TRANSACTIONS
OF THE
ENTOMOLOGICAL
OF
SOCIETY
LONDON
■l
u
.1
*
THE
TRANSACTIONS
OF THE
3 \
ENTOMOLOGICAL SOCIETY
OF
LONDON
FOR THE YEAR
1910.
LONDON:
PRINTED FOE THE SOCIETY BY RICHARD CLAY AND SONS, LIMITED,
LONDON AND BUNGAY.
SOLD AT THE SOCIETY’S ROOMS, 11, CH ANDOS STREET,
CAVENDISH SQUARE, W.,
AND BY LONGMANS, GREEN AND CO.,
PATERNOSTER ROW, E.C. ; AND NEW YORK.
1910-1911.
r?,(ob
DATES OF PUBLICATION IN PARTS.
Part jj’|(Trans., p. 1-160, Proc., i-xxxii) was published 19 July, 1910.
ENTOMOLOGICAL SOCIETY OF LONDON
Founded, 1833.
Incorporated by Royal Charter, 1885.
OFFICERS and COUNCIL for the Session 1910-1911.
president.
FREDERICK AUGUSTUS DIXEY, M.A., M.D., F.R.S.
\Dice-prc0iDent0.
GEORGE T. BETHUNE-BAKER, F.Z.S.
Prof. T. HUDSON BEARE, B.Sc., F.R.S.E.
H. ROWLAND-BROWN, M.A.
Zrrea0iirer.
ALBERT HUGH JONES.
Secretarte0.
HENRY ROWLAND-BROWN, M.A.
Commander JAMES J. WALKER, M.A., R.N., F.L.S.
Xibrarian.
GEORGE CHARLES CHAMPION, F.Z.S., A.L.S.
©tber /llbember0 of Council.
MALCOLM BURR, D.Sc., F.L.S., F.Z.S.
HORACE ST. J. DONISTHORPE, F.Z.S.
ALBERT HARRISON, F.L.S., F.C.S.
Prof. SELWYN IMAGE, M.A.
KARL JORDAN, Ph.D.
HUGH MAIN, B.Sc.
ALFRED SICH.
HENRY JEROME TURNER.
ROWLAND E. TURNER.
JAMES W. TUTT.
Resident Librarian.
GEORGE BETHELL, F.R.Hist.S.
( vii )
CONTENTS.
Explanation of the Plates
List of Fellows...
Additions to the Library
Errata
MEMOIRS.
I. A Revision of the African Species of the Lycaenesthes group of
the Lycaenidae. By G. T. Bethunb-Baker, F.L.S.,F.Z.S. ... 1
II. On Callophrys avis, Chpm. By T. A. Chapman, M.D., F.Z.S.. 85
III. On the genus Deuterocnpus, Zeller. By T. Bainbrigge
Fletcher, R.N., F.E.S . 107
IV. Some Experiments with Ants' Nests. By Horace St. J. K.
Donisthorpe, F.Z.S. . 1-12
V. A few words respecting Insects and their Natural Enemies. By
Arthur G. Butler, Ph.D., F.L.S., F.Z.S., F.E.S. , M.B.O.U. 151
VI. Further Notes on two Osniia-STpecies of the adunca-growp. By
the Rev. F. D. Morice, M.A., F.E.S. ... ... ... 155
VII. A Preliminary Revision of the Labiduridae, a family of the
Dermaptera. By Malcolm Burr, D. Sc., F.L.S., F.E.S. ... 161
VIII. On the Geometridae of the Argentine Republic. By Louis B.
Prout, F.E.S . 204
IX. Third Paper on the Tetriyinae (Orthoptera) in the Oxford
University Museum. By J. L. Hancock, M.U., F.E.S. ... 346
X. Descriptions of Micro-Lepidoptera from Mauritius and the
Chagos Isles. ByE. Meyrick, B.A., F.R.S . 366
XI. The Behaviour of Coleoptera in time of Floods. By Norman
H. Jot, M.R.C.S., F.E.S . 379
XII. On the Position of the Rhopalosotmdae, with the Description of
a Second Species. By Claude Morley, F.Z.S., F.E.S. ... 386
XIII. Notes on the iScofiYdue. By Rowland E. Turner, F.Z.S. ... 391
XIV. New Fossorial Hymenoptera from Australia. By Rowland E.
Turner, F.Z.S . 407
XV. Descriptions of Malayan Micro-Lepidoptera. By E. Meyrick,
B.A., F.R.S., F.Z.S . 430
XVI. On Zizeeria (Chapman), Zizera (Moore), a group of Lycaenid
Butterflies. By T. A. Chapman, M.D. ... ... ... 479
viii
ix
xxvi
... xxxix
( viii )
XVII. Mr. Alfred D. Millar’s Experimental Breeding from the Ova of
the Natalian forms of the Nymphaline Genus Euralia. By
Roland Teimen, M.A., F.R.S., etc . « 498
XYIII. On a new Genus of Stylopidae from Australia. By Arthde M.
Lea, F.E.S., Government Entomologist, Tastnania . 511
XIX. Hymevoptera Aculeata collected in Algeria by the Rev. Alfred
Edwin Eaton, M.A., F.E.S., and the Rev. Francis David
Morice, M.A., F.E. S. Part IV. Descriptions of new
Spheyidae. By the late Edward Saunders, F.R.S . 517
XX. Descriptions of two New Species of Luciola in the Collection of
Mr. H. E. Andrewes. By Ernest Olivier, F.E.S . 532
Proceedings for 1910
President’s Address
General Index
Special Index
... i-lxxxiii
Ixxxv
cvii
cxii
EXPLANATION OF PLATES, TRANSACTIONS.
Plates LI-LX. See pages 495-497
Plates LXI-LXV. „ 512-313
Plate LXVI. „ 516
Proceedings.
Plate A See page Ixi
( ix )
yist of Jfjllotos
OF THE
ENTOMOLOGICAL SOCIETY OF LONDON.
HONORARY FELLOWS.
Election.
1900 Adrivillius, Professor Christopher, Stockholm.
1905 Bolivar, Ignacio, Paseo de Recoletos Bajo, 20, Madrid.
1901 Fabre, J. H., Serignan, Vaucluse, France.
1894 Forel, Professor Auguste, M.D., Chigny, pres Morges, Sioitzerland.
1906 Ganglbauer, Gustos Ludwig, Hof- Museum, Vienna.
1898 Grassi, Professor Battista, The University, Home.
1908 Oberthur, Charles, Rennes, Rle-et-Vilaine, France.
1906 Reuter, Professor Odo Morannal, The University, Helsingfors,
Finland.
1895 Scudder, Samuel Hubbard, Cambridge, Mass., U.S.A.
1885 Snellen, P. C. T., Rotterdam.
1893 Wattenwyl, Dr. Carl Brunner von, Schbnhurgstrasse 3, Vienna.
1898 Weismann, Di’. August, Freiburg, Baden.
FELLOAVS.
j\farked f have compounded for their Annual Subscrigjtions.
Marked * deceased during the year.
Date of
Election.
1908 Ackerley, F. B., c/o Imperial Tobacco Co., P. 0. Box 1159,
Johannesburg, South Africa.
1901 t Adair, Sir Frederick E. S., Bart., Flixton Hall, Bungay.
1877 Adams, Frederick Charlstroiu, F.Z.S., 50, Ashley-gardens, Victoria-
street, S.W.
1877 Adams, Herbert J., Roseneath, London-road, Erifield, N.
1902 Ad KIN, Benaiah Whitley, Trenoweth, Hope-park, Bromley, Kent.
1885 Adkin, Robert, Wellfield, Lingards-road, Lewisham, S.E.
1904 Agar, E. A., La Haut, Dominica, B. TV. Indies.
1904 Alderson, Miss E. Maude, Park House, TT^orksop, Notts.
( X )
lUlOt Andrewes, H. E., 8, North Grove, Hicjlujate, N.
1899 Andrews, Henry W., Shirley, Welling, 8.O., Kent.
1901 Anning, William, 39, Lime Street, E.C.
1908 1 Antram, Charles B., The Insectarium, Kanny-Koory, Silchar
P. 0., Cachar, India.
1907 Arnold, G., M.Sc., University of Liverpool, Liverpool.
1899 f Arrow, Gilbert J., 87, Union-grove, Glapham, S.W. ; and British
Museum (Natural History), Cromwell-road, S.W.
1907 f Ashby, Sydney ll, 119, Oreenvale-road, Eltham-park, Kent.
1886 Atmore, E. a., 48, High-street, King's Lynn.
1850 t Avebury, The Right Honble. Lord, D.C.L., F.R.S., F.L.S., F.G.S.,
etc.. High Elms, Farnborough, Kent.
1901 Bacot, Arthur W., York Cottage, York-hill, Loughton, Essex.
1904 1 Bagnall, Richard S., Penshaw Lodge, Penshaw, Durham.
1909 Bagwell-Pdrefoy, Capt. Edward, 34, Sloane-Court, S.W.
1903 Baldock, G. R., Oakbnrn Villa, Enfield Highway, Middlesex.
1886 Bankes, Eustace R., M.A., Norden, Corfe Castle, Wareham.
1890 Barclay, Francis H., F.G.S., The Warren, Cromer.
1886 Bargagli, Marchese Piero, Piazza S. Maria, Palazzo 'Tempi No. 1,
Florence, Italy.
1895 Barker, Cecil W., The Bungalow, Malvern, Natal, South Africa.
1902 Barraud, Philip J., Bushey Heath, Watford.
1907 Bartlett, H. Frederick D., 1, Myrtle-road, Bournemouth.
1894 f Bateson, William, M.A., F.R.S., Fellow of St. John’s College,
Cambridge, Merton House, Orantchester , Cambridge.
1908 Bayford, E. G., 2 Bockingham-street, Barnsley.
1904 Bayne, Arthur F., c/o Messrs. Freeman, Castle- street, Frumlingham,
Suffolk.
1896f Beare, Prof. T. Hudson, B.Sc., F.R.S.E., Vice-President, 10,
Regent Terrace, Edinburgh.
1908 Becher, Major Edward F., Cranfield House, Polzeath, St. Minver,
Cornwall.
1908 Beck, Richard, Red Lodge, Porchester-road, Bournemouth.
1905 Bedford, The Duke of, K.G., Pres. Z.S., etc., Woburn Abbey,
Beds.
1899 Bedwell, Ernest C., Bonnicot, The Grove, Coulsdon, Surrey.
1904 Bengtsson, Simon, Ph.D., Lecturer, University of Lund, Sweden ;
Curator, Entomological Collection of the University.
1906 Bentall, E. E., The Towers, Heybridge, Essex.
1885 Bethune-Baker, George T., F.L.S., F.Z.S., Vice-President, 19,
Clarendon-road, Edghaston, Birmingham.
1895 Bevan, Lieutenant H. G. R., R.N., 38, The Common, Woolwich.
1880 * Bignell, George Carter, The Ferns, Ilomeparlc-ruad, Saltash.
1891 Blaber, W. H., F.L.S., 34, Cromwell-road, Hove, Brighton.
1904 Black, James E., Nethercroft, Peebles.
1904 Blair, Kenneth G., 23, West Hill, Highgate, N.
( xi )
1889 Blandford, Walter F. H., M.A., F.Z.S., 12, Anmdel Gardens,
Ladbroke-grove, W.
1885 Blathwayt, Lt.-Col. Linley, F.L.S., Eagle House, Batheaston,
Bath.
1909 Blenkarn, Stanley A., Norham, Cromwell-road, Beckenham.
1904 Bliss, Maurice Frederick, Coningsburgh, Montpelier-road, Ealing, W.
1886 Bloomfield, The Rev. Edwin Newsou, M.A., Guestling Rectory,
Hastings.
1903 Bogue, W. a.. The Bank House, Watchet.
1907 Bonnet, Alexandre, 54, Boulevard Bineau, Neuilly-sur-Seine,
Seine, France.
1891 Booth, George A., Fern Hill, Grange-over -Sands, Carnforth.
1902 Bostock, E. D., Holly House, Stone, Staffs.
1904 Bourgeois, Jules, Ste. Marie-aux-Mines, Markirch, Germany.
1892 Bouskell, Frank, Market Bosworth, Nuneaton.
1888 Bower, Benjamin A., Langley, Willow Grove, Ghislehurst.
1894 f Bowles, E. Augustus, M.A., Myddelton House, Waltham Cross.
1910 Boyd, A. Whitworth, The Alton, Altrincham, Cheshire.
1852 t Boyd, Thos., Woodvale Lodge, South Norwood Hill, S.E.
1893 Brabant, fidouard. Chateau de Morenchies, par Cambrai (Nord),
France.
1905 Bracken, Charles W., B.A., 5, Garfrae Terrace, Lipson, Plymouth.
1907 Brain, Charles Kimberlin, Government Experimental Station, Rose-
bank, Cape Colony.
1904 Bridgeman, Lieut. The Hon. Richard 0. B., R.N., Weston Park,
Shifnal, Salop.
1877 Briggs, Charles Adolphus, Rock House, Lynmouth, B.S.O., N.
Devon.
1870 Briggs, Thomas Henry, SLA., TJoc/c House, Lynmouth, R.S.O., N.
Devon.
1894 Bright, Percy M., Fairfield, Wimborne-road, Bournemouth.
1909 Britten, Harry, Prospect House, Salkeld Dykes, Penrith.
1902 Broughton, Captain T. Delves, R.E., R. A. and R. E. Mess, Malta.
1878 Broun, Major Thomas, Mount Albert, Auckland, New Zealand.
1904 Brown, Henry H., Castlefield Tower, Cupar, Fife, N.B.
1910 Browne, Horace B., M.A., 118, Sunny Bank, Hull.
1909 Bryant, Gilbert E., Fir Grove, Esher, Surrey.
1898 1 Buchan-Hepburn, Sir Archibald, Bart., J.P., D.L., Smeaton-
Hepburn, Prestonkirk.
1907 Bulleid, Arthur, F.S.A., Wimboro, Midsomer Norton, Somersetshire.
1902 Buller, Arthur Percival, Royal Societies Club, S.W.
1896 1 Burr, Malcolm, D.Sc., F.L.S., F.Z.S., F.G.S., A.R.S.M., Castle
Hill House, Dover.
1909 Burrows, The Rev. C. R. N., The Vicarage, Mucking, Stanford-le-
Hope, Essex.
1868 f Butler, Arthur G., Ph.D., F.L.S., F.Z.S., The Lilies, Penge-road,
Beckenham.
( xii )
1883 Butler, Edward Albert, B.A., B.Sc., 56, Cecile-Park, Orouch End, N.
1902 Butler, William E., Hayling House, Oxford-road, Reading.
1905 Butterfield, Jas. A., B.Sc., Comrie, Eglinton Hill, Plumstead.
1904 Byatt, Horace A., B.A., Berbera (via Aden), Somaliland Protectorate.
1902 Cameron, Malcolm, M.B., R.N., H.M.S. “ Attentive," Home
Fleet.
1885 Campbell, Francis Maule, F.L.S., F.Z.S., etc., Byrnllwydimjn,
Machynlleth, Montgomerysh ire.
1898 Candeze, L4on, Mont St. Martin, 75, Lihge.
1880 Cansdale, W. D., Sunny Bank, South Norwood, S.E.
1889 Cant, A., 33, Festing-road, Putney, S.W. ; and c/o Fredk. Du Cane
Godman, Esq., F.B.S., 45, Pont-street, S.W.
1890 Capper, Samuel James (President of the Lancashire and Cheshire
Entomological Society), Huyton Park, Liverpool.
1894 Caracciolo, H., H.M. Customs, Port of Spain, Trinidad, British
West Indies.
1910 Carlier, E. Wace, M.D., F.R.S.E., Morningside, Granville-road,
Durridge, and The University, Birmingham.
1892 Carpenter, The Honble. Mrs. Beatrice, 22, Grosvenor-road, S.W.
1910 Carpenter, Geoffrey D. H., B.A., Uganda Medical Service,
Uganda Protectorate.
1895 Carpenter, G. H., B.Sc., Royal College of Science, Dublin.
1898 Carpenter, J. H., Redcot, Belmont-road, Leatherhead.
1868 Carrington, Charle.s, Meadowcroft, Horley, Surrey.
1895 Carter, Sir Gilbert, K.C.M.G., 43, Charing Cross, W.C., and
Government House, Nassau, Bahamas.
1906 Carter, H. J., B.A., Ascham, Darling Point, Sydney, N.S. Wales.
1900 Carter, J. W., 15, Westfield-road, Manningham, Bradford.
1900 Cassal, R. T., M.R.C.S., Ballaugh, Isle of Man.
1903 Cattle, John Rowland, Nettleton Manor, Caistor, Lincolnshire.
1889 f Cave, Charles J. P., Diteham Park, Petersfield.
1900 Chamberlain, Neville, Highbury, Moor Green, Birmingham.
1871 Champion, George C., F.Z.S., A.L.S., Librarian, IJeatherside,
Horscll, Woking ; and 45, Pont-street, S.W.
1891 Chapman, Thomas Algernon, M.D., F.Z.S., Betula, Reigute.
1910 Charnley, j. R., Lyndhurst, Fulivood, Preston.
1897 Chawner, Miss Ethel F., Forest Bunk, Lyndhurst, R.S.O., Hants.
1902 Cheesman, E. M., c/o Mr. John Garson, 150, Umhilo-road, Durban.
1908 Chetty, B. Chourappa, The Government Museum, Bangalore,
India.
1889 Christy, William M., M.A., F.L.S., Watergate, Emsworth.
1909 Chubb, Ernest C., Curator, Durban Museum, Natal, South Africa.
1909 Clark, C. Turner, F.Z.S., 90, The Mall, Newport, Isle of Wight.
1908 Clark, Edgar L., Congella, Natal.
1867 Clarke, Alex. Henry, 109, W'artvick-road, Earfs Court, S.W.
1908 Clutterbuck, Charles G., Heathside, 23, Heathville-road, Gloucester
( xiii )
1908 Clutterbtjck, P. H., Indian Forest Department, Naini Tal, United
Provinces, India.
1904 Cockayne, Edward A., 16, Camhridge-square, London, W.
1873 Cole, William, F.L.S., Springfield, Buckhurst Hill, Essex.
1899 Collin, James E., Sussex Lodge, Neiomarket.
1906 CoLLiNGE, Walter E., M.Sc., F.L.S., 55, Newhall- street, Bir¬
mingham.
1892 Cowan, Thomas William, F.L.S., F.G.S., F.R.M.S., Upcott House,
Taxmton.
1867 Cox, Herbert Ed., Claremont, Jamaica.
1895 Crabtree, Benjamin Hill, The Oaklands, Levenshulmc, Manchester.
1909 Crawley, W. C., Tollerton Hall, Nottingham.
1906 Crawshay, The Rev. George A., MA., Melchhonrne Vicarage,
Sharnbrook, S.O., Beds.
1890 Crewe, Sir Vauncey Harpur, Bart., Calke Abbey, Derbyshire.
1880 t Crisp, Sir Frank, LL.B., B.A., J.P.
1907 Croft, Edward Octavius, M.D., 28, Clarendon-road, Leeds.
1902 Cruttwell, The Rev. Canon Charles Thomas, M.A., Eivelme
Rectory, Wallingford.
1908 CuLPiN, Millais, M.B., F.R.C.S., The Palace Hotel, Shanghai.
1908 Curtis, W. Parkinson, Aysgarth, Poole, Dorset.
1901 Dadd, Edward Martin, Hohenzollernstrasse 18, Zehlendorf, bei Berlin.
1900 Dalglish, Andrew Adie, 21, Prince’ s-street, Glasgoio.
1907 Dames, Felix L., 13, Humboldt-strasse, Steglitz, Berlin.
1886 Dannatt, Walter, Donnington, 75, Vanbrugh Park, Blackheath,
S.E.
1905 Davidson, James D., 32, Drumsheugh Gardens, Edinburgh.
1910 Dawson, William George, 31, King’s Gardens, West End Lane,
West Hampstead, N.W.
1903 Day, F. H., 26, Currock-terrace, Carlisle.
1898 Day, G. 0., Sahlatston, Duncan’s Station, Vancouver Island, British
Columbia.
1905 Dewar, W. R., Government Entomologist, Fruit Land, Ontario,
Canada.
1875 Distant, Wm. Lucas, Shannon-lodge, Selhurst-road, South Norwood,
S.E.
1887 Dixey, Frederick Augustus, M.A., M.D., President, Fellow and
Bursar of Wadham College, Wadham College, Oxford.
1895 Dobson, H. T., Ivy House, Acacia Grove, New Malden, S.O.,
Surrey.
1909 Dobson, Thomas, Park Avenue, The Park, Sharpies, Bolton.
1905 Dodd, Frederick P., Kuranda, via Cairns, Queensland.
1906 Dollman, Here ward. Hove House, Newton-grove, Bedford-park, W.
1903 Dollman, J. C., Hove House, Neivton-grove, Bedford-park, W.
1906 Doncaster, Leonard, M.A., The University Museum of Zoology,
Cambridge.
( )
1891 Donisthorpe, Horace St. John K., F.Z.S., 58, Kensington-inansions,
South Kensington, S.W.
1908 Dodglas-Crompton, Sydney, Carlton House, Kew-green, S.W.
1910 Downes-Shaw, Rev. Archibald, Kettlestone Rectory, Fakenham,
Norfolk.
1884 Droce, Hamilton H. C. J., F.Z.S., 43, Circus-road, St. John's
Wood, N.W.
1867 Druce, Herbert, F.L.S., F.Z.S., 43, Circus-road, St. John's Wood,
N.W.
1900 Drury, W. D., Rocquaine, West Hill Park, Woking.
1894 Dudgeon, G. C., The Imperial Institute, South Kensington.
1906 Dukinfield- Jones, E., Castro, Reigute.
1883 Durrant, John Hartley, Merton, 17, Bur stock-road, Putney, S.W.,
and British Museum {Natural History), Cromwell- road, South
Kensington, S.W.
1910 Eales-White, J. Cushny, 21, Cardigan-road, Richmond Hill, S.W.
1890 Eastwood, John Edmund, Enton Lodge, Witley, Godaiming.
1865 Eaton, The Rev. Alfred Edwin, M.A., West House, Simondsbury,
Bridport, Dorset.
1904 Eckford, George, F.Z.S., c/o Sir Morgan Tiiite, Bart., Kilruune,
Nenagh, co. Tipperary, Ireland.
1902 Edelsten, Hubert M., The Elms, Forty Hill, Enfield, Middlesex.
1886 Edwards, James, Colesborne, Cheltenham.
1884 Edwards, Stanley, F.L.S., F.Z.S., 15, St. Germans-place, Black-
heath, S.E.
1900 Elliott, E. A., 16, Belsize Grove, Hampstead, N.W.
1900 Ellis, H. Willoughby, Holly Hill, Berkswell, Warrvickshire.
1886 Ellis, John W., M.B., L.R.C. P., 18, Rodney-street, Liverpool.
1903 Eltringham, Harry, M.A., F.Z.S., Eastgarth, Westoe, South Shields.
1878 Elwes, Henry John, J.P., F.R.S., F.L.S., F.Z.S., Colesborne,
Cheltenham.
1886 Enock, Frederick, F.L.S., 13, Tufnell Park Road, London, N.
1903 Etheridge, Robert, Curator, Australian Museum, Sydney, N.S.W.
1908 Eustace, Eustace Mallabone, B.A., Challacombe Rectory, Parra-
combe, R.S.O., N. Devon, and Wellington College, Berks.
1909 Evans, Frank J., The Botanical Department, Trinidad, B.W.I.
1899 Farmborough, Percy W., Lower Edmonton, Middlesex. ,
1907 Feather, Walter, 10, Station Grove, Crosshills, nr. Keighley, Yorks.
1900 Feltham, H. L. L., P. 0. Box, 46, Johannesburg, Transvaal.
1861 Fenn, Charles, Eversden House, Burnt Ash Hill, Lee, S.E.
1886 Fenwick, Nicolas Percival, The Gables, New-roud, Esher.
1908 Fenwick, Norman Percival, Junior, The Gables, New-road Esher.
1889 Fernald, Prof. C. H., Amherst, Mass., U.S.A.
1910 Feynes, a., M.D., 61, East Colorado-street, Pasadena, California,
U.S.A.
( XV )
1900 Firth, J. Digby, F.L.S., Boys Modern School, Leeds.
1874 Fitch, Edward A., F.L.S., Brick House, Maldon.
1905 Fleet, Wilfred James, Imatra, King’s Road, Bournemouth.
1900 Flemyng, The Rev. W. Westropp, M.A., Goolfin, Portlaw, Go.
Waterford.
1898 Fletcher, T. Bainbrigge, R.N., Agricultural Research Institute,
Pusa, Bengal, India.
1883 t Fletcher, William Holland B., M.A., Aldwick Manor, Bognor.
1905 Floersheim, Cecil, 16, Kensington Gourt Mansions, S.W.
1885 Fokker, a. J. F., ZieriJczee, Zeeland, Netherlands.
1900 Foulkes, P. Hedworth, B.Sc., Harper-Adams Agricultural Gollege,
Newport, Salop.
1898 Fountains, Miss Margaret, 1, The Studios, Sheriff-road, West
Hampstead, N.W.
1880 Fowler, The Rev. Canon, D.Sc., M.A., F.L.S., Earley Vicarage,
near Reading.
1908 Fraser, Frederick C., M.D., I.M.S., Hyderabad, Sind, India.
1896 Freke, Percy Evans, Southpoint, Limes-road, Folkestone.
1888 Fremlin, H. Stuart, M.R.C.S., L.R.C.P., Mereworth, Maidstone.
1903 French, Charles, F.L.S., Government Entomologist, Melbourne,
Victoria, Australia.
1910 Frisby, G. E., 40, Windmill-street, Gravesend.
1908 Froggatt, Walter W., F.L.S., Government Entomologist, 138, George-
street, Sydney, New South Wales.
1891 Frohawk, F. W., Ashmount, Rayleigh.
1906 f Fry, Harold Armstrong, P.O. Box 46, Johannesburg, Transvaal
Golony.
1900 Fryer, H. Fortescue, The Priory, Ghatteris, Gambs.
1907 Fryer, John Claud Fortescue, The Priory, Ghatteris, Gambs.
1876 Fuller, The Rev. Alfred, M.A., The Lodge, 7, Sydenham-hill,
Sydenham, S.E.
1898 Fuller, Claude, Government Entomologist, Pietermaritzburg , Natal.
1887 Gahan, Charles Joseph, M.A., Whyola, Lonsdale-road, Bedford
Park, W. ; and British Museum {Natural History), Cromwell-
road, S.W.
1910 Garcke, Emile, M.I.E.E., Witton House, Maidenhead.
1892 Garde, Philip de la, R.N., Abbotsjield, Braunton, N. Devon.
1890 Gardner, John, Laurel Lodge, Hart, West Hartlepool.
1901 f Gardner, Willoughby, F.L.S., Deganwy, N. Wales.
1910 Geary, T. H., Enderby, Leicestershire.
1899 Geldart, William Martin, M.A., 27, Beaumont-street, Oxford.
1906 1 Gibbs, Arthur Ernest, F.L.S., F.R.H.S., Kitchener’s Meads, St.
Albans.
1908 Gifb'ard, Walter M., P.O. Box 308, Honolulu, Hawaii.
1907 Giles, Henry Murray, Perth, W. Australia.
1902 Gillanders, A. T., Park Gottage, Alnwick.
( xvi )
1904 Gilliat, Francis, B.A., 81, Philbeach-gardens, S.W.
1865 f Godman, Frederick Du Cane, D.C.L., F.R.S., F.L.S., F.Z.S., South
Lodge, Lower Beeding, Horsham; and 45, Pont-street, S.W.
1890 Goldthwait, Oliver C., Berisal Lake, near Sandoion, I. of Wight.
1886 f Goodrich, Captain Arthur Mainwaring, Brislington House, near
Bristol.
1904 Goodwin, Edward, Canon Court, Wateringbury, Kent.
1898 Gordon, J. G. McH., Coj'semaZs/e, Whauph'ill, R.S.O., Wigtownshire.
1898 Gordon, R. S. G. ^icH. ,Corsemalzie, Whaiq}hill,R.S.O., Wigtownshire.
1855 Gorham, The Rev. Henry Stephen, F.Z.S., Highcroft, Great
Malvern.
1910 Gottmann, a. M. S., 94, Niddastrasse, Frankfurt-am-Main, Germany.
1909 Gowdey, Carlton C., B.Sc., Uganda, British East Africa.
1891 f Green, E. Ernest, Government Entomologist, Royal Botanic
Gardens, Peradeniya, Ceylon ; and Mote Hall, Bearsted, Kent.
1910 Green, Herbert A., The Central Fire Station, Durban, Natal.
1894 Green, J. F., F.Z.S., West Lodge, Blackheath, S.E.
1898 Greenshields, Alexander, 38, Blenheim-gardens, Cricklewood, N.W.
1893 t Greenwood, Henry Powys, F.L.S., Whitsbury House, Salisbury.
1888 Griffiths, G. C., F.Z.S., Penhurst, 3, Leigh-road, Cliftoii, Bristol.
1894 Grimshaw, Percy H., Royal Scottish Museum, Edinburgh .
1905 Grist, Charles J., Elgin House, Knockholt, Sevenoaks.
1869 Grose-Smith, Henley, J.P., B.A., F.Z.S., The Priory, nr. Ryde,
Isle of Wight.
1909 Grosvenor, G. H., M.A., .3, Blackhall-ruad, Oxford.
1906 Gurney, Gerard H., Kesunck Hall, Normch.
1910 Gurney, Wm. G., Asst. Govt. Entomologist, Department of Agri¬
culture, Sydney, Australia.
1906 Hall, Arthur, 7, Park-lane-mansions, Croydon.
1890 1 Hall, Albert Ernest, Cranfield House, Southwell, Notts.
1885 Hall, Thomas William, Stanhope, The Crescent, Croydon.
1898 Hamlyn-Harrls, R., D.Sc., F.Z.S., F.R.M.S., Director of the Queens¬
land Museum, Brisbane, Australia.
1891 Hampson, Sir George Francis, Bart., B.A., F.Z.S., 62, Stanhogw-
gardeus, S.W.
1891 Hanbury, Frederick J., F.L.S., Brockhurst, E. Grinstead.
1905 t Hancock, Joseph L., 5466, Lexington Avemie, Chicago, U.S.A.
1903 Hare, E. J., Dunham, Boscombe, Hants.
1904 Harris, Edward, St. Conan’s, Chingford, Essex.
1897 f Harrison, Albert, F.L.S., F.C.S., Delamere, Grove-road, South
Woodford, Essex.
1910 Harwood, Philip, 23, Northgate End, Bishop’s Stortford.
1910 Hawkshaw, j. C., Hollycombe, Sussex.
1910 Hedges, Alfred van der, 42, Kensington Park Gardens, W.
1910 Henderson, J., Clifton, Ashbourne, Derby.
1881 Henry, George, Ivy Bank, 112, London-road, St. Leonards-on-Sea.
«
( xvii )
18!)8 Heron, Francis A., B.A., British Museum (Natural History),
Cromwell-road, S.W.
1903 Herrod, William, W.B.C. Apiary, Old Bedford-road, Luton, Beds.
1908 Hewitt, C. Gordon, D.Sc., Central Experimental Farm, Dept, of
Agriculture, Ottaiva, Canada.
1876 t Hillman, Thomas Stanton, Eastcjate-street, Leioes.
1907 Hoar, Thomas Frank Partridge, Mercia, Alhany-road, Leighton-
Buzzard.
1888 Hodson, The Rev. J. H., B.A., B.D., JRhyddington, Clifton Drive,
Lythgm.
1902 Hole, R. S., c/o Messrs. King and Co., Bombay.
1910 Holford, H. Oliver, Elstead Lodge ,Godalming, Surrey.
1887 Holland, The Rev. W. J., D.D., Ph.D., bth Avenue, Pittsburg,
Penn., U.S.A.
1898 Holman-Hunt, C. B., Asst. Entomologist, Department of Agri¬
culture, Kuala Lumpur, Federated Malay States.
1910 Holmes, Edward Morrell, Ruthven, Sevenoaks.
1901 Hopson, Montagu F., L.D.S., R.C.S.Eng., F.L.S., 30, Thurlow-road,
Rosslyn Hill, N.W.
1897 Horne, Arthur, 60, Oladstone-place, Aberdeen.
1903 Houghton, J. T., \, Portland-place, Worksop.
1907 t Howard, C. W., Entomologist Department of Agricxdture, Box 255,
Loxirengo Marques, Portuguese East Africa.
1900 Howes, George W., 812, George-street, Dunedin, New Zealand.
1907 Howlett, Frank M., M.A., Wymondham, Norfo%.
1865 tHDDD, A. E., 108, Pembrolce-road, Clifton, Bx'istol.
1888 Hudson, George Vernon, Hill View, Karoi'i, Wellington, New
Zealand.
1907 Hughes, C. N., 3, Wyndham Place, Bryanstoxi-square, W.
1897 Image, Prof. Selvvyn, M.A., 20, Fitzrny-street, Fitzroy-squax-e, W.
1908 Irby, Captain Leonard Paul, Brook House, Eastry, S.O., Kent.
1891 IsABELL, The Rev. John, Sunnycroft, St. Sennen, R.S.O., Cornwall.
1907 Jack, Rupert Wellstood, Assistant Entomologist, Department of
Agriculture, Salisbury, Rhodesia.
1907 Jackson, P. H., 112, Balham-park-i’oad, S.W.
1907 Jacobi, Professor A., Ph.D., Director of the R. Zoological and
Anthrop.-Ethnographical Museum, Dresden, Saxony.
1910 Jacobs, Lionel L., S.E. corner of Church and Wellington-streets,
Sault Ste. Marie, Ontario, Canada.
1869 Janson, Oliver E., Cestria, Claremont-road, Highgate, N.; and 44,
Great R ussell-street, Bloomsbury, W.C.
1898 Janson, Oliver J., Cestria, Clareiuont-road, Highgate, N.
1886 Jenner, James Herbert Augustus, East Gate House, T^ewes.
1899 Jennings, F. B., 152, Silver-street, Upper Edmonton, N.
1909 Jepson, Frank P., Department of AgricuUxix-e, Suva, Fiji Islands.
1886 John, Evan, Llantrisant, R.S.O., Glamorganshire.
h
( xvlii )
1907 Johnson, Charles Bh elding, Mayfield, Brinimjton Crescent, Stockport.
1889 Johnson, The Rev. W. F., M.A., Acton Rectory, Poyntz Fuss,
Co. Armagh.
1908 JoiCEY, Janies J., The Homestead, Bourne End, Bucks.
1888 Jones, Albert H., Treasurer, Shrublands, Elthum, Kent.
1910 Jones, Ernest P., 7, Shenoin-street, Nantwich-road, Creioe.
1894 t Jordan, Dr. K., Vice-President, The Museum, Trimj.
1910 Joseph, E. G., Lincoln College, Oxford.
1910 Joy, Ernest Cooper, 2, St. Kilda's-road, Stoke Newington, N.
1902 Joy, Norman H., M.R.C.S., L.R.C.P., Bradfield, Reading.
1884 Kane, W. F. de Vismes, ]\I.A., M.R.I.A., Drumleaske House,
Monaghan.
1884 Kappel, a. W., F.L.S., Linnean Society, Burlington House, W.
1876 f Kay, John Dunning, Leeds.
1896 f Kaye, William James, Caracas, Ditton Hill, Surbiton.
1907 Kelly, Albert Ernest McClure, Assistant Entomologist, Department
of Agriculture, Natal, S.A.
1902 Kemp, Stanley W., The Indian Museum, Calcutta.
1890 Kenrick, Sir George H., Whetstone, Somerset-road, Edghaston,
Birmingham.
1904 Kershaw, G. Bertram, Ingleside, West Wickham, Kent.
1901 Kershaw, John C., Ingleside, West Wickham, Kent.
1906 Keynes, John Neville, M.A., D.Sc., 6, Harvey-road, Cambridge.
1900 Keys, James H., Morwell, Freedom-villas, Lipson-road, Plymouth.
1889 King, J. J. F. X., Lecturer on Economic Entomology at the West of
Scotland Agricultural College, 1, Athole Gardens-terrace, Kelvin-
side, Glasgow.
1861 KiRBY,William F.,F.L.S.,77iWe», 46, Court-road,Chiswick,W .
1893 * Kirkaldy, George Willis, 2553, Punnin-avenue, Honohdu, Hawaii.
1889 Klapalek, Professor Franz, Karlin 263, Prague, Bohemia.
1887 f Klein, Sydney T., ICL.S., F.R.A.S., Haiherlow, Raglan-road,
Reigate.
1908 Knudsen, Jens Marius, Noerre Nebel, Denmark.
1910 Laidlaw, William, The Cedars, Romford, Essex.
1910 Lakin, C. Ernest, M.D., F. R.C.S., 2, Park-crescent, Portland-place, W.
1868 Lang, Colonel A. M., C. B., R.E., Bo.v Grove Lodge, Guildford.
1901 Lathy, Percy I., Fox Hall, Enfield.
1895 Latter, Oswald H., M.A., Charterhouse, Godaiming.
1908 Lawn, G. W., l\hdor House, Wealdestone, Harrow.
1899 Lea, Arthur M., Government Entomologist, Hobart, Tasmania.
1901 Leigh, George F., 45, Cuthbert’s Buildings, West-street, Durban,
Natal.
1910 Leigh, H. S., The University, Manchester.
1909 Leigh-Clare, Reginald L., c/o Messrs. Allen A Gledhill, Solicitors,
Singapore.
{ xix )
1892 Leslie, J. Henry, 84, IlTiron-road, Tooting Common, S.W.
1898 Lethbridge, Ambrose G., Guards Club, Pall Mall, S.W.
1903 t Levett, The Rev. Thomas Prinsep, Frenchgate, Richmond, Yorl-s.
1898 Lewis, E. J., F.L.S., Stainmore,'’ South Rill, Bromley, Kent.
1876 Lewis, George, F.L.S., 87, Frant-road, T'tinbridge Wells.
1908 t Lewis, John Spedan, Spedan Totoers, Hampstead, N.W., and 277,
Oxford-street, W.
1892 Lightfoot, R. M., Bree-st., Cajje Toivn, Cape of Good Hope.
1908 Lister, W. K., Street End House, Ash, near Dover.
1903 Littler, Frank M., Box 114, P.O., Launceston, Tasmania.
1865 f Llewelyn, Sir John Talbot Dillwyn, Bart., M.A., F.L.S. ,
Peullergare, Swansea.
1881 f Lloyd, Alfred, F.C.S., The Dome, Bognor.
1885 f Lloyd, Robert Wylie, 1, 5 and 6, The Albany, Piccadilly, W.
1903 Lofthouse, Thomas Ashton, The Croft, Linthorpe, Middlesbrough.
1908 Longsdon, D., 20, Holland-park, W.
1904 1 Longstaff, George Blundell, M.D., Highlands, Putney Heath,
S.W.
1899 Lounsbdry, Charles P., B.Sc., Government Entomologist, Cape
Town, S. Africa.
1894 Lowe, The Rev. Frank E., M.A., St. Stephens Vicarage, Guernseg.
1893 Lower, Oswald B., St. Oswalds, Bartley Crescent, Wagville, South
Australia.
1901 Lower, Rupert S., Davonpovt-terrace, Wayville, South Anstrcdia.
1909 Lucas, Dr. T. P., Wakejield-buildings, Adelaide-street, Brisbane,
Australia.
1898 Lucas, William John, B.A., 28, Knight’s Park, Kingston-on-Tharnes.
1904 * Luff, W. A., La Chaumih'e, Brock-road, Guernsey.
1880 Lupton, Henry, Devonia, Torquay.
1903 Lyell, G., Junr., Gisborne, Victoria, Australicc.
1901 Lyman, Henry H., M.A., F.R.G.S., 74, McTavish-street, Montreal,
Canada.
1909 Lyon, Francis Hamilton, Addlestone, Surrexj.
1906 McCarrison, D. L., Indian Police Forces, Madras Clxih, Madras.
1887 M‘Dougall, James Thomas, Dunollg, Morden-road, Blaclcheath, S.E.
1910 Macdougall, R. Stewart, M.A., D.Sc., F.R.S.E., Edinbxirgh Uni¬
versity.
1888 Mackinnon, P. W., Lgnndale, Mussoorie, N. W.P., India.
1900 Mackwood, The Hon. F. M., M.L.C., Colombo, Ceylon.
1899 t Main, Hugh, B.Sc., Almondale, Buckingham-road, South Woodford,
N.E.
1905 Mally, Charles Win., M.Sc., Graham’s Town, Cape Colony.
1887 Mander.s, Lieut. -Colonel Neville, R.A.M.C., F.Z.S., c/o Sir C.
McGrigor, 25, Charles- street, St. James’s- square, S.W.
1892 Mansbridge, William, 4, Norxvich-road, Waver tree, Liverpool.
1894 f Marshall, Alick, Auchinraith, Bexley, S.O., Kent.
( XX )
1895 Marshall, Guy Anstruther Knox, F.Z.S., 6, Chester-place, Hyde
Park-square, W.
1896 Marshall, P., M.A., B.Sc., F.G.S., University School of Mines,
Dunedin, Neiv Zealand.
1856 f Marshall, William, V.M.H., F.R.H.S., Auchinraith, Bexley, S.O.,
Kent.
1897 Martineau, Alfred H., Solihull, Birmingham.
1910 t Mason, C. W., St. Denis, Shaftesbury, Dorset.
1895 jMasset, Herbert, Ivy-Lea, Burnage, Didshnry, Manchester.
1865 Mathew, Gervase F., F.L.S., Paymaster-in-chief, R.N., Lee House,
Dovercourt, Harwich.
1887 Matthews, Coryndon, Stentaivay, Plymstock, S. Devon.
1900 Maxwell-Leproy, H., Imperial College of Science and Technology,
South Kensington, S.W.
1899 May, Harry Haden, Blackfriars’ House, Plymouth.
1904 Meade-Waldo, Geoffrey, Hever Warren, Edenhridge, Kent, and
British Museum {Natural History), Cromwell-road, S.W.
1872 f Meldola, Professor Raphael, F.R.S., F.C.S., 6, Brunswick-square,
W.C.
1885 Melvill, James Cosmo, M. A., F.L.S., Meole Brace Hall, Shretcsbury.
1887 Merrifield, Frederic, 14, Clifton-terruce, Brighton.
190G Merriman, Gordon, The Quick Laboratory, New Museums, Cambridge.
1905 Merry, Rev. W. Mansell, M.A., St. MichaeVs, Linton-road, Oxford.
1888 Meyer-Darcis, G., 5, Viide Poggio Imperiale, Florence.
1880 Meyrick, Edward, B.A., F.Z.S., F.R.S., Thornhanger, Marlborough.
1894 Miall, Louis Compton, F.R.S., Norton Way, N. LetcJnvorth.
1908 Middleton, Ivan E., 11, High-street, Serampore, Bengal.
1883 Miles, W. H., The New Club, Calcutta.
1910 Millar, F. Graliame, The Tang^a Batu Rubber Co., Malacca,
Straits Settlement.
1906 Mitchell-Hedges, Frederic Albert, Denbigh House, 25, Chepstow-
villas, Baystvater, W.
1905 Mitford, Robert Sidney, C.B., 35, Redcliffe Square, S.W.
1896 Moberly, J. C., j\I.A., Woodlands, Bassett, Southampton.
1879 Monteiro, Dr. Antonio Augusto de Carvalho, 70, Rua do Alecrinar,
Lisbon.
1902 Montgomery, Arthur Meadows, 34, Shalimar Gardens, Pembridge-
road. North Acton, W.
1899 ]\Ioore, Harry, 12, Loioer-road, Rotherhithe.
1907 Moore, Mrs. Catharine Maria, Ilolmeficld, Oakholme-rd., Sheffield.
1886 Morgan, A. C. F., F.L.S., 135, Oakwood-court, Kensington, W.
1889 f Morice, The Rev. F. D., M.A., Fellow of Queen’s College, Oxford,
Brunsivich, Mount Hermon, Woking.
1895 f Morley, Claude, The Hill House, Monk’s Soham, Suffolk.
1910 Morner, Count Birger, Consul for H.M. the King of Sweden,
Sydney, Australia.
1907 Mortimer, Charles H., Royton Chase, Byfleet, Surreij.
( xxi )
1893 Morton, Kenneth J., 13, Blackford-road, Edinburgh.
1910 Mosely, Martin E., 13, Addison-road, W.
1900 Moser, Julius, 59, Bulow-strasse, Berlin.
1882 Mosley, S. L., The Museum and Technical College, Huddersfield.
1907 Moulton, John C., The Hall, Bradford-on- Avon, Wilts., and The
Museum, Harawak.
1901 t Muir, Frederick, H.S.P. A. Experiment Station, Honolulu, Oahu,H.T.
1869 1 Muller, Albert, F.K.G.S., c/o Herr A. Miiller-Mechel, Gren-
zacherstrasse, 60, Basle, Switzerland.
1906 Muschamp, Percy A. H., Institut, Stafa, nr. Zurich, Sivitzerland.
1909 Musham, John F., 53, Brook-street, Selby, Yorks.
1903 Neave, S. a., B.A., Mill Green Park, Ingatestone.
1901 Nevinson, E. B., Morland, Cobham, Surrey.
1907 Newman, Leonard Woods, Bexley, Kent.
1909 Newstead, Alfred, The Grosvenor 3Iuseum, Chester.
1890 Newstead, R., Johnston Tropical Laboratory, University, Liverpool.
1909 Nicholson, Gilbert, M.A., M.D., University College Hospital, W.C.
1886 Nicholson, William E., School Hill, Lewes.
1906 Nix, John Ashburner, Tilgate, Crawley, Sussex.
1878 Nottidge, Thomas, Ashford, Kent.
1895 Nurse, Lt.-Colonel C. G., Timworth Hall, Bury St. Edmunds.
1908 Nurse, H. A., Botanical Department, Trinidad, B.W.I.
1877 Oberthur, Rene, Rennes {Ille-et-Vilaine), France.
1893 f Ogle, Bertram S., Steejile Aston, Oxfordshire.
1910 Oldaker, Francis A., M.A., The Red House, Haslemere.
1873 Olivier, Ernest, Ramillons, pres Moulins {Allier), France.
1895 Page, Herbert E., Bertrose, Gellatlg-road, St. Catherine's Park, S.E.
1907 Pead, Clement H., Box 252, Bulawayo, South Africa.
1883 P^ringuey, Dr. Louis, South African Museum, Cape Toion, South
Africa.
1903 t Perkins, R. C. L., M.A., D.Sc., F.Z.S., Park Hill House, Paignton,
Devon, and Board of Agriculture, Division of Entomology,
Honolid'u, Hawaii.
1879 Perkins, Vincent Robert, Wotton-under-Edge.
1907 t Perrins, J. A. D., 3rd Seaforth Highlanders, Fort George, N.B.
1897 Phillips, Hubert C., M.R.C.S., L.S.A., 262, Gloucester-terrace, Hyde-
park, W.
1903 t Phillips, Montagu A., F.R.G.S., F.Z.S., 22, Petherton-road, High¬
bury, New Park, N.
1901 Pickett, C. P., 28, Colwith-road, Leytonstone, S.E.
1891 Pierce, Frank Nelson, 1, The Elms, Dingle, IJverpool.
1901 * PiFFARD, Albert, Felden, Boxmoor, Hemel Hempstead.
1903 Pilcher, Colonel Jesse George, I.M.S., F.R.C.S., 133, Gloucester-
road, Kensington, S.W.
1910 PiLLAi, A. Raman, The University, Edinburgh.
( xxii )
1885 Poll, J. R. H. Neerwort van cle, Driehergen, Netherlands.
1870 f PoRRiTT, Geo. T., F.L.S., Mm Lea, Dalton, Huddersfield.
1884 f PouLTON, Professor Edward B., D.Sc., M.A., F.R.S., F.L.S., F.G.S.,
F.Z.S., Hope Professor of Zoology in the University of Oxford,
Wylceham House, Banhury-road, Oxford.
1905 Powell, Harold, 7, Rue Mireille, Hyeres (Var), France.
1906 Pratt, H. C., Government Entomologist, Federated Malay States,
Kuala Lumpur, Malay States.
1908 Pratt, William B., 10, Lion Gate Gardens, Richmond, Surrey.
1878 Price, David, 48, West-street, Horsham.
1908 Prideaux, Robert M., Woodlands, Brasted Chart, Sevenoaks.
1904 Priske, Richard A. R., 9, Melbourne Avetiue, West Ealing.
1893 Prout, Louis Beethoven, 62, Graham-road, Dalston, N.E.
1910 PuNNETT, Professor Reginald Crundall, M.A., Gains College,
Cambridge.
1900 Rainbow, William J., The. Australian Museum, Sydney, N.S. If '.
1907 Rayward, Arthur Leslie, 3, Albert Mansions, Lansdowne Road,
Croydon.
1874 Reed, Dr. E. C., Director del Museo de Concepcion, Concepcion,
Chile.
1893 Beid, Captain Savile G., late R.E., The Elms, Yalding, Maidstone.
1898 Relton, R. H., c/o Perkins and Co., Ltd., Brisbane, Queensland.
1898 Reuter, Professor Enzio, Helsingfors, Finland.
1910 de RhjS-Philips, George, Examiner of Accounts, Eastern Bengal
State Railicay, Calcutta.
1894 Riding, William Steer, B.A., M.D., Standlands, BuckereU, Honiton.
1908 Rippon, Claude, M.A., Spring^vood, Abingdon, Berks.
1905 Robinson, Herbert C., Curator of State Museum, Kuala Lumpur,
Selangor.
1892 Robin.son, Sydney C., 10, Inchmory-road , Gatford, S.E.
1869 1 Robinson-Douglas, William Douglas, M.A., F.L.S., F.R.G.S.,
Orchurdton, Castle Douglas.
1908 Rogers, The Rev. K. St. Aubyn, M.A., Rabai, Mombasa, British
East Africa.
1909 Rollason, Wm. Alfred, Lamorna, Triiro, Cornwall.
1886 Rose, Arthur J., Glanmor, Berkhamstead.
1907 Rosenberg, W. F. H., 57, Haver stock-hill, N.W.
1868 Rothney, George Alexander James, Pembury, Tudor-road, Upjyer
Norwood, S.E.
1894 f Rothschild, The Honble. Nathaniel Charles, M.A., F.L.S., F.Z.S.,
Arundel-house, Kensington Palace Gardens, W.
1888 f Rothschild, The Honble. Walter, D.Sc., F.L.S., F.Z.S., Zoological
Museum, 'Bring.
1890 Routledge, G. B., Tarn Lodge, Heads Nook, Carlisle.
1887 Rowland-Brown, Henry, M.A., Vice President and Secretary
Oxhey-grove, Harroio Weald.
1910 Rudge, Miss Carlotta, 1, Hamilton House, Grove-end-road, St.
John's Wood, N.W.
( xxiii )
1910 Rudge, Charles Henry, 1, Hamilton House, Grove-end-road, St.
John's Wood, N.W.
1898 Russell, A., IVilverley, Dale-road, Parley .
1892 Russell, S, G. C., 19, Lombard-street, E.C.
1899 Ryles, William E., B.A., 14, Arthur-street, Nottingham.
1905 St. Quintin, W. H., Seampton Hall, Pdllington, York.
1906 Sampson, Colonel E. Winns, 20 Arundel Mansions, S.W., and Junior
Carlton Club, Pall Mall, S.W.
1865* Saunders Edward, F.R.S., F.L.S., St. Ann’s, Mount Hermon,Woking.
1861 * Saunders, G. S., F.h.S., Littlethorpe, Furze Hill, Burgh Heath, Surrey.
1910 Saunders, H. A., Brookfield-house, Swanage.
1886 Saunders, Prof. Wm., Central Experimental Farm, Ottawa, Canada.
1901 ScHAUS, W., F.Z.S., 97, Elm Park Gardens, S.W.
1907 ScHMASSMAN, W., Beulah Lodge, London-r< ad, Enfield, N.
1881 ScoLLlCK, A. J., 8, Mayfield-road, Merton Park, Whnbledon.
1909 Scott, Hngh^ B.A., University Museum of Zoology, Cambridge.
1862 Sharp, David, M.A., M. B., F.R.S., F. L.S., F.Z.S., Lawnside,
Brockenhurst , Hants. ; and University Jduseum of Zoology and
Comparative Anatomy, Cambridge.
1902 Sharp, W. E., 9, Queen’s-road, South Norwood, S.E.
1886 Shaw, George T. (Librarian of the Liverpool Free Public Library),
]]'illiam Brown-street, Livergjool.
1905 Sheldon, W. George, Youlgreave, South Croydon.
1901 Shelford, Robert, M.A., F.Z.S., University Museum {Hope
Department), Oxford.
1900 t Shepheard-Walwyn, H. W., M.A., Dahvhinnie, Kenley, Surrey.
1887 Sigh, Alfred, Corney House, Chiswick, W.
] 909 SiLVERLOCK, Oscar C., c/o P. M. O. Livingstone, Esq., N. W. Rhodesia,
S. Africa.
1904 Simmonds, Hubert W., c/o Messrs. Kircaldie and Starnes, Limited,
Wellington, Neiv Zealand.
1902 Sladen, Frederick William Lambart, The Firs, Ripple, Dover.
1904 Slipper, The Rev. T. J. R. A., i\I.A., Tivetshall Rectory, Nortvich.
1902 Sloper, Gerard Orby, F.Z.S., J.P., Badminton Club, Piccadilly, W.
1907 Sly, Harold Baker, Mapledecm, Ringley -avenue, Horley.
1906 Smallman, Raleigh S., Homeside, Devonshire Park, Eastbourne.
1901 Smith, Arthur, County Mitseum, Lincoln.
1898 Sopp, Erasmus John Burgess, F.R.Met.S., 9, Merridale-road,
Wolverhampton.
1885 South, Richard, 96, Drakefield-road , Upper Tooting, S.W.
1908 Speyer, Edward R., Ridgehurst, Shenley, Herts.
1889 Standen, Richard S., F. L.S., Townlands, Lindfield, Sicssex.
1910 Stanley, The Rev. Hubert George, Marshfield Vicarage, Cardiff.
1898 Stares, C. L. B., M.R.C.S., L.R.C.P., The Limes, Swanley Junction,
Kent.
1898 Stebbing, Henry, Norham House, Sheen-road, Richmond, Surrey.
1910 STENTON,fRupert, Southivell, Notts.
( xxiv )
1910 Stoneham, Hugh Frederick, Lieut. E. Surrey Regt., Kingswear,
Streatham-pai'k, S.W.
1896 Stricici.and, T. A. Gerald.
1900 Studd, E. a. C., Kerremens, British Columbia.
1890 Studd, E. F., M.A., B.C.L., Oxton, Exeter.
1882 SwANZY, Francis, Stanley House, Granville-road, Sereuoals.
1908 SwiERsTRAY, Cominr. T., 1st Assistant, Transvaal Museum, Pretoria.
1884 SwiNHOE, Colonel Cliarles, M.A., F.L.S., F.Z.S., 6, Ounterstone-
roa'l, Kensington, W.
1894 SwiNHOE, Ernest, G, Gunterstone-rowl , Kensington, W.
1876 SwiNTON, A. H., c/o Mrs. Callander, Vineyard, Totnes.
1910 Tait, Robt., junr., Roseneath, Har ho rough-road, Ashton-on- Mersey.
1908 Talbot, G., 11, Palace Gardens, Enfield.
1893 Taylor, Charles B., Gap, Lancaster County, Penn., U.8.A.
1903 Taylor, Thomas Harold, M. A., Yorkshire College, Leeds.
1910 Terry, Frank Wray, P.O. Box 411, Honolulu, Haivaii.
1909 Tetley, Alfred, M.A., 22, Aven^ie-road, Scarborough.
1910 Theobald, F. V., M.A., Agricultural College, Wye.
1901 Thomp.son, Matthew Lawson, 40, Gosford -street, Middleshorough.
1892 Thornley, The Re\L A., M.A., F.L.S., “ Huyhenden,” Coppice-road,
Nottingham.
1907 Tillyard, R. J., B.A., The Grammar School, Sydney, New South
Wales.
1897 Tomlin, J. R. le B., M.A., Stoneley, Alexandra-road, Reading.
1907 Tonge, Alfred Ernest, Aincroft, Reigate, Surrey.
1907 Tragardh. Dr. Ivar, The University, Upsala, Sioeden.
1859 f Trimen, Roland, M.A., F.R.S.,F.L.S., Southbury, Lawn-road, Guild¬
ford.
1906 Tryhane, George E., Pedro Miguel Canal Zone, Panama.
1906 Tulloch, Captain James Bruce Gregorie, The King’s Own Yorkshire
Light Infantry; c/o Messrs. Cox & Co., 16, Charing Cross, S.W.
1895 Tunaley, Henry, 13, Begmead-avenue, Streatham, S.W.
1910 Turati, Conte Emilio, 4, Piazza S. Alessandro, Milan, Italy.
1898 Turner, A. J., M.D., Wickham Terrace, Brisbane, Australia.
1893 Turner, Henry Jerome, 98, Drakefell-road , St. Catherine's Park,
Hatch am, S.E.
1906 Turner, Rowland E., 21, EmperoTs Gate, S.W.
1894 Turner, Thomas, Cullompton, Devon.
1886 * Tutt, James W., Rayleigh Villa, Westcomhe Hill, S.E.
1893 Urich, Frederick William, C.M.Z.S., Port of Sjfain, Trinidad,
British HT.s^ Indies.
1904 t Vaughan, W., Badnlla, India.
1866 Verrall, George Henry, Sussex Lodge, Newmarket.
1909 VlDLER, Leopold A., The Carmelite Stone House, Rye, Sussex.
( XXV )
1895 Wacher, Sidney, F.R.C.S., Dane John, Canterhuri/.
1901 Waddington, Jolin, Park Holme, Harehill-avenue, Leeds.
1899 Wade, Albert, 12, Cadogan-place, Preston, Lancashire.
1897 Wainwright, Colbran J., 45, Ilandsworth ]Vood-road, Handsworth,
Birmingham.
1878 Walker, James J., M.A., R.N., F.L.S., Secretary, Aorangi,
Lonsdale-road, Summertoicn, Oxford.
1863 f Wallace, Alfred Russel, O.M., D.C.L., Oxon., F.R.S., F.L.S.,
F.Z.S., Broadstone, Wimborne, Dorset.
1866 f Walsingham, The Right Honble. Lord, Natural History Museum,
Cromwell -road, S.W.
1910 Ward, John J., Rusinurbe House, Somerset-road, Coventry.
1908 Warren, Brisbane 0. S., The Avenue, Amersham.
1886 Warren, Win., M.A., 33, Western-road, Tring, Herts.
1869 Waterhouse, Cliarles 0., Vice-President, Ingleside, Avenue-
gardens, Acton, W. ; and British Museum (Natural History),
Crom well-road, S.W.
1901 Waterhouse, Gustaviis A., B.Sc., F.C.S., Boycd Mint, Sydney, New
South Wales, Australia.
1904 AVatson, The Rev. N. Beresford, St. Martins Vicarage, St. Philip,
Barbados, W. Indies.
1893 AVebb, John Cooper, 218, Upland-road, Dulwich, S.E.
1908 AVellman, F. Creighton, M. D., U.S. Museum, \]'cishington, U.S..i.
1876 f AVestern, E. Young, 36, Lancaster Gate, Hyde Park, W.
1886 AA^heeler, Francis D., M.A., LL.D., Paragon House School, Norwich.
1906 AA^heeler, Tlie Rev. George, M.A., 37, Glo'ucester-place, AV.
1910 AVhite, Edward Barton, M.R.C.S., Cardiff City Mental Hospital,
Cardiff.
1907 AA'hite, Harold J., 42, Nevern-sq., Kensington, S.W.
1906 AA’ickwar, Oswin S., Charlemont, Gregnry-road, Colombo, Ceylon.
1903 AA^iggins, Clare A., M.R.C.S., Entebbe, Uganda.
1896 AVileman, A. E., H.B.M. Consul, Manilla, Philijjpine Islands.
1910 WiLLCOCKS, Frank C., Entomologist to the Khedivial Agricultural
Society, Cairo, Egypt.
1910 AVilliams, James M., Howard Motor Garage, Brithdir, Cardiff and
Canford Cliffs, Hants.
1894 AVolley-Dod, F. H., Millarville P. 0., Alberta, N.W.T., Canada.
1900 AVood, H., Kennington, near Ashford, Kent.
1881 Wood, The Rev. Theodore, The Vicarage, Lyford-road, Wandsworth
Common, S.N.
1905 WooDBRiDGE,Francis Charles,27ie Briars, Gerrard’s Cross, S.O., Bucks.
1888 Yerbury, Colonel John AV., late R.A., F.Z.S., Army and Navy
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1892 Youdale, AVilliam Henry, F.R. Al.S.,21, Belle-Isle-street, Workington.
( xxvi )
ADDITIONS TO THE LIBRARY
Dubinq the Yeak 1910.
Ahlwakth (K.). [See Coleopterorum Catalogus.]
Ainslie (G. G.)- The Cowpea curculio.
[U. S. Dept. Agric., Bureau Entom., Bull. No. 85, Pt. 8, 19^0.]
- - The Larger Coru-Stalk Borer (Diatroea sacchamlis. Fab.).
[U. S. Dept. Agric., Bureau Entom., Circular No. 116, 1910.]
U. S. Dept. Ayrir.
Akrow(G. J.). Fauna of British India. Coleoptera. Lamellicoruia(Cetoniinae
and Dyuastinae). 1910. The India Office.
Back (E. A.). The Woolly White-fly {Aleyrodes howardi, Quaint.).
[U. S. Dept. Agric., Bureau Entom., Bull. No. 64, Pt. 8, 1910.].
U. S. Dept. Ayric.
B.^gnai.l (R. S.). Contribution towards a knowledge of the Neotropical
Thysaiioptera.
[Journ. Linn. Soc., Zool., Vol. XXX, 1910.] Lord Avehury.
Banks (Nathan). Catalogue of Nearctic Spiders.
[U. S. Nat. Mus., Bull. No. 72, 1910.]
The Smithsonian Institution.
Bahgagli (Piero). L’Opera degli iusetti nella formazione dell’ Humus.
[Atti R. Accad. Georgofili, Vol. VII, 1910.] The Author.
Beahe (Prof. T. Hudson). Retrospect of a Coleopterist for 1909.]
[Entom. Record, Vol. XXII, No. 1, 1910]. The Author.
- and Evans (W.). Coleoptera from Mole’s Nests in the South-East
of Scotland.
[Ann. Scott. Nat. Hist. 1909.] The Authors.
Bericht iiber die wisseuschaftlichen Leistungen im Gebiete der Entomologie
wahrend des Jahres 1905 imd 1906. Published in 1910. By
K. Griiuberg, R. Lucas, H Schouteden, and G. Seidlitz.
Purchased.
Bernhauer (M.). [See Coleopterorum Catalogus.]
Bern.stein (Dr. Julius). Summary of literature relating to the bionomics
of the parasitic fungus of flies {Empusa muscae).
[Reports to L.G.B. on Public Health and Medical Subjects, N. S.
No. 40, 1910.] H.JI. Stationery Office.
Bickhardt (H.). [See Coleopterorum Catalogus.]
Blatchley (W. S.). On the Aleocharinae known to occur in Indiana.
[Bull. Indiana Dept. Geology and Natural Resources, No. 1, 1910.]
A. Fenyes.
Brown (H. Rowland-). Notes on a Butterfly Hunt in France, 1910.
[Entomologist, Nov. and Dec. 1910.] The Author.
Browne (H. B.). Descriptive Catalogue of the Dobree Collection of
European Noctuae, 1909. The. Author.
( xxvii )
Burr (Malcolm). A Preliminary Revision of the Labiduridae, a Family of
Dermaptera.
[Trans. Entom. Soc. 1910.]
- Dermaptera (The Percy Sladen Trust Expedition to th'^ Indian
Ocean in 1905).
[Trans. Linn. Soc., Zool., Vol. XIV, Pt. 1, 1910.] The Author.
- The Fauna of British India, including Ceylon and Burma. Derma¬
ptera (Earwigs). The India Office.
Busck (August). New Moths of the Genus Trichostihas,
[Proc. U. S. Nat. Mus., Vol. XXXVIll, 1910].
- Notes on a Horn-feeding Lepidopterous larva from Africa.
[Smithsonian Misc. Coll., Vol. LVI, No. 8, 1910.]
The Smithsonian Institution.
Carpenter (G. H.). Injurious Insects and other Animals observed in
Ireland during the year 1909.
[Econ. Proc. Royal Dublin Soc. 1910.] The Author.
Casey (Thos. L.). Memoirs on the Coleoptera, I, 1910. [Staphylinidae.]
Lancaster, Pa. The Author.
Champion (G. C.). [See Godman (F. D.), Biologia Central!- Americana.]
- and Lloyd (R. W. ). Some Interesting British Insects, I-III.
[Ent. Monthly Mag., 1909, 1910.] The Authors.
Chittenden (F. H.). Notes on the Cucumber Beetles.
[U. S. Dept. Agric., Bureau Entom., Bull. No. 82, Pt. 6, 1910.]
- The Oak Primer.
[U. S. Dept. Agric. , Bureau Entom., Circular No. 130, 1910.
U. S. Dept. Agric.
Cockerell (T. D. A.). Some Bees of the Genus Augochlora from the West
Indies.
[Proc. U. S. Nat. Mus., Vol. XXXVII, 1910.]
- North American Bees of the Genus Nomia.
[Proc. U. S. Nat. Mus., Vol. XXXVIll, 1910.]
The Smithsonian Institution.
Coleopterorum Catalogus. Junk (W.) editus a Schenkling (S.). Berlin,
1910-11.
Pars 9. Olivier (E.). Lampyridae.
„ 10. Olivier (E.). Rhagophthalmidae, Drilidae.
„ 11. Leveille (A.). Temnochilidae.
„ 12. Csiki (E.). Endomychidae.
„ 13. Csiki (E.). Scaphidiidae.
„ 14. Pic (M.). Hylophilidae.
„ 15. Gebien (H.). Tenebrionidae, I.
„ 16. Pape (P.). Brachyceridae.
„ 17. Zaitzer (Ph.). Dryopidae, Cyathoceridae, Georyssidae
Heteroceridae.
,, 18. Csiki (E.). Platypsyllidae, Orthoperidae, Phaenocejihalidae
Discolomidae, Sphaeriidae.
„ 19. Bernhauer (M.) et Schubert (K.). Staphylinidae, I.
,, 20. Schmidt (A.). Aphodiinae.
., 21. Ahlwarth (K.). Gyrinidae.
„ 22. Gebien (H.) Tenebrionidae, II.
„ 23. Schenkling (S.). Cleridae.
„ 24. Bickhardt (H). Histeridae.
„ 25. Dalla Torre (K. W. von). Cebrionidae.
„ 26. Pic (M.). Scraptiidae, Pedilidae. Purchased.
Copeman (Dr. S. Monckton). Note as to work in hand, but not yet pub¬
lished, and as to proposed further work in reference to Flies as
Carriers of Infection.
[Reports to L.G.B. on Public Health and Medical Subjects, N. S.,
No. 40, 1910.]
H.M. Stationery Office.
( xxviii )
CoQUiLLETT (D. W.). Type-species of the North American Genera of
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[Proc. U. S. Nat. Mus., Vol. XXXVII, 1910.]
The Smithsonian Institution.
Crawford (.J. C.). New Hymenoptera from the Philippine I.slands.
[Proc. U. S. Nat. Mus., Vol. XXXVIII, 1910.]
- Three new Genera and Specie.s of Para.sitic Hymenoptei’a.
[Proc. U. S. Nat. Mus., Vol. XXXVIII, 1910.]
The Smithsonian Institution.
CsiKi (B.). [See Coleopterorum Catalogus.]
Dalla Torre (K. W. von). [See Coleopterorum Catalogus.]
Dean (W. Harper). The Sorghum Midge (Contarinia [^Di^ilosis"] sorahirota,
Coq.).
[U. S. Dept. Agric., Bureau Entom., Bull. No. 85, Pt. 4, 1910.]
U. S. Dept. Ayric.
Distant (W. L ). Rhynchota: Heteroptera. Appendi.K. Fauna of British
India, Vol. V. London, 1910. The India Offic'’.
- - Fauna Transvaaliensia. Part X, 1910. Purchased.
Dognin (Paul). Heteroceres nouveaux de I’Anierique du snd. Fasc. 1, 1910.
The Author.
Dyar (H. G.). Descriptions of some new Species and Genera of Lepido-
ptera from Mexico.
[Proc. U. S. Nat. Mus. Vol. XXXVIII, 1910.]
The Smithsonian Institution.
Evans (W.). [See Beare (T. Hudson)].
Fouel (A). Fourmis des Philippines.
[Philippine Journ. Science, July, 1910.]
- Note sur Quelques Fourmis d’Afrique.
[Ann. de la Soc. Entom. de Belgique, 1910 ] The Author.
Foster (S. W.). On the Nut-feeding Habits of the Codling Moth.
[U. S. Dept. Agric., Bureau Entom., Bull. No. 80, Pt. 5, 1910.]
- and Jones (P. E.). How to Control the Pear Thrips.
[U. S. Dept. Agric., Bureau Entom., Circular No. 131, 1910.]
U. S, l>ept. Ayric.
Gadeau (Henri). Voyage Zoologique en Khroumirie (Tunisie), Mai-Juin,
1906. Paris, 1908. Dr. 31. Burr.
Gerien (H. ). [See Coleopterorum Catalogus.]
Ghosh (Cham C.). Insects of (on) Crops. Written on the basis of “ Indian
Insect Pests ” of H. Maxwell Lefroy and D. N. Pal. Pusa, 1909.
(Written in Bengali, with notes in English.) The Author.
Godman (F. D.). Biologia Centrali-Americana, Pts. CCVII, CCVIII, 1910 :
Diplopoda, by R. I. Pocock ; Insecta, by G. C. Champion and
Lord Walsingham. The Editor.
Gold Coa.st, Government of the. Report of the Agricultural Department for
the year 1909. Gold Coast, 1910. The Director.
Graham-Smith (Dr.). Observations on the ways in which artificially infected
flies {3Iusca domestica) carry and distribute pathogenic and
bacteria.
[Reports to L.G.B. on Public Health and Medical Subjects, N. S.,
No. 40, 1910.] H.3I. Stationery Office.
Green (E. Ernest). Coccidae of Ceylon, Pt. IV. London, 1909.
- Entomological Notes.
[Tropical Agriculturist, Vols. XXXIV, XXXV, 1910; Spolia Zeyl.,
Vol. VII, 1910.]
- On some Coccid Pests of Economic Importance.
[Journ. Economic Biology, Vol. V, Pt. 1, 1910.] 7'he Author.
( xxix )
Grossbeck (J. a.). Studies of the North American Geometrid Moths of the
Genus Pero.
[Proc. U. S. Nat. Mus., Vol. XXXVIII, 1910.]
The Smithsonian Institution.
Hagedorn (M.). [See Wytsman’s Genera Iiisectorum.]
Hammar (A. G. ). Life history of the Codling Moth in North-Western
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[U. S. Dept. Agric., Bureau Entom., Bull. No. 80, Pt. 6, 1910.]
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Hampson (Sir G. F.). Catalogue of the Lepidoptera Phalaenae. Vol. IX
(text and plates), 1910. By E.xchanye.
Hofeneder (Karl). Mengenilla, n. g., chohautii, n, sp. Eiue neue
Strepsiptere aus Nordafrika.
[Ber. naturw.-med. Vereins, Jahr. XXXI, 1910.] The Author.
Hope Reports. Vol. VII, 1908-10. Edited by Prof. E. B. Poulton, 1910.
Hope Dept., Oxford Univ. Mus.
Hopkins (A. D.). Insects which kill forest trees.
[U. S. Dept. Agric., Bureau Entom., Circular No. 125, 1 910.]
- Insect injuries to the wood of Dying and Dead Trees.
[U. S. Dept. Agric., Bureau Entom., Circular No. 127, 1910.]
- Insect Injuries to Forest Products.
[U. S. Dept. Agric., Bureau Entom., Circular No. 128, 1910.]
- Insects in their relation to the reduction of future supplies of timber
and general principles of Control.
[U. S. Dept. Agric., Bureau Entom., Circular No. 129, 1910.]
- Contributions toward a monography of the Bark-weevils of the
Genus Fissodes.
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1910.] U. S. Dept. Agric.
Howard (L. O.). Parasites reared or supposed to have been reared from the
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Hunter (W. D.). The Status of the Cotton Boll Weevil in 1909.
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U. S. Dept. Agric.
Hyslop (J. a.). The Smoky Crane-Fly.
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U. S. Dept. Agric.
Je.annel (R.) Essai d’une uouvelle classification des Silphides Cavernicoles.
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- et Racovitza (E. G.). Enumeration des Grottes visitees 1908-09.
(S'"®- serie).
[Arch. Zool. Exper. Gen., Tom. V, 1910.] The Authors.
Johnson (F.) and Hammar (A. G.). The Grape root-worm.
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U. S. Dept. Agric.
Jones (P. R.). Tests of sprays against the European fruit Lecanium and
the European Pear scale.
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Junk (W.). [See Coleopterorum Catalogus.]
( XXX )
Kerremans (Ch.). Mouographie des Buprestides. Vol. IV, Livr. 3 to
completion. Vol. V, Livr. 1-8, 1910. Purchased,
Kertesz (C.). Catalogus Dipterorum.
Vol. II. Cecidomyiidae, Limuobiidae, Tipulidae, Cyliiidrotomidae,
1902.
Vol. III. Stratiomyiidae, Erinnidae, Coenomyiidae, Tabanidae,
Pautophthalmidae, Rliagionidae, 1908.
Vol. IV. Oncodidae, Nemestriiiidae, Mydaidae, Apioceridae, Asilidae,
1909.
Vol. VI. Empididae, Dolicbopodidae, Musidoridae, 1909.
Vol. VII. Syrphidae, Dorylaidae, Phoridae, Clythiidae, 1910.
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Saltatoria, Pt. 2 (Locu.stidae vel Acridiidae), 1910.
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Klapalek (F. ). Metamorphose der Trichopteren, 1888.
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Lefroy (H. Maxwell-). Life-historie.s of Indian Insects, Coleoptera I.
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Leon (N.). Studii asupra Culicidelor din Romania. Bucuresti, 1910.
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L6veille (A.). [See Coleopterorum Catalogus.]
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Ldndbeck (Wm.). Diptera Danica, Pt. Ill, 1910. The Author.
Marlatt (C. L.). The Horn Fly {Haematobia serrata, Rob.-Desv.)
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Marsh (H. O.). Biologic and Economic Notes on the Yellow- Bear Cater¬
pillar.
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- - — Biologic Notes on species of Diahrotica in Southern Texas.
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Martelli (G.). Parassiti indigeni ed esotici della “ Diaspis pentayona ”
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( xxxi )
OsHANiN (B.). Verzeichnis der Palaearktischen Hemipteren. Baud III.
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Schaeffer (C.). Additions to the Carabidae of North America, with notes
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ScHENKLTNG (S.). [See Coleopterorum Catalogus.]
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Snyder (T. E.). Damage to Chestnut Telephone and Telegraph Poles by
AVood-boring Insects.
[U. S. Dept. Agric., Bureau Entom., Bull. No. 94, Pt. 1, 1910.]
U. S. Dept. Agric.
Spuler (A.). Die Schmetterlinge Europas.
[4 vols.. Plates. Stuttgait, 1901-10.] Purchased.
Theob.vld (E. V.). Monograph of the Culicidae of the World. Vol. V,
1910. By E.rchange.
- South-Eastern Agricultural College, AVye.
[Report on Economic Zoology for year ending Sept. 30, 1910.
London, 1910.] The Author.
Turati (Emilio). La Zygaena transedpina, Esp. e le sue forme Italiane.
1910. The Author.
- Note Critiche sulla Pieris ergane, H. G.
[Atti Soc. Ital. Sci. Nat., Vol. XLIX, 1910.] The Author.
Tutt (J. W.). British Lepidoptera. A^ol. X, 1908-09. Purchased.
Vickery (R. A.). Contributions to a knowledge of the Corn Root- Aphis
{Aphis maidiradicis, Eorbes).
[U. S. Dept. Agric., Bureau Entom., Bull. No. 85, Pt. G, 1910.]
U. S. Depit. Agric.
Viereck (H. L.). Descriptions of new species of Ichneumon-flies.
[Proc. U. S. Nat. Mus., Vol. XXXVIII, 1910.]
The Smithsonian Institution.
Walker (James J.). Second Supplement to the Preliminary List of the
Coleoptera of the Oxford district, 1910. The Author.
xxxiii )
Walsingham (Lord). [See Godman (F. D.). Biologia Centrali-Americana.]
Webster (F. M.). The Clover Root-Borer {Hylastimis obscurus).
[U. S. Dept. Agric., Bureau Entom., Circular No. 119, 1910.]
U. S. Dept. Agric.
Webster (F. M.) and Reeves (G. I.). The Western Grass-stem Fly
{Cephus occiderdalis).
[U. S. Dept. Agric., Bureau Eutom., Circular No. 117, 1910.]
U. S. Dept. Agric.
WiLDEu.MUTH (V. L.). Clover-root Curculio (Sitones hispidultcs. Fab.).
[U. S. Dept. Agric., Bureau Entom., Bull. No. 85, Ft. 3, 1910.]
U. S. Dept. Agric.
WvTSMAN (P.). Genera Insectorum. Fasc. LXXX&, C-CXIIa, 1910.
E. A. Elliott.
Zaitzkr (Ph.). [See Coleopterorum Calatogus.]
C
( xxxiv )
Periodicals and Publications of Societies.
AMERICA (NORTH).
CANADA.
London, Ontario. The Canadian Entomologist. Vol. XLII, 1910.
By Exchange.
Ontario. Entomological Society of Ontario. 40th Annual Eeport. 1909.
By Exchange.
UNITED STATES.
New York. New York Entomological Society. Journal. Vol. XVIII, 1910.
Purchased.
Philadelphia. Academy of Natural Sciences of Philadelphia. Proceedings.
Vol. LI, Part 3, 1909, Vol. LII, Parts 1 and 2, 1910.
By Exchaiige.
Entomological News, Vol. XXI, 1910. By Exchange.
AYashington. Smithsonian Institution. Annual Report, 1909.
United States National Museum. Proceedings. Vol. XXXVII, 1910.
Proceedings of the Entomological Society of Washington. Vol.
XI, 1910. Purchased.
AMERICA (SOUTH).
BRAZIL.
Para. Boletim do Museu Goeldi. Vol. VI, 1909. The Museum.
Rio de Janeiro. Memorias do Instituto Oswaldo Cruz. Tom. II, fasc. 2,
1910. The Institute.
WEST INDIES.
Barbados. West Indian Bulletin. The Journal of the Imperial Agricul¬
tural Department for the West Indies. Vol. X, Parts 3 and 4;
Vol. XI, Part 1.
Agricultural News. Vol. IX, 1910.
The Agricultural Department.
ASIA.
INDIA.
Bombay. Natural History Society. Journal. Vol. XX, 1910.
By Exchange.
Colombo. Spolia Zeylaudica. Vols. I-VI, 1903-10. The Colombo Museum.
PusA. Report of the Agricultural Research Institute and College, Pusa,
1909-10. The India Office.
List of Names used in India for Common Insects. Agric. Research
Inst. Pusa, 1910. The India Office.
( XXXV )
AUSTRALASIA.
New Zeaeakd. New Zealand Institute. Transactions and Proceedings.
Vol. XLII, 1910. The. Institute.
Perth. Journal Agricultural Department of AVest Australia. Vol. XVIII,
1910. The Socieli/.
Sydney. Linueau Society of New South Wales. Proceedings, 1910.
Bi/ Exchav(je.
EUROPE.
AUSTPvI A-H UNGARY.
Budapest. ArcLivum Zoologicum. Vol. I, 1909-10.
Ldh. ZooJ. Honyroise.
Brunn. Verhand. der uaturf. Vereines in Briinn. Bd. XIWII.
By Exchanye.
Vienna. K.-k. zoologische-botanische Verein (Gesellschaft) in Wien.
Verhandlungen. Band LX, Heft. 1-8. By Exchanye.
AViener entomologische Zeitung. Bd. XXIX, 1910.
Purchased.
BELGIUM.
Brussels. Societe Entomologique de Belgique. Ann.ales. A''ol. LIA', 1910.
Meinoires. A'ol. XVIII, 1910. By Exchanye.
FRANCE.
Caen. Societe Fran9aise d’Entomologie. Revue. Tome XXA'III, Parts
3-C>, 1909-10. Purchased.
Ch.ateauroux. Le Frelon. XA'II, Parts 7-11, 1910. Purchased.
Paris. Soc. Entom. de France. Aunales, A'ols. LXXVIII, Parts 3 and 4 ;
LXXIX, Parts 1 and 2, 1910. By Exchanye.
Toulouse. Bulletin de la Soc. d’Hist. Nat. de Toulouse. Tome XLII, Parts
1-4, 19(M). By Exchanye.
GERMANY.
Berlin. Deutsche entomologische Gesellschaft. Deutsche entomologi.sehe
Zeitschrift. Bd. LIA', 1910. By E.rchanye.
Deutsche entomologische National-Bibliothek. Band I, 1910.
By E.rchanye.
Entomologischer A'erein in Berlin. Berliner entomologische
Zeitschrift, 1907-10. By Exchanye.
Dresden. “ Iris.” Deutsche entomologische Zeitschrift. Band XXIA'.
Beiheft 1, 1910. • By Exchanye.
Munich. Mitteilungen der Miinchuer entomologischen Gesellschaft. 1910.
The Puldisher.
Stettin. Entomologischer A^ereiu. Entomologische Zeitung. Jahr. LXXI
1910. By E.rchanye.
Wiesbaden. Nassauischer A'erein fiir Naturkunde. Jahrbiicher. Jahr-
I.XIII, 1910. By E.rchanye.
( XXX vi )
GREAT BRITAIN AND IRELAND.
Gi.asgow. The Glasgow Naturalist. Vol. I, 1908-09. The Publisher,
Liveri’OOi.. Lancashire and Cheshire Entomological Society. 03rd Annual
Report and Proceedings. 1910. The ISociety.
London. Annals and Magazine of Natural Historj’. 1910. Purchased,
Athenaeum. 1910. The Publishers.
Bulletin of Entomological Research. Part 1-4, 1910-11.
Purchased.
City of Loudon Entomological and Natural History Society. Trans¬
actions. 1909. 'The Society.
Entomologist (The). 1910. li. South.
Entomologist’s Monthly Magazine. 1910. The Editors.
Entomologist’s Record and Journal of V.ariatiou. Vol. XXII, 1910.
Purchased.
Liuneau Society of London. Zoology, Transactions, Journal and
Proceedings. 1910. Jiy Exchange.
Nature. 1910. The Publishers.
Quekett Microscopical Club. Journal. 1910. The Club.
Royal Agricultural Society. Journal. Vol. LXX, 1909.
'The Society.
Royal 31icroscopical Society. Journal. 1910. By Exchange.
Royal Society. Proceedings. 1910. By Exchange.
Royal Society. Philosopliical Transactions. 1910.
South Loudon Entomological and N. H. Society. Proceediu g.s.
1909-10. The Society.
Zoological Society. Transactions and Proceedings. 1910.
Jiy Exchange.
Zoologist (The). 1910. 'I'he Publisher.
Manchester. Manchester Entomological Society, Reports for 1909 and 1910.
'I'hc Society.
HOLLAND.
The Hague. Tijdschrift voor Entomologie. Jahr. 1910. Jiy Exchange.
Eutomologische Berichten. Vol. II, Parts 49-54. 1909-10.
Jiy Exchange.
ITALY.
PoHTici. Bollettino del Laboratorio di Zoologia Geuerale e Agraria. Vol.
IV, 1910. Jiy I-ixchange.
PoRTici and Florence. Redia, Giornale de Entomologia. Vol. VI,
P'asc. 2. 1910. Purchased.
PORTUGAL.
S. Fiei.. Broteria. Revista de Sciencias Naturaes <lo Gollegio de S. Fiel.
Vol. IX, 1910. Purchased.
( xxxvii )
RUSSIA.
-Mo.scow. Societe Impemle ties Nafcuralistes tie !Moscou. Biilletiu. 1909.
/>’// Exchaiiije.
St. Peterskckg. Aunuaire du Musee Zoologiijue. . Tome XV, I'ai-ts 1, 2,
1910. F. ]). Godinan.
Horati Societatis Eutomologicae Kos.sicae. Tom. XXXIX, 1910.
J)\>/ Exchaiifje.
Revue Russe il’Entomologie. 'I’ome IX, 1909.
“ Table.s gtiiierale.s ties publieatious ” ttf tlie Russian Knttnnological Society.
]So9-1908. 1910. Socift!/.
SWEDEN.
Stockholm. Arkiv for Zoologi. Vol. VI, 1910. dii/ ExchaiHfe.
Entomologisk Titlskrift. Arg. 31. 1910.
/>';/ Exchani/f.
SW1TZERL.\NI).
tiKNKV.v. .Societe tie Physique et tPHistoire Naturelle. Memoires. Vol.
XXXVII, Fasc. 2-4, 1910. />'y Exchaiuje.
.ScHAFFH.\usKX aiul liERXK. Scliweizprisclie entomolttgische Gesellschaft.
Mittheilungen. Bd. XII, Hft. 1, 1910.
/>’// Exchoiitie.
ERRATA.
TRANSACTIONS.
Page 3, line 13 from bottom, for scintilulla read scintillula.
Page 481, line 11 from tog, for A^itizera read Actizera.
Page 494, top line, for ^rtffcera read Actizera.
Also in legend of Plates LII, LIX, LX, /or Antizera read Actizera
PROCEEDINGS.
Page xxi, line 12 from top, for ariyustatus read angustatus.
THE
PROCEEDINGS
OF THE
ENTOMOLOGICAL SOCIETY
OF
LONDON
For the Year 1010.
Wednesday, February 2nd, 1910.
Dr. F. A. Dixby, M.A., M.D., F.E.S., President, in the Chair.
Nomination of Vice- Presidents .
The President announced that he had nominated as Vice-
Presidents for the Session 1910-1911 Professor T. Hudson
Beare, B.Sc., F.B.S.E., Mr. G. T. Bethune-Baker, F.Z.S.,
and Mr. H. Rowland-Brown, M.A.
Election of Fellows.
Mr. Edward Morrell Holmes, of Ruthven, Sevenoaks ;
Mr. E. G. Josephs, of Lincoln College, Oxfoi’d ; Mr. Ernest
Cooper Joy, of 2, St. Hilda’s Road, Stoke Newington, N. ]
Mr. John J. Ward, of Rusinurbe House, Somerset Road,
Coventry; and Mr. Frank C. Willcocks, Entomologist to
the Khedivial Agricultural Society, of Cairo, Egypt, were
elected Fellows of the Society.
Entomological Congi'ess, 1910.
Dr. M. Burr gave an account of the forthcoming Entomo¬
logical Congress to be held in August next, and appealed to
PROG. ENT. SOC. LOND., I., II. 1910.
A
( ii )
all Fellows for their support, as well as to the Local Natural
History Societies throughout the United Kingdom. He men¬
tioned also that the County Council of Essex were sending a
delegate, and he hoped that the other Counties would recognise
in the same way the value of the services to be rendered to
science by the Congress.
Mr. F. Merrifibld, as an official of the East and West Sussex
County Councils, said the subject Avoukl undoubtedly be con¬
sidered by those bodies, and probably by others interested,
and Dr. K. Jordan gave an outline of the programme of
papers already arranged to be delivered.
Entomological Society of Russia.
The Secretary said that a letter had been received from
the Entomological Society of Russia, inviting tbe Society to
send a delegate to the forthcoming Jubilee Celebration in
St. Petersbiirg.
The President having intimated that the Council Avould be
glad to receive the name of any Fellow able to undertake the
journey, on the motion of Mr. J. W. Tutt, seconded by Mr.
H. Rowland-Brown, it Avas resoh-ed unanimously to send a
letter of congratulation to the Society in honour of the
occasion.
Exhibitions.
Collecting in the Rhone Valley. — Mr. John Alderson,
Avho Avas present as a visitor, exhibited the j-esults of six
weeks’ collecting in the Rhone Valley, SAvitzerland, in May
and June last, representative in all of one hundred and tAvo
species of Rhopalocera.
New Earavig from Borneo. — Dr. K. Jordan exhibited
two specimens of the new kind of earAvig, Arixenia esau, lately
described by him in Nov. Zook, p. 313, PI. xvi-xviii (1909).
The insect was discoA^ered in the breast-pouch of a specimen
of the “naked bat” obtained in Sarawak. Under the micro¬
scope Avere shoAvn the mandible and maxilla of Arixenia,
together with the mandible of Ileinimerus.
A CONVENIENT METHOD OF STORING BUTTERFLIES IN PaPER
Envelopes. — Mr. E. E. Green sent for exhibition boxes
( iii )
to illustrate a convenient method of storing butterflies in
paper envelopes. This consists of a box made of tin-plate,
with partitions dividing it into trough-shaped spaces, as shown
in the accompanying figures. The triangular envelopes rest
edgeways in the troughs. The box is fitted with two lids —
above and below.
Fig. 1 shows a box with the upper lid removed and the
lower one in place. The box measures 9 in. by 6 in. by 3 in.
The upper space contains a single trough (a), and carries
envelopes with a base of 5| inches.
Fig. la represents the reverse of the same box, with
two smaller troughs (b, c) to contain envelopes of half the
size.
Fig. 2 shows a box of the same size, but designed for the
smaller-sized envelopes alone, and containing four troughs
K 9)-
A 2
( iv )
Fig. 3 is a larger box, of just double the depth of the others,
measuring 9 in, by 6 in. by 6 in., with a diagonal partition
forming a single trough above and below (/«, i), capable of
carrying envelopes of a larger size.
Fig. Irt.
(In Figs. Iffi, 2 and 3, the two lids have been omitted, for
sake of clearness.)
These boxes are designed for three sizes of envelopes, which
gives a sufficient range for butterflies of any size. Size 1 is
made from a rectangle 8 in. by 5 in. Size 2, from a rect¬
angle 6 in. by 4 in. Size 3, from a rectangle measuring 4 in,
by 2| in.
( V )
It is found, in practice, that a box made according to Fig. 1
will carry — without overcrowding — from 100 to 130 full
envelopes in the larger trough, and from 175 to 200 in each
of the two smaller spaces. Design 2 will hold, in each of the
four spaces, 225 Lycaenidae, making a total of 900 insects.
Design 3 will hold 75 or more filled envelopes in each of the
two spaces.
For convenience of examination, the insects should be
arranged in families : the genera alphabetically in each family.
Fig. 3.
and the species alphabetically in each genus. Subsequent
additions can be slipped into their places without disturbing
those already in position. To keep the envelopes in place,
when the troughs are only partly occupied, triangular blocks
of cork — about f in. thick — can , be employed. For use as
collecting boxes, the troughs can be charged with empty
envelopes, and the cork triangles will serve as markers to
separate off the unused envelopes as they are filled.
The boxes illustrated are of the simplest design — as made
by a local tinsmith in Ceylon. They can be improved by a
coating of black japan on the outside.
( Vi )
A narrow partition of perforated zinc, as in the one
executed by Messrs. Watkins and Doncaster, may be fixed at
one end of each trough, to contain a bag of naphthalin ; the
lids hinged, and fastened with a spring clasp.
Mr. Green in his letter commiTnicating the above said that
he had found that envelopes stored in this way occupy very
much less space than if placed haphazard in a plain box, and
that the insects are less liable to damage. The ease with
Avhich any individual specimen can be found instantly is a
convenience that cannot he overestimated. He had over 6000
butterflies stored in this manner, the collection occupying only
a dozen of these small boxes which, being of uniform size, can
be stacked in quite a limited space.
Discussion of the Affinities of Agriades thbtis
(bellargus) and a. coridon.
The discussion of the affinities of Agriades thetis {bellargus)
and A. coridon, adjourned from the December meeting, was
resumed by Mr. J. W. Tutt, who exhibited five cabinet drawers
of the two species demonstrating in particular the several
forms of A. coridon as occurring in the Palaearctic region. He
pointed out in some detail the nearness of the two Agriadid
species in the structure of their eggs, larvae, pupae, imagines,
especially noting in the latter case the similarity in the $
genitalia. Observing that it had generally been assumed that
closely-allied species had been maintained if not developed by
the setting up of some ellective boundary in their environ¬
ment or morphological structure, resulting in isolation, it was
remarkable that no such boundaries could be found in these
species. Not only was their environment similar, their
morphological structure almost identical, but their actual
range * was practically the same.
The facts of their distribution appeared capable of tabulation
as follows : —
Mr. Betliune-Baker has two undoubted S A. coridon in his collection
sent from America, and supposed to come from Arizona. The species is
not known otherwise as American, the two examples have a quite European
facies, and the discontinuous distribution involved in the supposition that
they are N. American is very remarkable. We suspect some error.
( vii )
Southern range— 35° N. lat.
'A. thetis into Mauretania (withoiit
A. coridon).
A. coridon into Syria (without A.
I. thetis).
(Both sjiecies occur on the north
shores of the Mediterranean, in
Spain, Italy, Greece and Asia
Minor. )
Eastern range — 50° E. long. — Both species in Persia.
(A, thetis the Ufa dist. 56° N. lat., 55° E.
long.
A. coridon the Orenburg dist. 55° N. lat.,
55°-59° E. long.
f A. thetis — 56° N. lat. in the Ufa dist.
{A. coridon — 57° N. lat. in the "Viatka dist.
(Both speeies then fall to 53° N. lat. in Northern
Germany, A. thetis extending into Denmark
(without A. coridon), both pass into Belgium
and then into England to about 52° N. lat. {A.
thetis) and 53° N. lat. {A. ooridon).)
Western range. — The Atlantic littoral for both species, in England,
France and Spain.
North-eastern range.
Northern range.
One quite appreciates the value of a “physiological factor”
as an effective “ isolation ” barrier. This cannot arise over
the foodplant which, in Britain, and certainly in most con¬
tinental localities is restricted to Hijr^jocrejns comosa. Zeller
has recorded {Stett. Ent. Ztg.) Coronilla vciria as the foodplant
of A. coridon, but Krodel asserts {Allg. Zeits. far Ent.) that
larvae of A. coridon that he had, prefeimed to starve rather than
eat G. varia, although feeding ravenously again as soon as ff.
comosa was obtainable. The larva of A. thetis, too, is reported
as refusing to feed on anything but H. comosa.
We have already dealt with the long lists of reputed food-
plants of these species (possibly nearly all erroneous) in A
Natural History of the British Butterflies, vols. iii and iv.
But two “isolating” factors occur in the habits of the
species : —
(1) A. thetis hybernates as larva in the 3rd instar. A. coridon hyber-
nates as fully-formed larva inside the eggshell, and does not hatch till
about end of March.
(2) A. thetis is throughout its whole range doirble-brooded, not failing
in its most northern habitats. A. coridon is [with the exception of a
limited area on the Riviera, and possibly in some lowlying (coast) districts
of Italy and Spain] single-brooded.
The result of these factors is such that the single brood of
A. coridon (July-September), roughly falls between the two
broods of A. thetis (May -June and August-October), with
( viii )
considerable overlapping of the later A. coridon, and the earlier
second brood examples of A. thetis in late August and
September.
This overlapping has led to a remarkable result, for there
can be little doubt that occasional cross-pairings take place
{teste Sabine). Many are recorded in other butterfly species
(see A Nat. Hist. Brit. Lep., vol. v, chap, i, “ Mongrelisation
and Hybridisation in Lepidoptera”), but these are from one
case or other inoperative or ineffective in nature. In these
two species, however, there can be no doubt that natural
hybrids occasionally result, to which the identity of habitat,
overlapping in time of appearance, similarity in habitat,
almost structural identity of genitalia, actual identity of food-
plant, all contribute.
The first of these hybrids was described by Zeller {Stett.
Ent. Ztg., 1845), from Posen, as poloniis, and is now in the
Brit. Mus. coll. Two others were known to Zeller. The other
known examples ai-e:—
(1) Kerasdere, June 23rd, 1875, Staudinger (Hor. Soc. Ent. Ross., xiv,
p. 244).
(2) Pomeranian border of Silesia, Staudinger (7/or. Soc. Ent. Ross.).
(3) Cuxton, Kent, May 20th, 1893, Tutt (Proc. Ent. Soc. Loud.,
1894).
(4) Airolo, Ticino, June 1907, Dadd [Proc. Ent. Soc. Land., 1908).
(5-6) Pyrenees, June 25th, 1907, Keynes [Ent. Rcc., xx., p. 178).
(7) Plain of Mugnone, end July, Stefanelli (Bull. Soc. Ent. Hal., xxxii,
p. 339).
(8) Fiesole, early August, Stefanelli [Bull. Soc. Ent. Bah, xxxii, p.
339).
(9) Chateau de la Batiaz, Favre {Mitt. Schw. Ent. GeselL, xi, supp.
p. 4).
(10) Vernet les Rains, Oberthiir {Etudes, xx, p. 21).
(11) Cauterets, Oberthtir {Etudes, xx, p. 21).
(12) ? Bella Tola, 1878, Forbes {Ent. Mo. Mag., xv, p. 277).
(13) Digne, Muschamp {Ent. Rec., xxii, p. 30).
(14) Pfyn, July 1900, Rosa {Ent., xxxv, p. 96).
(15) Brides-les-Bains, July 19th, 1904, Reverdin {Ent. Rec., xxii, p.
30).
(16) Brides-les-Bains, July 1st, 1891, Blachier {Eat. Bee., xxii, p. 30).
(17) Bois-des-Freres, Geneva, Blachier {Ent. Rec., xxii, p. 30).
(18) Foot of the Saleve, Blachier {Eat. Rec., xxii, p. 30).
(19) Fei.stenberg, June 19th, 1900, Presseicker (as hafneri, Veidi. z.-h.
Ges. Wien, 1908, p. 68).
(20) Oberfeld, near Wippach, June 28th, 1907, Presseicker (as hafneri,
Verh. z.-b. Ges. JViea, 1908, p. 68).
(21) Shar Deresy, North Syria {see Tutt, Eat. Rcc., xxii, p. 30).
(22-26) Assisi, June and July 1909, Wheeler {Eat. Rec., xxi, p.
250).
( ix )
Only one 9 is known. This is in onr own collection, and
was captured at Neu Spondinig, August 12th, 1909. The cJ s
have been recorded under various names — caucasica, cory-
donius, hafneri, etc.
Great trouble has occurred owing to Staudinger in his
Catalog, having erroneously referred the bright blue Asia
Minor forms — caucasica, Led., corydonius, H.-)Sch., syriaca,
Tutt, and the Spanish form — hispana, H.-Sch. — to polonus, Zell. ,
as a variety of A. thetis, whilst retaining the various names
for other forms to which the names were not originally given.
This confusion we have already cleared up in A Xat. Hist, of
the British Lepidop)tera, vols. x. and xi.
The general variation of the two Agriadids is well known,
as is also the parallelism of the forms in the 9 s. This latter
is very remarkable, and extends from 9 s entirely brown
without markings on the upper side {A. thetis ab. unicolor,
Tutt; A. coridon ab. unicolor, Tutt) to forms well-banded
with orange lunules (A. thetis ab. marginata, Tutt ; A. coridon
ab. aurantia, Tutt), and in another direction to forms in which
the surface of the wings is so entirely blue-scaled that they
might be mistaken for c^s (A. thetis ab. coelestis, Obth. ; A.
coridon ab. excelsa, Tutt). The extreme forms of blue-scaled
9s sometimes occur discontinuously without intermediates.
At other times they occur with well-defined intermediate blue-
scaled aberrations. It may be advisable to note that the
bluest (^s of A. thetis occur either in spring or in those autumns
following summers in which the meteorological conditions have
been bad during the feeding-up of the summer brood. The
bluest 9s of A. coridon similarly occur either among the very
latest that put in an appearance, or after wet and cold
summers, or in bleak and exposed places ; in some localities,
e.g., Royston Heath, Herts, the semisyngrupha form is almost
racial.
The racial forms of A. thetis are very restricted; —
(1) var. pitnctifera, Obth. — Mauretania and Spain.
(2) var. coelestis, Obth.^ — Western France, is almost racial
in the 9-
The racial forms of A. coridon, chiefly confined to its most
( X )
south-western (Spain) and south-eastern (Asia Minor, Syria,
etc.) habitats are most interesting, and show a remarkable
parallelism in development, whilst not at all tending to
become identical. This parallelism is shown chiefly in the
coloration of the s. In the south-western and south-eastern
areas, the following pai’allelism in the forms occur : —
COLOUK, BRIGHT BIUTR (APPROACHING THAT OF A. TIRETIS).
{var. caucasica, Led.— Caucasus,
var. coryclonius, H.-.Scli. — Taurus nits,
var. syriaca, Tutt. — Lebanon nits.
South-western race var. hispana, H.-Sch. — Aragon, etc.
Colour, pale.
South-eastern race
South-western races
/var. olympica, Led. — Brussa, Aiuasia, Sliar
t Deresy (North Syria).
(var. arragonensis, Gerh. — Aragon,
ab. caerulescens, Tutt {albirxtns, H.-Sch.). —
Aragon.
var. albicans, Bdv. — Sierra Nevada.
Little is yet known of the Rivieran races, of which three
have recently been brought forward ; —
(1) var. meridionalis, Tutt. — Hyferes, Draguignan, Ste. Maxime, etc.
April-May.
(2) var. constanti, Reverdin (not yet published). — Pardigon, Canadel,
Cavalaire. April.
(3) var. rezniceki, Bartel. — Italian Riviera, Nice, Mentone. June.
The first-named seems to be the general variety or local race
throughout the Riviera, the two others localised forms of this
race so that Reverdin suggests the names should be applied
as : —
var. meridionalis, Tutt, forma constanti, Reverdin (dark underside).
„ ,, forma rezniceki, Bartel (pale underside).
Besides these in certain parts of Germany, Austria-Hungary,
Bosnia, etc., a form with very wide marginal band, extending
to the discoidal along the costa, becomes racial, although, in
England, Fi’ance, Switzerland and most parts of Central
Europe it occurs only aberrationally. This is var. {et ab.)
inarginata, Tutt (1896) \_ = horussia, Dadd (1909)].
The normal underside spotting is very similar in both
species, and its modifications almost identical. The underside
form of A. corido7i, with only discoidal and marginal spots
( xi )
present = corydonis, Bergstr., was erroneously printed
corydonius in the Proc. Ent. Soc. Lond., 1909, p. Ixxx.
Exhibitions were also made by Mr. G. T. Bethume-Baker
of varieties of A. coridon from Spain, Greece, Syria, Asia
Minor, and Persia, and of A. thetis (bellargus) from Algeria,
Greece, and Persia ; by the Rev. George Wheeler of examples
from Italy and Central Europe, and by Miss M. E. Fountaine
of A. coridon var. olympica, taken by herself at Amasia, Asia
Minor, and of A. thetis var. syriaca from the Lebanon.
At the end of Mr. Tutt’s remarks, the discussion was con¬
tinued by Mr. A. L. Rayward, Mr. Hamilton H. Druce, the
Rev. G. Wheeler, Dr. T. A. Chapman, Mr. W. G. Sheldon,
Mr. H. Rowland-Brown, Miss Fountaine, and other Fellows,
the President paying a special tribute to Mr. Tutt’s lucid
explanation and diagnosis of the various forms of the two
closely-allied butterflies.
Paper.
Dr. Malcolm Burr, D.Sc., F.L.S., F.Z.S., communicated a
paper entitled, “A Revision of the Labiduridae, a Family of
the Dermaptera."
Wednesday, March 2nd, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the Chair.
Obituary.
A vote of sympathy and condolence was passed unanimously
with the family of the late Edward Saunders, F.R.S., and of
the late George W illis Kirkaldy, Fellows of the Society. The
decease was announced, also, of George Carter Bignell and
Albert Piffard, Fellows of the Society.
Entomological Society of Russia.
It was announced that the Society would be represented at
the forthcoming Jubilee Celebration of the Entomological
Society of Russia by Dr. Malcolm Burr, D.Sc., F.L.S., F.Z.S.,
etc., and that he would deliver the following address of con¬
gratulation : —
( xii )
‘ ‘ To THE Entomological Society of Russia'.
“ The Entomological Society of London offers its most cordial
congratulations to its colleagues of St. Petersburg on the
occasion of the Fiftieth Anniversary of the foundation of the
Entomological Society of Russia. During the years that have
elapsed since the entomologists of Russia constituted them¬
selves as an organised body, the science to which the Societies
of London and St. Petersburg are equally devoted, has made
unexampled progress. The scientific value of entomological
study and research is noAV universally recognised. The light
which has been shed by the labours of entomologists upon
questions of general biology, especially in the department of
bionomics, is of the highest importance ; and the services
rendered by the economic branches of our science have been,
and will continue to be, of the utmost benefit to civilisation.
In all these forms of scientific activity the Entomological
Society of Russia, illuminated by many great names, has
borne a worthy part, and the Entomological Society of London
gladly welcomes the present opportunity of offering to its
brethren in Russia this expression of hearty goodwill, and of
congratulating them on their successful pursuit of a science
which will do much to encourage and foster the Comity of
Nations.
“On behalf of the Entomological Society of London,
“F. A. Dixey, M. a., M.J)., F.R.S.,
President.
“ H. Rowland Brown, M.A.,
“Comm. James J. Walker, M.A., R.N., F.L.S.,
ii'ecretaries.
“Feb. 28, 1910.”
Election of a Fellow.
Mr. John C. Eales-White, of 47, Chester Terrace, Eaton
Square, S.W,, was elected a Fellow of the Society.
Exhibitions.
Lycaenid and Argynnid Forms. — Mr. W. G. Sheldon
exhibited various forms of Agriades (Eolgommatus) coridon
from Southern Europe, including A. var. arragonensis, and its
form caerulescens from Albarracin, Spain ; also a pair of
( xiii )
Brenthis selene from La Granja and a pair of B. hecate from
Hungary, showing the remarkable approximation of the two
species in the markings of the under-side.
