segment; in the position of the spiracles on the
petiole, and in having the stump of a nervure on the
disco-cubital nervure, it agrees with the Jchnewmonina.
Buathra rufiventris, sp. nov.
Black ; the abdomen from the apex of the petiole and the legs
ferruginous; the antennal scape in the middle below ; the inner
orbits narrowly above, broadly below—the band becoming broader
beyond the middle and narrowed obliquely towards the apex—a
mark, rounded at the apex, in the centre below the antenne, the
clypeus, mandibles, except the teeth, the palpi, the outer orbits
narrowly, except above, the tecule and tubercles, yellow. The
four front cox and trochanters are yellow in front and at the sides ;
the middle trochanters are black behind ; the hind cox and base
of trochanters black ; the apical four joints of the hinder tarsi
spinose. Wings hyaline, the nervures and stigma black. ¢.
Length, 15 mm.
Hab, Kuasta Hitis. Coll. Rothney.
Head densely covered with glistening, silvery-white pubescence.
Face strongly punctured. Median segment with a gradually
rounded slope to the middle, the apex oblique. Thorax covered
with glistening white hair, closely and strongly punctured, the
pleurz more strongly than the mesonotum ; the metapleure at the
apex coarsely obliquely striated. The punctures on the scutellum
large and clearly separated ; the basal depression large and with
some stout strie. Post-scutellum longitudinally striated in the
centre. Petiole strongly aciculated except at the apex, as are also
the sides and ventral surface.
AMPULEX KHASIANA, Sp. nov.
Length, 16mm. 9.
Hab. KHAsta Hints. Coll. Rothney.
Agrees closely with A. longicollis, Cam., but is smaller,
the prothorax is not quite so long nor so distinctly narrowed
at the base; the apex of the median segment is not
reticulated at the end of the transverse striations; the
fovea at the apex is neither so deep nor so clearly defined,
the narrowed basal part is longer and more distinctly
separated, and the bordering nervure on the radial nervure
is not so curved at the apex.
Antennz black, moderately stout, the 3rd joimt about twice the
length of the 4th. Head dark purple, coarsely punctured above the
Species of Ichneumonide from India. 235
antenne, sparsely covered with pale hair. Clypeus smooth, blue
above, the middle sharply carinate, the apex with an oblique short
tooth on either side ; the labrum smooth, glabrous. Front keeled in
the centre, the keel bordered by some oblique one. Temples largely
developed, roundly narrowed. Prothorax smooth, longer than
broad, narrowed at the base, which is raised and distinctly separated ;
the middle of the pronotum forms almost a triangle. Mesonotum
with large, deep, scattered punctures; the furrows are narrow,
complete. Scutellumat the apex bordered with large, deep, irregular
punctures ; there being also a few in the centre; the sides of the
post-scutellam with two large punctures, The three central keels
on the median segment are distinct and extend to the base of the
apical third, where the central keel roundly bifurcates ; the central
space is stoutly transversely striated ; the inner lateral area is finely
and closely, the outer more stoutly and widely striated ; at the apex
are three transverse keels, the apical being more widely separated
from the posterior than the latter are from each other; from the
hinder keel another projects backwards into the bifurcated apex of
the central keel ; the projecting apex in the centre is smooth,
broader than long, and has the apical third depressed ; the lateral
teeth are large and somewhat triangular. The apex of the segment
has an almost perpendicular slope, is transversely striated, rather
irregularly at the base, more closely and regularly in the middle
and apex, where it is thickly covered with long white hair. Meso-
pleure stoutly, deeply, irregularly punctured ; the tubercles large,
smooth and roundly convex ; the metapleure with two stout longi-
tudinal keels ; the space between these is smooth at the base, the
rest with stout perpendicular keels ; the base in the centre is smooth,
its lower part and the apex stoutly, irregularly reticulated. Meso-
sternum irregularly punctured ; the process is long, narrowed at the
base, slightly narrower and rounded at the apex ; the inner side is
slightly raised, the centre hollowed, Legs black, the four hinder
cox green, the basal half of the hinder femora red ; their apex and
the hinder tibiz purple ; the tarsi thickly spinose ; the tooth on the
claws is placed shortly beyond the centre. Wings deep smoky-
violaceous ; the stigma and nervures black ; the apex of the radius
is rounded ; the appendicular cellule is almost open at the apex,
elongate ; the basal abscissee of the radius of equal length; the
Ist recurrent nervure is received in the middle, the 2nd near the
apex of the basal third of the cellule. Abdomen shining, very
smooth, purple ; the narrowed basal part of the petiole is as long
as the widened apical half ; the 2nd segment is slightly longer than
all the succeeding segments united. The sides and lower part of the
petiole are stoutly striated ; the strize on the sides are oblique.
236 Mr. P. Cameron on Twelve New Genera and
The pro- and mesonotum are largely tinted with green; the
metanotum is deep purple, blue at the base and apex and, to a less
extent, on the sides.
AMPULEX ASSAMENSIS, sp. nov.
Length,13mm. 4
Hab. Kuasta Hits. Coll. Rothney.
Antenne black, the apical joints perceptibly curved, the scape
thickly covered with pale pubescence; the 3rd joint slightly,
but distinctly, longer than the 4th. Head coarsely rugosely
punctured, the front transversely striated in the middle, the temples
obliquely narrowed ; the antennal tubercles stout, longish, narrowed
above, slightly curved; the temples narrowed. Clypeus green,
thickly covered with long white hair ; its apex ends in a triangular
tooth, there being no lateral ones. Labrum brownish, thickly
covered with long white hair. Mandibles black, the apex rufous,
the base punctured. Eyes only very slightly converging above,
where they are separated by slightly more than the length of the
2nd and 38rd antennal joints united. The thorax coarsely punctured,
the prothorax and sides of the mesonotum blue, the middle of the
metanotum indigo-blue; sparsely covered with long black hair.
The middle lobe of the median segment is stoutly striated ; the
striz distinctly curved, those at the base irregularly curved ; the
central keel is distinct; the outer are are more closely and regularly
striated ; the projecting apex is deeply hollowed, the hollow being
wider than long and bounded behind by three fovez, there being
another fovea behind the outer of these. The apex of the segment
is largely, irregularly reticulated, the upper reticulations being the
larger, the lower of the two teeth is small and triangular. Pro- and
mesopleure distinctly punctured, except on the middle of the former
and apex of the latter; the metapleure strongly reticulated. The
mesosternal process large, its sides oblique on the inner, rounded on
the outer-side. The four hinder femora are bright red ; the anterior
femora and tarsi of a duller red ; the femora are blue behind ; all
the tarsi black ; the hinder claws are stoutly bifid, the hinder claw
the stouter and shorter. The first two abscissee of the radius are
equal in length ; the Ist transverse cubital nervure is only indicated
slightly below. The narrowed basal part of the petiole is short, the
middle triangular and with a curved furrow on the sides at the base ;
it is sparsely punctured laterally ; the apex is more strongly punc-
tured ; the 2nd segment is more closely punctured all over ; the
apical segments are coarsely and deeply punctured below; the
Species of Ichneumonide from India. 23
punctures on the 2nd segment are more scattered, especially at the
base ; the other segments are coarsely and closely punctured.
AMPULEX CARINIFRONS, sp. Nov.
Length, 15 mm,
Hab. KHasta HILus. Coll. Rothney.
Agrees in coloration and closely, in some other respects,
with A. compressa, but may be known from it by the
parapsidal furrows being less clearly defined, by the raised
apex of the pronotum being smoother above and deeply
furrowed throughout, by the raised apex of the median
segment heing roundly depressed in the middle, and by
the 2nd abdominal segment being much more strongly
punctured.
Antenne black, the scape punctured, metallic green, the 3rd joint
not quite the length of the 4th. Head deeply punctured, the face
thickly covered with long white hair; the front and vertex with
brassy tints ; the front in the middle with an enclosed area, its keels
enclosing the front ocellus, the basal half of the area being broader
than the apical ; the keels bounding the latter are curved inwardly ;
the vertex has a furrow in the centre. Pronotum stoutly transversely
striated ; the furrows behind the middle are more widely separated ;
the apex is distinctly raised, smooth and furrowed down the middle.
Mesonotum with large, deep punctures which are more widely
separated in the middle at the apex ; the sides near the apex are
deeply hollowed, the inner-side of the hollow being stoutly striated,
the outer smooth. Scutellum more closely punctured than the
mesonotum ; the post-scutellum aciculated ; its sides, except at the
apex, with large punctures. Median segment transversely striated,
the striz on the middle more widely separated ; the middle keel is
straight, and reaches close to the apex ; the lateral are curved and
reach shortly beyond the middle ; the inner outer keel originates
from the outermost near its base. Tubercles stout, rounded at the
top; the keel bounding the apex is depressed in the middle. The
apex of the segment is irregularly reticulated all over and thickly
covered with long white hair. The upper part of the propleure is
strongly punctured, the lower smooth, faintly aciculated; the
mesopleure coarsely punctured except at the apex, where it is
aciculated ; the upper half striated below, reticulated above.
Mesosternum coarsely punctured, the central furrow not widened at
the apex as in A. compressa. Wings fusco-violaceous, darker along
the nervures, of which there are three; the cubital nervure is more
rounded upwards on the 2nd cubital cellule than in A. compressa,
TRANS. ENT. SOC. LOND. 1903.—PART Il. (JUNE) 16
238 Mr. P. Cameron on Twelve New Genera, ete.
and the 2nd recurrent nervure is received nearer the middle. The
base of the abdomen is distinctly narrowed and furrowed down the
middle. The base of the petiole is distinctly narrowed and furrowed
down the centre; the apex and the 2nd segment are distinctly
punctured all over, but not closely, the apical segments are more
closely and strongly punctured. The four hinder cox are brassy ;
the four posterior femora bright red, black at the apex ; the anterior
femora green, dull rufous in front, the tibiae are black in front, blue
or green behind ; the tarsi black ; hinder claws with a tooth at the
base. The eyes at the top are separated by the length of the 4th
antennal joint.
As the above concludes the descriptions of the new
species of Ampulex known to me from the Khasia Hills, it
may be useful to give a list of the Khasia species described
by me. In the Ann. and Mag. Nat. Hist. July 1899, p.
57, Ampulex (Rhinopsis) nigricans ; lc. 1890, pilosa, p.
37; pulchriceps, p. 38; lc. 1902, Rothneyi, p. 545 trichio-
soma, p. 55; ruficoxis, p. 56, /.c. 1903, montana, p. 319;
himalayensis, p. 320; interstitialis, p. 321; longicollis, The
Entomologist, p. 263, 1902; trigona, p. 264; brevicornis, p.
312. Described here, khasiana, assamensis, and carinifrons.
The known species from Khasia are latifrons, Kohl, and
compressa, Fab., in all seventeen species.
Bingham (Fauna of British India, Hym.) describes eight
Indian species only, but one of these (Cognata, Kohl)
appears to have been included in error. Cf. Cameron,
Ann. and Mag. Nat. Hist. 1900, p. 39.
JUNE 2, 1903.
XIV. On a Collection of Lepidoptera from Arctic America.
By Henry Joun ELweEs, F.R.S., etc.
[Read February 4th, 1903.]
PLATE IX.
THE collection of which I give a list was made by Mr.
David Hanbury, who has appended notes on the localities
and habits of the insects, which give an exceptional value
to it.
Though small in number of species, 1t is the most
interesting Arctic collection I have yet seen, and most of
the specimens are in beautiful condition. The variation
in some of the species is extraordinary.
Considering the difficulties under which collecting is
carried on in such a region, and that Mr. Hanbury had
not previously any experience in collecting, this collection
does him the highest credit.
He has been good enough to present the greater part
of it to the National Museum.
List oF Mr. Hanspury’s ArcTIC COLLECTION.
1. Hrebia fasciata. (Plate IX, fig. 11 g, 12 2.)
E. fasciata, Butler, Cat. Sat. B.M., p. 92, Pl. II, fig. 8
(1868).
Several specimens in beautiful condition ; from Point
Epworth, 11, vii; Cape Barrow, 30, vi ; Chapman Island,
27, vi; Gray’s Bay, 1, vii. These agree with the type in
the British Museum from Cambridge Bay, and vary con-
siderably in the amount of rufous in the fore-wing above,
which in the females extends to the base of the wing.
The fringe in quite fresh specimens is grey.
2. Hrebia disa.
Papilio disa, Thunberg, Diss. Ins. Suec. il, p. 37
(1791).
Three males and a female from Point Epworth, 11, vii.
These resemble specimens from Finland much more closely
than they do specimens of the var. mancinus, Hew., from
TRANS. ENT. SOC. LOND. 1903.—PART II. (ocT.) 17
240 Mr. H. J. Elwes on a
Alberta, in having the band of the hind-wing below well
marked.
3. Hrebia rossit.
Hipparchia ross, Curtis, Ross’ 2nd Voy. App. Nat.
Hist., p. 67, Pl. A, fig. 7 (1835).
A pair from 140° W., 67° 40 N., 14, vii, and one from
Point Epworth, 11, vii, are perfectly fresh, and. seem to
show that this species is barely separable from the Asiatic
form, evo, Brem. Cf. Trans. Ent. Soc. Lond., 1899, p. 347.
I previously had only bad specimens from Hudson Bay
for comparison. Recently I have received a fresh female
taken by Mr. Sampson in Frobisher Bay, Baffin’s Land,
14, vii, 02.
The fringes of these three are all grey, which is not the
case In any of my Altai specimens however fresh, though
slightly evident in some from Transbaikalia.
4. ners bore, var. taygete.
(H. taygete, Hibner, Samml. Ex. Schmett. (1816-
1824).
Several pairs in beautiful condition from Barren Grounds,
Gray’s Bay, and Point Epworth; vary a good deal in the
breadth, shape, and distinctness of the bands on hind-wing
below. ‘Two show the marginal row of whitish spots on
hind-wing very distinctly, these are usually faint or absent
in Labrador specimens.
5. Gneis semidea, var. vel crambis, var. (Plate IX, fig.
9 2,10 9)
Hipparchia semidea, Say, Amer. Ent. IIT, Pl. 50
(1828).
Chionobas crambis, Freyer, Neuere Beitr. V, Pl. 440,
figs. 3-4 (1844).
Five specimens from Barren Grounds and one from Point
Epworth, fresh and in good order, must, I think, be referred
to one of these species. I might call them peartiv, Edw.,
or assimalis, Butl., but they are intermediate between the
types of those two forms in the British Museum, being
rather less conspicuously banded on hind-wing below than
the former, and rather more so than the latter.
Some of them show more or less trace of the marginal
row of grey spots on hind-wing, which at first led me to
Collection of Lepidoptera from Arctic America. 241
think that they were cranbis; but in a fresh state they
are much blacker than any of the faded specimens of
crambis I have before me. Whether that species, which I
only know certainly from Labrador, occurs also in Arctic
America, and whether when we know it better it will be
possible certainly to distinguish it from semidea, are points
which at present remain obscure.
6. Caenonympha tiphon, var. niaturata.
C. tiphon, var. mixturata, Alpheraky, Rom. Mem. sur.
Lep. IX, p. 326 (1897).
Two males and a female from Dismal Creek, taken
30, vil, are in bad condition, but are sufficient to show
that the form found here, like that from Alaska, is nearer
to the Kamschatkan variety than to any other.
7. Argynnis pales,
Papilio pales, W. V., p. 177 (1776).
Three males and a female from the Barren Grounds,
taken 16 and 18, vi, are the first specimens of this species
I have yet seen or heard of from America, where I have
long expected to hear of its discovery.
The males are quite typical, and could not be dis-
tinguished from some Alpine specimens.
The female is like some I have from Northern Siberia.
8. Argynnis polaris.
A, polaris, Boisduval, Ind. Meth., p. 15 (1829); ad.
Icones, Pl. XX, figs. 1, 2 (1833).
Specimens were taken in all the localities visited in the
first half of July, and are quite typical.
9. Argynnis chariclea. (Plate IX, figs. 6, 7, 8.)
Papilio chariclea, Schneider, Neuest. Mag. V, p. 588
(1794).
The most extraordinary variation is shown by the
specimens of this species, which occurs in all parts of
Arctic America, and was taken by Mr. Hanbury at all the
places where he collected.
Among them a male from Chapman Island is almost
black. Another from Dismal Creek is very small and
pale, but a female from Point Epworth is a wonderful
242 Mr. H. J. Elwes on a
aberration, and I cannot say positively whether it 1s
polaris or chariclea, though the size and the shape of the
wings indicate the latter species.
10. Argynnis frigga, vax. umproba.
Papilio fr igga, Thunberg, ¢.¢., p. 33.
Argynnis vinproba, Butler, Ent. Mo. Mag. XIII,
p. 206 (877), ;
Several from the Barren Grounds and one from Point
Epworth are like the type, and show but little variation.
ll. Lycena orbitulus, var. franklinit.
Papilio orbitulus, Esper, Schmett. I, 2, Pl. CXII, fig.
4 (1800).
Lycena franklinii, Curtis, t.c., p. 69, Pl. A, figs. 8, 9.
A pair from the Barren Grounds are not so distinct
from the Arctic form found in Europe, var. agwilina, Ster.,
= aquilo, Bdv., as those from Labrador, and are perhaps
nearer to those I have taken in the Rocky Mountains
near Laggan.
12. Oolias hecla.
C. hecla, Lefebvre, Ann. Soc. Ent. Fr., 1836, p. 383,
Pl. IX, B, figs. 3-6. -
Four males and three females from the Barren Grounds,
114° W., 67° 40 N., 13-16, vii. Agreeing well with other
specimens from Arctic America, some of which were
called glacialis by McLachlan.
Staudinger now catalogues the Lapland form as var.
sulitelma, Auriv. The specimens in Mr, Hanbury’s collection
differ inter se to a remarkable extent in the colour of the
borders and discal spots of the wings above.
13. Colias boothii. (Plate IX, figs. 1-4 2,5 9.)
C. boothit, Curtis, t.c., p. 65, Pl. A, figs. 3-5.
This was represented by several fresh specimens, which
enable me to confirm the opinion formed on very in-
sufficient previous knowledge, that it is a species perfectly
distinct from the last. The variation in this species 1s so
great that I have had to figure five specimens to give a
fair idea of it; some of them would be supposed by their
markings to be females, but though the abdomens are
Collection of Lepidoptera from Arctic America. 243
difficult to examine, owing to their hairy covering and
being somewhat compressed in packing, I can find only one
undoubted female among them. None of the specimens
sent are quite what Curtis figures as chione, in which the
marginal band is faint or absent.
The species seems to have been fairly common at Point
Epworth, Gray’s Bay, and on the Barren Grounds.
14. Colias pelidne.
C. pelidne, Boisduval, Icones, Pl. VIII, figs. 1-3.
Three pairs from Point Epworth, Barren Grounds, and
Dismal Creek, of which the females differ dnter se a good
deal, one being white and two lemon-yellow.
15. Colias nastes.
C. nastes, Boisduval, /.c., figs. 4, 5.
Four males and two females from Barren Grounds, all
varying. These might be called rossti, Guen., or moina,
Streck., by those who hke to try and distinguish local
forms, a very uncertain task in the case of Arctic insects.
Heterocera by Sir GEorGE F. Hampson, Bart., B.A., F.Z.S.
NOcTUID&.
THypsophila zetterstedti, Ster., 4° 67 40".
GEOMETRID®.
Aspilates orciferaria, Wik., 114° 67° 40°.
Cidaria, sp. 114° 67° 407,
TORTRICID A.
= = ni .
gen. sp., Point Epworth.
FIG.
1
2
355
4D a5;
Os;
6
iis
8.
9.
1. 5,
11
EXPLANATION OF PLATE IX.
oP)
”)
”»
”?
oP)
. Hrebia fasciata, B
. Colias boothii, Curtis, ¢. Point Epworth.
¢. Barren Grounds.
3.
bP]
9)
gd. Gray’s Bay.
Oe
. Arygynnis chariclea, Schn., ab.
P
utl.,
9
Point Epworth.
Bathurst Inlet.
Dismal Creek.
Barren Grounds.
Point Epworth.
Cape Barrow.
Point Epworth.
XV. A contribution to the life history of Orina (Chryso-
chloa) tristis, Fabr., var. Smaragdina, Weise. By
THomMAs ALGERNON CHAPMAN, M.D., F.ZS.
[Read April Ist, 1903.]
PLATES X AND XI.
IN our observations on some species of Orina presented
to the Society by Mr. Champion and myself in December
1900, besides several viviparous species, we noted Orina
tristis to have a habit that was not viviparous, nor yet
strictly oviparous in the ordinary manner. Unfortunately
our material of this species consisted of only one female
beetle and our observations were accordingly somewhat
restricted, so that I was pleased to meet with the beetle
in some abundance on May 30th, 1902, near Pino, on Lago
Maggiore. The beetles were disporting themselves, often
a good number together, in the sunshine on a rather tall
upright species of Centaurea that looked otherwise very
like our nigra. I brought a number of beetles home and
had them alive throughout the summer. They laid eggs
freely, but began to die off about the middle of September.
In the first week in October egg-laying seemed to be
stopped, but a small batch of seven was laid October 18th,
at which date only that number of beetles remained alive.
Two males were still living on February 7th, 1903. I
noticed in several beetles that died that the posterior
tibiz and tarsi were damaged or wanting and must have
been eaten by the other beetles; whether this was a
cause of death or only occurred after mortal illness had
supervened I do not know. I twice found a beetle so
damaged, that was still very decidedly alive.
I may note parenthetically that at the end of June we
(Mr. Champion and myself) met with Orina rugulosa at
San Sebastian. We found both beetles and larvae, and
observed the egg-laying to follow almost precisely the
same methods as in 0. smaragdina, indeed the eggs were
almost indistinguishable from each other (I had some of
the latter with me). They were on a very nigra-like
species of Centaurea; the impossibility of obtaining
TRANS. ENT. SOC. LOND. 1903.—PART III. (OCT.)
246 Dr. T. A. Chapman’s
Centaurea at the places we visited in Spain brought to
a very early conclusion my attempts to rear the two
species side by side.
The account given in 1900 of the egg of O. tristis and
of the hatching of the larva is correct as far as it goes,
but wants much supplementing. When the larva is ready
to emerge it shows very visibly through the egg-shell
three black patches on either side, just above the spiracular
line, on the 2nd and 83rd thoracic and Ist abdominal seg-
ments; except the spiracles, eyes, some mouth parts, and
finally the cutaneous hairs it is otherwise colourless.
These three patches are apparently ordinary portions of
the cuticular covering that become hardened and matured
at this early period for a special function, just as the Ist
spiracular region and some other parts are solidly chitinised
with special objects in various Lepidopterous pup, whilst
the rest of the pupal integument is still soft. These three
hard black patches are slightly convex and project centrally,
but I cannot make out that they have any sharp or
angular point.
Their function is to determine the lines of rupture of
the egg-shell. The force used to open the egg-shell is
one that is in common use with many insects, especially
for emergence from the pupa. Here it 1s used for emerg-
ence from the egg. It is the secretion of air into the
interior of the larva; the large air bubbles look as if in
the general body cavity of the insect, and I am not positive
that they are not, but as some air often escapes by the
mouth and anus, when the insect is flattened, I incline
to believe it is in the alimentary canal. The effect of
this air being so secreted is to enlarge the bulk of the
larva and exert a bursting tension in the egg-shell. If
this tension became sufficient the shell would burst by
explosion in some irregular manner, but the presence of
these three hard points on either side of the larva is to
increase the tension along a line passing along them, and
so, long before the strain is sufficient to produce an explo-
sive rupture, the egg-shell splits along each side over these
points. The modus operandi is in principle precisely the
same as that by which the pupz of the Lepidopterous
genera Limacodes and Hriogaster open their cocoons; in these
a pressure, produced by air secretion within the insect, is
localised as to the severest point of strain by a pupal
beak, and so a lid splits off at the so-determined line.
Contribution to the life history of Orina tristis. 247
When the egg-shell is split, a portion of the larva slightly
protrudes through each slit, but the larva soon manages
to turn round and escape through one of them head first.
Since the larva can emerge through only one of them,
why should there be two slits?) These do not reach the
top of the shell and are still further from reaching the
bottom, but now and then they deviate from this symmetrical
arrangement, one may be longer than the other, and this
deviation may account for one slit only existing and that
one reaching to the top of the egg-shell. This is com-
paratively rare, but still not infrequent, and was the only
form I noticed in the few eggs I had in 1900. The two
slits are probably useful in affording the larva escape by
one, when the other is obstructed by the egg being
against one of its neighbours, or possibly even glued to
it or some other object, by some accident, either in the
disposal of the parental glue that fastens the egg to the
surface on which it is laid, or by a neighbouring egg
being injured and its contents acting as a cement, as
must not infrequently happen to eggs laid naturally on
a leaf. The eggs are laid in regular order, in several
adjacent rows, and are slightly inclined to the surface of
attachment, instead of being perpendicular to it, so that
one side of the egg is tolerably free, the other almost or
even actually touches its neighbours. The larva could
hardly hope to escape, if a solitary slit occurred on this
side, I had made many observations of these beetles, of
their egg-laying, of their hatching, had fed many of them
from hatching to full-fed larvze, and was still of the opinion
that this species could not be 0. tristis, because I had
observed O. tristis lay eggs quite ready to hatch, whilst
my insects from Pino laid eggs quite undeveloped. I was,
however, puzzled to find that some eggs hatched in a few
days whilst others did not do so for over a week. Some
showed the imaginal jaws through the shell very early
and soon. Still all the eggs I examined when new laid
showed no coloured indications of the young larva.
On August 29th, however, a new experience awaited
me. I found several eggs that had been laid since the
26th were already hatched. I determined therefore to more
closely scrutinise the periods from laying to hatching of
future eggs. Further light was closer at hand than I had
expected. At 11 am., August 29th, the beetles were fed
and as usual placed in a clean jar with fresh food, clean paper,
248 Dr. T. A. Chapman’s
etc. At 2 p.m. I found three small batches of eggs laid,
two presenting the usual clear aspect, but the third, which
the beetle was at the moment depositing, consisted of two
egos that were actually hatching and of six others which
presented the imaginal eyes well developed, but no jaws,
spiracles, or other larval parts visible.
The eggs that I had supposed to be quite undeveloped,
usually presented a rather opaque aspect, except at the
unattached (head) end, where about one-fourth or one-
fifth of the egg was apparently occupied with clear yellow
fluid, and it had struck me as curious that the eye-spots,
which were the first imaginal parts to be seen, occurred
on this transparent area. I now, however, understood that
this transparent region had no relationship to a similar
one that occurs in the newly-laid eggs of Zygaenids
(Anthrocera) amongst the Lepidoptera, and that it was
really the larval head, after considerable development of
the embryo had taken place. Most of the eggs are laid
at this stage, when the embryo is well-developed but
before the eye-spots appear. I now carefully examined
other eggs and found some with a fairly uniform aspect,
from end to end. I saw several such eggs actually laid,
and examined them immediately, and found I could treat
them as transparent objects sufficiently well to see that
the greater part of their contents were in small rounded
masses, but that at the free end there was within the
shell a delicate membrane and inside this several large
rounded processes, one nearly half the diameter of the egg
across, another smaller, with a clear angular space between
them, and smaller ones extending down the egg. I make
no doubt these are the early cephalic lobes of the embryo
with further segmental divisions.
So far as I have noticed, these are the youngest eggs
that are laid, the greater number are rather more advanced,
and they may even be so far advanced as to show the eye-
spots, and as an exceptional occurrence, they may be just
ready to hatch.
T regret that Iam ignorant of the methods of examining
the younger eggs, and of overcoming the difficulties pre-
sented by the tough leathery egg-shell and the very
delicate contents, but I feel satisfied that what I have been
able to observe justifies the conclusion just stated, that
the eggs have always undergone some development when
laid and sometimes have made a near approach to hatching.
Contribution to the life history of Orina tristis. 249
_Those of 0. tristis I watched during August two years
ago were well advanced when laid, none were so this year
till towards the end of August, so that it may be, that
only towards the end of its egg-laying are the ova retained
until the larva is well-developed. But it is also certainly
the case that the ovum is not fertilised at the moment
of deposition, but at some interval previous to that time,
notwithstanding that the egg-shell is a solid structure
similar to that of eggs that have (as, in fact, these always
have) to exist externally for some time before hatching.
They have, in fact, taken the first step towards becoming
viviparous, and it is of interest to note that the step is
not a definite and constant one, but varies from a habit
nearly that of ordinary egg-laying insects, to one that is
almost that of viviparity, not in different races of the
beetle, or even in different individuals, but apparently in
the same individual possibly according to its age. Not
certainly according to differences of treatment, as all my
beetles were kept together, yet their habits were not
always all alike.
It is interesting to notice that Mr. W. M. Wheeler in the
Journal of Morphology (Boston), vol. iii., 1889, figures
the young larva of Doryphora decemlineata as existing
within the egg-shell as almost identical in appearance with
that of O. tristis. The only difference I note in his figure
or description is that the hatching spines in D. decemlineata
have definitely sharp apices, those of 0. tristis being com-
paratively flat and blunt. He describes the hatching as
taking place by the contained larva moving within the
egg-shell so as to cut it with these spines, and says that it
cuts the egg on both sides and that these two incisions
meet over the apex, making one large opening. I should
certainly like to see this process take place; still I see no
reason to doubt that it is very possible for similar struc-
tures in allied species to exercise the same functions, or at
least to attain the same objects in very different ways. A
remarkable instance of this I mentioned lately to the
Entomological Society in the case of two species of Lepi-
doptera, Hh ybocompa milhausert and H. dryinopa, in which
a similar pupal spine is used in quite a different way in
each species for opening the cocoon.
In 0. tristis it 1s certainly the case that no movement of
the young larva takes place before the opening in the egg
is formed, and that it takes place by rupture under tension
250 Dr. T. A. Chapman's
from within, and merely the place where it shall occur, but
not the actual rupture, being determined by the hatching
spines.
It is certainly matter of some interest from a classi-
ficatory point of view, to find the egg and the embryo
within it and the method of hatching (hatching spines,
ete.), so very nearly identical in Doryphora decemlineata
and in Orina tristis, Wheeler's observations especially
relate to the development of the embryo; and as his
observations were made entirely on eggs already laid, and
he begins with the egg segmentation preceding the embryo,
it is clear that in Doryphora there is no quasi-viviparity as
in 0. tristis, There is, in his paper, no suggestion of the
possibility of such a thing. He would ‘certainly have
noticed and recorded it had there been any trace of it,
although it was not strictly the subject of his paper.
I believe Doryphora and Orina are not classified very
closely together. It may be that this method of hatching
is common to many species of Chrysomelids; if so, record
of it ought certainly to be more frequent than it is.
Immediately after each of its three subsequent moults,
the skin of the larva is quite colourless, exhibiting all its
interior anatomy, trachez, alimentary canal, etc., but it is
sufficiently loaded with yellow fat and fluids to make
details difficult to observe. It rapidly (in half-an-hour or
so) becomes black. At these moults there is no trace of
these hatching spines to be noticed. Two circumstances
attracted my notice; one was that the larva used inflation
to assist the rupture of the effete larva skin. This is not
pushed away towards the tail and so stretched in front till
it splits as is the method in Lepidoptera, but the whole
larva skin is fully on the stretch, and finally shts down the
dorsum, the new white larva protruding at once and then
creeping out slowly, leaving the empty larval skin slightly
contracted, but still looking uncommonly hke a living
larva, and nowhere with its segments crushed together.
It merely takes up the aspect of a younger larva with its
subsegmental ridges, when the tension of a full-fed larva
within it is withdrawn.
The larva when newly moulted and still colourless is
seen to be especially translucent in the thoracic region,
and several larger and smaller globules of air are seen to
1 Mr. Jacobi informs me that the Colorado beetle is not a Dory-
phora, but belongs to a genus very near to Orina.
Contribution to the life history of Orina tristis. 251
occupy this region; that they are in the alimentary canal
seems almost certain, as they escape by the mouth when
the larva is subjected to pressure.
The other circumstance connected with moulting that
seemed to me curious, is no doubt so, only because my
experience in watching the moulting of larve has been
almost entirely amongst the Lepidoptera; in these, the
larva always completely voids the alimentary canal at a
moult. A newly-moulted larva has no food contents at any
stage. In 0. tristis the anterior part of the alimentary
canal contains no food, but the hinder portion, in several
loops, in the abdominal segments, is more or less loaded.
The effete larval skin appears to be held in place so that
the larva may crawl out of it by a more or less half-dried
faecal deposit glueing the-anal extremity to the leaf.
The rupture in the larval skin divides the head into two
lateral (epicranial ?) pieces, and the clypeus with the mouth
parts and slits down the thoracic and two first abdominal
segments dorsally as if along a dorsal suture; the third
abdominal segment is often “slit in the same way, and I
have once seen the fourth slit also, but irregularly towards
one side, as if any dorsal suture was here certainly absent,
as it probably is in the third abdominal segment.
At the moult to pupa the skin collapses, and so far as
can be judged by the condition of the cast skin the process
of moulting is very similar to that in Lepidopterous larvie.
The eggs are not absolutely uniform in size and shape.
Their length i is about two-and-a-quarter times their width ;
their sides are not quite straight but curved, so that the
middle of the egg is the w idest part; sometimes the
longitudinal section is an ellipse, at others there is some
tapering towards one or both ends (as in the small end of
an egg).
The actual dimensions of four eggs measured was :—
owes) eal
eas PRIA 5 ABV A ana
Wider Osage ao 10;
25
94 ofan sO Pa) 7 O97 mm.
It is to be noted, that as the embryo is always developed
with the head to the free end of the egg, and has already
advanced in its development so far as to already have its
head to that end when laid, the orientation of the embryo
in the egg must be determined by its relations to the
ovarian tubuli in which it develops, and as these are coiled
in all directions, gravity has no share in the determination,
252 Dr. T. A. Chapman’s
but simply which is the end of the egg nearest the top of
the tube.
Though a dissected beetle shows the tubules loaded
with eggs of all sizes, my observations go no further in
this direction; it would be of interest to know whether
the eggs attain their full size before they are fertilised and
development begins, or whether they grow after develop-
ment of the embryo is begun. Probably not, but as the
embryos of the viviparous species grow considerably whilst
still in the tubules, only definite observation can determine
what happens in this species.
When hatched the larva of O. tvistis 1s colourless, except
for the eyes, jaws, spiracles and the six black spots that
may be seen through the egg-shell, but it very soon
becomes quite black, with the exception of the fine brown
hairs which are plentifully scattered all over it. The head
and prothorax are at this stage exceptionally large,
forming fully two-fifths of the length of the larva. The
legs are nearly colourless and the under-surface is pale,
the larva is blacker than at later stages, the blackness but
encroaching a long way on to the ventral surface.
The intersegmental depression divides the segment
deeply, but terminates some way above the spiracle, the
anterior and posterior subsegments meeting as a raised
ridge, below which the spiracle lies in a depressed
triangular area that extends to the front of the segment.
Length 2°0 mm., greatest breadth 1:0 m.
At the first moult, 7.e. in the second instar, the head
and prothorax remain dirty yellow for some time after the
abdomen has become black, and they remain a little paler
throughout the instar. The subsegments still meet in a
definite ridge above the spiracle, which is now hardly in a
depressed area, but has the diagonal grove separating Ist
and 2nd flange below it; the anterior branch of the sub-
segmental incision exists, but is not at all pronounced or
visible in all attitudes of the larva, Length 3:0—4:0 mm.,
width up to 2°0 mm., when contracted is almost a sphere
of 2°5 mm. in diameter.
In the third instar the larva is very similar to the
second, the head and prothorax are still a little paler than
the abdomen. The branch of the subsegmental incision
is very short, but distinct in the 2nd instar, and is
very similar in the third. Length 5:°0—8:0 mm., width
2°5—3'0 mm.
Contribution to the life history of Orina tristis, 258
The full-grown larva has the general aspect and structure
of Orinas, as described (Ent. Soc. Trans., 1901, p. 18). The
thorax is relatively smaller than in some other species, and
looks even less than it is, by comparison, being of the same
dark colour as the rest of the dorsum, without any kind of
paler tinting. The colour of the dorsum is black with a
bronzy shade, due either to texture and polish, or also
perhaps to a certain amount of the olive or brownish tint
that appears laterally and predominates ventrally, declaring
itself in the finer depressions of the minute wrinklings.
These wrinklings are apparently the same as in the other
species described. They are generally in transverse lines,
each elevated line being somewhat broken by partial or
complete sulci in more or less diagonal directions which are
sometimes independent, but are sometimes the transverse
sulci anastomosing. So far as it is possible to count them,
there are about thirteen or fourteen transverse ridges on
each segment. The transverse depressions, which are
most marked about the fourth or fifth abdominal segment,
and indicate subsegmentation of the segments, and
varied somewhat in each species previously described, are
well marked, and are probably characteristic. They smooth
out nearly completely in some attitudes. Dorsally it is
about central, the anterior margin highest. Rather lower
than half-way down to the spiracle, it sends back a branch
from its anterior margin, that fades out before reach-
ing the dorsum. The spiracle is a minute raised ring
surrounded by a small smooth surface; just below this is
a very definite diagonal groove, passing backwards and
ventrally. In certain attitudes the dorsal subsegmental
groove passes down as little more than a fine line in front
of the spiracle, and seems continuous with this diagonal
groove. This diagonal groove is in fact the demarcation
between the spiracular and subspiracular flange. The
segmental incision in the subspiracular flange is double, a
sulcus from below passing up behind one descending from
the dorsum. The marginal (3rd) flange is simple, its
upper and lower grooves being longitudinal.
The colour of the lower surface is yellowish-brown, over-
laid by a clouding of small blackish spots. The head and
prothorax has a tolerable coating of fine brown hairs; these
have to be looked for, the surface having at first glance a
glabrous aspect. After they are seen, one decides that the
254. Dr. T. A. Chapman’s
abdomen really is glabrous, till a closer search shows that
it carries a large number of fine brown hairs, less than half
the length of those on head and prothorax.
The legs and incisions are of the same olive-brown
already noted.
Length 10-14 mm. Width head 2°3 mm.
prothorax 4:0 mm.
mesothorax 3°8 mm.
5, Oth abl. '5°0=6:0 mm:
The colour of the larva varies a good deal. Taking it
as typically black above and pale luteous olive below, the
whole of the mouth region of the head belongs to the pale
lower region; the demarcation between the two regions
being the lower margin of the subspiracular flange (middle
ridge of the triple marginal flange). In many specimens,
however, the anterior portion of the larva, especially the
prothorax, is dorsally paler, without being pale enough to
suggest the yellow colouring this segment has in some
other species. The marginal area may also be a little
more invaded by the ventral paleness.
The blackness of the dorsum is also variable. In a
majority perhaps it is a brownish-olive, approaching black,
witb a bronzy aspect in some lights, its paler tones being
due to lighter colourmg (or comparative thinness of
pigment) in the bottoms of the minute sulci of the finely-
wrinkled surface. In some specimens the blackness is
dense and undoubted over the whole dorsum.
The most remarkable fact as to the colouring observed
was as to two specimens, differing apparently in no way
from any of the others, that formed part of a number
turned out in the field: these two were placed on open
flowers of the Centaurea when goimg out, and were still
there half-an-hour later when returning, but in the sun
were now of nearly as bright a metallic green as that of
the imago. I have been unable to induce others to exhibit
this colouring.
According to its attitude the segmentation of the
larva looks rather different. The prothorax is of course
always very evident, and when the full-fed larva is
active and lively the mesothorax is a good long segment,
much like the others, but with the two ridges lower,
especially the anterior one. When at rest, however, and
especially if the larva be a little sulky and only half-grown,
bb)
Contribution to the life history of Orina tristis. 255
the mesothorax partially or wholly disappears beneath the
prothorax, which may even slightly overlap the metathorax.
The 2nd thoracic segment is in these circumstances easily
overlooked, and owing to the backward direction of the
cox it 1s even difficult to believe that the 2nd and 3rd
pair of legs do not actually belong to the 3rd thoracic and
Ist abdominal segments, taking each of these in fact for
the segment in front of it.
The 7th abdominal segment is the last coloured like the
others, and has the appearance of a broad anal plate, the
8th is somewhat coloured, but is simple in structure and
retractile, the 9th and 10th are colourless and retractile,
the 9th forming a foot used in all progression.
The hairs, which seem to disappear as the larva passes
to its later instars, do not in fact do so. On the contrary
they persist, though the larva grows however, they do not,
but maintain almost precisely the same size and distribu-
tion they had in the Ist instar. Their length in the Ist
instar is tolerably uniform all over the larva, viz. ‘25 to
‘30 mm. in length. In the 2nd and 3rd instars they are
almost precisely the same, but of course make relatively a
much less show on these larger larve. In the last instar
they remain of almost identically the same length on the
head, over the dorsum and posteriorly, but laterally they
have actually dwindled to a length of only from 0°06 to
0°20 mm.
The hairs on the appendages and many of those on the
head appear like the ordinary tactile hairs terminating in
a solid sharp point, but the great majority, and nearly all
those on the general surface, are expanded at the tips, and
appear to be hollow, making the extremity trumpet-shaped,
but not so widely expanded perhaps as this suggests.
The arrangement of the hairs may be described as
irregular, merely, no doubt, because it is too complex to
describe ; on the 2nd thoracic and following segments they
are placed on the summits of the two subsegmental ridges,
on 2nd thoracic almost in one row on each, on 3rd the row
on front ridge is a little irregular, on the back one they are
better described as in two rows. On the front ridges of
the Ist and 2nd abdominal they are in two rows, the hairs
in the rows alternating, they are more numerous on the
back ridges, whilst on both ridges of 5 and 6 they may be
described as in three or four rows, but in all cases without
TRANS. ENT. SOC. LOND. 1903.—PART UL. (ocT.) 18
256 Dr. T. A. Chapman’s
the strict regularity that the idea of rows implies; they are
wanting near the incisions, both segmental and subseg-
mental. The 8th and 9th abdominal segments possess
hairs, especially a row of strong ones along their hind
margins, and 10 is not without finer hairs, Beneath the
abdomen each segment possesses shorter hairs, about ten
on each side, placed with some but not absolute bilateral
symmetry. The subspiracular flange on each segment has
a separate chitinous plate carrying four or five longer and
four or five shorter hairs.
The hairs on the prothorax are very numerous, and
except a certain regularity round the margin, are equally
but irregularly distributed over the whole surface, they
number about 110 on either side.
The pupa of O. tristis is 10 mm. long and 4°5 deep from
back to front across the 3rd abdominal segment, and 6°5
wide from knee-joint to knee-joint of 2nd legs, but this
when they are a little set forward, the natural position
would give perhaps nearer 5°3 mm. This is from a small
specimen, a larger one would probably be 12 mm. long.
The colour is pale whitish tending to flesh-colour, and
deepening as the beetle matures.
The head is well bent forward, 2. ¢. ventral, so that the
face is directed ventrally, and the abdomen is also curved
forwards, so that the ventral line is nearly straight, running
from the frons down the clypeus, labrum, labium, tarsi to
the anal point. The dorsum, on the other hand, presents a
curve from one of these points to the other, and distant
from it (as above noted) 4°5 mm. at the widest point.
The antennz pass first backwards then down behind
the Ist femora, their apices pointing forwards between the
1st and 2nd knee-jomts. The wings pass down between
the 2nd and 3rd legs, and curving round to the front, hide
the greater part of the 3rd femora and tibiw, and do not
meet in front by about 26 mm. The first wings (elytra)
show several longitudinal ribs or veins of which one on
either side is marginal, and three over the centre are more
marked than the others. The 2nd wings (wings) are visible
dorsally in a narrow slip passing under the Ist wings,
and their apices also appear beyond the first over the 3rd
tarsi.
The jaws, labrum, labium with palpi, the maxillary palpi
of three (?) joints passing straight backwards are very
Contribution to the life history of Orina tristis. 257
distinct, but these, with the legs and wings, are thickened
and fleshy (compared with the imago), but present no
spines or other definite armature except a few fine brown-
ish hairs on the upper outer exposed aspects of the femora.
The ventral aspect of the abdomen is equally smooth, but
dorsally all the segments have a greater or less supply of
short pale brown hairs. Over the front of the head and
the prothorax these are especially regularly and thickly
scattered. The prothorax is a large, somewhat square plate,
55x45 mm., as in the larva and imago. The meso-
thorax is narrow, 13 mm., witha projecting angle at centre
of its posterior margin ; just in front of this is a small
raised boss, which carries a few, the only, hairs on the
segment. The metathorax is again rather wide, 1°7 mm.
(the projecting point of mesothorax makes these measures
unduly favourable to that segment), with a dorsal longi-
tudinal grove, it is very smooth and polished, and possesses
only some half-dozen scattered hairs on each side, two on
either side near the middle of the hind margin being
especially strong.
The first abdominal segment 1s narrow (about 0°6 mm.),
and has a central dorsal grove, which on the second is
replaced by an elevation, the remainder having no dorsal
peculiarity. The first six abdominal segments are narrow
(about 5-0 mm. together), each however rather broader
than the one in front. The next is longer (2:0 mm.) “
is apically produced to an angle. Each of these (1-7
abdominal) has a spiracle, a small round chitinous ring,
about 1:0 mm. across, below which the dorsum of the
segment is rounded off, forming a lateral flange. 1-6
carry a fringe of hairs along their posterior margins, which
are carried round the margin of the flange, and where more
especially they are less a fringe, and more uniformly
scattered over the surface. Abdominal 7 has hairs scattered
all over it except its extreme anterior margin.
Dorsally 8 and 9 (abdominal) rapidly taper to a fine
brown chitinous point with which 9 terminates, 9 has a
very few inconspicuous hairs, on 8 they are more numerous
and evident but less so than on 7.
Ventrally 8, 9, and 10 are marked by sundry elevations
no doubt marking the genitation.
On March 4th, 1903, there still remained alive two
males of the beetles taken at Pino on May 30th, 1902. It
is not, however, the case that these represent the amount
bo
On
o2)
Dr. T, A. Chapman’s
of survival amongst the beetles taken. Comparing these
with the numbers that died, and eliminating those that
escaped, that were given away, or were otherwise disposed
of, these two represent the survivors rather of from eight
to ten beetles living in August 1902.
About this date (March 4th, 1903) I examined the pot
in which a number of larvee had gone down in September
and October, and was vexed to find that the earth was
apparently quite dry, and two larvee that I unearthed from
near the surface were apparently quite dead and dried up,
with the ventral and dorsal surfaces almost in contact.
Placing these in warm water, after twenty-four hours both
were alive and active. Though still looking somewhat
starved and contracted, they obviously had a good deal
of tissue within their skins.
I was, therefore, encouraged to damp the pot of earth
and put it in a warm place. Unfortunately again, I
overdid this.
On March 23rd I found a perfect beetle had emerged,
and forthwith examined the pot and contents. I found
it contained several beetles ready to emerge, one living and
one dead pupa, a number of dead larve, distended almost
to bursting by excess of imbibition but really close on
pupation, and others that had died at an earlier stage.
Disappointing as this result was, disappointing also
because so much due to my own want of intelligent care,
still, it enables me to say that I have reared the beetle
from the egg, and also to describe and figure the pupa.
The hibernating chamber of the larva, in which it
pupates in the spring, remaining probably as a pupa
from two to four weeks, according to temperature, 1s
nearly spherical, and about 2 inch in diameter, with the
interior wall pressed smooth.
The following data, giving dates of hatching of batches
of eggs laid at the same time, give some indication of the
various degrees of maturity of the eggs at the date of
laying.
Laid September 22-23 Laid September 12-13
3 hatched ae 26 1 hatched 3 15
pee fe 28 oe fe 17
22, a 29 1 a f 18
OW, cs, October 1 OW) 3 tas Way, os 23
|
Sie, 30 | geen Dy, Dame 21
leaving 14 still to hatch. | 5 mature eggs remaining.
Contribution to the life history of Orina tristis.
Laid September 17
9 hatched fs 24
Gor to ad 26
15 ” ) 27
2 i) 38
2 addled
Laid August 5-6
1 hatched He 8
2 ’9 ” 10
1 ” ” 13
We C4215
10 left of which the last
hatched ily
Larve 3 have moulted
Ist on August 19
All e 24
4 have moulted
2nd on August 26
All on 29
Many have moulted
3rd on September 4
Most 5 7
All 3 9
Nearly all full-grown 15
259
Laid September 13-15
1 hatched 3 17
1 ” #) 18
12 have died and blackened
that were ready to hatch
when laid.
5 hatched September 21
a a 23
ees ” 24
4 ” ” 26
After no eggs laid for 10
days, 7 were laid on
October 18th, 1 near hatch-
ing and 6 with top end
transparent. These show
eye-spots on 20th on which
date one more egg laid
with all spots visible.
The following actual notes show the time taken by the
larvee to feed up.
Batch laid August 12
4 hatched ie WW,
Many ° 19
Several lst moult ,, 24
All : s 29
Several 2nd moult ,, 31
Most 5, september 4
Some 3rd moult a 4
Nearly all full-grown ,,
Hatched August 12
Most in 2nd skin __,, 19
Six in 3rd skin ‘ 26
Some in 4th skin ,, 29
Nearly all in 4th skin 31
All September 4
Full-fed = 9
Hatched August 3-4
1st moult . 12
3 specimens 2nd moult 17
3 specimens 3rd moult 24
Eating little and very
fat 3 31
Had eaten nothing for some
time and allowed to go
down on September 9.
Laid August 7-8
1 hatched ,, 12
2 ” ” 13
OMe ame e if 15
7 in 2nd skin 21 rest removed
2 moulted to 8rd August 26
All in 8rd skin A 29
5 in 4th skin September 2
All a Ps 4
Full-fed i 13
260 Dr. T. A. Chapman’s
Hatched September 18 | Eggs laid August 3-4
Full-fed 2nd skin 3 27 Those (11) hatched 9-12 removed
All in 8rd skin October 4 The rest hatched August 13-15
2 in 4th skin Se BiG Several in 2nd skin ,, 17
All in last skin (4th) ,, 12 All . _ omer
Full-fed a eal Some in 3rd 5 ast O26
Going down ie le Some in 4th 29
Nearly all in 4th akin Sept. 2
Hatched September 13 | All ‘ is yon
Most in 2nd skin _,, 21 |» Full-fed sy als
BHHOl "ps 7 ih
Many in 4th skin __,, 30
It may be useful to summarise the items of the life
history of O. tristis noticed above. The beetles emerge from
the ground in May, and eating freely of the food- -plant
(Centaurea sp.) and pairing frequently, lay eggs abundantly
during June, July and August, and even on to November,
some specimens living (in confinement) until the following
March.
The eggs are rarely laid singly, usually in small batches
up to twelve or twenty, or even considerably more. When
laid the eggs have already undergone some development,
sometimes only to the mulberry stage, usually the larva is
already largely developed, and more rarely the eggs may
be ready to hatch. This seems to be more frequent later
in the season. The eggs hatch by two slits forming, under
internal pressure caused by the secretion of air in the
alimentary canal of the young larva, their site being
determined by the presence of “hatching spines.” The
larvee moult three times, the effete skim being similarly
split by pressure from “ inflation.”
They feed up in about a month, rather longer in cool
weather, shorter probably in hot, and then bury themselves
in the earth to a depth of one to five or six inches, remain-
ing unchanged to the following March or April, when the
further changes are rapidly gone through. It seems
possible, but I have no evidence whatever to make it
at all probable, that the earlier larvae feed up and may
emerge the same year,
Postscript, June 11th, 1903.—As an indication of the
date of the appearance of the beetle, on April 27th, 1903,
at the habitat of the beetle at Pino, only two specimens of
the beetle could be found, these happened to be a male
and female; the Centaurea was only some twelve inches
high. On May 30th, 1902, it was three to four feet high,
Contribution to the life history of Orina tristis. 261
but not in flower, and the beetles were plentiful. On
May 30th, 1903, two beetles were found at Reigate in my
garden, from larvee turned out last autumn, and one or
two a day have appeared since. These are all soft and
Immature.
The Orina taken at Pino April 27th, paired on May 7th
and May 8th, but not for long periods. They were placed
again in the same jar May 24th, and remained paired for
forty-eight hours on May 25th, 26th and 27th. The
female again paired with one of the Reigate beetles May
30th and 31st. On June 4th she laid two eggs ready
to hatch, and two more on the 5th. Of these one failed
to hatch, the others hatched on 5th and 6th, and larvee are
now feeding.
These are probably the result of the pairing of May 7th
and 8th, but this is by no means certain. The beetles do
not seem happy or to feed freely, when kept solitarily.
EXPLANATION OF PLATES X AND XI,
PRAtH xe
Fia. 1. Egg ready to hatch, showing “hatching spines” and other
parts of contained larva x 13.
. Egg-shell showing slits made at hatching x 13.
. Larva in Ist stage x 30.
5 2nd
55 Bye B IICO)
Athy; 3 6:
x dorsal view 6.
x 6,
x 6.
>t) ” ”
. Pupa lateral view
» ventral ,,
ODAAAS ww
PLATE xo
Fia. 1. Newly-hatched larva squeezed flat, showing hatching
spines and air bubble in gullet, ete.
. Another specimen, air bubble pressed out, now entangled
with legs.
i)
XVI. The Butterflies of Chile. By Henry JOHN
ELwEs, F.R.S., F.L.S., ete.
[Read June 4th, 1902.]
Puates XII, XIII, XIV, anp XV.
Our present knowledge of the butterflies of Chile consists
almost entirely of bare descriptions published by authors
who had little knowledge of the country or of the climatic
conditions which have tended to make the fauna of this
country so interesting and peculiar.
Guérin in the ‘ Voyage de la Coquille,’ published in 1832,
was the first entomologist who seems to have received any
butterflies from Chile, except a few of the very common-
est, which Molina, Drury, and Hiibner had already
described.
Blanchard in Gay’s ‘Fauna Chilena, published in 1852,
has described and figured more of the commoner ones.
Philippi has described others in ‘ Linnzea Entomologica’
in 1860, but though an excellent botanist he has paid
little attention to the Lepidoptera of his adopted country.
feed, an English naturalist long resident in Chile,
published in 1877 a small work in Spanish, ‘ Mariposas
Chilenas,’ with indifferent figures and descriptions of
several of the rarer ones, but gives little information as to
- their habits and distribution.
Wallengren in the ‘Wiener Ent. Monatschrift,’ and
‘Eugenie’s Resa,’ and Felder in the ‘Reise der Novara,’ add
a few more to the list, but the first serious attempt at a
Catalogue of the Lepidoptera of Chile was that by Butler
in this Society’s Transactions for 1881.
This paper was based on a collection made in Chile by Mr.
Thomas Edmonds, and describes a good many new or sup-
posed new species, giving a full synonymy of those already
described, with some useful but rather fragmentary notes
by Mr. Edmonds on the localities in which they were
found.
A translation of this paper into Spanish, with a list of
the Microlepidoptera of Chile described by Ragonot and
Zeller, was published by Mr. Bartlett Calvert, a member
TRANS. ENT. SOC. LOND. 1903.—PART Il. (OCT.)
264 Mr. H. J. Elwes on
of our Society, in the ‘ Anales de la Universidad de Chile’
in 1895.
None of these writers, however, have told us much of
the nature of the country which produces these insects, or
described the very peculiar geographical and physical con-
ditions which have influenced their variation and distribu-
tion, and no one has realized the great amount of variation
which is found in many of the species. Therefore in
attempting to give a better idea of the Butterflies of Chile,
I have had to rely mainly on my own observations.
I left England in November 1901, and arrived at
Buenos Ayres on December 2nd. Here I endeavoured
to find a companion who knew the language, and who
had travelled in the Southern Andes, which until quite
recently have been unvisited and undescribed by natural-
ists. I arranged with Mr. Arneberg, a Swedish engineer,
naturalized in Argentina, to accompany and assist me, but
when I reached “Santiago de Chile, I found that the
strained political relations between Chile and Argentina,
which very nearly led to war, and which delayed my start
for over a mouth, made it undesirable to carry out this
plan, and I eventually secured the companionship and
assistance of Mr. Bartlett Calvert of Quillota, who speaks
Spanish like a native, and who is a well-known ento-
mologist and an accomplished photographer.
I arrived at Puente del Inca, to which point the Trans-
andine railway is now open, on December 10th, and
stayed twonights in the excellent hotel which belongs toDr.
Cotton, and which stands close to the celebrated Natural
Bridge and warm baths, at an elevation of over 9000
feet. Here I was able to gain some idea of the flora and
butterflies of the high Andes, though the weather was
extremely dry, and the high wind from the west, which is
a daily feature of this locality after nine a.m., makes
collecting difficult.
I found at the entrance to the valley called Horcones,
and round the Laguna de Horcones, a few very interesting
species, including Scolitantides andina, a Colias which I
thought at the time was near cuxanthe, Feld., Argynnis
lathonioides, and Phulia nymphula, all of which are
peculiar to the high Andes of Chile and Bolivia.
I was unfortunate enough to lose the bag which con-
tained most of these specimens when star ting in the dark
5
the next morning to cross the pass into Chile, but I saw
The Butterflies of Chile. 265
enough to convince me that the envirous of Puente del
Inca are well worth a week’s stay in January by a passing
entomologist, and that a few of the species described by
Staudinger, in Iris Vol. vii, from the Bolivian Andes would
probably be found here. The pass between Puente del
Inca and Chile is over 12,000 feet, but very easy to cross
on mule-back except for three or four months in winter, and
the vegetation of the Chilean side from 10,000 down to
about 6000 feet is much richer than that of the eastern
valleys. Juncal, where there is a comfortable inn kept by
a Frenchman, would also be a desirable halting-place for
any one who wished to collect on the western side of the
pass. I was obliged, however, to hurry on, and could do
little in the way of collecting on this occasion. When I
returned at the end of February the season was far
advanced, and I was again unable to stop as I had to
catch the steamer at Buenos Ayres. From Juncal a good
carriage road leads down to Salto de Soldado, whence there
is a railway connecting at Los Andes with the Chilean state
railway to Valparaiso and Santiago, making it possible to
go in one day as I did from Puente del Inca to Santiago.
In this beautiful city I met with the greatest hospitality
and civility from many men of science and distinction,
among whom I should especially mention Prof. Federigo
Philippi, the director of the Museum, a most distin-
euished botanist, whose father, the describer of so
many Chilean plants, still enjoys life at the age of ninety :
Dr. Barros the Rector of the University, who was kind
enough to give me an introduction to a family from whom I
afterwards received the greatest possible kindness and help ;
and Sefior Dr. Vicente Isquierdo, who has the most complete
collection of Lepidoptera in Chile, and who has been good
enough to send some of them to the British Museum for
identification. The brother of the latter, Sefor Salvador
Isquierdo of Santa Ines, has a large and most interesting
collection of trees, and is developing the rising fruit-growing
industry in Chile in a manner worthy of the best EKuropean
horticulturists. Sefiora Ana de Jordan, the hostess of Miss
North, was kindness itself, and, ike many of the members
of the old Chilean families, gave me a charming idea of the
state to which progress and civilization have reached in her
delightful and to most Europeans little-known country.
Whilst waiting until the question of peace or war should
be settled I visited the Baiios de Cauquenes, a well-known
266 Mr. H. J. Elwes on
watering-place in a valley three hours south of Santiago
by rail, which has been described by Miss North, by Mr.
Ball, and by Darwin. Here I stayed four days and
collected all the species which were then out, but
failed in an attempt to reach the highest part of the
valley, where there is said to be good collecting ground
at 5000 to 6000 feet. This valley, however, is well
known in comparison with the mountains I afterwards
visited farther south, and has been visited by Reed and
others. The country around was dried up, hot and windy,
and the variety of insects not great, though I got one
species which seems to be undescribed.
I then went by rail to Concepcion on the coast, three
hundred miles south of Santiago, and visited the beautiful
park and gardens of the late Sefora Cousinho at Lota,
where Mr. O'Reilly, the superintendent of the gardens
and plantations, was good enough to show me everything
he could. I found few butterflies, however, in this neigh-
bourhood though there is some very good-looking collect-
ing ground near Coronel, and some trees and shrubs
characteristic of Southern Chile, which do not occur much
farther north, are found on this part of the coast.
On December 21st I left Concepcion by train, and got
to Chillan by 11 o'clock. From here there is a carriage
road to the celebrated Baths of Chillan situated in a
wooded valley of the Andes about sixty miles distant.
Starting in the afternoon ina carriage, I reached the second
stage of the j journey at dark, and slept in a fair roadside
inn. From here the road to the Baths passes through a
country which was once covered with forest, and is still in
many parts most beautifully wooded, the last stage from
Las Trancas to the Baths being specially fine, through
splendid forests of beech, of which three or four different
species constitute the principal forest trees in this latitude.
At the Baths I found excellent accommodation and a
rich flora. This was Mr. Edmonds’ best collecting ground,
and during January and February most of the peculiar
Chilean forest species may be found in the woods below
the Baths, and many of the Alpine species on the bare
mountains above them. I spent four days here, and got
the greater part of the species found by Edmonds, though
I was too early for two or three of the rarest. The snow
still lay at an elevation of 6000 to 7000 feet in shady places
near the Baths, and the weather, which had been wet
The Butterflies of Chile. 267
previously, was fine and hot, though, as usual in the
Andes at this season, very windy in the afternoon.
On December 28ta I returned to Santiago to make
arrangements for my journey to the south, and as our
Minister, Sir B. Cusack Smith, advised me not to take an
Argentine subject as my companion, I went to Quillota,
where I arranged with Mr. Calvert to meet me as soon as
he could get away on January 9th. Quillota has been so
well described by Darwin* that I need say nothing of it.
There is no good collecting ground within four or five
hours’ ride, but I got a few specimens here and at
Llai-llai, the junction between the lines to Santiago and
Valparaiso. Mr. Paulsen, who lives at Quillota, and
Mr. Calvert have both collected here, especially Coleoptera,
and the latter showed mea very nice collection mainly
of Coleoptera.
Leaving Santiago on January 3rd, I slept at San
Rosendo, where the line to what is called “la Frontera ”
diverges from the line to Concepcion and Lota. Here I
had only a morning in which to sample the environs, and
next day reached Victoria, a large new town to which most
of the live stock raised in the frontier districts of Argen-
tina comes over the Lonquimay Pass to market. Most of
the country round here has been cleared of forest near the
railway, but at Temuco further south, and at Tolten which
was then the terminus of the line now being extended
to Valdivia, I found the virgin forest which covers the
greater part of Chile south of the Biobio river, and soon
became convinced that, though the moths might afford a
rich harvest to a resident collector, the south of Chile, like
the centre, is, away from the mountains, very poor in
diurnal Lepidoptera. Returning to Victoria I engaged a
German anda Chilean as servants for my Andean journey,
and went on by a branch line to Mulchen, whence I drove
twenty miles east through a country recently cleared of
forest to the hacienda of San Ignacio, the property of
Sefiora Bussey, who received me with the greatest hospi-
tality,and to whose husband, George Bussey, Esq., I am in-
debted forinvaluable assistance in engaging reliable men and
mules for my journey. Whilst these were. being brought
in I collected in the neighbourhood, but found that though
the valley of the Renaico river, near which the hacienda is
situated, is mostly virgin forest with the rich evergreen
* Darwin’s ‘Journal,’ New Edition, Murray, 1890, pp. 269-286.
268 Mr. H. J. Elwes on
vegetation peculiar to Southern Chile, very few butterflies,
and those the same as I had previously taken, were to be
found. Moths, however, are abundant at light, and a
good many were collected. San Ignacio lies some little
way from the foot of the Pemehue Mountains, which
constitute an outlying group on the west of the Biobio
Valley, and the country round the hacienda is now mainly
under wheat cultivation.
I was here delayed more than a week by a sudden and
severe attack of illness, from which I was fortunate enough
to be cured by the medical skill of Senora Bussey’s brother,
Dr. Puelma, and was nursed with as much care and atten-
tion as if I had been at home. I cannot speak too grate-
fully of the kindness and hospitality of this charming
family, who, like many of the upper-class Chileans, are,
though living in a country only recently conquered from
the Indians, as civilized and well educated as any people
in Europe.
I was at last able to make a start on January 22nd, a
month later than I had hoped to do, and rode in two days
up the valley of the Renaico river through a beautiful
country mostly covered with virgin forest, and then over
the Sierra de Pemehue to Lolco, a hacienda belonging to
Senor Manuel Puelma, another brother of Senora Bussey’s.
This is a beautiful place near the Biobio river, and near it
I got some of the best insects I found on my journey. No
one except Mr. Calvert had ever collected Lepidoptera on
this road before, and if I had not been so pressed for time
I should have stayed longer.
On January 27th we started early from Lolco on a very
cold morning, and rode first through grassy valleys which
reminded one of Mongolia, and splendid araucaria and
beech forests, over a very striking pass about 8000 feet
high, to Lonquimay, which is the Chilean outpost and
custom-house, on the main road from Victoria to Argentina,
and from here in one and a half days up the head-waters
of the Biobio we reached the Argentine frontier, which is
an open bare ridge about 5000 feet high, and not the
least like the northern passes over the Andes, the higher
mountains in this latitude being isolated volcanoes; which
lie well to the westward of the watershed. Close to the
pass is an outpost of Argentine cavalry at Los Arcos, where
I was civilly received by the lieutenant in command, and
from here we turned more to the southward, and reached
The Butterflies of Chile. 269
the beautiful Lake of Aluminé, one of the long line of lakes
which are found mostly on the east side of the watershed, and
which form the head-waters of many rivers, some of which
run to the Pacific and some to the Rio Negro, and which
are in the frontier region, of which parts were in dispute.
This country has been described by Senor Moreno.* From
here my route southwards for three weeks lay through
Argentine territory, but I soon realized that there is no
well-marked natural boundary between Chile and Argen-
tina in this latitude, and that the influence of the rainfall,
which rapidly becomes less when the watershed is crossed,
is the real factor in determining the boundary between the
wet sub-evergreen forest which clothes the western slopes
of the mountains, and the dry grassy hills and valleys on
the east, which gradually fall away into the great arid
pampas of Patagonia that now form part of the ‘Argentine
territory of Neuquen.
It also became evident that nearly all the mountain
and forest species of butterflies which hitherto have been
considered peculiar to Chile occur in Argentine territory as
well, and that there are few species peculiar to the Pampas
in the country which [ passed through.
On reaching Lake Quillen im lat. 41° S. I made an
attempt to re-cross into Chile by a track which formerly
existed through the forest to Villarica, but it had become so
much blocked by fallen trees that we could not get through,
and after two days in the forest I turned south from Lake
Quillen, crossing a pass over a lateral ridge about 6000 feet
high to Junin de Los Andes, where an attempt has been
made to found a frontier town in a very sparsely inhabited
but fine cattle-ranching country. Up till now the weather
had been splendid, too hot for comfort in the middle of the
day, when high wind from the south usually prevailed, but
now we had a spell of bad weather which very much
hindered my collecting. At San Martin on Lake Lacar
we found a military settlement commanded by Col. Perez,
from whom, as well as from the other officers of the 3rd
Argentine Cavalry, we had a most hospitable reception.
This would be a good collecting station, as, though the
elevation is under 3000 feet, there is a heavy snowfall in
winter and a rich forest vegetation on the shores of the
lake, whilst a range of mountains about 7500 feet high,
* Notes préliminaires sur une excursion aux territoires du Neuquen
Rio Negro, etc., de Francisco P. Moreno. Musée de la Plata, 1898.
270 Mr. H. J. Elwes on
and wooded up to about 6000 feet, which is the approxi-
mate timber-line in this latitude, is within a ride of the
town. Here I got a good Indian guide, and left on
February 8th, intending to reach the great Lake Nahuel-
huapi, which is the largest of all the lakes on this part of
the frontier.
Our route through the mountains was extremely inter-
esting and picturesque on account of the extraordinary
volcanic rock formations which occur in these valleys, but
the heavy rain which came on soon after leaving San Martin
swelled the numerous mountain streams to an extent which
made some of them difficult and dangerous to ford with
loaded mules.
From San Martin to Nahuelhuapi I was able to do but
little insect collecting, having long marches and_ broken
weather, but the few “butterflies which I saw showed that
there was not much change in the fauna, and the country
began to assume an autumnal appearance. At the point
where the great river Limay runs out of Lake Nahuel-
huapi, I found a ranch and store kept by a Scotchman
named Neil, who is in partnership with Mr. Jones, one of
the earliest and principal ranch owners of the district ;
here I sold my mules and horses, and had intended to
take a boat and follow the river Limay down to its
confluence with the Rio Negro, about five days’ journey by
river and then by rail to Buenos Ayres. But I heard such
good accounts of the pass into Chile which is called the
pass of Perez Rosales that I gave up the idea of returning
by this route, and went on to a settlement called San Carlos,
founded by the German firm of Hube and Achelis, which
is on the south shore of Lake Nahuelhuapi. From here
there is a regular route to Puerto Montt by small steamers
on the lakes of Nahuelhuapi, Todos Santos, and Llanquihue,
which are connected by good mule-roads, and where lodgings
can be got in new houses built and managed by this enter-
prising and obliging firm. Anything more beautiful,
interesting, or novel than this route, which can be traversed
in about three days, cannot be imagined, and it must some
day be the favourite tourist resort of South America.
Though very profitable from a botanical pomt of view, I
was too much pressed for time to be able to add much to
my entomological collections on this part of the trip, and
after being delayed a day on Lake Llanquibue, only arrived
at Puerto Montt just in time to catch the steamer to Con-
The Butterflies of Chile. 201
cepcion, whence I had to hurry back to Santiago, and after
packing and drying my botanical collection, part of which
had been left at the museum in charge of Dr. Philippi,
I re-crossed the Andes by the Mendoza Pass, and reached
Buenos Ayres in time tocatch the mail steamer on March Ist.
Before giving a list of the butterflies I collected, which
include almost all the species hitherto described from Chile,
I must say a few words as to the climate of different parts
of the country. From a naturalist’s point of view Chile
may be divided into three regions.
First, Northern Chile, which consists of the long strip
of country comprising the provinces of Tacna, Tarapaca,
Autofagasta, Atacama and Coquimbo, from 18° to 32°S.,
between the sea and the Andes,about which I know nothing
personally. It includes the nitrate districts which are
absolutely arid, and most of the best mining districts ;
and from a zoologist’s pomt of view is poor, on account of
the very slight rainfall and absence‘of vegetation, except in
the valleys of the few perennial streams and tracts which
are cultivated solely by means of irrigation. It includes a
great tract of high mountains 1m the district of Autofagasta
and Tarapaca, which are, as far as I can learn, almost if
not entirely destitute of trees, and which have therefore
afforded no route by which tropical forms could extend
from the north, as they might have been expected to do if
the country had not been too dry.
Secondly, the coast region and valleys of Central Chile
from about lat. 32° down to about the latitude of Con-
cepcion and the river Biobio. This part of Chile has a
climate much like that of Southern Spain, damper on the
coast and drier in the great central valley between the
coast range and the Andes, and is the most populous and
richest part of Chile from an agricultural point of view,
being fertilized by the deposit from numerous mountain
streams which are made to irrigate large tracts of highly
fertile country. Forest is now found in Central Chile only
at and above elevations of 3000 to 6000 feet, and in the
upper parts of the Andean valleys; and I doubt whether
there was ever much true forest in the neighbourhood of
Santiago, where the hills are covered with shrubs and
thorny bushes, and where rain usually falls only during
twenty to thirty days in winter.
In the latitude of Chillan heavy forest is found, or rather
was found until it was destroyed by fire, along the foot and on
TRANS. ENT. SOC. LOND. 1903.—PART Ill. (ocT.) 19
paler Mr. H. J. Elwes on
the outer valleys of the Andes, but on the coast there is much
brushwood, and in narrow gorges and valleys some timber,
though the plains are still very dry. South of the Biobio
the climate changes very suddenly, and when Valdivia is
reached the rainfall is so great that the country is covered
with a forest so dense that one can hardly penetrate it;
where ferns grow twelve feet high, and a bamboo-like
grass climbs the trees to a height of forty feet.* Inland,
however, there is some open savannah and marshy
country at the foot of the volcanoes, south of Lake
Lianquihue.
This region has a tew peculiar Satyridze and Hesperidee,
but is extremely poor in diurnal Lepidoptera, though very
little collecting has been done except about Valdivia and
La Union.
Lastly, I take the region south of lat. 42° down to the
Straits of Magellan, which is familiar to us from the
writings of Dar win, Cunningham, and other naturalists, and
which includes a ereat number of forest-clad islands and
unexplored mountains, of which our zoological knowledge
at present is very limited. Some parts of the coast are
fairly well known to naturalists, and the district lying
south of Lake Nahuelhuapi and east of the watershed,
which has been described by Moreno and more recently
by Prichard,+ is beginning to attract colonists, especially at
the head-waters of the Chubut river, where there is some
good ranching country now occupied by Welsh settlers
from the east coast. This region will probably be found
to contain many of the species which I found farther north,
but the west coast and Straits of Magellan have such a
very wet and inclement climate that there can be but few
butterflies, and those few nearly allied to or identical with
those of Chile and Argentina.
The list of butterflies which follows is based on what
is by far the most complete collection of Chilean butterflies
ever brought together in Europe, comprising my own
collection, that of Mr. Edmonds, which was contained in
the British Museum, and Mr. Godman’s collection, and the
specimens taken by our fellow, Mr. J. J.Walker, R.N., when
serving on the coast at Coquimbo, Valparaiso and Concep-
f=)
cion. I have also been able to examine the types of
* See Darwin’s ‘Journal,’ New Edition, Murray, 1890, pp. 318-
322.
f+ Prichard, ‘Through the Heart of Patagonia,’ London, 1902.
The Butterflies of Chile. 273
several species described by Mabille, Felder, and Reed,
and have added some specimens from the late Mr. Crowley’s
collection now in the British Museum, where the whole
of this material is now placed.
There are, no doubt, in the collections of Mr. Paulsen
and Dr. Isquierdo many specimens which would have
been invaluable in studying the variation of Chilean
butterflies, and perhaps some new species, but as they have
had no means of identifying their specimens with the
types, which are all in Europe, and I was too ignorant of
the Chilean species to make notes of them when I saw
them, this list must not be taken as more than an
attempt to improve the foundation of what is still very
fragmentary and incomplete knowledge.
What is wanted are resident collectors, especially in the
south, who will observe the seasonal and local variations,
and bring together much larger series than now exist;
by which means alone my conclusions as to specific values
can be proved or disproved.
It will be seen from this list :—
First, that the number of species is extremely small,
being, if the diverse conditions of climate are considered,
the smallest found in any country in the world of equal
length from north to south.
Secondly, that the number of butterflies which are not
endemic in Chile, if the Argentine as well as the Chilean
slopes of the Andes are included, is very small.
Thirdly, that the scarcity of Nymphalidze and Lyccenide
is extreme; only six species of the former and seven of
the latter occurring in this immense tract of country, a
smaller number than could be taken in a single day in
almost any part of North America, Europe, or the
temperate region of Asia. Not a single Lyccenid and
only one or two Nymphalid butterflies seem to have been
taken south of Valdivia; a case quite unparalleled in
other parts of the world.
The Chilean butterflies consist almost entirely of
Satyridz and Hesperiidee, many of which are confined to
the forest region of the centre and south, and there is
a marked absence of alpine species ; only
2 Pierids
Nymphalids
Lyccenid, and
or 3 Satyrids
re po
Ww
274 Mr. H. J. Elwes on
can be considered as strictly alpine butterflies, though
several of the common species of the valleys and forest are
found up to, and even above, timber-line. Many of the
forest insects seem to be found only where Chusquea, a
bamboo-like grass, is abundant, but some others, such as
Argyrophorus argenteus, Cosmosatyrus leptonewroides, and
Neomenas simplex, are found on grassy downs and hills, and
never enter the forest so far as I observed. .
In the arrangement of the Hesperiideze I have availed
myself of Watson’s ‘Classification’ of that group (P.Z.8.
1893, pp. 8-132).
I have not studied the generic characters, as it seemed
to me that this could not profitably be done without
undertaking a study of the allied species from other parts
of 8. America.
SATY RIDA.
1. Hina lefebvrer.
Satyrus lefebvrei, Guérin, Voy. de la Coquille, p. 281
(1829).
S. montrolvi, Feisthamel, Mag. Zool., Ms., Pl. 20 (1839).
Laswmmata montrolu, Westwood, Gen. Diurn. Lep.,
p. 387 (1851).
Q Elina montroli, Blanchard, in Gay’s Fauna
Chilena, vii, p. 29, Pl. V, fig. 7 (1852).
A well-known and very distinct species, the largest of
the Chilean Satyrida. I found it common at the Baiios
de Cauquenes, where it flies in shady woods and settles
on the trunks of trees. It was also common at Temuco
and near San Ignacio, but I did not see it in the mountains.
The southern form is darker than that found at Valparaiso
and Quillota.
2. Hlina vanessordes.
Elina vanessoides, Blanchard, t. ¢., p. 28, Pl. V, figs. 5, 6.
Also a very distinct species which I did not take myself,
but which is common in the neighbourhood of Valdivia.
3. Hlina neomyrioides. (Plate XIV, fig. 5 3.)
2 Satyrus neomyriordes, Blanchard, t.c., p. 33, Pl. IT,
figs. 6, 7.
I doubt whether this species belongs to the same genus
as the last. I took it at San Ignacio in January, and at
The Butterflres of Chile. 275
Port Blest on Lake Nahuelhuapi in February. I have
figured a male of this species as it appears to be
undescribed.
4. Elina calvertiu, n. sp. (Plate XIV, figs. 3 ¢, 4 2.)
This species was not uncommon in the forest below the
Bafios de Chillan in December, when I took five males
and one female in fair condition. I also found a single
female in the thick forest on Lake Quillen in Argentina
on February 3rd; and there is in the British Museum a
female taken by Edmonds below the Baths of Chillan,
which Butler has mistaken for the female of neomyrioides
and marked as such. There is a distinct patch of velvety
androconia on the fore-wing in this species and the last
which is not found in the species of Elina, and they are
probably not congeneric. The base of the fore-wing below
in L. calvertii is, like that of #. edmondsit, fulvous, which
distinguishes it at once from neomyrioides, and the lower
part of the band on the hind-wing below is also much less
defined and does not form a conspicuous white patch on
the costa. On the upper-side the band is also much less
distinet forward.
5. Pedaltodes flora.
2 Satyrus flora, Philippi, Linnea Entomologica, xiv,
p-. 267 (1860).
Hipparchia ? flora, Butler, Cat. Sat., p. 58 (1868).
Pedaiiodes oaxes, d., Cist. Ent., i, p. 25 (1870).
Stibomorpha tristis, id. (nec Guerin), Lep. Exot.,
p. 180, Pl. LXII, fig. 3 (1874).
Satyrus tristis, Reed, Mon. Marip. Chil. Pl. ILI,
fig. 4 (1877).
Stibomorpha veedi, Reed (nec Butler), ¢t.c. explic. de
las laminas, lam. iu, fig. 4 (1877).
Neither im its appearance, habits, nor flight has this
species any affinity with lina lefebvrer or with any other
species of Elina that I observed. I found it abundant in
marshes overgrown with reeds, bushes, and great tufts of
Gunnera scabra on the east shore of Lake Llanquihue, where
it was fresh in February. It has a slow flight amongst
the rushes and bushes, and is very easy to take. The
females differ but little from the males. Edmonds found
it common in marshes at Valdivia.
276 Mr. H. J. Elwes
6. Hpinephele edmondsii.
EF. edmondsit, Butler, Trans. Ent. Soc. Lond., 1881,
p. 451, Pl. X XI, fig. 2.
This seems to be a distinct species, which was described
from one specimen only, taken near the Baths of Chillan
in March by Edmonds. This specimen is in the British
Museum, and agrees very closely with two in that collec-
tion from Cordoba and two from Uruguay sent by Berg to
Zeller.
7. Epinephele anirioides.
E. janirioides, Blanchard, t.c¢., p. 34, Pl. I, fig. 8 2.
Satyrus limonias, Philippi, ¢.¢., p. 268 ¢; Reed, ¢.¢.,
Pie 7.
Var. Epinephele dryas, Felder, Reise der Nov. Lep.,
ill, p. 492 (1867).
This was common at the Banos de Cauquenes on dry
hill-sides covered with bushes from the 18th to 21st of
December, and at San Ignacio in January. It is common
at Valparaiso in November and December, and has been
taken at Taleahuano and at Valdivia. The specimens in
the British Museum from this locality are smaller than
mine but not otherwise different. As I think Blanchard’s
figure of this species is unmistakable I adopt his name.
8. Epinephele monachus.
Satyrus monachus, Blanchard, ¢. ¢., p. 35.
Epinephele valdiviv, Felder, t.¢., p. 493.
Pedaliodes lugubris, Butler, Cist. Ent., 1, p. 25 (1870).
Stibomorpha monachus, id. Lep. Exot. p. 179,
Pl, XII, fig. 2 (1874): Reed}z.<., Pi Ul, figs5;
and explic. de las laminas, lam. 11, figs. 5, 6.
Satyrus luctwosus, Reed, t.c., Pl. I, fig. 6 2.
This species was very abundant in the heavy virgin
forest south of Temuco, where it was fresh out on January
10th. It was also very abundant amongst the dense
bamboo undergrowth in the Upper Renaico Valley at
3500 feet at the end of January, and was common in
similar forest on Lake Quillen and at Port Blest in the
middle of February, when it seemed to be nearly over.
I cannot distinguish the insect which Reed figures badly
under the name of /uctwosus. I have compared the type
of valdiviv, Feld., with my specimens from Temuco, and
believe it to be identical.
The Butterflies of Chile.
i)
“I
~I
9. Hpinephele tristis.
? Satyrus tristis, Guérin, Voy. Coq., p. 281.
? Argynnis tristis, rd., t.c., Atlas. Ins. Pl. XVI, fig. 5
(1832).
? Epinephele cocter, Reed, t. ¢., expl. de las laminas, lam.
ay stTe 3)
2S. pales, Philippi, t. ¢, p. 268.
Guérin’s figure being, like his description, very bad,
there is much confusion as to the synonymy of this species,
which being a common insect in all the low country varies
a good deal in size and in the markings on both sides.
I recognize two allied species only, of which this is the
larger. It was common amongst bushes at Bafios de
Cauquenes on December 15th and up to about 3000
feet; also amongst bushes at San Rosendo on the lower
Biobio on January 5th. I found it nearly over at
Llai-llai on January Ist, and took a single female at over
4000 feet in the Chillan Valley on December 22nd.
10. Epinephele coctet.
H. coctei, Guérin, t.c., id., Mag. de Zool. Ins., Pl. XI
(1839).
Erebia cocter, Westwood, t. ¢., p. 380.
¢? Hpinephele coctet, Butler, Cat. Sat., p. 68 (1868) ;
2 Satyrus tragiscus, Reed, t.c., Pl. IL], fig. 3.
Guérin’s figure of this species is I think unmistakable.
It is a smaller species than the last, and probably distinct,
though it may be only a mountain form of it. I did not
find it anywhere in the low country, but first saw it
abundantly flying amone the scrubby bushes and grass
tufts in the Upper Biobio Valley above Lonquimay on
January 28th. From here as far south as the Rio
Limay it was common on suitable ground from 2000
up to about 4000 feet. The males are always, when
fresh, darker than the last, and the under-side is less
variegated, but I can find no distinction that I could
consider specific between the two. The only specimen
I find in the British Museum marked cocter is a female
from Talcahuano sent by Edmonds; Butler says he thinks
this is only a dwarfed form of ¢ristis.
11. Neomenas cenonymphina.
N. cenonymphina, Butler, Trans. Ent. Soc, Lond.,
1881, p. 454, Pl. X XI, fig. 4.
278 Mr. H. J. Elwes on
This is a species somewhat like the last above, but
perfectly distinct by the broad pale band on the hind-wing
below, and on both wings in the female above, which
Butler’s figure does not well show. According to Edmonds
it is local at Valparaiso. I did not take it myself.
12. Neomenas fractrfascia.
N. fractifascia, Butler, t.c., p. 455, Pl. XXI, fig. 3.
A very distinct species which I found scarce at San
Martin, near Lake Quillen, and in the forest at Port Blest
in February. It frequents open places in the forest, and
was also taken by Edmonds near the Baths of Chillan in
March. The female has much more red on both wings
above, but both sexes vary in this respect.
13. Neomenas servilia.
N. servilia, Wallengren, Kon, Vet. Akad. Forh., p. 78
(1858); id., Wien, Ent. Mon., iv, p. 36 (1860) ;
wd., Kug. Resa, p. 354, Pl. VI, fig. 1 ¢ (1861).
Stibomorpha decorata, Butler, Ent. Mo. Mag., x,
p. 205 (1874); zd., Lep. Exot., p. 179, Pl. LXII,
fig. 3 2 (1874).
I took one only at Cauquenes. It is not uncommon
at Valparaiso and elsewhere, and frequents bushy un-
cultivated ground, but is not, I think, found in the forest
region.
14. Neomenas wallengreni.
N. wallengrenii, Butler, Trans. Ent. Soc. Lond., 1881,
p. 456, Pl. XXTI, fig. 5.
A rare species, which seems distinct from the last, and
is represented in the British Museum by two bad speci-
mens taken by Edmonds in the woods below the Baths
of Chillan in March. I did not find it myself.
15. Neomenas ? inornata, n. sp. (Plate XIV, figs. 1 7,
2 ¢.)
This species seems to be most nearly allied toservilia, Wall.,
with which it agrees in size and shape, but it is darker in
colour above and has the under-side of the fore-wimg in
both sexes of a deeper rufous colour. The hind-wing below
is perfectly plain olive-grey with a darker marginal line.
The female has on the fore-wing above an indistinct black
ocellus, and both sexes have on the fore-wing below an
The Butterflies of Chile. 279
ocellus like that of servilia. As in that species there
is no sexual patch on the fore-wing of the male. It is
also somewhat like Argyrophenga simplex, Butler, but
larger and much more rufous above, and has the hind-
wings of a different shape. I can find no described species
to which it can be referred in any collection. 1 took two
males and one female at the Banos de Cauquenes on
December 15th, flying on a bushy hillside above the
Baths and settling on low trees.
16. Neomenas ? edmondsit.
Argyrophenga edmondsii, Butl., t.c., p. 457, Pl. XXI,
fig. 6.
I know this only from the very bad specimen in the
British Museum which Edmonds took in March 1880 in
woods below the Baths of Chillan.
17. Neomenas ? humilis.
Stygrus humilis, Felder, t.c., p. 489.
Butler identifies as above a distinct species with the
same markings as ambioriz, but the ocelli much smaller
and fainter. It can be distinguished, however, I think
from ambiorix, certainly by the absence of any chocolate
colour in the fore-wing below, and by the rounder and less
pointed fore-wings. Edmonds found it common in woods
near Valdivia. I only took two or three males in forest
at Quillen and near Lake Aluminé at about 3000 feet;
these agree with Edmonds’ specimens, which I have
compared with Felder’s type and find identical.
18. Cosmosatyrus leptoneuroides. (Plate XV, figs. 3 2, 4 2,
6 2.)
Cosmosatyrus leptoneuroides, Felder, t.c. p. 495, &.
Satyrus antarctica, Reed, t.¢., Pl. I, fig. 4.
Tetraphlebia germaini, id., t.c., explic. de las laminas,
lam. 11, fig. 4.
2=T. plumbeola, Butler, Cat. Sat., p. 95, Pl. I, fig. 11
(1868).
I first took this species in the Renaico Valley at
Maitenes, a farm of Mr. Bussey’s, about 2500 fect
elevation, when it was fresh out on January 24th, and
common in grassy openings in the woods. Afterwards it
280 Mr. H. J. Elwes on
became abundant everywhere, and was generally dis-
tributed at elevations of 3000 to 5000 feet in all suitable
situations. I have figured three specimens to show the
great variation which exists in the species. The type of
leptoneuroides is a very large male, and agrees well with
my largest. Though at first I thought that the androconia
on the male fore-wing of some specimens would distinguish
them, yet on careful comparison of my very large ‘series I
am unable to separate what Butler calls plwmbeola, which
he thinks the mountain form of Jleptonewroides. This,
which is often smaller, was found by Edmonds at 6000 feet
in January, but was not out when I left Chillan at the end
of December. The species does not seem to occur in the
low country or on the coast of Chile, as Reed says he has
never seen it. It is probable that what he figures as
S. antarctica is the same as p/winbeola Butler, described
from the Straits of Magellan, which however he does not
allude to in his paper on Edmonds’ collection. This is also
found at Port Famine, and was placed with plumbeola in
the British Museum by Butler. Staudinger also figures as
Erebia plumbeola, var. Duseni, a variety of the same form
from the Rio Aysen in 8. Chile, showing that the species
has a continuous distribution from about the latitude of 38°
to the far south of Chile. From the figure of Duseni I
do not see much to distinguish it, and it is to be hoped
that the practice of giving varietal names to specimens of
whose distribution and variation so little is known will not
be adopted as largely by authors as it has been in the
Holarctic Butterfly Fauna.
19. Cosmosatyrus chiliensis. (Plate XV, figs. 9 J, 10 2.)
Satyrus chiliensis, Guérin, t.c., p. 280; Atlas, Ins.,
Pl. XVI, figs. 4, 5 (1832).
Stibomorpha veedii, Butler, Lep. Exot., p. 180 (1874).
This is one of the common species of Chile which I
first took in the Chillan Valley at about 3000 feet in
December, when it was just coming out. Afterwards I
found it on the coast near Coronel rather worn, and later
it was common in the Renaico Valley at 2000 feet, and
was found almost everywhere up to about 3000 to 4000 feet
and as far south as Nahuelhuapi, where, however, the
specimens show differences which might enable those
from Argentina to be separated from those taken in
Chile. In order to show these differences clearly, and
The Butterflies of Chile. 281
also how chiliensis can be separated from the next
species, I have figured a male and female from the
east side of Lake Nahuelhuapi, the farthest point south
which I visited. These were flying on grassy pampas
where the rainfall is very much less than in Chile and
the snowfall and cold of winter much greater, and theit
markings show a good deal more resemblance to those of
monticolens. It is possible that this form may prove
distinct from chiliensis.
20. Cosmosatyrus monticolens. (Plate XV, figs. 7 ¢, 8 §.)
Satyrus monticolens Butler, Trans. Ent. Soc. Lond.,
1881, p. 484, Pl. XXI, fig. 1.
I first found this at the head of the Lolco Valley on the
road to Lonquimay, at about 5000 feet, and was at once
certain from its flight that it was a species new to me,
Instead of a slow short flight among bushes like that of
chilvensis, it has a rapid straight flight of 20 to 50 yards
backwards and forwards over wet subalpine meadows always
amongst grass and stones, and was in consequence much
harder to catch. It was afterwards found between the
Aluminé Lake and Pulmari on similar ground at about
4500 feet, and also near San Martin on a mountain side at
6000 feet. It varies considerably, and I have figured a
pair from Pulmari to show the differences between it and
chiliensis. These agree fairly well with Butler’s type,
which came from the mountains above the Baths of
Chillan, where it had not yet appeared at the end of
December.
Cosmosatyrus williamsianus.
Arge williamsianus, Butler, Cat. Sat., p. 159, Pl. IV,
fig. 1 (1868).
(Hneis antarcticus, Mabille, Nouv. Arch. Mus. (3), 1,
p. 142, Pl. X, figs. 5, 6.
The type of williamsianus in the British Museum is a
female in bad condition from Port Famine, Patagonia, and
has been compared with the type of Geis antarcticus,
which appears to be identical. I believe that it is very
nearly allied to, if not identical with, monticolens ; the
shape of the wings and the white veins on the hind-wing
below being similar, and the difference not more than one
might expect in the species when starved and dwarfed by
an ungenial climate.
282 Mr. H. J. Elwes on
21. Argyrophorus argenteus.
A, argenteus, Blanchard, Fauna Chil. vu, p. 30.
Chionobas argenteus, vd., t.c., Pl. II, figs. 9-11.
This is one of the most beautiful and unique butterflies
in Chile, or I may say in the world, and as its habits are
undescribed I will give an account of them, as I had ample
opportunities of observing it. Though found at various
places in the mountains it seems local. Mr. Calvert has
taken it on the Campana Mountain near Quillota, and
Edmonds says he took it near La Union in the province
of Valdivia, but at what elevation he does not mention.
I first found it in the Villacura Valley east of the Pemehue
range at the end of January, where it was abundant at
about 3000 feet on grassy hillsides and flats covered with
long tufted herbage. In the morning when it first begins
to move, and before the wind has become strong, it may be
taken without much difficulty, though even then it 1s very
shy. Later it flies in the sun with such rapidity that it is
only by waiting in the line of flight that you can take them,
The brilliance of the shining silvery wings of this butterfly
make it a most beautiful and striking object when fresh,
but they soon become worn and broken, and a very large
proportion of those I took were not worth keeping. All
along the upper valley of the Biobio and on the Argentine
side of the frontier about Lake Aluminé I found it common
up to about 4000 feet, always on the grassy hill-sides but
never in the forest. It settles on the ground amongst tufts
of grass, and the larva is no doubt a grass-feeder. It was
common about San Martin and as far south as the valley
of the Limay, and when I re-crossed the Andes at the end
of February I saw one or two in the Aconcagua Valley
at about 6000 feet elevation.
22. Faunula stelligera. (Plate XV, figs. 1 f, 2 2.)
fF. stellagera, Butler, Trans. Ent. Soc. Lond., 1881,
p. 460, Pl. XXI, fig. 10.
I found this species common on the grassy hillsides and
ridges above timber-line near the Bafios de Chillan at
6000 to 7000 feet in December, when many specimens
were already worn. The flight and habits are essentially
like those of the Alpine grass Erebias, and as it never
occurs far from the dwarfed form of “ colihue” (a bamboo-
grass very like the Arundinarias of the Himalayas) which
covers large areas of these mountain-sides, I have little
The Butterflies of Chile. 283
doubt that the larve feed on this plant. The specimens
taken here average much larger than those which I after-
wards found abundant in January and February above
Loleo and near Pulmari, Quillen, and San Martin in
Argentina, always at elevations of 4000 to 6000 feet. They
vary a great deal, and some do not show as plainly as
others the toothed band on the hind-wing below which is
characteristic of the species, though Butler’s figure does
not show it. I have figured a pair from Lolco.
23. Faunula leucoglene. (Plate XIV, fig. 6 f.)
F. leucoglene, Felder, t.¢., p. 488.
This is essentially a high Alpine species, which I only
took myself on the top of the pass from Loico to Lonquimay
on January 27th at 8000 feet. Here it flew among stones in
a very exposed situation among a rich variety of Alpine
plants, sheltering itself from the high wind behind stones,
and so difficult to approach that in an hour's work I only
took one pair. It also occurs in the Cordillera near Santiago
at Condes, at 6000 feet according to Calvert, at 7000 to
10,000 according to Edmonds.
This species 1s curiously similar in appearance to Hrebiola
butleri, an Alpine species from New Zealand.
I saw a butterfly which looked like this on the Argentine
side of the Mendoza Pass near Las Cuevas, at about 10,000
feet, but failed to catch it.
24. Neosatyrus ambiorir.
N. ambioriz, Wallengren, Wien. Ent. Mon., iv, p. 36
(1860); 2d., Eug. Resa, Pl. VI, fig. 2 (1861).
2N. minimus, Butler, Trans. Ent. Soc. Lond., 1881,
p. 461, PIXE fies 7
This is a common forest insect wherever dense growth
of bamboos is found, and usually occurs m great numbers,
though hard to get in really fresh condition. I found it
at Bafios de Cauquenes and Bajios de Chillan up to 6000
feet in December, in the Pemehue Mountains at the head
of the Renaico Valley in January, and all along the
Argentine frontier at 3000 to 4000 feet in February. Both
sexes vary a good deal in the ocelli of the under-side, and
those taken at Cauquenes may belong to a different form
from those taken in Argentina and Pemehue, the males
when fresh having a distinct fulvous tinge at the base of
the wings above which is not seen in those from Pemehue
284. Mr. H. J. Elwes on
and Argentina. The females also of the Cauquenes form
are brighter-coloured above, much more yellow below, and
with larger ocelli. 1. minimus appears to be a small
starved variety of doubtful origin, and impossible to
describe from the specimen which is in the British
Museum collection.
25. Neosatyrus ? simplex.
Argyrophenga simplex, Butler, Trans. Ent. Soc. Lond.,
1881, p. 458.
Specimens of this agree with Butler’s type in the
British Museum taken by Edmonds above the Baths of
Chillan in March. I found it only in one place in a grassy
valley at about 5000 feet on the road from Lolco to
Lonquimay on January 27th. I took six or seven males
all in the same place which vary a good deal, most of them
being without the white dash on the hind-wing below which
exists in the type. It flies among the low bushes like an
Epinephele, and should be placed, I think, near Neosatyrus
ambworix, which it resembles in form and flight.
26. Neosatyrus vesagus. (Plate XIV, figs. 9 $,10 @.)
Erebia vesagus, Doubleday and Hewitson, Gen. Diurn.
ep, Panny ies also):
Homeonympha pusilla, Felder, t. ¢., p. 487.
Neosatyrus ochreivittatus, Butler, Trans. Ent. Soe.
Lond., 1881, p. 462.
NV. violaceus, id., t. c., p. 463, Plate XXI, fig. 8.
? N. hahni, Mabille, Miss. Cap. Horn. Lep., p. 3, Pl. I,
fig. 3.
? Erelna boisduvaliz, Blanchard, ¢. ¢., p. 32.
I am by no means sure of the above synonymy, as the
specimens before me vary a good deal and may belong to
two species.
First I have the type of vesagus in the Hewitson collection
from South America, which is undoubtedly the same as
violaceus and ochreivittatus, of which the types from Chillan
and Chile are in the British Museum. Also a pair of the
same from Valparaiso (Walker) in Mr. Godman’s collection ;
and a pair which I took at Coronel on December 19th, in
which the bands of the hind-wing below are less distinct.
Then I have a pair which I took near the Baths of Chillan
where Edmonds got the type of violaceus ; these, though
otherwise very like vesagus, have a distinct marginal band
The Butterflies of Chile. 285
of spots on hind-wing below, and seem to agree very
nearly with Felder’s type of pusilla. I have figured these
as none of the figures cited are satisfactory.
Then I have what seems a smaller species which has the
bands less distinct and the spots more so, which Butler calls
hoisduvalii, and of which two males and a female are in
Mr. Godman’s collection from Concepcion (Walker) close
to Coronel, where I took the larger form above-men-
tioned.* Then I have a single male from some part
of Chile which comes very near the figure of Hahnu
from Punta Arenas, which looks like, and probably is, a
starved southern form of the same. Until we get much
more ample material from intermediate localities in the
south the specific identity of these forms must remain
undecided.
27. Neosatyrus nycteropus. (Plate XIV, figs. 7 3, 8 2.)
Neosatyrus nyctexopus, Reed, t. c., Plate II, fig. 2.
I can find no other name for or description of this
species. What Reed speaks of in the explanation of the
plates as Hipparchia boisduvalvi is the same, but that name
cannot be identified certainly, and the figure Reed gives
of the under-side cannot I think be mistaken for that of any
other species. I can distinguish this from all others in Chile
known to me, by the distinct scalloped band on the hind-
wing below outwardly edged with white,in which Neosatyrus
hahnii, which I have referred to vesagus, resembles it more
than any other. On the upper-side both sexes usually (but
not always the male) have a fulvous submarginal band on
the hind-wing above not reaching either the costa or
inner margin of the wing.
I know this insect from two pairs taken at Coquimbo
by Walker in Collection Godman, a male and female
given me by Mr. Paulsen of Quillota, and three males
which I took on the scrubby hill-sides near Llai-llai on
January Ist. It seems, therefore, to be confined to the
coast region of Central Chile.
28. Neosatyrus reedii. ‘
N. reedit, Butler, Trans. Ent. Soc. Lond., 1881, p. 463,
Pl. XXI, fig. 9, var. fuscescens, id., t. ¢., p. 485.
This is a distinct species, which I did not take. It was
* What Blanchard called Boisduvalii came from the Straits of
Magellan and is probably same as Hahnui, but I cannot recognize it
by the description.
286 Mr. H. J. Elwes on
described by Butler from Reed’s collection without definite
locality, and is distinguished by the broad pale band on
hind-wing below. The figure of /Janiroides in Gay’s plate
(Fauna Chilena, Pl. III, fig 2), which Butler gives as a
synonym, is I think impossible to identify.
29. Tetraphlebia germainii. (Plate XV, fig. 5 2.)
T. germainii, Felder, t. ¢., p. 488.
Satyrus promaucana, Reed, t. ¢., Plate III, fig. 5.
I took the male of this in the woods below Chillan at
about 4000 feet at the end of December, and a single worn
female in the Villacura Valley at 3000 feet on January
25th. Itis a very distinct species. Reed says that it is
not scarce in the central provinces, but specimens are rare
in collections.
NYMPHALID.
30. HLuptoieta hortensia.
Argynnis hortensia, Blanchard, ¢.¢., p. 24.
This species does not seem common anywhere, but I
took specimens at Quillota, San Rosendo, and San Ignacio
in January.
31. Argynnis cytheris.
Papilio cytheris, Drury, Il]. Exot. Ent., ii, Pl. IV, figs.
3, 4 (1773).
Argynnis siga, Hiibner, Zutr. Exot. Schmett, figs. 677,
678 (1832).
One of the commonest butterflies all over the country,
and found from the sea-level up to at least 7000 to 8000 feet
near the Baths of Chillan. It extends from about Copiapo
in the north to as far south as the Straits of Magellan ;
and though it has several synonyms I cannot distinguish
any marked local varieties, except perhaps the one found
in the Falkland Islands. The largest and palest I have
is a female from 7000 feet elevation taken above timber-
line at Chillan, though probably it was bred in the valley
below. The smallest are those from the low forest country
near Mulchen and Tolten, though Edmonds says the con-
trary is the case. It occurred in Argentina as far south as
Nahuelhuapi.
32. Argynnis lathonioides.
A. lathoniwides, Blanchard, ¢. ¢., p. 22, Pl. H, figs. 1, 2.
The Butterflies of Chile. 287
? A. anna, id., t.¢., p. 23.
? A. dexamene, Boisduval, Bull. Ent. Soc. Fr., 1859,
De low
The only place where I took this species was in the
Horcones Valley near Puente del Inca in Argentine terri-
tory at about 10,000 feet, on December 12th, 1901, where
it was not uncommon. Most of my specimens were lost,
but Fitzgerald took one probably in the same locality, now
in the British Museum, and Edmonds found it in the
mountains above the Baths of Cauquenes at 6000 feet in
January. It is very rare in collections, and the females
might easily be mistaken for pale examples of the same
sex of cytheris. The male, however, is very unlike
the male of that species and very similar to its own
female. It seems to be a true mountain species confined
to the northern parts of the country, and perfectly distinct
from either cytheris or modesta.
33. Argynnis modesta.
A, modesta, Blanchard, ¢. ¢., p. 24, Pl. IT, figs. 3, 4.
I found this first above the Banos de Chillan at 7000 to
8000 feet on bare stony ridges above timber-line. Here
it was very hard to take, as it flew very fast aud quite close
to the ground, and was also very shy. I spent at least an
hour in securing two specimens, though they kept returning
to the same spots on little bare sandy places close to the
peaks but a little sheltered from the high wind. I did not
take the female, but have one from Edmonds, who found
it in the same place as the last but 2000 to 4000 feet higher ~
up. Ialso saw it and caught one specimen in Argentine
territory near Lake Aluminé at about 5000 feet on January
30th. It froze hard the previous night in our camp but
was very hot in the afternoon.
34, Pyramets carye.
Hamadryas decora carye, Hiibner, Samml. Exot.
Schmett, i, Pl. XLV (1806).
Common in the low country, but not found by me in
the mountains.
35. Pyrameis terpsichore.
Vanessa terpsichore, Philippi, Lin. Ent., xiv, p. 266
(1860).
TRANS. ENT. SOC. LOND. 1903.—PART III. (OCT.) 20
288 Mr. H. J. Elwes on
Taken at San Rosendo, San Ignacio, Lolco, and on
Lake Llanquihue in December, January, and February,
and at San Martin in Argentina up to about 3000 feet.
LYCHANIDA.
36. Scolitantides collina.
Lycena collina, Philippi, t. ¢., p. 270.
LL. lyrnessa, Hewitson, Ent. Mo. Mag., xi, p. 107
(1874).
? Scolitantides plumbea, Butler, Trans. Ent. Soc. Lond.,
1881, p. 486.
This belongs to a group which is represented in the
Andes of Bolivia by S. speciosa, Stgr., in Peru by vapa, Ster
and in Ecuador by an unnamed species of which there
are specimens in the British Museum. I am uncertain
whether plwmbea, Butler, is identical; Edmonds thought it
was only a variety, and though the type is larger and
duller in tint, I find no character in it to enable me to say
decisively. I took a single specimen exactly like the
type of plumbea, but am uncertain of the locality. I found
the species very abundant at 5000 to 6000 feet near the
Baths of Chillan in December, and also took it in the
Sierra de Pemehue in January. It frequents bushy ground
and also the bare ridges above timber-line. Philippi says
he took it on the hills near Santiago.
37. Scolitantides andina.
S. andina, Calvert, An. Univ. Chile, xxxiv, p. 832
(1894).
¢ Lycena endymion, Blanchard, t.c., p. 37, Pl. II,
fig. 3 a, b.
I cannot identify this species with certainty, as, if
Blanchard’s plate is correct, the under-side is different from
that of Calvert’s species which I know from two females sent
by him to the British Museum, where they stood without
name. I found the same species common near Puente
del Inca on December 11th, flying on bare ground among
grass tufts at about 9500 feet near the entrance to the
Horcones Valley. It differs from collina in the colour above,
which is more greenish in the male and grey in the female.
Beneath the markings are very like those of collina, but
the female has no red on the under or upper surface of
The Butterflies of Chile. 289
the wings. Calvert’s species was taken near Condes above
Santiago at 3000 metres elevation.
Endymion is said by Blanchard to come from Coquimbo.
It appears that Kirby in Cat. Diur. Lep., p. 377, gave
the name of Szbyl/a to Blanchard’s figure, because the name
endymion was pre-occupied. I prefer to use the name of
andina.
38. Scolitantides chilensis.
Lycena chilensis, Blanchard, ¢. c., p. 37, Pl. II, fig. 4
CaO!
2 Polyommatus atahwalpa, Wallengren, Wien. Ent.
Mon., iv, p. 37 (1860).
I found this species common at Banos de Cauquenes in
December. It is also found, according to Edmonds, at
Valparaiso and Copiapo.
39. Lampides trigemmatus.
L. trigemmatus, Butler, Trans. Ent. Soc. Lond., 1881,
p: £68.
This species seems to be peculiar to the north of Chile.
Besides the types in the British Museum there are four
specimens from Tarapaca. It is nearly allied to L. telicanus
of Europe.
40, Thecla bicolor.
Lycxna ? bicolor, Philippi, t.¢., p. 269.
2 Theela quadrimaculata f , Hewitson, Ent. Mo. Mag.,
xi, p. 106 (1874).
I am not convinced of the specific distinction of this
species from the next; it may be a mountain form of it.
The only specimen I have is labelled Puente del Inca,
but Iam inclined to think that this label has been misplaced.
The specimen is smaller, and on the under-side somewhat
different from qguadrimaculata. The males of both forms
have a large sexual patch in the fore-wing, which has led
Butler to put them in the genus Callipsyche, Scudd. The
type was taken near Santiago.
41. Theela quadrimaculata.
2 7. quadrimaculata, Hewitson, /. ¢.
Hewitson appears also to have doubted the distinctness
of this from the last, as he has put the female type as the
290 Mr. H. J. Elwes on
male of bicolor, I took two males of what Butler calls
quadrimaculata at Coronel and one at Banos de Cauquenes,
both at low elevations.
42. Strymon americensis.
Thecla americensis, Blanchard, t.c., p. 38.
I took a pair on the road up to the Baths of Chillan at
about 2000 feet on December 20th, and found it also near
Temuco and at San Ignacio in January. Edmonds took
it at Valparaiso, Valdivia and Cauquenes. The male
has the same sexual patch as bicolor, with which I think
it is congeneric,
PIERIDA.
43. Heliochroma leucothea.
Papilio (D.) lewcothea, Molina, Saggio sulla Storia
Naturale del Chili, libr. iv, p. 347 (1782).
Pieris gayi, Blanchard, t.c., p. 10, Pl. I, fig. 4.
Though Edmonds says this is common in the Cordillera
of the Cauquenes hacienda in January, I was informed
by M. Germain, the veteran entomologist of the Museum
at Santiago, who knows the insects of Chile very well
though he has never written on the Lepidoptera, that the
species was confined to the coast, where it comes out early
in spring. I only saw it near Coronel in December flying
among bushes where I was unable to take it.
44. Terias chilensis.
fT’. chilensis, Blanchard, t.c., p. 17, Pl. J, fig. 5 a, 0.
I only saw it near Llai-llai on January Ist in irrigated
fields. It seems to be common at low elevations.
45. Callidryas drya.
Papilio drya, Fabricius, Syst. Ent., p. 478 (1775).
Callidryas drya, Butler, Lep. Exot., p. 61, Pl. XXIII,
figs. 5-8 (1871).
C. amphitrite, Blanchard, t. ¢., p. 20, Pl. V, figs. 1, 2.
A few were seen in the lower part of the Aconcagua
Valley. Edmonds says it is common at Valparaiso and
is found at Copiapo, and occurs almost throughout the
year. He found the larva on Cassia.
The Butterflies of Chile. 291
46. Colias lesbia.
Papilio lesbia, Fabricius, t.¢., p. 477.
This was abundant and very variable at Buenos Ayres,
and was found as high as 9000 feet near Puente del Inca.
I also took a single albino female on the Chilean side of
the pass at about 9000 feet on December 18th. I cannot
find that the species has been recorded previously west of
the Andes, and I did not find it anywhere in Argentina
along the frontier.
47. Colias vauthiert.
C. vauthieri, Guérin, Voy. de la Coquille, Pl. XV,
fig, 2 (1829).
a C. rutilans, Boisduval, Sp. Gen. Lep., p. 642, Pl.
XIX, fig. 3 (1836).
C. minuscula, Butler, Trans. Ent. Soc. Lond., 1881,
p. 470; Pl xox fo
C. cunninghami, id., t.c., p. 471.
Very abundant everywhere in Chile and in Argentina
along the frontier from sea-level up to 8000 feet and
probably higher. Extremely variable in size and in the
colour and markings of the female; of which, however,
I have never seen an orange form. I could not see that
climate had any influence on the species, those from the
cold and windy plains about Lake Nahuelhuapi being
as large and bright as those from the damp warm forest
near Temuco.
The spring brood, however (minuscula, Butl.), which I
did not find, is small, and has a narrower black border on
both wings, and Cunninghami, Butl., is a narrow-winged
southern form from Sandy Point.
48, ? Colias cuwanthe.
Colias ewxanthe, Feld. Reise Nov. Lep., u, p. 196
(1865).
I took what I believe to be this species in the Horcones
Valley near Puente del Inca at about 10,000 feet on
December 12th, but the specimens were unfortunately lost
on the road when starting for Chile, and I have never
been able to recover the bag which contained them. As
it is a native of the mountains of Bolivia and Peru, there is
nothing improbable in its extending southwards.
292 Mr. H. J. Elwes on
49, Phulia nymphula.
Pieris nymphula, Blanchard, t.c., p. 14, Pl. I, fig. 3
C10:
? Phulia nymphea, Staudinger, Iris, vii, p. 49 (1894).
Taken near Puente del Inca in the Horcones Valley
at 10,000 feet on December 11th, and on the pass in
February at 11,000 feet. Chilean specimens are some-
what larger than those in the British Museum taken in
Bolivia by Sir M. Conway, and what Staudinger describes
as nymphea seems inseparable.
50. Tatochila autodice. (Plate XII, figs. 1—4.)
42 Synchloe autodice, Hiibner, Samml. Ex. Schmett,
i, Pl. CXX VII, 1—4 (1816).
2 Pontia mercedis, Eschscholtz, Kotzebue’s Reise,
ili, p. 215, Pl. IX, figs. 22 a, b-—36 (1821).
var.? Pieris microdice, Blanchard, t. ¢., p. 14. (Plate
MT igs) 5,76.)
I have had great difficulty in naming the Tatochilas I
found in Chile on account of their variation, and because
Blanchard has described three forms without figures which
I cannot identify certainly.
The first is a well-known species which occurs all over
Argentina and Chile, and is known as autodice, Hiibn.
This I took near Buenos Ayres. When I crossed the pass
into Chile, I saw flying on the head of the valley at about
9000 feet a small silvery form, of which I have figured a
f on Plate XII, fig. 5. This had a rapid flight and
appears to be distinct, and to be what Blanchard called
microdice. I could not find the female here, but two
months later found what I think is the same species fairly
common at 4000 feet and upwards on the Argentine
frontier near Lake Aluminé, and as far south as Lake
Nahuelhuapi. Of these I have figured a female from
Lake Quillen, fig. 2, one from Nahuelhuapi, fig. 6, and
one from the Sierra de Pemehue, in Chile, fig. 1. A male
from Pulmari is like the figure which Mabille calls theodice
in Miss. Cap. Horn Lep., xi, fig. 2.
51. Tatochila demodice. (Plate XII, figs. 7 2, 8 ¥.)
2 Pieris demodice, Blanchard, ¢.¢., p. 13.
Tatochila argyrodice, Staudinger, Hamb. Magal.
Sammelreise Lep., p. 14, fig. 11 (1899).
? Pieris theodice, Boisduval, Voy. Astr. Lep., p. 51
(1832), 2 dimorph. apud Staudinger, /. ¢.
The Butterflies of Chile. 293
This species was common at Bajios de Cauquenes in
December, in the Pemehue range in January, and down
south as far as Nahuelhuapi in February, practically at
all the localities where I found the last species. I have
figured a pair from San Martin, Pl. XII, figs. 7 ¢,
8 ?. What I take to be a form of the same species is
found as far south as the Straits of Magellan, where it
is known as argyrodice, Stgr. Of this I have figured a
pair (Pl. XII, figs. 9 ¢, 10 2) taken by Walker at Punta
Arenas. These are probably if not certainly the same as
what Staudinger calls theodice, Bdv., of which he makes
demodice, Blanch., a synonym, but a large series are
necessary to understand the variation of this species.
52. TLatochila theodice.
Preris theodice, Blanchard, t.c., p. 12, Pl. I, fig. 1 a, 0.
Tatochila blanchardii, Butler, Trans. Ent. Soc. Lond.,
1881, p. 472, Pl. XXI, fig. 15.
Lastly we have a species which is perhaps a second
brood of one or the other species usually identified with
theodice, Blanch. (nec Bdv.) = Blanchardii, Butl.
This is easily distinguished by the double bar at the
end of the cell, and extends from as far north as Islay in
Peru to about lat. 38°, where I took it at San Ignacio in
January. It seems to be common at Valparaiso, ‘and is in
Mr. Godman’s collection from Valdivia. Xanthodice, Lucas,
is another mountain species which is common in Ecuador
and Bolivia, but has not been found in Chile, though
Mabille figures under this name in the Miss. du Cap.
Horn. Lep., Pl. I, fig. 1, a species which appears to me
very near argyrodice, Steger.
PAPILIONIDA.
53. Papilio bias,
P. mas, Roger, Bull. Soc. Linn. Bord., 1 (1826).
I saw this in the Botanical Gardens at Santiago, but
not elsewhere, though Edmonds records it as common at
Valparaiso in successive broods from October to June.
HESPERIIDAL.
54. Hesperia fusca. (Plate XIII, figs. 1 f, 2 2.)
HT, fusca, Reed, Mon. Marip. Chil., p. 81 (1877).
This was common near Lolco in one place at the
confluence of the Lolco with the Biobio river at about 2500
294 Mr. H. J. Elwes on
feet. It seems a rare species, but I have a pair from
Copiapo, and Reed says it occurs in the province of
Valdivia. The male figured is from Lolco and the female
from Copiapo.
55. Hesperia americanus.
Syrichthus americanus, Blanchard, ¢. ¢., p. 44, Pl. II,
fig. 10.
I took this at the Baths of Cauquenes and at Coronel,
but it does not seem common anywhere.
56, Hesperia fulvovittatus.
Pyrgus fulvovittatus, Butler, Trans. Ent. Soc. Lond.,
1881, p. 475.
This was described from a single specimen in Edmonds’
collection, locality unknown. There are some from Callao
collected by Walker in the British Museum collection
which seem very close, and it may not be a Chilean species.
57. Hesperia trisignatus. (Plate XIII, figs. 3 2, 4 2.)
Scelothrix trisignatus, Mabille, Bull. Ent. Soc. Fr.,
1875, p. cexiv.
? Hesperia notata, var. valdiviana, Reed, l.c., p. 81.
I took a single female at Quillota and another near
Llai-llai on January Ist. Afterwards I found it abundant
on a dry bushy plain below the Quillen lake in Argentina
on February 2nd at about 3000 feet. Here it was flying
about small bushes close to the ground. The male figured
is from Quillen and the female from Quillota. There are
specimens in the British Museum collection from Callao
and Coquimbo taken by Walker.
A co-type of valdiviana in coll. British Museum looks
like a variety of the same species but may be distinct.
58. Hylephila fasciolata. (Pl. XIII, fig 7 2.)
a Hesperia fasciolata, Blanchard, ¢.¢, p. 42, PI. II,
hie: (:
DME SLONALH, 10., UC.
IT am not sure that I took this myself, though I found
a pair in one of my boxes without locality. It seems
rarer than the next species, though Kdmonds took it at
Copiapo, Cauquenes, Valparaiso and Valdivia.
The Butterflies of Chale. 295
59. Hylephila fulva. (Plate XIII, figs. 5 f, 6 9, 8 9, var.)
Hesperia fulva, Blanchard, t.c., p. 48, Pl. III, fig. 8.
I found this common in many places and have specimens
from Coronel, Cauquenes, Llai-llai, San Ignacio and
Lolco. The female from the Renaico Valley (fig. 8)
belongs. to the form mentioned by Butler as being larger
and with more markings than the other. The originals of
figs. 5 and 6 are from Lolco.
The species comes near phylewus, Drury, which has a
very wide range in America and extends as far south as
Buenos Ayres. A specimen from thence in the British
Museum seems very near fulva, Both these species
frequent grassy places in the open.
60. Argopteron aureipennis.
Syrichthus aureipennis, Blanchard, t.c., p. 40, Pl. I,
figs. 5 a, 5 b and 6.
I took this in the dense forest on the shores of Lake
Lacar near San Martin on February 8th, where it settles
on the bamboo and has exactly the same habits as the next
species. On the under-side the male is like puelmex, but
the female has the fore-wing below much more like
Jruticolens. 'The white spots on the under-side shown in
Gay’s fig. 5b are not in my specimens, and as Gay says, “sin
mancha alguna,” I presume this is an error of the artist.
I think I also saw this species at Puerto Blest on Nahuel-
huapi, and Edmonds found it at Valdivia.
61. Argopteron puclme. (Plate XIII, figs. 11 g, 12 2.)
Cyclopides puelmex, Calvert, Ent. Mo. Mag., xxv, p. 34
(1888).
I found this beautiful species very abundant among the
bamboo in the dense forest on the Pemehue range at
3000 to 4000 feet between Maitenes and Chilpa on January
26th. The flight is quite peculiar, and the insect settles on
bamboo leaves and also the orange flowers of Alstremeria
aurantiaca in the sunny openings of the forest. It seems to
represent aureipeniis in the north, but has a limited range.*
62. Butleria fruticolens. (Plate XIII, figs. 9 f, 10 2.)
Cyclopides fruticolens, Butler, Trans. Ent. Soc. Lond.,
HSSI- pp. 477, E> Xox tomes
* It seems impossible to reproduce by chromo-lithography the
shining gold of the under-side which makes this such a conspicuous
insect when flying.
296 Mr. H. J. Elwes on
Steropes tripunctatus, Mabille, C. R. Ent. Belg., xxxv,
p. lxiv (1891).
I found this along the frontier at 3000 to 4000 feet in
forest from the Pemehue range as far south as Puerto Blest.
The female figured is from the latter locality and the
male from Chilpa. It does not seem to vary in the
mountains so much as on the coast, where Edmonds found
it at Corral in March. Butler has described three varieties
as tractipennis, quadrinotatus and pulcher, which differ in
the number of spots on the upper-side and in other minor
characters. In Mr. Godman’s collection is a male from
Chile marked “tripunctatus=type, Mab.” This is described
as coming from high mountains.
63. Butleria philippit.
Cyclopides philippui, Butler, l.c., p. 479, Pl. XXI,
3.
o
>
Very near the last but has an additional silver stripe
on the hind-wings below which I have never found
in fruticolens. I did not take this species, which occurs
at Las Zonas in Valdivia.
64. ? Butleria sotoi.
Cyclopides sotoi, Reed, Mon. Mar. Chil., p. 86 (1877).
The only specimen of this in British Museum marked
“type” seems distinct from either of the last by the
yellowish colour at base of both wings below, and the
presence of a spot on the hind-wing below. Reed
took it in the hacienda of Cauquenes, but does not
say at what elevation. I found a single worn speci-
men, which I believe is identical, in thick forest by the
waterfall at Las Trancas, the last station below the Baths
of Chillan, at about 4000 feet, on December 24th, but
neither this nor the type are in good enough condition to
figure.
65. Butleria flavomaculatus.
2 Syrichthus flavomaculatus, Blanchard, ¢.c¢., p. 44, Pl.
LUBE ree SR oni
? Butleria vicina, Reed, t.c., p. 88.
The next three or four species are nearly allied to each
other, but I have not sufficient material to define them
accurately, and their synonymy is doubtful.
What seems to agree best with Blanchard’s plate is a
The Butterflies of Chile. 297
small species which I took on the edge of the forest at
Lake Quillen, and which is distinguished from the others
by the spots of the under-side being yellow and not white
or silvery. This is in Mr. Godman’s collection from
Staudinger as paniscoides, Blanch. The description of
this is not sufficient in my opinion to identify it by, or to
separate it from flavomaculatus on the material before me.
66. Butleria valdivianus. (Plate XIII, figs. 15 2,16 9,
ae 2.
Syrichthus valdivianus, Philippi, Linn. Ent., xiv, p.
272 (1860).
? Carterocephalus exornatus, Felder, Reise Nov. Lep.,
Pp; 02 PEG XERING fies; FS; 19° (1867):
? Butleria paniscordes, Reed, t. ¢., p. 82.
I found this species common both in the Pemehue range
at about 3000 feet, and in the marshy places on the edge
of the forest at Quillen, from which locality I figure a pair
(figs. 15 and 16). The original of fig. 17 is from Lolco.
It varies extremely in the spots of the hind-wing below,
which, however, seem to me to distinguish it from the
last species by their silvery white colour, and from the
next by the irregularity of the marginal series, which are
sometimes faint or absent. On the inner and costal
margins of the hind-wing below, these spots often coalesce
into a streak; as is also sometimes the case in the next
species. Hxornatus is described as from Valparaiso.
67. Butleria polyspilus. (Plate XIII, fig. 18 2.)
2? Carterocephalus polyspilus, Felder, Verh. Z. B. Wien.,
xu, p. 495 (1862).
I identify this with great doubt. In Mr. Godman’s
collection there are three’ specimens named C. exornatus,
Feld. I am not certain whether it is distinct from the last,
but have found it in much more open grassy places, and
took it in the valley of the Traful river in Argentine
territory as well as at Quillen. The specimen figured is
from the Traful Valley.
68. Butleria bissexguttatus. (Plate XIII, figs. 13 Z, 14 2.)
2 Steopes (sic) bissexguttatus, Philippi, Linn. Ent., xiv,
p. 272 (1860).
I found this first in the forest at Temuco on January
3rd, and afterwards at several places in the Pemehue range
List OF CHILEAN BUTTERFLIES SHOWING
298 Mr. H. J. Elwes on
and Argentina always in or close to the heavy forest. It is
easily distinguished by the colour of the under-side from the
allied species above. The specimens figured are from Chilpa.
69. Thanaos funeralis.
Nisoniades funeralis, Scudder and Burgess, Proc. Bost.
Nat. Hist. Soc., xiii, p. 293, fig. 7 (1870).
Butier includes this on Edmonds’ authority, but no
locality is given for it, and I think the identification highly
dubious. In the forest on the Pemehue range I am
certain that I saw a Hesperiid belonging to this group,
but did not succeed in taking it.
THEIR DISTRIBUTION AND RANGE.
OOD CON He
25
26
| 27
28
29
|
A E | ELSE-
a | CENTRAL. ze WHERE.
4 | n |
SATYRIDE. |
Elina lefebvret, Guer..............++- | x low dese |
», wanessoides, Blanch. .........| ... |x low ese |
», neomyriotdes, Blanch........ | |x middle eases |
MEA COLUCTIU UH S senate teeters | ... |x middle
Pedatliodes flora, Phil. ............... |... |x low bes a
Epinephele edmondsii, Butl......... ... |x middle ... | Argentina
e janirioides, Blanch. ... |x low Peers
a monachus, Blanch...... ... |x lowand middle
tristis, Guér. ............| x |x low and middle |
a COCLEU, (GUI aecceneneeere ; ... |X lowand middle| ...
Neomenas cenonymphina, Butl...| ... |x low
5 Fractifascia, Butl.......| ... |x middle
- servilia, Wall............ PS nLOW, [Nera
ae wallengrenii, But). ...) ... |x middle ea
an WNOyNAtH, TM. S......0.06- (ieeeal scal ow: eae |
f.. edmondsii, Butl......... x middle leita
humilis, Feld Pp seaccaic ... |x lowand middle| x |
Cosmosatyr us leptoneuroides, Feld.| ... | x middle ligeee
», var. plumbeolus, Buti.| ... |x middle Weer
es chiliensis, Guér...... |... |x lowand middle} x |
aA monticolens, Butl....| .... |x middle Sea ||
», var. williamsianus, Butl. | ... ane x
Argyrophorus argenteus, Blanch. | x? x middle
Faunula stelligera, Butl............. | ... |x middle
es leucoglene, Feld. ......... eo. [Selene |
Neosatyrus ambiorix, Wall. ...... | ... |x lowand middle] ... |
:: simplex, Butl. ......... | ... |x middle es,
+ vesagus, Doubl.......... |... |x low Saal
- nycteropus, Reed. ......| ... |x low 486
si TCCUUYsWDUble weeeasecece x low WSS
Tetraphlebia germainii, Feld. x middle
|
The Butterflies of Chile. 299
Zz CENTRAL. z WHERE
; eres
— = = — | —
NYMPHALID. |
30 | Euptoicta hortensia, Blanch. ...... x low | aise 7
\¢ ee Se : { Falklands and’
31) Argynnis cytheris, Drury ......... x x lowand middle| x “(Magellan
32 - lathonioides, Blanch. ... |x middleand high we
33 modesta, Blanch. ......... --. |x high
34 | Pyrameis carye, Hiibn. ............ x |x low
35 of terpsichore, Phil..........| ... |x low and middle |
LYCENID. |
36 | Scolitantides collina, Phil. ......... |x middleandhigh| ...
37 me andina, Calv. ......... x high eee
38 a chilensis, Blanch. ...| ... | x low
39 | Lampides trigemmatus, Butl....... | x | me
£00), Thecla, bicolor. Phil sseesesee sere: |... |x middle? high #
pal > quadrimaculata, Hew. =] ee) oo low
42 | Strymon americensis, Blanch....... x low |
PIERID&. | | |
43 | Heliochroma leucothea, Mol. ...... x low
AS) Tervas chitensts, Blanche... ee |< lows:
45 | Callidryas drya, Faby. ............ x |x low
CNT CONTA UAREUO, MEN OTES pacecece ecohosece | ... |x high? Ex
47 >, vauthiert, Guér. .............. | Seale low and middle x ae
48 55 hetnantie, Belds sesssss.-- x?) x high Bolivia
49 | Phulia nymphula, Blanch..........| -.. | x high | do.
50 | Tatochila autodice, Hiibn. ......... x lowand middle| ... | Argentina
or var. ? microdice, Blanch. x high bax
51 53 demodice, Blanch. ...... |x lowand middle| x
Bz xp theodice, Blanch. (nec
Bdvi)fe- 2 ee eee x |x low Ed)
PAPILIONIDS.
) 0s) | Papilio nas; Roger .......<..:-.0--- x low
|
HESPERIIDE.
bf | Hesperia fusca, Reed. .2.-.2.-..+s-- | x |x lowand middle
55 a americanus, Blanch. ...| ... | x low
56 5 Julvovittatus, Butl. ...... [esate a8:
57 33 trisignatus, Mab. ......... | ... |x lowand middle 3
58 | Hylephila fasciolata, Blanch....... | .- | x low Me
59 Meteo pice, blanch. 5...2:2..-6. | +? x lowand middle | Ba
60 | Argopteron aureipennis, Blanch.. x middle
61 - puelmex, Calv. ......... x middle
62 | Butleria fruticolens, Butl. ......... x middle
63 so SUC LOD, Naa eeeece babe x low
64 35 GOLOTMILCED Sore oe eniccanc rose . x middle
65 3 flavomaculatus, Blanch. _x middle |
66 re valdivianus, Phil.......... _... x lowand middle
67 #5 ? polyspilus, Feld.......... bE: ... x lowand middle
68 bissexguttatus, Phil....... ee fe low and middle
2 Thanaos Juneralis, Scudd.
(
EXPLANATION OF
300 )
Prates XTI—XYV.
Pisce Xe
Fic. 1. Latochila autodice,
Lo
” ”
3.
4, on 5
5, a microdice,
6. bP) 9)
ile democlice,
9. a argyrodice,
10. “
9, Pemehue.
@, Quillen.
d, Cauquenes.
Se ? oP]
¢, Above Juncal.
9, Nahuelhuapi.
¢, San Martin.
2 P) of} 7
¢, Punta Arenas (Walker).
2,
bh) 9
Prare Xin:
Fia. 1. Hesperia fusca,
2. 9 0
3. cs trisignatus,
A, ” ”
5. Hylephila fulva,
6. ? ”
ie Reis
dee *s fasciolata,
8. 5 fulwa, var.?
9. Butleria fruticolens,
10. ” ”
11. Argopteron puelme,
12. »
13. Butleria ? bissexquttatus,
14. ” ”
15. a valdivianus,
16. % s
Wet?” s, rs
18. y } polyspilus,
3d, Loleo.
2, Copiapo.
d, Quillen.
2, Quillota.
g, Loleco. -
oa
4g, Quillota or Cauquenes.
2, Renaico.
9, Port Blest.
3, Chilpa.
f, Quillen.
9,
3;
2 >} 5
g, Lolco.
OF erat:
7
inigey, 1h,
SS
SE
2 gm
10,
Explanation of Plates. 301
BEATE: ORVE
” bP 9
. Elina calvertii, n. sp.,
1) ” ”
5 neomyriordes,
. Fawiiula lewcoglene,
. Neosatyrus nycteropus,
. Neomenas ? inornata, n.sp., ¢, Batios de Cauquenes.
g ) re) yy) ”
3d, Banos de Chillan.
g b] ” 29 ”
3g, Chile Mus. Crowley.
, Lolco Pass. ~
6, Llai-lai.
Pol ies
» ”
55 vesaagus, 3d, Banos de Chillan.
ye) ~ ” 2? ) ” ” ”
PLATE eX Ve
Faunila stelligera,
” 9
Cosmosatyrus leptonenroides, &,
” 9
” ?
Tetraphlebia germainii,
Cosmosatyrus monticolens,
3, Lolco.
e 9
2
2, Aluminé.
?, Villacura Valley.
¢, Chillan Valley.
dé, Pulmari.
? > ”
” ”
He chiliensis, ? var. ¢, Lake Nahuelhuapi.
?
” ” ? 9) >
—— rs <A te.
2 eee a | ip aS aT Eg
7 = : 7 / e
oe
(=-303) )
XVII. On the genus Theodosia and other Eastern Golia-
thides, with descriptions of some new species. By
OLIVER E. JANSON, F.ES.
[Read June 3rd, 1903.]
THEODOSIA, Thoms.
Westwoodia, Cast., Helionica, Thoms., Atopocerus, Kz.
THIS genus was established by Castelnau in 1873 under
the name of Westwoodia, with howitti, Cast., as its type.
His generic name being preoccupied, Thomson in 1880
substituted that of Theodosia and at the same time erected
what he considered a distinct genus, Helionica, for the
reception of the species westwoodi, Thoms. ; the characters,
however, that he relied upon to separate it from Theodosia
are certainly not of generic value, and the two genera
must therefore be merged into one. In 1888 Kraatz,
evidently ignorant of the fact that Thomson had already
eight years previously re-named the genus, published the
name Atopocerus as a substitute for the preoccupied one
of Westwoodia, his name therefore must also be sunk as
a synonym.
The genus is very closely allied to Phedimus, Waterh.,
of which the type is cwmingi, Waterh., and Bates has
regarded them as generically inseparable, but this view
has not been adopted by subsequent authors, and I
certainly think they are sufficiently distinct to warrant
separation; besides a very different general facies and some
minor characters, in the male of all the Theodosia species
the head is flat with the horn arising from the extreme
front margin of the clypeus, the thorax has no distinct
lateral margin on the anterior part to separate the upper
and under surfaces, and the horn is formed as a prolonga-
tion of the exceedingly convex disk, the anterior tarsi are
also slender and longer than the tibize, whereas in Phedimus
the head is excavated and has the clypeus projecting
beyond the base of the horn, the thorax is strongly and
completely margined at the sides and with the horn arising
from close to the anterior margin, and the anterior tarsi
are stout and distinctly shorter than the tibiez.
