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TRANSACTIONS
OF THE
ENTOMOLOGICAL SOCIETY
OF
LONDON.
ERRATA.
TRANSACTIONS.
Page 9, line 7 from bottom, for multas paleas read multis paleis.
Page 38, line 2 from top, for July 8th read July 8th.
Page 45, line 28 from top, for regentsteinensis read regenstetnensis.
Page 51, line 4 from bottom, for cwrulea read cerulea,
Page 94, line 5 from bottom, for Epinephale read LEpinephile.
Page 109, line 5 from bottom, for Cephanodes read Cephonodes.
Page 113, line 16 from bottom, for charonia, Dru. read canace, Linn.
Page 115, line 17 from top, for wndularis read caudata, Butl., an aberra-
tion with a broad fulvous shade from the base parallel to the inner
margin, approximating to the colouring of the °.
Page 125, line 7 from top, for Calysime read Calysisme.
Page 127, line 24 from top, for ariadne read méznorata, Moore.
Page 128, last line but two, for daret?s, Hew. read drypetes, Hew.
Page 129, lines 5,6, 7 from top, insert a comma after “took,” and de/ete
“a single... also.” The sentence will then read—. . . “I took,
settled on aleaf...,a specimen... .”
Page 136, line 14 from top, for shakra read schakra.
Page 378, line 22 from top, for phlwas read phlwas.
Page 400, line 21 from top
Page 404, line 6 from top } for Mount read Md.
Page 406, line 19 from top
Page 406, line 2 from bottom
é } for Bon read Bou
Page 408, line 22 from top
PROCEEDINGS.
Page xxxv, line 16 from top, and line 26 from top, for Syrtcthus read
Syrichthus.
Page xliii, line 13 from top, for G. album read C. album.
Page liv, line 2 from top, for Zemophilus read Lemophleus.
THE
TRANSACTIONS
OF THE
ENTOMOLOGICAL SOCIETY
LONDON
FOR. THEY EA
1905.
LONDON:
PRINTED FOR THE SOCIETY BY RICHARD CLAY AND SONS, LIMITED,
LONDON AND BUNGAY.
SOLD AT THE SOCIETY'S ROOMS, 11, CHANDOS STREET,
CAVENDISH SQUARE, W.,
AND BY LONGMANS, GREEN, AND CoO,,
PATERNOSTER ROW, E.C.; AND NEW YORK.
1905-1906.
QL 4el
Res
DATES OF PUBLICATION IN PARTS.
Part I. (Trans., pp. 1-202, Proc., i-xvi) was published 20th May, 1905
y: LS Fos 203-268, ,, xvil—xliv) bs 15th July, _,,
ed. iC os, 269-364) sa » 4th Oct., 55
3 DE rs 365-438, ,, xlv—lxxvi) 3rd Jan., 1906
, Vv. (—_—-—————-_,,, | l xxvii-cxlv) 3) eh Mer...
ar FA 1
_
Vusects ,
ENTOMOLOGICAL SOCIETY OF LONDON.
FouNDED, 1833.
INCORPORATED BY RoyaL CHARTER, 1885.
OFFICERS and COUNCIL for the SESSION 1905-1906.
President.
FREDERIC MERRIFIELD.
Vicc=Presidents,
Dr. THOMAS ALGERNON CHAPMAN, M.D., F.Z8.
Dr. FREDERICK AUGUSTUS DIXEY, M.A., M.D.
Pror. EDWARD B. POULTON, D.Sc., M.A., F.R.S.
Treasurer,
ALBERT HUGH JONES.
Secretaries,
HENRY ROWLAND-BROWN, M.A.
CoMMANDER JAMES J. WALKER, M.A., R.N., F.LS.
Librarian,
GEORGE C. CHAMPION, F.ZS.
Other Members of Council.
GILBERT JOHN ARROW.
CoLonEL CHARLES T. BINGHAM, F.Z.S.
JAMES EDWARD COLLIN.
HAMILTON H.C. J. DRUCE, F.ZS.
HERBERT GOSS, F.L.S.
WILLIAM JOHN LUCAS, B.A.
LOUIS BEETHOVEN PROUT.
EDWARD SAUNDERS, F.R.S., F.LS.
CotonEL JOHN W. YERBURY, R.A., F.Z.S.
Resident Librarian.
GEORGE BETHELL, F.R. Hist. S.
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CONTENTS.
Errata
Explanation of the mletes
List of Fellows... ae
Additions to the Library
MEMOIRS.
I. The Life History of pes chinensis, Felder. ry Joun C. W.
KersHaw, F.L.S., F.E ade aoc
IL. Butterfly-destroyers in eee China. . Byd Joun C. W. Kersuaw,
BLS; BSS. .
Ill. On Erebia palarica, un. sp., and Evrebia stygne ; chiefly { in Bay
to its association with E. evias, in Spain. ved THomas A.
CHApMAN, M.D., F.Z.8. nae Boe
IV. Another Entomological Excursionto Spain. By GEORGE CHARLES
Cuampion, F.Z.S., and Tuomas ALGERNON CHAPMAN, M. iD
E.Z8.; with descriptions of two new ose of Hemiptera, by
’ Prof, O. M. REvrER 7
V. The Egg and Early Larval gah of a Coreid aad probably
Dalader acuticosta, Amyot et Serv.; with a note on its Hymeno-
pterous Parasite. By Nrtson ANNANDALE, B.A., ake
Superintendent of the Indian Museum, Calcutta eos
VI. Notes on the Butterflies observed in a tour through India and
Ceylon, 1908-4. By G. B. Lonesrarr, M.D., Oxon.
VII. The genus Criocephalus. By D.Suarp, M.A., F.R.S. With Notes
on the habits of Asemum striatum and Croaptate Mite
By F. Girsert Smite L
(VIIa. The Habits of Asemwm striatum and Cr Nias ferus) .
VIII. On the matrivorous habit of the pee Hetereynis, Rmbr
By T. A. Cuapman, M.D., F.Z.S. ...
IX. Descriptions of some new species of Satyride from coe
America, By Freperick Du CANE ai oe DiCilic. HRS;
etc.
X. Additions to a 2d beatbales of the e Homopterou Fanily € Cicadidz.
By W. L. Distant
XI. Ox ihe sg irl crea of Pits in T. A. Coarman, M.D.,
ZS.
37
55
61
. 165
al it
. 185
. 191
. 203
XII. ae on aw Zealand Lepidoptera. By E. Mevnie BA,PRS,
F.ES. . 219
XIII. On three remarkable new mens of _Mierotepidoptra a's SIR
: . 245
GrEOoRGE F. Hampson, Bart., B.A
ot
( viii )
PAGE
XIV. Descriptions of some new species of Diurnal Lepidoptera, collected
by Mr. Harold Cookson, in Northern Rhodesia, in 1903 and
1904. By Herpert Drvce, F.LS. ees and H. legs
by Hamitton H. Druvce, F.ZS. . 251
XV. Pseudacrea pogger. and Limnas Gaeiee the numerical pro-
portion of mimic to model. By Horace A. Byart, B.A,
F.E.S. With a note by Professor E. B. Pour cae 'D. Se.” 5
My ANs Heh s)tCs sexe sue Dae
XVI. A Monograph of me eee Our ws, ct GeOoRGE T. BETHUNE-
Baker, F.L.8., on see 369 me ..- 269
XVII. The structure and life history of Peyeheas sexpunctata, Curtis.
By Jon ALEXANDER Det, B.Sc. Communicated ye Prof.
L. C. Mratt, F.R.S.,, F.E.S. a ne . 293
XVIII. New African Lastocampide in the British Museum. Fe Prof.
Cur. AURIVILLIUS, Hon. F.E.S,, F.M.Z.S8., etc. a . old
| XIX. Memoir on the Rhynchota collected by Dr. ArTHUR WILLEY,
: F.R.S., chiefly in Birara Pos ae and Lifu. ey G. W.
KirKaLpy, F.E.S. ... . 827
XX. The Blind Coleoptera of Australia and Tasmania. ee ARTHUR
M. Lea, F.E.S., Government Entomologist, Tasmania .«» 300
XXI. On a Collection of Butterflies and Moths made in Marocco, in
1900-01-02. By E. G. B. Merapr-Watpo. Communicated
by H. J. Ewes, F.RS., etc. ... fie ie «-» OOD
XXII. A new species of the Hymenopterous Genus JMZegalyra, West-
wood. By J. Cuester Bravery, Ithaca, N.Y., U.S.A.
Communicated by Col. C. T. BincHam, F.Z.S. ban we 895
XXIII. Hymenoptera Aculeata collected in Algeria by the Rev. A. E.
Eaton, M.A., F.E.S., and the Rev. Francis Davip Morice,
M.A., F.E.S. Part II. Pee mE EpwWARD SAUNDERS,
F.RS., ete, oo. s 4c ws» 399
XXIV. On the Ichneumonidous ae Tr ani ae Palmer
With descriptions of new species. By CiaupE Mor ey,
Proceedings for 1905 ep oe ise aks i sk .. i—)xxvi
Annual Meeting ... ies Re ses ise aS bee .. Ixxvii
President’s Address oe de ict ae 33 bis hoa a lberocyin
General Index nee ain a és ee BS oe ae CXa
Special Index Aa aa oes ae “ee Se ase reap CACVILE
EXPLANATION OF THE PLATES.
Plate I. Seepage 4 Plate XII. See page 218
Plates II to VI. - 34—35 Plate XIII. " 262
Plate VII. i 54 Plate XIV. 5 as 268
Plate VIII. 2 59 Plate XV. Fs 292
Plate IX. S 164 Plate XVI. a 326
Plate X. $> 190 Plate XVII. LA ik 363
Plate XI. - 202 Plates XVIII, XIX. ,, 393
Pist of Fellows
ENTOMOLOGICAL SOCIETY OF LONDON.
Marked * have died during the year.
Date of HONORARY FELLOWS.
Election.
1900 AURIVILLIUS, Professor Christopher, Stockholm.
1905 Bottvar, Don Ignacio, Paseo de Recoletos Bajo, 20, Madrid.
1901 Fasre, J. H., Sérignan, Vaucluse, France.
1894 Foret, Professor Auguste, M.D., Chigny, prés Morges, Switzerland.
1906 GaneLBAvER, Professor Ludwig von, Hof Musewm, Vienna.
1898 Grassi, Professor Battista, The University, Rome.
1884 Osten SackeENn, Baron C. R., Bunsenstrasse 8, Heidelberg.
1884 * PackarD, Dr. Alpheus S8., Providence, Rhode Island, U.S.A.
1906 ReuTER, Professor Opo Morannat, The University, Helsingfors,
Finland.
1872 * SaussurE, Henri F. de, Tertasse 2, Geneva.
1895 ScuppER, Samuel Hubbard, Cambridge, Mass., U.S.A.
1885 SNELLEN, P. C. T., Rotterdam.
1893 WarrenwyL, Hofrath Dr. Carl Brunner Von, Lerchenfeldstrasse 28,
Vienna.
1898 Weismann, Dr. August, Freiburg, Baden.
FELLOWS.
Marked + have compounded for their Annual Subscriptions.
Date of
Election.
1901 + Aparr, Sir Frederick E. 8., Bart., Flixton Hall, Bungay.
1877 Apams, Frederick Charlstrom, F.Z.S., 50, Ashley-gardens, Victoria-
street, S.W.
1877 Apams, Herbert J., Roseneath, London-road, Enjield, N.
1902 ADKIN, Benaiah Whitley, Trenoweth, Hope-park, Bromley, Kent.
1885 ADKIN, Robert, Welijfield, Lingards-road, Lewisham, 8.E.
1904 Agar, E. A., La Haut, Dominica, B. W. Indies.
1904 AtpERson, Miss E. Maude, Park House, Worksop, Notts.
1899 AnpRews, Henry W., Shirley, Welling, S.O., Kent.
1901 Awnnine, William, 39, Lime Street, E.C.
eet)
1899 + ARRow, Gilbert J., 87, Union-grove, Clapham, S.W.; and British
Musewm (Natural History), Cromwell-road, S.W.
1886 Armorg, E. A., 48, High-street, King’s Lynn.
1850 t AvEBuRY, The Right Honble. Lord, D.C.L., F.R.S., F.LS., F.G.S.,
etc., High Elms, Farnborough, Kent.
1901 Bacor, Arthur W., 154 Lower Clapton-road, N.E.
1904 f BaGNALL, Richard 8., South Hylton, nr. Sunderland.
1903 Baupock, G. R., Oakburn Villa, Enfield Highway, Middlesex.
1886 Bankzs, Eustace R., M.A., Norden, Corfe Castle, Wareham.
1890 Barcuay, Francis H., F.G.S., The Warren, Cromer.
1886 Barcaauti, Marchese Piero, Piazza S. Maria, Palazzo Tempi No.1,
Florence, Italy.
1895 Barker, Cecil W., Rownham, Malvern, Natal, South Africa.
1887 Barker, H. W., 147, Gordon-road, Peckham, S.E.
1902 Barraup, Philip J., Bushey Heath, Watford.
1894 ¢ Bateson, William, M.A., F.R.S., Fellow of St. John’s College,
Cambridge, Merton House, Grantchester, Cambridge.
1904 Baynn, Arthur F., Gerencia, Ferro Carril del Sud, Plaza Constitu-
tion, Buenos Ayres.
1896 | BEarg, Prof. T. Hudson, B.Sc., F.R.S.E., 10 Regent Terrace,
Edinburgh.
1851*}Braumont, Alfred, Gosfield, Halstead, Essex.
1905 Brprorp, The Duke of, K.G., Pres. Z.S., etc., Woburn Abbey, Beds.
1899 BEDWELL, Ernest C., Elmlea, Clevedon-roud, Norbiton, Surrey.
1903 Brni-Maruey, H. W., c/o Messrs. Chiazzari and Co., P.O. Box 3,
Point-street, Natal.
1904 Brnersson, Simon, Ph.D., Lecturer, University of Lund, Sweden ;
Curator, Entomological Collection of the University.
1897 Brnnert, W. H., 15, Wellington-place, Hastings.
1885 Breraune-Baker, George T., F.L.S., 19, Clarendon-road, Edgbaston,
Birmingham.
1895 Bevan, Lieutenant H.G.R.,R.N., Fairfield, Weymouth.
1880 Brieneu, George Carter, The Ferns, Homepark-road, Saltash.
1895 Briyenam, Lieut.-Col. Charles T., F.Z.S., Bombay Staff Corps,
6 Gwendwr-road, West Kensington, S.W.
1891 Buaser, W. H., F.L.S., 12, Great Castle-street, Regent-street, W.
1904 Brack, James E., Nethercroft, Peebles.
1894 + BuackBURNE-Mazg, W. P., Shaw House, Newbury.
1904 Buarr, Kenneth G., 23, West Hill, Highgate, N.
1889 BLanpForD, Walter F. H., M.A., F.Z.S., 12, Arwndel Gardens,
Ladbroke-grove, W.
1885 Buarawayt, Lieut.-Col. Linley, F.L.S., HagleHouse, Batheaston,
Bath.
1904 Briss, Maurice Frederick, Coningsburgh, Montpelier-road, Ealing, W.
1886 B.oomriznp, The Rev. Edwin Newson, M.A., G'uestling Rectory,
Hastings.
Cet
1903 Bogus, W. A., Wilts and Dorset Bank, Salisbury.
1891 Booru, George A., Fern Hill, Grange-over-Sands, Carnforth.
1876 * Borre, Alfred Prudhomme de, Villa la Fauvette, Petit Saconnea,
Geneva.
1875 Borrer, Wm., F.G.S., Pakyns Manor House, Hurstpierpoint,
Hassocks, R.S.O., Sussex.
1902 Bostock, E. D., Holly House, Stone, Staffs.
1904 Bourceors, Jules, Ste. Marie-aux-Mines, Markirch, Germany.
1892 Bouskett, Frank, Market Bosworth, Nuneaton.
1888 Bower, Benjamin A., Langley, Willow Grove, Chislehurst.
1894 + Bows, E. Augustus, M.A., Myddelton House, Waltham Cross.
1852 + Boyp, Thos., Woodvale Lodge, South Norwood Hill, S.E.
1893 Brapant, Edouard, Chateaw de Morenchies, par Cambrai (Nord),
France.
1905 Bracken, Charles W., B.A., 18, Whiteford-road, Mannamead,
Plymouth.
1904 BrincEeman, The Lieut. Hon. Richard O. B., R.N., Weston Park,
Shifnal, Salop, and H.M.S. ‘ Clio,” China Station.
1877 Brices, Charles Adolphus, Rock House, Lynmouth, R.S.0., N. Devon.
1870 Briees, Thomas Henry, M.A., Rock House, Lynmouth, R.S.0., N.
Devon.
1894 Briaut, Percy M., Chunar, Lansdowne-road, Bournemouth.
1897 Bricutwen, Mrs. E., The Grove, Great Stanmore.
1890 Bristowg, B. A., The Cottage, Stoke D’ Abernon, Cobham, Surrey.
1878 Broun, Capt. Thomas, Drury, Auckland, New Zealand,
1902 Brovuauton, Captain T. Delves, R.E., Alderney.
1904. Brown, egay H., Castlefield Tower, ae Fife, N.B.
1886 Brows, John, 5, Tainaia Parade, Ce ids idge.
1892 Browns, Lieut.-Colonel Clement Alfred Righy, BE. ,Caleutta,Fndia.
1898 + Bucnan-Hepsurn, Sir Archibald, Bart., J.P., D.L., Smeuton-
Hepburn, Prestonkirk.
1883 * Buckron, George Bowdler, F.R.S., F.L.S., Weycombe, Haslemere,
S.0., Surrey.
1902 Buuier, Arthur Percival, Wellington, New Zealand.
1896 + Burr, Malcolm, B.A., F.L.S., F.Z.S., F.G.S., Royal Societies Club,
St. James's, S.W., and 23, Blomfield Court, Maida Vale, W.
1868 + Butter, Arthur G., Ph.D., F.L.S., F.Z.8., The Lilies, Penge-road,
Beckenham.
1883 Burier, Edward Albert, B.A., B.Sc., 53, Tollington Park, N.
1902 Burier, William E., Hayling House, Oxford-road, Reading.
1905 BurrerFiIELD, Jas. E., B.Sc., Comrie, Eglinton Hill, Plumstead.
1904 Byart, Horace A., B.A., Berbera (vid Aden), Somaliland Protectorate.
1886 CaLvERT, Wm. Bartlett, Liceo de Quillota, Quillota, Chili.
1902 Cameron, Malcolm, M.B., R.N., R.N. Hospital, Chatham.
1885 CampBeLL, Francis Maule, F.LS., F.ZS., &., Brynllwydwyn,
Machynlleth, Montgomeryshire.
1898
1880
1889
1890
1894
1892
1895
1898
1868
1890
1895
1900
1900
1903
C0 aga '8
CanpeEzE, Léon, 64, Rue de Ouest, Liége.
CansDALE, W. D., Sunny Bank, South Norwood, 8.E.
Cant, A.,69, Hampden-road, Hornsey, N.; and c/o Fredk. DuCane
Godman, Esq., F.R.S., 10, Chandos-street, Cavendish-square, W.
Capper, Samuel James (President of the Lancashire and Cheshire
Entomological Society), Huyton Park, Liverpool.
Caraccioto, H., H.M. Customs, Port of Spain, Trinidad, British
West Indies.
Carpenter, The Honble. Mrs. Beatrice, 22, Grosvenor-road, 8. W.
CaRPENTER, G. H., B.Se., Royal College of Science, Dublin.
CarPENTER, J. H., Redcot, Belmont-road, Leatherhead.
CaRRINGTON, Charles, Hailey Hall, Hertford.
Carter, George Wm., M.A., F.L.S., Eecleshall Castle, Staffordshire.
CartER, Sir Gilbert, K.C.M.G., 43, Charing Cross, W.C.; and
Government House, Nassau, Bahamas.
Carter, J. W., 25, Gilenholme-road, Manningham, Bradford.
Cassau, R. T., M.R.C.8., Ballaugh,:Isle of Man.
Cattle, John Rowland, Nettleton Manor, Caistor, Lincolnshire.
1889 + Caves, Charles J. P., Ditcham Park, Petersfield.
1900
1871
1891
1902
1890
1897
1898
1902
CHAMBERLAIN, Neville, Highbury, Moor Green, Birmingham.
CHAMPION, George C., F.Z.8., Liprartan, Heatherside, Horsell,
Woking ; and 10, Chandos-street, Cavendish-square, W.
CHAPMAN, Thomas Algernon, M.D., F.Z.8., VicE-PRESIDENT, Betula,
Reigate.
CHARNLEY, James Roland, The Avenue, Moor Park, Preston,
Lancashire.
CHATTERTON, Frederick J.S.,5, Camden Studios, Camden-street, N.W.
CHAWNER, Miss Ethel F., Forest Bank, Lyndhurst, R.S.O., Hants.
CHawner, Lawrence C., Forest Bank, Lyndhurst, R.S.O., Hants.
CHEESMAN, E. M., c/o J. Garson, 63, Railway-street, Durban, Natal.
1891 + Currry, Arthur John, M.A., 27, Hereford-square, 8.W.; and Hunt-
1905
1889
ingfield, Faversham, Kent.
CHOPARD, Lucien, 98, Bd. St. Germains, Paris.
Curisty, William M., M.A., F.L.S., Watergate, Emsworth.
1886 + CLARK, John Adolphus, 57, Weston Park, Crouch End, N.
1867
1886
1891
1904
1873
1899
1901
1900
1892
CuaRKE, Alex. Henry, 109, Warwick-road, Earl’s Court, 8.W.
CuaRrkE, Charles Baron, M.A., F.R.S., F.LS., F.GS., 18, Kew
Gardens-road, Kew, 8.W.
CLARKE, Henry Shortridge, 2, Osborne-terrace, Douglas, Isle
of Man.
Cockayne, Edward A., 30, Bedford Court Mansions, W.C.
Cotez, William, F.L.S., Springfield, Buckhurst Hill, Essex.
CoLttin, James E., Sussex Lodge, Newmarket.
Connotp, Edward, 7, Magdalen Terrace, St. Leonards-on-Sea.
Corton, Dr. John, 126, Prescot-road, St. Helens.
Cowan, Thomas William, F.LS., F.G.S., F.R.M.S., 8, Henrietta-
street, Covent Garden, W.C.
1886
1867
1895
1888
1890
(| xili )
CowetL, Peter (Librarian of the Liverpool Free Public Library),
William Brown-street, Liverpool.
Cox, Herbert Ed., c/o Mrs. Eve, 125, Harley-street, W.
CRABTREE, Benjamin Hill, The Oaklands, Levenshulme, Manchester.
Creeor, J. P., Tredinick, Mayow-road, Sydenham, 8.E.
Crews, Sir Vauncey Harpur, Bart., Calke Abbey, Derbyshire.
1880 + Crisp, Frank, LL.B., B.A., J.P., F.L.8., 17, Throgmorton-avenue,
1902
1901
1873
1900
1886
1905
1903
1898
1905
1875
1887
1895
1905
1903
1891
1885
E.C., and Friar Park, Henley-on-Thames.
CruTtwELL, The Rev. Canon Charles Thomas, M.A., Hwelme
Rectory, Wallingford.
Davp, Edward Martin, Bismarckstrasse 1, Charlottenburg, Germany.
Daz, C. W., Glanville’s Wootton, Sherborne, Dorset.
Daueisu, Andrew Adie, 21, Prince’s-street, Glasgow.
Dannatt, Walter, Donnington, 75, Vanbrugh Park, Blackheath, 8.E.
Davinson, James D., 32, Drumsheugh Gardens, Edinburgh.
Day, F. H., 12, Goodwin-terrace, Carlisle.
Day, G. O., Parr’s Bank-house, Knutsford.
Dewar, W. R., Government Entomologist, Orange River Colony.
Distant, Wm. Lucas, Steine House, Selhurst-road, South Norwood,S.E.
Dixey, Frederick Augustus, M.A., M.D., Fellow and Bursar of
Wadham College, Vicz-PResIDENT, Wadham College, Oxford.
Dosson, H. T., Ivy House, Acacia Grove, New Malden, S.O., Surrey.
Dopp, F. P., Kuranda, vid Cairns, North Queensland.
Doutmay, J. C., Hove House, Newton-grove, Bedford-park, W.
DonistHorPE, Horace St. John K., F.Z.S., 58, Kensington-mansions,
South Kensington, S.W.
Donovan, Major Charles, M.D., R.A.M.C., c/o Messrs. P, Macfadyen
& Co., Winchester House, Old Broad-street, E.C.
1845 * Doveras, John Wm., 61, Craven Park, Harlesden, N.W.
1899
1884
1867
1900
1894
1883
1890
1865
1904
1902
1886
Drewitt, Frederick G. Dawtrey, M.A., M.D., F.R.C.P., F.Z.8., 14,
Palace Gardens-terrace, Kensington, W.
Druce, Hamilton H. C. J., F.Z.S., 43, Circus-road, St. John’s
Wood, N.W.
Drucs, Herbert, F.L.S., F.Z.S., 43, Circus-road, St. John’s Wood, N.W.
Drury, W. D., Rocquaine, West Hill Park, Woking.
Dupeexon, G. C., The Imperial Institute, South Kensington.
Durrant, John Hartley, The Cottage, Merton Hall, Thetford.
Eastwoop, John Edmund, Enton Lodge, Witley, Godalming.
Eaton, The Rev. Alfred Edwin, M.A., Pentlands, Mill-road, West
Worthing, Sussex.
EckrorpD, George, F.Z.S., c/o Sir Morgan Tuite, Bart., Kilruane,
Nenagh, co. Tipperary, Ireland.
EDELSTEN, Hubert M., The Elms, Forty Hill, Enfield, Middlesex.
EpwWarDs, James, Colesborne, Cheltenham.
1884
1900
1900
1886
1903
1878
1886
1903
1899
1890
1900
1861
1886
1889
1898
1878
1900
1874
1886
1905
1900
( xiv )
Epwarps, Stanley, F.L.S8., F.Z.8., 15, St. Germans-place, Black-
heath, S.E. j
Exxiort, E. A., 16, Belsize Grove, Hampstead, N.W.
Exuis, H. Willoughby, Knowle, Birmingham.
Exuis, John W., M.B., L.R.C.P., 18, Rodney-street, Liverpool.
ELTrRInGHAM, Harry, M.A., F.Z.S., Eastgarth, Westoe, South Shields.
Etwes, Henry John, J.P., F.RS., F.LS. F.Z.S., Colesborne,
Cheltenham.
Enock, Frederick, F.L.8., 42, Salisbury-road, Bexley, Kent.
ETHERIDGE, Robert, Curator, Australian Musewn, Sydney, N.S.W.
FarmporouaH, Perey W., F.Z.S., Lower Edmonton, N.
Farn, Albert Brydges, Mount Nod, Greenhithe, Kent ; and Medical
Department, Local Government Board, Whitehall, 8.W.
FreitrHam, H. L. L., P. O. Box, 46, Johannesburg, Transvaal.
Fenn, Charles, Hversden House, Burnt Ash Hill, Lee, 8.E,
Fenwick, Nicolas Percival, The Gables, New-road, Esher.
FERNALD, Prof. C. H., Amherst, Mass., U.S.A.
Fiter, F. E., 122, Stockwell Park-road, Brixton, S.W.
Finzi, John A., 53, Hamilton-terrace, N.W.
Firta, J. Digby, F.L.8., Boys’ Modern School, Leeds.
Fircn, Edward A., F.L.S., Brick House, Maldon.
Frrcu, Frederick, Hadleigh House, Highbury New Park, N.
Freer, Wilfred James, Imatra, King’s Road, Bowrnemouth.
Fremyne, The Rev. W. Westropp, M.A., Ooolfin, Portlaw, Co.
Waterford.
1898 + FietcueEr, T. Bainbrigge, R.N., H.M.S. “Sealark,” Special Service.
1883 + FiercHeER, William Holland B., M.A., Aldwick Manor, Bognor,
1905
1885
1900
1898
1880
1883
1896
1888
1903
1891
FLoERSHEIM, Cecil, 16, Kensington Court Mansions, S.W.
FoxkeEr, A. J. F., Zierikzee, Zeeland, Netherlands.
Foutxkss, P. Hedworth, B.Sc., Harper-Adams Agricultwral College,
Newport, Salop.
FountaIng, Miss Margaret, 7, Lansdowne-place East, Bath.
Fow.er, The Rey. Canon, D.Sc., M.A., F.L.S., Harley Vicarage,
near Reading.
FREEMAN, Francis Ford, Abbotsfield, Tavistock.
FREKE, Percy Evans, Southpoint, Limes-road, Folkestone.
Fremiin, H. Stuart, M.R.C.S., L.R.C.P., Mereworth, Maidstone.
Frencu, Charles, F.L.S., Government Entomologist, Victoria,
Australia.
FroHawk, F. W., Ashmount, Rayleigh.
1855 * Fry, Alexander, Thornhill-house, Dulwich Wood Park, Norwood, S.E.
1900
1884
1898
1904
Fryer, H. Fortescue, The Priory, Chatteris, Cambs.
Funtirer, The Rey. Alfred, M.A., The Lodge, 7, Sydenham-hill,
Sydenham, 8.E.
FuLuer, Claude, Government Entomologist, Pietermaritzburg, Natal.
FuRNIVAL, Thomas F., 63, Coleman-st., E.C.
( xv)
1887 GanHan, Charles Joseph, M.A., Whyola, Lonsdale-road, Bedford
Park, W.; and British Museum (Natural History), Cromwell-
road, S.W.
1892 Garpbs, Philip de la, R.N., Manor-house, Shaldon, nr. Teignmouth.
1890 Garpner, John, 6, Friars-gate, Hartlepool.
1901 GarpnerR, Willoughby, F.L.S., Deganwy, N. Wales.
1899 Gaywner, Francis, Oxshott, Surrey.
1899 GeLpaRt, William Martin, M.A., Trinity College, Oxford.
1902 GrinLanpERS, A. T., Park Cottage, Alrwick.
1904 GurLr1At, Francis, B.A., Forest Dene, Worth, Sussex.
1865 + Gopmayn, Frederick Du Cane, D.C.L., F.R.S., F.L.S., F.Z.S., South
Lodge, Lower Beeding, Horsham; 7, Carlos-place, Grosvenor-
square; and 10, Chandos-street, Cavendish-square, W.
1890 Goxpruwalrt, Oliver C., 5, Queen’s-road, South Norwood, S.E.
1886 + Goopricw, Captain Arthur Mainwaring, Lennox Lodge, Malvern
Link, Malvern.
1904 Goopwin, Edward, Canon Court, Wateringbury, Kent.
1898 Gorpon, J. G. McH., Corsemalzie, Whauphill, R.S.O., Wigtownshire.
1898 Gorpon, R. 8. G. McH., Corsemalzie, Whauphill, R.S.O., Wigtown-
shire.
1855 Goruam, The Rev. Henry Stephen, F.Z.S., Highcroft, Great Malvern.
1874 Goss, Herbert, F.L.S., The Avenue, Surbiton-hill, Surrey.
1891 + GrrEN, E. Ernest, Government Entomologist, Royal Botanic
Gardens, Peradeniya, Ceylon.
1894 Green, J. F., F.Z.S., West Lodge, Blackheath, S.E.
1850 GREENE, The Rev. Joseph, M.A., Rostrevor, Clifton, Bristol.
1898 GREENSHIELDS, Alexander, 38, Blenheim-gardens, Willesden, N.W.
1899 GREENWOOD, Edgar, 49, Melrose-avenue, Willesden Green, N.W.
1893 + GREENWooD, Henry Powys, F.L.S., Sandhill Lodge, Fordingbridge,
Salisbury.
1888 GrirFiTHs, G. C., F.Z.S., 43, Caledonian-place, Clifton, Bristol.
1894 GrimsHaw, Percy H., Royal Scottish Museum, Edinburgh.
1905 Grist, Charles J., Apsley, Banstead, Surrey.
1900 Groom, Prof. Percy, M.A., F.L.S., Royal Indian Engineering
College, Cooper’s Hill, Staines.
1869 Gross-SmirH, Henley, J.P., B.A., F.Z.S.,5, Bryanston-square, Hyde
Park, W.
1899 GuNnNING, Montague, Narborough, Leicester.
1897 Haauz, Henry, 2, Woodland Avenue, Glen Ridge, N.J., U.S.A.
1890 + Hatt, A. E., Norbury, Pitsmoor, Sheffield.
1885 Hatt, Thomas William, Stanhope, The Crescent, Croydon.
1898 Hamuiyn-Harris, R., D.Sc., F.Z.S., F.R.M.S., Toowoomba Grammar
School, Queensland, Australia.
1891 Hampson, Sir George Francis, Bart., B.A., F.Z.8., 62, Stanhope-
gardens, 8.W.
1891 Hansory, Frederick J., F.L.S., Stainforth House, Upper Clapton, N.E.
(xt) )
1905 + Hancock, Joseph L., 3757, Indiana Avenue, Chicago, U.S.A.
1903 Hars, E. J.,. Dunham, Boscombe, Hants.
1904 Harris, Edward, St. Conan’s, Chingford, Essex.
1897 + Harrison, Albert, F.L.S., F.C.S., Delamere, Grove-road, South.
Woodford, Essex.
1889 Harrison, John, 7, Gawber-road, Barnsley.
1905 Harrison, T. W. H., B.Se., The Avenue, Birtley, R.S.O. Durham.
1881 Henry, George, Ivy Bank, 112, London-road, St. Leonards-on-Sea.
1903 Herron, William, W.B.C. Apiary, Old Bedford-road, Luton, Beds.
1898 Heron, Francis A., B.A., British Museum (Natural History),
Cromwell-road, 8.W.
1888 Huicas, Martin Stanger, F.C.S., F.G.8S., Mine Office, Venterskroon,
Transvaal.
1876 + Hittman, Thomas Stanton, Eastgate-street, Lewes.
1888 Hopson, The Rev. J. H., B.A., B.D., Rhyddington, Clifton Drive,.
Lytham.
1902 Hotz, R. 8., Indian Forest Service, c/o Messrs. King, King and
Co., Bombay.
1887 Houtanp, The Rev. W. J., D.D., Ph.D., 5th Avenue, Pittsburg,
Penn., U.S.A.
1898 Hoxman-Hount, C. B., Talawakelle, Ceylon.
1901 Horson, Montagu F., L.D.S., R.C.S.Eng., F.L.S., 30, Thurlow-road,
Rosslyn Hill, N.W.
1897 Horne, Arthur, 60, Gladstone-place, Aberdeen.
1876 ¢ Horniman, Fredk. John, M.P., FLS, F.ZS., &., c/o The
Librarian, Horniman Museum, Forest Hill, 8.E.
1903 Hovueuron, J. T., 1, Portland-place, Worksop.
1900 Howss, George H., Box 180, Dunedin, New Zealand.
1865 + Hupp, A. E., Clinton, Pembroke-road, Clifton, Bristol.
1888 Hupson, George Vernon, The Post Office, Wellington, New Zealand.
1902 * Huron, Captain Frederick W., F.R.S., Director of the Canterbury
Museum, Christchurch, New Zealand.
1897 Imaas, Selwyn, M.A., 20, Fitzroy-street, Fitzroy-square, W.
1893 *Irpy, Lieutenant-Colonel Leonard Howard Loyd, F.L.S., F.Z.S.,
14, Cornwall-terrace, Regent's Park, N.W.
1891 IsapELL, The Rev. John, Sunnycroft, St. Sennen, R.S.0., Cornwall.
1886 Jacosy, Martin, 1, The Mansions, Hillfield-road, West Hampstead,
N.W.
1869 JANSON, Oliver E., Cestria, Claremont-road, Highgate, N.; and 44,
Great Russell-street, Bloomsbury, W.C.
1898 Janson, Oliver J., Cestria, Claremont-road, Highgate, N.
1886 JENNER, James Herbert Augustus, 209, School Hill, Lewes.
1899 Jenninas, F. B., 152, Silver-street, Upper Edmonton, N.
1886 Joun, Evan, Llantrisant, R.S.O., Glamorganshire.
Cavin)
1889 Jounson, The Rev. W. F., M.A., Acton Rectory, Poyntz Pass,
Co. Armagh.
1888 Jones, Albert H., Treasurer, Shrublands, Eltham, Kent.
1894 } Jorpan, Dr. K., The Museum, Tring.
1902 Joy, Norman H., M.R.C.S., L.R.C.P., Bradfield, Reading.
1884 Kanz, W. F. de Vismes, M.A., M.R.I.A., Drumleaske House,
Monaghan.
1884 Kappet, A. W., F.L.S., Hilden, 18, Sutton Court-road, Chiswick, W.
1876 F Kay, John Dunning, Leeds,
1896 + Kayz, William James, Caracas, Ditton Hill, Surbiton.
1902 Kemp, Stanley W., Trinity College, Dublin.
1890 Kenrick, G. H., Whetstone, Somerset-road, Edgbaston, Birmingham,
1904 KersHaw, G. Bertram, Ingleside, West Wickham, Kent.
1898 KersHaw, J. A., Morton Banks, Lewisham-road, Windsor,
Melbourne, Victoria,
1901 Kersnaw, John C. W., Macao, China.
1900 Krys, James H., Morwell, Freedom-villas, Lipson-road, Plymouth.
1889 Kine, J. J. F. X., Lecturer on Economic Entomology at the West of
Scotland Agricultural College, 1, Athole Gardens-terrace, Kelvin-
side, Glasgow.
1861 Kirsy, William F., F.L.S., Hilden, 18, Sutton Court-road, Chiswick, W.
1893 KirKaupy, George Willis, Board of Agriculture, Division of
Entomology, Honolulu, Hawaii.
1905 Kuironen, Vernon Parry, The Priory, Watford.
1889 KtapAuEK, Professor Franz, Karlin 263, Prague, Bohemia.
1887 + Kuetn, Sydney T., F.L.S.,F.R.A.S., Hatherlow, Raglan-road, Reigate.
1876 Kraarz, Dr. G., 28, Link-strasse, Berlin.
1901 Lanz, E. W., Parkholme, 40, Fletching-road, Clapton, N.E.
1868 Lana, Colonel A. M., R.E., Box Grove Lodge, Guildford.
1900 Lane, The Rev. H. C., M.D., All Saints’ Vicarage, Southend-on-Sea.
1901 Laruy, Percy I., Penton House, Cheshunt.
1895 Latrer, Oswald H., M.A., Charterhouse, Godalming.
1899 Lexa, Arthur M., Government Entomologist, Hobart, Tasmania.
1900 Lerroy, H. Maxwell, B.A., Muzaffarpur, Behar, Bengal.
1901 Lexieu, George F., Woodside, off Umbilo-road, Congella, nr. Durban,
Natal.
1883 Lemany, Fredk. Charles, Blackfriars House, Plymouth.
1898 Lrraprivgr, Ambrose G., Nordrach-on-Dee, Banchory, N.B.;
Guards Club, Pall Mall, S.W.
1903 ¢ Levert, The Rev. Thomas Prinsep, Frenchyate, Richmond, Yorks,
1898 Lewis, E. J., F.L.S., 4, Clements Inn, W.C,
1876 Luwis, George, F.L.S., 87, Frant-road, Tunbridge Wells.
1902 Lewis, J. H., Ophir, Otago, New Zealand.
1892 Licurroor, R. M., Bree-st., Cape Town, Cape of Good Hope.
1903 Lirrter, Frank M., Althorne, High-street, Launceston, Tasmania.
b
(peu)
1865+ Lurwetyn, Sir John Talbot Dillwyn, Bart, M.A, F.LS.,,
Penllergare, Swansea,
1881 + Luoyp, Alfred, F.C.S., Zhe Dome, Bognor.
1885 + Luoyp, Robert Wylie, St. Cuthberts, Thurleigh-road, Balham, 8.W.
1903 LorrHouss, Thomas Ashton, The Croft, Linthorpe, Middlesbrough.
1904 + Lonestarr, George Blundell, M.D., Wighlands, Putney Heath, S.W.
1899 Lounspury, Charles P., B.Se., Government Entomologist, Cape
Toun, S. Africa.
1894 Lows, The Rev. Frank E., M.A., St. Stephen’s Vicarage, Guernsey.
1893 Lower, Oswald B., St. Oswalds, Bartley Crescent, Wayville, South
Australia.
1901 Lower, Rupert 8., Davonport-terrace, Wayville, South Australia.
1898 Lucas, William John, B.A., 28, Knight's Park, Kingston-on- Thames.
1904 Lurr, W. A., La Chawmicre, Brock-road, Guernsey.
1880 Lupron, Henry, Lyndhurst, North Grange-road, Headingley, Leeds.
1903 Lyext, G., Junr., Gisborne, Victoria, Australia.
1901 Lyman, Henry H., M.A., F.R.GS., 74, McTavish-street, Montreal,
Canada.
1902 Macponatp, George B. Douglas, M.B.
1887 M‘Douaatt, James Thomas, Dunolly, Morden-road, Blackheath, 8.E.
1888 Mackinnon, P. W., Lynndale, Mussoorie, N.W.P., India.
1900 MacKkwoop, The Hon. F. M., M.L.C., Colombo, Ceylon.
i898 Mappison, T., South Bailey, Durham.
1899 + Mary, Hugh, B.Sc., Almondale, Buckingham-road, South Woodford,
N.E.
1905 Matty, Charles Wm., M.Sc., Government Entomologist, Grahams-
town, Cape Colony.
1887 Manpers, Lieut.-Colonel Neville, R.A.M.C., Curepipe, Mauritius.
1892 Mawnspring8, William, 27, Elmbank-road, Sefton-park, Liverpool.
1894 ¢ MarsHALL, Alick, Auchinraith, Bexley, S.O., Kent.
1895 MarsHaut, G. A. K., P.O. Box 149, Salisbury, Mashonaland,
S. Africa.
1896 MarsHatt, P., M.A., B.Sc. F.G.S., University School of Mines,
Dunedin, New Zealand.
1856 | MarRSHALL, William, Auchinraith, Bexley, S.O., Kent.
1897 Martineau, Alfred H., Solihull, Birmingham.
1895 Massry, Herbert, Zvy-Lea, Burnage, Withington, Manchester.
1865 Maruew, Gervase F., F.L.S., Paymaster-in-chief, R.N., Lee House,
Dovercourt, Harwich.
1887 Marruews, Coryndon, Stentaway, Plymstock, Plymouth.
1899 May, Harry Haden, 12, Windsor Terrace, Plymouth.
1904 Mrapr-Waxpo, Geoffrey, Stonewall Park, Edenbridge, Kent.
1872 + Metpoua, Professor Raphael, F.R.S., F.C.S., 6, Brunswick-
square, W.C.
1885 Metvit, James Cosmo, M.A., F.L.8., Meole Brace Hall, Shrewsbury.
1887 MeERRIFIELD, Frederic, PRestpENT, 24, Vernon-terrace, Brighton.
——
WES)
1905 Merry, Rey. W. Mansell, M.A., St. Michael's, Oxford.
1888 Meryrr-Darcis, G., c/o Sogin and Meyer, Wohlen, Switzerland.
1880 Meyrick, Edward, B.A., F.Z.S., F.RS., Thornhanger, Marlborough.
1894 Mratt, Professor Louis Compton, F.R.S., 1, Richmond Mownt,
Headingley, Leeds.
1883 Minus, W. H., The New Club, Caleutta.
1905 Mrrrorp, Robt. Sidney, C.B., 35, Redcliffe Square, S.W.
1896 Moszrty, J. C., M.A., Woodlands, Bassett, Southampton.
1879 Mownrerro, Dr. Antonio Augusto de Carvalho, 70, Rua do Alecrinar,
Lisbon.
1902 Monreomery, Arthur Meadows, 34, Shaliman Gardens, Pembridge-
road, North Acton, N.
1853 Moorz, Frederic, D.Sce., ATS:, F.2.8..17, Mayle-road, Penge, 8.E.
1899 Moors, Harry, 12, Lower-road, Rotherhithe.
1886 Morean, A. C. F., F.LS., 135, Oakwood-court, Kensington, W.
1889 ¢ Morice, The Rey. F. D., M.A., Fellow of Queen’s College, Oxford,
Brunswick, Mount Hermon, W. oking.
1895 | Morey, Claude, The Hill H. ouse, Monk’s Soham, Suffolk.
1893 Morvron, Kenneth J -» 13, Blackford-roud, Edinburgh.
1900 Moser, Julius, 60, Bulow-strasse, Berlin.
1901 + Murr, Frederick, EZ. and S.A frican Telegraph Co., Mozambique.
1869 | Mitier, Albert, F. R.G.S., c/o Herr A. Miiller-Mechel, Gren-
zacherstrasse, 60, Basle, Switzerland.
1904 Mumrorp, Frank S., 10, Mountfield Gardens, Tunbridge Wells.
1872 | Murray, Lieut.-Col. H., 43, Cromwell Houses, Cromwell-road, S.W.
1903 Nuavs, S. A., B.A., Mill Green Park, Ingatestone.
1896 Nusuawm, Robert, Utrecht House, Queen’s-road, Clapham Parl: S.W.
1889 Nevinson, Basil George, M.A., F As as eR edworth-square,
Chelsea, S.W.
1901 Nevrinson, E. G. Tas. Dy Bentinck-terruce, Regent’s Park, N.W.
1890 NewstxEap, R., Johnston Tropical Laboratory, University, Liverpool.
1900 NicHont, Mrs. M. Dela B., Merthyr Mwor, Bridgend, Glamorganshire.
1904 Nicuotson, W. A., 36, Promenade, Portobello, N.B.
1886 NicHoxson, William E., School Hill, Lewes,
1893 Nonrrigp, A. F., Rakonitz, Bohemia.
1886 Norris, Herbert E., 15, Market-place, Cirencester.
1878 Norripax, Thomas, Ashford, Kent.
1895 Nursg, Major C. G., Timworth Hall, Bury St. Edmunds.
1869 OxseErRtHiir, Charles, Rennes (Ille et Vilaine), France,
1877 OxseErtHir, René, Rennes (Ille et Vilaine), France.
1893 + Octx, Bertram S., Steeple Aston, Oxfordshire.
1893 Ontver, John Baxter, 22, Ranelagh Villas, Hove, Brighton.
1873 Ottvirr, Ernest, Ramillons, prés Moulins (Allier), France.
1895 Pagan, Herbert E., Bertrose, Gellatly-road, St. Catherine's Park, S.F,
Gis y)
1898 PauusER, H. G., Holmwood, Addlestone, Surrey.
1901 * PeAL, Henry Woolner, Indian Musewm, Calcutta.
1883 PsRINGUEY, Louis, South African Musewm, Cape Town, South Africa.
1903 + Perkins, R. C. L., B.A., Board of Agriculture, Division of Ento-
mology, Honolulu, Hawaii.
1879 PeERKINs, Vincent Robt., Wotton-under-Edge.
1900 Puiuips, The Rev. W. J. Leigh, The Cottage, Parkwood-road,
Tavistock.
1897 Puitirps, Hubert C., M.R.C.S.,M. and L.8.A., 262, Gloucester-terrace,
Hyde-park, W.
1903 ¢ Putuuirs, Montagu A., F.R.G.S., F.Z.S., 22, Petherton-road, High-
bury, New Park, N.
1901 Picker, C. P., 99, Dawlish-road, Leyton, Essex.
1891 PreRcr, Frank Nelson, 1, The Elms, Dingle, Liverpool.
1901 Pirrarp, Albert, Felden, Boxmoor, Hemel Hempstead.
1903 PincuErR, Colonel Jesse George, I.M.S., F.R.C.8., 133, Gloucester-
road, Kensington, W.
1885 Pout, J. R. H. Neerwort van de, Driebergen, Netherlands.
1870 + Porrirr, Geo. T., F.L.S., Mayfield, Edgerton, Huddersfield.
1884 + Pouuton, Professor Edward B., D.Sc., M.A., F.R.S., F.L.S., F.G.S.,
F.Z.S., Hope Professor of Zoology in the University of Oxford,
Vicr-PRESIDENT, Wykeham House, Banbury-road, Oxford.
1905 Powett, Harold, 7, Rue Mirelle, Hyéres (Var), France.
1878 Pricr, David, 48, West-street, Horsham.
1904 Prisks, Richard A.R., 66, Chaucer-road, Acton.
1893 Prout, Louis Beethoven, 246, Richmond-road, Dalston, N.E.
1900 Rarnsow, William J., The Australian Musewm, Sydney, N.S.W.
1874 Reep, E. C., Director del Museo de Concepcion, Concepcion, Chile.
1900 Rerp, Percy Charles, Feering Bury, Kelvedon, Essex.
1893 Retp, Captain Savile G., late R.E., The Elms, Yalding, Maidstone.
1898 Reuron, R. H., c/o Perkins and Co., Ltd., Brisbane, Queensland.
1890 ReENpLESHAM, The Right Honble. Lord, Rendlesham Hall, Wood-
bridge.
1898 Reuter, Professor Enzio, Helsingfors, Finland.
1894 Riprina, William Steer, B.A., M.D., Buckerell Lodge, Honiton.
1853 Ripon, The Most Honble. the Marquis of, K.G.,'D.C.L., F.R.S., F.LS.,
etc., 9, Chelsea Embankment, S.W.
1905 Rosrnson, Herbert C., Curator of State Museum, Kuala Lumpur,
Selangor.
1892 Rosrnson, Sydney C., Goldsmiths’ Hall, E.C.
1869 + Ropinson-Doveias, William Douglas, MA, F.LS., F.R.GS.,
Orchardton, Castle Douglas.
1890 Rogson, John Emmerson, 15, Northgate, Hartlepool.
1886 Ross, Arthur J., 15, Boxwell-road, Berkhamstead.
1868 RorHney, George Alexander James, Pembury, Tudor-road, Upper
Norwood, 8.E.
ei, |)
1894 + Roruscaitp, The Honble. Nathaniel Charles, M.A., F.LS., F.Z.S.,
148, Piccadilly, W.; and Tring Park, Tring.
1888 + Roruscainp, The Honble. Walter, D.Sc., M.P., F.L.8., F.Z.8., 148,
1890
1887
1903
1898
1892
1899
1905
Piccadilly, W.; and Tring Park, Tring.
Rovutieper, G. B., Tarn Lodge, Heads Nook, Carlisle.
Rowtanp-Brown, Henry, M.A., Secretary, Oxhey-grove, Harrow
Weald.
Rowianps, Osbert William, Lickey Grange, nr. Bromsgrove.
RussEtn, A., The Limes, Southend, Catford, 5.E.
RussExt, S. G. C., 19, Lombard-street, E.C.
Ryxes, William E., B.A., 14, Arthur-street, Nottingham.
Sr.-Quintin, W. H., Scampton Hall, Rillington, York.
1865 + SaunpeERs, Edward, F.R.S.,F.L.S., St Ann’s, Mount Hermon, Woking.
1861 + SaunpeRs, G. S., F.L.S., 20, Dents-road, Wandsworth Common,
1886
1901
1881
1864
1862
1902
1883
1905
1901
SW.
SaunpErs, Prof. Wm., Central Experimental Farm, Ottawa, Canada.
Scuauvs, W., F.Z.S., Trentham House, Twickenham. |
Sconuick, A. J., 8, Mayfield-road, Merton Park, Wimbledon.
Semper, George, Klopstoch-strasse 23, Altona, Hibe, Germany.
SHarp, David, M.A., M.B., F.BS., F.LS., F.Z.8., Lawnside,
Brockenhurst, Hants. ; and University Museum of Zoology and
Comparative Anatomy, Cambridge.
SHarp, W. E., 9, Queen’s-road, South Norwood, 8.E.
Suaw, A. Eland, M.R.C.S., Overdale, Laxey, Isle of Man.
SHELDON, W. George, Youlgreave, South Croydon.
SHELFORD, R., M.A., C.M.Z.S., University Museum (Hope Depart-
ment), Oxford, Hythe, Kent.
1883 + SHELLEY, Capt. George Ernest, F.G.S., F.Z.S., 39, Egerton-gardens,
1900 + SHEPHEARD-Watwyy, H. W., M.A., Dalwhinnie, Kenley, Surrey.
1887
1904
1901
1902
1904
1902
1901
1901
1895
1898
1885
1889
Stcu, Alfred, Corney House, Chiswick, W.
Srmmonps, Hubert W., 17, Awrora-terrace, Wellington, New
Zealand.
SKERTCHLY, Ethelbert Forbes, c/o ‘ Penang Gazette,’ Penang, Straits
Settlements.
SLaADEN, Frederick William Lambart, 2, Sydney-road, Walmer,
Deal.
Supper, Rev. T. J. R. A., M.A., Tivetshall Rectory, Norwich.
SLOPER, Gerard Orby, Westrop House, Highworth, Wilts.
SmiruH, Arthur, 16, Edward-street, Grimsby.
SmirH, W. G., Mount Side, Bushey Park, Bristol.
Smita, W. W., Ashburton, Canterbury, New Zealand.
Sopp, Erasmus John Burgess, F.R.Met.S., 104, Liverpool-road,
Birkdale, Lancashire.
Sourn, Richard, 96, Drakefield-road, Upper Tooting, S.W.
STANDEN, Richard S., F.L.S., Townlands, Lindfield, Sussex.
( xxii )
1898 Srarms, C. L. B., M.R.C.S., L.R.C.P., The Infirmary, Wandsworth,
S.W.
1890 Stearns, A. E., 99, Gloucester-terrace, Hyde-park, W.
1897 Srepsine, E. P., Indian Forest Service, c/o King, Hamilton and
Co., Calcutta.
1898 Sressrne, Henry, 6, The Chase, Warley Mount, Brentwood, Essex.
1889 Srraton, C. R., F.R.C.S., West Lodge, Wilton, Salisbury.
1896 Srrickuanp, T. A. Gerald, Darlingworth House, Cirencester.
1900 Srupp, E. A. C., Kerremens, B.C.
1895 Supp, E. F., M.A., B.C.L., Oxton, Exeter.
1903 Swate, Harold, M.B., Arawa House, Rotorua, New Zealand.
1882 Swawnzy, Francis, Stanley House, Granville-road, Sevenoaks.
1884 SwiInHoE, Colonel Charles, M.A., F.L.S., F.Z.S., 42, Campden House
Court, Campden Hill, W.
1894 SwinHok, Ernest, 6, Gunterstone-road, Kensington, W.
1876 Swinton, A. H., c/o Mrs. Callander, Vineyard, Totnes.
1893 Taynor, Charles B., 22, Rae-street, Rae Town, Kingston, Jamaica.
1892 Taytor, The Rev. George W., F.R.S. (Canada), St. Mutthew’s
Rectory, Wellington, British Columbia.
1903 Taytor, Thomas Harold, M.A., Yorkshire College, Leeds,
1901 THompson, Matthew Lawson, 2, Thorncliff Villas, Saltburn-by-the-Sea.
1892 THorNuEy, The Rey. A., M.A., F.L.S., The Gables, Hacknall-road,
Nottingham.
1897 Tomuin, B., M.A., Mathow Lodge, West Malvern.
1859 f TRIMEN, Roland, M.A., F.R.S., F.L.S., Ovingdean, King Charles’-
road, Surbiton, Surrey.
1895 Tunauey, Henry, 13, Beemead-avenue,, Streatham, 8.W.
1897 Tunstati, Wilmot, Caerleon, Greenlaw Drive, Paisley.
1898 Turner, A. J., M.D., Widsham Terrace, Brisbane, Australia.
1893 TuRNER, Henry Jerome, 98, Drakefell-road, St. Catherines Park,
Hatcham, S.E.
1894 TurRNER, Thomas, Cullompton, Devon.
1886 Torr, James W., Rayleigh Villa, Westcombe Hill, S.E.
1904 Ty.Lecorr, Edward F. S, M.A., Ravensdene, Warwick Park,
Tunbridge Wells.
1893 Uric, Frederick William, Port of Spain, Trinidad, British West
Indies.
1904 f Vaueuan, W., Cocogalla, Madulsima, Ceylon.
1866 VERRALL, George Henry, Sussex Lodge, Newmarket.
1897 Vick, William A., M.B., 19, Belvoir-street, Leicester.
1895 Wacuer, Sidney, F.R.C.S., Dane John, Canterbury.
1901 Wapprineton, John, Park Holme, Harehill-avenue, Leeds.
1899 Wank, Albert, 52, Frenchwood-street, Preston, Lancashire.
( xxiii)
1897 Watnwriaut, Colbran J., 45, Handsworth Wood-road, Handsworth,
Birmingham.
1870 * WatkER, The Rev. Francis Augustus, D.D., F.L.S., Dun Mallard,
Cricklewood, N.W.
1878 Waker, James J.. M.A, R.N., F.LS., Secretary, <Aorangi,
Lonsdale-road, Summertown, Oxford.
1863 | WatLacr, Alfred Russel, D.C.L., Oxon., F.R.S., F.LS., F.Z.S.,
Broadstone, Wimborne, Dorset.
1866 + WatsineHaAM, The Right Honble. Lord, M.A., LL.D., F.R.S., F.LS.,
F.Z.S., High Steward of the University of Cambridge, Merton
Hall, Thetford ; and 66a, Eaton-square, 8.W.
1886 Warren, Wm., M.A., 61, Wilton-avenue, Chiswick-lane, W.
1869 WatERHOUSE, Charles O., Ingleside, Avenue-gardens, Acton, W. ;
and British Museum. (Natural History), Cromwell-road, S.W.
1901 WarrrHovuss, Gustavus A., B.Sc., F.C.S., Royal Mint, Sydney, New
South Wales, Australia.
1900 Warkins, C. J., King’s Mill House, Painswick, Stroud, Gloucestershire.
1904 Watson, Rev. W. Beresford, St. Martin’s Vicarage, St. Philip,
Barbados, W. Indies.
1893 Wess, John Cooper, 218, Upland-road, Dulwich, S.E.
1876 ¢ WestERN, E. Young, 36, Lancaster Gute, Hyde Park, W.
1886 WHEELER, Francis D., M.A., LL.D., Paragon House School, Norwich.
1884 Wut, William, 75, Thwlow Park-road, West Dulwich, 8.E.
1903 Wiaarns, Clare A., M.R.C.S., Kiswmu, Lake Victoria Nyanea,
British East Africa.
1896 Wiemay, A. E., c/o H.B.M.’s Consul, Kobe, Japan.
1894 Witson, Edwin, Mill-lane, Cambridge.
1904 Winterscats, J.C., F.Z.S., Karangari, Kedah, c/o Messrs. Patterson,
Simons and Co., Penang, Straits Settlement.
1894 Wo.Ey-Dop, F. H., Millarville P. 0., Alberta, N.W.T., Canada.
1900° Woop, H., 9, Church-road, Ashford, Kent.
1881 Woop, The Rev. Theodore, The Vicarage, Lyford-road, Wandsworth
Common, S.W.
1905 Woopsrines, Francis Charles, Northeroft, Cornwall-road, Uxbridye.
1901 Wooprorps, F. C., Market Drayton.
1899 Woottey, H. 8., 7, Park-row, Greenwich, S.E.; and P. 0. Box
1047, Waterbury, Conn., U.S.A.
1891 Wrovucuron, R. C., Inspector General of Forests, Indian Forest
Service, c/o Army and Navy Co-operative Society, Ltd., 105,
Victoria-street, S.W.
1888 Yersury, Colonel John W., late R.A., F.Z.S., Army and Navy
Club, Pall Mail, S.W.
1892 Youpats, William Henry, F.R.M.S., Daltonleigh, Cockermouth.
1904 Youne, L. C. H., 10, Harnam-street, Fort, Bombay.
( =v)
ADDITIONS TO THE LIBRARY
Durine THE YEAR 1905.
[The works marked with an asterisk (*) were presented by Mr. Henry
McLachlan, and belonged to the family of the late R. McLachlan; those
marked with a dagger (+) are second copies of works already in the library. |
Apricn (J. M.). Catalogue of North American Diptera.
[Smithsonian Mise. Coll., No. 1444, 1905.]
The Smithsonian Institution.
AsumEAD (W.H.). Descriptions of New Genera and Species of Hymeno-
ptera from the Philippine Islands.
[Proc. U.S. Nat. Mus., Vol. XXVIII, 1905.]
Additions to the recorded Hymenoptera of the Philippine Islands,
with descriptions of new species.
(Proc. U. S. Nat. Mus., Vol. XXVIII, 1905.]
New Hymenoptera from the Philippines.
[Proc. U.S. Nat. Mus., Vol. XXTX, 1905.]
New Genera and Species of Hymenoptera from the Philippines.
[Proc. U. S. Nat. Mus., Vol. XXIX, 1905. ]
The Smithsonian Institution.
Avurivituius (Chr.). Cerambyciden aus Bolivien und Argentina, Gesammelt
von Freiherrn Erland Nordenskiold.
[Entom. Tidskrift, XX V, 1904.]
Tutt’s British Lepidoptera, Vol. TV (Review)
{Entom. Tidskrift, XXV, 1904.]
Svensk Insektfauna. 13. Steklar. Hymenoptera, I. Gaddsteklar,
Aculeata,
[Entom. Tidskrift, XXV, 1904.]
Eine interessante neue Papilio-Art aus Afrika, von Papilio
Schultzet.
[Ins. Borse, 1904.]
Lepidoptera Heterocera.
[Arkiv fiir Zoologi, Band 2, No. 4, 1904.]
Verzeichnis von Lepidopteren gesammelt bei Mukimbungu am
unteren Congo von Hernn HE. Laman.
[Arkiv fur Zoologi, Band 3, No. 1, 1905.]
Lieut. A. Schultze’s Sammlung von Lepidopteren aus West-Afrika.
[Arkiv fiir Zoologi, Band 2, No. 12, 1905.] The Author.
* AUSSERER (Carlo). Neurotteri Tirolesi colla diagnosi di Tutti i Generj
Kuropei. Parte 1.
[Ann. Soc. Nat. Modena, IV, 1869.]
C ae)
Baker (C. F.). The Classification of the American Siphonaptera.
[Proc. U. S. Nat. Mus., Vol. XXIX, 1905,]
The Smithsonian Institution.
Barcacui (P.), Oration on Stefano Bertolini,
(Bull. della Soc. Entom. Italiana, XXXVI, 1904.] The Author.
*+ Barrett (C.G.). The Lepidoptera of the British Islands. Vols. I-VIII,
8vo, London, 1893-1902.
Bare (T. Hudson). Notes on some Coleoptera from the Flannan Islands,
[Annals of Scottish Nat. Hist., Jan., 1905. ]
Retrospect of a Coleopterist for 1904.
[Reprinted from the Entomologist’s Record, Vol. XVII, No. 2.]
The Author.
* Brecuer (Hduard). Zur kenntniss der Mundtheile der Dipteren,
[Denkschr. Math.-Naturwiss. Classe K. Akad. Wissensch., Vol. XLV,
1882. ]
Becker (Th,), Brzzt (M.), Kerrisz (K. and Stern (P.). Katalog der
Palaartischen Dipteren, Band I
Oyclorrhapha Schizophora ; are von Th. Becker.
Cyclorrhapha Schizophora: Pupipara von Dr. M, Bezzi.
Purchased.
Bett (R. G.). [See Kerroce (V. L.).]
BercenstammM (J. Halen v.). [See Braver (F.).]
Bezzi (M.). [See Becker (Th.).]
Buanprorp (W.F.H.). [SeeGopman (F. D.). Biologia Centrali-Americana. ]
* Borrvar (Ignacio). Sindpsis de los Ortépteros de Espana y Portugal.
4 parts.
[An. Soc. Esp. Hist. Nat., V. pp. 79, 259, VI. p. 249, VII. p. 63,
1876-1878. ]
* Analecta Orthopterologica.
[An. Soe. Esp. Hist. Nat., VII, 1878.]
* Orthoptéres recueillis en Portugal et en Afrique par M. C, Van
Volkem,
{Ann, Soc. Ent. Belg., 1878.]
* Catalogus Orthopterorum, Europe et Confinium.
[An. Soc. Esp. Hist. Nat., VII, 1878.]
* Artrépodos del Viaje al Pacifico verificado de 1862 4 1865. Insectos,
Neurépteros y Ortépteros. 4to, Madrid, 1884.
* Hemipteros Nuevos del Museo de Madrid,
[An. Soc. Esp. Hist. Nat., VIII, 1879.]
* Observations sur les Orthoptéres d’Europe et du Bassin de la
Méditerranée.
[Compt. Rend. Soc, Ent. Belg., 1884.]
* Monografia de los Pirgomorfinos. 8vo, Madrid, 1884.
*———_-y CuicoTr (C.). Enumeracion de los Hemipteros observadas en
Espana y Portugal.
[An. Soc. Esp. Hist. Nat., VIII, 1879.]
Brants (A.). Nederlandsche Vlinders, Beschreven en Afgebeeld. Afi. 1 and 2,
’s Gravenhage, 1905.
[Derde Serie van Sepp’s Nederlandsche Insecten. ] Purchased.
(ax o)
* Braver (F.). Beitrige zur kenntniss Aussereuropaischer Oestriden und
Parasitischer Muscarien.
[Denkschr, Math.-Naturwiss. Classe K. Akad. Wissensch., LXIV,
1896. ]
*——____und Bercrenstamm (J. Edlen v.). Die Zweifliigler des Kaisler-
lichen Museums zu Wien. Parts 1-7.
[Denkschr. Math.-Naturwiss. Classe K. Akad. Wissensch., 1880-
1894. ]
*——_—__und Low (Franz). Neuroptera Austriaca. Die im Erzherzogthum
Oesterreich bisjetzl aufgefundenen Neuropteren nach der analy-
tischen Methode zusammengestellt. 8vo, Wien., 1857.
Bruner (L.). [See Gopman (F. D.). Biologia Centrali-Americana.]
* + Bucxton (G. B.), The Natural History of Eristalis tenax or the Drone-fly.
8vo, London, 1895.
Observations on some undescribed or little-known species of Hemi-
ptera-Homoptera of the Family Membracide.
[Trans. Linn. Soc., Zool., 2nd ser., Vol. IX, Part 9, 1905.]
The Author.
Butter (A. P.). Notes on the occurrence of some rare Species of Lepidoptera.
[Trans. New Zealand Inst., Vol. XXXVII, 1904.]
Burr (Maleolm). Memoir of Henri de Saussure (1829-1905).
[Entom. Record, Vol. XVII, No. 7.]
Earwigs of the Indian Museum, with descriptions of New Species.
[Journal and Proc. Asiatic Soc. of Bengal (New Ser.), Vol. I, No. 2,
1905.]
Notes on the Forficularia, IX: On New Species, with Synonymic
Notes,
[Ann. and Mag. Nat. Hist., ser. 7, Vol. XVI, Nov. 1905.]
The Author.
* CatveRT (P. P.). The Odonata of Baja California, Mexico.
[ Proc. Calif. Acad. Sci., ser. 2, Vol. IV, 1895.]
[See Gopman (F. D.). Biologia Centrali-Americana.]
*+CamBripce Naturat History (The). Edited by S. F. Harmer and A.
E. Shipley. Vols. V and VI. 8vo, London, 1895-99.
Vol. V, Peripatus, by A. Sedgwick. Myriapods, by F. G. Sinclair.
Insects, Part 1, by D. Sharp (1895).
Vol. VI, Insects, Part 2, by D. Sharp (1899).
CaMERON (P.). Descriptions of New Species of Sphegidz and Ceropalide,
from the Khasia Hills, Assam.
[Ann. and Mag. Nat. Hist., ser. 7, Vol. XV, 1905.]
G. A. James Rothney.
*CARPENTER (G.H.). Insects, their Structure and Life. A Primer of Ento-
mology. 8vo, London, 1899.
Campion (G.C.). [See Gopman (F. D.). Biologia Centrali-Americana.]
Cuicors (C.). [See Borivar (Ignacio). ]
* Comstock (J. Henry). Report upon Cotton Insects, prepared under the
Direction of the Commissioner of Agriculture. 8vo, Washington,
1879.
(> xxvii ©)
*Costa (A.). Rapporto preliminaire e sommario sulle ricerche Zoologiche
fatte in Sardegna durante la primavera del 1882.
[Rend. R. Accad. Sci. Fis. e Mat. di Napoli, 1882. ]
* Nota intorno i Neurotteri della Sardegna.
{Rend R. Accad. Sci. Fis. e Mat. di Napoli, 1884. ]
* Notizie ed Osservazioni sulla Geo-Fauna Sarda. Memoria Terza.
[Atti R. Accad. Sci. Fis. e Mat. Napoli, ser. 2, Vol. I, 1884.]
* Notizie ed Osservazioni sulla Geo-Fauna Sarda, Memoria quinta.
[Atti R. Accad. Sci. Fis.e Mat. Napoli, ser. 2, Vol. II, 1886.]
* Notizie ed Osservazioni sulla Geo-Fauna Sarda, Memoria sesta.
[Atti R. Accad. Sci. Fis. e Mat. Napoli, ser. 2, Vol. II, 1886.]
Currie (Rolla P.). Catalogue of the Exhibit of Economic Entomology at the
Lewis and Clark Centennial Exposition, Portland, Oregon, 1905.
[U.S. Dept. of Agriculture, Bur. of Ent., Bull. No. 53, 1903.]
* Date (C. W.). The Lepidoptera of Dorsetshire ; or, A Catalogue of the
Butterflies and Moths found in the County of Dorset. 2nd ed.,
8vo, Dorchester, 1891.
Davis (K. C.). [See Nerpuam (J. G.).]
Disrant (W. L.). A Naturalist in the Transvaal. London, 1892.
The Publisher.
[See Gopman (F. D.). Biologia Centrali-Americana. |
Dixon (H. H.). Note on the Supply of Water to Leaves on a Dead Branch.
[Scientific Proc. Royal Dublin Soc., Vol. IX (New Ser.), No. 2, 1905. ]
i The Author.
Druce (Herbert). Description of some new species of Noctuide from
Tropical South America. as
[Ann. and Mag. Nat. Hist., ser. 7, Vol. XV, 1905, ]
Description of some new species of Syntomide and Arctide from
Tropical South America.
[Ann. and Mag. Nat. Hist., ser. 7, Vol. XV, 1905.]
The Author.
Dyar (H. G.). A Descriptive list of a Collection of Early Stages of Japanese
Lepidoptera.
[Proc. U.S. Nat. Mus., Vol. XXVIII, 1905.]
A list of American Cochlidian Moths, with descriptions of new
genera and species.
[Proc. U.S. Nat. Mus., Vol. XXIX, 1905.]
New genera of South American Moths.
[Proc. U. 8. Nat. Mus., Vol. XXIX, 1905.]
The Smithsonian Institution.
*+ Epwarps (James). The Hemiptera-Homoptera (Cicadina and Psyllina)
of the British Islands. 8vo, London, 1896.
* Epwarps (William H.). Butterflies of North America. Second series.
4to, Boston, 1884.
Butterflies of North America. Third series. 4to, Boston and New
York, 1897,
* EHRENBERG (C. G.) et Hempricu (F. W.). Symbole Physicz, seu
Icones et Descriptiones corporum Naturalium novorum aut
minus cognitorum que ex itineribus per Libyam, Aigyptum,
Nubiam, Dongalam, Syriam, Arabiam, et Habessiniam, etc.
(Neuroptera, by F. Kiue.) Folio, Berolini, 1828-45.
(>), sexval
* Enoyctoprpie Mérnopiaue. Histoire Naturelle—Text. Tomes iv-x.
Insectes. By Mauduyt, Olivier, Latreille, Godart, Saint-Fargeau,
Serville et Guérin. Histoire Naturelle—Plates. Crustacés,
Arachnides et Insectes, By Latreille. 2 vols., 4to, Paris, 1789-
1825.
* ENDERLEIN (Dr. Gunther). Zur kenntnis der Insekten Deutsch-
Ostafrikas.
[Mittheil. aus dem Zool. Mus. zu Berlin, IT, 1902.]
Eustace (H. J.). Winter Injury to Fruit-trees.
[New York Agric. Exp. Stn., Bull. No, 269, Oct. 1905.]
The Author.
*Frtr (H. P.). Insects Injurious to Elm-trees,
[Fifth Ann. Report Fisheries, Game and Forest Commission, State
of New York. ]
Insects affecting Forest Trees.
[Reprinted from Seventh Report Forest, Fish and Game Commis-
sion, State of New York, 1900. ] The Author.
* FERNALD (Mrs. M.E.). A Catalogue of the Coccide of the World. 8vo,
Amherst, Mass., 1903.
[Special Bulletin, Hatch Experiment Station of the Massachusetts
Agricultural College. |
* Finort (A.). Les Orthoptéres de la France. Perce-Oreilles, Blattes, Mantes,
Criquets, Sauterelles et Grillons. 8vo, Paris, 1883.
* Forpes (Henry O.). The Natural History of Sokotra and Abd-el-Kuri.
Being the Report upon the Results of the Conjoint Expedition to
the Islands in 1898-9, by Mr. W.R. Ogilvie-Grant, of the British
Museum, and Dr. H. O. Forbes, of the Liverpool Museums,
together with information from other available sources, forming
a Monograph of the Islands. 8vo, Liverpool and London, 1903.
* Forpss (S. A.). 17th, 19th, 21st, 22nd Reports of the State Entomologist
on the Noxious and Beneficial Insects of the State of Illinois.
8vo, Springfield, Chicago, and Champain, 1891-1903.
Foret (A.). Miscellanea Myrmécologiques, IT.
[Ann. Soc, Ent. Belg., XLIX, 1905. ] The Author.
[See KRAEPELIN (Karl). ]
Fowter (W. W.). [See Gopman (F. D.), Biologia Centrali-Americana. ]
* Frisco (J. L.). Beschreibung von allerley Insecten in Teutschland nebst
niitzlichen Anmerkungen und nothigen Abbildungen von die-
sem kriechenden und fliegenden inlandischen Gewiirme. Thiele,
1-10, 13, sm. 4to, Nicolai, 1720-38.
* Frocecarr (N. W.). Scale Insects that produce Lac. 8vo, Sydney, 1900.
* Notes on Australian Coccidz (Scale Insects). S8vo, Sydney, 1900.
(Dept. of Agric. N. 8. Wales, Misc. Publ. No. 358.]
* Plague Locusts. 8vo, Sydney, 1900.
[Dept. of Agric. N.S. Wales, Misc. Publ. No. 363.]
* The Hessian Fly (Cectdomyia destructor, Say) and Allied Grain
Pests. 8vo, Sydney, 1900.
[Dept. of Agric. N. 8. Wales, Mise. Publ. No. 369. ]
* Insects and Birds. 8vo, Sydney, 1900.
[Dept. of Agric. N. 8. Wales, Mise. Publ. No. 387.]
* Insects living in Figs, with some account of Caprification. 8vo,
Sydney, 1900.
(Dept. of Agric, N.S. Wales, Misc. Publ, No. 388.]
(°eeix,”':)
* Fiurnroure (A. H.). Naturhistorische Topographie von Regensburg. In
Verbindung mit Forster, Herrich-Schaffer, Koch, v. Schméger
und vy. Voith. 3 vols. 8vo, Regensburg, 1838-40.
* Furter (C.). First and Second Reports of the Government Entomologist
for Natal, 1899-1901. 8vo, Pietermaritzburg, 1901-2.
Fourth Report of the Government Entomologist, Natal, Depart-
ment of Agriculture, 1903-4. 8vo, Pietermaritzburg, 1905.
* Gay (Claudio). Historia fisica y politica de Chile, segun documentos
adquiridos en esta republica durante doce anos de residencia en
ella y publicada bajo los auspicios del supremo gobierno por C.
Gay. (Zoologia.) Orthopteros and Neuroptera (Atlas only).
Folio, Paris, 1847-54.
* GirARD (Maurice). Catalogue Raisonné des Animaux Utiles et Nuisibles
dela France. 2 parts, 8vo, Paris, 1878.
Fasc. 1. Animaux Utiles, leurs services et leur conversation.
Fasc. 2. Animaux Nuisibles dégats qu’ils produisent, moyens de
les detruire.
* + Le Phylloxera de la Vigne, son organisation, ses mosurs choix
des procédés de destruction. 38rd ed., 8vo, Paris, 1880.
Gopman (F. Ducane). Biologia Centrali-Americana. Parts OLXXXVI—
CLXXXIX. The Editor.
Arachnida Araneidea and Opiliones, by F. O. Pickard-Cambridge.
Insecta, by W. F. H. Blandford, L. Bruner, P. P. Calvert, G. C.
Champion, W. L. Distant, W. W. Fowler, and D. Sharp.
Gortpr (Emilio Augusto). On Mosquitos in Para.
Memorias do Museu Goeldi (Museu Paraens) de Historia Natural e
Ethnographia, 1905. ] The Author.
GREEN (Ernest E.). Toxorhynchites immisericors (Walker), The Elephant
Mosquito.
[Spolia Zeylanica, Vol. II, Part VIII, 1905.] The Author.
* Hacen (H. A.). Phryganidarum synopsis synonymica,
[Verhandl. K. K. zool.-bot. Gesellsch. in Wien, 1864. ]
Hammonp (A. R.). [See Mratt (L. E.).]
Hampson (Sir Geo. F.). Catalogue of the Noctuids in the Coll. of the
British Museum, Vol. V and one vol. of Plates, 1905.
In Exchange.
Hancock (J. L.). The Habits of the Striped Meadow Cricket (Gcanthus
fasciatus, Fitch). :
[American Naturalist, Jan. 1905.] The Author.
The Tettigidz of Ceylon.
[ “Spolia Zeylanica,” Vol. II, Part 7, Oct. 1904.] The Author.
Hanpuirscu (Anton). Friedrich Moritz Brauer (Obituary Notice).
[Verhandl. K. K. zool.-bot. Gesellsch. in Wien, 1905. ]
The Author.
Hempricu (F. W.). [See EnrENBERG (C. G.).]
Herzoc (M.). The Plague: Bacteriology, Morbid Anatomy and Histopa-
thology, including a consideration of insects as plague carriers.
(Dept. of Interior, Bureau of Gov. Lab., Manila, No. 23, 1904. ]
The Bureau of Gov. Lab.
(- *ogscx. 2)
Hinps (W. C.) and Hunter (W, D.). The Mexican Cotton Boll Weevil.
[Dept. of Agric., Div. of Entom., Bull. No. 45, also No. 51, a revision
of the above. | U.S. Dept. of Agriculture.
Hotiann (W. J.). Moth Book. 8vo, New York, 1903. The Author.
* Ho~mGrEN (Aug. Emil.). Ichneumonologia Suecica. Tom. iii. Ichneu-
monides Pneustici. Svo, Holmiz, 1889.
* Hupson (G. V.). New Zealand Neuroptera. A popular introduction to the
life-histories and habits of May-flies, Dragon-flies, Caddis-flies,
and allied Insects inhabiting New Zealand, including notes on
their relation to Angling. 8vo, London, 1904.
* + InpEx Zootocicus. An Alphabetical List of names of genera and sub-
genera proposed for use in Zoology, as recorded in the
“Zoological Record,” 1880-1900. By Charles Owen Water-
house. 8vo, London, 1892,
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OF
LONDON
For THE YEAR. 1905.
I. The Life History of Gerydus chinensis, Felder. By
JOHN C. W. KerrsHaw, F.L.S., F.E.S.
[Read November 16th, 1904.]
PLATE I.
ALTHOUGH the imago of Gerydus chinensis is peculiar,
with its very long body and broad, flattened legs, yet its
early stages are stili more curious. The butterfly is fairly
common throughout the year in certain localities near
Macao and Hongkong, but it keeps more or less strictly to
these shady and usually damp places, and is to a certain
extent crepuscular, though it is also on the wing during
the day.
It lays its eggs towards evening and till night fairly sets
in, on stems and leaves of plants and trees ‘infested with
Aphid es or Hemiptera and overrun by a host of ants of two
species (Polyrrhachis dives, Sm., and Dolichoderus bitu-
berculatus, Mayr.), both Aphides and ants apparently feeding
on juice exuding from the plant. Any vegetation subject
to these secretious seems to be attractive to them, the ants
not appearing to use the Aphides as ant-cows, though of
this lam not quite certain. The Aphides swarm so thickly
that the plant-stem or leaf is invisible, and, barring heavy
rain, they are as a whole stationary for days together,
though slowly changing their positions individually, and
going through various ‘transformations, fresh lots continu-
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 1
2 Mr. J. C. W. Kershaw on
ally replacing the old. The plants they seem specially to
affect are various species of bamboo, lichee, Stillingia
sebifera and a species of bur-marigold, Bidens pilosa, repre-
sented in the drawing. This latter is a very common
plant here, possessing bunches of spiky seeds with branched
ends armed with microscopic spines pointing downwards,
which catch in every imaginable thing that touches them.
The 2 butterfly, after almost interminable dancing up
and down and wandering hither and thither, finally alights
after two or three attempts in the midst of the Aphides
and ants, which she thrusts aside with a brushing move-
ment of her tail, immediately laying a single egg. She
then generally moves slightly and remains for some time
sucking up the exuding juice of the plant; both ¢ and ?
are very fond of it: half-a-dozen may occasionally be seen
close together on one leaf or stem, drinking this sap,
thrusting their tongues into any interstices left by the
Aphides, The ants do not appear to meddle either with
the butterflies or the eggs, though ants are very destructive
to eggs of most butterflies, and I have just recently seen
two butterflies (a Hesperid and a Neptis) seized by the
tongue as they probed a flower, and dragged off by this
same species of ant. Nor do they seem to interfere with
the larve. Probably they are too much occupied in
drinking the sap to trouble about eggs, ete,
The egg is circular and flat, of a pale green, ringed cir-
cumferentially, the edges of the rings milled like a coin.
It hatches in four days, the issuing larva being nearly
cylindrical at first, not assuming its slug-shape till a later
stage. It is light yellow, with a distinct purplish dorsal
line, and a few light-coloured hairs chiefly at head and
tail, the head dark. Later on it becomes limaciform, yellow
or greenish-yellow and banded longitudinally with purple-
brown, the segments well-defined, the first segment swollen
and produced, so that the head can be withdrawn entirely
into it, as it usually is when the larva is resting.
The larvae feed on the Aphides, sometimes pressing them
against the plant with head and forelegs, sometimes hold-
ing them in the forelegs quite away from the plant. <A
few bites disposes of an Aphis, and the larva then licks
and cleans its legs, just as a Mantis does. Some Aphides
must have a better flavour than others, as the larvee pick
and choose, moving their heads up and down over the
backs of the insects, evidently smelling them. As a rule
The Life History of Gerydus chinensis. 3
the creatures seem to make little attempt to escape till
they are actually bitten, when struggling is useless,
When not engaged in feeding, the larve rest amongst the
Aphides or crawl leisurely about between or over them,
and the Aphides do likewise, the larve being sometimes
covered with them. The eggs of the butterfly, too, are
often hidden under a mass of Aphides.
When nearly full-grown the larve lose most of their
sparse hairs and their colours fade, and the bands and four
blotches on the seventh and eighth segments become less
distinct. When about to pupate they walk about at quite
a fair rate of speed, and having chosen a spot, spin a few
threads a little distance from head and tail. My larve
pupated on a piece of bark, one putting a band round its
middle and pupating horizontally, the other vertically and
without a band, though both had the threads (not con-
nected to the pupz) at head and tail. The tail of the
pupa is cut off squarely, forming a disc, thus securing a
good hold, probably sufficient without the band round the
middle. It seems to be affixed with some secretion, not
actual threads. There is a small process each side of the
third segment from the tail. The pupal state lasts about
ten days. Both larva and pupe strike one as being small
compared with the size of the butterfly. I have only
found the larve feed on two species of Aphis, but they
doubtless eat other kinds. One was slate-coloured with
white efflorescence, the other greenish with four dark-green
patches, some of them being fringed with white, probably
moulted skin.
If the butterflies were only numerous enough they
would certainly prove a blessing to the gardener, for many
trees here, especially the lichee (a fruit tree) suffer very
much from “bugs” of all sorts which, even when they do
not entirely eat away the leaves, seem to blight them or
cause them to shrivel up. The larval state lasts about
fifteen days, and I reckoned on the average, from first to
last, the larva would eat some twenty Aphides per day,
but it would require many larve to make much impres-
sion on the crowds of Aphides one sees; perhaps a yard
of bamboo stem two or three inches in diameter being
absolutely covered with these disgusting insects.
I am not sure, but have some reason to think that the
larvee tend to assimilate in colour to the Aphides they
happen to be feeding on. At first, as the eggs were laid
4 Mr. J. C. W. Kershaw on Gerydus chinensis.
amongst the green Aphides, I fed the larvee thereon, but a
typhoon having washed away the food supply, I could
only find the slate-coloured species, which, however, the
larvee seemed to enjoy quite as much. But I noticed soon
after that the colours faded and the purple-brown tended
to mingle with the yellow. This is a question it will be
interesting to decide. One of the features of this curious
life history is the calm way in which the larva moves
amongst the Aphides and selects his prey, and the in-
difference with which the latter apparently accepts his
fate.
I have since found swarms of a bright yellow Aphis
on some Asclepias plants, on which I hope to rear some
larvee, with the object of testing their adaptability of
colour to their surroundings. Gerydus chinensis lays at
intervals all through the year, but these eggs were taken
in July and August.
EXPLANATION OF PLATE I.
Life History of Gerydus chinensis, Felder.
Butterfly laying egg amongst Aphides.
Fia. 1. Egg, enlarged.
2. Larva just hatched, enlarged.
3. ,, nearly full-grown, natural size.
4, Pupie, natural size.
5. Larva eating Aphis, enlarged,
6. Aphides, two species, enlarged.
Il. Butterfly-destroyers in Southern China. By Joun C.
W. Kersuaw, F.LS., F.ES.
[Read November 16th, 1904. ]
PERHAPS the creatures which in this district destroy
or injure the largest number of butterfly imagines are
lizards, particularly one species (Calotes versicolor, Daud.),
which climbs up into the clumps of Lantana camera, a
profusely flowering plant or shrub which grows here on
most waste ground. Probably the greater part of their
prey consists of Hesperiidx, three or four species of which
haunt the flowers of the Lantana in immense numbers.
Other reptile foes to butterflies are probably the tree-
frogs, and possibly the very numerous small snakes which
are ‘always climbing in the tops of the bushes.
Spiders’ webs account for some mishaps to butterflies,
but they are not invariably eaten; at other times species
which are reckoned most distasteful as food of other
animals are seized and eaten at once, according, I suppose,
as the owner of the web is hungry or otherwise. I have
seen our large black-and-yellow spider (Hpeira maculata
of Donovan’s “ Insects of China”) eating Huplea amymone,
Godt., caught in his web. He had attacked and consider-
ably injured the lower side of the abdomen and part of the
thorax ; but when I took the butterfly from him, examining
it Jaid Hat in my hand, the insect suddenly flew up to the
top of a large banyan tree, where it settled. One would
hardly imagine even a Huplwa would have much life in it
after being so badly mauled. The same spider I have
seen eating Papilio dissimilis, L. (= clytia, L.,=panope, L.),
the black-and-yellow form. Also at different times Veptis
eurynome, L., and Huplea midamus, L., but on the whole
it is not common to see a butterfly in a spider’s web. I
might mention that I saw this spider devouring a small]
bat, about eight inches across the wings, caught overnight
in his web.
There is also here a small but thickset white spider
which usually conceals itself in white flowers, and which
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY)
6 Mr. J. C. W. Kershaw on
I have two or three times seen capture the little black-
and-yellow Skipper, Z'elicota bambusw, Moore, by the head.
The large green Mantis, Mierodula saussurei, W. F.
Kirby, our commonest species, apparently catches anything
he can get hold of. I have kept Mantidse and fed them
with many species of butterflies, none of which they
refused, but data obtained from creatures in captivity are
not, I think, very reliable, as under these circumstances
they often take food which in a natural state they would
probably reject. But I have seen this Mantis in the open
on a shrub, eating the two species of Huplea before men-
tioned. He is a very bold insect, and, even in a wild
state, seldom refuses any butterfly offered to him.
One of the worst enemies of the Hesperiidx is a large
hairy species of (Asilid ?) fly, which seizes them and drives
its proboscis into the thorax. Either it is very fearless or
it cannot easily disengage its weapon from its prey, for I
have often kept stirrmg one up with a stick, when it
would merely fly to another leaf close by, still holding its
victim. These flies also kill a moderate-sized Cicada in
great numbers.
In the last two months (August and September) I have
three times seen butterflies seized by the tongue by ants,
as they probed a flower. On one occasion an ant seized
Neptis eurynome by the tip of the tongue, and the butter-
fly immediately flew away with the ant hanging on it.
About half-an-hour afterwards I captured the Neptis with
the ant still clinging to its tongue. The other instances
were both Zelicota bambusx, and each was caught by the
tip of the tongue by an ant.
I should be inclined to think that the birds here account
for very few imagines, comparatively speaking, whatever
they may do in the larva and pupa line. The only birds
I have seen attack butterflies, and generally miss them,
are sparrows (our common sparrow is Passer montanus),
the green Bulbul, Pycnonotus sinensis, the black Drongo,
Buchanga atra aud the Paradise-flycatcher, Terpsiphone
princeps. During five years I have, perhaps, seen a dozen
attacks on butterflies by birds, and only seen them cap-
tured two or three times. Speaking from my own limited
experience I should say imagines here have few enemies,
and that those few inflict very slight damage. Unfavour-
able climatic conditions seem to be most inimical to
them. (Cf. Trans. Ent. Soc. 1895, pp. 437-8.)
Butter fly-destroyers in Southern China. 7
But in the egg, larva and pupa state, butterflies here
have a host of enemies, the eggs especially being destroyed
by ants, which also attack and carry off young larvae and
pupe. Ants almost certainly account for the greater
part of casualties amongst eggs, because of their being
ubiquitous and in countless swarms. One small orange
plant, which I often examined many consecutive mornings,
was much frequented by 2? 2 of Papilio polytes, L., and P.
helenus, L., also P. sarpedon, L. I have frequently seen the
? Jay an egg and fly off, and a moment after an ant, waiting
below the leaf, would carry the egg away. Sometimes I
have noticed a particular leaf and stem with three or four
egos close together on them, and returning an hour or
so later have found them gone, almost certainly taken by
the ants which were invariably climbing over the plant.
Centipedes kill pupz, twining round them, boring a
tiny hole, and appearing to suck out the contents.
Some of the Hemiptera or “ bugs” (Capside, I think)
probably account for the destruction of some pups, as I
have seen one with its trunk driven into a chrysalis, and two
or three times have seen a small red-and-black bug, about
an inch long, force its proboscis through the thin shell of
a small snail and eat or suck out the inmate.
Amongst birds the Cuckoos especially must destroy
very great numbers of butterfly larvee during their short
summer stay here, Cuculus micropterus probably inflicting
most damage. The Cuckoo, like the Mantis, will eat
almost anything, and if a larva is particularly hairy it
rubs it up and down on the ground or tree trunk till it
breaks off most of the hair. It seems very fond of the
larvee of Rhopalocampta benjamini, Guer., a bright yellow-
and-black larva with a red head, and during May and June
its stomach generally contains several, judging from those
shot. Wasps, of course, carry off many larvee, but chiefly,
I think, those of the Pieride, which are usually fairly
smooth-skinned. I wasrearing a lot of Catopsilia pyranthe
larvee on the roof of the house, on a Cassia plant, and
the wasps carried off so many I was obliged to cover
them up. The ants, too, killed many of the newly-
hatched larve, and as soon as pupating began, started
carrying off the pupeze.
Ichneumonide I pass over, all kinds of larve in all
parts of the world being presumably attacked by them,
but some of the Pieride here seem especially subject to
8 Mr. J. C. W. Kershaw on Butter fly-destroyers.
their attacks. Of about a score pupee of Delias aglaia, L.,
found on a shrub, three produced butterflies, the rest
ichneumons. Of the same number of young larve reared,
all fed well till about half-grown, then all but six dwindled
away to mummies, two died when full-grown; the rest
pupated, but only two butterflies appeared, the other
pupe shrivelling up. They were supplied with fresh food
every day, and till half-grown appeared very healthy.
There seem to be special checks on the increase of some
species of Pieride.
No doubt the injuries done to the wings of butterflies
are often the result of attacks by various enemies, but I
think the greater part are caused by the insects themselves
flying through thick cover, where one may often distinctly
hear and see the wings of a Papilio strike leaves and
twigs ; or by getting into and escaping from the numerous
spiders’ webs, and threads stretched from tree to tree; or
by simple stress of weather. Several times I have seen
what I took, at first sight, to be a new species of Papilio
which when taken proved to be either P. polytes or P.
dissemilis with the hind-wings covered with the red or
yellow pollen of flowers, generally the large flowering tree
Bombax ceiba. These specimens inv ariably had the wings
torn, and as the pollen is sticky it probably was partly the
cause of various rents and ragged margins.
One often sees a Papilio, generally either P agamemnon,
L., or P. sarpedon, which haunts a certain locality or “ beat ”
for days together, flying at a good height and seldom
settling ; these “solitaries” chase away other butterflies and
even dragon-flies, and I have seen Hupleing and Neptis
chase dragon-flies, which swarm here. Of all insect foes I
should have thought that dragon-flies, from their very
swift flight and powerful jaws, would have been most
dangerous to butterflies, but I have never seen one attack
a butterfly, though they sometimes kill and eat each other.
My few notes would, in short, point to butterflies which
have escaped the sundry and manifold dangers of the egg,
larva and pupa stages having collectiv ely, comparatively
little to fear in the perfect state.
Go's)
III. On Erebia palarica,* n. sp., and Erebia stygne ;
chiefly in regard to its association with K. evias,
in Spain. By Dr. THomas A. CHAPMAN, M.D.
[Read December 7th, 1904.]
Puates: IT, TEZEV, Vv; Va;
In 1902 I reported that Z. stygne, Ochs., had not been
recorded as occurring in Spain. In this I am not quite sure
that I was not correct, but at any rate, if not recorded, the
Entomologists of Madrid were certainly aware that the
species occurred in the Sierra Guadarrama, as they have
specimens in their collections, and have it noted in their
MS. lists. I am not aware that even now any Spanish
localities are known beyond this one, except these recorded
in our Proceedings by Mrs. Nicholl and myself.
In 1902 and 1903 I met with forms of Z. stygne in Spain,
that led me to make sundry observations on that species
to the Society, in connection with specimens exhibited.
Mrs. Nicholl also met with the species in 1902, and also
Prof. Poulton, and I was able to make some remarks on
their specimens which were exhibited at our meetings.
Amongst the general remarks I hazarded, I expressed the
opinion that Hrebia stygne in Spain was well worthy of
further observation, and study. My observations during
1904 have, I think, fully confirmed the soundness of this
opinion, but though adding something to our knowledge,
they leave the necessity for further research in regard to
the species and those allied to it, at least as cogent as
before.
It may perhaps be well, before relating last season’s
* Palarica, from habitat, Pajares. Pajares is said to mean a place
with much fodder and litter, as if Paleares = ewm multas paleas; a
bad (and I am not sure that this is a bad) excuse for getting rid of
the j, is better than none, as the Spanish j = Scotch ch, is shibboleth
to the South Briton ; Spanish j is often = Latin], e.g. mejor—melior.
T should have liked to have given the name nicholli, in honour of
Mrs. Nicholl, who first took the species two years ago, but this would
lead to confusion with Erebia glacialis, var. nicholli, Oberth.
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY)
10 Dr. T. A. Chapman on
experience, to recapitulate our previous knowledge of Z.
stygne as a Spanish species.
I found in 1902 a very large and brightly-coloured race
of stygne at Bejar, which I named bejarensis,* and ex-
hibited to the Society on Nov. 5th. At the same meeting +
Mrs. Nicholl exhibited a large and small form of stygne
from the Picos de Europa. Of the large form which is no
doubt identical with palarica, there were only two poor
specimens, and all the evidence went to show they were a
form of stygne, but coming from the same locality as a
small form, it added an important item of interest to the
species.
In the following year, I took at Canales de la Sierra a
fairly large form of stygne, associated mimetically with a
small form of £#. evias, nearly, if not absolutely identical
with Zupater’s evias, var. hispanica, so I gave the same
varietal name to the form of stygne. These facts are
reported in the Proc. Ent. Soc. 1903, pp. xlv ef seq.
Stull later the President enabled me to show the Society
(Proc. 1904, p. xlvi) two Erebias from the Guadarrama,
which confirmed the existence of stygne there, and also
showed that it was there accompanied by a specially
modified form of £. evias.
It further happens that #. evias accompanied the small
stygne taken by Mrs. Nicholl at the Picos, though this had
not been recorded when I examined the specimens with a
view to the present report.
All the forms of stygne so far known from Spain there-
fore seemed to be no doubt varieties of that species, some
of them modified to agree with #. evias. There was
nothing in any of the forms that did not appear to be well
within the limits of variation.
Mrs. Nicholl has an interesting note on the stygne of the
Picos in Ent. Record, vol. xvi, p. 48. I may quote some
items of this, as I cannot find any details of Mrs. Nicholl’s
observations on the Picos Erebias in the Society’s Trans-
actions.
July 10th. “EHncamped at a height of nearly 5000 ft. on
the southern face of the chain, and found the mountain
pastures just above this level, swarming with 4H. stygne.
Both sexes were out and in good order, though some of
the males were slightly worn. I caught a good many
and kept a few; they were quite typical specimens, per-
* Proc. Ent. Soc. 1902, p. xxxvi. PL Aep. LEV,
Erebia palarica and Erebia stygne. 11
haps a trifle larger than those from Dauphiné and the
Pyrenees.”
On the 19th, “stygne was nearly over.” Mrs. Nicholl
tells me that she also took evias (‘‘mostly worn, rather
large type, and high up, none low down”).* I have
already noted two specimens of evias are mixed with the
stygne (no doubt some of the few noted above) from the
Picos in the drawers at South Kensington. I incline to
think they were not so mixed by Mrs. Nicholl, but, whoever
did so, afforded a proof that the high-level July evzas from
the Picos does, in some of its specimens at least, closely
resemble the stygne of the same time and place. The
note in the “Record” proceeds: “On July 22nd, at a
height of 3000 to 3500 ft. I saw several large Erebias, quite
fresh out. I caught six or seven specimens, all males, and
considered them to be Z#. xthiops and therefore only kept
acouple, . . . they turned out to be very large specimens of
E. stygne.” These two specimens are Z. palarica. Both
specimens are, however, poor, the, one in the British
Museum, the best, is very far from fresh, so that the
difference of date between /. yalarica and E. stygne is less
than Mrs. Nicholl’s note implies. They are no doubt on
the wing at the same time at the Picos, as I found them
this year at Pajares. I must take my share of responsi-
bility for declarmg these specimens of palarica to have
been stygne. ‘They were very much the same size as my
EE. stygne, var. bejarensis,+ with which I was then busy. So
that their large size did not suggest specific difference,
whilst the genitalia of the specimen I examined, though
not quite typical stygne, were within the limits of variation
found in these appendages in other species of the genus.
With the small amount of material available the conclusion
seemed clear that it must be regarded as a var. of stygne,
possibly near var. bejarensis, both being very large, but
certainly requiring further investigation.
We now come to my observations of the present year
(1904). From July 8th to July 22nd, Mr. Champion and
I stayed at Puerto de Pajares, which is the pass across the
Cantabrian Mountains followed by both the road and the
railway from Leon to Gijon, at an elevation of about
4500 ft., and at several places in the neighbourhood we
* See Notes on evias later.
{ (The specimen in B. Mus. is 56 mm.; small for palarica,
& maximum for bejarensis.)
12 Dr. T. A. Chapman on
found a large and a small form of stygne, which, there can
be little doubt, with probably some trifling local variations,
are the same two forms as those taken by Mrs. Nicholl at
the Picos de Europa. In one respect our observations,
though confirming Mrs. Nicholl’s, show that the apparent
conclusion to which they pointed is incorrect. Mrs. Nicholl
found one form at a high, and the other at a low level, and
the deduction was, that it was the habit of the two forms
to be separated by being adapted to living at different
elevations, even if the lower one were not a second brood
possible at the warmer station,
Well, we found one form at a low, and the other at a
high level, but then again we found precisely the contrary
case, in fact we found both forms at all levels from about
4000 ft. to over 6000 ft. The small form maintains
its characters both at a high and low level, as does also
the large one.
We found, however, that each species had definite local-
ities, within which the other did not oceur, and in the
case of two such localities for the small species, and also of
two for the large, the localities seemed to be very definitely
circumscribed. The species were found in other places in
which also the habitats may have been as definitely
marked out, but it did not happen that our examination of
them was sufficiently minute to justify an opinion. In-
deed, of those I regarded as well defined, this is of course
only true inregard to certain directions, but these directions
were sufficiently numerous, being in fact the directions
from which we turned back when collecting them, because
no more were to be seen, to justify the broad statement.
I must go into a little more detail as to the reasons that
led me to consider the large form palarica, to be a distinct
species from the small one, which I regard as a local form
of stygne very close to my var. hisganica.
The first and obvious reason is the difference in size.
Size one regards usually as of little value as a specific
character, and if it were in this case to stand alone, I
should, as I did in the case of var. lejarensis, attach no
importance to it. The size is, nevertheless, in this case a
very marked and unmistakable character. I have measured
all the specimens I brought home, and the result of
measuring 55 f 11 @ of stygne from Pajares, and 115 f
and 37 2 of palarica is, that the most dwarf palarica is
1 mm. larger than the most giant stygne (hispanica), and
Erelia palarica and Erebia stygne. 13
that the mean expanse of the two species differs by no less
than 11 mm., nearly half-an-inch. The smallest ? ranging
in size very close to stygne, var. hispanica, has a facies quite
characteristic of palarica, so that no one could confuse them
together.
Palarica is indeed the largest of all the Erebias, averag-
ing 59:0 mm. in expanse, and ranging from 55 mm. to
64 mm.
The largest Hrebia according to Ruhl is parmenio, to
which he gives an expanse of 50 mm. to55mm. 4. cyclopius
is as large, and embla very nearly so. I find, however,
that evias goes to 54, mm., but this is a maximum.
Palarica has 59 mm. as an average.
In size, stygne, at Pajares, and palarica do not overlap ;
the largest of the one is smaller than the smallest of the
other.
The next point is that each was, wherever we closely
observed it, strictly confined to its own habitat, into which
the other did not trespass ; this was very remarkable at two
pots where the areas occupied by each approached to
within a few dozen yards of each other. At one of these
places opportunity served for me to notice how sharply
defined was the margin of the territory of each species, and
how a specimen driven over the border, came back after a
very short detour. Nor did I ever find one butterfly where
I found the other though it might not be very far off.
Still it is difficult to suppose, considering how common
they were in places, each in its own area, and how close
these areas often were, that specimens did not sometimes
visit the habitats of the other species, though I did not
meet with such a case. And if they did and they were
really all one species some crossing was to be expected, and
intermediate forms ought to have occurred. Yet I must
have seen many hundreds of specimens altogether of both
species. In most places there was some difficulty in taking
specimens, and sometimes only two or three were taken
out of a score seen, and of these taken, though many
wretched specimens were retained, a large number in poor
condition were discarded. Those brought home must
therefore be but a portion of the number of individuals from
which I draw conclusions. Yet there never was the
slightest difficulty in saying at once, of which species any
particular specimen was. There was no trace of interme-
diate or transitional forms or of hybrids. Considering how
14 Dr. T. A. Chapman on
easy, even inevitable, crossmg must have been, had they
been merely forms of one species, I see no means of avoid-
ing the conclusion that the two forms are asyngamic.
Such an attitude towards each other, seemed to be regarded
as the truest test of specific distinctiveness of two forms,
when the subject of “what is a species” was debated at
our meeting last Spring. The difficulty is to put it to
experiment. In the case of Hrebia palarica nature has
provided the experiment, and the answer is distinct.
Usually she separates the subjects of experiments so
widely by time, place, and season, that the answer we have
to form is a purely personal one, viz. [ think if the experi-
ment were made the answer would be so and so. It was
in this way that I concluded, and still hold that all the
Spanish forms of séygne previously known to me are of
one species.
When we examine the insects themselves for confirma- —
tion of this conclusion, we first have size, usually a poor
specific character, but here so marked and constant as to
have some weight. Thenas to markings, the rusty blotches
in both species are divided into sections by the more or
less dark lines of the nervures, and each section in the
interneural spaces of the fore-wing has a definite form that
differs completely in the two species, and is constant in
every one of the large number I have examined. In stygne
the interneural blotch on its basal margin is stretched out
centrally into a more or less full convex margin, as if the
nervures carried the dark ground-colour into the blotch,
and so forced it to swell out into the intervals. In
palarica, each interneural blotch falls away from the base
in its mid-neural line, forming a more or less deep notch.
This is most marked in the blotch between nerves 3 and 4,
where the ocellus is weak or absent. This blotch is also
shorter than the others as if it failed with the ocellus,
whilst in stygne, though the ocellus may be wanting, this
blotch stretches inwards at least as strongly as the others.
The result is an hour-glass shape of the rusty mark on the
fore-wings, contrasting with the characteristic outline in
stygne. In the ¢ the rusty blotch is continued up to the
costa, with a deflection basewards, by a whitish-grey shade,
of which any trace is wanting in stygne. The blotches on
the hind-wings have a similar character. The blotch of
each ocellus is much the same in both species when it is
reduced, but when it is pronounced and large, it differs
Erebia palarica and Erebia stygne. 15
markedly in the two species. In stygne it extends basally
in a rounded or even pointed projection in the middle of
the interneural space, leaving the nervures, pronounced as
sharp angular interruptions. In palarica the blotches
extend inwards rather along the nervures than centrally,
and the ocelli never bear the aspect of being at the
outer end of the blotch as they so often have in stygne.
Palarica never follows stygne in this character of the hind-
wing, but a few stygne have a little of the tendency of
palarien.
I have one specimen, and one only of £. stygne 2, that
so far traverses some of these distinctive points, that it may
really be a hybrid. It is small like stygne, but has the
inner margin of the blotches of the hind-wings like palarica.
The blotches of the fore-wings are not distinctive, but the
large double apical ocellus is placed in the blotch very
much as in palarica. In stygne these ocelli are usually
nearer the outer than inner margin of the blotch, in
palarica they are quite median.
To resume the distinctive characters of palarica and
stygne, the under-side shows some slight differences. In
both, the under-side of the hind-wing in the ¢ is often
smooth, black and polished, with little or no marking, this
is in fact the rule in stygne, and it is often difficult to make
out the central darker band, and when seen it appears to
have rather a smooth margin. This form is rare in
palarica and generally the central band is quite conspicuous,
and in a few instances has some white markings just out-
side its outer border. This border is always carried
outwards in the interneural spaces, and the general facies
is much more that of evias than of stygne. The markings
are usually very distinct, with stygne one cannot find a
specimen marked enough to make a satisfactory comparison.
The 2 palarica under-side differs from that of stygne, var.
hispanica, in being less brown, more grey, and in being
more distinctly marked, the basal margin of the central
band being obscure or wanting in most stygne. There is,
however, much variation in both species. If the difference
be noted as a slight resemblance in the facies of the under-
side of the hind-wings to evias, wanting in stygne, then
we find that var. penalan ‘w@ of stygne goes nearly as far in
this direction as palarica, and var. bejarensis goes much
further.
A distinctive character is the structure of the clasps of
<r Dr. T. A. Chapman on
the male appendages. If there were no other reason to
separate palarica from stygne than the outline of the
clasps, I should certainly say they were not distinct. Still
there is a difference by which palarica differs from all
forms of stygne I have examined, and in which bejarensis
is distinctly stygne and not palarica. The difference is
shght and difficult to seize, but in some specimens all
mounted in the same way, and at the same time,* of
palarica, and of Pajares stygne, the difference is constant.
In specimens otherwise mounted it would probably express
itself differently. In these preparations, the head of the
clasp in stygne expands gradually from the neck, and is
therefore long and not very broad; the back of the clasp
is In one continuous curve, and there is a distinct notch or
step at the back of the head. In palarica the neck is
narrower and continues narrow, and the back of the clasp
being, for some distance above the neck to the end,
straight and having no notch at the end, the head instead
of having nearly parallel sides, and so of quadrilateral form
as 1n stygne, has the front line at a considerable angle to
the back, and the head looks somewhat triangular. The
terminal serrations are less visible in palarica, being forced
under one edge, instead of being marginal. This shows
that the differences are due in some degree to a different
amount of twisting in mounting, caused by a difference
in form, not necessarily, however, that presented in the
preparations.
The form of stygne with the nearest approach of clasp
form to that of palarica is var. pyrenaica, but it is
distinctly stygne and not palarica.
In palarica the side processes of the tegumen are
constantly though shghtly longer than in stygne. Not-
withstanding the relative sizes of the insects, the clasps of
stygne and of palarica are of almost identical size, that
of stygne, var. bejarensis, being longer.
This is decidedly another character making Mearensis
a variety of stygne, and palarica distinct. Contrary to
what perhaps one would expect, when geographical forms
differ in size, the size of the clasps differs also ; this is very
marked in xhiops where the large continental form has a
clasp large in proportion compared with the British form.
* By dividing the chitinous ring in the central line between the
clasps, and opening it out on the slide, so that the two clasps are at
each end of the preparation.
Krebia palarica and Erebia stygne. i
So if palarica were a variety of stygne, one would expect
to find the clasp large proportionally, instead of, as it is,
just the same size. The ? appendages also differ a little,
especially a hemispherical hollow is rugose in palarica,
much smoother in stygne; but I know so little of these
appendages, not even the names of the several parts, that
I can give no opinion as to the value of the difference, nor
have I examined examples enough to know whether they
are constant,
Palarica being thus differentiated from stygne, one for
the moment forgets the many points of resemblance,
especially the close resemblance of the appendages, and
the general scheme of colour and markings. In both these
respects even, it 1s, however, more distinct from stygne
than ewryale is from ligea, or nerine is from melas (not
lefebvret, which is a very different thing). Are all the
other Spanish forms of stygne, stygne, or are any of them
palarica? When in the field I thought that palarica was
possibly an extreme form of bejarensis trusting merely to
memory, the two points of large size, and dissociation from
evias in which they agreed carried too much weight. A
mere glance at the specimens when together is enough to
show that bejarensis and all the others are stygne, bejar-
ensis is certainly extreme, but perialare, though differing
in some directions, is fairly intermediate between bejarensis
and hispanica.
IT am not sure that I have not too much laboured the
distinctions between pularica and stygne, as 1t would not
surprise me to find that a majority of Rhopalocerists,
looking at the specimens in my boxes, where the constancy
of the two forms in good series is so manifest, and thei
facies so different, would off-hand say they are unquestion-
ably distinct.
The small stygne of the Puerto de Pajares is intermediate
between what I take to be typical stygne and the var.
hispanica. I do not think it enough removed from either
to require a varietal name, though it is a fine, large, bright
form, and is rather hispanica if a name be necessary.
The keynote of ZH. stygne, var. hispanica, was its ap-
proximation in size and markings to a form of evias that
met it half-way in this respect, and that flew along with it.
I believe Mrs. Nicholl’s specimens in the B, M. show that
stygne and evias are similarly associated at the Picos de
Europa, they ought therefore to be similarly associated at
TRANS. ENT. SOC. LOND. 1905—PART I. (MAY) 2
18 Dr. T. A. Chapman on
Pajares. Probably they are. I took one f specimen of evias
at Pajares; it was a form that might very well be associated
with stygne, but as a matter of fact I took it some 200
yards from the nearest stygne ground, at about 5000 ft.,
and I took but the one. Why? it was not in fine
condition but not so bad as to show that the species was
over.
I also took one 2 specimen of evias at a long distance
from the { specimen. It was associated with small stygne
at the highest point at which I found them, a small colony
at 6000 ft. on very steep impracticable rocks, so that I only
took two specimens. vias may have been in greater
proportion to the stygne here as I found one evias to two
stygne, but the whole colony was a very small one, as,
though I could not traverse the rocky slope, I easily got
round it except at one side where it ended in a precipice.
This ground was about 50 ft. above the upper margin of
one of the highest localities for palarica.
Why was ‘evias so rare? Was this a bad season for
high-level evzas? I will explain immediately what I mean
by high-level evias. It was not for want of going over
plenty of ground that I found only these two.
On going rather later (July 24th) to the Guadarrama,
we huuted well the slopes of the Penalara, whence our
President brought one specimen each (taken in 1902) of
stygne and evias, taken together, and so alike as to leave
no doubt they were mimetically associated on the Penalara
as we found them on the Sierra de la Damanda at Canales.
On the Pefialara we found LZ. stygne, var. peralarex, in
considerable numbers, unfortunately in a very poor con-
dition, except on some very rough ground near the top,
where they were less common and almost impossible
to take. As I brought home some 30 specimens I must
have handled 50 or 60 specimens. Amongst them was not
even one evias. I can only suppose evias was for some
reason very scarce. We were not too late to have seen
some old worn specimens, and I specially took many worn
individuals in hopes they might afford an evias; they were
all, however, stygne.
In what I have just said about evias, var. pewialarx, some
one may perhaps think I am casting some doubt on the
President’s specimen, that he got it somehow mixed up.
If I thought so, I should say so, and no one who knows the
President would think such a thing likely. But the
Erelna palarica and Erebia stygne. 19
specimen itself is abundant reply to all such doubts, it is
just such an evias as one would expect to find on Pefialara
with stygne and is not precisely like any other race of
evias. Why did I get none? Why did I get only two at
Pajares? I incline to believe that evias was scarce this
season, but I don’t know. This brings us to another point
which concerns evias, which I cannot do more than open
up, my own observations being too few to settle anything.
It is this, we have found at three or four places in Spain,
a small form of evias associated with stygne, at a fairly
high level, and in each place the two species havea special
similar facies. This form occurs about mid-July. But at
the end of May there occurs apparently over a large part of
Spain, a low-level evias. I have never taken this form,
being always too late for it on my visits to Spain. But I
saw a number this year in collectionsat Madrid. Like the
high-level form, it very probably varies a good deal at
different stations, but those I saw at Madrid were large and
brightly coloured, like large Swiss specimens, such as I
have taken at Locarno, with the rusty marks bright and
ruddy, and not yellowish, as in var. hispanica, and probably
four or five mm. more in expanse than that var. and five
or six more than var. pefalarx. Sr. Zapater records
both forms from the Teruel district, and Mrs. Nicholl has
reported low-level evias from various localities, and there
are other records. What is the relation of these high- and
low-level forms of evias to each other? The low are large,
bright, early, and self-dependent; the higher, smaller,
yellower, later, and associated with stygne. Are they
syngamic ? With only our present light on the matter, I
incline to answer “yes,” but with hesitation; we have no
experiment naturally provided as in the case of £. stygne
and palarica.
I can only repeat that #. stygne and £. evias in Spain
still present many interesting questions for investigation.
High- and low-level evias are to a great extent evias
hispanica and evias evias, but I do not think we have
evidence to justify such an identification. In fact, Mrs,
Nicholl takes evias evias at high levels, and evias hispanica
in Albarracin is not decidedly a high-level form.
Appended is a table of the wing expanse of the varieties
of stygne and evias I have met with, and of palarica—in
some instances founded on too few specimens to be trust-
worthy, still the best I can obtain for comparison.
20 Dr. T. A. Chapman on
I must note that most of my measurements were made
from the insects as set, and are probably two or three mm.
too small. I notice that Ruhl’s measurements quoted seem -
small, and I suspect were made in the same way (from tip
to tip) in insects with the wings much advanced and
therefore in some cases quite five mm. too small.
I add also a note of the varieties of the three species
known to me from Spain.
I was successful in getting both Z. styyne and palarica
to lay eggs freely by sleeving them over grass.
Unfortunately, being on the move prevented my making
such notes of the eggs and young larve as I should have
liked. I sent a number of eggs to Mr. H. Powell, of
Hyeres, whose knowledge of Satyrid larvee is more extensive
than that of any other of my entomological friends. I
append his notes and my own. ‘The difference between
the eggs struck me at first as being considerable, the one
have the minute dots that form the coloured patches in few
and large groups, the other in smaller but more numerous
ones: this conclusion was reached on the first eggs laid by
each species, and were those of one female of each. Later
when I got eggs laid by several others, I found that these
differences were not specific but individual. Each female
laid eggs all alike in these markings, and differing more or
less from those laid by others, but each species seemed to
have practically an identical range of variation in this
respect.
Mr. Powell agrees with me in two points. First, the
eggs of the larger species are rather the smaller, and,
secondly, there would be no difficulty in separating the newly
hatched larvee of the two species if mixed. Curiously,
however, Mr. Powell says that the larva of palarica is
darker than that of stygne (though he says that within the
egg it is lighter), whilst I note the pale lateral stripes as
being rather paler. JI agree with him, however, in the
stripes being better marked.
I agree with him also in another point that is probably
of considerable importance. I supplied both larvee with a
grass (Hesiuca ovina, or something like it) from stygne
ground, and stygne did well on it, and when (at La
Granja) I had to find a substitute (I fancy another var. of
the same grass), they ate that also, and I succeeded in
bringing them home, and believe I have some alive now
(Dec. 1904). The palarica, however, did badly on the
Erebia palarica and Erebia stygne. 21
same grass, and at La Granja, gradually starved themselves,
and died. I got none home alive. This clearly indicates a
different taste in food-plant, as Mr. Powell also points out,
though he was handicapped by a Riviera summer, practi-
eally making grass unobtainable.
Small stygne eggs laid July 14th and 15th, are pearly
white when newly laid, but soon become slightly brownish.
This is due to a general change of colour, but chiefly to a
development of brown patches, each consisting of an
agglomeration of dark dots. These vary very much in their
disposition. In one or two cases the dots are nearly
uniformly distributed (not in patches). The patches may
be four or five in the length or width of the egg, with
small spaces between, or they may be much smaller, so that
there are eight or ten to the length of the egg. On some
the dots are distinctly in regular rows in each patch.
They vary a little in size and shape, about 1°3 mm. high,
and 0°8 wide, a little narrower at the top, but maintaining
width to close to each end. There are 22 ribs, varying
from 20 (one counted) to 24 or perhaps 25. The ribs are
high but not sharp, and may be a little waved. They
never branch or anastomose, but end at top by merging in
an area that looks beaded (high power not available).
The secondary ribs are poorly marked but very distinct
when a suitable light falls on them.
Large stygne (palarica) eggs laid July 16th and 17th,
same as small. They are perhaps those of one @ only, but
they are very uniform in size of red-brown patches, viz.,
about 5 or 6 across egg and ribs seem most usually 24,
but one is found with 21,
In the individual variations of the eggs it is difficult to
be sure of size, but that of palarica seems to be fraction-
ally less than of stygne—1-2 instead of 13 mm. Each ?
seems to lay eggs of a similar facies, with the later
smaller and even stunted and deformed. Later layings by
other 22 of palarica showed the range of variation in the
egg markings to be practically the same in both species.
The eggs of both hatched between July 31st and
August 2nd. The young larvee are very nearly identical.
The large palarica seem the paler in having the dorsal
band slightly narrower and the lateral line is lighter in
colour. It might be called white in the larger (palarica),
yellow in the small (stygne), but this would exaggerate
the difference, |
to
22 . Dr. T. A. Chapman on
The following description was taken some time after
returning home ; it may have some value as a description
of first stage of stygne, but palarica having all died, it is
useless for comparison with that species, still less, of course,
will the later stages have the value hoped for when I
thought I might rear both species.
Erebia stygne (small form), Aug. 24th. Larva full-grown in first
stage, length 4°8 mm., head looks very small, about 0°5 mm. wide,
forward segments being quite 0°9, It tapers very steadily from 2nd
or 3rd abdominal to tail, 9th abdominal segment being about 0°5 mm,
across, 10th smaller.
Ground-colour whitish terra-cotta, a narrow dorsal red-brown band
(or line) broadest at 5th or 6th abdominal. Then a broad pale band,
which includes both I. and II., which are widely apart, one at front,
other at hind-margin of segment, and II. quite twice as far from
middle line as I. Next, a narrow red line, then a pale one, nearly
white, 7. e., decidedly lighter than general ground-colour. Then a
broader dark band, paler than the others owing to a good deal of
marbling of pale ground-colour in it, in this is III. Then a very
narrow whitish band and a very narrow brown (or reddish-brown)
line in which is spiracle. Then a broad yellowish-white band in
which are IV. and V. The anterior much the lower and the line of
spiracle just between them. Then a narrow reddish band, a slightly
broader pale one, and a narrow weakly-coloured dark one. Below
this is the proleg with two hairs at margin, and on Ist and 2nd
abdominal one hair in place of proleg, and another more ventral
(this may be more in situation of proleg), a minute one still more
ventral. No hair or tubercle is found on the three lines below the
band carrying IV. and V. Legs and prolegs pale fleshy, four hairs
and three dots like hairless tubercles on claspers. Head round, with
fine wrinkling, making it rough. The anterior ocellus very large
and prominent ; the second smaller, but very similar and very close
to it, the others flat and inconspicuous. The two first have pigment
in the epicranium, as well as the mass beneath. The head also
carries a number of hairs of the same colourless texture with rough
surface, curved and clubbed, as those of the ordinary tubercles.
The anal plate carries four hairs on conical tubercles along its
posterior margin, and one in either side higher up. On the pro-
thorax is a small plate on either side with four hairs, two in front
and two further back. Further down in line with plate, and well
above, and in front of spiracle, isa double tubercle of which the
front hair is short and curved like the others, but the second though
iransparent and rough (not spiculated) like the others is straight,
Erebia palarica and Evrebia stygne. 23
and four or five times their length. Below isa tubercle with one
very short clubbed hair. This is as much below spiracle as the
double one is above it. Thoracic 2nd and 3rd have I. and IT. single
and in the same transverse line as is III. which is however double,
it is exactly in line with III. abdominal. Below and quite to front
of segment, is a single tubercle. This is below line of spiracle but
above IV. and V. Lower is a single tubercle at base of leg (on pro-
thorax two). Prolegs have six hooks in single line, claspers eight.
The terminal bristle of the antenna rather long, three times the rest
of the antenna, or nearly so. The bristles are rather longer than
elsewhere, round the mouth region. The hairs of tubercles are very
small, curved backwards and slightly clubbed, transparent, and with
a roughened surface, in length, perhaps 0:06 mm.
The following are Mr. Powell’s notes.
“ Hrebia stygne.
“Ova received from Dr. Chapman, July 21st, 1904.
“ The eggs are fixed to blades of grass.
“ Shape. Oval, with a flattened base concave in the centre. The
top is only slightly flattened.
“ Appearance. Pearly with a pinkish tinge, blotched with reddish-
grey. Vertical ribbing quite distinct under hand-lens. Height,
12mm. Greatest width, -9 to 1 mm.
“ Under Microscope x 50. There are 21 (sometimes 22) vertical ribs
running from the edge of the base to the top, where they become
rather lumpy and broken, dying out on the nearly smooth area
around the micropyle. They are blunt-edged, with gently sloping
sides. The cross ribs are numerous, and of course very much
smaller and lower than the others. They constitute the long sides of
cells, the tops of the vertical ribs forming the short sides. On the
top of the egg the cells are large and irregular in shape, triangular,
lozenge-shaped, ete. They diminish very much in size as they near
the centre, in which is the invisible micropyle. There is no depres-
sion at the top, but in a few specimens the centre is occupied by a
low, roughened pimple formed by a bunching-up of the lumpy ends
of the ribs. The blotches observed under the hand-lens are seen to
be composed of pinkish or chocolate-coloured specks arranged in
loose groups.
“These groups are beneath the eggshell. That they
are directly connected with the living contents of the egg
24 Dr. T, A. Chapman on
there is no doubt, for they can be seen to move back-
wards and forwards together, always keeping the same
relative positions. The movement is a fairly brisk one,
reminding one of a shrug of the shoulders. It can be
induced by breathing upon the egg, or by giving a
tap to the support on which it rests, but it occurs from
time to time even when the egg is unmolested. Many
other (perhaps all) Zyvelix ova develop these pinkish
blotches after a few days. I have seen something similar
in Cenonympha. ‘The base of the egg is covered with
large shallow cells of irregular shape. Before hatching the
colour darkens to dull fleshy grey, there being darker and
lighter patches. The groups of specks disappear when
this change takes place, and the shape and markings of
the larva are seen through the shell. The head occupies
all the upper part of the egg, while the body is curled
round horizontally with the extremity turned downwards.
The numerous pits on the head show through as brown
dots. The two groups of beads (ocelli? one on each side)
are very clear, so is the mouth and the arch above it.
The markings of the body (lines and tubercles) are quite
distinct. The larvae commenced to hatch out on July
23rd, and all were out by the evening of July 24th.
They eat away the top of the egg in a circle, emerge
through the hole made, and eat up the rest of the egg or
part of it,
“The newly-hatched larva is 28° mm. long when still, but can reach
3mm. when walking. Width of the head 0°7mm. It is a large
head, and is higher and broader than the body, which tapers to begin
with, down to the forks. The colour is that of most newly-hatched
Satyrids, a sort of pale straw. The shape and markings of the
head are typically Satyrid. It is rounded, rather flattened in front ;
much resembles a lemon rind in pitting and polish. The pits are
large. Depression between the lobes very shallow. The side
‘beads’ (ocelli?) are large, dark brown, and shiny. The usual
small, brown seta patches are very distinct. Sets short, transparent,
curved forward. Those just over the mouth are longest.
“The Ist division of the 1st thoracic segment has 16 small beady-
brown tubercles arranged transversely but not in an exact line. The
2nd and 3rd thoracic segments have both a dorsal dark-brown
tubercle-bead outside the dorsal stripe. In a line with it on the
sub-dorsal stripe they have another bead. The abdominal segments
have the first bead on the 1st sub-division, and in a line with those
Erebia palarica and Erebia styqne. 25
on the thoracic segments. They have the second bead at the other
end of the segment and just above the sub-dorsal stripe. Another
line of large tubercle-beads (one in the centre of each segment and
nearly in a vertical line with the spiracles) runs down the larva
between the spiracular and supra-spiracular stripes. These last
beads are doubled on the 2nd and 3rd thoracic segments. They
generally approach the supra-spiracular stripe, entering its lower
edge on the 6th abdominal segment and remaining in the stripe
afterwards. Above the lateral border are two small beads, the first
in the front of the segment, the other farther back and higher. All
beads have short transparent hairs. The feet and claspers have each
two or three beads bearing longer hairs than those on the body, with
the exception of those growing from the tubercles on the. 8th
abdominal and anal segments.
“There are three conical tubercles on each fork (last segment) point-
ing backwards and giving rise to single, rather long, transparent
hairs. One of these tubercles terminates the fork ; the two others
are lateral. The ground-colour of the body is a pale straw-grey.
The stripes are pinkish-brown. Dorsal line weak on the thoracic
segments, well-marked on abdominals. Sub-dorsal line narrower
but better marked on the thoracic segments. Supra-spiracular and
spiracular both well marked, the former thickening as it runs down
the body, and continued along the edge of the forks. The space
between the lateral edge and the spiracular line is very light.
Beneath the edge, and therefore within the limit of the ventral
surface, is a very narrow pinkish-brown line. The stripes are
broken by the segmental and sub-segmental incisions. The dark
colour does not penetrate the crevices. This is most marked in the
case of the sub-dorsal and supra-spiracular stripes. Ventral surface,
claspers and feet of the usual pale straw-colour. Forks: short,
thick, and lying close. Claspers on last segment large, with several
hair-bearing tubercles. The spiracles are round, dark-brown beads,
no larger than the tubercles above them except on the Ist thoracic
and 8th abdominal segments, but they are rather lighter in colour.
“The larvee ate the grass I was able to give them, but it
was too tough and dry to nourish them properly, so they
soon died. No good grass was obtainable owing to the
drought. In course of time I was able to improve the
grass by watering, but it was then too late.
“Erebia palarica.
“Ova received from Dr. Chapman, July 21st, 1904,
“The eggs are fixed to blades of grass,
26 Dr. T. A. Chapman on.
“As the egg is much like stygne’s I will not go through all the
details again, but will note the points of difference.
“Size. All those observed were distinctly smalier. Height,
10mm. toll mm. Greatest width, 0°85 mm.
“Colour. Whiter.
“ Number of vertical ribs, 22 to 24, none with less than 22.
“The arrangement of cells at the top of the egg is on the whole more
regular, and the top looks smoother, but this is not always the case.
None of the eggs have anything like a central pimple, however, as
seen ina few stygne. I do not consider this is a good distinction,
for I have noticed that in some Satyrids the top of the egg varies in
the same species. S. cordula, for instance, has sometimes a decided
cone around the micropyle owing to the bunching of the lumpy
extremities of the ribs; in other specimens the top is fairly level.
“The specks beneath the shell are less numerous, and rather
smaller. The groups are better defined, not so straggling; they
are browner, not so pink. The same movement is observable.
“Before hatching, the eggs underwent similar changes to those
noted in stygne but without becoming quite so dark. Hatching
commenced on July 28rd, and all the young larve were out by the
afternoon of the next day.
“Larva is smaller than stygne. Length, when first hatched,
2°6 mm., when walking, about 2°8mm. Width of the head, about
0°6 mm.
“The larva is darker in appearance than stygne and
that is on account of the stripes which are more strongly
marked, especially the supra-spiracular stripe. The
grcund-colour is the same in both. If the larve are
mixed together it is quite easy to separate them without
any possibility of a mistake.
“T gave these larvee Brachypodium pinnatum, but was
unable to get any with fresh shoots. I also tried them
with a Festuca which I had in a pot, but it had only a
very few green blades. I tried them with a third grass,
but nothing suited them, and they quickly dried up, with-
out having eaten anything at all, as far as I could see.
They perished several days before the stygne larve.
Stygne ate Brachypodiwm and another grass, and if they
could have got some fresh shoots I have no doubt they
would have lived.”
It may be proper to give a short diagnosis of :
Erebia palariwa,
Colour and markings like EH. stygne, except in the inner margins
Erebia palarica and Erebia stygne. ae
of the elements of the red blotches, being notched and receding,
instead of convex and advancing, in the under-side of ¢ hind-wing
being often more distinctly marked, and in the red blotch in the
Q upper wing being continued to costa as a whitish or greyish
shade. Appendages, head of clasper more triangular and more
definitely marked off by a narrower neck. Expanse 53-64 mm,
Habitat, Cantabrian range at Puerto de Pajares, and Picos de
Europa, and probably elsewhere.
I will place type specimens in the National Collection.
Hrebia stygne, var. pevialare, is diagnosed in Proc. Ent.
Soc., 1904, p. xlvi, from one specimen only. The series
taken this year shows that the race is one of the most
variable of stygne. Some are not very different from
those taken at Pajares. Others are like that described,
and not a few vary even more, so as to seem to be quite
on the way to a form like bejarensis. The 22 show more
markedly than the males the remarkable inward extension
of the red blotches, which is carried so far in bejarensis.
In expanse it is smaller than the hispanica form from
Canales, but larger than that from Pajares.
The two evias taken at Pajares are small, and are of
different types, but are within the limits of variation of
Li. evias, var. hispanica, but a good way from its average
type. The high-level 2 is so very lke the 2 of stygne
taken at the same time and place, that I did not recognize
the specimen as evias, even when I had got it home and
set it, and only discovered it on critically examining the
specimens afterwards.
Mrs. Nicholl has kindly lent me three specimens, which
I may note as a specimen of JZ. stygne of fairly ordinary
type, 48 mm. in expanse, one of several taken on Mont
Seny, 6000 feet, near Barcelona, date not given. Mont
Seny is almost a spur of the Pyrenees; this specimen,
with several taken by Mr. Burr, one over the frontier at
Sallent, May 29th, 1904, proves, what we had no record of
before, but which nevertheless every one supposed to be
the case, that #. stygne occurs on the south as on the
north slope of the Pyrenees, and no doubt more or less
throughout the range, and probably abundant enough.
The other two specimens are in response to my request
to see the most extreme form of evias from the Albarracin
district. One of these, expanse 50 mm.,is a male taken early
in July on the summit of Sierra Camarena (Javalambre,
28 Dr. T, A. Chapman on
6000 feet). This specimen is identical with not a few of
those from Canales. The other is a very dwarf 2 from
Tragacete, June 29th. This is probably a genuine aberration
as regards size, being only 40 mm. in expanse as compared
with smallest from Canales,44 mm., or both measured as
set 38mm. and 41 mm. Otherwise, as regards form,
colour, markings, etc., it agrees exactly with various Canales
specimens. These from their locality are true var. hispanica,
Zapater, and go to show that hispanica is a high-level form
of evias, in the Teruel district as elsewhere.
What we know of the geographical distribution of
these butterflies is too fragmentary to take us very far, but
it is sufficient to afford a few interesting considerations,
Assuming the Erebias to have reached Spain vid the
Pyrenees, whilst it is Just possible that zapateri may have
reached the localities where it has developed its peculiar
character by way of the coast hills, for the most part we
find the Pyrenees so absolutely cut off from the rest of
Spain, so far as mountain forms are concerned, by the
wide and low valley of the Ebro, that the Erebias must
first have travelled westward into the Cantabrian range
before they could circle round the head waters of that
river. It was probably at a very early date that palarica
broke away from stygne, probably as early as the parting
of zapatera from neoridas.
It is, however, with stygne and evias that we are more
concerned. At the Picos de Europa, 180 miles west of
Pyrenees, the two species are still but little differentiated
from their mid-European types, nor is it clear that evias
has a high- anda low-leve! form distinguishable from each
other.
Mrs. Nicholl at Aliva on the high slopes of Pefia vieja,
5000 feet, met on “July 12th with one poor evias, many
stygne. July 14th, one nice 2 climbing Pefia vieja. July
17th, on the Col de las Nieves (at least 6500 feet) a few
evias. July 18th, above Aliva, about 6000 feet, evias
much battered, could scarcely get any good ones.” She
writes that she has in her collection only four specimens
of evias from the Picos, “of which three are 2 9, large and
much spotted; I caught many that I did not keep as they
were over, exceptafew very highup. They are very differ-
ent to the small form from Aragon. The type of stygne was
very common on all the Picos, up to the middle of July,
on the southern and eastern sides of the range; I saw few
Erebia palarica and Erelia stygne. 29
or none afterwards on the west and north faces of the
same mountains, and I saw remarkably few butterflies on
the west and north of the Picos.”
There are, as I have already noted, two Picos evias in
the B. M. collection ; unfortunately they have only general-
ized labels, but one seems to be like the low-level forms,
the other approaches the high-level one.
Going sixty miles further west to Pajares, we find stygne
less abundant, its habitat even restricted, but common
enough where it occurred. The three localities I most
closely examined were, first, the high-level one already
noted (over 6000 feet), I met with no other so high; with
these a definitely high-level form of evias. Second, at the
Puerto and extending for a mile along the north (Asturias)
side of the col, but not apparently reaching very far
either up or down from the 4500 feet level. The aspect
would be N. and N.W. The third locality was nearly two
miles below the col on the south side (Leon) at about
4250 feet, a comparatively small patch of a few hundred
acres, with a westerly aspect.
We took specimens at a few other localities, but not in
circumstances to enable us to say what might be the
extent of the habitat, but generally these places had an
easterly aspect and were at about 4500 or 5000 feet. One
evias of doubtful high-level type not closely associated
with these. These stygne from Pajares make a certain
approach to the hispanica form, so that it is convenient to
call them so, but many individuals are little if at all
removed from the ordinary type.
It would be interesting to know what forms occur
further west in the corner of Spain to the north of
Portugal and in Portugal itself, but for the present this is
a blank. The next point is at Canales to the south of the
Ebro. Here stygne and evias are both modified by way
of approach to each other, and fly together in about equal
numbers. vias probably has a low-level form here. ‘The
high-level one with stygne is of the form hispanica, but
only a few reach typical hispanica, others still retain
something of ordinary evias. The stygne is less vari-
able than at Pajares, is two mm. larger and averages
larger than the associated evias. Forty miles east of
this, at Moncayo, the same form of evias is found, but we
saw no stygne; possibly it does not occur there, as stygne
seems to fail eastwardly, evias towards the west.
30 Dr. T, A. Chapman on
Passing now to the Guadarrama, 8. W. from Canales some-
thing more than 100 miles, and at an elevation of 6000-
8700 feet, we find a still more specialized form of stygne,
very variable, from forins nearly typical to the peculiar
penalare and to forms as bright as bejarensis. These are
associated with an equally specialized high-level evias of
rather small size, and very different from the low-level
form of the same region.
A hundred miles to the west of this is Bejar, at about
5000 feet. Stygne occurs in a very large highly-coloured
form, but there is no record of evias with it. Neverthe-
less the association of the two species in the Pefialara
and the approach made by many specimens of stygne
there to the var. bejarensis, seems to lend a little further
plausibility to my suggestion that stygne, var. bejarensis,
owes its largeness and brightness to association with
evias, though that species appears to have failed to go
(at least in a high-level form) so far to the south and
west.
A hundred miles east of Madrid in the Albarracin and
neighbouring Sierras occurs evias apparently in ordinary
large red form and in the special small yellow form,
hispanica. I have no information to show whether these
are in any sense a high- and low-level form. It seems
certain that there is no stygne here. And a high-level
evias apparently implies stygne. Nevertheless, I will
venture the suggestion that evias hispanica in the Teruel
district has, as it were, got there vid stygne, even though
that species could not follow, just the converse of what I
suggest about stygne bejarensis to the west. The resem-
blance of the two forms of stygne, and the fairly con-
tinuous range of Sierra, from Pefalara to Bejar, would
suggest that Bejar was reached vid Guadarrama; whilst the
Teruel district was reached from the Canales area, vid the
head-waters of the Duero and the Jalon, would be the
conclusion for similar considerations in the case of evias.
I saw no stygne in the Albarracin Sierra, and incline to
believe it is really absent, the Sierra not being lofty
enough, as it hardly comes below 7000 feet on the Guad-
arrama in same latitude; so far east in this latitude it is
also possibly too dry.
Erebia palarica : Picos de Europa, 1902. Nicholl.
is 43 , : Puerto de Pajares. 1904, T. A.C,
Lrebia palarica and Erebia stygne. 31
Erebia stygne stygne
” ” ” or \
ah » Aispanica
” ” ” x
” ” ” °
3 » petalare ++
Erebia evias evias
” » hispanica -x
” ” ” OTs
” ” 5» OO.
” ” penalare ++
» stygne bejarensis oo
x associated.
ie) bb)
In Pyrenees (Spanish side), Burr, 1904.
Picos de Europa. Nicholl.
Puerto de Pajares. T. A. C.
Canales de la Sierra. T. A. C.
Guadarrama. Spanish Collection.
Poulton. 1902. T.A.C. 1904.
Picos, Guadarrama, Teruel and probably
many other places in Spain,
Approaching Puerto de Pajares. T. A.C,
Canales de la Sierra.
Teruel.
Guadarrama. Poulton, 1902.
Bejar. T. A.C.
oo apparently unassociated with any form of the other species.
I add a Table of the expanses of the various forms of
these species so far as my material allows.
32 Dr. T. A. Chapman on
* * * * *
Stygne. | Stygne. | Evias. | Stygne.| Evias. | Stygne.| Evias. | Evias. Stygne.
SwIss. CANALES. | PENALARA. Prcos DE Europa, PAJARES.
RP RCTES SGA AS Ae lod: oe? | aaa Grete erent cena
38 1 nh sa |
39 |
40 = |
5
41 ea |. ood
42 Bnet | 1
43 | ee 4
44 1 ian || i
45 1 eal Nak ate | 1 el 9
46 A al ou eZ 1 ° lo (ies:
| @
t )
47 2 pil 8 wad Ne 8} le 2 13° 3
| @
°
48 as 11 TS I Depart § 4
e °
49 A 6@ 2| 15 Md pote 4 | 4
®
50 Begs yh ak Veal ele ae 1 an
51 peealt a? tiie dal | | @ 1
52 3 2 4 | te
53 1
54
55
56
57 |
58 |
59
60
61
peek aes | 50 a7. | ae eae Pe mee 55 11
specimens
Mean [42-0 46-0/49-0 49-4148-3 48-2147-2 47-847-0 — |4e-2 — lag-5 — 46 45 46°24 47-0
expanse
* In these columns measurements are taken without allowance for setting, true expanses are therefore nearly 2 mm,
Mean expanse, or a solitary specimen.
*
Stygne. | Bejarensis.
Hicu
LEVEL BEJAR.
PAJARES.
See Sy) Se
Bs
if
@
al a0 2
le 4
e
3
il 3
1 1 e
2h) 2,
2
] 3
‘Picos.
Br 8 1
3 9 1
J 6
7 a
: 3
Besa 47°649°7 52°
|
Evias.
ACCORDING 'TO
RvuHL.
Swiss.
er 8 se
a
Ee
on Se
a ete By
Sy) hare Se cen
3 | 38 |3,\=
a 2 oS 8
Oo =~
=A Mar eg
i=)
|
er]
ie od
9 |)
Vit Patt
|
; ye
2 + |
ee | |
52-0 49 |
iter than set down. c.g. Several palasica accurately me
+ Two largest Swiss in B.M.
TRANS. ENT. SOC. LOND. 1905.—PART I.
Erebia palarica and Erebia stygne. 33
Small Swiss stygne
in B.M.
in B. ‘
e evias penalare
type 47.
° stygne penalare
type 44.
Largest Pyrenean (Irench)
stygne in B.M.
°
vasured are found to be 63 and 64 mim. in expanse.
(MAY) 3
34
idives Th.
Fie.
Euplanation of Plates.
EXPLANATION OF PuiatEs II, III, IV, V, VI.
Puate IT,
Erebia palarica, 4 upper side.
2, . ‘i 4 MLCT age
3. ” ” S upper ”
4. 3 a », under |,
5 » stygne from Puerto de Pajares, ¢ upper side.
6. ” ”? ” ” ” ” under ”
if 0 ” ” be] ” ? upper ”
8. » » ” ” Under,
9. 5 evias, var. hispanica, from Canales, 2.
10. * s 5 53 » Pajares, 9, only specimen
of high-level form taken.
PLATE ITI.
1. LHrebia stygne, 6 Swiss, upper side.
2. » evias, g@ Digne Bae 5
3. » stygne, var. hispanica, ¢ , Canales, upper side.
= +b) 9 ) ” g ” oy) ”
5. » evias, var. hispanica, g ‘. 3 -
6. ” ” ” ” 2 ” ” ”
ae » stygne, var. pefialare, g upper side.
8. ” ” ” ” g ” ”
9. - % ik * Q under ,,
10. a3 » var. bejarensis, ¢ upper ,,
ll. ” ” ” ” g ” ”
12. ¥ a _ - ? under ,,
Explanation of Plates, 35
PLATES,
As a Plate this is open to criticism, but it shows what it is desired
to do. Figs. 1 and 2 are LH. styyne, and 3 and 4 E. evias, taken
by Mrs. Nicholl at the Picos de Europa in 1902. The two species
show some little approach to each other, but one at least of the
evias is clearly the ordinary form, and these and other specimens
leave it doubtful whether H. evias has here made any recognizable
progress into the division into a large, bright, low-level, early
form (evias) and a small, pale, high-level, late form (hispanica).
Figs. 5 and 6 are Professor Poulton’s two specimens, one evias,
one stygne, from Penalara. The evias (var. pefialare) is clearly a
high-level form, but specially varied to agree with the stygne (var.
penalare) rather than quite like var. hispanica.
Prare: Vv,
Sketch map of the northern half of Spain, showing habitats of Erebia
palarica and of EH. stygne and evias and their varieties. The lines
connecting the stations for evias and stygne mark the progressive
variation of EH. stygne to successively vars, hispanica (Canales),
penalare (Penalara) and bejarensis (Bejar), and of evias to vars, -
hispanicw (Canales and Albarracin) and pejialare (Pefialara).
Low-level (type) evias is more widespread (as the others probably
are also) than is indicated on the map.
Prarn Vi.
Fic. 1. Eggs of Erebia palarica x 10.
2. ,, stygne (Puerto de Pajares) x 10, both from
phoioarapie by A. E. Tonge, Esq.
Clasp of E. palarica x 24.
4, - Ei. stygne x 24.
oo
ohdeg hen
Bane hail’ ject ,
J. Oot |
' Hay
ae | Soar
Hee eel»
5
Say x ws ast bas
4
Crate)
IV. Another Entomological Excursion to Spain. By GEORGE
CHARLES CHAMPION, F.Z.S., and Dr. THomas
ALGERNON CHAPMAN, M.D., F.Z.8.; with descrip-
tions of two new species of Hemiptera, by Prof.
O. M. REUTER.
[Read February Ist, 1905.]
Puate VII.
THE Puerto de Pajares is by no means unknown to
naturalists, and even entomologists have visited the
district, but so far as we know no lepidopterists. have
examined it. Herr Lucas von Heyden gives a rather full
note on it in the account of his journey in Spain in 1865,
in company with other entomologists (all coleopterists),
which was published in 1870 by the Berlin Entomological
Society. He especially dwells on the amenities of his
quarters at Arbas, which was his pied-d-terre for the region,
which reads very humorously. He found adequate quarters
in the muleteers’ cabin opposite the priest’s residence,
where men and beasts shared the lodging, and, the door
being open, sudden charges of cats, fowls, and pigs were
matters of course throughout the night. Arbas, Albas,
Arvas, Abbas, seem to be varied spellings, of which Arbas
is probably most correct and Arvas would give in English
nearly the correct pronunciation. This place is about a
mile to the south of the Puerto by the roadside, and
consists of an old church or chapel with a priest’s residence,
a variety of cowsheds, of which one long row has living-
rooms over and looks like a row in.a colliery district. The
Puerto de Pajares is at an elevation of about 4500 ft. and
we selected it for our excursion, hoping to get lodgings at
Arbas or at Busdongo (the town and railway station to the
south of the Pass is at about 4200 ft.), or perchance at
Pajares (the station to the north side of the Pass, the
town being, however, below the station and rather below
4000 ft.).
On arrival at Busdongo, we heard we might perhaps be
put up at the “tienda” of Sefior Francisco Alonzo, at the
Puerto itself, and it turned out that this establishment
possessed three bedrooms for travellers, and though small
and over the stables and piggeries, they served our purpose,
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY)
388 Mr. G. C. Champion’s and Dr. T. A. Chapman’s
with less call on our philosophy than Herr von Heyden
must have suffered. We remained here from July 3th—
22nd.
Coming up from the south at Leon, the railway follows
the valley, and has to make hardly any twists or spirals,
to reach the Perruca tunnel, by which it pierces the col.
But on the north face it emerges on a steep slope, and
has to make many curves and zigzags to fall some 2500 ft.
in half-a-dozen miles, running in fact over twenty actual
miles of line. At the Puerto the road south follows the
bottom of the valley at a regular and easy gradient. On the
north side the floor of this valley is some 2000 ft. below
and looks as if one could throw a stone into it. The
mountains on the north side have very steep slopes, with
bold outlines and rocky and precipitous summits, and
there are many picturesque rocky outcrops. Although
there are some bold rocks here and there in the southern
valleys, generally speaking the hills are rounded ‘and
grassy, but often with very steep flanks. They have very
little wood on the south side. On the north there is much
beech wood, mixed with birch in the higher ground, little
larger than coppice towards the Puerto, but with actual
timber lower down. ‘The region seems to be destitute of
pine forests.
The general aspect of the country, as to physical
features and vegetation, was more like portions of the
Scotch Highlands than anything in the central portion of
Spain that we visited. The country is, in fact, Atlantic
and not Mediterranean, and has a rainfall that is not
lacking at any season of the year, more than occasionally
happens even in Scotland. Going into detail, however,
the flora of course presented many plants unknown in the
north, though heather, gorse, various pasture and bog
grasses and other plants, such as Senecio, etc., were actually
or nearly identical.
During most of our stay, we were much interested for
practical as well as other reasons in a curious daily cycle
that obtained. The early morning would be brilliantly
fine, but by 8 or 9 o’clock, though it continued so to the
south, the northern valleys would be filled by a sea of fog
to within 500 or 1000 feet of the level of the Pass, with
mountains and ridges standing out of it like islands, a
light south wind blowing over the col; gradually the fog
would rise like a flowing tide, and at length would reach
Yntomological Excursion to Spain. 39
the level of the col, when it would pour over and down
the valley with a north current. This would take place
about 1 or 2 o'clock, earlier or later on different days.
The fog would still rise with a steady north wind,
gradually covering everything; in an hour or so the fog
would be very wetting, and a little later would be actual
rain, and this would usually continue till late in the night.
The change of wind from south to north when the fog
reached the level of the Pass was quite sudden and
remarkable.
On the north side, an easy walk took us down to quite
low ground, affording such insects as MZ. galathea, 4.
hyperanthus, L. argiades, and other low-level species, but
we did not find anything of note. It was not therefore
an attractive excursion, and, easy as it was to go down,
the return was a very different matter.
In any other direction, one could not descend more than
two or three hundred feet and then only to Busdongo, and
usually we went up the valleys, on to the cols and hill tops
above them, that ran down towards the main valley, of
which there were several joining near Arbas, which was,
in fact, very centrally situated for the best of these.
Apart from Hrebia stygne, E. evias, and EF. palarica
noted elsewhere, we met with Z. epiphron, H. ewryale, and
E. tyndarus; of the latter some specimens were of fairly
normal (Swiss) form, but a fair proportion had more than
the usual twin eye-spots, and one or two were larger and
made a fair approach to var. ottomana. Of twenty-six
specimens brought home, thirteen have more than the
twin eye-spot, sometimes the one beneath these, sometimes
the next lower, sometimes on one side only. Two specimens
have both these spots and one of these measures 44 mm.
in expanse. In six specimens these extra eye-spots are
white-pupilled. The normal double spot has large white
pupils in all specimens.
The Z. ewryale are 45 mm. in expanse (42 mm, smallest,
48 mm. largest). In two (out of twenty-two all ? g) there
is a trace of white pupils to the eye-spots, two have four
eye-spots in fore-wings and one has only two, the rest have
the usual twin spots, one or two have the orange-red much
as in var. circellaris.
The £. epiphron are rather close to var. cassiope, but are
large for that variety.
Amongst the moths observed was Oleogene peletieraria
40 Mr. G. C. Champion’s and Dr. T. A. Chapman’s
the white female of which was rare, but the nearly black
males were frequent enough at various points near the
Puerto amongst gorse and heath. This species has
previously been recorded apparently only from the Pyrenees
and on the French side.
Platyptilia isodactyla was common in the bogs near the
Puerto, and some larve were reared from the egg to nearly
full-grown, but died of starvation at La Granja.
Parnassius apollo was seen in many places, but was
common on the débris from the railway tunnel at about
4200 feet. It was rather over (July 11th).
The abundance of a Psyche larva on certain slopes and
pastures at about 6500 to 7000 feet was phenomenal, the
cases were very like those of Pachytelia villosella, but are
believed to be Amicta febretta; a number were brought
home, but none have been reared, the larve moult into
a colourless instar, preliminary to (hibernation and ?)
pupation. One became a pupa. The package containing
these larvee was lost on the way home and turned up three
weeks later; it is comforting to attribute our want of
success with them to this circumstance, but it is not
perhaps certain. Perhaps they meant hibernating, and we
may have some still alive doing so. It was a remarkable
sight to see forty or fifty of these large cases within
a square yard or so, either on the grass or on shrubs of
Cytisus purgans; the larvee did not seem very. particular
as to what they ate.
Mecyna polygonalis was one of the commonest moths
seen, and varied from having the upper wings very dark,
almost black, to pale yellowish, and some specimens
were beautifully marked.
The following is a list of some of the species of
Lepidoptera observed at Puerto de Pajares, and it will
suffice to show the absence of specially Spanish forms.
Melanargia galathea Vanessa polychloros
Erebia palarica Argynnis selene
» stygne » aglaia
» evias » lathonia
» euryale Oolias edusa
» tyndarus uh hyale
epiphron Pieris rapx
Pararge mera » nape
Cenonympha arcanius » Drassice
%, pamphilus 5, daplidice
Aphantopus hyperanthus Leptidia sinapis
Epinephele tithonus Parnassius apollo
Entomological Excursion to Spain. 41
Polyommatus gordius Cyaniris argiolus
iy phlxas Hesperia malve
“ virgaurex, var. 9 abverns
miegit Carcharodus alcec
tf dorilis Adopxa actxon
Lampides telicanus
» betica Procris geryon
Lycena astrarche Nemeophila plantaginis
» hylas Lasiocampa quercus
» corydon Plusia iota
a anton Heliothis peltigera
» argiades Catocala conversa
» warus Hepialus velleda
At La Granja (San Ildefonso, July 23rd—Aug. 3rd)
we were too late for most things, but the impression left
was that it was a most prolific station for many interesting
species, and varieties, some of which are not very common
but would be regarded as rarer but for their abundance in
this fairly well-known locality. Argynnis adippe, var.
chlorodippe and cleodippe, and A. paphia, var. anargyra,
were both abundant but nearly over. elanargia lachesis,
var. cataleuca, perhaps a dozen seen. Polyommatus virgauree,
var. miegii,rather a brilliant form. Lvosopis roboris, common
in the Palace grounds. Cawnonympha iphioides, at 5000—
6500 feet.
Satyrus actwe abundant at Navacerrada and on a slope
of Pefialara at nearly 7000 feet. At this station one very
large specimen (rather worn while the others were fresh)
was taken, that suggested (in size) cordula much more
than actwa.
Orgyia aurolimbata, larvee, pup, and imagines were
common at the Puerto de Reventon (7000 feet).
Erebias and Heterogynis have been elsewhere referred to.
Cledeobia angustalis and Rhodaria sanguinalis were in
many places perfect pests making it difficult to detect
amongst them any other species.
On obtaining eggs of P. virgaurex, var. miegii, we found
that its food-plant, Rwmea acetosa, was over for the season
and growing material was obtained with difficulty. The
butterflies, however, avoided the green plant and laid their
eggs on the most mature and actually dead stems. This
habit accords with the fact that though the larva develops
in the egg in the autumn, the latter does not then hatch,
but remains over till the spring.
The following are some of the butterflies seen at La
Granja:
42. Mr, G. C. Champion’s and Dr. T. A. Chapman’s
Argynnis aglaia Ceenonympha arcanius
. lathonia - iphioides
» paphia, var. anargyra Thecla ilicis
5 adippe, vars. chloro- » Quercus
dippe and cleodippe Lxosopis roboris
- niobe Polyommatus hippothoe
daphne “4 gordius
Melanarg gia lachesis 5 phleas
» var. cataleuca ' virgaurex, Var.