Association of Ants. — Mr. H. St. J. Donisthorpe brought
for exhibition a case containing a small nest of about 30-40
living examples of Formica rujibarhis, vax.fusco-rujibarbis, from
Whitsand Bay, August 1909, into which had been introduced
a queen of Formica rufa from Nethy Bridge (May 1909).
The queen had been accepted by the other ants in a few days.
Male Beetle with Female Antennae. — Mr. C. 0. Water-
house, on behalf of Mr. J. C. Moulton, of the Sarawak Museum,
exhibited a Longicorn beetle of the genus Cliloridolxmn, in
which the right antenna was much shorter than the left.
The specimen appeared to be a male in every respect except
in the length of the right antenna, the joints of which were
a trifle shorter than those of a female. Mr. Moulton was
inclined to think the specimen might be gynandromorphous.
Mr. Waterhouse suggested that it might rather be regarded
as a malformation,
Teratological Example of a Beetle. — -Mr. W. E. Sharp
exhibited an example of Calathus mollis, having the right
anterior tibiae and tarsus in triplicate. He said that such
reduplication was not uncommon, but that it was seldom so
perfect in detail. The occurrence raised the whole question
of the cause of the reduplication of such parts, whether it was
due to hereditary defect, or to external stimulus exciting cell
reduplication. A discussion folloAved in which Professor E.
B. PouLTON, F.R.S., Dr. T. A. Chapman, Mr. J. W. Tutt, the
Rev. F. D. Morice, and Professor Hudson-Beare and other
Fellows participated.
Zygaenid Forms. — Mr. L. W. Newman showed a case contain¬
ing the following forms of Zygaena liippocrepidis, all taken in
June 1908 in one small field in the neighbourhood of Bristol,
by Mr. H. J. Smallcombe : (a) analogous to A. fili'pendidae,
var, chrysanthemi, {b and c) examples with the normal carmine
coloration replaced by yellow, analogous to A. Jilipendulae, ab.
jiava, Robson, and by pink ; and {F) an example with yellowish
spots, otherwise normal. Also a specimen of Z. lonicerae
captured at the same place, with the markings broadly confluent.
( xiv )
Lasiocampid Nests. — Mr. G. Bethune-Baker brought for
exhibition three nests of a species of Lasiocampid moth from
Mount Elgon, Albert Nyanza, belonging to the same group as
2'haumatopoea in’ocessionea. He had been consulted relative
to the possibilities of using the silk of which the long pouch¬
like “nests” wei’e constructed. Mr. G. A. K. Marshall said
that they belonged to a species of Anaphe.
Professor E. B. Poulton, F.R.S., read the following —
Preliminary Note on Mr. A. D. Millar’s experimental
BREEDING OF FORMS OF THE NyMPHALINE GENUS EUKALIA IN
Natal. — By Boland Trimen, M.A., F.K.S.
There are three forms of Eurcdia which inhabit the coast-
land of Natal, vid. E. vmhlheryi, Wallengr., E. mivui, Trim.,
and E. decoptor, Trim. The first and second of these, without
being numerous, are met with not uncommonly ; the third,
first brought to notice in 1869, and described by me in 1873
from a single has hitherto remained very rare in the
(Colony. Each of the three forms in both sexes unmistakably
and very closely mimics a Danaine of the genus Amauris
prevalent in the same district : — thus, E. loahlbergi mimics
A. niavius, sub-sp. dominicanus, Trim. ; E. mima mimics
A. alhimacidata, Butl., as well as A. echeria, Stoll. ; and E.
deceptor mimics A. ochlea, Boisd. The intimate afiinity of the
three Eurcdiae is apparent, although their wing colouring and
marking present such Avide differences. To this I called atten¬
tion when describing (Trans. Ent. Soc., 1873. p. 107 and foot¬
note) E. deceptor, as well as to the fact that E. viiina 3 Rud
E. wcddberyi $ had been taken paired.* Since then Mr. A. I).
Millar, Mr. C. N. Barker, Mr. Guy A. K. Marshall, Mr. G. F.
Leigh, and other entomological observers in Natal have brought
to notice sever.al other _cases of the pairing of these tAvo forms,
and also various details of habits, distribution, etc., Avhich sup¬
ported the vieAv of their species-identity. Recently Mr. Millar,
having di.scovered the foodjilant — a stinging-nettle recognised
by Mr. J. Medley Wood as a species of Fleurya — has been
enabled to test matters by breeding from ova laid respectively
by a wcddberyi, a mima, and two deceptor. These four 9 9
Avere all taken toAvards the end of March 1909 — the loahlbergi
* See also “South Afr. Butt.,” I, pp. 282, 283 and 285 (1887).
( XV )
and the two deceptor on the 21st, and the viima between that
date and the 26th.
The results were as follows, viz. —
(a) Fi’om 10 ova laid by one wahlbergi were bi’ed 4 wcMhergi
(2 c? , 1 $ , 1 deformed, sex undetermined), and 5 niima
{4: 6 6,1 9)-
(h) From 39 ova laid by one mima were bred 33 mima
(24 6 6,^ 9 9)-
(c) From 139 ova laid by two deceptor were bred 127
decepto^' (70 6 6,57 9 $ )•
Experiment (a) proves the species-identity of wahlbergi and
mima ; while experiment {b) seems to indicate the probable
predominance — as in the case of the $ forms of Papilio
dardanios, sub sp. cenea — of the models Amaitris echeria and
albimacidata in affecting the mimetic pattern and colouring.
Experiment (c) affords no evidence of deceptor belonging to
the same species as the other two forms, all the bred examples
of both sexes being recorded by Mr. Millar as true to type.
The case of Euralia ivahlbergi and mima is in agreement
with that of Charaxes zoolina and neanthes* as regards the
dimorphism including both sexes ; but there is a striking
difference to be observed, in that the double mimicry so
highly developed in both sexes of the Euralia finds no place
in the Charaxes, whose colouring in both forms, but especially
in neanthes, is by no means conspicuous and on the under-side
apparently cryptic.
Mr. Millar has furnished photographs of the three sets of
Euralia progeny bred by him, with many most valuable details
as to transformations, etc., and also a selection of specimens
of the larvae, pupae, and imagines. With all these data I
propose to deal fully in a later communication to the Society. t
* See Mr. G. F. Leigli’s interesting account (Proc. Ent. Soc. Lond. for
21st October, 1908, p. Ixiv, and 6th October, 1909, p. xlix) of his experi¬
mental breeding of these forms from the ova.
t Since the meeting on 2nd inst., I have received from Mr. A. D. Millar
the result of a second experiment in breeding from tbe ova of the mima
form, ditfering remarkably from that of the first (b) described above. From
a ? mima’s 11 ova, laid on 21st November, 1909, were bred 8 mima
(5 8 8,3? 9 ) and 3 wahlbergi (8 8 ). This rounds off the case very
satisfactorily. As in result (a), the toahlbergi and mima of this last family
are respectively all true to type, and not one example is at all intermediate
between the two forms. — R. Trimen, March 15, 1910.
( xvi )
Professor Poulton reminded the Fellows of Mr. Guy A. K.
Marshall’s conclusion that E. locUdbergi and mima were forms
of the same species (Trans. Ent. Soc. Lond., 1902, pp. 491,
492), and of the exhibition of specimens relied on as evidence
(June 6, 1906 : Pi’oc. Ent. Soc. Loud., 1906, pp. liii, liv). He
felt that Mr. A. I). Millar was to be greatly congi’atulated on
his success in obtaining this long-wished-for proof of a most
impoi’tant and probably far-reaching conclusion.
Professor Poulton considered that the uniform offspring
of the mima parent (/>) could not be explained in the same
manner as the predominant cenea offspring of P. dardaims in
Natal ; for according to Mr. G. F. Leigh (Proc. Ent. Soc.
Lond., 1906, p. Ivii) mima is the rarer of the two forms in
Natal, whereas in the same area cenea is by far the commonest
of the dardaims females.
The comparison of the results obtained by Mr. Millar in (a)
and (b) suggested a Mendeliau relationship between the two
forms.* It Avas interesting to compare the records of two
broods obtained from llypolimnas misipjms (Proc. Ent. Soc.
Lond., 1909, pp. xxxvi, xxxvii).
Professor Poulton ahso exhibited a set of 6 Euralia anthedon,
iJoubh, and 4 E. dnbia, Beauv., captured in the same locality,
Oni, 70 miles east of Lagos, by Dr. W. A. Lamboru (Dec. 1908-
Dec. 1909). Anthedon was the western representative of
niahlbergi, dubia of mima, and noAv, after the proof obtained
by Mr. Millar, it became almost certain that the Av^estern
butterflies also were the dimorphic foi’ms of a single species.
He trusted that Dr. Lamborn Avould be able to obtain indis¬
putable proof of this conclusion.
Professor Poulton considered that it was convenient at this
point to correct an error for Avhich he Avas responsible in Rev.
K. St. Aubyn Rogers’ paper in Trans. Ent. Soc. Lond., 1908, p.
489. Euralia hirbyi, But!., mentioned and figured (PI. XXVII,
fig. 5) in this memoir, Avas not a distinct species, but the male
* Since tlie date of the meeting I have had the chance of discussing
the facts witli Mr. L. Doncaster, Avho agrees that the two forms are
probably a Mendeliau pair, but considers that we have not at present
sufficient evidence to decide whether viahlbergi or mima is dominant.
Mr. Doncaster agrees that the last result, recorded by Mr. Trimen in the
foot-note on p. xv, makes it probable, but by no means certain, that mima
is the dominant. — E. B. Poulton, March 21,1910.
( xvii )
of E. deceptor, Trim. The specimens sent by Mr. St. Aubyn
Rogers had been compared with tlie British Museum collection
in 1907, when the males stood under kirbyi, the females under
deceptor. When, however, in 1909, a number of the specimens
of decepitor bred by Mr. Millar were examined, the error
became manifest. In the mean time the mistake had been
independently detected and set right in the collection of the
British Museum. The male of E. deceptor was described by Mr.
Trimen in Trans. Ent. Soc. Bond., 1873, p. 105, and both sexes
in “ South African Butterflies,” 1887, 1, p. 286. The male was
described by Dr. A. G. Butler as kirbyi in P.Z.S., 1898, p. 51.
Professor Poulton exhibited the female of A2xitiiropsis
cleocharis, Hew., taken Feb. 1, 1909, by Mr. C. F. M.
Swynnerton on the outskirts of Mt. Chirinda Forest (3,800
ft.), MeLsetter, Gazaland, S.E. Rhodesia. The exhibitor did
not know of another example of the female of this rare
butterfly, which differs from the male to a remarkable extent,
being far larger and possessing a rounded hindwing without
the produced anal angle which is so characteristic of the
opposite sex. Professor Poulton also exhibited an example of
the male captured by Mr. G. A. K. Marshall, Oct. 19, 1905,
in the same locality.
rajxrs, etc.
The following papers, also, were read : —
“Third Paper on the Tetriyinae (Orthoptera) in the Oxford
University Museum,” by J. L. Hancock, M.D.
“ Descriptions of new Algerian Hymenoptera-Aculeata
{Spheyidae) ” by the late Edward Saunders, F.R.S., and the
Rev. F. D. Morice, M.A.
“ On Zizeeria, Chpmn. ( = Zizera, Moore), a group of Lycae-
nid Butterflies,” by Dr. T. A. Chapman, M.D., F.Z.S.
“Further notes on two Osmia species of the group,”
by the Rev. F. D. Morice, M.A.
“A Few Words respecting Insects and their Natural
Enemies,” by Arthur G. Butler, Ph.D., F.L.S., F.Z.S.,
M.B.O.U.
The Rev. F. D. Morice, M.A., also delivered an address
“On the Saws of Saw-flies,” illustrated by many lantern slides
prepared and arranged by him.
PROC. ENT. soc. LOND., I., II. 1910.
B
( xviii )
Wednesday, March 16th, 1910.
Dr. F. A. Dixey, M.A., M.D., F.K.S., President, in the
Chair.
Address to the Entomological Society of Russia.
Dr. Malcolm Burr, D.Sc., gave the following account of
his reception in St. Petersburg by the Entomological Society
of Bussia, to whom he had presented the Addre.ss of Congratula¬
tion, published at the last meeting.
The Entomological Society of Ptussia, one of the oldest
scientific bodies in the Empire, celebrated its Jubilee on
February 26th (Old Style) = March 11th (New Style), in the
Ministry of Agriculture, at St. Petersburg.
The chair was taken by the venerable President, Piitr
Petrovich Semenoff-Tian-Shansky, the eminent explorer and
traveller in Central Asia, discoverer of the Tian-Shan Moun¬
tains, President also of the Russian Imperial Geographical
Society, and member of the Imperial State Council. There
were also present Mr. Kozloff, another distinguished explorer
in Central Asia, Colonel G. V. Kakhovsky, an enthusiastic
coleopterist who has made important collections in Abyssinia,
Professors Rimsky-Korsakoll, Inostranstseff, and Shimkievich;
Academicians Borodnik, and Karpinski, Prince Massulsky, Mr.
Dostoievsky, secretary of the Russian Imperial Geographical
Society and kinsman of the famous author, Messrs. Alferaki,
Adelung, Bianki, Skarikoff, KuznetsofP, and E. E. Groum-
Grgimailo, lepidopterist and zoogeographer.
The President opened the proceedings by reading a letter
from H.I.H. Grand Duke Nicolas Mikhailovich, regretting
his inability to attend personally, and sending greetings to
the Society with his congratulations upon its fiftieth anni¬
versary. The President then greeted the Society, and
especially congratulated Baron Th, R. von Osten-Sacken, the
only surviving original member.
One of the Hon. Secretaries, Mr. G. G. Jacobson, then gave
a brief outline of the Society’s history, after which the Vice-
President, Mr, Andre Petrovich Semenoff-Tian-Shansky, one
{ xix )
of the foremost Entomologists in Russia, scarcely less dis¬
tinguished by his work upon the Chrysididae and Der)natoj)tera
of the Russian fauna than by his studies upon the Coleoptera,
read a paper upon the scientific achievements and literature
published by the Society,
The Secretary then announced the names of the Honorary
Members elected in celebration of the Jubilee.
Delegates from various institutions next presented their
addresses. The only foreigner present, Dr. INIalcolm Burr,
congratulated the Society on behalf of tlie Linnean, Entomo¬
logical, and Zoological Societies of London, and presented their
Addresses. The references in the Addresses to the bonds of
scientific good-fellowshiji which does so much to promote the
comity of nations, were received Avfith enthusiasm by the meet¬
ing, and in accepting the Addresses, the President charged Dr.
]\Ialcolm Burr to convey to the Council and Fellows of the
Societies w'hich he represented their deep appreciation of the
compliment paid by English men of science in sending a repre¬
sentative so far, and his hearty concurrence in the value of
such international courtesies in the scientific world, which
strengthened the bonds uniting friendly peoples.
Dr. Poppius and Prof. Sahlberg, Delegates of the Finnish
Geological Society and University of Helsingfors, read an
Address in French, after which followed a number of Addresses
from numerous Russian and learned bodies and Institutions
of Agi'icultiu’e and Forestry.
jMr. Oscar I. Jon, As^tant Hon. Secietary, then proceeded
to read very many letters and telegrams of congratulations
from all parts of the world including the Rothschild Museum
at Tring, the Senckenburgische Katurforschergesellschaft,
and the Raturforscherverein of Steiermark, the Royal Society
of Naples, and the Entomological Societies of Belgium,
Holland, Bohemia, Ontario, Switzerland, and Bulgaria, the
University of Upsala, the New York Academy of Sciences,
the Smithsonian Institution of Washington, the Indian
Museum of Calcutta, the “ Iris ” Society of Dresden, the
Geological Society of France, and from very numerous private
individuals.
B 2
( XX )
The Society and its guests were then photographed, and
after some light refreshments, the meeting terminated.
In the evening, the delegates were entertained at a banquet
as guests of the Russian Society. Numerous speeches were
made, one of the most striking of which was that of Mr.
Kuznetsoff, Editor of the Revue ruase d' Entomologie, who
spoke in admirable English, addressing his remai-ks through
Dr. Burr to the English nation, ever leaders, as he .said, in
action and in thought. Since Newton laid his foundation of
IModern Mathematics and Darwin revolutionized the theory
of modern Biology, he said, England has continued to })our
forth many leaders of Science, and in the domain of Entomology,
Stainton, Mac Lachlan, Saunders, Boulton, Dixey and Tutt.
Dr. Burr briefly thanked him on behalf of his fellow-countrymen.
Election of Felloios.
Mr. Edmond Wage Carlier, M.Sc., M.D., F.B.S.E., of the
University, Birmingham ; Mr. Herbert Alfred Green, of
the Central Fire Station, Durban, Natal; Mr. Philip Har¬
wood, of 23, Northgate End, Bishop’s Stortford ; Mr. J.
Henderson, of Clifton, Ashbourne, Derby; Mr. Lionel Leslie
Jacobs, Shelford, Copers Cope Road, Beckenham ; Mr. Wil¬
liam Laidlaw, 73, Endsleigh Gardens, Ilford, Essex, and 74,
Great Tower Street, E.C. ; Mr. H. S. Leigh, of the University,
Manchester ; Mr. F. Graham Millar, of Seafield, Batu Tiga,
Selangor; Mr. Francis Allcock Oldaker, M.A., of the Red
House, Haslemere; Mr. Aiyappa Raman Pillai, Trivandrum,
India, and 13, Buccleuch Place, Edinburgh; Profe.ssor Reginald
Crundall Punnett, M.A., of Gonville and Cains College,
Cambridge; and Mr. James M. Williams, F. R. Scott, G.S.,
of the Howard IMotor Garage, Cardiff, and Canford Cliffs,
Hants, were elected Fellows of the Society.
Exhibitions.
Ant and Mimic Beetle. — Mr. H. St. J. Donisthorpe
exhibited examples of Cremastogaster scutellaris Colohopsis
truncxtns and a beetle Forniicomus pedestris, a good mimic
of the latter ant, all taken in virgin cork at Kew, May 1909.
( xxi )
Living Examples of Holoparamecus. — Commander J. J.
Walker. R.N., exhibited living specimens of Holojxiramecus
caularum, Aube, taken commonly in haystack refuse at Water
Eaton, Oxon., on the 14th of March.
Ova and Appendages op Tapinostola. — Mr. H, M.
Edelsten brought for exhibition photographs of the anal
appendages of Ta2nnostola helhnanni, T. concolor, and 2'.
fulva, showing their remarkable similarity ; also ova of the
same species in situ. The photographs were the work of Mr.
H. Main, and Mr. A. E. Tonge.
Rare British Coleoptera. — Dr. G. W. Nicholson showed a
specimen of Dyschirius caigustatus, Ahrens, from Littlestone,
Kent, taken in July 1906; two specimens of Bembidium 4-
jmstulatum, Dj., taken at Pulborough in June 1909 ; and two
specimens of C onosomahipunctatum, Gr., found by Mr. Jennings
and himself at Broxbourne, Essex, in January of this year.
Notes on Bred Specimens of Callophrys avis. — Dr. T. A.
Chapman exhibited series of Gcdlophrys avis bred this spring
together with series of C. rubi. In order to get C. avis for¬
ward he had placed them at the beginning of February at a
temperature of 70° to 72° Fahr. He said a curious circum¬
stance was that two specimens emerged four days after being
brought from a temperature of 46°, one five days after, two
six days after, two seven days after, and so on. It seems
certain that those emerging so soon after removal to a warmer
temperature must already have made much advance to matur¬
ity at the lower/temperature, possibly ruuch earlier in the
Avinter, when the thermometer Avhere they were sometimes
reached 50°.
The specimens show much uniformity in size, more on
measurement than in appearance (due to setting 1) the range
being 33 to 36 mm.
The specimens placed together show a very bright and fairly
uniform red tint, markedly contrasting with the ruddiest rubi
I have, and looking brilliant beside them. The females are
rather larger than the males.
The range of A^ariation is not great. The males are darker
than the females, nearly all the males show the conspicuous
dark veins of the forewings characteristic of the species. In
( xxii )
the females this is much less marked. The males are al¬
together darker than the females. One or two males are
very dark, one or two of the females rather light. One or
two females have a distinctly darker margin just within the
fringe, with a paler one just within it.
The male androconial brands vary from nearly black to a
pale reddish not far from the ground-colour. In not a few
specimens they look almost white, and this was noticed to be
the case in several specimens almost immediately on emergence.
In some of these with white brands, the colour is white when
looked at from one angle, dark from another, and often one
brand looks dark, the other white, reversing when seen from
another angle. The effect seems to be due to a variation in
the degree to which the scales are erect on the wing, so that
in some lights, not the true scale colours, but only reflected
light from their sui'faces is seen. The ordinary scales lie very
flat, the androconia are raised at a considerable angle, and are
twisted so that often portions of the scales are almost at right
angles to the wing surface.
In the underside there is also some variation. Some are
of a fairly deep-blue-green, in others, and this is more frequent
in the females, there is quite a brownish tone in the green,
and in one or two specimens the brown is very evident
towards the fringes and along the nervures of the anterior
wings ; there is also an extension in a considerable proportion
of specimens of the brownish-grey area of the forewing where
the hind wing covers it. There are variations precisely parallel
to all these in C. ruhi.
The white line is less variable than in C. ruhi, but this is
very possibly due to all the C. avis being from one locality,
the C. ruhi I have from very numerous ones.
In only one or two is the line obsolete on the forewing, in
no specimen on the hindwing.
On the hindwing it is frequently obsolete or nearly so on
the inner half of the wing. The position (nearer the base)
and character of the markings is very different from C. ruhi
in all examples.
The individual elements of the white line, i. e. the portion in
each interneural space, are not more or less square, but dis-
( xxiii )
tinctly lunulated as in Neolycaena lunulata, tengstroemi, or
sinensis, and in one or two specimens one almost sees the
cusps running basally for a small distance, especially in the
first costal spots of the hindwing. One variation is specially
to he noted, two or three specimens have dark inner margins
to the lunules, as is usual in C. ruhi hut which I had not seen
in my captured specimen of C. avis. In one or two specimens
there may he seen a trace of the silvery eyebrows, so character¬
istic of C. ruhi.
On emergence the dehiscence of the pupa differs in one respect
from that of C. ruhi. In both species the prothoracic plates
sometimes separate from the rest of the pupa case, sometimes
they remain attached to the anterior border of the mesothorax ;
the difference between the two species is, that it is rather the
rule for them to separate in C. avis, rather the exception in
C. ruhi.
The few empty pupa cases also exhibited show this, so far
as a few can do so ; they also display the brown colouring of
the wing cases in C. ruhi, and their nearly black in C. avis, a
difference rather more marked when they are alive.
South African Lycaenidae. — Mr. Hamilton Druce drew
the attention of Fellows to his recently published book on
“ Some South-African Lycaenidae'' a copy of which he
exhibited.
Canadian Coleoptera. — Mr. G. C. Champion called atten¬
tion to a paper of interest to British Coleopterists, published
in Canada, entitled “ Some Guests at the Banquet of Blossom,”
by T. J. A. Morris, read at the Annual Meeting of the En¬
tomological Society of Ontario, November 4th, 1909, and
published in the “Canadian Entomologist,” December 1909,
pp. 409-418.
Papers.
Separation of British Species of Hydroecia. — Mr. J. W.
Tutt delivered a short address on “ The species of the nictitans
group of Hydroecia." He discussed the several forms of
Hydroecia occurring in Britain, and illustrated the superficial
differences where discoverable, and the marked difference in
the anal appendages of the several species, Hydroecia nictitans,
( xxiv )
II. paludis, H. lucens, and II. crinanensis. He pointed out that,
as far back as 1888, he had differentiated three species among
the specimens usually united under the name nictitans, viz.
Ilydroecia nictitans, a generally distributed inland species ;
Ilydroecia paludis, a paler coast species ; and Ilydroecia lacens,
a moss and moorland species. In liis “ British Noctuae and
their Varieties,” vol. i, pp. 58-64, he described these three
species at length, and gave a long account of their variation.
Recently the examination of the genitalia had proved the
distinctness of these species and had discovered a fourth
species, Ilydroecia crinanensis, the facies of whicli was prac¬
tically inseparable from II. hicens. Long series of the four
known British species were exhibited. Mr. Tutt, then, by
I'eference to some large, clearly-drawn diagrams, explained the
structure of the ancillary parts on which the differentiation
was based, and pointed out the differences in the four species
in the structure of the harpe, corona, clasper and clavus
respectively. He then dealt with the female' genitalia, and
discussed the differences exhibited in the lodix and genital
plate in the same four species. The ^ ancillary appendages
of Ilydroecia atlanticas were then discussed, and their distinct¬
ness from those of the four British species demonstrated. He
made an appeal to Fellows to try, during the coming season,
to obtain eggs, larvae and pupae of the four British species for
comparison, and to discover, if possible, structural differences
in these stages, in support of those shown by the superficial
imaginal characters and those of the ancillary appendages in
U. nictitans, II. p)cdudis,a\\d H. lucens, but more particularly to
detect, if possible, differentiating characters in the early stages
of the two species II. lucens and II. crinanensis, the genitalic
characters of which are so entirely different in both sexes
and yet whose facial characters appear as yet practically
indistinguishable. Excellent photographs of the genital
organs of the four British species (both sexes) made by
Mr. F. N. Pierce, were handed round for examination.
The Rev. C. R. N. Burrows, to whose research the dis¬
covery of H. crinanensis was really due, stated that he had
little to add to what Mr. Tutt had said ; it seemed to him
amazing that two species showing so much difference as
( XXV )
Hydroecia lucens and U. cvinanensis in theii' genitalia, should
present no definitely marked superficial character in the imago.
Dr. Karl Jordan considered that the four species, as
proved by the differences in the genitalic structures, were
abundantly distinct, and brought forward a parallel case
among the Attacids. Here, two species with almost identical
imaginal facies were separated first of all on the ^ genitalia,
and afterwards proved by the larvae and pupae to be essentially
different in all their stages. He considered the differences in
structure of the $ genitalic organs as being particularly
interesting.
Dr. T. A. Chapman, M.D., F.Z.S., read the following notes
entitled “ Xanthandrus comtus, Namr., a Correction.”
On June 6, 1906, Prof. Poulton read a paper on “ Predaceous
Insects and their Prey,” which appears in our Transactions
for that year. On p. 396 are some remarks on Xanthandrus
comtus, which involve cei’tain misapprehensions as to facts,
and consequently some probably erroneous deductions. I
pointed these out to Prof. Poulton at the time, and they were
to be corrected in a further paper on the same subject shortly
to appear. It seems, however, that the pressure of other
mattei’S is postponing indefinitely the completion of that paper
by the professor. I think, therefore, it is desirable that the
correction should not be further delayed.
I had recorded observations on X. comtus, showing that
the larva fed on the larvae of certain Tortrices, which were
gregarious in this modified sense that, though not necessarily
in association, a number were always found on the same plant,
probably all from one batch of eggs.
Prof. Poulton reports an observation by Mr. Lyle of the find¬
ing of a full-fed larva of X. comtus, which had reduced to a dry
and empty skin a larva of Scotosia, undulata,, a Geometrid moth.
The misapprehensions that Prof. Poulton adds to this fact
relate both to A", comtus and to S. undulata.
First, as to the habits of the larva of X. comtus. When this
larva finds itself amongst a number of larvae suitable for food
and of considerable size, as I have not unfrequently found it
amongst those of Hastula hyerana, it does not mind the size of
its victim, and often attacks one much too large for it. Of this
( xxvi )
the proof consists in finding, in plants with many larvae of
H. hyerana and only one or two of the fly, not a few larvae of
H. hyerana^ dead and blackened, but only partially collapsed,
the result of being killed by a larva of X. comtus much too
small to do more than suck a portion of its juices. In
captivity I have several times observed that the attacked
larva dies very soon after it has been seized by that of the fly.
These circumstances prove that a larva of X. coinUis could not
possibly, immediately it hatched, attack a larva enough to
contain a life ration for it, and yet have that ration remain
fresh during the whole period of its growth.
I ought, perhaps, to dwell on this point, as it is one that
might have occurred to the professor. Doubtless he had in
mind the habits of the fossorial wasps, whose prey remains
fresh for a very considerable time, fresh because it remains alive,
though paralysed. The rapid death of its prey and its subse¬
quent I’apid decomposition quite difPerentiate the case of X.
comtus from that of the wasps.
Mr. Lyle’s larva had no doubt already disposed of, at least,
several larvae of the Scotosia.
Then as to S. unchdata, several points deserve notice. One
is that its full-grown larva is of much the same bulk as that
of H. hyerana, and of H. hyerana a nearly full-grown larva of
X. comtus will attack and demolish a full-grown specimen,
and even more than one.
Perhaps the most important point in connection with S.
undidata is that the larva is gregarious in precisely the same
sense as the larvae of the Tortrices I have observed in this
connection are. Whether in their younger stages they are or
are not literally so, I do not know, but think probably they are.
At any rate, it is the case that several or many larvae are
always to be found on the same bush ; doubtless they have all
emerged from one batch of eggs, as in the case of the Toi trices.
The only difference, then, that exists between the cases of
S. undulata and that of the Tortrices, on which I have seen
A', comtus parasitic, is that they are Tortrices and it is a
Geometer, a condition, I should imagine, of infinite indifference
to the larva of X. comtus. A further point is not, however,
without interest and of some direct bearing on the subject,
( xxvii )
viz. that the lai’va of S. undulata infolds itself in leaves in
much the same manner as Tortrices do. It seems very probable
that the shelter thus afforded may be of some importance to
the X. comtus larva, which has a much more delicate cutaneous
envelope than have most of the Syrphid larvae feeding exposed
on Aphides that I happen to know. The shelter must be of
use, as it is to its victims, to hide it from its enemies, and the
delicate skin may be a result or a further cause of the sheltered
existence. Probably the larva searches for the shelter of its
next victim during the night when neither sun nor predaceous
enemies are to be dreaded.
It follows from all these facts and considerations that
Pi’ofessor Poulton’s beautiful theory as to the alternative
instinct displayed by the female of A", comtus, though of the
very highest interest if it could be established, may be regarded
as without the slightest foundation.
The remai’ks to the disparagement of the egg-laying instincts
of the Spliingidac must, of course, fail as a matter of compari¬
son with the instincts of X. comtus, and per se, the presence
of an unduly large number of larvae on a very small bush is
an occurrence that one not unfrequently meets with in other
insects and elsewhere.
It is the result of one particular bush presenting flavours
or other attractions to more than one moth, or to the same
moth over and over again, after it has gone in search of a
suitable place elsewhere. . A striking and not infrequent
instance is where a young pine-tree is nearly or actually
killed by a large number of nests of C. ^ntyocampa, whilst
numerous trees around, to our eyes much more tempting,
are untouched. A similar common case which most of us
have met with is in the case of Aglais urticae, where a brood
of larvae may be found on a patch (if it amounts to a
patch) much too small to nourish them all to full-growth.
Such a case usually occurs when the butterfly is very common,
and I have seen even large areas of nettles with a number of
broods more than sufficient to demolish it. All these cases
are, broadly, the results of over-population.
Mr. J. 0. Kershaw contributed a paper on “ The Oothecae
of an Asilid (Promac/ms sp.).”
( xxviii )
Wednesday, April 6th, 1910.
Mr. H. Rowland-] >RO\VN, M.A., Vice-President, in the
Chair.
Congress Delegates.
The Vice-Prestdent announced that the Society would send
as delegates to the International Congress of Entoiuolgy at
Brussels Dr. F. A. Dixey, M.A., M.D., F.R.S., Mr. II. St. J.
Donistiiorpe, F.Z.S., and one other Fellow ; to the Zoological
Congress at Gratz, Dr. K. Jordan, Ph.D.
Election of Fellows.
Mr. Horace B. Browne, M.A., of 118, Sunny Bank, Hull ;
Mr. William George Daw'Son, of 31, King’s Gardens, West
End Lane, West Hampstead, N.W. ; Mr. Alfred Hander
Hedges, of 42, Kensington Park Gardens, W. ; the Rev.
Gilbert H. Raynor, M.A., of Hazeleigh Rectory, Maldon,
Essex ; the Rev. Hubert George Stanley, of Marshfield
Vicarage, Cardiff ; and Mr. Rupert Stbnton, of Southwell,
Notts, were elected Fellows of the Society.
Exhib{tio7is.
Butterflies from the Carpathians. — Mr. W. G. Sheldon
exhibited several series of the butterflies taken by him last
July in the Hohe Tatra region of the Carpathians, Eastern
Hungary. They included examples of Melitaea dictynnoides,
Hormuzaki, with M. aw’elia and M. dictynna for comparison ;
Brenthis pales var. arsilache from the forest zone at 3000 feet,
and a form of B. pales from 5000 ft., with the upper-side
approaching in colour and markings to var. arsilache, but of
smaller size, the under-side being typical ; also Swiss examples
for comparison : Parnassius apollo, var. carpaticics, Aigner,
and P. apollo from the Swiss Alps for comparison ; also Erehia
medasa, var. hipponiedxisa, E. ligea, and Coenonympha Im'o.
The Rev. G. Wheeler expressed a decided opinion that
J/. dictynnoides constitutes a good species, and is not a form
of M. aurelia.
Rare British Beetle. — Mr. P. Harwood brought for ex-
( xxix )
hibition an example of Strangalia revestita, taken on a flowor-
head near Andover in 1909.
Mimetic Lepidoptera. from the Cameroons. — Mr. W.
F. H. Kosenberg exhibited (a) a “ combination ” consisting
of a i^ympbaline butterfly, Euphaedra ruspina, and three
species of moths belonging to as many different families,
viz. Phaegorista similis (Ilypsidae), Xanthospilopteryx ptoggei
{Agaristidae), and a Geometer, Aletis helcita. These insects
bear a close superficial resemblance to each other in colour and
pattern of markings, the wings being tawny-orange, with
black marginal borders and white apical and marginal
spots : (b) a pair of the Nymphaline butterfly, Ilarma theodota,
a strikingly dimorphic species, the female of which bears some
resemblance, especially on the upper side of the hindwings, to
a moth, Nyctemera hesperia, of the family Lymantriadae. The
specimens were taken in the same locality and at the same
time of year •, and (c) five species of Planema (family Acraeidae)
and an equal number of species of Pseudacraea (family Nym-
pkalidae) mimicking them, the supei’Acial resemblance being
very close in each case. Special attention was called to the
specimens of Planema plagioscia, the males of which, with
tawny bands on the forewings, are mimicked by the males of
Pseudacraea hohleyi, whilst the females of the Planema, with
white bands are mimicked by the females of the same species
of Pseudacraea. With these specimens were exhibited several
Elymnias, and specimens of Papilio zenohia, cynorta, and
another species of the same group, all mimicking certain species
of the Planema. The whole of these specimens were taken in
the same months and place.
Mr. J. W. Tutt considered the exhibition made by Mr.
Kosenberg one of the most interesting that had been pre¬
sented at any meeting of the Society. Here we had for
certain species belonging to widely dilferent groups of Lepi-
doptera — butterflies and moths — occurring at the same time
and in the same place with close resemblances on both
upper- and under-sides. It occurred to him that the upper-
side resemblance would be of value only during flight or
movement, and the under-side only during rest. The usual
habits of butterflies on the one hand, and Geometrids and
{ XXX )
Laclmeids on the other, were very different, both as to their
movements and their manner of resting, the moths flying by
night, the butterflies by day ; the moths resting with the
upper-sides, the butterflies with the under-sides of the wings
exposed. It would be very interesting to know whether in
the cases exhibited the butterflies and moths both flew by
day, and both rested in the same manner. This might not be
difficult to ascertain from the collector who supplied the speci¬
mens, and would be of the greatest value.
Beetle Parasite. — Mr. II. St. J. Donisthorpe exhibited
examples of Methoca ichneumonides, parasitic on the larva of
the Tiger Beetle, taken by him in the Isle of Wight. He
said that it was apparently unknown in this country that
Methoca ichneumonides is parasitic on this larva. Its life
history has been worked out by Bonuman (Tijd. voor Entom.,
1909), and Adlery (Arkiv. for Zook, 1906). They record it
pai’asitic on Cicindela campestris and C. sylvatica. He had
taken it at Blackgaug Chine, where it would be parasitic on
C, germanica. With these examples he also showed the
hosts, C. caiupestris, C. sylvatica, C. germanica, and a Tiger
Beetle larva.
Mr. Donisthorpe also exhibited an example of Ptinella
hritannica, Matt., found in a mole’s nest at Burwell Fen last
month. This is only the third British specimen yet recorded,
and apparently the fourth only in Europe, one having been
reported from France.
Papers.
Mr. Norman H. Joa', M.E.C.S., read a paper “ On the
behaviour of Coleoptera during Floods,” and exhibited living
specimens to illustrate the remai'kable power of Dianous
coerulescens in '• skimming ” on the surface of water.
The following papers were also read : —
“ A Revision of the genus Diplatys, Serv.,” by Malcolm
Burr, D.Sc., F.L.S., F.Z.S., etc.
“ On the Geometridae of the Ai’gentine Republic,” by Louis
B. Prout.
( xxxi )
Wednesday, May 4th, 1910.
Dr. F. A. Dixey, M.A., M.D., F.E.S., Pi’esident., in the
Chair.
Obituary.
The decease was announced of Mr. G. S. Saunders, F.L.S.,
a Fellow of the Society.
Exhibitio7is.
Rare Ichneumon. — Mr. John J. Ward brought for exhi¬
bition an example of the ichneumon-fly, Rhyssa jJersuaso^'ia, $ ,
together with a photograph of the living insect. The speci¬
men was captured at Coventry j but whether that place was
its natural habitat is doubtful, as some packing straw from
Geneva was lying in the vicinity.
Beetle new’ to Science. — Mr. H. St. J. Donisthorpe ex¬
hibited, on behalf of Dr. Nicholson, Mr. Dollman, and himself,
examples of a species of new to science, and Scotch
specimens of Olojyhrum fuscum, its nearest ally, for com¬
parison. The new beetle was taken b}’ Dr. Nicholson and
himself in Wicken Fen under sedge refuse, and subsequently
by Mr. Dollman in some numbers in the same locality.
Mr. Donisthorpe pointed out the chai’acters in Avhich it
differed, and said he had described it, and proposed the name
for it of Olojihrum nicholsoni.
Edibility of Lepidopterous Larvae. — Mr. H. Eltringham
stated that in reference to his previous paper describing experi¬
ments on the edibility of certain lepidopterous larvae (Trans.
Ent. Soc., 1909, pp. 471-478), the caterpillars there referred to
as Boarmia rhomboidaria had proved to be Odontopera bideii-
tala. Further, that some of the moths had been bred from
larvae fed exclusively on ivy, and though similar larvae had, as
explained in the paper referred to, proved extremely distaste¬
ful to the lizards with which he had experimented, the moths
were found to be palatable.
His lizards having failed to survive the winter, he had sent
the moths to the Zoological Gardens, where Mr. Pocock had
( xxxii )
given one to a bird and two to some lizards, Lacerta viridis.
The result was described by INIr. Pocock as follows : —
“Commander J. J. Walker sent me, about a week ago, one
of the moths you are interested in, and I gave it to a bird, a
black-winged Crackle (Graculijira melanojjtera), a kind of
starling from Java. I have tested this bird before with
insects and have known him reject some which other birds will
eat, thus showing that he is to a certain extent fastidious and
not like some birds which will eat almost anything, He took
it at once and after pecking it and pulling it about for a little
time, swallowed it. He showed no sign of finding the moth in
any way distasteful — that is to say he did not once wipe his
beak, or shake his head, or behave as other birds do when they
taste anything nasty. Having confidence in his powers of
discrimination, 1 had no hesitation in concluding the moth to
be palatable.
“This was borne out by my experiment with the two yoii sent
me this morning. I threw one into a cage of green lizards.
One of them seized it at once and swallowed it in ten seconds.
I then threw in the other ho})ing that a second lizard would
take it, but the first was on to it in a moment and ate it as
soon as ever he had adjusted it in his mouth so as to swallow
it head first. I noticed that he squeezed some fluid out of its
posterior end, but even when this came into contact with his
palate and tongue he gave no sign of tasting anything
disagreeable. The lizards were Lacerta viridis. There was a
saucei’ half full of mealworms in the lizards’ cage, so there is
no reason to suppose that they were specially hungry.”
Mr. Eltringham added that he was greatly indebted to Mr.
Pocock for so carefully carrying out and recording the experi¬
ment. The result ajipeared to show that the distastefulness of
the larvae was due to the particles of the foodplant contained
in the digestive tract. The inedibility, though not in this case
due to peculiar properties elaborated by the metabolism of the
larva, might nevertheless, under conceivable circumstances,
have considerable importance for the larva itself. Distaste¬
fulness must have a beginning, and so long as it was a question
of surviving or being eaten, the fact rather than the cause of
the inedibility would be of first impoi’tance.
( xxxiii )
Paper.
Mr. E. Meyrick, B. A., F.R.S., communicated a paper entitled
“Descriptions of Micro- Lepidoptera from Mauritius and the
Chagos Isles.”
Wednesday, June 1st, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the Chair.
Address to the King.
The President proposed that an Address of Condolence and
Congratulation should be presented by the Society to His
Majesty King George V. on his accession to the throne,
and an Address of Condolence to Her Majesty the Queen-
Mother, Queen Alexandra. The proposal was seconded by
Mr. H. Rowland-Brown, one of the Hon. Secretaries, and
carried unanimously, all Fellows present standing.
“To THE King’s Most Excellent Majesty.
“ The Humble Address of the President, Council, and Fellows
of the Entomological Society of London.
“Most Gracious Sovereign,
“We, Your Majesty’s loyal and dutiful subjects, the
President, Council, and Fellows of the Entomological Society
of London, beg leave to tender to Your Majesty the ex¬
pression of our deep and sincere sympathy in the great loss
which you have recently suffered by the death of your august
Father, our late beloved Sovereign, King Edward VII. The
grief which has fallen upon your Royal House is shared with
the subjects of Your Majesty’s Empire by all friendly Nations,
and has cast its shadow upon the world at large.
“ Remembering the care which Your Majesty has con¬
tinually shown for all that tends to advance the best
interests of your people, and in particular for the progress of
scientific study, we feel that we have especial cause to join
with this expression of condolence in your sorrow our most
sincere and heartfelt congratulations upon Your Majesty’s
accession to the Throne of your Ancestors.
PROC. ENT. SOC. LOND., III. 1910.
c
( xxxiv )
“We pray that Your Majesty may long and gloriously
rule over a people united under the Ci’own in loyalty,
gratitude, and affection.
“Given under the Common Seal of the Society this fifteenth
day of July, one thousand nine hundred and ten.
“Frederick Augustus Dixey, M.A., M.D., F.R.S.,
‘ ‘ President.
“Henry Rowland-Brown, M.A.,
“James J. Walker, M.A., R.N., F.L.S.,
“ Secretaries.”
“To Her Most Excellent Majesty the Queen Alexandra.
“ The Humble Address of the President, Council, and Fellows
of the Entomological Society of London.
“ Madam,
“ We, the President, Council, and Fellows of the
Entomological Society of London, beg leave humbly to offer
Your Majesty the expression of our heartfelt sorrow at the
grievous loss sustained by Your Majesty in the death of our
beloved and lamented Sovereign Lord King Edward VII.
His memory will ever be gratefully cherished by all ranks and
classes both at home and abroad, and not least by those whose
devotion to the cause of learning and science has always been
accustomed to meet with his sympathetic recognition and
approval.
“Given under the Common Seal of the Society this fifteenth
day of J uly, one thousand nine hundred and ten.
“Frederick Augustus Dixey, M.A., M.D., F.R.S.,
‘ ‘ President.
“Henry Rowland-Brown, M.A.,
“James J. Walker, M.A., R.K., F.L.S.,
“Secretaries.”
Election of Fellows.
Mr. Arnold Whitworth Boyd, The Alton, Altrincham,
Cheshire; Mr. Emile Garcke, M.I.E.E., Witton House,
Maidenhead ; Mr. Henry Oliver Holford, Elstead Lodge,
Godaiming; Count Birger Morner, Consul for 11. M. the King
( XXXV )
of Sweden, Sydney, New South Wales; Mr. C. W. Mason,
S.E.A.C., St. Denis, Shaftesbury, Dorset ; Mr. Martin E.
Mosely, 13, Addison Road, London, W. ; Mr. Robert Tait,
junior, Roseneatb, Harboro’ Road, Ashton-on-Mersey, Cheshire;
Mr. Frank Wray Terry, The Planters’ Association, Honolulu,
Hawaiian Islands; Professor F. V. Theobald, M.A., South
Eastern Agricultural College, Wye Court, near Ashfoi’d, Kent ;
Mr. Charles Henry Rudge, Assoc. M. Inst., C.E., 15, Newton
Road, Bayswater, W. ; and Miss Carlotta Rudge, 15, Newton
Road, Bayswater, W., were elected Fellows of the Society.
Postponement of the Conversazione.
The President announced that the Conversazione, post¬
poned from Friday, May 27th last, by reason of the general
mourning for his late Majesty King Edward VII., would be
held during the forthcoming session on some date not earlier
than the last week in November.
International Congress of Entomology.
Mr. F. Merrifield, and Mr. R. Trimen, M. A., F.R.S., F.L.S.,
were appointed to act as additional delegates to the Inter¬
national Congress of Entomology at Brussels.
Exhibitions.
Rare British Beetles. — Commander J. J. Walker ex¬
hibited examples of Ceuthorrhynchideiis mixtus, Muls., and
Ceuthorrhynclms ]yiloselli(,s, Gyll., taken by him during May
last at Tubney, Berkshire.
Rare Hymenopteron. — The Rev. F. D. Morice showed a
specimen of the Pompilid Clavelia pompiliformis, Luc. $ (prob¬
ably the only fossorial wasp with pectinated antennae), taken
by him this spring, in the province of Oi’an, Algeria, from
which it was originally described.
Oviposition of Saw-fly. — The Rev. F. D. Morice also ex¬
hibited an example of the saw-fly Phymatocera aterrima, King.,
killed by ether in the act of ovipositing on a stem of Poly-
gonatum officinale (“Solomon’s Seal”). He showed also a
photograph of this curiosity, and others illustrating the same
subject — the structure of the saws, the egg-pouch made by
c 2
( xxxvi )
them, etc., and gave an account of the way in which the saws
are employed for the purpose. Instead of cutting vertically
the saws are turned sideways, a characteristic method employed,
as Mr. F. Enock pointed out by many of the Homoptera.
Both insects exhibited have been presented to the South
Kensington Collections.
Mr. C. 0. Waterhouse and Dr. T. A. Chapman also joined
in the discussion, and Mr, A. Sich mentioned that some
years since many larv'ao which appeared identical with those
of P. aterrvma, occurred on the same plant in his garden
at Chiswick.
Glow'-worm pupa. — Mr. H. Main brought for exhibition an
empty larva skin of a c? “ glow-worm,” Lampyris noctiluca,
with a living pupa, which was seen to be intermittently
luminous.
Melanism of Ematuega atomaria. — Mr. L. Newman
showed a case containing a long and varied series of Ematurga
atomaria bred from a melanic $ taken in cop. with a dark
typical S at Bury, Lancashire. It was noticeable that, of the
offspring, melanic and semi-melanic forms predominated.
Hybrid Lycaenidae. — Mr. Newman exhibited and ?
forms of Agriades thetis (bellargus), taken wild in North Kent,
June 1909, by Sergt. -Major W. Crocker, B.E. He said they
resembled examples taken on the same spot about sixteen
years ago by the late Mr. E. Sabine, and suggested they might
be hybrids thetis x coridon.*
Ova of Sesia andreniformis. — Mr. Newman further ex¬
hibited ova in situ of Sesia andreniformis ; and IMr. A. E.
Tonge a photograph ( x 26) of the same.
Cetoniidae from Uganda. — Mr. O. E. Janson showed a
remarkable gynandromorphous example of Goliatlms giganteus
and other Cetoniidae recently collected by Mr. E. Brown in
Uganda, British East Africa, including both sexes of the rare
Foimiasimus 7'ussus. Nearly all the species exhibited were
West African forms, proving the great similarity of the Central
* Mr. J. W. Tutt, who has since had an opportunity of considering
this exhibit, expresses his opinion that the butterflies are simply aberra¬
tions of A. thetis. The hybrid, =polonus, Zell., was recently discussed by
him, and a list of known examples given (cp. Proc. Ent. Soc. 1910, pp. viii,
ix).
( xxxvii )
African fauna, extending over a district of two to three
thousand miles across that continent.
Variation in Smerinthus. — The Rev. G. Wheeler brought
for exhibition a case containing many examples of Smerinthus
populi, taken in Lancashire, showing a wide range of variation,
bred from identical parents ; also a curious pale dwarf example
of S. ocellaius from the same locality.
Beetles from Crop of a Pheasant. — Mr. C. 0. Water-
house exhibited specimens of a beetle of the family Chryso-
melidae, Crosita altaica, Gebl., found by a poulterer at Bourne¬
mouth in the crop of a pheasant from Russia. He remarked
on the brilliancy of the metallic coppery-red and green colours.
It has been held by some that bright metallic colours were
warning-colours.
Agriades coridon Double-brooded. — Dr. T. A. Chapman
exhibited specimens of the spring emergence of double-brooded
Agriades coridon, taken in April and May last at various
places in the St. Tropez district of the Riviera, viz. le Canadel,
Pardigon, Gassin, and several different points near Ste. Maxime
(Var), at various dates from April 23 to May 11.
The specimens displayed considerable variation, but all
appeared to be of one race : very few showing any red colour
on the upper side.
The original locality where the species was abundant several
years ago near Ste. Maxime, now afforded only a specimen or
two, the Hippocrepis, that abounded there having nearly died
out, largely smothered by growth of Cistus mo7ispeliensis. In
each place where the species appeared Uippoci'epis was present
over areas of only a few dozen square yards, so that one
wondered how the butterfly could maintain itself. No llippo-
c/'cyjfs was seen except where the A. coi’idon was found. The
butterfly must be able to wander (often for some miles) from
one such locality to another, or it could hardly maintain its
existence as it does.
Living Larvae from the South of France, and Imago. —
Dr. Chapman also showed («) living larvae of Thestor hallus in
the last instar, feeding on flowers of Ulex em’opaeus ; {h) a larva
of Agi'iades coi'idon var. constanti, from eggs laid at Ste. Maxime
at the beginning of May, and now in the third instar ; and (c) a
( xxxviii )
living imago of Callophrys avis, Clipmn., a somewhat belated
specimen, emerged June 1st, 1910; the delay no doubt due to
an unsuccessful attempt at forcing in February.
Kare Insects from Portugal.— Dr. K. Jordan exhibited
a live specimen ( d ) of a species of Tnixalis obtained by him
at Portimao, South Portugal, and also showed some live larvae
and the cocoon of a moth, Diplura loti. The larvae were found
on Cistus in the Serra de Monchique, Algarve, South Portugal,
on May 13th, and ai’e being fed upon Ileliantheinum. They
resemble the caterpillar of Eriogaster lanestris so closely that a
generic separation is hardly justified.
A LyCAENID in attendance on an IIOMOPTERON. — Mr.
Hamilton H. C. DaucEread the following notes communicated
by Mr. J. C. Moulton. — “Some little time ago while working
out some species of Lycaenidae, I noticed an interesting illustra¬
tion in Col. Bingham’s volume on this family in the ‘ Fauna
of British India’ series (Vol. II, p. 287); this was an illus¬
tration of a Lycaenid — Allotinus horsjieldi, INIoore — attending an
Aphide. The drawing is about a half larger than natural size,
and seemed to me so clear, that I showed it to my two museum
collectors — Dyaks — telling them to look out for an example
of this in real life when collecting in the jungle. I further
explained that the species figured was a common one in
Sarawak, and that there was no reason why they should not
be able to observe this phenomenon if they waited and watched
the insect settle. For a month or more they were unsuccessful,
although collecting four or five days a week. At last, on
December 31st, 1909, one of the collectors, Bigi by name,
came back triumphant with two examples of Allotinus
nivalis, Druce (a smaller species than A. horsjieldi, but
nearly allied), together ■with an Ilomopteron. He told me he
had found the two butterflies hanging downwards from the
underside of a thin twig of a ‘ kapor ’ tree, facing towards
each other, some two inches apart ; between them were two
Homoptera ; and each butterfly was engaged in slowly stroking
the Homopteron nearest it with its tongue. (He illustrated the
action with his finger for my benefit.) The antennae were in
an upright position as shown in Col. Bingham’s illustration.
In reference to that picture it is interesting to note that the
( xxxix )
Aphide is there shown firmly held between the fore-legs of the
Lycaenid as if the ‘ milking ’ operation was not particularly
voluntary on the part of the Aphide. My Dyak, on the other
hand, was quite positive that the fore-legs were not used for
this purpose at all in the instance he witnessed ; and that the
Homopteron was quite free. With one sweep of the net he
captured the two Lycaenids and one of the Homoptera, the
other escaping. I inquired if he saw any other Homoptera
on the branch, but he replied not, remarking though that
there were plenty of ‘ sSmut ’ (ants) on it, though they did
not seem to disturb the butterflies. The note by Col. H. J. W.
Barrow, R.A.M.C., quoted by Col. Bingham (Z. c.) similarly
remarks on this point. He says : ‘ Its legs (A. horsjielcli) are
immensely long, and I discovered why. It settles over a mass
of Aphides, and then tickles them with its proboscis, just as
ants do with their antennae, and seems to feed on their
exudations. . . . But it would settle calmly over largish ants,
and did not mind one or two actually standing up and examin¬
ing its legs to see who was there. The ants did not attack it
in any way.’
“The branch on which these two specimens of A. nivalis
were taken was some four to five feet from the ground in old
jungle. In conclusion I may say that I know my Dyak well,
and am quite sure of the accuracy of his statements. Col.
Bingham remarks at the end of his note {1. c.) ; ‘ So far as I
know this is the first recorded instance of butterflies being in
attendance on Aphides as ants often are.’ And this i-einai'k
led me to think that perhaps another record of it might not be
without interest to entomologists.”
A Further Note on a Lycaenid in attendance on an
Homopteron. — “My first note dated January 3rd, 1910, was
entitled ‘ A Lycaenid in attendance on an Aphide.’ * Mr. Druce
very kindly communicated this note for me to the Entomo¬
logical Society on Feb. 2nd, at which meeting it was pointed
out by Mr. Druce and other entomologists, that the ‘ Aphide ’
exhibited was not an Hemipteron but an Homopteron ; and
in view of the fact that no instance is apparently known of a
* Subsequently altered to “A Lycaenid in attendance on an Homo¬
pteron.” [H.H.D.]
( xl )
butterfly in connection with an ITomopteron, members of the
Society were inclined — naturally enough — to doubt the ac¬
curacy of a single observation made by a native. However,
further evidence has since been afforded by the same native
observer which to my mind can leave no doubt as to the
accuracy of his observation.
“On December 31st, 1909, he brought in his first cajffuresof
2 A. nivalis with one Homopteron (the other having escaped).
About January 20th, 1910, he again brought in this species
of Allotinus together with an Homopteron, telling me that
he had watched them together for a long time, and that he
was convinced that the Lycaenid in no way held the Homo¬
pteron. I was particularly anxious to get at the truth on this
point in view of Col. Bingham’s picture which shows an
Aphide apparently being held by the Lycaenid. On February
2nd, 1910, he again brought in an Allotinus sp. nr. nivalis
which he had found in similar attendance on the same*
species of Homopteron (which insect he also brought in).
As before, he tells me that he -watched them for some time to
see if there was any restraint on the Homopteron, but he was
quite sure it was free.
“ In I’eply to a suggestion that the insect he caught may not
have been the same as that which he saw in connection with
the Lycaenid, he was most emphatic on the point, and con¬
sidering that he has brought in the same species on three
separate occasions, I cannot but feel that there is no doubt
on this score.
“ About February 10th he brought me an example of
Allotinus horsfieldi which he had taken in attendance on a
‘ Heteropterous larva h ’ (I would ask some entomologist to
verify (1) this title for me from the material sent).
“The point about this last observation is that he reports
having noticed large numbers of this insect on the tree, in
company with many ants which latter appeared to be bother¬
ing the Lycaenid, although he stuck to his work in spite of
the ants, until the collector’s net removed him. In the
* I say same species of Homopteron, for they appear to me to be the
§ame under a lens, and from memory I should say the same as that sent
to Mr. Druce in January last for exhibition.
( xli )
previous observations none, or but very few, of the Homoptera
were seen on the same tree.
“ I send for exhibition another example of the Homopteron
which was taken with A. nivcdis and four examples of the
‘ Heteropterous larvse ’ which were taken off the same branch
as A. horsjieldi.
“ As these four observations have all been made in a period of
about six weeks, out of which the observer was collecting for
thirty days at most, owing to bad weather, one may suppose
that this phenomenon is by no means uncommon, and if other
collectors in the Tropics would look out for it, more interesting
occurrences with different species would probably be recorded.”
Error in the account of Mr. Leigh’s Breeding Experi^
MENT WITH Charaxes (Pi’oc. Ent. Soc., Lond., 1909, p. xlix).
Professor Poulton, F.E.S., who was unable to be present,
sent the following correction of an unfortunate error which
has crept into the Proceedings of last year : —
“ Mr. Leigh is stated, on p. xlix, to have exhibited the $
parent and 21 specimens of the offspring of Charaxes zoolina
neanthes.
“ On the other hand it is also stated that ‘ the ova deposited
by the zoolina form of the $ . . . pi’oduced 4 cJ cj s and 2 $ $ s
like the parent, and 15 d c? s and 9 $ $ s of the neanthes form,’
or 30 offspring in all. I have examined the specimens, which
are now in the Hope Department, and find 21 offspring, but
also observe that there are among them 6 and not 15 d (? s of
the neanthes form. It is to be hoped that this correction will
meet the eye of any naturalist who has been interested in the
published statement.”
Papers.
The following papers were read : —
“ Mr. A. D, Millar’s Experimental Breeding of Euralia ”
by Roland Trimen, M.A., F.R.S., F.L.S.
“Notes on the Scoliidae,'’ and “New Fossorial Hymeno-
ptera from Australia,” by Rowland E. Turner.
“On the position of the Rhopalosomidae, with description
of a second new Species,” by Claude Morlet, F.Z.S.
( xlii )
“Descriptions of Malayan Micro-Lepidoptera,” by Edward
Meyrick, B.A., F.R.S., F.Z.S.
“ On the Specific Distinctions between Acraea lycoa, Godt.,
and Acraea johiostoni, Godm.,” by Harry ELTRiNonAM,
M.A., F.Z.S.
Wednesday, October 5th, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the Chair.
Address to the Kivg, and Queen Alexandra.
The Secretary read the following reply to the Address to
the King, and to Queen Alexandra : —
“ Home Office, Whitehall,
“ 8th Atigust, 1910.
“ Sir, — I am commanded by the King to convey to you
hereby His Majesty’s Thanks for the loyal and dutiful Address
of the President, Council, and Fellows of the Entomological
Society of London, expressing their sympathy with His
Majesty on the occasion of the lamented death of His late
Majesty King Edward the Seventh, and congratulation on
His Majesty’s Accession to the Throne.
“ I am to say that the expression of sympathy with Queen
Alexandra has been laid before Her Majesty, who desires me
to convey to you Her Thanks.
“ I am. Sir,
“ Your obedient Servant,
“Winston Churchill.
“ The Secretary,
“ The Entomological Society of London,
“11, Chandos Street, Cctvendish Square, IV."
C onversazione.
The President announced that the Conversazione postponed
from last May was unavoidably postponed until next year.
Seal of the Entomological Society of London.
The President exhibited and passed round for inspection
an impression of the new seal of the Society, prepared from
( xliii )
a design made by Professor Selwyx Image, M.A., Slade Pro¬
fessor of Fine Art in the University of Oxford, and a Fellow
of the Society.
On the motion of the Peesident a cordial vote of thanks to
Professor Image for liis services was passed unanimously.
International Congress of Entomology.
Dr. M. Burr sent for exhibition a photograph of the
members attending the International Congress of Entomology
at Brussels in August last.
Election of Fellows.
Mr. George William Yitalli de Rhe Philiee, of the
Public Works and State Railways Department of the Govern¬
ment of India, Calcutta ; and Dr. Charles Ernest Lakin,
M.D., M.R.C.S., F.R.C.S., of 2, Park Crescent, Portland
Place, W., were elected Fellows of the Society.
Exhibitions.
Rare British Beetle. — Mr. G. C. Champion brought for
exhibition two living examples of Melanophila acuminata,
captured at Woking on September 1 7th last.
Rare British Hemiptera. — Mr. E. A. Butler exhibited
specimens of thi-ee species of I’are British Hemiptera, viz.
{a) Mesovelia furcata, M. ct R., and (6) Cicadxda cyanae, Boh.,
both from leaves of Potamogeton natans in Epping Forest ; and
(c) Cyrtorrhinus geminus, Flor., from Broxbourne — a recent
addition to the British List, only two British specimens being
at present known. He also exhibited living examples of
Mesovelia furcata, and drew attention to the extraordinarily
rapid movements of the species, which quite defy the eye.
Rare Australian and British Beetles. — Commander J.
J. Walker exhibited (a) a “co-type” of Austrostglops gra-
cili])es, Lea, from Bridgetown, W. Australia ; [h) a series
of Carabus violaceus, L., var. e.vasperatus, Curtis, taken by
Mr. F. C. Woodforde at sugar at Bude, North Cornwall,
where this well-marked race appears entirely to replace
the ordinary form of violaceus ; and (c) four examples of
( xliv )
Ilaemonia appendiculata, Panz., taken in an affluent of the River
Cherwell, near Kidlington, Oxon., August, 1910; also {d) a
specimen of Cryptophagus suhdepressus , Gyll, from Wytham
Park, Berks, with the right antenna duplicated from the third
joint.
Life Tenacity op Abyssinian Beetle. — Mr. W. F. H. Rosen¬
berg showed a living beetle of the genus AlindiHa, received in
a collection of preserved Coleoptera fi’om Abyssinia. The
specimens were rolled up in small paper cylinders, and when
they were opened the insect was found to be alive. The
collection must have been despatched from Abyssinia at least
a month before the opening of the package. The insect has
the habit of simulating death on being alarmed, which probably
accounts for the collector having packed it up without noticing
that it was alive.
Living Ants and their Nests. — Mr. H. St. J. Donisthorpe
exhibited two nests of ants to illustrate his observations on
the founding of nests of Formica ritfa by a ^ of that species
in nests of Formica fusca, and also made some remarks on
what he had observed in nature, both with F. rufa and F.
exsecta. He had observed a 9 '^’ufa in Pai’khurst Forest on
May 15th, running about near the entrances to a fusca nest.
The 9 fought with some of i\\e fusca, and eventually entered
one of the openings in the ground used by the fusca. On
May 27th, in company with Mr. E. R. Bankes, he found a small
nest of F. exsecta at Bournemouth, which on being investigated
was found to contain many 9 ? of F. fusca.
Exhibit Nest No. 1 consisted of 9 9 oi F . rufibarhis y . fusco-
rufibarhis, and a 9 of rufa. This was exhibited to the
Society on March 2nd, when the rufa 9 (which had been
obtained at Nethy Bridge in May 1909) had just been accepted
by the fusco-rufibarbis 9 (sent to him by Mr. Keys from
Whitsand Bay in July 1909) after having been introduced
into their nest. From March 7th to April 4th the 9
eggs, and larvae hatched by May 1st; these were fed and
attended to by the 9?- June 1st many changed to
pupae, nearly all being naked. By June 20th some of the
pupae hatched, and on July 20th all had hatched. Unfor¬
tunately they all died, the 9 ? uot having sufficiently removed
( xlv )
the pupal skins. These young rufa ^ ^ were exhibited
mounted on card.
Nest No. 2 consisted of a mixed nest of Formica rufa and
F. fusca. This nest which was very small but built of F.
rifa materials, was dug up in Parkhurst Forest on August
21st, 1910. It was found to contain 1 rufa $, some rufa
5 ^ , and many fusca 5 ? > ^ number of cocoons — these
as they hatched proving to be rufa 5 5- It is therefore clear
that this o'ufa $ must have founded her nest in a fusca nest-
The fusca were underground with the rufa ? and cocoons,
but no fusca $ was present.
PlERIDAE FROM FRANCE AND ItALY. — Mr. A. H. JoNES
showed a series of Pieris manni, males only, from the valleys
of the lower slopes of Mont Canigou, near Veimet-les- Bains,
taken at the end of June last, with examples of P. rapae,
and P. ergane from Italy for comparison.
Mr. P. J, Barraud brought for exhibition a case containing
Pieridae from the neighbourhood of Formia, Central Italy,
including Pieris rapae, L., and var. metra, Stephens ; Pieris
manni, Mayer, and var. rossii, Stefanelli, and ab. erganoides,
Stefanelli ; and Pieris ergane, Hiibn. With these he showed
also enlarged photographs of male and female examples of
the summer forms of the above-named species to illustrate
the difference in the character of the markings and shape of
the wing ; and enlarged photographs of the male genitalia of
each, exhibiting the difference in the shape of the clasper.
Butterflies from the Abruzzi. — The Eev. G. Wheeler
gave an account of an entomological excursion made by him
in the Abruzzi, and exhibited two drawers containing speci¬
mens taken at Assergi, Sulmona, Aquila, Roccaraso, Palena,
Scanno, and Villalago ; and a few also from Subiaco in the
Latian Apennines. Pieris ergane was found to be somewhat
widely distidbuted, having been taken at Sulmona, 1,300 feet ;
above Subiaco, about 2,000 feet ; and at Roccaraso, over
4,000 feet. There was also a series of very strongly mai'ked
Hesperia carlinae var. cirsii, some of Avhicli approached somewhat
closely to H. cynarae. Polyommatus eros, of a much deeper blue
than is usual in the Alps was abundant at and above 4,000 feet,
i.e, nearly 2,000 feet lower than is normal in the Alps ; other
( xlvi )
unexpected captures were F. amanda and Limeniiis Camilla,
both at over 4,000 feet ; one of the former was almost as
large as the Tyrolean form, and the latter Avere all exception¬
ally large, the species being no doubt single-brooded at that
altitude. By way of contrast a second-brood S from Assisi
was shown, these being very small. P. ineleager, which was
very common, showed three very different shades of blue,
though all came from Boccaraso, and P. icarus was of the
same deep hue as at Assisi, some of the having black
marginal spots on the upperside of the hind-wing. A. medon
{astrarche) had the unusually large, the 59 being small,
with large and brilliant orange spots. P. coridon was very
pale in colouring, cori-esponding with Zeller’s description of
var. apennina, but not with his type specimens. As a con¬
trast, two other forms from the Apennines, the much bi-ighter
blue from Assisi, and the somewhat leaden-looking form from
Fiesole Avere shown. The other species exhibited were :
Erimiys lavaterae (very pale), E. althaeae and E. alceae ; Hesperia
alveiis, //. carthami (very small), II. onopordi and II. serra-
tidae ; Ileodes virgaureae (large and bright, only), Loweia
gordiiis, L. dorilis, Rumicia p)ldaeas var. eleus, Lycaeaa avion,
Aricia eumedon (very small), Polyommatus escheri (small and
bright), P. Jiylas, P. thetis {hellargus), Plebeius aegon (small),
and P. argyrognomon, Everes alcetas, Cnpido minwms (avIucIi
seems to be scai’ce), and Celastrina argiolus', Klugia spini,
Nordmannia ilicis and i\\ acaciae (the commonest of the
Theclids) ; Ay)oria crataegi, Pieris rapae (from the same
ground at Subiaco as P. ergane, and nearly twice the size),
P. najii var. napaeae (very large, and as strongly marked
on the upperside as P. manni var. rossii), Argynnis aglaia
and A. niobe (from Boccaraso), A. adippe var. cleodoxa (a
very fine specimen from Subiaco, not taken in the Abruzzi),
Issoria lathonia (very small), Melitaea phoebe (from Palena,
over 4,000 feet, small and dark, contrasted with Spanish-
looking specimens from Assisi), M. didyma, large and pale,
II. parllienie (from Boccai’aso and Palena, single-brooded of
course at this eleAmtion, and coming in appearance between
the two broods at Assisi) ; Pararge maera (not of the adrasta
form), with a fine 5 aberration fi’om below Scanno ; Coeno-
( xlvii )
nympha pamphili^s, C. arcania (contrasted with the large var.
instihricci from the N. Italian Alps) ; and Melanargia galatea
(showing in the lowland specimens a strong tendency to
obsolescence of the underside markings, in contrast to the
somewhat strongly marked mountain forms). Besides these
was a Pontia dapliclice $ from Home, showing distinct indica¬
tions of a black spot near the middle of the inner margin of
each fore-wing on the upperside.
Rare British Xylophasia. — Mr. J. W. Tutt brought for
exhibition a fourth British example of Xylophasia zollikoferi
sent him for determination, taken in September, 1905, at
Norwich, by Mr. Plankett. He said he had dealt with
the first two examples captured in Britain at length, in his
“ British Noctuae and their Varieties,” i, p. 71. The first was
captured in October 1867, by Mr. Harding at Deal ; the
second by Mr. Tait .at Inverurie, in September, 1871 ; the
third by I\Ir. Lofthouse at Linthorpe, Middlesborough, Sep¬
tember 2nd, 1903. The first of these is in the Doubleday
collection. The preceding British examples have been of the
pale form (ab. pallida, Brit. Noct., i, p. 72). Mr. Plunkett’s
example is of the remarkable radiate form, represented in
Herrich-Schaffer’s “ Schmett. v. Europa,” Fig. 104. There
are only half-a-dozen examples of this species in the British
]\[useum Collection, all pale, and all males. The radiate form
figured by Herrich-Schaffer is a $ , so also is Mr. Plunkett’s,
and this suggests that the difference is sexual. It is really a
very rare species — as rare on the Continent as in Britain, on
this side of Hungary; getting more frequent in the Urals, and
extending into Asia to Turkestan, but evidently nowhere com¬
mon, so far as is known. We have no knowledge of its life
history. Several entomologists have remarked the superficial
resemblance of the pale forms to Nonagria arundinis, and for
this reason there may be others undetermined in British
collections.
Southern Forms op Agriades coridon. — Dr. T. A. Chap¬
man exhibited several cases containing series of Agriades cori-
don, var. meridional is, Tutt, 6 { = constanti, Reverdin), and
made the following remarks upon them ; —
“It is probably accepted now that this form is double-brooded.
( xlviii )
though even Mr. Tutt was sceptical until I reared larvae from
eggs laid in April, which were full-grown at the end of J une.
Unfortunately, taking these from home with me, they all died
of starvation, as their food-plant was unobtainable.
“ This year, however, I obtained eggs in the spidng from
Ste. Maxime specimens, and have reared the specimens now
exhibited. These emerged during August, and were dealt
with by Mr. A. E. Tonge, who kindly took charge of the
pupae when I went to Switzerland to find the larva of
Latiorina {Lycaena) orbitulus.
“ He endeavoured to obtain eggs from some specimens, hut
owing to an accident was apparently unsuccessful, though
in mid-September in looking over the plant on which specimens
were caged, I noticed several leaves eaten precisely as is done
by first-stage larvae of A. coridon. As the plant was indoors
and not accessible to any other flies {P. icarns, or anything
else), I have little doubt that eggs were laid, and one at least
hatched, the others being destroyed by a predaceous enemy.
“ In any case, making every allowance for the mild Eivieran
winter, there can be no doubt that the larvae do some feeding
in autumn and probably even duidng winter, in order to
emerge in April. It certainly seems very improbable that
they pupate in autumn and hibernate as pupae, but this is a
possibility to be kept in view in further investigation — most
likely the life circle is much the same as is that of A. thetis
(bellaryus) with us. Whether this summer brood has been
captured before or not, though one can hardly doubt that it
has, we have no record of it, or of whether and how it differs
from the spring form.
“ The specimens exhibited afford at least a partial answer
on this point. They are on the whole larger. The males have
no trace of a discoidal spot : instead of only a few, nearly three-
fourths of them are obviously of the ab. suavis, i. e. have
orange spots on the upper surface, and this, although the
margin of the hind-wing, is less darkly marked than in the
spring brood. Beneath, the orange spots of the hind-wing are
larger and of more bright and intense colour, and the line
of chevron markings on the fore-wing is generally a nearly con¬
tinuous dark band, as it is only occasionally in the spring brood.
( xlix )
“ The females differ from the spring form, on the upper
side, by the greater range of the orange lunules, which
extend all round both wings, though the apical two or three
ai-e often indistinct by being pale or small, but even so are
almost always present. The discoidal spot is small and usually
very obscure, in two (out of twenty-three) instances only, with
a trace of white border. Beneath, the surface is on the whole
darker than in the first brood, the chevron line of the fore-wings
is a black band, and in most specimens the orange spots form a
complete series round both wings, the apical spot only being
usually doubtful.
“ In both sexes the frequency of extra spots, and spots length¬
ened, or coalesced, is rather greater than in the spring brood.”
Mr. J. W. Tutt who exhibited long series of the spring
brood of the meridionalis form from Nimes, Hyeres, Ste. Maxime
Pardigon, Draguignan, etc., said the Riviei’an meridionalis
race of Agriades coridon was exceedingly interesting, as pro¬
viding the only case recorded where the species is certainly
double-brooded. This race appeal’s to extend from Kimes
almost to Genoa, perhaps beyond. Its superficial characters
are sufficiently striking to give it racial value, and consist
more particularly in the obvious features of (1) a peculiar
dull silvery-blue ground-colour on the upper-side of the
(2) a very wide dark outer-marginal band in the (3) a
strong tendency to the development of oi’ange lunules on the
upper-side of the marginal spots on the hind-wings also in
the S S- The upper-side of the $ ? is usually without any
blue scaling and deep fuscous brown in colour. The under¬
side of the c? is variable, extending from whitish to dark
grey and hence resembling that of A. thetis, whilst that of the
$ is very dark grey-brown, the ocellated spots in both sexes
being particularly large and Avell-developed, and the mai'ginal
lunules of the fore-wings strongly blackish-grey.
This meridionalis race has been divided into two forms, chieffy
chai’acterised by the intensity of the ground colour of the under¬
side, especially in the The form with pale under-side has
been described as rezniceki by Herr Bartel, his typical specimens
having come from Bapallo (Italian Riviera), June Ist-lOth,
1901, etc.
PROC. ENT. .SOC. LOND., IV. 1910. D
( 1 )
Quite recently (Bull. Soc. Lepid. de Cien^ve, Vol. ii, fasc. 1.^
June 1910) Dr. J. L. Keverdin has described, under the name
constanti, a section of the meridiotudis race on the strength
of the under-side of the d is, so far as can be seen in
the specimens under observation, the sole difference between
rezniceki and constanti.
As regards the specimens exhibited (captured in the spi-ing)
both forms appear to occur on the same ground in the districts
betAveen Hyeres and St. Raphael, with a large proportion of
dark-coloured under-side examples. Whether anywhere in the
area covered hj meridionalis the form rezniceki {$$ with pale
under-side) and the form constanti (d d 'with dark undei'-side)
become quite locally racial has not yet been determined. It
would be interesting for solving this problem to state, when
dealing with the specimens from any particular place, on what
geological formation the examples ax’e captured.
It is peculiar that M. Charles Oberthiir, in his excellent ork
(Lep. Comp., fasc. iv., p. 281), distributed so recently as early
August last, notes a connection between what may be called a
hypothetical var. apennina from Florence and rezniceki, Bartel^
but not between rezniceki and constanti, really, with very slight
differences, the same race ; hypothetical apennina because it
is clear that the apennina spoken of by M. Oberthiir are the
usual summer form taken in the Apennines, and have little,
if anything, in common with the specimens in the British
Museum Collection, named apennina by Zeller.
Protective attitude of a Mantis from Borneo. — Mr. G.
Meade-Waldo read the following note received by him from
Mr. J. C. Moulton, and exhibited an example of Ilymenopus
h icornis : —
“ This curious Mantis was given to me on June 2nd, 1910,
by Mr. E. Parnell, of the Sarawak Government Civil Service,
who found it in his garden. I kept it alive for twenty-three
days, and was thus able to make some observations on its
customary attitudes.
“ Generally it was to be seen in the usual ‘ praying ’ atti -
tude, i.e. with head, thorax and fore-legs up- raised. But one
day, on disturbing it, I noticed an improvement on this
position, which gave it the most remarkable resemblance to
( li )
some orchid. This was brought about by a very unnatural
(and one would think extremely uncomfortable) arrangement
of the legs. The head and thorax were raised, with f ore-legs
tucked up under the head as usual ; abdomen turned upwards
and end curving over, and immediately under it the left hind¬
leg, turned over so as to fully exhibit the flat lateral expansion
of the femur. On looking down upon the insect the left
middle-leg appeared under the head and rather to the left, the
right middle-leg immediately under the head, and the right
hind-leg a little to the right corresponding to the position of
the left middle-leg. Each of these three was so turned in
order to give full effect to the petal-like effect of the lateral
femoral processes.
“I have set the insect so as to show this curious position,
the particular specimen being kept for twenty-three days in a
flower-pot in company with Lepidopterous larvae which were
feeding on some low plant. I have heard of another example
which was kept in a glass jar for three weeks in this country
without giving it any possible food.
“ This comparative longevity and tenacity of life, together
Avith its sluggish habits, ha\'e made it necessary to produce
this wonderful protective deAuce in order to escape the
numerous enemies of Mantidae."
Commenting on this exhibit, Mr. Meade-Waldo said that
the insect sent was in the larval state. Wood-Mason (Proc.
Ent . Soc. Lond., 1877, p. xxix), mentions the extraordinai’y
likeness of the larval II. hicornis to blossoms, and describes
the attitude. He had received tAvo larvae from Assam, the
smaller of Avhich was rose-pink, and the larger one white,
evidently similar to the specimen now exhibited.
At that time the insect AA^as only known from JaA'a. There
are also specimens in the British Museum from the Khasi
Hills, Malacca and Selangor. Evidence has been brought to
prove that insects are attracted to the coloui-ed Mantidae as
they are to floAvers, and according to A. Pv,. Wallace, Avho sup¬
plied the evidence required, a small Mantis, exactly resembling
a pink Orchis-flower, was shoAvn to Sir Charles Dilke in Java.
This species not only attracted insects, but the kinds of insects
(Butterflies) which it allures and deA'ours Avere mentioned.
D 2
{ lii )
No previous writer appears to have observed that this
attitude is more rarely adopted than the usual “ praying ”
attitude.
Improved Photographic Process for Lepidoptera, etc. —
Mr. E. Dukinfield Jones brought for exhibition lantern
slides of Lepidoptera in natural colours, photographed by the
Dufay Dioptichrome process. The advantages over the Auto¬
chrome plates are the greater transparency and the facility of
working ; they are in fact very little more trouble than an
ordinary negative. The ti’ansparency is by no means perfect,
and the reds do not come out as well as one could wish ; but
on the whole the results are remarkably good.
Four slides were shown : —
(1) Containing Gonepteryx clorinde, Moipho ega, Catopsilia
philea, C. argante, Pajnlio ancanius, Morpho acJiilles.
(2) Some species collected at Castro Parana, South
America.
(3) Caligo arispe, wwdiev-Bide •, J natural size.
(4) Catocala ni'pta, slightly reduced.
Papers.
Miss Margaret E. Fountaine communicated a paper on
“ Descriptions of some hitherto unknown, or little known
Larvae and Pupae of South African Rhopalocera, with notes
on their Life Histories.”
Professor A. Jacobi communicated a paper, “ Remarks on
the Cicadoid Genera Lembeja, Dist., and DrejmnopsaJtria,
Bredd.”
Mr. Arthur M. Lea communicated a paper, “ On a new
Genus of Htylopidae from Australia.”
Wednesday, October 19th, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the Chair.
Election of Felloios.
Di\ A. Feynes, M.D., of 61, East Colorado Street, Para-
dena, California, U.S.A. ; Mr. Thomas Henry Geary, of
( liii )
Enderby, Leicestershire ; and Mr. Edward Barton White,
M.E.C.S., L.R.C.P., of the City Mental Hospital, Cardiff, were
elected Fellows of the Society.
Ohituary.
The decease was announced of Mr. Oliver C. Goldtiiwait,
and of Mr. W. A. Luff, Fellows of the Society.
Exhibitions.
Abnormal Beetles. — Mi-. A. M. Lea sent for exhibition
two interesting beetles of the genus Lissotes, which he had
presented to the British Museum (Natural History) Terato-
logical Collection ; L. curvicornis, Boisdv., var. 3, with an
additional leg jutting out from the left front coxa ; and L.
punctatus, Lea, an hermaphrodite having the left side $ and
the right $.
New British Braconid. — Mr. H. St. J. Donisthorpe
exhibited an example of Helcon ruspator, L., a Braconid new
to Britain, taken at Cannock Chase on the 16th July last, in
a cell of Strangalia ^-fasciata in a fallen birch tree, and an
example of the host captured at the same time. He pointed
out that this very fine addition to the British List is recorded
as parasitic on the same beetle on the Continent.
New Forms of Melitaea aurinia. — Mr. P. J. Barraud
showed examples of two new forms of Melitaea aurinia from
Italy, (a) var. aurunca, Turati, from the Aurunci mountains,
southern central Italy, first discovered in May 1909, by
Signore Orazio Querci, of Formia, and named by Count Turati
of Milan. The most striking feature is the wide black median
band on the upper-side, contrasting with a rather pale ground¬
colour. (h) var. comacina, Turati, from above Como, north
Italy, the examples given the exhibitor by Count Turati.
Summer Butterflies from Algeria. — Mr. A. E. Gibbs
brought for exhibition a case of butterflies containing a
representative collection of the 28 species met with by
him at Blidah and Hammam E’lrha, etc., during an
entomological excursion to Algeria made at the end of May
and the beginning of June this year. The weather was
extremely bad throughout, being cold, wet and windy ; the
most interestiug species taken in the first-mentioned locality
were Euchloe eupheno, and Coenonympha arcanioides ; in the
latter Dry as pandora, and Melitaea aetheria, var. algerica.
Mimics and Models from South America. — Mr. W. J.
Kaye exhibited remarkable examples of wasp-like and beetle¬
like Syntomidae, with their models, the w'asps and beetles from
British Guiana, S.E. Brazil and Venezuela, many having
been observed personally by himself. Attention was par¬
ticularly drawn to the wonderfully close habits of the one
to the other, so much so that it was impossible in some
instances to distinguish the moth from the model until it
settled. This was the case with Pseudosphex noverca and the
wasp Zethus binodis which Mr. Kaye had caught on Ageratum
fiowers at Fernandes Pinheiro in Parana. It was true that
there was a better model where the likeness was even
more complete in the Vespid Polyhia nigra, and Mr. A.
Bertoni had taken the Pseudosphex with the Polyhia in Para¬
guay. A new species of Pseudosphex was taken with the
Vespid IfeganthojMS cassununga. These two insects so closely
resembled one another that even at a distance of eighteen
inches or less it was not an easy matter to distinguish them.
Besides the general appearance in build, the wings, and the
markings, the antennae were remarkable in having exceed¬
ingly fine white thread-like ends, which became invisible when
the insect settled on the white Ageratum flowers, so that only
the black pectinated portion was visible, wdiich exactly re¬
sembled the antennae of the wasp. Many species of Alacro-
cneme had developed this remarkable character in the antennae.
Several, if not all the species of this genus also carried their
hind-legs extended out behind them wKen in flight, and this
gave them an extraordinarily close resemblance to the Pom-
pilidae, which had the same habit. The species Macrocneine
lades and M. leucostigma had the hind tibia and tarsus
heavily clothed with black hair, and this accentuated the
likeness to the heavy black legs of various species of Salius.
The heavy flight of such species as Macrocneme lades and
M. leucostigma, combined with other characters, made them
exceedingly close mimics of various species of Salius among
the wasps.
( )
Other species of Syntomidae exhibited were the following ; —
From Potaro River, British Guiana, Isanthrene melas,
Trichura inathina, Sphecosoma angustatum, and Sphecosoma
testaceiini, both the latter found with a Pompilid wasp of the
genus Pseudagenia, also shown. Gorrebidia n. sp., with the
Lycid beetle Calopteron serratmn. Pterygopterus clavipennis,
with the Pompilid wasp Salius kirbyi.
From Castro Parana, Rhynchopyga meisteri, with one of the
Braconidae. Paraethria triseriata, with the Coleopterous
Astylus antis. Although the last did not look much like the moth
when dead in a box, on the wing it was wondei’fully protected.
From Guaruja, near Santos, were exhibited two undescribed
species of Trichura which resembled moths in a remarkable
way, but the wasps they imitated were, unfortunately, not
secured. The very unmothlike habits of moving the abdomen
backwards and forwards, the waving of the antennae, and the
vibrating of the wings was commented on. The remarkable
tail-like appendage in the A) one of the species, as in the
well-known species Trichura cerberus, represented undoubtedly
the ovipositor of an ichneumon. The male sex being the better
protected, if protection was the explanation, was very excep¬
tional, as in the vast majority of cases the female was the
better protected. From Caracas, Venezuela, were shown the
Syntomid Macrocneme lades, with a Pompilid wasp of an
undescribed species. The two insects had been caught flying
together.
In the discussion which followed, the President said that
-cases of close similarity with insects of different Orders were
always welcome, and that in this instance Mr. Kaye had
shown that the resemblance extended not only to outside
appearances, but to the habits of the several mimics and their
models. Arguments based upon cabinet specimens alone as to
the supposed i-esemblance of originals in the field were to be
accepted with caution.
Mr. J. W. Tutt remarked that the exhibit was of great
interest, particularly in the fact that the specimens exhibited,
that resembled each other, were caught at the same time and
in the same place by the exhibitor, and not only bore a great
.superficial resemblance to one another, although belonging to
( Ivi )
diffei'ent Orders, but also in the fact that they still more
resembled each other in their habits, either in movement or
rest. In some instances, this general resemblance was so
great that it was dilEcult at a casual glance and at a short
distance to distinguish them at all accurately, even when dead
in the box and removed from those surroundings which aided
the greater resemblance, as Mr. Kaye had pointed cut,
presented by them when alive in their natural habitats. He
trusted that when Mr. Kaye illustrated these, as he hoped he
would, that he would do so in those positions in which tlie
resemblance struck him as being most marked in nature, as in
most cases the resemblance was entirely lost when the insects
were set. He observed that a case of two insects of quite
different appearance when looked at even casually with their
wings spread, but of remarkable general similarity when at
rest on flowers of thyme, had come under his notice in the
Sarnthal in August 1909, the one the common steel-blue form,
with yellow spots and yellow abdominal belt, of Anthrocera
ephialtes, the other a Hymenopteron of similar steel-blue colour
and yellow abdominal bands and spots. It was really in¬
credible that two such different insects could appear so similar
as they did in their normal position of rest.
Mr. C. J. Gahan remarked that, with the exception of one
or two, which were amongst the most beautiful of their kind
he had ever seen, the examples of mimicry shown by Mr.
Kaye were not more striking or more convincing than
many others already known, in which the facts in regard
to the habits of the insects, and their very close and deceptive
I’esemblance when seen under natural conditions, had been
observed and put on record. He thought, therefore, that the
eulogistic terms in which Mr. Tutt had referred to the
exhibition, as if facts of that kind had only now for the first
time been brought to notice, were scarcely justified. The
exhibition was no doubt highly interesting, but its especial
interest, in his opinion, was to find the field naturalist confirm¬
ing those cases of “ museum-made ” mimicry which appeared
to excite Mr. Tutt’s scepticism.
]Mr. G. C. Champion said that almost every case exhibited
had already been observed and described by the various authors
( Ivii )
interested with the insects of this particular region : and
the Eev. F. D. Morice suggested that, speaking from a Hvmen-
opterist’s point of view, he would not have the least difficulty
in distinguishing the Lepidoptera from the Pompilids, and
that in the Sahara region, where some models abound, none of
the Lepidoptera exist at all.
Mr. A. Sicu, Dr. T. A. Chapman, and other Fellows also
joined in the discussion.
Variation in Xanthia ocellaris. — Mr. E. D. Nevinson
showed bred series of Xanthia ocellaris, developing three
distinct aberrant forms, and examples of X. fulvarjo, and X.
yilvago for comparison, the exhibit demonstrating the apparent
transition from one species to the other through the typical
and variant forms. The only other series bred from British
ova by Mr. Mills in 1908 displayed no variation of any kind,
and were all typical specimens.
British Anthrocerids. — The Hon. X. C. Eothschild e.x-
hibited a number of Anthrocerids captured in Great Britain,
and called attention to some remarkable specimens secured at
Ashton Wold, Oundle, which belonged to the form known as
Anthrocera hippocrepidis. The exhibitor also showed some
enormous specimens of A. filipendulae from the same locality,
and pointed out that this large race had apparently exter¬
minated A. hippocrepidis in a locality where that species had
only recently appeared.
Commenting on Mr. Eothschild’s exhibits, Mr. J. W. Tutt
said that Anthrocera hippocrepidis, Stphs., was the most elusive
and least known of the British species of the group. It bore
certain superficial resemblances to Anthrocera irifolii-minor
and A. filipendxdae, the particularly being prone to smaller
size and five-spottedness, by losing the 6th (lower outer) spot
on the upper-side, the larger and, in the most developed
specimens, strongly six-spotted and scarcely, if at all, different
superficially from A. filipendidae. Like Mr. Eothschild, he
also found the insect confined to rough pastures, and also with
a tendency to die out in one spot and appear in a similar at no
great distance. It was usually on the wing in June, early or
late, according to season, but, in the same season well ahead
of A. filipendulae. He had already included details of its life-
( Iviii )
history and habits in his “ Nat. Hist. Brit. Lep.,” vol. i, pp. 532-
538. The two other sets of Anthrocerids exhibited by Mr-
Rothschild he thought were A. filipendulae. Dr. W. Bateson,
F.R.S., had examined the genitalia and pronounced A.
hippocrepidis as being nearer A. filipendulae than A. trifolii.
Papers.
Sir George H. Ivenrick communicated a paper “ On some
Rare and Undescribed Butterflies from Dutch New Guinea.”
Mr. A. E. WiLEMAN read a paper on “New Species of
Heterocera from Japan.”
Wednesday, November 2nd, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the
chair.
Election of Fellows.
Mr. H. E. Andrewes, of 8, North Grove, Highgate, N. ;
Mr. J. R. Charnley, of Lyndhurst, Fulwood, Preston ; the
Rev. Archibald Downes-Shaw, of Kettlestone Rectory,
Fakenbam, Norfolk ; Mr. G. E. Frisby, of 40, M^indmill
Street, Gravesend; Mr. O. M. Schmidt Gottmann, of 2, Forest
Villas, Whipps Cross Road, Leytonstone, N.E; Mr. Ernest
Purnell Jones, of 7, Nantwich Road, Crewe; and Count Emilio
Turati of 4, Piazza S. Alessandro, Milan, were elected Fellows
of the Society.
Fahre Memorial.
The President handed round for inspection a copy of the
plaquette designed in honour of M. J. H. Fabre, of Serignan,
Vaucluse, an Honorary Fellow of the Society, to whom also
a presentation was made recently by a number of Entomo¬
logists of all nations in appreciation of his services to Science.
Exhibitions
New and Rare Beetles. — Professor Hudson Beare
exhibited a specimen of Pterostichus aterrinms, Pk., taken
by himself under cut sedges near Stalham, Norfolk, on
19th April, 1910.
( lix )
Many years ago this insect was apparently fairly common
in the Fen District in the East of England, but there ai-e no
records of its capture for a great number of years, and in fact
it was supposed to be extinct in this country.
Commander J. J. Walker brought for exhibition the follow¬
ing rare Coleoptera : — (a) a specimen of Latlirohium longipenne,
Fairm., a beetle recently introduced as a British species, taken
at Tubney, Berks., July 29, 1909 ; {b) a specimen of
a remarkable ants’-nest beetle from the Atherton district,
New South Wales, described by Mr. A. M. Lea under
the name Tretothorax cleistostoma, and regarded by him as
representing a new family of Coleoptera, the Tretothoracidae ;
also the ant, Odontomachus coriarius, Mayr., with which
the beetle was found. These specimens were communicated
by Mr. C. French, a Fellow of the Society and Government
Entomologist, Victoria, New South Wales ; (c) two specimens
of Thomosis guanicola, Broun, a beetle allied to Sphaeridium,
etc., taken by Dr. L. Cockayne among penguin guano on the
Bounty Islands, 490 miles south-east of New Zealand.
Mr. G. C. Champion was of opinion that Tretothorax was
a member of the Heteromerous section of the Coleoptera, and
Mr. G. C. Gahan that it belonged to the newly-constituted
Family RhysojKiussidae.
Mr. J. R. LE B. Tomlin brought for exhibition examples of the
following British Coleoptera : («) Maci'onychus ^-tuherculatus,
Miill., recently rediscovered in the River Teme, and previously
only known from the River Dove, and not taken for at least
40 years ; (6) Enicmus histrio, Joy and Tomlin, a new species
described in the November number of the E.M.M. ; (c)
Laccohius regularis, Rey, from small “ sphagnum” pools at New¬
bury ; introduced recently by Dr. Sharp as L. scutellaris,
]\Iots., but now considered by him as mox^e correctly
named regidaris, Rey ; (d) Cionus lo7igicollis, Bris., taken at
Harewood Forest on Vei-hascimi thapstis on June 26, 1909,
the only previous capture in Britain being by Mr. Moncreaff
•at Portsmouth in 1871 ; and (e) Bemhidium tihiale, Duft.,
a melanic example taken this summer by the River Monnow.
Mr. Tomlin also exhibited on behalf of Dr. David Sharp,
F.R.S., examples of (/) Laccohius ytenensis, Sharp,
a new
( lx )
species also described in the current PIM.M. ; and {g) Orepi-
(lodera hnp)ressa, Fab., a littoral species recently introduced
to the British list by Dr. Sharp from Hayling Island.
It has now been found in an exactly similar locality at
Poole Harbour by Colonel Yerbury.
Asilid and Prey. — Mi-. G. T. Bethune Baker showed an
Asilid which be had taken at IMacugnaga in August with a
dead $ NomkuJes semiargus in its mouth. The Asilid held
the Lycaenid until it was boxed. At the same time' another
Asilid was observed with a beetle, which on being captured
was relinquished and lost in the grass.
Profe.ssor E. B. Poulton, F.R.S., said that undoubtedly
the Asilids inject a poison into their prey through the
proboscis which kills them immediately.
Ant Host and Guest. — Mr. H. St. J. Donisthorpe showed
a specimen of Claviger longicornis, Miill., with its
host Lasius timh'aUis, Nyl., taken by Father Schmitz in
Germany. He said this species should occur in Britain
with the same ant, and that Father Schmitz had told him
that April was the best month in which to look for it in
the nests of umhrahis under deeply embedded, heavy stones.
Teratological specimens. — Dr. T. A. Chapman exhibited an
example of Pterostoma palpma, one of a number of similar
specimens bred by Mr. L. W. Newman fi-om one brood of
larvae, and which may be called as a varietal (or aberrational 1)
name, var. hrevipennis. It is characterised by the extreme
shortness and (comparatively) great breadth of the wings, a
forewing measures 15 mm. long and 12 mm. from apex to
tornus; a normal specimen taken at i-andom has these measure¬
ments, 24 mm. long 10 mm. wide. Dr. Chapman expressed
regret that such specimens, involving structural variation, are
despised in favour of mere colour fluctuations. In the pi-esent *
instance there is a remarkable congenital variation tending to
the formation of a race, by way of discontinuity. For com¬
parison a specimen of Libythea celtis with a shortened wing was
shown. It is impossible in the specimens without the histories
to say that they were not of identical origin, yet the one is
congenital (or even racial), the other the result of a displaced
antenna in the pupa.
Proc. Ent. Soc. Land., iQio, Plate A.
Photo, F. N. Clark.
MALFORMATION IN ACRONYCTA TRIDENS. See p. Ixi.
1. The specimen X a. '1. Extremity, X Iti. ■’>. Interior jiortiun, X 10. 4. Xormali;
a]i])endages lateral view to coni})are with tig. 1.
( Ixi
Dr. Chapman also exhibited on behalf of Rev, C. R. N.
Burrows a specimen of malformation of the male appendages
in Acronycta tridens. No similar specimen appears to have
been recorded. Instead of occupying their proper situation,
the clasps, penis, and penis-sheath formed a mass in the
interior of the abdomen, and were accompanied by two,
apparently chitinous, bodies, that could hardly be anything
else than the two tubercles that represent these parts in the
pupa. The specimen suggests in the strongest way that the
parts placed internally are really internal parts, that become
external at pupation, and have in this specimen remained
internal through an arrest in the development of the indi-
Diagr itn to show the parts in noriral relations to each other and to the pupal skin
A pupal skin ; B parts external in specimen ; C parts internal in specimen.
vidual. It may bear on this that the spines on the eversible
membrane {vesica, Pierce) are poorly developed, though the
rest of the parts are fully foi’med, but owing to their situation
poorly displayed. On the other hand, all the authorities
agree that the clasps at least are dermal appendages, and
are developed from dermal structures of the 9th abdominal
segments, though any clear demonstration of this appears to
be lacking ; but in support of this view (the dermal origin of
the clasps) they possess scales and hairs which can hardly be
possessed by processes from the interior. Another hypothesis
would be that the clasps are dermal, but become internal for
a time (not unlike the wings) and here failed to return. The
specimen in fact raises more questions than it settles.
( Ixii )
Bred Ledcania l-album. — Mr. H. M. Edelsten exhibited
a bred example of Leucania l-album of which he gave the
following account : —
“In the ‘Entomologist,’ vol. xlii, p. 322 (1909), Mr. E. P.
Sharp recorded the capture of a $ specimen of Lmcania l-album,
from which he obtained a few ova. On November 4, he noticed
they had begun to turn colour and on the 5th they went
quite black and begun to hatch in the evening. The larvae
ate up the empty egg-shells anti some infertile eggs, but
refused any other food, though he tried almost everything.
They wandered about, and on being supplied with a section
of an old reed stem they promptly entered it, and on finding
their way into the inner lining commenced to hibernate.
“ The larva was about 2^ mm. in length, of a brownish
grey colour, head and plate on prothoracic segment yellowish
browD, and it had a few small bristles from tubercles.
“ Mr. Sharp very kindly sent me five larvae, and on
February 22 I noticed three had come out of the reed
and wex'e moving about (the other two had perished). They
at once changed their skins, and when this was over they
started to nibble some Poa atiuaa which I gave them. Two
more died dui-ing the second moult, but the remaining one
grew up slowly, pupated on June 11, and produced a fine
$ on July 14. Mr. Sharp unfortunately lost all his. This
species is double-brooded on the Continent, emerging in May-
June and August-September. I think we may overlook the
first brood in this country. The larvae seem hardy enough,
mine were kept in an old outhouse facing north.”
Variation of Luperina gueneei. — Mr. R. South showed an
exceedingly interesting and rather variable series of Luperina
gueneei, Doubleday, sent him by Mr. W. Yates, of St. Anne’s-
on-Sea, who obtained them, chiefly this year, on the Lancashire
coast. Commenting on his exhibit he said — “ The first
specimen in the series (No. 1) agrees well with the original
description of L. gueneei, the others show more or less of
the typical ochreous coloration but are variable in marking.
Mr. Yates considered the four specimens. Nos. 3-6, to be
melanic forms of L. gueneei, but I find that they are certainly
dark aberrations of L. testacea, probably referable to ab.
( Ixiii )
nigrescens, Tutt. The colour of the forewings, however, is not
‘ blackish grey ’ neither is it ‘ black,’ but ranges from greyish,
brown (No. 3) through dark brown (Nos. 4-5) to blackish brown
(No. 6) ; the pale edging of the transverse lines is distinct in
all the specimens.
“ Recently there has been a report that melanic specimens of
L. gueneei had been obtained this year on the Lancashire coast.
The darkest specimen of the species that has come under my
notice is No. 15 in the series now exhibited, and even this-
is very little darker than tyjiical gueneei, but there is little
ochreous in its coloration. Probably the rumour arose from
some error in identification, as in the present case.
“ The two species are somewhat similar in a general kind of
way, but among other points of difference that might be
mentioned, two stand out clearly, and study of these should
enable us to easily distinguish one species from the other.
“ In testacea the subterminal line is excurved between veins
5-7, outwardly angled at vein 3, thence incurved to the
dorsum ; the area beyond the line is nearly always darker
even in dark specimens, than the general colour. The same
line in gueneei (usually edged with white on the costal area) is
not angulated at vein 3, and the outward turn between veins
5-7 is not curved, but forms a square bracket; the area beyond
the line is not darker than the general colour, but often much
paler, even whitish.”
Oria musculosa in Britain. — Mr. South also exhibited
three of six specimens of Oria {Synia) musculosa, taken in the
Salisbmy district, in August 1909, by Mr. H. Haynes, who
captured three others in August of the present year. The hind-
wings of the specimens exhibited are darker than are those
of any other British or Continental specimen seen by the
exhibitor, with the exception of one rather small specimen
from Capri, in the British Museum Collections.
The species is undoubtedly scarce in Britain, and most of the
specimens recorded, or about which anything is definitely
known, have been taken in Sussex, chiefly on the coast, and
at long intervals between the years 1855 and 1883. Single
examples have been recorded from Kent and S. Devon, the
latter in 1899. The range of 0. musculosa would seem to
( Ixiv )
extend westward from Syria and Asia Minor to Spain,
Algeria, and the Canaries, and northward to Germany, France,
England and Denmark.
Aberrant Lepidoptera. — Mr. F. C. Oldaker showed a case,
containing various aberrant forms of Lepidoptera, viz. : — [a)
an example of Argynnls aglaia, a very dark form taken at
Chatelard, Valais, Switzerland, on August 16, 1909 ; (/;)
examples of Folygonia C-allmvi bred from ova, September
16-21, 1910, sent by Mr. L. W. Newman from the Wye Valley,
including one specimen, a $, of a very pale form ; (c) a series
of Noctua ditrapezium, including a form in which the ground¬
colour of the fore-wings is almost uniform dark reddish brown,
the usual black markings being only slightly darker than the
rest of the wing, and very faintly discernible ; and (d) a
series of Epione advenaria, bred from ova at Haslemere, 1907 ;
one of a remarkable form very much smaller than usual, and
of a uniform dull brown colour, with white fringes, but no
markings on the wings. The specimen is a $, and is recorded
in the “Entomologist,” Vol. xli, p. 157, being similar to the
male recorded (Joe. cit.) Vol. xliii, p. 201, taken April 8,
1907, by CajDtain Cardew.
Melitaeid Variety. — Mr. A. H. Jones exhibited a series
of Melitaea dictynna, var. vernetensis ^Ohth., taken by him this
year at Vernet-les-Bains, Pyrenees-Orientales, and said that
in his opinion the so-called variety was probably a distinct
species, having regard to its general appearance, and the
surroundings in which it is taken. With it he also showed
examples of M. dictynna type, and M. athalia for comparison.
Papers.
The Rev, F. D. Morice, M. A., read a paper entitled “ Hymen-
optera Aculeata collected in Algeria : The Sphegidae” being
Part V. of the work commenced by the late Edward Saunders,
F.R.S., F.E.S., in the TYans. Ent. Soc., 1904. p. 515.
Professor E. B. Poulton, D.Sc., M.A., F.R.S., communicated
a paper entitled “ Experiments with the larva and jjupa of
Uropteryx sambucaria in connection with their Colour Sur¬
roundings,” by Elizabeth Bridges.
The President, at the close of the discussion which followed,
( Ixv )
proposed a special vote of thanks to Miss Bridges, who was
present, and this was carried unanimously.
Dr. Malcolm Burr, D.Sc., M.A., communicated a paper
entitled “ A Revision of the Labiidae, a Family of Dermaptera.”
Wednesday, November 16th, 1910.
Dr. F. A. Dixey, M.A., M.D., F.R.S., President, in the
Chair.
Nomination of Officers and Council for 1911.
Mr. H. Rowland-Brown, one of the Secretaries, announced
that the Council had nominated the following Fellows to act
as officers for 1911 : President, Mr. J. W. Tutt ; Treasurer,
Mr. A. H. Jones; Secretaries, Commander J. J. Walker,
M.A., R.N., F.L.S., and the Rev. G. Wheeler, M.A., F.Z.S. ;
Librarian, Mr. G. C. Champion, F.Z.S., A.L.S. ; and as other
members of the Council, Mr. R. Adkin, Professor T. Hudson
Beare, B.Sc., F.R.S.E. ; Mr. G. T. Bethune-Baker, F.Z.S. ;
Dr. M. Burr, D.Sc., F.L.S., F.Z.S.; Mr. H. St. J. Donis-
thorpe, F.Z.S. ; Mr. J. H. Durrant, Professor Selwyn Image,
M.A. ; Dr. K. Jordan, Ph.D. ; Mr. A. Sich, Mr. J. R. le B.
Tomlin, M.A., and Mr. H. J. Turner.
Appointment of Auditors.
The following Fellows were appointed to act as auditors of
the Society’s account for the current year : Mr. H. St. J.
Donisthorpe, F.Z.S., Dr. T. A. Chapman, M.D., F.Z.S. ; Mr.
R. Wylie Lloyd, Mr. A. Sich, Mr. H. J. Turner, and Mr.
C. 0. Waterhouse.
Darwin Aledal.
The President having announced that the Darwin Medal
of the Royal Society was to be presented to Mr. Roland
Trimen, M.A., F.R.S., a Past-President of the Society, it was
agreed unanimously to convey to the recipient the heaity
congratulations of the Entomological Society of London.
Election of Fellows.
Dr. Geoffrey Douglas Hale Carpenter, M.A., M.B.,
Uganda Medical Service, Uganda Protectorate ; Mr. William
PROC. ENT. SOC. LOND., V. 1910.