TRANS. ENT. SOC. LOND. 1903.—PART Ill. (ocr.) 21
304 Mr. O. E. Janson on the genus
Considerable trouble has been experienced in the identifi-
cation of the species of this genus largely owing to the
confusion occasioned by Dr. Kraatz in using the preoccu-
pled name westwoodi, and in giving distinctive names to
slight variations and degrees of development. After a
careful study of the rather large material I have had an
opportunity of examining and the various descriptions, I
am only able to adopt five out of the nine names that
have been published, as representing distinct species, and
these only, with the two new species now added, are in-
cluded in the following tabulation which I have drawn
up of their most prominent male characters. Female
specimens are rarely received, and it is very difficult, or
almost impossible, to decide with any amount of certainty
to which species they pertain.
1. Cephalic horn acute at apex.
A, Anterior tibiex bare.
a. Angles of clypeus strongly
produced, spiniform. . . howitli, Cast. 8. Borneo.
b. Angles of clypeus only
slightly produced, obtuse.
aa. Thoracic horn without
prominences near apex. westwoodi, Thoms, N. Borneo.
bb. Thoracic horn with a
small prominence on
each side near apex. . magnifica, Roths. N. Borneo,
B. Anterior tibiae densely pilose
oninnerside. . . . . . perakensis, Moser, Perak.
2. Cephalic horn bifid at apex.
A, Anterior tibiz densely pilose
oninnerside. . . . . . rothschildi, Jans. N. Borneo.
B. Anterior tibize bare.
a. Thorax green, sub-opaque. telifer, Bates. N. Borneo.
b. Thoraxbrassy, very polished auwrata, Jans. N. Borneo,
The other names that have been given in the genus
are :—
T. westwoodi, Kz. This is evidently identical with magni-
fica, Roths., the latter name having priority. I think this
species may perhaps prove to be merely a variety of west-
woodi, Thoms., when we have an opportunity of examining
a larger series.
H., waterstradti, Kz. This is the larger highly developed
Theodosia and other Kastern Golrathides. 305
male form of westwoodi, Thoms., and should not bear a
separate name.
H., denticornis, Kz. This is a slight variety of the same,
and unworthy of a distinctive name.
HT, viridicollis, Kz. This is described as a variety of
westwoodi, Thoms., but is stated to have the angles of the
clypeus acutely produced and to come from South Borneo,
in which respects it agrees with howitti, Cast., and I think
it is probably a small and lesser developed form of the
male of this species.
Theodosia rothschildi, n. sp.
gd. Viridis, subnitida, elytris flavescentibus ; capite cornu sat
brevi, apice profunde bifido, supra igneo-eupreo ; thorace cornu
robusto, apice bifido, dilatato, igneo-cupreo ; tibiis anterioribus intus
dense flavo pilosis.
Long. (cornu excepto) 25 mm.
Mount Kina-balu, N. Borneo,
This species most nearly resembles 7’. ¢elifer, Bates, but
is to be at once distinguished from all the other known
members of the genus, except perakensis, Moser, in having
the inner-side of the anterior tibize densely clothed with
fine pale yellow pubescence. The horn on the head is
a little broader than in ¢elzfer and the thoracic horn is
less flattened and more dilated at its apex, the sides of
the thorax are a little more prominent in the middle, the
scutellum is much broader at the base, as in magnifica,
Roths., the elytra are broader and more rounded at the
sides before the middle, and the mesosternal process is
broader and more rounded at its apex. The sculpture is
very similar to that of felifer except that the punctuation
of the scutellum is very dense and confluent. The legs
are deep green with the upper-side of the femora flaves-
cent, otherwise the colour is similar to that of ¢elifer. The
length of the prothorax including the horn is 17 mm., the
cephalic horn is 6 mm. in length.
T. perakensis, Moser (Berl. Ent. Zeit., 1902, p. 379), is
the only other described species having pubescent anterior
tibize, but in that species the cephalic horn is acute at
the apex, not bifid as in rothschildt.
Theodosia aurata, n. sp.
¢. Aureo-viridi-flavescens, micans ; capite igneo-cupreo, cornu
brevi, apice bifido ; thorace dorso subtilissime punctato, nitidissimo,
306 Mr. O. E. Janson on the genus
cornu brevi, apice paullo emarginato, igneo-cupreo; scutellum
magnuim, basi latiusculum.
Long. (cornu excepto) 20 mm.
Mount Kina-balu, N. Borneo.
Most nearly allied to 7’. telifer, Bates, but differs in
having the thorax shorter and much broader, especially
across the middle, and more abruptly narrowed in front,
the dorsal surface is very shining, being much less closely
and more finely punctured, the horn is short, flattened
and with a slight notch, but not dilated, at its apex, the
scutellum is much broader at its base than in felifer and
less strongly punctured, the elytra are also more feebly
punctured and have the apical sutural angles strongly
rounded. The colour is a rather golden yellow with a
shght metallic greenish tint, the thorax is of a more brassy
tint and very polished on the dorsal part, with the horn
and median anterior part fiery coppery, whilst in tedifer
it is green and the entire surface sub-opaque by reason of
the dense granulose punctuation. The head is coppery
with the sides of the clypeus green and the horn greenish
brassy and darker at its apex. The under-side is more
strongly tinged with green and the legs are of the same
colour as the elytra. The second and third abdominal
segments are deeply impressed in the centre. The length
of the cephalic horn is 4 mm., and of the prothorax,
including the horn, 10 mm.
Mycteristes imermas, vi. sp.
3d. Gracilis, fuseo-olivaceus, opacus, parcius flavo-squamulosis ;
capite sub-nitido, inermi, clypeo apice medio anguste reflexo, ro-
tundato ; elytris planiusculis, sub-carinatis; subtus nitidus, viridi-
tinctus, griseo-squamulosis, Long. 14 mm.
Mount Kina-balu, N. Borneo.
Narrow, elongate, above dull olivaceous brown with sparse ap-
pressed, yellowish, coarse, squamiform sete. Under-side and legs
red-brown, shining, and with a greenish or brassy tinge in places,
more thickly clothed than the upper-side with finer and longer grey
setae, tibiae coppery.
Head shining, coarsely punctured, a small shghtly elevated smooth
spot near the base, elypeus convex in the centre, the side margins
elevated in front of the eyes, the apical margin narrowly reflexed
and rounded in the centre. Thorax much narrower than the elytra,
the basal two-thirds parallel-sided, then obliquely narrowed to the
Theodosia and other Eastern Goliathides. 307
front, strongly but rather narrowly lobed behind, the basal angles
right angles, the front margin slightly raised in the centre and green,
the disk with a slight curved carina on each side, convergent in
front, the elevated parts almost impunctate, the other parts with
rather close coarse irregular semi-circular punctures. Scutellum
large, depressed and smooth in the centre, a small group of coarse
setiferous punctures near the apex and on each side at the base, the
apex acute. Elytra depressed, narrowed behind, the apex slightly
rounded with the sutural angles a little produced, a sinuous dorsal
carina on each from the base to the apical tubercle, and a short
outer one from the humeral tubercle to about the middle, these
carinz almost impunctate, the depressed portion between the dorsal
carina and the suture, behind the middle, longitudinally strigose,
the other parts with rather close coarse semi-circular punctures.
Pygidium strongly and somewhat concentrically strigose. Under-
side and legs coarsely punctured and strigose. Mesosternal process
conoid, the apex rather acute. Abdomen with a strongly punctured
central impression. Anterior tibie with two equidistant lateral
teeth, posterior tibie strongly channelled, the inner-side fringed with
long grey hair, the outer-side with a strong spinose tooth below the
middle.
The absence of any form of cephalic or thoracic horn in
the male at once separates this interesting species from
all others hitherto described. It is most nearly allied to
khasiana, Jordan, and microphyllus, Wood-Mason, but
besides the absence of horns it differs from both species
in many other points that I have given in the description.
The female is unknown to me.
Cephalocosmus moewist, Kraatz (Deuts. Ent. Zeit., 1895,
p. 107), from the description, is evidently identical with
M. microphyllus, Wood-Mason, this latter name having
fourteen years’ priority.
INGRISMA, Fairm.
Ann. Soc. Ent. Belge, 1893, p. 292.
This genus established only in a note, and omitted in
the Zool. Record (though included in the recent ‘ Index
Zoologicus’), has been generally overlooked; rasuta, Fairm.,
is the type and cupreola, Fairm., from the, description
would appear to be merely a colour variety of the female
of the same ‘species. 7. whiteheadi, Waterh., has been
since added, and besides the two additions I now make,
the Heterarrhina euryrrhina, Gestro, and Coryphocera
308 Mr. O. E. Janson on the genus
tonkinensis, Moser (Berl. Ent. Zeit., 1901, p. 525), must
also be referred to this genus, but judging from the
description the latter is apparently identical with rasuta,
Fairm., in which case the genus would now comprise five
species, all of which are known to me.
Ingrisma binghami, n. sp.
Cyanea vel viridis, nitida, coxis posticis marginibus pedibusque
rufis, tarsis rufo-piceis, elytris striato-punctatis.
d. clypeo elongato, medio constricto, apice valde dilatato, margine
antico medio late reflexo ; tibiis anticis tenuoribus, lateribus
inermibus. Long. 30 mm.
2. clypeo piceo, apice leviter dilatato, rotundato ; tibiis anticis
brevioribus, latioribus, lateribus apiceque valde dentatis.
Long. 26 mm.
Tenasserim.
Deep purple blue, or green with a reddish-golden tint in certain
lights, the sides of the head and thorax, the base of the pygidium
and the abdomen slightly rufescent, the hind margin and exposed
upper-side of the meta-coxee, sides of the abdomen and the legs red,
the femora tinged with blue or green, the knees, extreme apex of the
tibia and the tarsi reddish piceous ; in the female the anterior part
of the clypeus is also piceous. The whole of the upper surface and
the abdomen have a very fine and dense punctuation, which is only
discernible under a lens.
Head rugosely punctured, the base smooth, clypeus strongly
constricted about the middle and very much dilated in front, the
side margins strongly elevated and acute, strigose on the outer side,
the apex broadly reflexed with the entire apical margin strongly and
uniformly rounded. Thorax a little rounded at the sides and
narrowed from the base, the basal margin distinctly emarginate
above the scutellum and with the lateral angles rounded, finely and
sparsely punctured on the disk and at the base, more strongly
punctured towards the sides. Scutellum broad and _ triangular,
remotely punctured. Elytra broader than the thorax at the base,
the disk obsoletely bicostate and with rows of coarse semi-circular
punctures, five of which are regular and continuate, the others
being irregular, the sides more finely and irregularly punctured, and
strigose towards the apex, the sutural angles almost right angles.
Pygidium very finely and closely strigose, slightly impressed on
each side near the apex. Beneath coarsely strigose at the sides:
mesosternal process long, broad and flattened, obliquely divergent
and slightly curved, its apex rounded ; anterior tibiz slender and
Theodosia and other Eastern Goliathides. 309
strigose, the outer edge without any indication of teeth, intermediate
and posterior tibize fringed with golden hair, the latter with a slight
sub-median tooth.
The female is smaller than the male, and altogether more strongly
punctured, the clypeus is scarcely constricted and very much less
dilated, with the side margins less elevated, the apex much less
produced and more narrowly reflexed, the legs are shorter, the
anterior tibiz are very broad with a large obtuse sub-apical tooth
and a strongly produced apex.
Allied to ewryrrhina, Gestro, but larger than that species, and
with the mesosternal process broader and more strongly divergent,
the clypeus in both sexes is longer, more rugosely punctured, and
with the side margins more strongly raised, in the male it is also
much more dilated and broadly reflexed at the apex.
A pair of this fine species have been given to me by
Col. C. T. Bingham, who took them on the flowers of
Ficus religiosa in the Thaungyin Valley, Tenasserim, in
August 1894. The male is of a beautiful purple blue
colour and the female is green, but this difference is
without doubt an individual variation, such as is seen in
nearly all the green Heterorrhiny and in other genera of
this family.
Ingrisma femorata, n. sp.
dg. Elongata, convexa, viridis, nitida, coxis posticis extus tibiisque
rufis, tarsis piceis ; clypeo antice abrupte dilatato et medio profunde
bi-foveolato, margine antico arcuato, reflexo, angulis lateralibus sub-
acutis ; elytris fere levibus, lateribus transverse plicatis ; pedibus
anticis longioribus, femoribus dentatis, tibiis curvatis, intus tenuiter
serratis, Long. 28 mm.
Korat, Siam.
Bright bluish, green, very shining, the exposed margin of the
meta-coxee and the tibia red, femora green changing to reddish on
the upper parts in certain lights, knees, extreme apex of the tibi
and the tarsi reddish piceous.
Head slightly convex in the centre between the eyes, the entire
surface, except at the base, rather closely covered with minute black
tubercles which become rather larger, and elongate in form, towards
the front, clypeus nearly straight at the sides with the margin
scarcely elevated, strongly and abruptly dilated just before the apex,
and with two large deep fovez in front separated by a slightly raised
carina, the apical margin broadly reflexed and regularly arcuate
with the lateral angles sub-acute and elevated. Thorax very convex,
310 Theodosia and other Hastern Goliathides.
the sides regularly rounded and narrowed almost from the base,
basal margin almost straight with a slight emargination above the
scutellum and the lateral angles obtuse, the disk extremely finely
and sparsely punctured, more coarsely so towards the sides. Scutel-
lum large elongate-triangular, almost impunctate. Elytra without
any trace of coste, the surface a little uneven but impunctate with
the exception of an almost obsolete sutural row on the basal part,
the inclined lateral part transversely wrinkled in the middle and
coarsely strigose behind, the sutural angles distinctly produced and
obtuse. Pygidium coarsely and irregularly, but not closely, strigose.
Beneath remotely strigose at the sides, the mesosternum punctured,
and with some fine pale pubescence in front ; mesosternal process
long, obliquely divergent, narrow and slightly curved inwards
towards the apex, which is obtuse. Anterior legs long, the femora
very stout and furnished with two closely approximate teeth on the
under-side about one-third from the apex, the tibiz slender, curved
and finely rugulose, slightly dilated and finely serrate on the inner-
side from a little below the base, the outer side without any indica-
tion of teeth and the apex but shghtly produced ; intermediate and
posterior tibiz fringed with hight brown hair, the latter with a very
slight tooth a little below the middle.
The form of the clypeus, the very convex thorax
strongly deflexed in front, the almost impunctate elytra
and remarkable anterior legs are sufficient to at once
distinguish this species from 7. binghamt, to which it is
most nearly allied.
(Padtl =)
XVII. Leperiments in 1893, 1894, and 1896 upon the
colowr-relation between lepidopterous larve and
their surroundings, and especially the effect of
lichen-covered bark upon Odontopera bidentata,
Gastropacha quercifolia, etc. By Epwarp B.
PovuLton, D.Sc., M.A., LL.D. (Princeton), F.R.S.,
etc., Hope Professor of Zoology in the Univer-
sity of Oxford, Fellow of Jesus College, Oxford.
[Read June 3rd, 1903. |
Pirates XVI, XVII, anp XVIII.
THE circumstances under which the experiments re-
corded in the present memoir were undertaken, afford a
good example of the stimulus and encouragement to work
rendered possible by that mutual intercourse and exchange
of experience and ideas which are promoted by meetings
of scientific societies.
In the year 1892 I conducted an extensive series of
experiments upon the adjustment of the colours of the
larvee of Amphidasis betularva to those of their environ-
ment (Trans. Ent. Soc. Lond., 1892, pp. 337-369). Living
examples of the chief results obtained were shown at the
meeting of Section D of the British Association at Kdin-
burgh, on August 9th (Report of the 1892 Meeting, p.
786, where however the word “pups” is erroneously
printed instead of “larve”). After the larvz had been
exhibited, Dr. Stacey Wilson, of Birmingham, asked if I
had tried the effect of lichen-covered bark. Dr. Wilson
stated that he had once beaten the larva from a food-
plant with twigs covered by lichen, and that its appear-
ance was entirely different from that usually borne by
betularia. He looked upon it, in fact, as the larva of
some other species, and was only convinced by breeding
the moth (Trans. Ent. Soc. Lond., 1892, p. 360). The
idea of making use of an environment of lichen-covered
bark had not occurred to me, and I determined to try
TRANS. ENT. SOC. LOND. 1903.—PART III. (OCT.)
312 Professor E. B. Poulton on colowr-relation
the experiment on this and other suitable larvae on the
first opportunity.
I propose to state the results of these experiments
forthwith, referring to the plates which accompany the
paper. Hence it will be possible to gather the conclusions
by looking at the few first pages, while those who desire
to study the evidence in detail will find it recorded in the
later part of the memotr.
The first larva which appeared suitable for the purpose
was Odontopera bidentata, and Mr. G. T. Porritt very kindly
consented to look out for eggs. Mr. Porritt had himself
suggested to me that the larva would probably prove to
be especially suitable for the purpose of this enquiry, and
he wrote on May 9th, “the larva varies so very much in a
wild state according to its food, that I fancy it will form an
interesting subject for your experiments.” * On May 9th,
18938, he kindly sent me from Huddersfield a batch laid by a
single female, and, on June 13th, asecond consignment laid
by two females, from Sledmere on the Yorkshire Wolds.
The first set afforded the material of Experiments I to IX
(including VA) described in this paper. The second mixed
set formed the subject of Experiments X to XVIII.
The chief results of both experiments will be gathered by
a glance at Plate XVI, in which figs. 1 to 5 represent
larvee from the first set of eggs, figs. 6 to 11 larvee from the
second set. At the same time results like those shown in
the former figures were produced in larvee of the second set
and like those in the latter figures, in larvee of the first set.
The detailed account of the experiments shows the
number of days which elapsed before the influence of each
environment became visible, and the time which was
necessary in order to produce the full effect. A very large
number of records proves that the larvae, im the great
majority of cases, rested by day upon the object which
* Dr. T. A. Chapman wrote to me (June 14, 1903), concerning the
forms and habits of bidentata :—‘ About forty years ago, I took
three or four beautifully green latticed larve off the lichen-covered
trunk of an old alder-tree in Glen Messen (Argyleshire). They
were quite new to me, and though very like (of course) bidentata,
I thought they must be something else, a lichen-feeder. When they
produced bidentata, I got no further than wondering whether bidentata
was sometimes a lichen-feeder. I remember well their resting-place
was near the ground, many feet from any leaves,—as long a journey
for feeding as the larva of Aprilina makes,”
between lepidopterous larve and their surroundings. 313
they afterwards came to resemble. This, however, is pro-
bably not the case in the earliest stages, when the larvae
doubtless rest on the leaves and stalks.
The extreme sensitiveness of this larva is clearly shown
by a glance at Plate XVI. The four first figures indicate
a power of adjustment about equal to that of the most
sensitive larva hitherto known, Amphidasis betularia (com-
pare Trans. Ent. Soc. Lond., 1892, Plate XIV). The effect
of green leaves and shoots, shown in fig. 5, is however
very inferior to that produced upon betularia, which
becomes bright green in this environment. The effect of
green leaves alone upon /identata is the same as that
observed in many other larvee, Noctuze as well as Geometre,
viz. the reduction of the brown ground-colour to a very
pale tint which would be far-.less conspicuous than the
more ordinary appearance. The contrast between the
results of an effective environment of green, and nothing
but green, as shown in fig. 5, and of green scattered over a
brown background of bark, as shown in figs. 6 to 11, is very
striking, and suggests renewed experiments with an artificial
arrangement of combined colours. Another interesting fact,
suggesting the restriction of larval susceptibility to the
immediate surface upon which the resting periods are
passed, is the entire absence of any effects traceable to the
green leaves of the food-plant when present with the
other forms of environment employed in these experiments.
The complex nature of the result produced is well seen in
the oblique white lateral marks which are found in larve
with the green lichen-like patches (figs. 6, 9), and probably
assist in the general effect by breaking up the larval
surface. The green markings are developed in the vicinity
of and include the projecting ridges, ete——a fact which
is of obvious significance in promoting the resemblance
to small scattered masses of lichen. ‘The various kinds
of lichen made use of did not produce corresponding
effects.* Thus the green dorsal patches shown on the
larvee which had been exposed to orange lichen (figs. 8,
9) did not differ in any marked degree from those in
which a green lichen had been chosen. It is by no means
* My friend Professor 8. H. Vines, F.R.S., has very kindly given
me the probable names of the lichens made use of. They will be
found in the detailed account of the experiments, and in the
description of Plates XVI to XVIII. The names could not be given
with certainty because the specimens themselves had been lost.
314 Professor E. B. Poulton on colour-relation
unlikely, however, that under entirely normal conditions
special detailed adjustments of this kind may be brought
about. With regard to the sensitiveness to lichen, bidentata
appears to be as superior to betu/aria, as it is inferior to the
latter larva in sensitiveness to green leaves, so that the
two species may be considered about equal in the power
of colour adjustment. It is interesting to observe that
dark purplish-brown twigs with white spots, although
producing lighter larvee than those upon unspotted but
otherwise similar twigs (compare figs. 4 and 3), did not
lead to the appearance of white marks upon the larvae
(fig. 4).
Dr. Stacey Wilson’s experience led me to try the same
experiments with an environment of lichen in the case of
A, betularia, My friend Mr. Arthur Sidgwick kindly gave
me a small batch of eggs in the summer of 1893, and the
fourteen young larvee which hatched from them were sub-
jected, together with lidentata, to this form of environ-
ment, in Experiments XII to XV. It will be seen
however that eleven of the resulting larvee were yellowish-
green, two brownish-green, and one grey mottled with
brown.
The same experiments produced the larvze of bidentata
of which typical examples are represented in Plate XVI,
figs. 6—11. So far as any conclusion can be drawn from
these four small experiments, betu/aria does not seem to be
nearly so sensitive or so specialized to this form of environ-
ment as bidentata. At the same time lichen must have
been the cause of the etularia larvee, with one exception,
becoming green; for ordinary bark tends strongly to
the production of dark forms of this species, even in
the presence of a great preponderance of green leaves
(Trans. Ent. Soe. Lond. 1892, pp. 381, 332). It will be
of interest to repeat these experiments upon a much
larger scale, and to introduce the larvee immediately after
hatching ; but it does not appear to be probable that this
species often exhibits the kind of susceptibility to lichen
observed by Dr. Wilson; for (1) it is remarkably sensitive
to other surroundings almost throughout its life-history
(see pp. 318—320), and (2) the four small experiments,
conducted in 1893, do prove considerable sensitiveness to
lichen although they did not lead to the production of
lichen-like larvee.
The fortunate discovery of a company of young larvwe
between lepidopterous larve and their surroundings. 315
of Gastropacha quereifolia by Mr. W. Holland, on July
22nd, 1893, enabled me to experiment on this interesting
species, which is well known to present grey and lichen-
like forms. The company, evidently the product of a
single batch of eggs, was so numerous that I was able to
start four experiments with fifteen larvae im each, on
July 28th. All were fed on hawthorn, the food-plant on
which the larvee had been found. This in three cases was
intermixed with environments more or less harmonizing
with known varieties of the larva—the rough black-barked
twigs of the Turkish oak, bramble-stems of a rich reddish-
brown colour, and sticks bearmg an abundant growth of
lichen (probably Ramalina farinacea in all cases). In the
fourth case the larvee were as far as possible restricted to
the green leaves and youngest shoots of their food-plant.
It was, however, impossible altogether to exclude shoots
of greyish and reddish-brown shades, and these probably
produced some effect.
At first the young larve rested chiefly on the food-
plant, but soon preferred the bark of the older wood. The
change took place simultaneously in each of the three
sets containing dark bark and lichen, as will be seen by
a glance at the following summary of Mr, Holland’s careful
notes :—
| | I. | IL, IIL.
| etes Food- Black Food- | Lichen: | ood | Reddish-
| plant. bark plant. | oe yer plant. brown bark,
| Aue. Tae 3 i ie Cee ie tea
PSS Var 2 cee oa feet ow GS te {are A ee!
Aug. 5. \ i es ee 5 a *
| Aug. 9 3 ee 1 | 14 1 13 |
* 3 on muslin roof. + 1 missing,
There is no reason to suppose that these effects were
due to any gradual recovery from disturbance. The re-
cently hatched larvze were found at the tip of a young
shoot on July 22nd, and it is probable that by August 5th
the period had been reached when they begin to seek the
older wood for the diurnal rest. After August 9th only
single larvee were found except upon the environments
which had been provided, and it is probable that, under
316 Professor E. B. Poulton on colour-relation
entirely natural conditions, larvee of the same age would
never be found upon the leaves or green shoots.
It is of interest to note that the larvee never rested upon
the lichen itself, but upon the bark of the sticks between
the masses of lichen. This position is consistent with
the larval appearance, which is that of bark partially
grown over with lichen.
There can be little doubt that the larva is influenced by
the colours of the environment from the time at which it
first seeks the older wood, but a certain period is required
before the effects become visible. A very obvious adjust-
ment to the three forms of environment was recorded on
August 14th—so obvious indeed that the first trace of a
visible result might probably have been detected some few
days earlier. The adjustment continued to become more
complete right up to the beginning of hybernation. On
Aug. 31st it was noted that the effects of the three environ-
ments had greatly increased. On Sept. 21st a careful
comparison of all the larvze was made upon a white paper
background. It was then thought that the adjustment
was as complete as it was likely to be before hybernation,
and for many larve this conclusion was justified. In
others however the effects continued to deepen right on
into October, as will be seen in the complete account of
the experiments. The latest changes probably took place
after the larve had ceased to feed; indeed they had eaten
very little for some time previous to October 3rd. The
degree of cryptic adjustment to the three environments
which had been reached by the beginning of hybernation
can be seen by a glance at the upper part of Plate
XVII, where examples of all the types of colouring are
represented.
In arranging the larve for hybernation many of the
environments were shifted, in order to test the existence
of any larval susceptibility during this period; and, as no
effects were visible when the larvee were compared after
the winter, these same surroundings were continued in
each case, right up to the time when the nearly mature
larvee were sent to Lord Walsingham for preservation, in
May. ‘The results of various comparisons point to the
conclusion that the larve of G. quercifolia are not sus-
ceptible to the colours of the environment after the begin-
ning of hybernation. Thus Plate XVIII, fig. 1 represents
a nearly mature larva, of which the appearance before
between lepidopterous larve and their surroundings. 317
hybernation is shown in either fig. 4 or fig. 9 on Plate
XVII. The latter appearance was a response to an en-
vironment of lichen-covered sticks; but after October 16th
these were replaced by black-barked twigs, which it is seen
produced no effect at all. Again, Plate XVIII, fig. 2 repre-
sents a later stage of Plate XVII, fiz. 6. Here too the
resemblance between older and younger larve is very close,
although the former had been subjected to the same black
environment after October 16th. The negative result
of a transfer experiment in the opposite direction is seen
in Plate XVII, fig. 14, the representation of a larva which
had been exposed to lichen-covered sticks after October
3rd. All the larvee did not remain as uniform throughout
their life-history as these three. Thus Plate XVIIL fig.
3 represents a nearly mature larva of which the appear-
ance before hybernation is seen in Plate XVII, figs. 4
or 9. In this case the larva darkened considerably after
the winter, although its environment had not been shifted,
but consisted of lichen-covered sticks throughout.
It is probable that the power of adjustment to environ-
ment possessed in so marked a degree by this species is
specially directed to protection during hybernation, when
the food-plants are leafless, and when enemies are often
pressed by hunger. But it is doubtless also of importance
later on when the larva becomes so much larger and would
on this account be far more conspicuous. It is probable,
however, that the caterpillar does not wander from its
food-plant, and that complete adjustment to the old wood
before hybernation is an adequate defence in the following
spring and summer. If this be correct there would be no
advantage in a prolonged larval susceptibility.
The same relationship between susceptibility and the
particular needs of each species is seen in the effect of
an environment of green leaves and shoots upon G. que7-
cifolia, O. lidentata, and A. betularia, The first-named
probably invariably rests by day, except for a brief period
after leaving the egg, upon the older wood, and the power
of adjustment to leaves and young shoots, being altogether
useless to it, has never been acquired. The last-named,
with its remarkable range of food-plants, including many,
such as broom or rose, in which green shoots are a
prominent feature, is frequently in a position in which
a green colour would best conceal its nearly smooth and
cylindrical form; and we find that, as a matter of fact,
318 Professor E. B. Poulton on colowr-relation
it always responds in this way to an environment of the
kind described above. Sidentata doubtless occupies an
intermediate position between the other two species in
this respect. The occasions are probably rare, but not
altogether wanting, in which it is compelled to develop in
a green environment. We find that it has the power of
making some considerable approach towards such surround-
ings, but not of attaining any high degree of resemblance to
them. It is probably the case, however, that the tint which
it produces on green leaves and shoots is of great value
on a pale yellowish-brown bark, which may often form its
environment; and it may well be that it is something
in common between the light reflected from this and from
green leaves which explains the similarity in the effects
produced upon the larve.
Typical examples of all the forms of quercifolia larvae
produced in these experiments were shown alive at the
meeting of the Entomological Society of London on May
2nd, 1894, and also at the Soirée of the Royal Society in
the same month. A brief account of the exhibit is printed
in the Proc. Ent. Soc. Lond., 1894, p. xvi. It is also referred
to in Mr. C. G. Barrett’s “Lepidoptera of the British
Islands” (Lond., 1896, vol. 1, p. 45).
The nearly mature larve of quercifolia, forming the
subject of the experiments described in this memoir, were
in almost every case sent to Lord Walsingham, and, with
the exception of one which was spoilt, were kindly preserved
by him. The specimens are now to be seen in the Hope
Department, Oxford University Museum, and in the British
Museum of Natural History
The last series of experiments described in this paper
grew out of the surprising restriction of susceptibility to
the younger stages of G. quercifolia. The results naturally
suggested further experiments upon other species well
known to be highly sensitive, and I immediately fixed
upon Amphidasis betularia as the most suitable for the
purpose. The investigation was carried out entirely by
the present writer, in the laboratory at Wykeham House,
Oxford. The results are clearly shown in the accompany-
ing diagram and summary. The Roman figures represent
the corresponding stages of larval life. The shaded squares
indicate stages passed in a black environment, the unshaded,
stages passed in the green surroundings,
between lepidopterous larve and their surroundings. 319
Experi-
migritl 116 II Ill. IV. Vs VI. Colour of mature larve.
B 5 green, 1 intermediate.
7 i Se eae
Ba
Us i ren smoky-black.
WY GY jj Rs igek (1 dark oi
B2 , 7 J) 3 eee greyish patches).
_ yo // 4 black.
B
eA YY | 5 black (overspread with |
grey).
| 1 brownish-black. |
6 dark (with grey
; C | patches).
| Y | | 3 intermediate.
y WIM) | 3 dark (respectively
yj Y | overspread with grey,
Yy Uy | greenish on sides, and
A GY YY YY with pale yellowish spots).
7 1 green (with brown
| | /dorsal line and lateral
YY, WY | patches).
| B; 2 black. |
5) Yy Y 2 dark overspread with
A* | Y | yy grey. |
Z; Ye VU, 1 bright green.
\ | 2 green, sprinkled with |
7) | grey, 1 greenish inter-
| jj mediate.
| Ae | Uj
1 whitish, 2 blackish
with grey markings.
1 distinct but dull green
with brown broad dorsal
line and slight lateral traces.
A glance at the diagram and results proves conclusively
that there is no restriction of susceptibility to the younger
stages of this species.
Experiment B* shows the strong
influence of a black environment applied only to: the two
latest stages, while A! shows considerable effects traceable
to black which was present in the 2nd stage only. When
TRANS. ENT. SOC. LOND. 1903.—PART II. (oOcT.) 22
320 Professor E. B. Poulton on colowr-relation
we furthermore take into: account the more intense effects
which were produced as additional stages were exposed to
a dark environment, we may feel confident that every stage
except the Ist and the 5th or 6th, is sensitive. These
require further experimental testing.
A very interesting and unlooked-for effect was produced
in many of the transferred larvee, viz. an overspreading
ereyness or the appearance of grey patches. Thus, although
the effect of the earlier surroundings appeared at first
sight to be entirely obliterated, the larvae were nevertheless
unable to develop their full and characteristic response to
the later environment. Details will be found in the account
of the experiments. It only remains to point out that
experiment A! probably indicates that these larvae are
susceptible to the colours of the branches at a period when
they are at any rate chiefly to be found upon the leaves
and leaf-stalks, and that there was some evidence to show
that the influence of environment may be largely a question
of time, so that of several larvae passing the same stages in
given surroundings, those which grow most slowly are, on
the whole, the most affected.
Finst EXPERIMENTS WITH LARVZ OF
O. BIDENTATA (1893).
The larve from the first set of eggs sent by Mr. Porritt
from Yorkshire hatched at about the same time, so that
nearly all the experiments recorded on pages 322—325 in
a tabular form were started on the same day, May 22nd.
The observations were in part conducted by Mr. Holland and
in part by me, as is indicated by the initials or name under
the dates in the left-hand column of the table. Hence in
the account of each experiment there is the opportunity of
comparing two independent sets of observations.
‘he food-plant made use of in all these experiments was
the black poplar (Populus nigra).
A careful comparison of the results of the 10 sets of
experiments (viz. I to IX, including VA) was made by the
present writer on July 6th, 1893, all the larvee being placed
on a background of white. At this time all except one
were in the last stages, and many in all the cylinders were
approaching maturity. Of the forms of environment made
use of, 7 had produced dark larve, and 3 light.
between lepidopterous larvee and ther surrowndings. 321
A. DARK LARV OF BIDENTATA ON JULY OTH.
I. Black sticks Fourteen out of the 15 larvae were very
black, the exception being quite small and probably in the
3rd stage. These larvee were the darkest of all the sets.
II. Deep blue paper spills—These 12 larvee were very
dark, coming next to those upon the black twigs in this
respect. The dark purplish-black colour was also very
uniform over the whole larval surface corresponding to the
unvarying tints of the environment.
I. Purplish-brown twigs.—These 15 larvee were slightly
more variegated than II, in correspondence with the less
uniform darkness of their environment. Except for this
slight introduction of rather lighter shades these larve
were as dark as II.
IV. White spotted purplish-brown twigs.—These 16 larvee
were dark, but distinctly lighter than those of the three
previous sets. They also varied a little, whereas the
groups first described were more uniform. Although the
relative lightness and the darkness of these larve, as
compared with III, corresponded to the general effect of
their respective environments, there was no marked
resemblance to the special details of the twigs which
had been carefully selected for Experiment IV.
V. Brown twigs—The 17 larvee were distinctly brown
and not nearly so black or purplish as the preceding sets.
The shade of brown varied, being much hghter in the
smaller larvee. The brown larval surface was also some-
what variegated with different shades of the same colour.
Va. Bark overspread with a bright yellowish-green powdery
lichen—The 18 larve varied very greatly, some being as
dark as the blackest of set 1; many were variegated with
shades of brown, harmonizing well with the environment ;
for the lichen soon lost its green tint and became various
shades of brown. This is the only note relating to this
experiment which has been found. There is no doubt that
the young larvee were introduced on May 22nd or 28rd.
VI. Lichen-covered sticks—The lichen was probably dead
and. had become much paler. These larvze too were much
lighter ‘than any of the previous sets, although a few
were quite dark. Nearly all were light brown and much
variegated with shades of brown, harmonizing well with
the environment.
Of the 7 sets of larvae described above, the degrees of
Professor E. B. Poulton on colowr-relation
|
| : ae | Dal paorplheb -
t 1393 Blick iica ot D P blue > aper | Dark purplish-brown White- spotted
ae Terk otk, | Pap 7) ey Sige pote meine ee
_ — - | —
May 22 15 Young larve, (May 23.) 15 15 Jlarve in-| 16 young larvee
| (E. B. P.) | introduced |young larve in- troduced (just introduced.
| troduced. hatched).
June 11. (tne f Re- fed. | 12 larve counted; (June 16). All 15 counted.
(EB .P;)) 7) eee very 15 dark like the |
| dark. twigs, but not |
| markedly purplish. |
a | | = = = ae} =
June 19 | (June 17.) All; All 12. very All 15 on twigs, 12 larve on
(E. B. P.) | 15 very black Hs) dark, very dark and/ twigs, 4 on green.
‘the twigs. purplish. | All dark, but
/greyer than those
/in III, and not so
| dark as latter.
June 25. | (June 22. ) As| As before. All |
(E. B. P.) | before. All at rest | on spills, |
(on twigs.
July 1. | As before: all | (June 27.) All (June 27.) Re-fed.
(E. B. P.) | on twigs. (15 dark brown, |
|
| | |
July 6. 2 at rest on| 9on aig, lon) All15 ontwigs. 13 on twigs, 3.
Careful com- | green, 13 on twigs. | green, 2 not noted. | All very dark. on green. Dark,
parison on = All very black. — | All 12 of a uniform | but much lighter
white paper. | ‘dark purplish- | and less constant
(EB E)) | black. | than III.
ae ere eee at ij | |
| July 10. 2 at rest on! Sonspills,4don| 14 on twigs, 1 | 15 on twigs, 1 |
| (W. Holland. )
July 12.
(W. Holland. )
green, 13 on twigs.
| All very black.
15 on twigs.
/green, All very|ongreen. Alldark on green. The,
dark. purplish-brown. larve remain
lighter in tint
‘than 1a,
9 on spills, 30n| 13 on twigs, 2 |
green. ; on green.
/on leaves.
|
13 on twigs, 3
between lepidopterous larve and their surroundings.
323
V.
Reddish-brown
twigs.
17 young larve
introduced,
|
Vile
Lichen-covered
sticks,
|
| 14 young larve
introduced.
AUT
| Weathered pale grey
barkless twigs.
VIII.
Green leaves and
shoots of food-plant
(Populus nigra).
|
|
ix |
Orange paper
spills.
(June 3.) 15) 15 young larve
young larve in-/| introduced.
troduced,
15 young larvee
introduced.
__ All 17 evidently
_ becoming
like the twigs.
brown |
| iN
| (June 9.) Re-fed.
Apparently be-
coming light
/brownish and ap-
proaching colour
of twigs.
Brown like the
twigs.
As before. 14
on twigs,
green, 1 uncertain,
2 on!
14 larvee ; vari-
able, with no
marked resem-
blance to lichen.
iG}
brown,
on twigs, 1 on
muslin roof, 1 on
green. All closely
resemble twigs.
All 15 very | |
pale brown
(June 17.) All
very
pale
some
faintly greenish.
6 on spills, 1
on muslin root,
8 on green. All
very light brown.
on green. Some
distinctly varie-
gated light and
dark brown, but
no green marks
yet.
10 on sticks, 4
(June 22.) 10
on twigs, 1 on
muslin, 3 on green,
before.
(June 22). As
14 on spills, 1
on green. Colour
as before.
a
|__ All 17 on twigs.
They become much
‘lighter when
changing skin.
9 on sticks, 5
on green. Larvee
still very varie-
gated.
13 on twigs, 1
on green.
17 on twigs.
Distinctly brown,
and not purplish
like III and IV.
12 on sticks, 2
on green. <A few
dark but mostly
light and _ varie-
gated with shades
of brown.
All 14 light grey
and harmonizing | very
perfectly with
twigs. | greenish,
16 on twigs, 1
on green. All
brown as before.
9
12 on sticks, 2
on green. Two
larvee variegated
with dark green:
the rest as before,
| but darker brown.
on green,
16 on twigs, 1
13 on sticks, 1
on green, 4 larvee
with dark green
patches, strongly
developed in 2 of
them.
10 on twigs, 4
on green. 1 larva
injured. Appear-
ance as before,
11 on twigs, =
on green.
14 larve. All
light
brown, but not
All 15 on spills.
Larve resemble |
VIII, only not,
quite so light.
11 on spills, 4 |
on green. Very
light yellowish-
brown.
12 on spills, 3
on green,
324 Professor E. B. Poulton on colowr-relation
| | 4 | 11. IL. AY:
Dates in 1893. Black twigs of | Deep blue paper eee Pp urplishs Tew wi hite-spotted :
Tuyleishi ale spills. glossy twigs probably purplish-brown twigs
| of birch. of birch. |
| ; | :
| July 15. | 15 on twigs. All| 10 on spills, 2} 14 on twigs, 1] 12 on twigs, 4
“Ow. Holland.) | very black.
on green.
dark.
All very
on green. All very
dark purplish-
brown.
on green, Colour
darkish brown but
| distinctly paler
i: than IIT.
As before.
July 18. As before.
(W. Holland.)
—| |
July 23. | As before.
_(W. Holland.)
— ee = =
Some mature
and spinning up.
July 27. Some pupating.
6 larvee remain,
As before.
As before.
As before,
Allavery uniform
brown, almost
black, but not so
much so as those
with the
spills.
all on — twigs.
(July 30.) A
typical larva
painted ; shown in
Plate XVI, fig. 3.
blue
9 still feeding : |
As before.
|
All dark greyish-
brown and very.
uniform.
12 still feeding :
all on twigs. |
(July 30.) AY
typical larva}
painted ; shown in
Plate XVI, ze: 4,
Careful ecom- | All black, but one nearly mature. All| dark purplish-
| parison ‘has very faint | very dark purplish,
_ (KE. B. P.) | traces of green on almost black, as
| 'sides anteriorly, on many previous
| These larve_ occasions.
lar ‘blacker and_ less
purple than those
| ‘with blue — spills
| (II).
| |-
|
July 28. | 8 Jlarve still 5 still feeding.
|(W. Holland.) feeding. All on 4 on spills, 1 on
| twigs. leaf,
Aug. 3. (July 31.) A
|
| (W. Holland.) |
|typical larva|
painted ; shown in
| Plate XVI, fig. 1.
(Aug. 4.) Re-fed. |
| (Aug. 8, | Re-fed.. One or
two still feeding.
|W. Holland.) |
5 still feeding.
on twigs.
\4 on twigs, 1 on
| green.
|
As before. All 5 |
6 feeding. All
on twigs.
5 feeding: all,
'on twigs. Colour)
las before. (Aug.
/19.) 8 still feed- |
ing g.
between lepidopterous larve and their surroundings.
Ve
VI.
Reddish-brown
twigs.
Lichen-covered
sticks.
10 on twigs, 7 on
VIL.
Weathered pale grey
barkless twigs.
VIII. |
Green leaves and |
shoots of food-plant|
(Populus nigra).
IX.
Orange paper
spills.
10 on sticks, 4
12 on twigs, 1
11 on spills, 4
very pale brown, |
| 4 feeding on |
| Aug. 26, “and 1|
| or more on Aug. |
oe
green. All brown | on green, 6 with | on green, All very | on green, All
like twigs. ereen patches, pale and extremely | | pale yellowish-
‘very marked in 2. | like twigs. | brown. |
| ne
As before. As before. Some | As before. As before.
larvee mature.
Some laryw]| Others mature. As before, | Some larvee |
mature. mature,
|
i | |
Some larve have! 3 or 4 with, All now much) Larve very! Very light.
become of a much | ereen patches, 3 darker, but still|light brown, brown, like VIII.
darker brown or 4 with brown. like the darker) like IX.
lately. The change twigs in the!
probably due to |eylinder. Some | |
maturity. mature. |
|
- ae aie ieee | ee
10 still feeding. 7 feeding. All| 10 feeding ; all | (July 31.) Aj; 7 feeding, all}
8 on twigs, 2 on on sticks. ontwigs. A typical} typical larva on spills. |
green, larva was painted ;| painted ; shown
shown on Plate lon Plate XVI,
| | XVI, fig. 2. | fig. 5.
a p Mth Bt [eae Asa is eee
4 feeding. All. 2 feeding. | 6ontwigs,lon) 6 feeding. All 5 feeding. 4 |
on twigs. | green. | pale brown. on spills, 1 on.
| muslin.
(Aug. 19.) 1 (Aug. 18.) All| 7 still feeding. hoe feeding, 1| As before. All.
still feeding. _pupated, |(Aug. 19.) 2 still | dead. (Aug. 19.) 2 aa spills. (Aug.
| | feeding. '5 feeding ; ait 0.) All pupated.
|
|
326 Professor E. B. Poulton on colowr-relation
darkness follow in the same order as that in which the
results are recorded, except that V and VA were much
alike, the brown differing in tint rather than in depth.
VI were much lighter than any of the others.
B. LIGHT LARVE OF BIDENTATA ON JULY OTH.
VII. Weathered grey barkless twigs—The larve were
very light; of a distinct grey colour, harmonizing perfectly
with the environment.
VIII. Green leaves and shoots of food-plant—The 14
larvee were all very light brown but not at all greenish.
IX. Orange paper spills—The 15 larve closely re-
sembled VIII, but were not quite so light.
Some of the results of this comparison have been
incorporated in the tabular statement. Another careful
comparison was made on July 27th, but in this case it
was possible to include the whole in the table.
SECOND EXPERIMENTS WITH LARV OF O. BIDENTATA.
EXPERIMENTS WITH LICHEN ON LARV OF
A. BETULARIA (18938).
In sending the second mixed set of eggs of bidentata,
Mr. Porritt wrote on June 13th, 1893—“TI certainly had
no expectation of seeing any more Odontopera bidentata
this season. However, when collecting on Saturday, at
Sledmere, in a high wood on the Yorkshire Wolds, I found
several! Two of the females have deposited a few eggs,
which I forward at once with this. Sledmere is on the
chalk, and bidentatw there is quite of a different type to
our West Riding moth, being of the pale, ochreous banded,
distinctly southern form.”
The eggs were placed in a single cylinder, and as soon
as a sufficient number of larvae had hatched, I started
Experiments X to XV, between June 27th and July 3rd,
the 14 larvie of A. betularia received from Mr. Arthur
Sidgwick being divided between Experiments XII to XV.
Experiments XVI to XVIII were started on August 3rd
by Mr. Holland, with the latest larvae of bidentata. All
the observations on Experiments X to XVIII recorded
in the tabular statement on pages 328—331 were made by
Mr. Holland. The food-plant used throughout was Populus
nigra. When flat pieces of bark were introduced (XII,
between lepidopterous larve and ther surroundings. 327
XIII, XIV, as well as VA in the first series of experiments),
they were tied together in pairs with the lichen-covered
surfaces outwards.
EXPERIMENTS XVI, XVII, and XVIII with THE LARVzA
OF O. BIDENTATA UPON GREEN LEAVES AND SHOOTS
OF THE FooD-PLANT (Populus nigra).
The results of these experiments were uniform, and
require so little description that a tabular form of
presentation is unnecessary.
Experiments XVI (10 young larvee), XVII (17 larvee),
and X VITI (14 larvae) were begun by Mr. Holland on August
3rd, 1893. He recorded that the larvee of XVI were pale
brown, 2 of them rather variegated; while of XVII, 14
were pale brown and 3 rather variegated. Of XVIII no
record was made. The larvze were the last hatched from
the mixed batch of eggs which supplied the material for
Experiments X to XV.
August 8th. All re-fed. Sixteen larvee in XVII.
August 12th. XVI and XVIII re-fed ; the latter noted
as nearly all pale brown, a few rather variegated.
August 15th and 19th. XVI and XVIII re-fed; larve
10 and 14 respectively.
August 17th. XVII re-fed; 16 larve.
August 21st. XVII re-fed; 8 larve had escaped. The
10 larvee in XVI were all of a pale brown colour.
August 22nd. _ XVIII, 13 larve pale brown, 1 darker
with a few green markings.
August 26th and 31st. All re-fed; larvee 10, 8, and 14
respectively.
September 5th. XVI, becoming mature (2 still feeding
on 9th and 13th ; no further notes on this set). XVI, 7
larvee, all pale brown. XVIII, 14 larve.
September 8th. XVII, 2 mature (all mature on 13th).
XVIII, 1 mature, 1 dead. Of 12 still feeding, 10 pale
brown, 2 rather darker brown. ‘T'wo out of the 12 slightly
tinged with green.
September 13th. XVIII, 6 still feeding. On the
18th they were neither pupating nor feeding, and on the
23rd the last died. It appears possible that there was
a tendency towards hybernation on the part of the larvae
with the slowest rate of growth.
328 Professor E. B. Poulton on colowr-relation
XI.
X. | XI. Bark covered with
Dates in Dark purplish-brown | Reddishebrown bright yellowish-green
1893. glossy twigs, | twigs. powdery lichen, the colour
probably of birch. of which faded and left
the dark bark,
(July 2.) 13 small) (July 3.) 20 larve| 3 betwlaria, smaller
| larvee introduced, introduced, at begin-|than in XIII and
. |mostly in 2nd stage, ning of second stage.|XV, introduced. 15
June 27. | The smallest about} bidentata, similar to
5‘5 mm. long, and/those introduced in
| none much longer, XIII.
| |
> jinw A. a SSS es = Sa EE aoa!
Half of the larve| 20 counted. Afew| 3. betularia, pale
_becoming purplish. | becoming reddish- | yellowish - green. 15
July 12. | _ brown. bidentata variegated. |
| |
| |
| |
July 15. Re-fed. | Re-fed. 20 counted. Re-fed. 3 and 15!
counted, —
eee SUE FS aes |e ck
| 9 on twigs, 4 on 10 on twigs, 8 on) 2 betwlaria, mature. |
July 18. green. -green. Larvie becom-|14 bidentata, all |
ing dark reddish-| darkish, variegated.
brown and variegated.
| (July 24.) 12 on | 15 on twigs, 1 on| Last betularia
‘twigs, 1 on green. | green, 1 on muslin. | mature. 10 bédentata ,
| Almost all dark, | As before. on bark, 1 on green, 3
July 23. | on muslin. All varie-
July 28. | variegated; 3 paler on muslin. 8 changed
|
12 on twigs, 1 on
green. 10 dark brown, |
and less variegated,
|
gated brown, slightly |
touched with green |
and with white dashes
(July 27.) 12 on|
twigs, 4 on green, 1
skin and of a more
uniform brown colour.
along sides.
13, all on bark (as
also on Aug. 1). About
half darker, but all
still variegated. All
except 2 with more |
or less green. |
Aug, 3.
|
| i11-.on twigs, 2 on
green. 11 dark brown
‘and more uniform.
than before, 2 rather
|paler. One or two
have patches of pale
| green,
(July 31.) 14 on|
twigs, 1 on green.
8 brown like the
twigs; 4 paler and
Jess uniform ; 3 varie-
gated and patched
with green. (Aug. 4.)
12 on bark, 1 on
muslin. As before,
but have become
darker. This is the
darkest set of Experi-
ments XII to XV.
12 on twigs, 2 on
green, 1 on muslin.
'As before. |
\
eh ea i
Se a ee
between leprdopterous larve and their surroundings.
XIII.
Bark covered with
bluish-green lichen, probably
Physcia pulverulenta.
XIV.
Bark covered with orange
lichen, probably Physcia
parictina, combined with
P. pulverulenta.
XV.
Lichen-covered sticks.
The lichen probably
Ramalina farinacea.
3 betularia, about 12)
mm. long, introduced.
12 bidentata, about 6)
mm. long, introduced,
mostly changing skin.
4 betularia and 10
about 7 mm. long, intro- | bidentata introduced.
duced. 1 changing first; Both similar to those in
or second skin, others | XIII.
rather larger. 8 bidentata, |
‘about 6 mm. long, in-|
troduced. |
a | | ——~ = ll ——— —————— fa
3 betularia pale, 3 betularia yellowish-
2 betularia yellowish- |
| green, 1 brownish-green.
mn
(June 25.) 4 betwlaria,
yellowish-green, 1 grey FE green, 1 dark brownish-
110 = bidentata much | mottled with brown. 2. green. Bidentata
‘variegated, much smaller than others. | mottled.
| Bidentata rather pale, |
yariegated. | |
2 Sy
Re-fed. 3 and 10; 2 yellowish - green) 4 and 10 counted,
counted. | betudars ia still feeding. |
| Betularia as before.
Betularia, all mature. | <All betularia mature.| 3 betwlaria mature, |
9 bidentata much | 4 bidentate on bark, 3,1 escaped. 10 bidentata
variegated. on green. All brightly. “dark, mottled; some
variegated. green on sides. |
8 on bark, 1 on green. | 4 on bark, 3 on green. (July 24.) 8 on sticks, |
2 seemed unhealthy, All |
much variegated
pale brown ‘and green.
Small white dashes on |
sides.
7 on bark, 1 on green. |
Colours as before. Typical
larve painted, Aug. Ist
-and 3rd, shown in ” Plate |
| XVI, figs. 6 and 7.
8 on bark. 1 unhealthy,
6 brown, lighter than in
XII, brightly mottled
‘with paler brown and
green. 1 paler, less varie-
‘gated, with little green.
_1 small and pale, without
green. White lateral
| dashes only on former 6.
with | with shades of brown,
All brightly variegated} 1 on green, 1 on muslin. |
“Nearly all patched a
and patched with green. "green.
|
|
6 on bark, 1 on green.! 10onsticks. 6 darkish |
As before. brown mottled with green |
and paler brown: white |
dashes on __ sides. 3 |
\similar but paler. 1
eseoue green.
7onbark. All Siena 7 oo more or less_
skin. 1 blackish-brown, | patched with green, some,
4darkish brown, 2lighter| very strongly. 1 grey,
brown; all variegated | and brown strongly |
with much green, and the | patched with pale green. |
darker ones also with 2 pale brown with little
paler brown. All with} green. Most show white
lateral white dashes, ‘lateral dashes.
330 Professor E. B. Poulton on colour-velation
| XIL.
| DG XI Bark covered with
| Datesin Dark purplish-brown cea Ss bright yellowish-green
1893. glossy twigs, Boden ie own powdery lichen, the colour
ES of which faded and left
| probably of birch.
the dark bark.
9 on twigs, 4 on (Aug. 7.) 14 on 10 on bark, 3 on
green. All dark brown, twigs, 1 on green. green. 10 of a rich
some with a little dark brown patched
green. | with lighter brown
and green: 3 lighter
Broree brown variegated with
aa | still paler brown and |
'green. All but one of
_ these paler larve with |
little white dashes on |
sides. The green marks |
| less pronounced than |
}in XIII and XIV.
|
| (Aug. 10.) 11 on} (Aug. 10.) 13 on 10 on bark, 2 on |
| | twigs, 2 on green. | twigs, 2 on green. | green, 1 on muslin, |
/11 purplish- brown, 2 All brown, mostly
Aug. 12. | rather paler. Several | rather darker than
| larve with green twigs, but 4 rather
'patches on sides. ‘paler. Several patched
with green,
12 on twigs, 1 on| 12 on twigs, 3 on| - (Aug. 14.) 10 on.
green. As before. green. As before. bark, 2 on green, ol
but 2very dark brown,
fo)
| Aug. 15. with far less green
! than XIII and XIV.
10 on twigs, 2 on 10 on twigs, 5 on! 12 larvie, as before.
green. Purplish- | green. All _ brown,
Aug. 19. brow n, several with a several patched w ith
little green. 2 rather green.
paler brown.
8 on twigs, 4 on| (Aug. 22.) 12 on (Aug? 9222) 29) ton
green. 10 purplish-| twigs, 3 on green. | bark, 3 on green.
brown, 2 brown. Only | All brown, 8 resemb-
Aug. 21. | 1 or 2 slightly marked | ling twigs, 2 rather
with green. dark ker, 2 rather
‘lighter.
Rani 256 (Aug. 25.) 12 on| Re-fed. 10 on bark, 2 dead.
eam (Stleks:
9 on sticks, 2 on| 2 mature, and 2|} 6 on bark, 4 on
green. 1 mature. more on Aug. 31. green. The colour of
Aug. 30. the lichen had faded,
and left the dark
bark.
Mostly mature. 2 | (Sept. 4.) Nearly} (Sept. 4.) Larvee |
feeding Sept. 8. All|all mature. Only 1| becoming mature. A
Sept. 5. | mature Sept. 18. feeding Sept. 9. few oy feeding Sept.
9 and 13.
between lepidopterous larve and their surroundings. 331
XIV.
Bark covered with orange
lichen, probably Physcia
parietina, combined with
P. pulverulenta.
XII.
Bark covered with
| bluish-green lichen, probably
| Physcia pulverulenta.
Lichen-covered sticks.
The lichen probably
Ramalina farinacea.
|
| Bon bark, 2 on green.| 7 on bark. As before.| 10 on sticks. As”
As before. Typical larvee painted | before. |
Aug. 4 and 5, shown in
}
|
Plate XVI, figs. 8 and 9.
| (Aug. 14.) 7 on bark. (Aug. 10.) A typical
As before, except that larva painted, shown in
‘smallest larva now marked | Plate XVI, fig. 10. |
with green.
(Aug. 14.) 7 on bark. (Aug. 14.) 10 onsticks.
Lighter and greyer|8 grey-brown mottled
ground - colour. Green| with darker brown and
patches very marked. much green, so that some |
|are half green ; 2 similar
_ but browner.
7 on bark. Brown, 5 on bark, 2 on green. 10 on sticks. Colour
variegated with paler| Colour as before. _as before.
brown and with green
(strongly 5, less strongly
2).
6 on bark, 1 on green. 6 on bark. Allvarie-| 10 on _ sticks. As|
Colour as before. gated greyish - brown, | before. |
strongly patched with
green. White lateral |
dashes. |
| (Aug. 25.) 60nbark,| (Aug. 25.) 6 on bark. (Aug. 25.) 9 on twigs,
1 on green. 1 on floor. |
2 mature. <A typical} 6 on bark. Colour as, Larve becoming mature. |
‘larva painted Aug. 31, | on Aug. 21. |
shown in Plate XVI,
fot Tl. |
aa pis
(Sept. 4.) All mature. All mature except one. | All mature.
|
Last mature Sept. 8. |
332 Professor E. B. Poulton on colour-relation
EXPERIMENTS UPON THE LARVA OF GASTROPACHA
QUERCIFOLIA (1893-4).
On July 22nd, 1893, Mr. W. Holland found a company
of young larvee of this species crowded together towards
the end of a shoot of hawthorn near Steeple Aston, Oxford-
shire. They were all the same size, and had evidently not
long before hatched from a single batch of eggs.. Thus was
afforded the opportunity of tryimg another set of experi-
ments upon a larva which is known sometimes to assume
a lichen-like appearance. Furthermore, there was the
additional interest of testing larval susceptibility to en-
vironment before, during, and after hybernation. The
food-plant employed throughout was hawthorn. The
present writer is responsible for the starting of the experi-
ments, the arrangements for hybernation,and the comparison
of larvze on September 21st, October 3rd to November 3rd,
March 26th, April 27th, May 7th and 25th. The other
records were chiefly made by Mr. W. Holland, in much less
part by the present writer.
LG. queresfolia.
July 28th, 1893. Fifteen larvae of quercifolia placed
with intensely black twigs of the Turkish oak (Quercus
cervis), intermixed with their food-plant, hawthorn. They
were re-fed and examined on the following dates.
August 1st. Twelve larve on the green leaves and
shoots, 3 on the black twigs. All blackish-grey in colour.
August 5th. Four larvze on the leaves and shoots, 11 on
the black twigs.
August 9th. Three larve on the leaves and shoots, 12
on the black twigs.
August 14th. Fifteen larvee counted; all on the black
twigs. The colours were black and white with a little
grey.
August 19th. All 15 larvee on black twigs.
August 22nd. All 15 larve on black twigs.
August 31st, All 15 larve on black twigs. The larvee
were now much blacker, but white markings were still ~
present.
September 4th. Fifteen counted.
September 7th. Fifteen larvee on black twigs.
September 9th. Fifteen larvee on black twigs.
between leprdopterous larve and their surroundings, 333
September 14th. Fifteen larvee on black twigs, 2
blackish, 18 black with white markings.
September 16th. Fifteen larvee on black twigs.
September 18th. Fifteen counted.
September 21st. Fourteen larvee on twigs, and 1 little
one on leaf. The larva were re-fed on September 26th
and 30th.
IL. G. queretfolia.
July 28th, 1893. Fifteen larvee of quercifolia placed with
sticks profusely covered with lichen (probably Ramalina
farinacea was employed thoughout) intermixed with their
food-plant, hawthorn. They were re-fed and examined on
the following dates.
August 1st. One larva on lichen-covered stick ; all the
others on leaves and leaf-stalks. All blackish-grey in colour.
August 5th. Twelve larve on lichen-covered sticks ; 3
on food-plant.
August 9th. Fourteen larve on lichen-covered sticks ;
1 on food-plant.
August 14th. Fourteen larvee on lichen-covered sticks,
Colours black with white and grey markings; the latter
larger than on the larvee of the other experiments.
August 19th. Thirteen larve on sticks; 1 on food-
plant.
August 22nd, Thirteen larvee on sticks.
August 26th. Thirteen larvee on sticks.
August 31st. Thirteen larvee on sticks. The white
and erey markings had now developed to a much greater
extent.
Septenber 4th, Thirteen larvee on sticks.
September 9th. Thirteen larvee on sticks.
September 14th. Twelve larvae on sticks Colours
white and grey and black, except in the case of 2 larvae
without any black.
September 16th. Twelve larvee counted.
September 18th. Twelve larvee on sticks.
September 21st. Nine larvee on sticks; 3 on the twigs
of hawthorn. At this time of the year it was almost
impossible to get a sufficient quantity of the green shoots
of the hawthorn, so that darker twigs were sometimes
included, and upon these the larvae occasionally rested.
The larvae never rested upon the lichen itself, but on the
surface of the bark between the masses.
334 Professor E. B. Poulton on colowr-relation
The 12 larvze were re-fed on September 26th and 30th,
and on October 3rd and 7th.
IIL. G. quercifolia.
July 28th, 1893. Fifteen of the quercifolia placed with
red-brown stems of the bramble, intermixed with their
food-plant, hawthorn. They were re-fed and examined
on the following dates.
August 1st. Only 1 resting on bramble stem, others on
leaf-stalks. Blackish-grey in colour.
August 5th. Seven on stems, 5 on green parts of food-
plant, 3 on gauze top of cylinder.
August 9th. Thirteen on stems, 1 on green parts of
food-plant, 1 missing. .
August 14th. Fourteen on stems. Some black with
white markings, some grey and some mottled reddish-
brown.
August 19th. Fourteen on stems.
August 22nd. One missing, all on stems.
August 26th. Thirteen counted.
August 31st. Thirteen on stems. All were now grey-
brown with black and white markings.
September 4th. Thirteen on stems.
September 7th. Thirteen on stems.
September 9th. Thirteen on stems.
September 14th. The colours were now as follows :—
1 brown and grey with white markings, 2 brown with white
markings, 3 brown with various shades of grey, 3 grey and
brown with white markings, 2 grey with white markings
and brown patches, 2 dark grey with white markings.
September 16th. Thirteen counted.
September 18th. Thirteen on stems.
September 21st. Twelve on stems; 1 on hawthorn
twig.
They were also re-fed on September 26th, 30th, October
3rd and 7th.
IV. G. queresfolia.
July 28th, 1893. Fifteen larvee of guercifolia placed
with green leaves and shoots of the hawthorn. They were
re-fed and examined on the following dates.
August 4th. Fifteen counted. Blackish-grey in colour.
August 8th. Fifteen counted.
between lepidopterous larve and their surroundings. 335
August 14th. Fifteen counted. Rather variable in
colour, but somewhat greyer than at first.
August 19th. Fifteen counted.
August 22nd, Fifteen counted.
August 26th. Fourteen counted.
September 1st. Fourteen counted.
September 5th. Fourteen counted.
September 9th. Fourteen counted. -
September 13th. Fourteen counted. Colour variable:
3 black and white; 9 grey and black-and-white ; 2 grey
and brown-and-white.
September 16th. Fourteen counted.
September 18th. Fourteen counted.
September 21st. Fourteen counted. All on the shoots
of the hawthorn.
September 26th. Fourteen counted.
October 37d. Fourteen counted.
October 7th. Fourteen counted.
October 18th. Fourteen counted. Colour unchanged,
but rather darker than when last noted. Arranged for
hybernation in two muslin bags on the hawthorn tree
described on p. 337; one bag containing the 7 darkest, the
other the 7 lightest.
First GENERAL COMPARISON (SEPTEMBER 21ST), ALL
THE QUERCIFOLIA LARV# BEING PLACED UPON A
UNIFORM BACKGROUND OF WHITE PAPER.
I. G. quereifolia.
As regards the larvee with black twigs of Turkish oak, it
should be noted that the moisture in the glass cylinders
had encouraged the growth of small whitish spots of mould
upon the dark bark, and it is possible that some etfect may
have been produced by their presence in the environment.
Of the 15 larve, 6 were remarkable for the very slight
development of light markings upon the intense black
ground-colour: in one larva mdeed the light markings
were altogether wanting. The remaining 9 larve were
black, chequered with white markings, which were however
far less developed than in the former group which had
been with lichen-covered sticks. The ground-colour of
these 15 larve differed strongly from that of all the others
in its deep black shade.
_ TRANS. ENT. SOC. LOND. 1903.—PART II. (OCT.) 23
336 Professor E. B. Poulton on colour-relation
Two of the 6 caterpillars described above, including the
uniformly dark one, were selected for painting, together
with a representative of the 9 chequered larvee. The larva
without white markings was painted by Mr. Bayzand on
September 23rd, and is shown on Plate XVII, fig. 2; the
one with minute white markings on September 26th
(Plate XVII, fig. 3), and the chequered larva on September
25th (Plate XVII, fig. 1).
Il. G. querectfolia.
By far the most marked result was seen in the 12 larvee
with an environment of lichen-covered sticks. Not one of
the 12 could be mistaken for any larva of the other
groups. In the case of 4 larvee the effects were especially
marked, these being of a light colour over nearly the whole
of the exposed surface, the black ground-colour quite
replaced by a pale brownish tint in one and by grey in the
others. The 8 remaining larvee closely resembled each
other, being dark chequered with white; but the ground-
colour, although dark, was not black like that of the larvae
with Turkish oak (I), while the white markings possessed
a greyish tinge in one larva and a bluish in another.
The 4 larvee first mentioned were separated from the
others for painting at this date, together with the one in
which the markings were bluish-white, and another repre-
sentative of the commonest type of appearance, viz. a dark
ground-colour with white markings.
IL. G. quereifolia.
The ground-colour of all the larvee with brown bramble-
stems was very dusky and brownish. Five of them had a
distinct brown patch on each side of the 2nd abdominal
segment, partially extending on to the Ist, and another
patch on the dorsal surface of the 5th abdominal. The
dorsal hump on the 8th abdominal, and in some larvee the
area surrounding its base, were also brown. In the remain-
ing larve the light markings were more generally greyish
than white, and were often evanescent. Of the first-named
5 larvee only one had the light markings well developed, and
this was selected for painting together with another in which
these markings were almost absent. The remaining larve
were similarly represented by the larva with the white
markings best developed and the one in which they were
feeblest,
between lepidopterous larve and their surroundings. 337
IV. G. quercifolia.
The larvae with green leaves and shoots were very
varied, showing tendencies in the direction of all the other
groups, but chiefly resembling the chequered black-and-
white larvee with the black twigs of Turkish oak, although
the ground-colour was not nearly so dark. A few tended
towards the greater development of the hght markings
which was characteristic of the group with the lichen-
covered sticks, while all of them approached the larve
with the brown stems in the dusky tint of the dark ground-
colour. There was no reason for painting any of these
larvee.
ARRANGEMENTS FOR HYBERNATION AND FINAL
COMPARISON OF THE QUERCIFOLIA LARVAE
BEFORE HYBERNATION.
I. G@. quereifolwa.
October 3rd. Larvee with black twigs. It was evident
that the larvee were ready to hybernate. ‘They had eaten
very little, and some of them were spinning a foot-hold.
The size was that shown on Plate XVII, figs. 1 to 3. A
great change had taken place since the last comparison on
September 21st: only 3 larvee instead of 9 were now
chequered black-and-white, 6 having become nearly as dark
as 5 out of the 6 darkest members of the group. The
white patches of the chequered individual painted on
September 25th were now far duller and less conspicuous.
The larve of this and all the other series were specially
arranged so as to test for the existence of any susceptibility
to the colours of the environment during hybernation. All
were enclosed in muslin sleeves upon a large hawthorn
tree, with pink double flowers, in the garden of Wykeham
House, Oxford. It should be noted that the branches
enclosed in the sleeves were very dark, although not so
intense a black as the twigs of the Turkish oak.
A. The 3 chequered black-and-white larvee were enclosed
with an abundance of black twigs of Turkish oak.
B. The 5 darkest were similarly placed with lichen-
covered twigs.
C. Six dark larvae, but not quite so dark as the last lot,
were similarly enclosed with black twigs,
338 Professor E. B. Poulton on colowr-relation
Il. G. quercifolia.
October 16th. Larvze with lichen-covered sticks. Many
of these larvae had also undergone considerable changes
since September 21st ; so much so indeed in certain cases,
that two of the examples set aside for painting had to be
changed for others which better represented the appear-
ance. borne by the former when they were selected. Of
the 4 lightest larvee on September 21st only one remained
greyish (painted October 2nd, shown in Plate XVII, fig. 6);
2 were now of a brownish tinge (painted October 3rd, fig.
4, and October 7th, fig. 9, Plate XVII); while the 4th
had become so much darker that another brownish larva
was substituted for painting (painted October 6th, fig. 5,
Plate XVII). The latter was not, however, as light as any
of the other 3 set aside on Sept. 21st. Of the other 2
darker larve previously selected for painting, the one with
bluish-white spots had altered, and another more like its
former appearance was substituted (painted October 9th,
Fig. 8, Plate XVII). The remaining 6 dark, white-marked
unpainted larve had not changed, except that the white
marks upon two of them had become smaller and less
conspicuous.
These 6 larvae, with the dark, white-marked one which
was painted on October 10th, and is shown on Plate XVII,
fig. 7, were divided into two groups for hybernation.
D. Four dark, white-marked larve, including the darkest
larva of all the 7 with the smallest white spots, were
enclosed with lichen-covered sticks.
E. Three dark, white-marked larve, including the
darkest larva but one of all the 7, were enclosed with
black twigs of the Turkish oak.
The 3 lightest larva were thus treated.
F. One larva was enclosed with lichen-covered sticks
(Plate XVII, figs. 4 or 9).
G. Two larve, including the greyish one (Plate XVII,
fig. 6), were enclosed with black twigs. The other larva
is represented in Plate XVII, figs. 4 or 9.
H. The two remaining larvee which had been substituted
for painting, viz. the brownish larva (Plate XVII, fig. 5),
and the dark one with bluish-white marks (Plate X VII,
fig. 8), were enclosed with lichen-covered sticks.
hetween lepidopterous larve and their surroundings. 339
Ill. G. quercifolia.
November 3rd. The larvee with brown bramble-stems were
compared and arranged for hybernation. The larve were
more distinctly brown and more uniformly so than when
last compared. The brown patches on three parts of the
surface of certain larve, viz. the 2nd, 5th, and 8th
abdominal segments, were still evident and more distinct
than ever. On some individuals they had increased in
size. In only a single larva were the light patches at all
large and conspicuous. Seven larvee were very uniform
in appearance—dark brown with light brown patches
generally present. The remaining 4 larve were also dark
brown, but 1 was distinctly marked with white, 2 less
distinctly marked with brownish-white, while the dorsal
surface of 1 was overspread with grey. The latter larva
was painted on October 17th (Plate XVII, fig. 12). One
of the brownish-white-marked larvee was painted on October
13th (Plate XVII, fig. 10); and 2 of the 7 first described
were painted on September 30th (Plate XVII, fig. 11) and
October 16th (Plate XVII, fig. 13).
This group of larve was arranged for hybernation as
follows :—
I. Four of the 7 uniform larve were enclosed with
lichen-covered sticks.
J. The remaining 3 were enclosed with black twigs.
K. The 4 more spotted or hghter larvee were enclosed
with brown stems.
First GENERAL COMPARISON AFTER HYBERNATION,
Marcu 26th, 1894.
The weather was very warm on March 25th and 26th,
and the buds of the hawthorn were well out.
I. THE 14 QUERCIFOLIA LARV2 ON BLACK TWIGS BEFORE
HYBERNATION.
A. The 3 chequered larve with black twigs.—All three
larvee were quite healthy, and freely moved about when
disturbed. Two were at rest on the black twigs, 1 was
walking about, probably disturbed by the examination.
They had eaten many of the buds, and one had changed
340 Professor EK. B. Poulton on colowr-relation
its skin probably since hybernation: the old skin was
found in the muslin bag.
B. The 5 darkest larve with lichen-covered sticks. —All
the larvee were in the same healthy and vigorous condition
as those just described, and had eaten the buds of the
hawthorn. Every trace of the winter torpor had dis-
appeared, and they moved freely when touched. The
larvee were uniformly dark brown, and quite unaffected by
the presence of the lichen. Four were resting on a branch
of the hawthorn, while one was on the muslin.
C. The 6 dark larvx with black twigs—Three were dead
and dried up, 2 in the bag and 1 still fixed to a branch of
the hawthorn. Of the living larvae, 2 were fixed to the
muslin and 1 to a branch. One larva appeared to be
unhealthy. One had changed its skin, and the hawthorn-
buds had been eaten. All the larvee, including the dead
ones, were uniformly dark.
It was clear from this comparison that no change had
been wrought by the winter surroundings.
II. THE 12 QUERCIFOLIA LARVA WITH LICHEN-COVERED
STICKS BEFORE HYBERNATION.
D. The 4 dark, white-marked larve, including the darkest
of the 12, with lichen-covered sticks——These larvee had
evidently eaten, and were healthy and easily disturbed.
Three were on branches of the hawthorn, and 1 on a
lichen-covered stick, but all had left their silken foot-holds.
All retained the appearance they possessed before hyberna-
tion: the contrast between dark ground-colour and lght
markings was weakest in the larva on the lichen-covered
stick.
E. The 3 dark, white-marked larve, including the darkest
but 1 of the 12, with black twigs——All 8 larvee were on the
hawthorn branch, and were in the same condition as the
above-described set (D). One was resting on a silken
foot-hold, but this was probably accidental, as there was
no trace of torpidity. All possessed the lchen-like
appearance borne before hybernation.
F. One of the 3 lightest larvx, with lichen-covered sticks,—
The larva was resting on the muslin. It was doubtful
whether anything had been eaten, but the larva readily
moved on disturbance, and was not resting upon a foot-hold.
It still remained one of the three lightest larva, and had
not undergone any change during hybernation.
between lepidopterous larve and their surrowndings. 341
G. Two of the 3 lightest larvx, with black twigs —Both
larvee were healthy and irritable; they had evidently eaten.
One was resting on the hawthorn branch and 1 on a
black twig. They still remained very light, like that
described above (F), and were quite unaffected by their
dark surroundings during hybernation.
H. The 2 larve—the brownish one and that with bluish-
white spots—with lichen-covered sticks,—It is improbable
that anything had been eaten, and these larvee did not
appear to have emerged from hybernation. The one with
bluish-grey marks was upon a lichen-covered stick: it was
shrunken, and it appeared doubtful whether it would
survive. The other was still attached to its foot-hold on
the hawthorn branch, and had evidently not moved during
the winter. It was very lichen-like, and entirely unchanged
by its winter environment.
III. tHe 11 QUERCIFOLIA LARVZ WITH BROWN
BRAMBLE-STEMS BEFORE HYBERNATION.
I. Four of 7 uniformly brown larve, with lichen-covered
sticks— All were healthy and had eaten freely: 3 were
on the hawthorn branch, 1 on the muslin. All were
distinctly brown.
J. Three of 7 uniformly brown larve, with black twigs.—
One larva was dead, while 2 healthy ones had evidently
eaten. Both were on the black twigs, and possessed the
same dark brownish ground-colour with brown patches as
the 4 last-mentioned larvee (1).
K. The 4 most distinctly spotted or lightest larve, with
brown stems.—One larva was dead, while 3 healthy ones
had evidently eaten. All 3 were on the hawthorn-branch.
This set still included the most distinctly white-spotted
individuals of the whole 11. It was evident that no
change occurred during hybernation.
IV. THE 14 QUERCIFOLIA LARVZ UPON GREEN LEAVES
AND SHOOTS OF THE HAWTHORN.
L. Seven larve enclosed in a muslin bag containing a
branch of the hawthorn.—One larva was dead, 3 were on
the branch, 2 on the muslin, while | became detached in
removing the bag. All had left their foot-holds, with the
doubtful exception of one on the branch. All 6 were
healthy and irritable, and most of them had evidently fed ;
. 342 Professor E. B. Poulton on colour-relation
in fact one of those on the branch was eating when the
examination was made. The larve were on the whole
darkish but very variable.
M. Seven larve arranged as in the last set (Li). One
larva was dead, 2 were on the branch, and 4 were on the
muslin. All had left their foot-holds and were healthy
and irritable. They had evidently eaten. Their appear-
ance was similar to that of the set last described.
A complete history of the larve of each set, I to IV,
subsequent to hybernation, will now be given, followed by
an account of the careful comparisons of the whole which
were made from time to time, all the larve being then
placed on a uniform background of white paper.
I. THE QUERCIFOLIA LARVAZ ON BLACK TWIGS BEFORE
HYBERNATION.
A. The 3 chequered black-and-white quereifolia larve on
black twigs throughout.
April Tth—-Re-fed. Three black larve with small
white patches. Unchanged when re-fed on the 10th: all
on twigs.
Apr il 16th—Re-fed. Two larve on twigs, 1 on haw-
thorn: appearance unchanged, as also when re-fed on 20th,
23rd, 27th, and May 3rd: all larve invariably resting on
the twigs.
May Tth—Re-fed. One larva unaltered, while 2 had
changed skin, and the white markings had become less
bright. In one of these the old skin was still adherent
anteriorly. This larva was removed to cylinder C (p. 344).
The other older darker larva was sent to Lord Walsingham
on May 11th.
May 11th.—Re-fed. _ The single remaining larva was at
rest on twig; it was still black with white markings, and
was unaltered when re-fed on the 17th, 22nd, and 25th,
its position on the twig being noted on the two former
dates. It was sent to Lord Walsingham on May 31st.
B. The five darkest quercifolia larve with lichen-covered
sticks during and after hybernation.
April 7th. Re-fed. All 5 larve black, 3 of them with
white spots.
hetween lepidopterous larve and their surroundings. 343
April 10th. Re-fed. Four larve on sticks. Appear-
ance unchanged.
April 16th, Re-fed. Four on sticks, 1 on hawthorn.
Only 1 larva was now entirely black, the remaining 4 had
acquired large white patches. Two larve, including the
first-mentioned, were removed to cylinder B}.
April 20th. Re-fed. All 3 on sticks, appearance un-
changed and on sticks, as also when re-fed on the 23rd.
April 27th. Re-fed. Two on sticks, 1 on hawthorn.
It was noted that im one of the larve the white marks
were less developed than in the others.
May 1st. Re-fed. The last-mentioned larva had
changed skin and the white markings were reduced to a
single anterior pair and were quite small. The 2 others
remained as before, and no further change was seen when
the 3 larvee were re-fed on the 3rd, 11th, 17th, 21st, and
25th. They were noted as at rest on the sticks on all these
dates, except the first and last. The darkest larva was
painted by Mr. Bayzand on May 19th, and is shown in
Plate XVII, fig. 14. Of the two others, the one with the
more intensely black ground-colour was painted by him
on the 22nd, and is shown in fig. 15 of the same plate.
Both these figured larvee were sent to Lord Walsingham
on May 25th, and the third was sent on May 31st.
Bt. The two quercifolia larve separated yrom cylinder B on
April 16th.
Apri 16th. The larvee were respectively entirely black,
and black with white patches.
April 20th. Re-fed. Both on sticks, colour unchanged.
Similar results were noted when re-fed on the 23rd and
27th.
May 1st. Both larve had apparently changed skin.
The darker one had gained a pair of small white marks
anteriorly. On May 5th the larve were brought back
from London. No further change.
May 11th. Re-fed. The black-and-white larva was
sent to Lord Walsingham. The single dark larva, on stick ;
appearance unchanged. Similar notes on the 17th, 22nd,
and 25th, except that it rested on the hawthorn on the
first-named date, and its position was not recorded on the
last-named. The larva was sent to Lord Walsingham on
May 31st.
»
344 Professor E. B. Poulton on colour-relation
C. Three dark quercifolia larve (out of 6) on black twigs
throughout.
April Tth. Re-fed. Two larve dead. The third at rest
on twig; black. When re-fed on the 10th it was on twig
and unchanged.
April 16th. Re-fed. At rest on twig. It had changed
skin and was black with a few whitish spots. When
re-fed on the 21st, 28rd, 27th, and May 3rd it was un-
changed, and on each occasion resting on a twig.
May 7th. Re-fed. At rest on twig; unchanged. A
rather darker larva from cylinder A was added. Part of
the skin of the previous stage was adhering to the anterior
part. When re-fed on the 11th, 17th, 21st and 25th,
these larvae were unchanged in appearance. Upon all
dates except the last it was noticed that they were both
upon twigs. Both larvae were sent to Lord Walsingham
on May 31st.
Pete QUERCIFOLIA LARVA ON LICHEN-COVERED
STICKS BEFORE HYBERNATION.
D. Four dark, white-marked quercifolia larve, including the
darkest of U1, on lichen-covered sticks throughout.
April Tth. Re-fed. Four larvae, of which one appeared
to be unhealthy. All were darkish with large white
patches.
April 10th. Re-fed. One larva had died, 3 upon sticks.
Two had changed skin, 1 remaining unaltered, the other
with a further extension of the white patches.
April 16th. Re-fed. ‘lwo larve upon sticks, 1 upon
hawthorn. All were dark with large white patches, and 1
larva was very small.
April 21st. Re-fed. The 2 larger larve on sticks, the
small one on hawthorn. The former dark grey and pale
greyish-brown respectively; both with very large white
patches. The small larva was brownish-grey with pale
marks. When re-fed on the 23rd all were on sticks and
unchanged, as also on the 26th, except that the small larva
was on the hawthorn.
May 3rd. Re-fed. The pale greyish-brown larva had
changed skin and was now a darker blackish-brown, with
large pale markings. ‘The other larvae were unaltered.
May 11th. Re-fed. Two on sticks, the third, the large
between lepidopterous larve and their surroundings. 345
dark grey larva, having been sent to Lord Walsingham on
this date. The remaining large larva was dark brown
with large yellowish markings; the small larva had
changed its skin and was bluish-grey. The larvee were
re-fed on the 17th and 22nd, when they were on sticks
and unchanged in appearance. The larger larva appeared
to be unhealthy on the 22nd and died on the 28rd.
May \5th. Re-fed. The colour of the small larva was
unchanged, as also on June Ist and 7th. It continued to
grow slowly without further change, and finally spun up
on July 28th.
K. Three dark, white-marked quercifolia larve, including
the darkest but one in II, on black twigs during and after
hybernation.
April Tth. Re-fed.
April 10th. Re-fed. All 3 larvee on twigs. Two larvee
had changed skin, but the appearance of all 3 remained
the same, viz. black with very conspicuous white markings.
They were unchanged and at rest on the twigs when
re-fed on the 16th, 21st and 23rd.
April 26th. Re-fed. All on twigs. The dark colour of
the larve appeared to have become more distinctly grey—
dark grey in two cases, grey in the third. As before, the
dark shade was combined with white markings.
May 2nd. One dark grey and white larva had changed
skin and was blackish and white.
May 11th. Re-fed. The largest larva with a rather
darker ground-colour than others was sent to Lord
Walsingham, The 2 remaining larvee on twigs; both
blackish and white. Re-fed again on the 17th, when they
were on the twigs and unchanged. The smaller larva did
not seem healthy and had not grown like the other. The
larger larva, the darkest of all in set II, was painted on
May 17th, and is represented in Plate XVIII, fig. 4.
May 22nd. Re-fed. Two larvee on twigs. The appear-
ance unchanged, but neither looked healthy. The larger
larva died on the 25th and the smaller on June Ist.
F. One light quercifolia larva on lichen-covered sticks
throughout.
April Tth. Re-fed. White and pale grey.
April 10th. Re-fed. On stick. The larva had changed
its skin, and had become white with markings of two
346 Professor E. B. Poulton on eolowr-relation
shades of brown. It was re-fed on April 16th, 21st (mark-
ings of two shades of pale brown), and 23rd, and was
always found on sticks with appearance unchanged.
April 26th. Re-fed. On stick. The larva was now
white with grey and brownish markings. On May 38rd it
was re-fed, the appearance being unchanged.
May 7th. The larva had changed skin; the white parts
had become yellower. It was re-fed on May 11th, 17th,
and 22nd, and was noted as at rest on stick on the 11th
and 22nd. It remained to the end of a yellowish-white
with markings of grey and brown.
May 25th. The larva was painted by Mr. Bayzand and
is shown in Plate XVIII, fig. 3. It was sent to Lord
Walsingham on the same date.
G. Two light quercifolia larve on black twigs during atd
after hybernation.
April Tth. Re-fed. Both lichen-like.
April 10th. Re-fed. Both at rest on twigs. One had
changed skin, and the greyish markings had become rather
darker. Both were at rest on twigs with appearance
unchanged when they were re-fed on the 16th. _
April 21st. Re-fed. Both on twigs. One larva was
white with pale grey markings, the other creamy-white
with markings of a brownish-grey. Unchanged and on
twigs when re-fed on the 23rd.
April 26th. Re-fed. Both on twigs. The white colour
of the larva first mentioned on the 21st had gained a faint
bluish tinge. The other larva unchanged.
May 5th. Re-fed, after having been brought back from
London. The creamy-white larva had changed skin and
was rather paker in tint. Both were on twigs and un-
changed when re-fed on the 11th. The creamy-white
larva was painted by Mr. Bayzand on the 14th, and is
shown in Plate X VITI, fig. 1.
May 17th. Re-fed. Both on twigs. One larva large
and pale creamy-white with greyish-brown markings.
The other much smaller, white with bluish-grey markings,
but with tints duller than they were formerly. It had not
grown much, nor changed its skin. Both larvee were un-
changed in appearnce, and were resting on twigs when
they were re-fed on the 22nd. The creamy larva was sent
to Lord Walsingham on May 25th. The smaller bluish-
between lempidopterous larve and their surroundings. 347
grey larva was painted on May 28th, and is shown in Plate
XVIII, fig. 2. It was sent to Lord Walsingham on
May 31st.
H. The brownish quererfolia larva and the larva with blursh-
white marks on lichen-covered sticks throughout.
April Tth. Re-fed. One larva dead; the other grey
and white. The larva was re-fed on the 10th, 16th, 20th,
23rd, 26th, and on May 38rd. Its appearance remained
unchanged. On all these dates except the last it was
noted that the larva was at rest on the sticks. On May
5th it was dead.
III. THE QUERCIFOLIA LARVZ ON BROWN BRAMBLE-
STEMS BEFORE HYBERNATION.
I. Four uniform quercifolia larve on lichen-covered sticks
during and after hybernation.
April Tth. Re-fed. All 4 dark brown, 2 rather darker
than the others.
April 10th. Re-fed. All 4 at rest on sticks: 2 un-
changed, and 2 had changed skin, becoming respectively
dark grey with white patches and blackish-grey with white
patches and brown marks.
April 14th. Re-fed. Larve on sticks. The larger ones
unchanged. The 2 smaller had now changed skin, be-
coming respectively very dark blackish-brown with white
marks, and dark brown mottled with pale grey.
April 16th. Re-fed. Larvze on sticks. The 2 larger
darker larvee were now placed in another cylinder. The
two remaining were respectively—mingled shades of dark
and light grey with brown patches, and a uniform grey
with large white patches. These 2 larve were re-fed on
April 21st and 23rd without change. They were always
at rest on the sticks.
April 26th. Re-fed. Both larve on sticks. One larva
retained same appearance, grey with brown patches; the
other was of a pale brownish-grey with paler marks.
May 1st. The latter larva had become of a much darker
brownish-grey with brown and pale marks. On May 38rd
they were re-fed, and both were on sticks: appearance
unchanged.
May 11th. Both larve on Sticks. The last described
348 Professor E. B. Poulton on colour-relation
larva unchanged. The other had now also darkened,
becoming blackish-grey with paler marks. The larve
were re-fed on May 17th, 21st, and 25th, without further
change in appearance. It was noted that they were at rest
on the sticks on the 17th and 21st. On May 31st both
larvee were sent to Lord Walsingham.
I. The two larger darker quercifolia larve separated from
I on April 16th.
April 16th. One larva was now bDlackish-brown with
white marks, the other very dark grey with white and
brown marks.
April 21st. Re-fed. One larva on sticks, 1 on twigs
of hawthorn. Appearance unchanged, as also on April
23rd and 26th, when they were re-fed and both found on
sticks. .
May 1st. The dark grey larva had changed skin and
become blackish-brown with pale and brown spots. On
May 3rd they were re-fed and found unchanged in appear-
ance on the sticks.
May 11th. The larva with pale and brown spots was
sent to Lord Walsingham. The remaining larva was at
rest on stick, and blackish with two pale marks. It was
re-fed on May 17th (on stick) and 25th. On the 31st it
was sent to Lord Walsingham, There was no further
change in its appearance.
J. Two uniform quercifolia larve on black twigs during
and after hybernation.
April Tth. Re-fed. One larva purplish-brown and black,
the other had changed skin and was black with small
white patches and minute brown points. Both were resting
on twigs and unchanged in appearance when re-fed on
the 10th.
April 16th. Re-fed. Both on twigs. The purplish-brown
and black larva appeared to be even darker, the other
unchanged. Both were on twigs and unaltered when
re-fed on 21st and 25rd.
April 26th. The black larva with small white patches
and minute brown points was dying.
April 27th. The last-mentioned larva was dead. The
other purplish-black larva was at rest on a twig.
May 1st, The larva had changed skin, and was black with
hetween lepidopterous larve and their surroundings. 349
a pair of small white marks. It was on a twig unchanged
when re-fed on the 3rd.
May 25th. The larva was sent to Lord Walsingham.
Mr. Bayzand completed his painting of it on the 11th, and
the larva is represented on Plate X VIII, fig. 6.
K. Three of the more spotted or lighter quercifolia larve on
brown bramble-stems throughout.
April 7th. Re-fed. One larva brown, 1 brown with few
white marks. The third grey-brown one had changed
its skin and become brown-grey with white patches.
April 10th. Re-fed. Larve resting on stems. The
brown larva had become brown and very dark grey with
white patches. The other two remained the same.
April 16th. Re-fed. Larve resting on stems. The brown
larva with a few white marks had become darker, and was
now black along the median dorsal area and very dark
brown on the sides: the white patches large. The two
other larvee remained the same.
April 20th, 23rd, and 27th. The larve were re-fed on
each of these dates, and were invariably found upon the
stems. The appearance was unchanged.
May 1st. One of the brownish-grey larve changed its
skin, becoming rather browner in shade.
May 3rd. Re-fed. Three larvee resting on stems. Ap-
pearance unchanged.
May 11th, Re-fed. Two larvee on stems. Appearance
unchanged. Mr. Bayzand finished painting the brownest
larva which is represented on Plate XVIII, fig. 5. The
brownish-grey larva sent to Lord Walsingham for pre-
servation. Another brown one was sent on the 12th.
May 17th, 22nd, and 25th. The single remaining larva
was re-fed on each of these dates. It was noted as at rest
upon a stem except on the 25th. Its appearance was the
same on all occasions, viz. blackish with pale grey marks
and brown spots. It was sent to Lord Walsingham on
May 31st.
IV. THE QUERCIFOLIA LARVZ ON LEAVES, TWIGS, OR
SHOOTS OF THE HAWTHORN THROUGHOUT.
L
April 7th. Re-fed. One brown larva dead. Appearance
af others unchanged, viz. 8 brown and 2 blackish-brown,
350 Professor E. B. Poulton on colour-relation
April 10th. Re-fed. Three larvee unchanged. One brown
and 1 blackish-brown larva had changed skin and become
blackish-grey with large white patches.
April 16th. Re-fed. The two largest larvee last described
removed to another cylinder, L'. The three others un-
changed. One of two smaller brown larve appeared
unhealthy.
April 20th. Re-fed. One larva had changed -skin and
grown much. Of the others, neither of which had changed
skin, one was dead and one appeared to be dying.
April 23rd. The single healthy larva was brownish-black
with white marks.
April 27th, The small larva was dead. The appearance
of the other unchanged, as also on May 3rd and 11th, when
it was again re-fed. On May 17th the larva had died.
i
The two larva separated from L on April 16th were
re-fed and examined on April 21st, 23rd, and 27th, the
appearance remaining the same throughout, viz. dark
grey with large white patches. On May 3rd the larger of
the two larve looked sickly ; on May 5th it was dead and
the smaller one seemed unhealthy. The latter died on
May 7th.
M
April 7th. Re-fed. Five larve alive, all greyish-brown.
One of two unhealthy-looking larve had died.
April 10th. Re-fed. Four larvee unchanged; the fifth
had changed skin and had developed larger white patches.
April 16th. The 2 largest larvae were now separated
and placed in another cylinder, M'. The remaining 3
were brownish-grey, 2 of them ‘with pale patches. The
smallest was browner than the others and appeared to be
unhealthy.
April 21st. Re-fed. The last-mentioned larva was dead ;
the others unchanged in appearance although one had
changed skin during the interval.
April 23rd. Re-fed. Unchanged.
April 27th. Re-fed. One larva had become darker, viz.
blackish-grey with white patches. The larva were re-fed
and compared on May 8rd, 11th, and 17th, without change
in appearance.
May 22nd. Re-fed. The brownish-grey larva had died,
the darker one unchanged.
between lepidopterous larve and their surroundings. 351
May 25th. The last-named larva had died. Neither of
these larvee had grown large.
M"
The 2 largest larvee separated from M on April 16th.
Both were greyish-brown with distinct white patches.
April 21st. Re-fed. Unchanged. The larve were also
re-fed on the 28rd, 27th, May 3rd, 11th, 16th, and 22nd,
and examined on each occasion. The appearance remained
the same throughout. On the 25th both were sent to
Lord Walsingham.
SECOND GENERAL COMPARISON AFTER HYBERNATION,.
April 27th, 1894. The larvee were carefully compared
and were all placed upon a background of white paper.
Nearly all of them were sluggish, probably preparatory to
the last ecdysis.
I. THE QUERCIFOLIA LARVZ WITH BLACK TWIGS BEFORE
HYBERNATION.
A. The three chequered larve with black twigs—These
larvee were unchanged and still remained the lightest of
the whole of series I. Hence the black twigs had _pro-
duced no effect during hybernation.
B. The five darkest larvxe enclosed with lichen-covered sticks
during hybernation.—One was very dark, the darkest of
the whole group; 1 was dark with faint light spots; 3
were black chequered with white markings, which however
were less developed than in A.
It is possible that these last-mentioned three larvee may
indicate some susceptibility to the effect of the lichen-
covered sticks after they were enclosed upon the tree and
before the commencement of hybernation.
C. The six dark larve with black twigs—Only one
larva was alive, and this was dark with very faint dull
white spots.
All the above described 9 larve were healthy and
well up to the average size.
II. THE QUERCIFOLIA LARVZ WITH LICHEN-COVERED
STICKS BEFORE HYBERNATION.
D. The four dark, white-marked larvx, with lichen-covered
sticks——Three larvee were alive and unchanged: one was
very small.
TRANS. ENT. SOC. LOND. 1903.—PART Ill. (ocT.) 24
352 Professor E. B. Poulton on colowr-relation
E. The three similar larve, with black twigs—These also
were practically unchanged and like D. One of these
was now the blackest larva, but the smallest in D was
almost exactly the same. It is possible that some very
slight effect was produced by these black twigs before
hybernation.
F. One of the three lightest larve with lichen-covered sticks.
—The larva was quite unchanged.
G. Two of the three lightest larve with black twigs.—No
effect had been produced by the black surroundings. The
bluish-grey larva remained very distinct.
H. Z'wo larve with lichen-covered sticks —The brownish
larva had died. The other still remained the darkest
individual of the lighter part of the group (F, G, H) as
it was when the arrangements for hybernation were made.
Comparing these two important groups I and II as a
whole, it was seen that the lightest larva of I (@n A) was
almost precisely similar to the darkest of I (in E) :—in
fact they could not have been distinguished as regards the
size of the hght patches. On the other hand, the larva in
I was healthy and of the average size, while that in II
was rather small. The smallest in D was not considered in
this comparison, as it had grown but little and was a stage
behind the others. Its light patches, although very dull
and grey, were almost exactly the same size as those of the
darkest larva in D. The remaining nine larve in II were
all large and healthy, and much lighter than the lightest
larva in I.
III. THE QUERCIFOLIA LARV2 WITH BROWN BRAMBLE-
STEMS BEFORE HYBERNATION.
Comparing these as a whole with sets I and IJ it was
obvious that the lightest of them was distinctly darker
than the darkest of the group just described (IT) and exposed
to lichen before hybernation. The larve were perhaps as
dark as those in group I, exposed to Turkish oak before
hybernation, but they were not so black, and, except in two
larvee, the light markings were less white, being greyish
and clouded over.
I. The four uniform dark larve » with lichen-covered sticks.
—Two larve were very dark with a dull blackish ground-
between leprdopterous larve and thei surroundings. 353
colour. The two others were not so dark and bore brown
points and patches, which tended to fuse at their edges
with the dark greyish ground-colour.
J. The three similar larve with black twigs—Only one
larva was alive, and this was rather darker than the darkest
of the set just described (1), having a dull blackish ground-
colour. One dead larva had a very black ground-colour
with brown dorsal points and distinct although small white
patches. It is probable that some slight effect was
produced by the black twigs just before hybernation
began.
K. The four lightest and most distinctly spotted larva,
with brown twigs.—Three larve were alive. ‘The smallest
one possessed the blackest ground-colour and the whitest
patches of any in the whole series (III), The other two
closely resembled the two lighter larve in I.
IV. THE QUERCIFOLIA LARVZ UPON GREEN LEAVES AND
SHOOTS OF THE HAWTHORN,
L, M. There was no distinction between the two lots.
The larvee, as before hybernation, presented a great range
of variation, but the ground-colour was upon the whole
greyish. The lightest individual was rather lighter than
the darkest of those upon lichen before hybernation (II),
while the next in order was about the same as the darkest
of II. Four others were rather less light, while a fifth was
a distinctly dark form. Hence the set was upon the
whole intermediate between I and II and distinctly
lighter than III.
April 28th. The length of all those larvee which had
ceased to feed preparatory to a change of skin was about
560 mm. A few days later the cast skins were examined,
and were seen to possess the white markings as well as the
dark ground-colour. Hence these characters are in part,
if not entirely, cuticular in position.
THIRD GENERAL COMPARISON OF THE LARVA MADE
AFTER HYBERNATION, MAy 7TH.
or ene QUERCIFOLIA LARVZ WITH BLACK TWIGS BEFORE
HYBERNATION.
A. The 3 chequered larve with black twigs—Two larve
were dark, and 1 black chequered with white markings,
which were larger than those of any other larve in I, but
354 Professor E. B. Poulton on colowr-velation
much smaller than any in II, except the small larva in
E and 1 in D.
B. The 5 darkest larve with lichen-covered sticks.—Three
larvee, 1 of which was changing its last skin, were black
chequered with white markings, which were not quite so
large as those of the larva in A. Two larve were very
dark and unspotted.
C.. The single dark larva with black twigs was changing
its last skin, and was still dark.
Hence probably no etfect had been produced by the
surroundings to which the larvie had been exposed since
hybernation.
II. THE QUERCIFOLIA LARVA EXPOSED TO LICHEN-
COVERED STICKS BEFORE HYBERNATION.
D. The 3 dark, white-marked larve with lichen-covered
sticks.—Two larvee were large in the last stage. One of them
became much darker after changing its last skin, but still
remained black with light markings. The effects of their
dark and light tints were more brownish and yellowish,
and on the whole darker than those of the 2 large larvie
in K. In fact, a comparison of D and E did not support
the conclusion that the larve were sensitive to their
environment after hybernation.
The small larva in D which had lagged behind the
others had changed its skin and was rather hghter. It
was still in an earlier stage than any of the others, but
apparently healthy.
EK. The 3 similar larve with black twigs during and
since hybernation—Two larvie were large in the last stage
and remained black-and-white. The third larva was smaller
and had been injured. It was probably unable to change
its skin in consequence.
F. One of the lightest larve with lichen-covered sticks
throughout.——This larva was in last stage, and its lght
markings had become much darker in tint, being of a
yellowish-brown colour.
G. Two of the lightest larve with black twigs during and
since hybernation.—The lighter of the 2 remained about
as in the previous stage; the other, the bluish-grey larva,
had not yet changed its last skin, but was apparently
between lepidopterous larve and their surroundings. 355
rather darker. These 2 larvee were upon the whole
somewhat lighter than those in F and H.
H. The larve with bluish-white spots exposed to lichen-
covered sticks throughout—This larva was dying. It was
now lighter than the larva in F, but this difference was
entirely due to changes in the latter.
III. THE QUERCIFOLIA LARVA WITH BROWN BRAMBLE-
STEMS BEFORE HYBERNATION.
I. The 4 wniform larve with lichen-covered sticks during
and since hybernation.—Three larvee were in the last stage,
2 of them brownish with small white spots like those in K,
but with the ground-colour darker. The 3rd was con-
siderably darker. The 4th larva was changing its skin.
It possessed a deep brownish-black ground-colour, which
appeared to be overspread with grey.
J. The uniform larva with black twigs during and since
hybernation—The larva was very large in the last stage,
and very uniformly dark and unspotted, although rather
less black than those which had been exposed to black twigs
before hybernation (1).
K. The 3 spotted or lighter larve upon brown stems
throughout.—Two larve in the last stage were dark
brownish with small light patches. The third, in the last
stage but one, was more black-and-white, resembling the
larve which had been exposed to black twigs before
hybernation (1).
Compared as a whole the larvee of I were blacker than
II, although these were very dark. The latter were
distinguished by greyish-brown shades absent from the
ground-colour of I. The lght patches, when present,
were distinguished in III by a brownish tinge, and were
more clouded and less distinct than in I.
Comparing carefully the darkest larva of II (viz. the
4 darkest in D) with those of I, it was seen that the pale
patches were of the same size as those of the larvze in which
they were most developed of all exposed to black twigs
(viz. the lightest larvee in A). But although the patches
were of the same size, those of the former were yellowish-
brown and clouded, and those of the latter white. The
ground-colour of the larva in II was, however, much lighter,
being a brownish-black, than that of the lightest larva in
I. Hence the darkest larva in II was distinctly darker
356 Professor E. B. Poulton on colowr-relation
than the lightest in I as regards its pale patches, but
distinctly lighter as regards its ground-colour. And this
comparison only holds for a single exceptional dark larva.
All others in II were far lighter? than any in either [ or ITT.
IV. THE QUERCIFOLIA LARVZ EXPOSED THROUGHOUT TO
THE LEAVES AND SHOOTS OF THE HAWTHORN.
Only 6 larve remained alive, and of these but 2 were in
the last stage. Four larve possessed a ground-colour very
like that of series III, but the white patches were far
larger, and tended to spread as a greyish shade down the
sides. The light patches were, however, much smaller
than in the larve of series II. One of the smallest larvee
was dark with very small white patches, like one of the
darkest of III. A dying larva, unable to change its skin,
was intermediate between this latter and the 4 first-
mentioned larve.
The colours of these larve in series IV seem to have
been influenced by the brownish and greyish twigs and
shoots of the hawthorn.
Last GENERAL COMPARISON OF THE QUERCIFOLIA
LARVZ AFTER HYBERNATION, MAy 25TH.
Of series I it was recorded that the single larve in A
and 2 in B were black with conspicuous white markings ;
while 1 in B, 1 in B!, and 2 in C were dull black with
only a pair of small inconspicuous white markings.
Of series IT nothing is recorded which is not contained
in the description of cylinders D to G.
Of the 4 larve then remaining in III it was noted that
the larve in K much resembled the black-and-white ones
in A and B; while the 3 in I and [' were dull blackish
with the pale markings very inconspicuous.
Only 2 larve remained alive in IV. The brownish
ground-colour of both was much clouded and overspread
with grey, and the pale patches tended much to spread
downwards, becoming grey and clouded especially towards
the ventral surface.
TRANSFER EXPERIMENTS WITH THE LARV2 OF
AMPHIDASIS BETULARIA IN 1896.
A female moth, captured at Oxford, laid a small batch
of eggs, which provided the material for the following
between lepidopterous larve and their surroundings. 357
experiments, undertaken in order to attempt to ascertain
the susceptible stages of this highly-sensitive larva.
The experiment started on May 10th, when the young
larvee were first fed upon the leaves of Populus nigra, all
dark twigs and branches being at first rigidly excluded.
It must be remarked, however, that for a few days after
May 10th the leaves had only just expanded and were
somewhat brownish. Various sets of larvae were withdrawn
from the stock, and the transference experiments were then
conducted in the following manner. All measurements
were taken when the larve were stretched and straight.
A convenient summary of the results of the following
experiments will be found in the table on page 319.
A. A. betularia.
May 16th. Twenty betularia larve at the end of the Ist
stage, and nearly all changing the first skin, and 5°0 mm.
long, were transferred from green to black surroundings
(the twigs of the Turkish oak). Up to this date the green
leaves of P. nigra upon which they had been placed on
May 10th were somewhat brownish, because the buds had
only just opened.
May 20th. The larvae were examined from time to time
between the 16th and this date, and had always been found
upon the leaves and never upon the twigs. On the 20th
every single larva was found upon the leaves. They even
avoided the stem of the food-plant. Ten larve changing
the 2nd skin, and 8°5 mm. long, were re-transferred into
green surroundings (A'). The 10 remaining larve had
attained various degrees of development in the 2nd stage,
3 being at the end of it, but not yet changing the 2nd
skin. From this date to the 29th these 10 larve were fed
upon Balsam Poplar, but from the latter date onwards
upon P. nigra.
May 25th. Only 1 on the twigs, the rest on the leaves.
One larva was changing the 2nd skin, and none had yet
entered the 3rd stage.
May 29th. Two larvee on the twigs, and both these
were changing the 3rd skin. Of the rest 1 was changing
the 8rd skin and 125 mm. long, and 2 had just changed
it. These 5 were re-transferred to green surroundings
(A2). The remaining 5 were in the 3rd stage, 4 nearly at
the end of it, and 1 very small.
May 31st. Only 1 on twigs. Four changing 3rd skin,
358 Professor E. B. Poulton on colour-relation
the 5th still small in the 3rd stage. The black twigs were
removed at this date, and the 5 larve by this means
re-transferred to green surroundings.
June 2nd. Four in 4th stage, and 1 nearly at the end
of 3rd stage. Four distinct medium brown colour; 1 very
black.
June 5th. One changing 4th skin and light reddish-
brown; 2 in 4th stage, both darkish brown ; 1 at beginning
of 5th stage and medium brown; the 5th small one was
only 9:0 mm. long.
June 14th. One small in 6th stage, and dark brown
with distinct grey markings prominent on it; 1 changing
5th skin and light brown with ventral surface rather
greenish; 2 at end of 5th stage, 1 intermediate and 1
similar to but rather darker than the larva changing its
skin.
June 20th. Four in 6th stage, 2 dark, overspread with
greyish, 1 green with brown dorsal line and _ lateral
patches, 1 dark form becoming greenish on the sides.
June 26th. One dark larva mature and removed.
July 2nd. The green larva and the one with greenish
sides mature and removed. The remaining larva was very
dark, with distinct sharply-marked pale yellowish spots on
its sides, and one on each side of the dorsal surface of each
segment.
July 12th. The larva described above had been
accidentally drowned.
CONCLUSIONS.
The effect of the dark surroundings is evident. The
green environment of the three last stages was doubtless
the cause of the greyish tint, the greenish sides, and the
yellowish spots on the 3 dark larve. In the case of the
4th larva the effects of the latter surroundings were pre-
dominant, although the larva still retained strong traces of
its earlier environment in the brown markings. Comparing
this result with that of A‘, the relative unimportance of
environment in stage IT becomes clear.
Al. A, betularia.
May 20th. The 10 betularia larve re-transferred from
black into green at the end of the 2nd stage, changing the
2nd skin, and 8°3 mm. long.
between lepidopterous larve and their surroundings. 359
From this date to the 29th the larve were fed upon
Balsam Poplar, and then again upon Populus nigra.
May 25th. Most of the larve were changing the 3rd
skin, and all were apparently dark.
May 29th. All had changed the 3rd skin except 2,
which were changing it. All were dark brown except
the 2 latter, which, together with 1 which had just changed,
were light brown. The average length was 14°25 mm.
June 2nd. Four were changing the 4th skin; 5 were
at various points in the 4th stage, with an average length
of 20°75 mm.; 1 was in the Sth stage. All were brown,
although not very dark.
June 5th. Four were changing the 5th skin, 33:0 mm.
long, and 1 at end of 5th stage. The latter was medium
brown, the others 3 light brown and 1 green with light
brown markings. The remaining 5 were much smaller,
being all in the 4th stage, 3 dark brown and 2 light brown.
June 14th. Five were nearly mature, 2 green sprinkled
with distinct greyish dots much more numerous in the
larger larva, which was practically mature and 55:0 mm.
long. The ground-colour of this latter larva was bright
green, the dorsal tubercles dark grey. Of the other 3
large larvee, 1 was very light grey, almost whitish with
darker dots and mottling, 2 were much darker, blackish
rather than brown, with ight grey markings. The largest
of the set of smaller larve was changing its 5th skin and
greenish-brown, 1 smaller in the 5th stage was green with
dorsal brown line, 1 smaller still, chocolate brown. Two
larve were still quite small, viz. 12°25 mm., and probably
still in the 4th stage. Both were dark brown, but they
appeared to be unhealthy.
June 15th. The greenest, the whitish, and 1 dark larva
had become mature and were removed.
June 20th. One green larva mature and removed. Two
in 6th stage, 1 dark with greyish markings not greatly
developed, 1 intermediate, brownish-green; 1 changing
5th skin, green, 2 in 5th stage and lightish brown.
June 22nd. The dark larva in 6th stage mature and
was removed. Two larve in 6th stage, 1 on green side of
intermediate, viz. green clouded over but not entirely
obscured by brown, 1 distinct green with a brown dorsal
line and a little brown on the sides. Of the 2 smaller
larve, 1 was in the 5th stage-and brownish-green, 1 in
the 4th and light brown.
360 Professor E. B. Poulton on colour-relation
June 26th. The brownish-green larva in the 5th
stage was dead. The 2 large ones as last described.
July 2nd. The greenish intermediate larva had become
mature and was removed. The distinct green one was
very large, being 64°6 mm. long. The green ground-colour
was somewhat dull, and the brown dorsal line broad but
not dark. The remaining larva was in the 5th stage and
intermediate.
July 12th. The last-mentioned larva was dead. The
other had pupated.
CONCLUSION.
The effect of a dark environment during the 2nd stage
alone, although slight, is very remarkable. The grey
which overspread the green forms may be compared to the
grey patches on the dark form in this and so many of the
other experiments.
A’, A. betularia.
May 29th. The 5 betularia larvee re-transferred to green
surroundings after they had been in black during the 2nd
and 3rd stages.
May 31st. All 5 larvee were advancing in the 4th stage,
and all dark or distinct brown.
June 2nd. Two larve were changing the 4th skin and
22:0 mm. long; the 3 others advancing in the 4th stage.
Appearance unchanged.
June 5th. One larva changing 5th skin and medium
brown; 2 half-grown in 5th stage and 1 nearly at end of
it; 2 dark brown, 1 of the larger pair light brown. The
5th larva in 4th stage and dark brown.
June 14th. Four approaching the end of the 6th stage ;
2 very black and 2 similar, except that the ground-colour
was overspread with light grey, in one case slightly, im the
other thoroughly. The latter was nevertheless a darkish
larva. The 5th larva was changing the 5th skin, and
green with a brown median dorsal line.
June 15th. One dark larva had become mature and
was removed.
June 20th. One dark larva mature and removed. The
remaining 3 larve were all in the 6th stage. The grey
colour of one of the dark larve was still very distinct.
The brown dorsal line was pronounced upon the green
larva.
between lepidopterous larve and their surroundings. 361
June 26th. The 2 dark larve mature and removed.
The 5th larva had now became a typical and distinct
bright green form, the brown dorsal line having almost
disappeared.
July 2nd. The green larva mature and removed.
CONCLUSIONS.
This is a deeply interesting little set, showing the effect
of the dark surroundings persisting unaltered in the 2
larvee which were first to pupate, the green environment
producing some effect in the greyness overspreading the
next 2, and predominating altogether in the last.
B. A. betularia.
May 20th. The 51 betuwlaria larve remaining in the stock,
with green leaves and shoots of Populus nigra, were care-
fully examined. Two larve had just entered the 3rd stage,
having changed the 2nd skin, 5 were small in the 2nd
stage, while all the rest were changing the 2nd skin, and
thus at the end of the 2nd stage. Twenty of these latter
were transferred to a cylinder (B') with an abundance of
black twigs of the Turkish oak, while the remainder were
put back into the green environment. From this date to
the 25th all the larvae were fed on Balsam Poplar.
May 25th, Thirty larvee counted at this date. Of these,
15 changing the 3rd skin were transferred to a cylinder
(B*) with black twigs. The remaining larve were smaller,
but even in the 3rd stage a small proportion of green in-
dividuals had begun to appear among the brown. All
the larvae were fed on Populus nigra from this date
(May 25th) onwards.
May 29th. The 15 larve remaining in the green environ-
ments were mostly at the end of the 3rd stage and many
were changing the 3rd skin. They were mostly light
brown but some were green.
June 2nd. Six larve had reached the end of the 4th
stage, although they were not changing their skins. Of
these 5 were green with slight traces of brown, while the
6th, although green, retained a larger amount of the darker
shade. Six larve were smaller, having reached various
points in the 4th stage. Of these 2 were green with
slight traces of brown, while four were brownish-green or
hght greenish-brown, Three larve were still in the 3rd
362 Professor E. B. Poulton on colour-relation
stage, and of these the smallest was dark brown while the
other 2 were light greenish-brown.
June 3rd. The five largest larvae were changing the 4th
skin, and were placed in a cylinder (B*) with black sticks.
Their colour was as described on June 2nd, and their
length 20°5 mm. The colours of the remaining 10 larvee
had not altered.
June 5th. Only 9 larvee were found. Two larve were in
the 5th stage and bright green. Five were more or less
advanced in the 4th stage, and were distinctly green with
a variable degree of development of brown patches. Two
were much smaller in the 4th stage and light greenish-
brown in colour.
June 20th. Three larve were large in the 6th stage, and
all very bright green with only a trace or no trace at all
of a brownish tint along the median dorsal line. The other
6 larvze were not noted on this date.
June 26th. Only 8 larvee were found. Two of the
largest green larve had become mature, and were removed
for pupation. The remaining 6 were of various sizes, but
all were bright green except one.
July 2nd. One green larva was mature and was re-
moved. Three were in the 6th stage,2 bright green, one of
them with a little brown on the sides and a brown dorsal
line ; the 3rd was intermediate, with a brown dorsal line,
and green and brown patches alternating on the lateral
surfaces. One was changing the last skin and one in the
5th stage, both bright green.
July 12th. One green larva mature and removed.
The 2 small ones were dead. Of the two remaining larvee
in the 6th stage 1 was bright green and 1 intermediate.
CONCLUSIONS.
The only point which calls for remark is the occurrence
of a single intermediate larva. This was a probable
result of the large numbers of the young larve in a single
cylinder: so that some effect in a specially susceptible
individual followed from the presence of other young
brownish caterpillars.
B!. A. betularia.
May 20th. The 20 betularia larve changing the 2nd skin
transferred from green leaves and shoots to an environment
between lepidopterous larve and their surroundings. 363
of black twigs. The larve were of a very uniform length
of 8°55 mm.
May 25th. Six were found on the black twigs, the rest
on the leaves. Ten larve, all about 140 mm. long, at the
end of the 3rd _ stage, and mostly changing the 3rd skin,
were transferred to green surroundings (C). The remain-
ing larvee were fed on Balsam Poplar from May 20th until
the 29th. Green forms had begun to appear among the
brown.
May 29th. Only 1 on black twigs. Seven larvae were at
the beginning of the 4th stage, 6 dark and 1 greenish.
Two were changing the 3rd skin and 1 nearly mature in
the 3rd stage.
June 2nd. Four larve on the twigs. Two were chang-
ing 4th skin, and § at various degrees of development in
the 4th stage. All brown and the largest larve very dark
brown.
June 3rd. Only 1 on the twigs. Two had now changed
4th skin on the 2nd, and were transferred to green sur-
roundings (C'): 1 was very black, the other a medium
brown.
June 4th. Three on twigs. All larve in 4th stage, all
brown of various shades: those on the twigs very black.
June 5th. Only 1 on twigs and that a light brown one.
One light brown larva changing 4th skin and transferred to
green surroundings (C'). Five in the 4th stage, including
1 dark brown larva which had just died, 1 in 3rd stage.
All brown (more or less dark).
June 7th. Two on twigs, dark and medium brown. The
latter was changing its 4th skin and was transferred to
green (C'), together with another dark brown larva
which had just entered the 5th stage. The remainder
were as last described, and all were advancing in the 4th
stage.
June 14th. Two were on the twigs; both in the 5th
stage, one dark greyish-brown and the other dark chocolate-
brown. Two were on the leaves, both in the 5th stage and
greyish-brown, lighter than the former two.
June 20th. Two large in the 6th stage and very dark.
June 26th. Four in the 6th stage and all very dark
indeed.
July 2nd. Two of the dark larve had become mature
and were removed. The remaining larve were very black
and nearly mature.
364 Professor E. B. Poulton on colour-relation
July 12th. The 2 last larve had forced their way through
the hole in the plate and were drowned. Their appearance
had not changed, and it is probable that they had become
mature and began to wander.
CONCLUSIONS.
In the final result we probably see the full and char-
acteristic effect of the black twigs unmodified by the green
environment in which the two youngest stages were
passed.
B2. A. betularia.
May 25th. Fitteen betularia larve transferred from green
to black; all changing the 3rd skin and 140 mm. long.
Some were becoming greenish, but most were brown.
May 29th. Allexamined: the colour varied very greatly,
but none were altogether green although there was much
green ground-colour on some. All were in the 4th stage;
1 only on the black twigs and that happened to be a
particularly green one. Many light brown and many dark
brown.
June 2nd. Five on twigs, 3 in the 5th stage, 1 chang-
ing 4th skin, and 1 nearly at end of 4th stage. The
latter and 1 in the 5th stage dark greenish-brown, the
remaining 3 very dark brown. On the leaves and green
stems there were in the 5th stage, 1 green, 3 light brownish-
green, and 2 light brown (1 slightly greenish); in the 4th
stage, 2 brownish-green and 2 medium brown.
June 5th. On the black twigs there were 8 larve, 7 in
the 5th stage (3 of them changing the 5th skin), 5 dark
brown, 1 bright green (changing skin), and 1 brownish-
green (half-grown in stage); 1 in 4th stage and brownish-
green. On the light brown stem of P. nigra was a single
larva which harmonized with it very exactly. On the
leaves were 5 larvee, 2 small in the 5th stage and light
brown, 38 in the 4th, 1 dark (changing the 4th skin), 1
brownish-green, and 1 greenish-brown.
June 14th. Six larve in the 6th stage were on the
twigs and very deep black, some of them with a small
amount of greyish markings. One similar larva on
green: 1 larva in the 6th stage on green was of a
uniform dull light grey tint: of 2 larve in the Sth stage,
1 was changing the 5th skin and light brown overspread
between lepidopterous larve and their surroundings, 365
with grey, 1 smaller and darker with less grey. Two
smaller larvee, probably in the 4th stage, or perhaps at
the beginning of the 5th, were respectively lightish and
darkish-brown.
June 15th. One of the darkest larvae had become mature
and was removed.
June 18th. One of the darkest larvee become mature and
was removed,
June 20th. Two very dark larve mature and removed.
One very small one dead. Ofthe 6 in the last stage, 4
were very black, in 3 cases overspread with dull greyish
patches; 1 was greyish on the dorsal surface, lighter
grey elsewhere; the 6th and smallest somewhat resembled
that last described.
June 26th. All had become mature except that last-
mentioned, which had become a dark brownish-black.
July 2nd, No further change.
July 12th. The larva had become mature and was
removed.
CONCLUSIONS.
The power of the black surroundings is evident, the in-
fluence of the green being only seen in the occasional grey-
ness on the black larvz,and especially in the one larva which
was entirely grey. The brownish shade of the larva which
was the last to reach maturity is unusual on the twigs of
Turkish oak. These probable effects of the green on the
larvee which had been longest exposed to the influence of
black (being the last to pupate), are contrary to the results
observed in the other experiments.
B3. A. betularia.
June 3rd. Five betularia larve in green surroundings
up to the end of the 4th stage were transferred to
black. They were changing the 4th skin and 20°5 mm.
long.
June 5th. One larva, a green one, was on a black twig,
the others on the leaves. Four were brownish-green and
1 medium brown, All were advancing in the 5th stage
and about the same size.
June 20th. Three large in the 6th stage, 1 dark smoky-
black ; 1 greyish smoky-black, and 1 intermediate, greenish-
brown. One small one dead, 1 missing.
366 Professor E. B. Poulton on colowr-relation
June 26th. One had pupated, and 1 was mature and
removed. The 3rd larva was greyish smoky-black.
July 2nd. The last larva had become mature and was
removed.
CONCLUSIONS.
The great power of a black environment is well shown,
in the production of 2 dark larvee and 1 intermediate.
At the same time the dark larvee were not quite the
characteristic forms produced by black-barked twigs.
C. A. betularia.
May 25th. Ten of the 20 betwlaria larvie transferred from
green to black for the 3rd stage, and re-transferred to
green at the end of it when they were changing the 3rd
skin and 140 mm. long.
May 29th. All in 4th stage and all dark.
June 2nd. Four in Sth stage, 5 changing 4th skin, 1 not
quite mature in 4th stage. All dark brown.
June 5th, Four changing 5th skin and 33:0 mm. long ;
2 nearly mature in 5th stage; all lightish brown over-
spread with a greyish cloud. Three small in the 5th stage,
2 of them as above and 1 dark brown. One in 4th stage
and very black.
June 14th. Six nearly mature in the 6th stage and all
very dark smoky-black with a pair of distinct grey patches
on the dorsal surface of each segment. Three in the 5th
stage, 2 as above and 1 lighter and really intermediate.
a
June 15th. Three dark larvee had become mature and
were removed.
June 18th. One dark larva mature and removed.
June 20th. Two larvee large in the 6th stage, very dark
smoky-black with the paired segmental light grey patches
distinct.
June 26th. The 2 larve above described had become
mature and were removed. Of the 3 remaining larvie, 2
were large in the 6th stage, one bright green with brown
dorsal line and a small brown patch on anterior part of
each side of the segments, the other smaller and darker
with more brown upon it, but still with a bright green
eround-colour. The third larva in the 5th stage and
chocolate-brown.
July 2nd. All 3 in 6th stage, but the smallest was now
intermediate. In view of the considerable development of
between lepidopterous larve and their surroundings. 367
brown markings the other 2 can only be considered as
rather on the green side of intermediate.
July 12th. The smallest larva was still feeding and still
intermediate. The other two had become mature and
were removed. There was no further change.
CONCLUSIONS.
These results are deeply interesting. The 6 larve
which first became mature were certainly influenced by the
green environment of the three last stages, inasmuch as
the final appearance was a dark smoky-black with a pair
of distinct grey patches on each segment, instead of the
well-known intense dead black which is the characteristic
effect of the black-barked twigs of Turkish oak. At the
same time, the remarkable susceptibility to this stimulus is
seen in the pronounced darkness of these 6 larvee after only
a single stage (of 5 days’ duration) had been passed among
black twigs. It is interesting to note that the 3 inter-
mediate larvee grew more slowly after May 25th, and
thus passed a relatively longer time in the green
environment.
Cl. A. betularia.
June 3rd. Two betularia larve at the beginning of the
5th stage re-transferred to green surroundings after they
had been in black for the 3rd and 4th stages.
June 5th. A third larva, changing its 4th skin, was
similarly re-transferred. It was of a light brown colour.
Of the 2 former, 1 was nearly at the end of the 5th stage
and dark brown, the other rather smaller and darkish
brown overspread with grey.
June Tth. A fourth larva changing its 4th skin and a
fifth at the beginning of the 5th stage were similarly
re-transferred.
June 14th. Three larve in 6th stage, smoky-black with
prominent light grey markings especially distinct in one
of them. One in Sth stage and 1 changing 5th skin, both
dark chocolate-brown with a little grey.
June 20th. Four large in the 6th stage, smoky-black
overspread with grey. The black ground-colour resembles
that of the larvee still in black surroundings.
June 23rd. Two had become mature and were removed.
June 26th. Two mature and removed. The 5th was
TRANS, ENT. SOC. LOND. 1903.—PART UI. (ocT.) 25
368 Professor E. B. Poulton on colour-relation, ete.
now large in the 6th stage, a dark brownish-black rather
than the dead black of the other 4 and those still on the
black twigs.
July 12th. Mature and removed. No further change in
appearance.
CONCLUSIONS.
In this case the grey which overspread. the black
ground-colour of 4 larve must be regarded as an effect of
the green environment during the 2 last stages. The 5th
larva took longer to develop, and there was a slight
departure from the characteristic dead black of the other
tour.
Explanation of Plates. 369
EXPLANATION OF PLATE XVI.
Results of Experiments in 1893 upon the colour-relation between
the larve of Odontopera bidentata and their environment.
All the figures are of the natural size.
Fie. 1.
Nearly mature larva of 0. bidentate showing the effect of
an environment of black-barked twigs (Quercus cerris).
This typical example of the results of Experiment I was
painted by Mr, P. J. Bayzand on July 31st, 1893. All the
larve figured on this plate were fed upon the leaves of
Populus nigra, but they nearly always rested by day on
the twigs or pieces of bark made use of in the experi-
ments here illustrated. The larva represented in this
figure is not in its normal diurnal resting position, having
been disturbed ; and the same is more or less the case
with Figs. 3, 5, 7 and 9.
Nearly mature larva showing the effect of an environment
of weathered pale grey barkless twigs. This typical
example of the final results of Experiment VII was
painted on July 28th. A little earlier the larve had
been rather paler and resembled more closely the majority
of the twigs made use of.
Nearly mature larva showing the effect of an environment
of dark purplish-brown, glossy twigs, probably of birch.
This typical result of Experiment III was painted on
July 30th.
Nearly mature larva showing the effect of an environment
of white-spotted, purplish-brown twigs of birch. This
typical result of Experiment IV was painted on July
30th. Although the details of the environment were
not reproduced, the larva was distinctly less dark than
that shown in Fig. 3.
Nearly mature larva showing the effect of an environment
of green leaves and shoots. The food-plant (Populus
nigra) was employed for this purpose, all dark-barked
twigs being carefully excluded. A comparison of this
figure with the others on the same plate indicates that
the leaves of the food-plant produce no effect when they
are combined with dark twigs or lichen-covered bark ;
while a reference to Experiments I to XV shows that
the great majority of the larve rest by day upon these
latter objects in preference to the leaves. This typical
result of Experiment VIII was painted on July 31st.
370 Explanation of Plates.
Fic. 6. About half-grown larva showing the effect of an environ-
ment of bark covered with bluish-green lichen, probably
Physcia pulverwlenta. This typical result of Experiment
XIII was painted on August Ist.
About half-grown larva showing the effect of the environ-
ment last described. This second typical example of the
results of Experiment XIII was painted on August 3rd.
8. Nearly mature larva showing the effect of an environment
of bark covered with orange lichen, perhaps Physcia
parietina, probably combined with P. pulverulenta. This
typical result of Experiment XIV was painted on
August 4th.
9. Nearly mature larva showing the effect of the environment
last described. This second typical example of the
results of Experiment XIV was painted on August 5th.
10. Nearly mature larva showing the effect of an environment
of lichen-covered sticks. The lichen was probably
Ramalina farinacea. This typical result of Experiment
XV was painted on August 10th.
11. Nearly mature larva showing the effect of the environment
described in Fig. 6. This third typical example of the
results of Experiment XIII was painted on August 31st.
Comparing the last six figures of larve together with the repre-
sentation of the various forms of lichen-covered bark employed in
the experiments, it is seen that there was no special resemblance to
the characteristic features which distinguished one form of environ-
ment from the others. Thus the orange colour of the lichen did not
produce any corresponding effect upon the larve shown in Figs. 8
and 9.
The whole results prove that bidentata is a larva with remarkable
susceptibility to the colour of its environment. In this respect it
is equal to the most sensitive of all larve hitherto tested—Amphi-
dasis betularia. The latter is more susceptible to green leaves and
shoots, becoming bright green when restricted to their influence.
When exposed to lichen-covered bark, however, bidentata was shown,
in Experiments XII to XV, to be far more sensitive.
=I
EXPLANATION OF PLATE KV,
Results of Experiments in 1893-4 upon the colour-relation between
the larve of Gastropacha quercifolia and their environment.
All the figures are of the natural size, and all represent the
normal resting position, except that the larvee more frequently rest
Explanation of Plates. 371
with the head downwards than is shown in the plate. Perhaps the
young larvee under normal conditions invariably rest in this position,
Figs. 1—13 represent the larve in the autumn of 1893, just before
the beginning of hybernation.
Figs. 14 and 15 represent the larva, nearly mature in the last
stage, in May 1894.
Fia. 1. Larva of Gastropacha quercifolia just before hybernation,
showing the effect of an environment of black-barked
twigs (Quercus cerris). This typical example of the nine
black larvee, chequered with white, described on Septem-
ber 21st, 1893, was painted by Mr. Bayzand on September
25th. Although these larve had eaten very little, and
had not grown appreciably by October 3rd, the white
marks on six out of nine of them had become reduced
almost to the condition represented in Fig.3. The white
patches on the larva shown in Fig. 1 had also become
much duller and less conspicuous. The persistence of
the process of colour-adjustment right up to the begin-
ning of hybernation is very interesting, and contrasts
remarkably with its entire cessation during and after
hybernation. The difficulty with which the contour of
the larvee could be made out against the black bark is
correctly rendered in Figs. 1—3.
2, Larva at the same period and exposed to the same environ-
ment as that shown in Fig. 1. The figure represents the
darkest larva, without any trace of white markings,
described on September 21st. The painting was made
on September 23rd.
3. Larva at the same period and exposed to the same en-
vironment as that shown in Fig. 1. This typical ex-
ample of five out of the six darkest larvee, described on
September 21st, was painted on September 26th.
4, Larva just before hybernation, showing the effect of an
environment of lichen-covered sticks. The lichen was
probably Ramalina farinacea. The figure represents
one of the four lightest coloured larve produced
in this environment and separated for painting on
September 21st. The larva was painted on October 3rd.
By October 16th, when the larve of series II were
arranged for hybernation, many changes had taken place,
but this larva remained among the lightest throughout.
5. Larva at the same period and exposed to the same environ-
ment as that shown in Fig. 4. The figure represents
372 Explanation of Plates.
one of the four lightest larvee on October 6th when it
was painted. It was not among the four lightest larvae
on September 21st, but changes took place after this date.
Fic. 6 Larva at the same period and exposed to the same en-
vironment as that shownin Fig. 4. The figure represents
one of the four lightest larvee separated for painting on
September 21st, and the only one which remained of a
greyish tint, the others becoming brownish. It was
painted on October 2nd.
7. Larva at the same period and exposed to the same en-
vironment as that shown in Fig. 4. The figure repre-
sents one of the commonest types of appearance on
September 21st, viz. the group of six dark white-marked
larve. The drawing was made on October 10th.
8. Larva at the same period and exposed to the same
environment as that shown in Fig. 4. Among the
eight darkest larve of this series II on September 21st,
was one in which the white markings possessed a bluish
tinge. This was set aside for painting. But changes
took place later on, and one of the other seven larve was
found more nearly to represent the previous appearance
of the separated larva. The former, which had become
greyish, was therefore painted on October 9th.
9. Larva at the same period and exposed to the same en-
vironment as that shown in Fig. 4. The description of
Fig. 4 applies in every respect except that the larva here
represented was painted on October 7th.
10, Larva just before hybernation, showing the effect of an
environment of reddish-brown stems of bramble. The
larva represented was one of the two mentioned on
October 16th, in which the light patches were well
developed and of a brownish tint. It was painted on
October 13th.
11. Larva at the same period and exposed to the same en-
vironment as that shown in Fig. 10. The larva repre-
sented was the lightest of the seven very uniform dark
brown larve with lighter brown patches and small white
marks mentioned on October 16th. These light patches,
which are not very distinct in this figure and in Fig. 13,
were generally present on the 2nd, 5th, and 8th abdominal
segments. The larva was painted on September 30th.
12. Larva at the same period and exposed to the same en-
vironment as that shown in Fig.10. The larva repre-
sented was the brown individual with the dorsal surface
Explanation of Plates. 373
overspread with grey mentioned on October 16th. The
greyish appearance does not come out in the figure, the
effect being merely to render the brown of a paler tint-
The larva was painted on October 17th.
Fic.13. Larva at the same period and exposed to the same envi-
ronments as that shown in Fig. 10. The description of
Fig. 11 applies in every respect, except that the larva
here represented was a specially dark example, and was
painted on October 16th.
14. Larva, nearly mature in the last stage, showing the effect
of an environment of black-barked twigs up to the
beginning of hybernation. During and after hybernation
the larva was placed (I, B) in an environment of lichen-
covered sticks, but, as the figure indicates, it had ceased
to be susceptible to such influences, and no effect was
produced. Before hybernation it had been one of the
five darkest larvee. The painting was made on May 19th.