5 dapygia miegit
Hrebia stygne, var. peiialare Lampides betica
EKpinephele tithonus as telicanus
Satyrus semele Lycena icarus
» actxa » astrarche
» statilinus 3 argyrognomon
Canonympha pamphilus Auyiades comma
Amongst the Coleoptera observed at the Puerto de
Pajares, Lylia vesicatoria (which is usually of a cupreous tint
in these southern localities) was perhaps the most in
evidence, this insect swarming in places on the flowers of the
Genista, as did two or three species of Zonabris. On this
plant, too, or on broom, there were also to be found, Corym-
bites hematodes, var., Athous reynose?, various Telephori,
Lthagonycha, and Dasytes, Cryptocephalus lusitanicus ( vary-
ing to almost entirely black, and sometimes seen on heath
also), C. vittatus, Gynandrophthalma concolor and G. reyt,
Anthophagus muticus and sp.? (in numbers, insects not
seen on the Bejar or the Guadarrama, or on Moncayo),
Antholium spp., Phyllobius tuberculifer, Strophosomus spp.,
etc. About the patches of snow, under stones, etc., were,
Lemlidium pyreneum (which seems to be scarcely more
than a form of the Alpine B. glaciale), in abundance,
Baris nivalis, Deltomerus nebrioides (very rarely), Zabrus
neglectus ?, Steropus lacordairer, Ch ymindis melanocephala,
Stenus guynemeri, ete. On a mountain ridge, or Puerto,
at about 6500 feet elevation, amongst short turf, under dry
dung, etc., many interesting insects occurred, as Homa-
pterus punctulatus (taken by us previously on the Bejar),
Pedilophorus metallicus, in abundance, but mostly dead,
Helophorus porculus, Hypera biglobosa (singly), Otiorrhynchus
asturiensis, a Timar cha, Chrysomela rufofemorata and C.
cantabrica, ete. Under stones in various places on the
mountains, were Carabus macrocephalus (very rarely ),
Harpalus cardioderus, Nebria sobrina, various Calathi,
Synuchus nivalis, Amara equestris, Notriophilus aquaticus,
etc.; and on the paths, or amongst the heath, Cicindela
Entomological Excursion to Spain. 43
sylvatica, two species of Dorcadion (one of them worn
and nearly over), Platycerus spinifer (one specimen,
probably knocked off the Genista), and others. In
dung, various Aphodit were to be found, as A. obscwrus,
A. scrutator, A. bonvouloiri, and others, also Ammecius
frigidus, Geotrupes pyrenvus, Emus hirtus, and Staphylinus
pubescens. Boggy places on the heath produced Hlaphrus
uliginosus, Anchomenus sexpunctatus, Carabus arvensis %, ete.
On the Pass, two species of Rhizotrogus, R. felicitanus and
R. fuscus, were occasionally seen in abundance on the wing
in the forenoon, disappearing after midday. A descent
into the valley cn the northern side resulted in the capture
of Perileptus arcolatus, on the banks of the river, Aromia
moschata, var. thoracica, Leptura fontenayi, Strangalia
attenuata, etc. As usual, in all these Spanish mountains,
a Henicopus (H. heydeni) abounded, the males clinging
tightly to the grass stems and sometimes accompanied by
individuals of the opposite sex.
List of CoLEOPTERA found at the Puerto de Pajares.*
Cicindela sylvatica, L., not rare, and once seen captured
by a large Asilid ; C. campestris, L. Carabus macrocephalus,
Dej., three specimens, on the mountains ; C. deyrollez, Gory
(? = arvensis, L.), in the marshy ground near the road;
C. lineatus, Dej.?, fragments of a single specimen found on
the mountains. Notiophilus aquaticus, L., and N. palustris,
Duft. * Hlaphrus uliginosus, F., one specimen, in the
marshy ground near the road: we have not seen a record
of this species from Spain. Nebria sobrina, Schauf., rarely.
Bembidium lampros, Herbst; B. pyreneum, Dej., in
abundance, at the edge of the snow on the mountains; B.
stomoides, Dej.; B. nitiduluwm, Marsh.; B. sp. n. Perileptus
areolatus, Creutz., on the banks of the river below Pajares.
* Deltomerus nebrioides, Vuillefr., two specimens, on the
mountains, near running water, issuing from the lowest
patches of snow. * Platynus sexpunctatus, L., not rare in the
marsh by the road. Synuchus nivalis, Panz., Pterostichus
nigrita, F.; P. cristatus, Duf., var. cantabricus, Schauf.
Haptoderus cantabricus, Schauf. Orthomus barbarus, Dej.
Pecilus cupreus, L. Steropus lacordairei, Putz. Amara
equestris, Duft. * Zabrus neglectus, Schaum, var. ?, not rare,
* The species marked with an asterisk are not included in
v. Heyden’s Asturian list.—Deutsche ent. Zeitschr. xxiv, pp. 281-
304 (1880).
44° Mr. G. C. Champion’s and Dr. T, A. Chapman’s
under stones on the mountains; Z. astwriensis, Heyd., one
specimen. * Harpalus cardioderus, Putz.; H. honestus,
Duft.; H. rubripes, Duft.; H. psittaceus, Fourer., etc., under
stones, on the mountains. * Chlenius velutinus, Duft.
Incinus planicollis, Fauv. Cymindis melanocephala, De}.
Limnebius nitidus, Marsh.? * + Helophorus porculus,
Bedel, at roots of grass, high up on the mountains; ZH.
glacialis, Villa, a species occurring on many of the
mountains in Spain.
Aleochara fuscipes, F. *Staphylinus pubescens, De G.
* Emus hirtus, L. * Leistotrophus murinus, L. * Actobius
prolixus, Er, Philonthus splendens, F., in dung; * P.
Sulvipes, F., var. varipes, Rey, in a marshy place. * Stenus
guynemert, ‘Duv., one specimen, near running water, as
usual; SS. carbonarius, Gyll. Oxytelus laqueatus, Marsh.
* Anthophagus muticus, Kies., and A. n. sp.? (near pyrenxus,
Bris.), both in numbers, on Genista, etc., females only of
the second species obtained. Omaliwm florale, Payk.; 0.
cxsum, Grav. Lesteva longelytrata, Goeze. Philorhinum
nigriventre, Rosenh. Anthobiwm torguwatwm, Marsh., and
A. adustwm, Kies., in plenty on Genista.
Sulpha nigrita, Cr. Ledilophorus metallicus, Chevr., in
numbers (but mostly dead) on a grassy mountain ridge,
under dry dung, stones, ete. Byrrhus pilula, L., and B.
depilis, Gr aells: this latter seems -to be nothing more than
an abraded form of B. pilula. Hister unicolor, L. ; H.
carbonarius, Ill. Saprinus eneus, F.
* Platycer us spinifer, Schauf., one specimen: apparently
very rare in the Cantabrian Mountains, as it is not
mentioned by v. Heyden. Onthophagus Sracticornas,
Preyssl., O. lemur, F., O. furcatus, F., Aphodius fuossor, L.,
A. scrutator, Herbst, A. fetens, F., A. hemorrhoidalis, L.,
A. rufescens, F., A. obscwrus, F. (in profusion), A. rufipes,
L., A. depressus, Kug., A. bonvouloirt, Harold, more or less
common in dung on the mountains. Ammecius frigidus,
Bris., in abundance, with the preceding. Geotrupes
p yrenseus, Charp. ; G. typheus, L. Lhizotrogus felicitanus,
Reitt., and LZ. fuscus, Scop., both in profusion on the wing
in the hot sun in the forenoon, apparently males only
obtained ; &. solstitialis, L. * Hymenoplia chevrolati, Muls.
* Anisoplia betica, Er.
Betarmon vittatus, Cand., six specimens on Grenista.
t This insect also occurs at Albarracin. It has been recorded by
Bedel from Scotland !
Entomological Excursion to Spain. 45
Melanotus tenebrosus, Kr. Athous reynosex, Bris.?, A. cervicolor,
Heyd., and various other species of the genus. Limoniis
nigripes, Gyll. Corymbites latus, F., C. pyrenxus, Charp. 2,
and * (. paulinoi, Desbr., rarely. C. hematodes, F., var. ?,
about a dozen specimens, all females, varying greatly in
size. Hydrocyphon deflexicollis, Miill., two specimens, by
the stream in the valley.
* Tampyris noctiluca, V.., females only obtained. Tele-
phorus abdominalis, F., T. rufus, L., 7. bivittatus, Mars.,
Rhagonycha cantabrica, Heyd., Rk. geniste, Kies. R.
hesperica, Baudi, and others, mostly on Genista. Antholinus
amictus, Er., on Genista. Henicopus heydeni, Kies., in
plenty, on grass-stems: there seem to be two forms of the
female of this species wherever it occurs—one with black,
the other with grey hair. Dasytes spp.?, Danacxa spp. ?
Feliopates lusitanicus, Herbst. ? * Cteniopus sulphureus,
L. Jsomira hispanica, Kies.?, I. murina, L., var., both on
Genista. Mordella aculeata, L. Anthicus antherinus, L.
* Meloe brevicollis, Panz. Zonabris sobrina, Graells, and
* Lytta vesicatoria, L., in profusion, on Genista. Cdemera
nobilis, Scop., *O. podagrarix, L.
Otiorrhynchus astwriensis, Chevr., not rare on the
mountains ; O. pyrenvus, Gyll., one specimen. * Homapterus
punctulatus, Bris., not rare, on a mountain ridge, under
dry dung, ete.; we had previously met with this insect on
the Sierra de Bejar.t Phyllobius tuberculifer, Chevr., on
Genista. Strophosomus spp. Barynotus pyreneus, Bris.,
one worn specimen. Sitones regentsteinensis, Herbst.
Hypera biglobosa, Kirsch, one specimen. Cleonus nagrosu-
turatus,Goeze. Anisorrhynchus hespericus, Desbr. * Baris
nivalis, Bris., one specimen. Ceuthorrhynchus eric, Gyll.
Cionus blattari#, F. Apion ulicis,Forst. Rhynchites betule, L.
Leptura fontenayi, Muls., Strangalia attenuata, L., and
Avromia moschata, L., var. thoracica, Fisch., on the road to
Puente de los Fierros. Dorcadion castilianum, Chevr. 2,
and D. sp.? (mostly rubbed and over), occasionally on
the mountain paths.
Lema lichenis, Voet. Labidostomis lusitanica, Germ.
* Gynandrophthalma concolor, ¥., common, and * G. reyi,
Bris., rarely, on Genista. Cryptocephalus lusitanicus, Suttr.,
many specimens, on Genista, ete., all belonging to dark
varieties, some of them wholly black, with the exception
of two minute spots on the head, the lateral margins of the
+ Recorded by us from Bejar under the name of H. affinis, Chevr.
46 Mr. G. C. Champion’s and Dr. T. A. Chapman’s
prothorax and the scutellum; *C. vittatus, F.; C. sericeus,
L.; C. tebialis, Bris.; C. mystacatus, Suffr.; C. moran,
L., and vars. Gastroidea janthina, Suffr., with its larva,
on Rumex, along the roadside, at the summit of the Pass.
Timarcha asturvensis, Kr. Chrysomela hemoptera, L., C.
rufofemorata, Heyd., C. cantabrica, Heyd., on the moun-
tains; C. menthastri, Suffr. * Orina cacaliw, Schr., two
specimens, * Phytodecta variabilis, Oliv., P. olivacea, Forst.
Phyllodecta vitelline, L. Luperus nigrofasciatus, Goeze,
common on Grenista. Luperus sp. * Malacosoma lusitani-
cum, L. Haltica sp.?, common on heath.
List of HEMIPTERA-HETEROPTERA found at Puerto de
Pajares,
Carpocoris purpuripennis, De G. Therapha hyoscyam,
L. Lygeus equestris, L., L. pandurus, Scop. Geocoris
ater, F., under stones, on the mountains. Orthostira
macrophthalma, Fieb., with the preceding. Nysius
senecionis, Schall. Stygnocoris fuligineus, Fourer. Hm-
blethus angustus, Mont. Aphanus pint, L. Heterogaster
artemisie, Schill. Harpactor iracundus, Poda. Nabis
rugosus, L. Calocoris sexguttatus, F. Pachytomella cursi-
tans, Reut., n. sp., in profusion, amongst short grass on the
mountains. Capsus scutellaris, F. Strongylocoris leucoce-
phalus, L., var. steganoides, Sablb. Heterocordylus tibialis,
Hahn.
In the immediate vicinity of La Granja there were but
few beetles to be found at the end of July, and even on the
mountains the characteristic Dorcadions were almost over.
We obtained, however, an interesting Cryptocephalus, C.
5-punctatus, and Cneorrhinus pyriformis, on young oaks,
on the “flat” east of the Palace Gardens. At the sources
of the small streams, in the upper part of the pine forest,
under chips and stones, we found the very local Leistus
constrictus, Carabus ghilianu, C. guadarramus, and C. helluo,
Steropus ghilianii, Haptoderus nemoralis, and others.
Higher up, above the pine forest, at or near the sources
of the streams issuing from the patches of melting snow,
we again met with the Lezstus, and here accompanied by
Nebria vuillefroyi (mostly immature), Zabrus seidlitzi,
Otiorrhynchus truncatellus, Bembidium ibericum, B. laterale,
ete. Around the Laguna de los Pajaros, on the south side
of the Pefalara, on which there were various patches of
snow, a variety of interesting species were obtained,
Entomological Excursion to Spain. 47
including a Pedilophorus (apparently new), Bembidiwm
carpetanum (at the edge of the melting snow, in abundance),
Dichotrachelus graellsi, Trechus pandellei ?, 2, Cathormiocerus
chevrolati, etc., and in the lagoon itself Esolus angustatus,
Lareynia carinata, Deronectes griseostriatus, and others.
Myriads of winged ants were sometimes found floating on
the surface of this lagoon, probably carried there by the
wind, On the short turf hereabouts, from which a large
number of vultures (looking like a flock of sheep, as seen
from the ridge above) were disturbed on one of our visits,
a Timarcha abounded, and the two local Dorcadions (so
abundant in early summer) were still represented by a few
worn examples. Near the Puerto de Reventon the brilliant
Geotrupes coruscans was seen in numbers, flying in the hot
sunshine, and at the Puerto de Navacerrada Cyrtonus
montanus, Misolampus scabricollis, ete.,occurred under stones.
The pine stumps and logs produced three species of Liodes
(as in Scotland), Zemosthenes pinicola, Spondylis bupres-
tordes, Sphindus dubius, a Bothrideres, a Plegaderus, Paro-
malus flavicornis, Enicmus rugosus, Lygistopterus sanguineus,
ete. About a waterfall, on the way to the Reventon,
Geodromicus nigrita was taken, in company with various
species of Hydroporus , Helophorus, Bembidium, ete. Chlenius
dives was occasionally seen, and a 7 hylacites occurred not
uncommonly, as usual under stones exposed to the full glare
of the sun. On flowers but few insects were to be found
beyond Cerocoma schreberi, one or two Zonabris, and Leptura
stragulata. ‘Though too late for most of the beetles, we
were, however, at the right time for the Capsidex, various
interesting species of Phytocovis being plentiful on oak, etc.,
though very difficult to catch when beaten into an umbrella.
Part of our success here was due to the kind assistance of our
old friend Sefior Ignacio Bolivar, whom we were fortunate
enough to meet on one of our excursions to the Sierra.
List of CoLEOPTERA found at La Granja (including the
Peijialara).
Carabus ghilianii, Laf., C. guadarramus, Laf., and
C. hellwo, Dej., under stones, especially about the
sources of the small streams, at the upper limits of the
pine forest, the first-mentioned species the most common.
Leistus constrictus, Schauf., a few specimens, at the sources
of the small streams, and also about the lowest patches of
snow ; it is exceedingly active, and difficult to secure in
43. Mr. G. C. Champion’s and Dr. T. A. Chapman’s
perfect condition ; on one occasion about a dozen examples
were found under the bark of a pine stump, on the banks
of a small stream, but few of them, however, were captured
without injury. Nebria vuillefroyi, Chaud., not rare, but
mostly immature at this season, under stones and moss, in
the streams issuing from near the melting snow. Zachypus
cyanicornis, Pand. Bembidium pyrenzum, De}. (var. carpeta-
num, Sharp), abundant, at the edges of the snow patches
on the Pefialara; B. decorum, Duft.; B. ibericum, Pioch. ;
B. laterale, Dej.; B. hypocrita, Dej.; B. toletanum, Perr.
Tachys parvulus, De}. Trechus pandellei, Putz. ?, plentiful,
at the edges of the snow on the Pefialara. Platyderus
montanellus, Graells. Haptoderus nemoralis, Graells, plenti-
fully, with the Leistus. Lemosthenecs pinicola, Graells.
Steropus ghilianiw, Putz., not rare, under stones. Zabrus
seidlitz, 58chaum,common on the mountains. Chlenius dives,
Dej., on the mountain slopes, running about in grassy places.
Deronectes griseostriatus, De G., Hydroporus flavipes,
Oliv., etc., Agabus heydeni, Wehncke, A. guttatus, Payk.,
A. conspersus, Marsh., Helophorus glacialis, Villa, Hydreia
sp., Hsolus angustatus, Miill., Lareynia carinata, Perez,
Limnius sp., ete., i the Laguna de los Pajaros, on the
Pefalara, or in the streams lower down.
Echidnoglossa glabrata, Kies. one specimen, on the
mountains. <Atheta luctwosa, Rey, and others of the genus,
at the edges of the snow, on the Pefalara. Baptolinus
alternans, Gravy., in the pine stumps. Geodromicus nigrita,
Miill., at ‘the edge of a waterfall near La Granja.
Liodes humeralis, Kug., L. castanea, Herbst, and ZL.
glabra, Kug., Agathidiwm piceum, Er., Enicmus rugosus,
Herbst, Bothrideres interstitialis, Heyd., Plegaderus sauctus,
Er., Paromalus flavicornis, Herbst, under bark of pine
stumps. Phalacrus maximus, Fairm. Olibrus bisignatus,
Mén., O. bimaculatus, Kiist., O. pygmeus, Sturm, on flowers,
near La Granja. Pedilophorus n. sp. (belonging to the sub-
genus Trichobyrrhulus, Ganglb.), two specimens, near the
snow, Pefialara.
Copris lunaris, L., Aphodius fetens, F., A. scrutator,
Herbst, A. bonvouloirt, Harold, A. obscurus, F., Ammecius
Srigidus, Bris., Onthophagus schreberi, L., Oniticellus flavipes,
F. , Gymnoplenrus flagellatus, F. , Geotrupes coruscans, Chevr.,
ete., in dung on the mountains, the Geotrupes frequently
taken on the wing in the hot sun. JLhizotrogus pyqialis,
Muls. Serica mutata, Gyll.
Entomological Excursion to Spain. 49
Corebus xneicollis, Vill. Anthaxia millefolii, F. Melan-
otus dichrous, Er.
Lygistopterus sanguineus, L.,on the pine stumps. 7%77-
chodes amnvios, F., 7’. apiarius, L. Sphindus dubius, Gyll.,
in powdery fungus on pine stumps.
Micrositus ulyssiponensis, Germ., under stones. JLisolam-
pus scabricollis, Graells, under stones, Puertode Navacerrada.
Lagria rulida, Graells (parvula, Perr.). Mordellistena
neuwaldeggiana, Panz. Zonabris hieracw, Graells, Z. 4-
punctata, L., Z. variabilis, Pall., Z. dejeani, Gyll., Z. geminata,
F., etc., and Cerocoma schreberi, F., more or less abundant
on flowers. Cdemera podagrariz, L.
Otiorrhynchus truncatellus, Graells, not uncommon on
the Guadarrama, occurring even on the summit of the
Pefialara. Strophosomus erinaceus, Chevr.; S. ebenista,
Seidl., a common insect on the Guadarrama, and often
found in numbers under stones. Cathormiocerus chevrolati,
Seidl, and Dichotrachelus graellsi, Perris, near the snow,
Laguna de los Pajaros. Cneorrhinus pyriformis, Boh., not
rare, on oak bushes near the town: two forms of the female
were met with (as in the species found on Genisia at
Piedrahita in 1902), one green, the other grey, like the
male, the latter being less elongate than the corresponding
sex of the Piedrahita insect ; the scales of the green females
from La Granja are opaque, while in those from Piedrahita
they are metallic.* Zhylacites sp., common, under stones,
in bare hot places. Brachyderes lusitanicus, F., B. incanus,
L. Pachytychius scabricollis, Rosenh. Polydrosus setifrons,
Duv. Brachytemnus porcatus, Germ., in the pine stumps.
Ceuthorrhynchus biscutellatus, Chevr. Balaninus pellitus,
Boh. Orchestes avellanx, Don., O. tlicis, F., O. quercus, L.
Rhamphus flavicornis, Herbst. Apion atomariwm, Kirby,
A. cracce; L., A. pomonw, ¥., A. vorax, Herbst, etc.
Attelabus curculionoides, L. Hylastes palliatus, Gyll.
Spondylis buprestoides, L., a common species in the pine
forest. Vesperus brevicollis, Graells, two males beaten from
oak. Criocephalus sp. in the pine forest. Leptura
stragulata, Germ., LZ. rubra, L., L. livida, F. Dorcadion
graellsi, Graells, D. hispanicum, Muls., Laguna de los
Pajaros, Pefialara, not rare, but worn, both species being
almost over in July.
Titubea biguttata, Oliv. Lachnea pubescens, Duf., L.
sexpunctata, Scop. Cryptocephalus 5-punetatus, Harr.,
* Of. Trans. Ent. Soc. Lond. 1903, pp. 176, 177.
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 4
50 Mr. G. C. Chanipion’s and Dr. T. A. Chapman's
sparingly, on young oak bushes on the “flat” outside La
Granja: we have not seen a record of this conspicuous
species from Spain, and it is not represented in the Madrid
Museum ; C. sewmuculatus, Oliv., C. globicollis, Suffr., C.
morei, L. Cyrtonus montanus, Graells, Puerto de Nava-
cerrada, nearly over in July. Z%imarcha sp., in abundance,
Laguna de los Pajaros. Chrysomela hemoptera, L.; C. ru-
foxnea, Suffr. Hispa atra, L. Cassida margaritacea, Schall.
Halyzia 12-guttata, L, H. 18-guttata, L. Adonia
mutabilis, Scriba. Coccinella 14-pustulata, L. Exochomus
4-pustulatus, L., LE. awritus, Scriba.
List of HemirrEra-HETreROPTERA found at La Granja.
Hurygaster maura, L. Graphosoma lineatum, L. Sehirus
dubius, Scop., S. maculipes, M. and R. Alia rostrata,
Poh. Neottiglossa leporina, H.-S. Staria lunata, Hahn.
Hurydema festivum, L., E. oleraceum, L. Tropicoris rufipes,
L. Carpocoris purpuripennis, De G. Verlusia quadrata, F.
Centrocoris spiniger, F. Gonocerusjunipert, H.-S. Therapha
hyoscyamt, L. Corizus erassicornis, L., C. parunypunctatus,
Schill. C. tégrinus, Schill. Lygevus equestris, L., L. pandurus,
Scop., LZ. superbus, Poll. Nysius thymi, Wolff. Cymaus glandi-
color, Hahn, C. melanocephalus, Fieb. Heterogaster arte-
misiv, Schill. Macroplax fasciata, H.-S. Microplax inter-
rupta, Fieb. Aphanus pini, L. Taphropeltus championi,
Reut.,n. sp. Beosus maritimus, Scop. LHinblethis angustus,
Mont. Jonanthia echii, Wolff. Serenthia Jleta, Fall.
Phymata crassipes, F. Aradus flavicornis, Dalm., one
specimen, in the Laguna de los Pajaros. Coranus sub-
apterus, De G. Nabis ferus,L. Salda cocksi, Curt. Cryp-
tostemma alienum, H.-S. Piezostethus terricola, Reut., under
a stone on the mountains. Jliris levigatus, L., M. cal-
caratus, Fall. Megalocerea erratica, L. Lopus gothicus, L.
Phytocoris meridionalis, H.-8., abundant, and P. vittiger,
Reut., P. varipes, Boh., P. femoralis, Fieb., P. abeillet, Puton,
and P. albofasciatus, Fieb., more rarely, on oak, etc., but
very difficult to secure. Iegalocelum infusum, H.-S. Adel-
phocoris lineolatus, Goeze. Peciloscytus unifasciatus, F¥.
Brachycoleus triangularis, Goeze. Cyphodena instabile, Luc.
Camptobrochis lutescens, Schill. Capsus ruber, L. Pilo-
phorus cinnamopterus, Kb., and P. perplecus, Scott, on oak.
Oncotylus boliwari, Reut. Sthenarus ocularis, M. and R,
Dicyphus genieulatus, Fieb. Coriaa mesta, Fieb,
Entomological Excursion to Spain. 51
From La Granja we went on to Madrid, spending a day
or two there to visit the Museum, etc. Leaving Madrid on
Aug. 5th, we proceeded northward to Navalperal (4165 ft.),
in the Province of Avila, and at the western extremity of
the Guadarrama. Here we were joined by the Spanish
entomologist, Manuel Escalera, who accompanied us to
the “Pinar” of Las Navas, ete. The partially-dried-up
streams at this place furnished a large number of water-
beetles, Deronectes carinatus being particularly abundant,
and on the sandy banks we found Acylophorus glabricollis,
various Lembidia, etc. In the pine forest we obtained
divers Longicornia and other beetles, from the earthen-
ware cups placed on the pine trunks to catch the exuding
resin, such as Nothorrhina muricata, Ergates faber, Tem-
nochila caerulea, etc. Our time, however, at Navalperal
was limited, and we finished our collecting here by
catching several males of Vesperus brevicollis, just before
commencing our long homeward journey. It may be
worth noting, perhaps, that at Irun, on July 7th, we
observed Hylotrupes bajulus in numbers on the telegraph
posts, in which the females were depositing their eggs.
List of CoLEoprers found at Navalperal.
Bembidium elongatum, Dej., B. octomaculatum, Goeze.
Bidessus geminus, ¥., B. minutissimus, Germ. Hydroporus
halensis, ¥., var. fuscitarsis, Aubé, in profusion, H. flavipes,
Oliv., H. varius, Aubé, H. lepidus, Oliv., H. marginatus,
Duft., AH. litwratus, Brullé, H. discretus, Fairm., ete.
Deronectes carinatus, Aubé, in abundance. <Agabus brun-
neus, F., A. didymus, Oliv., A. chalconotus, Panz. Ilybius
fuliginosus, F. Dytiscus pisanus, Lap. (2), D. marginalis,
L. (f and $), D. punctulatus, F. Gyrinus wrinator, Ul.
Hydrophilus flavipes, Stev. Limnoxenus oblongus, Herbst.
Hydrobius fuscipes, L. Helochares'sp. Paracymus xneus,
Germ. Hydrochus sp. Hydrena testacea, Curt., H. riparia,
Kug., H. nigrita,Germ. Pelochares versicolor, Walk. Acy-
lophorus glabricollis, Lac. Philonthus swavis, Bris., P. wm-
bratilis, Grav., P. quisquiliarius, Gyll. Phalacrus maximus,
Fairm., not rare on flowers. Meligethes exilis, Sturm, with
the preceding. TZemnochila cerulea, Oliv., Anthaxia 4-
punctata, L., Melanotus dichrous, Er., Rhizotrogus neglectus,
Perez, Nothorrhina muricata, Dalm., Ergates Jaber, L., and
Criocephalus sp., Hylurgus ligniperda, L., Tomicus seaden-
ay, Prof. O. M. Reuter’s Descriptions of two
tatus, Boern., in or about the earthenware cups fastened on
the pines to catch the exuding resin; many of the speci-
mens had been thus trapped, and were so coated with resin
that they had to be immersed in benzine or turpentine
before they were of any use. Lygistopterus sanguineus, L.
Clerus formicarius, L. Cerocoma schreberi, F. Zonabris
geminata, F., Z. variabilis, Pall. Brachyderes suturalis,
Graells, on pines. Cleonus marmottani, Bris. Lixus iridis,
Oliv., in the stems of a large Umbellifer; Z. ascani, L.
Pissodes notatus, F. Rhynchites cwruleocephalus, Schall., on
Cistus. Vesperus brevicollis, Graells, males attracted to an
acetylene lamp placed by Sefior Escalera outside a house
in the village for the purpose of catching moths.
List of HemMiprerA-HETERoPTERA found at Navalperal.
Ancyrosoma albolineatum, F. Ailia acuminata, L. Hury-
dema festiwum, L. Phyllomorpha laciniata, Vill. Centro-
coris spiniger, F. Syromastes marginatus, L. Camptopus
lateralis, Ger. Neides tipulariws, L. Lygeus saxatilrs,
Scop. Nabis ferus, L., WN. rugosus, L. Gerris gibbifer,
Schml. Coriza sahlbergi, Fieb., C. transversa, Fieb., C.
semistriata, Fieb.
Descriptions of two new species of Hemiptera-Heteroptera
Jrom Spain. By Prof. O, M. REUTER.
Laphropelius championi, n. sp.
Fusco-ferrugineus, 3 apicalibus marginum lateralium pronoti an-
guste pallidis, 4 horum basali nigro-fusco ; clava tertiaque basali
parte, corii testaceis, illo apice sat late nigro-fusco, corio cetero nigro-
fusco, macula marginali fere in tertia apicali parte posita albido-
flavente, membrana infuscata, venis obscurioribus, macula oblonga
exterlore suturam membrane terminante ; antennis modice gracili-
bus, obscure ferrugineis, articulo secundo apice tertioque nigro-fuscis,
quarto flavo-testaceo, basi nigro-fusco; pedibus cum coxis flavo-
testaceis vel flavo-ferrugineis, femoribus anticis (?) modice incras-
satis, spinulis duobus instructis et inter eas et apicem subtiliter
denticulatis, spinula posteriore supra medium posita, tibiis anticis
sat fortiter arcuatis.
Long. 2 23 mm.
Hab. SIERRA DE GUADARRAMA, La Granja.
new species of Iemiptera-Heteroptera from Spain. 53
L. contracto, H.-Sch., minor, colore corporis, antennarum et
pedum, pronoto magis quali, lobo antico ejus parum convexo,
sulco transversali minus distincto, spinula posteriore femorum
anticorum basi magis appropinquata mox distincta. Caput omnium
ereberrime et subtilissime punctatum. Antennz articulo secundo
latitudini verticis oculique unici que longo, ultimis apice ejus
paullo crassioribus, tertio seeundo circiter + breviore, quarto secundo
fere zque longo. Rostrum pallide flavens, coxas intermedias attin-
gens. Pronotum sulco transversali pone medium sito, parte apicali
parum convexo, creberrime subtiliter punctata, parte basali concolore,
minus crebre et paullo fortius punctata. Scutellum ut pars basalis
pronoti punctatum. Clavus triseriatim punctatum, Corium apicem
versus obsoletius punctatum.
Two specimens.
Pachytomella cursitans, n. sp.
Nigra, nitens, pronoto postice hemielytrisque omnium subtilissime
cinereo-pubescentibus ; antennis pedibusque utriusque sexus totis
nigris ; vertice haud vel obsoletissime marginato et utrinque ad
oculum obtuse depresso, fronte fortiter convexa, clypeo basi a fronte
impressione bene discreto; antennis articulo secundo latitudine
verticis interoculari parum longiore (¢) vel hac circiter 4 breviore,
gracili (9); hemielytris crebre sat fortiter coriaceo-punctulatis ;
mare feminaque (plerumque) brachypteris ; capite forma brachy-
pterze basi pronoti #que lato; dorso abdominis femine leviter
zenescente,
Long. ¢ ler, 2 14mm.
Hab. CANTABRIAN Mountatns, Puerto de Pajares.
P. doriz, Ferr. et Reut., affinis et similis, corpore superne niger-
rimo haud chalybeo-nitente, quamvis valde nitido, glabro, solum
pronoto postice hemielytrisque omnium brevissime et subtilissime
einereo-pubescentibus, his fortius coriaceo-punctatis, antennis
brevioribus pedibusque utriusque sexus totis nigris divergens.
Corpus forme brachypterze breviter ovale (¢) vel late ovatum (9 ).
Caput forme brachypteras a supero visum pronoto longitudine
equale vel subequale, ab antico visum latitudini verticis oculique
unici fere eeque longum, vertice oculo fere 24 (¢)—-triplo (?)
latiore, clypeo retrorsum vergente, genis altis, gula haud distin-
guenda. Oculi retrorsum vergentes, angulis anticis pronoti incum-
bentes. Antenne ad (4) vel paullo infra(?) apicem oculorum
interne inserte, articulo primo apicem clypei haud attingente,
secundo versus apicem sensim incrassato, femine quam maris
54 Two New Species of Hemiptera-Heteroptera.
graciliore, duobus ultimis simul sumtis secundo paullo longioribus,
quarto tertio breviore. Pronotum forme brachyptere basi longitu-
dine duplo (¢) vel magis quam duplo (?) latius, apice quam _ basi
parum angustius, disco horizontali, antice foveolis quatuor in arcum
positis, postice subtiliter striguloso, margine basali late sinuato.
Hemielytra forme brachyptere apicem segmenti quarti (3) vel
basin segmenti tertii (2) dorsalis attingentia, tota coriacea, com-
missura scutello distincte longiora (¢)-vel huie eque longa (2),
margine apicali valde obliquata ( ¢) vel subrecte truncata (9), angulo
exteriore subrecto (¢ ) vel acutiusculo, rotundato (¢).
Many specimens.
EXPLANATION OF PLATE.
PuaTEe VIL.
The Monastery of Santas Arbas, about a mile south of the Pass
of Pajares, and near the confluence of several valleys on the southern
side,
ro
Or
Oc
w
V. The Egg and Karly Larval Stages of a Coreid Bug,
probably Dalader acuticosta, Amyot et Serv.; with a
note on its Hymenopterous Parasite. By NELSON
ANNANDALE, B.A., Deputy Superintendent of the
Indian Museum, Calcutta.
[Read November 16th, 1904.]
PLATE VIII.
So little is known regarding the life history of the
Heteroptera that I have ventured to bring forward certain
observations on this subject which are confessedly most
incomplete. Possibly my figures and descriptions may be
of value to some naturalist more fortunate in rearing the
insect to be considered than I have been.
I.
On August 14th last, at Alipur in the suburbs of
Calcutta, I found, attached by a spider’s web to a tree-trunk,
a dead leaf on which were 14 eggs. They were arranged
separately, roughly in parallel rows but in no very definite
manner. Their length was about 2°75 mm.; their height
19 mm.; their colour a rich golden-brown, shining and
lustrous. As the embryo developed they became darker.
Examination with a lens showed that the surface was
patterned with a minute reticulation, and that a cordiform
or sub-oval area was marked off at the broader end. This
proved to be an operculum. Another area, of greater
extent than the last, but having identical limits above was
indicated. It was evidently due to internal structures
partially revealed by the translucency of the egg-shell.
In this region there were several extremely minute punc-
tures, which were only seen under a fairly high power of
‘the microscope. They were arranged in an are round the
upper limits of the operculum and were undoubtedly
micropyles: they are too small to appear in the figure.
The eggs were fastened to the leaf by a drop of a gummy
substance less brittle and more elastic than that of the
egg-shell. It was situated near the centre of the flattened
base.
TRANS. ENT, SOC, LOND, 1905.—PART I, (MAY)
56 Mr. N, Annandale on the
During the night between August 18th and 19th, each
egg produced a young Heteropteron. The operculum was
pushed out bodily and did not remain attached at any
point to the remainder of the egg-shell. What I take to
be the embryonic exuvize were lett hanging from the upper
part of the aperture thus formed in a heart-shaped mass,
the free apex of which was produced into a delicate
filament. ‘The mass was also attached to the base of the
ego-shell by a stouter filament, which originated above
as a ridge on the inner surface of the mass. On the
external face was a very conspicuous black spot, the nature
of which I have been unable to determine.
In what manner the operculum was pushed out I cannot
say, as I did not see the hatching. It appeared to be very
firmly attached to the remainder of the shell although its
limits were clearly defined to the eye. There was nothing
of the nature of a hardened projection on the head of the
larva when hatched.
The young bugs in their first instar measured 4°5 mm.
in length. There was no visible difference in size between
the different individuals of the brood. The head was
squarish, but shghtly rounded behind ; the eyes were fairly
large, but not prominent; I could detect no ocelli. At
first sight the antennz appeared to be five-jointed, but a
closer examination showed that the first apparent joint
was really a projection from the head: they were inserted
into the upper part of the head. The rostrum originated
close to the anterior margin of the head and was freely
movable; only three joints could be detected in it with
certainty, The tarsi appeared to have only two joints.
The form and proportions of the different parts of the body
are well shown in fig, 4, The colour was leaf-green,
marked with purplish-brown, which changed to olive-zreen
on the extremities of the limbs and antennz. The dorsal
“stink-glands”’ were rendered conspicuous by their promi-
nence and dark colour.
The larvee were sluggish, but they fed readily on the
juices of various leaves, standing with their rostra vertically
inserted into the vascular parts of the leaves, generally
into the midrib, Their abdomens rapidly became almost
globular.
During the night between August 21st and 22nd, the
first ecdysis took place. “A remarkable change both in
structure and demeanour was at once apparent. * The larvee
Log and Early Larval Stages of a Cored Bug. 57
were now about 7 mm. in length, but some individuals
were distinctly larger than others. The abdomen had
become narrower in proportion to its length; the separa-
tion between it and the thorax was now less marked; a
raised Y-shaped area had appeared on the dorsal surface of
the head ; a third joint to the tarsus was feebly indicated
(less feebly on the third pair of legs than on the first and
second), and the darker markings on the body had become
more extensive. The most notable change, however, was
intheantennsx. The first and second joints had lengthened
very considerably ; the penultimate joint had become
broad and flat, with a longitudinal ridge on its dorsal
surface ; the distal, increasing little in actual dimensions,
was now pear-shaped. The expansion of the penultimate
joint was rendered more conspicuous by the pale colour of
the distal adding a purplish suffusion on its own distal
half.
In the first instar, then, the antennz were of a general-
ized type ; in the second they had become highly specialized.
After the ecdysis the demeanour of the larvee became
much less sluggish. They ran about continually, agitating
their antennz, which were not held in the same attitude
as in the first instar (compare figs. 4and5). They refused,
however, to feed on the juices of leaves, and consequently
died, in the course of a few days, of starvation.
I have already noted that the individuals differed from
one another in size. With the exception of one, which
died immediately after getting rid of its old skin, the larger
individuals succumbed after the smaller. The difference
I attribute not directly to innate physical characters but
to the fact that on hatching all the young larve did not
show the same aptitude in finding the leaves provided for
their food. Some wandered about for several hours with-
out settling down to suck, while others commenced to do
so at once. All, however, apparently underwent ecdysis
at the same time. I may mention that when the skin was
cast the expanded joint of the antennz was drawn out
through the narrow aperture at the base of the organ, just
as the claws of crabs are drawn out or the expansions on
the femora of some Mantodea (e.g. Hymenopus bicornis).
As regards the systematic position of these Heteroptera,
I have no hesitation in assigning them to be Coreid, and
very little to the genus Dalader, which is remarkable for
having antenne with an expanded, ridged penultimate
58 Mr. N. Annandale on the Stages of a Corcid Bug.
joint and a pale-coloured, pear-shaped distal one. If this
attribution is correct the species is almost certainly Dalader
acuticosta, Amyot et Serv.,as this is the only representative
of the genus known from Lower Bengal, where it is not
uncommon,
II.
On August 20th I found seven more eggs of the same
species attached directly to the bark of the same tree at
Alipur. A few days later I dissected six of them. Each
was occupied by a small, black Hymenopteron, already
fully formed but enveloped in a delicate membrane. It
lay on its belly im the egg, with its head bent down beneath
the level of its thorax. The anterior edge of the thorax
was directed towards the operculum but was not in contact
with it. A sub-triangular, whitish mass covered the dorsal
surface of the abdomen, but was not in organic connection
with it. Possibly this was all that remained of the proper
occupant of the egg-shell which the Hymenopteron had
devoured. A similar parasite was hatched from the seventh
egg but unfortunately escaped.
In the collection of the Indian Museum I have recently
come across a third clutch of eggs, attached to the leaf of a
sugar cane. With them are the Hymenoptera bred from
them, evidently belonging to the same species. By a very
natural error the egg-shells are entered in the Museum
register as being the cocoons of the parasite.
I do not know of any other case in which a Hymeno-
pteron has been recorded as parasitic on the eggs of a
Heteropteron, though other members of the same family
are known to infest the egg-cases of cockroaches. The
eggs of the species under consideration are probably very
minute indeed and may be introduced through one of the
micropyles. No other aperture could be detected in
parasitized eggs. It is evident that the shells are firm
when the infested eggs are laid, as their base does not
preserve a cast of the surface to which they adhere.
The specimens of the parasite examined are in a poor
state of preservation. They belong to the family Chalcidide
in its wider application : further it would be ridiculous for
one who is not a specialist in the group to go.
Haplanation of Plate. 59
EXPLANATION OF PLATE VIII.
Fre. 1. Parasitized Egg of Heteropteron (? Dalader acuticosta),
from above.
” 9 ”
from one side.
3. Egg from which Heteropteron has hatched.
4, Larva in Ist instar.
” ” 2nd ”
” ”
Fies. 1 to 3 x 8; fig. 4 x 4; fig. 5 x 3.
In fig, 5 the artist has tipped the head of the insect up slightly so
that the proximal extremity of the mouth parts appears.
= ? ; i fa Aya, ; Lue ie th rae pa
% ¥ yOns Sit t clang es 1’ (> enh
1 gist Aka Oak a
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( 61")
VI. Notes on the Butterflies observed in a tour through India
and Ceylon, 1903-4. By G. B. Lonasrarr, M.D.,
Oxon.
[Read December 7th, 1904. ]
INTRODUCTORY.
Wuat follows is an account of the entomological ex-
periences of a “jglobe-trotter,” that is, of a traveller whose
main object was to take an all too rapid glance at the
scenery, the peoples, and the architecture of the places
visited, and whose route was planned with that object.
That I was able to give so much time to collecting was
due to the fact that, whereas my daughter and -her com-
panion felt the heat so much that they usually kept within
doors from about 10 a.m. to 8 p.m., I, for my part, protected
by a “sola tope” of the “ pigsticker” type, and a spinal
pad to my coat, suffered no serious inconvenience from the
sun’s rays so long as I took active exercise.
I sailed from England in September 1903 without the
slightest intention of collecting, and started accordingly
with no entomological outfit save half-a-dozen pill-boxes.
Not only was I without net and killing-bottle, I was with-
out books, and worse still, was in woeful ignorance of the
Rhopalocera of the Oriental Region.
The day after landing we took train for Simla, and a
little south of Jhansi I was struck by the large numbers of
bright yellow butterflies along the railway banks—in all
probability Zerias hecabe, L. October 5th found us at Kalka,
at the foot of “The Hills.” Fortunately the new railway
was not yet open, so we had to be driven up the 58 miles
to Simla in a “tonga,” or post-cart, by a wild-looking
hillman who handled the ponies magnificently. ‘To one
fresh from Europe the sights on the road were truly mar-
vellous: long trains of wagons drawn by humped oxen
or by buffaloes; natives in divers strange costumes, or
lack of costume; flocks of goats and herds of cattle ; strings
of pack-mules, and, to crown all, long lines of solemn
camels, always hideous, yet always picturesque. However,
amid all these strange sights there was one other which
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY)
62 Dr. G. B. Longstaff’s Notes on the Butterflies
interested me if possible even more, I ean the multitude,
the variety, and above all the beauty of the butterflies.
The first sight of such a thing as the big Hypolimnas
bolina, L., black flashing with violet-blue, excited an emotion
better imagined than described. At all events, the
creatures took me fairly by storm: collect I must!
The resources of the bazar at Simla ouly produced a
child’s butterfly-net, a mere toy, scarce twelve inches in
diameter and of a pale yellow colour! Armed with this
and a tin cigarette-box filled with triangular envelopes I
took the field. To this scanty equipment was shortly
added a cyanide bottle. It was two months before the
toy-net was superseded by an umbrella-net from Watkins
and Doncaster. This last is a very convenient weapon for
use in towns, or when travelling or sight-seeing. It is
inconspicuous when rolled up, but can be quickly brought
into action ; it is however inadequate for serious work. In
Calcutta I purchased a large Y-net with jointed canes,
and had it fitted to the end of a landing-net stick that was
made in two pieces that were six feet long when joined.
A fair-sized net is required for large and swift butterflies,
while for the many that habitually fly high and settle far
from the ground, six feet is none too long a stick, though
in narrow woodland paths it will be found unwieldy.
Another time I should travel with a jointed stick of three
segments, each three feet long. J may here add that
mosquito netting is far more serviceable than leno, owing
to its superior power of resistance to thorns of all sorts.
Mr. Otto Moller, of Darjiling, told me that he found
it best to pinch all butterflies, even the smallest, but I
found that Blues and Skippers were best “ bottled.” No
doubt it would have been better to pin many moths, but
my things were all enveloped in the way usual for butter-
flies, and the dates, localities, and any remarks that sug-
gested themselves were inscribed upon the papers. [These
data are still attached to the specimens.] Since getting
back to England the insects have been serially numbered,
and the data copied into a note-book. On another
journey I should number the insects at the time, and
while putting the more important data on the papers as
before, copy these, amplified where necessary, into a book.
This would, I am quite sure, save much time and result
in a more complete record.
The insects were sent to England from time to time
Observed in a towr through India and Ceylon. 68
by parcel post in small cigar-boxes, each enclosing a ball
of naphthalene. In no case do they appear to have sus-
tained any injury on the way. They have been beautifully
set at Oxford by Mr. A. H. Hamm, and all that are worth
preserving will be placed in the Hope Collection, while
the explanatory note-book will be deposited in the library
of the Department.
The total number of specimens sent home was as
follows :—
All countries. India and Ceylon only.
Butterflies . ; 1867 . : ; 1494
Moths : : 206 . : : 125
Coleoptera . : 7 ee ‘ : 15
Hymenoptera. BT : : 27
Neuroptera ; i : ; 5
Diptera. 5 bar : : 5
Hemiptera . . LO" d : 8
Orthoptera . : is: : 3
2164 1692
In round numbers, I was five months in India and three
weeks in Ceylon—say, six months together; during this
time I took nearly 1700 specimens, of which 1500 were
butterflies belonging to 204 species.
To these may be added the results of a fortnight in
China, a month in Japan, and a fortnight in Canada, viz.
500 more specimens, and 64 additional species of butter-
flies, which are only incidentally alluded to at the end of
this paper.
Naturally in a rapid tour of this kind there is small
probability of turning up anything new, but it is hoped
that some of the observations made (even on the com-
monest species) may throw a glimmer of light on some
of those questions of Bionomics which are now attracting
attention.
Simla, lat. 31° N., alt. 7200 ft.
In reference to the seasonal variation of many species
it may be remarked that at Bombay on October 2ad and
3rd there was heavy rain, the tail-end of the monsoon.
It was held to be a very late season, the rain had lingered
and the cold weather was delayed.
My collecting at Simla was confined to a riding ‘expedi-
tion along the old Hindustan-Tibet road. This is an
64 Dr. G. B. Longstaff’s Motes on the Butterflies
excellent riding-path along the watershed of the Sutlej
and Jumna, cut at one time on the hot and dusty southern
side of the mighty ridge, where the terraced slopes are
covered with crops of maize, or ruddy millet ; anon crossing
to the northern side which is mostly clothed with fine
forest of spruce, deodar, holm-oak and rhododendron—thus
winding in and out, but for the 50 miles that we traversed
always maintaining an altitude of from 7000 to 9000 ft.
We went by way of Fagu, Theog, Matiana and Narkanda to
Baghi, returning by the same route, except that from
Baghi to Narkanda we walked over Mt. Huttu, 11,000 ft.*
The expedition occupied eight days, but for simplicity
of description I shall not distinguish between outward
and return journeys.
The general aspect was decidedly autumnal; the nights
were chilly and most flowers had gone to seed. There
was more cloud than usual, and there were occasional
slight thunder-showers. Most of the butterflies seen
appeared to have been out some time, and were much
battered. Two circumstances tended to restrict the bag—
one, the fact that collecting was for the most part confined
to a narrow mountain road, bounded by a precipice on the
lower, and a cliff upon the higher side ; the other, a limit-
ation of wide application, that a tropical sun is not
conducive to rapid pursuit.
Simla, alt. 7200 ft., to Fagu, alt. 8200 ft.
October 10th and 17th.
Gonepteryx rhamnt, L., var. nipalensis, Dbl., was abundant
throughout the journey, and so was Awlocera swaha, Koll.,
though in very poor condition ; both occurred in Simla
itself. The last named is a Satyrid having on the upper-
side a resemblance to our White Admiral, flying also with
much of the grace of that favourite butterfly. It loves
open spaces in woods, returning to the same spot when
disturbed. When it settles on the ground, a rock, a
flower, or a tree trunk, it often goes over on one
side as much as 45° or even 50°. I saw one of these
butterflies make three successive efforts, getting further
over each time! On two distinct occasions I watched a
butterfly settle twice, turning the first time over to the
* In Indian names “4” is pronounced as “a” in father, “wu” as
“oo” m boot, “a”? or “uu” as“u7 insbut
Observed in a tour through India and Ceylon. 65
right, the second time to the left. I think there is no
doubt that this “list” makes the insect less conspicuous.
Pararge schakra, Koll. is another common roadside
butterfly in the Simla district; it closely resembles our
P. megera, L., but islarger. Chrysophanus phleas, L., var.
timeus, Cr., was also common ; Colias fieldii, Mén., is sufti-
ciently like our C. edusa to readily pass for that species ;
Polyommatus beticus, L., is also suggestive of our South
Downs, where it has been seen ; the same may be said of
Argynnis lathonia, L. Again, Pyrameis indica, Herbst., is
very like our P. atalanta, L., though not so handsome and
scarcely as graceful in its movements. So far there was
plenty to bring to mind the fact that one was still within
the Palearctic Region. There were however a few insects
to suggest the close proximity of the great Oriental Region,
for if Atella phalanta, Diu., is very like a Fritillary (at least
on the upper-side), and Jlerda sena, Koll., closely resembles
a Hairstreak, on the other hand, the under-side of Belenois
mesentina, Cr., is decidedly more brilliant than our Whites,
while there is no denying that Precis orithyia, L., is quite
Oriental in its aspect. This insect had struck me with
admiration at Solon on the way up to Simla, and is called
by the school-boys of India’s summer capital, “The Ladies’
Fancy.” With the habits of a Vanessa or Pyrameis, there
is something about the shape of the wings, the prominent
ocelli, the brilliant blue of the hind-wings, and the leaf-like
colouring of the under-side which gives it a very “ tropical ”
appearance. However, one soon learned to look upon it as
one of the most familiar butterflies of Northern India. At
Fagu it was common, but like most butterflies which have a
proclivity for settling on the ground, not too easy to catch.
At Fagu another buttertly of European aspect was
common, Vanessa kashmirensis, Koll.; this is no credit to its
name, but looks like an wrtica, L., that had been born and
bred in the “ Black country.” But the Chalcosiine day-
flying moth, Agalope hyalina, Koll., elegant in shape and
quiet in colour, white, shaded with grey towards the tips,
ochreous at the base, was quite a stranger.
Fagu, 8200 ft., to Theog, 7400 ft.
October 11th and 16th.
Before our start in the morning I found abundance of
Chrysophanus pavana, Koll, in dry weedy corners of
TRANS, ENT, SOC. LOND. 1905.— PARTI. (MAY) 5
66 Dr. G. B. Longstaff’s Notes on the Butterflies
cultivated ground; this is sufficiently distinct from C.
phiwas, but has no especial oriental glamour.
From the ground by the roadside I picked up a large
newly-emerged Bombyx with the awe-strikimg name of
Trabala vishnu, Lefevre ; it was unfortunately a good deal
damaged in the killing, through having no oxalic acid
available. Two Blues, Cyaniris vardhana, Moore, and Zizera
maha, Koll., var. diluta, Feld., together with the Hairstreak,
Llerda sena, completed the bag for this stage.
Theog, 7400 ft., to Matidna, 7700 ft.
October 11th and 16th.
At Theog, our first halting-place, Gonepteryx rhamni, var.
nipalensis, was especially common, and here I took my
first Athyma opalina, Koll., a Vanessid resembling on both
upper and lower surfaces Limenitis sibylla, L., an insect to
which it is closely allied in structure and habits.
On the road, besides Jlerda sena, Chrysophanus pavana,
Precis orithyia, and Argynnis lathonia, var. issva, Moore,
several things turned up. Of Pyrameis cardwi, L., a fresh
brood appeared to haveemerged onthe 15th or 16th October,
and was common at the flowers of a straw-coloured thistle.
I saw a few more Athyma opalina, and secured one. In
their elegant floating flight one seems to see through the
white markings of the butterflies of this genus. The
Simla school-boys call them “Sailors,” but to me the name
“Ghosts” would seem more appropriate. They settle on
the leaves of trees or shrubs, rarely affecting flowers. Here
I got my first Precis lemonias, L., an insect with the habits
(and structure) of a Vanessid, but with much the appearance
of Pararge xgeria, L. A specimen of Terias libythea, F.,
taken on the return journey, bears the note “easy to
catch,” which is true, but at the time I do not think that
I distinguished it from the much commoner 7. hecabe,
which it closely resembles.
At Matidna I found Chrysophanus phlxas, var. timeus ;
Vanessa kashmirensis; Precis orithyia ; and two specimens
of Huphina nerissa, F., both males. This last is a somewhat
glorified P. napi, L.; one of them appeared to have a
slight scent which I could not describe, but certainly it
was not that of the male napv.
Perhaps the most abundant butterfly at Matiana, and
indeed throughout the woods of the district, was Cyaniris
Observed in a tour through India and Ceylon. 67
singalensis, Moore, very like our argiolus; it was in poor
condition, flying about the tops of tall shrubs, but not
seeming to affect either ivy or holly, although both were
there.
Pararge schakra was especially abundant at Matidna
and on the road thence to Narkanda, It differs from our P.
megzxra, L.,in being larger and having more striking ocelli,
though these are variable, one of my specimens, a female,
having the ocellus near the tip of the fore-wing far larger
than the rest. In its habits this insect sometimes reminds
one of P. megera, sometimes of Satyrus semele, L, It abounds
along roads and in bare places, alighting almost always onthe
earth or on rocks, with its wings expanded (as with megevra),
but when it settles down to rest the wings are raised, the
fore-wings drawn back within the hind-wings, all that
remains visible being the colour of dust. In no case did I
see it turn on one side as S. semele does, but three times I
observed it settle with its back to the sun, so as to reduce
its shadow to a mere line; unfortunately I made this
observation towards the end of my acquaintance with the
butterfly, so was unable to make sure whether this was a
mere chance or a definite habit. At any rate, I did not
observe any instances to the contrary. I suspected in
.P. schakra the existence of a very slight sweet scent, that
appeared to be unlike that of any other species.
At Matiana I beat out of alders a number of geometers,
three Philereme variegata, Warr., and one Cidaria niphonica,
Butl.; they had a jerky flight, which saved many of them
from capture. A Deltoid, Hypena tristalis, Leder., came to
light at night.
Matiana, 7700 ft., to Narkanda, 8800 ft.
October 12th and 15th.
Many of the same insects were met with as on the
previous stage, but the following may be noted: Terias
hecabe, my first specimen of the commonest species of a
very characteristic Indian genus; Ganoris canidia, Sparrm.,
a White like P. rape, L., but with bigger black spots; and
Lelenois mesentina, flying fast and going straight ahead in a
purposeful manner. Here I may remark that the swift
flight of the Whites generally has much impressed me;
it is evidently closely related to the fact that they are
68 Dr. G. B. Longstaff’s Notes on the Butterflies
quite the most conspicuous of all butterflies, especially at a
distance. Argynnis lathonia, var. issva, was again well to
the front; another Athyma opalina was securely “ papered,”
and a specimen of Precis lemonias was taken in which the
anal angle of both hind-wings had been bitten off nearly
symmetrically. Amongst many of the argiolus-like
Cyaniris singalensis, Moore, one C. vardhana, Moore,
was taken; also Chrysophanus pavana. Three geometers,
Philereme variegata, Warr., Docirava xquilineata, Walk.,
and the widely-distributed Polyphasia truncata, Hufn.
(immanata, Haw.), complete the list.
At Narkanda Argynnis lathonia, var. issea, was in
great numbers in the woods, so was the Chalcosiine moth
Agalope hyalina; this flies fast and always in the same
direction, in this case up-hill and against the wind, more
especially up certain gorges in the mountain side. It was
hard to catch, and on the wing looked much larger than it
is, but on settling vanished suddenly, burying itself in the
herbage.
Colias fieldii was common, but was not remarkable for
swift flight. Zerias hecabe was also to be seen.
Narkanda, 8800 ft., to Baghi, 8900 ft.
October 13th.
The road through the magnificent forest, whence gaps
in the towering spruces give inspiring glimpses of “The
Snows” lying far away across the deep valley of the Sutlej,
was at this season too dark and chilly to be the haunt
of butterflies. At Bagchi were Atella phalanta, Neptis
astola, Moore, worn specimens of Awlocera swaha, and
Agalope hyalina.
Baghi, 8900 ft., over Mt. Hutiw, 11,000 ft., to Narkanda,
8800 ft.
October 14th.
Baghi, our furthest point, is but 26 miles W.N. W. of
Simla, though by the winding mountain road it is fifty.
The steep footpath up Mt. Huttu, when it has attained an
elevation of a little more than 10,000 ft., emerges from the
forest on to a flowery clearing that bore evidence of former
cultivation. Here I saw Colias field, Atella phalanta,
Observed in a tour through India and Ceylon. 69
and Argynnis lathonia, and here I took two specimens of
Parnassius hardwickii, Gray, one worn, the other in beauti-
ful condition ; it is a lovely creature, but the under-side has
a curious resemblance to oiled paper. Delicate looking
though it be, it is strangely tenacious of life. The con-
currence of a“ Clouded Yellow,” a “‘ Queen of Spain,” and an
‘Apollo ” was very suggestive of the Alps. All too soon the
path plunged again into the now somewhat scrubby forest
to come out finally, at near 11,000 ft., on to the grassy,
flower-bedecked plateau in which the mountain culminates.
The Lha To, or rude altar of the degraded form of
Buddhism that is prevalent in “The Hills,” crowning the
highest peak, reminded one of the High Places of Baal.
The troops of butterflies seemed almost to rejoice in the
glorious panorama of “The Snows” spread far around.
The brilliant Argynnis lathonia was common, and the
dingy Vanessa kashmirensis quite abundant—probably
the more distant of yon white peaks to the left arises from
its name-place, Kashmir. Colias jieldii was also in large
numbers, a female exhibiting a symmetrical injury to the
hind-wings very suggestive of a peck by a bird. Precis
orithyia. was there too, but Aulocera swaha was conspicuous
by its absence. Of a humming-bird moth, much smaller
than ours, Rhopalopsyche nycteris, Koll., I netted three
specimens, one at the flowers of a delphinium, Herbula
cespitalis, Schiff., reminded me of home. The Blues were
represented by Cyaniris singalensis, Moore. A male Terias
hecabe was of the wet-season form. I noted that this
species is very easy to catch, and is brilliant on the wing ;
also that when settled on a shrub or flower it is usually
extremely conspicuous, but not so when it chooses as its
resting-place a certain low plant with oval leaves fading to
a yellow tint; then the rounded form of the wings greatly
aids its concealment. An old friend, Zwxoa corticea, Schiff,
was taken flying in the sunshine. I had several exciting
chases after a big yellow Swallow-tail, and eventually
secured one—my first Papilio! It proved to be our
machaon, L., var. asiatica, Mén. Here, as in Japan, it scorns
fens and dykes, glorying in mountain tops. b.
On the way down to Narkanda several Pyrameis indica
disputed the path with our party.
The great resemblance to European forms presented by
the bulk of the butterflies seen in this expedition cannot
fail to strike the reader,
70 Dr. G. B. Longstaff’s Notes on the Butterflies
Solon, circa 5000 ft., to Kdlka, 2184 ft.
October 20th, 1903.
Starting from Simla by starlight, soon after 5 a.m., we
got to Solon by breakfast-time, and I there caught at 9
am. my first butterflies, two alsus-like Blues, Zizera
karsandra, Moore, and Z. maha, Koll. Also two flies, a
Musca of the domestica, L., group, and an Anthomyid.
On the drive from Solon to Kalka, by making the most
of stoppages to change horses, and by occasionally jumping
out of the carriage, I managed to secure quite a lot of
things. Among the commonest was the beautiful Precis
e@none, L., and with it P. orithyia and P. lemonias. Of Atella
phalanta, Belenois mesentina, 2, and Ilerda sena, I took
single examples. T'e7ias leta, Boisd., was rather common.
There were also Catopsilia pyranthe, L., the gnoma-form,
Terias hecabe, and Huphina nerissa. About two miles above
Kalka, say at about 2700 ft., I got a single Precis iphita, Cr.
At about the same place the great catch of the morning was
made, for I took my first Hypolimnas bolina, three males
and a female, believing them at the time to be two species.
Why does not this glorious insect retain its far more
poetical and more appropriate name, Diadema jacintha ?
Surely a black butterfly 34 inches in expanse with four
large glancing-blue spots, one on either wing, deserves to
be called after a gem. Anyway,I shall never forget the
impression produced by my first sight of its truly oriental
splendour; it was like Kingsley’s “ At last!”
On my way down I also saw Pyrameis indica, and missed
two Papilios, probably P. machaon.
At Kalka J got an hour and a half’s collecting late in
the afternoon ; it was partly on waste ground about the
station, but mainly in a field bearing a crop of some kind
of pulse with thin pods 4-5 inches long.
A black and brown Cantharid beetle, Mylabris sidx, Fab.,
was flying about flowers in the sunshine in large numbers.
The genus Precis was represented by orithyia and a@none ;
the genus Terias by hecabe, leta, and quite a number of
libythea. The inevitable Atella phalanta, never very
common, and Belenois mesentina were to the front again.
Ganoris canidia was fairly common; I noted that a male had
a “snuffy scent.” Single specimens of Jxias marianne, Cr.,
and Huphina nerissa, both males, were taken. Of Cato-
Observed in a tour through India and Ceylon, 71
psilia pyranthe 1 took two females, one of which had
suffered a symmetrical injury to both hind-wings. Three or
four Hypolimnas bolina, both sexes, were disturbed in their
first sleep, and being drowsy fell an easy prey. The Blues
were represented by several species—Zizera maha, Koll.;
Z, otis, Fab,, var. indica, Murray ; Catochrysops enejus, Fab. ;
and Nacaduba noreia, Feld. Two Pyrales, Hymenia
recurvalis, F., and Bradina admizxtalis, Walk., and a worn
Acidalid were picked up, A Sphinx, Nephele hespera, Fab.,
was taken during the afternoon at the flowers of a Lryonia,
A little later on, an Arctiid moth, Artaxva lunata, Walk.,
came to the lamp of the railway carriage, to which a Sphinx,
probably another JV. hespera, also paid a momentary visit,
Peshdwar, lat. 34° N., alt, 1165 ft.
October 22nd—25th, 1903.
This city is finely situated in the extreme north-west
of the great plain of the Panjab, or Five Rivers; the
mountains of the Sufid Koh and the foot-hills of the
Hindi Kush bounding the view to the west and north
respectively.
In the hotel garden I took a few things; Terias hecabe
was common, two of them lacked “the dog’s head mark.”
Belenois mesentina was represented by a solitary male.
One of three males of Ganoris canidia yielded a decided
scent, hard to describe but certainly not that of G. napz.
That dingy Skipper Parnara mathias, Fab., was abundant
at the flowers of Duranta. I missed several specimens of
a yellow Papilio, probably erithonius, Cr., and I believe
one allied to podalirius, L. Of the Blues I took one
Polyommatus beticus, and three Zizera karsandra, Moore.
Two moths came to light, Oligochroa akbarella, Rag.,
and Harias tristrigosa, Butl.
Near the waterworks at Bara, amidst a wilderness of
stones, I netted a female Belenois mesentina, three Blues,
Tarucus theophrastus, Fab. (2 $,1 9), and my first Zeracolus,
a female etrida, Boisd. Dr. Dixey tells me that he had no
idea that this species ranged so far north. <A strange-
looking grasshopper, Zruvalis nasuta, L., seemed well
adapted to its stony desert surroundings.
From ‘Peshawar my most interesting expedition, from
every point of view, was to Ali Musjid in the Khaibar
Pass. This tiny white building, said to be the first
72 Dr. G. B. Longstaff’s Notes on the Butterflies
Musjid erected in India by the invading hordes of Muham-
madan conquerors, stands about 2400 ft. above sea level.
Close to flows a little stream full of fish and frogs, which
produces an oasis among the hot dry rocks, where large
beds of a species of mint attracted a number of butterflies,
which I pursued under the strict and curious supervision
of those good-natured barbarians, the Afridis of the
Khaibar Rifles, who twice a week safeguard the caravans
as far as Lundi Kotal.
Limnas chrysippus, L., was fairly common. I took two
males and two females, one of the latter with pale ground-
colour of an umbreous tint and much shading along the
costa. Pyramets cardui was the commonest butterfly, mostly
in fine condition. I took one Ganoris brassice#, L., of the
form nipalensis, Gray, a female, and saw several CG. canidia,
which had all possibly strayed from a patch of cultivated
eround hard by. The Clouded Yellows were represented
by several Colias hyale, form erate, Esp. The beautiful Precis
orithyia was quite abundant ; P. almana, L., also occurred,
but was not common. I saw several Zerias hecabe.. The
Satyrids were the most interesting of all. A specimen of
Yphthima balanica, March, was my first acquaintance in
that elegant and delicately-made genus. Satyrus parisatis,
Koll, a handsome insect suggestive of Vanessa antiopa,
L., was rather common, but unfortunately much worn.
Very conspicuous on the wing it did not appear to be
attracted by the mint, but usually settled on the ground,
and was then very difficult to see. I also secured two
specimens of a very distinct pale Satyrid, much the colour
of C. pamphilus, L., but much larger and with dentate hind-
wings, Hpinephele davendra, Moore; they were both 2. I
took two Polyommatus beticus, but saw no Skippers.
Three of that widely-distributed beauty, Devopeia pul-
chella, L., were seen flying in the sun, and with them a
brilliant little Burnet, Zygena kashmirensis, Koll.
Among the outsiders were a locust, Pacilocera picta; a
beetle, Clinterta confinis, Hope; two bees, Bombus simal-
limus, Smith, f and 2; anda wasp, Vespa awraria, Smith (9).
Malakand, lat. 34° 30’ N., alt. circa 3000 ft.
October 28th and 29th, 1903.
By the kind hospitality of the Political Officer, Capt.
R. W. E. Knollys, I was enabled to get two days’ collect-
Observed in a tour through India and Ceylon. 73
ing at this remote frontier post. Perched on a saddle,
where the old Buddhist road crosses the foot-hills, looking
forward over the Swat valley, back over the dusty plain
of the Panjab, this isolated fortress affords a picture of
rocky desolation. The Pass is closed every night by
chevaux de frise, and the garrison is always prepared for
attack. When I went collecting it was deemed prudent
that I should be accompanied by a gigantic chuprassi, a
Pathan of the tribe of the Jusufsai, or Sons of Joseph.
Moreover, when scrambling over the hillsides, in addition
to the usual Indian thorns in all their varieties, wire
entanglements have to be negotiated !
The rocky hills seemed too dry and burnt up to harbour
many butterflies, but on the parched slopes of the fortified
erag, nicknamed Gibraltar, the pretty little Melitxa trivia,
Schiff., was almost abundant; ona glaucous shrub at the
foot of the same hill were numbers of a glaucous green and
yellow locust, Pacilocera picta, which though conspicuous
enough on the wing was decidedly cryptic. Other Orni-
thoptera were Quiroguesia blanchardianus, Sauss., and
Truxalis nasuta, L. I also took three wasps, two Vespa
velutina, Lep. (var. “ des Indes,” Sauss.) 9, and a § Polistes
hebreus, F.
In addition to the above a long and hot walk only
yielded one Ganoris canidia, $; two Terias hecabe, a f of
the variety without the ‘“‘dog’s head,” and a large but
otherwise normal 2; two Blues, a Zizera karsandra,
Moore, and a Z. maha, Koll. var. diluta, Feld.; one
Precis orithyia ; a dingy Skipper, Gegenes nostrodamus, Fab.,
and a micro, 7inxgeria, sp. Some puddles of water at
the baggage-mules’ drinking-place proved very attractive,
ylelding Argynnis niphe, a 9, Tarucus theophrastus, F.,
a ¢, and the conspicuous Hipparchia parisatis.,
The next day (Oct. 29th) I lighted upon an oasis in the
desert in the shape of the staff-sergeant’s garden, where
irrigation had produced a brilliant mass of flowers, some
vegetables, and a small field of lucerne. Here butterflies
abounded : Zerias hecabe, without the “dog’s head mark,”
was in plenty among the lucerne as well as at the marigolds ;
the lucerne also yielded both Colias fieldit and C. erate,
the eastern form of hyale. Among the Danaids Limnas
chrysippus was common, and D. genutia, Cr., abundant at
the marigold flowers, at which also one Z%rumala limniace,
Cr., was taken. Athyma perius, Linn., was rather com-
74 Dr. G. B. Longstaff’s Notes on the Butterflies
mon, but preferred the wet mud left in the irrigation
channels to any flowers. Argynnis niphe was also com-
mon, but had more refined taste, and was usually taken
on the marigold beds; its female was observed during life
to resemble Z. genutia. Precis almana was common; P.
orithyia very abundant at the same flowers, together with
a few P. enone, one of them very fine and large. As
usual in India Atella phalanta and Belenois mesentina put
in an appearance, the first at marigold, the second (a 2)
among the lucerne. The Hairstreak J/erda sena occurred
alike at marigold and high up on the mountain-side. The
flowers of Gaillardia proved more attractive to the smaller
fry than the coarser marigolds ; the brown Skipper, Parnara
mathias, Fab., was in abundance, so were the dingy Blues,
Lizera karsandra, Moore, and Z. maha, Koll., but the
latter and its variety diluta, Feld., preferred mud to any
flowers.
Other small things were Polyommatus beticus, and the
Skipper Gegenes nostrodamus, Fab., which was common at
the flowers of Gaillardia and marigold. I saw this species
at Malakand only, and unfortunately secured but two
specimens. Two or three Melitwa trivia also turned up
at these favourite flowers. Not far from the garden I took
two more Hipparchia parisatis ; this does not appear to be
much attracted by flowers, but settles on the ground and
is then often very hard to see. I observed it lean over
from 20° to 30°, and even saw it walking about with a
“list” of 20°.
Lahore, lat. 31° 35’ N., alt. cirea 700 ft.
October 31lst—November 4th.
At the capital of the Panjab, a city of the plains, my
chief collecting ground was the extensive Lawrence
Garden, which though full of flowers is, in parts, so wild
that, not to mention a mongoose, I even came across a
jackal at midday. The class of butterflies found here
differed widely from those met with at Simla and further
north, the predominant forms being Oriental. Here I
first captured Papilio erithonius, Cr., the “ tailless swallow-
tail,’ which I had perhaps seen at Peshawar ; this butter-
fly has a wide range in India and might almost be termed
abundant, it especially frequents the flowers of Zinnia,
Observed in a tour through India and Ceylon, 75
Lantana, and Bougainvillea. When feeding it settles for
a few moments only, fluttering with its wings the while ;
then it is not hard to catch, but when rushing from place
to place it is far otherwise. In colouring it is very like
P. machaon, but far less handsome; the yellow ground-
colour is often quite pale and bright when the insect is
fresh, but it usually turns much darker and duller; I sus-
pect that cyanide hastens this process. At Lahore also I
first came across another very common Indian butterfly,
Papilio pammon, L.; its graceful form and flight and rich
velvety-black coat at first excited me so much that I
had great difficulty in catching it! Naturally enough I
followed Linnzeus iv taking the sexes for different species :
he called the male pammon, and the very different female,
Wallace’s second form, pelytes. At Lahore it especially
affected the flowers of Bougainvillea and a shrub with
blossoms like in colour and scent to, but much larger than,
those of white jasmine. Like P. erithonius it flew rapidly
from flower to flower and fluttered while feeding, The
female taken here was of Wallace’s second form (polvtes) ;
among the males was a dingy variety with scarcely any
orange on the under-side of the hind-wings.
Inmnas chrysippus was abundant, more especially at the
flowers of Asclepias (the food-plant); amongst them was a
dwarf female. Tirwmala limniace was scarcely common.
Of Catopsilia pomona, F., 1 only netted one female, but
believe I saw others; it visits flowers high up on trees.
C. pyranthe was abundant; it flies fast and high and is
hard to catch; it was fond of settling on the flowers of
duranta on the tops of high hedges, forming a pretty
contrast with the lilac-blue racemes.
Terias hecabe, both sexes, was fairly common; it flew
slowly and near the ground. The black and white
Teracolus puellaris, Butl., was also fairly common; perhaps
it owes its name to the child-like simplicity of itsdress. It
flies near the ground, but so jerkily as to be somewhat
hard to catch. It has the habit of flying into bushes, by
preference those well provided with thorns, and not coming
out again. Of 7. protractus, Butl., I could only get two
specimens; its salmon-pink colour with broad black margins
dusted with blue-grey make it one of the most beautiful
little butterflies that I came across; its dress is all in
exquisite taste, the under-side being a quiet greenish-
yellow that must greatly protect it when at rest.
76 = Dr. G. B. Longstaff’s Notes on the Butterflies
White butterflies were not much in evidence; I took a
somewhat worn female of Appias libythea, Fab., also two
Belenois mesentina, both females. This last is another
common Indian butterfly; its upper surface reminds one of
P. daplidice, L., but beneath the hind-wings and tips of the
fore-wings are bright orange with brownish veins. _Experi-
ence at Lahore confirmed me in the opinion that “ Whites ”
of all sorts are most difficult to catch; they are shy, and
fly rapidly with a jerky vertical movement. ‘“ Whites”
are by far the most conspicuous butterflies, especially
when at a distance, and doubtless they need their swift
wings. The Catopsilias are nearly as conspicuous as the
true Whites, and they fly even more swiftly.
Among the Nymphalids the widespread Atella phalanta
was represented by a few specimens at marigold flowers.
Precis orithyia (an insect that suffers much loss of beauty
from grease) was not common, the same is true of P. almana ;
a few of each were taken at flowers. At zinnia flowers I
got my first Hypolimnas misippus, L.,a male ; it impressed
me as a most tropical-looking insect, though not so
gorgeous as H. bolina; it had both hind-wings clipped,
possibly by a bird.
The Blues were represented by two species—the neatly-
marked Tarucus telicanus, Lang., common at the flowers of
Plumbago, and the little greyish-blue Zizera maha, Koll,
abundant at the flowers of a species of millet and some herbs
of the labiate family; amongst them was a specimen of the
var. diluta, Feld. Blues swarm in India, many of the
species are small and dingy, so that they are hard to follow
on the wing, and their flight is even more jerky than that of
Whites. ‘They are often found on grassy banks as at home,
but are especially addicted to water-drinking and are
constantly present in irrigated fields and gardens. It
must be confessed that the abundance of bigger game often
led one to pass them by. Blues when killed are apt to
fold their wings the wrong way, and it is difficult to set,
them right; but if only kept a short time in the bottle
with a view to preventing this untoward result, they are
apt to recover and fly away when the paper is opened after
the day’s work.
The dull-coloured Skipper Gegenes nostrodamus, Fab.,
was common in the gardens, but I only took one female.
Small moths, especially Pyrales, were abundant in a patch
of long grass and herbage in a damp spot. One of these
Observed in a towr through India and Ceylon. 77
was Pyrausta incoloralis, Guen., another the tiny Gold-tail,
Porthesia marginalis, Wk., which was flying in the sun.
There was also the very widely-distributed Marasmia
trapezalis, Guen.; but by far the commonest was the pretty
little black-and-white Hymenia recurvalis, Fab. (very
suggestive of our H. cingulalis, L.).
In the gardens of the Shah Dara, Jehangir’s mausoleum,
four miles from Lahore, I saw at dusk a number of Hawk-
moths at the yellow tubular flowers of a small tree. My
short-bandled net only allowed me to catch two, which
proved to be beautiful specimens of Vephele hespera, Fab.,
and Cherocanvpa celerio, L.
In writing to Dr. Dixey from Lahore I made the
suggestive remark: “It is evident that being late in the
autumn many of the butterflies are old and much worn.
Curiously enough they are more often tattered and torn
than actually rubbed.” It is difficult to rightly apportion
the breaking of the wings between the work of thorns and
insectivorous foes. Certainly Indian butterflies fly into
and through bushes in a way that one does not see at
home.
In the Ajdib Ghar, or Wonder House of Lahore, Anglicé
Museum, well known to readers of “ Kim,” is a small
collection of insects. This was useful to me, but the
destruction wrought by Dermestes, etc., both among the
insects and the textile fabrics of the Industrial Collection,
is most sad to see. I trust Mr. Kipling will see to it.
Amritzar, lat. 81° 40’ N., alt. circa 750 ft.
November 5th and 6th, 1903.
At the sacred city of the Sikhs my collecting was
practically confined to two gardens close to the hotel.
Here a large dull brown butterfly, with somewhat of the
Vanessa habit, spread itself perfectly flat upon the surface
of the earth and more especially of the damp mud of the
little irrigation channels, lying so close to the surface as to
be with difficulty discerned, so exactly did it resemble the
tint of the mud. I secured three which proved to be
Euthalia garuda, Moore, all females.
Papilio pammon was common ; besides males I took one
female of Wallace’s Form I, which differs but slightly from
78 Dr. G. B. Longstaff’s Motes on the Butterflies
the male and is hence termed pammon pammon. Of
Precis almana 1 took one, of the ubiquitous Belenois
mesentina likewise one, a female, but I was somewhat
surprised to net a Colias fieldwi, 2, since the great plain of
the Panjab seemed an unlikely locality for a Colias.
Yphthima nareda, Koll, was scarcely common in the
hotel garden, it flew close to the ground. The list closes
with Polyommatus beticus and a grasshopper to which Mr.
Kirby cannot assign a name.
Delhi, lat. 28° 30’ N., alt. circa 700 ft.
November 7th—12th, 1903.
When collecting in the Kudsia Gardens at Delhi it was
impossible not to be impressed with the historic associa-
tions of the ground. Lying between the northern walls of
the city, the famous ridge, and the mighty Jumna, scarcely
more than a furlong from John Nicholson’s grave, stands,
nearly hidden by trees and flowering shrubs, all that is left
of the Summer Palace of the kings of Delhi. Its crumb-
ling walls, where not covered by Bougainvilleas or other
creepers, bear testimony by many a bullet-mark and round-
shot hole how fire-swept the place was during the long hot
days of 1857. Concrete blocks with suitable inscriptions
mark the sites of the breaching batteries of the last stages
of the siege—batteries placed strangely near the walls
when measured by the range of modern guns, for yon
breach in the Water Bastion is scarce two hundred yards
from the most advanced battery !
Here in a beautiful garden, the very ideal of quiet and
peace, where the numerous grey-striped squirrels are quite
tame and the greenest of parrots and the crested hoopoes
look as if war were unknown upon earth—here I watched
many gorgeous Papilio aristolochie, Fab., fluttering upon
the flowers, or sailing over the trees ; at one moment look-
ing like black crépe against the light, at another displaying
a circlet of brilliant rubies beneath. Once I had three
together in my net! With these were a few P. erithonius
and P. polytes, the latter females of Form II.
Limnas chrysippus was also common, one, a male, was
unusually small. Cvastia core, Cr., was common in shady
places under mango trees, but was rarely seen at flowers.
The pretty little black and salmon-coloured Teracolus
Observed in a tour through India and Ceylon. 79
calais, Cr., was abundant alike in the Kudsia Gardens and
close to the hotel, flying near the ground yet not so
easy to catch. One of them was very small. Of 7.
puellaris I only saw two. The “wet season” form of
Terias hecabe was abundant, flying low and about bushes.
Of the brilliant yellow and orange Jxias pyrene, L., 1
took but one; the less gaudy Orange-tip, 7. marianne, was
rather common, but some of them were worn and none very
easy to catch. The genus Catopsilia was represented by
one worn male pyranthe, and I took my first Delias eucharis,
Dru., a very worn female. The common Whites were
Huphina nerissa, all males, and Belenois mesentina, which
was abundant at flowers. The slender little Nychitona
wiphia, Fab., flitted weakly along close to the ground,
reminding me irresistibly of Zewcophasia sinapis, L., in
' spite of all structural differences. One of these ghostly
creatures was taken flying over a tablet that marked
the site of “Battery No. IV. Left attack; mortars.”
One wondered whether there were any butterflies in that
place during the terrible summer of 1857.
Three or four Precis lemonias, L., appeared to be rather
fond of shade, they settled upon the ground in preference
to flowers and then were hard to see. Of the gorgeous
Hypolimnas bolina I saw one of each sex; it needed an
effort to believe that they were one species.
The Blues included Catochrysops cnejus, Fab. ; Tarucus
theophrastus, Fab. ; Chilades varunana, Moore; and Chilades
laius, Cr., this last was common. The only Skipper taken
was Telicota augias, L.
A little geometer, like a Macaria, was common among
herbage, Semiothisa fidoniata, Guen., and one specimen of
Tephrinia disputaria, Guen., was taken in like situation.
Semiothisa jfidoniata also came to light, along with
Oligochroa akbarella, Rag. Can M. Ragonot have intended
a deliberate insult to the memory of the great Emperor
when he named this dingy and insignificant little Phycid
after him ?
Other moths taken in the Kudsia Gardens were the
tiny Noctuids Metachrostis badia, Swinhoe, and Larias
tristrigosa, Butl., which was common among bushes near
a back-water of the Jumna.
In the verandah of the hotel I took a fine Sphinx orien-
talis, Butl, the eastern form of convolvuli, L.; it had
probably been attracted by the lights the night before.
80 ~——- Dr. G. B. Longstaff’s Notes on the Butterflies
Ldlkot. November 10th.
Eleven miles south of Delhi lies this glorious city of
ruins, and there, under the shadow of the Kutb minar,
flying over the stones and amidst the thorny vegetation
were many Whites and Orange-tips. The butterflies
appeared especially to delight in flying about inside the
thorniest bushes, or even flying through and through
them, so that torn wings were almost the rule. Prominent
in the countless crowd of Belenois mesentina so employed
were Jxias marianne and I. pyrene; a female of the former
was distinguished by the substitution of cream-colour for
white in the ground-tint of the wings. The delicate-look-
ing Teracolus etvida, lover of ruins, was in abundance,
flying close to the ground.
I saw one black Papilio, one Limnas chrysippus and one
Precis lemonias.
Ndini Tal, lat. 29° 30’ N., alt. 6500 ft.
November 16th—23rd, 1903.
Unlike Simla and Darjiling, which stand astride lofty
ridges, Naini Tal lies in a basin by a lake, a situation
which, however pleasant it may be in summer, gives it in
late autumn a dank feel. In summer it affords good
collecting, but in November I found but few insects and
those mostly battered and forlorn looking. The fauna,
though more Oriental than at Simla, a degree and a half
to the north, was much more Palearctic than at Lahore,
which is yet half a degree north of Simla, but of course
upon the plain.
A very clear picture remains with me of a bright
sunny afternoon, with a raw chill in the air very sug-
gestive of home. On the one hand were rhododendrons and
Thujas growing as forest trees, and hard by cactus-lke
Kuphorbias some fifteen feet in height ; on the other, poplars
were shedding their golden leaves in bright contrast to the
crimson of the wild Ampelopsis (I cannot call it “ Virginian”
creeper), a ‘“‘ Brimstone”* butterfly dashes wildly past,
then a belated “'Tortoiseshell” + or “Red Admiral” + darts
* Probably Gonepteryx rhamm, var. nipalensis, but possibly a
Catopsilia,
t Vanessa kashmirensis. t Pyrameis indica.
Observed in a tour through India and Ceylon. 81
up from the path only to return again to the same stone,
while several “Small Coppers” (Chrysophanus pavana)
disport themselves about the autumn flowers on the
bank. Quite a Palearctic picture surely !
In addition to these I found at Naini Zerias hecabe, the
Hairstreak Zlerda sena, Moore, and the Blue Zizera maha,
Koll., also the Fossor, Pompilus analis, Fab.. 2, while
Agrotis fammatra, Guen., came to light.
A climb to the top of China Peak (pronounced Cheena),
8568 ft. produced two more Palearctic forms, Argynnis
lathonia, var. issva, and Lycena betica.
Five days were spent on horseback in an expedition
into Kumdaon as far as Ranikhet and Chaubattia, some
twenty-four miles north of Nainias the crow flies. The road,
at first slippery with ice so as to compel us to dismount, falls
rapidly to Khairna (Khairana, or Khyrna, for the spelling
seems uncertain). At about 6000 ft. Zlerda sena was again
met with, at about 4000 ft. Meptis astola, Moore, and
Terias hecabe. When near the bottom our eye was caught
by the fluttering down of shells from a large pod-bearing
tree. On looking up we saw about a dozen charming-
looking greenish monkeys, their little black faces set off
with most becoming white frills. It did not take them
long to strip that tree of every pod.
At Khdirna, 3200 ft. a tiny village in a deep and
narrow valley, I had a little time for collecting, but it was
limited by the steady march of the great chill mountain
shadow, which sent all butterflies quickly off to bed.
Precis orithyia was common, but the specimens were very
small; P. wnone, P. lemonias and P. iphita were also seen,
the latter at flowers, not a usual habit: of the species.
Several Athyma perius were seen, also several Catopsilia
pyranthe; of those taken one was the gnoma, the other
of the pyranthe form. Of Ganoris canidia and Turucus
telicanus, Lang, I took one each, but’ Zizera maha, Koll., was
in abundance. In addition to these were Deiopeia pulchella,
flying for short distances about low herbage according to
its wont, and a fly which hovered at flowers just like a
Sphinx—a Bombylius not in the National Collection. The
widely-distributed locust, Zhisoicetrus littoralis, Ramb.,
which was very common, had the curious habit after its short
flight of settling so brusquely upon a shrub as to make its
branches shake, but then quickly making its way to the
ground. I missed a Macroglossa twice at the same bush.
TRANS, ENT. SOC. LOND. 1905.—PART I. (MAY) 6
82 Dr. G. B. Longstaff’s Notes on the Butterflies
Late in the afternoon I took a Papilio pammon, a female of
Wallace’s Form II. polytes, which was flying about and
into bushes, apparently seeking for a resting-place for the
night, but possibly seeking where to lay its eggs.
Close to the village of Khairna I saw upon the cliffs by
the roadside several beautiful lizards, grey-spotted, with
bright blue legs.
On the long and hot way up again from Khairna to the
ridge on which stand Ranikhet and Chaubattia, a dwarf
Precis orithyia and a Neptis astola, Moore, were taken at
about 3500 ft. and at about 4000 ft. Belenois mesentina,
Pyrameis indica, and Llerda sena.
At Ranikhet, 6000 ft. (where, by the way, the cooking at
the Dak Bangla was the best that we came across in India),
monkeys were not uncommon in the woods, but unlike
our legumen-loving friends of Khairna, of a revoltingly
ugly type; butterflies, however, were scarce, and were
represented by Pyrameis cardui, Vanessa kashmirensis,
Tlerda sena, and Lycena maha, Koll., var. diluta, Feld.
At Chaubattia, four miles to the east of Ranikhet, and
at a height of about 6200 ft., the officers’ quarters com-
mand a most glorious panorama of Nanda Devi, 25,749
ft., Nanda Kot, 22,491 ft., and Trisul, 28,581 ft., mountains
of unsurpassed grandeur of form and held most sacred.
by pious Hindus as sources of Holy Ganges. These stand
between fifty and sixty miles away, yet shine forth as clear
and bright as if close to. Here there were rather more
butterflies, viz. our old friends Terias hecabe, Precis anone
and P. lemonias, Pyrameis cardui, and Chrysophanus
pavanda, and in addition something quite fresh, the Erycinid
Dodona durga, Koll., of which I got three specimens ;
though a small insect it proved tenacious of life. A little
beetle, Oides sp., was taken flying over the road.
On descending again from Naini to the plains one found,
as at Simla, that butterflies got more numerous and more
Oriental in character. At the top of the road the Hair-
streak, Zlerda sena, was common; at 5000 ft. Yphthima
philomela, Joh., was met with; at the Brewery, circa 4500
tt., butterflies were very common at a flowery turn of the
road, and I took Pyrameis indica, several Precis iphata,
P. lemonias, and a male Hypolimnas bolina, while I missed
a brown-and-white Neptis-like butterfly which may have
been Rahinda sinuata, Moore.
Observed in a tour through India and Ceylon. 83
Lucknow, lat. 27° N., alt. circa 500 ft.
November 24th and 25th, 1903.
Lucknow possesses a museum containing a fair collection
of insects, which would have been more instructive to me
if a majority of the species had been named.
My scanty collecting was almost confined to public
gardens. Near the hotel was a small institution, either a
children’s orphanage or hospital, and in the garden attached
thereto Hypolimnas misippus, J, was rather common,
but shy and worn; I took a battered one. Of H. bolina
I took a female. Of Delias eucharis the males were
common at Zinnia flowers. Odd specimens of Papilio
aristolochiv and Parnara mathias, Fab., also occurred.
In the beautiful garden of the Dilkusha Palace, where
Havelock fell sick of the illness that was to prove fatal in
the very hour of triumph, there was a great wealth of
flowers and consequently a great assemblage of butterflies.
Besides such things as Papilio erithonius; Argynnis niphe
(a 2); Hypolimnas misippus, several males; Crastra
core, both typical and the variety vermiculata, Butl.; and
a Catopsilia which evaded capture, I took there my first
Rapala melampus, Cramer. This is a small copper-coloured
butterfly belonging to a genus which, with its robust body,
sharp-cut wings, and curious anal lobe to the hind-wing,
looks very different from our Hairstreaks or Coppers. It
is neither easy to see on the small flowers which it
frequents, nor to catch.
Other butterflies taken in the same garden were the
Blues Catochrysops strabo, Fab., and Tarucus telicanus,
Lang, the latter abundant; Mycalesis perseus, and the
brilliant tawny Skipper Zelicota augias, L. A beautiful
little Noctua with yellow under-wings, Hyblea puwera,
Cram., was taken at flowers in full sunlight. The Blue
Zizera argia, var. similis, Moore, was in abundance. I also
took a locust, Gastrimargus marmoratus, Thun., a species
of wide distribution.
By the roadside between Dilkusha and La Martiniere a
few Chilades putli, Koll., a very small brown Lyceenid,
were obtained.
At the Alumbagh, ever to be remembered in connection
with Colin Campbell, the dry-season form of Terias hecabe
was flitting quietly about, and I netted Jxias marianne
(not so vulgar-looking as its name might lead one to
84 Dr. G. B. Longstafi’s Notes on the Butterflies
expect), also a variety of the female of J. pyrene without
the orange-tip. A male of the wet-season form of Hu-
phina nerissa was also taken, while Delias eucharis was
common, a specimen feeding on Zinnia flowers close to
Havelock’s grave. Deiopeia pulchella was flying com-
monly in the sun amongst the grass, and with it a specimen
of Argina cribraria, Clerck. ‘The Coleoptera were repre-
sented by Mylabris sidxv, Fab., and the Micros by a Pyrale,
Pyrausta juncturalis, Wk.
In the Presidency garden I took only a worn Acidalid
and the common Cantharid beetle, Mylabris sidw, Fab.,
which was seen in some numbers flying about the flowers
of a species of Hibiscus.
Benares, lat. 25° N., alt. 270 ft.
November 28th—December 2nd, 1903.
The sacred city of the Hindus proved more remarkable
for the number and variety of its pilgrims than for its
butterflies. In the hotel garden, where jackals howled
loudly by night, a few battered specimens of Papilio
ertthonius were seen by day, and the males of both species
of Hypolimnas were fairly common. Of HH. bolina I took
a fine female, while of mzsippus I also sent home a female
which is marked “common.” It is, however, certain that I
did not know this insect to be a Hypolimnas at the time,
since I only learned from the Calcutta Collection that the
female of misippus was brown! There is therefore little
doubt that I took it for a variety of Limnas chrysippus which
it mimics in such a surprising manner, and which certainly
was common enough in the same garden. It is one of
the inconveniences of the method of enveloping that so
much is left to memory, and the chances of comparing
insects are so very few. Dwarfed specimens of Precis
orithyia were now very common; P. wnone and P.
lemonias were less common but almost as small. Several
P. almana occurred. But in spite of the excessive drought
and the consequent occurrence of dwarfs, one of my
specimens of Zerias hecabe taken at Benares was quite of
‘“wet-season”” type. Catopsilia pomona was represented
by a very large male of the typical form and a smaller
female in fine condition, exhibiting the transition to the
catilla, Cr., or extreme dry-season form. Similarly C.
Observed in a tour through India and Ceylon. 85
pyranthe was represented by a male of the typical and
a female of the gnoma form.
Together with the above were several smaller things:
among the Blues Polyommatus bxticus occurred, whiie
Catochrysops strabo, Fab., and the tiny Chilades putli, Koll.,
were both common. The Skipper Parnara mathias, Fab.,
was also common, and I took one Telicota augias, L. The
little Pyrale, Hymenia recurvalis, was in some numbers in
one small flower-bed. Of the long-waisted wasp, Humenes
esurtens, F., I saw but one 2.
APHNAUS ELIMA, Moore.
Enlarged from sketches from the living butterfly.
Vj (Sif ES
cn. Apex OF FW.
Mpex of FW. x.
PW iniclose apncsitior™’| Wim ap he (ETN h) 1 nee HW. in close apposition
Longitudinal fol
mm HW. Longitudinal
fold in HK : Anterror tails
Anterior Fajls_..-.....
Fverted anal lobe... ap a Posterior talls
SESS ves Everted aral lobe
Posterior tails.......
Diagrammatic view Diagrammatic view
from above. from behind.
Drawn at Benares, November 30th, 1903, by G. B. LoNnastarr.
But among the frequenters of the small garden adjoin-
ing the hotel those that interested me most were the
“lobed” and “tailed” Lyceenids, of which there were no
less than four species. Of Aphnaus ictis, Hew., I took
but one, a male, of 4. elima, Moore (which, however, De
Nicéville considered to be only a dry-season form of icézs),
I secured two, also males. Of the third species, Pratapa
deva, Moore, I took but one, and that had lost the anal
angles, with their appendages, and a large part of both
hind-wings, which had apparently been bitten off, abso-
lutely symmetrically, by a lizard. The fourth species,
Rapala melampus, Cramer, was common, and I secured
seven specimens, all, however, males.
86 Dr. G. B. Longstaff’s Notes on the Butterflies
Concerning &. melampus I wrote to Dr. Dixey at the
time: “The Tailed Copper (or Hairstreak) first seen at
Dilkusha, Lucknow, and found commonly here to-day,
greatly interests me. Not only is it very beautiful, but it
is surprisingly hard to see, especially when at rest. Then
the structure of the hind-wing is most strange; posterior
to the tail (the next interspace but one) a portion of the
wing nearly circular, with a very obvious fringe of large
scales, is set at right angles to the plane of the wing and
to the direction of the veins.”
According to Schatz and Rober* this “anal lobe”
occupies the space between the sub-median and inner mar-
ginal veins; the second anal and third anal of Comstock ;
1b and 1e of Meyrick; but I have not found in these authors
any allusion to the striking fact that this lobe is quite out
of the plane of the wing. This omission may be due to
the fact that the process of setting usually flattens the
lobe out so that it is hardly seen in cabinet specimens.
Jt did not occur to me at the time (and the suggestion
arrived by letter too late) that the object of this structure
is possibly to produce the appearance of a head in a non-
vital part, the tails representing the antenne. However,
drawings made at the time strongly bear out the suggestion.
The resemblance would be still more striking if these
Lyceenids, like so many of the family, habitually rest with
the head downwards.
In another letter from Benares I said: “Thorns are not
specially bad here, only that one does not know the look
of many thorny plants until too late. But, on the other
hand, burrs of every sort and kind abound to an incredible
degree and tangle up the net; much of one’s time is
spent in freeing net and breeches therefrom.”
It might have been added that at Benares I first made
acquaintance (somewhat intimate) with “spear-grass,”
which is yet more provocative of bad language than
either thorn or burr.
Some three miles from Benares, on the way back from
Sarnath, where Buddha first taught, I found Delias eucharis
in extreme abundance in a small field of the tall marigold
which is so much cultivated for the service of the temples.
A truly gaudy sight it was to see crowds of these white,
yellow, and scarlet butterflies upon the orange-coloured
blossoms.
* Die Familien und Gattungen der Tagfulter,
Observed in a towr through India and Ceylon. 87
Calcutta, lat. 22° 30’ N. Near sea-level,
December 4th—12th, 1903.
Naturally one could not expect to turn up anything new
at Calcutta, the capital of India, and long the home of De
Nicéville, that martyr to science who met his death in the
deadly Terai when in pursuit of his favourite butterflies.
However, I determined in the few days at my disposal to
get at any rate a sample of the fauna of Lower Bengal.
The Eden Gardens, abutting on the Maidan and close to
Government House, bear much the same relation to
Calcutta as Kensington Gardens to London, and from their
proximity to the hotel afforded a convenient collecting
ground for odd hours.
The Duranta was nearly over and the most attractive
feature proved to be a hedge of Lantana in full bloom.
These dissimilar plants both belong to the Verbenacew and
are both natives of the West Indies, although the latter
appears to have run wild in many parts of India. On that
hedge Limnas chrysippus was in abundance, accompanied
by L. genutia, which I had not seen since I was at
Malakand, while numerous Zirwmala limniace and Crastia
core completed the company of the Danaids. I was able
to confirm my Benares observation that the male of JL,
chrysippus has a slight but decided odour suggestive of
cockroaches, which is perhaps stronger when the scent
sacs on the hind-wings are opened, though of this I am
not sure. On the.other hand, the male of C, core has
a faint scent that suggested to me rancid oil, or old lamps,
So far as I could judge the scent is connected with the
hind-wings but not with the very conspicuous genital tufts,
At the Lantana flowers along with the Danaids were
abundance of Swastus gremius, Fab., a somewhat dingy
Skipper, also a few of the brilliant and conspicuous Delias
eucharis. The upper-side of the female of this species
faintly mimics 7irwmala; the male yielded on rubbing the
wings a sweet flowery scent, which I was not at first able
to describe, but later it struck me as resembling that of our
domesticated Ganoris rapx and suggestive of sweet-briar.
Dr. Dixey informs me that scent-scales are very numerous
in Delias.
In the shadier parts of the garden together with numer-
ous Zerias hecabe, one at least of markedly wet-season
type, and many Vphthima hiibneri, Kirby, several Mycho-
tona xiphia were found, which, as ever, reminded me of
88 Dr. G. B. Longstaff’s Notes on the Butterflies
Leucophasia sinapis, a slender form and fragile appearance
being in each case associated with a weak flight close to the
ground. One of the Y. hiibnert had the whole hind margin
of both secondaries bitten otf nearly symmetrically.
Catopsilia pyranthe and C. pomona were both met with,
the former the more frequently. No Papilio turned up
although I was told that P. pammon occurs in the garden.
Amongst young palms the males of Hlymnias undularis,
Dru., were occasionally disturbed, and a very striking
thing it is. Then Nepheronia hippia, F.,came along, flying
strongly, the male looking on the wing, or more especially
when settled on a flower with wings expanded, much
bluer than its cabinet appearance might lead one to sup-
pose. Three Limenitis procris, Cr., required some catching,
preferring the leaves of tall shrubs to flowers; but it is
scarcely as graceful on the wing as our White Admiral.
I took two specimens of Catochrysops pandava, Horsf.,
var. bengalia, De Nicév. (being the dry-season form); the
female is a dingy creature, but the male is of an iridescent
blue, bordered with black. Hypolimnas misippus, f,
Precis almana and P. lemonias completed the list of twenty
species taken in four visits to the gardens. With them
was a bee Lis thoracica, Fab., a 9.
Baligany.
At the truly splendid museum (where, by the way, I saw
a native artist at work producing some of the very best
coloured figures of beetles and butterflies that I have ever
seen), Mr. 8. E. Peal, besides helping me in other ways,
put me on the track of one of the late Mr. De Nicéville’s
favourite collecting-grounds, a rus in urbe, at Baliganj, a
suburb only three miles from the hotel. I visited this
place twice, on December 5th and 9th. It consists of a
large deserted garden long run wild; weedy meadows and
jungly woods are all that is left of trim lawns and ordered
shrubberies, while a palm avenue and several tanks covered
with a floating flower of the convolvulus order, harbouring
countless dragon-flies, complete the tale of departed great-
ness. Altogether it is full of sad beauty. Palms and
crotons with an undergrowth of ferns were the char-
acteristic plants, flowers were few, yet in certain favoured
spots butterflies were in quite bewildering swarms. The
quiet charm of this old garden was greatly enhanced by
the absence of curious natives and the (comparative) absence
Observed in a tour through India and Ceylon. 89
of burrs, that curse of ‘‘up-country ” collecting, thougli the
unsuspected prickles of innocent-looking palms to some
extent took the place of the latter.
Some of the species seen near the centre of the city, in
the Eden Gardens, were conspicuous by their absence,
e.g. Limenitis procris, Precis lemonias and H; ypolimnas
misippus.
The four common Danaids, Zirumala limniace, Crastia
core, Limnas genutia and L. chrysippus, were not so
common as might have been expected, probably owing to
the scarcity of the flowers they love. In the last-named
species I was able once more to confirm the presence of a
distinct, but not strong, odour suggestive of cockroaches.
A few Papilio polytes, both sexes, gave to the assemblage
that air of distinction which the genus always has. Among
the more sombre things, most frequent under the shade
of groves, were a number of Mycalesis indistans, Moore,
together with one JZ. perseus, Fab., which so far as observed
have no “list” when at rest. In the shade also were two
or three Melanitis ismene, Cr. Close down among the
herbage together with Yphthima hiibneri there were flying
large numbers of Y. philomela, certainly a gregarious
species.
Precis almana was noted and P. atlites, Joh., here came
under my observation for the first time, but in poor con-
dition ; it is then a rather ghostly-looking butterfly though
a somewhat strong flier; this last is also true of Atella
phalanta. A single specimen of my old Amritzar friend
Euthalia garuda was observed, as before, to settle with its
wings fully expanded and closely appressed to the ground.
Elymnias undularis was in abundance ; it is especially
addicted to the characteristically Indian butterfly habit of
flying into or through bushes, and even of flying about
inside bushes. It is clearly oregarious, several specimens
flying about and in one palm- bush, its food-plant. The
male is very striking on the wing, and when settled, even
though the under-side is somewhat leaf-like, it is yet
quite conspicuous. The female, on the other hand, ig on
the wing a very fair mimic of Limnas genutia, but its
flight is ‘weaker.
“Catopsilia pyranthe and C. pomona were both rather
common; TZerias hecabe was abundant, and, as usual,
gregarious,
In half-shaded spots an occasional Nychitona xiphia
flitted slowly along close to the ground, Lrgolis ariadne,
90 Dr, G. B. Longstaff’s Motes on the Butterflies
L,, was abundant, of HY. merione, Cr., two specimens were
secured. The butterflies of this genus settle with the
wings three-fourths expanded.
Nepheronia hippia was rather common; though its
female somewhat mimics Zirwmala limniace, the male,
when on the wing, looks much bluer than that insect.
A brilliant fulvous Skipper, Zelicota bambus#, Moore,
was the only representative of the group, but there were
several Blues, to wit Catochrysops strabo, Fab., which was
common ; Lampides celeno, Or., larger than usual, one being
of the form alewis, Stoll.; and Neopithecops zalmora, Butl.,
was common. A single example of Curetis thetys, Dru.,
fell to my net, apparently bitten by some enemy; its
silvery white under-side is very striking. But perhaps the
strangest-looking butterfly of the lot was Loxura atymnus,
Cr., of which I got two. Its wings are much plaited
longitudinally, and when at rest its extremely long tails,
crumpled look, and brown colour give it quite the look of
a dead leaf. A closer examination shows that the portion
of the hind-wing near the anal angle is bent down, or
back, nearly to a right angle; this bent portion is, how-
ever, relatively smaller, more oval and less sharply bent
than the rounder anal lobes of Rapala and Aphnxus ;
moreover it is not furnished with the very large marginal
scales which are so conspicuous in those genera,
Toliganj. December 7th, 1903.
About two miles from Baliganj, and due south of
Calcutta (about half-a-mile beyond the Sports Club), is the
locality referred to as Toliganj. Here too is an old
abandoned garden, but lacking the elements of departed
grandeur that give a poetic colouring to De Nicéville’s old
hunting-ground. The prominent features are a great
profusion of Lantana in full bloom, a bamboo grove and a
good deal of thorny jungle. The day that I was there the
Lantana was the chosen haunt of great numbers of the
bigger butterflies such as Delias ewcharis, Tirwmala limni-
ace, Limnas genutria, Papilio pammon, mostly worn, P.
arwstolochie, and a few P. erithonius, together with an
occasional Nepheronia hippia, with his broad wings proudly
expanded to view. ‘The sight of these big fellows, expand-
ing from three to four inches, quietly settled on the flowers,
or fluttering after the manner of Papilio, or grandly
sailing around gorgeous in their white, yellow and scarlet,
Observed in a tour through India and Ceylon. 91
black and grey blue, mahogany-brown and black, black and
cream colour, black and coral-red, black and yellow, or sky-
blue and black, afforded indeed a glorious sight not soon to
be forgotten. Alas! such a tropical glory takes much
colour out of the most vivid mental pictures of butterfly
life at home.
In.a shady grove not far from these flowers Limnas
genutia was simply swarming, as many as ten or even twenty
being in sight at once, for it is one of the most gregarious
butterflies that I have met with. A few observations on
this species and Z%irwmala limniace failed to detect any
odour, but it was far otherwise with Delias eucharis, of
which several specimens had a distinct sweet scent, very
like that of G. rape. My strong impression is that this
scent is confined to the male, but I cannot, unfortunately,
speak with certainty on the point. The male of Huphina
nerissa has a distinct scent, also like that of G. rape,
although the butterfly more resembles G. napi. The scent
of these two butterflies is neither so strong nor so unmis-
takably characteristic as that of G. napi, but its existence
is quite beyond question,
These scents are not easy to deal with. The human
nasal organ is but a poor affair at best, moreover scents
are very hard to describe, and these butterfly odours are
only suggestive of, certainly not identical with, those to
which I have, for want of any better standard, compared
them. Then the scents are transient and may easily be
scattered by the wind or overpowered by neighbouring
flowers. Again the scales, independently of any scent, are
irritating to the mucous membrane. Lastly, any one who
has tried to use the sense of smell for diagnostic purposes
must know how even the most volatile perfumeis apt to linger
on, lurking as it would appear in the cavernous recesses of
the nose. Of course it is much easier to determine in the
field whether or no a scent is sexual in those species in
which the sexes are distinguishable by very obvious
characters. Lastly, it should never be forgotten that in all
probability the scents described are far more obvious to
the insects themselves than to human observers.
Only a solitary representative of the Huplea group
appears among the Toliganj specimens, but its envelope
bears the note: “Common, has a slight peculiar scent,
rather disagreeable.” Most probably I believed this at the
time to be the common Calcutta species Crastia core, but
it turns out to be Pademma kollari, Feld., and it 1s now
92 Dr. G. B. Longstaff’s Notes on the Butterflies
impossible to say what those that I passed over or missed
were.
In variety the Toliganj flies were disappointing, but,
besides the above named, they included a very fine
female Jxias pyrene, the sole Orange-tip seen at Calcutta;
a few Catopsilia pyranthe; several Ergolis ariadne ;
Elymnias undularis, not common; plenty of that. very
distinct Blue, Neopithecops zalmora; a single specimen
of Loxura atymnus, and plenty of Yphthima hiibneri,
Y. marshalli, Y. philomela and Nychitona xiphia.
The list is closed by “ Melanitis ismene, lover of darkness,
as its name seems to say. It flitted about everywhere
dressed in all the tints of fallen leaves, or, alighting among
them, fell partly on one side and was one of them.” *
I quote the words of E. H. A., that keen observer and
telling writer. The few specimens that I saw that day
were very dark and of the “dry season” form. A note
made at the time says: “ This shade-loving species, which
only flies for a very short distance and settles on the
ground, has a ‘list’ to the right of 20-30°, making it very
like a dead leat.”
A parasitic bee, Crocisa histrio, Fab., was caught feeding
on the wing like a Sphinx.
On December 8th, I visited the grand Botanic Gardens
at Howrah, but it was too late in the day for many butter-
flies to be about. I noted, however, Zampides celeno, Cr. ;
Mycalesis indistans, Moore, a Delias and two or three Terias.
Late in the afternoon, just before leaving the gardens, I
noticed a few Limnas genutia fluttering about a palm-tree
prior to settling down for the night. On looking carefully
I noted on one of the huge leaf-stalks, some twelve or
fourteen feet from the ground, a cluster of the butterflies
hanging together like swarming bees. By pelting with sticks
and stones the cluster was broken up and proved to consist
of at least seven or eight individuals. Altogether there
were perhaps twenty in and about that tree. This cer-
tainly establishes for ZL. genutia the character of gre-
gariousness. Both Mr. 8. E. Peal and Mr. F. Moller told
me that they had never seen such a thing.
Darjiling, 27° N., alt. 7000 ft.
December 13th—22nd, 1903.
I set off to this celebrated hunting-ground with great
* “ A Naturalist on the Prowl,” p. 203.
Observed in a tour through India and Ceylon. 98
misgivings as to season, but still full of wild hopes. The
ascent by the cog-wheel railway took us through a most
interesting forest, where amongst other things we saw our
first tree-ferns. Near Tindaria, at about 3000 feet, I saw
several xias pyrene and succeeded in catching one from the
train while it was going at full speed—about seven miles
an hour! 'lhis was a male of the large form evippe, Drury.
Before reaching Kurseong, nearly 5000 feet, where I
had arranged to sleep with a view to getting a little
collecting, we got into the clouds, and at our destination it
was very cold, with an atmosphere only too like that of
the West of Scotland. During a gleam of sunshine I took
a Vanessa kashmirensis, a poor dull thing compared to
our urtice. At night two moths, an Acidaliid, Synegiodes
hyriaria, Walk., and Caradrina albosignata, Oberth.
(thought by Sir G. Hampson to be probably identical
with dincosa, Moore), came to light.
The following morning was brighter and we started
early to walk up to the next station, Toong, but though
the weather was more benignant, the railway ran through
a district devoted to tea-growing which did not promise
well. A few Vanessa kashmirensis, a Pyrameis indica and
a P. cardui flew along the road, the latter with both apices
of the fore-wings and one hind-wing near the anal angle
apparently bitten. Near 'Toong station, 5500 feet, in a
sheltered and flowery spot I took single specimens of the
Hairstreaks Jlerda epicles, Godart, a female, and Camena
cleobis, Godart; the latter on the upper surface like 7’.
quercus, but bluer, on the under-side almost white with a
practically black spot on the anal lobe. Here also I took
a male Hiposcritia lalage, Dbl., and a native caught in his
fingers a Dodona eugenes, Bates (an Erycinid), and the same
man brought me a fine Saturnid moth, khodia newarra,
Moore, 64 inches in expanse, apparently recently dead.
As the train rounded the last corner we came in full
view of the Kangchinjunga range, rising majestically full
four miles above us. No words can describe the grandeur
of the scene and we were fortunate indeed in having it
clear throughout our stay. Yet, entomologically speaking,
it was the saddest of disappointments, for it was as cold as
England in November and the local entomologists—Messrs.
Moller and Lindgren—assured me that Kallima was
hopelessly over, as indeed were most things. They, and
every one we met, spoke of the astonishing multitudes of
94 Dr. G. B. Longstaff’s Notes on the Butterflies
butterflies in the rainy season ; the harder it rained, they
said, the more insects there were in the short interludes
between the showers.
On the high ground I got little; Vanessa kashmirensis,
Pyrameis indica and cardui, none of them common, also
a brilliantly-coloured beetle, a species of Cassida. I saw a
school-boy catch Colias fieldit and a tailed “blue.” At
Jalapahar, 7500 feet, I got a female Huphina nadina,
Luce. [ =remba, Moore].
The only chance was to go down into the valleys, but it
takes long to descend, and as the butterflies are for the
most part only “at home” from 10.0 a.m, to 2.0 p.m., one
does not get many hours’ collecting ; moreover, from those
precious hours there are deductions to be made for cloudy
times, and for the shadows of woods, and the still deeper
shadows of lofty mountains.
My first expedition, lasting three days on horseback,
was to the Tista valley, lying to the east of Darjiling.
December 17th, we went to Pashdk, about 17 miles,
sleeping at the Public Works Department rest-house,
about 2300 feet above the river, and about 3000 feet
above the sea.
When we got down to about 4000 feet above sea-level
insects began to get fairly numerous, although it was late
in the day for butterflies. Vanessa kashmirensis was
common, and with them were several Pyrameis indica. I
secured two of the handsome White Hiposcritia lalage
[=argyridana, Butl.], both females. Several Neptis astola,
Moore, were seen, mostly worn. At a shady turn of the
road I got Lethe rohria, F., an xgeria-like Satyrid butter-
fly ; close by Arhopala areste, Hew., flashed azure in the
sunlight, but a specimen of another beautiful Lyczenid,
Spindasis vulcanus, F., was badly battered. Of <Abisara
flegyas, Cr., and A. fylla, Doub., I netted one each, and a
large bee, Bombus funerarius ?, Smith, a g, tempted me to
catch him.
In the wood in which the rest-house stands Mycalesis
indistans, Moore, was in abundance ; this is a typical shade-
lover; when kicked up from the herbage it flaps about
three yards, like Zpinephale janira, L., and then settles on
dead leaves or on the earth. Some of them had a shght
“list,” but this did not seem to be a marked habit, possibly
because this position is not so advantageous in shade as in
sunlight, though the habit was first noticed in Melanitis,
Observed in a tour through India and Ceylon. 95
a typical shade-loving genus. The existence of shade-
loving butterflies would seem to be correlated to a tropical
sun, but even in England P. wgeria and E. hyperanthus,
L., still more Z. sinapis, are what I should term “ partial
shade-lovers.”
In the same wood, also in the shade, Zerias leta and 7’.
hecabe were both common, and in sunnier glades the
common Indian Blue, ZLampides celeno, Cr., was both
abundant and gregarious. Ina young cinchona plantation
close by I found Ganoris canidia ; a Blue, Cyaniris puspa,
Horsf. ; and a handsome very large Skipper, Celenorrhinus
lewcocera, Koll. In the rest-house there was a dead
Pyrale, Lepyrodes geometralis, Guen.
The next day, December 18th, I sent my pony on ahead,
and walked down to the river collecting on the way. In
the cinchona plantation close to the rest-house I found
Zemeros flegyas ; lower down were Luthalia appiades, Mén.,
of which I saw several worn specimens; a Hairstreak,
Arhopala bazalus, Hew., like a glorified 7’. quercus; Lethe
verma, Koll.; and Melanitis ismene, the last as usual a
shade-lover.
A path leading off through the wood brought me to a
tea-garden, perhaps 1500 feet above sea-level, where I
lingered all too long. ‘Tea-gardens are not as a rule good
places for butterflies, and the flower of the tea-plant, then
just coming out, does not appear to have attractions for
them, but this particular garden, just at the edge of the
forest, and especially that corner of it where the little
stream runs in, was certainly very prolific.
Athyma ranga was in abundance, though worn; like-
wise its relatives of the genus Neptis, but the three
specimens that I sent home belonged to as many species:
NV. aceris, Cramer; N. astola, Moore; and N. varmona,
Moore ; the closely allied, but brown and black, Symbren-
thia hyppoclus, Cr., was almost as common among the tea-
bushes. Of the satin-winged Hiposcritia indra, Moore, I
took two females. Of the following I got single examples
only :—Caduga melaneus, Cr., and the very similar Par-
antica melanoides, Moore, two black and white Danaids ;
Athyma selenophora, Koll.; Yphthima philomela, Job. ;
Arhopala centauwrus, Fab.; and Castalius anawra, De
Nicév., 9, while another Blue, Zampides elpis, Godt., pale
and beautifully sheeny, was common. I also missed what
was, I believe, Lbythea rama, Moore. There were in
96 Dr. G. B. Longstaff’s Notes on the Butterflies
addition two moths, an Arctiid, Lewcoma submarginata,
Walk., and a Nyctemerid, the fuscous-and-white Zonosoma
cenis, Cram. ( = interlectum, Walk.), the former possibly,
the latter certainly a day-flyer.
At last I dragged myself away and an hour later reached
a most attractive flowery bank immediately above the
river. This was evidently a great place, for in a very short
time I secured two sadly battered Papilio memnon, L., of the
form agenor, L.; a large male Jaias pyrene with the fore-
wings almost symmetrically bitten near the tip of the costa ;
also an insect that I had greatly desired to take, the lovely
and delicate-looking “ map-butterfly,” Cyrestis thyodamas,
Bdv., in splendid condition. This, a Nymphalid, by the
possession of a well-marked anal lobe to the hind-wing
suggested the Rapala group of Lycznids, but a close
examination of the veins shows that neither lobe nor tail
is homologous in the two widely separated genera. In ad-
dition to the above I took a second Caduga tytia, Gray, the
first having been netted 1000 feet higher. This blue-and-
black Danaid is distinguished by having brown hind-wings.
Time was however getting on and my “sais” was waiting
with the pony by the little bridge, so I reluctantly mounted.
I had not ridden far when I caught a glimpse of Kallima
inachis, Bdv., flying by the roadside ; flinging myself out of
the saddle I was fortunate in netting the butterfly of all
others that I had wished to see alive. It proved to be a fine
female ; I could not afford to risk waiting to see her settle,
and alas! never saw another. A few minutes later my
sais brought me a damaged Huplwa with a lovely purple
gloss; seeing many about I foolishly did not keep it.
These things happened close to the Tista bridge, by which
the road to Lhasa crosses the river, here only some 650 feet
above the sea, so deeply are these Himalayan valleys cut
down. Sad to say in a few minutes the winding of the
road took me under the deep chill shadow of the mountain
and the purple-glossed Huplwas and nearly all the other
butterflies vanished for that day. <A solitary Neptis aceris,
Cr., together with a few Jxias pyrene, Huphina nerissa and
Lampides elpis, were all that I saw; with them was a.
Nyctemerid day-flying moth, 7rypheromera plagifera, Wk.
The rest-house at Riang was reached too late for any more
collecting, and I had to content myself with watching the
long trains of Colonel Younghusband’s bullock wagons
painfully dragging loads of compressed hay for the Tibetan
Observed in a tour through India and Ceylon, 97
expedition. Alas for the once fair road, now a foot deep
in white dust !
December 19th. From Riang by way of Mongpu and
Saretl back to Darjiling. This was such along march that
little time could be given to collecting, moreover many
hours were spent passing along a beautiful forest track in
the deep afternoon shadow of the mountain. At the start,
close to the river, the silvery-white <Acropteris vagata,
Moore, was conspicuously spread out upon a leaf, this was
the only Uraniid that I met with. Near Mongpu, at about
3000 feet, Hrgolis merione was very common about Ricinus,
the castor-oil plant, upon which its larva feeds, A little
higher up I came across Ticherra acte, Moore, a Lycenid
with very long tails that wave with the wind; it has a
swift jerky flight. The hind-wing of this species is much
plaited but the anal lobe is rudimentary.
Other captures were Huphina nerissa, a male; Ganoris
canidia, a female with all the hind margin of the hind-
wing gone; Zachyris hippo, Cr..a male; Avrhopala rama,
Koll. ; Neptis astola, Moore; Jlerda epicles, Godart, with all
the hinder part of the secondary apparently bitten off by a
lizard ; Cirrochroa aoris, Dbl., which I had seen at Pashék
on the previous day ; Lethe rohria, very like P. xgeria in its
habits and liking for partial shade; and Argynnis niphe,
this last in the cinchona plantation at about 3600 feet.
A large white butterfly, bright yellow underneath, flutter-
ing at the sweet white flower of the cinchona led me to
dismount, and it was well that I did so, for it turned out
to be Prioneris thestylis, Dbl., and fortunately a female,
which must be very much the less common sex, at any rate
the Hope Collection contained no female of the genus.
The next day, December 20th, I rode down to the
Ranjit River, the boundary of Sikkim, the great Papilio
country. Distance however reduced my actual collecting
to less than four hours.
At about 3000 feet I took two of the Erycinid Zemeros
Jlegyas, also Symbrenthia hyppoclus. The chief collecting-
ground was near the suspension bridge leading into
Independent Sikkim, closed this year to all Huropeans,
including entomologists, on account of the Tibetan difficulty.
It was trying to one’s European temper to be stopped by
a coloured policeman, while natives passed freely over!
Here, some 8000 feet above the sea, the first thing that
I happened upon was Limnas chrysippus in extreme
TRANS. ENT, SOC. LOND. 1905.—PART I. (MAY) 7
98 Dr. G. B. Longstaff’s Notes on the Butterflres
abundance in a very limited locality, it was in fact
decidedly gregarious. By the way, pinching and cyanide
are both but very imperfect ways of slaying these tough-
skinned Danaids.
Elymnias undularis, both sexes, was common, but I did
not see any L. genutia for its female to mimic; although
the under-side of this butterfly is “leaf-like,” it is, as a fact,
usually conspicuous when settled.
I caught distant glimpses of two Papilios and I missed
my first Hebomoia, in fact the things that I missed that
day would have made quite a good collection !
The following were all common: Huphina nadina and
H. nerissa, both males; Jxias pyrene, large; Neptis aceris,
Cr.; Precis iphita, spreading out like a Hupithecia when
settled; Symbrenthia hyppoclus, and Lampides celeno, Cr.,
while Zerias hecabe was very abundant and large.
Other things taken were YVphthima marshalli and
Mycalesis runeka, Moore, this last a very dingy species.
In marked contrast was Jamides bochus, Cr., the male
iridescent dark-blue above, quite gem-like, beneath dull
grey with a metallic ocellus at the anal angle of the hind-
wing; the female comparatively dull in colour.
For some reason I that day missed a larger proportion
than usual but managed to catch the following :—Prioneris
thestylis, a male ; Cirrochroa aoris, looking on the wing like
a big Argynnis, but settling with wings half-expanded,
several seen, but only one netted; a Charaxes athamas
taken on a flower was the only individual of the genus
that I got in all my travels. Another specimen of this
very distinct and beautiful species was soon after seen
feeding upon human ordure! Fear of fouling my net
prevented me from striking down upon it, and it suddenly
darted up, went twice round with a swift jerky flight and
then disappeared. Mr. Moller had indeed told me that
Charaxes was a very foul feeder.
The elegant day-flying moth Zrypheromera plagifera,
Walk., must be added to ry list, as well as the little
Geometer Psilocambogia memorata, Walk., which I found
dead, caught and set out upon a burr (of some composite
flower). Lastly a beetle, Mimela horsfieldi, Hope, of
brilliant green with coppery tinge.
The extraordinary abundance of dragon-flies of many
kinds at the Ranjit River was remarkable, yet I did not
once see a butterfly attacked by any of them.
Observed in a tour through India and Ceylon. 99
I left Darjiling on December 22nd, with much regret,
and a strong desire to return at a better time of the year.
On the way down, at about 4000 feet, Zvias pyrene was
common, while close to Tindaria station, at about 2900
feet, I netted from the train a pale Blue with a whitish
patch on each wing, Cyaniris dilectus, Moore, as well as
another Blue, Cyaniris ladon, Cr., form pseudargiolus,
Boisd., and an Acidaliid, Jdxa vremotata, Guen.
At Tindaria I left the train and walked down to Sukna.
The following things were met with: (1) At altitudes of
from 2800 feet to 2000 feet :—
The Erycinid, Zemeros flegyas, Cram., almost abundant,
but rather worn ; A/ycalesis indistans, Moore; Precis lemonias,
a small specimen in fine condition, also large ones worn
[this and P. wphita were the only species of the genus met
with in the Darjiling district]; single examples of Neptis
aceris, NV. astola and N. varmona; Symbrenthia hyppoclus,
common; Lampides celeno, Cr. form alexis, Stoll., also
common; Ganoris canidia, a female; Huphina nadina, a
male, about 3000 feet; H. nerissa, a male, about 2500 feet ;
Tachyris hippo, a very fine female; Zerias hecabe, abund-
ant, two males, one of them dwarfed, were of the variety
without the “dog’s head.”
(2) At altitudes of 2000 feet to 1500 feet :—Here I
took Catopsilia pyranthe, a male ; Yphthima marshalli, two ;
Huphina nerissa, worn males were common ; Precis lemonias,
several; Zerias libythea, common; and the Blue, Zizera
otis, Fab.
At about 3.30 p.m., I watched a fine specimen of Papilio
aristolochiz flying very slowly about herbage, apparently
seeking for a resting-place for the night, just as I had seen
P. pammon doing at Khairna on November 18th; near
the same place I missed two specimens of a black-and-
white Danaid.
(3) A little way above Sukna, perhaps at about 700 feet
above sea-level Orsotriana [Mycalesis] runeka, Moore, was
in the greatest abundance in a deeply-shaded wood ; this, a
typical shade-lover, is sluggish but is on the move later
than most things (for it was just before sundown), but
when kicked up from ferns or other low herbage it did not
fly more than two or three yards. It varies greatly in the
pale streak on the under-side which may be white and very
conspicuous or almost obsolete. One specimen exhibits a
well-marked bite on the hind margin of both hind-wings
100 Dr. G. B. Longstaff’s Motes on the Butterflies
above the anal angle, the injuries on both sides corre-
sponding closely.
It was dark when I reached Sukna station, 500 feet
above sea-level, and fireflies, Lwciola sp., were flitting about
on all sides. <A “flare” lighted just before the arrival of
the train attracted many moths, of which I secured a
large sharp-winged transparent Pyrale, Cydalima conchyla-
lis, Guen.; and the Noctua, Prodenia littoralis, Bdv. In the
train, immediately after starting I bottled a strange-looking
gf winged ant, Dorylus guvenculus, Shuck.
Thus closed my short Darjiling campaign, and leaving
behind with much regret the awe-inspiring Himalaya, we
steamed away into the darkness over the monotonous
plain of Bengal.
Bankapwr, lat. 25° 30’ N., alt. c. 250 ft.
December 22nd, 1903—January 3rd, 1904.
In absolute contrast to Darjiling, Bankapur, the civil
station of the great city of Patna, is situated on the level,
monotonous, and highly-cultivated plain of the Ganges,
affording little harbour for butterflies, so that a fortnight’s
stay with old friends at the hospitable parsonage yielded
small entomological results.
In spite of these unpromising surroundings, Limnas
chrysippus was common, and in company with it Hypolim-
nas misippus, of which I saw several males and secured one
female, which latter so closely mimics the former species
that even the small white spots on the thorax and head
are reproduced !
Of Tirwmala limniace I saw a solitary example, of
Crastia core, two; but the other very common Danaid,
Limnas genutia, was abundant in a mango orchard, and
distinctly gregarious in its habits. It has rather an
unpleasant scent, but whether or no it is confined to one
sex I regret that I failed to notice.
The Satyrids were represented by a solitary Mycalesis
perseus; the Swallow-tails by Papilio pammon, worn,
P. aristolochiv, and P. erithonius, the last a flower-loving
species. Precis was represented by four species: almana,
one of them with large pieces, in part corresponding, bitten
out of each hind-wing ; wnone, one; demonias, several ; and
orithyia, several, the latter all small. Single specimens of
the common and generally distributed Atella phalanta and
Observed in a tour through India and Ceylon. 101
Ergolis merione were seen in gardens. <A fine Limenitis
procris was taken sipping “toddy” from a palm; I missed
him at the first shot, but he foolishly returned to his fatal
liquor.
~ Of Catopsilia pyranthe I took two males and a female.
I held one of the former fluttering beneath my nostrils,
when it gave out a strong scent that instantly brought
greenhouses to my mind, then my own greenhouse, then
Polianthes tuberosa (barbarously termed by nurserymen
“tuberose ”), and lastly jasmine. I do not think that I
ever smelt so distinct a scent in a butterfly, always except-
ing the male of Ganoris napi. The other male pyranthe I
held under my nose while I stroked the “ feather-tufts ” of
the hind-wing ; this at once elicited the odour of jasmine,
further confirming the observation of Wood-Mason.
Two males of Huphina nerissa bear the following notes :
“Scented, not like napi, more like rape,” and “this
specimen had a scent like P. rape, i.e., of the sweet-briar
type.” Again a female of Delias eucharis (which was
common) bears the note, “has a scent much like rape,”
and the specimen appears to have been wilfully rubbed.
My observations on butterflies in England show that in
some cases females have a scent, but not like, or as strong
as the males. My strong impression is that the male of
D. eucharis has the rape, or sweet-briar scent.
The three species of Terias, viz. hecabe, libythea, and
lxta, were all common; one of the /xia appears to have
been bitten by a bird.
Nychitona xiphia was not uncommon, and several Jxias
marianne were seen. Chilades varwnana, Moore (according
to De Nicéville the wet-season form of C. laius, Cr.), was
common about irrigated flower-beds, indeed Blues are
wonderfully fond of water. The only butterfly seen at
Bankaptr that was at all out of the common, besides
Limenitis procris, was the large grey Lycsenid Virachola
isocrates, Fab., of which I took one at flowers in the
Commissioner’s garden. I noted that its hind-wings were
much folded posterior to the tails, the convexities of the
folds being towards the upper-surface. These foldings of
the wings are not well seen in set specimens.
Although Bankapur is far from being a good locality, it
will give some idea of the abundance of butterflies in India
when I say that in mid-winter, December 24th, I took in
a suburban garden within three-quarters of an hour no less
102. Dr. G. B. Longstaff’s Noles on the Butterflies
than ten species, some of them represented by numerous
individuals.
Buddha Gaya, lat. 24° 42’ N., alt. c. 500 ft.
December 30th and 31st, 1903.
The vicinity of the shrine and its sacred Bo tree was not
productive. All the butterflies that I saw there were one
Terias libythea, a number of Huphina nerissa (the male
yielding a distinct, but not strong, flowery scent), together
with a lot of the Lyceenid Zizera karsandra, Moore.
The next day, on a steep hill of red trap rock overlook-
ing the town, I saw for the first time the Acreeid J'elchinia
violx, reminding one on the wing of Argynnis ewphrosyne ;
it was locally abundant and gregarious, its tone of colour-
ing harmonizing with the red igneous rock. On the same
hill were two or three Precis wnone and several small
P. orithyia, while Zizera otis, Fab., was abundant.
In the course of this walk I noticed a Fakir, or religious
mendicant ascetic, watching my operations with evident
suspicion, probably owing to the reverence in which some
of these folk hold all animal life. Presently a small native
boy threw a stone at a squirrel. I thought better of the
Fakir when he cursed the boy so fiercely that he fled in
terror as fast as the squirrel, while I rolled up my
umbrella-net and passed on, trying to elude observation !
Mozufferpur, lat. 28° 8’ N., alt. c. 300 ft.
On a flying visit, January 2nd, 1904, to this place, nearly
north of Bankapur, I took in my host’s garden two Zizera
otis, Fab., and one Zizera maha, Koll.
Allahabdd, lat. 25° 30’ N., alt. 370 ft.
Here on January 4th I saw a few of the very commonest
Indian butterflies in the public garden. The railway
carriage before leaving in the evening produced a grass-
hopper, Atractomorpha (Perena) sp., and Prodenia littoralis,
a Noctua that came to light. This last proved tenacious
of life, it laid a number of eggs in its paper which batched
on the voyage, the young larv perishing miserably.
Observed in a tour through India and Ceylon. 103
Shdnsi, lat. 25° 30’ N., alt. c. 750 ft.
January 5th—13th, also 21st, 1904.
Situated on a sandy plain, broken by precipitous ridges
of igneous rock, Jhansi, something like 750 ft. above the
sea, 1s characterized by dryness, heat, and sparsity of cover.
A couple of, Papilio aristolochiv taken at flowers near
the lake were the sole representatives of their family.
Several Belenois mesentina were taken, but it was scarcely
common; the male had a distinct but faint, sweet scent;
on the ridge of Retribution Hill (where Sir Hugh Rose in
1858 slew 2000 mutineers), | took a female B. mesentina
in which the hind-margins of the secondaries had been
symmetrically broken off, probably by the bite of a lizard.
Of three specimens of Zerias hecabe, one, a sinall female,
was of the variety without the “dog’s head” notch. Of
T. libythea a single specimen was taken, but 7’. /wta was
common and of gregarious habits. Zeracolus etrida was
locally rather common, especially the female. On the other
hand, the male of Jxias marianne was rather common.
Two Atella phalanta were taken; the only Precis noted
was orithyia, and that very dwarfed, one measuring only
1-4 inches across the wings.
Telchinia viole was abundant at the foot of Retribution
Hill, and scattered specimens occurred elsewhere. This
insect, like the Danaids, has a tough skin which enables it
to resist pinching, and doubtless makes it indigestible.
When injured a yellow juice exudes; a minute drop of
this placed on the tongue tasted somewhat bitter and
disagreeable, but the flavour was by no means strong.
The Jhansi Lycenids were fairly numerous, but not
very brilliant, the most striking was Chilades putli, Koll.,
actually smaller and darker than our alsws; other species
were Chilades laius, Cr., which appears to have been
common, but of which I unfortunately took but one
specimen, and Catochrysops contracta, Butl., of which I took
two; Turucus theophrastus, Fab., of which the two sexes
are, on the upper surface at least, very different, was
common, but of 7. telicanus, Lang, I only secured one of
each sex, though noting it as common. Blues are very
abundant in India, but they are very much alike, so that
being ignorant of the distinctions between allied species,
one was but too apt to neglect them while in the eager
104 Dr. G. B. Longstaff’s Notes on the Butterflies
pursuit of larger game. For these reasons too much
weight should not be attached to the observation that
such and such a species was common or abundant, but the
qualification, “or something superficially like it,” should be
added.
Two moths came to light, an Agrotid Huxoa spinifera,
Hiibn., and the AMacaria-like Semiothisa frugaliata, Guen.
I also took a brown beetle, Bolboceras quadridens, Fab.
After prolonged drought there was a heavy rainstorm at
Jhansi on January 14th, and there was slight rain at
Gwalior on the 16th and 17th. With the exception of a
very few days when there had been clouds and occasionally
a few drops of rain, there had been almost uninterrupted
sunshine for three months, 2. ¢. since October 8th. On
January 20th, writing to Dr. Dixey, I said, “ There has
been a very cold ‘wave’ in Northern India with a few
showers of rain, but scarcely enough of the latter to affect
either vegetation or insects.” On January 28rd there was
gentle rain at Jhansi lasting several hours.
On January 21st I had another day’s collecting at
Jhansi, but the species taken were not such as to show
any effect in the way of change of type due to the rain,
even if such change had been possible. The insects met
with were B. mesentina, I. marianne, T. etrida, A. phalanta,
and Tarucus theophrastus,
Orcha.
On January 9th I had an hour's collecting in this inter-
esting deserted city, some eight miles to the east of Jhansi,
and took or saw Limnas genutia, Precis lemonias, P. enone,
and P. orithyia (this last in abundance), Atella phalanta,
an Inias, Teracolus etrida, a Terias, and several female
Belenois mesentina. Monkeys were almost as common as
butterflies among the ruined tombs.
Burwa Sagar.
On January 14th I got a couple of hours’ collecting in
the neighbourhood of the interesting and romantically-
situated old castle of this name, which lies some twelve
miles to the east of Jhansi.
Here I observed in two specimens of Limnas chrysippus
(of which certainly one was a male) a distinct cockroach-
like odour, sufficiently strong to be perceptible when the
Observed in a tour through India and Ceylon. 105
insect was fluttering in the net. Of Catopsilia pyranthe I
took a female of the gnoma form; of Terias hecabe a male,
the variety without the “ dog’s head” mark. 7. /wta was
quite abundant. ‘Two specimens of Huphina nerissa were
taken, one worn, the other a dwarf. The male of Belenois
mesentind was common, in two specimens I detected a
sweet scent like that of P. rapx, but more or less faint. A
Polyommatus beticus completes the list.
Agra, lat. 27° N., alt. 550 ft.
January 25th and 26th, 1904.
At the sight-seeing centre of India there was but little
time or opportunity for entomology. In the fair gardens
of the Taj Mahal Zimnas chrysippus was abundant, three
or four Papilio aristolochiw haunted the brilliant orange-
coloured flowers of Bignonia venusta, and a few Belenois
mesentina were flying around. In the Government garden
close by I also saw the Belenois, together with Huphina
nerissa, Precis orithyia, and P. lemonias, Limnas chrysippus,
and L. genutia, also. a Teracolus, and some Blues which
escaped capture.
Fathipur Sikre.
January 28th and 29th, 1904.
At the abandoned capital of Akbar the Great, the
Pompeii of India, some twenty-two miles west of Agra,
those ruin-frequenting butterflies, Belenois mesentina and
Tevacolus etrida, were both common, but all appeared to be
males. The Belenois had a faint, sweet, flowery scent,
which did not appear to me to be quite like that of any
other insect.
I also took one Zeracolus puellaris, a female, and a most
ferocious hornet, Ewmenes dimidiatipennis, Sauss., a ¢.
Jéipur, lat. 27° N., alt. 1600 ft.
February 2nd, 1904.
The fine public gardens of the enlightened Maharajah
are too well kept to be a good collecting-ground. Terias
leta was however to be had there [as well as at the
deserted capital Ambér, a few miles to the north and on
higher ground]; those taken were males; a very small
106 Dr. G. B. Longstaft’s Notes on the Butterflies
Limnas chrysippus seemed to bear evidence of the prolonged
drought. The genus Papilio was represented by avistolo-
chiv, and the Chrysid Stilbwm splendidum, Fab., did its
best to gratify the Rajputs’ love of brilliant colour.
Ajmir, lat. 26° 30’ N., alt. c. 1800 ft.
February 4th and 5th, 1904.
The most notable capture here was Teracolus fausta,
Oliv., of which I only got one male, a poor specimen,
missing two others; it has a very distinct orange look on
the wing, and I feel sure that I saw one on January 22nd
at Palipahari, the artillery camp near Jhansi.
Of 7. etrida I took two males, one of them had lost the
apex of the left fore-wing and all its hind-margin, as well
as the apex of the left hind-wing. This is notable as
possibly being an attack on a “direction mark.”
I saw several battered Precis wnone. The smaller fry
were represented by a very neat little chequered Skipper,
Hesperia galba, Fab. The emerald-like Stidbum splendidwm
again turned up.
On Taragarh, the precipitous hill that overtops the city
by perhaps 500 ft., I got only Belenois mesentina, Terias
lxta, and the long-waisted 2 wasp, Humenes dimidiatipennis,
Sauss.
Mt. Abu, lat. 24° 30’ N., alt. of civil and military
station c. 4100 ft.
February 6th—8th, 1904.
Insects were extremely scarce upon the sacred Jaina
mountain. The commonest butterfly was Terias leta ; ib
was abundant up to 4500 ft., and the only representative
of the genus seen. These, together with Belenois mesentina,
Huphina nerissa, a few Precis lemonias, and a couple of
tages-like Skippers (which I missed upon rocks at about
4400 ft), were the only butterflies that I saw on the
elevated plateau. One moth, the very widely-distributed
Crambus, Hromene ocellea, Haw., came to light.
At lower elevations, on the fine road up from the plain,
the following were met with: at about 3000 ft., Belenovs
mesentina, Taracus telicanus, and Polyommatus beticus, the
last. as usual in poor condition. From 3500 ft. down to
Observed in a towr through India and Ceylon. 107
2500 ft. a few Yphthima inica, Hew., were seen, and at
about the last-named elevation, among the rocks of a
nearly dry water-course, I saw two specimens of the
beautiful Nymphalid, Symphedra thyelia, Fab., but only
secured one. It has the habits of a Vanessa; unfortunately
time was pressing, or I might probably have taken more.
Bombay, lat. 19° N., near sea-level.
February 10th, 1904.
In an hour’s visit to the Victoria Gardens, where there
were a fair number of insects, I got Papilio erithonius,
P. aristolochix, Neptis varmona, and Nepheronia hippra, a
female, the last named mimicking 7vrwmala.
On February 15th I was much interested in watching
the movements of a solitary butterfly in the small public
garden of the University, in the heart of the city. It
appeared to be a large Catopsilia, possibly the catila form
of pomona, but at any rate of a general greenish-yellow
colour; when disturbed it invariably settled in one or other
of several small shrubs with yellow leaves, when it would
vanish quite suddenly. It was only after several attempts
that I succeeded in getting a glimpse of it when settled, so
strong was the protective resemblance.
Bijapur, lat. 17° N., alt. c. 1500 ft.
February 16th and 17th, 1904.
This was further south than I had yet collected, but the
scanty vegetation among the ruins seemed too parched to
yield very much. The most prevalent genera here, as at
so many places where thorns, burrs, rocks and ruins pre-
dominated, were belenois and Teracolus, the last a genus
which, though beautiful in the cabinet, is not effective on
the wing. .
Teracolus etrida was abundant, the males appearing to
be about twice as numerous as the females; they varied
greatly in size, so much so that among the males the
largest had nearly double the alar expansion of the
smallest. Of 7. dulcis I took one female, and of 7’. amatus,
var. modestus, two males.
The only Zerias seen was leia. Lelenois mesentina was
abundant; a slight sweet scent was detected in one
Specimen.
108 Dr. G. B. Longstaff’s Motes on the Butterflies
Catochrysops strabo, Fab., was common, also Polyommatus
beticus, one specimen having lost two-thirds of each hind-
wing, presumably the work of some enemy; of Zizera
karsandra, Moore, I took one.
At night several moths came to light, viz. the Ocneriad,
EHnome detersa, Walk., the Geometers, Tephrinia disputaria,
Guen., and Jdva fibulata? Guen. (worn), and the very
widely-distributed iiella zinckenella, Treit.
A ferocious-looking spider, a Solpuga, shared the Dak
Bungla with us.
Anantépur, lat. 14° 30’ N., alt. c. 1500 ft.
February 18th—238rd, 1904.
This small civil station, situated on an irrigated though
elevated plain devoted to the growing of cotton and rice, is
typical of Southern India.
A very hot walk to some small granite hills on the other
side of the lake produced little beside two males of Lvxias
marianne, and a solitary TZeracolus eucharis; the hills
seemed too hot, dry, and parched to harbour butterflies.
About the trees along the dam, or “ bandh,” were a few
Hypolimnas misippus, males, and abundance of Papilio
aristolochix,
In the cotton fields by the river Hypanis [Biblia] aithyra,
Dru., was to be got, but not plentifully.
The best collecting-ground was a very weedy nursery
garden and orchard. Here I one day had the advantage
of the assistance of my host, Mr. Edwin Scott, LC.S.,
whose keen appreciation of scents helped me greatly.
LIimnas chrysippus was abundant; of its scent Mr. Scott's
first impression was “some sort of dung,” then “a zoo” ;
later he said “ possibly like a cockroach, but more like a
musk-rat.” The scent is, I think, general, but is perhaps
stronger when the scent sacs on the hind-wings are opened :
a fact that I also observed at Calcutta.
Crastia core was common and gregarious, frequenting a
special mango-tree. When he smelt this insect Mr. Scott
at once cried out “acetylene,” adding that he would like
to put a lighted match to it to see whether it would burn!
Subsequent observations on the butterfly and the gas con-
vinced me of the accuracy of his comparison. The genital
organs appear to exude the scent, probably the long tufts
appended to them.
Observed in a tour through India and Ceylon. 109
At this place I confirmed in two specimens of Catopsilia
pyranthe the jasmine odour connected with the “scent
tufts” of the male, but did not find it as strong as in
specimens of the same species examined at Bankaptir six
weeks before. Mr. Scott agreed to the comparison with
jasmine, but thought the scent was perhaps even more
hke that of Polianthes tuberosa.
I also examined two males of Zirwmala limniace for
scent, but was unable to elicit any from the prominent sacs
on the under-side of the hind-wings, although I suspected
some to be emitted by the genital tufts.
Papilio erithonius was frequently met with, and P.
aristolochiz was common, but I only took a single P.
pammon. Although the male of Hypolimnas nvisippus was
fairly common, I only saw one worn female; this was of the
very marked variety inaria, Cramer, in which the white
marks near the apex of the fore-wing are entirely wanting,
and the black tip is reduced to a narrow border, so that it
closely mimics L. chrysippus, var. dorippus, Klug., a form
that is very rare in India. I several times saw the male
H. misippus reconnoitring L. chrysippus as if in doubt as to
its identity !
Of Precis enone I took but one, of P. almana two, but
P. lemonias was common. Of the following species I took
mostly single examples :—Limnas genutia; Hrgolis ariadne ;
Neptis varmona ; Polyommatus beticus ; Lampides celeno, Cr.,
form conferanda, Butl.; Catochrysops hapalina, Butl., two;
C. strabo, Fab.; Zizera otis, var. indica, Murray, two; and
the Skipper, Swastus gremius, Fab.
Of Melanitis ismene I took but a small fraction, for one
seldom sees a butterfly so battered, yet even this fraction
was found in the shade. In marked contrast are the habits
of Telchinia violx, since it haunts the most sun-scorched
places; it was not uncommon at Anantapur, but if
gregarious, as elsewhere, then I did not hit upon its
head-quarters.
I took one Terias libythea, and saw several 7. hecabe,
though it was but moderately common.
Hovering at flowers I two or three times saw, and once
caught, Cephanodes hylas, L., an insect very like Sesia
bombyliformis, Esp. There were also flymg in the sun
Deiopeia pulchella and Trigonodes hyppasia, Cr., a Noctua
very like Hydrelia unca, L, which reminded me of
Headington and old Oxford days,
110 Dr. G. B. Longstaff’s Notes on the Butterflies
Out of the grass I kicked up Zephrina catalaunaria,
Guen.,a pretty little Macariid Geometer Semiothisa subalbi-
tarta, Swinhoe, and Sterrha paullula, Swinhoe. The
common dragon-fly, Orthetrwm sabina, Dru., and a bug,
Lysarcocoris guttigera, Thunb., completed the tenants of the
garden.
A number of things came to light, viz. :—Deiopeia
pulchella, the Ocneriad, Enome detersa, Walk., a Noctua,
Ericeia inangulata, Guen., a Pyrale, Schenobius binwnctifera,
Walk. and a tiny Quadrifid Noctua Raparna lactea,
Swinhoe, as well as two bugs, Acanthaspis apicata, Dist.,
and Dieuches wniguttatus, Thunb., the former apparently a
scarce insect since the national collection contains the type
only. There was in addition to these a small ochreous
narrow-winged Geometer to which I have not been able to
assign a name, and an ichneumon, Henicospilus, sp. In
fact one evening swarms of insects came to light, including
many mosquitos, but these appeared to be all Culez,
fortunately no Anopheles.
Bangalur, lat. 13° N., alt. 3100 ft.
February 23rd, 1904.
Thke change of trains at this large military station gave
me an hour or two’s collecting in the extensive public
gardens. There was rather a high wind which was against
a good day, but the afternoon proved interesting since it
gave me the first glimpse at the “ Ceylon” fauna. Here I
saw for the first time that very striking black, white, and
orange Lyczenid, Zalicada nyseus, Guer., as well as the
huge and magnificent Papilio polymnestor, Cram. [parinda,
Moore], a truly gorgeous monster in which a pale lilac is
the prevailing colour trimmed with black.
The only other things noted were more ordinary, to wit
Catopsilia pomona, a female, Crastia core, several Telchinia
viole, Nychitona xiphia, an abundance of Neptis varmona,
and one Neptis gumba, Moore.
The Nilgiris, lat. 11° N.
February 24th—March 8rd, 1904.
The Nilgiris, or Blue Mountains, rising abruptly from
the plain, itself nearly 2000 ft. above the sea, form a
Observed in a tour through India and Ceylon. 111
rolling table-land with an average altitude of from 6500 ft.
to 7500 ft. This plateau consists for the most part of
grassy downs with here and there “sholas,” or thickets of
mixed growth, very beautiful at this time of the year
owing to the red colour of the young leaves of the pre-
ponderant tree. Unfortunately, alike for the entomologist
and the artist, these ‘“sholas”’ have been largely cut down
to make way for the extensive Government plantations
of eucalyptus, which are by comparison dreary and
monotonous.
On the way up the cog-wheel railway I saw on the side
of the cutting two beautiful blue-green Papilios, which
may have been either P. telephus, Feld., or P. teredon,
Feld. At about 4500 ft. I netted a Neptis varmona from
the train in motion.
It was evidently too early in the year to get many
butterflies at Utakamand, the elevation making the nights
cool, so it was necessary to seek out sheltered flowery banks
facing south, or preferably south-east. In two such spots
within a very circumscribed area Zalicada nyseus was
common ; a single example also occurred [along with the
inevitable Pyrameis carduc] onthe grassy top of an isolated
and exposed peak of about 8000 ft. This Lyczenid is quite
typical of “South India and Ceylon”; it is a conspicuous
insect on the wing, its tricolour of black, white and orange-
red, which should delight German entomologists, making
it look larger than it really is.
Terias hecabe was rather common, but worn. A female
Lycena betica and several Pyrameis indica were also old
friends, and the same applies to two or three Papilio
aristolochixy seen at flowers in the hotel garden, the latter a
good deal the worse for wear.
A few Yphthima chenut, Guér., occurred at about 7800 ft.,
the only Satyrid I met with at Utakamand. Ganoris canidia
flew up to 8000 ft.; a male had a distinct smell like that
of our Pieris rape. “I submitted ‘the living butterfly to
my daughter and her lady friend, who both noticed the
scent, though unable to describe it. When mignonette
was suggested for comparison they both said “No”; but
when sweet-briar was mentioned they said it was like
that, my daughter speaking the more confidently of the
two.
At about 7400 ft. I took a female Catophaga paulina, and
also a fine female of Hiposcritia narendra [Moore], quite a
112 Dr. G. B. Longstaff’s Notes on the Butterflies
Ceylon species. The specimen is labelled “flies fast :
rather common from 7400 ft. to 8400 ft.” It is but too
evident that I had not recognized that I was catching
anything out of the common, and it is more than probable
that I confounded the females of Catophaga and Tachyris
with Hiposcritia, so that I am not by any means disposed
to trust the statement that H. narendra was common then
and there. One necessary consequence of my complete
ignorance of the Indian fauna was that I did not know
what was most worth catching. These white butterflies
are as a rule by no means easy to catch and were often in
bad condition, but in each locality I used to endeavour to.
secure one or two good samples. Doubtless the Hiposeritia
passed for a very fine Zachyris.
Amongst herbage Mecyna polygonalis, Hb., was often
kicked up, having much the habits and appearance of my
old Bermuda and Mortehoe friend Stenopteryx hybridalis,
Hubn. (Nomophora noctuella, Schiff.), which too was fairly
common in exposed situations at about 8000 ft.; at a
similar elevation a single example of the Danaid Badacara
nilgiriensis, Moore, was taken.
Above the Botanic Garden on the road to Dodabetta, at
about 8000 ft., I several times saw, but missed, Vanessa
charonia, Dru., a butterfly that looks dingy in the cabinet,
but on the wing looks much brighter and bluer than would
be expected. It is sometimes called the Blue Admiral
(completing the trio), though in truth it is much more
like a tortoise-shell. Argynnis niphe was common at the
higher elevations, and in exposed situations up to 8500
ft., reminding one of A. aglaia, L. It flew up and down
the roads, returning again and again to the same spot.
_The commonest and most characteristic butterfly of
Utakamand was the pretty little Colias nilgiriensis, Feld.,
which was seen coursing over the grassy downs from 7300
—8600 ft. Its flight is moderately fast, but quite close
to the ground. It was somewhat startling, but in a way
refreshing, to come across this Arctic survival so far within
the tropics, associated moreover with species characteristic
of Ceylon. As it was especially abundant in the hotel
garden I took the opportunity of examining five males for
scent; in two cases I suspected the existence of a slight
scent, but_in the remaining three the result was negative.
From Utakamand I moved to Konur, which stands on
the southern edge of the plateau, overlooking the plain,
Observed in a tour through India and Ceylon. 118
It is at about 6500 ft. above sea-level, or 1000 ft. lower
than Utakamand. At this elevation Colias nilgiriensis was
not nearly so common as at the higher levels,
Worn Pyrameis indica, a few Neptis varmona and Precis
iphita turned up here and there. Of a pair of Terias
hecabe taken in copuld, the male proved to be of “inter-
mediate dry,” the female of “pronounced dry” type.
Opportunities of noting the pairing of the several forms
occurred very rarely. Here I secured one specimen of
Yphthima ceylonica, Hew., another foretaste of the great
southern island. Y. inica, Hew., turned up at about
5800 ft., but at about 6500 ft. in a clearing in a wood, I
found Y. hiibneri, Moore, together with Y. chenwi, Guér.,
and Y. philomela, Joh. There were swarms of these
Yphthimas on that sunny bank, but as I did not distinguish
the species at the time I cannot now say of what the bulk
of them consisted. Some of the specimens have injuries
to the wings, which from their shapes may have been
inflicted by birds, but I attach little importance to them,
especially as the injuries are unilateral, since the wings
of Yphthima (and to a somewhat less degree of Mycalesis
also) are so fragile that quite unbroken specimens are
exceptional.
The genus Papilio was represented by a couple of
ertthonius. As usual Argynnis niphe showed a preference
for lofty and bare places. On one occasion I watched a
female of this species for some time under the impression
that it was Limnas chrysippus ! The resemblance on the
wing is greater than might be supposed. Vanessa charonia,
Dru., which had before eluded me so often, fell a victim
at last; I secured two specimens on a shady road through
a wood. It settles on rocks or walls, a habit that makes
it hard to net, moreover it is shy and easily disturbed,
though usually coming back again to its resting-place.
Stenopteryx hybridalis was common in grassy places, and
I took the Boarmid Bilactis inceptaria, Walk., flying in the
hotel garden at dusk.
It was tantalizing to be told by the hotel manager at
Kontr of the immense number and variety of butterflies
there in the summer. I was, however, fortunate in
making the acquaintance of a dealer, named Solomon, a
coloured man, who told me that at that time of the year
it was no good collecting on the high ground, but for
a consideration he agreed to show me a very good place
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 8
114 Dr. G. B. Longstaft’s Notes on the Butterflies
near the foot of the hills. Accordingly I went with him
on March 2nd, and again alone on the following day.