E
( Ixvi )
B. Gurney, Assistant, Government Entomological Department
of Agriculture, Sydney, N.S.W. ; and Mr. J. C. Hawkshaw,
Hollycomb, Liphook, Sussex, were elected Fellows of the
Society.
ExhibUions.
Rare Palaearctic Moth. — The Hon. N. 0. Rothschild
brought for exhibition some examples of a rai'e Noctuid moth,
Oxytripia orhiculosa, Esp., collected by himself and Miss Sarolta
von Wertheimstein, at Puszta Peszer, in Hungary, during the
first week of October of this year, where examples of both
sexes were secured.
The exhibitor made some remarks on the curious habits of
the moth illustrated by photographs of one of the sandy
spots in the wood it frequents, and recalled the adventurous
history of its discovery and re-discovery in Hungary.
The Hon. N. C. Rothschild also exhibited examples of two
species of flea, Cte7iocephalus canis (dog-flea) and Ctenocephalus
fells (cat-flea), and stated that, though still frequently con¬
sidered to be identical, they were really quite distinct species.
The exhibitor also remarked that the two had been united by
Dr. Taschenberg under the name of se7'ratice])S, a, name which
most certainly could not be retained. Under the microscope
it was seen that whereas the head of the dog-flea was rounded,
that of the cat-flea was long and flat.
Rare British Beetles. — Dr. G. Nicholson showed the
example of Lath^'ohiitm longip67ine, hairm., taken by him at
Croydon, in May 1910, of which a specimen was exhibited by
Commander J. J. Walker at the last meeting of the Society.
Professor T. Hudson Beare exhibited specimens of three
species of beetles, all taken abundantly by him at Nethy Bridge,
Inverness-shire, during July and August 1910, viz. : (a) Erir-
rhmus aethiops, F.— This species occurred in great abundance
in a mere handful of flood refuse on the banks of the river
Spey. The only other record of its occurrence in numbers is
that given by the Rev. W. F. Johnson from the North of
Ireland ; (6) C i'iocephalus ritsticus, Dej. — This species was taken
in numbers in the stumps of, and in small standing Scotch fir-trees
in a portion of the pine woods which surround the village, and
( Ixvii )
which had been swept over by a forest fire some few years ago.
There was no doubt that the larvae had a pi’eference for this
burnt timber, as they were much more abundant in this area
than in similar timber in adjacent parts of the woods ; (c)
Zeugophora turneri, Pow. — This species was beaten in great
profusion from aspens growing near Loch an-Eilan. Many of
the specimens were slightly immature, and as the species had
been taken early in June by Mr. Donisthorpe, near Braemar,
there is little doubt that this insect is double brooded.
Experiments with Ants’ Nests. — Mr. W. C. Crawley exhi¬
bited a colony of the ant Lasius niger which had accepted as
queen a $ of Lasius umh'atus in 1908. Up to this autumn
the only ants which had come to maturity in the nest were
pure Lasius nigei', thus confirming Reichenbach’s experiments
(“ Biologische Centralblatt,” July 15, 1902, p. 461), that
Losing niger ^s are able to produce ^s parthenogenetically.
It was hitherto supposed that ^s of ants could only lay eggs
that produced i^s, on the analogy of bees. A similar colony
dating from 1896 gave similar results (‘' Science Gossip,” May
1900).
In connection with Mr. Crawley’s exhibit Mr. H. St. J.
Donisthorpe exhibited ^ winged and wingless $ $ and ^ ^
of Lasiits niger and L. umhratus for comparison. He remarked
that umhratus was a scarce but widely distributed ant of con¬
siderable interest. Mr. Barnes has recorded nine wingless $ $
in a nest of Formica sanguinea at Wellington College, and he
himself had found ^ ^ in some numbers on several occasions
with the same ant at Woking. Wasmann has recorded
umhratus with niger. It is probable that L. umhratus $ is
unable to found her own nests and is a temporary social
parasite on niger. Mr. Crawley’s observations and experiments
go to confirm this.
Mr. Donisthorpe also exhibited ^ winged and wingless $ $ ^
and ^ ^ of Lasius fuliginosus, and pointed out that it was now
proved that the $ $ of this ant often founded their colonies
with imih'atus. He quoted Wasmann, Forel, Emery, Crawley
and his own observations, and stated that Mr. Crawley and
he intended to make experiments with these ants next year.
In connection with the ^ ^ bred in Mr. Crawley’s nest from
E 2
( Ixviii )
parthenogenitic eggs laid by the niger ^ he added that
Reichenbach had bred 300 ^ ^ and several dozen $ ^ from eggs
laid by twelve niger ^ ^ from 1899 to 1902. Mrs. Comstock
had also reared ^ ^ in a $less nest of L. niger, var. americanus,
in 1902 in America.
These facts confute Dzierzon’s hypothesis, which has been
tacitly extended to ants, that parthenogenitic eggs laid by the
honey-bee ^ ^ only produce This has never been properly
demonstrated. Finally, he remarked that it was a pity that
9 ? ants had been treated, and expected to behave, like 9 9
bees, which was not the case, and it had no doubt retarded
their proper study.
Close Resemblance of Butterflies from South America. —
Mr. W. J. Kate exhibited specimens of Eueides pavana
[lleliconidae), Actinote thalia [Acraeidae), and Dismorphia
actinote from S. Brazil. Comment was made as to the very
close resemblance between the first two. The resemblance was
greatest on the underside, but the upperside also showed
considerable converge of colouring. The specimen of E. pavana
exhibited had been caught and papered by Mr. Kaye as an
example of the common Actinote thalia. The specimen of
Dismorphia actinote caught on the Corcovado at Rio de Janeiro,
was shown principally as a mimetic species, for which a sharp
look-out was kept, while the much more convergent Heliconidae
had been passed over, because unsuspected. The Dismorphia,
while only a partial approach to the Actinote on the upperside,
was extremely close on the underside, with the hindwing
brought well over the forewing in an attitude of rest.
Aberrant Lepidoptera. — Mr. L. W. Newman exhibited
examples of Abraxas grossulariata, bred October 1910 as a
second brood, including two ab. varleyata d d) one very finely
rayed with white both on the fore- and hindwings, and one with
the yellow band replaced by a very narrow pale lemon band,
the parents being typical forms, and grandparents varleyata d
X type 9- He also showed an interesting Lycaena supposed
to be a natural hybrid between Agriades thetis (bellargus) d
X Folyommatus icarus, 9; taken wild near Folkestone, on
September 10th last; the underside showing both thetis and
icarus characteristics, with those of icarus most pronounced,
( Ixix )
the upperside a fine thetis colour with white fringes, the shape
of wings being also curious, rather suggesting that of Agriades
coridon.
Specimens of A. coridon, A. thetis, and P. icarus were also
exhibited for comparison.
Mr. G, T. Bethune-Baker, having examined this exhibit,
gave it as his opinion that the butterfly was merely an
aberrant form of Polyonimatus icarus,
Italian Rhopalacera. — Mr. Philip J. Barraud brought
for exhibition a case containing several series of a large form
of Satyrus statilinus from the Aurunci Mts., Southern-Central
Italy; series of Parnassius mnemosyne, xav, friihstoferi, ivom.
Mt. Petrella, Aurunci Mts., 9,000 ft. ; series of Colias edusa
and ab. helice, from Formia ; a very small specimen of Gono-
pteryx cleopatra, measuring 37 mm., from Formia; and four
examples of a large form of Pamphila comma from Southern-
Central Italy.
Papers.
The following papers were read : —
“ On the early stages of Latiorina {Lycaena) cu'hitrdus, Prun.,
an Amyrmecophelous Plebeiid butterfly,” and “ On the larva
of Orgyia splendida, Rbr. {duhia)," by Dr. T. A. Chapman,
M.D., F.Z.S.
“ Notes on Insect Enemies in the Tropics, and their Influence
on Mimicry,” by Edward A. Cockayne, F.E.S.
“ New Genera and Species of Striphnopterygidae and
Lasiocampidae^^ by Professor Christopher Aurivillius, Hon.
F.E.S.
Wednesday, December 7th, 1910.
Mr. H. Rowland-Brown, M.A., Vice-President, in the
Chair.
Nomination of Officers and Council, 1911.
The Secretary again read out the names of the Fellows
nominated to serve as Ofllcers, and other members of the
Council for 1911.
( Ixx )
Election of Fellows.
Mr. 11. Stewart MacDougall, M.A., D.Sc., F.E.S.E., of
Edinburgh University; and Mr. Hugh Frederick Stokeham,
Lieutenant, East Surrey Regiment, of “ Kingswear,” Streat-
ham Park, S.W., were elected Fellows of the Society.
Letter of Congratulation to Mr. Roland Trimen, F.R.S.
“ Entomological Society op London,
“11 Chandos Street, Cavendish Square, W.
“Dear Mr. Trimen,
“ At the last meeting of the Entomological Society
of London, it was proposed from the Chair and carried
unanimously —
“ ‘ That the hearty congratulations of the Society be accorded
to Mr. Roland Trimen, M.A., F.R.S., on the occasion of the
award to him of the Royal Society’s Darwin Medal.’
“ In conveying to you this expression of the Entomological
Society’s appreciation of the honour conferred on a dis¬
tinguished ex-President, we should wish to add that, in our
opinion, this event reflects no less honour upon the donors than
upon the recipient of the Medal. You are known to all
generations of scientific men as a veteran worker among those
who have found their chief inspiration in the principles laid
down by Charles Darwin. Among living naturalists there
are few indeed whose merits as associates and fellow-woi'kers
with Darwin can bear comparison with your own ; and we
feel sure that all alike, in rejoicing at this public recognition
of your life-long services to Biological science, will agree that
the present honour could not have been more worthily
bestowed.
“ (Signed)
“F. A. Dixey, M.A., M.D., F.R.S., President.
“ H. Rowland-Brown, M.A.
“J. J. Walker, M. A., R.N.,F.L.S.
“ Roland Trimen, Esq., M.A., F.R.S., F.L.S.
“ December 1910.”
( Ixxi )
Travel Grant.
The Vice-President announced that Mr. F. Merrifield
had most generously offered to renew his donation to the
Travel Grant of the Society, and proposed a vote of thanks,
which was given unanimously.
Collection of North American Beetles.
The Vice-President announced that he had received from
Dr. A. Feynes, of Pasadena, California — who was recently
elected a Fellow — and exhibited on his behalf, four boxes con¬
taining an admirable collection of North American Aleocharine
Coleoptera, which the donor had kindly offered to the Society.
In the absence of any collections belonging exclusively to
the Entomological Society of London, however, he had asked
Dr. Feynes to authorise a transfer of the gift to the
British Museum (Natural History), and he therefore, with the
consent of the Meeting, handed it over to Mr. G. J. Arrow
for that purpose.
Mr. G. J. Arrow, on behalf of the British Museum, said
that the donation would be greatly appreciated, as it included
examples of many rare species of North American Aleocharinae
not represented in the National Collections.
Exhibitions.
Bare and Variant Diptera. — Mr. H. W. Andrews ex¬
hibited a short series of Carphotricha guttularis, Mg., a scarce
Trypetid, taken at Milford Haven in July last, and a specimen
of a unicolorous form of Prosena sybarita, F., from North
Kent, July 30, 1910.
Introduced and New Insects. — Commander J. J. Walker
exhibited (a) specimens of Syagrms intriulens, Wat., an
Australian weevil, which had been introduced into a fernery
at Glasnevin, co. Dublin, where it had done considerable
damage; communicated by Mr. J. N. Halbert, M.R.I.A. ;
and of (6) Conops signata, Wiedemann, ^ and a Dipteron
new to Britain, taken at Tubney, Berks, September 11,
1910, and exhibited on behalf of the captor, Mr. Joseph
Collins, of the Oxfoi’d University Museum.
Rare and Variant Coleoptera. — Mr. E. C. Bed well
( Ixxii )
brought for exhibition examples of {a) Bruclms pectinicornis,
L. This beetle, as a rule, is looked upon as being introduced
into Britain, and only to be met with in granaries and similar
places ; the specimen exhibited was, however, swept on an
open hillside at Chipstead, Surrey, August 14, 1910: also
(i) Badister bipustidatus, F., a variety of this common species,
the usual black patches on the elytra being reduced to two
small black dots; from Chipstead, May 8, 1910.
Brachypterous Ants. — Mr. W. C. Crawley showed, with
normal examples, a brachypterous $ of the ant Lnsius
found at Oddington, near Oxford, in August 1900, at which
locality about the same time were observed $ ? of L. niyer
with short wings. Professor Wheeler has found similar 9 9
of American ants with the abdomen containing a nematode
worm.
Mr. H. St. J. Donisthorpe remarked that Herr Mrdzek had
recently shown that the short wings in 9 9 of Lasius alienus
were caused by the ant being infested by a nematode worm
of the genus Mermis, and he called them “ Mermithogynes,” and
that Professor Wheeler had found this to be the case with
short- winged 9 9 of L. neoniger in America. He now exhibited
a short-winged $ of Teclinomyrmex alhipes, Smith, together with
an ordinary winged ^ which he had recently taken at Kew, and
suggested that the former might be caused in the same way :
also ergatoid of the same species taken at the same time.
Forel has described this form from specimens captured by
the exhibitor, and suggests that they may be caused by the
presence of parasite guests entertained by the Technoviyrinex
in the same way as Pseudogynes are caused by the presence
of Lomechusa with F. sanguinea.
He further exhibited two forms of Prenolepis hraueri, sub-
sp. donisthorpei, Forel, taken at Kew. A black form ^ ^ and
taken in the Fern House and a red form from the Palm
House.
Bred Rare British Lepidoptera. — Mr. H. M. Edelsten
exhibited series of the following rare British Heterocera, viz. :
(a) Dianthoecia luteago, var. harrettii, bred 1910, from Devon
larvae ; (6) Tapinostola extrema, bred from Northamptonshire
larvae, July 1910 ; (c) Tapinostola hellmanni, bred from larvae
( Ixxiii )
taken in Wicken Fen, June 1910, and {d) a pale variety of
Meliana Jlammea, bi’ed from larvae collected in the Norfolk
Broads, together with a type specimen for comparison.
Butterfly and Mimic. — Mr, H. H, C, Druce brought for
exhibition a remarkable Nymphaline butterfly from the
Himalayas, Parhestina jermyni, n. sp., with Aforia agathon, var.
phryxe, the Pierid it mimics closely — the subject of a paper
read by him.
Melanism in Melitaeids. — Mr. H. Rowland-Brown ex¬
hibited, together with typical examples for comparison, two
fine melanic aberrations of Alelitaea parthenie, Bork., one of
which resembled ab. rhoio, Oberthiir (Lepid, Comparee, fasc.
iv. pi. xliv), taken at 2,800 ft., at Clelles, Isere, France, July
23, 1910; and also two very remarkable black aberrations
of Melitaea varia, taken by him in company with Mr, C. F.
Johnson, July 20, 1907, at the top of the Simplon Pass,
7,000 ft.
Papers.
Mr. Hamilton H. C. Druce, F.L.S., F.Z.S., read a paper
entitled “ Description of a new Nymphaline Butterfly from
British India.”
Mr. H. St. J. Donisthorpe, F.Z.S., read a paper entitled
“ Further Observations on Temporary Social Parasitism, and
Slavery in Ants.”
Dr. T. A. Chapman, M.D., F.Z.S., read a paper on “Two
new species of Lycaenopsis from Sarawak, Borneo.”
M. Ernest Olivier communicated a paper entitled
“ Description of two new species of Luciola in the Collection
of Mr. H. E, Andrewes.”
( Ixxiv )
ANNUAL MEETING.
Dr. F. A. Dixby, M.A., M.D., F.R.S., President, in the
Chair.
Mr. li. Wylie Lloyd, one of the Auditors, read the
Treasurer’s Balance Sheet, showing a balance of £30 11s. lOcZ.
in the Society’s favour. On the motion of Mr. R. S. Standen,
seconded by Mr. F. Enock, the Balance Sheet was adopted
unanimously.
Mr. H. Rowland-Bbown, one of the Secretaries, then read
the following : —
Report of the Council.
During the Session 1910-11 nine Fellows have died, viz.
Mr. C. G. Bignell, Mr. G. W. Kirkaldy, Mr. Oliver C.
Goldthwait, Mr. George Henry, Mr. W. A. Luff, Mr.
Albert Piffard, Mr. George S. Saunders, F.L.S., Mr. Edward
Saunders, F.R.S., F.L.S., and Mr. James William Tutt; ten
Fellows have resigned, and the names of eleven Fellows have
been removed from the list.
The number of Fellows deceased is about the average, the
number of names removed from the list slightly above it ; but
it is highly satisfactory to observe that the number of elections
into the Society is the highest recorded in our annals ; the
long list of candidates for election in the early part of the
current year indicating that we may look for an annual
increase on this scale to become normal in the future — the
wider range of research adopted by our Fellows, and the
more general recognition by the public as a consequence of
Entomology as a practical science, being responsible in our
opinion for this highly satisfactory state of things.
At present the Society consists of twelve Honorary Fellows,
and five hundred and forty-two ordinary Fellows, making a
total of five hundred and fifty-four.
The Transactions for the year 1910 form a volume of five
hundred and thirty-three pages, containing twenty Memoirs
( Ixxv )
by the following authors : George T. Bethune-Baker, F.Z.S.,
Arthur G. Butler, Ph.D,, F.L.S., F.Z.S., Malcolm Burr, D.Sc.,
F.L.S., F.Z.S., Dr. Thomas Algernon Chapman, M.D.,
F.Z.S. (two), H. St, J. Donisthorpe, F.Z.S., T. Bainbrigge
Fletcher, R.N., Dr. Joseph Lane Hancock, Norman H. Joy,
M.R.C.S., L.R.C.P., Arthur M. Lea, Edward Meyrick, B.A.,
F.R.S., F.Z.S. (two), the Rev. Francis David Morice, M.A. (part
author of one paper with the late Edward Saunders, F.R.S.)
(two), Claude Morley, Ernest Olivier, Louis B. Prout, Roland
Trimen, M.A., F.R.S., F.L.S., and Rowland E. Turner (two).
Of these, eight papers relate to Lepidoptera, three to
Coleoptera, six to Hymenoptera, two to Orthoptera, and one
is of general entomological interest.
The Memoirs above referred to are illustrated by sixty
plates — a record number — of which eight are coloured.
Towards the cost of Plates I-XIII Mr. G. Bethune-Baker
has contributed £5 ; towards the cost of Plates XIV, XV
(chromo) Dr. T. A. Chapman has contributed half the ex¬
penses, and also has given the cost of the blocks for Plates
XVI-XLIII, and LI-LX, together with that of Plate A
in the Proceedings ; for the Plates XLIV and XLVII Dr.
Malcolm Burr supplied the drawings, and Mr. L. B. Prout
the drawings for Plate XLVIII (chromo) ; towards the cost
of Plate L (chromo) Mr. Rowland E, Turner has contributed
the sum of <£10.
The volume of Proceedings consists of seventy-three pages
in all, containing many notices as well as several short papers
of scientific interest, one of which is illustrated by a half-tone
block, the gift of Dr. T. A. Chapman.
Mr. F, Merrifield, one of our past Presidents, again having
volunteered a handsome donation to the Travel Fund, we
were in a position last summer to send one of our Fellows,
Mr. B. C. S. Warren, abroad for a prolonged entomological
tour in South-eastern France. We feel sure that the con¬
ditions of this grant have only to be brought home to
Fellows to be more generally appreciated, and utilised by
those who are anxious to pursue their entomological studies
outside the United Kingdom.
At the first Congress of Entomology, held at Brussels in
( Ixxvi )
August last, we were represented officially by Dr, F. A.
Dixey, M.A., M.D., F.E.S., Mr. Roland Trimen, M.A., F.R.S.,
Mr. H, St. J. Donisthorpe, F.Z.S., and Mr. F. Merrifield ; the
great success of the meeting being in no small degree due to
the energy and interest displayed by the Fellows of our
Society, of whom the Hon. Walter Rothschild, M.A., D.Sc.,
and Dr. Karl Jox’dan, Ph.D. (Chairman of the Executive
Committee), Pi’ofessor Poulton, F.R.S., Dr. F. A. Dixey,
F.R.S., Mr. H. Rowland-Brown, IM.A., Mr. G. A. K. Marshall,
and Dr, Malcolm Burr, D.Sc. (General Secretary of the
Executive Committee), were elected permanent members of
the International Council.
We are interested to know that a second Congress of
Entomology is to be held next year at Oxford, under the
Presidency of Professor E. B. Poulton, D.Sc., F.R.S,, Hope
Professor of Zoology in the University of Oxford, and past
President of our Society.
At the International Congress of Zoology, held at Gratz in
August last, our delegates were the lion. Walter Rothschild,
M.A., D.Sc., and Dr. Karl Jordan, Ph.D.
The Treasurer reports that after carrying forward to 1911
£23 2s. for subscriptions paid in advance in 1910, and in¬
vesting £47 5s, in Consols for three Life Compositions received
during that period, making the total sum so invested
£1,045 6s. 3(7., there remains a genuine cash balance in the
Society’s favour of £30 11s. lOcZ.
An analysis of these figures shows that the new system of
combining the printing and distribution accounts ensures a
considerable saving of money, and that we are now well able
to discharge all liabilities incurred for the cost of producing
the Transactions and Proceedings within the current session.
It must be borne in mind also that the general expenditure
of the Society for the year includes also the cost of production,
and of the design of the new seal of the Society, which will
henceforth be affixed to all official documents issued by the
Council,
This improved condition of our finances is due in great
measure to the services rendered by the late Mr. J. W. Tutt,
by whose untimely death the Council and the Society suffer
( Ixxvii )
an irreparable loss in the management and control of their
business affairs.
In conclusion the Librarian announces that thii’ty volumes,
and a large number of Separata, together with the usual
Periodicals, have been added to the Library, and that greatly
increased use has been made of it for purposes of study and
reference by Fellows ; also the issue of books for home work
has been considerably extended.
H. Kowland-Brown,
J. J. Walker, R.N.,
Secretaries.
Entomological Society of London,
11, Chandos Street, Cavendish Square, W.
1 A January, 1911.
On the motion of Mr. R. Adkin, seconded by Mr. J. H.
Durrant, the Report was adopted unanimously.
The Secretaries also presented the following Report of
the Brussels Intexmational Congress of Entomology : —
THE INTERNATIONAL CONGRESS OF ENTOMOLOGY.
After preliminary meetings in London and Brussels
extending over close upon two years, the first International
Congress of Entomology assembled at Brussels, and continued
in session from August 1 to August 6. Our Society, of whom
the Executive Officers, the Hon. Walter Rothschild, Dr. Karl
Jordan, Dr. G. B. Longstaff, Professor Poulton, F.R.S., Mr.
G. A. K. Marshall and Dr. M. Burr, had already served upon
the General Committee, was specially represented by Dr. F. A.
Dixey, F.R.S., Mr. Roland Trimen, F.R.S., Mr. F. Merrifield
and Mr. H. St. J. Donisthorpe, while these and other Fellows
had been chosen by the following Public Bodies, Universities
and Societies to represent them respectively.
Colonial Office : The Hon. N. C. Rothschild, M.A., F.L S.,
F.Z S. (also delegate for the Tropical African Entomological
Research Committee).
Board of Agriculture: Dr. R. Stewart MacDougall, M.A.,
D.Sc.
( Ixxviii )
Commonwealth of Australia : Professor F. V. Theobald,
M.A.
Commonwealth of South Africa : Mr. Roland Trimen,
M.A., FR.S., F.L.S.
British Museum (Natural History) : The Hon. Walter
Rothschild, D.Sc., F.Z.S., F.L.S.
Royal Society and Linnean Society : Professor Poulton,
D.Sc., F.R.S.
Zoological Society : Mr. G. A. K. Marshall, F.Z.S,
University of Cambridge ; Professor R. C. Punnett, M.A.
University of Oxford: Dr. F, A. Dixey, M.A., M.D.,
F. R.S. ; Professor Poulton, D.Sc., M.A., F.R.S. ; Dr. Malcolm
Burr, D.Sc.
University of Edinburgh : Dr. R. Stewart MacDougall,
M.A., D.Sc.
University of London : Professor F. V. Theobald, M.A.
Royal British Arboricultural Society : Mr. A. T. Gillanders.
Agricultural Research Institute of Pusa, Bengal : Mr. F.
]\1. Hewlett, M.A., while numerous other Local Bodies, Insti¬
tutes and Associations were also represented by Fellows and
others : the total membership of the Congress amounting to
292, of whom 67 were British subjects.
Proceedings began with an informal reception on Sunday,
July 31, at the Taverne Roy ale, given by the Entomological
Society of Belgium, at which Professor Lameere, the President
of the Congress, received the guests, assisted by the Baron
G. de Crombrugghe de Piquendaele, ably supported by Madame
Lameere, M. Severin, to whose energy and courtesy the success
of the meeting was so largely due, Madame, and Mdlle. Severin,
and a number of other Belgian entomologists and their friends,
each member of the Congress receiving a prettily designed
badge in white metal, and a free pass into the Exhibition
for the week.
But it was not until the following day that the session was
opened in the Salles des Fetes, which had been put at the
disposal of the Congress by the Committee of the Brussels
Exhibition. Here Professor Lameere, Rector of the Fx'ee
University of Brussels, delivered an address of welcome, and
as soon as the subsequent formalities had been complied with,
( Ixxix )
Professor Y. Sjostedt, of the Stockholm 'Natural History
Museum, gave a vivid account, with lantern illustrations, of
the entomological discovei’ies, and the work done by the
Swedish Expedition to Kilimandjai’o, of which he had been
leader ; while other papers were read in the adjoining rooms
assigned to the several Sections detailed for the day.
During the Session the following papers were read or
communicated by Fellows of our Society : —
Section op Economic and Pathological Entomology. —
< ‘ The Artificial Distribution of Insect Pests ” (Professor F. V.
Theobald) ; “ Galerucella lineola, its Life History and Habits,
with notes on Preventive and Remedial Researches ” (Dr. R.
Stewart MacDougall); “The Distribution of the Yellow Fever
Mosquito, Stegomyia fasciata ” (Professor F. V. Theobald) ;
“ Economical Questions in Bengal ” (J. M. Hewlett).
Section of Bionomics, Physiology and Psychology. —
“ Note on the Chemical Constitution of the red-coloured
Secretion of Timarcha tenebricosa ” (communicated by Profes.sor
E. W. Carlier and C. L. Evans) ; “ Les Accouplements
Anormaux chez les Insectes ” (E. Olivier).
Section of Evolution and Mimicry. — “ Mimicry” (Dr. F. A.
Dixey, F.R.S.) ; “ Mendelism and Lepidoptera” (Professor
R. C. Punnett) ; “ Experimental Entomology ” (F. Merri-
field) ; “ A quoi sert le Mimetisme 1 ” (W. Schaus) ; “ The
Systematics of certain Lepidoptera which resemble each other,
and their bearing on general questions of Evolution ” (Dr. K.
Jordan) ; “ Mr. C. A. Wiggins’s Researches on Mimicry in the
Forest Butterflies of Uganda, 1909, 1910” (Professor E. B.
Poulton, F.R.S.).
Section op Nomenclature and of Bibliography, and
Papers of General Entomological Interest. — “The Im¬
portance of the Uniform Use of Technical Terms referring
to Types, Co-type, etc., and on the Eight of Authors to change
the Names given by other Authors ” (H. H. Lyman) ;
together with the following papers on subjects of general
entomological interest, the majority of which, with the above,
will appear, as we are informed, in the Transactions of the
Congress presently to be published. “ La G^ographie et la
Phylogenie des Fourmis” (A. Forel, Hon. F.E.S.) ; “Ants
( Ixxx )
and their Guests,” with lantern demonstration (H. St. J.
Donisthorpe) ; “ On the Conservation of Types in Museums ”
(the Rev. W. J. Holland, Hon. F.E.S.); “The Preservation
of Collections in Tropical Climates ” (J. M. Hewlett) ; “ Dis¬
tribution Geographique, et Physiol ogie des Coleopteres Lam-
pyrides ” (E. Olivier). While among the papers read by others
of our countrymen who are not Fellows may be mentioned
“ The Disinfection of Imported Seeds of Plants, and the Use
of Insecticides,” by Sir Daniel Morris, K.C.M.G., D.Sc.,
D.C.L., F.R.S., representing the Royal Colonial Institute,
and the Imperial Department of Agriculture in the West
Indies; and “Notes on the (Estrides (Diptera),” by Dr. G. H.
Carpenter, representing the Royal College of Science, Dublin.
At the final general meeting the President, Professor
Lameere, having briefly summed up the work done, and con¬
gratulated the Congress upon the unqualified success of its
first session, it was decided that a second Congress should
be held in the summer of 1912. As delegate of Oxford Uni¬
versity, Professor Poulton, supported by his co-delegates. Dr.
F. A. Dixey and Dr. Malcolm Burr, with the other Oxford
Graduates present, cordially invited the members to the
University city, an invitation which was accepted with
enthusiasm.
Meanwhile the more serious business had been relieved by
excursions to various places of interest and beauty in the
neighbourhood, including the Congo Museum of Tervueren
and the Brussels National History Museum. The visit to the
battlefield of Waterloo, under the able direction of M. Charles
Kerremans, who has an unrivalled knowledge of the site
and details of the battle, was very popular. On Friday,
August 5, the success of the Congress was also celebrated
by a banquet, where there gathered together many leaders of
the Science, wearing numerous brilliant orders conferred by
Sovereigns and other heads of States. The celebrations ter¬
minated with a reception, given by M. Max, Burgomaster of
Brussels, in the Hotel de Ville. And there, amid a scene of
brilliance and splendour, entomologists of all nations, old
friends as well as new, speaking many languages, bade one
another farewell. A detailed Report of the Congress by Dr.
( Ixxxi )
M. Burr was published in “The Times,” August 10, 1910,
and of the work of the British and Irish members in the
“ Entomologist,” Vol. xliii, p. 233, by Mr. H. Eowland- Brown,
Of the British contingent attending (67 in all) the following
are Fellows of the Society : — G. J. Ari’ow, R. S. Bagnall, Dr.
M. Burr, G. C. Champion, Dr. F. A. Dixey, F.R.S.; H. St. J.
Donisthorpe, H. Eltringham, G. C. Gahan, A. J. Gillanders,
Sir A. Buchan-Hepburn, Bart.; |^Dr. W. J. Holland, J. M.
Howlett, A. H. Jones, Dr. K. Jordan, E. G. Joseph, Dr. G. B.
Longstaff, H. H. Lyman, Dr. R. S. MacDougall, G. A. K.
Marshall, G, Meade Waldo, F, Merrifield, Professor E. B.
Boulton, F.R.S., Professor R. C. Punnett, the Hon. H. 0.
Rothschild, the Hon. Walter Rothschild, H. Rowland-Brown,
W. Schaus, Professor F. V. Theobald, R. Trimen, F.R.S. ; C. J.
Wainwright ; and ladies, Mrs. Malcolm Burr, Mrs. Donis¬
thorpe, Mrs. Longstaff, Miss Merrifield, Mrs. and Miss Poulton,
Miss Rowland-Brown, and Mrs. Trimen.
The Secretaries not having received any notice proposing
to substitute other names for those contained in the list
prepared by the Council, the following Fellows, on the motion
of Mr. H. Rowland-Brown, seconded by Mr. C. 0. Water-
house, were declared elected to serve on the Council for
1911-12: — Robert Adkin, Geoi’ge T. Bethune-Baker, F.Z.S.,
Professor T. Hudson Beare, B.Sc., F.R.S.E., Dr. Malcolm
Burr, D.Sc., F.L.S., F.Z.S., George Charles Champion, F.Z.S.,
A.L.S., Dr. Frederick Augustus Dixey, M.A., M.D., F.R.S. ,
Horace St. John Donisthorpe, F.Z.S., John Hartley Durrant,
Professor Selwyn Image, M.A., Albert Hugh Jones, Dr. Karl
Jordan, Ph.D., Alfred Sich, Mr. J. R. le B. Tomlin, M.A.,
Henry Jerome Turner, Commander J. J. Walker, M.A., R.N.,
F.L.S., and the Rev. George Wheeler, M.A., F.Z.S.
The following were also elected officers : — Treasurer, Albert
Hugh Jones; Secretaries, Commander James J. Walker,
M.A., R.N., F.L.S., and the Rev. George Wheeler, M.A.,
F.Z.S. ; Librarian, George Charles Champion, F.Z.S., A.L.S.,
but owing to the lamented death of the President-nominate,
Mr. J ames William Tutt, it was announced that the election of
a President would be postponed to a Special General Meeting
PROG. ENT. SOC. LOND., V. 1910. F
( Ixxxii )
to be held upon some convenient date after the first Ordinary-
Meeting of the Society in March.
Dr. F. A. Dixby, the President, then delivered an Address,
at the close of which a vote authorising the printing of the
Address, coupled with thanks to the President for his
Address, and for his services as President during the past
Session, Avas proposed by Mr. C. 0. Waterhouse, seconded
by the Rev. F. D. Morice, and carried unanimously.
Dr. G. B. Longstaff then pi’oposed a vote of thanks to
the other officers of the Society. This was seconded by Mr.
A. E. Tonge, and carried unanimously. The President, Mr.
A. H. Jones, Mr. H. Rowland-Brown, Commander J. J.
Walker, and Mr. G. C. Champion replied.
At the request of some of the Fellows present, Mr.
Rowland-Brown explained the procedure under the Bye-
Laws for the forthcoming election of a President to succeed
Dr. F. A. Dixey.
( Ixxxiii )
ENTOMOLOGICAL SOCIETY OF LONDON.
Balance Sheet for the Year 1910.
Assets.
£ s. d.
Subscriptions in arrear
considered good . 79 0 0
Cost of £1,045 6s. 3d.
Consols. Present value
at the price of 791 on
31st December, 1910,
£829 14s. 4d . 996 18 0
Cost of £239 12s. 4d.
Birmingham 3 per cents.
Present value at the
price of 86 on 31st
December, 1910, £206
Os. 9d . 250 0 0
Balance in hand . 30 11 10
Additional Assets : —
Contents of Library, and
unsold Stock.
£1,356 9 10
Liabilities.
Cost of printing, etc.. Parts 4 and 5.
Audited, compared with vouchers and
found correct —
T. A. Chapjian.
Horace St. J. Donisthorpe.
Hy. J. Turner.
Chas. O. Waterhouse.
R. W. Lloyd.
Less depreciation of £211 2s. lid. in the value of the Stocks.
A. Hugh Jones, Treasurer.
4th January, 1911.
( Ixxxv )
THE PKESIDENT’S ADDEESS.
Ladies and Gentlemen,
Our Anniversary Meeting last year took place in the
midst of a General Election. We may be permitted to hope
that these political tournaments are not about to become as
regularly annual as our own celebration, but the fact remains
that this year we have only escaped a similar coincidence by
the space of a few weeks. Our feelings with regard to the
recent contest are probably mixed ; but I am on safe ground
in asserting that we one and all regret that the House of
Commons has lost, even if only for a time, the services of a
distinguished representative of Entomology.
The year just finished has been an eventful one. It has
been marked by the passing of a beloved Sovereign, who,
during his all too short reign, more than fulfilled the hopeful
anticipations with which his accession was greeted. In his
Successor we welcome a Euler who has already given evidence
of the solicitude with which he watches over the best interests
of his people, and who inspires us with confidence that in him
the cause of science and learning will find an earnest advocate
and promoter. In this he will but be following the tradition
set by an illustrious Father and Grandfather, whose example
is destined to bear fruit for many years to come.
It will be within the recollection of all the Fellows of this
Society that loyal Addresses were presented on their behalf to
King George on his Accession, and to the widowed Queen
Alexandra. The text of these Addresses, together with that
of the Royal Reply, is to be found in our Proceedings.
At a time of general mourning it was obviously fitting that
we should defer the holding of our proposed Conversazione.
( Ixxxvi )
It was eventually decided to forego this entertainment
altogether for the year just ended ; and we may hope
that the social and scientific gathering, when it does take
place, may be for entomologists not the least interesting and
brilliant of the functions marking the season of the coming
Coronation.
If the public bereavement has been severe, our private
losses have also been heavy. All who had the privilege of
knowing Edward Saunders will preserve an abiding memory
of his striking, semi-ascetic countenance, his unfailing courtesy,
his quiet, unassuming but forceful character, his deep learning,
and the mitis safientia that pervaded in him both word and
action. One who has a much better right than myself to
speak of our departed friend has written an eloquent and
touching tribute to his memory ; and those of our number
who would wish to gain an idea of Avhat manner of man
Edward Saunders was, should turn to the account of his life
by the Kev. F. D. Morice. As a Fellow of the Royal Society
he worthily upheld the dignity of Entomology among the
Sciences of Life ; and his loss has been keenly felt, not only
by his immediate associates, but throughout all entomological
circles at home and abroad.
His death was shortly followed by that of his brother,
George Sharp Saunders, F.L.S., also a keen entomologist, and
one who had made valuable contributions to our knowledge of
noxious insects.
It was with unusually great regret that we received the
news of the untimely death of George Willis Kirkaldy,
which took place at San Francisco, in the thirty-seventh year
of his age. It was some five or six years ago that he met with
the accident, while riding, from the ultimate effects of which
he died. Educated at the City of London School, he early
developed a taste for entomology. His first paper, “A
Revision of the Notonectidae,” was published in 1897. The
hemipterous portion of the Zoological material collected by Dr.
R. C. L. Perkins was worked out by him, and the results
published in 1903 as part of the “ Fauna Hawaiiensis.” In
the same year Mr. Kirkaldy was appointed Assistant-
Entomologist to the Hawaiian Territorial Board of Agricul-
( Ixxxvii )
ture and the Hawaiian Sugar Planters’ Association, this office
giving him the opportunity of continuing his studies on the
Hemiptera, and especially on the Fulgoridae. His “ Catalogue
of the Hemiptera,” which was to have included the whole
Order, has been published in part, and it is understood that
further instalments will be issued under the auspices of
M. J. R. de la Torre Bueno, to whose charge Kirkaldy’s
unpublished manuscripts and notes have been committed.
Mr. F. W. Terry, to whom I am indebted for many of the
details just given, has written as follows of his deceased
associate : “ A voluminous writer and wide reader, a staunch
friend and genial companion, he was always ready to give
others the benefit of his udde bibliographical knowledge.
His optimistic and kindly personality will be greatly missed
by his friends and colleagues.”
George Carter Bignell, known for his earlier work on
Lepidoptera and his later researches on various forms of
parasites, especially those infesting the Aphides, died early
in the year at the ripe age of eighty-four.
Other losses have been suffered by our Society in the deaths
of Albert Piffard, a well-known traveller, whose entomo¬
logical interests were chiefly engaged in the collection and
study of Coleoptera and Diptera ; and in those of Oliver
C. Goldthwait, George Henry, and W. A. Luff.
But unfoi’tunately the death-roll of 1910 does not exhaust
the list of our losses. Within the last few days we have
received the sad intelligence that James William Tutt, a
member of our Council and President-nominate, is no moi’e.
This is not the time or the place to attempt a full apprecia¬
tion of his personal qualities, or of the services which he has
rendered to entomology in general and to our own Society in
particular. But I may at least be permitted to recall his
untiring industry, his exemplai-y thoroughness, his contagious
enthusiasm, and that over-mastering love of his subject which
constrained him to devote the scanty leisure of a busy pro¬
fessional life to the single-minded pursuit of his entomological
studies. To what good purpose he used his opportunities we
all know ; and those of us who have had occasion to serve
with him on the Council and Publications Committee will be
( Ixxxviii )
especially sensible of what his loss means to the Society in its
business relations. He has deserved well of us, and his place
will not easily be filled.
Outside our own ranks the year has witnessed the decease
of the well-known collector H. McArthur, who by his labours
in Central Asia, Kashmir and India, as well as nearer home
in the Orkneys, Shetlands and Hebrides, has widened the
bounds of entomological knowledge.
The death of Gustav Beeddin, which took place during the
last days of 1909, should not be allowed to pass without
notice. A recognised authority on the Hemiptera, he was
distinguished by the specially scientific character of his work
upon the group.
Lastly must be mentioned the death of Gustav Fischer,
the well-known publisher of Jena. Not specially an entomo¬
logist, he has yet deserved well of the entomological fraternity
on account of the numerous works, of high interest to all
students of insects, which under his auspices have seen the
light.
In the course of the year several of our Fellows have been
the recipients of well-merited honours. It is a matter of
great satisfaction to us all that the deserts of my immediate
predecessor in this Chair have met with recognition in the
highest quarters. As a Companion of the Imperial Service
Order, Mr. C. 0. Waterhouse will carry into his retirement
from the public service the affectionate goodwill of all
who know him ; and we may hope in this Society to
enjoy the benefit of his long experience and matured
wisdom for many years to come. The University of
Oxford conferred distinction at once on itself and on our
ex-President Professor Meldola, F.B.S., by bestowing on him
the honorary degree of Doctor in Science, and by appointing
him Herbert Spencer Lecturer for 1910. Mr. Selwyn Image,
a member of our Council, has been elected to the Slade
Professorship of Fine Art in the same Univei’sity. Those
who, like myself, have sat at the feet of John Buskin,
will agree that no more worthy occupant could have been
found for the Chair that will ever be linked in association
with the memory of that great teacher and prophet. With
( Ixxxix )
Ruskin, nature and art, though not identical, were in¬
separable ; and in his successor to-day we find a happy illus¬
tration of that successor’s favourite maxim, “ Ars est homo
additus naturae.” We may congratulate ourselves that the
artistic skill of our Slade Professor, already exemplified in
the Addresses which his handicraft has helped to make
worthy at once of the Society and of their recipients, will
leave with us a permanent memorial in the shape of
our new seal, in truth a Krrjfjia es dei for us and our
successors.
This leads me to speak more particularly of the occasion of
one of the Addresses to which I have just alluded. At the
Anniversary Meeting of the Royal Society, the Darwin Medal
was presented to our valued Fellow and ex-President, Mr.
Roland Trimen, M.A., F.R.S. The event was felt to be a
subject of special congratulation to entomological interests in
general as well as to the recipient in particular. It will be
remembered that the Society passed unanimously a special
resolution asking the officers to convey to Mr. Trimen “ the
hearty congratulations of the Society on the occasion of the
award to him of the Royal Society’s Darwin Medal.” I think
it will be of interest to put on record the terms of the
Address, or rather Letter, in which this was done, and also
Mr. Ti’imen’s reply. The Address, after quoting the Resolu¬
tion, proceeds as follows : —
“ In conveying to you this expression of the Entomological
Society’s appreciation of the honour conferred on a distin¬
guished ex-President, we should wish to add that, in our
opinion, this event reffects no less honour upon the donors
than upon the recipient of the Medal. You are known to all
generations of scientific men as a veteran worker among those
who have found their chief inspiration in the principles laid
down by Charles Darwin. Among living naturalists there
are few indeed whose merits as associates and fellow-workers
with Darwin can bear comparison with your own ; and we
feel sure that all alike, in rejoicing at this public recognition
of your life-long services to Biological science, will agree that
the present honour could not have been more worthily
bestowed.”
( xc )
The letter was signed, on behalf of the Society, by the
President and Secretaries. Mr. Trimen’s reply was as
follows ; —
“ Southbury, Guildford,
‘'December 1910.
“To the Entomological Society op London.
“ It is difficult to give adequate expression to my grateful
sense of the honour done me by the Resolution of the Ento¬
mological Society of London, according its hearty congratu¬
lations on the recent award to me of the Darw'in Medal of
the Royal Society, and by the highly appreciative and cordial
terms of the letter of the President and Secretaries trans¬
mitting that Resolution.
“ Such commendation from a Learned Society with which I
have been associated for over fifty years is no slight one, and
greatly enhances the distinction of the Medal in question. I
cannot say how much I owe to the kindness and encouragement
I have always met with from the Society collectively and from
so many individual members of it, and I recognise that the
Medal is as much due to the countenance and aid thus
generously given me as to any efforts of my own.
“ I find pleasure in thinking that Mr. Darwin himself would
have approved of the award this year to an entomologist whom
he knew, and who had the happiness of rendering him some
slight aid in his researches. For was not that greatest of
naturalists’ first love beetles 1 — and has he not himself recorded
that his favourite toast at Cambridge was ‘ Floreat Entomo-
logia ! ’ 1 All of us heartily re-echo this aspiration, and it
might well be adopted as the motto of our Society,
“ Your faithful and attached Colleague,
“Roland Trimen.”
Another great naturalist, w’hom we may claim as an
Honorary Fellow of our Society, is the veteran French ento¬
mologist, J, H. Fabre, of Serignan, Vaucluse, whose eightieth
birthday was signalised by a festival held in his honour at
Orange, on the 3rd of April. The committee formed to
promote this object contained some of the names most
( xci )
illustrious in the Science and Literature of France ; nor were
our own countrymen forgotten. A presentation was made to
the hero of the day, including a fine plaquette, a duplicate
copy of which I had the pleasure of exhibiting lately
in this room. Several Fellows of our Society contributed
towards the fund raised for the occasion, and I cannot resist
quoting a few words from the letter of acknowledgment
written by Dr. Legros, the Secretary of the movement : —
“ Je vous prie, Monsieur le President, de recevoir pour vous-
meme et de vouloir bien transmettre aux genereux donateurs
I’expression de la profonde gratitude des organisateurs du
comite. La noble Angleterre vient de nous donner une preuve
de plus de ses traditions de courtoisie et de solidarite.”
This, I think, may be taken as something more than mere
conventional politeness. I prefer to see in it evidence of what
I am convinced is a fact, viz. that nothing is more apt to
promote feelings of international goodwill than community of
scientific interests.
If we wished for another indication of the truth of this
opinion, we should find it in the excellent and interesting
account of the Jubilee of the Entomological Society of Pussia,
furnished to our Proceedings by Dr. Malcolm Burr, who so
worthily represented us on that occasion.
Our representatives at the Zoological Congress at Graz were
the Hon. \V. Rothschild and Dr. Karl Jordan. The entomo¬
logists there present were few, but made up in quality what
they lacked in number. Many of our Society will be pleased
to know that the long-continued experimental work of Pro¬
fessor Max Standfuss was recognised at the Congress by the
bestowal on him of a special honour.
But if entomologists at Graz were few and far between,
they had mustered in good force at Brussels for the First
International Congress of Entomology. The success of this
undertaking was undoubted. The social and scientific aspects
of the Congress were both felt to be of high value ; and the
fact that Brussels was en fete over the Exhibition with a gaiety
not yet sobered by the deplorable conflagration, while it
may have led here and there to a little crowding and incon¬
venience, yet provided ample compensation, especially for the
( xcii )
youngei’ and more sportive spirits amongst us. The Entomo¬
logical Society has every reason to be satisfied with the part
played by its Fellows at the Congress. Not only Messrs.
Trimen, Merrifield and Donisthorpe, who, with your Presi¬
dent, were specially deputed as representatives, but Professor
Poulton, Professor Punnett, Hon. C. and Hon. W. Roths¬
child, Dr. Karl Jordan, Dr. Burr, and others too numerous to
mention, were prominent in the proceedings. Brussels has
given a good lead, which we may feel confident will be well
followed up in this country next year under the auspices of
Professor Poulton. When the International Congress has
become an established and venerable institution, I trust it
will not be forgotten that the impulse which set it going
oi’iginated from Tring.
While on the subject of the Congress I recall with pleasure
the fact that a most agreeable feature was supplied by the
appearance, during its progress, of Mr. Eltringham’s splendid
work on Mimicry in African Butterflies, a book which I
cordially commend to the notice of all those who are in any
way interested in the subject of mimicry.
Before turning to the more special portion of my Address,
it now only remains for me to expi-ess my very grateful thanks
to the Society in general, and to the Business and Publications
Committee, the Council, and the Officers, in particular, for the
pleasant term of office which I have enjoyed. The experience
of another year in the Chair which my predecessor found
materially, though I trust not morally, uncomfortable, has
only served to deepen my former impression of the magnitude
of the obligation which the Society owes to its Librarian, its
Treasurer and its Secretaries. To repeat what I have said on
a former occasion is unnecessary, but I cannot allow this
opportunity to pass without adding a word of personal appre¬
ciation of the services of our retiring Honorary Secretary, Mr.
H. Rowland-Bi’own. I count myself fortunate to have had
the benefit of his assistance and advice, not only during my
raw novitiate, but up to the end of my tenure of office. As
the last of a long succession of Presidents whom he has helped
to bring up in the way they should go, I congratulate him on
his well-earned retirement, from which I venture to hope
( xciii )
he will one day emerge into an oflBcial position which he has
not yet held, but which he, if any one, is fully competent to
fill.
Entomology and Evolution.
In my former Presidential Address I took as a subject
a matter of special morphological interest. On the present
occasion I propose to ask your attention to a topic of
somewhat wider scope. It is not often that the opportunity
arises of dealing in a general way with the methods and
objects of entomological science. Such treatment is rightly
considered to lie outside the proper range of our Transactions,
nor would it be more in place in the record of our less formal
discussions, or in the midst of those discussions themselves.
But a Presidential Address affords a legitimate occasion for
such generalities. It is possible there, without impertinence
or irrelevance, to give expression to personal views of a larger
application than to any one department of our subject, or to
any particular example of the advancement of entomological
knowledge. I wish, then, to devote a little time to a brief
consideration of some of the matters with which we entomo¬
logists are brought into contact, in their bearing on biological
science in general ; and incidentally to notice the relation
which obtains, or should obtain, between different sides of
entomological study.
With the recent Darwin celebrations still so fresh in our
recollection, it is scarcely necessary to remind ourselves that
most biological pursuits derive their main interest and im¬
portance from the light they are able to throw on the process
of organic evolution. This is perhaps especially the case with
entomology. The intrinsic interest of the study is great, and
has been so felt by many lovers of nature in days before
Darwin. But it needed the Darwinian touch to transfoi’m
the dry bones of description and classification into a living
and breathing organism. Under the stimulating influence of
the theory of selection, all the scattered items of entomological
knowledge were found to have their proper place and signi¬
ficance in relation to the whole structure, and every addition
to our store of entomological fact was welcomed as marking a
( xciv )
fresh stage of growth in our comprehension of at least one
portion of the evolutionary domain. And the peculiar value
of entomological data for research on Darwinian lines was
speedily recognised. The material with which we entomo¬
logists have to deal is for the most part abundant, of manage¬
able size, easy of manipulation, and favourably constituted for
experiment. Hence it has followed that many of the most
important steps in the progress of our evolutionary knowledge
have depended directly or indirectly on the study of insects.
Let me take as an example the present state of scientific
opinion with regard to the transmission of acquired char¬
acters, or, as some prefer to put it, the inheritance of somatic
modifications. It is hardly necessary for me to say that
neo-Lamarckians are still on the look-out for an instance of
such transmission, and that neither among insects nor any¬
where else have they been able to find one. The antecedent
improbability of the alleged phenomenon has been well
shown by many writers, and by none more convincingly than
by Professor Poulton, in one of the admirable Presidential
Addresses delivered by him from this Chair. But the failure
of the neo-Lamarckians to establish the principle of specific
transmission must not blind us to the fact that the germ-
plasm is far from possessing the stability and inviolability
with which it was once supposed to be endowed. Weismann
himself, with great candour, admitted at a comparatively
early stage in his work that the germ-plasm was not in all
cases out of the reach of external influences. It is of special
interest to us as entomologists that he was led to this
conclusion by experiments on a butterfly — the well-known
“Copper” {Chrysophanus phlaeas). Exposing pupae of the
northern golden-red form of this insect to an abnormally
high temperature, he found that many of the resulting
butterflies were slightly dusted with black. On the other-
hand, pupae bred from eggs sent from Naples, which would
under ordinary circumstances have produced butterflies of the
dark southern summer form, were subjected to a relatively
low temperature and gave rise to perfect insects in which the
normal dark coloration was sensibly diminished. These
results, as we shall doubtless all remember, agree with those
( xcv )
obtained by our ex-President, Mr. Merrifield. So far they
show, as many similar investigations by other naturalists
have done, that it is possible by varying the external con¬
ditions to which a butterfly is exposed in its earlier stages, to
induce a well-marked alteration in its final aspect. This is
the first and obvious conclusion to be derived from these
experiments. But Weismann, as I need hardly remind you,
was not content to stop at this point. The fact "was un¬
deniable that warmth tended to produce in the northern form
of the species an approach towards the darkened aspect of
the southern race ; and conversely, refrigeration brought
about in the latter a tendency towards the familiar golden-red
of German or English specimens. From this the further
inference seemed clear that the difference in aspect of the
two geographical races was a direct result of the temperature
conditions to which they were respectively exposed during
the course of their normal life-history. But it was further
to be remarked that in neither series of experiments was the
transformation complete. In each instance several members
of the brood appeared to be unaffected, and in none did the
artificially-produced lightening or darkening reach the same
pitch of intensity as in the local races corresponding to the
respective conditions of temperature. The most probable
interpretation of this result seemed to be that, in Weismann’s
words, the difference between the two geographical races
originated in consequence of “ a gradual cumulative effect of
the climate, the slight effects of one summer or winter having
been transmitted and added to from generation to generation.”
At this point we may imagine the neo-Lamai’ckian asking,
“What further^proof can be required of the possibility of the
transmission of acquired characters'?” It is admitted, he
would say, that the difference between the northern C. j)hlaeas
and the southern var. eleits is in some way or other the result
of climatic conditions. It has also been shown that these
conditions are only partially operative for a single generation.
The effect is therefore cumulative. It has not been suggested
in this case that the change is an adaptation ; no reason has
been alleged why in the one instance the lighter and in the
other the darker coloration should have been selected as being
( xcvi )
in any respect serviceable to the life of the individual. All
the successive steps must have resulted directly from certain
external conditions ; and each step, as it was gained, must
have been passed on to the next generation by hereditary
transmission, until, in their respective geographical habitats,
the lighter and darker coloured races became ultimately fixed.
Could there, he might say, be a much clearer case of the
validity of Lamarckian, or rather Buffonian, factors 1
Clear as the case might seem, it did not satisfy Weismann.
He took into account a further fact, viz. that this butterfly in
a certain part of its range is seasonally dimorphic, alternating
in successive broods between the pale northern and the dark
southern form. This state of things he found hard to reconcile
with the theory of transmitted somatogenic modification ; for
if, as he argued, the colour of the race were becoming
gradually darker by the inheritance of each step in the
process as it appeared in the individual, no light-coloured
brood could be produced from dark parents, and vice versa.
There was still another featui’e in the case to be reckoned
with. Weismann’s own experiments on Araschnia prorsa-
levana had taught him that the susceptibility to the influence
of temperature is not uniform throughout the whole course of
the life-history, but belongs in greatest measure to one
particular period — in the case of A. lyt'orsa-levana, as he
thought, to the early part of the pupal stage, I need here
do no more than mention incidentally that this observation
has been abundantly confirmed and extended, with modifica¬
tions, by other investigators, notably by Mr. Mei’rifield ; and,
I may add, by Mr. G. A, K. Marshall, whose results, like those
of Lt.-Col. N. Manders, are especially valuable as having
been obtained under the peculiar diflSculties of work in the
tropics, and as bringing in the operation of another factor,
namely, that of moisture.
There must, then, be some constituent in the composition
of the individual which undergoes some successive changes
during the period of the individual life-history, and which
at one or more stages of its ontogenetic development is capable
of being acted upon by external influences in such a way as
to produce an obvious alteration in the final condition. But
, ( xcvii )
we have already seen that this effect is not purely a matter of
influence upon the individual. The apparent fact of accumu¬
lation from generation to generation shows that behind the
individual change there must be some element of heredity.
Where is this to be sought 1 Weismann is ready with an
explanation which is consonant with his general theory, and
which, at any rate, has the merit of accounting for the facts.
The modifying influence, he points out, though specially
effective at a certain stage in the ontogeny, is not entirely
inoperative at other periods. The phenomenon of seasonal
dimorphism, as exemplified in the Ligurian race of
Clirysoplmmis plilaeas, seems to suggest that the suscept¬
ible constituent, whatever it may be, has passed through a
preparatory change under the influence of the climate of that
region, and so has been brought up to a point where it needs
only the finishing touch of the active influence at the specially
susceptible ontogenetic .stage to push it over into the fully
modified condition. This finishing touch is absent during the
life-history of the early spring brood, but is supplied by the
heightened temperature at which the summer brood pupates ;
hence the difference in aspect between the two emergences.
But where does this store, so to speak, of partly-prepared
material reside 1 AYeismann answers, “ in the germ-plasm.”
The constituent in question, or the antecedents thereof, exists
within the germ-plasm in a condition which allows of gradual
modification by heat, cold, or whatever the influence may be.
But inasmuch as it has not reached the specially susceptible
stage, the modification may become only slightly manifest, or
may even find no visible expression at all. Still it is there;
and inasmuch as it belongs to the germ-plasm, it forms
necessarily a part of the inheritance of the next generation,
and as such may be capable of still further advance in the
same direction ; this advance being of course limited by the
potency of the external influence and by the time during
which it has worked. AVe have here no reflection of somatic
change upon the germ-plasm, such as would be supposed to
take place under Darwin’s pangenesis and similar theories,
but a common action upon the antecedents of the final
coloration, whether these antecedents are to be found in the
PROG. ENT. SOC. LOND., V. 1910. G
( xcvili )
reserve stock of germ-plasm, or in the soma during its
ontogenetic development. Thus, according to Weismann, the
case of Chrysophanus phlaeas, instead of supporting the theory
of the inheritance of somatic modification as at first sight it
may seem to do, in reality gives additional weight to the
theory of the continuity of the germ-plasm, and its independ¬
ence of somatic change. But it may properly be called an
instance of the inheritance of an acquired character, though
not in the Lamarckian sense. The transmissible character
has been acquired ; not, however, through the medium of the
soma, but directly by the germ-plasm itself. The germ-plasm,
therefore, is indeed continuous, but not unassailable by
external influences.
This important biological result was, as we have seen,
reached by Weismann in consequence of a combined investiga¬
tion by observation and experiment on a species of butterfly ;
and it is particularly worthy of note by entomologists that all
the data which he could obtain respecting the local and
seasonal variations of the form in question proved to be of high
significance in the solution of his problem. Without them he
could not have reached his conclusion ; with further informa¬
tion on these and similar points, he could no doubt have made
his explanation still more complete. This is a good illustration
of the way in which observation in the field, experiment in the
laboratory, and reasoning in the study can combine to establish
a result of great scientific importance.
It will no doubt have occuried to you that Weismann’s
view as to the nature of the seasonal dimorphism manifested
in the Ligurian form of C. phlcLsas ought to be tested by
exposing the immature stages of the spring brood of this form
to a high temperature. So far as I am aware the experiment
has not been tried ; and until it has, this part of Weismann’s
conclusion cannot be accepted without reserve. But as to his
view of the susceptibility of the germ-plasm, there has now
been abundant confirmation ; again, be it noted, by work
done on insects.
Standfuss, as Weismann notes, bred some specimens of
Vanessa urticae under conditions of ai’tificially lowered
temperature. The perfect insects showed a departure from
( xcix )
the normal which was no doubt due to the refrigeration, — in
this case an increase of dark pigment. Eggs were laid by
some of these abnormal specimens, the resulting larvae were
reared under normal conditions, and the butterflies on
emergence again showed a divergence from the normal form
which, though slight, was in the same direction as that of
their parents. Similar results with the same species were
afterwards obtained by Weismann himself. A still more
marked confirmation was supplied by Fischer. Pupae of
Chelonia caja were subjected to an artificially lowered
temperature, and some of the resulting moths showed an
extreme degree of melanism. Two of these artificially-produced
aberrations were paired, and their offspring were reared
under normal conditions. Among the latter were specimens
which exhibited in greater or less degree a tendency towards
the artificially-induced melanism of their parents. Here we
have an excellent case of the apparent transmission of an
acquired character. In the light, however, of Weismann’s
acute reasoning on the subject of G. phlaeas, there is every
justification for believing that the same interpretation is valid
in the present case. The modification is not inherited from the
soma of the parent, but is consequent upon the direct action of
the external influence upon that parent’s germ-plasm.
But all this pioneer work, impoi’tant and excellent as it is,
must yield the palm for completeness and cogency to the
laborious researches conducted by William Lawrence Tower,
whose treatise entitled “ An Investigation of Evolution in
Chrysomelid Beetles of the Genus Leptinotarsa ” is a perfect
storehou-se of valuable biological material, all derived from
the minute study of a single genus of Coleoptera. This
admirable publication, which appeared in 1906 under the
auspices of the Carnegie Institution of Washington, has
perhaps in this country hardly met with the attention which
it deserves ; though now that Professor Bourne, in his recent
Presidential Address to the Zoological Section of the British
Association, has drawn public attention to the work, it may
be expected that greater appreciation will be shown of the
extremely interesting results which it contains.
Among the numerous and carefully-planned experiments
G 2
( c )
undertaken by Tower, there are some which have a direct
bearing on the matter under consideration. Of these I propose
to give a brief account. It should be premised that the
beetles of the genus Leptinotarsa lay their eggs in batches,
each batch being laid before the next begins development.
Experiment 1. — Males and females of L. decemlhieata were
exposed to extremely hot and dry conditions, accompanied by
low atmospheric pressure, during the growth and fertilisation
of the first three batches of ova. These were reared from the
outset in normal conditions. Out of 96 specimens so obtained,
82 were of the varietal form pallida. After the deposition of
the first three batches of ova, the same parent beetles were
placed in normal conditions, and two more batches were laid.
These, reared under natural conditions, gave normal decem-
lineata, which continued to breed true to type. The origin of
the pallida forms may therefore fairly be ascribed to the
abnormal conditions to which their parents were for a time
exposed. Two of these pallida males were mated with decem-
lineata females ; the resulting hybi’ids resembled normal
decemlineata. Kow comes an interesting point. This genera¬
tion of apparently normal decemlineata gave, when bred inter
se, offspring consisting of pallida, decemlineata, and decem-
lineataA\kQ hybrids in Mendelian proportion. The pallida of
this generation bred true, being evidently extracted recessives ;
the decemlineata and hybrids broke up again in Mendelian
fashion. This most interesting experiment points to at least
three results of great biological importance. The first, which
is that in which we are more immediately interested, is the
confirmation of Weismann’s view as to the accessibility of the
germ-plasm to external influences. The second is the demon¬
stration that just as the somatic plasm has a period of highest
susceptibility, which fact was known to Weismann at the
time of his G. jMaeas experiments, so also the germ-plasm
itself has a similar period. This was not known to Weismann ;
if it had been, it might have led him to modify in some
respects his conclusions from those experiments. The third
point is the establishment of the fact that the type-form
of a given species may stand in Mendelian relation with a
variation of that species artificially induced by interference
( ci )
with its gem-plasm. The interest and importance of these
results is manifest, and it is well to remind ourselves once
more that they all emerge from the careful study of an
insect.
Experiment 2. — Males and females of L. decemlineata were
subjected to hot dry conditions, and afterwards to normal
surroundings. During each period they deposited eggs.
From the eggs deposited in the second peidod, that of normal
conditions, pure decemlineata were bred for ten generations.
But from the eggs of the first period came many aberrant
forms, together with a certain number of apparently typical
decemlineata. These latter were reared side by side with the
decemlineata of the second period, which, as we have seen,
behaved in a pei’fectly normal manner throughout. But the
decemlineata of the first period, after giving rise to a genera¬
tion of apparently quite normal forms, suddenly developed into
a race with an annual cycle of five generations instead of the
normal two, and continued to exhibit the same abnormality
up to the fourteenth generation, when the experiment ceased.
Here we have a physiological instead of a structural pecu¬
liarity imposed upon the germ-plasm by artificial means, and
giving rise to a permanent race. It should be mentioned
that, in the words of Tower, “ none of the beetles of the
lineata group, to which this beetle belongs, have more than
two, or rarely three, generations per year, and there are none
known in the genus that have over three.” Another point
that emei’ges is that the application of the stimulus need not
always be immediately followed by the obvious response. In
the present experiment one whole generation elapsed before
the race began to show the effect of the abnormal treatment
to which its progenitors had been subjected.
Experiment 3. — In another experiment with the same
species the conditions were varied by using heat in combina¬
tion with moisture. In other respects the procedure was the
same. From the eggs of tlie second period eight successive
generations were reared, giving nothing but pure decemlineata.
From those of the first period, that of abnormal conditions,
came a large number of unmodified decemlineata and a certain
number of the form melanicum. Both of these forms bred
( cii )
true in normal surroundings. The generation also included a
single male specimen of the form tortnosa, which crossed with
a female decemlmeata yielded hybrids with decemlineata
dominant. These in succeeding generations split up in
Mendelian fashion, the tortuosa (extracted recessives) breeding
true.
Experiment 4. — The subject of this experiment was the
tropical species Leptinotarsa muUitaeniata. Here, again, the
parent beetles were exposed during the first period of egg-
laying to artificial conditions of heat and humidity, while
during the second period the surroundings were normal. The
results obtained were analogous with the preceding. The
second batch produced nothing but nmltitaeniata for four
generations, when the experiment ceased. The first batch gave
together with typical midtitaeniata a much larger number of
the form melanothorax. The two forms were se})arated, and
each continued to breed true under normal conditions.
Experiment 5. — In this experiment, the subject of which
was again L. nmltitaeniata, the egg-laying period was divided
into three instead of two portions. The first and third were
passed under normal conditions, the second under the artificial
conditions of the last experiment. The first and third
batches of eggs gave nothing but typical muUitaeniata ; the
second consisted entirely of the forms melanothorax and
ruhicunda, each of which continued to breed true.
Experiment G. — Finally, a similar experiment was tried
with another tropical species, Leptinotarsa undecimlineata.
Again the result was the pi’oduction by artificial means of a
varietal form {angustovittata), which continued to give pure
descendants showing no tendency towards reversion.
After these experiments there seems no possible room to
doubt that the germ-plasm is accessible to external infiuences,
and that it may by these means be transformed in such a way
as to give rise to a permanent race of descendants showing
marked differences from the parent form.
I will not now stop to discuss Tower’s interpretation of the
results of these and his numerous other experiments. His
theoretical views seem to me to differ in value, some being
weighty in the extreme, and a few, to say the least, question
( Piii )
able. But there can be no dispute as to the high import¬
ance and interest of the facts that he has brought together,
and I trust that the examples I have given may induce
entomologists who are interested in the study of evolution,
and by whom these researches may hitherto have been over¬
looked, to make themselves acquainted with this mine of
valuable information. They may not find themselves able to
agree with the author at all points, but they will be intro¬
duced to a great mass of material with an important beax'ing
on the nature and causes of variation, the working of
heredity, the efficacy of selection, the significance of warning
colours, and many other matters essential to a proper
comprehension of the problem of evolution.
There is one other phase of this great problem about which
I should like to say a few words, because here again our
special entomological studies have an important part to play.
They have already contributed much towards the comprehen¬
sion of this side of the question, and I am convinced that they
are capable of leading to a still further advancement of know¬
ledge in the same direction. I refer to the psychic aspect of
evolution.
It was fully recognised by Darwin himself that mental no
less than physical characters are subject to evolution. The
same principle was adopted by Wallace, not, it is true, with¬
out reservation, and received at his hands,- some interesting
developments. But to Professor Mark Baldwin belongs the
principal credit of insisting on and driving home the fact that
evolution is psychophysical ; that, as he puts it, “there are not
two evolutions, one ‘ organic ’ and the other ‘ mental,’ but that
mind and body have evolved by one process and in one series
of graduated stages.” Now in order to illustrate in a forcible
manner the interdependence of physical and psychic pheno¬
mena in evolution, he has recourse to the theory of warning
colours in insects. This he expands in the following manner :
“ As preliminary to the theory there is the fact o coloration,
which is distinctly physical. The question is as to its origin.
The theory holds it to be due to the warning given to other
individuals that a particular colouring is distasteful or
poisonous. Now in order that this warning be given, the
( civ )
biologists tell us there is necessary a certain education of the
hostile individuals. The creatures have to learn the meaning
of the coloration; and this learning involves profiting by
experience, . . . Here is as distinctly a mental process in¬
volved as any one might cite. . . . The action of natural
selection, I may add for completeness, secures the survival of
the insects so coloured, seeing that being warned, their
enemies let them alone. The possibility of the evolution of
the definite coloration turns, in fact, upon this series of
psychological processes,”
Here the psychic endowment of the enemy is seen to be a
factor in the evolution of the prey ; and it may of course be
added that to ensure for the warning colour its full effect, it
is necessary that its owner should possess the appropriate
habits. Similar instances will at once occur to every field
naturalist. Protectively-coloured insects instinctively seek
corresponding surroundings. An interesting case of this is
the fact which now seems fairly well established of yellow and
white butterflies selecting for resting-places the neighbourhood
of leaves or flowers of similar colour. Dr. Longstaff’s excel¬
lent plate, drawn from nature, of the resting habit of Eronia
cleodora will occur to many of you. I ought to apologise for
dwelling on so familiar a topic, but the fact really requires
emphasising that the raw material of selection is neither
purely physical nor purely psychical, but a complex of the
two. The admission of this fact, so patent to evei’y observer
of insects, has important consequences. For as soon as we
can recognise a rudimentary form of mentality — the gaining
of a definite result by the method of trial and error — we have
a basis for habit, for memory, and eventually for the higher
psychological faculties. In the instances that I have men¬
tioned of the correlation of appropriate habits with a pro¬
tective colouring, there is no reason to suppose the presence
of anything like conscious accommodation ; the actions in
question are no doubt instinctive in the strictest sense. But
quite low down in the animal scale, and even, I think it will
be conceded, among the insects, we get indications of “ I’esidual
pi'ocesses left by actual experiences ” leading to what can only
be called memory. These processes, of undoubted utility, may
( cv )
no doubt serve in their day, as Professor Baldwin says, until
reinforced and possibly superseded by congenital variations
tending in the same direction. Memoiy thus established as
an inborn faculty, “the experimental use of memory images,
with corresponding success and utility, would be followed in
time by further variations, giving imagination and thought ”
(Baldwin).
In the higher animals we have no difficulty in perceiving
the co-existence of individual plasticity with the congenital
endowments of both mind and body. Instinct, one of these
congenital endowments, is not “ lapsed intelligence,” though
intelligence may be suffered to lapse if and when instinct is
ready to take its place. On the other hand, the reverse
px’ocess may and does occur. Instinct may fall away and
disappear before the encroachments of the increasing power
of individual accommodation, a power which is rooted in the
plasticity, and, to use Sir E. Bay Lankester’s term, the
“ educability,” of the psychic organisation. This final stage
is naturally of most frequent occurrence in man. The relative
proportions of the two principles are regulated by natural
selection ; the best combination in regard to any given pur¬
pose is the one that will win.
Now the special application of these considerations to
Entomology I take to be this. The later stages of psycho¬
physical development are comparatively accessible to our
observation. In the phylogenetic range of the higher verte¬
brates, and the ontogenetic history of man, we have much
material for the study of the interplay of plasticity, educability
and power of individual accommodation, with the congenital
endowments of a relatively fixed and stable character, such as
instinct and the fundamental features of bodily structure.
But the earlier steps in the process are involved in much
obscurity. What do we really know, for example, of the
relative value of the parts played by psychic plasticity and
fixed instinct in the case of the lowest organisms 1 And can
we be sui’e that to any large extent the growth of instinct has
been in their case actually screened by powers of functional
accommodation 1 What we have ascertained on these points
has been, I venture to say, largely or chiefly due to the
( t'vi )
labours of entomologists. Fabre, Avebury, Wasmaun, Forel
are names that will occur to everybody as those of men who
have done much in their several ways to help on an answer to
these and kindred (juestions, and who have shown the road
which future investigators may pursue with profit. And the
study of the social hymenoptera has a further bearing upon a
development of evolutionary theoi’y of the greatest importance,
for it introduces us to the conception of the yroitp as the
selective unit instead of the individual, and so leads on to the
evolutionary side of sociology and ethics, with their accom¬
paniment of organisation within the community, of imitation,
of tradition and the social sense.
This is a tempting subject, and I would gladly pursue it
further did time permit. But I think I have now said enough
to illustrate my contention that entomologists enjoy peculiar
advantages in the attack of the great outstanding problems of
evolution. It is unquestionable that such a mind as that of
Herbert Spencer, capable of welding together into a con¬
cordant whole a mass of facts and conclusions drawn from
every department of knowledge, is an asset of enormous value
to the cause of scientific progress. But it should never be
forgotten that the work of the specialist is equally essential ;
it is he who supplies material without which the generaliser
and unifier of knowledge could not work. And let us beware
of undervaluing kinds of work which do not happen to appeal
to our own individual tastes. I think no one can give atten¬
tive considei’ation to the topics that have been touched on
this evening without seeing that our study has need of the
morphologist, the physiologist, the field naturalist, the museum
worker, the laboratory experimenter, the systematist, and —
shall I add 1 — the arm-chair philosopher. Each one of these
has his contribution to make to the common fabric of organised
and unified knowledge.
{ cvii )
GENERAL INDEX.
The Arabic figures refer to the pages of the ^Transactions' ; the Roman
numerals to the pages of the ‘ Proceedings’
The President’s Address is not separately indexed.
GENERAL SUBJECTS.
Aberrant Lepidoptera, exhibited, Ixiv, Ixviii.
Abruzzi, butterflies from the, exhibited, xlv.
Abyssinian beetle, life tenacity of, exhibited, xliv.
Acraea It/coa, and A. johnstoni, on the specific distinctions between, xlii.
Africa, book on Lycaenidae from South, exhibited, xxiii ; descriptions of
some hitherto unknown, or little-known larvae and pupae of Rhopa-
locera from South, Hi ; revision of species of genus Lycaenesthes
from, 1.
Agriades coridon, double-brooded, exhibited, xxxvii ; southern forms of,
exhibited, xlvii.
Agriades thetis {hellargus) and A. coridon, discussion of the affinities of, vi.
Algeria, descriptions of new Hymenoptera-Aculeata (Sphegidae) from, xvii,
Ixiv, 517 ; summer butterflies from, exhibited, liii.
America, mimics and models from South, exhibited, liv, Ixviii ; beetles from
North, exhibited, Ixxi.
Anthrocerids, British, exhibited, Ivii.
Ants, association of, exhibited, xiii ; and mimic beetle, exhibited, xx ; and
their nests, living, exhibited, xliv, Ixvii ; host and guest, exhibited, lx ;
brachyterous, exhibited, Ixxii ; further observations on temporary social
parasitism, and slavery in, Ixxiii ; nests, some experiments with, 142.
Apaturopsis cleocharis, exhibited, xvii.
Argentine Republic, on the Geometridae of the, xxx, 204.
Argynuid forms, Lycaenid and, exhibited, xii.
Asilid (Promachus sp.), the oothecae of an, xxvii ; and prey, exhibited, lx.
Auditors for 1910-11, appointed, Ixv.
Australia, new fossorial Hymenoptera from, xli, 407.
Beetle, male with female antennae, exhibited, xiii ; teratological example of,
exhibited, xiii ; ant and mimic, exhibited, xx; rare British, exhibited,
xxviii, XXXV, xliii, Iviii, Ixvi ; parasite on, exhibited, xxx ; new, exhibited,
xxxi, Iviii ; from crop of pheasant, exhibited, xxxvii ; life tenacity of
Abyssinian, exhibited, xliv ; abnormal, exhibited, liii ; collection of
North American, exhibited, Ixxi.
Bignell, George Carter, notice of death of, xi.
( cviii )
Borneo, new earwig from, exhibited, ii ; protective attitude of Mantis from,
exhibited, 1 ; two new species of Lycaennpsis from Sarawak, Ixxiii.
Braconid, new Britisli, exhibited, liii.
Britain, Orta musculosa in, exhibited, Ixiii.
British, Coleoptera, rare, exhibited, xxi ; species of Hydroecia, separation of,
xxiii ; beetle, rare, exhibited, xxviii, xxxv, xliii, Ixvi ; Hemiptera, rare,
exhibited, xliii ; Xylophasia, rare, exhibited, xlvii ; Braconid, new,
exhibited, liii ; Anthrocerids, exhibited, Ivii ; Lepidoptera, bred rare,
exhibited, Ixxii.
Butterflies, from the Carpathians, exhibited, xxviii ; from the Abruzzi,
exhibited, xlv ; from Algeria, summer, exhibited, liii ; from Dutch New
Guinea, on some rare and undescribed, Iviii ; from South America, close
resemblance of, Ixviii.
Callopkrys avis, bred specimens of, exhibited, xxi ; on, 85.
C'ameroons, mimetic Lepidoptera from the, exhibited, xxix.
Canadian Coleoptera, paper on, xxiii.
Carpathians, butterflies from the, exhibited, xxviii.
Cetoniidae from Uganda, exhibited, xxxvi.
Chagos Isles, descriptions of Micro-Lepidoptera from Mauritius and the,
xxxiii, 366.
Charaxes, error in account of breeding experiment with, xli.
Cicadoid genera Lemheja, remarks on, lii.
Coleoptera, rare British, exhibited, xxi ; paper on Canadian, xxiii ; during
floods, on the behaviour of, xxx, 379 ; rare and variant, exhibited, Ixxi.
Congress in August, 1910, announcement as to Entomological, i ; delegates
to, xxviii, xxxv ; photogi-aph of members at, exhibited, xliii.
Conversazione, postponement of, xxxv, xlii.
Council for 1910-11, nomination of, Ixv, Ixix.
Darwin Medal, announcement as to, Ixv, Ixx.
Dermaptera, revision of the Laliduridae, a family of the, xi, 161 ; revision of
the Lahiidae, a family of the, Ixv.
Deuterocopus, on the genus, 107,
Diplatys, a revision of genus, xxx.
Diptera, rare and variant, exhibited, Ixxi.
Drepanopsaltria, remarks on, lii.
Earwig from Borneo, new, exhibited, ii.
Ematurcja atomaria, melanism of, xxxvi.
Envelopes for storing butterflies, exhibition of paper, ii.
Euralia in Natal, experimental breeding of forms of Nymphaline genus, xiv,
xli, 498.
Fabre Memorial, plaquette for, exhibited, Iviii.
Fellows, election of, i, xii, xx, xxviii, xxxiv, xliii, lii, Iviii, Ixv, Ixx.
Floods, on the behaviour of Coleoptera during, xxx, 379.
France, living larvae from South of, exhibited, xxxvii ; Pieridae from,
exhibited, xlv.
Geometridae of the Argentine Republic, on the, xxx, 204.
Glow-worm pupa, exhibited, xxxvi.
Goldthwait, O. C., notice of death of, liii.
Hemiptera, rare British, exhibited, xliii.
Heterocera from^Japan, new species of, Iviii.
( cix )
Holoparamecus caularum, living examples of, exhibited, xxi.
Hydroecia, separation of British species of, xxiii.
Hymenoptera, -Aculeata (Spheyidae), descriptions of new Algerian, xvii, Ixiv
from Australia, new fossorial, xli, 407.
Hymenopteron, rare, exhibited, xxxv; Lycaenid in attendance on, xxxviii,
xxxix.
Ichneumon, rare, exhibited, xxxi.
India, description of a new Nymphaline butterfly from British, Ixxiii.
Insects, and their natural enemies, a few words respecting, xvii, 151 ; from
Portugal, rare, exhibited, xxxviii ; in the tropics, and their influence on
mimicry, notes on enemies of, Ixiv ; introduced and new, exhibited, Ixxi.
Italy, Pieridae from, exhibited, xlv ; Ehopalocera from, exhibited, Ixix.
Japan, new species of Heterocera from, Iviii.
Jubilee Celebration of Entomological Society of Russia, announcement as to,
ii ; address to, xi ; account of, xviii.
King, address to the, xxxiii, xlii.
Kirkaldy, George Willis, notice of death of, xi.
Labiduridae, a family of the Dermaptera, revision of the, xi, 161.
Lahiidae, a family of the Dermaptera, revision of the, Ixv.
Larvae from South of France, and imago, living, exhibited, xxxvii.
Lasiocampid nests, exhibited, xiv.
Lasiocampidae, new genera and species of, Ixix.
Latiorina (Lycaena) orhitulus, an amyrmecophelous Plebeiid butterfly, on the
early stages of, Ixix.
Lemheja, remarks on the Cicadoid genera, lii.
Lepidoptera, from the Cameroons, mimetic, exhibited, xxix ; edibility of
larvae of, xxxi ; improved photographic process for, exhibited, lii ; aber¬
rant, exhibited, Ixiv, Ixviii ; bred rare British, exhibited, Ixxii.
Leucania l-alhum, bred, exhibited, Ixii.
Luciola, description of two new species of, Ixxiii, 532.
Luflp, W. A., notice of death of, liii.
Luperina gueneei, variation of, exhibited, Ixii.
Lycaenesthes, revision of the African species of the genus, 1 .
Lycaenid, forms, exhibited, xii ; butterflies, on Zizeeria, a group of, xvii,
479 ; in attendance on Homopteron, xxxviii, xxxix.
Lycaenidae, book on South- African, exhibited, xxiii ; hybrid, exhibited,
xxxvi ; revision of the African species of the Lycaenesthes group of
the, 1.
Lycaenopsis from Sarawak, Borneo, two new species of, Ixxiii.
Malayan Micro-Lepidoptera, descriptions of, xlii, 430.
Mantis from Borneo, protective attitude of, exhibited, 1.
Mauritius and the Chagos Isles, descriptions of Micro-Lepidoptera from,
xxxiii, 366.
Melanism, of Ematurga atomaria, xxxvi ; in Melitaeids, exhibited, Ixxiii.
Melitaea aurinia, new forms of, exhibited, liii.
Melitaeid, variety, exhibited, Ixiv ; melanism in, exhibited, Ixxiii.
Micro-Lepidoptera, from Mauritius and the Chagos Isles, descriptions of,
xxxiii, 366 ; descriptions of Malayan, xlii, 430.
Mimicry, mimetic Lepidoptera from the Cameroons, exhibited, xxix ; mimics
and models from South America, exhibited, liv, Ixviii ; notes on insect
^ ( cx )
euemies in the tropics, and their influence on, Ixix ; butterfly and mimic,
exhibited, Ixxiii.
Natal, experimental breeding of forms of Nymphaline genus Euralia in, xiv,
xli, 498.
New Guinea, on some rare and undescribed butterflies from Dutch, Iviii.
Nymphaline, genus Euralia in Natal, experimental breeding of forms of, xiv,
xli, 498 ; butterfly from British India, description of a new, Ixxiii.
Obituary. Edward Saunders, F.R.S., G. W. Kirkaldy, G. C. Bignell,
A. Piffard, xi; G. S.’ Saunders, F.L.S., xxxi; O. C. Goldthwait, \V. A.
Luff, liii.
Officers for 1910-11, nomination of, Ixv, Ixix.
Orgyia splenclida {cJubia), on the larva of, Ixix.
Oria musculosa in Britain, exhibited, Ixiii.
Orthoptera, third paper on the Tetriginae in the Oxford University Museum,
xvii, 346.
Os»refa-species of the adtoica-gvowp, further notes on two, xvii, 155.
Oxford University Museum, third paper on the Tetriginae (Orthoptera) in
the, xvii, 346.
Palaearctic moth, rare, exhibited, Ixvi.
Paper envelopes for storing butterflies, exhibition of, ii.
Parasite, on beetle, exhibited, xxx.
Pheasant, beetles from crop of, exhibited, xxxvii.
Photographic process for Lepidoptera, improved, exhibited. Hi.
Pieridae from France and Italy, exhibited, xiv.
Piffard, Albert, notice of death of, xi.
Portugal, rare insects from, exhibited, xxxviii.
President for 1910-11, nomination of, Ixv, Ixix.
Promachus, the oothecae of an Asilid, xxvii.
Ptinella britannica, example of, exhibited, xxx.
Rhone Valley, Rhopalocera from, exhibited, ii.
Rhopalocera, from Rhone Valley, exhibited, ii ; descriptions of some hitherto
unknown, or little-known larvae and pupae of South African, Hi ; Italian,
exhibited, Ixix.
Ehopalosomidae, with def5cription of a second new species, on the position of
the, xli, 386.
Russia, Entomological Society of, announcement as to Jubilee Celebration of,
ii ; address to, xi ; account of, xviii.
Saunders, Edward, notice of death of, xi.
Saunders, G. S., notice of death of, xxxi.
Saw-flies, on the saws of, xvii ; oviposition of, xxxv.
Scoliidae, notes on the, xli, 391.
Seal of the Society, new, exhibited, xlii.
Sesia andreniformis, ova of, exhibited, xxxvi.
Smerinthus, variation in, exhibited, xxxvii.
Sphegidae, descriptions of new Algerian Hymenoptera-Aculeata, xvii, Ixiv,
517.
StriphnopUrygidae, new genera and species of, Ixix.
Stylopidae from Australia, on a new genus of. Hi, 514.
Tapinostola, ova and appendages of, exhibited, xxi.
Teratological specimens, exhibited, lx.
( cxi )
Tetriginae (Orthoptera) in the Oxford University Museum, third paper on
the, xvii, 346.
Travel grant, announcement as to, Ixxi.
Trimen, K., letter of congratulation to, Ixx.
Uganda, Cetoniidae from, exhibited, xxxvi.
Uropteryx samhucaria in connection with their colour surroundings, experi¬
ments with the larva and pupa of, Ixiv.
Variation, in AS'mgj'tKtAMs, exhibited, xxxvii ; in Xanthia ocellaris, exhibited,
Ivii ; of Luperina gueneei, exhibited, Ixii.
Vice-Presidents for 1910-11, nomination of, i.
Xanthandrus comtus, notes on, xxv.
Xanthia ocellaris, variation in, exhibited, Ivii.
Xylophasia, rare British, exhibited, xlvii.
Zizeeria, a group of Lycaenid butterflies, on, xvii, 479.
Zygaenid forms, exhibited, xiii.
( cxii )
SPECIAL INDEX.
The Arabic figures refer to the pages of the ' I'lvinsactions' ; the Roman numerals
to the pages of the ‘ Proceedings'
abnormis (Cylindrogaster), 187
,, (Rhopalosoma), 386, 388
Abraxas, Ixviii, 152, 153
Abraxinae, 280
abrupta (Tiphia), 395, 396
acaciae (Nordiuannia), xlvi
Acanthostethus, 428
Acasis, 243
acerbata (Anapalta), 249
achilles (Morpho), lii
Achlora, 211
Acidalia, 205, 214, 215, 216, 217, 219,
220, 222, 223, 224, 226, 227, 228,
2-29
Acidalidae, 213
Acidaliinae, 213, 274
Acniophyllum, 346, 349, 355
Acraea, xlii
Acraeidae, xxix, Ixviii
Acratodes, 222
Acrocercops, 373
Acronycta, Ixi
Actinote, Ixviii
actinote (Dismorphia), Ixviii
Actizera, 483, 494, 495, 496, 497
Aculeata, 389
acuminata (Melanophila), xliii
adela (Acidalia), 222, 223
,, (Haraalia), 222
,, (Lipomelia), 222
adelaidae (Tachysphex), 425
adherbal (Lycaenesthes), 33, 38, 40
adippe, var. cleodoxa (Argynnis), xlvi
adsciti (Ichneumones), 389
adumbrata (Spododes), 308
adunca (Osmia), xvii, 155, 158, 159
adusta (Tachyuomyia), 407
,, f. uniformis (Tachynomyia), 408
adustus (Thynnus), 407
advena (Anisolabis), 176
advenaria (Epioiie), Ixiv
aegon (Lycaena), 487
aegoii (Plebeius), xlvi
Aelurus, 407
aemula (Herbita), 310
aenea (Honorana), 319, 321
,, (Salpis), 320, 321
,, (Tachytes), 523
aenigmatica (Sibylliua), 388
Aeolothynnus, 413, 416, 417
aequalis (Hypsaeus), 347, 350
aerata (Synchlora), 211
aetheria, var. algerica (Melitaea), liv
aethiops (Eriirliinus), Ixvi
affiiiis (Stilicus), 384
affirmata (Calocalpe), 265, 266
,, (Scotosia), 265
afra (Lycaenesthes), 16, 27
africanus (Prototettix), 361
afrum (Ecliinosoma), 164
Agaristidae, xxix
agathon, var. phryxe (A])oria), Ixxiii
aggerata (Meridarchis), 430
agilis (Dielis), 404
aglaia (Argynnis), xlvi, Ixiv
Agi’iades, vi, vii, viii, ix, x, xi, xii,
xxxvi, xxxvii, xlvii, xlviii, xlix, 1,
Ixviii, Ixix
Agriomyia, 408, 409, 410
alaris (Discolia), 403
albarita (Heterolocha), 289
,, (Melinoides), 289
alberta (Lycaenesthes), 16, 28
albidulata (Acidalia), 227, 228, 229
,, (Craspedia), 227
albimaculata (Amauris), xiv, xv, 499,
506, 508, 510
albipes (Anchomenus), 381, 383
,, (Teclinomyrmex), Ixxii
albipunctaria (Salpis), 320
albipunctatus (Deuterocopus), 112, 122
albipuiictella (Sochchora), 141
albirenata (Ira), 310
albocoeruleus (Zizera), 486
( cxiii )
albosignata (Philereme), 259
,, (Scotosia), 259
albovittata (Anisolabis), 176
albulata (Perizoma), 269
alceae (Erinnys), xlvi
alcetas (Everes), xlvi
Aids, 335
Aletis, xxix
alienus (Lasius), Ixxii
Alindria, xliv
Allostethinae, 161, 162, 163, 176
Allostethus, 162, 163
Allotiiius, xxxviii, xxxix, xl, xli
Ahnodes, 209, 210
alniaria (Ennomos), 305
alope (Zizera), 486
alaus (Coenonympha), 496
,, (Zizera), 479, 480
altaica (Crosita), xxxvii
alternata (Coremia), 261
,, (Epirrhoe), 249
altliaeae (Erinnys), xlvi
alveus (Hesperia), xlvi
ainauda (Polyommatns), xlvi
amarah. (Lycaenesthes), 17, 38
Amaurinia, 279
Amauris, xiv, xv, 498, 499, 501, 506,
508
amberia (Boarmia), 339
,, (Hymenomima), 339
ambigua (Euborellia), 180
Amblurodes, 327
Ameria, 233
americana (Forcipula), 186
,, (Psalis), 181, 182
ainethystata (Bassania), 308
ammorrhoa (Eupithecia), 278
amnicincta (Azelina), 315
,, (Pero), 313, 315, 316, 317
amoena (Pacliropbylla), 241
Amphidasys, 343
amphitricha (Epiphractis), 373
amplificata ( Heterusia), 234
anacardii (Salamis), 511
anadema (Lycaenesthes), 16, 22
Anaitis, 236
Anapalta, 249
Anaphantis, 472
Anaphe, xiv
Andiomenus, 381, 383
Ancylometis, 371
andereggaria (Catasda), 330
andreinii (Anisolabis), 180
andreniforniis (Sesia), xxxvi
angulatus (Coptotettix), 363
,, (Hypsaeus), 347, 349
angulifera (Anisolabis), 176
anguliferata (Cidaria), 270
PROG. ENT. SOC. LOND. , V. 1910.