15. Larva at the same period and exposed to the same environ-
ments (I, B) both before and after hybernation as that
shown in Fig. 14. Although conspicuous white patches
appeared on this and other larvae subsequent to hyber-
nation, it is improbable that this effect was due to the
lichen which formed the surroundings after the begin-
ning of hybernation. The comparison of the whole
of the larve indicates that they had then ceased to be
susceptible to the colours of the environment. The
larva was painted on May 22nd.
EXPLANATION OF Puate XVIII.
Results of Experiments in 1893-4 upon the colour-relation between
the larve of Gastropacha quercifolia and their environment.
All the figures are of the natural size, and all represent the
natural resting position.
All the figures represent the larvee in the last stage, and all but
one nearly mature, in May 1894.
Fic. 1. Larva of Gastropacha quercifolia, nearly mature in the last
stage, showing the effect of an environment of lichen-
covered sticks up to the beginning of hybernation.
The lichen employed was probably Ramalina farinacea.
During and after hybernation the larva was placed in an
environment of black-barked twigs (II. G) which, it is
Fig; 2.
Explanation of Plates.
obvious, produced no effect whatever. The appearance
of this same larva just before hybernation is represented
in Plate XVII, fig. 4 or 9. The painting of the nearly
mature larva was made on May 14th.
Larva, small but probably in the last stage, exposed to the
same environments (II, G) both before and after hyber-
nation, as that represented in fig. 1. Here too it is
clear that the black-barked twigs which surrounded the
larva during winter and spring produced absolutely no
effect. The appearance of the same larva just before
hybernation is represented on Plate XVII, Fig. 6. The
painting of the more mature larva was made on
May 28th.
3. Larva, nearly mature in the last stage, showing the effect
of lichen-covered sticks throughout (II, F). The lichen
was probably Ramalina farinacea. The appearance of
this same larva just before hybernation is represented in
Plate XVII, fig. 4 or 9. The painting of the nearly
mature larva was made on May 25th.
4. Larva at the same period and exposed to the same
environments both before and after hybernation as that
represented in Fig. 1. The larva here represented (from
Il. E) was the darkest of all the mature larve which
had been exposed to an environment of lichen before
hybernation (series I]). There is no reason to sup-
pose that the black twigs produced any effect in winter
and spring. The larva was one of the seven darkest
in series I] before hybernation. The painting was made
on May 17th.
Larva, nearly mature in the last stage, showing the effect
of reddish-brown stems of bramble throughout (III, K).
The specimen represented was one of the four more
spotted or lightest larve before hybernation, and the
same relationship towards the other divisions of this
series (III) was maintained during and after hyberna-
tion. The painting was finished on May 11th.
G6. Larva, nearly mature in the last stage, showing the effect
reddish-brown stems of bramble before hybernation.
During and after hybernation the larva was placed in an
environment of black-barked twigs (III, J). It had
been one of the uniform brown larve before the winter,
and there is no reason for the belief that the black twigs
introduced later produced any effect. The painting was
finished on May 11th.
7-_~
oo
~T
OV
-
XIX.—A Revision of the Old World Lymeantriidz in the
National Collection. By CoLONEL CHARLES
Swinnog, M.A, F.LS., ete.
THE order in which the genera and species of this Family
are placed in this paper is the same as that in which they
now stand in the National Collection.
In working out the material, I have gone through the
whole of the accessible literature: I may have increased
the synonomy somewhat by describing species that have
been described before, but their careful descriptions will in
any case be useful.
The difficulties to be overcome in examining literature
published in German, Dutch, Swedish, and French is much
increased by the extraordinary system now prevailing in
the formation of indices to the various scientific serials.
With the exception of the “ Tring Journal,” the “ Journal
of the Bombay Natural History Society ” (and perhaps one
or two others), all the scientific pericdicals have generic
indices ; this may be sufficient for mammalia, ete., where
the genera are so few you can almost count them on your
fingers : but for entomological workers such indices are
merely an aggravation, and are absolutely useless with so
large a number of genera. In working out small insects
mistakes in the reference of species to genera are very
liable to be made, and it is next to impossible te decide
with any accuracy the genus in which an author has put
the species one is looking for. The result of this is that
in working out references it becomes necessary to plod
through the papers of each writer, species by species.
I have found it neces ssary to sink a number of so-called
genera, especially under the well-known genus Dasychira ;
and though I have twice examined every ‘type specimen of
every one > of these genera, I can detect no distinctive points
in any of them sufficient to differentiate it.
Both sexes of typical Dasychira have peculiar dorsal
tufts of hairs on the abdomen, and a second characteristic
is the enlarged and truncate formation of the anal segment
of the abdomen, especially noticeable in the female.
TRANS. ENT. SOC. LOND. 1903.—PART IL (OCT.)
376 Colonel C. Swinhoe’s Revision of the
Sir George Hampson, in his able work on the Moths of
India, has included in this genus, under separate sections,
moths otherwise agreeing, but without these particular
characteristics, and I have followed him; but I should here
point out that the species without dorsal tufts on the
abdomen have likewise the anal segments pointed in both
sexes, and that therefore these sections are to that extent
divergent from the type. I have, however, been content
to separate merely into sections those species with tufts
and those without, as a guide for future workers.
The species of the Australian genus Anthela = Colussa
= Darala = Newmania differ in the neuration of the
hind-wings from typical Lymantriide; veins 7 and 8 are
divergent from base, and never approximate at any point
of the cell. In one species, Anthela (Colussa) varia, the
type of the genus Colussa, these veins are united by a
strong bar. ‘This disposition of the veins 7 and 8 of the
hind-wings is the same as in the Hupterotide; but vein 5
of both wings rises below the middle of the discocellulars,
whereas in Lupterotide it should arise from, or from above
the middle. This Australian genus therefore appears to
claim a place for itself intermediate between the true
Lymaniriide and the Eupterotidxy, and though included in
the former in this paper, must be considered as partially
abnormal.
Genus DENDROPHLEPS, Hmpsn., Moths, India, i, p. 491
(1892).
DENDROPHLEPS SEMIHYALINA.
Dendrophleps semihyalina, Hmpsn., |. ¢.
1 Z, Khasia Hills (type). 1 9, Sikhim.
Genus CARAGOLA, Moore, Lep. Atk., p. 46 (1879).
CARAGOLA IMPRESSA.
Leucoma impressa, Snellen, Tijd. v. Ent., xx, p. 8, pl. 1, f. 1
(1877).
Redoa impressa, Kirby (part), Cat. Moths, 1, p. 435 (1892).
1 9, Sandakan. 1 9, Fergusson Isl.
Snellen’s type came from Java. It is just as distinct
from cygna, Moore, as rinaria, Moore, is from dica, Swinh. ;
I have both sexes from Java and Amboina; the male has
acute fore-wings like the males of ochripes, Moore.
Old World Lymantriide in the National Collection. 377
CARAGOLA RINARIA.
Redoa rinaria, Moore, Cat. Lep. E.1.C., u, p. 336 (1859).
Caviria rinaria, Hmpsn. (part), Moths, India, i, p. 490
(1892).
Leucoma margaritacea, Snellen, Tijd. v. Ent., xxix, p. 35,
pl. 1, f. 2, 2a (1886).
Arctornis snelleni, Kirby, Cat. Moths, i, p. 432 (1892).
1 ¢, Sumatra. 3 3,1 9, Java, including the type.
Snellen’s type came from Sumatra.
CARAGOLA DICA.
Redoa dica, Swinh., Trans. Ent. Soc., 1891, p. 478.
Caviria rinaria, Hmpsn. (part), Moths, India, 1, p. 490
(1892).
1 2, Khasia Hills (type).
I do not think the sinking of this form to the Island
species is justifiable ; dica is much larger, and has rounded
hind-wings, whereas the hind-wings of vinavia are more or
less square, the margin being straight from vein 2 to vein
7; Ihave both sexes from the Khasia Hills in my own
collection.
CARAGOLA CLARA.
fRtedoua clara, Walker, xxxii, 343 (1865).
Caviria clara, Hmpsn., Moths, India, i, p. 490 (1892).
i$) Hong: Kone... 3.2,
type. '
CARAGOLA CYGNA.
Caviria cygna, Moore, P. Z. §., 1877, p. 601.
Caviria eygna, Hmpsn., Moths, India, 1, p. 489 (1892).
1 g, Ceylon (type). 1 2, Andamans (type).
CARAGOLA MARIA.
Redoa maria, Kirby, Ann, Mag. N.H. (6), xviii, p. 383
(1896).
1 g, Mtzbe, E. Africa (type).
CARAGOLA SERICEA.
Stilpnotia sericea, Moore, Lep. Atk., p. 45 (1879).
Caviria sericea, Hmpsn., Moths, India, i, p. 490 (1892).
1 $,39,Masuri. 2 ¢,2 2, Sikhim, including the types.
42, Thibet. 4 9, Kashmir.
5 ¢, Sikhim, including the
378 Colonel C. Swinhoe’s Revision of the
CARAGOLA OCHRIPES.
Stilpnotia SEES) Moore, Lep. Atk., p. 45 (1879).
Denar ophleps ‘semil: yalina, o. Swinh, (nec Hmpsn. »; Thine
Ent. Soc., 1895, p. 14.
1 g, Moupin, W. China. 1 9, Chin Hills. 1 2, Khasia
Hills.
The type from Darjiling is in coll. Staudinger.
Genus Leucoma, Steph., Ill. Brit. Ent. Haust., ii, p. 64
(1829).
Rtedoa, Walker, iv, 826 (1855).
Kanchia, Moore, Lep, Ceylon, 1, p. 92 (1883).
LEUCOMA SUBMARGINATA.
Redoa submarginata, Walker, iv, 826.
Leucoma submarginata, Hmpsn., Moths, India, i, p. 487
(1892).
Redoa transiens, Walker, Journ. Linn. Soc. Lond., vi, p. 128
(1862).
Leucoma hipparia, Swinh., Ann. Mag. N. H. (6), xii, p. 214
(1893).
2 ¢, Sarawak, Borneo, including the type of transiens.
1 ¢,Sandakan. 1 ¢, Pulo Laut. 3 ¢,1 2, Singapore,
including the type of ese (as oh ok Se lan eon | ote
Fergusson Isl. 2 N. Guinea. 1 ¢, N.Ireland. 1 ¢,
NaBritam, 2 fy i Java. 1 ¢, Chusan Isl. 1 ¢,
Ichang, W. China. t,19, Darjiling. 2 3, 19, Cachar.
5 f, Trevandrum. 3 au Ceylon. 1 #, Maulmein. 1 @,
1 2, Silhet (types). 2 $, Burma. 1 ¢,Andamans. 1 @,
Nilgiri Hills.
LEUCOMA CYGNA.
Redoa cygna, Moore, P. Z. S., 1879, p. 401.
ftedoa cymbicornis, Butler, Ill. Het., v, p. 48, pl. 89, f. 2
(1881).
Redoa nigricilia, Swinh., Trans. Ent. Soc., 1881, p. 478.
1 9, India (type). 2 9, Yokohama. 1 ¢, Pu-tsu-fong,
China. 5 ¢,1 2, Sikhim, including the type of cymin-
cornis. 1 2, Khasia Hills (type nigricilia). 1 3, Nilgiri
Hills. 1 ¢, Andamans. 3 ¢, Penang. 1 ¢, Borneo.
1 gf, Singapore. 2 2, Queensland.
g,N
i
Old World Lymantriide in the National Collection. 379
LEUCOMA FLAVESCENS.
fedoa flavescens, Moore, P. Z. S., 1877, p. 600.
Leucoma flavescens, Hmpsn., Moths, India, i, p. 488 (1892).
Redoa sericea, Moore, |. c.
3 $, Andamans, including both types. 1 7,19, Nilgiri
Hills. 2 ¢, Travancore.
LEUCOMA PRUINOSA.
Leucoma pruinosa, Butler, Ann. Mag. N. H. (5), iv, p. 236
(1879).
Arctornis pruinosa, Kirby, Cat. Mothis, i, p. 433 (1892).
1 2, Madagascar (type).
LEUCOMA NITIDA, nov.
6, 2. Palpi, fore-legs and frons orange ochreous; top of head,
middle and hind legs, body and wings above and below pure white ;
antenne with the shaft white, branches ochreous grey ; thorax of the
male suffused with orange ochreous ; fore-wings with a beautiful
silvery sheen, with thin longitudinal curved waves in certain lights;
the first runs from base to apex, the second from hinder margin near
base to outer margin above the middle, and the third from the middle
of the hinder margin to outer margin below the middle. Costal line
and cilia ochreous grey ; hind-wings without the sheen and without
any markings.
Expanse of wings ¢ 1,8,, 9 2.3, inches.
gf 2, types, Old Calabar (S. D. Crompton and F. W.
Sampson). 1 2, Ogove River. 1 9, Lagos (H. Strachan).
1 2, Sapele, River Niger. 1 2, Gold Coast (W. Ef. Johnston).
LEUCOMA LUTEIPES.
Stilpnotia luteipes, Walker, 1v, 843, 2 (1855).
Homeomeria luteipes, Kirby, Cat. Moths., i, p. 437 (1892).
Redoa laba, Schaus and Clements, Lep. Sierra Leone, p.
25, pl. 1, f. 4 g (1893).
ftedoa ogovensis, Holland, Ent. News, Phil., 1893, p. 63, pl.
3, f. 12, 13.
2 $,2 2, Sierra Leone, including the type. 1 7, Old
Calabar. 1 2, Ogove River.
There can be no doubt that Jaa is its male; the locality
and description fit it exactly; and I have compared an
example from Mr. Druce’s collection, given him by Dr.
Holland as ogovensis, with Walker's type: these are also
identical.
380 Colonel C. Swinhoe’s Revision of the
LEUCOMA EGERINA.
Leucoma egerina, Swinhoe, Ann. Mag. N. H. (6), xu, p. 214
(1893).
4 4,12, Singapore, including the type.
LEUCOMA MARGINALIS.
Redoa marginalis, Walker, Journ. Linn. Soc. Lond., vi, p.
128 (1862).
Leucoma marginalis, Swinhoe, Cat. Het. Mus. Oxon., i, p.
202 (1892).
1 2, Singapore. 3 7,Sarawak. 1 7,1 2, Kuching.
The type from Sarawak is in Mus. oe
LEUCOMA ALBA
Aroa alba, Brem., Bull. Acad. Pet., iii, p. 478 (1861).
Aroa alba, Brem., Lep. Ost. Sib., p. 41, pl. 5, £ 18 (1864).
Leucoma alba, Leech, Trans. Ent. Soc., 1899, p. 143.
Redoa sinensis, Moore, Ann. Mag. N. H. (4), xx, p. 92
(1s77).
=e ie 2 China, including Moore’s type. 1 ¢
oS Oo}
Shanghai. ie 192, Fusan. 2 ¢,1 2,Gensan.
LEUCOMA MOOREI.
Leucoma moorei, Leech, Trans. Ent. Soc., 1899, p. 1453.
Redoa alba, Moore, Ann. Mag. N. H. (4), xx, p. 92 (1877)
(preoce.).
1 2, Shanghai (Moore’s type). 3 7, Ichang. 12, Chang
Yang. 2 2,12, Omeishan. 3 a , Moupin. 1 Q, Wa-
Shan.
LEUCOMA MINUTISSIMA, nov.
4. Pure white; frons, pectinations of the antennz, and the tarsi
tinged with greyish ochreous; a very small black dot at the end of
the cell of the fore-wings ; otherwise the insect above and below is
entirely without markings.
Expanse of wings 4 inch.
Hah, SARAwAk, Borneo (Wallace).
I should think it is the smallest species of the genus;
the example is in excellent condition.
LEUCOMA DIVISA.
Euproctis divisa, Walker, iv, 836 (1855).
1 Z, Silhet (type). 1 7%, Bhutan. 2 7, Donat Range,
Burma. 5 7,1 2, Singapore. 1 f,1 2, Penang, 17,
12, Java. 2 7, Pulo Laut. 1 27, Sarawak.
Walker described two perfectly distinct moths as male
and female of his species; I take his type to be the first
one, following his description; it isa female from Nepal,
a true Huproctis; the next is a male from Silhet, a
Leucoma, and as the name is not preoccupied and his
deseription is sufficient, it must stand.
LEUCOMA SILHETICA.
Penora sthetica, Walker, xxxii, 341 (1865).
1 7, Khasia Hills. 1 2, Silhet (type).
This is a common form in the Khasia Hills: it is almost
completely hyaline, and is quite distinct from divisa, which
is well clothed in both sexes.
LEUCOMA DIAPHANA.
ftedoa diaphana, Moore, Lep. Atk., p. 46, no. 1051, ¢
(1879).
Redo lactea, Moore, |. c., no. 1053 2.
3 2, Bengal 2 ¢, Kulu. 1 7,1 Q, Omeishan.
Moupin. 1 2, Kwei-Chow.
LEUCOMA PELLUCIDA, nov.
3, 2- White; palpi and top of head orange ; frons white ; antennz
and thorax tinged with orange in the males; wings very thinly
elothed, almost hyaline ; the costal line orange ; cilia tinged with
orange ; a small, rather promment, black mark in the middle of the
discoidal vein of fore-wings ; fore-legs orange, hind-legs white.
Expanse of wings ¢ 135, 2 2 3, imehes.
Hab. Kuasts HILts.
I have two from the same locality in my own collection ;
one I have had for some years unnamed.
LEUCOMA TIPHIA, nov.
3; 2. Antenne black, the shaft speckled with white; palpi
black with some ochreous hairs ; frons with some ochreous hairs in
the male ; pectus ochreous in the male, and the thorax below covered
with ochreous brown hairs ; legs white ; fore tibiz striped with black
in front in both sexes ; head, bedy, and wings above and below pure
white sparsely covered with shining scales, semi-hyaline : costal line
382 Colonel C. Swinhoe’s Revision of the
of fore-wings black for one-third of its length from the base above
and below, but below the whole costal space is also tinged with
ochreous ; no other markings.
Expanse of wings ¢ 1,%,, 2 144 inches.
Hab. Natropt Forest, Kikuyu, British E. Africa
(Crawshay), 4 $,1 @.
LEUCOMA USEBIA, nov.
¢. Antenne black, shafts whitish ; frons, fore-legs, and hind
tibia ochreous; otherwise the insect is pure white without any
markings whatever above or below ; veins 6 and 7 of the hind-wings
are on a long stalk.
Expanse of wings 1,3, inches.
5 ft, Lake Nyassa (de Jersey).
LEUCOMA CROCIPES.
Cypra crocipes, Boisd., Faune.Madag., p. 87, pl. 12, f. 2
(1833).
Cypra crocipes, Kirby, Cat. Moths, i, p. 438 (1892).
2 ¢, Madagascar.
LEUCOMA TAVETENSIS.
Leucoma tavetensis, Holland, Ent. Suppl. xxv, p. 93
(1895).
Antiphella telesilla, Druce, Ann. Mag. N. H. (7), iui, p. 469
(1889).
The type came from the Taveta Forest, Kilimanjaro
(Abbott), and is in the U.S. Nat. Mus.: in the B. M. there
isone male from the same locality (Hannington), and two
males from Lagos (Strachan); there is no apparent difterence
between the Eastern and Western examples ; Druce’s type
is from Zanzibar; it is rather larger than the continental
examples, but otherwise identical.
LEUCOMA GRACILLIMA.
Leucoma gracillima, Holland, Ent. News, Phil., 1893, p. 64,
pls, 1. o.
1 Z, Ogove River. 1 2, Old Calabar.
LEUCOMA VATA, nov.
¢. Antenne and frons orange ; legs white, orange-grey in front ;
head, body, and wings above and below silky white ; costal line of
fore-wings orange ; no other markings; fore-wings rather long and
Old World Lymantriide in the National Collection. 383
narrow, the outer margin being longer than the hinder margin, the
hinder angle well rounded.
Expanse of wings 1,5 inches.
Hab. River NIGER, Sapele (&. W. Sampson).
LEUCOMA RUFIMARGINATA, Nov.
S, °. Pure silky white ; palpi and branches of antenne, body
beneath, and legs ochreous grey, the last with some dark brown
spots ; costa of fore-wings rufous; a spot of that colour at the end of
the cell, and the outer margins and cilia of both wings rufous, except
at the angles.
Expanse of wings ¢ 1 inch, 2 1% inches.
2, 1-2, Pulo Laut:
Allied to LZ. submarginata, Walker, but has no white
frons, and no black and yellow spots and markings on the
fore-legs.
LEUCOMA PRIMULA, nov.
¢, ¢. Palpi chestnut-brown, whitish beneath and on the inner
sides ; antenne grey, the shafts pale primrose ; frons dark chestnut-
brown ; legs, body, and wings above and below of a uniform pale
primrose colour ; fore-wings with the apical third of the costa and
the outer margin, including the cilia, dark chestnut-brown, a small
space just before the hinder angle not coloured ; hind-wings with
the outer marginal line and cilia from a little before the middle to
near the anal angle similarly coloured ; a very small brown dot in the
middle of the discoidal vein of the fore-wings. The fore-wings are
broad, and the hinder margin is nearly as long as the costa.
Expanse of wings ¢ 1,4, 2 3; inches.
Hab, SANGIR (Doherty).
LEUCOMA MICACEA.
Redoa micacea, Walker, Journ. Linn. Soe. Lond., vi, p. 127
(1862).
Leucoma micacea, Swinhoe, Cat. Hee Mus. Oxon., 1, p. 203
(1892).
4%, Sarawak, Borneo, including the type.
LEUCOMA FLORA, nov.
¢. Palpi and antennx pale chestnut ochreous, shaft of antenne
grey speckled with white ; frons pure white ; top of head white ;
space between the antenne chestnut-brown ; fore-legs with brown
TRANS. ENT. SOC. LOND. 1903.—PART III. (OCT.) 26
384 Colonel C. Swinhoe’s Revision of the
patches, tarsi ochreous ; body and wings white tinged with primrose;
thorax and fore-wings slightly irrorated with chestnut-brown atoms ;
a suffused lunular mark of that colour at the end of cell; the costa
near apex and the upper middle portion of the outer marginal space
both rather darkly suffused with the same colour; on the hind-
wings, which are otherwise unmarked, there are triangular marks on
the outer margin, and the cilia are of paler chestnut-brown.
Expanse of wings 1,}, inches.
Pulo Laut, 2 $ (Doherty); Sandakan, 1 gf (Pryer).
LEUCOMA DISCIRUFA, nov.
Antenne, palpi, head, body, and fore-wings above ochreous fawn
colour ; fore-wings with a large ochreous red mark at the end of the
cell ; the costal margin and outer marginal line and cilia of both
wings of the same colour ; hind-wings with the costal half whitish,
the lower half the same colour as the fore-wings ; a few minute black
irrorations in the fore-wings ; underneath, body, legs, and wings pale
yellow, nearly white, the wings with the margins and cilia red.
Expanse of wings | inch,
Hab. PuLo Laut (Doherty).
LEUCOMA SATURNIOIDES.
Laelia saturniordes, Snellen, Tijd. v. Ent., xxii, p. 105, pl.
8, f. 7, 7a—c (1879).
Leucoma saturnioides, Snellen (part), Iris, vii, p. 139
(1895).
The type came from Celebes: it is in the B. M. from
Singapore.
The venation is well figured by Snellen ; it is not quite
typical, but very nearly so, vein 5 arising a ‘Little above the
lower end of the cell instead of from it: it certainly is nota
Laelia. Snellen says Leucoma fenestrata, Hmpsn., Il. Het.,
ix, p. 78, pl. 160, f. 6 2 (1893), is the same thing; but in
this I am convinced he is wrong; Hampson’s and Snellen’s
figures are both good ; both the types are females, so is the
example of satuwrnioides from Singapore ; and they appear
to me to be quite distinct one from the other.
LEUCOMA FENESTRATA.
Leucoma fenestrata, Hmpsn., Moths, India, 1, p. 489
(1892).
Macrauzata fenestrata, Hmpsn., Il. Het., ix, p. 78, pl. 160,
f. 16 (1893).
1 , Ceylon (type).
Old World Lymantriide in the National Collection, 385
LEUCOMA SUBVITREA.
Leucoma subvitrea, Walker, xxxil, 344 (1865).
Kanchia subvitrea, Moore, Lep. Ceylon, 1, p. 93, pl. 113, f.
5 (1883).
1 3,4 9,Hong Kong. 1, Bengal (type).
LEUCOMA L-NIGRA.
Bombyx l-nigrum, Miill., Faun. Fridr., p. 40 (1764).
Bombyx v-nigrum, Fabr., Syst. Ent., p. 577 (1775).
Leucoma v-nigra, Steph., Ill. Brit. Ent. Haust., ii, p. 64,
pl. 16, £2
&, 2, Tokio, Japan (Pryer), and many European
examples.
LEUCOMA COMMA.
Ocinara comma, Hutton, Trans. Ent. Soc., 1865, p. 330.
Leucoma comma, Hmpsn. (part), Moths, India, 1, p. 487
(1892).
2°, Masuri, including the type. 1 9, Darjiling.
LEUCOMA AURIFRONS.
Euproctis aurifrons, Moschler, Abh. Senck. Ges., xv, p. 75,
f. 3 (1887).
1 2, Eb Urru, British E. Africa (Betton).
Moschler’s type, a male, came from Aburi on the Gold
Coast. Though he called it a Huproctis the venation of his
figure is that of a Lewcoma; and as his description corre-
sponds exactly with the female Lewcoma from Eb Urru I
feel sure my identification is correct, though the localities
are very far apart; but we have many instances of Kast
and West Africa having the same forms of moths.
Genus CREAGRA, Wallengr., K. Vet. Akad. Handl., (2), v,
(4), p. 38 (1865).
CREAGRA DEALBATA.
Liparis dealbata, Herr.-Schaff., Ausser-Eur. Schmett, i, f.
111 (1854).
Creagra dealbata, Wallengr., |. ¢.
Creagra dealbata, Kirby, Cat. Moths, i, p. 460 (1892).
12 ¢,1 2,South Africa. 2 $, 2 9, Knysna.
386 Colonel C. Swinhoe’s Revision of the
Genus CaLruRA, Moore, P. Z.S., 1879, p. 401.
CALTURA ALBA.
Caltura alba, Moore, 1. c.
Caltwra alba, Moore, Lep. Ceylon, ii, p. 98, pl. 114, f.
2, 2a, b (1883).
Cispia alba, Hmpsn., Moths, India, i, p. 493 (1892).
3 gf, 1 92, Ceylon.
The type, from Ceylon, is in Mus. Dublin.
CALTURA FLAVIPES.
Cispia flavipes, Hmpsn., Moths, India, i, p. 493 (1892).
1 9, Sikhim (type).
CaLTURA PUNCTICILIA.
Naxa puncticilia, Moore, P. Z. 8., 1872, p. 575.
Cispia puncticilia, Hmpsn., Moths, India, i, p. 493 (1892).
Caltura puncticilia, Swinhoe, Cat. Het. Mus. Oxon., 1, p.
204 (1892).
1f,1 9, Nilgini Hills (co-types).
The types from the Nilgiri Hills, both males, are in
Mus. Oxon.
Genus Naroma, Walker, vii, 1744 (1856).
Hysibada, Walker, xxxii, 497 (1865).
Zarfa, Walker, Proc. Nat. Hist. Soc. Glasg., 1, p. 338
(1869).
NAROMA SIGNIFERA.
Naroma signifera, Walker, vii, 1744 (1856).
Naroma signifera, Kirby, Cat. Moths, i, p. 716 (1892).
Hysibada varipes, Walker, xxxii, 498 (1865).
Zarfa lunifera, Walker, Proc. Nat. Hist. Soc. Glasg., 1, p.
339.
1 g, Sierra Leone (type). 2 ¢, Accra. 2 J, Old
Calabar. 1 3,2 2, Natal, including type of varies. 19,
Congo (type lunifera). 5 f, 2 9, South Africa. 1 Z,
Uganda. 14,8. Nigeria.
Genus Hrmata, Moore, Lep. Atk., i, p. 57 (1879).
HIMALA ARGENTEA.
fiedoa argentea, Walker, iv, 827 (1855).
Old World Lymantriide in the National Collection. 38T
Himala argentea, Moore, 1. e.
Himala argentea, Butler, Ill. Hep., v, p. 49, pl. 89, f. 6 |
(1881).
Dasychira ilita, Moore, Cat. Lep. KE. I. C., u, p. 341
(1859).
1 ¢, Kangra. 1 gf, Dalhousie. 1 ¢, Dehra Doon.
1 g¢,1 , Darjiling, including Moore’s type 9. 1 4%,
Assam (type).
Genus GAZALINA, Walker, xxx, 398 (1865).
Oligoclona, Felder, Reise Nov., pl. 94, f. 10 (1868).
GAZALINA APSARA.
Dasychira apsara, Moore, Cat. Lep. E. I. C., u, p. 341
(1859).
Gazalina venosata, Walker, xxx1i, 398.
Oligoclona nervosa, Felder, |. c., pl. 95, f. 8.
1 9, Subathu. 1, Dalhousie. 1 ¢, Dharmsala. 1 9,
North India (type). 1 3,19, N.E. Himalayas. 5 ¢, 7 9,
Sikhim, including Walker’s type.
GAZALINA CHRYSOLOPHA.
Liparis chrysolopha, Kollar, Hiig. Kasch., iv, p. 470 (1844).
Dasychira antica, Walker, iv, 867 (1855).
Gazalina antica, Butler, Ill. Het., v, p. 49, pl. 89, f. 4
(1881).
Oligoclona chordigera, Felder, Reise Nov., pl. 94, f. 10
(1868).
1 g,N. India (type anizca). 1 2, Dalhousie. 1 ~,1 9,
Murree. I ¢, 1 $, Dharmsalay I 9, Kangra. 8 7,
2 ¢, Sikhim. I f¢, 1 2, Washan. 1 $, Chang Yang.
1 ¢, Pu-tsu-fong. 1 ¢, Kwei-chow.
GAZALINA INTERMIXTA.
Gazalina intermixta, Swinhoe, Ann. Mag. N. H. (7), vi, p.
306 (1900).
1 Z, Jaintia Hills.
A very distinct form; the wings are much shorter than
the wings of chrysolopha, the bands more erect, the inner
band almost upright and not oblique; I have a nice series
of both sexes.
388 Colonel C. Swinhoe’s Revision of the
Genus IVELA, nov.
Palpi short, covered with hair, last joint depressed ; abdomen
cylindrical, of male slender, just reaching the end of hind-wings in
both sexes; antennz bipectinate, the pectinations ciliated ; fore-
wings with the costa slightly arched, apex rounded, outer margin
convex, not oblique, hinder margin slightly rounded, nearly as long
as the costa, making the wing somewhat triangular ; vein 2 from
middle of cell, 3 from one-sixth before end, 4 and 5 from end,
discoidal angled inwards, 6 from upper end, 7, 8, and 9 stalked
from end of cell, 10 from one-third before end, 11 from a little
beyond the middle. Hind-wings rounded, 2 from beyond middle, 3
from one-sixth before end of cell, 4 and 5 from lower end, their
origins not quite touching, discoidal with the lower arm produced
and angled, its upper portion erect, 6 and 7 on a short stalk.
Type, Zvela auripes (Leucoma), Butler, from Japan.
IVELA AURIPES.
Leucoma auripes, Butler, Ann. Mag. N. H. (4), xx, p. 402
(1877).
Leucoma awripes, Butler, Ill. Het., u, p. 9, pl. 24, f. 1 (1878).
Sitvia denudata, Swinhoe (nec Walker), Cat. Het. Mus.
Oxon), p: 20214892).
S $, 5 9, Japan, including the type. 1 ¢, Omeishan,
W. China.
Genus STRACENA, nov.
Q. Palpi short, upturned, and hairy; antenne bipectinate with
cilia and fine spines at the ends of the branches ; fore-wings long,
rather narrow, costa arched before apex, which is rounded, outer
margin curved, oblique, hinder angle rounded, hinder margin nearly
straight ; vein 2 from near middle of cell, 4 and 5 from lower angle,
3 from half-way between 2 and 4, discoidal angled inwards, 6 from
upper angle of cell, 7, 8, 9, and 10 stalked, 11 from one-third before
end of cell ; hind-wings with the outer margin rounded ; vein 2 from
one-third before end of cell, 3, 4, and 5 from end, but not touching
each other at their origin ; discoidal having its lower half produced
and angled, its upper portion erect ; vein 6 and 7 from upper angle.
Type, S. fuscivena, nov.
STRACENA FUSCIVENA, nov.
2. Antenne, palpi, and legs black; head, frons, pectus, and
shoulders ochreous ; thorax, abdomen, and wings dull white, tinged
slightly with grey, the costa of fore-wings tinged with ochreous ;
Old World Lymantriide in the National Collection. 389
costal line, the median vein, discoidal, and veins 1, 2, 3, 4,5, 6 and
7 brown ; hind-wings with the veins grey near the outer margin ;
both wings are semi-hyaline, and have very minute grey irrorations ;
no other markings.
Expanse of wings 23 inches.
Hab. River Nicsr, Sapele, 5 2 (Sampson).
STRACENA PROMELENA.
Sulychra promelena, Holland, Ent. News, Phil. iv, p. 61,
jolla ae IDNs)
1 2 from R. Gaboon.
Genus SAPELIA, nov.
d, ¢. Palpi upturned and hairy, very short in the female ;
antenne bipectinated, with cilia and fine spines at the ends of the
branches ; fore-wings with the costa nearly straight but rounded
before apex, which is also rounded, the outer margin curved, not
very oblique, but nearly as long as the costa and rounded at the
hinder angle, the hinder margin also rounded ; vein 2 from middle of
cell, curved hindwards in its middle, 4 from angle of cell, 3 from
half-way between 2 and 4, 5 from a little above the angle, discoidal
angled inwards, 6 from upper angle, 7, 8 and 9 stalked, 10 from just
before end of cell, 11 from one-third; hind-wings rounded, vein 2
from a little beyond middle of cell, 3 from a little before end, 4 and
5 from the end, but their origins not quite touching, discoidal angled
inwards, 6 and 7 stalked, the stalk about half the length of the
veinlet.
Type, Sapelia limpida, nov.
SAPELIA LIMPIDA, nov.
¢, ¢. Antenne black, body above and below, legs and wings
pure white; wings of the male very thinly clothed, of the female
hyaline ; costa of fore-wings and veins of both wings more or less
covered with ochreous scaling, making them somewhat prominent ;
the male has a grey dot at the lower extremity of the end of the cell ;
no other markings.
Expanse of wings ¢ 1,%, 2 2:°; inches.
Hab. River NIGER, Sapele (Sampson).
SAPELIA FLAVIPECTUS, nov.
9. Like limpida, but the antenne are only blackish at the tips,
the rest being yellowish ; the head, frons, entire body below, and
390 Colonel C. Swinhoe’s Revision of the
femora are bright yellow, the tibize and tarsi white ; the veins of the
wings are more prominent and more ochreous, and veins 6 and 7 of
the hind-wings are on a long stalk ; in the other forms of this genus
the stalk is short.
Expanse of wings 24 inches.
Hab. SAPELE, Niger River (Sampson).
Genus Sitvia, Walker, xxxu, 387 (1865).
Kettelia, Butler, Trans. Linn. Soc. Lond. (2), 1,
p- 560 (1879).
SITVIA DENUDATA.
Sitvia denudata, Walker, xxxui, 388.
Kettelia lowii, Butler, |. c.
1 f#, Malacca. 1 9, Penang. 5 ¢, 1 2, Borneo, in-
cluding type of lowit.
Walker’s type from Malacca is in Mus. Oxon.
Genus OLaPa, Walker, iv, 823 (1855).
Antiphella, Walker, vii, 1743 (1856).
OLAPA FLABELLARIA.
Phalena jlabellaria, Fabr., Mant. Ins., 11, p. 188 (1787).
Liparis erocicollis, Herr.-Schaft., Ausser-Kur. Schmett., 1, f.
110 (1854).
Olapa temperata, Walker, iv, 823.
Antiphella vecontia, Druce, Ann. Mag. N. H. (7), ili, p. 469
(1899).
7 g, 10 2 from Abyssinia, Knysna, Natal, and the Cape.
Druce’s type came from E. Africa; it is smaller than usual,
but one of the Natal examples is still smaller.
OLAPA ARGENNA.
Cypra argenna, Mab., Ann. Soc. Ent. Fr. Ivin, p. 725
(1899).
1 gf, Madagascar.
Mabille does not give the sex of his type, neither does
he give the measurement, but his description exactly
corresponds with this example; it isa true Lymantrid, and
not a Cypra, Boisd. = Cozistra, Walker, the type of which,
delicatula, Boisd., is a Boarmid, as shown in my Memoir
on the Geometers in the National Collection, Trans. Ent.
Soc. Lond., 1902, p. 633.
Old World Lymantriide in the National Collection. 391
OLAPA (?) ATRINOTATA.
Antiphella atrinotata, Butler, P. Z. S., 1896, p. 854, pl. 43,
it,
1 ¢, Nyassaland (type).
This species does not appear to belong to this genus,
but as Sir George Hampson is writing a paper on the
moths of Africa south of the Zambesi, I leave it to him to
determine its genus.
Genus Ocoa, Walker, vii, 1768 (1855).
OGOA SIMPLEX.
Ogoa simplex, Walker, vii, p. 1764, 2.
1 g,1 2, Natal, including the type. 3 2, Brit. H. Africa.
Genus Cropera, Walker, iv, 825 (1855).
CROPERA TESTACEA.
Cropera testacea, Walker, iv, 826, 2.
1 g,2 2, But. Es Africas 5: #)6) 2 Nataltand Cape
Colony, including the type.
CROPERA ADSPERSA.
LIiparis adspersa, Herr.-Schaff., Ausser-Eur. Schmett., f.
109 (1854).
Laelia prolixa, Wallengr., Wien. ent. Mon., iv, p. 162
(1860).
Gr ee Natale
CROPERA FULVINOTATA.
Olapa fulvinotata, Butler, P. Z. S., 1893, p. 678.
5 g¢, Zomba, including the type. 7 ¢,1 2,58. Africa.
Genus CrorEeMA, Walker, iv, 811 (1855).
CROREMA MENTIENS.
Crorema mentiens, Walker, iv, 811.
Cispia (2) obliqua, Walker, vii, 1734 (1856).
Euproctis ampla, Walker, xxxil, 346 (1865).
1 3,1 9, Old Calabar. 4 7,3 9, Sierra Leone, including
all three types. 1 9,8. Nigeria. 1 #, Gold Coast.
392 . Colonel C. Swinhoe’s Revision of the
Genus TopomMEsa, Walker, xxxv, 1921 (1866).
TOPOMESA SUBINANIS.
Topomesa subinanis, Walker, xxxv, 1921.
Hmpsn., Moths, India, i, p. 469 (fig.),
(1892).
2 ¢, Java, including the type. J 2, Borneo. 2 ¢,
Singapore. 2 ¢, Tenasserim.
” )
TOPOMESA DISCOLOR.
Topomesa discolor, Hmpsn., Moths, India, iv, p. 490 (1896).
2 ¢, Ceylon, including the type.
Genus CoBANILLA, Moore, Lep. Ceylon, 11, p. 120 eee)
CoBANILLA MARGINATA.
Cobanilla marginata, Moore, |. ¢., p. 121, pl. 124, f. 4.
2 ¢, Ceylon, including the type.
Genus PorTHESIA, Steph., Ill. Brit. Ent, Haust., 11, p. 65
(1829).
Chionophasma, Butler, Trans. Ent. Soc., 1886,
p. 384.
PORTHESIA SIMILIS.
Phalena similis, Fuessl., Verz. Schweiz. Ins., p. 35 (1775).
Bombyx chrysorrhea, Esper. Schmett., 1, pl. 39, f. 1, 2
(1785).
Bombyx auriflua, Hiibn., Bomb., f. 68, 69 (1800).
9 ~,5 2, Japan and Corea. 1 3,Shanghai. 3 J, 1 9,
W. China. Besides many European examples.
PORTHESIA VIRGUNCULA.
Huproctis virguncula, Walker, iv, 836 (1855).
ELuproctis marginalis, Walker, vii, 1731 (1856).
3 ¢,5 92,N. India, including both types. 2 ¢,2 $,
Kutch. 1 2, Trevandrum: 34,22, Burma. 4°35 3°92
Java. 1 ¢, Sumatra,
?
PORTHESIA PARADOXA.
Chionophasma paradoxa, Butler, Trans. Ent. Soc., 1886,
Dao, pl..9; £2, 2 Cisse):
Porthesia panabra, Turner, Tr. Roy. Soc., S. Austral., 1902,
p. 176.
2 ¢,3 2, Queensland, including the type. 2 ¢,1 $,
Bondin Island. 29, Damma Island. 1 9, Queen’s Island.
Old World Lymantriide in the National Collection. 393
Hardly separable from virguncula, but uniformly smaller.
One of the males from Queensland was received from Dr.
Turner as panabra.
PORTHESIA GALACTOPIS.
Porthesia galactopis, Turner, Tr. Roy. Soc., S. Austral., 1902,
p. 176.
2 ¢, 1 9, Queensland.
PoRTHESIA ENTHYSANA.
Porthesia enthysana, Turner, Tr. Roy. Soc, 8S. Austral.,
1902, p. 175.
1 f#, Queensland (Turner).
PORTHESIA MIXTA.
Porthesia mixta, Butler, Ann. Mag. N. H. (5), ix, p. 88
(1882).
Leucoma mixta, Kirby, Cat. Moths, 1, p. 445 (1892).
2 9, Tasmania, including the type.
PORTHESIA FIMBRIATA.
Teara fimbriata, Lucas, Proc. Linn. Soc., N.S. W., 1891,
SD:
Porthesia fimbriata, Turner, Tr. Roy. Soc., 8. Austral., 1902,
[Oe Itsy:
2 ¢, Queensland (Turner).
PORTHESIA ALIENA.
Porthesia aliena, Butler, Trans. Ent. Soc., 1886, p. 386.
Leucoma aliena, Kirby, Cat. Moths, i, p. 445 (1892).
1 g, Peak Downs (type).
PORTHESIA MELANOSOMA.
Porthesia melanosoma, Butler, Ann. Mag. N. H. (5), 1x, p. 87
(1882).
Leucoma melanosoma, Kirby, Cat. Moths, i, p. 445 (1892).
1 g, Melbourne (type).
PORTHESIA NIGRIFINIS, nov.
¢. Antenne ochreous-grey, shafts white ; head, body above and
below, legs, and wings above and below pure white, without any
markings. There is a very slight ochreous tinge on the thorax and
394 Colonel C. Swinhoe’s Revision of the
the costal and basal portions of the fore-wings ; the last segment of
the abdomen is black above and golden yellow below, and there is a
black stripe underneath.
Expanse of wings 1,% inches.
Hab. Kikuyu, Machakos to Neugia (Crawshay).
The only African Porthesia in the National Collection,
and, I believe, the first one recorded.
PORTHESIA PRODUCTA.
Huproctis producta, Walker, P. Z. 8., 1863, p. 168.
Porthesia depauperata, Mab., Comptes-Rend. Soc. Ent. Belg.,
XXlll, p. xvii (1880).
1 ¢, 4 9, Madagascar, including the type.
PORTHESIA VARIA.
Nola varia, Saalm., Ber. Senck. Ges., 1880, p. 176.
& » Kirby, Cat. Moths, 1, p. 372 (1892).
Hab. Nosst BE.
Not in B. M.
PORTHESIA PULVEREA.
Porthesia pulverea, Ampsn., Monograph Christmas Isl.,
p09; pl. OF tI 00):
8 $,6 @, Christmas Isl., including the types.
PORTHESIA IRRORATA.
Huproctis irrorata, Moore, Cat. Lep. E. I. C., ui, p. 347
(1859).
Leucoma wrrorata, Kirby, Cat. Moths, i, p. 445 (1892).
1 2, Java (type).
PORTHESIA XANTHORRHGA.
Liparis xanthorrhwa, Kollar Hugel’s Kasch., p. 470 (1844).
Luproctis subdita, Moore, P. Z. 8., 1879, p. 400.
Luproctis flavonigra, Moore, l.c., pl. 32, f. 11.
Euproctis subnigra, Moore, Lep. Atk., i, p. 48 (1879).
74, Punjab. 3 ¢, N. W. India. 14%, Sultanpore. 2 ¢,
Umballa. 1 2%, Kangra, 2 $, Nepal, including type
of flavonigra. 1 ¢, Jawar Hills. 1 3, Khasia Hills (type
submgra). 1 3, Travancore. 3 f, Ceylon, including type
of suwbdita.
Old World Lymantriide in the National Collection. 395
PORTHESIA LUTEA.
Bombyx lutea, Fabr., Syst. Ent., p. 574 (1775).
Porthesia lutea, Turner, Trans. Roy. Soc.,S. Australia, 1902,
ps live
Artaxa chrysophila, Walker, xxxii, 334 (1865).
Porthesia iobrota, Meyrick, Trans. Roy. Soc., S. Australia,
1891, p. 194.
Artaxa wbrota, Lucas, Proc. Linn. Soc., N. 8. W. (2), vii,
p. 251 (1892).
2 9,1 2, Queensland. 3 $f, Australia, including
Walker’s type. 1 2, St. Aignan, Louisiade Islands. 2 2,
Kapaur, N. Guinea.
The Fabrician type and another are in the Banksian
Cabinet, both females. Artaxa varians, Walker, doubtfully
referred to this species by Turner, and luci/uga, Lucas, have
vein 5 of the hind-wings present, and are true Luproctis.
PORTHESIA FULVONIGRA, nov.
d 2. Antenne blackish; palpi, frons, head, thorax, and fore-
wings above ochreous ; hind-wings black with the border broadly
ochreous ; abdomen black, anal tuft of female yellow ; body below,
legs and wings yellow ; both wings with the interior portions suffused
with black. The males somewhat vary; one has a great deal of
blackish suffusion on the fore-wings, another has it slightly, and
there is a melanistic variety entirely black above and below, the
wings with some yellow in the cilia, the frons, palpi, and legs yellow.
Expanse of wings ¢ 1 inch, 2 1,%; inches.
Hab. GUADALCANAR ISLAND, Solomons (Jeck), 4 2,1 9.
PORTHESIA AURANTIACA.
Porthesia aurantiaca, Hmpsn., Moths, India, i, p. 485
(1892).
1 g, Sikhim (type).
PORTHESIA STIGMATIFERA.
Porthesia stigmatifera, Hmpsn., Moths, India, iv, App.,
p. #91 (1896).
1 2, Bhutan (type).
Genus Euproctis, Hiibn., Verz. bek. Schmett., p. 159
(1818),
Lacipa, Walker, iv, 790 (1855).
Artaza, Walker, iv, 794.
396 Colonel C. Swinhoe’s Revision of the
Antipha, Walker, iv, 806.
Dulichia, Walker, iv, 809.
Lopera, Walker, iv, 919.
Arna, Walker, v, 1176 (1855).
Somena, Walker, vu, 1734 (1855).
Utidava, Walker, xxvi, 1689 (1862).
Cozola, Walker, xxx1i, 390 (1865).
Adlullia, Walker, xxxu, 392.
Themaca, Walker, xxxii, 394.
Orvasea, Walker, xxxul, 502.
Bembina, Walker, xxxul, 505.
Microgymna, Wallern., K. Vet. Akad. Handl., 2 (v), 4, p. 38
(1865).
Gogana, Walker, xxxv, 1920 (1866) (preocc.).
Cherotricha, Felder, Reise Nov., pl. 98, Erk., p: 3 (1874).
Terphothriz, Holland, Psyche, vi, p. 474 (1893).
EUPROCTIS NEGRITA.
HLuproctis negrita, Hmpsn., Moths, India, 1, p. 471 (1892).
4 f, Sikhim.
The types ¢ 2 from Sikhim are in coll. Elwes.
EUPROCTIS JOSIATA.
Orgyia josiata, Walker, xxxii, 326 (1865).
Artasxa josiata, Kirby, Cat. Moths, i, p. 453 (1892).
Cherotricha nobilis, Felder, Reise Nov., pl. 98, f. 17 (1868).
1 g, Celebes (type).
EUPROCTIS SUBNOBILIS.
Porthesia subnobilis, Snellen, Tijd. v. Ent., xxiv, p. 128
(1881).
Artaxa subnobilis, Kirby, Cat. Moths, 1, p. 455 (1892).
Artaxa simulans, Butler, Ann. Mag. N. H. (5), xi, p. 200
(1884).
1 ?, Amboina (type simulans). 2 2, Key Island.
EUPROCTIS NIGRIBASALIS, nov.
2. Palpi, head, fore part of thorax and shafts of antenne orange
colour ; branches of antenne, rest of thorax, and abdomen above and
below black ; pectus and legs pale yellow; wings nearly white,
tinged faintly with primrose; a black spot at the end of the cell on
fore-wings, and some slight blackish suffusion at the base; on the
hind-wings nearly the entire basal half is black, in which the black
Old World Lymantriide in the National Collection. 397
cell spot can be traced ; the under-side is more distinctly primrose ;
there is a black spot at the end of each cell, and nearly the entire
basal half of both wings is suffused with black.
Expanse of wings 1,%; inches.
Hab. KINA Bau.
EUPROCTIS RENOMINATA,
Charnidas wniformis, Hmpsn., Ill. Het., viii, p. 56, pl. 140,
f, 4 (1891) (preocc.).
Euproctis renominata, Hmpsn., Moths, India, i, p. 471
(1892).
1 f, Nilgiri Hills (type).
EUPROCTIS LEUCOMELAS.
Huproctis leucomelas, Walker, iv, 838 (1855).
EHuwproctis obsoleta, Walker (nec Fabr.), iv, 839.
ss zs Swinh., Cat. Het. Mus. Oxon., i, p. 192
(note) (1892).
A long series from Hobart, Tasmania; N. Holland; and
Melbourne. The Fabrician obsoleta, the type of which is
in the Banksian Cabinet, is a Lelia, and was misidentified
by Walker ; dewcomelas is a very variable insect.
EUPROCTIS DIVISA.
Huproctis divisa, Walker, iv, 836 (1855).
i. » Hmpsn., Moths, India, i, p, 471 (1892).
Te Nepal (type), gs ieee pollsimiecin peo ae
W. China.
EUPROCTIS LATIFASCIA.
Leucoma latifascia, Walker, iv, 831, 2 (1855).
Euproctis latifascia, Hmpsn., Moths, India, i, p. 472 (1892).
Huproctis antica, Walker, iv, 835, 2.
Euproctis postica, Walker, xxxii, 348, f (1865).
Euproctis abdominalis, Moore, P. Z. S., 1888, p. 398, 2.
1 gf, N. India (type postica). 1 f,1 2, Dharmsala
(type abdominalis). 29, Kangra. 1 2, 2 9, Nepal (types
antica and latifascia). 1 9, Sikhim. 1 9, Bhutan.
EUPROCTIS SUBFASCIATA.
Artaxa subfasciata, Walker, xxxii, 332 (1865).
Euproctis subfasciata, Hmpsn., Moths, India, i, p. 472
(1892).
398 Colonel C. Swinhoe’s Revision of the
2 $2 9 Sikhims- 72/4, Assam. 179; Cachan, auld) any,
Nilgiri Hills.
The type from Darjiling is in coll. Staudinger.
EUPROCTIS SIGNATA.
Liparis signata, Blanch. Jacquemont, Voy. Inde, iv,
Ins., p. 24, pl. 1, f. 7 (1844).
1 ¢, Murree. 1: ¢, Punjab Hills. 1 ¢, Skardo.
EUPROCTIS CHIONITIS.
Euproctis chionitis, Turner, Trans. Roy. Soc., 8. Australia,
1902, p: 177.
2S, S. ED Australia. plas 2) @ucensland
(Turner).
EUPROCTIS VIRGO, nov.
¢. Antenne ochreous, shafts white; head, body, and wings
above and below uniform white, rather dull in colour; anal! tuft
ochreous ; a pale greyish suffusion on the costa of fore-wings above,
and a dark brown costal stripe below, though the margin itself is
white ; fore-legs striped with brown on their inner sides, tarsi with
grey marks.
Expanse of wings 1 inch.
1 f, Mandalay (type). 4 ¢ from Katha and Thyetmyo,
Upper Burma, collected by the late Capt. E. Y. Watson,
and 1 ¢, Rangoon.
EUPROCTIS TENUIS, nov.
¢ @. Antenne, frons, head, thorax, and legs greyish-ochreous ;
shafts of antennw white ; wings white, thinly clothed ; an ochreous
costal stripe on fore-wings in the male only ; a very slight tinge of
ochreous all over both wings ; no other markings.
Expanse of wings ¢ +4, @ 1 inch.
Hab. SAMBAWA (Doherty).
There is also a male from Kapaur, N. Guinea, collected
by Doherty, and a male from St. Aignan, one of the
Louisiade Islands, collected by Meek, which do not appear
to me to be separable from the Sambawa species, only
differing in being without the ochreous tinge.
EUPROCTIS BIMACULATA.
Euproctis bimaculata, Walker, iv, 836 (1855).
ELuproctis bigutta, Walker, 1v, 837.
Old World Lymantriudx in the National Collection. 399
EHuproctis lutescens, Walker, iv, 837.
Huproctis celsa, Walker, xxxv, 1915 (1866).
19, Shanghai. 5 ¢,49,W. China. 1 3, India (type
lutescens, f). 1 g, Poona. 2 3, 1 9, Kanara, including
type bigutta, g. 19, Nilgiri Hills. 1 9, Ceylon (type).
29,1, Burma. 1 9, Philippines (type ce/sa).
EUPROCTIS ALBESCENS. nom. nov.
Euproctis immaculata, Moore, Trans. Ent. Soc., 1884, p. 358
(preoce. ).
2 9, Sikhim, including the type. 1 g, Bhutan. 1 8,
Java. 1 2, Levuka.
This immaculate white insect must be distinct from
bimaculata.
EUPROCTIS TITANIA.
Luproctis titania, Butler, Ann. Mag. N. H. (5), iv, p. 237
(1879).
- - Kirby, Cat. Moths, 1, p. 444 (1892).
1 f, 1 2, Madagascar (types).
EUPROCTIS LUNATA.
Euproctis lunata, Walker, iv, 837 (1855).
if » Dubler dl -Hety vp. 50; pl 189, £9
(1881).
1 % Subathu. 29, Kangra. 1 ¢,2 9; Umballa. 1.2,
1 9, N. India (type). 19, Bengal. 42, Poona. 2 7, 2 9,
Madras.
EUPROCTIS CONSOCIA.
Euproctis consocia, Walker, xxxu, 347 (1865).
Artaxa modesta, Schaus and Clem., 8. Leone Lep., p. 26
(1893).
1 g,1 9, Sierra Leone (type). 2 g, Brit. E. Africa. 6 ,
1 2, Machakos.
EUPROCTIS SEMISIGNATA.
Cispia semisignata, Walker, xxxii, 3856 (1865).
Artaxa citrina, Moore, Ann. Mag. N. H. (4), xx, p. 344
(1877).
Artaxa leithiana, Moore, P. Z. 8., 1879, p. 399, pl. 32, f 9
Artaxa erecta, Moore, l.c., p. 399, pl. 32, f. 6.
TRANS. ENT. SOC. LOND. 1903.—PART III. (OCT.) 27
400 Colonel C. Swinhoe’s Revision of the
Artaxa brevivitta, Moore, |. c., p. 400, pl. 32, f. 10.
19 ¢,11 Q from various parts of India, including all
the types except citrina.
EUPROCTIS ILLANTA.
Euproctis illanta, Swinhoe, Trans. Ent. Soc., 1891, p. 13
1 f, Poona, 1 2, Alibagh, Bombay (types).
A small pure white form, without any markings.
EUPROCTIS HOWRA.
Artaxa howra, Moore, Lep. Atk., 1, p. 51 (1879).
Artaxa rhoda, Swinhoe, Trans. Ent. Soc., 1891, p. 138.
Artaxa obsoleta, Hmpsn., Ill. Het., vin, p. 57, pl. 140, f. 12
(1891).
1 g, Calcutta (type). 1 ¢, N. Kanara (type rhoda).
1 g, Nilgiri Hills (type obsoleta). 1 g, Burma. 2 ¢,
Andamans,
EUPROCTIS ENDOPLAGIA.
Euproctis endoplagia, Hmpsn., Journ. Bo. N. H. Soc., xi,
p. 295 (1897).
1 f, Khasia Hills (type). 1 $, Ichang. 1 $, Moupin.
EUPROCTIS PAUPERATA.
Euproctis pauperata, Leech, Trans. Ent. Soc., 1899, p. 138.
2 £,2 2, Moupin, including the types.
EUPROCTIS ZEBOE.
Artaxa zeboe, Moore, Cat. Lep. E. I. C., ii, p. 350, pl. 9a,
f. 7 (1859).
1 g, Java (type).
EUPROCTIS SULPHURESCENS.
Artaxa sulphurescens, Moore, P. Z. S., 1888, p. 399.
5 f,Subathu. 3 2, 2 9, Kangra, including the types.
1 2, Dharmsala. 1 ¢,Sikhim. 2 ¢,Sultanpore. 1 6,
Burma.
EUPROCTIS MONTIS.
Artazxa montis, Leech, Entom., xxiii, p. 111 (1890).
2 ¢, Chang Yang, including the type. 1 2, Chia-kow-
ho (type).
Old World Lymantriide in the National Collection. 401
EUPROCTIS INCONCISA.
Artaxa inconcisa, Walker, xxxii, 332 (1865).
5 ¢, 49, Sikhim, including the type. 1 2, Sultanpore.
1 2, Khasia Hills,
EUPROCTIS INCONSPICUA.
Luproctis inconspicua, Leech, Trans. Ent. Soc., 1899, p. 133.
1 2, Kia-ting-fu (type). 1 2, Omeishan.
EUPROCTIS VENOSA,
Artaxa venosa, Moore, Lep, Atk., p. 50, pl. 2, f. 5 (1879).
1 2, Tenasserim.
The type from Sikhim is in coll. Staudinger.
EUPROCTIS FULVIPUNCTA.
Euproctis fulvipuncta, Hmpsn., Moths, India, i, p. 474
(1892).
Lopera fulvipuncta, Hmpsn., Ill. Het., ix, p. 76, pl. 159,
ff. 2-8; pl. 175, f. 14 (larva) (1893).
1 f, 4 9, Ceylon, including the types.
EUPROCTIS BIFASCIA.
Luproctis bifascia, Hmpsn., Ill. Het., viii, p. 58, pl. 141, f. 18
(1891).
1 ¢, Nilgiri Hills (type). 1 2, Travancore.
EUPROCTIS IDONEA, nov.
g. Antenne (including shafts), palpi, frons, body above and
below, and legs ochreous ; wings above and below whitish, slightly
tinged with ochreous, the fore-wings rather darker than the hind-
wings, with the costal and hinder borders, the outer marginal line
and cilia darker ochreous ; two thin transverse bands, composed of
blackish-brown atoms, antemedial and discal, both stopping short of
the costa; under-side uniform pale ochreous white, the apical
portion of the fore-wings tinged with fuscous ; otherwise both wings
are unmarked.
Expanse of wings 1,2; inches.
Hab. SHERLOCK RIveR, W. Australia (Clement); two
examples.
EUPROCTIS IMPUNCTA.
Lacipa impuncta, Butler, P. Z.8., 1898, p. 427, pl. 32, f. 6.
3 ¢,5 9, Brit. E. Africa, including the types.
402 Colonel C. Swinhoe’s Revision of the
EUPROCTIS POSTINCISA.
Huproctis (?) postincisa, Moore, P. Z. 8., 1879, p. 400, pl. 32,
ioe
2 2, N.E. Bengal, including the type.
EUPROCTIS BIZONOIDES.
Lacipa bizonoides, Butler, P. Z. 8., 1893, p. 677.
1 2, Zomba (type).
EUPROCTIS KARGALIKA.
Huproctis kargalika, Moore, Ann. Mag. N. H. (5), 1, p. 231
(1878),
= k Moore, 2d. Yark. Miss. Lep., p. 7, pl.
Piel SAG S7T 9):
ie oe varkand., ol fralso-eiumai let. eee oaiie =
Kanda 2, Purkistan, | Wea eo? Amani
The types from Yarkand are in the Indian Museum,
Calcutta.
EUPROCTIS NOBILIS.
Panthea (?) nobilis, Herr.-Schiff, Ausser-Eur. Schmett,
f 980 (18a):
Lopera nobilis, Kirby, Cat. Moths, i, p. 463 (1892).
1 $, Cape Colony. 1 9, Knysna.
EUPROCTIS GEMMATA.
Lacipa gemmata, Distant, Ann. Mag. N. H. (6), xx, p. 200
(1897).
eee? Natal:
The types, from the Transvaal, in coll. Distant.
EUPROCTIS OSTRA, nov.
2. Fulvous above and below ; legs striped with dark brown above,
tarsi brown ; fore-wings darker than hind-wings ; orange spots on
the hinder portion of the wings, showing indications of three trans-
verse bands, subbasal, antemedial, and discal ; five black spots from
near the lower end of cell in a row towards apex, the outer orange
spots connected with this row of black spots hindwards ; under-side
with the interior of the wing suffused with brown; no other
markings.
Expense of wings 1;'5 inches.
Old World Lymantiriide in the National Collection. 403
1 2, Machakos (Crawshay) (type). 4 ¢, Kibanui, Brit.
E. Africa (Crawshay).
It belongs to the group of nobilis, Herr.-Schaff., and the
male has probably got the orange bands complete, the
outer one running through the row of black spots.
EUPROCTIS QUADRIPUNCTATA.
Lacipa quadripunctata, Dew., L.-C. Akad., xlu, p. 67, pl. 3,
f. 4 (1881).
Lacipa seepunctata, Distant, Ann. Mag. N. H. (6), xx, p. 201
(1897).
Lacipa quinquepunctata, Distant, |. c. (7), 1, p. 117 (1898).
5 $,3 2, South Africa.
EUPROCTIS FLORIDA, Dov.
¢d. Antenne grey, shafts white; palpi ochreous, black at the
base ; head and thorax white, pale yellow at the sides ; abdomen
yellow, with broad black segmental bands, anal tuft yellow ; thorax
beneath and legs grey, the latter striped with black ; fore-wings
silvery white ; costal and outer marginal line yellow; two thin
transverse orange bands, antemedial and discal ; the former bent.
outwards above its middle, the latter nearly straight, from hinder
margin one-fourth before the angle to the costa near the apex ; spots
deep black and prominent, one subbasal, below the costa, another at
one-fourth, below the middle, three, with indications of a fourth, at
the end of cell, two smaller spots on the interior margin of the outer
orange band a little above its middle, and a submarginal row close to
the margin ; hind-wings pale yellowish without markings ; under-
side pale yellowish ; fore-wings suffused with brown, hind-wings
with a brown spot at end of cell.
Expanse of wings 1,3, inches.
Hab. Natrosi Puains, Kikuyu, Brit. E. Africa (Craw-
shay).
Allied to £. quadripunctata, Dewitz.
EUPROCTIS SUNDARA, nov.
¢. Antenne ochreous grey, shafts white; palpi and frons dark
ochreous orange, the former striped with black below ; head orange,
collar white; thorax orange in front ; abdomen pale orange, the
base and middle marked with grey; under-side of body and ‘legs
ochreous; tarsi brownish ; wings white, not shining; fore-wings
with the bands and spots bright orange ; bands subbasal, antemedial,
and discal, all more or less sinuous and outwardly curved ; a row of
somewhat triangular marginal spots of a darker orange than the
404 Colonel C. Swinhoe’s Revision of the
bands, three (and indications of a fourth) similarly coloured smaller
spots at the end of the cell ; hind-wings with all but a broad outer
marginal space suffused with pale blackish-brown ; under-side white;
fore-wings with all but the margins suffused with darker blackish-
brown, and the hind-wings similarly suffused at the base.
Expanse of wings 5% inch.
Hab. Katesa, Uganda, Brit. E. Africa (Betton).
EUPROCTIS GRACILIS.
Lacipa gracilis, Hopff., Peter’s Reise Zool., v, p. 430, pl.
ff. 4, 5 (1862).
1 g, Brit. E. Africa, 1 9, Lake Nyassa.
Le)
fore)
EUPROCTIS PUBESCENS, nom. nov.
Lacipa pulverea, Distant, Ann. Mag. N. H. (7), i, p. 117
(1898) (preocc.).
4 g, Natal. 1 ¢, Zululand.
Pulverea is pre-occupied, Leech having employed the
name in 1888 for a Chinese Luproctis.
EUPROCTIS PICTA.
Liparis picta, Boisd. Delagorgue, Voy. Afr. Austr., 11, p. 599
(1847).
3 ¢,Knysna. 1 f, Cape. 2 f, Natal.
EUPROCTIS DERSA.
Euproctis dersa, Moore, Cat. Lep. E.I.C., ii, p. 347 (1859).
1 3, 2 9, Java, including the types.
EUPROCTIS HOLOXUTHA.
Euproctis holozutha, Turner, Trans. Roy. Soc., 8. Australia,
1902, p. 178.
1 g, Adelaide River. 3 2, Townsville, Queensland.
Dr. Turner has not described the female; it resembles
the male, except that the last four segments of the ab-
domen have broad black bands above, and the tip of the
anal tuft is pale blackish, or dark grey ; one example has
the fore-wings dark deep orange like the male; the other
two are paler, the fore and hind-wings being almost con-
colorous; it is quite distinct from crocea, Walker.
Old World Lymantriide in the National Collection. 405
EUPROCTIS PURA, nov.
3 ¢. Palpi, frons, pectus, under-side of body, and legs bright
ochreous ; antennz, thorax, and fore-wings of a uniform clear lilac
grey ; abdomen above with the base and anal tuft ochreous, the remain-
ing portions deep black ; actually the four last segments are banded
with black, the bands in the ¢ type specimen being quite close
together, while in the others there are slight indications of ochreous
segmental lines ; the hind-wings are ochreous, without any markings
in either of the males, and with some black suffusion in the
abdominal marginal third in the female ; the under-side is pale
uniform ochreous, with no markings in the males, and black suffusion
in the female as on upper side.
Expanse of wings g 14, ? 2,1; inches.
1 #,1 9, Townsville, Queensland, bred (Dodd). 1 f,
Queensland (Raynor coll.).
I cannot find that this has been described by any one in
Australia, probably because it stands in collections as £.
edwardsii, Newman; but we have Newman’s type, and in
my opinion it is quite a different insect, being more nearly
allied to E. holoxutha, Turner; Newman’s type is well
figured in Trans. Ent. Soc., 1856, pl. 18, f. 10, and any one
comparing it with this beautiful Queensland species will
see that it is quite different.
EUPROCTIS EDWARDSII.
Teara edwardsii, Newm., Trans. Ent. Soc., 1856, p. 284, pl.
Spt. Os LO:
1 ¢, Australia (type).
EUPROCTIS OCHREA,
Gogana ochrea, Butler, Ann. Mag. N. H. (5), i, p. 459
(1878).
Nygmia ochrea, Kirby, Cat. Moths, i, p. 449 (1892).
1 f,1 9, Madagascar (types).
EUPROCTIS CROCEA.
Teara crocea, Walker, xxxii, 355 (1865).
Nyginia crocea, Kirby, Cat. Moths, 1, p. 449 (1892).
Artaxa arrogans, Lucas, Proc. Roy. Soc., Queensland, xv,
p. 140 (1900).
Type 2, Moreton Bay.
Lucas’ types came from Cairns; his description exactly
suits Walker’s type.
406 Colonel C. Swinhoe’s Revision of the
EUPROCTIS PULVEREA.
Artazxa pulverea, Leech, P. Z. S., 1888, p. 623, pl. 31, f 5.
2 Z, 2 2, Satsuma, including the types. 2 ¢, Nagasaki.
2 Z,1 2, Loo Choo Islands. 1 ¢, Gensan. 12, Kia-
ting-fu.
EUPROCTIS FLAVINATA.
Artaza fiavinata, Walker, xxxii, 331 (1865).
Luprocis flavinata, Hmpsn., Moths, India, i, p. 475 (1892).
ELuproctis recurvata, Leech, Trans. Ent. Soc., ae p. 138.
; 2,1 2, Shanghai, including the type. 2 7, Chusan.
: $, Ningpo. 3 z, Chang Yang, ers peers type.
1 2, Sarawak. 2 7,1 9°, Nilgiri Hills. , 1 2, Ceylon.
1 2, Maulmein. .
EUPROCTIS ARGENTATA.
Euproctis argentata, Leech, Trans. Ent. Soc., 1899, p. 139.
1 27, Japan (type).
EUPROCTIS CALVA, nov.
3 ¢. Antenne, palpi, head, body, and fore-wings dark bright
orange ochreous without any markings; hind-wings pale whitish
ochreous, also without any markings. :
Expanse of wings ¢ 14, 9 15 inches.
1 7, Sambawa (Doherty) (type). 1 2, Bali, Low Country
(Doherty) (type). 1 2, Lombok
The species is much like £. holoxutha, Turner, from
Australia, but that species has a pale spot at the end of the
cell of the fore-wings, the hind-wings are dark orange
ochreous instead of being whitish, and the abdomen of the
female has broad black bands.
EUPROCTIS FULVA.
Artaxza fulra, Butler, Ann. Mag. N. H. (5), x, p. 227
(1882).
1 2, Duke of York Isl (type). 1 2,N. Britain. 1 7,
Port Darwin. 1 2, Queensland.
EUPROCTIS VARIANS.
Artaza varians, Walker, iv, 796 (1855)
Euprocis pygmxa, Moore, Lep. Atk, p- 48 (1879),
(preocc.).
Old World Lymantriide in the National Collection. 407
Artaza pusilla, Moore, Lep. Ceylon, 1, p. 86, pl. 112, f. 4
(1883).
26 ¢, 14 2 from various parts of China, India and
Ceylon, including the types of varians and pygmea.
EUPROCTIS PYGMAA.
Aroa pygmea, Walker, iv, 793 (1855).
5 f, Sierra Leone, including the type. 3 ¢, Accra.
1 g, Congo. 13, Gold Coast. 1 3, Old Calabar.
EUPROCTIS SCOTOCHYTA.
Euproctis scotochyta, Turner, Tr. Roy. Soc., S. Austral., 1902,
p. 178.
1 gf, Adelaide River.
EUPROCTIS CERVINA.
Artaxa cervina, Moore, Ann. Mag. N. H. (4), xx, p. 345
GUS:
6 $,3 2, Ceylon, including the type.
EUPROCTIS CHRYSOPHEA.
Orgyia chrysophea, Walker, xxx, 324 (1865).
Notolophus chrysopheus, Kirby, Cat. Moths, 1, p. 493
(1892).
Abyssinia, type 2.
Veins 3 and 4 of the hind-wings are on an unusually
long stalk.
EUPROCTIS MAZA, nov.
¢. Primrose yellow ; body and legs without markings ; fore-wings
darker and brighter yellow ; a large round subbasal red spot near the
hinder margin ; a broad red discal band, divided by the veins into
elongated marks, acutely angled in its middle towards the outer
margin, which it does not touch ; it also stops short of the costa >
there are likewise indications of a central band of scales darker than
the ground colour of the wing, in two examples more pronounced
than in the others.
Expanse of wings 3; inch.
Four examples from Kapaur, N. Guinea (Doherty) ; it is
closely allied to £. titania, Druce, from Fergusson Isl. ;
there is a female in the collection from N. Guinea with
erange streaks between the veins, but with some of the
408 Colonel C. Swinhoe’s Revision of the
transverse markings distinguishable ; it may be the female
of this species.
EUPROCTIS DRUCEI, nom. nov.
Euproctis titania, Druce, Ann. Mag. N. H. (7), ili, p. 469
(1899), (preocc.).
2 7, Fergusson Island.
Titania is pre-occupied in the genus by Butler fora
Madagascar Luproctis.
EUPROCTIS DANA, nov.
¢. Palpi, antennz, head, body above and below, and legs brown-
ish ochreous, paler and brighter below than above ; fore-wings much
the same colour but inwardly suffused with brown, with two in-
distinct transverse pale lines beyond the middle rather close together,
forming a band which in some examples is darker than the rest of the
wing, as in E. cervina, Moore, from Ceylon ; hind-wings blackish-
brown ; cilia of both wings rather long and bright ochreous.
Expanse of wings ,{, inch.
6 ¢, Dana, Kashmir, June 1888 (WeArthu7).
Two of the examples have the fore-wings almost clear
of the brownish suftusion and the transverse pale lines
nearly obsolete, the insect thus looking very like JZ.
chrysophea, Walker, from Abyssinia.
EUPROCTIS LUCIFUGA.
Artaxa lucifuga, Lucas, Proc. Linn. Soc., N.S. W., 1892, p.
250.
Luproctis chrysophea, Turner (nec Walker), Trans. Roy.
Soc., S. Australia, 1902, p. 178.
3 ¢, Mackay, Queensland.
This is not the same as Walker’s Abyssinian species, the
type of which is now before me, and it is hardly to be
expected that it would be ; the fore-wings of chrysophea
are perfectly smooth dull orange colour, and the under-side
of both wings pale bright orange without any suffusions,
whereas Juwcifuga has the fore-wings more or less covered
with brown irrorations, which give the wings a fuscous
orange appearance, and on the under-side both wings are
dark brown with orange borders.
EUPROCTIS FULVISTRIATA, Nov.
¢ ¢. Bright orange ochreous, with darker orange streaks in the
interspaces, leaving the veins pale yellow ; two indistinct pale yellow
Old World Lymantriide in the National Collection. 409
transverse lines, medial and discal, both curving outwards; abdomen
with whitish segmental lines, the last segment entirely whitish ;
anal tuft grey ; legs with the tarsi whitish ; under-side of wings pale
ochreous yellow without any markings.
Expanse of wings ¢ 13%, 2 1y‘s inches.
Hab. St. AIGNAN Isz., Louisiade Group (Meek).
EUPROCTIS ARCLADA, nov.
2. Much larger and darker in colour than filvistriata ; the anal
end of the abdomen is narrowly white ; the legs are orange ochreous
without any white on the tarsi ; the orange stripes on the fore-wings
are similar, but the hind-wings are much darker than the fore-wings,
being of a curious vermilion orange colour.
Expanse of wings 2,2; inches.
Hab. WooDLARK ISLAND (Jeez).
EUPROCTIS APICALIS.
Bembina apicalis, Walker, xxxu, 505 (1865).
1 $,2 2, Ceylon, including the type 9.
EUPROCTIS FERVIDA.
Artaza fervida, Walker, P. Z. S., 1863, p. 168.
. 2 Saalm., Lep. Madag., p. 185, pl. 7, ff. 115,
116 (1884).
1 $, Madagascar (type).
EUPROCTIS SUBFUSCULA.
Artaxa subfuscula, Hmpsn., ll. Het., viii, p. 56, pl. 140,
ff. 5-11 (1891).
3 9,3 2, Nilgiri Hills, including the type.
EUPROCTIS TERMINALIS.
Aroa terminalis, Walker, iv, 794 (1855).
9 9, 1 9, Natal, including the types.
EUPROCTIS SQUAMOSA.
Lopera squamosa, Walker, iv, 920 (1855).
1 ¢,1 2, Natal (types).
EUPROCTIS PUNCTIFERA.
Aroa punctifera, Walker, iv, 792 (1855).
Lopera gaudens, Walker, xxxui, 357 (1865).
1 f, Caffraria (type). 7 3,19, Natal, including the type
of gaudens.
410 Colonel C. Swinhoe’s Revision of the
EUPROCTIS CROCATA.
Liparis crocata, Boisd. Delagorgue, Voy. Afr. Austr., i, p.
599 (1847).
Ss » Herr.-Schiff., Ausser-Eur. Schmett., i, f.
112 (1854).
Sot, 2 Os, Airiea:
EUPROCTIS PALLIDA.
Cropera pallida, Kirby, Ann. Mag. N. H. (6), xviii, p. 384,
Peel ost ondisoG)):
1 ¢, Mozambique (type). 1 3, Mandala. 1 ¢, Delagoa
Bay. 47,5 9, Natal. “2 ¢, Mashonaland.
EUPROCTIS MONOSTICTA.
Lopera monosticta, Butler, P. Z.S., 1898, p. 428, pl. 32, f. 7.
1 ¢, Taru, E. Africa (type). 1. gf, Machakos.
EUPROCTIS FASCIATA.
Dulichia fasciata $, Walker, iv, 809 (1855).
Artaxa squamiplaga 9°, Walker, Proc. Nat. Hist. Soc.,
Glasgow, i, p. 338 (1869).
Euproctis susanna, Staud., Iris, vu, p. 258, pl. 9, f. 9
(1894).
Euproctis torrida f, Distant, Ann. Mag. N. H. (6), xx, p. 202
897):
Euproctis stellata 2, Distant, 1. ¢.-
Twenty-two examples of both sexes from Natal, Zulu-
land, Congo (including type of sqguamiplaga), Old Calabar,
Sierra Leone (type), Aden, and Palestine.
KEUPROCTIS JONASI.
Aroa jonasi, Butler, Ann. Mag. N. H. (4), xx, p. 402
sigh):
P > Butler, TI Het. ai, p, fOM pk Za ws 78):
8 ¢, Japan, including the type.
EUPROCTIS UNIPUNCTATA.
Euproctis wripunctata, Leech, Trans. Ent. Soc., 1899, p.
136.
2 g, 5 9, W. China, including the types.
Old World Lymantriide in the National Collection. 411
EUPROCTIS TRIFASCIATA.
Artaxa trifasciata, Moore, Lep. Atk., p. 51 (1879).
1 g¢, Kangra. 2 g, Umballa. 1 fg, Darjiling. 1 Z,
Assam (type). 2 $, Cachar. 1 g, Bhamo. 1 @, Chia-
kow-ho.
EUPROCTIS DIGRAMMA.
Bombyx digramma, Guérim, Icon. R. Anim., Ins., p. 508,
pl. 86, f. 4 (1830).
12, Hone Kone, see bhutan. 1 9 Nepal 254) 3
Ceylon. 1 ¢, Burma. 1 %,Sumatra. 3 f,1 9, Java.
Kirby puts this as a synonym to flava, Fabr.; I have
examined the two specimens in the Banksian Cabinet, put
with a query as types; they are too faded and discoloured
for purposes of identification.
EUPROCTIS UNIMACULA-
Artaxa unimacula, Moore, P. Z. S., 1879, p. 399.
1 f, 2 2, Khasia Hills, including the type.
EUPROCTIS GUTTATA.
Artaza guttata, Walker, iv, 795 (1855).
1 2, N. India (type). 2 2,1 2, Dharmsala. 1 9, Kangra.
EUPROCTIS PELONA.
Artaxa pelona, Swinhoe, Trans. Ent. Soc., 1891, p. 138.
2 $, Khasia Hills, including the type.
There is a form from Ceylon, which appears to me to be
different from either guttata or pelona, but we want more
material for decision.
EUPROCTIS FRATERNA.
Artaxa fraterna, Moore, Lep. Ceylon, ii, p. 85 (1883).
2 ¢,Subathu. 1 $,Sahibgunge. 1 ¢,1 2, Bombay.
5 f, 5 9, Ceylon, including the types.
EUPROCTIS SASTRA.
Artaxa sastra, Moore, Cat. Lep. E. I. C., 11, p. 351 (1859),
2 4, Java, including the type.
EUPROCTIS INCOMMODA.
Artaxa incommoda, Butler, Cist. Ent., 111, p. 11 (1882).
1 gf, Madagascar (type).
412 Colonel C. Swinhoe’s Revision of the
EUPROCTIS COMMUTANDA, nom. nov.
Aroa immaculata, Butler, Ann. Mag. N. H. (5), x, p. 227
(1882) (preocc.).
1 2, Duke of York Island (type).
EUPROCTIS CHEELA, nov.
¢. Ochreous yellow ; palpi, antenne, head, and body dark bright
chrome yellow; branches of the antenne, thorax, and basal third of
fore-wings suffused with pale brown; hind-wings pale yellow ;
under-side whitish, just tinged with yellow, a little blackish suffusion
on fore-wings near the base on the hinder margin ; no other markings
above or below.
Expanse of wings 1,%; inches.
Hab, SINGAPORE (fid/ey).
EUPROCTIS MUNDA.
Euproctis munda, Walker, Journ. Linn. Soc. Lond., vi, p.
129 (1862).
2 gf, Sarawak, including the type. 1 9, Nias. 1 f,
Singapore.
EUPROCTIS CIVITTA.
Euproctis civitta, Swinhoe, Ann. Mag. N. H. (7), xii, p. 195
(1903).
1 f, Kuching, Borneo.
EUPROCTIS LODRA.
Euproctis lodra, Moore, Cat. Lep. E. I. C., 1, p. 349, pl. 9a,
f. 6 (1859).
2 4 semper, Het. Philipp., p. 471 (1898).
1 9, Java (type).
EUPROCTIS INTENSA.
Artaza intensa, Butler, Ann. Mag. N. H. (4), xx, p. 402
(1877).
Butler, Il, Het! 11, p. 10) pl 23, f£ 12
(1878).
6 4 2 9, Central China, 6356 2% Japan. 5 ¢) 2-9,
Corea.
>P) ”?
EUPROCTIS VITELLINA.
Tiparis vitellina, Kollar, Hugel’s Kasch., iv, p. 471 (1844).
Euproctis gamma, Walker, vii, 1731 (1856).
Old World Lymantriide in the National Collection. 413
Artaxa princeps, Walker, xxxui, 331 (1865).
Themaca comparata, Walker, xxxu, 395.
3 2, 1 2, Murree, 4 2, Dalhousie. 1 2, Dharmsala.
26,3 9, Kanera. 2 ¢, 3 9, N. India, including all
Walker’s types. 1 3, 2 9, Sultanpore.
EUPROCTIS LIMONEA,
Cherotricha limonea, Butler, Cist. Ent., i, p. 11 (1882).
1 ¢, Madagascar (type).
EUPROCTIS STRAMINEA,
Euproctis straminea, Leech, Trans. Ent. Soc., 1899, p. 135.
1 2, Chia-kow-ho (type). 1 2, Omeishan (type).
EUPROCTIS CONSPERSA,
Artaxa conspersa, Butler, Cist. Ent., ii, p. 117 (1882).
Huproctis conspersa, Leech, Trans. Ent. Soc., 1899, p. 140.
5 ¢, 7 2, Japan, including the type.
EUPROCTIS INEPTA,
Artaxa inepta, Butler, Ann. Mag. N. H. (5), xix, p. 228
(1887).
1 g, 2 9, Alu, including the types.
EUPROCTIS ANGULIGERA.
Artaxa anguligera, Butler, P. Z.8., 1886, p. 385.
1 f, Murree (type). 1 ¢, Andamans.
EUPROCTIS AMPHIDETA,
Euproctis amphideta, Turner, Trans. Roy. Soc., S. Australia,
QO 2F pele
1 2, Townsville, Queensland.
Turner’s description is not quite clear, but I think I have
properly identified the insect, and the size and locality
agree ; moreover, as Turner says, this species has vein 7 of
the fore-wings absent: but in the specimen before me the
grey portion of the fore-wing is limited by a sordid pink
band, which runs along the costa with a spot in its centre,
then submarginally to the hinder margin, having a tooth
outwards in the centre, not at “two fifths of termen” as
Turner says.
414 Colonel C. Swinhoe’s Revision of the
EUPROCTIS PECLA, nov.
¢. Antenne, palpi, head, thorax, and fore-wings dark bright
yellow ; two bands composed of blackish-brown, minute scales ; first
before the middle, interrupted but erect, consisting of a small piece on
the hinder margin and a spot in the cell ; the other discal, also erect,
but dentated above the middle, with a prolonged tooth that reaches
the outer margin ; abdomen, hind-wings, and under-side whitish,
slightly yellow-tinged, with no markings,
Expanse of wings 1 inch.
Hab. Mune Bay, New Guinea (Meek).
EUPROCTIS LIVIA, nov.
do. Pale yellow ; palpi ochreous ; antennz grey ; head and body
pale yellow ; fore-wings with chestnut grey bands covering nearly
the whole surface ; the first occupying more than the basal third,
divided from the second by a thin yellow outwardly curved band ; the
second narrower, also with a narrow yellow outer margin, which is
acutely bent outwards above the middle ; there is a yellow spot on
the costa, attached to it, and the outer portion of the wing is grey on
the costa and at apex, and thinly on the outer margin ; its inner portion
yellow, with two prominent black spots at the end of the angle of the
outer margin of the second band, these spots being joined together by
a line of black scales ; cilia pinkish-yellow ; hind-wings and under-
side pale yellow, nearly white, without markings.
Expanse of wings 3% inch.
Hab. MILNE Bay, New Guinea (Jeek).
KUPROCTIS BASALIS.
Artaxa basalis, Moore, Lep. <Atk., p. 51, pl. 2, f. 16
(S79):
1 2, Khasia Hills.
The type from Dayjiling is in coll. Staudinger.
EUPROCTIS BIDENTATA.
Huproctis bidentata, Hmpsn., Journ. Bo. N. H. Soe., xi, p.
296 (1897).
1 ¢, Darjiling (type). 1 2, Yatong, Sikhim.
EUPROCTIS GENTIA.
Huproctis gentia, Swinhoe, Ann. Mag. N. H. (7), xii, p. 195
(1903).
1 2, Kina Balu (type).
Old World Lymantriide in the National Collection. 415
EUPROCTIS SEXMACULA.
Euproctis seemacula, Swinhoe, Ann. Mag. N. H. (7), xii
p. 195 (1903).
1 2, Kina Balu (type).
EUPROCTIS MIRABILIS.
d, @. Antenne, palpi, frons, and head pale ochreous; palpi
brown above ; thorax brown; abdomen ochreous; fore-wings with
the ground colour pale ochreous, but the wing is entirely covered
with dark brown scales, except at the costa and outer margin ; the
latter in the male is broadly yellow, with two short projections
into it from the brown portion, but in the female these projections
are broad and run into the outer margin ; there is also a transverse
ochreous thin band through the brown portion a little before the
middle of the wing, which contains a large ochreous spot above its
centre; abdomen of the female brown, anal tuft ochreous ; hind-
wings and under-side pale ochreous without any markings.
Expanse of wings ¢ 1 inch, 2 1; inches.
Hab. ANDAMAN ISLANDS.
EUPROCTIS UTILIS, nov.
¢, 2. Palpi brown above, white beneath; frons whitish with
some ochreous hairs above the palpi ; head, thorax, and fore-wings
pale yellow ; two broad transverse bands, medial and sub-marginal,
composed of blackish scales, not very thickly packed together and
divided by the veins; abdomen whitish, anal tuft fulvous ; hind-
wings, both wings below, body, and legs pure white, without any
markings.
Expanse of wings ¢ 1,4, @ 14% inches,
Hub. OLD CALABAR (Crompton).
Corresponds somewhat to Holland’s description of his
Artaxa melaleuca from the Ogove River, but his insect, a
male, measures only 15 mm. and has a basal band.
EUPROCTIS SUBFLAVA.
Aroa flava, Brem., Bull. Acad. Sci. Pet., ili, p. 479 (1861)
(preocc.).
Arou subflava, Brem., Lep. Ost. Sib. p. 41, pl. 3, f 19
(1864). ;
Leucoma subflava, var. piperita, Oberth. Et. d’Ent., v, p. 35
(1880).
TRANS. ENT. SOC. LOND. 1903.—PART Ill. (OCT.) 28
416 Colonel C. Swinhoe’s Levision of the
Porthesia snelleni, Staud., Rom. sur Lep., 11, p. 207, pl. 12,
f. 3 (1887).
3
6 2 9. Japan. 2d aw Ching,
EUPROCTIS ATROSQUAMA.
Goyana, atrosquama, Walker, xxxv, 1921 (1866).
Euproctis atrosquama, Semper, Het. Philipp., p. 464, pl. N,
ff. 11-13 (1898).
Cherotricha glandulosa, Felder, Reise Nov., pl. 98, f. 14
(1868).
1 ¢, Philippines (type).
EUPROCTIS PLANA.
Euproctis plana, Walker, vu, 1731 (1856).
Cherotricha plana, Butler, Ill. Het., v, p. 51, pl. 89, f. 13
(1881).
Euproctis discinota, Moore, P. .Z. 8., 1877, p. 601.
i ?, Houg Kone, 3 3,1 2, Dharmsalas aia; toe
Kanera. 2 $,1 2, Darjiling, including the type. 1 9,
Burma. 2 9, Omeishan, W. China. 1 Jf, 1 9, Andamans
(type discinota).
EUPROCTIS IMMACULATA.
Cherotricha immaculata, Butler, Ill. Het., v, pl. 89, f. 14
(1881).
EHuproctis immaculata, Leech, Trans. Ent. Soc., 1899, p. 137.
1 2, Darjiling (type).
There is no more constant character in the species of
this genus than the cell spot, and therefore 1 cannot
believe that this immaculate insect with white hind-wings
is the same as plana, as Hampson says.
EUPROCTIS CATALA, nom. nov.
Euproctis atomaria, Walker, iv, 837 (1855) (preocc.).
a . Moore, Cat. Lep. E. I. C., ii, p. 347,
pl.16, f. 1, la Garva) (1859).
. " Snellen, Tijd. v. Ent. xx, p. 11, f. 4, g
(1877):
Euproctis miilleri, Snellen, 1. c., f. 6, 2 (only).
6 $,5 2, Java, including the types.
Horsfield bred this species, and we have his examples
in the National Collection; therefore I feel no doubt
Old World Lymantriide in the National Collection. 417
that Snellen’s figure of his female mii/lert represents this
species; I have not been able to identify his male. The
name atomaria is preoccupied, Walker himself having
used it at p. 796 of the same vol. under his genus Artaza,
which is the same as Huproctis,
EUPROCTIS ANNA.
Huproctis anna, Swinhoe, Ann. Mag. N. H. (7), xii, p. 194
(1903).
1 ¢, 1, Kina Balu (types).
EUPROCTIS ORESTES,
Cherotricha orestes, Druce, P. Z. 8., 1887, p. 674.
ee, Aburi.n 274) eo Goldi@oast-
The type from Mongo-ma-Lobah is in coll. Druce.
EUPROCTIS MADANA.
Euproctis madana, Moore, Cat. Lep. E. I. C., 11, p. 348
(1859).
1 2, Darjiling (type).
EUPROCTIS ICILIA.
Bombyx icilia, Stoll, Suppl. Cram., p. 158, pl. 35, f. 5
(L791).
Cherotricha decussata, Moore, Ann. Mag. N. H. (4), xx,
p. 345 (1877).
x ” Moore, Lep. Ceylon, ui, p. 91, pl. 113,
f, 2, 2a, b (1883).
Zo, Wurbuncass to Kanwar g.Malabara| 179
Trevandrum. 1 ¢, 2 @, Ceylon, including the type of
decussata,
EUPROCTIS STIRASTA.
Adlullia stirasta, Swinhoe, Ann. Mag. N. H. (6), xii, p. 214
(1893).
1 ¢, Mone, Shan States (type).
I have a female from the same locality in my own
collection.
EUPROCTIS BARBARA.
Huproctis barbara, Swinhoe, Ann. Mag. N. H. (7), xii, p.
197 (1903).
1 g, Kuching, Borneo (type).
418 Colonel C. Swinhoe’s Levision of the
EUPROCTIS STAUDINGERI.
Cherotricha staudingert, Leech, P. Z. 8., 1888, p. 624, pl. 31,
i, 0:
5 f, 5 9, Japan, including the types.
EUPROCTIS VARIA.
EHuproctis (2) varia, Walker, iv, 840 (1855).
Moore, Cat. Lep. E. I. C., 1, p. 348,
pl. 9a, f. 5 (1859).
1 ¢, Subathu. 37,12, Sikhim, 1 9, Khasia Hills.
The type, without locality, is in Mus. Oxon.
” dy)
KUPROCTIS OREOSAURA.
Adlullia oreosaura, Swinhoe, Ann. Mag. N. H. (6), xiv,
p. 435 (1894).
1 ¢, 1 9, Khasia Hills (types).
There is a small male from Kuching, Borneo, which
much resembles this species.
EUPROCTIS NIPHONIS.
Cherotricha niphonis, Butler, Trans. Ent. Soc., 1881, p. 9, 2.
Cherotricha squamosa, Butler, 1. ¢., 9.
Porthesia raddei, Staud., Rom. sur Lep., iii, p. 207, pl. 17,
f. 3 (1887).
7 $, 5 9, Japan, including both Butler’s types. 2 2,1 9,
Amur.
EUPROCTIS PLAGIATA,
Cispia plagiata, Walker, iv, 858 (1855).
1 9, Nepal (type). 1 ¢, Mount Parisnath. 1 2, N.
India.
EUPROCTIS MARGINATA,
Cherotricha marginata, Moore, Lep. Atk., p. 49 (1879).
Cherotricha quadrangularis, Moore, |. ¢., p. 50, pl. 2, f. 25.
2 $, 2 9, Darjiling, including the type. 1 f, Kurse-
yong.
The type of quadrangularis, from Manipur, is 1n coll.
Staudinger.
Old World Lymantriide in the National Collection. 419
EUPROCTIS PRACURRENS.
Adlullia precurrens, Walker, xxxui, 392 (1865).
Cherotricha globifera, Felder, Reise Nov. Lep., pl. 98, f. 13,
@ (1868).
Nygmia globifera, Kirby, Cat. Moths, i, p. 449 (1892).
Euproctis guttulata, Snellen, Notes Leyden Mus,, viui, p. 7
(1886). .
Snellen, Tijd. v. Ent, xxix, p. 36, pl. 1,
f. 3 (1886).
Adlullia boleora, Swinhoe, Cat. Het. Mus. Oxon., i, p. 186,
pl. 6, £2, % (1892).
4 2,52 from Dinding, Sarawak, Kuching, and Singa-
pore; Felder’s type came from the Moluccas, and is
undoubtedly a female, though figured as a male; the types
of precurrens from Celebes and of boleora from Sarawak
are in Mus. Oxon.
”? »?
EUPROCTIS PALLIPES.
Euproctis pallipes, Snellen, Tijd. v. Ent., xxii, p. 108, pl. 9,
if, 8) (ESi7/8)h
1 2%, Celebes.
EUPROCTIS OSUNA, nov.
2. Dull ochreous tinged with chestnut colour ; fore-wings long,
costa arched, outer margin nearly as long as the costa, much rounded,
the curve taking in the short hinder margin to the base ; two macular
transverse chestnut-coloured bands, discal and marginal; the first
consisting of five or six square spots ; the other complete, the spots
well separated ; indications of another band towards the base, where
there is a small group of black irrorations ; all the spots covered with
these irrorations, as also are the cilia of both wings, the antecilial line
being pale; abdomen and abdominal half of hind-wings suffused
with black ; under-side much as above.
Expanse of wings 2 inches.
Hab. Kapaur, New Guinea (Doherty).
Somewhat like #. (Cherotricha) armandvillei, Oberth.,
Etud. Ent., xix, p. 35, pl. 5, f. 31 (1894).
EUPROCTIS EGREGIA, nov.
Q. Palpi ochreous with some brown hairs; head and thorax
ochreous; abdomen blackish-brown, tip ochreous; fore-wings
blackish-brown, with a very broad orange ochreous outer marginal
420 Colonel C. Swinhoe’s Revision of the
border, occupying quite one-fifth of the wing ; hind-wings with the
basal third black, the outer portion orange ochreous; cilia of both
wings ochreous ; under-side of body and legs blackish-brown ; wings
as above, but the brown part darker.
Expanse of wings 2;°5 inches.
Hab. BATJAN.
EUPROCTIS SERVILIS.
Huproctis servilis, Walker, xxx, 350 (1865).
Artaxa servilis, Swinhoe, Cat. Het. Mus. Oxon., i, p. 190
(1892).
Darala prima, Walker, xxxv, 1917 (1866).
Euproctis incompta, Snellen, Tid. v. Ent., xx, p. 9, pl. 1, f. 2
(1879).
Euproctis cinerea, Heylaerts, Ann. Soc. Ent. Belg., xxxvi,
p- 10 (1392).
Euproctis nurma, Druce, Ann. Mag. N. H. (7), m1, p. 469
(1899).
3 $,19, Java. 12, Sambawa (Doherty). 1 %, Celebes
(Doherty).
Type Celebes (Wallace) in Mus. Oxon., as is also the
type of prima from the same locality; Snellen’s and
Heylaert’s types came from Java, Snellen also records it
from Celebes; Druce’s type came from Timor. The colour
of fore-wings varies much, from very pale yellow to olive-
brown, and the hind-wings from yellow to white. I have
before me the two extremes from the same locality: the
female has all yellow fore-wings, white hind-wings, without
markings.
EUPROCTIS UNIFORMIS.
Cherotricha wniformis, Moore, Lep. Atk., p. 49 (1879).
1 2, Darjiling.
The type from Darjiling is in coll. Staudinger.
EUPROCTIS CALESIA.
Huproctis calesia, Swinhoe, Ann. Mag. N. H. (6), ix, p. 81
(1902).
1 9, Lawas, Borneo (type). 1 9, Sandakan.
Old Worid Lymantriide in the National Collection. 421
EUPROCTIS POSTNIGRA, nov.
@. Antenne, palpi, head, and thorax brownish-ochreous ; fore-
wings with the ground colour sordid ochreous, irrorated with minute,
deep black atoms; a whitish space at the end of the cell, in which
there is a somewhat large black spot ; a discal, indistinct, transverse
whitish band, outwardly curved above its middle; abdomen and
hind-wings black, anal tuft and cilia of both wings sordid ochreous ;
under-side of body and legs black ; wings blackish-brown, with the
margins narrowly sordid ochreous.
Expanse of wings 2,7; inches.
1 9, Matang, Borneo (type). 1 2, Pulo Laut (Doherty).
T have it in my own collection from Pulo Laut.
HUPROCTIS INDECORA.
Teara indecora, Walker, xxxu, 353 (1865).
1 2, Moreton Bay (type).
EUPROCTIS DEFICITA.
Teara deficita, Walker, xxxil, 352 (1865).
3 f, 2 9, Australia, including the type.
EUPROCTIS FUNERALIS, nov.
d¢, @. Palpi, frons, and anal tuft of abdomen ochreous, paler in
some examples than in others ; antennze ochreous grey, shafts white ;
head, thorax, and fore-wings ochreous, the ground colour of the fore-
wings is ochreous grey, and the wings are covered with brown,
minute irrorations ; one Sarawak male example has the fore-wings
much darker than the others, and a Sarawak female is the palest and
yellowest of the lot; the hind-wings of the ¢ type are nearly white,
the abdominal margin broadly suffused with blackish ; in the female
they are entirely blackish-brown with pale yellowish outer margins ;
abdomen above and below black; legs yellowish; wings below
whitish in the male with blackish internal suffusions, entirely
suffused in the female. The species varies much in size and in shade
of colour; in some of the females the veins of the fore-wings are.
yellow and prominent.
Expanse of wings ¢ 15%, 2:'s inches.
1 g,1 2, Singapore (types) (fidley). 4 9, Singapore.
1 ¢, 12, Sarawak. 1 2, Penang (Plower). 1 2, Java.
Was many years ago wrongly identified in the B. M. as
conspersa, Felder, Reise Nov. Lep., pl. 98, f. 12, but has
very little resemblance to it ; conspersa is not in the B. M.
422 Colonel C. Swinhoe’s Levision of the
EUPROCTIS FLAVOCILIATA.
Euproctis flavociliata, Swinhce, Ann. Mag. N. H. (6), vii,
p. 465 (1901).
1 9, Perak (type). 12, 1 9, Singapore.
EUPROCTIS TRANSVERSA.
Artava transversa, Moore, Cat. Lep. E. I. C., 11, p. 352, pl. 9a,
f. 8 (1859).
ss Ws Swinhoe, Cat. Het. Mus. Oxon., 1, p. 188
(1892).
Huproctis guttistriga, Walker, Journ. Linn. Soc. Lond., vi,
p. 129 (1862).
1 ¢,3 2, Singapore. 3 2, Sumatra. 1 dg, 1 9, Borneo.
2 2, Java, including the type.
Walker’s type from Sarawak is in Mus. Oxon.
EUPROCTIS LEUCOSPILA.
Cozola leucospila, Walker, xxx, 390 (1865).
Cherotricha leucospila, Felder, Reise Nov., pl. 98, f. 16
(1868).
1 2, Celebes (Doherty).
The types, from Celebes, are in Mus. Oxon. and coll.
Rothschild.
EUPROCTIS CHIRUNDA, nov.
?. Palpi, frons, head, and fore-part of thorax yellow ; thorax and
fore-wings dark chocolate brown ; costal margin yellow ; a patch of
that colour in the middle extending to the end of the cell, where it
is rounded, and contains a large black spot ; a square yellow patch on
the outer margin above the middle, and another at the hinder angle ;
abdomen black, with pale segmental lines and yellow anal tuft; hind-
wings blackish-brown, yellow on the outer margin; cilia of both
wings yellow ; under-side, body and legs yellow, wings blackish-
brown, with the yellow marginal marks much as above.
Expanse of wings 2,5 inches.
Hab. SANDAKAN.
EUPROCTIS PERPLEXA, nov.
2. Antenne, palpi, head, and tip of abdomen ochreous ; thorax
and fore-wings dark ochreous brown, the lower part of the wings the
darker ; a large black spot of embossed scales on a pale ochreous
ground on the hinder margin a little before the middle ; another
Old World Lymantriide in the National Collection. 423
just above it, the pale margins joining ; abdomen black ; hind-wings
blackish-brown, darker than the fore-wings ; cilia of both wings
brown, with some ochreous hairs ; underside of wings paler, margins
broadly ochreous ; body and legs also ochreous,
Expanse of wings 1,/; inches.
Hab, SINGAPORE (Lidley).
There is another female caught apparently at the same
time and place by Ridley, which differs from the above in
having the apex of fore-wings tinged with ochreous, and
containing a brown subapical spot with a brown dot close
above it.
EUPROCTIS MAGNA.
Somena magna, Swinhoe, Trans. Ent. Soc., 1891, p. 479, §.
Swinhoe, Ann. Mag. N. H. (7), xii, p. 197,
b dy
¢ (1903).
1 $,19, Khasia Hills (types).
I have one male and two females from the same locality
in my own collection.
EUPROCTIS BIPARTITA.
Cherotricha bipartita, Moore, Lep. Atk., p. 49, pl. 2, f. 4
(1879).
1 2, Darjiling.
The type from Darjiling is in coll. Staudinger.
EUPROCTIS HAMPSON], nov. nom.
Artaxa variegata, Ampsn., Ill. Het., vii, p. 56, pl. 140, f. 6
(1891) (preocce.).
1 g, Nilgiri Hills (type).
Variegata being prececupied by Moore for Daplasa
variegata, which is an Kuproctis, Hampson’s species must
have a new name.
EUPROCTIS LUTEIFASCIA.
Artaxa luterfascia, Hmpsn., Il. Het., viii, p. 57, pl. 141, f. 2
(1891).
1 ¢, Nilgini Hills (type).
EUPROCTIS ATRISIGNATA, nov.
g¢. Antenne, head, thorax, and fore-wings red-brown, tinged with
ochreous; fore-wings with three pairs of white spots on the outer
424 Colonel C. Swinhoe’s Kevision of the
margin, apical, medial, and on the angle ; a small white dot between
each ; faint indications of two transverse pale lines, the first out-
wardly curved, the other outwardly curved above its middle ; a large
prominent black spot at the end of the cell ; cilia with white spots ;
hind-wings blackish-brown, with ochreous white outer marginal
border and cilia; under-side, palpi, pectus, body, legs, and wings
ochreous white; fore-wings suffused with brown, except on the
margins ; hind-wings suffused on the abdominal half; a black spot
at the end of each cell of both wings.
Expanse of wings 1,3; inches.
Hab, SINGAPORE (Lidley), two examples.
EUPROCTIS SCINTILLANS.
Somena scintillans, Walker, xu, 1734 (1856).
Artaxa justiciw, Moore, Cat. Lep. E. I. C., ii, p. 352, pl. xvi,
ff. 6 (larva) (1859).
Orvasca subnotata, Walker, xxxii, 502 (1865).
EHuproctis mooret, Snellen, Tijd. v. Ent., xxii, p. 106, pl. 8,
f. 8 and 10 (1879).
Artaxa limbata, Butler, Il]. Het., v, p. 53, pl. 90, f.3 (1881).
Somena wrrorata, Moore, Lep. Ceylon, 11, p. 87, pl. 111, f. 2
(1882).
A long series from different parts of India, Ceylon,
Burma, and Singapore. I have it in my own coll. from
Bombay, Poona, Karachi, and the Khasia Hills. I do not
see how any of the above can be specifically separated.
EUPROCTIS SIMILIS.
Artaxa similis, Moore, Cat. Lep. E. I. C., 11, p. 351
(1859).
1 9, Java (type). 2 2, Singapore.
The ¢ type I cannot find.
EUPROCTIS BIPUNCTAPEX.
Somena bipunctapex, Hmpsn., Nl. Het., vil, p. 57, pl. 140,
f. 13 (1891).
29, Nilgiri Hills, including the type. 2 9, Kangra.
19, Burma. 1 3%, Singapore. 7 3g, 10 9, China.
EUPROCTIS OBSCURA.
Artaxa obscura, Moore, Cat. Lep. E. I. C., u, p. 351
(1859).
1 2, Java (type). 1 9, Singapore.
Old World Lymantriide in the National Collection. 425
EUPROCTIS ATOMARIA.
Artaxa atomaria, Walker, iv, 796 (1855).
Butler, Ill. Het., v, p. 58, pl. 90, f. 2
(1881).
19, N. India (type).
9 »
EUPROCTIS APICALIS.
Arna apicalis, Walker, v, 1177 (1855).
Artaxa apicalis, Moore, Lep. Ceylon, ii, p. 85, pl. 111, f. 4,
4a, b (1883).
2 2, Bombay. 2 f, 3 2, Ceylon, including the type.
I cannot agree with Hampson, that this is the same as
the northern form atomaria, which has no yellow costa.
EUPROCTIS ABJECTA.
Somena abjecta, Swinhoe, P. Z. 8., 1889, p. 405, pl. 43,
iL” 118
1 f, 4 @, Karachi and Kotri, Sind, including the types.
EUPROCTIS SAGROIDES.
Somena sagroides, Hmpsn., Ill. Het., viii, p. 57, pl. 140, £, 14
(1891).
1 g, Nilgiri Hills (type).
EUPROCTIS FLAVICOSTA.
Euproctis flavicosta, Hmpsn., Journ. Bo. N. H. Soc., xiii, p.
420, pl. B, f. 3 and pl. 2, f. 18 (1900).
1 f, Sikhim (type).
EUPROCTIS RENIFERA.
Luproctis renifera, Swinhoe, Trans. Ent. Soc. 1895, p.
LOE
is 2 Swinhoe, Ann. Mag. N. H. (7), xii, p.
196, f (1903).
1g, 2 ¢, Khasia Hills (types).
EUPROCTIS MINUTISSIMA, nov.
¢. Antenne, palpi, head, thorax, and fore-wings ochreous ; fore-
wings sparsely irrorated with grey atoms, and with a large black
sub-apical spot near outer margin ; hind-wings black, with ochreous
cilia ; under-side, body and legs ochreous-white, both wings covered
426 Colonel C. Swinhoe’s Revision of the
with blackish-brown suffusion, the borders ochreous white, rather
broadly so at the apex of the fore-wings.
Expanse of wings +, inch.
Hab. SINGAPORE (Ridley).
With the spot on the fore-wings as in rubiginosa,
Snellen, from Java, Tijd. v. Ent., xx, p. 107 plete
(1877).
EUPROCTIS LINTA.
Artaxa linta, Moore, Cat. Lep. E. I. C., ii, p- 351 (1859).
1 $, Java (type). 2 3, Sandakan.
EUPROCTIS KALA.
Artaxa kala, Moore, Cat. Lep. E. I. C., ii, p. 351 (1859).
1 9, Java (type).
EUPROCTIS XANTHOPERA:
Huproctis canthopera, Hmpsn., Journ. Bo. N. H. Soe., x1, p.
295 (1897).
1 Z, Khasia Hills (type).
EUPROCTIS ORMEA, nov.
2. Palpi yellowish white, grey above; frons and pectus ochreous ;
antenne and thorax greyish-ochreous ; fore-wings with the ground
colour whitish-ochreous, almost entirely suffused with ochreous-
brown, and irrorated with black atoms, leaving some apical streaks,
a longitudinal streak below on the outer margin, and the outer
marginal line pale; two transverse lines also of that colour both
slightly sinuous, the first a little before the middle, nearly erect,
curving somewhat outwards above the middle, the other more or less
dentated, starting from the middle of the hinder margin close to the
first line, then well curved outwards and inwards on to the costa ;
a pale spot at lower end of the cell; cilia, with ochreous spots; hind-
wings pale black, with ochreous cilia ; abdomen and body below pale
black ; under-side of both wings suffused with pale black, with
ochreous margins ; legs yellowish-white.
Expanse of wings 1} inches,
Hab. 8. HE. BoRNEO (Doherty).
EUPROCTIS CERASINA, DOV.
4. Palpi white, black on the inner sides ; frons, antenna, and head
ochreous ; thorax and wings pale primrose-yellow ; hind-wings
Old World Lymantriide in the National Collection. 427
without markings ; fore-wings covered with cherry-coloured scales,
irrorated sparsely with black atoms, the ground colour of the wings
showing through here and there ; two transverse pale lines, anti-
medial and discal, both elbowed outwards above the middle, the first
acutely, the second somewhat near the outer margin; the space
beyond these mostly primrose-yellow, with some cherry-coloured
scales and black irrorations: the marginal border with very pale
pinkish markings, and a biack mark on the cilia at the middle of the
margin ; abdomen ochreous with some brown suffusion ; under-side
of body and legs ochreous, tarsi whitish ; wings pale primrose-
yellow, nearly white, with a rather broad blackish suffused sub-
costal streak on fore-wings.
Expanse of wings 1,45 inches.
Hab. SAMBAWA (Doherty).
EUPROCTIS OLIVATA.
Huproctis olivata, Hmpsn., Journ. Bo. N. H. Soe., xi, p. 295
(1897).
1 f, 1 9, Khasia Hills (types).
EUPROCTIS SUBRANA.
Artaxa subrana, Moore, Cat. Lep. E. I. C., ii
(1859).
1 @, Java (type). 1 2, Singapore.
, p. ddl
KUPROCTIS DISTRACTA.
Artaxa distracta, Walker, xxxu, 333 (1865).
1 g, Sarawak (type).
The type specimen is much rubbed and _ hardly
recognisable.
EUPROCTIS ALBODENTATA.
Pida albodentata, Moore, P. Z. 8., 1879, p. 401.
1 9, N. W. Himalayas (type). 1 9, Burma.
EUPROCTIS FACETA, nov.
6. Palpi, frons, head, and thorax white ; fore-wings greyish-
olive, costa white; two transverse white lines, ante-medial and
discal ; the first, nearly erect, bends inwards on to the costa, consisting
of two largish lunules, the lower being the outside margin of a large
white spot in the interno-median interspace, and which contains a
428 Colonel C. Swinhoe’s Revision of the
red-grey spot, while below this is the half of a similar mark on the
hinder margin; the outer line is lunulate and dentate, and curves
outwardly in its centre ; the lunules of the inner line are bent out-
wards and of the outer line inwards ; the cilia are white with grey
marks at the ends of the veins; hind-wings and abdomen sordid
ochreous white, unmarked; under-side nearly white, unmarked,
except for the grey streaks below costa of fore-wings.
Expanse of wings 5 inch.
Hab. Kapaur, New Guinea (Doherty).
EUPROCTIS ATRIPUNCTA.
Euproctis atripuncta, Hmpsn., Journ. Bo. N. H. Soc., xi, p.
295 (1897).
1 f, 2 9, Khasia Hills (types).
EUPROCTIS RECRABA.
EHuproctis recraba, Swinhoe, Ann. Mag. N. H. (7), xii, p.
196 (1903).
1 ¢, 1 2, Java (types).
EUPROCTIS ALBA, nov.
d, 2. Palpi grey, white beneath ; antenne grey ; head, thorax,
and wings white; fore-wings with some indistinct red scales at the end
of the cell, and also at the base ; some black irrorations on the lower
portion of the wing, more dense in the female than in the male ; hind-
wings with some brown suffusion on the abdominal area in the
female only ; abdomen blackish above, except the base and the anal
tuft, which are ochreous ; under-side white without markings.
Expanse of wings 1,5 inches,
Hab, SAMBAWA (Doherty).
EUPROCTIS LANARIA.
Terphothrix lanaria, Holland, Psyche, vi, p. 474 (1593),
2 f, Ogove River (Holland).
EUPROCTIS VARIEGATA.
Huproctis variegata, Moore, Lep. Atk., p. 48, pl. 2, f 24
(1879).
Daplasa variegata, Ampsn., Moths, India, 1, p. 458 (1892).
1 f, Sikhim (Pilcher).
The type from Darjiling is in coll. Staudinger. |
Old World Lymantriide in the National Collection. 429
EUPROCTIS STRIGIFIMBRIA.
Antipha strigifimbria, Walker, Journ. Linn. Soc. Lond., vi,
p. 126 (1862).
Artaxa fracta, Walker, xxxii, 333 (1865).
6 f, Sarawak, including both types. 1 2, Pulo Laut.
EUPROCTIS COSTALIS.
Antipha costalis, Walker, iv, 806 (1855).
Utidava incomptaria, Walker, xxvi, 1689 (1862).
Lacida complens, Walker, xxxu, 336 (1865).
Rilia (2) illepida, Walker, xxxii, 436.
5 2, 2 2, Ceylon, including all the types except complens,
which was in coll. Layard, but the collection is lost.
EUPROCTIS ANTICA.
Lacida antica, Walker, iv, 802, J (1855).
Anaxila notata, Walker, iv, 919, &.
4 f, 2 2, Ceylon, including both types.
EUPROCTIS ANTIPHATES.
EHuproctis antiphates, Hmpsn. Moths, India, i, p. 478
(1892).
1 fg, Khasia Hills (type).
EUPROCTIS PHA.
EHuproctis phexa, Hmpsn., Journ. Bo. N. H. Soc., xiii, p.
234 (1900).
2 2, Khasia Hills, including type.
_ Genus Hapiopseustis, Meyr., Trans. Ent. Soc. 1902, p. 34.
Acnissa, Turner, Trans. Roy. Soc. 8. Australia, 1902,
p- 180.
HAPLOPSEUSTIS ERYTHRIAS,
Haplopseustis erythrias, Meyr., |. c.
Acnissa pyrrhias, Turner, |. ¢.
2 f, 2 2, Port Darwin, whence also Meyrick’s type.
The type pyrriias from Townsville, Queensland.
430 Colonel C. Swinhoe’s Revision of the ,
Genus PERINA, Walker, iv, 966 (1855).
PERINA NUDA.
Bombyx nuda, Fabr., Mant. Ins., 11, p. 119 (1787).
Stilpnotia subtencta, Walker, iv, 843, 9.
Perina basalis, Walker, iv, 966, 2.
Euproctis combinata, Walker, xxxii, 347, (1865).
2 t,3 2, Hong Kong, including type of combinata. 5 f,
2 9, Nepal, including type of basalis. 3 2, 1 2, Bombay.
12, Mhow. 292, Belgaum. 1 9, Nilgiri Hills. 1 4,3 9,
Ceylon.
The type of swbtincta from India is in Mus. Oxon.
PERINA PURA.
Perina pura, Walker, Char. Undescr. Lep. Het., p. 17
(1869).
1 Z, without locality (type) ex. coll. Norris. 2 2,39, N.
India. 1 9, Calcutta. 1 9, Cachar.
Genus Dapiasa, Moore, Lep. Atk., p. 51 (1879).
DAPLASA IRRORATA.
Daplasa irrorata, Moore, 1. ¢., p. 52, pl. 2, f. 17.
1 f, Khasia Hills. 2 3, Omeishan.
The type from Darjiling is in coll. Staudinger.
Genus Cispia, Walker, iv, 857 (1855).
CISPIA CHARMA.
Cispia charma, Swinhoe, Ann. Mag. N. H. (7), ui, p. 112
(1899).
3 7, 2 2, Karwar, including the types.
CISPIA PUNCTIFASCIA.
Cispia punctifascia, Walker, iv, 857 (1855).
3 2,1 9, Sikhim, including the types. 1 9, Ceylon.
CISPIA VENOSA.
Cismia venosa, Walker, Trans. Ent. Soc., 1862, p. 264.
Drs 20% Sikhim.
The type, marked India, is in Mus. Oxon.
Old World Lymantriide im the National Collection. 431
Genus HERACULA, Moore, P. Z.8., 1865, p. 804.
HERACULA DISCIVITTA.
Heracula discivitta, Moore, |. ¢., pl. 48, f. 2.
3 3,1 §, Sikhim.
The type is in coll. Russell.
Genus Pipa, Walker, xxxii, 399 (1865).
Locharna, Moore, Lep. Atk., p. 53 (1879).
PIDA APICALIS.
Pida apicalis, Walker, xxxui, 400.
a x Hmpsn., Moths, India,i, p. 457 (fig.) (1892).
1 g, Darjiling (type). 12, Nepal. 32, Sikhim. 17,
Silhet. 1 2, Khasia Hills.
PIDA DECOLORATA.
Cyclidia (2) decolorata, Walker, Char. Undescr. Lep. Het., p.
96 (1869).
1 f, Khasia Hills. 1 2, Darjiling. 1 ¢, Subathu.
The type from Benares is in the Devon and Exeter
Museum.
PIDA STRIGIPENNIS.
Locharna strigipennis, Moore, Lep. Atk., p. 53, pl. 3, f. 11
(1879).
19, Burma. 1 2, Shan States. 17, Omeishan. 1 9,
Ichang. 1 9, Chang Yang.
The type, from the Khasia Hills, is in coll. Staudinger.
Genus FoDINOIDEA, Saalm., Madag., 1, p. 154 (1884).
FODINOIDEA STAUDINGERI.
Fodinoidea staudingert, Saalm., |. c., pl. 5, f. 63, 63a.
2 f, Madagascar.
FODINOIDEA VECTIGERA.
Eusemia vectigera, Mab., Le Nat., 11, p. 100 (1882).
Fodinoidea maculata, Butler, Ent. Mo. Mag., xxi, p. 198
(1885).
2 f, Madagascar, including Butler’s type.
TRANS. ENT. SOC. LOND. 1903.—PART Ill. (OcT.) 29
4.32 Colonel C. Swinhoe’s Revision of the
Genus NuMENES, Walker, 101, 662 (1855).
Pseudomesa, Walker, iv, 928 (1855).
NUMENES PATRANA.
Numenes patrana, Moore, Cat. Lep. E. I. C., u, p. 367
(1859).
Numenes partita, Walker, xxxi, 290 (1864).
142, Bhutan (type). 7 2,49, Sikhim.
The type of partita from Darjiling is in coll. Staudinger.
NUMENES SILETTI.
Numenes siletti, Walker, 111, 663, 2 (1855).
Pseudomesa quadriplagiata, Walker, iv, 923, $ (1855).
2 2,39, Silhet, including both types. 2 2, Sikhim. 1 ,
Cachar. 6 9, Burma.
NUMENES INSIGNIS.
Numenes insignis, Moore, Cat. Lep. E. I. C., u, p. 367, pl.
10a, £. 6,? (1859).
Pseudomesa leta, Walker, xxxv, 1916, (1866).
2 ft, 4 2, Java, including both types.
NUMENES DISPARILIS.
Numenes disparilis, Staud., Rom. sur Lep., 1, p. 200, pl.
il, f 2a; bi(iss7):
Numenes disparilis, var. separata, Leech, Entom., xxii, p.
112 (1890).
Lymantria albofascia, Leech, P. Z. 8., 1888, p. 629, pl. 31,
i, 0, c
2,12, Chang Yang, including type of separata. 1 2,
Ohoyama (type albofascia), 19, Chia-kow-ho. 1 9
Kiukiang. 1 2, Moupin.
?
NUMENES LIBYRA.
Aroa libyra, Druce, Ann. Mag. N. H. (6), xvi, p. 352.
(1896).
Heteronygiia flammeola, Distant, Ann. Mag. N. H. (7),
vu, p. 360 (1899).
Druce’s types f 2? came from W. Africa, Distant’s type
? from the Transvaal ; the species is not in the B. M.
Old World Lymantriide in the National Collection. 433
NUMENES HYPOXANTHA.
Heteronygmia hypoxantha, 2, Holland, Psyche, vi, p. 416
(1893).
One female from the Ogove River, the locality where
Holland’s type came from ; it is closely allied to Numenes
libyra, Druce.
Genus MArpDARA, Walker, xxxii, 402 (1865).
Mahoba, Moore, Lep. Atk., p. 52 (1879).
MARDARA PLAGIDOTATA.
Cyclidia (?) plagidotata, Walker, xxv, 1483 (1862).
2 2, Darjiling, including the type.
MARDARA RUFICEPS.
Mardara ruficeps, Hmpsn., Moths, India, iv, p. 489 (1896).
1 f, Bhutan (type).
MARDARA CALLIGRAMMA.
Mardara calligramma, Walker, xxxii, 402 (1865).
7 2, 2 2, Darjiling, including the type. 1 2, Kurseyong.
1 2, Sikhim.
MARDARA AFRICANA.
Lepasta africana, Holland, Ent. News Phil., 1893, p. 343,
oi Abs. ai ile
1 Z, Sierra Leone. 1 2, River Niger.
Genus NycTEmERA, Hiibner, Verz. bek. Schmett., p. 178.
Otroeda, Walker, 11, 402 (1854).
NYCTEMERA HESPERIA.
Geometra hesperia, Cram., Pap. Exot., iii, pl. 251, f. A, B
(1780).
Otroeda hesperia, Walker, ii, 402.
14 3,19, W. Africa.
NYCTEMERA JONESI.
Otroeda jonesi, Sharpe, Ann. Mag. N. H (6), vii, p. 184
(1891).
1g, Congo. 4 2, W. Africa.
434 Colonel C. Swinhoe’s Revision of the
NYCTEMERA MANIFESTA, nov.
¢. Belongs to the hesperia group, and is closely allied to that
species, but there is no sign of orange or ochreous on the hind-wings ;
the submarginal spots on both wings are somewhat differently placed,
forming a continuous and evenly-disposed curve, the upper one the
largest, lessening in size hindwards on both wings; in hesperia and
its allies the subcostal spot of the fore-wings is displaced inwards,
making the row of spots angled at its upper part, and in the
hind-wings the second spot (and sometimes the fourth) runs into the
white of the wing.
Expanse of wings 3} inches.
Hab. CoNnco.
NYCTEMERA CAFRA.
Bombyx cafra, Drury, Il. Ex. Ins., iii, pl. 5, f. 1 (1780).
2 g, Sierra Leone. 1 g, River Niger. 1 ¢, 8. Nigeria.
NYCTEMERA OCCIDENTIS.
Otroeda occidentis, Walker, 11, 403 (1854).
1 f, Sierra Leone (type). 1 ¢,Cameroons. 1 ¢, R. Gaboon.
i ¢, Rio del Rey. 1 2,8. Nigeria.
NYCTEMERA VESPERINA.
Otroeda vesperina, Walker, ii, 403 (1854).
2 f, Congo, including the type.
NYCTEMERA VARUNAA.
Otroeda varunxa, Druce, P. Z. 8., 1882, p. 780.
1st. Wi Atrica.
The type, from Congo, is in coll. Druce.
Genus RAJACOA, nov.
¢. Antenne bipectinated to the tips, pectinations ciliated and of
moderate length ; palpi upturned, very short and hairy ; fore-wings
with discoidal nearly straight, vein 2 from middle of cell, 4 from
lower end, 3 from half-way between, 5 from below middle and some-
what curved, 6 from below upper end, 7 and 8 stalked, 9 from 10
anastomosing with 7 and 8 to form the areole, 11 from one-third
before end of cell ; hind-wings with lower half of the discoidal pro-
truding, vein 2 from one-third before lower end of cell, 4 from end,
Old World Lymantriide in the National Collection. 435
3 from half-way between, 5 from a little above end and curved, 6
and 7 on a long stalk.
Type £. forbesi, Druce (Cypra).
RAJACOA FORBESI.
Cypra (2) forbesi, Druce, Ann. Mag. N. H. (7), ii, p. 469
(1899).
2 3, Fergusson Isl. (Meek). 2 2, Milne Bay (Jeek).
RAJACOA ANTRA.
Rajacoa antra, Swinhoe, Ann. Mag. N. H. (7), xu, p. 194
(19038).
1 ¢, Humboldt Bay, New Guinea (Doherty).
I have also two males in my own collection from the
same locality.
Genus Crmo.a, Walker, iv, 817 (1855).
CIMOLA OPALINA.
Cimola opalina, Walker, iv, 817.
Anomeotes thymiathis, Druce, Ent. Mo. Mag., xx, p. 156
(1883).
3 $, Delagoa Bay. 5, Natal, including the type.
CIMOLA ELEUTEBRIA.
Bombyx eleuteria, Stoll, Suppl. Cram., pl. 36, f. 18 (1791).
Cypra eleuteria, Walker, iv, 816 (1855).
10 4 19, Sierra Leone. 17) “Ashanti, 12, Benue
River.
Genus Pirea, Aurivillius, Ent. Tidsk., xii, p. 192 (1892).
PIRGA LASEA.
Xenosoma (?) lasea, Druce, Ann. Mag. N. H. (7), iu, p. 468
(1899).
2 f, 1 2, Sierra Leone.
Druce’s type came from the same locality. The type of
the genus is P. mirabilis, Auriv., described in Ent. Tidsk.,
xiii, p. 192, but figured in vol. xii, p. 228, pl. 2, f. 2, and put
in the Family Vyctemeridx = Deilemeridx: Karsch, however,
in Ent. Nachr., xvi, p. 351 (1900), pointed out that this
genus along with Otroeda, Walker = Nyctemera, Hiibn.,
really belongs to the Lymantrude.
436 Colonel C. Swinhoe’s Revision of the
PIRGA MAGNA, nov.
3, 2. Palpi, frons, antennz, head, ‘and body deep black, collar
golden-orange ; all the markings on the body of the same colour; a
tuft on each side of the thorax, a band behind, and segmental bands
on the abdomen ending on each side in bright orange-red spots,
joined together on the abdomen below by thin orange-red lines ;
wings nearly hyaline, the veins dark brown ; fore-wings pale blackish ;
a sordid white sub-costal streak from the base ; the interior third of
the interno-median space of the same colour ; hind-wings sordid
white, with the outer third pale blackish narrowing hindwards ;
under-side of body and legs black ; wings as above.
Expanse of wings ¢ 2,°%,, 9 3 inches.
Hab, Kixuyu, British E. Africa (Crawshay).
Genus MARBLA, nov.
Antenne bipectinated to the tips, branches of male long, of female
a little less than half the length of the male, minutely ciliated, each
branch with two long and one short spine at itsend ; palpi long and
porrect, last joint very short and blunt; fore-wings with the costa
bent at the base, otherwise nearly straight ; apex rather produced but
rounded ; outer margin slightly curved, two-thirds length of costa ;
hinder margin of the male much rounded, of the female nearly
straight ; discoidal bent inwards in its centre, vein 2 from one-third
before end of cell, 3 from before end, 4 and 5 from end, 6 from a
little below upper end, 7 and 8 on a long stalk from upper end of
cell, 9 and 10 also on a long stalk arising from a little before the end,
11 from one-third before end ; hind-wings with the lower end of the
discoidal square and slightly projected, vein 2 from a little beyond
middle of cell, 3 from before end, 4 from the end, 5 from a little
above, 6 and 7 from the upper end.
Type JL. divisa, Walker (Hloria).
MARBLA DIVISA.
Eloria divisa, Walker, iv, 815 (1855).
Hylemera tenera, Holland, Ent. News Phil., iv, p. 61, pl. 3,
f, 7 (1893).
Walker’s type, a male, has no locality ; but there are
two male and seven female examples, the same as type,
from Sierra Leone and Lagos: Holland’s description and
figure fits this species very well.
Old World Lymantriidx in the National Collection. 437
MARBLA (?) NYSES.
Cypra nyses, Druce, P. Z. S., 1887, p. 678.
1 g, Old Calabar.
The type from Oid Calabar is in coll. Druce.
MARBLA (?) BIGUTTA.
Soloé bigutta, Holland, Ent. News Phil., 1893, p. 62, pl. 3,
i
1 7, 1 2, Ogove River (Holland).
Genus Pantana, Walker, iv, 819 (1855).
Lirnara, Butler, Trans. Linn. Soc. Lond. (2), 1
560 (1879).
PANTANA VISUM.
Liparis visum, Hiibner, Zutr., 11, p. 82, f. 543, 544 (1825).
Pantana dispar, Walker, iv, 820.
1 2, Maulmein (type dispar). 5 2, Burma. 1 2, Shan
States. 2 2%, Tonkin.
PANTANA AMPLA.
Pantana ampla, Walker, iv, 820 (1855).
1 , Hong Kong (type). 5%, Hainan.
PANTANA BICOLOR.
Orgyia bicolor, Walker, iv, 787 (1855).
Genusa delineata, Walker, iv, 818.
Genusa circumdata, Walker, iv, 819.
Stilpnotia sordida, Walker, vil, 1732 (1856).
Genusa comparata, Walker, xxxil, 340 (1865).
1 f, Kangra. 1 ¢, Dharmsala. 3 f, N. India, including
type of circumdata. 37, Silhet, including types of
delineata and sordida. 49,29, Sikhim. 2 3,1 2, Khasia
Hills. 17, Bhamo, Burma.
The type, marked India, is in Mus. Oxon.
PANTANA BASWANA.
Pantana baswana, Moore, Cat. Lep. E. I. C., ui, p. 336, pl.
Gay td (1859):
5 f, Java, including the type.
438 Colonel C. Swinhoe’s Revision of the
PANTANA SIMPLEX.
Pantana simplex, Leech, Trans. Ent. Soc., 1899, p. 122.
8 ¢, W. China, including the type.
PANTANA TERMINATA.
Genusa terminata, Walker, xxxii, 340 (1865).
1 g, India (type). 5 $,1 9, Burma.
PANTANA NIGROLIMBATA.
Pantana nigrolimbata, Leech, Trans. Ent. Soc., 1899, p. 121.
5 $,3 9, W. China, including the type.
PANTANA SINICA.
Pantana sinica, Moore, Ann. Mag. N. H. (4), xx, p. 92
(1877).
* » Leech, Trans. Ent. Soc., 1899, p. 121.
1 Z, Shanghai (type). 1%, Chekiang, 1 2, Fouchau.
PANTANA ALBIFASCIA.
Orgyia albifascia, Walker, xxx1i, 825 (1865).
1 f, Darjiling.
The type from Darjiling is in coll. Staudinger. I have
it from the Khasia Hills.
PANTANA SUBFASCIA.
Orgyra subfascia, Moore, P. Z. 8., 1865, p. 803.
1 f, Bengal (type).
Hampson puts this with a/bifascia, but it seems to me
to be a very good form; there are some examples from
Sikhim in coll. Pilcher.
PANTANA INTERJECTA.
Pantana interjecta, Swinhoe, Trans. Ent. Soc., 1891, p. 478,
[OG TEs a od
1 fg, Khasia Hills (type).
I have several examples from the same locality.
PANTANA LUTEICEPS.
Pantana luteiceps, Swinhoe, Ann. Mag. N. H. (6), xvii,
p. 361 (1896).
1 2, Khasia Hills.
Old World Lymantriidx in the National Collection. 439
PANTANA PLUTO.
Gynephora pluto, Leech, Entom., xxiii, p. 111 (1890).
2 ft, Ichang, including the type. 1 g, Moupin. 1 7,
Kwei-chow.
PANTANA EURYGANIA.
Pantana eurygania, Druce, Ann. Mag. N. H. (7), ii, p. 470
(1899).
1 ¢, Chang Yang.
The type from Szechuen is in coll. Druce.
PANTANA SEMILUCIDA, nom. nov.
Pantana bicolor, Walker, iv, 820 (1855) (preocc.).
1 f, without locality (type). 1 gf, Cambogia, Siam.
2g, Penang. 1 gf, Sarawak.
PANTANA LINEOSA.
Etobema lineosa, Walker, xxxui, 389 (1865).
Birnara nubila, Butler, Trans. Linn. Soc. Lond. (2), 1, p.
560 (1879).
2 ¢, Malacca, including Butler’s type. 1 2, Singapore.
1 9, Sumatra.
Genus L&LIA, Steph., Cat. Brit. Ins., 1, p. 52 (1827).
Charnidas, Walker, iv, 797 (1855).
Repena, Walker, iv, 799.
Anthora, Walker, iv, 801.
Lacida, Walker, iv, 802.
Procodeca, Walker, iv, 812.
Ricine, Walker, iv, 824.
Odagra, Walker, xxxii, 401 (1865).
Harapa, Moore, Lep. Atk., p. 47 (1879).
Lxlioides, Moore, Lep. Ceylon, ii, p. 83 (1883).
Hondella, Moore, 1. ¢., p. 144.
L#LIA SUBROSEA.
Anthora subrosea, Walker, iv, 801.
Lelia subrosea, Schaus and Clements, Sierra Leone Lep.,
We2Gply lt one (89s):
Lelia subrufa, Snellen, Tijd. v. Ent., xv, p. 39 (1872).
3 J, 1 2, Sierra Leone, including the type. 1 2, Natal.
Snellen’s type came from Lower Guinea.
440 Colonel C. Swinhoe’s Revision of the
LALIA FASCIATA.