This involved travelling by an early goods-train to Kallar,
the first station on the mountain railway above Mettu-
pilaiyam, about 2000 ft. above the sea, but only 200 ft. to
500 ft. above the plain. Here, as in other parts of India,
the best places for insects, at any rate in the winter season,
are to be found in the belt of jungle at the foot of the
hills, or in the woods on their lower slopes. But it is just
in these places where the dreaded Anopheles is as abund-
ant as the Rhopalocera, and the station-master at Kallar
told me that entomologists always slept at Kontr and
went up and down by train to avoid the nocturnal
terrors of the deadly malaria—the tiny, innocent-looking
Anopheles !
The collecting-ground was various, and included, besides
bushy jungle with plenty of flowers near the station, large
irrigated banana and betel-nut plantations as well as the
bed of the river with its bordering woods.
The first thing to catch the eye was Papilio hector, L.,
and very magnificent he looked fluttering at the flowers of
Lantana in his crimson-and-black suit set off with white.
This is indeed one of the most striking butterflies that I
met with in my travels, with its wings expanding four
inches and upwards. It proved to be distinctly common,
but one does not get within reach of every Papilio that
one sees, nor indeed does one succeed in netting all that
are struck at. PP. hector was accompanied by plenty of
P. pammon and a few P. aristolochie. One of the P. hector
brought home is remarkable for the fact that the whole of
the tips and half the hind-margins of both hind-wings
have apparently been bitten off, almost absolutely symmet-
rically, by some foe. If the red spots on the under-side
be really “ warning marks” this is the more noteworthy.
A boggy, but sunny, corner of an irrigated banana-
garden produced single specimens of the fine Skippers
Tagiades atticus, Fab. [? = TZ. menaka, Moore] and
Tagiades distans, Moore.
This same garden and the adjoining plantations of
betel-palm (Areca catechw) yielded a few Melanitis ismene,
a fair number of Jycalesis perseus, Fab., as well as
Yphthina marshalli and Y. philomela, Joh. [= baldus,
Fab.]; there was also abundance of the pretty and very
distinct Vphthima ceylonica, Hew., with its silvery-white
Observed in a tour through India and Ceylon. 115
hind-wings, which tried, not without occasional success,
to pass itself off as a Blue. A few hasty observations on
this species when at rest failed to detect any such “ list ”
as is common in many members of the family.
I sent home two specimens of Mycalesis perseus, one an
ordinary example of the dry-season form in which the
ocelli are indicated by faint dark dots, the other (unfortun-
ately very tattered) in which the full complement of
ocelli on the under-surface is indicated by conspicuous
chalky-white spots of varying sizes, to wit, two large and
one small on the primaries and two large and five small
on the secondaries. Three of these spots are faintly
visible on the upper-surface. There are no rings and no
pupils to the spots. It would appear to be a unique
aberration of the dry-season form. In the shade along
with the above Satyrids was the inevitable NMychitona
xipia and a solitary male Hlymnias undularis.
A weedy neglected field near the river yielded besides
Limnas chrysippus and Atella phalanta plenty of the
“orange-tip” Jaias marianne, as well as a smaller number
of the more gaudy Jxias pyrene. One proved to be a worn
specimen of the female lacking the orange tip, a distinct
and well-marked variety ; another was of the racial form
cingalensis, Moore. The “whites” Huphina nerissa and
Catophaga paulina were in plenty. A single Catopsilia
pomona was netted, a somewhat papery-looking insect,
especially on the under-side, also several C. pyranthe of the
“transitional Gnoma” form. In one of the latter (a male)
I detected a faint scent, but less like that of jasmine than
in the Catopsilias examined at Anantdpur. In the same
field Telchinia viole was abundant, while Hrgolis ariadne
was, as usual, common among Lzcinus.
But all this time Solomon was most anxious to get me
down to the river. This is a rapidly-flowing stream,
occupying perhaps half its bed, and having on either bank
sloping woods of mixed growth. Solomon sought out a
place where a tiny tributary emerging from a rushy swamp
trickled over the damp sand. He forthwith stuck into
the wet sand a foot or so from the rill and well clear of the
herbage, three or four large butterflies of which he had
netted worn or broken specimens; then he stood by to
watch. Nothing much happened, for unfortunately clouds
had come up and the afternoon was only partly sunny,
whereas to get many things at water, whether decoys be
116 Dr. G. B. Longstaff’s Notes on the Butter flics
used or not, it needs, as Solomon put it, to be “ plenty hot.”
It was indeed hot enough for most Europeans, but not up
to the exacting butterfly standard. However, next day
the conditions were more favourable, and I found near
what was left of Solomon’s decoys a number of ‘ whites”
and “ orange-tips.” Accordingly I put down a few more
decoys and walked away. After spending some time in
vain endeavours to catch the conspicuous Hebomota glau-
cippe, L.—giant of orange-tips—which was careering wildly
about in all directions, I returned to the decoy-place and
sat down just within the reach of my six-feet net-stick,
Catophaga paulina were there in abundance, but all males,
mostly sitting quite close together, almost touching, with
wings erect so that the “hook-tip” of the fore-wing was
very conspicuous; in another cluster close by were from
six to eight Jaias marianne.
It will perhaps give some idea of the numbers when I.
say that I quite easily netted five C, paulina in one swoop,
and seven in another.
Then Hebomoia glaucippe came along, reconnoitred the
position with great circumspection, and settled warily for a
second or two, but darted swiftly off at the least movement
on my part. Nevertheless, with care and patience, I
managed to secure a couple of specimens, My old friend
Papilio erithonius came next and soon settled down a short
distance away from the “whites,” he was shortly followed
by another and yet another: they all settled close together,
within a hand’s-breadth, forming an exclusive community
and continued to drink steadily. All at once a blue-green
flash, and Papilio telephus, Feld., sailed close past me ; again
and again he came, and finally, looking askance at the
vulgar assemblage of “ whites” and “ orange-tips,” settled
quite close to the P. erithonius, evidently preferring their
more select company. This occurred several times. LP.
telephus, when settled with wings erect, displayed an
unexpected beauty, for, in place of the ebony and emeralds
on the upper-surface, it shows beneath nothing but sheeny
mother-of-pearl picked out with tiny rubies. By patient
watching and judicious swooping I secured three specimens,
and, be it remembered, these were all I saw that day. So
much for water; what share the decoys had in my success
it is hard to say, but Mr. E. E. Green, of Peradeniya, told
me that decoys were efficacious, and Mr. Denton, of Regent
Street, says that he has used even paper decoys with success.
Observed in a tour through India and Ceylon. 117
The congregation of butterflies at damp sand was
observed by Bates on the banks of the Amazon in 1849.
He noted that they were all males, mostly of the genus
Callidryas.* Indeed Sir J. D. Hooker had the year before
noted butterflies sitting on damp sand “in thousands” in
the Ranjit valley, Sikkim.+
Mr. E. André noted a similar thing in Venezuela in
1897, where the attraction was the foul mud of a farm-
yard: the butterflies were chiefly Callidryas, with some
Heliconius, Papilio, Metamorpha and Cwrois. He says:
“Each species tried to herd with its own kind,” but he says
nothing as to sexes. There is a capital photograph in his
book of a group of Callidryas.t
Doubtless this habit of butterflies is well known to all
tropical collectors. I myself in Germany some 385 years
ago, noted swarms of Blues at small puddles in the road
several species together, including, so far as I could see, L.
alsus, L. arion, and L. betiea. A few days afterwards near
the same place and similarly occupied I caught Apatura
aris in my hat! This summer at Mortehoe, on the Ist of
August, in the early afternoon, I saw 14 or 15 G. napr
sitting close together on wet mud; they were all males.
A piece of waste ground adjoining the plantation of the
singularly graceful Areca palms, covered with Lantana in
full bloom, was crowded with butterflies such as Crastia core
and Narmada coreoides, Moore, one or both of which (for I
did not distinguish them when alive) was abundant;
several Neptis varmona, and two or three Nepheronia
ceylonica, Feld., another southern species. More striking
than all these were the swarms of Zirwmala limniace, a
big and handsome black and bluish-white Danaid, which I
found all over India but never saw elsewhere in anything
like such numbers as on that mass of Lantana.
Other things that turned up in the course of the two
days’ collecting were Tachyris hippo, two; Teracolus etrida,
one; Hypolimnas bolina, two males ; H. misippus, one male ;
Precis iphita, common; Caprona ransonnetiu, Feld., one;
Parnara mathias, one; Castalius rosimon, and plenty of
Lampides celeno, Cr., including the form conferanda, Butt.
Of the above the fine Skipper, Caprona, was seen to
settle, in full’sunshine, on the under-side of a leaf, with its
* “Naturalist on the Amazons,” Ist edn., p. 249.
t “Himalayan Journals.”
t “Naturalist in the Guianas,” p. 142.
118 Dr. G. B. Longstafi’s Notes on the Butterflies
wings fully expanded like a Geometer. I do not ever
remember seeing a butterfly do this before, but then we
are perhaps wrong in calling Skippers butterflies. One
of the Lampides, a female (?), was found settled close to
the ground, with all its wimgs erect as usual and close
together; it was however moving its hind-wings alternately,
in a rhythmical manner, in the plane of the wing, about
10-15° forward and then back. No other specimen of the
species was near it.
On the occasion of my first visit to Kallar, as we were
walking back to the station, Solomon suddenly darted off
like the wind, and I found that he was after a very large
Papilio which he had caught sight of flying about a puddle
in the road, some hundred and fifty yards off. He waited
long and patiently until it settled to drink and then popped
his net over it. It was Papilio polymnestor in splendid
condition, black and French grey, 5$ inches in expanse!
This haughty beauty was not kind to me at Kallar; many
a time I caught a glimpse of her flying about in a super-
cilious sort of way, but she never gave me a chance of
closer acquaintance. Solomon had the advantage of me in
many ways, first and foremost in years, next in his keen
sight, but he was also wily and skilful with his net.
During the day he took among other things a specimen of
Papilio agamemnon, L., a fine black-and-green fellow that i
too had seen; also one of that grand diamond-beetle green
butterfly Papilio erino, F., which I missed the next day at
Lantana flowers, as I believe, through sheer excitement !
Trichindpali, lat. 10° 50’ N., alt. c. 400 ft. or less.
March 4th and 5th, 1904.
My collecting here was almost confined to the banks of
an irrigation canal, where the genus Papilio was repre-
sented by P. hector, P. pammon, and P. aristolochix, of each
of which I saw several.
Limnas chrysippus was common; in the male of Zirwmala
limniace I detected a very faint scent, suggesting old cigar-
boxes.
Catopsilia pyranthe was rather common, the specimen
preserved was of the intermediate form; I noted a scent
in the male, but it was not so strong as in some of the
Bankapur specimens. Of WDelias eucharis I took two
Observed in a tour through India and Ceylon. 119
females, by far the less common sex, at all events in
collections. Of Huphina nerissa I took one of each sex.
Terias hecabe was abundant.
The most striking fact about the butterflies of Trichin-
apali was the predominance of the genus TZeracolus; of
these I met with three species: 7. ewcharis was in abund-
ance, but I find my specimens comprise eight males to
two females; of 7’. ctrida I took two males; and I was
greatly delighted to see here for the first time that truly
exquisite little gem the crimson-tipped 7. dande. It
proved to be rather common, and I secured two of each
sex. It is one of the most “elegant flies” that I have
ever seen alive.
Precis orithyia was common, the specimens small and
brilliant ; P. /emonias was in larger numbers than I met
with anywhere else, in fact quite abundant; Zrgolis
ariadne was common; I netted a pair of Hypanis ilithyia
in copuld, one was of the “wet-season” form, the other
“intermediate” tending to “wet.” As usual Telchinia
viole was common.
I took here one specimen of that beautiful Lyceenid
with the under-side striped like a tiger, Spindasis vuleanus.
This is one of the butterflies with an anal lobe to the
secondaries, but unfortunately I had not an opportunity of
observing it at rest. Lampides celeno, Cr., was common ;
some smaller and dingy Blues were abundant, Zizera otis,
Fab., var. indica, Murray, and Chilades varwnana, Moore,
thought by De Nicéville to be the wet-season form of
C. laius. I also took one specimen of a small bright
golden Skipper, Ampitiia mavo, Fab.
Tanjur, lat. 10° 47’ N., alt. 350 ft. or less.
March 6th, 1904.
The predominant genus of the plains of Southern
Madras would appear to be TYeracolus, which was repre-
sented in my envelopes from Tanjtr by a male 7’. etrida,
a pair of 7’. ewcharis, and five males and two females of
my favourite crimson-tip, 7. dande, which was quite
common.
Of Catopsilia pyranthe I took a dwarf male of the
intermediate form. Zertas hecabe was common, and I took
a very large female [over 1:8 inches in expanse, it was
120 Dr. G. B. Longstaff’s Notes on the Butterflies
“dry”]. Single specimens of the following were sent
home: Nychitona xiphia; Papilio pammon, male; Limnas
chrysippus, female ; Castalius rosimon and Lamprdes celeno,
Cr., of the form conferanda. Telchinia viole was common,
one being of a fine red colour.
Médura, lat. 9° 55' N., alt. 600 ft.
March 7th, 1904.
This was about the least productive place that I visited.
Limnas chrysippus was scarcely common. A male Huphina
nerissa gave out the sweet-briar scent quite strongly. I
saw several T'elchinia viole upon a railway bank. Precis
anone was fairly common, but P. almana was commoner
here, about the irrigation ditches bordering meadows, than
at any place I visited; they were of the “intermediate
dry” form. PP. lemonias was also abundant, some of them
being very brightly coloured.
In a grove of young palms near the river a singular
dragon-fly, Libellula variegata, Linn., was common; the
tips of its wings are transparent and colourless, but the
basal three-fifths of the primaries, and the basal five-
sixths of the secondaries, are light-brown with a bold
dark-brown pattern. I believe that I saw the same crea-
ture in the Kudsia Gardens at Delhi, flying near the tops
of trees, and then, as in the present case, took it for a
Helwonius-like butterfly, which it greatly resembles on the
wing. As I did not know that any butterfly of that shape
was found in India I was greatly excited at seeing it, and
proportionately disappointed when I at last effected its
capture. |
This was the last place at which I collected in India.
Ceylon, lat. 7° N.
All the places that I visited in this beautiful island were
within twenty miles north or south of the seventh parallel
of latitude. The luxuriance of the vegetation was an
immense relief after the parched plains of India. At the
lower elevations it was more distinctly tropical than any
that I had yet seen, but this character was lost at greater
altitudes.
Observed in a tour through India and Ceylon. 121
Peradeniya, alt. c. 1200 ft.
March 10th, 1904.
These justly celebrated gardens lie about four miles
south of Kandy near the centre of the island. Their
situation is beautiful and all the familiar “ hot-house
plants” grow luxuriantly. Splendid palms of many kinds,
huge bamboos almost as tall, Dracewnas, Crotons, Acalyphas,
Marantas, nutmeg, cinnamon, camphor, huge trees of
Ficus elastica with roots spreading far over the surface, etc.,
etc. Grass has been largely ousted by the sensitive-plant,
Mimosa pudica, which, introduced from South America,
has run wild. It grows about a foot and half high, and
when one walks through it a broad path is left owing to
the collapse of the leaves.
Here, well out of reach, I saw my first Ornithoptera,
truly it is well named ; I missed a second specimen through
sheer excitement. Several Catopsilia pomona were netted,
one was a female verging on the catila form, the others
were typical males, one of which had a slight scent. Of
three male Zerias hecabe two were wet-season, the other
of “intermediate wet” form. A male 7’. libythea was also
of wet-season type, a female was also taken. The only
Nymphalids noted were two Neptis varmona and several
Precis iphita. Here also I took my first Parantica
ceylonica, Feld., a Danaid found in abundance later.
Of Mycalesis mandata, Moore, I only saw one, but the
pretty little Yphthima ceylonica was swarming amongst
the sensitive-plants. Mr. E. E. Green, the entomologist
to the Ceylon Government, suggested that its colouring
might be indirectly protective, since on the wing it looks
much smaller than it is, only the white posterior two-
thirds of the hind-wings being conspicuous, and these the
least vitally important to the insect. The only Blue seen
was Zizera karsandra, Moore.
A second visit to the gardens, rather late one afternoon,
produced no insects, but gave me my only sight of a wild
cobra, about 24 feet long, with a very large “hood”; it
crawled quickly away into the roots of a “ travellers’ palm.”
Unfortunately for me Mr. Green was on the point of
going to England on leave, but though busy with his
preparations for departure, he was good enough to show
me several very interesting things, such as larve of the
122 Dr. G. B. Longstaft’s Notes on the Butterflies
leaf insect, young snakes, etc., and above all he gave me
some very useful advice. Peradeniya, he said, was not as
rich a locality as Kandy; and, as regarded the highlands
of Ceylon, he told me that, at any rate at that time of the
year, Lepidoptera were for the most part confined to
certain favoured spots, which it was unlikely that 1 should
hit upon. He therefore strongly recommended me to
concentrate my attention on “Lady Horton’s Drive” at
Kandy.
Kandy, alt. 1500 ft.
March 11th—15th, 1904.
On the south side of the artificial lake at Kandy stand
some low hills, covered for the most part with natural
forests, through which have been cut a number of roads
named after the wives of former governors. Lady Horton's
Drive is one of these, which runs about half-way up the
hill, winding around its southern and eastern slopes. A
wide road, bounded on either side with forest of rich
and varied tropical growth, lying fully open to the morn-
ing sun, commanding moreover a glorious view over
groves of palms to the bluest of distant hills, it affords an
almost ideal collecting-ground. The climate of Kandy,
so far as I experienced it, is delightful, tropical heat
tempered by elevation, and with a pleasant softness in the
air, yet free from the excessive damp of many places within
the tropics. Its vegetation is by far the richest that I had
seen. My pleasure in collecting in this earthly paradise
was greatly enhanced by the companionship of Mr. W. G.
Freedley, junr.,a Philadelphia gentleman who had been
collecting butterflies in Borneo, Celebes, Japan, Macao, etc.
In such a locality it was perhaps to be expected that
Pierines would not be dominant, at any rate so it was.
By far the commonest of the family was Catopsilia pomona,
ot which the males were very abundant, but strong fliers
and by no means easy to catch. We remarked that they
usually all flew in the same direction, and that uphill.
As the females were comparatively scarce one was not
surprised to see more than once signs of jealousy on the
part of the males. I detected a slight jasmine-like scent
in the male on stroking the “scent tufts” on the hind-
wings. A female Terias hecabe had apparently been
bitten in both hind-wings when at rest, the injuries being
Observed in a tour through India and Ceylon. 128
more or less symmetrical. Delias ewcharis was scarce,
Catophaga paulina more common. In one case I saw a
bird try to catch a specimen of the latter on the wing ;
the bird missed its quarry, but I was more successful ; it
proved to be a male.
The Danaids were well to the front, the commonest
species being Parantica ceylonica ; it is smaller and greyer
than Tirumala limniace and varies considerably in size, a
small male measuring only 2°5 in., a large female as much
as: 3-4 in. across the wings. It was curious that this
species became quite abundant late in the afternoons, as
other things were retiring. I was surprised to find that a
male when fluttering in the net gave out a strong scent
like that of Crastia core, i.e. very like acetylene. This
was noted in two or three specimens, and was quite un-
mistakable. Danais septentrionis, Butl., appeared to be
rather common, A female has the hind-wings much
broken, perhaps from the bite of a lizard, but the breakage
is only in part symmetrical. Limnas genutia, of which I
took a very small one, was very scarce, and I did not see
L. chrysippus at all. The genus Crastia was represented
by many individuals. I took five C. asela, Moore ; of one
of them I noted at the time, “has a scent as in core.”
But the most prominent group of butterflies at Kandy
was assuredly the Papilzonidx ; | met with six species. The
most remarkable was Ornithoptera darsius, Gray, peculiar,
I believe, to Ceylon, an insect that I had greatly wanted
to take ; it appears to be fairly common, as I saw two at
Peradeniya, seven or eight at Kandy, and two at Haragdma.
It sails about somewhat slowly and in a dignified manner,
looking very distinguished in its rich yellow-and-black
livery and impressive by its size, five and a half to six inches
expanse of wings! When it comes within reach it is not
hard to catch, and I secured two males and a female, but it
is a formidable-looking creature in the net, with a thorax
suggestive of a Bombyx. Mr. Freedley told me that the
males have a scent like sassafras, but I learned this too
late for confirmation. The male Papilio pammon was
common enough, one specimen was unusually small, mea-
suring under three inches. ‘wo specimens of P. aristo-
lochie (a distasteful butterfly) were brought home; one of
them has the tips of the hind-wings up to the tails bitten
off quite symmetrically, thus much resembling the muti-
lated specimen of P. hector taken at Kallar. Of the tailless
124 Dr. G. B. Longstatt’s Notes on the Butterflies
P. dissimilis, L., 1 took three, but probably saw more, since
it so very closely mimics Zirwmala linuniace or a large
Parantica ceylonica, as easily to pass for one of those
insects; it is indeed most easily distinguished from them
by its habit of fluttering while feeding on a flower. One
of my specimens has the anal angle and a great portion
of both hind-wings bitten off in an almost symmetrical
manner, suggesting the bite of a lizard. It should be
noted, as was observed long ago by the President, that,
whatever the cause may be, it is in the great majority
of cases the hind-wings that suffer these injuries; doubt-
less the framework of the fore-wings is the stronger,
but that does not seem to be a sutticient explanation,
since from their greater length they must be more exposed
to chance injuries from thorns and the like. Mr. Freedley
took a Papilio that mimicked Huplea, but I believe that
P. dassimilis is dimorphic, one form mimicking each genus.
Indeed it would appear that the name dissimilis implies
that its bearer is like anything rather than a Papilio.
In a shaded glen down which a tiny stream and a foot-
path strove for the possession of the ground, I took close
to the water a faded specimen of my Kallar acquaintance
Papilio telephus, and missed another that was drinking at the
mud. But far more exciting than all the before-mentioned
species was Papilio polymnestor, or as Moore has it, P.
parinda, a truly magnificent fly that dashed about in all
directions. It measures about 54 inches across the wings
and is rendered most conspicuous by its colouring—French-
grey and black. It rarely settled and was very hard to
catch; Mr. Freedley and I were constantly striking at
it, but it almost always eluded us. After many fruitless
attempts I succeeded in netting two, one so battered that
its powers of flight were seriously impaired; Mr. Freedley
was even less fortunate, probably because he had a very
small net.
There was yet another Papilio which eluded me alto-
gether. It was black-and-green and I feel pretty sure
P. agamemnon [which I also missed at Kallar in the
Nilgiris]. It had the extraordinary and most aggravating
habit of flying up and down, or rather backwards and
forwards, just like a sentry, over some small trees below
the road. Its path, if one may so call it, was about a dozen
yards in length, and it always turned round at the same
place, moving by a succession of jerks. I once actually
Observed in a tour through India and Ceylon. 125
watched it for twenty minutes so occupied, it then settled
for a moment on a Lantana flower; I struck at it and
missed, and the performance began again | Another day
T saw it at its post as before. Of all the Papilios that I
saw this species was by far the wariest.
In striking contrast to the Papilios in every way are
the Satyrids. A single Calysime [Mycalesis] perseus
[? = medus, Fab.] was a very dingy shade-lover. The bright
little Yj “phthima ceylonica was abundant; so far as obser ved
it sits upright. Nyssanga patnia, Moore, a very distinct
species, with leaden metallic lines on the under- surface,
was fairly common at the edges of woods, but I did not
meet with it on the “patnas” or grassy plains of the
highlands of Ceylon.
The Nymphalines met with included several interesting
species, notably Cynthia asela, Moore, of which I saw a
very fine example, but caught only a very tattered fellow.
It seems to like sailing about over the trees. With
Cethosia wietnert, Feld., Thad similar ill-luck. Of Cupha
placida, Moore, again I have but a very worn specimen
annotated thus: “ Has the swift flight and to some extent
the habits of Precis, but is fond of resting on the leaves
of trees.” These remarks are probably intended for, or
at least include, the allied Cirrochroa cognata, Moore,
which was certainly common, though very local; one of
my five specimens, otherwise in good condition, has two
snips taken out of each hind-wing, symmetrically, but it
appears to be an insect readily chipped. Both these
species have fulvous wings with black tips, so they are
readily confounded in the field.
Neptis varmona might be said to be abundant, while its
ally, the brown-and-black Rahinda sinuata, Moore, was
decidedly common. In this connection may be mentioned
the Erycinid Libythea rama, Moore, which appears to
mimic Rahinda. I secured one specimen at Kandy, and
believe that I missed another in the Pashdk tea-garden
near Darjiling.
I saw no females of Hypolimnas bolina at Kandy, but
took three males, one of which had lost both the anal
angles of the hind-wings, the injury being in part sym-
metrical. Precis iphita. was common, so was P. atlites,
looking on the wing like a dingy Neptis a new brood
appeared on March 14th. Zy ‘golis mervone Was common.
The Lycznids were not well represented, but I saw
126 Dr. G. B. Longstaff’s Notes on the Butterflies
several Lovwra atymnus, Cr., var. arcuata, Moore; as usual
Lampides celeno was common, almost spangling in the
sunlight, one specimen was so unusually brilliant as to
recall LZ. adonis; Talicada nyseus brought up the rear.
The Skippers were represented by single examples of
Parnara mathias, a dark fulvous Jambria salsala, Moore,
and the dingy Spalgis epius, Westw.
At rest on a fence outside the “ Queen’s Bath” I found
one morning a fine Sphinx, Meganotum melanomera, Butl.
A very distinct- looking Arctiid having a crimson body and
light pink fore-wings, P with a longitudinal fuscous streak,
Creatonotus interrupta, Linn., came to light, as also did
Eupterote diffusa, Walk., a Lasiocampid.
A few insects of other orders forced themselves upon
me, busily occupied as one was, ¢.g. a very large, black 2
carpenter-bee, Xylocopa tenuiscapa, West., with peacock-
green wings ; a very large, evil-smelling, brown bug, covered
beneath with a waxy substance that during life “clistened
like silver, Zessaratoma javanica, Thunb. Another bug,
Chrysocoris stockerus, L., was an intense metallic green with
black spots; yet more conspicuous than any of these was
the large Fulgorid Hotinus maculatus, Oliv., or so-called
Lantern-fly, expanding three inches across the wings. Its
fore-wings are black-and-white, the hind-wings light blue
with a very broad black border. This was fairly common,
flying high and settling on tree-trunks out of reach, but
easily disturbed, when it flies off to a similar resting-
place.
Haragama, 11 miles 8.E. of Kandy.
March 12th, 1904.
This appears to be locally recognized as a great place
for butterflies; the collecting-ground is along the course
of a rapidly-flowing stream with wooded banks, perhaps
500 ft. below Kandy, or say, 1000 ft. above sea-level.
Again, I had the advantage of Mr. Freedley’s company
on the occasion of my expedition. ‘The first thing to
catch our attention was ‘Hebomoia glaucippe careering
about in considerable numbers, but most unwilling to be
caught.
The pretty little Zalicada nyseus was literally swarming.
I cannot remember ever having seen a Lyczenid in such
numbers. I repeatedly observed this butterfly settle with
Observed in a tour through India and Ceylon. 127
its head upwards and immediately turn round so that its
head looked downwards. This habit of resting with the
head down is common, if not general, in the family,
and has an obvious bearing on the protective use of tails,
anal lobes, and directive marks.
Zizera otis, Fab., was also in abundance, and Lampides
celeno, Cr., was common.
Tirumala septentriovis was not common, and the speci-
mens netted were tattered males; one had a symmetrical
injury near the anal angle of the hind-wings, possibly due
to the bite of a bird. I saw a few specimens of Crastia
asela, Moore, and secured two males, which exhibited the
“acetylene odour.”
Neptis varmona was common, and I took a specimen of
the elegant Nyctemerid day-flying moth 7rypheromera
nigrovenosa, Moore, which seems to mimic it. Of the
beautiful Nepheronia ceylonica | took one male. Cirrochroa
cognata, Moore, was noted flying about a particular tree
and did not appear to be attracted by flowers; I only saw
it in that one spot, and the two taken were in poor
condition. Itis very like Cupha placida, Moore [ = Messaras
erymanthis, Stgr.|, of which I took a fine specimen close by,
at wet sand. I again confounded the two species as at
Lady Horton’s Drive. A few Hrgolis ariadne and several
Nychitona xiphia were seen. A male Huphina nerissa
had the sweet-briar scent ; a female was in fine condition.
At the furthest point reached in our walk, by a little
bridge, two or three spots in the damp sand appeared to be
very attractive. Besides the Cupha already mentioned
there was [xias pyrene, var. cingalensis, and Papilio pammon,
the male, was rather common. [It was also seen flying
about bushes, but not at flowers.| My Kallar friend
Catophaga paulina was literally in crowds; they were all
apparently males, sitting in dense clusters, their pointed
white wings suggesting to me toy encampments. I easily
netted ten at one swoop, while Mr. Freedley by a more
cunning movement succeeded in getting as many as thirty-
four into his net! In the same place I saw six or seven
of the beautiful Papilio telephus, Feld., settled quite close
together, and managed to secure three. It is a black-and-
green species not easy to distinguish from P. jason, L.
The females of Catophaga paulina were common at
flowers. One of the males, by the way, had a symmetrical
injury to the ¢ips of the hind-wings, but I can hardly see
128 Dr. G. B. Longstaff’s Notes on the Butterflies
how it could have been inflicted by an enemy without
simultaneous injury to the fore-wings.
This day I saw two Ornithoptera darsius, one quite out
of reach, the other I missed badly.
Hatton, alt. 4200 ft.
March 16th—18th, 1904.
In going up-country from Kandy when near Ullapdne
station [alt. c. 2500 ft.] I caught, from the train, Nar-
mada montana, Feld., and a little further on, ec. 3000 ft.,
a male Catophaga paulina, a species that is very abundant
in the Ceylon highlands.
Before Hatton is reached the line enters the tea country,
whence the glorious primeval forests have disappeared,
having been ruthlessly and completely cleared out to make
way first for coffee and later for tea. Though doubtless
“orateful and comforting,” the tea-plant is most un-
picturesque, only slightly surpassing the potato in that
quality. The Grevilleas with their light feathery foliage,
planted in regular rows to slightly shelter the tea from
sun and wind, do but little to relieve its stiffness, and are
a miserable substitute for the departed woodland glories.
About Hatton there are but scraps of the forest left on the
tops of the highest hills, and we were told that the tea-
planters are constantly urging the Forest Department to
allow these to be improved away. It results that what once
was doubtless a grand entomological locality is now a very
poor one.
Here for the first time I examined Catophaga paulina for
scent, and was surprised to find that the three males tested
had a scent nearly as strong as that of P. napi; it was
described at the time as “like sweet-briar, but sweeter and
more luscious,” and I wrote to Dr. Dixey the same evening,
adding “I had no doubt whatever.”
About the hotel garden Argynnis niphe was common, a
male had the fore-wings notably shorter and broader than
usual.
A short walk in what is left of the old forest, towards
the top of a high hill, say at about 4500 ft. produced
several specimens of Lethe daretis, Hew., a regular sylvan
Satyrid, repeatedly settling on the path, apparently always
erect. Two of them have lost large portions of the hind-
Observed in a tour through India and Ceylon. 129
wings near the anal angle, one symmetrically and in a way
to suggest the bite of a lizard. On the under-surface of
this butterfly the unusually large light-coloured scales on
a black ground near the hind margin of the hind-wings
are striking. In the same scrap of forest I took a single
specimen of another species of the same genus, L. drypetes,
Hew. | =embolina, Butl.]; also settled on a leaf of a tree
far from the ground, as is usual with the genus, a Limenitis
calidasa, Moore. Several Atella phalanta, a few Nepiis
varmona and Terias hecabe were also seen, a female of the
last-named being of the “ completely wet” form.
A specimen of Cyaniris singalensis, Feld.,is very like our
argiolus. With some ditticulty I secured a specimen of the
Jarge black-and-white Skipper, Celenorrhinus spilothrys,
Feld. This is the second Skipper [the other being Caprona
ransonnettii, Feld., at Kallar] that I have seen settle on the
under-side of a leaf during full sunshine, the wings being
expanded like a Geometer’s. Another specimen was
settled on a rock with its wings expanded in like manner.
Tn the hotel I found a fine Burnet (Syntomid) Huchromia
polymena, Linn., at rest on a wall, it has slender black
wings bearing orange spots, the body is blue, ringed and
collared with scarlet; and in my bedroom took a beautiful
little Tortrix-lke Noctua, Metachrostis incondita, Butl.,
measuring only 17 mm. across the wings; also a most
formidable-looking long-waisted wasp, Humenes jpetiolata,
F., a 2, and Pomasia psylaria, Guén., a pretty little yellow
Geometer with metallic markings, evidently attracted by
light.
"When coming down from Adam’s Peak on March 18th,
at the height of about 6000 ft., I saw several of the
Lithosiid, Aswra wnifermis, Hmpsn., but in the rough
scramble of the descent could only secure one; at about
4800 ft. were several Zalicada nysews, and a few hundred
feet lower down I bottled two green beetles, somewhat
resembling our Rose-beetles, but much more shiny,
Coryphocera elegans, Fab.
Nuwéra Hiiya, alt. 6200 ft.
March 18th—21st, 1904.
This Sanitarium is in some respects like Utakamund, it
is situated on a grassy plain forming a basin among
TRANS. ENT. SOC. LOND. 1905,—PART I. (MAY) 9
130 Dr. G. B. Longstaff’s Notes on the Butterflies
mountains. The “patnas” or grassy areas are bounded
by woods, which in their turn are fringed by somewhat
stunted scarlet rhododendrons. At the best season it
doubtless affords excellent collecting, but I found Mr,
Green’s statement, that I should be unlikely to light upon
the good localities, amply confirmed.
I saw several Papilio teredon, Feld., flying about, and
secured two that were drinking at wet mud. A female
Terias hecabe proved to be of the wet-season form. Of
Neptis varmona I took two. Ina sedgy place surrounded
by wood, a small “patna,” I took the Skipper Baracus
vittatus, Feld., curiously enough the only butterfly that I
had taken in a swamp up to that date. The streaky
markings of the under-side, following the veins, appeared
when the insect was settled on sedge to be strongly pro-
tective. Of Talicada nyseus I saw several, the only other
Blue seen was the argiolus-like Cyaniris lanka, Moore,
much battered.
Among moths I found one of the yellow Geometer
Corymica specularia, Moore, at rest on a tree-trunk, and
one Acidaliid Jdxa costata, Moore. Also on Mt. Pederuta-
lagalla, at about 8000 ft., the Skipper Baracus vittatus
among sedgy grass and Abraxas so1 ‘dida, Hmpsn., flying at
dusk, nearly uniformly dark fuscous. This last is
presumably a scarce insect since the British Museum
possesses the type only.
Hakgdla, alt. 4800 ft.
On March 19th and 21st I visited the beautifully-
situated and well-kept Botanical Garden at _Hakgala,
some five miles south of Nuwara Eliya and at a con-
siderably lower elevation.
Along the road Catophaga paulina was swarming, males
with their sweet-briar-like scent appeared to largely
predominate. They flew rapidly and always in the
same direction, roughly speaking from south-east to
north-west. They frequently flew in strings, just as
if they were tied together, and reminded me strongly
of the strings of floating stars that are dropped by a
certain kind of rocket; I often saw three, four or five,
and once even seven, so following their leader’s every
movement.
At a turn of the road close by the garden there was a
i
Observed in a tour through India and Ceylon. 131
small patch of a tall, but small-flowered composite plant
(of the Thistle-head sub-order) ; this plant did not appear
to be common in the district, but it was especially attractive
to a black Danaid, which was quite abundant within the
limits of this plant’s distribution. Chittira fumata, Butl., is
very distinct and handsome on the wing, its flight is slow
and it is easy to catch, but like all Danaids it has a tough
integument and is very tenacious of life. The favoured
flower was so attractive to the butterfly that it would even
go into deep shade to visit it. Chittira fumata may be
said to be gregarious; it has the “acetylene” odour cf
Crastia core, but not so strong and with a difference. J
made no observation as to the relation of scent to sex in
this species, which, by the way, I believe I missed at
Hatton.
Near this same spot I took two Crastia asela, Moore, and
saw others. I also got one Danas septentrionis, Butl.
The inevitable Lycenids were worn Talicada nyseus ;
Jamides bochus, Cr., one; and Polyommatus beticus, which
was common.
It was interesting to watch the pretty little honey-birds
feeding at some tall spikes of flowers.
Horton Plains, alt. 7000 ft.
March 23rd, 1904.
This beautiful district gives one some idea of what
Ceylon was before the era of tea-planting. Situated
about 2500 ft. above the railway and approached by steep
zigzag paths through rather poor woods, are extensive
rolling plains of coarse grass, locally called “patnas” ;
these are surrounded by woods having a general temperate
zone character, but with here and there an epiphytal orchid
to remind one that the latitude is but 7° N. In the
swampier parts of the patnas the devastating work of
wild pigs was evident enough, while the paths through
woods, and unmistakable droppings, proved that wild
elephants had passed not many days before. It was how-
ever not the season for butterflies, the air being too
exhilarating for their luxurious ways.
About half-a-dozen Chittira fumata were seen at
elevations of 6000—7000 ft,, mostly at their favourite
132 Dr. G. B. Longstaff’s Notes on the Butterflies
composite. On the patnas and among sedges in the woods
were « few of the Skipper Baracus vittatus, they were not
easy to see. J was surprised to come across no other
butterflies on these patnas, which seemed the very
place for such a Fritillary as JZ, aurinia, Rott., or such a
Satyrid as C. pamphilus, L., or at least for a ‘Blue, but
no, even the eponymous Nyssanga patnia was not to be
found,
In the woods I took two specimens of the beautiful
Lethe darctis, Hew., and saw two or three others. They
frequented shady paths and flew but a short distance,
settling upon a trunk or branch, reminding me strongly
of P. xgeria in my own carden at Mortehoe. The
only Argynnis seen here (or indeed in Ceylon) was
A. niphe; it was rather common in open spots in woods,
the female looking on the wing very hke Zimnas
chrysippus; a specimen taken, a female, had the apices
of both hind-wings and the anal angle of both fore-wings
symmetrically bitten.
Of Terias hecabe I found a few in a wood, of the
intermediate dry form. Neptis varmona was not uncommon
in the woods, as usual flying in a ghostly manner, and
usually settling upon leaves of trees. In the same woods
Cyaniris lanka, Moore, was common, but it was astonishing
to see so few insects in such a locality.
Haputéle, alt. 4500 ft.
March 23rd, 1904.
At this beautifully-situated Rest-house, overlooking the
plain and the old Boer prisoners’ camp, a great many
moths came to light.
Owing to its large numbers the most prominent of these
visitors was the small Noctuid, Ploteia frontalis, Walk., an
extraordinarily variable species; another Noctuid was
Cosmophila wanthindyma, Boisd.; there were two Deltoids,
Olybama lentalis, Guen., and Rivula basalis, Hmpsn.; the
Lymantriad Dasychira inelusa, Walk., and the extremely
widely-distributed Plemyria fluviata, Hiibn:
The formidable-looking beetle, Yylotrupes gideon, L., was
an uninvited visitor to my bath-room.
On the same day an Acidaliid, Idea costat, Moore, flew
into my face in a tunnel near Ohiya station, alt. 5000 ft,
Observed in a tour through India and Ceylon. 133
Colombo, at sea level.
March 25th and 26th, 1904.
Following Mr. Green’s advice I went to the Museum and
was well rewarded, though too pressed for time to reap all
that I might have got by a more deliberate examination of
the local collection of butterflies.
My collecting-grounds at Colombo were the Victoria
Park, much exposed to the sea wind; the old Cinnamon
Garden, said to be much worked for insects by the Museum
“boys”; and the old Dutch Cemetery. None of these
were very promising or very productive.
However, I saw here for the first time in Ceylon Limnas
chrysippus; I also netted one Parantica ceylonica, and
missed what I feel certain was a Hestia, probably jasonia,
Westw., which is, I believe, common at Colombo.
Precis atlites was common in the Dutch Cemetery but
worn, so was P. almana, nearly all of the wet-season form,
P. asterie, L.; one specimen however was dwarfed and
another was of the “dry” form with the ocelli rudimentary.
P. almana would appear to be the more prevalent species
in Ceylon and Southern India, where it replaces P. orithyia,
so universal in the North.
I saw several Delias ewcharis in the Victoria Park, and
once more noticed their fondness for lofty flowering trees ;
those taken were males. In the same place I missed what
I think must have been: the catilla form of Catopsilia
pomona; its congener pyranthe was common and I took
two males. Once more JVelchinia viole was common, but
of Papilio aristolochie I have only one to record. Of
Yphthima ceylonica I took two.
Of the Blues there were several species; Nacaduba
noreia, Feld., was very abundant and decidedly gregarious,
it positively swarmed in Victoria Park, though good
specimens were scarce. Lveres argiades, Butl., var. parhasius,
Fab., and Zizera karsandia, Moore, were also both of them
abundant. I took also a single worn specimen of Castalius
rosimun, Fab.
English is more spoken in Ceylon than in most parts of
India, but the Cingalese appear more noted for fluency
than accuracy; the inner meaning of the following
apparently strange request of a lad is easy to fathom:
“Master, buy some butterflies, ready-made.” On getting
back to the hotel from an entomological expedition one of
134 Dr. G. B. Longstaff’s Notes on the Butterflies
the messengers came up to me and said: “ Missie told you
to told me they had gone in.”
Animmature locust, Truzxalis nasuta, L., taken in Victoria
Park, completes the list of my captures in Ceylon, an island
that I was truly sorry to leave and that will always occupy
a treasured place in my memory.
SUMMARY OF BIONOMIC OBSERVATIONS.
Injurves by enemies.
Specimens of the following twenty-eight species were
taken which appeared to present injuries caused by the
bites of birds or lizards; save in the two cases specified the
hind-wings had borne the brunt of the attack.
Tirumala septentrionis, two.
Cirrochroa cognata.
Precis lemonias, two.
Precis almana.
Hypolimnas bolina, &.
Hypolimnas misippus, g.
Pyrameis cardui (fore-wings).
Vanessa hashmirensis.
Argynnis niphe, §.
Yphthima hiibnere.
Lethe daretis, two.
Curetis thetys.
Llerda epicles.
Pratapa deva,
Polyommatus beticus.
Colias fieldit.
Catopsilia pyranthe.
Ixias pyrene.
Tervas hecabe, two.
Terias leta.
Catophaga paulina.
Ganorts canidia.
Belenois mesentina.
Teracolus etrida (fore-wings).
Papilio hector
Papilro aristolochix
Papilio pammon.
Papilro dissimilis.
\ red marks on hind-wings attacked.
Observed in a tour through India and Ceylon. 1385
' It will be noted that this list includes no Zimnas, but
does include two Tirumala septentrionis, and two Papilios
with conspicuous red “ warning marks.”
In Ceylon a bird was seen to make a swoop at a male
Catophaga paulina, but missed it. I may here add that at
Yokohama, May 19th, 1904, I saw a dragon-fly of moderate
size, Orthetrum japonicum, Uhler, carry off a Blanaida
goschkevitschit ; this is a butterfly resembling a very large
Pararge megera ; it did not appear to struggle at all.
Sideways attitude or “list” when at Rest.
In December, 1878, Col. C. T. Bingham noticed this rest-
ing attitude in a species of JMelanitis, but the account was
not published till many years afterwards.* EH. H. A.’s
papers in the “Times of India,” which contained a refer-
ence to this habit in JZ. ismene (p. 203), reappeared as “ A
Naturalist on the Prowl” in 1894.
In the summer of 1903 Dr. Dixey and I noticed this
habit in several British Satyrids at Mortehoe, N. Devon.
In Epinephele janira the list may amount to 15°—30°; in
E. hyperanthus (G. B. L., 1894) to 20°; in Pararge xgeria
and P. megera to 25°; but in Satyrus semele it reaches 40°
or even 50°. This list may be to right or left in the same
individual. The insects appear to settle in the upright
position, then to draw the fore-wings partly within the
hind-wings, and by a third distinct movement to throw
themselves over to one side or the other.
To the above insects having this habit I can now add
the following Indian Satyrids :—
Mycalesis indistans, slight list.
Hipparchia parisatis, 20° to 30°.
Aulocera swaha, 45° to 50°.
In the last-named species the same individuals were
observed sometimes to go over to the right, sometimes to
the left ; one was seen to make three efforts, getting further
over each time. A specimen of H. parisatis was observed
walking about with a list of 20°.
To these observations I may add that at Yokohama, May
19th, 1904, Blanaida goschkevitschii, a Satyrid like a large
P. megera, was observed with a list of 40°.
There is no doubt that this sideways attitude makes the
insects less conspicuous when resting on a flat surface, but
* See extracts from Col. C. T. Bingham’s Diary for December,
1878, quoted in Trans, Ent, Soc., 1902, p. 363.
136 Dr. G. B. Longstaffs Notes on the Butterflies
I have satisfied myself from observations on English
Satyrids that the attitude is more often adopted by the
butterflies when sitting in sunshine than in shade. Now
if the list be away from the sun the shadow would be
increased, but if towards the sun it would be diminished,
in some cases even to extinction. Numerous observations
are required to determine whether the list has any relation
to the sun’s position. I would however remark that in the
case of a butterfly with cryptic colouring on the under-side
the shadow is in many cases far more conspicuous than
the butterfly itself, as I frequently observed in India.
Obviously, therefore, economy of shadow might be a
considerable protection. Now, near Simla in October,
1903, in the case of Pararge shakra, a butterfly closely re-
sembling P. megexra, I noted three individuals in succession
settled with their backs to the sun so as to reduce the
shadow to a mere line. This was unfortunately just as I
was leaving the locality where the species occurred, but I
did not observe any instances to the contrary. I should add
that in P. shakra I looked for, but did not find any list.*
Scents in Butterflies.
Owing to the imperfections of the human nose these are
very difficult to detect and to describe, nevertheless certain
definite results were obtained.
* Since the reading of the paper Prof. Poulton has called my
attention to the following interesting observation by Mr. E. E.
Green. “ M. ismene is an adept at concealing itself. It usually
pitches amongst fallen leaves where its form and coloration are
sufficient concealment. But even on bare ground the insect is often
extremely difficult to localize, though the approximate spot may have
been carefully noted. I have watched the fly, immediately after
pitching, alter its position so that its axis is directed towards the
sun, thus casting no shadow.”—“ Notes on some Ceylon Butterflies,
Spolia zeylanica,” vol. ii, pt. vi, Aug. 1904, p. 76.
For the following reference I am also indebted to Prof. Poulton :—
Prof. G. H. Parker has clearly established that in the United States
when Vanessa antiopa, L., after a flight settles in full sunshine with
wings expanded, it speedily so adjusts its position as to place the axis
of the body as near as may be parallel to the sun’s rays, with its head
turned away from the sun. Some of the genus Grapta have the
same habit. He thinks they do this to display their colouring to the
best advantage. The bearing of his interesting observations on the
cryptic attitude of Satyrids is that they prove decisively that a
butterfly can acquire the habit of definitely orienting itself. If one
can do this for one purpose, another species may do it for a different
purpose, e.g. concealment, Mr, Parker gives a Bibliography of the
subject.—“ Mark Anniversary Volume,” Cambridge University, Mass.,
U.S.A., Article xxiii, p. 453-469, 1903.
Observed in a tour through India and Ceylon. 137
(1) The rape scent. Dr. Dixey and I have observed a
slight scent in Ganoris rape well compared by Mr. Selwyn
Image to that of sweet-briar, though the comparison is not
exact. Curiously enough I have been able to prove to my
own complete satisfaction the existence of the same scent,
or one scarcely distinguishable therefrom, in several
Pierines, viz. Delias eucharis, Ganoris canidia, Huphina
nerissa, Catophaga paulina, and Belenois mesentina.
I think it is confined to the male sex, but cannot speak
very positively. .
(2) The brassicx scent. This is fainter than the pre-
ceding; 1 compare it to violet-powder. It is confined to
the male. This scent I did not find in any Indian
butterfly.
(3) The napi scent. Far the strongest, and quite
unmistakable. It is by common consent compared to
lemon-verbena, but it is not identical therewith. This I
did not meet with in India, but it was unmistakable in
the male of the Japanese Ganoris melete, Mén. [My
specimens were of what Leech calls the Japanese spring
form = G. aglaope, Motchulsky, = G. megamera, Butl.]
It proved equally distinct in the male of G. oleracea, Harr.,
a North American form of napi.
That three species of one genus have as many distinct
scents, but that one of these extends to members of several
widely-separated genera is very remarkable, and to me at
least totally unexpected. I cannot help thinking that
when these scents have been more studied and are better
understood they may prove of great value in the solution
of phylogenetic questions.
(4) Several Danaids of different genera have a strong and
distinct odour of a disagreeable character, very suggestive
of acetylene. That it is possessed by thie males I am
certain, but cannot say whether it is confined to that sex.
The species are Crastia core, C. asela, C. amymone (at
Macao), Lsamia midamus (at Hong Kong), Parantica
ceylonica, and Chittira fumata. A single specimen of
Pademma kollart had a somewhat similar odour. In
several cases (in at least three of the above), the scent
was so strong as to be distinctly perceptible when the
butterfly was fluttering in the net (as it is indeed in the
case of Ganoris napt).
(5) Limnas chrysippus has a faint unpleasant odour like
cockroaches, or musk-rats. I suspected it to come from
138 Dr. G. B. Longstaff’s Notes on the Butterflies
the pouches on the hind-wings of the male, but more
observations are needed.
In the case of Limnas genutia, Tirumala limniace,
Pararge shakra, and Colias nilgiriensis the existence of
scents was suspected, but the results were ambiguous.
(6) The observations of Wood Mason were confirmed
in Catopsilia pyranthe and C. pomona. The tufts on the
wings of the males gave out on stroking a scent that may
be compared to jasmine, though I think it more like
Polianthes tuberosa.
Seasonal Forms.
With a view to seeing what light, if any, my fragmentary
observations might throw upon this puzzling subject, I
have adopted the following method:
In the Register, or Index, of my captures I noted to
every Pierine Dr. Dixey’s estimate of its seasonal character,
and then made my own (far less weighty) estimates of the
seasonal characters of the genera Precis, Melanitis, Mycalesis,
and Vphthima, and then analyzed the results for localities,
or groups of localities. ‘The seasonal characteristics were
classed under the following five heads :—
(1) Wet-season form, including “ wet,” “very wet,” and
“extreme wet.”
(2) Somewhat wet form, including “intermediate in-
clining to wet.”
(3) Intermediate form.
(4) Somewhat dry form, including “intermediate in-
clining to dry.”
(5) Dry, including “ very dry” and “ extreme dry.”
Without prejudice, and for the purpose of this grouping
only, I took Catopsilia gnoma to represent the dry-season
form of C. pyranthe, and in like manner Catopsilia catilla
and pomona to be dry-season forms corresponding to a wet-
season form C. crocale.
It must be borne in mind that such a classification is
necessarily very vague, for while the extreme forms are
easy to place it is most difficult to assess the numerous
intermediate specimens.
Observed in a tour through India and Ceylon. 139
® 3 a
ge /elel
~ S 2
Precis orithyia 22 4
»> @none ee ili cce 6
5, lemonias . all ef 3
», tphita a 1
Catopsilia pyranthe 1 a 1 Simla and Kalka,
Ieias marianne . ar 1 Oct. 10-20, 1903.
Terias heeabe alien Ti oe Slight showers.
» lxta “ae 2 21 1
Huphina nerissa Su | aSuallsroaet
Tora 14} a] 4| 1) 15
Precis orithyia 3
>> enone 2
>» almana ne 4 Peshawar and
Yphthima balanica 1 Ne pets | (ne ca Malakand, Oct.
Terias hecabe Pitas 2 24) al 22-29, 1903.
Teracolus etrida 1 No rain.
TOTAT Ss PO te: Zaller LO
Precis orithyia poe 1
», amana age cal ase
>, lemonias . ies 4
Catopsilia pyranthe (eo
op pomona iL Aeon este. |) ee
Ixias marianne . || 1 a B Dangsatiiaetban
ea FES ia and Delhi,
Terias ee 5 Been ecens || sche Oct. 31-Nov. 12,
Teracolus etrida | 2 7 || @ 1903.
ri protractus . Da epeems ered ence n\\Gcoc i eerey
» puellaris LGN eco fe 8! ih) 3}
a calais Calan ats
Huphina nerissa 2
Appias libythea aes 530 1
ToraL 18; 5 5 | 5) |) alts!
140 Dr. G. B. Longstaft’s Notes on the Butter flics
Naini Tal, Luck-
now end Benares,
Nov. 16-Dec. 2,
1903.
No rain.
Caleutta, Dec.
4-12, 1903.
No rain,
g|/E |e 4]
2 ge ieeamer |e
FPilg|/e|2|4
Precis orithyia DS ret
y, @none 2
5, almana i 1
5, lemonias . 1 | | 2 |
5» tphita | 3
| |
“phihima philomela . i
Mycalesis perseus 1 || | 1 |
Catopsilia pyranthe it} oa | 9"
| |
f pomona | ! 1 | 1 |
Ixias marianne . | | 1 |
yy pyrene | | | 1 |
Terias hecabe 1| ! | 1
Huphina nerissa He Ath !
Toran 3/3 | 2 || 4 | 16
ee cabrierin ; : 1 | | 1 if
», lemonias . | | nooeyh |
|
5 alates 1 1 | ies al | |
Melanitis ismene | | 4 |
Mycalesis indistans | 2
Catopsilia pyranthe 2 | I) al |
4 pomona 1 | 4 | |
Ixias pyrene } 1 | |
Terias hecabe 1 | a 9 |
Huphina nerissa 1 | |
ToTaL 5 Ba Ball | aa
Observed in aw tour
Precis lemonias .
Melqnitis ismene
Mycalesis indistans
Catopsilia pyranthe
Ixias pyrene
Terias heeabe
», leta
Huphina nerissa
3 nating
Tachyris hippo .
Prioneris thestylis
Hiposcritia lalage
ToraL
P Hinge ia
y» enone
», almana
», lemonias .
Yphthima imica.
Catopsilia pyranthe
Ivias marianne .
Terias hecabe
Pa ccebay
Trracolus etrida
oA puellaris
Huphina nerissa
TOTAL
through India and Ceylon. 141
8 Ree
2 | @ || 3 || @ | §
g a || @ a 3S
a | 5 | 8 We | a
S (ER NBS eet |p
Fla) 4 lala
el +| |
| || | iL
| |} 1
al |
i | 8
| h 3]
|
Yel Belge
| | |] |
| | | “ | Y Darjiling, Dee.
| 1] 9 |
| es | 13-22, 1903,
1B ee AE | are
| i y No rain.
| | eee 2
| ne 1 |
| ma
| 2 4
Tee, Us| So, Ue | 25
fae = = — ; ees Tal —t eo _ ——$$$_—$—=
| |
1 |
eae 2. |
| | 1 | Bankipur, Jhansi,
| i |
2 Agra, Jaipur,
| ‘
1 : a 1/2 ‘Ajmir and Mt.
| Nye
| 2; 5 Abu, Dec. 24,
ay. i | | eg
| Nasa al eel 1903-Feb. 8, 1904.
|
3) |) SGnh nO ‘ 3
A Shght rain Jan.
1 Dol 2 3 |
14-23.
1
7
3 2 8 | 41 37
142 Dr. G. B. Longstaff’s Notes on the Butterflies
2iaié
S\E EES
2) 28 Wie wie |
Fla@a\|siesa
Precis anone i ig ie
» almana oa | D)
Melanitis ismene Heal nee
Hypanis tlithyia es AAllitee
Catopsilia pyranthe a Pa 3
E pomona | 1 Bijapur, Ananta-
Txias marianne . | 2 pur and Bangalur,
Terias hecabe | ox 1 Feb. 16-23,
» lata ‘ lie. 1904.
Teracolus etrida ? 2 Q9/| 5 No rain.
Pa dulcis He | ; 1
», amatus 2. ee
5, eucharis {sag Are 1
ToraL 3 4 || 11 | 12
Precis orithyia . (toes yl iieers
», @enone Hi) ee 1
», almane i ee
»» lemonias . ia Pe
Melanitis ismene 2
Mycalesis perseus 3
Yphthima hiibneri be | 1 é Mls
Hypanis ilithyia 1 laa fee Nilgiris, Trichin-
Catopsilia pyranthe 2/1 2 dpali, Tanjur,
cs pomone ce cM pe Madura, Feb. 24—
Ixias marianne . asf Veet) cise 1 March 7, 1904.
yy pyrene 9 || 2. 1 al Very slight rain
Terias hecabe a 2 Ill ax 2 in Nilgiris.
Teracolus elrida 1 a
i eucharis se {| Goo |h} Goo? IIe gba 1
an dance ‘ 5) 5 2 1
Huphina nerissa 1 3 ee
Catophaga paulina , 1 | asl 74
ToraL , 5 | 10 || 10 || 13 | 20
Observed in a tour through India and Ceylon. 143
sisi &
glaieial:
Precis almana . ; : £ 1
», atlites . ; Saal hee | AU See
Mycalesis mandata . Sh|| demaslal lege | goa) ill ane
Catopsilia pyranthe . : 2
A pomony . Sl) Ate wi diate 2 | ZN aa | Ceylon, March
Ixvias pyrene. : Mel eecien iss | Bee | Dll biees 10-26, 1904.
Terias hecabe ; ol) ie | 1 | 1 Several showers.
»» lata ; P 3 {eset Pear | sogullieees 1
Huphina nerissa : j ast eee | il | ea MiL all
Catophaga paulina | 2 | 2 |
ToTAL . : 8] 4] 6 ps)
There was a storm at Simla on October 10th, and a few
trifling showers during our expedition to Baghi, but we
saw no sign of rain after that, and indeed scarcely a cloud,
save at Kurseong, until January 14th, when there was a
thunderstorm at Jhansi. There were then several very
slight falls of rain terminating with a long but not heavy
rain on January 23rd. There was a very slight fall at
Konutr on the night February 29th—March Ist. Then no
further rain till Kandy, March 10th. There were several
showers in Ceylon.
At Simla the effects of the monsoon were not quite past,
and wet-season forms were slightly more numerous than
dry ; the same applies to Ceylon. At all the other places,
as might have been expected, dry-season forms predomin-
ated, Calcutta occupies an intermediate position.
It must however be admitted that to prove a species to
be dimorphic js not necessarily to prove that the forms are
associated with seasons, In the genus Precis, so far as my
very few observations (limited to the dry season) are worth
anything, the two forms ocellated and non-ocellated seem
144 Dr. G. B. Longstaff’s Notes on the Butterflies, ete.
to be closely associated with wetness and dryness respect-
ively. Catopsilia pyranthe, as Dr. Dixey has shown,
occupies a far less clear position, and I may add that
Terias hecabe did not appear to me to follow any rule. The
two forms were taken together in most places.
Many dwarfed specimens of the genus Precis were met
with as the season advanced; with the exception of one
P. almana, var, asterve, they were all of the dry type, most
of them markedly so. The smallest Z'erias hecabe was of
the dry form, so was a dwarf Teracolus dulcis; four dwarfed
T. etrida were half dry, half intermediate. A dwarf
Belenois mesentina was dry, but a dwarf Catopsilia pyranthe
and a dwarf Huphina nerissa were intermediate, while a
dwarf Teracolus calais was actually of the wet-season form.
In conclusion I have to thank the President for much
valuable assistance in many ways; I am greatly indebted
to Mr. Hamilton H. Druce for most kindly naming all my
Lyczenids and Hesperids, to Sir George Hampson for much
help in naming my moths, to Mr. W. F. Kirby for kindly
naming my Orthoptera and Neuroptera, to Col. C. T.
Bingham, Mr. G. E. Austen, and Mr. Claude Morley; to
the Rev. F. D. Morice for naming my Hymenoptera; to
Commander J. J. Walker, R.N., whose practical experience
in many lands was of much assistance, and to Mr. W.
Holland of the Hope Department for constant help, while
to Dr. F. A. Dixey I am indebted not only for the names
of all my Pierines and much information about them, but
for continual encouragement and inspiration.
VII. The genus Criocephalus. By D. SHarp, M.A., F.R.S.
With Notes on the habits of Asemum striatum and
Criocephalus ferus. By F. Ginuerr Smiru.*
Prare 1x
In the Munich Catalogue of Coleoptera (vol. ix, 1872,
p- 2789) Criocephalus is allotted ten species. Another
species has since been added by Herr Edmond Reitter,
C. syriacus, Wien. Ent. Zeit., 1895, p. 86. C. coriaceus,
Motsch., must be rejected as the description is totally
worthless (cf. p. 158), and no type is known. C. mex-
canus, Thoms., is treated as a synonym of C. obsoletus.
Great confusion has always existed as to the species,
and there are numerous errors in the Munich Catalogue.
A revision of the genus will therefore be useful as a step
towards the attainment of further knowledge.
I owe my thanks to Mr. Gahan for assisting me in the
examination of the specimens in the British Museum, and
to M. Réné Oberthiir, Herr Edmond Reitter, M. Louis
Bedel, Mr. G. C. Champion, and Mr. George Lewis for the
communication of specimens.
DISTRIBUTION.
The genus Criocephalus is distributed over the Northern
hemisphere, occurring in about equal numbers in the Old
and New Worlds; it does not pass the equator, and scarcely
touches the tropics, except in the highlands of their northern
parts.. This distribution is probably chiefly determined
by the distribution of coniferous trees, to which it is
believed the species of Criocephalus are confined, their
food being found in the wood of these trees. C. rusticus
has a natural distribution probably as wide as that of any
other Coleopterous insect, it is found from Norway to
* Although Mr. Smith has not had any part in actually drawing
up the systematic part of this paper, I have received very important
assistance from him in this matter. He first noticed the peculiarity
of the deficient spur in the group. The notes on the larve have
been made by us conjointly. I, of course, have had no part in Mr,
Smith’s account of his observations in the field. —D. 8,
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 10
146 Mr. D. Sharp on
Algeria, and in the other direction to Japan, and even in
Yunnan. It is probably occasionally carried about with
timber, but this cannot have much affected its wide dis-
tribution. There is no reason for supposing that any of
the other species are increased in the area of their dis-
tribution by commercial means, as timber is not imported
into most of the countries in which the species live.
TAXONOMY.
Criocephalus has been placed in the group Asemina
of the great subfamily Cerambycides, a group of only a few
genera, but as to the composition of which systematists
have very widely differed. Lacordaire (Gen. Col. viii,
p- 205, as Asemides) assigns to it five genera, Asemum,
Nothorhina, Cyamophthalmus, Tetropium, Criocephalus.
Thomson (Syst. Ceramb., p. 463, as Asemitex) places in it
nine genera, excluding Tetropium, but adding five other
European genera. Leconte and Horn (Class. Col. N. Amer.,
ed. i, 1883, as Asemini) adopt Lacordaire’s view,
Cyamophthalmus being, however, unknown to them; and
the same remark applies to Schiddte (Danmark’s Ceramb.,
p. 507, as Aseminz), who also did not include Nothorhina.
Duval (Gen. Col. Kur. Céramb., p. 125, as Criocephatites)
associated Criocephalus, Cyamophthalmus, Tetropiwm, placing
Asemum and Nothorhina in a distinct group (Asemites).
Ganglbauer and Bedel, who have, each one, published an
excellent work on the European Cerambycide, have not
dealt at all with the question of minor groups. Kraatz
has recently added the genus Megasemum (Berlin. Ent,
Zeitschr., 1879, p. 97) without, however, giving any char-
acter that would distinguish the genus from Criocephalus.
On examination it appears that none of the genera
hitherto associated with Criocephalus are closely allied to it,
except Megasemum. Criocephalus possesses the very ex-
ceptional character of having only one spur on the front
tibia, while the other genera usually associated with it
have two.
Criocephalus syriacus, Reitt., is here made a distinct
genus, Cephalocrius; it and Megasemum agree with Crioce-
phalus in the unicalearate tibia and are extremely closely
allied to it.
Megasemum was distinguished by Kraatz solely on the
strength of shorter antennz. This however does not hold
the genus Crrocephalus. 147
good for the male which has longer antennz.* Even in the
female the antennz are not shorter than they are in the
N. American Criocephalus asperatus, to which species
Megasemum has a great resemblance. Megasemum differs
however in the form of the prosternum, which is concave in
profile instead of convex as in certain Criocephalus, or flat
as in other species. It has moreover the clypeus much
more elongate, the antennze more distant from the man-
dibles and the eyes not extended to the under-surface of
the head.
I describe also a new genus, Cephalallus, that is closely
allied to both MMegasemum and Criocephalus. The four
genera in question form a natural group to be called
Criocephaline, distinguished from the Aseminew by the
presence of only one spur on the front tibie.
The four genera of Criocephaline may be distinguished
as follows—
(1) Criocephalus. Terminal joint of palpi only moder-
ately broad at tip; prosternum protuberant in
front, or flat ; clypeus short.
(2) Cephalocrius. Terminal joint of palpi very broad at
the apex; prosternum nearly flat; clypeus very
short.
(3) Megasemum. Terminal joint of palpi only moder-
ately broad; prosternum slightly descending ;
eyes not extended on under-surface of the head,
(4) Cephalallus, n.g. Terminal joint of palpi not very
broad at tip; prosternum descending ; eyes ex-
tending to under-surface of the head.
The Criocephalinx should come at the beginning of the
subfamily Cerambycides, as being one of the most primitive
forms of Longicorn Coleoptera. The only other genus of
Longicorns that I have found to agree with Criocephaling
as to the unicalcarate tibize is the genus Philus, which
forms an annectant link between Cerambycides and
Prionides. This character is not to be looked on as
primitive. The primitiveness of Criocephalus depends not
on such points as this, but on the fact that it differs but
* The male of Megasemuwm has not yet been described, It is
similar to the female but has the antenne longer and much more
hairy. I have seen only one specimen of this sex, it is in the British
Museum Collection, For my knowledge of the female I am indebted
to Mr. George Lewis, who met with the species in Japan.
148 Mr. D. Sharp on
little from Coleoptera of other families; that it lacks all
the specializations that are so remarkable in other divisions
of Longicorns, while but little changes would suffice to
make it a member of other divisions, either of Cerambycides
or of Prionides.
CEPHALALLUS, n. g.
Caput brevissimum.
Palpi perbreves, articulo ultimo modice dilatato.
Oculi magni, subtus conspicui.
Prosternum descendens, parum elongatuin.
I have only a single example, of the female sex, and in
a very mutilated state, before me; but it represents a
specics of so much interest that I do not hesitate to name
it and call attention to it.
A slight modification of the prosternum of this insect
would transform it into a Ciriocephalus, while an elongation
of the head would apparently make it a Megasemum.
The labrum is small and has a pencil of hairs near the
front as in Criocephalus ferus. Mandibles extremely short,
each with a single, strong tooth in the middle. Palpi very
short, the terminal joint only slightly, but still distinctly,
dilated at the tip. Ligula extending on each side as far
as the middle of first joint of labial palpus. Eyes convex,
extending on to the under-surface of the head, but widely
separated there as in Criocephalus rusticus.
Prosternum margined in front; behind the margin ex-
tending obliquely downwards; only very slightly separating
the front cox. Metasternum elongate, terminal segment
of the female less elongate than in Criocephalus. Hind
femora hairy behind, but not densely so.
If the genus be not maintained as distinct it will be
necessary to unite Megasemum and it with Crrocephalus.
CEPHALALLUS OBERTHURIL, sp. n.
. Elongatus, angustus, convexus, piceus, elytris ferrugineis ;
5 o] DS ’ ) ) ro) d
prothorace angusto, convexo, asperato, sculpturato, peropaco, medio
late leviter impresso; elytris dense fortiter punctatis, subtiliter,
sex-costatis. Long. 22 mm.
Hab. TsEkovu in North Yunnan. (Coll. R. Oberthiir.)
The general form is that of a narrow Criocephalus, but
from all the forms of that genus it is distinguished by the
very dense punctuation of the elytra, which renders them
the genus Oriocephalus. 149
finely rugose. The eyes beara few fine but moderately long
sete between the facets. The clypeal suture is very
deep, and forms an acute angle inthe middle. The thorax
is a good deal narrower than the elytra, and only slightly
broader than the head and eyes; it is scarcely transverse,
but is convex transversely, it bears uumerous sharp,
elevated granules, between which it is minutely rugose,
and has some fine hairs; there is a vague, broad depression
on the disc, and behind this a feeble prominence in the
middle just in front of the scutellum. The elevated lines
on each elytron are not very distinct, the two on the
middle do not reach the apex, and the lateral one cannot
be seen either at the base or apex, the sutural angle is
minutely spinose.
The legs and antenne are very mutilated; all that
remains of the former are some portions of femora, and
one front tibia. Six joints remain of one antenna: these
are rather slender, and bear numerous erect hairs; the
first joint is rather more than twice as long as the second ;
the third is not quite three times as long as the second,
the fourth is about as long as the first, the fifth is shorter
than the fourth, and the sixth shorter than the fifth and
but little longer than the first.
CEPHALOCRIUS, n. g.
Antenne fere corporis longitudine, longius hirtella.
Palpi articulo ultimo ad apicem latissimo.
Type Criocephalus syriacus, Reitter.
Closely allied to Criocephalus, the C. syriacus of Reitter
is well distinguished by the above characters. I have
seen only one example and am not able to describe the
parts of the mouth more fully. Herr Reitter informs me
that in the female the antenne are less hairy than they are
in the male. The really important character of the genus
is the form of the palpi.
CEPHALOCRIUS SYRIACUS, Reitter.
Criocephalus syriacus, Reitter, Wien. Ent. Zeit., 1895,
p. 86.
g¢. Angustus, subdepressus, ferrugineus, elytris bicostatis et crebre
punctatis, antennis corporis longitudine subtus hirsutis ; tarsis
posterioribus articulo tertio profunde fisso, articulo basali elongato,
Long. vix 15, lat. 3} mm.
150 Mr. D. Sharp on
This is very distinct by the strongly dilated terminal
joint of the palpi which is broadly securiform, both on the
maxilla and labium. The hind tarsi are long and slender,
with remarkably long basal joint. The eyes are rather
more prominent and have a greater extension on the
under-surface of the head than they have in C. rusticus.
The sculpture of the gular area is transversely rugose, the
impressed punctures not numerous. This insect was
kindly sent home by Herr Reitter with the locality “ Syria.”
In his description he mentions, Haifa, Akbés. There is also
a male example from Akbes in the Oberthiir Collection.
N.B.—The British Museum Collection contains a
specimen of this genus from the collection of Dejean. It
is in very bad condition, and I do not know whether it is
C. syriacus or another species. In Dejean’s collection it
was labelled “ rwsticws, Gall. mer.” The locality from
which this specimen really came is, I consider, extremely
doubtful.
CRIOCEPHALUS, Mulsant.
Criocephalus (Dej.), Muls, Longicornes, ed. i, p. 63.
I have already mentioned the characters that distinguish
this genus from the allied forms (p. 147).
The species present a great uniformity of appearance
and as they are fairly numerous it is not easy to distin-
guish them. Particular attention should be given to the
form of the thorax, the length of the small setz between
the facets of the eyes, and the form of the third joint of
the tarsus.
The genus has been subdivided by the eyes being
“hairy” or “bare.” But this is incorrect; all the species
have some sete between the ocular facets, and it is only a
question of longer or shorter. The same remark applies
to the lobing of the third joint of the tarsus, which differs
somewhat from species to species. The labrum apparently
offers a better character, but unfortunately it is not easy
to see without opening the mandibles and this it is some-
what difficult to do, as they are closed by very powerful
muscles, which cannot be ruptured after the insects are
dead and dried.
Group <A.
Labrum without pencil of hairs in fovea on the middle
of its area, Eyes with long hairs between the facets,
the genus Criocephalus. 151
CRIOCEPHALUS DECEPTOR, sp. n.
Depressus, fusco-niger, breviter griseo-pubescens, opacus ; thorace
fortiter transverso; elytris bicostatis fortiter punctatis, Long,
20 mm.
This species is an extremely distinct one, though it
looks at first sight very like a moderately-sized individual
of C. rusticus. It is however more depressed than that
species, and this character removes it from all the other
described species** From C. rusticus, OC. deceptor differs
strongly by the tarsal structure, the third joint being only
slightly lobed. The unique specimen is much damaged ;
the prosternum is not in the least degree protuberant or
convex, and the under-surface is remarkably flat. The
labrum is of the type of that of C. rusticus.
Hab. WESTERN TIBET, one specimen in the collection
of the British Museum,
CRIOCEPHALUS RUSTICUS, Dej.
Brunneo-ferrugineus, vel fusco-niger, subplanatus, elytris sat dis-
erete bicostatis, inter costas perspicue punctatis ; prothorace trans-
verso, ad latera fere regulariter rotundato, posterius plus minus
angustato. Long. 14-30 mm.
The male has the basal portion of the antenna, and the
legs markedly thicker than the female.
The synonymy of this species, as distinguished from
@. ferus, is very doubtful, Some of that given in the
Munich Catalogue is taken from writers who did not dis-
tinguish the two forms. The C. rusticus of Dejean’s
Catalogue and collection is however certainly this species,
as is also Mulsant’s C. rusticws, and var. B., |. c., p. 64, Also
C. rusticus, Woll. Cat, Col. Madeira (1857), p. 124 (nec
Col. Canar., for which see C. pinetorwm).
The species is readily distinguished by the strongly
transverse thorax, which is only one-fifth of the length of
the elytra, and by the gular area being very indefinitely
limited, rugose in front and feebly punctate behind, and
with only a few suberect long hairs, forming a strong
contrast to what exists in @. ferus. In finely-developed
* There is an undescribed (?) species in the collection of the
British Museum, coming from the Amanus mountains of Asia Minor,
which is also very depressed; but in other respects this insect is
very different from C. deceptor. .
152 Mr. D. Sharp on
specimens the sculpture of the gular area is more largely
developed, there being both more rugz and more punc-
tures, but the sculpture always retains the same character.
The sete between the facets of the eyes are so long that
they can be easily detected ; the third joint of the tarsus
is divided nearly to the base, and the lobes formed by the
division are large and symmetrical. The scutellum is
always a little impressed along the middle. The meta-
thoracic epimera are comparatively narrow.
The species is very variable in size, form and sculpture,
but after repeated examinations I fail to find any reliable
character to separate even the most extreme forms. The
length varies from 14-80 mm. and the width from
4-7} mm. The female is generally larger than the male
and is somewhat darker in colour, but is never quite black.
There are sometimes three depressions on the thorax, and
these in a few specimens are deep and well-marked. The
sculpture of the upper-surface also varies: it 1s sometimes
coarser and less dense, and the insect appears slightly
shining. The elytra usually lave the tip-sutural angle
sharp, or rectangular, but this also is variable. The rugee
on the gular area vary a good deal, but the sculpture there
never has the character of definite depressed punctures,
and it bears only an irregular, somewhat scanty pubescence,
never the regular dense pilosity of the C. ferus group of
species. The setosity on the labrum also varies.
I have examined the male organs in some of the most
remarkable of the varieties and find they agree. The two
closely appressed laminze that form the tip of the intromi-
nent organ are not of equal length, but the lower one is
the longer and is very sharp. In this respect C. rusticus
differs from C. ferus.
I think it quite possible that careful study of a series of
good examples of both sexes may show that there is really
more than one species mixed under C. rusticus. I should
recommend a thorough scrutiny of the labrum as likely
to give good results, but this is unfortunately attended
with great practical difficulties.
The species inhabits apparently all the temperate zone
of the Old World in the Northern hemisphere, wherever
there are suitable timbers; what species of conifer it may
prefer I donot know; but doubtless the Scotch fir (Pinus
sylvestris) is one of them. I have specimens before me
from Norway (Champion); Hanover; Vienna; S. France ;
the genus Criocephalus. 153
Spain, Bronchales, Tragacete, La Granja (Champion);
Corsica, Vizzavona (Champion); Akbés (Coll. Oberthiir) ;
Amurland ; Japan (Lewis); Tsekou in North Yunnan (Coll.
Oberthiir) ; and I have seen specimens from Madeira and
other localities.
The species has recently been discovered by Colonel
Yerbury at Nethy Bridge, near our wildest Highlands,
where the Scotch fir is still extant in its natural state.
The specimen he found was on a sprig of heather near a
large stump of the Scotch fir. It isa fine female and is
now in the collection at the British Museum,
This specimen is I think undoubtedly a native of
Britain, and it is of importance as showing that the dis-
covery for the first time of a Criocephalus m a locality is
not sufficient evidence of its being an introduction. Mr.
Gilbert Smith’s extremely interesting notes on the natural
history of C. ferws and Asemum striatum should also be
considered in this connection. There can however be
little doubt that this species is liable to be carried about ;
though probably this only occurs as an exception, even
when large blocks of timber are transmitted. A specimen
has been found in a coal-mine in Wales, where it was
pretty certainly carried in the wooden props; and _ if
foreign fir-wood be used for railway sleepers it may thus
be transmitted. The case recorded of its introduction by
means of timber for masts must also be noticed.
C. rusticus was not known to Wollaston when he wrote
the “ Insecta Maderensia” (published in 1854), but it after-
wards became common in the plantations of introduced
fir-trees, and in 1857 was recorded by him in the category
of “undoubtedly introduced ” species. [The “ C. rusticus”
of the Canary Islands is C. pinetorwm.] It appears to
have spread with rapidity in the Madeiran group, as in
1865 Wollaston recorded it from. the extreme summit of
the small island called Dezerta Grande. But this must
be received with some caution, as it 1s far from impossible
that C. pinetorwm occurs in Madeira as well as in the
Canaries.
CRIOCEPHALUS COREANUS, sp. 0.
Q. Fusco-niger, haud nitidus, antennis pedibusque piceis ; pro-
thorace parum transverso, densissime rugoso-punctato, ad latera
numerose muricato ; elytris costis duabus parum elevatis, punctisque
impressis per-numerosis, Long, 22 mm,
154 7 Mr. D. Sharp on
In this species the thorax is about one-fourth the length
of the elytra, whereas in C. rusticus it is about one-fifth.
On measuring two specimens of about equal size I find
them to be, C. coreanus: thorax 33, elytra 154 mm.
long. C. rusticus: thorax 2{, elytra 15 mm., and these
proportions persist (with some variation) in the individuals
of different sizes of C. rusticus. The head and thorax of
CO. coreanus are also considerably narrower, and the
antennz more slender. The length of the thorax is
42mm. The sculpture of this part is denser and finer
than it is in C. rusticus, The scutellum is narrower than
in ©, rusticus, almost flat, with a slender polished line
along the middle.
Although I have only one mutilated female of this form
I have little doubt that it represents a species closely allied
to C. rusticus but distinct, The structural characters
appear to agree with those of C, rusticus. It should be
noted that the Japanese form of C. rusticus is very
different from C. coreanus. :
CRIOCEPHALUS OBSOLETUS, Randall.
Criocephalus obsoletus, Leconte, J. Ac. Philad. ser. 2, II,
p- 196 [ =obscwrus, id. p. 36].
Criocephalus mexicanus, Thomson, Classif. Longic. 1860, p.
260; Bates, Biol. Cent. Amer. Col., V, p. 15.
This species is extremely near C. rusticus, but will
probably prove to be distinct. I have only one specimen
at my disposal. The sculpture of the upper-surface is
rather coarser. The thorax has a slight angulation at each
side ; the second joint of the antenna is a little longer,
and the scutellum is broadly impressed.
Hab. N. AMERICA: widely distributed and apparently
not uncommon in Mexico, “in pine forests at the higher
elevations.”
I have followed Mr, Gahan, in his arrangement of the
British Museum Criocephalus, in considering C. mexicanus
of Thomson and Bates to be a synonym of C. obsoletus.
For remarks on the North American species generally
see under C. nubilus.
GROUP 2.
Labrum with a pencil of hairs placed in a fovea in the
middle near the front,
the genus Criocephalus. 155
N.B.—I have not been able to examine the labrum in
C. exoticus and C. tibetanus, and one or both may therefore
prove not to belong to this division, Neither the hairs on
the eyes nor the lobing of the tarsi can be used as sectional
characters as there are intermediate conditions of both of
them. But in this section the ocular sete are usually
short, and the basal lobes short.
CRIOCEPHALUS PINETORUM, Woll.
g. Criocephalus pinetorum, Woll., Journ. Entom. II, p.
103 (1863); and Cat. Col. Canar., p. 388 (1864).
2. Criocephalus rusticus, id. Col. Atlant., p. 8345 (1865).
Wollaston first fell into the double error of considering
the sexes of C. pinetorum to be different species and of
determining the female to be C. rusticus. This was not
recognized by him in revising the work in 1865, and being
then baffled to distinguish the C. pinetorwm he sank the
species as being merely C. rwsticus, from which however it
is totally distinct. This does not complete the sum of
Wollaston’s confusion; for he considered the Madeiran C.
rusticus to be the same, though he noticed the different
habitat in the two islands. The insect found about
Funchal in Madeira in connection with the introduced,
lanted, conifers is really the C. rusticus of authors, while
the C. rusticus of Wollaston Coll. Atlant. (nec auctt.),
found in the native pinals of the Canary Islands, is
the female of C. pinetorum, Woll., and is a quite distinct,
precinctive species. As Wollaston found the two together
in the Canaries it appears remarkable that he should not
have considered them to be the sexes of one and the same
species. Probably he suspected them to be so, but not
having detected the characters that distinguish the female
from C. rusticus and C. ferus (which at that time were
generally confounded in collections), he finally came to the
erroneous conclusion that all were one species. Hence
this good species has since been lost sight of. Bedel how-
ever suspected some error; and in placing the name as a
synonym of C, rusticus added a note of interrogation.
This species has the eyes bare (2. e. with extremely short
setze between the facets) and the fourth jomt of the tarsi
intermediate between that of C. rusticus and C. ferus.
The sexes are very different, and hence Wollaston supposed
156 Mr. D. Sharp on
the female to be C. rusticus. The female differs from the
male by its large size, black colour, and the usual differ-
ences in the length of the antennz and the structure of
the apex of the abdomen.
“ Abounds in the Ancient Pinals of intermediate and
lofty elevations” (Woll. 1865), but nevertheless is exces-
sively rare in collections. Recorded from the islands of
Palma, Teneriffe, and Hierro.
The male is small (sometimes only 10 or 11 mm. long)
and of a brown colour, the gular area is very definitely
limited behind, almost semicircular, and is closely and
coarsely punctured, the bush of erect pubescence moderately
dense.
The female has not been described. I have seen only
two specimens of it, one in my own collection and one in
Wollaston’s Canarian collection, where it stands as C.
rusticus. It is more flat than the corresponding sex of C.
Jerus, and has a more transverse thorax. The punctuation
of the gular area is remarkably definite, and extends over
a large space ; the prosternum is but little convex in front,
and the metathoracic episternum is much narrower. In
these Jatter respects, as well as in the more deeply divided
third tarsal joints, it approaches (. rusticus.
CRIOCEPHALUS FERUS, Mulsant.
Transversim convexus ; mas fusco-niger, femina nigricans ; opacus,
elytris bicostatis, inter costas tantum subtilissime punctatus. Long.
10-30 mm.
The male is usually shorter and paler in colour than the
female and the prosternum is much less convex. This
species is rare in collections, and owing to this and to the
want of knowledge as to the sex-differences are due its
confusion with C. rusticus by entomologists. The two
forms are really profoundly different. The structure of the
labrum distinguishes the two absolutely, but there are
numerous other characters. In the male of C. ferus the
basal portion of the antenng is but little thicker than in
the female. In the female the front of the prosternum
is more convex, and the metathoracic episterna considerably
broader than in C. rusticus, The under-surface is altogether
more closely and finely punctate and pubescent and there-
fore less shining. The scutellum in C. ferus is never in
the least depressed along the middle; and the sculpture
the genus Crrocephalus. ta
and pubescence of the gular area are very different. In
C. rusticus the tarsi have the third joint divided almost to
the base, and the setz on the eyes are very conspicuous ;
while in C. ferus the eyes appear to be bare, and the tarsal
lobes are less perfect.
The species is extraordinarily variable in size, some
specimens of the female being quite three times as long as
the small males, and are broad in proportion; so that it
would take twenty or thirty of the small males to make up
the bulk of one of the largest females. There are also
some variations 1n the proportions of certain parts of the
body, so that I at first thought there to be more than one
species under the name of C. ferus. A series from Algeria
kindly communicated to me by M. Bedel has convinced
me that this is not the case. There is also great variation
in size independent of sex, some of the females being six
or eight times the bulk of other individuals of the same
sex. The distance between the eyes on the under-surface
of the head varies somewhat, and correlative with it there
is a difference in the sculpture of the gular area. I
distinguish the following variety with some doubt, having
seen only two examples.
C. FERUS, var. n. HISPANICUS.
Minor, praesertim angustior, elytris fere ecostatis. Long. ¢ 14,
? 16 mm.; lat. ¢ 9 4mm.
I have seen only a single pair found at Navalperal in
the Guadarrama range by Mr. Champion. They were in
company with Nothorhina muricata and are damaged by
turpentine. There is a slightly larger specimen from the
same locality in the Oberthiir collection. The sculpture
of the gular area appears to be more rugose and less definite
than in other forms of C. ferus, but the distinction in this
respect from certain other Spanish examples is not great.
It would take eight such females to attain the mass of a
well-developed female of C. ferzus.
C. ferus has a very wide distribution, but is rarely met
with, and then usually only in one or two individuals.
Lapland (Coll. Oberthiir from Coll. Thorey); Denmark ;
8. England; France (various localities including Fontaine-
bleau, in August 1898, Coll. Bedel); Portugal; Spain ;
Corsica; Algeria; Pireeus, Besika Bay ; Caucasus (Leder) ;
Akbes (Coll. Oberthiir) ; Madeira.
158 Mr. D. Sharp on
The synonymy of this species is a rather difficult
question. The oldest name that certainly applies to it is
that given by Dejean in his collection as a var. of C. rustacus,
and formally published by Mulsant in 1840. C. polonicus,
Motsch. (1845), is probably this species. As regards Calli-
dium triste, Fabr., Mant. Ins., p. 154, the brief description
gives no special reason for supposing it to be a Criocephalus,
except that he likens it to C. rusticus. The C. lugubre,
Gmelin, is simply an error; he copies Fabricius’ descrip-
tion of C. triste, and, as he gives no reason for changing the
name, he probably merely wrote “/ugubre”’ in error for
C. triste.
The following is therefore the only synonymy for which
I can venture to take any responsibility.
Criocephalus rusticus, var. ferus, Dej., Cat. Coll. (not
described) ; Mulsant, Long. France, ed. I, p. 4.
22 C7. coriaceus, Motsch, Bull. Mosc., 1845, 1, p. 89.
? C. polonicus, Motsch., l.c., p. 88.
C. epibata, Schiddte, Kroyer Tidskr. (3), ii, p. 521 (1864).
C. ferus, Kraatz, Berlin. ent. Zeitschr., 1863, p. 107, and op.
cit., 1872, p. 319; Bedel, Faune Col. Seine, v, pp. 23
and 69,
CRIOCEPHALUS NUBILUS, Lec.
Criocephalus nubilus, Leconte, J. Ac. Philad., ser. 2, II,
p. 36.
This species has strong punctuation on the elytra lke
C. rusticus; the tarsi have moderate lobes like C. pinetorwm.
The labrum, I believe, brings the species in the C. ferus
group. The sculpture of the gular area is coarse and deep,
but occupies only a small area. The eyes bear long sete.
The prosternum is slightly convex in front.
Hab, N. AMERICA.
N.B.—The North American Criocephalus require a
thorough revision, for which the material does not exist
in this country, so far as I know. Leconte, new species
North American Coleoptera, Part II, 1873, gives no less
than seven species as North American. He divides the
genus into two groups: (1) tarsi feebly lobed, C. productus,
(7. agrestis and C. asperatus; (2) tarsi deeply divided with
long lobes, C. montanus, n. sp., C. obsoletus, C. nubilus,
C. wustralis. The matter stands practically as he thus left
the genus Criocephalus. 159
it thirty years ago. C. montanus is perhaps a variety of
C. nubilus. C.obsoletus and C. mexicanus are treated in the
British Museum collection by Mr. Gahan as being the same
species, and he is probably right. C. australis I do not
know; it is probably another genus, as suggested by
Leconte. '
In the Bull. Brooklyn Soc., vol. vii, 1884, pp. 63, 64,
there is given a table of the North American Criocephalus,
but it is not of any importance. It is taken from Leconte;
and some synonymy is added that is apparently taken from
the Munich. Catalozue; there is also a figure of “ C. pro-
ductus,’ which is certainly not representative of that
species, but is more like C. asperatus.
C. nubilus differs from the first Lecontian group of
species by having shorter hairs on the metasternum, and
longer setze on the eyes. It differs from C. tebetanus by
the less transverse thorax and the much larger lobes of the
tarsus,
CRIOCEPHALUS TIBETANUS, sp. n.
@. Major, fusco-niger opacus, prothorace sat transverso, elytris
bicostatis, fortiter punctatis. Long. 24 mm., lat. 6? mm.
Although this insect looks like C. rusticws, and has
sculpture similar thereto, it is allied to C. ferus. No setz
are visible on the eyes, there is a bush of erect pubescence
on the gular area, and the third tarsal joint is only slightly
bilobed. The labrum I have not been able to examine,
so that the position of the species is doubtful. The gular
area is not definitely limited behind, it is very broad,
densely sculptured, rugose in front, rather finely punctured
behind.
Hab, 'TIBET.
The type is in the British Museum, and was found in
Kast Tibet by one of the collectors of the late Mr. J. H.
Leech. A very similar insect from Tibet is the C. deceptor
described on a previous page.
CRIOCEPHALUS EXOTICUS, sp. 0.
¢. Angustus, niger, prothorace haud transverso, profunde bi-
impresso ; elytris obsoletissime costatis et punctatis. Long. 17 mm.,
lat. 5 mm,
Described from a single male in bad preservation in the
collection at the British Museum. The eyes have no long
160 Mr. D, Shatp on
hairs. The gular area is deeply rugosely punctate, and
bears a bush of long black hair. The front of the proster-
num shows some deep rugae. The second joint of the
antennz is elongate, about as long as the first joint. The
third joint of the tarsus is very little emarginate. Struc-
ture of labrum not definitely ascertained. The species is
no doubt quite distinct ; it looks like the North American
C. agrestis or productus, but appears to be nearer to C. ferus,
notwithstanding the intense black colour of the male.
Hab. NortH Cuin Hits, Burma. Lieut. Watson in
1894.
CRIOCEPHALUS AGRESTIS, Kirby.
Angustus, sat elongatus niger, elytris fusco-nigris ; prothorace
bi-impresso ad latera vix angulato; elytris bicostatis, inter cosias
subtilissime sculpturatis. Long. 184 mm., lat. 5 mm.
This North American insect in the male sex greatly
resembles C. ferws; it is rather more slender and shining,
and has a smaller thorax, which has a very shght angular
prominence at each side of the middle. I can give no
information as to the distribution of this species in North
America.
CRIOCEPHALUS PRODUCTUS, Lec.
Gracilis, elongatus, niger, elytris maris dilutioribus ; prothorace
haud transverso, disco inaequali, tri- vel quadri- impresso, ad latera
subangulato ; elytris elongatis, costatis, inter costas crebre perspicue
punctatis. Long. 20-26 mm., lat. 43, 65 mm.
Readily distinguished from C. agrestis by the more
elongate form, the less transverse thorax and the more
developed sculpture of the elytra. The sculpture of the
gular area consists of fine rugosities, and there is not much
beard.
Hab. N. AMERICA.
The species is, I believe, not uncommon in N.W, North
America, I have specimens from Montana (Morrison) ;
Vancouver (Mathew); California (Hepburn).
CRIOCEPHALUS ASPERATUS, Lec.
Latior, robustus, niger, elytris interdum dilutioribus ; prothorace
disco inzquali, ad latera numerose muricato-asperato, lateribus angu-
latim prominalis ; elytris bicostatis, sculptura inter costas obsoleta ;
antennis articulis ultimis quatuor abbreviatis. Long. 21-27 mm.,
lat. 7} mm.
the genus Ciriocephalus, 161
This very distinct species cannot be mistaken for any
other, the structure of the antennz being very peculiar.
The terminal four joints are abruptly shorter than the pre-
ceding, the eighth joint being scarcely half as long as the
sixth. There is not much difference between the sexes.
The male has the legs and antenne rather thicker and the
terminal joints of the latter not quite so abbreviate. The
bush of hair on the gular area is very remarkable in this
species. And the clypeus is considerably larger than in
any other species; in this last character it approaches the
genus Megasemum.
Apparently common in N.W. North America; it ex-
tends at any rate from Montana in the north of the United
States to Guaymas in Northern Mexico (Mathew).
LARV.
We have found larve of at least three species of Longi-
corn Coleoptera in the wood of the Scotch fir-trees in the
New Forest, viz. Rhagiwm bifasciatum, Asemum striatum
and Criocephalus ferus. These three larvee are extremely
similar. The characters of the three forms have been
described by Schiddte, so that it is only necessary here to
give diagnostic characters. Schiddte and the other writers
on the subject have given no information as to the habits
of the three larve, and as these exhibit very striking
differences, Mr. Smith’s notes on this subject will doubtless
be acceptable to a wide circle of entomologists.
The three larve, as already stated, are of three different
genera, Asemum, Criocephalus and Rhagiwm. Mr. Smith
has never met with all three contemporaneously associated,
but Asemum is frequently found in the same stumps as
Rhagium; and in the treesin which Criocephalus has been
found in this country it is also associated with Asemum.
Notwithstanding the extreme resemblance of the three
larvee, they may be readily distinguished by the spines at
the anal extremity of the body. In Rhagiwm there is
only one spine (Plate IX, fig. 9); in Asemum there are
two spines placed near one another*and divergent (Plate
TX, fig. 8); in Criocephalus there are two spines placed
at a little distance from one another (Plate IX, fig. 7)
and parallel, or even at the tips very slightly convergent.
In profile the two Asemum spines are straight; in
Criocephalus they are slightly directed upwards.
TRANS, ENT. SOC. LOND. 1905.—PART I. -(MAY) 11
162 Mr. D. Sharp on
Asemum larva. The description and figures given by
Schiodte (Naturalist. Tidskr. (3) x, Plate xiv, pp. 443) are
correct so far as they go, but he has given of the whole
larva, and in his figure of the head has omitted all details
of sculpture and clothing. We have not been able to find
any other drawing of Asemwm larva, and have therefore
given a figure (Plate IX, fig. 6). Beyond the character
we have already mentioned, drawn from the spines, there
are only very slight marks of distinction from the larva of
Criocephalus ; Asemum larva has the surface slightly more
shining, the upper-surface of the head is conspicuously
shining, its colour is paler, and the whole head is less
robust and powerful than it is in Criocephalus.
Criocephalus ferus larva (Plate IX, fig. 5). Larva of
Criocephalus has been described by Schiddte, and some
of its parts figured by him, t.c., Plate xii, figs. 11-19.
His species he considered to be C. rusticus, and, notwith-
standing the confusion that has existed for so many years
as to C. ferus and C. rusticus, there is little doubt that he
was correct in his determination. The larva of C. ferus,
as found in the New Forest, agrees very closely with
Schiddte’s description of C. rusticus. If actual com-
parison could be made of the two larvee some distine-
tions would probably be found. The head of C. ferus
larva appears to be shorter than Schiddte’s figure, the
part behind the clypeus especially shorter, and destitute
of the peculiar lobe-like marks shown in Schiddte’s
figure.
The Jarvee of Asemum striatum (fig. 6) and of Cricoce-
phalus ferus (fig. 5) resemble one another extremely. C.
jerus larva is much larger when full-grown ; it attains a
leneth of 30 mm. or more, and a width across the thorax
of 9 or 10 mm.; the terminal segments are more ample ;
the mandibles and margin of the head are black; the
chitinized patches—darker in colour—are more definite
and conspicuous; the under-side of the head is rough
and pubescent, and darker in colour than in Asemwm ;
the antennze are shorter. in comparison; the two short
spines placed at a little distance from one another (Plate
IX, fig. 7) at the extremity of the body are, as already
pointed out, diagnostic. The spiracles of the Criocephalus
larva are more elongate (less circular) than they are in
Asemum. ‘There isa good deal of ditference in the way
these two larve distend their segments for the purpose of
the genus Criocephalus. 163
progression in their burrows, and for working at their
excavation.
Rhagium bifasciatum. This larva is abundant in the
New Forest, and unlike the larve of Criocephalus and
Asemum is easily obtained owing to its totally different
habits. The Rhagium lives only in the soft wood that has
commenced to decay.
I will here call attention to a very remarkable variation
that has been detected by Mr. Gahan during the progress
of this memoir. By the courtesy of the Director of the
British Museum (Natural History) I am able to figure it.
I have in the preceding pages formed the group Crioce-
phaline for certain forms that have only one spur on the
front tibia (Plate IX, fig. 14), instead of two as is usual in the
Cerambycidee (Plate IX, fig. 13). In the British Museum
collection Mr. Gahan has detected a female Criocephalus
rusticus having two spurs (Plate IX, fig. 15) on the tibia of
the left front-leg. Unfortunately the right front-leg is
missing in this specimen, but when the individual was first
observed it possessed both anterior legs, and Mr. Gahan
and myself are pretty certain that both legs possessed the
two spurs.
There appears to be nothing else abnormal about this
individual. It may be described in brief as a specimen
that departs from charactr of its species, genus and tribe ;
taking on the character of another tribe. The character
abnormal in this specimen is normal in Cerambycid
Coleoptera generally.
It would be almost useless to discuss the nature of this
variation without further information. Some will see in
it a case of “reversion.” Mr. Punnett has suggested to
me that it is probable that other specimens “similarly
“abnormal” may be found in the locality where this C.
rusticus comes from, and if so the case might be viewed as
one of more continuous heredity.
On the other hand, it must not be forgotten, that two
spurs are normal on the other legs of Criocephalus; so
that it is possible to view the abnormality as a case of
homeeotic variation (Bateson).
Summary of changes and additions made in the pre-
ceding paper :—
Criocephaline, new group separated from Aseminx.
Cephalallus, n, g., C. oberthuri, n. sp., Yunnan,
164
Mr. D. Sharp on the genus Criocephalus.
Megasemwm, defined and male of M. 4-costatwm described.
Cephalocrius, n. g. for Crio. syriacus, Reitter.
Criocephalus pinetorum, Woll., established as a valid species.
Female described for first time.
C. deceptor, coreanus, tibetanus, exoticus, n. spp.
C. ferus, var. hispanicus, n. var.
C. obsoletus, Rand. = (mexicanus, Th.), n. syn.
C. rusticus, description of an aberration.
FIG.
SO) See Nae
12,
EXPLANATION OF PLATE IX.
Under-side of head of Criocephulus ferus.
” 22) nb 2) ” rustics,
Labrum of a ferus.
#4 a 65 rusticus
Larva of Oriocephalus ferus nearly full-grown.
» 3 Asemum striatum, profile.
Spines terminating the body of larva of Criocephalus ferus
seen from above and in profile.
Do. do. Asemum striatum.
Spine terminating body of larva of Rhagiwm seen from
above,
Portion of larva of Criocephalus ferus, showing form and
position of spiracles and the vestigial legs.
One of the legs more highly magnified.
Egg of Criocephalus ferus.
Portion of front-leg of Asemum striatwm 2 showing the
two spurs.
Portion of front-leg of Criocephalus rusticus 2 showing the
single spur.
Portion of front-leg of Criocephalus rusticus @ abnormal
specimen from Amanus mountains, showing the existence
of two spurs as in Asemwm.
Vila. The Habits of Asemum striatum and Criocephalus
ferus.
[Read February Ist, 1905.]
Asemum striatum was, prior to 1902, known in Great
Britain almost exclusively as an inhabitant of Scotch fir
(Pinus sylvestris) forests in Scotland.
Dr. Sharp recorded a specimen from Dumfriesshire in
1879. Messrs Rye and Skinner in recording the capture
of a specimen in 1898,in the New Forest, said—“ We
believe this to be the earliest record of the species from
the South of the Border” (“ Entom. Mo. Mag.” xxx, 1894,
p. 277). Nevertheless we have no doubt that it has been
abundant in the New Forest for a long period of years,
and has only failed to be detected because of its habits.
During May and June in 1902, while staying at Bourne-
mouth, I made several entomological excursions, with the
express purpose of obtaining Longicorn beetles in the
larval and pupal stages. It is well known to be very
difficult to rear these insects, and it may therefore be
well to explain my method.
My articles of equipment for this work included a thin-
edged teaspoon for picking up larvee and pup; a small
tobacco-box fitted up with short lengths of bamboo-cane
of different diameters to serve as artificial burrows for the
pup. The nearer the bamboo sections correspond with
the burrows from which pups are taken, the greater are
the chances of rearing them.
Also a number of cylindrical strong glass bottles with
metal screw caps, each one of which is intended for a separ-
ate larva. The bottles are filled tightly with two or three
pieces of wood and wood refuse taken from the larval
burrows; a suitable cavity is made between the pieces of
wood and the larva is inserted. Care is necessary in
order that the larva is not too tight nor yet too loose.
Once the larva is well established in its new burrow there
is not much risk of its dying. M. Valery Mayet advocates
the keeping of the Longicorn larve in galleries bored by a
large gimlet in solid wood (Bedel, Fauna Col. Seine V,
p. 77). For larvee in their last year or so, however, the
TRANS, ENT. SOC. LOND. 1905.—PART I. (MAY)
166 Mr. F, G. Smith on the Habits of
method I describe is very useful, and with care quite
satisfactory. The chief thing to guard against is empty
cavities which encourage mildew.
I secured and reared at this time several of the more
common species, and at a place near the South Coast I first
made acquaintance with Asemum striatum.
Emerging from a Scotch fir wood which I had been
working for some hours, and feeling tired, I sat down on
one of a great number of stumps of fir-trees which had
been cut down three or four years previously.
The tops of the stumps rose several inches above the
roots. JI immediately noticed a number of distinctly oval
and clean-cut holes on the table-like surface of the stump
and concluded that they were the work of a Longicorn
unknown to me.
While examining these holes I noticed a new hole
actually beginning to form, and through it a mandible
of a beetle appeared. With my knife I enlarged the hole
until the boring itself was laid open to view. The beetle
however had vanished down its burrow. I therefore
brought a small axe into play. I soon found however
that the stump was perfectly sound and almost as hard as
my axe; added to which, the roots radiating in all direc-
tions made it impossible to split off large pieces. How-
ever, after about an hour’s hard work I came in sight of
the beetle, but not of the bottom of its hole, for it seemed
to be able to retreat indefinitely. I at last laid down my
axe and sticking a pin into the end of a thin twig hooked
the beetle out minus two legs. This was my first Asemum
striatum taken from the bottom of its gallery thirteen or
fourteen inches long excavated in solid Scotch fir. This
concluded my day’s work, but I revisited the spot next day,
and after investigation I found that most of the stumps
were covered with holes almost all of which were quite
recent, and I also discovered that if I approached very
cautiously I could see the heads of the beetles at the
mouths of their burrows ; in many cases they were widen-
ing them to the required size of exit. In some stumps I
thus observed as many as twenty beetles at once. But
to secure them was quite a different matter; for as soon
as I touched a stump every beetle disappeared down its
hole not to appear again for an hour or so. I spent
several days carefully investigating the situation; during
which time I was only successful in securing about
Asemum striatum and Criocephalus ferus. 167
half-a-dozen maimed specimens. One stray specimen I
found on the ground among bits of bark. I did not see
one on the wing although there was brilliant sunshine.
During these two or three days I must have seen many
hundreds of the insect, all in their holes. The explanation
of this is that the insect is purely nocturnal in its habits,
taking refuge in its burrows during the daytime when
possible. It would appear however that though nocturnal,
it likes the sunshine, as most Longicorns do. The odd
specimens which subsequent to 1902 have been found at
different places have been stray insects most likely near
very strong colonies, seeing that from the colony I refer
to I only found one straggler. During the last day I
spent at this place at this time I devised a very simple
but effective method of obtaining specimens, and I took
in half-a-day about eighty. It would not, I think, be
fair to the insect to make this method generally known.
Among this number there were two of the var. agrestis,
which is a true variety and not an immaturity, as has been
supposed.
The normal insect, while undergoing the process of
hardening never assumes the brick-red tint of var. agrestis,
but rapidly changes from various hues of dull brown to
pitchy black; some however are not so black as others,
while var. agrestis having once arrived at the brick-red
colour retains that hue. Dr. Sharp has taken a stray
specimen of the var. agrestis on the wing, which proves
its maturity.
The situation I have referred to fulfilled at the time
the conditions necessary for a successful colony, which are
as follows :—
i. That the wood shall be dead or enfeebled, but not
have commenced to decay.
ii, The wood must not have ‘been attacked previously
by Lhagium bifasciatum, as the two are not
friendly, though the existence of Asemwm facili-
tates that of Lhagium.
Probably the insect exists in the same stump (not includ-
ing the thin rootlets) during a maximum period of five
years. My supposition is based on the following :—
Throughout the three years during which the first
brood is maturing, there are no visible punctures on the
168 Mr. F. G. Smith on the Habits of
flat face of the stump, consequently the rain does not pene-
trate into the wood in quantities sufficient to hasten decay.
Moreover, when the tree was felled a coating of resin was
formed on the surface, which prevented the water from
sinking in. For three years therefore the wood was per-
fectly sound and free from superfluous moisture and quite
unsuitable for the accommodation of Asemwm’s greatest
enemy, viz. Lhagiwm bifasciatum.
During these three years successive broods of larvie
entered upon existence.
When at the end of this period the first brood emerged
as perfect insects, the second brood had to live the final
year of their lives under far less favourable conditions
than the first; for every time it rained during that
year, the open burrows of the larvae that had already
developed conducted the water into the centre of the
stump, everywhere spreading decay, which had the effect
of driving some of the larve deeper into the roots while
many perished.
This was proved by the fact that when I revisited this
spot the following year with my friend, Mr. Willoughby
Ellis, although a fortnight later than in the previous
year, there were but few recent though many old
burrows.
The third brood exists under very unfavourable con-
ditions ; for, in the first place, they were tiny one-year
larvee when the rapid disintegration, caused by the open
burrows conveying the rain into the stump, commenced.
In the second place, these unfortunate larvee before they
were much older found that their burrows were repeatedly
crossed with the ever widening ones of their great rival
Rhagium, which species, owing to the decay caused by
repeated rains, has now completely gained the mastery over
the stump, so that the third brood of Asemwm has very
little chance of completing its development.
A few however succeed in finding their ways to the
thin rootlets, and I have found in upturned stumps in the
New Forest, the insect in all its stages ; the rootlets sound
but sometimes no thicker than one’s little finger, the
whole of the decayed part of the stump including the
larger roots being given over to Rhagium.
Since the discovery of this colony I have found a great
number of colonies of Asemuwm scattered all over the New
Forest and in remote corners of Hampshire and Dorset,
Asemum striatum and Criocephalus ferus. 169
and I have always observed that the insect is closely
followed up by Rhagium.
It is thus clear that Asemum striatum and Rhagiwm
bifasciatum together constitute one of the most potent
(though unobtrusive) helps to forestry. For there is good
reason for supposing that there is never a considerable
number of Scotch firs felled in the New Forest but these
two insects turn the opportunity to good account and
speedily reduce the stumps to manure.
The former ever retreats before the progress of decay
until retreat is no longer possible, while the latter delights
in decay and feeds on decaying wood until it thus reduces
the whole stump to powder, with the exception of the
thinnest layer of outer wood which hides its depredation
within. It often happens that a stump, the upper-surface
of which is covered with Asemum holes, and would appear
to contain larve of that imsect, when tapped, completely
collapses and appears but a mass of dust and manure.
Rhagium is not what we may call a hard-working insect
and does not make a hole of exit unless necessary, but
uses any convenient fissure, or the holes of Asemum, for
the purpose of exit.
The life-cycle of Asemwm is as follows, viz. :—
At the end of May or early in June, a number of eggs
are inserted between the scales of the bark on the sides of
the stump or exposed portions of the thick roots. The
young larve soon hatch and for some time feed on the
bark itself. Having worked their way through the bark
they do not bore between the bark and the wood as the
Callidiide do, but penetrate through the outer layer of
wood, when, if they were deposited in the side of the
stump, they commence working downwards, just under
the surface of the wood, making as they proceed galleries,
the calibre of which is in one direction slightly larger
than their own bodies and in the other direction much
wider. These galleries wind about and their margins are
more or less tortuous, The larvee when about half-grown
have proceeded some distance down the stump, and they
then connect the burrows with the outside by means of
small holes. Up to this point, as they progressed, their
galleries were completely and tightly filled behind them
with wood-dust and refuse, which they had compressed
so tightly that when dry it is almost as hard as the sur-
rounding wood. Till this period, the decay of the stump
170 Mr. F. G. Smith on the Habits of
has not been at all hastened by these larvae, for they have
left behind them no communications that would facilitate
the entrance and growth of mildew.
There are no punctures in the bark, and when this is
removed there are generally no traces of borings on the
inside of the bark, nor on the surface of the wood. The
small borings through which the larva entered the wood
itself are filled up and become almost imperceptible.
Owing to this absence of orifices, even though a stump
may contain a great number of Jarvee in this stage, one
not acquainted with their habits would be most likely to
overlook them.
But having made its communication with the exterior
the larva’s efforts have an additional object, for from hence-
forth they are partly directed towards making suitable
provision for itself during its later stages. It for a time
continues to bore downwards but with a more inward
direction, and instead of compressing the refuse behind
it as it previously did, it forces it through the orifice
alluded to.
After pursuing this method for some time it concludes
its downward course and enlarges the end of the burrow
so as to form a sort of turning-space. This may be
anything between eight to fifteen inches below the flat
surface of the stump. The larva now changes its direction
and strikes upwards, with a burrow which has a calibre
somewhat more corresponding with the shape of its body,
and also straighter than its downward burrow.
An upward burrow may at once be distinguished from a
downward one by the fact that the latter is almost imper-
ceptible on account of its being tightly filled with refuse,
while the former is visible as it is free from devs.
When the larva has brought its gallery very close to the
outside of the wood, it discontinues the gallery and pushes
into the dome-shaped termination thereof a number of
long wood-fibres, which it tightly compresses together. If
this be not done to keep out the rain, I cannot suggest
any other purpose.
In this completed and unobstructed upward-burrow, the
larva lives for a considerable time, for | have found com-
pleted burrows in October, while the insect does not
pupate till the following spring.
During this period it gradually ceases its activity and
becomes whiter in colour. Jt pupates in May, remaining
Asemum striatum and Criocephalus ferus. rt
in the pupa state about three weeks. The pupa is very
active. It has no cocoon, but can use the whole of its
upward-burrow (wp and down which it frequently travels)
for the purpose.
The extremity of its abdomen is provided with two long
sharp spines which assist in this operation.
The way it progresses is interesting. It curls the
extremity of its body underneath it, with the result that
one of the spines becomes fixed into the side of the
burrow. It then straightens itself. This operation con-
tinually repeated constitutes its mode of locomotion.
Minute down-curved spines on the dorsal side of the
abdomen prevent it from slipping down its burrow when
stationary. A noteworthy point in reference to these
spines is that they are the only hard and dark parts in
the pupa immediately after its metamorphosis from the
larva.
Apparently its object in travelling up and down its
burrow is to prevent the growth of mildew upon its walls.
I have observed that in glass tubes containing these and
other Longicorn larve the galleries when occupied by
healthy larva or pupa are immune from mildew, while the
other parts of the wood may be badly infested.
It will be recalled that the pupa of the clear-wing
moths are provided with serrations on the margins of their
abdominal plates to assist them in travelling up and down
their burrows in a similar manner.
These spines serve an additional purpose, for they assist
the pupa in the initial stages of casting its skin, by enabling
it to hold its body more rigid and in greater tension, while
splitting open the thin skin at the back of its neck. It
will thus be seen how necessary it is in rearing these and
allied pupee, to put them in tubes which afford them the
same facilities for doing this as their native burrows do.
I have lost more pupze from exhaustion in seeking to
split this thin skin than from any other cause.
After undergoing its final metamorphosis Asemwm
matures rather rapidly.
The imago opens the roof of its prison when almost
mature.
Seeing that their burrows are the only places where one
can hope to find these insects, it seems pretty certain
that when possible they always hide in them during the
daytime, affording as they do such excellent retreats,
172 Mr. F. G. Smith on the Habits of
Asemum sometimes attacks standing trees, but only
when the trees’ vitality has been impaired. In such cases
the upward borings are often much shorter than when the
insect attacks stumps.
There is no fear of Asenvum spreading from sickly trees
to healthy ones, and the greatest inducement for it to
attack is if the trees are burnt round the roots.
CRIOCEPHALUS FERUS.
My investigations in connection with Asemum striatum
led to my exploring the Scotch fir woods of the New
Forest.
In June 1902, in a grove of large trees, I observed a
number of conspicuous holes in the trunks between a foot
and fifteen inches from the ground. They were very like
Asemum holes, distinctly oval and clean-cut, but the
majority were much larger. Although it was raining
heavily I proceeded to investigate the cause. In an hour
or two I was rewarded by securing fifteen or sixteen large
larvee which looked much like Asemwm larvee but which I
clearly saw were different. I supposed that the insect
must be Hylotrupes bajulus. I was therefore much sur-
prised when early in August I reared from them a long
brown insect which certainly was not Hylotrupes. I at
once searched the Birmingham libraries for information, and
found in W. Rothschild’s “ Musée Entomologique Illustré ”
a short description and figure of Criocephalus rusticus, I
concluded that that was my insect, and accordingly so
labelled it in my collection. Towards the end of the same
month another emerged, and the only surviving larva,
showing no sign of pupation, continued to burrow in its
piece of wood. The reason I lost so many larve at this
time was that at the time of capture I had not sufficient
receptacles to enable me to, keep each one separate. The
larvee proved to be very active and irritable, and in con-
sequence before I arrived home all except three had
received injuries.
Early in 1903 I made the acquaintance of Mr.
Willoughby Ellis and informed him that I had taken
Criocephalus in the New Forest, and after seeing my
specimens and the surviving larva, he accompanied me to
the habitat. He subsequently wrote an article on the
subject which appeared in the “Ent. Recd.,” 1903, xv, 259.
Asemum striatum and Criocephalus ferus. 173
In setting forth what I know of the life history of this
insect, with which I subsequently became more fully
acquainted, I will, in order to avoid repetition, mention
chiefly those respects in which it differs from that of
Asemum striatum.
In the first place, it attacks large trees that are standing
and growing. When I discovered its habitat, the trees
appeared in no respect to differ from the surrounding
ones, but we now believe the vitality of these trees was
somewhat impaired by some cause or other,
~The wood in which the larvz were boring was by no
means dead; for when cut, sap freely exuded.
A careful examination revealed the fact that some of
the roots were dead or dying and inhabited by Asemwm.
The holes of Criocephalus are usually further from the
ground than in the case of Asemum, when that insect
attacks trees.
Judging from the length of time that the larva appears
full grown and lives in its completed burrow it would
appear to be much longer-lived than Asemum.
The following, in addition to other evidence I have
gathered, proves that this larva lives for a very considerable
time in its completed burrow, viz. : early in August of last
year I carefully opened two completed burrows which ran
side by side; the unobstructed portion of each was of
similar length, and I did not observe any difference. Each
contained a larva; one of these was about to pupate, while
the other, judging from its sluggish movements, appeared
to be rapidly approaching that condition. 1 concluded
that both would provide imagos last year. The former
however pupated, while the latter simply underwent a
larval ecdysis, and I hope to rear it this year; also the
largest larva I have ever seen I procured at that time,
but it will not mature till next August.
I am of the opinion that Criocephalus while in the larval
state always makes a hole for its exit asan imago and stuffs
up the entrance with gnawed-down bark. I do not con-
sider however that I have sufficient evidence to be quite
positive on this point. It is interesting to note however
that. Criocephalus forces a very much larger quantity of
material into the end of its burrow than Asemwm does,
the stopped-up part sometimes extending right through
the bet and into the wood itself where long wood-fibre
is used,
174 Mr. F. G. Smith on the Ialits of
This wood-fibre is quite different from any refuse found
Jower down its burrow and is only used for this purpose.
When the bark is very thick the larva pupates in the bark
itself, stopping up this portion of its gallery at each end.
When it pupates in the wood it reserves a considerable
portion of its burrow for its final stages and the pupa
travels up and down as that of Asemwm does.
Another interesting point about the burrows of these
larvee, and one which also shows that the larva lives for a
considerable time in that portion of its burrow which may
be termed its pupa-case, is that near the hole of exit the
larva makes a short off-shoot, into which it pushes one of
its discarded skins, and carefully covers over the entrance
to it with wood-dust. The result of this performance is
to place out of reach an object which would encourage the
growth of fungi. In one of my specimens of these pupa-
cases the annex containing the skin is full of mildew and
putrefaction, while the pupa-case itself is quite free from
any such thing.
Another noteworthy fact in the economy of this insect
is the extreme lateness in the season at which the perfect
insect appears. It apparently never undergoes its final
metamorphosis in this country till about the end of July,
although Dr. Sharp reared a specimen in June of last
year. I think there is no doubt however that this was
due to the high temperature of the room it was kept in
and to its proximity to the fire. Having cast its pupal
skin it is at the very least a fortnight before it is ready to
leave its hiding-place. In large, well-developed specimens,
it takes quite a considerable time for the distended abdo-
men to sufficiently contract not to be an impediment to
the insect and to come within the limits of the elytra.
Having matured, it leaves its prison as the twilight is
giving place to darkness.
It has been thought that it is a sluggish insect, but so
far is that from the case that I believe it to be second to
none of the British Longicorns for fleetness of foot and
agility of wing. There seems no difference between the
sexes in this respect. They both continue to fly and run
about at a great rate late into the night.
I kept several alive last year and they all died about
one month after maturing. They came from their hiding-
places regularly at the same time every night, and once
or twice I looked at them after 12 o'clock, and found
Asemum striatum and Criocephalus ferus. Wig
them still active. I succeeded in breeding with one pair,
and have at present some of their offspring in a piece of
bark.
I have on two or three different occasions observed them
flying about in their natural habitat, and noted that the
time at which they appeared was the same as in the case
of specimens in captivity.
The eggs look exceedingly like those of the common
blow-fly and are thrust deep down between the laming of
the bark to which they are cemented. A female does not
deposit many more than fifty or sixty eggs.
During the daytime the insect is perfectly inert, almost
to the point of feigning death. It may be said to be at
large from the middle of August till the middle of
September, although I have specimens which did not cast
their pupal skins till September.
I consider that the following considerations are re-
sponsible for this insect not having been observed
previously :—
i. The comparative rarity of suitable habitats.
i. The great difficulty in securing the perfect insect
(though the habitat may be known) on account
of its nocturnal habits and its great resemblance
to the bark among which it hides.
i. The difficulty in securing the larve owing to the
very hard wood into which they bore.
iv. The unusual lateness in the season at which it
appears.
v. Its probable confinement to one spet for a great
number of years.
vi. The fact that pine woods in S. England have till
lately been almost entirely ignored as unproductive
by Coleopterists.
Most of the above considerations are doubtless respon-
sible for the fact that Asemuwm up till 1902 was almost
unknown as an inhabitant of Great Britain south of the
Border. But as it does not appear ‘‘out of season” as
Criocephalus does, and is so very much more common than
that insect, it is indeed remarkable that it has not been
better known by British collectors before.
Before bringing these notes on the habits of these two
interesting insects to a close I must take this opportunity
176 The Habits of A. striatum and C. ferus.
of expressing my great indebtedness to Dr. Sharp, who
has put every possible facility in my way to help me to
complete my knowledge of these insects, and apart from
whose generous assistance it would have been impossible
for me to have contributed this very small item to the
stock of entomological knowledge.
My thanks are also due to Mr. Ellis, Mr. Colbran J.
Wainwright, and Mrs. Lamb of Brockenhurst, for help I
have received in various ways,
VIII. On the matrivorous habit of the species of HETERO-
GYNIS, Rmbr. By T. A. CHAapman, M.D., F.ZS.
[Read February Ist, 1905.]
IN presenting these notes in extension of my previous
papers on Heterogynis, Rmbr., the regret, I must acknow-
ledge, in having to correct an error I had fallen into but
slightly modifies the satisfaction I have in having at
length made certain observations, which I ought no doubt
to have made before, and in fact narrowly escaped making.
The satisfaction results not so much from the somewhat ~
remarkable nature of the observations themselves, as from
the circumstance that they appear to explain the object to
be attained by, and therefore the forces in action that
evolved, the very anomalous specialization of the female
imago in this genus, and relieves us from having to regard
them as isolated and inexplicable phenomena.
In my paper on H. paradowa in the Transactions of the
Society, 1902, pp. 717-718, I said that the young larvee of
that species “hibernate by spinning a small cocoon in
some crevice of the food-plant or elsewhere. H. paradoxa
does this, I find, in the second instar; Mr. Fletcher found
Hf, penella did so in the third. Whether there is here a
real specific distinction I cannot say, or whether there may
be an error of observation on my part, or Mr. Fletcher’s.
The newly-hatched larva of H. penella is certainly much
smaller than that of H. paradoxa.” Again, p. 726, I wrote,
“The newly-hatched larvae present very important differ-
ences that have perhaps more specific value than any
others.” I then proceed to describe certain differences in
the possession of stellate hairs by paradoxa which are not
to be found in H. penella until the second instar.
This year I met with H. paradoxa at La Granja; I was
too late for larve or moths, but found cocoons, from some
of which the larve had already hatched, from others of
which they emerged after I took them. Perchance the
want of the more interesting stages made me attend more
closely to the material I had. The result of my observa-
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 12
178 Dr. T. A. Chapman on the
tions is that the passages I have quoted above from my
previous paper on the species, are quite correct, if, by
“hatching,” one means the emergence of the young larvee
from the pupa-case and cocoon of their mother. But as
the true meaning of the word is emergence from the egg
they are all wrong. Hine ille lachrymx, though I rejoice
more over the correction than I weep over the error.
The actual facts are that in all the species the newly-
Fie. 1.
hatched larve are very similar; in all, their first pious
duty is to eat the remains of their mother.
Having done this, H. penella and H. canalensis perforate
the maternal pupa-case with various openings and make
their escape. An examination of the empty pupa-case
they have left shows it quite clean and containing only a
very few threads of silk entangling a small but varying
number of small greyish pellets, which I take to be urates
or some similar effete product of the dead mother. In
H., paradoaa, for some reason, the procedure is different,
probably because the mother is much larger than in the
Matrivorous habit of the species of Heterogynis. 179
other species, and the difference in size is largely expressed
not in the size or number of her eggs, but in the mass of
eatable materials she represents after her decease. At
any rate the young larvee get larger than do those of the
other species, and undergo their first moult before penetrat-
ing the pupa-case and cocoon and venturing to appear at
large.
A pupa-case of H. paradoxa §, from which the young
Fic. 2.
larvee have emerged, presents a dirty and muddled aspect
due to the presence of a good deal of spinning by the
young larve, amongst which the skins they cast at their
first moult are entangled.
A very interesting point is that there is no trace of
larval frass present in either species. This one can more
easily understand perhaps in the case of H. penella, where
the total amount of maternal tissue is small, and each larva
gets but little, as evidenced by their small size on emerging
from the parent cocoon. But in the case of H. paradoxa
the amount must be considerable, as the larvae grow very
180 Dr. T. A. Chapman on the
appreciably, in fact about double their size, besides going
through a whole instar and making a moult.
I am afraid that in my previous observations, made in
the first place hurriedly in the field, and not afterwards
properly verified, I mistook the heads of the cast skins for
larval frass, and assumed some such amount of frass to be
what one would reasonably expect after their first meal of
the maternal tissues. In H. paradoxa one finds, however,
nothing but some threads, the cast larval skins (skins as
well as heads), and the pellets of urates, which from their
small number and comparatively large size are obviously
maternal and not larval débris. The young larve there-
fore go through a whole instar, more than double their
size, undergo a moult, and then perforate the pupa skin,
and make their way through the cocoon without ejecting
any frass or effete material. It seems to follow that the
maternal remains must consist almost entirely of com-
pletely assimilable materials. As the larve on facing the
world have to begin life in many cases by making a long
journey to find their food-plant, a meal before starting is
an obvious advantage, but this does not explain the
curious details of the process, or why it should differ in the
several species.
I should note that I have examined just now pupz of
paradoxa and canalensis, but have no adequate specimen
of penella by me. With regard to it, therefore, I merely
assume from my previous observations of it that it agrees
with canalensis.
It results that at corresponding instars the larvee of the
several species are very much alike, it also follows that
the discrepancy between Mr. Fletcher’s observations as
to the hibernating instar of H. penella and mine of
H. paradoxa does not exist, though it is my fault that
there appeared to be one.
The distinction I drew between the first instar larvee of
H. penella and H. paradoxa does not exist as a structural
difference at all ; nevertheless there is a difference between
them in habits and instincts, which is probably of quite as
great importance.
The only other matter I observed this year at La Granja
as regards H. paradoxa was that it had two widely-separated
habitats, one at about 5200 ft. elevation, where its food-
plant was <Adenocarpus hispanica, the other at about
7000 ft. on Cytisus purgans; the opportunity was wanting
Matrworous habit of the species of Heterogynis, 181
to determine whether there was any varietal distinction
between the two races, but this is very probable, as the
intermediate country and elevations were without any
“broom” capable of nourishing the species, so that the
two colonies were probably segregated with considerable
strictness.
The extreme development of the matrivorous habit in
H. paradoxa, and its obviously great importance in the
economy of the species, afford an explanation of the
remarkably aberrant habits and structure of the female
imago, and give us some hints as to the probable steps by
which they were evolved.
Though the latter are more or less hypothetical, and
therefore less certain than the former, we may take them
first, as they are so in point of time.
We may safely assume that the first steps in the process
of evolution were similar to those that obviously obtained
in Psyche and Orgyia. Firstly, apterousness of the 2 con-
sequent on laying the eggs, on or in the cocoon or close by.
Then eggs laid in the pupa-case. And somewhere at this
point the dominating conditions would probably be those
which I imagined still to obtain, as no doubt subsidiarily
they do, when I discussed this matter in connection with
Hf. penella (Ent. Trans., 1898, p. 46), viz. protection of the
eggs from enemies and from desiccation by the mother
dying in and blocking the open end of the pupa-case. At
this time the moth probably still retained some legs and
some scales or hairs. Both Psychids and Orgyias still
retain some hairs, so far as I know, in all cases.
The delicate nature of the eggs (as in the allied
Anthroceras) would make protection against drought a
strong selective agency, and there would now come in the
matrivorous instincts of the larva. These no doubt would
originate accidentally in the necessities of the larvze finding
a way of escape from the pupa-case, leading some of them
to do so by eating into, or at least nibbling the dead body
of the 2. So soon as matrivoracity became definitely an
instinct then several forces would come into action. As
these would be the same that now keep the arrangements
in their present perfection we may better consider them in
connection with the explanation they afford of existing
facts. The very complete closure of the pupa-case, so that
it looks as if it had never been opened, and no moth ever
emerged from it, becomes very important when we regard
182 Dr. T. A. Chapman on the
the body of the moth no longer as a second line of defence,
but as a store of food to be kept from enemies gross or
bacterial, from drying up, and from other dangers. The
precautions for the moth getting safely back into the
pupa-case, apparently rather a hopeless matter considering
its maggot-like structure, become more obviously matters
of necessity, leading up to the organic connection the moth
has with the pupa-case at the sites of the atrophied legs.
The brief time the moth remains out of the pupa-case, less
probably sometimes than five minutes, is not only import-
ant as minimizing the period of exposure to enemies, but
also as a period of deterioration of the moth as a food
material.
We next come to the extraordinary structure (or want
of structure) of the moth itself. Everything aims at the
whole available forces and materials possessed by the larva
when it spins its cocoon being devoted to egg formation on
one hand and larval food on the other, and further that the
last object hardly takes a second place. If we compare
the female Heterogynis with those females of the Psychids
in which the structures have most degenerated, we find
that in the Psychids everything has given way to egg
development. The protection of the eggs is achieved
chiefly by mixing them with the hairs from the maternal
surface, and the female drops out of the case after she has
laid her eggs as a mere sciap of chitin, with considerable
masses of urate sand some little fluid. There is, in fact,
nothing whatever edible. In Heterogynis no hairs are
used to protect the eggs, and not only is the 2 devoid of
hairs and scales, but the cuticle is a simple membrane
without traces of the bases of hairs or scales, without any
skin points or other structures, and if I said actually
without chitin, I think I should commit no large error
demonstrable in a chemical balance. In the next place,
the urates are very small inamount. The quantity which
most insects void on emerging from the pupa is very
considerable, partly left in the pupa-case, partly voided
after the wings are expanded; it is usually in solid
particles suspended in fluid; there is usually some excess
tluid to be got rid of after the wings are expanded. Why
are these urates less in the case of Heterogynis? If 1 am
asked are they really so, I must admit that I am_ not
prepared to meet the question, as having weighed the
material in question. But the thing seems to me to be
Matrivorous habit of the spectes of Heterogynis. 183
so obvious, comparing the few pellets remaining from
Fleterogynis, with the deposits usual in most moths of
similar size, that the matter is self-evident. The urates
represent the amount of tissue waste that has taken place.
Now in Heterogynis this tissue waste is that involved in
the muscular exertion of spinning the cocoon, in casting
the larva skin, and in the emergence and retreat of the
moth, and in laying her eggs. The whole process of
histolysis and development by which the larval structures
are absorbed, and the imaginal (wings, legs, hairs, scales,
antenne, etc.) are built up from their embryonal condition
is completely saved ; this process must in ordinary Lepido-
ptera require a good deal of expenditure of tissue material.
In Heterogynis not only are no imaginal structures de-
veloped, but the larval skin muscles and the larval
colouring remain unchanged. I have already noted the
skin to be devoid of the usual skin points, but there are in
fact no hard parts whatever, no head plate, no prothoracic
plate, no appendages, no solid parts to the ovipositor, etc.
Everything is eatable, and all is eaten; I do not think we
can find in the imaginal composition any explanation of
why the larvee void no excreta during a whole instar, and
until, during the second instar, they have made their
escape from the parental cocoon.
This is probably a matter of hygiene, by which the
presence of such excreta would be most undesirable
amongst the crowded larve, especially if their emergence
were delayed by any climatic or other causes. The case of
such larvee as those of Cnethocampa, whose nests are loaded
with frass, are hardly parallel, since these nests are very
roomy, and the portion into which the larvee crowd are
more or less free from frass, whilst in the Heterogynis
pupa-case the larvee are solidly wedged together with no
spare space and very few threads of silk. That there shall
be no trass, however, that the larvz shall not require to
void anything, it is no doubt necessary that the pabulum
shall be of a most concentrated and digestible nature.
Summarizing the facts now advanced, there is first the
correction of the error as to first stage of H. paradoxa,
due to the recognition of the circumstance that it does
not emerge from the maternal pupa-case and cocoon till it
has moulted into the second instar. Secondly, the import-
ance in the economy of the genus, and especially in
H. paradoxa, of the matrivorous habit, all the details
184 Dr. T. A, Chapman on Heterogynis,
connected with which are elaborated to a degree, and of
which the remarkable structure of the female and her way
of life, are items which it very largely explains. I cannot
call to mind any other Lepidopteron with such a matrivorous
habit.
DESCRIPTION OF FIGURES.
Pupa-cases of H. canalensis (fig. 1) and H. paradoxa
(fig. 2) longitudinally divided and placed on _ slides—all
contents preserved. Photographs by A. E. Tonge, Esq.
The amplification is four diameters,
The H. canalensis pupa has no contents, but a few grey
pellets of maternal urates.
The H. parodoxa has similar pellets, but also contains
larval skins, of which the heads are very conspicuous, cast
on their first moult by the young larve. There is no
larval frass. There is in neither case any trace of the
mother except the pellets of urates.
The more solid abdominal ends of the cases are split
irregularly,
185
IX. Descriptions of some new species of Satyride from
South America. By FREDERICK Du CANE Gop-
MAN, D.C.L., F.R.S., ete.
[Read March Ist, 1905.]
PLATE X,
I TAKE the opportunity of describing the following
apparently new species of South American Satyride, which
have been for some time unnamed in my collection, before
handing them over to the Natural History Museum.
Pierella chalybea, sp. n.
Very similar to P. lamia, Sulz., but differs in having rather more
than the basal half of the hind-wings in both sexes suffused with
steely-blue, the black rings round the submarginal row of white
spots is almost or quite absent, the outer margin of these wings is
very broadly infuscate, and the two inner lines crossing the disc are
faint,
Hab. Brazit, Chapada (H. H. Smith).
I have two males and one female of this Pierel/a, which
seems sufficiently distinct to require a separate name.
The steely-blue suffusion of the basal area of the hind-
wings is strongly marked in both sexes. This character
also distinguishes it from P. rhea, Fabr.
Euptychia analis,sp.n. (Plate X, fig. 1, f.)
dg. Very like EF. batesi, Butl. ; above faintly suffused with purple
and the hind-wings with a conspicuous patch of dark blue scales at
the anal angle, the two reddish-brown bands beneath broad and
visible from above.
Hab. AMAZONS, San Paulo and Tapajos (Bates); PERu,
Pebas (Hawksley).
Three males. This appears to be a form of H#. batesi,
which also occurs at Tapajos, differing from it as
indicated above.
TRANS. ENT. SOC, LOND. 1905.—PART I. (MAY)
186 Mr. F. Du Cane Godman on Descriptions of
Euptychia stigmatica, sp. n. (Plate X, fig. 2, f.)
¢. Very similar to £. cxrilea, Butl., but paler blue, the primaries
with an elongated brand about the middle of the inner margin
composed of intermixed white and reddish-brown scales, the broad
blackish border less sharply defined, the transverse blackish streak
beyond the cell distinct ; beneath bluish-grey, the lines crossing
the wings broader.
Hab. ARGENTINA, Entre Rios (HZ. H. Smith).
One specimen. This insect in size, colour, and marking
resembles the female of /. cvrulea, but wants the inner
submarginal line on the secondaries and the line crossing
the cell on the primaries. The male has a very con-
spicuous elongate brand at the middle of the inner margin
of the primaries, a character not found in £. cxrulea.
There is also a male of this species in the Hewitson
Collection at the British Museum; it is labelled “ Rio
River.” A female placed with it in the same collection
no doubt belongs here.
Huptychia penicillata, sp.n. (Plate X, figs. 3, 3a, $; 4, 2.)
¢. Greenish-plumbeous, the primaries with a dense pencil of long
coarse brown hairs below the origin of the first branch of the median
nervure ; beneath greenish-brown, crossed by two common dark
brown lines, and with the usual wavy submarginal line of the same
colour, the primaries with three small ocelli, the subapical one
conspicuous, the others indistinct ; the secondaries with a series of
five ocelli, the second and last large and the others small.
@. Uniform brown above ; beneath as in the male, but the
ground-colour pale brown, becoming grevish towards the outer
margin.
Hab, AMAZONS, Tapajos (Bates).
One pair, which I have had for many years unnamed in
my collection. The male has a tuft of hairs on the fore-
wings as in the same sex of £. pilata, Butl., the hairs
themselves being coarser than in 4. wmbrosa.
Luptychia scopulata, sp. n. (Plate X, figs. 5, 5a, f.)
4. Allied to E. penicillata and having a similar tuft of hairs on
the fore-wings, but of a browner hue above ; beneath greyish, the
two common lines crossing the wings broader and of a red-brown
colour, the primaries with two ocelli only.
some new species of Satyride from South America. 187
Hab. UPPER AMAZONS.
Dr. Staudinger has sent me a single male example of
this insect, which for some time has been placed with the
preceding species in my collection. Although very like
LE. penicillata it differs considerably on the under-side.
Euptychia mimas, sp.n. (Plate X, fig. 6, 3.)
¢. Uniform dark-brown ; beneath paler, crossed by two dark-brown
lines and with a narrow, wavy, similarly coloured submarginal line,
the primaries with a series of two or three small ocelli, that nearest
the apex the most distinct, the secondaries dusted with ochreous
seales and with a series of five bipupillate ocelli, the second and fifth
black within.
Hab. Boutvia, Coroico, 6500 ft. (Gar 7g
Three examples. Near /. phineus, Butl., from Venezuela.
Euptychia boliviana, sp.n. (Plate X, fig. 7, 2.)
¢. Bright-brown ; beneath paler, primaries with a broad ochra-
ceous band extending from the anal angle and becoming less distinct
towards the apex, a wavy submarginal dark line, and two or three
very minute white spots towards the apex ; secondaries, except at the
base and outer margin, thickly covered with whitish scales, crossed
by two irregular ochraceous submarginal bands, three strongly
angulate dark lines (two crossing the disc, the third submarginal),
and a series of minute white dots.
Hab. Bouivia, San Jacinto, 6000-8000 ft. (Garlepp).
Two specimens. This is a species of large size, uniform
bright brown above, and peculiarly marked beneath.
Euptychia (2) biocellata, sp.n. (Plate X, fig. 8, %.)
¢. Uniform brown, the primaries with a double black ocellus to-
wards the apex enclosed in an ochreous ring; beneath paler, the
ocellus more conspicuous aud surrounded by a dark line, which
becomes faint at the apex and forms an angle towards the inner
margin, the space between it and the outer margin marked with grey ;
secondaries with two broad grey bands crossing the wing, one just
beyond the cell (bordered internally by a dark tiney the other close
to the margin and extending along the inner margin to the base, the
cell also crossed by a faint dark line about the middle.
Hab. Boutvia, Tanampaya, 6000-8000 ft. (Garlepp).
Three specimens of this small species have been sent
me by Dr. Staudinger. Jt will perhaps prove not really
188 Mr. F. Du Cane Godman on Descriptions of
to belong to Huptychia. There are, however, several
members of that genus similarly coloured on the upper-
side, but the present insect differs from all of them in the
marking of the under-side of the secondaries.
Lymanopoda maso, sp. n. (Plate X, fig. 9, 3.)
¢. Uniform brown ; beneath as above, the primaries at the apex
and the secondaries towards the inner margin dusted with lilac
scales, the primaries with a submarginal series of minute white dots,
the secondaries with a curved series of six or seven whitish pupillated
spots running from the apex to the anal angle. Primaries truncate
at the apex, secondaries dentate on the outer margin.
Hab. Cotomsta, Frontino in Antioquia (7. K. Salmon).
Four specimens. The shape of the wings and the
conspicuous row of white spots on the secondaries beneath
readily distinguish this insect. L. albomaculata, Hew.
(= apulia, Hopff.), has a somewhat similar row of white
spots on the under-side of the secondaries, but in that
insect the wings are differently shaped.
Lymanopoda malia, sp. n.
Very similar to LZ. acreida, Butl., but the secondaries also with a
conspicuous submarginal row of fulvous spots.
Hab. PERv, Rio Colorado (Watkins and Tomlinson),
Pozuzo (Pierce).
This is a form of Z. acrvida, Butl., but as it appears to
be restricted to Peru I have ventured to give it a name.
LI. malia is not uncommon, though hitherto I have not
seen a female of it. The Ecuador specimens of L. acrexida
in my collection are much smaller than those from Bolivia,
though they do not otherwise differ.
Lymanopoda levana, sp.n. (Plate X, fig. 10, ¢.)
¢. Dark brown, primaries and secondaries each with a small
pupillated black spot near the anal angle, and one or two minute
white dots above it parallel with the outer margin; beneath,
primaries as above, but with the apex broadly ferruginous, second-
aries ferruginous, an ochreous band (outwardly dentate, inwardly
evanescent) crossing the wing from about the middle of the costa to
the anal angle, the dots as above.
Hab. Cotomsta, Bogota (Chapman and Wheeler).
some new species of Satyride from South America. 189
Two specimens. ‘This is a small species, with somewhat
pointed primaries, very dark brown above, and with the
secondaries peculiarly marked beneath.
Pedaliodes albonotata, sp. n. (Plate X, fig. 11, 2.)
g. Blackish-brown, primaries with a transverse quadrate white
patch crossing the cell obliquely, two spots beyond this (one towards
the apex, the other towards the anal angle) also white; beneath as
above, the apex of the primaries and the whole of the secondaries
mottled with white, the two outer spots on the primaries as above,
but smaller, the secondaries with an irregular transverse \W-shaped
mark at the apex, a spot about the middle of the costa and another
near the anal angle also white. The cilia of the primaries spotted
with white.
Hab. VENEZUELA, Merida (ex Staudinger).
Two males of this well-marked species were sent me
some years ago by Dr. Staudinger. One of the specimens
has, on the upper-side,a minute white dot between the
two submarginal spots on the primaries and another near
the apex of the secondaries. There are also two examples
of P. albonotata in British Museum, both from Venezuela,
one from Culata, the other from Pedregosa; these have
the two submarginal spots on the upper-side of the
primaries almost or quite obsolete.
Oxeoschistus duplex, sp.n. (Plate X, fig. 12, 2.)
¢. Dark-brown, an ochreous band running from near the apex of
the primaries and widening till it reaches the anal angle of the
secondaries, the band marked with a series of large rounded dark
brown spots, its outer margin strongly dentate on the second-
aries, thus forming lunules, which are bordered externally
with ochreous; beneath, primaries paler than above, ferruginous
at the apex, the costa towards the tip streaked with white and
yellow, the outer margin also faintly spotted with yellow ;
secondaries obscure ferruginous, a transverse white line from the
costa crossing the middle of the cell and extending to near the inner
margin, the submarginal band darker than above and also bordered
on both sides with white, the black spots pupillate.
Hab, Boutvia, San Jacinto (Garlepp).
The single example I possess of this species was sent
me long ago by Dr. Staudinger under the MS. name of
190
O. duplex. It is a close ally of O. puerta, Westw., from
Costa Rica, Colombia and Venezuela, but differs from it
chiefly in the very strongly dentate outer margin of the
band on the secondaries, and in the large size of the spots
on both wings. There are also two males of this insect
Explanation of Plate.
in the British Museum, both from Bolivia.
EXPLANATION OF PLATE X,
Fie. 1. Euptychia analis, ¢.
2: A
os A
3a. A
4, 35
5. ”
da. aS
6. 4)
ie ‘5
8.
stigmatica, 3.
penwcillata, g.
mi tuft of hairs on the fore-wing, ¢.
” 2:
scopulata, 3.
. tuft of hairs on the fore-wing, ¢.
mimas, ¢.
boliviana, ¢.
(?) biocellata, 3g.
9. Lymanopoda maso, ¢.
levanu, 3.
11. Pedaliodes albonotata, ¢.
12. Oxeoschistus duplex, g.
(, Tees)
X. Additions to a knowledge of the Homopterous Family
Cicadide. By W. L. Disranr.
[Read March 1st, 1905.]
PuatTE XI.
BEING engaged in an attempt to revise the classification
of the Cicadide and to prepare a synonymic catalogue of
the Family, I have been entrusted with much material
belonging to other collections in order that my purview
may be as extensive as possible. IJ have thus come across
a number of undescribed species, and this paper refers
only to those belonging to the subfamily Cicadine. Of
these I found in the collection belonging to the Paris
Museum, some beautiful species from Madagascar, which
are probably among the finest in the whole family
Cicadide. A striking species of Pycna from Natal, is from
the Stockholm Museum.
Family CICADID A.
Subfamily CICADIN_ 2.
Div. POLYNEURARIA.
Sadaka hyalina, sp. n.
2. Body brownish-ochraceous ; head with a large central spot, an
anterior transverse line and linear margins to front, and a somewhat
broken transverse fascia between eyes, black ; pronotum with the
lateral margins, a central longitudinal line widened anteriorily, a
central transverse spot in front of anterior margin, and the incisures,
black ; mesonotum with four obconical spots, the two central smallest,
a central lanceolate spot and a small spot in front of each anterior
angle of the cruciform elevation, black ; basal segmental abdominal
margins black ; the central sulcation to face, a fascia between
face and eyes, and apex of rostrum black; tegmina and wings
hyaline, venation ochraceous, inclining to fuscous outwardly ;
tegmina with the costal area and membrane ochraceous, the last
containing two piceous spots; upper half of basal cell, and
basal half of claval area piceous. Head including eyes about equal
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY)
192 Mr. W. L. Distant’s Additions to a
in width to base of mesonotum; head as long as pronotum exclud-
ing posterior margin, lateral pronotal margins angularly ampliate ;
rostrum reaching the posterior coxe.
Long. excl. tegm. 2 18 millim. Exp. tegm. 58 millim.
Hab. W. Arrica; Guinée Frangais (Paris Mus.).
Munza revoili, sp.n. (Plate XI, fig. 5, a, b.)
¢g. Head above piceous-black ; a spot at base of antenne, eyes,
ocelli, and a large transverse spot between area of ocelli and eyes,
ochraceous; pronotum ochraceous, the margins flavescent, a pos-
teriorily broad central triangular fascia to disk, the incisures, and
sometimes extreme lateral margins, piceous-black ; mesonotum
testaceous, with four large obconical black spots, the two intermediate
spots smallest ; abdomen above brownish-testaceous ; body beneath
and legs more or less testaceous ; an anterior black fascia between
face and eyes ; anterior tibiew piceous ; tegmina with about basal half
creainy opaque, with obscure pale fuscous spots or mottlings, basal
cell with its anterior and apical margins piceous, apical half of
tegmina subhyaline, tale-like ; a macula fascia exten ling across the
bases of 1-5 apical areas, a similar fascia at apices of apical areas,
broken on lower apical area, and more or less extending to outer
margin ; wings umber-brown and outwardly piceous for about
two-thirds their breadth from base, the anal area dark fuscous, the
venation piceous. Head including eyes slightly wider than base
of mesonotum, the lateral pronotal margins broad and medially
angulate ; abdomen much narrowed posteriorly ; rostrum reaching
the posterior coxe; opercula transverse, centrally overlapping,
margins moderately convex, just reaching base of abdomen.
Long. excl. tegm. ¢ 26 millim. Exp. tegm. 77 millim.
Hab, SOMALILAND; Ouarsangueli (Revoil—Paris Mus.).
Platypleura spicata, sp. n.
Q. Body brownish-ochraceous, thickly and longly greyishly pilose ;
head with the area of the ocelli, two transverse lines on the lateral
margins of vertex, and an anterior marginal line to front, black ;
pronotum with two central lines fused posteriorly but not extending
across the posterior margin, and the incisures, black ; mesonotum
with four medial obconical spots on anterior margin, an elongate
lateral spot on each side, and a spot at each anterior angle of the
cruciform elevation, black ; tegmina and wings pale hyaline, a little
tale-like ; tegmina with a spot on costal membrane and area near
knowledge of the Homopterous Family Cicadide, 198
base, a small spot just beyond basal cell, some short shadings on the
longitudinal veins to ulnar and apical areas and the same on the
transverse veins at bases of most of the apical areas black. Head
including eyes about as wide as base of mesonotum ; the lateral
margins of the pronotum strongly angularly amplhated, their apices
acutely spinous and a little anteriorily recurved ; the greyish
pilosity particularly developed on the mesonotum and abdomen, on
the last forming distinct long segmental margins ; the face is more
or less piceous, very strongly longitudinally sulcated, its lateral areas
transversely ridged ; rostrum reaching the posterior coxa, its apex
piceous; intermediate and posterior femora longitudinally and
linearly streaked with fuscous beneath ; anterior femora obtusely
spined beneath.
Long, excl. tegm. 9 22 millim. Exp. tegm. 76 millim.
Hab. MapacascaAR; Majunga and Pays Mahafaly
(Paris Mus.).
This species is allied to P. polydorus, Walk., of S. and
E. Africa, from which, apart from other characters, it 1s at
once separated by the largely spinous character of the
lateral pronotal margins.
Platypleura seraphina, sp. n.
9. Head, pronotum, and mesonotum dull ochraceous ; head with
two transverse black fascie, the first extending through area of ocelli .
to lateral margin in front of eyes, the other one connected with it,
a little more broken and passing through base of front ; pronotum
with the lateral and posterior marginal areas stramineous, a
central longitudinal fascia connected with an anterior transverse
curved line and the incisures black ; mesonotum with two central
anterior obconical spots, with a much longer fasciate spot on each
side of them crossing the whole of the lateral area, and a triangulate
linear spot in front of the cruciform elevation, black; abdomen
above pale castaneous with transverse segmental black fascize which
are mostly broken on disk ; body beneath and legs dull ochraceous,
sternum greyishly tomentose, lateral margins and striations to face,
and a fascia between face and eyes piceous or black. Tegmina
greyishly subopaque, the veins reddish-ochraceous, with pale fuscous
mottlings and shadings, a curved fascia crossing tegmina through
radial area, more or less connected with a large faciate spot which
more than occupies the whole of seventh, apical area, a transverse
spot at end of radial area, the transverse veins at bases of apical area
9-5, a double series of spots at apices of longitudinal veins to apical
areas, and basal cell, piceous; wings reddish-ochraceous ; a basal
TRANS. ENT. SOC. LOND. 1905.—PART I. (MAY) 13
194. Mr. W. L. Distant’s Additions to a
costal spot, margins to the abdominal areas, and the outer margin—
very wide at apex, piceous; marginal membrane pale hyaline.
Head including eyes about as wide as base of mesonotum ; pro-
notal lateral margins ampliate and rounded, obsoletely centrally
angulate ; anterior femora in ? practically unarmed.
Long. excl. tegm. @ 18 millim. Exp. tegm. 54 millim.
Hab. Frencu Guinea; Rég. de Kouroussa (H. Por-
beguin—Paris Mus.).
Allied to P. severini, Dist.
Platyplewra esa, sp. n.
Head ochraceous, base and anterior margins of front, a spot above
base of antennze, a broad transverse fascia between eyes, and margins
of eyes connected with the transverse fascia near area of ocelli,
black ; pronotum with the posterior margin stramineous, the disk
with a central fascia subtriangularly ampliated anteriorly, less
ampliated posteriorly, the incisures, and sub-lateral margins (broadly)
black ; mesonotum black, with the margins of four obconical spots
—the two central shortest—ochraceous ; cruciform elevation ochra-
ceous; abdomen black ; segmental margins ‘castaneous, tympanal
flaps stramineous ; body beneath ochraceous, sternum greyishly
tomentose ; anterior margin and central sulcation to face, a fascia
between face and eyes, basal areas of opercula, abdominal segmental
margins, and under-surface of anterior tibia black ; tegmina and
wings pale hyaline ; tegmina, with more than half of basal venation
ochraceous, remainder black; basal cell and dividing line between
costal membrane and area, black. Head including eyes slightly wider
than base of mesonotum ; lateral marginal areas of pronotum moder-
ately ampliate, but the margins almost straightly truncate, not
angulate ; opercula in male broad, reaching the second abdominal
segment, their apical margins obliquely convex, their inner margins
overlapping ; rostrum reaching the posterior coxe.
oD?
Long. excl. tegm. ¢ and 9 22-24 millim. Exp. tegm. 65 millim.
Hab. SOMALILAND ; Ouarsangueli and Obok (Paris Mus.).
Yanga bouviert, sp.n. (Plate XI, fig. 2 a, 0.)
d. Head, pronotum, and mesonotum greenish or brownish-
ochraceous ; front of head with anterior marginal lines and a
central spot widened and angulated posteriorly, black ; vertex with
the area of the ocelli, two transverse fasciz on its lateral area, and a
linear longitudinal spot on each side of disk, black ; pronotum with
the anterior margin, the incisures, and two central lines neither
knowledge of the Homopterous Family Cicadide. 195
reaching the anterior margin nor extending to the posterior marginal
area, black ; mesonotum with two short obconical spots on anterior
margin, a discal triangular spot and a transverse series of four small
spots across posterior area, black ; a long, pale castaneous obconical
spot on each lateral area which contains two small black spots at its
base and one at its apex ; abdomen above black, with a central basal
longitudinal ochraceous area, and the segmental margins more or less
greyishly pilose ; body beneath and legs greenish-ochraceous, the
sternum greyishly tomentose, the opercula palely piceous and
greyishly tomentose, their margins dull ochraceous; tegmina brown-
ish-ochraceous, costal membrane and area more or less greenish, radial
area with three pale spots, some obscure paler spots and mottlings
in ulnar and apical areas and alternate darker and paler outer
marginal spots; wings pale umber-brown, basal third blackish-
castaneous, a streak of the same colour both in and beneath radial
area, and much paler castaneous longitudinal rays between the veins
on the apical area, extreme outer margin also pale castaneous ; body
short, broad, and robust ; head including eyes as wide as base of meso-
notum ; lateral margins of pronotum angularly ampliate ; abdomen
about as long as space between apex of head and base of cruciform
elevation ; costal membrane of tegmina a little wider than costal
area ; rostrum extending to posterior cox ; opercula somewhat large,
their inner apical areas overlapping, their posterior margins slightly
convexly rounded, and their disks strongly foveately impressed.
Long. excl. tegm. ¢ 38-40 millim. Exp. tegm. 113 millim.
Hab. MADAGASCAR; Baie d’Antongil (Mocouerys—Paris
Mus.).
I have named this species after Prof. A. Bouvier, “Fon-
dateur et Conservateur ” of the Paris Museum, from whom
I have received much valued entomological assistance at
various times.
Yanga grandidiert, sp. n. (Plate XJ, fig. 3 a, 0.)
¢. Head, pronotum, and mesonotum green ; head with margins of
front, a spot behind anterior angles of vertex, the area of the ocelli
and a spot at inner margin of eyes, black; pronotum with the
incisures and two small central spots in front of posterior margin,
black ; mesonotum with a spot in front of cruciform elevation and a
transverse series of four spots behind it, black ; abdomen piceous, a
central longitudinal basal area, the tympanal coverings and posterior
segmental margins pale ochraceous; body beneath and legs pale
greenish or inclining to ochraceous ; tegmina green, much suffused
with bronzy-green, the costal area and membrane pale green, the first
196 Mr. W. L. Distant’s Additions to a
with a central fuscous spot, outer margin with distinct fuscous spots ;
wings ochraceous, posterior margin narrowly, and apex broadly,
castaneous.