angustatum (Sphecosoma), Iv
angustatus (Dyscbirius), xxi
angustivertex (Euparatettix), 360
Anisodes, 214, 215
Anisolabis, 163, 168, 169, 170, 171,
174, 175, 176, 177, 178, 179, 180,
181, 188
Auisoperas, 306, 307
anisojjhthalnia (Antizera), 494
annandalei (Euborellia), 181
annulata (Dielis), 405
,, (Elis), 405
,, var. ferrugineipes (Dielis), 405
,, „ (Elis), 405
anuulicornis (Anisolabis), 176
annulipes (Anisolabis), 175, 176, 177
,, (Coptotettix), 364
anomala (Haminaptera), 246
Anomalon, 389, 390
Anomphax, 212
antanossa (Zizera), 481, 482, 484
,, (Zizina), 483, 488, 492, 495,
497
antenna (Anisolabis), 176
antenuata (Salpis), 320
Anteois, 217, 220, 221
anteviridata (Cidaria), 273
anthedon (Euralia), xvi, 498, 499, 507,
509, 510
Anthobosca, 391, 397, 398, 399
Anthrocera, xiii, Ivi, Ivii, Iviii
Anticlea, 252
antipodes (Cerceris), 422
antis (Astylus), Iv
Antizera, 481, 494
antoni (Anisolabis), 176
Antygophanes, 320, 324
Apaturopsis, xvii
apicalis (Cymatophora), 341
,, (Melanolophia), 341
apicata (Haramaptera), 245
Apida, 301, 302, 303, 304, 305, 306
apiciata (Geometra), 211
Apiciopsis, 306
apicitnmcaria (Thysanopyga), 301
Apion, 385
Aplodes, 207, 210, 211
apollo (Parnassius), xxviii
,, var. carpaticus (Parnassius),
xxviii
Aporia, xlvi, Ixxiii
aporonoma (Anisolabis), 175, 176
apparitaria (Acidalia), 227
appendiculata (Haeraouia), xliv
approbata (Acidalia), 227
approximans (Atyria), 231
,, (Atyriodes), 231
apricaria (Heterolocha), 288
H
( cxiv )
apiicaria (Melinoides), 288, 289 '
,, (Metrocampa), 288 |
arabica (Aiitliobosca), 397, 399 |
arana (Caripeta), 326
,, (Eriloj)hodes), 326
,, (Leucolithodes), 326
arcauia (Coenonympha), xlvii
,, var. insubrica (Coenonympha),
xlvii
arcanioides (Coenonympha), liv
arciel (Epactiothynnns), 415
Ardiosteres, 476
arenisca (Macaria), 294
arenosa (Nematocampa), 287
arcnulavia (Sabulodes), 308
arescopa (Cnpidesthes), 6, 9
ar^ante (Cato]isilia), lii
argentilinea (Ratiaria), 287
argentina (Coremia), 257
,, (Ochyria), 256, 257, 269
,, (Rhopalodes), 240
ai'gia (Zizera), 486
argica (Eupithecia), 277, 278
argilai'ia (Cymatophora), 341
argiolus (Celastrina), xlvi
,, var. sikkima (Lycaenopsis),
490
argopa (Ethniia), 461
Argynnis, xlvi, Ixiv
argyrognomon (Plebeius), xlvi
argyrophaea (Tisis), 439
Argyroploce, 435, 436
Aricia, xlvi
aridnla (Cryptolechia), 456
arion (Lyeaena), xlvi
arispe (Caligo), lii
Arixenia, ii
Arizela, 207, 324, 325
Arlex, 189, 190, 191
armata (Enborellia), 180
armiger (Aeolotliynnu.^), 417
arnobia (Ani])hidasys), 343
,, (Phalaena Geometra), 343
,, (Thyrinteina), 343
aiticulatum (Bembidium), 383
arnndinis (Nonagria), xlvii
ascanius (Papilio), lii
asceta (Ptychopoda), 217
Ascotinae, 280
As('llodes, 213
asina (Drepanodes), 302
a.sinana (Herbita), 309
Aspilates, 208, 221, 273, 291, 326, 327
Aspitates, 221
assimile (Bembidium), 383
Astatus, 519, 520, 521
asthcnaria (Acidalia), 215, 216
,, (Deptalia), 216
Astheninae, 231
Asthenothynnus, 411, 413, 414, 415
astraea (Antizera), 494
astrarche (Aricia), xlvi
Astyliis, Iv
Atemeles, 148
aterrima (Phyniatocera), xxxv, xxxvi
aterrimus (Pterostichus), Iviii
athalia (Melitaea), Ixiv
Athrypsiastis, 457, 458
atlanticus (Hydroecia), xxiv
atoniaria (Acidalia), 227
,, (Ematurga), xxxvi
atrapex (Deuterocopus), 112, 115, 118,
120, 121, 135
atrescens (Coryptiluni), 475
atridiscata (Acidalia), 229
,, (Craspedia), 229
atrigemmata (Actizera), 483, 494, 496,
497
,, (Antizera), 481, 495
,, (Zizera) 482
atropesaria (Choerodes), 308
,, (Sabulodes), 308
atrosignata (Imma), 465
Atteria, 433
Atteva, 470, 471
Atyria, 207, 229, 230, 231
Atyriodes, 230, 231
Auophylla, 211
aurantiaca (Cerceris), 420
aurea (Lycaenesthes), 20, 59
aurelia (Melitaea), xxviii
auricularia (Forficnla), 179
aurinia, var, aurunca (Melitaea), liii
,, var. comacina (Melitaea), liii
auritula (Luciola), 533
aurora (Eudule), 233
aurulenta (Dielis), 404
australis (Cerceris), 422
,, (Spilomena), 418
austriacaria (Larentia), 252
Austrostylops, xliii, 514, 515
Autanepsia, 221
autononia (Brachmia), 369
avia (Labiduromnia), 18S
avis (Callophrys), xxi, xxii, xxiii,
xxxviii, 85, 86, 87, 88, 89, 90, 91,
92, 93, 94, 97, 98, 99, 100, 101, 102,
103, 104, 105, 106
Azelina, 288, 307, 312, 313, 314, 315,
316, 318, 320
azteca (Anisolabis), 169
baccata (Camptogramma), 261
Badister, Ixxii, 383
balanota (Atteva), 471
balistaria (Ptychopoda), 219
ballus (Thestor), xxxvii
( cxv )
balteolata (Larentia), 259
barberae (Lycaena), 495
Barylypa, 390
basina (Xanthyris), 230
Bassania, 308
Bassarodes, 459
bathychasma (Deuterocopus), 112, 130
batida (Eupitliecia), 275
batis (Hammaptera), 245
baynei (Pseudobrephos), 328, 329
Bayiiia, 208, 242, 243
Bedellia, 374
bellargus (Agriades), vi, xi, xxxvi,
xlviii, Ixviii
,, (Polyoinniatiis), xlvi
bellimargo (Eupithecia), 276
,, (Tephroclystia), 276
Bembidium, xxi, lix, 382, 383
benesignata (Almodes), 210
bengalensis (Labidura), 185
,, (Lycaenesthes), 1, 14
benigna (Scoparia), 366
bernsteiniella (Dasycera), 440
beryllitis (Tortyra), 462
bicineta (Coricia), 290
,, (Loniographa), 290
bicolorella (Tisis), 437, 440
bicornis (Hyineiiopus), 1, li
bicurvata (Pero), 314, 316
bidentata (Odontopera), xxxi
bifoveolata (Brachylabis), 196
,, (Metisolabis), 196
bigemma (Idioglossa), 373
bihe (Lycaenesthes), 17, 34
bilineatus (Coptotettix), 364
liiinaculata (Snellenia), 440
bimaculatus (Stenus), 383
bipartitus (Epactiothynnus), 416
bipunctata (Tetrix), 359
bipunctatum (Conosoma), xxi
bipunctatus (Stenus), 382, 383
bipustulatus (Badister), Ixxii, 383
biskrensis (Tachytes), 521
Biston, 152, 343
Bistoninae, 280
bitje (Lycaenesthes), 19, 50
bjerkandrella (Choreutis), 374
Boarmia, xxxi, 205, 209, 332, 333, 334,
335, 337, 338, 339, 343
Boannidae, 259
Boarmiinae, 208, 280
bogotata (Epione), 308
bolivari (Echinosoma), 164
boops (Astatus), 520, 621
Borellia, 168, 179
boi-mansi (Anisolabis), 176
Bormansia, 189
botydaria (Acidalia), 217
botydata (Acidalia), 224
,, (Hamalia), 224
Braccinae, 207
brachicera (Dielis), 403
Brachmia, 369, 450, 451
Brachylabidae, 202
Brachylabinae, 161, 162, 188, 189, 190,
191
Brachylabis, 190, 191, 192, 193, 195,
196, 198, 199, 201
brachymorpha (Platyptilia), 367
Braconidae, Iv, 389
branchiodes (Argyroploce), 435
brasiliensis (Thermastris), 166
brassicae (Ganoris), 152
,, (Mauiestra), 152
braueri (Isolabis), 196
,, sub-sp. donisthorpii (Preno-
lepis), Ixxii
Brenthia, 467, 468, 469
Bronthis, xiii, xxviii
brevibractea (Echinopsalis), 166, 167
brevilinea (Sarraceua), 273
britannica (Ptinella), xxx
Bronchelia, 207, 342, 343
Brotis, 301
Bruchus, Ixxii
brunnea (Lycaenesthes), 13
,, (Propyragra), 166, 167
brunneata (Sphacelodes), 301
brunneri (Anisolabis), 175, 177
bruuneus (Cupidesthes), 7, 13
,, (Lycaenesthes), 13
brunnipes (Stenus), 383
bryifera (Syrtodes), 327, 328
Bryoptera, 326, 332
buccholzi (Lycaenesthes), 19, 49, 50
bucculentus (Prototettix), 362
bucephala (Pygaera), 152, 153
bufo (Trachytettix), 348
buphthalmus (Stenus), 383, 384
burri (Psalis), 182
,, (Pseudisolabis), 188
buscki (Propyragra), 166, 167
butleri (Lycaenesthes), 34, 35
Byssodes, 286, 287
Cacoecia, 432
Cacogamia, 437
caeca (Anisolabis), 176
caelicola (Brenthia), 468
caementaria (Osmia), 155, 157
caerulea (Lycaenesthes), 82
,, (Triclema), 72, 82
Calathus, xiii
calbisaria (Apicia), 302, 304
c-album (Polygonia), Ixiv
Caligo, lii
callichalca (Monopis), 474
H 2
( cxvi )
Callimorpha, 345
Callipia, 207, 238
Callipseustes, 327
Callophrys, xxi, xxii, xxiii, xxxviii,
85, 86, 87, 88, 89, 90, 91, 92, 93, 94,
97, 98, 99, 100, 101, 102, 103, 104,
105, 106
Calocalpe, 265, 266
Calopteron, Iv
calycodes (TineaJ, 474
Calyptocome, 225
Caiiibogia, 226, 280
Camilla (Linienitis), xlvi
campestris (Cicindela), xxx
Campsomeris, 400, 403, 404
Camptogramma, 258, 259, 260, 261,
268
caiiadaria (Melanolopliia), 341
canaliculata (Drusilla), 385
,, (Tiphia), 397
canis (Ctenocephalus), Ixvi
capitalis (Tachysphex), 526
capitaria (Nipteria), 290
capona (Herbita), 310
capucina (Xyloloclia), 307
Capys, 3
Carabus, xliii
carcharias (Crocanthes), 444
Carcinophora, 181, 183
carcinota (Lyonetia), 374
carfinia (Thysanopyga), 301
Caripeta, 326
carlinae, var. cirsii (Hesperia), xlv
carnea (Catascia), 208, 330, 331
carneitincta (Salpis) 208, 320, 321,
822
Carphotricha, Ixxi
Carposina, 431
Carposiuidae, 430
Carsia, 236, 239, 257
carthami (Hesperia), xlvi
Cassiopeia (Pseudasellodes), 213
cassununga (Megauthopus), liv
Castalius, 83, 495
castetsi (Psalis), 182
castina (Xantliyris), 230
Catascia, 208, 330, 331
Cataspilates, 291
Catocala, lii
Catophoenissa, 328, 329
Catopsilia, lii
caudelli (Metisolabis), 196 198
caularum (Holoparamecus), xxi
cauqiienenais (Scotosia), 267
,, (Triphosa), 266, 267
, , inca (Triphosa), 267
Celastrina, xlvi
celosoides (Boarmia), 335
celosoides (Steiialcidia), 335
celtis (Libythea), lx
censoria (Platyptilia), 367
centrifuga (Tortyra), 463
ceiitrostrigaria (Coremia), 262
,, (Orthonama), 262, 263
cepliise (Callimorpha), 345
Cerberus (Trichura), Iv
Cerceris, 420, 421, 422, 423, 424
certata (Calocalpe), 265, 266
Cerura, 151
cervinalis (Calocalpe), 265, 266
cervinicolor (Anisoperas), 307
cervus (Lucanus), 153
Cetoniidae, xxxvi
Ceuthorrhynchideus, xxxv
Ceuthorrhynchus, xxxv
chalybaeella (Tisis), 437
chauchaui (Cymatophora), 331
,, (Gnophos), 208
chandala (Zizera), 486, 487
charadraea (Tisis), 439
Charaxes, xv, xli, 506
Chelidura, 194
chelophora (Procalyptis), 431
chephise (Melanchroia), 345
,, (Phalaena Geometra), 345
Chesias, 235, 236, 238
cliibcha (Atyria), 207, 229
,, (Flavinia), 229
,, ab. cuneifera (Cyllopoda), 229,
230
chilenaria (Colotois), 320
,, (Salpis), 320, 323
chilensis (Brachylabis), 190, 192, 193
chillanensis (Larentia), 266
Chilopora, 384
cliincha (Eupithecia), 276
chirinda (Lycaenesthes), 16, 27
chlamydaria (Sarracena), 273
Chloraspilates, 273
Chloridolum, xiii
chloroelystata (Amblurodes), 327
,, (Syrtodes), 327
chlorophaearia (Cidaria), 327, 328
Choerodes, 308
chomatias (Osphretica), 475
chontalia (Pyragra), 166
Clioreutis, 374
Chrismopteryx, 208, 252, 253, 331
chryseostictus (Neurellipes), 66, 67, 68
Chrysididae, xix
chrysitis (Plusia), 151
Chrysophanus, 3, 98
Ciampa, 255
Cicadula, xlii
cicatriculata (Hoplolygris), 207, 255
„ (Lygris), 255
( cxvii )
cicatriculata (Pehirga), 255
Cicindela, xxx
cicindeloides (Stenus), 379, 380, 381
Cidaria, 205, 244, 245, 248, 249, 250,
260, 261, 262, 270, 273, 327, 328
Cidariophanes, 327
ciucticollis (Psalis), 182, 183
cinerasceus (Tinissa), 476
Cionus, lix
cisplatensis (Eueupithecia), 274, 275
citrata (Dysstroma), 268, 271
,, (Geometra), 271
,, (Phalaena), 271
citrinata (Perusia), 289
citrogastei- (Leptodeuterocopus), 138
Cladonotae, 347, 355
Cladoramus, 346, 348, 354
claustrata (Lecithocera), 448
Clavelia, xxxv
Claviger, lx, 145, 146
clavipennis (Pterygopterus), Iv
clavipes (Rhopalotettix), 357
cleistostoma (Tretothorax), lix
cleochai'is (Apaturopsis), xvii
Cleopatra (Gonopteryx), Ixix
Cleoia, 340
clorinde (Gonepteryx), Hi
clypeatus (Piezotettix), 347
Clysia, 307
Cnemodes, 214
Cnephora, 318
Cnethocampa, xxvii
Coenocalpe, 262, 264
Coeuonymplia, xxviii, xlvi, xlvii, liv,
480
coerulea (Amaurinia), 279
coerulescens (Dianous), xxx, 379, 383,
384
Colias, Ixix
collaris (Myrmedonia), 385
Colohopsis, XX
colonella (Ethinia), 462
colorata (Erilophodes), 326
colossea (Anisolabis), 168, 175, 176
Colotois, 320
colubraria (Pantherodes), 291
Columbiana (Psalis), 183
combustaria (Eucosmia), 244
,, (Hammaptera), 244
,, (Larentia), 244
comitata (Pelurga), 255
comma (Pamphila), Ixix
comminuta (Marragodes), 281
,, (Narragodes), 281
commixtata (Iridopsis), 340
commotaria (Cymatophora), 341
,, (Stenalcidia), 333
compensata (Acidalia), 227
completa (Hypolepis), 272
compressa (Anisolabis), 177
comtus (Xanthandrus), xxv, xxvi,
xxvii
concophanes (Sobareiitis), 470
concolor (Echinosoma), 164
,, (Tapinostola), xxi
concoloraria (Acidalia), 226
concomitans (Dicliromatopodia), 216
confertaria (Acidalia), 226
confrater (Sphex), 420
Congo (Forcipula), 187
congoata (Scotosia), 265
congolense (Echinosoma), 164
Conops, Ixxi
Conosoma, xxi
conradti (Acmophyllnm), 349, 355
,, (Tiphia), 394
conspicua (Atteva), 471
consti'icta (Hammaptera), 245
continua (Myzine), 392
,, (Plesia), 392
contorta (Azelina), 288
,, (Semiothisa), 288
convexirs (Coptotettix), 364, 365
Coptotettix, 363, 364, 365
Coremia, 245, 257, 261, 262
coriacea (Brachylabis), 192, 193
coriacella (Stagmatophora), 372
coriarius (Odontomaclius), lix
Coricia, 290
coridon (Agriades), vi, vii, viii, ix, x,
xi, xii, xxxvi, xxxvii, xlviii,
Ixix
,, var. albicans (Agriades), x
,, var. arragonensis (Agriades), x,
xii
,, ab. aurantia (Agriades), ix
,, var. borussia (Agriades), x
,, ab. caerulescens (Agriades), x,
xii
,, var. caucasica (Agriades), ix, x
,, var. constanti (Agriades), x,
xxxvii, xlvii
,, var. corydonis (Agriades), xi
,, var. corydonius (Agriades), ix,
X, xi
,, ab. excelsa (Agriades), ix
,, var. hafiieri (Agriades), ix
,, var. hispana (Agriades), ix, x
,, var. et ab. marginata (Agri¬
ades), X
,, var. meridionalis (Agriades), x,
xlvii, xlix, 1
,, var. meridionalis, f. constanti
(Agriades), x, 1
,, var. meridionalis, f. rezniceki,
(Agriades), x, xlix, 1
( cxviii )
coridoii var, olympica (Agriadea), x, xi
,, var. rezniceki (Agriades), x
,, f. semisyngrapha (Agriades), ix
,, ab. suavis (Agriades), xlviii
,, var. syriaca (Agriades), ix, x,
xi
,, ab. unicolor (Agriades), ix
,, (Polyomrnatus), xii, xlvi
,, var. appeiinina (Polyoiniaatus),
xlvi, 1
Correbidia, Iv
corrivulata (Epirrhoe), 249, 250
,, (Euphyia), 249
corticeata (Camptogramma), 268
corvaria (Phalaena), 345
Corycia, 289
Coryptiliiin, 475
Cosilidae, 390
Cosiliuae, 397
Cosinopterygidae, 372, 452
Cosniopteryx, 372, 452
costata (Eudule), 231
costipuncta, ab. ocellata (Cyclomia),
300
Coydalla, 449
Crabro, 429, 525
crambalea (Epimactis), 458
Craspedia, 226, 227, 228, 229
Craspediopsis, 221
crataegi (Aporia), xlvi
crawshayi (Lycaenesthes), 18, 45
Cremastogaster, xx
crenulatus (Oladorainus), 348, 354
Crepidodera, lx
crinanensis (Hydroecia), xxiv, xxv
crinita (Dielis), 403
,, (Elis), 403
,, (Scolia), 403
Criocephalus, Ixvi
crisiifrons (Xerophyllum), 352
cristitrons (Xerophyllum), 351
cristulata (Pantelia), 348
Crocantbes, 442, 443, 444, 445
Crocota, 231
Crosita, xxxvii
cruciata (Scordylia), 234
cruciferaria (Odontothera), 208
cruciferia (Aspilates), 327
,, (Odontothera), 327
crypsichola (Carposina), 431
Cryptoleehia, 456, 458
Cryptopbagus, xliv
Cryptophasa, 457
Ctenisolabis, 190, 191, 195, 196, 197,
198
Ctenocephalus, Ixvi
cubitalis (Sapheneutis), 377
culta (Discolia), 404
culta (Scolia), 404
cultratus (Piezotettix), 347
Cupidesthes, 2, 4, 5, 6, 7, 9, 10, 11,
12, 13
Cupido, xlvi
cupraria (Crocota), 231
,, (Eudule), 231, 232
curriei (Dasyleurotettix), 359 ■
curvicornis (Lissotes), liii
curvifascia (Nipteria), 290
curvilinea (Broncbelia), 343
custodiata (Coenocalpe), 262
cyanae (Cicadula), xliii
cyanolycliiia (Placoptila), 453
cyanoplaca (Ptilosticha), 440, 441
cyanosoma (Eretmocera), 454
Cyclomia, 300
Cylindrogaster, 187, 197
Cyllopoda, 229, 230
Cyllopodinae, 213
Cymatophora, 331, 332, 333, 341
Cymopsis, 344
cynarae (Hesperia), xlv
cynorta (Papilio), xxix
cypridia (Tortyra), 462
Cyrtorrhinus, xliii
cytherea (Mennis), 232
cytis (Antizera), 494
dacryodes (Cosmopteryx), 372
daemona (Euralia), 509
daplidice (Pontia), xlvi
dardanus, sub-sp. cenea (Papilio), xv,
xvi, 508, 510
,, f. hippocoon (Papilio), 508,
510
,, f. trophonius (Papilio), 508
Dasycera, 441
Dasyleurotettix, 359
debilis (Psalis), 182
Decadarchis, 473
deceptor (Euralia), xiv, xv, xvii, 498,
499, 500, 501, 502, 503, 511, 512
deceptrix (Numia), 289
declaratus (Stenus), 383, 384
decolyi (Forcipula), 186
decreta (Zizera), 489
decurtaria (Ira), 310
Dielis, 402, 403, 404, 405
definita (Hammaptera), 245
,, (Lycaenesthes), 16, 26, 27
defixaria (Acidalia), 227
defixata (Rumia), 289
Deileptenia, 209
Deiliniinae, 280
delinquaria (Iridopsis), 340
delospila (Zizera), 495
demissaria (Ptychopoda), 217
Demogorgon, 185
( cxix )
densata (Hymenomima), 338
dentata (Thoradonta), 359
denticularia (Boarmia), 332
denticulata (Imma), 464
,, (Polygraphodes), 226
,, (Spartopteryx), 327
dentilineata (Monoctenia), 320
,, (Salpis), 320
dentipes (Dinetus), 530
Deptalia, 215, 216
derivata (Anticlea), 252
Dermaptera, xi, 161, 193
Dermatoptera, xix
designata (Ochyria), 257
desilineata (Orthonania), 263
despecta (Stenalcidia), 335
detexta (Bronchelia), 342
Deuterocopus, 107, 109, 110, 111, 112,
113, 115, 117, 118, 120, 121, 122,
124, 126, 127, 128, 129, 130, 131,
132, 133, 134, 135, 136, 137, 138
Deuteroscopus, 109, 111, 118
Devarodes, 344
Diadema, 498
diademata (Epeira), 152
Dianous, xxx, 379, 380, 383, 384
Dianthoecia, Ixxii
dibapha (Catophoeiiissa), 329
,, (Epimecis), 329
Dichromatopodia, 216
dictynna (Melitaea), xxviii, Ixiv
,, var. vernetensis (Melitaea),
Ixiv
dictynnoides (Melitaea), xxviii
didyma (Melitaea), xlvi
differens (Stenalcidia), 333
diffusa (Euralia), 509
Diliacos, 400
dilucida (Ancylometis), 371
diluta (Zizera), 486, 487
dilutata (Oporinia), 269
dimidiaria (Boarmia), 333
,, (Stenalcidia), 333
,, (Tephrosia), 333
dimidiatipemiis (Dielis), 403
dimidiatus (Aeolothynnus), 417
Dinetus, 530, 531
Diplatys, xxx, 189
Diplura, xxxviii
Discolia, 403, 404
discolor (Coptotettix), 364
discreta (Decadarchis), 473
discriminaria (Acidalia), 229
Dismorphia, Ixviii
dismutata (Rhopalista), 243
dispar (ObiJa), 264
distans (Parasemia), 293
distant! (Echinosoma), 164
Ditlialaraa, 221
Ditliecodes, 221
ditiorana (Glyphipteryx), 374
ditrapezium (Noctua), Ixiv
divisaria (Thysanopyga), 301
divulsata (Scotosia), 264
doddii (Crabro), 429
dohrni (Psalis), 182
,, (Pyragra), 166
doriae (Allostetlius), 163
,, (Forficula), 163
dorilis (Loweia), xlvi
doris (Melodryas), 472
Drepanodes, 301, 302
Drepanopsaltria, lii
drucei (Euralia), 499, 509
Drusilla, 385
Dryas, liv
dubia (Euralia), xvi, 498, 499, 505,
509, 510
,, var. Cerberus (Euralia), 499, 511
,, (Flavinia), 230
(Orgyia), Ixix
dubiferata (Scotosia), 265
dubitata (Triphosa), 267
dubronii (Anisolabis), 178
ducalis (Larrada), 425
,, (Liris), 425
dukinfieldia (Boarmia), 334
,, (Stenalcidia), 334
dulcis (Arizela), 207, 325
,, (Lycaenesthes), 37
durnfordi (Atyria), 207, 230
,, (Flavinia), 230
Dyschirius, xxi, 384
Dyscia, 300
Dysstroma, 268, 270, 271
eatoni (Harpactopus), 518
,, (Spliex), 518
,, (Tachysphex), 525
eburneata (Acidalia), 226
eclieria (Amauris), xiv, xv, 499
Echinopsalis, 165, 166, 167
Echinosoma, 163, 164
Echinosomatinae, 161, 163, 165
eclipes (Acidalia), 228
Ectropis, 342
edmondsii (Hypochroma), 257
,, (Xanthorhoe), 208, 257
edna (Phibalapteryx), 286
oduardi (Tachysphex), 527
eductata (Cidaria), 245, 249
,, (Euphyia), 249
edusa (Colias), Ixix
,, ab. helice (Colias), Ixix
effluata (Orthonama), 262
ega (Morpho), lii
egialea (Amauris), 499
( cxx )
eion (Flavinia), 230
electa (Goiiolabis), 169, 170, 171, 173,
174
electrica (Placoptila), 454
elegans (Tisis), 437
Elis, 401, 403
Elyinnia, xxix
Einaturga, xxxvi
emberizata (Cidaria), 245
,, (Hammapteia), 245, 249
Emmelesia, 270
Emmiltis, 226
emolus (Lycaenesthes), 1, 4, 14, 15, 30,
32
enana (Boarmia), 337
,, (Physocleora), 337
Eiiicmus, lix
ennomaria (Gynopteryx), 302, 303, 304,
305
Ennominae, 280
Ennonios, 205, 305
enotata (Macaria), 293
Entephria, 248
Eois, 217
Epactiothyniius, 415, 416
Epeira, 152
ephialtes (Antlirocera), Ivi
Ephyra, 215
epicurta (Cacoecia), 432
Epimactis, 458
Epiniecis, 329
Epione, Ixiv, 308
Epiphractis, 373
Epirrhoe, 248, 249
Erebia, xxviii
Eretmocera, 454, 455
ergaiie (Pieris), xlv, xlvi
Ergavia, 210
Erilophodes, 325, 326
erina (Achlora), 211
,, (Miantonota), 211
,, ab. disjuncta (Miantonota), 211
Erinnys, xlvi
Eriogaster, xxxviii
Eriopygidia, 272
Erirrhinus, Ixvi
Eristalis, 153
eros (Polyoniinatus), xlv
Erosina, 308, 309, 310
erosinata (Tetracis), 306
erschoffi (Plebeius), 494
erythrogaster (Crabro), 429
Erythrolophus, 221
erytliropoecila (Lycaenestlies), 57
erythropoecilus (Lycaenesthes), 20, 57,
58, 59
esau (Arixenia), ii
escharota (Argyroploce), 436
escheri (Polyommatus), xlvi
Esplialmeninae, 161, 163, 169, 202
Espbalmenus, 163, 170
eterononia (Anisolabis), 175, 177
Ethniia, 460, 461, 462
Ethniiadae, 460
Euacidalia, 220
Eubolia, 258
Euborellia, 168, 175, 176, 179, 180,
181
Eucestiinae, 231, 238
Euchloe, liv
euchroma (Cei’ceris), 421, 424
Eucosmia, 244, 266
Eucosinidae, 368, 434
Eucymatoge, 274
eudora (Microdontopera), 322
,, (Salpis), 320, 322, 323
Eudule, 231, 232, 233
Eudulinae, 231
Eudulophasia, 233
Eueides, Ixviii
Euephyra, 215
Eueupithecia, 274, 275
Eulepidotus, 286
euinedon (Aricia), xlvi
Euparatettix, 359, 360
Eupbaedra, xxix
Euplianessa, 231
eupheno (Euchloe), liv
Euphyia, 248, 249, 250, 251, 252, 260
Eupileta, 280, 284
Eupithecia, 205, 274, 275, 276, 277,
278, 279
Euprepia, 152
Euralia, xiv, xv, xvi, xvii, xli, 498,
499, 500, 501, 502, 503, 504, 505
506, 507, 508, 509, 510, 511, 512,
513
euria (Camptogramma), 260
enrylampis (Tisis), 438
eurytus (Pseudacraea), 498
Eustroma, 270
Evaesthetus, 384
Everes, xlvi, 494
exacta (Calocalpe), 266
exagitata (Camptogramma), 261
exangulata (Hymenomima), 339
exanthista (Tortyra), 464
excavaria (Anisodes), 215
excavatum (Acmophylluin), 355
excavatus (Zaspilothynnus), 420
exhonorata (Sabulodes), 307, 308
exiguata (Eupithecia), 278, 279
expolita (Chrismopteryx), 252, 253
expositata (Tanagra), 345
exsecta (Formica), xliv, 146, 147
exsulatum (Labiduroinma), 188
(
extensuin (Xerophyllum), 350, 351
extersaria (Hymenomima), 338
extrema (Tapinostola), Ixxii
fagi (Stanropus), 152
famulus (Deuterocopus), 112
fasciata (Lycaenesthes), 81
fasciatipennis (Discolia), 403
fasciatus (Lycaenesthes), 81
,, (Triclema), 72, 81, 82
fasciolata (Perizoma), 268
fausta (Lecithocera), 449
felderi (Azelina), 320
,, (Salpis), 320
felis (Ctenocephalus), Ixvi
felix (Anisolahis), 178
femoralis (Psalis), 182
fernandezi (Ctenisolabis), 197, 198
ferrugata (Ptychopoda), 217
,, (Xanthorhoe), 269
, , ab. unidentaria (Xanthorhoe),
268
ferruginata (Eois), 217
ferrugineipes (Dielis), 404, 405
,, (Scolia), 404, 405
festae (Anisolahis), 175, 177
festiva (Psalis), 181, 182
Fidonia, 300
Fidoniinae, 280
figinii (Paralabidura), 185
filamentaria (Nematocampa), 288
filipendulae (Anthrocera), Ivii, Iviii
,, var. chrysanthemi (Anthro¬
cera), xiii
,, ab. flava( Anthrocera), xiii
fimbriata (Calocalpe), 266
,, (Metachanda), 369
flammea (Meliana). Ixxiii
Flavinia, 229, 230
flavipes (Stenus), 382, 384
flavisquamata (Eriopygidia), 272
,, (Spargania), 272
flavomaculata (Lycaenesthes), 53
flavomaculatus (Lycaenesthes), 19, 53,
65
flavopicta (Anthobosca), 399
flavus (Lasius), Ixxii, 145, 146
fletcheri (Ctenisolabis), 197
flexicosta (Metasiopsis), 219, 221
,, (Ptychopoda), 219, 220, 221
flexifascia (Hamalia), 222, 223
floccosaria (Pterocypha), 264
,, (Scotosia), 264
floridensis (Sphacelodes), 301
fluvialis (Cerceris), 421
fluviata (Geometra), 261, 263
,, (Percnoptilota), 261
foedata (Camptogramma), 259
forbesi (Echinosoma), 164
cxxi )
Forcipula, 184, 186, 187
Forficula, 163, 179
Forticularia, 202
Forficulidae, 202
Formasimus, xxxvi
Formica, xiii, xliv, xlv, xlvii, Ixxii
Formicomus, xx
formosa (Dielis), 403, 404
,, vai-. maculiceps (Dielis), 403
,, (Scelolophia), 226
forticornis (Ilyobates), 381
fossulatus (Prototettix), 362
foveolatus (Claviger), 145
fragilis (Tephrinopsis), 295, 296
fragosa (Oinophila), 376
freygessneri (Tachytes), 521, 522, 523
fringillata (Cidaria), 245
,, (Hammaptera), 245
Frisilia, 446
frondaria (Aplodes), 210, 211
,, (Synchlora), 210, 211
fugax (Solenopsis), 146
fuliginosus (Lasius), Ixvii
,, (Quedius), 384
fulva (Tapinostola), xxi
fulvago (Xanthia), Ivii
fulvicornis (Tiphia), 396
fulvilinea (Drepanodes), 302
fulvimacula (Lycaenesthes), 68
fulvitincta (Cleora), 340
,, (Iridopsis), 340
,, (Melinoides), 288
fumida (Hammaptera), 246
,, (Nephodia), 290
,, (Nipteria), 290
,, (Upenora), 290
fumigrisea (Cymatophora), 331
fumipennis (Astatus), 519
furcata (Mesovelia), xliii
furciferata (Larentia), 245
furva (Calocalpe), 265
fur^dpennis (Eupithecia), 276
,, (Tephroclystia), 276
fusca (Azelina), 313
,, (Formica), xliv, xlv, 144, 146,
147, 148, 149, 150
,, (Pero), 313
,, (Psalis, 181
,, (Stenalcidia), 333, 336
fuscata (Narragodes), 281
,, (Pyragra), 165, 166, 167
fuscovariata (Disclisioprocta), 259
,, (Gnophos), 259
fuscum (Echinosoma), 164
,, (Olophrum), xxxi
,, (Xerophyllum), 351
fusilinea (Apicea), 305
fusilineata (Brotis), 301
( cxxii )
gal)raria (Azelina), 307
,, (Geitonia), 307
,, (Xylolocha), 307
gagatina (Psalis), 182, 183
gaika (Zizeeria), 481, 482
,, (Zizera), 479, 482
„ (Zizula), 483, 491, 492, 493, 495,
496, 497
galatea (Melanargia), xlvii
galeatuin (Xerophyllum), 351
Gaiioris, 152
gariazzi (Forcipula), 187
Gastrosericus, 529
gaudens (Anisolabis), 176
Geitonia, 307
Gelasma, 210
Gelechiadae, 369, 437
geminus (CyrtoiThinus), xliii
gemniata (Geometra), 261
,, (Ptychopoda), 217
gemmifera (Neurellipes), 67, 70
geniculata (Brachylabis), 195
,, (Chelidura), 194
genuini (Ichneumones), 389
genurnia (Hammaptera), 244
Geometra, 211, 261, 263, 271
Geoinetridae, xxx, 204, 205, 207, 255,
265, 268, 320, 344
Geometrinae, 210
germanica (Cicindela), xxx
giacomellii (Litliostege), 234, 235
,, ab, angulata (Lithostege),
234, 236
gigantata (Macaria), 292
giganteiis (Goliathus), xxxvi
gilesi (Cercei'is), 422
gilvago (Xanthia), Ivii
glabrifrons (Trypophyllum), 348
gladiaria (Gynopteryx), 302, 303, 304,
305, 306, 311
Glaucopteryx, 254, 268
glauculata (Amaurinia), 279
,, (Thalassodes), 279, 280
globosus (Dyschirius), 384
glyphicarius (Aspilates), 327
Glyphipterygidae, 374, 462
Glyphipteryx, 374, 470
gnamptographa (Melaiiolopliia), 341
gnoma (Anoinpbax), 212
,, (Omphax), 212
Gnophos, 205, 208, 259, 330, 331
Gnypeta, 385
Godana, 432
Goliathus, xxxvi
Gonaepa, 442
Gonepteryx, lii
Gonolabidae, 169
Gonolabina, 161
Gonolabis, 163, 168, 169, 170,171, 172
173, 174, 175
gonopteraria (Azelina), 313
„ (Pero), 313
Gonopteryx, Ixix
gordius (Loweia), xlvi
Gracilariadae, 373
gracililinea (Geitonia), 307
gracilipes (Austrostylops), xliii, 515
gracilitarsis (Tachysphex), 527
giamraica (Lycaenesthes), 83
granillosa (Cymatophora), 341
,, (Melanolophia), 341
Graphidipus, 239
Graphipidus, 239
greeiii (Euborellia), 180
grisea (Stenalcidia), 334, 335, 338
griseata (Azelina), 318
,, (Cnephora), 318
grosei (Lycaenesthes), 40
grossulariata (Abraxas), Ixviii, 152, 153
,, ab. varleyata (Abraxas),
Ixviii
grumata (Hammaptera), 245
guanicola (Thomosis), lix
gueneei (Luperina), Ixii, Ixiii
guttata (Echinopsalis), 167
,, (Psalis), 163
„ (Stenus), 382, 383
guttularis (Carphotricha), Ixxi
gyda (Nanagodes), 207, 281
Gynopteryx, 302, 303, 304, 305, 306,
311
Gypsochroa, 259
gyritis (Paramorpha), 431
hades (Triclema), 71, 74, 76
Haemalea, 222
Haemonia, xliv
haesitata (Gypsochroa), 259
,, (Scotosia), 259
Halictophagidae, 515
Halictus, 419
halisma (Hammaptera), 248
halixanta (Argyroploce), 435
Hamalia, 220, 222, 223, 224, 225
Hammaptera, 243, 244, 245, 246, 247,
248, 249, 269
Hanna, xxix
Harpaetopus, 518
Hasodima, 325
Hastula, xxv, xxvi
hecate (Amauris), 499
,, (Brenthis), xiii
helcita (Aletis), xxix
Helcon, liii
heliarcha (Gonaepa), 442
heliaula (Atteria), 433
Heliconidae, Ixviii
( cxxiii )
helioclina (Tisis), 437
heliodoxa (Atteva), 471
hellmaniii (Tapinostola), xxi, Ixxii
helopioides (Oodes), 381
hemiacma (Lecithocera), 448
heiiiicausta (Thriambeutis), 470
heinichlorata (Aspilates), 273
Hemimei'us, ii
Hemitheidae, 210
Hemitheinae, 210, 213
heniixysta (Tisis), 438
Heodes, xlvi, 3
Herbita, 309, 310
hermanui (Agriom3da), 408
hero (Coenouympha), xxviii
Hesperia, xlv, xlvi
hesperia (Nyctemera), xxix
hesperina (Eudule), 231
Heterolocha, 288, 289
heterolocha (Hyperythra), 302, 304,
305
Heterothops, 384
Heterusia, 233, 234
Heterusiinae, 231
hewitsoni (Lycaenesthes), 40
Hexadactilia, 107, 108
Hibernia, 205
Hieroxestis, 375
himalayensis (Tiphia), 396, 397
hippocrepidis (Anthrocera), Ivii, Iviii
,, (Zygaena), xiii
Hippodes, 346, 347
hirtaria (Biston), 152
histricus (Paratettix), 360
histrio (Enicmus), lix
hobleyi (Lycaenesthes), 16, 24, 25
,, (Pseudacraea), xxix
hoedularia (Azelina), 312
„ (Pero), 312
holcias (Lycaenesthes), 83
holdansi (Nannisolabis), 194, 195
Holoparamecus, xxi
Homalotae, 384
Homaloxestis, 446
Homona, 432
Honorana, 319, 321
Hoplolygris, 207, 254, 255
Horisme, 262
horridum (Echinosoma), 164
horsfieldi (Allotinus), xxxviii, xxxix,
xl, xli
howardi (Leptisolabis), 199
hnnaboldti (Dielis), 404
,, (Scolia), 404
humeralis (Luciola), 532
hyacinthina (Tisis), 439
hyberniaria (Tephrosia), 332
yberniata (Erosina), 309
Hydata, 213
Hydriomena, 263
Hydrionieninae, 231
Hydroecia, xxiii, xxiv, xxv
liyerana (Hastnla), xxv, xxvi
hylas (Polyonimatus), xlvi
Hylograptis, 450, 451
Hymenomima, 334, 338, 339
Hymen opus, 1, li
Hypera, 385
Hyperetis, 288
Hyperythra, 302, 304, 305
Hypochroma, 257
Hypolepis, 271, 272
Hypolimnas, xvi, 498
Hypsaeus, 346, 347, 349, 350
Hypsidae, xxix
ianthes (Ypsolophus), 452
icarus (Lycaena), 487
,, (Polyommatus), xlvi, xlviii,
Ixviii, Ixix
Ichneumones, 389
Ichneumonidae, 386, 389
Ichneumonides (Methoca), xxx
Idialcis, 208, 284, 285, 286
Idioglossa, 373
iduna (Perizoma), 270
ilicis (Nordmannia), xlvi
illineata (Stenalcidia), 334
Ilyobates, 381
imitata (Thamnonoma), 292
Imma, 464, 465, 466, 467
immanata (Dysstroma), 268
,, (Phalaena), 271
iminissus (Biston), 343
imparata (Acidalia), 215
impauperata (Scotosia), 259
imperatrix (Tisis), 440
1 impressa (Crepidodera), lx
,, (Luciola), 532
inipi’essus (Prototettix), 361
iinpromissata (Perizoma), 254, 267, 268
,, (Ypsipetes), 267
inaequalis (Hypsaeus), 347
inca (Gonolabis), 163, 170
incandescens (Ptilosticha), 441
incerta (Anisolabis), 175, 177
incolorata (Phrygionis), 286
incurvata (Stenodonta), 318
indefinita (Eupithecia), 276
,, (Lycaenesthes), 16, 25
indica (Zizera), 479, 481, 482, 484, 486,
488, 489
„ (Zizina), 483, 488, 489, 490, 491,
492, 493, 495, 496, 497
indicum (Allostethus), 163
ineffectaria (Tephrosia), 332
inermis (Labidura), 185
cxxiv )
inexcisa (Semiothisa), 294
inexpectata (Cerceris), 421
infelix (Anisolabis), 178
,, (Sabulodes), 320; 321
,, (Salpis), 320, 321
infuscata (Coptotettix), 365
insifinis (Prosopigastra), 528
insolita (Gnophos), 208, 331
,, (Scotopterix), 331
iiisulanuin (Echiiiosoma), 164
insularia (Acidalia), 215, 216
,, (Deptalia), 215
,, (Pleuropruuha), 215
,, (Ptychopoda), 216
iiifeularis (Liacos), 400
,, (Scolia), 400
iiitaminata (Piagetia), 426
intercalata (Cidaria), 245
intevguttella (Coydalla), 449
,, (Onebala), 449
interlineata (Cymopsis), 344
,, (Devarodes), 344
internexata (Craspedia), 227
interrupta (Plesia), 392
interruptata (Cidaria), 262
iiiteiTuptus (Euparatettix), 359
iiitestinata (Phibalaptertyx), 259
,, (Pliibalapteryx), 259
intrudens (Syagrius), Ixxi
intrnsata (Phibalapteryx), 261
invaria (Ameria), 233
,, (Eudule), 233
,, (Eudulopliasia), 233
invenusta (Stenalcidia), 334
iostrota (Limnoecia), 454
Ira, 310
Iridopsis, 340, 341
Isanthrene, Iv
Ischnopteryx, 327, 328
isis (Atyria), 230
isochrysa (Anaphantis), 472
I.solabidae, 189, 190, 202
Isolabis, 190, 191, 195, 196
Issoria, xlvi
ituria (Lycaenesthes), 16, 22
iveni (Catascia), 330
ja (Lycaeuestlies), 20, 58
jaciiitaria (Pachrophylla), 241
jacintha (Idialcis), 285, 286
jacintharia (Cidaria), 244
jacksoni (Deuterocopus), 124, 129
jacobsoni (Forcipula), 187
jamaicensis (Pero), 313, 314
Jamides, 1
janeirensis (Euborellia), 180
japonica (Zizera), 486
jatropharia (Cyllopoda), 230
javana (Anisolabis), 174
iavana (Gonolabis), 169, 170, 171, 172,
173, 174
jermyni (Parhestina), Ixxiii
johnstoni (Acraea), xlii
jouesaria (Azelina), 313
,, (Pero), 313
juba (Lycaenesthes), 20, 63, 64
jiigurtharia (Cidaria), 244
,, (Hammaptera), 244
junctilinea (Alcis), 335
,, (Stenalcidia), 335
juno (Stenus), 382, 384
kamilila (Triclema), 71, 76, 78
kampala (Lycaenesthes), 19, 52
karsandra (Zizeeria), 480, 481, 482,
484, 485, 491, 495, 496,
497
,, (Zizera), 479, 480, 481, 482
karschi (Piezotettix), 347
Karschiella, 189
kersteni (Lycaenesthes), 44
kirbyi (Anisolabis), 171
,, (Euralia), xvi, xvii
,, (Gonolabis), 169, 170, 171
,, (Salius), Iv
kiiviiyu (Lycaenesthes), 17, 37
Klugia, xlvi
knysna (Zizera), 486
korlana (Everes), 494
kudagae (Anisolabis), 178
kurandae (Nitela), 428, 429
kurandensis (Asthenothynnus), 411
labecnlata (Cerceris), 421
Labidura, 175, 183, 184, 185, 187
Labiduridae, xi, 161
Labidurinae, 162, 165, 183
Labidurodes, 168
Labiduromma, 188
Labiidae, Ixv
labilis (Gnypeta), 385
labradus, var. otis (Zizeeria), 481
,, (Zizera), 482, 484, 486, 489
„ (Zizina), 483, 489, 490, 491,
492, 495, 497
,, var. dryina (Zizina), 489,
490, 491, 492, 495
,, var. otis (Zizina), 489, 495
,, var. sangra (Zizina), 489,
495, 496, 497
Laccobius, lix
lachares (Lycaenesthes), 3, 18, 19, 47,
48, 50, 51. 53, 66
,, var. obsolescens (Lycaenes¬
thes), 48
lachrymosus (Aeolothynnus), 416
lacides (Lycaenesthes), 73
,, (Triclema), 71, 73
lactevirens (Eupithecia), 276
( cxxv )
lactevirens (Tephroclystia), 276
lacunata (Pseudasellodes), 213
lades (Macrocneme), liv, Iv
laeta (Anisolabis), 177
laetus (Astatus), 520
laevipennis (Apicia), 306
laevis (Narragodes), 280
laica (Cryptolechia), 456
1-album (Leucania), Ixii
lamias (Lycaenestlies), 79
,, (Triclema), 71, 72, 79
lamprocles (Lycaenestbes), 65
,, (Neurypexina), 49, 65
Lampyris, xxxvi
lanestris (Eriogaster), xxxviii
lapidata (Coenocalpe), 262
lapillata (Camptogramma), 261
laqueata (Taragmarcha), 370
Larentia, 244, 245, 252, 254, 259, 260,
265, 266, 273
Lareutiinae, 231, 235, 259, 269
Lariada, 425
larydas (Lycaenestlies), 3, 10, 15, 18,
43, 44, 45, 46, 50
,, (Pseudodypsas), 44
Lasiocampidae, Ixix
Lasiops, 319, 320, 321
Lasius, lx, Ixvii, Ixviii, Ixxii, 144, 145,
146, 147
lasti (Lycaenestlies), 17, 22, 29, 84
lateraria (Coremia), 245
laternaria (Asellodes), 213
latlionia (Issoria), xlvi
Lathrobium, lix, Ixvi
latifasciella (Tisis), 437
latioiata (Amelia), 233
Latiorina, xlviii, Ixix
latirupta (Hydriomena), 263
,, (Phibalapteryx), 262
lativalvis (Tachysphex), 524
lativentris (Gonolabis), 169
lavaterae (Erinnys), xlvi
Lecithocera, 447, 448, 449
Lembeja, lii
lembula (Meridarcbis), 430
lemnos (Lycaenestbes), 17, 30, 47, 83
leonina (Cupidestbes), 5, 6, 12
,, (Lycaenestbes), 12, 13
Lepidoptera, lii
leptines (Lycaenestbes), 20, 56, 57, 58,
64
Leptisolabis, 190, 191, 194, 195, 196,
198, 199, 200, 201
Leptodeuterocopus, 107, 138, 141 •
Leptomeris, 217, 220, 222, 226
Leptostales, 219
Leucania, Ixii
Leucocbesias, 208, 326
leucoclora (Lecitbocera), 448
Leucolitbodes, 325, 326
leucostigma (Macrocneme), liv
leukokescba (Lycaenestbes), 51
levis (Lycaenestbes), 18, 42
Liacos, 400
libanota (Tortyra), 463
librata (Parelliptis), 445
Libytbea, lx
ligea (Erebia), xxviii
lignata (Ortbonama), 262
,, (Pergama), 318
,, ab. obscurior (Pergama), 318
,, (Pero), 317, 318
ligulifera (Gynopteryx), 302, 304, 305
ligures (Lycaenestbes), 3, 16, 23, 25,
28, 83
limbata (Eudule), 232
,, (Eupbyia), 251
,, (Mennis), 232
,, (Nematocampa), 288
Limenitis, xlvi
liinitata (Epirrboe), 248
,, (Ptycbopoda), 217
Limnoecia, 453, 454
limosa (Melanolopbia), 342
,, (Tepbrosia), 342
lindenii (Dielis), 404
linearis (Xylolocba), 307
Linoclostis, 458
bodes (Lycaenestbes), 17, 32, 34, 38
liparis (Lycaenestbes), 47, 48
Lipomelia, 222
Liris, 425
Lissocblora, 211
Lissopsis, 239
Lissotes, liii
litbas (Cupidestbes), 6, 11
,, (Lycaenestbes), 11, 40
Litbina, 292
Litbostege, 208, 234, 235, 236, 237,
238
littorea (Anisolabis), 178
litura (Rbizobius), 385
liturata (Macaria), 293
livida (Lycaenestbes), 17, 34
lividipes (Paralabidura), 184, 185
lobifonnis (Eudule), 232
lobulatus (Prototettix), 362
locbias (Lycaenestbes), 31
locra (Lycaenestbes), 43
locuples (Lycaenestlies), 19, 52
lombokianum (Allostethus), 162
Lomecbusa, Ixxii, 144
Lomograpba, 290
longicollis (Cionus), lix
longicornis (Claviger), lx
„ (Tbynnus), 411
Y cxxvi )
ongicovnis (Zeleboria), 411
longipedata (Ptycliopoda), 218, 220
longipemie (Lathrobium), lix, Ixvi
longipeniiis (Stenorrboe), 274
longitarsis (Chilopora), 384
loiiicerae (Zygaena), xiii
lophopteryx (Deuterocopus), 112, 118,
121
lorquirjii (Zizera), 479
loti (Diplura), xxxviii
,, (Osmia), 155, 156, 157
Loweia, xlvi
Lozograiuma, 292
Lucanus, 153
lucens (Hydioecia), xxiv, xxv
lucida (Actizera),483, 494, 495, 496, 497
,, (Antizera), 481, 494
Luciola, Ixxiii, 532, 533
lucretia (Lycaenesthes), 80, 81
lucretilis (Lycaenesthes), 80
,, (Triclema), 2, 71, 72, 80, 81
luctuosus (Agriomyia), 409
lugeus (Sphex), 519
lulu (Zizera), 495
luuulata (Lycaenesthes), 11, 17, 18,
32, 40, 42
,, (Neolycaena), xxiii
Luperina, Ixii, Ixiii
lurida (Forcipula), 186
luscinata (Cidaria), 262
lusones (Lycaenesthes), 68
,, (Neurellipes), 66, 67, 68, 69
lustrator (Stenus), 383, 384
luteago, var. barrettii (Dianthoecia),
Ixxii
luteela (Tisis), 437
Lycaena, xlvi, xlviii, Ixviii, Ixix, 14,
495
Lycaenesthes, 1, 2, 3, 4, 5, 7, 8, 10,
11, 12, 13, 14, 15, 16, 17, 18, 19,
20, 21, 22, 23. 24, 25, 26, 27, 28,
29, 30, 31, 32, 33, 34, 35, 36, 37,
38, 39, 40, 41, 42, 43, 44, 45, 46,
47, 48, 49, 50, 51, 52, 53, 54, 55,
56, 57, 58, 59, 60, 61, 62, 63, 64,
65, 66, 67, 68, 69, 70, 71, 73, 74,
75, 78, 79, 80, 81, 82, 83, 84
Lycaenidae, xxiii, xxxviii
lycaenina (Lycaenesthes), 4
Lycaenopsis, Ixxiii, 480, 490
lychnaptes (Lycaenesthes), 20, 61, 62
lychnides (Lycaenesthes), 19, 53
lychnoptera (Lycaenesthes), 62
lycoa (Acraea), xlii
lycotas (Lycaenesthes), 51
Lygris, 255, 270
Lymantriadae, xxix
Lyouetia, 374
lyrifera (Iinma), 466
lysicles (Lycaenesthes), 3, 18, 46
lysigania (Tanychastis), 371
lysimon (Zizeeria), 480, 481, 482, 483,
484, 485, 486, 491, 495, 496,
497
,, var. knysna (Zizeeria), 496
,, (Zizera), 479
ly'zanius (Lycaenesthes), 2, 64, 65
,, (Neurypexina), 54, 64, 65, 66
inaartensi (Allostethus), 163
Macaria, 292, 293, 294
macra (Periacina), 455
macrochalca (Acrocercops), 373
Macrocneme, liv, Iv
Macrolyrcea, 319
Macronychus, lix
maculata, var. variegata (Agidoniyia),
410
maculicornis (Tachytes), 523
madagascariensis (Euralia), 509
maeander (Lycaenesthes), 69
,, (Neurellipes), 66, 69
inaera (Pararge), xlvi
,, f. adrasta (Pararge), xlvi
maha (Zizeeria), 480, 481, 482, 484,
485, 486, 487, 488, 490, 491, 492,
495, 496
inahallokoaena (Lycaena), 495
inahota (Lycaenesthes), 20, 59
major (Anisolabis), 176
,, (Platylabia), 162
inakala (Lycaenesthes), 20, 58
malacodes (Nothris), 451
maldama (Apicia), 302, 303, 304, 305
maleformata (Sebastia), 275
malgacha (Metisolabis), 196
inalvata (Larentia), 260
Maniestra, 152
manicata (Osmia), 155, 158, 159
manifesta (Agriomyia), 409
manni (Pieris), xlv
,, ab. erganoides (Pieris), xlv
,, var, rossii (Pieris), xlv, xlvi
mansueta (Scolia), 402
Mantidae, li
marginalis (Anisolabis), 176, 177
mai'ginata (Nii)teria), 290
,, (Zizej'a), 486
maritirna (Anisolabis), 178
Marmarea, 312
Marragodes, 281
marshalli (Dielis), 402
• ,, (Lycaenesthes), 79
,, (Scolia), 402
, ,, (Triclema), 72, 79
martena (Physocleora), 334
martharia (Acidalia), 227
( cxxvii )
Mascora, 326
mathewi (Deuterocopus), 124
mathilda (Azelina), 314
„ (Pero), 314
raathina (Tricliura), Iv
mauritanica (Anisolabis), 177
maurorufus (Quedius), 384
maxima (Anisolabis), 176, 178
Jlecoceratinae, 209
mediata (Phibalapteryx), 262, 263
Jledinopliyllum, 346, 349, 355
medon (Aricia), xlvi
medusa, var. hippomedusa (Erebia),
xxviii
megalorma (Cryptnpbasa), 457
Meganthopus, liv
meisteri (Rhyucliopyga), Iv
melambrota (Lycaenesthes), 50
melambrotus (Lycaenesthes), 19, 50,
78
uielanardis (Frisilia), 446
ilelanargia, xlvii
Melanchroia, 345
Melanippe, 247
melanocephalum (Pison), 419
Melanolophia, 333, 341, 342
Melanopbila, xliii
melanota (Deuterocopus), 112, 113
melas (Isanthrene), Iv
meleager (Polyommatus), xlvi
Meliana, Ixxiii
Melinoides, 207, 288, 289
meliorella (Tisis), 437
l\lelitaea, xxviii, xlvi, liii, liv, Ixiv,
Ixxiii
Melodryas, 472
menciana (Homona), 412
,, (Pandemis), 432
Mengeoidea, 515
Mennis, 232
Meridarchis, 430
meridionalis (Eubolia), 258
Mermis, Ixxii
meropia (Erosina), 308
Jlesa, 394
mesargyrata (Leucochesias), 208, 326
meskearia (Stenaspilates), 312
mesoscia (Tephrinopsis), 298
Mesovelia, xliii
messapus (Lycaena), 495
Metachanda, 369
Metacliandidae, 369
metagrammata (Seotosia), 265
Metasiopsis, 219, 220, 221
Methoca, xxx
Metisolabis, 191, 195, 196, 198, 200
metria (Idialcis), 284
Metrocaiiipa, 288
Miantouota, 211
micacearia (Sphacelodes), 301
michaelseni (Gonolabis), 169, 170, 174
,, var. dentata (Gonolabis),
174
Microdontopera, 320, 321, 324
Microgoiiia, 307, 310, 311
Micro -Lepidoptera, 366
Microxydia, 289
millari (Lycaenesthes), 17, 35
miltocosma (Polemograptis), 432
mima (Euralia), xiv, xv, xvi, 498, 499,
500, 502, 503, 504, 505, 506, 507,
508, 509, 510, 511, 512, 513
raimetica (Cupidesthes), 7, 13
minietis (Cosmojderyx), 372
Mimophyle, 208, 283, 284, 286
mimulus (Sphex), 419
minima (Anthobosca), 398
,, (Lycaenesthes), 17, 36
minimus (Coenonympha), 480, 496
,, (Cupido), xlvi
,, (Zizera), 480
I minor (Astatus\ 521
,, (Pyragra), 166
minorata (Acidalia), 229
minuata (Aplodes), 211
,, (Thalera), 210
minuscula (Cerceris), 422
minuta (Anisolabis), 180
minutissimus (Asthenothynnus), 413
minutus (Coptotettix), 363
misippus (Diadema), 498
,, (Hypolimnas), xvi
mixtus (Ceuthorrhynchideus), xxxv
mnemosyne, var. frlihstoferi (Pariias-
sius), Ixix
Mnesteria, 438
moesta (Euborellia), 181
molaria (Odysia), 332
mblleri (Parasapyga), 405
mollis (Calathus), xiii
mollita (Byssodes), 287
monastica (Imma), 465
moncus (Lycaenesthes), 43
Monoctenia, 320
monomatapa (Tiphia), 396
Monopis, 473, 474
monoplanetis (Atteva), 470
montana (Ctenisolabis), 197
monteironis (Lycaenesthes), 33
morawitzi (Osmia), 156, 157
raoricei (Gastrosericus), 529
mormopa (Imma), 467
Morpho, lii
mortuaria (Plemyria), 261
multiguttata (Cerceris), 421
multiplagiata (Auophylla), 211
( cxxviii )
munda (Gelasnia), 210
raundaria (Bronchelia), 343
mundata (Microgonia), 307, 311
,, ab. crassior (Microgonia), 311
,, (Oxydia), 311
nmsagetes (Lycaenestlies), 3, 16, 21, 23
miiscilineata (Ptychopoda), 218
muscosaria (Rhopalodes), 240
innscosata (Perizoma), 268, 269
inusciilosa (Oria), Ixiii
,, (Synia), Ixiii
musicodes (Linoclostis), 458
niutabilis (Cidaria), 250
,, (Ypsipetes), 251
M3"riapoda, 191
Myrmedonia, 382, 385
Jlyrmica, 146
inyrtillata (Catascia), 330
Myzine, 391, 392, 393
Myzininae, 391
Nacaduba, 1
Nannisolabis, 190, 191, 194, 195, 200
napi (Ganoris), 152
,, var. napaeae (Pieris), xlvi
Narragodes, 207, 208, 280, 281, 282,
283, 284
natalensis (Lycaenestlies), 30
,, (Tiphia), 396
nazadaria (Apicia), 302, 303, 304
,, ab. inapicata (Gynopteryx),
303
neanthes (Charaxes), xv, 506
neavei (Xerophyllum), 351, 352, 353
neglecta (Lycaenestlies), 31
Nematocainpa, 287, 288
Neochesias, 236, 237, 238
Neolycaena, xxiii, 88
neoniger (Lasius), Ixxii
Neopithecops, 495
Neotaxia, 286
nepalensis (Paralabidura), 185
nephalia (Boarinia), 338
,, (Hymenomima), 334, 338
Nepliodia, 290
Nephodiinae, 280, 290
Nepliogenes, 455
Neurellipes, 66, 67, 68, 69, 70, 82
Neurypexina, 49, 54, 64, 65, 66
niavius (Amauris), 499
,, sub-sp. dominicanus (Amauris),
xiv, 499, 506, 510
nicbolsoni (Oloplirum), xxxi
nictitans (Hydroecia), xxiii, xxiv
niger (Lasius), Ixvii, Ixviii, Ixxii, 144,
146, 147
,, var. americanus (Lasius), Ixviii
nigeriae (Lycaenestlies), 78
,, (Triclema), 72, 78
nigra (Bracliylabis), 199, 201
,, (Ctenisolabis), 197, 198
,, (Heterotliops), 384
,, (Metisolabis), 200
,, (Polybia), liv
,, (Psalis), 182, 183
nigrescens (Pliysocleora), 337
nigricans (Nitela), 428
nigricosta (Tliysanopyga), 301
nigriplena (Bronchelia), 343
nigrirostris (Hypera), 385
nigrita (Myzine), 391
nigrocaudata (Lycaenestlies), 26
nigrociliata (Hamniaptera), 247
nigrolineata (Hamniaptera), 246
nigropuiictata (Lycaenesthes), 17, 36
nigro-punctatum (Acmopliylluni), 355
nimbata (Microgonia), 310
,, (Oxydia), 310
niobe (Argyiinis), xlvi
Nipteria, 290, 291
nisa (Liponielia), 222
Nitela, 428, 429
nitida (Bracliystichia), 303
,, (Gynopteryx), 303
nitidus (Tacliyspliex), 524
nivalis (Allotinus), xxxviii, xxxix, xl,
xli
niveata (Perizoma), 269
niveigutta (Hamniaptera), 245
niveipennaria (Aspilates), 208, 326
noctiluca (Lampyris), xxxvi
Noctua, Ixiv
noctuata (Mimopliyle), 283, 284
nodosa (Lecithocera), 447
Noniiades, lx
Nonagria, xlvii
Nordmannia, xlvi
norma (Azeliiia), 316
,, (Pero), 313, 316, 317
nossima (Amauris), 499
notata (Macaria), 293
notaturia (Honorana), 319
Notliris, 451
novae zeelandiae (Parisolabis), 188
noverca (Pseudospliex), liv
nubigena (Imma), 466
Niimia, 289
nupta (Catocala), lii
Nyctemera, xxix
Nymplialidae, xxix
obfuscaria (Catascia), 330
Obila, 264
obliquaria (Anteois), 220
,, (Apiciopsis), 306
obliquata (Iridopsis), 340
oblita (Gonolabis), 169, 170, 171, 172,
173, 174
( cxxix )
obruptata (Coremia), ‘Jdl
obscura (Ephyra), 215
,, (Physocleora), 336
,, (Tricleina), 72, 82
obscurus (Pterophorus), 115
obsoletaria (Cidaria), 260
obstipata (Ortlionama), 261, 262
,, (Phalaena), 261
obtiisaria (Hammaptera), 248
occidentale (Echinosoma), 164
occidentalis (Anisolabis), 177
occultaria (Pterocypba), 264
ocellaris (Xaiitliia), Ivii
ocellata (Cyc-loiiiia), 300
ocellatiis (Smerinthus), xxxvii
ocblea (Amauris), xiv, 499, 501
ochribasis (Entepbria), 248
„ (Hammaptera), 248
ochrifascia (Thamnoimma), 299
ochritiucta (Perizoma), 268
ochroleucata (Acidalia), 229
Oehyria, 256, 257, 259, 269
octonias (Lecithocera), 447
oculata (Lobophora), 241
„ (Pat'liropbylla), 208, 241
oculatiis (Lycaenesthes), 78
,, (Triclema), 72, 78
odatis (Cambogia), 280
Odites, 458
Odontomacluis, lix
Odontopera, xxxi
odontota (Baynia), 242, 243
Odontothera, 208, 327
Odontothynnus, 399
Odysia, 332 '
Oecoplioiidae, 373, 455
Oecophylla, 15
Oenoclirominae, 209
oenopodiata (Acidalia), 214
,, (Semaeopus), 214
Oenothalia, 301
Oinophila, 375, 376, 377
olivacea (Pero), 315
olivata (Apicia), 305
Olophrum, xxxi
olympusa (Lycaenesthes), 38
omphacina (Larentia), 273
Omphax, 212
Onebala, 449, 450
onopordi (Hesperia), xlvi
Oodes, 381
opalina (Zizera), 486
operaria (Catascia), 330
Opisthograptis, 292
opistholeuca (Neocliesias), 238
Oporinia, 269
opposita (Cerceris), 420
oppositaria (Boarmia), 343
PROC. EIW. SOC. LOND., V. 1910.
Opticus (Stenus), 383, 384
orbata (Braclimia), 450
orbiculosa (Oxytripia), Ixvi
orbifera (Antygophanes), 324
,, (Salpis), 320, 324
orbitulus (Latiorina), xlviii, Ixix
,, (Lycaena), xlviii, Ixix
oreas (Gnophos), 208, 331
Orgyia, Ixix, 152
Oria, Ixiii
Orneodidae, 111
orsitaria (Microxydia), 289
,, (Stegania), 289
Orthonama, 261, 262, 263, 264
Orthoptcra, xvii, 346
Orthostixinae, 209
osiris (Atyria), 230, 231
,, (Cyllopoda), 230
Osmia, xvii, 155, 156, 157, 158, 159
Osphretica, 475
ossa (Zizeeria), 480, 481, 482, 484, 485,
486, 487, 488, 495, 496, 497
ossium (Stenus), 382
ossularia (Leptomeris), 220
otacilia (Lycaenesthes), 17, 36, 37, 38,
40
,, var, kikuyu (Lycaenesthes), 37
Otis (Zizeeria), 481, 484, 489, 490, 491
otophora (Rhopalodes), 208, 240, 241
Ourapteryginae, 280
Oxydia, 310, 311
Oxypoda, 384
Oxyptilus, 110, 111
Oxytelus, 385
Oxytripia, Ixvi
Pachrophylla, 208, 241
Pachycoj»sis, 211
pacifica (Anisolabis), 175, 176, 179
pales (Brenthis), xxviii
,, var. arsilache (Brenthis), xxviii
Palex, 162
Palicinae, 161, 162
pallescens (Iridopsis), 340
pallidata (Xenoecista), 299
palmirensis (Azelina), 314
,, (Pero), 314
palpina, var. brevipennis (Pterostoma),
lx
paludis (Hydroecia), xxiv
Palyadinae, 207
Pamphila, Ixix
pamphilus (Coenonympha), xlvi
panagaea (Actizera), 483, 494, 495,
496, 497
Pandemis, 432
pandora (Dryas), liv
paiinaria (Scelolophia), 226
panteata (Bryoptcra), 326
I
( cxxx )
jianteata (Leucolithodes), 326
Pantelia, 346, 348
Panthera, 291
pantlierata (Leucolithodes), 326
Pantherodes, 291
panzeri (Tachysphex), 527
,, var. oranieiisi (Tachysphex),
527
Paptilio, XV, xvi, xxix, lii, 508, 510
jiaracyrta (Oditcs), 458
paradoxata (Uyssodes), 286, 287
,, (^Eulepidotus), 286
,, ( Plirygionis), 286
. , incolorata( Phrygioiiis), 207,
287
Paraethiia, Iv
jiaiaguaj'eiisis (Propyragra), 166, 167
,, (Pyragra), 166
Paralabidura, 183, 184, 185, 187
parallelaria (Tephriiiopsis), 297
Paramorpha, 431
paraneiisis (Plemyria), 262
paraptera (Schoeiiotenes), 433
Pararge, xlvi
Parasapiyga, 405
Paraseniia, 293
Paratettix, 360, 361
Paratyria, 230
pai'dalaria (Panthera), 291
., ab. obliterata (Panthera),
291
,, (Pantherodes), 291
Parellipdis, 445
Parhestina, Ixxiii
parinotata (Haminaptera), 246
Parisolabinae, 161, 162, 188
Parisolabis, 188
Parnassius, xxviii, Ixix
jiarthenie (Melitaea), xlvi, Ixxiii
,, ab. rhoio (Melitaea), Ixxiii
parvulum (Echinosonia), 164
jiarvulus (Coptotettix), 365
pauperula (Lycaenesthes), 84
j)avana (Eueides), Ixviii
pax (Lithostege), 208, 235, 236, 237
p)ectinicornis (Bruchus), Ixxii
pectinipes (Tachysphex), 524, 526
pectoralis (Anisolabis), 176
pedestris (Formiconius), xx
,, (Tiphia), 396
Pelopaeus, 390
peltigerata (Macaria), 293
Pelurga, 255
pentacarpa (Oinophila), 376
peracutata (Cidaria), 261
percheron (Psalis), 181
Percnoptilota, 261
peregrina (Anisolabis), 176
I perfiiscimargo (llyiuenomima), 339
Pergama, 312, 315, 318
Periacma, 455
perichlora (Homaloxestis), 446
pericles (Lycaenestlies), 44
periploca (linma), 466
(lerirrorata (Ptychopoda), 219
Perizama, 208, 254, 267, 268, 269, 270
perkinsi (Ani.solabis), 176, 178
,, (Asthenothynnus), 414
,, (Cerceris), 420, 423
Pero, 311, 312, 313, 314, 315, 316, 317,
318
perornata (Phacelophora), 242
persiniilata (Acidalia), 215
perspiciiata (Cidaria), 245
])ersuasoria (Rhyssa), xxxi
Perusia, 289
peruviana (Euborellia), 180
peruviensis (Aniaurinia), 279
petraria (Lithina), 292
Phacelophora, 242
Phaegorista, xxi.x
phaeochalca (Hieroxestis), 375
jihaeoleuca (Athrypsiastis), 458
Phalaena, 261, 271, 345
,, Geoinetra, 343, 345
pharetrata (Mnesteria), 438
Phasis, 3
jdiasmatopa (Stenoina), 460
Phibalaptertyx, 259
Phibalapteryx, 259, 261, 262, 263, 286
pliilea (Catopsilia), lii
Philereme, 259, 260, 265
i philetas (Nanni.solabis), 194, 200
philippi (Tachysphex), 524
phlaeas var. eleus (Rumicia), xlvi
phoebe (Melitaea), xlvi
phoenicis (Lycaenesthes), 75
,, (Triclenia), 71, 75, 79
phorcaria (Scelolophia), 225
phrictodes (Tinissa), 477
Plirygionis, 207, 286, 287
Phymatocera, xxxv, xxxvi
I physalodes (Rhopobota), 368
Physocleora, 334, 336, 337, 338
Piagetia, 407, 426
piagetioides (Tachysiihex), 524
picata (Eiiphyia), 249
picea (Gonolabis), 169, 171, 174
picina (Psalis), 183
pieipes (Stenus), 382
pictus (Dinetus), 530, 531
I ,, (Paratettix), 361
Pieridae, xlv
I Pieris, xlv, xlvi
1 Piezotettix, 346, 347
; Pigia, 220
( cxxxi )
pif^rata (Coreinia), 261
piligera (Anisolabis), 163, 176
piligenim (Allostethus), 163
pilosellus (Ceutlion'liyuchus), xxxv
,, (Sphodrotes), 427
pilosulus (Aehu’us), 407
pil)erata (Ptyeliopoda), 216
Fisoii, 419, 427
pityocaiiipa (Ciiethocampa), xxvii
Placoptila, 453, 454
plagioscia (Planeuia), xxix
plana (Neotaxia), 286
Planema, xxix, 498, 499
planeta (Dentcrocopiis), 112, 131, 132,
133, 134, 138
platensis (Hanialia), 224
platinata (Ilatiaria), 287
,, (Urapteryx), 287
platycorys (Xeiopliyllnm), 351
Platylabia, 162
Platylabiinae, 162
Platyptilia, 110, 367
Plebeius, xlvi, 494
plebeja (Psalis), 182
plebejata (Apicia), 301
,, (Drepanodes), 301
Plemyria, 261, 262
pleniyraria (Acidalia), 220
,, (Ptychopoda), 219, 220
plemyrata (Cidaiia), 261
,, (Orthoiiama), 261
Plesia, 392, 393, 394
Pleui’opmclia, 215
pliiithiota (Stathniopoda), 454
plurilincata (Aspilates), 291
,, (Cataspilates), 291
Plusia, 151
Plutellidae, 470
pluto (Anisolabis), 176
pluvialis (Labidura), 185
jioeyi (Rhopalosoma), 388
poggei (Xanthospilopteiyx), xxix
Polemograptis, 432
Polochrum, 405
polonus (Agriades), viii, ix, xxxvi
Polyliia, liv
I)olydora (Epione), 308
,, (Sabulodes), 308
Polygonia, xliv
Polygraphodes, 226
Polyominatus, xii, xlv, xlvi, xlviii,
lx viii, Ixix
Polyijhasia, 271
Polysemia, 209, 210
Pompilidae, liv
poinpiliformis (Clavelia), xxxv
Pontia, xlvii
pontina (Haminaptera), 247
; pontina (.Melaiuppe), 247
1 populi (Smerinthus), xxxvii
. praeclara (Etlimia), 460, 461
j praedura (Cerceris), 422
i praeinatura (Hieroxestis), 375
Pi'enole})is, Ixxii
- pretiosa (Tortyra), 464
primaeva (Scardia), 478
I princeps (Cainpsoineris), 403
i ,, (Lycaeuestlies), 17, 31, 32, 39,
40
jn'ionograinma (Hypolepis), 271, 272
piivignaria (Bj'ssodes), 286, 287
probolactis (Lyonetia), 374
Procalyptis, 431
processionea (Thauinatopoea), xiv
prodigella (Tortyra), 464
profugaria (Larentia), 273
progressata (Scotosia), 265
Ib'oiuaclius, xxvii
Propyragra, 165, 166, 167
prosecusa (Everes), 494, 495
Prosena, Ixxi
Prosopigastra, 528, 529
I Prosopolophinae, 280
' protona (Anaphantis), 472
Prototettix, 361, 362
prouti (Coenocalpe), 264
,, (Ortbonama), 264
providus (Stenus), 383, 384
Psalinae, 162, 167, 183
Psaliodes, 271, 272
Psalis, 163, 167, 168, 175, 181, 182,
183
Pseudacraea, xxix, 498, 499
Pseudagenia, Iv
pseudaluina (Cataspilates), 291
Pseiidasellodes, 213
Pseudisolabis, 188
Pseudobreplios, 328, 329
Pseudodypsas, 14, 43, 44
Pseudosalpis, 319, 320, 321
Pseudosphex, liv
Psodopsis, 208, 282, 284
psycliidia (Xarragodes), 281, 282
psyttalea (Amauris), 499
Pterocyplia, 264, 265
Pterophoridae, 107, 111, 367
Pterophorus, 110, 115
Pterosticlius, Iviii
Pterostoma, lx
Pterygopterus, Iv
Ptilosticha, 440, 441
Ptinella, xxx
Ptoclioryctis, 458
Ptychopoda, 216, 217, 218, 219, 220,
221, 222, 224
ptychopoda (Hanialia), 223
( cxxxii )
pudibunda (Ortlionaina), 2G4
pudicaria (Bronchelia), 342, 343
l)uechi (Lasiops), 320
,, (Microdontopera), 321
,, (Salpis), 208, 320
piiella (Perizoina), 208
pwellaria (Bronchelia), 342, 343
,, ah. coiifcrtisltriga (Bronehe-
lia), 343
pugiiator (Tachys[iliex), 42r>
pugnax (Forcipula), 186
,, var. niiiior (Forcii)ula), 186
pulcher (Lycaenesthes), 4, 48 •
pullata (Ethmia), 461
pullatana (Tortrix), 432
jnilvillus (Euparatettix), 360
pnnctata (Brachylabis). 108
,, (Lcptisolahis), 190, 20o
punctatissinia (Prosopigastra), 020
punctatus (Lissotes), liii
pnnctilla (Physocleora), 337
,, (Stcnalciilia), 338
punctiini (Elis), 401
,, (Trielis), 401
pimctuosus (Sphodrotes), 428
pustulatuni (Bembidiiiin), xxi
Pygaera, 1.02, 153
pygidialis (Taclij^sphex), 527, 528
Pygidicraninae, 165
pygmaeata (Auteois), 217
])ygmaeus (Asthcnotliynnus), 414
Pyragra, 165, 166, 167
Pyragropsis, 165, 167
Pyragiinae, 161, 163, 165, 183
Pyralidae, 205
Pyraustidae, 366
Pyrinia, 216
pyrochorda (Gonaepa), 442
pyrnptera (Lycaenesthes), 20, 61
Pythagoras (Lycaenesthes), 20, 64
pythonias (Rhopobota), 434
qnadricostaria (Boarniia), 343
,, (Thyrinteina), 343
([uadrifasciata (Straiigalia), liii
([uadrilineata (Hyperetis), 288
,, (Ureihone), 288
(inadrituT)ercuiatus (Macronychus), lix
quadris])inosa (Forcipula), 187
quadruplicaria (Heternsia), 233
,, (Scord)’lia), 233
,, var. latior (Scoidylia),
233
,, var. latissima (Scor-
djdia), 234
Quedius, 384
quelchi (Forcipula), 187
(juerula (Lecithocera), 440
quinquefasciata (Dielis), 402
(juinquefasciata (Scolia), 402
([uisquiliaria (Stcnalcidia), 336
Bacheospila, 212
radialis (Astatus), 520, 521
radlata (Lycaenesthes), 10, 55
radiila (Dielis), 403
,, (Scolia), 403
,. (Tiphia), 403
ralaria (Cidaria), 248
,, (Hannnaptera), 248
ramparia (Macaria), 294
,, (Seiniothisa), 204
rapae (Ganoris), 152
,, (Pieris), xlv, xlvi
,, var. metra (Iheris), xlv.
Ratiaria, 287
rectilineata (Tephrinopsis'i, 294, 295,
296
rectura (Iridopsis), 340
recusataria (Acidalia), 227
regilla (Lycaenesthes), 65
regillus (Lycaenesthes), 64, 65
regularis (Laccobius), lix
regulata (Macaria), 293
rcpandaria (Hannnaptera), 243
,, (Rhopalista), 243
rejietita (Macaria), 292
resistaria (Nematocanq)a), 287
responsaria (Tcphrina), 332
revestita (Strangalia), xxix
Rhagigaster, 407
rhamnata (Phileretne), 265
Rhizobius, 385
Rhodostrophia, 221
rhombaria (Gynopteryx), 302, 304
rhoinboidaria (Boarniia), xxxi
Rhopalista, 243
Rhopalodes, 208, 240, 241, 242
Rhopalosoma, 386, 388, 389
Rhopalosoinidae, xli, 389
Rhopalotettix, 356
Rhopobota, 368, 434
rhotliia (Spilonota), 368
Rhynchopyga, Iv
Rhynchotettix, 357
Rhysopaussidae, lix
Rhyssa, xxxi
rigida (Tinissa), 477
riparia (Labidura), 175, 185
ritseinae (Deuterocopus), 112, 132, 134,
136, 137
robusta (Carcinophora), 183
,, (Cupidesthes), 2, 5, 6, 7, 11,
12
,, (Cymatophora), 341
,, (Melanolophia), 341
rosenbergi (Calocalpe), 365
„ (Psalis), 181
( cxxxiii )
roseoliva (Scelolopliia), 225
rosula (Rhopaloiles), 241
rotalis (Stagmatophora), 453
rotundiceps (Agriomyia), 409, 410
rubens (Salpis), 208, 320, 321, 322
nibi (( 'allophiys), xxi, xxii, xxiii, 85,
86, 87, 88, 89, 90, 91. 92, 93,
94, 97, 98, 99, 100, I'Ol, 102,
103, 104, 105
,, var. fei’vida (Callopbrys), 86, 87,
91, 92
,, var. suaveola (Callopliry.s), 91,
92
,, (Theda), 91
rubricatus (Spliodrotes), 426
j'ubricincta (Lyeaenesthes), 21
nibricinctus (Lyeaenesthes), 16, 21
rubrimaculata (Lyeaenesthes), 29, 84
nibripnneta (Cneinodes), 214
,, (Semaeopus), 214
rnbrodaetylns (Denterocopus), 112,126,
127, 131, 132, 134, 135
, , ( Deuleroseopus), 134,135,
136, 137
irubromaeulatus (Aeolotliynnus), 413
,, (Asthenothynnus), 413
,, (Thynnns), 413
nifa (Formica), xiii, xliv, 143, 144,
146, 147, 150
I’ufibarbis, var. fusco-rntibarbis (For¬
mica), xiii, xliv, 144, 145, 146, 147,
148, 149, 150
rnfieoesia (Anapalta), 249
,, (Enphyia), 249
ruficorne (Pison), 427
rnficorpus (Anapalta), 249
,, (Enj)hyia), 249
rnfifimln'iata (iMicroxydia), 289
rufifrons (My/.ine), 392
rnfinodis (Myzine), 392
nifipcs (Astatus), 520
,, (Coptotettix), 365
rutblnteus (Thynnns), 417
,, (Zaspilothynnns), 417
rntbmarginata (Lyeaenesthes), 19, 54,
56
rnfoplagata (Triclemal, 71, 72
Rumia, 289
Rnmicia, xlvi
rnspator (Helcon), liii
rnspina (Enpliaedra), xxix
Tussns (Formasimus), xxxvi
rnstieata ( Ptychopoda), 218
,, (Sabulodesl, 309
rnsticus (Criocephalus), Ixvi
rntilellum (Coryptilnm), 475
rutilus (Ohrysophanus), 98
xytinatettix, 362
Sabulodes, 307, 308, 309, 320
sabnlosa (Mimophyle), 283, 284
saeva (Cereeris), 422
salaconia (Brentliia), 4 88
I Salamis, 511
Salius, liv, Iv
Salpis, 208, 319, 320, 321, 322, 323,
324
sambnearia (Uraptcry.x), 152
,, (Uropteryx), Ixiv
I Sangaloi)sis, 207
sangra (Zizera), 481, 484, 486, 488,489
: ,, (Zizina), 488, 489, 490, 491,
492, 495
sanguinea (Formica), Ixvii, I.xxii, 144,
145, 146, 147, 148, 149,
150
,, (Lyeaenesthes), 18, 41
sangiiinipunctata (Lissochlora), 211
Sapheneutis, 377
Sapyga, 405
Sapygidae, 405
Sarisophora, 445
Sarracena, 273
saturata (Nipteria), 291
satyrodes (Scardia), 477
Satyrus, Ixix
sanssurei (Mesa), 394
,, (Plesia), 394
,, (Pyi'agra), 166
scabrosa (Tiphia), 396
Scardia, 477
sceletopa (Crocanthes), 445
Scelolopliia, 220, 225, 226
Schoenotenes, 433
scintillula (Lyeaenesthes), 3, 20, 60, 62
scioxantha (Crocanthes), 443
scodionoeata (Salpis), 320
Scolia, 400, 401, 402, 403, 404, 405
Scoliidae, xli, 390, 391
Scoliinae, 400
scolopaiea (Rronchelia), 343
Scoparia, 366
Scordylia, 207, 233, 234
Scotopterix, 331
Scotopterj'ginae, 280
Scotosia, XXV, xxvi, xxvii, 250, 258,
259, 264, 265, 267
scotti (Bi'aehylabis), 193
scudderi (Psalis), 182
scntellaris (Cremastogaster), xx
,, (Laccobius), lix
Sebastia, 275
seenra (Caeoeeia'), 432
sekalavnm (.Eehinosoma), 164
; selenc (Rrenthis), xiii
i Selidoseminae, 280
Semaeopus, 214, 220
^ cxxxiv )
scniiarf:;us (Noniiades), Ix
Heinicaiia (Ectropis), 342
seinidivisa (Atyria), 230, 231
,, (Cyllo])oda), 230
seiiiilugcns (Pluiathecia), 270
,, (Tepliioclystia), 276
Seiuiothm, 288, 293, 294
Seniiotliisinae, 280
seiniustus (Erythroloplais), 221
seiiescens (Tonica'), 456
septeniciiicta (ScoHa), 404
sercna ( Dy.scia), 300
scriavia (Gynopteryx), 302, 304, 305
Serieopliorus, 428
serraticeps (Ctenocephalus), Ixvi
seiTatilinea (Ilyjiolepis), 271, 272
,, (Psaliodes), 271
serratulac (Hesperia), xlvi
serratmu (Calopteroii), Iv
serrulata (Euepliyra), 215
servillei (Xeroiiliyllnm), 347, 350
Sesia, xxxvi
setaria f Ijozogramina), 292
setiger f Allostetlms), 163
seudora (Salpis ', 322
Sibylla (Eupithecia), 279
Sibylliiia, 386, 388, 389
siccata lOinopliila), 375
sicliela (Lycaeiiesthes), 33
sidoiiia fCrocaiitlies), 442
sigillaria (Racheospila), 212
sigillata (Dicliromatopodia), 216
sigiiata (Conops), Ixxi
,, (Scolia), 402
signataria (Cainptogramma), 261
,, (Tephriua), 332
silvaiius (Lycaeiiesthes), 15, 18, 4-"’>
,, (Pseiidodypsas), 43
Siniaetliis, 467
simile (Xeropliylluui), 351
similis (Pliaegorista), xxix
siiiilaciisis (Tiphia), 396, 397
simpliciata (Pterocyplia), 265
simiilicipes (Dinetiis), 530
simulaua (Godaiia), 432
sinensis (X'eolycaena), xxiii
Siosta, 207
siriaca (Bassarodes), 459
sitcllata (Gypsochroa), 259
sjoestedti (Arlex), 189
sjiistedti (Brachylabis), 191
sinaragdina (Oecophylla), 15
Smeriiithns, xxxvii
sinithi (Lyeacnestlies), 17, 31
smitliiana (Spilonota), 368
Snellenia, 440
Sobareutis, 469, 470
Soclichora, 141
sochclioroides(Lcptudeutei'oco]ius),138,.
140, 141
sociata (Epirihoi'), 249
socotrana (Dielis), 402
,, (Scolia), 402
socotranus (Dcnterocopiis), 112, 115,.
I 124, 127, 128, 129
soleata (Dielis), 402
Solenopsis, 146
somnnlentella ( Bedel lia), 374
, sordescens (Perizonia), 208, 269
sparattoidcs (Palex), 162
Spargania, 272
Spartopteryx, 327
spectabilis (Anisolabis), 176
S spernata (Acidalia), 217
,, (Ptychopoda), 217
Sphacelodes, 301
I Sphacridium, lix
Spliecosoma, Iv
Sphegidae, xvii, Ixiv, 390, 418
j Sphegophaga, 390
Sphenarches, 110
Sphex, 419, 420, 518, 519
Sphingidae, xxvii
Sphodrotes, 407, 426, 427, 428
I Spiloinena, 407, 418, 419
i Spilonota, 368, 434
I spilosata (Hydata), 213
Spilosoma, 152
, spini (Klugia), xlvi
j spintheritis (Brenthia), 469
j spleudida (Orgyia), Ixix
I Spododes, 308
sqnalida (Zizera), 486, 487
sqnamigera (Boarmia), 209
,, (I)eileptenia), 209
Stagmatophora, 372, 453
Stainnodes, 238, 239
j Stapliylinidae, 384
' Stathmopoda, 454
I statilinus (Satyrus), Ixix
1 staiidingeri (Lycaeiiesthes), 69
,, (Ncurcllipes), 67, 69, 82
Stanropus, 152
I stefania (Apicia), 306
! ,, (Apiciopsis), 306
I Stegania, 289
stellata (Actizcra), 483, 494, 495, 496,
497
! ,, (Cainptogranima), 258
,, (Gypsochroa), 259
,, (Scotosia), 258
stellidaria (Alniodes), 209
,, (Polysemia), 20i*
Stenaleidia, 331, 332, 333, 334, 335,.
336, 338, 341
Stenaspilates, 312
( cxxxv )
Stenoclonta, 312, 318
Stenoma, 460
Stenomidae, 459
Stenopla, 275, 276
Stenorrhoi^ 274
Steims, 379, 380, 382, 383, 384
stephanitis (Atteria), 433
stereodes (Themeliotis), 476
Steri'hinae, 213
stigma (Scolia), 401
,, (Trielis), 401
stigniatica (Gelasma), 21 1
Stilicus, 384
stollaria (Anisodes), 214
straminea (Pelurga), 255
Straiigalia, xxix, liii
strepsizona (Imnia), 467
sti'iaiia (Toi'tyra), 464
strigata (Thysanopyga), 300
strigosa (Braclimia), 450
Striplinopterygidac, Ixix
strumosa (Lomechusa), 144
Stylopidae, Hi, 514
subarmata (Anisolabis), 176
subcarnearia (Anisodes), 214
,, ab. grisea (Anisodes), 214
subcertaria (Larentia), 265
subcervinella (Hieroxestis), 375
subclararia (Choeiodes), 308
subclatlirata (Macaria), 294
,, (Semiothisa), 294
siibcoeruleus (Zizera), 495
subdepressns (Cryptopliagus), xliv
subnitens (Lycaenesthes), 81, 82
subopaca (Campsomeris), 404
,, (Dielis), 404
,, (Scolia;, 404
subopalaria (Clysia), 307
subpallida (Anisodes), 214
,, ab. figurata (Anisodes), 214
subpisciata (Graphidipus), 239
,, (Graphipidus), 239
subpiilverata (Hammaptera), 245
subqnadrata (Acidalia), 226
subscripta (Tephrinopsis), 208, 297, 298
subsordida (Hymenomima), 339
siibstrigata (Ptychopoda), 221
subulata (Tetrix), 359
subumbrata (Tephrinopsis), 207, 296,
297, 298
subusta (Lipomelia), 222
subvinosa (Hammaptera), 247
suffocata (Xantliovhoe), 257
snffusella (Cryptolechia), 458
,, (Epimactis), 458
sulphurata (Corycia), 289
sumatrana (Gonolabis), 169, 170, 172,
173, 174
sumatranum (Echinosoma), 164
superjecta (Orthonama), 263
suquala (Lycaenesthes), 17, 38
Syagrius, Ixxi
sybavita (jMonopis), 474
,, (Prosena), Ixxi
Sybarites, 273
Sybillina, 389
syllidus (Lycaenesthes), 43
sylvanus (Lycaenesthes), 30
sylvatica (Cieindela), xxx
Synchlora, 210, 211
Synemia, 312
Synia, Ixiii
Synnenria, 208, 239
Syntomidae, liv, Iv
syntricha (Oinophila), 377
syrniaria (Iridoj)sis), 340
I Syrtodes, 327, 328
I Tachynonyyia, 407, 408
Tachypori, 384
j Tachysphex, 425, 524, 525, 526, 527^
I 528
I Tachytes, 521, 522, 523
tamiodes (Sarisophora), 445
Tanagra, 345
Tanychastis, 371
Tapinostola, xxi, Ixxii
Taragmarcha, 370
tansalis (Stenus), 379, 382, 384
tarsata (Forcipula), 186
tartarea (Amauris), 499
tasmanica (Euborellia), 181
Technomyrmex, Ixxii
techowi (Scolia), 400, 401
,, (Trielis), 400, 401
tcctigera (Argyroploce), 436
tegularia (Cambogia), 280
teilinii (Anisolabis), 177
telysaria (Gynopteryx), 302, 304
tenieraria (Crocanthes), 443
tenax (Plristalis), 153
tenera (Hammaptera), 246, 269
,, (Psendisolabis), 188
tengstroemi (Deuterocopus). 110, 112,
115, 117, 118, 120, 124,„
126, 127, 135
,, (Deuteroscopiis), 118
,, (Xeolycaena), xxiii
tennicornis (Paralabidnra), 184, 185
tennimargo (Kacheospila), 212
tenuis (Cyllopoda), 229
Tephrina, 332
Tephrinopsis, 207, 208, 294, 295, 296,
297, 298
Tephroclystia, 276
Tephroclvstiinae. 231
Tephrosia, 254, 332, 333, 342
cxxxvi
tepidata (Glaucopteryx), 254, 268
,, (Larcntia), 254
ternata (l*achropliylla\ 241
tersata (Horisnie), 262
testacea (Lupcrina), Ixii, Ixiii
,, al). iii"i'escc‘us (Luperiiia), Ixii
testaceum (Splieoosoina), Iv
Tetraei.s, 306
tetrica (Scotosia), 250
Tetri.ginae, xvii, 346
Tetrix, 359
thalassitis (Spiloiiota), 434
Thalassodes, 279
Thaleia, 210
thalia (Actiiiote), Ixviii
'I’hainnonoraa, 292, 299, 300
Tliaiimatopoea, xiv
Tliecla, 88, 91
Tliemeliotis, 476
tlieodota (Hairaa), xxix
theoiiae (Leptisolabis), 198, 201
Therniastris, 166
Thestor, xxxvii, 88
Thestoridi, 88, 105
thetis (Agriades), vi, vii, viii, ix, x,
xi. xxxvi, xlviii, xlix, Ixviii,
Ixix
,, ab. coelestis (Agriades), ix
,, ab. inarginata (Agriades), ix
',, var. puiictif'era (Agriades), ix
,, ab. ixiiicolor (Agriades), ix
,, (Lycaeiia), 487
,, (Polyonimatus), xlvi
theto)ia (Nepliogenes), 455
thiantlia (Jlonopis), 473
Thomosis, lix
tlioracica (Platylabia), 162
,, (Psalis), 167, 183
Tiiorjidonta, 358, 359
thoraea (Ethmia), 461
thrasydora (Crocautlies), 444
Tliriambeiitis, 470
thryptica (Hylogi’aptis), 451
Thyuiiidae, 390, 407, 410, 416
Thyiiims, 407, 411, 413, 417
Thyrinteina, 343
tbyrsis (Cupidestbes), 6, 10, 11
, , (Lycaenesthes), 1 0
Tliysanopyga. 300, 301
tibiale (Heinbidiiini), lix
Tinea, 474, 475
Tiiieidae, 374, 473
Tineina, 430
Tiugentera, 437
Tinissa, 476, 477
Tipba, 437
Tiphia, 392, 394, 395, 396, 397, 403
Tipbiinae, 394
Tirallis, 437
tisamenus (Lycaenesthes), 74
,, (Tricleina), 71, 74
Tisis, 437, 438, 439, 440
Titanolabis, 167, 168, 176
togoensis (Ctenisolabis), 190, 197
Tomopteryx, 241
Toinopygia, 184, 187
Tonica, 456
toraula (Cosuiopteryx'i, 452
tortilinea (Psodopsis), 282
Tortricidae, 431
Tortricina, 430
Toi’trix, 432
Tortyra, 462, 463, 464
Trachytettix, 346, 348, 354
transitaria (Broncbelia), 343
transversata (I’liilereine), 265
transvisata (Macariu), 293
Tretothoracidae, lix
Tre to thorax, lix
triangularis (Anteois), 220, 221
;, (Jletasiopsis), 221
,, (Ptychopoda), 221
trias (Acidalia), 227
Tricentra, 220
Tricbopteryginae, 231
Triclioptilns, 367
Trichura, Iv
Triclema, 2, 3, 66, 70, 71, 72, 73, 74,
75, 76, 78, 79, 80, 81, 82, 83
tridens (Acronycta), Ixi
tridentata (Pachycopsis), 211
Trielis, 400, 401
trifolii-minor (Antbrocera), Ivii, Iviii
trigonodes ( Ancyloxnetis), 371
trilobata (Hexadactilia), 108
tripartita (Thamnonoina), 299
Triphosa, 260, 266, 267
'ITiptila, 242
trisecta (Callipseustes), 327
triseriata (Paraethria), Iv
trispiuosa (Forcipula), 186
tristani (Pyragropsis), 167
trixysta (Oinoi)hila), 376
Ti’ogopbloei, 384
truncata (Dysstroiua), 271
,, (Labidura), 185
,, rufibrunnea (Polypliasia), 271
truncatijxeunis (Eupithecia), 276
,, (Tephroclystia), 276, 278
truncatus (Colobopsis), xx
Truxalis, xxxviii
Trypopbylluni, 346, 348
turbata (Canibogia), 226
,, (Lycaenesthes), 65
,, (Scelolophia), 226
tui'batus (Lycaenesthes), 2, 65
( cxxxvii )
turbatus (Nemypexina), 65
turneri (Zcugophora), Ixvii
tzaddi (Litliostege), ’236
Uganda (Lycaenestlies), 17, 32
ukerewensis (Lycaenestlies), 23, 30, 83
umbratus (Lasius), lx, Ixvii, 144
umbrinata (Pterocy[)ha), 264
,, (Scotosia), 264
,, ab. velutina (Scotosia), 264
umbripennis (Aeolothynnus), 417
undiliiiea (Anisoperas), 306, 307
undularia (Tephrosia), 254
undulata (Scotosia), xxv, xxvi, xxvii
undiilatum (Acinophyllum), 349, 355
unicolor (Eudule), 233
,, (Euplianessa), 231
,, f. lyelli (Rbagigaster), 407
,, f. inutatus (Rbagigaster), 407
unicornata (Acidalia), 228, 229
,, (Craspedia), 228
uniformata (Carsia), 239
,, (Synneuria), 208, 239
uniformis (Pergama), 318
„ (Pero), 318
Upenora, 290
uranitis (Imnia), 465
Urapteryx, 152, 287
Urepione, 288
Uropteryx, Ixiv
urticae (Aglais), xxvii
usambarana (Leptisolabis), 198, 201
ustifuinosa (Biyoptera), 332
,, (Cymatophora), 33
,, (Stenalcidia), 332, 341
uxiaria (Apicia), 302, 304
vallonia (Urepione), 288
Vanessa, 151
varia (Luciola), 532
,, (Melitaea), Ixxiii
varians (Pachrophylla), 241
variaria (Pero), 313, 314
variegata (Mascora), 326
,, (Rhopalodes), 240
,, ab. derufata (Rhopalodes),
240
varipes (Cerceris), 421
venusta (Cerceris), 421
Verhoeffia, 191
verhoeffi (Gonolabis), 169, l7l, 175
versatilis (Lycaenesthes), 19, 49, 50
Vespidae, 389
vesta (Anisoperas), 307
vestita (Oenothalia), 301
vestitaria (Nematocampa), 287
vexans (Quedius), 384
vicarius (Hippodes), 347
vicina (Chrismopteryx), 253
victrix (Cerceris), 420, 422
vinosa (Cycloniia), 300
,, (Fidonia), 800
vinula (Cerura), 151
violacea (Campsomeris), 400
,, (Diliacos), 400
,, (Scolia), 400
violaceus (Carabus), xliii
,, var. exasperatus (Carabus),
xliii
virescens (Phacelophora), 242
,, (Stenopla), 276
j viretata (Acacis), 243
I virgaureae (Heodes), xlvi
virgellata (Synneuria), 239
I vittata (Oxypoda), 384
j vittulata (Ortlionania), 263, 264
I ,, (Pbibalapteryx), 263
' voeltzkowi (Brachylabis), 196
,, (Metisolabis), 196
voltae (Cupidesthes), 5, 6, 7, 11
,, (Lycaenestlies), 7
,, var. gabunica (Lj'caenestbes), 8
vosseleri (Anisolabis), 176
vulgata (Eupithecia), 277
vulneraria (Brotis), 301
,, (Sphacelodes), 301
waddyi (Psalis), 182
ivalilbergi (Echinosoma), 164
,, (Euralia), xiv, xv, xvi, 498,
499, 500, 502, 503, 504,
505, 506, 507, 508, 509,
510, 511, 512, 513
,, var. niadagascariensis (Eu¬
ralia), 499
,, (Tricboptilus), 367
walkeri (Forcipula), 187
westerinanni (Echinosoma), 164
westwoodi (Hypsaeus), 347
weyenbergliii (Eudule), 232
willeyi (Nannisolabis), 194
wilverthi (Paratettix), 361
woodwardi (Gonolabis), 169, 171, 173,
174
,, var. dentata (Gonolabis), 174
Xantbandrus, xxv, xxvi, xxvii
Xanthia, Ivii
xantbocblorata (Sarracena), 273
,, (Sybarites), 273
xanthonota (Eretmocera), 455
xanthopoecila (Ijycaenesthes), 51
xantbopoecilus (Lycaenestlies), 19, 51
52
xantbopus (Labidura), 185
Xantborboe, 208, 257, 268, 269
Xanthospilopteryx, xxix
Xanthyris, 230
xenia (Anisolabis), 176
Xenoecista, 299
( exxxviii )
xerophylloides (Acinopliyllum), doS
,, (Medinophyllum), 349
Xerophylluni, 34 d, 347, 350, 351, 352,
353, 354
xuthogastia (Cosmopteryx), 452
xylinata (Isclinopteryx), 328
,, (Syi'todes), 328 ,
Xylolocha, 307
Xylopliasia, xlvii
Xyloryctidae, 457
yorkense (Ecliiiiosonia), 104
Ypsi petes, 251, 267
Ypsolopliiis, 452
ytenensis (Laceoldus), lix
Zaspilothyniius, 417, 420
Zeleboria, 411
zeiikeri (Lycaeiiesthes), 20, 62
zenobia (Papilio), xxix
Zerene, 205
zeiitis (Heterusia^, 234
zeritis (Scordylia), 234
Zeugophora, Ixvii
Zizeevia, xvii, 480, 481, 482, 483, 484,
485, 486, 487, 488, 489, 490, 491,
492, 493, 494, 495, 496, 497
Zizeeriidi, 480, 481, 482, 483
Zizera, xvii, 479, 480, 481, 482, 483,
486, 495
Zizina, 482, 483, 488, 489,490,491,
492, 495, 496, 497
Zizula, 483, 492, 493, 495, 496, 497
zollikoferi (Xylopliasia), xlvii
,, ab. pallida (Xylopliasia),
xlvii
zooliiia (Cliaraxes), xv, xli, 506
,, f. neantlies (Cliaraxes), xli
zona (Euphyia), 250
,, (Scotosia), 250
Zygaena, xiii
TRANSACTIONS
OF THE
ENTOMOLOGICAL SOCIETY
OF
LONDON
For the Year 1910.
I. A Revision of the African species of the Lycaenesthes
group of the Lycaenidae. By G. T. Bethun e-Baker,
F.L.S., F.Z.S.
[Read October 6th, 1909.]
Plates I— XIIL
The genus Lycaenesthes was created by Moore in 1865
(Proc. Zool. Soc. Bond,, 1865, p. 773) for the reception of
his species L. hengalensis, i.e. emolus, Godart. The neura-
tion is quite ordinary, and is very closely allied to Nacaduha
and Jamides, from which however, it may be known by the
possession of three short hair-like tails in the secondaries,
this characteristic will also separate it from almost all
other genera of the Lycaenidae. The Oriental species
require a good deal of sifting, whilst those obtaining in
Africa are much in the same condition in spite of the
fact that Aurivillius dealt with the genus so admirably in
his able work, “Rhopalocera Aethiopica ” ; since the issue of
that book, however, a considerable amount of material has
come to hand, and we are thus in the position of being
able to elucidate many points and species. Africa may be
said to be the headquarters of the genus; it was the un¬
ravelling of many species from the West Coast, together
with Doherty’s collection from the Mau Escarpment in
the Tring Museum, that necessitated my taking up a more
detailed study of the genus, and thus led to the present
TRANS. ENT. SOC. LOND. 1910. — PART I. (JUNE) B
*2 ^Ir. G. T. ] 5{iiPinie-Baker's Revision of the
Revision of the African species of the group. In 1893
Kar.scli divided the genus into two, creating a second,
Triclema (Berlin Ent. Zeit., 1893, p. 227) with lucretilis,
Hew., as its type ; the difference in the two genera lying in
the subcostal veins, Lycaenesthes having four, Triclema only
three, whilst vein 11 anastomoses with 12 in Triclema
but not in Lycaenesthes. Aurivillius (in op. cit.) objects to
this subdivision on the ground that in one species vein 8
is very short and almost wanting. It is with very great
hesitation that I venture to differ from so able and revered
an authority as the celebrated Swedish professor, but the
fact remains that in the species he cites — turhatus, Grose-
Smith (a synonym that must fall before lyzanius, Hew.) —
the vein 8 remains : it is short, no doubt, but it remains,
and therefore it does not belong to Triclema. This opens
a wide subject, viz. •“ On what characteristics may we found
new genera?” It is now almost universally conceded that
where structure differs, another genus should, or may be
erected ; and the learned professor himself acts on this
theory, for in this group that we are dealing with he has
created the genus Cupidesthes (Ent. Tid., xiv, 1895, p. 215),
on the ground that the eyes are naked, that the shape of
the wings differs slightly, and that the species rohusta,
Auriv., is, as its names signifies, more robust. Now the
neuration of the Lycaenidae is one of their most constant
and reliable characters. Many years’ study of the group
from all over the world has certainly revealed a remarkable
constancy of this character, whilst the study of this one
genus only has revealed the fact that in more than one
species the eyes may be hairy or not in individuals of the
same species. I hold the view that neuration is a very
important structural point, and, generally speaking (apart,
of course, from those exceptional genera where tlie sexes
differ in neuration as also in some cases individuals like¬
wise, but these are probably instances of change; the
species and the genera being in a period mutation), is
sufficient of itself to establish the validity of a genus.
Under these circumstances I cannot do otherwise than
maintain the validity of Karsch’s genus; but more than
this, I am Avith some reluctance perhaps, raising two
additional genera for the group. In several species vein
11 and 12 anastomoses and there are four subcostal veins,
whilst in several other species veins 11 and 12 do not
anastomose, but there are only three subcostals. 1 do not
African species of the Lycaenesthes group of Lycaenidae. 3
consider that tliese can rightly be put in either Lycaenesthes
or Triclema, so that for the sake of the observer of the
future it appears to me to be advisable to further sub¬
divide the genus, then the man who follows me hereafter
may be able to show whether the group has become more
homogeneous, or whether it has been further subdivided
in its struggle for existence. The whole group has proved
to be very complex and most difficult to treat ; the under¬
side pattern separates itself into two well-marked sections,
the upperside separates itself into four or five equally well-
marked sections, whilst there are individual species that
stand out quite separate from these ; unfortunately for the
systematiser, however, these well-marked groups according
to colour or pattern do not run structurally along similar
lines, but fall somewhat indiscriminately into the various
genera. The genus Lyeaenesthes takes all the species
(sens, strict.) and forms a most homogeneous section, but
apart from these a large number of a quite different pat¬
tern and colour must be included in the genus in its wider
application, so that we find no less than 56 species falling
under its range. The other genera are more homogeneous
on the whole, the one or two divergences occurring will be
found mentioned under the respective species. I still con¬
sider that a genus should not be raised on the character of
pattern and colour only ; our knowledge of the physiological
changes required to produce an alteration in pigment colour
or pattern is almost nil, and therefore apart from structure
it should not be used. The development of pattern of the
underside is readily followed through the somewhat simple
markings (simple for Lycaenidae) of the musagetes-ligures
section, through the heavy pattern of the larydas group
into the complex design of the lysicles-scintihdla sub¬
sections. It is also interesting to find in tropical Africa
an almost precisely similar evolution of colour as obtains
in the temperate regions of the world; we have in this
small group blues of all shades, browns, and coppers — the
latter a colour quite separable from the coppers of the
Capys and Phasis genera of South Africa, and it also differs
from Heodes (if the species usually cited under it really
belong to that genus), which is more closely allied to our
Palaearctic Chrysophanus. The sexes are occasionally
somewhat different, this being more noticeable in the
species lachares originally described by Hewitson from a
female, and being brown with a largish yellow patch,
B 2
4
Mr. G. T. Be^^ine-Baker’s Revision of the
whose male was unrecorded until Grose-Smitli described
it under the name <pulchei\ this being of a beautiful sub¬
dued lustrous blue sometimes variegated with a more
subdued ochreous patch ; the underside of the female at
first sight is very different from the intricate and closely
packed pattern of the male, but careful examination reveals
the pattern in embryo, and its evolution into that of the
male can be easily imagined. The life history is not
known of any of the African species, but de ISiceville has
given us the life history of two of the Eastern species,
L. lycaenina (J. A. S. B., Ixix, p. 237) and L. emolus (But!.,
India, vol. iii, p. 129) ; they are both attended by ants.
The scales of the wings present no points that call for
special attention except for the fact that the long battle¬
dore scales are very few in number.
All the species of the group are robustly built, with
strong triangular primaries and fairly full roundish
secondaries, betokening a brisk, restless habit ; the thorax
is unusually robust considering the small size of the insects.
They are all strong-flighted, wary little creatures, fond of
sunninor themselves on the foliage of trees, and all — of
which we have observations — delighting in flowers and in
settling on damp spots for the benefit of the moisture they
can imbibe. Though small in size they are of a pugnacious
disposition, and will often take up a position on a twig
and give battle to any passer by, even though it may be
an insect much larger than itself, returning very frequently
to the same twig time after time. I am greatly indebted
to Dr. Holland for most beautiful coloured figures painted
amidst very great pressure of work, by his own hands of his
species now in the Carnegie Museum, which have saved me
from more than one probable error; also to Mr. H. H.
Druce for access at all times to his valuable collection,
and to Prof. Aurivillius and M. Mabille for the loan of
their types. Had it not been for the kindness of these
gentlemen, it is doubtful if some of the species would have
been properly identified, and I now accord them my hearty
thanks, as also to Mr. Heron of the British Museum, for
constant and unfailing courtesy and help at all times.
Genus Cupidesthes, Aurivillius.
Synonymy : Genus Cupidesthes, Auriv., Ent. Tid., 1895,
p. 215; id. Rhop. Aethiop., p. 345 (1898); Lycaenesthes,
African siJccies of the Lycaenesthes yroxi'p of Lycaenidae, 5
Hew., Ill. Diurn., p. 224 (1878) ; id. H. H. Druce, Ann.
Mag. N. H., 1890, p. 24 ; id. id. Sharpe, p. 105 (1890) ; id.
Crowley, Trans. Ent. Soc., Lond., 1890, p. 555; id. Smith
and Kirby, Rhop. Exot., p. 110 (1894); id. Auriv., 1. c. m
parte ; id. Beth. -Baker, Ann. Mag. N. H., 1903, p. 333.
In creating this genus Aurivillius stated that it was
very near Lycaenesthes, but differed through its quite
naked eyes, through the almost straight costa of the
primary, through the truncated inner edge of the second¬
ary, and through the much stronger build of the whole
insect.
I define it as follows —
Caput and frons hairy, eyes glabrous or hairy more or less, palpi
scaled, end segment longish, nearly naked, i.e. quite smooth. Thorax
and wings robust. Wings ; costa of primaries nearly straight, broad
and triangular ; secondaries less ample, termen truncated, anal angle
reduced. Neuration : primaries, vein 2 at a third from the lower
angle, 3 fr^m just before, 4 from the angle, 5 from about the middle
of the discocellulars, 6 from just beyond the upper angle of the cell,
8 stalked from 7 at half to a third from the apex, 9 absent, 10
and 11 free from the cell ; secondaries, with two inner veins, la
and 16 ; vein 2 given off from beyond the middle of the cell, 3 and
4 from the lower angle, 5 from about the middle of the cell, 7 from
the cell, 8 highly curved at the base and long. The genitalia are
also very different to those of the following genera, the clasps
especially being of a totally different structure, as will be seen from
the descriptions of these organs.
Type, Cupidesthes rolxista, Auriv.
Prof. Aurivillius lias been so kind as to lend me
his type, which is intermediate in size between the smaller
leonina xnihi and the larger voltae, Sharpe. I am including
several species in the genus with more or less hairy eyes
they so obviously are of the same type of insect, being
decidedly larger than Lycaexiesthes in one or both of the
sexes, with the same class of pattern on the underside
and similarly restricted blue, generally bright, on the
upperside, so that I find it impossible to locate them
elsewhere. I shall consider the eyes under each individual
species.
G
Mr. G. T. Bu^i no-Baker’s Revision of the
Key to Figure for Description of Genitalia.
a. Hai'pago (in pairs) ; also called clasps.
b. Ciugula ; consisting of two arms or a girdle confluent with
the tegumen, and jointed to the harpagines.
c. Tegumen ; always divided into two sides or cheeks in the front.
d. Falces (in pairs) ; also called hooks.
e. Furca ; consisting of two arms from a common base in the har¬
pagines, the support to the penis.
/. Penis sheath.
j KEY TO SPECIES.
A. Blue above.
1. Underside : primaries with a spot in
middle of cell, secondaries with a
series of sub-basal spots below
each other . rohusta.
2. Blue very restricted above in
primaries. Below with markings
large, postinedian stripe in
primaries very broad, no sub-basal,
series of spots in secondaries . . voltae.
3. Palest bluish green above restricted
in primaries . arescopa.
4. Small in size. Below pattern consist¬
ing of small spots . tJiyrsis.
5. Small in size. Below pattern with
larger spots, postmedian stripe in
primaries very broad . lythas.
6. Small in size. Brown with a deep
leaden blue lustre above, pattern
below very large on a dark ground
in , on a white ground in the 5 leonina.
African species of the Lycacncstlies group of Lycacnidac 7
B. Brown above.
1. Both wings uniformly brown above
and below, the undersurface having
numerous fine white lines . . . bninneus.
2. Both wings white above ; secondaries
with a terminal row of black spots.
Below white with pattern small
and broken up . mimetica.
Giqyidesthes rdbusta, Auriv, (Plate I, fig. 3.)
C. robusta, Auriv., Ent. Tid., 1895, p. 215; id. Rhop.
Aethiop., p. 845 (1898).
$ Eyes almost glabrous ; under a low-power lens ( x 6) the eyes
appeared to be glabrous, but under a higher power ( x 10) a few
isolated hairs were visible. Both wings brown with restricted bright
blue area. Primaries, with the blue occupying the cell, evenly curved
off to about a quarter from the cell on the inner margin. Secondaries
with the blue occupying the cell and nearly to the angle and termen,
but leaving the apex broadly brown ; the cell closed by a dark mark.
Underside very pale brown largely covered with white scales.
Primaries with a spot in the cell and a larger L-shaped one at the
end, postmedian stripe very irregular, first spot on the costa below
which are two at an angle follow’ed by a fourth shifted inwards, very
oblique, fifth again inwards less oblique, sixth scarcely traceable, a
very obscure subterminal scalloped stripe. Secondaries with three
sub-basal marks, below the third, which is a dash, are two round
spots on the inner margin, the upper one being very pale ; a large
oblique spot closing the cell ; postmedian stripe divided into three
parts, the first two spots near the large cell spot, spots 3 to 6 shifted
right out in an irregular row, spots 7 and 8 angled and shifted right
inwards ; a trace of a subterminal line •, marginal spots in the anal
area iridescent.
Expanse 36 mm.
Hah. Cameroons (Kitta), April.
Type, Mus. Holmiae (Stockholm).
Gupidesthcs voltae, E. M. Sharpe. (Plates I, fig. 1 ;
IV, fig. 1.)
Lycaenesthes voltae, Sharpe, Ann. Mag. N. H., 1890,
p. 105 ; id. Crowley, Trans. Ent. Soc., Lond., 1890, p. 555,
Pl. 18, f. 6 ; id. Auriv., Rhop. Aethiop., p. 349 (1898).
8
JMr. G. T. Be^iue-Baker’s Bevision of the
Primaries brown with the area below the cell and to about
half vein 2 palish violet blue. Secondaries entirely violet blue as in
the primaries with a short apical dark brown dash. Underside : both
wings greyish white, with the usual markings of the ground-colour
with a double edging first broadly of pale brown then finely of
white. Primaries, with the postrnedian stripe irregular, very broad,
the four subcostal spots confluent oblique, fifth spot ovate shifted
inwards, sixth spot very broad its inner margin shifted far
inwards, seventh spot waved quadrangular shifted well in, its
inner edge reaching the angle of vein 2. Secondaries with the post¬
median stripe twice fractured, not so broad as in the primaries, the
third, fourth, and fifth spots confluent shifted well outwards, sixth
spot large its inner edge shifted well inwards, seventh and eighth
angled spots inwards again — anal spots reduced to very small dimen¬
sions.
9 . Primaries brownish with a large white patch occupying
the cell the fold and lower radial area to the inner margin.
Secondaries entirely white with termen broadly brownish with an
obscure row of scalloped brown spots. Underside of both wings
white, pattern as in the male, but obscure with fine obscure outlines.
Genitalia. — Harpago very specialised, of very moderate breadth at
the base, very rapidly tapering down to a long, narrowish arm, the
whole of the upper edge straight, tip terminating in a blunt point
with its upper edge dentated. Cingula rather narrow, expanding
but little to the tegumen, which is well excised at the front of the
dorsum, cheeks with a straightish front edge. Falces very fine and
long, highly curved at a third. Furca with short arms curved
backwards rising directly from their base. Penis sheath very long
and exceedingly narrow, the hinder two-fifths being of moderate
width. Tegumen and harpagines with very long, strong bristles
extending far down the latter.
The genitalia alone of this genus are so specialised, especially in
the harpagones and penis sheath, as to be almost sufficient alone to
substantiate Aurivillius’ genus.
Var. gabunica, Auriv.
Differs only in that it has no white patch.
Ifah. Volta River ; Sierra Leone ; Ashanti; Uganda.
Type in the British Museum.
This species should be readily known by its large size
and palish violet blue colour. It avas originally described
from a female, the only male I have seen being that from
Ashanti in the British Museum.
African sjiccies of the Lycaenesthes group of Lycacnidae. 9
Cupidesthes arcscopa, sp, nov. (Plates I, fig. 2 ; IV, fig. 2.)
g . Both wings blackish brown. Primaries with a restricted area
of very pale silvery greenish blue, restricted to below the cell.
Secondaries with the pale bluish area below vein 6, extending well
beyond the cell and up to the abdominal margin and the tornus,
these latter areas becoming whitish. Underside : both wings pale
brownish almost entirely covered with white scales, so as to look
w'hitish, the spots being only marked out by their brown margins,
but being filled in with white. Primaries with a spot closing the cell ;
postmedian stripe very broad, the third and fourth spots confluent
shifted outwards, fifth and sixth each slightly inwards ; to this point
the spots form one continuous band, but the first spot on the costa
is absent, the seventh spot is isolated and shifted right inwards
so as to be below the spot closing the cell. Secondaries with a
sub-basal subcostal pale brown spot, an oblique spot closing the
cell ; postmedian stripe with the third spot shifted out on to the
outer edge of the second, fourth and fifth confluent shifted slightly
out, sixth oblique shifted slightly in, seventh and eighth angled
spot shifted far in below and almost to touch the spot closing the
cell, an indefinite subterminal brown line interrupted at the veins,
a trace of a preterminal interrupted line, a small black terminal
spot between veins 2 and 3 and a larger one at the angle, with a
trace of a few metallic scales and slight yellow internal edges.
5 . Both wings darkish brown. Primaries with a white central
roundish patch. Secondaries with a larger patch below vein 6 to
the inner margin, and occupying about half the radial area. Under¬
side as in the male.
Expanse $ 87, P 42 mm.
Ifab. Bitje; Ja Kiver, Cajieroons.
Type in the National Collection.
Mr. H. H. Druce has a series of this species from the
same locality with females, and my description of this sex
is from one of his specimens. I have no doubt in my
own mind as to the determination of the sexes.
Genitalia. — Harpago somewhat roughly and narrowly reniform at
the extreme base, suddenly reduced to an exceedingly fine, narrow,
long point, with long, strong bristles most of its length. Cingula
rather narrow, tapering wider towards the tegumen, which is well
excised in the front of the dorsum (or dorsal apex). Falces very long
and narrow, well curved at a quarter from their sockets. Furca
with short, narrow arms rising directly from the harpagines, and
10
Mr. G. T. B^une-Baker’s lievision of the
evenly curved backwards. Penis sheath with the hinder half
rather Indian-club shaped, the fore half very narrow with a
blunt, rounded tip.
Cupidesthes thyrsis. (Plate IV, fig. 3.)
Lycaenesthes thyrsis, Hew., Ill. Diurn. Lep., p. 224,
PI. 92, ff. 42-44 (1878) ; id. Auriv., Rliop. Aethiop., p. 349
(1898).
This species is easily distinguished by its smaller size, the blue
in the primaries is confined to the cell, to below vein 2 including
all the area below the cell, but not reaching into the tornus; in the
secondaries the whole of the wing is bright blue to vein 6. The
underside pattern is very typical, very irregular as to its postmedian
stripes, but not absolutely fractured as in the type of the genus.