Lxlioides fasciata, Moore, Lep. Ceylon, ii, p. 84, pl. 110,
f. 6 (1882).
Procodeca testacea, Moore, P. Z. 8., 1872, p. 574.
Leiroides rubripennis, Moore, Trans. Ent. Soc., 1884, p. 358.
1 g, Bengal (type testacea). 1 $,1 9, Barrackpore. 4 2,
2 2, Ceylon, including the type. 1 2, Burma (type rubri-
pennis). 1 2, Tongoo, Burma.
The female type of ¢estacea is in Mus. Oxon.
L&#LIA FIGLINA.
Lelia figlina, Distant, Ann. Mag. N. H. (7), iv, p. 361
(1899).
1 Z, 1 9, British E. Africa.
The type from the Transvaal is in coll. Distant.
L#LIA LILACINA.
Lelia lilacina, Moore, Trans. Ent. Soc., 1884, p. 357.
1 g, Nilgiri Hills (type).
LZLIA DEVESTITA.
Odagra devestita, Walker, xxxii, 402 (1865).
Lelia pallida, Moore, Trans. Ent. Soc., 1884, p. 358.
Lelvoides lactea, Moore, 1. ec.
1 2, Punjab Hills (type lactea). 19, Umballa. 2 f,
Karachi. 1 ¢, Bombay.
The type from Darjiling is in coll. Staudinger.
LALIA PROLATA.
Lelia prolata, Swinhoe, Cat. Het. Mus. Oxon., i, p. 195
(1892)
5 2, N. India, including the type. 1 2, 2 2, Maulmein.
1 Z, Kangra. 2 9, Borneo.
A perfectly distinct form.
LALIA OBSOLETA.
Bombyx obsoleta, Fabr., Ent. Syst., iii, 1, p. 463 (1793).
5 Sg, 49, Australia.
The type is in the Banksian Cabinet in the B. M.
©
Old World Lymantriide in the National Collection. 441
L&LIA SINENSIS.
Lelia sinensis, Walker, iv, 829 (1855).
Leucoma brevicornis, Walker, vii, 1729 (1856).
1 2, 2 2, Hong Kong, including both types.
LALIA CANDIDA.
Lelia candida, Leech, Trans. Ent. Soc., 1899, p. 121 (text).
3 2, Chang Yang, including the type.
It is a pure white form of cwnosa, Hiibner.
LAELIA SANGAICA.
Lelia sangaica, Moore, Aun. Mag. N. H. (4), xx, p. 92
(1877).
6 2,59, Japan. 1 2, Shanghai (type). 2 7, Ningpo.
L#LIA SUFFUSA.
Ricine suffusa, Walker, iv, 824, 9 (1855).
Procodeca angulifera, Walker, iv, 919, 2.
Lelia subrufa, Snellen, Tijd. v. Ent., xxxii, p. 105, pl. 8, f.6
(1879).
1 2, Maulmein (type 2, angulifera). 2 $, 3 9, Java,
including the tyye ¢.
Snellen has identified this species from Celebes as the
same as his African subrufa, but his figure represents the
Island form and not the African; this form is more closely
allied to Lelia sangaica, Moore, from China.
LALIA GIGANTEA.
Lelia gigantea, Butler, Cist. Ent., ili, p. 117 (1885).
1 2, 5 2, Japan, including the type.
LAELIA PUNCTULATA.
Lopera punctulata, Butler, Ann. Mag. N. H. (4), xvi, p. 400
(1875).
8 2, Natal, including the type.
LZLIA SETINOIDES.
Lelia setinoides, Holland, Psyche, vi, p. 431 (1893).
1 2, Ogove River (Holland). 1 9, Natal. 2 2, Kaniss,
Brit. E. Africa. 1 2, Shambe.
442 Colonel C. Swinhoe’s Revision of the
LAELIA CALAMARIA.,
Lelia calamaria, Ampsn., Journ, Bo. N. H. Soe., xiii, p.
234, pl. B, f. 19 (1900).
3 4, 2 9, Nilgiri Hills, including the types.
LA&LIA UMBRINA.
Procodeca wmbrina, Moore, P. Z. 8., 1888, p. 398.
Charnidas umbrina, Butler, Ill. Het., vii, p. 36, pl. 123, f. 8
(1889).
1 g, Kulu (type). 1 3, Subathu. 2 2, Dharmsala. 1 2,
Dalhousie. 2 2, Kangra. 1 2,1 2, Sultanpore.
LZLIA CARDINALIS.
Lelia cardinalis, Hmpsn., Ill. Het., ix, p. 74, pl. 158, f. 29
30 (1893).
2,1 2, Ceylon, including the type.
>
LALIA BUANA.
Phragmatobia buana, Moore, Cat. Lep. E. I. C., ii, p. 358
(1859).
1 f, Java (type).
LALIA FRACTA.
Lelia fracta, Schaus and Clements, Sierra Leone Lep., p.
26, pl. 1, f£. 12 (1893).
3 2,2 9, Sierra Leone.
LALIA VENOSA.
Lelva venosa, Moore, P. Z.8., 1877, p. 601, pl. 59, f. 1.
3 ¢, Andamans, including the type. 1 9, Singapore.
LA&LIA ATESTACEA.
Lelia atestacea, Hmpsn., Moths, India, 1, p. 443 (1892).
Harapa testacea, Moore, Lep. Atk., p. 47, pl. 2, f. 15, 2
(nom. preocc.).
1 g, Khasia Hills.
There is a female in coll. Pilcher from Sikhim, and both
sexes in my own coll. from the Khasia Hills. The type
came from Darjiling, and is in coll. Staudinger; it is the
largest species in the genus, even larger than adalia,
Swinhoe.
Old World Lymantriide in the National Collection. 443
LALIA LAVIA, nov.
¢. Olive-brown ; palpi with black hairs beneath ; frons, antenne,
head, and thorax dark brown ; shafts of antenne white ; fore-wings
pale olive-brown, darkest on the margins ; a discal outwardly curved
row of seven black spots, and a blackish lunule at the end of the cell ;
abdomen and hind-wings paler, evenly coloured ; a blackish lunule
at the end of the cell; under-side dull pale olive-grey ; a brown
lunule at the end of each cell ; legs ochreous with brown stripes.
Expanse of wings 1,%; inches.
2 ¢, Lavi, Brit. E. Africa (Betton).
L#LIA EXCLAMATIONIS.
Euprepia exclamationis, Kollar, Hugel’s Kasch., iv, p. 469
(1844).
Repena cervina, Walker, iv, 800 (1855).
Lacida rotundata, Walker, iv, 802.
Cycnia rubida, Walker, xxxi, 297 (1864).
Lymantria disjuncta, Walker, xxxii, 366 (1865).
9 f, 12 ¢, from various parts of India and Ceylon,
including the types of votwndata from Ceylon, and disjuncta
from South India.
The type of cervina from India is in Mus. Oxon., the
type of rubida from Ceylon was in Layard’s lost collection.
LALIA ADALIA.
Lxlia adalia, Swinhoe, Ann. Mag. N. H. (7), vi, p. 307
(1900).
1 f,1 ¢, Jaintia Hills (types).
Put with exclamationis in the B. M., but it is certainly
more nearly allied to atestacea; it is a good and distinct
form, and one of the largest of the genus. I have a nice
series of both sexes,
L2ALIA TESTACEA.
Cycnia testacea, Walker, 111, 683 (1855).
Lelia wniformis, Hmpsn. (2 only), Il. Het., viii, p. 56, pl.
140, f. 20 (1891).
Charnidas colon, Himpsn., l. c., f. 3, 19.
1 2, N. India (type). 1 9, Allahabad. 1 2, Bombay.
19, Mhow. 2 g, 4 9, Nilgiri Hills, including both of
Hampson’s types. 1 2, Travancore.
44.4 Colonel C. Swinhoe’s Revision of the
LALIA ADARA.
Procodeca adara, Moore, Cat. Lep. E. I. C., 11, p. 337 (1859).
1 9, Java (type).
LALIA JUVENIS.
Ptilomacra juvenis, Walker, v, 1099 (1855).
Fondella juvenis, Moore, Lep. Ceylon, ii, p. 144, pl. 137,
f. 4 (1883).
2 f, 2 9, Ceylon, including the type.
A very peculiar and distinct form.
LALIA LITURA.
Charnidas litwra, Walker, iv, 797 (1855).
2 4,Subathu. 1 gf Kangra. 1 g, Nepal (type). 2 3,
Sultanpore.
LA&LIA FURVA.
Ocneria furva, Leech, P. Z.8., 1888, p. 631, pl. 31, f. 10.
ea? Ichane:. 19s Pekin:
I cannot find the type.
LALIA HETEROGYNA.
Lelia heterogyna, Hmpsn., Moths, India, i, p. 443 (1892).
12 7, 6 2, Dras, Kashmi, including the types.
The females have aborted wings.
Genus DacryLoRHYNCHA, Hmpsn., Moths, India, i, p. 470
(1892).
DACTYLORHYNCHA PALLIDA.
Charnidas pallida, Umpsn., Ill. Het., vii, p. 56, pl. 140, f.
10 (1891).
2 ¢, Nilgini Hills, including the type.
DACTYLORHYNCHA LUTEIFASCIA.
Dactylorhyncha luteifascia, Hmpsn., Trans. Ent. Soc., 1895,
p. 292.
1 f, Pauk Yaw, Burma (type).
Old World Lymantrudx in the National Collection. 445
Genus ANTHELA, Walker, iv, 853 (1855).
Darala, Walker, iv, 886.
Colussa, Walker, xxi, 288 (1860).
Leptocneria, Butler, Trans. Ent. Soc., 1886, p. 386.
Newmania, Swinhoe, Cat. Het. Mus. Oxon., i, p. 199
(1892).
ANTHELA RUBICUNDA.
Darala rubicunda, Swinhoe, Ann. Mag. N. H. (7), ix, p. 419
(1st June, 1902).
Anthela phenicias, Turner, Trans. Roy. Soc. 8. Australia
(1st July, 1902), p. 182.
1 g, Roebourne (type). 4 g, Sherlock River. 1 ¢,
Queensland.
Turner's type came from Roebourne.
ANTHELA PUDICA.
Darala pudica, $, Swinhoe, Ann. Mag. N. H. (7), 1x, p. 419
(Ist June, 1902).
Anthela aspilota, 2, Turner, Trans. Roy. Soc. S. Australia
(Ist July, 1902), p. 182.
1 f, Roebourne (type). 6 ¢, 2 2, Sherlock River.
ANTHELA RUBESCENS.
Darala rubescens, Walker, xxxui, 370 (1865).
1 g, Australia (type).
ANTHELA ELIZABETHA.
Odonestis clizabetha, White, Grey, Journ. Exped. Austral.
ii, p. 478 (1841).
2 f, King George’s Sound.
ANTHELA ADRIANA.
Darala adriana, Swinhoe, Ann. Mag. N. H. (7), ix, p. 419
(1902).
3 ¢, Sherlock River, including the type.
ANTHELA PARVA.
Darala parva, Walker, iv, 892 (1855).
1 g, Sydney. 1 3, Tasmania (type). 2 3%, 6 9, SE.
Australia. 1 2, Victoria.
Very near fer7ruginosa.
44.6 Colonel C. Swinhoe’s Revision of the
ANTHELA FERRUGINOSA.
Anthela ferruginosa, Walker, iv, 854 (1855).
1 9, Tasmania (type). 1 9,8.E. Australia. 19, Sydney
ANTHELA ADDITA.
Darala addita, Walker, xxxii, 372 (1865).
1 9, Tasmania (type).
ANTHELA VENOSA.
Colussa venosa, Rosenst., Ann. Mag. N. H. (5), xvi, p. 384
(1885).
1 9, S. Australia (type).
ANTHELA CLEMENTI.
Darala clementi, Swinhoe, Ann. Mag. N. H. (7), ix, p. 81
(1902).
3 7, 2 2, Sherlock River, including the types.
ANTHELA GUENEI.
Teara guenéi, Newman, Trans. Ent. Soc., 1856, p. 284,
fol: US yats wo:
Newmania guenéi, Swinhoe, Cat. Het. Mus. Oxon., i, p. 199
(fig.) (1892).
1 f, 2 §, Australia, including the type.
ANTHELA FIGLINA.
Darala figlina, Swinhoe, Ann. Mag. N. H. (7), ix, p. 81
(1902).
3 7, Sherlock River, including the type.
ANTHELA BINOTATA.
Leptocneria binotata, Butler, Trans. Ent. Soc., 1886, p. 386,
Ole Beat oe
1 3, Peak Downs (type). 1 9, 2 9, Port Darwin.
ANTHELA REDUCTA.
Darala reducta, Walker, iv, 888 (1855).
4 7, 3 9, Australia, including the type 2.
Old World Lymantriide in the National Collection. 447
ANTHELA OCELLATA.
Darala ocellata, Walker, iv, 887 (1855).
Ommatoptera tetrophthalma, Herr.-Schiiff., Ausser-Eur.
Schmett., f. 506, 507 (1856).
7 f, 12 9, Australia, including the types.
It is a somewhat variable insect.
ANTHELA DENTICULATA.
Teara denticulata, Newman, Trans. Ent. Soc., 1856, p. 283.
Darala basigera, Walker, xxxii, 372 (1865).
Darala undulata, Felder, Reise Nov., pl. 98, f. 11 (1868).
1 2, Adelaide (type). 1 ¢, Melbourne.
ANTHELA OSTRA, nov.
¢. Antenne black, shafts white ; palpi, frons, head, and shoulders
bright ochreous ; thorax blackish-brown ; wings dark black ; fore-
wings with a minute spot in the cell, a round spot at end, both yellow,
as also are the costal margin, the outer half of cilia, and a sub-
marginal row of dots; hind-wings with the entire cilia yellow,
and arow of larger spots near the margin, and extending into it ;,
abdomen dark dull orange ; under-side of body and legs bright
ochreous ; fore-wings yellow, with the reniform distinct, the disc
blackish, intersected by a yellow transverse line, the outer margin
slightly suffused with black ; hind-wings deep black, with a yellow
cell spot, and broad marginal yellow border.
Expanse of wings 1,3, inches.
Hab. ADELAIDE RIVER.
ANTHELA UNISIGNA, nov.
¢. Antenne blackish-brown, shafts. whitish ; palpi and frons
blackish-brown ; thorax covered with dark bright chestnut-brown
hairs, variegated with pure chestnut colour ; abdomen pale crimson,,
anal tuft white ; fore-wings pale ochreous-grey ; a transverse erect
and straight brownish-grey band beyond the middle, in which is a
prominent pure white spot ringed with blackish-brown at the end of
the cell, another more attenuated similar band in the disc, followed
rather closely by a broad brownish-grey marginal band ; hind-wings
paler, nearly white, with a similar broad outer marginal band ;
under-side of wings white, with the first band of fore-wings and
cell spot, the cell itself suffused with ochreous ; hind-wings with a
TRANS. ENT. SOC. LOND. 1903.—PART III. (octr.) 30
448 Colonel C. Swinhoe’s Revision of the
small brown lunular mark at upper end of cell; pectus and legs
brownish-grey, with some crimson hairs; abdomen white.
Expanse of wings 2,3, inches.
Hab. SHERLOCK River, W. Australia (Clement).
Of the shape of stygiana, Butler.
ANTHELA STYGIANA.
Darala stygiana, Butler, Ann. Mag. N. H. (5), ix, p. 88
(1882).
1 $, Melbourne (type).
ANTHELA TRITONEA, NOV.
?. Antenne black, shafts pale red; palpi and frons black,
ochreous at the sides; head brown; thorax and abdomen covered
with very dark blackish-chestnut hairs with a bronzed tint ; in the
female the thorax is covered with grey and white hairs; the
abdomen is coloured like that of the male, but the segments are
ringed with long crimson hairs ; fore-wings black, minutely and
very densely irrorated with grey, crossed by three black bands rather
close together in the middle of the wing, the first outwardly and
irregularly curved, the other two nearly erect, the last including a
small pale spot at the end of the cell ; another transverse black band
in the disc, sinuated, almost crenulate, outwardly edged with white,
and close to it a prominent white semi-dentate band, which runs
from close to apex toa little in front of the hinder angle ; hind-
wings pale brown tinged with ochreous, especially towards the base,
where there is a bunch of dark chestnut hairs in the male ; a broad
darker brown transverse straight band before the middle, a duplex
whitish sinuated thin band before the blackish border ; under-side
whitish irrorated with pale black; a white spot in cell of fore-
wings, another at the end, both ringed with brown ; a brown even
band across end of cell, and continued across both wings; a discal
sinuate thinner band; an inner brown spot on hind-wings ; thorax
and legs dark chestnut-brown, with some crimson hairs ; abdomen
white,
Expanse of wings ¢ 4, @ 4 -¢ inches.
Hab, QUEENSLAND.
These examples were presented in 1896 by Mr. Wailley :
they seem to be allied to A. magnifica, Lucas, Proc. Linn.
Soc. N.S. W. (2), vi, p. 286, but differ in many characters
from Lucas’s description ; they are the largest examples of
the genus I think yet recorded.
Old World Lymantriide in the National Collection. 449
ANTHELA CONNEXA.
Darala connexa, Walker, iv, 898 (1855).
Darala Jervens, Walker, iv, 898.
Darala zonata, Felder, Reise Nov., pl. 99, f. 1 (1868).
5 $,5 9, Tasmania, including types of connewa and fervens.
The type of zonata from Australia is in coll. Rothschild.
ANTHELA POSTICA,
Darala postica, Walker, iv, 899 (1855).
1 2, Australia (type).
ANTHELA NICOTHOE.
Bombyx nicothoé, Boisd., Voy. de | Astrolabe, i, p. 226
(1882).
Darala adusta, Walker, iv, 897 (1855).
Lelia australasivw, Herr.-Schaff., Ausser-Eur. Schmett,, f.
386 (1855).
9 3, 8 9, Australia, including Walker’s type.
ANTHELA REPLETA.
Darala repleta, Walker, iv, 896 (1855).
1 f, 2 9, Tasmania, including type ff.
ANTHELA CINERASCENS.
Darala cinerascens, Walker, iv, 900 (1855).
2 J, 1 9, including type 9.
ANTHELA RUFIFASCIA.
Darala rufifascia, Walker, xxxui, 370 (1865).
2 , 1 9, Tasmania, including type 9.
ANTHELA FERRUGINEA.
Darala ferruginea, Walker, iv, 890 (1855),
Ommatoptera diophthalma, Herr.-Schiff., Ausser-Eur.
Schmett., f. 508 (1856).
4 ¢, Tasmania, including type.
ANTHELA ACUTA.
Darala acuta, Walker, iv, 889 (1855).
3. ¢, New Holland, including the type. 5 2 S.E.
Australia.
450 Colonel C. Swinhoe’s Revision of the
ANTHELA DELINEATA.
Darala delineata, Walker, xxxil, 371 (1865).
1 ¢, Melbourne. 1 @, Australia (type).
ANTHELA SUBFALCATA.
Darala subfaleata, Walker, iv, 894 (1855).
1 ¢,2 9, Tasmania, including the type ¢.
ANTHELA HAMATA.
Darala hamata, Walker, iv, 895 (1855).
Darala latifera, Walker, Trans. Ent. Soc., 1862, p. 266.
Colussa uvaria, Walker, xxxv, 1576 (1866).
12,3, Moreton Bay, including types of Jatifera and
wvaria. 1 9, Sydney (type).
ANTHELA VARIA.
Darala varia, Walker, iv, 890 (1855).
Colussa odenestaria, Walker, xxi, 288 (1860).
2 ¢, New Holland, including the type. 1 ¢, SE.
Australia, 2 9, Australia, including the type of odenestaria.
ANTHELA SIMPLEX.
Darala simplex, Walker, iv, 891 (1855).
1 ¢, without locality (type). 2 ¢, 3 9, Sydney.
ANTHELA PLANA.
Darala plana, Walker, iv, 892 (1855).
1 g, Australia (type).
In poor condition, and without antenne, doubtfully
distinct from simplex.
ANTHELA CANESCENS.
Darala canescens, Walker, iv, 901 (1855).
Dreata deficiens, Walker, xxx, 874 (1865).
1 2, Australia (type). 1 2, S.E. Australia. 2 9, Swan
River. 1 ¢, N. Australia (type deficiens). 1 2,Carnarvon,
W. Australia.
They are of different shades of colour, but not more
so than is common in the genus, and their markings are
all identical.
Old World Lymantriide in the National Collection. 451
ANTHELA CARNEOTINCTA, nov.
3g, 9. Of a uniform ochreous-grey, tinged with flesh colour, the
female rather darker than the male and tinged with pink ; antenne,
palpi, pectus, under-side of body, and legs more or less ochreous ;
on the sides of the abdomen of the male there are some brown
patches ; on the fore-wings a brown dot in the middle of the cell
and a brown spot at the end; a very indistinct, more or less
crenulated grey line across the dise of both wings and even with the
margins ; cilia of both wings ochreous; in the female there is, in
addition, an inner discal duplex band in the fore-wings and indica-
tions of asimilar band in the hind-wings, and very faint indications
of an inner band across both wings; the under-side is evenly coloured
like the upper-side, but yellower in the male and pinker in the
female, with a brown spot in the cell and another at the end in both
wings, and indications of the upper transverse bands.
Expanse of wings, ¢ 275, 9 33 inches.
2 f, 1 2, Freemantle, bred March, 1891 (Walker).
ANTHELA CRENULATA, nov.
2. Palpi brown-pink ; antenne, head, body, and wings above
and below of a uniform ochreous-grey colour, with a slight shade of
pink in it ; fore-wings with a small brown spot in the cell, and larger
spot at the end; an ante-medial transverse brown band, inside the
first spot, bent outwardly in an irregular manner in the middle, and
continued in a very indistinct form across the hind-wings ; a discal
band across both wings formed by a very evenly crenulated line ;
cilia of both wings ochreous ; under-side with only the crenulated
discal band, and a brown spot in the cell and another at the end in
all the wings; fore-legs brown above.
Expanse of wings, 2,55 inches.
Freemantle, bred March, 1891 (Walker).
Of a beautiful tint of colour; but this colour varies
much, and in two examples is more yellow, and in two
others bright yellow; the markings are identical, though
in some examples more distinct than in others. They were
all bred at the same time and place.
ANTHELA CONSPERSA.
Darala conspersa, Walker, iv, 891 (1855).
Darala quadriplaga, Walker, Trans. Ent. Soc., 1862, p.
269.
Ennomos (2) potentaria, Walker, xxvi, 1519 (1862).
452 Colonel C. Swinhoe’s Revision of the
1 ¢, without locality (type). 19, New Holland. 1 9,
S.E. Australia. 1 9, Australia (type potentaria).
The type of quadriplaga marked Australia is in Mus.
Oxon.
ANTHELA EXCISA.
Darala excisa, Walker, iv, 889 (1855).
4 $, 1 9, Sydney, including the type 2.
Very near conspersa.
ANTHELA CONSORS.
Darala censors, Walker, xxxii, 369 (1865), (ex err. typ.).
Darala consors, Walker, xxxv, 1917 (1866).
3 g, S. Australia, including the type %. 1 3, S.E.
Australia.
ANTHELA EXCELLENS.
Darala excellens, Walker, iv, 902 (1855).
1 Z, 1 2, Australia, including the type &.
ANTHELA INTEGRA.
Darala integra, Walker, iv, 893 (1855).
1 $, New Holland (type.) 1 2, Australia.
ANTHELA LIMONEA.
Darala limonea, Butler, Cist. Ent., i, p. 291 (1874).
1 Z, 1 2, Rockhampton (types).
ANTHELA INORNATA.
Darala inornata, Walker, iv, 901 (1855).
1 g, Swan River (type).
ANTHELA FLAVALA, nov.
@. Of a uniform rather bright yellow colour; upper-side of
palpi brown ; both wings with a grey spot in the cell, and another
at the end; indications of an inner outwardly curved transverse
line, and a post-medial broader line, followed by two thin crenulated
lines ; under-side same as upper-side.
Expanse of wings, 13’; inches.
Hab. SHERLOCK River, W. Australia (Clement).
Old World Lymantriide in the National Collection. 453
There is also a still brighter yellow female, collected by
Dr. Clement at the same place, with very similar markings
(which are however very indistinct), in which the cell
spots are somewhat closer together.
ANTHELA PINGUIS.
Darala pinguis, Walker, xxxii, 372 (1865).
1 f,1 9, Australia, including the type 9.
ANTHELA OBSCURA.
Trichiura obscura, Walker, vi, 1481 (1855).
1 gf, Australia (type).
Not a typical Anthela, quite a small insect ; as it is an
unique example, its proper position can only be determined
when more specimens are obtained.
Genus PTEROLOCERA, Walker, iv, 883 (1855).
PTEROLOCERA AMPLICORNIS.
Piterolocera amplicornis, Walker, iv, 884.
1 g, Adelaide (type). 13, Melbourne. 5 2, Australia.
PTEROLOCERA INSIGNIS.
Pterolocera insignis, Herr.-Schaff., Ausser-Eur. Schmett.,
f, 459 (1856).
1 Z, Australia.
Genus Aroa, Walker, iv, 791 (1855).
Baziza, Walker, xxx, 398 (1865).
AROA MAXIMA.
Aroa maxima, Hmpsn., Ill. Het., ix, p. 74, pl. 159, f. 9
(1893).
1 f, Ceylon (type).
AROA MAJOR.
Aroa major, Hmpsn., Ill. Het., ix, p. 74, pl. 159, ff. 3,5
(1893).
1 g, 1 9, Pundaloya, Ceylon (types). 1 g, Ceylon.
1 2, 1 2, Trevandrum.
454 Colonel C. Swinhoe’s Revision of the
AROA PLANA.
Orgyia (?) plana, Walker, iv, 786 (1855).
Charnidas junctifera, Walker, xxxui, 334 (1865).
1g; Ne india (type): 25a, 2 8, Nolo Ells, one
Trevandrum. 4 2, 1 2, Ceylon.
AROA OCHRACEA.
Charnidas ochracea, Moore, Lep. Atk., p. 44 (1879).
1 Z,1 9, Nilgini Hills. 1 2, Calcutta (type).
AROA SIENNA.
Avoa sienna, Ampsn., Ill. Het., viii, p. 55, pl. 140, f. 29
(1891).
23,19, Nilgiri Hills (type). 4 3, Ceylon.
AROA SUBNOTATA.
Lacida subnotata, Walker, iv, 803 (1855).
5 $, 39, Ceylon, including the type #. 12, Nilgiri Hills.
AROA FLAVICOLLIS.
Crinola flavicollis, Leech, Entom., xxiii, p. 111 (1890).
6 f, Chang Yang, including the type Z. 2 9, Chia-kou-
ho, including the type &.
AROA ATRELLA.
Aroa atrella, Hmpsn., Moths, India, i, p. 489 (1892).
1g, Sikhim. 1 2, Khasia Hills.
The type, from Margharita, is in coll. Elwes.
AROA ATRESCENS.
Aroa atrescens, Hmpsn., Journ. Bo. N. H. Soc., xi, p. 294
(1897).
1 g, Khasia Hills (type).
AROA CINNAMOMEA.
Charnidas cinnamomea, Moore, Lep. Atk., p. 44 (1879).
Charnidas aurantiaca, Warren, P. Z. S., 1888, p. 296.
3g, Kunawar, including the type. 1 3, Attock (type
aurantiaca).
Old World Lymantriide in the National Collection. 455
AROA SAGRARA.
Aroa sagrara, Swinhoe, P. Z.8., 1885, p. 299, pl. 20, f. 13.
4 2, Belgaum, including the type. 2 2, 1 9, Jubbulpore.
1 9, Rajputana.
AROA CLARA.
Aroa clara, Swinhoe, P. Z.8., 1885, p. 299, pl. 20, f. 10.
2 f, 2 2, Bombay, including the type.
AROA SIMPLEX.
Orgyia simplex, Walker, xxxui, 325 (1865).
1 3g, 8. India (type). 2 3, Nilgiri Hills.
AROA XERAMPELINA.
Gynxphora cerampelina, Swinhoe, P.Z.8., 1885, p. 299, pl.
Pak tis fe),
1 g, 2 $, Poona, including the types. 1 2, Deccan.
1 f, Bombay. 1 2, Nilgiri Hills.
AROA FLAVEOFUSGCA.
Aroa flaveofusca, Swinhoe, Ann. Mag. N. H. (7), ix, p. 80
(1902).
1 f, Lawas, Borneo (type). 1 ¢, Sandakan.
AROA MIRMA, nov.
¢. Orange-yellow ; branches of antennze black ; head and thorax
brown ; fore-wings with a rather large brown spot at the end of the
cell, and dark brown irrorations forming a broad band at the apex ;
some paler brown spaces on the costa, base, and hinder margin, the
irrorations being less thick, leaving only a small place in the middle
of the wing clear ; hind-wings with the costal space broadly dark
brown ; under-side without any irrorations, except at the apices of
both wings, but the brown spaces more limited on the hind-wings,
there being merely a brown apical patch ; body and legs pale whitish-
yellow.
Expanse of wings 1 inch.
2 f, Sumatra (Lricson),
With the wings rather broad and short and Wee apices
rounded as in flaveofusca, Swinhoe, from Borneo,
456 Colonel C. Swinhoe’s Revision of the
AROA COMETARIS.
Aroa cometaris, Butler, Ann. Mag. N. H. (5), xix, p. 223
(1887).
1 ¢, Alu (type). 2 3, Guadalcanar Island.
AROA DISCALIS
Aroa discalis, Walker, iv, 792 (1855).
14,8. Africa (type). 13, Kilimanjaro. 1 g, Zomba.
22 2, Natal, Mashonaland and Zululand.
Kirby, p. 463, sinks dificilis, Walker, to this species, but
difficilis is a Dasychira.
AROA BISTIGMIGERA.
Aroa bistigmigera, Butler, P. Z. S., 1896, p. 847, pl. 42,
Lea.
3%, Nyassaland, including the type. 2 g, Zululand.
2¢, Port Ugowe. 2%, Basutoland. 2 ¢, Natal. 1 3,
Transvaal. 6 ¢, Neugia, Brit. E. Africa.
AROA DREGEI.
Orgyia dregei, Herr.-Schiff., Ausser-Eur. Schmett., 1, f. 114
(1854),
2 g, Cape.
AROA MELAXANTHA.
Orgyia melaxantha, Walker, xxxii, 324 (1865).
1 ¢, Cape (type).
AROA SUBSTRIGOSA.
Aroa substrigosa, Walker, iv, 794 (1855).
MA 7 Butlers wy ket. vp 4) pl, COmatae®
(1886).
Aroa pyrrhochroma, Walker, xxxui, 329 (1865).
28 2, 5 2, from parts of North India and China, including
both types.
Typical pyrrhochroma with its broad black outer marginal
band looks very distinct from typical swbstrigosa, which
has no band whatever, but I have received both forms and
any number of intergrades from the Khasia Hills, where
the species is quite common, and I am convinced that they
are both one and the same very variable form.
Old World Lymantriide in the National Collection. 457
AROA SOCRUS.
Gynzephora socrus, Geyer, Zutr., v, p. 12, ff 837, 838
(1837).
4 f, Java.
The “East Indies,” which Geyer gives as his locality,
undoubtedly meant the Dutch East Indies; his figures
correspond with the Javan form, which is quite different
from the Indian.
AROA RISORIA, nov.
¢. Antenne black, shafts white ; palpi and frons ochreous ; head,
body, and fore-wings pink-grey, with pale blackish-brown indistinct
streaks along the interspaces above and below the cell, and near the
hinder margin ; hind-wings brownish-grey, paler towards abdominal
margin, in one example pale yellowish-white and very uniform in tint ;
cilia of both wings dark brown-pink ; under-side paler, both wings
uniformly coloured, the apex of fore-wings with an obscure grey
space ; body and legs grey, strongly tinged with pink.
Expanse of wings 2%; inches.
3 f, Arjuno, Java, 3000 feet (Doherty).
Belongs to the socrws group.
AROA CHARAX.
Neurophana charax, Druce, Ann. Mag. N. H. (6), xvii, p.
352 (1896).
1 g, Nyassaland.
The type from E. Africa is in coll. Druce.
AROA OCHRACEATA.
Aroa ochraceata, Walker, xxxil, 327 (1865).
1 9, Natal (type). 4, Salisbury, Mashonaland. 2 9,
Zomba.
AROA SIGNATA.
Aroa signata, Walker, xxxii, 328 (1865).
6 9, Natal, including the type. 1 2, Port Ogowe.
Genus TrIA, Walker, iv, 803 (1855).
TEIA ANARTOIDES.
Teia anartoides, Walker, iv, 804.
5 g, Tasmania, including the type. 2 2, S.E. Australia.
458 Colonel C. Swinhoe’s Revision of the
TEIA PUSILLA.
Tcia pusilla, Butler, Ann. Mag. N. H. (5), ix, p. 88 (1882).
1 g, Melbourne (type). 7 2g, Sydney. 1 2, SE.
Australia.
Genus OrGyYIA, Ochs., Schmett. Eur., i111, p. 208 (1810).
Notolophus, Germ., Prodr., 11, p. 35 (1812).
Gynephora, Hiibner, Verz. bek. Schmett., p. 161
(1818),
Acyphas, Walker, iv, 797 (1855).
Micropterogyna, Ramb., Cat. Lep. Andalusie, p.
281 (1866).
Clethrogyna, Ramb., |. ¢.
Thylacogyna, Ramb.,1. ec.
Apterogynis, Guen., Stat. Sci. d’Eure et Loire, p. 78
(1876).
ORGYIA NIGROCROCEA.
Orgyia nigrocrocea, Walker, Journ. Linn. Soc. Lond., vi, p.
124 (1862).
4, Sarawak, including the type. 1 2, Matang, Borneo.
ORGYIA NIGRIPLAGA.
Orgyia nigriplaga, Swinhoe, Ann. Mag. N. H. (7), xii, p.
194 (1908).
1 ¢, N. Borneo (Fruhstorfer) (type). 1 2, Perissiu,
Borneo.
ORGYIA TURBATA.
Orgyia turbata, Butler, Trans. Linn. Soc. Lond. (2), 1, p.
560 (1879).
1 g, Malacca (type). 1 g¢, Province Wellesly. 1 4,
Tenasserim. 1 g¢, Heipaw, Burma.
ORGYIA GONOSTIGMA.
Bombyx gonostigma, Fabr., Syst. Ent., p. 585 (1775).
Orgy ia approximans, Butler, Trans. Ent. Soc., 1881, p. 10.
1 gf, Tokio (type approvimans). 1 g, Oiwake.
Besides many European examples.
ORGYIA NUCULA.
Orgyia nucula, Swinhoe, Ann. Mag. N. H. (6), xiv, p. 435
(1894).
Old World Lymantrudxe in the National Collection. 459
Orgyia nucula, Swinhoe, Trans. Ent. Soc., 1895, p. 7, pl.
IL Se IB
1%, Khasia Hills (type). 1 g, Fort Stedman, Shan
States.
ORGYIA LEECHI.
Orgyia prisca, Leech, Entom., xxiii, p. 111 (1890)
(preoce.).
Notolophus leecht, Kirby, Cat. Moths, i, p. 495 (1892),
2 ¢, Chang Yang, including the type. 10 3, various
parts of Western China.
ORGYIA SENICA.
Orgyva senica, Hmpsn., Journ. Bo. N. H. Soc., xiii, p. 234,
pl. B, f. 23 (1900).
1 g, Chitral (type).
ORGYIA THYELLINA.
Orgyia thyellina, Butler, Trans. Ent. Soc., 1881, p. 10, ¢.
: » leech iRs ZS: 1883;) p625, ple sl f° 7,
(hy Sh
1 f, Tokio, Japan (type). 3g, Yokohama. 1 2, Oiwake.
5 f, 6 2, marked Japan.
ORGYIA VIRIDESCENS.
Acyphas viridescens, Walker, iv, 798 (1855).
2 $, Ceylon, including the type.
ORGYIA DEWARA, nov.
¢. Antenne, body, and wings dark brown-pink ; branches of the
antenne black; thorax and fore-wings with many pale pinkish
markings very difficult to describe ; fore-wings crossed by a number
of dark brown very irregular bands, and with a submarginal series
of large brown spots on a pale ground, all very obscure ; hind-wings
without markings, of a uniform colour, much darker than the fore-
wing, nearly black ; cilia of both wings with ochreous-white tips ;
under-side much paler brown ; both wings uniform in colour and
without markings ; palpi, body beneath, and legs dull ochreous, the
legs with some dark brown stripes.
Expanse of wings 1,5 inches.
Hab. Kapaur, N. Guinea (Doherty).
The fore-wings are narrower than is usual in this
genus.
460 Colonel C. Swinhoe’s Revision of the
ORGYIA POSTICA.
Lacida postica, Walker, iv, 803 (1855).
Orgyia postica, Hmpsn., Moths, India, i, p. 437 (fig.)
(1892).
r es Pag., Zoologica, xxix, p. 40 (1900).
Orgyia ceylanica, Nietner, Ed. New Phil. Journ., xv, p. 34
(1862).
Orgyia ocularis, Moore, Lep. Atk., p. 44 (1879).
Orgyia ludekingit, Snellen, Tijd. v. Ent., xxii, p. 104, pl. 8,
f. 5 (1879):
One from Hong Kong, 1 Formosa, 2 Calcutta, including
Moore’s type of ocularis, 5 Ceylon, 1 Maulmein (type), 1
S arawak, 5 Java, 1 New Guinea, 1 Yorubaland, all males.
ORGYIA AUSTRALIS.
Orgyia australis, Walker, iv, 787, 2 (1855).
Orgyia canifascia, Walker, xxxii, 325, $ (1865).
2 2,1 9, Australia, including both types.
ORGYIA TISDALA, nov,
¢. Brown, with a slight pink tinge ; palpi on the inner sides,
body beneath, and legs very pale, nearly white ; wings with the
veins brown, rather prominent, rather thinly clothed, the costal and
outer marginal spaces of fore-wings dark brown ; a large dark brown
spot at the end of the cell.
Expanse of wings 34 inch.
3g, Arjuno, Java (including type), (Doherty). 2 3,
Sumatra (de Nicéville).
A very small example of the genus; the outer margins
of fore-wings much rounded.
Genus Lacuana, Moore, P. Z.8., 1888, p. 397.
LACHANA LADAKENSIS.
Lachana ladakensis, Moore, 1. ¢., p. 398.
ms “ Butler, Ill. Het., vii, p. 30, pl. 121, f. 6
(1889),
1 g, Ladak (type). 1 3, Skardo.
Genus Icta, Walker, iv, 922 (1855).
IcTA FULVICEPS.
Icta fulviceps, Walker, iv, 922.
2 ¢, Australia, including the type.
Old World Lymantriide in the National Collection. 461
Genus VARMINA, Moore, P. Z. S., 1888, p. 405.
VARMINA INDICA.
Gluphisia (?) indica, Walker, v, 1039 (1855).
1 gf, Simla. 2 7,19, Kangra. 1 fg, 19, Dharmsala.
1 g, 1 3, Umballa.
The type marked India is in Mus. Oxon.
Genus CASAMA, Walker, xxxui, 611 (1865).
CASAMA VILIS.
Euproctis vilis, Walker, xxxii, 348, f.
Casama indeterminata, Walker, xxxii, 611, 9.
1 3,1 9,58. India (types $2). 1 g, Nilgiri Hills. 2%
Ceylon. 14,2 2, Muscat, Arabia. 1 3, Somaliland. 1 ¢
Abyssinia.
Hind-wings with costal and sub-costal nearly touching
at the middle, 3 and 4, and 6 and 7 stalked.
?
?
Genus TuracipAs, Walker, v, 1027 (1855).
THIACIDAS POSTICA.
Thiacidas postica, Walker, v, 1028, 2.
Dirymonia (2) denotata, Walker, xxxui, 414, 2 (1865).
FHeterocampa (2) nigroscripta, Walker, xxxii, 423 9.
Cnethocampa (?) curvata, Walker, xxx1i, 429, 2.
Cnethocampa (2) basifurca, Walker, xxxii, 430.
12 ¢, 11 9, from various parts of India, including all the
types.
Costal and sub-costal touching close to base, 3 and 4, and
6 and 7 from the angles of the cell.
Genus Dasycuira, Steph., Ill. Brit. Ent. Haust., ii, p. 58
(1829).
Psalis, Hiibn., Zutr., 11, p. 19 (1823) (non descr.).
Olene, Hiibn., 1. c. (non deser.).
Arestha, Walker, iv, 805 (1855).
Melia, Walker, iv, 808 (preocc.).
Anaxila, Walker, iv, 810.
Argila, Walker, iv, 811.
Nioda, Walker, v, 1069 (1855).
Dediama, Walker, v, 1074.
Rilia, Walker, v. 1075.
462 Colonel C. Swinhoe’s Revision of the
Phineca, Walker, vii, 1746 (1856).
Ilema, Moore, Cat. Lep. E. I. C., ii, p. 341
(1859) (preocc.).
Anchyneura, Felder, Sitz. Akad. Wiss. Wien., xliii,
p- 331 (1861).
Janassa, Walker, Journ. Linn. Soc. Lond., vi, p.
135 (1862).
Thelde, Walker, 1. c., p. 139.
Boreconia, Walker, xxxii, 460 (1865).
Turriga, Walker, Char. Undescr. Lep. Het., p. 15
(1869).
Tearosoma, Felder, Reise Nov., pl. 100, f. 6 (1868)
(non descr.).
Cadrusia, Moore, Lep. Atk., p. 54 (1879).
Calliteara, Butler, Trans. Ent. Soc., 1881, p. 12.
Ludasychira, Mosch, Abh. Senck. Ges., xv., p. 75
(1889).
Pseudonotodonta, Mosch., 1. ¢., p. 76.
cura, Holland, Psyche, vi, p. 389 (1893).
Euproctidion, Holland, |. c., p. 414.
Notohyba, Holland, |. c., p. 434.
Thamnocera, Holland, 1. c., p. 454.
Bathmochtha, Karsch, Ent. Nachr., xxi, p.368 (1895).
SECTION I, WITHOUT DORSAL TUFTS ON ABDOMEN.
DASYCHIRA SECURIS.
Psalis securis, Hiibner, Zutr., 111, p. 9, ff. 291, 292 (1823).
Arestha antica, Walker, iv, 805 (1855).
Rigema falcata, Walker, xxxii, 437 (1865).
Rigema tacta, Walker, xxxii, 438.
Anticyra approximata, Walker, xxx1i, 440.
14 2, 13 92, from many parts of India, Ceylon, Java,
Australia, and Africa, including all Walker’s types.
DASYCHIRA OLEARIA.
Olene olearia, Swinhoe, P. Z. 8., 1885, p. 297, pl. 20, ff. 14,
is
1 Z, 1 9, Poona (types). 5 9, Nilgiri Hills.
DASYCHIRA FUSCA.
Anthora fusca, Walker, iv, 918, 7 (1855).
Boreconia subviridis, Walker, xxxii, 460, 2 (1865).
Old World Lymantriidz in the National Collection. 463
17,1 2, South Africa (types $2). 127,Cape. 43,19,
Knysna.
The type example of subviridis is a female, not a male as
stated by Walker.
DASYCHIRA CCERULEIFASCIA.
Thamnocera ceruletfascia, Holland, Psyche, vi, p. 469
(1893).
12, Ogove River (Holland). 67, 22, River Niger
(Ff. W. Sampson).
Holland’s type, a female, came from the Ogove River.
DASYCHIRA ROBUSTA.
Aecyphas (2) robusta, Walker, iv, 799 (1855).
1 Z, Sierra Leone (type).
DASYCHIRA DELICATA.
Notohyba delicata, Holland, Psyche, vi, p. 451, pl. 18, f. 4
(1893).
\ /
1 Z, Ogove River (Holland).
The type from Ogove River is in coll. Holland.
DASYCHIRA CITANA.
Utidava (?) citana, Schaus and Clements, Sierra Leone
Lep., p. 26, pl. i, £10 (1893).
1 Z, Sierra Leone.
DASYCHIRA PUNCTIFERA.
Erastria (?) punctifera, Walker, xu, 809 (1857).
1 f, Congo (type).
DASYCHIRA PECULIABRIS.
Mardara (2) peculiaris, Butler, Ann. Mag. N. H. (5), iv, p.
240 (1879).
1 3, Madagascar (type).
DASYCHIRA BASISTRIGA.
Phineca basistriga, Walker, vu, 1747 (1856).
1 2 without locality (type).
TRANS. ENT. SOC. LOND. 1905.—PART UI. (ocT.) 31
464 Colonel C. Swinhoe’s Revision of the
DASYCHIRA ANTICA.
Dediama antica, Walker, v, 1074 (1855).
Bathmochtha albilunulata, Karsch, Ent. Nachr., xxi, p. 368,
pl. 4, £ 12 (1895).
1 2, without locality (type).
Karsch’s type came from the Cameroons.
DASYCHIRA LIGNEA.
Nioda lignea, Butler, Ann. Mag. N. H. (5), iv, p. 241
(1879).
1 Z, Madagascar (type).
SECTION II, WITH DORSAL TUFTS ON ABDOMEN.
DASYCHIRA MENDOSA.
Olene mendosa, Hiibner, Zutr., u, p. 19 ff. 293, 294 (1823).
Antipha basalis, Walker, iv, 806 (1855).
Nioda fusiformis, Walker, v, 1070 (1855).
Rilia lanceolata, Walker, v, 1075.
Dasychira sawanta, Moore, Cat. Lep. E. I. C., ii, p. 340, ¢
(1859).
Dasychira divisa, Walker, xxxii, 368, ¢ (1865).
Dasychira basigera Walker, xxxii, 363, 2.
Rilia distinguenda, Walker, xxxil, p. 435, 2.
Olene basivitta, Walker, xxx, 436 9.
Turriga invasa, Walker, Char. Undescr. Lep. Het., p. 15,
(1869).
9 f, 13 9, from various parts of India and Ceylon, includ-
ing the types of lanceolata, fusiformis, and basivitta. 1 f
Singapore, 1 f 1 ¢ Borneo, 2 f1 2 Java, including the
type of savanta ; 1 f Christmas Isl., 4 26 2 Australia, includ-
ing the types of basalis, divisa, invasa, distinguenda and
basigera.
DASYCHIRA BASALIS.
Argila basalis, Walker, iv, 811 (1855).
1 fg, Sierra Leone (type).
DASYCHIRA ORIMBA.
Olene orimba, Swinhoe, Ann. Mag. N. H. (6), xiv, p. 435
(1894).
1 g, 1, Khasia Hills (types).
Old World Lymantride in the National Collection. 465
DASYCHIRA INVARIA.
Repena (?) invaria, Walker, vii, 1724 (1856).
Dasychira inclusa, Walker, vii, 1737.
Thelde patula, Walker, Journ. Linn. Soc. Lond., vi, p. 140
(1862).
Dasychira dalbergix, Moore, P. Z.8., 1888, p. 399.
1 2, Philippines (type). 8 3, 8 2, Java, including type
Zt of inclusa. 29,Sambawa, 2 2, 3 2, Dharmsala, includ-
ing type f of dalbergix. 22%, Bengal. 1 9, Hong Kong.
DASYCHIRA ASVATA,
Dasychira asvata, Moore, Cat. Lep. E. I. C., ii, p. 340
(1859).
1 Z, 1 9, Java (types). 1 2, Dinding Isl., Malaya.
DASYCHIRA FLAVIMACULA.
Dasychira flavimacula, Moore, P. Z.8., 1865, p. 804.
1 f, Bengal (type). 43,2 9, Darjiling. 1 3,1 2, Yatong,
Thibet.
The Yatong examples are not quite typical; the mark-
ings are practically the same, but the fore-wings are
shorter and the outer margin rounder, especially in the
female, the wings not being elongated, with a very oblique
outer margin, as in the Sikhim and Bengal examples.
DASYCHIRA RENDALLI.
Dasychira vendalli, Distant, Ann. Mag. N. H. (6), xx, p.
203 (1897).
3%, Mashonaland. 1 7, Natal.
The type from the Transvaal is in coll. Distant.
DASYCHIRA MUNICIPALIS.
Lelia municipalis, Distant, Ann. Mag. N. H. (6), xx, p. 200,
ai (1897):
Lacipa diffusa, Distant, |. ¢., 2.
13 3,6 2, Cape. 19, Natal. 2 92, Nyassaland.
The types from the Transvaal are in coll. Distant.
DASYCHIRA EXTORTA.
Dasychira extorta, Distant, Arn. Mag. N. H. (6), xx, p. 203
Gisam):
4.66 Colonel C. Swinhoe’s Revision of the
Lymaniria hera, Druce, Ann. Mag. N. H. (7), i, p. 209
(1898).
1 Z, Brit. E, Africa. 17, Natal. 5 4192, Mashonajand.
The type from the Transvaal is in coll. Distant, and the
type of hera from British E. Africa in coll. Druce.
DASYCHIRA NUBIFUGA.
Notohyba nubifuga, Holland, Psyche, vi, p. 451, pl. 18, f. 33
(1893).
2, Ogove River (Holland).
The type from the Ogove River is in coll. Holland.
DASYCHIRA MISERATA.
Llema miserata, Holland, Psyche, vi, p. 471, pl. 17, f. 3
(1893).
1 2, Ogove River (Holland). 1 9, Lokoja.
The type from Ogove River is in coll. Holland.
DASYCHIRA OCELLIFERA.
Occura ocellifera, Holland, Psyche, vi, p. 454, pl. 18, f. 5
(1893).
1 3, Ogove River (Holland).
The label on this example has got on it Notopriota
ocellifera, Holland, co-type, but I can find no reference to
any such genus, except on Holland’s plate; the genus does
not seem to have been described. The type from the Ogove
River is in coll. Holland.
DASYCHIRA MALIGNA.
Parorgyia maligna, Butler, Cist. Ent., ii, p. 17 (1882).
1 f, Madagascar (type).
DASYCHIRA PHASIANA.
Parorgyia phasiana, Butler, Cist. Ent., i, p. 17 (1882).
1 $, Madagascar (type).
DASYCHIRA CHALANA.
Dasychira chalana, Moore, Cat. Lep. E. I. C., 1, p. 339
(1859).
Lacida costiplaga, Walker, Journ. Linn. Soc. Lond., vi, p.
126 (1862).
Old World Lymantriide in the National Collection. 467
47%, 3 9, Java, including the type 9.
The type of costiplaga f from Sarawak is in Mus. Oxon.
DASYCHIRA MISANA.
Dasychira misana, Moore, Cat. Lep. E. I. C., ii, p. 340, pl.
9a, t 271859):
2 f, Java, including the type.
_ DasYCHIRA WHITEI.
(Ecura whitei, Druce, Ann. Mag. N. H. (7), i, p. 209 (1898).
1 2, Old Calabar. 2 ¢, Natal. 1 ¢, Durban.
The type from W. Africa is in coll. Druce.
DASYCHIRA GOODII.
(Eeura goodii, Holland, Psyche, vi, p. 390, pl. 9, ff. 3, 4
(1893).
1 f, Ogove River (Holland). 3 9, Sierra Leone.
The types from the Ogove River are in coll. Holland.
DASYCHIRA COSTALIS.
Melia costalis, Walker, iv, 808 (1855).
3 2, 7 2, Java, including the type 2. 1 3, Heipaw,
Burma.
DASYCHIRA CROMPTONI, nov.
¢. Antenne grey, shafts white ; palpi, frons, and thorax greyish-
ochreous ; fore-wings ochreous fawn colour; two black subbasal
marks, a brown obscure spot in the cell, a white lunule at the end
with a white dot above and another below it, an antemedial semi-
dentate erect brown line, a postmedial brown band from centre of
hind margin straight to the costa one-fourth from apex, where it
curves in; within this band there isa very fine semi-dentate blackish
line with pale points; an oblique row of blackish-brown spots
parallel to the second line ; the lower portion of the space between
the first line and the band brighter ochreous than the rest of the
wing; hind-wings pale ochreous-grey, without markings ; under-side
of wings uniform ochreous-grey, with an oblique grey line in the disc
of each wing ; body and legs ochreous ; tarsi with black spots.
Expanse of wings 1,3, inches.
1 2, Old Calabar (Crompton) (type). 2 2, River Niger
(Sampson).
The Niger examples are without any ochreous tinge,
but the markings are identical.
468 Colonel C. Swinhoe’s Revision of the
DASYCHIRA TENEBROSA.
Dasychira tenebrosa, Walker, xxxii, 361 (1865).
1 fy) 9, Dalhousie, 5 45 Yatong, Ehibet.| Aras =,
Darjiling, including the type. 5 ¢, W. China.
DASYCHIRA BHANA,
Dasychira bhana, Moore, P. Z. §., 1865, p. 804.
4 ft, Darjiling, including the type.
The same size as tenebrosa, but can at once be distin-
guished by its large dull ochreous spot at the end of the
cell of fore-wings, instead of the black stigma of that
species.
DASYCHIRA NIGRITULA.
Dasychira nigritula, Walker, xxxii, 360 (1865).
1 3, 2 9, N. India, including the type.
Much larger than the two preceding forms, the ochreous
spot at the end of the cell very large, in some examples in
the form of a patch, prominent in fresh examples.
DASYCHIRA FEMINULA.
Mardara feminula, Hmpsn., Il. Het., viii, p. 58, pl. 141, ff 1,
7 (1891).
6 3, 7 9, Nilgini Hills, including the type.
This is the South India form of the bhana group; the
great difference in the female is quite enough to differ-
entiate it; the male is as large as migritula, and the
ochreous markings on the transverse bands of the fore-
wings are more distinct and conspicuous.
DASYCHIRA CHINENSIS, nov.
4, @. Greyish-brown ; palpi black above ; antennz with whitish
shafts ; fore-wings variegated with greyish-white in both sexes, more
especially on the outer half; four transverse, indistinct, brown lines,
subbasal and medial both sinuous, discal dentated and recurved,
submarginal line lunular, followed by a marginal row of lunules
with whitish inner parts, and brown spear-shaped marks pointing
inwards; hind-wings brown, with indications of a darker discal
band; cilia of both wings ochreous with brown patches ; under-side
pale olive-grey ; a double discal thin brown band across both wings ;
a large brown lunular mark at the end of the cell of hind-wings,
cilia as above.
Expanse of wings g 44mm., 2? 56 mm.
Old World Lymantrudx in the National Collection. 469
1 2,1 2, Moupin (types). 13,1 9, Ta-chien-lu, 10,000
feet (Pratt). 1 3, Kwei-chow.
Belongs to the bhana group, but is different from any of
the Indian forms; its coloration is peculiar, and the trans-
verse lines on fore-wings are differently shaped.
DASYCHIRA MAGNALIA.
Dasychira magnalia, Swinhoe, Ann. Mag. N. H. (7), xii, p.
198 (1903).
1 ¢, 1 2, Khasia Hills (types).
DASYCHIRA NIGRA.
Dasychira nigra, Hmpsn., Journ. Bo. N. H. Soc., xiii, p. 416,
pl 2a. 1941900):
1 2, Sikhim (type).
DASYCHIRA GWELILA, nov.
¢. Antenne black; palpi, frons, and head brown; body and
fore-wings very dark grey, nearly black, caused by very minute
black irrorations closely packed on a grey ground colour ; veins more
or less prominent ; faint indications of antemedial, postmedial, and
discal transverse bands, and a submarginal series of more prominent
black spots close to the outer margin; hind-wings white, without
markings ; under-side of fore-wings grey, becoming white hindwards ;
hind-wings white, with no markings; body and legs whitish, the
latter with some brown stripes.
Expanse of wings 1,', inches.
Hab. GWELIL, Brit. E. Africa (Betton).
DASYCHIRA ALBIBASALIS.
Llema albibasalis, Holland, Psyche, vi, p. 470 (1898).
1 f, Ogove River (Holland).
The type from Ogove River is in coll. Holland.
DASYCHIRA GABUNICA.
Euproctidion gabunica, Holland, Psyche, vi, p. 414, pl. 10,
f, 19 (1893).
1 g, Ogove River (Holland).
The type from the Ogove River is in coll. Holland.
470 Colonel C. Swinhoe’s Revision of the
DASYCHIRA GONOPHORA.
Ilema gonophora, Holland, Psyche, vi, p. 470, pl. 17, f. 1
(1893).
Lelia curvivirgata, Karsch, Ent. Nachr., xxi, p. 372, pl. 4,
f. 6 (1895).
1 f, Ogove River (Holland). 1 9, Cameroons.
The type from the Ogove River is in coll. Holland.
DASYCHIRA ALBINOTATA.
Thamnocera albinotata, Holland, Psyche, vi, p. 469 (1898).
1 f$, Ogove River (Holland).
The type from the Ogove River is in coll. Holland.
DASYCHIRA STRIATA.
Notohyba striata, Holland, Bee vi, p. 452, pl. 17, f. 28
(1893).
1 g, Ogove River (Holland).
The type from the Ogove River is in coll. Holland, very
close to if not identical with albinotata,
DASYCHIRA BRUNNEICOSTA.
Tlema brunneicosta, Holland, Psyche, vi, p. 471, pl. 17, f. 10
(1893).
1 f, Ogove River (Holland).
The type from the Ogove River is in coll. Holland.
DASYCHIRA HORRIDA, nov.
¢. Dark brown; thorax blackish-brown in front ; abdomen with
the first and last two segments smeared with white ; fore-wings
tinged with olive; an inner, outwardly curved, transverse, rather
sinuous blackish-brown line; a lunule at the end of the cell; a
nearly straight and erect discal line; marginal spots black, flecked
with white ; hind-wings a little paler ; marginal line dull ochreous,
otherwise without markings; under-side uniform pale brown; a
lunule at the end of each cell, a discal thin even band across both
wings, and indications of a similar band between that and the
margin.
Expanse of wings 1 inches.
Hab. OLD CALABAR (Crompton).
ee ee
Old World Lymantriide in the National Collection. 471
DASYCHIRA PLAGIATA.
Anaxila plagiata, Walker, iv, 810 (1855).
1 f, without locality (type).
DASYCHIRA VARIA.
Dasychira varia, Walker, iv, 868 (1855).
» Swinhoe, Cat. Het. Mus. Oxon., i, p. 219,
note (1892).
Dasychira maruta, Moore, Cat. Lep. E. I. C., ii, p. 339
(1859).
1 2, Darjiling (type maruta). 1 9, Subathu. 6 %,
Dalhousie.
DASYCHIRA LUNULATA.
Dasychira lunulata, Butler, Ann. Mag. N. H. (4), xx, p. 403
(1877).
i Butler, Ill. Het., ii, pl. 24, f. 8, oy
Dasi yohir a solitaria, Staud., Rom. sur Lep., iii, pl. 2,
2 (1878).
Dasychira acronycta, Oberth., Etud. d’Ent., v, p. 35, pl. 5,
tine, (LSSih):
3 2, 6 2, Japan, including the type ¢.
DASYCHIRA ALBESCENS.
Dasychira albescens, Moore, Lep. Atk., p. 59 (1879).
3,1 9, Darjiling.
The type from Darjiling is in coll. Staudinger.
DASYCHIRA CINCTATA.
Dasychira cinctata, Moore, Lep. Atk., p. 59 (1879).
ig 2 Darling:
The type from Darjiling is in coll. Staudinger.
DASYCHIRA AMPLIATA.
Dasychira ampliata, Butler, Ann. Mag. N. H. (5), ii, p. 460
(1878).
1 2, Madagascar (type).
DASYCHIRA ANGULATA.
Dasychira angulata, Hmpsn., Trans. Ent. Soc., 1895, p. 292.
ig, cikhim (ype). 1 2) Upper Burma. 1. 7, 2 9;
Khasia Hills.
472 Colonel C. Swinhoe’s Revision of the
I put the Khasia Hill specimens very doubtfully with
this form; they are white instead of dull ochreous, and
the lines seem to be different; but the lines are not very
distinct in the type, and we must await more Sikhim
examples.
DASYCHIRA PSEUDABIETIS.
Calliteara pseudabietis, Butler, Cist. Ent., iii, p. 118 (1885).
Calliteara abietis, Leech, P. Z.8., 1888, p. 631.
Dasychira pryert, Butler, l.c., p. 119.
Dasychira pudica, Staud., Rom. sur Lep., ii, p. 204 (1887).
Dasychira modesta, Kirby, Cat. Moths, 1, p. 483 (1892).
6 f, 4 2, Japan, including both of Butler’s types. 1 2,
1 $ from Amur.
Staudinger’s type came from the Amur; the name
pudica was preoccupied in the genus, and therefore Kirby
re-christened it modesta ; but the Amur and Japan examples
are not separable.
DASYCHIRA POSTFUSCA.
Dasychira postfusca, Swinhoe, Trans. Ent. Soc., 1895, p. 9,
Pl pr 2:
1 g, Khasia Hills (type).
DASYCHIRA THWAITESI.
Dasychira thwaitesi, Moore, Lep. Ceylon, 1, p. 98, pl. 116,
f. 1, la, b (1883).
Dasychira pudica, Moore, |. ¢., 11, p. 538 (1887).
1 g, 1 9, Ceylon (types), 1 3,1 9, Trevandrum. 1 32;
Nilgiri Hills.
The type of pudica is a dwarf female.
DASYCHIRA GROTEI.
Dasychira grote’, Moore, Cat. Lep. E. I. C., 11, p. 338
(1859).
Dasychira kausalia, Moore, P. Z. §., 1879, p. 401.
Dasychira varia, 2 only, Walker, iv, 868 (1855).
1 f, Kasaoli (type kausalia, ). 1 3, Subathu. 1 3,14,
Dharmsala. 1 ¢,1 9, Kangra. 1 9, N. India (type). 14,
Himalayas. 2 9, Darjiling. . 1 9, Calcutta. 1 9, Moupin,
W. China.
Old World Lymantrude in the National Collection. 473
DASYCHIRA HORSFIELDII.
Arctia horsfieldit, Saunders, Trans. Ent. Soc., 1851, p. 162,
play 2,, tie 1.2.
Dasychira arga, Moore, Cat. Lep. E. I. C., 11, p. 339 (1859).
Dasychira longipennis, Walker, Journ. Linn. Soc. Lond., vi,
p. 13 (1862).
Dasychira nilgirica, Ampsn., Il. Het., viii, p. 58, pl. 114,
ff. 13, 14 (1891).
1 2, 2 2, Nilgiri Hills, including Hampson’s types. 3 2,
29,Ceylon. 4 3,19, Java, including Moore’s types. 1 2,
2 9, Singapore.
Saunders’ and Walker’s types should be in Mus. Oxon.,
but I have been unable to find them; Walker's type was a
female, and his description well agrees with this form.
DASYCHIRA GEORGIANA.
Dasychira georgiana, Fawcett, Trans. Zool. Soc., xv, p. 314,
pl. 49, ff. 19, 20, 21 (1900).
2g, £ 2, Natal” 1 2) Wransvaal!
DASYCHIRA CERIGOIDES,
Janassa cerigoides, Walker, Journ. Linn. Soc. Lond., vi, p.
135 (1862).
a 5 Swinhoe, Cat. Het. Mus. Oxon., i, p. 297,
pled; & O1892):
1 2, Borneo.
The type $ from Sarawak is in Mus. Oxon.
DASYCHIRA GRANDIDIERI.
Calliteara grandidiert, Butler, Cist. Ent., 111, p. 14 (1882).
1 $, Madagascar (type).
DASYCHIRA VIBICIPENNIS.
Dasychira vibicipennis, Butler, Ann. Mag. N. H. (5), iv,
p. 239 (1879).
1 f, 1 2, Madagascar (types).
DASYCHIRA GENTILIS.
Dasychira gentilis, Butler, Ann. Mag. N. H. (5), iv, p. 239
(1879).
2 9, Madagascar, including the type.
474 Colonel C. Swinhoe’s Revision of the
DASYCHIRA ACRISIA.
Deiopeia (?) acrisia, Plotz, Stett. Ent. Zeit., xli, p. 83 (1880).
Dasychira acrisia, Kirby, Cat. Moths, i, p. 484 (1892).
Dasychira crausis, Druce, P. Z.S., 1884, p. 228, pl. 17, £5, 9.
1 f, Acera. 1 3, Old Calabar.
The type came from W. Africa, the type of crausis from
the Lower Niger.
DASYCHIRA ELEGANS.
Calliteara elegans, Butler, Cist. Ent., 11, p. 13 (1882).
1 f, Madagascar (type).
DASYCHIRA MGRENS.
Calliteara merens, Butler, Cist. Ent., 11, p. 14 (1882).
1 g, 1 9, Madagascar (types).
DASYCHIRA PUMILA.
Calliteara pumila, Butler, Cist. Ent., iii, p. 16 (1882).
2 ¢, Madagascar, including the type.
DASYCHIRA ARGENTATA.
Dasychira argentata, Butler, Trans. Ent. Soc., 1881, p. 12.
5 t, 1 9, Japan, including the type &.
DASYCHIRA ASPERSA.
Tearosoma aspersa, Felder, Reise Nov., pl. 100, f. 6 (1868).
2 2, Natal.
Felder’s type, a female without locality, is in coll.
Rothschild.
DASYCHIRA ROTUNDATA.
Teara rotundata, Walker, iv, 851 (1855).
1 gf, Tasmania (type). 1 3, Australia. 1 g, Moreton
Bay.
DASYCHIRA SUBLUTESCENS.
Dasychira sublutescens, Holland, Psyche, vi, p. 433 (1898).
1 2, Ogove River (Holland).
The types ¢ ¢ from the Ogove River are in coll. Holland.
Old World Lymantriide in the National Collection. 475
DASYCHIRA SUBFLAVA.
Anaxila subflava, Walker, iv, 918 (1855).
1 f, Ashanti (type).
The type has a female body put on it wrong side forward ;
it is a mere fragment, but was evidently once a Dasychira.
DASYCHIRA DIFFICILIS.
Aroa diffcilis, Walker, xxxii, 328 (1865).
3 2, Natal, including the type.
Walker described this type example as a male.
DASYCHIRA ENOS,.
Aroa (?) enos, Druce, Ann. Mag. N. H. (6), xvui, p. 353
(1896).
1 9, Sapele, Niger River.
The type, a female, from Old Calabar, is in coll. Druce.
DASYCHIRA STRIGATA.
Dasychira strigata, Moore, Lep. Atk., p. 58 (1879).
- fs Hmpsn., Moths, India, i, p. 449 (1892).
is » swithoe, Ann, Mag, N: . (7), xu,
p- 197 (1903).
Dasychira niveosparsa, Butler, Il. Het., v, p. 59, pl. 91, f. 7
(1881).
1 2, Masuri (type). 1 9, Darjiling (Butler’s type). 1 3,
Khasia Hills.
I have it also from the last-named locality; the male
does not differ much from the female; the transverse
duplex bands on the fore-wings are less sinuous, and the
abdomen and hind-wings are more ochreous. Expanse of
wings 2), inches.
DASYCHIRA VIOLA.
Calliteara viola, Butler, Ann. Mag. N. H. (5), iv, p. 240
(1879).
2 f, Madagascar, including the type.
DASYCHIRA CHLOROPTERA.
Dasychira chloroptera, Hmpsn., Moths, India, i, p. 450
(1892).
1 Z, Khasia Hills (type).
476 Colonel C. Swinhoe’s Revision of the
DASYCHIRA VIRESCENS.
Cadrusia virescens, Moore, Lep. Atk., p. 54, pl. 3, f. 16
(1879).
Dasychira virescens, Hmpsn., Moths, India, i, p. 451 (1892).
2 2, Darjiling, including the type.
DASYCHIRA MASCARENA.
Dasychira mascarena, Butler, Ann. Mag. N. H. (5), ui, p.
294 (1878).
Dasychira cangia, Druce, P. Z.8., 1887, p. 674.
¥ = Holland, Psyche, vi, p. 567, pl. 10, f. 12
(1893).
1 Z, 1 9, Madagascar (types).
The types f 2 of cangia from Delagoa Bay are in coll.
Druce; I do not see how they can be separated from
Butler’s species.
DASYCHIRA PASTOR.
Calliteara pastor, Butler, Cist. Ent., iii, p. 15 (1882).
1 Z,1 2, Madagascar (types).
DASYCHIRA CATOCALOIDES.
Mardara catocaloides, Leech, Trans. Ent. Soc., 1899, p. 126.
1 7, Moupin (type). 1 2, Ta-chien-lou.
DASYCHIRA PRASINA.
Calliteara prasina, Butler, Cist. Ent., iii, p. 16 (1882).
2 ¢, Madagascar, including the type.
DASYCHIRA BARUNA.
Somena baruwna, Moore, Cat. Lep. E.I. C., ii, p. 346 (1859).
x » Kirby, Cat. Moths, i, p. 55 (1892).
Dasychira viridis, Druce, Aun. Mag., N. H. (7), iii, p. 470
(1899).
3 2, Java (Horsjield), including the type. 1 7, Penang
(Flower).
The type of viridis came from Perak and is in Mus.
Druce.
Old World Lymantrudz in the National Collection. 477
DASYCHIRA GNAVA, nom. nov.
Pseudonotodonta virescens, Moschler, Abh. Senck. Ges., xv,
p- 77, f. 6 (1889) (preocc.).
1 f, Old Calabar (Sampson).
Moschler’s type, a female, came from Aburion the Gold
Coast: it cannot be generically separated from Dasychira.
DASYCHIRA INFIMA.
Somera infima, Holland, Psyche, vi, p. 479, pl. 17, f. 29
(1893).
1 f, Ogove River (Holland).
The type 2 from Ogove River is in coll. Holland.
DASYCHIRA REMOTA.
Dasychira (?) remota, Druce, P. Z. S., 1887, p. 675.
4 9, Niger River.
The type, a female, from Gambia is in coll. Druce.
DASYCHIRA VARIEGATA.
Dasychira variegata, Holland, Psyche, vi, p. 568, pl. 18,
f. 13 (1893).
1 2, Ogove River (Holland).
The types 7 2, from the Ogove River are in coll. Holland.
SECTION III, ABDOMEN CLOTHED WITH LONG HAIR AND
WITHOUT DORSAL TUFTS.
DASYCHIRA COMPLICATA.
Dasychira complicata, Walker, xxxii, 365 (1865).
Trisula pustulifera, Walker, xxxii, 576.
49,29, Darjiling. 17%, Sikhim. 2 g,N. India (types).
SECTION IV, ABDOMEN SMOOTHLY SCALED, WITHOUT DORSAL
TUFTS ; WINGS VERY BROAD; FEMALE WITH BRANCHES
OF ANTENNA UNUSUALLY LONG.
DASYCHIRA LINEATA.
Lymantria lineata, Walker, iv, 875 (1855).
1 9, Silhet, including the type. 1 2, N. India.
The type was described as a male; but the male of this
species is unknown to me.
478 Colonel C. Swinhoe’s Revision of the
Genus MunycurvylA, Walker, xxxii, 395 (1865).
MUNYCHRYIA SENICULA.
Munychryia senicula, Walker, xxxii, 396.
1 $, Moreton Bay (type).
Genus ANEPA, nov.
Palpi porrect, slightly bent downwards ; frons, head, and thorax
covered with long hair ; abdomen with dorsal tufts on each segment ;
antennee bipectinate to the tips, the branches long, ciliated, and with
bristles at their ends; wings shaped as in Dasychira of the Olene
group; fore-wings with veins 7, 8, and 9 stalked, 10 and 11 co-
incident and form the cell; hind-wings with veins 6 and7 on a long
stalk ; hind-legs with two pairs of spurs.
Type A. fusca, Walker (Acyphas).
ANEPA FUSCA.
Acyphas fusca, Walker, iv, 798 1855),
2 ia Kirby, Cat. Moths, i, p. 472 (1892).
2 f, Australia, including the type. 1 ¢, Tasmania.
Kirby has left out from his genus Acyphas the type
of the genus viridescens, Walker, from Ceylon, which is
an Orgyia.
ANEPA FULVICEPS.
Charnidas fulviceps, Walker, iv, 797 (1855).
4 7, Tasmania, including the type.
Genus Urocoma, Herr.-Schaff., Ausser-Eur. Schmett., 1
p. 82 (1855).
UROCOMA LIMBALIS.
Urocoma limbalis, Herr.-Schatf., 1. c., f. 389.
4g, 19, Sydney.
UROCOMA BALIOLALIS.
Urocoma baliolalis, «Swinhoe, Cat. Het. Mus. Oxon., 1,
pelo. pl6, ©.4 @s92):
1g, Moreton Bay. 1 2,29, Australia. 1 $, Sydney.
The type f from Australia is in Mus. Oxon.
Old World Lymantrudx in the National Collection. 479
UROCOMA MARGINALIS.
Trichetra marginalis, Walker, iv, 845 (1855).
1 f, Tasmania (type), 1 2%, Australia.
Genus ABYNOTHA, nov.
¢. Antenne bipectinated to the tips, the branches long, with
cilia and with spines at their ends; palpi upturned, thickly clothed,
the last joint short; fore-wings with vein 3 a little before end of
cell, 4 from end, and 5 from a little above it; 6 from the upper
angle ; 7, 8 stalked, 9 from 10, anastomosing with 7 and 8 to form
the areole ; hind-wings with veins 3, 4, and 5 as in fore-wings, 6
and 7 from upper angle of cell.
Types A. preussi, Mab. (Liparis).
ABYNOTHA PREUSSI.
Tiparis (?) preusst, Mab., Vuill. Nov. Lep., i, p. 57, pl. 9,
f. 5 (1892).
Lymantria (?) preussi, Aurivillius, Ent. Tidsk., xii, p. 194,
(fig.) (1892).
Phxgorista (?) preussi, Kirby, Cat. Moths, i, p. 918 (1892).
4 f from the Cameroons.
Genus PacuycispiA, Butler, Cist. Ent., iui, p. 11 (1882).
PACHYCISPIA PICTA.
Pachycispia picta, Butler, l.c., p. 12.
3 ¢, Madagascar, including the type.
Genus CiruNA, Walker, v, 1172 (1855).
Baryaza, Moore, Lep. Atk., p. 45 (1879).
CIFUNA LOCUPLES.
Cifuna locuples, Walker, v, 1173.
1 9, Silhet (type). 1g, N.E. Bengal. 2 ¢, Khasia Hills.
62,39, Japan. 1 fg, Gensan. 1 gf, Chang Yang. 4 f,
iS Acar
The Amur examples are very dark.
CIFUNA BIUNDULANS.
Cifuna binndulans, Hmpsn., Journ. Bo. N. H. Soce., xi,
p- 294 (1897).
1 f, Khasia Hills (type).
TRANS. ENT. SOC. LOND. 1903.—PART UI. (OCT.) 32
480 Colonel C. Swinhoe’s Revision of the
Genus HETERONYGMIA, Holland, Psyche, vi, p. 414 (1893).
HETERONYGMIA RHODAPICATA.
Heteronygmia rhodapicata, Holland, |. ¢., p. 416, pl. 10, f. 15.
1 2, Ogove River (/olland).
The type ¢ from the Ogove River is in coll. Holland.
HETERONYGMIA CHISMONA, nov.
g. Palpi covered with crimson hairs; antenne grey; head,
thorax, and fore-wings pale fawn colour, covered with very minute
grey atoms; three very indistinct transverse grey upright bands;
hind-wings white, without markings ; abdomen above and below
pale crimson ; under-side of wings white without markings ; pectus
and legs yellowish-grey.
Expanse of wings 1,5 inches.
2 g, Aburi, Gold Coast, 1880 (Carter).
Genus Lomaponta, Holland, Psy che, vi, p. 417 (1893).
LOMADONTA ERYTHRINA.
Lomadonta erythrina, Holland, |. ¢., pl. 10, f. 1.
1 g, Ogove River.
The types f 2 from the Ogove River are in coll. Holland.
LOMADONTA JOHNSTONI, nov.
¢. Antennz ochreous ; palpi, frons, head, and thorax with the
ground colour covered with crimson - hairs ; fore-wings ochreous,
washed with crimson, the costal and outer margins dark red ; two
semi-hyaline indistinct whitish spots in the lower disc ; the entire
wing covered with well-separated red striz ; a brownish line from
the costa a little beyond the middle to the upper whitish spot, where
it is outwardly angled, then straight down to the hinder margin ;
a submarginal line of brown lunules ; hind-wings white tinged with
crimson, especially towards the outer margin, which is dull crimson ;
cilia of fore-wings purplish, of the hind-wings crimson ; under-side
whitish ochreous, crimson-tinged, with the marginal lines dark ; a
dark lunule at end of cell of fore-wings, a spot at end of cell of
hind-wings ; pectus and fore-legs covered with crimson hairs.
Expanse of wings 14 inches.
Hab. Conco Forest, July, 1900 (H. H. Johnston).
Allied to LZ. erythrina, Holland, the type of the genus
from Ogove River.
Old World Lymantriide in the National Collection. 481
Genus L&vapia, Butler, Ann. Mag..N. H. (5), iv, p. 238
(1879).
LALAPIA NOTATA.
Lelapia notata, Butler, |. ¢.
6 7, 1 2, Madagascar, including the type @.
Genus ACLONOPHLEBIA, Butler, P. Z. S., 1898, p. 428.
ACLONOPHLEBIA FLAVINOTATA.
Aclonophlebia flavinotata, Butler, l.c., pl. 32, f. 8.
2 f, Brit. E. Africa, including the type.
Genus NUMENOIDES, Butler, Ann. Mag. N. H. (5), iv,
p- 238 (1879).
NUMENOIDES GRANDIS.
Numenoides grandis, Butler, 1. ce.
2 2, Madagascar, including the type.
Genus Maracuitis, Hmpsn., Trans. Ent. Soc., 1895, p. 291.
MALACHITIS MELANOCHLORA.
Malachitis melanochlora, Hmpsn., |. ec.
. _ Dudgeon, Journ. Bo. N. H. Soc.,
xi, p. 413, pl. 1, f. 7 (1900).
1 f, Bhutan (type).
Genus ENomE, Walker, iv, 883 (1855).
ENOME AMPLA.
Enome ampla, Walker, iv, 883.
4 ft, Ceylon, including the type. 3 #, Calcutta. 1 2,
Malabar. 3, Bangalore. 1 ¢, Burma.
ENOME INCERTA.
Lymantria incerta, Walker, iv, 880 (1855).
1 , N. India (type). 2 3, Kangra. 1 2, Jubbulpore.
5 2, Nilgiri Hills.
I cannot find the type of Lymantria aryama, Moore,
from Canara, which Hampson puts as a synonym to this
species.
482 Colonel C. Swinhoe’s Revision of the
ENOME DETERSA.
Lymantria detersa, Walker, xxxui, 365 (1865).
Enome detersa, Swinhoe, P. Z. 8., 1885, p. 300, pl. 21,
ff. 3, 4.
Lymaniria costalis, Walker, xxxii, 365.
2 4,8. India (types of both.) 7 3, 19, Mhow. 9 2,
Poona. 4, Belgaum.
The locality for detersa in Walker's Catalogue is entered
as Mauritius, the insect from the Entomological collection ;
but this is a mistake: the example came from the East
Indian Museum, and the locality label on it is India. This
form belongs entirely to South India; I have bred many
specimens from larvze in Poona.
ENOME OBFUSCATA.
Lymantria obfuscata, Walker, xxxui, 367 (1865).
1 9, N. India (type). 2¢ N.W. Himalayas. 9 3, Murree.
1 Z, Kangra. 1 3, Campbellpore, Punjab. 1 9, Kalu.
1%, Sultanpore. 2 2, Karis, Cashmir. 1 f, Skardo.
Genus LyMANTRIA, Hiibner Verz. bek. Schmett., p. 160
(1818).
Liparis, Ochs., Schmett. Eur., ii, p. 186 (1810)
(preoce.).
Porthetria, Hiibner, |. ¢.
Hypogymna, Steph., Il. Brit. Ent. Haust., 11, p. 55
(1828).
Psilura, Steph., |. ¢., p. 57.
Pegella, Walker, xxxv, 1922 (1866
Nagunda, Moore, Lep. Atk., p. 53
Barhona, Moore, 1. ¢., p. 55.
)
(1879).
LYMANTRIA MCRENS.
Dasychira (?) merens, Felder, Reise Nov., pl. 99, f. 4
(1868).
1 f, 1 2, Ceylon.
The type from Ceylon is in coll. Rothschild.
LYMANTRIA DISPAR.
Bombyx dispar, Linn. Syst. Nat., x, p. 501 (1758).
Porthetria umbrosa, Butler, Trans. Ent. Soc., 1881, p. 10.
2 4,3 2, Japan, including Butler's types.
Many examples from different parts of Europe.
Old World Lymantriide in the National Collection. 483
LYMANTRIA JAPONICA.
Liparis dispar, var. japonica, Motsch, Etud. d’Ent., 1860,
p. 31.
Porthetria hadina, Butler, Trans, Ent. Soc., 1881, p. 11.
8 2,5 2, Japan, including Butler’s types. 4 7,1 2,
ee
Though closely related to dispar it is a uniformly
distinct form.
LYMANTRIA EURYDICE.
Porthetria ewrydice, Butler, Cist. Ent., ii, p. 118 (1885).
Dasychira amata, Stand.,Rom.sur Lep., iii, p. 206, pl. 12,
f. 2 2 (1887).
1 7, Choyama (type). 1 2,1 9, Japan.
Staudinger’s type came from the Amur.
LYMANTRIA ATLANTICA.
Liparis atlantica, Ramb., Faune Ent. Andal., ui, pl. 15,
f. 7 (1842).
1 7, N. Africa.
LYMANTRIA FUMIDA.
Lymantria fumida, Butler, Ann. Mag. N. H. (4), xx, p. 402
(1877).
» Butler, TH, Het., 11, pl. 24, f. 4 2 (1878).
Lymantria sinica, Moore, P.Z. S., 187 9, p. a
4 7,4 9, Japan, including the types. t, N. China
(type sinica). 2 7, Formosa.
LYMANTRIA RHODINA.
eas rhodina, Walker, xxxii, 366 (1865).
= Sean Cat. Het. Mus. Oxon., 1,
p. "222 (note) (1892).
1 7, India (type).
LYMANTRIA CEREBOSA, nov.
¢d, 2. Antenne orange-ochreous ; palpi and frons dark brown
with some white hairs ; head, body, and fore-wings dark olive-grey,
irrorated with white atoms ; markings darker grey and very indis-
tinct ; a lunular double line at end of cell, and four transverse
duplex lines, more or less sinuous but erect, subbasal, antemedial,
484 Colonel C. Swinhoe’s Revision of the
discal, and submarginal ; whitish lunules on the outer margin ;
hind-wings dirty white or pale greyish-white ; a grey spot at the
end of the cell; under-side pale grey ; a lunular mark at the end of
each cell; an indistinct and incomplete grey discal straight band
across both wings ; tarsi with whitish spots.
Expanse of wings, ¢ 115, @ 2 inches.
Hab. Soon, Simla (Reed).
LYMANTRIA MOESTA, nov.
¢, @. Palpiand antenne brown ; head, thorax, and fore-wings
dark grey ; two transverse darker grey lines on the fore-wings, one
ante-medial, angled outwards above the middle and crenulated, the
other discal, even with the margin, rather close to it, and semi-
dentate ; a grey lunular mark at the end of the cell ; all these very
indistinct ; abdomen and hind-wings paler grey, tinged with pink,
without markings; under-side uniform pale grey, no markings.
Expanse of wings, ¢ 13, 9 11% inches.
1 2, Kasaoli (Reed). 1 9, Kangra Valley (Dudgeon).
LYMANTRIA FURVA.
Ocneria furva, Leech, P. Z. 8., 1888, p. 631, pl. 31, f. 10.
5 2, 6 2, Japan, including the type. 1 2, 2 2, W. China.
LYMANTRIA DISSOLUTA, nov.
¢, @. Antenne, palpi, frons, and head brown; thorax and
fore-wings dark brownish-grey ; transverse sinuous lines brown,
first antemedial, angled outwards above the middle, second discal
slightly recurved, dentated in parts, third submarginal ; a large
angled dark brown mark on the discoidal vein; cilia grey with
blackish-brown patches ; hind-wings and both wings on the under-
side of a uniform paler grey colour, without markings; on the
upper-side the wing is slightly darker towards the outer border in
some specimens ; abdomen pale dull crimson.
Expanse of wings, ¢ 14, 2 2 inches.
1 gf, 2 9, Hong Kong (Walker).
Belongs to the obsoleta group.
LYMANTRIA TURNERI, noy. nom,
Lymantria aurora, Walk. (?), Turner, Trans. Roy. Soe. S.
Australia, 1902, p. 181.
1 gf, Cape York.
Old World Lymantriide in the National Collection. 485
Butler in Ann. Mag. N. H. (4), 'xx, p. 403 (1877)
described a Lymantria aurora from Japan; the name
aurora for this form cannot therefore stand. The history of
this name is rather curious: on this insect, which was
purchased from Mrs. Higgins in 1867, is a Jabel with the
following, “Lymantria aurora, W.,n.s. named on a spec. of
Mr. Chapman.” Some one must have taken a specimen to
Australia which had been named from Mrs. Higgins’
Cape York example, and ever since this insect has been
called Lymantria aurora, Walker, in Australian collections;
but where the writer of the label containing the name
aurora, Walker, got it from it is impossible to say; cer-
tainly Walker never published any description of this form.
Turner has well described it; there are several examples
in coll. Druce from Cooktown. It is, however, only a form
of antennata, Walker, differing merely in colour.
LYMANTRIA ANTENNATA.
Lymantria antennata, Walker, iv, 881 (1855).
1 g, Richmond River (type). 2 ¢, Moreton Bay.
LYMANTRIA OBSOLETA.
Lymantria obsoleta, Walker, iv, 880, f (1855).
Lymantria bhascara, Moore, Cat. Lep. E. I. C., i, p. 345,
OGsa9)):
Lymantria vinacea, Moore, P. Z. 8., 1879, p. 408, §.
2° Hone Kone. 1 22) Bormosa. fg, N. india
(type). 2 9, Silhet. 19, N. India (type bhascara). 2 g,
Calcutta. I 3,1 9, Bombay. 19, Poona. 1 92, Madras.
29, Trevandrum. 1 JZ, Ceylon.
This is a South Indian form, and I doubt the localities of
Silhet and N. India on the old examples; I have never got
it from any northern locality; I cannot find the type ?
vinacea from Canara.
LYMANTRIA ALBOLUNATA.
Lymantria albolunata, Moore, P. Z.8., 1879, p. 405.
1 gf, 1 2, Dharmsala (type 3). 1 2, Simla (type).. 44,1 9,
Dalhousie. 3 2, Subathu.
The northern form of obsoleta, but easily separable :
there are two diminutive examples from Omeishan and
Moupin in the National Collection, which come very near
to this form.
486 Colonel C. Swinhoe’s Levision of the
LYMANTRIA SOBRINA.
Lymantria sobrina, Moore, P. Z. 8., 1879, p. 402.
1 f, Dharmsala (type). 1 9, N. India (type).
Quite distinct I think from obsoleta, where it is put in
the B. M. collection; I do not consider it to be near
enough allied to that species, even to put it in the form
of what is called a sub-species.
LYMANTRIA SEMICINCTA.
Alope semicincta, Walker, 111, 620 (1855).
Nagunda semicincta, Butler, Hl. Het., v, p. 54, pl. 90, f. 7
(1881).
1 9, N. India (type). 3 9, Darjiling.
I do not know the male of this form.
LYMANTRIA MONACHA.
Bombyx monacha, Linn., Syst. Nat., x, p. 501 (1758).
Af, 5 9, Japan.
Many examples from European localities.
LYMANTRIA CONCOLOR.
Lymantria concolor, Walker, iv, 876, 2 (1855).
F - Butler, Ill. Het., v, p. 55, pl. 90, ff. 8, 9
(1881).
Lymantria superans, Walker, iv, 876, 9.
Lymantria micans, Felder, Reise Nov., pl. 99, f. 2, 2
(1868).
Lymantria carnecolor, Moore, P. Z. 8., 1888, p. 399.
ig cimla, 1a Dalhousie, 1925272) Dharmsalas 3.7
3 2, Kangra, including type f carnecolor. 1 3,1 9, Sultan-
pore: 1 ):9) «Abbottabad. 1) 23. Manipur. ae
Sikhim, including both of Walker’s types. 1 ¢, Kujiar.
1 f, Jawai Hills.
This form is common in the Khasia Hills, from whence I
have received many examples of both sexes; Felder’s type
from Silhet is in coll. Rothschild.
LYMANTRIA TODARA.
Lymantria todara, Moore, P. Z. 8., 1879, p. 402, pl. 33,
f (6.
2 $,2 9, Nilgiri Hills, including the types ¢ 9. 1 2,
Shevaroy Hills. 1 9, Travancore. 1 9, Malabar.
Old World Lymantriide in the National Collection. 487
LYMANTRIA SIMILIS.
Lymantria similis, Moore, P. Z.8., 1879, p. 402, f.
Lymantria cara, Butler, Ill. Het., v, p. 56, pl. 90, f. 13, 2
(1881).
1 2, Calcutta (type). 1 2, Bengal. 1 9, Bhotan (type
CUT).
I think it may be fairly assumed that these are conspe-
cific, though the examples do not come from the same
locality; the differences in the sexes are similar to those
usually occurring in the concolor group, and the double
band on the hind-wings is similar in both sexes.
LYMANTRIA BEATRIX.
Bombyx beatriz, Stoll, Suppl. Cram., pl. 40, f. 2 1791).
1 9, Java.
Stoll’s figure is excellent; I have three females from
Java in my own collection, all identical with the B. M.
example and with Stoll’s figure, and have seen many
others ; it appears to me to be very distinct from the Indian
forms put under it in the B. M.
LYMANTRIA GANAHA, nov.
2. Fore-wings, upper-side of a uniform purplish-grey colour,
slightly tinged with pink, and with a glazed appearance ; markings
and hind-wings the same as beatria ; in beatrix the fore-wings are
pure white.
Expanse of wings 2,°,-—3 inches.
49, Java. 1 9, Kuching, Borneo.
A form of beatriz, Ihave 3 2 in my own collection from
Java and 3 7, and there are three Javan males in the
B. M.; but without breeding I do not think it is possible
to say whether they belong to this form or to beatriz, but
they are quite different from the males of various Indian
forms sunk under beatriz.
LYMANTRIA MARGINATA.
Lymantria marginata, Walker, iv, 877 (1855).
Lymantria pusilla, Felder, Reise Nov., pl. 99, f.3 (1868).
Lymantria nigra, Moore, P. Z. S., 1888, p. 399.
1 ¢, Bengal. 1 9, Silhet (type). 2 9, Sikhim. 1 9,
Burma. 3 2,3 9, Kangra (including Moore’s types). 2 2
Masuri. 2 Z, 2 9, Sikhim.
y
s
488 Colonel C. Swinhoe’s Revision of the
The males vary much in size, but are quite different
from the Javan males; they vary also much in shade of
colour, but none of them have any tinge of yellow; the
females vary in the width and intensity of the medial band
of the fore-wings ; their hind-wings are always pure white,
with a black border ; I have this form in my own collection
from Dharmsala, Sikhim, and the Khasia Hills.
LYMANTRIA FULIGINOSA.
Lymantria fuliginosa, Moore, P. Z. 8., 1883, p. 17.
5 f, 2 2, Bombay, including the types. 1 9, Ceylon.
I have it also from Bombay where I have bred this
form; there is a general yellow tinge in the male; it is
nearer to the Javan form than any of the other Indian
forms; the hind-wings are always more or less yellow ; the
hind-wings of the female are always entirely suffused with
black; I have never seen an exception. The type of
Moore’s wmbrina I have not seen, it is also included by
Hampson under beatriz without any mention of the differ-
ences between the forms, just as if they ran into each
other, but this they do not do.
There are 1 2,3 2 from Moupin, and 1 2 from Chang
Yang, and 1 ¢, 2 from Penang which appear to repre-
sent two more forms, and I have no doubt that there are
many otber forms of this species, or group, still unknown.
To call them all beatrix is very misleading.
LYMANTRIA VELUTINA.
Orgyia (Dasychira) velutina, Mab., Bull. Soc. Zool. Fr., iu,
p. 90 (1878).
1 gf, Madagascar.
LYMANTRIA AURORA.
Lymantria aurora, Butler, Ann. Mag. N. H. (4), xx, p.
403 (1877).
* - Butler, Il]. Het., ui, pl, 24, f. 5 (1878).
Lymantria aurora, var. fusca, Leech, P. Z. 8., 1888, p.
629.
1 gf 1 §, Japan (types). 13,19, Yesso. 4 3, Naga-
hama, including type fusca.
Sunk to mathura inthe B. M. collection, but it is a
distinct form, the male being uniformly blackish-brown.
Butler's male type example is an old worn and faded
Old World Lymantriide in the National Collection. 489
specimen ; whether his female type is really the female of
this form is I think very doubtful.
LYMANTRIA MATHURA.
Lymantria mathura, Moore, P.Z.8., 1865, p. 806.
1 ¢, N. India (type). I ¢, 1 2, Kangra. I 9, Dehra
Doon. 2 2)1 9) sikh 02) NH. Eimalayas: 93°) 02
Loo Choo Islands. 1 2, Chefoo, China. 2 9, Omeishan.
2 f, 3 Q, Japan.
LYMANTRIA VIOLA.
Lymantria viola, Swinhoe, P. Z. 8., 1889, p. 406.
1g, 9 2, Thanna, Bombay, including the types. 2 @,
Jubbulpore.
LYMANTRIA GRANDIS.
Lymantria grandis, Walker, iv, 874, § (1855).
* as Butler, Ill. Het., v, p. 57, pl. 91, ff. 1, 2
(1881).
Lymantria maculosa, Walker, iv, 881, ¢ (1855).
Lymantria metarhoda, Walker, xxxui, 365, f (1865).
1 ¢, 2 2, Ceylon, including the type. 1 ¢ marked
N.S. W.
The type of metarhoda from Ceylon is in Mus. Oxon.
LYMANTRIA SUBROSEA, nom. nov.
Lymantria rosea, Ampsn., Ill. Het., ix, p. 76, pl. 158, f. 28
(1893).
1 f, Ceylon (type).
Butler’s Lymantria rosea from Madagascar (1879) ante-
dates this.
LYMANTRIA LUCESCENS.
Porthetria lucescens, Butler, Trans. Ent. Soc., 1881, p. 11.
1 f, Hakodate, Japan (type). 1 ¢, Tokio. 2 2, Oiwake.
LYMANTRIA MANICATA.
Lymantria manicata, Aurivill., Ent. Tidskr., xiii, p. 193, 2
(1892).
2 9, Old Calabar. 1 9,Sapele, River Niger.
The type from Cameroons is in Mus. Stockholm.
490 Colonel C. Swinhoe’s Levision of the
LYMANTRIA XYLINA, nov.
¢. Antenne black ; palpi brown, white on the inner sides ; head,
body, and wings creamy white ; some crimson hairsat the base of the
antenn ; collar crimson ; fore-wings with a blackish-brown sub-
costal dot near the base, a small spot on the middle of the costa, a red-
brown, rather pale, sinuous, erect transverse thin band beyond the
middle ; hind-wings with some brown suffusion on the costal space ;
both wings with brown marginal dots between the veins; under-side
dirty white, with indications of a transverse band on both wings, ex-
tending to lower end of cell on hind-wings, and some brownish
suffusion in and near cell of fore-wings ; face, pectus, body, and legs
crimson.
Expanse of wings 2,%; inches.
Hab. FORMOSA.
LYMANTRIA GALINARA, Nov.
¢. Antenne ochreous-grey ; palpi black, white inside ; body and
wings white ; thorax with three black spots ; abdomen with blackish
segmental bands ; fore-wings with five transverse, upright, more or
less sinuous bands, similar to those in the fore-wings of J. lepcha,
Moore, from India ; hind-wings with a grey submarginal band and
some slight pinkish suffusion near abdominal margin ; under-side
white, without markings ; face black ; body and legs white.
2. Antenne ochreous-grey; palpi brown; head, body, and wings
dull white ; fore-wings rufous grey, irrorated with white atoms ;
indications of central, discal, and submarginal rufous grey transverse
bands; a ringed lunular mark at the end of the cell, and sub-
marginal whitish lunules ; hind-wings and under-side of both wings,
body, and legs dull white without markings.
Expanse of wings ¢ 2:45, 2 3;°5 inches.
1 2,19, Singapore (fzdley) (types). 1 2, Bali, low country
(Doherty). 1 %, Tambak, Borneo.
I have both sexes from Java, and a very large female
from New Mecklenburgh measuring 5,5, inches in expanse.
Semper records it from the Philippines under the name of
Lymantria lepcha, Moore ; and the female at p. 460 of Het.
Philippines he doubtfully identifies as the female of
Dasychira grossa, Pagenstecher.
LYMANTRIA LEPCHA.
Porthetria lepcha, Moore, Lep. Atk., p. 54, f (1879).
Barhona carneola, Moore, |. ¢., p. 56, 9.
3 2, 6 9, Sikhim, including both types.
Old World Lymentriide in the National Collection. 491
LYMANTRIA BRUNNEIPLAGA, nov.
¢. Palpi black with white thick hairs at the tips ; antenne# brown,
shafts white at the base ; head, thorax, and fore-wings white ; costa
with four brown spots, the first and second large, third small, fourth
composed of two streaks ; a large brown patch on the middle of the
hinder margin; some brown dots near the base: two dentated
transverse brown lines across the middle; another submarginal ;
some rather large brown spots on the outer margin ; abdomen pink,
with a dorsal row of brown spots ; hind-wings grey ; a slightly darker
marginal border and white cilia; under-side, pectus brown ; legs
white with black spots ; fore femora crimson and some crimson hairs
on the white thorax.
Q. With wings shaped as in the female of lepcha; head and thorax
chocolate brown ; fore-wings chocolate grey, the central and sub-
marginal transverse dentated lines of the fore-wings as in the male,
also the large brown patch on the middle of the hinder margin and
the brown spots on the outer margin, which however are on both
wings ; the hind-wings of a pale grey colour.
Expanse of wings ¢ 1,5, 2 2,5, inches.
1 $1 9, Java (types). 2 fg, Borneo. 1 J, Sumatra.
1 9, Penang.
Unfortunately in describing cara, Butler queried these
Bornean males as probably the males of his species, and
consequently these examples have remained over the cara
label ever since, and so stand in many collections ; but
they have no connexion with cara, which Hampson was
perfectly justified in putting as the female of similis.
LYMANTRIA LUNATA.
Bombyx lunata, Stoll, Pap. Exot., iv, pl. 369, f. C (1782).
Pegella wchorina, Butler, Ann. Mag. N. H. (5), xiii, p. 201
(1884),
2 2, 2 9, Amboina, including Butler's type. 1 2, Celebes.
1 9, Aru.
I have it also from Celebes, Amboina, New Guinea, and
Kast Java.
LYMANTRIA CURVIFERA.
Pegella curvifera, Walker, xxxv, 1922 (1866).
Lymantria curvifera, Swinhoe, Cat. Het. Mus. Oxon., i, p.
223 (note) (1892).
Lymantria lunata, Semper, Het. Philipp., p. 461, pl. N, ff.
5, 7 (larva and pupa) (1898).
492 Colonel C. Swinhoe’s Revision of the
1 2, 1 9, Philippines, including Walker’s type 2, which
Walker described as a male.
I have it from Gilolo and Batjan under Staudinger’s
MS. name batjana. There are apparently two forms of
lunata on all the different islands: the large typical male
with well-marked wings, and the small form lke the
Philippine male of curvifera, the bands in which are
straighter and all the markings ill-defined; the breadth of
the middle transverse bands on the fore- wings of the
female is very variable in both forms ; sometimes it is
a mere line, often broad and commences extended inwards
on the hinder margin.
LYMANTRIA PLUMBALIS.
Lymantria plumbalis, Hmpsn., Trans. Ent. Soc., 1895, p.
292.
ft, Tinlin- Yaw, Burma (type),
LYMANTRIA ASCTRIA.
Lymantria asetria, Hiibner, Samml. Exot. Schmett., u, ff.
1-4 (1818).
Dasychira antica, Walker, vii, 1739, 2 (1856).
Lymantria pramesta, Moore, Cat. Lep. EK. I. C., 1, p. 344,
pl. 9a, £. 3 (1859).
1%, Maulmein. 19,Thyetmyo. 62, 7 9, Java, includ-
ing the types of antica and pramesta.
LYMANTRIA NARINDRA.
Lymantria narindra, Moore, Cat. Lep. E. I. C., ii, p. 342
(1859).
Kirby, Cat. Moths, 1, p. 479 (1892).
Lymantri ia hilari is, Voll, Tijd. v. Hnt., vi, p. 143, pl. 10;
ff. 2, 3 (1863).
1,1 2, Java, including the female type.
I have in my own collection a female from Java and a
male from Sumatra (where Vollenhoven’s types ¢ 9 came
from); the two names undoubtedly belong to the same
species.
LYMANTRIA GRISEA.
Lymantria grisea, Moore, Lep. Atk., p. 55, pl. 3, f 5
(1879).
Old World Lymantrvide in the Naticnal Collecticn, 498
1 gf, N.E. India. 1 9, Darjiling (type). 1 3, N. Chin
Hills, Burma.
LYMANTRIA SILCA.
Lymantria silea, Swin hoe, Ann. Mag. N. H. (7), xii, p. 197
(1903).
1 f, 1 2, Fergusson Isl. (types). 12%, Fergusson Isl. 1 2,
Milne Bay. 1 2,19, Kapaur, New Guinea.
LYMANTRIA GANARA.
Lymantria ganara, Moore, Cat. Lep. E. I. C., ii, p. 344
(1859).
4 ft, 2 9, Java, including the type 2 1 2, Sumatra. 5%,
Borneo. 1 f¢, Singapore. 2 ¢, Burma.
LYMANTRIA BIVITTATA.
Pegella bivittata, Moore, Lep. Atk., p. 57 (1879).
1 2, Darjiling (type). 1 9, Silhet.
I do not know the male of this form.
LYMANTRIA VACILLANS.
Lymantria vacillans, Walker, iv, 873 (1855).
1 2, Congo (type).
LYMANTRIA DULCINEA.
Lymantria dulcinea, Butler, Cist. Ent., iii, p. 12 (1882).
1 Z, 2 $, Madagascar (types).
LYMANTRIA ROSEA.
Lymantria rosea, Butler, Ann. Mag. N. H. (5), iv, p. 239
(1879).
1 Z, 1 9, Madagascar (types).
LYMANTRIA LUGARDI, nov.
2, d. Palpi brown, under-side yellow ; antenne black ; frons
brown ; thorax chestnut-brown; abdomen crimson ; fore-wings
purple brown, in the female almost purplish slate colour; some
dark brown, irregular and wavy, transverse bands; one at the
base ; a very broad one in the middle, with very irregular sides,
and containing a large nearly white irregular square with its
upper edge against the costa, and in the middle of it a black
494 Colonel C. Swinhoe’s Levision of the
spot at the end of the cell ; a pale sinuous discal line, and some pale
yellowish elongated spots in the disc ; all these markings more or
less obscure and contracted in the male, prominent in the female ;
cilia yellow with brown spots; hind-wings ochreous without
markings,
Expanse of wings ¢ 1
2 $, 3 9, N’Gami Country (Lugard).
>
10 ? liv inches.
LYMANTRIA BANANA.
Lymantria bananx, Butler, P. Z.8., 1896, p. 848, pl. 42,
f. 9.
1 Z, Nyassaland (type).
LYMANTRIA TOTTEA, nov.
gd. Palpi with the lower and under parts white, tips and last
joints black, and a tuft of black hairs immediately below ; antennwe
grey ; frons brown, with the edges greyish-ochreous ; head and
thorax greyish-ochreous ; fore-wings pale brownish-grey with a tinge
of yellow ; a broad whitish band divided by the veins into eight
square-shaped large spots, and a large square spot at the end of the
cell, divided from the band by a thin band crossing the discoidal
vein ; a small round spot in the middle of the disc outside the band ;
abdomen pale greyish-yellow ; hind-wings white tinged with yellow ;
under-side white without markings ; pectus and fore-legs ochreous ;
legs white with black tips.
Expanse of wings 1,5; inches.
Hab. OLD CALABAR.
Allied to banane with which I found it, but very
different, except superficially.
LYMANTRIA MOSERA.
Lymantria mosera, Druce, Ann. Mag. N. H. (7), i, p. 208
(1898).
1g, 1 9, Delagoa Bay. 22,19, Natal. 1 ¢, Rhodesia.
LYMANTRIA EDDELA, nov.
¢. Antenne black, shafts white; frons, head, thorax, and fore-
wings pale pinkish buff; a black spot at base of fore-wings, and three
other spots close toit ; antemedial and discal transverse lines of black
spots, erect but slightly sinuous ; three black spots at the end of the
cell ; abdomen orange with a dorsal row of black spots ; hind-wings
paler than the fore-wings, nearly white; a broad marginal black
Old World Lymantriide in the National Collection. 495
band ; under-side of body and legs blackish ; both wings of the
colour of the fore-wings above ; fore-wings with an apical and outer
marginal grey band, which narrows hindwards ; hind-wings with a
broad black band as above.
Expanse of wings 1,5, inches.
Hab. Fwameo, Tanganyika,
LYMANTRIA CARRIALA, nov.
?. Frons, antenne, thorax, and fore-wings pale blackish-brown ;
two white spots on the thorax ; fore-wings with a large white patch
on the middle of the hinder margin, with a white spot immediately
above it, and with four small black spots inside it almost in the form
of a square ; two or three indistinct pale spots in the upper disc ;
hind-wings pale pink ; a rather broad apical pale blackish-brown
band, which stops abruptly half-way down the outer margin ;
abdomen yellow, with dorsal and lateral black spots ; under-side,
wings as above, very thinly clothed ; body and legs blackish.
Expanse of wings 1,5 inches.
Hab. ANGOLA.
Belongs to the group of LZ. mosera, Druce, from Natal.
LYMAXTRIA GONDONA.
?. Thorax and fore-wings pale grey, nearly white, the latter with
four darker grey transverse bands, limited by brown lines; the first
basal, the second before the middle and bread, the limiting lines
rather sinuous, the third discal, a little narrower than the second,
bent outwards above the middle, the limiting lines semi-dentate,
the fourth apical ; the inner margin very sinuous ; hind-wings white
without markings; cilia with pale brown spots ; abdomen pink.
Expanse of wings 1} inches.
Hab. SABAKI VALLEY, Brit. E. Africa.
Genus Dura, Moore, Lep. Atk., p. 56 (1879).
DURA ALBA.
Dura alba, Moore, |. e.
1 9, Darjiling (type). 2 f without locality.
The type ¢ from Dazjiling is in coll. Staudinger.
DURA ALBICANS.
Dasychira albicans, Walker, vii, 1739 (1856).
1 2, Celebes (type). 4%, Borneo. 1 2, Singapore.
TRANS. ENT. SOC. LOND. 1903.—PART TI. (OCT.) 33
496 Colonel C. Swinhoe’s Revision of the
Genus Imavs, Moore, Lep. Atk., p. 54 (1879),
IMAUS MUNDUS.
Lymaniria munda, Walker, iv, 875 (1855).
1 4 49; Sikhim. 1 9, Silhet (type). 29, Java.
Genus: PyrAMOocERA, Butler, Journ. Linn. Soe. Lond., xv,
p. 85 (1880).
PYRAMOCERA FULIGINEA.
Pyramocera fuliginea, Butler, |. ¢. (fig.).
1 f 1 Y, Madagascar (types).
Genus PotymMona, Walker, iii, 768, 2.
Morasa, Walker, iv, 859 (1855).
Sarothropyga, Felder, Reise Nov., pl. 100, f. 23, 3
(1868) (undeser.).
POLYMONA RUFIFEMUR.
Polymona rujifemur, Walker, ui, 768, &.
Morasa lorimeri, Butler, P. Z.S., 1878, p. 387, &.
1 & Africa (type). 2 2, Natal, including Butler's type. |
1g, Rhodesia. 1 3, Nyassaland. 1 2, Mashonaland.
POLYMONA MODESTA.
Morasa modesta, Walker, iv, 859 (1855).
Sarothropyga rhodopepla, Felder, Reise Nov., pl. 100, f. 23,
2 (1868).
1 9,8. Africa (type). 5%, 8 2, Natal.
Genus NatTaxa, Walker, v, 1179 (1855). |
Dicreagra, Felder, Reise Nov., pl. 100, f. 2 (1868)
(undeser.).
NATAXA FLAVESCENS.
Perna (2) jflavescens, Walker, v, 1128.
Natazxa flavijascia, Walker, v, 1179.
Diereagra ochrocephala, Felder, 1. ¢.
1 Z, Australia (type). 1 2, Tasmania (type flavifascia).
17,19, Melbourne. 5 9,S.E. Australia.
Old World Lymantriide in the National Collection. 497
NATAXA RUBIDA.
Nataxa rubida, Walker, xxxui, 512 (1865 ).
1 f, Australia (type).
It differs from flavescens in having no central band, only a
yellow spot on the costa where the band commences in the
other form.
The following genera have been removed from the
Lymantriide in the National collection, but I have been
unable to find them. The numbers are those of the genera
in Kirby :-—
44. Comana, Walker, xxx, 495.
90. Xanthodura, Butler, Ann. Mag. N. H. (5), v, p. 384.
103. GEnosandra, Newman, Trans. Ent. Soc., 1856, p. 286.
112. Murcipa, Walker, iv, 807.
156. Epipyrops, Westw., Proc. and Trans. Ent. Soc., 1876, pp. xxiv,
522,
163. Sarsina, Walker, iv, 800.
Genera in Kirby’s Catalogue wrongly included in the
Lynantrude :—
1. Geodena, Walker, vii, 1691 . Belongs to the Aganaide.
2. Soloé, Walker, 1, 557 é : Pe 1 é
3. Isine, Walker, 11, 545 = Chione-
ma, Herr.-Schiff., Ausser-
Eur. Schmett., p. 20 . d - PF Lithosiide.,
13. Lerna, Walker, xxxiil, 805=
Castulo, Walker, i, 562. 5 ss Fi
27. Cypra, Boisd., Voy. de l’Astro-
labe, p. 201 . ‘ : ‘ s » Boarmiide.
28. Deroca, Walker, iv, 822 . : Fe » Drepanulide.
41. Trichetra, Westw., Mod. Class.
Ins. Gen. Syn., p. 92 . ; s » Lupterotide.
42. Marane, Walker, xxxii, 397. iS 7 1
43. Apina, Walker, ili, 756 . : % 35 Noctwide.
51. Phiala, Wallengr., Wien. Ent.
Mon., iv, p. 165 . : : D » Lupterotide.
72. Raphipeza, Butler, Ann. Mag.
N.H. (5), v, p. 386. : 5s . Lasiocampide.
73. Chrysopsyche, Butler, 1. ¢. p. 387. 5 a +
Colonel C. Swinhoe’s Revision, ete.
. Trisula, Moore, Cat. . Lep.
Hol. Ci, ai, p. 420
. Trisuloides, Butler, Ann. Mag.
N. H. (5), vii, p. 36
. Rhanidophora, Wallengr., CEf-
vers. Vet. Akad. Forh., xv,
p. 213 .
. Amsacta, Walker, iv, 804
. Teara, Walker, iv, 846= Agla-
osoma, Scott, Austral. Lep.,
p. 14
. Ochrogaster, Felder, Reise Nov.,
pl. 94, f. 5
}. Sitina, Walker, iv, 854
. Callia, Walker, vi, 1482 .
. Amana, Walker, ii, 661 .
. Vanga, Walker, xxxii, 453=
Smyriodes, Guen., Phal., i,
p- 223
° Beralade, Walker, lv, 852
. Cebysa, Walker, 11, 486
. Castulo, Walker, 11, 561
. Cluaca, Walker, xxxi, 268
. Lecriolepis, Butler, Ann. Mag.
IN; @EE(5), Vv; Pp. Beco
. Laganda, Walker, xxxii, 389
. Ticilia, Walker, xxxii, 394
. Cosmethis, Hiibner, Verz.
Schmett, p. 179= Arycanda
Walker, vii, 1775
Belongs to the Noctwide.
” ”
” a0)
5 Arctiide.
» LHupterotide.
” ”
», Lasiocampide.
” aed)
* Uraniide.
55 Boarmiide.
,», Lasiocampide.
Tineide.
ss Lithosiide.
oe) ”
», Lasiocampide.
s Bombycide.
» Drepaniulide.
» Boarmiide.
OcToBER 5TH, 1903.
|
|
|
|
|
a a a
( 499 )
XX. On the antenne of Hepialidee—Lepidoptera Jugate.
By AMBROSE QUAIL, F.E.S.
[Read October 7th, 1903. ]
jy oe. © DD.
THE characters which separate Jugatx from other Lepido-
ptera in the imago stage are well known to be definite and
constant, being essentially the structure of the wings,
which have nervures more numerous than prevails with
other Lepidoptera, and a “jugum” or lobe near the
base of the fore-wings which may be noted among Neuro-
ptera, Diptera, and Hymenoptera, but not elsewhere among
Lepidoptera. The Hepialid group have neither maxillary
palpi nor tibial spurs, which characters are well developed
among other Jugate.
The species which comprise the well-defined Hepialide
have great diversity of antennal structure, which I have
examined, so far as my material permits, to observe whether
inter se there is evidence indicating which is the older and
more primitive form; I believe the subject will interest
others. To properly examine the antennal structure one
must remove and mount in balsam, or other media; for this
reason one is confined to insects in one’s own collection,
and in consequence many of my otherwise perfect specimens
are minus antenne.
In my previous yaper I did not note the presence of
scales on the Hepialid antenna, having devoted attention
to the pectination per se; the scaling, however, is very
obvious. The large proximal joint (scape) is invariably
scaled more or less completely, the scales are long—often
overlap one or more further segments—striated, slender,
attenuated throughout when viewed sideways, and look
very like a hair; in fact, as Bodine puts it, “the scale is
a hair flattened out.” The segment next the scape
(pedicel) is smaller and sometimes clothed with scales
similar to those of the scape, but usually with shorter and
broader scales approximating to the scales of the wings,
and this kind of scale prevails on the other (clavola)
segments of the antenna.
TRANS, ENT. SOO, LOND. 1903.—PART IV. (DEC.) 34
500 Mr. A. Quail on
On the surface of the segments there are minute “hairs,”
which may be sparsely scattered as on the scape, or closely
cover the surface of the segment as on the clavola; those
of the scape are perhaps more bristle-like, and remind one
of the minute hairs on the Hepialid wing membrane. One
cannot help thinking that these are the original antennal
“hairs,” antecedent to all other hairs and scales on the
Lepidopterous antenna; those which we find on the clavola
segments everywhere, scaled or not, are, I take it, these
‘primary ” hairs, perhaps somewhat modified ; they serve
probably the function of protection.
Amongst the “primary” hairs others arise which are
longer and tapering, not rigid, usually fine but sometimes
stout with socket rims at base; these are probably “sense ”
hairs, and appear to be in some way associated with reduced
scaling. The sensory hairs appear to be confined to the
clavola segments. One has, however, some diffidence in
deciding whether some forms of “sense” hairs are not
bristles. There are short, pointed, socket-rimmed bristles
at the base of both scape and pedicel, and scattered among
the scales are longer bristles; there are always bristles on
the clavola segments. The transition from pale-coloured,
fine scape bristles to red-tinted, robust, rigid clavolar
bristles, I have demonstrated on the antenna of Porina
fuscomaculata. The bristles on the antenna of one species
may resemble the “sense ” hairs on the antenna of another
species, but all the “sense” hairs on any given antenna
differ from the bristles of that antenna.
There appear to be good reasons for believing the
primitive form of Lepidopterous antenna to have been
one closely covered with scales—such forms one finds
among the lower groups of Lepidoptera—the scales after-
wards replaced by “‘sense” hairs, but amongst the scales
there were semi-erect bristles, and, it may be remarked,
similar bristles, whatever their functions, occur on different
parts of the “scale-covered” wings. The bristle is probably
older therefore than “sense ” hairs.
The scape and pedicel retain comparative rotundity with
remarkable constancy throughout the Hepialidz known to
me, with the exception of Oncoptera (intricata), in which
the scape is remarkably elongate. The scape and pedicel
are never pectinate in the Hepialide.
The segments of the clavola vary greatly in number,
form, and dimensions in Hepialide, The proximal joint
the antenne of Hepialide—Lepidoptera Jugatz. 501
is usually smaller than succeeding segments, being as it
were a transition between the pedicel and clavola, each with
its definite functions; this joint has usually only rudi-
mentary appendages on antenne with pectinations; rarely
is true pectination developed, and never does it attain
proportional length. The proximal joint of the clavola,
at least, lags behind the others in the onward march of
specialization.
The sexes on the whole agree as to number of clavola
segments per species, for instance Hepialus humula, 8, § ,
and var. Hethlandica, have an equal number; sometimes a
variation in number occurs, but such has been noticed
mostly in the female sex. It is an interesting fact that
sometimes segments are partially or wholly coalesced, the
sutures being obliterated, but hairs, bristles, etc., are
normal. I have noted this in Charagia virescens, C. daph-
nandre, Hepialus argenteo-maculata, Cibyra sylvinus. The
division of the clavola into more or less segments does not
seem of first importance, their functions being collective.
As far as possible several specimens of each of the following
species have been examined, to arrive at the probable
typical number of clavola segments.
HEPIALUS = unpectinate antennee.
humauli. 20 clavola segments, a few scales on five
basal segments.
velleda. 23 clavola segments, a few scales on first
basal segment.
lupulinus. 24 clavola segments, one or two scales
on first basal segment, also single
scales per segment on apical thirteen.
hectus 3. 12 clavola segments, one or two scales
on first and second basal segments,
Subg. STHENOPIS = unpectinate antenne.
argenteo-maculata. 21 clavola segments, a few scales on six
basal segments.
Subg. CHARAGIA = unpectinate antenne.
daphnandre. 28 clavola segments, numerous scales on
four basal segments and dorsal scales
on next fivesegments ¢ ; dorsal scales
on nine basal segments 9.
virescens. 39 clavola segments, numerous dorsal
scales on six basal segments ¢ ; few
dorsal scales on six basal 9.
502 Mr. A. Quail on
eximic, 28 clavola segments, dorsal scales on
four basal segments.
ramsayi. 35 Pa A ar A
lignivora. 24 clavola segments, few dorsal scales
on four basal segments.
Snbg. CipyrA = — Jatero-ventral enlargement of antennal segments,
sylvinus. 23 clavola segments, dorsal scales from
base to tip of antenna.
ONCOPTERA = unpectinate antenna.
intricata. 17 clavola segments, numerons scales
on nine basal segments, with dorsal
scales on others to apical segment.
( ?) = ventral enlargement of antennal segments.
bacotii 50 clavola segments, devoid of scales.
GORGOPIS = _ bipectinate antenne.
libania. 41 clavola segments, devoid of scales.
HECTOMANES= bipectinate antenne.
Susca. 30 clavola segments, dorsal scales from
base to tip of antenna,
simulans. 34 = - - es
erocet. 34 clavola segments, dorsal scales on
basal thirteen segments.
PORINA = semi-tripectinate antenne.
cervinata, 36 clavola segments, devoid of scales.
despecta. B10 on -s
umbraculata, 38 clavola segments, one or two dorsal
scales on three basal segments.
signata.
enysit ¢.
ee
40 clavola segments, devoid of scales.
49 clavola segments, one or two dorsal
scales on four basal segments.
45 clavola segments, a few dorsal scales
on six basal segments.
fuscomaculata ¢. 56 clavola segments, one or two dorsal
scales on first basal segment.
TRICTENA = tripectinate antennze 7, unipectinate 9.
labyrinthica. 71 clavola segments, dorsal scales on
four basal segments,
PIELUS = ventral enlargement of @ antennal
segments.
hyalinatus. 54 clavola segments, few dorsal scales
on seven basal segments.
Evriocrania admitted on other grounds—embryological
the antenne of Hepialide—Lepidoptera Jugatx. 503
and imaginal—to be more ancient than Hepialide, and
accepted as such by Bodine and Chapman in respect to
antennal structure, has (subpwrpurella) elongated scape
and fully-sealed clavola—forty- one segments in number—
the scales bidentate, alignment not definite, the outer row
of scales overlap base of next segment ; the segments are
circular in transverse section, and are clothed with a erowth
of primary hairs, the apical segment an elongated cone
with scales, “ primary ” hairs and numerous bristles. An
associated genus Mesarchea (hamadelpha) furnishes a form
with short scape, fully-scaled clavola, the scales bidentate,
overlapping. Palwomicra (chalcophanes) furnishes a form
with short scape curiously swollen at both ends, the clavola
having on each segment a few remarkable broad leaf-like
scales placed in pairs, one of each almost covering its
fellow; on the segments bristles rise from beneath, between
and beyond the scales. We have the support of our
authorities in selecting the eriocranid form of antenna
as being probably the most generalized.
Examining our material among the Hepialidx, Oncoptera
arrests our attention. Two or three basal segments are
not unlike the eriocranid in shape, circular in transverse
section, apparently fully scaled on all aspects ; the segments
beyond gradually become less eriocranid-like, shorter,
broader, and developing towards the apical segment
a ventral ridge. Completely scaled as are the basal
segments with. typical flat, striated, bidentate scales, align-
ment not definite ely two rows,* the outer scales overlapping
next segment, localization of hairs commences on a ventral
area of the third clavola segment, where there are only
“primary” hairs and one or two bristles, gradually the
scales are replaced on a larger ventral area, “sense” hairs
develop, the scales becoming dorsal only. We find associ-
ated with the scaled clavola of Oncoptera an elongate scape ;
we suspect that with fuller material others would be found
with elongate scape and somewhat fully-scaled clavola ;
meanwhile we look upon these as primitive characters
retained by Oncoptera, which has, however, somewhat
specialized on the other clavola segments.
The clavola segments of Hepialus and Charagia are
circular in transverse section, and the apical segment of
some species, like Hriocrania, isan elongate cone; H.humuli
* IT cannot detect alternate lines of scales and hairs, there appear
to be only scales except for occasional bristles.
504 Mr. A. Quail on
is typical in this respect. The majority of species of
Charagia have only a few dorsal scaleson not more than
half-a-dozen basal segments, the scales of the antenna
like those cf the wing being green in colour, striated, not
dentate, and the segments have numerous pits bearing
superficial resemblance to sockets of long-lost scales. C.
daphnandrex preserves some basal segments scaled dorsally
and laterally, with “primary” and “sense” hairs and
bristles located on a ventral unscaled area; the segments
beyond are devoid of scales, “sense” hairs are not numer-
ous on the dorsum, they are so, however, both laterally
and ventrally, with bristles on all aspects at the distal end
of the segments. C. lignivora has gone a long way
ahead, the scales having almost completely vanished, and
“sense ” hairs are numerous on all aspects.
European Hepialus have likewise gone a long way
towards complete loss of clavola scales. H. /upulinus
retains single scales per segment curiously enough on the
apical half of the clavola, the scales are not dentate and
are longer than ordinary clavolar scales; one finds here
“sense” hairs on the dorsum as on other aspects. The
antennal scales of H. lupulinus suggest that Hepialus,
which now so generally have only a few clavolar scales,
somewhere in the past had antennz scaled from base to
tip. The subgenus Cibyra preserves exactly this condition,
and must have separated from Hepialus when the antenne
were scaled to tip; it is not in the least likely that Cibyra
has since developed scales after Hepialus once lost them.
Cibyra then retains this character, but has developed latero-
ventral enlargement—the beginnings of pectination—there-
by increasing the “sensory” area, which indeed is wholly
covered with “primary” and “sense” hairs, there being
pone of the latter on the scaled dorsum.
The latero-ventral enlargement of Cibyra is not properly
unipectination although in profile it appears to be so, it
is a widening and ventral lengthening of the segment as a
whole. Definite ventral unipectination does exist else-
where among Lepidoptera (Jnewrvaria muscalella), and
may be observed among Diptera (Zipulidz).
Among the latero-ventrally enlarged forms known to me
the clavola segments of bacotii (Gorgopis ?) are most special-
ized; “sense” hairs completely cover the segments on all
aspects together with “primary” hairs, even the bristles
are reduced in size, so that the clothing of the segments 1s
Lepidoptera Tugate. 505
the antennx of Hepialide
remarkably uniform. We have here a form of Hepialid
antenna answering the Jordan requirements of an antenna
covered with hairs and devoid of scales ; this form, however,
from our standpoint 1 is reached by. specialization, and
therefore is not primitive. Indeed it seems abundantly
evident that the tendency in the Hepialide has been to
completely get rid of the clavola scales. This seems at
first sight somewhat at variance with the tendency shown
by the majority of the Lepidoptera, which generally preserve
the dorsal scales; but Hepialidx are a very distinct and
somewhat isolated group, moreover the antenne are
admitted to be “sense” organs, and it seems only reason-
able to conclude that on the antenne and on the wings the
function of scales is identical, and it can hardly be as
aiding any special sense, such as we believe to be centred
in the antenne, for similar sense can hardly be supposed
to exist in the wings. It is not illogical to believe that
when the primitive Lepidopteron developed scales—how-
ever far back we may have to go, the development com-
menced somewhere, and has since extended and become a
permanent and characteristic feature of the Lepidoptera—
the scales spread over the several organs, proving to be an
encumbrance on the antennae, although of some utility to
many groups of Lepidoptera, since not only are scales
retained on the dorsum of the shaft, but they are spread
over the dorsa of pectinations; but the effects of elimination
can be traced everywhere in the localization of the scales
instead of their general distribution over the clavola
segments, which is to be observed among the very primitive
groups, as for instance the eriocranid already discussed.
Complete loss of scales by elimination has been the
ultimate result on the antenne of many Hepialide.
Hectomanes have bipectinate antenne generally scaled
from base to tip on the dorsum of the clavola, the scales
not dentate, with “ primary ” hairs, ventral “sense” hairs,
and numerous ventral pits. The pectinations of Hecto-
manes are essentially lateral extensions, short and curved ;
one can hardly say whether highly specialized or no, judg-
ing from the scales they are not. The bipectinate, unscaled
antenna of Gorgopis (libania) is a very highly-specialized
form. The pectinations are long slender appendages,
provided with ‘ primary” hairs, “sense” hairs on the
inner side, and bristles on the outer side; on the dorsum
of the shaft the ‘primary ” hairs are modified or replaced
506 Mr. A. Quail on
by sharp-pointed spines, of which I do not know the like
elsewhere among Hepialidx. I erred in placing bacotii in
the genus Gorgopis; although both are highly specialized,
as regards the antennee the structure is so different.
The antennz of Porina are interesting. Viewed from
the side clavola segments are apparently uniform, with
sight ventral extension, “sense” hairs encircling the
anterior rim of the segment, and others gathered over a
ventral posterior area; a transverse view shows the seg-
ments are somewhat quadrangular with posterior append-
ages; viewed from above, the lateral angles of the quad-
rangular figure appear now to be the tips of lateral
bipectination, and the shaft of the segment is small and
constricted. The appearance of pectination is greater
beyond the middle of the antenna, where the shaft is
smaller and the lateral elongations are longer, still how-
ever preserving the quadrangular form tr ansversely ; the
apical segment presents a much-specialized structure.
Porina fuscomaculata has an antenna which in the ¢
has complete bipectination, which supports our view of
the specialized form being like Gorgopis; on the ventral
surface there is, however, a “sensory ” tubercle which
brings this definitely bipectinate Porina in line with the
other species of the genus. The bristles appear to be
always placed irrevularly on the different segments of the
clavola in Hepialide, and P. fuscomaculata is a convenient
species to examine in this connection. On the dorsum of
Ist clavola segment are some half-dozen bristles (slender) ;
2nd has several slender and one large bristle on dorsum,
and one large bristle on one of the pectinations ; 3rd, four
slender bristles on dorsum, two large bristles on one
pectination; 4th, two slender, two large bristles, on
dorsum; 5th, one slender bristle middle of dorsum, one
near pectination, three large bristles at intervals on pectin-
ation, and two ditto on opposite pectination ; 6th, two
slender bristles on dorsum, four large on pectination, one
on opposite pectination ; 7th, one slender bristle in front,
one large one at middle of dorsum, three large on one
pectination, one on opposite. In addition to large bristles
placed irregularly on the pectinations, succeeding seg-
ments have on the dorsum of shaft, one large bristle on
Sth; next has two large bristles, then three segments have
one bristle each; next has two bristles; thirteen consecutive
segments have one bristle each; next has two; next one;
Pam
the antennx of Hepialide—Lepidoptera Jugate. 507
next one; next two; four segments have one each; next
two, and so on. Not only is the irregularity in number
and position interesting, but the transition from slender
bristles to stout ones is also.
Nowhere in Porina do we find the ventral “sensory ”
area develop into a definite appendage, but the centring
of “sense” hairs on a ventral area which is a feature of
Porina antennal structure, suggests that such ventral
development might occur, and that we find such in
Trictena is most interesting.
The female antenna of 7'rictena (labyrinthica) is remark-
able in the appendage—distinctly ventral—being very
much larger than the shaft of the segment ; this antenna
presents a very near approach to unipectination in the
remarkable slender appendage. The swollen area above is,
however, the rudiment of lateral pectinations, which we
find highly developed in the male, which has probably the
most specialized Hepialid antennz. Decidedly the male of
Trictena, with its tripectinate antenna, is more highly
specialized than the female. We may then regard a
laterally-pectinated antenna as specialized, and look back
consistently, touching Porina by the way, to the latero-
ventral forms—Pvielus, bacoti, Cibyra, thence to Hepialus
and Oncoptera. We suggest divergence in several direc-
tions from some such form as Oncoptera (but not Oncoptera
itself). It is unlikely there has been convergence from a
number of points represented by several forms of pectinated
antenne, and some others.
We can indeed corroborate to some extent the conclu-
sions arrived at from a study of the antenne, by examination
of the wing neuration. Oncoptera, Hepialus, Charagia,
Gorgopis, bacotii, Trictena, are of one pattern; from a
previous paper I make the following quotation—“ It is
as nearly certain as anything of the sort can be that the
Hepialus-Trictena pattern of neuration is a generalized
pattern, from which the wing patterns of Hectomanes,
Palpiphorus, and of Hepialiscus, and Porina, have been
derived.”
Fig.
<3)
or
10.
Wik,
12.
13.
14.
15,
16,
ie
18.
MS).
20.
21.
(508)
EXPLANATION OF PLATE XIX.
Eriocrania subpurpurella.
”) »
Palxomicra chalcophanes.
Oncoptera intricata.
” 9
Charagia virescens.
% ”
” ”
2 97
» fuscomaculata.
Hectomanes crocea.
Gorgopis libania.
a hn
) bacolit.
)
Pielus hyalinatus.
Trictena labyrinthica.
” ”
Scape, pedicel, first clavola
segments x 200.
Clavola segment x 200
” ” )
Scape, pedicel, two clavola
segments x 100.
Clavola segment x 200
Scape, pedicel, first clavola
segment xX 50.
Base of scape showing bristles
x 200.
Clavola segment lateral view
x 200.
Clavola segment transverse
x 200.
Clavola seginent terminal
x 200.
Clavola segment lateral view
x 200.
Clavola sesyment transverse
x<e2 00;
Clavola segment beneath x
200.
Clavola segment terminal x
200.
Clavola segment x 50.
‘ ia x 200.
. » x50.
Spines on dorsum of shaft x
200.
Clavola segments lateral view
x 200.
Clavola segments transverse
x 200.
Clavola segments lateral view
x 100.
Clavola segments transverse
x 100.
Clavola segments ¢ x 50.
PS (0),
»P) )
Magnification quoted is approximate,
XXI. On the Laparostict Lamellicorn Coleoptera of Grenada
and St. Vineent (W. Indies). By GILBERT J.
Arrow, F.E.S.
[Read October 7th, 1903.]
In 1900 I published a paper in these Transactions dealing
with the Pleurostict Lamellicornia collected by Mr. H. H.
Smith in the islands of St. Vincent and Grenada. The
present paper contains an enumeration of the remainder
of the Lamellicornia of the same collection. The total
number of species of the Laparostict families is 22, making
in all 40 species of Lamellicorns now known froin the
islands. Of the present 22 species, 9 are known as in-
habitants of the mainland of America, 2 more have been
described from other islands of the Antilles, and 10 are
here described as new. One of the latter, <Atenius
tenebrosus, is also a widely-distributed form and, with the
exception of 3 represented by single specimens and 3 by
two specimens, only 3 of the whole number did not occur in
both islands; and two of these three occur elsewhere. The
families dealt with in my previous paper therefore differ
curiously from those now under consideration in the
localization of their species.
The fauna of St. Vincent has assumed a very special
interest since the volcanic catastrophe from which the
island has so recently suffered, and which can hardly have
failed to produce a permanent effect upon its fauna and
flora. It is fortunate that so excellent a collection was
brought from the island before the event, so that future
comparisons will be possible which should throw valuable
light upon biological problems.
I have also characterized in this paper a few Central
American allied forms, as to which confusion has previously
existed.
CopRID&.
Charidium tllesum, Harold.
This insect was found in decaying fruit and other
vegetable débris. It was very common in St. Vincent,
Grenada, and Mustique in the Grenadines. It has only
TRANS. ENT. SOC. LOND. 1903.—PART Iv. (DEC.)
510 Mr. G. J. Arrow on the Laparostict Lamellicorn
previously been recorded from Mexico, but will no doubt
be found in other West Indian islands.
Pseudocanthon chlorizans, Bates.
A specimen was found in Mustique, and another in
Union Island, in the Grenadines. This species also has
only so far been known from Mexico. The green tinge is
exceedingly faint, as in the specimens from Yucatan, and
traceable only on the head and thorax. The elytra are
quite black in all known examples except the type, found
in Juquila.