Q. Larger than ¢, abdomen above more castaneous, and with the
apical and anal segments cretaceous; tegmina less suffused with
bronzy-green, especially on basal area, where there are two distinct
fuscous spots in radial area, and about four similar ones beneath
them, the apical half is also much more broken up with pale green ;
wings as in ¢, but sometimes with the apical castaneous area only
indicated by two inner spots.
Head, including eyes, about equal in width to base of mesonotum,
its length equalling that of pronotum—excluding its posterior
margin ; opercula in ¢ very strongly concavely sinuate inwardly,
the posterior angles overlapping, and thus exposing a circular space
of the metasternum ; rostrum reaching the posterior coxa, its apex
piceous.
Long. excl. tegm. g 27-32, 9 29-30 millim. Exp. tegm. ¢
70-90, 2 90-100 millim.
Hab. MapaGcascar; Région du Sud-est (Alluaud—
Paris Mus.).
Allied to Y. guttularis, Sign., from which it is to be
differentiated apart from its very distinct coloration by the
more produced frontal area of the head and the very broad
central longitudinal sulcation to face.
I have dedicated the species to Mon. A. Grandidier,
whose name is a household word in the zoological literature
of Madagascar.
Umjaba alluaudi, sp. n. (Plate XI, fig. 1 a, 0.)
Head, pronotum, and mesonotum pale virescent ; head with the
anterior margins of front, two spots on lateral margins of vertex, and
the area of the ocelli, black ; pronotum with two oblique lines on
disk, and eight small spots—three on each lateral area, and two
central on posterior area, black ; mesonotum with four pale sub-
obsolete obconical spots on anterior area, and a transverse series of
four black spots on posterior area ; abdomen above piceous, centrally,
longitudinally castaneous on basal area; body beneath and legs
greenish-ochraceous, ventral segments margined with piceous ; tegmina
very pale tawny brown, the costal membrane and area virescent ; on
basal half there are two spots in radial area, and two or three spots
in each of the ulnar areas beneath it umber-brown, on apical half
there are many umber-brown and greyish mottlings, and a prominent
subcostal greyish spot between radial area and apex of wing ; wings
knowledge of the Homopterous Family Cicadide. 197
pale ochraceous, with an umber-brown submarginal macular fascia.
Head including eyes considerably narrower than base of meso-
notum ; lateral margins of pronotum slightly and obscurely angulate ;
costal membrane of tegmina much narrower than costal area ; oper-
cula with their margins rounded, discally foveate at base, their inner
apical angles overlapping; rostrum reaching posterior coxe.
Long, excl. tegm. ¢ 36 millim. Exp. tegm. 112 millim.
Hab. Mapacascar; “Vallee d’Ambolo”; Foret de
Sakavalana (Alluaud—Paris Mus.).
Allied to U. evanescens, Butl., and although very dis-
similar in colour, the markings of the tegmina are very
similar in pattern. The two species, apart from colour,
can be thus identified :
U. evanescens, g. U. alluaudi, .
Greatest width of tegmina nearly Greatest width of tegmina little
equal to half their length. more than one-third of their
Rostrum passing anterior mar- length. Rostrum only reach-
gins of opercula. ing anterior margins of oper-
cula,
Pyena natalensis, sp.n. (Plate XI, fig. 4 a, 6.)
Body above ochraceous; head with two central fasciw to front
which are connected at centre, a transverse spot at anterior angles of
vertex, and a fascia between eyes, black ; pronotum with a transverse
spot on anterior margin, and two oblique lines from near eyes which
converge on disk, black; mesonotum with four short anterior
obconical spots, of which the central two are largest, a central
lanceolate spot much ampliated at base, two small spots in front of
cruciform elevation, and a submarginal fascia on each lateral area,
black ; abdomen greyishly-pilose, segmental margins, and two
central lines on first and second segments, black ; body beneath and
legs ochraceous, femora streaked with piceous ; central sulcation to
face and spot between face and eyes black ; tibia castaneous, the
tarsi more or less piceous ; tegmina hyaline, the venation castaneous,
costal membrane and area ochraceous; about basal third creamy
opaque, with two hyaline spots in radial area, two in base of both
third and fourth ulnar areas, and a long hyaline streak in lower
ulnar area ; remaining hyaline area of tegmina with scattered fuscous
markings ; wings hyaline, venation ochraceous, about basal half
ochraceous, opaque, which is partially margined both outwardly and
posteriorly with fuscous. Opercula in male broad, transverse,
strongly overlapping at their inner margins, their posterior margins
198 Mr. W. L. Distant’s Additions to a
convex ; rostrum reaching the posterior coxe ; head including eyes
much less in width than base of mesonotum ; pronotal margins much
ampliated and medially angulate, reaching base of basal cell to
tegmina ; costal membrane prominently arched at base and broader
than costal area.
Long. excl. tegm. ¢ 30-31 millim, Exp. tegm. 90-92 millim.
Hab. NATau (Stockholm Mus.).
Div, CICADARIA.
Rihana martini, sp. 0.
Body above dark chocolate-brown, head, pronotum, and meso-
notum sometimes a little paler in hue ; head with the lateral areas of
front, a spot at apical angles of vertex, and a transverse fascia between
the eyes, black ; pronotum with the anterior margin and two central
longitudinal fasciz connected posteriorly, black ; mesonotum with
two anterior central obconical spots, a more indistinct lateral fascia
on each side, a central lanceolate spot, its base widened in front of
cruciform elevation, which has a spot on each of its anterior angles,
black ; abdomen greyishly-tomentose at base ; segmental margins—
very broad on second segment, and anterior margins of tympanal
coverings, black ; body beneath paler than above; head and sternum
greyishly-tomentose, legs and opereula brownish-ochraceous, abdomen
purplish-brown, margins of acetabule crimson; a transverse fascia
between eyes, central sulcation to face, and sometimes subapices to
femora, black; tegmina and. wings hyaline, extreme bases of
both sanguineous; venation ochraceous or greenish-ochraceous ;
tegmina with the transverse veins at bases of second, third, fourth
and fifth apical areas more or less infuscate, and a small fuscous spot
near apices of longitudinal veins to those areas ; opercula about half
the length of abdomen, inwardly overlapping, their lateral margins
coneavely sinuate, their apical margins rounded; rostrum just
reaching the posterior cox ; anterior femora with two strong black
spines beneath.
Long. excl. tegm. ¢ and 2 34 millim. Exp. tegm. 102 millim.
Hab, Mapacascar (Coll. Dist.); Région du Sud,
Andranomana (Alluaud—Paris Mus.).
Rihana hova, sp. n.
Body above and beneath pale brown, thickly finely greyishly
pilose ; head with the lateral striz to front, area of ocelli anteriorly
emitting a transverse linear fascia, lateral margins of vertex, and two
knowledge of the Homopterous Family Cicadide. 199
transverse oblique fascia before eyes, black; pronotum with the
lateral and posterior margins virescent, the disk with two central—
sometimes broken, longitudinal fascize united posteriorly, black ;
mesonotum with two central obconical spots, on each side of which
are two small oblong anterior marginal spots, a broad sublateral
fascia, and a large spot in front of cruciform elevation anteriorly
lanceolately produced, black ; abdomen with the segmental margins
obseure olivaceous; a central anterior spot to face, ochraceous
margined with black, and a black fascia between face and eyes;
tegmina and wings hyaline, the venation brownish-ochraceous,
tegmina with the transverse veins at bases of second and third apical
areas moderately infuscated ; length of head more than half the
breadth between inner margins of eyes, front moderately prominent ;
pronotum with the lateral margins angulate a little before the
posterior angles; opercula in male longer than broad, centrally
overlapping, their lateral margins a little sinuate, their apices
broadly rounded and not extending beyond basal abdominal
segment ; rostrum reaching the posterior coxs, its apex piceous ;
anterior femora armed beneath with two long spines and a smaller
one near apex.
Long. excl. tegm. ¢ 25 millim. Exp. tegm. 77 millim.
Hab, MApaGcaAscaR; Ikongo and Morondava (Paris
Mus.).
Cicada pulverulenta, sp. n.
Pale brownish-ochraceous, shortly but thickly griseously pilose ;
head with the lateral areas of front, lateral apical margins of
vertex, and a transverse fascia between eyes, piceous ; pronotum
with two central longitudinal lines—united posteriorly, and the
incisures, piceous ; mesonotum with four obconical spots of which
the two central ones are smallest, a central lanceolate spot, and a
small spot in front of each anterior angle of the cruciform elevation,
piceous ; abdomen above densely griseously pilose, the disks of first
and second segments distinctly darker in hue ; body beneath pale dull
ochraceous ; a fascia between eyes and a large spot on face, piceous ;
opercula greyishly tomentose; basal segmental margins pale cas-
taneous ; tegmina and wings pale hyaline; tegmina with the venation
and costal membrane ochraceous, both finely spotted with white ;
transverse veins at base of second, third, and fifth apical areas and
small spots near apices of longitudinal veins to apical areas, fuscous ;
wings with the veins ochraceous ; length of head not more than half
the width between eyes ; head including eyes about as wide as base
of mesonotum ; operculain ¢ not passing base of abdomen, sub-
200 Mr. W. L. Distant’s Additions to a
elongate, not overlapping internally, apical margins rounded, lateral
margins moderately concavely sinuate ; rostrum reaching posterior
cox,
Long. excl. tegm. g¢ and 9, 21 millim. Exp. tegm. 55 millim.
Hab. MADAGASCAR and SEYCHELLES (Paris Mus.).
Div. DUNDUBIARIA.
Cosmopsaltria alticola, sp. n.
6. Dull ochraceous ; head with the following black markings,
viz. a basal angulate spot, margins of pale central apical spot,
transverse striations, and basal lateral margins to front and area of
the ocelli, an oblique streak on each lateral area of vertex and a
basal spot before each eye ; pronotum with two central fasciw united
anteriorly and posteriorly, and the inner lateral margins, black ;
mesonotum with the margins of two central obconical spots, a central
lanceolate spot, and two small spots in front of basal cruciform
elevation, black ; abdomen with the segmental margins black, those
to the third, fourth and fifth segments more or less maculate ; body
beneath and legs ochraceous; a fascia between eyes, apex of face, sub-
apical annulations to anterior femora, anterior tibie excluding bases,
apices of intermediate and posterior tibiee, and apices of opercula,
black, the last posteriorly greyishly tomentose ; tegmina and wings
hyaline ; the venation ochraceous, transverse veins at base of second,
third and fifth apical areas to tegmina palely infuscate.
Long. excl. tegm. ¢ 40 millim. Exp. tegm. 108 millim.
Hab. BoRNEO; Kina Balu (Brit. Mus.).
Allied to C. montivaga, Dist., but with the opercula only
reaching the fifth abdominal segment; markings also
different.
Platylomia juno, sp. n.
Head, pronotum, and mesonotum dull ochraceous; head with a
large quadrangular spot on lateral margins of front, the area of the
ocelli connected with a transverse angulated spot on each side, and a
basal spot near inner margins of eyes, black ; pronotum with two
much angulated central fascia connected with a curved discal fascia
on each side, and a broad sublateral fascia, black; mesonotum with
two broken anterior obconical spots, connected with a central fascia
much widened posteriorly and occupying the whole anterior area of
the cruciform elevation, two sublateral fascia and a posterior
marginal spot, black; cruciform elevation centrally ochraceous ;
knowledge of the Homopterous Family Cicadide. 201
abdomen black, the anterior areas of the segments above somewhat
castaneous ; sternum and cox ochraceously tomentose ; transverse
striations and longitudinal area of face, a transverse spot between
face and eyes, and spots to cox black ; a basal spot to face, and the
rostrum excluding apex, ochraceous ; legs piceous or black, apices of
femora ochraceous ; opercula black with their disks dull ochraceous ;
tegmina and wings hyaline, the venation fuscous or brownish-
ochraceous, their bases narrowly piceous; tegmina with the
transverse veins at the bases of the second and third apical areas a
little infuscated. Face prominently tumid; rostrum extending a
little beyond the posterior cox, opercula only reaching the third
abdominal segment, situate widely apart and occupying the lateral
abdominal areas, sinuate on each side near base, narrowed and a
little rounded posteriorly. ;
Long. excl. tegm. ¢ 40 millim. Exp. tegm. 114 millim.
Hab. Se-TcHouEN; Ta-tsien-lou (R. P. Gros-Jean—
Paris Mus.).
Allied to P. umbrata, Dist.
Platylomia diana, sp. n.
Closely allied in general appearance and markings to P. juno, Dist.,
but larger, the opercula reaching the sixth abdominal segment
ro) d Pp to} 5 )
entirely ochraceous excepting their extreme margins and apex,
posteriorly globosely convex ; legs ochraceous, longitudinal streaks
to femora, bases and apices of the tibis, and the tarsi, black.
> ’ >}
Long. excl. tegm. ¢ 50 millim. Exp. tegm. 126 millim.
Hab. St-TcHouEN, Ta-tsien-lou (R. P. Gros-Jean—
Paris Mus.).
Oncotympana virescens, sp. .
dé. Body above black ; head with a large spot at base of front,
and a small spot on each side of ocelli, ochraceous ; pronotum with
a central lunulate ochraceous fascia, the whole of the lateral areas
piceous, the posterior margin inwardly pale-greenish ; mesonotum
with the lateral margins, the margins of two central obconical spots,
and two small spots in front of cruciform elevation, ochraceous ;
abdomen with the posterior margins of the second and third
abdominal segments, the tympanal coverings, disk beneath, and
opercula, pale bright virescent, sternum thickly ochraceously tomen-
tose ; legs black, longitudinally streaked with virescent ; tegmina
and wings hyaline, extreme bases of both virescent, the venation
fuscous ; tegmina with the transverse veins at the bases of the second,
202 Mr. W. L. Distant’s Additions, ete.
third, fifth, seventh, and eighth apical areas broadly piceous, and a
series of small piceous spots near apices of longitudinal veins to
apical areas ; opercula in male, broad, centrally overlapping, and
reaching base of second abdominal segment; rostrum reaching the
posterior coxe,
Long. excl. tegm. ¢ and 9 19-20 millim. Exp. tegm. 107-112
millim.
Hab. Tiset ; Tsekoo (Paris Mus.).
EXPLANATION OF PLATE XI.
Fig. 1. Umjaba alluaudi.
2. Yanga bouvieri.
3. 4, grandidieri.
4. Pycna natalensis.
5. Munza revoili.
May 20TH, 1905.
( .203 5)
XI. On the Pupal suspension of Thais,
By T. A. Cuapman, M.D., F.ZS.
[Read February 1st, 1905.]
PLATE XII.
IT is now some ten years since I ventured to question the
view quoted in Scudder’s great work on the American
Butterflies as to the suspension of the pupa of Thais
(Ent. Record, Vol. VI, pp. 125, 126), and somewhat later I
obtained (Ent. Record, Vol. VII, pp. 81, 82) tolerably
strong evidence that my opinion on the subject was correct.
Up till the present time, however, no one has chosen to
report actual observation of how the larva and pupa of
Thais actually manage to make the girth (incidentally
proving that it 7s the girth) leave its usual situation and
become attached to the nose-hooks. This Spring (1904) I
made an effort to supply the deficiency, and obtained a
number of larvae of Thais polyxena, Schiff., var. cassandra,
Mann, from Hyeres. I was successful in observing the
whole operation by one specimen or another; whether I
can successfully describe what I saw is I fear doubtful,
but I will make the attempt. I was so interested in the
matter, even someway outside the chief point in question,
that I also made a successful attempt to see Papilio
machaon and Pieris rape (as examples of Papilionide
and Pieridz respectively) make their silken holdfasts
for their pupee. Though there is no novelty about my
observations of these, there are one or two points that
may bear description again.
The first spinning done by the larva of Zhais is to form
what we must call a cocoon, though it consists merely of
three or four, or at most a dozen, rather strong silken
cables, sometimes simple, sometimes branching, tying
together the objects surrounding the position chosen for
suspension. This structure must be correlative with a
habit of retiring for pupation into a situation surrounded
by not very fixed materials, such probably as dead herbace-
ous material round shorter stems near the ground-level or
below it. Having prepared a carpet of silk of rather more
TRANS. ENT. SOC. LOND. 1905.—PART II. (JULY) 14
204 Dr. T. A. Chapman on the
than its own length, either on a flat surface or by prefer-
ence on a round one such as a stem, 1t makes the anal
pad. It may be noted here that in Thais and the other
girthed pupze observed, when this is completed the larva
takes its station with the claspers just in front of it, the
little mound of silk forming the pad being unused, and
lying immediately behind the claspers and beneath the tip
of the anal plate. In suspended pupze this pad is held by
the claspers, whilst the larva awaits pupation.
Before describing the spinning of the girth, it may be as
well to explain its position on the larva when it is finished
and the larva rests before pupating.
The girth arises well forwards from the carpet of silk,
and passes over the back of the first abdominal segment.
Kach segment of the larva has on either side three tall
processes or warts, which with their colours and spines
give the larva its special and beautiful aspect. These
three warts are a subdorsal one (= I. + II.?), which is the
largest, then a somewhat smaller one (= IV. + V.?), which
arises below the spiracle though one is inclined before
examining it to think it is subdorsal also, 7. e. above spiracle.
The third one is still smaller, and is marginal Cas Vit)
In passing over the segment, the girth reaches backwards
from its attachment, to behind the marginal process, and
passes up in the same line behind the subspiracular one ;
then it turns forward, and gets in front of the subdorsal
process crossing the dorsum therefore rather on the front
of the segment, proceeding down the other side of the
larva, by, of course, a precisely similar route, to its attach-
ment to the carpet on the other side. In taking this
course it makes several bends. First it inclines rather
backward from its attachment, has a bend at the marginal
process where it turns upwards, and two more bends in
passing from behind the subspiracular to the front of the
subdorsal process. It is fairly taut, so much so as rather
to bend forward the marginal process, against the similar
process of the meta-thoracic segment. Nevertheless the
curved or angulated route ensures that it shall be of
considerable length, longer, that is, than a merely simple
trausverse course across the back of the larva.
I succeeded in seeing this cable or girth manufactured
on several occasions, but that does not seem to help me in
describing it in words. As in Papilionidx and Pieride, it
is not spun where we see it when completed, but in a
Pupal suspension of Thats. 205
position that may be described as in front of the larva,
the head being thrown back, so that the legs are used as
hands, one might say, to hold it up. Not, however, the
claws, but the thick bases of the legs are used, the silk not
being on the legs proper, but rather in the incision in front
of them. This is the position when the spinneret is at the
middle of the girth. But as the head goes from one side
to another, the relations of parts is much changed, though
quite gradually and automatically. It is this that makes
it difficult to give a description easy to understand.
The actual line of the girth, at the middle of the move-
inent, when the larva is straight with the head and legs
well raised, is behind the marginal process of the Ist
abdominal segment, then forwards above the marginal
processes of 2nd and 3rd thoracic, and then across the
larval venter to the other side in the incision between the
first and second pair of legs. I have said the head is well
raised, and so it is, by the Sphinx attitude of the first
segments, but it is strongly bent forwards, so that the tip
of the spinneret reaches very closely to the position of the
girth in the incision behind the first pair of legs. I say
very nearly, for the girth when completed consists of a
number of quite separate threads, showing that each
thread is not spun along, and glued to those that preceded
it, and that therefore the extremity of the spinneret does
not actually reach and touch the previously spun threads,
which lie deeper in the incision between the segments.
As the larva moves its head from side to side in adding
each thread, the position of the girth differs from this
central position by being stretched along one side and all
but relieved from the other; when the head is round to the
left and the left end of the tliread is being fixed, the thoracic
and first two abdominal segments have their right sides
stretched so as to form the margin of nearly a circle,
whilst their left sides are so approximated as to be close
together at the centre of the circle. In this attitude the
line occupied by the girth above the marginal tubercles on
the stretched side of the larva is raised above the surface
on which the larva rests, and is on what for the moment is
rather the upper-side of the larva, though it would be
rather the under-side if the larva were im a natural resting
attitude, since, as I have above called attention to, the
larval warts are,and look, higher up on the larva than their
real anatomical positions would indicate.
206 Dr. T. A. Chapman on the
The following notes made whilst actually watching the
larva may help to explain the spinning of the girth, though
they have some incoherence from the circumstances of their
production.
Cassandra, when building its loop, has it quite ventral,
it passes round above the marginal wart of 2nd and 3rd
thoracic and 1st abdominal and between the Ist and 2nd
pairs of legs. ‘There is, however, only one phase in this
process to which this applies, viz. when a thread of silk is
being added to the loop at exactly its highest point. When
the spinneret is against either side of the loop or working
at the point of attachment, the loop is free from all contact
with that side of the larva, or just touches the marginal
wart of Ist abdominal, whilst it is fully stretched on the
other side, over the 2nd thoracic leg, the three warts (2nd,
3rd_ thoracic, Ist abdominal), and touches the ventral
prominence of Ist abdominal. The attitude of the larva
being that of a curve or twist that brings these parts
directly away from the twig of attachment, whilst those
that the loop is free from are crowded together towards
the twig, the three warts having their apices close together,
and so that on the curvature relaxing they expand
again into that side of the loop and hold it whilst the
other side contracts and frees itself from the loop in its
turn.
The larva moves very leisurely, and with some to and
fro movement, so that one traverse of the loop takes about
three minutes and the movements of fastening the end of
each thread to the twig about one minute; but between
each complete traverse usually at least one partial journey
is taken, 7.¢. from the twig for about one-third of its
length and then back again, and along this piece especially
towards the end of the process a good deal of local spmning
is done which covers this thicker portion of the loop with
an outside binding. When the loop is finished, the central
third consists of a number of threads more or less separate,
or at least apparently separate, straight, parallel and
uncomplicated. The end portions are thi¢ker, and bound
together as one strong strand.
As the larva moves its head from side to side, the loop
slips to and fro, or rather perhaps the larva slips to and
fro on it, the loop always taking the position described on
the convex side of the larva, and lifting out of it on the
concave, as the warts and legs are contracted together into
Pupal suspension of Thais. 207
one eminence. The head and prothorax are stretched out
when the attachment of the loop is being worked at, the
first pair of legs being one on each side of the strand ; when
the centre of the loop is under the spinneret the thorax is
bent back from the twig, but the head is bent down to
bring the spinneret against the loop in front of the 2nd
legs, and the Ist pair of legs become practically invisible.
In two specimens watched the whole process took about
an hour and a half,ina room at about 64°. One wondered
all the time how the loop when finished was to get back
to its place. This however is managed very simply and
very quickly. After finishing, as it had appeared to finish
several times before, one of the lappings of the side of the
loop with binding threads, instead of sliding slowly round
and bending the head down slowly as it went, to the posi-
tion it takes when at the central point, it gave it this
position at once, 7. ¢. before moving round, so that the head
went under the loop somewhat to one side, and as it then
gradually assumed the median position, the thread lay
across the middle of the front of the head. At the same
time, however, as it assumed the median position, it bent
back the head and curved the thoracic segments back wards,
so as to bring close together the back of the head and the
dorsal thoracic humps. Then the thread became slack
over the head, and slipped back into its place. In one
instance, the thread caught, on the side on which the
manoeuvre was made, the tops of the subspiracular warts
of the meso-and meta-thorax, bending them back, the larva
rested here a little, probably from some inadvertent move-
ment of mine that alarmed it, and then by asimple turning
and stretching movement the loop fell into its place. One
asks, Why does the loop always fall to this identical spot,
how does the larva manage to make the necessary move-
ment so exactly? This question is justifiable as an
expression of admiration that the larva should always
make precisely the same movement, but is foolish in not
seeing that the same movement will always produce the
same result. In fact a little, but very little, latitude is
possible, as one could see that the result would be the
same had the movement been a little more or less ample,
etc. Nevertheless, the length of the loop and the place
from which it starts must be very exactly related to the
length, thickness, and movements of each larva.
Another point may be noted, whilst the larva is at work
208 ' Dr, T. A. Chapman on the
on the loop, the thoracic segments are rather contracted
and the abdominal somewhat full, and when first the loop
falls into place, it runs more directly backwards than it
does later; the head of the Jarva being rather in front
of the points of attachment of the loop. But gradually
the abdominal segments shrink, the thoracic enlarge and
bend forwards, the whole larva thickens and shortens so
that the loop has the position described in my first note.
The amount of spinning for the anal hold varies a good
deal, but in two specimens in which it appears to be well
elaborated, it stands up as a somewhat flocculent little
mass, and the prolegs take hold not of this, but a little
way in front of it, suggesting that the pupa shall have
a freer access to it than if the prolegs held it
My special object being to see how the girth, now of
course clearly seen and proved to be a girth, as in other
Papilionide, got moved to the position and function of
a nose-cable, I watched carefully for the moult to pupa,
to see how this occurred. My notes on this throw no light
on the critical question, but may be worth giving on their
own merits.
A larva that was very close to the change was watched
on—
May 26, 9a.m. No very definite change noticeable.
10 a.m. Some movements observable and a slight change of
colour, 10.30am. All the red has gone from the warts,
except those of 8 and 9 abdominal. The others are shrivelled
and show no colour, except their black points, the rest
of the larva is slaty-grey, quite different to what it was
an hour or two ago, No trachea drawn out yet, 3rd
thoracic and 1st and 2nd abdominal are very narrow and
shrunk. 2nd thoracie large. A good many little wrink-
lings of empty skin are visible round warts as well as
elsewhere on segments. Girth hangs heavily on marginal
wart of 3, pressing it forwards, that of Ist abdominal seems
to have slipped behind it on one side, collapsed against
that of 3rd thoracic on the other. Wart of 9 still full and
red, the red would almost seem to depend on the contained
fluid, as it disappears with the shrinking and collapse of
the warts, which has just occurred. The fluid which filled
them must have been absorbed or evaporated very rapidly.
Disturbed the larva by placing it in a better position for
observation. 10.40. A few slight movements. 4th, 5th
and 6th abdominal seem large as also 7th and Sth; 3rd
Pupal suspension of Thars. 209
about normal, intermediate between these and the shrunk
Ist and 2nd.
10.45. A few more movements, the changes of dimen-
sions of segments have placed the mouth parts and the
legs below the girth. They were not very definitely
above or below it before, but rather unrelated to it, but
: strictly lateral view made it cross over 2nd pair of
egs.
11.35. Colour all but gone from warts of 9th abdominal
and claspers are almost collapsed, no other very obvious
change. Th. 64° Fahr. 11.50. Some movement occurring.
The large size of abdominal 4th, 5th, 6th and 7th very
conspicuous, the incisions being much expanded. The
appearance is as though the girth was holding up the skin
on Ist and 2nd abdominal and meta-thorax, but the
contents had gravitated back and distended these lower
segments. In this specimen the silk for the tail grip is a
little tag, a little loose and flocculent, shaped like a short
blunt thorn, of which the apex is at present opposite the
middle of the dorsal surface of 10th, the prolegs holding
the stem to which the larva is attached some 2°5 mm.
above it.
12.30 p.m. Length of different portions of larva:
Head and Ist and 2nd thoracic = 3 mm., but these are
curved and are much more along dorsum, 3rd thoracic,
Ist abdominal = 2 mm.,, 7.¢. 1mm. each. 2nd and 3rd
= 3 mm. = 1°'5 each. 4th, 5th, 6th = 7 mm,, 22.
2:3 mm. each; 7th, nearly as much. 7th, 8th, 9th, 10th
= .5) om,
12.45 p.m. Strong movements of contraction in 2nd, 3rd,
4th abdominal segments, but quiescence elsewhere. No
definite vermicular movements.
12.55. Regular vermiform movement from behind for-
wards, one every five seconds without so far much change
in skin. .
12.58. Tracheze drawn out along abdominal segments
2-7, loop drawn back, front segments straight.
1.2 p.m. Split down back in usual way. Skin slips back
under loop.
1.5. Skin behind loop. Scrap of broken-off tubercle
sticks to it; loop lies in 3rd and 4th abdominal incisions.
1.10. Skin gone. The movements of getting hold by
the cremaster move the loop to the thoracic abdominal
incisions, where it deeply indents wings during movements
910 Dr. T. A. Chapman on the
for getting rid of skin. Wings not yet descended fully
on to 4th abdominal.
1.15. Various movements apparently for forcing fluid
forward and expanding wings, etc.
1.18. The attachment of the loop is just opposite the
eyes of the pupa; wings, etc., and head above loop swollen
and knobbed, below smooth and tapered.
1.22. Occasional movements, but on the whole resting,
wings not quite down. Hangs much like an ordinary
Papilio, except for appareut strangulation of front.
1.24, Wings to place.
1.35. Ist spiracular hollow (with black spiracle at bot-
tom) sinking, dorsal eminence of meso-thorax and wing
angles becoming of more mature form and less like swellings
from strangulation, but loop still cuts deeply into wing
and across base of hind-wing.
1.50. The wings are much straightened out and the
loop cuts into them very little to what it did. Abdominal
segments are shortening and closing up, not yet quite so
much as in a mature pupa.
1.54. Is hanging with a strong sag (or bend) in
abdomen.
2.3. From head to end of 4th abdominal, now very long
and abdominal segments diminished. The loop is now
hardly buried and is seen to be on top of 3rd thoracic (not
in 3rd thoracic to 1st abdominal incision) at about 4 or}
of its width from front border.
2.15. Straightened itself a little and then fell back
again. Meso-thoracic dorsal ridges becoming sharper,
anterior end still obviously somewhat soft.
2.25. Straighter,—no deformity of wings can be seen
either where loop now crosses, or where it did before,
except it seems a little impressed on one ridge of the
venation.
2.30. A. lateral jerk or two.
3.40. Loop is still a girth, though pupa is apparently
mature as to form, and much dark shading has appeared.
5.45. No further change, still slung as a Papilio.
In both this and following specimens the girth is quite
loose and at liberty to slip in any direction. Moving the
pupa by touching the head enough to show the girth to be
quite free does not in any way alter its position.
It does not look possible for it to slip forwards, but a
twist of the pupa might catch it by the wing-spine and
Pupal suspension of Thais. 211
throw it forward. Something of this sort must occur,—
when ?
May 27. This specimen escaped observation but moved
the girth to nose-hooks some time between 1.15 and 3 p.m.,
twenty-four hours after moulting to pupa.
SECOND SPECIMEN.—May 26, 12.35 p.m. Subspiracular
tubercles of Ist, 2nd, 3rd and 4th abdominal are now colour-
less and collapsed, dorsal and marginal of same region also
changing. 3rd and 4th abdominal segments are beginning
to enlarge. Incisions 4th to 5th abdominal getting wide;
up till half-an-hour ago no change was visible and segments
were not of very unequal lengths.
1.16. Only a few anal warts still coloured.
1.35. Only wart of 9th abdominal segment still coloured
red.
1.50. Thermometer 74° Fahr. Abdominal segments large
and incisions wide, 3rd thoracic and Ist abdominal very
small. The higher temperature seems helping this
specimen on more rapidly than No. 1.
2 p.m. Abdominal incisions wide, lower end looks like
end of pupa with larva skin overlaid and claspers (shrunk)
stuck on outside.
2.20. 4th, 5th and 6th abdominal very large, incisions
wide ; some stretching, almost vermiform, movements.
2.43, Genuine rhythmical vermiform motion begins
and head rises a little.
2.50. Quiet again, the three free incisions (of pupa)
very wide; from abdominal 4th to 5th incision to head,
11 mm., to claspers, 9 mm.
2.59. Movement beginning again, in Ist, 2nd and 35rd
abdominal segments.
3.5. Regular vermiform movement from end to end.
3.6. Skin accumulated at tail.
3.7. Thoracic skin split, it slips down, the collapsed
warts passing under loop.
3.15. Process finished. The getting rid of larval skin
is delayed a little by its fluid adhesion to pupa. The
cremaster got hold at second try, and skin was shoved off
at two twists, after fluid adhesion gave in. Loop fell into
abdominal 3rd—4th incision as soon as skin left it, but in
twistings for shoving off skin it slipped quickly to back
of prothorax. To end of wings 8 mm., wings to cremaster,
12 mm.
3.22. Wings getting rapidly down to place. There are
212 Dr. T. A. Chapman on the
contracting and expanding (lengthwise) movements of
thorax.
3.30, Wings into place. 9 mm. head to end of wings,
11 mm. wings to cremaster = 20 mm.
3.40. Top of wings and meso-thorax, etc, have the
appearance of being ‘strangulated above loop, being swollen
out and lumpy.
3.45. On lateral view, pupa has very Papilio-like out-
line, say podalirius, head thrown back, curvature with
ventral projection of wings, etc., very different from the
straightness of the mature Thais pupa.
4.3. Is now hanging very much sagged and bent.
5.45. Seems nearly mature and has more dark colouring.
Head to end of wings 12 mm. Wings to cremaster 9°5,
May 27, 4.50. Was not looking at specimen but at
another two inches from it, when a sudden movement
attracted my attention to it.
I was in time to see the girth take hold of the nose-
spines, and see the pupa complete what appeared to be
two revolutions on its axis, but might have been one, or
three; the whole taking place in less than a second, per-
haps a third of a second, apparently as. rapidly as an
active pupa rotates his tail.
A very unexpected movement, as the pupa looked so
straight and stiff (and it is fairly stiff when one receives it
in autumn), and for a pupa with a girth to rotate on its
axis seems so unlikely, the way in which the girth is
twisted round the nose-hooks Jed me to expect a rotatory
movement, but without any change of orientation. What
struck me afterwards as remarkable was how in such a
sudden movement the pupa came to rest again exactly
facing its attachment. An examination of these two
pup shows that they made at least two revolutions, as the
girth has reached the nose-hook and one further revolution
has been made twisting it round once (quite) after it had
caught, and the cremastral silk also shows twisting.
I have not made it very clear how the revolution is
effected. It is by bending the tail to one side and then
twirling it round ; when I say to one side, I should in this
case say forwards, as the attitude during the movement
was that of bending away from the supporting twig.
Another specimen was observed to make a second
rotatory movement, on one occasion, some hours after the
cable had been moved forwards, but it would seem to be
Pupal suspension of Thais. 213
the rule that rotation is made once only for a fraction of
a second, about twenty-four hours after the moult.
The rotation is on an axis, z.¢. it does not sweep the
head round in a circle, maintaining the venter towards the
stem of attachment all the time, but presents towards the
stem successively one side, the dorsum, the other side, and
so on. The rotation is effected by the abdominal seg-
ments being bent forwards, 7.e. virtually making the pupa
preseat a deep curve towards the stem. Then by sweep-
ing round the abdominal segments in a way that is common
to many pupze the hollow of the curve remains towards
the stem, but affects successively each aspect of the pupa
as it rotates.
The mature pupa a few days later is very stiff and
straight, and its possessing such motility twenty-four hours
after moulting is somewhat unexpected.
In order to observe the way in which the girth is made
by the larva of Papilio, I obtained a few larva of P.
machaon, and kept rather a close watch on them, but four
succeeded in suspending themselves, girth and all, without
my having surprised them at work. They do it apparently
somewhat quickly, and give no clear indication beforehand
of when they are likely to do so. With the fifth I was
more fortunate, happening to look at it when the making
of the girth was under way, indeed more than half-finished ;
he was working at one end, passing the spinneret too and
fro about the attachment of the loop. The loop itself,
consisting of a number of quite detached threads, passed
between the fore-legs of the larva, forwards, the larva being
so bent round that its head and prothorax were directed
backwards. Then well above the meso-thoracic leg and
rather higher on the meta-thorax, then well above the
spiracle of the Ist abdominal, not quite so far above that
of the second, and then passed down to its other attach-
ment. The most anterior portion of the larva in this
position was the side of the meta-thorax and Ist abdominal
segment just below the loop. Then the larva began to
add another thread to the loop. The head and front seg-
ments are well raised and the loop falls into the incision
between the Ist and 2nd pair of legs, the head is pressed
down and the spinneret lies between the first two legs
and reaches very close to the loop; the larva sweeps the
head across from the one end of the loop to the other
slowly and by short jerks and stops, as though the silk
214 Dr. T. A. Chapman on the
had to be drawn out. As it does so, the loop which was
stretched over the sides of the four segments (2nd, 3rd
thoracic, Ist, 2nd abdominal) on one flank, gradually leaves
them and becomes stretched over the same line on the
opposite side. Then the process of fastening it and
spinning various sinuations of silk over the other place
of attachment of the loop is repeated. I saw it do this
several times, and then at one end it delayed a little, and
came up with the head not above the loop, still less the
front-legs, but with the loop across the front of the head
at the base of the labrum. When about the middle, it
made some movements and I thought it was going to
throw the loop backwards into place ; this, however, it did
not do, but continued to the other side ; so far as I could see
it spun no thread this time, it then twice repeated the
ordinary process of adding a thread, and again repeated
the passage with the thread above the labrum. It was
now making a good many contortions with the effect that
the anterior segments became decidedly diminished in
bulk ; as soon as it reached the end of the loop with it
above the labrum, it put the head with little difficulty under
the loop so that the loop was across the vertex, and then
making the passage across and raising the mouth end of
the head, the loop easily slipped backwards. It was not,
however, before some little time spent in twistings and
contortions, that looked as if intended to push the loop
back, but were really effective in again distending the
front part of the larva and diminishing the posterior that
the loop fell into its place between the 2nd and 3rd
abdominal segments, and the larva rested as quite satisfied.
Though the larva was slow and deliberate in its move-
ments this was all done ina very few minutes; the threads
of the girth seem to remain distinct and separate, and do
so to much nearer their attachment than in 7’. cussandra,
where the spreading of silk over the attachment includes
the lapping of the adjacent portion of the loop.
It may be noted that the position of the loop in the
larva between the 2nd and 8rd abdominal is constant, as I
saw it in all specimens, though I did not see it made. In
the pupa it is different, viz. just behind the middle of Bue
meta-thorax.
It sinks into the soft chitin of the newly-moulted pupa
which, as it were, flows over it in two places on each side,
there is in fact a lappet formed at each of these places,
Pupal suspension of Thais. 215
that passes over the loop and encloses it in a short tunnel,
Sometimes it is quite fixed here, at others it can be drawn
through these tunnels and removed, if it is first cut. The
two places where these lappets are found are on the hind-
wing, and on the fore-wing just below the cell some little
way basal to vein 2.
It is perhaps erroneous to say that the loop sinks into
the soft chitin, at least to say so may produce the false
impression that there is something accidental about it; it
is, however, a constant arrangement, and the lappets are
no doubt structures specially arranged for the purpose.
To test this a larva suspended for pupation was treated
by cutting the loop away. It was seen very shortly after
pupation, before the pupa had quite assumed its permanent
form and was still very soft. The very striking fact
appeared now, that across the wings there was a very
distinct depression marked by a central linear mark, as
if the girth were present and indenting the soft pupa.
When the pupa had attained maturity with a firm skin,
this appearance had gone, and at first glance there was no
evidence of the overlapping chitin that bridges over the
girth. A closer look, however, showed the prominence at
the base of the under-wing that forms the anterior over-
lapping margin, with a linear groove just behind it, and on
the middle of the fore-wing at the position of the tunnel for
the girth is a glazed line, but no definite overlapping. It
therefore seems that the pupa is prepared for the girth tak-
ing its proper position by a definite channel occurring across
the wings whilst the cuticle is still plastic; that the two
special places (on fore- and hind-wing) are prepared for
its inclusion beneath the surface; that of the hind-wing
which is a very strong lappet forms even if the girth is not
there; that on the fore-wing, the overlying lappet does
not form, unless it has the assistance of the downward
growth of the wings, which occurs during the maturing of
the pupa just before hardening, and as the lappet is not
formed, the walls of the tunnel that would contain the
girth are left exposed as a glazed line in this position.
As a subsidiary effect of cutting the girth the pupa did
not succeed in fastening the cremaster on the provided
anal pad of silk. When at rest for pupation the claspers
hold the silk in front of the pad, which stands up un-
secupied just behind the pupa, or rather behind the
claspers and under the end of the anal plate, and in
216 Dr. T. A. Chapman on the
moulting the cremaster is thrust back over the skin to
reach the pad. The loss of the girth, however, deprives
the pupa of the power of preserving a correct alignment,
and so the fulerum provided by the skin held in place by
the claspers cannot be efficiently utilized.
The whole process of spinning the girth seems therefore
to be identical in Papilio and Thais.
P.S.—The abortive journeys across the loop, between
the spinnings of its separate threads and the final journey,
in which the loop is thrown back, may be speculated on as
showing that the movement for spinning and that for
throwing back are variations of one and the same move-
ment, and that the definite distinction between them is
not so fixed, but that an intermediate movement may
occur, either as not being yet entirely eliminated or by
reversion.
The larva of rape makes the girth in a way that is
essentially the same as in Papilio, but yet with an amount
of variation that renders it actually very different.
Ersentially the girth is made in front of the larva and
between the head and first pair of legs (not between the
Ist and 2nd pair of legs, as in Papilio), but the raising of
the front segments of ‘the larva, which in Papilio may be
likened to the ‘ ‘Sphinx ” attitude, is In vapex carried to an
extreme, so that when the larva is adding to the middle
point of the girth the head is bent back so that the back
of the head touches the dorsum of the abdomen, about
the incision between 2nd and d5rd abdominal segments,
the ventral face of head and first thoracic segment beimg
directed exactly dorsal, the legs of 2nd and 38rd_ thoracic,
forwards. As the head is carried to either side, these for-
ward segments so rotate that the venter becomes ventral
over all segments, but the forward segments instead of
being bent dorsally, are bent laterally, and the head is
against the side of the 2nd and 8rd abdominal segments.
Tn all these positions the loop seems to be fairly tense.
When the head is bent to one side, the girth passes over
the middle of the 2nd abdominal segment and the middle
of the Ist thoracic, the portion of the larva between
these two positions being in front of the loop, the rest
behind it. In the median position, there is perhaps a
large proportion of the Ist thoracic segment in front of the
loop. Indeed the head only might be regarded as behind
the loop. It is observable, that during this process the
Pupal suspension of Thais. yA §
2nd abdominal segment seems small and contracted, the
abdominal segments behind and the thoracic in front
seeming relatively swollen.
The completion of the process, when the spinning is
finished, is really very different from that in Papilio. In
Papilio the front of the head is put forward under the loop
and it is slipped back into its place by a movement very
similar to that by which a thread is added to the girth,
In rape, at the end of fixing the last thread at the side,
the head is merely drawn forward from under the loop.
The references to the spinning of the loop in Pieridx
that I have met with, give the idea that it is spun from
the outside across the abdominal segments, the spinneret
being carried to and fro across the surface of the
segment—a feat that a moment’s reflection on the details
of such a process will show to be impossible. It is made
across the 2nd abdominal segment, but the larva is so
bent back that the loop passes at the same time round
its neck and the spinning is from the inner- or under-side.
When I say that the spinning from the outside 1s
impossible, it is perhaps going too far, for there is no
necessary limit to the amount of bending a larva may do,
but when one sees the amount of strain on the full-fed
larva of rapx to get the head as far back as it does, it is
seen that whatever it might be for some hypothetical larva,
it would be impossible for rapx to bend further back till
the spinneret touches the abdominal dorsum. Or if we
take the actual position when the larva is fastening one
end of the loop, if it thence carried the spimneret up the
side of the segment towards its dorsum, the combination
of lateral bending and longitudinal twist would stop the
process before the spinneret reached the dorsum, much less
reached the other side.
218 . Explanation of Plate.
EXPLANATION OF PuaTE XII.
SUSPENSION OF Z'HAIS.
Three several figures of the larvee of different individuals sus-
pended for change to pupa to show (which the smallest specimen
does most successfully) the position of the “ girth” at this stage,
and three of pupa, the first with the girth in the Papilio position it
oceupies during the first twenty-four hours after moult, the other
two with it in the well-known Thais position attached to the
nose-horn.
( 24901)
XII. Notes on New Zealand Lepidoptera. By E. Meyrick,
B.A, E.RS., F.ES.
[Read February 1st, 1905. |
For the material of the following notes and descriptions
Iam again indebted to the kindness and energy of Mr.
G. V. Hudson, of Wellington, except where it is otherwise
stated. In addition to describing the new species I have
made some corrections of synonymy and other notes.
HYDRIOMENID A.
Chloroclystis sandycias, sp. n.
Chloroclystis plinthina, Huds., N. Zeal. Moths, 41,
pl. vi, 8, nec Meyr.
49. 14-16 mm. Head and palpi ochreous-white, seldom
mixed with green, palpi 2%. Antennz white ringed with dark
grey, in ¢ ciliated with fascicles (3). Thorax whitish-ochreous
sprinkled with blackish. Abdomen ochreous-whitish, more or less
tinged with reddish-ochreous towards base and apex, and variably
sprinkled or mixed with blackish. Fore-wings somewhat elongate-
triangular, costa faintly sinuate, apex obtuse, termen bowed, rather
oblique ; ochreous-whitish to white ; basal area more or less tinged
with reddish-ochreous and suffusedly striated with blackish irroration,
usually extending considerably further on costal area than on dorsal;
median band almost always conspicuously pale, sometimes mixed
with green, generally striated with dark irroration on costa and
dorsum but seldom indistinctly throughout, posterior edge formed
by a double pale line prominently angulated in middle, more
than usually approximated to termen on lower half ; when a series of
blackish neural dots precedes this, it follows the angulation of the
line ; a blackish linear discal dot, sometimes indistinct ; terminal
area reddish-brown, seldom mixed with green, interrupted by a
pale patch opposite angle of median band, subterminal line pale,
waved ; an interrupted blackish terminal line: cilia whitish more
or less suffused with fuscous-reddish, basal half barred with
blackish, apical half less distinctly with grey. Hind-wings with
termen sinuate, rounded-prominent below middle ; whitish, towards
TRANS. ENT. SOC. LOND. 1905.—PART IL. (JULY) 15
220 Mr. E. Meyrick’s Notes on
dorsum obscurely striated with reddish and blackish irroration ; a
rather large blackish-grey discal dot ; terminal area with indistinct
grey lines, sometimes reddish-tinged ; cilia as in fore-wings.
Twelve specimens, Wellington. Although very variable,
this is a neat and easily recognised species; it is one of
the smallest, and the almost constantly conspicuous paler
median band is a striking feature; it is perhaps nearest
to C. plinthina, but in that species the palpi are much
longer (especially in the ?, in which sex they are 42), the
expanse of wing greater (19-20 mm.), termen of fore-
wings more oblique, post-median series of black dots
gently curved in disc, not angulated.
Xanthorhoe chlorias, Meyr.
Venusia princeps, Huds., Trans, N. Zeal. Inst. 1902,
244, pl. xxx, 1, 1s a synonym of this.
Aanthorhoe occulta, Philpott.
Nanthorhoe occulta, Philpott, Trans. N. Zeal. Inst. 1902,
248, pl. xxxii, 5. A specimen received ; it is a good
species, apparently allied to Yanthorhoe mnesichola.
NXanthorhoe periphea, sp. n.
dé. 26mm. Head, thorax, and abdomen fuscous sprinkled with
whitish. Fore-wings triangular, termen slightly bowed, oblique ;
light fuscous, sprinkled with darker fuscous and whitish ; two very
obscurely darker fascie indicating median band, first curved, pre-
ceded by several white dots, second irregularly curved outwards
on upper 3, followed by a series of white dots; some whitish dots
indicating subterminal line, edged with obscure darker shades : cilia
fuscous-whitish, with two fuscous lines. Hind-wings rather elongate,
light fuscous ; cilia as in fore-wings
One specimen; Humboldt Range, Lake Wakatipu, at
4000 ft. An obscure species, apparently nearest to
A. chionogramma.
Notoreas synclinalis, Huds.
Notoreas synclinalis, Huds., Trans. N. Zeal. Inst. 1902,
244, pl. xxx, 6.
¢d 2. 25-26 mm. Head and thorax grey-whitish, more or less
mixed with fuscous ; thorax with a blackish stripe on each side of
back. Fore-wings rather elongate-triangular, costa sinuate, termen
bowed, rather strongly oblique ; ochreous-brown ; a broad grey
New Zealand Lepidoptera. 221
streak along costa, in 2 mixed with whitish, connected at base with
a white longitudinal median streak reaching to ? of disc ; two longi-
tudinal black streaks above and below middle, curved upwards
posteriorly, not reaching termen, upper interrupted near base and
more broadly posteriorly, lower indented above on veins, margined
beneath by a white streak curved up round its extremity, indented
beneath extremity of upper and continued round it toapex of wing ;
a slender dark grey terminal streak, in Q mixed with whitish : cilia
dark grey, more or less mixed with white, especially in 9. Hind-
wings blackish-grey, with faint pale post-median shade.
Three specimens ; Seaward Moss, Invercargill. Allied
to V. insignis, but very distinct.
Notoreas mechanitis, Meyr.
Three specimens sent of a well-marked local form
occurring on Mt. Holdsworth, Tararua Range, north of
Wellington, at 4000 feet, characterized as follows :
25-27 mm. (typical form does not exceed 23 mm.) ; duller, with
less yellow intermixture ; specially characterised by the obsolescence
of the black lines on under-surface of hind-wings, which are strongly
marked in all South Island forms.
Notoreas isoleuca, Meyr. (the locality for which should, I
am informed, be Castle Hill, not Wellington), is, as I am
now satisfied from further material, only a form of J.
mechanitis, the chief differences being due to a diminution
of the yellow suffusion. The species is however in all
its forms always quite distinct from NV. paradelpha, in
which the wings are differently formed, the fore-wings
being somewhat more elongate, with the costa straighter,
and the hind-wings obviously more elongate in proportion
to the fore-wings, the cilia sharply barred, the median
band of fore-wings narrower and differently formed, its
anterior margin much less curved, and posterior less
angulated in middle, and regularly indented above and
below middle.
Notoreas ischnocyma, sp. Nn.
Notoreas isolewca, Huds., N. Zeal. Moths, 72, pl. viii,
27, nec Meyr.
¢. 20mm. Head and thorax dark fuscous. Fore-wings trian-
gular, costa slightly arched, termen obliquely rounded ; dark fuscous,
with some scattered yellow-whitish scales ; subbasal, first, median,
229 Mr. E. Meyrick’s Notes on
second, and subterminal lines slender, whitish, subbasal straight,
first nearly straight, somewhat irregular, median indistinct, curved
outwards in disc, second waved throughout, angulated in middle,
indented beneath middle, subterminal irregularly waved: cilia
white, basal half barred with dark fuscous. Hind-wings with
ground colour, median, second, and subterminal lines, and cilia as
in fore-wings.
One specimen, Castle Hill. This is easily known from
the allied forms by the slender waved second line.
Notoreas omichlias, Meyr.
36 2. 22-27 mm. In fresh specimens the fore-wings show a
bluish-slaty gloss; in Mt. Holdsworth examples the edge of basal
patch and margins of median band are marked by slender light dull
ochreous fascie, but these are hardly perceptible in my other
specimens,
Seven specimens, Humboldt Range, and Mt. Holdsworth.
MONOCTENIADA.
' PARAGYRTIS, gen. n.
Face obliquely prominent. Tongue developed. Antennz %, in ¢
bipectinated, towards apex simple. Palpi moderately long, porrected,
rough-sealed. Thorax not hairy beneath. Fore-wings: 10 anas-
tomosing with 9 above 7. Hind-wings: 6 and 7 approximated, 8
approximated to cell to beyond middle.
Allied to Theoxena.
Paragyrtis inostentata, Walk.
Panagra inostentata, Walk. Cat. Geom. 1012; Dichro-
modes griseata, Huds. Trans. N. Zeal. Inst. 1902,
244, pl. xxx, 5.
& @. 15-23 mm. Head and thorax white, more or less irrorated
with fuscous. Fore-wings rather elongate-triangular, costa straight,
termen straight, rather oblique, strongly rounded beneath ; ground
colour formed by fine transverse rows of dark fuscous scales tipped
with white, appearing grey, sometimes broadly suffused with white
beneath costa and beyond second line; a black discal spot ;
second line represented by a sinuate dark fuscous shade from
beneath costa near apex to dorsum beyond middle, sometimes
broadened on lower portion, more usually faint or wholly obsolete ;
in one specimen a dark fuscous pretornal suffusion : cilia dark
New Zealand Lepidoptera. 223
fuscous, tips white, round tornus lighter. Hind-wings elongate,
varying from whitish to grey, towards termen usually darker-
suffused ; in the most strongly marked specimen there are suffused
darker post-median and preetornal shades as in fore-wings; cilia grey,
extreme tips white.
Seaward Moss, Invercargill; described from one New
Zealand and twelve Australian examples. This species
was accidentally omitted from my paper on the Australian
species of this family, but it has long been known to me,
being in fact one of the commoner Australian species,
occurring at Duaringa and Brisbane, Queensland ; Sydney
and Blackheath, New South Wales; Melbourne, Victoria ;
Blackwood, South Australia; Perth and Albany, West
Australia; from September to January. I have not
however previously seen it from New Zealand. It is
extremely variable in the depth of colouring and intensity
of marking.
SELIDOSEMID &.
Selidosema pungata, Feld.
Selidosema pungata, Feld., Reis. Nov., pl. cxxxi, 23 ; S.
Jascialata, Philpott, Trans. N. Zeal. Inst., 1902, 248,
BL xcs
¢. 40 mm. Head and thorax brownish-ochreous. Fore-wings
somewhat elongate-triangular, costa gently arched, termen bowed,
little oblique ; light ochreous-brown, with scattered short fuscous
strigula marked with a few black scales; a rather broad dark
fuscous median band, edged narrowly anteriorly and more broadly
and suffusedly posteriorly with ochreous-whitish, anterior edge
slightly curved, posterior rounded-prominent near costa and more
broadly below middle, thus concave above middle ; subterminal line
waved, ochreous-whitish, edged with dark fuscous suffusion, rather
broadly interrupted in middle, posterior marginal suffusion running
to termen beneath apex instead of to costa. Hind-wings light
ochreous-yellow, towards termen deeper and more ochreous.
One specimen; according to Mr. Philpott, this species
occurs in several localities in Southland in February and
March, and the female does not differ from the male ex-
cept in being somewhat paler. It is very similar to
S. productata, but constantly distinguished by the peculiar
form of the posterior margin of median band of fore-wings.
Not knowing of the existence of this species, I formerly
attributed Felder’s figure to productata,
bo
to
=
Mr. E. Meyrick’s Notes on
PHYCITIDA.
SPOROPHYLA, gen. n.
Face rounded; tongue developed. Antenne 3, in ¢ simple,
shortly ciliated, basal joint moderate. Labial palpi moderately
long, obliquely ascending, second joint much thickened with dense
scales, terminal joint short, obtuse. Maxillary palpi rudimentary.
Fore-wings: 4 absent, 3 and 5 connate, 8 and 9 stalked. Hind-
wings : 2 almost from angle of cell, 4 absent, 3 and 5 stalked, 6 and
7 connate, 8 closely approximated to cell and anterior portion of 7.
Allied to Crocydopora, but distinguishable by the simple
antennz of the male; the labial palpi are also shorter
and stouter.
Sporophyla anospora, Meyr.
Crocydopora enospora, Meyr., Trans. Ent. Soc. Lond.,
1897, 388.
Having now received additional specimens, including
the male, which was previously unknown, I am enabled
to characterize the genus as above, and also to add the
following particulars to my specific description :
¢ ?. 15-17 mm. Head and thorax dark grey mixed or densely
irrorated with white, sometimes mixed with ferruginous, Fore-
wings as described, but more usually without ferruginous admix-
ture, varying considerably in the preponderance of white or blackish
scales, in one specimen densely irrorated with white throughout so
as to obscure the markings. Hind-wings varying from grey to dark
fuscous, usually darker terminally.
Three additional specimens, Ida Valley, Otago, taken
by Mr. J. H. Lewis. This species is evidently very
variable,
CRAMBID/L.
Orocrambus machexristes, sp. D.
49. 21-24 mm. Head and thorax dark brown mixed with
blackish, centre of crown and of thorax anteriorly and in @ face
whitish. Palpi blackish-fuscous, whitish internally. Antenne dark
fuscous. Abdomen dark fuscous, somewhat whitish-sprinkled. Fore-
wings elongate, moderately dilated, costa hardly arched, apex obtuse,
termen rather oblique, straight, rounded beneath ; brown, more or
less mixed with dark grey and grey-whitish (more strongly in @) ;
New Zealand Lepidoptera. 225
an ochreous-white median longitudinal streak from base to 4, dilated
and obliquely truncate posteriorly, edged beneath by a thick black-
ish streak which curves up round posterior extremity, where it is
interrupted by interneural lines of ground colour, and continued as
a series of diminishing spots to near costa; a black subdorsal
streak from base to }; a curved series of blackish marks on veins
from ¢ of costa to tornus: cilia grey, with darker basal shade, tips
ochreous-whitish. Hind-wings dark grey, becoming blackish-grey
towards termen ; cilia as in fore-wings.
Two specimens, Mount Earnslaw (5300 feet). <A
striking and distinct species.
Crambus xthonellus, Meyr.
The female differs considerably from the male, and may
be described as follows:
9. 20mm. Fore-wings brassy-bronze, suffused with dark fuscous
in disc and on veins posteriorly ; a narrow white sub-costal streak
from base to near apex, sometimes interrupted obliquely beyond
middle, broader and more suffused posteriorly, connected at base
with a moderate straight white median streak from base to termen ;
a white dorsal streak from near base to tornus, dilated posteriorly :
cilia white, mixed with fuscous at apex and below middle of termen,
Hind-wings rather dark fuscous, darker posteriorly ; cilia whitish-
ochreous, with a fuscous line.
I took both sexes in company at Invercargill, and have
since received a female taken in the same district by
Mr. Philpott.
Crambus heteraulus, sp. n.
g. 35mm. Head white, behind eyes and in centre of face
brownish-ochreous. Palpi nearly 4, bronzy-ochreous, white above
and at base beneath. Antenne dark fuscous. Thorax bronzy-
ochreous, with broad white dorsal stripe. Abdomen whitish-
ochreous. Fore-wings elongate, moderately dilated, costa moder-
ately arched, apex nearly rectangular, termen obliquely rounded ;
brownish-ochreous, with strong brassy reflections ; a rather narrow
white median longitudinal streak from base to beyond 4, thence
continued as three undefined white interneural lines to termen,
edged with some darker scales posteriorly above, and towards middle
beneath; dorsum narrowly white towards base; two elongate
wedge-shaped white interneural streaks resting on termen beneath
apex, uppermast longer ; cilia white. Hind-wings fuscous-whitish ;
cilia white,
226 Mr. E. Meyrick’s Motes on
One specimen, Humboldt Range (3600 feet). Nearest
to C. crenwus, from which it ditfers by the fore-wings being
broader, with more arched costa, the median white streak
incomplete posteriorly, and the white interneural streaks
between this and apex; from all other nearly allied species
it differs by the absence of any white costal or sub-costal
streak,
Crambus sophistes, sp, 0.
d.17 mm, Head, palpi, and thorax fuscous, suffusedly mixed
with whitish ; palpi5. Antenne dark grey. Abdomen whitish,
basal half brassy, Fore-wings very elongate, narrow, gradually
dilated, costa gently arched, apex somewhat acute, termen rounded,
rather strongly oblique ; fuscous, irregularly irrorated with grey-
whitish ; a short suffused dark fuscous mark along dorsum towards
base ; a slender white streak, edged with scattered dark fuscous
scales, along fold from base to middle, thence acutely angulated in-
wards to near dorsum, this portion edged posteriorly with some dark
fuscous suffusion ; two dark fuscous dots longitudinally placed in disc
about 3; two sub-terminal series of short blackish dashes on veins,
and a third less complete on termen: cilia whitish, with two inter-
rupted fuscous lines. Hind-wings fuscous-whitish ; cilia ochreous-
white.
One specimen, Ida Valley, Dunedin, taken by Mr. J. H.
Lewis. The species is intermediate between C. cyclopicus
and harpophorus, but cannot be confused with either,
Argyria strophea, sp. n.
d @. 15-18 mm. Head and thorax ochreous-whitish, partially
yellowish-tinged, and sprinkled with dark grey. Palpi 4, grey,
darker-sprinkled, whitish above and towards base beneath. An-
tenne ochreous-whitish, obscurely ringed with dark fuscous.
Abdomen ochreous-whitish irrorated with dark grey. Fore-wings
elongate-triangular, costa gently arched, apex obtuse, termen slightly
rounded, oblique, faintly waved ; pale brassy-ochreous, suffusedly
mixed with white, and irrorated with dark grey ; subbasal line
white, edged anteriorly with dark fuscous, angulated near costa,
obsolete towards dorsum ; first and second lines white, more or less
edged with dark fuscous, first obtusely angulated above middle,
second sinuate inwards towards dorsum, preceded on costa by a small
dark fuscous spot ; a small roundish dark fuscous spot in disc before
middle; a narrow white transverse mark in disc beyond middle;
a terminal series of dark fuscous lunulate marks: cilia whitish, with
New Zealand Lepidoptera. 227
a fuscous subapical line, basal half barred with fuscous, Hind-wings
grey, darker posteriorly; cilia as in fore-wings.
Five specimens, Wellington. It is extremely distinct,
bearing very little resemblance to the other New Zealand
species, A. pentadactyla, yet probably more allied to it
than to any other known to me.
Tauroscopa trapezitis, sp. n.
¢. 22mm. Head and thorax blackish-fuscous mixed with white,
orbits and cheeks naked, orange. Palpi dark fuscous, white on
upper longitudinal half and at base beneath. Antenne blackish,
towards base whitish-sprinkled. Abdomen bronzy-fuscous sprinkled
with white. Fore-wings elongate-triangular, costa faintly sinuate,
apex rounded, termen rather obliquely rounded ; grey, partially
slightly ochreous-tinged, densely and suffusedly mixed with whitish;
markings blackish, somewhat irregular-edged ; a streak along basal
third of dorsum, and a streak from base of costa equal and parallel
to this ; two small costal spots before and one beyond middle ; an
oblique narrow-trapezoidal spot in dise before middle, its posterior
edge formed by whitish first line, which is very acutely angulated
below middle; a reniform discal spot slightly beyond middle ;
second line whitish, obtusely angulated in middle and indented
beneath costa, edged with black anteriorly and by an oblique
black spot on costa posteriorly ; a terminal series of small black
spots: cilia grey, with dark grey basal shade, tips white. Hind-
wings rather dark grey ; cilia grey, mixed with white towards tips.
One specimen, Mount Earnslaw, at 5300 ft. This, the
second discovered species of the endemic genus 7'awroscopa,
bears a general resemblance to 7’. gorgopis, but is very dis-
tinct, especially by the peculiarly formed first line ; whilst
the singular naked orange cheeks and orbits are quite a
unique feature, of which there is no trace in the other
species,
Diptychophora microdora, sp. n.
g 2. 10-12mm. Head, palpi, antenne, thorax, and abdomen
dark fuscous ; palpi mixed with yellowish towards base. Fore-
wings triangular, costa gently arched, apex obtuse, termen bisinuate,
oblique, rounded beneath ; dark fuscous, bronzy-tinged ; first. and
second lines dark, angulated, edged on costa with yellow strigule,
first edged on dorsal half anteriorly by two confluent yellow spots
and posteriorly by a yellow line, second edged posteriorly towards
228 Mr. E. Meyrick’s Notes on
dorsum by a wedge-shaped yellow mark ; a yellow costal strigula
just before apex, terminated by a white dot ; cilia dark grey, with
an irregular white basal line round apex and upper part of termen
(imperfect). Hind-wings and cilia dark fuscous.
I took a specimen on Mount Arthur at 3000 feet in
January, and have received a second from Mr. Hudson,
but neither is in very good condition. The species is
nearly allied to pyrsophanes, but much smaller, darker,
and with more developed yellow markings.
PYRAUSTID.
Scoparia meliturga, sp. 0D.
¢ 2. 17-19mm. Head and thorax ochreous-white irrorated with
dark fuscous. Palpi 3, dark fuscous, whitish-mixed, white towards
base beneath. Antenne dark fuscous, ciliations in ¢ 4. Abdomen
pale grey, more or less suffused with ochreous-whitish. Fore-wings
elongate-triangular, costa gently arched, apex obtuse, termen slightly
sinuate, rather oblique, rounded beneath ; ochreous-whitish, more
or less suffused with pale ochreous-yellowish in disc, sprinkled with
dark fuscous ; first line ochreous-white, edged posteriorly with dark
fuscous suffusion, slightly curved, indented in middle; orbicular
and claviform usually quite undefined or represented only by some
dark fuscous suffusion, seldom orbicular centred with an ochreous-
whitish dot; discal spot 8-shaped, ochreous-whitish or whitish-
ochreous, more or less outlined finely with dark fuscous ; second
line ochreous-white, edged anteriorly with dark fuscous, moderately
curved outwards from } to }, sharply indented at } ; terminal area
irrorated with dark fuscous, subterminal line thick, suffused,
ochreous-whitish, touching second line in middle, sometimes
slenderly interrupted above this; an interrupted ochreous-white
terminal line: cilia whitish, with dark grey interrupted anterior
and light grey posterior line. Hind-wings without long hairs in
dise ; pale whitish-grey, suffused with grey posteriorly ; cilia whitish,
with grey subbasal shade.
Six specimens, Auckland and Wellington, taken by
myself in December and January; I have also received
the species from Mr. Hudson. Until lately I had mistaken
this for a form of S. philerga, but am now satisfied of its
distinctness; it is a neater and smoother-looking insect,
always recognizable by the well-marked yellowish tinge ;
moreover in S. philerga the orbicular is normally white
New Zealand Lepidoptera. 229
edged with black, and the subterminal line is more broadly
interrupted.
Scoparia thyridias, sp. n.
dg. 18mm. Head fuscous. Palpi 2, fuscous, white towards base
beneath. Antennz dark fuscous, ciliations }. Thorax brownish,
mixed with whitish and dark fuscous. Abdomen whitish-fuscous.
Fore-wings very elongate-triangular, costa gently arched, apex
rounded-obtuse, termen rather oblique, rounded ; brownish, densely
and suffusedly irrorated with dark fuscous ; first line white, nearly
straight, rather oblique, irregular-edged, becoming double towards
dorsum; an oblong pellucid patch in middle of disc, preceded and
followed by blotches of orange suffusion, anterior extended down-
wards along first line ; second line slender, waved, white, curved
outwards from 7 to ?, forming a spot on dorsum ; subterminal line
very undefined, whitish, broadly interrupted above middle: cilia
whitish-fuscous, with two darker shades. Hind-wings without hairs
in disc ; light grey, darker terminally ; cilia as in fore-wings.
One specimen, locality not specified but probably
Wellington. This is distinguished from all other New
Zealand species by the naked discal patch (possibly not
developed in @); a similar structure is found in certain
Hawaiian species, but I think it improbable that they are
nearly allied specifically, and regard the character as having
been developed independently in each ‘case. 8. thyridias
belongs apparently to the neighbourhood of S. philerga.
Scoparia dochmia, sp. n.
¢. 20 mm. Head brownish-ochreous, somewhat whitish-sprinkled,
Palpi 3, dark fuscous, white towards base beneath. Antenne dark
fuscous, ciliations }. Thorax fuscous, whitish-sprinkled. Abdomen
pale ochreous-grey, whitish-mixed. Fore-wings elongate-triangular,
costa gently arched, apex obtuse, termen faintly sinuate, rather
oblique, rounded beneath ; light brownish irrorated with darker,
veins and margins suffusedly irrorated with blackish, median area
irrorated with white ; first and second lines well-marked, white,
interiorly blackish-edged, first somewhat curved, rather oblique,
sinuate in middle and rather abruptly rounded-prominent below
this, preceded in middle by a spot of white and blackish irroration ;
second parallel to termen, slightly curved from } to 2; orbicular
and elaviform forming undefined spots of blackish suffusion resting
on first line ; discal spot X-shaped, blackish, ill-defined ; subterminal
line indicated by whitish irroration, entire, approximated to termey,
230 Mr. E. Meyrick’s Notes on
a
and not touching second line: cilia whitish-fuscous, with two rather
dark fuscous shades. Hind-wings without long hairs in dise; pale
whitish-fuscous, with a slight brassy tinge ; a grey discal spot ; an
undefined grey postmedian line ; termen suffused with grey ; cilia
whitish, with fuscous subbasal shade.
One specimen, Lake Wakatipu, at 1300 feet. Although
an ordinary-looking insect, without striking characters,
this species does not in fact at all nearly approach any
other ; possibly it has most resemblance to the Australian
S. eremitis,
Scoparia triclera, sp. n.
Q. 12mm. Head, thorax, and abdomen dark grey sprinkled with
whitish. Palpi 2?, dark fuscous, white towards base beneath.
Antennz dark fuscous. Forewings elongate-triangular, costa poste-
riorly gently arched, apex obtuse, termen rather oblique, faintly
sinuate, rounded beneath ; blackish-fuscous mixed with whitish ; a
small pale ochreous elongated subbasal spot in middle; a broad
ochreous-white postmedian band parallel to termen, somewhat
narrowed towards costa, mixed with ochreous in disc ; subterminal
line undefined, whitish : cilia dark fuscous, somewhat whitish-mixed.
Hind-wings without long hairs in disc ; dark fuscous ; cilia fuscous,
base pale.
One specimen, Wellington. In size, shape and general
markings so like S. chlamydota that 1 should have thought
it one sex of that species, of which however I have both
sexes, not differing at all from one another; I am there-
fore obliged to regard it as distinct. It is easily separated
by the dark fuscous terminal area of fore-wings, and the
dark fuscous hind-wings.
Scoparia phalerias, sp. n.
@. 23mm. Head ochreous, sprinkled with dark fuscous. Palpi
2%, ochreous, mixed with dark fuscous, white towards base beneath.
Antenne ochreous spotted with dark fuscous. Thorax brown mixed
with dark fuscous, edged laterally and posteriorly with ochreous-
white. Abdomen light greyish-ochreous. Fore-wings elongate,
gradually dilated, costa slightly arched posteriorly, apex obtuse,
termen rather oblique, faintly sinuate, rounded beneath ; dark
fuscous, with fine seattered whitish scales ; a pale ochreous patch,
mixed with ochreous-brown, extending along costa from 4 to , and
reaching nearly half across wing; a small round pale ochreous sp o +
New Zealand Lepidoptera. 231
beneath this before middle, and a narrow elongate one beyond middle:
cilia ochreous-white. Hind-wings without hairs in disc; pale grey,
darker towards apex ; cilia whitish.
One specimen, Wellington, in April. Not at all like
any other, but perhaps allied to S. swbmarginalis.
Scoparia siderasprs, sp. n.
& 2. 25-28 mm. Head dark grey mixed with whitish. Palpi 3,
blackish mixed with white, towards base white beneath. Antenne
dark fuscous, in ¢ pubescent-ciliated on whole surface. Thorax
shining bluish-bronze. Abdomen bronzy-fuscous, in ¢ suffused
with yellow-ochreous on sides and posteriorly. Fore-wings elongate,
moderate, suboblong, in ¢ more dilated posteriorly ; shining slaty-
bluish fuscous ; a faint darker undefined discal spot: cilia fuscous.
Hind-wings with long hairs in disc ; fuscous, darker posteriorly, in
2 more or less tinged or suffused with ochreous-yellowish anteriorly ;
cilia whitish-yellowish, with two fuscous lines. Under-surface,
especially of hind-wings, more or less wholly suffused with ochreous-
yellow.
Three specimens, Mount Earnslaw (5300 feet) and
Humboldt Range. The slaty-bluish colouring recalls 8.
cataxesta, to which however the species is not really allied,
but rather to the neighbourhood of S. nomentis, though
very distinct ; the ochreous-yellow suffusion of the under-
surface is a peculiar characteristic.
PTEROPHORID.
Platyptilia epotis, sp. n.
9. 25mm. Head white. Palpi 24, greyish-ochreous sprinkled
2d
with dark fuscous, white above. Antenne grey. Thorax whitish
suffusedly sprinkled with brownish-ochreous, patagia becoming clear
white posteriorly. Abdomen white, partially suffused with pale
brownish-ochreous except towards base. Legs white, anterior femora
and tibie fuscous. Fore-wings with apex pointed, subfalcate, termen
rather deeply concave ; pale brownish-ochreous densely irrorated
with white, broadly suffused with white towards costa and dorsum
on posterior half ; basal } of costa irrorated with dark fuscous, thence
a narrow dark fuscous costal streak reaching to #; a dot of two or
three dark fuscous scales in disc at 2; a short transverse blackish-
fuscous mark before cleft; a strong oblique blackish streak from
apex to anterior half of lower margin of first segment, where it is
232 Mr. E. Meyrick’s Notes on
broadest ; an undefined blackish-fuscous longitudinal dash in centre
of basal half of second segment: cilia whitish, with a dark fuscous
spot above apex, an interrupted blackish basal line on central third
of termen, and two or three blackish scales on dorsum at 3. Hind-
wings pale fuscous, second segment whitish-tinged ; cilia whitish-
fuscous ; two or three fine blackish hair scales in dorsal cilia of third
segment beyond middle.
One specimen, Humboldt Range, at 3600 feet. The
stalk of veins 8 and 9 of fore-wings is very short. The
species resembles generally Stenoptilia leucoxesta and S.
charadrias (probably with real genetic affinity), but is
distinct from everything by the well-defined blackish
subapical bar.
EPIBLEMID.
Noteraula sideritis, sp. n.
¢@. 15mm. Head aud thorax brownish-ochreous. Palpi moder-
ately long (23), ochreous-brown. Antennze whitish-ochreous, with
dark fuscous serrations. Abdomen grey, darker posteriorly, anal
tuft whitish-ochreous. Fore-wings elongate, somewhat dilated
posteriorly, costa moderately arched, apex obtuse, termen slightly
sinuate, rather oblique, rounded beneath ; whitish-ochreous finely
strigulated with dark fuscous, mixed in dise and towards apex
wholly suffused with ferruginous-ochreous ; two curved posterior
oblique purplish-leaden-metallic striz from costa at 3 and 3, terminat-
ing before and beyond tornus ; a series of undefined blackish marks
between these, starting from an oblique leaden-metallic costal streak ;
a short direct leaden-metallic costal streak before apex : cilia grey,
basal third with a blackish-grey line broadly interrupted with
brownish-ochreous, at tornus whitish-ochreous, Hind-wings grey,
terminal edge whitish ; cilia light grey, with darker basal line.
One specimen, Wellington (?). The discovery of a
second species of this genus is interesting ; the palpi are
much shorter and the termen of fore-wings much less
oblique than in J. stramanea.
Strepsicrates dolopxa, sp. n.
¢. 18 mm. Head, palpi, and thorax pale greyish-ochreous
irrorated with whitish. Antenne grey, suffused with whitish above,
notch at about } from basal joint. Abdomen pale greyish-ochreous.
Fore-wings elongate, narrow, costa moderately arched, apex round-
pointed, termen sinuate, oblique, rounded beneath, costal fold reach-
New Zealand Lepidoptera. 233
ing 2; pale greyish-ochreous, irrorated with whitish and strigulated
with fuscous, posteriorly more ochreous; an undefined patch of
fuscous suffusion extending along costa from 3 to ¢; margins of
ocellus, and an angulated stria beyond it leaden-metallic : cilia
pale grey irrorated with whitish. Hind-wings with 3 and 4 coinci-
dent ; whitish-grey ; cilia grey-whitish, with faint grey subbasal
shade.
One specimen, Wellington, Structurally similar to
S. zopherana, but much paler, and without any of the
dark markings of that species.
TORTRICIDA.
Dipterina hemiclista, sp. n.
¢. 15 mm. Head, palpi, and thorax grey mixed with dark
fuscous, palpi short, 13. Fore-wings elongate, considerably dilated
posteriorly, costa gently arched, apex obtuse, termen obliquely
rounded ; rather dark slaty-grey, with irregular angulated dark
fuscous striz, tending to break up into strigule ; edge of basal patch
dark fuscous, right-angled in middle ; an oblique dark fuscous blotch
on costa before middle, another at #, and a third apical: cilia grey
mixed with dark fuscous. Hind-wings fuscous, darker posteriorly.
One specimen, Wellington. Not like any of the other
New Zealand species (which also differ much among them-
selves), but with considerable resemblance to an undescribed
Tasmanian species.
Eurythecta zelea, sp. n.
@. 10-11 mm. Head ochreous, face and a central longitudinal
streak on crown suffused with white. Palpi 2, white, externally
suffused with ochreous. Antenne dark grey. Thorax fuscous mixed
with ochreous and whitish. Abdomen fuscous. Fore-wings lanceo-
late, round-pointed, obviously more elongate and narrower than in F.
robusta ; fuscous, more or less mixed irregularly with pale yellowish-
ochreous and white ; costa suffused with white, and marked with a
quadrate dark fuscous spot before middle, and three smaller posterior
spots ; a dark fuscous dorsal spot before middle, another on tornus,
and several more or less defined dots on termen: cilia fuscous, basal
third mixed with whitish, tips pale. Hind-wings rather dark fuscous,
darker posteriorly ; cilia fuscous, paler towards tips.
Two specimens, Ida valley, Otago, taken by Mr. J. H.
Lewis. This is closely allied to #. robusta, but differs in
234 Mr. E. Meyrick’s Notes on
the absence of the two dark fasciz, which though often
interrupted are always present and well-marked as fascize
in H. robusta, whereas in LH. zelea they are represented only
by small costal and dorsal spots; and also structurally in
the considerably narrower and more elongate fore-wings.
LE. robusta may be extinct; I believe it has not been seen
for very many years, though formerly locally abundant.
ASCERODES, gen. n.
Palpi moderately long, porrected, triangularly scaled with long
rough projecting hairs. Antenne in ¢ biciliated with fascicles of
long cilia. Thorax without crest. Forewings in ¢ without costal
fold ; 7 and 8 separate. Hind-wings without pecten of hairs on
lower margin of cell; 3 and 4 approximated at base, 5 somewhat
approximated to 4, 6 and 7 approximated at base.
Apparently most allied to Harmologa, from which it
differs by the absence of the costal fold; separable from
Proselena and Prothelymna by veins 6 and 7 of hind-wings
not being stalked.
Ascerodes prochlora, sp. n.
g. 18 mm. Head, palpi, and thorax dark fuscous mixed with
ferruginous-orange, palpi 3. Antenne blackish spotted with white,
ciliations 3. Abdomen dark grey, with whitish lateral streaks.
Fore-wings elongate, suboblong, costa moderately arched towards
base and apex, apex rounded, termen somewhat rounded, rather
oblique ; dark grey, densely overlaid with ferruginous-brown ; costa
broadly and dorsum narrowly suffused with pale yellowish-ochreous :
cilia ochreous-whitish, basal half suffused with dark grey. Hind-
wings dark grey, more blackish-grey posteriorly ; cilia whitish, basal
half greyish-tinged, with blackish-grey basal line.
One specimen, Humboldt range, at 4000 feet.
Tortrix indigestana, Meyr.
Tortriz indigestana, Meyr., Proc. Linn. Soc. N.S Wales,
1881, 520.
Of this widely distributed Australian species, not
previously recorded from New Zealand, I took a good
specimen at Whangarei in December, which I cannot
distinguish from Austrahan examples.
New Zealand Lepidoptera. 235
PHALONIAD.
Heterocrossa contactella, Walk.
Tinea contactella, Walk. Suppl. 1813.
‘9. 14-15 mm. Head white. Palpi 24, white, lower longitudinal
half blackish. Antenne white, obscurely ringed with fuscous.
Thorax whitish-ochreous mixed with dark grey, collar and patagia
white. Abdomen pale greyish-ochreous. Fore-wings elongate,
narrow, costa moderately arched, apex round-pointed, termen very
oblique, almost straight, rounded beneath; grey, irregularly irrorated
with dark grey and white ; a broad irregular-edged white suffusion
extending along anterior half of costa, and reaching 3 across wing ;
three or four small black dots on costa anteriorly, second forming a
short strigula ; a narrow oblique-transverse pale ochreous spot edged
with black below middle at }; a black dot above middle of disc, and
a small pale ochreous sometimes blackish-mixed spot below it ; three
small faint whitish-ochreous spots arranged in a triangle in disc
beyond middle; all these ochreous spots are ringed with white
suffusion ; an undefined angulated dark subterminal shade, marked
with black on veins ; a series of blackish dots on posterior half of
costa and termen: cilia light grey irrorated with white, basal half
obscurely barred with whitish. Hind-wings grey, paler anteriorly ;
cilia whitish.
Two specimens, Wellington. I have not actually com-
pared these specimens with Walker’s types, but from notes
I made on a previous inspection J think there is little
doubt of their identity; the species had not hitherto been
rediscovered. It is intermediate between adreptella and
the rest.
(ECOPHORID A.
Hypercallia aletis, sp. n.
¢d.13 mm. Head and thorax light fuscous sprinkled with
whitish-ochreous. Palpi whitish-ochreous, a subapical ring of second
joint, and terminal joint except apex somewhat infuscated. Antenne
greyish-ochreous, ciliations 3. Abdomen fuscous, Fore-wings elon-
gate, moderate, costa gently arched, apex round-pointed, termen
somewhat rounded, rather strongly oblique; greyish-ochreous
irrorated with fuscous ; some dark fuscous scales towards base of
costa ; first discal and plical stigmata very obscure, darker, plical
rather obliquely beyond first discal; second discal distinct, dark
fuscous, with some whitish scales beneath it: cilia greyish-ochreous
TRANS. ENT. SOC. LOND. 1905.—PART II. (JULY) 16
236 Mr. E. Meyrick’s Notes on
mixed with fuscous. Hind-wings rather dark fuscous, lighter ©
anteriorly ; cilia light fuscous, with darker subbasal shade, tips
whitish.
I took one specimen on Arthur’s Pass at 3000 feet, in
January. It is an insect of the most obscure appearance,
probably allied to the other New Zealand species of the
genus, H. amenena, but differing obviously in the very
much smaller size and dark hind-wings. These two out-
liers of a characteristic Australian genus are probably
amongst the few indications of an immigration by way of
Tasmania.
Proteodes profunda, sp. ua.
¢. 20mm. Head ochreous, mixed on sides with dark fuscous.
Palpi dark fuscous, extreme apex of second and terminal joints pale
ochreous. Antenne fuscous. _Thorax dark fuscous, somewhat
mixed partially with whitish-ochreous, Abdomen dark fuscous
mixed with ochreous-whitish ; at base a pair of oblique narrow
bladder-like membranes above. Fore-wings broad, suboblong,
rather dilated posteriorly, costa moderately arched, apex obtuse,
termen faintly sinuate, little oblique, rounded beneath ; fuscous,
with series of cloudy dots of black irroration on veins; markings
formed by whitish-ochreous suffusion, irregularly sprinkled with
dark ferruginous ; about six irregular subconfluent spots towards
base and anterior half of costa; a triangular subterminal patch
extending from apex to tornus and leaving a narrow terminal streak
of ground colour, its apex extending inwards to lower angle of cell ;
small discal spots of blackish suffusion before and beyond middle :
cilia whitish-fuscous, basal half mixed with dark fuscous and whitish-
ochreous, on costa with four pale dots. Hind-wings grey ; cilia
whitish-grey.
One specimen, Mount Holdsworth, in forest, at 2000
feet. This, the second discovered species of Proteodes,
differs a good deal superficially from P. carnifex, and has
a very Tortriciform appearance.
Trachypepla pheoptila, sp. n.
g. 15mm. Head ochreous-white, face and crown mixed with
dark fuscous. Palpi white, basal third and a subapical band of
second joint, and two bands on terminal joint blackish. Antenne
dark grey, ciliations #,even. Thorax fuscous sprinkled with whitish,
mixed with dark fuscous anteriorly. Abdomen rather dark fuscous.
New Zealand Lepidoptera. 237
Fore-wings elongate, costa moderately arched, apex obtuse, termen
oblique, rounded beneath ; fuscous, irregularly strewn with white,
ochreous-brown, and black scales; markings very undefined ; the
black scales indicate an angulated basal patch, a large irregular
blotch on middle of costa, some raised tufts in disc, and an apical
patch ; the anterior edge of the costal blotch is margined with white
suffusion ; an obscure roundish spot of brownish-ochreous suffusion
beneath dise beyond middle : cilia fuscous, basal half mixed with
white and blackish. Hind-wings dark fuscous ; cilia fuscous, with
dark fuscous subbasal shade.
One specimen, Mangaterera River, Mount Holdsworth.
Comparable with 7’. anastrella from the dark fuscous hind-
wings, but apart from the more blackish mixture of fore-
wings (which are also more elongate), differmg widely in
the antennal ciliations of ¢, which are short (#) and even,
whilst in anastrella they are long (3) and whorled; this
character 1s not noticed in my description of anastrella.
The length of ciliations differs considerably in various
species of Z’rachypepla, and appears to afford here specific
characters only.
Trachypepla lathriopa, sp. n.
gd. 14-16 mm. Head and thorax brown. Palpi brownish-
ochreous, sprinkled with dark fuscous, extreme tips of second and
terminal joints whitish. Antenne dark fuscous, ciliations 14, even.
Abdomen rather dark fuscous. Fore-wings elongate, costa moder-
ately arched, apex round-pointed, termen very obliquely rounded ;
light reddish-fuscous, irregularly sprinkled with brown aud dark
fuscous ; edge of basal patch indicated by a very obscure pale acutely
angulated narrow fascia; stigmata. dark fuscous, very undefined,
plical rather obliquely beyond first discal ; a subterminal series of
undefined dark fuscous dots, indented beneath costa; a series of
undefined dark fuscous dots along posterior part of costa and termen :
cilia whitish-fuscous, tinged with reddish, with a grey postmedian
shade. Hind-wings dark fuscous, lighter towards base ; cilia grey,
with dark grey basal shade.
Six specimens, taken by myself at Wellington, Nelson,
and on the Mt. Arthur plateau in January, and also
received from Mr. Hudson. Very like obscure examples
of anastrella, with which I have hitherto confused it, but
obviously longer-winged, and certainly distinct by the
much shorter and evenly-arranged ciliations of antenne.
238 Mr. E. Meyrick’s Motes on
Izatha (Semiocosma) metadelta, sp. n.
¢. 17mm, ¢. 19-25 mm. Head fuscous sprinkled with whitish,
with well-marked conical horny frontal prominence concealed in
scales. Palpi white, second joint mixed with dark fuscous, with
dark fuscous basal and subapical bands, terminal joint mixed with dark
fuscous at base, with dark fuscous median band. Antenne dark
fuscous ringed with whitish. Thorax whitish, mixed with fuscous
in ¢, on patagia ochreous-tinged, anteriorly suffused with dark
fuscous. Abdomen dark fuscous mixed with whitish, two basal
segments ferruginous-ochreous. Fore-wings elongate, moderate,
costa gently arched, apex obtuse, termen straight, rather oblique,
rounded beneath ; brownish-ochreous or fuscous, more or less
sprinkled with whitish and dark fuscous, darkest in ¢, towards costa
and dorsum suffused with whitish in ? ; dark fuscous marks at base
of costa and dorsum ; a small fuscous spot on costa at }, and a larger
triangular one on dorsum at 3, latter followed in 9 by a patch of
ochreous suffusion ; a dark fuscous subquadrate spot on costa before
middle, and a larger subtriangular one extending on costa from
middle to }; two dark fuscous transverse discal spots before and
beyond middle, touching these costal spots respectively, partially
edged with black and then with white, first suboval, mostly filled
with raised scales, second curved-reniform, lower posterior margin
broadly interrupted ; in ¢ two undefined longitudinal discal black-
ish streaks traversing these spots; an ill-defined cloudy whitish
curved subterminal line; a terminal series of small dark fuscous
spots: cilia fuscous sprinkled with whitish. Hind-wings in ¢ black-
ish-fuscous, in 2 dark fuscous, lighter anteriorly ; cilia fuscous, with
darker basal shade.
Three specimens, Wellington. Nearest to I. epiphanes,
but very distinct ; the conical horny frontal prominence is
a noticeable structure; it recurs (though previously un-
observed because slight and concealed) in a less developed
form in some but not all of the other species, and is
doubtless adaptive.
Gymnobathra bryaula, sp. n.
¢. mm. Head and thorax ochreous-whitish, somewhat mixed
with pale greenish-yellow, thorax with two small anterior dark
fuscous spots. Palpi whitish, base and a subapical ring of second
joint, and subbasal and subapical rings of terminal joint dark
fuscous. Antenne ochreous-whitish, basal joint dark fuscous above,
ciliations 2. Abdomen ochreous-whitish, second segment more
New Zealand Lepidoptera. 239
ochreous. Fore-wings elongate, somewhat dilated posteriorly, costa
moderately arched, apex obtuse, termen straight, oblique, rounded
beneath ; ochreous-whitish, mixed with light greenish-yellow and
sprinkled with dark fuscous, especially on veins ; markings fuscous
mixed with dark fuscous ; small spots on base of costa and dorsum,
a small double spot near base in middle, and three small double spots
representing stigmata, plical obliquely beyond first discal ; two
elongate marks above tornus ; a subterminal series of small subcon-
fluent spots, approximated to termen, acutely indented beneath apex;
five small spots on posterior half of costa: cilia ochreous-whitish,
basal half spotted with dark fuscous (imperfect). Hind-wings
ochreous-grey-whitish ; cilia whitish, with grey basal dots on veins.
One specimen, Wellington. Very distinct; might be
placed between sarcoxantha and thetodes.
Lorkhausenia (Ccophora) penthalea, sp. n.
d 9. 15-17 mm. Head pale brownish-ochreous, whitish-
sprinkled. Palpi ochreous-whitish, second joint with basal half
dark fuscous and a subapical ring of dark fuscous irroration, terminal
joint with subbasal and subapical rings of dark fuscous irroration.
Antenne grey, ciliations of ¢ 1. Thorax pale brownish-ochreous
sprinkled with fuscous, anteriorly suffused with dark fuscous.
Abdomen light grey. Fore-wings elongate, costa moderately arched,
apex obtuse, termen very obliquely rounded ; whitish-ochreous,
irrorated with light brownish, with some dark fuscous scales towards
margins, sometimes forming distinct patches of dark suffusion on
costa at base, +, and middle ; an oblique dark fuscous mark above
dorsum before middle; discal stigmata crescentic, dark fuscous,
hollow beneath, often ill-defined ; a moderately broad subterminal
fuscous or brown fascia, irrorated with dark fuscous, narrowed
towards costa and on tornus: cilia whitish-ochreous irrorated with
fuscous, at tornus with a darker bar above a pale spot. Hind-wings
light grey, darker towards apex ; cilia grey-whitish, with grey basal
shade.
Three specimens, Wellington. Allied to griseata; readily
distinguished by the smaller size and well-defined dark
fuscous subterminal fascia.
Borkhausenia chloradelpha, sp. n.
¢ 9. 18-20mm. Head whitish-ochreous. Palpi ochreous-whit-
ish, basal half of second joint sometimes irrorated with dark fuscous.
Antenne whitish ringed with dark fuscous, ciliations of ¢ 1,
240 Mr. E. Meyrick’s Notes on
Thorax whitish-ochreous, with brown subdorsal stripes. Abdomen
ochreous-whitish. Fore-wings elongate, costa moderately arched,
apex pointed, termen faintly sinuate, rather strongly oblique ; pale
whitish-ochreous, along dorsum and on posterior half more or less
partially brownish-tinged and sprinkled with brown and dark brown ;
a suffused brown streak, mixed with dark brown, along submedian
fold from base to tornus ; stigmata fuscous, often indistinct, plical
obliquely beyond first discal ; a subterminal line of dark fuscous
scales very indefinitely indicated : cilia pale whitish-ochreous, more
or less sprinkled with fuscous. Hind-wings and cilia ochreous-
whitish.
Four specimens, Wellington. Intermediate between
griseata and politis, but easily distinguished from both by
the whitish-ochreous colouring, brown submedian streak,
and whitish hind-wings.
Borkhausenia thranias, sp. n.
¢.10 mm. Head and thorax orange-yellow. Palpi whitish-
yellow, second joint irrorated with dark fuscous except towards
apex. Antennee dark fuscous, ciliations 1. Abdomen light silvery-
grey. Fore-wings elongate, costa moderately arched, apex round-
pointed, termen very obliquely rounded ; deep yellow, towards base
orange-tinged ; a well-defined rather dark fuscous streak along costa
from base to middle ; a few fuscous scales in dise beyond middle
and towards tornns: cilia light ochreous-yellow. Hind-wings light
grey ; cilia whitish-grey-ochreous.
‘One specimen, taken by myself at Whangarei in
December. Nearest horwa; characterised by small size,
well-defined costal streak reaching middle, and absence
of dark colouring on thorax.
Borkhausenia melanamma, sp. n.
@. 12-14 mm. Head, palpi, and thorax grey irrorated with white.
Antenne dark grey ringed with whitish, ciliations 1. Abdomen
grey, more or less mixed with ochreous-yellowish. Fore-wings
lanceolate, costa bent at 4; fuscous, irrorated with white, with a few
seattered dark fuscous scales; four oblique fasciz of dark fuscous
irroration from costa at base, 4, ?, and $ indicated or obsolete : cilia
light fuscous, some irroration and tips whitish. Hind-wings grey;
cilia light grey, with darker basal shade.
Two specimens, Ida Valley, Otago, taken by Mr. J. H.
Lewis. Near siderodeta; similar in form of wing but
differs by the white irroration (causing grey appearance),
New Zealand Lepidoptera. 241
absence of ochreous-yellow or ferruginous suffusion, and
lighter hind-wings. Both species evidently vary con-
siderably, but I have taken siderodeta in plenty, and in
all its forms it is clearly distinct from melanamma.,
Borkhausenia loxotis, sp. n.
dg. 11-12 mm. Head and palpi dark fuscous irrorated with
ochreous-whitish, Antenne dark fuscous, pale-ringed. Thorax and
abdomen dark fuscous. Fore-wings elongate, costa moderately
arched, apex round-pointed, termen very obliquely rounded ; dark
fuscous ; some scattered pale yellowish scales along submedian fold ;
a narrow straight pale ochreous-yellowish fascia, edged with some
black scales, from } of costa to 2 of dorsum ; a pale ochreous-yellow
dot in dise at ?, and sometimes others on costa beyond middle and
at tornus ; a more or less indicated subterminal line of pale yellowish
scales, starting from a small costal spot: cilia fuscous, irrorated with
yellow-whitish. Hind-wings dark fuscous ; cilia fuscous, with
darker basal shade.
I took one specimen at Wellington in January, and
have received another from Mr. Hudson. It is inter-
mediate between siderodeta and chryssgramma, but quite
distinct from either.
Borkhausenra pharmactis, sp. n.
g. 15mm. Head fuscous, sprinkled with pale yellowish hairs.
Palpi whitish-yellowish, sprinkled with dark fuscous. Antenne
whitish-yellowish, ringed with dark fuscous. Thorax ochreous-
yellow, anteriorly suffused with fuscous. Abdomen grey, Fore-
wings elongate, costa gently arched, apex obtuse, termen slightly
rounded, rather strongly oblique; deep ochreous-yellow, finely
sprinkled throughout with brown; extreme costal edge dark
fuscous towards base: cilia ochreous-yellow, somewhat sprinkled
with brown. Hind-wings dark grey ; cilia whitish-grey, with grey
basal shade,
I took one specimen on the Mount Arthur plateau, at
an elevation of 4000 feet, in January. Perhaps allied to
anema, but much deeper-coloured, and quite without the
dark costal streak and other markings.
PLUTELLIDA.
PHYLACODES, gen. n.
Head with appressed scales; ocelli present; tongue developed.
Antenne 3, in ¢ shortly ciliated, thickened above with scales on
242 Mr. E. Meyrick’s Notes on
basal half, basal joint moderate, without pecten. Labial palpi long,
recurved, with appressed scales, second joint somewhat roughened
beneath towards apex, terminal joint longer than second, acute.
Maxillary palpi short, slender, acute, porrected. Posterior tibize
smooth-scaled above, loosely haired on apical half beneath. Fore-
wings: 2 from very near angle of cell, 7 and 8 stalked, 7 to termen,
11 from middle of cell. Hind-wings slightly over 1, subtrapezoidal,
termen slightly sinuate, cilia #; 3 and 4 rather approximated at
base, 5, 6, 7 parallel.
Not obviously near to any described genus.
Phylacodes cauta, sp. n.
¢.°14mm,. Head and thorax light brownish-ochreous, slightly
purplish-tinged. Palpi whitish-ochreous, slightly fuscous-sprinkled,
especially beneath apex of second joint. Antenne pale greyish-
ochreous. Abdomen dark grey, anal tuft'white. Fore-wings elongate,
narrow, costa gently arched, apex round-pointed, termen very
obliquely rounded ; light greyish-ochreous, on costal half suffused
with ochreons-white, especially on veins ; several minute black dots
along submedian fold, dorsum, and termen, and towards apex ; a
larger round dot in dise at #: cilia ochreous-whitish. Hind-wings
grey, becoming blackish-grey towards apex ; cilia light grey, with
dark grey basal shade, tips whitish.
One specimen, Ida Valley, Otago, taken by Mr. J. H. Lewis.
Orthenches drosochalea, sp. n.
@4@.11mm. Head and thorax leaden-grey. Palpi dark fuscous,
inwardly and towards base white, terminal joint half as long again
as second (14), Antenne dark fuscous, ringed with white. Abdomen
grey, towards base pale ochreous, apex in ¢ whitish. Fore-wings
elongate, costa moderately arched, apex pointed, termen sinuate,
rather strongly oblique, rounded beneath ; 7 to termen; shining
coppery-bronze; four oblique fascia of white irroration, first
slender, second antemedian, broader, third angulated, considerably
enlarged towards costa, fourth forming an apical patch extended
along termen ; a spot on base of costa, an interrupted streak along
submedian fold, and a spot above middle of dise purple ; a dark
fuscous-purple transverse mark in dise at %,in third fascia: cilia
light grey, above apex spotted basally with purplish. Hind-wings
ovate-lanceolate, apex acute, termen sinuate; light grey, darker
posteriorly ; cilia whitish-grey.
I took a specimen in the Otira Gorge in January, and
have received another from Mr, Hudson, taken in Welling-
New Zealand Lepidoptera. 243
ton. Allied to porphyritis, and similar in form of wing,
but structurally distinct by vein 7 of fore-wings run-
ning to termen, not apex, and terminal joint of palpi 13
instead of 2, as well as by the clear coppery-bronze colour-
ing and lighter hind-wings.
Compsistis orthophanes, sp. n.
9. 9-10 mm. Head, antennex, and thorax dark bronzy-fuscous.
Palpi rather dark fuscous, internally whitish-ochreous, terminal
joint as long as second. Abdomen dark fuscous, apex whitish-
ochreous. Fore-wings elongate, narrow, costa gently arched, apex
round-pointed, termen very obliquely rounded; fuscous, slightly
purplish-tinged, irrorated with dark fuscous and blackish ; a straight
rather narrow fascia at }, a small spot in middle of disc, and opposite
subcostal and subdorsal spots at } pale ochreous-yellow, irregular-
edged : cilia fuscous, mixed with dark fuscous towards base. Hind-
wings dark fuscous ; cilia fuscous, with dark fuscous basal shade.
Two specimens taken by myself at Auckland and Nelson,
in December and January.
TINEIDZ.
Dryadaula myrrhina, sp. 0.
g¢. 9mm. Head, palpi, antenne, and thorax ochreous-whitish.
(Abdomen broken.) Fore-wings elongate, rather narrow, costa
gently arched, apex round-pointed, termen faintly sinuate, extremely
oblique; 7 and 8 separate ; shining, whitish-yellowish ; two very
oblique triangular fuscous spots on costa about } and before middle ;
a fuseous dot above dorsum at +; an inwardly oblique fuscous strigula
on dorsum at 2; a fuscous costal dot at 3}; some indistinct brownish
suffusion on dorsal half posteriorly and towards apex; a_ thick
black terminal line, interrupted below apex and in middle: cilia
whitish-yellowish. Hind-wings trapezoidal, termen faintly sinuate
below apex ; light grey ; cilia grey-whitish.
One specimen, probably from Wellington. Nearly allied
to the Australian D. glycinopa, under which name I now
believe I confused two distinct species; I therefore add
diagnoses of these, to show that all three are distinct.
Dryadaula glycinopa, Meyr.
Dryadaula glycinopa, Meyr., Proc. Linn. Soc. N.S.
Wales, 1892, 559 (part).
¢. 9-10. (Antenne broken.) Fore-wings with 7 and 8 stalked ;
shining whitish-yellowish ; very oblique ochreous-yellow fascia-like
244 Mr. E. Meyrick’s Notes on New Zealand Lepidoptera
streaks from costa at 1 and before middle, and a blotch on costa
about 2, all marked with dark fuscous on costal edge ; a suffused
ochreous-yellow streak along submedian fold, and indications of two
or three oblique streaks from dorsum coalescing with it ; an incom-
plete fine dark fuscous line from about ? of disc to apex ; an irregular
blackish line along termen, tending to be interrupted below apex and
in middle. Hind-wings subtrapezoidal, termen not sinuate ; grey,
lighter toward base.
Two specimens, Blackheath, New South Wales, in
February.
Dryadaula napa, sp. n.
Dryadaula glycinopa, Meyr., Proc. Linn. Soc. N.S.
Wales, 1892, 559 (part).
9.8mm. Antenne towards apex with three dark fuscous bands.
Fore-wings with 7 and 8 short-stalked; white; very oblique
ochreous-yellow fascia-like streaks from costa at { and 4, marked
with fuscous on costa, meeting in dise two shorter oblique ochreous-
yellow streaks from dorsum ; a deep ochreous-yellow streak along
submedian fold from base ; a triangular fuscous blotch extending
on costa from middle to 4, connected with termen by a yellow-
ochreous suffusion ; a fuscous spot on costa near apex ; an incomplete
line of dark fuscous scales from 2 of dise to termen beneath apex,
space beneath this mostly suffused with yellow-ochreous ; blackish
dots at apex and tornus, connected by an incomplete row of blackish
scales along termen. Hind-wings broadly lanceolate, pale grey.
One specimen, Deloraine, Tasmania, in November,
( 245 )
XII. On three remarkable new genera of Microlepidoptera.
By Sir Greorce F, Hampson, Bart., B.A.
[Read March Ist, 1905,]
PYRALIDA.
CHRYSAUGINA.
Genus PACHYPODISTES, nov.
Proboscis aborted ; palpi not projecting beyond frons and very
thickly clothed with hair; maxilliary palpi absent; frons with
thick tuft of hair; antenne ciliated ; legs of male thickly clothed
with hair, the fore tarsi fringed with hair, the mid and hind tarsi
with immense tufts of hair on upper-side, the spurs absent, female with
the tibize and tarsi smoothly scaled and very much swollen, the tarsal
joints indistinct, the spurs absent ; abdomen very long, especially in
female where the anal segment is twice the length of the others.
Fore-wing with the costa concave and the apex produced upwards,
especially in male, the termen strongly excurved ; vein 3 from angle
of cell; 4-5 stalked in male, approximated for some distance in
female ; the upper angle of cell produced, especially in male, 6 from
angle in male, in female shortly stalked with 7, 8, 9,10; 11 from
cell. Hind-wing with the inner angle thickly clothed with hair ;
vein 3 from angle of cell; 4,5 stalked in male, approximated for
some distance in female ; 6, 7 stalked, 7 anastomosing with 8.
The genus is best placed in the Chrysauginz in spite of
the absence of proboscis.
Pachypodistes goeldit, n. sp.
¢. Head and thorax greyish-ochreous mixed with fuscous-brown ;
patagia dull vinous-red irrorated with grey and edged with ochreous ;
hind tarsi tinged with vinous-red, the fringe of hair dark brown at
tips; abdomen ochreous-brown, the ventral surface tinged with
vinous-red. Fore-wing vinous-red irrorated with greyish-ochreous
especially on costa and inner margin ; an oblique grey antemedial
line joined at inner margin by a very obliquely curved line from
costa near apex below which it is slightly angled ; cilia brownish-
ochreous, Hind-wing vinous-red irrorated with greyish-ochreous,
TRANS, ENT, SOC. LOND, 1905.—PART II. (JULY)
9
od
46 Sir G. F. Hampson on three
an indistinct very obliquely curved postmedial greyish line ; cilia
brownish-ochreous ; the inner margin clothed with long brownish
hair.
?. Wings much brighter vinous-pink ; fore-wing with the post-
Pachypodistes goeldti, +
medial line oblique, not curved and well separated from the ante-
medial line at inner margin.
Larva a whitish stoutly-built grub with red-brown head, evidently
an internal feeder. Cocoon; red-brown, flattened, bivalvular, formed
remarkable new genera of Microlepidoptera. 247
in layers and rather resembling a mussel-shell, probably attached
to bark or between leaves.
Hab, AMAZONS, Pard (Goeldi), 1 f, 2 2, larva, pupa and
cocoons. Exp. ¢ 38,250 mm. Type in B. M.
ORNEODID.
Genus TRISCADECIA, nov.
Proboscis fully developed ; palpi with the 2nd joint porrect about
twice the length of head and broadly fringed with hair below, the
3rd oblique, moderate ; antennze of male ciliated ; tibia with pro-
Jecting tufts of hair at the spurs. Fore-wing divided into six plumes
to rather more than one-third length; the costa with six tufts of
scales ; the scaling rough ; vein 3 from well before angle of cell ;
Triscedecia dactyloptera, & 2.
4-5 from angle; 6 from below upper angle ; 7-8 stalked ; 9, 10, 11
from cell. Hind-wing divided into seven plumes to half length ;
veins 3-4 from angle of cell; 5 absent ; 6-7 from upper angle ;
8 free, from base.
This is the third known genus of the family, the others
being Orneodes, Latr., and Pelia, Wik.
Triscedecia dactyloptera, n. sp.
6. Head, thorax and abdomen clothed with whitish, pale brown
and black scales; antennee and legs mixed with whitish ; abdomen
with diffused blackish bands. Fore-wing clothed with greyish, pale
brown and black scales; the costa with whitish spots between
the black tufts of scales ; the cell with small black lunules followed
by whitish spots at middle and extremity ; traces of a diffused
oblique whitish band from lower angle of cell to inner margin ; an
indistinct slightly waved whitish postmedial line ; the plumes with
248 Sir G. F. Hampson on three
white spots at base and subterminal and terminal series of white
lunules. Hind-wing with the basal half white with diffused irregular
black antemedial and two medial lines on it, the 1st of the medial
lines arising from a discoidal bar ; a slightly waved whitish post-
medial line; the plumes with white spots at base and subterminal
and terminal series of white lunules.
Hab. CrYLon, Maskeliya (J. Pole), Ohiya (Gossage).
Exp. 26 mm. Type in B. M.
PTEROPHORID&.
Genus TITANOPTILUS, nov.
Proboscis fully developed ; palpi porrect, slender, projecting about
the length of head; antennee with the shaft roughly scaled, the
basal joint with large tuft; fore femora with tuft of scales near
Titanoptilus melanodonta, & 3.
extremity, the tibia with large tuft at extremity ; mid tibia with
tufts at middle and extremity ; hind tibize with three tufts ; abdomen
very long with lateral tufts of scales at extremity of segments.
Fore-wing bifid to near middle, the lower plume falcate at extremity ;
a very large scale-lobe on inner margin below end of cell; vein 3
absent ; 5 absent; 6 from. below upper angle of cell ; 7-8 stalked ;
9-10 absent; 11 from cell. Hind-wing trifid, the upper excision to
two-thirds, the lower to near base; the inner margin with large
remarkable new genera of Microlepidoptera. 249
scale-tooth at one-third and very large scale-lobe at two-thirds ;
veins 3, 5 and 6 absent.
Nearest to Z'richoptilus, Wlsm.
Titanoptilus melanodonta, n. sp.
¢. Head, thorax and abdomen clothed with grey, brown and
black scales. Fore-wing greyish suffused with dark brown and
irrorated with rough black scales ; a whitish mark at base of excision,
the fringes of scales below upper plume and above lower black and
whitish ; the lobe on inner margin formed of large black scales, the
lower plume fringed below with black and whitish scales and with a
scale-tooth before its falcate extremity. Hind-wing greyish suffused
with dark brown ; a few black scales in the fringes below and at
extremity of the two upper plumes; the inner margin with the
long spatulate scales of the scale-tooth and lobe pale at base, black
at extremities ; a tuft of black scales at extremity of plume with
the fringe on inner margin before it white.
Hab. British E. Arrica, N’dimu, Uganda Ry., mile
469. (Betton), f. Exp. 42 mm. Type in B. M.
‘all i rh
< P
(i: ADabiew)
XIV. Descriptions of some new species of Diurnal Lepido-
ptera, collected by Mr. Harold Cookson, in Northern
Rhodesia, in 1903 and 1904. By HERBERT
Drucsz, F.L.S. Lycznide and Hesperiide, by
Hamitton H. Drucez, F.Z.S.
[Read March Ist, 1905, ]
PLATE XIII.
THE collection contains a large number of species, the most
interesting being Planema pogget, Precis touhilimasa, Crenis
rosa § and 9, Pseudacrxa poggei, Huryphene plistonax,
Buryphene letitia fand 2, Crenidomema concordia & and 9,
Charazxes macclouni g and §, Teracolus regina, Papilio
ridleyanus, Papilio toboranus.
We have described 11 as new species, leaving several
others, of which only one specimen has been sent, for future
determination.
Family SATY RIDA.
Mycalesis cooksoni, sp. nov. (Plate XITI, fig. 1.)
d. Upper-side. Primaries and secondaries dark brown, the
fringes of both wings lighter brown. Primaries with a small in-
distinct apical ocellus, a large black ocellus near the anal angle with
a small white dot in the middle, a short oblique cream-coloured band
erosses the wing from the costal margin to the upper-side of the
black ocellus.
Under-side. Primaries greyish-brown crossed beyond the middle
by a narrow yellow curved line, the cream-coloured band as above but
smaller, the ocellus more distinct, two fine yellow lines close to the
base. Secondaries greyish-brown mottled with darker brown, a
blackish-brown line crosses the middle of the wing from near the
apex to the inner margin above the anal angle, a submarginal row
of minute black dots extending from near the apex to the anal angle
those near the anal angle the largest.
Expanse 2 inches.
Hab. NORTH-WEST RHODESIA.
This species is allied to A/ycalesis sciathis, Hew., from
Old Calabar, and MM. selous?, Buti.
TRANS. ENT. SOC. LOND. 1905,—PART 11. (JULY) 17
2
to
On
bo
Mr. H. Druce’s Descriptions of
Mycalesis haroldi, sp. nov. (Plate XIII, fig. 2.)
d- Head, antenne, palpi, thorax and abdomen brownish-black,
under-side of the abdomen and the legs pale brown. Upper-side :
Primaries cream-colour broadly shaded with brown at the base, the
costal margin apex and outer margin dark brown, a small indistinct
ocellus close to the apex, a large ocellus nearest the anal angle, a
faint submarginal line extends from the apex to the anal angle.
Secondaries cream-colour, the base and inner margin clouded with
brown, the apex and outer margin dark brown with a faint sub-
marginal line extending from the apex to the anal angle.
Under-side ; both wings pale brown. Primaries crossed beyond the
middle of a cream-coloured band which widens out to the anal
angle, two large ocelli close to the outer margin, the first near the
apex, the second above the anal angle ; a marginal and submarginal
pale greyish band extends from the apex to the anal angle. Second-
aries crossed about the middle bya rather wide cream-coloured band,
a submarginal row of six ocelli extending from the apex to the anal
angle, the first large, second and third very small, four and five large,
six small, The marginal lines the same as on the primaries.
Expanse 2 inches.
Hab. NORTH-WESTERN RHODESIA.
LYCANIDA.
Amongst the Lycwnide are many interesting species,
showing an admixture of Western and South Tropical
forms. The collection contains Alxna oberthuri, Auriv., a
species with uniform black hind-wings on the upper-side,
a good series of Pentila amenaida, Hew., showing consider-
able variation as regards the richness of colour and the
number of black spots : Mimacrea marshalli, Trimen, in
fine condition ; a largé series of the little-known Terias-
looking Liptena homeyeri, Dewitz.* A 9 specimen agree-
ing exactly with Dr. Trimen’s figure of Epamera trimeni,
Walleng., which if this is the true Zrimeni makes one
doubt whether the insects from the Transvaal referred by
Mr. G. A. K. Marshall to this species are correctly deter-
mined or whether Dr. Trimen’s figure really refers to
Triment or some other unnamed form. Zeritis sorhagent,
Dewitz, and Z. neriene, Boisd., agreeing exactly with
Boisduval’s figure, and in my opinion quite distinct from
Dr. Butler’s Z. amine. Aphnwus erikssoni, Trimen.
* Homeyeri, Dewitz, D. E. Z., 30, p. 429, t. 2, f. 5, 5a, 5b, 5e.
some new species of Diurnal Lepidoptera. 253
Spalgis lemolea, H. H. Druce, known from the Gambia.
Lycexenesthes adherbal, Mabille, Z. lasti, S. and K., LZ. Modes,
Hew., LZ. crawshayi, Butl, ZL. defina, Butl, and LL.
monteironis, Kirby, agreeing exactly with the type in the
Hewitson collection.*
Uranothauma autinorii, Oberth, U. falkensteinii,
Dewitz, and a large series of U. poggei, Dewitz, showing
scarcely any variation. Custalius hintza, Trimen, C. calice,
Hopff., and C. melena, Trimen. Catochrysops glauca,
Trimen, a good series, C. peculiaris, Rogenh., and the giant
C. stormsi, Robbe.
I recognize in all about sixty species, including the new
forms hereafter described.
Liptena eukrines, sp. nov. (Plate XIII, fig. 7.)
¢. Allied to L. prestans, Grose Smith,t from which it differs on
the upper-side by the black costal margin on the primaries being
broken beyond the end of the cell, and projecting downwards, thus
freeing the black apex ; the outer margin has only a linear black
edge and the secondaries are unmarked excepting at the base of the
cilia, where there appears to be a very narrow black line. Under-
side : primaries differ from those of L. prestans, as on upper-side ;
secondaries, ground colour pale grey dusted with brown and crossed
by numerous broken bands composed of irregular brown spots.
Head black ; eyes ringed with white ; legs yellowish ; abdomen
yellowish above, grey beneath ; antennw white-ringed, with black
clubs.
Expanse 1,', inch.
Hab. NORTH-EAST RHODESIA.
Three specimens showing no variation. This species
agrees in venation with Z. libyssa, Hew. {; the upper
radial on fore-wing originating at end of cell as in that
species. It has close superficial resemblance to Teriomima.
* Since the above was written, Mr. Bethune-Baker has shown me
M. Mabille’s type of L. adherbal, and pointed out that it is identical
with L. monteirowis, Kirby, and has priority. With this conclusion
I quite agree. The species I have in this paper referred to as L.
adherbal should be known as L. lamuwlata, Trimen.
+ Liptena prestans, Grose Smith, Rhop. Exot. African Lycenide,
pl. xxix, f. 15, 16, p. 141 (1901).
£ Liptena libyssa, Hew., Exot. Butl., Pentila and Liptena, t. 1, f.
5, 6 (1866).
254 Mr. H. Druce’s Descriptions of
Iridopsis cuprepes, sp. nov. (Plate XITT, fig. 8.)
gd. Closely allied to I. exquisita, Grose Smith.* Upper-side :
black margins more distinctly defined and rather narrower. Under-
side: primaries with four pale spots beyond the cell, the upper one
small, the two median quadrate, large and distinct, the lower spot
rounded and less distinct, below these there is an indication of a fifth
spot. There are three pale patches at the outer angle, large and
distinet. Secondaries ; the median band appears wider and more
broken, and the marginal coloured band containing the metallic
lunules is wider.
Expanse 132 inch.
Hab. NoRTH-weEst RHODESIA.
Mr. Cookson only sent one specimen. Although this
insect is undoubtedly very close to J. exquisita; I believe
that, judging from analogy in the genus Phytala, it will
prove to be distinct.
The species of this genus are very little known and
seldom seen in collections. We have in our possession a
specimen from Sierra Leone, which is exactly like Mr.
Grose Smith’s figure of J. ansorgei,t and I have no doubt
that it is the 2 of LZ. ineredibilis, Staud., the ¢ of which
is figured in Iris 4, t. 1, f. 7, 1891.
Spindasis kallimon, sp. nov. (Plate XIII, fig. 9.)
Q. Upper-side. Primaries blackish-brown with distinct orange
bands formed of quadrate spots. The Ist in, and at the end of the
cell, consisting of a quadrate spot and a triangular spot below it ; a
large quadrate spot beyond, joining at the median nervule to an
irregular band consisting of six confluent spots which reach from the
apex to near the outer angle. The inner margin from the base for
about two-thirds its length and extending to the lower median
nervule pale silvery blue. Secondaries pale grevish-brown with the
under-side bands showing through as darker grey patches ; a deep
black outer marginal line. Anal angle largely deep orange, contain-
ing a small black spot and two small patches. of metallic scales,
Cihe of both wings distinctly and evenly deep orange, except at the
anal angle where it is deep black. The tail on the lower median
* I. exquisita, Grose Smith, Nov. Zool., 5, p. 353 (1898), Rhop.
Exot. African Lycenidx, pl. xxviii, f. 12, 13 (1901).
+ I. ansorgei, Grose Smith, id. p. 354, Rhop. Exot. id. f. 10, 11.
some new species of Diurnal Lepidoptera. 255
nervule is black, tipped with white, that on the submedian nervures
being orange tipped with black. Under-side: Both wings pale yellow
with dark orange red-bordered bands, which incline to rounded
spots, thickly lined interiorly with metallic scales. At the base of
the fore-wing is a bright orange costal dash. There is a marginal
narrow red line and two submarginal more prominent red lines to
both wings.
A rather large marginal black spot at the anal angle below the
submedian nervule and a smaller one crowned with metallic scales
in the submedian interspace. Head dark orange without any
black markings. Thorax and abdomen blackish above, pale yellow
below; abdomen ringed with whitish.
Legs pale yellow; palpi pale yellow below, darker above with
black tips.
Expanse 1,3; inch.
Hab. NortH-west RuHopesIA, January 1903.
A very beautiful and distinct species, not nearly allied
to any that I can find described, but perhaps nearest to S.
homeyeri,* Dewitz, Mr. Cookson sent four specimens,
all females.+
Erikssonia cooksoni, sp. nov. (Plate XIII, fig. 4.)
¢. Allied to Hrikssonia acrxina, Trimen.{ Upper-side uniform
reddish-brown. Primaries with the costa (except at the base, which
is reddish-brown) increasingly broadly black for about three-fourths
its length when it becomes suddenly narrow and joins the broad evenly
black outer margin. An elongate black spotat the end of the cell. Cilia
deep black. Secondaries with a gradually increasing outer marginal
-black band, commencing beyond the middle of the costa and be-
coming narrow and ,broken about the region of the median nervure,
* Spindasis homeyeri, Dewitz, D. E. Z., 30, p. 429, t. 2, f. 5, 5a,
5b, 5e.
+ Since the above description was written, Professor Poulton has
sent me for examination a ¢ captured at Rabai, 14 miles N.W. of
Mombasa on June 18th, 1908, by Rev. K. SAubyn Rogers, and
presented by him to the Oxford Museum. It differs from the ? on
the upper-side by the orange bands on the primaries being narrower
and rather less conspicuous and by the hind-wing being suffused
with rich purple from the subcostal to the submedian nervure ; in
this respect somewhat resembling spindasis natalensis, Westw., which
has the inner margin of the fore-wing also broadly sutfused. On the
under-side it is identical with the 9.
t Hrikssonia acreina, Trimen, P. Z.8., 1891, p. 92, Pl. IX, f. 18,
19, 20.
256 Mr. H. Druce’s Descriptions of
and continuing narrowly to the anal angle where there is a black
line placed obliquely just above it. The cilia is black, whitish at
the base between the nervules. Under-side: Primaries rich orange,
paler along the inner margin; black spots arranged as in H. acreina
with an additional black spot below the cell placed below the central
cell spot. The outer margin is broadly paler, bordered inwardly
with a black line, heavily covered with metallic silver scales and
divided by the black nervules. Cilia black. Secondaries: Ground
colour pale buff with the black spots large and distinct, the basal
area suffused with bright carmine, and a bright carmine band placed
between the ultra-median row of black spots and the sub-marginal
black line. Outer margin and metallic scaling as in primaries.
Legs reddish-orange ; palpi orange below, black above. Antenne
black; brownish at the extreme tip. Head black, yellow between
the eyes. Abdomen deep orange-red above with a row of black
spots ; sides deep black with white spots ; under-surface clothed
with more or less orange hairs.
Expanse 1,'5 inches.
Hab. NortH-wEsT RHODESIA.
Although doubtless allied to #. acrxina this insect
appears to have many points of distinction, notably the
broad black outer marginal border to the fore-wing above;
the deep black cilia; and the carmine on the under-
surface of the hind-wings. It is also a larger insect.
COOKSONIA, gen, nov.
Allied to Hrikssonia,* Trimen, from which it differs in the fore-
wing by the 4th subcostal nervule being emitted nearer to the base
and by the subcostal nervure reaching the margin further below
the apex; and in the hind-wing the stalk of the submedian
nervure is much shorter. The antenne are longer and when viewed
from above appear to be but slightly thickened at the extremities,
but when examined from the side they present a broadly spatulate
club, in this respect entirely differing from Hrikssonia.t The palpi
which are not clothed with thick scales to anything like the
same extent have the terminal joint less than half as long ; almost
minute.
* Hrikssonia, Trimen, P. Z.8., 1891, p. 91. ‘
+ This may possibly be due to pressure in packing the specimen,
which, however, shows no signs of flattening as regards the abdomen
or thorax, and is in very perfect condition.
some new species of Diurnal Lepidoptera. 250
Type. Cooksonia triment, sp. nov.
I have compared this genus to Dr. R. Trimen’s Zriks-
sonia, but it presents important differences which may
prove more remarkable when the male sex is examined.
I dedicate it to Mr. Cookson, its discoverer.
Cooksonia triment, sp. nov. (Plate XIII, fig. 3.)
2. Upper-side. Orange-ochreous, suffused with reddish-ochreous,
deepening in colour towards the bases, brightest in the fore-wing.
Primaries : Costal margin narrow and evenly black, outer margin
rather more broadly black; apical third pure white, inwardly
bordered with a rather broad black band, comparatively straight and
even on its inner edge, and much suffused and irregular on its outer
edge, reaching from the costa to the outer margin, where it is
broadest ; the nervules crossing this white apex being distinctly
black and being more decidedly conspicuous as they recede from the
apex. A large black spot at the end of the cell and a smaller black
spot beyond the middle of the cell placed near to but apart from its
upper wall.
Secondaries ; With a black border, commencing very narrowly be-
yond the middle of the costal margin, gradually and evenly widening
towards the apex whence it becomes broken and though thickening
at the extremities of the median nervules is very narrow along the
inner margin and disappears before the base is reached. <A distinct
black spot at the end of the cell placed well above the median
nervure.
Under-side. Primaries brownish-ochreous, reddish in the cell ; a
distinct black spot at the end of the cell. The apical third consists
of a series of five narrowly black-bordered greyish conical spots placed
between the nervures, each one containing at its marginal base an
indistinct ochreous patch. The outer margin has a linear black
border.
Secondaries pearly-grey, with the inter-nervular spaces more or
less suffused with reddish-brown scales which are most conspicuous
just beyond the middle. The basal half contains about twelve large
black irregular white-ringed spots, several of which, close to the
base are confluent, and a median row of irregular black dots. The
outer margin is bordered with a linear black band; beyond which
are two rows of angular black markings placed between the
nervules, the outer row enclosing ochreous patches, the inner pale
grey.
Head black, with two white spots behind the bases of the antenne
and two white streaks between the eyes,
258 Mr. H. Druce’s Deseriptions of
The upper part and the sides of the abdomen are oclireots, but on
the under-side of each segment there is a large distinct, white, black-
margined spot. There are also white spots on the under-side of the
thorax. The legs are black with yellow tufts at the base,
Antenne black ; the shafts minutely spotted with white on the
under-surface.
Expanse 2} inch.
Hab. NortH-EAST RHopESIA. October 1903.
On the under-side this insect has a remarkable Acrxine
appearance, but although the upper-side has a general
likeness to that group I know of no species of the genus
Acrea which has the large white apex to the fore-wing.
Lycenesthes anadema, sp. nov. (Plate XIII, fig. 6.)
¢. Allied to L. lasti,* Smith and Kirby, but smaller. Upper-side:
dark purple with indistinct black marginal spots between the lower
median nervules on hind-wing. Under-side: ground colour rather
deeper in tone with the whitish chain-like bands less inclined to
break up into spots and with their enclosed spaces scarcely darker
than the surrounding ground colour. The red spots on the hind-
wing, with the exception of the one below the costal nervure near
the base, which is entirely absent, are dark, clearly defined, and con-
spicuous. The black, yellow-crowned marginal spot placed between
the lower median nervules is smaller, as is the orange anal-angular
streak. Thorax, abdomen, head and legs as in L. lasti. Cilia on
upper-side pale brown, whitish towards tip.
Expanse 17 inch.
Hab. NoRTH-WEST RHODESIA.
Can be distinguished by the smaller size and by the
absence of the red basal spot on hind-wings below.
It is also allied to L. hobleyi,+ Neave, but that species
had a red basal spot and also a red cellular spot in hind-
wings.
Catochrysops pampolis, sp. nov. (Plate XIII, fig. 11.)
¢. Upper-side: pale opalescent bluish brown witha linear brown
margin and brown nervules to both wings; a slightly darker brown
patch at the end of the cell of the fore-wing, containing a still darker
streak, A medium-sized round black spot close to the margin of the
* L. lasti, Smith and Kirby, Rhop. Exot., 27. Afr. Lye., p. 109,
t. 24, f. 1, 2, 1894.
{ L. hobleyi, Neave, Nov. Zool., v. xi, p. 339, 1904.
some new species of Diurnal Lepidoptera. 259
hind-wing, placed between the two lower median nervules. Cilia
white, indistinctly darker at the extremities of the nervules. Under-
side ; ground colour of both wings pure white with broad blotches.
Primaries ; a large quadrate blotch at the end of the cell ; costal margin
pale brown; an ultra-median brown band composed of confluent
quadrate spots, semi-circular in form, reaching from the subcostal
nervure, where it is widest, to lower median interspace where it is
narrowed ; beyond this a rather narrow brown band of confluent
inter-nervular patches, followed by a somewhat paler and narrower
brown band reaching only to the lower median nervules, and beyond
this a clearly defined, narrow, marginal line. Cilia white; dusky at
the ends of the nervules. Secondaries: costal margin pale brown ;
two large black spots close to the margin, one about the middle, one
half-way between this and the base ; a triangular black spot at the
base of the cell, and two more along the anal margin ; a large irregular
dark brown blotch, with several smaller blotches more or less
attached to it, occupying the central area, and a double submarginal
brown band more inclined to become fused towards the anal angle.
Close to the margin between the lower median nervules is a large
oval orange spot centered by a black spot which is crowned exteriorly
by rich metallic-blue scales. In the submedian interspace, close to
the submedian nervure is an orange streak bordered on either side
by a small black spot; cilia white, brownish at the extremities of
the nervules.
Head, thorax and abdomen brownish-grey above, white beneath,
Legs white ; palpi white, with black tips. Antenne black, white
ringed,
A short linear brown tail on the lower median nervule.
Q. Upper-side paler and brighter blue than ¢, fore-wing with
apex and outer margin broadly pale brown ; hind-wing with an
ultramedian, semicircular, irregular whitish band. Under-side: the
white areas more extensive than in ¢, causing the brown blotches to
appear more distinct; the costal margin and the basal area are more
broadly pale brown and the black spots appear white ringed.
Expanse ¢ 134 inch, ? 1,3, inch.
Hab. Norta-west RaopeEsia, November 1903.
Not very closely allied to any species that I can find
described.
Catochrysops skotios, sp. nov. (Plate XIII, fig. 12.)
d. Upper-side uniform dark olivaceus-brown, primaries with a
darker streak at the end of the cell. Cilia slightly paler brown.
Underside : ground colour olivaceus-brown, but slightly paler than
260 Mr. H. Druce’s Descriptions of
upper-surface, with sordid white-ringed spots and markings. Prim-
aries : a large spot at the end of the cell ; an ultramedian band of six
chain-like spots running almost in a straight line from the subcostal
nervure to the submedian nervure, the two lower links being placed
slightly nearer to the base, then two submarginal rows of pale
sagittate lunules and a pale marginal line. Secondaries: two
distinct black spots on the costal margin, another about the middle
of the cell, and two more on the anal margin, one about the middle,
the other near the base. A large sordid white-ringed brown spot
near the end of the cell, then an irregular, much broken band con-
sisting of sordid white-ringed spots, commencing below the black spot
near the middle of the costal margin and reaching to the submedian
nervure ; beyond this band are two rows of sagittate pale markings
and a pale marginal line as in the primaries. A deep black spot,
bordered outwardly with rich metallic scales and broadly crowned
and partially surrounded by rich orange, is placed between the lower
median nervules close to the base. Cilia of both wings brown.
Abdomen brown above, paler below; legs brown ; head brown,
eyes white-ringed. Palpi white, with black tips.
?. Differs only from the ¢ on the upper-side by having a large,
black, orange-crowned spot on the hind-wing near the margin, placed
between the lower median nervules. On the under-side of the fore-
wing there is a minute dot above the subcostal nervure at the end
of the chain-like band. The orange-crowned spot on hind-wing is
more conspicuous and there are traces of orange scales near the anal
angle.
Expanse ¢ 14 inch, 9 1% inch.
Hab. NorRTH-westT RHopESIA, October and November
1903.
This species, which at first sight appears of a general
type, is not very closely allied to any with which I am
acquainted,
HESPERIID.
Cyclopides cooksoni, sp. nov. (Plate XIII, fig. 10.)
Allied to Cyclopides formosus, Butl.* Upper-side blackish-brown
with bright yellow spots. Primaries: the basal third and the inner
margin thickly dusted with yellow scales. A minute spot about the
middle of the costa, and a rather larger one in the cell, below it. An
ultra-median band, consisting of various sized irregular spots, reaching
from the costa to the inner margin, the spot next to the costal spot
* Cyclopides formosus, Butl., P..Z..S., 1893,.p. 670, pl. Ix, £8:
some new species of Diwrnal Lepidoptera. 261
being placed right beyond the band, nearer to the outer margin.
Secondaries : a semicircular median band consisting of three large
irregular spots, beyond which, is a submarginal row of six (sometimes
seven) smaller spots. Cilia of fore-wing black ; of hind-wing black,
but bright yellow at apex and anal angle. Under-side. Primaries ;
costal margin yellow, divided just beyond the cell; a broad yellow
streak and a spot in the cell. The band of irregular spots as on
upper-surface is present with the addition of a large spot in the
interspace below the costal spot. Yellow marginal internervular
radiations and spots as in C. formosus, but more pronounced.
Secondaries much as in C. formosus, but the dark yellow spots deeper
in tone and larger.
Antenne, legs, palpi and head much as in OC. formosus.
Expanse 1%; inch.
Hab. NortH-wrEst RHopDESIA, December 19038.
One specimen which I take to bea ? has a submarginal
series of about eight minute yellow dashes on the upper-
side of the fore-wing, and on several other specimens I
can detect traces of some of these markings.
The hind tibiz have the two pairs of spurs, the upper
pair being smallest.
Mr. Cookson procured a good series of this species,
which appears to be quite distinct from any hitherto
described.
Fig. 1.
2.
Huplanation of Plate.
EXPLANATION OF PLATE XIII.
Mycalesis cooksoni, sp. n., p. 251.
“4 haroldi, sp. n., p. 252.
Cooksonia triment, sp. n., p. 257.
LHrikssonia cooksoni, gp. n., p. 255.
Liptena homeyeri, Dewitz, p. 252.
Lycenesthes anadema, sp. n., p. 258,
Liptena eukrines, sp. ty p. 253.
Tridopsis ewprepes, sp. n., p. 254,
Spindasis kallimon, sp. n., p. 254.
Cyclopides cooksoni, sp. 0., p. 260.
Catochrysops pampolis, sp. N., p. 258.
skotios, Sp. N., p. 259.
(9fQGBe a
XV. Pseudacrea poggei and Limnas chrysippus; the
numerical proportion of mimic to model. By
Horace A. Byatt, B.A., F.E.S. With a note by
Professor E. B. Pouuton, D.Sc., M.A., F.R.S., ete.
[Read April 5th, 1905. ]
PLATE XIV:
THESE butterflies were found among a collection of some
1200 specimens given to me by Pére Guillemé of the
White Fathers’ Mission to Central Africa, under whose
direction they were collected at his station at Kayambi,
in Awemba country, N.E. Rhodesia, near the sources of
the Congo, locally called the Chambezi, between October
1898 and January 1899.
His system was to send out a number of native school-
boys—his “ gamins,” as he called them—each armed with
a net and a book, and orders to capture anything and
everything that came in their way, placing their captures
between the leaves of the book for safe carrying home.
He particularly mentions that he told his boys to take
“des spécimens aussi variés que possible ;” and that they
would do this literally I know from my own experience of
natives, for I have fond them generally unable to dis-
criminate between species, and when sent out by me on
similar occasions they have returned with large numbers
of the msect most in evidence at the moment, and a pro-
portionally smaller number of others. It is, therefore,
allowable to suppose that the whole lot which came into
my possession gives a very fair idea indeed of the numerical
strength of the several species found in the locality.
On opening the papers and examining the specimens—
which have suffered a good deal from the damp and
neglect of seven years—it was found that roughly one-
third of the whole collection consisted of Limnas chrysippus,
L., and its mimics ; and among these latter were seventeen
specimens of Pseudacrea pogger, Dewitz,—many of them
in a fair state of preservation, though, with the rest, they
show signs of being unduly pressed between the pages of
the book, and are somewhat dulled in colour by damp.
TRANS. ENT. SOC. LOND. 1905.—PART II. (JULY)
264 Mr. Horace A. Byatt on
It is due to the suggestion of Professor Poulton that
this would be an excellent opportunity to ascertain what
numerical relation the Pseudacrea bears to its model
chrysippus that this paper has been hastily prepared in
the Hope Department at Oxford before my return to
Central Africa.
The country in which Kayambi is situated does not
differ greatly from the rest of the plateau to the W.
of Lake Nyassa and N.E. Rhodesia. Large stretches
of undulating plains, covered with thin scrubby bush
or dense tall grass, are intersected at intervals of 5-20 miles
by streams and rivers. In the latter months of the year,
when these insects were mostly collected, these plains
are bare, dry, and dusty, grass and bush bemg burnt up
by the annual bush-fires: and only along the streams is any
verdure found. Pére Guillemé describes the soil as fertile
along the course of the rivers, but elsewhere the district
is generally poor and sandy, and for this reason sparsely
inhabited, and he remarks on bush-fires being the cause
of the general scarcity of insect-life except along the
water-courses, where the vegetation is untouched by fires,
and where forest-giants, trailing creepers, and tree-ferns
flourish.
The altitude of Kayambi is about 3950 ft. above sea-
level; and its position roughly 9° 20'S. and 31° 50’ E,,
some two days’ march from Fife, and three from Abercorn,
on the Nyassa-Tanganyika plateau.
The respective numbers and species in the collection
were worked out in the Hope Department and are stated
in tabular form below :—
SPECIES. ' 3 Q |ToTAL
Limnas chrysippus ; «1 "288 TO 367
do. do. var. dorippus 8 a 12
Pseudacrexa pogger ; . — — 17
Hypolimnas misippus . = 36 7 ; p
“ BREED on V3 45
a » §@ Var. maria 2
|
Total number in group 441
From this table it will be seen that Pseudaerva pogger
is by no means so rare as has been hitherto supposed ;
its proportion to LZ. chrysippus is no less than 4°72 per
cent. The dorippus, K1., or klugit, Butl., form of chrysippus
Pseudacrxa pogge and Limnas chrysippus. 265
is found, but in this whole series of seventeen specimens
of pogget no individual shows any resemblance to dorippus :
it is purely a mimic of chrysippus and shows no approach
to dimorphism. This is explainable on the ground that it
is found only where chrysippus is the largely predominant
form, and, so far as is known, it does not occur in, or has
not yet reached, the parts where dorippus is relatively
abundant—that is, the desert strip along the E. Coast,
extending in the E, African Protectorate inland at least to
the shores of Victoria Nyanza. In this respect it com-
pares in an interesting manner with misippus ¢, of which
the inaria fori, mimicking dorippus, is found all over
Africa (Trans. Ent. Soc. Lond., 1902, pp. 473-484): and
also with Acrwa encedon, L., var. daira, G. and S., which
is only frequent in localities where its type dorippus 1S
predominant (l.c. pp. 473-484).
This occurrence in considerable numbers of what has
hitherto been regarded as the rarest species of Psewudacrea
supports the hypothesis that the mimics of this group are
Miillerian rather than Batesian. This has already been
argued for HH. misippus (cf. Rep. Amer. Assoc. Adv. Sci.—
Detroit, 1897; and Trans. Ent. Soc. Lond., 1902, pp. 480
and 483, with references) as well as for the genus Psewda-
crea and for A. encedon (pp. 480, etc.). The fact that
the latter species, belonging to a protected and much-
mimicked subfamily, is far more coincident geographically
with the corresponding forms of its model A. misippus
2 is obviously a powerful argument in favour of the
Miillerian interpretation.
Note sy Proressor E. B. Pouuton, F.R.S.
Ir is deeply interesting to compare the details of the
mimetic resemblance borne by Pseudacrea poggei to
Limnas chrysippus with those of the other great Nympha-
line mimic—Hypolimnas misippus 9. Almost all the
points in the following statement can be verified by
means of the half-tone reproductions of the three species
on the accompanying Plate XIV. It is to be observed,
however, that Fig. 1 represents a 2 chrysippus with five
wings, a second smaller left hind-wing concealing the
central portion of the normal hind-wing of the same side.
266 Mr. Horace A. Byatt on
As this rare monstrosity was found among the large
number of specimens tabulated by Mr, H. A. Byatt it
was thought well to select it for representation, inasmuch
as the teratological interest is simply an additional
advantage which in no way interferes with the bionomic
interest of the plate.
The subapical white bar on the fore-wing of chrysippus
is prolonged downwards and outwards with a slight in-
ward trend by means of a few small marginal spots and a
local intensification of the white elements in the fringe.
This character is very persistent, and is traceable in the
dorippus form when the band itself has, except for its
costal end, disappeared (compare Fig. 4 with 1). A similar
effect is produced in the 2 misippus (Fig. 2) by the
position of the last or fifth spot of the band, by a local
strengthening of the two rows of whitish hind-marginal
lunules, aud by the white elements of the frmge. The
three narrow interrupted white lines which are thus
formed parallel with the hind-margin, persist in the
inaria form when the band itself is only faintly trace-
able (compare Fig. 5 with 2). In both chrysippus and
misippus it is obvious, especially in the latter, that this
prominent subapical marking is in large part prolonged by
the local strengthening or the local persistence of elements
which are not part of the bar itself, but belong to the
category of marginal markings. In this respect Pseuda-
crea pogger (Fig. 3) stands in considerable contrast with
the other two members of the group; for its bar is
prolonged—and much more fully prolonged than in the
other species—by elements which have the appearance of
continuity with the bar itself. If these elements are
marginal markings as in chrysippus and misippus they
have been far more subordinated to the subapical bar
than in these species. The local strengthening of white
elements in the fringe is also somewhat less marked, and
plays a less important part in poyger than in the others.
As regards the few minute spots at the extreme apex
of the fore-wing of chrysippus, mimetic resemblance is
more honoured in the breach by poggei than in the too
emphasized observance by misippus ?—to say nothing of
the very different position of the marking in model and
mimic,
In spite of all these differences in detail, the two
mimics are by no means unlike; and in general effect
Pseudacrea pogge: and Limnas chrysippus, 267
each of them resembles the other more closely than it
resembles the model.
The internal contour of the black hind-marginal border
of both wings is prolonged inwards along the veins, produc-
ing a festooned appearance in pogget (Fig. 3) and, to a far
smaller degree, in the model (Fig. 1). In the 2 misippus
(Fig. 2) this feature is almost wanting. Apart from the
contour, the narrow black border of the hind-wings
of pogget more closely resembles the model than the
broader more interrupted and less sharply outlined border
of the 2 msippus. At the same time, the two mimics
resemble each other in this character more fully than
either of them resembles the model, which is widely
separated by the row of distinct white spots, which
however are very variable, and not infrequently barely
traceable. On the under-side of both wings the border
of misippus (Fig. 2a) reproduces the black and white
effect of the model (Fig. 1a), far better than poggei (Fig.
3a), in which the white marginal elements are confined
to the fringe. On the other hand, in the lighter tint of
the veins of the hind-wing under-side and in the colour
and texture of the ground-colour, pogget is by far the
better mimic of the two, while msippus is equally superior
in the tint of the apical area of the fore-wing under-side
beyond the bar. As regards the black discal spots of
the hind-wing under-side poggei is the closer mimic. The
development and shifting outwards of the peripheral spots
is an evident special modification, in the direction of the
model, of a characteristic feature of the genus Pseudacrea.
The two small spots nearest to the centre of the wing
(see Fig. 3a) were only seen in a single specimen out of
the seventeen, They indicate the existence of material
which may be developed into a still closer likeness to the
Danaine model.—E. B. P.
TRANS. ENT, SOC, LOND. 1905.—ParT Il, (JULY) 18
268
Explanation of Plate.
EXPLANATION OF PLATE XLV.
All the figures are two-thirds of the natural size.
Limnas chrysippus with its two chief mimics, from the sources of
the Congo.
1 eS ile
Limnas chrysippus, 9, wpper-side. The specimen possesses
a second smaller hind-wing on the left side.
1. Limnas chrysippus, 3 , under-side.
Hypolimnas misippus, 9 , upper-side.
* #4 © , under-side.
Pseudacrxa poggei, upper-side.
ss » under-side.
Limnas chrysippus, form dorippus (Klugii), g, wpper-side.
Hypolinnas misippus, 9 form inaria, upper-side.
JuLy 15TH, 1905,
( 269 )
XVI. A monograph of the genus Ogyris. By GrorGcE T.
BETHUNE-BAKER, F.L.S., F.Z.S.
[Read April 5th, 1905.]
PLATE XV;
THIS genus was established and described by Doubleday
and Westwood in their Gen. Diurn. Lep. II, p. 472, 1850,
where they figured the ? of 0. abrota and used this species
int heir description of the family. It is a well-marked
genus, and as a consequence later authors have had no
difficulty in their descriptions of new species, all having
been described under the one genus. Hewitson described
and figured his several species in his Exotic Butterflies, vol.
I, and his Spec. Cat. Lye. B. M.; whilst in his Ill. Diurn.
Lep. Lycenide, he again refers to the group and sinks his
orontas under his idmo. The Felders also use the genus
in the descriptions of their two species, whilst all later
authors have likewise used it.
The genus is confined to the Australian sub-region, all
the species but one being found on that continent
(Australia) : one species has been recorded from Kangaroo
Island as well, whilst one is confined to New Guinea. All
the species are beautiful in,colour, whilst several rival the
Morphos in the brilliancy of their blues.
I am indebted to Mr. G. A. Waterhouse of Sydney for
very many particulars relative to the life histories of the
species and also for the loan of a large number of specimens.
Mr. F. P. Dodd has also furnished me with many interest-
ing facts about O. zosine, Hew., and hewitsoni, Waterh.,
whilst Mr. R, E. Turner and Mr. H. H. Druce and Mr.
Tepper have kindly lent me specimens of 0. gxnone,
Waterh., O. barnardi, Miskin, and O. halmaturia Tepper.
The ova are either nearly spherical, or somewhat com-
pressed at each end of the axis, whilst some are strongly
reticulated. The larve, so far as is yet known, are all
TRANS. ENT. SOC. LOND. 1905.—PART II. (ocT.) 19
270 Mr. G. T. Bethune-Baker on
Loranthus feeders, and all feed either at dusk or at night
all are attended by ants of various species, some apparently
by different species in different neighbourhoods, whilst Mr.
Dodd informs me that 0. zosine G genoveva) i is attended by
two species of Camponotus and also by the small black ant.
He adds also that both it, orwtes, Hew., and hewitsoni,
Waterhouse, emit sounds when in the pupa state and when
accompanied by ants; if however the ants are taken away
the pupze remain silent—whilst if a pupa be found
unattended by ants, it is an almost certain indication that
it is dead; the sound is described as a distinct ticking,
occasionally accompanied by a soft humming. The pupz,
of which I have several before me, are of the usual Lyczenid
shape, and are found under stones or under loose bark,
fastened at the anal extremity and supported by a girdle
of silk around the middle, and generally assimilating in
colour with their surroundings.
The species of the parasitic genus Loranthus, on which
the larvee feed, generally grow high up on the tallest trees,
and as a consequence the perfect insect flies high and is a
strong flier; this fact possibly accounts for the scarcity of
the group in collections generally. The species are
probably on the increase, as Mr. Waterhouse informs me
that he now breeds ianthis in a spot over which during his
younger days he collected for years, and of which he knew
every inch of the ground, and during this time he never
saw a single specimen of this genus; this is however what
might be expected, for it appears quite certain that the
larvee and pupz are protected by the ants, in which case
they would be immune from a certain class of enemies, and
we might therefore look for the various species to increase
in numbers.
The various species form themselves into two obvious
sections, the females of which have a pale spot in the
primaries, or are without that spot; and in like manner
they fall into sub-sections also; the whole being closely
verified by the form of the genitalia of the males, descrip-
tions of which I append in the form of a table, though in
a few species that have been lent me I am unable to do
this.
The tegumen of the whole of the Lycwnide is furnished
with a pair of hooks at the lower extremities of the lateral
lobes, these I have designated by the term “ Falces”
(falx, a reaping-hook).
The genus Ogyris. 271
Key.
A. Upper-side generally purple; 2 with pale spot on the
primaries.
a. Sexes nearly the same colour.
al, Termen broadly brown, strongly
arched. : E é . waterhouseri.
a2. Termen quite narrow, nearly
straight . : : : . amo.
a3. Smaller, lighter purple, termen
narrowly brown, pattern of
secondaries beneath nearly
obsolete . : : : . otanes.
a4. Like 3, but termen broadly
brown, broader at apex and
tornus . : F : . halmaturia
b. Sexes dissimilar in colour.
b 1. Dull brownish-purple, termen
broadly brown at apex tapering
to tornus, anal angle of second-
aries produced into a short
broad tail : ; : . genoveva.
b 2. Brighter purple, termen nearly
uniform in width . : . genoveva-duaringa.
b 3. Rich purple, larger in size with
very large females . : . geivoveva-magna,
b 4. Very dark velvety-purple (almost
brown) with long tail, slightly
spatulate, pattern below strongly
spotted, no metallic markings
in the cell of primaries. . meeki.
b 5. Smaller, very deep velvety-blue,
termen black very broad, pale
large subovate lemon spot in
primaries : . 4 . abrotu.
b 6. Brilliant metallic-blue with broad
black costa and termen; @
primaries bright orange, with
broad borders . : : . ianthis.
272 Mr. G. T. Bethune-Baker on
B. 2 without a pale spot on the primaries.
a. 2 with no red spot beneath in the
primaries.
b. Both sexes dark purple.
a1. Purple area very small, nearly the
same in both sexes . . olane.
a2. Purple area brighter extending
up to termenin g . : . barnardi.
ce. Both sexes brilliant metallic-blue.
al. Discal band of primaries beneath
not fractured . : A . oretes.
d. Brilliant metallic-blue; 9 with red
in the primaries beneath.
al. Discal band of primaries beneath
fractured . : ; ; . hewitsoni.
a2. Smaller, deeper blue, with pale
metallic-blue spots in cell
beneath . : ; : . hewitsoni-meridionalis.
a 3. Primaries with very broad black
apex and termen, and very dark
under-surface . ; 4 . amaryllis.
a4, Brilliant silvery-blue with costa
and apex broadly black, be-
neath very pale grey, with
pattern small and isolated . enone.
See Tables, pages 290, 291.
Oayris, Doubleday, Westwood and Hewitson.
Ogyris, D. W. and H., Gen. Diurn., Lep. II, p. 472
(1850-1852).
Primaries broad, with costa slightly arched ; apex subacute, some-
times produced ; termen sometimes slightly excavated below the
apex, or nearly straight, or slightly convex ; inner margin straight.
Neuration, vein 1 waved, vein 2 arising beyond the middle of the
cell, 3 from nearer the lower angle of cell than from 2, 4 from the
lower angle, 5 from above the middle of the discocellulars, 6 from
the upper angle, 7 absent, 8 from directly behind the upper angle
of the cell, 9 from about midway between the end of the cell and
the apex of the wing, or from nearer the apex, 10 and 11 with bases
The genus Ogyris. 273
equi-distant from each other and the upper angle of cell, 11 short,
12 short never as long as the cell. Secondaries broad, sometimes
with a short broad spatulate tail at the end of vein 2, sometimes
with a short tail at end of vein 1b, or with termen crenulate— ?
sometimes with a short tail at the end of vein 4, with two internal
veins—vein 2 rising from just below the middle of the cell, 3 from
near the lower angle, 4 from the lower angle, 5 from the middle of
the discocellulars, 6 from the upper angle of cell. Palpi curved,
with middle segment long, end segment very short and slightly
deflexed.
The genus was created for the reception of two species
0. abrota, D. W. and H., and O. idmo, Hew., the diagnosis
being evidently made from the former species. It is
nearly related both in structure and pattern of wings to
the genus Arhopala, Feld.; both genera are arboreal, rarely
coming down to the ground, and when disturbed from
their resting place, they will frequently return to it
after the lapse of a little time; this, however, is a habit
well known to occur in other genera also.
OayRIs WATERHOUSERI, spec. nov.
¢. Both wings very dull brownish-purple, almost greasy look-
ing. Primaries with a broad even brown costa and a broad termen
likewise of even width. Secondaries with a broad dark brown costa,
and a broad brown termen of uniform width. Fringes white broadly
intersected with brown at the veins. Under-side. Primaries brown,
with three increasing cell spots, the first two divided by an obscure
bluish-white line, beyond the third a short broad oblique pale grey
stripe, beyond which is the catenulated, very irregular, dark brown
stripe, slightly curved extending from the costa to very near the
termen at vein 2; the apical area up to this stripe is densely
irrorated with whitish-grey fine scales. Secondaries, brown, finely
irrorated with brownish-grey, the three basal spots are confluent,
in the median row of three spots the two upper ones touch, the
second occupying the central part of the cell is large, the third on
the inner margin is isolated, the third row is very irregular, first
spot on the costa subreniform, the second spot very large, touching
the exterior edge of spot 1, <-shaped on its exterior margin, a large
spot beyond the cell touching the inner edge of spot 2, the fourth
spot angled with the third and receding inwards, the fifth spot
again angled internally confluent with spot 4, the posterior row of
four spots touches the exterior edge of the second large spot in the
274 Mr. G. T. Bethune-Baker on
previous row, and each spot is angled externally but is rounded in-
ternally, the upper three touch each other, the fourth being isolated on
the inner margin, the subterminal row is obscure consisting of little
more than an indefinite stripe.
2. Both wings brightish-purple with a blackish spot at the end of
the cell, that on the primaries being much larger than the one on the
secondaries, Primaries with a broad brown costa and termen and a
small (comparatively) lemon-coloured spot beyond the black spot
terminating the cell ; secondaries with a very broad brown costa and
a broad brown somewhat irregular termen. Underside primaries
as in the male, but the divisions between the cell spots are pale
blue and the lemon spot shows through, Secondaries as in the
male but greyer, the pattern therefore shows more distinctly and
the spots in the median area are more confluent.
Expanse ¢ 53, 9 52-53 mm.
Hab. Victoria.
This species is distinct from 0. idmo, Hew., to which it
is very nearly allied, the shape of the wings in the % is
quite different, the costa of the primaries is more curved,
and the termen is distinctly produced outwards between
veins 2 and 5; in idmo the termen is nearly straight,
receding from near the apex to the tornus, the shape and
width of the brown costa and termen are very different,
and below the posterior catenulated stripe is different in
position and shape whilst the under-side of the secondaries
differs considerably in pattern also.
OagyRis IpMo, Hew.
O. idmo, Hew. Cat. Lyc. B. M., p. 2, pl. I, figs. 3,4, 1862.
Ill. Diurn. Lep. Lye. p. 2, 1863; zd. Kirby, Cat. D. L.,
p. 425, 1871; id. Semper, Journ. Mus. God., p. 55, Heft
14, 1878; wd. Miskin. Proc. Lin. Soc. N.S.W., p. 24,
1890; id. idem. Ann. Queensl. Mus, p. 72, 1891; id. And.
and Spry. Vict. Butt, p. 104, 1894; id. Waterhouse,
P.L.S., N.S.W., p. 248, 1908; id. idem. Mem., N.S.W.
Nat. Club, p. 29, 1903; O. orontas, Hew., Cat. Lyc., B. M.,
p. 2, pl. I, figs. 8 and 9, 1862.
¢. Both wings dull brownish-purple or purplish-brown, in
certain lights the purple has an almost red lustre. Primaries with a
very narrow almost linear-brown termen. Secondaries with costa
brown to vein 7, termen very narrowly brown. Under-side, both
wings brown finely irrorated with pale grey. Primaries with three
The genus Ogyris. 275
increasing cell spots, edged with pale blue or blue and white lines ;
beyond the third spot, which is very large, is ‘a broad fascia of
greyish-white scales, followed by the posterior catenulated stripe
from the costa to vein 1, the spots on the costa being large and
tapering rapidly to the third spot, the least trace of a subterminal
line. Secondaries more thickly covered with pale grey irrorations,
the basal spots are obscure and only indicated by the very fine
darker encircling lines, the median row of three spots irregular
that in the cell very obscure ; the third series of spots is very con-
fluent, the one on the costa being isolated and the rest all confluent ;
the posterior row of four or five spots is irregular and fractured,
the first and second below vein 6 being confluent and touching the
previous series, together they form a reniform spot, spots 3 and 4 are
projected outwards, detached from 1 and 2, but touching each other,
spot 5 shifted inwards and isolated.
2. Both wings brown with a dull purplish lustre over a portion
of the wings. Primaries with the purplish lustre over half the cell
and extending broadly for two-thirds of the inner margin, a dark
large spot at the end of the cell followed by a larger lemon cream-
coloured spot. Secondaries with the purplish lustre over three-
fourths of the wing leaving merely a very broad brown border to the
costa and termen. Under-surface similar to the male but in the
primaries the pale cream spot shows through, and in the secondaries
the pattern is much more distinct.
Expanse ¢ 52-57, 2 57-60 mm.