The species should be easily recognisable. The female is entirely
brown above.
Hal). Gaboon, Fernando Po ; Ogowe, French Congo
{Coll. B.-B.).
Type in Mus. Brit.
Though this species is decidedly smaller than the type
species, I have no hesitation in placing it in this genus,
the pattern obtaining in precisely similar fascies ; the eyes
of my male, whilst not glabrous, yet are not hairy — in the
female they are hairy. I do not, however, trust this
character, my doubting it seriously has made me test it,
and I have found it very easily removed. Lycaenesthes
larydas, Cram., has densely hairy eyes; in one specimen of
this insect I removed the hairs with a gentle application
of a soft camel-hair brush, in two other specimens this
failed, but they immediately yielded to a quite gentle
treatment from a small hogs-hair brush. This shows that
as a character it should be used with very great caution
as it is open to accidental alteration.
Genitalia. — The harpago is reduced to an exceedingly narrow
process for the basal half being highly and sharply excavated on
its upper edge, forming a sharp tooth, the curve below being very
rapid, the lower portion is then produced forward into a long,
sharp point rather longer than the basal part ; the cingula is well
developed, its basal hind extremity being projected backwards
into a long, bluntly ended tube, its girdle expands rapidly to the
cheeks of the tegunien which are very fully developed with a fairly
A frican species of the Lycaenesthes group of Lycaenidae. 1 1
erect but uneven front edge ; the fakes are very long, sharply
curved, almost at a right angle at a third from their sockets, the
extremity forming a small talon ; the dorsal apex is very highly
excavated as to its fore margin only ; the furca is very short,
divided almost from its base, and sharply curved backwards
from its base also. The penis sheath is very “ Strymonid ” in shape,
its hinder two-fifths being broadish, the fore three-fifths tapering
narrowly to its orifice which suddenly expands and is somewhat
trumpet-shaped.
The cheeks of the tegumen are well furnished with strong, long
hairs, the harpagines being scantily furnished with them, a few of
which are, however, of exceptional length.
Cupidesthes lithas, H. H. Druce. (Plate IV, fig. 4.)
Lycaenesthes lithas, H. H. Druce, Ann. Mag. N. H.,
1890, p. 24; id. Smith and Kirby, Rliop. Exot., p. 110,
PI. 24, ff. 8, 4 (1894) ; id. Auriv., Rhop. Aethiop., p. 348
(1898).
A species very close on the upperside to the previous one, but
the blue is less bright, not lustrous, and in the secondaries it often
extends up to vein 7, and generally beyond vein 6. On the under¬
side it differs more, the pattern is not composed of small spots,
but of large ones, the postmedian fasciae, especially of the primaries
consisting of very broad conjoined spots after the style of G. voltae,
whereas G. thyrsis has the pattern more like that obtaining in
G. robusta. There is also the subcostal spot near the base on the
secondaries which is lacking in thyrsis. The blue of this species
may be variable, as I have a specimen in which it is quite mauve
in tone. The female is entirely brown in both wings, with a
trace of white showing beyond the cell in the primaries, and
a row of four scalloped marginal black spots in the anal area
of the secondaries, the underside being like the male but slightly
modified.
Hah. Sierra Leone ; Ashanti (Adda).
Type, Coll. Druce.
The insect described by Grose-Smith as the female of
this species is the male of lunulata, Trimen, a species
wddely distributed and equally widely confused by authors.
Writers, however, of even twenty years ago ought to be
excused much, the material available and the means of
identification were very different tben to Avhat they are
to-day. The female is brown without any blue, this
12
Mr. G. T. Jl^iiine -Baker’s Revision of the
obtains throughout the genus where both sexes are known,
and may be expected to prove to be the case in robusta
also.
Genitalia. — Harpago of moderate width at base, but rapidly
and evenly tapering to a point. Cingula well developed, the
basal bind extremity produced far backwards, and ending in a
small globose termination, the girdle or arms taper gradually up to
the cheeks of the tegumen, which have erect front edges ; the falces
are very long and fine, highly curved at about a third from their
socket ; the dorsal front apex is deeply excavated. Furca shortish,
with fine erect arms. Penis sheath somewhat “ Strymonid ” in
shape, with the hinder two-thirds hroadish, then gradually tapering
narrowly to an oblique apex ; both the cheeks of the tegumen and
the harpagines are well furnished with longish hairs, the latter
having them almost to the base.
Gupidesthes leonina, B.-B.
Lycaenesthes leonina, Beth. -Baker, Arm. Mag. N. H.,
1903, p. 333.
(J. Both wings dark brown. Secondaries wuth a leaden lustre in
the cell and below it, and with four or five terminal dark .spots
edged palely inteimally. Underside: both wdngs dark brown with
spots palely edged. Primaries with a spot closing the cell and
one above it. Postmedian stripe broad, third and fourth spots
conlluent shifted outwards, fifth well inwards, sixth inwards again
and largi.sh, seventh very large right inwards, terminal area broadly
whitish, with an internervular series of scalloped marks. Second¬
aries with a sub-basal spot below the costa and on inner margin, a
spot closing the cell ; postmedian stripe beginning and ending right
over and under the cell spot, second spot shifted outwards, third and
fourth confluent shifted outwards, the fourth being slightly further
out than the third, fifth and sixth small, and each shifted in, seventh
and eighth angled spot right in ; terminal area broadly whitish with
a lunular dividing line, a black terminal spot between veins 2 and
3, and at the angle edged internally wuth yellow, with a trace of
iridescent scales.
Expanse 32 mm.
The female is entirely brown, but I have a specimen from Busoga
w’ith a wUite postmedian area in the primaries and a whitish curved
postmedian stripe in the secondaries.
Hah. Sierra Leone ; Upper Congo ; Busoga ; Uganda
African species of the Lycaenesthes group of Lycacnidae. 13
{Mincliin)-, Gold Coast {Dudgeon)-, Volta River {in
]\his. Brit.).
Types, ^ in my collection, ^ in coll. Gator.
When I described this species originally, from Sierra
Leone, I was much puzzled as to the sex in the absence
of the forelegs ; since that time, however, I have received
an undoubted male from the Upper Congo, and the British
Museum have also received a male, which though entirely
brown above is certainly the same species, the latter mates
well with my Busoga
Gupidesthes brunneus, Smith and Kirby,
Lycaenesthes brunneus, S. & K., Rbop. Exot, p. 106,
PI. XXIII, £f. 18, 14 ; L. br'unnca, Auriv., Rhop. Aethiop.,
p. 348 (1898).
9 . Uppersicle uniformly brown in both wings. Secondaries
with a terminal row of black spots, decreasing rapidly to the apex
where they become minute, the row is finely edged laterally with
whitish. Underside uniformly brown with numerous whitish lines
in both wings. Secondaries with two small dark spots on the
costa.
Hab. Ogowe River.
Type in the Carnegie Museum.
I only know this species from Smith and Kirby’s figures,
but it is evidently near my C. leonina. The pattern
below is, however, quite different, and it has no white
ground-colour at all, either above or below. I think there
is not much doubt that I am right in placing the species
in this genus.
Gupidesthes mimetica, Druce. (M.S.)
This species ditiers from the preceding in being practically all
white both above and below, the pattern below is also different,
being much broken up.
Hab. Bitje ; Ja River, Cameroons.
Type in the Druce collection.
Mr. Druce has let me see this species before his descrip¬
tion has appeared. It is a very distinct insect, quite
different from others of the genus. Its clear, cream-white
wings should make it recognisable at a glance, and as I
14
Mr. G. T. l!^^une-Baker’s Revision of the
understand he will figure it in his paper, I would refer my
readers to his plate and description.
Genus Lycaenesthes, Moore.
Synonymy : Lycaena, Trimen, Rhop. Afr. Aust., p. 234
(1866) ; Lycacnedhes, Moore, P. Z. S., 1866, p. 773; id. Hew.,
Trans. Ent. Soc. Lend., 1874, p. 343; Ill. Diurn. Lep.,
p. 219 (1878) ; id. Moore, Lep. Ceylon, i, p. 87 (1881) ; id.
Distant, Rhop. Malay, p. 232 (1884); id. Trimen, S. A.
Butt., ii, p. 93 (1887) ; id. de Nic^ville, Butt. Ind., iii, p. 127
(1890) ; id. Karsch, B. E. Z., p. 214 (1893) ; id. Reuter, Act.
Soc. Sc. Fen., 22, i, p. 182 (1896) ; id. Auriv., Rhop.
Aethiop., p. 345 (1898) ; id. id. Arkiv. Zook, ii, p. 16 (1905) ;
id. Holland, Psyche, vi, p. 50 (1891); id. id. Ent. News,
iv, p. 25 (1893); id. Smith and Kirby, Rhop. Exot., ii, p.
97 (1893) ; id. id. iii, p. 137 (1901) ; Pseudodypsas, Reuter,
1. c.
“ Eyes hairy. Palpi long, compressed porrect ; third joint long,
attenuated, half the length of the second. Legs moderate ; femora
slightly pilose beneath ; mid and hind tibiae with two short apical
spurs. Antennae slender at the base, thickened near the end which
is finely pointed. Thorax and abdomen robust. Wings moderately
broad ; f.w. with costal margin arched at the base ; apex rather
acute ; exterior margin slightly oblique ; subcostal vein with
first branch arising at one-third the length of the wing, second
and third equidistant, fourtli remote, fifth joined at the base to the
third ; h.w. rounded exteriorly ; two small very fine tail-like
fascicles of hair near anal angle. Lycaenesthes bengalensis type( =
moZtts).”— (P. Z. S., 1865, p. 773.)
The African sjDecies agree entirely in neuration and
other generic characters with the type named by Moore,
whilst the species themselves (sens, strict.) follow the
general pattern of the underside in a remarkably constant
manner. The neuration has nothing remarkable about it,
and may be defined thus —
“ Primaries with vein 2 from well beyond the middle of the cell,
3 from in front of the lower angle, 4 from the angle, 5 from just
above the middle of the discocellulars, 6 from the upper angle, 7
and 8 stalked about a third from the apex, 7 being emitted from near
the end of the cell, 9 absent, 10 and 11 from the cell ; secondaries, 2
from just beyond the middle of the cell, 3 from close to the angle, 4
African species of the Lycaenesthes group of Lycaenidae. 15
from the angle, 5 from directly above the middle of the discocellu-
lars, 7 stalked from 6 from the cell.”
De Niceville describes the larva of the Eastern species
emolus, Godart, thus —
“ Larva when full fed ‘62 of an inch in length, somewhat dark
green in colour of a darker shade than most Lycaenid larva, smooth
and shining, the whole upper surface covered with minute pits to be
seen only under a strong magnifying glass. The head is very small
and retractile as usual, and of a pale green colour, the second segment
is unmarked, the third to sixth segments inclusive have some
obscure reddish-brown dorsal blotches, the three following segments
are unmarked, the tenth to twelfth have somewhat similar blotches
to those on the third to sixth, but they are more distinct and darker
in shade. There is a pale yellow lateral line just above the legs.
All the segments are irregularly and broadly pitted at the sides ;
these pits seem more or less to assume the form of a longitudinal
subdorsal depression, below which to the lateral line the colour of
the insect is slightly paler. The whole larva is much depressed,
somewhat wider than high, and seems to gradually increase in
breadth to the segment 10, the last segment is almost as broad and
rounded. The larva varies greatly in colour and markings, some
being pale green throughout and unmarked, others again are reddish
brown throughout. It feeds, in Calcutta, on Nephelium litchi.
Lamb, Cassia fistula, Linn., Heynea trijuga, Roxb., and not im¬
probably, as it feeds on so many bushes, it will eat others. Dr.
Forel identifies the ant which attends the larva as Oecophylla
smaragdina, Fab., the large red and green ant that makes immense
nests of growing leaves in trees.
“ Pupa '4 of an inch in length of the usual Lycaenid shape, the
tail pointed, the thorax slightly humped, and ending in a somewhat
sharp ridge line on the back ; it is coloured pale ochreous and bears
a prominent diamond -shaped mark posteriorly. It is smooth
throughout, reddish brown, sprinkled with minute darker dots.” —
(Butt. India, iii, p. 129.)
We have no information about the ova, and there are
one or two points in the brief life history that would be
interesting to elucidate by a more critical examination of
both larva and pupa — these gaps we hope by degrees will
be filled. Silvanus and larydas appear to be common in
many parts of Africa, so that it ought to be possible to
obtain a more detailed record of their metamorphoses ; at
present nothing is known of these details of any of the
IG Mr. G. T. H#^une-Baker’s Revision of the
Ethiopian species. I would appeal to observers in the field
to give us these records. Many Eastern species have been
described, some of which de Niceville deals with ; whilst he
admits three Indian ones he states he believes there are
only two, and it is to my mind clear that, when carefully
sifted, the number of species now standing in the Oriental
region will have to be considerably reduced. A study of
the genitalia of the section we are now considering has
been of great assistance in checking the natural groups
into which the individual species fall, and it bears out
previous experience that they are of the utmost value both
specifically and generically.
KEY TO SPECIES.
I. Lycaenesthes (.sens, strict.). Blue above, spotted below.
A. No basal spots in either wings.
1. Brown above, secondaries with broad
yellow terminal band . ruhricinctus.
2. Blue above, below spots plain almost
as ground-colour . musagetes.
3. Blue above, below spots much filled in
Avith reddish . itnria.
4. Blue above, below one red spot on
costa and one red one on inner
margin in secondaries . anadeuia.
B. Basal spots in secondaries only.
1. Deep blue above, below' spots almost
as ground-colour . ligurcs.
2. Deep blue above, below spots with
traces of red in primaries, much
red in secondaries . Iwblegi.
3. Deep blue above, postmedian stripe
broad almost confluent very slightly
fractured . indefinita.
4. Blue lilacine, below grey, no basal
spot in cell of secondaries . . . definita.
5. Mauve blue above, with fine termen in
both wings . chirirtda.
6. Dark blue above, with broad dark
apex and termen in both Avings . afm.
7. Dark blue above, Avith broad dark
apex and termen ; below spots large,
primaries Avith postmedian stripe
very oblique as to costal spots 1 to 4,
twice strongly fractured .... alberta.
African species of the Lycaencsthes group of Lycaenidac. 17
8. Dark blue above, secondaries with two
red costal and two red inner-margi¬
nal spots . lasti.
9. Mauve blue paler above, below with
three basal chestnut spots in
secondaries . lemnos.
10. Blue cupreous above, below no basal
spot in cell, costal spot bright blue smithi.
11. Cupreous lilac above, below secon¬
daries with two dark costal but
no basal spot in cell . princeps.
12. Leaden lilac above . Uganda.
13. Mauve blue above, below white ; sec¬
ondaries with no basal spot in cell,
spot at base of costa and inner
margin dark and definite . . . liodes.
14. Blue darkish above, apex broadly
brown, anal spots edged with red,
below no basal spot in cell . . . bihe.
15. Greyish blue above, apex and termen
broadly brown, below secondaries
with only one small dark basal
spot on costa . livida.
16. Blue very restricted above, below with
three small dark basal spots . . . millari.
17. Mauve blue above, below all marks
obsolete except dark basal spot
below costa . nigropunctala.
18. Brown above, pale brownish below
with pattern near to otacUia . . . minima.
19. Blue restricted to cell and below ; sec¬
ondaries below with four basal spots,
a dark one on costa, two paler in
cell, and one below . otaciliu.
20. Blue not restricted . kikuyu.
21. Brown above, underside like lunulata
without black spots . s}cquala.
22. Shining grey above, below primaries
with black basal bar below cell,
secondaries with three black costal
and four black basal or sub-basal
spots . amarah.
23. Lustrous blue above, below secondaries
with two strongly emphasised round
TRANS. ENT. SOC. LOND. 1910. — PART I. (JUNE) C
18
Mr. G. T.
une-Baker’s Revision of the
black or crimson subcostal spots
and one inner-marginal ....
24. Blue lilacine evanescent somewhat
restricted, anal spots surmounted
with orange ; below, costal and
inner-marginal spots small scarlet .
25. Blue restricted in both wings above.
In primaries below postmedian
stripe strongly fractured, costal part
very oblique ; in secondaries a
blackish subcostal spot and a
similar smaller spot on the inner
margin, no sub-basal spots in cell .
26. Deep purplish above. In primaries
below postmedian stripe irregular
but confluent ; in secondaries three
largish round sub-basal spots, post-
median stripe strongly accentuated
27. Blue dark indigo purple above. Prim¬
aries below with a large angled
basal mark, secondaries with four
largish sub-basal spots often con¬
joined, both wings much covered
with the pattern .
28. Blue pale lilacine above. Both wings
below pale with scarcely darker
spots somewhat similar in pattern
to larydas .
II. (Sens. lat.).
A. Above blue or brown, below with patches
as well as spots.
1. Both wings with blue patches, pri¬
maries with blue restricted to cell and
below ; secondaries, blue restricted
below cell interrupted by a black
bar. Below, subterminal line in pri¬
maries invaded obliquely by both
first and second portion of post¬
median stripe .
2. Blue lilacine paler somewhat tran¬
sparent above, primaries occasion¬
ally with a subdued yellow spot.
Below postmedian stripe not oblique,
subterminal line not invaded . . .
hinulata.
sanijuinea.
levis.
sihamis.
larydas.
cratvshayi.
lysicles.
lachares.
African species of the Lycaenesthes group) of Lycaenidae. 10
3. Brown above, with lustrous blue in
fold of primaries, trace of yellow in
blue below vein 2 ; secondaries
with a sub-lustrous blue stripe from
vein 2 to 5 . versatilis.
4. Brown above, secondaries with a blue
dash and an orange lunule . . . biichhohi.
5. Brown above, with orange spot in pri¬
maries and an orange transverse
dash in secondaries . bitje.
6. Brown aboA^e, with a distinct row of
terminal spots in the secondaries ;
below white, stripes disintegrated,
a row of terminal sjAots in each wing meUanJtrotiis.
7. Brown above, primaries Avith large
OA'al yellow patch, secondaries with
curved postmedian row of four
whitish blue small spots .... xanthopoecilus.
8. Brown above, primaries with a largish
spot of pale 3'ellowish below vein 2 kampala.
9. Brown above ; below Avith pattern
almost like lachares . hjchnides.
10. Brown, both wings al)Ove darkly
spotted, primaries with two obscure
yellowish spots on the fold and
slightly in the postmedian area,
secondaries Avith an obscure
yellowish shbterminal line . . . flavomaculatns.
11. BroAvn, primaries aboA^e with an
obscure orange patch beloAV the
angle of vein 3 ; secondaries Avith
a more or less j^rominent terminal
border of orange yellow tapering
finely toAvards the costa Avith black
internervular spots .
12. Brown, primaries above with veins
orange yelloAV ; secondaries Avith
broad terminal orange yelloAV
border .
B. Orange red above.
1. Primaries aboAm with a large orange
red j)atch, secondaries Avith a smaller
patch, below both Avings deep
cream, spots comparatively feAV . .
rufomdrginata.
mdiato.
locuples.
C 2
20
Mr. G. T. Bellume-Baker’s Revision of the
2. Botli wings with large red patches,
secondaries witli a prominent black
subterminal line . leptines.
3. Underside white, with orange mark in
fold of primaries . erythropoecilus.
4. Underside with large areas of orange
colour . jci-
5. Underside grey, postmedian stripes
with dark spots . malcala.
G. Eed area larger above, below with
closely placed darkish grey pattern mahota.
7. Golden orange above . aurea.
8. Golden red above, below dark grey . scintilhUa.
9. Both wings entirely golden red above,
primaries with apex broadly black,
tapering evenly off on the costa and
termen ; below yellowish orange,
primaries with upper part of post¬
median stripe very oblique invading
the subterminal dark line .... pyroplera.
1 0. Both wings entirely bright coppery red
above, secondaries witli costa and
termen very broadly black ; below
black pattern merely marked out
by rather fine white lines .... lychnuptes.
11. Both wings orange red above, pri¬
maries with very broad dark costa
and termen, secondaries very broadly
margined all round with dark
brown ; below with a large white
area in both wings at the end of the
cell . juha.
12. Primaries with a largish central round
orange patch, rest of wing black,
secondaries with an orange broad
dash in lower radial area ; below
both wings with two snow-white
patches each . zenkeri.
13. Deep orange red above, secondaries
with dark submarginal line ; below
after the pattern of jxiba .... pythagoras
African species of the Lycaenesthes group of Lycacnidae, 21
Lycaenesthes ruhricinctus, Holland.
L. ruhrieinetus, HolL, Psyche, p. 51 (1891); L. rubric
cineta, S. & K., Rhop. Exot., p. 100, PI. XXII, ff. 11, 12
(1893) ; id. Auriv., Rhop. Aethiop., p. 349 (1898).
^ . Primaries entirely uniform brown above. Secondaries uni¬
formly brown above, except for a very broad yellow terminal band
from the tornus up to vein 6.
Hob. Kangwe ; Ogowe River.
Type in the Carnegie Museum.
I know this insect only from Smith and Kirby’s figures,
and the upperside alone will serve to distinguish it from
all others in the whole group. The very broad yellow
marginal band of the secondaries is wholly unlike any
other species dealt with in this paper. It is apparently a
true Lycaenesthes (sens, strict.), and evidently belongs to
the first division, viz. species without any basal marks in
either wing.
Lycaenesthes musagetes, Holl. (Plate V, fig. 5.)
L. musagetes, Holl, Ent. News, iv, p. 25 (1898); id.
Auriv., Rhop. Aethiop., p. 349 (1898).
. Both wings above very dark indigo purple ; below uniformly
warm brown with spots no darker than the ground, or very little
darker, with no basal or sub-basal spots in either wings.
ITab. Cameroons ; Gaboon ; Sierra Leone ; Ogowe ;
French Congo ; Volta River and Angola {Mus. Brit.) ;
Rena Bendil (Congo); Makala, Beni, Mawambe
(Upper Congo); Uganda.
Type in the Carnegie Museum.
This species can easily be recognised from its allies by
the uniform brown colour, both of the ground and the
spots of the underside.
Genitalia. — Harpago very broad, the lower edge slightly curved,
the upper edge gradually rising in a strong curve to about the
middle, then tapering very slowly to the front upper apex, front edge
produced into three strong teeth, highly excavated between each, one
tooth being at the upper, one at the lower, apex, and one midway
between them. Cingula with long arms or girdle, the hind base
produced backwards into a blunt curved point, the arms of moderate
width, very gradually expanding into the cheeks of the tegumen,
22 Mr. G. T. I’^une-Baker’s Revision of the
which have an evenly curved front edge ; the dorsal apex of the
tegumen is deeply excavated in the front ; the falces are very strongly
formed, of unusual width at their sockets and well curved, ending in
a short talon ; the furca is short, with moderately broad arms, and
curved backv'ards. Penis sheath with the hinder two-thirds broad,
the front third being a somewhat curved tube of nearly even width,
slightly expanding at the orifice. The front edge of the cheeks of
the tegumen is thickly furnished with hairs, the harpagines having
but few.
Lycaenestlies anadema, H. H. Druce.
L. anadema, Druce, Trans. Ent. Soc. Loud., 1905, p. 258.
The blue of the upperside is scarcely distinguishable from its
close allies, but on the undersurface it is more easily separated ; the
costal spot of the postmedian stripe of the secondaries is red, whilst
the eighth spot, forming the lower angle of the angled spot, is like¬
wise red ; these characteristics separate it easily from its allies. At
first sight it looks very close indeed to lasti, but Druce’s species has
no basal spots in either wing, therefore it will not fall into that
section.
Hah. N. W. Rhodesia.
Type in the Druce coll.
Lyeaenesthes ituria, sp. nov. (Plates I, fig. 4 ; V, fig. 6.)
g . Upperside : both wings deep brownish purple, with fine linear
black margins ; fringes dark brown. Underside : both wings rufous
brown, with pattern scarcely darker than the ground-colour, but in
the secondaries somewhat filled in with reddish ; all the pattern has
paler edges. Primaries cell closed by a good-sized quadrate spot with
a pale central line ; an irregular postmedian band, slightly fractured
inwards at vein 4, and well fractured inwards below vein 2 ; a sub¬
terminal band from the apex to vein la. Secondaries with no spots
nearer the base than the quadrate spot closing the cell, which has a
paler middle line. The postmedian band is very irregular and
mostly filled in with red ; the second spot is shifted right outwards,
the third to fifth spot form a subconfluent series shifted right out from
2, l)ut the fifth is slightly shifted in from the fourth, sixth spot
oblique and shifted in, angled spot shifted in again, and very
strongly filled in with red ; beyond this another red spot on the
inner margin ; a trace of a subterminal band, a large, and prominent,
velvety black spot between veins 2 and 3, and a double prominent
one at the angle, all with bluish metallic scales, and edged broadly
African species of the Lycaenesthes group of Lycaenidae. 23
with bright red, this edging extending into a bright dash above the
angle.
Expanse 32-34 mm.
Hah. Beni, Makala and Mawambe (Congo Free
State), February.
Type in my collection.
I have a series of this species which is closely allied to
L. musagetes, but is readily distinguished by the orange
brown of the underside, by the markings being decidedly
reduced in size, and by the postmedian stripe of the
secondaries being largely filled in with orange red.
Genitalia. — Harpago broad, wall curved above and below, con¬
stricted slightly near the front, with upper and lower extremities
forming a strong, sharp, projecting tooth. Cingula widish, expanding
into the tegumen, which is deeply excised at its front apex, with the
cheeks gradually sloping off and not very broad. Falces strong,
curved near the middle. Furca with longish arms, curved backwards.
Penis sheath broadish, moderately even in width, with a curved,
oblique tip. Cheeks of tegumen and apices of harpagines well
supplied with long hairs.
Lycaenesthes ligures, Hew. (Plate V, fig. 7.)
L. ligures, Hew., Trans. Ent. Soc. Loud., 1874, p. 349 ; Ill.
D. Lep., p. 2'20, PI. 90, ff. 1, 2 (1878); id. Auriv., Rhop.
Aethiop., p. 849 (1898) ; L. ukerewensis, Strand, Ent.
Zeitsch, xxiii, p. 127 (1909) in parte.
Separable from its near allies by the three sub-basal spots across
the secondaries. The female, hitherto undescribed, is entirely pale
brown, with a broad, obscure, whitish terminal area in the second¬
aries, in which are two preterminal broadish lines, the outer one the
darker of the two, a prominent dark spot at the anal angle and one
above it. Below the colour is quite white, the pattern following that
of the male, but larger and broader. The Sierra Leone specimens are
very much greyer in shade and of a smoother texture than those
from Central or Eastern Equatorial Africa. In a series of the insect
thfey are recognisable at a glance.
Hah. Sierra Leone ; Ashanti ; Old Calabar ; Came-
ROONS ; Gaboon; Congo; Angola; Upper Congo
(Makala, Mawambe) ; Toro, Nandi.
Type in the British Museum.
Genitalia. — Harpago broad, similar in outline to L. musagetes, but
the front edge has but two teeth, the upper and lower extremity only
24< Mr. G. T. Dilhune -Baker’s Revision of the
being produced into a strong tooth. The cingula is also similar to
Holland’s species, hut broader and rather shorter ; the cheeks of the
tegumen and the tegumen as a whole are not developed very fully ;
the dorsal front apex is well excavated, and the cheeks are gradually
and evenly rounded on their front edge ; the falces are strongly
developed, and well though evenly curved ; the furca is not large,
the arms curved backwards being but moderately long. Penis sheath
very broadly developed for its hinder two-thirds, gradually rising to
its highest point, where it is suddenly reduced in front to a broadish
even tube for its front third. The front edge of the cheeks of the
tegumen is well furnished with hairs, the front edge of the harpagines
having a smaller supply.
Lycaenesthes hobleyi, Neave. (Plates I, fig. 9; VI, fig. 10.)
L. hobleyi, Neave, Novit. Zook, vol. xi, p. 339 (1904).
A species closely allied to the preceding one but separable at once
by the fact that nearly all the spots of the undersurface pattern are
more or less filled in with vivid red; the costa is also finely of the
same colour. The female is bluish in both wings ; primaries with
costa and termen very broadly brown, apex brown to the cell ;
secondaries with costa brown to the cell and vein 6; termen broadly
brown with a bluish interrupted dividing line ; a black spot at the
anal angle with a second above it internally edged with orange red.
Undersurface : both wings very pale greyish brown with the markings
as in the male but larger.
Hab. IMau Escarpment {Doherty).
Tring Museum ; and in my collection from the Kikuyu
District, also the Nandi country.
Type in coll. Trimen.
Genitalia. — Harpago broad, evenly curved above and below, the
curve of the upper edge being very bold ; about a third from the
front upper apex is a curious sqiiare process, unique so far as I am
aware in the genus, consisting of a strong, short tooth on each side
of the harpago, joined by a chitinous bar ; the upper and lower front
extremities are each produced into a strong tooth. Cingula with the
girdle strongly developed ; the lower extremity produced well back
into a curved hook-like projection. Tegumen raised at its hind
dorsal apex, highly excavated at its front dorsal edge, the cheeks
being unevenly rounded off. The falces are very strong and broad,
rising directly from the socket in an even curve upwards. Furca
moderately long, with narrowish arms slightly curved backwards.
Penis sheath of moderately even width throughout, narrow with the
African species of the Lycaenesthes group of Lycaenidae. 25
upper part of the tip cut off for nearly a third, and gradually taper¬
ing to the tip. Cheeks of tegumen well supplied with hairs, there
being but few on the harpagines.
Lycaenesthes indefinita, sp. nov. (Plates I, figs. 7 $-8
V. fig. 9.)
^ . Upperside : both wings deep brownish purple with linear black
margins with pale extremities. Underside : both wings brownish
grey with slightly darker pattern with pale edging. Primaries with
a large subquadrate spot closing the cell, the postmedian band con¬
sisting of seven spots, the first four confluent curved, the next two
confluent shifted inwards, the last spot shifted right in, the outer
margin being almost confluent with the inner margin of the sixth
spot ; a trace of a subterminal band. Secondaries with three sub-basal
spots, viz. one on vein 8, one in the cell, and one on the inner margin ;
a large quadrate spot closes the cell. Postmedian band with the
second spot shifted outwards, third and fourth almost confluent also
shifted out, fifth inwards, sixth inwards, angled spot inwards again ;
subterminal band more distinct, a black spot between veins 2 and 3
and at the angle, with metallic-blue scales, and edged above and
laterally with deep yellow.
Expanse 35 mm.
5 . Both wings brown. Primaries with pale bluish below the cell
and vein 3. Secondaries with the radial area up to the termen pale
bluish, the subterniinal band dividing it and the black spot between
veins 2 and 3 prominent, edged above with deep yellow. Underside :
both wings quite white, but with the pattern practically as in the ^ .
Hah. Kikuyu, 6,000 feet; Toro, February; Nairobi,
Uganda, Kikuyu, Njoro and Ruwenzori {British
Museum) ; Mau Escarpment {Tring Museum).
Types in my collection, also in the British Museum and
at Tring.
The species is slightly variable as to the fracturing of
the postmedian band of the primaries. It is more closely
allied to L. hohleyiy Neave, than to ligures, Hew., but it has
no red at all below, and the postmedian fasciae in both
wings are broad and confluent to a degree quite unattained
in either of those species.
Genitalia. — Harpago exceedingly broad, with the lower edge deeply
shouldered at a quarter from the lower apex and produced forward
into a gouge-shaped extremity, the upper edge being furnished with
two strong teeth at about a quarter from the apex. Cingula with a
short basal backward process, longish arms expanding but slightly ;
26 Mr. G. T. J »#Plune- Baker’s Revision of the
tegumen moderately developed, the dorsal front apex being well
excavated with a straightish edge on each side, slightly angled at the
true fore-apex ; the cheeks then descend in a fairly straight line ; the
falces are strong, evenly curved, ending in a very long talon ; the
furca has longish arms, not very wide and curved well backward.
Penis sheath gradually rising from the hind extremity to rather
beyond the middle, where it is very broad, then gradually tapering
to the tip, the front third being somewhat shagreened on its upper-
side. The tegumen and harpaginea are furnished with hairs.
Lycaenesthes deflnita, Butler. (Plates I, figs. 5^6^.)
L. definita, Butler, Ann, Mag. N. H., 1899, p. 342 ; id.
P. Z. S., 1900, p. 927 (1901); id. Druce, Trans. Ent. Soc.
Lend., 1905, p. 253; L. nigrocandata, Pagenstch., Jahrb.
Nassau, Iv, p. 156 (1902).
A species easily to be identified, as the blue is quite different,
much more lilacine, and decidedly paler than any of the very deep
blue species that are allied to it ; it is also very much paler on its
underside, being of a pale clear'grey, as are the markings also. The
female likewise differs, being pale blue in both] wings above, with a
prominent dark spot closing the cells, and a broad dark postmedian
foscia likewise in both wings, the termen being broadly brown ;
whilst in the secondaries there is an additional dark spot in the
middle of the cell.
Hal. Kikuyu ; Cape Colony ; Natal ; Mashonaland
(Umtali); Uganda; Fort Jameson (N. W. Rhodesia);
Nandi; Kilimanjaro; Mount Elgon; Nairobi.
Type in the British Museum.
Dr. Pagenstecher has been good enough to lend me his
type of L. nigrocaudata, and it is a specimen of the above
insect with the basal spot of the secondaries rather obsolete,
but it is quite visible in one wing under a low-power lens.
Genitalia. — Harpago almost elliptical, tapering rapidly beyond the
middle and terminating at the upper and lower extremity in a strong
tooth. Cingula produced well backwards at the lower base, with
longish arms, very narrow ; tegumen highly excavated at its front
dorsal apex with the fore edge of the cheeks very slightly curved ;
falces long, narrow, deeply curved at a quarter from the socket; furca
with longish arms, united for a fifth at the base, receding well back¬
wards. Penis sheath narrow, gradually increasing for two-thirds,
then tapering down into a trumpet-shaped orifice. Tegumen well
furnished with hairs ; harpagines but scantily supplied with short
hairs.
African species of the Lycaenesthes grcnip of Lycaenidae. 27
Lycaenesthes chirinda, sp. nov, (Plate I, fig. 10.)
$ . Upperside : both wings darkish mauve blue with the finest
possible dark linear borders to all the wings. Underside pale grey with
whitish fine edges to the spots. Primaries with spot closing the cell
largish, and a small costal one above it ; postmedian stripe near the
termen, consisting of six confluent spots, angled slightly at the fourth,
seventh spot shifted well inwards ; a subterminal row of interner-
vular broadish dashes. Secondaries with a sub-basal spot below vein
8, and a small one on the abdominal margin ; postmedian stripe sub¬
confluent, the second, third and fourth spots each shifted out rather
further than the predecessor, fifth and sixth each shifted slightly in,
seventh and eighth angled spot shifted further in ; a terminal row of
internervular lunules ; two black terminal spots with iridescent
scales and red inner margins.
5 . Uniformly brown above, except that the secondaries have a
broad white radial patch. Underside white with the postmedian
stripes in both wings rather more confluent than in the male.
Eah. Chirinda Forest, Gazaland, 3,600 feet,
October.
Types in my collection (2 pair).
This species is superficially very difficult to separate from
L. definita, the main difference being that the postmedian
stripes in both wings are decidedly nearer the termen than
in Butler’s species. The genitalia are, however, markedly
different. The female is also quite different, with no blue
and no spotted band above.
Genitalia. — Harpago subovate tapering from the middle to a blunt
apex without any dentation. Cingula produced w'ell backward at
the base with arms of moderate length and width ; tegumen
moderately developed, with a somewhat straight front edge ; falces
strong, slightly curved ; furca smallish, with arms narrow, inclined
strongly backwards. Penis sheath long, wide for the hind three-
fifths, then gradually tapering to a trumpet-shaped orifice. Tegumen
and apex of harpago furnished with longish hairs.
Lycaenesthes afra, sp. nov. (Plate I, fig. 12.)
d . Both wings deep indigo purple. Primaries with costa broadly
brown, termen broader still, and apex yet broader. Secondaries
with costa brown to the cell margin and vein 6, termen broadly
brown, two dark marginal spots [near the angle. Underside : both
wings grey, with scarcely darker markings margined with whitish.
Primaries with a spot closing the cell, having a paler central line, a
28 Mr, G. T. B^Piune-Baker’s Bevision of the
costal spot near the postmedian band, which latter consists of seven
more or less confluent spots, viz. a pair at the costa, another pair
curved and shifted outwards, fifth spot shifted inwards, sixth well
inwards, seventh right inwards, larger and curved ; an obscure scal¬
loped subterminal band. Secondaries with three quite small sub-basal
spots, viz. one on vein 8, one in the cell, and one on the abdominal
margin, a largish spot closing the cell with a pale central line ;
postmedian band consisting of eight spots, the seventh and eighth
forming the angled one, the second spot shifted outwards, third and
fourth confluent, shifted outwards, fifth inwards, sixth inwards,
oblique, angled spot right inwards ; a trace of a subterminal band,
with a scalloped whitish fine outer edging, below which is a black
spot with blue metallic scales, edged with orange colour, and a similar
smaller double spot, similarly edged at the angle.
Expanse 31 mm.
Hah. Makala, Congo Free State, July; Bitje,
Cameroons.
Type in my collection from Makala ; also in the British
Museum, and in the Druce collection from the Cameroons.
The broad black borders to both wings should suffice to
distinguish it from other species ; the underside is very
close to Sierra Leone specimens of ligures.
Lycaenesthes alberta, sp. nov. (Plate I, fig. 11.)
^ . Both wings deep brownish purple. Primaries with the costa
and termen broadly brown, the apex more broad, a black line closing
the cell. Secondaries with very broad brown costa tapering round
the termen to the angle, where it is moderately narrow. Underside :
both wings brownish with pattern edged finely with white. Primaries
with a large quadrate spot closing the cell with a pale central line,
a small costal spot above it and shifted slightly outwards. Post-
median band consisting of four confluent obliquely arranged spots,
each shifted slightly beyond its predecessor,’ spots five and six con¬
fluent shifted inwards, the sixth spot decidedly larger than the fifth,
seventh spot shifted well inwards and oblique ; a scalloped post¬
median band indefinitely edged with whitish, particularly on its
inner edge. Secondaries with three sub-basal spots, viz. one on
vein 8, one in the cell, and one on the inner margin. A large
quadrate spot closes the cell, finely dusted with whitish ; postmedian
band darkish with pale central lines ; spots two, three, and four, each
shifted slightly beyond each other, fifth spot shifted inwards, sixth
inwards, angled spot very obtuse, shifted slightly inwards ; subterminal
line obscure, scalloped, broadly and indefinitely edged internally
African species of the Lycaenesthes group of Lyeacnidac. 29
with whitish, less broadly externally ; a black spot with bluish scales
on the margin between veins 2 and 3, edged above with deep yellow,
a small double spot at the angle similarly scaled and edged. There
is a considerable amount of pale fine irrorations over a good deal of
the secondaries.
Expanse 30 mm.
Hal. Albert Edward Nyanza, October ; Entebbe,
Uganda ; Congo (Makala) ; Angola.
Type in my collection from Makala, and in the British
Museum from Entebbe (coll. Minchin).
This is exceedingly close to the preceding species above,
but below it is more easily recognisable ; the pattern is
decidedly larger, the postmedian stripe being very broad,
the fractures being different and the spots darker.
Lycaenesthes lasti, Smith and Kirby.
L. lasti, S. & K., Rhop. Exot., p. 109, PI. 24, ff. 1,
2 (1894) ; id. Auriv., Rhop. Aethiop., p. 348 (1898) ; id.
H. H. Druce, Trans. Ent. Soc. Bond., 1905, p. 253 ; L.
rubrimacidata, Strand, Ent. Zeitsch., xxiii, p. 129 (1909).
The upperside of .this species would be most nearly described as
plum colour; it is a distinct tone from other species. The underside
is pale grey with two red costal spots, and two red spots on the
abdominal margin, the upper one of which is sometimes inclined to
obsoletion. These characters will enable the insect to be easily
identified. The female is brown above in both wings ; the primaries
with a greyish white patch in the fold and the lower radial area ;
the secondaries with a similar stripe in postmedian area. The
Kambove form, as brought home by Mr. Neave, has the red spots
much reduced both in size and colour, the one at the beginning of
the postmedian line of the secondaries being often plain, not red.
Hal. Mombasa ; Chirinda Forest, Gazaland,
3,600 feet, October; Rhodesia, Kambove,
Type in coll. Grose-Smith.
Genitalia. — Harpago broad, evenly curved below, highly curved
above for its basal two-fifths, then nearly straight to the apex, which
is produced into two very strong teeth, the lower extremity being
produced into a three-toothed process. Cingula well produced
backwards at its base, with narrow arms of moderate length ;
tegumen with dorsum weakly developed, excavated in front, cheeks
very ample, with evenly curved front edge ; falces very broad
30
Mr, G. T. Be^une-Baker’s Revision of the
curved from the socket ; furca small, short in the arms, and narrow,
inclined well backwards. Penis sheath almost elliptical for its hinder
half, then tapering forwards, suddenly reduced near its tip, but
expanding equally suddenly into a trumpet-shaped orifice. Tegumen
and harpagines fairly well supplied with hairs.
Lycaenesthes lemnos, Hew. (Plate VI, fig. 11.)
L. lemnos, Hew., Ill. D. Lep., p. 221, PI. 90, ff. 13, 14
(1878); id. Holland, Proc. U. S. Nat. Mus., p. 241 (1895) ;
id. Auriv,, Rhop. Aethiop., p. 347 (1898) ; L. sylvanus, Hew.,
Ill. D. Lep., p. 222, PI. 92, f. 41 (1878); id. Trimen, S, A,
Butt., ii, p. 98 (1887); id. Staud., Exot. Schm., i, Taf. 94
(1887); emolus, Gerst. Deckens, Reise iii, p. 378, Taf. 15,
f. 4 (1878); natalensis, Stand., Exot. Scfim., i,p. 273 (1888);
id. Butler, P, Z. S., 1900, p. 927 ; ukereivensis Strand,
Ent. Zeitsch., xxiii, p. 127 (1909).
The undersides of the secondaries contain the best points of
difference between this and other species ; tliere are three sub-basal
almost round spots of a distinctly brown tone different from the
other markings, varying from chestnut colour to very dark brown.
No doubt it was a specimen -with this latter colour that made Trimen
confuse it with sylvanus, from which it is abundantly distinct. The
postmedian fasciae in both wings are very broad.
This is the species that was first confused with emolus,
Godart, and secondly with sylvanus, Drury. Now that we
have more material in our hands it has needed but little
to difierentiate it from either of the species mentioned ;
the ground-colour below is quite different from sylvanus, as
also is the pattern.
Ilab. Natal ; Delagoa Bay ; Mombasa ; Uganda ;
Upper Congo.
Type in the British Museum.
frenffaZm.— -Harpago broad, suboval with the lower extremity
forming a long, strong, double-toothed process, the upper edge being
very highly dentate for fully a tliird to the apex some of the teeth
being very highly developed. The cingula is well produced back¬
wards at its base, the arms being of moderate length and wfidth ; the
tegumen has very ample cheeks ; the front dorsal apex is deeply
excavated, rounded off into the cheeks, which have a fairly straight
front edge; the falces are long, of moderate widtli; the furca is long
A frican species of the Lycaenesthes group of Lycaenidae. 81
with narrow arms united for a third from the base. Penis sheath
very broad for its hinder three-fifths, then slowly tapering to a
blunt oblique tip. Tegumen well furnished with long, strong hairs,
harpagines with hairs shorter and finer.
Lycaenesthes smithi, Mabille. (Plate I, fig. 13.)
L. smithi, Mab., An. Eut. Fr. (5) 7, Bull. 72 (1877);
id. Hist. Madag. Lep., i, p. 213, PI. 27, ff. 9-11 (1887);
? lochias. Hew., Ill. D. Lep., p. 221, PI. 90, tf. 7, 8 (1878); id.
Auriv., Rhop. Aethiop., p. 348 (1898).
A very rare species, of a cupreous lilac in both wings with very
fine dark margins. The undersurface is pale brownish grey wdth
broad white margins to the spots. In the secondaries there is a
blue sub-basal spot below vein 8 white-edged, whilst the black
iridescent spot between veins 2 and 3 is edged internally with
crimson, a most unusual colour in this genus, orange and orange
red being almost universal.
Hah. Madagascar.
I am indebted to M. Mabille for the loan of this type,
which is in his collection. I have little doubt that it is a
local form of L. princeps, Butler,
Lycaenesthes princeps, Butler. (Plates I, fig. 17 ;
VII, fig. 15.)
L. princeps, Butler, Ann. Mag. N. H., 1876, p. 484; id.
Auriv., Rhop. Aethiop., p. 349 (1898) ; L. neglecta. Trimen,
Trans. Ent. Soc. Lend., 1891, p. 175 ; id. id. p. 132, PI. 8,
ff. 7, 8 (1893) ; id. Auriv., Rhop. Aethiop., p. 348 (1898).
cj. Both wings a delicate cupreous lilac colour with very fine
dark terminal lines ; the underside is a delicate shade of grey with
a cupreous lustre in parts in certain lights, the distinguishing
feature being two subcostal dark spots in the secondaries, the one
sub-basal below vein 8, and the second forming first spot of the
postmedian line ; there is also present a similar dark spot on the
abdominal margin ; this, however, is liable to obsoletion, and I
have one specimen in which it is totally absent. The female is
greyish brown with a very restricted bluish base in both wings,
the underside presenting no difference whatever to that of the
male.
32
Mr. G. T. B^iune- Baker’s Revision of the
Butler’s ^ type in tlie British Museum does not belong
to this species ; there is no question at all that it is the
$ of lunulata, Trimen.
Hah. Abyssinia; Niger; Somaliland; Natal; Zulu-
LAND; Zomba; Salisbury; Sierra Leone; Gold Coast
{Dudgeon) ; Delagoa Bay {Monteiro).
Type in the British Museum.
Genitalia. — Harpago roughly oval with the upper fore ex¬
tremity produced into a serrated point, llie front edge erect, slightly
excavated. Cingula well developed with the lower hind extremity
produced far backwards, the sides rapidly expanding upwards
towards the tegumen which is wholly excised as to its apex, the
neck being quite narrow at the back top of the cingula ; the falces
are slightly curved at the tip, of only modeiate length and curved
sharply near their socket ; the furca has its main stem long and
strongly curved forwards, the diverging arms then curve back and
are long and narrow ; the penis sheath is long and large for its
hindmost half being suddenly reduced in width, the fore half having
an open boat-shaped structure with a somewhat flat constricted tip ;
the tegumen is abundantly furnished with long, strong bristles,
those on the apex of the clasp being only few in number.
Lycaenesthes nganda, sp. nov.
Both wings upperside greyish lilac with white fringes.
Primaries with very fine linear black costa and slightly wider
termen. Secondaries with fine black termen. Underside ; both
wings pale ashy grey with darker ashy-grey markings prominently
edged with pure white. Pattern similar to L. princeps, Btl., but
larger and broader, with a wdiite band in both wings between the
postmedian and subterminal bands.
Expanse 31 mm.
Hah. Entebbe, Uganda, 3 specimens {Minchin).
Type in British Museum.
This is near to princess, Butler, and may prove to be
a local form of it; but the difference in colour is quite
marked, and there are also equally perceptible differences
of the underside pattern.
Lycaenesthes liodes, Hew. (Plate I, fig. 14.)
L. liodes, Hew., Trans. Ent. Soc. Lond., 1874, p. 349 ; id.
Trimen, S. Afr. Butt., ii, p. 100 (1887) ; id. Auriv., Rhop.
Aethiop., p. 349 (1898) ; id. H. H. Druce, Trans. Ent. Soc.
Lond., 1905, p. 253 ; eviolus, Triineu, Rhop. Afr. Aust.,
African species of the Lyeacnesthcs group of Lycaenidae. 33
p. 234, PI. 4, £f. 8, 9 (1866); id. Mab., Hist. Mad. Lep.,
p. 202, Pi. 27, ff. 1, 2 (1887) ; sichela, Hew., Ill. D. Lep.,
p. 222 (1878); id. Trimen, S. Afr. Butt., ii, p. 101 (1887);
adhcrhal, Mab., Bull. Zool. Soc. Fr., p. 217 (1877); id.
Butler, P. Z. S., 1893, p. 660 (1894) ; id. id. p. 120 (1896);
id. Druce, Trans. Ent. Soc. Lond., 1905, p. 253; id. Auriv.,
Arkiv. Zool., ii, p. 16 (1905) ; Monteironis, Kirby, Hew,, Ill.
D. Lep., p. 223 (1878); id. Druce, Trans. Ent. Soc. Lond.,
1905, p. 253.
Upperside : both wings slightly lustrous violaceous, with
tennen finely dark. Secondaries with a dark terminal spot between
veins 2 and 3. Undersurface whitish grey, almost dirty white, with
slightly darker spots of the usual pattern. Primaries with the post¬
median stripe fractured outw'ards below the second spot, inwards
below the fourth, and again inwards below the sixth spot. The
distinguishing point of the secondaries is the presence of two small
round dark grey sub-basal spots, one below vein 8 and one on the
abdominal margin. The female vai’ies considerably as to the extent
of blue on the upperside ; in some specimens there is only a trace
of pale blue scales in the fold of the primaries, and scattered scales
in the cell of the secondaries ; in others, the blue is quite marked
and occasionally there is a considerable area over both wings ; the
undersides, however, all agree and follow the male pattern entirely.
Expanse ^ 30, $ 28-34 mm.
Hak Gaboon; Calabar; Angola; Midodoni (B. E.
Africa), British Museum \ Sierra Leone; Northern
Nigeria, Bassa Prov. {Coll. Bctlmne- Baker) ; Cape
Colony; Kaffirland; Natal; Transvaal; Landana.
Type in the British Museum.
There has been considerable difficulty in elucidating
the various species, but the advent of a pair of specimens
of both sexes from Nigeria, together with the loan of
Mabille’s type, at once showed that the synonymy as given
above was quite correct. There should be little difficulty
in identifying the species in the future. It is widely
spread, but either very local in its distribution or very
retiring in its habits. It is interesting to note that a ^
and ^ in my collection of this species have glabrous eyes,
one ^ has eyes slightly hairy, and in another ^ the eyes
are more hairy. None of the specimens are in bad
condition, so that it is a reasonable deduction that the
eyes are in their natural condition.
TRANS. ENT. SOC. LOND. 1910. — PART 1. (JUNE) D
34
Mr. G. T. BdiTuue -Baker’s Revision of the
Lycaenesthes hike, sp, nov. (Plate I, fig. 15.)
Both wings brightish purple. Primaries with broad brown
costa and broader apex tapering very rapidly to about vein 5, from
whence the termen is quite fine. Secondaries with costa broadly
brown, termen linear, a black spot on the margin between veins 2
and 3, and another at the angle broadly edged above with orange.
Underside ; both wings pale grey with darker spots edged with white.
Primaries with a spot closing the cell with whitish central line ;
postmedian band consisting of seven small spots, two below the
costa, two shifted outwards, the fifth spot oblique right inwards,
sixth oblique also right inwards, seventh slightly curved far inwards
and isolated ; subterminal line definite, scalloped. Secondaries
with two basal sjjots, one small one almost tawny on vein 8 and one
blackish one on the inner margin ; a large quadrate spot closes the
cell witli a pale central line ; postmedian band very irregular in
pairs, a pair on the costa, a pair shifted right out detached, a pair
right inwards, the lower of which is further in, fourth pair forming
the angled spot right inwards and detached ; subterminal line fairly
distinct, a good deal of whitish between it and the postmedian line,
small black spot on the margin between veins 2 and 3, with a few
metallic scales margined with bright orange red, a similar minute
spot at the angle, wdth a largish round orange-red spot above it, over
which also are some metallic scales.
Expanse 31 mm.
Hah. Bihe District, Angola.
Type in my collection.
This is a well-marked species allied to liodes, Hew., but
recognisable by its darker and more purple colour, by the
red spots at the anal angle of the upperside, and by the
sub-basal spot on the undersurface of the secondaries being
red.
Lycaenesthes livida, Trimen. (Plate VI, fig. 12.)
L. livida, Trimen, Trans. Ent. Soc. Bond., 1881, p. 443 ;
id: S. Afr. Butt., p. 103, PI. 7, ff. 7-7a (1887); id. Auriv.,
Rhop. Aethiop., p. 348 (1898); L. hutlcri, Auriv., Rhop.
Exot., p. 348 (1898); Lycaena hutleri, Oberth., An. Mus.
Genov., p. 170, PI. I, f. 2 (1880).
^ . Both wings blui.sh grey. Primaries with costa broadly brown,
termen more broadly, and apex yet more so ; a dark dash closes the
cell. Secondaries, rather bluer than the primaries, with a sub¬
terminal scalloped line, the usual black anal spots bordered inter-
African species of the Lycaencsthes group of Lycaenidae. 35
nally with yellow ; a dark spot closes the cell. The underside may
be recognised at once by the fact that there is only one sub-basal
spot, a small dark round one below vein 8.
A large female from the Kikuyu District, agreeing
entirely Avith Oberthiir’s description and figure of hutUri,
differs somewhat from Trimen’s description on the upper-
side. The blue is brighter and the area more definite,
being confined to the cell, the fold and the lower
radial area below vein 3; there is a dark spot in the angle
of vein 2. In the secondaries the blue is as in the
primaries but diffused, with a short postmedian row of
confluent spots from vein 2 to 6. The underside is quite
typical, paler than the male.
Hah. Cape Colony ; Natal (Mulkolomba Mountains) ;
Kikuyu District {Coll. Bethune- Baker) ; Abyssinia.
Genitalia. — The harpago is very full with rounded outline above
and below, the fore part of the upper edge being strongly and evenly
excised ; the cingula of moderate development, somewhat produced
backwards at the base, developing more heavily towards the tegumen,
which is large and copious, well excised at apex in the front, less so
at the back ; the falces are long and well curved with the tip slightly
hooked ; the furca is longish with narrow arms and rising from near
the middle of the harpagines ; the penis sheath is long, highly
enlarged for two-thirds of its length in the middle with a trumpet-
shaped orifice.
Lyeaencsthes millari, Trimen. (Plate VI, fig. 13.)
L. millari, Trimen, Trans. Ent. Soc. Lend., 1893, p. 133,
PI. 8, f. 9 ; id. Auriv., Rhop. Aetliiop., p. 348 (1898).
^ . On the uppersurface this species is exceedingly close to
Uvula, Trim., but the blue is paler, less extensive, and more distinct ;
in its undersurface it can be at once recognised by the fact that it
has on the secondaries a series of three or four dark sub-basal small
spots, one below vein 8, one or two in the cell, and one on the
abdominal margin. This character is a prominent feature.
Hah, Natal (Tugela River).
Genitalia. — The harpago is a broad oblong with the upper front
edge rounded off to the lower extremity ; the cingula is broad, well
developed, the basal extremity being produced backwards in a long
horn ; the tegumen very ample in the hood, whose front edges are
nearly straight, moderately excised at the apex, scarcely at all at
1) 2
m
Mr. G. T. BcTTiune-Baker’s Revision of the
tlie back ; the falces are strong, deeply curved, almost at a right angle ;
the furca is short with strong arms ; penis sheath long, straight,
tapering gradually from the broad back extremity to the tip, w'hich
is bluntly pointed.
Lycaencsthes nigrojpunctata, sp. nov. (Plate I, fig. IG.)
Both w’ings darkish mauve blue with fine linear black
margins. Secondaries wdth two distinct black spots (one at the
angle and one between veins 2 and 3). Underside : both wings clear
French grey with the spots finely edged with greyish white.
Primaries with a small costal spot above the spot closing the cell ;
postmedian stripe with the costal four spots confluent, curved and
compressed, fifth spot shifted inwards, sixth inwards again as to its
inner margin only, seventh spot yet further in ; subterininal line
fairly distinct, consisting of a series of six internervular scallops.
Secondaries with a defined black sub-basal spot below vein 8 ; post¬
median stripe with the first costal spot darkish, second spot shifted
out, third and fourth confluent compressed shifted out, fifth com¬
pressed shifted inwards, sixth slightly further in, seventh and
eighth angled spot still further in ; subterminal line less defined,
scalloped ; anal sirot and that between veins 1 and 2 black with
metallic-blue scales, edged above with bright red.
$ . Entirely brown above. Below like the male, but the ground¬
colour whitish ash grey.
Expanse 27-29 mm.
Hal. Fort Jameson, N. E. Rhodesia; Lakes Bang-
WEOLO (4,200 feet) and Tanganyika (4,500 feet), N. W.
Rhodesia, June, July and August. (A. S. Neave.)
Type in my collection.
Lycaenesthes minima, Trimen.
L. minima, Trimen, Trans. Ent. Soc. Lond., 1893, p.
135, PI. 8, f 10 ; tf?. Auriv., Rhop. Aethiop., p. 347 (1898).
Trimen describes this species as brown with a slight
cupreous gloss ; the underside is very close to otacilia.
Trimen, the sub-basal series of spots of the secondaries
having an additional one on the abdominal fold. The
species, however, should be readily separated from otacilia,
for it is not blue at all on the upperside.
Hal. Natal ; Mashonaland ; Ndola District, N. E.
Rhodesia ; Ndzovuni (B.E.A.).
The specimens from Rhodesia show no trace of a
cunreous gloss.
A O
African species of the Lycacncsthcs group of Lycaenidae. 37
Lycaenesthes otacilia, Trimen. (Plates II, fig. 1 ;
VI, fig. 14.)
L. otacilia, Trimen, Trans. Eut. Soc. Lend., 1868, p. 90 ;
id. S. Afr. Butt., p. 102, PL 7, f. 8 (1887); id. Aiiriv.,
Rhop. Aethiop., p. 348 (1898) ; L. didcis, Pagenstch., Jahrb.
Nassau, Iv, p. 156 (1902).
d. With the blue of a dullish, sub-lustrous violaceous shade,
confined in the primaries to the cell, the fold, and below vein 3 in
the radial area ; in the secondaries the blue is evenly di.stributed
with a broad brown termen. The undersurface is warm greyish
brown with the spots closing the cells in both wings, and the post¬
median fasciae broadish and prominent ; the secondaries have a
prominent small dark round sub-basal spot below vein 8, another
paler one in the cell, and a third just below the cell.
I have a female which is entirely brown, without any
blue colour at all. It is a small, well-marked species,
widely distributed. Dr. Pagenstecher’s L. dulcis, the type
of which is before me, is only a small specimen of it.
Ilcch. Cape Colony ; Kaffirland ; Natal ; Zambesi ;
Dar es Salaam and Ruwenzoki (^n7. Mus.); Njoro and
Uganda (Coll. Bethune- Baker).
Genitalia. — The harp)ago is somewhat lozenge-shaped, with the
front termination divided into three deeply excised points or fingers.
The cingula is strongly developed, with the lower basal hind
extremity well produced backwards and strong, the sides approach¬
ing the tegumen broad ; the tegumen is well developed, very
deeply excised at the front apex, with the hind apex practically
level with the cingula ; the falces are long and broadish, very broad
for the size of the insect, well curved and tapering gradiially to the
extremity ; the furca is rather long with arms of moderate width.
The penis sheath is almost boat-shaped for nearly half its length ;
a short depression occurs after this point, but it rises again, and
then tapers to the front tip rapidly, the upper portion being more
or less open — it is unusually stout.
Lycaenesthes otacilia kikuyu, var. nov.
(Plate II, fig. 2.)
(J. Differs from otacilia. Trimen, in that all the wing is blue,
leaving only the apex of the primaries broadly brown tapering
rapidly along the termen to the torniis, whilst the secondaries have
also a larger blue area practically up to the termen, except just at
38
Mr. G. T, Betliune-Baker’s Bcvuion
tlie apex. Uiulersnrface : the colour is much paler than in the
parent form with the markings blending closely into the ground¬
colour, and the basal dark spots much less pronounced.
9. Strongly marked with iridescent blue in both wings. Under¬
surface as in the male.
JTah. Kikuyu District, January, 5,000 to 6,000 feet.
'J'ypes in my collection.
The female of otacilia is very rare, but those I have
seen are all entirely brown, one has the least trace of blue
scales at the extreme base of the secondaries.
Lycaencsthes stiqualct, Pagenstecher.
L. suquala, Pagenstch., Jalirb. Nassau, Iv, p. 155 (1902).
9 . Dark lustrous brown with termen blackish and white fringes.
Secondaries with an anal black spot bordered with yellowish red.
Underside : both wings with four dark brown whitish bordered
spotted lines with two black dots on the fore edge of the secondaries
and the usual spots in the anal area.
Jfah. N. E. Africa.
Type in Dr. Pagenstecher’s collection.
Dr. Pagenstecher, who has been good enough to lend me
his type of this species, describes it as near L. adherlal,
Mab. (i. e. liodes, Hew.). The specimen, possibly owing to
rough usage, is now in very poor condition, so that the
scales are mostly more or less absent, but there is a trace
of blue scaling on the uppersurface, whilst the underside
pattern, just discernible, is exceedingly close to L. otacilia,
Trimen ; but there is no trace whatever of two black spots
at the fore edge (“am Vorderrande ”) as described. Dr.
Pagenstecher, however, describes his species from a female,
whilst the specimen before me should be a male, judging
from the shape of the Avings ; the legs are unfortunately
absent. It may therefore be that I have not the original
type before me. The species under these circumstances
must be left in this somewhat uncertain condition.
Lycaenestlics amctrah, Guerin. (Plate VII, fig. 16.)
A. amarah, Guerin, Lefeb., Voy. Abyss., vi, p. 384, PI. 11,
ff. 5, 6 (1847) ; id. Wallengrn., Khop. Caffr., pj. 40 (1857);
id. Trimen, Rhop. Afr. Aust., p. 235 (1866) ; id. id. S.
Afr. Butt., ]). 94 (1887) and p. 411 (1889); 7c?. Anri v.,
Rhop. Aethiop.,}!. 347 (1898) ; id. id. Arkiv. Zool., ii, p. 12
(1905) ; olympusa, Walker, Entomologist, p. 53 (1870).
African species of the Lycaenesthes group of Lycaenidae. 39
There are two forms of this species, one apparently ob¬
taining in the hot arid parts of the south and one in other
districts ; the difference in colour is quite marked. The
former has both wings of a pale bright shining cupreous
grey with the red-edged black anal spots in marked con¬
trast, the underside is also slightly paler ; the other form
is dark greyish brown with scarcely any gloss, and the anal
spots much less marked — a natural result of the increased
tone of colour. In the underside this species stands quite
alone, as in addition to the usual Lycaenesthes pattern, it
has in the primaries a broad blackish bar in the fold ; in
the secondaries there are four bold blackish brown spots
below vein 8, the basal one being small, two similar sub-
basal spots, one in the cell, one below- it, and two small
ones on the abdominal margin.
It has been a matter of considerable difficulty to assign
the species its position ; the genitalia are nearest to L.
princeps, but the other characters seem to bring it more into
the present position, and on the whole I am inclined to leave
it here. It is very widely distributed over the continent,
the dark form prevailing throughout Eastern and Central
Equatorial Africa, throughout the whole of the West Coast ;
the pale form only occurring in the hot dry regions, so far
as my material enables me to judge ; the palest forms of
all coming from Bomba and Muscat, with the ^ almost
the same colour as the I have not, however, examined
a large number of specimens from the latter district.
Hah. Sierra Leone ; Angola ; Ovamboland ; Cape
Colony; Kaffraria; Natal; Transvaal; Mashuna;
British East Africa; German East Africa; Abys¬
sinia; Congo; White Nile; Bomba; Aden; Muscat;
Poona.
GenitnUa. — Harpago somewhat oval, slightly excised above and
below to the front edge, which is erect and slightly excised, the
bottom front extremity thus forms a small acute angle, whilst the
upper extremity is produced forward and upward into a long point;
the cingula is fairly developed, the lower extremity produced back¬
wards into a long, strong, curved process, the sides rapidly increasing
in width towards the tegumen, which is wholly excised on the apex ;
the neck being quite narrow and slightly elevated at the very back,
the fore edge bearing the falces being evenly rounded ; the falces are
large and strong, highly curved from the sockets with a formidable
hook at the extremity; the furca has an erect, tapering, short main
40
Mr. G. T. une-Baker’s Revision of the
stem with the diverging arms suddenly increasing in width forwards
and forming roughly an isosceles triangle. Penis sheath of a long and
irregular shape, the hind two-thirds broad, suddenly constricted, the
fore part being narrow with an elliptical orifice and a pointed
extremity.
Lycaenesthes Innulata, Trimen. (Plates II, fig. 4, ^ 5 $ ;
VII, fig. 17.)
L. lunulata. Trimen, P. Z. S., 1894, p. 51, PI. VI, f. 12 ;
L. princeps Butler, Ann. Mag. N. H., 1876, p. 484 (m
parte) ; L. otacilia, Hew., Ill. D. Lep., p. 228, PL 92, ff.
85-87 (1878); L. hewitsoni, Auriv., Rhop. Aetliiop., p. 347
(1898) ; L. litlias Smith and Kirby, Rhop. Exot., p. 110
{in parte), PI. 24, ff‘. 5, 6 (1894); L. grosei, Auriv., Rhop.
Aethiop., p. 848 (1898).
^ . The upperside of both wings is of a beautiful sub-lustrous
lilacine violet ; the primaries having a broad brown costa, a*broader
brown apex tapering rapidly to the tornus, the eye spot between
veins 2 and 3 is prominently edged internally with orange red ;
below, the prominent character is the two largish costal spots of the
secondaries below vein 8, these are sometimes black and sometimes
red in both sexes. Mr. Cator who took the species commonly in
Sierra Leone noticed the difference when capturing them, the red
spotted ones sometimes had the colour a brilliant blood red. The
female is like the male, but the blue of the upperside is brighter and
less extended.
Hob. Sierra Leone; Nigeria (Bassa Prov.); Came-
ROONs ; Angola ; Congo Free State ; Masiionaland ;
Gazaland ; Rhodesia ; L. Nyassa ; Mombasa ; Patigo
Kissegneis (Albert Edward Nyanza) ; Mazoe (Mashoiia-
land).
This species appears to have been very misunderstood,
and considering the very marked underside, it is some¬
what surprising. It is very distinct from the little L.
otacilia, Trimen, with which Hewitson confused it. Auri-
villius at once recognised its diversity, and his quick eye also
saw that Grose-Smith’s $ of lithas did not belong to that
species, hence his two new names ; but he did not succeed
in recognising Trimen’s figure, considering it to be a male
of Mabille’s adherhal, which had been described from a
female. I have been able to see most of the types, ami
African species of the Lycaencsthes group of Lycaenidae. 41
have no doubt as to the correctness of the synonymy of
the species.
Genitalia. — Harpago with its njiper part elongated into a somewhat
fan-shaped extremity with the outer and lower inner edges strongly
dentate, below this the front edge has a deep circular excavation
with the lower extremity terminating in a point. Cingula fairly
broad M’ith the hinder base produced broadly backwards ; girdle
expanding but little to the tegumen, which is of quite moderate
dimensions, with the front dorsal apex well excavated ; cheeks evenly
rounded off ; falces strong, highly curved at a third from their
sockets ; furca rising in two arms almost directly from the base
with a small patch of short strong bristles at the back at the base of
the arms, and a larger patch of longer bristles rather higher in the
front ; arms broad and inclined backwards. Penis sheath long and
narrow, waved with a slight expansion for the hinder half ; cheeks of
tegumen with long hairs, as also the extremities of the harpagines.
Lycacncsthcs sanguincct, sp. nov. (Plates II, fig. 3 ;
VII, fig. 18.)
. Both wings delicate lilac blue, somewhat fuller in tone when
held up directly facing the light, extending to the costa in the
primaries, but only to about vein 7 in the secondaries. Primaries
with the apex broadly brown tapering rapidly to the termen, and
much narrower from vein 3 to the tornus. Secondaries with a
black internervnlar marginal spot between 2 and 3 edged
above broadly with red. Underside : both wings very pale buff grey
with spots no darker than the ground-colour, faintly and finely
edged with whitish. Primaries with an obscure spot closing the
cell; posterior stripe consisting of three spots slightly curved below
the costa, two spots shifted right in so that their outer margin
touches tlie inner margin of the upjier three, sixth spot oblique
shifted right in precisely as the two middle spots were; a subterniinal
row of internervular dashes. Secondaries, with two quite small
spots below the costa almost scarlet with white irides, a sub-basal
usual-coloured spot on the abdominal margin. Cell closed by an
obscure spot ; posterior stripe with the first spot below vein 7 and
the second and third each shifted outwards beyond its predecessor,
fourth and fifth each shifted inwards in like manner, seventh angled
spot again inwards; a subterminal row of lunules, a black termina
spot as on the upperside but with metallic-blue scales and edged
above with orange red, a small similar spot at the angle,
$ . With the blue darker and brighter and only extending partly
42 Mr. G. T. PJ^mne-Baker’s Revision of the
into the cell of the primaries and below vein 3. Secondaries
with blue below vein 4, a marked series of terminal spots en¬
circled with whitish, the black spot edged with red as in the male.
Undersurface as in male.
Expanse $ 30, $ 29 mm.
Hah. Bihe District ; Angola ; N. E. Rhodesia.
Types in my collection.
This species I at first thought to be a local form of L.
lunulata, Trimen, but the advent of more specimens which
are very constant in their markings makes me more than
doubtful of it. The area and the tone of blue differ, the
latter very considerably, though this is less important than
the former ; the pattern and general appearance of the
undersurface is markedly diverse, and could be separated
at a glance from the very pronounced underside of
Trimen’s species, of which I have a good series, and of
which a very large number have passed through my
hands. It is certainly close to lunnlata^ and apparently
replaces that species in the Bihe District. Mr. Neave
thinks this may be the dry-season form of L. hinulata, but
my specimens from Angola show a distinct modification
of the genitalia ; the harpagines are not excavated to
nearly so great an extent ; the falces are decidedly more
slender, and the penis sheath is broader at the back and
decidedly more slender in the front. The genitalia com¬
pared are fully as large as each other, which points to the
modifications mentioned being more than merely indi¬
vidual differences.
Lycaenesthes levis, Hew. (Plate VII, fig. 19.)
L. levis, Hew., Ill. D. Lep., p. 221, PI. 91, ff. 21, 22
(1878) ; id. Smith and Kirby, Khop. Exot.,p. 118, PI. XXIV,
ff. 11-13 (1894) ; id. Auriv., Rhop. Aethiop., p. 348 (1898).
^ . Upper wings bright sub-lustrous mauve blue ; primaries with
blue restricted to cell and fold, secondaries to about vein 5 to 6,
termen very broadly brown with a fine scalloped interrupting sub¬
terminal line. Underside brownish grey with pattern large for the
size of the insect. In the primaries the postraedian stripe is com¬
posed of four pair of confluent spots, the second pair projected well
outwards, third pair inwards touching the cell spot, fourth pair well
inwards to beloAv and rather beyond the cell spot. The same pattern
obtains in the secondaries, and there is a prominent round sub-basal
Af rican species of the Lycaenesthes group of Lycaenidac. 43
subcostal spot, the ocellated spots at the anal angle have iridescent
scales and are edged internally with orange.
Hob. Gaboon ; Ogowe Kiver; Sierra Leone ; N’dola
District, N. E. Rhodesia (Neave).
Genitalia. — Harjjago roughly oblong with its front extremity
extended into two strong, long tusks, the upper one being the longest
and strongest ; the cingula is well developed with the hind lower
extremity projected backwards to a moderate length, the sides
developing well as they approach the tegumen, which is deeply
excised in the front, with the falces long and strongly arched and
having a hook at their extremities; the furca has its arms bent back¬
wards, of moderate length and width ; the penis sheath is long and
straight, the hind half being broad, tapering very rapidly in the
middle and narrowing to half its width or less, the orifice being tongue¬
shaped, with apparently a small opening ; the cheeks of the tegumen
are clothed with strong hairs, and the extremity of the harpagines
have also a few scattered hairs which are long and fine.
Lycaenesthes silvanus, Drury. (Plate VIII, fig. 20.)
L. silvanus, Drury, Ill. Exot. Ins., ii, p. 5, PI. 8, ff. 2, 3
(1773); id. Auriv., Ent. Ticl., xvi, p. 215 (1895); id. id,.
lihop. Aethiop., p. 347 (1898); $ moncus, Fab., Spec. Ins.,
ii, p. 113 (1781) ; id. Butler, Cat. Fab. Lep., p. 188, PI. II,
f. 10 (1870); silvanus, Herbst., Naturs, Schm., xi, p. 49,
PL 303, ff. 5, 6 (1804); syllidus, Hiibner, Verz., p. 76
(1826) ; larydas var., Godart, Enc. Meth., ix, p. 619 (1823) ;
$ locra, Plotz, S. E. Z., xli, p. 203 (1890); Pseudodypsas
silvanus, Staud., Exot. Schm., i, PI. 94, ii, p. 273 (1888).
^ . Both wings intensely dark purple blue with very fine dark
termen. Below both wings pale greyish brown, with the radial
area whitish ; the postmedian stripes very confluent, rather irregular,
very as to its outer edge in the secondaries, standing out very promi¬
nently on the pale ground ; a series of three round almost black sub-
basal spots, one below vein 8, one in the cell, and one on the
abdominal margin.
This is probably the largest of this genus, and its
strongly-marked underside should enable it to be recog¬
nised at once. On the West Coast it is common in many
places, but I have been unable to trace it into Central,
South or Eastern Africa.
41*
Mr. G. T. BiWunc-Baker’s Bevision of the
Hah. SiEiiiiA Leone; Volta River; Nigeria
(Northern and Southern) ; Togo ; Cameroons ; Gaboon ;
Lower and Upper Gongo; Angola.
Genitalia. — Harpago very broad, strongly convex on upper and
lower edges, specially so on the former, gradually tapering down to
the front edge, which is composed of two blunt teeth, the lower one
larger than the upper ; the upper edge is also heavily dentate for its
outer half. Cingula broadish, produced far back at its base. Tegumen
highly excised at its front apex, with broad cheeks. Falces broad,
curved evenly almost from their sockets. Furca rising from the
middle of the harpagines, arms long, broadish, well curved back¬
wards. Penis sheath broad, expanding highly at a fifth from the
base, suddenly contracted at a third from the semispherical top.
Cheeks of tegumen and harpagines with numerous hairs.
Lycaenesthes larydas, Cramer. (Plate VIII, fig. 21.)
L. larydas, Cramer, Pap. Exot., iii, p. 160, PI. 282, H.
(1780); id. Herbst.,Naturs. Schm., x, p. 287, PI. 290, f. 1
(1800); id. Godart, Enc. Meth., ix, p. 619 (1823); id.
Hew., Ill. D. Lep., p. 222, PI. 92, f. 40 (1878); id. Tri¬
men, S. Afr. Butt., ii, p. 96 (1887); id. Karscli, B. E. Z.,
p. 229 (1893); id. Auriv., Ptliop. Aethiop., p. 349 (1898);
Pseudodypsas larydas, Stand., Exot. Schm., ii, p. 273
(1888) ; 'pcrichs. Fab., Ent. S3'st., 3, i, p. 273 (1793); id.
Donov., Ins. India, PL 42, f. 4 (1800) ; kerstem, Gerst.,
Archiv. Naturg., 37, i, p. 359 (1871) ; id. Deckens, Reise
iii, p. 373, PI. 15, f. 5 (1873); id. Butler, P. Z. S., 1894,
p. 568.
. Both wings above very deep indigo blue. Underside dark brown
almost entirely covered with the pattern. Primaries with a broad
angled sub-basal double spot confluent at the lower margin of the
cell, above this and above the spot closing the cell several small
costal spots ; postmedian stripe broken up into three oblique parts,
the second part shifted inwards and the final lowest spot inwards
again. Secondaries with four or five sub-basal spots below each
other, generally touching each other ; spot closing cell large ; post¬
median stripe broad, more or less confluent, but irregular in outline.
These points are the main characters of differentiation
of the underside. Kersteni, Gerst., has the same pat¬
tern, but very broadly edged with wliite and with white in
the ground, which applies to the female as well, though by
African species of the Lycaencstlics group of Lycaeniclae, 45
no means to so great an extent, as the female of laryclas
(sens, strict.) has much more white below than the male.
In the male the two forms are easily separable, but they
fly together, and the genitalia are the same.
Hah. Sierra Leone ; Assinie ; Ashanti ; Togo ;
Nigeria; Old Calabar; Cameroons; Congo; An¬
gola; Natal ; Kaffraria ; Delagoa Bay; AIanica-
LAND; N. W. Rhodesia; Makala (Upper Congo);
Albert Edward Nyanza; Toro, Uganda; British
East Africa; German East Africa; Uganda;
Ruwenzori.
This species is probably the most widely distributed of
all, and the commonest. There should be no difficulty in
determining it at a glance, as, with the exception of L.
crawshayi — a smaller and much more delicate-looking
species — there is nothing quite like its underside pattern.
Genitalia. — Harpago shortisli with basal half broad gradually
tapering to a blunt fore extremity (ham-shaped), from the main con¬
striction a long, strong, horn-like tusk is emitted on the upperside
and (.‘xtends forward to the apex. Cingula shortish, strongly
developed, of moderate width, extended backwards at its lower
extremity into a short, strong process at right angles with the main
sides. Tegumen very specialised, highly excised in the front apex,
slightly so at the back with the neck raised into a high saddle ; the
cheeks are broad, extended well forward, terminating in a gradually
tapering blunt point above and below. The falces are moderately
short, strong and curved forward nearly from their origin. The
furca has rapidly-expanding arms, slightly excised on the hinder
eJge for the upper half, and are nearly erect. The penis sheath is
short for the size of the insect, very broad for its hinder half, sud¬
denly constricted and narrow for its apical portion, the orifice
ending in a tapered point. The cheeks of the tegumen are well
furnished with long, strong hairs, and the apex of the harpagines
has also a few hairs distributed at the end and its lower portion.
Lycaenesthes craiushayi, Butler. (Plates II, fig. 6 ;
VIII, fig. 22.)
L. craivshayi, Butler, Ann. Mag. N. H., 1899, p. 342.
. Upperside blue in both wings, sub-lustrous lilacine, with ter-
men finely dark brown. Secondaries with a dark spot at the anal
angle and another between veins 2 and 3. Underside pale greyish
4G Mr. G. T. Bet^l^ie-Baker’s Revision of the
brown with markings very similar to those in larydas, Cram., but
smaller.
9 . Dull grey with a bluish scaling at the extreme base of both
wings. Secondaries with a terminal row of small spots in a lighter
ground. The colour of the upperside varies somewhat ; in some it
is a full-toned lilacine, in others very pale, and in others, viz. the
type, somewhat cupreous. In some specimens the dark border is
decidedly wide, but generally it is only linear.
The species can be separated easily from L. larydas,
Cram., its nearest ally, by the colour of the upperside and
by the pale colour of the underside.
Hah. Sierra Leone; N. Nigeria; Nyassa;
Uganda; Nandi; Ruwenzori ; Lake Njoro ; Kili¬
manjaro.
Type ill the British Museum.
Genitalia. — Harpago very broad, roughly oval for its basal two-
thirds, erect the apical third, spread out into a hood whose
two front edges are strongly serrated, the lowest tooth being the
largest, below which is a deep circular excavation, the lower edge of
which is produced into a long, blunt, horn-like process. The cingula
is heavily developed, broad from its origin, tapering gradually and
more widely to the tegumen ; the basal hind extremity is produced
backwards into an egg-shaped cavity. Tegumen very highly exca¬
vated at its front apex and slightly at its back, the narrow neck
rising into a high, saddle-like process ; the cheeks are evenly
rounded in front, broad and well developed, with the falces long and
strong, highly curved some little distance up from their sockets ; the
furca is curved backwards with the arms expanding broadly for half
their length, then diminishing to a point. Penis sheath very broad
for the basal three-quarters, then suddenly reduced with a some¬
what trumpet-shaped orifice. Both the cheeks of the tegumen and
the end of the clasps are furnished with longish fine hairs.
Lycaenesthes lysicles, Hew. (Plates II, fig. 7 ; VIII, fig. 23.)
L. lysicles, Hew., Trans. Ent. Soc., Loud., 1874, p. 348 ;
Ill. D. Lep., p. 224, PI. 91, ff. 15, 16 (1878) ; id. Auriv.,
Khop. Aethiop., p. 350 (1898).
. Both wings blackish, with the cell and below vein 4 brilliant
lustrous purple blue ; in the secondaries there is a broad black dash
in the lower radial area. The underside of both wings is dark
brown with the pattern marked out by broad white lines. Primaries
with a large wedge-sliaped mark, a broken broad stripe at the end of
African species of the Lycaenesthes group of Lycaenidae. 4-7
the cell all across the wing ; postniedian stripe divided into two
parts, the costal portion oblique, and the lower part less oblique but
shifted right inwards, a double terminal line, the inner one being
broad. Secondaries, with a broad, irregular basal dash, a spot closing
the cell ; postmedian stripe very irregular, the first spot on the costa
large dark, second small and paler, third and fourth confluent
shifted outwards, fifth and sixth inwards and smaller, seventh and
eighth angled spot dark and shifted right inwards ; a strongly-curved
subterminal line, an anal black terminal spot with a second between
veins 2 and 3, with iridescent scales and the latter edged with
yellow’.
5 . Brown above. Below like the male, but the white is pre¬
dominant.
This species forms the first of the second division of
the genus. In structure it is a typical Lycaenesthes, the geni¬
talia are allied to the lemnos group ; but the pattern is
quite diverse, and is so different and so homogeneous with
others that follow that they form a well-marked sub¬
division so far as the pattern of the underside is concerned.
There is, however, no structural point of separation.
Hah. Sierra Leone ; Gaboon ; Old Calabar ;
Cameroons.
Type in the British Museum.
Genitalia. — Harpago roughly ovate, wdth a deep tooth about three-
quarters along the low’er edge, two Avell-developed deep on its front
extremity, and another on its upper edge near the tip. Cingula
well developed but narrow’, with the lower extremity produced back¬
wards into a strong lobe. Tegumen excavated on the apex in front
with the cheeks, having almost a straight front edge ; the falces being
very strongly developed highly curved near the sockets, and with a
strong hook at ’the tip ; the furca expands widely from the base,
suddenly contracts narrowly at a half, and bends backwards, the
penis sheath is small comparatively, of moderate width for its basal
half, gradually tapering to the apex, which is slightly bulbous, having
a shortly excavated extremity. Both tegumen and harpagines are
supplied with long, fine bristles.
Lycaenesthes lachares. Hew. (Plates II, fig. 9 $ ;
VIII, fig. 24.)
L. lachares, Hew., HI. D. Lep., p., 225, PI. 91, IT. 33, 34
(1878) ; ? id. Auriv., Eut. Tid., 1895, p. 216; id. Auriv.,
Ptliop. Aethiop., p. 350 (1898); L. liparis. Smith, Novit.
48
Mr. G. T. 1 *4^une-Baker’s Bcvision of the
Zool., V, p. 357 (1898); id. Aiiriv., Rhop. Aetliiop., p. 350
(1898); L. indclier, Smith and Kirby, Rhop. Exot., ]>. 105,
PI. 53, ff. 9, 10 (1893) ; id. Auriv., Rhop. Aethiop.,
p. 350 (1898); L. liparis, G.-Smith, Rhop. Exot., p. 138,
PI. XXIX, ff. 4, 5 (1901).
Both wings above a delicate sub-histrous violet colour with
broad brown margins ; the primaries with a largish subdued ochreous
patch about the end of the cell. Underside very similar to lysides,
Hew. ; the postmedian stripe of the primaries being moderately
confluent with spots three and four shifted outwards on their outer
margin.
9 . Both wings dark brown. Primaries with an ochreous yellow
.spot in the angle of vein 2 and extending well beyond. Secondaries
with a yellow dash in the lower radial area just over the yellow line
bordering the anal sjjots. Underside : both wings white with the
pattern of the male carried out, but in a somewhat vestigial manner.
o . Var. obsolescens, var. nov. (Plate II, fig. 8.)
Similar to the type form, but entii-ely without the ochreous patch
in the primaries.
Smith and Kirby were the first to describe the ^ of this
species under the name oi puhher (/. c.), and I have, there¬
fore, taken that as the type form. The unicolorous violet
form flies with the type species, and I have no doubt it is
only a variety of it.
Hewitson described it in the first instance from a 9- I
have a good series of both sexes from several parts of West
Africa, and have no doubt whatever that Smith and Kirby’s
species is the ^ of lacliares, Hew. I have, through the
kindness of the Tring Museum, also had the type of liparis,
S. & K., and it in all respects agrees with this species. It is
widely distributed in West Africa, but I can find no record
from other localities.
Hal. SiEREA Leone ; Old Calabar ; Ashanti ; Benin ;
Warri; N. Nigeria; Gaboon.
Type 9 in the British Museum.
Genitalia. — Harpago rouglily oblong willi the front upper apex
raised into a blunt cone, front edge straight, serrated, ending in a
sharp tooth, excavated shortly below in a curve, then receding in a
straightish shoulder to the lower edge, which is slightly hollowed at
this point. Cingula well developed with narrow sides, the lower
basal extremity produced backwards into a long, curved, blunt pro¬
cess, the sides expand rapidly at about halfwaj" into the tegumen,
A frican species of the Lycaencsihes group of Lycaenidae. 49
which is deeply excavated at front apex and less so at the back, the
narrow neck being slightly raised ; the cheeks are evenly rounded
off to their lower angle ; the falces being very long and strong, curved
sharply well above their sockets, and with the tip developed into a
sharp, strong hook ; the furca is very long, curved well back, with
arms long and narrow. Penis sheath very wide for its hinder half,
gradually tapering narrowly to the tip, whose orifice is excavated into
a hollow point. The cheeks of the teguinen are furnished with fine
hairs, and the upper apex has also a few shorter ones sparingly placed
down the front edge.
Lycaencsthes versatilis, sp. nov. (Plate II, fig. 10.)
^ . Both wings blackish bi’own. Primaries with the area below
the cell and between veins 1 and 2 dull lustrous purplish blue to a
quarter from the termen ; below vein 2 there is a trace of a yellow
dash showing through the blue. Secondaries with a short, dull,
lustrous blue stripe across the wing in the postmedian area from
vein 2 to 5. Underside white with greyish-brown markings.
Primaries with a basal subcostal dash, below which is a broad dark
mark along the inner edge, angled suddenly upwards at vein 2 ; cell
closed by a large spot, with another below it below vein 2 ; post¬
median stripe further out than usual, slightly curved, shifted out at
vein 6 ; subterminal line broad, touching the postmedian stripe at
vein 2. Secondaries with a broadish basal dash, having a dot below it
on the inner margin ; cell closed by a largish spot; postmedian stripe
so broken up as not to appear in sequence at all, viz. costal spot
very dark, with an indefinite mark below’ it ; a dark, confluent spot
between veins 4 and 6, shifted well out, follow’ed by a less dark spot
between veins 2 and 4 shifted iirwards ; angled spot below this very
dark ; subterminal line broadish ; a black terminal spot between
veins 2 and 3, and another at the angle, both with metallic scales
and margined with reddish.
Expanse 27 mm.
Hab. Gaboon.
Type in the National Collection.
I have described this underside as white ; it is no doubt
brown, almost entirely covered with white superimposed
scales, and there are therefore, strictly speaking, no white
lines around the spots. The species is in markings nearest
to Neurypexina lamproclcs, Hew., but the neuration is
different, and it will therefore, with L. huchholzi, Karsch,
hold a somewhat isolated position in this genus.
TRANS. ENT. SOC. BOND. 1910. — PART I. (.JUNE) E
50
Mr. G. T. l^Wume-Baker’s Revision of the
Lycaenestlies hLchholzi, Plotz.
L. htchholzi, Plotz, S. E. Z., p. 202 (1880) ; id. Auriv.,
Rhop. Aethiop., p. 350 (1898).
. Upperside : both wings uniform bluish black. Underside
similar to larydas, Cram., but with the base and the terminal areas
white, and tlie postmedian stripe oblique in the primaries.
$• Upperside : both wings blackish browm. Primaries with an
ovate orange spot. Secondaries with a fine blue stripe in the inter-
nervular spaces of veins 1 to 3, and between veins 2 and 3 a small
orange spot.
Hah. Cameroons (Victoria).
Type ill coll. Erliardt, Muiiicli.
I have not been able to see this species, but it is
evidently near iny versatilis.
Lycaenesthes hitje, Druce.
Mr. Druce is describing this species from the Cameroons ;
it is closely allied to L. versatilis, but has yellovv-oi'ange
spots not blue, and the underside also differs slightly.
Hah. Bitje; Ja River, Cameroons.
Type in the Druce collection.
Lycaenesthes nielamhrotus, Holland. (Plate II, fig. 11.)
Tj. melamhrotus, HolL, Ent. News, p. 27 (1893); L.
melamhrota, Auriv., Rhop. Aethiop., p. 350 (1898).
5 . Both wings brown on the upperside ; the secondaries with a
marked row of dark marginal spots with an outer white line bordering
them. Underside very similar to an obsoletely-inarked female
lacliares, with a terminal row of black-pupilled white spots in both
wings. I am greatly indebted to Dr. Holland for a coloured figure
of this species ; his figures of other species known to me are so
accurate that I have no hesitation in reproducing this figure. At
the same time it apj^ears to be probably a sport. The whole appear¬
ance of the insect leads me to this conclusion, and I should not be at
all surjtrised to find, if its genitalia could be dissected, that it w'as a
(jynandromorph.
Hah. Ogowe River.
Type in the Carnegie Museum.
African species of the Lycaenesthes group of Lycaenidae. nl
Lycaenesthes xanthopoccihis, Holland.
L. xanthopoecilus, Holl., Ent. News, p. 27 (1893) ; L.
xanthopoecila, Auriv., Khop. Aethiop., p. 352 (1898) ;
L. lycotas, Smith, Novit. Zool., p. 356 (1898) ; id. id.
Rhop. Exot., p. 137, PI. XXIX, ff. 1-3 ; id. Auriv., Rhop.
Aethiop., p. 350 (1898) ; L. leuhokescha, Karsch, Ent.
Nach., p. 295 (1895) ; id. Auriv., Rhop. Aethiop., p. 352
(1898).
d . Both wings brown above. Primaries with a large orange-
yellow nearly-round patch in the lower radial area. Secondaries
with a row of blue spots in the ^Jostmedian area from vein 1 to 5,
and a fine bluish subterminal line interrupted by the black spot
between veins 1 and 2. Underside white with dark spots, no scat¬
tered markings in the median area of the primaries from the sub-
basal wedge-shaped mark to the postmedian transverse stripe,
except a single mark at the end of the cell and one below it, between
veins 1 and 2 ; tranverse stripe confluent curved to vein 2, where it
nearly touches the thickened end of the subterminal line, wdiich is
broadish and evenly curved. Secondaries with a curved basal
costal line, a broad sub-basal Imnd ; a largish indefinite spot closes
the cell ; postmedian stripe broken into four pairs of spots, tlie costal
upper spot being large and dark, the lower spot paler and smaller,
third and fourth spots dark shifted right out, fifth and sixth small
shifted right inwards, seventh and eighth angled, isolated and
shifted far inwards ; subterminal line evenly curved, edging inter¬
nally the anal black-jjupilled yellow spots which have iridescent
scales.
9 . Like the male above, except that the yellow spot in the
primaries is paler, and the row' of blue spots in the secondaries is
replaced by a band of yellow'. Underside like the male, but the
markings reduced.
Hab. Ogovve River ; Bopoto ; Mukenga ; Congo
Free State.
Type (a ^) in the Carnegie Museum.
I am fortunate in having an excellent figure of Holland’s
and also of Karsch’s species, together with the Tring type,
and they all agree one with another ; Karsch’s .species is
smaller than the others, but that is all the difference.
This species is very similar to the $ of lachares, especially
to those that have much development of yellow on the
upperside of the wings ; the underside, however, differs
E 2
52
Mr. G. T. Bf^nume-Baker’s Brvision of the
decidedly in the paucity of markings in this species. Both
species seem to be very constant in the pattern of the
underside.
Lyeaencslhes locuples, G. -Smith.
L. locwpUs, Grose-Smith, Novit. Zook, p. 357 (1898); id.
Auriv., Rhoi). Aetliiop., p. 352 (1898) ; id. G. -Smith,
Rhop. Exot., p. 138, PI. XXIX, ff. G, 7 (1901).
d- Botli wings dark brown. Primaries with a very large deep
orange-red patcli occupying the median and radial areas. Second¬
aries with a large orange-red spot at the end of the cell and in the
radial area. Underside very deep cream with an orange tinge, the
pattern of both wings similar to xanthojwecilKs, Holh, birt with fewer
marks in the secondaries ; in the primaries the postmedian stripe is
curved, confluent, and not so near the termen; in the secondaries
the postmedian stripe is present only in vestigial remains, the seventh
and eighth angled spots being the only ones really well developed in
the usual way; the subterminal line is very evenly .curved like
Holland’s species but broader.
JIab. Warri, Niger District.
Type in the Tring Museum.
This is a very distinct species, and should be recognised
without any difficulty.
Lycacnesthcs kampala, sp. nov. (Plate II, fig. 12.)
^ . Both wings very dark brown. Primaries with a small chrome
yellow ovate patch between veins la and 2 in the radial area of the
wing. Secondaries with the white inner edge of the subterminal
line on the undei’surface showing through, a small black terminal
spot between veins 2 and 3, obscurely edged finely with yellow, a
trace of two more smaller black dots between this and the angle,
a preterminal fine white line interrupted by the veins. Undenside :
both wings dark sooty grey with the pattern marked out by broad
lines of pure white, the enclosed spots being only occasionally darker
than the ground-colour. Primaries with a sub-basal wedge-shaped
spot followed liy an irregular H-shaped mark across the outer part
of the cell, a broad dash from vein 5 to 2, from whence it descends
to la in a fine line ; beyond this from the costa is a broad-angled dash
to below vein 3, angled between 5 and 4, and succeeded by an outer
narrower dash from the costa to vein 5 ; a double subterminal line
rather irregular, the outer one being decidedly narrower than the
inner of the two. Secondaries with a broad l)asal dash from the lower
Af rican species of the Lycaenesthes grovp of Lycaenidae. 53
margin of the cell to the costa, curved at the costa ; sub-basal dash
broad, suddenly contracted about vein la ; cell closed by a narrow
spot ; postmedian stripe with the first spot large and dark, second
much smaller and not so dark, third and fourth confluent dark shifted
right outwards, fifth and sixth confluent darkish shifted inwards,
seventh and eighth angled shifted right inwards ; subterminal line
strongly marked followed by the least trace of a preterminal
interrupted line, a black terminal spot between veins 2 and 3, and at
the angle with metallic-bluish scales, the former also edged finely
with yellowish.
Expanse 28 mm.
Hah. Kampala ; Uganda, April.
Type in my collection.
Lycaenesthes lychnides, Hew. (Plate II, fig. 13.)
Ij. lyehnidcs, Hew., Ill. D. Lep., p. 224, PI. 91, f. 32, PI.
92, f. 38 (1878); id. Auriv., Rhop. Aethioj), p. 351 (1898).
^ . Upperside uniformly brown in both wings. Underside similar
to L. lachares. Hew.
? • Upperside uniformly brown above in both w'ings. Primaries
with a yellow dash between veins 1 and 2.
This species has not been recorded again since the
original description ; it is near to lacharcs, but its male
may be recognised by being brown above, not blue ; whilst
the female has much less white on its underside than is
usual in the female of lacharcs.
Hah. Old CalabaPv.
Type in the British Museum.
Lijcaenesthes fiavomaculatus, Smith and Kirbj''.
(Plate IX, fig. 25.)
Tj. fiavomaetdatus, S. & K., Rhop. Exot., p. 104, PI.
XXIII, ff. 7, 8 (1893) ; flavoniaculata, Auriv., Rhop.
Aethiop., p. 351 (1898).
$ . Upperside : both wings darkish brown with darker spots and
dirty yellowish-creamy small patches. Primaries with a spot closing
the cell ; postmedian stripe broken up into three nearly detached
parts, viz. a very oblique costal dash, two spots below it shifted
right inwards, and edged outwardly by a dirty deep creamy patch,
the third part being a single spot shifted right in and edged on
54 Mr. G. T. Betlnme- Baker’s Revision of the
each side l)y a deep creamy patcli, that on the outside being tlie
larger of the two. Secondaries with a spot closing the cell ; post-
median stripe very irregular, the spots small ; outside this a deep
yellowish-cream line almost angled at vein 5, a terminal row of
small spots decreasing towards the apex and edged internally by a
fine whitish line. Underside white with the usual markings of this
section of the group — the distinguishing feature being the exces¬
sively oblique costal part of the postmedian stripe of the primaries.
Hob. West Afeica; Bassa Prov. (N. Nigeria)
Bihe District, Angola.
Type in the British Museum.
The species is quite different from others of this genus ;
it is superficially nearest to Neurypexina lyzanius, Hew.,
but the neuration is that of a true Lycaenesthes.
Genitalia. — Harpago subovate, with the fore edge developing two
teeth, the upper one formed by a slight excision of the front line
and not extruded, the lower one at the apex longish. Cingula well
developed with a deep curve at the back of the girdle, making the
tegumen very narrow ; it (the tegumen) is deejjly excised with the
cheeks tapering off narrowly into an acute angle at their lower
extremity. Falces much exposed owing to the narrowness of the
cheeks of the tegumen, moderately strong and well curved at about
a third from their sockets. Furca short, almost erect with short
narrow arms. Penis sheath small, widest beyond the middle, i. e.
near the top, where it tapers down very rapidly to the oblique
extremity. Cheeks of tegumen with an abundance of long hairs,
harpagines with a more scanty supply.
This species is with some difficulty separable from
many specimens of Neurypexina lyzanius, Hew., if the
colour and pattern only are considered ; the genitalia, how¬
ever are decidedly different, so that with the neuration
differing also, we are abundantly justified in placing Hewit-
son’s species in another genus.
Lycaenesthes rufomarginata, sp. nov. (Plates II, fig. 14;
IX, fig. 26.)
(t. Botli wings uniform dark brown. Primaries markless.
Secondaries with three marginal spots edged all round, except on
the termen, with orange yellow, one between veins 3 and 4, one
between 2 and 3, and a twin spot at the angle. Underside ; both
wings pale greyish, with the pattern marked out by white lines not
darker than the ground-colour except in one or two places which
African species of the Lycaenesthes group of Lycaenidae. 55
will be specially mentioned. Primaries with a long, slightly wedge-
shaped sub-basal mark, followed by an irregular H-shaped mark
across the end of the cell ; postinedian stripe irregular, spots 1 and
2 small confluent just below the costa, 3 and 4 confluent dark
shifted well outwards, 5 and 6 confluent with dark externals shifted
right inwards, 7 confluent with 6 but no dark patch in it ; sub¬
terminal line somewhat scalloped. Secondaries with a long, irregular
sub-basal band, a spot closing the cell ; postmedian stripe not con¬
tinuous, first spot dark below vein 8, second obsolete, third and
fourth dark confluent shifted right outwards, fifth very small,
somewhat obsolete, confluent with the sixth, which is darker,
eighth and ninth angled and dark shifted far inwards ; subterminal
line farther from the termen than usual ; outside the terminal
pupilled spots, one of which lies between veins 2 and 3 and is black
with metallic-bluish scales edged with yellow, and a smaller similar
one at the angle.
9. Like the male, but with a very obscure tawny patch in the
median area of the primaries.
Expanse ^ 31, $ 29 mm.
Hob. Makala (Congo Free State), June, and Lake
Albert Edward, March.
Types in my collection, a short series.
In one ^ specimen there is a distinct small orange tawny
patch about the angle of veins 2 to 4.
Genitalia. — Harpago somewhat oblong with the upper edge
hollowed but rising to a head at the front apex, front edge serrated,
the teeth being short but strong, lower edge slightly hollowed near
apex, then recurved somewhat boldly. Cingula well developed,
■widening rapidly towards the tegumen ; the lower basal extremity
produced backwards into a long, broad, straight, horn-like process.
Tegumen with the fore apex deeply excised, the hind apex slightly
so ; cheeks with a straight front edge. Falces long, strong, highly
curved directly above their sockets, with talon-like extremities.
Furca with very long, narrow arms inclined backwards. Penis
sheath long, the hinder half hollowed, somewhat boat-shaped, rising
to a broad centre from whence it is gradually reduced into a
narrowish tube with a trumpet-shaped orifice. Both the cheeks of
the tegumen and the front edge of the harpagines are furnished with
longish hairs.
Lycaenesthes radiata, sp. nov. (Plate II, fig. 15.)
(J. Both wings brown, with yellow markings. Primaries with
veins la, 2, 3, 4, 6 and 7, and the lower margin of the cell, more or
56 Mr. G. T. I#! )une-Baker’s Eevidon of the
less completely marked out with yellow ; a marginal small patch of
yellow between veins 2 and 3. Secondaries with the termen yellow
from the apex, rapidly increasing towards the tornus ; a terminal
blackish spot between veins 2 and 3, and a minute one between
3 and 4. Underside : both wings greyish brown, with the pattern
more or less dark, edged broadly with white. Primaries with a large
wedge-shaped sub-basal mark, a squarish spot closing the cell ; post¬
median stripe most irregular, composed of three pairs of confluent
spots and a seventh ; the first pair on the costa small, second pair
dark, twice as large, and shifted well outwards, third pair dark,
smaller, shifted right inwards, seventh spot waved and oblique
again shifted right inwards ; an even subterminal stripe. Secon¬
daries with a broad basal band widest in the centre, a squarish .spot
closing the cell ; postmedian stripe composed of four pairs of spots —
first pair on the costa with upper spot blackish, second pair blackish
shifted well out, third j)air dark shifted well inwards, fourth pair
angled, dark, shifted well inwards again ; subterminal stripe
strongly arched ; a dark terminal spot between veins 2 and 3 and at
the angle overlaid with metallic blue and edged with orange
yellow.
Expanse 34 mm.
Hob. Sierra Leone, July.
Type in my collection.
Genitalia. — Very closely allied to L. rufomarginata ; the harpago
is slightly hollowed above and below beyond the middle, rising in
each case to the apex, which is curved and strongly serrated. Cingula
fairly developed, the basal backward process being broadly trian¬
gular ; the tegunien is well excavated at its fore apex only, and the
cheeks have a curved front line ; the falces being curved at nearly
right angles just above their sockets very strong, with talon-like
extremities ; the furca is very long with very narrow arms curved
backwards. Penis, with the hinder half somewhat hollowed and
apparently open above (probably this is only apparent), rising i;p-
wards towards the centre, where it reduces rapidly to the tip, the
orifice having a taper tip. The cheeks of the tegunien and the front
edge of the harpago is well furnished with longish hairs, those of
the harpagines being fine.
Lycaenesthe^ leptines, Hew. (Plates II, fig. 17 ; X, fig. 29.)
L. Ie2')tines, Hew., Trans. Ent. Soc., Lend., 1874, p. 348 ;
id. Ill. D. Lep., p. 226, PI. 91, ff. 23, 24 (1878) ; id. Auriv.,
Rhop. Aethiop., p. 352 (1898).
African species of the Lycacncsthcs group of Lycacnidae. 57
. Both wings with large orange patches above. Primaries with
broad borders to the costa and terinen, a very broad dark apex and
narrow dark inner margin. Secondaries wdth costa very broadly
dai’k, extending over the base to two-thirds of the cell ; abdominal
margin dark ; a dark defined subterminal line with dark spots at the
angle; cell closed by a dark .spot. Underside greyish with the
usual pattern, but little darker than the ground-colour.
Hah. Gaboon; Congo; Beni; Mawambe (Upper
Congo).
Type in the British Museum.
Genitalia.- — Harpago, rounded at first on its upper and lower edge,
at a quarter from the front ajiex it suddenly rises into a bluntly
shouldered extremity, the lower edge is equally suddenly produced
outwards into a broad large horn which is strongly excavated on its
front edge and is then developed into a .sharp tooth whence the
fore edge which is strongly serrated rises erectly to the upper apex.
Cingula moderately wide, expanding somewhat into the tegumen,
w’hich is well excavated at its front dorsum with the cheeks nearly
straight but gradually increasing to their lower extremity where
they are broadest. Falces strong highly curved at a third from the
sockets. Furca with longish arms inclined backwards. Penis sheath
broadest beyond the middle, the apical third being broad and with
a tapered extremity. Tegumen and harpagines with stiff fine hairs.
Lycacncsthcs crythropoccilns, Holland. (Plate III, fig. 1.)
L. crythropoecilus, Holl., Ent. News, p. 26 (1893);
L. crythropoccila, Auriv., Rhop. Aethiop., p. 352 (1898).
^ . Upperside almost exactly like L. leptines, Hew. ; the primarie.«,
however, have a dark subterminal line in the brown area, and the
secondaries have a fine preterminal line in addition to the sub¬
terminal one. Underside : both wings pure white, with the usual
pattern, the distinguishing characters being in the primaries ; the
postmedian line is confluent, slightly curved and not fractured until
the lowest spot, which is shifted right in and surrounded with
orange scales ; the postmedian stripe is followed by three finer dark
lines. Secondaries with the postmedian stripe much disintegrated,
the seventh and eighth angled spots shifted far in below the spot
closing the cell and separated from it by some orange scaling.
Hah. Ogowe River.
Type in the Carnegie Museum.
This species though near Icjdines, Hew., is evidently dis¬
tinct, and should he recognised at once by its distinctive
underside.
58
Mr. G. T. BGliune-Baker’s Revision of the
Lycaenesthes ja, Druce.
]\Ir. Druce is describing this species from the Ja River
in the Cameroons ; it is very near to L. erythropoecihis,
Holland, but the underside is almost entirely bright
orange, and the pattern also differs.
Hah. Bitje; Ja River, Cameroons.
Type in the Druce collection.
Lycaenesthes niahala, sp. nov. (Plates II, fig. 16 ; IX,
figs. 27-28.)
^ . Both wings blackish brown, with pale orange-yellow areas.