The angulate inner edge of the front tibia of the male is
the only substantial distinction between this genus and
Canthon, the parallel-sided thorax mentioned by its author
being only an optical effect produced in a particular position
of the specimen.
Uré..ys Vineentie, sp. n.
Parum elongatus, cupreo-niger, nitidus, capite polito, sat crebre
punctato, oculis supra angustis, clypei margine setoso, medio biden-
tato ; prothorace levissime punctato, lateribus valde angulatis, pro-
funde sulcatis, sulco ad basin attingente ; elytris extus carina integra
apice paulo depressa circumdatis, striatis, striis haud perspicue
punctatis ; pygidio polito convexo, es profunde bi-impresso ; tibiis
anticis gracilibus, tridentatis, dentibus omnibus multo pest medium
positis.
Long. 3-4 mm.
Hab. Sv. VINCENT, Cumberland.
Two specimens of this minute species were found beneath
rubbish on a damp rock near the sea. It has the form and
size of U. pygmeus, Harold, but the upper division of the
eyes is narrow, and there are no supplementary teeth on
the margin of the clypeus. The disk of the latter is with-
out any trace of transverse wrinkles. The slight depression
at the suture of the carina surrounding the elytra suggests
the origin of the apical processes which form such a
remarkable feature of some of the larger species of this
genus,
Onthophagus Antillarum, sp. n.
Fusco-niger, opacus, supra parce brevissime setosus, pygidio
femoribusque 4 posterioribus plerumque flavis, elytris basi nonnun-
quam flavo-maculatis ; clypeo integro, leviter punctato-rugoso, maris
Coleoptera of Grenada and St. Vineent (W. Indies). 511
vertice cornnbus 2 intus curvatis, cum carina connectiva cirenlum
fere formantibus, armato ; prothorace minute sat regulariter punctato,
antice utrinque profunde impresso, lobo crasso antico formante,
postice medio leviter canaliculato, margine angulato ; elytris subtiliter
striatis, striis haud punetatis, interstitiis minutissime et disperse
punctulatis, punctis setiferis, elytris nonnunquam minus nigris, basi
vage rufo-flavo-maculatis ; pygidio, abdominis lateribus, femoribusque
4 posterioribus plerumque flavis, pedum reliquis fusco-castaneis, tibiis
anticis acute 4-dentatis.
Long. 65 mm.
Hab. St. VINCENT; GRENADA, Balthazar, Mt. Gay
Estate.
The armature is described from the best-developed male,
in which the horns are about as long as the head and, with
the connecting ridge, form three-quarters of a circle. The
female is similar, but the head bears only two transverse
ridges, and the thorax shows only vestiges of the anterior
depressions. O, Antillarum is allied to O. marginicollis,
Harold, which js similar in size and form, while the traces
of yellow marking often present in the new species also
point to this affinity. It is less shining, however, the
striz are not punctured, and the horns of the male, although
not longer, are more slender.
APHODIID A.
Aphodius luridus, Olivier.
A single specimen of this almost universally distributed
species was found on the Leeward side of Grenada.
Aphodius cuniculus, Chevrolat.
This was found in numbers in St. Vincent, Grenada, and
Mustique, and is no doubt distributed throughout the
Antilles. It also occurs on the American mainland.
Atenius strigicauda, Bates.
This widely-distributed species was found in numbers
in St. Vincent and Grenada, and a specimen was also
brought from Becquia Island. It generally occurred
beneath rubbish in swampy places.
512 Mr. G. J. Arrow on the Laparostict Lamellicorn
Atenius frater, sp. n.
A. strigicaude forma et magnitudo, sed differt capite crebre
punctato, haud ruguloso, elytrorum interstitiis minus convexis,
sparsissime punctulatis, metasterno femoribusque omnibus disperse
et grosse punctatis.
Long. 5-55 mm.
Hab. St. VINCENT, South end; GRENADA, Leeward side.
At first sight this species is indistinguishable from
A. strigicauda, but a close examination reveals slight
differences which in combination sufficiently establish its
distinctness. The punctures on the head are very fine,
and, though not uniformly distributed, are everywhere
distinct, and do not change into rstaltdtes anteriorly as in
A, strigicauda. The elytral interstices show lateral strive
behind, as in that species, but unlike it the interstices are
strewn with minute punctures. The evenly-punctured
surface of the metasternum and middle and hind femora
perhaps constitute the most evident difference between
the two, these in A. strigicauda being smooth except for
clusters of about half-a-dozen punctures on each side of the
median furrow of the metasternum and at the distal
extremities of the femora.
This appears to be a very local species, but considerable
numbers were found at the southern end of St. Vincent
and a few specimens in Grenada (Grand Etang Lake and
Mount Gay Estate). It occurred under the same conditions
as the allied form, but in different places.
Atenius terminalis, Chevrolat.
Numerous specimens were found in Grenada of the form
referred to by Chevrolat as inhabiting most of the Antilles.
Its apparent absence from St. Vincent is rather remarkable.
All our specimens have the yellow border extending round
the elytra from shoulder to shoulder.
Atexnius tenebrosus, sp. 0.
Niger, opacus, griseo-vestitus, latus, parum convexus, capite densis-
sime rugose punctato ; anaiihannes creberrime sed distincte punctato ;
elytris striatis, striis haud profunde punctatis, interstitiis subcari-
natis, haud ie 3 prothoracis lateribus medio fere rectis, angulis
eee arcuatis,
Lat.2mm. Long. 4°5 mm.
Coleoptera of Grenada and St. Vincent (W. Indies), 513
Hab. GRENADA; TRINIDAD; BraziL, Marajo, ete.
A single specimen was found by Mr. Smith in a marshy
place on Mount Gay Estate, Grenada. It agrees with a
specimen in the British Museum from Trinidad and others
from Brazil bearing the unpublished name tenebrosus,
Reiche. The species is very near A. tmbricatus, Melsh.,
which has been recorded from Cuba by Harold under the
name of A. sordidus. A. tenebrosus, however, is a broader
insect, more finely and densely punctured on the head and
thorax, and without visible setze upon the elytral coste.
Alxnius Vincentix, sp. 0.
Valde elongatus, nigro-piceus, opacus, clypeo, prothoracis margine
antica pedibusque rufis ; clypeo bidentato, dentibus minutis, capite
antice nitido, transversimn rugato, postice crebre punctato ; prothorace
minute, fere confluenter, punctato, angulis omnino obliteratis, sulea
obsoleta media postice impresso; elytris anguste carinatis, ubique
punctato-rugoso.
Long. 3°3 mm.
Hab. St. VINCENT, Leeward side and South end.
This was always found near the sea, beneath vegetable
débris, It is a species closely allied to A. scalptifrons,
Bates, and A. Steinheili, Harold—especially the former. It
is more elongate, however, the posterior angles of the
thorax are completely obliterated, and the head is more
coarsely sculptured and shining,
Atenius polyglyptus, Bates.
A number of specimens agreeing with the variety
jalapensis of Bates occurred both in St. Vincent and
Grenada. They were also found under rubbish, but
generally beside forest streams at an elevation of 250—
1000 feet.
- Atxnius gracilis, Melsh.
Several specimens were found at Balthazar and Mount
Gay Estate in Grenada, and one in St. Vincent. The
species occurs throughout the Western Hemisphere.
Atzxnius, sp.
A single specimen from Mustique Island of a species
near A. polyglyptus, but probably new.
514 Mr. G. J. Arrow on the Laparostict Lamellicorn
Psammobius parvulus, Chevr.
This minute insect was very commonly found on the
wing at sunset in both St. Vincent and Grenada. The
species was described from a single Cuban specimen which
I have not been able to trace, but M. Oberthiir has
kindly sent me specimens from Cuba which there is no
reason to doubt belong to Chevrolat’s species, and I am
not able to distinguish our series from these. The
Psammobius from Mexico and Guatemala identified by
Bates as P. parvulus, although closely related, is not the
same, and the Amazonian specimens also ascribed to it by
him, and which I have seen in M. Oberthiir’s collection,
represent a third species.
For the sake of clearness it may be well to re-name the
Central American form shortly described by Bates, which
I will call P. Batesi. This is generally larger than
P. parvulus and considerably stouter, the width of the
elytra together being greater than that of the thorax,
whereas they are of equal width in Chevrolat’s species,
producing a very characteristic linear form. The head
and prothorax are less markedly darker than the elytra in
P. Batesi, and the thoracic puncturation is sparser.
Saprosites grenadensis, sp. 0.
S. parallelo affinis sed paulo minor, minus grosse punctatus. Rufo-
piceus, capite prothoraceque magis rufescentibus, prothorace haud
crebre sed irregulariter punctato, elytris profunde striatis, striis
fundo punctatis, metasterno late canaliculato, segmentis abdominali-
bus subtus minutissime punctatis; tibiis anticis extus minute
serratis, dentibus tribus majoribus.
Long. 4 mm.
Hab. GRENADA, Grand Etang (1900 feet).
A single specimen only was found under the bark of a
decaying log. Its colour is distinctly red on head and
prothorax, but this is probably not a constant specific
feature. It differs from S. parallelus, Harold, of Central
and South America, by its smaller size and finer punctura-
tion. The elytral strize are deeper, and the punctures of
which they are formed less distinctly traceable. The
abdomen, which is smooth in S. parallelus, is very thickly
and finely punctured. As in the other species, there is a
minute tooth between the two upper of the three large
teeth of the anterior tibia.
Coleoptera of Grenada and St. Vincent (W. Indies). 515
ORPHNID.
Hygidium vincentig, sp. n.
Parum elongatum, nigro-piceum, capite crebre punctato, clypeo
antice leviter acuminato, prothorace grosse punctato, (¢, medio
longitudinaliter excavato, minus dense punctato) lateribus valde
arcuatis, grosse crenulatis, setigeris, postice perspicue marginato ;
scutello levi, modice elongato ; elytris vix quam eapite et prothorace
longioribus, ical wbique ennnvethe: haud costatis, dense strigose
punctatis, punctis setigeris.
Long. 11 mm.
Hab. St. VINCENT, La Souffriere Volcano, Petit Bordelle
Valley, 1000—2500 feet.
Two specimens, one of each sex, were found under logs.
The male is probably almost at the minimum develop-
ment. The species is near 4. parvulum, Westw., but
rather more elongate, although considerably shorter than
4. colombianum, Westw. The elytral costze have almost
entirely vanished. The sides of the prothorax are much
more ‘distinctly and regularly crenated than in &. parvu-
lum, and the flattened hind margin is broader. ,
It must be noted here that the Central American insects
described by Bates as varieties of £4. colombianum are
quite distinct specifically from that form. Both sexes
are smaller and relatively shorter, much more strongly
punctured, especially upon the head and metasternum,
and with a less narrow and parallel-sided scutellum. In
the male the thorax is almost rugosely punctured, whereas
in Westwood’s species it 1s polished and shining, Although
in the latter there are large scattered punctures in the
dorsal cavity, they are of a different nature, and do not
interfere with the smoothness of the surface.
4{. colombiaxum seems to inhabit Western South
America as far south as Chili, from which country there
are two female specimens in our collection which cannot
be distinguished from it. The Central American species
ranges from Nicaragua to Bogota. It should be called
4h. cribratum, that name having been given to the typical
specimen figured by Batesas a variety of &. colombianwm.
It is curious that a remarkable sexual characteristic of
these insects has been overlooked, although Westwood has
figured the structure of the genus in such detail. The
front tibia of the female has three external teeth and a
very strong short spine internally. In the male this spine
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.) 35
516 Mr. G. J. Arrow on the Laparostict Lamellicorn
is wanting, but is replaced by a supplementary tooth
anterior to the others and directed internally, so that the
tibia becomes a broad digging instrument, recalling the
same member in the mole-cricket.
HyBosoriD&.
Celodes nigripennis, sp. n.
Castaneo-rufus, elytris nigris; clypeo breve, fere rugoso, fronte
polito, paulo tumido ; prothorace minutissime et sparsissime punctato
(¢ antice medio leviter excavato) ; scutello polito ; elytris lineato-
punctatis, lineis haud perspicue geminatis, linea suturali profunde
impressa ; pedibus corporeque subtus flavo-rufis, tibiis anticis parum
longis, extus haud copiose denticulatis, dentibus tribus majoribus,
quorum medio maximo,
Long. 4°5 mm.
Hab. St. VINCENT, Leeward side.
The species of Calodes are very much alike in their
general form and appearance, and require careful examina-
tion to di stinguish them, C., nigripennis is a small species
about the size of CL parvulus, Westw., but besides the
difference in the colour of the elytra these are rather more
finely and uniformly punctured, the rows of punctures
being less evidently paired. The clypeus is short, not
much narrowed anteriorly, and shining, but not smooth
The anterior tibia is not elongated, the apical tooth is
short and rounded, and the minute denticulations are few.
TROGID.
Trox suberosus, Fab.
Specimens of this were brought from St. Vincent by Mr.
Smith, who has noted it as occurring commonly in fungi.
It is strange that it was not found in Grenada, as it is a
very common species distributed over the greater part of
the New World.
Acanthocerus relucens, Bates.
The recorded range of this species is from Jalapa, in
Mexico, to Panama, but it will probably be found to have
a much wider distribution. Five specimens were brought
by Mr. Smith from St. Vincent agreeing with the type in
all essentials. The lines of punctures upon the elytra are
Coleoptera of Grenada and St. Vincent (W. Indies). 517
exceedingly faint, and the marginal line of the thorax
is rather further from the anterior edge at the angles.
But the species is in certain respects very variable. A
remarkable peculiarity of this genus, which has not
previously been noticed, is the extreme variability in size
of the division of the eye situated upon the upper part of
the head, which in this species diminishes from a large,
almost circular disc to a mere narrow vestige, and in a
larger series would probably be found to vanish altogether,
as it has done in some specimens of Acanthocerus vicarius,
Bates, which I have examined. ‘To show how remarkable
is this peculiarity I have sketched the heads of a series of
four specimens of Acanthocerus relucens, brought respect-
ively from St. Vincent, Mexico, Guatemala and Honduras.
As far as our materials show, individuals from the same
place are similar, but there is no real segregation into
geographical races, nor is the variation in the eyes corre-
lated with the variation in other respects to which
the species is subject. All the specimens of A. relucens
from St. Vincent have the eyes well developed above.
These specimens were found in different parts of the
island and at different times of year.
. Cleotus rufopriceus, sp. n.
Paulo elongatus, rufo-piceus, nitidus, capite strigoso-rugoso, medio
leviter elevato, clypeo antice late arcuato, medio vix angulato ; pro-
thorace erebre strigoso-punctato, undique marginato, angulis anticis
productis, posticis late arcuatis; scutello parce punctato ; elytris-
omnino punctato-striatis, postice multo profundius ; pedibus lte
rufis, tibiis anticis acute serratis, dentibus tribus apicalibus majoribus,
518 Mr. G. J. Arrow on the Laparostict Lamellicorn
tibiis intermediis et posticis latis, supra politis, subtus longitudinaliter
striatis, tarsis omnibus subtus dense fulvo-pilosis.
Long. 5 mm.*
Hab. St. VINCENT, Chateaubelair, Petit Bordelle Valley ;
GRENADA, Grand Etang (1900 feet).
This species has about the size and shape of C. brunnipes
and C. bidens, but is more strongly punctured. Its non-
metallic deep red colour distinguishes it from all the other
species known to me.
It occurred under bark and on brushwood.
Claotus crassicollis, sp. 0.
Niger, nitidus, prothoracis lateribus pedibusque rufopiceis, capite
tumido, rugoso, medio sulea transversa curvata levi impresso, clypeo
transverse strigoso, margine arcuato ; prothorace irregulariter punc-
tato, marginibus lateralibus (postice nonnihil extensis) valde incras-
satis, linea marginali impressa completa demarcatis, margine posteriori
medio fere angulato, angulis anticis et posticis curvatis ; scutello
parvo, polito, epimeris mesosternalibus supra vix perspicuis ; elytris
conjunctim exacte hemisphiericis, lineato-punctatis, punctis elongatis,
setas albas decumbentes ferentibus, interstitiis postice et lateraliter
fortiter et acute elevatis, costas interruptas formantibus, humeris
tuberculiferis; tibiis quatuor anterioribus paulo longis, anticis extus
minute serratis, singulo apice dentibus duobus majoribus approxi-
matis, intermediis leviter incurvatis (¢, intus apice fortiter curvato-
producto), posticis paulo brevioribus, subcompressis, posterioribus
quatuor longitudinaliter striatis.
Long. 6 mm.
Hab, Sv. VINCENT, Morne a Garon (1500 feet).
Two specimens of this, apparently representing the two
sexes, were found in rotten wood. It is a very well-
marked form, which may be compared with C. posticus,
Germ., although not closely related to that or any other
known species. It is of similar colour and form to
C. posticus, although rather larger, and the elytral cost
are similar, but much more prominent, and extending over
half the total area of the elytra. The lines of punctures
are fewer, and each puncture contains an elongate white
scale. The most distinctive feature, however, is the
* As these insects have been measured sometimes in the rolled-up
and sometimes in the extended condition, it is necessary to say that
I have given measurements of them in the condition of normal
extension.
Coleoptera of Grenada and St. Vincent (W. Indies). 519
conspicuous submarginal thickening of the pronotum
extending on each side from the anterior angle to about a
quarter of the length of the base.
While dealing with Tropical American species of Clwotus,
I may conveniently point out that Bates has wrongly
recorded C. metallicus, Har., from Central America. In
the Biologia Cent.-Americana he has referred a specimen
from Panama and another from Costa Rica to that species,
but the two are not conspecific, nor is either C. metallicus.
The latter was described from specimens collected by
Bates at Ega, of which a series is in the British Museum.
It is distinguished from both the other forms by the serrate
humeral margins of the elytra, the three acutely-raised
interstices at their apical end, and the narrower and more
curved intermediate tibiz, which have three or four strize
on their lower face.
The specimen from Bugaba, Panama, agrees with another
from the same locality referred by Bates to his C. vividi-
pennis, of which all true examples are from Chiriqui. The
distinctive characters of this undescribed species are shortly
diagnosed as follows :—
Claotus bugabensis, sp. n.
C. viridipenne valde affinis, differt magnitudine paulo minore, colore
magis cupreo, prothoracis disco impunctato, linea marginali antice
medio interrupto, tarsisque anticis tenuioribus.
Long. 5 mm.
Hab. PANAMA, Bugaba.
The Brazilian species which appears to have been
mistaken by Bates for C. metallicus, Har., and from which
I cannot distinguish the Costa Rican specimen mentioned
above, is also characterized here :-—
Cleotus acutipes, sp. 0.
C. metallico et viridipenne valde affinis, sed tibiis anticis fortiter
dentatis, dentibus tribus apicalibus majoribus: tibiis intermediis
paulo latis, subtus levibus, prothorace prope lateribus solum leviter
punctato ; elytrorum marginibus humeralibns haud serratis, inter-
stitiis apicalibus leviter hand acute elevatis.
Long. 7 mm.
Hab. Brazit, Rio de Janeiro; Costa Rica, Irazu.
This is no doubt a widely-distributed species.
520 Mr. G. J. Arrow on the Laparostict Lamellicorn, ete.
In conclusion I describe, supplementary to my previous
paper, a Pleurostict species belonging to a genus of Melo-
lonthidz not hitherto recorded from the West Indian
Islands, a single representative having been found in Mr.
Smith’s collection since the publication of my enumeration
of the Pleurostict Lamellicornia.
Faula insularis, sp. 1.
Ferruginea, prothorace nigro-maculato, elytris paulo’ infuscatis,
capite supra granuloso, elypeo arcuato; prothorace punctato-rugoso,
macula magna media duabusque parvis lateralibus nitidis, media
nigra ad margines antice et postice fere attingente, postice dilatata,
pronoti lateribus valde angulatis, angulis posticis acutis, margine
postica spina acuta medio instructa ; scutello fere circulari, grosse
punctato, antice profunde indentato ; elytris ad humeros quam pro-
thorace paulo Jatioribus, deinde leviter ampliatis, ante medium
usque ad apices angustatis, punctato-rugosis, dense breviter erecte
pilosis, vage costatis, humeris prominentibus, nitidis; pedibus graci-
libus, rufis, tibiis anticis tridentatis, dente superiori ad medium
posito.
Long. 10 mm. Lat. max. 5 mm.
Hab, Sv. Vincent, Leeward side (500 feet).
A single female specimen of this species was found. It
is easily recognizable by the smooth black area shaped like
a vine-leaf occupying the centre of the prothorax. In the
shape of the scutellum and the sharp tooth m the middle
of the hind margin of the thorax, which fits into the deep
notch in the former, it resembles Ceraspis and Ancistrosoma,
but it has the typical form and size of Fawla as well as the
entire claws distinctive of that genus.
XXII. Note on the habits of Chironomus (Orthocladius)
sordidellus. By THomas HaArotp Taytor, M.A.,
E.ES.
[Read October 7th, 1903. ]
THE larva of this insect lives submerged in brisk streams.
It has the usual structure of a bloodworm, but like other
surface-feeding Chironomus-larvee, it is greenish in colour,
and lacks the ventral gills on the last segment but one.
When full-grown it measures 7 mm. in length. This
larva constructs for its abode a hollow cylindrical tube, one
end of which it attaches to the surface of a stone, leaving
the other end free and open. The material of the tube is
furnished by the secretion of the salivary glands without
any admixture of foreign substances. Within the long
and flexible silken tube the larva may be seen creeping to
and fro, or, if stationary, bending its body up and down in
the undulations often practised by tube-dwelling larve,
as a means of renewing water which has been vitiated by
respiration. Pure water is admitted by special inlets,
which may be either mere slits or roundish holes. The
food of the larva consists of unicellular and filamentous
alge. Although these grow in abundance on the stones
around, the strength of the current, and the incessant
oscillations of the tube make it a matter of some difficulty
for the larva to browse upon them. But since the tube
itself soon becomes overgrown with diatoms and other
microscopic algze, the larva, thrusting its head out of the
terminal opening and holding on by its anal feet, is able
to search the exposed surface of its own tube. The range
of its body is of course very small, but the tube is flexible,
and admits of being bent upon itself or even doubled in
two. By reaching out, the larva can draw up to its mouth
any part of the surface of its tube, which thus not only
affords lodging, but board as well.
When the larva is full-grown, it transforms the old tube
into a pupal case. It fetains the free end for its abode,
and converts the rest into an anchoring strand by drawing
the sides together with threads of silk. The free end
becomes dilated into an oval chamber, the terminal aper-
ture of which is somewhat more narrowed than in the
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.)
522 Mr. T. H. Taylor’s Note on the
larval dwelling. The last thing the
larva does before pupation is to make
a new aperture at the fixed end,
which may be distinguished as the
front opening. This aperture is
seldom single; generally two, three
or even four small circular holes are
arranged in a ring around the attach-
ment of the stalk. The advantage of
several small holes over one large one
is obvious; they do not weaken the
strand so much, nor are they so
liable to be torn by the force of the
stream. Whenall is ready the larva
pupates ; the old larval skin is usually
allowed to float out of the tube, for it
is seldom seen afterwards.
The pupa of this species of Chirono-
mus has minute prothoracie horns in
which no apertures are visible, and
an effective tail-fin. It lies in the
chamber with its head towards the
attached end, that is, pointing up-
stream. It maintains an undulatory
movement with its body, which causes
water to be drawn in at the front
opening and passed out behind. As
the front opening looks up-stream
and the hinder down-stream, it follows
that the course of the respiratory cur-
rent is parallel to and in the same
direction as the flow of the stream.
When, in the course of a few days,
the fly is ready to emerge, the pupa,
by means of the hooks on its body,
forces a way through the front open-
ing. It then rises to the surface of
the water, when the skin at once
splits open along the back and allows
the fly to escape.
It frequently happens that while
the pupa is still within its case, the
six-legged larva of a water-mite is
found attached to its body, generally Fig. 1. Larval tube of
_ ; Chironomus (Orthocla-
on the upper part of the thorax. dius\ sondage!
habits of Chironomus (Orthocladius) sordidellus, 528
Its colour, which is bright red,
renders the mite conspicuous
even through the wall of the
tube. The mite does not appear
to do any injury to its host,
even when, as sometimes occurs,
two are present on the same
pupa. It has not been seen
upon the larva. When the pupa
is extricating itself from its
chamber, the mite occasionally
becomes dislodged, and is left
behind, evidently ill at ease, as
its agitated movements show.
Generally, however, the mite is
carried up to the surface of the
water together with the pupa
itself. And then an unexpected
thing happens. One might
imagine that when the fly
emerges, the mite would remain
attached to the pupal skin ; but
such is not the case. When the
pupa rises to the surface, the
mite is apparently aware of what
is going on, and, all alert, at the
very moment when the old skin
cracks, it releases its hold and
transfers itself to the body of
the half-extricated fly. It is
interesting to watch the mite,
hitherto perfectly passive, exe-
cute this rapid and sudden move-
ment. ‘Thus when the fly leaves
the water and rises into the air,
the mite, still a six-legged larva,
is borne along with it. The
ultimate fate of the mite can
only be guessed at.
My best thanks are due to
Professor Miall for his valuable ~
assistance during the preparation
of this note.
Fig. 2. Pupal tube of Chirono-
mus (Orthocladius)
sordidellus x 8.
( 525)
XXIIL. Additions to the Rhynchotal fauna of Central
America. By W. L. Distant, F.E.S.
[Read October 7th, 1903. ]
My last contribution to our knowledge of the Rhynchotal
fauna of Central America appeared-in the pages of our
Transactions for 1900 (p. 687). The following descriptions
and notes refer to a sinall collection recently sent me by
Prof. P. Biolley, and which was made in Costa Rica, It
contained three new species, and two others not enumerated
in the “ Biologia Centrali Americana.”
HETEROPTERA.
Family PENTATOMIDA.
CYDNINA.
Cyrtomenus vestigiatus, sp. 0.
Piceous, corium more or less piceous-brown ; legs, antennee and
rostrum, pale castaneous ; head broad, rounded, lateral lobes some-
what transversely rugose, lateral margins of the central lobe strongly
defined, lateral lobes not meeting in front of central lobe, and
thus cleft at their apices ; pronotum with a strong transverse im-
pression near middle, this impression and the lateral and anterior
margins, sparingly, strongly, linearly punctate, a few coarse punctures
on anterior disk of posterior lobe; scutellum very sparingly but
very coarsely punctate, its apex impunctate, and distinctly posteriorly
depressed ; corium somewhat thickly and finely punctate ; membrane
pale ochraceous, longly passing abdominal apex ; rostrum reaching the
intermediate coxee, its apex piceous; intermediate and posterior
femora with a few coarse lateral punctures ; tibize with outer series
of strong piceous spines, inwardly longly hirsute.
Long. 8 to 9 millim.
Hab. Costa Rica, San José (Biolley).
Taken at electric lamps.
Allied to C. mirabilis, Perty, from which it differs by its
shorter and more robust form, and its more sparsely and
deeply punctate scutellum.
TRANS. ENT. SOC. LOND. 1903.—PART Iv. (DEC.)
Mr. W. L. Distant’s Additions to the
Or
bo
lon)
PENTATOMIN A.
Hdessa cervus.
Kdessa cervus, Fabr., Mant. Ins., 11, p. 283 (1787).
Hab. Costa Rica, Surubres, near San Mateo (Biolley).
Not previously recorded from Central America.
Family LYGAIDA.
" APHANIN ES.
Gonatas costaricensis, sp. 0.
Piceous-brown ; head, anterior lobe of pronotum, scutellum, and
body beneath, piceous ; veins and lateral margins of corium, rostrum,
antenne, and tibie, luteous; antennw with the first, second, and
third joints hirsute, fourth pilose, second joint longest ; head glabrous
with a few long hairs at apex ; pronotum with the lateral margins
longly pilose, anterior lobe glabrous, posterior lobe sparingly punctate
and somewhat obscurely pilose, lateral posterior angles slightly
nodulose ; scutellum long, triangular, foveately discally ceed a
few scattered hairs on posterior half, anterior half glabrous ; corium
distinctly ochraceously pilose, and punctate between the veins,
margins of clavus, luteous ; intermediate and posterior tibia some-
what longly setose ; membrane subhyaline.
Long. 5 millim.
Hab, Costa Rica, Surubres, near San Mateo (Biolley).
Allied to G. diversus, Dist., but ditfering by the glabrous
head, anterior pronotal lobe, and anterior half of scutellum ;
corium shining, punctate, without spots, veins and lateral
margins luteous ; pronotum shorter, ete.
HOMOPTERA.
Family CICADID.
Odopea liolleyi, sp. n.
¢. Body dark chocolate-brown ; vertex of head—excluding a few
obscure spots, lateral and posterior margins of pronotum and two
oblique discal spots at both anterior and posterior margins, two
discal linear posteriorly broadly angulated spots, bordered on each
side by a curved oblong linear spot, and some obscure linear
markings to mesonotum, basal cruciform elevation, sternum, opercula
and a broad basal annulation to tibize, obscure olivaceous-green ;
Rhynchotal fauna of Central America. 527
abdomen beneath thickly, ochraceously, very finely pilose. Tegmina
pale dull ochraceous, semihyaline ; costal membrane and veins,
brownish-ochraceous, or very obscure olivaceous, apical two-thirds
somewhat thickly spotted with fuscous, the spots almost entirely
situate on the veins ; wings a little paler and subhyaline, the veins
brownish-ochraceous, but unspotted.
Long. excl. tegm. ¢ 32 millim, Exp. tegm. 85 millim.
Hab, Costa Rica, Cerro Carisia (Biolley).
Alhed to O. medea, Stal.; body much longer and more
robust; tegmina pale ochraceous and with the apical two-
thirds of venation somewhat thickly spotted; opercula
much more truncated interiorly, and therefore more widely
separated ; colour altogether darker, ete.
Carineta postica.
Carineta postica, Walk., Ins. Saund. Hom., p. 238 (1858).
Hab. Costa Rica, Cariblanco, Sarapiqui (Ch.
Lankester) ; COLOMBIA.
Not previously recorded from Central America.
XXIV. Notes on some Central and South American
Erycinidee, with descriptions of new species. By
FREDERICK Du CANE GopmAN, D.C.L., F.R.S., etc.
[Read October 7th, 1903, ]
PEATES" XOXOXO XN PAN De kee TE
THIS paper includes descriptions of various South American
Erycinide that have been accumulating for many years in
my collection. The opportunity is also taken to correct
certain synonymy, and of figuring a few forms of which one
sex only was previously known, or which had been figured
from specimens in bad condition.
Mr. Schaus [Proc. U.S. Nat. Mus. xxiv, pp. 397-406
(1902)] has recently described twenty-six species of this
group from Central or South America. One of these
(Hurygona tarinta) I had already characterized in MS.,
and a figure of this insect and three others are added
in the accompanying plates.
MESOSEMIA PHACE, sp. n. (Plate XX, fig. 1, t.)
¢. Alis nigro-fuscis, anticis ocello nigro ad cellule finem albo tri-
punctato, lineis quatuor transversis, una ad basin, secunda et tertia
ultra eam ocellum includentibus, quarta arcuata latiore a costa ante
apicem ad angulum analem extendente, viridibus (aut ceruleis) ;
posticis lineis “angustis septem transversis coloris ejusdem ; subtus
griseis, anticis ocello ut supra sed linea ochracea circumcincta, lineis
undulatis fuscis per alarum medium, altera latiore margini exteriori
propiore ; posticis lineis angustis variis undulatis coloris ejusdem,
anticis et posticis macula in medio marginis externi nigra, illa in
posticis multo majore.
Hab, British GUIANA, Roraima, Quonga (Whitely).
Four specimens from Guiana, all males, three from
Roraima and one froin Quonga. The stripes crossing the
wings seen in certain lights are of an emerald green
colour, while in other positions they are of a dark blue.
Allied to JZ. bella, Sharpe, but may be distinguished by
having more transverse bands on the upper side.
TRANS. ENT, SOC, LOND. 1903.—PART IV. (DEC.)
530 Mr. F. Du Cane Godman on
MESOSEMIA THERA, sp. n. (Plate XX, fig. 2, 2.)
¢. Alis nigro-fuscis, anticis ocello indistinctoad cellule finem nigro,
albo-punctato, fasciis tribus submarginalibus communibus exteriore
angustissima, costa anticarum et ad alarum basin céruleis; subtus
sriseo-fuscis, anticis ocello sicut supra, sed punctis tribus albis no-
tatis et lineis obseurioribus incertis circumcinetis, fascia lata sub-
marginali coloris ejusdem ; posticis lineis angustis undulatis variis
fuscis, puncto nigro in cellula ornatis.
Hab. BRazt., Chapada (H. H. Smith).
A single male much resembling J/. orbona described
below. It may, however, be readily distinguished from
that species by the ocellus in the primaries being smaller
and less conspicuous; the blue stripes, too, on both wings
are much narrower.
MESOSEMIA ORBONA, sp. n: (Plate XX, fig. 3, 2.)
d. Alis nigro-fuscis, anticis ocello magno ad cellule finem nigro
albo-pupillato et czeruleo circumcincto, fasciis duabus communibus
submarginalibus, tertia prope basin (in posticis angustissima) et ad
basin ipsam ceruleis ; subtus griseo-fuscis, anticis ocello sicut supra
sed albo-tripunctato, lineis duabus obscurioribus circumeinctis, fascia
latiore margini propiore coloris ejusdem ; posticis lineis variis an-
custis undulatis (marginem versus latioribus) obscuris, puncto in
cellula nigro.
Hab. SURINAM, Paramaribo (ea Semper’).
Dr. Semper sent us some years ago a single male
example of this species in bad condition, under the name
of IL. ephyne, Cram., but it does not agree with the figure
of that insect. It is not very unlike the Colombian J/.
thetis, G. and S., but the arrangement of the bands is
different, those towards the outer margins especially being
much wider and of a brighter colour. J/. orbona also
resembles J/. bella, Sharpe, from Brazil, but the blue at
the base of the wings on the upper-side is interrupted by
a dark transverse band, and it is likewise different beneath.
CREMNA MALIS, sp. n. (Plate XX, fig. 4, %.)
¢. Alis fuscis maculis albis transversis notatis, anticis exterioribus
lunulatis ; posticis maculis submarginalibus valde angulatis, aliis
irregularibus interioribus albis, inter lineas et in margine exteriore
some Central and South American Erycinide. 531
squamis ceruleo-griseis indutis ; subtus pallidioribus, colore albo
magis extenso et dimidio basali maculis nigro-limbatis,
Hab. W. Cotomsia, San Pablo, Rio San Juan (ex
Staudinger).
Very like C. beltiana, Bates, from North Brazil (of which
the female only is known), but the white spots on the
upper-side are much smaller, the marginal ones being
obsolete ; and the posterior wings towards the anal angle
are clothed with bluish-grey scales, which show a tendency
to form a band between the two submarginal white lines.
HYPHILARIA ORSEDICE, sp. n. (Plate XX, fig. 5, 9.)
Q. Alis pallide flavis, anticis costa et triente distali fuscis, linea
submarginali ad costam in maculas fracta, ad angulum analem angus-
tissima, notata; posticis margine externo late fusco, lineis duabus
(exteriore angustissima) pallide flavis ; subtus sicut supra sed anticis
prope marginem externum maculis variis quoque flavis notatis.
Hab, VENEZUELA (Mus. Brit.); BRITIsH GUIANA,
Roraima ( W/itely).
Similar to H. nicias (Stoll), but without the transverse
bands crossing the wings at the end of the cell. Two
specimens in our collection and one in the British Museum.
EURYGONA AURANTIACA.
Hurygona elmira, Hew., Ent. Monthly Mag. vi, p. 226 (1870).
In the “ Biologia” we omitted to note that Hewitson
had described /. aurantiaca, Godm. and Salv., under the
name of #, elmira; the former, however, has two years’
priority.
EURYGONA LICINIA, sp. n. (Plate XX, fig. 6, 2.)
?. Alis fuscis, posticis litura magna alba rotundata prope mar-
ginem externum ad angulum analem extendente ; subtus griseis,
posticis dimidio distali fere albo, linea communi a costa anticarum
ultra cellulam ad medium marginis interni extendente ferruginea,
ultra eam in anticis fascia transversa obscura, posticis macula ad
medium marginis externi, aliisque minutis angulum analem versus,
margine externo ipso flavo limbato. ;
Hab. BRITISH GUIANA, Roraima (Whitely).
On the under-side this insect closely resembles the cor-
responding sex of #, mys, H.S., but the large white patch
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.) 36
532 Mr. F. Du Cane Godman on
occupying nearly half the upper-side of the secondaries at
once separates it from that species. I have a very long
series of both sexes of #. mys, but none of them show any
white on the upper-side. £. effinia, Hew., from Ecuador,
somewhat resembles £. licinia above, but is quite different
beneath. One specimen.
EURYGONA AUTHE, sp. n. (Plate XX, fig. 7, 3.)
¢. Alis fuscis, anticis litura magna infra cellulam ad marginem
internum extendente rufo-aurantia, hine ad basin ferruginea ; pos-
ticis fere eodem modo notatis, sed litura aurantia ultra (nee infra)
cellulam posita ; subtus pallide fuscis, fascia communi a cellula anti-
carum ad marginem internum prope angulum analem posticarum
extendente alba, intus linea fusca limbata; posticis punctis in serie
submarginali nigris albo circumcincetis, illo in medio maximo,
Hab. BRazit, Chapada (1. H. Smith).
Very like £. eubule, Feld., from Central America, but
with the orange colour on the secondaries not extending
to the outer margin. On the under-side, too, there is a
broad white transverse band nearly crossing both wings.
EURYGONA RHODOGYNE, sp.n. (Plate XX, figs. 8, ¢; 9, 2.)
dé. Alisnigro-fuscis ceruleo tinctis, anticis fascia obliqua prope cos-
tam ultra cellulam fere ad medium marginis externi extendente rufo-
aurantiaca; subtus rufo-griseis, fascia ferruginea communi a costa
anticarum ultra cellulam ad medium marginis interni posticarum
extensa, altera sinuata (in anticis fere obsoleta) exteriore, anticis mar-
gine interno grisescentiore ; posticis ocello parvo, extus albo notato,
in medio marginis externi, aliisque minimis in serie angulum
analem versus.
@. Majore et multo pallidiore, fascia anticarum multo latiore,
Hlab, COLOMBIA (Wheele7).
One pair, not very unlike Z. ewhemerus, Hew., from the
Amazons, of which the female only has been described ; it
differs, however, in having the band on the primaries
longer and less graduate, and in the female reaching the
costal margin; the markings, too, on the under-side are
more distinct. The blue colour on the upper surface is
only visible in certain lights, and is entirely wanting in
the female. Of 4H. ewhemerus I have two females obtained
by Bates at Para,
some Central and South American Erycinide. 588
EURYGONA TARINTA. (Plate XX, fig. 10, 4)
Hurygona tarinta, Schaus, Proc. U.S. Nat. Mus. xxiv,
p. 398 (1902).
Hab, COLOMBIA.
I now have three males of this species, two sent by Dr.
Staudinger and one by Wheeler, the latter being labelled
as coming from the neighbourhood of Bogota. It is allied
to LH. euwpepla, G. and 8.,* but differs in having the blue
lustre on the upper-side brighter and on the primaries
confined to the outer half. This colour is seen only when
the insect is held in a certain light. The female is pro-
bably of a pale fuscous colour with a broad white band
crossing both wings, as in Z. ewpepla.
EKURYGONA FABIA, sp.n. (Plate XX, fig. 11, 2%.)
dg. FH. tarinte similis sed dimidio anticarum et margine externo
posticarum purpurascentioribus; subtus magis ferrugineis, fascia
externa fere obsoleta, posticis ocello ultra ineam albam absente.
Hab. EK. Peru, Pebas (ee Staudinger), Sarayacu
(Buckley).
Of this insect I have three males, and have seen others
from San Paulo in the late Dr. Staudinger’s collection.
E. fabia is a very close ally of H. ewpepla, from which it
differs in having the blue of the fore-wing less extended,
and on the under-side it is of a more ferruginous colour,
the ocellus of the hind-wing, too, is not edged with white
externally.
MESENOPSIS PULCHELLA, sp. n. (Plate XX, fig. 12, 9.)
2. Alis nigris, litura lata mediana a basi extensa (ultra cellulam
albo-terminata) rufo-aurantia ; posticis quoque litura mediana a basi
fere ad marginem externum extendente ; subtus sicut supra sed ciliis
ad angulum analem posticarum albis ; corpore supra nigro ; palpis
et pedibus ochraceis; capite, collo et ad abdominis basin aurantiacis,
Hab, AMAzONS, Massauary (ex Staudinger).
A very close ally of JZ melanochlora, G. and S., from
Central America, but differs from it in having the broad
* Mr. Schaus quotes this insect under the name of £. eupiola,
Hew. This is doubtless an error, as Hewitson described no such
species,
534 Mr. F. Du Cane Godman on
median streak on the primaries less extended outwardly
and in being white beyond the celi; the streak on the
secondaries, too, does not reach the outer margin. One
specimen.
CHAMAELIMNAS PANSA, sp. n. (Plate XX, fig. 13, 3.)
¢. Alis nigrescentibus, anticis fascia angusta obliqua a vena sub-
costali vix ad marginem externum prope angulum analem extendente
pallide flava ; subtus sicut supra, sed anticis margine interno paulo
pallidiore ; collo flavo.
Hab. Brazit, Chapada (/, H. Smith).
I have five males of this insect from Chapada and a bad
specimen from Semper’s collection. The species is very
like Felder’s figure of C. tivcis, from Bahia, but the trans-
verse band of the primaries extends almost -across the
wing. From the Amazonian C. iwris, Bates, it differs in
having a longer and paler band on tle primaries.
CHAMALIMNAS JOVIANA. (Plate XX, fig. 14, 2.)
Chamelimnas joviana, Schaus, Proc. U. 8. Nat. Mus.
xxiv, p. 401 (1902).
Of this species, which I take the opportunity of figur-
ing, I have numerous specimens from both Bolivia and
Brazil; some of these were sent by Staudinger under the
name of Jsapis joviana. C. similis, Schaus, which occurs
with C. joviana at the same locality (Bueyes in Bolivia),
appears to be only an extreme form of it, with the orange
band on the secondaries obsolete. Mr. Schaus’s types of
both insects were from Peru.
SISEME PEDIAS, sp.n. (Plate XXI, fig. 1, 2.)
¢. Alis nigro-fuscis, fascia communi in cellula anticarum angu-
lum analem posticarum versus (sed non attingente) extensa alba,
altera ultra eam cerulea ; posticis plaga aurantia ad angulum ana-
lem ; subtus fere ut supra, sed colore cieruleo, praesertim in anticis
multo latiore et ad basin regionis costalis extenso.
Hab, COLOMBIA (Wheeler).
This species differs from all others known to me be-
longing to the genus in having the outer blue band
crossing both wings. It is somewhat faintly shown on the
upper- -side, but beneath, especially on the primaries, it is
very brilliant, and occupies the space beyond the white
band, extending nearly to the outer margin.
some Central and South American Erycinide. 535
RiopinA (?) THEODORA, sp. n. (Plate X XI, figs. 2, 7; 3, 2.)
6 Allis fuseis, anticis macula (interdum duabus) in cellula et fascia
transversa subapicali albidis aut flavescentibus, duabus infra cellule
finem, aliis duabus subquadratis in margine externo ochraceis ; posti-
cis fascia interrupta a costa prope apicem ad marginem internum
extendente, maculis duabus prope marginem externum, echraceis ;
subtus fere ut supra, sed maculis anticarum majoribus ; posticis fascia
transversa pallidiore et magis distineta, maculis marginalibus absen-
tibus ; ciliis anticarum fuscis et albis ; capite et collo ferrugineis.
2. Mari similis, sed maculis duabus infra cellulam anticarum et
fascia transversa posticarum obsoletis ; subtus posticis immaculatis.
Hab. Brazit, Chapada (H. H. Smith).
This and the following species probably do not properly
belong to the genus Riodina, but they can be included in
it for the present ; both have much longer palpi than the
type of Riodina—R. lysippus (Linn.). 2. luctus, Berg, and
L. lysistratus, Burm., too, may have to be removed from it.
Three specimens.
RIoDINA (?) ALBOFASCIATA, sp. n. (Plate XXI, fig. 4, f.)
¢d. Alis fuseis, anticis macula ad cellule finem, altera majore trans-
versa apicem Versus, tertia (interdum quarta) infra cellule finem,
albidis ; posticis fascia venis interrupta, a costa prope apicem ad
cellule finem extendente, alba ; subtus fere ut supra, sed fascia com-
muni venis divisa a costa anticarum ad marginem internum postica-
rum extendente alba ; collo aurantio ; ciliis anticarum albis.
9. Mari similis, sed posticis fascia alba obsoleta.
Hab, ARGENTINE ReEpPuBLIC, Corrientes (Perrens), Cor-
doba (Berg, in Mus. Brit.); PaARaGuay (Coll. Crowley, in
Mus. Brit.).
I have long had numerous males of this peculiar species,
but have hitherto been unable to identify them. There
are specimens of it from Cordoba and also from Paraguay
in the British Museum; the former have the common
white band beneath broader than it is in our examples.
L. albofasciaia is undoubtedly a close ally of L. theodora,
but is much smaller and differently marked.
ANTEROS CARUS, sp.n. (Plate XXJ, fig. 5, ¢.)
g. Allis fusco-nigris, anticis ad basin et posticis (nisi apicem versus)
pilis ceruleis indutis, anticis maculis duabus, una ad cellule finem,
536 Mr. F. Du Cane Godman on
altera margini exteriori propiore, albo-hyalinis ; posticis punctis
apicem versus fere obsoletis coloris ejusdem; subtus leete castaneis,
anticis flavo, albo et fusco maculatis, posticis disco griseo irroratis,
basin versus et ad angulum analem flavo maculatis, fascia communi
angusta submarginali strigis et maculis variis interioribus metallicis ;
ciliis alterne flavis et fuscis ; cruribus pallide castaneis.
Hab, COLOMBIA, Bogota (Wheeler); BotitviaA, Tanam-
paya (Garlepp).
Dr. Staudinger sent me some years ago two specimens
of this species from Bolivia under the “MS. name of 4.
carus, and I have two others from Colombia in my collec-
tion, all males. The Bolivian insects differ from the others
in having a minute additional hyaline longitudinal spot
on the primaries beyond the cell, and the castaneous
colour on the under-side perhaps less extended. 4. carus
is a close ally of A. cavausius, Westw., with which it is
confused in the Hewitson collection, but the markings of
the under-side are very different, as mall be seen Sa the
figure here given. A. principalis, Hoptt., from Peru, 1s
unknown to me, but the description does not agree with
the present insect.
KMESIS EURYDICE, sp.n. (Plate XXI, figs. 6, $; 7, 2.)
¢. Alis glauco-cinereis, dimidio basali lineolis transversis nigris
irregulariter notatis, fasciis duabus exterioribus (presertim in anticis)
fere obsoletis ; subtus lete rufo-ferrugineis, lineolis nigris sicut in
pagina superiore, marginibus externis late nigro-fuscis.
2. Mari similis, sed majore et pallidiore, lineolis magis undulatis,
anticis plaga magna prope apicem, altera parva ad apicem ipsum,
albis ; subtus dimidio basali anticarum et posticarum ochraceo, aliter
fere ut supra.
Hab, ECUADOR, Sarayacu (buckley).
This is a form of #. fastidiosa, Mén., but the male differs
in having the ground colour of the upper surface uniformly
glaucous and both wings broadly bordered with fuscous
beneath. The female is very like that of the above-men-
tioned species, but the ground colour is ochreous beneath.
The broad dark border of the wings in the male beneath
separates it from both 2. godarti and £. aurimna, Boisd.
In the Hewitson collection #. ewrydice is confused with
EL. fastidiosa.
some Central and South American Erycinide. 537
EMESIS NEEMIAS. (Plate XXI, figs. 8, 7; 9, 9.)
Emesis neemias, Hew., Exot. Butt. v. Emesis, ff. 3, 4 (f).
d2. Alis eee tce lineolis variis transversis nigris (exterioribus
angulatis), viridi-ceruleo limbatis, punctis in serie communi sub-
marginali nigris ; subtus ferrugineis, maculis irregularibus elongatis
nigris,
2. Supra pallide fuscis, aliter mari similis, subtus albescentibus
lineis transversis paginz superioris rufescentibus et in costa, apicem
versus, squamis paucis metallacis notatis.
Hab. Boxtvia, Yungas (Garlepp); Brazir, Chapada
(H. H. Smith); Paracuay (Coll. Crowley, in Mus. Brit.).
Hewitson’s figure of this insect was taken from a worn
male from Brazil. I have now seen several good specimens
of both sexes, and take this opportunity of re-describing
the species. The metallic spots on the upper-side are
very similar to those of Symmachia fatima (Cram.). My
collection contains three males, two of which were received
from Dr. Staudinger.
SYMMACHIA PROGNE, sp.n. (Plate XXI, fig. 10, 7.)
g. Alis fuscis ferrugineo tinctis, undique nigro irregulariter lineo-
latis, anticis ad marginem externum punctis nigris in serie notatis,
fascia communi ultra cellulam anticarum feread marginem internum
posticarum extendente ochracea vix ceruleo lavata; subtus rufo
ferrugineis, lineis nigris sicut supra, margine interno fusco.
Hab. Bowtvis, Yungas (Garlepp); E. Perv, Pozzo
(Pearce).
This species is about the same size as S. emesina, Staud.,
from the Upper Amazons, but may at once be distinguished
by the absence of the ferruginous apical patch on the
primaries and the ochraceous band crossing both wings,
which in certain lights is shot with blue. I have also
seen a specimen of S. progne, from Chanchamayo, in Dr.
Staudinger’s collection. It is very like S. temesa, Hew.,
from Ecuador, but is differently coloured above and
beneath. Four males.
SYMMACHIA HIPPODICE, sp. n. (Plate XXI, fig. 11, 2.)
9. Alis aurantiis, anticis costa et margine externo (ad apicem
macula transversa excepta) late fuscis, margine interiore valde den-
tata ; posticis margine externo late (linea angustissima submarginali
excepta) fuscis, angulum analem versus maculis tribus obscuris
538 Mr. F. Du Cane Godman on
notatis ; subtus ut supra, sed colore fusco minus extenso et in
maculas fracto.
fab. BRAziL, Chapada (Hl. H. Sinith).
The above description is taken from a single female
specimen. It seems to be most nearly allied to S. hetwrina,
Hew., from the Lower Amazons, of which also the female
alone is known.
CARIA SMARAGDINA, sp. pn. (Plate XXI, fig. 12, ¢.)
¢. Alis fuscis, lineis submarginalibus nigris, anticis linea trans-
versa prope basin quoque nigra, anticis et posticis (marginibus ex-
ternis et in anticis fascia lata basin versus exceptis) squamis viridibus
nitentibus dense vestitis, linea communi angusta submarginali metal-
lica; subtus griseo-fuscis, lineis variis undulatis nigris transeuntibus,
anticis in costa tota maculis metallicis notatis; capite.et corpore
supra squamis viridibus indutis.
Hab. Bouivita, Coroico, 6,500 ft. (Garlepp).
Dr. Staudinger has sent me an example of this species
as a small variety of C. amazonica, Bates, but it differs
from that insect in having the upper surface almost
entirely clothed with green scales, and both wings have
a submarginal metallic line. C. smaragdina is also very
much smaller, and is really more nearly allied to C.
chrysame, Hewitson (?=C. psittacus, Hopff.). C. trochilus
occurs in the same locality, but is a much larger insect
and the green scales are more restricted.
CARIA MARSYAS, sp. n. (Plate XXI, figs. 13, f; 14, 2.)
¢. Alis fuscis, lineis angustis interruptis transversis nigris, anticis
ad basin, in area discali ad costam lata, et angulum analem versus
anguste, squamis viridi-aureis vestitis ; posticis squamis in dimidio
basali ad anguluin analem extendentibus coloris ejusdem, linea
angustissima communi submarginali metallica, punctisque nigris,
fulvo plus minusve circumcinctis, ultra eam; subtus pallide fuscis,
lineis nigrescentibus irregularibus transversis, anticis litura elongata
cellulam occupante coccinea, et maculis tribus aut quatuor notatis.
?. Mari similis sed pallidiore et minus viridi squamata ; subtus
lineis transversis in maculas fractis.
Hab. Brazit, Chapada (1. H. Smith); Paracuay,
Corrientes (Perrens).
This alone of the group of species including C. argiope,
Godt., C. amazonica, Bates, and C. lampeto, G. and §., has
=
ros) ert
some Central and South American Erycinide. 539
the red patch in the cell of the primaries beneath broken
by several transverse metallic spots, a character common to
the females of most of the allied species. The submarginal
metallic line is very narrow. The series before me con-
sists of ten males and one female. I also have three male
specimens from Corumba, in Brazil, which perhaps repre-
sent another form; in these the red spot on the primaries
beneath is dentate on the upper edge.
CHARIS ARCUATA, sp.n. (Plate XXII, figs. 1, $;-2, 2.)
4
¢. Alis nigro-fuscis, fascia communi arcuata a costa anticarum ad
marginem internum posticarum extendente rufo-aurantia, intus linea
angusta argentea limbata, linea altera argentea submarginali; subtus
ut supra sed fascia multo latiore, linea argentea absente sed macula
in margine interno posita.
@. Mari similis sed fascia latiore et in anticis magis arcuata ;
subtus sicut supra.
Hal, CoLoMBIA, Bogota.
This is a northern form of (. cwcias, Hew., differing from
it in the more arcuate fascia, which is specially noticeable
in the female. I have one pair only.
CHARIS DUKINFIELDIA. (Plate XXII, fig. 3.)
Charis dukinfieldia, Schaus, Proc. U. 8. Nat. Mus. xxiv,
p- 401 (1902).
A figure of this species is given from the specimen from
Parana, presented by Mr. Schaus to the British Museum.
MONETHE MOLIONE, sp.n. (Plate XXII, fig. 4, f.)
¢d. Alis ochraceis, anticis costa, apice late, et margine externo
nigris, macula elongata transversa angulum apicalem versus quoque
ochracea ; posticis margine externo nigro; subtus ut supra, sed
posticis costa, linea angusta breve basali et altera margini interno
subparalleli, nigris ; corpore supra ochraceo, nigro lineato ; pedibus
nigris,
Hab. KE, Peru, Pebas (Hahnel, ex Staudinger).
A very fine species, differing from Jf. albertus, Feld.,
and its allies in the large extent of the ochreous colour on
both wings. The antennz are Jong and very gradually
thickened towards the apex, so that there is no distinct
club. One specimen.
540 Mr. F. Du Cane Godman oz
CRICOSOMA ASCLEPIA. (Plate XXII, fig. 5, f.)
Symmachia asclepia, Hew., Equat. Lep. p. 51; Exot. Butt. v.
Erycinide, f. 9.
¢. Alis lewte aurantiis, costa anticarum et marginibus externis
maculis variis nigrescentibus, iis in costa transversis et in cellulam
extendentibus ; subtus fere sicut supra sed costa anticarum et
marginibus externis pallidioribus, maculis nigrescentibus ad_ basin
extendentibus.
Hab, Ecuapor, Curarai; E. PERU, Sarayacu (Buckley).
Hewitson’s type of this insect is in a very worn con-
dition, and it is scarcely re-cognizable from his figure. I
therefore take the opportunity of re-describing the species
from a better specimen in my own collection. CC. asclepia,
Hew., is a close ally of C. zypete, Hew.; the type of the
former is from Curarai.
CRICOSOMA IRRORATUM, sp.n. (Plate XXII, fig. 6, f.)
¢. Alis pallide fuscis, squamis sparse et maculis parvis obscuris
dense vestitis, iis ad margines externos flavo circumcinctis ; subtus
flavis, maculis obscuris sicut in pagina superiore.
Hab. BritisH GUIANA, Carimang River ( Whitely).
In general colour this species somewhat resembles C.
calligraphum, Hew., but it has less pointed wings, and
it is possible that the insect is not rightly placed in the
genus Cricosoma.
MESENE IASIS, sp. n. (Plate XXII, fig. 7, 3.)
¢. Alis nigris cyaneo tinetis, fascia lata infra et ultra cellulam
anticarum fere ad medium marginis interni posticarum extensa
coccinea ; subtus nigris leete cyaneo lavatis, anticis macula elongata
in margine interno ochracea.
Hab. EK. Peru, Sarayacu (Buckley).
A single male, closely resembling J/. sagaris, Cram., but
larger and blacker, and showing a brilliant blue lustre
both above and beneath. ‘The common band on the
upper-side is red, instead of being orange. The body
appears to want the ochreous band so conspicuous 1n
the male of Jf. sagaris.
MESENE EANES, sp.n. (Plate XXII, fig. 8, t.)
¢. Alis nigris, anticis litura magna subtriangulari basali a cellula
ad marginem internum extendente, posticis altera discali oblonga,
coceineis ; subtus pallide fuscis basin versus ochrascentibus, maculis
some Central and South American Erycinide. 541
parvis obscuris et griseis dense notatis ; corpore supra nigro, coccineo
late cincto.
Hab, Lower AMAZONS, Santarem (H. H. Smith).
Closely allied to JZ. debilis, Bates, from the Tapajos,
but differently shaped, and with the patches on the upper-
side of both wings red. One specimen.
LASAIA MERITA, sp. n. (Plate XXII, fig. 9, 3.)
dg. Alis glauco-viridibus, anticis lineis angustis transversis in
area costali, fasciis duabus submarginalibus exterioribus ad costam
latioribus, et margine externo anguste, nigris; posticis punctis in
serie submarginali et margine exteriore anguste quoque nigris ;
subtus griseis, maculis marginem externum versus sicut supra,
anticis dimidio basali pallide fuscis lineis transversis obscure notatis,
posticis dimidio basali lineis obscuris reticulatis; ciliis nigris et
albis.
Hab. Boutvia, Coroico, 6,500 ft. (Garlepp), Chairo
(Buckley).
This is a form of the very variable and widely distributed
LL. meris, Cram., but the wings are less spotted, both above
and beneath, the basal half of the under-side is more
reticulate, and the distal portion (the apical angle of the
primaries excepted) is principally grey. Three specimens,
two of which were sent me under the MS. name of Z.
merita by Dr. Staudinger.
LASAIA OILEUS, sp. n. (Plate XXII, figs. 10, f; 11, 9.)
¢. Alis pallide fuscis, punctis obscuris dense notatis, marginibus
externis (presertim in posticis) argenteo-czeruleo irregulariter notatis,
anticis punctis duobus in costam ultra cellulam albis; subtus
fuscis, griseo et albo dense variegatis ; ciliis alterne fuscis et albis ;
antennis albo annulatis.
?. Mari similis, sed brunnescentiore, magis maculato et colore
metallico fere obsoleto ; subtus alis brunneis albo variegatis.
Hab. Paracuay (Perrens).
The above description is taken from two males and one
female from Paraguay, and I have another male labelled
Cayenne, but this locality is probably incorrect. L. oilews
seems to agree sufficiently well with Z. mxris, Cram., to
be included in the same genus, though it is much smaller
than any of the other described species. It is also some-
what like Calydna euthria, Doubl. (=C. micra, Bates),
from the Amazons.
542 Mr. F. Du Cane Godman on
LEMONIAS ANNULIFERA, sp.n. (Plate XXII, fig. 12, f.)
¢. Alis fuscis, anticis annulis tribus in cellula et duabus infra eas
ceruleo-albis ; posticis maculis obscurioribus tribus, una in costa,
duabus in cellula, margine externo late albo; subtus ut supra sed
pallidioribus, anticis annulis quinque et posticis annulis sex rufo-
brunneis ceruleo circumcinctis.
Hab, BRITISH GUIANA, Quonga (MW hitely).
This species is a close ally of L. /ewcocyana, Hiibn., but
may easily be distinguished by the annulated base of the
wings.
LEMONIAS MHON, sp.n. (Plate XXII, fig. 13, ¢.)
¢. Alis rufo-fulvis, lineis brevibus dispersis transversis, punctis
marginalibus in serie fere obsoleta, nigris, anticis (nisi margine
interno) brunneo lavato, fascia indistincta submarginali obscuriore ;
subtus cinereis, marginibus externis late obscurioribus, maculis
transversis nigris sicut in pagina superiore, maculis indistinctis in
serie submarginali cinereis, in posticis nigro pupillatis; ciliis anti-
carum albo maculatis; corpore supra rufo-fulvo.
Hab. W. Cotomais, San Pablo, Rio San Juan (ex
Staudinger ).
A single example. Nearly allied to Z. galena, Bates,
from the Amazons, but differing in having the fore-wings,
except along the inner margin, much darker ; the narrow
whitish marginal line is absent. Beneath, the primaries
are cinereous, instead of ferruginous, and the black sub-
marginal spots on the hind-wings are less conspicuous and
farther from the margin.
L. idmon, 8. and G., from Chiriqui, is also very similar,
but has the ground colour of the fore-wings of a uniform
tint, as in LZ. galena.
LEMONIAS MONOIDES, sp.n. (Plate XXII, fig. 14, ¢.)
¢. L mexoni similis, sed maculis discalibus posticarum rotundatis
et punctis submarginalibus magis obviis ; subtus pallidioribus.
Hab. British GUIANA, Bergen-Daal (£//acombe).
A single specimen, differing from ZL. maxon as above
described.
LEMONIAS ELPINICE, sp. n. (Plate XXIII, fig. 1, 2.)
¢. Alis fuscis, anticis lineis angustissimis transversis cerules-
centibus, posticis dimidio distali aurantiis; subtus anticis palli-
some Central and South American Erycinide. 543
dioribus, dimidio basali lineis irregularibus transversis czerules-
centibus, dimidio distali colore eodem variegato, margine externo
punctis nigrescentibus ; posticis albis, costa late czerulescente,
lineolis et punctis fuscis notatis ; ciliis aurantiis,
Hab, ConomBia, Fusagusuga (Chapman).
On the upper-side the secondaries of this species are
broadly marked with orange as in LZ. florus, Staud., also
from Colombia; but on the under-side the general colour
resembles that of ZL. lewcocyana, Hiibn., L. violacea, Butl.,
and others. One specimen.
LEMONIAS MARTIA, sp.n. (Plate XXIII, fig. 2, f.)
¢. Alis anticis ceeruleo-violaceis, ad basin et in margine interno
brunneo-ferrugineis, dimidio basali nigro maculato, costa, margine
externo et fascia interrupta submarginali quoque nigris; postic's
brunneo-ferrugineis, dimidio basali lineolis transversis et margine
externo nigris, fascia lata curvata ab angulum analem fere ad apicem
extendente rufo-aurantia ; subtus pallide griseo-ceruleis, marginibus
externis late fusco suffusis, bitriente basali lineolis transversis nigris,
anticis maculis in serie submarginali (iis ad apicem et ad angulum
analem magis distinctis) nigrescentibus, posticis maculis elongatis
submarginalibus nigris czeruleo circumcinetis ; ciliis anticarum nigris
et albis.
Hab. CotompiA, San Pablo, Rio San Juan (ex
Staudinger).
Very like Z. florus, Staud., but much larger, being of
the same size as ZL. staudingerit. The hind-wings have
the orange band of a richer colour and narrower, the
ferruginous marking on the primaries extends well into
the cell, and the submarginal band of these wings is
broader towards the apex. It is somewhat remarkable
that these three insects all inhabit the same very humid
district on the Pacific slope. One specimen.
LEMONIAS STAUDINGERI, sp. 0.
Lemonias sudias, Staud., Exot. Schmett, p. 259, t. 92
(2) (nec Hew.).
¢. Alis ceruleo-violaceis, anticis maculis transversis in dimidio
basali, costa, apice et fascia submarginali, posticis area costali maculis
quinque transeuntibus, nigris ; subtus pallide fuscis, dimidio basali
nigro maculatis, posticis griseo lavatis, fascia irregulari submarginali
obseuriore, maculis elongatis ultra eam nigris,
544, Mr. F. Du Cane Godman on
Hab. Cotompia, San Pablo, Rio San Juan (ex
Staudinger).
The insect figured by Dr. Staudinger as LZ. sudias, Hew.,
of which he sent me an example, was wrongly identified
and requires a new name. Hewitson’s species is common
in the low country of Central America, extending from
southern Mexico to the republic of Honduras, the male
being very different from that of the Colombian insect.
LI. staudingeri 1s considerably larger than ZL. sudias, the
ground colour of the upper surface of the male is entirely
violaceous, and the orange border of the secondaries is
absent.
LEMONIAS PIRENE, sp.n. (Plate XXIII, fig. 3, 2%)
¢. Alis anticis nigro-fuscis, litura transversa magna ultra cellu-
lam et linea submarginali cxruleis, area marginis interni late
aurantia, maculis transversis nigris notata ; posticis aurantiis nigro
limbatis, ad apicem (et interdum in area discali) nigro punctatis ;
subtus griseo-ceruleis nigro maculatis, punctis elongatis in serie
submarginali griseo circumcinctis ; ciliis nigris et albis.
Hab. Ecuavor, Rio Napo (Whitely).
This is a form of LZ. rhesa, Hew., from the Amazons, but
differs from it in the less extension of the orange colour
of the primaries and the much larger blue patch beyond
the cell. On the upper-side it is not unhke Charis
amalfreda, Staud., from Pebas, which, however, is a much
smaller insect. Two specimens.
LEMONIAS NOMIA, sp. n.
Lemonias thara, Hew., var., Exot. Butt. v. Lemonias, t. 5,
ff. 41, 42 (ft, 9).
Hab. Guiana, Aunai, Essequibo River ( Whitely), Cay-
enne (mus. G. & S.).
The Guiana form of the Amazonian JZ. thara, Hew.
(= L. melia, Bates), requires a separate name. Hewitson
described it as a variety, but a comparison of his figures
of the male of each shows that they should be treated as
distinct. Z. nomia may be readily distinguished from
L. thara by the fulvous colour of the upper- -side, and also
by having an additional band of elongate black spots
crossing the wings beyond the cell.
some Central and South American Erycinide. 545
LEMONIAS FANNIA, sp. n. (Plate XXIII, fig. 4, ¢.)
¢. Alis nigro-fuscis, anticis ad angulum analem anguste ceruleis,
posticis dimidio distali pallide caruleis maculas duo nigro-fuscas ad
apicem includente ; subtus pallidioribus, lineis obseurioribus, trans-
versis communibus ad basin frequenter interruptis ; posticis
maculis obscuris in serie submarginali ad apicem et ad angulum
analem majoribus.
Hab. British Gutana, Aunai, Essequibo River
(Whitely).
A single rather worn specimen is all I have seen of this
species. It appears to be very distinct from any known
member of the genus, and is perhaps most nearly allied to
L florus, Staud., and L, lasthenes, Hew.
LEMONIAS (?) AXENUS. (Plate XXIII, fig. 5, 9.)
Lemonias axenus, Hew., Exot. Butt. v. Lemonias, ete.,
fT:
The male only of this species was described and figured
by Hewitson, and no locality was given. I have two
males of it from Chapada, Matto Grosso, and two females
from Estancia La Gama, Venado Tuerto, in the province
of Santa Fé, La Plata; these latter, one of which is
here figured, differ in having all the markings of the upper-
side straw colour. It probably does not belong to the
genus Lemonias. Beotis bifasciata, Mengel, from Paraguay,*
is no doubt a nearly allied form, but has fewer spots on
the outer margin.
APODEMIA GLAPHYRA.
Anatole glaphyra, Doubl. and Hew., Gen. Diurn. Lep. ii,
taniplente 3
Anatole modesta, Mengel, Ent. News, 1902, p. 176, t. 8,
ied 4,
There does not appear to be any reason for separating
the Paraguayan insect described and figured by Mr.
Mengel from A. glaphyra, Doubl. and Hew. Possibly the
American author was not acquainted with the figure of
the latter.
* “Ent, News,” 1902, p. 177, t. 8, f. 4.
546 Mr, F. Du Cane Godman on
APODEMIA MULTIPLAGA. (Plate XXIII, fig. 6.)
Apodemia multiplaga, Schaus, Proc. U.S. Nat. Mus. xxiv,
p- 404 (1902).
The specimen figured is from Rinconada, Mexico, given
by Mr. Schaus to the British Museum.
NYMPHIDIUM GRANDE, sp.n. (Plate XXIII, fig. 7, 9.)
®. Alis fuscis, fascia latissima a cellule finem anticarum ad
medium marginis interni posticarum extendente albida, linea
communi ferruginea submarginali exteriore, anticis macula elongata
costali venis divisa apicem versus, aliisque minoribus plus minusve,
lunulatis in serie angulum analem posticarum extendentibus, serie
altera margini exteriori propiore (in anticis fere obsoleta), albidis ;
subtus pallidioribus maculis omnibus albidis majoribus ; ciliis posti-
‘carum albis et nigris.
Hab, CoLomBta (Wheeler).
This fine species is even larger than WV. nycteus, G. and
S., from Chiriqui, but in colour it approaches the female
of N. imycone, Hew. It is probable that the male is
without the broad median band. <A single specimen.
ARICORIS PLAGIARIA. (Plate XXIII, fig. 8, 2.)
Aricoris plagiaria, Grose-Smith, Rhop. Exot. in,
Erycinide, p. 4, t. 1, ff 12, 13 (¢@).
9. Alis nigro-fuscis, anticis fascia a costee medio fere ad angulum
analem extensa sordide alba, anticis ad basin late posticisque (nisi
in area costali et margine externo) cxeruleo lavatis, posticis ad mar-
ginem externum venis, et lineis inter venas, obscuris notatis ; subtus
pallide fuscis, anticis fascia sicut supra, area cellulari indistincte
nigro maculata, posticis grisescentioribus maculis variis nigris, ad
margines elongatis, anticis ad basin et posticis in area _basali
ochraceo notatis.
Hab. E. Peru, Sarayacu (Buckley, ¢, 2); AMAZONS,
Santa Rosa (Stwart, @).
Mr. Grose-Smith described this species from two males
supposed to be from Buckley’s collection. We have both
sexes from Sarayacu, and the female, which we now take
the opportunity of figuring, approaches the same sex of A.
serica, Westw., but is considerably larger, the fascia, too, on
the primaries is whitish and more oblique, and the under-
surface 1s spotted.
some Central and South American Erycinide. 547
ARICORIS HIPPOCRATE, sp.n. (Plate XXIII, fig. 9, 9.)
?. Alis nigrescentibus, anticis fascia obliqua a costee medio fere
ad angulum analem extensa sordide alba; subtus sicut supra, sed
posticis margine externo angulum analem versus obsolete albis ;
ciliis posticarum albo-notatis,
Hab, Ecuapor, Rio Napo (Whitely).
The above description is taken from a single female much
resembling that sex of A. dagus, Cram.; but it is blacker, the
band on the primaries is whitish, and the secondaries are
without definite white spots on the outer margin beneath ;
the cilia of the hind-wings, too, are spotted with white.
A. hippocrate is not unlike the temale of A. phedon, but
the band of the fore-wings is whitish instead of orange,
and the secondaries are differently marked on the under-
side.
ARICORIS PHEDON, sp. n.
d. Alis nigrescentibus leete purpureo tinctis; subtus nitente
fuscis, inter venas ad basin et ad margines externos pallide ochra-
ceis,
Q?. Alis nigrescentibus, anticis fascia lata obliqua apicem versus
ochracea ; subtus fere ut supra sed posticis ad marginem externum
albescentibus.
Hab, Ecuapor, Rio Napo (Whately).
A close ally of A. velutina, Butl., from the Upper
Amazons, but smaller, the male with less elongate wings,
and the lustre of the upper-side is purple instead of blue,
as in A. velutina, and extends over the whole surface.
Five specimens.
ARICORIS MAIA, sp. n. (Plate XXIII, fig. 10, f.)
dg. Alis nigro-fuscis, anticis squamis sparsis czeruleis inter venas
maculas formantibus, ea angulum analem versus elongata ; posticis
(nisi area costali et venis omnibus) ceruleis ; subtus fuscis, ad angu-
lum analem anticarum et posticarum czeruleo indistincte maculatis.
Hab. W. CotomptaA, San Pablo, Rio San Juan (ex
Staudinger).
Very near Arioris alemevon, Hew., from Ecuador, but
differs in having the blue scales on the primaries forming
scattered elongate spots towards the outer, instead of
occupying a broad space along the inner, margin. Three
specimens,
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.) 37
548 Mr. F. Du Cane Godman on
ARICORIS HELICE, sp. 0.
Aricoris crucntata, Staud., Exot. Schmett., p. 264, t. 93
( nec 3.)
The insect figured by Staudinger as the female of 4.
cruentata, Butl., was wrongly identified, and a new name
is required for it. I have both sexes of A. erwentata,
from Ega, where the types of the latter were obtained.
Of A. helice I received two females from the Amazons,
from Staudinger. It appears to be allied to A. helvodora,
Staud.
ARICORIS (?) TERIAS, sp. n. (Plate XXIII, fig. 11, 2.)
9. Alis ochraceis, anticis apice et margine externo (intus valde
sinuato, maculam elongatam includente) fuscis, posticis fusco
limbatis ; subtus anticis ochraceis costa et margine externo fusces-
centibus et albo lineatis, posticis fuscis albo striatis.
Hab, PARAGUAY.
Of this extraordinary insect, which is very like a Verias
on the upper-side, I possess a single female only. It 1s
unfortunately imperfect, the legs and part of the antenne
being wanting, and its generic position in the absence of
the male is doubtful. On the under-side it somewhat
resembles the female of A. cepha, Feld., except that it
is much more strongly streaked with white.
some Central and South American Erycinide.
EXPLANATION OF PLATES XX—XXIIL.
Hie.
Fic.
dF
TID OT Bg bo
PLATE XX.
Mesosemia phace, &.
. thera, g.
3 orbona, 3.
ae om
Cremna malis, g.
Hyphilaria orsedice, 2.
Hurygona lieinia, 9.
5 authe, 3.
‘5 rhodogyne, 2.
” ” 2 O
3 tarinta, 3g.
3 fabia, 3.
Mesenopsis pulchella, 2.
Chamelimnas pansa, g.
n joviana, ¢.
PATE, DOG
Siseme pedias, 2.
Riodina (?) theodora, ¢@.
” ” 2.
» (2) albofasciata, 3.
Anteros carus, ¢.
Einesis eurydice, 3.
” ” g 0
» neenius, go.
” z) 2.
Symmachia progie, &.
i hippodice, @ .
Caria smaragdina, ¢.
» marsyas, 3.
a]
” ” 2 3
549
leis Te
PuatTe XXII.
Charis arcuatu, 2.
) rp) e ©
» dulanfieldia.
Monethe molione, 2.
Cricosoma asclepia, ¢.
- irroratimn, & .
Mesene iasis, g.
» «eanes, g.
Laswia merita, ¢.
5, ouleus, 3.
$3 ” g q
Lemonias annulifera, g.
5 mon, 3b.
- meonoides, ¢.
PLATE XXIII.
Lemonius elpinice, 4.
” martia, &.
x pirene, ¢.
Ba fannia, 6.
+ axenus, 2.
Apodemia multiplaya.
Nymphidiwm grande, @.
Aricoris plagiaria, ?.
5, hippocrate, &.
» maida, do.
» (©) teras, ¢.
( 551 )
XXV. Supplementary Note to a Paper entitled Hymenoptera
aculeata, collected by the Rev. ALFRED E. EATON,
M.A., in Madeira and Tenerife, in the spring of
1902. By EpwarD SAUNDERS, F.R.S., F.L.S.
[Read November 18th, 1903. ]
I REGRET to say that when I wrote the above paper I
entirely overlooked an article by Professor Perez of
Bordeaux, published in the Annales de la Société Ent.
de France for 1894, in which he describes the new species
of Hymenoptera taken in the “ Voyage de M. Ch. Alluaud
aux iles Canaries.’—In this article I find the species
which I have described under the name of Podalirius
canariensis described as Anthophora alluaudi, to which of
course canariensis must sink as a synonym.
It is curious that Professor Perez and myself should have
made almost identical remarks about the black variety of
Bombus terrestris with the white tail, both suggesting
that it probably represents what Brullé quotes as 2.
sovoensis. I can only apologize to Professor Perez for
unwittingly appropriating his observation.
It may be worth mentioning that Al/waudi, Per., is
omitted by Friese in his monograph of the Genus
Podalirius 1897.
TRANS, ENT. SOC. LOND. 1903.—PART IV. (DEC.)
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XXVI. Protective Coloration in its relation to Mimiery,
Common Warning Colours, and Sexual Selection.
By Apssorr H. THAYER. Communicated by
Pror. Epwarp B. Poutton, M.A., D.Sc., F.R.S,
[Read October 21st, 1903.]
THE following paper records an artist’s examination of the
principles of butterflies’ coloration, and shows how the
results tend to restrict the fields heretofore claimed for
Mimicry and Common Warning Colours, and to place
them on a basis of Concealing Coloration. It contains
also several arguments tending to restrict the hypothesis
of Sexual Selection.
It does not attack the obvious fact that every possible
form of advantageous adaptation must somewhere exist.
It is obvious to its writer that there must be unpalata-
bility accompanied by Warning Coloration,—as apparently
in the cases of the Hornbills and Wood Hoopoes reported
by Mr. Frank Finn, and probably in many Corvidee, for
instance,—and equally plain that there must be Mimicry,
both Batesian and Miillerian. Yet every case demands
special examination, for the reasons that I shall show
herein; and no apparent conspicuousness of coloration
is sure to prove such when examined on the principles
established in this article.
First, it seems necessary to establish the artist’s claim
to be the judge of all matters of visibility, and the effect,
upon the mind, of all patterns, designs, and colours. If
even the artist is limited in this, his own field, what
hope is there for others? Fullest wisdom on the part of
naturalists would make them adjourn all matters of
animals’ appearance to us artists, just as any wise ruler
gathers about him the most highly specialized minds, to
widen, through them, his own scope.
An artist reads design wherever it occurs, just as a
composer reads a score, without playing it, or hearing it.
He perceives that every juxtaposition of spots, or shapes,
or colours, or of dark and light, and of degrees of these,
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.)
554 Mr. A. H. Thayer on
is just so much representation of some structure, whether
the representation be accidental or intentional. He sees
at a glance in marble-veins, the grain of wood, etc., not
imaginary, but actual representations of natural objects
and perspectives, and weighs the correctness of these.
Nature has evolved actual Art on the bodies of animals,
and only an artist can read it. When he examines the
colour and colour-pattern of the animal kingdom, he sees
that zoologists are hopelessly off the track in Y their general
conception as to which coloration is to be called con-
spicuous, 7.c. rendering its wearer so. Any coloration
or pattern would be conspicuous somewhere, and Nature
cannot prevent animals from straying beyond the environ-
ments that would most perfectly harmonize with their
colour and pattern. But let us take the broadest possible
survey, and we cannot doubt that most animals wear on
their coats pictures of their habitat. As I before pointed
out, even the under-sides of the wings and tails of hawks bear
the general twig-patterns so common on forest birds, as if
Nature found it worth while to efface the white silhouette
their wings’ under-sides would make when they extended
them while perching. We see how completely such
patterns (when couched, of course, as they always are, in
the effacive gradation) ilo help to obliterate a partridge,
grouse, woodcock, hare, or any other of almost all the
species in every order; since they prove to be actual
animated pictures of hen environment. As I said before,
in my paper on so-called “ Banner-marks,” * these forest-
like patterns are found on forest creatures, and not on
desert creatures, or ocean creatures. Sand-birds are usually
marked in longitudinal, delicate patterns, very like those
the sand assumes when seen at the same angle at which
one observes the birds themselves. Tigers and zebras are
resolved into pictures of tall, strong ‘flags, grasses, and
bamboos, while the hon is a picture of The desert. (It
will some day be plainly understood that the effacive
gradation is the essence of the success of these pat-
terns. Were they not arranged to compose one perfect
counter-gradation, from top-dark to under-white, they would
appear merely as what artists call “lines of quantity,’
like the hoops of a barrel, emphasizing the rotundity, not
effacing it.)
Now, let me prove that aay pattern would somewhere be
* ©The Auk,’ vol. xvii, 1900, p. 108,
Protective Coloration in its relation to Mimicry, ete. 555
conspicuous. I once saw a skunk (Mephitis americanus)
crossing a snow-field near at hand. This animal is_black
(with the slight amount of effacive gradation found even
in black animals), with a large white pattern on top.
He was totally unrecognizable, because his white against
the snow was undistinguishable. His black was left to
form a most grotesque silhouette. Had he been against
black, it would have been this black part that disap-
peared, and one would have seen only an unrecognizable,
moving white thing. Naturalists’ lack of understanding
this principle’s immense import has gone far to strengthen
the present Mimicry and Warning-Colour theories, which
may prove to have been evolved, largely, in the eftort
to explain supposed conspicuousness, where such did not
exist. A tiger in the desert sands, though his gradation
would still, more or less, efface his solidity, would never-
theless show his pattern. His bamboo-vistas would be
plainly a failure against the sand. The lion in the bamboos
would, when not covered by them, tend to present an unac-
countable flat silhewette,—a lion-shaped section of desert-
landscape, out of place. On the same principle, a white
patch on striped cloth or a striped patch on white cloth
would be conspicuous. We see on all hands evidence that
Nature cannot help moving forward to the utmost com-
pleteness of protective devices ;—that, in fact, she cannot
grope or blunder. A marvellous, turquoise, emerald-green
and red-coral-marked Mediterranean fish looks conspicuous
on the fishmonger’s slab; but follow him to the sun-lt
ocean grottos which he inhabits, and of which he is a
wonderful picture! No, the whole use of the word con-
spicuous 1s mainly born of the zoologist’s lacking the
artist’s sight.
Let us now turn to the field in which the naturalists
are most conspicuously at fault, that of the butterflies and
moths. One glance of an artist,—that is, of an artist
accustomed to lifelong looking at vegetation and butterfly-
life,—at a world’s collection of butterflies, shows him that
they are mainly either flying pictures of various com-
binations of flowers and their backgrounds, pictures of the
shadow under foliage, with delicate patterns of vegetation
or flowers drawn across it, as, for instance, in the North
American Papilo polydamas, and the dark Satyrine,—or
that they are wonderful representations of flowers them-
selves, as in the Pierinx (all but their usually narrow dark
556 Mr. A. H. Thayer on
border), many of which even bear a representation of six
stamens (counting their two antennz), and, what is very
common in butterflies, a wonderfully perfect shading on
that part of the wings next the body, grading toward it
in a way that makes it appear like the bottom of a con-
cavity. My photographs of Limenitis (Basilarchia) arthemis
show the flower-form, the appearance of the mm of its
cup being carried across the butterfly, as in the species of
Precis which wear a large, bright semicircular bar, cutting
them as the skunk’s white cuts him.
I should have placed at the beginning this axiom:
Only wnshiny, bright monochrome is intrinsically a revealing
coloration. As soon as patterns begin, obliteration of the
wearer begins, as shown in the case of the skunk. Nature
does not blunder, and Natural Selection would evolve the
monochrome, instead of a patterned surface, were simple
conspicuousness her aim. Also, she would, if she used
patterns mainly as badges for identification of the wearer,
fo)
have omitted the delicate subtilties that go to make up
Ss
the patterns of most butterflies. Let us apply the skunk-
lesson to the many dark butterflies which wear more or
less bright, clean-cut patterns. As they rest on flowers,
their dark matches very closely the shadew-depths between
the flowers, especially when seen from above or outside the
flower-mass; and, in fact, the delicate general gradation
and faint detail existing even in these parts, appear to
an artist to represent the near vistas under the flowers;
while the bright pattern is likely to echo the notes of the
flowers themselves. Only artists understand this colour-
echoing. The artist’s sight is conscious, as it ranges over
a scene, of every recurrence of each colour-note. This
colour-note, wherever seen, seeks, as it were, i/s own, in
lis brain,—just as a violin-string rings when its note is
sung. In a book we are writing on protective coloration,
my son and I shall show larve that resemble things
(already well known), larvee that disappear, larvae that
appear to be extensions of leaves; and larve with many
other startling and dissimilar concealment-schemes. What
wonder if in butterflies there prove to be as many different
forms of concealment? It is impossible to lay too much
stress on the fact that all patterns which look so striking
and bizarre, when off duty, are, when on duty, up to the
moment of detection, precisely the workers of the magical
illusion that conceals, It is inconceivable that birds should
Protective Coloration in its relation to Mimicry, ete. 557
more easily recognize minute patterns than colour, when
we realize that the perfect colour-adaptation of innumerable
forms of life, from mammals to larvee, proves that the lower
animals see colour (since otherwise such adaptation would
not be necessary for their concealment). In each form of
protective coloration there exist cases so pronounced as
to leave no doubt of their use. Each of these has been
assumed to be mimicked, or, at least, echoed, for some
reason, by other species than the one in which it is most
perfect. Let us look at the dead-leaf pattern, i.e. the
pattern that represents, in the most minute degree,
substance of the colowr and thickness of dead leaves, and
lying as near the ground as dead leaves usually he. This
pattern is marvellously perfect on the Copperhead snake
(Trigonocephalus contortrix), on some Boas, on that form
of domestic cat which has the most tiger-cat-like black
and grey pattern (as well as, in fact, on tiger-cats them-
selves), and on several Sphinx moths. Of course, when
this leaf-representation occurs on the rotundity of animals’
bodies, as in the cats or snakes, it exists only in
co-operation with the regular effacive gradation, but on
the flat plane of a Sphinx’s upper-wing-surface it has
and needs no such co-operation. In the Sphinx-moth
photograph which I have sent Professor Poulton, this
reproduction of thin material casting a shadow on the
surface it lies on is past all mistaking. This artifice is
present on many moths, and its elements are traceable in
such butterfly genera as Vanessa, Grapta, and many
others. To know at what point in the long series of
somewhat similarly marked species the original function
has ceased, would require impossible study.
While it is plain that a hundred needs may each be
represented in the pattern- and colour-schemes of animals,
it is also plain to an artist’s eye that in most butterflies all
visible details of colour, pattern, and form are essential
parts of the representation of flower-scenery. And it is
surely conceivable that, in a certain region, one particular
form of flower-scenery-representation may furnish such
advantages to butterflies as to cause many widely-separated
species to become modified till they wear a common aspect;
and it is conceivable also that. there would be one common
form of wing which would best lend itself to this scheme.
Surely we do not know enough of the habits of these
insects or of the regions that may be their strongholds to
558 Mr. A. H. Thayer on
feel sure that this hypothesis is absurd; and were it
correct, it would complete a chain of seemingly perfect
evidence.
After we see how inexplicable it would be if butterflies
did not either resemble flowers, or represent some portion
of flower-scenery, why should we, in view of the endless
variety of flower-forms, stick at any form or pattern in
the butterfly that frequents them ? One must constantly
remember that any pattern is less conspicuous than bright,
unshiny monochrome. Therefore, “conspicuous” is not the
right word for the character of patterned butterflies,
Now since the Jthomiine, Heliconiny, and Danaine,
such for instance as the similarly coloured cow-red and
chrome-yellow, black-bordered J/elinwa, Heliconius, and
Lycorea (and equally, in other colour-schemes, all the
other so-called mimicking groups), are in every way com-
pletely painted by Nature into these three tones,—the
note of shadow under vegetation making their borders,
which it occupies, coalesce with the shadow under the
flowers, and disappear, while the red and chrome wonder-
fully reproduce the colours and patterns of such flowers
as Odontoglossum triumphans, who shall say that it is not
to this flower—which perhaps, by its abundance, dominates
the region—that these cow-red and chrome- yellow butter-
flies owe their common appearance? Some such flower
may be overwhelmingly attractive for its honey.
Perhaps the most conclusive of all our evidence is to
be seen in the transparent winged members of these
mimicry groups. Dismorphia orise, for instance, with its
green transparencies enclosed in a pattern of the same
velvety dark fuscous that I have already described. What
conceivable artifice could offer greater opportunity for
frequently remaining unnoticed amidst flowers and leaves ?
These httle green windows must of course allow any
bright object to show through them, while the fuscous
cuts the aspect to pieces by representing a shadow far
below the insect. The very word transparent wrecks any
theory of conspicuousness or adaptation suitable for a
badge. Add to this the present belief that the trans-
parency has been attained through selection, and ought not
those who hold this theory to “believe that concealment
was obviously the goal of a change toward invisibility ?
It is hard to conceive of a better device for representing
little green leaves than by these glossy green, leaf-shaped,
Protective Coloration in its relation to Mimacry, ete. 559
and leaf-veined windows, bordered with imitation back-
ground, and ever ready to look like glossy leaves the
moment they are extended over a bright flower or other
bright object.
Professor Poulton has already noticed the efficacy of
the imitation hole in the wing of Grapia (a device similar
in effect to the gold dots on some pupe).
During the writing of this article I have been learning
that iridescence itself is an immense factor of conceal-
ment, far greater than I at first realized. I have lately
had excellent opportunity to study several species of
golden-brown butterflies with sheeny black tips spotted
with white, and I begin to realize the wonderful power
of this combination. The white dots stand changeless,
while upon the black, in bright sunlight, faint rainbow
sequences dissolve the actually flat wing-surface into
liquid depths, apparently wholly detached both from the
insect and from the white spots, which appear, as I before
said, to be shiny points like dewdrops down in the spaces
below the buttertly.
If butterflies were mimicking each other, Mr. Blandford’s
objection (Proceedings of the Entomological Society, 1897)
that the resemblances would be hypertelic would seem
true. Since an attempt on Nature’s part to give common
colours and patterns to a group of insects involves no need
that any one of them shall have sharp delicate contours
of spots, or have subtle gradations, these species would,
if their object were to resemble each other in their colour
and markings, stop short of such sharp contours, ete.
On the other hand, if they are representing flowers or any
organic forms instead of merely patterns, etc., on forms,
they would profit by the utmost minute finish of every
part of their design, since just this finish, this microscopic-
ally perfect smoothness and minuteness of detail is an
essential characteristic of flowers and even of leaves.
Upon my hypothesis, the many “ warning-colour”
species that have dull-contoured spots instead of sharp
ones, would seem (as they do to the supporters of Mimicry)
to be species in process of adaptation, but to the aspect of
flowers, instead of to that of each other.
As soon as the advocate of the Mimicry theories sees
that to wear the region’s prevailing pattern tends to con-
ceal, his case looks bad; since we see throughout the
animal kingdom common coloration, and often common
560 Mr. A. H. Thayer on
form in widely separated orders, plainly accompanying
common environment and habits. The Salmon’s silver,
grading upward into dusky, and downward to purest
white, is identical with that of countless fish in many
groups, and no one doubts that environment and habits
are the cause. Among birds, Lmberiza miliaria, Anthus
pratensis, Alauda arvensis, and Alauda arborea are four
species of three genera for all four of which one minutest
colour-and-pattern-description would almost suffice; and
the same colour-scheme and pattern with slight varia-
tions is found on a great many other species throughout
the world, both of Passeres and even Scolopacide and
(alline, telling plainly of life on the grownd amidst
grasses. Among the Scolopacidx, many females and young
of the Anatidxy, and the Laridx, Nature betrays, in the
main, great lack of variety in design, easily accounted for
by the lack of variety im the aspect of the environment.
In a broad survey of the animal kingdom we perceive that
everywhere the degree of colour-and-pattern difference
between different members of an order, family, or genus
keeps pace with the degree of variation in their environ-
ment’s aspect.
Why may not the circumstances of a group of butterflies
furnish them similar needs to wear a common livery, even
if we cannot see the reason? Might they not tend also
to have their flavour similarly affected by similar food ?
The Spruce Grouse (Canachites canadensis) is saturated
with spruce flavour, and the world is full of such cases.
Even the amazing similarity between members of these
groups is no proof they may not, for reasons which we
have not discovered, profit each by exactly the same form
of concealing-coloration. It should be borne in mind that
it is not a flower that these mimics evidently represent,
but a certain combination of the flower’s aspect with that
of its surroundings. Hence there may be one best way
to render this. Butterflies on wing are conspicuous, but
are wonderfully protected by their jerky flight, which is
completed by their wings being so large as necessarily
to throw the body up and down at every movement.
This latter advantage, attainable by no other conceivable
means, may be a great factor in the whole matter. In
flight they are doubtless practically safe, 7.¢. too trouble-
some a quarry to be seriously decimated. I send, for
Professor Poulton to exhibit, photographs of a number
Protective Coloration in its relation to Mimicry, etc. 561
of so-called conspicuous butterflies (dead specimens), the
examples having been placed as far as possible without
an unfair attempt to favour my argument, except in a
few cases where the attempt is obvious. Surely they
speak eloquently. Could they be seen in their colour-
coalition, they would speak even more so, Any one care-
fully examining them will see that, in most cases, their
dark parts are not distinguishable from the background
(although the average person, wnaccustomed to analyze his
sight, will, by recognizing the butterfly through its pattern,
fancy he sees every part).
The very keynote of the zoologist’s error is psycho-
logical, One sees only what is out of place ;—that which is
in place is harmonious and unnoticed. We know how
many of these concealed animals we sce, but we do not
dream of how many we pass by.
By tracing back to so palpable an example as our
Sphinx-moth photograph, we see that the various com-
binations of sharp-edged markings with delicate blendings,
exactly resembling the combination of patterns made by
any sharp-edged fabric lying near a ground on which its
shadow falls, do vepresent such combinations of form ; so
that we must believe that so elaborate and delicately
complete a design would scarcely exist merely to identify
a species as unpalatable. We find on several Preces, as on
many Vanesse, and Papiliones, very highly developed
cases of the varied combinations of design worn by multi-
tudes of the most obviously protected birds, and other
animals ;—slight variations of representation either of near
objects casting a shadow on the background, as in the
cats, snakes, and moths mentioned, or of near objects
relieved against more distant, fainter ones, as in the
Kuropean ‘Woodcock’s wings, many female Pheasants, and
male Pheasants’ tails, such as that of the Copper
Pheasant. Doubtless each species has some particular
headquarters, as it were,—some region which it fits best,—
aud unless we chance to study it in this very region, and
at the most favourable season, we shall never witness the
full operation of its protective colour-scheme. Mr. Frank
M. Chapman has already pointed this out in a paper
entitled mOn the Birds of the island of Trinidad,” published
Feb. 1894, in the “ Bulletin of the American Museum of
Natural History,” a paper containing some very prophetic
glances into the future of protective coloration.
562 Mr. A. H. Thayer on
Apparently Nature has two main _protective-colour
schemes; one of which is closely imitative of the very near
environment of the animal, and applicable to such species
as sit close, and keep still, for concealment, as do the tree-
toads, moths, goatsuckers, certain snakes, and, among butter-
flies, the species of Grapta. (The latter, at least, keep very
still when resting, and expose at such times only the rock
or bark representation on the under-side of their wings.)
Among those butterflies, on the other hand, which have
no pronounced habit of protecting themselves in this
manner, Nature seems to have been forced to a_ bolder,
more positive way by furnishing them an upper-side
bearing a sort of conventionalized representation of the
predominant details among which they are destined to
move. Flowers, of course, must almost always be present.
And always the notes of the conventionalization are perfect.
Here is a most impressive argument, viz., so-called con-
spicuous butterflies have the body, head and all, exquisitely
effacively graded. Would it not be absurd for Nature to
spend energy in effacing the body while making the wings
conspicuous? The multitude of species, the world over,
whose main colour is largely the peculiar fuscous of
shadow under vegetation, have in most cases not merely
this shadow-colour, which so perfectly coalesces with the
shadow and apparently vanishes from the insect, but also
a system of exquisitely delicate perspectives within the
patches of shadow-colour; as in the genus Caligo es-
pecially. I mean that Calig go is an exquisitely developed
representation of the perspectives which an artist sees in
peering down through the openings between the flowers.
The parts of the world which I know well do not yet
furnish me a clear vision why so many butterflies, such as
several Preces, and Anosia plexippus, for instance, have
these delicate perspectives done in golden brown instead
of either shadow-colour or the more delicate flower-colour ;
but that this delicate design does represent perspective,
and would be wasted if used for any attempt at conspicu-
ousness, and that it is entirely akin to the perspectives
rendered on perfect shadow-colour in so vast a number of
species, is reason enough for’ trusting it to prove to be
some form of concealment device ; and on red flowers
these species show surprisingly little. I myself suspect
that butterflies of the A. plexippus type represent half a
concave flower. Watch any butterfly of this class, or any
Protective Coloration in its relation to Mimicry, ete. 563
of the classes in which the pattern, when the wings are
open, arranges itself in amphitheatre-like semicircles of
stripes or dots, ete. When such a butterfly rests with
open wings on a flower, its head is at the centre, its
antennz form two stamens, and these semicircles seem to
belong to half the flower of which its head is the centre.
In several Preces, and many other butterflies, there is a
general representation of something like a bunch of
stamens casting their shadow deep under them in the
flower’s cavity. Usually a butterfly’s upper-side has the
exact colour-note characteristic of flowers and flower-
scenery seen from right overhead (take, for example,
Papilio turnus) ; while its under-side is a picture of such
greater distance as would be seen from the s7de position
necessary for beholding it when the wings are in their
characteristic vertically-folded position; and this is the
position from which enemies on neighbouring bushes
would see it. So-called “conspicuous” butterflies have,
in short, their upper-side designed with the full strength
“values” of the nearest flowers looked into from above,
and their under surfaces designed in notes more delicate,
to counterfeit the distance, and a perfectly effacively-
graded body. Their under-side is also more delicately
finished, as if against the nearer inspection possible from
neighbouring bushes. In fact, they wear every conceivable
aspect to fit them into the background from each point of
view, and make you think you see through them; or else,
seen from above, to make you think, as in the case of the
Pierine, that you see a flower itself. How can such a
case call for a theory that is based on the hypothesis that
they are conspicuous? One very important fact is that
we have abundant proof that animals, including birds,
have totally different sight from ours; and the existence
of these patterns, etc., unless 1t can be denied that they
even tend to eftace, should be taken as proof that they
sufficiently succeed in effacing. Otherwise, why are they
there, when almost the whole animal kingdom does need
concealment? A fox,a deer, a bear, a grouse, a turkey,
or any small bird or mammal, may come almost to one’s
feet if one stay still, yet flee wildly on seeing any motion.
Ts not this sufficient proof that even if we were usually
able to detect a Papilio when it is effacively situated, it
is no sign that a bird could do so, if the insect kept its
place ?
TRANS. ENT. SOC. LOND. 1903.—PART IV. (DEC.) 38
564 Mr. A. H. Thayer on
Butterflies very often remain unobserved amidst flowers
or other vegetation, by any one approaching (especially
if he be not keenly in search of them) until once flushed.
Of course our yellow and our white Pierine are pretty
sure to catch the eye of the person approaching, if, as
very commonly, they are found amidst dark vegetation.
Yet their colours are precisely those of our most abundant
flowers, just as they are our most abundant butterflies.
This fact harmonizes with my argument that, however
conspicuous in many situations, few animals are so in the
place or region to which they doubtless owe their abund-
ance. We see largely the overflow individuals from a
concealing region into a less favouring one, and erroneously
think of the species as typical of the region where it is
visible to us. The gentle waving of the wings, so common
among butterflies when they are feeding, seems plainly a
protective imitation of the swaying of leaves and flowers
in the breeze. Any one who has photographed outdoor
vegetation knows how seldom it stands still.
To sum up, the general aspect of each animal’s environ-
ment, throughout the animal kingdom, is found painted
upon his coat,in such a way as to minimize his visibility,
by making the beholder think he sees through him. How
has it chanced that, while this fact has long been recog-
nized, in a crude way, in many fields of zoology, it has
remained essentially unnoticed in butterflies? Their
most critical moments being passed upon flowers, the
aspect of flowers combined in various proportions with
the dark vistas down among them to the shadowy earth
beneath, is exquisitely painted upon a vast majority of
the world’s butterflies, and on none more plainly than
on those called conspicuous. The Pierine are mainly
representations of flowers, though surrounded by a dark
border which appears to belong to the shadows beneath
it. On the other hand, there are a vast number of dark
species which represent a portion of this shadow-under-
vegetation, with bits of yellow vegetation, or of flowers,
seen against it (these of course being rendered by the
hight markings), Could small, bright | patterns on dark
possibly be more perfect generalizations of small blossoms,
buds, and stems ?
I cite the followmg examples of the various colorations
described.
Among the Brassoline, Caligo, eurylochus is a marvel of
Protective Coloration in its relation to Mimiery, ete. 565
wholly effacive design, so subtle as to make it absurd to
suppose that Nature could be trying to have him conspicu-
ous, or to use such delicate gradations for zdentification.
Caligo telamonius and Caligo demosthenes are even more
wonderful examples. Cynthia has a wonderful multiplicity
of perspectives represented on its surface. Black and
green Nymphaline are notably orchid-like in design.
Their dark tips disappear, uniting with the shadows.
Dione has good near-scenery on its upper-side, while
the silver spots of its under-side appear in a side view to
cut holes through its wings.
The Danaine butterfly ZLimnas chrysippus is covered
with design which I am not prepared to interpret.
Whether or not it is a flower, the four interior spots on
the upper-side of the hind-wings may pass for stamens, as
may also, of course, the antenne ; and whether or not the
yellow-red ground counterfeits the colour of a flower, it
represents a flower’s form. Caduga melaneus has the colour-
scheme of the skunk, with, of course, similar advantages.
The Satyrine, i.e. the dark ones, with strong, light
patterns, have also the skunk’s colour-principle. The
Danaine, Ithomiine, and Heliconine of South America,
Lycorea, Melinza, and Heliconius, for imstance, display
marvellous mutual resemblance, yet their likeness to
Odontoglossum triumphans, when their dark tips are cut
out by coalescing with the shadow, is most impressive.
Among the transparent Satyrine I may mention
Pierella nereis. Unmistakably the whole surface of this
insect (and likewise that of Citherias menander) pictures
a single flower.
Pierella astyoche represents flower-scenery (likewise
Pierella rhea).
In the Oriental Danaine genus Luploca we see exquisite
shadow-perspective over which white spots relieve. The
blue sheen, seldom or never occurring on both wings at
once, additionally effaces.
In the Lycenide the exquisite blue species represent
flower-cups, their black border of course detaching into
the background.
The above examples I have chosen from all the families
I have lately examined, which do not include the Skippers,
or the great mass of Papilionide.
Let me add a few more reflections, all harmonious with
my theory.
566 Mr. A. H. Thayer on
The act of flight tends to obliterate pattern, by the
too quick substitution of one colour for another before the
eye. A black-and-white butterfly, therefore, tends to
look simply grey in flight.
It is not necessary to conceive that a bird must find the
imitation flower on its proper plant, if the flower represent
a type common in the neighbourhood. A vast majority of
butterflies, including most members of Mimicry groups,
have the common dark wing-tips of the /wscows colour
which causes this portion to seem lacking from the butter-
fly, leaving the lighter-coloured parts to represent a more
flower-like form. The white dots, so common on these
black tips, surprisingly aid the representation of space
below the flower by supplying the average sharp details
that are to be seen down in the shady under-spaces,—
little glints of light on twigs, etc.—and their dark ground
is rendered additionally transparent in appearance by
iridescence.
If the foregoing arguments prove that the so-called
Warning-colours commonly cited do not exist mainly to
make their wearer conspicuous, it does not follow that
they may not still serve secondarily as Warning-colours.
When, for instance, they happen to fail to conceal, they
may then serve to warn. My main point is that they
first of all conceal. I suspect that the same principles
apply to striped wasps and hornets, and many other
insects called conspicuous. The yellow pattern unmistak-
ably allies their appearance to the pollen-covered flower-
interiors, making them far less conspicuous than an
unmixed need to be seen would have them. Yet when
seen, they may well profit by the pattern’s recognizability.
Can any one, once shown, as I here show, that butterflies’
patterns are 7o¢ intrinsically the thing to make the wearer
conspicuous, and shown that they ae wonderful representa-
tions of the flower-scenery I describe, believe that Natural
Selection has bungled, and wasted design of the most
intricate kind? No, it is the beauty of the whole thing
that absolute fitness is the goal of all changes by Natural
Selection :—is, in fact, the only motive-power; changing
all forms steadily toward itself.
We see, then, that butterflies are imitation flowers, or
pictures of flower and background. This has escaped the
eye of zoologists. They see that fish wear representations
of under-water scenery; that forest animals are forest-
Protective Coloration in its relation to. Minicry, etc. 567
patterned ; beach animals, beach-patterned, etc., through
the whole animal kingdom. But this other obvious case
has escaped them. What other equal hope were there for
insects that feed in full sunlight on masses of bright
flowers ?
In another paper I shall extend this criticism on the
animal-conspicuousness-theory to the field of birds, and to
strengthen the present paper by showing reasons to sus-
pect that this theory is also not well intrenched in the
bird part of its field, I append the following examples of
the material to be used in the next paper.
Several of the most apparently conspicuous details of
the exteriors of male birds can be shown to be such as
would aid them to escape their enemies, and it is plain
that simple life-preservation must for ever take precedence
in the scale of importance of animals’ needs. It is a mild
statement to say that if the animal kingdom is to survive,
females have greater need of the mere existence of mates
than of any particular attribute in them, and if this state-
ment is true, in all its immense import, it is among the
most primitive needs of the male, that we should search
for the explanation of his present attributes. All the
nuptial developments, either of feathers or fleshy growths
on beaks, etc., are much more rationally explicable along
the simple lines of utility, than those of direct Sexual
Selection, since it is apparent that every appendage, and
every brilliancy of colour or costume adds to the formid-
ableness of a warrior’s aspect. One male conquers another
partly through overawing him by superior splendour,
and actually looking larger by means of his appendages,
and when these gaudy-feathered braves flaunt before their
females, why are they not presumably appealing to the
females’ love of a good fighter,—a sentiment so dominant,
even in the human race,—and a simple sense of what con-
stitutes a husband full-equipped for the rough work
devolving on all feudal lords? In fact, from which end
of the animal scale is this human sentiment traceable ?
If from the lower, as seems obvious, it must exist there.
I believe that a material need for any existing thing will
always be found to precede the spiritual, just as simply as
a man must catch before he can eat, and will then think.
These arguments suggest, at least, that the nuptial
superficial developments are for the direct use of the male
who wears them. Let us look at the iridescent splendours
568 Mr. A. H. Thayer on
ot the Peacock family. An artist can see that whereas
unshiny monochrome reveals its wearer to the utmost,
iridescence, on the other hand, destroys visibility of surface,
by substituting for a normal light-and-shade gradation, a
totally new succession of colour and light notes, and above all
one that changes its character with every movement of the
bird, and every change of the beholder’s standpoint. Add
to this in the Peacock’s case, for instance, his habitual
resort to dense cover, and his gorgeous blue and green
gleams, through its interstices, present merely the aspect
of foliage-colours and hints of flower-masses. I feel sure
that Peacock hunters will testify that this bird is hard to
see when lying close.
Let us imagine an animal stalking this bird. He will
look wholly for motion :—(such at least is the habit of all
predatory creatures I know). Now it is the peculiar
property of sheen, that it will stand sti// while the thing it
is on moves. This means that a Peacock can move his
brilliant neck, while its sheen stands stil/—just as the
gleam on the telegraph wires keeps pace with the railway
train as one sees it from the window. And since this
gleam of the bird’s neck must be the most visible thing,
the possibility of the neck’s gliding along behind it, while
at stands still, must often save the Peacock ; (for the balance
between the evolved skill of the hunter and the evolved skill
of the hunted must always be close, and smallest advantages
must often tip the scale). While the fore-part of the bird
is beginning to move, unnoticed, his conspicuous tail, a
yard behind his vital parts, catches the tiger’s eye, in its
earliest motion, and the tiger, seeing no other part so
distinctly, springs at these long feathers, whose design is
arranged for conspicuousness in motion.
These gorgeous birds will prove to be additionally con-
cealed, not revealed, by their costumes. It is worth men-
tioning here, in connection with the Warning-Colour
theory, that while Peacocks and Pheasants are iridescent
plumaged birds, and would be called conspicuous in the
highest degree, they are not wapalatable ;—a fact that goes
to strengthen my argument.
The next thing to be pointed out is that the general
tendency of birds to wear longitudinal markings forward,
and transverse ones aft, is an important factor of protec-
tion, especially in the case of the Pheasants and Peacocks,
among whom this arrangement is very highly developed.
3
Protective Coloration in its relation to Mimicry, ete. 569
Any one who has tried to catch a snake in the grass will
see at a glance why Nature tries to direct an enemy's
attention behind the animal he is hunting. The snake
for ever proves to be further on. It is hard to set one’s
foot far enough ahead as he moves, just as a wing-shot
tends to shoot behind. Now Nature, realizing this, offers
the enemy the utmost inducement to strike too far back.
The strong cross-bars of the Reeves or the Copper Pheasant,
while visually they cut the tail to pieces when it is still,
are, as with the Peacock, by far the most visible part of the
bird as soon as he moves. The reason of this is that in
forward motion the longitudinal markings scarcely show,
while the transverse ones become conspicuous. To prove
this, any reader has only to blacken a few points an inch
or so apart on a white cord, and then move the cord longi-
tudinally, drawn tight across some aperture a few yards
away, the cord being only visible where it crosses the
aperture. He will see that its motion is distinguishable
much farther off when the spots are in sight than when
the wamarked cord is passing. The spots correspond to
the tail-marks of the Pheasant, and the cord where it is
not spotted represents the bird’s longitudinal markings, 7. e.
his body-markings.
Before closing I beg to say that I do not mean that I
am convinced that Mimicry and Common Warning Colours
have no hand in these resemblances. I merely point out
that the coloration of every individual of the “ mimicking
groups” of butterflies seems to be the best conceivable for
effacing the aspect of its wearer, and also that it is per-
fectly conceivable that an external influence, like super-
abundance of certain very sweet flowers, could do the
whole thing.
570 Professor E. B, Poulton on Ir. Thayer's
XXVII. A brief discussion of A. H. Thayer's suggestions as
to the meaning of colour and pattern in insect
bionomics. By PROFESSOR EDWARD B, PoULTON,
M.A., D.Sce., F.R.S., ete.
[Read October 21st, 1903.]
THE discoverer of the meaning of the white under-sides of
animals is entitled to a respectful hearing on any question
of animal coloration. Furthermore, by his discovery, he
has proved the benefits which the artist can confer on the
naturalist, benefits which we naturalists are only too
pleased to receive with gratitude. Our only difficulty is
that so few artists seem disposed to consider our problems
seriously. In order to be able to do so they must become,
at least in spirit, naturalists as well as artists. The more
numerous the men of creative power who can occupy, as
Mr. Thayer does, the double standpoint, the better it will
be for both domains. I therefore express my cordial
agreement with Mr. Thayer’s claim for the artist. I now
propose to make a few comments upon the details of his
interesting paper.
Every naturalist will agree that “any coloration or
pattern would be conspicuous somewhere.” We have often
called attention to the fact that colour, pattern, shape,
and attitude can only be understood in the natural environ-
ment. In fact, Mr. Thayer’s own suggestions are, I think,
most open to criticism when he is speaking of animals in
countries he has not visited; when, for instance, he sug-
gests the kind of concealment brought about by the stripes
of the zebra. The lion is the zebra’s great enemy, and in
spite of their very different kind of colouring they are both
adapted to the same general environment. The propor-
tion of dark and light stripes, Francis Galton told us long
ago, “is such as exactly to match the pale tint which arid
ground possesses when seen by moonlight.” So too the
suggestion that the groups of similar South American
butterflies have gained their resemblance by a common
(syneryptic) likeness to some flower which they chiefly
frequent would be more plausible if Mr. Thayer had
studied them in their native haunts. I have asked Mr.
suggestions as to insect colours and patterns, 571
W. J. Kaye if he can remember the colour of the
tlowers visited by the black, cow-red, and yellow Melinxa
group and its mimics in British Guiana, and he tells
me they are either white or cream-coloured. Further-
more, Mr. Thayer treats this group as though it
were uniform throughout tropical South America, dis-
regarding the extraordinary changes of colour and
pattern undergone by its representative species as we pass
from one part of the Neotropical region to another. It
is almost inconceivable that the following features, which
are characteristic of whole groups in particular areas, can
be due to the special flowers of those areas. The barred
form of Central America, Colombia, and Venezuela, the
black hind-wing of the Guianas, the bright yellow band of
Eastern Brazil, the chestnut ground-colour of Ega on the
Amazon, the black marked fulvous of the Napo River,
passing on into the black forms with fulvous marks which
constitute so large and characteristic a group in Ecuador,
Peru, and Bolivia. In all these cases, nothing short of
actual evidence on the spot can warrant the improbable
suggestion that we are dealing with syncryptic groups,
changing as the species of flowers are replaced by others
in passing from one district to another.
Moreover, the theory of a syncryptic resemblance to
flowers fails to account for certain broad characteristics of
the groups in question, which on the other hand receive
a ready explanation on the theory of common warning
(synaposematic) coloration. These are (1) the predomin-
ance of forms belonging to the sub-families Jthomiine,
with the allied Danaing, and Heliconine, with the allied
Acrwing : (2) the fact that the predominant members of
the chief groups in all the other tropical parts of the
world are also contributed by the Danaine and Acreine :
(3) the flaunting flight, exposure at rest, and general
want of alertness exhibited by the species of these sub-
families as compared with others: (4) the more or less
exact similarity of the pattern on the under to that on the
upper surface, an arrangement comparatively rare in other
Rhopalocera: (5) the experimental evidence of the un-
palatability of these very sub-families to a large number
of the enemies of insects.
Hence, until positive evidence is obtained on the spot
in favour of Mr, Thayer’s suggestion of syncryptic resem-
blance, I must regard such an interpretation as highly
572 Professor E. B. Poulton on Mr. Thayer's
improbable, in the case of the groups hitherto explained
by the Miillerian or Batesian theories. Of course close
syncryptic resemblances between bark-like moths, lichen-
like moths, grass-like and pine-needle-hke larve, etc.,
have been known and admitted for many years.
Leaving the tropics we find a beautiful example of
mimicry, Batesian, or more probably Miillerian, which has
arisen in Mr. Thayer’s own region, and has never wandered
much beyond it, an example moreover very well known to
the American artist-naturalist, viz. the resemblance of the
northern Limenitis (Basilarchia) archippus (misippus) to
the Danaine intruder from the tropical south, Axnosia
plexippus.
In this case there is little doubt that the Nymphaline
has been actually drawn away from an ancestral appear-
ance, much like that now borne by ZL. avthemis, explained
by Mr. Thayer as promoting concealment by likeness to
flower-masses and their background. If therefore Mr.
Thayer is compelled to admit all this effect produced by
the Danaine intruder in his own northern region, why
should he not be ready to accept far more extended effects
of the same kind in the crowded luxuriant life of the
tropics ?
I do not think that naturalists Lave so entirely misunder-
stood the principle of a cryptic pattern resembling some
object in the environment combined with the eftacive
gradation so admirably explained by Mr. Thayer. His
illustrations cf tiger, lion, brilliantly-coloured fish, appear-
ance of forest and shore birds, ete., all these are accepted
at once and have been accepted for a long time. But
naturalists have regarded the skunk as conspicuous, and I
feel sure that Mr. Thayer will admit that it falls into
another category from that which includes the forms just
named. If concealment is brought about by the beautiful
and delicately adjusted effacive gradation from upper dark
to under white, as is now generally admitted, surely the
“slight amount of eftacive gradation” of the black
skunk cannot be the same thing, or belong to the same
class.
We must admit Mr. Thayer’s main conclusion, that the
forms we call conspicuous might be more conspicuous, and
also accept the statement that a pattern is less conspicuous
than the monochrome.
Admitting all Mr. Thayer says, at least of the butterflies
suggestions as to insect colours and patterns. 573
he knows in the living state, and of the skunk, he cannot
contend, I think, that his criticisms are powerful enough to
transfer these examples into the bionomic group which con-
tains the well-known examples of cryptic colouring—the
skunk into the same category as the hare or ptarmigan, the
under-side colouring of the Danaine butterflies, or the Nym-
phaline genus Limenitis (Basilarchia) into the samecategory
with that of Grapta or Kallima, etc. I believe the whole
of his criticism of warning colours can be accepted, and
can be reconciled with the existing hypotheses. All
animals with warning colours have enemies, all are
liable to special attacks, in times of exceptional hunger,
by enemies which would at other times neglect them.
Even the skunk has special bird enemies. Provided such
forms are easily seen and avoided by enemies which
respect their special modes of defence, it is clearly an
advantage to be as far as possible concealed from those
which do not respect them. Hence conspicuousness, but,
as Mr. Thayer tells us, something very far short of ideal
conspicuousness. The black and white pattern of the
skunk is probably glaring and conspicuous enough to all
enemies near at hand, but at the immense distance covered
by the long-range sight of a predaceous bird it may melt
into an inconspicuous grey.
The same kind of interpretation probably holds for a
cryptic element whenever it exists in the appearance of
butterflies belonging to distasteful sub-families. It is the
probable meaning of the transparency so widespread in
the Ithomtine, although I do not think it is so effective
in concealing as Mr. Thayer supposes. We must remember
that many of these transparent species are excessively
abundant, flying in clouds often made up of the individuals
of several species and different genera. I quite recognize
that the transparency may protect such forms against
distant enemies, but I should be much surprised if the
species of Methona and Thyridia, as well as Dismorphia
orise, of which they are the models, are not rendered
extremely conspicuous to enemies close at hand, by their
numbers, habits of flight, and attitudes of rest. As Mr.
Thayer has said, the black and white markings will melt
into an elusive grey on a rapidly vibrating wing ; but the
specially protected groups have developed a sailing flight
which shows off the elements of pattern to perfection.
When the body in such groups is effacively graded the
574 Professor E. B. Poulton on Mr. Thayer's
explanation may well be that it is advantageous to direct
attention to the wings rather than the vital parts; but it
is precisely in these groups that the black body, and some-
times the head, are so often marked with white or red. <A
bright red or orange collar is found in several species.
Furthermore, it must be remembered that the body being
moved much less rapidly than the wings during flight is
more easily seen. The black and white apical area of the
fore-wing may help to conceal, as Mr. Thayer supposes,
under certain conditions, but the numerous examples of
injuries at this very spot, figured in Plates IX and XI of
our Transactions for last year, strongly support the hypo-
thesis that it is directive, and diverts the stroke of the
attacking enemy from the body.
Apart from the suggested interpretation of mimetic
resemblance, which I believe to be untenable, Mr. Thayer's
suggestions supplement and complete rather than oppose
existing hypotheses. The words he uses of the wasp may
in fact be employed of the skunk, and the well-known
distasteful Rhopalocerous groups, ete. The colours may
not be conspicuous to enemies at a great distance, “ yet
when seen they may well profit by the pattern’s recogniz-
ability.” We have rather insisted on this latter fact and
its advantage, and Mr. Thayer has done us good service
in calling attention to the other aspect of the appearance.
Ideas not dissimilar to those of Mr. Thayer’s upon
warning colours have for some time crossed my mind.
Thus last year I suggested as regards the abundant, much-
mimicked Limnas chrysippus, that its desert form dorippus
(Alugit) “is a development in a procryptic direction in areas
where the struggle” is especially severe (Trans. Ent. Soc.
Lond., 1902, p. 475).
Furthermore, the idea has often forced itself upon me
that the ground colour of the type form of this butterfly, as
well as of the Ethiopian Acrwine and Lycid beetles, may,
under certain conditions and at a certain distance, become
procryptic against the prevalent reddish tinge of the soil
of Africa.
The author’s suggestions of the resemblance of buttertly
patterns in general to flower-masses and the shadow-depths
between them; of the under-sides of Grapta and the
upper-sides of many moths representing dead leaves lying
on the ground and casting such shadows as they would
throw at their small distance from it; of the concealing
suggestions as to insect colours and patterns, 575
effect of iridescence ; of the overflow of individuals from a
concealing region into one less favourable—in all these we
have illuminating ideas which demand the fullest and
most respectful consideration. That they are sound prin-
ciples must, I think, be admitted at once ; but their relative
importance, the amount of ground which they cover, cannot
be decided offhand. I would only point out the extra-
ordinary frequency with which a continuous black colour-
ing unrelieved by pattern is accompanied by iridescence
or surface colours of some kind. In view of the whole
drift of Mr. Thayer’s interesting and most suggestive paper
it becomes probable that dead black would be too con-
spicuous even to many a well-armed aculeate or nauseous
Huploea, and that it is therefore modified so that it obtrudes
less upon the distant view of enemies which “ mean
business.”
Although I have criticized some of the details of Mr.
Thayer’s paper, I should wish again to point out that they
concern just those species which have not come under his
own eyes in the living state. Naturalists owe him a large
debt for the many new points of view and illuminating
suggestions contained in his memoir.
DrEcEMBER 24th, 1903.
*)
one
THE
PROCEEDINGS
ENTOMOLOGICAL SOCIETY
LONDON
For THE YEAR 1903.
February 4th, 1903.
Professor E, B. Poutron, M.A., D.Sc., F.R.S., President, in
the Chair.
Nomination of Vice-Presidents.
The PrestpENT announced that he had nominated the Rev.
Canon Fow er, M.A., D.Se., F.L.S., Professor RAPHAEL
Metpona, F.R.8., and Dr. Davip SHarp, M.A., F.R.S., F.L.S.,
as Vice-Presidents for the Session 1903-1904.
Election of a Fellow.
Mr. 'T. Asuton LorrHouse of the Croft, Linthorpe, Middles-
brough, was elected a Fellow of the Society.
Exhibitions.
Dr. Tt. A. CHapman exhibited two male specimens of Oring
(Chrysochloa) tristis, var. smaragdina, taken at Pino, Lago
Maggiore, on May 30, 1902, still alive; and living larve of
Crinopteryx familiella, second generation, bred at Reigate from
the egg. The parents were taken at Cannes in February 1901.
The Rev. F. D. Morice exhibited, with drawings of the
abnormal parts, a hermaphrodite of Hucera longicornis, Linn.,
showing one 2 antenna normal, and one gd antenna remarkably
PROC. ENT. SOC. LOND., I. 1903.
A
( ii)
shortened and with the joints greatly dilated; the clypeus
and labrum one half white (the ¢ character), and the other
half black as in the 2. In the abdomen and legs the ?
character predominated, but one half of the apical segments
and genitalia seemed to be 6.
In a discussion which followed on hermaphroditism, Dr.
Suarpv stated that Father Wasman had announced the
discovery that in certain Diptera, parasites of Termites, the
individual commences as a male and ends as a female—a
phenomenon entirely new to entomology, though paralleled in
some other groups. Father Wasman had examined a large
number of cases, and considered that the whole genital system
changes in the course of the imago life from male to female.
It was noteworthy that in this case of chronological her-
maphroditism protandry existed, as in all other cases of the
kind. The Prestpenr expressed the opinion that such a
discovery required the confirmation of several workers.
Mr. R. McLacunan, F.R.S., exhibited a living example of
Chrysopa vulgaris, Schnd., taken by Dr. Chapman in his
house at Reigate. The primary object of the exhibition was
to show the manner in which this species, which is ordinarily
bright green, assumes a brownish colour, the abdomen being
often marked with reddish spots in hibernating individuals.
Mr. W. J. Lucas submitted specimens of a bug—J/iris
calearatus—and some fruit of a grass, swept up together by
Mr. W. J. Ashdown from the canal side near Byfleet on July
14, 1902, on the occasion of the South London Entomological
and Natural History Society’s excursion. The similarity of
form and colouring constituted a probable case of protective
resemblance.
Major Nevitte Manvers exhibited two specimens of an
undescribed species of Ated/a from Ceylon ; and remarked that
it was a very local insect, and only found in the Nitre Cave
district, one of the localities most remote from civilization in
the island. It was probably a well-marked local race of
A, alcippe, but easily distinguished from any known species of
the genus by the apex of the fore-wing being entirely black.
The species is constant in colouring, but apparently varies
considerably in size.
( ii)
Mr. F. B. Jenninas exhibited British specimens of two species
of Hemiptera—Heteroptera : (1) two females of Drymus pilipes,
Fieb., a rare species of the family Lygwidz, which were found
among dead leaves on a hillside near Croydon in September
1901; (2) the black aberration of Miris levigatus, L., recorded
by him in the Entomologists’ Monthly Magazine for 1902,
p. 224. The species of Miris and the allied genus of Capside,
Megalocerwa, are ordinarily grass-green, or pale-yellowish.
Mr. H. J. Etwes, F.R.S., exhibited two cases of arctic
butterflies. The first contained specimens from a collection ~
formed by Mr. David Hanbury on the arctic coast of North
America, in the region where the Parry expedition was lost.
Of the butterflies observed—fifteen species in all—a large
number for such barren and inhospitable ground, two had not
been taken since they were first described by Curtis sixty
years ago. Among them was Clias boothii. This species, in
comparison with Colias hecla, Lef., is undoubtedly distinct
in both sexes, but it is most remarkable that the male
in coloration and markings appears to approximate more
closely to the characters usual in the females of other members
of the genus. The collection contained nothing new, but
included the rare and curious Argyunis improba, Butler,
hitherto taken only in Novaya Zembla (cf. Markham’s “ Polar
Reconnaissance,” p. 351), which Mr. Etwes believed to be
nothing more than a high arctic form of A. frigga, Thnb.; a
remarkable aberration of A. chariclea, Schn., in which the
black netting marks were resolved into smeared black lines ;
A, pales, for the first time from this region, precisely similar
to the form taken on the east of the Lena river in Siberia ;
and Caenonympha tiphon closely resembling the form from
Kamtschatka.
The second case contained specimens from a collection made
by a Russian, between Jakutske and Verkhojansk in north-
eastern Siberia at about the same latitude, 67°, as the
preceding exhibit. They included many species which occur in
the western palearctic region, such as Aporia cratxegi, Triphysa
phryne, Cenonympha iphis, Argynnis selene, A. ino, Melitza
phebe, etc., and, most remarkable of all, Veptis lucilla. Also
Parnassius delius, which Mr. Etwes said was the first
(ig @)
Parnassius he had seen from within the arctic circle, and
Colias viluiensis, Mén., an insect peculiar to Siberia, showing
remarkable female aberrant forms. The PRESIDENT observed
that these collections threw a great light on the butterflies
inhabiting the fringe of the holarctic region. .
Mr. C. O. WATERHOUSE gave an account of a nest of a bee,
Trigona collina, recently received from Malacca. The whole
resinous mass weighed 40 lbs. <A section of the nest which
was exhibited showed the cavities in the resin filled with
pollen. The central portion of the nest was constructed of
more waxy material, and contained numerous cells in which
were immature bees. The bees were still alive when the nest
arrived, and among them males as well as workers were
found. Specimens were exhibited, as were also males and
a worker of the much smaller species, Zrigona ruficornis,
Smith, received at the same time from Singapore, and sent by
Mr. H. N. Ridley.
Mr. W. J. Kaye exhibited two drawers containing Danaine,
Ithomiine and Heliconine species from British Guiana, all of
similar coloration, and forming a Miillerian association with
a black hind-wing. He pointed out that possibly the species
had converged, and that some were in advance of others.
The central species was considered to be a Melinxa
Melinxa crameri, a species with a constant pattern and a very
complete black hind-wing. Closely following this Melinwa
mneme (a very abundant and variable species) entered, drawing
in its turn a large number of different species of even widely
different families. A diagrammatic table was shown with the
exhibit, which included the following species: (1) [thomiine,
Melinxa crameri, M. mneme, M. egina, M. n. sp., Ceratinia
veritabilis, Ceratinia, sp., Mechanitis doryssus ; (2) Danaine,
Lycorea ceres, L. pasinuntia; (3) Heliconine, Heliconius
vetustus, H. numata, H. sylvana, KEueides, n. sp.; and (4)
Erycinide, Stalachtis calliope.
Papers.
The following papers were communicated :—
“On the Hypsid Genus Deilemera, Hiibner, by Colonel
CHARLES Swinuogk, M.A., F.L.S.”
vas)
“ An Account of a Collection of Rhopalocera made in the
Anambara Creek in Nigeria, West Africa, by Mr. P. J.
eAreny,7
‘Some Notes on the habits of Vanophyes duriewi,. Lucas, as
observed in Central Spain by Mr. G. C. Cuampton, F.Z.S8., and
Dr. T. A. Coapman, M.D., F.Z.8., with a description of the
larva and pupa by Dr. T. A. CHapman.”
March 4th, 1903.
Professor E. B. Poutton, M.A., D.Sec., F.R.S., President,
in the Chair.
Election of a Fellow.
Mr. Harry Exrrincuam, of Eastgarth, Westoe, South
Shields, was elected a Fellow of the Society.
Exhibitions.
Colonel C. T. BincHam sent for exhibition specimens of
Diptera and two Aculeates from Sikhim, constituting in the
banding of the wings and other characteristics a striking case
of mimicry. The Rey. F. D. Moricr drew attention to the way
in which the fly imitated with its tibia the tarsus of the bee.
Mr. A. J. Curry exhibited specimens of Atomaria rhenana,
K., taken by him out of some flood rubbish found near
Lancing, probably the same locality where the beetle was
discovered formerly by Dr. Sharp. The species is represented
in the Power collection, and it is also mentioned in Canon
Fowler’s ‘ British Coleoptera ” as having been taken at Bognor,
and in Wicken Fen. He also exhibited a Ptinus, apparently
new to Britain, found in a granary in Holborn in 1893,
where it had probably been introduced.
Mr. W. J. Kaye exhibited species of Lepidoptera from
British Guiana, forming a Miillerian Association in which all
but one were day-flying moths, the exception being an Erycinid
butterfly, Lsthemopsis sevicina. The moths, belonging to three
families, included Syntomide ; Agyrta micilia, and Huagra
*
(a5)
celestina, Hypside ; LIostola diwvisa, Geometride (1); Pseudar-
bessa decorata. It appears very evident from the specimens
collected over eighteen months in exactly the same place,
that the Syntomidx in being so numerous have acted as the
types, toward which the other species have converged. The
particular interest of the exhibit consisted in the association
being one of moths, a butterfly being the exception, and not
one of butterflies with perhaps a single moth, which latter is
so frequently the case in South America. The butterfly most
closely resembled Agyrta micilia, one of the Syntomide that is
perhaps the most abundant of all the groups.
Papers.
Mr. C. O. WATERHOUSE read ‘‘ Notes on the nests of Bees of
the Genus Trigona.”’
Mr. G. A. RoTHNney communicated a paper on “The
Aculeate Hymenoptera of Barrackpore, Bengal ;” and another
paper entitled ‘“ Descriptions of eighteen new species of
Larridx and Apidx, from Barrackpore, by Peter Cameron.”
Colonel CHARLES SWINHOE communicated a paper “On the
Aganiide in the British Museum, with descriptions of some
new species.”
Wednesday, March 18th, 1903.
Professor E. B. Poutron, M.A., D.Sc., F.R.S., President,
in the Chair.
Election of Fellows.
Mr. H. W. Betit-Mariey, Durban, Natal, Mr. J. C.
Dotuman, Newton Grove, Bedford Park, W., Mr. W. W.
Row.anps, Lickey Grange, near»Bromsgrove, and Prof. T. H.
Taytor, M.A., The Yorkshire College, Leeds, were elected
Fellows of the Society.
Hehibitions.
The Rey. F. D. Morice exhibited with drawings a dissected
gynandromorphous specimen of a bee (Osmia fulviventris,
Panz.). He said in this species the 9 is normally larger,
(ouaiachen))
less brightly eneous, and altogether darker-looking above
than the ¢. There are also of course the regular sexual
differences between them, which occur in all Anthophila,
as to antenne, number of abdominal segments, possession
of a scopa, genital armature, etc. The clypeus and mandibles
differ also in shape, though not in colour; the last or sixth
dorsal segment is simple (apically incised, as is the seventh
also, In the male) ; and the hind metatarsi are without a little
tooth, which in the ¢ is always present, though generally
quite concealed by longer and very dense hairs. In the
specimen exhibited, the general appearance above was (be-
fore dissection) completely 2; as were also (I believe) both
antenne, though unluckily one has been broken off and lost.
That remaining is obviously 9. The genital armature seems
to be entirely @ also, though not quite in a normal state,
as though somewhat shrivelled and distorted. But the
following parts are certainly ¢: the right mandible; the
right posterior metatarsus; the two last apical (sixth and
seventh) visible dorsal plates of the abdomen ; the right side
only of the third, fourth, and fifth ventral plates of the
abdomen. I think that the first and second ventral plates are
also ? throughout. The last or sixth ventral plate is rather
puzzling. It shows, on the right side only, what looks like
part of the 2 scopa; whereas, in the preceding segments,
the scopa-bearing @ side is, as has been said, the left. Thus,
on the ventral side of the abdomen ¢ characters largely
predominate (the @ with one doubtful exception occurring
only on three segments, and only on the left side of each of
these), The dorsal apex is also ¢ entirely. Yet the genital
armature seems to be completely @, and I can find no trace
of the concealed imperfectly chitinized segments which in
all Hymenoptera ensheath the armature of a 3. In the
head the general appearance, the clypeus, one antenna at
least (¢ both), and the left mandible certainly suggest a
2 :—perhaps too the thorax does so, but one ought to have
studied its sexual modifications more thoroughly than I have
done, to be certain as to this, The insect was kindly sent to
me (with the gynandromorphous Eucera which I exhibited last
month, and several other similar monstrosities) by M. Jean
(Ga)
Vachal of Argentat, France. The species is a common one ;
whether that called fulviventris in the British List is a variety -
of it, or a distinct species, is not yet finally decided. I have
handed the dissections to Prof. Poulton, for presentation to
the Hope Collection in Oxford.