The distribution of this species issomewhat uncertain,
but I believe that it is confined to Western Australia,
and that the species found in South Australia and Victoria
will all prove to be halmaturia and waterhousert, Br
I have seen no true idmo from either of these localities.
I have before me now Hewitson’s type specimen of orontas
and it is identical with idmo. I have pointed out the
differences between these species, and I would further state
that I regard Tepper’s species as distinct from ofanes, Feld.,
both of which types are now before me.
OGYRIS OTANES, Feld.
Ogyris otanes, Felder Reise. Nov. Lep. II, p. 217, taf. 28,
fig. 1-3, 1865 ; id. Kirby, Cat. D. L., p. 425, 1871; 2d.
Semper, Journ. Mus. God. p. 55, Heft 14, 1878 ; 7d. Miskin.
P.L,S8., N.S.W., Ser. 2, V, p. 23, 1890; zd. tdem. Ann.
Queensl. Mus. No. 1, p. 71, 1891; id. Waterh. (in parte)
276 Mr. G. T. Bethune-Baker on
P.L.S.,N.S.W., p. 249, 1908; cd. idem. Mem, N.S.W. Nat.
Club, No. 1, p. 29, 1903.
¢. Both wings dull reddish brown purple, primaries with the
brown termen narrow straight and of uniform width. Secondaries
with the costa and apical area very broadly brown, the latter taper-
ing into the wide brown termen, the tail at the tornus is somewhat
developed, but not sufficiently as to be called a tail. Under-side.
Both wings pale brownish, primaries with two obscure spots in the
cell, finely divided by bluish white lines, followed by a large black-
ish patch with an irregular pale bluish pupil. The posterior
catenulated stripe is composed of five irregular dark spots palely
edged, followed by an obscure trace of a sixth shifted inwards.
There is no trace of any subterminal or terminal line. Secondaries with
pattern very obscure, the basal series of spots is barely discernible, in
the median series the costal spot is very large, that in the cell much
smaller and irregular, that on the inner margin smaller still, the
series at the end of the cell is confluent irregular extending from the
costa to the inner margin, the second spot (from the costa) being very
large and touching the posterior series which is composed of a pair
of confluent spots followed by a second pair of confluent spots shifted
outwards, beyond which is an isolated spot shifted well inwards.
?. Both wings brighter purplish, primaries with the wing beyond
the cell and above vein 3 brown. A darker patch at the end of the
cell followed by a pale lemon-coloured smallish patch. Secondaries
with the costa above vein 6 brown and a broad brown termen.
Under-side like the male but irrorated with grey so that the pattern
is more distinct and the pale spot shows through in the primaries,
Expanse ¢ 45, 2 48 mm.
Through the kindness of Mr. Rothschild I have Felder’s
types before me and am therefore enabled to compare
them with specimens sent me from South Australia.
The shape of the insect is that obtaining in 7dmo, Hew.,
the apex of the primaries being somewhat acute and the
termen straight ; the colour also is nearest to that species,
but is paler and somewhat brighter, it is however smaller
and the under-side pattern is quite diverse. Mr. Water-
house has kindly sent me for examination two specimens
from Kangaroo Island with a query as to whether they
are Felder’s insect, but after a very careful comparison I
believe them to be distinct, and they are the form named
by Tepper halmaturia. I have now before me the type of
this species as well as Felder’s type (I must here express
The genus Ogyris. 277
my best thanks to Mr. Tepper for the loan of it), and I
consider that they are distinct forms; more material may
prove them to be sub-species, but they differ sufficiently to
warrant them being named. The locality for ofanes will
therefore be South Australia only.
OGYRIS HALMATURIA, Tepper.
0. halmaturia, Tepper Com. Ins. 8. Aust. II, p. 12,
1890. O. otanes, Feld., Waterh, P. L. S., N.S.W., p.
249, 1903 (part); id. idem. (part) Mem. N.S.W. Nat. Club,
p. 29, No. 1.
¢@. Both wings brighter purplish. Primaries with the termen
broadly brown, broadest at the apex and tornus, the purplish area
terminating in an even curve from the costa to the inner margin.
Secondaries with the purple area almost confined to the cell and
about two-thirds beyond it. Under-surface like otanes, Feld., but
much greyer. In the primaries the catenulated posterior stripe is
curved, not straight as in Felder’s species. In the secondaries the
pattern is more obscure and there is a broadish indefinite band of
darker shading outside the posterior stripe which is more marked in
the female than in the male.
Q. Like the male in all respects except that the colour is brighter
and there is the pale spot on the primaries.
Expanse ¢ 46, 2 50 mm.
The types from Kangaroo Island are in the 8. Australian
Museum. Mr. Waterhouse also has specimens from the
same locality.
This species may be distinguished from otanes, Feld., by
the marked arched and broad termen to the primaries, by
the broad brown termen in the secondaries, and by the
shape of the wings, the termen being arched and the
apex rounded.
It is very desirable that a trip should be made to
Kangaroo Island and also to the localities in South
Australia from Cape Willoughby to the south-west corner.
The species frequents broken country, thinly studded
with Melaleuca shrubs, between which, Mr. Tepper tells
me, they sailed in couples but were very wary and difficult
to approach. It is however much to be wished that a
good series of both these closely-allied species could be
obtained, so that we could see whether the distinctions are
constant,
278 Mr. G. T. Bethune-Baker on
OGYRIS MEEKI, Roths.
O. meeki, Roths. Nov. Zool. VII, p. 274, pl. V, fig. 1,
1900.
The female of this species is as yet unknown; we may
expect it to have three tails and to have a pale spot on the
primaries; the male is the largest of the genus and the
only species that has fully developed tails; the outline of
wings is different to all others inasmuch as the costa of
the primaries is more strongly arched and the apex
produced, with the termen slightly excavated below it,
thus giving it a very distinctive appearance from the rest
of its allies ; it may also be recognized by the uniform deep
purplish tone of its upper-side which is quite different from
the colour of any other species of the genus.
OGYRIS ZOSINE, Hew.
O. zosine, Hew., Exot. Butt., I, pl. I, figs. 3, 4, d, 1853.
id. Kirby, Cat. D. L., p. 423 (1871). id. Semper, Journ.
Mus. Godef. Heft 14, p. 55, 1878. id. Hew., Cat. Lye.
B. M., pl. I, fig. 7, ¢ (mec 2), 1862. 2d. idem. Ill, Diurn,
Lep., p. 2, 1863. 7d. Miskin, Ann. Queensl. Mus., No. 1,
p. 71,1891. 0. genoveva, Hew., Exot. Butt. I, pl. I, figs.
5, 6, 9, 1853. id. Semper, Journ. Mus. Godef. Heft 14,
p. 55, 1878. zd. Hew., Cat. Lyc., B. M., p. 3, 1862, ad. Il.
Diurn. Lep., p. 2,1863. id. Staud., Exot. Schmett. taf 96,
1888. id. Miskin, P. L.8., N.S.W., p. 23,1890. zd. Miskin,
Ann. Queensl. Mus., No.1, p. 71, 1891. 7d. Waterh., P. L.S.,
N.S.W., p. 245, 1908. id. idem. Mem. N.S.W. Nat. Club,
p. 29, 1903.
This species is probably the best known of the genus,
and is apparently becoming a dominant species; there are
already three distinct local races in different parts of the
country, that in the south being the brightest of all,
whilst the type form obtaining in Townsville and its
neighbourhood is the most sombre in colouring.
It has hitherto been known in all collections with which
IT am acquainted as O. genoveva, Hew. It is difficult to
discover how this has arisen, because Hewitson described
zosine (l.c.) previously, and also figured it previously on
the same plate. It is perhaps commoner in Queensland
than elsewhere. The § is dimorphic. I have therefore
retained the name genoveva for the pale blue 2 as described
The genus Ogyris. 279
by Hewitson. There are however two other easily recog-
eae forms obtaining in other localities in Australia, the
one from Coomooboolaroo, N. Queensland, and the other
from Brisbane; these appear to be good constant local
races, so I have no hesitation in naming them. Mr.
Waterhouse tells me that he has taken this species feed-
ing on Loranthus linophyllus, Feu., at the Richmond
River, and on JL. celastroides, Sieb., at Sydney, The egg is
plain, spherical, flattened at the top, and is usually laid on
the mistletoe stem near its origin with its host. The
young larvee are uniformly light brown (F. P. Dodd), and
subonisciform ; when full grown, however, iBey become less
woodlouse- like, as they then measure from 25 to 32 mm.,
but when in motion they are considerably longer ; in colour
they are dirty cream colour to ochreous yellow above, and
purplish below, with black spiracles, this being the form
that is found in North Queensland ; those found in the
South however differ, and are, according to Waterhouse,
brownish-red above and yellowish below. They hide in
the cracks of the bark of the host tree or in holes in the
mistletoe during the day, or beneath the surface of the
earth, coming out at dusk and feeding at night, at which
time I am informed the ants associated with them are
likewise on the move; the species of ants that Dodd has
found them with most commonly is Mcophylla virescens,
but several other species also associate with them. They
evidently protect the larvee, and have been observed to
milk them; in one instance an ant was observed to
approach a larva and wave its antennz over its terminal
segments, ae then to lightly touch it with its fore-leg
when a small globule of liquid was emitted from a small
retractible nipple-like organ on the dorsum which was at
once sucked up by the ant. The process was then repeated
on the other side, there being two of these organs, one on
each side of the dorsum ; this happened two or three times,
and the larva seemed quite composed and in no way
incommoded by the incident. I possess larvae in spirit
both from Queensland and from Sydney, and the form
from Sydney is darker and more highly coloured than that
from the North. The pupa, some of which I have before
me, is very dark brown, almost blackish, of the usual
Lycxnid form ; it attaches itself by its terminal segments
to the under-side of stones or loose bark and spins a girdle
of silk across its back about the end of the wing cases and
280 Mr. G. T. Bethune-Baker on
so undergoes its transformation, and is quite undisturbed
and unmolested by its ant hosts.
Mr. Waterhouse tells me that the form found at the
Richmond River (N.S.W.) is distinctly smaller than that
found both to the North and to the South of that locality,
he says also that he never sees males on the wing until
about 2 p.m. in the day, when they fly high (about thirty
feet) around the Hucalyptus trees, and are very difficult to
catch ; the females are very rarely seen on the wing.
This year the species has been recorded from South
Australia, Mr. Lyell having recorded it in the Victorian
Naturalist, vol. xxi, p. 166 and 167; from Dimboola,
larvee and pupz were taken in November and December
last, the perfect insect emerging in the latter month. Mr.
Lyell and his friend Mr. Fricot were hunting for the larvee
of O. idmo, but instead of finding it they discovered
caterpillars of this species; they confirm the observa-
tion that it feeds by night, and they also state that it
pupates at the foot of the tree below the surface of the
ground. In order to test the action of the attendant ants
one or two larvee were placed a couple of feet or so away
from the tree; they were however soon discovered and
dragged carefully back to the tree by the ants at a pace
much more rapid than their own rate ; pupz were likewise
carried back to the tree.
OGYRIS ZOSINE-DUARINGA, sub spec. nov.
Ogyris genoveva, Miskin, Trans. Ent. Soc. Lond., p. 343,
pl. xv.
¢. Both wings brightish purple. Primaries with a broad darker
but equally bright suffusion across the median area of the wing, the
brown termen is much narrower than in zosine and of equal width
from the apex to the tornus. Secondaries with a very narrow brown
termen, little more than linear. Under-side as in the type but with
the spots more distinctly outlined and with a velvety clouding near
the tail at vein 4.
?. Similar to the female of the form genoveva, Hew., but the blue
instead of cerulean is brilliant greenish-blue much more lustrous
than in Hewitson’s insect.
Expanse ¢ 54-58, 2 56-58 mm.
This subspecies was taken by Meek at Coomooboolaroo,
where it appears to be a constant race; there is a long
The genus Ogyris. 281
series of it in the Tring Museum; it is also the form
figured by Miskin (I. ¢.).
OGYRIS ZOSINE-MAGNA, sub spec. nov.
¢@. Both wings rich purplish-blue. Primaries with the apical third
of the costa and the apical half of the termen hoary. Secondaries
dark brown to vein 6. Tail at vein 4 not developed, the scallop
being but little longer than that at the end of veins 3 and 5.
Fringes white, interrupted at the veins, antennz bright chestnut,
very finely dark on the upper-side, the club being bright orange
chestnut. Under-side as type species but darker grey.
@. Both wings velvety black. Primaries with the basal area to
about half the cell and half the inner margin brilliant silvery
lustrous greenish-blue more greenish than in duwaringa, and with the
usual pale spot. Secondaries with the silvery lustrous greenish area
more reduced extending only to just beyond the cell and tapering in
an arc to three-fifths of vein 2. Veins 2 and 4 are developed into
strong broad tails, and vein 3 into a prominent tooth, at each of the
latter is a lustrous greenish blue spot. Under-side similar to the
type species but more strongly marked.
Expanse ¢ 66, 2 70 mm.
The types are from Brisbane, and are in my collection,
Mr. Waterhouse also has it from the same locality.
OGyRIS ABROTA, Doub. and Hew.
Ogyris abrota, Doubleday and Hew., Gen. Diurn. Lep.,
pl. 75, fig. 8 , 1850; zd. Hew. Exot. Butt. I, pl. I, figs. 1
and 2 f, 1853; id. zdem. Cat. Lyc., B. M., p. 2, 1862; 2d.
idem. Il, Diurn. Lep. Lyc., p. 2, 1863; id. Kirby, Cat.
D. L., p. 425, 1871; 2d. Semper, Journ. Mus. Godef., Heft
14, p. 55, 1878; zd. Miskin, P. L: S., N.S.W., p. 25, 1890;
id. idem. Ann. Queens. Mus., No. 1, p. 72, 1891; 7d. And.
and Spry., Vict. Butt. p. 109, 1894; id. Waterh.,
P.L.8., N.S.W., p. 247, 1903; zd. idem. Mem. N.S.W.
Nat. Club, p. 28, 1903. 0. damo, Doubld. List. Lep., B. M.,
pt. 2, p. 20.
This species is recorded from Victoria, New South Wales
and South Queensland. Anderson and Spry have described
its metamorphosis, they say that the ova are dull white and
quite globular and are laid on the inside of loose bark by
282 Mr. G. T. Bethune-Baker on
the food plant or upon its stems. The larvee feed chiefly
upon Loranthus pendulus, and are onisciform, broadened
and flattened at the anal segments and with a dorsal ridge ;
the general colour is dark fulvous paler along the ridge
with a series of oblique (? lateral) stripes; on the tenth
segment is a black irregular diamond-shaped patch pupilled
with greyish-white, the last segment having a dark T-
shaped mark, the spiracles are white encircled with black,
the surface generally being rugose and punctated with
minute bristles, with a subspiracular series of short bristles.
‘The larval state is said to last for some months and to be
delicate and difficult to rear; ants are always with them,
and whilst they appear to be free from attacks of insect
parasites, a fungoid disease at times kills large numbers
of them. When full grown the larva measures from 20
to 25 mm.
OGYRIS IANTHIS, Waterhouse.
O. ianthis, Waterhouse, P. L. 8., N.S.W., pp. 52-54,
pl. I, figs. 1-4, 1900; id. cdem. p. 841, 1902; id. p. 247,
1903; Mem. N.S.W. Nat. Club, No. 1, p. 29, 1903.
This species can readily be recognized from Water-
house’s description and from its rich orange-chrome-
coloured female; it is apparently confined to the neigh-
bourhood of Sydney. The ova are pale pinkish in colour,
in shape they are flattened somewhat at the top, and
reticulated all over. The larva, which hides at some depth
in ants’ nests during the day, does not move until night, and
feeds on Loranthus celastroides in the dark, always making
for a dark corner should a light be turned on it; it is
pinkish-brown with a much darker dorsal line and paler
lateral bands, one on each segment; the under-side is pale
cream colour. The pupa is reddish-brown of the usual
shape. I find that the organs on the last segment but one
through which the fluid so greedily sucked by ants is
passed are very marked in this species. I have a young
larva in formalin now in front of me, and I notice that
these organs (a pair) are situated on each side of the
dorsum, and consist of a prominent tubercle shagreened all
over, with a deep orifice in the centre from which the
retractile tube is everted or withdrawn at will; when at
rest the tube is contracted, and is apparently only emitted
when the larva is induced to do so.
The genus Ogyris. 283
OGYRIS OLANE, Hew.
Ogyris oiane, Hew., Cat. Lyc., B. M., p. 2, pl. I, figs. 10,
11, 1862; 7d. Ill. Diurn. Lep., p. 2, 1863; zd. Miskin,
Pobirs, NS. W., p. 27, 1890° cdi Karby, Cat” D! T., p.
425, 1871; id. Semper, Journ. Mus. God., Heft 14, p. 55,
1878; id. Miskin, Ann. Queensl. Mus. No. 1, p. 72,
1891; O. catharina, Feld. Reise, Nov. Lep. II, p. 218,
1865.
d+. Both wings darkish-brown. Primaries with a patch of dull
brownish-purple occupying the area between the median and sub-
median veins extending slightly into the cell and one-third the
space beyond below vein 2. Apical area and half down the termen
paler brown. Secondaries with the purple occupying the cell and’
one-third the space beyond, termen strongly crenulate.
Fringes white interrupted at the veins.
Under-side primaries dark brown with the cell spots defined by
five pale metallic-blue lines, a dark irregular oblique line midway
between the cell and the apex, beyond which the wing is suffused
with whitish-grey scales, an obscure subterminal band tapering
towards the apex extending into the grey area. Secondaries grey
darker towards the inner marginal and tornal area, spots slightly
darker defined by fine dark lines. Basal spots small, median series
large, the first below the costa narrow touching the large spot across
the cell which is confluent with the one below it, an isolated long
oblique spot from the costa to the large cell spot just mentioned,
third series, with the first and second spots confluent, the first
strongly oblique, the second an inverted A-shape confluent with the
two smaller spots below it, posterior series from vein 6, the second
and third spots confluent shifted outwards, the fourth still outwards
and slightly darker, fifth and sixth smaller and shifted inwards, a
trace of a subterminal dark irregular line, lower terminal area
darkly suffused.
Q. Like the male but with the purple area very slightly less in
extent, whilst the secondaries on the under-side have the spots
larger and darker thus forming a strong contrast with the grey
ground-colour.
Expanse: ¢ 42-44, ? 44-49 mm.
Hab. S. AUSTRALIA, VicToRIA and N. S. WALES.
This description is of the form now found in Australia ;
it does not agree with the colour on the upper-side of
Hewitson’s type, which is paler and bright blue, the under-
284 Mr. G. T. Bethune-Baker on
side agrees, however, fairly closely. I have no doubt that
the two insects are the same species, but that Hewitson’s
type may have undergone some change (possibly chemical)
which has effected the difference in colour. In my study
of the genus Arhopala I became convinced that the blues
of that genus, hitherto considered so very constant and
unchangeable in colour, are liable to alteration, this being
especially so where the colours are apparently aniline, and
further experience tends to confirm that opinion, though I
am not prepared to say in what manner the change of
tone is effected, but it apparently does not affect the scales
themselves. ;
Anderson and Spry describe the ova as of a pearly lustre
and flattened at the poles; they are laid on the edges of
loose bark, or on the stems of Loranthus pendulus and some-
times on the galls which furnish hiding-places for the
larvee ; these closely resemble the larvee of 0. abrota, Hew.,
but have a small black patch on the anal plate; they are
at times great wanderers, and are strictly nocturnal feeders
and have to travel long distances from their hiding-places
to their food, which in the summer months is often scarce,
as the Loranthus loses most of its leaves then; they appear
to be able however to withstand prolonged abstinence
and yet to undergo safely their metamorphoses, in these
cases however the imagines are of course smaller than
usual, The larve are also greatly subject to parasites, their
worst enemies being two species of Diptera, one of which
lays its eggs in its host, whilst in the case of the other fly,
apparently the larve is not a parasite, but is furnished
with pointed mandibles and is said to suck its victims dry.
This dipterous caterpillar being very active and voracious
works serious destruction amongst the larve of olane.
The pupa is somewhat delicate and sutfers from the rays
of the sun if in too exposed a situation; it remains about
a month in this state before emerging as the perfect insect.
Isolated specimens may be caught throughout the year,
but October and November, and February and March are,
according to Anderson and Spry, the months when they
usually fly, though it is difficult to take, owing to its habit
of flying around the topmost branches of the gum trees.
Mr. Spry informs me that he has never once seen this
caterpillar attended by ants, though he has studied it and
known it for years, the fact also that it is subject to the
attacks of parasites no doubt corroborates his observations.
The genus Ogyris. 285
OGYRIS BARNARDI, Miskin.
Ogyris barnardi, Miskin, Proc. Linn. Soc. N.S.W., p. 27,
1890 ; ad. idem. Ann. Queensl. Mus., No. 1, p. 72, 1891;
id. Waterhouse, idem. p. 248, 1908; id. Waterhouse, Mem.
N.S.W. Nat. Club, p. 28, 1903.
3. Both wings dull purple. Primaries with termen narrowly
brown, slightly broader at the apex. Secondaries with costa some-
what broadly brown and very narrow brown termen ; termen very
slightly crenulate. Under-side like olane, Hew., but the primaries
are paler brown, and in the secondaries the spots are smaller and
there is no dark suffusion.
Expanse ¢ 41-45,
Dawson River, Peak Downs (QUEENSLAND).
OGyRIS ORG@TES, Hew.
O. oretes, Hew., Cat. Lye. B.M., p. 3, pl. I, figs. 12
and 13, 9, 1862; dd. idem. Ill. Diurn. Lep., p. 2, 1868;
id. Kirby, Cat. D. L., p. 425, 1871; 7d. Miskin, P. L.S.,
N.S.W., p. 25, 1890; 2d. idem. Ann. Queens]. Mus., I,
p. 71, 1891; id. Waterh. P.L.S., N.S.W., p. 335, pl. XIV,
figs. 1 and 2, 1902; id. idem. p. 246, 1903; id. idem.
Mem. N.S.W. Nat. Club, I, p. 29.
The only locality of which we are certain at the present
time is Queensland. Mr. Waterhouse informs me that the
specimens mentioned by Miskin (l.c.) from W. Australia
and Victoria are hewitsoni, whilst the ? in the Australian
Museum is a f amaryllis. Dodd has bred the species
plentifully, and he tells me that the full-grown larve are
light yellowish-brown faintly tinged with green, and that
he has always found them among or close to communities
of ants. The pupz are dark brown, and are of the usual
shape; I have specimens now before me, but there is
nothing worthy of note in them. Dodd says they “ tick”
in the same way as genoveva, only decidedly more slowly
and not so loudly.
OGYRIS HEWITSONI, Waterh.
O. hewitsoni, Waterh., Proc. Linn. Soc. N.S.W., p. 338,
pl. XIV, f. 5-8, 1902; id. idem. p. 246, 1903; id. idem.
Mem. N.S.W. Nat. Club, p. 29, 1903; O. amaryllis, And.
and Spry, p. 102, 1894.
TRANS, ENT. SOC. LOND. 1905.—PART III. (00T.) 20
286 Mr. G. T. Bethune-Baker on
The female of this insect is unrecognizable on the upper-
side from the ? of the oretes, except that in the present
species the brilliant blue extends over the upper margin
of the cell, whilst in Hewitson’s species it does not.
Under-side, like the male, except that the pattern is
generally more distinct and isolated. In the cell of the
primaries are two large vermilion red spots edged on each
side with pale metallic bluish, and with traces of red
further along the cell.
This species seems to be the commonest of all the genus
with the exception possibly of genovera. It has been
recorded from al] the Australian States except Tasmania.
I am again indebted to that careful observer, F. P. Dodd,
for information as to the life history. He finds the larvee in
the same localities as orwtes always among or near ants;
when full fed they are grey or greyish-brown, and are
duller in colour than the other species he has taken; the
pup emit similar sounds, and like the preceding insect
“tick” more slowly than genoveva. ‘These sounds are not
continued for any length of time, but a gentle touch or
a shake will generally set them going again, and when
one specimen begins others in the vicinity. as a rule follow
its example. Mr. Dodd tells me that each of the three
caterpillars of this pens that he is acquainted with feed
at night only.
OGYRIS HEWITSONI-MERIDIONALIS, sub spec. nov.
¢. Both wings with the blue decidedly deeper and less silvery in
tone. Under-side altogether darker, the cell markings larger and the
white edgings tinged with blue. Secondaries browner, with none
of the strong contrasts that are so conspicuous in hewitsoni; the
spots are browner and not so broken up as in Waterhouse’s type
race.
9. Both wings with the blue of a lilac lustre, not silvery.
Under-side similar to the type form.
Expanse ¢ 40-41, 9 42 mm.
The Southern form from Victoria is very decidedly less
brilliant than Waterhouse’s species, it is also much smaller.
I have it from several localities, and as Mr. Waterhouse
says that these differences are constant, it seems to be
advisable to name the local race.
The genus Ogyris. 287
OGYRIS AMARYLLIS, Hew.
0. amaryllis Hew., Cat. Lyc., B. M., p. 3, pl. I, f. 5 and
6, 2, 1862 ; ad. adem. Il. Diurn. Lep., p. 2, 1863 ; ad. Kirby,
Cat. D.L., p. 425, 1871; id. Miskin, P.L.8., N.S.W., p.
26, 1890; 2d. idem. Ann. Queensl. Mus., p. 72, 1891; id.
Waterh., P.L.S., N.'S.W., p. 336, pl. XIV, f. 3, 4, 1902;
id. idem. Mem. N.S.W. Nat. Club, p. 29, 1903.
This species can easily be recognized from its pre-
decessor by the very broad borders to the primaries and
by the deeper tone of blue in both wings.
The range of the species appears to be restricted to the
neighbourhood of Brisbane (S. Queensland) and the
Richmond River in New South Wales, from which latter
locality Waterhouse has bred it. The ova are pinkish
grey, somewhat flattened with raised reticulations, and are
laid on the mistletoe knobs. The larvee feed on Loranthus
linophyllus, and when young are greenish, but later they
become dark grey assimilating very closely with the host-
plant of the Loranthus.
OGYRIS ZNONE, Waterhouse.
O. xnone, Waterh. P. L. 8., N.S.W., p. 339. pl. XIV, fig.
9 g, 1902; id. cdem. p. 246, 1908 ; id. idem. Mem. N.S.W.
Nat. Club, p. 29, 1903.
¢. Both wings brilliant lustrous morpho-blue, more brilliant and
lustrous than in any other of the genus, in certain side lights having
a metallic mauve lustre. Primaries with the costa broadly blackish
increasing at the apex, termen narrowly blackish increasing rapidly
towards the apex, and decreasing slightly towards the tornus.
Secondaries with the costa broadly dark greyish and termen very finely
black. Under-side. Both wings pale whitish dove-grey. Primaries
with three increasing cell spots palely edged, the basal spot pale
brownish, the second and third deep black with bluish white
margins ; below each of these latter a black spot, catenulated stripe
very irregular consisting of seven spots, the first two below the costa
quite pale, the third blackish and small, fourth slightly larger shifted
outwards, fifth shifted inwards, sixth very large and ovate inclined
outwards, seventh oval rather smaller shifted and inclined inwards,
a trace of a subterminal line. Secondaries rather darker than the
primaries with three basal spots, the first and third very small
median series widely isolated, a small darkish spot below the costal
288 Mr. G. T. Bethune-Baker on
vein, a larger pale grey spot on the inner margin of the cell with
two small spots below it (one on each side) followed by another pale
spot on the inner margin, third series very irregular, a longish
narrow spot below the costal vein, a pale round one in the angle of
vein 7, a large irregular one closing the cell below which is an
indefinite trace of a fourth followed by a larger and more distinct spot
on the inner margin, posterior catenulated series, irregular and
somewhat indefinite on its inner margin, the first spot large below
vein 7, second and third shifted right outwards, fourth and fifth well
inwards, sixth spot obscure, shifted inwards, an indefinite subter-
minal slightly dentate stripe, a trace of a brownish spot on the
slightly developed lobe at the tornus.
2. Both wings less lustrous and not quite so pale a blue as the
male with broader margins. Primaries with a black invading spot
at the end of the cell and a creamy yellowish costal patch in front of
the apex. Under-side. Primaries like the male but with all the
spots larger and darker, and between the cell spots large patches of
very pale (washed out) orange. Secondaries similar to the male but
with the spots darker, and a white oblique broken stripe right across
the wing across the middle of the cell edged by a broad indefinite
suffusion of golden brownish, termen with a similar suffusion.,
Expanse ¢ 52, 9 54 mm.
This species is only recorded from Cooktown; the
brilliancy of its upper-side and the paleness of the under-
side, together with the small and isolated arrangement of
spots beneath, will enable it to be readily separated from
any other of the group.
289
The genus Ogyris.
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Explanation of Plate.
EXPLANATION OF PLATE XV.
Janey te
La.) ie
ade 9
2a. +
9a.
10a: =, 4;
lla.
Ogyris amaryllis, genitalia profile
= penis.
otanes, genitalia.
» penis.
halmaturia, genitalia.
i penis.
oretes, genitalia.
» penis.
zosine, genitalia.
9 penis,
abrota, genitalia.
» penis,
barnardi, genitalia.
diz)
ff penis.
olane, genitalia.
» . penis:
ianthis, genitalia,
» penis.
idmo, genitalia.
» ~~: penis,
hewitsoni, genitalia
i, f eNIC,
( 9889
XVII. The structure and life history of Psychoda sex-
punctata, Curtis. By JoHN ALEXANDER DELL,
B.Sc. Communicated by Prof. L. C. MIALL,
F.RS., F.ES.
[Read May 3rd, 1905.]
Halitat and Mode of Life.
Psychoda in all stages of growth abounds throughout
the year at the Leeds Sewage Works, and advantage has
been taken of this circumstance to investigate more fully
the life history of the insect. It obtains its food from
heaps of coke, over which crude sewage is sprayed during
the purifying process. The life which flourishes on the
coke-heaps, and especially the bacterial life, effects the
reduction of the sewage to a harmless liquid. Alga,
among which Stigeocloniwm tenue is conspicuous, find here
congenial conditions, and supply nutriment to insects of
more than one kind. Besides Psychoda, a Chironomus and
a beetle (Platystethus) occur. A Scolopendra, Mites, and
Nais have also been observed on the coke-heaps.
The Psychoda-larva is minute, cylindrical, tapering
toward the hinder end, and bluntly rounded in front. It
may attain a length of 75 mm. The larva bears no
locomotor appendages, but moves in a vermiform manner.
On the fore part of the head are a pair of jaws, probably
mandibles, which can be extended forwards, or folded
backwards. By means of these and the recurved labrum
food is crammed into the mouth. The last segment is
usually bent upwards so as to keep its tip, which bears the
only functional spiracles, clear of the semi-liquid filth in
which the rest of the body is usually immersed. ‘lhe
pup are found together with the larvee in the sewage.
The fly is small and of grey colour. The female is 4 to
4°5 mm. long, the male only 25 to 3 mm., including the
wings. The male is further distinguished by the large
genital appendages, which project from the extremity of
the abdomen. The wings are longer than the body, which
they completely cover, and slope when at rest. Both
TRANS, ENT. SOC. LOND. 1905.—PART III. ( OCT. )
294 Mr. J. A. Dell on the Structure
body and wings are abundantly covered with grey hairs,
which give the fly at first sight the appearance of a small
moth, hence the name Psychoda. Round the edges of the
wings are a few black tufts, and the veins are thickly
covered with hairs. The hairy surface protects the body
from accidental wetting. A fresh-killed fly, when pressed
below the surface of the water, carries down with it so
large a quantity of air, that on being released, it shoots at
once to the top and remains floating. The flies may often
be observed resting on the windows of empty houses, and
are also abundant in privies, urinals, ete. At the Leeds
Sewage Works they cluster on the under-side of the pieces
of coke upon which the sewage is sprayed. They are
sluggish, and do not readily take to flight. In returning
from the sewage works to the University I have often
found that the walk of three or four miles was insufficient
to dislodge Psychoda flies which had clung to my dress.
They can, however, run fast, and they are sometimes
carried far by wind. The fly does not, so far as is known,
feed at all.
In copulation the male runs alongside the female,
stroking her with his antenne, while the wings, antennz
and halteres are thrown into spasmodic vibration. The
large forceps is then extended directly backwards, and the
abdomen of the male bent round so that the hinder ends
of the two bodies are brought into apposition. The
forceps then close upon the body of the female, unless
they miss, which not infrequently happens, as the male
fly is unable to see what he is doing. In such a case the
whole manoeuvre is repeated. The males are apparently
unable to distinguish which of their companions are
females, as they very frequently attempt to copulate with
one another. They have often been seen to die while
still attached to the female.
As usual in Nemocera the eggs are deposited in an
elastic, jelly-like mass. They are irregularly arranged in
it, and unconnected with one another. The egg-mass
thus formed is indefinite in shape, and may be fixed to a
stone or other solid object. The total length of an egg-
mass may be 1-2 mm., and such an egg-mass commonly
contains 35-40 eggs. The eggs are small (‘2 mm. to 3
mm. in length) and sausage-shaped ; they are opaque, and
loaded with yolk-granules. Hatching takes place 10-14
days after laying, but may possibly be more rapid in a
and Life History of Psychoda seapunctata. 295
warmer season than that in which WAN
my observations were made.
Systematic position of Psychoda
sexpunctata.
The long antennee of many similar
joints, and the slender flexible maxil-
lary palps, at once place Psychoda
among the Nemocera. The absence
of ocelli, the short and fleshy mouth-
parts, the bead-like joints of the
antenne, the prominent genital
forceps, and the structure of the j
wings characterize the Psychodide.
The pattern of the wing-veins suffi-
ciently indicates the genus Psychoda, /
while the black tufts on the margin
of the wing distinguish the species
sexpunctata.
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External features of the Larva
(Fig. 1).
The body consists of a head and
eleven segments. Lach of the first
three segments is imperfectly
divided into two annuli, and each
of the remaining ones into three.
The last segment is the only one
which is extensively modified. The
integument is covered with a thick
layer of chitin, which is in places
thickened into special protective
plates. In general there is a thin
membrane between the adjacent
segments, and a longitudinal thin
strip separating the dorsal from the
ventral armour. The cuticle of a
larva which has recently shed its
skin is white and transparent, and
such larvae can be easily studied
PR
IN WNT ne
alive as transparent objects. As rey
time goes on, however, the cuticle 4
thickens, and a larva which has a Fic. 1( x 20).
moult in prospect becomes com- Dorsal view of larva.
296 Mr. J. A. Dell on the Structwre
pletely brown and opaque. From the middle of each
segment a pair of lateral sete project, which are more
or less curved. There are also on all the segments
except the first and second numerous short, rather blunt
sete, arranged in transverse bands; there are two of these
bands on the third segment, and three on each of the
others. In the mid-dorsal line of the ninth and tenth
segments occur a series of black, oval, shining plates,
transversely elongated, but not reaching the sides of the
body. There are usually three on each segment, but they
are irregular in form and size, and occasionally one is
found on the eighth segment. The last or eleventh
segment is smooth and hard, and tapers backwards, though
Fig. 2 ( x 80).
Dorsal view of head of larva (to left); ventral view (to right).
the extreme tip is truncated. The anus opens on its
under-side, in a recess at the attached end of the segment.
There are two small appendages arising from the truncated
terminal surface of this segment. They are short, fringed
with setae, and somewhat like the corresponding structures
in the Pericoma larva, but not so much developed. As in
Pericoma, the larvee can hang from the surface-film by
means of the anal appendages. If depressed beneath the
surface, the anal appendages do not carry down a bubble
of air, with which the spiracles communicate, as in the
Pericoma larva, though small air bubbles often adhere to
the sete. When floating at the surface, the larva, if it is
able to reach the bottom, can drag itself along by means
and Life History of Psychoda seapwnctata. 297
of the mouth-appendages, which are rapidly extended and
closed, so as to scrape the bottom.
The head (Fig. 2) is subconical, much reduced, and
partly retractile. It can be withdrawn into the prothorax,
so that the head is concealed as far forward as the eyes.
This position is temporarily assumed during crawling, and
is normal in larve about to pupate. The head nearly
resembles that of Pericoma* and Maruina.t In each case
the mid-dorsal surface is occupied by the roughly triangular
clypeus, flanked on either side by the large epicranial
plates, which bear the black, shining, simple eyes, and the
antenne. The antennz are even more reduced than in
Pericoma, and are little more than knobs. The forward
prolongation of the clypeus is the labrum. This is an
elongated triangular organ ; its apex is turned underneath
in such a way that the tip lies almost directly beneath its
junction with the clypeus. A few hairs arise from its
anterior edges, and on the surface of the apical part, which
is morphologically dorsal, but actually ventral, there occur
a large number of backward-directed spines. On each
side of, and immediately behind the labrum is an appen-
dage, probably the mandible, which is short, and ends
bluntly. From each appendage arise two large plumose
sete, one running forward, the other outward. There are
also two bunches of shorter setae. Immediately behind the
supposed mandibles is a pair of semi-circular, plate-lke
appendages, which probably represent the maxilla.
Between them and in the median line is a comb-lke
organ with its teeth projecting forwards. A similar plate
is common in Nemoceran larvee, and has been identified
with the submentum.t
Nervous System of the Larva.
The nervous system (Fig. 3) consists of the usual cere-
bral ganglia and the ventral chain. The cerebral ganglia
are pear-shaped, and from their pointed ends arise a pair
of nerves, which run forwards into the head. The sub-
cesophageal ganglia are, as usual, connected with the
cerebral so as to form a ring round the cesophagus, through
which runs the dorsal vessel. Behind these is a series of
* Miall and Walker (1895). + Miiller (1895).
+ Miall and Hammond (1892 and 1900).
298 Mr. J. A. Dell on the Structure
eleven ganglia, not segmentally arranged. The hindmost
of the chain is small, and close to the one immediately in
front of it. The first body-segment contains, in addition
to the first ganglion of the ventral chain, the cerebral and
sub-cesophageal ganglia, which in most insects lie in the
head. The shifting of these ganglia into the thorax is no
doubt explained, as in the case of Chironomus, by the
reduction of the larval head. The large and complex
head of the fly cannot be developed within the
small larval head, and its rudiments, besides the
brain, which it will ultimately enclose, are trans-
ferred to the much more spacious prothorax.*
Alimentary Canal of the Larva.
The alimentary canal resembles that of
Chironomus + so closely that I shall merely
describe the points of difference between the
two. ‘The fore-part of the stomach is separated
as a cardia, but has no ceca. Between its wall
and the reflected wall of the cesophagus arises
a thin chitinous membrane, the “ peritrophic
membrane ” of Balbiani.+
The function of the peritrophic membrane is
to protect the mesenteric epithelium from
abrasion by inorganic particles, which the larva
swallows along with its food. It is apparently
cast periodically. In a living Psychoda larva
reversed peristaltic contractions have been
observed. The digested food has been seen to
be carried up into the space between the
peritrophic membrane and the mesenteric
epithelium by these peristaltic contractions. At
ae the hinder end of the mesenteron is a slight
‘dilatation, into which open the five Malpighian
Nerve-cord tubules. Five is an unusual number of tubules
ormve among insects, but it is found in Blepharoceride,
Culew, and Ptychoptera.§
There is a pair of salivary glands situated in the fore-
part of the thorax. They are roughly bean-shaped, and
the duct arises from the part corresponding to the hilum
* Miall and Hammond (1892, 1900).
+ Miall and Hammond (1900). { Balbiani (1890).
§ Miiller (1895).
and Life History of Psychoda sexpunctata. 299
of the bean. They are placed one on each side of the
alimentary canal, which they partly encircle. The two
ducts run forward beneath the alimentary canal, and unite
in the posterior region of the head. The structure of the
glands is very simple. They are hollow, the wall being
only one layer of cells thick. The cells are very large,
and possess very large nuclei, in which by suitable methods
a distinct chromatic filament and nucleus can be made
out. I have not been able to detect the elaborate
nuclear structure of the salivary cells of the Chironomus
larva.* Outside the cells there is a thin basement-
membrane.
Respiratory System of the Larva (Figs. 4-6).
The respiratory system of the larva is well developed.
The air is renewed mainly, if not altogether, by means of the
large posterior spiracles (Fig. 6). These are situated at the
apex of the last segment, close to the bases of the minute
analappendages. ach spiracle communicates with a large
longitudinal trachea. Immediately within the spiracle the
trachea is slightly dilated into an oval chamber, and the
centre of this is occupied by an ovoid mass of chitin, attached
to the walls of the chamber by chitinous threads. The
whole arrangement forms a kind of loose spongy plug. In
addition to this pair of large posterior spiracles, there are
an anterior pair borne by two short processes on the sides
of the prothorax (Fig. 4). They are not open, and pro-
bably not functional, since they are always immersed in
water or mud. Leading into each spiracle is a trachea,
which, as it passes into the process on which the spiracle
is borne, gradually becomes spongy and takes on a black
appearance as seen by the microscope, probably due to
contained air. |
There are two very large trachee (Fig. 4), each arising
from one of the posterior spiracles, and running forward
along the back. They are connected by a series of
commissures, one of which occurs at the hinder end of
every segment, except the last. In many cases small,
forward-directed branches arise from the commissures.
Some of these are connected with the alimentary canal,
and on this account the commissures are very slack,
especially those near the middle of the body. During the
* Balbiani (1881).
300 Mr. J. A. Dell on the Structure
Fre. 4 ( x 20).
Tracheal system of larva,
vermiform movements of the animal
there is a considerable amount of
sliding of the alimentary canal and
body-wall on one another, and a tight
commissure would be liable to be
snapped, or to tear the tissues. In
each segment, besides the commissures,
there is found a lateral branch running
downward and_ forward, and soon
splitting into three. Of these three
each posterior branch unites with the
fe HN MS A TEE
I
Fie. 5 ( x 20):
Anterior part of tracheal system of larva,
in side view.
anterior branch of the segment behind,
thus forming a loop, while the middle
branch breaks up into branches of
distribution. In the last segment,
immediately dorsal to the hinder part
of the heart, there arise from each
Fic. 6 ( x 100).
Posterior spiracles of larve,
and Life History of Psychoda sexpunctata. 301
Hie. 7 ( x 20).
Ventral view of pupa (to left) ; side view (to right).
TRANS. ENT. SOC. LOND. 1905.—PART UI. (00T.) 21
302 Mr. J. A. Dell on the Structure
longitudinal trachea a number of small branches, which
break up into branches of distribution in the neighbour-
hood of the pericardium.
Pupation: Haternal Features of the Pupa.
After reaching its full size, the larva becomes sluggish
and ceases to feed ; the head becomes retracted, so that the
part behind the eyes is concealed by the first body-
segment ; the last larval skin is then shed, and the pupa
emerges. This takes place while the animal is still buried
in slime. Ina short time the pupa makes its way to the
surface, where it les with the respiratory appendages
exposed, This facilitates respiration, and the escape of
the fly.
The pupa (Fig. 7) is, as usual, shorter and thicker than
the larva. When it first appears it is white, but the
cuticle soon thickens and turns brown. The head is not
externally marked off from the rest of the body, but the
compound eyes of the fly can be seen through the pupal
skin. From the front of the prothorax arise the two
respiratory appendages (Prothorakalhorne of De Meijere*).
They are rather less than one-third of the length of the
body, and transversely wrinkled. Scattered over their
surfaces, especially towards the distal end, occur numerous
transparent circles, which appear to be holes, but are really
thin spots in the cuticle. A shallow transverse furrow
separates the prothorax from the mesothorax. The three
pairs of leg-sheaths are straight, and reach the beginning
of the third abdominal segment, where the tips of the wing-
sheaths are also situated. All the exposed abdominal
segments, except the eighth or last, are protected by
dorsal and ventral plates, whose hinder margins end in
rows of backwardly-directed spines. There are also a few
scattered spines on both dorsal and ventral surfaces. Each
spine is a pointed process of the body-wall. Its apex bears
aspike set in a socket. The last segment bears four large
spines, two terminal and two subterminal. The spines
assist the pupa to move about in the filth in which it
is buried, and so keep the respiratory appendages exposed
to the air.
* De Meijere (1902).
and Life History of Psychoda sexpunctata. 303
Tracheal System of the Pupa (Fig. 8).
The tracheal system of the pupa is closed, the contained
Fic. 8 ( x 20).
Tracheal system of pupa.
air being renewed by means of the
prothoracic respiratory appendages,
a pair of hollow finger-shaped
processes, which project forward
from the prothorax. <A tracheal
extention (Fig. 9)—Hornfilzkammer
of De Meijere—lies within the
respiratory appendage. The structure
of the wall of the pupal respiratory
appendage is not unlike that of
Ptychoptera. In places the tracheal
extension bulges through the wall
of the respiratory appendage, giving
Fic. 9 ( x 40).
Tracheal extension of pupa.
Fia. 10 ( x 100).
Transverse section of tracheal extension of pupa,
304 Mr. J. A. Dell on the Structure
rise to the clear circular spots previously mentioned.
Except at these places the tracheal extension is separated
from the wall of the respiratory appendage by a space, in
which sections show the presence of a tissue, affected by
ordinary staining re-agents (probably the epithelium which
secreted the tracheal extension), A transverse section
shows scattered sete projecting inwards from the wall of
the tracheal extension (Fig. 10). Traced backwards, the
tracheal extension passes through the space between the
body of the fly and the pupal skin, and finally enters the
body in the fore part of the mesothorax. There are two
large, longitudinal tracheze (Fig. 8) communicating directly
with the tracheal extensions just described. They are
connected by a series of nine commissures, one in the
mesothorax, two in the metathorax, and one in each ab-
dominal segment, except the seventh and eighth. There
is also a series of lateral loops exactly like those of the
larva. The first, however, which arises in the metathorax,
runs through two segments; all the others run from one
segment to the next. The longitudinal trachez end in
tapering extremities in the last segment.
The tracheal system of the pupa is formed around that
of the larva, and consequently in a larva about to pupate
the walls of the tracheze appear double. The longitudinal
trachee of a late larva communicate with the pupal respi-
ratory appendages, which are formed during this stage.
The respiratory appendages are not modelled on any pre-
existing larval organ, hence it is not surprising to find that
the larval trachea, just behind the respiratory appendage,
comes out through the wall of the pupal traches, and
makes its way to the prothoracic spiracle of the larva. It
is also continuous with the base of the respiratory append-
age. This makes a sharp angle with the larval trachea,
and can be seen through the larval cuticle just in front of
the larval prothoracic spiracle. Its distal end turns down-
wards, and almost touches its fellow of the opposite side
in the mid-ventral line. In the pupa which has just shed
the larval skin, the respiratory appendages lie close to the
body, much as they do in the larva; but when the pupa
makes its way to the surface of the mud, the respiratory
appendages stiffen, and stand out clear of the surface.
Pupz can be easily detected in a’sample of sewage, by
their projecting respiratory appendages. A pupa, when
kept in a bottle of water, is unable to rise to the surface,
and Life History of Psychoda sexpunctata. 305
or to breathe the air dissolved in the water, and dies in a
few days. .
The Head and its Appendages in the Fly.
The head is almost vertical. (Figs. 11 and 12.) Seen
from above it is transversely oval ; the front surface is pear-
shaped, narrowing downwards to the labrum. The com-
Fic. 11 ( x 80).
Head of fly.
pound eyes are horse-shoe shaped, and nearly meet above.
They contain comparatively few ommatidia, not more than
five or six rows in the widest part, and possibly about
eighty in all.
Immediately within the lower ends of the horse-shoe-
shaped compound eyes can be seen in the openings of a
pair of chitinous tunnels which perforate the head. They
run through it obliquely, and their openings on the back of
306 Mr. J. A. Dell on the Structure
the head are nearer to one another than the openings on
the face. The tunnels are not of absolutely . uniform
diameter, though their variations are nowhere very great.
The antennary muscles, which are the only important
muscles in the head, are attached close to the hinder
openings of the chitinous tunnels, which seem to act as
struts, and prevent the head from yielding when a strain
Fie. 12.
Ventral view of mouth-parts of fly (to left); maxilla
in side view (to right).
is put upon it by the contraction of the antennary muscles.
Similar structures have been described in Anopheles,
where a pair of straight chitinous tunnels perforate the
head, and in Chironomus, where a pair of small openings
on the face open into a pair of somewhat irregular cham-
bers, which in turn open to the exterior by a pair of slit-like
apertures on the back of the head.*
* Miall and Hammond (1900).
and Life History of Psychoda sexpwnetata. 307
In the Chironomus fly the antennary muscles are
attached to the walls of the chitinous invaginations them-
selves, an interesting point of difference from Psychoda.
A somewhat similar pair of structures has been described
in some other insects, notably in Corydalis.*
Each antenna consists of fifteen joits; of these the
first is cylindrical, the second globular, and the others
flask-shaped, with the exception of the last three, which
are globular. From the globular part of each joint arise
a number of curved, radiating sete.
The fly apparently does not feed at all, since no traces
Hircomian (x20):
Side-view of male fly. The wing is extended and cut short; the legs cut
short. The lower figure shows the genital armature in plan.
of food have been observed in its reduced alimentary
canal, nor has it ever been seen to take in food. The
mouth-parts (Fig. 12) are somewhat similar in arrange-
ment to those of Culex, but are very largely atrophied. As
in most Diptera, there is a roughly conical rostrum, which
bears an elongated, triangular labrum, behind which is
the short, bilobed labium. This is fringed with numerous,
* Waterhouse (1895).
308 Mr. J. A. Dell on the Structure
short, finger-like processes, and its edges are rolled for-
wards so as to enclose a pair of maxilla, The maxille
are short, styliform, and fringed with sete ; near the base
they give rise to a pair of long, four-jointed maxillary
palps. The styliform maxilla probably corresponds to the
lacinia of typical forms; there does not appear to be any-
thing to correspond to cardo and stipes. On the front of
the labium is a minute, triangular hypopharynx or lingua.
The Thorax of the Fly.
As usual in Diptera, the thorax of Psychoda (Figs.
Fic, 14.
Longitudinal section of thorax of Psychoda (upper fig.); ditto of
Chironomus (lower fig.).
13, 14) consists of a small prothorax, an enormous meso-
thorax, and a small metathorax. ‘The prothorax is little
more than a ventral hoop, containing the muscles of the
first pair of legs. Ventrally and laterally its hinder
margin is indicated by a suture. Dorsally its boundary
is uncertain ; it certainly, however, lies in front of the
first spiracle,
and Life History of Psychoda sexpwnetata. 309
The relations of the component parts of the mesothorax
are peculiar, as will be seen by comparison with the cor-
responding structures in Chironomus (Fig. 14). In that
type the dorsal region of the mesothorax consists of a
large shield-shaped piece, the scutum; a transverse, semi-
cylindrical ridge, arching across the back, the scutellum ;
and behind this a large postscutellum, to which the
hinder ends of the longitudinal mesothoracic muscles are
attached.*
In Psychoda the same parts are found with the following
points of difference -—
(1) The scutum extends forwards above the head, so
that the head and prothorax are depressed to the ventral
side, and the face looks downwards.
(2) In Chironomus the post-scutellum has extended
backwards so far as to obliterate the dorsal part of the
metathorax. In Psychoda the post-scutellum extends back
quite as far as in Chironomus, but underlies the metathorax
and the dorsal part of the first and second abdominal
segments. It thus appears in longitudinal section as an
enormous dorsal invagination of the cuticle, running back-
wards and downwards into the body as far as the hinder
end of the second abdominal segment. J am informed
by Mr. T. H. Taylor that a somewhat similar state of
things is found in Stmulium.
The dorsal part of the metathorax is a narrow strip,
arching across the back immediately behind the scutellum.
At the sides of the metathorax are the club-shaped
halteres, or rudimentary second pair of wings, and the
metathoracic spiracles.
The mesothorax is chiefly occupied by the muscles of
flight, which are of extraordinary size, the longitudinal
muscles, for instance, being half as long as the body.
There are two pairs of spiracles. The anterior, which is
mesothoracic,} is formed at the place where the tracheal
extension from the pupal respiratory appendage enters the
imaginal body. Just at this point the cuticle of the fly
thickens into a partial ring, while immediately distal to it
* Miall and Hammond (1900).
+ Voss (1905, p. 739) concludes from his study of Gryllus
domesticus that the thoracic spiracles are probably prothoracic and
mesothoracic. Mial! and Hammond, on the other hand, find that
they are mesothoracic and metathoracic in Chironomus, while Taylor
(1902) comes to the same conclusion in his paper on Simuliwm,
310 Mr. J. A. Dell on the Structure
the tracheal extension is thinner than anywhere else, and
lacks the annular thickenings. This appears to be an
arrangement for snapping the tracheal extension just at
the required spot. Whether the pupal trachez persist as
those of the adult, or whether all those of the thorax are
withdrawn through the adult metathoracic spiracle, I am
unable to say, but in all pupal skins hitherto examined no
trace of trachez could be discovered, while the end of the
tracheal extension always appeared to have been snapped
across just where it originally entered the imaginal thorax.
The metathoracic spiracle is a small, circular opening
immediately below the base of the haltere.
The Abdomen of the Fly and its Appendages (Fig. 13).
The abdomen consists of eight segments. The first of
these is only distinguishable dorsally, being ventrally
obscured by the bases of the metathoracic legs. The
seventh segment shows some indication of being divided
into two parts. The last (eighth) segment in the male is
flattened from above downwards, and slightly bifurcated at
its hinder end, from which arises a long forceps, reaching
forwards when at rest as far as the middle of the seventh
segment. It is composed of two joints, a large basal and
a small terminal one. From the fore end of the eighth
segment arises a second forceps, also two-jointed. This runs
upwards and backwards, and is enclosed by the other.
There are also two processes arising from the hinder end
of the seventh segment in the middle line; one is dorsal
to the other. They are possibly analogous in function to
the penis and titillator of a cockroach,
In conclusion, I desire to express my indebtedness to
Professor Miall, at whose suggestion the work was first
taken up, and without whose assistance and encouragement
it could never have been carried out, and also to Mr. T. H.
Taylor for much helpful advice and criticism.
Libliography of Psychoda.
Baliani, #. G. (1881.) Sur la structure du noyau des
cellules salivaires chez les Jarves de Chironomus,
Zool. Anz. pp. 687-641 and 662-666, with figs. in
text.
(1890.) Etudes anatomiques et histologiques sur
le tube digestif des Cryptops. Arch. Zool, Expér.,
2° ser., VIII, pp. 1-82, Pl. I-VI,
and Life History of Psychoda sexpunctata. 311
- Bouché, P. F. (1834.) Naturgeschichte der Insecten,
Psychoda phalenoides, Meig. Pl. II, fig. 20-23.
Curtis, J. (1823-40.) British Entomology, XVI, p. 745.
Haton, fev. A. H. (1895.) Supplementary notes on Dr.
Fritz Miiller’s paper on a new form of larvae of
Psychodide from Brazil. Trans. Ent. Soc. Lond.,
Part IV, p. 489.
De Meyere, J. C. H. (1902.) Ueber die Prothorakal-
stigmen der Dipteren-puppen. Zool. Jahrb., XV,
pp. 623-692, Pl. XX XII-XXXV.
Leewwenhoek, A. (1692.) Figures the wing and antenna
of Psychoda. Arcana Nature Detecta, Ed. Leyden,
1722, vol. II, 283, and the opposite plate (un-
numbered), figs. 2, 3, 4.
Miall, L. C., and A. L. Hammond. (1892.) The develop-
meut of the head of the Imago of Chironomus. Linn.
Trans., Zool., 2nd ser., V, pp. 265-279, Pl. XX VIII-
XXXI.
(1900.) The Structure and Life-history of the
Harlequin fly.
Mall, L. C., and. N. Walker. (1895.) Life-history of
Pericoma canescens (Psychodide). Trans. Ent. Soc.
Lond., pp. 141-153, Pl. III, IV.
Miller, Dr. Fritz. (1895.) Contribution towards the
history of a new form of larvae of Psychodide from
Brazil. Trans. Ent. Soc. Lond., p. 479-481, Pl.
x, XI.
Osten Sacken, C. FR. (Baron) von. (1895.) Remarks on
the homologies and differences between the first
stages of Pericoma, Hal., and those of the new
Brazilian species. Trans. Ent. Soc. Lond., p. 483-487.
Taylor, T. H. (1902.) The Tracheal system of Stmuliwm.
Trans. Ent. Soc. Lond., pp. 701-716.
Voss, F. (1904-5.) Ueber den Thorax von Gryllus
domesticus. Zeits. fur wiss. Zool., Bd. LXXVITI.
Waterhouse, C. O. (1895.) Labium and submentum of
certain Mandibulate Insects.
Hy
“i hf re h® : \ 4
% cole ie yd Ie ' a
val abe tg him,
SQLs A) dng om
ah,
ayy
rans | hae he
. PNG),
XVIII. New African Lasiocampide in the British
Museum. By Cur. AuRIVILLIUS, Hon. F.E.S.,
F.M.Z.S., etc. .
[Read June 7th, 1905.]
PLATE XVI.
In a collection of seventy African Lasiocampide kindly sent
to me for determination by Sir G. F. Hampson, I have
found some new species described in this paper. Three of
the species belong to new genera, of which Haplopacha
and Chondrostegoides are of interest as representing the
aberrant genus Chondrostega in the Ethiopian region.
I have to thank the Entomological Society of London
for the beautiful figures illustrating my papers.
GONOMETA REGIA, Dn. sp.
(Plate XVI, fig. 3.)
¢. Ochreous-brown or yellowish-brown ; patagia, mesonotum,
legs and under-side of abdomen with a distinct violaceous or
purplish tint ; fore-wing ochreous-brown the basal fourth and four
transverse sinuous bands violaceous-brown, the outermost band end-
ing at vein 2, the median space between band 1 and 2 more or less
suffused with violaceous, outer margin darker than the ground-colour
between the bands ; hind-wing reddish-brown, broadly yellowish at
base of inner margin; under-side of both wings ochraceous, vio-
laceous-brown at costal margin, fore-wing behind middle with an
obsolete brownish transverse band.
Expanse 155 mm.
UcanpDa: Hoima (8S. C. Tomkins).
This giant moth is most nearly allied to G. titan, Holl,
which, however, has a much darker ground-colour without
violaceous tint and obsolete transverse bands of fore-wing.
TRANS. ENT. SOC. LOND. 1905.—PART III. (OCT.)
314 Mr. Chr. Aurivillius on New African
GASTROPLAKHIS RUFESCENS, n. sp.
(Plate XVI, fig. 10.)
Q. Head, thorax, legs and under-side of abdomen pale greyish-
brown with a pinkish tint ; abdomen above ochraceous-yellow darker
at base and without black transverse bands ; fore-wing pale greyish-
brown with a pinkish tint, two waved brown transverse lines before
middle and two such lines about middle, an obsolete angulated
brown submarginal line and a black dot at apex of cell, hind
border with long grey or blackish hairs and scales ; hind-wing pale
yellowish broadly suffused with fawn colour at apex and outer
margin ; wings below pale fawn colour, yellowish towards base
without markings.
Expanse 97 mm.
W. Arrica: Asaba.
PSEUDOMETA PATAGIATA, N. sp.
(Plate XVI, fig. 1.)
Dark umber-brown, with the patagia yellow ; fore-wing with a
white dot at end of cell and two yellow patches mottled with
reddish-brown, one at base of hind margin occupying the base of
the areas la, 1b, 2and 3, the other beyond the cell between veins 5
and 6 ; an irregular submarginal series of fuscous spots.
Expanse 37 mm.
Riv. Nicer: Sapele (2. W. Sampson).
This species differs from Ps, viola in the more elongate
hind-wings. Veins 9 and 10 of fore-wing are somewhat
longer than their common stalk.
«¢ ANADIASA CERVINA, 0. sp.
(Plate XVI, fig. 5.)
Fawn colour; abdomen paler, testaceous. ¢. Fore-wing fawn
colour with three fine, simple, fuscous transverse lines, the first sub-
basal nearly erect and very indistinct, the second a little behind
middle, evenly excurved from costa to vein 5, thence incurved to
middle of hind margin, third line submarginal, uneven and more or
less dissolved into spots, a small whitish dot at end of cell ; hind-
wing pale testaceous. @ differs by the fore-wing being darker and
Lasiocampide in the British Museum. 315
suffused with grey at hind margin and outer margin and by the hind-
wing being nearly as dark as fore-wing.
Expanse 33-35 mm.
NataL: Brit. Musewm; NYASSALAND and GERMAN
East AFrrica: Musewm Holmiez.
Nearly allied to A. (“Philudoria”) distincta, Dist., but
differing in the simple transverse lines of fore-wing and
by the median line not being strongly incurved at costal
margin.
TARAGAMA CAPENSIS, 0. sp.
(Plate XVI, fig. 11.)
Abdomen, under-side of thorax and legs greyish-brown ; palpi
yellowish ; head and upper-side of thorax whitish, the latter with
pale brown patches ; fore-wing fuscous-brown with two oblique,
very irregularly sinuated, white transverse lines, inner line from
middle of costal margin to hind angle of cell and thence following
the median vein to middle of cell and ending at hind margin near
base, outer line from costal margin to vein 3 nearly parallel to outer
margin but forming two distinct projections on inner-side at veins
5 and 8, from vein 3 running obliquely to middle of hind margin
and forming two lunules, a spot at base in 1b, a spot at end of cell
and a submarginal series of spots yellowish ; hind-wing fuscous with
transverse white line and a vellow spot at anal angle ; wings below
fuscous with an irregular transverse submarginal white line and one
or two yellowish spots near anal angle.
Expanse 45 mm.
Cape Cotony : Brak Kloof, April (White).
Allied to Targama cristata but smaller and also differing
in the yellow spots and the darker ground-colour of the
wings.
—TARAGAMA DISTINGUENDA, 0. sp.
(Plate XVI, fig. 13.)
¢. Body brown, darker on upper-side ; patagia deep castaneous
brown ; fore-wing dark castaneous brown paler towards outer margin,
an obsolete blackish dot at end of cell and beyond it two transverse
blackish-brown lines, the first arched and bent basally at costal margin,
the second nearly straight running from costal margin just before
apex to last third of hind margin ; the usual hind marginal area
316 Mr. Chr. Aurivillius on New African
lanceolate slightly curved on fore-side, and terminated by a greyish
line ; very strongly suffused with blackish except at hind margin ;
hind-wing and under-side of both wings dark brown with pale veins.
Expanse 44 mm.
9. Body fawn colour with the patagia deep castaneous ; fore-
wing castaneous, paler behind middle, a very strongly curved trans-
verse median line and another nearly straight oblique line from
costal margin just before apex to last third of hind margin, an
obsolete angulate submarginal line, hind marginal area lanceolate,
fawn colour without silvery streak at front side; hind-wing and
under-side of both wings fawn colour.
Expanse 63-65 mm.
Natau: Durban, October (G. F. Leiyh). A bred male
and female. Also a female from Manau, Nyassaland, in
Museum Holmie.
Very nearly allied to 7. grabert, Dewitz, and perhaps
only a southern form of that species. The female of
grabert has the body greyish, the fore-wing fuscous, greyish
at termen with the second transverse line running exactly
to apex and bent at vein 8; hind-wing and under-side
greyish, not fawn-coloured.
T. cuneata, Dist., has the same ground-colour, but is
easily distinguished by the cuneate form and the silvery
line at front-side of the hind marginal area of fore-wing.
The male has a greater resemblance to the male of
T. sjostedti, Auriv.
(Larva dark brown with numerous pale reticulate striz ;
the thoracic somites dorsally yellow in front, the abdominal
somites dorsally yellow’behind and with ill-defined dia-
mond-shaped dorsal marks; two fine rather irregular sub-
dorsal yellowish lines; the hair on dorsum black, the
lateral tufts white.—G. F. H.)
TARAGAMA FUSCA, n. sp.
(Plate XVI, fig. 8.)
¢. Dark blackish grey; patagia blackish brown; fore-wing
blackish, along hind margin with a lanceolate dark greyish patch,
which on its fore-side near base has a silvery angulated line, behind
middle a slightly curved, nearly parallel 3 mm. broad, blackish
band, an obsolete irregular curved angulated blackish submarginal
line ; hind-wing and under-side of both wings dark blackish grey
without markings.
Expanse ¢ 33, 2 50 mm.
Lasiocampide in the British Musewm. 317
Brit. EK. Arrica: Ndimu (C. 8S. Betton); Kikuyu
(Crawshay) ; Uganda, Kampala (Christy).
(2. Fore-wing more uniform in colour, the patch on
inner area paler.—G. F. H.)
The darkest species of Zaragama known to me.
OCINAROPSIS, 0. sp.
Eyes hairy. Palpi rather long, porrect, reaching beyond
the frons; last joint short, pointed. Antenne nearly alike
in both sexes, bipectinate to the apex; branches much
longer before middle, suddenly shorter behind middle, hind
row of branches at base much longer than the front row.
Legs hairy; tarsi scaled; front tibie unarmed; spurs of
hind tibiz wanting. ore-wing triangular, rather short ;
outer margin angled at vein 5; vein 2 from near base, 3
nearly from middle of cell, 4 and 5 from angle, 6, 7 and 8
stalked, 9 and 10 on a very long stalk, 9 running to the
outer margin well below apex. Hind-wing: costal margin
nearly straight, lobed at base; outer margin distinctly waved
with a longer tooth at vein 2; vein 2 from behind middle of
cell, 3 from before angle of cell, 4 and 5 on a short stalk ;
cell obliquely closed, vein 8 anastomasing with 7 near its
base and thus forming a small precostal cell, which has no
accessory veinlets.
Ocinaropsis obscura, 3 4.
- OCINAROPSIS OBSCURA, n. sp.
Dark blackish grey (9), or blackish brown (¢), thorax and fore-
wing with a reddish tint in the male ; fore-wing with very minute
white point at end of cell and a double submarginal row of obsolescent
black and yellowish minute dots ; hind-wing paler, blackish at anal
TRANS. ENT. SOC. LOND. 1905.—PART In. (ocT.) 22
318 Mr. Chr. Aurivillius on Mew African
angle ; terminal tooth of both wings much longer in male than in
female.
Expanse ¢ 27, 9 33 mm.
Nata: Durban (Leigh).
PHILOTHERMA SORDIDA, 0. sp.
(Pl, XViwis.<2.)
4. Very pale fawn colour or isabelline ; fore-wing pale fawn
colour, darker at base and behind discal line, whitish between dis-
cocellular spot and discal line, a nearly erect slightly waved blackish
line before middle and behind middle, a distinct oblique nearly
straight black discal line, which at vein 9 is suddenly bent towards
costal margin, a submarginal series of small black dots between the
nervules, a white, black-ringed spot at end of cell; hind-wing pale
fawn at costal margin, whitish towards inner margin, a fine discal
black line from costa to vein 6 and a submarginal series of small
black dots; under-side of both wings whitish isabelline with a
submarginal series of black dots on hind-wing, discal transverse line
wanting or obsolete, brownish.
Expanse 62-70 mm.
9. Differs from the male in both wings being more or less densely
sprinkled with blackish scales, especially at outer margin and
behind discal line ; transverse lines more distinct ; discal white
black-ringed spot of fore-wing wanting ; tarsi black.
Expanse 96 mm.
SrerRA LEONE: (Brit. Mus. and Staudinger’s Collection);
NIGER: Onitsba (Brit. Mus.).
Nearly allied to Ph. jacchus, Méschl., and rosa, Druce,
differing from the latter in the paler ground colour and
the straight discal line of fore-wing.
LECHRIOLEPIS PULCHRA, 0. sp.
(Plate XVI, fig. 12.)
Ochraceous yellow ; head, thorax and fore-coxz densely clothed
with long reddish brown hairs; antenne at tip with blackish
branches ; fore-wing yellow (nearly lemon yellow), base, a transverse
line before middle and another behind middle, and a submarginal
irregular row of eight large spots reddish-brown, the discal area
between the transverse lines except in cell and at costal margin
largely suffused with reddish-brown, a reddish-brown spot at end of
Lasiocampide in the British Musewm. 319
cell and between the discal line and the submarginal row, another
row of six reddish-brown spots from hind-margin to vein 7; veing
blackish in marginal area; hind-wing yellow with five reddish-
brown spots from vein 2 to 7, becoming larger and more distinct
towards costal margin, veins 4-7 blackish at termen ; wings below
yellow with submarginal row of brownish spots.
Expanse 72 mm.
Brit. CENTRAL AFRICA: Fwambo.
Nearly allied to Z. varia, Walk., but differing in the
reddish suffusion of the fore-wing and probably distinct.
BERALADE, Walk.
The genera Chilena, Walk. (1855), and Lahea, Wallengr.
(1865), are not structurally distinct from Beralade. Sena,
Walk. (1862), is also probably founded on a species of this
genus.
Some of the species of Beralade are very nearly allied,
and may perhaps only be colour-varieties. At present we
know however but little of their variability. The species
of Beralade are found in dry regions, and the larve feed
probably on thorny bushes in the desert.
I give here a key to the African species known to me,
including also the new ones described below.
KEY TO THE AFRICAN SPECIES.
A. Tarsi ringed with black. Fore-wing without silvery
mark at the end of the cell. Fore-wing with (1)-2
oblique dark stripes, one discal, the other submarginal
or marginal.
a. Costa of fore-wing distinctly but
narrowly ochreous yellow. Palpi
orange, black or blackish on outer
side.
*, Body and fore-wing whitish.
a, Stripes of fore-wing macular, com-
posed of small streaks on the
veins, or dentate. Discal stripe
ending at the apex of vein9 . 1. B. perobliqua,Walk.
320 Mr. Chr. Aurivillius on New African
b. Stripes of fore-wing continuous
and even.
1. Discal stripe of fore-wing very
oblique, ending at apex of vein
8; veins of fore-wing white,
distinctly paler than the
ground , : Yang me . 2. B. continua, Auriv.
2. Discal stripe of fore-wing less
oblique, ending at apex of fore-
wing. Veins of fore-wing not
paler than the ground . . 3. B. obliquata, Klug.
* *, Body and wings fuscous, mouse-
colour . ‘ . ; . 4. B. fumosa, Dist.
B. Costa of fore-wing not yellow. Palpi
not, or only pale orange. Discal
stripe oblique.
*, Body and wings milky-white or
greyish. Outer stripe narrow,
submarginal .. . . 5, B.wallengreni, Auriv.
**, Body and wings dark fuscous.
Stripes of fore-wing blackish ;
outer stripe broad marginal,
composed of large spots . 6. B. bettoni, Auriv.
B. Tarsi not ringed, unicolorous.
a. Fore-wing with a white or silvery
mark at end of cell.
*, Fore-wing reddish-fawn colour,
with three oblique darker
stripes. Discocellular white
mark of fore-wing short and
straight. 3 ‘ , . 7. B. prompta, Walk.
**, Fore-wing greyish or cinereous,
with only one oblique, nearly
submarginal stripe. Disco-
cellular mark angulated.
a. Discocellular white mark short,
not produced at hind angle
to the middle of cell; ob-
lique stripe waved and dent-
ate, not bordered with white. 8, B. levenna, Wallengr.
Lasiocanvpide in the British Musewm. 321
b. Discocellular white mark pro-
duced at hind angie towards
the base along the median
vein nearly to middle of cell ;
oblique stripe nearly even,
bordered with white on
inner side,
1. Fore-wing more or less
tinged with fawn ; oblique
stripe curved towards the
middle of hind margin.
Hind-wing without black-
ish patch at analangle . 9. B. donaldsoni, Holl.
2. Fore-wing brownish-grey or
greyish ; oblique stripe
nearly straight. Hind-
wing with a distinct black-
ish patch at analangle . 10. B. marshalli, Auriv.*
B. Fore-wing without white mark at
end of cell. Wings yellowish-
brown, fore-wing with two, hind-
wing with one dark transverse
line . ‘ 5 : : . 11. B. simplex, Auriv.
BERALADE CONTINUA, N. sp.
(Plate XVI, fig. 14.)
Milky-white or greyish-white ; palpi orange, black on outer side ; a
yellow hair-pencil at base of antennz ; antennze whitish with yellow
branches ; fore and middle tibiew yellow on inner side ; tarsi orange
ringed with black ; upper-side of fore-wing whitish (¢) or more
greyish (9), with the costa narrowly edged with yellow, two even
and continuous oblique darker stripes, one discal from the hind
margin before middle to apex of vein 8 (or 9) nearly straight or
slightly curved, the other submarginal more curved and sometimes
obsolete ; the areas between subcostal veins usually suffused with
fuscous especially in the female, a short blackish streak at the root ;
under-side suffused with dark fuscous, at least in anterior part.
Hind-wing on both sides whitish (¢) or fuscous grey (@).
Expanse 32-34 mm.
Macuakos (2. C. Crawshay); Muant (Betton); ATHI-
Y-MAWE (Betton); TAKAUNGA (Thomas); UGANDA (Christy).
* May be a local race of B. Donaldsont.
322 Mr. Chr. Aurivillius on New African
° BERALADE BETTONI, 0. sp.
(Plate XVI, fig. 9.)
Dark fuscous grey, upper-side of thorax somewhat paler; tarsi
ringed with black ; fore-wing fuscous, veins paler, a blackish stripe
in cell, a broad oblique, nearly straight, blackish stripe from apex to
middle of hind margin, a similar marginal stripe ; hind-wing and
under-side of both wings dark fuscous, unicolorous.
Expanse 32 mm.
Brit. East Arrica: Nakuro (C. S. Betton), February.
BERALADE (CHILENA) SIMPLEX, n. sp.
(Plate XVI, fig. 6.)
Bright yellowish brown ; fore-wing with two transverse, slightly
curved, somewhat oblique and nearly parallel blackish brown lines,
a minute brown dot at end of cell, cilia brown ; hind-wing with a
distinct, median, slightly curved, dark brown transverse line ; both
wings beneath with a transverse median brown line.
Expanse 27 mm.
NataL: Durban (Col. Bowker).
Differs from typical Chilena in vein 8 of hind-wing
anastomosing with vein 7 from its origin.
. BoOMBYCOPSIS CONSPERSA, 0. sp.
(Plate XVI, fig. 4.)
d. Yellowish, thorax above darker greyish yellow; fore-wing
greyish flesh-colour suffused with fuscous grey, with two nearly
straight transverse pale lines, one before middle very indistinct, the
other beyond apex of cell distinct and followed on outer side by a broad
fuscous grey transverse diffused band ; a very broken submarginal
fuscous line terminating on inner side the marginal area, which is
densely suffused with olivaceous grey, no dot at end of cell, the cell,
darker than the surrounding parts ; hind-wing pale yellowish with
an obsolescent pale flesh-coloured transverse shade in the middle.
Wings below ochreous yellow, slightly suffused with brownish or
fuscous at costal margins.
Expanse 45 mm.
Brit. Easr Arrica: N’gong (2. C. Crawshay).
Nearly allied to Bombycopsis indecora, Walk., and
b. venosa, Butl., and somewhat intermediate between
Lasiocampide in the British Museum. 323
these two species. B. conspersa is larger than indecora and
has a yellowish and not a grey ground-colour; from venosa
it differs in having the thorax and the fore-wings much
darker and suffused with grey or fuscous; the veins are
not so distinctly yellow as in venosa.
OLYRA RECTILINEATA, 0. sp.
(Plate XVI, fig. 7.)
Q. Dark reddish brown ; abdominal segments above with pale
hind marginal borders ; fore-wing above with two entirely straight
and erect transverse parallel pale yellowish lines, one before the
other behind middle, an irregular submarginal series of short
yellowish erect lines between the veins; hind-wing with a fine
yellowish line from costal margin at least to end of cell (sometimes
obsolete) ; fore-wing below much paler yellowish, reddish brown at
apex and at termen ; hind-wing below reddish brown with two, in
the middle very angulate, transverse, pale yellowish lines.
Expanse 39-40 mm.
GAZALAND: Mt. Chirinda (G. A. K. Marshall), Decem-
ber. MASHONALAND (Dobbie).
Two females, one in Brit. Museum, one in Museum
Holmiz.
HAPLOPACHA, nov. gen.
Palpi entirely hidden by long hairs ; frons with slight corneous pro-
minence. Antenne of male bipectinate to apex ; branches moder-
ately long. Head, thorax and legs clothed with hairlike scales,
Fore-legs slender, unarmed ; middle and hind tibiz with two apical
spurs. Abdomen short not reaching beyond tornus, Cell of both
wings open. Fove-wing short, triangular ; vein 2 from behind middle
of cell, slightly curved, 3 from slightly nearer to 4 than to 3, 4 and
5 from hind angle, 6 and 7 stalked, 8 from the long stalk of 9 and
10, 9running to termen near apex. Hind-wing short, obtuse ; costal
margin nearly straight; vein 2 from middle of cell, 3 nearer to 4, 4
and 5 from the same point, 7 well separated from 8, 8 anastomosing
with cell near base, forming a very narrow pre-costal cell, one
thick, curved accessory veinlet at base.
A very distinct genus, allied to Chondrostega and Chon-~
drostegordes, but differing from both in vein 8 of fore-wing
being emitted from the long stalk of 9 and 10 and vein 8
of hind-wing not anastomosing from base with the cell,
but forming a narrow precostal cell.
324 Mr. Chr. Aurivillius on Mew African
HAPLOPACHA CINEREA, 0. Sp.
¢. Cinereous grey ; fore-wing with two obsolete transverse fuscous
lines or shades, a dark dot at end of cell and a submarginal series of
Haplopacha cinerea, 6 }.
black dots ; hind-wing on both sides and fore-wing below unicolorous
fuscous grey. Antenne with the branches yellowish-brown.
Expanse 26 mm.
RHODESIA: Sebakwe.
A single, rather badly damaged specimen.
CHONDROSTEGOIDES, nov. gen.
Palpi very minute. Hyes small, hairy. Antenne bipectinate to
tip in both sexes, branches rather long in male, shorter in female.
Frons with a conical corneous prominence, short in male, much
longer in female. Fore tibi# normal, unarmed ; middle and hind
tibie without spurs. Cell of both wings open. Fore-wing with
costal and hind margins straight, outer margin regularly curved ;
vein 3 from middle between 2 and 4,6 and 7 stalked, 8 free from
middle between 7 and 9410, 9 and 10 stalked both running to
termen, 11 to apex. Hind-awing with vein 3 from middle between
2 and 4, 8 entirely anastomosing with cell to middle between
base and vein 7, two short accessory veinlets at base.
Nearly allied to Chondrostega (Led.) differing, in the
normal, unarmed tibize and the winged female. Frons
with a single conical prominence only.
CHONDROSTEGOIDES CAPENSIS, 0. Sp.
g. Ashy-grey, fore-wing above, with two white spots at costal
margin near apex, cilia of both wings chequered with whitish ;
Lasiocampide in the British Museum. — 325
under-side of fore-wing fuscous grey, with three white spots at costal
margin near apex separated by blackish streaks ; under-side of hind-
wing white, a spot at base of costa, the cell and the base of the
areas 2-4 brown ; two angulated blackish lines across the middle,
inner line indistinct in the cell.
Expanse 18 mm.
Chondrostegoides capensis, $ 4.
Q?. Unicolorous, dark grey. Hind-wing below with an obsolescent
fuscous median band. Abdomen without terminal tuft.
Expanse 22 mm.
Care CoLony: Deelfontein, October.
The male is in beautiful condition, but the female is
rather rubbed.
326 Explanation of Plate.
EXPLANATION OF PLATE XVI.
1. Pseudometa patagiata.
2. Philotherma sordida.
3. Gonometa regia.
4. Bombycopsis conspersa.
5. Anadiasa cervina.
6. Beralade simplex,
7. Olyra rectilineata.
8. Taragama fusca.
9. Beralade bettoni.
0. Gastroplakexis rufescens.
1!. Taragama capensis,
12. Lechriolepis pulchra.
13. Taragama distinguenda.
i4. Beralade continua.
(SRE)
XIX. Memoir on the Rhynchota collected by Dr. ARTHUR
WILLEY, F-.R.S., chiefly in Birara (New Britain)
and Lifu. By G. W. KirKaxpy, F.E.S.
[Read June 7th, 1905.]
PEATE: XV EE.
ALTHOUGH of an extremely interesting character, the
collection of Rhynchota made by Dr. Willey * is too
small, and too many of the forms described from neigh-
bouring localities are as yet insufficiently known to me,
to render a discussion upon their geographical distribution
profitable. However, as might have been expected, the
Rhynchotal fauna of Lifu partakes very closely of the
nature of that of New Caledonia, while that of Birara is
equally intimately related to the fauna of Papua (or
New Guinea), at the same time all these are closely inter-
related. Throughout I have employed the terms Lifu and
Birara, as it appears expedient to retain the native names
of such islands wherever these are distinctive, except
those which are extensively colonized by Europeans, as,
for example, New Zealand and even New Caledonia.
Birara (also spelt Berrara) was formerly known as New
Britain, and it was under this term that the bulk of Dr.
Willey’s researches have been published. The present
official term, since the acquisition of the group by Germany,
is Neu Pommern (or New Pomerania). The island of
Murua, frequently alluded to in the following pages, is
also known as “ Woodlark” and is termed “ Moiou ” by
Montrouzier.
The chief difficulty in the specific determination of
this fauna as regards the Ahynchota, is its close relation-
ship with that of Papua (or New Guinea), the latter being
as yet very imperfectly known, the latest descriptions
dating in some cases from Guérin-Meneville’s contribution
to the “Voyage of the Coquille,” and Boisduval’s
* This Memoir may be regarded as a supplement to the ‘ Zoolo-
gical Results based on material. . . collected by Dr. Willey,” published
in five volumes by the Cambridge University Press, 1898-1902.
TRANS. ENT. SOC. LOND. 1905.—PART III. (OCT.)
328 Mr. G. W. Kirkaldy’s Memoir on the
volumes on the ‘‘ Voyage of the Astrolabe.” The exami-
nation of large series of Rhynchota from the multitude of
islands comprising the Bismarck, Solomon, New Hebrides,
and Loyalty Archipelagoes would be exceedingly interest-
ing from a distributional point of view. Our principal
knowledge of these faunas is derived from the works of
Montrouzier (or Montrousier as he also terms himself), a
French priest, who resided in New Caledonia jand the
surrounding islands for many years. Dr. Willey has col-
lected some seventy-six species, falling into sixty-five genera.
Of these I have described as new five genera, one sub-
genus, twenty species and one variety, but the principal
interest of his labours lies in the large proportion of
immature forms, the consideration of which. is reserved
for a future paper.
My thanks are due to Mr. W. L. Distant, for kindly
assistance in the determination of several species, and for
help with the proofs in my absence from England; and
to Dr. D. Sharp for allowing me the opportunity of
examining so interesting a collection.
Sub-order HOMOPTERA.
Family CICADID.
Genus Urana, Dist. Ann. Mag. Nat. Hist. (7), xvi, p. 28.
Dr. Willey has collected fourteen specimens, which I
have ventured to divide into two species—though somewhat
doubtingly—neither of which can I refer to Cicada lifuana,
Montrouzier, though one of them is certainly the Ueana
lifuana as identified by Distant.
1. UEANA LIFUANA, Montr.
(2) Cicada lifuana, Montr., 1861, Ann. Soc. Ent. France, 4, 1,
p. 70.
Veana lifuana, Dist. Ann, Mag. Nat. Hist. (7), xvi, p. 29.
Montrouzier says, “ Veines de la premiere moitié des
elytres vert-bleu. Elytres irisées,”. . . . which does not
in the least apply to any of the specimens before me.
Montrouzier records it from Lifu.
Ehynchota collected in Birara and LInfu. 329
2. UEANA HARMONIA, sp. nov.
(Plate XVII, fig. 1.)
(2) Tibicen (?) lifuana, Dist., 1888, P. Z. S., p. 190.
Pale Iuteous, markings on head and pronotum, lateral third of
scutellum, a W-mark on anterior margin of scutellum, abdomen
above at base—pale luteo-ferruginous ; posterior margin of pronotum
pale lutequs. Costa, except lateral margin and basal nervures, pale
luteous; lateral margins of costa very narrowly, basal cell and
apical nervures fuscous. Tegmina immaculate. Ocelli yellow.
¢. Rostrum reaching scarcely beyond base of posterior coxee ;
opercula larger, distinctly reaching well beyond apical margin of
Ist (? 2nd) abdominal segment—genital segment apically narrowed,
acutangular. Metasternal process, roundly and smoothly tubercular.
@. Opercula smaller, not reaching to apical margin of first
segment, Ultimate segment acute.
Long. 234 mm, Expanse of tegmina 61 mm.
Liru (Willey).
3. UEANA POLYMNIA, sp. nov.
(Plate XVII, fig. 2.)
(2) Tubicen (?) lifuana, Dist., 1. ¢.
Darker, the luteous replaced by furruginous on head, pronotum
and ce except posterior margin of, and longitudinal stripe on,
pronoti®n, etc.
d. Rostrum reaching to apex of posterior coxe. Opercula smaller,
not reaching to apical margin of Ist segment. Metasternal tubercle
smaller, not smooth. Genital segment not apically truncate or
slightly rounded.
Q. Opercula as in g.
T. polymnia thus seems to differ—beyond coloration——
by the longer rostrum, the larger opercula (at least in the
male), the differently formed metasternal tubercle, and the
stronger nervures.
While in some specimens the ulnar nervures are quite
contiguous close to the base, they are clearly apart in others.
Distant records “ 7’. lifwana” from New Caledonia.
*%
Genus BzTurRiA, Stal.
Beituria, Stal, 1866, Hem. Afr., IV, p. 9.
Distributed through the Malayan Archipelago and the
South Pacific Ocean.
330 Mr. G. W. Kirkaldy’s Memoir on the
1. B. EXHAUSTA, Guérin.
Cicada exhausta, Guérin, 1838, Voy. Coquille, p. 181.
[Plate 10, fig. 6 (1831). ]
Beturia exhausta, Distant, Orient. ies py ee err a,
fig. 13.
Brrara (Willey); also recorded from New Guinea, Amboina,
Ceram and Buro (Distant); Ternate and Ulasser
(Breddin).
Genus DUNDUBIA.
Dundubia, Amyot and Serville, 1843, Hémipteres, p. 470.
Cosmopsaliria, Stal, Hem. Afr., IV, p. 5.
Distributed throughout Oriental and Australian regions.
DUNDUBIA OBTECTA, Fabr.
Tettigonia obtecta, Fabricius, 1803, Syst. Rhyng., p. 35.
Cosmopsaltria obtecta, Distant, Orient. Cic., p. 67. Pl. 5,
fig. 13.
BirARA and Liru (Willey); I have also seen it from
Ternate, Batchian, Amboina and Aru, and Distant
further records it from Salawati, Halmahera (or Jilolo),
Ceram, and Duke of York Island (in the Bismarck
Archipelago, I presume) ; Obi (Breddin).
Family CERCOPIDA,
Genus CLOVIA.
Clovia, Stil, 1866, Hémipt. Afr., IV, p. 65.
Distributed over the Ethiopian, Oriental and Australian
regions,
CLOVIA BIRARENSIS, sp. nov.
(Plate XVII, fig. 3.)
Closely allied to OC. separata, Walker, but differing in pattern,
which seems constant.
Shining black, whole surface covered with short yellowish pilosity,
more thickly on the vertex. The posterior margin of vertex and a
slightly rounded stripe connecting the intero-apical ocular angles ;
Rhynchota collected in Birara and Lifu. 331
a sub-anterior stripe across pronotum (not reaching lateral margin) ;
a stripe reaching from posterior angle of scutellum across clavus and
Corium, narrowing outwardly and not quite reaching externo lateral
margins of corium; a slightly curved stripe from externo lateral
margins of corium—a little posterior to the stripe last described—
almost to apex of tegmina (roughly at right angles to the clavo-corial
stripe) ; a stripe (broadening posteriorly) from postero lateral angle
of mesopleura, running sublaterally beneath the eyes and round the
antero-lateral margin of the frons, bright yellow. Lateral mar-
gins of posterior third of tegmina exteriorly, and posterior half in-
ternally, brownish hyaline ; abdomen above and connexivum beneath
dark brownish ; wings brownish hyaline. Frons, clypeus, rostrum,
anterior legs and intermediate and posterior tibia, abdomen more
or less, ferruginous. Sterna, coxee, posterior femora, pale yellow.
Spines and tarsi black. Genz, except the yellowish parts, black ;
a sublateral stripe on mesopleura interior to the yellow, and apex of
rostrum black. Head slightly declivous. Frons much swollen,
rostrum reacting to apex of mesocoxee. Ocelli very slightly nearer
to one another than to the eyes. Pronotum and tegmina strongly
and closely punctured, the former somewhat convex, nearly twice
as long as the head. Tegmina rounded laterally, apically sub-
rotundately acute.
¢. Genital plate long.
¢. Valves five or six times as long as the last abdominal segment.
Long. 12-12 mm. Breadth across pronotum 4 mm. ; across
tegmina 52 mm.
Hab. Brrara (Willey).
Var. The scutellum may be either partly or entirely
ferruginous.
CLOVIA DRYAS, sp. nov.
Pale ferruginous. Vertex and pronotum with similar markings
to O. birarensis, but apical margin of vertex (to eyes) also pale
yellowish. Anterior margin of scutellum pallid. Tegmina similarly
marked to C. birarensis except that the clavo-corial stripe practically
reaches the extero-lateral margin, the apical stripe is straighter, and
these marks are clouded exteriorly with dark brownish, instead of
being clear-cut as in C. birarensis. Beneath paler ferruginous, the
yellow stripe in ©. birarensis only faintly indicated here. Anterior
tibiz and all the tarsi blackish. Head with eyes wider than the
pronotum ; head and pronotum not or scarcely convex. Surface of
frons much flatter than in the other species. Pronotum striate-
punctate, very slightly longer than vertex. Ocelli a little closer
332 Mr. G. W. Kirkaldy’s Memoir on the
together than to the eyes. Tegmina more parallel-sided, striate-
punctate.
Long. 10: mm. Width 3 mm.
Hab. Brrara (Willey).
Genus AUFIDUS, Stal.
Aujsidus, Stil, Trans. Ent. Soc. London, 1863, p. 594.
AUFIDUS HYPERION, sp. nov.
(Plate XVII, fig. 4.)
Allied to A. tripars (Walker).
Head, thorax and basal third of tegmina, ventral surface, legs,
abdomen, etc., stramineous. Base of frons proper, extreme apex of
vertex, a small spot enclosing each of the ocelli, eyes, tarsi
apically, exterior margin of anterior tibiz, black. Apical two-thirds
of tegmina (except lateral margin of their basal two-thirds) blackish
fumate. Posterior margin of pronotum widely (except postero-
lateral angle) and pallid parts of tegmina, suffused with sanguineous.
Abdomen above, femora and posterior tibize more or less sanguineous.
Tegmina and pronotum posteriorly yellowish pilose ; head and pro-
notum anteriorly blackish pilose. Frons swollen, medianly flattened
and suleate; rostrum apically black, reaching a little beyond the
base of intermediate coxe.
Long. 103 mm. to apex of tegmina. Breadth 3 mm.
Hab. Brrara (Willey).
The abdomens of the two specimens (1 f 12) are
unfortunately a little distorted by alcohol.
Family TETTIGONIID.
Genus TErricontA, Geoffr.
Tettigonia, Geoffroy, 1762, Histoire abrégée, I, p. 429.
Tettigonides (part), Signoret, Ann. Soc. Ent. France, 1853,
p. 13.
The most satisfactory arrangement of this huge genus
or collection of genera yet made is that by E. D. Ball, but
it only applies, unfortunately, to North American
forms. So little is known of the Asiatic and Australian
Ethynchota collected in Birara and Lifu. 333
species that I have preferred to describe fully, as a
Lettigonia the following interesting. form, which appears
to be so far unknown. Tettigonia in its wide sense is
practically cosmopolitan.
T. WILLEYI, sp. n.
Head, thorax, and abdominal sternites, rostrum and legs, testaceous,
abdominal tergites a little darker. A black elongate spot on each
side of the middle line of the head, at the junction of frons and
vertex almost meeting in the middle and extending laterally as far
as the antennal ledges ; also a black spot immediately posterior to
each ocellus reaching from thence almost to the base of the vertex.
Apical margin of pronotum medianly black, also a number of
blackish-brown spots on the anterior quarter of pronotum ; scutellum
with a sublateral longitudinal black stripe on each side. Tegmina
dark purplish brown, interior fourth of membrane hyaline, more or
less fumate. Wings purplish brown, fumate. Anterior intermediate
tarsi more or less fuscous, anterior tibial bristles black. Abdomen
above medianly blackish. Head triangular, sub-convex, apically
acutangular ; vertex and pronotum forming an obtuse angle at their
junction. Seen laterally, the head from base of eye to apex of vertex
is distinctly longer than the pronotum; the antennal socket is
protected by a somewhat strong ledge, part of which is distinctly
visible from above, interrupting the curve of the head. Ocelli large,
about twice as far apart from one another as from the nearest eye.
Base of vertex roundly emarginate, carinately margined through its
entire breadth. Frons slightly swollen, the surface flattened, some-
what obsoletely granulate. Anterior tibia not (or exceedingly
slightly) dilated, not sulcate. First segment of posterior tarsi longer
than the other two together. Head and eyes scarcely so wide as the
base of pronotum. Vertex between the eyes two-thirds wider than
the eyes together. Pronotum quadrilateral, anterior margin rounded,
posterior margin slightly obtusangularly emarginate. Pronotum
obsoletely impressed transversely at its anterior third ; strong and
finely punctured (on posterior two-thirds at least), Elytra extending
far beyond apex of abdomen, apically rounded.
¢. Male plates much longer than the ultimate abdominal segment,
apically angular, the extero-lateral margins rotundate basally,
obliquely subtruncate apically, provided with short bristles basally
near the lateral margins. Pygofers dark, more than twice as long as
the plates, bristly.
?. Ultimate abdominal segment bisinuate apically, valves about
four times as long as the preceding, bristly.
TRANS. ENT. SOC. LOND. 1905.—PART Ill. (OCT.) 23
334 Mr. G. W. Kirkaldy’s Memoir on the
Long 8°4 mm. to apex of abdomen, 9°8 to apex of tegmina, width
across pronotum 2°3 mm.
Hab. Brrara (Willey).
THALATTOSCOPUS, gen. nov.
(Plate XVII, fig. 5.)
Very similar to Bythoscopus, Germ. (=Macropsis, Auctt.), but
frons, clypeus and gene anastomosed or only obsoletely sutured off.
Postmargin of eyes beneath, emarginate. Apical margin of head
widely rounded. Ocelli distant about two and a half times as far
from each other as from the eyes. Vertex very short, seven or eight
times as wide between the eyes as long, about twice as wide as the
eyes together, acutely margined basally. Antennal ledge truncate.
Pronotum and scutellum very distinctly transversely striated,
pronotum at base slightly wider than the eyes, lateral margin
somewhat rounded.
T. DRYAS, sp. nov.
(Plate XVII, fig. 5.) —
Above pale Iuteous, beneath testaceous. Eyes reddish-brown.
Apical margin of vertex widely rounded, rostrum slightly passing
anterior coxe. Anterior and intermediate femora incrassate, posterior
femora a little dilated, tibiz subobliquely flattened, sulcate beneath.
First segment of posterior tarsi equal to the other two together.
Tegmina reaching well beyond apex of abdomen.
2. Valves four times as long as preceding segment, exteriorly
subrotundate, five-eighths longer than maximum width, each valve
apically subacute, covered sparsely with short, pale luteous, bristles.
Long. 10} mm. to apex of tegmina, 9 to apex of abdomen, width
3} mm. across base of pronotum.
Hab, BIRARA (Willey).
Family FULGORID&.
Genus PHYLLYPHANTA, Am. Serv.
Phyllyphanta, Amyot and Serville, 1843, Hémipteres, p.
528. Melichar, 1902, Ann. Hofmus. Wien, XVII,
p. 54.
Cromna, Walker, 1857, Journ. Linn. Soc. Lond., I, p. 85.
Distributed over the Southern Oriental Region and
Malayan Isles.
Ehynchota collected in Birara and Lifu. 335
P. BIRARA, sp. nov.
Allied to P. producta (Spinola), but smaller and differs as
follows :—
Apparently immaculate, Frons distinctly carinate along the
middle. Pronotum and scutellum distinctly 3-carinate. Tegmina
narrower, intero-apical angle somewhat obtuse—not strongly acute as
in P. producta (Spin.) and P. dubia (Kirby).
Preserved in alcohol and not in good condition.
Long. 7 mm. (exclusive tegmina); 114 mm, (with tegmina) ;
expanse about 234 mm.
Hab. BrrARA (Willey).
Genus EuricantA, Melichar.
Huricania, Melichar, 1899, Ann. Hof. Wien, XIII, p. 255.
Distributed over the Oriental region up to Japan and
over the Pacific Islands.
EK. SPLENDIDA, Fabr.
Flata splendida, Fabr., 1803, Syst. Rhyng. p. 50.
Fiicania splendida, Guérin, 1838, Voy. Coquille, p. 191,
[Pl. 10, fig. 10 (1831)].
Huricania splendida, Melichar, 1899, Ann. Hofmus. XIII,
p- 262 (PI. 10, fig. 8).
Hab. BrrARA (Willey), two examples without the white
tegminal spots; also recorded from New Guinea, Dorei,
Triton Bay, Kimgunan, Meisol, Key and Sula (Jelichar).
Genus TARUNDIA, Stal.
Tarundia, Stal, 1859, Berlin Ent. Zeit. II], p. 325;
Melichar, 1899, Ann. Hofmus. Wien, XIII, p. 265.
Distributed over South Africa, the smaller Mascarene
Islands, New Guinea, the Solomons Islands.
T. GLAUCESCENS, Melichar.
Tarundia glaucescens, Melichar, 1899, Ann. Hofm. Wien,
XIII, p. 267.
Hab. Brrara (Willey), also recorded from New Guinea.
Hagar, gen. nov.
Allied to Tarundia, Stal, and Pochazina, Melichar, but dis-
tinguished by the basal cell emitting four nerves and by a different
disposition of basal nerves.
336 Mr. G. W. Kirkaldy’s Memowr on the
Frons not narrower than long, lateral margins of clypeus not
carinate, laterally rounded, subperpendicular. Tegmina wide tri-
lateral. The two claval nerves uniting at the middle of the clavus ;
six or seven tranverse nerves in clavus ; great claval nerve uniting
with the claval margin at the apex of the clavus. Radial and sub-
radial veins springing at the same point from the basal cell, both
running subparallel as far as stigma. Costal membrane much wider
than costal area, slightly arched exteriorly, several of the costal
nervures furcate. Longitudinal nerves in corium somewhat far
apart, few transverse nervures. Those always simple. Interior
ulnar nerve arising out of the basal cell at the opposite apical angle
to the exterior ulnar nerve.
HAJAR FORNICATUS, sp. nov.
Pale luteo-testaceous (greenish ?), Abdomen with some black
spots dorsally ; a few of the transverse corial nerves fumate. Tegmina
a little longer than wide, apically rounded. Vertex a little produced,
truncate apically, lateral margins carinate, at right angles to apical
margin, posterior margin excavated, pronotum short, but strongly
carinate medianly longitudinally. Scutellum tricarinate the lateral
keels meeting the middle keel apically at an acute angle. Frons
obscurely carinate longitudinally, strongly carinate laterally and
apically. Posterior tibiae with two spines.
Long. 5°2 mm. to apex abdomen ; expanse tegmina 18 mm.
Hab. Brrara (Willey), 1 2 damaged by alcohol.
PEGGIOGA, gen. nov.
Allied to Dictyophara, Germ., but differs by the acutangularly
emarginate bases of head and pronotum, the numerous transverse
nervures in the very narrow costal membrane, and the less reticulate
apical third of tegmina. rons tricarnate, clypews very short,
strongly tricarinate, posterior tibie 3-spinose. Anterior femora
unarmed.
PEGGIOGA FORMOSA, sp. n.
(Plate XVII, fig. 6.)
Stramineous (greenish ?) ; apex of head black, median line of the
various keels of the head; pronotum, scutellum, and elytral nervures,
more or less pale ferruginous. Stigma, intero-apical margin of
tegmina, exterior half of the transverse series of nerves leading from
the stigma, and the nerves in the costal membrane (and the nerves
Rhynchota collected in Birara and Lifu. 337
in the dark apical part aforementioned) blackish-brown or blackish,
Apex of femora and base of posterior tibia very narrowly blackish.
Abdomen above more or less blackish.
Vertex subfiliform, apex subrotundate-truncate, 24 times as
long as width of head including eyes—more than twice as long
as scutellum—slightly compressed medianly, as wide between the
eyes as the length of an eye (as seen from above), one-third wider
than the eyes together. Posterolateral margin of pronotum truncate,
at right angles to the insect’s longitudinal plane ; at its meeting with
the convex lateral margins of scutellum, it abruptly turns forward at
an obtuse angle, the two sides almost meeting at a right angle when
they abruptly turn forward again acuminately ; between the postero-
lateral angle and the apex of pronotum it is medianly and strongly
elevately carinate. The upper lateral margins are roundly sinuate
and do not quite reach the base, the lower lateral margins commence
carinately close to the eyes and reach the posterolateral angle
acuminately, Lateral part of scutellum between lateral margins and
the exterior keels is strongly punctured. Interior (great) claval
nerve meets the commisural vein almost at the same place as it
meets the claval sutural vein, forming a very long narrow cell.
Basal cell elongate gives off three nervures, the radial and subradial
arising at the same spot and proceeding somewhat divergingly to the
transverse veins, a little before which the radial forks; the ulnar
nerve arises at the opposite apical angle of the cell and continues
to the transverse veins, forking a little before them. All the apical
veins are furcate, sometimes reuniting ; all the veins are shortly,
somewhat sparsely, pilose, Rostrum reaches the base of posterior
coxee,
Long. to apex of abdomen 13 mm., to apex tegmina 17 mm.,
expanse tegmina 245 mm.
Hab, Liru (Willey).
Sub-order HETEROPTERA.
Family CAPSID/L.
IMOGEN, gen. nov.
Elongate. Head vertical, shining, polished, smooth, logitudinally
impressed near the base. Eyes shortly pedicillate and very promi-
nent, nearly twice as broad in profile as high, posterior margin
sinuate, Antennze inserted close to the inner angle of the eye, first
segment a trifle shorter than the profile-width of the eye. Second
incrassate towards the base, third segment very slender. Pronotum
338 Mr. G. W. Kirkaldy’s Memoir on the
smooth, shining, and polished, deeply impressed and constricted in
the middle, with a distinct annuliform collar, lateral margins of
anterior lobe rotundate, obtusangular, not reaching laterally as far
as the eyes, subtuberculate. Posterior lobe laterally subcarinate,
straight, widely diverging posteriorly. Scutellum smooth, shining,
anteriorly depressed. Elytra long, subparallel, clavus and corium
coarsely and thickly punctured ; cuneus somewhat deeply excised,
much longer than wide ; membrane with two cells. Femora not
incrassate ; third segment of posterior tarsi longer than the other
two together.
This genus probably belongs to the Capsaria, but has
apparently no near relations.
IMOGEN FORMOSA, sp. nov.
Head above and beneath (except at base of vertex), basal two
segments of antennz, posterior half of scutellum, elytra, posterior
margin of pronotum except medianly, basal two segments of rostrum,
abdomen above and beneath, sterna more or less, femora, ete. black-
ish or blackish-violet. Base of vertex, pronotum, anterior half of
scutellum, propleura, etc., apical two segments of rostrum, yellowish-
orange; anterior tibie, tarsi and third segment of antenne
pallid yellow. Posterior tibize and tarsi black annulated with whitish.
Vertex, between the eyes, about two-sevenths wider than the two eyes
together. Rostrum reaching to about the middle of the mesosternum.
Second segment of antennz three times as long as the first which
is two-fifths longer than the third.
Long. 5 mm. (to apex of elytra6 mm.). Breadth 2 mm.
Hab. Birara (Willey).
Family NAUCORID.
Genus Mononyx, Laporte.
Mononyx, Laporte, 1832-38, Essai, p. 16; Montandon,
1899, Bull. Sci. Bucarest, VIII, nos. 4 and 5, pp. 4 and 5.
Not found in America, north of Mexico; or in the
Palearctic Region.
M. mrxtus, Montandon.
Mononyx mixtus, Montand., op. cit. p. 15.
Rhynchota collected in Birara and Lifu. 339
Hab. BrrARA (Willey); also recorded by Montandon
from Australia, Amboina, and New Guinea.
Family NOTONECTID.
ENITHARES, Spinola.
Enithares, Spin., 1837, Essai, p. 60.
Distributed over Australian, Ethiopian, and Oriental
Regions. Also in Brazil.
ENITHARES, sp. nov.
A new species, scarcely in sufficiently good condition to
characterize, allied to ZL. abbreviata, Kirby.
Hab. Brrara (Willey).
Family REDUVIID Ai.
Genus PoLypbipus, Stal.
Polydidus, Stal, 1858, O. V. A. F., p. 448.
A few species distributed over Oriental, Australian,
and Ethiopian Regions.
POLYDIDUS ARMATISSIMUS, Stal.
Polydidus armatissimus, Stal, 1859, O. V. A. F., p. 376.
Hab, Brrara (Willey); also from India, Ceylon, China,
and the Philippines.
Genus PHysopERuS, Westwood.
Physoderus, Westwood, 18, Journ. Proc. Ent. Soc. Lond.,
1844, p. cxv.
Epirodera, Westwood, 18, Trans. Ent. Soc. Lond., 1847,
p. 247; Leth. and Sev., l.c., III, p. 96.
Madagascar, Philippines, and the Indomalayan Islands.
PHYSODERUS AZRAEL, sp. nov.
Belongs to Stal’s div. a.
340 Mr, G. W. Kirkaldy’s Memoir on the
Head, pronotum, scutellum (except the produced posterior part),
membrane, sterna (in great part), dark purple-brown, almost black,
dull. Scutellum posteriorly, abdomen above, etc., clear luteous.
Coriaceous part of elytra fulvofuscous. Connexivum blackish,
spotted with Iuteous. Abdomen beneath sordid yellowish-brown
with two sublateral (or submedian) subparallel longitudinal lines
formed of blackish-brown spots, the spotted connexivum forming a
third line. Legs pale, sordid yellowish-brown, somewhat obscurely
annulated with dark brown. Antenne pallid yellowish-brown.
Long. 8 mm.
Hab. Brrara (Willey).
Genus DARBANUS, Am. Serv.
Darbanus, Am. Serv., 1843, Hémipt., p. 370.
Huagoras, Leth. Sev., l.c., II, p. 184.
Distributed over the Oriental and Malayan Islands.
D. poLosus, Stal.
Huagoras dolosus (Stil), 1863, Ann, Soc. Ent. France,
p. 28.
Hab. Brrara (Willey); Ké Island (Stal); Aru (Distant).
Genus ONCOCEPHALUS, Klug.
Oncocephalus, Klug, 1830, Symb. Phip., p. 2, almost world-
wide in distribution.
ONCOCEPHALUS AMEN, sp. nov.
¢. Apparently allied to O. assimilis, Reuter, but differing by the
proportions of the antenns, pronotum, etc. Elongate, sordid
testaceous, head with a large black spot on the posterior area, prono-
tum with three double longitudinal lines which are fainter on the
posterior area. Scutellum with a pallid irregular central longitudinal
line. Head, pronotum, and abdomen irregularly marked with black
laterally. Antenne pallid, apex of first and of second segments black,
third and fourth fumate. Apex of the second and third rostral
segments black. Anterior and intermediate legs pallid, anterior
femora banded irregularly with brown beneath, anterior and inter-
mediate tibize with three blackish rings; intermediate femora apically
black. Posterior femora brownish-black, a little clouded with
pallid ; posterior tibize pallid, with two brownish rings near the
base. Elytra brownish testaceous, obscurely irrorated with brownish,
with two large velvety blackish-brown spots, the apical one with
Rhynchota collected in Birara and Lifu. d4]
a whitish wedge anteriorly. Head long and narrowed in profile,
anterior spines not very prominent; anterior area and first segment
of antennze subequal in length, the former distinctly longer than the
posterior area plus the eyes. Second segment of antennae one-third
longer than the first, 2} longer than the third, and a little more than
twice as long as the fourth ; first segment glabrous, second slender.
Eyes beneath strongly convergent, width of eye (as seen from beneath)
about three times (or more) as wide as the intraocular space. Gula
not tuberculate. Rostrum incrassate, first segment reaching to
anterior margin of eye. Eyes very large, occupying almost the entire
width of the head in profile. Anterior lobe of pronotum destitute
of discal spines, anterior angle minutely spinose. Pronotum about
the middle of the lateral margins with a small tubercle, a little
posterior to this somewhat deeply impressed transversely ; the lateral
margins of the anterior area are subparallel, of the posterior area
widely divergent. Posterolateral angles acute, subvertical, not ex-
tending anteriorly beyond apical margin of posternum, Scutellum
distinctly recurved posteriorly. Anterior femora, with a single row
of eleven to twelve small spines inferiorly, trochanters with two or
three small spines. Posterior tibia slightly pilulose, distinctly
longer than femora.
Long. 15 mm.
Hab, Brana (Willey).
LUTEVA, sp. nov.
Genus LutTEvA, Dohrn.
Luteva, Dohrn., Linn. Ent., XIV, pp. 218 and 242.
Two or three specimens in indifferent condition from
Lifu, probably referable to a new species.
Family GERRID.
Genus GERRIS, Fabricius.
Gerris, Fabr., 1794, Ent. Syst., IV, p. 188.
Limnometra, Mayr., 1866, Novara Reise, Hem., p. 174.
Cosmopolitan; the subgenus Limnometra occurs in
every region except the Palzearctic.
GERRIS, sp. (?).
Hab. Brrara (Willey). |
342° Mr. G. W. Kirkaldy’s Memoir on the
Family NEPID&.
Genus RANAtTRA, Fabricius.
Ranatra, Fabr., Syst. Rhyng, p. 108.
Cercotmetus, Amyot and Serville, Hist. Hém., p. 441.
Practically cosmopolitan.
RANATRA PARMATA, Mayr.
anatra (Cercotmetus) parmata, Mayr., Novara Reise, p.
192° Hab, Vot. 60.
Hab. Java, Tspopas (Willey).
Family ARADIDA.
Genus BraAcHyRuyncuus, Lap.
Brachyrhynchus, Laporte, Essai, p. 54.
Almost world-wide in distribution.
BRACHYRHYNCHUS TAGALICUS, Stal.
Arietus tagalicus, Stal., 1870, O. V. A. P.; p. 672.
Hab. BrrARA (Willey), also recorded from Philippines,
Burma, Java and Nias Island.
BRACHYRHYNCHUS MEMBRANACEUS, Fabr., var.
BRACHYRHYNCHUS ORIENTALIS, Laporte.
Aradus membranaceus, Fabr., 1803, Syst. Rhyng, p. 118.
Var. Brachyrhynchus orientalis, Lap.
Hab. Brrara (Willey); also recorded from South
Oriental region, New Guinea, Vanikoro, ete.
Genus NrEuROcTENUS, Fieber.
Neuroctenus, Fieber, 1861, Europ. Hem., p. 34.
Similar distribution to Brachyrhynchus, except the
Palearctic Region.
NEUROCTENUS, sp. (2).
Two specimens in indifferent condition (Wiley)
Ethynchota collected in Birara and Lifu. 343
Family PYRRHOCORIDA.
(= Pyrrhocoride + Lygacidx, Leth. and Sev.)
Genus PHYSOPELTA.
Physopelta, Ann. Serv., 1843, Hémiptéres, p. 271.
Tphita, Stal, 1870, Sv., Akad. Handl., 9, No. 1, p. 99.
Distributed through the Oriental, Ethiopian, and
Australian Regions.
PHYSOPELTA FIMBRIATA, Stal.
Physopelta fimbriata, Stal, 1863, Berl. Ent. Zeit., VII, p.
392.
Hab. Brrara (Willey) ; previously recorded from Timor.
P. FAMELICA, Stal.
Physopelta famelica, Stal, 1862, Berl. Ent. Zeit., VI, p. 391.
(?) Lygaeus woodlarkianus, Montr., 1855, l.c., p. 105.
Hab. Brrara (Willey); recorded also from Australia,
New Guinea, Ceram, ete., scarcely distinguishable from
P. gutta, Burm.
DYNAMENAIS, gen. nov.
Allied to Aschines, but distinguished by the glabrous
abdomen and prominent eyes; also allied to Lctatops, Bo.,
distinguished by the form of the head, the much more
prominent bucculze and the glabrous abdomen.
Head almost vertical in the front of the eyes, transversely im-
pressed behind them but not narrowed ; eyes large and very promi-
nent, shortly stylate, projecting above the surface of the vertex, as
seen in profile; base of eyes touching or slightly overlapping
pronotum. The base of the head between the eyes is also sulculate,
slightly obtuse angulately emarginate. Pronotum, with a well-
marked collar, divided from the anterior area by a punctured deeply
impressed line, anterior area separate from posterior area by a very
deeply impressed line, also punctured. Lateral margins and prono-
tum subvertical. The collar and the lateral margins are smooth and
polished, the rest of the pronotum is dull and sparsely punctured,
except the posterior margin which is polished and sparsely punctured.
First segment of antennz longer than second and third together
and longer than fourth. Buccule very prominent, acute. First
344 Mr. G. W. Kirkaldy’s 2Zemotr on the
segment of rostrum reaches to the base of head, or at least not
beyond base of collar. Stink-gland orifices with a short sulcus,
reaching along half the width of the pleuron, but very distinct ;
margins curved, callose. Wings with a hamus. Legs unarmed.
Abdomen glabrous beneath,
DYNAMENAIS VENUSTUS, Walker.
Ectatops venustus, Walk., 1873, Cat. Hem., VI, p. 17.
Dull black, marked with yellowish white. Head, except apex ;
pronotum, except collar, lateral and posterior margins ; scutellum ;
elytra, except costal margin, and the apical margin of corium—the
latter widely—and the apical margin of membrane; antenna, ex-
cept extreme base of first segment, and basal two-thirds of the fourth ;
first segment of rostrum; basal two-thirds of the femora ; apical
abdomen tergites ; sterna and pleura, the collar and the posterior
margin of pro- and metapleura excepted ; ambulacra ; basal half of
abdominal sternites in the middle and one or more of the other
abdominal sternites entirely ; black more or less dull, except the
more or less shining abdominal segments, the above exceptions are
all more or less yellowish-white except the sanguineous abdominal
tergites. Second, third and fourth rostral segments castaneous-
brown ; tibize and tarsi pallid or brownish. The prominent parts
of the buccule are ivory-white. Head dull, somewhat obscurely
rugulose, first segment of antennze a little more than twice as
long as the second, and one-sixth longer than the fourth ; second
one-third longer than third. First rostral segment a trifle longer than
the second ; the fourth reaching a little beyond the posterior coxa,
Lateral margins of pronotum sinuate; posterior margins of pro- and
metapleura, and the ambulacra smooth, polished, and punctured.
Elytra, exteriorly at least, punctured. Anterior and intermediate
femora somewhat incrassate. First segment of each tarsus longer than
the other two segments together ; third longer than the second.
Long. ¢ 7mm., 2? 9 mm.
Hab. Brrara (Willey); New Guinea (Walker).
Genus DinDYMUS.
Dindymus, Stal, 1861, O. V. A. F., p. 196.
Distributed throughout Oriental, Ethiopian and Austra-
lian Regions.
DINDYMUS FAMOSUS, Distant.
Dindymus fanosus, Dist., Trans. Ent. Soc. Lond., 1901,
p- 589.
Hab. Brrara (Willey).
Rhynchota collected in Birara and Lifu. 345
DINDYMUS PYROCHROA, Boisduval.
Dysdercus pyrochroa, Boisduval, 1835, Astrolabe Ent. IH, p.
642, Pl. XI, fig. 9.
Hab. Brrara (Willey), Aru, Moluccas, New Guinea,
Myso, ete.
Genus DysDERCUS, Serville.
Dysdercus, Serville, 1835, in Boisduval, Astrolabe, I, p. 64.
Laporte fixed the type of the genus Astemma in 1838,
“ Kssai,” p. 38.
“Le genre Astemma a été établi sur la Salda pallicornis
de Fab.”
Astemma, Kirk., olim.
DyspERCcUS sID&, Montrouzier.
Dysdecus (sic !) sidx, Montr. and Sign., 1861, l.c., p. 68.
In the Journal Bombay Nat. Hist. Soc, I lumped
together a number of forms under the name “ cingulatus.”
Dr. Breddin has since kindly sent me specimens of cingu-
latus and poecilus, and has explained the difference in the
Wiener. Ent. Zeit.
I now find among Dr, Willey’s captures what I believe
to be the true side. D. cingulatus is distinguished by the
shining, polished, immaculate head (except the base
behind the eyes which is generally very narrowly blackish),
while poecilus and sidx have dull heads. In both of the
latter the base of the head beyond the eyes is blackish,
though generally more largely in sidz. While in poecilus
(as in cingulatus), the tylus is red, in sidx it is blackish, the
black colour often encroaching well over the juga, etc. ;
the rostrum is also entirely black (except sometimes the
extreme apex of the first segment) in sidx, while in the
other two, at least the first segment is red. The red
colour in sidz is also darker and browner. These characters
seem constant, other colour characters are however un-
reliable in cingulatus and poecilus, particularly that of the
levigate anterior transverse band on the pronotum, Sida
is perhaps also more strongly punctured.
Dr. Willey has taken A. side at Lifu, the original
habitat noted by Montrouzier. I have it also in my
collection from New Hebrides, New Caledonia and Loyalty
Islands. Stil records it from North and West Australia.
346 Mr. G. W. Karkaldy’s Memoir on the
DYSDERCUS CINGULATUS, Fabricius.
Cimex cingulatus, Fabr. 1775, Syst. Ent., p. 719.
Dysdercus cingulatus, Kirk., J. Bombay Soe., XIV, p. 301,
pl. A., figs. 11 and 12 (except some of the synonymy).
Hab. Brrara (Willey). Widely distributed over the
Australian and Oriental Regions.
DyYsDERCUS POECILUS, Herr Schiff.
Pyrrhocoris poccilus, Herr Schaff., 1844, Wanz. Ins., VII,
p. 17, fig. 699; Breddin, Wien. Ent. Zeit., XX, p. 84.
Hab. BrrsARA (Willey). A very similar distribution to
that of cingulatus. 5
Genus PTocHIoMERA, Say.
Ptochiomera, Say, 1832, Descr. Heteropt. (New Harmony).
Plociomera, Leth. and Sev., |. c., I, p. 194.
Distributed over America; also Japan, ete.
PTOCHIOMERA KYDIPPE, sp. nov.
Belongs to Stal’s divisions dd. Covered all over, more
thickly on head and pronotum, with silvery hair.
Head, pronotum, scutellum and under-side dull black ; a silvery
posterolateral spot on anterior lobe of pronotum. Posterior margin
of pronotum narrowly fulvescent. Elytra pale yellowish-brown,
clouded and punctured with blackish-brown, Connexivum pallid
brown. Membrane fumate, irregularly marked with pallid. First
and fourth antennal segments, more or less blackish, 3rd and 4th
fulvous. Legs brownish, apical half of femora black. Rostrum
fulvous, first and fourth segments darker. Stink orifices black.
Head and eyes wider than anterior margin of pronotum. Fourth
segment of antennze twice as long as the first, a trifle longer than the
second, which isa quarter longer than the third. Pronotum somewhat
deeply impressed transversely, anterior areathree-fifths longer thanthe
posterior ; lateral margins rounded, lateral margins of posterior area
widely divergent roundly, base slightly sinuately emarginate.
Long. to apex of elytra 5 mm.
Hab. Liru (Willey).
Genus PacHyMERus, Auctt., olim.
Pachymerus, Lep. and Serv., 1895, Encycel., X, p. 322."
Aphanus, Leth. and Sev., |. ¢., I, p. 215 (nec Laporte).
Distributed throughout the Old World.
Rhynchota collected in Birara and Lif. 347
I have used the preoccupied name “ Pachymerus” be-
cause I do not know what the correct name of the genus is.
PACHYMERUS NEREIS, sp. nov.
(Plate XVII, fig. 7.)
Although the fourth segment of the antenne is partly
pallid, this species seems to be a Pachymerus, but I have
seen only 3 $9.
Head, first segment of antennz (except extreme apex), extreme
apex of 2nd segment, the third (except extreme base) and apical
half of the fourth, disk of anterior lobe of pronotum, two spots (one
near apex, one near base) on the explanate lateral margin of pro-
notum, scutellum (exeept a proportionately long Y-mark at posterior
angle), two spots near and at apex of costal area ; abdomen above
entire ventral surface (except rostrum, trochanter, 2 ovipositor,
ambulacra and pleural lateral margin and connexivum), black or
blackish ; the exceptions all pale yellowish-brown except lateral
margins of pleura which are yellowish-white. Stink orifices black.
Ocelli close to the eyes, first segment of rostrum reaching to base of
head, 4th to about middle of intermediate coxes.. Second and fourth
segments of antenne sub-equal, each a trifle more than twice as long
as the first, and a quarter longer than the third. Disk of posterior half
of pronotum, scutellum, elytra, etc., very strongly punctured. Elytra
pale yellowish-brown, strongly and somewhat closely punctured
with brownish-black. Clavus triseriate punctured, also with some
irregular punctures between the two interior rows. Membrane
pale yellowish-white marbled with pale yellowish-brown. Lateral
margins of pronotum rounded, sufficiently widely explanate, more
narrowly posteriorly ; base truncate. Femora unarmed, except a
spine near apex of anterior pair.
?. Long. 6 mm.
Hab. Liru (226a Willey).
Genus OxyYCARENUS, Fieber.
Oxycarenus, Fieber, 1837, Weitenweber’s Beitrage, p. 339.
Distributed throughout the Old World; occurs also in
Peru.
OXYCARENUS ‘LIFUANUS, Sp. nov.
Is intermediate between Stil’s divisions b and bb.
Head, rostrum, antenne, pronotum, scutellum and clavus, black ;
orificial callosities white, orifices black; abdomen above black,
medianly sanguineous. Corium white, with a large black spot in
348 Mr. G. W. Kirkaldy’s Memoir on the
the middle and a black spot at apical angle of corium. Membrane
whitish hyaline with a large brownish-black spot. Legs black,
except posterior tibize which are white except at base and apex.
Ambulacra and posterior margin of metapleura white. Abdomen
beneath sanguineous along some of the sutures covered with stiff
whitish hairs, Head, pronotum, etc., strongly punctured. Clavus
punctured in three parallel rows, and there is also a row of black
punctures dividing the corium from the exocorium. First segment
of antennz scarcely reaching beyond the head, second segment half
longer than the first, a trifle longer than the fourth, which is a trifle
longer than the third. Rostrum reaching well beyond base of
abdomen.
Long. 3} mm.
Hab. Liru (Willey).
This may be Macroplax luctwosis, Montr., but that species
is described as finely granulated, with the clavus whitish
and the membrane brown. It is also a little larger than
0. lifuanus.
Genus Astacops, Boisduval. |
Astacops, Boisduval, 1835, Voy. Astrolabe, IT, p. 637.
Australia and the Malayan Archipelago.
ASTACOPS DoRYCUS, Boisduval.
Astacops dorycus, Boisd., l.c., p. 638, pl. xi, fig. 16.
Hab. Brrara (Willey); also from New Guinea and Mysol.
ASTACOPS, sp. nov.
Hab. Liru ( Willey).
This is most probably a good species, but I have re-
frained from describing it, as it belongs to a difficult group
largely characterized by colour-markings, and there is but
a single specimen before me.
Genus STALAGMOSTETHUS, Stal.
Stalagmostethus, Stal, 1868, Svensk. Vetens. and Akad.
Handl., 7, No. 11, p. 72.
Sub-genus Spilostethus, Stal, op. cit. = Lygaeus, Auctt.
nec Fabr.
Rhynchota collected in Birara and Lifu. 349
S. FAMILIARIS, Fabr., var. HOSPES, Fabr.
Cimex familiaris, Fabricius, 1781, Spec. Ins., I, p. 363.
Var. Lygaeus hospes, Fabr., 1794, Ent. Syst., [V, p. 150.
= I. squalidus, Montrouzier, 1861, A.S. E. F., 4, 1, p. 66.
Hab. Liru (Montrouzier and Willey); Brisbane (my
coll.) ; New Caledonia, Philippines, Malacca, China, India,
Africa, Europe.
Genus ONCOPELTUS, Stil.
Oncopeltus, Stal, 1868, Svensk., Vetensk., Akad. Handl., 7,
No. 11, p. 75.
ONCOPELTUS DISPAR, Walker, var.
(2) Lygaeus bicinctus, Montrouzier, 1861, A. 8. E. F., p. 67.
Lygaeus dispar, Walker, 1872, Cat., V, p. 60.
+ » (Willey, 332 n.); Moluccas, Borneo, etc.
Differs from the type by the pallid (sanguineous 2)
femora, The description of Z. bicinctus from New Cale-
donia agrees with this, but the size (7 mm.) is much too
small.
Genus GEOCORIS WILLEYI, sp. nov.
(Plate XVII, fig. 8.)
Allied to G. flaviceps,
Head, anterior and lateral margins of pronotum, posterior angle
of scutellum, first and apical two-thirds of third segments of
antenne, rostrum, legs, etc., pale yellow-testaceous. Rest of antennz
blackish or blackish-brown. Rest of pronotum, scutellum, elytra,
abdomen, etc., bluish-black. Eyes red. Basal two-thirds of
membrane fumate, apical third hyaline, colourless. Connexivum
flavous, spotted with black. Head straight between antenn and
eyes, shortly triangularly prominent in the middle, urpunctate ;
with the eyes wider than pronotum. Fourth antennal segment
fusiform, a little longer than the third, which is a little longer than
the second and twice as long as the first. 'The pronotum has a single
row of punctures near the anterior margin, not nearly reaching to
the lateral margin, basal two-thirds of pronotum and the entire
scutellum somewhat superficially and irregularly punctured ; lateral
margins of pronotum smooth, straight and sub-parallel. Clavus
TRANS, ENT. SOC. LOND. 1905.—PART III. (OCT.) 24
350 Mr. G. W. Kirkaldy’s Memoir on the
with two rows of parallel punctures, interior half of corium not (or
superficially) punctured, exterior half more strongly punctured.
Long. 5:4 mm. to apex of elytra; breadth across eyes 2-6 mm.
Hab. Brrara (Willey).
Family LYGAEIDA), Kirk.
(= Corrib, Auctt.)
Genus Lrerrocoris, Hahn.
Leptocoris, Hahn., 1833, Wanz. Inst., I, p. 200.
Serinctha, Spinola, 1837, Essai, p. 24:7.
Widely distributed through the Oriental, Australian,
Ethiopian and Nearctie regions,
LEPTOCORIS RUFOMARGINATUS, Fabricius.
Lygacus ruformarginatus, Fabr., 1794, Ent. Syst., IV,
p. 152.
Hab. Brrara (Willey); Puintppines (Stal).
Genus Myopocua, Latreille.
Myodocha, Latr., 1807, Gen. Crust. Ins., LT.
Leptocorisa, Latr., 1825, Fam. Nat., p. 421.
Almost world-wide in distribution except the Palwarctic
region.
MyopocHA BURMEISTERI, Montr.
Leptocorisa burmeistert, Montrouzier, 1865, Ann. Soc. Linn.
Lyon., XI, p.-227,
Hab. Brrara (Willey). Described from New Caledonia ;
“Se trouve & Kanala dans les champs, sur les graminées,
au milieu des herbes ” (Montrouzier).
Probably two species contained in Willey’s captures, but
the species of this genus are not very well characterized.
Genus RiIpTortus, Stal.
Riptorius, Stal, 1859, O. V. A. F., p. 460.
Distributed over the Oriental, Ethiopian and Australian
regions, also Japan,
Lhynchota collected in Birara and Lifu. 351
RIPTORTUS IMPERIALIS, sp. 0.
Allied to R. robustus, Dallas.
Head, beneath, juga, base of vertex, pronotum, scutellum, extreme
base of elytra, first, second and apical two-thirds of 3rd segment of
antenne, rostrum, sterna and pleura (except as below), coxa,
trochanters, and posterior femora, a broad median longitudinal
black band on abdomen below, spots on the apical segments of con-
nexivum above, etc., black or blackish-brown, the posterior femora
beneath very polished. Tylus, base of third antennal segment,
abdomen above, and laterally below, a broad almost continuous, very
smooth, stripe from apex of head to base of metanotum, reddish,
Apical segment of antenne, anterior and intermediate tibia, and all
the tarsi, a somewhat indefinite pale ochreous. Posterior tibia
blackish laterally, reddish above and below. Elytra pale ochreo-
fulvous, membrane yellowish-brown hyaline. Pronotal spines,
acuminate, prominent ; pronotum minutely granulate with black.
Rostrum reaching to nearly base, fine at sternum. Posterior femora
with three or four larger spines and a number of smaller ones,
Fourth antennal longer than the first and second together, second
and third subequal.
Long. 174 mm., lat. 4 mm,
Hab. Brrara (Willey).
(?) RIPTORTUS ATRICORNIS, Stal.
Riptortus atricornis, Stal, 1873, Sv. Vet. Ak Handl., 11,
No. 2, p. 94.
A specimen from Lifu (JVilley) is placed here with
some little diffidence. A. atricornis was described from
Australia and Java.
Genus CLETUS, Stal.
Cletus, Stal, Eugenies Resa, Hem., p. 236.
Peniscomus, Spinola, in Signoret, 1861, Ann. Soc. Ent.
France, 4, 1, p. 66 [nov. syn.].
Distributed over Oriental, Ethiopian, and Australian
regions, also from North America.
CLETUS AMYOTI, Montrouzier.
Gonocerus amyoti, Montr., 1861, l.c., p. 66.
Hab. Liru (Montrouzxier and Willey),
352 Mr, G. W. Kirkaldy’s Memoir on the
Genus LEPTOGLOSSUS, Guérin.
Leptoglossus, Guérin, 1838, Voy. Coquille, p. 174.
Widely distributed over the Ethiopian, Oriental and
Australian regions (including the Pacific Isles), America,
and the Canary Isles.
L. MEMBRANACEUS, Fabricius.
(2?) Cimex australis, Fabr., 1775, Syst. Ent., p. 708.
Cimex membranaceus, Fabr., 1781, Spec. Ins., I, p. 351.
Anisoscelis (Lygaeus) bidentatus, Montr., 1855, 1. ¢., p. 101.
Solomons Islands, Guodaleanar (Willey, June 11, 1897);
also from Woodlark, New Caledonia and San Cristoval,
North Australia, Philippines, Africa, India, Ceylon, Canary
Tsles, etc. Willey’s examples are typical membranaceus ;
L. australis, which is doubtfully distinct, is from Tahiti.
“Vit sur les Cucurhitacées. On le trouve aussi sur certains
acacias ” (Montrouzier).
Genus Mictis, Leach.
Mictis, 1814, Leach, Zool. Misc., L, p92.
Oriental, Australian, and Ethiopian regions.
MICTIS PROFANUS, Fabricius.
Lygaeus profanus, Fabricius, 1808, Syst. Rhyng., p. 211.
Nematopus profanus, Montr., 1855, |.c., p. 102.
Hab, Liru (Wiiley); also from Australia, Woodlark,
New Caledonia, Viti, Samoa, San Cristoval, etc. “ Vit
sur les mimosa” (Montrouzier).
Genus PRIOCNEMICORIS, Costa.
Priocnemicoris, Costa, 1863, Rend. Acad. Napol., I, p. 253.
New Guinea and neighbouring islands.
PRIOCNEMICORIS ALBITHORAX, Boisd., var. PATRICIUS, nov.
I have not been able to examine P. albithorax, Boisd.,
1835 (= flaviceps, Guérin, 1838), and am therefore unable
to differentiate this form specifically. It differs from the
typical form, as figured by Boisduval and Guérin, by the
unicolorous pronotum.
Head, rostrum, entire ventral surface (except the exterolateral
margins of the pleura), connexival tergites, legs (except posterior
Rhynchota collected in Birara and Lifu. 353
femora, ete.) olivaceo-testaceous. Eyes, ocelli, antennae, etc., dark
reddish-brown ; 4th segment of antenne fulvous. Pronotum,
scutellum, exterolateral margins of the pleura, clavus, corium, and
abdomen above metallic shining green. Posterior femora (except
at the base) and base of posterior tibie shining greenish-black.
Membrane irridescent dark olivaceous, tinged with purple. There is
a pale luteous band on the pleura interior to the lateral metallic
margin, and a sublateral pale luteous spot on each abdominal tergite.
Rostrum reaching to the middle of the mesosternum, which is
channelled so far. Fourth segment of antenne about one-half
longer than the first, which is one-sixth longer than the second,
which is one-half longer than the third. Pronotum transversely
striolate, anteriorly obsoletely callose, sulcate medio-longitudinally.
Clavus and corium strongly and finely punctured. Each abdominal
segment posterolaterally minutely spined.
. 6th (?) abdominal tergite very slightly rotundately emarginate ;
posterior femora somewhat incrassate, with one larger, median, and
three to five smaller spines, near the apex.
9. 6th (?) abdominal tergite roundly emarginate; posterior
femora much less emarginate, lacking the median larger spine.
Long. 19 mm., lat. 53-53 mm.
Hab. Brrara (Willey), The typical form is from New
Guinea.
Genus PTERNISTRIA, Stal.
Pternistria, Stal, 1873, Svensk. Vetensk. Akad. Hand], p.
43.
Australia, New Guinea, Birara, and Aru.
PTERNISTRIA, Sp.
Specimens from Birara (Willey) closely agreeing with
the description and figure of macromera (Guérin), which,
however, is unknown to me. The latter has been recorded
from New Guinea and Aru.
Family CIMICID.
Genus MrcyMeEnvuM, Laporte.
Megymenum, Lap., 1833, Essai, p. 52.
Megalymenum, Burmeister, Handb. Ent., I, p. 349 (note),
1835.
Distributed throughout the Australian and Oriental
regions up to Japan.
354 Mr. G. W. Kirkaldy’s Memoir on the
M. AFFINE, Boisduval.
Megymenum affine, Boisd., 1835, Voy. Astrolabe, II, p. 633,
Pl. 11, f. 12; Montrouzier, 1855, 1. ¢., p. 101.
M. montrouziert, Leth. and Sev., 1893, Cat. Hem., I,
p. 239.
These two names appear to belong to the same form.
Stil says of the Montrouzierian, “a JZ. affine, Boisd., vix
differt,” though the former is in his division “cc” and the
latter in “c.” Dr. Willey collected specimens from Birara.
It has also been recorded from Moiou (Montrouzier) and
New Guinea.
Genus AGAPoPHYTA, Laporte.
Agapophyta, Lap., 1833, Essai, p. 63.
The genus contains one species.
AGAPOPHYTA BIPUNCTATA, Boisduval.
A gapophyta bipunctata, Guérin, 1838, Voy. Coquille, p. 168.
[Pl. 11, f. 5 (1831), not f.15, as say Leth. and Sev.]
Hab. Brrara (Willey); also from Australia, Tasmania,
New Guinea, Woodlark Island, Havre Carteret, Buru and
Sarawatte Islands, Moluccas, ete.
“Vit en troupe sur une espece d’acacia & Woodlark ”
(Montrouzier).
Genus CATACANTHUS, Spinola.
Catacanthus, Spinola, 1837, Essai Hémipt., p. 325.
Distributed from India and Ceylon to Toga and from
Australia to Japan.
C. puNctuM, Fabricius.
Cimex nigripes, Subzev., 1776, Gesch. Ins., p. 96, Pl. 10,
fig. 9 (nec Fabr.).
C. punctum, Fabricius, 1787, Mant. Ins., II, p. 291.
C’. carrensi, Le Guillou, 1841, Rev. Zool., p, 262.
Pentatoma tricolor, Montrouzier, 1855, l. c., p. 96.
Catacanthus nigripes, Leth. and Sev., 1898, Cat. I, p. 163.
Hab. Brrara (Willey; typical pwnetum); Woodlark,
Art, Australia, Philippines, Java, ete.
OU
Lthynchota collected in Birara and Lifu. 35:
Genus AGONOSCELIS, Spinola.
A gonoscelis, Spinola, 1837, Essai, p. 327.
Widely distributed over Ethiopian, Oriental and Aus-
tralian Regions.
AGONOSCELIS RUTILUS, Fabricius.
Cimeaw rutilus, Fabricius, 1775, Syst. Ent., p. 714.
Hab, BrraArA (Willey); also recorded from Australia,
Moluccas, Java, Celebes, Batchian, Banda, ete.
Genus PLAUTIA, Stal.
Plautia, Stil, 1864, Hem. Afric., I, p. 191.
Distributed similarly to the foregoing genus.
PLAUTIA BRUNNIPENNIS, Montrouzier.,
Pentatoma brunnipennis, Montr., 1861, |. ¢., p. 63.
Hab. Brrara (Willey); Liru (Stal); New CALEDONIA
(Montrouzier).
- A very variable species. In one specimen the posterior
half of the pronotum is black and there is a large black
spot towards the posterior angle of the scutellum. The
corium may be immaculate or heavily spotted with black.
Venter usually immaculate, but in the example first
mentioned, there are three black spots on each segment.
Differs from P. jimbriatus, Fabr., by the absence of the
black thread-line on the lateral margins of the pronotum ;
scarcely separable from P. affinis, Dallas, from Australia,
except that the pronotum is more coarsely punctured in
the present species.
Genus ANTESTIA, Stil.
Antestia, Stal, 1855, O. V. A. F., p. 54
Similar distribution to the foregoing.
ANTESTIA CHAMBERETI, Le Guillon.
Strachia chambereti, Le Guillon, 1841, Rev. Zool., p. 262.
Hab, BIRARA (Willey) ; also from New Guinea, Wood-
lark, and Waigiu.
356 Mr. G. W. Kirkaldy’s Memoir on the
Genus Stenozyaum, Fieber.
Stenozygum, Fieber, 1861, Europ. Hem., p. 345.
Oriental, Ethiopian and Australian regions, also Soutb-
eastern Palearctic.
J. STENOZYGUM ERYTHRASPIS, Boisduval.
Pentatoma erythraspis, Boisd., 1835, Voy. Astrolabe, pt. 2,
p. 629, Pl. 11, fig. 8:
Hab. BrraRA (Willey); also recorded from Moluccas,
Ceram, Batchian, Halmahera, Carteret, Ké and New
Guinea.
Genus VITELLUS, Stal.
Vitellus, Stal, 1865, Ann. Soc. Ent. France, 4, 5, p. 170.
Distributed over Australia and the Malayan and Pacific
Isles.
VITELLUS PUNGENS, Montrouzier.
Lhyncocoris pungens, Montr., 1861, 1. c., p. 66.
Vitellus pungens, Stal, 1865, 1. c., p. 172.
Hab, Brrara (Willey); one ex. measuring 16 mm. long ;
WoopDLarK (Montrewzier).
Genus ZANGIS, Stal.
Zangis, Stal, 1867, O. V. A. F., p. 514.
Distributed over Australian, Ethiopian and Oriental
Regions.
ZANGIS MONTROUZIERI, Leth, and Sev.
Lhaphygaster sulcatwm, Montr., 1861, 1. ¢., p. 65.
Zangis montrouziert, Leth. and Sev., 1898, |. ¢., p. 170.
Doubtfully distinct from Z. swlcatum (Montr.), Stal.
Hab. BrrRaARA (Willey); New CaLEponta (Stal) ; Liru
(Montrouzier).
Genus Cuspicona, Dallas.
Cuspicona, Dallas, 1851, List., p. 296.
Oriental and Australian regions.
Rhynchota collected in Birara and Lifu. 357
CUSPICONA LAMINATA, Stal.
Pentatoma viride, (p.) Montrouzier, 1855, 1. c., p. 98.
Cuspicona viride, (p.) Montrouzier and Signoret, 1861, l.c.,
p. 65.
C. laminata, Stal, 1876, Svensk. Akad. Handl., 11, No. 2,
p. 102.
Hab. Liru (Willey and Montrouzier); WooDLARK
(Montrouzier), and New CALEDONIA (Séd/).
Genus EURINOME, Stal.
Lurinome, Stal, 1867, O. V. A. F., p. 516.
New Caledonia, Lifu and the Philippines.
EURINOME BASIVENTRIS, Signoret.
Pentatoma inconspicua, Montrouzier, 1858, 1. ¢., p. 249 (nec
Dallas).
P. basiventris, Signoret, 1861, Ann. Soc. Ent. France, p. 63.
Hab. Liru (Willey) ; Balade “commune dans les herbes,
& terre sous les végétaux en décomposition ” (Montrouzer).
Genus CocToTERIS, Stal.
Coctoteris, Stal, 1858, O. V. A. F., p. 435.
Coccoteris, Leth, and Sev., 1893, 1. c., I, p. 100.
New Guinea, Birara and Woodlark.
CocToTERIS EXIGUUS, Distant.
Coctoteris exiguus, Distant, Trans. Ent. Soc. Lond.
A very variable species, the ground-colour ranging from
pale greenish-cinereous to a fine rosy hue.
Hab. Brrara (Willey); New Guinea (Distant).
Genus Eysarcoris, Hahn.
Lysarcoris, Hahn, 1834.
Universally distributed.
EYSARCORIS MEGASPILOIDES, Tryon.
Hab. Birara (Willey); originally described from New
Guinea.
358 Mr. G. W. Kirkaldy’s Memoir on the
Genus ApRISA, Am. Serv.
Adrisa, Amyot and Serville, 1843, Hémipteres, p. 89.
Geobia, Montrouzier, 1858, Ann. Soc, Linn, Lyon., p, 245,
“Vit toujours & terre sous les plantes, les végétaux en
décomposition, et répand une odeur tres fétide” (M.).
ADRISA WILLEYI, sp. nov.
Differs from the other Australo-Polynesian species by its more
elongate and parallel-sided form ; from A. similis, Signoret by the
prosternum being only obsoletely punctured, different ostiolary
structure, also the tylus not at all surpassed by the juga and the
elytra reaching far beyond apex of abdomen; from A. nwmeensis,
Montrouzier by the longer membrane, more feebly punctured
pronotum and different ostiolary structure. Black (somewhat dull),
lateral margin of pronotum very narrowly fulvous, posterolateral
angles yellowish. Elytra strongly punctured with black, clavus
internally and the nervures of the corium, castaneous; exterior one-
third of corium and apical margin of the same narrowly, yellowish.
Membrane yellowish hyaline, mottled with pale brown. Basal two
segments of antenne, rostrum, and anterior tibia, fulvous ; third
segment of rostrum fumate, fourth segment, intermediate and
posterior tibizw and all the tarsi, yellowish ; coxew, femora, entire
ventral surface and tibial spines, black. Head smooth, somewhat
irregularly and obsoletely striolate. Pronotum somewhat feebly
punctured, except on the smooth anteromedian area. Scutellum
similarly punctured, more strongly and regularly at the sides.
Elytra strongly and closely punctured, more strongly so at the base ;
apical margin of corium sinuate. Membrane reaching well beyond
apex of abdomen. Second segment of antenne 3} times as
long as the first, and one-third longer than the third which is
equal to the fourth. Second and third segments of rostrum sub-
equal, each two-fifths longer than fourth. Anterior femora incrassate.
Long. 114-124 mm. (to apex of elytra), lat. 6~6% mm. across widest
part of elytra.
Hab. Liru (Willey, 226a, 212b, 319).
Grotomus, Mulsant and Rey.
Geotomus, M. and R. 1866, Punaises Pentat., p. 84.
Signoret, 1883, Ann, Soc, Ent. France, p. 33.
Almost world-wide.
Rhynchota collected in Birara and Lifu. 359
GEOTOMUS PYGM&US, Dallas.
Cydnus pygmevus, Dallas, 1851; List, p. 129.
Geotomus pygmeus, Signoret, 1883, 1. ¢, p. 51, Pl. 3, fig.
160.
Hab. Lrru (Willey); described under eleven names;
from India to New Caledonia and the Hawaiian
Archipelago,
GEOTOMUS, sp. ?
Probably a new sp. but I have inadequate material
before me,
Genus CHRYSOCORIS.
Chrysocoris, Hahn, 1834, Wanz, Inst., II, p. 38.
Distributed over the Oriental and Australian regions.
CHRYSOCORIS SEXMACULATA, Leach.
Scutellera seamaculata, Leach, 1815, Zool. Misc., II, p. 36,
PRX,
S. arrogans, Montr., 1858, 1. c., 2, V, p. 258.
S. grandis, Montr., Ann. Soc. Ent. France (1861), p. 60
(nec Thunberg).
Chrysocoris (Hucorysus) sexmaculatus, Stal, Svensk. Akad.
Handl., p. 19.
Hab. Liru (290a Willey); also recorded from New
Caledonia, Art, etc.
Genus PHILia, Schiddte.
Philia, Schiddte, 1842, Kroyer’s Nat. Tidsskr., IV, p. 279.
Distributed over Oriental Islands and Australian region.
PHILIA FEMORATA, Walker, var. AUREOCINCTA, Walker.
Callidea aureocincta, Walker, 1867, Cat. Hem. Het., I,
p. 41.
Hab. Brrara (Septr., Willey); the variety is recorded
from New Guinea; the species is also recorded from
Moluccas and Ceram. The placing of awreocincta as a var.
of femorata is on Distant’s authority.
360 Mr. G. W. Kirkaldy’s Memoir on the
PHILIA LEUCOCYANEA, Montrouzier.
Scutellera lewcocyanea, Montr., 1855, |. c¢., p. 95.
Philia leucocyanea, Stal, 1868, Svensk. Vetensk. Akad.
Hand), 7, No: Lit, ae
Differs from Montrouzier’s description by the pronotum
being of the general ground-colour, with the anterior and
posterior margins metallic (green or blue). Femora usually
crimson. Posterior angle of scutellum always blackish.
Hab. Liru and Birara (Willey); also recorded from
Murua, Solomons and Duke of York’s Island. One
example from Lifu has the scutellum largely clouded
with blackish, and several specimens from Birara have a
nebulous blackish median line down the scutellum.
Var. EBENINA, J. Martin.
Philia ebenina, J. Martin, 1898, Bull. Soc. Ent. France, p.226.
P. leucocyanea, var. ebenina, Distant.
Hab. Brrara (Paris Mus, and Willey).
One specimen has four yellowish-brown spots on the
scutellum ; the anterior lateral margins of the pronotum
are also yellowish-brown. It would perhaps be more
correct to state the ground-colour as yellowish-brown, with
blackish-brown markings.
Genus TEctTocoris, Hahn.
Tectocoris, Hahn, 1834, Wanz. Inst., II, p. 33.
This genus contains one species only.
I
TECTOCORIS LINEOLA, Fabricius.
Cimex lineola, Fabr., 1781, Spec. Inst., I, p. 340.
Tectocoris lineola, Leth. and Sev., 1893, Cat. Hem., I, p. 19.
Scutellera banksii, Montrouzier, 1855, Ann. Soc. Afric.
Lyon (2), VII, p. 92, and 1858, Ann. Soc. Linn. Lyon
(2), V, p. 243.
Tectocoris bunksit, Montrouzier, 1861, Ann. Soc. Ent.
France, p. 60.
The synonymy of this variable species is given by
Lethierry and Severin, the variations are described at length
by Montrouzier, Stil (1873, Svensk. Vet. Akand Handl.,
11, No. 2, p. 11), and Vollenhoven (1863, Essai Faune Ent.
Archip. Indo-néerl., I, p. 7). Dr. Willey has collected two
Ehynchota collected in Birara and Lifu. 361
examples, (a) allied to var. tonge, Boisduval; bluish, a
small median spot on the pronotum anteriorly, a spot near
each anterolateral angle of pronotum, anterior margin
prosternum, base of abdomen and incisures of abdominal
sternites (also apparently part of beneath abdominal
tergites) sanguineous. (8) very near typical banksvi,
Montrouzier (which is not var. banksit, Donovan, but is
near diophthalmus, Thunberg). Pale luteous above, testa-
ceous below; lateral margins of tylus, vertex behind the
eyes, two anterior submedian wedge-shaped spots on
pronotum—brownish-black with a suspicion of metallic
green. Connexivum (except extero-lateral margin
narrowly), antennz, a sublateral spot on each pleuron,
apex of femora, the tibiz and tarsi metallic green.
Hab. Liru (No. 319, Willey); extended over the whole
of Malaysia from Java eastward, Australia, Polynesia, ete.
(Woodlark) ; Isle of Pines, New Caledonia, Art, Isles to
the north of New Caledonia, etc., on Hibisews tillaceus
(Montrouzier); “Elle vit & Woodlark en troupes, dépose
un trés-grand nombre d’ceufs en forme de barillet,
@abord blancs, et prenant une teinte de plus en plus
rougéatre & mesure quils approchent du temps de
Véclosion” (Montrouzier).
Genus CANTAO, Am. Serv.
Cantao, Amyot and Serville, 1843, Hist. Nat. Hem., p. 29.
Distributed over the Oriental region up to Formosa and
to Australia and Polynesia.
1. CANTAO VARIABILIS, Montrouzier.
Scutellera variabilis, Montr., 1855, 1. c., (2) VIL, p. 93 [not
(2) I, as in Leth. and Sev. Cat.]
Cantao variabilis, Stil, 1873, Svensk. Akad. Handl., p. 10,
11, No. 2.
Hab. Brrara (Willey); Moiou (Montrouzier).
This is a very variable species.
Montrouzier in speaking of the Seutellerine of Oceania
(I, p. 91), says: “Les moeurs de ces Scutelléres sont
interessantes. Plusieurs d’elles déposent leurs ceufs sous
le revers des feuilles pour les mettre & V’abas de la
pluie, les coudrent de leur corps, et quand ils sont éclos,
conduisent leurs petits et les protegent jusqu’a ce quwils
362 Mr. G. W. Kirkaldy’s Memoir on the Rhynchota.
aient acquis leurs ailes. On voit leur nombreuse famille
les suivre le long des arbres et se presser vers elles au
moindre danger. Plusieurs aussi sont parées des plus
vives couleurs, et je n’ai encore rencontré chez aucune
Yodeur fétide que laissent échapper les Pentatomes. .. . .
Quant aux lieux ot on les trouve, ils varient selon l’espéce.
Il est des Scutelléres comme ma 8S. metallica (= Philia
senator, Fabr.,) var., qui vivent sur les buissons exposés au
soleil. D’autres, comme ma S. splendida (= Calliphara
billiardiens, Fabr.), choisissent les lheux ombragés ot
croit une espece de myrtacée. La S. banksii (= Tectocoris
lineola, Auctt.) aime beaucoup les endroits ot croissent les
ketmies & feuilles de tilleul.”
Genus CoprosomA, Laporte.
Coptosoma, Lap., 1832-3, Essai, p. 73.
Distributed throughout the Old World.
CoPpTOSOMA SPHARULA, Germar.
Coptosoma spherula, Germ., 1839, Zeitschr., Ent. I, p. 25.
Hab. Brrara (Willey).
Described under thirteen names, from India, Siam, Java,
Ceylon, China, Celebes, Sula, Borneo, Papua, ete.
Genus BRACHYPLATYS, Boisduval.
Brachyplatys, Boisd., 1835, Voy. Astrolade, Ent. I], p. 627.
Platycephala, Lap., 1832-3, Essai, p. 74.
Plataspis, Hope Cat., 1837, I, p. 16.
Similar distribution to Coptosoma except that it does not
enter the Palearctic region.
BRACHYPLATYS PUSILLUS, Tryon.
Brachyplatys pusillus, Tryon, 1892, Ann. Mus., Queensland,
p. 13.
Hab. BrrARA (Willey); NEw GUINEA (Zryon).
Kindly identified by Mr. Distant.
Explanation of Plate. 363
EXPLANATION OF PLATE XVII.
F . Ueana harmonia.
a
ce
» polymnia,
. Clovia birarensis,
. Aufidus hyperion.
. Thalattoscopus dryas.
. Peggioga formosa.
. Pachymerus nereis.
nA DT SB w
. Geocoris willeyt.
OctroBER 4TH, 1905.
XX. The Blind Coleoptera of Australia and Tasmania.
By Arraur M. Lea, F.E.S., Government Ento-
mologist, Tasmania.
[Read October 5th, 1905.|
BLIND Coleoptera in Australia are principally to be taken
under stones, or at the roots of beach-growing plants. In
other parts of the world many species have been taken in
caves, but so far no one has so taken them in Australia,
although they have been specially searched for on several
occasions. I have myself searched for them without
success in caves in Western Australia, New South Wales,
and Tasmania. ‘The total number of blind species
(including one now first described) recorded from Aus-
tralia and Tasmania is but eight (four from Tasmania,
two from New South Wales, and two from Western
Australia, a number probably far short of the total to be
obtained by diligent searching. All the species are of
small size with, as elsewhere, the body apterons and elytra
soldered together.
Following are given notes on all the known species.
CARABIDA.*
ILLAPHANUS STEPHENS!, Macl.
In company with Messrs. H. J. Carter and E. Ferguson
at Otford, my brother (Mr. A. H. T. Lea) at Watson’s Bay
near Sydney and Mr. R. Helms, also at Watson’s Bay, I
have recently (April 1905) taken specimens of this inter-
esting little species. The original locality was Wollongong,
but the late Rev. R. L. King also took the species at
Parramatta and Mr. H. W. Cox has taken it at the Lane
Cove River. It is thus fairly widely distributed, and that
* Steganomma porcatum, Macl., was recorded (P. L. S., N.S.W.,
1904, p. 60) as blind, but it really has eyes, although these are small
and very difficult to find.
TRANS, ENT. SOC. LOND. 1905.—PART IV. (DEC.) 25
366 Mr. A. M. Lea on the Blind Coleoptera
it has not been taken more frequently must be put down
to its small size and to its habit of clinging to the under
surface of deeply buried stones, which in the ordinary
way would seldom be turned over in the search for insects.
We found it necessary in searching for it to kneel on the
ground and closely examine every inch of the surface of
the stones turned over, as the insects, although fairly
active when in motion, are not always ready to move, and
their minute size and dingy colour render them very incon-
spicuous. The stones favoured by them are nearly always
of fairly large size and are always deeply buried in damp
(but not wet) places, and it generally takes two men to
turn them over.
The upper surface is clothed in sparse short pale
pubescence, but this is almost invisible, except from the
sides. The original figure is not a very good one.