Primaries with a large pale orange-yellow patch occupying nearly
all the cell to the inner margin and nearly all the radial area, this
area being defined outwardly in a bold curve; termen narrowly
brown. Secondaries with the wing beyond the cell orange from
vein 1 to 6 up to the termen. Underside : both wings dull grey,
not pale, with more or less darker markings edged with creamy
white. Primaries with a darkish long wedge-shaped sub-basal
mark ; a large spot closing the cell ; postmedian stripe with two
small confluent pale spots on the costa followed by a pair of large
dark spots projecting well outwards, third pair smaller shifted well
inwards, seventh spot erect shifted right inwards ; subterminal line
somewhat scalloped increasing in width to the tornus, between this
and the termen is a second narrower line. Secondaries with a basal
band of nearly even width ; a spot closing the cell ; a postmedian
stripe of four pair of spots, the first pair on the costa with the upper
spot large and dark, second pair shifted right out with the outer
half of the spots dark, third pair smaller shifted well in, fourth pair
angled sliifted far in, isolated ; a row of subterminal lunular marks ;
a black spot between veins 2 and 3, and a small double one at the
angle each overlaid with bluish-metallic scales and edged with
orange yellow.
Expanse 32 mm.
Hah. Beni and Makala; Congo Free State,
January — April.
Type in my collection.
This species is near leptincs. Hew., but the yellow
area is much larger and the underside different in many
particulars.
Genitalia. — Harpago broad with the lower edge nearly straight,
the upper edge strongly curved and gradually rising to the upper
African species of the Lycaenesthes group of Lycaenidae. 59
extremity, where it suddenly ascends into a high collar, front edge
strongly serrated, the lower extremity being deeply but shortly
excavated and forming two strong long teeth ; cingula narrow with
the lower extremity produced backwards into a broad process, the
arms expand but little imtil they fuse with the tegumen, which is
very deeply excavated at the front apex only, the cheeks being
gradually rounded off in the front ; the falces are long and strong,
curved at right angles at near their middle, and terminating as usual
in a talon-like hook ; the furca has a short base with long, narrow
arms receding backwards ; the penis sheath is twice cxirved, the hind
extremity being blunt and narrow, slightly angled at a little distance
along when it suddenly expands and is deeply curved, and hollowed
on its upper edge, at about three-fifths it is again curved and gradu¬
ally tapers in an even tube, to the orifice, having a short tapered
extremity. The cheeks of the tegumen and the fore edge of the
harpagines are sparingly furnished with fine, long hairs.
Lycaenesthes mahota, G. -Smith.
Z. mahota, G.-Smith, Ann. Mag. N. H., 1887, p. 65 ; id.
Trimen, S. Afr. Butt., iii, p. 390 (1889) ; id. Monteiro,
llelagoa Bay, Frontisp. f. 8 (1891) ; id. Smith and Kirby,
Bhop. Exot., p. 99, Pi. XXII, ff. 7, 8 (1893) ; id. Auriv.,
Rhop. Exot., p. 352 (1898).
d. Both wings with large deep orange patches and broad dark
borders. Primaries with costa and termen broadly dark brown, the
orange patch not rounded at the apex but of the same shape as the
wing only smaller. Underside : both wings greyish white with the
usual pattern of this section. The postmedian stripe is fractured,
the third and fourth spots confluent and shifted out, fifth and sixth
each shifted slightly in, seventh right inwards ; subterminal line
broadish and interrupted at each vein. Secondaries with the spots
of the postmedian stripe isolated and very dark.
JIah. Delagoa Bay; Cameroons.
A species very near to Z. erythropoecihis, Holl., but
the shape of the orange patch of the primaries above is
different, and the postmedian stripes in both wings are
different, as already described.
Lycaenesthes aurea, sp. nov. (Plate III, fig. 2.)
d. Both wings deep golden yellow (not red). Primaries with
costa and termen finely black. Secondaries with the base and costa
broadly black, abdominal fold broadly black decreasing towards the
60
Mr. G. T. Bi^unc-Baker’s Revision of the
angle, termen finely black, tlie least trace of a dusky subterniinal
line, more defined at the a2)ex and the angle. Underside : both
wings with all the dark markings broadly edged with deep cream
colour. Primaries with a dark wedge-shaped basal j^atch, a largish
dark spot closing the cell, with another below it below vein 2 ; post¬
median stripe with the first three scoots oblique, fourth and fifth
shifted well inwards ; snbterminal stripe broad. Secondaries with
a broad unbroken basal band ; a large spot closing the cell ; post¬
median stripe with the second and third sjiots shifted outwards,
fourth, fifth and sixth shifted well inwards and vertical, seventh and
eighth angled and shifted right inwards; subterminal line broad,
strongly curved, black terminal spots between veins 2 and 3 and at
the angle with metallic scales and margined with red.
Ex^^anse 28 mm.
Hah. Begoro ; Goi.d Coast (Dudgeon).
Type in the National Collection.
Lycaenesthes scintilkda, Holland. (Plates III, fig. 3 : X,
figs. 30-31.)
L. scintillida, Holl., Psyche, p. 50 (1891) ; id. Ent.
News, p. 26 (1893) ; id. Smith and Kirby, Rhop. Exot.,
p. 98, PI. XXII, ff. 3 and 4 (nee 5 and 6) (1893) ; id. Auriv.,
Rhop. Aethiop., p. 352 (1898).
6 • Ujri^erside. Primaries brilliant lustrorrs coppery orange, base
very restricted black, costa linear black, aj^ex broader black, termen
very narrowly black. Secondaries as primaries, but costa base and
abdominal fold broadly deep black, an isolated black terminal sjjot
between veins 2 and 3. Underside darkish brownish grey with
darker sj)ots with cream-coloured edges. The 2)Ostmedian stripe is
composed of three sections, the first to the fourth spot confluent
gradually incre§sing in width and oblique, fifth and sixth erect
shifted right in on to the inner edge of the fourth, the seventh large
shifted right in on to the inner edge of the sixth. Subterminal line
broadish almost broken at vein 4 and again at vein 2. Secondaries
with the postmedian stripe composed of four pair of spots, all very
dark except the fifth and sixth; the second pair is shifted well out¬
wards and is erect, the third pair right inwards and oblique,
fourth pair deeifiy and acutely angled far in to below the spot
closing the cell ; the ground between the first and the third jiair
which are below each other is deep cream.
5 . Similar to the male, but with very broad borders to the costa
and termen and a narrow dark inner-marginal border in the prim-
African species of the Lycacncsthcs group of Lycaeniclae. Gl
aries. The secondaries with a broadish dark subterminal line, the
dark brown extending to the termen along the veins. The under-
side of both wings is just like that of the male but paler.
Hah. Ogowe River ; Sierra Leone ; Gaboon ;
Ruwenzori ; Beni ; Mawambe ; Congo Free State,
February.
Type in the Carnegie Museum.
This species is easily recognised by its very brilliant
colour above and by the entire w'ing being coppery red.
The description of the female as also the figures of that
se.x given by Smith and Kirby (Rhop. Exot., p. 98, PI.
XXir, If. 5, 6) do not belong to tins species ; I have both
sexes and they cannot be mistaken. The figures are, I
have no doubt, the female of L. lychnaptes, Holl.
Genitalia. — Harpago broad, nearly straight as to its lower edge,
but rising up to a high ridge at its upper extremity, front edge highly
serrated from apex forming a long, strong, straight 2)oint at about a
quarter from the lower extremity, immediately below which it is
highly excavated and terminates in a heavy j)ointed shoulder.
Cingula heavily developed at the base with the backward projection
broad, the arms forming the girdle rather short, narrowish, expand¬
ing but little at the tegumen. Tegumen deeply excavated at the front
apex, cheeks gradually taj^ering to a straight front edge and not
large ; the fakes large and strong, terminating in a heavy talon
curved at right angles at about a quarter from their sockets. Furca
with base small, the arms long, very narrow, receding well back¬
wards. Penis slightly hollowed for its hinder half, the fore half
gradually reduced to the orifice, which suddenly expands slightly,
terminating in a tapering trumjjet-sliaped tip. The cheeks of the
tegumen are well furnished with strong hairs at their lower angle,
the hairs on the harpagines being very few.
Lycaenesthes pyroptera, Auriv.
L. ygroptera, Auriv., Ent. Nachr., p. .382 (1895); id. id.
Rbop. Aethiop., p. 352, PI. 6, f. 3 (1898).
(?• Ujq^erside : both wings brilliant coi)2)ery orange. Primaries
with a broad blackish ajiex and somewhat broadish termen. Second¬
aries with black base reduced to a minimum, the red extending even
into the abdominal fold. Underside : both wings jjale tawny with
whitish dividing lines. The characteristic feature of both wings is
the 2)ostmedian stripe, wdiich has a series of small oval black ^jupils
02
Mr, G. T. Btii^ine-Ba.ker’s Revinion of the
ill the more or less confluent spots ; the shajie of this line together
with the black pupils distinguishes the species from all others.
Hob. Banana; IMompono; Congo Free State;
Bitje; Ja River, Cameroons {Bruce Coll).
This is a very pretty and distinct species,
Lycaencsthes lychnr.qjtcs, Holland.
L. lychncqMes, Holl., Psyche, p. 51 (1891) ; id. Ann. Mag,
N, H,, 1893, p. 250 ; id. Auriv., Rhop. Aethiop., p, 352
(1898); L. lyclinoptera, Smith and Kirby, Rliop, Exot.,
]), 97, PI. XXII, ff. 1, 2 (1893) ; L. scintilluld S. & K.,
p. 98, PI. XXII, ff. 5, 6 (1893).
d . Upperside : both wings bright orange yellow. Primaries with
costa and termen broadly and evenly dark brown. Secondaries with
costa, fold and termen to about vein 3 broadly dark brown. Under¬
side very dark blackish brown with creamy fine lines marking out
the usual pattern of this section of the genus; the markings are all
so confluent and solid, that the description can be.st be realised by
detailing the lines. Primaries with an oblique creamy line at the
base, two short parallel ones at the end of the cell, below which are
three oblique ones; from the costa midway to the cell are two more
short lines at converging angles, with a longer curved line below, and
extending between, two subterminal lines. Secondaries with two
sub-basal lines, another interrupted across the wings at the end of
the cell, double and angled below the cell ; three or four short dashes
in the radial area, a curved subterminal line, and an interrupted
terminal one.
Hah. Ogowe River, West Africa.
Type in the Carnegie Museum.
The broad borders of the upperside, and the very black
and solid pattern of the underside, will serve to discriminate
this species from others, as already mentioned. I have
little doubt that Smith and Kirby’s ffgures 5 and G {1. c ),
called by those authors scintillula, are really tliis species.
Lycaencsthes, zenkeri, Karsch. (Plate III, fig. 4.)
L. zenkeri, Karsch, Ent. Nadir., ji. 293 (1895); id. Auriv.,
Rhop. Aethiop., p. 253 (1898).
^ . Primaries in fresh specimens almost black, with a good-sized
orange patch nearly in the centre of the wings. Secondaries black
with a broad irregular orange dash in the po.stmedian area from
African apecies of the Lycamcstlics group of Lycaenidac. G3
vein 2 to 6. Termen with a whitish line interrupted at the veins.
Underside ; both wings blackish brown with white patches and
stripes. Primaries with a white basal dasli, a large white patcli in the
centre, a broad subapical dash to about vein 5, two fine pretenuinal
lines. Secondaries with a broad suboval white mark across the cell
to the abdominal edge and a longer curv'ed broad dash just outside
it, a trace of a black terminal spot between veins 2 and 3 edged
with reddish with a trace of iridescent scales, a preterminal fine
scalloped line.
Hah. Yaunde ; Bitje, Cameroons.
Type iu the Berlin Museum.
The underside pattern of this species will at once
separate it from its nearest ally, viz. L. juba, Fab.
Lycacncstlics juba, Fabricius. (Plate X, fig. 32.)
L. juba, Fab., Mant. Ins., p. 82 (1787); id. Auriv., Ent.
Tid., 1895, p. 219 ; id. Rhop. Aethiop., p. 353 (1898).
(J. U])perside’: both wings dark brown with large orange-red
patches. Primaries with the orange patch occupying all the median
and radial areas. Secondaries with the patch occupying irregularly
the postmedian area. Underside dark brown with white patches
and marks. Primaries with a broad basal dash along the U2)per
margin of the cell, a very large irregular white patch in the radial
area and along the fold nearly to the tornus, but being invaded on
the fold by the highly-curved postmedian broad line which is con¬
fluent and nearly semicirculai' ; a white curved subapical broad dash
terminates at the point where the broad dark subterminal line
touches the postmedian stripe. Secondaries with a large white patch
in the median and radial areas terminated by the curved postmedian
broadish stripe ; beyond this stripe is a broadish white one inter¬
rupted about veins 2 and 3 by the subterminal line, which is also
broad.
? • Upj)erside like the male, but paler with smaller spots. Under¬
side just like the male.
Hah. Sierra Leone; Bonny; S. Nigeria.
Genitalia. — Harpago broad somewhat saddle-shaped, but cut olf
straight at the bottom edge, fore apex rising into a saddle peak
rounded and slightly hollowed below on its front edge. Cingula
strongly produced backwards at its base with the girdle or arms
shortish. Tegumen well developed, deeply excavated at the front
64
Mr. G. T, Bf^^une -Baker’s Revision of the
dors.il apex, with the cheeks somewhat rounded. Falces long, narrow,
highly curved. Furca moderate in size with narrow arms inclined
backwards. Penis sheath broadish for its hinder part, then tapering
gradually to its trumpet-shaped orifice. Tegumen and harpagines
with strongish hairs.
Lycaencsthes 'pythagoras, Fabricius.
L. Pythagoras, Fab., Eiit. Syst., iii, p. 259 (1893) ; id.
Donov., Ins. Incl., PI. XXXIX, f. 8 (1800); tV?. Godart, Enc.
^[eth., p. 619 (1823).
(J. Upperside very close indeed to L. leptines, II ew. Underside
not unlike L. juba, Fab., but the secondaries apparently have all the
basal marks confluent, with a band of white beyond, following which
is the postmedian line. I have been quite unable to discover where
this type is or whether it exists. I therefore only know the insect
from the figure in Donovan’s work {1. c.), and I havevei’y little doubt
that the artist used both his imagination and his colour so as to
improve upon his model. This is specially evident in the underside.
Hah. “ In Indis ” ? ?.
Neurypexina, gen, nov.
The general neuration is similar to that of Lycaencsthes,
but in the primaries 11 and 12 anastomose; veins 7 and
8 are on a very long stalk branching to the apex and costa,
very near the latter, so that 8 is quite a short vein ; the
fork, however, is distinct under quite a low-power lens.
Eyes hairy. Palpi thickly scaled with hairs, end segment
long, smooth.
Type, Neurypexina lyzanius, Hew.
KEY TO SPECIES.
1. Blackish, above with blacker spots and less
dark intersjjaces. Below dark pattern
closely compressed . lyzanius.
2. Brown above, more uniform. Below white,
spots and marks well separated, decidedly
fewer markings than in lyzanius .... lamprocles.
Neurypexina lyzanius. Hew. (Plate XI, figs. 33-34.)
Tj. lyzanius. Hew., E. M. Mag., 1874, p. 86 ; Ill. H. Lep.,
p. 226, PI. 91, ff. 27, 28 (1878) ; L. regillus, Holland, Psyche,
African species of the Lycaenesthes yronp of Lycaenidae. 65
p. 51 (1891) ; L. regilla, Auriv., Khop. Aetbiop., p. 351
(1898); L. turlahis, Smith and Kii'by, Rhop. Exot., p.
105, PL XXIII, ff. 11, 12 (1893) ; L. turhata, Auriv., RIiop.
Aetbiop., p. 851 (1898).
^ . Both wings brown with darker spots ; these spots are produced
by the under.side pattern showing through, though in an intensified
and more definite form. Primaries with the basal wedge, the spot at
the end of the cell, and the postmedian stripe present ; the latter
broken up into three ; a sliort costal oblique dash ; two spots .shifted
well inwards, the lower one much the larger of the two, and a large
spot in the fold again shifted well in, each of the three being practi¬
cally isolated. Secondaries with the spots less evident, one at tlie end
of the cell ; the postmedian stripe irregular, composed of three pairs of
spots, the middle pair being shifted right outwards ; the seventh and
eighth angled spots though very definite on the underside do not
appear at all on the upperside. Undersurface very close to Lycae¬
nesthes flavomucxdai'as, S. & K., but with a dark line in tlie primaries
between the wedge spot and that closing the cell, also the costal part
of the postmedian stripe is much less oblique.
Hal. Old Calabar ; Ogowe River ; Cameroons ;
Gaboon ; Sierra Leone ; Bassa Province and Oruga
(N. Nigeria); BiHE District (Angola).
Type in tbe British Museum.
It has been somewhat difficult to unravel this species.
Fortunately, however, I possess N. turlatus, sent me by
Staudinger before his death. Dr. Holland has been good
enough to send me most beautiful and evidently most
accurate coloured figures of his regillus made by his own
hand, and a very careful comparison showed them both
to be Hewitson’s species lyzanius. The question then,
however, arose as to the relationship with L. flavomaculatus,
S. & K., to which it is very closely allied superficially ; the
underside pattern, however, shows definite differences as
already mentioned, and added to these the neuration of the
two species also differs, thus separating them generically.
L. flavomaculatus is a true Lycaenesthes, whilst in lyzanius
veins 11 and 12 anastomose.
Neurypexina lamp'ocles, Hew.
L. lamprocles, Hew., Ill. D. Lep., jx 225, PI. 91, f. 81
(1878) ; id. Auriv., Rhop. Aethiop., p. 351 (1898).
TRANS. ENT. SOC. LOND. 1910. — PART I. (JUNE) F
06
Mr. G. T. Bi^uiie-Baker’s Revision of the
. Both wings nnifonn dark brown above. Primaries below with
a large spot closing the cell and a still larger one below it below
vein 2. Postinedian line less oblique and continued down to vein 2.
PTudersurface near lachare-% Ilew., but the markings are decidedly
fewer tlius showing more white of the ground-colour.
Hah. West Africa; Cameroons.
Type in the British Museum.
This is an obscure species, again very closely allied to a
true Li/caencsthcs, viz. ^ lachares, but it lias fewer spots and
they are more localised. From the ^ of lachares it can be
easily separated by its brown upperside and by the fact
that the underside is like the ^ and not like the ^ of that
species. From N. lyzanius it may be recognised by the
underside having decidedly fewer markings, which are also
more separated.
Genus Neurellipes.
Differs from Tricleina, Karscli, in that veins 11 and 12
do not anastomose ; the neuration is otherwise similar to
Triclema, in which veins 8 and 9 are absent. Eyes hairy.
Palpi roughly scaled with hairs, end segment of moderate
length, somewhat robust.
Type, Neurellipes lusoncs, Hew.
KEY TO SPECIES.
1. Both wings blackish brown ; primaries
with an oval orange-coloui-ed spot
between veins Ici and 2. Below
blackish, spot on primaries above
does not show through ....
2. Both wings blackish brown ; jirimaries
M'ith a largish round orange spot
in lower radial area. Below brown
spot on primaries above showing
through .
3. Both wings dark brown ; primaries
with a largish round orange spot ;
secondaries with a short blue dash
in the internervular spaces of veins
1-3 .
cliryseosticUis.
lusones.
maeander.
African si^cdcs of the Lucaenesthes gron]} of Lycaenidae. G7
4. Blackish brown above ; j^rimaries with
basal half of fold blue. Below
strongly patched with white . . . standingeri.
5. Warm brown above, spotted like Ly-
caenesthes (sens, strict.) below. Sec¬
ondaries with a marginal row of
black spots with metallic scales . yemmiferu.
JVcurellipcs chri/seostictiis, sp. nov. (Plates III, fi" 5;
XII, figs. 37-38.)
(J. Both wings on the upperside blackish brown; primaries with
an orange-red oval spot in the fold between veins la and 2, beyond
the end of the cell, rarely extending very slightly over vein 2
Underside like N. lusones, Hew., but the colour is not brown but
blackish, and the orange patch of the primaries does not show through,
the white margins to the marks are whiter and broader ; with the
exception of these differences the underside pattern is very closely
similar. There are certain divergencies, however, in the structure of
the pattern and in the look of it that render the separation of the
two comparatively easy.
5 . Like the male both in the upper- and undersides ; whereas the
$ of lusones is paler above, and in the secondaries has a pale sub¬
terminal line, whilst below it is decidedly paler, with the mark¬
ings reduced to a certain extent by the broadness of the white
lines.
Hctb. Sierra Leoxe; Bass a Province (N. Nigeria).
Types ill my collection.
This species and N. lusones, Hew., are found flying
together from November to February, and it is possible
that they may be in many collections mixed together. The
spot in the primaries of lusones is, however, much larger
and is always more or less round ; in my species it is much
smaller and always oval. The genitalia also differ in
several points.
Genitalia. — Harpago broad, evenly excurved as to its lower edge,
gradually rising on its upjier edge to near the apex, when it suddenly
rises again into a high ridge ; front edge erect, at a third very highly
excavated, fornang a sharp long tooth, the previous part being
slightly serrated, the lower extremity being produced into a chisel¬
like edge. The cingula is of moderate width, well produced back¬
wards at its lower edge, gradually tapering to the cheeks of the
F 2
(38
tej,uiinen, which is well excavated at its front apex, tlie cheeks oradii-
ally curved to their base, and somewhat serrated at the edge. Falces
long, gradually curved from their sockets. Furca rising from a high
base, with fine arms, nearly straight, inclined backward.s. Penis
sheath long, broadish for the hinder half, then nearly straight to the
tip, which is somewhat trumpet-shaped.
Neurdli'pes lusones, Hew. (Plates III, fig. G ; XII, figs.
85-36.)
Lycaenesthes lusones, Hew., Trans. Eat. Soc. Load.,
1874, p. 347; id. Ill. Diara. Lep., p. 227, PI. 91, if. 17,
18 (1878) ; id. Dewitz., N. Acta. A. N. C., p. 202, PL 26, f. 8
(1879); id. Auriv., Rliop. Aethiop., p. 352 (1898); X.
fidvimacula, Mab., An. Eat. France, p. 24, PI. II, f. 5 (1890);
id. Auriv., Rhop. Aethiop., p. 352 (1898).
(J. Both wings brown above. Primaries with a large orange -yellow
spot in the lower radial area and extending into the fold. Underside :
both wings brown, with the pattern marked out by greyish-white
lines. The main distinguishing feature is that the orange-coloured
spot of the upperside shows well through in tlie primaries on the
underside.
9 . Paler brown above. Secondaries with a pale subterminal
line ; the underside decidedly paler brown, with the lines much
whiter and broader than in the male.
Hob. Sierra Leone; Ivory Coast; N. Nigeria;
Bassa and Kabba Provinces ; Cameroons ; Gaboon ;
Congo Free State ; Angola.
Type in the British Museum.
The differences between this species and the previous
one (chryseostictus) have already been referred to under
the latter description. M. Mabille has been so good as to
lend me his type of fidvimacula from the Ivory Coast, and
it agrees in all particulars with Hewitson’s.
Genitalia. — Harpago broad, with the upper edge gradually rising,
then very slightly hollowed, from whence it rises into a highish
process, the front edge being quite straight ; at its lower extremity
it is very highly excavated, the lower extremity thus forming a
longish tooth or point, the excavation being gradually rounded off
to the lower edge, which is slightly hollowed at first, and then
nearly straight to its origin with the cingula ; this is w'ell developed,
rapidly expanding towards the cheeks of the tegumen, and having
its lower hind extremity produced backwards in a long, hollowed
African species of the Lijcacncsthcs group of Lycaenidae. 60
formation ; the tegiunen is deeply excavated at its fore apex, the
clieeks are gradually rounded off with their lower edge straightish ;
the falces, with talou-like tips, are highly curved at about a third
from their sockets ; the furca is long, with a high, narrow, pyramidal
base, and longish narrow arms curved backwards. Penis sheath of
moderate length, hollowed for its hinder third, and gradually tapering
from its highest point at two-thirds to the orifice, which is trumpet-
shaiied.
Neurellipes maeandcr, Plotz.
Lycacnestlics maeandcr, Plotz, Stt. Eut. Zeit., 1880, p
202; id. Auriv., Eiit. Tid., 1895, p. 218; id. id. Rhop.
Aethiop., p. 351 (1898).
^ . Both wings uniformly blackish, with an orange spot in the
primaries, which spot does not show through on the underside. In
the secondaries there is a fine blue streak in cells 1 to 3.
5 . Like the male, but the orange S2)ot of the primaries shows
through on the undenside.
Hob. Einingo ; Ogowe River ; Gaboon ; Cameroons.
I have been unable to find the type of this species.
Herr Plotz’s collection was unfortunately disposed of
piecemeal, so that I have not seen the species. It is
evidently near N. lusones, Hew.
Neurellipes staudingeri, Smith and Kirby. (Plate XI,
fig. 39.)
Lycaencstlies staudingeri, S. & K., Rhop. Exot., p. 112,
PI. XXIV, ff. 9, 10 (1894); id. Auriv., Rhop. Aethiop., p.
150 (1898).
d. Both wings brownish black above. Primaries with a deep
sub-lustrous blue dash in the fold below the cell, and extending
slightly below vein 2. Underside blackish, with the usual marks
typical of this section of the group ; the separating lines are very
fine and interrupted, and there are several white patches which are the
distinguishing feature of this species. Primaries with an oval white
patch across the end of the cell ; two subapical white spots. Second¬
aries with a large white spot at the end of the cell, and a larger one
at the apex.
$ . Brown above. Primaries with a whitish spotted subterminal
line. Underside palish brown in both wings with the pattern as in
the male.
Hah. SiERHA Leone ; Ogowe River ; Ashanti ;
Upper Kasii District; Congo Free State.
70
ilr. G. T. I une-Baker’s Revision of the
This species, of which I have a series from different
localities in Sierra Leone, can be recognised immediately
by its underside pattern, which is very distinct in its
prominent white spots from any other of the family.
Genitalia. — Harpago verp specialised, consisting of a narrow bifid
arm, the upper portion of which is an erect, curved, very narrow arm,
the Iqwer arm being bent, Avith a knob-shaped extremity, with a
blunt, broad tooth at a quarter from the tip, the edge being well
excised from the tooth. The cingula is produced far backwards at
its basal extremity, the girdle being strongly curvmd, expanding
slightly to the cheeks of the tegnmen ; this is deeply excavated at
the front apex, the cheeks being moderately evenly curved ; the
falces are narrow, somewhat Aveak, and curA'ed Avell above their
sockets ; the furca rises as a narrow-nrmed fork directly at its base,
and is inclined gently baseAvards. Penis sheath broaclish for its
hinder half, Avell curved on its loAver margin, and forming a gradualh"-
tapering tube as to its fore half. The hairs on the cheeks of the
tegumen are someAvhat restricted, and those on the harpagines are
much restricted.
Neurelli'pes gemmifera, N eave.
Mr. Neave took iu the East LoangAva district in 1905 Iavo
specimens of a broAvn “ Lycaenesthid ” Avbich form a most
interesting species, truly tyjDical Lycaenestlies (sens, strict.)
beloAv, except that they have a terminal roAv of largish
black spots on the secondaries, margined finely Avith bright
orange and with bronzy-green metallic scales ; it is quite
remarkable in appearance, whilst also in the secondaries
each of the veins form at the termination a short, fine, hair¬
like tail as at anal angle.
The neuration is that of this genus.
Hah. Petauke ; N. E. Rhodesia, 2,400 feet, March.
Type in the British Museum.
Triclema, Karsch.
Triclema, Berliner Stt. Ent. Zeit, p. 214 (1893); id.
Reuter, Acta. Soc. Sc. Fen., p. 182 (1896) ; id. Auriv.,
Rhop. Aethiop., p. 345 (1898); Lycacncsthes {in parte),
Auriv., Rhop. Aethiop., pp. 345-353 (1898), trf. ; Ark.
Zook, II, p. 16 (1905).
Head roughly haired ; palpi A^ery hairy ; end segment of moderate
length ; eyes hairy ; neuration similar to Lycaenestlies, but with veins
S and 9 of the primaries absent, Avhilst veins 11 and 12 anastomose.
African r,<pccic& of the Lycacncstlus gvonf of Lycacnidac. 71
Type, Triclema lucretilis, Hew.
Karscli was the first to divide the somewhat large and
intricate genus Lycaenesthes into two, when he separated
the species in which vein 8 was absent and in which the
costal vein anastomoses with 11. However closely a
group of insects may be allied together by a merely super¬
ficial likeness of pattern, yet when it is seen that a certain
number have structural characters differing along the
same line, it is evident that the family is in the process
of change, and that in the evolution of the group it is
splitting up itself into different genera. Under such
circumstances it appears to me not only justifiable but
necessary to adopt the lines that are being worked out by
the insects themselves, and when structure in the imago
changes then that species should form or be placed in
another genus, and this should apply in all cases, except
where it is known that such structure varies inter se in the
species of the genus. The genus diagnosed by Kai’sch
should thei’efore be adopted. In the species T. lamias,
whilst the neuration of the male is constant that of the
female is not so, for out of a fairly long series I have two
in which veins 11 and 12 do not anastomose, though even
here there is a possibility of an aberration.
KEY TO SPECIES.
A. Upper side brown.
t. Primaries above with a tawny patch . rufoplagata.
2. ,, „ uniform dull brown . lacides.
.3. „ with no basal spots in
either wing . tisamenus.
4 Primaries above blackish ; markings
below very dark . hades.
5. Primaries above dull brown, dark spot
at end of cell, dark subtermiiial line in
both wings. Below pattern markedly
divided with white . phoenicis.
6. Both wings warm brown above ; primaries
with a trace of a short paler subter-
ininal line ; secondaries with a distinct
pale subterminal line. Below warm
brown with broad white dividing
lines . lamilila.
72 Mr. G. T. I '^Piune-Baker’s Revision of the
7. Both wings blackisli above witli obscure
subterniinal lines. Below greyish
with obscure whitish-grey dividing
lines . tiUjeriae.
8. Both wings brownish above ; secoiulai’ies
only with a pale subterniinal line.
Below whitish with pattern broken
up into more or less parallel stripes . orulatvs.
9. Both wings brownish above ; secondaries
with a pale postmedian line and a
terminal row of small spots. Below
white with the usual pattern . . . marshalli.
10. Both wings brownish above with pale
lilac-blue spots . Inmias.
11. Both wings black with bright blue lines ;
primaries with a bright blue sub-basal
figure 8. Below black with white
patches . h(cretili>t.
12. Both wings blackish; 2)rimaries with
mauve-blue reflections in lower part
of cell and in fold. Below black with
white dividing lines . fanciatus.
13. Both wings with blue patches above.
Underside with less white .... ohscura.
14. Both wings blue above . caernlea.
Triclcvia o'vfoitlagata, sp. nov. (Plates III, fig. 8 ; XI,
fig. 40.)
^ . Both wings brown ; primaries only with a pale tawny chest¬
nut patch around the angle of veins 3 and 4, extending but little into
the cell and well into the postmedian area. Underside : both wings
dark greyish brown with darker spots edged with whitish. Primaries
with a broad, wedge-shaped sub-basal spot, an oblique spot closing the
cell, with a small one above it; postmedian strij^e irregular with two
costal spots, followed by two slightly angled shifted well outwards,
below which are two shifted well inward.s, the lower of which is the
larger, and having its inner edge shifted further in, seventh spot
oblique shifted right in with its outer whitish edge confluent with
the inner edge of the previous sjjot; subterminal line increa.sing in
width towards the tornus. Secondaries with the large basal band
broken up into two spots, the upper one ovate, the lower irregular,
large, projecting outwards in the cell to touch the somewhat oblique
African sjJccics of the Lycacncdhcs gronjj of Lycaenidae. 73
spot closing the cell. Postmeclian stripe irregular, with the second
spot smaller than the first and shifted slightly outwards, third and
fourth larger, confluent, fifth spot small shifted slightly inwards,
sixth larger shifted well inwards, seventh and eighth strongly angled,
confluent, and shifted right inwards; subteruiinal line broadish,
increasing in width towards the angle ; a black terminal spot between
veins 2 and 3, and a double one at the angle, each with bluish-
metallic scales and edged above and laterally with orange red.
$ . Like the male in all particulars, except that the chestnut
patch on the upperside of the primaries is larger.
Expanse ^ 28, 9 29 mm.
Hctb. Sierra Leone.
Types in my collection — a series from ditferent places
in the Colony.
Genitalia. — Harpago somewhat pear-shaped, tapering to a narrow
extremity, strongly serrated and curved downwards, the upper edge
is evenly curved and wmll studded with minute tubercles and teeth ;
the cingula is erect, narrow with long arms expanding very little at the
cheeks of the tegumen ; the base is produced backwards into a long,
blunt, tube-like process ; the tegumen is fairly developed, excavateil
somewhat at the front apex, with cheeks having evenly-curved rather
long fore edges; the fakes long and strong with small talon-like tips,
and are highly curved almost from their sockets ; the furca has longish
narrow arms curved well backwards. Penis sheath broad, hollowed
for the hinder two-fifths, and gradually tapering from thence to the
orifice, which is thickly studded for a fifth with fine, close serrations,
reminding one closely in arrangement of the teeth in the palate of a
pike. The cheeks of the tegumen and the upper edge and extremity
of the harpagines are furnished with long hairs.
Triclema Ictcidcs, He^v. (Plate II F, fig-. 7.)
Lycacnesthes lacides, Hew., Traus. Ent. Soc. Lend., 1874,
p. 348; Ill. D. Lep., p. 227, PI. 91, ff. 19, 20 (1878); id.
Auriv., Rhop. Aetliiop., p. 351 (1898).
5 . Both wings uniform dull brown above. Ltndersurface : both
wings similar to above, but decidedly warmer in tone, with markings
scarcely darker than the ground-colour. Primaries with the upper
four spots quite confluent and nearly straight, not touching the
white subterminal line. Secondaries with the basal band not broken
into spots, a whitish patch near the apex above the third spot of
74
Mr. G. T. luric-Baker’s Bxvision of the
the postniediau stripe. The otlier spots have little to call for
dilTerentiatioii.
llab. Angola.
Type in the Britisli Museimi.
Triclema iisamcnus, Holland. (Plate III, fig. 13.)
Lycacnesthes iisamcnus, Holl., Psyche, p. 52 (1891); id.
Aiiriv., Rhop. Aethiop., p. 849 (1898).
g. Both Avings uniform dull brown above. Underside pale
brownish grey with no basal spots in eitlier wing. Primaries with
a spot closing the cell ; postmedian stripe conlluent and evenly
curved to vein 2 ; the sjiot l)elow this is shifted inwards, a narrow
dark subterminal line. Secondaries with a sub-basal spot below
the costa and a small one on the abdominal margin; a small spot
near the end of the cell; postmedian stripe with two confluent
spots on the costa, below which it is resolved into an indefinite
waved line ; subterminal dark line slightly waved.
Hal. Ogowe River.
Type in the Carnegie Museum.
The absence of basal spots should enable this species to
be recognised very easily.
Trichma hades, sp. nov. (Plates III, fig. 9 ; XI, fig. 41.)
g . Both wings uniform dark brown. Primaries with an obscure
dark spot closing the cell and an obscure dark subterminal arched
line of even width, and not parallel with the termen from the costa
to the .inner margin. Underside : both wings dark brown with
slightly darker spots finely edged with whitish. Primaries with a
lu’oad, wedge-shaped sub-basal mark, an olilong spot closing the cell,
above which is a small costal spot ; postmedian stripe irregular, first
and second spots confluent, the second the larger of the two, third
and fourth spots larger, confluent, slightly angled internally, the
fourth the larger with its outer edge shifted outwards, both being
shifted well out from the second, fifth and sixth large shifted right
in, extending below the spot closing the cell and touching it, sixth
spot the larger, its inner edge extending rather beyond the fifth,
seventh spot very oblique shifted right in so that its inner edge
nearly touches the wedge-shaped mark, a broadish arched sub¬
terminal line (showing through on the upper side), followed by a
fine, even, preterminal line. Secondaries with the basal broad band
broken up into two, the upper part subovate, the lower part
African sjjccies of the Lycacnesthes gronyj of Lyccicnidae. 7 5
irregular, broken outwards at the lower margin of the cell and
constricted below this, an oblique oblong spot closing the cell ;
postiuedian stripe irregular, the first two spots confluent, the first
the larger and the darker of the two, third and fourth confluent
shifted well outwards, the fourth being shifted slightly further out as
to its outer margin, fifth spot very small shifted well in, sixth larger
with its inner margin beyond and touching the spot closing the cell,
seventh and eighth shifted right in, strongly angled, almost touching
the sub-basal band ; a well-arched subterminal broadish line, followed
by a series of four small sublunular dark terminal spots from the
apex to the blackish spot with metallic-blue scales edged with deep
orange red, at the angle is another similar spot but smaller.
Expanse 25 mm.
Hal. Sierra Leone.
Typo in my collection.
This species is near T. 'pliocnicis, Kar.sch; the dilferences
are very difficult to describe in words, though real in
effect ; the figures will show them better than words.
Genitalia.— Harpago broad, hollowed, its lower edge slightly
curved, its upper edge tapering down to three quarters, where it is
excised and suddenly elevated into two long sharp teeth, from
where it rapidly tajiers to the extremity which is develo])ed into a
very long fine curved tusk-like jirocess. Cingula with narrow
arms expanding somewhat into the cheeks of the tegumen, the basal
process projected backwards into a gradually tapering formation.
Tegumen moderately developed, the whole of the front apex slightly
and gradually excavated, cheeks evenly rounded from the excavation,
then straight, with the falces long, strong, slightly hooked at the
extremities, highly curved about a quarter from their sockets.
Furca with base short, longish narrow arms curved backwards.
Penis sheath with the hinder half broadish somewhat oval, gradually
tapering at the middle to the tip, tbe extremity of which is exca¬
vated, the apical fifth being roughly shagreened.
Trichvia inlioenicis, Karseb. (Plates III, fig. 10 ; XIII,
fig. 42.)
Lycacncsthcs iiliocnids, Karseb, B. E. Z., 1893, p. 238 ;
id. Auriv., Rbop. Aetliiop., p. 351 (1898) ; id. id. Ent. Tid.,
1901, p. 118.
g. Both wings uniformly brown above. Secondaries with an
obscure dark terminal spot between veins 2 and 3. Underside : both
wings very pale brownish with the usual Triclema pattern divided
7G
Mr. G. T. 1 >^iune-Ijaker’s Revision of the
l)y whitish lines; postmedian stripe with the costal four .spots nearly
confluent very slightly oblique, the third and fourth spots dark,
fifth and sixth also dark and shifted far inwards, seventh spot
dark and again shifted far in ; an obscure darkish subterminal line
increasing in width somewhat towards the tornns. Secondaries with
the basal I'and divided into spots touching each other; postmedian
stripe, with first costal spot longish and dark ; second, small, same
colour as the ground; third and fourth shifted right out, dark; fifth
and sixth shifted right in, slightly darker than the ground ; seventh
and eighth angled spot dark and shifted far inwards ; anal spots
blackish and one above it, edged internally with reddish and having
metallic scales.
$ . Upperside paler brown than the male. Secondaries with a pale
subterminal curved line interru^sted at the veins. Undersurface like
the male, but the white dividing lines broader.
Hah. Togo; Cameroons; Ogruga (Nigeria); Sierra
Leone; Patigo (Acholi Country); Uganda Protect¬
orate ; N. E. Rhodesia (Neave).
Type in the Berlin Museum.
A species very near T. hades, B.-B., but less dark above,
whilst below the pattern is less prominent and smaller
altogether.
Genitalia. — Harpago broad, hollowed, both upper and lower edges
minutely serrate, the lower developing into a strong tooth in front
of the extremity, the lower apex forming a chisel-like projection
with a serrated edge, the upper apex forming a longish strong tooth.
Cingula well developed with the lower base produced well back¬
wards, the girdle narrow, expanding but slightly at the cheeks of
the tegumen, which is very narrow, slightly hollowed in front, with
small cheeks. Falces long and strong, well curved at a third from
their sockets. Furca with long narrow arms well apart. Penis sheath
a long ellipse for three-quarters its length, suddenly narrowing to
an oblique extremity. Hairs on cheeks of tegumen and end of
harpagines long, but not numerous.
Triclema kamilila, sp. nov. (Plates III, fig. 11; XIII,
fig. 43.)
g. Both wings brown. Primaries with a dark dash closing the
cell, around which is a small patch of a lighter tone, an obscure
series of internervular marginal dashes, obscurely and palely edged
internally. Secondaries with a strongly-ai’ched series of subterminal
internervular whitish-grey dashes from the costa to vein In, followed
African species of the Lycaenesthes group of Lycieniclac. 77
by a series of terminal black points gradually developing into spots
towards the angle ; ternien finely black with grey fringes. Secondaries ;
both wings coffee brown with darker spots broadly edged with white.
Primaries with the usual wedge-shaped sub-basal spot, the oblong
spot closing the cell with a small costal one above it ; the postmedian
stripe irregular, the first four strongly oblique, of which the lower
two are shifted well outwards but continuing the oblique position,
fourth and fifth confluent, slightly oblique, shifted right in and
touching the cell spot, seventh oblique shifted far in so that its
outer margin is confluent with the inner margin of the sixth spot ;
subterminal line very fine in its upper radial area, increasing in
width in the tornal area, broadly edged laterally with white ; a fine
preterminal line. Secondaries with the sub-basal band broken up
into three spots, the upper one ovate, the middle one reniform
divided into two by a white line, the third and lowest one con¬
stricted ; cell closed by an oblique rectangular spot with a fine
central line ; postmedian stripe with the first spot dark, second paler
shifted outwards, third, fourth, and fifth confluent arched, shifted
well out, the third and fourth being dark, the fiftli paler and smaller,
sixth shifted inwards, seventh and eighth angled and shifted right
in ; subterminal line obsolescent, a terminal series of obscure small
dashes, the subterminal area being broadly whitish, a dark spot
between veins 2 and 3 and a smaller one at the angle, well covered
with bluish-metallic scales and broadly edged with bright orange.
Ternien very finely dark.
5 . Just like the male.
Expanse 28 mm.
Hah. Kamililo; Nandi, June; Sierra Leone.
Type in my collection.
Tlie type was taken in tlie Nandi country ; but I have
a female from Sierra Leone that exactly corresponds with
this male, and as it suits no other species I conclude it
must be this.
Genitalia. — Hariiago broad, well hollowed, upper apex sharply
triangular, lower apex produced into a longish blunt-edged point
finely serrated, lower edge developing a longish fine tooth just
beyond the middle. Cingula broadly produced backwards at its
base, girdle longish, narrow, expanding slightly to the tegumen,
which is narrow, very slightly hollowed, with rather wider cheeks.
Falces long, narrow, well curved at a third from the sockets. Furca
with long, narrow arms strongly curved backwards. Penis sheath
a gradually tapering ellipse suddenly hollowed at a quarter from the
78
Mr. G. T. I une-Baker’s Revision of the
tip, then expanding into a .sliglitly trumpet-shaped orifice. Cheeks
of teguincn and apex of harpago scantily supplied with hairs.
Triclema nigeriae, Auriv.
LycaenestlLCs nigeriae, Auriv., Arkiv. Zool., ii, p. 16, PI. Ill,
f. 4 (1905).
Both wings sooty blackish above; each with a terminal
broadish border obscurely edged by palish scalloped lines. Under¬
side pale greyish with pattern very similar to Kamilila, B.-B., but
the spots smaller and the pattern somewhat different.
Hah. Benue; Lokoja; Nigeria.
It is very difficult to describe the differences between
this and my species. Kamilila, my insect, is a warm
brown above and below, not at all black ; whilst the
pattern described in words would be quite similar, the
appearance is very different, the whitish terminal areas,
the iridescent scales to the anal spots of the secondaries
edged above with bright pale orange, and the somewhat
different spots, make the insect look very different.
Triclema oculatus. Smith and Kirby.
lAicaenestkcs oculatus, S. & K., Bhop. Exot., p. 101,
PI. XXIII, ff. 1, 2 (1893); id. Auriv., Rhop. Aethiop.,
p. 350 (1898).
9 . Upperside : both wings entirely pale brown above. Secondaries
with a brown subterminal line edged on each side rather broadly
with deep cream colour. Underside: both wings whitish crossed by
numerous brown lines. Primaries with an oblicpie basal line, two
transverse median, a postmedian and a subterminal brown line.
Secondaries with the radial area crossed by numerous short, very
fine dark dashes, the median area pale brownish with obscure traces
of tlie white ground ; the usual anal spots are present but small.
Hah. Ogowe River.
I do not consider that this species has anything to do
with L. mclamhrotus, Holl.; they belong to different genera
and are widely separated in pattern. It is an obscure little
species, and may be best recognised by the fact that the
usual pattern is converted into a series of stripes ; there is
no veal postmedian line present, this applies in a special
degree to the secondaries.
African species of the Lycacnesthes group) of Lycaenidae. 79
Triclcma marshalli, Betliune-Baker. (Plate III, fig. 12.)
Lycacnesthes marshalli, B.-B., Ann. Mag. N. H., Ser. 7,
Vol. XII, p. 332 (1903).
Differs from T, phoenicis, Karscli, in the underside, which is
browner, the basal spots in both wings are very dark brown; in
Karsch’s species they are the same colour as the ground, viz. grey ;
all the spots are more filled in with brown, whilst the angled seventh
and eighth spots of the secondaries are entirely dark in my species,
but in 2)hoen}cis, of which I have seen a long series, they are quite the
same as the ground-colour.
Hah. Sierra Leone; Ogruga (Nigeria).
Type in my collection.
Genitalia. — Harpago broadish with lower extremity forming a long,
sc^uare serrated point, the lower edge being produced into a blunt
point at a quarter from the apex, between which it is well excavated,
upper edge develoi:)ed into a short, sharp tooth. Cingula with base
well produced backwards, girdle very narrow. Tegumen of even
width with slightly expanding cheeks. Falces long of moderate
strength, highly curved at two-fifths from their sockets. Furca with
very long arms inclined forwards, the arms are broad at first then
suddenly constricted very narrowly. Penis sheath broadish, hollowed
for the hinder half, then gradually tapering down to a suddenly
expanding trumpet-shaped orifice. Tegumen and harpagines well
supjjlied with long hairs.
Triclcma lamias, Hew. (Plates III, fig. 14; XIII, fig. 44.)
Lycacnesthes lamias, Hew., Ill. D. Lep., p. 227, PL 91,
ff. 25, 26 (1878); id. Auiiv., Rhop. Aethiop., p. 351
(1898).
g . Both wings sooty brown above. Primaries with i)ale bluish
spots in the postmedian area of the wing, and a double one in the
middle of the fold, and a horizontal dash further out near the tornus.
Secondaries with a pale bluish po.5tmedian line interrupted at the
veins, and an obscure fine subterminal scalloped one enclosing an
obscure row of small spots. The characteristic feature of the under¬
side in the primaries is the postmedian stripe, which is reduced to
an oblique confluent stripe from the costa to vein 3, the whole of
the lower part being absent; in the secondaries the basal band is
irregular but confluent, and is not broken up into separate spots ;
the postmedian stripe is divided into three pair of spots, the fifth
and sixth spots being reduced to a minimum ; the second pair is
80
IVlr. G. T. rtW lUiie-Baker’s Herismi of the
shifted right out, and the seventh and eighth angled spots being
shifted right below the sj)ot closing the cell and touching it ; the
dark subterininal stripe edged with white is broadish and distinct ;
the anal spots of the secondaries are present as usual.
9- Entirely pale brown above in both wings. Underside like the
male, but with broader wliite lines.
Jlal). Sierra Leone; Nigeria; Cameroons; Gaboon.
Type in the British Museum.
This species, a common one in Sierra Leone, can
easily be recognised by the pale whitish-blue marks of the
upperside ; they are, liow^ever, somewhat fugitive, the super¬
posed scales being very easily rubbed off ; they leave,
however, behind a slight trace of their former presence.
Genitdlia. — Harpago deeply split from the fore edge, the lower
part forming e.xteriorly an angled sharp-pointed curved knife with
a minutely-serrated edge, the upper extremity being somewhat
hammer-headed. Cingula well developed with the base produced
backwards and the arms fairly wide. Tegumen narrowish with
wider cheeks. Falces short, curved at their middle. Eurca with
narrow waved arms inclined backwards. Penis sheath very broad at
the base, gradually tapering to a trumpet-shaped orifice. Tegumen
and harpagines fairly supplied with hairs.
Tridema lucretilis, Hew.
Lycacnesthes lucretilis, Hew., Trans. Ent. Soc. Lond., IST-f
p.349; id. Bl. D. Lep., p. 228, PI. 91, fif. 29, SO (1878) ;
id. Dewitz., N. Acta. Acad. N. Cur., p. 180, PL 26, f. 4
(1879) ; id. S. & K., Rhop. Exot., p. 103, PI. XXIII, ff. 5, 6
(1893) ; id. Auriv., Pdiop. Aethiop., p. 351 (1898); id. id.
Arkiv. Zook, ii, p. 16 (1905) ; Lycaenesthes lucretia. S. & K.,
Rhop. Exot., p. Ill, PI. XXIV, ff. 7, 8 (1894) ; il Auriv.,
Rhop. Aethiop., p. 351 (1898); Tridema lucretilis, Karsch,
B. E. Z., p. 214 (1893); id. Reuter, Act. Soc. Sc. Fenn., p.
182 (1896).
8 • Upperside : both wings black with bright blue marks.
Primaries with a brilliant fine blue figure 8 in the median area, two
fine blue lines more or less parallel beyond it from vein 1 to 5 or
6, a subterminal fine double blue line. Secondaries with a fine blue
circle in the cell, which is clo.^ed by a parallel double blue short line ;
postmedian stripe marked out by a double line of broken fine blue
short dashes ; termen very finely blue preceded by a similar line
African species of the Lycaencsthcs group> of Lyeacnidae. 81
just beyond tlie postmedian. Underside blackish brown with the
pattern common to this section marked out by white lines. Primaries
with a white spot closing the cell, and below it two broad short white
dashes in the fold. The white subterminal line is completely inter¬
cepted by the exceedingly oblique costal part of the postmedian
stripe. Secondaries with a white spot in the cell near its end, and a
smaller one at the apex.
Hob. Sierra Leone; Gaboon; Ogowe Kiver;
Chinchoso; Angola.
Type in the British Museum.
This is a very distinct species, both as to its upper- and
undersides. It is the species for which Karsch created the
genus. Smith and Kirby’s L. lucretia is really only
recognisable by the absence of the red edging to the anal
spots of the underside of the secondaries; this is a feature
that is variable in Incretilis, being sometimes reduced to a
very fine line. I have little doubt that it is Hewitson’s
species ; the iridescent anal spots, both in size and width
of edging and also in the metallic-blue scales, are very
variable and cannot safely be used in species making.
Triclcma fasciatus, Auriv. (Plates III, figs. 15 16 $;
XIII, fig. 45.)
L. fasciatus^ AxinY., Ent. Tid., 1895, p. 218; L.fasciata
id. Rhop. Aethiop., p. 350 (1898); id. id. Arkiv. Zook, ii, p. 16
(1905); L. suhnitens, B.-B., Ann. Mag. N. H., 1903, p. 332.
^ . Both wings blackish above. Primaries with the fold deep
lustrous violaceous blue to a quarter from the tornus, with discoidal
area of the same colour. - Underside: primaries with the postmedian
stripe twice strongly fractured, the four costal spots conliuent very
oblique, each spot rather further out than its predecessor, the fifth
and sixth spot confluent vertical shifted well in, seventh spot oblique
shifted far in, white subterminal stripe unbroken. Secondaries with
basal band of even width and unbroken ; postmedian stripe fractured
three times, the second pair of spots shifted right out, third pair and
fourth pair each well in from its predecessor, subterminal white line
highly curved with a marked indentation oi;twards at about its
middle.
9 . Both wings uniformly brown. Primaries with a postmedian
spotted palish line. Secondaries with a pale postmedian and subter¬
minal line, somewhat interrupted at the veins. Underside as the
male, but white lines broader.
TRANS. ENT. SOC. LOND. 1910. — PART I. (JUNE) G
82 Mr. G. T. lune-Baker’s Revision of the
Hob. Cameroons ; Congo Free State ; Sierra
Leone.
Type ill the Stockholm Museum.
Prof. Aurivillius has been good enough to let me
see the type of his T. fasciatus, and it is the same as my
siLhnitens, only in his specimen the blue is considerably
faded; it is highly to be desired that, in genera wherein the
species are so very closely allied, a figure of the species
sliould be given.
There should be little difficulty in recognising this
insect ; the uj^perside is somewhat near N. staudingeri,
S. & K., but there are no white patches below, and the
neuration of staudingeri is different, so that it has been
necessary to place it in another genus.
Genitalia. — HAi'i^ago very deeply bifid aluiost to the base, tlie
upper arm being well curved, narrow, and tongue-shaped, the lower
very narrow, slightly increasing to a knobbed extremity ; at a quarter
from its end is a short blunt tooth, from whence to the tip it is well
excised. Cingula well produced backwards at the base, narrow,
expanding to the cheeks of the tegumen, which is deeply excised at
its front apex, v’ith the fr.mt line oblique not curved until the lower
extremity. Falces longish, narrow, strongly curved at about a quarter
from their sockets. Penis sheath very narrowly elliptical, tapering
very narrowly from the middle to an oblique tip. Lower extremity
of harpagines and cheeks of tegumen -well supplied with longish
hairs.
Triclcma ohsciira, Druce.
Tiiis species is very close superficially to Neurcllipes
staudingeri, S. & K., but has blue on its secondaries as
well as the primaries, whilst the underside has very much
less white, and has practically only one white patch in the
secondaries.
Hah. Bitje ; Ja River, Cameroons.
I am indebted to Mr. H. H. Druce for the loan of this
species, which he is about to describe; it may be at once
distinguished from N. staudingeri by the underside, for in
the primaries the oblique costal part of the postmedian
stripe does not invade the white subterminal line.
Triclema cacrulea, Auriv. (Plate III, fig. 17.)
Lgcaenestltes cacrulea, Auriv., Ent. Tid., 1895, p. 217 ;
id. id. Rhop. Aethiop., p. 350 (1898).
African species of the Lycaenesthes group) of Lycaenidac. 8!^
8 . Both wings mauve blue. Primaries with apex broadly dark
brown gradually tapering along the termen to the tornus. Secondaries
with termen narrowly brown with a dark line of subterininal
dashes between the veins, costa brown above vein 8. Underside :
both wings w^arm brown with the usual Triclema 'pattevn. Primaries
with the postmedian stripe reduced to a curved tapering confluent
band to vein 2. Secondaries with the basal band divided into three
sjrots touching each other. Postmedian band fractured thrice below
each pair of spots, second pair shifted well out, third pair well in
and almost unicolorous with the ground-colour, fourth angled pair
shifted right below the spot closing the cell and almust confluent
with it.
ITah. Cameroons.
Type in the Stockholm Museum.
I am again indebted to Prof, Aurivilliiis for the loan of
this species ; it will be readily distinguished from all others
of the genus by its pale mauve blue colour extending
almost entirely over both wings.
Species Omitted.
I. liolcias, Westwood.
The type of this species has not yet been discovered
and as the late Hope Professor stated in his description
“ cauda mdlaf I have no hesitation in eliminating it.
L. yrammica, Smith and Kirby.
I have seen this type, and it is a Castcdius.
Since the foregoing paper went to press I have received,
through the kindness of Herr Embrik Strand of the
Berlin Zool. Museum, a copy of his paper describing three
species of this group, and he has made for me accurate
coloured figures now in my possession. I would tabulate
them as follows —
Lycaenesthes uhereiuensis, Strand
I cannot regard this as otherwise than a variety of
lernnos ; this species gradually changes as it approaches
Equatorial Africa, and I have no doubt it is Hewitson’s
insect.
The however, is without any question ^ ligures. I
have several exactly the same with their males from the
Victoria Nyanza.
G 2
84 Mr. G. 'J\ Betiiune-Baker’s Revision of Lycaencsthcs.
a.b. imuycrula, Strand.
This I believe to be a distinct species, and is veiy close
to iny cliirinda ; the postniedian band in the primaries is
fractured differently to mine, so that until more material
comes to hand we must leave them distinct standing next
one another.
Lycaencsthcs nibrimaculata, Strand.
This cannot be other than Grose-Smith’s lasti.
Explanation of Plates I — XIII.
[Nee Exphincdion facing tite Plates.]
Explanation of Plate I.
Tran s Ent. Soc. Loivd,.,] 91 0. PL. T.
E.C. Knight del.etlith.
West, Newman chr.
AFRICAN SPECIES OF LYCAENESTHES &c.
I
r
. I
•f
!
1
H
I
'■ -
is-
Explanation of Plate II.
Fig. 1. Lycaenedhes otacilia^, p. 37.
2.
3.
4.
5.
6.
7.
8.
9.
10.
11.
12.
13.
14.
15.
16.
17.
kikmju ^ , p. 37.
sanguiikea ^ , p. 41.
lunulata^, p. 40.
lu'imlata 9 , p. 40.
crawshayi^i p. 45.
lysicles ^ , p. 46.
lachares ohsolescens ^ , p. 48.
lachares ^ , p. 47.
versatilis^^ p. 49.
melamhrohis (1 sex), p. 50
kampala $ , p. 52.
lychnides^, p. 53.
rufomarymata^, p. 54.
radiata^, p. 55.
makala^, p. 58.
leptines , p. 56.
Trans. Ent. Soc. Load.,! 910. PL. II.
•C.Kniglit del etlith.
West.Newman chr.
AFRICAN SPECIES OF LYCAENE S THE S .
.1'
I ^ 1 •
■i
-•/ 11^
Explanation of Plate III.
Fig. 1. Lyc(tenedhes erythropoecilus^ , Y*-
Trans. Eat. Soc. Lon d., 1910. PI. III.
'..Kraght deLecliui,
West.Newmar. chr.
AFRICAN SPECIES OF LYCAENESTHES &c.
3
I
i-
\
I
Explanation of Plate IV.
Fig. 1. Ciopidesthes voltae. Genitalia in profile, p. 8.
„ 2. „ arescopa „ „ „ p. 9.
,, 3. ,, thyrsis ,, ,, ,, p. 10.
,, 4. ,, litJius ,, ,, ,, p. 11.
All the i^rofile preparations of the genitalia have been mounted
with as little pressure as jiossible so as to examine the organs as
near the state of nature as could be. The few vertical views have
been cut and then spread out absolutely flat. All have been
magnified 50 diam., then reduced a third.
Trans. Ent. Soc. Land., IQTO, Plate IV.
(J
1
ANCILLARY APPENDAGES OF CUPIDESTHES.
Explanation of Plate V.
Fig. 5. Lycaenesthes mtisagetets. Profile view of genitalia, p. 21
Trans. Knt. Soc. Land., igio, Plate V
ANCILLARY APPENDAGES OF LYCAENESTHES.
Explanation of Plate VI.
Fig. 10. Lycacnedlies hobleyi. Profile view of genitalia, p. 24.
Tracis. Ent. Sac. Lond., iQio, Plate VI.
s?
S
ANCILLARY APPENDAGES OF LYCAENESTHES.
Explanation of Plate VII,
ANCILLARY APPENDAGES OF LYCAENESTHES.
■-S
►i
;vi
4
•7*
7
Explanation of Plate VIII.
Profile view of genitalia, p. 44.
» ,, ,, P- 45.
5) )) P- 46.
„ M V p. 47.
51 5, „ p. 48.
ANCILLARY APPENDAGES OF LYCAENESTHES.
Explanation of Plate IX.
Fig. 25. Lycaenesthes Jlavomaculatus. Genitalia, profile view, p. 54.
„ 26. „ rufmiarginata „ ,, „ p. 55.
,, 27. „ makala „ „ „ p. 58.
„ 28. „ makala „ vertical view pressed
fiat, p. 58.
I rails. iLiit. ,boc. Lona., IQto, I'laie ja..
■S
ANCILLARY APPENDAGES OF LYCAENESTHES.
Explanation of Plate X.
Trans. Ent. Soc. Loud., ipio, Plate
ANCILLARY APPENDAGES OF LYCAENESTHES.
Explanation of Plate XL
Fig. 33. Nenrypexina lyzanius.
34
5) ))
,, 39. NeurelUpes dandingeri
„ 40. 'Tridema rnfoplagata
„ 41. „ hades
Genitalia, profile view, p. 64.
„ vertical „ flat, p. 64.
,, profile „ 70.
„ » » P- 73.
>) >1 P-
Trans. Ent. Sac. Land.., 1910, Plate XI.
'Cliel. p/iu/o, A. H
ANCILLARY APPENDAGES OF NEURYPEXINA, NEURELLIPES, AND TRICLEMA.
t
Explanation of Plate XII.
Fig. 35. Neurellipes lusunes. Genitalia, profile view, p. 68.
,, 36. „ ,, ,, vertical „ Hat, p. 68.
„ 37. „ dbryseodict’us „ profile „ p. 67.
„ 38. „ ,, ,, vertical „ fiat, p. 67.
95
Trans. Ent. Sac. Land., /gio, Plate XII.
G
ANCILLARY APPENDAGES OF NEURELLIPES.
Explanation of Plate XIII.
Fig. 42. Triclema phoenicis. Genitalia, profile view, p. 76
,, 43. „ kamiUla „ „ „ p. 77
„ 44. „ lamias „ ,, „ p. 80
„ 45. „ faseiatus „ „ „ p. 82
Trans. Ent. Soc. Land.., iQio, 1 laie XIII.
Sr-
ANCILLARY APPENDAGES OF TRICLEMA.
( 85 )
II. On Callophrys avis, C%p'm. By T. A. Chapman,
M.D., F.Z.S.
[Read November 17th, 1909.]
Plates XIV— XLIII.
It is a remarkable circumstance that this very interesting
and pretty butterfly should have remained quite unknown
until so late a date as this, a species somewhat widely
distributed, occurring in France, Spain, Portugal, Morocco
(probably Algeria) * and Tunis. It is not a matter of dis¬
tinguishing or establishing a species previously confounded
with another or regarded as a mere variety, but a species
of which so far as I have been able to ascertain, only some
half-dozen specimens, beyond my own capture.s, exist in
collections, and of these, three, I think, were captured in
the present year. The species, as observed by me, is
extremely localised in each habitat, and this is no doubt
true throughout its whole distribution.
One circumstance about the specimens of C. avis that I
have ascertained to exist in collecbons is somewhat striking,
that is, that they all or nearly all are solitary specimens,
each from a different locality. My two HyO'es specimens
were taken in two different years at two different places.
Then there is one from Tangier in the British Museum,
one taken by Prof. Reverdin on the Riviera, one taken by
Mr. Rowntree at Bussaco, one by Prof. Tavares at Jerez, and
M. Oberthiir’s specimens (two, not one) from Tunisia. But
that my curiosity about the insect had been aroused, there
would probably have been a further one from AmMie-les-
Bains.
The explanation is, I think, simple. Most collectors
more or less ignore C. rnhi as beneath their notice,
o ^ '
perhaps pick up half-a-dozen. C. ruli has habitats by
the square mile. Tlie habitats of C. avis may usually, I
imagine, be measured by acres, or even yards. In picking
up his half-dozen G. ruhi, he may happen to pick up an
odd specimen of C. avis, but before he captures another,
* Under date November 24th, M. Oberthiir tells me that Mr, H,
Powell took two (^s at Khenchela (Province of Constantine).
TRANS, ENT. SOC. LOND. 1910.— PART IL (JUNE)
8G Dr. T. A. < TTapman on CaUoplirys anis.
be lins quite left the C. avis habitat, and that odd one is
all he has. With no clue but my first capture and think¬
ing it was possibly an aberration of C. niM, I failed to recog¬
nise the habitat though I knew approximately where it
was, and in my second year’s search, picked up another in
quite another spot, which unfortunately did not make me
work this second locality, but rather, at the time, confirmed
the idea that it was a sporadic aberration. Not perhaps
an unnatural result, as those were the only two known
specimens.
An account of C. avis involves at many points a
comparison with C. rnln, and it will be found that the
following notes deal largely with C. rubi. I have been
somewhat struck, however, by the circumstance that
many of the details as to G. riibi with which I have to
deal, are more or less new, and, at any rate, have not been
dealt with adequately, if at all, in any British medium.
The original description of Callophrys avis is as follows
(from “ Ent. Record,” vol. xxi, p. 130) —
“ ]\Iuch resemliles Cullojihrys ruhi. C. oris is larger, .32 min. to
36 mm. in expanse ; C. rubi rarely exceeds 32 mm. It has hardly
any trace of tails. The upper surface has a ruddy tint, in excess
usually of that of G. rubi var. fervida, and the venation is often,
especially veins three, four, and five of the upperwing in the s.
marked by rather broad dark lines as if raised, dift’ering from the
narrower flatter lines seen in G. rubi. A marked character is that
the head has a long ruddy fur, replacing all trace of the silver lines
round the e3'es so conspicuous in G. rubi. The androconial brand
on the cJ forewing is triangular, perhaps a shade broader than in G.
rubi, but of only about half the length along the line of the veins,
that it has in G. rubi, in which it is oval or fusiform. The club of the
antenna is red or flesh-colour, all along the lower inner side, a colour
confined in G. rubi to a few terminal joints of the club, and the same
on all aspects of the antenna. There is rather a different shade
of green on the underside, and the white line has quite a different
character from that in G. rubi. It is narrow, but continuous ; it is,
in fact, usually broken bj’ each vein, but looks cbntinuous compared
with G. rubi, in which the line breaks up into spots, rather than
become narrow as in G. avis. Either really, or as an effect of its
narrowness, it has a suggestion of being faintly tinted green. It is
entirely without the dark scales along its inner margin that are so
constant in G. rubi. The portion in each interneural space is curved.
It occupies all the spaces on each wing, from the costa to the space
87
Dr. T. A. Chapman on Calloj)lirys avis.
in front of vein two, but is bright towards the costa (space between
six and seven), and fades towards the inner part of the wing. The
row of spots in C. rubi is much more irregular. The first spot on
the hindwing slopes inwards, in C. rubi it slopes outwards, giving
the second spot the appearance of being displaced inwards in G. rubi,
outwards in G. avis. The appendages have only slight differences.
I have not examined sufficiently numerous specimens to be able to
assert that these are constant.
“ It specialises in its foodphuit, instead of being quasi-omnivorous
like G. rubi.
“Habitat, Southern France (Var, and Pyrdnees-Orientales), Morocco
(Tangier). The only specimen I have seen, not in my possession, is
one in the Brit. Mus. Collection, ranged with G. rubi var. fervida,
and labelled ‘ Tangier, Elwes Coll.’ ”
My original description does not need much amplifica¬
tion, but one or two points deserve notice, perhaps the
most important is in the lengths of the palpal joints, a
reference to Plate XXI will show that in C. avis the second
joint is rather longer and stouter than in G. ruhi, but the
last joint definitely shorter’.
The white line on the underside is really often slightly
tinted green, especially beneath the forewing ; sometimes
it is continuously a line, at others, and more usually, a
little broken up. Sometimes it is nearly a straight (or
curved) line. In other specimens, on the hindwing, tlie
costal spot, for example, inclines inwards, the next three are
nearly in line quite outside this, and the fifth and follow¬
ing spots are again further out. In C. rubi, the spots are
always in a continuous chain, sometimes in a regular line
(or curve), at others with some zigzag.
In G. avis, apart from some variation on the forewing,
all the spots are present and the total variation is com¬
paratively small. In C.-ruhi, as is well known, they vary
to the greatest extent, as to some, more, or all being absent.
The general position of the line is much nearer the base
of the wing than in G. rubi. In (7. avis it is, for example,
on the hindwing very near the end of the ceil; in C. rubi
it is much nearer half-way between the cell and the hind
margin.
The red colour beneath the knob of the ’antenna strikes
one as a novelty compared with the more familiar G. rubi,
and one is inclined to assume that the black area in G. rubi
is scaled. It is, however, the case that the scaled and
88
Dr. T. A. l^iiman on Callophrys avis.
unsealed areas are almost identical in the two species. The
unsealed area corresponds to the red area in C. avis. The
rest of the antenna is of the same colour if the scaling be
removed ; this is also the case in C. ruhi, the difference
between the two antennae, denuded of scales, being the
blackness in C. rubi of the black part of the unsealed
portion (Plate XIV).
The face differs less than one at first supposes. C. avis
has rather more and longer liairs, but the great difference
is the absence of the silver lines of ruhi (from which the
genus is named), these scales being red in G. avis.
The brilliance of these lines in C. ruhi makes the hairs
curiously inconspicuous, an effect aided by their duller
colour, which varies from grey (sometimes almost brown)
to black (Plate XIV).
In regard to the dark margins to the white spots in
C. ruhi, it is worth noting that in the genus Ncolycaena,
there is a very similar row of white spots with very similar
dark margins. Taking into account the very similar
androconial brand, there is no doubt that hail Neolycaena
had a green undenside (after all a really trivial difference),
we should have considereil the species to belong to Callo-
2^hrys. Staudinger places the Neolycamas in Theda,
though he separates ruhi under Callophrys. Curiously
Bingham retains Neolycaena amongst the “Blues,” although
Bethune-Baker, so long ago as 1892, showed (in our
Transactions) their true position. Callophrys, Neolycaena,
and Thestor iorva a very natural tribe (7’/;e.s’to?’hf'i). 1 would
venture to predict that the larva of Neolycaena will be
found to possess a honey-gland. I have noted the brand
in C. avis as triangular, really it is polygonal and might
almost as well be called circular. Its lower margin is
rather angular. The relative measurements in the two
species are; longitudinal, i. e. in line of veins, 0. ruhi, 1'7
Tum. ; C. avis, 1'2 inm. ; transverse, C. ruhi, 0‘75 mm.; C.
avis, 1'05 mm.
It will be seen on reference to my figures (Plate XVI),
that the difference in the form of the brand is not so much
due to a difference in the portion of the wing occupied by
it, as to a marked difference in the arrangement of tlie
veins underlying it — a difference not easy to describe but
obvious on inspection. These figures are rough and dia-
"rammatic, but are nevertheless sketched under camera,
and may be trusted as correct on this pomt.
89
Dr. T. A. Chapman 07i Calloj)hrys avis.
The androconia (Plate XVI) of both species are parallel¬
sided with rounded extremities, those of C. avis are
apparently rather more loaded with pigment. There is,
however, a very notable difference in form between them.
Those of G. avis are short and broad, of C. ruli long
and narrow, i. e. compared each with the other.
The following figures are the result of measuring
thirteen scales of each, taken at random, just as they
came. They are of course not so accurate as if a much
larger number were taken, but are probably not far out,
as they agree with one’s impressions on looking over a larger
number.
The relative measurement of the palpi (Plate XXI,
fig. 1) in the two species is interesting, especially in view
of Rambur’s note on his Andalusian specimens of C.
Tuhi (?).
The actual lengths of the three palpal joints in the two
species are in millimetres.
The second joint is also slightly more robust in G. avis.
In G. rubi its end is nearly transverse ; in G. avis it is
sloping, so that its upper margin has much the aspect of
ending in a spine. In both, the end of the third joint
looks truncate and has the well-known terminal tube.
The male ancillary appendages (Plate XX) are very
similar, quite within limits that are exceeded by mere
variation in many species, but we know that in several
groups of Lycaenids, specific and even generic, differences
are often very small.
In this case all the G. rubi examined absolutely agree
90
Dr. T. A. il?rij)mau on Collo'plirys avis.
together, and the one C. avis I have mounted differs in
several points from these.
The proximal ventral projection (saccus) is in G. avis
broader but with straighter sides, and therefore sharper
point, than it is in C. ruin. The dorsal hooks are at their
bases suddenly bent rather than curved and are thicker,
especially towards their ends ; their to.pering extremities
being shorter in C. avis than in G. rubi. The exserted
portion of the shaft of the aedoeagus is rather more slender.
The dense portion of the clasps is very distinctly more long
and slender in G. avis (as 7 to 6).
In the medio-dorsal line the proximal margin is a sharp
angle in G. avis, an open arch in G. ruin, whilst the distal
margin is narrower in G. avis and the denser chitinous
elements more solidly knitted together.
The preparations I have made of the female abdomen
(Plate XXT, fig. 2 ; Plate XXII) are far from being as satis¬
factory as I should like, but one or two points of difference
between the two species, G. avis and G. rubi, are evident
enough.
The long chitinous tube in the eighth segment (corre¬
sponding to the very long aedoeagus of the male Theclid),
is narrower in G. avis (two specimens examined) especially
towards its lower end, so that in G. avis it gradually in¬
creases in width from its lower end forwards. In G. rubi
the narrowest part is about the middle, thence enlarging
towards either end. The tube is also distinctly longer in
G. rubi than in G. avis, though I have a specimen of rubi
that approaches G. avis more nearly than the one photo¬
graphed. The two remarkable spinous organs in the wall
of the bursa seem to be nearly alike, viz. each with two
long sharp spines ; in G. riibi there seems to be much
variation in the length of the second spine ; in G. avis they
are both equal, but an examination of further specimens
may (or may not) show them to be as variable as in G. rubi.
The reason for supposing they are not so variable in G. avis
is, of course, that in other characters in which the two
species differ, G. rubi is always the more variable.
I took my first specimen in the little range of hills
between Hyeres and Carquieranne ; of these hills the
best known is the Gollinc, des Oiscanx, near Costcbelle, and
so, assuming this fly to be one of the birds for which that
summit is famous, I have given it the specific name of
Avis.
91
Dr. T, A. Chapman on Calloplir.ys avis.
My interest at the time in Callophrys rnti was stimu¬
lated by Mr. Tutt’s impending treatment of that insect in
his “ British Butterflies,” in connection with Avhich he
expected me to supply him with various important
observations. Possibly without this stimulus the specimen
of C. avis would not have been taken. It puzzled me very
much ; I hardly thought it could be a distinct species, but
there were almost greater difficulties in supposing it to be
a hybrid with some other Theda or an aberration of C. rnhi.
The specimen is noted in Tutt’s “ British Butterflies ”
(vol. ix, p. 90) under the head of C. rnhi, C vav. fervicla.
“ A very remarkable example of this form, taken by Cliapraan at
Hyhres in April 1906, is very large, 36 mm. ; has a very narrow and
yet almost continuous white line on the undenside of the wings ;
has the underside of the antennal club largely red, a character that
appears to be very rare, judged by our long series, and we believe
not before noticed by any lepidopterist.”
The colour of the upper surface would lead to its being
placed with G. onihi var. fervida, but no notices of fervida
I have come across report specimens possessing the
peculiar characters of C. avis, and the localisation of C. avis
and its rarity even where it occurs make it unlikely that
many specimens exist unrecogni.sed in collections. Even
in his great collection, where one would expect to find it,
if anywhere. Mods. C. Obertbiir tells me that he cannot
detect one.
Rambur’s note on Theda ridn in his “ Catalogue of the
Lepidoptera of Andalusia” leaves on my mind little doubt
that some, at least, of his specimens were C. avis. He
says the specimens are distinguished “ by the ochreous of
the upper wings and by the continuity of the white line
beneath the lower ; the last joint of the palpi is also
shorter.” The last item at least is definite, and it is the
case that this last joint is shorter in C. avis than in C. rnhi
(see ante and Plate XXI).
Staudinger’s var. suaveola of 0. rnhi agrees with
C. avis in its large size and in the want of tails. In all
other respects it agrees with C. rnhi. I have examined
two specimens at the Brit. Mu.s., South Kensington, from
Artrabad and Shahkuh, and one from M. Oberthiir’s col¬
lection, from Syria.
The last palpal joint is more like that of C. rnhi than
92
Dr. T. A. (#lhpman on Gallo'phrys, avis.
of C. avis, and the green of the underside lias a little of
the tone that distinguishes C. avis, but a similar tone is
not uncommon in 0. rnhi. The underside spots have the
dark scales at their inner margins as in C. ruhi. On the
meagre amount of material examined, I do not feel at all
positive that suaveola is not perhaps a species, distinct
from C. ruhi, as it certaiidy is from C. avis.
The var. fcrviila of C. ruhi, with which I was at first
inclined to agree with Mr. Tutt in placing my first odd
specimen of C. avis, agrees with C. avis only in approach¬
ing its ruddy tint, 1 have seen no sj)ecimen reaching the
rich tones of a really line G. avis', and in a little approach
also in many specimens in the tint of green on the under¬
side — in all other respects it differs from G. avis as does
ordinary G. ruhi. It was the prevailing form of G. ruhi
at Amelie-les-Bains.
I have not met with any further reference to the species,
except my own note on the capture of my second Hyeres
specimen in 1907, which appeared in “ Ent. Record,” vol.
xix, p. 152, until I described the insect as a new species
in the “ Ent. Record,” vol. xxi, p. 130 (see ante), and ex¬
hibited specimens at the Society’s meeting on June 2nd
last (Proc. 1909, p. xxix).
I was enabled to do this in consequence of having taken
a few specimens in the Pyrenees-Orientales (at Amelie-les-
Bains), and luckily observed a ovipositing, and thus suc¬
ceeded in obtaining eggs, and have reared the insect, up to
the present, of course, only so far as the pupal stage.
Since the description of the species was published Prof.
Reverdin tells me he took a specimen some 20 kilo¬
metres east of Hyei'es, and I have seen a specimen in the
collection of Mr. A. S. Tetley, of Scarborough, taken by
Mr. Allan Rowntree at Bussaco, Portugal (within the old
monastery pi'ecincts), on May 27th, 1904. The specimen
is a ^ of 34 mm. expanse, in fair condition, but sufficiently
worn to have perhaps been sometime on the wing. Still,
the date seems later than one would expect, but the
Atlantic side of the Peninsula probably does not warm up
in spring so rapidly as the Mediterranean littoral.
The specimen is very interesting as showing a wider
distribution of the species than merely the western end of
the Mediterranean basin.
This Portuguese specimen led to my writing to my
friend Prof. C. Mendes at Sao Fiel, asking him to look
])r. T. A. Chapman on CaUo'phrys avis. 93
over his series of C. ruhi in search of any intrusive
C. avis.
He writes me that he “ has found one specimen taken by
his colleague Prof. Tavares in May at Jerez. It is one of
two specimens, of which one is C. ruhi, the other C. avis,
with the hindwings somewhat worn. The distinction is
perfect. I have also examined more than twenty speci¬
mens from the neighbourhood (Sao Fiel), from Lisbon and
from Algarve, all of which are G. o'lchi.”
Spain can thus be added to the habitats of C. avis.
On October 26th, 1909, M. C. Oberthiir wrote me that he
had just received from Tunisia two specimens that appear
to be G. avis, and later, after actual comparison, he tells
me that they certainly are G. avis (2 ^ from Aiu-Draham
(Kroumirie), Tunis.
Unfortunately, by the time I liad “ spotted ” my butter¬
fly at Amelie-les-Bains the weather became dull and
unsettled, so that I can say little of its habits. It can fly
strongly in good weather, but does not stray far from its
proper habitat. It affects (for feeding probably) flowers of
a species of Etqihorhia, where it is sometimes accom¬
panied by specimens of C. riibi. Owing to weather, I
had no opportunity of seeing it orient itself in the sun
as G. rubi does.
The butterfly I saw ovipositing on April 20th was lay¬
ing her eggs on the flowers (?), calyx really, of Goriaria
myrtifolia.* I found at this date that two $ $ that I had
placed on Gytisus spinosibs (or closely allied species), which,
by their joint distribution, seemed to be the chief food-
plant of G. rubi at Amelie, three days before, had laid
only two eggs. They began to lay at once when given
Goriaria. Un April 30th an egg laid wild, hatched ; eggs
laid in confinement began to hatch on May 8rd.
The following are my notes of experiments on the food-
plants of G. rubi and G. avis. They show that G. rubi
would not touch, or only sparingly and ineffectually, the
foodplaut of G. avis. Whilst G. avis absolutely refused all
the ordinary pabula of G. rubi.
May 26th. — Placed some newly- hatched larvae of
G. rubi, j^arents taken amongst Goriaria at Vernet-les-
Bains, on Goriaria, and found that they refused for 24
* The local botanists (Messrs. Raine, Powell, Jehandiez, etc.) say
that Goriaria myrtifolia does not grow near Hybres, it follows that
C. ori.s must there subsist on some alternative foodplant.
94 Dr. T. A. o% CalloplirTjs avis.
liours to touch it. When jhaced on Ulex europaeus flowers
they at once settled down and soon began to feed.
H aving done so twice before, I again placed a half-
grown larva of 0. avis on mixed flowers of Ulex europaeus and
Cytisus scoparius, and another on flowers of Cornus san-
guinea. Left for 24 hours the larvae on each occasion starved.
Larvae of C. ruhi, newly hatched, placed on flowers of
Cornus sanguinea, went for the open flowers and did some
eating ; next day, however, they had eaten very little, and
were very restless, three of them being down on the stem ;
they became quite pacified and settled down when given
blooms of Ulex.
Larvae of C. rubi placed on Heliantheimiin at once
attacked the flower-buds and appeared to thrive on them ;
one then placed on Eroclimii attacked a flower-bud, but,
that demolished, and there being no other, it and a second
larva starved rather than touch the leaves or any other
part of the plant. Larvae given buds of bramble, made
deep holes in the buds, but after a time and after seeming
to be quite at home, left them and seemed very discon¬
tented. First-instar larvae placed on Goriaria. starved
rather than eat it ; last-stage larvae ate it sparingly, but
in two days were smaller rather than larger, got on a little
better for a day or two, but then were obviously doing so
badly that they were restored to broom flowers.
C. avis refused any of the ordinary flowers loved by
ruhi, such as gorse, broom, Sp)artiuvi jttnceum, etc.
I fed my young larva on the flowers, and they seemed
somewhat inditferent what portion they ate, but as the
male flowers rapidly become useless on the stamens burst¬
ing, I found them to take more especially to the female
flowers, clearing out the immature fruits. When they
were in their second instars I found them eating such
portions of leaves as I had given them with the flowers,
and on giving them leaves they seemed to prefer them to
the now not too tender flowers (or rather fruits). Thereafter
I fed them on leaves, and I owe many thanks to Monsieur
Boixo, the “ Roi du Canigou,” for sending me supplies of
leaves on which to feed my larvae after my return to England.
The last larva hatched on May loth, and on May 18th
the first larva is supposed to be in third instar.
In the first instar the larva is a greyish green, or even perhaps
cinereou.s, but pale and in a sense colourless, tlie head black and the
95
Di'. T. A, Chapman on CallofUrys avis.
hairs also, except just at emergence. The youngest larvae are now
(May 18th) somewhat grown and are of a pale brownish tint, consist¬
ing, however, of the effect of the sparse black hairs, with a skin in
two colours. The black hairs are the long dorsal ones, the, nearly
as long, lateral ones, are pale or colourless.
Tlie ground-colour is a light ochreous yellow with a trace of green
and the reddish-brown markings are a light or diluted maroon.
The prothoracic plate is dark but not black, the anal plate darkly
tinted but pale by comparison with the prothoracic plate. The
reddish markings are a line down each side of the dorsum, not quite
central to the dorsal flange but rather to its inner side, and slightly
interrupted at the incisions. A similar subspiracular line, leaving
the dark dots of the spiracles on a pale line or area above it. Above
the spiracles a line or band broadest at the front margin of each
segment, leaving another pale area between this and the subdorsal
line, somewhat triangular on each segment, from being broadest at
its posterior margin, complementarj- to the red line beneath it, and
centrally in this pale area a triangular red mark, with its posterior
margin not quite coincident with the posterior margin of the seg¬
ment. On dorsal view the prothorax often looks dark (pale Indian
ink) from the black head retracted into it. The larva is of the
usual onisciform shape, about 2 mm. long and 0'7 mm. broad, fairly
equal in width, but perhaps slightly tapering backwards, the ends
little more than semicircles. On end view, the lateral flanges are
marked, but the dorsal ones are the rather sharp margins of a flat
dorsal area.
In the second instar the larva is at first reddish in general tone,
about 3'5 mm. long. The ground-colour is a green slightly toned
with yellowish brown, but the greater part of the surface is occupied
by pale reddish brown, viz. two brown lines (or bands) on each side
of the dorsal line (left of the yellowish green ground-colour) and
just inside tubercles I and II. The broad brown flange line below
the spiracles, along the prominent bosses of IV and V forming the
flange. The spiracular level is of pale ground-colour, but above it
is a darker area, higher on the front of each segment, leaving a pale
area narrow in front and broader behind, above it containing a
darker spot in its centre.
As it grows in this instar it becomes wholly green, the change in
size and colour being so remarkable that I had some difficulty in
satisfying myself that an unobserved moult had not taken jjlace. The
length reaches 4'5 to 5'0 mm. The previous markings are just
indicated in darker green. The head is dark except the mouth
parts and just above the labrum.
In the third instar (in which were still a few on June 3rd) the
96
Dr. T. A. on Callophrys avis.
larva attains 12'0 nun. when stretched in walking with head ex¬
tended ; the width about 3'0 mm., and of head about 0’6 mm. Tlie
head black, except where the vertex is usually within prothorax,
labruni whitish. The rest of the larva is wholly of a transparent
green, except a very narrow lateral line, yellow, and the prothoraeic
plate of a purplish pink, with a faint pale median line. The s])iracles
are darker dot.®, apj)arently very narrow rings of deep brown.
When about to moult the larva spins a few threads to surrounding
leaves as well as a silken pad.
In the last instar when full grown (Plates XV, XVII, XVIII) the
larva Avhen active attains a length of about 20 mm., but when resting
or sulky only some 15 or 16 mm., but is then some 5'5 mm. wide, and
5 mm. high ; short and thick, thickest about first and second
abdominal segments.
The larva is of a uniform transparent green, with a yellow lateral
line and a brilliant red (purplish pink) prothoraeic plate. The
depression of the prothoraeic plate is conspicuous, lying deeply
under mesothorax with the front of the prothorax rising above it.
The broad red stri})e across the prothorax, and the uniformity of
all the larvae in the light transparent green colour, broken only by
the plate and by the thin lateral line, render the larvae conspicuously
different from any other I have seen.
The red stripe is like nearly all colouring in Lycaenid larva, a
little way below the cuticle, and though one supposes at first it is a
colouring of the prothoraeic plate, this is not so, as it e.xtends widely
to each side beyond the plate, almost right across the prothorax.
Oidy one larva varies from this, in having a broad lateral rusty red
band, obscuring the yellow lateral line, which seems deeply buried,
but has some of the rusty tinting at the incisions. The rusty tint
is superficial and involves the whole flange practically up to the
spiracles. The prothorax is ruddy tinted and there is a slight tint
on the dorsum of the seventh abdominal segment.
I copy tlie above as it appears in my notes, but furtlier
notes show that this larva was in the first stage of the
change of colour that is undergone when the larva is pre¬
paring to search for a pjlace for jrupation.
The larva never goes anywhere without making a silken
path and often quite a pad, on the leaf it is eating. When
recently moulted into the last instar, there is quite a
ruddy halo due to the closeness of the reddish hairs.
When the larva is full-fed the hairs become so far separated
as to be hardl}^ visible.
On June 7th, I noted that on each of the past three
Dr. T. A. Chapman on Callophrys avis. 97
days some larvae have assumed a darker tint, gradually
deepening to a dark olive green, almost black, as com¬
pared with the bright light green of the feeding larvae.
They have the yellow lateral line changed into a
red, almost brick-red one, looking dee23ly sunk in the
tissues.
In looking for a place to pupate, the larvae of C. avis
were quite disconsolate when I provided them with some
loose earth with a little loose rubbish on top, such as I
had found suited G. riibi admirably. With such provision
G. o'uhi goes out of sight and pupates, if not strictly under¬
ground, still beneath the loose surface material. On
providing G. avis with bits of stick, leaves, paper and so
on, I found tliat after a period of wandering, that is
common to so many larvae that have to travel often some
distance in search of a puparium, they settled down, on
some object an inch or two above the surface, sometimes
in a little hollow, sometimes between two surfaces such as
two dead leaves, etc., and here did some slight spinning,
amongst which some few threads were usually disposed,
though often separatel}^ as a girth, and there was also a
more or less distinct anal pad. Not one example \vent
into even the most superficial rubbish.
In comparing the larvae of C. avis and C. mhi, we find that in
C. rithi, in the first instar wlien fairly grown the larva has the white
patches below the dorsal line Ijroad, very white (not ochreons) and
without any brown patch in it on each segment as in C. avis.
In the second instar (June 2nd) it looks not unlike C. avis in the
same instar, but is a little greyer ; as it grows it loses all likeness to
first-stage larva, at first rather striking, and becomes green, but with
very marked darker and lighter green on pattern of the brown
and white, and reaches a length of 6 mm.
In the third instar (June 8th) (Plate XXXII), it is green with a
very marked pjale dorsomedian line, dorsal flange line and oblique
stripes, these are quite yellow in some specimens, pale green in others,
but seldom quite as bright as the yellow lateral line. The hairs are
bltick, not quite so in some, but without the rich ruddy tint
they have in avis in the later instars. The prothoracic plate is
clearly seen brownish-black, the colouring being confined to the
plate itself. Length 10-12 mm., according to how much extended.
The larvae appear frequently to eat the cast skins and
apparently the heads with them, if one may judge by the
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE) II
98
Dr. T. A. on Callophrys avis.
difficulty of finding tlie cast heads. A circumstance that
adds mucli difficulty to counting the moults.
When the young (first instar) larva of G. ruhi attains its
colour, ■which it does a shade earlier than does G. avis, the
colours are not pale ochreous green and brown, but a
white that is almost that of porcelain and a more ochreous,
less red, brown. The dorsal brown lines are broader.
The white below it looks broader and perhaps is, but its
greater effect is more due to its whiteness, aud to its
wanting the brown patch on each segment, the sub¬
dorsal hairs (ill ?) are visible with a hand lens easily,
being black as are the lateral (llange) hairs (colourless
in avis).
In the last instar, the uniform green tint of G. avis con¬
trasts in its light transparent tone with that of G. rubi
(Plate XV), in the same stage, and in its uniformity, quite
unbroken by any trace of the oblique stripes, which are
always more or less present in G. rnbi. Any appearance
of stripes in G. avis is the effect of the illumination on
the irregularities of the “ upholstering ” of the sides.
G. avis has its yellow lateral line and its red mark across
the prothorax, but no other colouring, aud each larva is
like every other without any variation. G. rubi always
has the oblique stripes, sometimes as a different tone of
green, sometimes as more or less bright yellow, and
sometimes further emphasised by some red margins,
making a very handsome and gaudy larva. I did not
meet with this brilliant form this year, so have not got it
figured. An essential point, moreover, is not merely the
j)resence of these stripes in some form, but the great
range of variation that the larva presents, as compared
with the uniform G. avis. It never has any approach to
the red thoracic mark, and may even seem almost without
a thoracic plate ; the mark here in G. rnbi being merely
the very small central brownish portion of the plate,
Avithout any further colouring.
Another marked difference between the two ' species in
the last instar, is the much greater dorsal swelling of the
segments and the correspondingly greater depth of the
nicisions dorsally in G. rubi. So that on lateral view,
G. rubi has a very serrate dorsal outline, as compared
with the tolerably smooth aspect of G. avis, the latter
might, in I'act, wlien compared with G. rubi, almost be
taken for a Chrysophanid like G. rutihis.
99
Dr. T. A. Chapman on Calloiihrys avis.
Callophrijs avi.‘i lias a pupa (Plate XIX) very similar to that of G.
a little larger and stouter, length 10 mm. ; measurement does not,
however, show that it is any stouter. In this living pupa the pale
(brown) ground-colour seems more easily seen and more abundant
than on ruhi, and the two tints brown and black seem more dis¬
tinctly separated. The abdominal dorsum distinctly looks paler than
in ruhi, owing to the gi'eater amount of brown ; this may easily,
however, be a matter of racial or even individual variation.
The spiracles (abdominal) are more obvious in 0. ar is, being nearly
white on a very small brown area, whilst in C. rubi they are brown
on a (usually) rather larger brown area. The hair and surface
sculpturing present no differences to be detected in the living pupa.
The cremaster appears to possess a much larger number of hooks,
corresponding with the more frequent occurrence of a practicable
girdle.
On August 26th, 1909, I overhauled pupae of C. avis
and C. rubi. Those of C. avis have wings very black and
opaque ; those of rubi are brown, in some, suggesting
that there was clear fluid under a brown skin, but in
others (to my surprise) the brown was paler and opaque
suggesting wing-development going on, and in these the
eye-spots were also blackening. The eyes and wings of
C. avis were too black and opaque to make it probable
similar changes, if going on, would be visible. Kept several
advancing rubi up in warm room to see if they would
force, but without effect after several weeks. With hardly
an exception the C. rubi were all on the earth at the
bottom of the jars, or even in if not below it, and without
an adhesion to anything. The C. avis were equally away
from the ground, an inch or more, and with some attach¬
ment in man}^ cases, and in one or two with something like
a distinct girth. The decay of material to which they
were attached made this impossible to verify in probably a
good proportion of cases in which it had existed.
The egg (Plate XXIII, et seq.) is laid in the flowers of
Coriaria my7'tifolia, those I saw laid naturally on the calyx of
the male flower, and providing them with similar material I
found those in captivity did the same, varying a little by
laying occasionally on the stamens, as well as on the short
flower-stalks, and even on the stems at the base of the
flowering racemes.
In no case was an egg attached by anything but its
lower surface, and though put to some extent out of
II 2
loo
l)r. T. A. ^apnian on CaUo'phrys avis.
sight was not often wedged into any sort of crevice, differ-
ing therein much from the egg of C. win, which is almost
as a rule pushed in between two surfaces, and adheres by
its top to the upper one, only a little less than by its base.
The ejig is extremely like that of G. ruhi (Plate XXIII) ; it may
he a shade larger, 0'7 mm. in diameter, or sometimes rather wider. In
colour it is like G. ruhi, green, and in some respects nearly white,,
but on the whole it is less green and has usually a whiter appearance.
The sculpturing is almost alike. The actual egg is in fact green, but
the superficial adventitious coating is white, and differs from that in
G. ruhi in consisting not merely of the columns and ribs, but has
also a slight laj'^er over the bottom of the cells that seems to be
wanting in G. rxhi.
In photographing the micropyles of the two species, Mr.
Clark found such a difference in the nature of this adven¬
titious coating, that he made various prej^arations and quite
a research to elucidate the differences. He notes that
“ 0. ruhi has apparently a single coat, which is quite
transparent when mounted in glycerine ( Farrant’s medium),
and shows the network structure of cells perfectly, either
in this way or when mounted as a dry object.” On
the other hand, “ G. avis has apparently two coats, tlie
outer one bears the sculpture of network, is granular,
especially under pressure, and separable from the inner
coat ; the latter being transparent and devoid of any
special structure or trace of cell formation. The sculp¬
ture is fairly well seen mounted as a dry specimen,
whilst in fluid (Farrant’s) it becomes a granular amor¬
phous mass.” He also notes that “ the basal portion of
these eggs is of simple cells ” (i. e. without adventitious
coat), “ which terminate abruptly when the side of the egg
is reached.” This is the case in all Lycaenid eggs I have
examined, though the cellular structure of the base is often
so faint as to require some looking for. In Calloplirys it is
very distinct. Mr. Clark also notes that “ the micro pylar
structure is best seen when mounted and pressed quite flat
in Farrant’s medium. I could detect no ‘ pores ’ as is the
case in some eggs.”
The adventitious coat of Lycaenid eggs, as a rule,
breaks up into an amorphous pulp when treated with
any medium such as alcohol, benzole, glycerine, etc.,
so that it is useless to try to preserve or observe them
101
Di'. T. A. Chapman on Callophrys avis.
in any way but as dry objects. Mr. Clark’s observa¬
tion on C. avis eggs shows tliat it follows this usual rule,
and in glycerine it is at once seen that the eggshell has
two layers — the true shell which is persistent and the white
coating which disintegrates. His remarkable discovery is
that in the egg of C. ruhi the adventitious coat differs
entirely from that of C. avis, in some chemical or other
. quality, as to be uninjured by Farrant’s medium, and acts as
if the true eggshell and the coating were one homogeneous
layer. One would hardly expect so considerable a
difference to exist between two so closely allied species.
I ought perhaps to say, though it is carrying coals to
Newcastle, that I do not report Mr. Clark’s discovery
without having repeated and verified it m3’^self. As per¬
haps having some connection Avith this point, I may
observe that an egg of C. avis is easily obtained, pure and
simple, an egg of C. ruhi has nearly always attached to it
some hairs or filaments belonging to the plant on Avhich it
Avas laid (see Plate XXIII). A corollary no doubt of the cir¬
cumstance that the egg of C. avis is attached merely by its
base, Avhilst that of C. rubi adheres also, if possible, by
some part of the upper surface. This is probably asso¬
ciated with a different composition of the white coating,
a difference revealed in another Avay by Mr. Clark’s
experiment.
The eggshell of G. ruhi comes out little if at all in¬
jured after successive treatment by water, by alcohol, and
by benzole, that of G. avis is decidedly damaged both by
alcohol and by benzole, but by Avater it is much altered,
and the water around it exhibits a multitude of minute
colourless spindle-shaped bodies of about O'OOl mm. in
length.
In all cases, the base, Avhich has no coating, is unaffected,
and retains its network unaltered.
Plates XXV and XXVII shoAv the eggshell of G. rubi Avith
the mesh of ribs arranged in triangles, so that they Avould
obviously fall into hexagons but for the exigencies of a
curved surface. The columns are end on, and so do not
.^ihoAv their height. The double outline of the ribs is, no
doubt, an optical effect in the dry shell of the empty egg ;
it is seen also in G. aws and in other Lycaenid eggs. Fig. 2
is a portion of the eggshell of G. avis dry, and not
pressed.
In Plates XXVI and XXVII Ave have three specimens of
102
Dr. T. A. <^Phpman on Callophrys avis.
the eggshell of C. avis showing the inicropylar area, and
showing also the effect of pressure in Farrant’s medium
in destroying the meshwork of the surrounding area, and
in the example of C. rnli (Plate XXVII, fig. 1) it appears
unaltered by this treatment, except a little distortion by
pressure and some blurring due to the high magnification
( X 350) making its thickness too great for being in focus
tluoughout. Though Mr. Clark says there are no pores
visible, I think I detect them in the preparations from
which Plate XXVII, fig. 2 {avis) and same Plate, fig. 1 {ruli)
are taken. They have probably become more evident by
lapse of time, as often happens to objects in Farrant’s
medium. In Plate XXVI, fig. 2, there are five cells to the
inicropylar rosette (four and six respectively in the other
two figures of C. avis), and apparently five pores ; but it is
also the case that in the specimen of C. rwii! (Plate XXVII,
fig. 1), which has seven cells to the rosette, there are
equally five pores, arranged most regularly in the centre.
The breaking down of the adventitious coat, especially in
Plate XXVi, fig. 2, and Plate XXVII, fig. 2, in G. avis
makes the distinction between the cells of the microiiylar
area (without adventitia) anA those surrounding it very dis¬
tinct, and shows that two rows with a few odd ones surround
the rosette. Though the demarcation is not so fully
demonstrated in the preparation of C. rubi, it is neverthe¬
less very obvious, and more so in the actual preparation
than in the photograph, that there are about four rows of
cells round the rosette in that species, and that as the
cells do not differ much in size, the area (or “depres¬
sion ”) is larger in C. ridn than in C. avis. In dealing
Avith the details of hairs, lenticles, etc., of the larva at
different stages, I may refer to my notes on C. rubi in
Tutt’s “British Lepidoptera,” vol. ix, p. 104 ct seq.,
which are fairly full, and also to the photographs by Mr.
F. N. Clark, accompanjdng this paper, from specimens I
have prepared. This will save much verbosity.
In comparing the larvae in the first instar, C. avis
(Plate XXVIII) seems smaller and more delicate (though
the egg seemed, if anything, larger), certainly it has a smaller
head, and certain hairs, present in C. rahi (Plate XXIX),
are wanting and others less developed. The most notable
of the absent hairs are those (ill ?) between the sub¬
dorsal lenticles and the spiracle. Of these C. ridn has two in
each of the abdominal segments one to seven. They are
103
Dr. T. A. Chapman on Calloplirys avifi.
well-developed hairs, nearly half as long as the long ones
of tubercle i. These are wanting in C. avis absolutely in
all of nearly a dozen specimens examined, except in one
that has a hair here on the first and second abdominal
segments, and on one or two others,
Another hair well developed in C.ruhi, absentia C. avis,
is one in front of the long dorsal hair (i ?) on seventh
abdominal segment. The short hair in front of I on the
abdominal segment is little more than half tlie size in
C. avis that it is in C. rnhi.
Plate XXVIII is taken from the specimen of C. avis in
which the supra spiracular hairs (ill ?) happen in a few
cases to be present.
The prothoracic plate has four short hairs along the
front margin in C. rnhi that are wanting in C. avis. This
is in accordance with the general stronger development of
hairs in C. ruhi.
In the first instar there is no trace of a honey-gland.
In the second instar (Plate XXXj less difference
between the panoply of hairs is discoverable than in the
first, between the two species. C. rtihi (Plate XXXI) is,
however, rather more strongly haired. The longest hairs
are about 0'37 mm. long in C. avis, and few reach this ;
the corresponding figure for C. ruhi is 0‘‘i2 mm. The two
photographs (Plates XXX and XXXI) give a not very
unfair indication of the difference between the two,
although it is much emphasised by accidental differences
in preparing the specimens.
In these photographs, as in several of the others, it is
necessary to point out that in Plate XXXI the prothoracic
plate is properly displayed, in Plate XXX its posterior
margin is bent under.
In both species there is in this instar a first indication
of the honey-gland on seventh abdominal segment, a slight
failure of the ordinary hairs appear, and there are three or
four lenticles in line, marking the posterior lip of the
opening, which I suppose is always present, though I have
only managed so to speak to “ glimpse ” it in two or three
specimens. Each pad of the prolegs and claspers, which
in the first instar had two hooks, now has five — two longer
and three shorter.
To complete the view of these crochets it may be noted
that in both species eleven seems to be the number of
hooks to each pad in the third instar, eleven on one pad
104 ])r. T. A. (^^pinan on Calloinlirys avis.
and nine on the other being not uncommon, a larger
number, twelve or thirteen, is rare ; they alternate large
and small, with small ones at each end, sometimes the end
of each series is smaller than the middle ones.
In the last instar they are much more numerous but
much alike and variable in the same wa}'’ in the two
species ; twenty-one to twenty- three is about the average ;
one may find twenty-three on one pad and seventeen on
the other, and as many as twenty-five occur — sometimes
the front, sometimes the posterior, pad has the fewer
crochets. They are alternately longer and shorter, but the
longer set often varies again into longer and shorter.
In the third instar (Plate XXXII) the hairs are again
just appreciably longer and stronger in C. ruhi (Plate
XXXIII). The honey-gland (Plate XXXVI, fig. 2) is now
quite distinct in both species by a flight of lenticles
along both lips and by the recession of the principal
hairs.
In the last (fourth) instar, the honey-gland (Plate
XXXVII) is very obvious; a space is made for it by the
loss of the longer dorsal hairs. The appearance shown in
Plate XXXVII, fig. 1, of four circles right down in the
fjland, is one I am familiar with in the larvae of “ Blues,”
but what precisely they represent, functionally or homo¬
logically, I do not know.
I suspect them, however, to represent the two pairs of
dorsal hairs (1 and II) changed into glandular structures.
The hairs at all stages (if we except the curving in the
first stage) are simple and straight and armed with fine
spicules ; in the last stage, however, these have so
dwindled that it may be said they have disappeared,
traces of them are more easily detected in C. ruhi than in
C. avis.
At all stages the bases of the hairs are spread out and
divided by lines, so as to take the floral aspect they have
in many other Theclids.
I have already noted the extra hairs that C. ruhi has on
the prothoracic plate in the first instar. In the later
instars (Plates XXXIV, XXXV, and XXXVI) the hairs and
lenticles of the 2:»rothoracic plate vary so much that it is
difficidt to find two individuals with precisely the same
dispositions, so that the differences one may note in a few
specimens between C. avis and C. ruhi are more ^mobably
individual than specific. The special angular hair does
105
Dr. T. A. Chapman on Calloplvrys avis.
not declare itself in the first instar, but afterwards is
present equally in both species and is well seen in the
photographs of those of the second and third instars
(Plates XXXIV and XXXV).
In the last instar (Plate XXXVI) they are not so easily
found, as the outer angle of the plate on which they are
is not at all chitinised, and is, in fact, like the rest of the
skin. I believe I am correct in saying that the red mark
in C. avis goes far beyond the plate, but I may not have
made enough allowance for this non-chitinisation of its
lateral wings. In Plate XXXVI, fig. 1, the special hairs
would be about half-an-inch outside the picture.
So far as I know the existence of a honey-gland in the
Thestoridi has not been previously reported. It
certainly escaped the notice of both Mr. Tutt and myself,
both in his researches in the literature of the species and
my (I fear rather perfunctory) observations of the living
larvae. There is no mention of it in Tutt’s account of
Calkyjhrys ruhi in his “ British Lepidoptera,” vol. ix.
I may add that I have searched for indication of the
fans or scent (?) glands of the eighth abdominal segment
and have not succeeded in discovering any ; I incline to
believe they do not exist, still, if they are rudimentary,
they might be present and easily elude me.
So far I have had no empty pupa shell of C. avis to
mount. The only one I have mounted had to be cleaned
by some maceration in potash, which has had the effect of
softening the hairs and rendering them useless for de-
scrijDtive purposes. In other respects I believe the
specimen is trustworth}^ I believe the hairs are practi¬
cally identical with those of C. riiin, as the result of such
observations as can be made on the living pupa. Nor in
other respects have I been able to seize any decided
differences. Both species, for example, appear to have
structural provision for movement between the fifth and
sixth abdominal segments. Both have the peculiarity that
I have only discovered in Theclids of having not only the
eighth but also the seventh abdominal spiracle merely
cicatricial. In neither is there any scar of the honey-
gland, present in not a few “ Blues.”