Mr. A. Bacor exhibited a number of specimens of J/ala-
cosoma neustria x castrensis in various stages, including a
series of six ¢ ¢ and sixteen 9 9 imagines reared during
1902 from one batch of ova laid by a 2 Castrensis, which
had been mated with a ¢ Neustria, and two 2 @? reared from
another batch of ova the result of a similar cross ; also blown
larvee of hybrid parentage, and twigs showing attempts at
ovipositing on the part of 9? 9 hybrids that had paired with
hybrid ¢ ¢ of the same brood; also a series of I. wWeustria,
M. Castrensis and the hybrid moths reared during 1901 for
comparison. He said that the larvee of the 1902 broods, as
stated in the Society’s Proceedings for June 4th last, exactly
followed those of the previous year in respect of their division
into “ forwards” and “laggards,” the former again producing
only females, and the latter males. By forcing the pupe of
the laggards it was found possible to synchronize the emer-
gencies this year, and pairings between the hybrid moths
were obtained. The females attempted egg-laying, adopting
the position and motions of normal females of Castrensis, but at
each opening of the ovipositor they produced only the small drop
of cement which accompanies the egg in the normal oviposition
of the parent species, resulting in a more or less perfect spiral
band of cement upon the twigs. The length of time occupied
in producing this result was about the same as that required
by MW. Castrensis 2 for depositing her normal batch of eggs,
which are laid at the rate of about eight or nine a minute.
Pairings or attempted pairings were also obtained between
the hybrid males and females of both the parent species, but
only a very few eggs were laid by these females, who subse-
quently recommenced calling. Perhaps the most interesting
feature of the exhibit was the great variability shown by the
specimens comprising the larger of the 1902 broods compared
with the remarkable uniformity of the hybrid moths reared
during the previous year. Such uniformity appears the more
( ix)
remarkable if we remember the wide range of variability
shown by both the parent species, and that a very wide, if
not the entire local range, may be found within the limits of
a single brood of either species.
Mr. H. St. J. DontstHorre exhibited specimens of 7rimium
brevicorne, Reich., from Chiddingfold, Surrey, an unusually
southern locality for this species,
Mr. C. P, Pickerr specimens of /Hybernia leucophearia and
Phigalia pedaria taken at Chingford on February 14th. With
regard to the resting habit of the former species he said it
was somewhat curious. The bodies lay in a parallel position
to the bark of the trees on which they were found ; the cross-
markings of the wings in an upright position corresponding
closely with the lines of bark. Their colour also harmonized
remarkably well with the surroundings. He also exhibited
ova of Hndromis versicolora on birch twigs, laid March 17th.
The parent moths paired on March 16th at 1.20 p.m., remain-
ing in cop. thirty-three and a half hours, until 10.55 p.m. the
following day. The female then commenced crawling about
the cage to find some suitable place for laying, and on the
introduction of some birch twigs deposited thirty-eight ova
in ten minutes upon them, On the morning of the 18th she
had laid 171 ova, which were cream-coloured and shiny in
appearance, but after two or three days assumed the colour
of the birch twigs upon which they were placed near the
buds. The female in the act of oviposition prefers to rest
head downwards, and sometimes uses the back legs for
arranging the ova.
Mr. G. C. Cuampion exhibited a long series of specimens of
a species of Cneorrhinus (¢ pyriformis, Boh.) from Piedrahita,
Spain, and called attention to the great dissimilarity between
the sexes, and also to the possibility of the females being
dimorphie, one form clothed with green scales, and the other
with grey scales like the male. He also exhibited Dorcadion
dejeani, Chevr., from the Sierra de Bejar, a species peculiar
to that district.
Mr. k. McLacutran, F.R.S., exhibited a dragonfly belonging
to a small species of the genus Orthetrum, attacked by a fly,
almost as large as itself, of the family Asilide, taken in
( x )
Persia in June 1902, by Mr. H. F. Witherby, F.Z.8., the
well-known ornithologist. The fly had inserted its proboscis
at the junction of the head and prothorax, a vulnerable point.
Mr. G. H. Verraxn said that he had seen a moderate-sized
Asilid in Darenth Wood attack an insect much larger than
itself, and that Lord Walsingham had received a species of
the same family which attacked large dragonflies. Mr.
McLacatan also exhibited a female specimen of.a large
Aischnid dragonfly, Hemianax ephippiger, Burm., captured
in a street at Devonport on February 24, 1903, by a tram-
car driver as it was flying about his vehicle. ‘The species
occasionally visits southern Europe in migratory swarms or
sporadically, but is especially African, and its presence at
Devonport in February might possibly be due to the example
having flown on board a vessel off the African coast, and
then becoming lethargic until roused by the unusually high
temperature prevailing in the south of England at that time.
The species has once been observed on the Continent as far
north as Brussels. Mr. F. Mrerrirrerp suggested that there
might be some connection between the appearance of the
insect in England and the reported showers of fine dust
which are generally supposed to have come from the Sahara.
Professor E. B. Poutron, F.R.S., exhibited seasonal forms
of Precis antilope, parent and offspring, bred in 1902 by
Mr. G. A. K. Marshall in South Africa, showing the remark-
able dimorphism of the species, which was especially notice-
able in the protective colouring of the under-side of the dry-
season form as compared with the startling conspicuousness
of the under-side of the wet. Healso exhibited Precis celestina,
captured by Dr. C. A. Wiggins in the Victoria Nyanza region,
with the dry-season form of that species, now taken probably
for the first time. The resemblance of the under-side of the
latter to dead leaves was very marked. Professor Pounton
also showed lantern-slides of the same two species.
Mr. W. J. Lucas exhibited with the lantern a slide showing
the larva of Cossus ligniperda in its gallery in a tree trunk,
Dr. T. A. CoapmMaANn exhibited with the lantern a series of
slides illustrating the life history of Liphyra brassolis, Westw.,
a Queensland species, the larva of which lives in ants’ nests,
( xi)
and feeds upon the ant larve. ‘The imago on emergence from
the pupa is clothed with scales highly distasteful to the ant,
which thus protect it from attack during emergence, and
until such time as it is able to fly, when the scales drop off.
(Cp. Entomologist, xxxv, pp. 1538, 184.)
Papers.
Mr. G. C. Cuampion, F.Z.8., read a paper on “ An Entomo-
logical Excursion to Bejar, Central Spain.”
Mr. Epwarp Saunpers, F.R.S., F.L.8S., communicated a
paper on ‘ Hymenoptera Aculeata collected by the Rev. A. E.
Eaton, M.A., in Madeira and Teneriffe, in the spring of
1902."
Dr. Frepertck A. Dixey, M.A., M.D., read a paper, illus-
trated by lantcrn slides, “On Lepidoptera from the White
Nile, collected by Mr. W. L. S. Loat, F.Z.S.; with further
Notes on Seasonal Dimorphism in Butterflies.” He said that
the collection of butterflies which had been made at intervals
by Mr. Loat during his tenure of office under the Egyptian
Government, was of special interest on account of the accurate
data which accompanied the specimens. Mr. Loat’s collecting
grounds were in the neighbourhood of Kaka, about 11° N. lat. ;
and of Gondokoro, about 6° further south. The meteorologi-
cal conditions at the time of collecting were generally those of
the dry season, though at Kaka the rains were just beginning.
Most of the examples of seasonally dimorphic species belonged
to the “dry-season” phase ; but there were some curious ex-
ceptions. Perhaps the most remarkable of these was Zeracolus
daira, Klug, specimens of which caught in January, during the
height of the dry season, were of the full “ wet-season ” colour-
ing ; while some of those taken at the beginning of the rains
were much “drier.” The large proportion of Pierinx to the
whole number of captures was noticeable, as also was the
general likeness of the whole assemblage to the butterfly fauna
of Aden; the different forms of L. chrysippus, for example,
were found by My. Loat all flying together at the same spot,
just as is the case at Aden. The collection brought to light
no new species ; it contained, however, a single example of
the male of Pinacopteryx venatus, Butl., of which only two
(2 sa)
specimens, including the type, both females, have hitherto been
known to science.
Mr. Loat’s series did not seem to favour the opinion that
had been held that Zeracolus evayore, as described and figured
by Klug, was the dry-season form of 7. yerburit, Swinh. It
appeared from this and other evidence that Mr. G. A. K.
Marshall was right in dissociating the two forms. The weak- -
ness of the reasons given for the contrary view had lately been
pointed out by Col. Yerbury.
With regard to the general question of Seasonal Dimorphism,
a point that deserved notice was the greater intensity and
greater persistence of the cryptic dry-season coloration of the
under surface, which often characterizes the female sex. This
might be illustrated from among Mr. Loat’s specimens ; but
the principle was of wide application, and was operative in
both hemispheres. In the genus TZeracolus especially, the
‘‘wet-season ” female often retained some of the ‘“ dry-season ”
garb, and in certain cases (as in 7. puellaris and 7. phisadia)
the female could scarcely be said to have a ‘“ wet-season ”
phase at all. The significance of these facts lay no doubt in
the special need for protection experienced by the female sex.
Prof. Poulton had lately given strong grounds for believing
that on the whole concealment was a more efticacious means of
defence for moderately distasteful forms than the display of
warning colours, especially when the pursuit was keen ; and
the instances here adduced seemed to show that it might in
some cases be of advantage for the female of a given species to
remain cryptic in the wet season, even though the male should
assume brighter colours with the advent of a more copious
supply of insect life. An interesting parallel with the seasonal
changes in Precis antilope and P. archesia, so carefully worked
out by Mr. Marshall and Prof. Poulton, was furnished by the
Central and South-American Proterpia pyrisitia, Fabr. (a
Pierine form allied to Terias), with what is doubtless its dry-
season phase, ?. gundlachia, Poey. Here, as in Precis, the
dead-leaf appearance of the under surface in the dry-season
form is enhanced by the faleation of the forewings and the
development of “tails.” These changes of shape are found
in the gundlachia form of both sexes, but are intensified in
( ‘xiii )
the female; in the wet-season or proterpia form they are
retained by neither sex, but the under-surface of the female
is duller than that of the male.
The simultaneous occurrence in generally dry localities, such
as Aden, of forms which in other places are associated with
contrasting seasons, was not easy to explain. Prof. Poulton
had shown that in several species of Precis the dry-season form
was larger than the wet, and had on that fact founded the
inference that the dry-season form must have been predeter-
mined in the larval stage. But there was reason to believe
that in many genera, and perhaps even occasionally in Precis,
the assumption of the characteristic seasonal garb was not
determined until a later period—in some cases, the last few
days before emergence from the pupa. If it might be assumed
that the Aden species in question were in a state so sensitive
to meteorological conditions as to respond almost immediately
to a few heavy showers, such as were reported to fall there
not unusually from January to May, the intermixture of
“wet” and ‘‘dry-season,” which in many cases meant an
intermixture of aposematic and cryptic forms, might possibly
be accounted for. This suggestion could only be* verified by
observers on the spot.
Wednesday, April Ist, 1903.
Professor E. B. Poutron, M.A., D.Sec., F.R.S., President,
in the Chair.
Message to the Entomological Society of France.
The PRESIDENT announced that he was going to Paris at
Easter, and proposed that the Society should authorize him to
deliver a greeting to the Entomological Society of France,
before whom he was to read a paper. The meeting acceded
unanimously to the President’s proposal.
Exhibitions.
Mr. M. Jacosy exhibited specimens of Rhagiosoma mada-
gascariensis, Heyd., from Madagascar,and Carpophagus banksix,
PROC. ENT. SOC. LOND., II. 1903. B
(C= xlva.)
McLeay., and Mecynodera coxalgica, Boisd., from Australia.
In appearance they presented many characteristics not usually
associated with phytophagous coleoptera.
Mr. C. P. Pickerr exhibited specimens of Dilina tilix
forced from Essex pup during February and March. In
two @ @ the usual rust-coloured markings on the fore-wings
were abnormally pale, and the hind-wings were black. In
another 2 the rust-red hue pervaded the whole wing area,
the four normal green blotches being a deep reddish-brown,
corresponding with a form of Smerinthus populi frequently
bred. A third ? displayed light-brown hind-wings ; while one
4 was of the normal 2 coloration.
Mr. W. J. Lucas exhibited lantern slides of the specimen of
Hemianax ephippiger, and of the Orthetrwm species attacked
by an Asilid fly, shown by Mr. R. McLachlan at the last
meeting.
Mr. W. J. Kaye exhibited a lantern slide to illustrate the
collecting grounds visited by him in British Guiana.
Papers.
Dr. T, A. CHAPMAN read a paper entitled ‘ Contributions
to the Life History of Orina (Chirysochloa) tristis, var.
smaragdina.”
Sir GeorGe Hampson read a paper on “ Apoprogones hes-
peristis, a remarkable new lepidopterous insect from Zulu-
land.” He said that the genus must be referred to the family
Euschemonide which is represented by the single species
Euschemon rafflesia, Westw., from Australia. In what quarter
of the globe the family originated it was impossible to say,
but the appearance of the species in question suggested that
it was a survival of the scattered remnant of the Antarctic
fauna. It was, however, most remarkable that the genus
should occur in Africa and Australia alone. The PresipENnt
remarked that the wide spread of the Hesperiade indicated a
group to which a local origin could only be assigned with
great doubt. Mr. M. Jacosy mentioned that there were
similarly divided coleopterous genera occurring simultaneously
in South America and South Africa only.
( xv )
Mr. F. Enock gave the following account of ‘‘The Life
History of Cicindela campestris,” illustrated with lantern slides.
“Tt was only after many years’ watching that I observed
the female, boring into the sand at Hampstead Heath, de-
positing a single egg in a tiny burrow one-sixteenth of an
inch wide by a quarter of an inch deep, I was not successful
in rearing the eggs, but from observations found that they
hatched in from a week to ten days—according to the weather,
The young larva is decidedly like the mature one, both in
form and temper, which may be described as ‘nasty.’
“In certain parts of Hampstead, where the sand is free of
moss and grass, I have frequently seen dozens of burrows—
of all sizes—from the one inhabited by a baby larva to that
of a full-grown one, just the diameter of an ordinary pen-
holder. On cautiously approaching the ground the larva may
be seen ‘sitting,’ the head filling up the hole, which is finished
off with a saucer-shaped concavity of three-eighths of an inch
diameter : a mere hollow, just sufficient to cause an unwary
insect to stumble on a quick run, and—fall into the jaws of
the larva.
“To dig these holes wp and turn out the larva is no easy
task, unless the following plan be adopted. Cut a grass
stem a foot long, letting it down the hole until it will go no.
farther ; if the top end is agitated, the other is probably
between the mandibles of the irate larva. Keep hold of the
end, and with a three-fourth-inch half-round file bent round
an inch at the tip, dig away the sand from around the stem,
keeping on until you come to the end, where the larva is seen
sitting, its body turned into a perfect zigzag, which is soon
straightened out on being placed on level ground. The
strange appearance of the creature can then be studied,
with its huge head and receding ‘frons,’ the mandible turn-
ing wp instead of down, the immensely developed mentum,
then the large semi-circular chitinous thoracic plates, followed
by the soft, white flaccid abdomen, until the fifth segment
is reached. This is swollen into a large dorsal hump capped
by two short and powerful vertical spikes ; and in front, but
lower down, two longer ones pointing forward, each having
a somewhat twisted or corkscrew form—the tips diverging.
(xavier)
The remaining segments gradually taper to the anus, each
having tubercles of long spines, those at the anus shorter
and stronger. The legs are decidedly wiry, and the claws
long and sharply pointed.
‘On touching its tail, the larva under observation immedi-
ately turned its head over its back, seized its own tail,
suddenly let it go, and sprang a distance of seven to ten
inches.
‘The larva uses its wedge-shaped head as a shovel, digging
into the sand, which it throws over its back right out of its
way. It soon disappears, though the continual upheaval of
the sand shows that it is still at work. Occasionally there is
a great uprising of sand, followed by the appearance of the
head of the larva, which clears away the accumulated heap.
This it accomplishes by thrusting its head into the mass,
and quickly jerking it over its back, sending the sand flying
a distance of several inches. Many hours are occupied in
excavating the burrow, which sometimes reaches a depth of
ten inches.
“As soon as the required depth has been reached and the
hole cleared out, the larva ascends slightly above the level,
and taking mouthfuls of sand from the edge, hurls them
clear ; and this is continued until a rowgh excavation is made
around the entrance. At this point the larva ascends a little
higher, and hammers all round the burrow with its chin,
until it has beaten the sand into a smooth saucer-shaped
excavation. Satisfied with its labours, it rests therefrom in
the following manner. The head is drawn up level to the
top, just fitting one-half the entrance, the other half being
exactly filled by the first semi-circular plate on the thorax,
thus completely blocking the entrance—the colour of the
head and thorax giving it natural protection.
“ By filling a large bell-glass with sand to within three inches
of the top, and with a cane three-sixteenths of an inch diameter,
I made holes down the sides, so that the glass formed one side.
I then placed a larva in each hole, with the result that some
escaped and tried to enter that of another larva, an act of
trespass for which its life was forfeited. Others made them-
selves at home after trying to plaster the glass with wet sand ;
(i Sarat)
but when this happened—and some kept it up for a fortnight
—I dug them up and started them in a fresh hole, and at
last they adapted themselves to circumstances. I was then
enabled to watch their habits underground. They deepened
the burrow by using their mandibles as picks and their
heads as shovels and elevators, for as they excavated heads
down, they drove their long-pointed claws into the sides above,
and making a larger excavation on one side they passed the
sand above by simply turning up their heads, ramming the
sand past and above their tails; until, having a sufficient
weight or load, the larva reversed its position, put its head
and first thoracic plate underneath, and simply pushed and
pushed until it raised the sand to the ground-level, where a
large heap accumulated, which from time to time was shot
away.
‘“When the desired step was reached and the pitfall set at
the entrance, the larva quietly put its head and thoracic plate
to the top to bar the entrance, the feet were driven into
the sides below the mentum, and the fifth abdominal segment
drawn up until the hump was wedged close up and under the
thorax, bringing the two vertical dorsal spines at right angles to
the burrow, into the sides of which the points were driven; the
remaining abdominal segments were curled round until the anal
spines touched the opposite side, into which they were driven,
thus securely holding the larva, in its peculiar zigzag form,
ready at a moment’s notice to drop down to the bottom of
the burrow by releasing the anal spines, and straightening
the abdomen. Occasionally the larva in its deepening would
meet with a small quartz pebble, which it seized between its
mandibies and carried to the top, where, with a rapid back-
ward movement of its head, it jerked the pebble away a
distance of ten inches.
“Watching the larva through the glass, I pushed a dead fly
near. When it was within five-eighths of an inch it seemed to
suddenly disappear, and appear at the bottom of the burrow
in company of the larva. The movement was almost too
rapid to follow and observe how it was made, but after end-
less watchings I was able to follow its movements. I caught
a Blow Fly, killed it, and tied a thread of floss silk to its leg,
(seve)
then pushed it toward the waiting larva, at the same time keep-
ing the thread ‘taut.’ On reaching to within five-eighths of an
inch of the larva it struck out,and I held on to the thread. The
larva would not let go, and I was able to see that in springing
out the larva had sprung up and backward (just the move-
ment of a gymnast turning a back somersault), throwing all
its legs out clear of the burrow, the abdomen forming a semi-
circular arch between the hole and the seized fly. I gently
pulled my thread, so did the larva, when I was enabled to
see that in throwing itself out of the burrow it had made
a centre of the two vertical spines, which, as the head turned
over, were levered out of the sand; while the points of the
other longer pair in front of the dorsal hump were brought
into contact with the sides of the burrow, into which they
were deeply driven and firmly ‘anchored’ the larva, whilst
it seized its prey. I tried this experiment a number of
times, fully confirming my observations.
“On the approach of cold weather in October the larvze
begin to deepen their burrows, ramming the excavated sand
above them, so closing all entrance and exit. Without food of
any kind the first winter was spent underground. During
March the larva commences to re-excavate the sand above, a
laborious task of biting it down until, in April and May, the
surface was reached, and another year or six months spent
in feeding. For the second time in September or October the
larva retired to the bottom of its burrow, after filling in with
sand above it.
“On the approach of warm weather it once more re-ascended
to the surface, and for a few months feasted upon earwigs,
etc. Several times I found remains of lepidopterous larva.
Once also I observed a White butterfly caught, and on
another occasion a fully-developed Puss moth.
“In August of the third year the larva excavates an oval
chamber at the bottom of the burrow, sometimes in a slanting
position, but more frequently horizontal—an inch and a half
long by three-quarters diameter. The burrow is rammed full
of sand to within an inch of the chamber; the larva then
rests upon its back, but only the thorax dorsal hump touches
the sand, the anal segments being turned up. In a week it
(S26)
changes to a pure white pupa, the dorsal hump disappears,
and in its place a large fleshy process is evolved at each side,
terminated with a brush of strong spines. A similar but
much smaller pair appears on the Ist, 2nd, 3rd and 4th seg-
ments, each process diverging outwards, the brushes of stiff
hairs giving a firm hold, and in this trussed position the pupa
rests safe from contact with the sand.
“In less than a week the limbs become suffused with a
delicate tint of green, which increases in intensity until the
legs are distinctly visible under the pellicle. The mandibles
as well as the eyes are brown. At the end of four weeks
the pupa becomes restive. At last it turns over, frees itself
of the membrane enveloping its whole body, and there appears
a very delicate-looking Tiger Beetle, but so weak that it can-
not bear its own weight. Ina day or two it attains its full
colour and character.
“‘ After this it quietly rests in a semi-torpid condition until
the following April, when it sets out for the small shaft left
by the larva as a final instruction. This it proceeds to enlarge
by biting the sand, which it removes grain by grain until its
labour is ended, and it reaches the surface.”
A discussion followed as to how far the abundance of food
in the larval state affects the development of insects, in which
the Presmpent, Mr. W. E. Swarp, and other Fellows took
part. Mr. Enock said that where the food supply happened to
be insufficient, neuropterous nymphs would continue two years
in that stage, and Mr. C. O. WatERHOUSE mentioned a case
reported to him of the larve of Vanessa urtice which, having
exhausted their summer pabulum, retired to hibernate until
the following year. Mr. A. J. Currry said he had observed
that some coleopterous larve under similar circumstances would
consume flies; while Mr. H. Sr. DonistHoRPE mentioned that
he had bred successfully a species of the same order on paper.
(nex)
May 6th, 1903.
Professor E. B. Poutron, M.A., D.Se., F.R.8., President, in
the Chair.
The PrestpENT announced that he had received a generous
welcome from the Entomological Society of France on the
occasion of his recent visit to Paris. He remarked that the
Entomological Society of London, as now constituted, came into
existence two years after that of France, and exhibited one of
the original invitations to join the Society issued in 1834, and
signed “G. R. Gray, Secretary pro tem.” The Meeting
signified approval that this document should be reproduced
in lithograph, and placed in the Library.
The Prestmpenr also announced that a Suggestion Book
for subjects of discussion and ,exbibition would be placed
in the Library for the use of Fellows.
Exhibitions.
Mr. WititouGHBy GARDNER exhibited nest cells of Osmea
wxanthomelana from Conway, North Wales. He said the
species, one of our rarer mason bees, places its beautifully
constructed pitcher-shaped cells at the roots of grass, usually
four or five together. The bee itself is always scarce in
Britain, and there is no previous record of the nest having
been found since Mr. Waterhouse discovered and described it
from Liverpool about sixty-five years ago.
Mr. M. Jacosy exhibited Avsoa longimana, Fairm., and A.
aranea, from Madagascar, the only other specimens of these
species he knew of being in the British Museum collection.
He also exhibited Megalopus melipona, Bates, and IM. pilipes
from the Amazon, which bore a remarkable resemblance to a
bee, so much so that Bates always mistook it for one when
settled on a twig.
The PrestDENT remarked that two species of Cetoniidve
observed by him in Majorca, in 1900, were very like humble-
bees in their mode of settling or rising from flowers.
Mr. A. J. Currty exhibited a water-beetle new to Britain,
viz. Hydroporus bilineatus, Sturm., discovered by Mr. Edward
Waterhouse among some specimens of Hydroporus given by
( 3eah))
Mr. Chitty to him as H. granularis. The specimens were
taken at Deal in 1891, and probably all records of granwlaris
from Deal relate to this species. He also exhibited a specimen
of the rare 7rechus rivularis (incilis of Dawson), taken at
Wicken Fen in August 1900. The only other British speci-
mens of this insect are believed to be the original specimens
taken at Whittlesea Mere, and some afterwards found by Dr.
Power at Holme Fen.
Mr. O. E. Janson exhibited specimens of Neophxedimus
melaleucus, Fairm., a goliath beetle from Upper Tonkin, and
remarked that the white colouring was derived from a dense
clothing of peculiar semi-transparent coarse scales which were
apparently easily removed by abrasion, and seemed to partake
of the nature of the “fugitive” scales found upon freshly-
emerged specimens of Hemaris and other Lepidoptera.
The PRrEsIDENT read the following communication on “ Pro-
tective Resemblance and other modes of Defence adopted by
the Larve and Pupe of Natal Lepidoptera” from Mr. G. F.
LEIGH.
“ Having devoted a great deal of time during the past two
years to finding and preserving the larvee of Natal Lepidoptera,
it appeared to me that some account of the most striking
examples, as well as of the conclusions to which I have been
led, would be useful.
“ Among the butterflies the larvee of Papilios demodocus, poli-
cenes, brasidas and nireus do not appear to conceal themselves,
and I have never seen birds, hornets or spiders attacking
them. Papilio morania, until the last change of skin, is also
very conspicuous ; but then the larva becomes of a perfectly
green colour, exactly resembling the food-plant, and is very
difficult to detect except during movement. The larva progresses
ina characteristic jerking manner, spinning a web all the time.
Papilio dardanus (cenea) is however quite different: in the
first and second stages the larva is chocolate and white, and
always feeds on the leaves near the ground and among the
undergrowth. During the third and fourth stages, on the
other hand, the larva is green with small white and blue
markings. It is then extremely difficult to find, so closely
does it resemble the colour of the older leaves of its food-plant,
(>-Sie™ ))
upon which it now feeds. These older leaves are blue-green
in colour and covered with white spots. The resemblance of
the pupa to the leaf is so perfect, that I have failed to notice
them upon twigs which I had picked for food, until they
were revealed upon the bare stems from which all the leaves
had been stripped. I believe that Col. Fawcett has described
and illustrated this example of protective resemblance.
“The larva of Charaxes citheron exactly resembles its food-
plant in colour and markings, whereas C’. varanes has yellow
markings, which render it very conspicuous. CC’. ethalion also,
with its one or two black-bordered yellow bands, is conspicuous,
as is C. brutus with its large orange spot. Of all the Charaxes
larve with which I am acquainted none equal C. neanthes in
the perfection of its protective disguise. This larva when at
rest upon the stem—its characteristic position during the last
two stages—is so well concealed by colour and markings that
it is almost impossible to detect. The small green dumpy
pupe of this species exactly resemble some of our berries.
“The larve of Acreas buxtoni, cabira, petrea, natalica,
encedon, and doubledayi, as also that of Limnas chrysippus, use
no disguise, and are certainly unpalatable and avoided by
birds, hornets, spiders, and Mantides.
“Tt is however in the larvee of moths that the cryptic char-
acters are chiefly seen, and, strange as it may appear, most of
all in those that congregate together during the day. J/eta-
nastria aculeata, for instance, exactly resemble the bark of the
tree upon which they rest by day, crawling up to eat the
leaves at night. The larvee of Musgravia leight also vary in
colour—grey, green, yellow or brown—according to the colour
of the bark or lichen upon which the hours of daylight are
passed. Their food is not even supplied by the leaves of the
tree, but by a mistletoe growing upon it. Another species of
Metanastria (as yet unnamed) rests by day upon the trunks of
trees in large patches of one hundred to two hundred larve.
Each patch so exactly resembles the bark that it entirely escapes
notice. In nearly every instance where I have found them it
has been owing to the conspicuous appearance of the indi-
viduals attacked by Ichneumonide, and covered with their small
white larve. When desirous of rearing any of these, I visit
(exalt!)
the patch each morning and remove all the infected individuals.
It is almost impossible to induce these Jarvee to feed in con-
finement, and I conclude their long journey each night is
necessary for their health. The larvee of Matarbela sp. exactly
resemble the droppings of birds, and is also armed with a tele-
scopic appendage that les prone upon the back when at rest.
As soon however as the caterpillar is touched, it whips this
structure backwards and forwards, behaving like the larve of
Cerura vinula, although the appendage is in quite a different
position and placed far more anteriorly. This caterpillar is the
most curious of any I have yet seen. The larva of Homoptera
glaucinans—a green and silver half-looper—almost exactly
resembles the stem of the Acacia upon which it feeds. The
caterpillar of Vephele variegata in its young stages is reddish-
brown, exactly like the young leaves of its food-plant, the
fig-tree. It turns green in its last stage, thus reversing the
history of the colour-changes in almost all the Sphingide. In
spite of its strong cryptic colouring it is one of the most
ichneumoned larve with which I am acquainted. Bunxa
caffraria and belina are most conspicuous larve, and birds will
not eat either, although they are very foud of another species
of the same genus, Buna tyrrhenx, which exactly resembles
the colouring of the leaves of its food-plant. In spite of its
size this caterpillar is so difficult to find that it is necessary to
look for the feces upon the ground, and then carefully to
search the under-side of the branches over the spot. It is inter-
esting to note that the eggs of this species are laid upon the
upper-side of the leaves, whereas those of the other Bunzeas
with freely-exposed larve are always placed upon the under-side.
The caterpillar of Argema mimosx is also very difficult to
detect and exactly resembles its food-plant in colour, although
the beautiful silvery cocoons are very conspicuous upon the
bare twigs of the trees during the winter months. The larve
of Gonometer postica, Metanastria cuneilinea, Jana edulis, Ludia
delagorguet, and L. smilax are all extremely conspicuous, and
sting very badly. They are attacked by ichneumons, but birds,
etc. always avoid them. In fact I do not know any yellow
and black or black and white larve which birds will eat, and
these are by far the commonest colours for larve out here.
(| ty)
Stauropus mediata is another very brilliant larva which
nothing will eat. Its colours are green, bright yellow,
blue and black. The moth is however far from common,
owing to the attacks of Ichneumonide. The larve of
Hublemmistis chlorozonea is also well concealed in the last
stage by the adoption of allocryptic methods. It feeds upon
lichen and is white in colour: when full-fed it completely
covers itself with the lichen, and uses the same to form its
cocoon, which is suspended from the branches or trunk of the
tree upon which its food-plant grows. The moth itself when
at rest exactly resembles the lichen, and is most difficult to
find,
“The larva. of Lophostethus dumolinii is certainly a distinct
case of mimicry, as it is far more likea Saturniid larva than one
of the Sphingide, and does not even rest in the manner usual
among Sphinx larvee. It is greatly persecuted by ichneumons,
and very difficult to rear unless taken full-fed or at any rate
in the last stage.
‘*Ophiusa mormoides and echo, large half-loopers, almost
exactly resemble the bark of the trees upon which they
rest, and their colour varies so as to match that of the
particular tree on which the larve are found. Ophiusa
indeterminata, on the other hand, is conspicuous, and does not
possess the power of colour adjustment, so that three or four
different forms may be found upon the same food-plant : it is
moreover refused by birds. The very common larve of J.
armigera also appears in three or four different forms, which
are relished by birds. These larvee however prefer to feed
concealed inside lilies and the pods of peas. The conspicuous
black and white larva of Diaphone dominica also feeds inside
the thick leaves of lilies, but this garden pest is unfortunately
refused by birds. _
‘‘Geometers I need not refer to, as they are much the same
as at home, only far less numerous.
“There are a great many other larve which might be
mentioned, but, as they produce moths as yet unnamed, I
will leave for another occasion, when the species have been
determined or described.
“The general conclusion may be accepted that black and
( xv)
yellow, and black and white larve are distasteful to birds,
Mantides, hornets, etc., as also are moths, butterflies, beetles,
and flies of these colours in almost every instance.
“The greatest enemies of larve here after birds are Fossorial
wasps and Mantides. I have found nests of the Fossores full
of larvee, generally but a single species ina single nest. Some-
times as many as twelve green geometrid larve may be
found in one nest. Spiders of the family Attide are also
frequently found eating larve. When disturbed the Attid will
jump away, carrying its caterpillar prey with it.
“In conclusion I desire to express my thanks to Sir G. F.
Hampson for naming the moths which I have succeeded in
breeding. A great deal of work still remains to be done in
this direction in Natal. Not one quarter of the larve are
at present known, and yet as a result of the work that is pro-
ceeding, new species of moths are appearing almost every
week.”
The PresipENT said that it was of great interest to hear
the conclusions, as to the general meaning of the colouring
of Natal larvee, reached by one of their Fellows who had
so wide and intimate an experience as Mr. Leigh. It was
interesting to observe that Mr. Leigh considers the larva
of Papilio nireus to be conspicuous, when its pupa possesses a
wonderful power of colour adjustment, as was first shown and
figured by Mrs. M. E. Barker in our Transactions (1874, p.
519). The first account of the larve and pupe of Papilio
dardanus (Merope, cenea) was given by Mr. Mansel Weale
(Trans. Ent. Soc. Lond., 1874, p. 131). The combination of
many larve to producea patch of colour like the bark on which
they rest is very interesting, as also is the “homing” instinct
which leads the larvze to return to the same spot at the end of
each night. It would be useful to experiment in order to
ascertain the cause which enables them to return and collect
together, whether following the clue afforded by a thread or
otherwise. A gregarious habit in order to promote concealment
by the production of a brown patch has been described in the
young larvee of Mania typica (Proc. Zool. Soc. Lond., 1887,
p. 241).
The adjustment of the colours of the larva of JL. leighi to
PROC. ENT, SOC. LOND., III. 1908. C
(xxvii ®)
the trunk of a tree on which it does not feed is a further
and very interesting proof to the many already supplied that
this colour-relation is in no way associated with food.
A careful drawing of the larva of Matarbela sp. and an
investigation of the structure of its dorsal appendage would
be valuable.
When Mr. Leigh considers that the conspicuously-coloured
Heterocerous larve are the most abundant in Natal, is it
not likely that sufficient allowance is not made for the far
greater difficulty in finding the others and for the immense
number of cryptic species of which the larve are as yet
undiscovered !
It is probable that the spines of the larva of Lophostethus
dumolinii are not mimetic of Saturniide, but an indication
of real affinity. All Smerinthine larve and many. other
Sphingidee may be seen by the lens to be covered with spines
during the first stage, and these spines are often forked. A
drawing of the young larva of Macroglossa fuciformis,
magnified a few diameters, would show a most astonishing
form apparently very different from anything that we are
accustomed to regard as a Sphinx. (See Trans. Ent. Soe.
1887, pp. 568-74, together with the references cited. )
Professor PouLron then exhibited the cocoonsof Lublemmistis
chlorozonea sent by Mr. G. F. Leigh in illustration of his paper.
He also showed a specimen of Polygonia (Grapta) C-album
in the attitude of prolonged repose, together with specimens
of Anxa moeris set in different ways to illustrate its probable
resting position, and upon these specimens he read the
following notes :—
“Many years ago I came to a conclusion as to the
probable meaning of the ‘C’ or ‘comma’ on the under
surface of the hind-wings in butterflies belonging to the genus
Polygonia (Grapta). 1 believe that it represents, in bright,
strongly-reflecting ‘body-colour, the light shining through
a semi-circular rent in a fragment of dead leaf,—the rent pro-
duced when a little segment of leaf has broken away along
a curved line, but still remains connected with the rest
across the chord of the arc. Unless such a segment remains
precisely in the plane of the leaf, light may pass through
© xxvil)
a curved and often a semi-circular slit-like window. Such
curved cracks are extremely common in old weather-beaten
dead leaves. They are probably produced by drying and
shrinkage after much wetting and some decay.
“On April 23rd last I had the opportunity of testing how far
the whole attitude of Polygonia C-album, during profound re-
pose, is consistent with the interpretation suggested above. By
a curious coincidence I had been speaking of the differences
between temporary and prolonged resting attitudes in butter-
flies, at the meeting of the Entomological Society of France
on the evening of April 22nd, and the very next morning saw
for the first time in my life the position of this species during
complete repose. The day was excessively cold for this time
of the year, and the butterfly was hanging perfectly torpid
from the horizontal rail of a wood fence in a street at Passy.
Several excellent but very small photographs were taken with
my daughter’s camera: enlargements have been made, and
from these the actual specimen has been set and photographs
of the natural size taken by Mr. Robinson in the Oxford
University Museum.
“The specimen which is now exhibited affixed to a piece of
stick in precisely the same manner in which it hung from
the rail, shows that the two anterior wings are held so far
forward that a deep wedge-shaped notch appears between them
and the hind-wings. On each side of this notch the well-
known ragged outline of the wings is extremely distinct.
The two posterior pairs of legs by which the butterfly clings
to the supporting surface are light-brown in colour and
unexpectedly conspicuous. The antennz are concealed, and
the contour of the head does not break that of the costal
margin of the anterior wings so as to interfere in any way
with the general effect. The whole appearance is consistent
with a single interpretation—concealment effected by resem-
blance to a weather-beaten fragment of dead leaf, deeply
notched and ragged, and hanging by two denuded fibro-vascular
‘veins’ standing out far beyond one of the edges. The kind
of injury suggested by the ‘comma’ only adds another
convincing detail to a perfectly harmonious cryptic effect.
“Tt is interesting to compare this mode of concealment with
(| xxvii)
that which is far commoner in Nymphaline genera (Kallima,
Doleschallia, Anxa, Precis, ete.), viz. the resemblance not to a
fragment but an entire dead leaf, with midrib and suggestion of
lateral oblique venation. In this latter form of disguise, holes
are frequently suggested in the apparent leaf, either by opaque
‘body-colour’ as in Doleschallia, by transparent windows as
in Kallima, or by actual discontinuity, as is probably the case in
certain species of Anwain which the deeply-cut bay in the inner
margin of the fore-wing may be converted into the likeness
of a hole by closure along its open side by the costal margin
of the hind-wing, in the manner indicated in one of the
specimens exhibited. In certain parts of the under surface of
Kallima a hole is suggested by ‘body-colour,’ in other parts
by transparency, and the latter is undoubtedly the more recent
and more highly-specialized method ; for when the transparent
window is examined under the microscope scattered opaque
white scales can still be seen in abundance over its surface,
not thickly placed so as to prevent the passage of light, but
witnesses to an earlier and less perfect representation of light
shining through a hole.
“Tt is interesting to note that the holes represented in these
apparent dead leaves seem to have been produced by gnawing,
whereas in the leaf-fragment suggested by C-album the
forces of the inorganic environment, which by their prolonged
action have produced the wear and tear of the margin,
have also been responsible for the more centrally-placed dis-
continuity. Comparing various species of the genus Polygonia
(Grapta), it is seen that the curved C-like window occurs in
several; in some the suggested rent is V-like, while occasionally
the mark appears to represent a hole of a reniform shape.”
Professor Poutron also exhibited a pair of Hypolimnas
misippus taken “in coitu” on Feb. 3rd, 1903, by Mr. Horace
A. Byatt, B.A. (Lincoln College, Oxford), near his highland
home ata height of 4500 to 5000 feet, in Dedza,Central Angoni-
land, British Central Africa. The specimens are remarkable
in that the female is excessively worn and old, far more so
than the male. Such an observation tends towards the con-
clusion that pairing occurs more than once in the life of an
individual of this species.
( sax 3)
Mr. Byatt writes (Feb. 15th, 1903) concerning the species
—‘‘Close round my house 7. misippus is in vast numbers just
now, but other species are not very numerous. You will
see that at Dedza ZL. chrysippus is quite rare—at least at this
season. I have sent you only two, I think, and my eyes are
always open for it.” This observation of relative abundance
certainly suggests the Miillerian rather than the Batesian
interpretation of the mimicry of the former for the latter
species ; although quite near to Dedza the proportions may be
entirely different. The following passage shows how rapidly
My. Byatt can pass from one kind of area into another. ‘ You
must understand that I have two distinct climates to work in.
T am about 4500 to 5000 feet up—the top of Dedza is 7000
and I drop straight down into what is really tropical Africa
on the lake level : tropical foliage, swamp, damp atmosphere
and intense heat. This station might be in S. Africa
the Orange River Colony. Consequently Lepidoptera are widely
different: in this open country they are fewer and more sober
say
in colouring ; down below they are plentiful, more varied,
larger and more gaily coloured. Pyrecis so far seems to be
below 3000 feet, and extremely rare up here.”
The observations of another friend further to the north in
Africa also show the great abundance of H. misippus and
how far it is from occupying the subordinate numerical
position assigned to mimics by the late H. W. Bates. Between
January 20th and 25th of the present year Mr. C. A. Wiggins
captured at Kisumu, near the terminus of the Uganda line,
on the N.E. shore of Lake Victoria Nyanza, the following
specimens of this model and mimic :—
Limnas chrysippus. Hypolimnas misippus (females).
Type-form 10, Type-form 27.
klugii-form 20. inaria-form 16.
And again at the end of January and the beginning of
February :—
L. chrysippus. H, misippus (females).
Type-form 73. Type-form 18.
Hdugiiform 85. | inaria-form 15.
These results are roughly compiled from the unset specimens,
(7 xx @)
but no serious modification of the proportions is to be expected.
Mr. Wiggins’ very large and interesting series of captures,
bearing upon many bionomic problems of the highest interest,
are now being studied in the Hope Department by Mr. 8. A.
Neave, B.A., of Magdalen College.
Paper.
Mr. G. A. J. RotHney communicated ‘ Descriptions of
twelve new genera and species of Jchneumonide, and three
new species of Ampulex from India,” by Peter Cameron.
Wednesday, June 3rd, 1903.
Professor E. B, Poutron, M.A., D.Sc., F.R.S., President,
in the Chair,
Exhibitions.
Mr. G. C. CHampion exhibited numerous specimens of
Coccinella distincta, taken during the past few days in the
pine woods of Woking. They were found, as usual, running
about the ground in company with Formica rufa, and were
perhaps wanderers from some other locality. Mr. Donts-
THORPE said the species was still common at Weybridge in
the nests of Formica rufa, and that he had observed it
also at Bexhill, while Mr. Cuartrry noted its former occurrence
in Blean Woods in great numbers. The life history of the
larva, he said, had not been worked out. Mr. CHAMPION
suggested that it might pick up stray aphids dropped by the
ant.
Mr. H. Sr. J. DonistHorre exhibited a very remarkable
melanic form of Halyzia 18-quttata, L., black with white
spots, the type, which was also exhibited, being light brown
with white spots. The former was taken at Oxshott on the
22nd May. He also exhibited Stilieus fragilis, Gr., a
melanic form with a black thorax instead of red as in the
type, taken at Shirley on May 15th; and Staphylinus fulvipes,
Scop., taken by himself at Bamber Forest on June Ist, a new
locality for this rare beetle.
(Gazi ®))
Dr. T. A. CHapman exhibited two full-grown larve of
Thestor ballus, sent by Mr. H. Powell, from Hyeres. In its
previous stages the back, except the dorsal line, is bright
yellow, from the metathorax to the sixth abdominal segment,
the ends being reddish, giving the larva a very brilliant
appearance, while the conspicuous black spiracles on the top
of the seventh and eighth (abdominal) segments give very
much the appearance of a mammalian head to these segments.
In this last skin the colours are much darker and more red-
purple, and the posterior spiracles less obvious. The front of
the larva has now much more the aspect of a head, say of a
wild boar, largely owing to the uniform colour, black spiracles,
and depression of the prothoracic plate which characterises the
larva throughout. There are two eversible glands behind the
last pair of spiracles as well as the glandular line across the
preceding segment. The lenticles are very inconspicuous.
The Prestpent remarked that the terrifying appearance
usually occurred in large insects.
Dr. CHapmaN also exhibited a larva of Heterogyna paradoxa,
full fed, reared from the egg at Reigate, and a cocoon of Orgyia
auro-limbata, with parasite Braconid. In this instance a
larva produced an imago and the parasite. The cocoon when
opened last October showed the cocoon of a Braconid within
it ; adense oval-ribbed cocoon of whitish silk, with longitudinal
darker flutings. One compartment of the O. awro-limbata
cocoon was quite empty and flattened, the other contained
larva-skin of Orgyia, pupa-skin of Orgyia,a small shrivelled 9
of Orgyia denuded of wool and containing eggs (perhaps a dozen),
and the microgaster cocoon, which was well coated with and
entangled amongst the loose wool of the moth. The cocoon
looked as if made first and mixed up in the hairs afterwards.
If this be correct, then the microgaster larva emerged from
the pupa, and the moth nevertheless emerged afterwards. As
against this, the cocoon was loosely, if at all, attached to
the eccoon of the moth, as one would expect it to be if it
emerged from the pupa. In that case it must have emerged
from the moth. Im any case, the microgaster and the
moth both came from the same larva, and the moth, though
containing few eggs and (not being fertilised) laying none,
(xo)
was nevertheless energetic enough to denude herself of all
her clothing. An imago and a parasite from the same larva
have not infrequently been recorded, but very often doubt
has been thrown on the occurrence. In this instance this is
impossible, since the apterous female of O. aurolimbata not
only never leaves her cocoon, but never makes an opening in
it, that being done by the male. In this case the cocoon had
no opening in it, and nothing had gone in or out since the
larva spunit. The miserable development of the female showed
that it had suffered some serious malady, and there was
no trace of a second larva having been in the same cocoon.
Some Fellows may like to be assured that the wretched
fragment exhibited is really the imago of a moth. Such as it
is, it is precisely like the ordinary female of O. awrolimbata
after she had laid her eggs and denuded herself of her wool
and died in the cocoon. The only difference is that this
specimen never had any eggs to lay.
The Presipent exhibited the dry form of Precis actia bred
by Mr. Guy A. K. Marshall from an egg laid by a female
of the wet form. The parent was captured by Mr. Marshall
at Salisbury, Mashonaland (5000 ft.), on February 14th, 1903 ;
the egg was laid on the following day. It hatched February
20th, the larva pupated March 16th, the perfect insect, a
male, emerged March 28th. The differences between these
two forms are as astonishing as those between the two phases
of Precis antilope, bred, the dry from the wet, by Mr. Marshall
last year (Trans. Ent. Soc. London, 1902, pp. 418-20). In
fact, upon the upper surface of the wings the differences are
much greater than in this latter species, the dominant colour
upon the black background of the dry form of actia being
blue, as it is in the dry form of sesamus, while the red is of
so deep a shade as to be sombre and inconspicuous. In the
wet form these blue markings are only represented by
marginal, submarginal, and apical traces, while the dull red
becomes a bright and vivid reddish-brown, which forms a
startling contrast with the background and the row of black
spots which crosses both wings. Within these spots, in the
position of the chief blue band of the dry form, the reddish-
brown band of the wet form passes into a brilliant pearly white
@ Fcc)
in the female, into a pale reddish-brown in the male,—forming
in each case a startling contrast with the nearly black basal
half of both wings against which it terminates abruptly.
Intermediate forms are probably much commoner than in
P. sesamus. In one dry male out of four in the Hope Depart-
ment, the chief blue band is in large part replaced by an
intrusion of dull red. The extraordinary differences in shape
are the same as those between the two forms of P. antilope
(compare Fig. 3 on Plate XII of Trans. Ent. Soc. Lond., 1902,
with Figs. 3a and 36), and much greater than those between
the forms of P. sesamus (compare Fig. 1 with la or 2 with 2a
on the same plate). In the dry phases of actia and antilope
the hooked tip of the fore-wing is even more recurved than
in that of sesamus, while the prolongation of the anal angle
of the hind-wing which is so marked a character in the two
former is wanting in the latter species. These characteristic
features in the contour of the wings in the dry antilope and
actia are related to the beautiful and detailed resemblance of
their under-sides to dead leaves, while the greenish-black under-
side of the dry sesamus is well concealed by a general harmony
with the dark shady environment which it seeks for pro-
longed rest, rather than by any detailed special protective
resemblance. Hence the necessity for a profound modification
of shape is far less imperative in this latter, It has been
pointed out that the upper-side differences between the two
phases of actia are greater than in antilope. As regards
their under-sides the reverse is the case, because this surface
is so much less conspicuous in the wet phase of the former
when compared with that of the latter species. It is, however,
very far from being cryptic, attaining nearly the same degree
_of conspicuousness as Precis triment which Mr, MarsHa.y
considers to be another of the wet forms of antilope (/.c. pp.
419, 420). The representation of a dead leaf by the under-
side of the dry acéia is slightly more elaborate than in antilope.
Both species have an equally beautiful mid-rib-like stripe, but
the former alone possesses the representation of two holes, the
posterior minute, near the tip of the simulated leaf—due to
white semi-transparent spots. KEqually elaborate detail is seen
in P. cwama, of which P. triment is the wet form. In Mr,
PROC. ENT. SOC. LOND., Iv. 1903. D
(Saxe)
Marshall’s opinion, however, P. antilope and P. cuama are
two dissimilar dry forms and simia and triment two dissimilar
wet forms of a single species. It is much to be hoped that
the point will soon be settled by breeding.
Mr. Marshall is to be warmly congratulated on this third
South African species of the genus Precis, in which he has
produced incontrovertible evidence of the specific identity
of forms widely separated in colours, patterns, shape, relation
of upper- to under-side, etc., and even instinct, including
the selection of a particular type of country.
The PresIDENT also showed a small series of ants, part of
a much larger collection made by the late W. J. Burchell in
Brazil between the years 1825 and 1850. They were obtained
with his other vast zoological and botanical collections at Rio
or its neighbourhood, or in the course of the long journey from
Santos to Para. Considering their great age the specimens
were wonderfully well preserved and are accompanied by
remarkably exact and detailed data, and, in many cases,
interesting notes on habits, instincts, etc. Hardly anything
in the whole of the zoological material, all of which was pre-
sented by Miss Anna Burchell to the University of Oxford
in 1865, has as yet been published. Arrangements were now
being made to ensure that these interesting results may, with
as little delay as possible, be given to the scientific world.
The PrestpEnt then gave a summary of his paper on the
effect of lichen-covered bark, etc., upon certain Lepidopterous
larve. He explained that these results were now being
brought forward about ten years after the experiments had
been begun. The delay was to be explained by the dis-
organization of the library and papers of the Hope Department
during the building operations in 1894, As a result the notes
of the Professor and Mr. Holland, as well as the beautiful
water-colour drawings made by Mr. P. J. Bayzand, were mis-
laid, and when everything necessary had been recovered the
press of other work for a time prevented this memoir from
being undertaken,
The chief object of the investigation was to test the efliciency
of lichen-covered bark as a stimulus for the production of a
lichen-like appearance in certain larve. It was found that
(oe xxocven)
Gastropacha quercifolia was highly sensitive to such a stimulus
up to the beginning of hybernation, but that during and
after hybernation, all such susceptibility entirely ceased. The
larvee were also sensitive, during the same period, to brown
and black bark, which caused the appearance of corresponding
tints. In all experiments the food made use of was the same,
viz. the leaves of the hawthorn.
Odontopera bidentata was also sensitive to the same sur-
roundings, lichen in the environment producing the well-
known green patches on the dorsal surface.
The rigid restriction of the sensitive period to the earlier
part of larval life in the case of quercifolia suggested a set
of transference experiments on the highly sensitive larva of
Amphidasis betularia, carried out in the summer of 1896. The
result was to prove that this species remains sensitive, at least
to the strongest stimuli, viz. those provided by dark bark, for
nearly the whole of larval life.
Papers.
Mr. O. E. JANson communicated a paper “On the genus
Theodosia, and other Eastern Goliathides, with descriptions of
some new species.”
Colonel C. SwinHor communicated a paper on ‘ New
genera and species of the family Lymantriide in the National
Collection.”
Mr. G. W. Kirkatpy communicated a ‘Memoir on the
Rhynchota collected by Dr. Arthur Willey chiefly in Berara
and Lifu.”
Professor E. B. Poutron read an account of ‘‘ Experiments
in 1893, 1894, and 1896 on the colour-relation between certain
lepidopterous larve and their surroundings, and especially the
effect of lichen-covered bark upon Odontopera bidentata and
Gastropacha quercifolia.”
Wednesday, October 7th, 1903.
Professor E. B. Poutton, M.A., D.Sc., F.R.S., President,
in the Chair.
(e xxoay laa)
Election of Fellows.
Mr. F. M. Lirriter, Althome, High Street, Launceston,
Tasmania; Mr. H. Sware, M.B., Arawa House, Rotorua,
New Zealand ; Col. Jesse Gricas Piucuer, J.M.S., F.R.C.S.,
133 Gloucester Road, Kensington, 8.W.; Mr. 8. A. Nzavs,
B.A., Magdalen College, Oxford; and Mr. C. A. Wiaarns,
Kisuma, Lake Victoria Nyanza, British East African Pro-
tectorate, were elected Fellows of the Society.
Hehibitions.
Mr. G. C. CHampion exhibited on behalf of Professor
Hudson Beare some specimens of a Viptus new to the British
list, captured at Messrs. Horsnaith and Reynolds’ granary,
Strood, on May 11th, 1901. The specimens were found crawling
about on empty sacks inside the building, and were probably
numerous ; the insect being mistaken for another, only a few
were taken.
Mr. C. O. Warteruouse exhibited on behalf of Mr.
Charles Pool specimens of a beetle of the genus Niptus
closely resembling WV. crenatus, but with distinct shoulders,
and more parallel elytra which also are less strongly striated.
They were found in large numbers in a corn-chandler’s at
Edmonton. The insect had no doubt been introduced, but
whence it was impossible to say. It was new to the Museum
collection, and Mr. Warsrnousr had not been able to find —
it described.
Mr. H. Sr. J. DonistHorre exhibited specimens of Apha-
nisticus emarginatus, a beetle new to the British list, from
Parkhurst Forest, where it occurred plentifully this year, and
a Scymnus new to science, from Yarmouth, Isle of Wight.
Mr. M. Burr exhibited a living adult male earwig, Labidura
riparia, Pall., captured near Boscombe at the end of August
1903. He said that the very noticeable pale coloration be-
comes darker after death, sometimes nearly black, which
might account for some of the numerous ‘ colour-varieties.”’
He had observed that the insect used its forceps with great
agility to seize a very active bluebottle which was placed with
it. It greedily devours flies, leaving the chitinous shell and
horny parts sucked dry.
(xxxvil ”)
Dr. Norman Joy exhibited a specimen of Argynnis selene,
taken last year in Berkshire, showing a remarkable tendency
to melanism. He also exhibited Coleoptera taken in the
same county during 1903, including Dinarda dentata, from
a nest of Formica sanguinea at Wellington College; and
Notothecta confusa, Miirk, from the nest of Formica fuliginose
from the same locality. Also Aleochara mexrens, Gyll., from
fungi; Philonthus fuscus, Grav., from a Cossus-infected tree
at Bradfield ; Scopxus sulecicollis, Steph., from a sandpit at
Bradfield; Diplocelus fagi, Gaer., from Streatley ; Apha-
nisticus pusillus, Ol., from Aldermaston ; Colon dentipes, Sall.,
from Bradfield; and later elongatulus, Ol., from an old fir-
stump at Mortimer.
Sir Georce Hampson exhibited a collection of Norwegian
butterflies made by him on the Dovrefjeld, on the Alten fiord,
at Bossekop, and other localities this year, and remarked how
greatly the dates of emergence appeared to differ from those
experienced by Staudinger and other collectors. The speci-
mens included fine series of Colias hecla, Lef., Chrysophanus
hippothie, var. stieberi, Gerh., Qneis norna, Thnb., Melitea,
var. Norvegica, Auriv., the Norwegian form of J/. aurelia,
Argynnis freyqa, Thnb., and A. frigga, Thnb., a Labrador,
arctic, and North American species, now found further south
at Kongsvold for the first time.
Mr. A. H. Jones exhibited examples of Lrebia christi,
taken this summer in the Laquinthal, and of the species of
Erebia to which it is allied; Satyrus actxa, var. cordula,
captured last July at Sierre, having four equal-sized pupilled
eyes on the fore-wings, probably a local form peculiar to this
warm locality ; a short series of Chiysophanus dorilis (type)
and (. var. subalpina from the Laquinthal, with P. hippothée,
var. eurybid, showing the strong resemblance on the upper
surface, which the 9? of this latter species bears to the 9
subalpina.
Mr. A. J. Curry exhibited specimens of a Proctotrupid
which he said approached Ponera constricta, Laty., in appear-
ance, and might be an Jsobrachium. If so, it was new to the
British list. He also showed a specimen of Pieris nape, and
the head of a Bombus, severed from the body when found by
C xxxvin 3)
him, but still alive, and opening and shutting its jaws,
illustrating the manner in which these insects are attacked
by their enemies.
Mr. H. Witiovensy Exuis exhibited Criocephalus polonicus,
Motsch, a Longicorn beetle new to Great. Britain, and also
specimens of all stages from the egg to the imago, to illustrate
the life-history of the species which he explained. The insects
were taken in Scotch fir-trees this year in the New Forest ;
and the fact that the larva was found plentifully seemed to prove
that the colony had been in existence for some considerable
time. He also exhibited specimens of Asemum striatum, L.,
with larva and pupa, accounted heretofore rare in the New
Forest, but this year occurring in abundance.
Mr. AmprosE QUAIL exhibited cases showing the life-history
of some Australian Hepialide.
Dr. D. SHarp, F.R.S., exhibited specimens illustrative of
the egg-cases, and life-histories of eight species of South
African Cassididx as described in a paper by Mr. F. Muir,
and himself. The larvee displayed, with one exception, the
peculiarity of retaining the cast larval skins as accumulations
on the long anal processes with which the larve are provided.
The exception is the larva of Basipta stolida. In this species
the anal tails are more robust and better developed than usual,
but they do not carry the exuvie, and are probably used for
some other purpose. The egg-cases showed a very interesting
series of degrees of perfection; some of them consisting
merely of a few membranes enclosing two or three eggs, and
covered with a patch of excrement; while in the case of
Aspidomorpha punceticosta the odtheca is among the most
remarkable and perfect structures produced by any animals.
Both the larve and eggs are extensively destroyed by parasitic
Hymenoptera.
Mr. W. L. Distant also showed the pupa cases of some
African species of the genus Aspidomorpha, South, with the
cast heads of the larve. Sir Grorce Hampson said that in
the case of some Nolide these cast skins acted as a protection,
the enemy attacking them instead of the insects themselves.
Mr. Rouanp Trimen, F.R.S., exhibited some cases of mimi-
cry between butterflies inhabiting the Kavirondo-Nandi
(cod be 1)
district of the Uganda British Protectorate, particularly that
in which Planema poggei, Dewitz, is imitated by an apparent
variety of Pseudacrxa kiinowit, Dewitz, and also by a hitherto
undescribed form of the polymorphic 2 Papilio merope, Cram.
He mentioned that both Planema poggeit and Pseudacrea
kiinoww were described and figured by Dewitz in 1879 from
single specimens taken by Dr. Pogge in Angola, and added
the interesting fact that the only other example of the unde-
scribed mimicking form of the Q Papilio merope known to
him—in the Hope Department of the Oxford University
Museum—is ticketed ‘‘ Angola; Rogers, 1873.” Now, all
three butterflies had been found as far to the eastward as
Uganda, the Planema apparently not uncommonly, but its
two mimickers very rarely indeed, by Mr. C. W. Hobley, of
Kisumu.
This case wasof special interest because it was the first brought
to notice of the mimicry of one of the Acrvinxe by a 2 Papilio
of the merope-group of the genus, members of the Danaine
genera Amauris and Danais being the known models copied.
Yet it must not be forgotten that the extremely rare form
of 9 of the Abyssinian Pap. antinorii, Oberthiir, named
Ruspine by Kheil, mimicked the moth Aletis helcita more
closely than it did Danais chrysippus ; and the large number
of variations in various directions from the principal and
pronounced mimicking forms of the merope-group indicated
plainly how plastic and adaptable they remain for modification
in any advantageous direction.
Planema pogget was one of the largest species of its genus,
and must be very conspicuous in life owing to the very broad
postmedian transverse rich ochre-yellow band of the fore-
wings in contrast with the white median band of the hind-
wings. The Pseudacrea—which Mr, Trimen referred with
some hesitation to Ps. kiinowii, on account of the different
markings on the under-side of the fore-wings in the premedian
area
except in the narrower white band of its hind-wings. In
Mr. Hobley’s large collection only one example (2) of this
Pseudacrexa occurred, and similarly only one of the undescribed
form of the 9 Pap. merope.
was an excellent mimic of the Planema in every way,
( 3%)
The characters of the last mentioned were the following,
vid. :—
Fore-wing: markings rich deep ochre-yellow,—the sub-apical
bar, disco-cellular streak, and much narrowed, but superiorly
prolonged inner-marginal patch being confluent into a wide
discal band very irregular in outline; some slight fuscous
sealing on yellow band between 2nd and third median nervules
indicates the normally wide separation between the sub-apical
and inner-marginal markings ; ordinary apical spot wanting ;
two small sub-marginal spots respectively above and below
2nd median nervule of the same ochre-yellow as the discal
band. Hind-wing: white patch from base outward much
restricted, barely reaching middle and extending only just
beyond extremity of discoidal cell ; sub-marginal spots ali dull
ochre-yellow except the three next apex which are white.
Under-side: Apex of fore-wing and all outer area of hind-wing
of a paler brown than in the form Hippocoon. Fore-wing :
ochre-yellow bar as above but rather paler, and with scarcely
any fuscous scaling between 2nd and 3rd median nervules ;
an additional sub-marginal very small ochre-yellow spot close
to posterior angle. Hind-wing: restricted white patch as
above, but much duller.
The second example (from Angola) of this form of the 9
merope agreed very nearly with the Uganda specimen above
described, only differing in the rather duller ochre-yellow of
the band in the fore-wings ; in the presence in the same wings
of a very small ochre-yellow apical spot, and of a similar spot
close to the posterior angie on the upper-side ; and on the under-
side the increase of the fuscous scaling about 2nd median
nervule, so as almost to interrupt the yellow discal band.
This made the fourth pronounced known form of the 9
Papilio merope. The usual and generally distributed form
of this sex throughout Tropical Africa was that named Hippo-
coon, by Fabricius—an excellent mimic of Amauris niavius, L. ;
all the other forms appeared to be very rare, and two of them
—Dionysos, Doubl., and the form from Zanzibar described in
Mr. Trimen’s Presidential Address to the Society on January 19,
1898—were not direct mimics of any other butterflies, but were
least divergent from the non-mimetic coloration and pattern
es
(Sexe *)
of the male. The form which he now brought to notice was,
on the contrary, a direct and unmistakable mimic of Planema
pogget; and, as it was inconvenient to refer to the mimetic
forms without assigning names to them, he proposed to style
this form Planemoides.
The PresipENT congratulated Mr. Trimey on the exhibit,
and the special interest attaching to an interpretation of this
remarkable form of the female merope. The Society would
sympathize with the feelings of the author of the original
discovery brought before the scientific world, more than
thirty-five years ago, of the most remarkable of all examples
of mimicry in Rhopalocera, that of merope and its allies,
as he saw before him for the first time this mysterious
form accompanied by its model. At the same time it was
only just to a younger worker, who had been in great part
guided by Mr. Trimen’s classical monographs, to point out
that the interpretation so convincingly illustrated that evening
had been made out last spring by Mr. 8. A. Neave, B.A., of
Magdalen College, Oxford, who had just become a Fellow of
their Society. Mr. Neave had exhibited this form of the
female merope together with Planema pogget as its model at
both soirées of the Royal Society in May and June, a time
when Mr. Trimen’s absence from England unfortunately
prevented him from seeing them. Mr. Neave had shown
at the same time another most striking and interesting
member of the same group, a species of Hlymnias, almost
certainly a form of F. phegea, Faby. He had formed the
group in the course of an investigation into the bionomics of
a very fine set of butterflies from the north-eastern shores of
Lake Victoria Nyanza, presented to the Hope Department by
Mr. C. A. Wiggins. Still later Mr. Wiggins had captured
and presented further consignments, including a small but
very interesting set of specimens from Entebbe. Among the
later series, and especially that from Entebbe, Mr. Neave had
another
found additional members of the same group:
Acreine butterfly, viz. A. auwrivillii, Staud., synaposematic
with the principal model, P/anema poggei; the Pseudacrea,
exhibited by Mr. Trimen, P. kiinowii, Dewitz, var.; and a
second probably undescribed species. All the above are
(xii)
obvious and strongly characterized members of the “ Planema
pogget group”: a more outlying, but apparently distinct
member was found in the female of Precis rawana, Grose-
Smith.
Dr. T, A. CHApMAN exhibited butterflies taken last summer
near Biarritz, and in Spain, in the Logrono Sierra.
These he suggested were probably examples of homceochro-
matism. Little attention has been directed to homeceochro-
matism in European butterflies, and these were certainly not
examples of the detailed mimetism we are now familiar with
in Miillerian groups from the African and neotropical regions.
The first of these sets of associated species has, however,
some of the aspects of a Miillerian group. It contains three
species, which are all on the wing together in very restricted
habitats, into which the numerous other butterflies flying
at the same time in the immediate vicinity do not intrude.
They have the same general appearance and coloration when
flying; a peculiarity which none of the outside species happen
to possess,
They are dark, almost black on the. upper-side, a colour
rather unusual amongst European butterflies, and besides
these three, only found, I think, in Aphantopus hyperantus,
and in Satyrus actea and S. cordula; in the two latter
without the dingy brownish tone of the species exhibited.
The locality is at Guéthary, a most pleasing little watering-
place near Biarritz, and the date July 29-30, 1903.
The three butterflies are Canonympha wdipus—of course
of the southern (western) French form, rather larger and
better marked than the type—J/eteropterus morpheus, and
Satyrus dryas. ‘They are confined to small boggy or swampy
hollows an acre or two in extent—rarely more than five or
six—often a mile or two apart. The two first species do not
leave these hollows, and if by any chance a specimen is driven
a few yards up the surrounding slopes, it at once begins to
work its way back again.
The third species, S. dryas, is extremely abundant in some
of these swamps, and seems to be really equally confined to
them, though its abundance and comparative large size and
powers of flight lead to its making short excursions up the
( xiii)
slopes, and an odd one or two was even seen on the levels
above.
The only other butterflies seen in the bogs at the same time
are comparatively rare—viz. a few Brenthis euphrosyne and
Augiades sylvanus, The country around and above abounds
in various common butterflies ; the common “whites,” Lep-
tidia sinapis, Colias edusa, and C. hyale, several Vanessas,
Satyrus arethusa abundant, and Lpinephele tithonus in swarms ;
Pararge xgeria, Cenonympha arcania, Chrysophanus phleas,
ete. None of these visit the swamps except by rare accident,
C’. edusa, which goes straight across everything, being in fact
almost the only species doing so.
Last year, at St. Jean de Luz, we found S. dryas and
H. morpheus on very similar ground, but C. e@dipus was
wanting. This was in a swamp or bog at nearly sea-level
in the open river valley. C. @dipus appears to require the
protection afforded by the steep slopes, surrounding on all
sides the hollows in which we found it. These hollows are
the upper ends of small streams, perhaps a couple of miles
from the sea, and the little valleys wind enough for one to
say that the swamps are surrounded on all sides by very steep
slopes of 100 to 300 feet, rising to the general low undulating
country behind Guéthary, the highest point of which cannot
be of more than 500 feet elevation.
I have found S. dryas in various localities, generally on
low-lying or shady, and, I imagine, more or less boggy
ground. Here and there, as at Aix, I have met with it where
I thought it abounded on dry slopes, but I suspect it must
really have belonged to damp places adjacent. Nor have I
anywhere else than at Guéthary been so impressed by its
slow, floppy flight. I must confess, however, that I have
usually regarded it as too common a butterfly to be worthy
of much attention.
In the Entomologist’s Monthly Magazine for 1894, p. 221,
Mr. W. E. Nicholson notes the association of these three
butterflies in a marsh near the Lac de la Négresse, near
Biarritz. He places with them S. arethusa and S. statilinus,
not, perhaps, thinking it necessary to mention, what he tells
me is stated, however, in his diary, that these two butterflies
(OP xiv)
occupied drier ground close by. At Guéthary S. arethusa was
abundant on the higher open ground. The Lac de la Négresse
is well sheltered from the sea by an intervening hill.
I do not know, but the European distribution of morpheus
and edipus suggests them as frequent associates, whilst dryas
is probably always common enough to join the group. If not
so rare in the §.E. of France as they are (@dipus is, I believe,
absent), it is certainly not so widely distributed there as
elsewhere. Without forming any theory as to the cause of
the homeochromatic separation of these butterflies from the
other species of the district, it seems difficult to believe that
it is merely fortuitous. Assuming for the sake of argument
that it is an instance of Miillerian association, several points
may be worth noting. The three butterflies are of very much
the same colour on the wing, and no other of the butterflies
flying in the country around at the same time are at all like
them. They keep together, and are not mixed up with their
neighbours. They fly in a slow, floppy manner, and are
captured with the greatest ease. Not to be misunderstood,
I must add that even C’. wdipus can mend its pace at need,
whilst H. morpheus has the capacity for a sudden and almost
mysterious disappearance that many skippers possess, and
S. dryas is, when it likes, a very strong flier.
To associate the three forms together, as differing from
everything else, must be as inevitable to the bird or other
predaceous enemy, as to the entomologist. Equally, however,
it must happen that such enemy, if it were useful to him to
do so, could not possibly fail to discriminate between them
with the greatest facility. S. dryas is at once separated by
its great size, whilst the peculiar hopping, dancing flight of
H. morpheus, in which it shows the colour and even marking
of its under-side, is unmistakable. We have here none of the
detailed agreement of colour and markings that we are now
so familiar with in neotropical groups.
I do not know of any association of European butterflies
that is quite parallel to this one, nor has it before occurred to
me to regard any other association of butterflies I have seen,
as being of homeecchromatic significance, whatever that may
‘be. “ Blues” often swarm together, but they seem to be blue
( oxlv. )
because they can’t help it, and not of any set purpose.
Similarly one finds Chrysophanus virgaurexe and C. var. rutilus
together, and Z'hecla spini and 7’. ilicis, and so on.
The Hrebias are perhaps less accidental: several of the
“grass” Hrebias are generally found together, usually I
imagine fortuitously; but . eriphyle, which I found in
Carinthia established in localities by itself, rarely occurs in
Switzerland except with, and possibly protected by, 2. pharte.
Again a few years ago I found at Guarda (Lower Engadine)
E. pharte and FE, manto, forms usually abundantly distinct,
equally common. on the same ground, and most specimens of
such approximating forms as to require some little care to
’ Erebias are
discriminate them. But all these “ grass’
apparently very closely related. This year I found associated
in Spain two Hrebias that belong to very different sections
of the genus—viz. H. evias and L. stygne.
In the district examined these were always on the same
ground; if one was found you might be sure very soon to
see the other. . evias is an early species, At Digne and
Locarno I have found it in April, #. stygne usually at the
beginning of July. But these dates must be largely a matter
of elevation in each locality. Central European specimens
are certainly rarely taken together, and they differ much
in habits of flight and especially in size. Riihl gives stygne
as 38-42 mm., and evias 45-49 mm., and though both
vary rather more, and evias is usually 50-55 mm., this
fairly represents their ordinary relative size. Our specimens
from Canales and elsewhere present the two species as
very much the same in size, stygne indeed being if anything
the larger and closer together in colour and marking than the
usual Swiss or French specimens.
Evias was perhaps a week earlier than stygne, but
both flew together all the time we observed them, perhaps
about a fortnight. The evias, though not quite agreeing with
description, are no doubt substantially of Zapater’s var.
hispanica, whose leading difference is its smaller size than the
type. The stygne are not so large, or otherwise quite the
same as my var. bejavensis, but are rather intermediate
between the type and that local race.
( xlvi_)
Zapater says nothing of stygne, which was not reported
from Spain, till taken last year by Mrs. Nicholls and by us.
We did not see evias last year when taking stygne, var.
bejarensis. So that the association of the species in Spain is
far from universal. I may remark, by the way, that the
ground where bejarensis was found was very different from
that affording the species this year. Nevertheless, it seems
highly probable that this association has something to do with
Spanish evias being small and stygne larger.
The homeochromatism here consists in these two Lrebias,
so very distinct in their ordinary habitats and belonging to
different sections of the genus, when they oceur in Spain on
the same ground, doing so at the same time, and making a
considerable approach to each other in markings and an almost
absolute coincidence in size. I have placed with them some
bejarensis for comparison. It will be noticed that it goes
much beyond the Canales form in the direction in which that
has varied. Looking at the under-sides, one would say the
2 bejarensis were forms of evias and not of stygne. Might
one suggest that, in Spain, evias attracted stygne and led to
its variation to the form before us, and by this assistance
enabled it to get over the intermediate ground, and to pass
on when dissociated again from evias to the bejarensis form ?
I note some specimens of evias vary very much towards the
form, size, and colour of EZ. zapateri. There is no evidence
known to me of these species ever being associated (zaputert
is much later than evias), so that this resemblance must be
one entirely due to climate and locality.
Erebia stygne in Spain will obviously repay more study ;
there is the curious problem of the large (high level)
and small (low level) forms taken last year at approximately
the same dates, and in adjacent localities near the Picos de
Europa by Mrs. Nicholls. Has association with £. evias
anything to do with this as yet unexplained phenomenon ?
I propose that the present form of Z. stygne be called var.,
or at least local form, Azspanica, as parallel to H. evias, var.
hispanica, It is rather darker and more strongly marked
than the type, and has an average expanse of 48 mm., ranging
from 45 mm. to 51 mm.
( xlvii_ )
Dr. CuapmAN also exhibited living imagines of Crinopteryx
familiella. These had just emerged at Reigate; where they
and their parents, descended from pup brought from Cannes
in March 1901, had lived out of doors during their active
existence, being brought into the house only during their
pupal estivation. This seemed noteworthy in so southern
(Mediterranean) a species. ‘The experiment seemed quite
likely to continue successful for the next generation.
Papers.
Mr, Amprose Quaru read papers “ On the antenne of the
Hepialide,” and “ On Epalxiphora axenana, Theyr.”
Mr. Gitpert J. Arrow read a paper “On the Laparestict
Lamellicorn Coleoptera of Grenada and St. Vincent, West
Indies.”
Mr. Thomas Haroup Taytor, M.A., communicated ‘ Notes
on the Habits of Chironomus (orthocladius) sordidellus.”
Mr. F. Du Cane Gopman, D.C.L., F.R.S., communicated
‘* Descriptions of some new species of Lrycinidx.”
Mr. W. L. Distant communicated “ Additions to the
Rhynchotal Fauna of Central America.”
Dr. D. Sarr, M.A., F.R.S., read a paper ‘‘On the E
cases and Karly Stages of some Cassididx.”
oo-
foto)
Wednesday, October 2ist, 1903.
Professor EH. B. Poutron, M.A., D.Sc., F.R.S. (President),
in the Chair.
Election of a Fellow.
Mr. Montacue Austin Puiuuips, F.R.G.S., F.Z.S., of 22
Petherton Road, Canonbury, N., was elected a Fellow of the
Society.
Kaxhibitions.
Mr. J. H. Keys sent for exhibition a black variety of
Carabus nemoralis, Miill., from Dartmoor, recently recorded
by him in the Entomologist’s Monthly Magazine.
Mr. G. C. Cuampron exhibited a series of Rosalia alpina,
-( xlvii )
Linn., found by himself on old beech-trees at Moncayo, North
Spain, in July last.
Mr. A. J. Cairry exhibited a larva of Drilus flavescens,
taken by Mr. Pencott, the schoolmaster of Eastling, Kent,
near the school buildings.
Dr. T. A. CaapmMan exhibited an album containing several
series of photographs of the development of the embryo within
the egg of Psammotis hyalinalis, taken in 1901-2 by My.
W. H. Hammond of Canterbury, and Mr, W. R. Jeffrey. The
depth of focus obtained, and the consequent amount of detail
exhibited in the photographs was remarkable.
Col. J. W. Yersury exhibited (1.) 4 9 2 of Gastrophilus
nasalis, Linn., taken at Torcross, Devonshire, from the 19th
to the 3lst of August last. He said that as this rare species
differed in a marked degree in its mode of flight, ete., from the
common Horse Bot-fly, Gastrophilus equi, it would be as well
to draw attention to these differences. Gastrophilus equi
when flying round a horse visits as a rule the belly and the
fore legs. The @ carries her ovipositor almost horizontal,
and she looks when on the wing like the lower two-thirds of
the letter Z (L). G. nasalis, on the other hand, carries the
ovipositor tucked under the belly and almost parallel to
the axis of the body, this gives her when on the wing a
peculiar ball-like appearance; G. nasalis too always flies to
the horse’s head. While at Torcross during August and
September 20 Bot-flies, 4 9 92 G. nasalis, and 12 92 9,
4 & 6 of G. equi, were taken round the same horse. The flies
came up wind to the horse, G. equi always appearing under the
belly between the fore and hind legs, while G. nasalis would
appear in the triangle formed by the fore legs, the neck, and
the ground. As a rule, the horse paid no attention to G. equi,
but G. nasalis caused him great alarm. The eggs of G. equi
were in hundreds on the shoulders and fore legs of this cart-
horse, but although the face and nostrils were searched carefully
no signs of eggs or larvee could be found thereon. Exhibiting
(2.) Chersodromia hirta, Walker, Col. Yerbury said these
little Empids were common on the shore near Prawle Point :
some were obtained by sweeping over seaweed, while others
were taken running about over the sand. Although these
(Gexiise ”)
insects were loth to take flight, still they were by no means
easy to catch as they ran very fast, and even when covered
by a glass-bottomed box stuck to the ground rather than run
up the box to the glass ; they were therefore very difficult to
box. Col. Yersury also exhibited (3.) Pamponerus germanicus,
Linn., from Barmouth, taken on the 27th and 30th of June
(1 ¢ and 4 9 9), and from Porthcawl ¢, on the 3rd of the
same month. He said this rare species appeared to be struggling
to keep its place in the British List. Mr. Verrall took it
some years ago in fair numbers at the first-named locality,
while Mr. Dale reported it as taken by Capt. Bloomer at
Bridgend fifty or sixty years ago. The second locality was
probably the same as Capt. Bloomer’s. This insect appeared to
frequent the marram grass on the sand hills,and a ? taken at
Barmouth 27th June was preying on a beetle.
Mr. A. H. Jones exhibited specimens of Melitaa deione from
the Basses Alpes, Jf athalia from the Cévennes, Upper
Engadine, Lago di Loppio near Riva, and JM. var. navarina
from Lugano.
Mr. H. Rownanp-Brown exhibited IM. didyma, M. athalia,
M. deione from Digne ; and M. aurelia, M. parthenie and var.
varia, with M. asteria, from various alpine localities.
Dr. T. A. Coapman exhibited specimens of the same genus
from Spain, and Mr. R. W. Luioyp specimens from the
Engadine, and eastern Switzerland.
A discussion on the probable affinities of the several
named species took place.
Dr, CHapMAN said that he was really very ignorant of these
species, but had come to the conclusion that M. athalia, M.
aurelia and M. parthenie, and even perhaps JM. asteria, were
local or seasonal forms of the same species quite distinct in
some localities, difficult to separate in others. The larve as
described vary very little. The difference of the under-sides
in the specimens examined by him was quite within the
ordinary limits of variation. His own specimens of W/. athaha
taken in Spain appeared to differ but very little from the
typical British form.
Mr. J. W. Turr said he was quite unable to distinguish
specifically between various specimens that had been sent to
PROC. ENT. SOC. LOND., Iv. 1903. E
Cle)
him from Switzerland at various times uuder the names of
Melitxa athalia, M. parthenie and M. aurelia, and he was
inclined to think that almost all the specimens caught in the
Rhone valley proper and the lower lateral valleys branching
therefrom were referable to one species. This, however, he
was unprepared to assert, as I. parthenie was always recorded
as a double-brooded species, and MW. athalia as a single-brooded
one, and only those lepidopterists who could deal with the
life-histories of these two so-called species in these districts
(1) by breeding, (2) by comparison of the eggs, larvee and pupe,
could determine the matter. It was to be remembered that
in the Rennes district (teste Oberthiir) and Havre district
(teste Dupont) typical M. parthenie occurred as a double-
brooded species in April, May and August, with typical J/.
athalia, appearing, practically on the same ground, as a single-
brooded species between these broods, viz. in June. It was also
recorded as occurring similarly in Fontainebleau Forest, and
certainly, in June 1897, he had found typical J/, athalia in
great abundance there, and, in August 1899, two or three
specimens of what might be well considered the double-brooded
parthenie, but this was the extent of his practical experience
with the species in this locality. His doubts as to these
species were, however, most increased by the capture of a large
number of a Jelitxa,in the meadows of Grésy-sur-Aix, in late
July and August, 1894-1900. Here the time of appearance
suggested that the specimens captured must be the second-
hood of AL. parthenie, and this view was further strengthened
by the fact that in early May 1897, Dr. Chapman had taken a
fine large typical J/, athalia in the same place, showing that, at
any rate, J/, athalia occurred here earlier in the year ; but the
July—August specimens of Grésy-sur-Aix, whilst furnishing
some examples with the particular facies that experts associate
with MW, parthenie (and existing in the most marked manner in
the specimens obtained in the higher alps and known as var.
varia), are, on the whole, particularly characteristic JZ. athalia,
and, so far as can be judged, most are quite indistinguish-
able from British and Continental specimens of undoubted J/.
athalia ; nor mustit be overlooked that Dr. Chapman found, as
just noted, a large typical MV. athalia in early May at Grésy,when
(i)
M. parthenie should have occurred, for, at this time, Callophrys
rubi, Syrichthus malve, and the ordinary early spring butterflies
were only just showing there ; it is therefore most difficult to
come to a conclusion on the July—August specimens captured,
and it really becomes a matter of first importance that some
lepidopterist with leisure should stay at Aix-les-Bains one
season long enough to determine whether (1) J. parthenie (as
the species is known at Rennes and Havre) occurs at Greésy-
sur-Aix in April—May, (2) JL parthenie is followed by the
single-brooded MV. athalia of typical form, (3) J. parthenie or
M. athalia produces the second brood which has been found so
abundantly in the district in July—August. The only place
in which he had captured what he considered more or less typical
M. parthenie in spring was at Digne in April 1897, but he un-
fortunately knew nothing about the J/. athalia reputed to occur
at Digne in May—June, or whether a second brood of what
he presumed to be JZ. parthende occurred in July. These points
exhibit the weakness of any scientific conclusions based on the
haphazard way in which British lepidopterists work on the
Continent. We go to the Continent, collect “holiday ” series,
sort them out maybe according to Dr, Lang’s plates, in some
cases put on one side the doubtfuls and fill a cabinet series of
very typical-looking examples. That at least is the general
conclusion one comes to after examining many series in the
cabinets of British collectors. What we want are longer series
of specimens, and longer rests in the same district. As to MW,
berisalensis, which has heen referred to J. athalia, the facies of
three very fine examples from Martigny given Mr. Turr by Mr.
Sloper, had led him to suggest that the insect does not even
appear to belong to the athalia group. His impression was
that it came very close to J/. deione, although at present he
knew too little of both insects, to wholly support Mr. George
Wheeler, who in his recently-published work on ‘“ The Butter-
flies of Switzerland and the Alps of Central Europe,” had
come to the conclusion that berisalensis was a mere Swiss form
of the southern species.
Mr. H. Rowtanp-Brown said that he did not remember
having observed J/. athalia and JM. deione flying together at
Digne in June, but that he had noticed MW. athalia and M.
(Cain)
aurelia on the same slope if not actually overlapping in the
mountains about Susa, Piedmont.
The PRESIDENT; who also exhibited some forms of J/. aurinia
taken by Mr. A. H, Hamm at Basingstoke and elsewhere,
and specimens of MV. athalia, M. didyma, and M. phoebe from
Asia Minor and Persia, continued the discussion, in which
Mr. M, Jacosy and other Fellows joined.
The Presipent inquired of any Fellow present whether it
was the case that the dead leaves upon the ground in tropical
countries tended to warp and curl in the dry season, but lay
flat in the wet. He suggested that the remarkable tendency
in the dry phases of many species of butterflies, with dead-
leaf-like undersides, to develop an elongated and hooked
or bent apex to the fore-wing, and a greatly produced anal
angle to the hind-wing, might thus receive its interpretation.
The development certainly could not be explained by affinity,
occurring as it did in the Nymphaline genera Aallima and
Precis, the Satyrine genus Melanitis, and as Dr. Dixey has
recently shown in the Pierine genus.
The Prestpent further stated that Mr. H. C. Robinson had
informed him that it was certainly the case in northern
Australia that the dry-season dead leaves were warped and the
wet ones flattened.
In reply Mr. W. J. Kaye stated that he had been in
Trinidad in the dry season, and noticed that the dead leaves
were curled and bent, whereas in British Guiana which he
visited in the wet season they were flat like damp blotting-
paper.
Paper.
The Presipent gave an account of a paper by Mr. Anporr H.
THAYER on “ Protective Coloration in its relation to Mimicry,
Common Warning Colour, and Sexual Selection,” and made
comments thereon.
Wednesday, November 4th, 1903.
Professor E. B. Poutton, M.A., D.Sc., F.R.S., President,
in the Chair.
( hii )
Election of Fellows. :
Mr. W. A. Boausr, Wilts and Dorset Bank, Shepton Mallet ;
Mr. G. R. Batpocx, 71 Hertford Road, Lower Edmonton ;
Mr. Rogert Erueripce, Junior, Curator of the Australian
Museum, Sydney, New South Wales; Mr. Cuarites FREncuH,
F.L.S., Government Entomologist, Victoria, Australia; Mr.
J. T. Hoventon, Worksop, Notts; Mr. G, Lykrtn, Junior,
Gisborne, Victoria, Australia; and Mr. Wriii1am Herron,
the Horticultural College, Swanley, Kent, were elected
Fellows of the Society.
The SecreTary announced that in some copies of Part TI
of the Transactions recently issued, the numbers to the
figured species on Plate IX were accidentally omitted, and
that Fellows returning Part III to Messrs. R. Clay & Sons,
Bungay, might have the same printed in without additional
expense,
Exhibitions.
Mr. H. J. Ewes, F.R.S., exhibited a small collection of
butterflies made in July last in Saltdalen, Norway, within the
Arctic Circle, Owing to the continued dull, wet weather and
extreme lateness of the season he had not procured some of
the arctic fjeld butterflies which occur there, such as Colias
hecla, C'. werdandi and Lycena aquilo.
He said that it was remarkable to find such plants as Cypre-
pedium caleeolus, and such butterflies as Pararge mxra in this
high northern latitude close to such arctic insects as Hrebia
disa, of which he took a fine series on the one fine day that
occurred during his stay. He also took a series of Cartero-
cephalus sylvius, which, so far as he knew, had been taken no-
where else in Norway. Pieris rape was in the female sex
extremely variable, here some of them, though much suffused,
being more like Scotch specimens than the var. bryonix which
is found in the Alps and in the extreme north of Norway.
Mr. A. J. Cutrry exhibited living specimens of Anthribus
albinus, Showing the way in which this beetle mimics its sur-
roundings. The resemblance to the lichened bark was most
striking, the species choosing the inside holes of hazel twigs
intertwined in hedges, and generally covered with lichen.
@ hy 4
Mr. J. W. Turr exhibited a number of series of the genus
Melitza to illustrate his remarks made at the last meeting.
The discussion on the affinities of the several named species
was continued, and in reply to a question by Mr. Turr, Mr.
H. J. Etwes said that so far as he knew no systematic
examination of the genitalia had yet been made. The
PRESIDENT suggested that the real physiological test by which to
determine and differentiate the several species of the genus
was by breeding, and he impressed upon entomologists resident
in favourable localities, as in Switzerland, the desirability of
making these experiments,
Mr. H. J. Etwes mentioned that he was at present engaged
in the classification and arrangement of the Meliteas and
Argynnids in the British Museum, and appealed to collectors
to bring their series there to be looked over, and to present such
specimens as might be useful for the completion of the group.
The Prestpenr exhibited a set of 323 butterflies from
British Guiana, all captured on one day, August 28th,
1903, between the 9th and 10th mile from the Potaro
River to the gold-mines. The road starts from the
river-side at a point about 30 miles above the confluence
with the LEssequibo. This opportunity of studying the
proportions of the various constituents of the Miillerian
group was owing to the kindness of Mr. W. J. Kaye. The
specimens constituted the entire catch of a single day, and
all were taken by the road-side, on the white blossoms of a
large-leaved plant which springs up wherever the bush is
cut down. The catch represented a full day’s work. August
28th was a particularly dry day in one of the driest months
in the year. The butterflies were most plentiful from 6 to
11 am. and from 3 to 6 p.m., retiring into the thick bush
during the hottest part of the day.
The dominance of the black-hind-winged group is seen in the
fact that it included no less than 295 specimens belonging to
the following species :—
ITHOMIIN®.
Melinxa mneme—253. Mechanitis polymnia—9.
> erameri—8, n. sp.—10.
be)
» egina—d9.
CMa)
DANAIN.
Lycorea ceres—1.
- pasinuntia—3.
HELICONINE.
Heliconius vetustus—1.
Eueides, n. sp.—l.
Thus a single species, Jf, mneme, entirely dominates the
group. Beautiful series exhibiting transition from the barred
to the black-hind-wing were seen in this species, and in the
3 individuals of LZ. pasinuntia. The single ZL. czres was an
intermediate example. One specimen of Melinza crameri
was broadly barred, and another faintly so. In Mr. Kaye’s
experience this species had hitherto always been black.
Mechanitis n. sp. included a fine transitional series, but, as
in other examples from this district, the black mark-
ings were very heavy even in the lightest forms. Mechanitis
polymnia was, as usual, an antithesis to the last-named, the
blackest hind wings being still distinctly barred.
Comparing these and other specimens in the Hope Depart-
ment, and Mr. Kaye’s fine series, with Oxford specimens of
the same or representative species from Surinam, it appears
certain that the Potaro district must stand on the fringe of
the area where this black-hind-winged group is developing. The
ancestral barred pattern and the various grades of intermediates
which occur so abundantly with the black on the Potaro road
are apparently far less common in Surinam, and are probably
less common still in French Guiana. We do not, however,
know the distance to which the group extends along the coast
or into the interior. The apparent anomaly of the dominant
Melinxa mneme exhibiting the most ancestral series of any
species in the group may be merelya result of this position on
the fringe of the area. Extended observations are greatly
needed ; for, so far as it is possible to judge from the facts
before us, more could be learnt of the origin and more inferred
as to the bionomic significance of this black-hind-winged
group of the Guianas than any other in the world.
The remaining specimens, with the exception of a single
Hesperiid, were all Jthomiine. They included 5 Scada theaphia,
( lvi )
but no other member of its group; | Jthomia zarepha, but in
this case also no other member of the group; 16 ceratinia
vallonia, 1 Napeogenes pheranthes, another obvious member of
the same group, and 4 ceratinia bari, a more outlying member.
The Hesperiid Hesperia syrichthus was the only butterfly out
of the 323 which did not fall into one of the Ithomiine
combinations,
Mr. J. C. KersHaw communicated a note on the larva and
pupa of Clerome ewmeus, Drury, as follows :—
‘“No recent description has, I believe, been published on
the habits of Clerome ewmeus, Drury, and, so far as I am
aware, the larva has never been figured or described.
“For the following references to the literature on this
insect I am indebted to Mr. Francis A. Heron :—Pupa
described by J. J. Walker in Trans. Ent. Soe., 1895, p. 450 ;
Drury’s ‘Illus. Exotic Ent.,’ vol. i (1773), and Cramer’s
‘Pap. Exot.,’ vol. 11 (1777), Trans. Ent. Soc., 1858, p. 183.
The larva is smooth, cylindrical, Indian or salmon-red on the
entire upper surface, the articulations of the segments marked
in black. Sides and back sprinkled with whitish hairs. Under
surface and legs black. Head black, upper part of clypeus
bifid. About 1} inches in length when full-grown. The
spiracles indistinctly marked. The larva feeds on Smilax
lancexfolia, 2 common climbing plant in South China, growing
as a rule amongst thick jungle—a smooth-stemmed creeper,
with large lanceolate shiny green leaves, broad and rounded
at the base, of thick texture and glaucous beneath.
“The larve usually feed in little colonies, and have a habit —
when not feeding of resting side by side on the under surface
of a leaf, perhaps half-a-dozen together, so closely packed as
to touch one another.
“The pupa is smooth, of a light pea-green, the apex of
abdomen, from which the pupa is slung without a girdle,
blue bifid, tips yellow. The insect is more or less on the wing
throughout the year, having several broods, some emerging in
January and February, the larvee feeding in November and
December.
‘““Clerome ewmeus is very common in the districts round
Hongkong and Macao, and may sometimes be seen in the
( Ivii )
streets of those towns. It haunts damp and shady localities,
flitting between the tree-trunks and undergrowth, and often
settling on the ground amongst dead leaves. It is to some
extent crepuscular, though it is on the wing throughout the
day in shady spots. There is no noticeable variation in the
insect between the wet and dry seasons, The butterfly appears
to be distasteful to birds and lizards ; though very common in
parts of South China and fairly conspicuous both flying and at
rest, when it closes its wings, I have never seen it attacked.
Its flight, though slow and rather flabby—the wings of this
butterfly are what may perhaps be termed ‘limp ’—is erratic,
and it delights in threading its way through thick cover,
being a wandering insect, and seldom returning to the same
resting-place after once leaving it.”
Papers.
Mr. W. J. Kaye contributed “A Catalogue of the Lepi-
doptera-Rhopalocera of Trinidad, with an appendix by G,. L.
Guppy.”
Mr. P. I. Laruy, F.Z.S., communicated a paper “On some
Aberrations of Lepidoptera.”
Wednesday, November 18th, 1903.
Professor E. B, Poutton, M.A., D.Sc., F.R.S., President, in
the Chair.
Election of Fellows.
Mr. Jonn Rowtianpd Carrie, of Nettleton Manor, Caistor,
and 59 Chancery Lane, E.C., and Mr. E. J. Hare, of 8
Hillsboro’ Road, East Dulwich, S.E., were elected Fellows of
the Society.
Obituary.
Mr. G. C. Caampion announced the death of Mr. Pariip
Brookes Mason, M.R.C.S., F.L.8., one of the oldest members
of the Society, and on the motion of Canon W. W. Fow ter,
seconded by Mr. Cuampton, it was unanimously resolved to
express on behalf of the Society sincere sympathy with Mrs.
Mason in her bereavement,
( lviii)
Nomination of Officers and Council for 1904.
Mr. H. Goss, one of the Secretaries, then announced that
the Council had nominated the following Fellows as Officers
and members of the Council for 1904, in accordance with the
new Bye-laws :—President, Professor E. B. Poulton, M.A.,
D.Se., F.R.S. Treasurer, R. McLachlan, F.R.S., F.LS.
Secretaries, H. Goss, F.L.S., and H. Rowland-Brown, M.A.
Librarian, G.C. Champion, F.Z.8. Other Members of the Council,
Colonel C. T. Bingham, F.Z.S., Dr. T. A. Chapman, M.D.,
BZ... A.J. Chitty, MAS J. Hy Collin, Dro Wa AS Dixey,
M.A., M.D., Hamilton C. J. Druce, F.Z.S., W. J. Lucas, B.A.,
the Rev. F. D. Morice, M.A., the Hon. N. Charles Rothschild,
MEAS) hi S28 254) Dr. Da Sharp.. MEAG Heh ee balnos.
Colonel C. Swinhoe, M.A., F.L.S., F.Z.S., and Colonel J. W.
Yerbury, R.A., F.Z.S.
Hehibitions.
Mr. G. C, Cuampion exhibited numerous specimens of both
sexes of Vyleborus dispar, from Moneayo, Spain, taken out of
beech-stumps. Canon Fow er said that the late Miss Ormerod,
who found this beetle abundantly in orchards at Evesham, had
prophesied that it would become a pest. Asa matter of fact
the insect is now extremely rare, and there appear to be
hardly any Qs in the collections of the United Kingdom.
Mr. O. E, JANSEN said he had found it in pear-trees.
Mr. F. B. Jennines exhibited (1.) on behalf of Mr. H.
Britten, of Great Salkeld, Cumberland, a specimen of Z'ropi-
phorus tomentosus, Marsh., from Great Salkeld, showing the
deciduous false mandibles intact ; (2.) a 9 specimen of Ancho-
menus parumpunctatus, F., from the same locality, showing a
malformation of the middle right tibia which was abnormally
thin, and bent in the centre, but thickened at the base; the
right antenna also had the last seven joints flattened and
dilated. Myr. Jenninas also exhibited on his own behalf
Apion sanguineum, De G., taken at Brandon, Suffolk, in
August last, on Rumex. Commenting on (2) the Rev. F. D.
Morice remarked that similar malformations occurred in
Hymenoptera. The abdomen in the specimen exhibited
appeared to be normal, but he had found in hymenopterous
(dlike %))
insects, segments of the abdomen distorted in such a manner
as hardly appeared consistent with life. Mr. JENNINGS
said that some years ago he had taken a weevil, Sitones
regentsteinensis, with a double club to one antenna, and
Mr. M. Burr said that it was not uncommon to find mal-
formations in the forceps of Earwigs which were not also
hermaphrodites.
Later in the discussion, replying to Mr. Morice and Dr.
CHAPMAN, the President said that during the past summer he
had been experimenting on the eyes of the larvee of Hnnomos
autumnaria. In the attempt to ascertain the physiological
significance of the eyes, some of these larve had been blinded
with a photographic varnish rendered opaque with lamp-black.
It seemed impossible to imagine a more innocent material,
and furthermore the application was but of short duration,
for the varnish did not adhere well to the smooth chitin,
and was soon rubbed off—probably an accidental result of
the ordinary movements of the larve. Nevertheless, when
the corresponding imagines emerged the speaker was intensely
surprised to find that the majority of them were devoid of
eyes, and that the antenne were generally rudimentary. He
could only suppose that something in the varnish, perhaps
the spirit, penetrated pores in the chitin and injured the
subjacent tissues.
Mr. H. St. J. K. DonistHorre exhibited Apion sorbi, 2,
taken this year at Freshwater, Isle of Wight, and said that
the ¢ of this species was extremely rare.
Mr. M. Burr exhibited two 9s and two ds of the largest
known earwig, Anisolobis colossea, Dohrn., from New South
Wales, representing the extremes of size, the average size
being between these two extremes.
Mr. A. J. Currry exhibited a specimen of the beetle
Homalium testaceum taken in Blean Wood in 1900, and a pair
of bees Nomada qguttulata, of which the ¢ has never been
recorded hitherto in Britain, taken by him at Huntingford,
Kent, in May last.
Dr. Norman Joy exhibited (1.) Huconnus Miklini, Mannerh.,
taken at Bradfield in July 1901, new to the British list of
Coleoptera, and (2.) a series of beetles taken at Bradfield at the
( Ix )
exuding sap of trees attacked by Cossus ligniperda; includ-
ing Philonthus fuscus, Grav.; Thamiarea hospita, Mirk. ;
Thalycra sericea, Sturm. ; Cryptarcha strigata, F.; Cryptarcha
imperialis, F.; a series of Epurxa 10-guttata, F., and £.
diffusa, Bris.; and some apparently intermediate forms ;
Quedius ventralis, Ar.; and Atomaria elongatula, Ky.
Colonel J. W. Yrersury exhibited (1.) ZLeptopa filiformis,
Zett. He said that as this insect stood in italics in the
last edition of Verrall’s List, the confirmation of its occur-
rence within the British Isles was worthy of note. Haliday
(‘ Entomological Magazine,’ Vol. iv. p. 150, 1837) described
it under the name of Cordylura flava, from specimens taken at
Holywood near Belfast in the month of June, and its record
as a British insect probably rests on this, for no subsequent
account of its occurrence can be traced, nor do specimens of it
exist in any of our great collections. Leptopa filiformis was
not uncommon at Porthcawl from May 26th to June 4th of
this year; it frequented meadow-sweet under the shade of
some old poplars, where the capture of a chance specimen
led to a thorough overhaul of the neighbourhood both by
searching and sweeping, resulting in a bag of twelve
specimens (10 ¢ and 2 9). Though a shade-loving insect,
it was only to be caught between the hours of 12 noon and
2 p.m., many attempts made both earlier and later in the
day being alike unsuccessful. Exhibiting (2.) Thyreophora
Jurcata, F., Col. Yersury said that though probably not an
uncommon insect, this was one of the least known of our
native Diptera; it stood in italics in the first edition of
Verrall’s List, but was confirmed from specimens taken in
Mount Edgecumbe Park (17th and 24th April, 1889), on a
dead donkey. The late Dr. Meade (E. M. M., Oct. 1889, p. 224)
referred to a specimen found by the Rey. L. Jenyns near Ely,
and to this specimen its reputed occurrence in the British
Isles was probably due. Dr. Meade was apparently ignorant
of its subsequent capture, though the three specimens referred
to above had been in the British Museum collection since
1893. The attractions for Thyreophora furcata appear to be an
open space and the carease of a large animal; it occurred in
some numbers on the Kenfig Sand Hills (Porthcawl) from 13th
(eli)
to 18th May, 1903, on the carcass of a cart-mare ; 7 oy 2
being taken between those dates. In Verrall’s List this species
was included among the Cordyluridx, but as its affinities
appeared to be with the Helomyzidex near Blepharoptera modesta,
probably the best plan would be to give it a family heading
Thyreophoride all to itself. Hendel (Zeit. f. Hym. u, Dipr.
iii, Jahrg Heft. iv, p. 215, July 1903) had recently pro-
posed the new genus “ Centrophlebomyia ” for this species, and
with reason, for the two species Thyreophora furcata and 7’
cynophila seemed to be generically distinct. In our present
knowledge of diptera, where genera could only be looked upon
as artificial groups got together for the purpose of facilitating
identification, this separation of single species from numerically
weak genera was of doubtful utility.
As regards collecting 7’. furcata, it might be pointed out
that a stern sense of duty was necessary to keep one for any
length of time in the neighbourhood of its haunts.
Colonel YErsury also exhibited 13 Pelidnoptera nigripennis,
F’., and said that although only two or three records existed
of the occurrence of this species within the British Isles, still
it was very abundant at Porthcawl from 24th May to 3rd June
of this year. The species was first recorded as British by Mr.
Verrall (E. M. M., July 1894, p. 145) from two specimens taken
as follows: Dolgelly, June 13th, 1887, and Muchalls, near
Aberdeen, June 4th, 1884; a third specimen taken May 17th,
1893, has been in the B. M. collection since 1893. In April
1899 (E. M. M., p. 102), under the heading ‘“ British Diptera
unrecorded or undescribed by English Authors,” Dr. Meade
recorded a specimen taken by Miss Prescott-Decie at Chagford,
Devon, in 1881. He had apparently overlooked Verrall’s
previous record, while it is probable that he was ignorant of
the existence of the third specimen referred to above. Of
(4) Lucina fasciata, also shown by him, Col. YERBury said
that this was another rare insect which had almost been lost
sight of. Recently however Mr. Lamb had met with it in
some numbers at Padstow and Weston-super-Mare, while it
was stated still to exist in Haliday’s old locality, Portmadoc
near Dublin. The specimen exhibited was taken at Porthcawl
(11th May, 1903), seated on a blade of marram grass, but
( Ixii_ )
neither searching nor sweeping produced further specimens,
though the attempt was repeated again and again.
Dr. T. A. CHapman exhibited specimens of Chiysophanus
philxas—(1.) a ? captured at Reigate; small spots and narrow
hind margin to fore-wing, broad copper band to hind-wing
running up the veins, the copper bright and rather pale.
(2.) 15 specimens reared from eggs laid by this 9, and kept at
a temperature of 85° Fahr. (egg and larva), rising to 95° for
the pupe. These were a little smaller than the parent. The
black spots were much enlarged, forming in some specimens
an antemarginal band by confluence, and much restricting
the copper area. Hind-margin broader than in the parent, but
still not broad. Copper of hind-wing more or less reduced, in
one specimen almost to four or five spots. The tails in nearly
all specimens were very long; the copper bright, somewhat
suffused basally, but in only two or three was it suffused on
the apical half of the wing. The dark costal shade almost
wanting in the parent was marked, and to the series of three
most apical spots two more were added between the more
costal veins ; of these the parent had no trace. All the spots
were in most specimens very well defined—three specimens
whose pup were kept very damp are not distinguishable from
the others whose pupe were kept very dry. The exhibit also
included (3.) three specimens taken at Locarno in May 1902,
just after a very cold, wet spell, that affected the whole district.
These had the spots small; the copper area large. One large
specimen (34 mm.) was reminiscent of the Lapland form
which Staudinger identifies with the American representative
hypophixas. (4.) Three specimens from Locarno, April 1903,
after ordinary weather, one of which, with the copper a
little reduced, might be an ordinary British specimen. The
other two had the spots more or less confluent, the margins of
the fcre-wing are very wide, leaving very little copper outside
the spots. The tails also were short and there was little
suffusion. (5.) 16 specimens from various parts of Spain
illustrating various forms of suffusion and darkening, but
none perhaps typical eleus. The darker ones differed from (2)
and (4) in the spots being ill-defined, surrounded by a ring of
less deep black (suffused), and in the black hind-margin of the
(baie)
fore-wing being very broad. In few were the tails as well-
developed as in (2). In the darkest the copper still preserved
some brightness along the discal cell and the two cells beyond
it, whilst two or three might have been ordinary English
specimens.
Mr. G. J. Arrow showed specimens and diagrams in
illustration of a point mentioned in his paper on “The
Laparostict Lamellicorn Coleoptera of St. Vincent and
Grenada.” This had reference to a remarkable kind of
variability noticed in beetles of the 'l'rogid genus Acantho-
cerus. These beetles have the faculty of rolling themselves
into a ball in the interior of which all the vulnerable parts are
enclosed. The head forms a large triangular plate in which
the eyes appear half on the upper and half on the lower
surface. In some examples of the species exhibited (Acantho-
cerus relucens, Bates) the upper division of the eyes forms a
large, nearly circular mass, while in others it is reduced to
a mere thin vestige, and in extreme examples of another
species of the genus Mr. Arrow had even found it to vanish
altogether.
The Presipent showed an exhibit sent by Mr. A. H. Thayer,
of Monadnock, N.H., U.S.A. The greyish silhouettes of two
butterflies were represeuted in a tint nearly the same as the
background, but sufficiently distinct to be easily recognizable.
On one side of one silhouette a row of white spots had been
placed in a submarginal position. It was evident that the
adjacent border was thereby rendered far less distinct than
that of the opposite side of the silhouette, or of both sides of
the other silhouette. The spots in position and shape were
approximately as in Papilio polydamas, and Mr. Thayer con-
sidered they possessed a similar significance in this butterfly.
The dark ground-colour of many Rhopalocerous species he
thought represented shadow under vegetation, the white sub-
marginal lines and dots a generalization of flowers and flower-
masses, But these markings also had a second meaning in
that they tended to obliterate the tell-tale margin of the wings.
Professor Pounron also exhibited specimens of Drurya
antimachus, together with the butterflies which he suggested
as forming a group synaposematic with it. The central species
(“Ixy >)
appeared to be Acrwa egina, round which clustered a number
of other species of the same genus so much alike as to be
probably indistinguishable upon the wing. Examples of these
were exhibited, viz. A. zetes, perenna, rogerst, and pharsalus.
Another beautiful Papilionine member of the group, P.
ridleyanus, was also shown. Its pattern, in both sexes, was
nearest to that of the male A. egina. In fact, so close was
the resemblance that Godart had been entirely misled by it,
and had described the Papilio under the name of ztdora as
the female of Acrea eyina. My. Roland Trimen, F.R.S., had
recently called the speaker’s attention to this, and had in-
formed him that the specimen of the Papilio in the Dufresne
Collection at Edinburgh bears the MS. label “ z¢dora, fem.,
Egina, Cram.,” probably in Godart’s handwriting. Godart’s
mistake had been recently pointed out by Mr. Percy H.
Grimshaw, and Mr. Trimen had himself recognized it from
Godart’s description, and had made a note of it in his copy
of the work.
An obvious Nymphaline member of the group was Pseud-
acrea boisduvalii, the male, like the last-named Papilio,
resembling most closely the male of Acrea egina. It was the
under-side of the female Psewdacrea which first suggested to
the speaker the idea that antimachus was a member of the
same group. While the upper-side of the Papilio seemed
obviously mimetic of the male of Acraa egina, the under-side
of its hind-wings possessed a remarkable and characteristic
ochreous ground-eolour distinguishing it from any other
member of the group except the female Psewdacrexa, in which
a distinct resemblance was manifest. That the approach has
been from the side of the latter seemed clear on comparing the
female of the western form with that of its south-eastern close
ally Pseudacrea trimeni, in which no trac of this peculiar
tint was to be found. Such deutero-synaposematic resemblance
between these two mimics of the egina type of colouring and
pattern had been doubtless encouraged by the fact that they
were the two largest members of the whole group, the female
Pseudacrxa serving as a link between the immense Papilio and
the comparatively small but dominant and central Acrzine
members. The inclusion of antimachus, in spite of its size, in
a.
( lxv ¥)
this powerful combination seemed more satisfactory than Mr.
Trimen’s supposition in 1868 that it “is possibly an instance
of special modification in imitation of some gigantic Acrexa as
yet unknown, or perhaps extinct”? (Trans. Linn. Soc., vol.
xxvi, 1870, p. 503). Professor Poutron suggested that it was
possible that the remarkable bluish-grey patches on the under-
side of the fore-wing of antimachus were traces of descent from
an ancestor common to it and the other equally extraordinary
and equally isolated species of the genus Drurya,—D. zalmoxis.
In the discussion which followed the exhibit Professor
PouLton suggested that the struggle for existence against the
attacks of young, inexperienced enemies,—the kind of selective
attack to which ex hypothesi Miillerian (synaposematic) resem-
blance was due—was in reality far more severe than appeared
at first sight because of the pressure of the struggle upon the
enemies themselves. This pressure was chiefly felt by the young,
and it was so excessive that comparatively few individuals in
the fresh wave sent forth at each breeding season, survived
to become mature and experienced. It followed from this fact
that the amount of selective pressure exerted by inexperienced
enemies of insects was ten, twenty, a hundred, at any rate
many times as great as that which was due to the educational
period of the mature enemies existing at any moment.
With reference to the PresmpENT’s remarks, on the great
size of Drurya antimachus compared to that of the other
members of the synaposematic group, Mr. F. A. Heron sug-
gested that, in the recognition of prey by sight, size,—within
considerable limits,—might be of minor importance to color-
ation,—the term being used to cover every kind of pattern
and marking. The size of an insect, as correlated with the
idea of its distance, was, in natural surroundings, under
varying atmospheric conditions, extremely difficult of exact
estimation, though it might be easily observable in a group
of other insects of known size in a standard cabinet drawer.
Distance, and its correlative size, might perhaps be especi-
ally hard of determination by animals which, like the
majority of insect-hunting birds, had their eyes placed
somewhat laterally and not frontally, as in the anthropoids
in carnivora and in owls and similar predatory birds,
PROC. ENT, S80C, LOND., Vv, 1903, F
( Ixvi )
for, unless the two eyes could be simultaneously focussed
on the same object the estimation ef distance, which deter-
imined the idea of size, could only take place by the knowledge
of the effort made to secure the focus of one eye, instead of
by the system of unconscious trigonometrical survey, which
was one of the main sources of the knowledge of distance
employed by frontal-eyed animals; unless, indeed, it were
considered that the angle was measured first with one then
with the other eye by quick movement of the head.
With monocular vision, where the distance was not exactly
known, a small object nearer the beholder might subtend a
greater angle than a larger similar one further off, and, if
unfamiliar, be mistaken for the greater.
The idea of distance was one of the more slowly-acquired
concepts ; but the eyes of the young of all animals were quickly
taken by conspicuous pattern.
Coloration, at one extreme, served to break up the apparent
mass and protectively obscure it, while at the other it invited
attention as some glaring label ; a Porson label, which would
denote danger to the consumer of the contents of the object
bearing it, and, as in the case of the Porson label, the danger
would exist irrespective of the size of the ‘“label’’ and the
object it protected. Perhaps it might not be too strong an
assumption to consider that the young inexperienced enemy,
tasting the gaudily-coloured, distasteful Acrawa, would be
impressed, more by the coloration—by the Portson label—
than by the size of the object, and afterwards would avoid
similarly-coloured objects which crossed its field ‘of vision,
without taking any conscious account of their size.
Paper.
Mr. Epwarp Saunpers, F.R.S., contributed “A Supple-
mentary Note to a Paper entitled ‘Hymenoptera Aculeata
collected by the Rev. A. E. Eaton, M.A., in Madeira and
Tenerife, in the Spring of 1902.’”
Wednesday, December 2nd, 1903.
Professor Epwarp B. Pouuron, M.A., D.Se., F.R.S., Pre-
sident, in the Chair,
( ixvii )
Election of Fellows,
Mr. F. H. Day, of Carlisle, The Rev. Tuomas Prinsrp
Levert, of Frenchgate, Richmond, Yorkshire, and Parkington
Hall, Lichfield, and Mr. Roserr C. L. Perkins, B.A., of
Honolulu, were elected Fellows of the Society.
Nomination of Officers and Council for 1904.
Mr. H. Goss, one of the Secretaries, again read the names
of the Officers and Members of the Council proposed for
election at the General Meeting.
Exhibitions.
Mr. G. T. Porrirr exhibited, on behalf of Mr. T. Ashton
Lofthouse, a specimen of Xylophasia zollikoferi taken at
Sugar, near Middlesbrough, Yorkshire, on the 26th September
last. He said he believed that this was only the second
specimen which had been recorded as having been taken
in Britain. Mr. Mclacuian, F.R.8., said the strongest
evidence existed that a very large immigration of insects
from the nearest Continental coast took place during the
exceptional (for this year) spell of warm and calm weather
prevailing towards the end of September, and he was of opinion
that the specimen of NXylophasia zollikoferi, taken by Mr.
Lofthouse in Yorkshire, formed an item in this migratory
swarm, Mr. Kacue Crarke had witnessed such immigration
when staying on board the Kentish Knock lightship for the
purpose of studying bird-migration, He had witnessed a
considerable immigration of Vanessa cardui, for instance,
amongst many other insects, and not the least remarkable of
his observations was the fact that V. cardui flies at night
during migration as well as by day. Mr. MclLacu.an re-
marked that the laws governing migration in insects were
at present little understood, and urged upon entomologists
the necessity of obtaining a clearer insight into their working.
The Presiwent and Mr. Jacosy continued the discussion.
Mr. Matcotm Burr exhibited, and remarked on, a specimen
of Dinarchus dasypus, llig., belonging to a family of five or
six species confined to the Balkans,
The Prestpent exhibited a series of photographs sent by
Mr. A. H. Thayer to illustrate his views on the significance
( Ixviii_ )
of the colours and patterns of butterflies’ wings. The insects
had been photographed on masses of foliage and flowers, and it
was obvious that the dark ground-colour harmonized with the
dark shadow behind and under the vegetation, while the
light markings stood out as conventionalized representations
of single flowers and flower-masses.
The PrestpENt also exhibited the eyeless imagines and
pupa-cases of Hnnomos autumnaria, in illustration of his
remarks at the meeting on November 18th. Imagines pro-
duced by unblinded larvee were also shown for comparison.
Dr. CHAPMAN made some remarks on the specimens
exhibited by the President.
Papers.
The Rev. Francis D. Moricr, M.A., read a Paper entitled
“ Tllustrations of the male terminal ségments and armatures
in thirty-five species of the Hymenopterous genus Colletes.”
ANNUAL MEETING.
January 20th, 1904.
Professor Epwarp B. Poutton, M.A., D.Se., F.R.S., President,
in the Chair.
Mr. Ropert Wynter Lioyp, one of the Auditors, read the
Abstract of the Treasurer’s accounts, showing a balance in the
Society’s favour of £47 2s. Td.
Mr. Herpert Goss, one of the Secretaries, read the
following :
Report of the Council.
During the Session 1903-1904 five Fellows have died, viz.
Mr. Frederick Bates, the Rev. John Hocking Hocking, M.A..,
the Rev. Thomas A. Marshall, M.A., Mr. Philip Brookes
Mason, M.R.C.S., F.L.S., and Mr. John S. Stevens; eight
Fellows have resigned; and twenty-three new Fellows have
been elected.
As was the case last year, the number of Fellows who have
died is below the average, as is also the number of those who
have resigned ; whilst the number of new Fellows is the same
( Ixix )
as last year, and six less than in 1901. The Council hope that
the Fellows will during the coming Session display more
energy and secure a considerable addition to the Society’s list
of members.
At present the Society consists of twelve Honorary Fellows,
and four hundred and seventy-one Life and Subscribing
Fellows, making a total of four hundred and eighty-three,
which, notwithstanding the losses by death and resignation, is
an increase by eleven on last year’s list, and represents a
membership greater than in any previous year of the Society’s
existence of seventy-one years.
The Transactions for the year 1903 form a volume of 575
pages, containing twenty-seven Memoirs, contributed by the
following authors: Mr. Gilbert J. Arrow, Mr. Peter Cameron
(two papers), Mr. G. C. Champion and Dr. T. A. Chapman, Mr.
G. C. Champion, Dr. T. A. Chapman, Mr. Lionel Crawshay,
Mr. W. L. Distant, Dr. Frederick A. Dixey, Mr. Henry J.
Elwes, F.R.S. (two papers), Mr. F. Du Cane Godman, F.RB.S.,
Sir George Hampson, Bart., Mr. Martin Jacoby, Mr. Oliver
Janson, Mr. Percy I. Lathy, Professor Edward B. Poulton,
F.R.S. (two papers), Mr. Ambrose Quail, Mr. G. A. James
Rothney, Mr. Edward Saunders, F.R.S. (two papers), Colonel
Charles Swinhoe (two papers), Mr. T. H. Taylor, Mr. Abbott
H. Thayer, and Mr. Charles Owen Waterhouse.
Of these twenty-seven papers, six relate to Coleoptera, one to
Diptera, one to Hemiptera, six to Hymenoptera, and nine to
Lepidoptera. In addition, there are four papers which cannot
be identified with any one order of insects, viz. Mr. Champion’s
paper entitled “ An Entomological Excursion to Bejar,” which,
in addition to an account of his travels with Dr. Chapman in
Central Spain, contains lists of his captures of Coleoptera and
Hemiptera-Heteroptera ; Professor Poulton’s very interesting
paper on his experiments upon “The Colour Relation between
Lepidopterous Larve and their Surroundings,’ Mr. Abbott H.
Thayer’s paper on “ Protective Coloration in its Relation to
Mimicry, Common Warning Colours, and Sexual Selection,”
and Professor Poulton’s paper entitled ‘‘ A Brief Discussion of
A. H. Thayer’s Suggestions as to the meaning of Colour and
Pattern in Insect Bionomics.”
C Alix?)
It is to be regretted that no papers on Neuroptera or
Orthoptera have been published during the year, and only one
paper on Diptera and one on Hemiptera ; but it is satisfactory
to have published so many papers relating to Coleoptera,
Hymenoptera, and Lepidoptera, in addition to several papers
of interest to students of insect bionomics.
The Memoirs above referred to are illustrated by twenty-
three plates, of which fifteen are coloured. Towards the cost
of Plate VIII Mr. Herbert Adams and Mr, Lathy contributed
the greater portion. Half the cost of Plates IX, XII, XIII,
XIV, and XV was paid by Mr. H. J. Elwes, F.R.S., two-
thirds of the cost of Plates XVI, XVII, and XVIII have
been contributed by Professor Poulton, F.R.S., and the entire
cost of Plates XX, XXI, XXII, and XXIII has been
defrayed by Mr. F. Du Cane Godman, F.R:S.
The interest shown by the Fellows during the past year by
their exhibitions and discussions has enabled the Secretaries to
double or treble the Reports of the Proceedings, which up to
the present date extend to sixty-four pages, and thus compare
very favourably with those for any previous year of the
Society’s existence.
During the past year twenty-six volumes—in addition to
periodicals, pamphlets, and reprints—have been added to the
Society’s Library. The increased use of the Library is proved
by the fact—according to the Resident Librarian’s Report —
that three hundred and thirty-four volumes have been borrowed
by Fellows during the year.
The Treasurer reports that after carrying forward to 1904
the sum of £18 18s. Od. for subscriptions paid in advance in
1903, and investing in Consols the sole Life Composition
received during the year, making the total sum so invested
£744 18s. Od., there remains a genuine cash balance in the
Society’s favour of £47 2s. 7d. The subscriptions received for
1903 slightly exceed those for 1902, and are therefore the
highest on record. The amount of arrears received has
exceeded expectations. ‘The admission fees have been less
than usual, not because a fewer number of new Fellows were
elected, but because many of these are resident in the Colonies,
and are therefore exempt from admission fees. The most
(> ixxis 9}
notable feature of the balance sheet is the large sum received
for Sales of Publications. The increase under this heading
came opportunely during the last quarter. On the whole the
financial position is perfectly satisfactory, and the Fellows
have to be congratulated on the increased and increasing
prosperity of the Society.
ENTOMOLOGICAL Society,
11, CHANDos STREET, CAVENDISH SQUARE, W.
20th January, 1904.
The Secretaries not having received any notice proposing to
substitute other names for those contained in the list prepared
by the Council, the following Fellows constitute the Council
for 1904-1905: Lieut.-Colonel Charles Bingham, F.ZS. ;
George C. Champion, F.Z.8.; Dr. Thomas A. Chapman, M.D.,
F.Z.S.; Arthur John Chitty, M.A.;.James Edward Collin ;
Dr. Frederick A. Dixey, M.A., M.D.; Hamilton H. C. J.
Druce, F.Z.S.; Herbert Goss, F.L.8.; William John Lucas,
B.A. ; Robert McLachlan, F.R.S. ; The Rev. Francis D. Morice,
M.A.; Professor Edward B. Poulton, M.A., D.Sc., F.R.S. ;
The Hon. N. Charles Rothschild, M.A., F.L.S.; Henry Rowland-
Brown, M.A.; Dr. David Sharp, M.A., F.R.S. ; Colonel Charles
Swinhoe, M.A., F.L.S.; Colonel John W. Yerbury, R.A.,
F.Z.8.
The following are the Officers elected: President, Professor
Edward B. Poulton; 7reaswrer, Robert McLachlan; Secretaries,
Herbert Goss and Henry Rowland-Brown ; Librarian, George
C. Champion.
The Balance Sheet and Report having been unanimously
adopted, Professor Poulton, the President, delivered his
Address, A vote of thanks to the President for his Address
and for his services as President during the past year was
moved by Dr. F. A. Dixey, seconded by Canon Fowler, and
carried. The President replied. Mr. G. H. Verrall moved a
vote of thanks to the other Officers of the Society. This was
seconded by Mr. A. J. Chitty, and carried. Mr. Goss and
Mr. Rowland-Brown replied.
(
Ixia) ))
ENTOMOLOGICAL SOCIETY OF LONDON.
Balance Sheet for the Year 1903.
RECEIPTS. PAYMENTS.
G5 Gh iS Gh Oh
Balance in hand, Ist Jan., Printing Transactions, ete. 242 5 6
1903 ‘ ... 2610 7 | Plates, etc. e epee
Subscriptions for 1903 . 870 13 O | Rent and Office Ex-
Arrears 33 12 0 penses 5 IBY)
Admission Rees 33 12 0 | Books and Binding... 2817 1
Donations ... 59 12 2 | Investments in Consols... 15 15 0
Sales of Transactions, etc. 117 7 10 | Subscriptions in advance,
Interest on Investments :— per contra, carried to
Consols ... ... £18 4 8 1904 1818 0
Westwood Bequest6 16 0 Balance 47 2 7
25.0 8
Subscriptions in advance 1818 0
Life Composition ilsy lbs 2 (0)
£701 1 3 £701 1 3
ASSETS.
Subscriptions in arrear considered good (say) £21 0 0
Investments :—
Cost of £752 6s. 6d. Consols ... F .. £744 18 0
Cost of £239 12s. 4d. Birmingham 38 per Sea 250 0 0
Balance in hand 47 2 7
NO ASCERTAINED LIABILITIES.
Ropert McLAcunan,
Treasurer.
13th January, 1904.
Examined and found correct.
A. Huau Jonss.
GEO. S. SAUNDERS.
Hamitton H. Druvce.
J. W. YERBURY.
R. Wyuiz Luioyp.
( Ixxii )
THE PRESIDENT’S ADDRESS.
GENTLEMEN,
It is a great pleasure to congratulate the Society at the
close of another successful year. The repetition of this con-
gratulation in successive Annual Addresses is happily almost
monotonous. It is a monotony which will never weary us,
and in itself an indication that no other monotony has
prevailed.
The meetings have been well attended, there have been
numerous, varied, and interesting exhibits leading to animated
discussions, Our Transactions do not reach the phenomenal
dimensions attained in 1902, but still form a noble volume,
containing 23 plates and well over 600 pages. There is a
pleasing variety in the papers, and the domination of the
Lepidoptera is less pronounced than usual. An important
share of the space is occupied by memoirs on the Coleoptera,
Hymenoptera, and Insect Bionomics, while the Diptera and
Rhynchota are also represented.
I should wish to refer again to the warmth of the greeting
received as your President at a meeting of the Entomological
Society of France on April 22nd. The cordial friendship
between the followers of science in all lands is of happy augury
for the advancement of the researches in which we find common
aims and mutual sympathy and respect.
On this, the first occasion on which I have the honour of
addressing you formally, I cannot resist the temptation of
calling attention to a remarkable coincidence of a personal
nature—the fact that the present occupant of this Chair and
his immediate predecessor should be members not only of the
same University, but of the same College, and that not a large
one. When this fact was explained to a friend he said it
was easily understood, because the study of natural history is
infectious. This suggestion, plausible as it is, fails to account
for the fact ; inasmuch as Canon Fowler left Jesus College,
( lxxiv. )
Oxford, in June 1873, while I did not matriculate until October
of the same year, so that, as undergraduates, we never saw
each other.
Before speaking of the losses which have fallen so heavily
upon our community during 1903,—the brother Fellows who
have gone from our midst,—I feel bound to allude to the
grief which we share with the whole intellectual world at
the passing away, towards the close of the old year, of
the great thinker to whom we owe far more than we can
realise. I well remember the sudden access of light received
when, between the age of seventeen and eighteen, Herbert
Spencer’s works were first placed in my hands. The whole of
science seemed illuminated, the whole outlook broadened. It
was the most sudden and by far the greatest intellectual
awakening I have ever experienced. And, as we know well,
it has been the same with thousands. After Shakespeare, no
man has done more to bring together the English* of the
Old World and the New. And not only among ourselves,
but everywhere in the civilised world the writings of Herbert
Spencer have stirred enthusiasm and compelled admiration,
They have left strong, indelible, beneficent after-effects even
in those who are unable to believe in the enduring stability
of the Synthetic Philosophy—a fabric as fair and stately as
any created by the mind of man.
Since the above paragraph was written Mr. Herbert
Spencer’s will has been made known in the 7imes for January
14th. I am sure that every Fellow of our Society keenly
appreciates the expression of confidence which is implied in
the gift which will hereafter be offered to us by the Trustees
of the will—a gift which we shall regard as a solemn trust,
to be so carried out as to secure the greatest possible advantage
to the science we serve.
Freperick Bates, F.E.S., joined the Society as a ‘‘sub-
seriber” in 1867. Subsequently withdrawing, he again
entered the Society as a Fellow in 1897. He was born at
Leicester in 1829, and his death occurred at Chiswick on the
6th of October, in his 74th year. Like his distinguished
* For the justification of this use of the word see Sir Michael Foster’s
Presidential Address to Section D of the British Association at Toronto
(Report for 1897).
(© lxxy=))
brother, H. W. Bates, F.R.S., he was especially devoted to the
Coleoptera, although his interests were wide and embraced
many aspects of natural history, both zoological and botanical
He was the author of many papers, chiefly dealing with the
Heteromera, in our Transactions and in the ‘ Entomologist’s
Monthly Magazine.” His exceedingly fine collection of
Heteromera is nowin the British Museum, while his collection
of British Coleoptera was a gift to his intimate friend Mr.
Horace Donisthorpe. Many friends mourn the loss of a keen
and able naturalist, a many-sided and genial personality.*
THe Rev. Joun Hocxine Hockine, M.A., J.P., F.ES.,
Rector of Copdock-with-Washbrook, near Ipswich, was elected
a Fellow in 1896. His death occurred on the 10th of
December last, at the age of 69. He was an ardent collector
of the Lepidoptera, but having only recently joined the Society
was unfortunately known to but few of the Fellows.t
THe Rev. THomas AnseLL Marsuatu, M.A., F.E.S., joined
the Society in 1865. By his death on April 11, 1903, at
Ajaccio, one of the few authorities upon the parasitic Hymen-
optera is lost to science. Mr. Marshall was born at Keswick
on March 18, 1827, the son of Thomas Marshall, an original
member of the Entomological Society. He took a scholarship
at Trinity College, Oxford, and passed through the Classical
Honours course. With great powers asa linguist, and a student
of Hebrew and Sanskrit, he worked for a time on the staff of
the British Museum Library. Subsequently he took Holy
Orders, and after engaging in scholastic work, held livings in
various parts of England, interrupted only by his appoint-
ment as Bishop’s chaplain in Antigua. In this island he was
bereft of his wife, and was himself in serious danger from an
attack of fever. Upon his return to England he was presented,
in 1889, to the living of Botus Fleming, Cornwall, which he
retained until 1897, when he retired to Corsica, and devoted
the remainder of his life to his favourite science. T. A.
Marshall’s earliest important work dealt with the Coleoptera
(Journ. Linn. Soc. 1865). The first of the series of memoirs
* See the Obituary notice in the ‘‘ Entomologist’s Record,” vol. xv, No.
12, by Horace Donisthorpe ; and that in the ‘‘ Entomologist’s Monthly
Magazine,” Nov. 1903, p. 286.
+ See also the Obituary notice in the ‘‘Entomologist’s Monthly
Magazine ” for Jan. 1904, p. 19.
( Ixxvi )
by which his name will be chiefly known was published in
1870, ‘“Ichneumonidum Britannicorum Catalogus,” followed
by the valuable monographs on the British Parasitic Hymen-
optera, which appeared in the Transactions of this Society
between 1872 and 1889. He published an important volume
on a portion of the Braconide in André’s “Species des
Hyménopttres d’Europe,” and was still at work on the subject
at the time of his death. He was an accomplished draughts-
man and a clear and admirable writer. The loss of so able
a student of an important but much-neglected group will be
long and deeply deplored.*
Puitie Brookes Mason, M.R.C.S., F.LS., F.E.S., a Fellow
of our Society since 1874, died on November 6, 1903, at
Burton-on-Trent. His death is a sad loss to his profession, to
the neighbourhood in which he laboured, and to a wide circle of
naturalist friends. Mr. Mason was born at Burton on January
2, 1842. After a medical education of unusual distinction
and variety of valuable experience, he made his permanent
home at his native town in the Midlands. The British fauna
and flora formed the chief interest of his life, and he possessed
magnificent collections of both. With his sympathetic genial
nature, it was his delight to welcome his brother naturalists to
share in the well-nigh unique advantages which he possessed.
And as he was a skilled and honoured member of the healing
profession, so he was ever ready to lend the weight of his
influence and the power of his persuasion to promote peace
and friendliness. As the Society concerned with the branch of
natural history to which he was chiefly devoted, we recognise,
with grief, that a strong influence for good has passed
from us.t
Joun Sanpers Srevens, F.E.S., became a Fellow in 1862.
His death at Woking, on July 15, makes a sad break in
the ranks of the senior Fellows of the Society.
* See also the Obituary notice in the ‘‘ Entomologist’s Monthly Maga-
zine,” June 1903, p. 152, by R. McLachlan, F.R.S. ; and that in the ‘‘ Ento-
mologist’s Record,” vol. xv, No. 7, p. 190, by G. C. Bignell.
+ See also the Obituary notice in the ‘‘Entomologist’s Monthly
Magazine,” Jan. 1904, pp. 17, 18, by the Rev. Canon W. W. Fowler;
also ‘“‘The Lancet” for Nov. 13, 1903.
~ See also the Obituary notice in the ‘“‘Entomologist’s Monthly Maga-
zine,” Sept. 1903, p. 229.
( Ixxvii )
Outside the number of our own Fellows, we miss four well-
known names from the ranks of British entomologists :—
Witiram Duppa Crorcn, M.A., F.L.S., a keen student of the
Lepidoptera, Coleoptera and Hemiptera; Epwarp Roserr
Date, son of the eminent J. C. Dale, and himself an eager
entomologist in his younger days; THe Very Rey. Canon
BernarD Smita of Great Marlow, an enthusiastic collector and
breeder of the British Lepidoptera ; Samce: James WILKINSON,
author of the celebrated “ British Tortrices,” published in
1859.
We sympathise deeply with our brethren on the continent
in their grief for the eminent men who have passed away in
1903 :—Jonannes Faust, the eminent authority upon the
Curculionide, whose collection contained over 13,000 species,
of which more than 2000 were described by himself ; Pro-
FEssor Aucustus Rapcuirre Grote, A.M., the celebrated
student of the Lepidoptera.
“WHAT IS A SPECIES?”
The late Professor Max Miiller, in an eloquent speech
delivered at Reading in 1891, spoke of the necessity of
examining, and, as time passes by, re-examining the meaning
of words. He referred as an illustration to the man at the
railway station who taps the wheels with his hammer, test-
ing whether each still rings true or has undergone some
change that may mean disaster. In almost the same way,
the speaker maintained, a word may slowly and unobtrusively
change its meaning, becoming, unless critically tested to
ascertain whether it still rings true, a danger instead of an
aid to clear thinking, a pitfall on the field of controversy.