Some specimens taken at Watson’s Bay appear to
represent a variety, as they differ from normal specimens
(in the company of which they were taken) in being
decidedly larger (much smaller than macleayi, however),
darker in colour, more parallel-sided, the cephalic impres-
sions less marked and the median line of the prothorax
more distinct.
ILLAPHANUS MACLEAYI, n. sp.
Pale testaceous, appendages still paler. Clothed with short and
very sparse pubescence ; a long seta on each side of the prothorax
towards both apex and at base; head with two sete on each side
behind antennz.
Head slightly longer than wide, obtusely longitudinally impressed
on each side of middle, sides from antenn to base gently curved.
Antenne extending to hind coxz, two basal joints stout, first almost
as long as second and third combined, second distinctly longer and
stouter than third, third to tenth more or less head-like, eleventh
slightly wider and about twice the length of tenth. Prothorax dis-
tinctly wider than head, not much wider than long, apex incurved to
its middle, sides rounded in front and slightly incurved to base, base
strongly bisinuate ; median line narrow and distinct ; impunctate.
Elytra distinctly wider than prothorax, more than twice as long as
wide, not quite covering tip of abdomen, sides gently diminishing in
width to base and rather more noticeably to apex, margins finely
channelled throughout, densely but obscurely punctate. Mesosternal
epimera wide. Legs strong ; front tibie strongly dilated and deeply
of Australia and Tasmania. 367
notched towards apex, the others slightly dilated and not notched ;
tarsi rather short. Length 14 to 2mm.
Hab. N.S. WALES, Otford. (A. J Carter and A. M.
Lea.)
The male differs from the female * in being smaller,
with stouter legs, longer antennz, and wider front tarsi.
The two specimens described were taken under small but
deeply buried stones in a small gully near the. Otford
railway station. The pubescence can only be seen dis-
tinctly from the sides; the sete are also by no means
distinct.
Readily distinguished from J. stephensi by its much
greater size, and by the absence of the strong elytral striz
so noticeable in that species. Other differences are that
in stephensi the prothorax at its widest is very little wider
than the head or narrower than the elytra, and that its
apex is considerably wider than its base and its sides with
less sinuous outlines.
COLYDIID.
ANOMMATUS 12-sTRIATUS, Mill.
This minute British species I have taken at the roots of
grass in Hobart. Specimens must be common, although
seldom seen, as [ obtained nineteen living specimens and
fragments of many others by the use of sieves. The plan
adopted was to break up the earth for about three inches
from the surface, this was then thrown into water and
stirred about; every thing that floated was picked out,
dried, and afterwards put through sieves, the rubbish that
came through the finest sieves being gone over on white
paper in the ordinary way. ‘The specimens would often
remain for minutes together without moving, and even
when they did move would walk at a very slow rate.
SCARABAEID&.
PHYCOCHUS GRANICEPS, Broun.
In Hobart, obtained at the roots of plants (usually the
bracken fern) growing close to the sea-beach. I have seen
* The type female has been returned to Mr. Carter.
368 Mr. A. M, Lea on the Blind Coleoptera.
very few living specimens, but many dead ones. Mr. J. J.
Walker (who first took it at Hobart, in June 1901) informs
me thatin New Zealand he obtained many specimens under
old logs partially buried in the sand of sea-beaches.
PHYCOCHUS SULCIPENNIS, Lea.
Obtained at Hobart in company with the preceding
species, than which it is rather more numerous.
CURCULIONIDA.
HALORHYNCHUS GENICULATUS, Lea.
Numerous specimens of this species were obtained at
the roots of a small species of salt-bush (Aériplex) growing
just above high-water mark on the outer beach at Gerald-
ton (Champion Bay). They remained quite motionless
for a considerable time after being exposed.
HALORHYNCHUS Cacus, Woll.
I repeatedly searched for this species at Cottesloe and
other beaches close to Fremantle (the original locality) but
never found more than two specimens; these were taken
at the roots of a species of spinifex grass.
TASMANICA MYRMECOPHILA, Lea.
The type specimen of this species was obtained in
Hobart under a stone in the nest of an ant. No other
specimen has been recorded, although the species has been
repeatedly searched for; but there is a minute specimen
in the Australian Museum (from the late Rev. R. L. King’s
collection, without locality label, but probably taken about
Parramatta) which may belong to the species. The
specimen in question is old and dirty, but I could see no
eyes in its head, and from memory it struck me as
probably being 7. myrmecophila.
«(°° 369. J
XXI. On «@ Collection of Butterflies made in Marocco, in
1900-01-02. By E.G. B. MrapEe-WAxpo. (Com-
municated by H. J. Eiwes, F-.R.S., etc.)
[Read October 18th, 1905. ]
Plates XVIII, XIX.
THouGcH the butterflies of the neighbourhood of Tangier
and of some other points on the coast of Morocco have been
collected by the late Mr. J. H. Leech and Mr. J. J. Walker,
yet no really important collection has been made in this
little-explored country. The following collection was made
during an eighteen months’ residence in Marocco, Col-
lecting at and in the neighbourhood of Tangier, viz. within
20 miles, and on a long excursion which started from Tan-
gier on May 8th, 1901, and lasted until August 21st. We
went by way of Busharin to Rabat, thence by Fedulla, and
striking inland over the central plains to Beni Meskin,
crossed the Ouad Moorbey, Oom-er-rebia of the maps, and
went eastward, spending some time at a tiny tent village
on the desert called Oolad Lasara. Throughout the central
plains the heat was great, often reaching 116° in the shade,
with hot nights, and owing to this, and also to a previous
visitation of locusts which had consumed almost all vegeta-
tion, nothing much could be done except on the banks of
the river. From here we went slowly south to Marrakesh
(Marocco City), and then into the Great Atlas, which we
entered at Agiirgir; from here we went, first southward
until we struck the valley of the Ouad Nyfys, which we
followed until we got to the watershed above Tsigidir-el-
bor, then retracing a short way struck westward until we
reached the “ Amsmiz.” Here we encamped for some time
at a Berber village called Sould Jedid, which was delight-
fully situated about 6,000 feet up, and with most pleasant
inhabitants. The mountains here only rise to about 9,000
feet and are to a considerable extent covered with damp
forest on their northern slopes; this forest consists largely
of Arar (Callitris quadrivalvis), Prickly Oak, Holly, Laurus-
tinus, Arbutus, etc. We did not come to Pine (Pinus
halepensis) until we reached the Imentalla valley a day’s
journey further on. After spending some days collecting
in this neighbourhood, we started light, with only a little
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.)
370 Mr. E, G. B. Meade-Waldo on a
food, our mules and one pack mule, up a lateral valley so
as to get nearer to a great peak, which the natives called
Tizi Gourza, but which according to Thompson’s map must
be Jebel Ogdimt. We bivouacked at the last hamlet at
the top of this valley, a place called Imi Ouern. This
place, which appeared to be “the end of all things,” would
have been splendid for a prolonged stay, but no food
could be procured, shooting was not safe, and we were a
very small party. It was most beautifully situated and
appeared to be highly productive in butterflies and moths.
We ascended the mountain and made a most interesting
collection of butterflies; from here we returned to Imen-
talla, and by slow stages worked our way northward and
westward by way of Anzoot and Tafegar to Mogador, which
we reached on August 21st. In the early spring of 1902 I
paid a visit to the forest of Marmora, the only tract of
primeval forest in the north of Marocco. It lies between
the Seboo and Bou-reg-reg, east of Rabat. It is entirely
unexplored, and would prebably be very productive, it is
however populated only by bandits, and anything like
shooting or collecting is almost impossible. However, I
managed to collect a considerable number of moths, and a
few butterflies. The primary object of these excursions
were ornithological, but I collected both butterflies and
moths all the time, and paid special attention to the high
mountain species. An analysis of this list will show how
very few species there are in these remote regions that are
not also to be found in the Mediterranean region, and that
these mountains have developed comparatively few Alpine
species,
I was accompanied on both these excursions by M. Henri
Vaucher of Tangier, as taxidermist and interpreter. M.
H. Vaucher is a Swiss gentleman who has lived most of
his life in Marocco, is an enthusiastic naturalist himself,
and such success as attended these expeditions is almost
entirely due to his great tact and intimate knowledge of
how to treat the extremely difficult inhabitants of this
fascinating country. It is to Mr. H. J. Elwes that I must
tender my thanks for the trouble he has taken in going
over the collection of butterflies and in assisting me with
naming those that are new, and to Sir George Hampson
for going through and naming the moths.*
* The remarks by Mr. Elwes are in brackets signed “H. J. E.”
Collection of Butterflies made in Marocco, 1900-2. 371
1. Papilio podalirius, L. and var. feisthameli, Dup.
March 12th, 1901, bred, fed on Cherry.
Tangier, March 22nd, 1901. Amsmiz, June 27th, 1901.
Imentalla, 5,500 feet, July 9th, 1901.
Is universally distributed, and on the wing the whole
summer, very fond of the tops of low mountains of about
5,000 ft. elevation, where it may be seen in numbers on
the leeward side during the heat of the day. The larva
feeds on Cherry. Both P. podalirius and var. feisthamelia
occur at Tangier in March,
2. P. machaon, L.
Tangier, January 19th, 1901. Just emerged. Amsmiz,
June 27th, 1901. Tangier, August 29th, 1901. Forest
of Marmora, March 28th, 1902.
On the wing at intervals from January until September.
3. Thais rumina, L.
Tangier, February 27th, 1901. Tangier, April 8rd,
1901. Tangier, March 9th, 1901. Tangier, March 19th,
1901 (var. canteneri, Hey.).
This variety is not very uncommon late in the season,
and only occurs as far as I could seein the . The perfect
insect is only on the wing from the end of February until
the beginning of April. Feeds on Aristolochia betica.
4, Preris brassice, Li.
Only taken at Tangier in all months from December to
August. I did not see it on the wing in the summer
anywhere in the central plains.
Tangier, January 17th, 1900, December 25th, 1900;
August 26th, 1901; January 17th, 1902.
[“The specimens are large but do not approach var.
cheiranthi, Hiibn., and the seasonal differences are slight.
“The form known as var. wollastont, Butler, from Madeira
closely resembles some of these.’—H. J. £.]
5. Preris rape, L.
On the wing all the year round in the north. Idid not
see it in the Great Atlas, but throughout the plains
wherever there was moisture diminutive forms were
ubiquitous in summer.
Tangier, March 13th, 1901. Klatsa, May 13th, 1901.
Saffi, August 27th, 1901.
ate Mr. E. G. B. Meade-Waldo on a
6. Pieris daplidice, L.
Universal in summer, taken only in the Atlas from 6,500
to 9,000 ft.in July in the central plains. I saw many
diminutive individuals.
Wad Moorbey, June 16th, 1901. Sould Jedid, July 3rd,
1901. Tsauritz Entsagauz, 9,000 feet, July 13th, 1901.
7. Luchloe belemia, Esp.
Extremely abundant and universal, and on the wing
almost the whole year. I did not see many in the height
of the summer.
Tangier, December 8th, 1900; March 18th, 1902.
8. Huchloé eupheno, Esp., and var. 2 androgyne, Leech.
(Plate XIX, fig. 7, var. 2 androgyne.)
Extremely abundant and universal during March and
April in the north of Marocco. I saw very few in the
forest of Marmora, and the only one, a 9, which I caught,
is an androgyne. :
Tangier, March 9th, 1901. Tangier, April 15th, 1901.
Hawara, May 4th, 1901. Forest of Marmora, March 28th,
1902, 2, var. androgyne, Leech.
9. Teracolus daira, Klug., var. nowna, Lue.
(Plate XIX, fig. 8, 3, 9, .)
I only saw this species at Agirgtir, where it was abund-
ant in a damp wood at an elevation of about 3,500 ft.
I saw one torn specimen near Tsigidir-el-Bor about
30 miles south.
Agtrgir, June 24th, 1901. Tsigidir-el-Bor, June
25th, 1901.
10. Colias edusa, L.
On the wing practically all the year. Extremely abund-
ant in summer, var. /elice, Hiibn., not uncommon. I took
one specimen with one forewing, /elice, the other normal.
Very variable.
} Tangier, December and January. Interior, May and
une.
ll. Gonepteryx rhamni, L.
Is common in the north of Marocco, a very large
form. I did not see any in the south; it is on the wing at
intervals during the winter and spring, and again in the
Collection of Butterflies made in Marocco, 1900-2. 3873
Jate summer ; both it and the next species feed on Khamnus
alaternus.
Tangier, March 22nd, 1901. Tangier, December 28rd,
1902.
12. Gonepteryx cleopatra, L.
Is commoner, and more widely distributed than the last.
I saw it throughout the country.
Tangier, January Ist, 1901. Fedulla, May 28th, 1901.
Moorbey, June 5th, 1901. Sould Jedid, 6,500 feet, July
1901.
13. Charaxes jasius, L.
Locally common, somewhat of a mountain species, but
the males travel long distances; frequents fig-trees when
the fruit is ripe and becomes stupefied. A few on the
wing in April and May, and common in August and
September. Some specimens are very large.
Tangier, August 16th, 1901, September 15th, 1901.
14, Pyrameis atalanta, L.
On the wing nearly all the year.
Tangier, Feb. 16th, 1901. Tangier, March. Amsmiz,
June 19th, 1901. Imentalla, 5,500 feet, July 8th, 1901.
15. Pyramers cardut, L.
On the wing all the year, late autumn brood small and
dark.
16. Vanessa c. album, L.
I saw two or three specimens high up on Tizi Gourza,
and took one somewhat worn, large pale specimen on July
11th, at about 12,000 feet.
[I am not aware that this has been taken in North
Africa before.’—H. J. E.]
17. Melitxa didyma, O.
T did not see this species until I got to Rabat, after that
I saw it at intervals throughout the country, both in the
plains and at a considerable height in the mountains.
Rabat, May 16th. Agurgtr, July 6th, 1901. Tsauritz
Entsagauz, 9,000 feet, July 4th.
374 Mr. E. G. B. Meade-Waldo on a
[“ This form agrees with the specimens from Kabylia
and the oasis of Biskra known as var. deserticola, Obth.;
it does not vary appreciably between the plains and high
mountains.”—Z, J. E.]
18. Argynnis lathonia, L.
I only saw this in the Atlas, where it was not uncommon.
Sould Jedid, July 1901. Tsauritz Entsagauz, July
1901.
19. Argynnis pandora, Schiff.
I only saw two worn specimens in the north of Marocco ;
it is certainly scarce on the low ground ; it was abundant
in the Great Atlas, and I saw it at an elevation of over
12,000 feet.
Sould Jedid, July 2nd, 1901. Tizi Gourzd, July 11tb,
1901.
20. Melanargia ines, Hoftm.
I only saw it at one spot on the leeward side of a stony
peak, everywhere else the following was abundant.
Tsauritz Entsagauz, July 6th, 1901.
21. Melanargia lucasi, Ramb.
Abundant in the Great Atlas, most frequent in the
woods.
Tsauritz Entsagauz, July 4th. Imentalla, July 8th,
1901. Sould Jedid, July 2nd.
A very large and pale form was taken in the central
plains in June, and also on the high table-land above
Agirguir, on June 23rd.
22. Satyrus briseis, L.
Locally common high up in the Atlas, but not on the
highest. tops.
Tizi Gourza, July 11th, 1901, about 9,000 feet.
L“ Very variable, and large, under-side of hind-wing
very strongly marked.”—ZZ, J. £.]
23. Satyrus semele, L.
I only saw a few in the mountain woods.
Imentalla, July 6th, 1901. Tsauritz Entsagauz, July
4th, 1901.
Collection of Butterflies made in Marocco, 1900-2. 375
24. Sutyrus mniszecht, H.S., var. maroccana, Meade-
Waldo, n. var.
Plate XIX, fig. 3, g, 4, 9.)
With the exception of one individual I only saw this
insect on one stony slope at the top of one of the peaks
of Tsauritz Entsagauz. There it is abundant.
Tsauritz Entsagauz, July 6th and 7th, 1901, 9,000 ft.
[“ The occurrence of a form of this purely eastern group
so far from any region where any have been previously found
is very curious, but it may be constantly distinguished
from the Syrian or Persian forms, by the different form of
the sexual band on the fore-wing of the ¢, which instead
of forming a broad velvety patch of raised scales extend-
ing almost to the hind margin of the fore-wing, as in the
Syrian form of the species, is a comparatively narrow
band. Another point of distinction is in the cilia of the
hind-wings which are very much less scolloped than in
mniszechi or in the various forms of S. pelopea. On the
under-side, though the median bands of the fore-wing are
better marked than usual, yet the position of all markings
are identical in Syrian specimens, and I do not therefore
think it justifiable to treat this as a distinct species.’— _
ed, Li]
25. Satyrus abd-el-kader, Pier.
I saw this fine insect commonly on Tizi Gourza, but on
such bad ground I was only able to take two worn
females.
Tizi Gourza, July 11th, 1901.
26. Satyrus actea, Esp.
Was common and only just on the wing in the Great
Atlas, at from 8,000 to 10,000 ft. mostly on open ground.
Tsauritz Entsagauz, July 6th, 1901, also on Tizi Gourza.
27. Pararge mexra, L,
I only saw a few specimens on Tizi Gourzi, and no-
where else in Marocco. I am not aware that this species
has been taken previously in North Africa.
Tizi Gourzd, at about 10,000 ft., July 11th, 1991.
28. Pararge megera, L.
Not by any means abundant, but out at intervals
throughout the year.
376 Mr. EF. G. B. Meade-Waldo on a
Tangier, December 3rd, 1900. ‘Tsauritz Entsagauz
July 4th, 9,000 ft.
29. Epinephile jurtina, L., var. fortunata, Alph.
Very generally distributed all through the summer.
Klatsa, May 14th, 1901. Ras Doura, May 19th, 1901.
Tsauritz Entsagauz, July 7th.
30. Lpinephile lycaon, Kiihn, var. mauwritanicus, Ober.,
also var. lupinus, Costa.
Common in the Atlas woods. Not seen anywhere else.
Tsauritz Entsagauz, July 6th, 1901. Imentalla, July
9th, 6,500 ft.
31. HLpinephile ida, Esp.
Common all the summer and universally distributed.
Rabat, May 25th, 1901. Tsauritz Entsagauz, July 4th.
Tangier, Sept. 1901,
32. Hpinephile pasiphaé, Esp.
Very common but local. I did not see it in the south.
El Mediar, May 11th, 1901. Klatsa, May 13th, 1901.
33. Canonympha arcanioides, Pierr.
Locally common in the north of Morocco. I did not see
it in the south:
Hawara, April 1901. El Mediar, May 12th, 1901.
34. Cenonympha fettigi, Obth.
I only saw this insect at one spot above Imentalla,
where however it was abundant.
Imentalla, 5,500 ft, July 8th and 9th, 1901.
35. Cenonympha pamphilus, L.
I took typical C. pamphilus, in the plains of the north
of Morocco in May 1901, also on the downs by the sea
between Mehedia and Rabat in March 1902.
In spring in the north of Marocco, where there appears
to be no summer brood.
36. Cenonympha lyllus, Esp.
Very numerous south of Rabat, and throughout the
south of Morocco, ascending the Great Atlas to at least
10,000 ft. during the summer.
Collection of Butterflies made in Marocco, 1900-2. 377
Imentalla, July 9th, 1901, 5,500 ft. Amsmiz, June
16th, 1901.
37. Cenonympha vauchert, Blachier. Bull. Soc. Ent.
France, 1905, p. 213. (C. meade-waldoi, n. sp. Elwes MS.)*
Plate XIX, fig. 1, ¢, 2, 2.
It was abundant on the more barren stony places near
the top, but none occurred amongst the broom-covered
slopes lower down.
Fairly common on Tsauritz Entsagauz, and very
abundant on Tizi Gourzd up to the top of the mountains.
Seen nowhere else. Tsauritz Entsagauz, July 6th. Tizi
Gourza, July 11th, 1901.
I may incidentally add that the top of Tizi Gourza is
entirely stone boulders and great slabs of rock for the
last 500 ft., and on reaching the extreme summit we saw
a hollow place beneath a great slab, in this were some
bits of rag held down by stones, and from the top fluttered
some white rags from a stick jammed in between the
rocks. It was a holy place, and our susi, Hammoo, the
only one of our Moors who would accompany us, crept in,
lay down on his face and prayed.
[This is a very distinct species which can be mistaken
for no other, and on account of the extreme difficulty of
reaching the place where it was found is likely to remain
very rare in collections. I do not know with what species
to compare it, and as the figures will show its characters
better than any description I will only say that it seems
to be the most Alpine species found in the Southern
Atlas, for although Mr. Meade-Waldo saw many other
insects as high up on the mountain, he never saw this
butterfly except at a very considerable elevation, certainly
not below 8,500 ft."—H. J. £.]
38. Thecla ilicis, Esp., var. mauritanica, Stgr.
Abundant everywhere where cork oak and prickly oak
occur.
Busharin, May 16th. Ras Doura, May 18th. Tsauritz
Entsagauz, 1901.
* Mr. Meade-Waldo’s specimens of this new and very interesting
Satyrid in the Natural History Museum, from which the figures on
Plate XIX have been drawn, agree in all respects with M. Blachier’s
description of C. vaucheri, and I have therefore adopted this name.
—J. J. W.
378 Mr. E. G. B. Meade-Waldo on a
[“' This seems to agree best with the form described by
Staudinger from North Africa ‘minor subtus obscurior fere
unicolor, but it varies greatly, and some specimens might
be referred to 7’. acacix.”—H. J. E.]
39. Callophrys rubi, L.
I only saw it in the north of Marocco, where it was
common in early spring almost everywhere.
Tangier, March 26th, 1901; March 18th, 1902.
40. Chrysophanus thersamon, Esp., var. omphale, Klug.
I only took one worn specimen on the outskirts of
Marocco City on June 12th. This species has not, as
far as we know, been previously taken in Africa.
41. Chrysophanus alciphron, Rott., var. gordius, Sulz.
T only saw it in the Atlas, where it did not seem
common; it was probably going over, as I only took two
specimens that were fresh.
Tizi Gourza, July 11th. Tsauritz Entsagauz, July 6th,
ils OM ts
[“ This also has not previously been recorded from North
Africa; the male has no violet sheen on the upper
surface.’—H. J. H.]
42. Chrysophanus phleas, L.
Common and universal, extremely abundant locally,
subject to great variation, and is out all the year.
Tangier, March 10th, 1901. Tangier, Feb. 27th, 1901.
Marrakesh, June 10th, 1901.
43. Thestor mauritanicus, Luc.
Common, but extremely local; is on the wing for only
a short time in early spring.
Tangier, March 11th. Bubana, March 27th. Hawara,
April 4th, 1901. Larva feeds on the pod of a leguminous
plant. ;
44, Thestor ballus, Fabr.
Is as the last, but not quite so local, and is also on the
wing earlier and a little later.
Tetuan, January 16th. Tangier, March 6th. Hawara,
April 4th, 1901.
Collection of Butterflies made in Marocco, 1900-2 379
45. Lampides beticus, L.
Fairly common and on the wing all the summer. I
saw a worn specimen in December 1900.
46. Lampides telicanus, Lang.
Commoner than the last, and out earlier.
Tangier, July 13th. Rabat, May 26th. Rehamma,
June 9th, 1901.
47. Lampides theophrastus, Fabr.
I only saw this species south of Ouad Moorbey from
where it reached the foothills of the Atlas, and was abund-
ant wherever the Jujube bush (Zizyphus lotus) grew.
Moorbey, June 6th. Anzoot, July 14th, 1901.
48. Lampides jesous, Guér.
The distribution the same as the last, but most common
in the valley of the Ouad Moorbey.
Moorbey, June 5th. Below Agtrgir, June 21st, 1901.
49, Lycena lysimon, Hiibn.
I saw a few at Busharin, and it swarmed over some
pools of water outside Marrakesh.
Busharin, May 16th. Marrakesh, June 12th. Amsmiz,
June 27th, 1901.
50. Lycena baton, Berg., var. abencerragus, Pier.
I only saw one specimen below Sould Jedid.
Amsmiz, June 28th, 1901,
51. Lycena wcarus, Rott.
Fairly universal, but not remarkably common. From
Tangier in August, and various places up to 9,000 feet.
Tangier, August 17th. Busharin, May 16th. Sould
Jedid, July 13th. Imentalla, July 8th, 1901.
[“ Many specimens have spots on the hind margins, and
might be called var. celina, Aust., but this var. is evidently
inconstant.” —H. J. E.]
52. Lycena astrarche, Bergstr.
Almost universal at all elevations, and in the plains,
positively swarming in some woods.
380 Mr. E. G. B. Meade-Waldo on a
Bershid, May 30th. Imentalla, July 9th. Ouad
Moorbey, June 9th. Tangier, April 3rd, 1901. Only
found in summer, and as usual in hot regions very brown -
below (var. calida, Bell).
53. Lycena hylas, W. V.
I only saw this butterfly on Tsauritz Entsagauz, and at
Tmentalla, where it was extremely abundant on the top,
of a low mountain, 5,500 feet.
Imentalla, July 8th, 1901.
(Plate XIX, fig. 5, 3, 6, 2.)
[“ Var. atlantica, Elwes, n. var.
“This is quite an unexpected discovery in Marocco, and
a very interesting one, because though a variety of this
species occurs in the higher region of the Sierra Nevada,
and in some places in the mountains of eastern Spain, yet
the male of this form (nivescens, Kef.) is of a grey-blue
colour above, whilst the colour of the male of atlantica
is to my eye like that of typical hylas, though Mr, Meade-
Waldo sees a difference.
“Some males have marginal black spots on the hind-wing
above, and both sexes have the white stripe on hind-wing
beneath, broad and well marked. All the females have a
broad band of orange marginal markings on the fore-wings
which I have never seen so well marked in any European
hylas.”—H. J. E.|
54. Cyaniris argiolus, L.
Appears first in February, and at intervals throughout
the summer.
Tangier, February 20th. Amsmiz, June 26th. Tangier,
August 28th, 1901.
55. Adopexa thaumas, Hiibn.
Tsauritz Entsagauz, 9,000 ft., June 1901. Common.
56. Adopxu actxon, Rott.
Ras Doura, May 1901. Common.
57. Adopxa hamza, Obth.
Rabat, May 26, 1901. Common.
58. Parnara zelleri, Led.
Local and scarce, frequents wet places in woods, flies
extremely fast.
Bashasin, May 20th. Tangier, August 30th, 1901.
Collection of Moths made in Marocco, 1900-2. 381
59. Parnara nostrodanus, Fab.
Extremely abundant locally in late summer in open
stony places.
Tangier, August 28th, September 2nd, 1901.
60. Carcharodus alcew, Esp.
Imentalla, 5,000 ft. July 1901, Common.
61. Carcharodus althex, Hiibn.
Plains of Marocco, rather scarce.
Mehedia, May 1901.
62. Hesperia ali, Obth.
I only saw a few on low ground.
Tangier, August 30th, 1901.
63. Hesperia serratule, Raimb.
[“ Though I do not profess to know how serratule can
with certainty be distinguished from a/vews without study-
ing the genitalia, which in this case I have not done, yet
these specimens from Imentalla and Agurgur appear to
me to belong to serratulx, though according to Staudinger
onopordt of Rambur is the form found in Marocco.”—HH.J. £.]
Moths collected in Marocco. By EK. G. B. MEADE-WALDO.
ARCTIADA.
ARCTIANA,
1. Huprepia cribraria, Linn., Syst. Nat.,i, p. 507 (1758).
Forest of Marmora. The variety punctigera, Frr., L.
candida, Cyr.
March 26th, 1902. Ensar, Kabyla amar, Forest of
Marmora.
2. Arctia villica, Linn., Syst. Nat., i, p. 501 (1758).
All the specimens belong to the form onewkat, Freyer.
Abundant in April at Sharf-at-Akab, Tangier.
3. Cymbalophora pudica, Esp., Schmett., iii, p. 177,
pl. 33, f. 1 (1784).
Abundant in August and September. The larva, which
feeds on various grasses, spins a slight cocoon in the
TRANS. ENT. SOC. LOND. 1905.—PART Iv. (DEC.) 26
382 Mr. E. G. B. Meade-Waldo on a
bottom of thick herbage, but remains in its larval state
until within about fourteen days of emerging.
4. Utetheisa pulchella, Linn., Syst. Nat., i, p. 534 (1758).
Locally and sporadically very abundant in August and
September.
NOCTUIDA..
AGROTINE.
5. Chloridca dipsacea, Linn., Syst. Nat., xu, p. 856 (1766).
Forest of Marmora.
Very common the end of March. Flying by day, 1902.
6. Chloridea peltigera, Schiff., Wien Verz., p. 89 (1776).
Distributed throughout the central plains of Marocco.
June and July, 1901. .
7. Huxoa segetis, Schiff., Wien Verz., 11, 81, 252, ff. 3. a.
b. (1776).
Common at Tangier in spring and autumn.
8. Huxoa spinifera, Hiibn., Samml. Eur. Schmett. Noct,
f, 389 (1827).
Tangier.
9. Agrotis ypsilon, Rott., Naturf., ix, p. 141 (1776).
Tangier, May 1901.
10. Agrotis comes, Treit., Schmett. Eur., v, 1, p. 254
(1825).
Tangier. One specimen,
11. Agrotis pronuba, Linn., Syst. Nat., i, p. 512 (1758).
The only one seen, September 1901.
Tangier. Two specimens.
12. Agrotis c-nigrum, Linn., Syst. Nat., x, p. 516 (1758).
Bred from dug pupa, March 1902.
13. Hpisilia faceta, Treit., Schmett. Eur., x, 2, p. 35
(1836).
Very common at Tangier throughout the winter.
Collection of Moths made in Marocco, 1900-2. 383
14. Lycophotia margaritosa, Haw., Lep. Brit., p. 218
(1809).
Bred from dug pupa, March 1902.
HADENIN.
15. Mamestra oleracea, Linn., Syst: Nat., ed. x, p. 517
(1758).
Bred from dug pupa, March 1902.
16. Mamestra chrysozona, Bork., Eur. Schmett., iv,
p. 264 (1792).
Forest of Marmora, Ensar. Forest of Marmora, April
1902.
17. Leucania l-album, Linn., Syst. Nat., ed. xii, p. 850
(1766).
Ensar. Forest of Marmora, April 1902.
18. Leucania loreyt, Dup., Lep. Fr., vii, 1, p. 81, pl. 105,
er bes).
Central plains, June 1901.
19. Glottula pancratii, Cyr., Ent.Neap., pl. 12, f. 4 (1787).
Tangier.
CUCULLIANZ.
20. Cucullia chamomillx, Schitf., Wien Verz., p.73 (1776).
Very abundant in January and February, 1902.
21. Cucullia tanacett, Schiff., Wien Verz., p. 73 (1776).
Tangier.
22. Bombycia viminalis, Fabr., Gen. Ins., p. 284 (1777).
Forest of Marmora.
Came to light on March 28th, 1902.
23. Cleophana dejeani, Dup. Lep. Fr., vii, p. 115 (1827).
Came to light at Sharf-at-Akab, Tangier, April 1901.
24. Cleophana pauli, Staud., Iris., iv, p. 306, pl. 4, f. 4
(1891).
(Plate XIX, fig. 11, 9.)
Forest of Marmora, April 27th. Flying by day.
384. My. E. G. B. Meade-Waldo on a
25. Cleophana difiuens, Staud., Berl. Ent. Zeit. 1870,
p. 121.
(Plate XIX, fig. 12, var. maroccana, §.)
Forest of Marmora, at light March 27th, 1902. The
variety maroccana.
26. Metopoceras felicina, Donz., Ann. Soc. Ent. Fr., 1844,
paleo; plG, £2.
Forest of Marmora, at light March 27th, 1902.
ACRONYCTINE.
27. Brotolomia meticulosa, Linn., Syst. Nat., p. 513 (1758).
Only saw three in Marocco, March 1901.
28. Huplexia solrerr, Boisd., Ind. Meth., Errat., p. 4 (1829).
Tangier, September 1901.
29. Acronycta runicis, Linn., Syst. Nat., p. 516 (1758).
Tangier.
30. Acronycta megacephala, Fabr., Mant. Ins., p. 175
(1787).
Tangier, September 1901.
31. Laphygma quadripunctata, Fabr., Syst. Ent., p. 594
(1775).
Tangier.
32. Amphipyra eriopoda, Herr.-Schif., Kur. Schmett,
li, p. 413, f. 596 (1851).
Tangier.
CATOCALINA.
33. Catocala conversa, Esp., Schmett., f. 105 (B) 1-2
(1787).
Tangier. The variety ayamos, Hiibn.
34. Catocala oberthuri, Aust., de. Nat. 1879, p. 85.
Very common, August and September. Tangier.
35. Ophiusa tirrhea, Cram., Pap. Exot. ii, p. 116,
pl. 172, E (1780).
Not common, Sharf-al-Akab. Tangier, April 1901.
36. Ophiusa algira, Linn., Syst. Nat. ed. xii, p. 836
(1766).
Tangier.
Collection of Moths made in Mavocco, 1900-2. 385
37. Ophiusa lunaris, Schiff., Wien. Verz., p. 94 (1776).
Forest of Marmora. March 28th, 1902.
38. Grammodes geometrica, Fabr., Syst. Ent., p. 599 (1775).
Tangier.
39. Grammodes stolida, Fabr., Syst. Ent., p. 599 (1775).
Tangier.
PLUSIANZ,
40. Plusia gamma, Linn., Syst. Nat., i. p. 513 (1758).
Abundant everywhere.
NoctTuIn&.
41. Apopestes spectrum, Esp., Schmett., iv, p. 131,
pl. 100, ff. 3, 4 (1786).
Common at Tangier in summer. These specimens
were bred from pupa found in May, near Fedulla, on the
West Coast. ‘The white silk cocoons were abundant, and
attached to Huphorbia on the sea coast.
HRASTRIAN A.
42. Callopistria latreilli, Dup., Lep. Fr., vii, Pp. a29,
plet20.f 2 (1827):
Taken on Olive, above Amsmiz in the Great Atlas (1901).
43. Tarache lucida, Hiibn., Berl. Mag., iii, p. 302 (1767).
Very common, on the plain near Fedulla in May 1901.
The variety albicollis, F.
44. Tarache luctwosa, Esp., Schmett., pl. 88, f. 4 (1786).
Tangier.
45. Eublemma ostrina, Hiibn., Eur. Schmett. Noct.,
1; ao) (1827).
Tangier, February 1902.
HYPENINA.
46. Zanclognatha crinalis, Treit., Schmett. Eur., vu,
p. 17 (1829).
Tangier, February 1902.
386 Mr. E. G. B. Meade-Waldo on a
47. Hypena obsitalis, Hiibn., Eur. Schmett. Pyr.,
ff. 164-165 (1827).
Tangier.
48. Hypena lividalis, Hiibn., Beitr., 11, 4, pl. 1, E (1827).
Tangier, December 1901.
LYMANTRIADA.
49. Huproctis chrysorrhaa, Linn., Syst. Nat., p. 502
(1758).
Tangier.
50. Lymantria dispar, Linn., Syst. Nat., p. 501 (1758).
Tangier,
51. Lymantria atlantica, Ramb., Faun. And., pl. 15,
f, 7 (1838).
Tangier,
SPHINGIDA.
ACHERONTIANA.
52. Acherontia atropos, Linn., Syst. Nat., i, p. 490
(1758).
Tangier.
The only one seen or heard of, September 1901.
AMBULICINE.
. Smerinthus populi, Linn., Syst. Nat., p. 489 (1758).
53
Tangier. he variety austauti ab. tncarnata.
CHG@ROCAMPINE.
54, Deilephila lineata, Fabr., Syst. Ent., p. 541 (1779).
April and May, flying by day as well as in evening.
Tangier.
55. Cherocampa celerio, Linn,, Syst. Nat., 1, p. 491
(1758).
Very common. ‘Tangier in September, comes to P/wm-
bago at sunset.
Collection of Moths made in Marocco, 1900-2, 387
MACROGLOSSIN A.
56. Macroglossa stellatarwm, Linn., Syst. Nat., 1, p. 493
(1758).
Very abundant everywhere.
NOTODONTID.
57. Dicranura vinula, Linn., Syst. Nat., i, p. 499 (1758).
Tangier, 1901.
1 .
Saw many old cocoons on Aspen and Willow, but only
two containing living pups, from one of which this species
emerged.
58. Phalera bucephala, Linn., Syst. Nat., p. 508 (1758).
Tangier. The variety bucephalina.
GEOMETRIDA.
BOARMIANZ.
59. Opisthograptis luteolata, Linn., Syst. Nat., p. 525
(1758).
Tangier.
60. Eurranthis pennigeraria, Hiibn., Eur. Schmett.,
Geom., f. 363 (1827).
Akba Hamra. The variety chrysitaria, H. G.
61. Aspilates ochrearia, Rosst., Mant., i, pu.od* pl.f,
N (1794).
Forest of Marmora. “Metradia. Ras Doura.
Abundant; a day-flyer. March 1902.
62. Amygdaloptera testaria, Fabr., Ent. Syst, p. 143
(1794).
Forest of Marmora.
Abundant; a day-flyer. March 1902.
63. Fidonia famula, Esp., Schmett., iv; pl OG AE 4
(1787).
Forest of Marmora. In extraordinary abundance.
March 1902.
388 Mr. E. G. B. Meade-Waldo on a
64. Thamnonoma gesticularia, Hiibn., Eur. Schmett.
Geom., ff. 472-3 (1827).
The only specimen seen. February 1902. Tangier.
65. Thamnonoma semicanaria, Frr., Beitr., 78, 2, 1,
p. 145 (1833).
Tangier, Bubana. March 1901.
66. Selidosema ericetaria, Vill. Linn., Ent., 11, p. 329,
pl. 6, f. 9 (1789).
Tangier.
67. Gnophos asperaria, Hiibn., Eur. Schmett., Geom.,
f. 484 (1827).
Tangier. March 1902.
68. Boarmia (?) gemmaria, Brahm., Ins, Kal., 11, p. 255
(1791).
More uniform above, and more fulvous below, than the
typical form.
The only one seen March 1902. Tangier.
69. Boarmia abietaria, Goeze, Beitr. iii, 3, p. 439 (1781).
Forest of Marmora. March 27th, 1902.
70. Boarmia abruptaria, Thnb., Diss. Ent., iv, p. 59,
play, 1.3 G07 92).
Very common in winter at Tangier.
71. Boarmia atlanticaria, Staud., Stett. Ent. Zeit., 1859,
jteeralkey
Only one taken, March 1902. Tangier.
LARENTIAN.
72. Sterrha sacraria, Linn., Syst. Nat., ed. xii, p. 863
(1766).
Tangier.
73. Anaitis plagiata, Linn., Syst. Nat., i, p. 526 (1858).
Common at Tangier in winter.
Collection of Moths made in Marocco, 1900-2. 389
74. Cidaria malvata, Rmbr., Ann. Soc. Ent. Fr., 1832,
p. 43, pl. 2, f. 5.
Tangier. October 1901, only one seen.
75. Cidaria siterata, Hiibu., Berl. Mag,, iv, p. 522 (1769).
Tangier, February 4th, 1902. Given me by Lord
Walsingham.
76. Cidaria basochesiata, Dup., Lep. Fr., vin, 1, p. 559,
pl. 210, f. 5 (1831).
Tangier. February 2nd, 1902. Given me by Lord
Walsingham:
77. Eupithecia oblongata, Thnb., Diss. Ent., 1, p. 14,
f. 12 (1784).
Near Tangier, at Ain Zeitun. February 1902. Common.
GEOMETRIN&.
78. Pseudoterpna coronillaria, Hiibn., Eur. Schmett.,
Geom., ff. 479-482 (1827).
Near Tangier, at Ain Zeitun. February 1902.
ACIDALIAN.
79. Rhodostrophia sicanaria, Zell., Stett. Ent. Zeit., 1852,
p. 180.
Atlas Mountains, Imentalla. 5,000 feet. July 9th, 1901.
80. Ephyra pupillaria, Hiibn., Eur. Schmett., Geom., f.
69 (1827).
Bred from larva. March 1901.
81. Acidalia perpusillaria, Ev., Bull. Mosce., 1847, ill,
peee pl. 6, f 7.
Given me by Lord Walsingham, February 1902.
82. Acidalia elongaria, Rmbr., Ann. Soc. Ent. Fr., 1838,
a6, pl. 2, f 20.
Tangier. March 1901.
83. Acidalia subsericeata, Haw., Lep. Brit., p. 352 (1809).
Forest of Marmora. March 1902.
390 Myr. E. G. B. Meade-Waldo on a
LASIOCAMPID.
84. Taragama repanda, Hiibn., Eur. Schmett., Bomb.,
ff. 274-5 (1827).
The larva of this species may be found in all stages of
growth all the year round, with perhaps the exception of
the months of August and September ; it varies remark-
ably in colouring, individuals being found in all shades of
grey and brown to black with violet dots. It will feed on
almost anything, but a prickly broom, Grenista tricuspidata,
appears its favourite food plant ; it is very abundant in the
neighbourhood of Tangier, and I also saw it at Tetuan and
Rabat.
85. Pachygastria trifolii, Schiff, Wien Verz., p. 57
(1776).
The same remarks practically apply to this larva, which
may be found universally in numbers all the year round.
It is much more abundant than the last, and varies greatly
in colour ; it will feed on almost anything.
86. Macrothylacia rubi, Linn., Syst. Nat., ed. x, p. 498
(1758).
Subsp. digramma, Meade-Waldo, subsp. n.
(Plate XIX, fig. 10, §.)
Deep red-brown: antenne ochreous-white above. Fore-wing
with fine oblique white antemedian line from subcostal nervure to
vein 1, very slightly exeurved below the cell: a fine white post-
median oblique line arising from vein 10, incurved to above 7,
excurved to vein 2, then straight to inner margin. Differs from the
typical form in the uniform dark chocolate colour, the white lines of
fore-wing being fine and farther apart, and the uniform chocolate
ciliz of both wings. Exp. 86-94 mm.
ITab. TANGIER.
I also took plenty of larvee at Babara. It frequents large
tracts of Zentiscws and ilex scrub. The larva feeds on
Pistacia lentiscus and Quercus ilex; it hibernates curled
up in December, spins a slight cocoon in March, and
emerges the beginning of April. Although carefully
sought for we never saw a male. I only saw one form
of larva, dark chocolate brown.
Collection of Moths made in Marocco, 1900-2. 391
87. Malacosoma franconica, Esp. Schmett., pl. 26, f. 1-2
(1784).
Tangier.
COSSIDA.
88. Zeuzera pyrina, Linn., Faun. Suec., p. 306 (1761).
Tangier.
PSYCHID.
89. Oreopsyche albida, Esp., Schmett., 11, p. 391, pl::78,
te a 187).
The variety lorquiniella, Bruand.
Common in the forest of Marmora, March 1902, flying
by day, and attracted apparently by the dead birds, ete.,
that were lying by us as we sat at lunch, hovering over
them and possibly mistaking their smell for that of the
female.
ZYGANID.
90. Zygena carniolica, Scop., Ent. Carn., p. 189 (1768).
The variety orana, Dup.
Laraishe, April 3rd, 1902.
91. Zygena favonia, Frr., Beitr. Kur. Schmett., v,
p. 76, pl. 428, f. 1 (1845).
Zygena aurata, Blachier, Bull. Soc. Ent. Fr., 1905, p. 213.
Two varieties (1) awrata mostly clothed with fulvous
to whitish hair, the abdomen with two crimson bands on
terminal segments, the fore-wing pale brassy golden yellow.
(2) favonia: thorax with few whitish hairs ; abdomen with
one crimson band, fore-wing blackish. Very abundant at
great heights on the Atlas, up to 12,400 ft., in July on
Tizi Gourza.
92. Procris orana, Aust., de. Nat. ii, p. 284 (1880).
Taken on grass at Imi Ouern on a branch of the Ouad
Amsmiz, July 11th, 1902, about 9,000 ft.
PYRALID.
ScHANOBIAN.
93. Scinpophaga prelata, Scop., Ent. Carn, No. 198
(1763).
Ras Doura, April Ist, 1902.
392 Mr. E. G. B. Meade-Waldo on Moths.
PHYCITINE.
94. Myelois cribrella, Hiibn., Eur. Schmett., Tin., f. 67
(1827).
Ras Doura. April Ist, 1902.
95. Acrobasis porphyrella, Dup., Lep. Fr., x, p. 191,
pl. 279, f. 2 (1836).
Forest of Marmora. March 7th, 1902.
PYRALINA,
96. Cledcolia interjunctalis, Guen., Lue, Expl. Alg.,
p. 398 (1848).
Ras Doura. April 1st, 1902.
HYDROCAMPIN&.
97. Nymphula stratiotata, Linn., Syst. Nat., i, p. 529
(1758).
Forest of Marmora. March 1902.
PYRAUSTINA.
98. Phlycteenodes paltealis, Schiff, Wien Verz., p. 123
(1776).
Forest of Marmora. March 1902.
99. Mecyna polygonalis, Hiibn., Vog., p. 76 (1827).
Forest of Marmora. March 1902.
100. Pyrausta flavalis, Schiff., Wien Verz., p. 121 (1776).
The Atlas.
Explanation of Plates. 393
EXPLANATION OF PLAtES XVIII, XIX.
Puate XVIII.
Sketch map showing Mr. Meade-Waldo’s route in Marocco.
PLATE XTX:
Fic. 1. Cononympha vaucheri, g, p. 377.
2 O peeks.
3. Satyrus miriszechi, var. maroccanw, ¢, p. 375.
4, ” ” ” ” 2 ’ p. 375.
5. Lycena hylas, var. atlantica, 3, p. 380.
6 ” bb} ” ” 23 p- 380.
7. Huchloé ewpheno, Q var. androgyne, p. 372.
8. Teracolus daira, var. nouna, go, p. 372.
9 ” ph 2, ” 2, p- 372.
10. Muacrothylacia rubi, subsp. digrumma, ? , p. 390.
11. Cleophana pauli, 9, p. 383.
12. Cleophana diffluens, var. maroccana, ¢, p. 384.
ve | Bee a ue ae
ite 1 * dosti)
Ci egnty)
XXII. A new species of the Hymenopterous Genus
Megalyra, Westwood. By J. CHESTER BRADLEY,
Ithaca, N.Y., U.S.A. Communicated by Col.
C. T. Binauam, F.Z.S8.
{Read November Ist, 1905. ]
I FIND among some material from the American Museum
of Natural History an undescribed species of Jegalyra.
In order that its position may be readily seen I append
a translation of a table to the genus published by
Szepligeti in the “ Természetrajzi Fiizetek,” xxv, p. 526,
adding the new species in its proper place.
1. Wings black, short, scarcely longer than the head above:
Mutilis, Westwood.
Wings not shorter than normal. ! : : ; (2)
2. Wings black or brown . : : : : : (3)
Wings hyaline, with brown scoters : : : ‘ (5)
3. Wings black, almost opaque, with a hyaline spot ; forehead and
vertex longitudinally grooved . . Shuckardiit, Westwood.
Wings more or less brown, without a hyaline spot; forehead
and vertex not longitudinally grooved : ; : (4)
4. Entirely black ; head, venter, legs, and propodeum heavily
clothed with grey hair; a hairy spot on the side of each
abdominal segment : ; Melanoptera, Schletterer.
Base of the antenne, legs beyond the femora, and the abdomen
red ; not heavily clothed with hair; the abdomen without
hairy spots . s : : : Rufiventris, Szepligeti.
5. The posterior ocelli further removed from each other than from
the compound eyes. ; (6)
The posterior ocelli equally Seibel ae iis gekipieitied eyes
and from each other . d : : é (7)
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.)
396 Mr. J. C. Bradley on a New Species of the
6. Entirely black ; scutellum coarsely punctured, smooth mesally ;
the first and a narrow margin on the second and third dorsal
segments smooth and ipa the rest shagreened by minute
puctulations . 3 : z Szepligetti, n. sp.
Dark brown with dee areas, the anterior legs beyond the
femora and the posterior beyond the tibiae reddish-brown ;
scutellum longitudinally wrinkled ; abdomen not sculptured
as above . : : : : 5 Caudata, Szepligeti.
7. Cheeks thickly and rather coarsely punctured ; terebra about
three times as long as the body. . Fasciipennis, Westwood.
Head equally punctured all over, finer than on the mesonotum ;
head red ; terebra four times as long as the body.
Longiseta, Szepligeti.
MEGALYRA SZEPLIGETIL, n. sp.
?. Entirely black. Slightly pubescent, the truncature of the
propodeum tomentose. Head globular, the occiput broad behind
the eyes, the posterior margin rounded ; antenne inserted below the
base of the eyes, a lateral oblique groove on each side marked by a
high ridge above receiving the scape when pressed downwards, and
separating the face from the cheeks and forehead ; head except face
rather coarsely closely not deeply, umbilicately reticulate-punctate.
Prothorax concealed ; mesonotum convex, produced laterally into
two anterior blunt angles ; dorsum punctured similarly to the head,
but punctures larger, a sub-smooth area in the middle of the
scutellum ; pleura more finely punctured, venter much more finely
and shallowly punctured ; distinct oblique grooves in the pleure
for the reception of the anterior and middle femora; propodeum
indefinitely shallowly reticulate; seen from above the posterior
margin is concave, the lateral angles sharp ; the posterior face of the
propodeum is concave, the abdomen fitting into it.
Posterior legs covered evenly with well separated punctures ; the
coxe and trochanters short, the femora stout, the single spine on
the apex of the tibia reduced to a mere rudiment, the metatarsus
longer than the other tarsal joints united, the fourth joint about one
half as long as the third ; middle and posterior and to some extent
the anterior legs covered with sparse long white hairs, about as long
as the third tarsal joint.
Wings hyaline, a broad brown band extending all the way across
at the stigma, a slightly dark spot at the apex ; the free part of M,
and the spur of the transverse part of M, wanting.
Abdomen sessile but constricted, the first dorsal segment and a
very narrow apical margin on the second and third smooth,
Hymenopterous Genus Megalyra. 397
impunctate, and polished, the remaining dorsal segments shagreened
by very minute punctulations,
Length 12 mm. ; of the head and thorax 6°5 mm. ; of the abdomen
55mm. ; of the fore-wing 9mm. ; of the ovipositor 34mm.
Hab. SoutH AUSTRALIA.
Type—1 $ in the collection of the American Museum
of Natural History in New York City.
A fine specimen of Megalyra melanoptera, Schletterer,
from New South Wales is also contained in that collection.
TRANS. ENT. SOC, LOND. 1905 —PART Iv. (DEC.) 27
| in [DS Meare
Palig W8 toy
4
Deere had” REG fut " a
ss ea 4
. # Munk Veet! w iy" ‘re Wi
d eo Guttsulles, taht f ti] ne v4 Hii hele a aie
( 399.)
XXIII. Hymenoptera aculeata collected in Algeria by the
Rev. A. E. Eaton, M.A., F.E.S., and the Rev.
Francis Davip Moricr, M.A., F.E.S. Part
II. Diploptera. By Epwarp SAUNDERS, F.RS.,
ete.
[Read November 15th, 1905. ]
JUGURTHIA NUMIDA, Sauss.
g. Biskra, in grounds of Chiteau Landon, 8.10 a.m., 24,
v, 93. (A. #. £.)
& » between Fort St. Germain and the brick kiln
on Ammi visnaga in the afternoon, 19, v, 97.
(A. HE. #.)
2 ; between Fort St. Germain and the brick kiln
on Ammi visnaga, in the afternoon, 18, vi,
SWRA es iw oc)
9. Médéa, Route d’Alger, north of town, on low plants,
such as Convolvulus ar VENSIS, Malva s ylvestr is
and Hchiwm, 26, vi, 93. (A. #. £.)
Fe aie as OOM towards Kef-el-azeri, on Daucus setifolius,
2, vill, 93. (C4. a.)
CELONITES FISCHERI, Spin.
®. Koudia Sma, a hill east of Médéa, alt. abt. 3,180 ft.,
on Microlonchus salmanticus, 15, vu, 93. (A. #. £.)
2. Constantine, visiting Hehiwm italicwm, “rolls itself
into a.pill and shams death,” 21, v, 95. (A. #. #.)
fg. Biskra, on Hehiwm humile, 19, v, 97, and 18, vi, 97
(A. #. £.)
QUARTINIA MAJOR, Kohl.
94. Sidi Ferruch, about sixteen miles from Algiers
westwards on the coast, on the glacis of the fort,
resting on flowers of Asteriscus maritimus, 8, v,
93. (A. #. £.)
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.)
4.00 Mr. E. Saunders on
QUARTINIA DILECTA, André.
? 1. Biskra, on Picridiwm tingitanwm, 3,iv, 97. (A. HL. E.)
EUMENES SICHELII, Sauss.
7? 1. Biskra, cornlands bordering the route des Zibans on
Ammi visnaga, Lam., 25, v, 93. (A. #. #.)
by 4; hollows between the railway near Kilométre
199, and the scrub bordering the pasture,
6.7, 07. (CAL BLE)
91. ,, scrub bordering the pasture, on Antirrhinum
ramosissimum, 13, v, 97. (A. #. £.)
2? 2. Tunis in the Cimetiére de Sidi-ben-Hassen, 20, xii,
93. (4. #. £)
EUMENES NIGRA, Brullé.
9 2. Road to Hamman-es-Salahin, opposite the new
Beni Mora racecourse, visiting Peganum harmala,
21, iv, 95. (A. #. #.)
EUMENES COARCTATA, F.
Various localities and dates from April to November.
(A. #. #. and F. D. M.)
EUMENES COARCTATA, F., var. dubius, Sauss.
2 1. Ravines, near Mount Ben Omar, north of Médéa,
alt. 2,730-3,130 ft., on Hryngium triquetrum, 11,
vil, 93. (A. #. #.)
2 1. Bone, on Mentha rotundifolia, 10, viii, 97. (A. #. £.)
EUMENES PICTETI, Sauss. var. (?)
91. Biskra, amongst Tamarix, 30, 1x, 97. (A. #. £.)
PSILOGLOSSA ALGERIENSIS, n. sp.
Nigra; rufo flavoque picta ; pronotum antice truncatum angulis
rectis ; post-scutellum sub-lamelliforme lateribus elevatis, apice
emarginato, lingua elongata ad coxas intermedias attingens, abdomen
segmento primo cupuliformi, margine apicali subelevato.
¢. Black, mandibles, clypeus, scape of antenne in front, and face
from just above the sinus of the eyes, except two short black divergent
lines originating at the insertion of the antennz flavous, flagellum
of antennx beneath, a spot behind each eye, the pronotum except at
Hymenoptera aculeata collected in Algeria. 401
its posterior angles, the tegule, a spot below the tubercles, the pos-
terior angles of the scutellum, and the raised margin of the post-
scutellum flavo-testaceous, abdomen with the Ist segment, except
a basal triangular spot, testaceous, the other segments black
with a trisinuate testaceous apical band, legs with the apices of the
femora, the tibie and tarsi rufotestaceous. Head closely and coarsely
punctured, mandibles tridentate, clypeus deeply and angularly
emarginate, apical joint of the antenne small, slightly curved and
subtruncate; thorax punctured very similarly to the head, prothorax
truncate in front, its angles not produced and nearly rectangular ;
wings slightly dusky especially along the costal margin, nervures
brown, clear testaceous towards the base of the wings ; 2nd recurrent
nervure received near the base of the third submarginal cell ;
post-scutellum flattened and produced horizontally over the pro-
podeum, slightly emarginate posteriorly, its margins reflexed, a
strong carina runs along the sides of the propodeum, below which
the surface is bright and shining ; abdomen strongly punctured
lst segment cupuliform, its apical margin slightly raised.
9. Very like the ¢ and similarly punctured, but with the base
and under-side of the antenne, the mandibles, the head posteriorly,
and a spot in the sinus of each eye rufotestaceous ; on the face, its
apex between the insertion of the antenne, is a triangular spot of a
paler testaceous colour, clypeus only slightly sinuate at its apex,
pronotum truncate with its angles more produced than in the ¢, and
if anything slightly less than a right angle, flavotestaceous in front
blending through a brownish colour into black posteriorly ; wings as
in the ¢. Sides of the propodeum rufotestaceous, abdomen with the
basal segment rufotestaceous, its apex of a more flavous tint, its
disc with a round blackish spot, the rest of the segments black,
with a testaceous trisinuate apical band, more or less margined with
rufous, apex of terminal segments entirely testaceous ; legs as in
the ¢.
Long. 7-8 mm.
g 1. Biskra, on Antirrhinum ramosissimwm, 21, ii, 97.
(A. #. #.)
SBP 3 022, mep OG AG ee sy
22. » on Abractylis serratuloides, 10, v, 97.
(A. EH. E.)
an ip » on Deverra chlorantha, 13, v, 97. (A. #. £.)
Apparently allied in colour to P. pulchra, Mor., but
differing in the shape of the pronotal angles and that of
the lamelliform post-scutellum.
4.02 Mr. E. Saunders on
RAPHIDOGLOSSA RUFESCENS, n. sp.
Rufotestacea, nigro-variegata, clypeo, abdominisque segmento
primo et secundo rufotestaceis primo fascia apicali nulla, ? clypei
marginibus elevatis, nigris ;
Rufotestaceous, head and thorax largely and closely punctured,
the former black with the clypeus, a large somewhat triangularly-
shaped spot, its base situated just below the anterior ocellus and its
apex just above the insertion of the antenne, and a large spot behind
each eye rufotestaceous, mandibles in the centre and antenne at the
base of the same colour, mandibles with four blunt teeth, elypeus in
the ¢ bidentate, with a small, deep emargination between the teeth,
? with the free sides of the clypeus widely reflexed, shining and
black, the apical margin which lies between them slightly emargin-
ate, with two small prominent teeth near the centre. Thorax above,
black only on the mesonotum (in one specimen entirely red),
and the base of the scutellum and post-scutellum, beneath black
with the exception of a red spot on the mesopleura. Pronotum
truncate in front meeting the sides in angles slightly less than right
angles, the sides slightly sinuate immediately behind the angles
which are consequently somewhat produced, post-scutellum truncate
posteriorly wings slightly smoky especially towards the costa,
veins rufotestaceous, 2nd submarginal cell sub-triangular, very
narrowly truncate at its upper margin ; propodeum black in the
centre, red and clothed with silvery hairs at the sides ; abdomen
dull, finely punctured and clothed with very short white pubescence,
with the basal and 2nd segments entirely rufotestaceous, the
former strongly punctured, with a large round impression near the
apex, and in some examples with the latter more or less suffused with
black, following segments black, their posterior margins more or
less rufous, sometimes widely so, the colour band of a more or less
bisinuate form ; legs entirely rufotestaceous except the anterior coxx
in front.
Long. 15-16 mm.
¢ 2. Biskra, near railway, kilométre 199, “asleep on
Deverra scoparia, 25, v, 94, holding on to the extreme tips
of the stems by the mandibles only, with wings folded
longitudinally and widely spread, the legs tucked up close
to the body not touching the stems, which waved about in
the breeze.”
d 9. Biskra; same locality on Zizyphus lotus, 21 and 29,
iv, 95.
Hymenoptera aculeata collected in Algeria. 403
3 2,25. Biskra, on Atractylis serratuloides and Antir-
rhinum ramosissimum, 6-13, iv, 97.
g 2. Biskra, on Amberboa lippii, 8, iv, 97.
Several of the above specimens are stylopized (Xenos?).
PTEROCHILUS ATERRIMUS, 0. sp.
Ater, atrohirtus, punctatus, mandibulis, capite utrinque pone
oculos, et in femina interdum clypeo et oculorum orbitis interioribus,
tibiis anterioribus antice, articuloque apicali palporum labii, rufo-
ornatis ; alis valde infumatis.
Black, densely clothed with long black hairs, especially in the ¢,
mandibles more or less red, generally nearly entirely so, but in one 9
with only the extreme apices of that colour, a spot of variable size
behind!each eye, and in one female two large spots on the clypeus and
the inner orbits of the eyes red. Anterior tibie in front in both sexes
and the much curved, flattened apical joints of the labial palpi of
the same colour, wings dark brown with purple reflections. Head
and thorax rugosely punctured, somewhat shining, anterior margin
of the clypeus narrowly emarginate in the ¢ semicircularly rounded
and entire in the ? ; apical joint of labial palpi in the ? flat and
dilated, fringed with very long hairs, in the ¢ sub-cylindrical with
hairs as long as those of the 9, but finer and less regular. Antenne
in the ¢ slightly rolled at the apex. Penultimate and apical joints
very sinall, especially the latter, posterior wings clearer than the
anterior ; legs shining, punctured, the inner sides of the first and
third pairs of tibiaee densely clothed with fine pubescence in both
sexes ; propodeum with an angular tooth on each side; abdomen
nigro cyaneous, largely punctured on its basal segment which bears
a slightly raised tubercle in the centre of its basal brow, finely
punctured on the rest, first and second segments in the ¢ first only in
the 2 clothed with long hairs, apical margins of the ventral segments
with a row of exceedingly short bristles.
Long. 14-16 mm.
Biskra, 2 1, amongst hummocks of Limoniastrum guyo-
nianum on the Dunes by the route des
Zibans, 19, ii, 94.
2 1, visiting Brassica napus, 8, ii, 97.
gf 1, between Biskra and Hamman-es-Salahin.
g 3, at rest on dead twigs of Demia cordata at
5 p.m., holding on by the mandibles and
legs; wings divergent, 14, ii, 97.
404 Mr. E. Saunders on
PTEROCHILUS GRANDIS, Lep.
9 Médéa, about a mile along the road to Lodi 12.30 to
2.30 p.m., on Hryngium triquetrum, 28, vi, 93. (A. #. £.)
2 1, hill north of railway station, alt. 3,000 feet, 10,
vii, 93. (A. #. £.)
® 1, ravines, near Mount Ben Omar, on Lryngium
triquetrum, alt. 2,730—8,130 ft., 11, vii, 938. (4. #. #.)
2 1, on Koudia Sma, a hill to the east, of an altitude of
about 2,700 ft., about 4 p.m., 15, vii, 93. (4. #. £.)
PTEROCHILUS COCCINEUS, André.
2 1. Fontaine Chaude, near Biskra, on the dunes near
the stream among Limoniastrum guyonianum, 3 to 5 p.m.,
16, v, 94.
PTEROCHILUS LINGUARIUS, Nn. sp.
Niger albo et ferrugineo variegatus, lingua longissima ad apicem
abdominis extensa, abdominis segmento primo cupuliformi apice
elevato, ‘secundo margine posteriore membranaceo, ad basin seri-
atim punctato-impresso.
Black with white and ferrugineous markings. Head black, shin-
ing and strongly punctured, mandibles, scape of the antenne and
the flagellum beneath, testaceous, the latter darker on its upper-side ;
Clypeus in the male flavous, in the 9 sometimes with a testaceous
spot, semicircularly emarginate in the ¢,in the ? with its free sides
reflexed, face densely clothed with silvery hairs in the ¢@, sparsely
so in the 2. Tongue very long reaching in repose to the apex of the
abdomen, labial palpi slightly flattened and plumose in the 9, nearly
simple in the ¢: behind the eye in both sexes is a very small whitish
spot, antenne in the ¢ with a sharp reflexed apical joint thorax
black, shining, rather remotely punctured pronotum except at its
extreme basal angles, tegule, a spot on the mesopleura, scutellum
except a triangular patch at its base and post-scutellum, ivory white ;
prothorax truncate in front, its angles nearly rectangular, legs tes-
taceous, the femora darker at the extreme base, propodeum clothed
with silvery white hairs, excavated posteriorly and with a long sharp
spine at each side of its extreme apex especially in the ¢ ; on each
side there is a testaceous spot of variable size ; abdomen sparsely
clothed with silvery hairs, largely and remotely punctured especially
on the basal segment which is campanulate in form, its apical margin
raised and rounded, entirely testaceous or with its base more or less
suffused with black, the raised apical ring ivory white, 2nd segment
black with its apical margin flavous the colour spreading at the sides
Hymenoptera aculeata collected in Algeria. 405
and sometimes produced on to the disc in the form of two spots, the
sides of the segment are generally more or less brown, the apical
margin of this segment is produced into a rather wide membrane
bearing a row of punctures at the base especially noticeable in the
d, the following segments are black or piceous with pale apical
bands, apical segment black.
Long. 7-8. mm.
f1. Fontaine Chaude, near Biskra, on the dunes near
the stream, 9, v, 94. (4. #. £.) :
£1,292. Biskra, Hamman-es-Salahin, on the sandhills,
avy ols (A BAB)
93. Biskra, 20 and 31, v, 98. (F% D. IL)
A small species superficially like a small Chevrieranus
but easily distinguished by its extremely long tongue, a
character which almost warrants the erection of a new
genus for its reception.
PTEROCHILUS AFFINIS, 0. sp.
Niger flavopictus, clypeo maris emarginato, femine punctato,
sine rugis longitudinalibus, apice truncato, alis subluteis, segmentis
ventralibus maris 3-5 dense aureofimbriatis, segmenti secundi
feminze disco valde sed sparse punctato.
Black with the following parts yellow, mandibles except their
piceous teeth and apices, the entire clypeus in the ¢, the base
only in 2, a spot between the antenne and a spot behind each eye
and another in the sinus of each in both sexes, and the scape of
the antenne in the ¢ a somewhat triangular spot on each side of
the prothorax not reaching either the anterior or posterior angle
a spot below the wing, the tegule, two spots on the scutellum, the
post-scutellum and two small spots on the propodeum, the borders
of the abdominal segments more or less trisinuately except that of
the basal which is straight, and the apical, which has a central spot
only ; beneath with a bisinuate band on the second and following
segments, except the apical ; all the above markings are the same
in both sexes ; legs testaceous-yellow, coxe, trochanters and bases
of the femora black. Head and thorax dull, closely punctured,
clothed with long hairs, brownish-red in the ¢, nearly black in
the 9. Clypeus semi-circularly emarginate in the 2, narrowly
truncate in the 2, spot between the antenne in the ¢ slightly
raised, wings slightly brownish especially along the costa, the
nervures testaceous-red. 2nd submarginal cell, receiving both
recurrent nervures ; posterior tarsi with their 2nd and claw-joints
4.06 Mr. E. Saunders on
subequal in length, the 4th joint about two-thirds as long as the
third. Abdomen somewhat shining, very densely and microscopic-
ally punctured, the basal segment sparsely clothed with rather long
brownish hairs, 3rd, 4th and 5th ventral segments in the ¢ with
a dense fringe of short golden hairs.
Long. 13-14 mm.
Bone, f 1. 28, iv, 96. (A. #. £.)
Alger, 2 1.29, iw,.98.;5CBo) Dial)
Apparently closely allied to hellenicus of Morawitz, but
the @ differs in having the clypeus destitute of longitu-
dinal rugosities, in having a central spot between the
antennz instead of two side by side, and in having the
second segment of the abdomen beneath, with larger
punctures scattered on the disk and without the larger
punctures on the apical margin. The g of hellenicus is
not described. The head and thorax are also clothed with
longer hairs than in hellenicus.
OpDYNERUS (/oplopus) LUTEOLUS, Lep.
Gf 2. Médéa, ravines near Mount Ben Omar, alt. 2,730-
3,130 feet, 8 to 10 am., on Hryngium triquetrum, 11, vii,
93. (A. #. #.)
g 1. Constantine, 16, vi, 98. (FD. I,
ODYNERUS (Hoplopus) SPIRICORNIS, Spin.
9 1. Constantine, on M’cid, 21, vi, 94. (A. #. #.)
Pe un OR Wowk
ODYNERUS (Hoplopus) CONSOBRINUS, Duf.
g 2. Algiers, 14 and 17, ii, 93. (4.4. #,)
g 1. Biskra, about walls of “adobe,” village Negre,
13,1, 94.. (A. #. #.)
g 1. Biskra, visiting Moricandia arvensis, 3, ii, 97.
(A. #. FE.)
? 1. Biskra, 15, ii, 94. (4. #. #)
91. Bone, host of Phileremus oraniensis, 2, iii, 96.
(A. #. #.)
3 3,29. Aigiers, March 1898. (£. D. IL)
ODYNERUS (Hoplopus) CAROLI, Mor.
g 2. Bone, footpath and river-bank by the Bon
Djemma, 24, iii, 96. (A. #. £.)
Hymenoptera aculeata collected in Algeria. 407
g 1., Alger, 27, iv, 98. (#.D. IL)
(2) 2 1. Constantine, on M’cid, 22, v, 95. (A. #. £.)
(2?) 21. Alger, 27, iv, 98. (F. D. dL)
The ? of this species is known at once by the spine at
the base of the anterior femora (beneath), The ? which
I have assigned to it agrees with it in size and general
coloration, and has the post-scutellum rather sharply
elevated as inthe #. It lacks the long hairs of conso-
brinus and rotwndigaster, but has the yellow band of the
second abdominal segment dilated at the sides as in those
species, although more distinctly so. The scutellum and
post-scutellum are entirely black.
ODYNERUS (Hoplopus) DECEPTOR, n. sp.
O. variegato simillimus, ab illo mas coxis intermediis nigro
spinosis facile distinguitur, femina, angulis propodei subproductis,
segmento secundo abdominis latiori subtus testaceo, minus crebre sed
obsoletius punctato aegre distinguenda.
Resembling O. variegutus in colour but at once recognizable by the
black spines on the intermediate cox of the ¢ and the very largely
laterally produced bands of the abdomen which on the Ist and 2nd
abdominal segments leave only the central basal portion black. In
the 2(?) of this species the lateral angles of the propodeum although
not acute are more produced than in variegatus, the 2nd abdominal
segment is wider in proportion to the lst and on its under-side is
testaceous, more shining than in variegatus less closely and more
obsoletely punctured, but only having one example of this 9 I am
unable to say how far these characters are constant.
$. Head and thorax black, closely punctured, dull, clothed rather
sparingly with greyish-brown hairs, clypeus yellow, shining, deeply
and semicircularly incised at the apex, labrum yellow, mandibles
black, spotted with yellow at the base, a somewhat straight yellow
spine projects from the gena on each side of their extreme base ;
this spine is slightly curved towards the apex and viewed laterally
appears nearly truncate, a transverse tubercle between the antenn»
and the scape in front testaceous, spire entirely black, pronotum with
a ferruginous band not extending to its posterior angles, anterior
angles slightly produced and sharp, tegule ferruginous, wings smoky,
especially along the costal margin, scutellum slightly raised, post-
scutellum nearly perpendicular, ferruginous, slightly concave and
very shining, sides of the propodeum obtusely angled, its concavity
dull, diagonally striate, widely channelled, with a narrow longi-
tudinal carina running along the bottom of the channel—legs
408 Mr. E. Saunders on
testaceous femora black at the base, intermediate cox each bearing
a black, slender, nearly straight spine. Abdomen slightly shining
very finely rugulose with the first segment clothed with hairs as the
thorax, the rest glabrous, all the segments bearing broad bisinuate
apical testaceous bands, the bands of the 1st and 2nd segments so
produced at the sides towards the base, as to leave a square black
central emargination, segments beneath very finely and closely
punctured, 2nd with a free testaceous spot on each side in some
specimens, often wanting.
@ (2) The female which I associate with the ¢ has exactly similar
bands on the abdomen, and is very similar otherwise, with the
exception that the scutellum is ferruginous except at its extreme
base and that the sides of the propodeum have the angles rather
produced but obtuse and widely yellow and the 2nd segment of the
abdomen beneath is nearly entirely ferruginous, shining, and rather
obsoletely punctured, there is also a testaceous line on the lower
internal orbit of the eye and a large spot behind nearly on the vertex
scarcely indicated in the ¢ and a spot on the mesopleura under
the wing.
Long. 10-12 mm.
gf 4. Bone, 20, iv, 96. (4. # &.)
é 3. ,, foot-path and river bank by the Bon
Djemma, 24, ui, 96. (4. #. £#)
2 1. Médéa, hill north of railway along the edge of the
enclosure on the west side, on Hryngiwin
triquetrum, 27, vi, 93. (A. #. £.)
ODYNERUS (Hoplopus) VARIEGATUS, Fab.
° 4, Constantine, 15, 16, vi, 98. (# D. dL)
OpyYNERUS (Hoplopus) ANGUSTIOR, Nn. sp.
Angustus, parce griseo-hirsutus niger, scapo antennarum antice
flavo, flagello toto nigro, clypeo, tuberculo transverso inter antennas,
prothoraceque antice flavis, scutello nigro, immaculato, post-scutello
flavo, plano, sub-levi, basi solo punctato, abdominis segmentis
apicibus late flavo-fasciatis, subtus segmento secundo crebre et minute
punctato, punctis majoribus intermixtis.
d- Black, clypeus, labrum, a line along the anterior lower orbit of
the eyeand a spot behind theeye yellow, antenne with theseape in front
yellow, otherwise entirely black. Thorax with the anterior margin
of the pronotum, the colour widened at the sides but not produced
to the posterior angles, the tegule and post-scutellum testaceous,
abdomen with a wide, angularly emarginate band on the basal
Hymenoptera aculeata collected in Aigeria. 409
segment, a wide nearly straight band on the 2nd and a very slightly
bi-sinuate band on the 3rd to 6th testaceous, These bands are
all continuous on the ventral segments. Legs testaceous yellow,
base of the anterior femora above, about three-fifths of the inter-
mediate femora above, and the posterior femora, except at the apex,
black.
Head and thorax closely and rugosely punctured, clothed with short
greyish white hairs, clypeus deeply and semicircularly emarginate,
pronotum rather produced at the sides and sharply angulated, post-
seutellum shining, propodeum very declivous, its sides rounded, not
much excavated posteriorly, the sides of the excavation irregularly
and diagonally striate, wings-nearly hyaline, but smoky along the
costal margins, nerves and stigma dark brown, abdomen coarsely and
irregularly punctured on the basal segment, finely but irregularly on
the second, finely closely and regularly on the rest, beneath closely
and finely punctured throughout, and with a larger (but still rather
fine) puncturation scattered over the surface—observable chiefly on
the 2nd and 3rd segments—legs simple.
Long. 9 mm.
Aslenderer species than usual but bearing no very marked
characters, its entirely black flagellum combined with the
wide abdominal bands separate it from any other species I
can find described.
gee iskra, 215 it, 95," (Can ee 2)
ae 23, 11, 97 on Tamarix. (A. #. Ff.)
at) Bone? - Tv 9G. Ce EE.)
ODYNERUS (Ancistrocerus) ATROPOS, Lep.
f? 2. Biskra, sandy banks bordering the fallow
near the market garden by the river and
railway near kilométre 199, 17, ii, 94,
ft 4,2 4. Biskra, near the Barrage, 22, 11i—iv, 97.
“Makes a curved. tubular portico to its
burrow with irregular elougate longitudinal
loopholes ; tube 18 mm. long and 3°5 mm,
external diameter.”
In one $ there are 6 submarginal cells in the right
wing and 4 in the left, all the other specimens have the
normal 3. <A character observable in the 2 which does not
appear to have been noticed in descriptions is that the
clypeus is practically 3-dentate—as there is a very small
tooth in the centre of the emargination.
410 Mr. E. Saunders on
ODYNERUS (Ancistrocerus) PARIETUM, Linn.
Various localities; Biskra; Bone; Algiers; Azazga,
alt. 1,900 ft. (4. # #. and F. D. IL)
ODYNERUS (Lionotus) CHLOROTICUS, Spin.
© 9) Biskra, 31, v; 98: “(De M)
ODYNERUS (Lionotus) EGREGIUS, H. Sff.
O10 Biskray 22 av O72 Cae ee)
OUP tents og hy AOR eee eee)
21. Médéa, 28, vi, 93. (4. #. #.)
ODYNERUS (Lionotus) SAUSSUREI, André,
21,92. Biskra, on Ammi visnaga, 18-22, vi, 97.
ODYNERUS (Lionotus) DANTICI, Rossi.
gf 2. Biskra, on Ammi visnaga, 18, v, 93. (A. #. £.)
2 . Tizi-Ouzou,on Zizyphus lotus, 14, vi, 93. (A. L. £.)
2. Constantine, on Ferula scabra, 22, v, and 2,
vi, 95.. (A. # £)
ik, he on Zizyphus lotus, 15, v, 97. (A. #. #.)
1. Bone, 25, vu, 97. (A. #. £.)
2; PG. Biskra, 13, v-10, vi, 98. (#. D. I)
1. Philippeville, 20, vi, 98. (#. D. M)
ODYNERUS (Lionotus) BIDENTATUS, Lep.
¢@ 1. Biskra, railway, near kilométre 199, 31, iii, 94.
(A. #. E.)
a IP i Fontaine Chaude, on Tamarix paucé-
ovulata, 19, iv, 94. J.Gay MS. (4. # #.)
Bul. is ridge of hills north-east of Hamman-es-
Salahin, visits Serula vesceritensis,
5, 1V, 95.0) (Ane)
be, ky visiting Antirrhinum vranosissimum
21,11, 07 Arn es
el 5 on Moricandia arvensis, 12, iv, 97
(A. EB)
Hymenoptera aculeata collected in Algeria. 411
ODYNERUS (Lionotus) SPECULARIS, n. sp.
O. bidentato colore assimilis minus hirsutus, post-scutello arcte
elevato, crista media subdentata, postice perpendiculari, nitidissimo,
propodeo linea dorsali ad basin explanata nitidissima, lateribus
utrinque bi-angulatis, squamis magnis subquadratis.
Group of dantici and allied to bidentatus, Lep.
¢ - Black, clothed with short grey hairs, those on the head and front
of the thorax rather longer. Head and thorax rugosely punctured,
clypeus and labrum flavous, punctured, the former deeply and semi-
circularly emarginate, clothed with silvery pubescence, mandibles
testaceous, brown at the apex, a spot between the antenne, the lower
anterior orbits of the eyes, a spot behind each eye, and the first two
antennal joints flavo-ferrugineous, apical joints of the antenne
including the recurved hook testaceous, at least beneath ; pronotum,
except its posterior angles, the tegule, two transverse spots along
the apex of the scutellum, and a spot on each side of the propodeum
along the lateral margin yellow, anterior margin of the pronotum
very slightly produced in the centre, its angles slightly prominent
and rectangular, tegule punctured, wings including nerves anteriorly
and on the apical half, suffused with brown colour becoming yellower
towards the base, post-scutellum sharply elevated with a shining
semi-circular area on its perpendicular posterior side, scarcely
erenulate, but with a slight central tooth, propodeum angulated at
the sides with a strong emargination beneath the larger angle,
depression rugosely punctured, but with a very bright shining sub-
triangular central space meeting the shining area of the post-scutellum,
apical scales very large and subquadrate ; abdomen thinly clothed
with fine grey pubescence, giving it a greyish look in some lights,
tugosely punctured, all the segments with broad yellow trisinuate
apical bands, that on the first segment very wide, only leaving a
sub-triangular black spot at the base; beneath shining, strongly
punctured, with a band on the second and third segments only, basal
crenatures of the second segment long and well developed, legs yellow,
coxx and trochanters black, all the former spotted with yellow in front.
Long. 13-14 mm.
¢@ 1. Biskra, to the north of railway, kilométre 199,
4.30-6 p.m., on Teucriura polium, 28, v,
94. (A. #. £.)
$1. 4, on Echinops spinosus. (A. EL. £.)
I have described the bands, etc. of this species as
yellow, as they are evidently discoloured in both my
examples.
412 Mr. E. Saunders on
ODYNERUS (Lionotus) BLANCHARDIANUS, Sauss.
ahs Tart) 27, vi, 26:1 A eee)
f 2. Biskra, on Ammi visnaga, 15, 17, v, 97. (A. £. £.)
f 1. Bone, on Luphorbia paralias, 7, vii, 97. (A. £. #.)
fl. 4, 4, Mentha rotundifolia, 10, vii, 97. (A. #. £.)
ODYNERUS (Lionotus) sp. (?)
9 1. Biskra, on Tamarix, 28, iv, 97. (A. #. #,)
ODYNERUS (Lionotus) sp. (2)
2 1. Bone, 10, vi, 96. (4. #. £.)
ODYNERUS (Lionotus) TRIPUNCTATUS, Fab.
© 1. Biskyra, 23; 4; 9f.cu( Ay Bad)
2 1. Biskra, on Amma visnaga, 24, v, 97. (A. #. £.)
ODYNERUS (Lionotus) REGULUS, Sauss.
? 3. Bone, on Luphorbia paralias, 7 and 23, viii, 97.
(ALE)
a1. 4 ,, Bubus discolor, 13\-vin, 97. (A. 2. 2.)
f1. 4, 5, Mentha rotundifolia, 10, vii, 97.. (A. #. £.)
Ol. ,, ,, Juncus acutus, 25, vii, 97. (A. #. £.)
f 1. La Calle, 4, vii, 96. (A. #. Ff.)
d 1, Biskra, ‘7, 1v, Of: (CA)
ODYNERUS (Lionotus) EATONI, n. sp.
Citrinus, glaber, fortiter punctatus, capitis vertice, mesonoto,
sternis, et propodei postice macula centrali nigris; abdominis
segmentis primo et secundo nigro maculatis, tegulis magnis, valde
punctatis.
Bright citron yellow, glabrous, very largely punctured, head in
both sexes with only the vertex black, clypeus narrowly emarginate
and bidentate, antennz sometimes darker towards the apex,
apical recurved joint in the ¢ very small; thorax with the
mesonotum, except a quadrate spot in front of the scutellum, and
the sterna black, propodeum black with its sides yellow, tegule
large very strongly and closely punctured, wings very slightly
dusky rather more so along the costal margin, nervures brown, post-
scutellum slightly raised and crenulated, sides of the propodeum
rounded and crenulated, apical scale produced above into a long sharp
Hymenoptera aculeata collected in Algeria. 413
spine ; abdomen closely but not quite so coarsely punctured as the
head and thorax, posterior margin of the Ist segment very slightly
and of the 2nd very distinctly raised, a spot on the apical margin
of the first segment, a transverse spot on the disk of the 2nd towards
the apex black, apical segments sometimes more or less infuscate ;
beneath with the 2nd segment convex, its basal costs rather long
but not very strongly developed, legs with the cox at the base
black.
Long. 8 mm.
A very distinct species by the strongly punctured
tegule, the peculiar coloration, somewhat resembling
chloroticus, and the long spines of the apical scales of the
propodeum.
$1, 24, Biskra, cornlands bordering the route des
Zibans, on Ammi visnaga, 10.30 am. and
1-3 p.m., 18 and 24, v, 98. (A. #. £.)
¢ 2. Biskra, outskirts of the oasis, 17 and 19, v,
Of (ALE ES
ods. 5... Biskra,»5, v=9, vi, 98. (Ch. D117)
ODYNERUS (Lionotus) DOURSH, Sauss.
¢ 1. Constantine, visitng Marubium vulgare, 29, v, 95.
(A. EB. EB)
ODYNERUS (Lionotus) PONTEBH, Sauss.
f 1. Médéa, 28, vi, 98. (4. # #)
g¢ 1. Constantine, 18, vi, 98. (/ D. IL)
ODYNERUS (Lionotus) PARVULUS, Lep.
Numerous specimens of both sexes from Biskra, Le Tarf,
Hippone, Bone, frequently on Ammi visnaga, iv—
vill. (A. #. #. and & D. J.)
ODYNERUS (Lionotus) BISPINOSUS, Lep.
9 2. Biskra, 28, v, 3, vi, 98. (A. #. #.)
gl.” Algiers) 13) 1v,798, (F. D! AL)
2 1. Constantine, 17, vi, 98. (/ D. I)
2 1. Biskra, 24, v, 98. (/ D. I)
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.) 28
414 Mr. E. Saunders on
ODYNERUS (Lionotus) HANNIBAL, Sauss.
¢@ 1. Constantine, entering a snail shell “on busi-
ness,” 16, v, 95. (A. #. #.)
df 2,91. Biskra, visiting flowers of Tamarix brachystylis,
4 and 8, iv, 95. (A. # #)
24,94. 4, visiting flowers of Tamarix brachystylis,
29, in, 18,40, Or (a, Ae
Ovens fp SLD eo oa ee ir)
2 1. Constantine, 18, vi, 98. (Ff. D. IL)
ODYNERUS (ZLionotus) ALPESTRIS, Sauss.
2 1. Biskra, Fontaine Chaude, 4, iv, 94. (4. #. #.)
Sa hs visiting Moricandia arvensis, 25, ii, 95.
(A. #. £.)
Oran e § visiting Antirrhinum ramosissimum, 1
and 21, 11,97. (4. #. #)
21. Bone, visiting Huphorbia paralias, 23, viii, 97.
(A. #. £.
ODYNERUS (Lionotus) FLORICOLA, Sauss.
92. Bone. 6, vii, 96, and 29, vii, 97. (A. #, #.)
$9. Algiers, 22 and 30, iv, 98. (/ D. JL.)
G1. Le Tarf. 26, vii; 965 (42s
ODYNERUS (Lionotus), sp. (2)
G1. Biskra and Oued Biskra, above the Barrage,
amongst Z'amarix scrub, 4,iv, 97. (A. #. £.)
This male is peculiar in having the claw joints of the
intermediate and posterior tarsi black.
ODYNERUS (Lionotus), sp. (?)
9 2. Biskra,a little above the Barrage, amongst Tamarix,
30, iv, O0.* CA, eee
nL , between the railway and the scrub border-
ing the pasture, 10, v, 97. (A. #. £.)
This may prove to be the 2 of the preceding, but the
tarsi are entirely pale, and both my specimens are much
discoloured; they are very closely allied to floricola, but
the post-scutellum is slightly raised and the mesopleurz
are not carinated under the yellow spot as in that species.
-
Hymenoptera aculeata collected in Algeria. 415
OpYNERUuS (Lionotus), sp. (?)
? 1. Constantine, on Jnwla viscosa, 9, x, 93. (A. #. E.)
9 1. Philippeville, 20, vi, 98. (Ff. D. J.)
Apparently only differing from the preceding by having
the pronotal emargination narrower.
These three species are so closely allied that I do not
feel that it would be safe on the material before me to
describe them as new.
ODYNERUS (Microdynerus) PEDUNCULATUS, 0. sp.
Niger, flavo rufoque ornatus ; segmento primo abdominali maris
petiolato (Eumeniformi), foeminz forma consueta, in utroque sexu
plus minus rufotestaceo, apice flavo fasciato,segmento secundo nigro
fascia apicali pallida, ipso apice membranaceo, punctorum serie
impressa,
Black, head and thorax shining, strongly punctured, face entirely
clothed with silvery hairs in the ¢ ; mandibles piceous in the ¢,
fulvo-testaceous in the ? , clypeus white in the ¢ except at the apex.
Scape of the antenne in front white in the ¢, fulvo-testaceous in the
¢, recurved apical joint in the ¢ testaceous, terminating in a black
knob, reaching to the base of the 11th joint, which is testaceous
beneath, the very short 12th joint being entirely of that colour.
Anterior margin of the pronotum truncate, with an entire narrow
pale band in the ¢, two spots in the 9, angles nearly right angles,
sides of the pronotum nearly parallel, very long in the Q ; tegule
whitish in the ¢, fulvo-testaceous in the 9, wings slightly smoky,
darker along the costa and in the radial cell, scutellum nearly square
in the 2 and fulvo-testaceous at the apex, transverse in the ¢, with
two apical pale spots, post-scutellum black in the ¢, concolorous
with the scutellum in the 2, propodeum black with a strong central
impression, its sides clothed with silvery hairs, the impression
shining in the @, sides of the propodeum gradually converging
towards the apex in the male and terminating in two rather long
pale spines which somewhat embrace the petiole of the body, sides
much more rounded in the 9, the spines short, and further apart
not conspicuously pale, but piceous in colour, propodeal scales small,
not spinose ; abdomen slightly shining, punctured and clothed with
greyish, very fine pubescence, with the basal segment fulvo-testaceous
in both sexes, very narrow and petiolate in the ¢, of ordinary form
in the 2, the extreme apex with a pale whitish band, in the ¢ the
base of the segment is black, and the apical margin is raised, callose,
416 Mr. E. Saunders on
and densely fringed posteriorly with very short fine white hairs,
second segment with a pale apical band beyond which is a mem-
branous appendix, which in the ¢ bears a row of elongate
punctures ; beneath with the second segment more shining than the
rest and strongly punctured, with the apex pale as on the upper-side.
Legs fulvo-testaceous, paler in the ¢, extreme base of the femora
piceous.
Long. 7-8 mm.
91. Biskra, near railway, kilométre 199, in the scrub
between the fringe of reeds and the farm mule-track,
DOA OTs p(t Lede)
$1,¢91. The same place, 25, iv, 97. (A. #. £)
The ¢ and 2 are so different in form that I feel great
diffidence in uniting them, at the same time their colours
are so similar and the locality identical that I hope I
have done right; the 2 is of the ordinary form of a
Microdynerus, but the $ more resembles those species of
the minutus group, with constricted petioles, cruentatus,
eatont, etc., but is much more elongate.
POLISTES.
Of this widely-distributed genus, numerous specimens
have been brought home by Messrs. Eaton and Morice,
all of which belong to the gallica group—but the
majority are females which in the present state of our
knowledge cannot be referred for certainty to their
respective males. Dr. F. F. Kohl (Ann. d. K. K. Naturhist,
Hofinuseums, Wien, Band xii, 1898, p. 87, taf. 111)
describes and figures the males of five species of this group,
which appear to be easily distinguishable, and of these
Mr. Eaton has brought home two.
22 are recorded from Biskra, Bone, Médéa, and Mustapha
Supérieur. (A. #. #. and F. D. M.)
PoLISTES DUBIA, Kohl.
2? 2. Biskra, in the grounds of Chateau Landon, and in
the fields between that and The Negro Village, on Amma
visnaga, 24, v, 93 ;“ eyes” in life “ subolivaceous.” (A. #. £.)
POLISTES FG@DERATA, Kohl.
g1. Bone, near the Plage, 28, xi, 93. (A. # £.)
Hymenoptera aculeata collected in Algeria. 417
VESPA GERMANICA, F.
gf 1. Constantine, 12, xi. 94.
9 1. Mustapha Supérieur, 25, 1, 93.
01. Colonne Voirol, 4, v, 98.
O 1. Médéa, 1, vu, 93.
This is the only species of Vespa sent home.
g% ue
a a Ven ws
i
( 419 )
XXIV. On the Ichnewmonidous Growp Tryphonides schizo-
donti, Holmer., with Descriptions of New Species,
By CLAUDE Mor ey, F.E.S.
[Read December 6th, 1905.]
THESE injurious parasites have upon several occasions
come before the notice of our Society. Thomas Desvignes
published (Trans. Ent. Soc., 1862, pp. 215-222) descriptions
of ten supposedly new species of the genus Bassus, unfor-
tunately with no knowledge that Holmeren had previously
brought out (Sv. Ak. Handl.,1855, pp.353-371) an elaborate
revision of the group, wherein all such species, indicated
by the former, as had been unknown to Gravenhorst (Ichn.
Europ., 1829, pp. 310-357) are fully dealt with. Desvignes’
names have ever since remained unsynonymized, a mere
encumbrance to catalogues, and it is only now that they
are for the first time relegated to their true positions.
Bridgman also described five supposititiously new species
in our Transactions (1882, p. 161; 1883, p. 170; 1886,
pp. 364-5 ; 1887, p. 875), of which there are still three
considered to be good. No reliable mention of British
representatives of this group was made till the publication
in 1856 of Desvignes’ “Catalogue of British Ichneumonide,”
wherein are recorded eighteen Gravenhorstian and one
new species. Of these, B. rufipes is no more than a variety
of B. biguttatus, and B. insignis of B. exultans; the same
author’s paper of 1862 added four species under preoccupied
names, leaving the total at twenty-one kinds. In 1870
the Rev. T. A. Marshall’s “Ichneumonidum Britannicorum
Catalogus” enumerated thirty-nine species, of which
thirteen are now regarded as synonyms. Kirchner’s
“Catalogus Hymenopterorum Kurop” of 1867 mentions
sixty-two kinds, among which, however, at least twenty-
four are synonyms and three of the Fabrician titles apper-
tain to other groups. In 1872, the Entomological Society
published “A Catalogue of British Hymenoptera,” which
has ever since, though now sadly obsolete, been the basis-
list of entomophagous work in Britain. In it we find
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.)
4.20 Mr. Claude Morley on the
Bassus rufiventris incorrectly placed in the genus Poly-
blastus, and Desvignes’ names are still treated as valid ;
the number of species is placed at forty, though thirteen
of these are now considered synonymous. ‘To these I was
enabled to compile nine additions, in the paper I had the
honour of reading before you on March 6th, 1901, which
brought the actual total to thirty-six indigenous species.
The palearctic fauna now includes little more than sixty
species (a few of which I do not know), and of these I am
herein able to add eight to our catalogue, and to describe
four kinds which appear to be new.
PHTHORIMUS ANOMALUS, Sp. Nn.
Head not narrowed posteriorly, entirely black and subglabrous
with the strongly pilose maxillary palpi apically white; vertex
posteriorly entire; frons nitidulous, shortly pilose, sparsely and
obsoletely punctulate, centrally subcarinate with the scrobes large
and glabrous ; face strongly nitidulous; epistoma somewhat convex
and distinctly disereted from the short, broad, apically strongly
emarginate clypeus by a straight fossa which is laterally produced
upwards to the orbits ; cheeks much shorter than basal width of the
stout and laterally margined mandibles. Antenne short, not reaching
beyond apex of thorax; black with the pedicellus pale above;
flagellum possessing about nineteen joints, ferrugineous beneath with
apices of the joints subnodulose and infuscate ; apical joint conical
and obtuse. Thorax strongly nitidulous, immaculate ; mesonotum
anteriorly subconvex, evenly and finely punctate, with distinct
notauli ; mesopleurz sparsely and evenly punctate, basally impressed
in the centre; metathorax very strongly and evenly punctate with
the petiolar area subobsolete and spiracles circular ; areola glabrous,
parallel-sided and twice longer than broad, with lateral costze dis-
tinct, but costula: wanting ; metapleuree sparsely pilose. Scutellum
black, hardly convex, nitidulous and obsoletely punctate ; post-
scutellum very small. Abdomen longer than head and thorax,
immaculate, very strongly and evenly punctate throughout, with
only the apices of the segments glabrous, and broadest at the apex
of the first ; second segment with distinct thyridii ; terebra exserted,
as long as the abdomen, with spicula flavous and strongly acuminate
apically, valvule black and very strongly setiferous-pilose through-
out. Legs clear red with all the coxe and basal joint of trochanters
black, with all the apical tarsal joints infuscate ; hind metatarsi,
except apically, flavescent and four times longer than their con-
Ichneumonidous Group Tryphonides schizodonti. 421
colorous ecalearia; tarsi not longer than the tibiz, of which the hind
pair is very slightly intumescent before the flavescent base. Wings
hyaline with the stigma luteous, tegule and base of costa pale
flavous ; areolet sessile, twice broader than long ; nervellus opposite
and intercepted distinctly below the centre.
Length,6 mm. (Terebra,4mm.) @ only.
There can be no doubt that this anomalous species,
from the conformation of the mandibles and spatulately
compressed abdomen, belongs to this genus as set forth by
Thomson (Opusc. Ent., xiv, 1474); and perhaps also to
Phthorima, Forst. (Verh. pr. Rheinl., 1868, p. 162), but the
latter’s diagnosis is too short and he presents no type. It
is closely allied to P. compressus, Desy., but differs abund-
antly in the posteriorly broader head, distinct notauli,
the metathoracic structure, abdominal puncturation,
strongly exserted terebra and very much broader areolet.
The Rev. F. D. Morice took a single specimen at Long-
cross, on June 17th, 1904; the type is in my collection.
PROMETHUS DODSI, sp. n.
Black ; legs, scutellum, post-scutellum and centre of abdomen
broadly, red; antennze beneath, part of the face and of the mouth
flavous ; mesonotum closely and coarsely punctate, dull ; antenne
as long as the body ; clypeus bifoveolate.
Length, 6 mm. ¢.
This species is so closely allied to P. scwtellaris, Bridg.,
as to require no detailed description. Therefrom it differs
in its twice longer antenne (which possess twenty-four,
not twenty-two, joints), its bifoveate clypeus which is sub-
dentately produced apically in the centre and distinctly
impressed transversely before the base; in its dull and
coriaceously punctate mesonotum, distinct and discally
coalesced notauli, strongly transverse areola; in the broadly
flavous facial orbits, entirely rufescent clypeus, apically
black epistoma, the antennz entirely flavidous beneath,
darker scutellum and red _post-scutellum, immaculate
trochanters, testaceous stigma; and in having the abdomen
and especially its basal segment stouter, with segments
three, four, base of the fifth and. apex of the second entirely
clear red, the last-mentioned being striolate from its base
nearly to its apex. In the dull and confluently punctate
mesonotum it resembles P. pulchellus, from which the
422 Mr. Claude Morley on the
entirely pale cox and scutellum, as also the clypeal
structure, at once distinguish it.
Taken by Dr. Capron, probably in Surrey; the type is
in my collection.
Uxort mihi hoc insectum dicatum volo.
HOMOPORUS NIGER, sp. n.
Head as broad as thorax, closely and finely punctate, somewhat
dull; black with face, mouth except apices of mandibles, frontal
orbits and the cheeks shortly, stramineous ; vertex narrow and not
posteriorly emarginate ; frons strongly canaliculate centrally and the
epistoma subdeplanate ; clypeus convex, apically margined, evenly
rounded and hardly emarginate apically in the centre ; cheeks as
long as breadth of mandibles at their base. Antenne slightly longer
than head and thorax, filiform, black above, scape and pedicellus
stramineous and flagellum ferrugineous beneath. Thorax stout,
somewhat nitidulous, strongly and evenly but not confluently
punctate ; notauli anteriorly wanting, though traceable from the
proad and hamate pale line before the wing to the subdeplanate disc
of the mesonotum ; dots beneath both pairs of wings as well as
before the anterior pair pale, as also is the mesosternum anteriorly ;
metathorax scabriculous, with the lateral cost, petiolar and sub-
quadrate basal areze entire. Scutellum strongly deplanate, black ;
deeply and evenly but not confluently punctate. Abdomen sub-
parallel-sided, entirely black ; the two basal segments coarsely and
confluently punctate, with their ventral plica dull testaceous ; the
first basally bicarinate nearly to its centre and the second basally
irregularly striolate; the third segment basally scabrous, apically
(with the following) nitidulous, obsoletely punctate and pilose ;
sixth apically truncate and not compressed, the two last incon-
spicuous and hardly exserted. Legs clear fulvous ; all the coxe and
trochanters clear stramineous with the hind coxe basally black ;
hind tarsi not infuseate, with calcaria one-third the length of their basal
joint. Wings normal, radix stramineous, stigma fulvous ; areolet
sessile with the outer nervure sub-obsolete ; nervellus sub-opposite
and intercepted below its centre.
Length,i mm. ¢ only.
This species is certainly allied to H. brevicornis, Thoms.
(only known in the ¢ sex), and it is just possible that it
may represent its male, though differing in the short
petiolar caring, posteriorly entire vertex, entirely pale
stigma, substriate second segment and the colour of the
Ichneumonidous Growp Tryphonides schizodonti. 423
legs. At first I had thought it possibly the unknown f
of H. longipes, Holmgr., to which the colour of the legs is
more nearly referable, but the comparatively short antenne,
entirely black abdomen and fulvous hind tarsi seem to
preclude such a situation. In its distinct metathoracic
are, it appears related with the Aniarophron-group, with
which, however, it has no other character in common.
I swept this species from rank herbage in Wroxham
Broad in the evening of June 14th, 1901; the typé is in
my collection.
HOMOPORUS REFLEXUS, sp. n.
Head closely and confluently punctate, dull, black; vertex not
broad, posteriorly entire ; frons centrally distinctly, but not deeply,
canaliculate ; face coriaceously punctate somewhat broader apically,
with the epistoma a little convex and quadrately pale in the centre ;
clypeus testaceous, deplanate, not laterally elevated, but with the
apical margin entire and strongly reflexed ; palpi and base of the
stout mandibles flavescent, latter slightly broader than the length of
the cheeks. Antennz longer than head and thorax, and slightly
pilose, filiform, entirely black with the apices of the seven basal
flagellar joints subnodulose. Thorax black, with a hamate line and
a dot before with a longitudinal callosity beneath the front wings,
and the basal margin of the mesopleurz flavous ; mesothorax some-
what nitidulous, finely and evenly punctate throughout with the
notauli entirely wanting ; metathorax evenly scabriculous throughout
with minute circular spiracles, and the apex centrally substrigose
between two broad and shallow fovee. Scutellum subdeplanate,
evenly and closely punctate, somewhat nitidulous, with the apical
margin, together with that of the post-scutellum, transversely flavidous,
Abdomen elongate-oval, immaculate, centrally as broad as thorax,
finely alutaceous and dull, becoming nitidulous towards the slightly
compressed anus ; basal segment quadrate, laterally marginate with
no carinw, the following transverse with the second obsoletely
aciculate at its base ; terebra reflexed. Legs clear red, with the tibia
and front tarsi testaceous; the hind tibiz becoming gradually
nigrescent from centre to apex, and theirtarsi entirely and the anterior
at the apex, black ; hind calcaria testaceous and not reaching centre
of the metatarsus. Wings hyaline, with tegule white, stigma
testaceous ; areolet subpetiolate, broad with the outer nervure well
defined ; nervellus a little postfurcal and intercepted only slightly
below the centre.
Length, 6} mm. @ only.
4.2.4 Mr. Claude Morley on the
From all the other members of the genus bearing an
areolet and no petiolar carine, the present is very distinct
in its apically entire and strongly reflexed clypeus. The
alutaceous abdomen, thoracic coloration and general con-
formation ally it with H. crassicrus, Thoms., from which it
is sufficiently distinct in its clypeal and vertical structure,
canaliculate frons, apically explanate face, the antennz
entirely and scutellum laterally immaculate, the intercep-
tion of the nervellus, acutely margined basal segment, the
dull testaceous hind tibize and abdominal plica.
The type, which is in my collection, was taken by
Dr. Capron probably in Surrey; I also took this species
in a greenhouse at Ryde in the Isle of Wight in
August 1902.
Homoporus INcIsus, Thoms. ~.
Its author only knew the female of this species ; the male differs
in nothing but size and coloration. Epistoma, clypeus and mouth
excepting apices of mandibles, stramineous, orbits and cheeks
immaculate : a broad subhamate line before and a callosity beneath
the front-wings, with the basal margin of the mesopleure stramineous;
apices of scutellum and of post-scutellum, with the sides of the
former somewhat broadly flavous ; legs pale red with all the coxe
and trochanters, and the hind tibiz (except at their extreme base
and apex), whitish ; abdomen subparallel-sided, broadest behind
the centre.
Length, 54 mm.
A SYNOPSIS OF THE PALAARCTIC BASSIDES.
(2). 1. Metathoracic spiracles large and
pale ; scutellar fovea costate. TRICHOMASTIX.
(1). 2. Metathoracic spiracles small and
immaculate ; scutellar fovea
simple.
(8). 3. Face dull and punctate, not dis-
tinctly impressed longitudi-
nally.
(5). 4. Notauli deeply impressed ; scape
excised to its centre . . . ZOOTREPHUS.
ra
or
. Notauli obsolete; scape not
deeply excised.
Ichneumonidous Group Tryphonides schizodonti. 42:
(7).
(2).
(1).
a os
6. Anterior segments transversely
impressed ; tibia white ...
7. Anterior segments not impressed
or tibiz not white
8. Face shining and sah pleest
bicarinate lloeetea eee!
9. Areolet wanting; abdomen not
entirely black.
Areolet present ; abdomen Wino
and spatulately compressed .
TRICHOMASTIX, Voll.
A large black, shining species,
with legs red; scutellum
laterally white
ZooTREPHUS, Thoms.
1. Flagellum 18-jointed; anus
broadly black; metathorax
more rugose ae
2. Flagellum pe eialed: anus
* narrowly black ; metathorax
less rugose .
Bassus, Fab.
1. Anterior cox not entirely pale ;
abdomen rarely red-marked.
2. Hind coxe red ; basal segment
apically pure white . ;
3. All the coxe black ; basal seg-
ment not white-marked.
4, Apex of hind tibize black ; 2
epistoma white-marked
5. Apex of hind tibiz ferrugin-
eous; 2 epistoma immaculate
6. Anterior coxze entirely pale ;
abdomen usually centrally red.
7. All the cox pale.
8. Abdomen not broadly red cen-
trally ; hind tibiz bicolored .
9, Abdomen broadly red centrally;
hind tibiz tricolored :
10. Hind cox basally black ; cen-
tral segments red-banded .
Bassus.
Homoporus.
PROMETHUS.
PHTHORIMUS.
1. flavipes, Holmgr.
. rufiwentris, Grav.
. holmgreni, Bridg.
. albosignatus, Grav.
. varicoxa, Thoms.
. annulatus, Grav.
. tricinctus, Grav.
. letatorius, Fab.
. multicolor, Grav.
Or
426
. Notauli wanting ;
Mr. Claude Morley on the
Homororvus, Thoms.
. Areolet wanting (?SyRPHOCcTO-
nus, Forst.).
. Metathoracic arez not entirely
wanting.
. Second segment transverse ;
anus often somewhat dull.
. Basal area distinct and entire ;
hind tibiz not white.
g; scutellum
pale.
. Femora entirely, abdomen not
broadly, red’ < weve) ee.
. Femora basally black ; abdomen
usually broadly red .
. Notauli distinct; scutellum
black.
. Hind femora black ; orbits
alone pale .
. Hind femora red ; face im-
maculate
. Basal area obsolete ; hind tibiz
white.
. Scutellum, and often the meso-
notum and pleure, red
. Scutellum centrally black, apex
or also the sides flavous
. Second segment quadrate ; anus
strongly nitidulous.
. Clypeus convex; antenne
shorter than expanded wings
. Clypeus deplanate ; antennez
as long as expanded wings.
. Coxe red; epistoma immacu-
late ; second segment striolate
. Coxee black ; epistoma pale ;
second segment not striolate .
. Metathoracic aree entirely
wanting.
. Scutellum subtumidous, im-
maculate
. Scutellum normal, nearly
always pale.
1. cinetus, Grav.
2. bizonarius, Grav.
3. graculus, Grav.
4. brevitarsis, Thoms.
5. pectoratorius, Gr.
6. alpinus, Holmer.
7. caudatus, Thoms.
8. longipes, Holmgr.
9. punctiventris, Th.
10. borealis, Holmgr.
Ichneumonidous Group Tryphonides schizodonti. 42
(29).
(28).
(27).
(26).
(25).
(24),
(23).
(22),
(1).
(60).
(41).
(36).
(35).
(34).
(33).
(40).
(39).
(38).
Ci).
(56).
22.
23.
24,
25.
26.
27.
28.
29.
30.
31.
32,
. 42,
43.
Vertex of head more or less
emarginate.
Scutellum only partly pale;
hind tibie usually black
with base white.
Meso-humeral marks, and usu-
ally the epistomal, wanting.
Metathorax not apically red
Metathorax apically red .
Meso-humeral marks, and the
epistoma always pale
Scutellum entirely pale ; hind
tibizw red with apex black
Vertex of head not emarginate ;
scutellum laterally pale
Areolet present or abdomen
broadly fulvous.
Abdomen with no segment
entirely red; hind tibiz
usually white.
Petiolar carinz distinct and
usually elongate ; tibize not
mainly white.
. Hind tibiz basally white.
. Petiolar carinz parallel .
. Petiolar carine divergent
(?ENIZEMUM, Forst.)
. Hind tibiz not white-marked.
. Clypeus not apically excised ;
scutellum black ; areolet weak.
. Anterior cox and hind tarsi
black.
. Anterior coxs and hind tarsi
| ft) Ee chee
. Clypeus apically excised ; scu-
tellum pale ; areolet strong .
. Petiolarcarine shortor wanting ;
hind tibize usually mainly
white (? HomoTROPUS, Forst.)
Scutellum not entirely pale, nor
the second segment strongly
elongate.
Scutellum of normal size and
convexity.
11. biguttatus, Gray.
12. abdominator, Bridg.
13. favolineatus, Gr
14. tarsatorius, Panz,
15. fissorius, Grav.
16. ornatus, Grav.
17. deplanatus, Grav.
18. brevicornis, Thoms.
19. niger, Morl.
20. sundevalli, Holmer.
7
. 64.
BOD:
Mr. Claude Morley
. Clypeus apically reflexed and
entire
. Clypeus apically fom reflemed
nor entire.
. Clypeus apically strongly ex-
cised and laterally elevated
: Clypeus apically emarginateand
not laterally elevated.
. Clypeus laterally foveate.
. Hind tibize basally infuscate.
. Scutellum apically white; 9
flagellum red .
51. Scutellum usually dssevallly
white ; ? flagellum black
. Hind tibia often externally,
though hardly basally, in-
fuscate .
. Clypeus not frau patie
. Hind tibiz normal and basally
infuscate . .
. Hind tibiz stout and na asalty,
infuscate
. Scutellum large and FORE
strongly convex.
. Metathorax rugulose ; second
segment longitudinally stri-
gOSe .- 8
. Metathorax Gnely, punctate ;
second segment not strigose .
. Scutellum entirely pale ; second
segment strongly elongate
. Abdomennearly always broadly
red centrally; tibiz not
white.
. Scutellum not entirely black or
basal segments scabrous.
. Scutellum at most apically pale;
second segment punctate .
. Scutellum mainly pale ; second
segment subglabrous.
Areolet present ; ‘second seg-
ment elongate pene
Areolet wanting ; second seg-
ment transverse
22.
30.
the
. reflecus, Morl.
incisus, Thoms.
. ruficornis, Holmer.
. pumilus, Holmer.
. dimidiatus, Schr.
. longiventris, Thoms.
. crassicrus, Thoms.
. strigator, Fab.
. megaspis, Thoms.
. xanthaspis, Thoms,
. elegans, Grav.
. pallidipes, Grav.
pulcher, Holmer.
Ichneumonidous Growp Tryphonides schizodonti. 429
(61). 66.
(68). 67.
(67). 68,
(12). 1.
(7), 2:
(Gio
(5). 4.
(4). 5.
(3).
(2).
(11). 8
(10). 9.
(9). 10.
-~I &
(8). 11
(1). 12.
(14), 13.
(13). 14.
(16). 15.
(15). 16.
(18). 17.
(19), 218:
(20). 19.
(19). 20.
Scutellum entirely black ; basal
segments not scabrous (? ANIA-
ROPHRON, Forst.)
All the cox pale ; areolet pre-
sent . .
Hind coxee Pact pldek's areo-
let wanting “ss.
34, signatus, Grav.
35. hygrobius, Thoms.
PrRomETHUS, Thoms.
All the coxe pale; petiolar area
not coarsely sculptured.
Scutellum pale.
Abdomen broadly red centrally.
Mesonotum dull; second seg-
ment mainly striolate .
Mesonotum shining; second
segment basally striolate .
. Abdomen nearly entirely black
- Scutellum black.
. Notauli present ; coxze flavous.
Abdomen nearly entirely black
Abdomen broadly red in the
centre
. Notauli wanting; coxe Hrlaite
Hind coxe mainly black ; peti-
olar area finely sedivearest
Basal segment twice longer than
broad ; coxa white
Basal segment not elongate ;
coxe flavescent.
Third segment with a basal
fascia or lateral gutta citri-
nous ‘
Third segment eth no Teite
nous rane
Mesonotum dull and confiuently
punctate
Mesonotum shining a cara
punctate.
Stout ; second segment apically
glabrous eA Tas 7
Slender; second segment en-
tirely scabrous
1. dodsi, Morl.
2. scutellaris, Bridg.
3. nigriventris, Thoms.
4, melanaspis, Thoms.
5. sulcator, Grav.
6. albicoxa, Thoms.
7. cognatus, Holmer.
8. laticarpus, Thoms.
9. pulchellus, Holmgr.
10, festivus, Fab.
11. dorsalis, Holmgr.
TRANS. ENT. SOC. LOND. 1905.—PART IV. (DEC.) 29
430 Mr. Claude Morley on the
PuatTHoRIMus, Thoms,
(2). 1. Areola transverse ; temples nar-
row ; abdomen not strongly
punctate ... + + « « «+ Il. compressus, Desv.
(1). 2. Areola elongate ; temples broad ;
abdomen strongly punctate . 2. anomalus, Morl.
Synonymy, Economy AND DISTRIBUTION.
During the course of the last ten years I have been
enabled to amass a little over seven hundred specimens of
this group in Britain, and perhaps it may not be out of
place to here indicate their relative frequency in our islands.
The species bearing an asterisk have not yet been noted as
indigenous; of the rest there are only three species not
represented in my collection, from the closely examined
specimens of which alone this summary is drawn up, the
respective number of specimens there representing each
species is indicated in parentheses. A dagger indicates that
the species is new to Britain.
T. flavipes, Holmer. (1.) Very rare ; my only specimen was
taken by Dr. Capron, probably about Shere in Sur-
rey; Bridgman records it from Worcester in May, and
doubtfully from Norfolk. It has been bred from a
Syrphus pupa. [=T7. polita, Voll.; =. tibialis,
Bridg.; =Z. pallipes (sic), Thoms.]
Z. rufiventris, Grav. (7.) Not rare at Brandon in Suffolk
in June 1903; Lowestoft on umbelliferous flowers,
August; Wicken in Cambs., June; Reigate in July.
[ =sulcator, var. 2, Gray., ¢.]
Z. holmgreni, Bridg. (16.) Somewhat common in the
Breck district of Suffolk, June to August; Surrey
(Capron) ; Greenings (W. Saunders) ; Harting in May
(Beaumont); Point of Aire (Tomlin). [I have fol-
lowed Thomson in considering the species of Zootre-
phus as distinct; it is quite possible that Bridgman
did not know Z. rufiventris, which stood under an
incorrect genus in the British list. The relative
rugosity of the metathorax and rufescence of the
abdomen are the only features of separation which to
me appear to be at all constant. Thomson separates
them thus :—
Ichneumonidous Group Tryphonides schizodonti. 431
HOLMGRENI, @°. RUFIVENTRIS, @.
Flagellum ferrugineous, 22-jointed infuscate above, 18-jointed.
Metathorax more rugose less rugose.
Basal segment longer.
rugose with carinze distinct less rugose with shorter
carine.
2nd segment fulvous or (var. b) black with red line thyridii
basally black with thyridii obsolete large and subcircular.
Qnd to 4th with epipleurz inflexed 2nd and 3rd with epipleure
inflexed.
3rd and 4th always red red, discally black-marked.
Hind legs elongate less elongate.
femora red red with black dot beneath.
trochanters citrinous stramineous, basally black.
coxe basally black black to beyond their centre.
Intermediate coxz citrinous basally black. ]
B. albosignatus, Grav. (4.) Certainly rare, occurring only
in July; Cadney, Lines. (Peacock); Nairn (Yerbury) ;
Southwold on flowers of Heraclewm sphondylium ;
Kessingland, Suffolk, flying on the beach. It has
been bred from Syrphus sp.[ = albosignatus, Grav.,
2 et var. 3, $; Holmer., excl. var. 3.]
B. varicoxa. Thoms. (17.) Common, June to August;
Suffolk, Cambs., Herts., Poimt of Aire. [=<a/lbosig-
natus, var. 3, Holmer. = ? albosignatus, var. 1, Grav.]
B. annulatus, Grav. (9.) Not very common from middle of
July to end of September; Suffolk, Surrey. [= ?
albosignatus, var. 2,Grav. It differs from B. varicoxa,
Thoms., in the less deeply impressed incisures,
entirely () and internally (f) ferrugineous apices of
hind tibiz; smaller humeral and scutellar pale marks
and immaculate 2 epistoma. The latter is probably
no more than a variety.]
B. tricinctus, Gray. (43.) Abundant, May to September ;
Isle of Mull (Zomlin); Yorks. (Askrigg, ;£iliott),
Gloucester, Hereford, Herts., Suffolk, London, Surrey,
Sussex, Hants., Devon. [= albosignatus, var. 4, ,
Grav. = nemoralis, Holmgr. This species differs from
nemoralis in nothing but the abdominal coloration ;
the latter is a var.]
B. letatorius, Fab. (90.) Abundant, May to September ;
Norfolk, Suffolk, Essex, Lincs., Herts., Gloucester,
432 Mr. Claude Morley on the
London, Kent, Surrey, Sussex, Isles of Wight and
Purbeck, Cornwall (Cremyl, Newbery), Also from
Bucarest. It is said to have been bred from such
varied hosts as Adimonia rustica, Tortrices, Syrphus
balteatus and S. pinasiri (= ? corollx);—cf. also
Entom., 1884, p. 167.
B. multicolor, Grav. (18.) Very local but common where it
occurs, in marshes, June; Horning Ferry in Norfolk
and Tuddenham Fen in Suffolk; Sussex, Surrey.
[= deletus, Thoms, ¢ .]
HT, cinctus, Grav. (18.) Uncommon in woods in May and
June, also found in autumn; Lines., Herts., Suffolk,
Surrey. [= lateralis, Grav. = albicinctus, Desv., 3 ;
var. = scabriculus, Holmgr.]
HT, bizonarius, Grav. (15.) By no means common, June to
September; Herts. Norfolk, Suffolk, Essex; ap-
parently commoner in Surrey. Also from Nantes.
= cingulatus, Holmgr. = ? frontalis, Brisch. This
species greatly resembles Bassus multicolor superfici-
ally in its more or less distinct tranverse segmental
impressions, entirely white scutellum and usually
brightly cinctured central segments; the coloration
of the hind tibia, however, readily distinguishes it
from that genus. The outer recurrent nervure is
rarely entirely wanting. |
H. graculus, Grav. (18.) Not uncommon in boggy spots in
May, July and August; Suffolk, Herts., Gloucester,
Surrey, Conwall. [ B. graculus, Grav., 2 = obscuripes,
Holm., 2 = rufocinctus, Desv., f 2.)
H. brevitarsis, Thoms.* Switzerland.
H. wectoratorius, Grav. (11.) Local in woods in May and
August; Suffolk, Surrey, Isle of Wight.
H. alpinus, Holmgr. 2 [= ? monticola, Voll., $.J* Scandi-
navia and ? Switzerland.
H. caudatus, Thoms.* (1.) The only specimen I have seen
was taken by Capron in Surrey.
H. longipes, Holmgr.* Svandinavia.
H. punctiventris, Thoms.+ (2). Felden near Boxmoor, in
Herts. (Piffard) ; Cornworthy near Totnes, in Devon
(Marshall). i
H, borealis, Holmgr.* Lapland.
H. bigutiutus, Grav. (9.) Probably not common; I only
possess it from Surrey and Herts. [= var. rujipes,
Grav., Holmgr. = var. (scut. apically white) deplanatus,
Ichneumonidous Group Tryphonides schizodonti. 433
Grav. et Holmer. excl. f ( = diguttatus, var. e,
Thoms.); = confusus, Woldst. This is a most
variable species and the sexes are dissimilar. |
H. abdominator, Bridg. The unique 2 was taken by
Bignell at Dousland (Devon.), August 23rd, 1884, and
is in his collection. [This is probably no more than
a colour variety of H, biguttatus, though considered
valid by Thomson. ]
I. tarsatorius, Panz. (52.) Alindanti in woods and marshes
in May and June, also occurs on flowers of Angelica
sylvestris in the autumn; Yorks. (Askrigg, Zil/cott),
Lincs., Suffolk, Herts., Surrey, Sussex, Isle of Wight.
Also from Bucarest. It is said to have been bred
from Syrphus balteatus, Depressaria angeliella and
Coccinella sp. [ = insignis et exultans, Grav. et
Holmgr. = pulchellus, Desv.,. (nec Holmer.) =
flavus, Desv., f (var. c, alis areola completa, Thoms.).
This is the only species I know which may or may
not possess an areolet; when present it is very small,
oblique, consisting of little more than the duplication
of its basal nervure ; only one ? of my fifty specimens
possesses it.]
HT, flavolineatus, Grav. (11.) Not very common, June and
August; Suffolk, Herts., Surrey, Devon. [= inter-
ruptus, Holmer., J = bimaculatus, Holmer. f ; nec Zett.]
H, fissorius, Grav. (9.). Uncommon and always occurs
singly, May, July, and September; Kent (Hunting-
field, Morice), Surrey, Hants., Suffolk, Herts.,
Gloucester. [ = punctatus, Bridg.; nec fissorius,
Holmer. |
Hf. ornatus, Grav. (7.) Very uncommon, May, August,
and September; Suffolk (Lowestoft and Southwold),
London (Blackheath and Plumstead, Beaumont),
Hants. (Lyndhurst, Adams), Isle of Wight (Ryde).
It has been bred from Syrphus sp. [ = frenator,
Desv., 2. = ornatus, Thoms., 2. = var.?2; deplanatus,
Holmer., “3.”]
HT. deplanatus, Grav. (3.) In Dr. Capron’s collection from
Surrey. It has been bred from Syrphus sp. [deplan-
atus, Grav. et Holmgr., excl. 2 = nigricornis, Thoms.,
g. Gravenhorst’s g differs from that of B. ornatus
(with which it is incorrectly synonymized by Thomson)
in its immaculate face, red antennz, immaculate
sternum, black-marked front cox, simply white-
43.4 Mr. Claude Morley on the
circled hind tibize, the pale marking of which does
not extend down the leg, etc.]
HI. brevicornis, Thoms.* France.
FH, niger, Morl.+ (1.) Norfolk.
H, sundevalli, Holmgr. I have not seen this species;
recorded from Devon. in September (cf. Trans. Devon.
Ass., 1898, p. 501). [= scabrosus, Desv., 3.]
H, reflexus, Morl.+ (2.) Isle of Wight and Surrey.
H. incisus, Thoms.+ (4.) Apparently rare ; Surrey and the
Isle of Wight, in August.
HI, ruficornis, Holmgr. [= ? suspiciosus, Brisch.]* Sweden
and ? Prussia.
HT, pumilus, Holmgr. (15.) Common from June to Sep-
tember; Cheshire, Herts., Suffolk, London (Wimble-
don and Blackheath), Surrey, Isle of Wight. [=
thoracicus, Desv., 2 = ? pictus, Grav. part. One f I
examined possessed a distinct areolet in the left wing
but no trace of one in the right.]
Hf, dimidiatus, Schr. (45.) Abundant, May to September ;
Scotland (Crockston and Bonhill, Dalglish), Suffolk,
Herts., Sussex, Surrey, Hants., Isle of Wight and
Cornwall. [= pictus, Grav., Holmgr., Thoms, =
planus, Desv., 2 (2 = var. b, Thoms.).|
FT. longiventris, Thoms.*+ (1.) The only male I have seen
was swept in a marsh at Brandon in Suffolk, July 4th,
1903. [= pumilus, Holmer. part.]
FT, crassicrus, Thoms.t (6.) Apparently not common,
August and September; Suffolk, Sussex, Surrey.
[= fissorius, Holmgr., nec Grav.]
H, strigator, Fab. (1.) The only specimen of this species
I have seen was swept from herbage in Henstead
marsh (Suffolk), August 28th, 1898. Said to have
been bred from larve preying upon Siphonophora
absinthi. [This f is very like that of H. tarsatorius
in facies, but—besides the distinct areolet and
scutellar coloration—the second segment is longer,
coarsely longitudinally strigose with the thyridi
obsolete; metathorax rugulose and much narrower
above the hind coxas, with the petiolar area centrally
striolate and bearing some traces of a basal area ;
head narrower behiad the eyes, with the vertex
much less emarginate ; face more distinctly punctate
with the epistoma more prominent and cheeks
shorter; clypeus similarly excised centrally but
Ichnewmonidous Group Tryphonides schizodonti. 485
acuminately explanate at the sides; antennz shorter
and consisting of 21 (not 20,as in the latter) flagellar
joints and the scutellum is more convex and coarsely
punctate (cf. also Brischke, Schr. Nat. Ges. Danz.,
1878, no. 6, p. 112). The areolet and sculpture will
also distinguish it from H. fissorius. It appears to
only differ trom H. megaspis, Thoms., in the sculpture
of the metathorax and second segment, and in the
colour of the hind tibie.]
HI. megaspis, Thoms.* Bavaria.
H. xanthaspis, Thoms.+ (4.) Rare; three from Surrey in
Capron’s collection and one I swept in Tuddenham
Fen (Suffolk), August 23rd, 1905.
Hi, elegans, Grav. (35.) Abundant; June to October; London
(Blackheath), Norfolk, Suffolk, Herts., Kent, Sussex,
Surrey, Isle of Wight, Devon. Also from Nantua and
Nantes. It has been bred from Bombyx querecus,
and Gracilaria phasianipennella. [= elegans, Grav., 2
(f in error), Holmgr., Desv. = rufonotatus, Holmgr.
= nigritarsus, Grav., Holmgr., = picitans, Desv., $2
(¢ = var. f., Thoms.).]
HI. pallidipes, Grav. I have not seen this species; re-
corded from Devon in August (¢f. Trans. Devon.
Ass., 1898, p. 501).
H. pulcher, Holmgr. (2.) Very rare; two from Surrey in
Capron’s collection.
Hf, signatus, Grav. (28.) Abundant, June to September ;
Lincs., Norfolk, Suffolk, Herts., Surrey, London,
Hereford, Ireland (Rossbeigh in Kerry). It has
been bred from Syrphus sp.
Hf, hygrobius, Thoms. (13.) Not uncommon, May to October;
Surrey, Suffolk, Lines., Scotland (Giffnock). Also
from Nantes. [= festwus, Holmer., part; nec Grav. ]
P. dodsi, Morl.+ (1.) Surrey.
P. scutellaris, Bridg. (4.) Certainly rare ; all my specimens
are from Surrey ; Bridgman describes it from Devon.
[B. scutellaris, Bridg., 2 (nec f), Thoms., f 2. ]
P. mgriventris, Thoms.* Sweden.
P. melanaspis, Thoms.* Bavaria.
P. sulcator, Grav. (78.) Abundant, May to October ; Devon.,
Hants., Worcester, Herts., Suffolk, Norfolk, Cambs.,
Scotland (Crockston, Dalglish). It has been bred
from Syrphus sp. [sulcator, Grav., excl. var. 2-5 et
1 g = areolatus, Holmer. ]
436 Mr. Claude Morley on the
P. albicova, Thoms.+ (9.) Probably not uncommon, June
to August; Stafford., Herts. Norfolk, Surrey, Isle of
Wight. [= ? sulcator, var. 5, Grav., ¢.]
P. cognatus, Holmgr. (20.) Common, May to September ;
Suffolk, Norfolk, Herts., Surrey, Sussex, Cornwall
(Botusfleming, Marshall).
>. laticarpus, Thoms.t (5.) Rare, in boggy situations; 2 2
swept in Henstead marsh (Suffolk), 12, vii, 1900,
and Rockland Broad (Norfolk), 10, vi, 1901; f 2@
at Metton (Norfolk), 30, viii, 1903; and Greenings
(Surrey) from W. Saunders’ collection, viii, 1871.
Also from Nantes. [= ? gracilentus, Holmer. ]
P. pulchellus, Holmgr. (54.) Very common May to Octo-
ber; Suffolk, Norfolk, Northants., Herts., Wilts., Wales
(Trefriw, Newbery), Devon., Cornwall, Isle of Wight.
[=sulcator, var. 1f,3 et 4, Grav. = festivus, var.
2 et 3, Grav. =/fraterculus, Forst.]
festivus, Fab. (20.) Uncommon, July only; Suffolk,
Herts., Surrey, Kent. It has been bred from Syrphus
sp., and Heliodines roesella (ef. Isis, 1848, p. 986).
[=/festivwus, Grav., $2, excl. var., 2 et 3; Grav. var.
1, facie tota nigra, 9.|
P. dorsalis, Holmgr. (9.) Not common, in marshy places,
June and July; Suffolk, Surrey. [B. dorsalis, Holmgr.
2 = maculatus, Desv. f §.]
P. compressus, Desv.(1.) Very rare; I took a female at
Felden in Herts., while staying with Mr. A. Piffard,
flying on a chalky hillside, on August 9th, 1903—
probably the only localized British specimen. I
noticed that in life the abdomen was no _ less
compressed. [= Bassus ibalioides, Kriech.]
P. anomalus, Morl.t (1.) Longeross.
I propose to conclude this paper with the description of
four little-known species of Bassus (sensu, Grav.), which
have been utterly ignored by all systematists :—
MY
SS
BASSUS HISPANICUS, Spinola.
Ann. Soc. Fr., ser. ii, I (1843), p. 118.
Dull and strongly punctate, black. Face deplanate, clypeus
apically truncate and not discreted. Antenne filiform, black, 32-
jointed, and as long as the body ; scape obconical and deeply
excised ; flagellar joints cylindrical with the second and third ob-
conical, very short and the fourth elongate. Metathorax with a
Ichnewmonidous Group Tryphonides schizodonti. 437
central transverse carina and the petiolar area basally rounded.
Seutellum and post-scutellum subconvex, immaculate. Abdomen
black with segments two to four red ; the basal longer than broad
and apically explanate ; second and third uniformly convex, with
no transverse impressions, Legs red with cox, base of the front
trochanters and apices of their tarsi, black. Wings hyaline ;
nervures basally, and tegule, red.
Length, 9 mm.
Southern Spain (Victor Ghiliani).
BASSUS ATHALIZPERDA, Curtis.
Farm Insects (1860), 53.
Black, minutely punctate and finely pubescent. Face and mouth
white, with epistoma longitudinally, and the clypeal fovez black ;
labrum and apices of mandibles dull ferrugineous, Antenne as long
as the body. Legs red and somewhat stout; coxe flavous; tarsi
and basal half of hind tibize fulvous, the latter apically and their
tarsi black. Wings iridescent ; costa and stigma fulvous, nervures
piceous ; areolet wanting.
Length, 6 mm.
England ; bred from Athalia spinarum.
BASSUS CARINULATUS, Ruthe.
Stett. Ent. Zeit. xx (1859), p. 373, f 9.
Black. Palpi infuscate; ¢ with face, mouth and apices of cheeks
stramineous ; 2 with clypeus and mandibles ferrugineous. Antennz
of g as long as the body, basally fulvous, with the scape clear
stramineous, beneath ; of 9 shorter, with the flagellum subrufescent
beneath. Thorax of 2? immaculate, with mesosternum closely
punctate; of ¢ with lines before and beneath the radix, and the
mesopleure anteriorly flavous. Scutellum black. Abdomen imma-
culate, with the basal segment rugosely punctate, bearing two sub-
parallel carinze ; the second rugosely striate ; anus of 2 closely and
finely punctate, of ¢ sparsely punctate and nitidulous. Legs pale
red, with the hind tarsi and tibiz black, the latter basally more or
less broadly white; front coxce of 2 black, of ¢ stramineous.
Wings with tegule of ¢ flavous ; areolet triangular, petiolate, entire.
Length, 5-6 mm.
The 2 resembles B&B. deplanatus, but is much more
slender; the # is nearer B. ornatus.
Iceland (Dr. Staudinger).
438 Mr. Claude Morley on Tryphonides schizodonti.
BASSUS PERONATUS, Marshall.
E. M. M.,, xii (1876), p. 194, 2
Shining, punctulate, black. Head transverse and broader than
thorax ; face deplanate and not centrally canaliculate. Antenne
as long as the body, infuscate, testaceous beneath. Thorax nitidu-
lous; mesosternum fulvous; metathorax rugose with distinct areex,
areola elevated and acutely margined. Scutellum convex, testaceous
with its centre and two elongate lateral marks infuscate. Abdomen
shining, with all the segments narrowly white apically, and not
transversely impressed ; basal segment broad, depressed, and sub-
contracted in front, with carine extending a little beyond its centre.
Legs fulvous, hind ones stout ; hind femora infuscate with their
tibize, except basally, and tarsi black. Wings with tegule white ;
areolet wanting ; nervellus intercepted far below the centre.
Length, 6 mm.
This species is said to be closely related to B. pector-
atorius, differing therefrom in its immaculate pleure and
metathoracic coste. I very strongly suspect, however,
that it represents the unknown 2 of B. strigator, Fab.
England; bred from Nematus cadderensis (cf. E. M. M.,
xi, p. 127).
IT have quite failed to discover where Bassus pipize,
Gir., is described ; it is mentioned as parasitic upon Pipiza
noctiluca in one of that author’s posthumous works (¢/.
Laboulbéne in Ann. Soc. Fr., 1877, p. 408). Dr. Giraud’s
memoirs are numerous, extending from 1854 to 1871
according to Mocsary, but this species is not mentioned in
the Annales from 1852 (when he joined the Society) to
1872, nor do I find it in the Zoological Record; it may
appear in one of his contributions to the Verh. Wien.
z. b. Ver., but it is more probably a MS. name (c¢7. Mar-
shall, Bracon. d’Europ. i, 199).
JANUARY 38RD, 1906.
THE
PROCEEDINGS
ENTOMOLOGICAL SOCIETY
LONDON
For THE YEAR 1905.
February 1st, 1905.
Mr. F. Merrirrenp, President, in the chair.
Mr. Merririexp, taking the chair as President, said it would
be in accordance with some recent precedents if he expressed
his acknowledgments for the honour which had been conferred
on him. It was also a responsibility. He succeeded one
under whose Presidency the Society, which had been steadily
rising for years in numbers, in utility, and in scientifie estim-
ation, had reached the highest point yet attained. At the
last meeting, when Professor Poulton vacated the office which
he had filled with so much distinction, two past Presidents of
the Society appropriately bore testimony to his services ; it
would not be inappropriate if he took the opportunity which
this meeting afforded him of saying how cordially he con-
curred in all that was then said. The Presidency of one so
widely known, and so appreciated. in all parts of the world
where biological research, united with powers of exposition,
was valued, reflected honour on the Society with which he
was thus associated. In the speaker’s little research work he
had had the kindest help from Professor Poulton, and he might
add that also of the many other Fellows who had taken the
leading part in the work of the Society. With their con-
tinued aid and support he hoped he might be able to perform
the duty which was incumbent on every one who filled that
PROC. ENT. SOC. LOND., 1. 1905. A
ae)
chair—to do all in his power to promote the interests of the
Society and the advancement of science, and, as he intended
to take some trouble towards these ends, he hoped he might
attain some of the moderate success which usually attends on
well-meant exertion.
Nomination of Vice-Presidents.
The President then announced that he had appointed Dr.
THomas ALGERNON CHapMaN, M.D., F.Z.8S., Dr. FrepErick
Avueustus Dixry, M.A., M.D., and Professor Epwarp B.
Poutton, M.A., D.Sc., F.R.8., as Vice-Presidents for the
Session 1905-6.
Huhibttions.
Mr. H. Sr. J. DonisrHorrs exhibited specimens of Oligota
granaria found in a granary in Holborn, the only other
localities reported hitherto being Shoe Lane, London, and
Scarborough.
Mr. W. J. Kays exhibited a specimen of the Erycinid butter-
fly Mesosemia ewmene pinned in its natural position of rest to
show its resemblance to the head of a small mammal, such as
a mouse. The eye-spots of both fore- and hind-wings came
together in such a way as to resemble the two eyes of the
animal, while the outline of the wings so closed constituted
a family class-likeness to the sharp outline of the mouse’s
head.
Professor Poutron remarked that, as the eye-spot was
placed close to the body near the base of the wings, the
ordinary protective value thereof might be explained in some
such way.
Dr. T, A. Cuapman exhibited a variety of the female of
Lycxena melanops. As a mere aberration it was interesting ;
but it was of value as showing that the position in the genus
for long accorded to the species, whether by accident or design,
close to the arion—euphemus group, was correct.
The considerable extension of the blue in this specimen
showed up certain black spots on the upper surface of both
upper and lower wings, strictly similar to these characteristics
of the arion—euphemus group. He placed with it for illus-
Crit)
tration specimens of arion, euphemus, alcon and arcas. Also a
fairly typical 2 melanops from Digne, and another from Spain,
perhaps more usual in colour, viz. with very little blue, but
of large size ; also two other specimens from Digne, in which
traces of the black spotting were easily seen, after the variety
exhibited had shown what to look for. Dr. CHapman pro-
posed to name the variety, which seemed to be undescribed,
Lycxna melanops, Boisd., var. wheeler?, Chapm.
( x 2)
var. wheeleri, in recognition of the work of the Rev. George
Wheeler on Alpine Butterflies.
Mr. F. Enock exhibited a living 9 Hybernia defoliaria,
taken as late as February Ist, and another ? taken January
28th, both at rest on north side of an oak-tree, in the same
wood at Bexley. He also exhibited on behalf of Mr. Leonard
Newman, of Bexley, two fine hybrids bred from a 4 Woto-
donta ziezac, and a 2 NV. dromedarius, the colour being that
of dromedarius, while the markings were those of ziczac. He
also showed two varieties of WV. ziczac var. bred from larve
( iv )
taken in Kent, which had the pebble marking much suffused,
and the ground-colour darker than the types.
Mr. 0. E. Janson exhibited a living specimen of Acridiwm
egyptium, L., found in a cauliflower in Bloomsbury, and
probably imported from Italy. Professor Poutron said he
had also received similar examples introduced in the same
way.
Mr. G. C. Cuampion exhibited two examples of Malachius
barnevillei, Puton, captured by Mr. Thouless at Hunstanton,
Norfolk, in June 1899, a recent addition to the British List.
Mr. H. W. AnpreEws exhibited a ¢ and 2 of Machimus
rusticus, Mg., a rare Asilid, taken in cop. at Freshwater, Isle
of Wight, on August 13th, 1903.
Mr. W. J. Lucas exhibited a 2 Panorpa cognata taken
at Byfleet Canal on August 23rd, 1904. The species occurs
at Folkestone, and is said to be found in the New Forest.
It is a little difficult at times to identify the ? alone, but
Mr. K. J. Morton also had determined the insect in question
to be P. cognata. For comparison he brought also ?
specimens of P. communis and P. germanica.
Papers.
Mr. T, Givsert Smiru read a paper entitled ‘“‘ A revision of
the genus Criocephalus, with notes on the habits of Asemum
striatum and Criocephalus ferus,’ by himself and Dr. D.
Suarp, M.A., F.R.S., and exhibited specimens to illustrate his
remarks.
A discussion followed in which Mr. DonistHorPE remarked
that he had bred Criocephalus polonicus, and Mr. A. J. Curry,
commenting on the life history of these beetles, said that he
thought the longevity of some Coleopterous insects had hardly
been appreciated. He possessed a Dytiscus still alive which
he had taken in October 1902, while Mr. DonisTHorPE
mentioned the case of a Longicorn bred by Mr. C. O.
Waterhouse which lived for twenty-one years. Commander
Waker, Mr. G. C. Coampion, and other Fellows continued
the discussion.
Papers were also read by Dr. T. A. Cuapman, M.D., F.Z.S.,
‘On the Matrivorous Habit of Heterogynis,” and “On the
Ga)
Pupal Suspension of Thais,’ the author exhibiting a number
of specimens of /Teterogynis from Spain, and elsewhere.
Mr. E. Meyricxs, B.A., communicated a paper on “ Lepi-
doptera from New Zealand.”
Mr. G. C. Cuampion, F.Z.S., contributed a paper entitled
« Another Entomological Excursion to Spain (with description
of two new species of Hemiptera by Dr. O. M. Reuter),” by
Dr. T. A. Cuapman and himself.
Wednesday, March Ist, 1905.
Mr. F. Merrrrrecp (President) in the Chair.
Election of Fellows.
The Duxe of Beprorp, K.G., President of the Zoological
Society, ete., of Woburn Abbey, Beds., and 15 Belgrave
Square, 8.W.; M. Lucren Cuoparp, Membre de la Société
Entomologique de France, of 98 Boulevard St. Germain, Paris ;
Mr. Winrrip Furet, F.R.A.S., of “ Imatra,” Bournemouth ;
and Mr. Rosert Smwney Mirrorp, C.B., of 35 Redcliffe
Square, S.W., were elected Fellows of the Society.
Obituary.
The decease of M. Henri F. ve Saussure, of Geneva,
Honorary Fellow, of the Rev. Francis Avueustus WALKER,
D.D., and of Mr. ALEXANDER Fry, F.L.8., was announced.
Exhibitions.
Mr. H. Sv. J. Donistuoree exhibited an example of Oxypoda
sericea, Heer, taken in Dulwich Wood, June 17th, 1904, a
species new to Britain; also 0. nigrina, Wat., with a type
lent by Mr. E. A. Waterhouse, to demonstrate that it is not
synonymous with sericea as stated on the Continent ; and 0.
exigua, which is also regarded there as synonymous with
nigrina.
Cat)
Mr. Hueu Main and Mr. Apert Harrison exhibited a
long series of Colias edusa, with var. helice (bred from one ?
helice sent by Dr. T. A. Chapman from the South of France),
to show the proportion of type and variety obtained. The
percentage worked out as follows :—
Notal'specimens :¢ G ="... Bay,
type: 9 9 = Bay f holes
re 47
Of the 9 @ typ,=19 . . . QTY
Vanieby) 12. 2404 1S ees
71 100
They also showed the results of similar experiments with
Amphidasys betularia, bred from a ¢ var. doubledayaria,
and a type ? taken in cop. at Woodford, Essex, in 1903,
as follows :—
Type. Var. Type. Var.
3d 22 21° = oY 207,
2 9 35 26° "= B47, 257,
57 AT 55 45
Of the 43 ¢ ¢ 22=517/%=type, following the ? parent.
21= 49% = var. 9 3 ”
Of the 61 9 2? 35=57%7 =type, following the ? parent.
26 = 43 = var. ” 3 ”
As the result of “ assembling” at Woodford, in June 1904,
it was found that: 17 = 6 type = 637,
10 = var. doubledayaria = 377
27 100
Mr. R. A. R. Priske exhibited a specimen of Helops striatus,
with a photograph, showing an abnormal formation of the
Crem)
right antenna, which was divided into two branches from the
fifth joint.
Mr. Percy H. Grimsnaw showed examples of Hyddrotza
pilipes, Stein, ¢ and 9, the latter sex being previously un-
known. The species was first recorded as British by the
exhibitor in July 1904, the first specimens having been taken
at Aberfoyle by Mr. A. E. J. Carter in July 1903. Since that
date it has also been discovered by Col. J. W. Yerbury at
Porthcawl. Mr. GrimsHaw also exhibited several specimens
of Hydrotxa tuberculata, Rond, not hitherto recorded as
British, captured by Mr. C. W. Dale and Dr. J. H. Wood in
various localities.
Dr. F. A. Dixty exhibited some cocoons and perfect insects
of hybrid Swturnias, and made the following observations :-—
“T ventured some time ago (see Proc. Ent. Soc. Lond., 1897,
pp. iii) to draw attention to the experiments in hybridisa-
tion made by Dr. Standfuss, of Ziirich, and to lay before the
Entomological Society a brief summary of his principal results.
In the year 1898 I published in ‘Science Progress’ a more
extended account of Standfuss’s work in this direction, giving
particulars of several of the hybrid forms obtained by hin,
and adding comments of my own on what appeared to be their
biological significance. This paper is included in vol. ii of
the ‘ Hope Reports’ (1901), and portions of it may be found,
though (doubtless from inadvertence) without due acknowledg-
ment, in the pages of an important work on Lepidoptera, now
in course of publication. An excellent translation, by Mr.
E. M. Dadd, of Dr. Standfuss’s account of some later results
than those recorded in his ‘Handbuch’ (Jena, 1896), was
also published in the ‘Entomologist’ for 1900 and 1901. It
is illustrated by four well-executed photographic plates.
“Dr. Standfuss has been good enough at various times to
present me with some of his specimens; these are now, with
two exceptions, in the Hope Collection at Oxford.
‘“The examples I exhibit to-night consist of hybrids of two
kinds. The first is a pair, male and female, of the hybrid
between Saturnia pavonia, Linn., ¢,and S. pyri, Schiff, 9 (see
‘Science Progress,’ 1898, pp. 188, 189; pp. 4, 5 of the paper
as included in the ‘Hope Reports,’ vol. 1, 1901). I have
( viii)
added specimens of both sexes of the parent forms for com-
parison, and it will be seen that the cross-product (which
Standfuss calls S. emilix) resembles a large S. pavonia rather
than a small S. pyri; except that the sexual disparity in size
and colour, which is so marked in S. pavonia as compared with
S. pyri, is very little developed in the hybrid. In most of
Standfuss’s specimens of this hybrid, some of the veins are
forked terminally. This it will be seen is the case with one
vein of the right fore-wing in the male exhibited.
“The second form is a hybrid with a somewhat more
complex ancestry. It consists of three males and three females
of which the female parent is S. pavonia, and the male parent
a hybrid between S. pavonia, 3, and S. spini, Schiff, ?, 7. e.
the cross-product to which Standfuss has given the: name
S. bornemanni. The present form (called by Standfuss 5S.
schaufusst) is therefore, in the common way of reckoning,
three-quarters S. pavonia and one-quarter S. spini. Dr.
Standfuss’s first attempt to rear it ended in failure, his speci-
mens all dying in the larval stage. Subsequent trials, however,
were more fortunate, and some examples of Standfuss’s own
rearing are figured in the ‘ Entomologist’ for 1900, Pl. VII,
figs. 6, 7, 8. The six individuals I now exhibit were reared
from eggs kindly sent me by Dr. Standfuss in 1895, and are
some of the actual specimens mentioned in the footnote on
p- 189 (p. 5 in the ‘Hope Reports’) of the paper above
referred to.
“Their history is briefly as follows :—
1895.
May 11. Eggs received from Dr. Standfuss.
» 23. Eggs nearly all hatched. lLarve black, hairy.
Fed on whitethorn. Show distinctly gregarious
habit.
June 3. lLarve undergoing first moult. Some in second
stage show a rather indistinct yellowish-brown
lateral line. They are still gregarious.
» 14. Larve in second moult. In third stage still
black, but some have a yellow or orange lateral
line.
June 24,
a5 (208
July 5.
4 20:
ee 2h.
js ees
St OR
Ca)
Third moult. Larve in fourth stage are still
black ; hairs whitish. Front of head pale green ;
some show small portions of green colour on body.
Eighteen larve in fourth stage given to Professor
Poulton, for the Hope Department. Five others
were afterwards given to another friend. The
subsequent history refers only to six specimens
finally retained by me.
Fourth moult. In fifth stage the ground-colour is
generally black; a variable amount of green may
be present in addition ; in two out of the six the
green predominates. Tubercles yellow, deepening
to orange.
First larva began to spin cocoon.
Three larvee now spinning. The cocoons are those
marked 1969 in the exhibit. These three larve are
black with orange tubercles, one is mottled with a
little dull green. All the larve have throughout
been variable, but especially so in the last stage,
varying from bright green with black rings (like
S. pavonia) to uniform black. Tubercles orange
in all.
A bright green larva with definite black rings,
began to spin. This cocoon is the one marked
1443,
Another larva, resembling the last, now spinning
(cocoon numbered 1327). The last larva retained
by me, a specimen with much less green ground-
colour than the two just noted, also spinning
to-day (cocoon 1273 in the exhibit).
“These larve all had a profuse diarrheal discharge before
spinning ; the discharge was clear and colourless when emitted,
afterwards becoming brown. A period of quiescence, lasting
for twenty-four hours or thereabouts, intervened between the
discharge and spinning. During this period the larve seemed
shrunken and looked torpid and unhealthy. The cocoons were
spun among twigs of the food-plant (whitethorn) with few or
no leaves, in a glass cylinder resting on a white glazed plate.
(3°)
They were all brown in colour, two (Nos. 1230 and 1327) being
a little paler than the rest.
‘“‘The first emergence took place on March 17, 1896. It is
the female numbered 1327 in the exhibit, and resulted from
the bright green pavonia-like larva which spun on July 27,
making a rather light-brown cocoon. Only one more of my
six specimens reached the perfect state, and this did not
appear until Dec. 1, 1896. It came from the bright green
larva which spun a dark reddish-brown cocoon on July 24,
1895, and is the male numbered 1443. The other four
specimens having shown no sign of emergence, I opened and
examined the cocoons in April, 1898. Three of them (num-
bered 1969), all dark brown, each contained a dead larva.
The fourth (the last one to be spun) was a somewhat pale-
brown cocoon of a curious shape, having two orifices for
emergence, only one of which was furnished with converging
bristles (No. 1273). It contained the cast larval skin and a
dead pupa.
“The four remaining examples of this hybrid now shown
belong to that portion of the original batch which was con-
signed on June 29 to the Hope Department. They were
reared with the rest, as I have related, up to the fourth larval
stage. I have no record of the time of their emergence.
“Tt will be seen that in this form (S. schaufussi) there is far
less difference between the sexes than in S. pavonia. In spite
of the fact that only one grandparent is S. spinz, the influence
of this latter species is seen both in the reduction of the sexual
disparity and also in the prevailing aspect of the larve in their
last stage. The difference between the male (No. 1443) and
those which emerged in the Hope Department is very notice-
able. A similar variability in the males of the same hybrid
was observed by Standfuss (‘ Entomologist,’ 1900, p. 346,
note ; and Pl. VII, figs. 6, 7). I attribute the semitranspar-
ency of most of the specimens to the want of vigour char-
acterising the batch generally, and showing itself in another
way by the number of larve that failed to attain the perfect
condition.
“While on the subject of Saturnia I may perhaps be allowed
to mention that a passage in Mr. Tutt’s very complete account
a
Gree")
of the larva of \ pavonia (‘British Lepidoptera,’ vol. iii,
pp. 325, 326) may give rise unintentionally to a false impres-
sion. The eighty larve there spoken of as having been
received from Norfolk were reared by myself, and not, as
might appear from Mr. Tutt’s account, by Professor Poulton,
who indeed to the best of my belief never saw them. I must
therefore assume the entire responsibility for the description
of their markings which Mr. Tutt quotes. In the succeeding
year I raised a fresh brood, numbering 120, from two of the
pink-tubercled larvae of 1885, the parent imagines being still
in my possession; 40 of these 1886 larvee were reared and
described by myself, and 80 were given by me to Professor
Poulton, who also duly noted their appearance in the last
stage, as recorded by Mr. Tutt. My original account will be
found in Trans. Ent. Soc. Lond., 1887, pp. 310, 311, having
been kindly incorporated by Professor Poulton, at my own
request, with his ‘Notes in 1886 upon Lepidopterous Larve,’
On reference to this account it will be seen that Professor
Poulton fully acknowledges the source of the description.”
The PrestpENT exhibited some Pyrameis atalanta, some of
them lent by Mr. R. 8. Mitford, bred from pupz found by
Mr. Harwood, Colchester, which emerged on the last half of
November and the first week of December last. Others in
large numbers which had emerged earlier presented no
remarkable features, but many of the later ones presented
considerable differences, especially the following:—On the upper
surface a patch or streak of blue or grey-blue scales on the fore-
wings above the scarlet band and just below the large white spot
on the costa; also a narrowing and partial interruption of the
scarlet band by the invasion of black scales ; on the under-side
of the hind-wings a considerable increase of variegation, with
an increase of the light areas, especially towards the outer
margin. ‘These effects are no doubt owing to the greater cold
to which the pups were exposed than are the ordinary
summer pup, as they resemble in their general nature the
effects caused by artificially low temperatures ; two specimens
of which the President showed obtained by him ten years
since ; also, for comparison with these and Mr. Harwood’s, a
specimen at normal temperature, and a specimen bred at
( “xii, )
80°-90° F., both also obtained ten years ago. Mr. Harwood
kept no exact records of temperatures or dates, but states
that most emerged in November, the last six in December;
severe frost having set in early in November, when some just
emerging died, apparently from this cause, and they were then
brought into a fireless room ; after a milder interval severe
frost again set in, which caused them to be removed into a
room with a fire. He remarks that probably all the very
late larvee and pup perish in this country under natural
conditions; in which the President agreed with him—he
never could get the pupe to survive ten or eleven weeks of
cooling, and believed that, as with Pyrameis cardut, the
swarms of atalanta that decorate our gardens in the summer
are all immigrants or their descendants; in the South of
England there are certainly two broods in the year, perhaps
more in hot summers. Mr. Harwood’s experience was
interesting, as it showed that there are natural variations of
temperature which may produce ettects on the facies of a
butterfly corresponding with some of the results obtained by
artificial temperatures ranging from that of an ordinary winter
to that of a hot summer.
Commander J. J. WALKER drew attention to the white spot
on the scarlet band that most showed, which had been thought
to indicate the female sex. The PresiIpEnT said he thought all
the specimens he had bred showed traces of the white spot on
the under-side, so that he did not think it was an indication
of sex; he had understood that the American examples were
without the white spot.
Professor E. B. Pourron, F.R.S., exhibited (1) Groups of
Synaposematic Hymenoptera and Diptera captured by Mr.
A. H. Hamm, of the Hope Department, Oxford University
Museum, and (2) Three specimens of Papilio hesperus, taken
at Entebbe in 1903, by Mr. C. A. Wiggins. The attention
of the exhibitor had to be called by Mr. W. Holland, of the
Hope Department, to the fact that the tails of the hind-wing
had not been broken off in these excessively worn and torn
specimens. The evidence supports the conclusion that the
tails of a Papilio, if untouched by enemies, can endure a great
deal of wear. (3)"Professor Poutron also showed Nymphaline
( xiii)
butterflies from Northern China, apparently mimetic of the
male Hypolimnas misippus, which is not known to occur in
this region.
A. discussion followed, in which Sir Groregz Hampson, Col.
J. W. Yepsrury, Mr. A. Bacov, and other Fellows joined.
Papers.
Sir Grorce Hampson read for Mrs. De ta B. NicHoin a
paper on “ Butterfly-hunting in British Columbia and Canada,”
illustrated by numerous examples of the species captured
during the summer of 1904.
Sir GEorcE Hampson, B.A., F.Z.S8., also communicated a paper
“On three remarkable new Genera of Micro-lepidoptera.”
Mr. Hersert Deuce, F.L.S., F.Z.8., communicated a paper
entitled ‘“‘ Descriptions of some new species of Diurnal Lepi-
doptera, collected by Mr. Harold Cookson in Northern
Rhodesia in 1903-4: Lycxenide and Hesperitdx, by HamMiILton
H. Drucs, F.Z.S.”
Mr. F. Du Cane Gopmay, F.R.S., D.C.L., communicated a
paper entitled ‘“ Descriptions of some new species of Satyridx
from South America.”
Mr. W. L. Distanr communicated a paper entitled,
‘“‘ Additions to a knowledge of the Homopterous family of
Cicadide.”
Wednesday, March 15th, 1905.
Mr. F. Merrirrecp (President) in the Chair.