The general characters will be better gathered from the
few photographs reproduced than from long descriptions
(Plates XTX, XXXVIll, and XLIII).
P.S. — A few further notes will be found in Proceedings
O
106
Dr. T. A. O^pPJ^nnan on Gallo2')hriis avis.
under date March 16, 1910. M. Oberthur figures and
describes C. avis in Fasc. IV of his “ Etudes de
Lepidoidcrologie Com'pa.ree'G which will be published about
the same time as this. Prof. C. Mendes has a note on
the species in P>roteria 1910, p. 67.
Explanation of Plates XIV — XLIII.
[.See Explanutiun facing the Plates.]
Explanation of Plate XIV.
The figures of the imugines of C. avis and G. rnhi explain
themselves.
The figures of the heads are fairly accurate, the palpi are of course
not so accurate as those on Plate XXI, nor does the ruddiness in
C. avis give any idea of how dominant this feature is when a living
C. avis is looked in the face.
The undersides of antennae are open to the criticism that at this
magnification the individual segments are visible.
F
Trans. Ent. Soc. Lorbd.,1910,Pl.JCIV.
Avis.
Avis. Rizbi.
E.C. Knight del.
West, Newman chr.
CALLOPHRYS AVIS AND C. RUBI.
Explanation of Plate XV.
Five larvae of C. nvis on Coriaria and four of C. rubi on Cytia^ix,
last instar x 2.
Trans. Ent. Soc. Lend. ,191 0,PL. JCV.
E.C.Knight del.
West, Newman chr.
LARVAE OF CALLOPHRYS AVIS AND C.RUBI.
Explanation of Plate XVI.
Region of androconial brand, roughly sketched under camera x 16.
Fig.
5?
55
1.
2.
3.
4.
Callophrys rubi.
„ avis.
C. avis']
^ . y Androconial scales x 150.
C. rub I f
Trans. Ent. Sac. Land.., iQio., Plate XVI.
C. Hen/st'hel.
ANDROCONIAL BRAND-(i) RUBI, (2) AVIS.
ANDR0C0NIA-(3) avis, (4) RUBI.
Explanation of Plate XVII.
Larvae of G. ruhi, pliotograplied by A. E. Tonge.
The upper on growing point of gorse are in third instar magnified
X 5, the lower in last instar, natural size, on Spartium junceum.
Tnitis. E7it. Soc. Lo>td., ^910, Plate X V JI.
C. Hentsckel. Photo, A. E. Touge.
LARVAE OF C. RUBI -THIRD INSTAR ON CORSE x 5,
LAST INSTAR ON BROOM, NAT. SIZE.
r
i
'i
f '
’nl
i
'i
:S
>
1
• ^
Explanation of Plate XVIII.
Larvae of G. avis on Curiaria, last iiistar, natural size, photographed
by IMr. Main and Mr. Tonge. The figures on the right .show very
well the level back, as compared with the serrations of C. ruhi.
Trans. Ent. Soc. Lond.., iQio, Plate XVI II.
C. Hentschel. Photo, A. E. Tonge and H. Main.
LARVAE OF C. AVIS- LAST INSTAR, NAT. SIZE.
Explanation of Plate XIX.
Fig. 1. Pupa ol' C. avis, dorsal view x 4. i tt •
, w 1 . Q Photo, IT. Main.
„ 2. „ „ lateral „ x 4-8. I
„ 3. „ „ liead and portion of front X 30.
Photo, F. N. Clark.
Trans. Ent. Soc. Land.., 1910, Plate XIX.
C. Hentschel.
C. AVIS— PUPA X 4, AND HEAD x 30.
Explanation of Plate XX.
Male ancillary appendages of
(1) 0. avis X 20,
(2) 0. r'libi X 20.
I selected the preparation of G. ruhi from others a.s likely to make
the best photograph, though it is imperfect in having lost the
proximal end of the Aedoeagns.
Trafis. Ent, Soc. Land., 1910, Plate XX.
Hentschcl. Photo, F. N. Clark.
ANCILLARY APPENDAGES x 20— (i) C. AVIS, (2) C. RUBI.
Explanation of Plate XXI,
Fig. 1. Two pairs of palpi of G. rubi and one pair of C. avis x 13.
The differences are obvious. One pair of the C. rubi are more
denuded of scales than the other.
Fig. 2. Terminal abdominal segments of C. avis 9 , opened dorsally
and spread, x 18.
A portion of the same specimen is shown on Plate XXII.
Trans. Ent. Soc. Lond., igio, Plate XXI.
C. Hentschel. Photo, F. N. Cl^rk.
PALPI OF C. RUBI (2 PAIRS) AND C. AVIS.
OVIPOSITOR, ETC., OF C. AVIS.
Explanation of Plate XXII.
Portion of eighth abdominal segment 9 of
(1) G. avis X 30,
(2) G. rnbi x 30,
showing chitinous tube and spines in the wall of Bursa. The two
spines in the specimen of G. avis are so nearly .superposed that
tiiey will probably appear as one in the reproduced photograph.
These spines are so variable in G. rn&f, that probably no specific
differences in them can be relied on.
Trans. Ent. Sac. Land., iQio, Plate XXI 1.
C. Hentschel.
CANAL AND BURSA OF (i) C. AVIS 9, AND (2) C. RUBI 9 , x 30.
Explanation of Plate XXIII.
Stereoscopic view of the egg of C. avis x 60. The pictures are
placed sufficiently close together to combine ea.sily without a
stereoscope. An injury to the adventitious coat, a portion being,
as it were sliced up, is obvious ; probably occurring In journey of
egg from South France.
The lower figures ( x 60) are non-stereoscopic views of C. rubi ova.
The adherence of the hairs of the plant (Ulex) to the eggs is to be
noted. Photos by A. E. Tonge.
Trans. Tnt. Soc. Lo^id.., 1910^ Plate XXIll.
C. Hentschcl. Photo, A. E. Tonge.
STEREOSCOPIC VIEW OF EGG OF C. AVIS ^
TWO VIEWS OF EGG OF C. RUBI^^
1
m
• «
4
Explanation of Plate XXIV.
Fig. 1. Eggs of G. avis x 40.
„ 2. Egg when pressed x 180.
Trans. Ent. Soc. Land., ipio, Plate XXIV.
2
C. Hentschel.
(I) EGGS OF C. AVIS x 40.
(2) EGG PRESSED x 180.
Explanation of Plate XXV
Portions of e^oshells of
(1) C. rnhi
(2) C. avis /
not pressed x 200
Imrans. Ent. Soc. Lond., iQio, Plate XX P.
C. Hentschel.
EGGSHELLS (NOT PRESSED) x 200.
(I) C. RUBI, (2) C. AVIS.
Photo, F. N. Clark.
Explanation of Plate XXVI.
Two photographs of iiiicropyle of egg of G. avis x 385.
Coating shown broken down by Farrant medium — less so in part
of Fig. 2, where the coating has only been partially wetted by the
medium.
Trans. Ent. Soc. Lend., igio, Plate XX FI.
C. Hentschel. F. X. Clark.
MICROPYLAR AREA (2 SPECIMENS) OF EGG OF C. AVIS
IN FARRANT’S MEDIUM x 385.
Explanation of Plate XXVII.
Micropylar areas of eggs of
(1) G. nibi, (2) G. avis,
mounted in Tarrant’s medium x 350. Shows also the breaking
down of the adventitious coat of the egg in G. avis in this medium
which that of G. rubi resists.
Trans. Ent. Sac. Loud., igio, Plate XXVII.
C. Hcnlschcl. Photo, /•'. Clark.
MICROPYLAR AREAS OF EGGS OF
(I) C. RUBI, (2) C. AVIS, X 350, IN FARRANT.
SHOWS RESISTANCE TO DISINTEGRATION IN C. RUBI.
■ ,'<• r .
■(t
_ tiJ-
Explanation of Plate XXVIII.
Skin of first-stage larva of G. avis, opened ventrally and spread
out X 36.
Trans. Ent. Soc. Lana., ipio, Plate A'A^ VIII.
C. Hentschel. Photo, F. N, Clark.
SKIN OF C. AVIS LARVA, FIRST INSTAR x 36.
Explanation of Plate XXIX.
Skin of larva of G. rubi in first instar x 36.
Tra7is. Ent. Soc. Land., ipio, Plate XXIX,
C. Hentschel. Photo, F.
SKIN OF C. RUBI LARVA, FIRST INSTAR x 36.
i"". Clark.
Explanation of Plate XXX.
Skin of C. avis in second instar x 24.
Trans. Ent. Sac. Loud.., iQio, Plate XXX.
C. Henfschel. Photo, F. N. Clark.
SKIN OF C. AVIS LARVA, SECOND INSTAR x 24.
u
n
Vj'is
3
'V
'■ ' f ,_•
^ I
» . * V ’ .
;> T r'\r ’'.
^ *1*.
•>V-J
/•
<1^ -
.'M> ■...v'*
\ta
Jv;' ■
' . -<
/
at-di
Explanation of Plate XXXI.
Larva skin of C. rnhi in second instar x 24.
Trans. Ent. Soc. Land.., 1910, Plate XXXI.
C. Hentschel. Photo, F. N. Clark.
SKIN OF C. RUBI LARVA, SECOND INSTAR x 24.
Explanation of Plate XXXIL
Skin of C. avis in third instar x 24.
/
Trans. Ent. Soc. Lond., iQio, Plate XXXII.
C. Heutschel. Photo, F. N. Clark.
SKIN OF C. AVIS LARVA, THIRD INSTAR x 24.
r:
t
-• ; '
; *• ^ ?♦ ■ '
v'' i
'y :.
*3>
/
•? *
*
;:t'-
r:<^
j^.
- V .-■'«
‘ ^
i
Explanation of Plate XXXIII.
Skill of C. ruhi in third instar x 24.
Trans. Ent. Soc. Land., iQio, Plate XXXI II.
C. Henfschel.
Photo. F. X. Clark.
SKIN OF C. RUBI LARVA, THIRD INSTAR x 24.
Explanation of Plate XXXIV.
Prothoracic plate of larva of
(1) C. rubi, (2) G. avis, in second instar x 150.
The latter has posterior margin folded under, as has also Plate
XXXV, fig. 1.
Trafis. Ent. Soc. Loud., igio, Plate XXX IV
C. Ilenischel. Clark.
PROTHORACIC PLATES OF (i) C. RUBI, (2) C. AVIS,
SECOND INSTAR x 150.
Explanation of Plate XXXV.
Prothoracic plate of larva of
(1) C. r'lihi, (2) G. avis, in third instar x 150.
The special hairs at outer angles are well shown on both this and
previous plate.
Trans. Ent. Soc. Loud.., K^io, Plate XXXV.
C. Hcntschel. F. N. Clark.
PROTHORACIC PLATES OF (i) C. RUBI, (2) C. AVIS, IN
THIRD INSTAR x 150,
Explanation of Plate XXXVI.
Fig. 1. Prothoracic plate of C. avis in last instar x 150, or rather
of the central chitinised portion of it, the ends which are here like
the rest of the skin in texture, witli the special angular hairs, are
outside the figure.
Fig. 2. Honey-gland area of C. avis in third instar x 100.
Tra?is. Ent. Soc. Lend., ip/o, Elate XXXVI.
C. Hcntschel. Photo. F. Clark.
C. AVIS.
(I) PORTION OF PROTHORACIC PLATE, LAST INSTAR x 150,
(2) HONEY-GLAND AREA, THIRD INSTAR x 100.
r
Explanation of Plate XXXVll.
Honey-gland area of
(1) C. rubi, (2) C. avis, in last instar x 100.
Trans. Ent. Soc. Lond., igio, Plate XXXVII.
'C. Hentschel. Photo, F. X. Clark.
HONEY-GLAND AREA IN LAST II N STAR x lOO.
(I) C. RUBI, (2) C. AVIS.
\
1
• r
/
wmw.
f
i
j.
Explanation of Plate XXXVIII.
Portions of cremastral region of pupae of
(1) G. avis X 200,
(2) G. rubi x 200.
These show the greater length and slenderness of the hooks in
C. avis, despite the maceration that has injured the hairs, which are
much alike in both species. A fine specimen is seen in the G. rubi
photograph.
Trans. Ent. Soc. Land.., iQio., delate X\X VII 1.
C. Hentschel. P-
CREMASTRAL REGIONS (PORTION) OF (i) C. AVIS,
(2) C. RUBI.
Explanation of Plate XXXIX.
Pupa of C. ruhi.
Portions of pupa, chiefly to show hairs, which are almost identical
with those of C. avis, in my preparations of which they happen to
be destroyed or absent, enabling, however, the other features to be
more easily seen.
Fig. 1. End of prothorax x 80.
Fig. 2. Portion (dorsal) of 5-6 abdominal incision x 45, to show
functional intersegmental membrane. It is nearly identical in C.
avis, but was not so well shown in specimen.
Trans. Tnf. Sac. To?id., iQjIO, Plate XXXIX.
C. Hcntschel. Photo, F. N. Clark.
PORTIONS OF PUPA CASE OF C. RUBI.
Explanation of Plate XL.
Pupa of G. avis.
Fig. 1. Prothorax of pupa x 18, a portion of mesothorax with
spiracle covers also shown.
Fig. 2. Half of prothorax x 36.
N.B. — Hairs destroyed.
Trans. Ent. Soc. Land., ipio, Plate XL.
C. Hentschel. Photo, F. N. Clark.
PORTIONS OF PUPA CASE OF C. AVIS.
Explanation of Plate XLT.
Pupa 6f G. avis.
Fig. 1. Portion of mesothorax x 30. Spiracle cover in front,
dorsum on the right, forewing on the left, of picture. The change
of skin sculpture from the body to the wing is very distinct, the
former having hairs, lenticles and rosettes, the latter only network.
It is noteworthy that the .spiracle cover belongs to the wing area.
Fig. 2. A portion of wing x 50 to show sculpture.
Trans. Enf. Soc. Loud., igio, Plate XLI.
C. Hentschel. Photo, F. X. Clark.
PORTIONS OF PUPA CASE OF C. AVIS.
Explanation of Plate XLII.
Pupa of C. avis.
Fig. 1. Portion of antennae, legs and maxillae x 50. Chiefly to
show lenticles at tibio-tarsal (?) joints ; the only jdace where the
appendages (wings, legs, antennae, etc.) are not absolutely without
any hairs, lenticles, rosettes, or any skin process but network.
Fig. 2. Metathorax of one side x 35.
Trans. Ent. Soc. Land.., iQ^o, Plate XL II.
C. Hentschel.
Photo, F. N. Clark.
PORTIONS OF PUPA CASE OF C. AVIS.
1
■* ?
f
i
\
-■3
'4
3
r
r
\
I
Explanation of Plate XLIII.
Pupa of C. avis.
Fig. 1. Portions of sixth, seventh and eighth abdominal segments
X 50, to show spiracle on sixth and only scars on seventh and
eighth. The obsolete seventh abdominal spiracle being characteristic
of Theclid pupae.
Fig. 2. A spiracle and its surroundings of third abdominal segment
X 100, showing lenticles and rosettes and hairs, the latter as already
explained damaged in preparation.
Trans. Ent. Soc. Lond., ipio, Plate XLIII.
C. Hentschel. Pholo, /•'. .V. Clark.
PORTIONS OF PUPA OF C. AVIS.
I
( 107 )
III. 071 the, genus Deuterocopiis, Zdlc7\ By T. Bain-
BRiGGE Fletcher, R.N., F.E.S.
[Read Decembei’ 1st, 1909.]
Plates XLIV, XLV.
The present paper deals with those species of the
Ftco’ophoo'idae in which the forewing is cleft into three
lobes. Hitherto all these forms have been included in
the single genus Deute7'oc(ypus, 2i., but, owing on the one
hand to the very distinctive appearance of members of
this group, and on the other hand to the insufficiency of
former descriptions based on very scanty material, great
confusion has resulted, three or four distinct species often
being referred to under one common name (and that often
wrongly applied to any one of them). Tliis paper is an
attempt at a revision, not only of the species of these
little Plume-moths so far as ive know them at present,
but also of all the previously published literature concerning
this group.
I may say at once that such a task would have been
quite impossible without the kind help rendered by Lurd
Walsingham, who most generously put at my disposal the
whole of his unrivalled series for examination and de¬
scription, and I take this opportunity of acknowledging my
indebtedness.
So far as we know them at present, the Plume-moths
with trifid forewings are included in three genera, two of
which are here described as new, and consist of only 13
species. Their head-quarters appear to be in the Indo-
Malayan region, but further research will probably show
that these little moths are widely distributed throughout
the Tropics.
The three genera may be artificially tabulated as
follows : —
Fo7'ewing with tlwee scginents.
2. f
\
General appearance Trichoptiline, first cleft from about
abdomen long and slender .... Hexadactilia
General appearance Oxyptiline, first cleft from beyond i,
abdomen short and stout. ....... 2
Second cleft at least {y length of first . . Deuterocopns
Second cleft not more than J length of first Leptodeiiterocopxs
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE)
108 Mr. T. B. FlCTclier on the genns Dcntcrocojpns.
Hexadactilia, n. g.
Lilbial pal})! longer than length of heiul. Alalonien long and
slender. Posterior tibiae with three tufts of s[)inoi;s hairs. Wings
Fio. 1. — Hexadactilia trilobata.
long and slender. Forewing cleft from about 1, first segment very
narrow, second segment again cleft from within ^ of its length, sub¬
segments linear. II indwing cleft from about and from near base,
segments very slender and linear, third segment with a large apical
scale-tooth.
Type : — 11. trilobata.
Hexadactilia trilobata, ii. s.
(Plate X LI V, fig. 1.)
Expanse 15 mm. Labial palpi long, semi-erect ; second joint
slightly dilated ajjically, light fuscous annnlated with darker ; third
joint rather over half-length of second, smoothly scaled, dark-fu.s-
coiis. Antennae dark-fuscous, dotted with whitish above. Head
and thorax tawny-fuscous. Abdomen long and slender, tawny-
fuscous, sufl'used with dark-fuscous ; fourth segment with jialer /\
markings, posterior segments margined with whitish. Foreleg dark-
fuscous : tibia short, dilated liy a tuft of blackish scales banded
and tipped with white ; tarsus narrowly banded with white at joints.
!Midleg long, slender, tawny-fuscous : tibia with scale-tufts at | and
apex, the latter tuft emitting a pair of long spurs. Hindleg long
and slender, tawny-fuscous, irrorated above with dark-fuscous :
tibia with pale scale-tufts at L and apex, second and third tufts
each emitting a pair of long slender blackish-tipped spurs ; tarsal
joints slightly thickened with scales apicall3^
Forewing long, slender, cleft from about I, second segment again
cleft from about J of its length ; first segment very narrow, cleft
wide and conspicuotts ; subsegments linear, first subsegment parallel
Mr. T. B. Fletcher on the genus Deuterocopus. 109
to first segment, divergent from second su’osegment ; pale tawny-
fuscous, suffused with fuscous ; base of cleft preceded and edged by
a longitudinal blackish discal spot ; first segment with central third
pale-tawny, cut at | by a narrow transverse white line, area beyond
this pale-tawny ; second segment pale-tawny at base of second cleft ;
subsegments cut at of their length by a narrow white line, area
beyond this paler. Cilia on costa fuscous-brown, pale-ochreous
opposite paler areas, a whitish subapical dot ; on hiirder-margin of
first segment whitish-ochreous mixed with a few whitish liairs and
scattered black scales, on third fourth blackish, on outer fourth
whitish-ochreous ; on fore-margin of second segment pale-ochreous,
blackish on central third of segment ; orr hinder-margin of first sub¬
segment whitish-ochreous with a few scattered black scales ; on
fore-margin of second subsegirreut whitish-ochreous orr first forrrth of
segment, beyond J pale-firscous with a few scattered black scales ,
on dorsrrrn ochreous-whitish berreath bases of both clefts, otherwise
blackish, with minute black scale-tufts at f and apex.
Iliudwing cleft from within J arnl from near base, segments very
slender and lirrear j dark-fuscous. Cilia dark-fuscorrs, third seg
nrerrt orr both margins with a large black apical scale-tooth and a
few scattered black scales between this and base.
Type ^ (No. 41944') in Walsingham Coll., and Co-t3^pe
(No. 8527) in Bainbrigge Fletcher Coll. ; both from Ron,
New Guinea, in July 1897 {Doherty). A third example
in Lord Walsingham’s collection was collected by Doherty
at Humboldt Bay, Northern New Guinea, in April 1893.
Deuterocopus, Z.
Deuterocopus, Zeller, Linn. Ent., vi, 402 (1852) ; Bull. Soc.
Nat. Mosc., XX vi, pt. ii, PI. 4 ff. 17-20 ; Walker,
Cat., XXX, 951 (1864) ; Meyrick, T.E.S., 1886, 8 ; Rebel,
Denk. Matli-Nat. K.K. Ak. Wiss., Ixxi (ii), 85-87
(1907) ; Fletcher, Spolia Zeylan., vi, 16-19, PI. B
f. C. (1909).
Deuteroscopus, Hofmann, Iris, xi, 329 (1898) ; Pagen-
stecher, Abh. Ges. Zook, xxix, 241.
The following is a translation of Zeller s original
diagnosis of the genus : —
“ Deuterocopus, nov. geii. (From bevrepos, second, and
KOTiro), I cut.)
Forewings semi-trifid (== bifid, the second segment also bifid).
Hindwings trifid, the third segment very .short. Abdomen stout.
Anteiiiiae rather sliort, lilifonu. liej^s short, tibiae willi knots of
scales, posterior tibiae with long spurs.
“ A very distinct genus, at first sight in appearance a
Pterophorus of Section B. {Oxyptilns, Z.] But distin-
guished from that genus by the fact that the. lower segment
of the forevnng is cleft from half its length into two linear
divergent subsegments and that the third segment of the
hindxoing is only half the length of the second segment.
Besides these points the abdomen is very thickset and finely
pointed posteriorly ; the antennae are unusually short, the
legs short and stout and thickened with knots of very
enlarged long scales at the roots of the strong, long spurs,
as well as at the ends of all the tibiae.”
Fig. 2. — Deiiterocopus tengstroeini.
In Trans. Ent. Soc. Bond., 1880, p. 8, Mr. Meyrick
diagnosed the genus Deiiterocoims as follows
“ Forehead without tuft. Auteniiae in male — (?). Palpi moder¬
ate, ascending, 2nd joint with appressed scales, terminal joint
moderate, acute. Tibiae thickened on origin of spurs with dense
spreading whorls of scales ; posterior tarsi with similar smaller
whorls on apex of two basal joints. Forewings trifid, cleft centrally
from middle and lower segment cleft from two-thirds ; 2 and 4
from point of angle of cell, 3 absent, 5 and 6 very short, 7 from
below angle long, 8 and 9 long-stalked, 10 from near angle, 11 from
near 10. Hindwings trifid, 3rd segment unusually short, terminating
in a tuft of black scales ; 2 from middle of cell, 3 absent, 5 and 6
very short, 7 to apex.
“ I have examined D. tengstroeini, the only known
species. The genus is derived from Plcdyptilia, and other¬
wise most nearly allied to Sphenarches.” It may be added
that the species examined by Meyrick was evidently not
tengstroeini, and that he was incorrect in giving as a
generic character the absence of vein 3 of the forewing.
Mr. T. B. Fletcher on the genus Deuteroco'pus. Ill
In his monograph of the Palaearctic Orneodidae, Dr. O.
Hofmann remarked : —
“Tlie genus Deuteroscopus from Java, with three lobes to the
fore wing, which Zeller (Lin. Ent., vi, p. 402) on its foundation
alleged to have a relationship with the Orneodidae, is according to
investigation of its neuration a typical Pterophorid standing next to
the genus Oxyptilus, Z. The cleft of the lower segment of the
forewing is only a continuation, stretching deeper into the wing-
membrane, of the forking which in Oxyptilus is found between the
extended branches IIP of the Medius [vein 4] and IV^ of the
Cubitus [vein 3]. Also the club-scales on the branches of the
Cubitus of the hindwing, characteristic of the Pterophoridae, are in
Deuteroscopns also present and indeed in two rows on the base of
both branches, only in one row beyond that in the periphery itself.”
In these observations Dr. Hofmann was perfectly correct
— except for his misquotation of the name of the genus —
and I quite agree tliat Dcuterocopus may be regarded as
an offshoot from the Oxyptiline stem.
In “ Spolia Zeylanica," vol. vi, pp. 16-17, I made some
remarks on the Sinhalese members of the genus, but at
that time I had concluded that the different forms occurring
in Ceylon all belonged to one highly variable species.
This is an opinion which I have now seen reason to alter,
and my previous remarks under this heading must be
considered as cancelled.
The following characters are now given as a revised
definition of the genus : —
Labial palpi upturned, third joint about 4 length of second. Haustel-
lum developed. Antennae moderate, minutely ciliated in Head
with erected scales on crown. Abdomen short, thickset, third seg¬
ment usually edged posteriorly with a raised band of scales. Fore¬
tibia with apical scale-tuft, mid-tibia with a scale-tuft and pair of
spurs at apex, hind-tibia very stout with a moderate whorl of scales
near base and two large whorls, first beyond half, second apical, each
emitting a pair of very long spurs, posterior tarsi with small whorls
of spinous hairs on apex of two basal joints. Forewing moderately
broad, cleft from about second segment again cleft, subsegments at
least half length of second segment. Hindwing cleft from about f
and from near base, segments linear, third segment with large apical
scale-tooth and smaller tooth on dorsal margin at about half.
Neuration ; — Forewing : 2 from lower edge of cell, 3 and 4 from
beneath base of first cleft, 5 and 6 very short or absent, 7 from above
112 Mr. T. B. Fiff •her on the genns B enter oco’pxts,
main cleft to liimler-niargin of first segment near apex, 8 and 9 long-
stalked, 10 from near angle of cell, 11 from near 10, 12 into costa
before cleft. Hind wing : 1*’ to apex of third segment, 2 from middle
of cell, short, 3 absent, 4 to apex of second segment, 5 and 6 very
short or absent, 7 to apex, 8 into costa at about half. Spina of
frenulnm single in 5 .
Synopsis of species of Deuteiiocopus.
1. -!
2.
3.
4
6.
f Blackish species, f.w. and abdomen witbout any ferruginous-
I orange markings ........ 2
I Not blackish species, f.w. and abdomen with ferrnginous-
[ orange markings ......... 3
JCo.sta of f.w. with scale-tuft at about f . . . melanota
\ Costa of f.w. without scale-tuft at about f . . famulus
f Pectus and venter whitish . . . 4
\ Pectus and venter jiale-yellowish ...... 9
rSecond cleft only about ^ length of first . . . . .o
\ Second cleft at least f length of first . . . . .0
j'Dorsum of f.w. with a small l)lack scale-tuft beneath base of
I first cleft . . atrapex
1 Dorsum of f.w. without scale-tuft beneath base of first
1^ cleft . . . . . . . . . tengstroemi
f Costa of f.w. with tuft of scales at about f . . lophojdergx
\Costa of f.w. without tuft of scales at about f ... 7
[Dorsum of f.w. with dark scale-tuft beneath base of first cleft 8
J Dorsum of f.w. without dark scale-tuft beneath ba.se of first
cleft ........ (dhipunctatus
(Expanse 10 mm. ; posterior tarsi ringed with whitish or
paler . socotran us
Expanse 12 mm. ; posterior tarsi unicolorous golden-
ferruginous ....... bathychasma
Second cleft f length of first ; no scale-tuft in dorsal cilia of
f.w. beneath base of first cleft ..... planeta
Second cleft less than f (usually about |) length of first ; a
dark scale-tuft in dorsal cilia of f.w. beneath base of first
cleft .... ritsemae (including rubrodacti/his)
Dcutex'oeopus famulus, Meyr.
(Plate XLIV, fig. 2.)
Deuteroco'pus famxdus, Meyrick, T.E.S., 1907, 474.
“ 9 . 15 mm. Head, palpi and antennae [evidently a lapsus calami
for ‘thorax’] dark coppei’y-fuscous w'ith a few _ white scales.
Mr. T. B. Fletcher on the genus Deuterocofus. 1 Bj
Antennae dark-fuscous. Abdomen blackish-fuscous, first .segment
white except a dorsal stripe, second segment wholly ochreous- white,
third ochreous-white on sides, ventral surface wholly ochreous-white.
Legs dark coppery-fuscous, anterior and middle pairs lined and
sprinkled with -white, posterior tibiae banded with white, -with
expansible wdiorls of scales on origin of spurs (tarsi broken). Fore¬
wings cleft firstly from before second segment cleft from its middle ;
dark prismatic coppery-fuscous a few' irregularly scattered white
scales ; cilia fuscous, towards apex of segments suffused with dark-
fuscous, beneath apex of first and second segments with a slender
white bar, on dorsum with a wdiite space extending from middle to
I, including a central fuscous bar. Hindwings cleft firstly from 4,
secondly from h, segments slender ; dark-fuscous ; a pellucid streak
in disc ; two or three wdiite scales in middle of third segment • cilia
pale pri.smatic purplish-fuscous, darker towards tips of segments,
apex of third segment with a small projection of black scales above
and beneath.
“ Kei Islands, in May ; one specimen.”
I have examined three examples of this species in the
collection of Lord Walsinghara. They are labelled : —
Fergusson Id., D’Entrecasteaux Is., 8-30, ix, 1894, Meek,
No. 17328; Patani, Halmahera, Doherty 1897, No,
419C0; and Batian, viii, 1897, Doherty, No. 41948.
Dcuterocoims mclanota, n. s,
(Plate XLIV, fig. 3.)
Deutej'oeopus mclanota, Wlsm., MS. (ined.)
(J. Expanse 11 mm. Palpi erect, smoothly scaled, third joint
filiform, acuminate, shorter than second ; greyish-wdiite, banded
with brown at joints and at apex. Antennae brown, dotted with
wdiitish above. Head dark-browm, sprinkled with wdiitish ; some
anteriorly-erected scales on top of crown. Thorax blacddsh-brown
sprinkled wdth greyish-ochreous ; joectus whitish. Abdomen blackish-
brown, sjDrinkled with greyish-ochreous especially towards base ;
second segment with two short submedian posteriorly-divergent
wdiitish streaks, apex of segment edged with a few scattered wdiitish
scales ; apex of third segment edged wdth a raised band of ochreous-
whitish scales intermixed with blackish-brow'n ; fifth segment wdth
a pair of whitish submedian dots separated by a darker space ; anal
tuft long, blackish-brown : venter wdiitish, fourth segment crossed
TRANS. ENT, SOC. LOND. 1910. — PART II. (JUNE) I
114 Mr. T. F!(i?l ler on the fjenus Devteroco'pns.
l)y a l)lackisli-bro\vn bar. Foreleg fnscous-brown : tibia short and
tliickened with scales ; tarsus whitish, banded with blackish-brown
in middle and at apex. (Mid legs broken.) Hindleg blackish-brown ;
tibia with slight whorl of scales at ^ and moderate whorls at f and
apex, broadly lined with whitish at base and below whorls ; tibial
spurs strongly -developed, dai’k -brown, wdiitish in centre and at apex,
posterior edges roughly haired, spurs of proximal pair longer than
those of distal pair ; tarsus blackish-brown, apices of joints with
gradually decreasing clusters of spinous hairs, first tarsal jointbroadly
lined with white above, third tarsal joint suffused with greyish.
Forewing cleft from within |, segments narrow, second segment
again cleft from about J of its length, subsegments linear : blackish-
coppery-brown, irrorated with whitish-ochreous : a narrow whitish
subdorsal dash at J, a broad whitish subcostal spot at about reach¬
ing nearly to dorsum but separated from base of cleft by a narrow
black interval ; first segment cut at about J and f by ill-defined
whitish bars of which the inner is the more distinct ; first and second
subsegments with a few whitish scales a little beyond Cilia on
costa blackish-brown with a conspicuous projecting tuft of black
hair-scales above base of first cleft and slight tufts of black hair-scales
at about * ; at apex black ; on hinder-margin of first segment
brownish, black from about f length of segment to just before apex,
beneath apex greyish-ochreous, base of cilia black and forming a
small posteriorly-projecting tooth ; on fore-margin of second segment
blackish-brown, beyond 4 black ; on apex of first subsegment greyish-
ochreous ; on hinder-margin of first subsegment greyish-ochreous,
blackish beyond | to just before apex, beneath apex a small posteriorly-
projecting scale-tooth ; on fore-margin of second subsegment blackish-
brown to 4, beyond that blackish ; on apex of second subsegment
greyish-ochreous ; on dorsum greyish-ochreous, with a small black
scale-tooth at about f (beneath base of first cleft), beyond f
blackish-brown, beneath tornus a small black posteriorly-projecting
scale- tuft.
Hindwing cleft from -I and from near base, segments linear ;
blackish-coppery. Cilia blackish-brown, on hinder-margin of second
segment greyish-ochreous at about 4, third segment with a large
spatulate tooth of black scales occupying exterior fourth of both
margins and a small black scale- tooth on dorsum slightly within
Type ^ (No. 42101) in Walsingham Coll.; Perak (3000
feet), October 1890 {Doherty).
Mr. T. B. Fletcher on the genus Deutcrocoims. 115
Deutcrocojms tcngstroemi, Zell.
(Plate XLIV, fig. 4.)
Deuierocopus tengstroemi, Zeller, Linn. Ent., vi, 402-404
(1852) ; Zeller, Bull. Soc. Imp. Nat. Mosc., xxvi, pt. ii,
t. iv. It. 17-20, p. 516 ; Walker, Cat., xxx, 951 (1864).
nee
Meyrick, T.E.S., 1886, 8 ; Pagenstecher, Abb. Ges. ZooL,
xxix, 241 Q.atrapex)] Meyr., J. Bombay N. H. Soc.,
xvii, 134 { = socotranus, Ptbl.) ; Meyr., T.E.S., 1907,
474 { — ato'apex, Fletcher); Fletcher, S^^ol. Zeylan., vi,
16-17,18,19-20 (1909) {— soc otr anus, Rbl.) ; Max-
well-Lefroy, Indian Insect Life, jd. 528 (1909).
The following is a translation of Zeller’s original descrip¬
tion : —
“ Body and wings ferruginous, cilia of the latter spotted with fus¬
cous ; third segment of hindwing with black scaling in middle of
dorsum and around apex (d ? )•
“ Size of a small Pt. ohscnrus. Body and wings rust-yellow, the
former rather shining and somewhat browned on the thorax. An¬
tenna in 5 half-length of forewing, moderately thick, ringed
with white and brown, basal ly white above ; antenna of ^ somewhat
longer, less ringed, above dilated and with broader white markings ;
terminal joint somewhat thickened, apex acuminate. The hairs on
the hinder margin of the head erected and inclined forwards. Palpi
as long as the head, moderately thin, horizontal ; the terminal joint
pointeil, white above, before the tip and on the underside yellow-
brown. Haustellum yellowish, as long as the thorax. Pectus
white-spotted on the side. Venter white, the edges of the first tAvo
segments with small rust-yellow bands enlarged laterally. Forelegs
rather shining on the femora and base of tibia ; the end of the tibia
is thickened knottily with unequally-long, very plentiful, rust-brown
scales; out of the knots project the long wliitish spurs, blackish on
one side ; the tarsi are whitish above, the first joint thickened ajAically
with scales and black, the three folloAving not densely scaled beneath,
the two last suffused with black. In the second pair of legs the tip
of the tibia is only slightly thickened, the tarsi whitish blackened at
the tip. [N.B. — By some error Zeller has evidently transposed the
descriptions of the fore- and mid-legs, and this must be borne in
mind when reading the above ti’anslation. — T.B.Pb] In the posterior
legs the tibiae are extraordinarily thick, with three thickenings; the
I 2
16 Mr. T. B, Flff^ber on the fjeniLS T) enter oco'puH.
weakest is near tlie base ; tlie second is strong, milled up in the
shape of a funnel, sprinkled with brown at its l)ase, and is situated
at tlie root of the first pair of spurs ; the third is still stronger and
longer, is likewise ruflled up and thickly suffused with blackish and
is situated at the extremity of the tibia : the upper pair of spurs
reaches somewhat beyoncVthe extremity of the tibia and is thickened
with hairs, rusty-yellow on one side, on the other white and blackish-
brown at the tip ; the second pair of spurs is as long as the first tw’o
tarsal joints and is also hairy on one side, rusty-yellow', black at the
root and in the middle, and at the tip wdiite with a small black spot.
The first three tarsal joints are rusty-3'ello\v, the first wdth a some¬
what weak apical hair-funnel, tlie second with a weaker hair-funnel
white apically, the third wdth a still weaker browm hair-fnnnel ; the
last tw'o joints are black.
“Forewdngs 2| to 2^ lines long, moderately slender ; the fissure
stands somewhat separated. The first segment, which is almost
half as long as the wdng, is brown and strewn longitudinally wdth
blackish scales. A bright yellowdsh transverse streak cuts off the
beginning of the darkening to a narrow' bar. The costal cilia are
bright rust-yellow wdth three black longitudinal streaks at. equal
distances up to the apex. The yellowdsh inner-marginal cilia are
blackish at the mouth of the fissure ; betw'een these places and tlie
black streak coming out to the tip of the w’ing the fringes are light-
yellow on the blackish edge-line. The second segment is browned
on its outer three-fourths, darker than the first segment ; at its
middle it splits into two divergent, linear subsegments, of wdiich the
anterior is the longer ; the cilia of both subsegments are of the ex-
tremest pale-yellow at the tip ; moreover, the first subsegment is
black-fringed at the extremity of its fore-margin and also on the
inner half of its lower-margin ; and all the cilia of the second subseg¬
ment are also so coloured ; but both these subsegments have black
scales at their apices at the base of the pale cilia.
“ The first tw'o segments of the hindwing are cleft up to three-
fourths of the wdng-length ; they are faintly browned posteriorly
and the lower margin of the second segment has thei)alest yellowdsh
cilia. The third segment is so extraordinarily short that it only
reaches a little beyond the cleft between the first tw'o segments ; it
is very slender, and has in the middle of the dorsal margin a small
deep-black scale-tufr,, and at its apex is edged w'ith a large rounded
black scale-spot enclosing both margins.
“ Underside coloured almost like the upperside.
‘^Habitat. — Java. I have named the species after its discoverer,
Herr J. af Tengstrbm, the airthor of the ‘ Bidrag till Finlands Fjiiril-
Fauna.’ ”
]\Ir. T. B. Fletcher on the gmns D enter ocojjns. 117
In the following year Zeller published figures of this
species, but without any text beyond description, in Bull.
Soc. Nat. Moscou. Figure 17 shows the whole insect,
but it is to be noted that the second segment is here
shown as cleft from about whereas figure 20 (which
gives a more enlarged view of two wings) shows this
cleft from h only in accordance with the description.
Figures IS and 19 show the raid- and hind-leg.
The following is a redescription of one of Zeller’s original
specimens of D. tengstroemi, now preserved in Lord Wal-
singham’s collection. It is labelled “ Java, Tengstr.” and
also “ Type 2/2 ” : —
5. Expanse 12 mm. Palpi long, slender, semi-erect, pale-ferru¬
ginous ; second joint very slightly dilated at apex, where it is banded
with dark-ferruginous ; third joint slightly over half length of
second, aexuninate, broadly banded with dark-ferruginous at base
and apex. Antennae dark-ferruginous, dotted with whitish above.
Head light-ferruginous, top of crown with long anteriorly-erected
scales. Thorax light-ferruginous ; pectus marked with whitish.
Abdomen stout, light-ferruginous with traces of whitish submedian
spots on second segment, and of a narrow transverse white band
along posterior edge of third segment ; ventral surface white, mottled
on sides with ferruginous-golden, fourth segment margined posteriorly
with ferruginous-golden. Foreleg whitish-ferruginous, tibia with
apical scale-tuft of darker ferruginous scales (but no “ white spines’’).
Midleg whitish-ferruginous, tibia with apical scale-tuft of dai’ker-
ferruginous scales (spurs broken). [Hindlegs missing.]
Forewing cleft from rather within f, first segment moderately
broad with slight hinder-angle, second segment expanded posteriorly
and again cleft to one-half of its own length, subsegments linear,
first parallel to first segment, divergent from second : bright ferru¬
ginous-golden slightly suffused with light ferruginous : first segment
lightly suffused with ferruginous except on costa at i length of seg¬
ment, cut at about | by a nearly obsolete inwardly-oblique whitish
line ; second segment lightly suffused with ferruginous, first subseg¬
ment with a minute whitish dot near base. Cilia on costa light-
ferruginous, slightly darker on base of first segment, on first segment
black at f as a slight bar, with slight blackish admixture at | and | ;
at apex black ; on hinder-margin of first segment whitish-ochreous
from I to immediately before hinder-angle blackish-ferruginous, a
hinder-angle a small blackish wisp preceded by a very narrow
ferruginous wisp, on termen pale-ferruginous with a blackish basal
line ; on fore-margin of second segment whitish-ochreous, beyond |
118 Mr. T. B. Flmcher on the genus De^derocoims.
blackish-ferruginous, at apex of first subsegment whitish-oebreous
with a broad blackish basal line surrounding apex of subsegment and
forming a very small posteriorly-directed scale-tooth ; on hinder-
margin of first subsegment dark-ferruginons to i, beyond | whitisli-
ochreous ; on fore-margin of second subsegment dark-ferruginous to
i, beyond I blackish ; at apex of second subsegment whitisli-ochreous
with a black basal line produced into a small posteriorly-directed
tornal scale-tooth ; on dorsum whitish-ochreous, a small blackish
wisj) beneath base of second cleft, cilia beyond this darker.
[N.B. — The cilia on the forewing are in very good order, and tliere
is no sign of there ever having been a scale-tooth beneath the base of
the fird cleft.]
Ilindwing cleft from about | and from near base, segments linear ;
liright ferruginous-golden slightly suffused with light-ferruginous.
Cilia whitish-ochreous ; third segment with a large apical blackish-
ferruginous scale-tooth on both margins, a few small scattered black
scales on fore-margin, and a small blackish tooth on dorsum slightly
before i.
llahitat. — Zeller’s original specimens, one of which I
have examined, were from Java. The only other examples
which I have seen are two Irom Queensland in Lord
Walsingham’s collection, one from Cedar Bay {Meeh, 1894),
the other — very worn and possibly not really tengstroemi,
its condition making its identification doubtful — from
Geraldton, near Cairns (J/cc/r, 1894).
Deuteroco'pus atrapex, Fletcher.
(Plate XLIV, fig. 5.)
Dcuterocopus atrapex, Fletcher, Spolia Zeylanica, vol. vi, p.
19 (1909).
Dcuterocopus tengstroemi, Meyrick, T.E.S., 1907, 474 (piec
Zeller).
? Deuteroscopus tengstroemi, Pagenstecher, Abh. Ges. ZooL,
xxix, 241 {nee Zeller).
In my original brief description I had confused this
species with D. lopihoptcryx and therefore I now give the
following redescription of the type of D. atrapex : —
Exp. 13 mm. Palpi erect, smootlily-scaled, sickle-shaped,
reaching vertex ; third joint about one-half length of second, acumi¬
nate : whitish-ochreous, banded with ochreous-brown at joints.
Head dark-brown sprinkled with dark-ochreous ; a few anteriorly-
erected scales on crown. Haustellum well developed. Antennae
Mr. T. B. Fletcher on the gmns Deiiterocopus. 119
ferruginous-brown with a series of whitish dots above. Thorax
golden-ferruginous, wholly suffused with dark-brown ; pectus
whitish. Abdomen rather stout, golden-ferruginous lightly suffused
with dark-brown, across third segment a broad transverse dark-brown
bar ill-deffned posteriorly and tending to be continued as a short
subdorsal stripe on either side ; anal tuft short, concolorous with
abdomen ; venter whitish with a ferruginous transverse bar across
fourth segment. Foreleg : femur ferruginous ; tibia short, ferrugin¬
ous, banded with whitish at base and provided apically with a large
flattened brush-like tuft of dark -ferruginous scales ; tarsi-whitish,
dark-ferruginous at apex of first joint and beyond ^ of third joint.
Midleg ferruginous : tibia with a large apical tuft of dark-ferruginous
scales below which projects a pair of moderate spurs ; tarsi lined
with whitish above, first tarsal joint incrassated apically with short
divergent posteriorly-projecting spinous hairs. Hindleg bright
golden-ferruginous minutely irrorated with blackish : tibia large and
stout with whorls of scales at i, f and apex, the second and third
whorls subequal, much larger than first, and each emitting a pair of
long whitish slightly curved spurs which are banded with blackish -
ferruginous at their bases and before their apices and which are
roughly haired all along their posterior edges ; the inner proximal
spur is slightly longer than the outer, those of the distal pair are
equal and are each rather shorter than the outer proximal spur ;
tarsal joints with very small apical whorls of spinous hairs, these
whorls decreasing in size on successive tarsal joints, second joint
moderately, third and fourth heavily, suffused with blackish.
Fore wing rather short, cleft from second segment again cleft
from about i of its length, segments sublinear without any appreci¬
able angles : bright golden-ochreous, suffused with ferruginous
especially evident between i and f : extreme base of wing blackish,
a whitish subcostal patch at f ; first segment wholly black, cut at
^ of its length by a narrow transverse ferruginous fascia most
evident costad ; second segment sharply cut at base by a line extend¬
ing obliquely outwards from base of cleft to dorsum, the whole area
outside of this line (and including both subsegments) wholly black.
Cilia golden-ochreous on costa, ferruginous opposite ferruginous
suffusion and with traces of three pale-ferruginous patches (sometimes
well marked) between f and apex ; at apex black ; on hinder-margin
of first segment blackish with a few heavy black scales towards base,
at 5 a moderate patch of pale-ochreous and a narrow patch of pale-
ochreous just before termen, where the cilia are black at their base
and pale-ochreous apically ; on fore-margin of second segment
blackish ; on termen of first subsegment p ile-ochreous, blackish at
their base around apex of subsegment ; within second cleft blackish ;
120 Mr. T. B. Filler on the genus Deuteroco'pus.
at tornus pale-oclireous ; blackish at their base ; on dorsum ochreous-
yellow with a small blackish scale-tuft at f (beneath base of first
cleft), blackish beyond I.
liindwing cleft from about -f and 1, segment linear: golden-
ferruginous irrorated with blackish. Cilia pale-ferruginous, darker
around apices of segments and at ^ on hinder-margin of second seg¬
ment, third segment with a large terminal patch of blackish scales
occupying exterior fourth of both margins, a small black scale-tooth
on dorsum at i, and a few scattered blacki.sh scales on anterior
margin.
Type No. 6650, in Bainbrigge Fletclier Coll.: Ceylon,
Central Province, Kandy (1600 feet), May 1907 {Tl. Ernest
Green).
I have examined examj)les from the following localities : —
Ceylon — C. Prov., Kandy, May 1907 {E. E. Green)
[Fletcher Coll., No. 6650, Type atrapex] ; S. Prov.,
Galle, June 19, 1908 {Fletcher), one.
Assam — Khasi Hills, Dec. 1906 [Meyrick Coll., one].
Selangor — Batu Estate, July 27, 1902 {H. E. Durham)
[Brit. Mus., one].
Tenasserim — Mergui, 1888 {Doherty) [Wlsm. Coll, one].
S. E. Borneo — Pulo Laut, 1891 {Doherty) [Wlsm. Coll,
one].
Tern ATE — 1892 {Doherty) [Wlsm. Coll, one].
Amboyna — 1892 {Doherty) [Wlsm. Coll, seven].
Batian — August 1897 {Doherty) [Wlsm. Coll, one].
S. Celebes — low country, 1896 {Doherty) [Wlsm. Coll,
one]; Macassar, 500 feet, 1891 {Doherty) [Wlsm.
Coll, one].
Sanguir — 1892 {Doherty) [Wlsm. Coll, one].
Ombay (= Allor) — Oct., Nov. 1891 {Doherty) [Wlsm. Coll,
two].
Halmahera — Patani, 1897 {Doherty) [Wlsm. Coll, one].
Neu Pommern — Kinigunang {C. Eihbe) [Wlsm. Coll, one
example labelled tengstroemi topotype,” so that
atra2)eo'y may be the species recorded from Kinigunang
by Pagen steelier as tengstroemi, Z.].
N. New Guinea — Dorey, 1897 {Doherty) [Wlsm. Coll,
one].
Kei Isds. — Little Kei Id., April 23,1898 [Meyrick Coll,
one].
D’Entrecasteaux Isds. — Fergusson Id., Sept. 8-30, 1894
{Mcele) [Wlsm. Coll, two].
Mr. T, B. Fletcher on the gemis Deuterocoi^ns. 121
BeuUroeogns lophopteryx, n. s.
(Plate XLIV, fig. 6.)
D enter ocoims atrapex, Fletcher, Spolia Zeylanica, vi, pp.
18-19 (gmrtim).
$ . Exp. 12 mm. Palpi erect, slender, smoothly-scaled, third
joint about half length of second ; ferruginous, banded with darker
apically; third joint ochreous-whitisli broadly banded with blackish
beyond base and at apex. Head brownish-ferruginous, back of head
with some anteriorly-erected scales. Haustellum well developed.
Antennae minutely ciliated (about J)) ferruginous, aniiulated
with darker. Thorax dark brown ; i^ectus whiti.sh. Abdomen
rather stout, dark ferruginous-brown with a nai’row darker dorsal
line, first segment whitish-ochreous dorsally, second segment with
divergent submedian whiti.sh-ochreous patches, fifth, seventh and
eighth segments with small submedian whitish patches ; venter
whitish with a blackish bar across fourth segment. Foreleg pale-
ferruginous dusted with whiti-sh ; tibia with a large apical flattened
tuft of blackish scales ; tarsus darker apically. Midleg pale-fer¬
ruginous ; tibia with a large apical tuft of blackish scales beneath
which projects a pair of moderate spurs ; tarsus lined with whitish
above, darker posteriorly. Hindleg ochreous-whitish heavily dusted
with dark-golden-feiTUginous ; tibia large and stout, provided at
I and apex with whorls of blackish-coppery scales, of which whorls
the second and third are subequal and larger than the first and each
emits a pair of long .slightly-curved blackish-brown spiirs banded
with whitish at middle and at apex and moderately clothed with
short hairs on their i)osterior edges ; of these spurs, the inner
proximal spur is longer than the outer proximal, both tliese being
longer than either spur of the distal pair which are equal in
length.
Forewing short, cleft from |, second segment again cleft from f of
its length ; segments narrow, sublinear : golden-ferruginous much
suffused with blackish-brown ; a whitish patch before base of cleft
surrounded by a more extended area of golden-ferruginous scales ;
first segment with a small ill-defined patch of lighter scaling at of
its length and a minute white dot at f ; both subsegments with a
minute white dot at ^ of their length. Cilia pale-yellow on costa
with a projecting patch of long black hair-scales at f, black between
f and apex but tipped with ochreous immediately before apex ; on
hinder-margin of first segment black with a moderate pale-ochreous
wisp at h and a narrow pale-ochreous wisp before termen ; on fore-
122
Mr. T. B. FiCTclier on the (jenus Deuteroco'j^us.
margin of second segment black with a small pale-ochreons wisp at
on termen blackish, black at base of cilia, the black continued
around apex as a small posteriorly-projecting scale-tuft ; on hinder-
margin of first subsegment black with a pale-ochreous preterminal
wisp ; on foremargin of second subsegment black, tipped with pale-
ochreous on termen ; on dorsum pale-ochreous with a broad blackish
wisp at I and small black scale-tufts at (beneath base of tirst cleft)
and at apex.
Hindwing cleft from about f and segments linear : golden
ferruginous, thickly suffused with blackish. Cilia blackish, a broad
pale-ochreous wisp on hinder-margin of second segment at on third
segment pale-ochreous ; third segment with a large black scale-tooth
occupying exterior fourtli of both margins and a moderate black
scale-tooth slightly before ^ of dorsum.
Type ^ (No. GG52) iu Bainbiigoe Fletcher Coll. :
Ceylon, Central Province, Kandy (IGOO feet), May 1907
{E. Ernest Green).
Besides the type I have seen thirteen other specimens
from the following localities : — -
Ceylon — C. Prov., Kandy, May 1907 {Greeni) [Fletcher
Coll., one] ; Maskeliya (4000 feet) [de Mowbray Coll.,
one]; S. Piov., Yala, Feb. 9, 1909 [Fletcher Coll.,
one].
Amboyna — 1892 {Doherty) [Wlsm. Coll., seven].
Sanguir — 1892 {Doherty) [Wlsm. Coll., one].
S. Ce[.EBES — Macassar (500 feet), 1891 [Wlsm. Coll., one].
Halmahera — [Wlsm. Coll, one labelled “Bernstein, Noord
Halmaheira (Zell. Coll.”)].
Denteroeofus alhiininctatus, n. s.
Deuteroco2nis alh^ninctatus, Wlsm. MS. {ined.).
(Plate XLIV, fig. 7.)
8. Expanse 12 mm. Palpi ei’ect, smoothly-scaled, third joint
shorter than second ; golden-ochreous, ringed with ferruginous
before apex of second joint and at base and apex of third joint.
Head golden-ochreous, top of crown with some anteriorly-erected
scales. Thorax golden-ochreous, suffused with ferruginous ; pectus
whiti.sh. Abdomen ferruginous-golden (details obscured fjy mould),
anal tuft ferruginous ; venter whitish, fourth segment thickly flecked
with golden. Foreleg golden-ferruginous, tibia dilated with an apical
scale-tuft, tarsus lined with whitish above. l\Iidleg ferruginous-
Mr. T, B. Fletcher on the genus Deuterocoims. 123
golden ; tibia with a large projecting apical scale-tuft beneath which
is emitted a pair of moderate spurs of which the inner is the longer ;
tarsus light golden-yellow. Hindleg ferruginous-golden : tibia with
a small whorl of scales at and at f and apex larger subequal whorls
each emitting a pair of long spurs which are banded with whitish on
their second and exterior fourths and very slightly rough-haired on
their posterior edges, tibia banded with whitish beyond first and
second whorls ; tarsus with decreasing posteriorly-directed tufts of
spinous hairs on apices of tarsal joints, first and third joints whitish
on their basal halves.
Fore wing cleft from within first segment narrow, second seg¬
ment again cleft from about of its length, subsegments linear :
ferruginous-golden ; a small longitudinal discal dash at a trans¬
verse whitish area at base of cleft, costal area sufi'used with ferruginous
to beyond J ; first segment cut at J by an ill-defined inwardly-oblique
whitish bar, at b\'’ an ill-defined outwardly-oblique whitish line
and with a minute whitish dot on hinder-margin at f of its length ;
second segment with a whitish spot at base of second cleft ; first
subsegment crossed at J and at apex by narrow whitish bars ; second
subsegment cut at ^ by a narrow whitish bar. Cilia on costa fer¬
ruginous to f, opposite first white bar paler preceded and followed
by blackish, on exterior fourth of costa ferruginous-golden ; on apex
blackish ; on hinder-margin of first segment pale golden-ochreous,
with dark-ferruginous scales beyond } ami between f and 4 fer¬
ruginous, and with a narrow wisp of ferruginous before apex ; on
fore-margin of second segment pale golden-ochreous, on outer third
of segment ferruginous ; around apex of first subsegment pale golden-
ochreous, with a narrow ferruginous posteriorly-directed scale-tuft
beneath apex; on hinder-margin of first subsegment pale golden-
ochreous, with a ferruginous wisp beyond ^ ; on fore-margin of
second subsegment pale golden-ochreous, beyond 5 ferruginous ; on
tornus pale golden ochreous; on dorsum pale golden-ochreous, suf¬
fused with ferruginous at about 4 and with a narrow ferruginous
posteriorly-directed scale-tuft beneath tornus.
Hindwing cleft from about f and from near base, segments linear ;
bright ferruginous-golden. Cilia greyish-golden; apex of third seg¬
ment surrounded by a large spatulate tuft of ferruginous scales
occupying exterior fourth of both margins, dorsum with a large
tuft of ferruginous scales at about i.
Type $ in Walsingliam Coll. ; Japan, Naga, Tokei.
A second specimen, also in Lord Walsingbani’s collection
(No. 62353), was collected in Korea by Ito iu 1887 {ex
Leech Coll).
124 Mr. T. B. Flt^P.ier on tlt.c genus Deuteroco^nis.
The National Collection also contains an example from
the Snowy Valley, near Chekiang (JF. B. Fryer).
Deuterocopus socotranns, Kebel.
(Plate XLIV, fig. 8.)
Deuterocoinis socotranus. Rebel, Denk. Math-Nat. Ak. Wiss.,
Ixxi, part ii, 85-87 (1907).
Dcuterocoyus tengstroemi {ncc Zeller), Meyrick, T.E.S., 188G,
8 {partim)-, Meyrick, J. Bomb. N.H.S., xvii, 134;
Fletcher, Spol. Zeylan., vi, 16-20.
Denterocopus mcdliewi, Wlsm. MSS. {ined.).
Betdeo^ocopius jachsoni, Wlsm. MSS. (ined.).
This clingy little short-winged species, which seems to
be one of the most widely- distributed members of its
genus, appears to have been first collected in New Guinea
by G. F. Mathew in 1884, and in the same year Lord
Walsingham stated (Notes Leyden Mus., vi, 243) that he
had lately received specimens from West Africa. Under
these circumstances it is perhaps somewhat unfortunate
that its first validly published name should have been
derived from its occurrence within the circumscribed area
of Sokotra.
In some examples there is a great extension of the
white scaling on the forewing, giving at first sight a
different facies, but this variation appears to be confined
to individuals and not to coincide with geographical
distribution.
Professor Ur. Rebel, who has very kindly compared some
Sinhalese examples with the original types of this species
from Sokotra, writes that in all essential points they are
in full agreement and that he considers the two forms as
conspecific, although he adds that in socotranus the palpi
appear slightly longer and the coloration a little darker
(less bright red-yellow). This latter point is unimportant,
but the slight difference in the length of the jjalpi, if
constituting a constant difference between the two forms,
may perhaps indicate that we are here dealing with true
local races (incipient new species) ; at joresent, liowever, I
think that we must look on these races (from South aud
West Africa, Sokotra, Ceylon, etc.) as only forming a
single “ species ” in the ordinary acceptation of that word.
The following is a translation of Prof. Ur. Rebel’s original
description of 1). socotranus : —
Mr. T. B. Fletcher on the genus Deiderocoims. 125
“ • Of this distinct genus, whose hitherto known representa¬
tives belong to the Oriental and Papuan Regions, Prof. Simony
brought back from Sokotra three specimens ( d $ ) of a new species,
and of these the least good specimen was sacrificed to the design of
making a microscopic preparation of the neuration. The specimens
were obtained on January 11, 1890, in the mangrove-swamps in
Western Sokotra by beating the flowers of Avicennia marina.
“ Since hitherto there has only been published a short account of
the system of neuration by Meyrick (T.E.S., 1886, 8), and that in
many respects not concordant with the aforesaid preparations, nor
has a figure of the insect or a sketch of its neuration been published
before, both are here given for the first time. [Dr. Rebel has
evidently overlooked Zeller’s figures in Bull. Soc. Mosc. — T. B. F.]
“ The genus is easily distinguished by the thickset build of the
abdomen and the short antennae, which in the are somewhat
roughly scaled towards the tip. The face is closely scaled, the palpi
are slender, their pointed terminal joint half as long as the curved
middle-joint. The legs also are unusually short and stout, the tibiae
with knot-like dilatations of scales, on the fore-tibia only at the
apex, on the mid-tibia at i and apex, on the hind-tibia at J and also
at the origin of both pairs of the very long spurs. The apices of the
tarsal joints also are more moderately dilated with scales, only the
terminal joints remaining bare.
“ The lower fissure of the trifid forewing does not reach so far
basewards as the upper one. The lengths of both clefts are in the
proportions of 3:2. In the neuration of the forewing, the discal
cell is not closed and veins 5 and 6 are so far wanting that they are
reduced to a quite short termination running into the first cleft.
Veins 8 and 9 are long-stalked. Vein 10 arises separate from these,
vein 11 remote from 10.
“In the hindwing the third segment is only half as long as the
first, the cleft between the first and second segments is to the greatest
length of the hindwing as 27 : 43. In this wing also the cell is not
closed, vein 2 also is wanting [the figure, however, clearly shows
that 2 is present and that it is vein 3 which is absent. — T. B. F.],
Amin 3 does not reach as far as the apex of the second segment, Avhose
main support is formed by A'ein 4, 5 and 6 are much reduced, vein 7
traverses the first segment to its apex, A’ein 8 is free.
“ Mej’rick (1. c.), Avho must liaAm described the neuration after only
slight clarification, makes veins 2 and 4 of the foreAving arise from
the angle of the cell (which scarcely agrees with the present species),
vein 3 AV’anting, 5 and 6 A’ery short, vein 11 arising near 10. In the
hindwing A’ein 3 is Avanting, according to his account.
“ Unless another system of neuration be admitted for the example
120 Mr. 'r. B. i^’4^rher on the cjcnus Dcnicrocoims.
of Deulerocupiis Imgstroemi (probably from Java) examined by him,
tliere appears to be an error on Ins ]iart easily arising under the
al)ove-mentioned circumstances.
“So far as at present concerns the species from Sokotra, in any
case it stands very near to both the hitherto-described species, namely
tengstroemi, Z., and mhrodadylus, Pag. Perhaps upon the whole we
have to admit of only one widely-distributed species which has
formed local races.
“ In this species also the general coloration is a bright rusty-
brown, the antennae ringed with black-brown towards the base.
The second and third palpal joints with broad central bands. The
pectus is wdiitish, especially laterally. The ground-colour of the
legs is also reall}' whitish with increasing thicker rusty-brown irrora-
tion towards the ends of the joints, noticeably on the tibial dilata¬
tions. The inner side of the hind-femur as well as the ventral surface
of the abdomen is pure white, the latter with brown edging to the
segments in the 5 only. The naked, very long spurs of the hind-
tibiae are white with a blackish central band especially distinct on
their inner side and blackish tips. Head and thorax thickly sprinkled
with rusty-brown, the abdomen much lighter at the base, yellowish,
in the ^ less thickly sprinkled with rust-brown, in Imth sexes with
light margins to the segments. The anal tuft of the seems to
cover a long uncus.
“The rust-brown irroration of the forewung is in the 9 niuch
more uniform, in the ^ strongly intermixed with light yellowish
scales. Beyond the cleft there lies in the first segment a constantly
distinct pale (yellowish) transverse bar, wdiich terminates on the
costa as pure white. Before and beyond this the first segment is
more or less sufl’used with Idackish. On the hinder-margin of the
first segment the cilia are broadly suffused with blackish and they
are also Ijlackish beneath the apex. The first subsegment shows in
the ^ a distinct yellowi.sh patch before i, and beyond this the cilia
on both margins are blackish. On the apex [of the first subsegment]
lies a deep-black scale-tuft which is surrounded with white posteriorly.
The second subsegment is darker, without markings, with broad
black cilia on both margins ; here also a black point occurs on the apex.
“The segments of the hindwing are rusty-brown with lighter
cilia glistening-yellowish basally, the third segment with a small
black cilial scale-tuft in the middle of the dorsal margin and a
similar broad apical scale-tuft surrounding both margins.
“ The under-side of the wdngs is red-brown with a white-yellow
costal spot at the termination of the transverse band of the ujjper
side and a small bright-yellow spot before the apex of the first
segment. Length of forewing 4*5 mm., expanse 9'5 mm.
Mr. T. B. Fletcher on the genus Deuterocofus. 127
“ Di.stinguished from Deutewcopus tengstroenii by the non-hairy
hinder-tibial spurs and the not bright rust-yellow costal cilia of the
first segment, from Deuterocopns ruhrodadylus by the obvious scale¬
thickening at the end of the fore-tibia.
“As already remarked, it may be that the three described forms
of Deuterocopus (tengstroenii, rubrodadgliis and socotranus) are only
local races of one species, but that can only be proved by comparison
of sufficient material.”
As regards the figure of I), socotranus given by Dr.
Bebel, it is particularly to be reinarketl that the slight
scale-tuft in the dorsal cilia of the forewing beneath the
base of the first cleft has been omitted in error ; this tuft
is actually present in the specimens themselves.
The following is a general description of the species : —
Expanse 9-10 mm. Paljii erect, third joint about half length of
second ; very pale -ferruginous, apices of joints banded with darker.
Antennae above blackish dotted with white, beneath pale-ferruginous.
Head and thorax dark brownish-ferruginous, intermixed with darker
and lighter scales ; pectus white. Abdomen dark ferruginous-
brown, first two segments ferruginous-ochreous variably mixed with
whitish, third segment edged posteriorly by a transverse raised bar
of whitish-ochreous scales, fourth, sixth and .seventh .segments with
narrow submedian longitudinal whiti.sh stripes ; ventral surface
white, fourth segment edged posteriorly with red-brown. Legs
ferruginous-brown, paler or whitish towards base and on tarsi : fore¬
tibia with acuminate scale-tuft on apex ; mid-tibia with apical scale-
tuft and a pair of long spurs ; hind-tibia with moderate whorl of
scales at J and larger whorls at f and apex, the.se two whorls
emitting each a pair of long, naked spurs which are banded with
blacki.sh before apex ; the posterior edges of these whorls of scales
are usually paler or whiti.sh and the hind-tibia is more or less
banded with whitish beyond first and second whorls ; posterior
tarsus with first, third and fourth joints, banded with paler or
whitish.
Forewing cleft from about f, first segment moilerately broad,
second segment again cleft from about \ of its length : very dark
brownish-ferruginous intermixed with paler or whitish scaling some¬
times forming a patch at base of first cleft : costal area suffused with
pale-ochreous, especially evident at about ; first segment cut at
about by an inwardly-oblique transverse whitish line preceded by
pale-ochreous, at f by a .similar but more distinct white line not
reaching costa, segment beyond this much suffused with pale-
128 Mr. T. B. l^'^^cher on the genus Deidcroeopus.
ochreous ; both sulisegmeiits cut at about ^ by transver.se white
lines. (Note. — All the white markings on segments appear to be
variable in extent of development.) Cilia on costa ochreous, blackish-
ferruginous above base and middle of first segment ; on apex
blackish ; on ternien iiale-yellowish ; on hineler-margin of first
segment pale-yellowish to I, beyond that blackish, with a small
black posteriorly-directed scale-tooth beneath apex preceded by a
narrow ochreous wisp ; on upper margin of first .subsegment pale-
yellowish mixed with white, beyond |- blackish continued around
apex of sub.segment to a small posteriorly-directed scale-tooth beneath
apex, on lower-margin pale-yellowish to beyond that blackish ;
on upper margin of second subsegment pale yellowish mixed with
white to about |, lieyond that blackish ; on dorsum pale-yellowish,
with a small blackish scale-tuft at about f (slightly lieyond base of
first cleft), blackish on outer J of second subsegment.
Hindwing cleft from about f and from near base, segments linear ;
brownish-cupreous-ferruginous. Cilia pale-yellowish, darker on
outer half of wing ; third segment with a small blackish scale-tooth
on dorsal margin slightly within J and a large apical blackish-
ferruginous scale-tooth on both margins.
Larva. — The larva feeds in Ceylon on the flowers of the
square-stemmed jungle vine ( Vitis quadrangularis) so
characteristic of the dry districts of that island. The
following is a brief description made on October 20, 1908,
from a living larva found at Hambantota : —
Length 7 mm., stout, stoutest at about miil-length, decreasing
rapidly anally ; head capable of retraction into or under prothorax.
Fig. 3.— Dent, socotraiius.
Incisions between segments distinctly marked. Colour a uniform
pale-green ; head yellowish-brown and prothorax dark blackish-
purple. Prothoracic legs pui-ple, other legs and prolegs pale-green ;
legs and prolegs rather short and stout. To the naked eye no hairs
are visible except two paii’s of short whitish curved hairs on the
anal segment and a pair of short submedian hairs, directed forward,
on each thoracic segment. Spiracles high-placed, about half-way up
the side, fairly conspicuous from being outlined in a slightly lighter
green tint than that composing the general colour of the larva.
Mr. T. B. Fletcher mi the genus Deuterocop^is. 129
Movements slow and deliberate, spinning a thread as it moves along
and when it drops. Under the microscope the skin is seen to he
covered with minute skin-points as if shagreened.
Ptfjoa.— The pupa is attached to a flower, flower-stalk or stem of
the food-plant, or more rarely to a leaf of the same, and is usually
enclosed in a very flimsy cocoon composed of a few silken threads.
It is possible however that these threads are merely fortuitous,
having been spun by the larva during its search for a suitable
pupation-place or whilst preparing its cremastral pad. The pupa
is about G mm. long, stout, smooth, rounded and blunt at the
capital extremity. Its usual colour is a pale apple-green, marked
Fig. 4. — Dent, socotranus.
with dark- or purplish-red on the dorsal surface, the markings usually
consisting of (i) a narrow median thoracic stripe broadening
posteriorly into a transverse bar extending obliquely downwards to
about the edge of the wing-covers, and (ii) a series of sulnnedian
patches on the second to fifth abdominal segments forming a more
or less interrupted longitudinal stripe. Some pupae, however, which
had pupated in my boxes, were wholly of a dark -grey colour. The
moth emerges in the early morning.
Distribution. — As already remarked, the geographical
range of this species is very wide. Localities known to
me are : —
West Africa — Gambia, Bathurst, 1885-87 {Carter^
[Wlsm. Coll., four].
South-East Africa — Delat^oa Bay, 1884 (Druce) [Wlsm.
Coll., three].
East Africa — Ibea, Ukambani, Dec. -Jan. 1889 {Jackson)
[Wlsm. Coll., No. 6330, one J > fyp® jacksoni,
Wlsm. MS.].
SoKOTRA — Western Sokotra, Jan. 11, 1890 {Simony)
[Vienna Mus., ^ and ^ types oi socotranus, Rbl.].
India — Ganjam, Aug. 1882 {Minchin) [Wlsm. Coll., one] ;
Bombay, Surat {Lcfroy) [Meyrick Coll.].
TRANS ENT. SOC. LOND. 1910. — PART II. (JUNE). K
1 oO Mr. T. B. I^TTlc'-ber on the genus D enter oeo'pus.
Ceylon — N. W. Brov., Piittalam, Feb. 1890, March 1902
{Pole) [Wlsm. Coll., four] : E. Prov., Trincoinali,
July 20, 1890 [Brit. Mus., one]: S. Prov., Hamban-
tota, abundant in Oct. and Nov. 1908, about seventy
{Fletcher)-, Yala, larvae common Feb. 9, exc. Feb. 20,
1909, five {Fletcher) : Uva Prov., Madulsima, Coco-
ofalla Estate, 4000 feet, Oct. 1907 and Jan. 1908
(fF. Vaughan) [Fletcher Coll., two].
SuMBA — below 2000 feet, 189G {Doherty) [Wlsm. Coll.,
two].
Tamboha — low country, 1896 {Doherty) [Wlsm. Coll.,
one].
Amboyna — 1892 {Doherty) [Wlsm. Coll., one].
New Guinea — Port Moresby, 1884 (G'. A’. Matheio) [Wlsm.
Coll., two specimens including No. 2287, type ^
mathewi, Wlsm. MS.] [Meyrick Coll., one] ; Humboldt
Bay, April 18, 1893 {Doherty) [Wlsm. Coll., three].
Queensland — Townsville {Dodd) [Meyrick Coll.].
Deuteroeapus hathyclmsma, n. s.
(Plate XLIV, fig. 9.)
9 . E.xpanse 15 mm. Palpi upturned, smootldy-scaled, third
joint about half length of second, second joint somewhat enlarged
apically, tliird slender, acuminate ; golden-ferruginous, ringed with
wliitish-ochreous. Head golden-ferruginous, with some anteriorly-
erected scales on back of crown. Antennae golden-ferruginous,
lined above with whitish, extreme tijjs whitish. Thorax golden-
fernxginous with some whitish-ochreous scales intermixed : pectus
whitish. Abdomen rather stout, light golden-ferruginous, with a
few whitish scales tending to form longitudinal lines especially
evident laterally ; venter whitish, fourth segment with a transverse
ferruginous bar. Foreleg golden-ferruginous ; tibia short, with an
apical tuft of scales. Midleg golden-ferruginous; tibia with an
apical scale-tuft emitting a pair of moderate spurs. Hindleg bright
golden-ferruginous : tibia stout, with whorls of whitish-tipped scales
at I, I and apex ; tibial spurs long, strong, slightly curved, ferru¬
ginous, ochreous-whitish in centre and apically, thickly clothed with
short hairs on posterior edge ; the distal spurs are slightly unecpial,
the inner being the longer.
Forewing rather short, cleft from slightly within f, second
segment again cleft from rather within J of its length ; iirst segment
narrow, subsegments linear : bright golden-ferruginous : base of
wing slightly suffused with blackish tending to form longitudinal
Mr. T. B. Fletcher on the genus Dmterocopus. 181
lines before base of cleft, a white dot in disc at another slightly
before and above base of cleft, and a third before base of second
cleft ; first segment with a white dot at | and another on its hinder-
margin at I, exterior half blackish-suffused ; both subsegments with
a white dot a little beyond i, outer third of first subsegment and
W’hole of second subsegment blackish-suffused. Cilia golden-ferru¬
ginous ; on costa blackish at about f and J ; on hinder-margin of
first segment with a few scattered black scales basally, a blackish-
brown wisp at f and a small black preapical scale-tuft ; on fore¬
margin of second segment blackish-brown beyond f ; apex of first
subsegment with a small blackish posteriorly- directed scale-tuft ;
hinder-margin of first subsegment with a small blackish -brown scale-
tuft at f ; foremargin of .second subsegment blackish-brown beyond
I ; on dorsum with a small brownish-ferruginous scale-tuft at f
(below base of first cleft), a dark-brown wisp along middle third of
second subsegment, and a small blackish tornal scale-tuft.
Hindwing cleft from about f and ^ ; bright golden-ferruginous.
Cilia golden-ferruginous ; third segment with a large scale-tooth
occupying exterior fourth of both margins, a moderate triangular
dorsal scale-tooth at about and a few scattered scales on fore¬
margin.
Type ^ (No. 42084) in Walsingliam Coll. ; Sumbawa,
Tambora (3000 feet), 1896 {Doherty).
Co-type (No. 8528) in Bainbrigge Fletcher Coll., also
collected by Doherty in Tambora (3000 feet).
Two other examples in Lord Walsingham’s collection are
from Batian (August 1897 ; Doherty) and from Kapaur,
S.W. New Guinea (sea-level to 2000 feet ; Dec. 1896 —
e. February 1897 ; Doherty).
Deuterocopus planeta^ Meyr.
(PI. XLIV, fig. 10.)
Deuterocopus planeta, Meyrick, T.E.S., 1907, 4713-4.
Deuterocopus ruhrodactylus, Fletcher {nec Pag.), Spolia
Zeylan., vi, 20, t. E., f. 7.
I am indebted to Mr. E. Meyrick for the opportunity of
examining the type of this species of which his original
description runs as follows : —
“ (^. 10-11 mm. Head and thorax deep ferruginous, sprinkled
with yellow-whitish. Palpi ferruginous, with several whitish rings.
Antennae whitish, with a biserrate fuscous line. Abdomen whitish-
K 2
132 ]\Ir. T. B, inetcher on the genus Deuteroeopus.
yellow, ubove with three suffused stripes and several transverse
bands ferruginous, third segment with a semicircular silvery-white
spot on posterior margin, anal valves and tuft very lotig and slender.
Legs ferruginous ringed with whitish, posterior pair with whorls of
ex])anded scales at origin of spi;rs and apex of tarsal joints. Fore¬
wings cleft firstly from t [? misi)rint for f], secondly from -j, first
segment narrow, second and third linear ; deep ferruginous, sprinkled
with whitish-yellowish ; undefined slender irregular white bars on
first segment before ^ and beyond -|, before base of second cleft, and
on middle of second segment ; cilia pale-ferruginous, on costa
mostly dark-fuscous, with dark-fuscous bars at apex of segments, and
dark -grey patches or bars on lower margin of first segment towards
apex, ujiper margins of second and third segments towards apex, and
lower margins of second and third segments about middle. Hind-
wings cleft firstly from secondly from near base, segments linear ;
ferruginous, first two segments suffused with dark-fuscous posteriorly;
cilia light-ferruginous, third segment’with a dot of one or two black
scales on dorsum before its middle, and a moderate apical scale-tooth
of black and ferruginous scales extending above and beneath.
“ CooRG (Kuti) (Netveome), Assam (Kliasi Hills); in
October and November ; two specimens. Very similar to
rubrodactylus, but the silvery-white spot on abdomen is an
easy distinction. In D. ritsemae, Wals., the abdomen is
also marked with white, but differently, according to the
description.”
To the above description I would add that the ventral
surfaces of the thorax and abdomen are pale-yellow, and
that the white spot on the abdomen is often more or less
obsolete. D.planeta,\\o\veYe\\ is easily distinguished from
ritsemae or ncbrodaetylus by the greater length of the
second cleft and by the absence of any scale-tuft in dorsal
cilia of tlie forewing beneath the base of the first cleft.
Early Stages. — These were briefly described by me in
“ Spolia Zeylanica ” under the name of rubrodaetylus, and
for the sake of completeness the descriptions are here
reproduced : —
Ovum. — The egg is about '44 mm. long by about ’20 mm. broad ;
in shape it is ovo-cylindrical, the ends rounded and subequal, the
micropylar area distinctly depressed ; the surface is very smooth and
shining, of a very pale-orange colour, suffused with red at either
pole.
Larva.- A larva found at Galle on June 18, 1908, feeding on the
flowers of Leea samhucina (Sinh. “ Bouroula ”) was described as —
Mr. T. B. Fletcher on the genus Dcuterocojjus. 133
•• Pale-green without any markings except red suffusion at either
extremity. The skin is roughened into minute knobs (like shark
skin) everywhere, but especially on the ventral region. A distinct
subsegment is formed on the posterior ventral region of abdominal
segments. The hairs, excejjt (i), are very short and inconspicuous ;
(i) is short, less than breadtli of segments. The hairs are trans¬
parent whitish (glassy) and the tubercles very indistinct. The hairs
are longest on thoracic and anal regions. The legs are extremely
short and inconspicuous. There are no secondary hairs, these seem¬
ing to be reduced to skin-points or rather rugosities of the skin.” A
sketch of the arrangement of the tubercles was given in figure 7 of
Plate E of the publication quoted.
Pupa. — A pupa, also found at Galle on June 18, 1908, was brown
with a broad lighter ochreous-fuscous central band ; very few hairs
Fig. 5. — Deut. planeta.
or projections. It was suspended anally to a flower-stalk within a
slight attempt at a cocoon — a few silken threads spun around it to
form a spacious but flimsy enclosure, in wdiich the pupa was fully
visible. The cast larval skin remained at the anal extremity of the
pupa.
Specimens examined by me are from : —
Assam — Khasi Hills, Nov. 1906 [type of planeta in
Meyrick Coll.].
Burma — Bernardmyo, Ruby Mines (5000-7000 feet), June
1890 {Doherty) [Wlsm. Coll., one].
Ceylon — S. Prov., Galle, June 3, June 21, June 23,
October 26, 1908 [Fletcher Coll., four].
Portuguese Timor — 1892, Doherty [Wlsm. Coll., one].
Tenimber (= Timor Laut) — 1892, Doherty [Wlsm. Coll.,
two].
134 Mr. T. B. iMetclier on the genus Benterooo^jus.
New Guinea — Little Kei Isd., April 11, 1897 [Meyrick
Coll., one*]; Feb. 2 and Feb. 14, 1900 {H. Kiihn)
[Brit. Mus, Coll., two].
Deuterocopus ritsemae, Wlsra.
(Plate XLIV, figs. 11, 12.)
Deuterocopus ritsemae, Wlsm., Notes Leyden Mus., vi, 243
(1884) ; Fletcher, Spol. Zeylan., vi, 21 (1909).
Deuteroscopus ruhrodactylus, Pag., Abb. Ges. Zool., xxix, 241.
't. Deuterocopus ruhrodactylus, Meyr., T.E.S., 1907, 473.
Deuterocopus ruhrodactylus, Fletcher, Spol. Zeylan., vi, t. B,
f. C {nee l.c., p. 20).
Lord Walsingbam’s original description is as follows : —
“ Head ferruginous, palpi ferruginous, annulated and tipped with
white. Antennae dull ferruginous, delicately spotted along their upper
side with white. Thorax ferruginous with two ill-defined yellowish-
white spots anteriorly. Forewings bright shining ferruginous, with
two white spots before the fissure, one on the dorsal margin nearer
to the base than to the fissure, another on the middle of the wdng
nearer to the fissure than to the base, a brownish spot at tlie base of
the fissure preceded by a few brownish scales : the costal lobe
clouded about its middle, but not at its base, with bronzy -brown and
having two blackish streaks on the costal margin and another at the
extreme apex. The central bronzy-brown shade is preceded and
followed by a silvery-white spot. The divided lobe is also some¬
what clouded with bronzy-brown but not at its base, a silvery- white
spot precedes the dark colouring and a second smaller one is about
the middle of the upper division, a dark streak in the fringes at the
apex of each division. In the pale yellowish fringes on the dorsal
margin is a tuft of bronzy scales below the first fissure and another
larger bronzy cloud below the second fissure. Hindwings dark-
ferruginous with purplish tinge in the fringes especially towards the
apex of the lobes ; there is a small tuft of darker scales on the dorsal
margin before the middle of the posterior lobe, a spatule of bronzy-
brown scales at its apex. Abdomen bright ferruginous, with a raised
band of shining silvery scales above the middle, a conspicuous
yellowish-white spot preceding the anal segments. The posterior
legs are bright-ferruginous, banded with white on the tibiae and first
* Mr. Meyrick does not agree that this specimen is conspecific
with his type of planeta, but I have been unable to detect any
difference of specific importance.
Mr. T. B. Fletcher on the genus Deuterocojyus. 135
tarsal joint ; with long tibial spurs also indistinctly banded, the
raised tufts at their bases rather less conspicuous than in the allied
D. tengstroemi. Under-side of throax and abdomen very pale
yellow ; the anal tuft ferruginous.
“ Expanse of forewings 11 mm.
“Java (Muli4) ; a single specimen.
“ This species differs from D. tengstroemi, Z. . . . in the presence
of the white spots and in the pale bases of the lobes of the fore¬
wings, also in the less thickly scaled posterior legs and in the absence
of scales in the spurs themselves. . .
The following is a translation of Pagenstecher’s original
description of the form ruhrodactyhis : —
“ In the Ribbe collection there are several specimens of a new species
from Neu Pommern (Kinigunang) which is closely allied to the
preceding (U. tengstroemi, Zeller [= ? atrapex — ^T. B. F.j) and which
Herr Snellen regards as tengstroemi. Dr. Hofmann, however, makes
the following comments
“ ‘ Does not agree with the description of D. tengstroemi, Zeller,
Linnaea Ent., vi, p. 402. The belly, the coxae, the sides of the
pectus and the frontal area are bright sulphur-yellow, as is the
upper-side of the two first segments of the abdomen. The forelegs
have no scale thickening at the end of the tibiae and no spurs, as
described for tengsU-oemi. [As already pointed out, Zeller’s state¬
ment to this effect is evidently incorrect. — T. B. F.] The very
long spurs of the hind tibiae are not thickened wdth hairs. Fore¬
wings brilliant red-brown, with black longitudinal spots and tips
dusted with black. Three segments in the hindwing as in tengstroemi.
The species should be easily recognisable by means of the characters
here indicated.’”
In spite of Dr. Hofmann’s concluding remark, the above
description is quite insufficient for the recognition of this
form, and I have therefore drawn up the following descrip¬
tion from one of Ribbe’s original specimens now preserved
in Lord Walsingham’s collection and labelled as the type
of ruhrodactyhis (No. 73186) : —
5. Expanse 12 mm. Palpi long, smoothly-scaled, erect ; second
joint whitish-ferruginous, broadly banded with dark-ferruginous at
base and before apex; third joint acuminate, rather more than half
length of second, whitish-ferruginous, broadly banded with dark-
ferruginous at base and before apex. Antennae blackish-ferruginous,
dotted with white above. Head ferruginous, summit of crown with
136 Mr. T. B. Fl^R*lier on the genus Deutcrocopus.
erected scales. Thorax golden-ferruginous, mottled with darker ;
pectus pale-yellow. Abdomen large and stout, golden-ferruginous
sprinkled with darker ; first two segments pale-yellow, third segment
with indications of a whitish semi-circrrlar spot on posterior margin
preceded by darker, posterior margins of third and succeeding seg¬
ments faintly outlined in whitish scaling ; ventral surface pale-
yellow. Foreleg whitish-ferruginous-golden, tibia with slight indi¬
cation of scale-tuft at apex. Midleg bright ferruginous-golden, tibia
lined with whitish beneath and banded with whitish before apex
where a dark-ferruginous scale-tuft emits a pair of short equal spurs ;
tarsi whitish. [Hindlegs broken.]
Fore wing cleft from about first segment narrow, second again
cleft from slightly beyond half its own length : bright ferruginous-
golden : costa blackish to J, a short blackish shade along dorsum,
a blackish longitudinal subcostal spot at a blackish suffusion along
costa at ^ leading into first segment and broadly continued down¬
wards into disc but not reaching dorsum ; two black dots preceding
cleft, one above base of cleft, the other below ; first segment with
costal area suffused with black to cut at ^ by an inwardly-oblique
black line, area beyond this wholly black except for a small whitish
dot on hinder-margin at | ; second segment with a longitudinal
whitish spot at area beyond this (including both subsegments)
suffused wdth blackish ; first subsegment with a whitish spot at ^ of
its length, second subsegment with a few whitish-ochreous scales near
its base. Cilia on costa blackish, on first quarter of first segment
ferruginous-golden ; within cleft blackish, w'hitish-ochreous in central
third and with a narrow jjale-ochreous wisp at hinder-angle of first
segment ; on hinder-margin of first subsegment blackish to i, then
whitish-ochreous to just before apex, where blackish ; on fore-margin
of second subsegment blackish ; on dorsum blackish with black scale-
tufts beneath base of each cleft, area between these tufts whitish-
ochreous
Hindwing cleft from | and from near base ; golden-ferruginous,
first two .segments suffused with blackish. Cilia blackish ; on third
segment whitish-ochreous with a large blackish-ferruginous apical
scale-tooth on both margins, a few scattered blackish scales on fore-
margin and a small blackish scale-tooth on dorsum slightly before
Although the two forms, ritscmae, Wlsm., and rubro-
dactylus, Pag., appear abundantly distinct when contrasted
with one another, a long series soon reveals a complete
gradation between them, nor does the development or
suppression of the pure-white markings on the forewing
and abdomen seem to follow the geographical distribution
Mr. T. B. Fletcher on the gemis Dcuterocopus. 137
of the species, specimens with these markings w^ell de¬
veloped or quite obsolete being found side by side in
Assam, Ceylon, Batian, Tambora, etc.
I have examined 37 examples, as under : —
In my oivn collection ; —
Ceylon — C. Prov., Peradeniya, July 1907 {E. E. Green),
one; S. Prov., Galle, June 5, 1907, June 4-24, 1908,
eight, including three bred specimens.
In Lord Walsmgham's collection : —
Assam — Margherita, 1889 {Doherty), two.
Tenasserim — Mergui, 1888 {Doherty), one.
S.E. Borneo — Pulo Laut, 1891 {Doherty), one; Riam
Kiwa, 500-1000 feet, 1891 {Doherty), one.
Tambora — low country, 1896 {Doherty), three ; 3000 feet,
1896 {Doherty), three.
Batian — August 1897 {Doherty), eight.
Bali — 1896 (Doherty), four.
Ternate — 1892 {Doherty), one.
Halmahera — Gani, 1897 {Doherty), one.
N. New Guinea — Humboldt Bay, April 1893 {Doherty),
two.
Neu Pommern — Kinigunang {C. Bibhe), one (type of
rnhrodactylus, Pag.).
Early Stages. — These are as yet practically unknown.
The moth was bred by me from pupae found at Galle,
Fig. 6. — Deut. ritsemae.
Ceylon, suspended anally from the upper surface of leaves
of Leea sambucina, which is evidently the foodplant. The
different method of suspension, as compared with the pupa
138 T. B. FleTTlier on the genus Dmtci'ocopus.
of D. 2')laneta, is noteworthy. On tlie same bush I found
a larva feeding inside an unopened flower-bud ; it appeared
to be very similar to that of but wanted the
terminal red suffusion ; unfortunately I failed to rear it,
Leptodeuterocopus, n. g.
Labial palpi long, smooth-scaled, erect, sickle-shaped. Abdomen
rather short and stout. Legs strong ; posterior tibiae with tufts of
Fig. 7.— Lept. citrogaster.
spinous hairs at about 4 and apex, second and third tufts each
emitting a pair of long spurs. Forewing cleft from about f ; first
segment rather narrow, without hinder-angle ; second segment ex¬
panded posteriorly, again cleft from about of its length. Hindwiug
cleft from about ^ and | ; second segment with a slight scale-tuft
about centre of its hinder-margin ; third segment with a large scale-
tooth surrounding its apex and a small scale-tooth on dorsal margin
nearer the base.
Type : — L. citrogaster.
The two species described below are readily separable
by the following character ; —
Dorsal cilia of f.w. with a large white pi’etornal patch . sochchor aides
Dorsal cilia of f.w. without a large white pretornal patch citrogaster
The slightly different shape of the second cleft in these
species may indicate a generic difference ; otherwise they
appear very closely allied, agreeing with one another and
differing from Dcuterocopus, in the different positions of
the scale-tufts on hind tibiae, the less deeply cleft hind¬
wing, etc.
Leptodeuterocojms citrogaster, n. s.
(Plate XLIV, fig. 13.)
g. Expanse 14 mm. Palpi long, slightly curved, erect, smoothly
scaled; second joint ferruginous-yellow, banded exteriorly before
Mr. T. B. Fletcher on the genus Deuterocoims. 139
apex with dark-ferruginous; third joint more than half length
of second, acuminate, pale yellow, broadly banded at base and
before apex with dark-ferruginous. Antennae dark-ferruginous,
dotted above with pale-yellow. Head dark-ferruginous intermixed
with paler, back of crown with long anteriorly-erected scales.
Thorax dark -ferruginous ; pectus pale-yellow. Abdomen large and
stout, golden-ferruginous, pale-yellow on two basal segments; third
segment edged posteriorly by a narrow band of large pale-yellowish
scales, .sixth segment pale-yellowish except narrowly on anterior
margin ; venter -pale-yellow ; anal tuft dark golden-ferruginous.
Foreleg pale- ferruginous ; tibia short with large apical tuft of dark-
ferruginous scales. Midleg dark golden-ferruginous ; tibia with
apical scale-tuft emitting a pair of unequal spurs ; tarsus whitish,
ferruginous at apex of first tarsal joint and lightly sufifused with
ferruginous on fourth and fifth joints. Hindleg pale-yellowish ;
tibia large and stout with tufts of spinous hairs at about |, t and
apex, broadly suffused with ferruginous- golden above second and
third tufts which each emit a pair of long slightly-curved spurs
broadly banded wdth dark-ferruginous at base and before apex and
roughly clothed with short hairs on their posterior edges.
Forewing cleft from about f, first segment rather narrow, second
segment expanded posteriorly and itself cleft from | of its length,
the second cleft as wide as deep : bright golden-ferruginous : basal f
slightly irrorated with dark-brown ; first segment preceded by a
patch of blackish suffusion from costa to base of fir.st cleft, cut at J
by a narrow inwardly-oblique white line ended on hinder-margin of
second as a small longitudinal white spot, outer half of segment
lightly suffused with dark-brown ; second segment lightly suffused
with dark-brown, with whitish dots on dorsum near base and at J
length of segment and a short white longitudinal streak on fore¬
margin at I length of segment, base of second cleft narrowly edged
with blackish. Cilia on costa ferruginous-golden, on outer half of
first segment blackish to just before apex ; on lower margin of first
segment ferruginous-ochreous, on terminal third blackish to apex;
on fore-margin of second segment ferruginous mixed with blackish
scales, apex of first subsegment with a small posteriorly-directed
blackish scale-tuft ; within second cleft pale ferruginous-ochreous ;
on dorsum ferruginous-ochreous, blackish beyond f, with a small
tuft of blackish scales at about | (just beyond base of first cleft)
followed by a broad wisp of pale ferruginous-ochreous and a small
blackish scale-tooth at tornus.
Hind wing cleft from about | and from near base, golden-ferruginous
irrorated with ferruginous. Cilia ferruginous-ochreous ; second seg¬
ment with a small blackish scale-tuft in centre of hinder-margin ;
140 Mr. T. B. Fl< m ler on the genus Dcuterocopus.
tliird segment with a large blackish apical scale-tooth on both
margins, a few scattered blackish scales on fore-margin and a small
black triangular dorsal scale-tooth at about
Type (No. 42125) and Co-type (No. 41929), both in
Walsingliain Coll, and both collected by Doherty in
Ainboyna in 1892.
Leggtodeuteroco'pus sochchor aides, n. s.
(Plate XLIV, fig. 14.)
Sex d ( D- Expanse 13 mm. Palpi upturned, long, slender,
smoothly scaled, reaching vertex; second joint long, sickle-shaped ;
third joint rather short, straight, acuminate : ferruginous-orange,
banded at joints and striated exteriorly with yellowish- white.
Antennae ciliate (about D> blackish-fuscous, basal third lined above
with short pale yellow streaks, apical portion wholly greyish. Head
bright purplish-golden-reddish, with two lines of pale-yellowish
scales between the eyes, one before and the other behind origin of
antennae ; eyes margined j)osteriorly with pale-yellow. (Abdomen
broken.) Foreleg bright reddish -orange ; tibia short, with two ill-
defined pale-yellowish bands, apex dilated into a small blackish-
tipped scale-tuft ; tarsi darkened apically. Midleg pale-yellowish
suffused and banded with golden-orange ; tibia with a small golden-
ferruginous scale-tuft before 4, apex dilated with a small dark-golden-
ferruginous tuft emitting a pair of long blackish white-banded spurs
of which the inner is the longer; tarsi whitish, golden-fuscous at
apices of joints. Hindleg golden-orange; tibia with a minute golden-
ferruginous tuft at i and large tufts at |- and apex, these two large
tufts emitting pairs of long smoothly-scaled spurs of which those of
the proximal pair are slightly unequal (the inner spur being the
longer), those of the distal pair equal, areas between all these three
tufts narrowly banded with pale-yello%v ; tarsi blackish-fuscous with
a slight golden tinge, first two tarsal joints with small apical tufts of
hair-scales.
Forewing rather long and narrow, cleft from about f, second
segment again cleft from about f of its length : golden-ferruginous
suffused exteriorly with reddish-fuscous : two ill-defined yellowish
spots at I and f, the first costal, the second midway between costa
;uid base of cleft, the space betw'een them blackish-fuscous; first
segment blackish on costa, with a silvery-white square spot on costa
at I length of segment, a white dot in disc at 4, an inwardly-oblique
narrow white line from below costa to hinder-margin at J and a
white costal dot in extreme apex ; second segment broader than first,
expanded posteriorly, with a shallow cleft on termen barely reaching
Trans. Enl. Soc. Loud., igio. Plate XLIV.
H. Knight, del.
SPECIES OF DEUTEROCOPUS.
Andre & Sleigh. Ltd.
Explanation of Plate XLIV.
. <
Trans. Ent^h\ Loud., ipio, Plate XL V.
C. Hentschel.
APPENDAGES OF DEUTEROCOPUS.
Explanation of Plate XLV.
Fig. 1. Denterocopus socotranus, Rebel, genitalia, x 40 (from a specimen
from Hambatonta, Ceylon).
„ 2. Deuterocopiis planeta, Meyr., genitalia, x 40 (from a specimen
from Galle, Ceylon).
f
I
Mr. T. B. Fletcher on the genus Deuteroeopus. 141
to f length of segment, base of first subsegment cut transversely by
a narrow inwardly-oblique white line, second segment with a large
and conspicuous patch of pure white at about \ length of segment
resting on dorsum but not extending to fore-margin of segment ;
dorsum of wing with a few scattered wliitish scales between J and
white ])atch. Cilia on costa and at apex blackish, dotted with
whitish on basal half, wliite opposite white spots ; within first cleft
dark reddish-ferruginous with a purplish tinge, with a few white
scales around base of cleft and some darker scales exteriorly ; on
termen blackish ; within second cleft pale reddish-fuscous on
basal half of lower-margin of first subsegment, followed by a
narrow whitish patch ; around tornus blackish ; on dorsum fuscous-
ferruginous, with a slight purplish-fuscous scale-tuft at f, followed
by a broad patch of pure white extending half-way to tornus.
Hindwing cleft from about f and i, first and second segments
sublinear, second segment with hinder angle nearly obsolete, third
segment linear : golden-ferruginous : first segment wholly blackish,
with a broad white spot at about | opposite white patch on forewing ;
second segment wholly blackish ; third segment about | length of
second, apical fourth dark-purple. Cilia blackish on costa ; white at
apices of first and second segments, pale-fuscous within cleft, pale-
ferruginous on basal half of lower margin of second segment, on
second segment with a narrow ferruginous-fuscous scale-tuft on
lower margin at i, on third segment pale-ferruginous, its apex
broadly surrounded by a dark purplish-ferruginous scale-tooth, and
a few scattered dark purplish-ferruginous scales between scale-tooth
and base on fore-margin and dorsum.
Type in British Museum Collection.
Habitat. — Brazil, Ega {Bates).
Observation. — In size, colour and general appearance
L. sochehoroides exhibits a great resemblance to a species of
Soclichora {S. albipunetella) which was also collected at Ega
by Bates. Both show the same coppery-brown forevving,
and both have a large and conspicuous white spot in about
the same position, but whilst this spot in the Leptodeutero-
copus is on the dorsal margin of the forewing, in the
Soehchora it is on the tip of the first segment of the hind¬
wing. Thus the same effect is produced by two different
methods.
Explanation of Plates XLIV, XLV.
[Nee Explanation facing the Plates.]
( 142 )
IV. Some Eo:periments with Ants' Nests. By Horace St.
J. K. Donisthorpe, F.Z.S.
[Read December 1st, 1909.]
A COLONY of ants may be founded in several ways —
(1) The most simple and ordinary method is that in
which the queen ant, after her marriage flight, starts the
colony herself. She relieves herself of her wings, either
by brushing them off with her feet, or, as I have sometimes
seen myself, by grasping them with her jaws, and removing
them with a jerk. Selecting a suitable spot, she digs a
small chamber in the ground or under a stone, and laying
her egffs she tends them till the first batch of workers are
hatched.
(2) The female, again, may obtain admission into a
small queenless colony of a different species, and there
bring up her offspring. When the host species has died
out, there will remain a pure colony of the queen species.
This has been called “ Temporary Social Parasitism.”
(3) The queen may also enter a small colony of another
species, and killing the workers, take possession of the
pupae. When these have hatched and have helped her
to bring up her own brood, the mixed character of the
nest is kept up by raids on the host species, which is
commonly known as “ slavery ” in ants.
(4) A female may obtain admission into the nest of
anotlier species, and there permanently reside with
her offspring, this has been called “ Permanent Social
Parasitism.”
Now, as is well known, ants, as a rule, strongly object
to the intrusion of strange ants, either of their own or of
another species. Touch and smell are the two principal
senses in ants, and the antennae are the chief organs in
which they chiefly reside. Forel says the members of a
colony know each other by smell and contact. Wasmann
has called their antennae “ touching noses,” and says they
do not know each other personally, but recognise each
other by an intelligent “ parole,” a recognised form of
antennae stroke. Miss A. W. Fielde has carried out a
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE)
Mr. H. St. J. K. Donistliorpe on Ants’ JVests. 143
number of experiments to prove that each of the different
joints of an ant’s antennae has a different function. For
example, the 12th or final joint recognises the home or
nest odour, the 11th recognises personal relations, the
10th the path or track, etc. The authoress also concludes
that the whole nest aura changes every two or three
months. Though these experiments were very carefully
elaborated, I do not think too much importance should be
attached to them ; and this, I believe, is the opinion
of both Father Wasmann and Prof. Wheeler. The
subject is far too difficult and intricate to be settled at
once. Herr Bethe wished to prove that it was only by
smell that ants knew each other, and he found that when
they were washed in alcohol and water, dried, and bathed
in a liquor of crushed ants from another nest, they were
received by that nest. This, however, is only the case
for a short time, the strangers being eventually killed.
Also ants returned after similar treatment to their own
nest are not recognised for a long time. Lord Avebury
has pointed out that ants that had been soaked in water
were not at first recognised by their friends.
Any careful experiments with ants’ nests are therefore
of the greatest value and interest, as a means of helping
those who are endeavouring to clear up these difficult prob¬
lems. I will now give the results of some experiments
with ants’ nests, which touch on the different points
discussed above.
On April 2ncl, 1907, I established a nest of Formica rufa, from
Oxshott, in my study. It contained 12 ^ $ and many ^ 5, etc.
On April 12tli I brought up from the same nest at Oxshott, some
more 5 ? and ^ 9 • They were at once recognised and received
with pleasure, the $ $ being cleaned and led into the nest. On
April 26th, I brought up a $ and some ^ 9 from anotlier nest at
Oxshott, far removed from the first nest. These also, to my surprise,
were equally well received.
These ants must have sprung from the same stock,
since and from Weybridge and Bournemouth were
attacked, and dragged about and killed. I have also been
in the habit of obtaining ants in the spring from the same
nests I took them from the year before and introducing
them into my observation nests, and always found them
well received and undoubtedly recognised. I extract the
144' j\Ir. T^St. J. K. Doiiisthorpo on
following from one of my note-books — “May lOtli, 1907.
Took part of a nest of F. rufa at Weybridge then later
on, “April 7tb, 1908. Got some more ddhris from the
same nest at Weybridge, 4^$, ^ etc.; ants all well
received.” This is the rufa nest I have still, which is
doing very well.
Mr. Keys, of Plymouth, when starting some observation
nests of Formica rufibarlis v. fusco-rujibarhis from Whit-
sand Bay, told me that he mixed ^ ^ and ^ ^ from different
nests in that locality, and that they agreed perfectly well
together. These facts look as if the “recognition method”
is inherited in a common stock, and also appear to
disagree with the theory of the progressive odour of ants.
In the “Ent. Mo. Mag.” for April and May 1909, Mr.
Crawley publishes some experiments with Lasius species,
ants which found their colonies in the simple or primitive
method. He records cases where queens of Lasius
umbratus were accepted by colonies of L. nigcr.
On May 17th, 1907, I obtained and fixed up in a large glass bowl,
a nest of Formica sanguinea from Woking, which contained very
few slaves, and all the ^ ^ were of a small type. (The nest con¬
tained over 60 specimens of Lomechusa strumusa, wliich may account
for the small size of the ^ 5 ? though no pseudogynes had yet been
produced.) Large sangninea ^ ? taken from a nest at Woking,
tpiite near to this one, were all dragged about and killed when
introduced into this observation nest.
lu this case workers of the same species from another
nest in the same locality were attacked and killed.
On April 17th, 1909, I took a small nest of Formica rufibarbis v.
fusco-rufibarbis at Whitsand Bay. It contained a $ and about 25
^ 5 , and I put them into a small plaster nest on April 22nd. No
eggs were ever laid by this $ . On June 1st I removed some of the
9 ^ and introduced them into a small bowl which contained sand
and a 9 of A’, fusca taken at Bradgate Park on May 3rd, 1909.
This queen had laid a few eggs in a small chamber underneath a
piece of damp sponge. On June 27th, I introduced the rest of the
rujibarbis v. fusco-rufibarbis 9 9 • The queen was not attacked, and
on July 4th all the 9 9 collected under the sponge with
the queen. On August 3rd I liberated them all at Eyde in the
Isle of Wight.
Ill this experiment a fusca 9 was adopted by workers of
a dilferent race from a different locality.
145
Some Experiments with Ants Nests.
On .hily I4th Mr. Keys sent me up several different nests of
rufibarhis \ . fusco-rvfibarbis from Whitsand Bay, which contained
many pupae. On August 10th I allowed two of these colonies,
which I had placed in separate compartments of a combined Fielde
and Jannet nest, to mix, by removing the obstruction in the passage
between the two compartments. They were all quite friendly, and
eventually collected all the pupae that were left (many ^ ^ had
hatched from the others) in one side of the nest.
Here ttvo colonies from different nests in the same
locality combined at once when allowed to mix with each
other.
On May 9th I took 5 $ $ and a number of 5 from a nest of
Fonnica-sangmnea at Woking. These I eventually put into a large
bowl with sand, and a damp sponge. The ants burrowed into the
sand under the sponge. On July 23rd I introduced many winged
$ $ and ^ ^ , some pupae and a few ^ 5 from a sangninea nest at
Bewdley Forest. None of these were attacked ! On July 25th the
Woking 9 $ were up under the sponge, and all the ants were
together with the pupae.
Here ants of the same species from quite a different
locality mixed quite peaceably together. This is very
strange ; it may be that as the first colony were under the
sand, and did not come up till tw'o days after the second
lot of ants had been introduced, , the latter may have
acquired the smell or nest aura. Also the first nest was not
very strong as many of the ^ ^ had died. In any case I can
only state what actually occurred.
On April 21st I put a number of ^ 5 of Lasius flavus, which I
had brought up from Whitsand Bay, into a glass bowl with sand.
On May 6th I introduced two $ 5 flnvus from Bradgate Park, these
were accepted by the § ^ on May 8th eggs were laid in a small
chamber under a bit of damp sponge, and the 5 $ were attended by
the 9 • A $ flavus from Portland was attacked and killed when
introduced. The nest was eventually destroyed by mould.