He then went on to say, that Darwin had written a great
work upon the Origin of Species, and had never once explained
what he meant by the word Species. So decided an utterance
—the statement was made emphatically—ought to have in-
volved a careful and critical search through the pages of
the work that was attacked. However this may be, it is
quite certain that the search was unsuccessful; and yet a
few minutes’ investigation brought me to a passage in which
the meaning attached by the author to the term Species is set
( Ixxvili )
down in the clear, calm, and simple language which did so
much to corvinee an unwilling world.
Darwin is speaking of the revolution which the acceptance
of his views will bring about. “ Systematists will be able to
pursue their labours as at present; but they will not be
incessantly haunted by the shadowy doubt whether this or
that form be in essence a species. This, I feel sure, and I
speak after experience, will be no slight relief. The endless
disputes whether or not some fifty species of British brambles
are true species will cease. Systematists will have only to
decide (not that this will be easy) whether any form be suffici-
ently constant and distinct from other forms to be capable of
definition, and if definable, whether the differences be sufficiently
important to deserve a specific name. This latter point will
become a far more essential consideration than it is at present ;
for differences, however slight, between any two forms, if not
blended by intermediate gradations, are looked at by most
naturalists as sufficient to raise both forms to the rank of
species. S/ereafter we shall be compelled to acknowledge that
the only distinction between species and well-marked varieties is,
that the latter are known, or believed, to be connected at the
present day by intermediate gradations, whereas species were
formerly thus connected. Hence, without quite rejecting the
consideration of the present existence of intermediate grada-
tions between any two forms, we shall be led to weigh more
carefully, and to value higher, the actual amount of difference
between them. It is quite possible that forms now generally
acknowledged to be merely varieties may hereafter be thought
worthy of specific names, as with the primrose and cowslip ;
and in this case scientific and common language will come
into accordance. In short, we shall have to treat species in
the same manner as those naturalists treat genera, who admit
that genera are merely artificial combinations made for con-
venience. ‘This may not be a cheering prospect, but we shall
at least be freed from the vain search for the undiscovered
and undiscoverable essence of the term species.” I have
quoted from pages 484, 485 of the original edition (1859),
and have italicised the sentences in which Darwin defines a
species and distinguishes it from a variety.
(lexi 9)
Max Miiller’s special criticism falls to the giound, but his
general exhortation remains, and I think we shall do well to
be guided by it, and attempt to apply it to this difficult and
elusive word SPECIES.
The passage I have quoted was Darwin’s prediction of the
meaning which would be attached to the word “species ”’ by
the naturalist of the future. Nearly half-a-century has passed
since those words were written. For more than a generation
the central ideas of the “Origin” have been an essential part
of the intellectual equipment, not only of every naturalist, but
of every moderately intelligent man. What then is the
meaning of the word “species” to-day, and how does it differ
from that of the years before July 1, 1858, when the Darwin-
Wallace conception of natural selection was first launched
upon the world ?
The present occasion is especially favourable for this inquiry,
because we have just been given two additional volumes of
the letters of Charles Darwin. After the- three volumes
published in 1887, naturalists were certainly unprepared for
the welcome revelation of such a mine of wealth. The work
is all the more valuable because it contains many letters from
Alfred Russel Wallace and Sir Joseph Hooker, thus giving
both sides of a part of their correspondence with Darwin.
Then in 1900 the “ Life and Letters of Thomas Henry Huxley ”
appeared, so that we are now admitted “behind the veil,” and
can read, as never before, the central thoughts of the great
makers of biological history. On the publication of the last-
named work, I took occasion to combat the view that the
thousand closely-printed pages might have been reduced by
omitting and condensing many of the letters. The serious
student of those stirring years requires the opportunity of
thinking over and comparing all the available thoughts and
opinions of the chief actors in the memorable scene ; and the
very repetition of certain ideas, which proves their persistence
and dominance in the writer’s mind, is a matter of deep
importance and interest. However it may be to the general
reader, the student would deprecate the omission or condensa-
tion of any of the writings of Darwin or Huxley. The special
interest and value in the letters of these men depend on the
(Or lxeaxa)
fact that their inmost convictions on matters of the deepest
scientific importance are to be read, often in the compass of
a brief sentence. There we find, as we cannot find in any
other way, the real core of the matter, with all accessory and
surrounding considerations stripped away from it.* A care-
ful study of the two recent volumes of Darwin’s letters, and
a re-study of the three earlier volumes, with a view to this
Address, have shown how Darwin’s thoughts were again and
again occupied upon subjects bound up with the problem I
have ventured to bring before you this evening. The interest
reaches its height when we find that strongly-marked differ-
ences of opinion on fundamental questions are threshed out
in the correspondence, when we see, as I shall have occasion
to point out in greater detail in the later pages of this Address,
Darwin differing sharply from Huxley on the one hand, and
with Wallace on the other, as to the significance and history
of sterility between species.
In such episodes we are permitted to become the witnesses
of a supremely interesting struggle, where the central figure
of modern biological inquiry is contending with his chief
comrades in the great fight,—with the co-discoverer of natural
selection, with the warrior hero who stood in the forefront of
the battle.
The correspondence of Charles Darwin has a further deep
interest for us. We see the means by which a gentle,
sympathetic, intensely human nature overpassed the stern
limits imposed by health, and was able to impart and to
receive fresh ideas, and a stimulus ever renewed—the im-
pulse to varied and unceasing research. I have lately been
studying with keen interest the life of another great English-
man, William John Burchell,t than whom no better equipped
or more learned traveller ever explored large areas in two
continents. When I state that searching inquiry has only
brought to light a dozen of his letters, and that he was
known to hardly any of the great naturalists of his day, we
see the reason for the sad, unproductive, brooding close of a
career which opened with almost unexampled brilliancy and
* “Quarterly Review,” January 1901, p. 258.
+ ‘Ann, and Mag. Nat. Hist.,’’ January 1904, p, 45.
(lex 3) ;
promise. The time which we give to Societies such as this—
time we are sometimes apt to grudge—is well spent. Here,
and in kindred communities, a ‘‘man sharpeneth the counten-
ance of his friend,” and there is born of the influence of mind
upon mind thought which is not a mere resultant of diverse
forces, but a new creation.
The scientific man who shuts himself away from his fellow-
men, in the belief that he is thereby obtaining conditions the
most favourable for research, is grievously mistaken. Man,
scientific man perhaps more inevitably than others, is a social
animal, and the contrast between the lives of Darwin and
Burchell shows us that friendly sympathy with our brother
naturalists is an essential element in successful and continued
investigation.
I do not suppose that it is necessary to justify a discussion
of the term “species” as the subject of the Anniversary
Address to the Entomological Society of London. The students
of insect form and function hold an exalted place among
naturalists. The material of their researches enables them,
almost compels them, to take the keenest and most active
interest in broad questions affecting the history and course of
life on our planet. Naturalists engaged upon other groups
may reasonably inquire why insects, above all other animals,
should be so especially valuable for the elucidation of the
larger problems which deal, not only with the species of a
single group, but with every one of the innumerable and
infinitely varied forms, vegetable no less than animal, in
which life manifests itself. ‘The answer is to be found in the
large number of offspring produced by each pair of insects,
and the rapidity with which the generations succeed each
other, many cycles being completed in a single year in warm
countries; in the severity of the struggle for life which
prevents this remarkable rate of multiplication from becom-
ing the cause of any progressive increase in the number of
individuals; and finally, in the character of the struggle
itself, which is precisely of that highly specialised kind
between the keen senses and activities of enemies, and the
means of concealment or other modes of defence of their
insect prey, which leads, by action and answering reaction, to
PROC. ENT. SOC. LOND., Vv. 1903. G
( Ixxxi )
a progressive raising of the standard in both pursuer and
pursued. This is why it is that insects mean so much to the
naturalist or to the philosopher who desires to look beneath
the surface for the forees which have moulded existing forms
of life out of earlier and very different forms. The wings of
butterflies, it has been said, serve as a tablet on which Nature
writes the story of the modification of species.* But the
careful study of insects tells us even more than this; for it
gives us the clearest insight we as yet possess into the forces
by which these changes have been brought about. Light is
thrown upon the causes to which organic evolution is due no
less than upon the course which organic evolution has pursued.
And I think we shall find that a consideration of the
numerous distinct categories of forms presented by the insect
world is especially advantageous in an attack upon the difficult
question—“ What is a species!”, while properly-directed
observation of insects, and experiments upon insects afford
the most hopeful prospect of a final answer.
And here I am compelled to say a word in defence of the
Lepidoptera from this point of view. Undoubtedly it is most
unfortunate that the obvious attractions of the group have
led entomologists to neglect other Orders ; for this can be the
only explanation why naturalists have so often preferred to
do over again what others have done already, apparently
oblivious of fields comparatively empty and unexplored. It
must further be admitted, that the greater visibility of
structure, and the more urgent necessity for the study
of structure in other groups, render them better instruments
of zoological education. But although the Lepidoptera are
inferior in this respect, although they lack the unique interest
of the Hymenoptera and the social Neuroptera, and can-
not claim any of the respect due to venerable age like the
Aptera, Orthoptera and Neuroptera—in spite of their many
demerits they stand at the head, not only of all insects, but
* H. W. Bates, quoted by A. R. Wallace in ‘‘ Natural Selection,”
London, 1875, p. 132. The original passage may be found in ‘‘ The
Naturalist on the Amazons” (London, pp. 347, 348 of the 1879
edition).
+ This justification for the study of insects was urged by the present
writer in the Hope Reports, vol. iii, 1903, preface, pp. 4, 5.
(> ia.)
of the whole organic world, as the registers of subtle and
elusive change—ever going on, yet never seen,—by means of
which forms are slowly becoming different from what they
have been in the past. It is the existence of a complex
pattern composed of several colours, which renders butterflies
and to a less extent moths such a remarkably delicate record
of change. As we trace the representative individuals of a
community of butterflies over any wide range, the trained eye,
and often the inexperienced eye, can detect differences which
are not seen to anything like the same extent in the individuals
of other Orders with corresponding ranges. If the wings of
Hymenoptera, Diptera, or Orthoptera possessed the same
elaborate patterns as the Lepidoptera, we cannot doubt that
they too would exhibit the same differences in various parts
of their areas. These continual changes which we find as we
study the distribution of Lepidopterous forms in space, is
undoubtedly a measure of the speed with which they have
occurred in time. Rapidity of change is essential if it is
to keep its adjustment with nicety to the fleeting details of
distribution.* Hence we may confidently believe, that if we
* It is to be observed that I speak of the details as fleeting. The
aeneral area of distribution is doubtless extremely ancient in most cases.
Thus, although a species of Heliconius, etc., may have originated within
the South American tropics, and never have wandered beyo..d them, the
complex shape of its actual area of distribution at any one time cannot be
regarded as fixed or ancient. Yet in many a species the variation of the
constituent individuals is adjusted with precision to the geographical details
of the existing range.
Mr. Roland Trimen, on reading the above footnote, writes to me
January 24, 1904 :—‘‘ Your note reminds me of the recent appearance
on the Natal coast of several conspicuous East-African butterflies, vid. :
Pieris spillert, Crenis rosa, and Godartia wakefieldii, all of which are
shown to have not only extended their range to a point where they were
previously quite unknown, but to have also established themselves in the
fresh area. This is a good case, as Durban has had, for the last twenty-
five years at least, a number of keen collectors of Lepidoptera, whom such
conspicuous forms could not possibly have escaped had they inhabited the
neighbourhood. Besides these species, the last butterfly that my friend
and collaborator, the late Colonel Bowker, sent to me (1898) was the large
and extremely conspicuous black-and-white Acrea satis, which he took
at Malvern, near Durban. This is the only example known to me to have
occurred in Natal; but Bowker, who noted the resemblance on the wing
to Papilio morania, wrote that he had seen one other for certain, and
thought that he might very possibly have passed over more examples for
the common Papilio named, This last case is of special interest (should
it prove one of extended range like the three mentioned), because the
Acree are so exceptionally slow-flying and gregarious, that they must
spread very slowly indeed into fresh areas.”
(7 Uisxcxay =)
could wake up in say a thousand years, we should be able to
detect changes in the patterns of some butterflies. Although
T am afraid the advance of science is not likely to be sufficiently
rapid in our time for me to hold out any prospect of such an
experience for any of you, there is every reason why we should
afford this opportunity to posterity. A critical examination
of the fragments of many species of butterflies captured ninety
years ago by Burchell in 8. Africa, and gnawed to pieces
during his Brazilian travels from 1825 to 1830, renders it
probable, nay, almost certain, that with moderate care, insect
pigments will endure for an indefinite period in our museums.
One important justification for the great and permanent
outlay required to bring together and maintain large collec-
tions of insects is, that we are allowing our successors the
chance of detecting and measuring the rate of specific
change.* And, as I have already said, for this purpose the
Lepidoptera stand pre-eminent.
For the purpose of the inquiry this evening, our instances
will be drawn from the Lepidoptera rather than other Orders
of insects, because of the numberless examples of subtle
distinction between forms which but yesterday, so to speak,
became separate ; because of our knowledge, insufficient but
considerable, of their geographical ranges; because of our
experience, excessively imperfect and scanty, but still much
larger than in other Orders, of inter-breeding and of descent
from parent to offspring.
First among the attempts to define species must be placed
that which we rightly associate with the name of Linnzus.
It has been admirably pointed out by the late Rev. Aubrey
L. Moore, + that the dogma of the fixity of species is entitled
to none of the respect which is due to age. “It is hardly
credible to us,” he wrote, ‘that Lord Bacon, ‘the father of
* Karl Jordan argues with great force in favour of specialisation in
this direction by our museums. (See ‘‘ Novitates Zoologice,” vol. iii,
December 1896, pp. 431-433.) The Burchell collection from Brazil is
only seventy-four to seventy-nine years old, but the species are numerous,
and often represented by long series. An account of the butterflies by
Miss Cora B. Sanders will shortly appear in the ‘‘ Annals and Magazine of
Natural History”; and it will then be seen that the evidence of change
in certain forms is by no means wanting.
+ ‘Science and the Faith,” London, 1889, pp. 174 et seq.
( Ixxxv )
modern science’ as he is called, though he was only a school-
man touched with empiricism, believed not only that one
species might pass into another, but that it was a matter of
chance what the transmutation would be. Sometimes the
medieval notion of vivification from putrefaction is appealed
to, as where he explains the reason why oak boughs put into
the earth send forth wild vines, ‘which, if it be true (no
doubt),’ he says,* ‘it is not the oak that turneth into a vine,
but the oak bough, putrefying, qualifieth the earth to put
forth a vine of itself.’ Sometimes he suggests a reason which
implies a kind of law, as when he thinks that the stump of a
beech tree when cut down will ‘put forth birch,’ because it
is a ‘tree of a smaller kind which needeth less nourishment.’ t
Elsewhere he suggests the experiment of polling a willow to
see what it will turn into, he himself having seen one which
had a bracken fern growing out of it!{ And he takes it as
probable, though it is inter magnalia nature, that ‘whatever
creature having life is generated without seed, that creature
will change out of one species into another.’ Bacon looks
upon the seed as a restraining power, limiting a variation
which, in spontaneous generations, is practically infinite, ‘for
it is the seed, and the nature of it, which locketh and boundeth
in the creature that it doth not expatiate.’” And the author
also shows that much earlier than the date at which Bacon
wrote, theologians were by no means unanimous in accepting
“ special creation ”’ ; that St. Augustine even distinctly rejected
it, and propounded an idea which was evidently considered
tenable by the greatest of the schoolmen, St. Thomas Aquinas,
St. Thomas’ words, quoted by Mr. Aubrey Moore, are as
follows :—‘‘ As to the production of plants, Augustine holds
a different view. For some expositors say that, on this third
day (of creation), plants were actually produced each in his
kind—a view which is favoured by a superficial reading of
the letter of Scripture. But Augustine says that the earth
is then said to have brought forth grass and trees causaliter—
z.e. it then received the power to produce them.” §
* “CO Nat. Hist... Cent. vi, 522, fol. ed:
tf 2. ¢. p.-523: ae Up jay WUE
§ St. Thomas Aquinas, ‘Summa Theol.” Prima Pars. Quaest., ]xix,
Art. 2.
( lxxcevai 5)
How then did the fixity of species become an article of
belief in later years? Aubrey Moore traces it to the influence
of Milton’s account of creation in the seventh book of
“Paradise Lost” (1. 414, e¢ seg.), and Professor Huxley
had still earlier suggested the same cause in his ‘‘ American
Addresses.” I cannot help thinking that the belief had even
more to do with the spirit of the age which spoke, and spoke
for all time, with Milton for its interpreter,—the spirit of the
Puritan movement, with its insistence on literal interpreta-
tion and verbal inspiration.
John Ray was Milton’s younger contemporary, and many
writers, including Aubrey Moore, have thought that with him
began the idea of the fixity of species. Sir William Thiselton
Dyer has, however, recently pointed out, that a conception
similar to Ray’s may be traced to Kaspar Bauhin (1550-1624)
and to Jung (1587-1657).*
From Ray we pass to Linneus with his often- quoted
definition, “Species tot sunt, quot diversas formas ab initio
produxit Infinitum Ens, quae formae, secundum generationis
inditas leges produxere plures, at sibi semper similes.” Of
the Ray-Linneus-Cuvier conception of species, which found its
most precise and authoritative expression in the above-quoted
latin sentence, Dr. F. A. Dixey has well said that it ‘‘ left
order where it found confusion, but in substituting exactness
of definition for the vague conceptions of a former age, it did
much to obscure the rudimentary notions of organic evolution
which had influenced naturalists and philosophers from
Aristotle downwards.” + At the same time it is by no means
improbable, as Dixey has suggested, that the Linnean concep-
tion “of the reality and fixity of species perhaps marks a
necessary stage in the progress of scientific enquiry.” {
The Linnean idea of special creation has no place in the
realm of science ; it is a theological dogma. The formation
of species, said Darwin in a letter to Lyell, ‘has hitherto
been viewed as beyond law ; in fact, this branch of science
* «The Edinburgh Review,’ Oct. 1902, p. 370.
+ ‘‘Nature,” June 19, 1902, p. 169. For the history of these early
ideas upon evolution see ‘‘ From the Greeks to Darwin,” by H. F. Osborn,
New York, 1894.
+ ‘Church Quarterly Review,” Oct. 1902, Art. II, p. 28.
( Ixxxvii )
is still with most people under its theological phase of develop-
ment.” * And this explains the intense opposition at first
encountered by the principles of the “ Origin.”” The naturalist
whose genius sympathised most fully with the Linnean con-
ception would feel that he was admitted, like a seer of old,
into the presence of the Maker of the Universe. His
)
convictions as to species were to him more than the conclu-
sions of the naturalist; they were a revelation, stirring him to
“break forth and prophesy.” Do we not sometimes recognise
a lingering trace of this phase of thought in the serious shake
of the head and tone of profound inner conviction with
which we are sometimes told that the speaker is decidedly of
the opinion that so-and-so is a perfectly good species ?
We recognise the same sharp antagonism between two
irreconcilable sets of ideas when the late W. C. Hewitson
expressed such horror at Roland Trimen’s remarkable dis-
covery of the polymorphic mimetic females of the Papilio
merope group. The wonderfully acute detection of minute but
significant resemblance hidden under the widest possible
superficial difference, which enabled the great South African
naturalist to unravel the tangled relationships, was to Hewitson
but one of ‘‘ the childish guesses of the . . . Darwinian School.”
To meet the carefully-thought-out argument, the only objec-
tions that could be urged were that the conclusion stretched
too severely the imagination of the writer, and that it
shocked his notion of propriety ! 7
* Letter 132 to C. Lyell, Aug. 21, 1861. ‘‘More Letters of Charles
Darwin,” London, 1903, i, p. 194.
+ See an account of the controversy in Trans. Ent. Soe. Lond., 1874, p.
137. The passages I have alluded to are as follows :—‘“‘ P. merope, of
Madagascar, has a female the exact image of itself; and it would require
a stretch of the imagination, of which I am incapable, to believe that the
P. merope of the mainland, having no specifie difference, indulges in a
whole harem of females, differing as widely from it as any species in the
genus. . . . In the two species of Papilio which have lately been united,
Torquatus and Candius, and Argentus and Torquatinus, though much
unlike each other, there is quite sufficient resemblance not to shock one’s
notions of propriety.” A little later Mr. Hewitson himself received
evidence of the truth of the conclusion he so disliked; for he told how
his collector Rogers had sent ‘‘ Papilio merope and P. hippocoon, taken
by him in copulation, another illustration of the saying that ‘truth is
stranger than fiction.’ I find it very difficult (even with this evidence) to
believe that a butterfly, which, when a resident in Madagascar, has a
female the image of itself, should, in West Africa, have one without any
resemblance to it at all” (‘‘ Entomologist’s Monthly Magazine,” Oct.
1874, p. 113).
(Gibse cain)
In leaving the dogma of “special creation,” and the assump-
tion of “ fixity of species” with which it is bound up, it is
only right to point out how completely the logical foundations
of both were undermined by the great thinker who has just
passed away. Years before the appearance of the Darwin-
Wallace essay, and of the “Origin,” Herbert Spencer wrote
on “The Development Hypothesis.” * Although of course
wanting the great motive power to evolution supplied by
natural selection, this essay is a powerful and convincing
argument for evolution as against special creation. It is
astonishing that it did not produce more effect. I may appro-
priately conclude this section of the Address by quoting the
results of Herbert Spencer’s critical examination, from every
point of view, of the Linnean conception of species. ‘ Thus,
however regarded, the hypothesis of special creations turns
out to be worthless—worthless by its derivation ; worthless in
its intrinsic incoherence; worthless as absolutely without
evidence ; worthless as not supplying an intellectual need ;
worthless as not satisfying a moral want.” ¢
If then the Linnean conception of species—separately created
and fixed for all time at their creation—has been abandoned,
what have we to put in its place? In a letter to Hooker, Dec.
24, 1856, Darwin gave a list of the various definitions he had
met with. “I have just been comparing definitions of species,
and stating briefly how systematic naturalists work out their
subjects. . . . It is really laughable to see what different
ideas are prominent in various naturalists’ minds when they
speak of ‘species’; in some, resemblance is everything, and
descent of little weight—in some, resemblance seems to go
for nothing, and creation the reigning idea—in some, descent
is the key—in some, sterility an unfailing test, with others it
is not worth a farthing. It all comes, I believe, from trying
to define the indefinable.” {
As regards the work done by the systematist, we find that
Darwin did not agree with those of his friends who thought
* In the Leader, between January 1852 and May 1854, reprinted in
‘*Kssays Scientific, Political, and Speculative.” London, 1868, vol. i,
De Ode
+ ‘*The Principles of Biology.” London, 1864, vol. i, p. 345.
+ “‘Life and Letters of Charles Darwin ” London, 1887, vol. ii,
p. 88.
(Gocxix)
that a belief in evolution would entirely alter its character.
Thus he wrote to Hooker, Sept. 25, 1853 :—“ In my own
work I have not felt conscious that disbelieving in the mere
permanence of species has made much difference one way or
the other ; in some few cases (if publishing avowedly on the
doctrine of non-permanence) I should ot have affixed names,
and in some few cases should have affixed names to remark-
able varieties. Certainly I have felt it humiliating, discussing
and doubting, and examining over and over again, when in my
own mind the only doubt has been whether the form. varied
to-day or yesterday (not to put too fine a point on it, as
Snagsby would say). After describing a set of forms as
distinct species, tearing up my MS., and making them one
species, tearing that up and making them separate, and then
making them one again (which has happened to me), I have
enashed my teeth, cursed species, and asked what sin I had
committed to be so punished. But I must confess that perhaps
nearly the same thing would have happened to me on any
scheme of work.” *
The essentially subjective character of the results reached by
the systematist stands out with remarkable force in this as in
other passages of Darwin’s letters.
A few years later, on July 30, 1856, he wrote to the same
friend :—‘‘I differ from him [Lyell] greatly in thinking that
those who believe that species are not fixed will multiply
specific names: I know in my own case my most frequent
source of doubt was whether others would not think this or
that was a God-created Barnacle, and surely deserved a name.
Otherwise I should only have thought whether the amount of
difference and permanence was suflicient to justify a name.” T
Disregarding for the moment the term species, it is
convenient to consider the various groupings of individual
animals and plants,
1. Forms having certain structural characters in common
distinguishing them from the forms of other groups. Groups
thus defined by Diagnosis may be conveniently called Syndi-
agnostic (ovv, together ; diudyvwors, distinction).
* «Tife and Letters,” vol. ii, p. 40.
+ bid. vol. ii, p. 81.
( xc )
2. Forms found together in certain geographical areas and
not in other areas. Such groups may be called Sympatric (ovv,
together ; zarpa, native country). The occurrence of forms
together may be termed Sympatry, and the discontinuous
distribution of similar forms Asympatry.
3. Forms which freely inter-breed together. These may
be conveniently called Syngamic. (cvv, together; ydpos,
marriage). Free inter-breeding under natural conditions may
be termed Syngamy; its cessation or absence, Asyngamy
(equivalent to the Amiwxia of Weismann).
4, Forms which have been shown by human observation
to be descended from common ancestors. Such groups may be
called Synepigonic (ovv, together; ériyovos, descendant). Breed-
ing from common parents may be spoken of as Epigony or
the production of Epigonic evidence.*
My friend, Professor E. Ray Lankester, to whom I owe
so much, in this as in many other subjects, is inclined to
think that we should discard the word species not merely
momentarily but altogether. Modern zoology having aban-
doned Linnzeus’ conception of “species” should, he considers,
abandon the use of the word. In his opinion the “ origin”
of species was really the abolition of species, and zoologists
should now be content to describe, name, draw, and catalogue
forms. Furthermore, the various groups of forms briefly
defined above should be separately and distinctly treated by
the zoologist, without confusion or inference from one to the
other. The systematist should say, “I describe and name
certain forms a, 6, ete.” ; and then he or another may write
a separate chapter, as it were :—‘‘I now show that the forms
ab, ac, ad (form names) are syngamic:” at another time he
may give reason for regarding any of them as related by
epigony.
I fear that this suggestion is a ‘counsel of perfection,”
impossible of attainment, although there would be many
* My friend Mr. Arthur Sidgwick has kindly helped me by suggesting
the appropriate Greek words. The use of émiyoves I owe to my friends
Mr. Arthur Evans and Mr. R. W. Macan. The adjectival termination is
made -ic throughout for the sake of convenience, although Sympatriote or
Sympatrid would have been more correct.
(xc)
and great advantages in thus making a fresh start and in
the abandonment of “species,” or the restriction of the
word to the only meaning it originally possessed before
it was borrowed from logic to become a technical term in
zoology.*
Professor Lankester in former years published (I cannot
at this moment lay my hands upon the communication) the
suggestion that the term species should be limited to a group
which includes all the forms derived from common ancestors
within human experience, or inferred to be so derived within
the possible period of human observation. Thus if the common
ancestry of two forms has to be traced back to a period be-
yond the late pre-historic times (or beyond any other arbi-
trary line which is agreed upon), then they are not members
of the same species. Professor Lankester is the first to admit
that the practical application of this as of every other con-
ception of species would very often mean a great deal more
than we can prove, in fact, hypothesis.
It is evident too that Darwin regarded persistence of form
as an important criterion of aspecies. We recognise this in the
definition I have quoted from the “ Origin ” (see p. Ixxviii), and
it is stated with even greater force in the following passage,
where persistence is placed beside other distinguishing marks
of a species and given the pre-eminence. In a letter to Hooker
(October 22, 1864) Darwin says :—‘‘I will fight to the death
that as primrose and cowslip are different in appearance (not
to mention odour, habitat, and range), and as I can now show
that, when they cross, the intermediate offspring are sterile
like ordinary hybrids, they must be called as good species as
aman and a gorilla. The power of remaining for a good long
period constant I look at as the essence of a species, combined
with an appreciable amount of difference.”
It is now necessary to examine in some detail the most
usual conception of a species, a conception based upon
distinguishing structural characters, or diagnosis.
This idea of a species is clearly expressed by Sir William
Thiselton Dyer, when he speaks of the older writers who
* See F, A. Dixey in ‘‘ Nature,” June 19, 1902, p. 169.
t ‘More Letters,” vol. i, p. 252, Letter 179.
( xeii )
employed “the word species as a designation for the
totality of individuals differing from all others by marks
or characters which experience showed to be reasonably
constant and trustworthy, as is the practice of modern
naturalists.” *
This conception of a species is founded upon transition.
Whenever a set of individuals can be arranged, according to
the characters fixed upon by the systematist, in a series
without marked breaks, that set is regarded as a species.
The two ends of the series may differ immensely, may
diverge far more widely than the series itself does from other
series; but the gradual transition proclaims it a single
species. If transitions were all equally perfect of course there
would be no difficulty. But transitions are infinite in their
variety ; while the subjective element is obviously dominant
in the selection of gaps just wide enough to constitute
interspecific breaks, just narrow enough to fuse the species
separated by some other writer,—dominant also in the choice
of the specific characters themselves.? Looking back upon the
interval between Linnzus and Darwin, it seems remarkable
that the mutability of species was not forced upon systematists
as the result of their own labours. It is astonishing that
many a naturalist was not driven by his descriptive work to
the conclusion which Darwin stated to Asa Gray on July 20,
TS SG as an honest man, I must tell you that I have
come to the heterodox conclusion, that there are no such
things as independently created species—that species are only
strongly defined varieties.” {
For, as I have said above, every describer of species made
continuity and transition in characters the test of a variety,
discontinuity the test of a separate species. And in difficult
cases no two of them agreed in their conclusions. Many
passages in Darwin’s correspondence convincingly prove how
essential an element is this continuity, and how inevitable
B jls Cop. O05
+ How important this choice may be is well shown by Karl Jordan in
‘* Novitates Zoologice,”’ vol. iii, Dec. 1896, pp. 428-430. Characters are
subject to independent variation as well as correlated variation. Hence
there may be the widest discrepancy between the transitions constructed
by naturalists making use of different characters.
+ ‘Life and Letters,” vol. ii, p. 79.
( xelll )
is the dominance of the subjective element. Thus he writes
about his descriptive work on Cirrhipedes to Hooker, October
12, 1849 :—‘T have of late been at work at mere species
describing, which is much more difficult than I expected, and
has much the same sort of interest as a puzzle has; but I
confess I often feel wearied with the work, and cannot help
sometimes asking myself what is the good of spending a week
or fortnight in ascertaining that certain just perceptible
differences blend together, and constitute varieties and not
species. As long as | am on ahatomy I never feel myself in
that disgusting, horrid, ewt bono, inquiring humour.” *
On another occasion, when Darwin was anxious to ascertain
the ‘‘ close species” in the North American Flora, and wrote
for information to Asa Gray, he frankly adopted the sub-
jective criterion in order to explain exactly what he meant.
He wrote, June 8, [1855]:—‘“The definition I should give
of a ‘close species’ was one that you thought specifically dis-
tinct, but which you could conceive some other good botanist
might think only a race or variety ; or, again, a species that
you had trouble, though having opportunities of knowing it
well, in discriminating from some other species,” +
Asa Gray’s reply is also very interesting from the same
point of view. He-wrote, June 30, 1855 :—“‘Those thus
connected” [he had bracketed the “ close species” in a list
of the Flora], “ some of them, I should in revision unite under
one, many more Dr. Hooker would unite, and for the rest it
would not be extraordinary if, in any case, the discovery of
intermediate forms compelled their union.” {
Darwin was evidently in high spirits when he wrote the
following passage which bears on the same subject. The
“Origin”? had been published on November 24, 1859, and
the whole edition of 1250 copies sold on the day of issue.
On November 29 he wrote to Asa Gray :—‘ You speak of
species not having any material base to rest on, but is this
any greater hardship than deciding what deserves to be called
a variety, and be designated by a Greek letter? When I
* “TVife and Letters,” vol. i, p. 379.
+ Ibid., vol. ii, p. 64.
+ ‘More Letters,” vol. i, p. 421, Letter 324.
( “uxcive #>)
was at systematic work I know I longed to have no other
difliculty (great enough) than deciding whether the form was
distinct enough to deserve a name, and not to be haunted with
undefined and unanswerable questions whether it was a true
species. What a jump it is from a well-marked variety, pro-
duced by natural cause, to a species produced by the separate
act of the hand of God! But I am running on foolishly.
By the way, I met the other day Phillips, the paleontologist,
and he asked me, ‘ How do you define a species?’ I answered,
‘T cannot.’ Whereupon he said, ‘ At last I have found out
the only true definition—any form which has ever had a
specific name !’”’ *
The idea of a species as an inter-breeding community, as
syngamie, is, I believe, the more or less acknowledged found-
ation of the importance given to transition. This will become
clearer from the consideration of a concrete example. The
common black-and-white Danaine butterfly, Amauwris niavius
of West Africa, is represented on the East and South-East
Coasts by a very similar butterfly, distinguished by the greater
size of the largest white patch, and of the white spot in the
cell of the fore-wing. Both forms are very constant in the
areas over which they were known, and on these constant
easily recognisable characters the eastern butterfly was
described as a distinct species under the name of A. domini-
canus. Aurivillius, however, in his valuable Catalogue refuses
to recognise this latter as a distinct species, and considers it
as the dominicanus variety of niavius. Through the kind-
ness of Mr, C. A. Wiggins and Mr, A. H. Harrison, the Hope
Department has recently been presented with an exceedingly
fine series of butterflies from both east and west of the
northern shores of Lake Victoria Nyanza. These have been
carefully studied by Mr. 8. A. Neave, B.A., of Magdalen
College, Oxford, who finds that the typical néaviws occurs in
great abundance to the west of the lake, while on the east he
meets, in both collections, with varieties beautifully inter-
mediate between it and dominicanus. These varieties,
occurring precisely in the zone where the eastern form meets
the western, complete for the systematist the transition which
* ** More Letters,” vol. i, p. 127, Letter 79.
(= xev) >)
renders dominicanus a variety of niavius and not a distinct
species. But it is clear that they do more than this; they
make it almost certain that the two forms freely interbreed,
and constitute but a single syngamic community.
This is one of the remarkably clear examples. In many
cases we know the transition, but the extremes are not sorted
out in different parts of the total area of distribution. Never-
theless if complete enough the transition of forms on the
same area always raises the strong presumption that we are
dealing with a syngamic community.
Probably the most remarkable series of transitional varieties
ever depicted is that shown in the eleven quarto plates of
the last part of Monsieur Charles Oberthiir’s great “ Etudes
d’Entomologie,” entitled “ Variation des Heliconia thelxiope et
vesta”’ (Rennes, February, 1902).
The method of diagnosis, at its clearest and simplest, is
always consistent with, and often strongly suggests, an under-
lying syngamy. There are, however, numberless examples
belonging to various categories in which a rigid adherence
to diagnosis cannot avail. In these cases the systematist
frankly appeals to syngamy or synepigony as decisive; and
if he has not direct: proof of the existence of either of these,
indirect evidence is, at any rate provisionally, regarded as
sufficient.
I. Dimorphism, Polymorphism :—In an _ ever-increasing
number of examples an assemblage of individuals is regarded
as a single species, although split up into two or more widely
different and sharply separated groups, between which transi-
tional varieties are excessively rare or even unknown. For
instance, the extremely abundant, widely distributed butterfly
Limnas chrysippus wncludes among other forms one in which
the black-and-white tip is wanting from the fore-wing, the
dorippus (=klugit) form. This variety is sharply cut off from
the type form. Although faint traces of a former white bar
can be made out in dorippus, I have never seen, among
thousands of individuals, the material out of which a good
transitional series between it and chrysippus could be con-
structed. In this case the evidence of syngamy is strong and
complete ; for Col. Yerbury has recorded the fact that the
( xevi )
two forms certainly occur im copuld.* But if this evidence
were wanting there would still be strong presumptive evi-
dence that the forms are associated by syngamy and synepigony.
Thus, so far as our knowledge extends, dorippus occurs as the
only form in certain parts of N.E. Africa alone. From this,
its metropolis, dorippus spreads on all sides, its individuals
existing intermingled with those of chrysippus, becoming less
and less numerous until they finally die out. Thus if we
trace the two forms eastward we find them both abundant
at Aden; further east, at Karachi, dorippus is well known,
but very scarce as compared with chrysippus; in Southern
India it is a great rarity, if indeed it is known at all on the
mainland ; in Ceylon a single specimen was captured by Col.
Yerbury in 1891, and since then others have been taken.
Further east I have never heard of a specimen. Similarly
when it is traced southward in Africa, dorippus is dominant
in the coast strip of British East Africa, where it constitutes
about three-quarters of the total number of individuals.
Further to the south it becomes rarer and rarer, until in
Natal and the Cape, if it occurs at all, it is even rarer than
in Ceylon.{ Such a distribution is consistent with the inter-
pretation that dorippus and chrysippus are two forms in one
syngamic community. It is difficult on any other hypothesis
to account for the facts which we observe on the outskirts of
* Speaking of his experience at Aden, Col. Yerbury says: ‘‘I have
taken them [the forms of chrysippus] i coitw in every possible com-
bination.” (Journ. Bomb. Nat. Hist. Soc., vii (1892), p. 209.)
+ See Major N. Manders, F.Z.S., in Journ. Bomb. Nat. Hist. Soc., xiv
(1902), p. 716 :—
‘The first specimen of this insect [dorippus=klugit] in Ceylon was
captured by Lieut.-Colonel Yerbury at Trincomalie, April 15th, 1891. . .”
Of five or six more recent examples Major Manders writes, ‘‘ These speci-
mens were captured by Mr. Pole at Puttalam on the east coast and Ham-
bantotte on the south coast in the dryest and perhaps most arid portion of
the island. It is evidently widely distributed in the desert portion of the
island and is possibly not uncommon.”
‘The distribution of this insect in India cannot yet be fully known ; it
is rare in Canara, but is not yet reported from the plains of the Deccan,
or Southern India, so far as I am aware, though it probably exists.” The
occurrence of dorippus at Bombay, Kutch, and Sind had been previously
published by Major Manders and the late Mr. de Nicéville in Journ. As.
Soc. Bengal, vol. Ixviii, Pt. ii, No. 3, 1899, p. 170.
~ Mr. Roland Trimen tells me that he knows of only three South-
African dorippus :—two from Durban and one from Pretoria. The latter
and one of the former were taken by Mr. W. L. Distant (Ann. Mag. Nat.
Hist. (7), vol. i, 1898, pp. 48, 49).
Caxcvin =)
the range of dorippus—the occasional appearance of single
individuals in the swarms of the type form. And if the two
are syngamic on the outskirts, the gradual transition in pro-
portions towards the metropolis of dorippus suggests that
they are syngamic throughout. Common as the species is—
probably the commonest butterfly in the world,—the evidence
from epigony has never been obtained, although from the
point of view of heredity the investigation promises to be of
the deepest interest.
The remarkable forms of the females of the Papilio merope
group already alluded to afford another excellent example,
although in this case good transitional series can be constructed.
The evidence of syngamy was first obtained by Hewitson (see
p- Ixxxvii), but is now well known. The evidence of epigony has
fortunately been obtained in 1902 and again within the last
few weeks by one of our Fellows at Durban, Mr. G. F. Leigh.
Eggs from a female of the commonest cenea form yielded a
synepigonic group, including a large majority of forms like the
parent, but also examples of the very different hippocoon form.
Still more recently seven eggs from the rarest of the forms,
trophonius, produced, in addition to males, two females of the
cenea variety, and not one resembling the parent.
These differences, although only of colour and pattern,
greatly exceed those between ordinary close species. When we
deal with other kinds of dimorphism or polymorphism involv-
ing important structural differences, such as those of the social
Hymenoptera and Neuroptera, the discriminating characters
between nearly related genera are commonly equalled or
exceeded.
IL. Seasonal Dimorphism :—In certain exceedingly interest-
ing examples of dimorphism the relation between the forms is
epigonic and not syngamic ; for rare and occasional inter-breed-
ing is not syngamy. I refer to the most strongly-marked cases
of seasonal dimorphism in butterflies, especially the wonderful
examples proved to be epigonic by Guy A. K. Marshall. In
some of the forms the two seasonal phases were not even
regarded as closely related species. In these extraordinary
cases, where the widest difference in colour and pattern exists,
in combination with others which are far more deep-seated,
PROG. ENT. SOC. LOND., v. 1903. H
(, Exc ey)
IT urged upon Mr. Marshall that the few recorded examples
of capture or observation 77 coitw were insufficient evidence
of specific identity, and that nothing short of epigony would
suffice.
In seasonal dimorphism, in the dimorphism of social insects,
and doubtless in a large proportion of other examples, it is
probable, indeed often certain, that the different forms are
produced in response to some stimulus which acts at a speci-
ally susceptible period of the life-history ; but from the point
of view of the systematist the mature individuals can only be
known as forms which, structurally widely different, must
nevertheless be placed within the limits of a single species.
The investigation of the probable physiological causes of differ-
ence is, however, of the utmost importance from other points
of view. Altogether apart from*its bearing upon dimorphism,
the effect of individual susceptibility to stimulus requires
treatment in a separate category.
Il. Individual Modification :*—One of the most striking
developments of recent years has been the growth in the
number of these very cases in which an individual animal
or plant has been rendered by natural selection susceptible
to some stimulus associated with each one of its possible
normal environments. Every individual of such species
comes into the world with two or more very distinct and
very different possibilities before it, each of which will be
realised only in the appropriite environment—realised as the
response to some stimulus provided by the environment itself.
We can see clearly that this idea was in Darwin’s mind,
although there were then but few facts which pointed in
its direction. Thus in Schmankewitsch’s experiments
Crustacea of the species Artemia salina were described as
gradually changing in the course of generations, as the result
of a progressive freshening of the water in which they were
kept, until they took on the characters of the genus Bran-
chipus. On this subject Darwin wrote to Karl Semper,
February 6, 1881 :—‘‘ When I read imperfectly some years
* “A structural change wrought during the individual’s lifetime (or
acquired), in contradistinction from variation, which is of germinal origin
(or congenital).”” Dict. of Phil. and Psych., ed. by J. Mark Baldwin,
New York and London, vol. ii, 1902, p. 94.
(e excixe™)
ago the original paper, I could not avoid thinking that some
special explanation would hereafter be found for so curious a
case. I speculated whether a species very liable to repeated
and great changes of conditions might not assume a fluctuating
condition ready to be adapted to either conditions.” *
I venture to express the prediction that this class of cases,
already very numerous, will hereafter be immensely enlarged,
and will become especially important in the vegetable king-
dom.t Although Hooker at one time took the opposite side,
and thought that plants were never ‘‘changed materially by
external conditions—except in such a coarse way as stunting
or enlarging,” | Darwin considered that ‘“ physical conditions
have a more direct effect on plants than on animals.” § Un-
doubtedly the view at the time was that of Buffon, the idea of
an operation of the environing forces almost as direct as
those which produce the weathering of rocks or the whitening
of an exposed flint. But it is probable that the more in-
timately we know of the conditions of plant-life, the more
fully it will be recognised that all such changes are adaptive.
* “More Letters,” vol. i, p. 391, Letter 303.
+ See ‘‘Stimulus and Mechanisin as Factors in Organisation” by J.
Bretland Farmer, F.R.S. (the New Phytologist, vol. ii, Nos. 9 and 10,
Novy. and Dec. 1903), Professor Farmer speaks of the probable prevalence
in the plant-world of ‘‘a constant specific mechanism that is able to be
actuated in different ways by different kinds of stimuli.” Although for
the purpose of his paper Professor Farmer is concerned with the train of
physico-chemical sequences which is set going, utility orno utility, when-
ever the mechanism of an individual is stimulated, he fully admits that
the mechanism itself has come to be a character of the species by the oper-
ation of natural selection. ‘‘ Naturally,” he says, ‘‘only those species
whose inner character expressed itself in making these ‘suitable’ adjust-
ments to the environment were able to survive.”
Toward the close of his paper Professor Farmer seems to bring the con-
siderations that have regard to the species into somewhat unnecessary
conflict with those that have regard to the individual. Thus he says
that ‘‘current literature still teems with teleo’ogical explanations that
realiy explain nothing, but rather bar the way of scientific enquiry.”
A properly loaded, well-constructed modern gun goes off, for disadvan-
tage no less than for advantage, when its trigger is pulled ; but the very
existence of the gun depends upon a long succession of past stages, eich of
which was more advantageous than its predecessor. The recognition of
this history does not bar the way of enquiry, but rather stimulates and
suggests a searching and intelligent study of the latest mechanism with
all its intricacy.
+ See the letter from Hooker to Darwin, March 17, 1862, in ‘* More
Letters,” vol. i, p. 197.
§ See the letter from Darwin to Lyell [June 14, 1860], ‘‘ Life and
Letters,” vol. ii, p. 319.
(oh)
I will mention merely by way of illustration, that my attention
has been called in recent years to the dwarfing effect of the
prevalent south-western winds on the vegetation of the
exposed chalk downs of the Isle of Wight. It has occurred
to me as a mere suggestion, but one worth investigating, that
the effect of wind upon a tall flower-head might be such as to
render less easy and less frequent the visits of insects. If
this were so, it would perhaps explain why certain species of
entomophilous plants liable to grow in such situations have
gained a special susceptibility to the stimulus provided by
constant winds during some particular period of growth.
The absence of this stimulus would also correspond to a
condition in which the plants would gain in the conspicuous-
ness brought about by increased height.
The further growth of a class already proved to be large,
would play havoe with a definition of species rigidly based
upon discriminating structural characters alone.
IV. Geographical Races or Sub-Species :—If we depend upon
unaided diagnosis there is no means of discriminating between
species and those sub-species of which the whole mass of in-
dividuals are distinguished by recognisable characters. Here
again the mere beginning of the difficulty is in sight; for as
museums recognise more and more the necessity for long series
of specimens with exact geographical data, so will the compara-
tively simple conception of the single species be replaced again’
and again by the far more complex but much truer idea of
sub-specific groups still fused by syngamy into a single species,
but as it were trembling on the edge of disruption, ever ready,
by the development of pronounced preferential mating or by
the accumulated incidental effects of isolation prolonged beyond
a certain point, to break up into distinct and separate species.
V. Results of Artificial Selection ;—These obvious difi-
culties encountered by a mechanical adherence to defini-
tion by diagnosis naturally lead to the consideration of
the further difficulties presented by domestic races of
animals and plants. The wide structural differences be-
tween the forms accumulated by human selection greatly
impressed Darwin. Thus he wrote to Hooker, September
8, [1856]:—* By the way, I have been astonished at the
( ci)
differences in the skeletons of domestic rabbits. I showed
some of the points to Waterhouse, and asked him whether he
could pretend that they were not as great as between species,
and he answered, ‘ They are a great deal more.’ How very odd
that no zoologist should ever have thought it worth while to look
to the real structure of varieties. ...”’* Then again, the differ-
ences between many of our domestic breeds, and between them
and the nearest wild species, are, as is well known, generic rather
than specific. Why do we not consider such races to be of
different species and genera? Because of the criterion sug-
gested by Lankester; because we have reason to believe in
their descent from common parents within the historic period ;
because, in spite of their wide differences, they are still
syngamic.
What is the practical bearing of these criticisms upon the
definition of species by diagnosis and diagnosis alone? The
systematist, confronted by his series of specimens in a museum
cannot do otherwise than arrange them in groups which he
will describe and name as species. But much would be gained
if he admitted at the outset that his conclusions are provisional,
if he said with Dr. Karl Jordan, “The actual proof of specific
distinctness the systematist as such cannot bring; ... we
work, or we ought to work, with the mental reservation that
the specific distinctness of our species nove deduced from
morphological differences will be corroborated by biology.” T
The advantage of thisattitudeis obvious. Work would goon
as at present. Powers of acute observation and good judg-
ment would still furnish descriptions of species to be hereafter
confirmed, or confirmed at the time by observation and experi-
ment upon the living material. But the systematist would
not only receive our gratitude for the performance of these
important and necessary duties: he would also be seeking in
every direction for the evidence of syngamy and of epigony.
The museum would become a centre for the inspiration of
researches of the highest interest to the investigator himself,
of the greatest importance to the whole body of naturalists.
* “¢ More Letters,” vol. ii, p. 210, Letter 543.
+ “Novitates Zoologice,” vol. iii, Dec. 1896, pp. 450, 451. I here
desire to express my indebtedness to the author of this learned and valuable
paper.
(Ee ))
We now turn to the consideration of interspecific sterility,
which many have supposed to be an _ infallible criterion.
Huxley himself felt this so stron gly that he was, in consequence,
never able to give his full assent to natural selection.
The grounds of his objection were the subject of prolonged
correspondence with Darwin. In order to prove that natural
selection has produced natural species separated rigidly, as he
believed, by the barrier of sterility, Huxley maintained that we
ought to be able to produce the same sterility between our artifi-
cially selected breeds ; and until this had been done he could
not thoroughly accept the theory of natural selection. This
objection he expressed, or implied, in many speeches and
writings up to within a few months of his death. One of
the simplest statements is contained in a letter to the
late Charles Kingsley. Huxley wrote, April 30, 1863,
“ Their produce [viz. that of Horse and Ass] is usually
a sterile hybrid.
“So if Carrier and Tumbler, e. g., were physiological species
equivalent to Horse and Ass, their progeny ought to be
sterile or semi-sterile. So far as experience has gone, on the
contrary, it is perfectly fertile—as fertile as the progeny of
Carrier and Carrier or Tumbler and Tumbler.
“From the first time that I wrote about Darwin’s book
in the 7%mes, and in the Westminster, until now, it has been
obvious to me that this is the weak point of Darwin’s
doctrine. He has shown that selective breeding is a vera
causa for morphological species; he has not yet shown it a
vera causa for physiological species.
“ But I entertain little doubt that a carefully devised system
of experimentation would produce physiological species by
selection—only the feat has not been performed yet.”*
It was against this same view, as expressed in Huxley’s
‘Lectures to Working Men” in 1863, that Darwin argued
with convincing force in many letters. The main facts with
which he confronted Huxley again and again were the
artificially selected races of certain plants which are sterile
inter se. The position is clearly expressed in the following
amusing, vehement passages from two letters :—
* «Tife and Letters of Thomas Henry Huxley,” vol. i, p. 239.
( ciii_ )
“ Dec, 18, [1862.]
“Do you mean to say that Gartner lied, after experiments
by the hundred (and he a hostile witness), when he showed
that this was the case with Verbascwm and with maize (and
here you have selected races): does Kolreuter lie when he
speaks about the varieties of tobacco? My God, is not the
case difficult enough, without its being, as I must think,
falsely made more difficult? I believe it is my own fault—
my d
fuss about these most careful experiments.” *
d candour: I ought to have made ten times more
‘*{ Jan. ]10, [1863.]
“Tn plants the test of first cross seems as fair as test of
sterility of hybrids, and this latter test applies, I will maintain
to the death, to the crossing of varieties of Verbascwm, and
varieties, selected varieties, of Zea. You will say, Go to
the Devil and hold your tongue. No, I will not hold my
tongue; for I must add that after going, for my present
book [Variation under Domestication], all through domestic
animals, I have come to the conclusion that there are almost
certainly several cases of two or three or more species blended
together and now perfectly fertile together. Hence I
conclude that there must be something in domestication,—
perhaps the less stable conditions, the very cause which
induces so much variability,—which eliminates the natural
sterility of species when crossed. If so, we can see how
unlikely that sterility should arise between domestic races.
Now I will hold my tongue.” fT
Darwin made attempts to ‘‘produce physiological species
by selection,” and thus meet his friend’s criticism. He
thought out and suggested a plan of experiment to W. B.
Tegetmeier,{ and gave a brief account of the scheme to
Huxley, December 28, [1862]:—“I have
[Tegetmeier] the result of my crosses of the birds which he
proposes to try, and have told him how alone I think the
experiment could be tried with the faintest hope of success—
namely, to get, if possible, a case of two birds which when
given him
* “More Letters,” vol. i, p. 230, Letter 156.
+ Ibid. vol. i, pp. 231, 232, Letter 157.
+ Ibid. vol. i, pp. 223, 224, Letter 153, [1862, Dec. ] 27.
(cia)
paired were unproductive, yet neither impotent. For instance,
I had this morning a letter with a case of a Hereford
heifer, which seemed to be, after repeated trials, sterile with
one particular and far from impotent bull, but not with
another bull. But it is too long a story—it is to attempt to
make two strains, both fertile, and yet sterile when one of one
strain is crossed with one of the other strain. But the
difficulty . . . would be beyond calculation.” *
The experiment was evidently unsuccessful,—perhaps was
never seriously undertaken,—and a few years later Darwin
added the following postscript to a letter to Huxley, January 7
[1867].
‘““P.S.—Nature never made species mutually sterile by
selection, nor will men.” t
This was probably only an offhand expression of opinion,
not intended to be taken seriously. An altogether hopeless
attitude would not be reasonable until the suggested scheme
had been applied many times, and in several parts of the
animal and vegetable kingdoms.
But the positive results demanded by Huxley, even if
obtained, would by no means justify his far-reaching
conclusions, If the barrier of sterility were thus artificially
produced, we should be very far from the proof that its exist-
ence in nature is due to the same kind of cause, viz. selection.
If Darwin was right in his controversy with Wallace, if
‘““Nature never made species mutually sterile by selection,”
the suggested experiment would merely do by artificial selection
what is not done by natural selection.
It is by no means difficult to understand the mutual sterility
which is usual between natural species as an incidental result
of their separation by asyngamy for a long period of time.
In the process of fertilisation a portion of a single cell nucleus
from one individual fuses with a portion from another in-
dividual, the two combining to form the complete nucleus of
the first cell of the offspring, from which all the countless
cells of the future individual will arise by division. Each
part-nucleus contains the whole of the hereditary qualities
* “* More Letters,” vol. i, pp. 225, 226, Letter 154.
+ Ibid. vol. i, p. 277, Letter 197.
xs)
received from and through its respective parent, and must
therefore be of inconceivable complexity. We can only speak
in generalities about processes of which so little is known,
but we cannot be wrong in assuming that sterility is some-
times due to the fact that the complex architecture of one
part-nucleus fails in some way to suit the equally complex
structure of the other. The individuals of an inter-breed-
ing community form a biological whole, in which selection
inevitably keeps up a high standard of mutual compati-
bility between the sexual nuclei. Individuals whose sexual
nuclei possess a structure which leads to sterile combinations
with those of other individuals are excluded from contributing
to the generations of the future. As soon, however, as a
group of individuals ceases, from any reason, to breed with
the rest of the species, there is no reason why the compati-
bility of the sexual nuclei of the two sets should be retained.
Within each set, selection would work as before and keep
up a high standard of compatibility ; between the sets, com-
patibility would only persist as a heritage of past selection,
gradually diminishing as slight changes of structure in either
or both of the sets rendered them less and less fitted to
produce fertile combinations.*
It is probable that of all the nice adjustments required in
the living organism, the mutual adjustment of these incon-
ceivably complex part-nuclei is the most delicate and precise.
Now, delicately adjusted organs, such as those of sight, rapidly
become incapable of performing their functions when in any
species they have been withdrawn from the operation of
natural selection ; similarly it is suggested, that the adjustment
of sexual nuclei to each other would sooner or later give way
* T must guard against the inference that the only explanation of
sterility is here set forth. It is indeed maintained that incompatibility
of the sexual part-nuclei is the inevitable outcome of enduring asyngamy,
and is the almost certain cause of the sterility of hybrids. And it may
be suggested that sterility is a result of the combination of two incom-
patible germ-plasms in the sexual cells of the hybrid. When the
incompatibility is not strongly marked we can understand how such
sexual cells may be capable of fertile fusion with the cells of either
parent, but not with those of another hybrid.
But short of these ultimate effects it must not be forgotten that there
are many obscure factors of asyngamy—causes of various kinds which
interfere with the fusion under normal conditions or entirely prevent the
meeting of the sexual cells.
(eva)
when no longer sustained by selection, If, then, mutual
fertility be the result of unceasing selection, and mutual
sterility the inevitable, even if long-postponed, consequence
of its cessation, it is obvious that Huxley’s difficulty is solved,
while his suggested experimental creation of sterility by
selection would not reproduce any natural operation: it would
afford a picture of a natural result but would be produced
in an unnatural way. This criticism of Huxley’s contention
was advanced by the present writer three years ago,* the
final conclusion being stated in the paragraph printed
below :—
“Tf, then, we cannot as yet reproduce by artificial selection
all the characteristics of natural species-formation, but can
only imitate natural race-formation, we can nevertheless
appreciate the reasons for this want of success, and are no
more compelled to relinquish our full confidence in natural
selection than we are compelled to adopt a guarded attitude
towards evolution because our historical records are not
long enough to register the change of one species into
another.” f
It was therefore with intense interest and pleasure that I>
read the following sentences in a letter written by Darwin to
Huxley, Dee. 28, [1862]—sentences which show that criticism
practically identical had been made by the illustrious naturalist
nearly forty years earlier.
‘“We differ so much that it is no use arguing. To get the
degree of sterility you expect in recently formed varieties
seems to me simply hopeless. It seems to me almost like
those naturalists who declare they will never believe that one
species turns into another till they see every stage in
progress.” {
After reading, in the first volume of ‘More Letters,’’ the
often-repeated refutation of Huxley’s objection so clearly and
strongly expressed in letters received by the objector himself,
it is surprising that no effect was produced, and that reference
should have been nearly always made to this supposed flaw in
the theory of natural selection, whenever the great compara-
* “The Quarterly Review,” No. 385, January 1901, pp, 368-371.
ds Ch Ps Ol Ls
~ ‘More Letters,” vol. i, p. 225, Letter 154,
( evii )
tive anatomist had occasion to speak or write on the broader
aspects of biological inquiry.*
Darwin also considered that there was something in the
very conditions of domestication which tended to promote
fertility between races and even between distinct species.
Thus he followed Pallas in believing that the domestic dog
has been derived from more than one wild species, although he
did not trace existing differences to this cause but to artificial
selection.t However, as regards the origin of the dog, “the
evidence is, and must be, very doubtful,” as he wrote to Lyell,
August 11, [1860]. The fact which Darwin “ considered the
most remarkable as yet recorded with respect to the fertility of
hybrids,” was the fertility of the offspring of the Common and
Chinese Goose, originally described by Eyton, and confirmed by
Goodacre and by Darwin himself. ‘The two species of goose
now shown to be fertile inter se are so distinct that they have
been placed by some authorities in distinct genera or sub-
genera.” }
Another interesting and excéedingly difficult experiment in
hybridisation has been carried through by the Rev. P. St. M.
Podmore, F.Z.S., who in Sept. 1899, after numerous failures,
succeeded in rearing a healthy male hybrid between the
Ring Dove (Columba palumbus) and the domestic pigeon.
On May 27, 1903, this male was mated with a Blue Homer
hen, which produced healthy offspring. §
* For several instances see Poulton’s ‘‘ Charles Darwin and the Theory
of Natural Selection,’ Lond. 1896, pp. 124-141.
+ “Though I believe that our domestic dogs have descended from
several wild forms, and though I must think that the sterility, which
they would probably have evinced, if crossed before being domesticated,
has been eliminated, yet I go but a very little way with Pallas & Co. in
their belief in the importance of the crossing and blending of the
aboriginal stocks.
* * * * * * *
‘Although the hound, greyhound, and bull-dog may possibly have
descended from three distinct stocks, I am convinced that their present
great amount of difference is mainly due to the same causes [artificial
selection] which have made the breeds of pigeons so different from each
other, though these breeds of pigeons have all descended from one wild
stock ; so that the Pallasian doctrine I look at as but of quite secondary
importance.”
**More Letters,” vol. i, pp. 127, 128, Letter 80, to Lyell, Oct. 31,
[1859].
+ “‘ Life and Letters,” vol. ili, p. 240.
§ ‘*The Zoologist,” Nov. 1903, p. 401.
( eviii_ )
A comparison between the difficulty of producing such a
cross and that of obtaining hybrids between the Ring Dove
and the Rock Pigeon, the ancestor of the domestic breeds,
would probably throw much light on the Pallasian hypothesis.
If the view here proposed be sound—that syngamy lies
behind, and is at least provisionally implied in the transition
which means so much to the systematist, and is his only real
evidence when the structural test breaks down, the conclusion
is suggested that the real interspecific barrier is not sterility
but asyngamy. Nevertheless, as argued on pages civ-cvi,
asyngamy will infallibly lead to sterility, although the result
may be long delayed. This latter view, which was that of
Darwin, is the exact opposite of the “ physiological selec-
tion” of Romanes, in which sterility is supposed to arise
spontaneously, asyngamy being not the cause, but the
consequence.
Asyngamy may be brought about in various ways, of which
the most obvious is geographical separation. But asyngamy
is by no means the necessary result of geographical discon-
tinuity or asympatry. Thus Darwin considered that there
is regular inter-breeding between Madeiran and continental
birds of the same species. He wrote to Hooker, August 8
[1860]: “I do not think it a mystery that birds have not
been modified in Madeira. Pray look at p. 422 of Origin
[ed. iii]. You would not think it a mystery if you had seen
the long lists which I have (somewhere) of the birds annually
blown, even in flocks, to Madeira. The crossed stock would
be the more vigorous.” * An even more striking case is that
of Pyrameis cardui, which ranges over nearly the whole world.
The singular absence of local geographical races in this
abundant butterfly is almost certainly due to the astonishing
powers of dispersal which enable intermittent syngamy to
prevail over the whole vast area of its distribution.
An interesting and curious cause of persistent asyngamy
is the ‘‘ Mechanical Selection” so thoroughly explained and
abundantly illustrated by Karl Jordan.f The complex genital
armature of Lepidoptera is during syngamy kept constant by
* “More Letters,” vol. i, pp. 487, 488, Letter 370.
+ de p. 518-522,
(seix )
unceasing selection. Comparatively brief isolation of a group
of individuals may lead to a departure from the specific type
of apparatus prevalent in other areas, and may thus mechanic-
ally prevent syngamy if from any cause members of the
group became again sympatric with those of the parent
species.
A very different but exceedingly interesting origin of
asyngamy is suggested by observations which support the
conclusion that varietal forms may show a tendency towards
preferential inter-breeding.
H. W. Bates believed that he had strong evidence for the
existence of this tendency in the races of certain tropical
American butterflies. He stated this in his epoch-making
paper on the butterflies of the Amazon valley,* and it is
interesting to observe in the published letters how Darwin
instantly fixed upon the point and tried to elicit the data
upon which the conclusion was formed. Thus he wrote to
Bates, Noy. 20 [1862] :—“ No doubt with most people this
[viz. the interpretation of Mimicry] will be the cream of the
paper; but I am not sure that all your facts and reasonings
on variation, and on the segregation of complete and semi-
complete species, is not really more, or at least as valuable, a
part. I never conceived the process nearly so clearly before ;
one feels present at the creation of new forms. I wish,
however, you had enlarged a little more on the pairing of
similar varieties ; a rather more numerous body of facts seems
here wanted.” +
Then a few days later we find Darwin still thinking of the
subject, and writing to Hooker [1862, Nov.] 24:—“TI have
now finished his [Bates’] paper . . .; it seems to me admir-
able. ‘lo my mind the act of segregation of varieties into
species was never so plainly brought forward, and there are
heaps of capital miscellaneous observations.” {
He also again wrote to Bates, probably on the following
day, Nov. 25 [1862 4], asking for the solid facts which are so
greatly wanted :—
‘Could you find me some place, even a footnote (though
* Trans. Linn. Soc., vol. xxiii (1862), p. 495.
+ ‘‘ Life and Letters,” vol. ii, p. 392.
‘* More Letters,” vol. i, p. 214, Letter 147.
( ex )
these are in nine cases out of ten objectionable), where you
could state, as fully as your materials permit, all the facts
about similar varieties pairing
at a guess how many you
caught, and how many now in your collection? I look at this
fact as very important ; if not in your book, put it somewhere
else, or let me have cases.” *
Remembering that Mr. Roland Trimen, F.R.S., had expressed
the same opinion as the result of his wide and long experience
of South African butterflies, [ asked him if he would kindly
furnish me with a statement. His reply, dated Dec. 28, 1903,
is as follows :—
“* Dec. 28, 1908.
“T have noticed the tendency of the sexes of a variety to
pair together rather than with other varieties in the numerous
cases of captured pairs sent to me by correspondents in South
Africa, and sometimes in cases of the same kind which occurred
to myself when collecting. The species which particularly
attracted my notice in this way during my visit to Natal was
Hypanis acheloia (= Gétzius, Herbst, part), which is curiously
variable on the underside, from pale creamy to deep chocolate.
I did not know of its seasonal variation at the time, but I was
in Natal just at the change of season from wet to dry, when
the intermediate gradations were about, and I was struck with
the close resemblance of the sexes in pairs that I caught. I
am sorry to have nothing more definite to give on this head ;
it is a point much requiring exact and prolonged observation.”
Mr. Trimen furthermore entertains no doubt that much, if
not all, of the material upon which he based the conclusion
that the individuals of the same race tend to interbreed,
exists, distinctively labelled, in the South African Museum,
at Cape Town. It is greatly to be hoped that collectors will
in future carefully label all specimens captured im cottu, and
that the fact will be recorded on the labels in museums and
in private collections. It is tantalising to reflect upon the
number of interesting and important questions which could
be now decided if this practice had prevailed during the past
fifty years. The question of the possible origin of species
* More Letters,” vol. i, p. 215, Letter 148.
(Ga!)
from races by preferential syngamy is of such high import-
ance that we may confidently hope that the attention here
directed to the question, and especially the quotation of
Darwin’s letters to Bates, may lead to that “exact and
prolonged observation,’ accompanied by careful records, with-
out which a safe decision cannot be reached. In the
meantime the decided impressions of two such naturalists as
H. W. Bates in South America and Roland Trimen in South
Africa render it in every way probable that the conclusion
will be established on a firm foundation.*
It is also possible that asyngamy may be brought about by
the breaking of what we may call “a syngamic chain.” In the
case of large and widely-distributed interbreeding communities,
it is an open question whether syngamy would freely take
place between the most distant of the outlying sections if
directly brought into contact, and whether, even if syngamy
prevailed, there wotld be any diminution in fertility.
Limnas chrysippus, perhaps the commonest butterfly in the
world, forms a probably continuous syngamic chain stretching
from the Cape of Good Hope at least as far as Southern China.
It is even reported from Japan, The far Eastern forms are
readily distinguishable by the greater size of a single white
spot, giving quite a different appearance to the fore-wing.
If pup or eggs were transferred from Hong-Kong or Macao
to South Africa, would the perfect butterflies freely interbreed
* Dr. T. A. Chapman sends me the following interesting and suggestive
note :—
‘‘T met lately with a curious instance that deserves following up, of
some bearing on the question of selective mating of varieties.
‘*T saw some broods of P. phlxas lately that differed from each other,
but each brood was remarkably uniform. There were three broods, all
bred in the same conditions, in a greenhouse (by Mr. Carpenter of
Leatherhead). It seems difficult to explain this, unless both parents of
each brood were very nearly identical.
‘‘Mr. Frohawk, who has bred the species largely, tells me he has
noticed similar facts.
‘*When I bred Acronycta tridens and psi largely, some fifteen or more
years ago, I noticed that each brood had its own pairs, and suggested that
tridens was now trying to break up into separate species just as some
ancestor split into psi, tridens and cuspis.
** Another fact I observed in Acronycta rather bears on the other side
of the question. Of A. strigosa I reared a large brood, which paired
readily and frequently together, but no eggs were laid. I then got some
captured males, which paired with equal readiness with the bred females,
and as a result obtained plenty of fertile eggs.”
(exit, 9%)
with the indigenous forms of chrysippus ? We do not know ;
but it is an experiment well worth trying, and one which
would yield results valuable in many ways. If inter-breeding
did not take place, or if the unions were sterile, then we
should have the interesting case of a single species which
would instantly become two if through any circumstance a
central link dropped out of the chain. Even if chrysippus
yielded negative evidence in this respect, it is highly probable
that other widely-distributed species would, under these cir-
cumstances, fall into two or more groups, each held together
by inter-breeding, and divided from others by asyngamy.
Sterility, if present in any degree, would have been brought
about quite independently of selection ; for in such cases each
link of the chain would be freely syngamic with the links on
either side, and asyngamy or sterility would only be. revealed
by artificially bringing together the widely-separated ends of
the chain.
I cannot but think, therefore, that such experiments made
upon many carefully-selected species would probably bring
important additional evidence to bear upon the controversy as
to whether sterility between species is, as Wallace believes, a
selected quality, or, as Darwin held, an incidental one. The
deep interest of this question is realised when we thus re-
member that the two discoverers of natural selection held
widely different opinions about it. We cannot read the letters
on both sides, printed in the first volume of “ More Letters,”
without realising how deeply this divergence—one of the
principal differences between them—was felt by the two great
naturalists.
This is one of the many reasons for which I plead with
Mr. Roland Trimen for the establishment of tropical bio-
logical stations where work of the kind could be carried on.
Such establishments should be associated with and be under
the control of museums at home, where the experiments
could be directed and the results studied and made available
for all time for the researches of the naturalist. Just as
Harvard has her main Observatory at the University, but also
maintains an outlying institution in the Peruvian Andes,
where certain kinds of research, unsuited to New England,
(" cx)
can be carried on under the most favourable conditions, so
our chief museums should be provided with the means of
establishing temporary stations in the most favourable parts
of the tropics. When I say temporary, I do not refer to the
means, but to the position of the station, which should be
freely movable in response to the call of important problems
as they present themselves for solution in other localities.
Another urgent reason for the establishment of biological
stations is forced upon us by the inadequacy of diagnosis for
the separation of very variable species, such as many of the
African Acreinx. I cordially agree with the view often
expressed to me by my friend Mr. F. A. Heron, that we shall
never reach a secure foundation until synepigonic series have
been obtained on a large scale. To achieve this end a
temporary station would be required. In this way our
museums could receive, and should keep for permanent study,
the whole of the offspring reared from the eggs of a single
parent. If several species were thus represented by one or
more large synepigonic series, we should know what to expect
and what to allow for; and diagnosis in general would gain
the most helpful guidance.
Asyngamy, as regards particular lines of union, has also
been incidentally brought about by certain adaptations
for cross-fertilisation in plants, and such asyngamy has in
some cases persisted long enough to have led to sterility in
greater or less degree. Of all Darwin’s work, that upon
the fertilisation of heterostyled plants threw most light, he
considered, upon sterility between species. As Francis Darwin
has stated, “ He found that a wonderfully close parallelism
exists between hybridisation and certain forms of fertilisation
among heterostyled plants. So that it is hardly an ex-
aggeration to say that the ‘illegitimately ’ reared seedlings are
hybrids, although both their parents belong to identically the
same species. In a letter to Professor Huxley, given in the
second volume [of ‘Life and Letters’], p. 384, my father
writes as if his researches on heterostyled plants tended to
make him believe that sterility is a selected or acquired
quality. But in his later publications, e.g. in the sixth
edition of the ‘Origin,’ he adheres to the belief that sterility
PROC. ENT. SOC. LOND., v. 1903. I
(cxv >)
is an incidental rather than a selected quality. The result of
his work on heterostyled plants is of importance as showing
that sterility is no test of specific distinctness, and that it
depends on differentiation of the sexual elements which is
independent of any racial difference.” *
The different forms of a heterostyled plant are adapted for
cross-fertilisation by insects, and each individual of each form
is by the same means excluded more or less completely from
fertilisation by another of the same form. In the former case
the sexual cells and the accessory apparatus have been kept
by selection during long generations of syngamy in a high
state of mutual compatibility ; in the latter asyngamy, partial
or complete, has produced a large measure of the sterility
which is its inevitable even if long-delayed result.
This argument has, I admit, carried me much further than
I originally intended, and it will be a pleasure to me if the
following criticism can be overthrown.
If the special adaptation of heterostyled plants for particu-
lar lines of syngamy has incidentally resulted in lessened
fertility, when the unions discouraged by these adaptations
are artificially secured, and in this case without appeal to the
physiologically injurious effects of self-fertilisation, why should
we not similarly explain these effects whenever manifest in
the self-bred 7 offspring of any plant especially adapted for
cross-fertilisation ?
Darwin tells us in the Autobiography that as soon as his
‘attention was thoroughly aroused to the remarkable fact
that seedlings of self-fertilised parentage are inferior, even
in the first generation, in height and vigour to seedlings of
cross-fertilised parentage,” | he entered upon a series of
experiments which lasted eleven years, appearing in 1876 as
‘‘Kittects of Cross and Self-Fertilisation in the Vegetable
Kingdom.” Of this work he wrote in 1881, “the results
there arrived at explain, as I believe, the endless and wonder-
ful contrivances for the transportal of pollen from one plant
to another of the same species.’’§ It is here suggested that
* “Life and Letters,” vol. iii, p. 296.
+ See Francis Darwin on ‘‘The Knight Darwin Law,” Nature, October
27, 1898, p. 680.
t ‘‘ Life and Letters,” vol. i, p. 96. Siibids; Viol. 1p Oe
(Grex s
these injurious results have been not the cause but the con-
sequence of specialisation for cross-fertilisation. In such
plants fertilisation is mainly brought about along the line
for which special adaptation is made: self-fertilisation is
relatively infrequent, often very rare, sometimes perhaps
absent altogether. May not the less successful results have
followed from a condition in which self-fertilisation is but
little tried by the fires of selection? * It would be of much
interest to compare a long series of experiments on the cross-
fertilisation of plants which are habitually self-fertilised, and
on the self-fertilisation of plants in which the adaptations
for cross-fertilisation are made use of in widely different
degrees.
This criticism, should it be sustained, would of course throw
much light upon the case of the Bee Orchis and the numbers
of tropical Orchidacez, etc., which are now known to be
regularly self-fertilising without apparent physiological injury.
It might also have a bearing upon an intrusive set of facts
which must often have weighed upon the minds of naturalists,
as they reflected upon the commonly received hypothesis
that assumes the dangers of continued breeding between
near of kin. A. R. Wallace speaks of these facts in
“Darwinism,” f and I have drawn attention to them in dis-
cussing the meaning of insect migration, although, as will
be seen in the following passage, without any serious doubt
as to the physiological significance of cross-fertilisation. {
““We may well inquire why it should be necessary for such
emigration, with a possible successful issue in colonisation, to
require the services of countless individuals when the importa-
tion of half-a-dozen rabbits or a few specimens of Pieris rape
will, for the naturalist, change the face cf a continent. The
results of these unintentional, or intentional but ill-considered,
experiments do indeed shake the belief in the paramount
necessity for crosses and the dangers of in-and-in breeding ;
but the end is not yet, and the teeming colonies which have
arisen from such small beginnings may in time vanish from
the operation of deep-seated causes. The varied adaptations
for cross-fertilisation and the prevention of in-and-in breeding
* See also A. R. Wallace in ‘‘ Darwinism,” London, 1889, pp. 321-326.
+ p. 326. ~ Trans. Ent. Soc. Lond., 1902, pp. 460-465,
(Svexvil))
are so evident in nature, that we are compelled to believe that
they meet and counteract serious dangers which sooner or
later would menace the very existence of the species. And
among other adaptations it is significant that the instinct
under discussion should lead to the streaming of large popula-
tions, and not of small batches of individuals, from an area of
high-pressure.” *
It is impossible to consider the advantages which may
have favoured cross-fertilisation, if hereafter the generally
accepted physiological necessity turn out to be a delusion.
Brief reference may, however, be made to the special advant-
ages of community which are possible through syngamy alone.
By inter-breeding the favourable variations arising in one
direction are combined with others arising in different direc-
tions ; by the kaleidoscopic changes produced by inter-breeding
more varied results are presented for selection, and the bene-
ficial qualities arising in one part of the mass may quickly
become the heritage of the whole; by inter-breeding excessive
spontaneous variation is checked, and the whole community
of the species advances surely and with stability into adjust-
ment with the progressive changes of the environment.
Weall remember Darwin’s beautifully elaborated metaphor fT
by which the past history of evolution is shown forth in the
form and branching of a great tree. Darwin represented
species by the “green and budding twigs,” and we may
suppose that the leaves stand for individuals, and that syn-
gamy is represented by the contact of leaf with leaf when the
branches sway in the wind. And just as contact may run
through large and small, irregular and compact masses of
leaves, so syngamy binds together groups of varying size and
distribution. So too a mass of foliage breached by a sudden
storm pictures for us the splitting of a syngamic chain into
two species by the disappearance of an intermediate link.
It has been a pleasure to me that the central idea which I
have endeavoured to bring before you should be represented,
I trust without violence to the imagery, by means of ‘the
great Tree of Life, which fills with its dead and broken
branches the crust of the earth, and covers the surface with
fy
its ever-branching and beautiful ramifications.” }
* le. p. 464. + ‘‘Origin of Species,”+1859, p. 129. t Zc. p. 130.
(
cxvli_ )
INDEX.
The Arabic figures refer to the pages of the ‘ Transactions’; the Roman numerals
to the pages of the ‘ Proceedings’
The President’s Address is not separately indexed.
COLEOPTERA.
ABACETUS, 171
abdominalis (Dibolia), 4
(Hyperacantha),
aby ssinica (Megalognatha), oe
Acanthocerus, Txiii, 516, 517,
Acmeeodera, 169, 173
aculeata (Mordella), 175
acuminata (Akis), 175
acuticollis (Dapsa), 172
acutipes (Clueotus), 519
Adalia, 179
Adimonia, 179
adspersula (Acmeodera), 174
{egidium, 515
wneicollis ey
zenescens (Olibrus),
zeneus (Paracymus), 1
wthiops (Apion), 178
affinis (Chrysobothrys), 174
(Dibolia), 5
(Gynandrophthalma), 1
(Homapterus), 176
», (Polydrusus), 167, 176
africana (Lypnea), 14
Agabus, 168, 172
Agapanthia, 169, 178
Agrilus, 167, 174
Airaphilus, 172
Akis, 175
albinus (Anthribus), lii
albipilis (Haplocnemus), 174
albonotatus (Lionychus), 171
aleyoneum (Apion), 178
Aleochara, xxxvii, 170, 172
alpina (Rosalia), xlvii
174
172
=
(2
99
78
99
29
amabilis (Gynandrophthalma), 16
Amara, 166, 169, 171
Amauronia, 167, 174
americana (Chrysomela), 179
ammios (‘Trichodes), 174
Ammeecius, 173
amori (Zonabris), 176
_aquila (Triodonta), 173
| aranea (Arsoa), xx
amplicollis (Corymbites), 174
(Hister), 173
Anaspis, 175
Anchomenus, lviii
Ancistrosoma, 520
angustulus (Agrilus), 174
Anisoplia, 173
Anisorrhynchus, 177
Anomala, 169, 173
Anthaxia, 168, 173
| Anthicus, 175
| Anthobium,
172
Authribus, iii
antillarum (Onthophagus), 510, 511
Aphanisticus, xxxvi, xxxvii, 174
| Aphodius, 167, 169, 170, 173, Gilat
Aphthona, 9, 10, 11, 179
apicipes (Hyperacantha), 18
53 (Longitarsus), 8
Apion, lviii, lix, 87, 167, 178
Apophylia, 22, 23, 24, 25, 27
Apteropeda, 179
ey
arcuatus (Plagionotus), 178
», (Scymnus), 179
areolatus (Perileptus), 171
argentatum (Apion), 167, 178
argus (Epilachna), 179
Arsoa, XX
| ascanii (Lixus), 177
Asemum, Xxxvili
Asida, 166, 167, 169, 175
| Aspidomorpha, xxxviil
Astenus, 172
Atactogenus, 176, 177
| Ateenius, 509, 511, 512, 513
; 178 |
| Ateuchus, 167, 173
ater (Stenopterus), 178
| Athous, 166, 167, 174
| Atomaria, v, Ix
| Atopocerus, 303
atra (Hispa), 179
-atraphaxidis (Clythra), 178
atratus (Philonthus), 172
atripes (Danacia), 174
» (Lagria), 175
Attagenus, 172
Attalus, 174
Aulacophora, 17
aurata (Theodosia), 304, 305
auritus (Exochomus), 179
avellane (Rhynchenus), 178
azare (Pterostichus), 166, 171
beetica (Anisoplia), 173
balyi (Eurycycla), 32
banksiz (Carpophagus), xiii
Baris, 178
barkeri (Aphthona), 9, 10
»» (Exosoma), 25
», (Longitarsus), 7
,, (Weiseana), 16
Barypithes, 167, 176
Basipta, xxxvili
batesi (Psammobius), 514
Bembidium, 166, 170, 171
bicostata (Megalognatha), 30
bicostatus (Deronectes), 168, 172
bidens (Cleeotus), 518
bidentata (Diacantha), 17
Bidessus, 168, 172
biguttata (Titubcea), 178
biguttatus (Agrilus), 167, 174
bilineatus (Hydroporus), xx, 168
billbergi (Coryna), 176 :
bimaculata (Aphthona), 11
KA (Dibolia), 4, 5
bimaculatus (Cryptocephalus), 179
sf (Drasterius), 174
binghami (Ingrisma), 308, 310
bipunctata (Megalognatha), 29
A (Paleophylia), 23
bipunctatus (Cryptocephalus), 179
bipustulatus (Agabus), 172
biseutelatus (Ceuthorrhynchus), 1
bisignatus (Olibrus), 172
bispinosa (Diacantha), 17
exvill_)
|
|
bubalus (Bubas), 173
Bubas, 173
bugabensis (Clceotus), 519
Buphonella, 37
Buphonida, 37
Byrrhus, 166, 173
cerulea (Monolepta), 34
czesus (Pleurophorus), 173
Calamobius, 178
campestris (Cicindela), xv
Candezea, 35, 36, 37
_cantabricum (Apion), 178
capensis (Candezia), 35
,, (Iphidea), 35
Carabus, xlvii, 171
Cardiophorus, 167, 169, 174
cardui (Agapanthia), 178
Sp bixus) Sali /7/
carinatus (Deronectes), 168, 172
carinipennis (Crepidodera), 12
carinula (Sciaphilus), 167, 176
carpetanum (Bembidium), 171
Carpophagus, xiii
castanipes, (Melanotus), 174
eastellana (Asida), 166, 169, 175
celtibericus (Cryptocephalus), 179
Cephalocosmus, 307
Ceraspis, 520
Cerocoma, 168, 169, 170, 176
cervus (Lucanus), 173
Cetonia, 173
Ceuthorrhynchidius, 177
Ceuthorrhynechus, 177
Cheetocnema, 179
| Charopus, 174
_Chasmatopterus, 167, 173
| Chlenius, 166, 169, 171
| chlorizans (Pseudocanthon),
_chlorophana (Cheetocnema),
510
179
_chloroptera (Apophylia), 22
if |
|
|
|
bituberculata (Hyperacantha), 17, 18
blattariz (Cionus), 178
Bledius, 169, 172
bohemani (Megalognatha), 27
bonvouloiri (Aphodius), 167, 169, 173
borrei (Paleeophylia), 23
Brachyderes, 167, 176
braunsi (Candezea), 35, 36
brevicorne (Trimium), ix
brevipennis (Aleochara), 172
brevirostre (Apion), 178
Brontes, 172
brunneus (Agabus), 168, 172
brunnipes (Clceotus), 518
a (Silaria), 175
Cheeridium, 509
Chrysobothrys, 174
Chrysochloa, i, xiv, 245-261
Chrysomela, 179
Cicindela, xv, 171
cinctus (Agrilus), 174
Cionus, 178
circumfusus (Luperus), 179
Cleeotus, 517, 518, 519
clypeatus (Ccelometopus), 175
Clytanthus, 169, 170, 178
Clythra, 167, 178
Cneorane, 27
Cneorrhinus, ix, 167, 169, 170, 176,
He fe,
coarctata (Tachyusa), 170, 172
Coccinella, xxx, 179
Ccelodes, 516
Ciexice s)
Ccelometopus, 175
Ceelostoma, 172
ccerulescens (Cryptocephalus), 179
colombianum (Aigidium), 515
Colon, xxxvii
concolor (Gynandrophthalma), 178
confluens (Polydrusus), 176
confusa (Notothecta), xxxvii
constrictus (Zabrus), 166, 171
Coptocephala, 178
Coreebus, 167, 168, 174
coruscans (Geotrupes), 169, 173
Corymbites, 166, 167, 174
Coryna, 176
Coryphocera, 307
coxalgica (Mecynodera), xiv
crassicollis (Cleeotus), 518
crassus (Cryptocephalus), 179
crenatus (Niptus), xxxvi
- crenulatus (Peecilus), 171
Crepidodera, 12, 13, 14
eribratum (Agidium), 515
crinitus (Sitones), 176
Criocephalus, xxxviii
eristatus (Sciaphilus), 176
Cryptarcha, lx
Crypticus, 175
Cryptocephalus, 167, 168, 169, 179
Ctesias, 173
eumingi (Pheedimus), 303
cuniculus (Aphodius), 511
cuprea (Haltica), 2, :
cupreola (Ingrisma), 307
eyanescens (Apion), 178
eyanicollis (Haltica), 3
cyanocephala (Lebia), 171
eyanoptera (Psylliodes), 179
Cymindis, 166, 169, 171
Danacwa, 167, 174
Dapsa, 172
Dasytes, 167, 174
10-guttata (Epurea), Ix
decemlineata, 249, 250
dejeani (Dorcadion), ix, 166, 167, 169,
178
,, (Zonabris), 176
Dendarus, 175
dentata (Dinarda), xxxvii
denticornis (Helionica), 305
denticulatus (Ceuthorrhynchus), 177
dentipes (Colon), xxxvil
,, (Otiorrhynchus), 166, 176
depilis (Byrrhus), 166, 173
Deronectes, 168, 172
Diacantha, 17
Mibolia, 3; 4; 5; 179
Dichillus, 175
didymus (Agabus), 168, 172
didymus (Scopzeus), 172
diffusa (Epureea), 1x
| Dima, 170
Dinarda, xxxvii
| Diploccelus, xxxvii
| discoidea (Acmeodera), 174
discolor (Donacia), 178
_ dispar (Cneorrhinus), 176
tI}
(Xyleborus), lviii
var. meleagris (Cneorrhinus),
177
| distineta (Coccinella), xxx
| dives (Chlenius), 166, 169, 171
Donacia, 178
Dorcadion, ix, 166, 167, 169, 170, 178
Dorcus, 173
Doryphora, 249, 250
Drasterius, 174
dregei (Diacantha), 17
Drilus, xlviii, 39-51
dubia (Scraptia), 175
dunbrodensis (Longitarsus), 8
dunbrodyensis (Luperus), 20
durieui (Nanophyes), v, 87-91, 168, 178
duvivieri (Apophylia), 24
,, (Hyperacantha), 18
Dytiscus, 172
Ebeus, 167
Elater, xxxvii, 167, 174
elegans (Amauronia), 174
re (Megalognatha), 27, 31
elegantula (Apophylia), 27
3 (Atomaria), Ix
elegantulus var. gracilis (Ceuthorrhyn-
chus), 178
elongata (Buphonella), 37
elongatissimum (Apion), 178
elongatulus (Elater), xxxvii
elongatum (Bembidium), 171
elongatus (Aphanisticus), 174
emarginatus (Aphanisticus), xxxvi, 174
Emblethis, 169
Emenadia, 175
Epicometis, 173
Epilachna, 179
Epurea, 1x
equiseti (Cardiophorus), 174
Ergana, 38
erinaceus (Strophosomus), 167, 176
erosus (Cryptocephalus), 169, 179
estrellanus (Zabrus), 171
Euconnus, lix
Euryeycla, 32
euryrrhina (Heterarrhina), 307
EKutornus, 6
Exochomus, 179
Exosoma, 25
fagi (Diploccelus), xxxvii
(Sexy)
fasciatus (Corcebus), 168, 174 Gymnopleurus, 167, 173
Faula, 520 Gynandrophthalma, 167, 178
femorata (Ingrisma), 309 Gyrinus, 168, 172
fenestratus (Ilybius), 172 hemapterus (Corymbites), 166, 174
ferrugineus (Aphodius), 173 hematura (Candezea), 36
festiva (Diacantha), 17 Haliplus, 172
ficus (Hypoborus), 178 Halticay 1 25,3, 167,09
filum (Calamobius), 178 | Halyzia, ex
flagellatus (Gymnopleurus), 173 | Haplocnemus, 167, 174
flaveola (Candezea), 36 Harpalus, 169
flavescens (Drilus), xlviii, 39-51 Helhonica, 303
Ap (Sitones), 176 Heliopates, 166, 175
», var. cinnamomeus (Sitones), | Helochares, 172
6s © Helophorus, 172
flavicornis (Hemixantha), 32 Helops, 167, 175
5 (Rhamphus), 178 hemisphxricum (Nanophyes), 88
flavipes (Hydroporus), 168, 172 Hemixantha, 32
», (Oniticellus), 173 Henicopus, 167, 174
fontenayi (Leptura), 169, 178 Hespera, 5, 6 2
foveicollis (Cneorane), 27 Heterarrhina, 307
foveolatus (Olocrates), 175 Heterocerus, 170 ;
fracticornis (Bledius), 172 heydeni (Agabus), 163, 172
fragilis (Stilicus), xxx », (Henicopus), 174
frater (Ateenius), 512 hirta (Epicometis), 173
fulvipes (Ergana), 38 -,, (Lagria), 175
», (Staphylinus), xxx | hirticollis (Megalognatha), 28
funerula (Anthaxia) 173 | hirtulus (Chasmatopterus), 173
furcatus (Onthophagus), 173 hispanicum (Anthobium), 172
fuscipes (Aleochara), 172 Fa (Bembidium), 171
», (Pederus), 172 2% (Calostoma), 172
fuscitarsis (Diacantha), 17 hispidulus (Bledius), 172
fuscus (Meligethes), 172 es (Chasmatopterus), 173
,, (Philonthus), xxxvii, lx Hister, 173
geminata (Zonabris), 176 Homalium, lix
generosa (Diacantha), 17 Hoplia, 173
geographicus (Ceuthorrhynchus), 177 | hospita (Thamiarza), 1x
Geotrupes, 167, 169, 173 howitti (Theodosia), 304
ghiliani (Steropus), 169, 171 », (Westwoodia), 803, 304
gibbicollis (Corcebus), 174 hungarica (Anthaxia), 168, 173
glaciale (Bembidium), 166, 171 hungaricus (Onitis), 173
clobicollis (Cryptocephalus), 168, 179 | hybrida (Cicindela), 171
gloriosa (Diacantha), 17 Hydrena, 172
Glyptolus, 22 Hydrochus, 172
godarti (Athous), 174 Hydrocyphon, 174
Gonocephalum, 175 Hydroporus, xx, 168, 172
goudoti (Asida), 175 Hymenalia, 175
gougeleti (Ceuthorrhynchus), 178 Hymenoplia, 173
Ss (C Rote, 167, 174 Hypebeeus, 174
gracilis (Ateenius), 5 Hyperacantha, 17, 18, 19
graellsi aoe 174 Hyperaspis, 179
»5 (Cneorrhinus), 177 hyperici (Agrilus), 174
granularis (Hydroporus), xxi Hypoborus, 178
granulicollis (Megalognatha), 28 | hypocrita (Geotrupes), 175
granulosa (Apophylia), 25 hypomelena (Anthaxia), 173
a (Paleophylia), 24 ibericuin (Bembidium), 171
grenadensis (Saprosites), 514 ilieis (Rhynchenus), 178
grenieri (Lagria), 175 illesum (Cheeridium), 509
gressorius (Sitones), 176 imbecilla (Megalognatha), 28
imbricatus (Atenius), 513
immaculata (Pseudolognatha), 31
immune (Apion), 178
imperialis (Cryptarcha), 1x
ap (Cryptocephalus), 179
impressipennis (Podagrica), 11
inequalis (Hister), 173
incanus (Brachyderes), 176
», (Limnichus), 173
incertus (Omophlus), 175
incilis (Trechus), xxi
inconspicuus (Luperus), 22
indica (Podagrica), 12
inermis (Mycteristes), 306
infuscata (Candezea), 36
Ingrisma, 307, 308, 309
inornata (Monocida), 26
insularis (Faula), 520
Iphidea, 35
iridis (Cixis), 177
khasiana (Mycteristes), 307
kiesenwetteri (Corymbites), 174
koyi (Cryptocephalus), 167, 179
kraatzi (Apion), 178
», (Crypticus), 175
Labidostomis, 178
Laccobius, 172
Lachnea, 167, 178
leeve (Lasioderma), 174
lievicollis (Notothecta), 172
leevigatum (Apion), 178
levigatus (Geotrupes), 173
leviuscula (Clythra), 178
Lagria, 168, 175
laminatus (Philonthus), 172
Lampyris, 174
Larinus, 177
larvata (Emenadia), 175
larvatus (Ceuthorrhynchus), 177
Lasioderma, 174
lata (Lagria), 175
laterale (Bembidium), 171
Lathrobium, 170, 172
laticollis (Ateuchus), 173
aon (Elelops) iy i75
latus (Carabus), 171
», (Corymbites), 174
Lebia, 167, 171
lepidus (Hydroporus), 168, 172
leprieuri (Stenus), 172
Leptura, 169, 178
Leucocelis, 173
leucopsideus (Trichodes), 174
Limnebius, 172
Limnichus, 173
Lionychus, 171
Liopterus, 172
lividus (Luperus), 179
(
exxi )
Livolia, 15
Ibreqney, 1/7
longimama (Arsoa),
longitarsis (Stenus),
Longitarsus, 7, 8, 179
Lucanus, 173
Luperodes, 20, 35
Luperus, 20, 21, 22, 167, 179
luridus (Aphodius), 511
20.
172
| lusitanica (Labidostomis), 178
lusitanicum (Malacosoma), 179
Lypnea, 14
lythri (Nanophyes), 87, 178
Lytta, 167, 176
maculata (Dibolia), 4
a (Strangalia), 178
maculatum (Bembidium), 171
maculicollis (Hespera), 5, 6
a6 (Paleophylia), 23
madagascariensis (Rhagiosoma), xiii
mierens (Aleochara), XxxXvii
magnifica (Theodosia), 304, 505
majalis (Meloe), 167, 175
major (Hister), 173
miklini (Eucounus), lix
Malachius, 167, 174
Malacosoma, 25, 179
malvernensis (Haltica), 1
Malaxia, 22, 23
Malthinus, 167, 174
malvernensis (Luperus), 21
marginalis (Dytiseus), 172
marginata (Apophylia), 24
marginatus (Platynus), 171
| marginicollis (Onthophagus), 511
;marshalli (Crepidodera), 13
Mastigus, 172
Mecynodera, xiv
Megalognatha, 27, 28, 29, 30, 31
| Megalopus, xx
melaleucus (Neophedimus), xxi
melancholicus (Carabus), 171
| melanocephala (Cymindis), 171
Ar (Megaloenatha), 29, 30
melanostictus var. murinus (Ceuthor-
rhynchus), 177
Melanotus, 167, 174
Meligethes, 172
melipona (Megalopus), xx
Meloe, 167, 175
metallicus (Cloeotus), 519
/microphyllus (Mycteristes), 307
Micrositus, 175
nulitaris (Hyperacantha), 19
millefolii (Anthaxia), 173
minutissimus (Bidessus), 168, 172
minutum (Anthobium), 172
moewisi (Cephalocosmus), 307
(
Mombascia, 37
Monocida, 26
Monolepta, 33, 34
montanum (Bembidium), 171
mori (Cryptocephalus), 179
Mordella, 175
Mordellistena, 175
morio (Cetonia), 173
murina (Apophylia), 23
mutabilis (Adalia), 179
mutata (Serica), 168, 173
Mycteristes, 306
Nanophyes, v, 87-91, 168, 178
nanus (Stenus), 172
natalensis (Megalognatha), 29
Nebria, 166, 170, 171
neglectus (Zabrus), 171
nemoralis (Carabus), xlvii
Neophedimus, xxi
niger (Aphodius), 173
», (Luperus), 179
», (Nanophyes), 178
5, (Orphilus), 173
nigerrimus (Elater), 174
nigra (Strangalia), 178
nigripennis (Ccelodes), 516
nigrita (Pterostichus), 171
nigritarsis (Paleophylia), 23
nigritula (Longitarsus), 7
nigrocerulea (Candezea), 37
nigrofasciata (Megalognatha), 30
nigrofasciatus (Luperus), 167, 179
nigrosuturalis (Candezea), 35
nigrotibialis (Luperodes), 20
Niptus, xxxvi
nitidus (Agabus), 172
nivicola (Corymbites), 174
nobilitata (Apophylia), 23
Notothecta, xxxvii, 172
Notoxus, 175
oblonga (Cetonia), 173
Ochthenomus, 175
18-guttata (Halyzia), xxx
octomaculatum (Bembidium), 171
octopunctatus (Trichodes), 169, 174
(Kdemera, 176
Olibrus, 172
olivacea (Phytodecta), 179
Olocrates, 166, 167, 169, 175
Omophlus, 175
Oniticellus, 173
Onitis, 173
Onthophagus, 173, 510
Oochrotus, 175
ooptera (Amara), 166, 169, 171
Orchestes, 168
Orectochilus, 168, 172
Orina, i, xiv, 170, 245-261
CxxH }))
| Orphilus, 173
Otiorrhynchus, 166, 176
Pachybrachys, 168, 179
Pachytychius, 168, 177
Pederus, 169, 170, 172
Paleophylia, 23, 24
pallida (Hespera), 6
paludosus (Agabus), 168, 172
pandellei (Trechus), 166, 171
papalis (Mastigus), 172
Paracymus, 172
parallelopipedus (Doreus), 173
parallelus (Saprosites), 514
Parnus, 170, 172
parumpunctatus (Anchomenus), lviil
parvula (Lagria), 175
parvulum (Aégidium), 515
parvulus (Ccelodes), 516
(Psammobius), 514
, . (Tachys), 171
paulinoi (Bembidium), 171
pazi (Nebria), 171 ;
perakensis (Theodosia), 304, 305
Perileptus, 169, 171
| Pheedimus, 303
Philhydrus, 172
Philonthus, xxxvii, lx, 170, 172
Philorhinum, 172
_Phylobius, 167, 176
Phyllotreta, 179
Phymatodes, 178
| Phytodecta, 179
Phyteecia, 178
picea (Diacantha), 17
piceipes (Hemixantha), 33
,, (Omophlus), 175
picturatus (Eutornus), 6
pilipes (Megalopus), xx
pilularius (Gymnopleurus), 173
Pimelia, 167, 175
pineticola (Rhizotrogus), 173
Plagionotus, 178
planatus (Brontes), 172
Platycerus, 167, 178
platycerus (Notoxus), 175
Platynaspis, 179
Platynus, 171
Platyxantha, 33
Pleurophorus, 173
| Podagrica, 11
Pecilus, 171
politus (Philonthus), 172
polonicus (Criocephalus), xxxXviil
Polydrusus, 167, 176
polyglyptus (Atéenius), 515
posticus (Cleeotus), 518
preeusta (Tetrops), 178
(Zonitis), 176
99
(Sexxin =)
preustus, var. aurileculus (Elater), 167,
1
Psammobius, 514
Pseudapophylia, 23
Pseudocanthon, 510
Pseudolognatha, 31
Pseudophlceus, 169
Psylliodes, 179
pteromelas (Pachybrachys), 179
Pterostichus, 166, 171
Ptinus, v
pubescens (Lachneea), 178
punctatum (Lathrobium), 172
puncticosta (Aspidomorpha), xxxvill
punctipennis (Monolepta), 33
punctulatum (Bembidium), 171
punctus (Philonthus), 172
pusillus (Aphanisticus), xxxvil
putoni (Apion), 178
pygialis (Rhizotrogus), 173
pygmeus (Cryptocephalus), 179
3 (Uroxys) 510
pyriformis (Cneorrhinus), ix, 176, 177
pyritosa (Haltica), 2, 3
14-pustulata (Coccinella), 179
quadriguttatum (Bembidium), 171
quadriguttatus (Anthicus), 175
quadripunctata (Zonabris), 176
quadripunctatus (Cryptocephalus),
quadripustalatum (Bembidium), 17
Quedius, Ix
quercus (Rhyncheenus), 178
rasuta (Ingrisma), 307
reflexa (Trachys), 174
regentsteinensis (Sitones), lix, 176
relucens (Acanthocerus), Ixiil,
517
reppensis (Hyperaspis), 179
Rhagiosoma, xiil
Rhagonycha, 167, 174
Rhamphus, 178
rhenana (Atomaria), v
Rhizotrogus, 173
Rhynchenns, 178
rivularis (Trechus), xxi
rodriguesi (Anthicus), 175
Rosalia, xlvii
roscidus (Agrilus), 174
rothschildi (Theodosia), 304, 305
rubida (Lagria), 168, 175
rubricus (Nanophyes), 178
ruficollis (Cymindis), 171
(Liopterus), 172
9p (Megalognatha), 31
(
(
516,
Omophlus), 175
oF Peederus), 170, 172
ruficornis (Clytanthus), 178
rufipennis (Aleochara), 170, 172
179
1
rufipes (Cardiophorus), 169, 174
», (Hymenalia), 175
rufiventris (Megalegnatha), 28
rufopiceus (Cloeotus), 517
rugatipennis (Anomala), 169, 173
rugosicollis (Henicopus), 174
rugulosa (Orina), 170, 245
rutilipennis (Silesis), 174
sacer (Ateuchus), 173
salisburiensis (Pseudolognatha), 31
salzmanni (Abacetus), 171
sanguineum (Apion), viii
Saprosites, 514
Sardoides, 26, 27
saxeticola (Olocrates), 175
saxicola (Olocrates), 175
scabricollis (Pachytychius), 168, 177
scalptifrons (Atzenius), 513
Seaurus, 167, 175
schiefferi (Sisyphus), 173
schdnherri (Apion), 178
schreberi (Cerocoma), 168, 169, 176
», (Onthophagus), 173
Sciaphilus, 167, 176
Scopeus, xxxvil, 172
scopolina (Coptocephala), 178
Seraptia, 175
scrofa (Aphodius), 173
scutellata (Candezea), 36
Seymnus, 179
sedi (Apion), 87, 88
semirugosa (Apophylia), 24
senegalensis (Aphthona), 10
Serica, 168, 173
sericea (Asida), 175
pee (CUbalyera)) sel
serra (Ctesias), 173
setifrons (Polydrusus), 176
sexmaculata (Titubcea), 178
sexmaculatus (Cryptocephalus), 179
sexpunctata (Lachneea), 178
6-pustulata (Diacantha), 17
sexstriatus (Tachys), 171
Sibinia, 178
siculus (Nanophyes), 88
signatus (Cardiophorus), 167, 174
Silaria, 175
Silesis, 174
Supha, 172
silphoides (Zabrus), 171
Silvanus, 172
similis (Aphthona), 10
simplex (idemera), 176
sinuatus (Hister), 173
Sisyphus, 167, 173
Sitones, lix, 167, 176
smaragdina (Apophylia), 22, 23
ns (Orina), i, 245-261
(
smaragdinipennis (Pseudapophylia),
Smicronyx, 168
solieri (Agrilus), 174
sorbi (Apion), lix, 178
sordidum (Philorhinum), 172
sordidus (Atzenius), 513
sparsus (Rhynchenus), 178
sparsutus (Pachytychius), 168, 177
spartii (Lixus), 177
spinifer (Platycerus), 167, 173
spinipes (Cneorrhinus), 177
spinosa (Diacantha), 17
squamulatus (Ceuthorrhynchus), 177
Staphylinus, xxx
steinheili (Atenius), 513
Stenoplatys, 17
Stenopterus, 178
Stenus, 172
Steropus, 169, 171
stictica (Leucocelis), 173
Stilicus, xxx
stolida (Basipta), xxxviil
Strangalia, 178
striatum (Apion), 178
5 (Asemum), XXxxviil
striatus (Scaurus), 175
strigata (Cryptarcha), lx.
strigicauda (Atenius), 511, 512
Strophosomus, 167, 176
Stylosomus, 179
suavis (Philonthus), 172
subcostatus (Dichillus), 175
suberosus (Trox), 516
sulcicollis (Livolia), 15
ss (Scopzeus), XXXvil.
sulcifrons (Barypithes), 167, 176
sulphuripennis (Luperodes), 20
Sunius, 172
suturalis (Megalognatha), 29
Tachys, 171
Tachyusa, 170, 172
teniata (Acmeodera), 169, 173
tamarisci (Nanophyes), 88
tanaceti (Adimonia), 179
telephii (Nanophyes), 87, 88
Telephorus, 174
telifer (Theodosia), 304, 305, 306
tenebrosus (Atenius), 509, 512, 513
ae (Melanotus), 174
Tentyria, 166, 167, 174
terminalis (Atenius), 512
tessellatus (Hydroporus), 172
testaceum (Homalium), lix
testaceus (Phymatodes), 178
testudinaria (Diacantha), 17
Tetrops, 178
thalassina (Donacia), 178
Thalyera, 1x.
Cxxiva5)
23 | Thamiarea, lx
Theodosia, xxxv, 303-310
thoracica (Dibolia), 3
», (Monocida), 26
Thylacites, 170, 177
| Timarcha, 166, 179
| timida (Baris), 178
| 4, (Dibolia), 179
| Titubcea, 178
| tomentosus (Tropiphorus), lviii
_tonkinensis (Coryphocera), 308
torquatum (Anthobium), 172
| Trachys, 174
| transvalensis (Sardcides), 26
Trechus, xxi, 166, 171
Trichodes, 169, 174
tricincta (Diacantha), 17
tricolor (Apophylia), 22, 25
», (Paleophylia), 23
trifasciata (Silaria), 175
trifasciatus (Attagenus), 172
55 (Clytanthus), 169, 170, 178
ee (Notoxus), 175
trimaculata (Lebia), 167, 171
Trimium, ix
_Triodonta, 173
'tristis (Anthicus), 175
», (Orina), i., 245-261
ys Var. smaragdina (Chrysochloa),
i, xiv, 245-261
»» var. smaragdina (Orina), i, xiv,
245-261
| Tropiphorus, lviii
Trox, 516
| tuberculifer (Phyllobius), 176
tugelaensis (Luperus), 21
turbatus (Thylacites), 177
ulyssiponensis (Micrositus), 175
11-punctata (Diacantha), 17
unicolor (Oochrotus), 175
/ unidentatus (Silvanus), 172
| unifasciata (Diacantha), 17
_unifasciatus (Ochthenomus), 175
_uniformis (Crepidodera), 14
unipunctata (Diacautha), 17
urens (Ceuthorrhynchidius),
urinator (Gyrinus), 168, 172
Uroxys, 510
variabilis (Coccinella), 179
| (Phytodecta), 179
varians (Aphodius), 173
varicornis (Platyxantha), 33
_variolosa (Cymindis), 171
varius (Hydroporus), 172
var, bimaculatus (Philonthus),
—_
~sI
~I
| 5,
172
velutinus (Chlenius), 171
| ventralis (Quedius), 1x
verticalis (Luperus), 21
verticicornis (Onthophagus), 173
vescicatoria (Lytta), 167, 176
vicarius (Acanthocerus), 517
vicina (Lachneza), 178
villosa (Platynaspis), 179
villosulus (Chasmatopterus), 173
villosus (Orectochilus), 172
vincentie (Agidium), 515
SG (Ateenius), 513
5 (Uroxys), 510
violaceus (Cryptocephalus), 179
virescens (Phytcecia), 178
viridicollis (Helionica), 305
.; (Sardoides), 37
viridicupreus (Platynus), 171
viridiniteus (Paleophylia), 23
viridipennis (Clcotus), 519
viridissimus (Pachybrachys), 179
waterstradti (Helionica), 304
Weiseana, 16
westwoodi (Helionica), 303, 304, 305
eA (Theodosia), 304, 305
Westwoodia, 303
whiteheadi (Theodosia), 307
Xyleborus, lviii
Xylopertha, 174
Zabrus, 166, 169, 171
Zonabris, 167, 170, 176
Zonitis, 176
HYMENOPTERA.
abdominale (Rhynchium), 106
aberrans (Cremastogaster), 98
absoluta (Prosopis), 107
Acantholepis, 99
acasta (Polyrhachis), 99
acutipennis (Xylocopa), 109
adusta (Nomada), 108, 116
advena (Nomada), 108
edilis (Pompilus), 102
edipus (Mutilla), 100
fnictus, 96
zstuans (Xylocopa), 109, 116
affinis (Pheidologeton), 98, 110
s, (Vespa), 107
agelia (Mutilla), 101
agilis (Astata), 103
albescens (Halictus), 132
albifrons (Megachile), 108
albipes (Technomyrmex), 98
albofimbriata (Nomia), 108
alecedo (Halictus), 214
allaborans (Sima), 98
Allodape, 108
alluaudi (Anthophora), 551
»> (Podalirius), 551
cxxv )
| Alyson, 105
| amatorius (Gorytes), 105, 113
Amblyopone, 97
| Amblyteles, 230
_amethystina (Xylocopa), 109
| Ammophila, 104, 210
| Ampulex, xxx, 105, 110, 112, 234-238
analis (Liacos), 101
» (Mutilla), 100
», (Pompilus), 102
| anceps (Iridomyrmex), 98
Ancyra, 228, 229
| Andrena, 215
angustifrons (Halictus), 215
annulata (Elis), 101
annulipes (Alyson), 105
Anochetus, 96
Anthidium, 108
Anthophora, 95, 109, 116, 551
anthracina (Notogonia), 104, 117
antoni (Odynerus), 107
apicalis (Megachile), 216
Apis, 109, 116
| Aporus, 102
_appendiculata (Cenolarra), 104
| appendiculatus (Pison), 104
_architectus (Eumenes), 106
| arctifrons (Halictus), 215
arcuata (Eumenes), 106
_ardens (Crabro), 106
argentatum (Rhynchium), 106
argentatus (Crabro), 106
| argentea (Polyrhachis), 99
| argenteobalteata (Nomia), 108
| argenteofacialis (Lyroda), 103
_argentifrons (Ceelioxys), 108
_argentipes (Mutilla), 100
argyreus (Tachysphex), 103
ariadne (Pompilus), 102
ariel (Mutilla), 101
,, (Pseudagonia), 102
| armata (Solenopsis), 110
| arrogans (Camponotus), 99
| artifex (Iearia), 107
_assamensis (Ampulex), 236, 238
Astata, 103
atomus (Monomorium), 97
Atopomyrmex, 98
atra (Prosopis), 212
atripes (Ammophila), 104
atroalba (Podalirius), 216, 217
atropus (Salius), 103
auratus (Liris), 104
/ aureipennis (Scolia), 101
aureobalteata (Nomia), 108
aureo-rubra (Mutilla), 100
auriceps (Tachysphex), 103
auripennis (Xylocopa), 109
(
aurulentus (Sphex), 105, 112
balearicus (Podalirius), 217
basalis (Ammophila), 104
(Ccelioxys), 108
(Notogonia), 104
(Vespa), 107
9)
3
basimacula (Rhynchium), 107, 115, 114
bellicosa (Prosopis), 107
bellus (Crabro), 106
Bembex, 105, 113
bengalense (Rhynchium), 107
bengalensis (Aporus), 102
- (Halictus), 108, 131
(Melipona), 109
(Notogonia), 104, 123
(Oligomyrmex), 98
(Plesia), 101
5 (Prenolepis), 99
Benyllus, 232
bicincta (Anthophora), 109, 116
bicolor (Megachile), 108, 116
(Meranoplus), 97, 110
(Methoea), 101
(Polyrhachis), 99
5, (Trypoxylon), 106
bidens (Mutilla), 100
bimaculata (Andrena), 215
binghami (Gastrosericus), 104, 113
bipartita (Acantholepis), 99
5p (Andrena), 215
bipartitus (Salius), 103
bipustulatus (Odynerus), 107
bituberculata (Tachysphex), 127
blanda (Pseudagenia), 102
blandinus (Stizus), 105
Bombus, xxxvii, 207, 217, 218, 551
Bothriomyrmex, 98
Bothroponera, 96
Brachyponera, 97, 109
brevicornis (Ainictus), 96
3 (Ampulex), 238
brevipennis (Elis), 101
a (Tachytes), 103
- (Tiphia), 101
brunnea (Myrmicaria), 97, 110
brunneum (Rhynchium), 106
bryorum (Xylocopa), 109
Buathra, 233, 234
buddha (Bembex), 105, 113
(Crabro), 106
», (Eumenes), 106
_;, (Trypoxylon), 106
Ceenolarra, 104
cerulea (Pseudagenia), 102
>, (Trirhogma), 105
calopteryx (Stizus), 105
Camponotus, 95, 99, 111, 208
canaliculatum (Trypoxylon), 106
>)
oe)
9?
+]
be]
2)
exxvi )
canariensis (Podalirius), 216, 551
canescens (Oxybelus), 106
canifrons (Pompilus), 102
capensis (Acantholepis), 99
capitata (Scola), 101
cara (Mutilla), 100
carbonaria (Parevaspis), 108, 116
earbonarium (Monomorium), 209
Cardiocondyla, 98
carinifrons (Ampulex), 237, 238
Ap (Halictus), 132
Caspipina, 219 ,
Cataulacus, 97
cellularis (Pompilus), 102
| Cemonus, 105
| Ceratina, 108
Cerceris, 105, 106, 113
ceylonica (Nomada), 108
_ceylonicus (Atopomyrmex), 98
i (Eumenes), 106
tes (Halictus), 131
chinensis (Lobopelta), 97
Chrysis, 96
cincta (Anthophora), 109, 116
5, (Vespa), 107, 114
cinerascens (Sphex), 105
ciris (Halictus), 108, 130
clavipes (Dolichurus), 105
cleonyma (Mutilla), 100
clypeata (Nomia), 108
clypeatum (Rhynchium), 107
| Ccelioxys, 108, 109
| cognata (Ampulex), 238
| cognatum (Trypoxylon), 106
collaris (Xylocopa), 109
Colietes, xviii
collina (Trigona), iv, 133, 134, 136
Colobopsis, 99
ecompactilis (Mutilla), 100
compressa (Ampulex), 105, 112, 237,
238
(Sima), 98
99
compressus (Camponotus), 95, 99, 111
cona (Mutilla), 100
confinis (Ponera), 97
», var. aitkeni (Ponera), 97
confusa (Ceelioxys), 108
conica (Eumenes), 106, 113
constancez (Rhinopsis), 113
constricta (Ponera), xxxvil
contemta (Crematogaster), 98
contracta (Ponera), 209
coromandelicum (Sceliphron), 104
cotesi (Aporus), 102
(Chrysis), 96
» (Mutilla), 101
Crabro, 106
crassicornis (Pison), 104
2?
( exxvii )
crassicornis (Sphecodes), 107 ferruginea (Icaria), 107
Cremastogaster, 95, 98 festinata (Pseudagenia), 102
Crocisa, 95, 109, 116 fianna (Mutilla), 100
eruentatus (Odynerus), 212 flavinerva (Leptolarra), 104
cuneata (Ceelioxys), 108 flavipes (Oxybelus), 106
curvipes (Nomia), 108, 115 flavomaculata (Ancyra), 229
cylindricus (Halictus), 215 54 (Stelis), 108
Darachosia, 221, 222 flavomarginatum (Rhynchium), 107
deceptrix (Pseudagenia), 102 flavopicta (Cerceris), 105
denticulatum (Tetramoriuin), 97 Pe (Eumenes), 106
depredator (Philanthus), 105 flavus (Salius), 103
destructor (Monomorium), 97 florea (Apis), 109, 116
Diacamma, 95, 96, 97, 109 floricola (Monomorium), 97
Didineis, 105 fodiens (Myrmicaria), 97
diffinis (Odynerus), 107 Formica, xxx, Xxxvii
dilecta (Mutilla), 100 forticeps (Losgna), 230
dimidiata (Ammophila), 104 fortinata (Mutilla), 100
5 (Plesia), 101 frauenfeldi (Acantholepis), 99
_ diminuta (Lobopelta), 97, 109 frederici (Nomia), 108
discipiens (Celioxys), 108 fulgidipennis (Salius), 103
discreta (Mutilla), 100 fuliginose (Formica), xxxvii
disjuncta (Megachile), 108, 116 fulvipennis (Salius), 103
disparilis (Chrysis), 96 fulvipes (Darachosia), 222
dissimilis (Xylocopa), 109 ,, (Legnatia), 226
diversus {Pheidologeton), 98 fulviventris (Osmia), vi, viii
dives (Mutilla), 101 fulvopicta (Liacos), 101
Dolichoderus, 99 fulvopilosus (Oxybelus), 106
Dolichurus, 105 funebrana (Mutilla), 100
dorsata (Apis), 116 funeraria (Mutilla), 100
Dorylus, 96, 109 furiosa (Chrysis), 96
dryta (Mutilla), 100 fuscipennis (Cemonus), 105
durga (Mutilla), 101 op (Ccelioxys), 109
eatoni (Miscophus), 209 - (Notogonia), 104
egregia (Mutilla), 100 fuscistigma (Notogonia), 104, 122
electus (Salius), 103 gallicus (Polistes), 211
Elis, 101, 112 Gastrosericus, 104, 113
elliotii (Nomia), 108 geminata (Solenopsis), 95, 98, 110
emancipata (Mutilla), 101 geniculatum (Trypoxylon), 106
emarginata (Crocisa), 109, 116 germanica (Vespa), 211
Enchisiades, 220 gnoma (Mutilla), 101
erraticum (Tapinoma), 208 Gorytes, 105, 113
erxia (Mutilla), 100 gracilipes (Dolichoderus), 99
erythrocephala (Ammophila), 104 guttulata (Nomada), lix
erythrogaster (Larra), 104 hematodes (Odynerus), 212
erythropoda (Notogonia), 104 hemorrhoidale (Rhynchium), 106
93 (Nysson), 105, 113 Halictus, 108, 130-132, 208, 213, 214,
escuriens (Eumenes), 106 215
Eucera, i hebreus (Polistes), 107, 114
Eumenes, 106, 113 hecate (Pompilus), 102
Eutanyacra, 227 Hedychrum, 96
Evirchoma, 222, 223 hero (Pompilus), 102
excellus (Salius), 103 Heterogyna, xxxi
excisus (Iridomyrmex), 98 hieroglyphica (Ceratina), 108
fallax (Anthophora), 109 himalayensis (Ampulex), 238
feai (Prosopis), 107 hindostanus (Odynerus), 107
femorata (Megachile), 108 hirsuta (Ammophila), 210
fenestrata (Xylocopa), 109 » (Elis), 101
ferruginea (Caspipina), 219 », (Psammophila), 210
(
histrio (Crocisa), 109, 116
Holcomyrmex, 97, 110
hortorum (Bombus), 217
humbertiana (Cerceris), 105
humeralis (Scolia), 101
Icaria, 107
Ichneumon, 231
idyia (Mutilla), 100
illa (Mutilla), 100
implactibilis (Pompilus), 102
incognitus (Pompilus), 102
indica (Apis), 109, 116
(Bembex), 105
(Notogonia), 104, 120
(Pheidole), 98
(Prenolepis), 99
5, . (scolia), 1005 112
instabilis (Cerceris), 105
intermedia (Notogonia), 104, 118
interrupta (Mutilla), 100
interstitialis (Ampulex), 238
Be (Halictus), 108, 130
iridipennis (Larra), 103
(Melipona), 109
_ (Salius), 103
Iridomyrmex, 98
Isobrachium, XXXvVil
itinerans (Halictus), 108, 150
jaculatrix (Notogonia), 104, 126
javana (Pheidole), 98
jerdoni (Brachyponera), 97, 109
junctus (Camponotus), 99
khasiana (Ampulex), 234, 238
kitteli (Lobopelta), 97
labiena (Mutilla), 100
laboriosa (Notogonia), 126
laboriosus (Pheidologeton), 98, 110
lata (Mutilla), 1012
letus (Halictus), 214
leevissima (Polyrhachis), 99, 111
lamellata (Nomia), 108
lanata (Megachile), 108, 116
Larra, 103, 104
lascivus (Pompilus), 102
Lasius, 208
lateralis (Stizus), 105
latifrons (Ampulex), 258
latinoda (Monomorium), 97
,, (Pheidole), 98
latipes (Nomina), 108
5, (Xylocopa), 109
latreillei (Osmia), 216
latro (Oxybelus), 210
Legnatia, 222, 225, 226
lena (Mutilla), 100
Leptolarra, 104
Leptothorax, 98, 209
lethargia (Mutilla), 101
29
9)
be)
29
9
CXxvili_)
Liacos, 101
liodomus (Halictus), 131
Lioponera, 97
Liris, 104
lobatus (Sphex), 105, 112
Lobopelta, 97, 109
longicollis (Ampulex), 234, 238
longicornis (Eucera), 1
a (Larra), 103
~ (Prenolepis), 99
longipes (Plagiolepis), 99
longitarsis (Leptolarra), 104
longitarsus (Lioponera), 97
Lophomyrmex, 98
Losena, 229, 230
ludovica (Mutilla), 100
lugubre (Hedychrum), 96
lunata (Bembex), 105, 113
luteipennis (Sphex), 105, 112
Lyroda, 103
Macromeris, 102
maculicornis (Mutilla), 100
maculipes (Pompilus), 102
maculitarsis (Tachytes), 103
maderz (Ammophila), 210
,, (Podalirius), 216
madraspatanum (Sceliphron), 104, 112
madraspatanus (Salius), 103
mandibularis (Pramha), 231
marcia (Mutilla), 100
marginata (Allodape), 108
marginella (Elis), 101, 112
martialis (Mutilla), 100
megacephala (Pheidole), 209
Megachile, 95, 108, 116, 216
melanocephalum (Tapinoma), 98
Melipona, 109, 135
melleus (Stizus), 105
mendicalis (Chrysis), 96
Meranoplus, 97, 110
Methoca, 101
minchini (Cremastogaster), 98
5, (Lobopelta), 97
minutula (Andrena), 216
mirandus (Salius), 105
Miscophus, 104, 209
mithila (Mutilla), 101
mitis (Camponotus), 99
», var. fuscithorax (Camponotus),
99
modesta (Tachytes), 103
monetaria (Tachytes), 103
Monomorium, 97, 209
montana (Ampulex), 238
morio (Halictus), 214
morna (Pseudagenia), 102
Mutilla, 95, 100, 101, 111
mutua (Pseudagenia), 105
(Gmrexexix yh)
Myrmicaria, 97, 110
Myzine, 95, 101
nanus (Crabro), 106
niger (Lasius), 208
nigra (Sima), 98, 110
nigricans (Ampulex), 238
5 CAlstata) 03
»» (Megachile), 108
nigripes (Ammophila), 104
nigritarsis (Polistes), 107
nigriventris (Larra), 104
nitidulum (Rhynchium), 106
nitidus (Crabro), 106
nodicornis (Steganomus), 108, 115
Nomada, lix, 108, 116
Nomia, 108, 115
Notogonia, 104, 117-126
nuda (Cardiocondyla), 98
Nysson, 105, 113
obesum (Tetramorium), 97
ocellata (Mutilla), 101
ocellifera (Lobopelta), 97
», (Pheidologeton), 98
oculata (Chrysis), 96
odontophorus (Crabro), 106
Odynerus, 107, 129, 212
(cophylla, 95, 99, 111
Oligomyrmex, 98
orientale (Monomorium), 97
orientalis (Bembex), 105
5 (Cereeris), 105
a (Chrysis), 96
ee (Crabro), 106
55 (Didineis), 105
+ (Dorylus), 96, 109
af (Methoca), 101
5 (Pison), 104
= (Planiceps), 102
,» (Pompilus), 102
5, (Vespa), 107, 115
ornatipes (Tachytes), 103
Osmia, vi, xx, 216
ovalis (Odynerus), 107
oxybeloides (Nomia), 108
Oxybelus, 106, 210
pachycerus (Mnictus), 96
pallidicoxis (Eutanyacra), 227
pallidimaculata (Evirchoma), 223
pamphia (Mutilla), 100
paradoxa (Heterogyna), xxxi
Parapison, 104
Parevaspis, 108, 116
paria (Camponotus), 99
Parnopes, 96
parthenia (Mutilla), 101
parva (Notogonia), 104, 119
,, (Pheidole), 98
parvula (Stelis), 108
PROC, ENT. SOC. LOND., Vv. 1903.
Passaloecus, 105
pedalis (Pompilus), 102
pedunculata (Pseudagenia), 102
pentadonta (Cerceris), 106
peregrina (Mutilla), 101
peregrinus (Salius), 103
perfecta (Chrysis), 96
perturbans (Pompilus), 102
perversa (Mutilla), 100
petiolata (Eumenes), 106
Pe (Plesia), 101
pheenna (Mutilla), 100
pharaonis (Monomorium), 97
Pheidole, 98, 110, 209
Pheidologeton, 98, 110
Philanthus, 105
Piagetia, 103
picipes (Notogonia), 104, 125
pictipes (Prosopis), 212
pictiventris (Cerceris), 105
pictus (Gorytes), 105, 113
pileatum (Trypoxylon), 106
piliventris (Notogonia), 104, 118
pilosa (Ampulex), 238
,, (Notogonia), 104, 125
pinguis (Bembex), 105
Pison, 104
Plagiolepis, 99, 208
Planiceps, 102
Platythyrea, 96
Plesia, 101
Podalirius, 216, 551
poesia (Mutilla), 100
Polistes, 107, 114, 211
Polyrhachis, 99, 111
Pompilus, 102
Ponera, xxxvli, 97, 209
Pramha, 231
Prenolepis, 95, 99
principalis (Chrysis), 96
propinquus (Sphecodes), 107
Prosopis, 107, 208, 212
pruinosus (Sphex), 105
Psammophila, 210
Pseudagenia, 102, 103
Pseudamblyteles, 227
pulcherina (Mutilla), 100
pulchra (Cerceris), 105
pulchriceps (Ampulex), 238
punctata (Ammophila), 104
Ap (Eumenes), 106
puncticeps (Tachysphex), 103, 127
punctiventris (Anochetus), 96
. (Lobopelta), 97
punctum (Odynerus), 107
purpureolineata (Nomia), 108
pygmea (Notogonia), 104, 124
», (Plagiolepis), 208
K
pygmeum (Trypoxylon), 106
4-carinata (Mutilla), 100
quadrifasciatus (Podalirius), 216
quadrimaculata (Icaria), 107
quadripustulata (Scolia), 101
quadrispinosa (Kumenes), 106
4-spinosus (Lophomyrmex), 98
quadristrigatus (Halictus), 213
rabula (Cremastogaster), 98
rasorum (Anthidium), 108
redacta (Mutilla), 101
redtenbacheri (Scolia), 101
reflexus (Pompilus), 102
reculatum (Rhynchium), 106
reticulata (Leptolarra), 104
reticulatus (Passaloecus), 105
reversus (Stizus), 105
Rhinopsis, 105, 110, 113, 238
rhombinoda (Pheidole), 98, 110
Rhynchium, 106, 107, 113
robustus (Oxybelus), 106
rogenhofferi (Cremastogaster), 98
rothneyi (Amblyopone), 97
oe (Ampulex), 238
Pr) (Cerceris), 105
(Colobopsis), 99
* (Cremastogaster), 98
oe (Gastrosericus), 104, 113
. (Miscophus), 104
_ (Mutilla), 101
. (Oligomyrmex), 98
Be (Parapison), 104
3 (Plagiolepis), 99
ia (Polistes), 107
os (Pompilus), 102
is (Salius), 103
He (Stigmatomma), 97
(Tachytes), 103
rubiginosa (Scolia), 101, 112
rufa (Formica), xxx
rufescens (Stizus), 105
ruficornis (Ampulex), 110
. (Melipona), 135
vs (Piagetia), 108
i (Rhinopsis), 105, 113
= (Trigona), iv, 135
ruficoxis (Ampulex), 238
rufipes (Enchisiades), 220
», (Larra), 104
rufitarsis (Spanolarra), 104
rufiventris (Buathra), 234
rufo-facies (Sycaonia), 224
rufoglaucus (Camponotus), 208
rufolineatus (Polistes), 107
rufonigra (Sima), 95, 98, 110, 113
rufus (Benyllus), 232
rugosus (Nysson), 105, 113
,» (Pison), 104
bed
le)
al
va
a
~-
sabellica (Mutilla), 100
saggitarius (Polistes), 107
Salius, 103
salomonis (Monomorium), 97
nO r. subopacum (Monomorium),
209
sanguinea (Formica), xxxvil
scabiose (Halictus), 213
scabriceps (Haleomyrmex), 97, 110
Sceliphron, 104, 105, 112, 210
Scolia, 101, 112
seulptum (Diacamma), 96, 97
scutellata (Nomia), 108
selma (Mutilla), 100
serena (Mutilla), 100
sericeus (Camponotus), 99
»> opaceventris (Camponotus), 99
sexmaculata (Ccelioxys), 108
a (Mutilla), 100, 111
sibilans (Odynerus), 107, 129
sicheli (Odynerus), 107
signata (Prosopis), 208, 212
Sima, 95, 98, 110
simillima (Larra), 103
simillimum (Tetramorium), 97, 209
simillimus (Pompilus), 102
simonyi (Tachysphex), 209
simplex (Polyrhachis), 99, 111
sinensis (Tachytes), 103
smaragdina (cophylla), 99, 111
smithi (Tetramorium), 97
smithii (Melipona), 135
| smythiesi (Pheidole), 98
AA var. bengalensis (Pheidole),
98
Solenopsis, 95, 98, 110
soroensis (Bombus), 207, 551
Spanolarra, 104
spathifera (Pheidole), 98
speculare (Monomorium), 97
Sphecodes, 107
Sphex, 105, 112
spinigera (Polyrhachis), 99, 111
splendidum (Stilbum), 96
x var. amethystinum (Stil-
bum), 96
squamosus (Oxybelus), 106
Steganomus, 108, 115
Stelis, 108
stigma (Polistes), 107
Stigmatomma, 97
Stilbum, 96
Stizus, 105
strenua (Prosopis), 107
striaticollis (Notogonia), 104, 121
striatidens (Triglyphotrix), 97
striativentris (Pheidole), 98
striolata (Tachysphex), 103, 126
(Ge 1c xexext a)
subaurata (Halictus), 215
subearinata (Myrmicaria), 97, 110
submicans (Osmia), 216
subnuda (Cremastogaster), 98
subtessellata (Notogonia), 104, 125
suspiciosus (Pison), 104
Sycaonia, 222, 224
Tachysphex, 103, 126-128, 209
Tachytes, 103
Tapinoma, 98, 208
taprobanie (Cataulacus), 97
rf (Dolichoderus), 95
tarsata (Tachytes), 103
taylori (Camponotus), 99
A (Leptothorax), 98
Technomyrmex, 98
tegularis (Oxybelus), 210
tenuscapa (Xylocopa), 109
terrestris (Bombus), 207, 208, 218, 551
tesserinoda (Bothroponera), 96
Tetramorium, 97, 209
thoracica (Elis), 101
” (Nomia), 108
thrinax (Polyrhachis), 99, 111
tibialis (Polyrhachis), 99
tincta (Pseudogenia), 102
Tiphia, 95, 101
tisiphone (Larra), 103
trepanda (Bembex), 105, 113
trichiosoma (Ampulex), 238
Triglyphotrix, 97
Trigona, iv, 183-136
trigona (Ampulex), 238
3-maculata (Mutilla), 101
Trirhogma, 105
tristis (Cerceris), 106
Trypoxylon, 106
tubericeps (Polyrhachis), 99
tubifex (Sceliphron), 210
tydei (Ammophila), 210
», (Psammophila), 210
umbripennis (Megachile), 108
umbrosus (Sphex), 105, 112
unifasciata (Leptothorax), 209
unifasciatus (Pompilus), 102
vagabundus (Pompilus), 102
vagans (Diacamma), 96, 109
valida (Mutilla), 101
varihirta (Tachysphex), 103, 128
varipilosa (Notogonia), 104, 122
vastator (Monomorium), 97
veda (Pseudagenia), 102
versicolor (Diacamma), 97
AS (Megachile), 216
verticalis (Xylocopa), 109
Vespa, 107, 114, 115, 211
vicina (Tachytes), 1038
vicinus, var. rothneyi (Sphex), 105
}
victoriz (Platythyrea), 96
viligans (Cerceris), 106
villosulus (Halictus), 214
violacea (Anthophora), 109
+9 (Macromeris), 102
violaceum (Sceliphron), 105, 112
viridis (Halictus), 214
,, (Parnopes), 96
viridissima (Ceratina), 108
vischnu (Cerceris), 105
AA (Pompilus), 102
vishnu (Halictus), 130
vivax (Pompilus), 102
vulgaris (Vespa), 211
watsoni (Pheidole), 98
westwoodii (Nomia), 108
wroughtoni (Bothriomyrmex), 98
xanthomelana (Osmia), xx
xanthopterus (Sphex), 105
Xylocopa, 95, 109, 116
yerburyi (Prenolepis), 99
zebrata (Nomia), 108
zebrus (Halictus), 213
zeus (Pompilus), 102
zonata (Anthophora), 109
zonatulus (Halictus), 108
LEPIDOPTERA.