Election of an Honorary Fellow.
SeNor Don Ianacto Bortvar, of Paseo de Recoletos Bajo,
20, and Calle Jorge Juan, 17, Madrid, was elected an Honorary
Fellow of the Society, in the place of Professor F. M. Bravurr,
deceased.
Election of Fellows.
Mr. Frank P. Dopp, of Kuranda, vid Cairns, Queensland ;
Mr. Ceci, Fiorrsuerm, of 16 Kensington Court Mansions,
(2 yay?)
S.W.; Mr. JosepH Lane Hancock, of 3757 Indiana Avenue,
Chicago; and Mr. Herserr C. Rosson, Curator of the State
Museum, Kuala Lumpur, Selangor, were elected Fellows of
the Society.
Mr. C. O. WaterHovusr announced that the late Mr.
Alexander Fry, a Fellow of the Society, had bequeathed his
large and important collections of Coleoptera to the British
Museum.
Exhibitions.
Dr, F. A. Dixzy exhibited some butterflies from Natal which
had been presented by Mr. G. A. K. Marshall, F.E.S., to the
Hope Department at Oxford, and read the following note :—
“Tt will be remembered that some few years ago Mr.
Marshall conducted certain experiments with a view to
ascertain whether the assumption of the wet- or dry-season
form of various African butterflies could be controlled by
exposure in the pupal state to artificial conditions of tempera-
ture and moisture. Some of the results of these experiments
were recorded and discussed by Mr. Marshall in Ann. Mag.
Nat. Hist., 1901, vol. ii, p. 398. Others were dealt with in a
paper published in Trans. Ent. Soc. Lond., 1902, p. 189. Most
of the material produced in the course of this research was
exhibited on the occasion when the last-named paper was read ;
the present specimens, however, though duly recorded in that
paper, had not then reached the Hope Department. They
were therefore not included in the comments added by me to
Mr. Marshall’s account of his experiments, and they have not
previously been seen by the members of this Society. This
note and exhibit may accordingly be taken as a kind of
appendix to the paper in the 1902 volume of our Transactions.
“The specimens now shown are as follows :—
““(1) Four specimens of Crenis boisduvalii, Wallgrn.
“These form part of the series recorded by G. A. K.
Marshall in Trans, Ent. Soc. Lond., 1902, p. 206. They consist
of one ¢ reared under normal conditions; two ? ? exposed
in the pupal condition to excessive moisture; and one 9
similarly exposed to dry heat. They accord fairly with the
seven specimens of the same series, reared under correspond-
( xv )
ing conditions, which were presented by Mr. Marshall to the
Hope Collection in 1897. The statements made (loc. cit.,
p. 209) with regard to the original seven are confirmed by the
new accessions, with the exception that the upper surface of the
male kept in normal surroundings is perhaps somewhat lighter
than that of the dry-heat male already in the Collection. The
under surface, however, of this male, and the general aspect of
all the females support the conclusion before arrived at that
the intermediate or early dry-season form of this butterfly can
be at least slightly influenced in the direction of the dry- or
wet-season phase respectively by artificial conditions of dry
heat and moisture. ;
‘“©(2) Three specimens of Pinacopteryx pigea, Boisd.
“‘These three males are the specimens referred to by Mr.
Marshall (oc. cit., pp. 206-208) as d of the first series, and
g, & of the second series. Like the previous examples result-
ing from the same experiment, which are nine in number and
all females, they tend to show that under conditions of moisture
a certain approach is made towards the wet-season form of this
species ; this, however, is less conspicuous in the present group
of males than in the females just referred to.
“©(3) A male specimen of Veracolus anne, Wallgrn.
‘This specimen is spoken of by Mr. Marshall (loc. cit.,
p. 201) as the one example which emerged out of two that
were kept as pupx for nine days in a damp jar. The present
specimen shows a nearer approach to the full wet-season form
than the example in the Hope Collection which was left under
moist conditions for seven days instead of nine. It is, however,
less ‘wet’ than another male bred at nearly the same time
under normal conditions (loc. cit., p. 202). In the case of this
species the numbers experimented with are probably too small
to warrant a definite conclusion as to its susceptibility.
‘*(4) A male specimen of Teracolus awaxo, Luc.
“This very interesting butterfly is the last of the three
individuals mentioned by Mr. Marshall (loc. cit., p. 201) as
having been bred from eggs laid by Zeracolus topha, Wallgrn.
The example previously presented to the Hope Collection is
quite of the auxo form, though not extreme; but the present
specimen, as noted by Mr. Marshall (Joc. cit.), shows a still
(¢ 9a»)
closer approach to the fullest @uxo condition, which is known
to be characteristic of the wet or summer season. The two
specimens now referred to are of great value as having afforded
the actual synepigonic proof of the specific identity of the two
phases 7’. auxo and 7’. topha. This identity had been suspected
so long ago as 1877 by Mansel Weale, who placed some of his
specimens of these forms in the Hope Collection. But, as the
writer has elsewhere stated, Mr. Marshall’s experiments, the
results of which are now at Oxford, remove the subject of the
specific identity of these two forms from the region of probable
conjecture to that of actual proof.”
Mr. W. E. Swarr exhibited a specimen of the North
American Longicorn, Neoclytus erythrocephalus, F. He said
the species had been discovered in a sound ash-tree seven inches
from the bark, grown in the neighbourhood of St. Helens,
Lancashire. Some palings of American ash in the vicinity
suggested the origin of the progenitors of the colony ; but it
was not known how long they had been erected. Mr. SHarp
also showed examples of Amara anthobia, Villa, with a series
of A. familiaris, Duf., and A. lucida for comparison. They
had been sent him by the Rev. G. A. Crawshay from
Leighton Buzzard, where they occurred not infrequently at
the roots of grass in sandy places. Thinking their identity
with A. ducida doubtful, he had sent them to M. Bedel, of
Paris, who confirmed them as A. anthobia, Villa, a species not
hitherto recorded in the British Lists.
Mr. M. Burr exhibited a number of mutilated Stenobothrus
from the Picos de Europa, Spain. He said that these grass-
hoppers were taken at a height of about 1300 metres, on turfy
ground exposed to north wind from the Atlantic, and covered with
tufts of a short, dense, tough, and spiky shrub, together with
heather. Of the grasshoppers occurring on this spot, almost
every specimen had the wings and elytra more or less mutilated,
sometimes actually torn to shreds, entirely altering their
appearance. A notable exception was St. bicolor, of which no
single specimen was found mutilated. This species also
frequently indulged in flight, which the others were unable to
do ; and he suggested that its immunity might be due to the
vitality which has enabled it to become the most abundant and
(\} xvii 2)
widespread grasshopper in Europe. Two ? @ of a species
peculiar to the mountain, S¢. cazurvoi, were also shown, one
torn, one not torn. But of the common St. viridulus almost
every specimen was mutilated.
Mr. F. W. Pierce exhibited drawings of the genitalia of
Noctuid moths, and also with the lantern a number of slides
showing the respective peculiarities of many members of the
Genus. Among other things he drew attention to the fact
that in the case of the Zweniocampide the genitalia were
widely dissimilar, while his investigations had led him to
conclude that ashworthii, at present ranked as an Agrotis,
should more properly be included in the Noctua group.
Mr. L. B. Prout suggested that the arrangement was a
physiological necessity for the separate existence of the various
species of Txniocampa, as they all observed practically the
same habits and emerged simultaneously,
A discussion followed upon the value of the genitalia for
the purpose of distinguishing species, in which Mr. G. C.
Cuampion, Mr. A. W. Bacor, and other Fellows joined.
Wednesday, April 5th, 1905.
Mr. F. Merririetp, President, in the Chair.
Obituary.
The decease was announced of Dr. ALPpHEUS 8, PAcKARD, an
Honorary Fellow, and of Mr. Atrrep Braumont, and Mr.
ALFRED PREUDHOMME DE Borrg, Fellows of the Society.
Lxhibits.
Mr. H. Sr. J. DonistHorpe exhibited a melanic specimen
of a Grammoptera, discovered by Mr. C. J. C. Pool at Enfield,
which appeared to be quite distinct from any member of the
genus recorded in Britain, though Mr. E. A. Waterhouse had
taken a similar example thirty years ago in Yorkshire. Mr.
C. J. Gahan, to whom the specimens had been referred,
considered them to be a form of G@. ruficornis, F.
PROC. ENT. SOC. LOND., 11. 1905. B
( wail)
Mr. M. Jacopy brought for exhibition a specimen of Mega-
lopus melipona, Bates, an insect which so much resembles a
bee that Bates had said they were indistinguishable in nature.
Professor E. B. Poutron remarked that the species recalled
a little black 7rigona from Borneo, recently discovered there
by Mr. R. Shelford, but in this case the mimic was not of a
stinging but of a biting model.
Mr. A. Bacor exhibited on behalf of Dr. Culpin specimens
of Papilio macleayanus and Hypocista metirius captured in
Queensland, illustrating the use of “ directive”” markings in
the Rhopalocera in influencing their enemies to attack non-
vital parts. All the specimens were taken on the same
occasion, and the fresh condition of the Papilio was in marked
contrast to the specimens of another species recently exhibited
by Prof. Poulton, to show that the tails were not always the
first portion of the wings to be lost.
It was noticeable that two out of the six specimens showed
signs of an attack directed at the lobes of the hind-wings,
situated at the base of the tails, and judging by the unset
specimens, that these lobes were roughly suggestive of a head
when the insect is at rest. In the other species eight out of
the seventeen specimens showed damage at or near the small
ocelli on the upper or under side of the hind-wings.
Mr. G. J. Arrow exhibited an example of Ceratopterus
stahli, Westw., a beetle from Australia possessing notable
powers of crepitation.
Mr. A. H. Jones and Mr. H. Row1ianp-Brown showed a
series of Hrebia alecto (glacialis), var. nicholli, Oberth., taken
by them at about 8000 ft. at Campiglio, South Tyrol, with
specimens of Dasydia tenebraria, var. wockearia, caught in the
company of the Hrebias in the same localities ; when upon the
wing the two species were not dissimilar. Mr. Jones also
exhibited examples of Hrebia melas from the Parnassus Mount-
ains, Greece, for comparison, and fine forms of butterflies found
at Mendel, near Botzen, including very large Apatura iris 9 9°,
Vanessa antiopa, and Satyrus semele.
Mr. W. J. Kaye exhibited a 2 Morpho adonis from British
Guiana together with several males. The female sex is
extremely rare, and was only definitely known in 1881,
URE)
when it was described and figured by Mr. W. L. Distant in
the Transactions of the Society for that year. Mention was
made as to how the vast difference in the sexes arose, and it
was suggested as probable that originally both sexes were blue
with a broad white band, as is the case with the allied
Morpho cypris and Morpho helena. The 3 had, however, now
lost all trace of the white band, while the ? had retained
the white band and suppressed all the blue scaling. In
Morpho cacica, 6, normally the white band was absent, but as
a rare aberration a well-defined band was present, thus show-
ing that this phase of development was frequent in the genus.
Amongst the male specimens were two that had apparently
been attacked by natural enemies. One showed a large beak-
shaped rent in one of the fore-wings, indicating attack while
the insect was sailing with outstretched wings. Another had
a large piece removed from the anal angle of both hind-wings ;
the damage on both sides coinciding when the insect was
sitting with folded wings.
Dr. F. A. Dixy exhibited the social web and pupal shells
of Kucheira socialis, Westw., together with specimens of the
perfect insect, and made the following observations :—
‘“‘ By the kindness of Professor Poulton I am able to show
the common larval habitation of the remarkable gregarious
Pierine Lucheira socialis, Westw. This is the actual nest,
from Mexico, which was described and figured by Westwood
in Trans. Ent. Soc. Lond., 1836, p. 38, and Pl. VI, figs. 1 and
2. The longitudinal incision is the one originally made by the
late Hope Professor; the two transverse cuts have recently
been added by myself, with the object of displaying the interior
of the nest more clearly. The figure in our Transactions shows
the upper part of the receptacle, and the twig which descends
through its neck, thickly covered with the pupz of the butter-
fly suspended by their tails—a most unusual mode of attach-
ment among the Pierine, though not entirely unexampled.
At some time in the period of over seventy years during which
this specimen has been preserved, the dried pupze have been
-attacked by some cabinet pest, as a result of which many have
been detached from the wall of the nest, and several have
crumbled away. Such of these loosened pupe and their
( xx)
fragments as could be collected are now cemented to a card
and shown beside the nest, while among the pups still in situ
will be seen the little bosses of silk in which the anal hooks of
the detached pupz were once engaged.
“Similar nests, presumably of this species, have been
described by many travellers, Westwood (loc. cit.) quotes from
Hardy’s ‘Travels in the Interior of Mexico’ one such account ;
and another, by A. Sallé, will be found in the ‘Annales de la
Société Entomologique de France,’ 1857, p. 20. The latter
observer, who discovered his nests on the branches of a small
Arbutus, quotes from Humboldt, Hssai politique sur le royawme
de la Nowvelle-Espagne, Paris, 1827, p. 28, under the name of
Capullos de madrogno, a description of nests which must, it
seems, also have belonged to this species. Humboldt gives a
short account of the larva, which, however, he considered to
be a ‘ Bombyx.’
‘Tn the year 1900, Dr. Dyar exhibited two of these nests,
which had been sent to Washington by Dr. Alfredo Dugés, of
Guanajuato, Mexico. Dr. Dyar at the same time showed
specimens of the larva, of which he has published a minute
description (Proc. Ent. Soc. Washington, vol. iv, 1901, pp. 419,
420). The food-plant was stated to bea species of Arctosta-
phylos. In the discussion which followed this exhibit, Dr.
Dyar said that this was the only social butterfly known to
him. It is no doubt rare in the extreme for the gregarious
habit and the construction of a common abode to be carried so
far in the case of butterflies as in the present example, but
there is possibly at least one other instance to be found
among the Pierines. I refer to MNeophasia terlootii, Behr,
which also inhabits Mexico and Arizona, and is stated by its
describer to feed on an Arbutus, the larvee forming common
habitations in which they pupate gregariously. I cannot,
however, avoid the suspicion that the nest of Hucheira has
by some mistake been attributed to Veophasia, though the
testimony of Behr with regard to J. terlooti, originally pub-
lished in 1890 and repeated in a letter to Dr. Skinner in 1900,
certainly appears to be explicit enough, This latter butterfly
is especially interesting as possessing a female which closely
resembles some of the mimetic forms of Huterpe. (See Proc.
€ fxr}
Calif. Acad. Sci., 2nd series, vol. ii, 1890, p. 91; Ent. News
of Philadelphia, vol. xi, 1900, pp. 331, 413, 538, Pl. TI, fig.
28, and Pl. XIV.) Iam not aware that the method of pupal
suspension in this species has been recorded, but the pupa of
the other species of Veophasia, viz. V. menapia, Feld., is stated
by H. Edwards to be invariably attached with the head up-
wards. (Proc. Calif. Acad. Sci., vol. v, 1875, p. 165.) Behr
is of opinion that the two known species of Veophasia should
be referred to the genus Lucheira ; this, as I have elsewhere
stated (Trans. Ent. Soc. Lond., 1904, p. 304), seems to rest
on insuflicient evidence, though there is doubtless some
affinity between the two genera, and both belong to an early
Pierine stock. When the life history of the primitive Pierines
(Metaporia, Pontia, etc.) of Central Asia is better known, it
will be interesting to see whether the social habit and com-
munity of larval shelter will be found to prevail with any of
these in a form at all comparable with those of their relatives
on the other side of the Pacific. As I have before suggested
(Trans. Ent, Soc. Lond., 1904, p. 303), it seems not at all
unlikely that the common larval habitation of Pontia (Aporia)
crategi, rudimentary as it is, and belonging only to the early
larval stages, may be a degenerate or undeveloped form of
the elaborate silken nest constructed by the not very
widely-removed Lucheira socialis.’”
Dr. W. J. Hotanp, of Pittsburg, Pa., U.S.A., said that
although he had been a Fellow of the Society for many years,
this was the first meeting which he had ever attended, and he
regarded it as a rather peculiar coincidence that the subject
under discussion should be one from the study of which he
himself had just freshly come. Continuing his remarks, Dr.
Honanp said :—“ In the early part of the past winter, the
president of one of the Mexican railways showed me some
pieces of a white silken web, remarkably tough and durable,
which represented the covering of what he called ‘a great
cocoon,’ abundant on the branches of the ‘madrofia’ trees
in the State of Durango, and always full of a multitude of
caterpillars. I asked him to write at once to Mexico and
request the superintendent of his railway to ship me a number
of these silken bags. In due course of time I received a large
(. =xii ))
erate filled with them, and, fortunately, when the bags
arrived, the caterpillars having passed the final moult, some
of them were already pupating, and I was able to watch the
process, The butterflies subsequently emerged, and as I
had imagined from the outset, the insect proved to be
Eucheira socialis, Westw., the males appearing in advance
of the females. Oviposition took place within the silken
bags in a number of instances, the females not coming
forth, and I noticed that a number of the females, which
appeared to be sluggish in their movements, did not attain a
perfect normal expansion of the wings, as if there were already
a tendency toward the development of a weakly-winged or
possibly an ultimate apterous form; a phenomenon which is
well known in the case of the ‘bag-moths.’ This abortion of
the wings, however, may have been purely an accident, but it
was rather remarkable that while all the males emerged
with finally perfect wings—and there were some hundreds of
them—a very heavy percentage of the females were either
under-sized or partially aborted, crumpled, and unable to
attain full development. Not having the proper food-plant, I
was, of course, unable to work out the life history of the
larvee, although I preserved a large number of them both in
spirits and inflated. That this insect is congeneric with
Neophasia menapia and N., terlootic I do not believe, and in
this respect I quite agree with Dr. Dixey.
“In conclusion, it may be interesting to recall that my
friend, Professor Carl Lumholz, in his recent book on Mexico,
figures a group of Indians engaged in extracting the caterpillars
from those silken tents. They are a staple article of diet
among some of the mountain tribes, and I am told that the
forests in places are literally white with the big silken webs,
many of which are five or six times greater than the original
specimen described and figured by Professor Westwood, and
now before us.
‘‘Meanwhile I have written a fully-illustrated article on
the subject, which will shortly be published, covering my
observations upon this interesting subject.”
Professor E. B, Pouuron, F.R.S., read the following note
recently received from Mr. S. A. Neave, B.A., F.E.S. The
( xmin |)
observation supplies further interesting evidence of the super-
stitious dread of larve with terrifying eye-like markings
(Trans. Ent. Soc. Lond. 1902, pp. 399-401) :—
“Petauke, Fort Jameson,
N.E. Rhodesia.
“ January 4-5, 1905. While here my boys brought me a
specimen of Sphinx larva with terrifying markings. I think
this must be the Chewrocampa osiris mentioned by Marshall
(Trans. Ent. Soc. vide supra), but I have written a description
of it in case it should prove to be different. I can add some
further interesting evidence as to the value of its terrifying
markings. It most successfully imposes upon the natives, who
will not actually handle it at any price, and say they are afraid
of its ‘eyes.’ I have asked I should think fifty natives and
they all consider the ocellated spots to be eyes. They have
also a superstition that it jumps out of the bush and upa
man’s nostrils. This is all the more interesting when we
remember that a// other species of Sphinx larve that I have yet
seen, besides many other Lepidopterous larvee, are eaten by the
natives and much searched for, being considered a great delicacy.
They have different names for different species, which shows
their great interest in them.”
“Jan. 14. I have several of the Cherocampa larve in
captivity and three have just pupated. They spin a very
slight web, with large meshes of tough silk around them. The
natives were unaware of the metamorphoses of these larve
and were very astonished to see the pupx. They are inclined,
I think, to attribute it to magic on my part!”
Paper's, etc.
The Present read the following note on experiments
conducted by him to ascertain the vitality of pup subjected
to submersion in water :—
“The popular impression, as often shown in the news-
papers, that severe winters are specially destructive to insects
is well known to be unfounded, as our native insects in the
hibernating stage will bear lower temperatures than they
are ever exposed to, but | am not aware that there is any
(| xxiv )
positive evidence as to the effect upon them of floods. As
regards many pup, which bury themselves in earth in low-
lying localities it is certain that lengthened submergence
cannot be injurious, for they must frequently be many days
under water, yet survive. It does not however follow that
pupe which habitually are high and dry (and most of the
common lepidoptera having subterranean pupe evince a
preference for such situations) would similarly escape. With
the object of learning something definite on this point, I tried
some experiments in the winter of 1901-2, supplementing
them in the winter of 1904-5.
“JT was able to obtain the following species for the pur-
pose :—Phalera bucephala, which buries without a cocoon ;
Drepana faleataria (with which a few D. curvatula were
mixed) and Selenia bilunaria, which both form close but thin
cocoons between united leaves; Cochlidion limacodes, which
forms a dense egg-shaped cocoon; and Araschnia levana,
which is naked, and, being attached, as I have observed in
the many hundreds I have bred, to deciduous nettle leaves
and leaf-stalks, must pass the winter on or near the ground
among the leaves and refuse which gather at the bottom of
the stumps of dead stems.
“The pupe were kept in unglazed earthenware jars which
were placed in an ordinary flat sponging bath which stood on
a table close to an open window facing north-west. Many
were kept thoroughly submerged during the whole of their
periods of immersion, being deprived of any opportunity of
floating by means of earth in the case of bucephala, of
thoroughly soaked moss in the other cases. In 1902 they
were thus submerged from the 2nd February onwards, the
weather being very cold, so that ice formed several times
upon the water in which they were plunged.
“The treatment was as follows :—
Not
Submerged, Submerged. Removed from the water.
bucephala 73 10 6 day by day for 12 successive days.
Saleataria 127 27 6 x5 ass 21 a 4
bilunaria 18 20 about 4 HA Pe 5 4 sa
limacodes (As 12 5 bt Hs 15 x ap
levana 84 20 5 ae i 17 < AD)
(az)
Of the bucephala submerged, 24 emerged after 1, 2, 3, 4, 5, 6,
7,10, 11 and 12 days respectively, many more afterwards ; of
the unsubmerged 8 emerged. Of the fulcataria and curvatula
at least 66 emerged after 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12,
13, 14, 15, 16, 17, 18, 19 and 20 days (10 of the pup sub-
merged had first slipped out of their cocoons, and 8 of these
emerged) ; of those unsubmerged about the same proportion
emerged ; of the limacodes, nearly all emerged; of the sub-
merged levana only 3 emerged, after 1, 3 and 9 days
respectively.
“Of the submerged bilunaria not one emerged, but of the
20 unsubmerged 13 emerged. That result was so surprising
as*compared with the other submerged pupex that I thought
the fatal consequences might have been owing to the fact that
the 18 pup had been deprived of the protection of their
cocoons (though this had not affected the fulcataria). There-
fore in the winter of 1904-5 I tried the experiment with
this species on a large scale. Of 50 submerged in their
cocoons and taken out in lots of about 4 for 12 successive
days nearly all emerged, very few being cripples, but of 23
submerged naked and taken out in from 1 to 5 days, only 2
emerged, both cripples. Of 34 other naked pup not sub-
merged the great majority have emerged, about a quarter
of them cripples, while of 50 left in their cocoons, but
unsubmerged, nearly all have emerged, very few of them
being cripples.
‘«‘ The inference to be drawn from these experiments appears
to be that in the case of subterranean naked pupe sub-
mergence for 20 days is not injurious. Of moth pupe that are
not subterranean, such as fulcataria, limacodes and bilunuria,
submergence for 12 days or more is not injurious, with the
proviso that if biuwnaria is without its cocoon it will almost
always be killed even if the submergence is only for a day.
As to levana the pupe were probably unhealthy, as shown by
the large number of the unsubmerged that died ; the numbers
of both that survived are too few for any decided inference,
but it is established that this naked pupa can survive
submergence for 9 days.
“The power of resistance of pupx to submergence seems
(( xa)
very superior to that of larve, many of which I have found
drowned after a very few hours in water. Of course the
number of species experimented on by me is much too small
for a very wide generalization ; still it has been shown to be
a fact that several kinds of lepidopterous pupe, fair samples, I
think they may be called, will survive submergence for many
days in water, and my figures show that just as large a pro-
portion survive many days of it as survive a single day. It
should perhaps be observed that the submergence of all the
pup tried was completed many weeks before the imago was
due to emerge, with the partial exception of the biuwnaria tried
in 1904-5, a very few of which, kept in a cold, fireless rcom,
emerged before and during the term of experimentalism ; it
does not follow that submergence would have been as harmless
after the pupa had got over its winter lethargy and was
moving towards emergence; this might be tested by
experiments on short-lived swomer pup.”
A discussion followed in which Mr. A. W. Bacor, Mr. R.
ADKIN, Professor E. B. Pouuron and other Fellows participated.
Mr. G. T. Beruune-Baker contributed “A Monograph of
the Genus Ogyris.”
Mr. H. A. Byart, B.A., read a paper on ‘ Pseudacraa
pogger and Limnas chrysippus; the Numerical Proportion of
Mimic to Model.”
Wednesday, May 3rd, 1905.
Mr. F. Merririecp, President, in the Chair.
Election of a Fellow.
Mr, J. Burrerwortu, B.Se., of “Comrie,” Eglinton Hill,
Plumstead, 8.E., was elected a Fellow of the Society.
Exhibitions, ete.
Mr. M. Jacosy exhibited a series of Nenarthra cervicornis,
Baly, from Ceylon, and drew attention to the curious structure
of the antenne of the ¢ those of the 2 being quite simple. It
( axa )
now being generally considered that the antenne are olfactory
organs, the complication of joints was all the more remarkable.
Mr. G. T. Porrirr exhibited specimens of Tephrosia con-
sonaria, ab. nigra, and melanic examples of Loarmia consortaria,
all from a wood in West Kent, by Mr. EK. Goodwin. These
forms were exactly on the same lines as the melanism in West
Yorkshire, and it is curious they should occur in such widely
separate localities. The two genera, however, are evidently
prone to melanism, as Mr. Porritt had now seen black or
almost black specimens of all the British species except
Tephrosia punctulata.
Commander J. J. Waker exhibited (1) two specimens of
the very rare Staphylinid, Medon castaneus, Grav., taken in
the Oxford district during the last week of April 1905; (2)
several examples of both sexes of the giant flea Hystrichopsylla
talpx, Curtis, from field-mouse nests in the same district ; and
(3) the type-specimen of the Bostrichid beetle Dinoderus
ocellaris, Steph. (taken by the late Prof. Westwood at ‘“ Little
Chelsea” previous to 1830), from the Hope Collection at
Oxford.
With reference to the last exhibited he also contributed the
following note :—
“« Dinoderus ocellaris, Steph. (Ill. Mandib. III., p. 352,
footnote [1830]), + Sp. 2, ocellaris. Nigro-piceus, subnitidus,
antennis pedibusque piceis, elytris punctatis, punctis ocellatis
in striis digestis. (Long. corp. 1} lin.) Di? ocellaris mihz.
_ Pitchy-black, slightly glossy; thorax tuberculate in front,
punctate posteriorly; elytra rather deeply punctate, the
punctures disposed in striz, distinctly ocellated, and largest
posteriorly ; antenne and legs piceous. I have little doubt of
this insect being congenerous with the foregoing, but the
antenne are broken, Taken at ‘ Little Chelsea’ in July last.
—Mr. Westwood. It is probably exotic, having been found
in a cup of coffee,”
“The British species” (of the family Bosrricuip™) ‘are
few in number, of small size, and considerable rarity ; indeed
it is not improbable that some of them are not really in-
digenous, but have been imported in timber, etc., from abroad,
as in the instance of Dinoderus ocellaris, described by Mr.
( xxviii )
Stephens, from my collection, which I found floating in a
cup of coffee.””— Westwood, Int. Mod. Classif. Insects, L.,
p- 278 (1839).
Professor E, B. Poutron, F.R.S., read the following note
on “Heliotropism in Pararge and Pyrameis,” communicated
by Dr. G. B. Lonestarr, M.D.:—“Following up my observa-
tions on the attitude at rest of Pararge schakra, Koll., made
near Simla in October 1903,* I paid a good deal of attention in
February and March of the present year to P. meone, Cram., a
butterfly that I found in varying numbers in all the parts of
Algeria that I visited. This is either a southern form of P.
ageria, L., or a closely allied species in which the yellowish
spots are replaced by fulvous. It is fond of settling on sandy
roads, rocks, walls, or the leaves of trees or shrubs, compara-
tively rarely visiting flowers. It first pitches, invariably I
might say, with its wings about three-quarters expanded, and
in the vast majority of cases with its back to the sun, the axis
of the body being rarely more than 45° to either side; im-
mediately after settling it more often than not adjusts itself,
by a quick movement, so as to make its tail point fairly
accurately tothe sun. After this adjustment, if at all, it closes
its wings over its back, and as a necessary result its shadow
is reduced to, or approximates to, a mere line. There can, I
think, be no doubt that this habit is a great protection to the
insect, since when resting on fairly flat surfaces the shadow of
a Satyrine or Nymphaline butterfly with cryptic under-side is
often more conspicuous than the fly itself.
“IT quote two cases from my notes :—
‘Feb. 8, Guyotville. Watched a specimen settle about
twenty times, The wings were always at first expanded about
three-fourths, an adjustment of its position was in most cases
made immediately, and after that the wings were raised over
the back. In about twelve cases the orientation was perfect
and the shadow a minimum ; in seven or eight cases the orien-
tation was imperfect, with a maximum error of about 45°, but
usually much less.
“ Feb. 25, Biskra. Watched one settle three or four times,
always with a shadow near the minimum.
* See Trans. Ent. Soc. Lond., 1905, p. 67 and 136.
(°° xxx)
“Only one specimen of P. meone was ever seen to settle
facing the sun ; it did so three times, a fourth time turning its
tail in the usual way. This was at Biskra on March 5th, and
I noted at the time that the sun was not shining strongly.
Two only were observed to settle with the axis of the body at
right angles to the sun. One of these appeared to be crippled
in the legs, since it alone always sat on one side. A third
specimen, feeding on laurustinus flower in the Chabet Gorge
on March 18th, did not appear to orient itself by the sun.
“Tn conclusion, except early in the day, or when the sun is
dull, or when feeding on flowers, P. meone settles with the axis
of the body turned so that its tail points more or less accurately
to the sun, therefore when the wings are raised, in the attitude
of repose (as is more common towards the afternoon), the
shadow is reduced to insignificant dimensions.
** Pyrameis cardui, L., is an abundant butterfly in Algeria.
It is not so easy to watch as P. meone, for it is a strong flier
and much more wary ; moreover, many of the specimens were
seen in exposed places during windy weather, Nevertheless,
T can say confidently that it generally settles with its tail to
the sun, though it does not do this with the regularity of
meone. 1 saw two specimens turn their faces to the sun, and
saw a third settle twice with its body axis at right angles,
though the third time it settled normally. The first two
freshly emerged specimens, at Hammam Pvihra, March 25th,
did not orient as well as those that had hybernated.
“These observations confirm those of Professor G. H. Parker
on Vanessa antiopa, L.,and on a Grapta in the United States,*
and supply a reason for the habit that does not appear to have
occurred to him, namely, concealment when in repose.”
A discussion followed in which Dr. W. J. Hotuanp said
that he had not noticed this tendency to orientation. In the
case of Colias philodice and other Pieridx, he had observed
that they assumed all manner of positions, so that it was not
safe to extend the observation presented by Professor Poulton
to such species. The Prestpent asked for information on the
resting habit of Selenia bilunaria, and remarked that it differed
in his observations from that of S. tetralunaria and the other
* Referred to Trans. Ent. Soc, Lond., 1905, p, 136.
( ‘sae j
members of the genus. Mr. C. O. Waternouse remarked that
he had noticed that butterflies always turn ¢o the sun, so as to
expose the greatest amount of surface to the warmth. Mr.G.
C. Cuampion also mentioned that when collecting in the
tropics he found that a dead butterfly exposed to the sun’s
rays at once shrivelled up. Mr. C. J. Ganan, Dr. F. A.
Drixey, and other Fellows also joined in the discussion.
Papers.
Professor L, C. Mraun, F.R.S., communicated a paper on
“The Structure and Life History of Psychoda sexpunctata,
Curtis,” by John Alexander Dell, B.Sc.
Dr. D. H. Hurcntyson gave an address on “ The Three-
colour Process as applied to Insect Photography,” illustrated
by lantern slides of British and Indian Rhopalocera, the
exhibits showing a marked advance in accuracy of wing
coloration to any yet shown at the Society’s meetings. The
PRESIDENT at the close of the Proceedings heartily con-
gratulated Dr. Hurcurnson upon the results of his work.
Wednesday, June 7th, 1905.
Mr. F. Merririevp, President, in the Chair.
Obituary.
The decease was announced of Lieut.-Col. L. H. L. Irsy,
ELS., PZ:
Election of an Honorary Fellow.
Herr Lupwic von Ganenpaurr, of the Vienna Museum,
was elected an Honorary Fellow of the Society,
Election of Fellows.
Mr. Cuartes J. Grist of “ Apsley,” Banstead, Surrey; Mr.
Vernon Parry Krrcnen of the Priory, Watford, Herts.; and
the Rev. W. Mansectt Merry, M.A., of St. Michael’s, Oxford,
were elected Fellows of the Society.
( toomxi -)
Exhibitions.
Mr. M. Burr exhibited an earwig, Apterygida arachidis,
Yers., found by Mr. Annandale of Calcutta, in a box of
specimens received from the Andaman Islands. When placed
in a small box, it was alone, but next morning there were five
larve present ; two disappeared, apparently being consumed
by the parent ; and the remaining three were those exhibited.
Mr. Burr also showed a Locustid of the family Psewdophyllidx
taken in Queensland by Mr. H. W. Simmonds, a Fellow of the
Society, among twigs and plants which it greatly resembled,
together with a photograph of the insect in its natural
position.
Mr, E. C. Bepwet showed (1) three examples of Gnorimus
nobilis, Li, taken at Woolwich on May 20th last under the
bark of an old dead cherry tree, a beetle of which Canon
Fowler writes that it seems to be becoming extinct in
Britain ; and (2) a malformed specimen of Lochmea suturalis
which had the left posterior tibia bifid for about one-third of
its length, and two tarsi, one of which had the joints
considerably enlarged.
Mr. O. EK, Janson brought for exhibition a living specimen
of Omophlus betule, Herbst, a beetle not known to occur in
Britain, found by his son near Covent Garden, and probably
imported,
Mr. W. J. Lucas exhibited one ¢ and three ? 9 of Agrion
armatum taken this year by Mr. F. Balfour Browne and sent
to him alive. This dragonfly, searce everywhere, was added
to the British list by Mr. Browne about two years ago.
Mr. G. C. Cuampton showed four specimens of the rare
Acrognathus mandibularis, Gyll., captured on the wing towards
sunset near Woking at the end of May.
Mr. Setwyn ImaceE exhibited two aberrations of Biston
hirtarius, both females, taken by himself at rest on tree-
trunks at Mortehoe, North Devon, April 23rd, 1905. The
first aberration was tolerably normal in general coloration,
but the anterior half of the fore-wings was much suffused
with fuscous, and at the costa broadly emphasized with rich
{ mex /)
black. The transverse lines on the hind-wings, all unusually
distinct, were also dark, and broad throughout. The second
aberration was semi-transparent black all over both fore- and |
hind-wings, the veins strongly delineated with black, powdered
with ochreous. All traces of transverse lines and bands were
obsolete in both wings, except a narrow pale-ochreous sub-
terminal band, and this also nearly obsolete in hind-wings.
The head and thorax were black dusted with ochreous, the
abdomen totally black, and the legs and antennz immaculate
black.
Mr. W. J. Kaye showed a number of empty pupa-cases of
Zonosoma pendularia to demonstrate the wide variation of
methods in the ‘placing of the silken girth round the pupa.
The two extreme instances were—(1) where the girth went
over the meso-thorax, and (2) where the girth was round the
fourth abdominal segment, ‘The average was close to the
third abdominal segment. Allusion was made to Prof. EK. B.
Poulton’s experiment with Pieris brassice and Pieris rape,
as given in the Presidential address delivered in January
1905. This experiment went far to prove that those species
were incapable of showing variation in this particular respect of
pupal girth, because there was a specially strengthened groove
to receive the silken strand and it was proved that the ordinary
tegument of the pupa was unable to withstand the strain, at
least when the pupa was in an unnatural position. But in
the case of Zonosoma pendularia there is no groove present
and the pupa is able to undergo development with the silk
girth placed very variously, and it is only necessary for the
pupa to be secure, the girth being in reality loose and having
no definite position rather than a variable one. In one of the
pup the excretions were at the head end, showing that the
pupa must have been placed upside down.
Professor E. B. Poutron, F.R.S8., exhibited leaves of straw-
berry, Berberis japonica, and cherry-laurel which had been
sent to him by Mr. W. B. Grove, of Handsworth, Birming-
ham. The leaves had been attacked by minute fungi which,
in the strawberry and erberis, had been identified by Prof.
S. H. Vines, F.R.S., as Phyllosticta fragaricola and Phyllosticta
japonica, respectively. The clean round holes in the laurel
( Xxxili )
leaves had been caused by a fungus identified by Mr. George
Massee as Cercospora circumscissa, Sacc.,—the ‘“ shot-hole
fungus.” The attack was local and followed by the death
and disappearance of the central portion of the leaf-tissue of
each patch, leaving a roundish or oval window outlined with
brown, sometimes in the form of a narrow line, sometimes
spreading peripherally into the leaf for a greater or less
distance. In the strawberry the edges of the windows were
somewhat ragged, but those of the other two leaves had
smooth contours, and strikingly resembled the oval trans-
parent areas upon the fore-wings of Aallima inachis, paralekta,
etc.—surrounded most conspicuously with a marginal zone of
modified colour varying greatly in different individuals as
regards both tint and breadth. Professor Pounron had
believed that these “windows” of Aallima represented holes
gnawed by larve and that the altered marginal zone repro-
duced the effect of the attacks of fungi entering along the
freshly exposed tissues of the edge. But he now desired to
withdraw his earlier hypothesis in favour of the more pro-
bable and convincing suggestion made by Mr. Grove. The
origin of the suggestion is of some interest in relation to the
meetings of their Society and other associations which pro-
mote the intercourse of naturalists. Professor Pouiton in
the course of the ‘‘ Huxley Lecture,” recently delivered by
him in the University of Birmingham, had explained his
hypothesis and illustrated it upon the screen. Mr. W. B.
Grove heard the lecture and forthwith proceeded to develop
a sounder hypothesis.
Professor Poutron also showed a photograph of the fungus-
like marks on the wings of the Oriental Kallimas, prepared
under his direction by Mr. Alfred Robinson of the Oxford
University Museum. The photograph was taken with oblique
illumination, and shows, somewhat magnified, the tall up-
standing scales which form the centre of each well-marked
patch, as well as the pronounced shadow cast by them. They
doubtless represent, in form as well as in colour, the fructifica-
tion in the centre of a patch of leaf-attacking fungus, perhaps
the very kinds which at a later stage of their development
produce the “windows” represented on another part of
PROC. ENT. SOC. LOND., 11. 1905. 9)
( jee”)
the wing-surface, perhaps some other common tropical
Cryptogam.
Dr. Kart Jordan communicated the following note upon
the Variability of the Genitalia in Lepidoptera :—
“For a long time it was the opinion of systematists that
the organs of copulation in insects were practically constant
within a species, and that therefore a form of insect which
was found to be different in these organs was a distinct
species. I demonstrated about ten years ago with Lepido-
ptera, and have since done so on several occasions, that there
is a certain amount of individual variability, and that this
variability in the organs of copulation is independent of the
variability in other organs, for instance in the wings. Indi-
viduals which are aberrant in pattern are normal in the
copulatory organs, and specimens with marked deviation from
the typical in these organs have normal wing-patterns.
Dimorphism in the wings, so strongly marked in many
Lepidoptera, is not accompanied by differences in the organs
of copulation.
**On the other hand, I have found that there is often a more
or less marked geographical variability in the organs of
copulation accompanying variability in the wings, a geo-
graphical variety of a butterfly or moth being in most cases
characterised by some distinction in the wing and the organs
of copulation. The bearing, on the evolution of species, of
this contrast of geographical and non-geographical variation
is obvious. It appears to me easy to understand why the
specimens of the same locality, which copulate together, are
on the whole the same in the organs of copulation, but it is
more difficult in the case of seasonal varieties. If the causes
of seasonal variation have anything to do with the origin of
new species, one should expect that seasonal forms, which are
often so very different from one another in the wings, were
also different in the organs of copulation. I have examined
many seasonally dimorphic species without result. Lately,
however, I have come across a solitary instance of seasonal
variability in the organs of copulation. The spring form of
Papilio xuthus is slightly but distinctly and almost constantly
different from the summer form in the ‘‘harpe” of the
(cose)
clasper, the dentate portion of the ‘ harpe” being propor-
tionately longer in the summer form than in the specimens of
the spring brood.
“This case of variability is of importance for the system-
atist as well as the evolutionist. It disposes of the opinion
still held by many systematists that a form with some dis-
tinction in the organs of copulation is a species, and it
demonstrates at the same time that a species can be split up
by natural causes into forms which differ morphologically like
species in organs of such importance for the preservation of
the species as are the organs of copulation.”
Mr. J. W. Turr remarked that among the “Skippers”
there are one or two species ranked as such on the variation
of the genital organs, and he drew special attention to one
described by Mr. H. J. Elwes and Mr. J. Edwards allied
closely with Syricthus malvx, viz. var. malvoides, Elw. and
Edw., Trans. Zool. Soc. Lond. xiv, p. 160, Pl. XXIII, figs.
27-27a. He also suggested that a more definite statement
as to whether the genitalia vary side by side with wing differ-
ences would be useful to determine whether the individual
form was merely a form or constituted a distinct species.
Mr. H. J. Etwes said that in the work alluded to Mr.
Edwards was responsible for the parts dealing with the
genitalia as a means of identification of species. As a rule
he was loth to accept the genitalia alone as proof, but in the
species of Syricthus mentioned Mr. Edwards had actually
found the genitalia distinct.
The subject was further discussed by Mr. G. C. CHampion,
Mr. M. Jacosy, the President, and other Fellows.
Dr. G. B, Lonestarr read the following observations on
scents in the male of Gonepteryx :—
‘At Hammam Meskoutine, Algeria, on March 15th, 1905,
while examining my captures towards the close of the day
prior to writing the data on their envelopes, I was struck
with the sweet scent of a ¢ Gonepteryw cleopatra, L. All
the three dead specimens which I had taken that day had
the scent, but in two it was faint. On March 19th, at
Bougie, | confirmed this in a living specimen, describing the
scent at the time as ‘sweet, rich, thick—suggesting [’reesia,’
( xxxvi )
At Hammam R’ihra I submitted living ¢ cleopatra to four
ladies ; one could not detect the scent, another could not
describe it, a third compared it to primrose, the fourth com-
pared it first to gorse, then to faint ‘Syringa’ (meaning
Philadelphus coronarius). Personally after more experience I
hesitate between Freesia and ‘Syringa.’ Altogether up to
March 31st, when it was getting over, I examined nineteen
male (f. cleopatra and found the scent quite distinct in all save
one.
“Though not as abundant in Algeria as G. cleopatra, our
more familiar G. rhamni, L., is sufficiently common, and
naturally I examined that species, or form, for scent. ‘To
my great surprise out of ten specimens examined not one had
a scent at all like that of cleopatra, indeed in most of them I
could detect no scent whatever! One day in the hotel garden
at Hammam R’ihra, I caught within a space a few yards in
extent, and within a quarter of an hour, five butterflies in the
following order :—1 ¢ rhamni, 3 ¢ cleopatra, 1 ¢ rhamu ;
all the three cleopatra had the scent, but neither of the rham.
“The one specimen of cleopatra in which no scent was
detected was tested on a day when my nasal mucous membrane
was somewhat inflamed, moreover a neighbouring pig-stye
was distinctly a disturbing element. For these reasons I do
not include in the above numbers 2 ¢ rhamnt examined
under the same conditions with negative results.
“The living Gonepteryx can be easily held by the thumb
below and the forefinger above the thorax, the wings being
expanded, and so examined without appreciable injury to the
specimen, ina way that the more delicate structure of most
butterflies renders impossible.
“These surprising results struck me so much at the time
that I took care to assure myself that I was not deceived, but
I trust other entomologists will repeat the observations when
opportunity offers. Such a difference in the scent of the two
forms must imply a physiological difference that would point
to a specific distinction.
“In North Devon on the 29th of last month I examined a
3 G. rhamni but could detect no scent.”
Dr. F. A. Drxey exhibited specimens of several forms of
( {exmvii ))
Gonepteryx occurring in the Palearctic region, and commenting
upon Dr. Longstaff’s observations, said :—
“The facts relating to the scent of Gonepteryx cleopatra
and G. rhamni which have just been laid before us by Dr.
Longstaff are of very great interest, and certainly point in the
direction of a real distinction between the two forms. I have
occasionally detected a slight scent in British specimens of G.
rhamni 4, as recorded in our ‘ Proceedings’ for 1904, p. lviii,
but nothing like what Dr. Longstaff describes in G. cleopatra.
“Tt is by no means easy to determine the relation in which
the forms of the genus Gonepteryx stand to one another, and I
hope I shall not be thought to be wasting the time of the
meeting if I say a few words on this subject.
“In the Canary Islands we have (@. cleobule, Hiibn., which
is no doubt a distinct insular form. The fore-wings in the
male are of a brilliant orange right up to the margin, while the
hind-wings of that sex and the whole upper-surface of the female
are tinged with paler orange. In G. maderensis, Feld., from
Madeira, the males more nearly resemble G. cleopatra, Linn.,
and the females are also more deeply coloured than in G. rhammnit.
G. cleopatra, as is well known, is chiefly characteristic of the
Mediterranean subregion. Its male shows a brilliant orange
flush on the fore-wings which varies somewhat in size, but is
always, I think, inferior in area to that of G. maderensis. In
Syria, Asia Minor, and the Island of Cyprus occurs the form
G. taurica, Stdgr. (@. antonia, Butl.), which is like G. cleopatra
with its orange flush in a pale, washed-out condition. G.
rhamni, Linn., occurs generally in Europe, including the parts
of the Mediterranean subregion inhabited by G. cleopatra.
Mr. Elwes records it also from the Altai. In the Levant,
and as far east as Turkestan (Elwes), is found a form, G.
farinosa, Mann., in which the wings of both sexes have a
mealy appearance, due to semi-erect scales like those which
characterise the borders of the wings in many species of
Catopsilia and Callidryas. G., nipalensis, G. R. Gray, of which
G. carnipennis, Butl., is perhaps a dry-season phase, is the
form inhabiting northern India and the Himalaya. The male
is of a deeper and richer yellow than that of G. rhamni, and
both sexes often have the orange discoidal spots, especially of
(| xxxviii )
the hind-wing, greatly increased in size. A similar form from
China has an indication of an orange flush on the fore-wings
in a still more attenuated form than G@. tawrica. This Chinese
form appears to be a seasonal phase of G. acuminata, Feld.,
which inhabits the same districts, together with Burma, and
is marked by extreme angulation of the wings. G. zaneku,
Moore, is a similar highly-angulated form from northern India.
G. aspasia, Ménétr., is an acuwminata-like type from Amurland,
and a large race of acuminata from Japan has been named
G. maxima by Butler.
‘* Additional data concerning the local and seasonal occur-
rence of these various forms are much needed. In the mean-
time certain points appear to be tolerably clear. Each of the
forms maxima, acuminata and aspasia seems to occur under two
phases ; one distinguished by a lower degree of angulation
and a comparatively smooth surface in the wings of both
sexes, and by a richer and deeper yellow in the colouring of
the males ; the other having the fore-wings strongly uncinated,
with the wings showing a mealy surface much like that of G.
farinosa, and the colouring in the male generally paler. There
appears to be some reason to suppose that these parallel phases
in the three forms have a seasonal significance, and they were
arranged by Mr. Butler in the National Collection in accord-
ance with that view; the smooth-surfaced, deeply-coloured,
less highly-angulated phase being considered to belong to the
wet season, and the other to the dry. If this be so, it is
interesting to observe that the same relation between dry
season and uncination of the fore-wing obtains in the case of
this genus as in the other instances from Pierine genera
(Pyrisitia proterpia and Teracolus auxo) to which I drew
attention in the volume of our Transactions for 1903, p. 157,
and Plate VII, figs. 1-8. It is also significant in relation to
the principle there laid down, that the most highly angulated
forms to be met with in the genus are the supposed ‘ dry-
season’ females, especially, it may be noted, those of @.
acuminata. Now whatever be the relation between the two
‘seasonal’ phases of the three forms just mentioned, I have
little doubt that the same relation holds between G. nipalensis
and G. zaneka; and I am quite prepared to find that these two
(i) eee.)
are simply phases of the same species, the more markedly
uncinate form G', zaneka belonging to the dry season, and the
more deeply coloured form G. nipalensis to the wet.
“We have then, in passing from west to east, a chain of
forms as follows:—G. cleobule (Canaries), G. maderensis
(Madeira), G. cleopatra (Mediterranean), G. tawrica (Levant),
G. nipalensis and G. zaneka (N. India), G. acuminata (China
and Burma), G. aspasia (EK. Siberia), and G. maxima (Japan).
These appear to stand to one another in the relation of more
or less distinct geographical races ; in some cases, as in that
of G. cleobule, doubtless deserving the name of representative
species. In passing from G cleobule to G. cleopatra the orange
flush diminishes in area but becomes heightened in colour ;
from G. cleopatra to G. tauwrica the area of the flush remains
the same, but its intensity is lessened. In each of the forms
ranging east of the habitat of G. taurica there are, as we
have seen, two phases, perhaps seasonal in significance ; one
of which phases, resembling G. cleopatra more closely in
contour, recalls it also by the reappearance, at least in the
far eastern forms, of an indication of the orange flush.
“What, it may be asked, is the relation of G. rhamni to
the other forms of the genus? If after studying the Asiatic
forms in their ‘seasonal’ phases, we were suddenly con-
fronted with G. cleopatra and G. rhamni for the first time, I
believe we should be inclined to regard the two latter as also
phases of each other, parallel with G. nipalensis and G. zaneha,
or with the two forms of G. aewminata. But we know that
whatever may be the relation between G. rhamni and
G. cleopatra, it is not one of regular seasonal alternation.
Are they to be regarded as completely distinct? If so, this
would seem to carry with it a presumption that the supposed
seasonal forms of G. aspasia, G. maxima, ete., have been
wrongly associated; in which case the strongly acuminate,
‘wet-season’ forms, with G'. zaneka, must be looked upon as
geographical representatives of one species, viz:, G. rhamind ;
and the less acuminate, ‘dry-season’ forms, with G. nipal-
ensis, as the same of another species, viz., G. cleopatra.
What evidence is there of a synepigonic kind? The state-
ment met with in popular books, that G. rhamni and G.
rae 7
cleopatra have been reared from the same parent, appears to
rest on a passage in Westwood’s ‘ British Butterflies and
their Transformations, 1841, p. 13, in which he mentions
that M. Boisduval had informed him ‘that he had reared
G. rhamni and CG. cleopatra from eggs deposited by the female
of the former.’ He adds a reference to Boisduval’s Hist.
Nat. Lepid., vol. i, p. 602, where, however, I find no state-
ment about synepigony, though the author points out that
the difference in shape which usually characterises G. cleopatra
as compared with G. rhamni is not of constant occurrence.
Specimens of G. cleopatra occasionally observed in Great
Britain have doubtless been introduced ; but Curtis’s figure
of G. rhamni with slight orange markings may probably
represent a really British individual. On this part of the
question, however, it will be allowed that Boisduval’s alleged
experience requires confirmation. As to syngamy, we have
the interesting testimony of Mr. Bagwell-Purefoy (‘ Entomolo-
gist,’ 1902, p. 304), who found that home-reared G. rhamni 9 ,
emerging in August, were persistently courted by G. cleopatra
3, ‘though whether any results were obtained is more than
doubtful.’ I would suggest that the reluctance of the female
G. rhamni which Mr. Bagwell-Purefoy speaks of may have
been due to the fact that this form, at any rate in the British
Islands, does not pair until after hibernation.
“In favour of the view that G. rhamni and G. cleopatra are
entirely distinct, we have the fact, explicitly stated by the
last-named observer, that his G. cleopatra reared in Ireland
under natural conditions were double-brooded, whereas G.
rhamni is well known to be single-brooded, at least in a large
part of its range. There is also the difference in contour,
G'. rhamni being much more acuminate than G. cleopatra.
The food-plant, again, is said to be different. It would seem,
however, that not one of these points is absolutely free from
doubt. It is positively stated that G. cleopatra is single-
brooded in some localities; and if it should turn out that, as
has been asserted, G. farinosa is a second brood of G. rhamni
in the eastern Mediterranean subregion, the supposed dis-
tinction would break down on both sides, Then with regard
to contour, though the difference is usually well marked, it is
( xii )
possible, as Boisduval pointed out, to find specimens of
G. rhamni which are less acuminate than some specimens of
G. cleopatra. We often find females which belong unmistak-
ably to the cleopatra type, both in form and in colour ; but
beside these we get other females which in shape appear to be
of the one form, and in colour of the other. Finally as to
the food-plant. This in the case of G. cleopatra is said to be
Rhamnus alaternus or R. alpinus. Mr. Bagwell-Purefoy’s
cleopatra, however, would not eat &. alaternus; but throve,
like rhamnt, on R. catharticus.
“Dr. Longstaff’s very interesting observation must of
course be allowed full weight. In reference to this it is
remarkable that I have not found any scales in either
G. rhamni or G. cleopatra which appear to be specialised for
scent production; nor do I find any microscopic difference
between the ordinary scales of these two forms, except in the
matter of pigmentation. Further information as to all these
forms would be very welcome, especially in the case of G.
forinosa and its possible relation with G@. rhamni and G. taurica,
in company with the latter of which forms it was found flying
by Mrs. Nicholl on the west side of Lebanon towards the
end of June (Trans. Entom. Soc. Lond., 1901, p. 81). Mean-
while, I think it may be considered as open to question
whether in a portion of their range G. rhamni and G. cleopatra
may not to some extent intergrade, though for the most part
they certainly appear to be distinct enough.
“ By the kindness of Professor Poulton I am able to
exhibit to-night specimens from the Hope Collection of many
of the forms that have been mentioned in the course of these
remarks.”
Mr. H. J. Exwes, F.R.S., read the following note on the
Geographical Affinities of Japanese Butterflies, of which he
also exhibited numerous specimens :—
‘“Twenty-three years ago (P. Z. 8. 1881, p. 856, et seq.), I
wrote my first entomological faunistic paper, ‘On the Butter-
flies of Amurland, North China and Japan,’ basing it upon
the collections of Messrs. Jonas, Pryer, and Fenton. I then
attempted to analyse the distribution of the species, which
I classified as follows :—
( ala. 4
Species common to and characteristic of the
Palearctic region ! ; oie
Peculiar to the Kastern part of ae region,
but mostly belonging to genera of Palearctic
affinity : j : \ 4 5 262
Common to Indian region, or cosmopolitan in
Old World . , : 2 ‘ ‘ . 44
1a
‘Last summer I made a short trip to this wonderfully in-
teresting country, and though my objects were principally
the study of trees and forests, I was able to collect in some
parts of Japan little known to the ordinary tourist, a number
of butterflies of great interest. Though I had carefully
studied the best Natural History books on the country I had
not in the least realised -what were the climatic conditions
which prevail in summer, which have had a much greater
influence upon the character both of the vegetation and the
insects than I could have believed without seeing it. On
many occasions I was struck by the remarkable similarity of
the forest and the forest insects, even as far north as central
Hokkaido (usually known in England as Yezo) to the middle
region of the Sikkim Himalayas at about 8-10,000 feet ; and
even in the low country in the months of July and August
there is more to remind one of India than of Europe. The col-
lections which I made, though very scanty in point of numbers,
fairly represent the character of the Butterfly fauna, which,
as you will see, contains a great number of Himalayan
genera and very few species really characteristic of Northern
Kurope and Asia. No doubt in the spring this would not
have been the case ; for though Pryer has stated in ‘ Rhopala-
cera Nihonica’ (p. 2) that the Fauna is decidedly Palearctic,
and my own analysis indicates a greater aflinity to the
Palearctic region than to the Indian, yet the species which
were most numerous in the forests almost all belong to
Indian genera; characteristic Palearctic genera and species
with the exception of Argynnis and Pieris were almost
always few in number, and the most generally distributed
and common species of Europe and Northern Asia were
( xiii)
either absent, or so much altered in appearance, that they
might almost be treated as different species. A short analysis
of my collection shows this clearly. Amongst the Satyride,
for instance, there is not a single species found in Europe
except Satyrus dryas and Pararge deidamia, the latter only
taken in Hokkaido; whilst three species of Lethe, one Neope,
one Ypthima and the large Pararge schrenkii—elsewhere only
found in Amurland, which has very much the same faunistic
vharacteristic as Japan,—were far more abundant. I did not
see an Hrebia, of which only one species is found in Japan, a
Canonympha or an Lpinephele, three of the most characteristic
Palearctic genera. In the Nymphalide, Vanessas except
V. urtice and G. album were rare in Hokkaido. Jelitxa is only
found in the higher mountains, all the smaller Argynnis, so
characteristic of Europe, are absent ; and Veptis, which is much
more an Indian than a Kuropean genus, was almost the only
abundant Nymphalid genus in Central Japan. Though Apatura
and Limenitis are both common, yet Dichorrhagia and Luripus
are quite as much so. Amongst the Pieridz it is true the
species are rather Palearctic than Oriental, but Terias is
quite as commonly seen as any of them, except in Hokkaido.
In the Papilionide the only European species is P. machaon,
whilst three or four truly Oriental forms are much more in
evidence, and may be seen hovering about the roadsides and
settling in front of the houses as in the Himalayas.
‘“* Amongst the Lycenid the only common European form
is ZL. argus or xgon. Though forms of several wide-ranging
species such as argiolus and argiades are yet present, in Central
and Southern Japan purely Oriental genera such as Curetis,
Niphanda, Dipsas, Amblypodia, and in Hokkaido the green
Theclas all give a very Himalayan character to the Butterflies.
But it is amongst the Hesperiide that the truly Oriental
character of the fauna is most strongly shown. Of the species
I show to-night only one is really European, all the rest
belonging to characteristic Indian genera; and this was
equally the case in Hokkaido as in Central Japan. In the
only day’s collecting I had in the neighbourhood of Kioto
I saw not one single European species except the ubiquitous
Colias hyale, and if I had had time to visit the southern
( ai)
islands of Kiusiu and Shikoku I have no doubt that the
same feature would have been even more striking. This is
not so much to be wondered at when one considers that for at
least five months in the year, from the beginning of June to
the end of October, the climate is really much more tropical than
temperate, and enables tropical plants, such as Caladiums,
Ginger, and Rice to be cultivated generally. It seems strange
that this remarkable fact has not been previously brought
out, or made clear by authors resident in Japan, and the
tourist who visits the country usually in the autumn and
spring does not realise the complete change that takes place
during the rainy season. Briefly, one may say that though
during the winter and spring months the plants and insects
of Japan are, like the climate, Palearctic in character, yet
during the summer and autumn they are Tropical; and
though, no doubt, we must not generalise too hastily from
such a short experience as I had, yet it serves to show that
one cannot realise the climatic and faunistic peculiarities of
any country thoroughly without having personally visited it.”
A discussion followed in which Mr. R. Trimen, Mr. J. W.
Turr and other Fellows participated.
Papers.
Professor CuristoPpHER AURIVILLIUS communicated a paper
on “ New African Zas/ocampidx in the British Museum.”
Mr. G. W. Krrkatpy communicated a “ Memoir on the
Rhynchota taken by Dr. Willey chiefly in Birara and Lifu.”
Cake)
Wednesday, October 5th, 1905.
Mr. F. Merrirrecp, President, in the Chair,
Election of a Fellow.
Mr. J. R. Davipson, of Drumsheugh Gardens, Edinburgh,
was elected a Fellow of the Society.
Obituary.
The decease was announced of Mr. Jonn Witir1am Douctas,
the oldest Fellow of the Society, who was elected in 1845 ;
Mr. GeorcE BowpLer Bucxkton, and Mr. AmBprosE QUAIL.
Honorary Degree.
The PresipENT said that since the last meeting the
University of Oxford had conferred upon Commander J. J.
Watker, R.N., one of the Secretaries, the degree of M.A.
honoris causa for his services to entomological science. After
the announcement, which was received with loud applause,
Commander WALKER thanked the Fellows present for the kind
way in which they had expressed their approval and interest.
Exhibitions.
Mr. Epwarp Harris showed some living larvee of the Lon-
gicorn beetle Cordylomera suturalis, Chevr., taken from a log
of mahogany imported from the Sekondi district of the Gold
Coast, together with the perfect insect, which was dead at the
time the discovery was made.
Mr. A. T. Rose exhibited a remarkable melanic specimen of
Catocala nupta taken by Mr. Lewis in his garden at Hornsey,
N., in September. The coloration of the lower wings was of a
dull brown, and all the markings of the upper wings were
strongly intensified. Mr. J. H. Carpenter said that a some-
what similar form had been taken by the late Mr. Mark Winkley
at Mitcham, Surrey, on August 22, 1892. (Cp. Entomologist,
xxv, p. 243.)
PROC. ENT. SOC. LOND., Iv. 1905. D
( gyi)
Mr. Norman H. Joy brought for exhibition Coleoptera taken
during a three days’ trip to Lundy Island in August.
Melanophthalma distinguenda, Con., a species new to Britain ;
Stenus ossium,var. insularis,a variety apparently new to science;
series of Psylliodes lwridipennis, Kuts., and Ceuthorrhynchus
contractus, var. pallipes, Crotch, peculiar to the island. One
hundred and sixty-three species were taken on the island,
about eighty of which are not recorded in Wollaston’s and
F. Smith’s lists of Lundy Coleoptera.
Mr. Atrrep Sicu showed examples of Argyresthia illwmina-
tella, Z., two of the four specimens taken near Hailsham,
Sussex, on June 15th this year. They were beaten off Pinus,
and until examined with a lens were supposed to be Oecnero-
stoma piniariella, of which species two were also exhibited for
comparison. Afterwards they were thought to be A. illumina-
tella, and two specimens were sent to Mr. Meyrick, who has,
by request, recorded the occurrence of the species in the
Entomologists’ Monthly Magazine.
Mr. W. J. Lucas exhibited the larva, cocoon, and the
subsequent imago of an ‘‘ant-lion,” Myrmeleon formicarius, from
two Spanish larve given him by Dr. T. A. Chapman last
autumn, ‘The difference in size between the small larva and
the large perfect insect was remarkable. He also showed a
living 9 of the rather scarce grasshopper Stenobothrus rufipes,
taken in the New Forest at the end of August, and kept alive
feeding on grass.
Mr. G. C. Cuampion exhibited several examples of Lymeay-
lon navale, L., from the New Forest, whence it had not been
previously recorded. Mr, H. St. J. DonisrHorre said that he
had been the first Coleopterist to take it there.
Mr. A. H. Jones showed series of Lycxna argus, L. (L. xgon,
Schiff.) var. hypochiona, a South European form, taken on
the 16th July last on the North Downs; also for comparison
specimens of L. argyrognomon, Brgstr. (L. argus, Esp.) from
Sierre in the Rhone Valley, to which species they have a far
greater resemblance on the under-side both in regard to
coloration and the arrangement of the spots, than to typical
L. argus.
Colonel J. W. Yersury, who exhibited specimens of
(xiv, )
Hammerschmidtia ferruginea, Fln., said that, although Mr.
Verrall (“ British Flies,” p. 671) had recorded his opinion
regarding the probable occurrence of one or more species
of this genus in the British Isles, these were the first
authentic British specimens. At Nethy Bridge this year Mr.
Lamb of Cambridge and himself obtained during the month of
June three ¢ ¢, three 9 2. ‘The insects were found resting
on aspen logs in the birch woods, and were chiefly remarkable
for the quiet manner in which they sat and allowed themselves
to be boxed. Mr. Verrall (l.c., p. 475) expresses doubts as to
the claims of Hammerschmidtia to rank as a genus. It is,
however, a good one, and perfectly distinct from Brachyopa,
though it seems to be very imperfectly known everywhere.
With regard to Microdon latifrons, Lw., which he exhibited
also, Col. YERBuRY said that at the meeting of the Society held
November 3rd, 1900, he had shown a specimen of a Microdon
taken at Nethy Bridge June 18th, 1900, which specimen he
wrongly identified as JZ. devius, and under this name it was
recorded in Verrall’s ‘ British Flies,” p. 662; it seemed advis-
able, therefore, to exhibit a second specimen from the above
locality, and to draw attention to the previous error. Only
four examples of this species have so far been obtained within
the United Kingdom. Of Chamezsyrphus scxvoides, Fln., the
single specimen shown was swept on June 15th, 1905, in
the Abernethy Forest near Forest Lodge, while regarding
Cynorrhina fallax, L., this insect occurred in some numbers
at Nethy Bridge during the same month, numerous specimens
having been taken by Mr. Lamb and himself.
Mr. H. J. Turner exhibited the imagines bred from the
larvee of three of the species of Coleophoridx, exhibited by him
during the year 1904, together with the leaves of their respect-
ive food plants, to show the character of the mines made by
the larve. The larval cases were also shown mounted in the
position they normally retain during life. The species
exhibited were C. alcyonipennella, C. lixella,and C’. albitarsella,
as well as a similar life-history of a species he had not shown
before, viz. C. badiipennella, the larve of which feed on elm.
He also exhibited the living larve and their cases of three
other species, which he had received the previous day through
( xlviii )
the kindness of Mr. Eustace R. Bankes of Corfe Castle, and
read extracts from the remarks which had accompanied them.
(1) Goniodoma limoniella, Stn. ‘The larve you will find in
plenty on the bunch of seed-heads of Statice limoniwm in their
very inconspicuous cases consisting of the old flower and
calyx: there are a few cases also on the stalks. You probably
know that when full-fed the larva descends the stalks of the
seed-head, takes up its position on some spot on the main stem
(sometimes high up, often very low down on it), bores into it,
hibernates therein, and pupates the following late spring
or early summer. The case remains outside the stem, but,
being only lightly attached, drops off in the course of the
autumn or winter, so that, although one can collect the stems
in the spring, one then gets no cases.” (2) Coleophora obtusella.
“ These larvee are very difficult to detect, the case being made
of the seed-husks of Juncus maritimus on which the larve
feed,” and (3) Coleophora glaucicolella. ‘* Larve of what I
expect will prove to be this species, on seed-heads of Juncus
glaucus, to which they show a decided preference.” All three
species were obtained by Mr. Bankes in the Isle of Wight.
Paper.
Commander J. J. WALKER read a paper by Mr. A. M. Lea
entitled “The Blind Coleoptera of Australia and Tasmania,”
and exhibited specimens of Jilaphanus stephensi, Macl., from
Watson’s Bay, Sydney, N.S.W., and Phycochus graniceps,
Broun, and P. sweipennis, Lea, from Hobart, Tasmania.
Wednesday, October 18th, 1905.
Dr. T. A. Cuapman, M.D., F.Z.S8., Vice-President, in the
Chair.
Election of Fellows.
Mr. Cuartes WitiiAM Bracken, B.A. (Lond.), of 18
Whiteford Road, Mannamead, Plymouth, and Mr. Wr1tam
Husert Str. Quentin, of Scampton Hall, Rillington, York,
were elected Fellows of the Society.
( xlix, )
Donation.
Commander J. J. WALKER announced that Mr. Charles D.
Douglas had presented to the Society a framed photograph of
his father, the late Mr. John William Douglas. <A vote of
thanks to the donor was carried unanimously.
Exhibitions.
Mr. H. Rownanp-Brown exhibited a small collection of
Erebias taken this year in the Pyrenees, including Hrebia
lefebvrei, with the varieties pyrenxa, Obth., from Mt.
Canigou, E. Pyrenees, and var. intermedia, Obth., from
Gavarnie. He also*showed for comparison Z. alecto, Hb.,
var. nicholli, Obth., from Campiglio, which at one time was
supposed to be identical with /efebvrei, then considered the
Pyrenean form of 2. melas. With them were also shown
specimens of Z. gorgone from the Lac de Gaube, Cauterets, and
from Gavarnie, and EF. gorge from the same localities. Mr.
Row1anp-Brown remarked that whereas lefebvrei appeared
to be confined to the stony detritus just below the snow line,
where it flew with gorge, he only found gorgone on the
grass pastures at an altitude of about 6,000 feet ; and there
it occurred in the Vallée d’Estaubé in some numbers on
July 23rd.
The Vicr-Prestpent, who showed a diagram of the clasps
of lefebvrei and melas, said that in a previous year of the
Transactions he had published the results of his anatomic
examinations of the species, which showed that /efeburei ap-
proached pronoe and neoridas, while the clasps of melas and
nerine (diagram also exhibited) were practically identical.
He also drew attention to the remarkable superficial resem-
blance of the under-side of the 9 gorgone to that of the ¢
goante, a species not hitherto recorded from the Pyrenees,
Mr. Rowianp-Brown included in his exhibition short series
of Lycxna orbitulus from the Centrai Alps, Z. orbitulus, var.
oberthuri, Stgr., L. pyrenaica, and L. pheretes from the Brenner
and Cortina districts. He said that with regard to the first
three butterflies his observations confirmed those made by
M. Rondou in his ‘‘ Catalogue Raisonnée des Lepidoptéres des
Pyrenées,” the orbitulus, which he found on the Col de
(9
Puymorens, closely resembling the central Alpine form, while
all those taken in the Central Pyrenees at Gavarnie and the
Lac de Gaube were the darker and more robust var. oberthurt.
He further suggested that, as between those species enumerated,
there seemed to be a greater superficial affinity between
pyrenaica and pheretes (not reported from the Pyrenees) than
between pyrenaica and orbitulus. At Gavarnie the two latter
were taken flying together, but were hardly emerged this year
in the last week of July, and no 9 @ were observed.
L. orbitulus, 8. L. pyrenaica, 8. L. var. oberthuri.
Mr. E. C. Bepwext exhibited eight specimens of Apion
levigatum, Kirby, one of the rarest indigenous Apions, found
on August 31st, sheltering under plants of Hchiuwm vulgare in
the Lowestoft district. The one ¢ taken was exceedingly
diminutive, being only about 1} mm. in size. Commander
WaAtker said that he had this year seen one example of this
beetle taken at Oxford, where it was known to occur.
Mr. R. SHetrorp showed several insects from Sarawak,
Borneo, including (i) a Lygeid bug which had been taken
burrowing in decayed wood: the fore-limbs exhibited a
remarkable adaptation for fossorial habits, comparable with
the modified fore-limbs of the mole-cricket (Gryllotalpa), which
insect in Borneo is frequently found in decayed wood ; (ii) a
Brenthid beetle, with a deep channel running along the dorsal
a a
part of the prothorax, the lips of which channel are nearly
apposed, so that the channel communicates with the exterior
merely by a narrow slit, and constitutes a nearly cylindrical
chamber ; this chamber is occupied by Acari; (iii) two Bren-
thide with deeply sulcated prothorax, in which Acari occur ;
the sulci in these two species not being ‘“‘roofed in” as in
the preceding species: and (iv) an Anthribid beetle with a
crescentic sulcus on the prothorax.
Mr. C. J. Ganan, on behalf of Mr. C. O. Waterhouse,
exhibited a living example of Phaneroptera quadripunctata,
which species had been found in some numbers in a vinery
near Chester. It is a Southern European species, and is not
known as British. But a single specimen of an allied species
Phaneroptera falcata was once taken near Land’s End by
Dr. Mason.
Mr. W. J. Kaye brought for exhibition a long variable
series of Heliconius numata from the Potaro River, British
Guiana. These included a very large number of different
forms, from some with a narrow black bar across the hind-
wing, to others with almost the whole of the wing of a deep
black. The series clearly proved that these very variable
forms were only aberrations, and were not sub-species, at least
in this locality, as had been described by Riffarth, Weymer
and others.
A pair of Heliconius silvana were also shown with two
rare aberrations, showing the black area of the hind-wing
divided; and examples of Veliconius vetustus, it being
remarkable that although similar to nwmata it was neverthe-
less a distinct species.
Mr. A. H. Jones exhibited a collection of Lepidoptera made
by him in Majorca during the first half of last June, and
remarked upon the almost total absence of Lepidopterous life
in the island. Only thirteen species of butterflies were
observed, all of the commonest kinds and without any indica-
tion of variation, with about six species of moths (all occurring
in Britain), including Agrotis saucia, Acidalia ochrata, and
A. degeneraria, the latter, interesting in point of colour,
being much redder. He also exhibited Melanargia lachesis, var.
canigulensis, from Vernet-les-Bains, showing on the under-
|
(Tay
side in the males a strong resemblance to M. galathea, also
Melitxa aurinia, var. iberica, Obth.—(desfontainii, Rbr.) from
Montserrat, near Barcelona, and a melanic specimen of Frebia
stygne, taken by Mr. R. 8. Standen last June at St. Martin
du Canigou, Vernet-les-Bains.
Papers.
Mr. Frank P. Dopp communicated a paper “ On a parasitic
Lepidopteron from Queensland, Australia.”
Commander J. J. WALKER read a paper by Mr. E. G. B.
Merapre-Watpo, ‘On a Collection of Butterflies and Moths
made in Marocco, 1900-01-02.” The species enumerated
included a Caenonympha new to science, and a Satyrus not
hitherto figured. But for soluxuriant a country as that visited
it was remarkable how few butterflies and moths there were.
Of the former only sixty-three were observed ; of the latter
one hundred, including a new sub-species of Macrothylacia rubt.
Wednesday, November Ist, 1905.
Mr. F. Merririevp, President, in the Chair.
Election of a Fellow.
Mr. J. W. H. Harrison, B.Sc. (Lond.) of The Avenue, Birtley,
R.8.0., Durham, was elected a Fellow of the Society.
Exhibitions.
The Rev. F. D. Morice exhibited (1) Panurgus moricei,
Friese, a species of bee new to science taken by him near
Gibraltar, of which it was remarkable that whereas usually
the species of this genus are entirely black, in this species
the ¢ face entirely and the ? partly were bright yellow, the
legs partly yellow, and the abdomen spotted down each side,
very much as in Anthidiwn, and (2) the unique type-specimen
of Heriades fasciatus, Friese, a ¢ of the Chelostoma group,
Cle}
taken by him at Jericho in 1899, in which again, while all
its congeners are practically unicolorous, the abdomen is
banded, as in a wasp, with yellow. A discussion followed
as to the reason of the peculiar coloration in the species under
review, the exhibitor pointing out that the colour-mimicry in
these species could not be due to parasitism, both Panurgus
and Heriades being industrious genera. He also invited any
information or suggestion which might explain the pheno-
menon, so common in various Aculeate genera, of a white or
yellow face in the ¢, represented only by two spots or streaks,
or often not at all, in the corresponding ?. Professor HE. B.
Poutton, F.R.S., expressed his opinion that the species shown,
though industrious, were mimics ; and that the question as to
the ¢ face was not at present answered, or answerable, but
that it might perhaps be ascertained by experiment in what
way it was of advantage to the ¢ to have this character which
the ? didnot need. The pale face in these and other ¢ bees
could not be explained on the lines of mimicry, but might
be used as a method of recognition. In the case of some
Algerian Aculeates the light pubescent colouring of the head
might assist as a protection to the insect when looking out
of its hole in the sunshine. The coloration of the abdomen
however suggested a case of mimicry. Mr.C. O. WATERHOUSE
mentioned that with some Buprestidz the front of the head in
the ¢ was light, but unicolorous in the ? ; a peculiarity
also observed by Mr. M, Jacosy in the genus Cryptocephalus.
Mr. W. J. Lucas showed a ¢ specimen of the earwig
Forficula auricularia taken at Warwick in September last,
with a drawing of the cerci (forceps), which were entirely
abnormal, the broader basal part of the two appearing to be
more or less fused together, while the legs of the forceps were
jointed to the basal part. The case, he said, was interesting
because in cockroaches, etc., the cerci are regularly jointed.
Mr. G. C. Cuampion exhibited various interesting insects
from Guatemala recently received from Sefior Rodriguez, in-
cluding Heterosternus rodriguezt, Cand., Pantodinus klugi,
Burm., Plusiotis adelaida, Hope, and a species of Orthoptera
greatly resembling a dead withered leaf, possibly a new species
of Mimetica.
€ Yr 5
Mr. Norman H. Joy showed two species of Coleoptera new
to the British Islands :—Lamophilus monilis, F., taken in
the neighbourhood of Streatley, Berks; and Dacne fowleri,
n. sp., from Bradfield, with specimens of D. humeralis, and
D. rufifrons for comparison, He pointed out that /owleri
differed not only in colour and structure from the two other
species of the genus, but also in habits, as it was a quick
runner.
Mr. H. Sr. J. DonistHorre showed a specimen of a new
British Agathidiuwm (Badium, Er.) discovered last year in
Cumberland, and now taken by him at Gibside, Durham, and
specimens of Prionocyphon serricornis with a drawing of the
hitherto unknown larva which he said he had taken under
water in the hollow bole of a tree, in the New Forest, last
July.
Dr. F. A. Dixry exhibited some specimens of African
Pierine butterflies, together with alcoholic extracts of the wings
of Mylothris agathina, Cram., 3, and Belenois severina, Cram., 3,
and remarked upon them as follows :—
“Tt may be remembered that some little time since (see
Proc. Ent. Soc. Lond., 1904, pp. lvi-lx) I gave a short account
of various observations made by me in 1899, and subsequently,
on the scents of several of our British butterflies, exhibiting
at the same time preparations of some of these perfumes which
still retained their distinctive qualities. During the present
year I have had the opportunity of testing many of the
African butterflies in the same way, and I now propose to
give details of some of the facts that I have been able to
observe in relation to their property of scent-production. In
this investigation I have had the great advantage of the co-
operation of Dr. Longstaff, who had already, at my suggestion,
made a careful examination of the odours of many Asiatic
butterflies, as recorded in the volume of our Transactions for
1905, pp. 61-144. Iam further indebted to him for permission
to include his own impressions of these perfumes.
“The scents now dealt with are of two kinds; these
may be roughly distinguished, in reference to their probable
functional significance, as attractive and repulsive.
“The first class is, I think, of importance in courtship as a
ee)
means of sexual attraction and possibly of recognition. The
odours belonging to this class, in all the species with which I
am here concerned, are confined to the male sex. They have
a general family likeness, recalling in most cases the perfumes of
flowers, or of various aromatic vegetable products such as lemon-
peel, spices, vanilla and chocolate. They are, I think, in nearly
every instance agreeable to the average human perception.
“The scents included in the second class are probably made
use of for defensive purposes. They occur, as a rule, in
species which we have grounds for believing to be dis-
tasteful to some at least of the usual enemies of butterflies.
They are found, as would be expected on this supposition,
in both sexes; and they are for the most part disagreeable
or even disgusting to the human observer. It is perhaps
conceivable that both kinds of odours may occur in the
same individual; though such instances, if they exist at all,
would appear to be rare.
“The Pierine scents that have been examined belong to
the first class—probably serving the purpose of sexual
attraction and perhaps also of recognition. I subjoin a list
of the African species in which they have been noted.
“Catopsilia florella, Fabr. When the tufts of hair-like
scales present in the male are exposed by separating the
fore- from the hind-wing, they emit a very strong, sweet
scent, which Dr. Longstaff compares to that of ‘tuberose’
or Freesia, and to my mind suggests jasmine. (A similar
scent was detected by Wood-Mason in the Indian C’. gnoma
[pyranthe|. See Journal of the Asiatic Society of Bengal,
vol lv., *Part' IT; 1886; No?4,"p.°37!;)
“Colias electra, Linn. The wings of a male specimen were
separated, and the costal patches of the hind-wing smelt,
with a negative result. The left costal patch was then
scraped with a penknife and the detached scales tested,
again with a negative result. The right costal patch was
then gently scraped and the patch itself smelt. I immedi-
ately perceived a distinct flowery fragrance reminding me
of heliotrope, with the added suggestion of chocolate. The
scent, which was quite pleasant, corresponded very fairly
with my recollection of that of Colias edusa, Fabr., obtained
( ay )
in a similar manner (Proc. Ent. Soc. Lond., 1904, p. lvii).
In the latter case, however, I was able to detect the odour
in the detached scales.
“Jn two males similarly examined by Dr. Longstaff, the
scent, although sweet, contained a disagreeable element
which led him to compare it to that of ‘bad sweetmeats.’
This was not the case with the specimen above recorded
by me, though I can confirm the observation with regard
to one of Dr. Longstaft’s males.
“Two specimens of Zeracolus achine, Cram., 3, emitted a
distinct flowery perfume, something like that of honeysuckle.
Another seemed to be devoid of odour; the female also was
found to be scentless.
“The scent of TZeracolus annex, Wallgrn., ¢, is faint,
but quite unmistakably present in some individuals. It is
flowery, with a slight roughness or spiciness suggestive of
heliotrope. Dr. Longstaff failed to detect a scent in one of
his specimens, but found it very faintly present in a second.
He is inclined to compare it with that of G. rape.
“In Teracolus auxo, Luc., 6, the fragrance may be strong,
in which case it resembles that of some heavily-scented
white flowers such as ‘Syringa’ (Philadelphus) or jasmine.
There is no suggestion of either lemon or rose—both of
these being odours not uncommonly met with among Pierines.
Dr. Longstaff, who found the scent faint (as I did in some
instances), characterizes it as ‘clean, flowery’; and compares
it with that of Ganoris brassice, Linn.
“ Teracolus eris, Klug, was only encountered in small
numbers. A male caught by me on the banks of the Zambesi
had a distinct flowery scent when fresh. A corresponding
observation was made by Dr. Longstaff.
“In Zeracolus omphale, Godt., ¢, 1 was invariably able to
detect an odour, always well-marked and often strong. This
odour in the first place resembled that of 7’. auxo in its
likeness to the fragrance of Philadelphus ; a possibly nearer
comparison, however, being with the white lily. But in
addition to this ‘ white-flower perfume’ there was nearly
always present an aromatic, almost ‘snuffy’ constituent of
different character, hard to define; this gave to the scent a
( Pr?)
certain roughness—not, as in G. rapx, the roughness of
sweetbriar—and suggested at one time chocolate, at another
musk, while itself quite distinct from either. Two females of
this species were found to be completely devoid of odour.
Dr. Longstaff’s results with 7. omphale ¢ were largely nega-
tive. In three specimens, however, he found a fairly strong
scent which he compares to that of Freesia or honeysuckle.
“The scent in Teracolus ione, Godt., ¢, cannot always be
detected. When present, it is sweet, flowery, and apparently
free from all traces of ‘ snuffiness.’
“ Eronia cleodora, Hiibn., 3. In this form the intensity of
the odour varies greatly; the scent is sometimes absent
altogether, but when present may be strong. Its basis isa
flowery perfume like that found in many species of Teracolus,
but to this is superadded an aromatic element giving a distinct
suggestion of sweet chocolate. Dr. Longstaff, who found the
scent very faint in all his specimens, considers it to belong to
the G. brassicx type.
“ Belenois gidica, Godt., 3, appears in many instances to be
without perceptible odour. Occasionally, however, I found it
to possess an odour of roses, which in one of my specimens was
strong. Dr. Longstaff’s results were mainly negative, but in
a few instances he reports a slight, sweet flowery perfume.
“ A specimen of Belenois mesentina, Cram., 5, emitted, when
fresh, a scent much like that of B. gidica. In a second
specimen no odour was detected.
“ Belenois severina, Cram., ¢ shows, like Hronia cleodora,
much individual variation in the matter of scent. An odour
is, however, nearly always present in greater or less intensity,
and to my mind is of the same general character as that of
G. rapx, giving a decided suggestion of sweetbriar. Dr.
Longstaff, who also finds great individual differences in
intensity, considers the scent to be ‘like that of G. brassice,
but stronger and more luscious.’ I have tested the female,
with negative results.
“ Belenois thysa, Hopff., ¢, has a strong, distinct odour,
which appears to me to be like that of roses. Dr. Longstaff
notes it as a very Sweet scent; in one case comparing it to
that of Scilla nutans, in other cases to that of Freesia.
( viii)
“The scent of Belenois zochalia, Boisd., ¢, is not as a rule
very powerful. It seems to me to be of the sweetbriar order,
including the scent of the flower as well as of the foliage. It
was more distinct in a worn specimen than in two fresh ones
examined on the same day. Dr. Longstaff found a ‘faint
flowery scent like that of G. brassicx’ in the single individual
he tested.
“ Pinacopteryx charina, Boisd., 3, has a perfume like a whiff
of mignonette, seldom strong and often absent. Of nine males
tested at different times by Dr. Longstaff, only one had a
decided odour, which he records simply as ‘ flowery.’
“In Pinacopteryx pigea, Boisd., ¢, the scent is distinct, in
some cases strong. It appears to me to resemble honeysuckle
—a comparison in which Dr. Longstaff concurs. One of my
specimens had an odour more suggestive of Philadelphus, and
the scent of one of Dr. Longstaff’s was noted by him as ‘sweet
and spicy.” The female was found by me to be odourless.
“The scent of Mylothris agathina, Cram., ¢ , though varying
somewhat in intensity in different individuals, is generally one
of the strongest and most distinct in the whole range of
Pierine odours so far as I am acquainted with them. To both
Dr. Longstaff and myself it appears to be exactly that of
sweetbriar. It is a pleasant and refreshing perfume, and is
easily perceptible even when the butterfly is held in the hand
unrubbed. The female was always found by me to be scent-
less ; but one tested, after death, by Dr. Longstaff, had a
strong ‘mousy’ odour, like henbane ; an observation which I
can confirm.
“The scent of Mylothris riippellii, ¢,is in the opinion of
both of us indistinguishable from that of I. agathina, ¢. A
female specimen, when dead, was noted by Dr. Longstaff as
having a slight ‘black-beetle’ odour ; this, however, I failed
to detect.
“Tn nearly all the specimens of Mylothris trimenia, Butl., 3,
which I was able to test, there was a strong, pleasant,
characteristic odour, not easily described, and quite distinct
from that of I agathina. It contains a predominating
sweet flowery constituent, perhaps a little like that of Pina-
copteryx pigea, but reminding me rather of sweet-pea than of
@ liz»)
honeysuckle. Besides this there seems to be an element of
sweetbriar, but without the characteristic roughness—
recalling rather the rose than the foliage of that plant. The
scent also conveys a suspicion of orange or lemon-peel—nearer
orange. The scent of nearly all Dr. Longstaff’s specimens
was noted by him as being like that of clover ; in one, how-
ever, it was thought to be like that of G. rapx. The female
was found by me to be scentless.
““Only one specimen of Glutophrissa saba, Fabr., 3, was
tested ; no odour was detected.
“A few specimens of Synchloé hellica, Linn., § , examined by
me gave a distinct, though not very strong scent like that of
gorse. The females were odourless, as indeed were many of
the males. A male tested by Dr. Longstaff was recorded by
him as having ‘a very slight, heavy, flowery odour.’
“‘Tt is worthy of notice that although in all these species
the intensity of the scent varies, sometimes greatly, as between
different individuals, this variation bears no definite relation
to the condition as to freshness of the specimen examined. I
have no clear evidence that the scent is discharged more
copiously during courtship; though this may quite possibly
be the case. The perfume may persist for some little while
after the death of the insect, but has generally become much
impaired or has disappeared altogether by the time that the
butterfly is stiff. So long as stiffening has not taken place,
the scent is more easily detected and estimated at home
than in the field. Rubbing the wings is in most cases un-
necessary.
“T exhibit this evening spirit extracts of the wings of
Mylothris agathina and Gelenois severina, which will be found
easily distinguishable by the sense of smell from each other
and from the pure spirit. On a future occasion I hope to
say something of the odours as they exist in other groups of
butterflies.”
A discussion on the presence and use of scents in various
orders of insects followed, in which the President, Professor
Poutton, Col. C. T. BineHam, and other Fellows joined,
ae
Papers.
Mr. P. I. Latuy, F.Z.S., communicated “A Contribution
towards the knowledge of African Rhopalocera.”
Col. C. T. BrncHam communicated a paper entitled “ A New
Species of the Hymenopterous genus Megalyra, Westwood, by
J. Cuester Bravery, Ithaca, N.Y., U.S.A.”
Wednesday, November 15th, 1905.
Mr. F. Merririetp, President, in the Chair.
Obituary.
The decease was announced of Captain FREDERICK Wo L-
LAston Hurron, F.R.S., Director of the Canterbury Museum,
Christchurch, New Zealand.
Election of Fellows.
Mr. W. R. Dewar, Government Entomologist, Orange
River Colony; Mr. Witi1am Groree SHeELpon, of Youl-
greave, South Croydon ; and Mr. Francis C. Wooppripes, of
Northeroft, Cornwall Road, Uxbridge, were elected Fellows
of the Society.
Nomination of Officers and Council for 1906.
Mr. H. Rowianp-Browy, one of the Secretaries, announced
that the Council had nominated the following Fellows to
act as Officers for 1906:—President, Mr. F. Merrirrerp ;
Treasurer, Mr. A. H. Jones; Secretaries, Mr. H. Row1anp-
Brown, M.A., and Commander J. J. Waker, M.A., R.N.;
Librarian, Mr. G. C. Cuampton, F.Z.8. ; and as other Members
of the Council, Mr. G. J. Arrow, Mr.’ A. J. Cuirry, M.A.,,
Mr. J. E. Contin, Dr. F. A. Dixny, M.A., M.D.; Mr. H.
Goss, F.LS., Ma, W. J. Kaye Mr Ws Lots eae,
CY Tal )
Professor E. B. Poutton, D.Sc., M.A., F.R.S., Mr. L. B.
Prout, Mr. E. Saunpers, F.R.S., F.L.S., Mr. R. 5. Sranpen,
F.L.S., and Mr. C. O. WATERHOUSE.
Exhibitions.
Mr. G. J. Arrow exhibited a flower-frequenting beetle from
the Transvaal, illustrating a remarkable device for the cross-
fertilization of flowers, one of the front feet being tightly
clasped by the curiously-formed pollinia of an Asclepias. Mr.
Arrow remarked that he had seen no similar instance amongst
Coleoptera, although in some insects the phenomenon was no
doubt familiar to those who had had opportunities of observ-
ation in the countries affected by these plants. Professor EK, B.
Povuron stated that he had often found such pollinia attached
to Hymenoptera.
Mr. W. J. Kays showed a remarkable specimen of Agrotis
tritici, bearing a close resemblance to A. agathina. It had
been taken this year at Oxshott flying over heather in company
with agathina, and was a good example of syncryptic resem-
blance brought about by the common habit of resting on
heather. The specimen agreed with agathina in ground
colour ; in the pale costa; in the distinct dark wedge-shaped
marks near the outer margin ; and in the distribution of the
stigmata. These characteristics are simply a strong develop-
ment of characters which recur in the species, and it was the
simultaneous development of all of them which gave the insect
its strong likeness to A. agathina.
Mr, W. J. Lucas exhibited a specimen of Forficula auricu.
laria taken by Mr. R. A. R. Priske at Deal in September
1905, having the left cercus normal, while the right was that
of var. forcipata.
Dr. F. A. Dixsey exhibited some specimens of African
Pierine, and remarked upon them as follows :—
“The recent visit paid by Dr. Longstaff and myself to
South Africa fell, so far as Natal and Rhodesia are concerned,
well within the limits of the dry season. I have brought
here for exhibition representative specimens of four species
of Yeracolus, viz, 7. annx, Wallgrn., 7. omphale, Godt.,
PROC. ENT. SOC. LOND., Iv. 1905. E
( ixii )
T. achine, Cram., and JT. auxo, Luc., all taken by us in those
localities during August and September of the present year.
Wherever possible, I have included the male and female of
both the Natal and the Rhodesian form of each species
exhibited. I also show, for comparison, male and female
specimens of the same four species, taken in the same localities
as our own, but during the wet season of the year instead of
the dry. As none of the specimens exhibited was selected for
the purpose of proving any point, but each is a fair repre-
sentative of the category to which it belongs, the whole
assemblage may be taken as an illustration of the general
correspondence of the variation of these particular forms with
the meteorological conditions prevailing at the time of year
when they are respectively found. So different in appearance
are the ‘wet season’ and ‘dry season’ phases of these butter-
flies, that, as has also happened in many other cases, they
have been described under different names, and are even now
pretty generally ranked as separate species. It may therefore
not be amiss to recall the fact that in the case of three out
of the four it has been absolutely proved by Mr. G, A. K.
Marshall, who has bred them through, that the two seasonal
forms are conspecific (see Trans. Ent. Soc. Lond., 1902,
pp. 200, 209-212). ,
“There remain two further points that seem worthy of
note. One is that, as will be easily evident on inspection,
our Natal forms, though distinctly of the ‘dry season’ type,
are appreciably less extreme in that direction than are our
Rhodesian representatives of the same species. This corre-
sponds with the much more markedly dry meteorological
conditions that we found prevailing in the latter locality.
The other point is that, even in cases where the wet-season
males of two species, such as 7’. omphale and 7’. achine, are
quite unlike one another, the resemblance between the dry-
season males of the same two species may be so close that
in the field they can only be distinguished with difficulty.”
Colonel J. W. YersBury said that in his opinion the term
‘‘seasonal” when applied to developments of this kind was
misleading. The variation of the forms seemed to depend
not upon fixed seasonal conditions, wet or dry weather, but
(| Ixiii )
upon the amount of moisture and light to which the species
were subjected, probably in the pupal stage.
Professor E. B. Pounron understood that the term ‘ season ”
as used in this connection, referred to the climatic conditions
which prevailed at the time. He considered that “ wet
form” and “dry form” were more accurate expressions than
“wet season form” and “ dry season form.”
Mr. Setwyn ImaceE exhibited a 3 specimen of Tortrix
pronubana, Hb., taken by Mr, Harold Cooper in his drawing-
room at Kastbourne, either at the end of September or the
beginning of October last, and sent to him on October 12th.
The insect, which is new to his British list, is placed by
Staudinger in his Catalogue next to 7. viridana, L., and is
reported by him from South Europe, North-West Africa, and
Asia Minor. It is about the size of J. bergmanniana, L.,
the upper wings of a soft purplish-brown, the lower a rich
orange terminally suffused with fuscous.
Commander Waker said that the capture this year of
another specimen of pronwbana had been reported in the
Worthing district, and suggested that there might have been
an immigration.
Mr. Epwarp Harris exhibited a long series of Hemerophila
abruptara bred for the purpose of determining the effects of
heredity through successive generations. He said :—
“ On November 2nd last year I showed a small brood of
abruptaria including a few melanic forms (Proc. 1904, p. lxxii).
The two pup that went over emerged April 4th and 5th this
year, one a light male and the other a light female. Both are
remarkably larger than those that emerged in 1904 from the
same brood. From the brood of larve, also mentioned, 57
pupated and the following imagines emerged between March
24th and May 6th, 1905 :—
12 dark males, 16 dark females, 5 light males, and 6 light
females.
“The remaining 18 did not break through their silken cocoons,
but upon examination I think they would have been 5 dark
males, 6 dark females, 3 light males, and 4 light females.
‘* From the above moths I paired dark male with dark female,
light male with light female, dark male with light female, and
( tny 3
light male with dark female, of which the following is the
result :—
“The dark male and dark female have thrown all dark, 34
males and 34 females ; 9 pup have not changed.
“The light male and light female have thrown all light, 9
males and 9 females ; 3 pupx have not changed.
“The dark male and light female have thrown 8 dark males
and 16 dark females, 3 light males and 3 light females; one
pupa has not changed.
‘The light male and dark female have thrown 19 dark males
and 15 dark females, 7 light males and 8 light females ; 13
pupe have not changed.
“The Jast moth emerged November 6th.”
Paper.
Commander J. J. WALKER communicated a paper entitled
“Hymenoptera Aculeata collected in Algeria by the Rev.
A. E. Eaton, M.A., and the Rev. F. D. Morics, M.A.; Part
ii, Diploptera,” by Epwarp Saunpers, F.R.S., F.L.S.
Wednesday, December 6th, 1905,
Mr. F. Merrirrevp, President, in the Chair.
Election of an Honorary Fellow.
Doctor O. M. Reuter, of Helsingfors, Grand Duchy of
Finland, was elected an Honorary Fellow of the Society.
Election of Fellows.
Mr. Caartes Witiram Matty, M.Se., Associate of the
Society of Economic Entomology of Washington, U.S.A.,
Government Entomologist for the Eastern Province of the
Cape Colony, and Mr. Harotp Powe tt, of Rue Mireille,
Hyéres, France, were elected Fellows of the Society.
( tev 2
Nomination of Officers and Council for 1906.
Mr. H. Rowianpv-Brown, one of the Secretaries, again read
the list of Fellows nominated to act as Officers and as other
Members of the Council for 1906.
Awditors.
The PresipEeNtT announced that the following Fellows had
been appointed Auditors of the Society’s Accounts for the
current year. Mr. R. Apxiy, Dr. T. A. Capmany, Mr. R. W.
Luoyp, Mr. L. B. Prout, Mr. H. Rownanp-Browy, and Mr.
R. Sours.
Lahibitions.
Dr. K. Jorpan exhibited a series of varieties of the
Mediterranean Curabus morbillosus, showing all intergradations
from the ordinary morbillosus with broad prothorax and costate
and catenulate elytra to the Moroccan awmonti which has a
narrow thorax and smooth elytra. It is one of the most
striking cases of geographical variability.
Mr. H. St. J. DontsrHorre showed specimens of Péinus
pusillus, Sturm, recently discovered in a corn factor’s shop at
Edmonton. This species, which is common in France and
Germany, has not been recorded hitherto in Britain.
Mr. A. J. Carrry exhibited (1) a hermaphrodite Proctotrupid
belonging to the Spilomicrine, Aspn., probably one of the
Ceraphronine—a hermaphrodite of Diapria verticillata is men-
tioned in Marshall’s Catalogue, but the specimen exhibited
appeared to belong to the other subfamily of the Diapriide ;
(2) a sand-wasp without wings taken by Mr. Poole, running
on a beech trunk, and probably Didineis lunicornis, Fab. ; (3)
the 3$ specimen of Apion semivittatum, Gyll. (germari, Walt.),
taken many years ago by Mr. Walton near the Tivoli Gardens,
Margate, together with a 9 specimen of the same species
discovered while sweeping long grass near the Chequers Inn,
Deal, on September 26th, 1904, on which occasion also Mr.
Tomlin had made a similar capture.
Mr. F. B. Jenntnes exhibited (1) a ¢ and 9 example of
the Dipteron Helophilus transfugus, L., taken from thistle-
heads in the marshes at Edmonton last July, and (2) a
( ‘levi )
specimen of Stenopteryx hirundinis, a parasite on swallows and
martins, found on Box Hill, Surrey, in August.
Mr. G. T. Porrirr brought for exhibition specimens of
Odontopera bidentata ab. nigra, and stated that the form was
rapidly increasing in the Wakefield district of South Yorkshire.
Apparently, in a few more years it would oust the ordinary
form prevailing there completely.
Dr. F. A. Dixry exhibited examples of five species of South
African Pierine, in further illustration of geographical and
seasonal variation. The exhibit, which was supplementary to
that shown on November 15, comprised specimens of Lvronia
cleodora, Hiibn., from East London and Natal, Belenois severina,
Cram., from Natal and Rhodesia, Teracolus speciosus, Wallgrn.,
Pinacopteryx pigea, Boisd., and P. charina, Boisd., all from
Natal. In every instance the contrast between forms captured
during the dry months of the year and specimens of the same
species} taken in the same localities during the rains was very
distinct. Two points were mentioned as being especially
worthy of note. The first was the extremely “ dry ” character
of the examples of H. cleodora taken both in August and
September at East London as compared with those taken in
August at Durban. The difference was so well marked and
so constant as to suggest the existence in the region south-
west of Natal of a distinct local race. The second point was
the appearance presented by the “dry season” examples of
B. severina from Rhodesia, in which the dark veining of the
under-side of the hind-wing tended to become faint or to
disappear ; this disappearance of the veining being in Natal
specimens a mark of the “ wet season” phase of the species,
while the “dry season” forms from Durban were in nearly
every instance heavily veined beneath. Dr. Dixey further
remarked that the seasonal relations of the various forms of
B. severina had always been somewhat of a puzzle, and the
difficulty seemed to be increased rather than diminished by a
consideration of the present specimens. More data were
needed in regard both to this species and to the nearly-allied
B. mesentina, before any satisfactory conclusion could be
reached. With regard to the exhibit as a whole, he was well
aware that by selecting instances it was often possible to make
( Ixvii_)
out a better case than the facts really warranted. He had,
however, done his best, in arranging the present series, to
choose not extreme, but average representatives of each form
of the various species exhibited.
Mr. O. E. Janson exhibited a ¢ and 2 specimen of Orni-
thoptera chimera, Rothschild, and some remarkable species of
Delias collected recently by Mr. A. S. Meek in the mountain
region of British New Guinea. With regard to the former he
observed that it was the only known Papilio with a ringed
body, while the strongly pubescent body of the 2 was also an
exception to the general rule.
Commander J. J. Wa.ker, on behalf of Mr. A. M. Lea,
Government Entomologist of Tasmania, showed a specimen of
the Buprestid beetle Cyria imperialis, Don., having, in addition
to the normal fore-leg on the left side, two supplementary
fore-legs originating from separate coxe. One of them was
equally well-developed with the normal leg ; the other some-
what shorter, owing to abbreviation of the tibia and tarsus,
all the parts in these legs being present.
Mr. G. C. Cuampron exhibited living ¢ and ? examples of
Tetropium crawshayt, Sharp, bred by the Rev. G. A. Crawshay
from eggs deposited in July last in the bark of larches at
Leighton Buzzard.
Mr. E. R. Bankes showed (1) the specimen, unique
as British, of Depressaria emeritella, Stn., on which the
species was added to the British List by Stainton (Sup. Cat.
Tin. Pter. p. 4) in 1851; (2) a specimen of Cerostoma
asperella, L., captured by Mrs. Hutchinson near Leominster,
on September 21st, 1881, and only taken, as regards Britain,
in Dorset (not since 1831), and Herefordshire very rarely ;
and (3) various specimens labelled by Haworth himself,
showing the method of explaining his identification of the
species, noticed by him in his ‘‘ Lepidoptera Britannica,”
published 1803-1828. The exhibitor hoped that a more
general acquaintance with these labels would lead to the
rescue, from oblivion or destruction, of other of these valuable
Haworthian types and identifications.
Mr. A. Bacor exhibited a series of Vryphxena comes repre-
senting three generations, bred by Messrs. J. E. Gardner,
( Ixviii_ )
A. Harrison, T. H. Hamling, L. W. Newman, L. B. Prout,
the Rev. G. H. Raynor, and himself.
PARENTS.—The parents were selected from moths reared
from wild larve collected at Cluny, Aberdeenshire, by Mr.
James Duncan in the spring of 1904. And these, together
with the remainder of the moths reared from these larve,
were also exhibited.
From the wild larve three melanie specimens alone were
reared out of about fifty ; these were all males. Among the
non-melanic specimens one was of a very dark slate colour,
all the remainder being paler, comprising very varied shades
from dull grey through ochreous and bright red to a deeper
almost chestnut red.
Only the bright red and melanic forms, and the dark slate-
coloured specimen above referred to were used as parents.
The red forms, save for the purposes of a very careful
classification, are uniform, but the melanic specimens reared
include a range from rusty red powdered with black scales to
velvety black; there is, however, a quite definite line of de-
marcation between “ red” and “ melanic” forms, no doubt as to
the correct position of any particular specimen having occurred.
In contradistinction to Mr. Prout’s Forres race (Ent.
Record, vol. xv, p. 217 ; vol. xvi, pp. 1-5) there is no trace of
melanic suffusion on the hind-wings of any of the specimens
of Cluny parentage.
The first generation took from the time of hatching in July
1904 until about the middle of January 1905 to complete
their cycle; the second generation, December 1904 to April—
May 1905, except those reared by the Rev. G. H. Raynor,
which were allowed to develop more slowly, emerging in July
and August 1905. The third generation, and last,* occupying
from May to September 1905.
* Pairings between moths of the third generation were very difficult to
obtain, and when obtained there were usually very few, if any, ova laid
by the female. In two of the three instances in which fairly normal
numbers were laid only three or four larvee hatched, and these did not
survive the first moult. The one exception, a pairing between moths of
the B x C stock, resulted in a fair-sized brood of larve, but they were
all weakly and died off before their fourth moult.
In this connection it must be remembered that the B x C stock was
less pure than the others as it resulted from a cross between two different
stocks in the first generation, whereas all the other stocks arose from a
single pair of the moths reared from the wild larve.
( ‘tbsex: 4)
During late autumn, winter, and early spring, those reared
by myself were fed for the most part on cabbage and brussels
sprouts with some admixture of dock. During the summer
the larve were supplied with various low plants as: well as
sallow and willow, but no cabbage.
The imaginal life of the species is long with healthy
specimens, varying according to conditions of temperature
and moisture from three weeks to a month in the summer,
and considerably longer during the winter—of course within
doors—the females being longer lived as a rule than the
males.
Pairing in my experience does not take place in less than
a week from date of emergence under the most favourable
circumstances (warm moist summer weather) with the males,
and from two to three weeks in the case of the females. In
one or two instances the early death of the specimen selected
for male parent necessitated the careful choice of a second as
nearly identical with the first as possible, in one case that of
Brood C 2, a third male had to be introduced, fortunately
owing to the peculiarity of the pairing habit above referred
to, and the fact that the replaced males became paralyzed or
died within two or three days of emergence, I feel quite
confident that only the one pairing took place.
The experiment was not made in the first place with a view
to testing the Mendelian theory, but in order to ascertain the
relative proportions of melanic to non-melanic forms, and the
possible range of variation to be obtained from a single pair.
Apart from the relative strength or weakness of the melanic
strain the heredity from parents to offspring would seem to
be very strict, and this result is quite in conformity with
previous experiments conducted with the same species, and
also with other Noctuid moths in which great diversity of
forms can be obtained from a very restricted area. This
purity of strain suggests how easily a series of local races
developing into sub-species might be caused by isolating
factors such as diverging habits, etc.
But the result is in contrast with experiments in rearing
broods of Malacosoma castrensis when a very wide range of
forms was obtained from a single batch of eggs or from a
(plea? “)
pairing between a particular male and female. And Mr.
Prout informs me that this is sometimes, though not as a rule,
the case with the Geometrid stirps.
All the results of the second and third generations seem
to be capable of Mendelian explanation if we call the melanic
strain the Dominant and the red forms Recessives. For this
purpose all the parents of the melanie broods of the second
generation must be assumed to be Dominant Recessives, while
the male parent of brood C 7 x C 7 (third generation—
(melanic male paired with a red female) must be assumed to
be a pure Dominant, as all the offspring were melanic. And
again both the parents of brood C 9 x C 7 (third generation)
must also have been Dominant Recessives. On the other
hand, at least one of the parents of the remaining melanic
broods of the third generation, broods GC 9x C 9 and
C 7 x C 2, must have been for Mendelian purposes a pure
Dominant.
The red parents throwing all red offspring will of course
be Recessives. So far all is straightforward, but if we
attempt to apply the principle backwards instead of forwards,
we come to a serious difficulty in respect to the first genera-
tion, as the results do not tally with either the supposition
that the male parent was “pure Dominant” or Dominant
Recessive.
FIRST GENERATION
ACTUAL RESULTS CORRECT MENDELIAN PROPORTIONS
Brood B. Male parent dark slate (non-melanic) If male D. R. x female R. Result should be
Female Red (non-melaniec)
Result. Most of the larvee were lost owing to an
accident but one slate-coloured speci-
men and one red one were bred. The
“Red” specimen of this brood was the
male parent of Brood B x C.
Brood C. Male Melanie. Female red, Or if male D.D. x female R. Result should be
Result, 21 Melanic(=40%) 32 Red (=60%) 100% Melanic.
50% Melanic, 50% Red.
SECOND GENERATION
Brood C 2. Male Melanic. Female Melanic. Brood C. If both parents D. R. Result should be
Result. Bacot 60 Melanie 26 Red. 75% Melanie, 25% Red.
Gardner 21 Melanic 5 Red.
Prout 17 Melanie 5 Red.
98 = 737 36 = 277
( vie )
ACTUAL RESULTS
Brood C 7. Male Melanie. Female Melanic.
Result. Bacot 50 Melanic 14 Red.
Newman 8 Melanic 1 Red.
Raynor 22 Melanic 4 Red.
80 = 81% 19 = 19%
Brood C 9. Male Melanie. Female Melanic.
Result. Bacot 19 Melanic 8 Red,
Prout 15 Melanie 8 Red
34 = 68% 16 = 32%
Brood C 8. Male Red. Female Red.
Result. Bacot 19 Red
Gardner 10 Red
29 = 100%
Brood B x C. Male Red. Female Red.
Result. Bacot 131 Red
Gardner 77 Red
Raynor 48 Red
256 = 100%
Brood C7 x C 2.
Result. Bacot
Male Melanic.
CORRECT MENDELIAN PROPORTIONS
Brood C7. If bothparents D.R. Result should be
75% Melanic, 25% Red.
Brood C9. If both parents D. R. Result should be
75% Melanic, 257% Red.
Brood C3. In agreement.
Brood B x C. In agreement.
THIRD GENERATION
Female Melanic.
24 Melanie
Harrison 21 Melanic
Brood C9 x C7.
Result.
Brood C9 x C9.
Result. Bacot
Brood C7 x © 7.
Male Melanic.
Prout 16 Melanic(=76%) 5 Red(=24%)
Male Melanic.
Male Melanic.
Harrison 15 Melanic
45 = 100%
Female Melanic.
Female Melanic.
7 Melanic (=100%
Female Red.
Hamling 2 Melanic
Result.
Brood (B x C)x (Bx C)
Result. Bacot
17 = 100%
Male Red. Female Red.
20 Red = 100%
Brood C3 xC8. Male Red. Female Red.
Result. Bacot 6 Red = 100%
Brood C7 x C2. Ifone parent D. D. In agreement
Brood G7 x C9. If both parents D. R. Result should
be 75% Melanic, 25% Red.
Brood C9 x C9. If one parent D. D. In agreement
Brood C7 x C7. Ifmale parent D. D. In agreement.
Brood (B x C) x (B x C). In agreement.
Brood (C3 x C3), In agreement.
( tse 9
Mr. R. Saetrorp exhibited the larve of Collyris emargin-
atus, Dej., and of Mormolyce, and read the following notes :—
‘““In ‘ Mededeelingen uit ‘Slands Plantentuin,’ xliv, 1901,
p. 118, Dr. J. C. Koningsberger, of the Buitenzorg Zoological
Museum, publishes a brief notice of the larva of the Cicindelid
beetle, Collyris emarginatus, Dej., burrowing in the twigs of
coffee-shrubs. I noticed a preparation illustrating this remark-
able habit for a Cicindelid larva in the museum at Buitenzorg
in March of this year, but it was inside a locked case, and as
Dr. Koningsberger was on leave in Europe, I was unable to
make a close examination of the larva and its burrow. In
answer to a request for material and information on the
species, Dr. Koningsberger has kindly sent me the specimens
which I-now have pleasure in exhibiting to this Society. Dr.
Koningsberger tells me that the larva feeds on the ants and
aphides that crawl over the coffee-twigs ; pupation takes place
in the burrow ; ovipositio# has not been witnessed, nor have
any but full-grown, or nearly full-grown, larvee been found, so
that it is not known if the burrow is enlarged to allow of the
increase in size of its occupant, or if it is originally made large
enough to accommodate the larva throughout its life. A
figure of the larva is published in the above-mentioned work
(Fig. 59), but it is evidently only a copy of the figure of a
Cicindelid larva in Packard’s “‘ Guide to the Study of Insects,”
and is quite inadequate. The Collyris larva differs from the
larvee of the British Cicindelx by its flattened femora, small
size and more cylindrical body ; the tubercles of the fifth
abdominal segment bear a comb of three or four short teeth
instead of one long hook, and these combs are better adapted for
getting a purchase on the walls of a wooden burrow than
would be the long hooks of some sand-burrowing Cicindele.
I intend to make a careful and critical study of the mouth-
parts; as it is certainly unusual to find a predaceous larva
with mouth-parts qualified to excavate burrows in wood. The
mouth of the burrow is countersunk, and Mr. A. H. Hamm
informs me that the same feature can be seen in the sand-
burrows of the British Cicindele. I have no doubt but that
the lower surface of the head of the Collyris larva fills com-
pletely the orifice of the burrow when the insect is awaiting
( Ixxiii )
its prey, the jaws projecting into the countersunk area. The
adult Collyris emarginatus is arboreal in its habits, is remark-
ably fleet, and readily takes to wing; in Borneo, as I have
shown (P. Z.8., 1902, vol. ii, p. 264), it is mimicked by a
flower-haunting fly of the genus Sepedon; it feeds on small
insects, and the statement in the Deutsche Entomol. Zeitschr.
1905, p. 172, that it is herbivorous is erroneous, and is due to
a misunderstanding.
“JT have also for exhibition some larve and pup of
Mormolyce, together with a specimen of a Polyporus fungus
split open to show the lenticular chamber excavated by the
larva. The adults, male and female, are usually found resting
on the under surface of these fungi, or on the tree trunk from
which the fungus projects ; oviposition has not been witnessed,
but I believe that only one egg (or at most two) is laid at a
time, for I have not been able to find more than one larva in
the chamber,* though on one occasion I took a newly-emerged
adult from the surface of a fungus in which was hollowed out
a chamber containing one larva. Access to the larval chamber
is attained by an orifice so small that it is surprising that the
newly-emerged beetle can squeeze through it to the exterior, this
orifice is situated on the upper surface of the fungus. It will
be seen that there are no features of particular interest about
the external appearance either of larva or pupa; these pup
do not show the elongation of the head and thorax character-
istic of the adult. Overdyk, who gives a bionomical account
of this species in Ned. Ent. Vereen, vol. i, 1857, states
that the larve are predaceous. The larva and the pupa have
been figured and described by Verhuell (Ann. des Sciences
Naturelles, 1847.)”
Professor E. B, Poutton communicated the following note
by Mr. A. H. Hamm, of the Hope Department, Oxford
University Museum. ‘The observations extend and support
those recorded by him in “Proc. Ent. Soc. Lond., 1904,”
p. Ixxv.
“The following observations made at Oxford during the
* The short note on this species which I published in British Assoc.
Reports, 1901, was largely based on information supplied to me by a native
collector, and this information, I regret to say, was proved afterwards
by my own observations to be inaccurate.
( Ixxiv )
present year, tend to confirm the opinion that Pieris rapx
chooses for prolonged rest, a surface upon which it will be
concealed. Although only five individuals were observed,
these had, without exception, selected a position of consider-
able protective value. The first butterfly was seen near
Cowley Marsh on May 7th, at about 4p.m. The afternoon
was dull and cloudy, and the insect was resting in a semi-
torpid state among the flowers of the white dead-nettle
(Lamium album) in a hedge. A week later, on May 13th,
between 4 and 5p.m., when searching the same hedgerow,
another individual was found in an almost identical position.
Hanging in the midst of the largish white flowers of the
Labiate the butterflies were beautifully concealed, and were
indeed only found because the plants were carefully examined
for other insects.
“On August 17th, between 7 and 8 p.m., when looking for
larve on the hedge bordering my garden, near the Cowley
Road, I found two rapx within a few inches of each other.
Both were resting on the silvery under-sides of bramble leaves.
In both cases, however, the leaves were twisted and the lower
surface had become the upper. Again, on August 20th, in a
lane leading to Temple Cowley, at 5.30 p.m., I watched an
individual flying slowly along a hedgerow, every nowand then
resting for a few seconds. Finally, after many attempts, it
settled down, apparently for the night, and on going to look
more closely, I found that it also had chosen the under-side
of a bramble leaf. Although not quite so well concealed
as among the white flowers, these last examples blended
wonderfully well with their environment.
“ After some years of close observation of the resting habits
of insects in general, I feel convinced that they possess an
inherent tendency to select an environment that aids in con-
cealment ; and as the various species differ in appearance so do
they select diversified situations suited in each case to the
particular requirement of the insect.”
A discussion followed, Mr. H. J. Etwes expressing his
conviction that the use of surroundings by insects, and Lepi-
doptera especially, for protective purposes simply, was still an
open question. Colonel J. W. YERBuRY mentioned the case
Conley *)})
of LHuchloé euphenoides and Zegris eupheme roosting at
Granada on plants of Biscutella, the appearance of the insects
at rest closely approximating to that of the flower heads.
Mr. H. Rowianp-Brown observed that he had noted a similar
resemblance between a specimen of Pieris napt and the flower
of Leucojuns on which it rested for the night. Mr. G. C.
CuHampion said that he had noticed specimens of Gonepteryx
vhamni seek the under-sides of bramble leaves.
Dr. T. A. Cuapman said that Col. Yerbury’s observation on
Euchloé euphenoides resting on the flowers of Biscutella is one
that may be often made in the South of France, and few
English Entomologists have not observed 2. cardamines at
~ rest on Alliaria flowers, or on some Umbellifer that roughly
resembles it. These facts seem to give the clue to how the
practice of these Crucifer-feeding Pierids of affecting objects
of their own colour for resting arose. In so many of these
cases, the food-plant is in flower when the butterfly is on the
wing, and in the case of Huchloé the flower head is the
place selected for egg-laying. It comes about, then, that the
butterflies frequent the flowers of their food-plant, both for the
honey they afford and for egg-laying. That it would often
happen, therefore, to be the resting-place of the butterfly
when overtaken by a sudden failure of sunshine is obvious.
Nothing further is needed as a basis for natural selection to
gradually accumulate in association, the characters of resting
on the flower of the food-plant, and assimilation to it in colour.
A butterfly, searching for a flower head to rest on, will act
of course in the same way as one searching for it for any other
purpose, viz., it will discover at a distance an object of the
desired colour, and will be able to verify what it really is,
only after a close approach. If it wants to suck honey, or to
lay an egg, such verification is imperative, and all collectors
are familiar with butterflies closely approaching a piece of
coloured paper, a dead leaf or other object, and only when
quite close discovering that it is not the desired mate, or
flower, and at once departing. But if a resting place for the
night is desired, such verification is not absolutely essential,
and a place must be chosen, so that if after several tries a
near approach only produces disappointment, an occasion at
(C tee.)
length occurs, when advancing evening makes it necessary to
appropriate the resting place without the final verification.
Thus a Pieris or a HLuchloé will learn to accept as a resting
place any white object, even if it be not a flower head of a
Crucifer.
But if it be objected that cabbage and turnip, the usual
foods of our “‘cabbage” butterflies, have yellow flowers, the
reply is that these foods, for these butterflies, are entirely
human inventions, and therefore comparatively modern.
Their natural foods have chiefly white flowers, such as,
Nasturtium, Arabis, Draba, Cochlearia, Thlaspi, Lepidiwm,
Turritis, Alliaria, and many others, the two last mentioned
being the favourite foods of ZH. cardamines. It may also be
suggested that the frequently very yellow under-sides of our
common ‘“‘cabbages” are due to the yellow flowers of our
cultivated Brassicas, and of such wild Crucifers as Raphanus
sinapis, and Barbarea.
The Presipent, Mr. A. J. Cuirry, Dr. F. A. Dixny, Pro-
fessor E. B. Poutton, and other Fellows offered observations
on the subject.
Papers.
Mr. Wiuuiam Joun Lucas exhibited diagrams of the instars,
and also of the mouth parts of the imago, to illustrate the
paper read by him ‘On the Emergence of Jyrmeleon
Sormicarius.”
Mr. Martin Jacosy communicated a paper entitled
“Descriptions of New Species of African Haltictne and
Galerucine.”
Mr. CraupE Mortry communicated a paper “On the
Ichneumonidous group Zryphonides schizodonti, Holmegr.,
with Descriptions of New Species.”
(| Ixxvii )
ANNUAL MEETING.
January 17th, 1906.
Mr. F. Merririetp, President, in the Chair.
Mr. R. Wvytitr Lioyp, one of the Auditors, read the
Treasurer’s Balance Sheet, showing a balance of £50 3s. 5d.
in the Society’s favour.
Mr. H. Rowrtanp-Brown, one of the Secretaries, read the
following
Report of the Council.
During the Session 1905-1906 eleven Fellows have died, viz.
two Honorary Fellows, M. Henri de Saussure and Dr. Alpheus
8. Packard, and nine Fellows, Mr. Alfred Beaumont, Mr.
George Bowdler Buckton, F.R.S8.,M. A. Prudhomme de Borre,
Mr. John William Douglas, Lieut.-Colonel L. H. Loyd Irby,
Captain F. Wollaston Hutton, F.R.S., Mr. Alexander Fry,
Mr. H. W. Peal, and the Rev. Francis Augustus Walker,
D.D.; seven Fellows have resigned, or have been struck off
the list, and twenty-three new Fellows have been elected.
The number of Fellows deceased is, happily, not above the
average; nor is the number of those who have resigned, or
for other causes have been removed from the list. The
number of Fellows elected, however, shows a considerable
falling off from last year, though with candidates nominated
and still to be elected it compares favourably with most
previous years. The Council, however, are of opinion that
the number might still be considerably augmented if Fellows
would bring somewhat more energetically to the knowledge of
their entomological friends the advantages of Fellowship of
the Society.
At present the Society consists of twelve Honorary Fellows,
and five hundred and four Life and Subscribing Fellows,
making a total of five hundred and sixteen. These figures
show that, for the first time in the history of the Society, the
number of ordinary Fellows has reached five hundred, while
the total fellowship also surpasses that of any previous year.
The Transactions for the year 1905 form a volume of four
PROC, ENT. SOC. LOND., v. 1905. F
(> iezvanl*)
hundred and thirty-eight pages, containing twenty-four
Memoirs, contributed by the following authors: Mr. Nelson
Annandale, Professor Christopher Aurivillius, Mr. George
Bethune-Baker, Mr. Horace A. Byatt (with a note by Pro-
fessor E. B. Poulton), Mr. J. Chester Bradley, Mr. George C.
Champion and Dr. Thomas A, Chapman (with description
by Professor O. M. Reuter), Dr. T. A. Chapman (three papers),
Mr. John Alexander Dell, Mr. W. L. Distant, Mr. Herbert
Druce and Mr. Hamilton Druce, Mr. F. DuCane Godman, Sir
George F. Hampson, Bart., Mr. John C. W. Kershaw (two
papers), Mr. G. W. Kirkaldy, Mr. Arthur M. Lea, Dr.
George B. Longstaff, Mr. E. G. B. Meade-Waldo, Mr. E.
Meyrick, Mr. Claude Morley, Mr. Edward Saunders, and Dr.
David Sharp and Mr. F. Gilbert Smith.
Of these twenty-four papers no less than fourteen relate to
Lepidoptera, three to Khynchota, three to Hymenoptera, two
to Coleoptera, and one to Diptera, while Mr. Champion and
Dr. 'T. A. Chapman’s paper, “ Another Entomological Excur-
sion to Spain,” deals with Coleoptera, Hemiptera-Heteroptera,
and Lepidoptera.
The Memoirs above referred to are illustrated by nineteen
plates, including two maps, of which eight are coloured. The
entire cost of Plates XVIII and XTX has been defrayed by
Mr. E. Meade-Waldo, and of Plate X by Mr. F. DuCane
Godman. Dr. T. A. Chapman has given the whole cost of
Plate VIII, and half the cost of Plates II, ITI, IV, V, VI,
and XII, and Professor E. B. Poulton half the cost of Plate
XIV. Blocks inserted in the text of the Transactions and
the Proceedings have also been given by Dr. G. B. Longstaff,
Dr. T. A. Chapman, and Mr. H. Rowland-Brown. The
volume of the Proceedings, consisting of about eighty pages
in all, has been well kept up, thanks to the increased number
of interesting exhibitions made by Fellows, and the notes read
by them at the meetings, which also show a satisfactory
average of attendances,
During the past year eighteen volumes—in addition to one
hundred and eighty-seven periodicals, pamphlets, and reprints
—have been added to the Society’s Library, the increased use
of which is demonstrated by the fact that, according to the
( Texte’)
Assistant Librarian’s Report, two hundred and eight volumes
have been taken out by Fellows during the year, and the
number of Fellows using the Library has largely increased.
There have also been placed in the Library, under the gift
of the Representatives of the late Mr. Robert McLachlan,
F.R.S., not included in the aforesaid donations and purchases,
two hundred and ninety-two volumes and pamphlets, to
which a distinguishing mark is affixed in the list of additions.
During this year, also, it has been arranged that the Library
shall be open daily from 9 a.m. to6 p.m., except on Saturdays,
when the hours are from 9 a.m. to 2 p.m,
The Treasurer reports that, after carrying forward to 1906
the sum of £8 8s. for subscriptions paid in advance in 1905,
and investing £47 5s. for three Life Compositions received
during the year, making the total sum so invested £857 10s.
7d., there remains a genuine cash balance in the Society’s
favour of £50 3s. 5d.
Arrangements have been made during the year for the sale
of back numbers of the Transactions to Fellows and non-
Fellows at a considerable reduction, and the report of the sales
thus effected, and for more recent volumes, shows a satisfac-
tory result. On the whole, therefore, we have to report that
our financial position is well established, and Fellows may
again be congratulated on the flourishing condition of the
Society.
EntTomo.LoaicaL Soctety or Lonpon,
11, CaHanpos Street, CavenpisH Square, W.
17th January, 1906.
The Secretaries not having received any notice proposing
to substitute other names for those contained in the list
prepared by the Council, the following Fellows constitute the
Council for 1906-1907 :—Gilbert J. Arrow; George C.
Champion, F.Z.8.; Arthur John Chitty, M.A.; James E.
Collin; Dr. Frederick Augustus Dixey, M.A., M.D. ; Herbert
Goss, F.L.S.; Arthur Hugh Jones; William James Kaye ;
William John Lucas, B.A.; Frederic Merrifield ; Professor
Edward B. Poulton, D.Sc., M.A., F.R.S.; Louis Beethoven
( deez’)
Prout; Henry Rowland-Brown, M.A.; Edward Saunders,
F.R.S., F.L.S.; Richard 8. Standen, F.L.S.; Commander
James J. Walker, M.A., R.N., F.L.8.; Charles O. Water-
house.
The following are also re-elected as officers :—President
Frederic Merrifield; Treasurer, Albert Hugh Jones ; Secre-
taries, Henry Rowland-Brown, M.A., and James J. Walker,
M.A., R.N., F.L.S.; Librarian, George C. Champion, F.Z.8.
The Balance Sheet and Report having been adopted, Mr.
F. Merrifield, the President, delivered an Address. A vote of
thanks to the President for his Address, and for his services
as President during the past year, together with those of the
other officers, proposed by Professor Meldola, and seconded
by Mr. Charles O. Waterhouse, was carried unanimously. Mr.
H. Rowland-Brown, Commander J. J. Walker, and Mr. A.
H. Jones replied.
( Texxt’)
ENTOMOLOGICAL SOCIETY OF LONDON.
Balance Sheet for the Year 1905.
RECEIPTS. PAYMENTS.
By Seads Ee asond:
Balance in hand, Ist Jan., Printing Transactions, etc. 242 138 2
1905, and at Bankers’... 18 15 5 | Plates,etc. ... ... ... 105 5 4
Subscriptions for 1905 ... 392 14 0 | Rent and Office Ex-
AY PORES?) 62 5-)u cist) on bye, Oly LORE O WENSES, (2c. «22, cae) ae COAL: 6
Admission Fees ... ... 29 8 O | Books and Binding... ... 44 2 8
Donations... ... ... ... 3 5 9 | Investment in Consols ... 47 5 O
Sales of Transactions ... 144 14 4 | Subscriptions in Advance,
Interest on Investments :— per contra carried to
Consols ... ... £1917 1 1906% Ui. 9s, aipen 8) '8% 0
Westwood Bequest 6 16 6 Balance in hand and at
——26 13 7 Bankers?! 7 a5) ses) 2-5. 00) ar 8
Subscriptions in Advance 8 8 0
Life Compositions ... ... 47 5 O
£702 14 1 £702 14 1
In account with the late Ropert McLacutan, Esq., Treasurer.
Balance at Bankers’ due Paid by cheque to his
to his Representatives Representatives ... ...£100 0 0
brought forward from
last year) c/s 1 SLOO) O70
Testimonial to the late Mr. W. R, Haut, Resident Librarian.
Further subscriptions Paid by cheque to his
Tecelved esse sec | c-ye ee oe Representatives ... ... £1 1 0
ASSETS. ADDITIONAL ASSETS.
Be send.
Subscriptions in arrear Contents of Library, including
considered good (say)... 50 0 0 | books from the library of the
Cost of £857 10s. 7d. late Mr. McLachlan and unsold
Consols... ... «. +» 889 8 O | Pyblications.
Cost of £239 12s. 4d.
Birmingham 3 per cent. 250 0 0
T. A. CHAPMAN.
Balance inhand ... ... 50 3 5
R. W. Lioyp.
No ASCERTAINED LIABILITIES. Roasan ApEn,
A. Huau JongEs, RicHarp Soutu.
Treasurer. H. RowLanD-BRown.
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( * bein: )
THE PRESIDENT’S ADDRESS.
LapIES AND GENTLEMEN,
The Report presented by the Council will have
informed you of the prosperous state of the Society, and
of the record it has during the past year established in the
number of its Fellows. When one considers the vast number
of species of insects, as compared with that of other classes
of animals, their immense antiquity, the perfection they have
reached in their physical equipment and in their instincts,
their exhaustless variety of form and function, and the scope
which, by their metamorphoses and in other ways, they
afford for the solution of problems in which all biologists
are interested, one can but expect that the Society will
become more and more important, and that its members will
continue to increase in number and, as far as indications
appear, certainly with no deterioration in quality. The
interest which draws its members together is one that is
concerned with every part of the world. It is by a collation
of the species and varieties which all the different parts of
the world have to present that the science will be advanced,
and from this point of view it is most satisfactory to note
how largely the Society is constituted of Fellows resident
abroad. In looking through the list of Fellows at the
beginning of the year 1905, 1found that these mounted up
to more than one-fifth of the whole membership. Fifty-nine
are resident in the British colonies and dependencies, and
forty-six in foreign countries, of whom twelve are honorary
members.
It is of interest to observe that about half the papers
(| Ixxxiv )
contributed to the Transactions are of a biological character,
and that many of the papers contain, besides the full
descriptions of new species that are not only important but
necessary, notes on their life histories and habits.
One of the most satisfactory features of the volumes published
of late years is the amount of space given to the Proceedings.
These record the periodical meetings of the Society, when,
by means of the objects exhibited and of the observations
that arise out of them and out of the papers read, many
valuable facts are made known ; they frequently give rise to
discussions of the greatest interest, and it is important that
the advantage of these should not be lost to those who are
not present at the meetings when they take place. It is of
course impossible that your Honorary Secretaries, to whose
skilled and assiduous labours we are so greatly indebted, should,
unassisted, record with sufficient fulness all the salient points,
but if those who bring forward objects for exhibition or take a
part in the discussions will always render assistance by handing
in notes of their contributions and of their observations, a
satisfactory record can be preserved.
I hope it will not be out of place to express the satisfaction
with which the Fellows of the Society have observed that
some of their body, including two members of the Council,
Professor Poulton and Dr, Dixey, were among those who
took part in the expedition to South Africa of the British
Association for the Advancement of Science. Wecongratulate
them on their successful journey and their safe return in
vigorous health, and I suppose we may say with even enhanced
knowledge, from which our discussions have already derived
benefit. I think I may add that our gratitude to them in-
cludes a prospective element, for we know that they have
brought back much that is of great value in the way not only of
specimens but of recorded observations, from which the future
Transactions and Proceedings of the Entomological Society,
of which they are such important members, cannot fail to
derive much advantage.
I follow the time-honoured practice of referring to some
of the Fellows of the Society whose loss during the past year
we have to deplore.
(i isxxy 5)
Dr. ALpHEUS Sprinc Packarp, an Honorary Fellow of this
Society, died at Providence, Rhode Island, on the 14th
February, 1905. He had held the position of Professor of
Zoology and Geology in Brown University since 1878. No
one did mare than he to advance the knowledge of entomology
in the United States. Among his voluminous contributions
to science, the list of which fills nearly ten pages of the
catalogues of the library of the Entomological Society, may be
singled out for mention his monograph of the “ Bombycine
Moths,” and the Text-book of Entomology published in 1898,
in which he sums up the existing knowledge on the anatomy,
physiology, embryology, and metamorphoses of insects, with
full bibliographical lists; this work may be described as
supplying a well-ordered guide to all that was known on the
wide range of subjects of which it treats. Mr. Packard was
well known for his advocacy of “ neolamarckian ” views.
The loss of Honorary Fellows during the past year includes
another distinguished name, that of M. Henri F. pe SaussuRg,
of Geneva, eminent in many departments of life, grandson of
one whose name was once famous throughout Europe as
having made the ascent of Mont Blanc in 1787. Born in
1829, he wrote at an early period of life his monograph of
the “Solitary Wasps,” which he completed in Paris, where he
became known to some of the distinguished men of science.
In 1854 he was granted the diploma of Doctor by the
University of Giessen. In the same year he started on a
course of travel, which he pursued through the West Indies
and parts of the North-American continent, and after his
return to Europe in 1856 he began the publication of a vast
number of works on a great variety of subjects—geological,
zoological, ethnological, geographical and historical. Especially
large were his contributions to Entomology, and among these
the Hymenoptera and the Orthoptera in particular claimed his
attention. He died on the 20th February, 1905, lamented for
his deep and wide knowledge and for his attractive personal
qualities.
A veteran English entomologist, Jonn Witi1AM Douec.as,
elected in 1845 a Fellow of this Society, of which he became
President in 1861, died on the 28th August, 1905, in his
(° ixxevi ))
ninety-first year. As a young man he obtained a situation in
the Customs House, where he rose to a high position, retiring
at the age of seventy after more than fifty years’ service. His
first entomological paper was published in the ‘‘ Entomological
Magazine” for 1837. In 1874 he became one of the joint
editors of the Entomologist’s Monthly Magazine, retaining
that position for thirty years. Associated with the leading
English entomologists of his day, including Stainton, whose
residence was near his own, he extended his studies from the
Lepidoptera, with which he began, to the Coleoptera and other
orders, rendering important assistance in the production of
Stainton’s “ Natural History of the Tineina” in 1856, and
in 1865 he was one of the joint authors of Douglas and Scott’s
“ British Hemiptera.” His works on the Coccide are very
valuable. During his long life he did.much to promote the
knowledge of our favourite science, and always held a high
position in the esteem and affection of his friends.
LiEUTENANT-CoLONEL LeonaRpD Howarp Loyp Irpy died last
April at the age of sixty-nine. A Fellow of the Linnean and
Zoological Societies, he joined this Society in 1893. Retir-
ing after service through the Crimean War and the Indian
Mutiny, he devoted himself to several branches of natural
history, especially ornithology, on which he published a work,
“The Ornithology of the Straits of Gibraltar,’ that went
through several editions. He was an enthusiastic collector
of Lepidoptera, a first-rate shot, and his friends found him a
most charming and genial companion.
ALEXANDER Fry, born in 1821, went in 1838 to Rio de
Janeiro, where his father had a mercantile house. There he
devoted much time to collecting Coleoptera, After 1854,
barring occasional visits to Rio, he resided in England, adding
by purchase and otherwise to his collection, which at his death
reached the enormous number of some 200,000 specimens, all
of which, well arranged and for the most part named, he
bequeathed to the British Museum, where they occupy 29
cabinets, containing 348 drawers, and 400 boxes. He became
a Fellow of this Society in 1855, and died 26th February, 1905.
Grorce Bowpier Buckton, F.R.8., had been a member of
the Entomological Society from 1883. Incapacitated by an
a
( Ixxxvil )
accident in early life from active pursuits he turned his
attention to science, and in 1867 carried out some important
original work, in recognition’ of which he was that year made
a Fellow of the Royal Society. In 1865 he settled at
Haslemere, where he died on the 25th September, 1905. Mr.
Buckton was the author of valuable entomological works. His
monograph of the “ British Aphides,” in four volumes (1876-
1883), was followed in 1890 by his illustrated monograph of
the “British Cicade or Tettigide,” and afterwards by the
“Natural History of Hristalis tenaa,’’ and by various pub-
lished papers. Most of the plates in these works were drawn
and in some cases lithographed by himself. Mr. Buckton was
a member of the Linnean Society, the Chemical Society,
and of several foreign scientific societies, many of the meetings
of which he attended, and by the members of which, as well
as his numerous private friends, his kindly and courteous
presence will be missed.
I ask leave this evening to bring under your notice a
subject on which entomologists have thrown some light, but
which by common consent stands in need of much more ; to
call attention to some of the effects on living things of that
chief element in climate, temperature, and to the remarkable
limitations imposed on those effects by the nature of the living
thing operated on, these two opposing forces giving rise to a
state of conflict between the outside power and the organism.
General operation of temperature.
The general effect on a growing organism of an augmented
temperature, within the limits of healthy vitality, and
associated with other wholesome conditions, is well recognized,
as promoting physiological activity, heightening energy and
accelerating development. The whole practice of “forcing,”
as applied in many industries, by florists, market-gardeners,
chicken-farmers, and others, is founded on this principle, and
entomologists every day come upon illustrations of it.
( “lxxxviii )
Insensitiveness of many oxganisms.
And yet we know there are myriads of insect organisms
in an immature condition, apparently waiting for nothing but
development, which are absolutely insensitive to the influence
of a heightened temperature, nay, in some cases, as it were,
resent that influence, and are moved by it in the opposite
direction, that of delay or postponement. This radical dis-
tinction in physiological character is one that is met with
throughout the Lepidoptera, and, of course, elsewhere in the
animal kingdom. Individuals belonging to the same family,
the same genus, the same species, even the offspring of the
same parents, are found separated into these two distinct
physiological classes.
Regular response of oné class.
The first class—those that yield to temperature—seem
usually to respond to it with a regularity which is noteworthy,
in many instances all emerging to the very day. I give
examples, summer pupe of some common English species,
showing this.* (See Appendix, Table I.)
Resistunce of another class.
As examples of the other class, that on which warmth is
expended in vain, we have such species as Huchloe cardamines,
Araschnia levana (winter phase), Pieris brassice (winter phase),
Selenia tetralunaria (winter phase), some experiments on most
of which are described later,
I am not suggesting that the second class, those non-respon-
sive to temperature, are thus insensitive throughout their lives ;
the insensitiveness may be confined to certain metamorphic
stages, or even to certain periods comprised in those stages ;
when, for example, any of them shake off their long torpor
* In my experience eggs of most Lepidoptera that are laid at one time
in summer hatch out nearly all together, in this respect differing greatly
from many of the wintered eggs, such as those of the genus Ennomos,
and of Liparis monacha, which usually spread their hatching over a very
long period with little regard to temperature. In this respect there is a
parallel between the egg and the pupa.
al
ao
( ( leexix) )
and begin to move actively towards emergence, the pace at
which they do so is accelerated by warmth, and retarded by
cold.
Nor do I suggest that all individual Lepidoptera belong,
throughout their lives, or even throughout any one meta-
morphic stage, to one or other of these two classes, nor that
the element of personal variability does not come in largely,
so that there are many species and individuals which may be
described as intermediate, though often very discontinuously so.
Physiological dimorphism.
I shall have to advert, rather fully, to that very numerous
and remarkable class called double-brooded (digoneutic and
bivoltine), which in one of their ‘“ phases”—to adopt the
term suggested by Professor Poulton—belong to the sensi-
tive, in the other phase to the insensitive, thus exemplify-
ing the kind of diversity to which Dr. Dixey has given the
apt name of “ physiological dimorphism.”
Nonseasonal Lepidoptera.
The climates of the world may be divided into the non-
seasonal and the seasonal, the former consisting of those equa-
torial or tropical regions where, with a temperature averaging
something over 80° F. (27° C.),* not much above and very little
below, and with adequate humidity all the year round, these
conditions continuing the same year after year, species abound
whose “ little lives are rounded” in a month, regions where
at all times and in all their stages—egg, larva, pupa, and
imago—they are to be found simultaneously and in nearly
equal abundance—such species as the Heliconias and Ithomias
of tropical South America, and the Ypthimas, etc., of the Indo-
Australian region. I do not know the life histories of these
species, but, looking to the regularity with which generation
succeeds generation, it may perhaps be assumed that so far as
the operation of the temperature—itself so regular—is con-
cerned, there is nothing in a nature bred under these con-
ditions to prevent that general effect which I have adverted to
* For convenience I have in all cases given, approximately, the
Centigrade degrees,
( xe )
—i.e. that warmth heightens and accelerates, and coolness
retards, in other words, that these species would respond in a
normal manner to change of temperature within the limits
of health.
Seasonal influence— Temperature.
The rest of the world is all more or less “ seasonal” and the
main distinction between the seasons is that of temperature.
In many countries containing an abundance of Lepidoptera the
principal climatic distinction is that between dry and wet
seasons, these, however, being often more or less associated
with changes of temperature. The recorded accurate observa-
tions on the effects of temperature in relation to Lepidoptera
are far more considerable than those on the effects of wet and
dry seasons, and, without in any way seeking to minimize the
operation of these, I practically limit myself to the former,
not only because I think it the most far-reaching and
penetrating influence, but also because the materials accessible
for the purpose are so much more abundant.
Seasonal Lepidoptera.
There are, as we know, places extreme in climate, such as
North-Eastern Siberia, where the annual range of temperature
is over 100° F. (56° C.), from about — 50° F. (— 46°C.) in
January to over + 50° F. (10° C.) in July, and the polar regions,
where some of the Argynnids and Satyrids are found, and
where the actual life of a butterfly is limited to about a month,
though the whole duration of its life may extend over several
years, and the food plants are covered with ice and snow
except for about six weeks. ‘Though the range of temperature
in the seasonal world in general does not approach such
extremes, the seasons are strongly marked in most of it, and
whenever the winter is sufficiently severe, either by its direct
effect or its indirect effect in destroying green food, to impose
a lengthened period of rest, the Lepidoptera are, like the
climate, seasoned in their habits ; and, the seasons having an
annual cycle, the normal life of a lepidopterous insect is
annual also, or at least is comprised within a year, its rate of
(*ixer 3)
development, in conformity with the general law adverted to,
being, as a rule, rapid in the warm season, slow or even
arrested during the colder part of the year.
Year-lived Lepidoptera.
The more usual case, at least in our latitudes, may be taken
to be that of insects which live a whole year, constituting those
seasonal species whichare usually calledsingle-brooded, but which
it might be more convenient, as their whole life is in question
and the term ‘‘ brood” is often used in a different sense, to
call “year-lived.” These, in contrast with the non-seasonal
species, are in general, in their four several stages—those of
egg, larva, pupa, and imago—associated respectively with
particular seasons of the year, and any of their stages, varying
however according to species, may be associated with any
season.
It will be convenient here, as the foundation for some of
the observations that follow, to refer to a few familiar
examples of the extreme persistence in their habits, in disregard,
apparently, of very great differences of temperature, of some
of the “‘ year-lived ” class, so numerous in temperate countries,
and to select for that purpose some that pass the winter in
varying stages. Huchloé cardamines ranges over Europe and
a large part of Asia, and is found plentifully at low elevations
as well as up to 7,000 feet above the sea level. Appearing
on the Riviera in February or March, and in the mountain
districts found occasionally as late as August, it pupates usually
in the early part of summer and then (with rare exceptions, if
any) quietly bides its time through all the hottest part of a
South European summer until the spring of the following
year. This is an insect which has a winter pupa. The
same rigid adherence to the one-year cycle is shown generally
by the Erebias and other Satyrids, chiefly mountain insects,
but many of them found in the warmer plains; these pass
the winter in the larval stage, as does Aporia crategi, a
very abundant species with wide geographical range. <A
similar fixed yearly life is shown by species which, like
Argynnis adippe, Augiades comma, Lycena xgon (argus), the
( ) mei: \)
Theclas, and the Geometrid moth Hnnomos angularia, pass
the winter as eggs, and by Scoliopterya libatrix, and many
other moths which hibernate in the winged stage. So the cold-
weather moths of autumn, winter, and early spring, the four
common Hybernias and Cheimatobia brumata and Phigalia
pedaria, rapidly run through the stage of larva in about two
months, and lie all through the greater part of the year,
including its hottest period, before they emerge as moths.
Cold required to mature winter pupe.
A continuation by artificial means of heat not only fails to
accelerate but often retards or kills, and in some cases throws:
the emergence over to another year, so that in these cases
cold is necessary to accelerate or cause maturity. Four Thais
polyxena, a year-lived spring butterfly of Southern Europe, were
placed by me at 80° F. (27° C.) on each of the following dates,
12th November and 31st December. All were dead by 14th
March except one of the first lot, which emerged a cripple
28th December. Of four placed there on 14th March and
two at 92° F. (33° C.) on 25th March, all emerged ; of some
that were left out of doors all, or nearly all, emerged 16th to
30th May. Ten Rumia luteolata placed by me on the 18th
October at 80° F. (27° C.) were all dead by March ; ten placed
there on the 20th March all died except three, which emerged
from the 4th April to the 2nd May, one being a cripple; of
twenty placed there on the 2nd May, twelve emerged between
the 6th and 22nd May; many left out of doors emerged be-
tween the 17th and 3lst May, and a few later, Of twenty-
five Hybernia marginaria placed on the 19th October at the
more moderate temperature of 63°-70° F. (18°-21° C.),
not one had emerged by the Ist March, by which time many
at the temperature of a cold room had emerged. Of seventy-
five C’. brumata brought indoors when the weather began to be
cold in October, and on the 2nd November placed at 63°-
70° F. (17°-21° C.), not one had emerged by the 12th
January, when many at the ordinary temperature had
emerged ; some of them had been killed by the warmth indoors,
( xcii )
but most apparently had been sent over by it to wait for a
second winter.*
There are many other species which, though not so obstinate
as those which have been mentioned nor perhaps so ready to
die or to postpone emergence for a year, show an equally deter-
mined resistance for a shorter period. These species probably
belong properly to a somewhat warm climate and are attuned
to a shorter winter.
Mediterranean species.
Wholly opposed in their constitution to the seasonal year-
lived species are many belonging to parts of Southern Europe
and of Northern Africa, bordering the Mediterranean, where
the summer is warm and long, the spring and autumn mild,
and the winter not severe enough to prescribe any lengthened
rest, and where accordingly, though no rapid multiplication
of generations approaching to that of the equatorial regions
exists, the year, with its marked seasons, is no longer with
many species the usual period of life, but they breed on,
generation after generation, continuously, getting in thvee or
four or more generations annually.j This is exemplified with
such species as Pyrameis cardui, Colias edusa, Phragmatobia
fuliginosa, and Plusia gamma. These would appear physio-
logically to belong rather to the equatorial non-seasonal than
to the seasonal class, and at all events seem to have little of
the seasonal character stereotyped upon them.
The seasonally double-brooded.
It is now time to refer to that remarkable class usually spoken
of as the seasonally double-brooded, producing two or sometimes
more generations in the year; one, the “ winter phase,” that
which in our climate lives about nine months, including the
winter, the other, the ‘“‘summer phase,” living about three
months, May to August; the one having a pupal period of
* Similar occurrences are frequently recorded in entomological publica-
tions. (See Dr, Chapman, Ent. Record, vol. i, p. 127, and the Ent,
Record, passim,)
+ See Tutt in Ent. Record, vol. x, pp. 211, 212.
PROC. ENT. SOC. LOND., v. 1905. G
{ ae )
about seven to eight months, the other of afew weeks, and the
two phases, as will be seen, differing to such a degree physio-
logically, that the winter phase is killed by the temperature in
which the summer phase flourishes; often differing also in
size, shape and colour, and I believe frequently in habits.
The winter phase appears to have much more in common
physiologically with the ‘ year-lived” species, the summer
phase belonging on the other hand to the equatorial or the
Mediterranean type.
Examples of winter pupx killed by warmth.*
Among these let me first mention the famous Araschnia
levana, much the most striking of seasonally dimorphic
Kuropean forms of butterfly ; the winter phase being on its
upper surface of a bright orange brown, mottled with black,
having a general resemblance to some of the little ‘‘fritillaries”’
(Melitxas and the smaller Argynnids) so abundant in spring,
the summer phase black, with an oblique white band running
down both pairs of wings, much as it does with the common
Limenitis sibylla. The pup of the summer phase respond to
temperature most readily, those forced emerging in about
six days with great regularity; if on the other hand
cooled, they are retarded for a period of, it may be,
many months. To the effect of such cooling on the
facies of the imago allusion will be made later. In 1895,
between the 22nd and 29th August, I received from
various parts of Germany, North, Central, and South, many
hundreds of larvee of A. levana of various sizes from half-
grown or less upwards; many were very kindly sent to me
from Freiburg by Prof. Weismann. These pupated from about
the 28th August to the 2nd September. A large number
were on pupation placed in the refrigerator at about 46°—50°
F, (8°-10° C.) for eight weeks, @.e. till 26th October, when
they were placed out of doors, batches being transferred at
about fortnightly intervals to incubators (at 92° F.(33° C.) and
80° F, (27° C.)—it made no difference which). Some were left
out of doors from pupation, and 53 of these emerged between
* See Table II in Appendix, Experiment 1.
( %ev,)
2nd April and 7th May. It will be seen by the tables in the
Appendix that all those transferred to the incubators in less
than 13 (perhaps 15) weeks after pupation died ; after that a
proportion died, gradually diminishing until the 29th week,
after which all survived.
A large number during the first eight weeks were left in
the room at a temperature which may be taken at 5°-10° F.
(3°-6° C.) above that of outdoors. These were afterwards
transferred in batches to the incubators. All so transferred
up to the 19th week died ; after which a proportion, gradually,
but irregularly, increasing, but never the whole of those
transferred, survived.* (Of some left in the room until
emergence, 23 emerged between 24th March and 31st May.)
Of a third lot left in the room until 13th January (19 weeks),
when they were placed out of doors and from 7th February
(24 weeks) transferred as before to incubators, some of each
batch transferred survived, the proportion being larger than
in the last experiment.t (Some of them, left out of doors
from the time they were placed there, emerged, to the number
of about 20, between 26th April and 19th May.)
Experiments with other seasonal double-brooded species
show an equal resistance to a heightened temperature on the
part of the winter phase. Thus, of 40 English pupz of P.
brassicx forced at 80° F. (27° C.) from the 29th November,
1897, all but 7 were dead by 23rd March, 5 more by 18th April,
the remaining 2 on 22nd May, one of these last in attempting
to emerge. Eighteen of the same batch were iced 29th Novem-
ber, and forced on the 23rd March; 11 of these emerged
between the 3rd and 6th April. Another lot of 86, received on
the 30th December, 1900, were kept out of doors ; of 10 forced
lst March only one emerged, on 21st March; of 6 forced
19th April only 2 emerged, on 30th April ; of 6 kept outdoors
all emerged from 30th May to 2nd June.
Many experiments tried by me on Selenia tetralunaria, some
of which are recorded in the Transactions and Proceedings
of this Society, show somewhat similar results. As to the
summer phase of this species, when the pupa is forced at about
* See Table II in Appendix, Experiment 2.
+ Lbid., Experiment 38.
(- sevi )
80° F. (27° C.) it emerges with great regularity in about 8
or 10 days; but the winter phase behaves very differently. Of
16 forced at 80° F, (27° C.) on the 26th November, 1890, and
11 more forced 14th December, all were dead by Ist February ;
of 16 kept at the more moderate temperature of 60° F. (16° C.)
on 26th November all were dead by 16th March ; many were
iced 26th November to 24th December ; of 8 of these forced
24th December and 7th January at 80° F. (27° C.) 6 died and
2 emerged, one a cripple; while of those not forced until the
21st January or later, nearly all emerged and with fair
regularity.
Tolerance of cold by summer pupe.
In contrast with winter pupe killed by the absence of cold,
reference may be made to the much greater tolerance of an
unaccustomed temperature shown by the summer pupa of S.
tetralunaria and of A. levana, which I have kept, retarded and
changed in facies but uninjured by a low temperature, for
more than four months, that of Z. punctaria for more than
six months.
Three or more broods.
The resemblance of the summer phase to the continuously
brooded is perhaps also indicated by its readiness to repeat
itself by interpolating one or more broods between its
first and latest broods, which is so commonly the case
in a hot season, all but the last of them showing the
characteristic quality of the larve of the summer phase,
i.e. the feeding up fast. The entomological periodicals con-
tain frequent instances of this. A. Jevana in a hot summer
often produces a third brood, and S. bilunaria (illunaria) from
eggs of the spring brood this year, has given me four
subsequent complete generations all of the summer phase,
having been kept by me at 70°-80° (21°-27° C.) during
their whole lives, the time occupied by them from the laying
of the first egg to the emergence of the last imago having
been only about six months, ze. from 3lst March to
2nd October. Thus, moths of five generations succeeded each
7
( xevii )
other within about six months. They then came to an end
without giving rise to a winter phase.
« Splitting.”
_ I have treated the tendency to be single-brooded, or the
reverse, as usually congenital in the individual ; and I think
this is proved by the common phenomenon which I have called
“splitting,” met with in nature and in home-breeding, when
part of a brood from the same parent, all bred under similar
conditions, divides, one portion feeding up hastily, and within
the year, the other portion feeding slowly and the pupa
resting peacefully till the following year. This separation, I
think, most usually shows itself in the larval stage, when the
“forwards,” to use Dr. Chapman’s name for them in his
experiments with Arctia caja,* feed up rapidly and voraciously,
the others biding their time ; his explanation being that A. caja,
as we find it, is a mongrel having two or more races mingled
together in its composition.
Though the character of the winter phase is in many cases
impressed so deeply on the individual as to render it immov-
able in its habits, there is a vast number of cases in which
it is not so. The books are full of accounts of Lepidoptera,
ordinarily single-brooded in England, which give out a second
or even a third brood in a hot summer, and the degree of
instability in this respect varies greatly not only as between
different species, but as between different individuals.
Conversion of one phase into the other.
Reverting to those individuals on which the physiological
character, 7.e. that of the summer or winter, is strongly
impressed, it seems proper to say something on the difficult
problems connected with the conversion, complete or partial, of
one of these phases into the other,
Conversion in the pupal stage.