Abantis, 203
abdera (Acrea), 186
! abdominalis (Euproctis), 397
abietis (Calliteara), 472
Abisara, 194
abjecta (Cyclopides), 204
(Euproctis) 425
», (Somena), 425
abraxas (Pentila), 196
abraxata (Nyctemera), 61
abraxoides (Deilemera), 59, 60
3 (Pitasila), 59
absurda (Deilemera), 79
absurdum (Leptosoma), 79
5 (Nyctemera), 79
Abynotha, 479
abyssinica (Belenois), 145
accepta (Deilemera), 79
acceptum (Leptosoma), 79
acheemenes (Charaxes), 206
achine (Teracolus), 157
Aclonophlebia, 481
Acnissa, 429
Acrea, xxii, xli, lxiv, Ixv, 149, 150,
151, 155, 185, 186, 206
acreina (Deilemera), 67
(Nyctemera), 67
be)
2?
acrisia (Dasychira), 154, 474
», (Deiopeia), 474
acronycta (Dasychira), 471
acta (Satyrus), xi
actia (Precis), XXxli, XXXill
Actinote, 195
actinotina (Telipna), 194, 206
aculeata (Metanastria), xxii
acuta (Anthela), 449
5, (Darala), 449
Acyphas, 458, 459, 463, 478
adala (Leelia), 443
adamsi (Jolaus), 199, 206
adara (Lelia), 444
», (Procodeca), 444
addita (Anthela), 446
», (Darala), 446
Adlullia, 396, 417, 418, 419
admatha (Acreea), 185
adriana (Anthela), 445
», (Darala), 445
adspersa (Cropera), 391
5 (Liparis), 391
adusta (Charaxes), 193
»» (Darala), 449
wgeria (Pararge), xliil
worota (Deilemera), 56
wgrotum (Leptosoma), 56
a (Nyctemera), 57
wres (Deilemera), 62
,, (Leptosoma), 62
africana (Lepasta), 433
» (Mardara), 433
agagles (Leptosoma), 84
agatha (Neptis), 150, 151, 189
», (Papilio), 189
Aglages, 68
Aglaosoma, 498
agraphis (Monotrichtis), 185
Agrotis, lxvii
Agyrta, v, vi
alba (Aroa), 380
(Caltura), 386
(Cispia), 386
(Deilemera), 83
», (Dura), 495
(Euproctis), 428
(Leucoma), 380
(Nyctemera), 83
(Redoa), 380
alberti (Baoris), 204
albertus (Monethe), 539
albescens (Dasychira), 471
5 (Euproctis), 399
albibasalis (Dasychira), 469
i> (Ilema), 469
albicans (Dasychira), 495
9?
(
var. cordula (Satyrus), XXXVil
CXXxil )
albicans (Dura), 495
albifascia (Orgyia), 438
or (Pantana), 438
albilunulata (Bathmochtha), 464
albinotata (Dasychira), 470
5 (Thamnocera), 470
albipuncta (Deilemera), 56, 85
_albodentata (Euproctis), 427
56 (Pida), 427
| albofascia (Lymantria), 432
| albofasciata (Riodina), 535, 549
albolunata (Lymantria), 485
alcesta (Leptosia), 201
», (Papilio), 201
alcippe (Atella), ii
alcippina (Acrea), 186
nsw
alemzeon (Ariecoris), 547 .
| Aletis, xxxix
aliena (Leucoma), 393
_;, (Porthesia), 393
Alope, 486
alternata (Deilemera), 70
os (Nyctemera), 70
aluensis (Deilemera), 73, 76, 78
(Nyctemera), 76
as var. (Deilemera), 77
amalfreda (Charis), 544
Amana, 498
/ amarah (Lycenesthes), 148
amata (Dasychira), 483
; Amauris, xxxix, xl, 184
amazonica (Caria), 538
ambiorix (Neomcenas), 279
35 (Neosatyrus), 283, 284
amelia (Teracolus), 147
amenaida (Pentila), 195
americanus (Hesperia), 294, 299
- (Syrichthus), 294
americensis (Strymon), 290, 299
so (Thecla), 290
amica (Deilemera), 68
», (Nyctemera), 68
amicus (Aglages), 68
amosa (Deilemera), 59, 85
Amphidasis, xxxv, 311-367
amphideta (Euproctis), 413
amphitrite (Callidryas), 290
ampla (Enome), 481
(Euproctis), 391
;, (Pantana), 437
amplata (Dasychira), 471
amplicornis (Pterolocera), 453
amplificata (Deilemera), 82
(Tanada), 82
(Tripheromera), 82
99
99
| 9
99
alcippoides (Limnas), 148, 145, 149
alcippus (Limnas), 143, 145, 146, 149,
{ exxxii )
Amsacta, 498
amulia (Crenis), 189
», (Papilio), 189
anacardii (Papilio), 188
of (Salamis), 188
Ana, Xxvili
anartoides (Teia), 457
Anatole, 545
Anaxila, 429, 461, 471, 475
Anchyneura, 462
andina (Scolitantides), 264,
Anepa, 478
angulata (Dasychira), 471
angulifera (Procodeca), 441
anguligera (Artaxa), 415
5 (Euproctis), 413
angustata (Catuna), 189
angustatum (Euomma), 189
anna (Argynnis), 287
5, (Kuproctis), 417
annulata (Deilemera), 68
5 (Nyctemera), 68
annulatum (Leptosoma), 68
annulifer (Oxypalpus), 204
annulitera (Lemonias), 542
Anomceotes, 435
Anosia, 562, 572
antalus (Deudorix), 198
», (Dipsas), 198
antarctica (Satyrus), 279, 280
antareticus (CEneis), 281
antennata (Lymantria), 485
Anteros, 535, 549
anthedon (Diadema), 188
90 (Hypolimnas), 188
Anthela, 376, 445-453
Anthora, 439, 462
anthracinum (Leptosoma), S4
Anthrocera, 248
antica (Arestha), 462
», (Dasychira), 464, 492
(Dediama), 464
», (Kuproctis), 397, 429
(Gazalina), 387
», (Lacida), 429
auticlea (Charaxes), 193
», (Papilio), 193
Anticyra, 462
antifaunus (Hypolyciena), 199
antigone (Teracolus), 142, 161
Antigonus, 203
antilope (Precis), X, Xil, XXXil, Xxxili,
20.0.5 Ilsy/
antimachus (Drurya), Ixiti, lxiv, Ixv
antinorii (Deilemera), 67
(Nyctemera), 67
(Papilio), xxxix
Antipha, 396, 429, 464
>
antiphates (Euproctis), 429
| Antiphella, 382, 390, 391
antra (Rajacoa), 435
aolwnsis (Nyctemera), 72
apensis (Nyctemera), 84
Aphantopus, xlii
Aphneus, 199, 200, 206
apicalis (Arna), 425
(Artaxis), 425
(Bembina), 409
(Deilemera), 66
» (Euproctis), 409, 425
(Nyctemera), 66
a) (aida). 43m
Apina, 497
Apodemia, 545, 546, 550
Apoprogones, xiv, 137, 138
Aporia, 1
Appias, 202
approximans (Orgyia), 458
approximata (Anticyra), 462
aprilina (Odontopera), 312
apsara (Dasychira), 387
», (Gazalina), 387
Apterogynis, 458
aquilina (Lycena), 242
aquilo (Lycena), liii, 242’
arcania (Ceenonympha), xliii
archesia (Precis), xii, 157
archippus (Basilarchia), 572
sa (Limenitis), 572
arclada (Euproctis), 409
arctata (Deilemera), 64
» (Nyctemerw), 64
Arctia, 473
Arctornis, 377, 379
arcuata (Charis), 539, 550
arcuatum (Nyetemera), 78, 83
Arestha, 461, 462
arethusa (Satyrus), xliii, xliv
arga (Dasychira), 473
Arge, 281
Argema, Xxiil
argenna (Cypra), 390
», (Olapa), 390
argentata (Dasychira), 474
- (Euproctis), 406
argentea (Himala), 386, 387
A (Redoa), 386
argenteo-maculata (Hepialus), 501
(Sthenopis), 501
274,
+B)
29
99
99
argenteus (Argyrophorus),
298
3 (Chionobas), 282
argia (Eronia), 206
Argila, 461, 464
argiope (Caria), 588
| Argopteron, 295, 299,
Nee
282,
300
(
Argynnis, lii, xxxvii, 241, 242, 244, |
264, 277, 286, 287, 299
argyrocyclus (Aphneus), 200
argyrodice (Tatochila), 292, 293
Argyrophenga, 279, 284
Argyrophorus, 274, 282, 298
Aricoris, 546, 547, 548, 550
Arna, 396, 425
Aroa, 380, 407, 409, 410, 412, 415,
432, 453, 454, 455, 456, 457, 475
arrogans (Artaxa), 405
Artaxa, 395-427
artemis (Nyctemera), 56
arthemis (Basilarchia), 556
a (Limenitis), 556, 572
aryama (Lymantria), 481
Arycanda, 498
asclepia (Cricosoma), 540, 550
55 (Symmachia), 540
Aslauga, 197
ascetria (Lymantria), 492
aspersa (Dasychira), 474
a (Tearosoma), 474
Aspilates, 243
aspilota (Anthela), 445
assimile (Deilemera), 70, 71
(Leptosoma), 70
(Nyctemera), 70
assimilis ((Hneis), 240
asteria (Melitiea), xlix
astyoche (Pierella), 565
asvata (Dasychira), 465
atahualpa (Polyommatus), 289
Atasca, 54, 55, 56, 57
Atella, ii, 142, 187, 206
Aterica, 189
atestacea (Leelia), 442
athalia (Melitea), xlix, 1], li, li
,, var, navarina (Melitiea), xlix
atlantica (Liparis), 483
5 (Lymantria), 483
atolmis (Acriea), 155
atomaria (Artaxa), 425
re (Euproctis), 416, 425
atralba (Nyctemera), 53, 80
atrella (Aroa), 454
atrescens (Aroa), 454
atrinotata (Antiphella), 391
% (Olapa), 391
atripuncta (Euproetis), 428
atrisignata (Euproctis), 423
atrosquama (Huproctis), 416
Pe (Gogana), 416
Augiades, xiii
aurantiaca (Charnidas), 454
(Eurygona), 531
“ (Porthesia), 395
aurata (Eupheedra), 191
9?
93
99
| aurelia (
cxxxiv )
aureipennis (Argopteron), 295, 299
(Syrichthus), 295
Melitea), xlix, 1, hi
» var, norvegica (Melitea), xxxvii
aureofasciata (Huphdra), 191
auriflua (Bombyx), 392
aurifrons (Euproctis), 385
ap (Leucoma), 385
aurimna (Emesis), 536
aurinia (Meliteea), li
auripes (Ivela), 388
;, (Leucoma), 388
aurivillii (Acrea), xli
rs (Nyctemera), 83
aurivina (Ceenina), 154
auro-limbata (Orgyia), Xxxi, XxXii
aurora (Lymantria), 484, 485, 488
,, var. fusea (Lymantria), 488
australasiz (Lelia), 449
australis (Orgyia), 460
authe (Hurygona), 532, 549
autodice (Synchloe), 292
5 (Tatochila), 292, 299
autumnaria (Ennomos), lix, lxvili
auxo (Teracolus), 157, 162
axenana (Epalxiphora), xlvii
axenus (Lemonias), 545, 550
Axiocerses, 200
Azanus, 148
bacotii (Gorgopis), 504, 507
», (Oncoptera), 502, 504, 508
Beotis, 545
bieticus (Cupido), 201
(Papilio), 201
», (Polyommatus), 144
baliolalis (Urocoma), 478
ballus (Thestor), xxxi
bammakoo (Elymnias), 184
a (Melanitis), 184
banane (Lymantria), 494
Baoris, 204, 205, 206
barbara (Euproctis), 417
Barhona 482, 490
barii (Ceratinia), lvi
baruna (Dasychira), 476
», (Somena), 476
Baryaza, 479
basalis (Antipha), 464
(Argila), 464
(
(
(
39
Artaxa), 414
Dasychira), 464
Euproctis), 414
5, (Perina), 480
basifurea (Cnethocampa), 461
basigera (Darala), 447
», (Dasychira), 464
Basilarchia, 556, 572, 573
basistriga (Dasychira), 463
2?
9
9?
( cxxxy, )
basistriga (Phineca), 463
basivitta (Olene), 464
baswana (Pantana), 437
Bathmochtha, 462, 464
batjana (Lymantria), 492
baulus (Deilemera), 55, 75, 76
», (Leptosoma), 75
beatrix (Bombyx), 487
(Lymantria), 487, 488
beckeri (Cymothoé), 192
», (Diadema), 192
Belenois, 143, 144, 145, 147, 148, 150,
151, 152, 154, 155
belina (Buna), xxiii
bella (Mesosemia), 529, 530
beltiana (Cremna), 531
Bembina, 396, 409
Beralade, 498
berisalensis (Melitza), li
betularia (Amphidasis), xxxv, 311-367
bhana (Dasychira), 468
bhascara (Lymantria), 485
bias (Papilio), 293, 299
bibulus (Lachnocnema), 206
bicolor (Lyczna), 289
», (Orgyia), 437
5, (Pantana), 437, 439
», (Thecla), 289, 290, 299
Bicyelus, 184
bidentata (Euproctis), 414
5 (Odontopera), xxxv, 311-370
bifascia (Euproctis), 401
bifasciata (Beeotis), 545
biformis (Deilemera), 66
», (Nychthemera), 66
», (Nyctemera), 66
bigutta (Euproctis), 398
», (Marbla), 437
», (Soloé), 437
bijunctella enone) 58, 60
(Pitasila), 5
bimaculata (Deudorix), S6
Ls (Euproctis), 398
- (Myrina), 198
binotata (Anthela), 446
», (Leptocneria), 446
bipartita (Cherotricha), 423
PP (Euproctis), 423
bipunctapex (Euproctis), 424
(Somena), 424
Birnara 437, 489
bissexguttatus (Butleria), 297, 299,300
55 (Steropes), 297
bistigmigera (Aroa), 456
biundulans (Cifuna), 479
bivittata (Lymantria), 493
» (Pegella), 493
bizonoides (Euproctis), 402
bizonoides (Lacipa), 402
blanchardii (Tatochila), 295
boguensis (Belenois), 147, 148
boisduvalii (Diadema), 189
(Erebia), 284, 285
(Hipparchia), 285
(Pseudacreea), lxiv, 189
boleora (Adlullia), 419
Bombyx, 385, 392, 395, 411, 417, 430,
434, 435, 440, 449, 458, 482, 486,
487, 491
bonasia (Acreea), 186
», (Papilio), 186
bodpis (Precis), 142
boothii (Colias), 111, 242, 244
borbonica (Parnara), 206
bore, var. taygete ((neis), 240
Boreconia, 462
brahami (Aphneeus), 199
brasidas (Papilio), xxi
brassolis (Liphyra), x
brenda (Terias), 202
Brenthis, xliii
brevicornis (Leucoma), 441
29
| brevivitta (Artaxa), 400
brigitta (Terias), 150, 151, 155, 206
brunneicosta (Dasychira), 470
“3 (Ilema), 470
brunneiplaga (Lymantria), 491
brutus (Charaxes), 193
», (Papilio), 193
buana (Lielia), 442
», (Phragmatobia), 442
Bunea, xxi
buquetii (Leuceronia), 152
burica (Deilemera), 58
», (Nyctemera), 58
butleri (Erebiola), 283
Butleria, 295, 296, 297, 299, 300
buxtoni (Acriea), xxii
Byblia, 143, 189
cabira (Acriea), XX1i
Cadrusia, 462, 476
Caduga, 565
eecias (Charis), 539
excilia (Acreea), 206
Ceenina, 154
caffra (Nyctemera), 53
», (Otroeda), 53
caffraria (Buna), xxiii
cafra (Bombyx), 434
,, (Nyctemera), 434
calais (Teracolus), 158
calamaria (Lielia), 442
c-album (Grapta), xxvi
» (Polygonia), xxvi, xxvii, xXxvili
calesia (Euproctis), 420
caliginosa (Deudorix), 197, 206
( exxxvi )
Caligo, 562, 564, 565
Callia, 498
Callidryas, 290, 299
calligramma (Mardara), 433
calligraphum (Cricosoma), 540
calliope (Stalachtis), iv
pe tbevene) 289
Calliteara, 462, 472, 473,
Callophrys, li
Caltura, 386
calva (Euproctis), 406
calvertii (Elina), 275, 298, 301
Calydna, 541
calypso (Papilio), 202
san (eieris) 202:
camiena (Acrea), 185
camerona (Deudorix), 197
camerunica (Planema), 187
camillus (Cyrestis), 189
ao (Papilio), 189
candida (Lelia), 441
pee (Anthela}, 450
Ae (Darala), 450
cangia (Dasychira), 476
canifascia (Orgyia), 460
cara (Lymantria), 487, 491
Caragola, 376, 377, 378
carausius (Anteros), 536
cardinalis (Lelia), 442
cardui (Papilio), 187
», (Pyrameis), 187
», (Vanessa), Ixvii, 144
Caria, 538, 549
carissima (Deilemera), 63, 64
carmentis (Acrzea), 186
carnecolor (Lymantria), 486
carneola (Barhona), 490
carneotincta (Anthela), 451
earriala (Lymantria), 495
Carterocephalus, lili, 297
carus (Anteros), 535, 536, 549
carye (Pyrameis), 287, 299
Casama, 461
castor (Charaxes), 193
»» (Papilio), 193
castrensis (Malacosoma), viii
Castulo, 497, 498
Catacroptera, 188
catala (Euproctis), 416
catilla (Catopsilia), 157
catocaloides (Dasychira), 476
x, (Mardara), 476
Catopsilia, 156, 202
Catuna, 189
Caviria, 377, 378
cebrene (Precis), 143, 145, 206
cebron (Pieris), 202
Cebysa, 498
Celenorrhinus, 206
cellularis (Pseuderesia), 196
celsa (Deilemera), 81
5, (uproctis), 399
,, (Nyctemera), 81
celsum (Leptosoma), 81
cenea (Papilio), xxl, xxv
cenis (Deilemera), 55, 68
», (Phalena), 68
censors (Darali), 452
cepha (Aricoris), 548
cerasina (Euproctis), 426
Ceratinia, iv, lvi
cerebosa (Lymantria), 483
ceres (Kuphedra), 190
», (Lycorea), iv, lv
», (Papilio), 190
cerigoides (Dasychira), 473
(Janassa), 473
Cerura, xxiii
cervina (Artaxa), 407
», (Euproctis), 407
», (Repena), 443
cervinata (Porina), 502, 508
cerymica (Hesperia), 205
», (Ploetzia), 205
ceylanica (Orgyia), 460
Cheerotricha, 396, 413, 416, 417, 418,
419, 420, 422, 423
chalana (Dasychira), 466
chalcis (Euryphura), 192
», (Harma), 192
chalcophanes (Paleeomicra), 503, 508
chalybe (Ismene), 205
», | (Rhopalocampta), 205
chameleon (Pamphila), 205
at (Platylesches), 205
Chamvelimnas, 534, 549
Chapra, 146, 204
Charagia, 501, 503, 504, 507
charax (Aroa), 457
,, (Neurophana), 457
| Charaxes, xxii, 193, 194, 206
chariclea (Argynnis), ili, 241, 244
5, (Papilio), 241
Charis, 539, 544, 550
charma (Cispia), 430
Charnidas, 397, 489, 442, 443, 444,
454
| cheela (Euproctis), 412
| Chilades, 143, 145
| chilensis (Lyczna), 289
» (Scolitantides), 289, 299
», (Terias), 290, 299
chiliensis (Cosmosatyrus), 280, 281, 298
», (Satyrus), 280
», var. (Cosmosatyrus), 301
chinensis (Dasychira), 468
i
(| exxxvil
Chiotiema, 497
chione (Colias), 243
chionitis (Euproctis), 398
Chionobas, 240, 282
Chionophasma, 392
chirunda (Euproctis), 422
chismona (Heteronygmia), 480
chloris (Mylothris), 201
», (Papilio), 201
chloroptera (Dasychira), 47
chlorozonea (Eublemmistis)
chordigera (Oligoclona), 38
chorimene (Precis), 188
93 (Vanessa), 188
christi (Erebia), xxxvii
chromis (Deilemera), 67
», (Nyctemera), 67
chrysame (Caria), 538
chrysippus (Danais), xxxix, 184
(Limnas), xi, xxii, xxix, 142,
143, 144, 145, 146, 149, 150,
151. 152, 565, 574
7 (Papilio), 184
chrysolopha (Gazalina), 387
ee (Liparis), 387
chrysophia (Euproctis), 407, 408
i (Orgyia), 407
chrysophzeus (Notolophus), 407
Chrysophanus, xxxvii, xliii, xlv, Lxil
chrysophila (Artaxa), 595
Chrysopsyche, 497
chrysorrheea (Bomby x 5)
Cibyra, 501, 502, 504, 507
Cidaria, 243
Cifuna, 479
Cimola, 435
cinctata (Dasychiva), 471
cinerascens (Anthela), 449
fj (Darala), 449
cinerea (Euproctis), 420
> cinnamomea (Aroa), 454
5 (Charnidas), 454
cireumdata (Genusa), 437
Cispia, 386, 391, 399, 418, 430
citana (Dasychira), 463
», (Utidava), 463
Citheerias, 565
citheron (Charaxes), xxii
citrina (Artaxa), 399
Citrinophila, 196
civitta (Kuproctis), 412
clara (Aroa), 455
», (Caragola), 377
», (Caviria), 377
clathrata (Deilemera), 70
clathratum (Leptosoma), 70
a (Nyctemera), 70
celaudianus (Euryphura), 192
F
.
XX1V, Xxvl
?
/
2)
)
| clelia (Papilio), 187
», (Precis), 151, 187
clementi (Anthela), 446
», (Darala), 446
cleodora (Eronia), 148
Clerome, lvi
Clethrogyna, 458
cloantha (Catacroptera), 188
», (Papilio), 188
Cluaca, 498
Cnethocampa, 461
Cobanilla, 392
coccinata (Cymothoé), 193
59 (Harma), 193
coctei (Epinephele), 277, 298
», (Erebia), 277
ccelestina (Euagra), vi
oy _{ IPAREXGrIS)), 5:<
ccenobita (Hesperia), 189
», (Pseudoneptis), 189
Ccenonympha, iii, xlii, xliii, xliv, 241
ccenonymphina (Neomeenas), 277, 298
cceruleifascia (Dasychira), 463
(Thamnocera), 465
coleta (Deilemera), 54, 55, 64
(Nyctemera), 53, 64
», (Phalena), 64
Colias, iii, iv, xxxvii, xliil,
243, 244, 264, 291, 299
collina (Lycena), 288
», (Scolitantides),
colon (Charnidas), 443
columbina (Atella), 206
Colussa, 376, 445, 446, 450
Comana, 497
combinata (Euproctis), 430
cometaris (Aroa), 456
comma (Leucoma), 385
», (Ocinara), 385
commutanda (Euproctis), 412
comparata (Genusa),437
5p (Themaea), 413
complens (Lacida), 429
complicata (Dasychira), 477
concolor (Lymantria), 486
confluens (Nyctemera), 83
confusum (Leptosoma), 70
conica (Nyctemera), 68
connexa (Anthela), 449
», (Darala), 449
consanguinea (Planema), 187
consobrina (Deilemera), 78
Pr (Nyctemera), 78
consocia (Euproctis), 399
consors (Anthela), 452
Darala), 452
Deilemera), 65
Leptosoma), 65
be)
liii, 242,
288, 299
+)
(
Se an
(
(
>
(| exxxvill )
conspersa (Anthela), 451 Cyenia, 443
a (Artaxa), 413 cydippe (Nyctemera), 82
(Darala), 451 | eyena (Caragola), 377
5 (Euproctis), 413, 421 | 5, (Caviria); 376; 347
contracta (Nyctemera), 83 », (Leucoma), 378
cordula (Satyrus), xlii », (Redoa), 378
corynetes (Eresina), 197 cymbicornis (Redoa), 378
Cosmethis, 498 _cymodoce (Kallima), 188
Cosmosatyrus, 274, 279, 280, 281, 298, rr (Papilio), 188
301 Cymothoé, 192, 193
Cossus, x, xxxvii, lx cynorta (Papilio), 202
costalis (Antipha), 429 Cynthia, 565
»» .(Dasychira), 467 cynthia (Charaxes), 193
5, (Euproctis), 429 Cypra, 382, 390, 435, 437, 497
», (Lymantria), 482 Cyrestis, 189
,, (Melia), 467 eytheris (Argynnis), 286, 287, 299
costiplaga (Lacida), 466 », (Papilio), 286
Cozola, 396, 422 Dactylorhyncha, 444
crambis (Chionobas), 240 deedalus (Hamanuinida), 190
,, (Ckneis), 240 », (Papilio), 190
crameri (Melinza), iv, liv, lv daira (Acrva), 151
crateegi (Aporia), ii », (Teracolus), 145, 146, 147, 155,
crausis (Dasychira), 474 160, 161, 162
Creagra, 385 dalbergiz (Dasychira), 465
Cremna, 530, 549 .| dana (Euproctis), 408
Crenis, 189 danae (Teracolus), 157
crenulata (Anthela), 451 Danais, xxxix, 184
creona (Papilio), 202 daphnandre (Charagia), 501, 504
5, (Pieris), 202 Dapidodigma, 199
crescens (Nyctemera), 80 Daplasa, 428, 430
Cricosoma, 540, 550 Darala, 376,420, 445-453
Crinola, 454 dardanus (Papilio), xxi, xxv, 202
Crinopteryx, 1, xlvii darwinia (Mimacriea), 196
crocale (Catopsilia), 156, 157 Dasychira, 154, 375, 387, 461-477, 482,
crocata (Euproctis), 410 483, 488, 492, 495
», (Liparis), 410 dealbata (Creagra), 385
crocea (Euproctis), 405 », (Liparis), 385
», (Hectomanes), 502, 508 debilis (Mesene), 541
», (Nygmia), 405 decolorata (Cyclidia), 431
,, (Teara), 405 5 (Pida), 431
crocicollis (Liparis), 390 decora carye (Hamadryas), 287
crocipes (Cypra), 382 decorata (Pseudarbessa), vi
s, (Leucoma), 382 », (Stibomorpha), 278
crockeri (Euphiedra), 191 decussata (Cheerotricha), 417
cromptoni (Dasychira), 467 Dediama, 461, 464
Cropera, 391, 410 deficiens (Dreata), 450
Crorema, 391 deficita (Euproctis), 421
cruentata (Aricoris), 548 » (Teara), 421
cuama (Precis), Xxxili, xxxiv Deilemera, iv, 53-85
cuneilinea (Metanastria), xxiii | deione (Melitzea), xlix, li
cunninghami (Colias), 291 Deiopeia, 154, 474
Cupido, 201, 206 ' delagorguei (Ludia), xxiii
curvata (Cnethocampa), 461 | delicata (Dasychira), 463
curvifera (Lymantria), 491 », (Notohyba), 463
», (Pegella), 491 delicatula (Cozistra), 390
curvivirgata (Lielia), 470 delineata (Anthela), 450
Cyclidia, 431, 433 45 (Darala), 450
Cyclopides, 204, 295, 296 (Genusa), 437
(
delius (Hypanartia), 187
», (Papilio), 187
», (Parnassius), lil
demodice (Pieris), 292
», (Tatochila), 292, 293,
300
demodocus (Papilio), xxi, 202
demosthenes (Caligo), 565
Dendrophleps, 376, 378
denotata (Drymonia), 461
denticulata (Anthela), 447
. (Teara), 447
dentifascia (Nyctemera), 84
denuba (Antigonus), 203
», (Eagris), 203
denudata (Sitvia), 388, 390
depauperata (Porthesia), 394
dermaptera (Myrina), 198
Deroca, 497
dersa (Euproctis), 404
desjardinsii (Terias), 202
_ (Xanthidia), 202
desolata (Monotrichtis), 206
despecta (Porina), 502
detersa (Enome), 482
,, (Lymantria), 482
Deudorix, 197, 198, 206
devestita (Lielia), 440
aA (Odagra), 440
dewara (Orgyia), 459
dexamene (Argynnis), 287
Diadema, 188, 189, 192
diaphana (Leucoma), 381
», (Redoa), 381
Diaphone, xxiv
dica (Caragola), 377
,, (Redoa), 376, 377
Dicreagra, 496
didyma (Melita), xlix, hi
Diestoyyna, 192
difficilis (Aroa), 456, 475
»» (Dasychira), 475
diffusa (Lacipa), 465
digramma (Bombyx), 411
_ (Euproctis), 411
Dilina, xiv
Dione, 565
dionysos (Papilio), xl
diophthalma (Ommatoptera), 449
Dipsas, 198
disa (Erebia), liii, 239
», (Papilio), 239
disealis (Aroa), 456
discinota (Euproctis), 416
discirufa (Leucoma), 384
discivitta (Heracula), 431
discolor (Topomesa), 392
disjuncta (Lymantria), 445
Cxxxix |)
299,
Dismorphia, 558, 573
dispar (Bombyx), 482
,, (Deilemera), 73
», (Lymantria), 482
», (Pantana), 437
», var. japonica (Liparis), 483
disparilis (Numenes), 432
Ye var. separata (Numenes), 432
disrupta (Deilemera), 58, 60
dissoluta (Lymantria), 484
distincta (Deilemera), 58, 69, 72, 85
», (Nyctemera), 69
», (Osmodes), 204
distinguenda (Rilia), 464
distracta (Artaxa), 427
», | (Euproctis), 427
divisa (Dasychira), 464
.,, (Hloria), 436
,, (Euproctis), 380, 397
5, (lostola), vi
5, (Leucoma), 380
», (Marbla), 436
Doleschallia, xxviii
doleta (Ypthima), 185
dominica (Diaphone), xxiv
doriz (Nyctemera), 57
dorilis (Chrysophanus), xxxvil
», var. subalpina (Chrysophanus),
XXXViL-
dorippus (Limnas), 142, 143, 149, 151,
574
dorothea (Monotrichtis), 185
re, (Papilio), 185
doryssus (Mechanitis), iv
doubledayi (Acreea), xxii
s (Nyctemera), 68
Dreata, 450
dregei (Aroa), 456
», (Orgyia), 456
drucei (Deilemera), 73
5, (Kuproctis), 408
Drurya, ]xiii, xv
drya (Callidryas), 290, 299
5, (Papilio), 290
dryas (Epinephele), 276
», (Satyrus), xli, xliii, xliv
dryinopa (Hybocampa), 249
Drymonia, 461
dryope (Eurytela), 188
», (Papilio), 188
dubius, (Hypolimnas), 188
», (Papilio), 188
dukinfieldia (Charis), 539, 550
dulcinea (Lymantria), 493
Dulichia, 396, 410
dumolinii (Lophostethus), xxiv, xxvi
Dura, 495
dynamene (Teracolus), 158
Eagtis, 203
eanes (Mesene), 540, 550
echo (Ophiusa), xxiv
eddela (Lymantria), 494
edmondsii (Elina), 275
a (Epinephele), 276, 298
BS (Neomcenas), 279, 298
edulis (Jana), xxiii
edusa (Colas), xliii
edwardsii (Euproctis), 405
re (Teara), 405
effinia (Eurygona), 532
egerina (Leucoma), 380
egesta (Cymothoé), 192
», (Papilio), 192
egina (Acrea), Ixiv, 185
»» (Melinzea), iv, liv
so) CRapilio) 185
egregia (Kuproctis), 419
eleala (Deudorix), 197
5, (Hypolycena), 197
elegans (Calliteara), 474
», (Dasychira), 474
elegantula (Abantis), 203
eleus (Chrysophanus}, 1xii
», (Eupheedra), 190
,, (Papilio), 190
eleuteria (Bombyx), 435
», (Cimola), 435
», (Cypra), 435
Elina, 274, 275, 298, 301
elizabetha (Authela), 445
Ae (Odonestis), 445
elmira (Eurygona), 531
Eloria, 436
elpinice (Lemonias), 542, 550
Elymnias, xli, 184
emesina (Symmachia), 537
Emesis, 536, 537, 549
encedon (Acriea), xxii, 151, 186
», (Papilio), 186
endoplagia (Euproctis), 400
Endromis, ix
endymion (Lyciena), 288, 289
Knnomos, lix, lxvili, 451
Enome, 481, 482
enos (Aroa), 475
»» (Dasychira), 475
enotrea (Ergolis), 188
», (Papilio), 188
enthysana (Porthesia), 393
enysil (Porina), 502
epiea (Papilio), 186
», (Planema), 186
Epalxiphora, xlvii
epaphia(Appias), 202
», (Papilio), 202
ephyne (Mesosemia), 530
exl )
| epigone (Teracolus), 147, 148
epijasius (Charaxes), 193
Epinephele, xliii, 276, 277, 298
Epipyrops, 497
Erastria, 463
Erebia, xxxvii, xlv, xlvi, lili, 239, 240,
244, 277, 284
Erebiola, 283
erecta (Artaxa), 399
Eresina, 197
Ergolis, 188
Eriocrania, 502, 503
Eriogaster, 246
eriphyle (Erebia), xlv
eris (Teracolus), 157
‘| ero (Erebia), 240
Eronia, 148, 206
erythrias (Haplopseustis), 429
erythrina (Lomadonta), 480
Esthemopsis, v
etesipe (Charaxes), 193-
», (Nymphalis), 193
ethalion (Charaxes), xxii
-etheocles (Charaxes), 194
| », - (Papilio), 195
| ethosea (Mesoxantha), 188
», (Papilio), 188
Etobema, 439
Euagra, v
Eublemmistis, xxiv, Xxvi
eubule (Kurygona), 532
eucharis (Teracolus), 157
Eudasychira, 462
Eueides, iv, lv
euhemerus (Eurygona), 532
eumeus (Clerome), lvi
Euomma, 189
eupale (Charaxes), 194
», (Papilio), 194
eupepla (Kurygona), 533
Eupheedra, 190, 191, 192, 206
euphrosyne (Brenthis), xliii
eupiola (Hurygona), 533
Eupleea, 565, 575
eupompe (Teracolus), 146, 147, 148,
150, 152
Euprepia, 443
Euproctidion, 462, 469
Euproctis, 380, 385, 391, 392, 394-450,
460
Euptera, 193
Euptoieta, 286, 299
| eurinome (Euxanthe), 195
| 37 (Papilio)5 193
eurissus (Jolaus), 199
se (Bapilio) 3199
eurydice (Emesis), 536, 549
(Lymantria), 483
+]
(exli >)
eurydice (Porthetria), 483
eurygania (Pantana), 459
Eurygona, 529, 531, 532, 533, 549
eurylochus (Caligo), 564
Euryphene, 192, 206
Euryphura, 192
Eurytela, 188
eurytus (Papilio), 189
», (Pseudacrea), 189
Euschemon, xiv, 138
Eusemia, 431
euthria (Calydna), 541
Euxanthe, 193
euxanthe (Colias), 264, 291, 299
evagore (Teracolus), xii, 142, 160, 161,
162
evarne (Teracolus), 147, 148, 152
evergista (Deilemera), 54, 61, 62
aA (Phalena-Geometra), 61
evias (Erebia), xlv, xlvi
ve hispanica (Erebia),
evippe (Papilio), 202
»» (Teracolus), 202
excellens (Anthela), 452
3 (Darala), 452
excisa (Anthela), 452
>, (Darala), 452
exclamationis (Euprepia), 443
oe (Leela), 443
eximia (Charagia), 502
exornatus (Carterocephalus), 297
extendens (Deilemera), 72
Aa (Nyctemera), 72
extorta (Dasychira), 465
fabia (Eurygona), 533, 549
faceta (Euproctis), 427
falcata (Rigema), 462
falkensteinii (Cupido), 201
fallax (Deilemera), 65
», (Nyctemera), 65
familiella (Crinopteryx), 1, xlvil
famula (Nyctemera), 53
fannia (Lemonias), 545, 550
fasciata (Deilemera), 75
(Dulichia), 410
(Erebia), 239, 244
(Euproctis), 410
(Leelia), 440
(Lielioides), 440
», (Nyctemera), 75, 83
fasciolata (Hesperia), 294
55 (Hylephila), 294, 299
fastidiosa (Emesis), 536
fatima (Symmachia), 537
Faunula, 282, 283, 298, 301
faunus (Oxylides), 198
», (Papilio), 198
feminula (Dasychira), 468
9 xlv, xlvi
39
be)
be)
29
feminula (Mardara), 468
fenestrata (Leucoma), 384
(Macrauzata), 384
| ferruginea (Anthela), 449
5 (Darala), 449
ferruginosa (Anthela), 446
fervens (Darala), 449
fervida (Artaxa), 409
», (Euproctis), 409
figlina (Anthela), 446
(Darala), 446
», (Lelia), 440
fimbriata (Porthesia), 893
(Teara), 393
flabellaria (Olapa), 390
a (Phalna), 390
flammeola (Heteroygmia), 432
flava (Aroa), 415
flavala (Anthela), 452
flaveofusca (Aroa), 455
flavescens (Leptosoma), 84
(Leucoma), 379
(Nataxa), 496
(Perna), 496
. (Redoa), 379
flavicollis (Aroa), 454
A (Crinola), 454
flavicosta (Euproctis), 425
flavifascia (Nataxa), 496
flavimacula (Dasychira), 465
flavinata (Artaxa), 406
», (Euproctis), 406
flavinotata (Aclonophlebia), 481
flavipectus (Sapelia), 389
flavipes (Caltura), 386
», (Cispia), 386
flavociliata (Euproctis), 422
flavomaculata (Lycenesthes), 200
LB)
99
99
by)
flavomaculatus (Butleria), 296, 297,
299
(Syrichthus), 296
flavonigra (Euproctis), 394
flesus (Papilio), 203
», (Tagiades), 203
flora (Hipparchia), 275
(Leucoma), 383
», (Pedaliodes), 275, 298
», (Satyrus), 275
florella (Catopsilia), 202
», (Papilio), 202
florida (Euproctis), 403
florus (Lemonias), 543, 545
Fodinoidea, 431
forbesi (Cypra), 435!
», (Rajacoa), 435
forestan (Papilio), 205
», (Rhopalocampta), 205
fracta (Artaxa), 429
2?
( exlii_ )
fracta (Lelia), 442
fractifascia (Neomcenas), 278, 298
franklinii (Lyceena), 242
fraterna (Artaxa), 411
,, (Kuproctis), 411
freija (Argynnis), XXxvil
frigga (Argynnis), ili, XXXxvii
», (Papilio), 242
5, var. improba (Argynnis), 242
fruticolens (Butleria), 295, 299, 300
3 (Cyclopides), 295
Fa var. pulcher (Cyclopides),
296
. var. ae ae (Cyclo-
pides), 2
3 var. Pe ate (Cyelo-
pides), 296
fuciformis (Macroglossa), xxvi
fuliginea (Pyramocera), 496
fuliginosa (Lymantria), 488
fulleri (Deilemera), 67
5, (Nyctemera), 67
fulva (Artaxa), 406
», (Euproctis), 406
», (Hesperia), 295
(Hylephila), 295, 299, 300
,, var. (Hylephila), 300
fulvescens (Charaxes), 194
fulviceps (Anepa), 478
AD (Charnidas), 478
ae (Icta), 460
fulvinotata (Cropera), 391
re (Olapa), 391
faleinincts (aproctin’ 401
es (Lopera), 401
fulvistriata (Euproctis), 408
fulvonigra (Porthesia), 395
fulvovittatus (Hesperia), 294, 299
(Pyrgus), 294
fulvus (Oxypalpus), 203, 206
fumida (Lymantria), 483
funeralis (Euproctis), 421
- (Nisoniades), 298
‘i (Thanaos), 298, 299
furva (Lelia), 444
», (Lymantria), 484
», (Ocneria), 444, 484
fusca (Acyphas), 478
:, (Anepa), 478
», (Anthora), 462
», (Dasychira), 462
,, (Heetomanes), 502
», (Hesperia), 293, 299, 300
fuscipenne (Leptosoma), 84
fuscivena (Stracena), 388
fuscomaculata (Porina), 502, 506, 508
fusiformis (Nioda), 464
gabunica (Dasychira), 469
gabunica (Euproctidion), 469
galactopis (Porthesia), 393
galbana (Deilemera), 80
galbanum (Leptosoma), 80
= (Nyctemera), 80
galena (Lemonias), 542
galene (Aterica), 189
5, (Papilio), 189
galenus (Celenorrhinus), 206
galinara (Lymantria), 490
gamma (Euproctis), 412
ganaha (Lymantria), 487
ganara (Lymantria), 493
Gastropacha, xxxv, 311-373
gaudens (Lopera), 409
gayi (Pieris), 290
Gazalina, 387
Gegenes, 143, 144
gemmata (Euproctis), 402
A (Lacipa), 402
gentia (Euproctis), 414
gentilis (Dasychira), 473
Genusa, 437, 438
Geodena, 497
Geometra, 433
georgiana (Dasychira), 473
germainii (Tetraphlebia), 279, 286, 298,
301
gerontes (Abisara), 194
» (Papilio), 194
gerra (Deilemera), 63, 85
gidica (Belenois), 145, 147, 150, 152
gigantea (Lelia), 441
glacialis (Colias), 242
glandulosa (Cheerotricha), 416
glaphyra (Anatole), 545
3H (Apodemia), 545
glaucinans (Homoptera), xxii
globifera (Cherotricha), 419
(Nygmia), 419
Gluphisia, 461
glycera (Teracolus), 142, 143, 144, 145,
146, 147, 148, 150, 152, 161
gnava (Dasychira), 477
Gnophodes, 184
godarti (Emesis), 536
Gogana, 396, 405, 416
gondona (Lymantria), 495
Gonometa, Xxill
gonophora (Dasychira), 470
- (Ilema), 470
gonostigma (Bombyx), 458
(Orgyia), 458
goodii (Dasychira), 467
», (CKeura), 467
gordoni (Psenderesia), 196, 296
Gorgopis, 502, 504, 505, 506, 507
Gorgyra, 203
( .cxlii )
gotzius (Byblia), 189
», (Papilio), 189
gracilis (Euproctis), 404
», (Lacipa), 404
gracillima (Leucoma), 382
grande (Nymphidium), 546, 550
grandidieri (Calliteara), 473
“a (Dasychira), 473
grandis (Lymantria), 489
(Numenoides), 481
29
Grapta, xxvi, xxvill, 557, 559, 562, 573,
574
grisea (Lymantria), 492
grossa (Dasychira), 490
erotei (Dasychira), 472
guenéi (Anthela), 446
(Newmania), 446
,, (Teara), 446
gundlachia (Precis), xii
‘ (Pyrisitia), 157, 162
guttata (Artaxa), 411
,, (Euproctis), 411
guttistriga (Euproctis), 422
guttulata (Euproctis), 419
cuttulosa (Deilemera), 61
(Nyctemera), 61
- (Pitasila), 61
gwelila (Dasychira), 469
Gynephora, 439, 455, 457
hadina (Porthetria), 483
hahni (Neosatyrus), 284, 285
hamadelpha (Mesarchea), 503
Hamadryas, 287
Hamanumida, 190
hamata (Anthela), 450
hampsoni (Euproctis), 423
Haplopseustes, 429
Harapa, 439, 442
harea (Deilemera), 81
, (Leptosoma), 81
Harma, 192, 193
harpalyce (Euphedra), 191
3 (Papilio), 191
harpax (Axiocerses), 200
», (Papilio), 200
hatita (Hypolycena), 198
heeate (Amauris), 184
,, (Danais), 184
hecla (Colias), ili, xxxvii, liii, 242
Hectomanes, 502, 505, 507
hectus (Hepialus), 501
helcita (Aletis), xxxix
helice (Aricoris), 548
Heliconius, iv, lv, 558, 565
Heliochroma, 290, 299
heliodora (Aricoris), 548
Hemaris, xxi
Hepialiscus, 507
9
99
Hepialus, 501, 503, 504, 507
hera (Lymantria), 466
Heracula, 431
herce (Nyctemera), 81
_herklotsii (Nyctemera), 75
Hesperia, Ivi, 189, 199, 201, 203, 204,
205, 298, 294, 295, 299, 300
hesperia (Geometra), 433
(Nyctemera), 53, 433
», (Otroeda), 53, 433
hesperistis (Apoprogones), xiv, 137,
138
hetiwerina (Symmachia), 538
Heterocampa, 461
heterogyna (Lelia), 444
Heteronygmia, 432, 433, 480
Heteropterus, xlii, xliii, xliv
hiarbas (Eurytela), 188
», (Papilio), 188
hilaris (Lymantria), 492
5, themis (Najas), 190
Himala, 386, 387
Hipparchia, 240, 275, 285
hipparia (Leucoma), 378
hippocoon (Papilio), x1
hippocrate (Aricoris), 547, 550
hippocrates (Cupido), 206
hippodice (Symmachia), 537, 549
hippothoe, var. eurybia (Chrysophanus),
XXXVli
var. stieberi (Chrysophanus),
XXXVil
holoxutha (Euproctis), 404
Homceomeria, 379
Homceonympha, 284
Homoptera, xxiii
Hondella, 439
horites (Nyctemera), 72
horrida (Dasychira), 470
horsfieldii (Arctia), 473
(Dasychira), 473
hortensia (Argynnis), 286
5 (Euptoieta), 286, 299
howra (Artaxa), 400
», (Kuproctis), 400
humilis (Neomcenas), 279, 298
», (Stygnus), 279
humuli (Hepialus), 501, 503
var. hethlandica (Hepialus),
>
99
Dh)
501
hyale (Colias), xliii
hyalinalis (Psammotis), xl viii
hyalinatus (Pielus), 502, 508
Hybernia, ix
Hybocampa, 249
Hylemera, 436
Hylephila, 294, 295, 299, 300
hymen (Dapidodigma), 199
( yexliv 5)
hymen (Papilio), 199 indeterminata (Casama), 461
Hypanartia, 187 (Ophiusa), xxiv
hyperantus (Aphantopus), xlii indica (Gluphisia), 461
Hyphilaria, 531, 549 », (Varmina), 461
Hypogymna, 482 inepta (Artaxa), 413
Hypoleucis, 204 », (Euproctis), 413
Hypolimnas, xxviii, xxix, 142, 188 infima (Dasychira), 477
Hypolycena, 197,198, 199 ;, (Somena), 477
hypophleus (Chrysophanus), [xii infuscata (Leptosoma), 70
hypoxantha (Heteronygmia), 433 a (Nyctemera), 70
(Numenes), 433 ino (Argynnis), iii
Hypsophila, 243 inornata (Anthela), 452
Hysibada, 386 > Warala), 452
iwris (Chamielimnas), 534 ;, (Neomeenas), 278, 298, 301
iasis (Mesene), 540, 550 insignis (Numenius), 432
ichorina (Pegella), 491 s, (Pterolocera), 453
icilia (Bombyx), 417 insulare (Leptosoma), 65
,, (Euproctis), 417 insularis (Deilemera), 65
Ieta, 460 integra (Anthela), 452
idmon (Lemonias), 542 », (Darala), 452
idonea (Euproctis), 401 »5 (Deilemera), 76, 85
ignita (Oxypalpus), 204 », (Leptosoma), 76
ignobilis (Mycalesis), 185 | ,, (Nyctemera), 76
Ilema, 462, 466, 469, 470 intensa (Artaxa), 412
ilicis (Thecla), xlv », (Euproctis), 412
ilita (Dasychira), 387 | intercisa (Deilemera), 62
ilithyia (Byblia), 143 Fe (Nyctemera), 62
illanta (Euproctis), 400 interjecta (Pantana), 438
illepida (Rilia), 429 interlectum (Nyctemera), 68
illustris (Deilemera), 77 intermixta (Gazalina), 387
Imaus, 496 intricata (Oncoptera), 500, 502, 508
immaculata (Cherotricha), 416 invaria (Dasychira), 465
3 (Euproctis), 399, 416 », (Repena), 465
impressa (Caragola), 376 invasa (Turriga), 464
.s (Leucoma), 376 iobrota (Artaxa), 395
as (Redoa), 376 ,, (Porthesia), 395
improba (Argynnis), ili, 242 ione (Teracolus), 157
impuncta (Euproctis), 401 lostola, vi
ae (Lacipa), 401 iphis (Ceenonympha), iil
inaria (Hypolimnas), xxix », (Papilio), 205
incerta (Enome), 418 »» (Rhopaloeampta), 205
5, (Lymantria), 481 irrorata (Daplasa), 430
;, (Pamphila), 205 », (Euproctis), 394
5, (Pardaleodes), 205 5» (Leucoma), 394
inclusa (Dasychira), 465 ,, (Porthesia), 394
incommoda (Artaxa), 411 ;, (Somena), 424
(Euproctis), 411 irroratum (Cricosoma), 540, 550
incompta (Euproctis), 420 Isapis, 534
incomptaria (Utidava), 429 isca (Liptena), 196
inconcisa (Artaxa), 401 ,, (Pseuderesia), 196
,, (Euproctis), 401 Isine, 497
inconspicua (Euproctis), 401 isis (Cupido), 201
inconstans (Deilemera), 57, 60 », (Papilio), 201
x (Nyctemera), 78 Ismene, 205
3 (Pitasila), 57 Ithomia, lvi
Incurvaria, 504 itonia (Ypthima), 206
indecora (Euproctis), 421 Ivela, 388
a (Teara), 421 Ixias, 156
( exly*)
Jana, xxili
Janassa, 462, 473
janetta (Euphiedra), 191
»- (Romalwosoma), 191
janirioides (Epinephele), 276,
Janiroides (Neosatyrus), 286
japonica (Lymantria), 483
jesous (Azanus), 148
jodutta (Acriea), 186
jodutta (Papilio), 186
johnstoni (Lomadonta), 480
Jolaus, 199, 206
jonasi (Aroa), 410
», (Kuproctis), 410
jonesi (Nyctemera), 433
», (Otroeda), 433
josiata (Artaxa), 396
», (Euproctis), 396
», (Orgyia), 396
-jJoviana (Chamelimnas), 534, 549
»» (Isapis), 534
junctifera (Charnidas), 454
Junonia, 187
justicie (Artaxa), 424
juvenis (Hondella), 444
», (Lelia), 444
», (Ptilomacra), 444
kala (Artaxa), 426
5, (Deilemera), 75
», (Euproctis), 426
,, (Leptosoma), 75
Kallima, xxviii, lii, 188, 573
Kanchia, 378, 385
kapaurensis (Deilemera), 73, 85
kargalika (Euproctis), 402
kausalia (Dasychira), 472
keiskamma (Teracolus), 157
Kettelia, 390
kinabalina (Deilemera), 81
3 (Nyctemera), 81
kinagananga (Deilemera), 76
klugii (Limnas), xxix, 142, 143, 144,
145, 146, 149, 150, 151, 152, 574
kondeka (Deilemera), 72, 77
kondekum (Leptosoma), 77
kiinowii (Pseudacriea), xxxix, xli
laba (Redoa), 379
labdaca (Libythea), 194
labyrinthica (Trictena), 502, 507
Lachana, 460
Lachnocnema, 206
Lacida, 429, 439, 443, 454, 460, 466
Lacipa, 395, 401, 402, 403, 404, 465
lactea (Lielioides), 440
», (Redoa), 381
lacticinia (Deilemera), 74
os (Leptosoma), 74
3 (Nyctemera), 53
298
PROC, ENT, SOC. LOND., Vv. 1903.
lacticinia (Phalena-Geometra), 74
ladakensis (Lachana), 460
Lelapia, 481
Lelia, 384, 391, 397, 439, 440, 441,
442, 443, 444, 449, 465, 470
Lielioides, 439, 440
leeta (Pseudomesa), 432
letitia (Euryphene), 192
Laganda, 498
lagus (Aricoris), 547
lampeto (Caria), 538
Lampides, 289, 299
lanaria (Kuproctis), 428
» (Terphothrix), 428
lanceolata (Rilia), 464.
laodice (Charaxes), 194
», (Papilio), 194
», (Precis), 187
laon (Jolaus), 199
larydas (Lyceenesthes), 200
», (Papilio), 200
Lasaia, 541, 550
lasea (Pirga), 435
», (Xenosoma), 435
Lasiommata, 274
lasthenes (Lemonias), 545
latemarginata (Deilemera), 68, 85
(Nyctemera), 68
lathonioides (Argynnis), 264, 286, 299
latifascia (Euproetis), 397
Fe (Leucoma), 397
3 (Nyctemera), 84
latifera (Darala), 450
latistriga (Deilemera), 78, 79
7 (Leptosoma), 78
_ (Nyctemera), 75, 78
var. fasciata (Nyctemera), 75
laufella (Hesperia), 205
»» (Pteroteinon), 205
lavia (Lelia), 443
lebona (Hypolycena), 199
Lecriolepis, 498
leda (Melanitis), 184
»» (Papilio), 184
leechi (Notolophus), 459
2” (Orgyia), 459
lefebvrei (Elina), ee role 298
45 (Satyrus), 27
leighi (Mania), xxv
», (Musgravia), xxii
leithiana (Artaxa), 399
Lemonias, 542, 543, 544, 545, 550
leo (Teracolus), 145, 146, 148
leonidas (Papilio), 203
Lepasta, 433
lepcha (Lymantria), 490
», _ (Porthetria), 490
Leptidia, xliii
( exlvi )
Leptoeneria, 445, 446
leptoneuroides (Cosmosatyrus), 274,
279, 280, 298, 301
50 yar, plumbeolus (Cos-
mosatyrus), 298
Leptosia, 201
Leptosoma, 55-84
Lerna, 497
lesbia (Colias), 291, 299
», (Papilio), 291
Leuceronia, 152
leucocyana (Lemonias), 542, 543
leucoglene (Faunula), 283, 298, 301
Leucoma, 376, 377, 378, 379, 380, 381,
382, 383, 384, 385, 388, 393, 394,
397, 415, 441
leucomelas (Euproctis), 397
leuconoe (Deilemera), 55, 65
(Leptosoma), 65
ns (Nyctemera), 65
leucopheearia (Hybernia), ix
leucospila (Cheerotricha), 422
(Cozola), 422
(Euproctis), 422
leucospilota (Deilemera), 60
(Nyctemera), 60
A (Pitasila), 58, 60
leucostigma (Leptosoma), 84
leucothea (Heliochroma), 290, 299
(Papilio), 290
leuctra (Deilemera), 62, 63, 85
libania (Gorgopis), 502, 505, 508
libyra (Aroa), 432
», (Numenes), 432
Libythea, 194
licinia (Eurygona), 531, 532, 549
lignea (Dasychira), 464
», (Nioda), 464
ligniperda (Cossus), x, 1x
lignivora (Charagia), 502, 504
lilacina (Leelia), 440
liliana (Pinacopteryx), 153
Limacodes, 246
limbalis (Urocoma), 478
limbata (Artaxa), 424
Limenitis, 556, 572, 573
Limnas, xi, xxii, xxix, 142, 148, 144,
145, 146, 149, 150, 151, 152, 565,
574
limonea (Anthela), 452
(Cherotricha), 413
(Darala), 452
», (Euproctis), 413
limonias (Satyrus), 276
limpida (Sapelia), 389
jineata (Dasychira), 477
5, (Lymantria), 477
lineosa (Etobema), 439
2?
2?
2)
99
Js)
|
lineosa (Pantana), 439
lingeus (Cupido), 201
», (Papilio), 201
linta (Artaxa), 426
,, (Euproctis), 426
Liparis, 385, 887, 390, 391, 394, 398,
404, 410, 412, 437, 479, 482, 483
Liphyra, x
Liptena, 196, 197, 206
litura (Charnidas), 444
»» (Lelia), 444
livia (Kuproctis), 414
livida (Dendorix), 198
I-nigra (Leucoma), 385
l-nigrum (Bombyx), 385
Locharna, 431
locuples (Cifuna), 479
lodra (Euproctis), 412
Lomadonta, 480
lombokiana (Deilemera), 71
Be (Nyctemera), 71
longipennis (Dasychira), 473
Lopera, 396, 401, 402, 409, 410, 441
Lophostethus, xxiv, xxvi
lorimeri (Morasa), 496
lorisona (Deudorix), 198
»» (Myrina), 198
losinga (Euphiedra), 192
», (Romaleosoma), 192
lowii (Kettelia), 390
| lucescens (Lymantria), 489
% (Porthetria), 489
lucifuga (Artaxa), 395, 408
,, (KEuproctis), 408
lucilla (Neptis), iii
lucretia (Papilio), 189
oy (Pseudacriea), 189
luctuosa (Deilemera), 80
luctuosum (Leptosoma), 80
luctuosus (Satyrus), 276
luctus (Riodina), 535
ludekingii (Deilemera), 67
(Leptosoma), 67
» _ (Orgyia), 460
Ludia, xxiii
lugardi (Lymantria), 493
lugubris (Pedaliodes), 276
29
/lunata (Bombyx), 491
(Euproctis), 399
», (Lymantria), 491
lunifera (Zarfa), 386
lunulata (Dasychira), 471
Iupulinus (Hepialus), 501, 504
lutea (Bombyx), 395
», (Porthesia), 395
luteiceps (Pantana), 438
luteifascia (Artaxa), 423
(Dactylorhyncha), 444
99
2?
luteifascia (Euproctis), 423
luteipes (Homceomeria), 379
5 (Leucoma), 379
», (Stilpnotia), 379
lutescens (Euproctis), 399
Lycena, liii, 201, 242, 288, 289
Lyczenesthes, 148, 200, 206
lycoa (Acreea), 186
Lycorea, iv, lv, 558, 565
Lymantria, 432, 443, 466,
481-496
lyrnessa (Lyczena), 288
lysimon (Cupido), 206
lysippus (Riodina), 535
lysistratus (Riodina), 535
mabillei (Nyectemera), 66
me kieana (Nyctemera), 84
macklotti (Leptosoma), 84
Maecrauzata, 384
Macroglossa, xxvi
maculata (Deilemera), 62
5 (Fodinoidea), 431
a (Nyctemera), 62
maculosa (Lymantria), 489
55 (Nyctemera), 64
maculosum (Leptosoma), 58
madagascariensis (Junonia), 187
(Precis), 187
madana (Euproctis), 417
mieon (Lemonias), 542, 550
meonoides (Lemonias), 542, 550
mera (Pararge), lili
mieris (Lasaia), 541
magna (Euproctis), 423
» (Pirga), 436
», (Somena), 423
magnalia (Dasychira), 469
magnifica (Anthela), 448
Mahoba, 433
maia (Aricoris), 547, 550
major (Aroa), 453
Malachitis, 481
Malacosoma, viii
malathana (Cupido), 201
4% (Lyceena), 201
maligna (Dasychira), 466
33 (Parorgyia), 466
malis (Cremna), 530, 549
malve (Syrichthus), li
mancinus (Erebia), 239
Mania, xxv:
manicata (Lymantria), 489
manifesta (Nyctemera), 434
manto (Erebia), xlv
Marane, 497
Marbla, 436, 437
Marcipa, 497
mardania (Euryphene), 192
exlvii_)
477, 479,
mardania (Papilio), 192
Mardara, 433, 463, 468, 476
margaritacea (Leucoma), 377
marginalis (Euproctis), 392
a (Leucoma), 380
ap (Redoa), 380
- (Trichetra), 479
Be (Urocoma), 479
marginata (Aslauga), 197
es (Cheerotricha), 418
Pr (Cobanilla), 392
- (Euproctis), 418
3 (Liptena), 197
Ke (Lymantria), 487
maria (Caragola), 377
», (Redoa), 377
marianne (Ixias), 156
marsyas (Caria), 538, 549
martia (Lemonias), 543, 550
maruta (Dasychira), 471
mascarena (Dasychira), 476
Matarbela, xxill, xxvi
mathias (Chapra), 146, 204
- (Hesperia), 204
mathura (Lymantria), 488, 489
maxima (Aroa), 453
maza (Euproctis), 407
Mechanitis, iv, liv, lv
mediata (Stauropus), xxiv
medon (Hupheedra), 191
», (Papilio), 191
medusa (Leptosia), 201
», (Papilio), 201
melaleuca (Artaxa), 415
melaneura (Deilemera), 65
_ (Leptosoma), 65
x var. melas (Nyctemera), 83
melaneus (Caduga), 565
Melanitis, lii, 184
melanochlora (Malachitis), 481
ss (Mesenopsis), 533
melanosoma (Leucoma), 393
a5 (Porthesia), 393
melaxantha (Aroa), 456
(Orgyia), 456
Melia, 461, 467
melia (Lemonias), 544
melicerta (Neptis), 189
55 (Papilio), 189
Melineea, iv, liv, lv, 558, 565, 571
Melita, ili, xxxvii, xlix—lii, liv
menander (Cithierias), 565
mendosa (Dasychira), 464
Fe (Olene), 464
menes (Nyctemera), 84
menestheus (Papilio), 203
mentiens (Crorema), 391
mercedis (Pontia), 292
(Vexlyin se)
meris (Lasaia), 541
merita (Lasaia), 541,
Merope, xxv
merope (Papilio), xxxix, xl, xli
Mesarchiea, 503
Mesene, 540, 550
Mesenopsis, 533, 549
550
mesentina (Belenois), 143, 144, 145,
147, 148, 151, 155
mesolychna (Nyctemera), 83
Mesosemia, 529, 530, 549
Mesoxantha, 188
metallica (Neocastnia),
Metanastria, xxii, xxiii
metarhoda (Lymantria), 489
Methona, 573
micacea (Leucoma), 383
», (Redoa), 383
micans (Lymantria), 486
», (Parnara), 204
micilia (Agyrta), v, vi
micra (Calydna), 541
microcale (Teracolus), 147
microdice (Pieris), 292
55 (Tatochila), 299, 300
Microgymna, 396
Micropterogyna, 458
micyelus (Cupido), 201
es (Papilio), 201
milhauseri (Hybocampa), 249
milonia (Precis), 187
milyas (Monotrichtis), 185
», (Mycalesis), 185
Mimacriea, 195, 196
mimosie (Argema), xxii
minimus (Neosatyrus), 283
minuscula (Colias), 291
minutissima (Euproctis), 425
(Leucoma), 380
mirabilis (Euproctis), 415
36 (Pirga), 435
mirma (Aroa), 455
mirza (Cupido), 201
misana (Dasychira), 467
miserata (Dasychira), 466
. (Ilema), 466
misippus (Basilarchia), 572
(Hypolimnas), xxviii,
142, 188
(Limenitis), 572
= (Papilio), 188
mixta (Leucoma), 393
» (Porthesia), 393
mneme (Melina), iv, liv, lv
mocquerysii (Gorgyra), 203
modesta (Anatole), 545
(Argynnis), 287, 299
(Artaxa), 399
39
2?
29
29
KK,
| modesta (Dasyehira), 472
(Morasa), 496
3 (Polymona), 496
meerens (Calliteara), 474
(Dasychira), 474, 482
», (Lymantria), 482
moeris (Anza), XXvVi
moesta (Lymantria), 484
moina (Colias), 243
molione (Monethe), 539, 550
monacha (Bombyx), 486
*5 (Lymantria), 486
monachus (Epinephele), 276, 298
(Satyrus), 276
ue (Stibomorpha), 276
Monethe, 539, 550
monosticta Gee 410
(Lopera), 410
Monotrichtis, 185, 206
monticolens (Cosmosatyrus), 281, 298,
301
A (Satyrus), 281
montis (Artaxa), 400
», (Euproctis), 400
montroilii (Elina), 274
(Lasiommata), 274
(Satyrus), 274
29
9?
99
99
99
| Monura, 194
moolaica (Pitasila), 57
moorei (Euproctis), 424
», (Leucoma), 380
morania (Papilio), xxi
Morasa, 496
mormoides (Ophiusa), xxiv
morpheus (Heteropterus), xlii,
xliv
mosera (Lymantria), 494
mozambica (Aphneus), 200
(Spindasis), 200
xiii,
| miilleri ( (Deilemera), 63
(Euproctis), 416
», (Leptosoma), 63
multiplaga (Apodemia), 546, 550
multipunctata (Pentila), 195, 206
munda (Euproctis), 412
,, (Lymantria), 496
mundipicta (Deilemera), 55, 74
(Leptosoma), 75, 76
55 (Nyctemera), 75
mundus (Imaus), 496
Munichryia, 478
municipalis (Dasychira), 465
PA (Leelia), 465
muscalella (Incurvaria), 504
Museravia, xxii
mutabilis (Deilemera), 62
2?
29
3 (Nyctemera), 62
Mycalesis, 184, 185
(
mycone (Nymphidium), 546
Mylothris, 201
Myrina, 198, 206
mys (Eurygona), 531, 532
Nagunda, 482, 486
Najas, 190
nape (Pieris), XXXVii
Napeogenes, lvi
narindra (Lymantria), 492
Naroma, 386
nastes (Colias), 243
natalensis (Precis), 160
natalica (Acreea), Xxil
Nataxa, 496, 497
Naxa, 386
neanthes (Charaxes), xxi
nebulosa (Monotrichtis), 185
35 (Mycalesis), 185
neemias (Emesis), 587, 549
negrita (Euproctis), 396
nemetes (Neptis), 189
Neocastnia, 139
Neomeenas, 274, 277, 278, 279
301
neomyrioides (Elina), 274, 275, 298,
301
3 (Satyrus), 274
Neosatyrus, 283, 284, 285, 298, 301
Nephele, xxii
Neptidopsis, 188
Neptis, i, 150, 151, 189
nereis (Pierella), 565
neriene (Zeritis), 206
nerina (Nyctemera), 53
», (Otroeda), 53
nervosa (Oligoclona), 387
netopha (Baoris), 205
», (Hesperia), 205
Neurophana, 457
neustria (Malacosoma), viil
x eastrensis (Malacosoma), |
vill
Newmania, 376, 445, 446,
niavius (Amauris), xl, 184
5, (Papilio), 184
nicevillei (Neocastnia), 139
nicias (Hyphilaria), 531
nicippe (Xanthidia), 156
nicothoé (Anthela), 449
5 (Bombyx), 449
nigra (Dasychira), 469
», (Lymantria), 487
nigribasalis (Euproctis), 396
nigricilia (Redoa), 378
nigrifinis (Porthesia), 393
nigriplapa (Orgyia), 458
nigritula (Dasychira), 468
bP)
exis)
nigrocrocea (Orgyia), 458
nigrolimbata (Pantana), 438
nigroscripta (Heterocampa), 461
nigrovena (Deilemera), 74, 85
nigrovenosa (Deilemera), 64
nilgirica (Dasychira), 473
Nioda, 461, 464
| niphonis (Chierotricha), 418
», (Euproctis), 418
nireus (Papilio), xxi, xxv, 202
nisa (Deilemera), 77, 85
Nisoniades, 298
‘nitida (Leucoma), 379
by
niveosparsa (Dasychira), 475
nobilior (Ploetzia), 205
nobilis (Cheerotricha), 396
(Euproctis), 402
(Lopera), 402
(Panthea), 402
2)
| nohara (Acrea), 155
Nola, 394
'nomenia (Deudorix), 206
298, |
nomia (Lemonias), 544
/nomion (Deudorix), 197
_numata (Heliconius),
/oaxes (Pedaliodes),
nigrocilia (Euphedra), 190, 206
norna ((Eneis), Xxxvil
nostrodamus (Gegenes), 143, 144
notata (Anaxila), 429
(Leelapia), 481
», var. valdiviana (Hesperi
Notohyba, 462, 463, 466, 470
Notolophus, 407, 458, 459
nouna (Teracolus), 162
noviespunctatum (Nyctemera), 62
nubecula (Leptosoma), 84
nubifuga (Dasychira), 466
», (Notohyba), 466
nubila (Birnara), 439
nucula (Orgyia), 458, 459
nuda (Bombyx), 430 ©
9
a), 294
Jy 2?
iv
Numenes, 432, 433
numenes (Charaxes), 194
Numenoides, 481
nurma (Euproctis), 420
Nychthemera, 66
Nyctemera, 53-84, 433, 434
nycteropus (Neosatyrus), 285, 298, 301
nycteus (Nymphidium), 546
Nygmia, 405, 419
| Nymphalis, 193
nymphea (Phulia), 292
Nymphidiuin, 546, 550
nymphula (Phulia), 264, 292 299
a (Pieris), 292
nyses (Cypra), 437
(Marbla), 437
a7
af
obfuscata (Enome), te
obfuscata (Lymantria), 482
obliqua (Cispia), 391
obseura (Anthela), 453
(Artaxa), 424
(Euproctis), 424
», (Trichiura), 453
obsoleta (Artaxa), 400
(Bombyx), 440
(Euproctis), 397
(Lelia), 440
», (Lymantria), 484, 485
obtusa (Deilemera), 78
»». (Nyctemera), 78
occidentis (Nyctemera), 434
AS (Otroeda), 53, 434
ocellata (Anthela), 447
ee (Warala) 447
ocellifera (Dasychira), 466
(Notopriota), 466
», (Oecura), 466
ochracea (Aroa), 454
», (Charnidas), 454
ochraceata (Aroa), 457
ochrea (Euproctis), 405
(Gogana), 405
5 (Nygmia), 405
ochreipennis (Teracolus), 163
ochreivittatus (Neosatyrus), 284
ochripes (Caragola), 378
(Caviria), 378
», (Stilpnotia), 376, 378
ochrocephala (Dicreagra), 496
Ochrogaster, 498
Ocinara, 385
Ocneria, 444, 484
octavia (Papilio), 187
»» (Precis), 187
ocularis (Orgyia), 460
Odagra, 439, 440
odana (Deudorix), 197
odenestaria (Colussa), 450
Odontopera, xxxv, 311-370
(Ecura, 462, 466, 467
cedipus (Cceenonympha), xlii, xl, xliv
(Kneis, xxxvii, 240, 244, 281
Cinosandra, 497
Ogoa, 391
ogovensis (Redoa), 379
ogrugana (Baoris), 204, 206
oileus (Lasaia), 541, 550
Olapa, 390, 391
olearia (Dasychira), 462
», (Olene), 462
Olene, 461, 462, 464
Oligoclona, 387
olivata (Euproctis), 427
Ommatoptera, 447, 449
omphale (Teracolus), 157
99
9
2)
29
9
39
29
3)
cl
Oncoptera, 500, 502, 503, 507
onetha (Deilemera), 80
,, (Leptosoma), 80
opalina (Cimola), 435
ophione (Neptidopsis), 188
», (Papilio), 188
| Ophiusa, xxiv
ophiusa (Hesperia), 204
», (Hypoleucis), 204
optata (Deilemera), 82, 85
orbitulus (Papilio), 242
" var. franklinii (Lycena), 242
orbona (Mesosemia), 530, 549
orcas (Aphneeus), 199
», (Papilio), 199
orciferaria (Aspilates), 243
oreosaura (Adlullia), 418
s, (Euproctis), 418
orestes (Cherotricha), 417
», (Euproctis), 417
orestia (Acriea), 186
Orgyla, xxxi, xxxil, 396, 407, 437, 438,
454, 455, 456, 458, 459, 460, 488
orientalis (Tascina), 139
orimba (Dasychira), 464
,» (Olene), 464
orise (Dismorphia), 558, 573
ormea (Euproctis), 426
oroya (Deilemera), 59, 85
orsedice (Hyphilaria), 531, 549
Orvasea, 396, 424
osiris (Cupido), 201
», (Lyceena), 201
Osmodes, 204
ostra (Anthela), 447
5, (Euproctis), 402
osuna (Euproctis), 419
otlauga (Liptena), 197
Otroeda, 58, 54, 433, 434
ovada (Deilemera), 82
Oxylides, 198
Oxypalpus, 203, 204, 206
Pachycispia, 479
pagenstecheri (Nyctemera), 83
Paleeomicra, 503
pales (Argynnis), iii, 241
», (Papilio), 241
», (Satyrus), 277
Palla, 194
pallens (Leptosoma), 84
pallida (Charnidas), 444
(Cropera), 410
(Dactylorhyncha), 444
(Euproctis), 410
», (Lelia), 440
pallipes (Euproctis), 419
Palpiphorus, 507
Pamphila, 205
99
”)
(eli es)
panabra (Porthesia), 392
paniscoides (Butleria), 297
pansa (Chameelimnas), 534, 549
Pantana, 437, 438, 439
Panthea, 402
Papilio, xxi, xxv, xxxix, xl, ]xiii, lxiv,
150, 184-205, 239, 241, 242, 286, |
290, 291, 293, 299, 555, 561, 563
paradoxa (Chionophasma), 392
a (Porthesia), 392
Pararge, xliii, ii
Pardaleodes, 205
parmeno (Gnophodes), 184
Parnara, 204, 206
Parnassius, iii, iv
Parorgyia, 466
parrhasia (Acreea), 186
35 (Papilio), 186
parthenie (Melita), xlix, ], li
Re var. varia (Melita), xlix, ]
partita (Numenes), 432
parva (Anthela), 445
», (Darala), 445
», (Telipna), 194
pasinuntia (Lycorea), iv, lv
pastor (Calliteara), 476
,, (Dasychira), 476
patrana (Numenes), 432
patula (Thelde), 465
pauli (Pentila), 195
pauperata (Euproctis), 400
peartize (Cneis), 240
pecla (Euproctis), 414
peculiaris (Dasychira), 463
se (Mardara), 463
Pedaliodes, 275, 276, 298
pedaria (Phigalia), ix
pedias (Siseme), 534, 549
Pegella, 482, 491, 493
pelarga (Papilio), 187
ee (recs) sal(Sif
pelidne (Colias), 243
pellex (Atasca), 56
», (Deilemera), 56
,, (Leptosoma), 56
pellucida (Leucoma), 381
pelona (Artaxa), 411
», (Euproctis), 411
peneleos (Acrea), 186
Penora, 381
pentapolis (Acraea), 186
Pentila, 195, 196, 206
perenna (Acrea), lxiv
Perina, 430
Perna, 496
perplexa (Euproctis), 422
perspicua (Deilemera), 66
59 (Nyctemera), 66
petiverana (Danais), 184
petreea (Acriea), xxii, 155
petulca (Phalena), 80
| phace (Mesosemia), 529, 549
| phzea (Euproctis), 429
pheedon (Aricoris), 547
Phiegorista, 479
Phalena, 64, 68, 80, 390, 392
», (Geometra), 61, 74
», (Noctua), 56
phalantha (Atel!a), 142, 187
a (Papilio), 187
| phalanthus (Bicyelus), 184
phaola (Appias), 202
»> (Pieris); 202
pharsalus (Acriea), Ixiv, 186
pharte (Erebia), xlv
phasiana (Dasychira), 466
», (Parorgyia), 466
phegea (Elymnias), xli, 184
5, (Papilio), 184
pheranthes (Napeogenes), lvi
Phiala, 497
| Phigalia, ix
philippii (Butleria), 296, 299
+5 (Cyelopides), 296
_philippus (Hesperia), 199
», (Hypolycena), 199
Phineca, 462, 463
phisadia (Teracolus), xii, 145,146, 157,
163
phleas (Chrysophanus), xliii, bxii
phlegyas (Teracolus), 146, 157
pheebe (Melitiea), iii, li
pheenicias (Anthela), 445
Phragmatobia, 442
phranza (Euryphene), 206
phryne (Triphysa), iit
Phulia, 264, 292, 299
phyleus (Hylephila), 295
picata (Deilemera), 81
picatus (Secusio), 81
picta (Euproctis), 404
», (Liparis), 404
,, (Pachycispia), 479
Pida, 427, 431
Pielus, 502, 507
Pierella, 565
Pieris, xxxvil, lili, 202, 290, 292, 293
Pinacopteryx, xi, 142, 152, 153, 154
pinguis (Anthela), 453
», (Darala), 453
pirene (Lemonias), 544, 550
Pirga, 435, 436
Pitasila, 54, 55, 56, 57, 58, 59, 60, 61
plagiaria (Aricoris), 546, 550
plagiata (Anaxila), 471
», (Cispia) 418
( eli
plagiata (Dasychira), 471
- (Euproctis), 418
plagiatum (Leptosoma), 68
plagidotata (Cyclidia), 433
7 (Mardara), 433
plagifera (Deilemera), 67
», (Nyctemera), 67
», (Tripheromera), 67
plana (Anthela), 450
», (Aroa), 454
., (Cherotricha), 416
(Darala), 450
(Euproctis), 416
(Orgyia), 454
Planema, D@;o:ab.gy ond hy ove BIE
planemoides (Papilio), xli
Plastingia, 205
Platylesches, 205
plautilla (Euryphene), 192
», (Euryphura), 192
pleione (Teracolus), 146
plexippus (Anosia), 562, 572
plinius (Cupido), 201
», (Hesperia), 201
ploetzi (Hesperia), 203
Ploetzia, 205
plumbalis (Lymantria), 492
plumbea (Scolitantides), 288
plumbeola (Tetraphlebia}, 279, 280
- var. duseni (Erebia), 280
pluto (Gynzphora), 439
53 (Pantana), 439
poggei (Planema), xxxix, xli, xhi
polaris (Argynnis), 241
policenes (Papilio), xxi, 203
polydamas (Papilio), Ixili, 555
Polygonia xxvi, XXVii, XXvViii
polymnia (Mechanitis), liv, lv
Polymona, 496
Polyommatus, 144, 145, 150, 289
polyspilus (Butleria), 297, 299, 300
ae (Carterocephalus), 297
pomona (Catopsilia), 156
Pontia, 292
popiya (Deilemera), 69, 85
populi (Smerinthus), xiv
Porina, 502, 506, 507
Porthesia, 392, 393, 394, 395, 416, 418
Porthetria, 482, 483, 489, 490
postfusca (Dasychira), 472
postica (Anthela), 449
(Darala), 449
(Euproctis), 397
,, (Gonometa), xxili
,, (Lacida), 460
(Orgyia), 460
s, (Thiacidas), 461
postincisa (Euproctis), 402
+B]
186, 187
be)
”)
)
postnigra (Euproctis), 421
potentaria (Ennomos), 451
preecurrens (Adlullia), 419
5 (Euproctis), 419
pramesta (Lymantria), 492
prasina (Calliteara), 476
;,» (Dasychira), 476
Precis) xX, Kil, SGKViily Kx OCD aTe
lii, 142, 143, 145, 151, 157, 158, 159,
160, 187, 188, 206, 556, 561, 562,
563
preussi (Abynotha), 479
(Liparis), 479
(Lymantria), 479
», (Phegorista), 479
prima (Darala), 420
primula (Leucoma), 383
princeps (Artaxa), 413
- (Lycenesthes), 206
principalis (Anteros), 536
prisca (Orgyia), 459 ~
Procodeca 439, 440, 441, 442, 444
producta (Euproctis), 394
», (Porthesia), 394
progne (Symmachia), 537, 549
prolata (Lelia), 440
prolixa (Lelia), 391
promaucana (Satyrus), 286
promelena (Stracena), 389
< (Sulychra), 389
propria (Deilemera), 71
proprium (Leptosoma), 71
», (Nyctemera), 71
Proterpia, xl
preterpia (Pyrisitia), 157, 162
protoclea (Charaxes), 193
>
9
| protomedia (Teracolus), 145, 147, 152
pruinosa (Arctornis), 379
(Leucoma), 379
pryeri (Dasychira), 472
Psalis, 461, 462
Psammotis, xlviii
pseudabietis (Calliteara), 472
% (Dasychira), 472
Pseudacreea, xxxix, xli, lxiv,
Pseudarbessa, vi
pseudegina (Acrea), 186
Pseuderesia 196, 206
Pseudomesa, 432
Pseudonotodonta, 462, 477
Psilura, 482
189
| psittacus (Caria), 538
| psyttalea (Amauris), 184
_ Pterolocera, 453
Pteroteinon, 205
Ptilomacra, 444
pubescens (Euproctis), 404
pudica (Anthela), 445
(
pudica (Darala), 445
5, (Dasychira), 472
puellaris (Teracolus), xii, 157, 163
puelme (Argopteron), 295, 299, 300
», _ (Cyclopides), 295
pulchella (Deiopeia), 154
3 (Mesenopsis), 533, 549
pulver ea (Artaxa), 406
», (Euproctis), 406
pulverea (Lacipa), 404
- (Porthesia), 394
pumila (Calliteara), 474
», (Dasychira), 474
punctatus (Cupido), 201
puncticilia (Caltura), 386
An (Cispia), 386
vie (Naxa), 386
punctifascia (Cispia), 430
punctifera (Aroa), 409
A (Dasychira), 463
- (Erastria), 463
pe (Euproctis), 409
punctulata (Lelia), 441
% (Lopera), 441
pura (Euproctis), 405
,, (Perina), 430
pusilla (Artaxa), 407
», (Homceonympha), 284, 285
,, (Lymantria), 487
5, (Teia), 458
pustulifera (Trisula), 477
pygmea (Aroa), 407
s (Euproctis:, 406, 407
pylades (Papilio), 150, 203
Pyrameis, 187, 287, 299
Pyramocera, 496
pyrene (Ixias), 156
Pyrgus, 294
Pyrisitia, 157, 162
pyrisitia (Proterpia), xii
pyrrhias (Acnissa), 429
pyrrhochroma (Aroa), 456
quadrangularis (Cherotricha), 418
quadriguttata (Deilemera), 71
quadriguttatum (Leptosoma), 71
(Nyctemera), 71
cliii )
}
|
quadrimaculata (Thecla), 289, 290, 299
quadriplaga (Atasca), 57
i (Darala), 451
ms (Deilemera), 57
quadriplagiata (Pseudomesa), 432
quadripunctata (Euproctis), 403
on (Lacipa), 403
quaternarium (Nyctemera), 83
quercifolia (Gastropacha), xxxv,
373
quinquepunctata (Lacipa), 403
raddei (Porthesia), 418
311-
radiata (Deilemera), 70
,, (Leptosoma), 70
», (Nyctemera), 70
», (Pentila), 195, 206
_rafflesia (Euschemon), xiv, 138, 139
Rajacoa, 434, 435
_ramsayi (Charagia), 502
| rapee (Precis), liii
,, var. bryonie (Pieris), liii
| Raphipeza, 497
rasana (Nyctemera), 84
rauana (Piecis), xlii
recraba (Euproctis), 428
| recurvata (Euproctis), 406
Redoa, 376,
383, 386
377, 378, 379,
/reducta (Anthela), 446
” (Darala), 446
regina (Teracolus), 157
_regularis (Deilemera), 81
,, (Leptosoma), 81
,, (Nyctemera), 81
| reichenowi (Pardaleodes), 205
(Plastingia), 205
remota (Dasychira), 477
rendalli (Dasychira), 465
renifera (Euproctis), 425
renominata (Euproctis), 397
Repena, 439, 443, 465
repleta (Anthela), 449
,, (Darala), 449
restricta (Deilemera), 66
restrictum (Leptosoma), 66
9 (Nyctemera), 66
reedi (Neosatyrus), 285, 298
,, (Stibomorpha), 275, 280
;, var. fuscescens (Neosatyrus), 285
Rhanidophora, 498
rhea (Pierella), 565
rhesa (Lemonias), 544
rhoda (Artaxa), 400
rhodapicata (Heteronygmia), 480
rhodina (Lymantria), 483
rhodogyne (Eurygona), 532, 549
rhodope (Appias), 202
;, (Papilio), 202
rhodopepla (Sarothropyga), 496
Rhopalocampta, 205
Ricine, 439, 441
ridleyanus (Papilio), lxiv, 203
Rigema, 462
Rilia, 429, 461, 464
rinaria (Caragola), 377
», (Caviria), 377
;, (Redoa), 376, 377
Riodina, 535, 549
risoria (Aroa), 457
robusta (Acyphas), 463
380, 381,
robusta (Dasychira), 463
rogersi (Acreea), Ixiv
Romaleeosoma, 190, 191, 192
rorus (Teracolus), 163
rosea (Lymantria), 489, 493
rossi (Colias), 243
» (Erebia), 240
», (Hipparchia), 240
rotundata (Dasychira), 474
3 (Lacida), 443
be (Teara), 474
rubescens (Anthela), 445
55 (Darala), 445
rubi (Callophrys), li
rubicunda (Anthela), 445
as (Darala), 445
rubida (Cyenia), 443
», (Nataxa), 497
rubripennis (Leelioides), 440
ruficeps (Mardara), 433
rufifascia (Anthela), 449
a (Darala), 449
rufifemur (Polymona), 496
rufimarginata (Leucoma), 383
rumia (Kallima), 188
ruspina (Eupheedra), 190
ss (Romaleosoma), 190
ruspine (Papilio), xxxix
rutilans (Colias), 291
safitza (Monotrichtis), 185
», (Mycalesis), 185
sagaris (Mesene), 540
sagrara (Aroa), 455
sagroides (Euproctis), 425
an (Somena), 425
Salamis, 188
sandace (Monotrichtis), 185
», (Mycalesis), 185
sangaica (Leela), 441
Sapelia, 389
Sarangesa, 203
Sarothropyga, 496
Sarsina, 497
sastra (Artaxa), 411
5, (Euproctis), 411
saturnioides (Lelia), 384
of (Leucoma), 384
Satyrus, xxxvii, xlii, xlii, xliv,
275, 276, 277, 279, 280, 281, 286
saucia (Agrotis), Ixvii
sawanta (Dasychira), 464
saxeus (Teracolus), 142, 161
Seada, lv
Scelothrix, 294
scintillans (Euproctis), 424
56 (Somena), 424
Scolitantides, 264, 288, 289, 299
scotochyta (Euproctis), 407
(
cliv )
-sexmaculatum (Leptosoma),
secundaria (Deilemera), 68
secundarium (Leptosoma), 68
securis (Dasychira), 462
», (Psalis), 462
seis (Acreea), 185
selecta (Deilemera), 60
3 (Nyctemera), 60
selene (Argynnis), iii, xxxvil
Selenia, 159
semicincta (Alope), 486
(Lymantria), 486
(Nagunda), 486
semidea (Hipparchia), 240
(CEneis), 240, 244
9)
9
semihyalina (Dendrophleps), 376, 378
semilucida (Pantana), 439
semire (Papilio), 189
», (Pseudacreea), 189
semisignata (Cispia), 399
op (Euproctis), 399
semperi (Deilemera), 57°
senegalensis (Terias), 150, 152
senica (Orgyia), 459
senicula (Munychryia), 478
separata (Nyctemera), 56
serica (Aricoris), 546
sericea (Caragola), 377
oa (Cawaitia). 304
»» (Redoa), 379
», (Stilpnotia), 377
sericina (Esthemopsis), v
servilia (Neomcenas), 278,
servilis (Artaxa), 420
5, (Euproctis), 420
298
| sesamus (Precis), xxxii, xxxiil, 160
setinoides (Lielia), 441
severina (Belenois), 147, 148, 152, 154,
155
sexmacula (Euproctis), 415
sexmaculata (Deilemera), 72
Land
76
sexpunctata (Lacipa), 403
_sibylla (Scolitantides), 289
sienna (Aroa), 454
_siga (Argynnis), 286
-signata (Aroa), 457
274,
| siletti (Numenes),
», (Deilemera), 56
;, (Euproctis), 398
» (Hesperia), 294
(Liparis), 398
», (Nyctemera), 56
(Porina), 502
signifera (Naroma), 386
silea (Lymantria), 493
silenus (Myrina), 198
(Papilio), 198
432
silhetica (Leucoma), 381
99
(
silhetica (Penora), 381
simana (Pinacopteryx), 154
simia (Precis), xxXiv
similis (Artaxa), 424
», (Chamelimnas), 534
», (Citrinophila), 196
», (Euproctis), 424
», (Lymantria), 487, 491
» (Phalena), 392
simplex (Anthela), 450
», (Argyrophenga), 279, 284
a (Atroa)s) 455)
», (Atasca), 57
», (Darala), 450
», (Neomeenas), 274
», (Neosatyrus), 284, 298
», (Nyctemera), 57
» (Ogoa), 391
», (Orgyia), 455
(Pantana), 438
simplicia (Liptena), 196
= (Ypthima), 185
simulans (Artaxa), 396
a (Hectomanes), 502
simulatrix (Deilemera), 74
a (Leptosoma), 74
As (Nyctemera), 74
sinapis (Leptidia), xliii
sinensis (Leelia), 441
», (Redoa), 380
sinica (Lymantria), 483
», (Pantana), 438
sirene (Euptera), 193
Siseme, 534, 549
Sitina, 498
Sitvia, 388, 390
smaragdina (Caria), 538, 549
Smerinthus, xiv
smilax (Ludia), xxiii
Smyriodes, 498
snelleni (Arctornis), 377
5 (Porthesia), 416
sobrina (Lymantria), 486
socrus (Aroa), 457
5, (Gynephora), 457
solitaria (Dasychira), 471
Soloé, 437, 497
Somena, 396, 423, 424, 425, 476, 477
sontica (Deilemera), 79
sonticum (Leptosoma), 79
a (Nyctemera), 79
sophia (Papilio), 187
», | (Precis), 187
sophus (Euryphene), 192
», (Papilio), 192
sordida (Stilpnotia), 437
sotoi (Butleria), 296, 299
, (Cyclopides), 296
clv_ )
speciosa (Scolitantides), 288
specularis (Deilemera), 61
0 (Nyctemera), 61
(Pitasila), 61
spica (Mylothris), 201
Spindasis, 200
spini (Thecla), xlv
squamiplaga (Artaxa), 410
squamosa (Cheerotricha), 418
3 (Euproctis), 409
(Lopera), 409
Stalachtis, iv
statilinus (Satyrus), xliii
staudingeri (Chierotricha), 418
- (Euproctis), 418
rH (Fodinoidea), 431
a (Lemonias), 543, 544
Stauropus, xXiv
stellata (Euproctis), 410
stelligera (Faunula), * 282, 298, 301
Steropes, 296, 297
Sthenopis, 501
Stibomorpha, 275, 276, 278, 280
stigmatifera (Por thesia), 395
Stilpnotia, 377, 378, 379, 4380, 437
stirasta (Adlullia), 417
5, (Euproctis), 417
Stracena, 388, 389
straminea (Euproctis), 413
striata (Dasychira), 470
(Notohyba), 470
», (Pseudacrea), 189
strigata (Dasychira), 475
strigifimbria (Antipha), 429
si (Euproctis), 429
strigipennis (Locharna), 431
- (Pida), 431
Strymon, 290, 299
stygiana (Anthela), 448
», (Darala), 448
stygne (Erebia), xlv, xlvi
5, var. bejarensis (Hrebia),
xlvi
Stygnus, 279
subdita (Euproctis), 594
subfaleata (Anthela), 450
(Darala), 450
subfascia (Orgyia), 438
3s (Pantana), 458
subfasciata (Artaxa), 397
(Euproctis), 397
subflava (Anaxila), 475
», (Aroa), 415
», (Dasychira), 475
», (Euproctis), 415
?
xlv,
» var. piperita (Leucoma), 415
subfuscula (Artaxa), 409
os (Euproctis), 409
(
subinanis (Topomesa), 392
sublutescens (Dasychira), 474
submacula (Liptena), 196, 206
submarginata (Leucoma), 378, 383
FS (Redoa), 378
subnigra (Euproctis), 394
subnobilis (Artaxa), 396
ce. (Euproctis), 396
<5 (Porthesia), 396
subnotata (Aroa), 454
3 (Lacida), 454
as (Orvasea), 424
subornata (Myrina), 198, 206
subpurpurella (Eriocrania), 503, 508
subrana (Artaxa), 427
», (Euproctis), 427
subrosea (Anthora), 439
», (Lelia), 439
,, (Lymantria), 489
subrufa (Leelia), 439, 441
substrigosa (Aroa), 456
subtincta (Stilpnotia), 430
subvelata (Deilemera), 70
“5 (Nyctemera), 70
subyelatum (Leptosoma), 70
subviridis (Boreconia), 462
subvitrea (Kanchia), 385
a (Leucoma), 385
sudias (Lemonias), 543, 544
suffusa (Lelia), 441
», (Ricine), 441
sulitelma (Colias), 242
sulphurescens (Artaxa), 400
5 (Euproctis), 400
sumatrensis (Nyctemera), 80
sundara (Euproctis), 403
superans (Lymantria), 486
Surattha, 154
susanna (Euproctis), 410
sybaris (Tarucus), 148
sylvana (Heliconius), iv
sylvanus (Augiades), xliii
54 (Lycnesthes), 200
a (Papilio), 200
sylvinus (Cibyra), 501, 502, 508
sylvius (Carterocephalus), lii
Symmachia, 537, 549
Synchloe, 292
synestalmenus (Antigonus), 203
5 (Sarangesa), 203
Syntomis, 154
Syrichthus, li, 294, 295, 296, 297
syrichthus (Hesperia), Ivi
syrnia (Deilemera), 72, 85
tacta (Rigema), 462
teenias (Mycalesis), 184
Tagiades, 203
Tanada, 82
elvi )
tarinta (Eurygona), 529, 533, 549
Tarucus, 143, 144, 148, 151
Tascina, 139
Tatochila, 292, 293, 299, 300
tavetensis (Leucoma), 382
taygete (CEneis), 240
Teara, 393, 405, 421, 446, 447, 474,
498
Tearosoma, 462, 474
Teia, 457, 458
telamonius (Caligo), 565
telesilla (Antiphella), 382
telicanus (Lampides), 289
ka (Tarucus), 144
Telipna, 194, 206
temesa (Symmachia), 537
temora (Salamis), 188
temperata (Olapa), 390
tenebrosa (Dasychira), 468
tenera (Hylemera), 436
tentyris (Euryphene), 192
tenuifascia (Nyctemera), 84
tenuis (Euproctis), 398
Teracolus, xi, xii, 142, 148, 144, 145,
146, 147, 148, 150, 152, 155, 157,
158, 160, 161, 162, 163, 202
terea (Papilio), 187
», (Precis), 187
Terias, 150, 151, 152, 155, 202, 206,
290, 299
terias (Aricoris), 548, 550
terminalis (Area), 409
ar (Euproctis), 409
| terminata (Genusa), 438
Ha (Pantana), 438
Terphothrix, 396, 428
terpsichore (Acrea), 186
As (Papilio), 186
(Pyrameis), 287, 299
5p (Vanessa), 287
tertiana (Nyctemera), 75
testacea (Cropera), 391
°F (Cyenia), 443
(Harapa), 442
(Lelia), 443
,, (Procodeca), 440
tetralunaria (Selenia), 159
Tetraphlebia, 279, 286, 298, 301
tetrophthalma (Ommatoptera), 447
Thamnocera, 462, 463, 470
Thanaos, 298, 299
thara (Lemonias), 544
theaphia (Scada), lv
Thecla, xlv, 289,290, 299
Thelde, 462, 465
thelestis (Acriea), 186
Themaca, 396, 413
themis (Euphedra), 190
hed
3
”)
( elvii )
theobene (Cymothoé), 192
a (Harma), 192
theodice (Pieris), 292, 293
», (Tatochila), 292, 293, 299
theodora (Riodina), 535, 549
theognis (Kuryphene), 192
theophrastus (Tarucus), 143, 144, 148,
151
thera (Mesosemia), 530, 549
Thestor, xxxi
thetis (Mesosemia), 530
Thiacidas, 461
thwaitesi (Dasychira), 472
thyellina (Orgyia), 459
Thylacogyna, 458
thymiathis (Anomeceotes), 435
Thyridia, 573
Ticilia, 498
_tilie (Dilina), xiv
tiphia (Leucoma), 381
tiphon (Ceenonympha), iii
» var. mixturata (Coenonympha),
241
tircis (Chamelimnas), 534
tiridates (Charaxes), 194
», (Papilio), 194
tisdala (Orgyia), 460
titania (Huproctis), 399, 407, 408
tithonus (Kpinephele), xlii
todara (Lymantria), 486
topha (Teracolus), 157, 162
Topomesa, 392
torrida (Euproctis), 410
tottea (Lymantria), 494
tragiscus (Satyrus), 277
transiens (Redoa), 378
transversa (Artaxa), 422
$3 (Euproctis), 422
Trichetra, 479, 497
Trichiura, 453
tricolor (Deilemera), 66
», (Leptosoma), 84
Trictena, 502, 507
trifasciata (Artaxa), 411
3 (Euproctis), 411
trigemmatus (Lampides), 289, 299
trimeni (Precis), XXXili, XXX1V
», (Pseudacreea), lxiv
Tripheromera, 55, 56, 64, 67, 82
Triphysa, iil
tripunctaria (Deilemera), 80
35 (Nyctemera), 53
35 (Phalena), 80
tripunctatus (Steropes), 296
trisignatus (Hesperia), 294, 299, 300
“3 (Scelothrix), 294
Tristania, 56
tristis (Argynnis), 277
tristis (Epinephele), 277, 298
», (Satyrus), 275, 277
5, _ (Stibomorpha), 275
| Trisula, 477, 498
| Trisuloides, 498
trita (Deilemera), 69
,, (Nyctemera), 69
tritoides (Nyctemera), 84
tritonea (Anthela), 448
tritum (Leptosoma), 69
trochilus (Caria), 538
3 (Chilades), 143, 145
_turbata (Orgyia), 458
turneri (Lymantria), 484
turnus (Papilio), 563
Turriga, 462, 464
typica (Mania), xxv
tyrrhenz (Buna), xxiii
| umbra (Papilio), 187
», (Planema), 187
umbraculata (Porina), 502
umbrina (Charnidas), 442
. (Leelia), 442
5p (Lymantria), 488
5 (Procodeca), 442
umbrosa (Porthetria), 482
undularis (Liptena), 197
undulata (Darala), 447
uniformis (Cherotricha), 420
x (Charnidas), 397
aA (Deilemera), 84
“ (Euproctis), 420
Re (Lelia), 443
unimacula (Artaxa), 411
v (Euproctis), 411
uniplaga (Deilemera), 61, 85
unipunctata (Kuproctis), 410
unisigna (Anthela), 447
Urocoma, 478, 479
urticee (Vanessa), x1x
usebia (Leucoma), 382
Utidava, 396, 429, 463
utilis (Euproctis), 415
uvaria (Colussa), 450
vacillans (Lymantria), 493
vagata (Nyctemera), 83
valdivize (Epinephele), 276
valdivianus (Butleria), 297, 299, 300
ss (Syrichthus), 297
vallonia (Ceratinia), lvi
Vanessa, xix, lxvii, 144, 188, 287, 557,
561
vanessoides (Elina), 274, 298
vapa (Scolitantides), 288
varanes (Charaxes), xxii, 194
», (Papilio), 194
», var. fulvescens (Palla), 194
varia (Anthela), 450
varia (Darala), 450
(Euproctis), 418
(Nola), 394
,, (Porthesia), 394
varians (Artaxa), 395, 406
,, (Deilemera), 57
,, (Kuproctis), 406
», (Nyctemera), 57
oa (Pitasila) 957
variegata (Artaxa), 425
(Daplasa), 428
(Dasychira), 477
(Euproctis), 428
a (Nephele), xxiii
variolosa (Deilemera), 60
(Nyctemera), 60
Be (Pitasila), 60
varipes (Hysibada), 386
Varmina, 461
varuniea (Nyctemera), 434
55 (Otroeda), 434
vashti (Amauris), 184
,, (Danais), 184
vata (Leucoma), 382
vauthieri (Colias), 291, 299
vecontia (Antiphella), 390
vectigera (Fodinoidea), 431
velans (Deilemera), 71
(Leptosoma), 71
», (Nyctemera), 71
velleda (Hepialus), 501
velutina (Aricoris), 547
= (Dasychira), 488
* (Lymantria), 488
a (Orgyia), 488
2)
9?
venatus (Pinacopteryx), xi,
153
venosa (Anthela), 446
;, (Artaxa), 401
», (Cispia), 430
», (Colussa), 446
», (Euproctis), 401
», (Lelia), 442
venosata (Gazalina), 387
veritabilis (Ceratinia), iv
veronica (Diestogyna), 192
», (Papilio), 192
versicolora (Endromis), ix
vesagus (Erebia), 284
», (Neosatyrus), 284,
301
vesperina (Nyctemera), 434
ss (Otroeda), 53, 434
vestalis (Acrea), 187
», (Planema), 187
vetustus (Heliconius), iv, lv
vibicipennis (Dasychira), 473
(Dasychira), 471, 472
(. clviii_ )
| vicina (Butleria), 296
vilis (Casama), 461
,, (Euproctis), 461
viluiensis (Colias), iv
vinacea (Lymantria), 485
vinidia (Acrea), 149, 150, 186
vinula (Cerura), xxiii
viola (Calliteara), 475
5, (Dasychira), 475
,, (Lymantria), 489
violacea (Lemonias), 543
violaceus (Neosatyrus), 284
virescens (Cadrusia), 476
35 (Charagia), 501, 502
54 (Dasychira), 476
i (Pseudonotodonta), 477
virgaureze (Chrysophanus), xlv
5 var, rutilus (Chrysophanus),
xlv
virgo (Euproctis), 398 _
virguncula (Euproctis), 392
<s (Porthesia), 392
viridescens (Acyphas), 459, 478
- (Orgyia), 459
viridis (Dasychira), 476
visum (Liparis), 437
», (Pantana), 437
vitellina (Euproctis), 412
(Liparis), 412
vollenhovii (Nyctemera), 84
vulgaris (Monotrichtis), 185
», (Mycalesis), 185
Vunga, 498
wallengrenii (Neomcenas), 278, 298
werdandi (Colias), liii
whitei (Dasychiva), 467
», (Ccura), 467
williamsianus (Arge), 281
a (Cosmosatyrus), 281,298
Xanthidia, 156, 202
xanthodice (Tatochila), 293
Xanthodura, 497
xanthopeva (Euproctis), 426
xanthorrhcea (Liparis), 394
. (Porthesia), 394
Xenosoma, 435
xerampelina (Aroa), 455
Pe (Gyniephora), 455
xylina (Lymantria), 490
Xylophasia, xvii
xypete (Cricosoma), 540
yerburii (Teracolus), xii, 160, 161, 162
Ypthima, 185, 206
zalmoxis (Drurya), lxv
zapateri (Hrebia), xlvi
zarepha (Ithomia), lvi
Zarfa, 386
zeboe (Artaxa), 400
cl
zeboe (Euproctis), 400
zerenoides (Deilemera), 64, 67
55 (Tripheromera), 64
Zeritis, 206
Zetes (Acrvea), Ixiv, 185
», (Papilio), 185
zetterstedti (Hypsophila), 243
zidora (Papilio), Lxiv
zingha (Monura), 194
», (Papilio), 194
zollikoferi (Xylophasia), Ixvii
zonata (Daraia), 449
Zonosoma, 55, 56
DIPTERA.
3slepharoptera, Lxi
Chersodromia, x]viii
Chironomus, xlvii, 521, 5
Cordylura, 1x
eynophila (Thyreophora), 1xi
equi (Gastrophilus), xlviii
fasciata (Lucina), 1xi
filiformis (Leptopa), 1x
flava (Cordylura), 1x
furcata (Thyreophora), 1x, 1xi
Gastrophilus, xlvili
germanicus (Pamponerus), xlix
hirta (Chersodromia), xlviti
Leptopa, lx
Lucina, lxi
modesta (Blepharoptera), Lxi
nasalis (Gastrophilus), xlviii
nigripennis (Pelidnoptera), 1xi
Orthocladius, xlvii, 521, 522, 523
Pamponerus, xlix
Pelidnoptera, 1xi
sordidellus (Chironomus), xlvii, 521,
522) 523
(Orthocladius), xlvii, 521,
522, 523
Thyreophora, lx, lxi
99. 5
aay
23
99
HEMIPTERA.
Adelphocoris, 181
aduncus (Neides), 180
wgyptius (Coranus), 181
Ablia, 180
agilis (Stenocephalus), 180
albomaculatus (Lygzeus), 180
andrei (Notochilus), 181
angustus (Emblethis), 181
anomalus (Lasiocoris), 181
Aoploscelis, 180
Aphanus, 181
ix)
arenarius (Trapezonotus), 181
artemisiz (Heterogaster), 180
aterrima (Brachypelta), 180
auriculata (Phyllontocheila), 181
Beosus, 181
Berytus, 180
biolleyi (Odopcea), 526
bivirgatus (Aoploscelis), 180
bolivari (Megalocoleus), 181
Brachycoleus, 181
Brachypelta, 180
brevicornis (Prionotylus), 180
brevipennis (Plinthisus), 180
cerulea (Zicrona), 180
_calcaratus (Miris), ii
| Calocoris, 181
| Camptobrochis, 181
Camptopus, 180
Capsus, 181
cardui (Phyllontocheila), 181
| Carineta, 527
Carpocoris, 180
carthusianus (Catoplatus), 181
Catoplatus, 181
Centrocoris, 180
cerinthe (Psacasta), 180
cervus (Edessa), 526
championi (Systellonotus), 181
cocksi (Salda), 181
Conostethus, 181
Coranus, 181
cordiger (Capsus), 181
», Var. fastidiosus (Capsus), 181
Coreus, 180
Corizus, 180
costaricensis (Gonatas), 526
crassicornis (Corizus), 180
Cyllocoris, 181
Cymodema, 180
Cymus, 180
Cyrtomenus, 525
decolor (Onychumenus), 181
Dictyonota, 181
Dicyphus, 181
Dimorphocoris, 181
discolor (Triphleps), 181
discrepans (‘linicephalus), 181
ditomoides (Metopoplax), 180
diversus (Gonatas), 526
Drymus, iii
dubuis (Sehirus), 180
echii (Monanthia), 181
Edessa, 526
Emblethis, 181
erratica (Megalocerza), 181
Kurydema, 180
Kurygaster, 180
exoletus (Phytocoris), 182
(
falleni (Pseudophlceus), 180
fasciata (Macroplax), 180
femoralis (Phytocoris), 182
ferus (Nabis), 181
festivum (Kurydema), 180
fossularum (Micrelytra), 180
fuliginosa (Dictyonota), 181
geminatus (Ischnorrhynchus), 180
geniculatus, var. disjunctus (Dicyphus),
181
Geocoris, 180
Geotomus, 180
Gerris, 181
Globiceps, 181
Gonatas, 526
gothicus (Lopus), 181
graminicola (Nysius), 180
grammicus (Odontotarsus), 180
Graphosoma, 180
Grypocoris, 181
Harpactor, 181
helferi (Sciocoris), 180
Heterogaster, 180
hirticornis (Berytus), 180
33 (Coreus), 180
histrionicus (Cyllocoris), 181
Homodemus, 181
hyalinipennis (Dicyphus), 181
hyalinus (Corizus), 180
Hydrometra, 181
Hypsitylus, 181
interrupta (Microplax), 180
iracundus (Harpactor), 181
Ischnorrhynchus, 180
levigatus (Miris), iii
Lasiocoris, 181
lateralis (Camptopus), 180
lativentris (Nabis), 181
leporina (Neottiglossa), 180
leucocephalus (Strongylocoris), 181
lineatum (Graphosoma), 180
lineatus (Nysius), 180
lividipennis (Dimorphocoris), 181
Lopus, 181
lutescens (Camptobrochis), 181
Lygzeus, 180
Macroplax, 180
maculipennis (Piezostethus), 181
merkeli (Pithanus), 181
maritimus (Beosus), 181
maura (Eurygaster), 180
medea (Odopeea), 527
Megalocerea, iii, 181
Megalocoleus, 181
melanocephalus (Cymus), 180
Metopoplax, 180
metriorhynchus (Platycranus), 181
in-flavum (Homodemus), 181
cha)
Micrelytra, 180
Microplax, 180
mirabilis (Cyrtomenus), 525
Monanthia, 181
montivagus (Berytus), 180
Nabis, 181
najas (Gerris), 181
nanus (Ochetostethus), 180
nebulosa (Rhaphigaster), 180
Neides, 180
Neottiglossa, 180
nigrocucullata (Kurygaster), 180
Notochilus, 181
noualhieri (Grypocoris), 181
Nysius, 180
obscurus (Strongylocoris), 181
Ochetostethus, 180
ocularis (Sthenarus), 181
Odontotarsus, 180
Odopcea, 526
oleraceum (Eurydema), 180
Onychumenus, 181
| Orthocephalus, 181
pallicornis (Dicyphus), 181
pandurus (Lygeus), 180
parumpunctatus (Corizus), 180
Peribalus, 180
Peritrechus, 180
Phyllontocheila, 181
Phytocoris, 182
picteti (Globiceps), 181
Piezostethus, 181
pilipes (Dryimus), iii
pilosella (Salda), 181
pini (Aphanus), 181
Pithanus, 181
plagiata (Microplax), 180
Platyeranus, 181
Plinthisus, 180
postica (Carineta), 527
prasinus (Hypsitylus), 181
Prionotylus, 180
Psacasta, 180
Pseudophleeus, 180
Pterotmetus, 180
punctipennis (Nysius), 180
punctulatus (Geotomus), 180
purpuripennis (Carpocoris), 180
quadrata (Verlusia), 180
quadratus (Aphanus), 181
reuterianus (Nabis), 181
Rhaphigaster, 180
Rhyparochromus, 180
roseomaculatus (Calocoris), 181
roseus (Conostethus), 181
rostrata (Alia), 180
ruber (Capsus), 181
rufus (Corizus), 180
rugosus (Nabis), 181
sabulicola (Rhyparochromus), 180
Salda, 181
saltator (Orthocephalus), 181
sanguineus (Harpactor), 181
saxatilis (Lygeus), 180
Sciocoris, 180
scutellaris (Capsus), 181
Sehirus, 180
siculus (Geocoris), 180
sphacelatus (Peribalus), 180
spiniger (Centrocoris), 180
stagnorum (Hydrometra), 181
staphylinoides (Pterotmetus), 180
Stenocephalus, 180
Sthenarus, 181
strichnocera (Dictyonota), 181
Strobilotoma, 180
Strongylocoris, 181
sulcicornis (Verlusia), 180
sulphureus (Calocoris, 181
superbus (Lygeus), 180
sylvestris (Peritrechus), 180
Systellonotus, 181
tabidum (Cymodema), 180
terricola (Piezostethus), 181
- Tinicephalus, 181
tipularius (Neides), 180
(
cha)
Trapezonotus, 181
triangularis (Brachycoleus), 181
Triphleps, 181
typheecornis (Strobilotoma), 180
ullrichi (Trapezonotus), 181
vandalicus (Adelphocoris), 181
Verlusia, 180
vestigiatus (Cyrtomenus), 525
vittiger, var. (Phytocoris), 182
waltli (Pseudophlceus), 180
Zicrona, 180
NEUROPTERA.
Chrysopa, li
ephippiger (Hemianax), x, xiv
Hemianax, x, xiv
Nemoptera, 168
Orthetrum, ix, Xiv
vulgaris (Chrysopa), li
ORTHOPTERA.
Anisolabis, lix
colossea (Anisolabis), lix
Labidura, xxxvi
riparia (Labidura), xxxvi
Marcu 16, 1904
TRANS. ENT. SOC. LOND., Vv. 1903. M
CHARTER AND BYE-LAWS
OF THE
ENTOMOLOGICAL SOCIETY OF LONDON.
FOUNDED IN 1833.
INCORPORATED BY ROYAL CHARTER IN 1885.
CONTENTS.
te
Page
CHARTER : : : : Vv
BYE-LAWS.
CHAPTER I.—Object of the Society : . Vili
™ I1.—Constitution of the Society ; : vill
» Ill.—Management . , é : : vill
+3 IV.—Officers . ; ; . Vili
. V.—Removal or Resignation of Officers. — vill
= VI.—Of the President : : 3 : 1x
,, WII.—Of the Vice-Presidents ? f ; 1X
VIII.—Of the Treasurer. ix
es IX.—Of the Secretaries. : : : x
: X.—Of the Inbrarian . ; i
s X1L.— Library Regulations ; ‘ x1
,» XNII.—EHlection of Fellows . : : : xi
», NITL—O; the Admission Fee and Annual
Contribution ; : ; xii
XIV.— Withdrawing and Removal of Fellows xii
9
1V CONTENTS,
CHAPTER XV.—Privileges of Fellows
5 XVI.—Honorary Fellows
i XVII.—Ordinary Meetings of the Society .
5» NVIITI.—Special Meeting
ss XIX.—Annual Meeting .
55 XX.—Transactions and Journal of Pro-
ceedings
5 XXI.—Alteration of the Bye-Laws .
THE SCHEDULE
Containing Form of Balloting List for the Election of
Officers and Council
XV
xvii
XVIll
CHARTER.
———
Victoria, by the Grace of God of the United Kingdom
of Great Britain and Ireland, Queen, Defender of the Faith.
TO ALL TO WHOM these presents shall come Greeting :
WueEreAS JOSEPH WILLIAM DuNnnNING, of Lincoln’s Inn, in
the County of Middlesex, Barrister-at-Law, Esquire, Master
of Arts, formerly Fellow of Trinity College, Cambridge,
Fellow of the Cambridge Philosophical Society and of the
Linnean and Zoological Societies of London, has by his
Petition humbly represented unto US, That in the year 1833
certain of our loyal subjects formed themselves into a Society
for the Improvement and Diffusion of Entomological Science,
and subscribed and expended considerable sums of money for
such purposes, and have collected and become possessed of a
valuable library and other property, and have been and
continue to be actively employed in promoting the objects for
which the said Society was founded, especially by the
publication of Volumes of Transactions composed of Original
Memoirs, read before the Society. AND wHeEREAS the said
Petitioner, believing that the well-being and usefulness of
the said Society would be most materially promoted by
obtaining a Charter of Incorporation, hath therefore, on
behalf of himself and the other Members of the said Society,
most humbly prayed that WE would be pleased to grant a
Royal Charter for incorporating into a Society the several
persons who have already become Fellows, or who may at
any time hereafter become Fellows thereof, subject to such
Regulations and Restrictions as to US may seem good and
v1 CHARTER.
expedient. NOW KNOW YE that WE, being desirous of
encouraging a design so laudable, and of promoting the
improvement and diffusion of Science in all its branches,
have of Our especial Grace, certain Knowledge and mere
Motion, given and granted, and We do hereby give and
grant, That the said JosepH Witi1Am Dunnine and such
others of Our loving subjects as are now Fellows of the said
Society, or who shall at any time hereafter become Fellows
thereof in pursuance of the provisions of this Our Charter
and according to such Bye-Laws as are hereinafter men-
tioned, shall be a Body Corporate by the name of “ The
Entomological Society of London,” having perpetual succes-
sion and a common seal, with power to sue and to be sued in
their Corporate name, and to acquire and hold any goods and
chattels whatsoever.
And our Will and Pleasure is, That JoHN OBADIAH
Westwoop, Esq., Master of Arts, Hope Professor of Zoology
in the University of Oxford, shall be Honorary President of
the said Corporation during the term of his natural life.
And that Roperr MacLacuian, F.R.S., shall be the first
President of the said Corporation and shall continue such
until the Annual Meeting to be held in the month of January
next.
And our Will and Pleasure is, And we do hereby declare,
That there shall always be a Council to direct and manage
the concerns of the said Corporation. And that the thirteen
persons, who were elected to form the Council of the said
Society at the Annual Meeting held in the month of January
last, shall form the first Council of the said Corporation, and
shall continue in Office until the Annual Meeting to be held
in the month of January next.
And our Will and Pleasure is, And we further grant and
declare, that the existing Bye-Laws of the said Society, as
revised and amended at a General Meeting held on the 2nd
day of May, 1883, shall be the Bye-Laws of the said
Corporation, until the same shall be revoked or altered as
hereinafter mentioned. And that it shall be lawful at
General Meetings of the said Corporation to revoke or alter
any former Bye-Laws, and to make such new Bye-Laws as
CHARTER. vil
shall be deemed useful and necessary for the regulation of
the said Body Corporate.
Provided always: And we lastly declare it to be our Royal
Will and Pleasure, That no Bye-Law or Resolution shall, on
any account or pretence whatsoever, be made by the said
Corporation in opposition to the general scope, true intent,
and meaning of this our Charter or the Laws and Statutes of
this Realm, and that if any such Bye-Law or Resolution
shall be made, the same shall be absolutely null and void.
In Witness whereof We have caused these our Letters to
be made Patent. __
Witness Ourself at Westminster the A
twentieth day of July, in the Forty-ninth
year of Our Reign. THE SEAL.
By Warrant under the Queen’s Sign
Manual. NY
ae
( viii )
BYE-LAWS.
As AMENDED AT A SPECIAL MEETING HELD 3RD DECEMBER, 1902.
Cuap. I. Object.
Tue EntomoLocicaL Socrety or Lonpon is instituted for
the improvement and diffusion of Entomological Science.
Cuap. II. Constitution.
The Society shall consist of Honorary and Ordinary
Fellows.
Cuap. III. Management.
The affairs of the Society shall be conducted by a Council
consisting of the Officers of the Society hereinafter mentioned
—other than the three Vice-Presidents—and of twelve
ordinary Members to be chosen annually from among the
Fellows. No Fellow shall be eligible as an ordinary Member
of the Council for more than three years successively. Five
shall be a quorum.
Cuap. IV. Officers.
The Officers of the Society shall consist of a President ;
three Vice-Presidents ; a Treasurer ; two Secretaries; and a
Librarian. The Officers shall be chosen annually. No
Fellow shall be President, or a Vice-President, more than
two years successively.
Cuap. V. Removal or Resignation of Officers.
1. For any cause which shall appear sufficient to a majority
thereof, the Council shall have power to suspend any Ofticer
of the Society from the exercise of his office, or to remove
him and declare such office vacant.
BYE-LAWS, 1x
2. In the event of any vacancy occurring in the Council
or Officers of the Society, at the next meeting of Council
after such vacancy has been made known, the Council shall
elect some Fellow to fill the vacancy till the Annual Meeting.
Cuap. VI. President.
1. The duty of the President shall be to preside at the
Meetings of the Society and Council, and regulate all the
discussions and proceedings therein, and to execute or see to
the execution of the Bye-Laws and orders of the Society.
2. In case of an equality of Votes the President shall have
a double or casting Vote.
CuHap. VII. Vice-Presidents.
1. The Vice-Presidents shall be nominated by the Presi-
dent, from amongst the members of the Council. Such
nomination shall be declared at the Ordinary Meeting next
after the election of the President in every year.
2. In the absence of the President a Vice-President shall
fill his place, and shall for the time being have all the
authority, power, and privilege of the President.
3. In the absence of all the Vice-Presidents a Member of
the Council shall preside ; and if no Member of the Council
be present at an Ordinary Meeting, the Fellows present shall
appoint by a majority to be Chairman such Fellow as they
shall think fit; and the Member of Council so presiding, or
the Fellow so appointed, shall for the time being have all the
authority, power, and privilege of the President.
Cuap. VIII. Treasurer.
1. It shall be the duty of the Treasurer to demand and
receive for the use of the Society all sums of money due or
payable to the Society, and to disburse all sums payable by
the Society out of the Funds in his hands.
2. No payment exceeding £5, excepting for rent or taxes,
shall be made by the Treasurer without the consent of the
Council,
Xx BYE-LAWS.
3. The Treasurer shall keep a book of Cheque Receipts for
admission fees and annual payments ; each Receipt shall be
signed by himself, the date of payment and name of the
Fellow paying being written both on the Receipt and on the
part of the Cheque which is left in the book.
4, The Treasurer shall demand all arrears of annual
payment after such payment shall have been due three
months.
-5, The accounts of the Treasurer shall be audited annually,
previously to the Annual Meeting, by a Committee of six
Fellows (of whom three shall be Members of the Council),
to be appointed by the President at the Ordinary Meeting
in December, of which Committee three (or two, provided
that one of them is not a member of the Council) shall be a
quorum. The Treasurer shall furnish the Auditors with a
detailed account of all receipts and disbursements down to
the 31st December.
Cuap. IX. Secretaries.
1. It shall be the duty of the Secretaries to keep a list of
all the Fellows of the Society, together with their addresses ;
to summon Meetings (when necessary) of the Society and the
Council ; to conduct and produce to the Council all corre-
spondence in any way connected with the Society at the next
Meeting after such correspondence shall have been received or
taken place; to take Minutes of the Proceedings at Meetings
of the Society and the Council; to edit the Transactions and
Journal of Proceedings; and, generally, to act under the
direction of the Council in all matters connected with the
welfare of the Society.
2. In the absence from any Meeting of the Society, or the
Council, of both the Secretaries, Minutes of the Proceedings
shall be taken by a Fellow whom the President shall appoint
for the occasion.
Cuap. X. Librarian.
1. It shall be the duty of the Librarian to take care of the
Library and MSS., and keep a Catalogue thereof, with the
names of the Donors; to call in all Books borrowed, and see
that the Library regulations are carried into effect.
BYE-LAWS. at
2. The Council may employ a Sub-Librarian, who shall
receive such remuneration as the Council shall from time to
time determine, and shall be subject to such Rules and
Orders as shall from time to time be given to him by the
Council.
Cuap. XI. Library Regulations.
1. No Fellow shall, without special permission of the
Council, be allowed to borrow from the Library more than
four volumes at one time, or without leave of the Librarian,
to retain any volume longer than one month.
2. If any book be torn, injured, lost, or not forthcoming
when demanded by the Librarian, full compensation shall be
made for the same by the borrower.
3. The Librarian shall call in all books borrowed from the
Library on the 5th day of January and 5th day of July in
each year ; and in case the same be not returned on or before
the Ordinary Meeting of the Society in the following month,
notice thereof shall be given by him to the Council, who
shall then direct a second notice to be sent to the Fellow
retaining any book, and in case the same be not returned
within the further space of four weeks from the date of such
second notice so sent, such Fellow shall in future be dis-
qualified from borrowing books from the Library without the
special permission of the Council.
4. Subject to such Regulations as may be made from time
to time by the Council, the Library shall be open to the
Fellows between the hours of one and six p.m. on every
week-day, except Saturday, and on that day between one and
three p.m.
5. Nostranger shall be allowed access to the Library unless
introduced by a Fellow ; but a note addressed to the Librarian
or Secretary shall be deemed a sufficient introduction.
Cuap. XII. Election of Fellows.
1. Every Candidate for admission into the Society shall be
proposed by three or more Fellows, to one of whom he shall
be personally known, and they must sign a Certificate in
recommendation of him. The Certificate shall specify the
name and usual place of residence of the Candidate.
xl BYE-LAWS.
2. The Certificate having been read at one of the Ordinary
Meetings shall be suspended in the Library, read again at
the following Ordinary Meeting, and the person therein re-
commended shall be balloted for at the next Ordinary Meeting.
3. The method of voting shall be by ballot, and two-thirds
of the Fellows balloting shall elect.
4. Fellows shall sign the Obligation Book of the Society
at the first Ordinary Meeting of the Society at which they
are present, and shall then be admitted by the President.
Cuap. XIII. Admission Fee and Annual Contribution.
1. The Admission Fee shall be £2 2s., the Annual Con-
tribution £1 1s.
2. Fellows permanently resident out of the United King-
dom shall pay the Annual Contribution, but shall be exempt
from payment of any Admission Fee.
3. The composition for Life Fellowship, in lieu of the
Annual Contribution, shall be £15 15s.
4. The Annual Contribution shall become due on the Ist
day of January in advance; any Fellow elected after Sep-
tember will not be called upon for his Contribution for
that year.
Cuap. XIV. Withdrawing and Removal of Fellows.
1. Every Fellow, having paid all sums due to the Society,
shall be at liberty to withdraw therefrom upon giving notice
in writing to the Secretary.
2. Whenever written notice of a motion for removing any
Fellow shall be delivered to the Secretary, signed by the
President or Chairman for the time being on the part of the
Council, or by six or more Fellows, such notice shall be read
from the Chair at the two Ordinary Meetings immediately
following the delivery thereof, and the next following Ordinary
Meeting shall be made a Special Meeting and the Fellows
summoned accordingly, when such motion shall be taken into
consideration and decided by ballot ; whereat if a majority of
the Fellows balloting shall vote that such Fellow be removed,
he shall be removed from the Society.
3. In the month of November in each year the Council
BYE-LAWS. Xill
shall cause to be suspended in the Library of the Society a
list of the Fellows who owe more than two Annual Contri-
butions. If the Contribution due from any Fellow named in the
said list shall not have been paid within three months after
the first suspension of the list, the Council may remove such
Fellow from the Society, but notwithstanding such removal
any Fellow so removed shall continue liable to pay, and may
be sued for the recovery of any money due from him to the
Society. The Council may remit wholly, or in part, the
entrance fee payable by any former Fellow rejoining the
Society.
Cuap. XV. Privileges of Fellows.
1. Fellows have the right to be present, to state their
opinions, and to vote, at all General Meetings; to propose
Candidates for admission into the Society; to introduce
Visitors at General Meetings of the Society ; to have personal
access, and to introduce scientific strangers, to the Library ;
and Fellows who have paid the Annual Contribution for the
year shall be entitled to receive a copy of the Transactions
published during the year.
2. A Fellow shall not be eligible to any office in the Society,
or to the Council, unless he shall have paid his Annual
Contribution for the year previous to that in which the
Election takes place.
3. A Fellow shall not be entitled to vote on any occasion
until he shall have paid his Contribution for the year last
past.
Cuap. XVI. Honorary Fellows.
1. Every person proposed as an Honorary Fellow shall be
recommended by the Council; and shall be balloted for, and,
if elected, be liable to be removed in the like form and manner,
and be subject to the same rules and restrictions, as an
Ordinary Fellow.
2. Honorary Fellows shall be exempt from the payment of
Fees and Contributions, and shall possess all the privileges
of Ordinary Fellows.
3. No British Subject shall be an Honorary Fellow.
4. The number of Honorary Fellows shall not exceed twelve.
X1V BYE- LAWS.
Cuap. XVII. Ordinary Meetings of the Society.
1. The Ordinary Meetings of the Society shall be held on
the first Wednesday in each month (except January, July,
August and September), beginning at eight o’clock in the
evening, or at such other days or times as the Council shall
from time to time direct.
2. At the Ordinary Meetings the order of business shall be
as follows :—
(1.) The names of the Visitors present at the Meeting
shall be read aloud by the President.
(2.) The Minutes of the last Meeting shall be read
aloud by one of the Secretaries, proposed for
confirmation by the Meeting, and signed by the
President.
(3.) The Presents made to the Society since the last
Meeting shall be announced and exhibited.
(4.) Certificates in favour of Candidates for admission
into the Society shall be read, and Candidates
shall be balloted for.
(5.) Fellows shall sign their names in the Obligation
Book, and be admitted.
(6.) Exhibitions of specimens, &e., shall be made.
(7.) Entomological communications shall be announced
and read either by the Author or by one of the
Secretaries.
(8.) Business not specified in the above order and dis-
cussions arising out of the exhibitions and com-
munications shall be taken at such times and in
such manner as the President shall direct.
3. All Memoirs which shall be read at any Meeting of
the Society, and accepted for publication, shall become the
property of the Society, unless otherwise stipulated before
the reading thereof.
4. No Motion relating to the Government of the Society,
its Bye-Laws, the management of its concerns, or the election,
appointment, or removal of its Officers, shall be made at any
Ordinary Meeting.
BYE-LAWS. XV
Cuap. XVIII. Special Meeting.
1. Upon the requisition of six or more Fellows, presented
to the President and Council, a Special General Meeting of
the Society shall be convened ; a notice thereof shall be sent
to every Fellow whose last known residence shall be in the
United Kingdom, at least seven days before such Meeting
shall take place: and any motion to be submitted to such
Meeting which involves a substantive proposition and is not
of the nature of an amendment shall be stated at length in
such notice.
2. No vote shall be taken at any Special Meeting unless
nine or more Fellows shall be present.
Cuap. XIX. Annual Meeting.
1. The Annual Meeting of the Society shall be held on the
third Wednesday in January.
2. The objects of the Meeting shall be to receive from the
Council, and hear read, their Annual Report on the general
concerns of the Society ; and to elect the Council and Officers
for the ensuing year.
3. The Council for the time being shall annually cause to
be prepared a list which shall contain the names of such
Fellows being eligible as they shall recommend to fill the
offices of President, Treasurer, Secretaries, and Librarian for
the year ensuing ; and also the names of such Fellows as they
shall recommend to be re-elected, and of other Fellows to
be elected into the Council. The list shall include the names
of not less than twelve Fellows recommended as ordinary
members of the Council.
4. The list prepared by the Council shall be read at the
Ordinary Meeting next but one before the Annual Meeting.
At the Ordinary Meeting preceding the Annual Meeting, the
names of other candidates to fill any of the offices, or to
serve as Members of the Council (each proposed and supported
by at least four properly qualified Fellows of the Society), ,
shall be received. Nominations by post made by four pro-
perly qualified Fellows, and received prior to this Meeting,
shall also be accepted. These shall be added to the Council’s
xvi BYE-LAWS.
list, and copies of the complete list shall, before the 20th
December, be transmitted to every Fellow whose last known
residence shall be in the United Kingdom, and who shall
have paid his subscription for the current year.
5. The election shall be by Ballot at the Annual Meeting,
and copies of the final list shall be used as Ballot papers.
The President shall appoint two or more Scrutineers from the
Fellows present, not being Members of the Couucil, to super-
intend the Ballots and report the results to the Meeting.
The Secretaries, assisted by the Treasurer, shall prepare a list
of the Fellows entitled to vote, and each Fellow voting shall
give his name to the Scrutineers to be marked on the said
List, and shall then put his Ballot paper into the respective
receptacles to be provided for such occasion.
6. A Fellow voting may strike out any name or names on
the said List, and the Scrutineers shall record the votes
accordingly, but any Ballot paper which shall contain a
greater number of names for any Office or Position than the
number to be elected to such Office or Position shall be wholly
void, and be rejected by the Scrutineers. No Ballot shall be
taken unless nine or more Fellows shall be present.
7. If from any cause any election which ought to take
place at the Annual Meeting shall not take place, then such
election shall be adjourned until the next convenient day, of
which notice shall be given in like manner as is directed for
the Annual Meeting.
8. The form given in the Schedule shall be used, with such
variations as may be considered by the Council necessary or
desirable.
CuHap. XX. Transactions and Journal of Proceedings.
1. The Transactions shall consist of such papers commu-
nicated to the Meetings of the Society as the Council shall
order to be published therein.
2. The Transactions shall be published quarterly, or at
such other times, and at such prices as the Council shall
direct for each Part or Volume.
3. Authors of Memoirs published in the Transactions shall
be allowed twenty-five copies of their communications gratis.
BYE-LAWS. Xvil
If any additional number be required, the permission of the
Council shall be first obtained, and the entire expense thereof
shall be paid for by the Authors,
4, A Journal of Proceedings of the Society shall also be
published, containing Abstracts of the Papers read and
Notices of other Matters communicated at the Ordinary
Meetings of the Society. The Proceedings shall be bound
up with the Transactions.
Cuap. XXI. Alteration of the Bye-Laws.
Any of the Bye-Laws of the Society may at any time be
repealed or altered, or others adopted in lieu thereof, at a
Special Meeting of the Society, to be held after a Notice
given to the President and Council, signed by six Fellows at
least, and specifying the intended repeal or alteration, has
been read at three Ordinary Meetings of the Society.
XVlll BYE-LAWS.
THE SCHEDULE REFERRED TO IN CHAPTER XIX.
Balloting List for the Election of Officers and Council.
Office. List.
President. Li
Treasurer. MWA, IDE
| Secretaries. | T. G. |
Librarian. N. M.
| Council. | Ae.
5: Ee:
Wi
IW oe Xe.
Instructions.
In using the above list as a Ballot paper a Fellow voting
may strike out any name or names printed therein, but any
Ballot paper which when placed in the receptacle shall contain
a greater number of names for any Office or Position than the
number to be elected to such Office or Position shall be wholly
void and rejected by the Scrutineers.
Trans Ent. Soc Lond 1903 .Pt.1.
L.R.Crawshay del. Mintern Bros. lith.
Transformations of Drlus flavescens.
. | Trans Ent. See Lond 1903. PUI.
L.R.Crawshay del. Mintern Bros. hth.
Transformations of Drilus flavescens.
Zrans. Ent. Soc. Lond. 1903. Pl. 117.
André & Sleigh, Limited
NEW SPECIES OF DEILEMERA.
* *
Trans. Ent. Soc. Lond. 1g03.. Pl. IV.
Z
André & Sleigh, Limited
NEW SPECIES OF DEILEMERA.
Trans. Ent. Soc, Lond.1903.P1l. V.
E Wilson, Cambridge.
Nanophyes durieui,
with its larva, pupa ke.
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Nest of Trigona collina.
Carl Hentschel, sculp.
et imp.
VII.
PAS
Lond., 1903.
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7
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Trans. Ent. Soc Lond .1903. PLVUL.
R.Mintern del-et lth, Mintern Bros.Chromo
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“Westwood Bequest.” Trans. Ent. Soc. Lond. 1908. Pl. XI.
Isis Al.
F. N. Clark, Phot. Mintern Bros., Eng.
Orina tristis, var. smaragdina.
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Horace Knight, del. André & Sleigh, Limited.
Chilean Butterflies.
Trans Ent. Soe Lond. IW Pl NIM.
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Chilean Butterfhes.
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Trans. Ent. Soc.
Chilean Butterflies.
Lond., 1907. Plate XIV.
André & Sleigh, Limited.
Horace Knight, del.
Trans. Ent. Soc. Lond., 1903. Plate XV.
André & Sleigh, Limited.
Chilean Butterflies.
Trans. Ent. Soc., Lond., 19037. Pl. XVI.
P.J. Bayzand, del. André & Sleigh, Limited
Results of Experiments in 1893 upon the Colour-relation between the
larve of Odontopera bidentata and their environment.
Trans. Ent. Soc., Lond., 1903. Vet QW.
P. J. Bayzand, del. André & Sleigh, Limited.
Effects of lichen, of black twigs, and of reddish brown sticks upo:
the larve of Gastropacha quercifolia (1893-4).
Trans. Ent. Soc., Lond., 1903. Pl. XVIII.
P. J. Bayzand, del. André & Sleigh, Linut
Effects of lichen and of reddish brown sticks upon the larve o
Gastropacha quercifolia (1893-4).
Trans.Ent.Soc.Lond 1903. Pl. XIX.
A.Quail del Mintern Bros. hth.
Segments of Antenne of Hepialide.
race Knight ad nat. lith
(“ ant ] ann Sanit ft \merinrgay } Tay a aia ia & 4
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