To deal in the first instance with conversion in the pupal
stage, let me point out that it appears to be usually easier to
convert the summer phase into the winter than the opposite,
* Ent. Record, vol. iv, pp. 265 et seg., and vol. v, p. 33 et seq.
(| xeviii )
often found impossible, and that the conversion can be made
very thoroughly so far as many characteristics go, for example
as to colour and markings (or pattern). For results obtained
in this way, I may refer to papers contributed by me to the
Society’s Transactions, which Dr. Dixey has done me the honour
to epitomize most accurately in an article in “Nature” of
23rd December, 1897.*
Some observation has been made j on the distinction I have
drawn between “markings” (pattern) and “colouring,” but I
do not think any one who has seen the results of my experi-
ments can doubt that the distinction is a real and very
substantial one. To speak first of the markings (pattern), a
striking change in these is caused by the exposure for many
weeks of the summer pupa of S. tetralunaria to a low tempera-
ture ; further, this change is distinctly in the direction of (L
do not say it exactly reproduces) the pattern which the spring
emergence bears. It begins to show itself after about six
weeks of the low temperature, and becomes the prevailing
pattern after about eight weeks, though even after twenty
weeks there is an occasional reversion to the usual summer
pattern,
Standfuss and Fischers experiments.
Probably many of my hearers are conversant with the
wonderful results (far surpassing mine) of high and low
temperatures in altering the colouring and patterns of Lepi-
doptera, especially the Vanessids, obtained by Prof. Standfuss {
* It may illustrate the plastic character of the summer compared with
the winter phase of S. tetralunaria, to mention that with the winter pupa
one can only alter the colouring, but with the summer pupa one can alter
either colouring or markings, or both. Where extreme temperatures are
used, some of the effects are doubtless to be ascribed to physiologically
injurious conditions ; and in the converse case, where the artificial condi-
tions are specially favourable to vigorous development, such as warmth com-
bined with moisture, the resulting imagines may be expected to attain large
size and great brilliancy, as they would in nature. (See Mr. Elwes, Proe.
Ent. Soe. for 1891, p. viii.) Length of time in feeding up (and in pupa)
seems to be connected with the large size and strong marking of the
winter phase of seasonal double-brooded Geometrids.
+ Weismann’s ‘‘ Neue Versuche, ete.,” translation by W. E. Nicholson,
p. 59, reprinted from ‘‘The Entomologist,” January—August, 1896,
t A translation by E. M. Dadd, F.E.S., of Prof. Standfuss’s ‘* Synopsis
of Experiments to the end of 1898,” appeared in ‘*The Entomologist,”
1900-1901, and has been reprinted. I have seen no translation of Dr.
Fischer’s papers, but hope to have an opportunity of referring more fully
to the valuable researches of these gentlemen at some future time.
(( (eid)
and Dr, K. Fischer, who used for the most part temperatures
more extreme than those which I employed. This is not the
occasion for setting forth the different theories ably stated by
those gentlemen in explanation of the results obtained, both
invoking to some extent “arrested development” as their
cause, and, as development is arrested by abnormal heat as
well as by abnormal cold, the theories are strongly supported
by the fact that there is a striking similarity between
many of the results of the application of directly opposite
temperatures.
The difference in size between the two phases is often so
great that in the pupal stage there can be no complete
conversion in this respect, and I think it may be doubted
whether in this stage even the summer phase is ever converted
physiologically, 7.e. so as to give to a summer pupa the ten-
dency to lie over for many months, as distinguished from the
mere temporary retardation imposed as the normal effect of
a low temperature.*
Conversion in earlier than pupal stages.
There is evidence, however, that in stages earlier than the
pupal the conversion of individuals may be complete. Dr.
Chapman in the paper I have adverted to mentions that he finds
about 5 per cent. pretty regularly of the larva of a brood of
A. caja to be “ forwards,” ¢. e. to be ready (under the very slight
encouragement it would appear of an indoor life, with its
somewhat higher temperature, and perhaps without it) to feed
up rapidly in about three months, and emerge the same
summer instead of hibernating as larvee ; and he records that,
of a brood which he sent to me for experiment and which
were forced by me, 75 per cent. so fed up in about two
months.
But it must not be assumed from this that conversion from
the winter to the summer phase can generally succeed with
half-grown larve. In 1895, from the 22nd to the 29th
* As regards size, it is noteworthy that the summer insect is often the
smaller, usually so, I think, in the Geometrid moths ; in the Sedenias the
size is much less. In A. devana, on the other hand, the butterfly from
the summer pupa is the larger.
(te)
August I forced at 70°-80° F. (21°-27° C.) a large number,
nearly 100, of larvee of A. levana, many not more than half
grown, the pup being from pupation kept for months at about
70° F. (21° C.) and removed in batches at about fortnightly
intervals to 80°-90° F. (27°-32° C.) or a little higher ; not one
was converted by the process into the prorsa form, though one
was of the porima form; all the rest died.*
I am satisfied that to cause conversion in many cases the
temperature must be applied in a very early period in the
life of the individual. I have been trying experiments with
a view of ascertaining the exact period—varying doubtless
in many species—in which to be effective the change of
temperature ought to be administered, without having yet
attained sufficiently definite results to lay before the Society.
Tam not sure that in some cases it would not be necessary
to make the application of temperature as far back as
the period when the egg is laid, and perhaps at a time when
it is still unlaid, so as to affect the ““germ plasm” of the
parents or one of them,
Races of Bombyx mori.
The case of the common silkworm moth, Bombyx mori, a
species which hibernates in the egg, is interesting, as showing
how deeply ingrained the tendency to single-broodedness may
be as a hereditary character implanted in different races of a
single species. The race ordinarily cultivated in the south
temperate zone of the silk-producing countries of France,
Italy and Central Asia, and in Japan, has many varieties,
but (with the exception of a few bivoltine, trivoltine and even
quadrivoltine races locally cultivated in parts of Italy) their
cycle is annual. And yet there is a race usually cultivated in
Bengal which is polyvoltine, producing ten or eleven genera-
tions ina year. All the races interbreed freely. So essential
is a certain degree of cold to the univoltine race, that in order
that the eggs should hatch out healthily when transported to
the Punjab and Dehra Dun, which has been done to improve
* Of 24 not thus forced as pupe but kept out of doors 16 emerged in
the spring.
GMet#)
the Indian breeds, the eggs have to be sent up annually to the
Himalayas for the necessary cold.*
Single and double-brooded —Difficulties of transformation.
It is sometimes thought that a fixedly single-brooded species
has only to move south sufficiently far to easily establish itself
there as a double-brooded one. But is this so? If you try
to work out, as I have done, the case of such a single-brooded
insect going south, I think you will find the difficulties arising
from want of fit between the seasons and the habit of the species
(to say nothing of the competition it would have to meet) such
as to render its task in establishing itself a lengthened and by no
means an easy one. The opposite case of a double-brooded southern
species going north would, I think, be a little less difficult,
because a species having the inherent capacity of being double-
brooded can better adapt itself to circumstances, though even
here I think most of its attempts would be failures.t I believe,
however, that an altogether non-seasonal species where all the
pup are of the same physiological type, regularly responsive
to temperature, would find the change more easy.{ Supposing
an equatorial species having ten or twelve generations in a
year to flow over northwards, as it appears frequently to do, §
it would only have to reduce the number of these gradually
in natural response to the lower temperature, as it spread
over cooler regions. When they came down to three or four,
one of which would have to encounter a very mild winter, to
prolong this one in accordance with the lower temperature
of that season over three or four months, would gradu-
ally bring it to the South European or Mediterranean
continuously-brooded type. Finally, having regard to the
plastic character of its pupa, it might, after a long period of
probation and many vicissitudes, become seasonally double-
brooded. ||
* See ‘‘Silkworms in India,” by E. C. Cotes, Indian Museum Series,
vol. i. I am indebted to Major Alcock, F.R.S., C.I.E., Superin-
tendent of the Calcutta Museum, for these volumes.
+ On this-see Tutt in Ent. Record, vol. x, pp. 211, 212.
+ See Tutt, Ent. Record, vol. vii, p. 6.
§ See Ent. Record, articles on Migration, vol, xiv.
|| In dry countries the peried of hot weather resting, estivation, would
come in as an element, as in the case of Ocnogyna betica, mentioned by
( wa)
The abundance of Lepidoptera in warmer regions, and the
vast number of their individuals, arising out of the frequency
of their broods, would tend to overspills, and the evidence of
actual migration is in favour of its having been mostly in a
direction that would point to these regions as the source.*
I do not suggest that in any of the cases the difficulties are
insuperable, and one can see that even when there is so great
a physiological gulf as that between the summer pupa of a
double-brooded insect and the winter pupa of a single-brooded
one, there are stepping-stones across it, such as variability of
climate in the same locality, variability in individuals, and
possibility of overlapping resulting in fusion by mating of
different generations. And many species are probably at all
times in an unfixed, perhaps intermediate and transitional
condition, such as those single-brooded species, which frequently
under the stimulus of a hot season bring forth wholly or
partially a second brood, often ineffectively, it is true.t
Altering pattern by temperature.
I have referred in general terms to the effect of the
application of temperature in the pupal stage in altering
the colour and pattern of the wings of Lepidoptera. To
attain this result in a high degree it is usually necessary to
apply the temperature from the time when the newly-formed
Dr. Chapman, Ent. Record, vol. xvii, p. 266. As to hibernation, it is not
suggested that this is only caused by cold, or eestivation only by heat or
draught. There is an interesting discussion on this by Mr. Tutt, Mr.
de Vismes Kane, Dr. Ridding, Dr. Dixey and others in Ent. Record, vols.
vii and viii.
* See articles by Tutt in Ent. Record, vol. xiv, pp. 262, 292, and
315. ‘‘ Broadly speaking, the dispersal movements of insects... are...
from sub-tropical to temperate regions. . .. The general trend . . . in the
Northern hemisphere is northerly and in the Southern hemisphere is
southerly.” I have only mentioned migration, which is operative before
our eyes, but I am not unmindful of another potent way in which climate
may affect organisms, and one that has doubtless been highly effective in
the past, and that is where, instead of the organism migrating to the
climate, the climate comes to the organism, in the great secular changes
which have taken place; heat and cold having at different periods char-
acterized most of the Palearctic region, and having had their turn in
starting or moulding the forms that have descended to our times.
+ Warm climates where there are considerable mountains would seem
suited to qualify species for acclimatization, though, as we see, many
of the species generated in such countries remain obstinately single-
brooded,
( ciii_ )
pupa hardens until near the end of the pupal period.
Experiments tend to show that a winter pupa is in a state of
almost absolute torpor for a long period,* perhaps up to the
time when it has been subjected to cold sufficiently long
to bear the application of a forcing temperature, but
that, after that time, though the cold may continue, the
development begins to move very slowly forward, for it will
emerge in a shorter time if the forcing is deferred until long
after that critical period has been surmounted, than if forced
at once. In the case of summer pup, there appears, however,
to be but a short period, if any, of torpor ; my experiments with
S. tetralunaria have shown that the pupal period of this
summer phase can, by icing, be protracted to 22 weeks, and
that changes in the facies of the moths that came from pup
transferred to a warmer temperature began in six weeks, 7. e.
long before the middle of the pupal period.
“ Direct” effect of temperature.
But besides these changes, which—at least most of them—
cannot be ascribed to the direct effect of cold, there are others
which may be called by way of distinction direct, and these I
have found to be produced mainly (in the case of some species
I may say exclusively) in the latter part of the pupal period.
I do not think sufficient regard has been given to what may
thus be called the “direct” effect of temperature during the
pupal period on the colouring of Lepidoptera. But there is
no manner of doubt that it has such an effect, and in many
cases this effect is so considerable that it may have protective
value. As already stated, many of the seasonal differences,
and those much the more striking, must find an entirely
different explanation, that is to say, in the hereditary pre-
disposition of the organism called into operation by the
appropriate stimulus (as the fur of the Arctic hare is whitened
by the advent of winter cold) ; for example, the differences in
A. levana and, to take a marvellous example, those in a species
of Precis, the one phase, P. octavia-natalensis, being on the
* The torpor often lasts for years, as in the numerous cases where
winter pup he for several years in that stage.
(‘ay °)
upper side deep salmon-red with black borders and spots, the
other, P. sesamus, violaceous blue streaked with black, and
with a continuous series of salmon-red spots.*
Affects general colouring.
The effect to which I am calling attention is one which
shows itself especially in the ground colour and general
colouring of the wings and body. It may be stated as
follows :—The higher temperature tends to produce a light
yellowish brown passing on to chestnut or ochreous and
orange, the low temperature dark, and in places blackish
brown, the darkening effect showing itself in general, not
merely in the ground colour, but also in some or most
of the clouds and spots of different colour that diversify it,
giving it in some cases, where these spots consist of single scales,
the appearance of increased irroration. This kind of difference
(frequently associated, in the facies presented by an insect
subjected to severe or lengthened exposure, with the more
deep-seated and often much more striking change of pattern
and colour above referred to as arising out of a hereditary
predisposition that is merely awakened by the stimulus) I
have found in Lepidoptera, so widely separated from each
other as P. machaon, A. cratexgi, EF. cardamines, L. sybilla,
P. atalanta, P. cardui, V. urtice, V. polychloros, Polygonia
c-album, Melitxa aurinia, M. cinxia, M. didyma, Argynnis
paphia, Zephyrus betule, Orgyia antiqua, Lasiocampa quercus
* Mr. Trimen’s Presidential Address, Proc. Ent. Soc., 1898, p. lxxii,
which gathers together all the knowledge on “‘ seasonal dimorphism” up
to that date, since which important additions have been made especially
by Mr. Guy K. A. Marshall,1896-1901. See the joint paper below referred
to pp. 287-585 ; also Dr. Dixey, in Trans. Ent. Soc. 1902, pp.189-218, and
Proc. Ent. Soc. 1902, pp. xv—xviii, on Mr. Loat’s collection, Proc. Ent. Soc.
1903, pp. xi-xiii, and Trans. pp. 141-164; Poulton, Proc. Ent. Soe.
1903, pp. xxxii-xxxiv ; and Lathy, Trans. Ent. Soc. 1903, pp. 183-206.
As to the protective value of the different colouring—extending also to
habits—of the two forms of this species of Precis, see Mr. Guy A. K.
Marshall and Professor Poulton’s joint paper, Trans. Ent. Soc., 1902, p.
424 et seg. In the case of A. levana, [ do not know that more can be
said than that there is probably something advantageous to a summer
butterfly in the general colour and pattern on the upper side, combined
with powerful flight, that is common to so many of them (Limenitis,
Apatura, Satyrids), and that the summer form, prorsa, of Zevana, has a
great resemblance to them in all these respects. And such of these external
appearances as are not in themselves useful, may be the outward and
visible signs, the correlatives, of physiological adaptations beyond our ken.
(See Professor Meldola’s Presidential Address, Proc. Ent. Soe., for 1896.)
Coen)
(and var. callune), Saturnia pavonia, Drepana falcataria,
Z. punctaria (both broods), S. bilunaria, S.- tetralunaria,
S. lunaria (both seasonal forms of these three species), and
Ennomos autumnaria. With several other species I have
found no appreciable difference of this general character
caused by temperature, and in those I have mentioned it
varies very greatly indegree. In some of these, e.g. the Selenias,
especially S. tetralunaria, the difference is striking, being
equal to that between a dead leaf ina dry August and the same,
moist and darkened by a wet winter; corresponding, in fact,
very closely to the difference in appearance between the summer
and spring phases of this insect, either of which can be con-
verted into a colouring approaching that of the other by the
appropriate temperature, a few days being sufficient for the
purpose where it is a case of high temperature which greatly
shortens the pupal stage. I do not hesitate to say that in
many species much the most conspicuous difference caused by
special temperature, experimentally applied, is that which is
caused by this “direct” effect during the latter part of the
pupal period, and a given temperature must have the same
effect in nature if the only difference between the two cases is
that in the one it is applied by man in a laboratory, and in
the other by nature in a field.
In using the term “direct,” I do so by way of dis-
tinguishing it from that previously spoken of as based on
hereditary predisposition. It is not at all the same thing as,
for example, the direct effect of heat in coagulating albumen,
or of light falling on a photographic plate, or developing green
chlorophyll, or the bright colour of a budding flower. The whole
effect on the colour of the imago is caused before its colour begins
to appear through the transparent skin of the pupa. Remove
the pupa into a normal temperature at this time, and the
colour comes out just the same as if the removal had not taken
place. It must therefore be caused through some vital
process * acted on by the temperature at a somewhat earlier
period, though in general later than that at which the pattern
is determined.
* Of a nature, I suggest, resembling that which Professor Poulton has
shown to be concerned in the colouring of pupe by the surroundings
of the larva before pupation.
( evi )
It is perhaps to be expected that this alteration of colour in
the wing should take place during the latter part of the pupal
stage, for it is at this period that the hemolymph or blood
from which the pigment of the scales is formed flows into
them.* This hemolymph, at first yellow and then drab, can
by various chemicals be made to assume many colours, in-
cluding reds, yellows and bright carmine. We have thus the
raw material provided by nature out of which various colours
can be elaborated.
Is tt protective ?
Is this difference in ground colouring which I have de-
scribed protective? 1 think there is ground for saying that it
may be so. It has a strong likeness to the difference between
the general aspect of vegetation in different seasons, between
a hillside parched by an August sun and the dark stems of
the undergrowth of wet woodland ; and the lighter coloured
insect should be better adapted for concealment in seasons
or in regions where the colour of dead vegetation is the
prevailing one.
Can these temperature effects become hereditary ?
There is weighty opinion in favour of the view that some of
the effect produced by abnormal temperature on the facies of
a butterfly of which the pupa has been exposed to it for a
single generation can be passed on to the progeny. Professor
Standfuss adduces evidence to that effect in the case of V.
urtice. Dr. E. Fischer has stronger evidence, derived from
some experiments on Arctia caja. Professor Weismann
accepts the conclusion that in these cases the aberrations
of the parents induced by cold descended in a considerable
degree to the offspring, explaining this by the view that
the germ plasm is affected, some of the “ determinants” being
strengthened, others weakened by it;f and he elsewhere
* A. G. Mayer, ‘‘ Development of Wing Scales,” etc., Bull. Mus. Comp.
Zoology, Harvard College, June 1896, as to the vital structure developing
in the pupa earlier than in the wings. See Ent. Record, vol. i, p. 298.
+ Weismann ‘‘The Evolution Theory,” translated by Mr. and Mrs.
Thomson, vol, ii, pp, 267 and 275, also p. 137. See also his observations
on my experiments on C. phlwas, Neue Versuche, etc., English translation,
p- 10
(‘exit )
observes, ‘“‘it is indubitable that external influences such as
those emanating from the environment or media in which
the species live, are able to cause direct variation of the
germ plasm that is permanent because hereditary.”
But when the effects on the individual are only transient
there would still remain scope for the great factor in evolution,
natural selection, because the tendency to respond in general
colouring to the temperature undoubtedly varies, and, if it
be protective, an individual ready to respond in this way
would have an advantage which natural selection could secure
to the species,
Two varying aspects of Nature.
But, do we find any correspondence between the colouring
produced by high and low temperatures respectively on
Lepidoptera and the general colouring of those found in warm
and cold countries respectively? I think we do.* I have
looked through some of the families genera and species that are
spread over cold regions as well as over temperate and warmer
parts, among them Argynnids and Satyrids, in the Natural
History Museum at South Kensington, and they seem to me to
show the following results. In those from very cold regions
the ground colour tends to a uniform dulness with sparseness
or diminution in size of scales, and consequent “greasy”
appearance, owing to the shining membrane of the wings
showing between the scales, and paucity or indistinctness or
want of force in the markings. In those taken in what may
be described as the cold temperate regions, there is a greater
tendency towards bright light brown ochreous or fulvous, the
sort of colour so very common in butterflies, shown for exam-
ple by the bright parts of the wing of Pararge megexra, or the
female of Epinephile jurtina, and to greater variety of marking.
In those from the warmer temperate countries there is an
* P. cardui is much darkened by exposing the pupa to a low tempera-
ture. On the other hand, specimens placed at a high one have a beautiful
light brown golden hue extending over much the greater part of the hinder
wings and a large part of the forewings, but without a trace of the rosy
colour on the upper wings which gives such a charm to ordinary English
examples when fresh. I was interested in seeing by a notice in the Ent.
Record, vol. xii, p. 156, that an African type is much less rosy than the
ordinary European form.
( eviii )
increased tendency towards brightness in place of sombreness,
and an increasing variety of marking and colour. My know-
ledge, however, of the butterflies of the world is so limited that
I should have distrusted my own conclusions had I not found
support for them in the opinions of several others better
acquainted with the subject whom I have consulted. Perhaps
on this point I may refer to the remarks of your former Pre-
sident, Lord Walsingham (Proc. of Ent. Soc. 1890, p. liv),
where he speaks of the general tendency of Arctic Lepidoptera
to a certain suffusion of markings and to an increase in the
proportion of dull and dingy scales.*
But, whether it is owing to the direct effect of temperature
on the individual, or whether it is fixed by natural selection as
being transmissible and of protective value, or whether there is
a combination of these causes and perhaps others, I think
there is no doubt that in temperate countries as a general rule
the southern species and the summer broods do in their general
colouring vary rather in the direction of light brown, and the
more northern species and the spring broods in the direction
of darker colouring.
And now let me conclude with a few words of apology for
the many imperfections arising from my want of knowledge
on points beyond those which I have found opportunity per-
sonally to investigate. I think you can rely on my state-
ments of fact, culled from a very large number of experiments,
the results of which, however, as a whole, I have not yet had
time to co-ordinate, and one can hardly either gather or put
forward the facts without some attempts toexplainthem. As
to my ventures in this direction, they may serve as materials for
the consideration of others better capable of dealing with such
a subject. I hope what I have put forward may be acceptable
as a contribution, however slight, to the elucidation of some
interesting phases in that perennial conflict between the phylo-
geny of the organism and its environment which prevails
wherever there is life, the ceaseless struggle by which nature
* There seems an exception in the colour orange which is so highly
developed in some of the Argynnids of the Arctic circle; it seems a
brighter kind of orange, however arrived at, from that reached through light
qrown referred to at p. civ.
( -eix 5)
achieves her ends, and from which occasionally some nascent
species, the outcome of submerged and slaughtered millions,
arises, endowed with vigour enough to seize and hold a place
in the select company of the fit ones who survive.
APPENDIX.
TasiE I (p. 1xxxviii).
Regularity of development of summer pupx.
Number Number Days
of indi- Temperatures emerged in pupa
viduals
Pyrameis atalanta
ii 80°-90° F. (27°-32° C.) il 6
2 62°-69° F. (17°-21° C.) 2 18-19
(average 64°)
4 About 56° F. (13° C.) 4 33-85
5 Average about 54° F. (13° C.) 5 43-45
Chrysophanus phlxas
10 80° F. (27° C.) 10 6-74
10 | 90°F, (32° C.) 10 ae
6. | 70° B..(21° G.) 6 11-15
6 Average about 58° F, (14° C.) 6 22-30
10 Average about 56° F. (13° C.) 10 29-33
10 | 27>. (8° G) 3 57-59
Vanessa polychloros
20 85°-90° F. (29°-32° C.) 14 63-7
10 Average 62° F. (17° C.) | 10 | 20-22
10 56° F. (13° C.) 10 40
Zonosoma punctaria
21 90° F. (32° C.) 21 4-5
22 70° F. (21° C.) 22 10-11
17 56° F. (13° C.) 17 22-27
13 45H. (i C)) 10 57-70
Pieris napi
4 90° F. (32° C.) 4 6
4 67> H. (u9F Ch) | 3 | 12-13
PROC. ENT. SOC. LOND., v. 1905. H
TABLE IT (pp. xciv, xev).
( ex )
Forcing winter pupx of A. levana, 1895-6.
EXPERIMENT 1. —Pupated 28 Aug. to 6 Sept. as they did so placed in
refrigerator 46°-50° F. (8°-10° C.) till about 26 Oct.
Transferred
thence to incubator [80°-92° F. (27°-33° C.)] in both at intervals.
Number
transferred,
Dates of
transfer.
16 2?
1 March
ios,
Weeks from
pupation.
Number
emerged.
a |
DOPORATOMONWONDOOMN
Number of days after
transfer until emergence.
| All died ; those on
| 1 Dec. prob-
| ably killed by
accident.
9
104
10-11
7-93
83-9
7-8
6-9
5-6
4-7
4-53
34-44
14-43
EXPERIMENT 2.—Pupated 28 Aug. to 6 Sept. Left in room (rather
cold and fireless one) till 26 Oct., then transferred to incubators
80°-92° F. (27°-33° C.) in lots at intervals.
Number Dates of Weeks from | Number Number of days after
transferred. transfer. pupation. emerged. | transfer until emergence.
26 Oct. 8
3 Nov 9 All died ; those on
25: 12 24 Nov. and 8
8 Dec 14 Dec. probably
Poe 16 killed by acci-
5 Jan 18 dent.
i ae 19
4 2 Feb 22 1 12
4 ON a5 23 1 10
4 1G oe 24 i 11
4 23's 55 25 3 7-11
8 1 March 26 4 3-10
4 sia at 1 7
4 ie ey 28 3 5-11
4 Da 9 29 2 2-5
4 29 30 2 5-10
4 5 April 31 2 3 hours—4} days.
( ex
)
EXPERIMENT 3.—Pupated 28 Aug. to 6 Sept. Left in room till 13 Jan.
(19 weeks), then placed out of doors, and thence transferred to
incubators at 80°-92° F. (27°-33° C.) in lots at intervals.
Number Dates of Weeks from | Number Number of days after
transferred. transfer. pupation. emerged. | transfer until emergence.
4 7 Feb 24 2 10
8 Pee 26 3 S24
8 1 March 27 3 9-11
8 Shick 28 5 10-14
8 Lote 29 5 or 6 7-10
8 yas 30 5 5-9
8 29 ,, 31 5 4-64
8 5 April 32 2 8-9
The results of the above three experiments may be compared with
Prof. Weismann’s experience (Neue Versuche), where of 120 A. levana
pupe placed on the 10 Jan. at the temperature of 27°-30° C., 30 emerged
at different dates between 18 Jan. and 25 Feb.
« ami )
GENERAL INDEX.
The Arabic figures refer to the pages of the * Transactions’; the Roman
numerals to the pages of the ‘ Proceedings.’
The President’s Address is not separately indexed.
GENERAL SUBJECTS.
Acari, exhibition of beetles bearing, li.
Acridium xgyptium, found in Bloomsbury, exhibited, iv.
Acrognathus mandibularis, taken at Woking, exhibited, xxxi.
Africa, exhibition of Pieridx from, liv, lxi, lxvi; contribution to the know-
ledge of the Rhopalocera of, lx; descriptions of new Halticinw and
Galerucine from, lxxvi; new species of Lastocampide from, 313.
Agathidium, exhibition of a specimen of, liv.
Aygrion armatum, exhibition of British specimens of, xxxi.
Agrotis ashworthi, to be ranked as a Woctua, xvii.
» tritict, exhibition of aberration of, lxi.
Algeria, on Hymenoptera-Aculeata collected in, lxiv, 399.
Amara anthobia, from Leighton Buzzard, exhibited, xvi.
America, on new Satyridz from South, xiii, 185.
Amphidasys betularia, exhibition of specimens bred from 4. betularta var.
doubledayaria, vi.
Anthribid beetle, exhibition of specimens with crescentic sulcus on the
prothorax, li.
Apatura iris, exhibited, xviii.
Apion levigatum, exhibited, 1.
35 semzvittatum, exhibited, lxv.
Apterygida arachidis, larve and perfect insect exhibited, xxxi.
Argyresthia tlluminatella, exhibited, xlvi.
Asemum striatum, on the habits of, iv, 165.
Auditors for 1906 appointed, lxv.
Australia, the blind Coleoptera of, xlviii, 365.
Belenois, on the scents of species of, lvii; on variation in, Lxvi.
» severina, on the scent of, and exhibition of alcoholic extracts of the
wings of, liv.
Birara, on Rhynchota from, xliv, 327.
Biston hirtarius, exhibition of aberrations of, xxxi.
Boarmia consortaria, melanic examples exhibited, xxvii.
Bolivar, Senor I., elected Honorary Fellow, xiii.
Brauer, Herr F. M., notice of death of, xiii.
a
( ex )
Brenthidex, exhibition of specimens of, bearing Acari, 1.
British Columbia, butterfly-hunting in, xiii.
Buprestidx, on the coloration of some, liti.
Butterflies facing sun when at rest, xxx.
Butterfly-destroyers in South China, 5.
Canada, butterfly-hunting in, xiii.
Carabus morbillosus, exhibition of a series of, to show its variability, lxv.
Catocala nupta, melanie specimen exhibited, xlv.
Catopstlia florella, on the scent of, lv.
Ceraphronine, hermaphrodite specimen of, exhibited, lxv.
Ceratopterus stahli, exhibited, xviii.
Cerostoma asperella from Leominster, exhibited, Ixvii.
Ceuthorrhynchus contractus var. pallipes, from Lundy Island, exhibited, xlvi.
Ceylon, on Butterflies observed during a tour through, 61.
Chalcididx, on a specimen of, parasitic on Dalader acuticosta, 55.
Chamesyrphus scxvoides, exhibited, xlvii.
Cherry-laurel, leaves of, attacked by fungus, resembling spots on wings of
Kallima, xxxii.
China, on Butterfly-destroyers in South, 5.
Cherocampa ostris, superstitious dread of the larva of, xxiii.
Cicadidx, additions to the knowledge of the, xiii, 191.
Coleophora, exhibition of imagines of species of, and of leaves showing the
mines made by the larve of, xlvii.
Coleoptera, the blind Coleoptera of Australia and Tasmania, x] viii; exhibition
of specimen of, showing device for cross-fertilization of flowers, lxi.
Colias edusa, exhibition of a series bred from C. edusa var. helice, vi.
», electra, on the scent of, lvi.
», philodice, on the resting habit of, xxix.
Collyris emarginatus, larvee of, exhibited, xxii. :
Cordylomera suturalis, exhibition of larve taken from mahogany, and of
a perfect insect of, xlv.
Council for 1906, nomination of, Ix, Ixv.
Crenis boisduvaltt, exhibition of and remarks upon, xv.
Criocephalus, revision of the genus, iv, 145; C. ferus, on the habits of, iv,
165; C. polonicus, on the life history and longevity of, iv.
Cynorrhina fallax, occurrence of, at Nethy Bridge, xlvii.
Cyria imperialis, with supernumerary legs, exhibited, Ixvii.
Dacne fowlert, from Bradfield, exhibited, liv.
Dalader acuticosta, on the egg and early larval stages, and on the Hymeno-
pterous parasite of, 55.
Dasydia tenebraria var. wockearia, taken in company with Erebia alecto var.
nichollz, exhibited, xviii.
Delias, exhibition of species of, Ixvii.
Depressaria emeritella, exhibition of unique British specimen of, Ixvii.
Didinets lunicornis, without wings, exhibited, Lxv.
Dinoderus ocellaris, exhibition of the type specimen of, and remarks upon,
XXVil.
Diploptera collected in Algeria, Ixiv, 399.
Diptera, exhibition of synaposematic, xii.
Douglas, J. W., donation to the Society of a photograph of the late, xlix.
( ey )}
Dytiscus, on the longevity of, iv.
Evebia, exhibition of specimens of various species of, taken in the Pyrenees,
xlix.
» alecto var. nicholli, taken in company with Dasydia tenebraria var.
wockearia, exhibited, xviii.
» gorgone and Evebia goante, resemblance between, xlix.
>, lefebvrez, on the clasps of, xlix,
» melas, exhibited, xviii; on the clasps of, xlix.
» palarica, association of, with LE. evias, in Spain, 37.
» stygne, melanic specimen, exhibited, lii; association of, with E, evzas,
in Spain, 37.
Eronia cleodora, on the scent of, lvii ; on variation in, lxvi.
Eucheira soctalis, exhibition of the social web, pupal shells and perfect
insects of, xix.
Euchloé ewphenoides, on the resting habit of, lxxv.
Fellows, election of, v, xiii, xxvi, xxx, xlv, xlviii, lii, lx, lxiv.
Forficula auricularia, with abnormal cerci, exhibited, liii, lxi.
Fry,A., bequest of his collection of Coleoptera to the British Museum, xiv.
Galerucine, descriptions of new African, lxxvi.
Ganglbauer, Herr L. von, elected Honorary Fellow, xxx.
Genitalia of Noctuid Moths, drawings and slides of, exhibited, xvii; on the
value of the, for the purpose of distinguishing species, xvii; on the
variability in Lepidopterous, xxxiv.
Geographical affinities of Japanese Butterflies, xli.
Gerydus chinensis, the life history of, 1.
Glutophrissa saba, on the absence of scent in, lix.
Gnorimus nobilis, exhibited, xxxi.
Gonepteryx, on scents in the male, xxxv; exhibition of several forms of, xxxvi.
5 rhamnt, resting on underside of bramble leaves, Ixxv.
Gontodoma limoniella, exhibition of the larvz and their cases of, xlviii.
Grammoptera ruficornis, melanic specimen of, exhibited, xvii.
Guatemala, exhibition of insects from, liii.
Haltictnz, descriptions of new African, lxxvi.
Hammerschmidtia ferruginea, exhibition of British specimens of, xlvii.
Haworth, exhibition of specimens labelled by, Ixvii.
Helicontus, exhibition of a series of species of, showing variation in colour, li.
Heliotropism in Pararge and Pyramets, xxviii.
Helophilus transfugus, exhibited, Ixv.
Helops striatus, with abnormal antenna, exhibited, vi.
Hemerophila abruptaria, exhibition of series to show effects of heredity, lxiii.
Hemiptera, description of two new species from Spain, v.
Heriades fasciatus, type-specimen, exhibited, lii; on the coloration in, liii.
Hermaphrodite specimen of Spilomicrinz, exhibited, lxv.
Heterogynis, on the matrivorus habit of, iv, 177; exhibited, v.
Heterosternus rodriguezt, exhibited, liii.
Honorary Fellows, election of, xiii, xxx, Lxiv.
Hyhernia defoliaria, exhibited, iii.
Hydrotwa pilipes, exhibition of both sexes of, vii; H. tuberculata, specimens
captured in Britain, exhibited, vii.
Hymenoptera, exhibition of synaposematic, xii,
( exyy
Hymenoptera-Aculeata, collected in Algeria, Ixiv, 399.
Hystrichopsylla talpx, exhibited, xxvii.
Lliaphanus stephenst, exhibited, xlviii.
India, on butterflies observed during a tour through, 61.
Japan, on the geographical affinities of butterflies of, xli.
Kallima, spots on wings of, resembling leaves attacked by fungus, xxxiii.
Lexmophleus monilis, from Berkshire, exhibited, liv.
Larva of Cherocampa, superstitious dread of, xxiii.
Lasiocampidx, on new African, in the British Museum, xliv, 313.
Lepidoptera, from New Zealand, v, 219; on the variability in the genitalia of,
xxxiv; exhibition of a collection of, from Majorca, li; ona Lepidopterous
parasite from Queensland, lii; on a collection of, made in Maroceo, lii,
369.
Lifu, on Rhynchota from, xliv, 327.
Limnas chrysippus and Pseudacrea pogge?, the numerical proportion of mimic
to model, xxvi, 263.
Lochmexa suturalis, malformed specimen, exhibited, xxxi.
Lundy Island, exhibition of Coleoptera taken on, xlvi.
Lycena, exhibition of specimens of various species of, taken in the Alps, xlix.
3 argus var. hypochtona, taken on North Downs, exhibited, xlvi.
rf argyrognomon, from Rhone Valley, exhibited, xlvi.
= melanops var. wheelert, exhibited and described, ii.
Lygxide, specimen of, exhibited, 1.
Lymecxylon navale, from the New Forest, exhibited, xlvi.
Majorca, exhibition of a collection of Lepidoptera from, li.
Malachius barnevillet, taken in Norfolk, exhibited, iv.
Marocco, on a collection of butterflies and moths made in, lii, 369.
Medon castaneus, exhibited, xxvii.
Megalopus melipona, exhibited to show its resemblance to a bee, xviii.
Megalyra, on a new species of, Ix, 395.
Melanargia lachests var. canigulensts, exhibited, li.
Melanism in Tephrosia and Boarmia, xxvii; in Catocala nupta, xlv; in Erebia
stygne, lii.
Melanophthalma distinguenda, fron Lundy Island, exhibited, xlvi.
Melitzxa aurinia var. cberica, exhibited, li.
Mesosemia eumene, exhibited, ii.
Microdon latifrons, taken at Nethy Bridge, exhibited, xlvii.
Microlepidoptera, on three new genera of, xiii, 245.
Mimetica, exhibition of specimen of, lili.
Mormolyce, larvee of, exhibited, lxxii.
Morpho adonis, a female, exhibited, xviii.
Mylothris, on the scents of species of, lviii.
is agathina, on the scent of, and exhibition of alcoholic extracts of
the wings of, liv.
Myrmeleon formicarius, larva, cocoon, and imago, exhibited, xlvi; exhibition
of diagram of instars and mouth parts of, lxxvi; on the emergence of,
Ixxvi.
Natal, exhibition of butterflies from, xiv.
Neoclytus erythrocephalus, taken from the trunk of an ash tree, exhibited,
XVi.
( cxvi_ )
New Britain, on Rhynchota collected in Birara and Lift, xliv, 327.
New Zealand, on Lepidoptera from, v, 219.
Noctuid Moths, drawings and slides of the genitalia of, exhibited, xvii.
Notodonta ziczac and NV. dromedarius, hybrid between, exhibited, iii.
Ss ztezac var., exhibited, iii.
Nymphalidx, from N. China, apparently mimetic of Hypolimnas misippus,
exhibited, xiii.
Obituary. Henri F. de Saussure; Rev. Francis Augustus Walker; Alexander
Fry, v; Alpheus 8. Packard; Alfred Beaumont, Alfred Preudhomme de
Borre, xvii; Col. L. H. L. Irby, xxx; John William Douglas, George
Bowdler Buckton, xlv; Capt. Frederick Wollaston Hutton, Ix.
Odontopera bidentata ab. nigra, exhibited and remarked upon, Ixvi.
Officers for 1906, nomination of, Ix, Ixv.
Ogyris, monograph of the genus, xxvi, 269.
Oligota granaria, exhibited, ii.
Omophlus betulx, taken at Covent Garden, exhibited, xxxi.
Ornithoptera chimera, with ringed body, exhibited, lxvii.
Orthoptera, exhibition of a specimen of, resembling a dead leaf, liii.
Oxypoda sericea, taken in Dulwich Wood, exhibited, v; O. négrina, exhibited,
v; O. exigua, exhibited, v.
Panorpa cognata, exhibited, iv.
Pantodinus klugi, exhibited, liii.
Panurgqus moricet, exhibited, lii.
Papilio hesperus, exhibition of specimens from Entebbe with unbroken tails, xii.
», macleayanus, exhibition of specimens to illustrate the use of directive
markings, xviii.
Pararge, heliotropism in, xxviii.
Parasitic Lepidopteron from Queensland, lii.
Phaneroptera quadripunctata, from Chester, exhibited, li.
Photography, on the three-coloured process of, xxx.
Phycochus graniceps and P. sulcipennis, exhibited, xlviii.
Pieridx, ou the resting habit of, xxix; exhibition of African specimens of, liv,
lxi, Ixvi; on the scents of, liv.
Pieris napt, on resemblance of, to flower of Leucojum, Ixxv.
» vap#x, on the resting habit of, Lxxiii.
Pinacopteryx, on the scents of species of, lviii; on variation in, Ixvi.
ss pigea, exhibition of, and remarks upon, xv.
Plusiotis adelaida, exhibited, liii.
Prionocyphon serricornis, perfect insects and larva, exhibited, liv.
Pseudacrxa poggei and Limnas chrysippus, the numerical proportion of mimic
to model, xxvi, 263. .
Pseudophyllidex, exhibition of a specimen of, xxxi.
Psychoda sexpunctata, on the structure and life history of, xxx, 293.
Psylliodes luridipennis, from Lundy Island, exhibited, xlvi.
Ptinus pusillus, from Edmonton, exhibited, lxv.
Pupe, vitality of when submerged in water, xxiii.
Pyrameis atalanta, bred in winter, exhibited, xi; on the white spot on
specimens of, xii; heliotropism in, xxviii.
Rhodesia, on new species of Lepidoptera from Northern, 251,
Rhopalocera, contribution to the knowledge of African, lx.
( ‘exvii )
Rhynchota, collected in Birara and Lifu, New Britain, xliv, 327.
Sarawak, exhibition of insects from, 1.
Saturnia, cocoons and perfect insects of hybrid species of, exhibited, vii.
Satyridex, descriptions of new 8. American, xiii, 185.
Satyrus semele, exhibited, xviii.
Scents, in the male of Gonepteryx, xxxv; of African Pieridex, liv; in various
insects, lix.
“Season,” on the word as applied to forms of insects, lxiii.
Selenta bilunaria, on the resting habit of, xxix.
Spain, entomological excursion to, v, 37; description of two new species of
Hemiptera from Spain, v.
Spilomicrinz, hermaphrodite specimen of, exhibited, Ixv.
Stenobothrus, exhibition of specimens taken in Spain, mostly with mutilated
wings and elytra, xvi.
s rufipes, exhibited, xlvi.
Stenopterys hirundinis, exhibited, Ixvi.
Stenus osstum var. nsularts, from Lundy Island, exhibited, xlvi.
Strawberry, leaves of, attacked by fungus, resembling spots on wings of
Kallima, xxxii.
Synchloé hellica, on the scent of, lix.
Tzeniocampide, on the dissimilarity in genitalia of, xvii.
Tasmania, the blind Coleoptera of, xlviii, 365.
Tephrosta consonaria ab. nigra, exhibited, xxvii.
Teracolus, on the scents of various species of, lvi; exhibition of African species
of, lxi; on variation in, lxvi.
x annex, exhibition of, and remarks upon, xv,
fp auxo, exhibition of, and remarks upon, xv.
Thais, on the pupal suspension of, v, 203.
Tortrixz pronubana, exhibition of, and on its occurrence in England, Ixiii.
Tryphsena comes, exhibition of three generations of, Ixvii.
Tryphonides schizodonti, on the group, Ixxvi, 419.
Vanessa anttopa, exhibited, xviii.
Vice-Presidents, nomination of, ii.
Vitality of Pupz submerged in water, xxiil.
Walker, Commander J. J., degree of M.A. conferred upon, xlv.
Xenarthra cervicornis, exhibited, xxvi.
Zegris eupheme, on the resting habit of, Ixxv.
Zonosoma pendularia, pupa cases of, exhibited, to show the variation in the
position of the silken girth, xxxii.
( exyii ))
SPECIAL INDEX.
The Arabic fiyures refer to the pages of the * Transactions’ ; the Roman numerals
to the pages of the ‘ Proceedings,
abbreviata (Enithares), 339
abd-el-kader (Satyrus), 375
abdominalis (Telephorus), 45
abdominator (Homoporus), 427, 433
abeillei (Phytocoris), 50
abietaria (Boarmia), 388
Abisara, 94
Abraxas, 130
abrota (Ogyris), 269, 271, 273, 281,
284, 289, 290, 292
abruptaria (Boarmia), 388
’ (Hemerophila), 1xiii
absinthii (Siphonophora), 434
acacie (Thecla), 378
Acanthaspis, 110
aceris (Neptis), 95, 96, 98, 99
Acherontia, 386
achine (Teracolus), lvi, 1xii
Acidalia, li, 389
Acraa, 265
acreida (Lymanopoda), 188
acreina (Erikssonia), 255
Acridium, iv
Acrobasis, 392
Acrognathus, xxxi
Acronycta, 384
Acropteris, 97
actea (Satyrus), 41, 42, 375
actzeon (Adopiea), 41, 380
acte (Ticherra), 97
Actobius, 44
aculeata (Mordella), 45
acuminata (Aflia), 52
= (Gonepteryx), XXXViii, Xxxix
acuticosta (Dalader), 55-59
Acylophorus, 51
adelaida (Plusiotis), liii
Adelphocoris, 50
adherbal (Lycenesthes), 253
adippe, var. chlorodippe (Argynnis),
41, 42
adippe, var. cleodippe (Argynnis), 41,
42
admixtalis (Bradina), 71
Adonia, 50
adonis (Morpho), xviii
Adopza, 41, 380
adreptella (Heterocrossa), 235
Adrisa, 358
adustum (Anthobium), 44
wgeria (Pararge), xxvili, 66, 135
wgon (Lyciena), xliii, xlvi
egyptium (Acridium), iv
Alia, 50, 52
zneicollis (Corcebus), 49
zneus (Paracymus), 51
,» (Saprinus), 44
enone (Ogyris), 269, 272, 287, 289
wequilineata (Docirava), 68
fEschines, 343
wthiops (Erebia), 11
zthonellus (Crambus), 224
affine (Megymenum), 354
affinis (Pterochilus), 405
Agabus, 48, 51
Agalope, 65, 68
agamemmnon (Papilio), 8, 118, 124
Agapophyta, 354
Agathidium, liv, 48
agathina (Agrotis), 1xi
hs (Mylothris), liv, lviii, lix
ageliella (Depressaria), 433
aglaia (Argynnis), 40, 42, 112
»» (Delias), 8
aglaope (Ganoris), 137
Agonoscelis, 355
agrestis (Criocephalus),
167
Agrion, Xxxi
Agrotis, li, xi, 81, 382
akbarella (Oligochroa), 71, 79
Alena, 252
158, 160,
€ ‘ex |)
albicinctus (Homoporus), 432 amymone (Eupleea), 5
albicoxa (Promethus), 429, 436 amyoti (Cletus), 351
albida (Oreopsyche), 391 », (Gonocerus), 351
», var. lorquiniella (Oreopsyche), | anadema (Lycenesthes), 258, 262
391 Anadiasa, 314, 326
albitarsella (Coleophora), xlvii aneema (Borkhausenia), 241
albithorax (Priocnemicoris), 352 Anaitis, 388
on var. patricius (Priocnemi- | analis (Euptychia), 185, 190
coris), 352 », (Pompilus), 81
albofasciatus (Phytocoris), 50 anastrella (Trachypepla), 237
albolineatum (Ancyrosoma), 52 anaura (Castalius), 95
albomaculata (Lymanopoda), 188 Anchomenus, 43
albonotata (Pedaliodes), 189, 190 Ancistrocerus, 409, 410
albosignatus (Bassus), 425, 431 Ancyrosoma, 52
alcez (Carcharodus), 41, 381 angustalis (Cledeobia), 41
alciphron, var. gordius (Chryso- | angustatus (Esolus), 47, 48
phanus), 378 angustior (Hoplopus), 408
alcon (Lyczena), iii 33 (Odynerus), 408
aleyonipennella (Coleophora), xlvii angustus (Emblethus), 46, 50
alecto (Erebia), xlix Anisoplia, 44
», var. nicholli (Erebia), xviii, | Anisorrhynchus, 45
xlix Anisoscelis, 352
Aleochara, 44 anne (Teracolus), xv, lvi, ]xi
aletis (Hypercallia), 235 annulatus (Bassus), 425, 431
algeriensis (Psiloglossa), 400 anomalus (Phthorimus), 420, 430, 436
algira (Ophiusa), 384 Anommatus, 367
ali (Hesperia), 381 Anopheles, 110, 114, 306
alienum (Cryptostemma), 50 ansorgei (Iridopsis), 254
alluaudi (Umjaba), 196, 197, 202 Antestia, 355
almana (Precis), 72, 74, 76, 78, 84, 88, | Anthaxia, 49, 51
89, 100, 109, 120, 133, 134, 139, | antherinus (Anthicus), 45
140, 141, 142, 143 Anthicus, 45
alpestris (Lionotus), 414 anthobia (Amara), xvi
», (Odynerus), 414 Anthobium, 42, 44
alpinus (Homoporus), 426, 432 Antholinus, 45
alternans (Baptolinus), 48 Anthophagus, 42, 44
althee (Carcharodus), 381 antinorii (Uranothauma), 253
alticola (Cosmopsaltria), 200 antiopa (Vanessa), xviii, xxix, 72,
alveus (Hesperia), 41, 381 6
Amara, xvi, 42, 43 antonia (Gonepteryx), XXXvli
amaryllis (Ogyris), 272, 285, 287, 289, | aoris (Cirrochroa), 97, 98
291, 292 Apatura, xviii, xliii, 117
amatus (Teracolus), 142 Aphantopus, 40
», var. modestus (Teracolus), 107 | Aphanus, 46, 50, 346
Amblypodia, xliii Aphneus, 85, 90, 252
amen (Oncocephalus), 340 Aphodius, 43, 44, 48
amenaida (Pentila), 252 apiarius (‘Trichodes), 49
amenena (Hypercallia), 236 apicata (Acanthaspis), 110
Amicta, 40 Apion, 1, Ixv, 45, 49
amictus (Antholinus), 45 apollo (Parnassius), 40
amine (Zeritis), 252 Apopestes, 385
ammios (Trichodes), 49 Aporia, xxi
Ammeecius, 43, 44, 48 appiades (Kuthalia), 95
Amphidasys, vi Appias, 76, 139
Amphipyra, 384 Apterygida, xxxi
Ampittia, 119 apulia (Lymanopoda), 188
Amygdaloptera, 387 aquaticus (Notiophilus), 42, 43
amymone (Crastia), 137 arachidis (Apterygida), xxxi
( exe: )
Aradus, 50, 342
Araschnia, xxiv
arcanioides (Coenonympha), 376
arcanius (Ceenonympha), 40, 42
arcas (Lycena), iii
Arctia, 381
areolatus (Perileptus), 43
», (Promethus), 485
areste (Arhopala), 94 ©
argia, var. similis (Zizera), 83
argiades (Lycena), xliii, 39, 41
», var. parrhasius (Everes), 133
Argina, 84
argiolus (Cyaniris), 41, 67, 380
», (Lycena), xliii
argus (Lycena), xlili, xivi
», var. hypochiona (Lycena), xlvi
Argynnis, xlii, xliii, 40, 41, 42, 65, 66,
68, 69, 73, 74, 81, 83, 97, 98, 102,
112, 113, 128, 132, 134, 374
Argyresthia, xlvi
Argyria, 226
argyridana (Hiposcritia), 94
argyrognomon (Lycwna), 42
Arhopala, 94, 95, 97
ariadne (Ergolis), 89, 92, 109, 115, 119,
127
Arictus, 342
arion (Lyciena), il, iii, 41
aristolochiz (Papilio), 78, 83, 90, 99,
100, 103, 105, 106, 107, 108, 109,
111, 114, 118, 123, 138, 184
armatissimus (Polydidus), 339
armatum (Agrion), xxxi
Aromia, 48, 45
arrogans (Scutellera), 859
Artaxa, 71
artemisiz (Heterogaster), 46, 50
arvensis (Carabus), 43
ascanli (Lixus), 52
Ascerodes, 234
Asclepias, lxi
asela (Crastia), 123, 125, 127, 131, 137
Asemum, iv, 145-176
ashworthii (Agrotis), xvii
65 (Noctua), xvii
aspasia (Gonepteryx), xxxvili, xxxix
asperaria (Gnophos), 888
asperatus (Criocephalus), 147, 158, 159,
160
asperella (Cerostoma), lxvii
Aspilates, 387
assimilis (Oncocephalus), 340
Astacops, 348
Astemma, 345
asterie (Precis), 133
astola (Neptis), 68, 81, 82, 94, 95, 97,
99
astrarche (Lyczena), 41, 42, 379
var. calida (Lycena), 380
asturiensis (Otiorrhynchus), 42, 45
=, (Timarcha), 46
oe (Zabrus), 44
Asura, 129
atalanta (Pyrameis), xi, xli, 65, 373
Atella, 65, 68, 70, 74, 76, 89, 100, 103,
104, 115, 129
ater (Geocoris), 46
aterrimus (Pterochilus), 403
athalizeperda (Bassus), 437
athamas (Charaxes), 98
Atheta, 48
Athous, 42, 45
Athyma, 66, 68, 73, 81, 95
atlantica (Lymantria), 386
atlanticaria (Boarmia), 388
atlites (Precis), 89, 125, 183, 140,
143
atomarium (Apion), 49
atra (Hispa), 50
Atractomorpha, 102
atricornis (Riptortus), 351
atropos (Acherontia), 386
(Ancistrocerus), 409
», (Odynerus), 409
Attelabus, 49
attenuata (Strangalia), 43, 45
atticus (Tagiades), 114
atymnus (Loxura), 90, 92
Ss var. arcuata (Loxura), 126
Aufidus, 332, 363
Augiades, 42
augias (Telicota), 79, 83, 85
Aulocera, 64, 68, 69, 135
aumonti (Carabus), lxv
auraria (Vespa), 72
aurata (Zygena), 391
aureocincta (Callidea), 359
auricularia (Forficula), liii, xi
var. forcipata (Forficula),
29
>
1xi
aurinia, var. iberica (Melita), lii
auritus (Exochomus), 50
aurolimbata (Orgyia), 41
australis (Cimex), 852
os (Criocephalus), 158, 159
auxo (Teracolus), xv, xvi, xxxviii, lvi,
lxii
avellanze (Orchestes), 49
azrae] (Physoderus), 339
Badacara, 112
badia (Metachrostis), 79
badiipennella (Coleophora), xlvii
badium, liv
beetica (Anisoplia), 44
(Lycena), 81, 111
93
( text) )
beeticus (Polyommatus), 65, 71, 72, 74,
77, 85, 105, 106, 108, 109, 131, 134
Beturia, 329, 330
bajulus (Hylotrupes), 51, 172
balanica (Yphthima), 72, 139
Balaninus, 49
baldus (Yphthima), 114
ballus (Thestor), 378
balteatus (Syrphus), 432, 433
bambuse (Telicota), 6, 90
banksii (Scutellera), 360, 362
», (Tectocoris), 360
Baptolinus, 48
Baracus, 130, 132
barbarus (Orthomus), 43
Baris, 42, 45
barnardi (Ogyris), 269, 272, 285, 289,
291, 292
barnevillei (Malachius), iv
Barynotus, 45
basalis (Rivula), 132
basiventris (Eurinome), 357
5 (Pentatoma), 357
basochesiata (Cidaria), 389
Bassus, 419, 420, 425, 431, 432, 436,
437, 438
batesi (Euptychia), 185
baton, var. abencerragus (Lycna), 379
bazalus (Arhopala), 95
bejarensis (Erebia), 10, 11, 16, 17, 27,
30, 33
belemia (Euchloé), 372
Belenois, liv, lvii, lviii, lix, lxvi, 65,
67, 70, 71, 76, 78, 79, 80, 82, 103,
104, 105, 106, 107, 134, 187, 144
Bembidium, 42, 43, 48, 46, 51
benjamini (Rhopalocampta), 7
Beosus, 50
Beralade, 319-322, 326
Betarmon, 44
bettoni (Beralade), 320, 322, 326
betulee (Omophlus), xxxi
», (Rhynchites), 45
betularia (Amphidasys), vi
‘5 var. doubledayaria (Amphi-
dasys), vi
Biblia, 108
bicinctus (Lygzus), 349
bicolor (Stenobothrus), xvi
bicornis (Hymenopus), 57
bidentata ab. nigra (Odontopera), Ixvi
bidentatus (Anisoscelis), 352
ys (Lionotus), 410
by (Lygeus), 352
5 (Odynerus), 410
Bidessus, 51
bifasciatum (Rhagium), 161, 163, 167,
168, 169
biglobosa (Hypera), 42, 45
biguttata (Titubcea), 49
biguttatus (Bassus), 419
a (Homoporus), 427, 432
$3 var. deplanatus (Homo-
porus), 432
3 var. rufipes (Homoporus),
432
Bilactis, 113
billiardiens (Calliphara), 362
bilunaria (Selenia), xxiv, xxv, XXvi,
XX1X
bimaculatus (Homoporus), 483
nF (Olibrus), 48
biocellata (Kuptychia), 187, 190
bipunctata (Agapophyta), 354
bipunctifera (Schcenobius), 110
birare (Phyllyphanta), 335
birarensis (Clovia), 330, 331, 363
biscutellatus (Ceuthorrhynchus), 49
bisignatus (Olibrus), 48
bispinosus (Lionotus), 413
PP (Odynerus), 413
Biston, xxxi
bituberculatus {Dolichoderus), 1
bivittatus (Telephorus), 45
bizonarius (Homoporus), 426, 432
Blanaida, 135
blanchardianus (Lionotus), 412
3 (Odynerus), 412
i (Quiroguesia), 73
blattariz (Cionus), 45
Boarmia, xxvii, 388
bochus (Jamides), 98
beetica (Lampides), 41, 42
beeticus (Lampides), 379
boisduvalii (Crenis), xiv
Bolboceras, 104
bolina (Hypolimnas), 62, 70, 71, 76, 79,
82, 83, 84, 117, 125, 134
bolivari (Oncotylus), 50
boliviana (Euptychia), 187, 190
Bombus, 72, 94
Bombycia, 383
Bombycopsis, 322, 326
bombyliformis (Sesia), 109
bonvouloiri (Aphodius), 43, 44, 48
borealis (Homoporus), 426, 482
Borkhausenia, 239, 240, 241
bornemanni (Saturnia), viii
Bothrideres, 47, 48
bouvieri (Yanga), 194, 202
Brachycoleus, 50
Brachyderes, 49, 52
Brachyplatys, 362
Brachyrhynchus, 342
Brachytemnus, 49
Bradina, 71
( oma )
brassice (Ganoris), lvi, lvii, lviii, 72
(Pieris), xxxii, 40, 371
var. cheiranthi (Pieris), 371
», var. wollastoni (Pieris), 371
brevicollis (Meloe), 45
e5 (Vesperus), 49, 51, 52
brevicornis (Homoporus), 422, 427, 434
brevitarsis (Homoporus), 426, 432
briseis (Satyrus), 374
Brotolomia, 384
brunneipennis (Pentatoma), 355
J (Plautia), 355
brunneus (Agabus), 51
bryaula (Gymnobathra), 238
bucephala (Phalera), xxiv, 387
var. bucephalina (Phalera),
99
99
387
buprestoides (Spondylis), 47, 49
burmeisteri (Myodocha), 350
x (Leptocorisa), 350
Byrrhus, 44
Bythoscopus, 334
cacaliz (Orina), 46
eacica (Morpho), xix
cadderensis (Nematus), 438
Caduga, 95, 96
Ceerois, 117
cerulea (Euptychia), 186
ceerulea (Temnochila), 51
ceruleocephalus (Rhynchites), 52
cresum (Omalium), 44
calais (Teracolus), 79, 1389, 144
Calathus, 42
c. album (Vanessa), xlili, 373
calcaratus (Miris), 50
calice (Castalius), 253
calidasa (Limenitis), 129
Callidea, 359
Callidium, 158
Callidryas, xxxvii, 117
Calliphara, 362
Callophrys, 378
Callopistria, 385
Calocoris, 46
Calysime, 125
Camena, 93
Campanotus, 270
campestris (Cicindela), 43
Camptobrochis, 50
Camptopus, 52
canalensis (Heterogynis), 178, 180, 184
canescens (Pericoma), 311
canidia (Ganoris), 67, 70, 71, 72, 73, 81,
95, 97, 111, 134, 137
cantabrica (Chrysomela), 42, 46
5 (Rhagonycha), 45
cantabricus (Haptoderus), 43
Cantao, 361
:
capensis (Chondrostegoides), 324, 325
$5 (Taragama), 315, 326
Caprona, 117, 129
Capsus, 46, 50
Carabus, Ixv, 42, 43, 46, 47
Caradrina, 93
carbonarius (Hister), 44
ss (Stenus), 44
Carcharodus, 41, 381
cardamines (Euchloé), lxxv
cardioderus (Harpalus), 42, 44
cardui (Pyrameis), xii, xxix, 66, 72,
82, 94, 111, 134, 373
carinata (Lareynia), 47, 48
carinatus (Deronectes), 51
carinulatus (Bassus), 437
carnifex (Proteodes), 236
carniolica (Zygiena), 391
5 var. orana (Zygzena), 391
carnipennis (Gonepteryx), xxxvil
caroli (Hoplopus), 406
5, (Odynerus), 406
carpetanum (Bembidium), 47, 48
Carpocoris, 46, 50
carrensi (Cimex), 354
Cassida, 50
Castalius, 95, 117, 120, 188, 253
castanea (Liodes), 48
castaneus (Medon), xxvii
castilianum (Doreadion), 45
castrensis (Malacosoma), ]xix
Catacanthus, 354
catalaunaria (Tephrina), 110
cataxesta (Scoparia), 231
eatharina (Ogyris), 283
Cathormiocerus, 47, 49
eatilla (Catopsilia), 84, 138
Catocala, xlv, 41, 384
Catochrysops, 71, 79, 83, 85, 88, 90,
103, 108, 109, 253, 258, 259, 262
Catophaga, 111, 112, 115, 116, 123,
127, 128, 130, 134, 185, 187, 142,
143
Catopsilia, xxxvii, lv, 7, 70, 75, 79, 80,
81, 84, 88, 89, 92, 99, 101, 105, 107,
109; 110; 115, 118, 119, Moi;e22,
133, 134, 138, 139, 140, 141, 142,
143, 144
caudata (Megalyra), 396
caudatus (Homoporus, 426, 432
cauta (Phylacodes), 242
cazurroi (Stenobothrus), xvii
Celenorrhinus, 95, 129
celeno (Lampides), 90, 92, 95, 98, 109,
117, 119, 120, 126, 127
var. alexis (Lampides), 90, 99
»» Var. conferanda (Lampides), 109,
117, 120
2?
( ‘oxxin. )
celerio (Chcerocampa), 77, 386
Celonites, 399
cenis (Zonosoma), 96
centaurus (Arhopala), 95
Centrocoris, 50, 52
Cephalallus, 147, 148, 163
Cephalocrius, 146, 147, 149, 164
Cephanodes, 109
Ceratopterus, xviii
Cercotmetus, 342
Cerocoma, 47, 49, 52
Cerostoma, Ixvii
cervicolor (Athous), 45
cervicornis, (Xenarthra), xxvi
cervina (Anadiasa), 314, 326
cespitalis (Herbula), 69
Cethosia, 125
Ceuthorrhynchus, xlvi, 45, 49
ceylonica (Nepheronia), 117, 127
i (Parantica), 121, 123, 124,
1133), 137
a (Yphthima), 113, 114, 121,
133
chalconotus (Agabus), 51
chalybeea (Pierella), 185
Chamesyrphus, xlvii
chambereti (Antestia), 355
5p (Strachia), 355
chamomille (Cucullia), 383
championi (Taphropeltus), 50, 52
charadrias (Stenoptilia), 232
Charaxes, 98, 251, 373
charina (Pinacopteryx), lviii, Ixvi
charonia (Vanessa), 112, 113
chenui (Yphthima), 111, 113
chevrolati (Cathormiocerus), 47, 49
ee (Hymenoplia), 44
Chilades, 79, 83; 85, 101;
119
Chilena, 319, 322
chimera (Ornithoptera), xvii
chinensis (Gerydus), 1-4
chionogramma (Xanthorhoe), 220
Chironomus, 293, 298, 306, 307, 308,
309, 310, 311
Chittira, 131, 137
Chlenius, 44, 47, 48
chlamydota (Scoparia), 230
ehloradelpha (Borkhausenia), 239
chlorias (Xanthorhoe), 220
Chloridea, 382
Chloroclystis, 219
chloroticus (Lionotus), 410
ae (Odynerus), 410
Choerocampa, xxili, 77, 386
Chondrostega, 313, 323, 324
Chondrostegoides, 313, 323,
325
103,
324,
chrysippus (Limnas), xxvi, 72, 73, 75,
78, 80, 84, 87, 89, 97, 100, 104, 105,
106, 108, 109, 113, 115, 118, 120,
123, 132, 133, 137, 263-268
an var. dorippus (Limnas), 109,
264, 265, 266, 268
35 var. klugi (Limnas), 264,
268
Chrysocoris, 126, 359
chrysogramma (Borkhausenia), 241
Chrysomela, 42, 46, 50
Chrysophanus, 65, 66, 68, 81, 82, 378
chrysorrheea (Euproctis), 386
chrysozona (Mamestra), 383
Cicada, 199, 328, 330
Cicindela, 42, 43
Cidaria, 67, 389
Cimex, 346, 349, 352, 354, 355, 360
cinctus (Homoporus), 426, 432
cinerea (Haplopacha), 324
cingulatus (Cimex), 346
s (Dysdercus), 345, 346
5 (Homoporus), 432
cinnamopterus (Pilophorus), 50
Cionus, 45
Cirrochroa, 98, 125, 127, 134
Cistus, 52
Cledeobia, 41, 392
cleobis (Camena), 93
cleobule (Gonepteryx), Xxxvii, xxxix
cleodora (Eronia), lvii, Ixvi
Cleogene, 39
Cleonus, 45, 52
cleopatra (Gonepteryx), XXXV, XXXvVi,
LOCA M Soba all, db, Bis)
Cleophana, 383, 384, 393
Clerus, 52
Cletus, 451
Clinteria, 72
Clovia, 330, 331, 363
clytia (Papilio), 5
enejus (Catochrysops) 71, 79
Cneorrhinus, 46, 49
Cnethocampa, 183
c-nigrum (Agrotis), 382
coarctata (Eumenes), 400
an var. dubius (Eumenes), 400
Coccinella, 50
Coccoteris, 357
Cochlidion, xxiv
cocksi (Salda), 50
Coctoteris, 357
coecus (Halorhynchus), 368
Coenonympha, xliii, li, 24, 40, 41, 42,
376, 377, 393
cognata (Cirrochroa), 125, 127, 134
», (Panorpa), iv
cognatus (Promethus), 429, 436
( cGxxiv )
Coleophora, xlvii, xlviii
Colias, vi, xxix, xliii, lv, 40, 65, 68,
69, 72, 78, 78, 94, 112, 113; 372
Collyris, xxii, xxiii
comes (Agrotis), 382
. 9, (Tryphena), lxvii
comma (Augiades), 42
communis (Panorpa), iv
compressus (Phthorimus), 421,
436
Compsistis, 243
conchylalis (Cydalima), 100
concolor (Gynandrophthalma), 42, 45
concordia (Crenidomema), 251
confusus (Homoporus), 433
consobrinus (Hoplopus), 406
5; (Odynerus), 406
consonaria (Tephrosia), xxvii
consortaria (Boarmia) xxvii
conspersa (Bombycopsis), 322
conspersus (Agabus), 48
constrictus (Leistus), 46, 47
contactella (Heterocrossa), 235
“e (Tinea), 235
continua (Beralade), 320, 321
contracta (Catochrysops), 103
contractus (Taphropeltus), 52
5 var. _pallipes
rhynchus), xlvi
conversa (Catocala), 41, 3884
a var. agamos (Catocala), 384
convolvuli (Sphinx), 79
cooksoni (Cyclopides), 260
s (Erikssonia), 255, 262
i (Mycalesis), 251, 262
Cooksonia, 256, 257, 262
Copris, 48
Coptosoma, 362
Coranus, 50
cordula (Satyrus), 26
Cordylomera, xlv
core (Crastia), 78, 83, 87, 89, 91, 100,
108, 110, 117, 128; 187
»> var. vermiculata (Crastia), 83
coreanus (Criocephalus), 153, 154, 164
coreoides (Narmada), 117
coriaceus (Criocephalus), 145, 158
Corixa, 50, 52
Corizus, 50
Corcebus, 49
corolla (Syrphus), 432
coronillaria (Pseudoterpna), 389
corticea (Euxoa), 69
coruscans (Geotrupes), 47, 48
Corydalis, 307
corydon (Lyczna), 41
Corymbites, 42, 45
Corymica, 130
430,
(Ceuthor-
Coryphocera, 129
Cosmophila, 132
Cosmopsaltria, 200, 330
costata (Idea), 130, 132
cracce (Apion), 49
Crambus, 225, 226
crassicornis (Corizus), 50
crassicrus (Homoporus), 424, 428, 434
crassipes (Phymata), 50
Crastia, 78, 83, 87, 89, 91, 100, 108,
110, 117, 123, 127, #81, 137
erategi (Aporia), xxi
>» (Bontia)> xx
crawshayi (Lycznesthes), 253
(Tetropium), Lxvii
Creatonotus, 126,
crenzus (Crambus), 225
Crenidomema, 251
Crenis, xiv, 251
cribella (Myelois), 392
cribraria (Argina), 84
», (Euprepia), 381
»» var. punctigera (Euprepia),
381
crinalis (Zanclognatha), 385
crino (Papilio), 118
Criocephalus, iv, 49, 51, 145-176
cristata (Taragama), 315
cristatus (Pterostichus), 43
»» var. cantabricus (Pterostichus),
43
crocale (Catopsilia), 138
Crocisa, 92
Crocydopora, 224
Cromna, 334
Cryptocephalus, liii, 42, 45, 46, 49,
50
Cryptops, 310
Cryptestemma, 50
Cteniopus, 45
Cucullia, 383
Culex, 110, 298, 307
cuneata (Taragama), 316
Cupha, 125, 127
cupreus (Peecilus), 43
curculionoides (Attelabus), 49
Curetis, xliii, 90, 134
cursitans (Pachytomella), 46, 53
curvatula (Drepana), xxiv, xxv
Cuspicona, 356, 357
Cyamophthalmus, 146
eyanicornis (Tachypus), 48
Cyaniris, 41, 66, 68, 69, 95, 99, 129,
130, 132, 380
cyclopicus (Crambus), 225
Cyclopides, 260, 262
cyclopius (Erebia), 13
Cydalima, 100
(-texxv +)
Cydnus, 359
Cymbalophora, 381
Cymindis, 42, 44
Cymus, 50
Cynorrhina, xlvii
Cyphodema, 50
cypris (Morpho), xix
Cyrestis, 96
Cyria, lxvii
Cyrtonus, 47, 50
Dacne, liv
dactyloptera (Trisceedecia), 247
daira, var. nouna (Teracolus), 372, 393
Dalader, 55-59
damo (Ogyris),
Danaca, 45
daniie (Teracolus), 119, 142
Danais, 73, 123, 1381
dantici (Lionotus), 410
», (Odynerus), 410
daphne (Argynnis), 42
daplidice (Pieris), 40, 372
Darbanus, 340
daretis (Lethe), 128, 182, 134
darsius (Ornithoptera), 123, 128
Dasychira, 132
Dasydia, xviii
Dasytes, 42, 45
davendra (Epinephele), 72
deceptor (Criocephalus), 151, 159, 164
AA (Hoplopus), 407
is (Odynerus), 407
decorum (Bembidium), 48
defina (Lycenesthes), 253
deflexicollis (Hydrocyphon), 45
defoliaria (Hybernia), iii
degeneraria (Acidalia), li
deidamia (Pararge), xliii
Deilephila, 386
Deiopeia, 72, 81, 84, 109, 110
dejeani (Cleophana), 383
,, (Zonabris), 49
281
Delias, lxvii, 8, 79, 83, 84, 86, 87, 90,
91, 92, 101, 118, 123, 133, 137
Deltomerus, 42, 43
depilis (Byrrhus), 44
deplanatus (Bassus), 437
ar (Homoporus), 427, 438
Depressaria, Ixvii
depressus (Aphodius), 44
Dermestes, 77
Deronectes, 47, 48, 51
desfontainii (Melitza), lii
detersa (Enome), 108, 110
deva (Pratapa), 85, 134
devius (Microdon), xlvii
deyrollei (Carabus), 48
Diadema, 70
PROC. ENT. SOC. LOND., v. 1905.
diana (Platylomia), 201
Diapria, lxv
Dichorrhagia, xliii
Dichotrachelus, 47, 49
Dichromodes, 222
dichrous (Melanotus), 49, 51
Dicraneura, 387
Dictyophara, 336
Dicyphus, 50
Didineis, lxv
didyma (Melitea), 373
Pe var. deserticola (Melitéa), 374
didymus (Agabus), 51
Dieuches, 110
diffluens (Cleophana), 384
ae var. maroccana (Cleophana),
384, 393
diffusa (Eupterote), 126
dilecta (Quartinia), 400
dilectus (Cyaniris), 99
dimidiatipennis (Eumenes), 105, 106
dimidiatus (Homoporus), 428, 434
Dindymus, 344, 345
Dinoderus, xxvii
diophthalmus (Tectocoris), 361
dipsacea (Chloridea), 382
Dipsas, xliii
Dipterina, 233
Diptychophora, 227
discretus (Hydroporus), 51
dispar (Lygzeus), 349
», (Lymantria), 386
5, (Oncopeltus), 349
disputaria (epbrinta); 79, 108
dissimilis (Papilio), 5, 8, 124, 134
distans (Tagiades), 114
distincta (Anadiasa), 315
a (Philudoria), 315
distinguenda (Melanophthalma), xlvi
(Taragama), 315, 326
dives (Chlenius), 47, 48
Fie (Polyrrhachis), 1
dochmia (Scoparia), 229
Docirava, 68
Dodona, 82, 93
dodsi (Promethus), 421, 429, 435
Dolichoderus, 1
dolopea (Strepsicrates), 232
dolosus (Darbanus), 340
», (Kuagoras), 340
domestica (Musca), 70
domesticus (Gryllus), 309, 311
donaldsoni (Beralade), 321
Dorecadion, 43, 45, 46, 47, 49
doria (Pachytomella), 53
dorilis (Polyommatus), 41
dorsalis (Promethus), 429, 436
dorycus (Astacops), 348
I
Dorylus, 100
doursii (Lionotus), 413
»» (Odynerus), 413
Drepana, xxiv
dromedarius (Notodonta), iii
drosochalea (Orthenches), 242
Dryadaula, 243, 244
dryas (Clovia), 331
», (Satyrus), xliii
,, (Thalattoscopus), 334, 363
drypetes (Lethe), 129
duaringa (Ogyris zosine-), 280
dubia (Phyllyphanta), 335
», (Polistes), 416
dubius (Sehirus), 50
», (Sphindus), 47, 49
dulcis (‘Teracolus), 107, 142, 144
Dundubia, 330
12-guttata (Halyzia), 50
12-striatus (Anommatus), 367
duplex (Oxeoschistus), 189, 190
durga (Dodona), 82
Dynamenais, 343, 344
Dysdereus, 345, 346
Dytiscus, iv, 51
Farias, 71, 79
eatoni (Lionotus), 412
», (Odynerus), 412
ebenina (Philia), 360
ebenista (Strophosomus), 49
Echidnoglossa, 48
echii (Monanthia), 50
Ectatops, 348, 344
edusa (Colias), vi, lv, 40, 65, 372
», var. helice (Colias), vi, 372
egregius (Lionotus), 410
», (Odynerus), 410
Elaphrus, 43
electra (Colias), lv
elegans (Coryphocera), 129
», (Homoporus), 428, 435
elima (Aphnzus), 85
Elis, 88
elongaria (Acidalia), 389
elongatum (Bembidium), 51
elpis (Lampides), 95, 96
Elymnias, 88, 89, 92, 98, 115
emarginatus (Collyris), Lxxii
embla (Erebia), 13
Emblethis, 46, 50
‘embolina (Lethe), 129
emeritella (Depressaria), Ixvii
-emilize (Saturnia), viii
Emus, 43, 44
encedon (Acriea), 265
Enicmus, 47, 48
Enithares, 339
Enome, 108, 110
( Yexxvi}’)
Epamera, 252
Epeira, 5
Ephyra, 389
epibata (Criocephalus), 158
epicles (Ilerda), 93, 97, 134
Epinephele, xliii, 40, 42, 72,94, 135, 376
epiphanes (Izatha), 238
epiphron (Erebia), 39, 40
Epirodera, 339
Episilia, 382
epius (Spalgis), 126
epotis (Platyptilia), 231
equestris (Amara), 42, 43
(Lygeeus), 46, 50
erate (Colias) 73
Erebia, xviii, xliii, xlix, lii, 9-35, 39,
40, 41, 42
eremitis (Scoparia), 230
Ergates, 51
Ergolis, 89, 92, 97, 101, 109, 115, 119,
125, 127
ericee (Ceuthorrhynchus), 45
Ericeia, 110
ericetaria (Selidosema), 388
erikssoni (Aphneus), 252
Erikssonia, 255, 256, 262
erinaceus (Strophosomus), 49
eriopoda (Amphipyra), 384
eris (Teracolus), lvi
erithonius (Papilio), 71, 74, 78, 88, 84,
90, 100, 107, 109, 118, 116
Eronia, lvii, lxvi
erratica (Megalocerea), 50
erymanthis (Messaras), 127
erythraspis (Pentatoma), 356
(Stenozygum), 356
er ythrocephalus (Neoclytus), xvi
esa (Platypleura), 194
Esolus, 47, 48
esuriens (Eumenes), 85
Etiella, 108
etrida (Teracolus), 71, 80, 108, 104,
105, 106, 107, 117, 119, 134, 139,
141, 142, 144
Euagoras, 340
Eublemma, 385
eucharis (Delias), 79, 88, 84, 86, 87, 90,
91, 101, 118, 123, 133, 137
5, (Teracolus), 108, 119, 142
Eucheira, xix
Euchloé, lxxv, Ixxvi, 372, 393
Euchromia, 129
Eucorysus, 359
eugenes (Dodona), 93
eukrines (Liptena), 253, 262
eumene (Mesosemia), ii
Eumenes, 85, 105, 106, 129, 400
eupheme (Zegris), 1xxv
( cxxvii )
se tee
eupheno (Euchloé), 372
», var, androgyne (Euchloé), 372,
393
euphenoides (Euchloé), lxxv
euphrosyne (Argynnis), 102
Eupithecia, 389
Euplexia, 384
Eupleea, 5, 6, 91, 96, 124
euprepes (Iridopsis), 254, 262
Euprepia, 381
Euproctis, 386
EKupterote, 126
Euptychia, 185, 186, 187, 190
Euricania, 335
Eurinome, 357
Euripus, xliii
Eurranthis, 387
euryale (Erebia), 17, 39, 40
Eurydema, 50, 52
Eurygaster, 50
eurynome (Neptis), 5, 6
Euryphene, 251
Eurythecta, 233
Euterpe, xx
Euthalia, 77, 89, 95
Kuxoa, 69, 104, 382
evanescens (Umjaba), 197
Everes, 133
evias (Erebia), 9-35, 39, 40
», var. hispanica (Erebia), 10, 19,
27, 28, 30, 31, 34, 35
», var, penalare (Erebia), 18, 19, 31,
33, 35
exhausta (Boeturia), 330
55 (Cicada), 330
exigua (Oxypoda), v
exiguus (Coctoteris), 357
exilis (Meligethes), 51
Exochomus, 50
exoticus (Criocephalus), 155, 159, 164
exquisita (Iridopsis), 254
exultans (Bassus), 419
Eysarcocoris, 110
Eysarcoris, 357
faber (Ergates), 51
faceta (Episilia), 382
falcata (Phaneroptera), li
falcataria (Drepana), xxiv, xxv
falkensteinii (Uranothauma), 253
fallax (Cynorrhina), xlvii
famelica (Physopelta), 343
familiaris (Amara), xvi
939 (Cimex), 349
‘; var. hospes (Stalagmoste-
thus), 349
famosus (Dindymus), 344
famula (Fidonia), 387
farinosa (Gonepteryx), Xxxviii, xl
fascialata (Selidosema), 223
fasciata (Macroplax), 50
fasciatus (Heriades), lii
fasciipennis (Megalyra), 396
fausta (Teracolus), 106
favonia (Zygzena), 391
febretta (Amicta), 40
felicina (Metopoceras), 384
felicitanus (Rhizotrogus), 48, 44
femoralis (Phytocoris), 50
femorata, var. aureocincta (Philia), 359
ferruginea (Hammerschmidtia), xlvii
ferus (Criocephalus), iv, 145-176
», (Nabis), 50, 52
»» var. hispanicus
157, 164
festivum (Eurydema), 50, 52
festivus (Homoporus), 435
s, (Promethus), 429, 436
fettigi (Ceenonympha), 376
fibulata (Idea), 108
Fidonia, 387
fidoniata (Semiothisa), 79
fieldii (Colias), 65, 68, 69, 73, 77, 134
fimbriata (Physopelta), 343
fischeri (Celonites), 399
fissorius (Homoporus), 427, 433
flagellatus (Gymnopleurus), 48
flammatra (Agrotis), 81
Flata, 335
flavalis (Pyrausta), 392
flaviceps (Geocoris), 349
a, (Priocnemicoris), 352
flavicornis (Aradus), 50
ay (Paromalus), 47, 48
a (Rhamphus), 49
flavipes (Hydrophilus), 51
», (Hydroporus), 48, 51
», (Oniticellus), 48
», (Trichomastix), 425, 430
flavolineatus (Homoporus), 427, 433
flegyas (Abisara), 94
5, (Zemeros), 95, 97
florale (Omalium), 44
florella (Catopsilia), lv
floricola (Lionotus), 414
5, (Odynerus), 414
fluviata (Plemyria), 132
foederata (Polistes), 416
foetens (Aphodius), 44, 48
fontenayi (Leptura), 43, 45
Forficula, liii, lxi
formicarius (Clerus), 52
a (Myrmeleon), xlvi, lxxvi
formosa (Imogen), 338
», (Peggioga), 336, 363
formosus (Cyclopides), 260
(Criocephalus),
( exxviii )
fornicatus (Hajar), 336
fossor (Aphodius), 44
fowleri (Dacne), liv
fracticornis (Onthophagus), 44
franconica (Malacosoma), 391
fraterculus (Promethus), 436
frigidus (Ammeecius), 43, 44, 48
frontalis (Homoporus), 432
+9 (Ploteia), 132
frugaliata (Semiothisa), 104
fuligineus (Stygnocoris), 46
fuliginosus (Ilybius), 51
fulvipes, var. varipes (Philonthus), 44
fumata (Chittira), 131, 137
fumosa (Beralade), 320
funerarius (Bombus), 94
furcatus (Onthophagus), 44
fusca (Taragama), 316, 326
fuscipes (Aleochara), 44
», (Hydrobius), 51
fuscus (Rhizotrogus), 43, 44
fylla (Abisara), 94
galathea (Melanargia), lii, 39, 40
galba (Hesperia), 106
gamma (Plusia), 385
Ganoris, lvi, lvii, lviii, lix, 67, 70, 71,
G2, Wd; SL, -8¥E9915495597, 1995 1015
111, 184 187
garuda (Euthalia), 77, 89
Gastrimargus, 83
Gastroidea, 46
Gastroplakeis, 314, 326
Gegenes, 73, 74, 76
geminata (Zonabris), 49, 52
geminus (Bidessus), 51
gemmaria (Boarmia), 388
geniculatus (Dicyphus), 50
os (Halorhynchus), 368
geniste (Rhagonycha), 45
genoveva (Ogyris), 270, 271, 278, 280,
286
7 -duaringa (Ogyris), 271
i -magna (Ogyris), 271
genutia (Danais), 73
», (Limnas), 87, 89, 90, 91, 92,
98, 100, 104, 105, 109, 128, 138
Geobia, 358
Geocoris, 46, 349, 363
Geodromicus, 47, 48
geometralis (Lepyrodes), 95
geometrica (Grammodes), 385
Geotomus, 358, 359
Geotrupes, 43, 44, 47, 48
germanica (Panorpa), iv
% (Vespa), 417
germari (Apion), lxv
Gerris, 52, 341
Gerydus, 1-4
geryon (Procris), 41
gesticularia (Thamnonoma), 388
ghilianii (Carabus), 46, 47
5s (Steropus), 46, 48
gibbifer (Gerris), 52
gideon (Xylotrupes), 132
gidica (Belenois), lvii
glabra (Liodes), 48
glabrata (Echidnoglossa), 48
glabricollis (Acylophorus), 51
glaciale (Bembidium), 42
glacialis (Erebia), xviii
», (Helophorus), 44, 48
a var. nicholli (Erebia), 9
glandicolor (Cymus), 50
glauca (Catochrysops), 253
glaucescens (Tarundia), 335
glaucicolella (Coleophora), xlviii
glaucippe (Hebomoia), 116, 126
globicollis (Cryptocephalus), 50
Glottula, 383
Glutophrissa, lix
glycinopa (Dryadaula), 243, 244
gnoma (Catopsilia), lv, 138
Gnophos, 388
Gnorimus, xxxi
goante (Erebia), xlix
goeldii (Pachypodistes), 245, 246
Gonepteryx, xxxv-xli, 64,66,80,372,373
Goniodoma, xlviii
Gonocerus, 50, 351
Gonometa, 313, 326
gordius (Polyommatus), 41, 42
gorge (Erebia), xlix
gorgone (Hrebia), xlix
gorgopis (Tauroscopa), 227
goschkevitschii (Blanaida), 135
gothicus (Lopus), 50
graberi (Taragama), 316
gracilentus (Promethus), 436
graculus (Homoporus), 426, 432
graellsi (Dichotrachelus), 47, 49
», (Doreadion), 49
Grammodes, 385
Grammoptera, xvii
granaria (Oligota), ii
grandidieri (Yanga), 195, 202
grandis (Pterochilus), 404
; (Scutellera), 359
graniceps (Phycochus), xlviii, 367
Graphosoma, 50
Grapta, xxix, xliii, 136
gremius (Suastus), 87, 109
griseata (Dichromodes), 222
»» (GKeophora), 239, 240
eriseostriatus (Deronectes), 47, 48
Gryllotalpa, 1
Gryllus, 309, 311
v
()) @xxix. ')
guadarramus (Carabus), 46, 47
guttatus (Agabus), 48
guttigera (Kysarcocoris), 110
guttularis (Yanga), 196
guynemeri (Stenus), 42, 44
Gymnobathra, 238
Gymnopleurus, 48
Gynandrophthalma, 42, 45
Gyrinus, 51
hzematodes (Corymbites), 42, 45
hemoptera (Chrysomela), 46, 50
hemorrhoidalis (Aphodius), 44
Hajar, 335, 336
halensis, var. fuscitarsis (Hydroporus),
51
halmaturia (Ogyris), 269, 271, 275, 276,
277, 289, 290, 292
Halorhynchus, 368
Haltica, 46
Halyzia, 50
Hammerschmidtia, xlvii
hamza (Adopea), 380
hannibal (Lionotus), 414
ss (Odynerus), 414
hapalina (Catochrysops), 109
Haplopacha, 313, 323, 324
Haptoderus, 43, 46, 48
hardwickii (Parnassius), 69
Harmologa, 234
harmonia (Ueana), 329, 363
haroldi (Mycalesis), 252, 262
Harpactor, 46
Harpalus, 42, 44
harpophorus (Crambus), 235
Hebomoia, 98, 116, 126
hebreus (Polistes), 73
hecabe (Terias), 61, 66, 67, 68, 70, 71,
72, 73, 75, 79, 81, 82, 83, 84, 87, 89,
95, 98, 99, 109, 111, 113, 119, 121,
122, 129, 130, 132, 134, 139, 140,
141, 142, 143, 144
hector (Papilio), 114, 118, 123, 134
helena (Morpho), xix
helenus (Papilio), 7
helice (Colias), vi
Heliconius, li, 117
Heliopates, 45
Heliothis, 41
hellenicus (Pterochilus), 406
hellica (Synchloé), lix
helluo (Carabus), 46, 47
Helochares, 51
Helophilus, lxv
Helophorus, 42, 44, 47, 48
Helops, vi
Hemerophila, lxiii
hemiclista (Dipterina), 233
Henicopus, 43, 45
Henicospilus, 110
Hepialus, 41
Herbula, 69
Heriades, lii, lit
hespera (Nephele), 71, 77
Hesperia, 41, 381
hesperica (Rhagonycha), 45
hespericus (Anisorrhynchus), 45
hesperus (Papilio), xii
Hestia, 133
heteraulus (Crambus), 224
Heterocordylus, 46
Heterocrossa, 235
Heterogaster, 46, 50
Heterogynis, iv, 41, 177-184
Heterosternus, lil
hewitsoni (Ogyris), 269, 270, 272, 285,
289, 291, 292
-meridionalis (Ogyris), 272,
286, 289
heydeni (Agabus), 48
ae (Henicopus), 43, 45
hieracii (Zonabris), 49
Hierodula, 6
hintza (Castalius), 253
Hiposcritia, 93, 94, 95, 111, 112, 141
Hipparchia, 73, 74, 135
hippia (Nepheronia), 88, 90, 107
hippo (Tachyris), 97, 99, 141
hippothoe (Polyommatus), 42
hirtarius (Biston), xxxi
hirtus (Emus), 43, 44
hirundinis (Stenopteryx), Ixvi
Hispa, 50
hispanica (Erebia), 12, 17, 27, 28, 29,
30, 38
3 (Isomira), 45
hispanicum (Doreadion), 49
hispanicus (Bassus), 436
Hister, 44
histrio (Crocisa), 92
hobleyi (Lyczenesthes), 258
holmgreni (Zootrephus), 425, 430, 431
Homapterus, 42, 45
homeyeri (Liptena), 252, 262
= (Spindasis), 255
Homaporus, 422, 423, 424, 425, 426,
432, 433, 484, 435
honestus (Harpalus), 44
Hoplopus, 406, 407, 408
hora (Borkhausenia), 240
horsfieldi (Mimela), 98
hospes (Lygeeus), 349
Hotinus, 126
hova (Rihana), 198
hiibneri (Yphthima), 87, 88, 89, 92,
113, 184, 142
humeralis (Dacne), liv
9
( mes. )
humeralis (Liodes), 48 idmo (Ogyris), 269, 271, 273, 274, 275,
Huphina, 66, 70, 79, 91, 94, 96, 98,] 276, 280, 289, 290, 292
99, 101, 102, 105, 106, 115, 119,|Ilerda, 65, 66, 70, 74, 81, 82, 93, 97,
120, 127, 187, 189, 140, 141, 142,] 134
143, 144 ilicis (Orchestes), 49
hyale (Colias), xliii, 40 », (Thecla), 42
»» var. erate (Colias), 72 var. mauritanica (Thecla), 377
hyalina (Agalope), 65, 68 ilithyia (Biblia), 108
», (Sadaka), 191 », (Hypanis), 108, 119, 142
Hybernia, iii Illaphanus, xlviii, 365
Hyblea, 83 illuminatella (Argyresthia), xlvi
hybridalis (Stenopteryx), 112, 113 Ilybius, 51
Hydrena, 48, 51 immanata (Polyphasia), 68
Hydrelia, 109 Imogen, 337, 338
Hydrobius, 51 imperialis (Cyria), Ixvii
Hydrochus, 51 - (Riptortus), 351
Hydrocyphon, 45 inachis (Kallima), xxxiii, 96
Hydrophilus, 51 inangulata (Ericeia), 110
Hydroporus, 47, 48, 51 incanus (Brachyderes), 49
Hydrotea, vii inceptaria (Bilactis), 113
hygrobius (Homoporus), 429, 435 incisus (Homoporus), 424, 428, 434
hylas (Cephonodes), 109 inclusa (Dasychira), 132
», (Lyczena), 41, 380 incoloralis (Pyrausta), 77
», var. atlantica (Lycena), 380, 393 | incondita (Metachrostis), 129
Hylastes, 49 inconspicua (Pentatoma), 357
Hylotrupes, 51, 172 incredibilis (Iridopsis), 254
Hylurgus, 51 indecora (Bombycopsis), 322
Hymenia, 71, 77, 85 indica (Pyrameis), 65, 69, 70, 80, 82,
Hymenoplia, 44 93, 94, 111, 113
Hymenopus, 57 indigestana (Tortrix), 234
hyoscyami (Therapha), 46, 50 indistans (Mycalesis), 89, 92, 94, 99,
Hypanis, 108, 119, 142 135, 140, 141
Hypena, 67, 386 indra (Hiposcritia), 95
Hypera, 42, 45 ines (Melanargia), 374
hyperanthus (Aphantopus), 40 infusum (Megaloccelum), 50
i (Epinephile), 135 inica (Yphthima), 107, 113, 141
Hypercallia, 235 inostentata (Panagra), 222
hyperion (Aufidus), 332, 363 - (Paragyrtis), 222
Hypocista, xviii insignis (Bassus), 419
hypocrita (Bembidium), 48 », (Notoreas), 221
Hypolimnas, xiii, 62, 70, 71, 76, 79, | instabile (Cyphodema), 50
82, 83, 84, 88, 89, 100, 108, 109, 117, | interjunctalis (Cledeobia), 392
125, 134, 264, 265, 268 interlectum Zonosoma), 96
hyppasia (Trigonodes), 109 iaterrupta ‘Creatonotus), 126
hyppoclus (Symbrenthia), 95, 97, 98, 99 a (Microplax), 50
hyriaria (Synegiodes), 93 interruptus (Homoporus), 433
Hystrichopsylla, xxvii interstitialis (Bothrideres), 48
Iambrix, 126 ione (Teracolus), lvii
ianthis (Ogyris), 270, 271, 282, 289, | iota (Plusia), 41
290, 292 iphioides (Ccoenonympha), 41, 42
iapygia (Melanargia), 42 Iphita, 343
ibalioides (Bassus), 436 iphita (Precis), 70, 81, 82, 98, 99, 113,
ibericum (Bembidium), 46, 48 117, 121, 125, 139, 140
icarus (Lycena), 41, 42, 379 iracundus (Harpactor), 46
»» var. celina (Lyczna), 379 iridis (Lixus), 52
ictis (Aphneeus), 85 Iridopsis, 254, 262
ida (Epinephile), 376 iris (Apatura), xviii
Idea, 99, 108, 130, 132 Isamia, 137
( ‘eres )
ischnocyma (Notoreas), 221
ismene (Melanitis), 89, 92, 95, 109, 114,
135, 136, 140, 141, 142
isocrates (Virachola), 101
isodactyla (Platyptilia), 40
isoleuca (Notoreas), 221
Tsomira, 45
Ixias, 70, 79, 80, 83, $4, 92, 93, 96, 98,
99, 101, 103, 104, 108, 115, 116, 127,
134, 139, 140, 141, 142, 143
jacchus (Philotherma), 318
jacintha (Diadema), 70
Jamides, 98, 131
janira (Epinephele), 94, 135
janthina (Gastroidea), 46
japonicum (Orthetrum), 135
jasius (Charaxes), 373
jasonia (Hestia), 133
javanica (Tessaratoma), 126
jesous (Lampides), 379
Jugurthia, 399
jumba (Neptis), 110
juniperi (Gonocerus), 50
juno (Platylomia), 200
jurtina, var. fortunata (Epinephile),
376
juvenculus (Dorylus), 100
Kallima, xxxiii, 93, 97
kallimon (Spindasis), 254, 262
Karsandra (Zizera), 70, 71, 78, 74, 102,
108, 121, 133
kashmirensis (Vanessa), 65, 66, 69, 80,
82, 93, 94, 134
ne (Zygeena), 72
klugi (Pantodinus), iii,
kollari (Pademma), 91, 137
kydippe (Ptochiomera), 346
Labidostoinis, 45
lachesis (Melanargia), 42
», Var. canigulensis (Melanargia),
li
» var. cataleuca (Melanargia),
41, 42
Lachnea, 49
laciniata (Phyllomorpha), 52
lacordairei (Steropus), 42, 43
lactea (Raparna), 110
ladon, var. pseudargiolus (Cyaniris), 99
Lemophleus, liv
Laemosthenes, 47, 48
Leosopis, 41, 42
leta (Serenthia), 50
», (Terias), 70, 95, 101, 103, 105, 106,
107, 134, 139, 141, 142, 143
letatorius (Bassus), 425, 431
letitia (Euryphene), 251
levigatus (Miris), 50
Lagria, 49
Lahea, 319
laius (Chilades), 79, 101, 108, 119
lalage (Hiposcritia), 93, 94, 141
l-album (Leucania), 383
lamia (Pierella), 185
laminata (Cuspicona), 357
Lampides, 41, 42, 90, 92, 95, 96, 98,
99, 109, 117, 118, 119, 120, 126, 127,
379
lampros (Bembidium), 43
Lampyris, 45
lanka (Cyaniris), 130, 132
Laphygma, 384
laqueatus (Oxytelus), 44
Lareynia, 47, 48
Lasiocampa, 41
lasti (Lyceenesthes), 253, 258
laterale (Bembidium), 46, 48
lateralis (Camptopus), 52
sp (Homoporus), 432
lathonia (Argynnis), 40, 42, 65, 69, 374
» var. isseea (Argynnis), 66, 68,
81
lathriopa (Trachypepla), 237
laticarpus (Promethus), 429, 436
latifrons (Microdon), xlvii
latreilli (Callopistria), 385
latus (Corymbites), 45
Lechriolepis, 318, 326
lefebvrei (Erebia), xlix, 17
var. intermedia (Erebia), xlix
a var, pyrenza (Erebia), xlix
Leistrotrophus, 44
Leistus, 46, 47
Lema, 45
lemolea (Spalgis), 253
lemonias (Precis), 66, 68, 70, 79, 80,
81, 82, 84, 88, 89, 99, 100, 104, 105,
106, 109, 119, 120, 134, 139, 140,
141, 142
lemur (Onthophagus), 44
lentalis (Olybama), 132
lepidus (Hydroporus), 51
leporina (Neottiglossa), 50
Leptidia, 40
Leptocoris, 350
Leptocorisa, 350
Leptoglossus, 352
Leptura, 43, 45, 47, 49
Lepyrodes, 95
Lesteva, 44
Lethe, xliii, 94, 97, 128, 132, 184
Leucania, 383
leucocephalus, var. steganoides (Stron-
gylocoris), 46
leucocera (Celenorrhinus), 95
leucocyanea (Philia), 360
or var. ebenina (Philia), 360
99
(Gun)
leucocyanea (Scutellera), 360
Leucoma, 96
Leucophasia, 79, 88
leucoxesta (Stenoptilia), 232
levana (Araschnia), xxiv, XXV
», (Lymanopoda), 188, 190
levenna (Beralade), 320
Libellula, 120
libyssa (Liptena), 253
Libythea, 95, 125
libythea (Appias), 76, 139
» (Terias), 66, 70, 99, 101, 108,
109, 121
lichenis (Lema), 45
Licinus, 44
lifuana (Cicada), 328
», (Tibicen), 329
,, (Ueana), 328
lifuanus (Oxycarenus), 347
ligea (Erebia), 17
ligniperda (Hylurgus), 51
limacodes (Cochlidion), xxiv, xxv
Limenitis, xliii, 66, 88, 89, 101, 129
Limnas, XXVi, Hex 73, 75, 78, 80, 84,
87, 89, 90, 91, 92, 97, 98, 100, 104,
105, 106, 108, 109, 113, 115, 118,
120, 128, 132, 138, 1385, 187, 138,
263-268
Limnebius, 44
limniace (Tirumala), Hy Gm 87, 89, 90,
91, 100, 109, 117, 118, 128, 124, 138
Limnius, 48
Limnometra, 341
Limnoxenus, 51
limoniella (Goniodoma), xlviii
Limonius, 45
lineata (Deilephila), 386
lineatum (Graphosoma), 50
lineatus (Carabus), 43
lineola (Cimex), 360
5, (Tectocoris), 360, 362
lineolatus (Adelphocoris), 50
lineosa (Synegiodes), 93
linguarius (Pterochilus), 404
Liodes, 48
liodes (Lyceenesthes), 258
Lionotus, 410-415
Liptena, 252, 253, 262
littoralis (Prodenia), 100, 102
se (Thisoicetrus), 81
lituratus (Hydroporus), 51
livida (Leptura), 49
lividalis (Hypena), 386
lixella (Coleophora), xlvii
Lixus, 52
Lochinewa, xxxi
longelytrata (Lesteva), 44
longipes (Homoporus), 423, 426, 432
longiseta (Megalyra), 396
longiventris (Homoporus), 428, 434
Lopus, 50
loreyi (Leucania), 383
loxotis (Borkhausenia), 241
Loxura, 90, 92, 126
lucasi (Melanargia), 374
lucida (Amara), xvi
»» (Tarache), 385
», var. albicollis (Tarache), 385
Luciola, 100
luctuosa (Atheta), 48
~ (Tarache), 385
luctuosis (Macroplax), 348
lugubre (Callidium), 158
lunaris (Copris), 48
», (Ophiusa), 385
lunata (Artaxa), 71
» (otaria), 50
lunicornis (Didineis), lxv
Luperus, 46
luridipennis (Psylliodes), xlvi
lusitanica (Labidostomis), 45
lusitanicum (Malacosoma), 46
lusitanicus (Brachyderes), 49
. (Cryptocephalus), 42, 45
(Heliopates), 45
luteolata (Opisthograptis), 387
luteolus (Hoplopus), 406
», (Odynerus), 406
lutescens (Camptobrochis), 50
Luteva, 341
Lycena, ii, xliii, xlvi, xlix, 1, 39, 41,
42, 81, 82, 111, 379, 380, 393
Lycenesthes, 253, 258, 262
lycaon, var. Iupinus (Epinephile), 376
», var. mauritanicus (Epinephile),
Lycophotia, 383
Lygeus, 46, 50, 52, 343, 348, 349, 350,
352
Lygistopterus, 47, 49, 52
lyllus (Cenonympha), 376
Lymanopoda, 188, 190
Lymantria, 386
Lymexylon, xlvi
lysimon (Lycena), 379
Lytta, 42, 45
macclounii (Charaxes), 251
macheristes (Orocrambus), 224
machaon (Papilio), xliii, 70, 75, 203,
213, 371
a var. asiatica (Papilio), 69
Machimus, iv
macleayanus (Papilio), xviii
macleayi (Illaphanus), 366
macrocephalus (Carabus), 42, 48
Macroglossa, 81, 387
( ‘exxxiil } )
macrophthalma (Orthostira), 46
Macroplax, 50, 348
Macropsis, 334
Macrothylacia, lii, 390, 393
maculata (Kpeira), 5
maculatus (Hotinus), 126
a (Promethus), 436
maculipes (Sehirus), 50
maderensis (Gonepteryx), XXXVil, XXX1X
inera (Pararge), 40, 375
maha (Zizera), 70, 71, 74, 76, 81
», var. diluta (Lycena), 82
.% (Zizera), 66, 73, 74, 76
major (Quartinia), 399
Malachius, iv
Malacosoma, Ixix, 46, 391
malia (Lymanopoda), 188
malvee (Hesperia), 41
», var. malvoides
XXXV
malvata (Cidaria), 389
Mamestra, 383
mandata (Mycalesis), 121, 143
mandibularis (Acrognathus), xxxi
Marasmia, 77
margaritacea (Cassida), 50
margaritosa (Lycophotia), 383
marginalis (Dytiscus), 51
(Syrichthus),
55 (Porthesia), 77
marginatus (Hydroporus), 51
a (Syromastes), 52
marianne (Ixias), 70, 79, 80, 83, 101,
103, 104, 108, 115, 116, 139, 140,
141, 142
maritimus (Beosus), 50
marmoratus (Gastrimargus), 83
marmottani (Cleonus), 52
maro (Ampittia), 119
marshalli (Beralade), 321
x3 (Mimacreea), 252
Ff (Yphthima), 92, 98, 99, 114
martini (Rihana), 198
Maruina, 297
maso (Lymanopoda), 188, 190
mathias (Parnara), 71, 83, 85, 117, 126
maura (Kurygaster), 50
mauritanicus (Thestor), 378
maxima (Gonepteryx), XXXviil, Xxxix
maximus (Phalacrus), 48, 51
meade-waldoi (Ceenonympha), 377
mechanitis (Notoreas), 221
Mecyna, 40, 112, 392
Medon, xxvii
medus (Calysime), 125
meeki (Ogyris), 271, 278, 289
megacephala (Acronycta), 384
megera (Pararge), 65, 67, 185, 136, 375
Megalocerea, 50
Megaloccelum, 50
Megalopus, xvill
Megalymenum, 353
Megalyra, Ix, 395, 396
megamera (Ganoris), 137
Meganotum, 126
Megasemum, 146, 147, 148, 161, 164
megaspiloides (Eysarcoris), 357
megaspis (Homoporus), 428, 435
Megymenum, 353, 354
melena (Castalius), 253
melampus (Rapala), 83, 85, 86
melanamma (Borkhausenia), 240
Melanargia, li, 39, 40, 41, 42, 374
melanaspis (Promethus), 429, 435
melaneus (Caduga), 95
Melanitis, 89, 92, 94, 95, 109, 114,
135, 136, 1388, 140, 141, 142
melanocephala (Cymindis), 42, 44
melanocephalus (Cymus), 50
melanodonta (Titanoptilus), 248, 249
melanoides (Parantica), 95
melanomera (Meganotum), 126
Melanophthalma, xlvi
melanops (Lyczna), ii, iil
Fr var. wheeleri (Lycéena), iii
melanoptera (Megalyra), 395, 397
Melanotus, 45, 49, 51
melas (Erebia), xviii, xlix, 17
melete (Ganoris), 137
Meligethes, 51
melipona (Megalopus), xviii
Melita, xliii, lii, 73, 74, 373
meliturga (Scoparia), 228
Meloé, 45
membranaceus (Aradus), 342
3 (Cimex), 352
(Leptoglossus), 352
var. orientalis (Brachy-
rhynchus), 342
memnon, var. agenor (Papilio), 96
memorata (Psilocambogia), 98
menaka (Tagiades), 114
menapia (Neophasia), xxi, Xxil
menthastri (Chrysomela), 46
meone (Pararge), Xxvlii, xxix
meridionalis (Phytocoris), 50
merione (Ergolis), 90, 97, 101, 125
mesentina (Belenois), vii, Ixvi, 65, 67,
105, (ll, G45 (Osiiie ones 82, 103,
104, 105, 106, 107, 134, 137, 144
Mesosemia, ii
Messaras, 127
Metachrostis, 79, 129
metadelta (Izatha), 238
(Semiocosma), 238
metallica (Scutellera), 362
metallicus (Pedilophorus), 42, 44
re)
( \iexxxiv ))
Metamorpha, 117
Metaporia, xxi
meticulosa (Brotolomia), 384
metirius (Hypocista), xviii
Metopoceras, 384
mexicanus (Criocephalus), 145, 154,
159, 164
Microdon, xlvii
microdora (Diptychophora), 227
Microdynerus, 415
Microplax, 50
Micrositus, 49
Mictis, 352
midamus (Euplcea), 5
B3 (Isamia), 137
millefolii (Anthaxia), 49
Mimacrvea, 252
mimas (Euptychia), 187, 190
Mimela, 98
Mimetica, liii
minutissimus (Bidessus), 51
Miris, 50
misippus (Hypolimnas), xiii, 76, 83, 84,
88, 89, 100, 109, 117, 134,
264, 265, 266, 267, 268
»» var. inaria (Hypolimnas),109,
264, 265, 266
Misolampus, 47, 49
mixtus (Mononyx), 338
mnesichola (Xanthorhoe), 220
mniszechi, var. maroccana (Satyrus),
375, 393
meesta (Corixa), 50
Monanthia, 50
monilis (Lamophleus), liv
Mononyx, 338
montana (Narmada), 128
montanellus (Platyderus), 48
montanus (Criocephalus), 158, 159
is (Cyrtonus), 47, 50
monteironis (Lycznesthes), 253
monticola (Homoporus), 432
montivaga (Cosmopsaltria), 200
montrouzieri (Megymenum), 354
(Zangis), 356
morei ’(Cryptocephalus), 46, 50
morbillosus (Carabus), ]xv
Mordella, 45
Mordellistena, 49
moricei (Panurgus), lii
Mormolyce, |xxii, lxxiii
Morpho, xviii
moschata, var. thoracica (Aromia), 43,
45
multicolor (Bassus), 425, 432
a (Homoporus), 432
Munza, 192, 202
muricata (Nothorrhina), 51, 157
murina (Isomira), 45
murinus (Leistotrophus), 44
Musca, 70
mutabilis (Adonia), 50
mutata (Serica), 48
muticus (Anthophagus), 42, 44
mutilis (Megalyra), 395
Mycalesis, 83, 89, 92, 94, 98, 99, 100,
114, 115, 121, 125, 135, 138, 140,
141, 142, 148, 251, 252, 262
Myelois, 392
Mylabris, 70, 84
Mylothris, liv, lviii, lix
Myodocha, 350
myrmecophila (Tasmanica), 368
Myrmeleon, xlvi, lxxvi
myrrhina (Dryadaula), 243
mystacatus (Cryptocephalus), 46
Nabis, 46, 50, 52
Nacaduba, 71, 133
nadina (Huphina), 94, 98, 141
Nmatopus, 352
Nais, 293
napa (Dryadaula), 244
napi (Ganoris), 71, 91, 101, 117, 187
», (Pieris), Ixxv, 40, 66
nareda (Yphthima), 78
narendra (Hiposcritia), 111, 112
Narmada, 117, 128
nasuta (Truxalis), 71, 73, 1384
natalensis (Pyena, 197, 202
navale (Lymexylon), xlvi
Nebria, 42, 43, 48
nebrioides (Deltomerus), 42, 43
neglectus (Rhizotrogus), 51
bs (Zabrus, 42, 43
Neides, 52
Nemeophila, 41
nemoralis (Bassus), 431
5 (Haptoderus), 46, 48
Neoclytus, xvi
Neope, xliii
Neophasia, xx
Neopithecops, 90, 92
neoridas (Erebia), 28
Neottiglossa, 50
Nephele, 71, 77
Nepheronia, 88, 90, 107, 117, 127
Neptis, xliii, 2, 5, 6, 8, 81, 82, 94, 95,
96, 97, 98, 99, 107, 109, 110, 11],
117, 121, 125, 127, 129, 130, 132
nereis (Pachymerus), 347, 363
neriene (Zeritis), 252
nerine (Erebia), 17
nerissa (Huphina), 66, 70, 79, 84, 91,
96, 97, 98, 99, 101, 102, 105, 106,
115, 119, 120, 127, 137, 139, 140,
141, 142, 143, 144
Saag
( \exuxvy )
Neuroctenus, 342
neuwaldeggiana (Mordellistena), 49
newarra (Rhodia), 93
nicholli (Erebia), 9
nietneri (Cethosia), 125
niger (Homoporus), 422, 427, 434
nigra (Humenes), 400
nigricornis (Homoporus, 433
nigrina (Oxypoda), v
nigripes (Catacanthus), 354
i! (Cimex), 354
Bie (Limonius), 45
nigrita (Geodromicus), 47, 48
»» (Hydreena), 51
5, (Pterostichus), 43
» (Silpha), 44
nigritarsus (Homoporus), 435
nigriventre (Philorhinum), 44
, (Promethus), 429, 435
nigrofasciatus (Luperus), 46
nigrosuturatus ( (Cleonus), 45
nigrovenosa (Tr oe 127
nilgiriensis (Badacara), 112
Br (Colias), 112, 138
niobe (Argynnis), 42
nipalensis (Ganoris), 72
mi (GonepteryXx), XXXVI, XXXViii,
XXX1X
Niphanda, xliii
niphe (Argynnis), 73, 74, 83, 97, 112,
118, 128, 132, 134
niphonica (Cidaria), 67
nitidulum (Bembidium), 43
nitidus (Limnebius), 44
nivalis (Baris), 42, 45
,, (Synuchus), 42, 43
nobilis (Gnorimus), xxxi
», (Cidemera), 45
noctiluca (Lampyris), 45
Bi (Pipiza), 438
noctuella (Nomophora), 112
nomentis (Scoparia), 231
Nomophora, 112
noreia (Nacaduba), 71, 133
nostrodamus (Gegenes), 74, 76
(Parnara), 381
notatus (Pissodes), 52
Noteraula, 232
Nothorrhina, Gp Gy aly
Notiophilus, 42, 43
Notodonta, iii
Notoreas, 220, 221, 222
nubilus (Criocephalus), 154, 158, 159
numata (Heliconius), li
numida (Jugurthia), 399
nupta (Catocala), xlv
Nychitona, 79, 87, 89, 92, 97, 101, 110,
115, 120, 127
nycteris (Rhopalopsyche), 69
Nymphula, 392
nyseus (Talicada), 110, 111, 126, 129,
1380, 131
Nysius, 46, 50
Nyssanga, 125, 1382
oberthuri (Alena), 252
a (Catocala), 384
(Cephalallus), 148, 163
obliquata (Beralade), 320
oblongata (Eupithecia), 389
oblongus (Limnoxenus), 51
obscura (Ocinaropsis), 317
obseurus (Aphodius), 43, 44, 48
obsitalis (Hypena), 386
obsoletus (Criocephalus), 145, 154,
158, 159, 164
obtecta (Cosmopsaltria), 330
», (Dundubia), 330
», (Tettigonia), 330
obtusella (Coleophora), xlviii
occulta (Xanthorhoe), 220
ocellaris (Dinoderus), xxvii
ocellea (Eromene), 106
ochrata (Acidalia), li
ochrearia (Aspilates), 387
Ocinaropsis, 317
Ocnerostoma, xlvi
18-guttata (Halyzia), 50
octomaculatum (Bembidium), 51
ocularis (Sthenarus), 50
Odontopera, lxvi
Odynerus, 406-415
(Ecophora, 239
(Ecophylla, 279
(Kdemera, 45, 49
cenone (Precis), 70, 74, 81, 82, 84, 100,
102, 104, 106, 109, 120, 139, 140,
141, 142
cenospora (Crocydopora), 224
ie (Sporophyla), 224
Ogyris, xxvi, 269-292
olane (Ogyris), 272, 283, 289, 291, 292
oleracea (Ganoris), 137
+3 (Mamestra), 383
oleraceum (Eurydema), 50
Olibrus, 48
Oligochroa, 71, 79
Oligota, ii
olivacea (Phytodecta), 46
Olybama, 132
Olyra, 3238, 326
Omalium, 44
omichlias (Notoreas), 222
Omophlus, xxxi
omphale (‘Teracolus), lvi, 1xi, lxii
Oncocephalus, 340
Oncopeltus, 349
( exxxvi )
Oncotylus, 50
Oncotympana, 201
Oniticellus, 48
onopordi (Hesperia), 381
Onthophagus, 44, 48
opalina (Athyma), 66, 68
Ophiusa, 384, 385
Opisthograptis, 387
orana (Procris), 391
orbitulus (Lycena), xlix, 1
5 var. oberthuri (Lyczna), xlix, 1
Orchestes, 49
Oreopsyche, 391
Orgyia, 41, 181
orientalis (Sphinx), 79
Orina, 46
orithyia (Precis), 65, 66, 69, 70, 72, 73,
74, 76, 81, 82, 84, 100, 102, 103, 104,
105, 119, 1838, 139, 140, 141, 142
ornatus (Bassus), 437
;, | (Homoporus), 427, 433
Orneodes, 247
Ornithoptera, Ixvii, 121, 123, 128
Orocrambus, 224
oreetes (Ogyris), 270, 272, 285, 286,
289, 291, 292
orontas (Ogyris), 269, 274, 275
Orsotriana, 99
Orthenches, 242
Orthetrum, 110, 135
Orthomus, 43
orthophanes (Compsistis), 243
Orthostira, 46
osiris (Chcerocampa), Xxili
ossium, var. insularis (Stenus), xlvi
ostrina (Eublemma), 385
otanes (Ogyris), 271, 275, 277, 289,
290, 292
Otiorrhynchus, 42, 45, 46, 49
otis (Zizera), 99, 102, 127
»» var. indica (Zizera), 71, 109, 119
Oxeoschistus, 189, 190
Oxycarenus, 347
Oxypoda, v
Oxytelus, 44
Pachygastria, 390
Pachymerus, 346, 347, 363
Pachypodistes, 245
Pachytelia, 40
Pachytomella, 46, 53
Pachytychius, 49
Pademma, 91, 137
Peelia, 247
palarica (Erebia), 9-35, 39, 40
palealis (Phlyctenodes), 392
palliatus (Hylastes), 49
pallidipes (Homoporus), 428, 435
pallipes (Trichomastix), 480
palustris (Notiophilus), 43
pammon (Papilio), 75, 77, 88, 90, 99,
100, 109, 114, 118, 120, 123,
127, 134
», var. polytes (Papilio), 82
pamphilus (Ceenonympha), 40, 42, 72,
376
pampolis (Catochrysops), 258, 262
Panagra, 222
pandava, var. bengalia (Catochrysops),
88
pandellei (Trechus), 47, 48
pandora (Argynnis), 374
pandurus (Lygzeus), 46, 50
panope (Papilio), 5
Panorpa, iv
Pantodinus, liii
Panurgus, lii, liii
paphia, var. anargyra (Argynnis), 41,
42
Papilio, xii, xviii, xliii, Ixvii, 5, 7, 8,
69, 71, 74, 75, 77,.78, 80, 82, 83,
84, 88, 89, 90, 96, 99, 100, 103, 105,
106, 107, 108, 109, 110, 111, 113,
114, 116, 117, 118, 120, 123, 124,
127, 130, 133, 134, 208, 213, 216,
217, 251, 371
Paracymus, 51
paradelpha (Notoreas), 221
paradoxa (Heterogynis), 177, 178, 179,
180, 181, 183, 184
Paragyrtis, 222
paralekta (Kallima), xxxiil
Parantica, 95, 121, 123, 124,'133, 137
Pararge, xxviii, xliii, 40, 65, 66, 67,
135, 136, 138, 375
pancratii (Glottula), 383
parietum (Ancistrocerus), 410
3 (Odynerus), 410
parinda (Papilio), 110, 124
parisatis (Hipparchia), 73, 74, 135
», (Satyrus), 72
parmata (Cercotmetus), 342
», (Ranatra), 342
parmenio (Erebia), 13
Parnara, 71, 74, 83, 85, 117, 126, 380,
381
| Parnassius, 40, 69
| Paromalus, 47, 48
parumpunctatus (Corizus), 50
parvula (Lagria), 49
parvulus (Lionotus), 413
cs (Odynerus), 413
es (Tachys), 48
pasiphaé (Epinephile), 376
patagiata (Pseudometa), 314, 326
patnia (Nyssanga), 125
pauli (Cleophana), 383, 393
(. | \@xeexvat '.)
paulina (Catophaga), 111, 115, 116,
123, 127, 128, 130, 134, 135, 137,
142, 143
paulinoi (Corymbites), 45
paullula (Sterrha), 110
pavana (Chrysophanus), 65, 66, 68, 81,
82
pavonia (Saturnia), vii, viii, ix, x, xi
pectoratorius (Bassus), 438
(Homoporus), 426, 432
peculiaris (Catochrysops), 253
Pedaliodes, 189, 190
Pedilophorus, 42, 44, 47, 48
pedunculatus (Microdynerus), 415
(Odynerus), 415
Pegsioga, 336, 363
peletierari ia (Cleogene), 39
pellitus (Balaninus), 49
Pelochares, 51
peltigera (Chloridea), 382
39 (Heliothis), 41
pefalare (Hrebia), 17, 30
pendularia (Zonosoma), xxxii
penella (Heterogynis), 177, 178, 179,
180, 181
penicillata (Euptychia), 186, 187, 190
Peniscomus, 351
pennigeraria (Eurranthis), 387
58 var. chrysitaria (Kurran-
this), 387
pentadactyla (Argyria), 227
Pentatoma, 354, 355, 356, 357
penthalea (Borkhausenia), 239
- (cophora), 239
Pentila, 252
Perena, 102
Pericoma, 296, 297, 311
Perileptus, 43
peripheea (Xanthorhoe), 220
perius (Athyma), 73, 81
perobliqua (Beralade), 319
peronatus (Bassus), 438
perplexus (Pilophorus), 50
perpusillaria (Acidalia), 389
perseus (Calysisme), 125
», (Mycalesis), 83, 89, 100, 114,
115, 125, 140, 142
petiolata (Kumenes), 129
pheoptila (Trachypepla), 236
Phalacrus, 48, 51
phalenoides (Psychoda), 311
phalanta (Atella),
89, 100, 103, 104, 115, 129
Phalera, xxiv, 387
phalerias (Scoparia), 230
Phaneroptera, li
pharmactis (Borkhausenia), 241
phasianipennella (Gracilaria), 435
65,68, 70, 74, 76,
pheretes (Lycsena), xlix
Philereme, 67, 68
philerga (Soparia), 228
Philia, 359, 360, 362
philodice (Colias), Xxix
philomela (Yphthima), 82, 89, 92, 95,
113, 114, 140
Philonthus, 44, 51
Philorhinum, 44
Philotherma, 318, 326
Philudoria, 315
Philus, 147
phineus (Euptychia), 187
phleas (Chrysophanns), 66, 378
», (Polyommatus), 41, 42
», var, timeus(Chrysophanus), 65,
66
Phlyctzenodes, 392
Phthorimus, 420, 425, 480, 436
Phycochus, xlviii, 367, 368
Phylacodes, 241, 242
Phyllobius, 42, 45
Phyllodecta, 46
Phyllomorpha, 52
Phyllyphanta, 334, 335
Phymata, 50
Physoderus, 339
Physopelta, 343
Phytocoris, 47, 50
Phytodecta, 46
piceum (Agathidium), 48
picitans (Homoporus), 435
picta (Peecilocera), 72, 73
picteti (Humenes), 400
pictus (Homoporous), 434
Pierella, 185
Pieris, xxxii, xlii, Ixxiv, lxxv, Ixxvi,
40, 101, 203, 371, 372
Piezostethus, 50
pigea (Pinacopteryx), xv, lviii, Ixvi
pilata (Kuptychia), 186
pilipes (Hydrota), vii
Pilophorus, 50
pilula (Byrrhus), 44
Pinacopteryx, xv, lviii, lxvi
pinastri (Syrphus), 432
pinetorum (Criocephalus),
155, 158, 164
pini (Aphanus), 46, 50
piniariella (Ocnerostoma), xlvi
pinicola (Lamosthenes), 47, 48
pipize (Bassus), 438
pisanus (Dytiscus), 51
Pissodes, 52
placida (Cupha), 125, 127
plagiata (Anaitis), 388
plagifera (Trypheromera), 96, 98
Planema, 251
151, 153,
( exxxviii )
planicollis (Licinus), 44
plantaginis (Nemeophila), 41
planus (Homoporus), 434
Plataspis, 362
Platycephala, 362
Platycerus, 43, 44
Platyderus, 48
Platylomia, 200, 201
Platynus, 43
Platypleura, 192, 193, 194
Platyptilia, 40, 231
Platystethus, 293
Plautia, 355
Plegaderus, 47, 48
Plemyria, 132
plinthina (Chloroclystis), 219
plistonax (Euryphene), 251
Plociomera, 346
Ploteia, 132
Plusia, 41, 385
Plusiotis, liii
Pochazina, 335
podagrariz ((idemera), 45, 49
podalirius (Papilio), 71, 371
s var. feisthamelii (Papilio),
371
Peecilocera, 72, 73
Peeciloscytus, 50
Pecilus, 48
pecilus (Dysdercus), 345, 346
», (Pyrrhocoris), 346
poggei (Planema), 251
», (Pseudacrea), xxvi, 251, 263-
268
», (Uranothauma), 253
Polistes, 73, 416
polita (Trichomastix), 430
politis (Borkhausenia), 240
polonicus (Criocephalus), iv, 158
polychloros (Vanessa), 40
Polydidus, 339
polydorus (Platypleura), 193
Polydrosus, 49
polygonalis (Mecyna), 40, 112, 392
polymena (Euchromia), 129
polymnestor (Papilio), 110, 118, 124
polymnia (Ueana), 329, 363
Polyommatus, 41, 42, 65, 71, 72, 74,
78, 85, 105, 106, 108, 109, 131,
134
Polyphasia, 68
Polyrrhachis, 1
polytes (Papilio), 7, 8, 75, 78, 89
polyxena (Thais), 203
Pomasia, 129
pomona (Catopsilia), 75, 84, 88, 89,
110, 115, 121, 122, 133, 138, 139,
140, 142, 143
pomona, var. catilla (Catopsilia), 107,
P21 ASS
pomone (Apion), 49
Pompilus, 81
pontebze (Lionotus), 413
ss (Odynerus), 413
Pontia, xxi
populi (Smerinthus), 886
95 var. austauti ab.
(Smerinthus), 386
porcatum (Steganomma), 365
porcatus (Brachytemnus), 49
porculus (Helophorus), 42, 44
porphyrella (Acrobasis), 392
porphyritis (Orthencha), 243
Porthesia, 77
prelata (Scirpophaga), 391
prestans (Liptena), 253
Pratapa, 85, 134
Precis, 65, 66, 68, 69, 70, 72, 73, 74,
76, 78, 79, 80, 81, 82, 84, 88, 89, 98,
99, 100, 102, 103, 104, 105, 106, 109,
118, 117, ‘119, ° 120,191, ‘135,183;
134, 138, 139, 140, 141, 142, 143,
144, 251
princeps (Venusia), 220
Priocnemicoris, 352
Prioneris, 97, 98, 141
Prionocyphon, liv
prochlora (Ascerodes), 234
Procris, 41, 391
procris (Limenitis), 88, 89, 101
Prodenia, 100, 102
producta (Phyllyphanta), 335
productata (Selidosema), 223
productus (Criocephalus), 158, 159,
160
incarnata
profanus (Lygieus), 352
», (Mictis), 352
on (Neematopus), 352
profunda (Proteodes), 236
prolixus (Actobius), 44
Promethus, 421, 425, 429, 435, 436
prompta (Beralade), 320
pronuba (Agrotis), 382
pronubana (Tortrix), Lxiii
Proselena, 234
Proteodes, 236
proterpia (Pyrisitia), xxxvili
Prothelymna, 234
protractus (Teracolus), 75, 139
Pseudacriea, xxvi, 251, 263-268
Pseudometa, 314, 326
Pseudoterpna, 389
Psilocambogia, 98
Psiloglossa, 400
psittaceus (Harpalus), 44
Psyche, 40, 181
( (exaxix +)
Psychoda, xxx, 293-310
psylaria (Pomasia), 129
Psylliodes, xlvi
Pternistria, 353
Pterochilus, 403, 404, 405
Pterostichus, 43
Ptochiomera, 346
Ptychoptera, 298, 303
pubescens (Lachneea), 49
He (Staphylinus), 43, 44
pudica (Cymbalophora), 381
puellaris (Teracolus), 75, 79, 105, 139,
141
puera (Hyblea), 83
puerta (Oxeoschistus), 190
pulchella (Deiopeia), 72, 81, 84, 109, 110
», (Utetheisa), 382
pulchellus (Promethus), 429, 436
pulcher (Homoporus), 428, 435
pulchra (Lechriolepis), 318, 326
», (Psiloglossa), 401
pulverulenta (Cicada), 199
pumilus (Homoporus), 428, 434
punctatus (Homoporus), 433
punctiventris (Homoporus), 426, 432
punctulata (Tephrosia), xxvii
punctulatus (Dytiscus), 51
ae (Homopterus), 42, 45
punctum (Catacanthus), 354
», (Cimex), 354
pungata (Selidosema), 223
pungens (Rhyncocoris), 356
», (Vitellus), 356
pupillaria (Ephyra), 389
purpuripennis (Carpocoris), 46, 50
pusillus (Brachyplatys), 362
3 _(Ptinus); lxv
puspa (Cyaniris), 95
putli (Chilades), 88, 85, 103
Pycna, 191, 197, 202
pygialis (Rhizotrogus), 48
pygmeeus (Cydnus), 359
+ (Geotomus), 359
5 (Olibrus), 48
Pyrameis, xi, xxviii, 65, 66, 69, 70, 72,
80, 82, 93, 94, 111, 113, 134, 373
pyranthe (Catopsilia), lv, 7,71, 75, 79,
81, 85, 88, 89, 92, 99, 101,
105, 109, 115, 118, 119,
133, 134, 138, 139, 140,
141, 142, 148, 144
», var. gnoma (Catopsilia), 70,
81, 85, 105
Pyrausta, 77, 84, 392
pyreneum (Bembidium), 42, 48, 48
pyreneus (Anthophagus), 44
5, (Barynotus), 45
aa (Corymbites), 45
pyrenzeus, (Geotrupes), 43, 44
FF (Otiorrhynchus), 45
pyrenaica (Lyciena), xlix, |
pyrene (Ixias), 79, 80, 84, 92, 93, 96,
98, 99, 115, 1384, 189, 140,
141, 142, 143
»» var. cingalensis (Ixias), 115, 127
»> var. evippe (Ixias), 93
pyri (Saturnia), vii, viii
pyriformis (Cneorrhinus), 46, 49
pyrina (Zeuzera), 391
Pyrisitia, xxxviii
pyrochroa (Dindymus), 345
An (Dysdercus}, 345
Pyrrhocoris, 346
pyrsophanes (Diptychophora), 228
quadrata (Verlusia), 50
4-costatum (Megasemum), 164
14-pustulata (Coccinella), 50
quadridens (Bolboceras), 104
4-punctata (Anthaxia), 51
quadripunctata (Laphygma), 384
53 (Phaneroptera), li
4-punctata (Zonabris), 49
4-pustulatus (Hxochomus), 50
Quartinia, 399, 400
quercus (Bombyx), 435
», (Lasiocampa), 41
», (Orchestes), 49
», (Thecela), 42
Quiroguesia, 73
quisquiliarius (Philonthus), 51
5-punctatus (Cryptocephalus), 46, 49
Rahinda, 82, 125
rama (Arhopala), 97
5, (Libythea), 95, 125
Ranatra, 342
ranga (Athyma), 95
ransonnettii (Caprona), 117, 129
rape (Ganoris), lvi, lix, 87, 91, 101,
137
», (Pieris), xxxii, lxxiv, 40, 67, 105,
203, 216;1217,)/37a
Rapala, 83, 85, 86, 90, 96
Raparna, 110
Raphidoglossa, 402
rectilineata (Olyra), 323, 326
recurvalis (Hymenia), 71, 77, 85
reflexus (Homoporus), 4238, 428, 434
regensteinensis (Sitones), 45
regia (Gonometa), 313, 326
regina (Teracolus), 251
regulus (Lionotus), 412
», (Odynerus), 412
remba (Huphina), 94
remotata (Idea), 99
repanda (Taragama), 390
revoili (Munza), 192, 202
( wexl> 2) i
reyi (Gynandrophthalma), 42, 45 rufomarginatus (Leptocoris), 350
reynose (Athous), 42, 45 rufonotatus (Homoporus), 435
Rhagium, 161, 1638, 164, 167, 168, | rufus (Telephorus), 45
169 rugosus (Enicmus), 47, 48
Rhagonycha, 42, 45 », (Nabis), 46, 52
rhamni (Gonepteryx), Xxxvi, xxxvii, | rumicis (Acronycta), 384
xxix, xl li laxver ss 2 rumina (Thais), 371
oa. Wale nipalensis (Gonepteryx), var. canteneri (Thais), 371
64, 66, 80 runeka (Mycalesis), 98, 99
Rhamphus, 49 s, (Orsotriana), 99
Rhaphygaster, 356 riippellii (Mylothris), lviii
rhea (Pierella), 185 rustica (Adimonia), 432
Rhizotrogus, 43, 44, 48, 51 rusticus (Criocephalus), 145, 148, 150,
Rhodaria, 41 151, 152, 153, 154, 155, 156, 157,
Rhodia, 93 158, 159, 162, 168, 164, 172
Rhodostrophia, 389 »» (Machimus), iv
Rhopalocampta, 7 », var. ferus (Criocephalus), 158
Rhopalopsyche, 69 rutilus (Agonoscelis), 355
Rhynchites, 45, 52 », (Cimex))'355
Rhyncocoris, 356 saba (Glutophrissa), lix
Ricania, 335 sabina (Orthetrum), 110
ridleyanus (Papilio), 251 sacraria (Sterrha), 388
Rihana, 198 Sadaka, 191
riparia (Hydrena), 51 sahlbergi (Corixa), 52
Riptortus, 350, 351 Salda, 50, 345
Rivula, 132 salsala (Iambrix), 126
roboris (Leosopis), 41, 42 sandycias (Chloroclystis), 219
robusta (EKurythecta), 233 sanguinalis (Rhodaria), 41
robustus (Riptortus), 351 sanguineus (Lygistopterus), 47, 49,
rodriguezi (Heterosternus), liii 52
rohria (Lethe), 94, 97 Saprinus, 44
rosa (Crenis), 251 sarcoxantha (Gymnobathra), 239
5, (Philotherma), 318 sarpedon (Papilio), 7, 8
rosimon (Castalius), 117, 120, 133 Saturnia, vii
rostrata (Ailia), 50 Satyrus, xviii, xliii, lii, 26, 41, 42, 67,
ruber (Capsus), 50 72, 1385, 374, 375, 393
rubi (Callophrys), 378 saucia (Agrotis), li
», (Macrothylacia), li, 390 saucius (Plagaderus), 48
», digramma (Macrothylacia), 390, | saussureii (Hierodula), 6
393 + (Lionotus), 410
rubida (Lagria), 49 Fe (Odynerus), 410
rubra (Leptura), 49 saxatilis (Lygeus), 52
rubripes (Harpalus), 44 scabricollis (Misolampus), 47, 49
rufescens (Aphodius), 44 *n (Pachytychius), 49
oy (Gastroplakeis), 314, 326 scabriculus (Homoporus), 432
eS (Raphidoglossa), 402 scabrosus (Homoporus), 434
ruficornis (Grammoptera), xvii sceevoides (Chamesyrphus), xlvii
ss (Homoporus), 428, 434 schakra (Pararge), xxvill, 65, 67
rufifrons (Daene), liv schaufussi (Saturnia), viii, x
rfipes (Aphodius), 44 Scheenobius, 110
», (Bassus), 419 schreberi (Cerocoma), 47, 49, 52
5, (Stenobothrus), xlvi »» (Onthophagus), 48
», (Tropicoris), 50 schrenkii (Pararge), xliii
rufiventris (Bassus), 420 sciathis (Mycalesis), 251
be (Megalyra), 395 Scirpophaga, 391
a (Zootrephus), 425, 430, 431 | Scolopendra, 293
rufozenea (Chrysomela), 50 Scoparia, 228-231
rufofemorata (Chrysomela), 42, 46 scopulata (Euptychia), 186, 190
a
(
scrutator (Aphodius), 48, 44, 48
scutellaris (Capsus), 4
re (Promethus), 421, 429,
435
Scutellera, 359, 860, 361, 362
segetis (Euxoa), 382
Sehirus, 50
seidlitzi (Zabrus), 46, 48
selene (Argynnis), 40
Selenia, xxiv, xxix
selenophora (Athyma), 95
Selidosema, 223, 388
selousi (Mycalesis), 251
semele (Satyrus), xviii, 42, 67, 135,
374
semicanaria (Thamnonoma), 388
Semiothisa, 79, 104, 110
semistrista (Corixa), 52
semivittatum (Apion), lxv
Sena, 319
sena (Ilerda), 65, 66, 70, 74, 81, 82
senator (Philia), 362
senecionis (Nysius), 46
separata (Clovia), 330
Sepedon, lxxiii
septentrionis (Danais), 123, 131
4; (Tirumala), 127, 134,
135
seraphina (Platypleura), 193
Serenthia, 50
Serica, 48
sericea (Oxypoda), v
sericeus (Cryptocephalus), 46
Serinetha, 350
serratule (Hesperia), 381
serricornis (Prionocyphon), liv
Sesia, 109
setifrons (Polydrosus), 49
severina (Belenois), liv, lvii, lix,
Ixvi
severini (Platypleura), 194
sexdentatus (Tomicus), 51
sexguttatus (Calocoris), 46
sexmaculata (Chrysocoris), 359
Fi (Scutellera), 359
sexmaculatus (Cryptocephalus), 50
(Eucorysus), 359
sexpunctata (Lachnea), 49
3 (Psychoda), xxx, 293-311
sexpunctatus (Anchomenus), 43
a (Platynus), 43
shakra (Pararge), 136, 138
shuckardii (Megalyra), 395
sibylla (Limenitis), 66
sicanaria (Rhodostrophia), 389
sichelii (KEumenes), 400
side (Dysdereus), 345
», (Mylabris), 70, 84
PROC. ENT. SOC. LOND., v. 1905.
exli }
sideraspis (Scoparia), 231
sideritis (Noteraula), 232
siderodeta (Borkhausenia), 240, 241
signatus (Homoporus), 429, 435
Silpha, 44
silvana (Heliconius), li
simillimus (Bombus), 72
simplex (Beralade), 321, 322, 326
», (Chilena), 322
Simulium, 309, 311
sinapis (Leptidia), 40
», (Leucophasia), 79, 88
singalensis (Cyaniris), 67, 68, 69
sinuata (Rahinda), 82, 125
siterata (Cidaria), 389
Sitones, 45
sjostedti (Taragama), 316
skotios (Catochrysops), 259, 262
Smerinthus, 386
sobrina (Nebria), 42, 43
,, (Zonabris), 45
socialis (Eucheira), xix, xxi, xxii
solieri (Euplexia), 384
Solpuga, 108
solstitialis (Rhizotrogus), 44
sophistes (Crambus), 225
sordida (Abraxas), 130
», (Philotherma), 318, 326
sorhageni (Zeritis), 252
Spalgis, 126, 253
speciosus (Teracolus), lxvi
spectrum (Apopestes), 385
specularia (Corymica), 130
specularis (Pippaipe), 411
Hs (Odynerus), 411
sphierula (Coptosoma), 362
Sphindus, 47, 49
Sphinx, 79
spicata (Platypleura), 192
Spilostethus, 348
spilothrys (Celcenorrhinus), 129
spinarum (Athalia), 437
Spindasis, 94, 119, 254, 255, 262
spini (Saturnia), vill, x
spinifer (Platycerus), 48, 44
spinifera (Euxoa), 104, 382
spiniger (Centrocoris), 50, 52
spiricornis (Hoplopus), 406
(Odynerus), 406
splendens (Philonthus), 44
splendida (Euricania), 335
>> (Blata),, 3385
5 (Rieania), 335
- (Scutellera), 362
splendidum (Stilbum), 106
Spondylis, 47, 49
Sporophyla, 224
( exlu »)
squalidus (Lygzus), 349
stahli (Ceratopterus), xviii
Stalagmostethus, 348, 349
Staphylinus, 43, 44
Staria, 50
statilinus (Satyrus), 42
Steganomma, 365, 366
stellatarum (Macroglossa), 387
Stenobothrus, xvi, xvii
Stenopteryx, Ixvi, 112, 113
Stenozygum, 356
Stenus, xlvi, 42, 44
stephensi (Illaphanus), xlviii, 365,
367
Steropus, 42, 43, 46, 48
Sterrha, 110, 388
Sthenarus, 50
stigmatica (Euptychia), 186, 190
Stilbum, 106
stockerus (Chrysocoris), 126
stolida (Grammodes), 385
stomoides (Bembidium), 43
stormsi (Catochrysops), 253
strabo (Catochrysops), 83, 85, 90, 108,
109
Strachia, 355
stragulata (Leptura), 47, 49
straminea (Noteraula), 232
Strangalia, 43, 45
stratiotata (Nymphula), 392
Strepsicrates, 232
striatum (Asemum), iv, 145-176
striatus (Helops), vi
strigator (Bassus), 438
af (Homoporus), 428, 434
Strongylocoris, 46
strophea (Argyria), 225
Strophosomus, 42, 45, 49
stygne (Erebia), lii, 9-35, 39, 40
»» var. bejarensis (Erebia), 11, 15,
16, 30, 31, 34
», var. hispanica (Erebia), 13, 15,
17, 18, 31, 34, 35
var. pefialare (Erebia), 15, 18,
27, 31, 33, 34, 42
», var. pyrenaica (Erebia), 16
Stygnocoris, 46
Suastus, 87, 109
suavis (Philonthus), 51
subalbitaria (Semiothisa), 110
subapterus (Coranus), 50
submarginalis (Scoparia), 231
submarginata (Leucoma), 96
subsericeata (Acidalia), 389
sulcator (Promethus), 429, 435
5, (Zootrephus), 430
sulcatum (Rhaphygaster), 356
sulcipennis (Phycochus), xlviii, 368
sulphureus (Cteniopus), 45
sundevalli (Homoporus), 427, 484
sttperbus (Lygzeus), 50
suspiciosus (Homoporus), 434
suturalis (Brachyderes), 52
~ (Cordylomera), xlv
»» (Lochmea), xxxi
swaha (Aulocera), 64, 68, 69, 135
sylvatica (Cicindela), 43
Symbrenthia, 95, 97, 98, 99
Symphedra, 107
Synchloé, lix
synclinalis (Notoreas), 220
Synegiodes, 93
Synuchus, 42, 43
syriacus (Cephalocrius), 149, 164
», (Criocephalus), 145, 146, 149,
150
Syrichthus, xxxv
Syromastes, 52
szepligetii (Megalyra), 396
Tachypus, 48
Tachyris, 97, 99, 112, 117, 141
Tachys, 48
Teniocampa, xvii
tagalicus (Arictus), 342
», (Brachyrhynchas), 342
Tagiades, 114
Talicada, 110,
131
talpz (Hystrichopsylla), xxvii
tanaceti (Cucullia), 383
Taphropeltus, 50, 52
Tarache, 385
Taragama, 315, 316, 326, 390
tarsatorius (Homoporus), 427, 433
Tarucus, 71, 73, 76, 79, 81, 83, 103,
104, 106
Tarundia, 335
Tasmanica, 368
taurica (Gonepteryx), XXXvil, xXxxviii,
Xxxix, xli
Tauroscopa, 227
Tectocoris, 360, 362
Telchinia, 103, 109, 115, 119, 120,
133
Telephorus, 42, 45
telephus (Papilio), 111, 116, 124,
127
VL, 26) 129. S05
telicanus (Lampides), 41, 42, 379
», (Tarucus), 76, 81, 83, 108,
106
Telicota, 6, 79, 83, 85, 90
Temnochila, 51
tenebraria, var. wockearia (Dasydia),
xviii
‘enebrosus (Melanotus), 45
enuiseapa (Xylocopa), 126
( cxliii_ )
Tephrinia, 79, 108, 110
Tephrosia, xxvii
Teracolus, xv, xxxviii, lvi, lvii, 1xi,
ibe Meera, 7, day 9h tS salto
104, 105, 106, 107, 108, 117, 119,
134, 139, 141, 142, 144, 251, 372,
393
teredon (Papilio), 111, 130
Terias, xliii, 61, 66, 67, 68, 69, 70, 71,
79, 73. 75. 79, 81. 82, 88, 84, 87, 89,
92, 95, 98, 99, 101, 102, 103, 104,
NOS MOG LOM. LOOs ATS Weiss Lor
121, 122, 129, 130, 132, 134, 139,
140, 141, 142, 143, 144
Teriomima, 253
terlootii (Neophasia), xx, xxii
terricola (Piezostethus), 50
Tessaratoma, 126
testacea (Hydrena), 51
testaria (Amygdaloptera), 387
tetralunaria (Selenia), xxix
Tetropium, Ixvii, 146
Tettigonia, 330, 332, 333
Tettigonides, 332
Thais, v, 203-218, 371
Thalattoscopus, 334, 363
Thamnonoma, 388
thaumas (Adopea), 380
Thecla, 42, 377
theophrastus (Lampides), 379
3 (Tarucus), 71, 78, 79, 108,
104
Therapha, 46, 50
thersamon, ver.
phanus), 378
Thestor, 378
thestylis (Prioneris), 97, 98, 141
thetodes (Gymnobathra), 239
thetys (Curetis), 90, 134
Thisoicetrus, 81
thoracica (Elis), 88
thoracicus (Homoporus), 434
thranias (Borkhausenia), 240
thyelia (Sympheedra), 107
Thylacites, 47, 49
thymi (Nysius), 50
thyodamas (Cyrestis), 96
thyridias (Scoparia), 229
thysa (Belenois), lvii
tibetanus (Criocephalus), 155, 159,
164
tibialis (Cryptocephalus), 46
», (Heterocordylus), 46
Tibicen, 329
Ticherra, 97
tigrinus (Corizus), 50
omphale (Chryso-
Timarcha, 42, 46, 47, 50
Tinegeria, 73
Tinea, 235°
tipularius (Neides), 52
tirrhea (Ophiusa), 384
Tirumala, 73, 75, 87, 89, 90, 91, 100,
107, 109, 117, 118, 128, 124, 127,
134, 1385, 138
titan (Gonometa), 313
Titanoptilus, 248
tithonus (Epinephele), 40, 42
Titubcea, 49
toboranus (Papilio), 251
toletanum (Bembidium), 48
Tomicus, 51
topha (Teracolus), xv, xvi
torquatum (Anthobium), 44
Tortrix, lxili, 234
touhilimasa (Precis), 251
Trabala, 66
Trachypepla, 236, 237
transfugus (Helophilus), lxv
transversa (Corixa), 52
trapezalis (Marasmia), 77
trapezitis (Tauroscopa), 227
Trechus, 47, 48
triangularis (Brachycoleus), 50
Trichobyrrhulus, 48
Trichodes, 49
Trichomastix, 424, 425, 430
Trichoptilus, 249
tricinctus (Bassus), 425, 431
triclera (Scoparia), 230
tricolor (Pentatoma), 354
trifolii (Pachygastria), 390
Trigona, xviii
Trigonodes, 109
trimeni (Cooksonia), 257, 262
», (Epamera), 252
», (Mylothris), lviii
tripars (Aufidus), 332
tripunctatus (Lionotus), 412
Hp (Odynerus), 412
Trisceedecia, 247
tristalis (Hypena), 67
triste (Callidium), 158
tristrigoa (arias), 71, 79
tritici (Agrotis), 1xi
trivia (Melita), 78, 74
Tropicoris, 50
truncata (Polyphasia), 68
truncatellus (Otiorrhynchus), 46, 49
Truxalis, 71, 73, 134
Tryphena, lxvii
Trypheromera, 96, 98, 127
Tryphonides, Ixxvi
tuberculata (Hydrotea), vii
tuberculifer (Phyllobius), 42, 45
tyndarus (Hrebia), 39, 40
var. ottomana (Erebia), 39
( exliv )
typheus (Geotrupes), 44
tytia (Caduga), 96
Ueana, 328, 329, 363
ulicis (Apion), 45
uliginosus (Elaphrus), 43
ulyssiponensis (Micrositus), 49
umbrata (Platylomia), 201
umbratilis (Philonthus), 51
umbrosa (Euptychia), 186
Umjaba, 196, 202
unca (Hydrelia), 109
undularis (Elymnias), 88, 89, 92, 98,
115
unicolor (Hister), 44
unifasciatus (Pceciloscytus), 50
uniformis (Asura), 129
uniguttatus (Dieuches), 110
Uranothauma, 253
urinator (Gyrinus), 51
urtice (Vanessa), xliii
Utetheisa, 382
vagata (Acropteris), 97
Vanessa, xviii, xxix, xliii, 40, 65, 66,
69, 72, 80, 82, 98, 94, 107, 112, 113,
134, 136, 373
vardhana (Cyaniris), 66, 68
varia (Lechriolepis), 319
variabilis (Cantao), 361
», (Phytodecta), 46
- (Scutellera), 361
aA (Zonabris), 49, 52
varicoxa (Bassus), 425, 431
variegata (Libellula), 120
», (Philereme), 67, 68
variegatus (Hoplopus), 408
x (Odynerus), 408
varipes (Phytocoris), 50
varius (Hydroporus), 51
varmona (Neptis), 95, 99, 107, 109,
110, 111" 11s. T77. vary 196,137,
129, 180, 132
varunana (Chilades), 79, 101
vaucheri (Coenonympha), 377, 393
velleda (Hepialus), 41
velutina (Vespa), 73
velutinus (Chlznius), 44
venosa (Bombycopsis), 322, 323
venustus (Dynamenais), 344
», (Ectatops), 344
Verlusia, 50
verma (Lethe), 95
versicolor (Pelochares), 51
verticillata (Diapria), lxv
vesicatoria (Lytta), 42, 45
Vespa, 72, 73, 417
Vesperus, 49, 51, 52
vetustus (Heliconius), li
villica (Arctia), 381
villica, var. konewkai (Arctia), 881
villosella (Pachytelia), 40
viminalis (Bombycia), 383
vinula (Dicranura), 387
viola (Pseudometa), 314
viole (Telchinia), 102, 103, 109, 110,
115, 119, 120, 133
Virachola, 101
virescens (Gicophylla), 279
», (Oncotympana), 201
virgaurex, var. miegii (Polyommatus),
41, 42
viride (Cuspicona), 357
,, -(Pentatoma), 357
viridulus (Stenobothrus), xvii
vishnu (Trabala), 66
vitellinz (Phyllodecta), 46
Vitellus, 356
vittatus (Baracus), 130, 131
», (Betarmon), 44
», (Cryptocephalus), 42, 46
vittiger (Phytocoris), 50
vorax (Apion), 49
vuillefroyi (Nebria), 46, 48
vulcanus (Spindasis), 94, 119
wallengreni (Beralade), 320
waterhouseri (Ogyris), 271, 273, 275,
289
willeyi (Adrisa), 358
s, (Geocoris), 349, 363
>, (Tettigonia), 333
woodlarkianus (Lygzeus), 343
xanthaspis (Homoporus), 428, 435
xanthindyma (Cosmophila), 132
Xanthorhoe, 220
Xenarthra, xxvi
xiphia (Nychitona), 79, 87, 89, 92, 101,
110, 115, 120, 127
xuthus (Papilio), xxxiv
Xylocopa, 126
Xylotrupes, 132
Yanga, 194, 195, 202
Yphthima, 72, 78, 82, 87, 88, 89, 92,
95, 98, 99, 107, 111, 113, 114, 121,
125, 138, 184, 188, 189, 140, 141,
142
ypsilon (Agrotis), 382
Zabrus, 42, 48, 46, 48
zalmora (Neopithecops), 90, 92
Zanclognatha, 385
zaneka (Gonepteryx), Xxxviii, xxxix
Zangis, 356
zapateri (Erebia), 28
Zegris, 1xxv
zelea (Eurythecta), 233
zelleri (Parnara), 380
Zemeros, 95, 97
Zeritis, 252
( exlv )
Zeuzera, 391
ziczac (Notodonta), ii
zinckenella (Etiella), 108
Zizera, 66, 70, 71, 73, 74, 76, 81, 83,
99, 102, 108, 109, 119, 121, 127,
133
zochalia (Belenois), lviti
Zonabris, 42, 45, 47, 49, 52
Zonosoma, xxxii, 96
Zootrephus, 424, 425, 430
zopherana (Strepsicrates), 233 x
zosine (Ogyris), 269, 270, 278, 289,
290, 292
», -duaringa (Ogyris), 280, 289
», -magna (Ogyris), 281, 289
Zygena, 72, 391
Marcu 7TH, 1906.
Yee me ty ty hy
a rr HE wings eas £ 7 7” sey
oe: US neapttats eae Fy Ar ne pre.
My Ree et 7 on haz iy
| ord
ys me es %
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yt
=
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i a oe
Trans. Ent. Soc. Lond., 1905. PI. I.
2
SERN Ce * oa 7
3 ec
ta
Andre & Sleigh, Ltd.
Life-History of Gerydus chinensis, Feld.
ivans Ent. Soc. onda 905m eel
Erebia palarica, etc.
Erebia stygne
Trans. Ent.
and E. evias.
Soc.
Lond.
1905.
Br:
UE
Pia a i> ;
¥ a F.
es
5 ~
+ pete tS
Trans. Ent. Soc. Lond., 1905. PL. IV.
. evias.
Erebia stygne and E
“SPIqeIA eos jo UOHNGIysIp yyw ‘uredS useYyZION ‘dey ‘Yp324S
Shiavedez'7 SBIAT TT 20lse/eg q ‘edjuedsiy seA Sela] —Q—o— aushig —o—o— Pr
\ noasiigyo >)
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Trans. Ent, Soc. Lond., 1905.
© puoseie
telat
\\ Wee
Trans. Ent, Soc. Lond., rogs. Pi. Vi.
Erebia palarica and E. stygne.
Trans, Ent. Soc. Lond., 1905. Pi. VII.
Monastery of Arbas.
Trans. Ent. Soc Lond. 1905.Pt. VIL.
West, Newman lith.
Overand arvesto: Dalader acuireosta.t:
me os
v lv
axle : ? zB .
M.A Sharp,del.et lith.
CRI CHE PHA
AND ASEMUM.
Beara (4
od tan
al
Me ae
E.Wilson,Cambridge.
Trans. Ent. Soc. Lond., 1905. Plate X.
Horace Knight. del. Andre & Sleigh Limited.
New species of Satyridz from South America.
a
« S eae
. 1 ae
"9 bie
i *
to
Trans. Ent. Soc. Lond., 1905. Plate XI.
Horace Knight, del. Andre & Sleigh, Ltd.
New Species of Cicadide.
Trans. Ent. Soc. Lond. 1905. Plate XII.
lugh Main, Esq., F.E.S. Andre & Sleigh, Ltd.
Pupal Suspension of Thais.
Trans. Ent. Soc. Lond. 1905 Pl. XI.
Horace Knight del West, Newman chromo.
NEW RHODESIAN BUTTERFLIES.
PUSXIV:
Trans. Ent. Soc. Lond. 1905.
‘O8U0D 9} JO S9DANOS 9Y} WIOJJ ‘SOLUITU JOIYD OA} S} UIA snddiskayd SDUULT
‘azis joAnjDOU ay} fo § aAv sSaanslf ay} IP
"PIT ‘YSIRIS B orpuy : ‘ojoyd ‘uosuiqoy peaTy
ees
Paik OE eer 5
Trans. Ent.Soc. Lond. 1905. PL XV.
E.C.Knight ad nat.lith. West, Newman imp.
GENITALIA OF OGYRIS.
- odes sed? Dea Weel s
Knight, del. Andre & Sleigh, Ltd.
New African Lasiocampide.
Trans. Fint. Soc. Lond. 1905. Pl. XVI.
M.A.Sharp del, et lith. Edwin Wilson, Cambridge.
Rhynchota from New Britain etc.
AROSS A: \
aa duz
eee = if 15 500
Trans. Ent. Soc. Lond. 1905, Pl, XVII
Tizi Wado
ae
tiny
q
Jes
i ‘Erdax XQ
ii sana : .
Pduz es Tizi- -n- Prine
JWirsan_ ==
+ ne
X
Ny \
William Stanford & Company, Ltd, SCALE OF MILES
Mk
Tizi Gourza
12,004 4
8° Long. West of Greenwich
The Oxford Geographical Institute.
° 5 10
MEADE-WALDO’S ROUTE IN MAROCCO.
Plate XIX.
nt. Soc. Lond., 1905.
E
Trans.
Andre & Sleigh, Ltd.
i. Knight, del.
Marocco.
Lepidoptera from