In this experiment $ ? of the same species were accepted
by from a different locality in a nest without a
On June 1st Mr. Forsyth sent me up from Portland a large nest
of Lasius flavus, which contained 3 9? many ? ?, eggs and pupae,
and some 50 Claviger foveolatus. I kept the main nest in a large glass
bowl with sand, and put 2 of the ? ? and a dozen ? ? and
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE) L
146 Mr. ^ St. J. K. Donisthorpe on
Clavigers into a small plaster nest for observation. From June 1st
till the end of August I kept introducing $ 9 from the main nest
into the small plaster nest, and they were always recognised and
well received. On August 9th I introduced ^ 5 oi Solenopsis fugax
taken with Lasins niger at Sandown. These were all killed by the
Jiavus 9 9 •
Here we see that ants from the same nest, separated
for some time, were recognised and well received when
brought together again. The experiment with Solenopsis
was perhaps too severe a test, as the little parasitic ants
had nowhere to hide in the plaster nest.
I now come to my experiments with nests of Formica
fusca and miJibarUs v. fusco-rnjiharhis and $ $ of Foomiica
sanguinea. The modern view of the foundation of colonies
by the Formica rufa, sanguinea and exsecta group supposes
that the $ after her marriage flight enters a small nest of
F. fusca, or one of its races, and takes possession of the
pupae, being accepted by the woi’kers, or killing them if
they prove to be antagonistic. This opinion is held, I
believe, by both Father Wasmann and Prof. Wheeler. It is
certainly the case that no one has ever witnessed, either
in Europe or Ameiica, a $ of the rufa group founding a
colony by herself, as may be seen in Zccsius and Myrmica,
etc. I have observed quite small nests of F. rufa at
Weybridge, which appear to have been quite recently
formed, but I believe these to have split off from older
nests, of which there are large numbers in the locality.
I have also seen individuals of this species at Buddon
Wood moving the whole nest to a new situation ; the
pupae and entire contents of the nest and most of the nest
materials being carried bodily away. For fifteen years I
have known a very large nest at Weybridge. A few years
ago a part of the ants in this nest moved to a spot close
at hand. This year the ants in the old portion have
moved to another spot near to the first new settlement,
the old nest being deserted. Nests may spread in this way,
but this has nothing to do with the founding of a colony
by a single queen. I have no doubt some of the young
queens return to the old nest after their marriage flight,
but the problem is to ascertain the fate of those that do
not. In order to test this question in the most exhaustive
manner, we require a young female just after her marriage
flight, and also a small, or impoverished, fusca nest. The
147
Some Experiments with Ants' Nests.
latter, as Prof. Wheeler quaintly remarks, are as rare
as “hens’ teeth” when one starts to look for them. As
to the former, I have personally never seen or come across
a marriage flight of either rufa, sanrjuinea, or exsecta. I
therefore made up small colonies of fusca and ruflbarhis v.
fusco-rufiharhis by putting a limited number of ^ ^ and pu j)ae
into a combined Fielde and Jannet nest, and introduced
^ $ of sanguinea. I used both old dealated, and doubt¬
less impregnated, $ and young winged virgin $ taken
from sanguinea nests. From the latter I removed the
wings, as Wheeler has shown that when the wings are
removed the ^ acquires the instincts of an impregnated
female.
Before describing my experiments, I must mention that
I kept sanguinea ^ ^ in bowls of sand for months, alone
and together, and they never attempted to lay eggs or
start a colony. When, however, a few pupae, of this or
another species, were introduced, they sometimes collected
them together and sat upon them.
No. 1. A small nest of F. /(tsca $ ^ and larvae taken at Sherwood
Forest on June 13th. On June 24th I introduced a $ sanguinea,
which I had taken from a nest at Aviemore on May 17th. She still
retained one wing, which I removed. The fusca $ 9 ran away at
first when the 5 approached them, but later attacked her. The $ bit
at the /uscas when attacked. In the evening they were still fighting.
The $ did not pay any attention to the larvae and did not try
to conciliate the $ but ran away from them. By June 25th
5 ? 5 had been killed by the $, and the rest were in the passage
between the two compartments with the larvae. June 27th, $ still
attacked a little, several more 5 9 killed. I introduced some
large niger pupae, which the/wscos collected with their larvae. On
J line 28th, the $ appeared to be accepted by the ^ ^ , &s they were
all sitting together, and several § ^ were cleaning the 5 . On
July 2nd, the 9 dead, no doubt from injuries received in the
previous encounters.
In this experiment the ^ was finally accepted by the ^
although she died from injuries received.
No. 2. July 4th, introduced dealated $ saiiguinea, taken at
Woking, May 5th, into small fusca nest with pupae. The 9
apjiroached the pupae and tapped them with her antennae, evidently
much interested in them. The 9 9 removed them, but the 9
148
Mr. H. St. J. K. Donisthorpe on
was little attacked and repulsed ^ . I gave them a little honey,
and the $ and ^ ^ fed side by side. July 5th, the $ had collected
all the pupae into a corner and sat on them, 2 5 9 with her,
but several others were dead and injured. July 6th, 5 on guard
over all the pupae in one corner, all 9 9 killed but 3. These
try to remove i^upae one by one. $ brings them back again. I
introduced some pupae and larvae from a fusca nest from Wey-
bridge. 5 collected them all together into her corner. July
15th, only 2 9 9 left, quite friendly with 9? all sitting together
on the pupae. July 18th, all well and friendly together.
This experiment was quite successful, the 9 took posses¬
sion of the pupae, killed § ^ when attacked, and eventually
became friendly with the remaining two.
No. 3. On July 12th Mr. Hamm sent me up a small fusca nest
from Shotover. It contained many pupae, 9 9 and 2 Atemeles
larvae. I placed them all in a combined Fielde and Jannet nest.
July 15th, introduced a dealated sanguinea $ taken at Woking,
May 9th. The 9 was at once fiercely attacked by the 9 9 • She
was not very aggressive herself when attacked. In the afternoon
she was still being attacked, and held by her legs and antennae by
many 9 9- 16th, 9 "o longer attacked, but has lost an
antennae. July 17th, 9 iiot attacked, in corner by herself. July
18th, 9 dead.
In this experiment the 9 took no notice of the pupae,
did not resist much when attacked, and finally died from
injuries received. One difficulty in these experiments is
that it is not possible to provide a means for the 9 to
escape, if she wished to do so, as she could in nature.
No. 4. July 15th, Mr. Keys sent me up several F. rujibarbis v.
fusco-rufibarbis nests from Whitsand Bay. July 17th, introduced
dealated snagfumeo 9 taken at Aviemore, May 17th. She approached
the p)upae, when she was fiercely attacked by the workers, and killed
the same day.
No. 5. Another rufibarlns v. fusco-rufibarbis nest. July 17th,
introduced dealated sanguinea 9 j from Aviemore. Imme¬
diately attacked by 2 9 9) with which she fiercely grappled and
killed both during the day. July 18th, 9 dead !
Nos. 6 and 7. July 23rd, introduced virgin sanguinea 9 ?
(having removed their wings), taken at Bewdley, July 21st, into
Sovie Experiments with Ants Nests. 149
two rufibarbis v. fusco-rujibarbis nests. Both 5 ? killed the same
day.
No. 8. July 23rd, introduced virgin sanguinea $, from Bewdley,
having removed her wings, into a /usca nest. 5 attacked
and killed same day.
No. 9. July 2.3rd, removed the wings from a virgin 5 sanguinea,
taken at Bewdley, and placed her in a tin with some pupae. July
24th, introduced this $ into the fusca nest from Shotover.
Attacked by 2 ^ 5 , which she killed. Later 5 captured some of
the pupae and sat on them in a corner. The ^ ^ collected the rest
of the pupae into another corner. Later 5 injured another ^
killed one that fastened on to her leg, and captured more of the pupae.
July 25th, all 9 $ killed but one, $ sitting on all the pupae in one
corner. July 26th, 5 carrying pupae about and arranging in
corner, where she sat upon them. I introduced a few rufibarbis v.
fusco-rujibarbis 9 ^ into the nest. When these approached the
pupae the 9 sprang forward and seized them and shook them as a
terrier shakes a rat, and killed them all.
This experiment was quite successful. The ^ killed all
the ^ and took possession of all the pupae.
No. 10. July 25th, introduced virgin 5 sanguinea, from Bewdley,
into a rufibarbis v. f'usco-rufibarbis nest. On J uly 23rd I had removed
the wings and placed her in a small dark tin with a few pupae. 9
was attacked and killed in two hours.
No. 11. July 25th, introduced a virgin 9 sanguinea, from
Bewdley, which had been treated like the last one, into a rufibarbis
Y. fusco-rujibarbis nest. 9 attacked and killed several 9 9- July
26th, 9 dead.
No. 12. July 28th, introduced virgin 9 from Bewdley,
after same treatment, into a rufibarbis v. fusco-rujibarbis nest. Much
attacked by 9 9- July 29th, 9 dead.
No. 13. July 28th, separated 6 rufibarbis y. fusco-rujibarbis 9 9
with pupae into one compartment of nest. August 9th, introduced
virgin 9 sanguinea, from Bewdley, which had shed her wings.
Immediatel}^ attacked by 2 of the 9 ^ > und killed the same
morning.
It will thus be seen that in no single case was a 9
sanguinea accepted by rufibarHs v. fusco-rujiharhis ^ the
9 always being killed, and generally the same day she was
introduced. This ant is a much bolder race than pure
fusca, the latter being a timid and cowardly species. In
150 Mr. H. St. Donisthorpe on Ants' Nests.
nature when a fusca nest is disturbed, the ants imme¬
diately scatter and run away, all disappearing in a very
short time. This is by no means the case with rufibarhis
V. fusco-ru fibarhis.
In future for further experiments I shall only use fusca
^ and shall next try ^ ^ of i^. rnfa.
The two successful experiments -with fusca show that it
is quite possible for a sanguinea ^ to start a colony in this
way, but it seems rather a precarious method to depend
upon.
( 151 )
V. A few words respecting Insects and their Natured Enemies.
By Arthur G. Butler, Ph.D., F.L.S., F.Z.S., F.E.S.,
M.B.O.U.
[Read March 2nd, 1910.]
I WAS much interested in the perusal of Mr, H. Eltring-
ham’s account of his experiments on the edibility of
Lepidopterous larvae by lizards ; for, although it is now
many years since I made experiments with these charming
reptiles, I have for a long time — since 1883 in fact — kept
a large family of living birds, with which I have frequently
experimented.
That a bizarre appearance or startling colours afford
considerable protection to certain caterpillars, in the case
of all birds not intimately familiar with them, is an un¬
doubted fact ; though in the case of birds which see them
daily they afford the caterpillars no protection whatever,
but rather serve as an attraction. Thus many birds which
naturally seek their food upon the ground will not look at
or will be nervous of seizing the caterpillar of Cerura
rinula, w'hereas those birds (like the Tits) which seek for
much of their insect-food among the branches of trees
seize it and tear it to pieces without a moment’s hesitation.
Bright colours, and especially metallic colours, are very
attractive to birds ; consequently the golden chrysalides of
some of the Vanessae and the more metallic Pkisiae, such
as P. chrysitis, are eagerly devoured by many birds ; a fact
w'hich used to be doubted, on the supposition that a bird
would mistake the colouring for actual metal. That birds
reason I have no doubt; but, when in search of food, they
test an object first, and reason about it afterwards when
they have found it unpleasant.
Touching so-called warning colours I am very sceptical ;
a bird has a great appreciation of form and outline, and,
however coloured, rarely attempts to eat a really nauseous
insect after the first trial ; though in the case of some
moderately unpleasant caterpillars, rejected at first per¬
haps because their flavour w’as unusual, I have known a
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE)
on
152 I'^^Arthur G. Butler
hungry bird to take them a second time and acquire a
taste for them.
To many birds, though by no means to all, the cater¬
pillars of Ganoris hrassicac are objectionable, and are
systematically rejected, whereas those of G. rapae and
7iapi are greedily eaten by all, and those of Mamesti'a
hrassicae by most. In like manner the hairy caterpillars
of Orgyia, S])ilosoma and Eiiprcpia are eaten by most of
the larger birds after the longer hairs have been rubbed
off ; but the smaller birds as a rule refuse them.
The larger Geometrid larvae are, I believe, protected
quite as much by their thick skins as by their twig-like
appearance and stiff immobility during the daytime. I
have seen a bird try to break the skin of Biston hii'taria
or TJraptoryx samlucaria by banging it on the ground ;
but, owing to the rigidity of the long body and its lack of
weight in proportion to its length, making no impression
and rejecting it in disgust. Caterpillars fed upon ivy
probably carry the rank smell of the leaves about them,
and this may be offensive to birds apart from the flavour
of the larvae themselves; so that the same caterpillars
when fed upon apple would naturally be accepted without
hesitation by a bird strong enough to break them up.
The caterpillars of Abraxas grossulariata are usually
rejected by birds on account of their acid or acrid flavour,
but Mr. W. T. Page, a member of the Avicultural Society,
found that his Weaver-birds ate them without hesitation,
and the imago of the same species is greedily accepted by
several birds : this may also be the case with the generally
rejected larvae of the Cinnabar Moth, but of late years I
have had no opportunity of testing this point.
Most birds reject with scorn the caterpillar of Pygaera
bucepliala, yet it is the favourite food of the Cuckoo, and
Jays will accept it, although they do not seem to care very
much for it ; if hungry, no doubt they would eat it freely.
I should expect all birds to eat caterpillars of Staui'opus
fagi, on account of its somewhat spider-like aspect when
viewed from the front ; spiders are the favourite food of
all insectivorous birds and are snatched up greedily even
by the tiniest birds, the little Waxbills of Asia and Africa
attacking and devouring full-grown females of Epeira
diademala with avidity.
For a sick insectivorous bird there is no better medicine
than half-a-dozen good-sized spiders for two or three days
Insects and their Naturcd Enemies. 153
ruiiuing. I have on two, if not three, occasions saved the
life of one of my Scarlet Tanagers by supplying it with
this medicine.
Touching terrifying attitudes in larvae I have no faith
whatever, A Sphinx-larva is attacked at once whatever
its attitude. On the other hand, if its anterior segments
are so ornamented as to resemble the head of a venomous
snake I do not for a moment doubt that, whatever its
attitude, a bird would avoid it.
Birds are not terrified by attitudes, and a hungry bird
is rarely deceived by the resemblance of an insect to a leaf ;
he sees the legs and immediately approaches and pecks it,
which usually settles the matter to his satisfaction. It is
against the passing bird not pressed by hunger that the
insect’s resemblance to surroundings is a protection.
In the case of a Stag-beetle, a Mantis, or probably of
some of the larger tropical spiders, the upright attitude
with ojaen claws ready for defence naturally make the
attacks of a bird wary : he leaps from side to side, getting
in a peck and jumping back out of harm’s way until he
has capsized his small opponent, and then (for a time at
any rate) he has it at an advantage. It is a most entertain¬
ing sight to put a good-sized Lucanus cervus into the cage
of a Thrush in which there is space for free movement :
the attitude of the little creature does not alarm but
simply makes the bird cautious and cunning. It always
seems to me that the more enthusiastic of the advocates
of protective assimilation are in too great a hurry ; they
wish to prove that the advantages of protection are far
greater than they actually are ; as though it did not come
to the same thing in the end if an animal’s disguise or
nastiness served at times to protect it as it would do if it
were universally efficacious : it is merely a question nf
time, and Nature has had any amount of time to work
her miracles in. That some caterpillars are more pro¬
tected than others doubtless explains the fact that they
have become a general nuisance, like those of Pygaera
huccphala and Abraxas grossidariata.
Nastiness is the factor which protects insects best from
birds : they will rarely touch the common Soldier-beetle ;
but by far the greater number will freely eat the buzzing
bee-like Eristalis temcx or the wasp-like Sun-flies ; yet
among the larger birds there are not a few which feed
freely upon wasps, like the Bee-eaters, the Jay-Thrushes,
154 Dr. A. G. B^er on Insects and their Enemies.
and others. In the case of Dryonastes (Jay-Thrushes) the
wasp is seized and its tail rubbed backwards and forwards
between the tail-feathers of the bird, in order either to
break the sting or exhaust its venom, before it is eaten.
That many small birds are not afraid of wasps is
evidenced by the fact that they build their own nests
close to those of these much-shunned insects because of
the protection thus afforded against predacious mammals.
Insectivorous birds as a rule do not touch wasps because
they object to the sting, and doubtless the intelligent
little insects are well aware of the fact and therefore do
not object to them as neighbours : possibly an occasional
dead youngster tossed out of the nest may afford a wel¬
come feast for the wasps.
Neither birds nor insects are credited generally with so
much reasoning power as they undoubtedly possess, though
the fact that they do reason is beginning to dawn upon
men’s minds.
( 155 )
VI. Further Notes on two Osmia-sj?ecies of the adunca-^row^j
By the Rev. F. D. Morice, M.A., F.E.S.
[Read March 2nd, 1910.]
In a paper read on December 5th, 1900, and published in
the Society’s transactions for the following year, I offered
some remarks on such (9s?Rm-species of the adunca-gxoxx^ as
were then known to me, and endeavoured to clear up
certain difficulties about their synonymy. Among the
forms discussed in that paper were two, of which I knew
$ $ only. One was so large and striking a form that
it surprised me to find it apparently undescribed. But
undescribed it seemed to be, and I named it manicata n. sp.
The other I ventured to identify with a species described
by Morawitz under the name loti, which name was pre¬
sently “ sunk ” by Gerstaecker (owing in my opinion to a
mistake), and appears in later works only as a “ synonym ”
— the ^ of one • species, the ^ of another. Having since
become acquainted with the ? ? of both these forms, and
having taken manicata, both sexes together, in several
new localities, I offer the Notes here following as a kind of
supplement to my former paper.
1. Osmia loti, Morawitz.
According to Gerstaecker, loti, Mor., is merely the
well-known caementaria, Gerst. But Morawitz expressly
notes that his species was exclusively attached to Lotus,
whereas caementaria, so far as is known, visits only Fchium.
However much, therefore, the description of loti ^ may
have suggested caementaria, it was surely a rash assumption
on the part of Gerstaecker, who (N.B.) does not seem to
have actually examined Morawitz’s types, that the two
must necessarily be identical. If it can be shown that in
or near the region whence loti was described a distinct
species exists, attached to Lotus and not to Fchium, and
having in both sexes the characters assigned by Morawitz
TRANS. ENT. SOC. LOND. 1910. — PART II. (JUNE)
156 Rev. F.^. Morice’s furtlur Notes on
to his loti, the whole theory of Gerstaecker seems to me
to break down. There is no longer any reason for doubting
that Morawitz’s $ and ^ belonged to the same species, nor
for altering the name in either sex. The name “ mora-
witzi, Gei'st,,” must become a mere synonym of loti ^ ; and
if the laws of priority will allow any species to bear still
the name morcmitzi (as to which experts must decide), it
must be applied in future to the morawitzi of Perez and
Schmiedeknecht — a species which I believe I have shown
to be distinct from loti, and consequently from “ morawitzi,
Gerst.,” which is simply the latter re-named for no good
reason !
I now proceed to offer further evidence in support of my
view that such a species does in fact exist, and that the
name of loti, Mor., should therefore be restored to the list of
European 0s7nm-species.
In 1900 I knew only that a ^ Osmia answering as well
as, and in my opinion even better than, morawitzi, Perez, to
the original description of loti was to be found on the hill
called Petit Saleve, within a walk of Geneva, but on the
French side of the frontier, I had a vague impression
that I had taken my specimens on LoUis, and was pretty
sure that they had not occurred on Echium. But I
abstained from mentioning this point till I could verify it,
and rested my argument solely on consideration of the
characters assigned to loti by its author. Since my paper
appeared I have twice been in the neighbourhood of the
Saleve, but once only Avhen the Lotus was in bloom, viz. in
the spring of 1908, On the latter occasion I made frequent
excursions, sometimes alone, and sometimes in company
Avith my old and revered friend the veteran Swiss hymeno-
pterist, M. Emil Frey-Gessner, of the Geneva University
Museum, on purpose to clear up, if possible, the mystery
as to the habits of loti^, and to discover its $. In the first
object I succeeded, again taking ^ ^ with all the peculiar
characters described in my former paper, and finding that
they did occur, as anticipated, only on Lotus cornicula-
tus. But Ave sought in vain for ^ And, although on
examination of M. Frey-Gessner’s cartons containing his
captures of recent years, and also in the collections of the
late M. Tournier, certain $ ^ from the Saleve and else-
Avhere occurred, resembling, but apparently distinct from,
morawitzi. Per., some of Avhich Avere actually ticketed as
found visiting Lotus coo’nictdatus, there Avas no actual proof
tm Osmia-spccies of the adunca-group. 157
that these were the $ $ of my Saleve $ though we both
thought it very likely that they might be so.
Being obliged to return to England I had to bequeath
my problem to M. Frey-Gessner for further investigation,
and he continued to make excursions to the Saleve with
that object in the spring and summer of 1908, which,
however, met with no success, till at last on June 28th, in
company with Dr. H. A. Schulz, he found both sexes of an
Osmia visiting the Lotus — the ^ J agreeing with those
taken by myself, and the $ $ with those which we had
expected would prove to be their partners ! He has
recorded these captures, making kind allusion to my paper
of 1901 and expressing his agreement with its views, in
the Transactions of the Swiss Entomological Society (July
1909), and has also most kindly presented me with several
of the ^ which now' lie before me.
0. loti ^ much resembles caementaria in sculpture and
pilosity, and like tliat species has pale calcaria. But it is
even more like morawitzi, Per., and might easily be mis¬
taken for it without most careful examination. It seems,
how'ever, to be a smaller insect than either morawitzi or
caementaria — at least I have seen no specimen of either
sex more than 8 mm. long, a size which is generally a good
deal exceeded in both the other species. The best char¬
acter, however, by which it can be at once separated from
either morawitzi or caementaria, and which originally led
M. Frey-Gessner to set it apart in his collection, is to be
found in the sculpture of the clypeus. This in the other
species is evenly punctured all over, but in loti is bisected
longitudinally by a smooth and shining carina which is
uniformly developed, and quite unmistakable when once
noticed, in every specimen that I have seen. Nothing of
the sort seems to exist in any other ^ ^ of the group.
And this fact, coupled with the characters of the ^
antennae and 6th ventral-plate, which my former paper
describes in detail, to my mind fully justifies the retention
of this as a distinct species.* Unfortunately Morawitz says
nothing as to the clypeus of his loti ^ ; but, notwithstand¬
ing this omission, I feel practically certain that his species
* M. Frey-Gessner has lately written to me that he finds the usual
habitats of morawitzi and loti differ, the former occurring chiefly
on the higher Alps, the latter on mere hills and in the valleys. Yet I
have also taken morawitzi in North Italy near the sea and at no great
height above it, I think on Echiwm.
3 58 Rev. F^. Morice’s further Notes on
and the present were the same. He gives no character
for either sex that I cannot recognise in the Saleve insects ;
and liis statement that the species is attached exclusively
to Lotus is borne out by all the facts that have come before
me.
2. Osmia manieata, Morice.
In 1900 I could only record two examples of this species
(both ^ ^), one from Algeria, the other taken by the late
Sir S. S. Saunders probably in the Ionian Islands. I
have now quite a long series of both sexes, and can record
it from the following additional localities : Spain (Granada),
South Italy (Taranto and Brindisi), Greece (neighbourhood
of Athens and Olympia, both sexes common in May 1901),
Asia Minor (Smyrna ^ and ^). Its range therefore extends
over the Avhole length of the Mediterranean.
0. mcinieata, in both sexes, is generally at once recog¬
nisable simply by its great size. Its length may extend
to 13 or even 14 mm. (that of adunca only from 9 to 11),
Its breadth is still more remarkable, quite twice (!) that
of a normal cidunea in all my specimens. This regular
difference in size, and still more in proportions, makes
it perfectly easy to separate examples of the two forms ;
and, as shown in my former paper, the ^ ^ differ entirely
in the structure of the concealed 6th ventral-plate. In
the $ $, however, I have quite failed to recognise any
points of detail on which a reliable “ character ” for their
separation can be based. The calcaria, indeed, are usually
(perhaps always) somewhat rufescent in manieatet (black in
adunea), and the antennae also tend in the former species
to show rufescence beneath, but the extent of this rufes-
cence varies. The normal number of wing-hoohs seems to
be greater in manieata than in adunca (13-14 against
11-12); but, as we commonly find large and small exam¬
ples of a single species differing in this wa}^ 1 have some
hesitation in suggesting that such a difference may be
here “specific.” Still when, as in my collection, a long
series of manieata and another of adunca from many
localities are exhibited side by side, the general “ habit ” of
the two forms is so obviously dissimilar that no amount of
common characters can make them seem identical ; and
there can at least be no doubt that the ^ ^ differ markedly
and regularly (for I have dissected many specimens of both)
in the paradoxically developed 6th ventral-plate of the
tivo Osiiiia-spccics of the adunca-group. 159
abdomen. I feel justified therefore in upholding manicata
as a form differing sufficiently from adimca to deserve a
separate name.
Being certainly no rarity in several Mediterranean
countries, it probably figures as a variety of adunca in
many collections, as it did in my own, until I examined
the concealed $ ventral segments.
July 19, 1910.
( 101 )
VII. A Preliminary Revision of the Labiduridae, a family
of the Dermaptera. By Malcolm Burr, D.Sc.,
F.L.S., F.B.S.
[Read February 2nd, 1910.]
Plates XLVI, XLVII.
The family Lahiduridae was erected by me in a recent
paper (1909^) as one of the five families into which I
divide the order Dermaptera. It is well characterised by
the peculiar form of the pygidium ; this organ is i*epre-
sented by a flap of the dorsal sclerite of the last abdominal
segment, which is bent downwards between the branches of
the forceps, so as to present a more or less vertical sur¬
face. The passage from the dorsal surface to the posterior
is marked by an angular fold which is usually sharp and
distinct; only in the curious Chilian genus Gonolahina,
Verhoeff, is the passage gradual and the line of division not
marked.
We may add that the antennae have generally numerous
segments ; the first few after the basal one being as a rule
short.
In the paper referred to above, the group is sub-divided
into six sub-families; this I now propose to raise to nine.
The Echinosomatinae are separated from the Pyragrinae, the
Parisolabinae from the Brachylabwae, and a new sub-family,
the Palicinae has been since added, with a single species.
The arrangement of these sub-families is as follows ; —
1. Corpus valde deplanatum . 1. Paiucinae.
1.1. Corpus baud valde deplanatum.
2. Mesosternum valde angnstatum . . 2. Allostethixae.
2.2. Mesosternum hand angnstatum.
3. Prosternum valde angustatum . 3. Esvhalmexinak.
3.3. Prosternum baud angustatum.
4. Metasternum postice sinuatum.
5. Prosternum antice rotun-
datum . 4. Echinosomatinae.
5.5. Prosternum antice acum¬
inatum . 5. Pyragrinae.
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) M
162 Hr. ]\ralcoln1^)Urr’s Vrdiminary Revision of the
4.4. Metasterniun postice tnm-
catuin, vel vakle productuiii
et convexum.
5. Mesosternum postice rotun-
datiini . G. 1’sai.ina.e.
5.5. Mesosternum postice trun-
catum.
G. Antennae 25-35 segmen-
tatae ; elytra perfecte e.x-
plicata . 7 Labidurinae.
G.6. Antennae ad 15 segmen-
tatae ; corpus apterum.
7. Segmentuni u 1 1 i in u m
dorsale truncatum . . 8. Parisolabinae.
7.7. Segmentuni ultiinurn
dor.sale emargina-
tum ac bilobum . . 9. Brachylabinae.
Sub-family l.—PALIGINAE.
I am inclined to think that it was this species that
Dohrn described under the name Platylabia major. It is
the creature whicli de Bormans took for P. major, as the
specimens recorded by him from Burma under that name
are Palex gjiaxattoides, and nothing else.
It is odd, but not incredible, that he should have
described it in 1900 as a new species, after he had already
liandled it under the other name. I have no doubt about
the identity of de Bormans’ Burmese specimens, since I
possess some of the original ones.
If this is correct, Palex sparatfoidcs, Borm., falls as a
homonym of Platylahia major, Dohrn, and the sub-family
must be named Platylabiinae, and a new generic name
formed for the group represented by PI. thoracica, as I
know of no other species congeneric with P. sparattoides.
For the present, however, I am in a position neither
to assert nor prove the identity, but I consider it exceed¬
ingly probable.
Sub-family 2.—ALLOSTETHINAE.
The sub-family contains the single genus Verb.,
characterised by the triangular prosternum and strongly
narrowed metasternum. I consider A. loiiibohianum,
Verb., a good species, as it appears to be restricted to
Lombok and has a perfectly distinctive appearance. A.
Labiduridae, a famil'y of the Dermaptera. 163
setiger, Verb., and A. maartensi, Verb., I regard as mere
varieties of A. indicum, Hag.
It is necessary to remove to this genus Psalis guttata,
Borm., wliicli is the same as Forficula doriae, Diibr., so the
correct name is now Allostethus doriae, Dubr.
Anisolabis piligera, Borm., has been looked upon as an
aberrant form of the genus Anisolabis, but the pro- and
mesosternum are strongly narrowed, so it must be removed
to the Allostethinae. It may be the nymph of A. indicum,
but as the three syntypes in my possession appear to be
adult, we may provisionally call it Allostethus piligerum,
Borm. ; if it really is mature a new genus must be formed
for its reception, since it is completely apterous.
Sub-family Z.—ESPHALMENINAE.
This group was recently revised by me (1901^), and
I have nothing to add, except that Gonolabis inca,
Burr, from Peru, should be included here ; it is a true
Esplialmenns.
Sub-family 4:.—EGHINOS()MA TINAE.
This group I have recently separated from the Pgra-
grinae, in which Verhoeff included them, on the strength
of the form of the prosternum ; in the former group this
plate has the anterior margin rounded ; in the latter it
is more or less bluntly pointed. The Ecliinosomatinae
is formed only for Echinosoma, Serv., which is essen¬
tially an Old World group; the forceps of the male are
invariably remote, cylindrical and arcuate, and the whole
body is covered with stiff hairs and numerous, blunt,
dilated bristles. All the species have a strong family
likeness, and doubtless several may be advantageously
reduced to the rank of local races or varieties.
A number of the existing species can only be discrimin¬
ated by coloration, as a glance at the synoptical table of
de Bormans will show. Unfortunately, sufficient material
is not yet available to establish a satisfactory arrange¬
ment.
The following notes, however, may be of use for the
identification of species, and may contribute to the estab¬
lishment of a rational arrangement based on structural
characters.
One species has the segments of the antennae clavate ;
M 2
164 ])r. MalcolilFlltirr’ s PrcUniinurij Jievlsion of the
this is £J. parvulum, ])ohrn, the smallest known species,
apparently confined to Ceylon. Brachypterous and mac-
ropterons forms occur. In all other species known to me
the antennal segments are quite cylindrical.
Two species have the sides of the 7th-9th abdominal seg¬
ments in the male acute and carinulate; these are E.for-
hcsi, Kirby, from Dinner Island, and E. distanti, Burr,
from the Transvaal. In all other species known to me the
sides of these segments are rounded or convex, but not
acute nor carinulate.
Three species, all Ethiopian, have the penultimate ven¬
tral segment of the male entire, that is, neither sinuate
nor emarginate. E. wahlhergi, Dohrn, occurs in the African
Continent; the female has a transverse pygidium, and the
pronotum and wings are spotted.
The other two, with the pygidium of the female narrow,
are E. sehalavum, Borm., a small variegate<l species, and
E. holivari, Rodz, a large black species with uniform
tawny wings. Both are confined to Madagascar and
the adjacent islands.
The remaining species known to me all have the
penultimate ventral segment of the male sinuate or
emarginate.
The pygidium of the female is transverse and the
abdomen of the male almost parallel in E. afrum, Beauv.,
and E. horridum, Dohrn. The former Ethiopian, the
latter a Javanese species. In the former the penulti¬
mate ventral segment of the male is gently sinuate, in
the latter decidedly emarginate.
The abundant Oriental E. sumatranum, Haan, with
which I sink E. ivestermanni, Dohrn, as a mere colour-
variant, has a narrow pygidium in the female and the
abdomen of the male is moderately dilated about the
middle : but the pygidium is truncate at the apex. It is
either acute or blunt at the apex in E. Juscum, Bor., and
E. insulanum, Karsch, neither of which are known to
me. I have not yet been able to examine in both sexes
authentic specimens of E. occidentale, Borm., from West
Africa ; E. congotcnse, Bor., from the Congo ; E. yorkeoise,
Dohrn, from Cape York, and E. concolor, Bor., from West
Africa, but I hope before long to be able to compare them
with the other forms.
labid'itridac, a family of the Dcrma'ptera.
105
Sub-family b.-PYRAGRINA E.
This sub-family requires a revision and I now offer the
following observations. The group represents the passage
from the Pygidicra.ninae to the Lahidurinae-, both de
Bormans and Verhoetf have given too much weight to the
superficial resemblance to the former family. But apart
from the essentially Labidurine form of the pygidium, the
smooth and rounded femora, the sternal plates, and the
elytra, are quite distinctive.
The anal or axilliary angle of the elytra is weak, and
consequent!}^ a small scutellum is sometimes exposed, but
this apparently Pygidicranine feature is inconstant. De
Bormans examined 150 specimens of Pyrayra fiiscata, and
found this minute scutellum visible in about half of them.
The elytra are not keeled in any known species; the
body is usually strongly pubescent, but never has the
dilated bristles which characterise the Echinosomatinae ;
the lobe of the metasternum has a sinuous posterior margin
as in that group, and as in the Pygidicraninae, but the
anterior margin of the prosternum is always more or less
pointed.
I include here four genera, all exclusively Neotropical,
of which one is new.
TABLE OF GEN EH A.
1. Pronotuin sublongius quain latius,
antice aiigustatuni. (Segnientum
pemiltimuni ventrale ^ emargina-
tum ; tarsi longi.) . 1. Pyragm, Serv.
1.1. Pronotum transversuni, margine
antico recto.
2. Segmentum pemiltimuni ventrale d
emarginatum . 2. Propyragra, n.
2.2. Segmentum penultimum ventrale
^ integrum.
3. Caput depressum ; tarsi breves . 3. Pyragropsis, Borelli.
3.3. Caput tumidum ; tarsi longi . 4. Echinopsalin, Bonn.
To supplement this table we may add that the head is
depressed and the femora rather heavily thickened in all
the genera, except Echino^isalis ] while tlie tarsi are long
in all, except Pyragropsis.
160 Dr. Malcolm Burr’s Preliminary Revision of the
Genus 1. — Pyragra, Serv.
The genus Pyrayra, as thus restricted, contains three
species, as follows : —
1. Elytra unicoloria, fusca.
2. Statura minore (18 mm.) ; forceps
asymetricns . 1. minor, Borelli.
2.2. Statura majore (23 mm.) ; forceps
symetricus . 2. fuscxita, Serv.
1.1. Elytra fulvo-vittata . 3. dohrni, Scudd.
All authors are agreed in sinking Thermastris, Scudd.,
its type, 27i. hrasiliensis, Gray, being in my opinion,
synonymous with Pyragra fuscata, Serv., the type of
Pyragra ; I also agree with Rehn that Serville’s descrip¬
tion and figure of P.fnscata is perfectly clear, so that it is
unnecessary to employ any other name ; P. ckontalia,
Scudd., and P. sanssurei, Dohrn, are only based on wing-
coloration, and so I sink them as mere trifling variations
of a con.mon and widely distributed species.
I think it probable that the creature described by
Rehn (1903^, p. 300), under the name of Rchinopsalis
brevibractea, is an immature specimen of the same species.
It extends from Mexico to Paraguay and appears to be
common. Pyragra minor, Borelli, is a small species allied
to P. fuscata, occurring in Costa Rica.
Genus 2. — Propyragra, n. g.
Differs from Pyragra in the almost rectangular transverse pronotum,
which is not narrowed anteriorly. In the ^ , the sides of the 6th
and sometimes of the 7th, 8th, and 9th, abdominal segments are
sharply pointed posteriorly, the line joining the points being carried
along as a more or less distinct keel on the sides of the last dorsal
segment, corresponding with the outside ridge of the forceps. In
other respects it resembles Pyragra ; the penultimate ventral .segment
of the male is ample, broad, the angles broadly rounded, and the
posterior margin gently emarginate in the middle. The species are
generally smaller than those of Pyragra.
The type of the genus is Pyragra paragnayensis, Borelli.
The apparently trivial feature of the coloration of the
pronotum appears to be a constant feature. P. paraguay-
ensis seems to be a southern form. P. brunnea is only
known from Peru. P. buscJci is known from Jamaica and
167
Labiduridae, a family of the Dermajptera.
Cuba ; the type is preserved in the collection of the
United States National Museum.
TABLE OF SPECIES.
1. Valcle pubescens : segmentum ulti-
nuini dorsale ^ supra utrinque carina
acuta instructuin.
2. Pronotum antice fuscum, postico
fulvo maculatum ac lineolatum . 1. paraguayensis, Bor.
2.2. Pronotum fuscum, utrinque
maculata singula fulva ornatum 2. brunnea, Burr.
1.1. Corpus fere glabrum ; segmentum
ultimum dorsale ^ carinula super¬
ior! nulla . 3. buscl'i, Caudell.
Genus 3. — Pyragropsis, Bor.
This is a monotypic genus founded by Borelli for F.
tristani, from Costa Rica. It is a handsome species, of a
strikingly contrasted black and yellow coloration.
Genus 4. — Echinopsalis, Borm.
This genus still contains only E. guttata, Bonn., as
E. hrevihractea, Rehn, I consider to be a larva of Pyrogra
fuscata (q. v., ante, p. 166).
I have in my collection a single female from Peru,
which may be referable to this genus; the head is de¬
pressed and brick-red in colour, the pronotum, tawuy, with
black band, as also the elytra and wings ; the tarsi are
Possibly Fsalis thoracica, Serv., is referable here ; it is
certainly a Pyragrid. The specimen figured by de Borman
(1893, p. 4, PI. I, fig. 7) is a female ; it is very small, being
only 6.5 mm. long.
Sub-family Q.—FSALINAE.
This sub-family includes those genera in which the
metasternum is truncate posteriorly, but the mesosternum
rounded, except in the monotypic genus Titanolahia, in
which the metasternum is produced into a lobe, rounded
at the extremity. All the genera include rather stoutly
built species ; the forceps are strong and trigonal, never
very long, nor slender, often subcontiguous in the male,
168 ]^i‘. Malcol in Barr’s Freliminary Revision of the
and frequently asymmetrical, rarely toothed ; many of the
genera are apterous.
TABLE OF GENERA.
1. Melusterjium ])ostice in lobum angustuin,
apice rotunclatuin, productmn ; (corpus
apterurn; abdomen subparalleluni)
1.1. Metasternum postice tnincatiiin.
2. Abdomen d hasi ad apicein am-
})liatum ; (forcipis braccliia d hasi
remota; corpus apterurn) . . . .
2.2. Abdomen pone medium plus minus
diktat um, apice subangustius.
3. Elytra omnino desunt .
3.3. Elytra rudimentaria, vel perfecta.
4. Elytra rudimentaria ; alae nullae .
4.4. Elytra libera, perfecta ; alae
saejrius adsunt.
5. Antennae segmentis 17-22 . . .
5.5. Antennae segmentis 20-30 . .
]. Titfinolobis, n. g.
2. Gonolabis, Burr.
3. Aiiisoldbis, Fielj.
4. EuboreUia, Burr.*
5. Faalis, Serv.
6. Lahicliirodef^, Borm.
Genus 1. — TitanolohiSj n. g.
Cum genere Anisulabide congruet; ab eo dift’ert pedibus brevibus,
crassi.s, metasterno margine postico in lobulo apice rotundatum
producto.
Type of tlie genus. — Anisolahis colossca, Idohrn.
The powerful build and the great size attained of this
giant among earwigs have always led me to consider it as
forming a distinct group; an examination of the sternal
plates shows that the posterior margin of the metasternum
IS produced with a long, parallel-sided, apically rounded
lobe, which is quite distinctive.
I have never examined one of the Burmese specimens
reported by de Bormans, but I doubt it they are referable
to the true A. eolossea.
The range of size of this creature is very remarkable.
I have in my own collection a female whose length,
inchiding the forceps, which are always short, is 22 o mm.,
and another, which attains 51 mm., both from New South
* Borellia being praeoccujiied by Rebn (1906, Proc. U.S. N. M.,
XXX, p. 379), 1 have projiosed the new name EnboreUia, Burr. See
Proc. U.S. Nat. Mus., xxxviii, p. 448 (1910.)
Lahiduridae, a family of the Bermmptcra. 169
Wales. Perhaps we may later be able to discriminate two
species.
Genus 2.— Gonolabis, Burr.
The genus Gonolalm has been subject to two distinct
sources of confusion.
First Verboeff's characterisation of his sub-family Gono-
laMdae, with G. lativentris as type, excluding by this very
characterisation G. javana, which liad been originally
chosen as type.
Secondly the doubt as to the identity of the true
G. javana, under which name no less than three species
have been confused in collections.
The first confusion has been cleared up by the separation
of Verhoeff’s '' Gonolahidae” into a separate sub-family, the
Esplialmcninac. (See Burr, 1909^.)
The second confusion is now cleared up, thanks to the
courtesy of Dr. Gestro of Genova and Dr. Holdhaus of
Vienna, who have kindly communicated the respective
types of G. sumatrana, Borm., and G. javana, Borm.
Thus I have been able to examine authentic specimens
of every known species of the genus, the catalogue ot
which now stands as follows : —
1. G. Idrhyi, Burr, Java. Type examined.
2. G. electa, sp. n., Java-Ceylon. Type examined.
3. G. sumatrana, Bonn., Java and Sumatra. Type
examined.
4. G. ohlita, sp. n., Java. Type examined.
o. G. javana, Bonn., Java. Type examined.
6. G. michaelscni, Burr, Australia. Type examined.
7. G-incea, Borelli, E. Africa. Syntype examined.
8. G. moodwardi. Burr, Australia. Type examined.
9. G. verhopffi., Burr, Australia. Type examined.
From this list will be observed that I have been able
to examine and compare the types of every known species,
except G. picea, and of that authentic syntypes kindly
given me by Dr. Borelli, which is almost the same thing.
It will also be noticed from the above list that the
genus is chiefly characteristic of the Oriental and Aus¬
tralian regions, a single species occurring in the Etliiopian
region.*
* Since writing the above I have seen the type of Anisolabis
azteca, Dobrn, from Mexico : it is a female, but evidently a Gonolabis.
It will be discussed in a later paper.
170 Dr. Malcolm Durr’s Preliminary Revision of the
It will be observed that the Neotropical G. inca, Burr,
is omitted. This species is a true Psphalvienus, and
should have been included in the revision of that group.
(Burr, 1909h)
The genus Gonolabis is akin to Anisolabis, differing in
the apically dilated abdomen of the male, and fewer
antennal segments.
TABLE OF SPECIES.
1. Segmentum penultimum ventrale
late rotundatnm (necnon apice ipso
subrecto).
2. Segmentum penultimum ventrale ^
medio depressum, utrinque carina-
tum ; forceps asymetricus ; (species
javana) . 1. kirbyi, Burr.
2.2. Segmentum penultimum ventrale
5 planum, baud carinatum;
forceps symetiiciis.
3. Abdomen segmentis 7-9 d lateribus
acutis ; (statura ininore ; abdomen
^ valde dilatatum ) . 2. electa^ Burr.
3.3. Abdomen segmentis 7-9 lateribus
rotundato-convexis.
4. Segmentum ultimum dorsale
margine postico truncato, rectis-
simo ; (colore rufo, pedibus uni-
coloribus) . 3. sumatrana, Borm.
4.4. Segmentum ultimum dorsale ^
margine postico subsinuato ;
(colore atro, pedibus annulatis). 4. oblita, sp. n.
1.1. Segmentum penultimum ventrale ^ $
angustum, lateribus obliquis, apice
subtruncato, fere triangulare.
2. Segmentum penultimum ventrale
medio carinato ; (species javana) . . 5. javaim, Borm.
2.2. Segmentum penultimum ventrale ^
laeve.
3. Segmentum ultimum dorsale
utrinque plica cristata instruc-
tum ; (statura majore ; species
westralica) . . 6. michaelseni. Burr.
Lahid^iridae, a family of the Dermaptera, 171
3.3. Segmentum ultimiim dorsale ^
plica cristata nulla.
4. Forcipis bracchia ^ nec dentata
nec denticulata ; (colore ater-
rimo ; species africana) .... 7. picea, Borelli.
4.4. Forcipis braccliia ^ ornata ;
(species australicae).
5. Forcipis bracchia prope basin
siiperne crenulata . 8. woodwartli, Burr.
5.5. Forcipis bracchia (J prope basin
superne dente forti armata . . 9. verhoefi. Burr.
1. Gonolahis kirhyi, Burr.
Anisolabis? Icirbyi, Burr (1897'^), p. 311.
This Javan species is well characterised by the form of
the penultimate ventral segment of the male, which is
broad and rounded but depressed in the middle, with a
well-marked ridge on each side.
The abdomen is also more strongly and more abruptly
dilated towards the apex than in most other species, thus
resembling G. electa, and the forceps are asymetrical, the
right branch being bowed inside the left.
It is so far only known from the unique male in my
collection, taken by Friihstorfer at Pengalengan, in
Western Java, at an elevation of 4000 feet.
2. Gonolabis electa, Burr.
Gonolabis electa. Burr (1910^), p. 79, fig. 21.
Gonolabis javana, A., Burr (1908^^), p. 78.
This species is probably represented in many collections
under the name of G. java.na, together with G. oblita.
Several years ago Dr. Borelli called my attention to the
fact that apparently two species were confused in col¬
lections under this name, and inquired my opinion. It
was not, however, until the opportunity of examining de
Bormans’ type of G. javana presented itself, that it
became obvious that neither of these two was the real G.
javana, and consequently new names became necessary
for them both.
G. electa, which occurs in Ceylon and in Java (c.m.) is
one of the smallest of the genus, and has the most strongly
dilated abdomen, in correlation with which we find the
172 Di\ Malcolm Burr’s Frdiminary lirvidon of the
sides of the 7th-9th abdominal segments strongly acute in
the male. The female may be recognised by her small
size, deep colour, and broad abdomen,
3. Gonolahis sumairana, Bonn,
(tonolabis sumatrana, Bonn. (1900^), p. 452.
This species is discriminated by de Bormans from G.
javana only by the coloration. From the true G. javana
it differs in the form of the penultimate ventral segment
of the male, but it is probable that in the synoptical table
given by de Bormans (1900^), p. 451, he includes in G.
javana the common species wliich we discriminate under
the name G. ohlita.
A comparison of the type of G. sumatrana with a good
series of G. ohlita shows that though quite different in
appearance, tliere are few structural characters to separate
them.
G. sumatrana is somewhat larger and of stouter build,
of a uniform reddish brown, with plain feet and antennae ; *
the pimctulation of the abdomen is finer, and the pubes¬
cence is very dense in the type, but is apparently worn off
in a second specimen from Java, in the Leyden Museum.
4. Gonalahis ohlita, sp. n,
Gonolahis javana, Burr (1902), p. 479.
Gonolahis javana, B., Burr (1908^’)) P-
Statiira parva ; colore nigro-castaneo ; anteiinai' pedescpie annu-
lati ; abdomen vix dilatatum ; segmentum penultinnini ventrale ^
late rotundatum ; segmentum ultimum dorsale laeve, margine pos-
tico subconcavo ; forciins braccliia d tyjiica, arcuata.
d ?
Long, corporis . . . 9-10 mm. . . . 9'5-12 inm.
Long, forcipis . . . „ ... 1-2 „
Of medium size and rather slender build ; colour deep shining
reddish black.
Antennae black, ringed with white before the apex ; with 13-14
segments.
Head smooth, tumid ; sutures obsolete.
Pronotum subquadrate, slightly longer than broad, and slightly
173
Lahiduruhtc, a fumihj of the Derma ptcro .
widened posteriorly, sides straight ; prozona soniewluit tumid,
separate<l by a transverse depression from the metazona.
Meso- and metanota typical.
Sternal plates typical, pale yellow.
Legs yellow, femora broadly banded with black ; tarsi very slender.
Abdomen punctulate, gradually and gently widened towards the
apex ; sides of 7th, 8tb, and 9th segments in ^ convex but not
acute. Last dorsal segment quite smooth, shining, transverse,
with a sharp low keel on each side in correspondence with the outer
edge of the forceps, and a slight protuberance on each side of the
hinder margin, in correspondence with the upper edges of the
forceps ; between these two points, tlie hinder margin is subconical ;
in female similar, but decidedly narrowed apically and simple.
Penultimate ventral segment broadly rounded, in ^ rather
obtusely triangular.
Pygidium narrow, comj)ressed.
Forceps with the branches remote, stout, trigonal, unarmed,
straight at first and tapering ; apical third smooth and arcuate ;
in $ nearly straight, contiguous, stout, trigonal, and tapering.
Java (c.m.), Buitenzorg (c.in.), Malang (e.rn.), Am-
barawa (c.m.).
This species has invariably been considered to be de
Bormans’ G. javana, but the penultimate ventral segment
of that species is absolutely distinctive. Under the
common name of G. javana it has been confused with G.
electa, but the gradually and gently dilated abdomen and
longer body give it a totally distinct appearance.
It is structurally nearest to G. snmatrana ; the deep
red-black colour, black-banded femora, and pale ringed
antennae give it a different appearance ; it is smaller in
size and slenderer in build ; the sculpture of the abdomen
is coarser ; the abdomen also is less pubescent ; the forceps
in G. oblita are arcuate, that is, bent inward at a curve,
whereas in G. snmatrana, they are more usually bowed,
that is, bent at an angle.
This species has a strong superffeial resemblance to
G. woodwarcli, but differs in the totally unarmed forceps,
apart from the different penultimate ventral segment of
the male.
It appears to be common in -Java.
De Bormans himself confused it with G. javana, as I
possess a female which came from that collection under
that name. •
174 Dr. Malcoln'^iurr’s Preliminai'y Revision of the
5. Gonolabis javana, Bonn.
Anisolahis javana, Bonn. (1883), p. 63, PI. II, fig. 4,
(19002), p. 27.
Anisolahis java7ia, Bonn. (1900^), p. 451 partim.
Gonolabis javana, Burr (lOOO^), p. 49 (nec 1902, p. 479).
Most collections have specimens under the name of
G. javana, as we have seen under G. electa and G. oblita,
but it is probable that no authentic specimen exists
beyond the type in the Brunner collection, now in the
Vienna Museum.
In size and general appearance it closely resembles
G. suniaGrina.
The head and pronotum are brick red, with black
markings, the general colour is a dull blackish red.
Absolutely distinctive in the form of the penultimate
ventral segment, which is figured by de Bormans ; it is
in the form of a blunt pointed obtuse triangle, with a
prominent central compressed crest or ridge.
The tyi^e is from Java, and is numbered 14, 6996.
6. Gonolabis 7iiichaelse7ii, Burr.
Go7iolabis 7nichaelse7ii, Burr (1908^^), p. 73, PI. I, fig. 4.
„ „ do. va7\ de7itata, op. cit. p. 7 5,
PI. I, fig. 5.
This striking species, the giant of the genus, is described,
figured and discussed in detail in the work quoted.
7. Go7iolabis picea, Borelli.
Gonolabis picea, Borelli (19072).
This, the only known Ethiopian species, was discovered
by H.R.H. the Duke of the Abruzzi at Butiti, in Eastern
Africa.
It is remarkable for its intense pitch black colour.
8. Go7iolabis Tvoodioardi, Burr.
Go7iolabis 7ooodwa7'di, Burr (1908^^), p. 75, PI, I, fig. 1.
„ „ var. dentata, op. cit., PI. I, fig. 2.
This species, which is abundant in Western Australia,
is discussed, figured, and described in the Avork quoted.
Lahiduridae, a family of the Dermaptera. 175
9. Gonolahis verhoeffi.,* Burr.
Gonokibis verhoeffi, Burr (1905^), p. 487.
This species is well marked by the prominent teeth on
the upper surface of the forceps of the male, near the base.
In addition to the two males in the Hope Museum,
Oxford, I have received a third from Kuranda, Queensland,
sent me by Mr. Simmonds.
Genus 3. — Anisolabis, Fieber.
This genus contains about forty species, even after the
separation of those furnished with rudimentary elytra into
the genus Enhorellia. It requires a thorough revision, as
a good many names will require to be sunk in synonymy,
and perhaps one or more new genera may be necessary.
For instance, the gigantic Anisolabis colossea, Dohrn, from
New South Wales, may be conveniently separated owing
to its produced and apically rounded metasternal lobe.
I will not, however, offer a revision yet, as the material is
not complete.
Immature specimens of Psalis are frequently mistaken
for some of the less well-marked species of Anisolabis.
Some of the smaller kinds seem to pass insensibly from
one form to another, with subtle distinctions that are
difficult to express. My personal inclination is to sink all
species which are not clearly defined upon some well-
marked structural character, though it is not always easy
to say which forms should be fused. In this respect, as
in the analogous case of Labidura riparia, I assume the
attitude of a frank “lumper.” j-
The genus Anisolabis has become less unwieldy now
* Since writing the above, I have identified this species with
Anisolabis brunneri, Dohrn, of which I have seen the type. A.
bnmneri of Burr is a totally different creature.
f Since going to the press, further material has been examined,
which will be dealt with in a later paper. But it may be mentioned
here, with reference to the above table, that A. incerta, Borm.,
appears to be a melanic form of /estae, Borelli ; that vl. etero-
nonm, Borelli, is in my opinion indistinguishable from A. anmdipes,
Luc., as also A. aporonoma, Borelli.
The species referred toby me as A. brunneri, Dohrn, and recorded
and figured by me in (1908*9) P- I* hg-
brunneri of Dohrn, which is the same as Gonolahis verhoeffi, Burr ;
the A. brunneri figured by me is a well-known species which requires
a new name, unless it can be identified with A. pacijica, Erichson.
170 ]^r. Malcoln^^ inn’s Prclbiiinanj Revidnn of iltc
that nine species with nuiiinentary elytra have been
removed to form the genus Eiiborellia, and A. colossea
separated in a new genus Titcmo/abis.
The number is furtlier reduced by the removal of
ijcra to the Allostethinae, and by the sinking of a number
of names, such as A. hormand, Scudd., A. iintoni, Dohrn,
and other well-known synonyms, under A. annulipe!^.
There still remain a good many whose exact position is
still doubtful ; such are A. coporonoina, Bor., A. (jaiulens,
Burr, A. spectahilis, Phil., A. pacijica, Erichs., A. anrjulifera,
Dohrn, A. pectorcdis, Etsch., A. annuHcornis, Blanch.
I am not yet certain which of tw'o kindred forms is the
true A. margvnalis of Dohrn.
A. alhovittata, Burr, is probably immature. A. advena,
Mein., A. subarmata. Kirby, and A. antenna, Kirby, are
unrecognisable ; as also A. peregrina, Mjbberg, A. pluto,
Rehn, which is the female of A. angidifera, Dohrn. .T.
major, Brulle, is probably the larva of A. maxima, Bridle.
The following table is not intended for a scientific
discrimination of species, but purely as a provisional
arrangement, as help-notes for identification.
TABLE OF SPECIES.
1. Oculi desunt . 1. caeca, Bor.
1.1. Oculi adsunt.
Abdomen d segmentis 6-9 lateribus
rotundatis, vel convexis, neqne
acuminatis, necpie carinulatis, nec
striolatis.
3. Statura gracili ; forcipis braccbiis
elongatis, attenuatis ; anten-
narum segmentis 4 et 5 cylind-
ricis.
4 . Statura minore ; species bawai-
ensis . 2. perkinsi, sp. n.
4.4. Statura maj ore; species africana 3. i:osseleri, Burr.
3.3. Statura robustiori ; forcipis brac-
chiis curvatis vel subrectis, hand
elongatis ; antennarum segmentis
4 et 5 fere globularibus.
4. Forcipis braccliia d margine in-
terno excavata . 4. xenia, Kirby.
4.4. F'orcipis braccliia d margine in-
terno baud excavato.
Lahkluridae, a family of the Dermaptera.
177
5. Forcipis bracchia ^ dentata.
6. Colore fusco-castaneo ; capite
rufescenti ; species austra-
lica . 5. occidentalis, Kirby.
6.6. Colore piceo ; species equa-
toria . 6. incerta, Borm.
5.5. Forcipis bracchia inermia.
6. Femora fusco-annulata ; spe¬
cies japonica . 7. mory/maiis, Dohrn.
6.6. Femora fulva ; species ha-
waiensis . 8. eteronoina, Bor.
2.2. Abdomen segmentis 6-9 lateribus
acutis vel valde convex is.
3. Abdomen ^ segmentis 6-9 lateribus
striolatis et carinulatis.
4. Forcipis bracchia ^ basi remota,
valde arcuata.
5. Forcipis bracchia intus den¬
tata.
6. Corpus atrum ; species aus-
tralica . 9. hrunneri, Dohrn.
6.6. Corpus pallescens; species
equatoria . 10. festae, Bor.
5.5. Forceps ^ inermis.
6. Statura minore ; femora
maculata ; species africae
orientalis . 11. teilinii, Bor.
6.6. Statura majore ; femora
fulva ; species algerica . 12. mauritanica, Luc.
4.4. Forcipis bracchia ^ basi sub-
contigua, paullo arcuata.
5. Segrnentum ultimum dorsale
(J lateribus externis baud
carinatis; caput rufum; pedes
unicoloribus ; species africae
orientalis . 13. laeta, Gerst.
5.5. Segrnentum ultimum dorsale
margine externo carinu-
lato.
6. Caput pallidum ; pedes uni¬
colores ; species africae
orientalis . 14. compressn, Bor.
6.6. Caput nigrum ; pedes annu-
lati ; species cosmopoli-
tana . 15. anmtUpes, Luc.
TRANS. ENT. SOC. LOND. 1910. — PART III. (nOV.) N
178 Dr. Malcolm Burr’s Preliminary Revision of the
3.3. Abdomen ^ segmentis 6-9 lateri-
bus punctatis vel stiiolatis, baud
carinulatis.
4. Segmentum ultimum dorsale ^
utrinque tuberculatum.
5. Forceps ^ valde arcuatus ;
species ceylonica .... 16. liudagae, Burr.
5.5. Forceps ^ elongatus, apice
tantum arcuatus ; species
canariensis . 17. maxima, Brulle.
4.4. Segmentum ultimum dorsale
inerme.
5. Forcipis braccbia ^ basi remota,
fortius arcuata, symmetrica.
6. Forcipis bracchio dextro ^
dente rectangulari armato.
7. Caput nigrum ; species
cosmopolitana . . . .18. maritima, Bon.
7.7. Caput rufum ; species
afric£E orientalis . . .19. felix, Burr.
6.6. Forceps inermis ; species
australica . 20. littorea, White.
5.5. Forcipis braccbia basi sub-
contigua, vix arcuata.
6. Forcipis braccbia ^ intus
crenulata ; caput rufum ;
species birmanica . . .21. duhronii, Kirby.
6.6. Forcipis braccbia ^ baud
crenulata ; caput nigrum ;
species africana . . . .22. infelix, Burr.
Anisolabis perhinsi, sp. n.
Statura mediocri, gracili ; colore fusco-castaneo et nigro ; antennae
segmentis 4 et 5 baud globularibus ; abdomen gracile, vix dila-
tatum, segmentis lateribus laevibus, baud acutis ; pedes longi,
graciles ; forcipis braccbia ^ $ subcontigua, triquetra, subarcuata,
attenuata, inermia.
c? ?
Long, corporis . . . . 13 mm . 15 mm.
j, forcipis .... 3 „ .... 3*5 „
Of medium size; build slender; general colour dark chestnut
brown to black.
Labiduridae, a family of the Dermaptera. 179
Antennae with 16 segments, dark brown ; first long and slender,
apically clubbed ; third long and slender, cylindrical ; 4th and 5th
cylindrical, each about half as long as the 3rd, the rest gradually
lengthening, all slender and cylindrical, the apical segments about
as long as the third.
Head dark brown, smooth and shining ; eyes small.
Pronotum slightly longer than broad ; gently widened posteriorly,
all margins straight, sides gently refiexed.
Meso- and metanota transverse, typical : all the thora.x dull deep
brown, with a few brown bristles ; smooth.
Sternal plates lighter brown ; prosternum almost parallel sided;
mesosternum rounded ; metasternum truncate posteriorly.
Legs long and slender, dull brown ; tarsi very slender.
Abdomen slender, feebly dilated beyond the middle and apically
attenuate in both sexes, exceedingly finely punctulate; sides of
segments 6-9 in ^ rounded and smooth.
Last dorsal segment ^ 5 slightly narrowed apically, with a
median sulcus, posterior margin subsinuate, with a slight convexity
corresponding to the upper ridge of the forceps.
In forceps with the branches ^ 5 subcontiguous at the base, tri¬
gonal, rapidly attenuate, feebly arcuate, more so in the ^ than in
the $ , slender ; black, with a fulvous spot on the outer face near
the base.
Sandwich Islands: Kaui, Koholuamano, 4000 ft., and
Waimea, iv/95 and vii/96 (Perkins).
This is evidently the species quoted by Brunner (P.Z.S.,
1895, p. 892) and by Perkins (Fauna Haw., 1899, p. 4)
as A. pacifica, Erichs., but it is not “ half the size of
F. aiLricularia," nor do the forceps correspond to Erichson’s
description.
There are specimens in the British Museum, and a pair
in my collection, the male of which is my type.
The two sexes are very much alike ; the slender body
and limbs distinguish it, as much as the relatively long
fourth and fifth segments of the antennae.
According to Perkins, it is found only in the mountains
of Kaui.
Genus 4. — Euborellia, Burr.
This genus was erected recently, under the name
Borellia, by me (1909^, p. 325) for those species, formerly
included in Anisolabis, with distinct rudimentary elytra,
named in honour of my good friend Dr. Alfredo Borelli, of
N 2
180 Dr. Malcolm Burr’s Prcli'iiiinary Revision of the
Turin, who has clone valuable work upon the Dermapterous
fauna of the Neotropical and Ethiopian Regions, but as
the name is praeoccupied by Rehn (1906, Proc. U. S. N. M.,
XXX, p. 379), I have suggested a new name Enborellia. It
contains at present nine * species, which may be arranged
as follows : —
1 . Elytra libera per luar^iiieni suturaletii,
usque ad apicem contigua, sub-
quadrata ; species americanse.
2. Pedes uuicolores, testacei; anten-
narum segiuenta 1 et 2 Hava,
ceteris flavo-brunneis .
2.2. Femora fusco-aiinulata ; anten-
iiarum segmenta 1 et 2 rul'e-
scentia, ceteris brunneis . . .
1.1. Elytra baud contigua, lateralia, vel
saltern per miuiniam partem mar-
ginis suturalis contigua.
2. Elytra in parte basali angnsta,
me.sonotum in modum scutelli
liberantia ; in parte postica valde
dilatata, ad suturam attingentia,
metanotum obtegentia ; species
ceylonica .
2.2. Elytra angustata, lateralia, nequa-
quam contigua.
.3. Forceps ^ dentatus; species
americanae
4. Forceps d dente magnoarmato ;
segmentum ultimum dorsale
lateribus plicis 2 instruc-
tum ; colore aterrimo . .
4.4. Forceps dente parvo arma-
tus ; segmentum ultimum
dorsale lateribus tuberculis
2 instructum .
1. iimhigna, Borelli.t
2. jayi.eirensis, Dobrn.
3. (jreeni, Burr.
4. peruviana, Borm.
b. armata, Borelli.
* Anisolabis andreinii, Bor., from Eritrea bas rudimentary elytra,
and so tbis species also must be included bere, and also Anisolabis
minuta, Caudell.
■f Since writing tbe above, I bave compared syntypes of E. ambkjna,
Borelli, with autbentic specimens of E. ianeirensis, Dobrn, with
wbicb I now sink it as a bomonym.
Lahiduridac, a family of the Dcrmaytera.
181
3.3. Forceps $ inerniis ; species
mundi antiqui.
4. Forceps ^ valde asymetricns ;
species tasmanica .... 6. tasmanica, Borm.
4.4. Forceps ^ regiilariter vel fere
regulariter arcuatus vel
curvatus.
5. Femora liaud annulata.
6. Abdomen laeve ; pronotum
planum; caput thoraxque
fusca ; species europaea . 8. moeda, Gbnb.
6.6. Abdomen ptunctatum ;
pronotum pi’ozona tu-
mida ; caput tlioraxque
pallescentia ; species
indica . 9. anncmdalei , Burr.
Genus 5. — Psalis, Serv.
This genus only differs from Anisolahis and Euborellia
in having fully developed organs of flight. Carcinophora,
Scudder, must fall, the only real difference from typical
Psalis being the abbreviation of the wings alone ; as the
character is insufficient to justify specific rank, it cannot
support a genus.
I now include it in the following sixteen species : —
1. Pronotum parallelum ; species ameri-
canae.
2. Elytra abbreviata, scutello saepius
j'atenti ; elytra maculata ....
2.2. Elytra perfecte explicata.
3. Elytra maculata.
4. Statura mediocri; (14-18 mm.);
elytra ad humeros maculata.
5. Corpus glabrum ; abdomen seg-
mentis 6-9 lateribus acutis .
5.5. Corpus pilosum ; abdomen
segmentis 6-9 baud acutis
4.4. Statura majore ; (18-25 mm.)
Elytra disco maculata . . .
3.3. Elytra unicoloria.
4. Caput longurn . . .
4.4. Caput brevius.
1. fediva, S]>. n.
2. perchewn, Guer.
3. roKenhergi, Burr.
4. americana, Beauv
5. fusca, Borelli.
182 Dr. MalcoH^]’) hit’s rreliminary Revision of the
5. Antennae seginentis 4-5-6
longis ; pronotuin longius
qnain Latins . 6. ciagatina, King.
5.5. Antennae scginentis 4-5-6
globularibns ; pronotuin
aeqne longnm ac latum . 7. nifjra, Caiul.
1.1. Pronotuin postice ampliatum.
2. Elytra triangularia ; species orientalis 8. castetsi, Bob
2.2. Elytra rectangularia.
3. Antennae tenues, segmentis apice
valde clavatis.
4. Alae nullae; species americana 9. imddyi, Burr.
4.4. Alae perfecte explicatae ;
species orientalis . 10. femoralis, Dohrn.
3.3. Antennae fortiores, cylinclricae
vel subcylindricae.
4. Antennae segmentis 4-5-6 globu-
laribus, omnibus brevibus . . 11. Dohrn.
4.4. Antennae .segmentis, 4 brevi,
5 et 6 longioribus, ceteris
subcylindricis.
5. Antennae segmento primo
brevi ; species americana . .12. burri, Borelli.
5.5. Antennae segmento primo
longo.
6. Antennae segmento 3 brevi.
7. Colore fulvo-castaneo,
hand metallico ; species
americana . 13 satdderi, Bonn.
7.7. Colore atro caeruleo-
nitenti ; species ori¬
entalis . 14. duhrni, Kirby.
6. Antennae segmento 3 longo ;
S2iecies africanae.
7. Antennae segmento 4 glo¬
bular! ; pronoto brevi,
scutello parvo liberanti . 15. debilis, Burr.
7.7. Antennae segmento 4
cylindrico ; pronoto
super elytra producto 16. cincticollis, Gerst.
Rsalis f estiva, sp. n.
Allied to P. americana ; differs-in the smaller size, more
slender build, abortive wings, and strongly abbreviated
183
Lahiduridae, a family of the Dermaptera.
elytra. Length of body 18’5-23 mm, ; of forceps S’oOmm.
^ ^ unknown.
Brazil, Bahia (Mus. Vienna),
This species is represented by three females in the
Brunner von Wattenwyl collection, now in the Hof-
museiim, Vienna, and will be fully discussed in a forth¬
coming work on the material of that Museum.
Psalis gagatina, Klug.
I have recently discussed the identity of this species
(1909®, p. 126). I consider that Carcinophora rohusta,
Scudder, with its synonymous Psalis columhiana, Borm., is
the real P. gagatina of Klug.
Psalis nigra, of Caudell, from Trinidad, is only known
from the female ; the almost square pronotum and short,
thick antennal segments appear to be characteristic
features.
Psalis cinciicollis, Gerst., is discussed elsewhere in a
recent paper (1909®, p. 113). Psalis picina, Kirby, falls as
a synonym.
Psalis thoracica, Serv., I believe to be referable to the
Pyragrinae (q.v,, ante, p. 167).
Sub-family l.—LABIDURINAE.
The group includes Lahidura, and its immediate allies ;
in structure it agrees in many respects with the Psa-
linae, but the mesonotum is less rounded posteriorly. The
organs of flight are all well developed ; the antennae are
multisegmentate ; the forceps rather slender and usually
remote at the base.
TABLE OF GENERA.
1. Pedes breves ; femora postica pronoto
baud lougiora ; tarsorum posti-
corum segmentis 1 et 3 subaeque
longis, pronoto unitis baud longi-
oribus (pronotum postice ampli-
atum) . 1. Paralabidura, n. g.
1.1. Pedes longi ; femora tarsique pos-
tici pronoto longiores ; tarsorum
posticorum segmentum primum
tertio longius.
184 Dr. Malcolm Burr’s Preliminary Revision oj the
2. Pronotum postice anipliatuni, loii-
"ius (piam latius ; sciitello nullo ;
femora postica pronoto circa 1^
longiora.
.3. Abdomen lateribus integris . . 2. Labidura, Leach.
.3..3. Abdomen lateribus spinulis vel
plicisconstatis armatum . . 3. Forc^yida, ’Bo).
2.2. Pronotum quadratum ; .scutello
magno ; femora postica pronoto
triplo longiora . 4. Tomopyijia, Burr.
Genus 1. — Paralabidura, n. g.
Cum genere Labidura congruet ; differt statura minore et graci-
liori, tarsis brevioribus, posticis pronotum longitudine baud vel vix
superantibus, segmentis primo et tertio subaequantibus.
Differs only from Labidura in the smaller size and slender build,
and shorter tarsi, the posterior pair scarcely, or not at all, exceeding
the pronotum in length, the first and third segments approximately
equally long, and the third segment slightly exceeding the first.
This genus is raised for the reception of the small, dark
slender species, hitherto included in Labidura.
In all known species the colour is dark chestnut.
The posterior tarsi are of different lengths in the dif¬
ferent species, but in no case does the posterior pair
exceed the length of the pronotum, and first, second and
third segments are either equal, or the third a little
longer than the first ; in Labidura and Forcipuki the first
segment is the longest.
The type of the genus is Paralabidura lividipcs, Duf.
TABLE OF SPECIES.
1. Forcipis bracchia margine interno
baud laminato . 1. lividipes, Dufour.
2. Tar?i postici pronoto breviores ; for¬
ceps arcuatus, hand depressup.
2.2. Tarsi postici quam pronotum
aeque longi ; forceps ^ elonga-
tns, depressus . 2. tenuicornis, Bonn.
1.1. Forcipis bracchia margine interno
laminato.
185
Lahiduridac, a family of the Dermwptera..
2. Forcipis braccliia ^ parte laminata
parallela, apice rectaiigula, species
indica . 3. nepalensis. Burr.
2.2. Forcipis bracchia parte laminata
in dentem valde aentnm pro-
ducta, species africana .... 4. Jigini% Burr.
Faralabidura lividipes, Duf., is a widely distributed and
familiar species ; the development and armature of the
forceps vary considerably, as would be expected, and
consequently there are several synonymous names.
P. tenuicornis, Bonn., is a larger, but allied, species from
Sumatra.
P. nepalensis, Burr, has laminate forceps ; it is only
known from Nepal, and is figured in a recent work (Burr,
I910h fig. 30).
P. fginii. Burr (1908^-, p. 176), also has laminate
forceps : it is based on a pair from Eritrea, in the Genoa
Museum. It is interesting to note that it has a strong
resemblance to some undescribed species preserved in
Baltic amber of Oligocene age.
Genus 2. — Labidura, Leach.
This genus now includes only L. hengalensis, Dohrn,
and the polymorphic L. riparia, with its numerous forms,
mutations, varieties, races, sub-species, or even species,
according to the individual opinion of every author. The
synonymy is consequently very involved, and it is diffi¬
cult to assign the correct name to the various forms.
Kirby (1908) has given a useful preliminary arrange¬
ment; he separates the apterous Neotropical race into a
distinct genus, Demogorgon, and is followed by Borelli, but
I can hardly agree with either. Personally I cannot
allow more than specific rank to these large, pale, apterous
forms, and not even that with a feeling of real conviction.
I can only say that certain well-marked geographical
races stand out, as pluvialis, Kirby, from New Guinea ;
truneata, Kirby, from Australia; a small apterous form
from India, that I have provisionally identified with “ var.,
inermis" of Brunner, and xanthopus, Stal, from South
America ; hengalensis, Dohrn, from Bengal, is of no higher
rank.
186 Dr. Malcoll^lj urr’s Preliminary Revision of the
Genus 3. — Forcipula, Bol.
This is a well-marked genus, containing several large
earwigs, occurring in tropical parts of Asia, Africa and
America, having the sides of the abdomen furnished with
spines, tubercles or crests in the male, and elongate
slender forceps. I offered a revision of the genus in
1904^, p. 288, and now suggest the following modified
arrangement : —
FORCIPULA. •
1.1. Abdomen segmentis nonnnllis lateri-
bus crLstis obliqnis serratis in-
structis.
2. Colore nigro, elytris rufis ; species
americana . 1. americaim, Borm.
2.2. Coiore fulvo ; species indica . . 2. hirida, Bol.
1.1. Abdomen lateribus segmentis non-
nullis spinis armatum.
2. Abdomen lateribus segmentis 3-6
spinis binis armatum ; elytra
brevia ; alae abbreviatae ; species
indica . 3. decolyi, Borm.
2.2. Abdomen segmentis nonnullis
spinis singulis armatum.
3. Elytra costa carinula nulla ;
species indicae.
4. Statura majore 3.0-43 mm. (alae
longae ; abdomen spinis 4
armatum) . 4. pugnax, Kirby.
4.4. Statura minore (16-20 mm.,
alae breves ; abdomen spinis
2 vel 3) . 4a. var. minor. Burr.
3.3. Elytra costa carinula instructa.
4. Elytrorum carinula debili, fere
laevia.
5. Segmentum ultimum dor-
sale inerme ; forceps con-
strictus ; abdomen spinis
3 ; colore fusco ; species in¬
dica . 5. irispinosa, Dobrn.
Lahiduridae, a family of the Dermaptera. 187
5.5. Segmentum iiltimum dor-
sale utrinqiie tnberculo
erecto magno armatum ;
forceps sinuatus ; colore
fulvo ; species africana . 6. congo, Burr.
4.4. Elytra granulosa, carinula
fortiori.
5. Forceps constrictus ; species
orientates.
6. Forceps ad angulum den-
tatus . 7. tarsata, Westw.
6.6. Forceps ad angulum in-
ermis ...... 8. quadrispinosa, Dohrn.
5.5. Forceps sinuatus.
6. Abdomen spinulis 3 ; pygi-
dium trispinulosum ;
forceps unidentatus ;
species americana . . 9. quelchi, Burr.
6 6. Abdomen spinulis 4 ;
abdomen inerme.
7. Statura majore t34-38
mm.) ; forceps uni¬
dentatus ; species afri¬
cana . 10. gariazzi, Bor.
7.7. Statura minore (21-22
mm.) ; forceps basi
denticulatus ; species
orientales.
8. Forceps rectus ; abdo¬
men granulosum,
spinulis recurvis . 11. jacobsoni, Burr.
8.8. Forceps undulatus ;
abdomen punctul-
atum ; spinulis
rectis .... 12. walker i, Kirby.
Genus 4. — Tomopygia, Burr.
This genus was erected by me in 1904, p. 287, for the
curious creature from Java described by de Bormans
under the name Cylindrogaster abnormis. It is undoubtedly
allied to Labidura and Paralabidura, but the abbreviated
elytra, broad short scutellum, small, square pronotum and
exceedingly long and slender legs, are all strong characters.
188 Dr. Malcolm Burr’s Prcliminfir]i Bevidon of the
Sub-family ^.—PABISOLABINAE.
This group was formed by Verhoeff for a district New
Zealand species which he described under the name Pari-
solahis novae zee-landiae (1904, p. 120). I have recently
added the genus Pseudisolahis (1908^, p. 254) for P.
walkeri, also from New Zealand. Later still, I have
raised the groiip to the rank of sub-family (1910\ p. 102).
Tlte genus Pseudisolahis now contains also/*, hurr\ Borelli,
and P. tenera., Burr, both Indian species.
Sub-family 9. —BRA CHYLABINAE.
This interesting group was separated by de Bormans
from Anisolabis, with which it had been unnaturally
arranged previously, but it has been little known until
recently.
The group is interesting on account of several appar¬
ently primitive features. In a number of genera we find
that the eyes are not spherical, as in most earwigs, but
ovate or elliptical, and very large, extending from the
insertion of the antennae backwards almost to the hinder
margin of the head ; at the same time, the head itself is
not pentagonal, but almost triangular, so that in these
particular genera the eyes may be said to be truly
lateral.
That this large size and lateral position of the eyes is
a primitive feature is indicated by the discovery of a
number of fossil earwigs in the famous Tertiary Lake
Basin of Florissant, in Colorado, from which IScudder
described and figured no less than eleven species.
These all show an approach to a primitive type in the
uniformity of character, simplicity of structure, and general
similarity, but the most remarkable feature is the great
size and lateral position of the eyes. This is best seen in
his figs. 2 and 3 (Z. avia), fig. 12 (Z. cxsulatum). (Scudder
1890, Tert. Ins., PI. XVI.)
So much was Scudder impressed by this feature that he
erected a species genus, Lahiduromma, for their reception.
There are other features, too, in the Brachylahinae that
appear to be primitive ; for instance, the forceps are in¬
variably simple ; they are, in fact, the simplest imaginable
type of forceps, the branches being cylindrical, that is
circular in cross section, absolutely unarmed, tapering and
Liihiduridae, a family of the Dcrmaidera. 189
very gently arcuate. The only sexual dilference in the
forceps is that in the female the branches are somewhat
closer together at the base than in the male.
The pauci-segmentate antennae may perhaps be looked
upon as another primitive feature, as also the spindle-
shaped body. In many or all of the species there is a pair
of curved impressions on the frons between the eyes ; can
these be vestiges of ocelli ? These organs are unknown in
earwigs.
The total absence of organs of flight is probably to be
explained rather by degeneration than by non-develop¬
ment, since one African form, Arlex sjoestedti, Borg.,
possesses signs of rudimentary elytra on the mesonotum
analagous to the same rudiments in Karschiella, which is,
moreover, also a primitive type, its larva probably having
segmented caudal setae instead of forceps, as in its near
ally Borniansia, and the not distantly related Diplatys.
The legs in the Brachylahinae are very simple, and show
neither keels on the femora nor specialisation in the seg¬
ments of the tarsi. The second segment is decidedly
longer than in most other earwigs. I can detect no trace
of pulvillus between the claws, nor are the claws different
from those of other earwigs. The Brachylahinae all have a
strong family likeness and cannot be confused with any
other group.
Apart from the various features enumerated above, the
last dorsal segment has the posterior margin emarginate
or concave, and the two lobes are pointed and slightly
produced over the forceps. All known species are deep
dull black in colour, and probably are clothed in life with
a long velvety pubescence, which is generally worn off in
cabinet specimens.
They appear to be rare insects, but are widely distributed
throughout the tropical regions, occurring in Java, Burmah,
India, Ceylon, Madagascar, Africa, Guatemala, Brazil, Peru,
etc. This wide distribution of so well-marked a type of
earwig is probably also an archaic feature.
Verhoeflf was the first to draw attention to the great
size of the eyes in certain Brachylahinae : in his paper on
“ Neue ungefluegelte Eudermapteren-Gattungen” (SB. Ges.
Naturfr. Fr. 1901, p. 10) he makes a special point of this
feature in distinguishing his family Isolahidac. 'I'he words
he uses are : “ Augen sehr gross, hoechstens um ^ ihres
Durchmessers von Hinterhaupte entfernt.”
190 Dr. Malcoli^Burr’s Preliminary Revision of the
In a recent paper I revised tlie Bracliylabinae (Ann.
Mag. N. H. (8) vol. ii, pp. 246-355, 1908), showing that
Verhoeff’s Isolahidae coincide with the already known
Brachylabinae.
In that paper I sunk Ctenisolabis, VerhoefiF, in Brachy-
lahis, Dohrn, not because I considered C. togoensis (type of
Ctenisolabis) congeneric with B. chilensis (type of Brachy-
labis), but because Verhoeff’s diagnosis of Ctenisolabis gives
no character not common to Brachylabis.
But the consideration of the eyes, suggested by the
study of fossil forms, suggests a modification of this
arrangement, because B. chilensis, the type of Brachylabis,
has eyes which are large, it is true, but do not reach
back near to the posterior margin of the head, being
more normally situated.
Most of other species of Brachylabinae known to me
have lateral eyes, and this appears to be the case in Cteni¬
solabis togoensis, Verb., judging from a sketch of the type
made for me by Herr Flandetsky in the Berlin Museum.
It would accordingly appear necessary to eon^we Brachy¬
labis to its type, B. chilensis, and to revive Ctenisolabis
with its type, G. togoensis, Verb., and also for some of the
other species formerly included by me in Brachylabis,
separating others into a new genus which stands in exactly
the same relation to Isolabis that Ctenisolabis stands to
Leptisolabis.
This modification of my former views, suggested by an
examination of further material, was first put forward in a
recent work on the Earwigs of India, the faunistic scope of
which, however, prevented the treatment of the whole group.
I accordingly now offer a revised system, with sundry
observations, upon all forms known to-day.
TABLE OF GENEEA.
1. Oculi parvi, anteriores.
2. Elytra rudimentaria adsuiit . . . 1. Arlex, n. n.
2.2. Elytra omnino desunt.
3. Mesonotum carinatuni ... 2. Brachylabis, Dohrn.
3.3. Mesonotum hand carinatuni . 3. Nunnisolabis, Burr.
1.1. Oculi magni, laterales.
2. Antennarum segraentum 3 elon-
gatuin, duplo longius quam
latius ; 4 sat longum hand
globulare.
191
Labiduridae, a family of the Dermaptera.
3. Mesonotum liautl carinatum . 4. Isolabis, Verb.
3.3. Mesonotum carinatum ... 5. Metisolabis, Burr.
2.2. Antennarum segmenta brevia,
3 baud vel vix longiori quam
latiori, 4 globulari.
3. Mesonotum carinatum ... 6. Gtenisolabis, Verb.
3.3. Mesonotum baud carinatum . 7. Leptisolabis, \ erh.
SCHEME OF GENERA OF THE BRACHYLABINAE.
A.
Rudimentary Elytra present . . Arlex, n. n.
B.
No rudiments of Elytra.
Mesonotum.
Genus 1. — Arlex, n. n.
Verhoefia, Burr, 1908^, p. 248.
This is the only member of the family showing any
trace of rudiments of elytra. This feature is well shown
in Borg’s figure (1904, p. 568, Taf. XXVI, fig. 2), as also
the normal situation of the eyes.
The genus is monotypic for Brachylahis sjostedti, Borg.
(/. c.), from the Cameroons.
The name Verhoeffia is praeoccupied by Broleman in
Myriapoda in 1895, so I proposed the new name
Arlex.
Genus 2. — Brachylabis, Dohrn.
Brachylahis, Dohrn (partim) (1884), p. 297.
„ Bonn. (1883), p. 64.
„ Burr (1908»), p. 248.
192 Dr. Malcolm Burr’s 1 Preliminary Revision of the
The type of this genus is Brachylahis chilensis, Blanchard,
from Chile,
It is very unfortunate that the true Chilian B. chilensis
is not better known ; it is not represented in any collection
that I have examined, and I have not been able to find
the types in the Paris Museum, where, however, Dohrn
found them and redescribed them.
Now Dohrn’s description is important, for it gives
several features which preclude that species which is
generally labelled B. chilensis in collections, but comes
from Brazil, and not from Chile.
It is a pity that Dohrn does not refer to the size, nor
form of the eyes, nor to the presence or absence of a keel
on the mesonotum.* These are, of course, essential fea¬
tures, and as B. chilensis is the type-species of the group,
it is most desirable tliat the doubt be removed as soon as
possible.
The Brazilian species commonly known as B. chilensis,
being distinct therefrom, requires a new name, for which I
propose coriacea, as some specimens are thus named in
MS. in the Brunner collection.
The points in which B. cormcca, sp. n., differs from
B. chilensis, according to Dohrn’s description, are as
follows : —
B. chilensis, Blanch.
(From Chile ; based on Dohrn’s
description.)
Pronotiim so breit als lang.
Pronotuni in der Mitte quer
geteilt durch eine Sattelfoermige
Vertiefung.
Das 2te und 3te Segment des
Abdomens mit deutlicher Falte.
. . . die folgenden Segmente
beim ^ an der Seite winklig,
nach hinten in eine stumpfe
Spitze ausgezogen.
B. coriacea, sp. n.
(From Brazil.)
Pronotum about H times
longer than broad.
Pronotum not so divided ; no
transverse sulcus.
Tubercles feeble.
Sides of these segments in the
^ rounded.
* Blanchard’s description and figure are equally silent on these
essential points ; his figure is very sketchy, and probably untrust¬
worthy ; it gives small eyes and smooth mesonotum, but we must
not attach too much importance to it.
Lahiduridae, d family of the Derma^dcra. 193
Tlie above discrepancies are to my mind quite decisive,
and I liave no doubt that the Brazilian B. coriacea is quite
distinct from B. chilensis. But as it is generally known
under that name in collections, it is convenient to define
the typical genus Brachylabis by the main generic features
of B. coriacea, as I have done in the preceding table, i. e.
eyes small and normal, and mesonotum keeled; the an¬
tennae segments are cylindrical, not globular, the fourth
being a little shorter than the third.
It is quite possible, or even probable, that when the
true B. chilensis is re-examined, it will be found to be
generically different from B. coriacea. It will then be
necessary to re- characterize the genus, and to erect a new
one for B. coriacea.
B. coriaxca has all the characteristics of the group in
colour, appearance, texture and general structure ; the
antennae have 15-16 segments, and are paler at tlie apex;
the pronotum is about If times as long as broad, gently
widened posteriorly ; the mesonotum has a distinct keel
running its entire length ; the legs are yellowish brown ;
the forceps of the male are arcuate towards the tips, and
sometimes have a blunt tooth near the base.
I possess specimens from Espirito Santo, and in tlie
Bnmner collection there are specimens from Santa
Catherina and Novo Friburgo.
I may add that these are not common insects, and it is
not probable that any one species should be common to
Brazil and to Chile.
Brachylabis scotti, sp. n.
Statura minore, robustiori ; oculi parvi, anteriores; antennae
seginentis 10 elongatis, conicis ; mesonotum baud carinatum ; abdo¬
men dilatatum; forceps paullo arcuatus. $
Long, corporis . ... 6 mm.
„ forcipis . 1‘5
Hah. Seychelles.
I describe this species here briefly ; it will be more fully
discussed in a work upon the Dermaptcra of the Seychelles,
on material obtained by the Gardiner-Scott expedition.
We must provisionally place it in this otherwise Neo¬
tropical genus ; when more material is discovered, its
true relationship will doubtless be better defined.
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) O
104 ])r. Malcolm llurr’s J^reliiiiinarii Herisum of tho
Perhaps the creature described by Montroiisier, 18G4,
p, 272, under the name of Glididuvct gmiculata, from New
Caledonia, is to be referred here. De Bormans’ collection
contained a specimen, now in the British Museum, from
New Caledonia, which is decidedly a Brachylabid. The
antennae are missing, and the specimen is a female ; but
the eyes are normal, and the keels of the mesonotum are
very sharp and distinct. The pronotum is a little longer
than broad. It is described by de Bormans, 1000^, p. 54.
Genus 8. — Nannisolabis, Burr.
Nannisolahis, Burr, 1910b p. 106.
I erected this genus for two Cingalese species which
have normal, small, anterior eyes and globular antennal
segments. The mesonotum has in its anterior portion a
transverse depression which is bounded by a raised tumid
ridge. Though somewhat compressed at the shoulders,
this ridge is not sharp.*
In the globular antennal segments and non-keeled
mesonotum it approaches Le^ytisolahis, but differs in the
normal eyes.
The type of the genus is Nannisolahis loilleyi, Burr, the
other species being N. idiiktas. Burr, 1901, p. 322, PI. VIII,
fig. 7. The figure shows the peculiarities referred to.
Unfortunately, of the original pair of this latter species,
owing to an accident, all that is left is the head and thorax
of the male, but this, with the original description and
figure, is enough to allow its position being satisfactorily
determined.
I place here also, provisionally, a very distinct new
species from Australia.
TABLE OF SPECIES.
1. Pronotum longius quam latius ; species
orientales.
2. Pronotum parallelum, punctatum ... 1. philefas, Burr.
2.2. Pronotum postice ampliatum, laeve . . 2. ivilleyi, Burr.
1.1. Pronotum transversum ; species australica 3. hohlansi, sy. n.
* Perhaps some of tliese peculiarities are due to immaturity of the
specimens.
Lahitlvriilac, a familij of fhe Derma'ptera .
195
Nannisolabis hohlausi, sp. ii.
Long, corporis
„ forcipis
13 Him.
•2
Of rather stout build, for this genus large ; colour deep reddish-
lu’own or blackish, scarcely punctate.
Head broad, sutures distinct ; occii)ut rectangular ; eyes anterior
and relatively small.
Antennae with 16 segments, two or three praeapical segments
yellowish ; 3rd segment long and cylindrical ; 4th ovate, about half
as long as the 3rd ; the rest gradually lengthening, the apical segments
about equalling the 3rd.
Pronotum distinctly broader than long, rectangular.
Mesonotum a little shorter and a little broader than the pronotum,
with no keel.
Legs long and slender, typical of the sub-family, yellow, the femora
and tibiae banded with black.
Abdomen somewhat dilated about the middle, tapering apically,
typical.
Penultimate ventral segment narrowly rounded.
Forceps subcontiguou.s, rounded, rapidly tapering.
Australia : Queensland, Cooktown (coll. Brunner,
No. 20, 162, 1 ? ).
Although this specimen is a female, and no male is
known, the characters are so Avell marked that the species
can be easily recognized, apart from the fact that it is the
only known Australian member of the sub-family, except
the doubtful B. geniculata.
I place it in Nannisolabis on account of its small eyes
and smooth pronotum, but in its broad head and transverse
pronotum it differs from the other two known species, as
also in the long antennal segments; a new genus must be
erected for it when the male is discovered. This new genus
will stand in the same relation to Nannisolabis that Leptiso-
labis stands to Isolabis and Cte7iisolabis to Mdisolahis.
The type is in the Brunner collection, now in the Hof-
mnseum in Vienna.
(.) 2
196 Br. Malcolm r)urr’s Preliminary Revision of the
Genus 4. — IsoLABis, Verhoeff.
Isolahis, Verb., oj). cit., p. 14.
Jsolabis, Burr (op. cit.).
This genus is monotypic for J. hraneri, Verb., from
Africa. It is somewhat larger than its allies.
Genus .5. — Metisolabls, Bun-.
Metisolabis, Burr, 1910^, p. 108.
Antennae segmentis longis, omnibus longioribus quain latioribns ;
mesonotum lateribus carina, interdam obtusa, instructum ; ceteris
cum Ctenisolabide congruet.
I formed this genus for those species whicli liave the
antennal segments elongate, cylindrical, always longer than
broad, even the fourth. Thus it approaches Isolabis,
V'^erli., to which it stands in the same relation as Ctenisola-
bis to Lcptisolabis^
The type of the genus is Prrachylabis voeJtzhoun, Burr,
from Nossi-be, though that species has the keels of the
mesonotum decidedly blunt and rounded, thus approaching
holabis.
I place Br. hifoveolcUa here because Bolivar figures it
witli clearly elongate antennal segments and well-marked
keels.
TABLE OF SPECIES.
1 . Pronotum longius quam latius ;
species madecassae.
2. Mesonotum carinis obtusis, usque
ad marginem posticum productis 1. •mabjacha, Burr.
2.2. Mesonotum carinis acutis, ante
marginem posticum evanescen-
tibus . 2. voeltzkoivi, Burr.
1. Pronotum vix longius quam latius y
species orientales.
2. Pedes unicolores ; abdomen seg-
mento 4 plicifero . .3. bifoveolata, Bob
2.2. Pedes fulvo-annulati ; abtlomen
segmentis 3 et 4 plicit'eris . . 4. caudelli, Burr.
Lcibiduridae, a family of the De,rma])Ura, 197
Genus 6. — Ctenisolabis, Verb.
This genus is now restricted to those species with
short and almost globular antennal segments and keeled
mesoDotum, with large, lateral eyes.
The type of the genus is Ctenisolabis toejoensis, Verb.
TABLE OF SPECIES.
1. Carinae mesonoti acutae, usque ad
inarginem posticuin percurrentes.
2. Frous punctis impressis postice cou-
fluentibus ; species africana . . 1. togoensis, Yerh.
2.2. Frous punctis impressis baud
couHuentibus ; species americae
meridionalis . 2. nigra, Scudd.
1.1. Carinae mesonoti obtusae, antice
marginem posticuin evanescentes.
2. Corpus valde pilosum ; species
orientalis . 3. fetcheri. Burr.
2.2. Corpus fere glabrum ; species
americanae.
3. Pronotuin vix longius quam
latius ; statura min ore (8 mm.),
robustiori . 4. inontaua, Bor.
3.3. Pronotuin distincte longius
quam latius; stature paullo
majore (11'5 mm.), graci-
liori . 5. fernandezi, Bor.
Ct. torjoensis, Verb., is only known to me from Verhoeff’s
description and a sketch of the type by Herrn Flandezky
of Berlin. It appears to resemble its allies very closely,
but is at present the only known African member of the
genus.
Ct. nigra, Scudd., appears to be relatively common in
Brazil, and extends into Paraguay. It was originally
described by Scudder as a Cylindrogaster , with which
genus, of course, it has no affinities. But in those
days this group was exceedingly imperfectly known, and
Scudder only had a single female.
Ct. fietcheri, Burr, is the only known Oriental species.
Ct. montana, Bor. (1909^, p. 5), from Costa Rica, closely
resembles Ct. nigra and Ct. fietcheri in appearance. It is
198 Dr. Malcolm'^iurr’s Prc/iiiilnar// Jievision of the
well described, but not tigured, by Borelli, It must be
carefully distingiiisbed from Ct. ni(jra. The antennal
segments are rather longer than the other species, thus
affording the transition to Mctisolahis.
Ct. fernandezi, Bor. (1909^, p. 4, fig. 2), is well described
and figured by Boielli. It occurs in Costa Rica. The
unique type has white eyes, which are very prominent, but
this colour is probably not a permanent feature. The body
is longer anti build slenderer than in the other species, and
the pronotum is much more finely punctulafe than in Ct.
montana.
All the species of Ctenisolahis have a strong family like¬
ness, and though when placed side by side tliey appear to
be quite distinct, it is difficult to express the difference in
words. As they are apterous, and rare forms, probably
each with a restricted distribution, the locality becomes an
important specific character.
The occurrence of such similar species in such widely
separated localities is probably an archaic feature, point¬
ing to the great age of a group, now dying out, and
which must formerly have been a dominant and widely
distributed type of earwig.
Perhaps more than one species is confused under
the name Ct. nigra, but I cannot distinguish between
Paraguayan and Brazilian specimens.
Genus 7. — Leptisolabis, Verb.
Le/disoUd/is, Verb., op. cit., p. 12.
JJrachykihis {partim), Borm.
This genus is characterised by the lateral eyes, non-
keeled mesonotum and globular 4th and 5th antennal
seofinents.
The type is L. usamharanu, Verb., from Usambara
(Africa). It includes also the true Br. punctata, of Dubrony
from Java, which is distinct from the species quoted by
him under that name from Burmah, which is Metisolahis
caudelli, Burr.
One of the chief features of the two African species
described by Verhoeff, L. usamharana and L. thcoriac., is
that the anterior border of the pronotum is produced into
Lahichiridae, a family of the Deomiaptera. 199
a naiTOw neck. This is not the case in L. 'punctata, but is in
L. howardi, Burr, from Guatemala.
As the two African forms are unknown to me in the
flesh, I cannot erect a satisfactory table of species, but it is
unlikely that a Central American insect can be confused
with African forms.
By good fortune I have been able to examine the
original insect quoted by Caudell from Guatemala as
Brachylabis nigra. Owing to the distance between Guate¬
mala and Para, I was not surprised to find on examination
that Caudell’s specimen is distinct. It is a true Leptiso-
lahis, as will appear from the following description.
Leptisolahis hoioardi, Burr.
Brachylabis nigra, Caudell, 1907^, p. 172.
Leptisolabis howardi, Burr, 1910, Proc. U.S. Nat. Mus.,
vol. 38, p. 449.
Statura minore, gracili ; colore atro, antennis pedibusque palles-
centibus ; corpus totum confertim punctulatum ; oculi magni, mar-
ginem posticum capitis fere attingentes ; prouotum elongatum et
postice anipliatum, margine antico in collem producto ; forcipis
bracchia cylindrica, basi baud coutigua, sensim arcuata.
Long, corporis . 8 mni.
„ forcipis . I’To „
Size small and slender.
Colour jet black, dull. The whole boily clothed with long golden
pubescence. The whole surface finely and densely punctulate.
Antennae with 1.3 segments, brownish grey, all the segments very
thick, 3rd scarcely longer than broad, 4th really broader than long ;
5th globular, the rest gradually lengthening and also thickening.
Head tumid, hinder margin straight, sides convex, and in front
triangular. The small marks on the frons indistinct. The punctu-
lations are exceedingly fine.
The eyes are very large and prominent, ovate in shajje, and extend
from the insertion of the elytra almost in the posterior margin of
the head, gently converging posteriorly.
Pronotuin about H times as long as the average breadth, all
margins straight, sides gently diverging as the pronotuin widens
somewhat posteriorly. The anterior margin has the middle portion
produced slightly, and carried a short but distinct cylindrical neck,
so that the head is distinctly separated from the pronotuin.
200 Dr, Malcolm Durr’s rrdiminary Revision of the
IMesonotum transverse, posterior margin truncate ; sides not keeled,
but raised with blunt, tumid ridges converging posteriorly.
Metanotum strongly concave posteriorly, almost entirely covering
the first abdominal segment.
The whole of the thoracic plates are dull black and densely and
finely punctulate.
Prosternum is double as long as wide, jiarallel.
Mesosternum and metastern uni somewhat broader than long, the
posterior margin truncate ; sternal plates reddish black and densely
and finely punctulate.
Legs typical, femora slender, black, yellow at the base and apex,
long and slender.
Tibiae long and slender, yellowish.
Tarsi yellowish, long and slender.
The second segment elongate and cylindrical, nearly as long as the
third ; the first is longer than the second and third combined.
Abdomen typical, of same colour and sculpture as the rest of the
body, passing to reddish beneath ; lateral tubercles on 3rd and 4th
segments faint.
Last dorsal segment very short and transverse, but narrower than
the abdomen ; posterior margin naiTowed, and concave in the
middle, leaving a short triangular lobe in each side projecting
slightly over the forceps.
Penultimate ventral segment short and broad, very obtusely
rounded.
Pygidium short, very narrow and compressed.
Forceps with the branches remote at the base, cylindrical, tapering,
straight at first, gently arcuate towards the apex.
Guatemala : Provincia de Alta Vera Paz, Trece Aguas,
Cacao, April 19 (Leg. Schwarz and Barbar) (in
U,S. N. M. vide Caudell, loc. cit.).
This species has a superficial resemblance to M. nigra.
It differs however in the generic characters, namely, the
non-keeled mesonotum; the 4th-Gth antennal segments
are much shorter, the pronotum longer, and the neck
distinct.
It is more difficult to distinguish from L. punctata,
Dubr., from Java, but is smaller, slenderer, and the
sculpture finer, the antennal segments shorter.
It is not quite so small as N.'philetas, Burr, from Ceylon,
the pronotum is much longer and not parallel, and the
mesonotum has no strong transverse depression.
I cannot compare it with Verhoeff’s two African species,
Lahiduridae, a family of the Dcrmaptera. 201
L. usambarana and L. theoriae, as these are only known to
me from the description.
As these creatures are totally apterous, it is very prob¬
able that the various records of “ Brachylabis nigra ” from
such various localities as Peru, Bolivia and Brazil refer to
distinct species.
It is dedicated to Dr. L. Howard.
Literature.
The following works are quoted in the preceding pages.
The enumeration is that of my complete bibliography, at
present in MS., from which this list is extracted for the sake
of uniformity.
Blanchard, E. (1851.) Orthoptera in: Gay, Historia fisica y
politica de Chile segun Documentos adquiridos . . . y pub-
licada bajo los auspicios del supremo Gobierno. (Zoologia,
vol. vi, and Atlas, Paris, 1851.)
Borelli, Dr. Alfredo. (1907^.) Spedizione al Ruwenzori di
S. A. R. Luigi Ainadeo di Savoia, Duca degli Abruzzi, xxvii,
Nuova specie di Forficola. (Terza note preventiva.) (Boll.
Mus. Tor., xxii, No. 572, 1907.)
- (1909^.) Forficola nuove o poco note di Costa Rica. (Boll.
Mus. Tor., xxiv, No. 611, 1909.)
Borg, Hjalmar. (1904.) Forficuliden aus Karaerun ; Beitriige
zur Kenntniss der Insektenfauna von Kainerun. (Arkiv for
Zoologi, Band 1, pp. 563-580, Tafel 26, 1904.)
Bormans, A. De. (1883.) Etude sur quelques Forficulaires
nouveaux on peu connues, precede d’un tableau synoptique
des genres de cette famille. (Ann. Soc. Ent. Belg., pp. 59-90,
pis. 2 et 3, 1883.)
- (1893.) Dermaptera : in Biologia Centrali-Americana. (Zoo¬
logia, Orthoptera, 1, pp. 1-12, pis. 1 and 2, 1893.)
- (1900h) Quelques Dermapt^res du Musee civique de Genes.
(Ann. Mus. Civ. Gen., (2) xx. pp. 441-467, 1900.)
- (1900-.) Forficulidae in “Das Tierreicli,” 11 Lief. Orthoptera,
pp. 1-142, 1900.
Brunner, V. Wattenwyl. (1895.) Orthoptera of the Sandwich
Igles. Proc. Z. S. Loud. 1895.
;’()2 Dr. Malcoli^lJurr’s Preliinmary Revidon of the
Bukh, Malcolm. (1897'*.) On new Species of Forficuloria. (Ann.
Mag. N. H., (fi) XX, pp. 310-.316, 1897.)
- - (1900'*.) Forficules exoti<iues du Mnse'e royal d’liistoire natur-
elle de Bruxelles. (An. Soc. Ent. Belg., xliv, pj). 47-0-1,
1900.)
- (1901.) The Earwigs of Ceylon. (Bombay Nat. Hist. Soc.,
xiv, pp. .59-78 and 316-33(5, pis. A and B. 1901.)
- (1902.) On the Forficularia of the Hungarian National
Museum of Budapest. (Termesz. Fiizetek, xxx, pp. 447-
489, pi. 20, 1902.)
- (1904*^.) Observations on the Dermaptera, including Revisions
of several genera and descriptions of New Species. (Tr. Ent.
Soc. London, pp. 277-322, 1904.)
- (1905^.) Notes on the Forficularia, ix. On new species with
synonymic Notes. (Ann. Mag. N. H., (7) xvi, pp. 486-496,
1905.)
- (loos'*.) Notes on the Forficularia, xiii — A revision of the
Brachylnbidae (Isulabichte). (Ann. Mag. N. H., (8) ii, p. 246,
1908.)
- (1908**.) Die Fauna Siidwest-Australiens. Ergebnlsse der
Hamburger siidwest-australischen Forschungsreise, 1905,
herausegegeben von Prof. W. Michaelsen und Dr. R. Haid-
meyer. Band II, Lief. 5 ; .lena, 1908. (Dermaptera, ])p. 69-80.
Taf. xi, 1908.)
- (1908*2.) Sopra alcuni Dermatteri del Museo Civico di
Genova. (Bull. Soc. Ent. Ital., ix, 1!>08, pp. 175-191.)
- (1909*.) Notes on the Forficularia, xv — The Esphalmeidnae.
(Ann. .Mag. N. H., (8) iii, pp. 249-253, 1909.)
- (1909'*.) Note on the Classification of the Dermaptera. (Deutsch.
ent. Zeitschr., pp. 320-328, pi. iv, 1909.)
- (1909**.) Notes on the Forficularia, xvii — On new species, a
new genus, and new synonomy. (Ann. Mag., N. H. (8) iv,
August 1909.)
- (1910*.) The Fauna of British India. Dermaptera (Earwigs),
London, 1910.
Caudell, Andrew Nelson. (19072.) On some Earwigs (Farjicn-
lidae) collected in Guatemala by Messrs. Schwarz and Barber.
(Proc. U. S. Nat. Mus., xxxiii, pp. 169-176, 1907.)
Dohkn, H. (1864.) Versuch einer Monographie der Dermapteren.
(Stett. Ent. Zeit., xxv, pp. 285 and 417.)
Kirby, W. F. (1903.) Notes on Forficulldae with descriptions of
new species in the Collection of tlie Natural History Museum,
South Kensington. (Ann. Mag. N. IL, (7) xi, 1903.)
Montruuzier. (1864.) Essai sur la faune entomologi(jue de
■TW. V
> A
■ ir.f .
" ^ ••>-
Trans^'jit. Soc. Lond., ipio, Plate XLVI.
E. Wilson del.
C. Hentschel.
DETAILS OF LABIDURIDAE.
Explanation of Plate XL VI.
Fig. 1. Allostethus indicum, Hag., sternum x
2. Echinosoma parvulum, Dohrn, sternum x
3. Gonolabis electa, Burr, sternum x
4. Pyragra dohrni, Scudd., sternum x
5. „ ,, „ tarsus x
6. Gonolabis oblita. Burr, apex of abdomen, ventral
view, ^ X
7. Euborellia ambigua, Bor., thorax x
8. Euborellia greeni, Burr, thorax x
9. Euborellia armata, Bor., thorax x
10. Palex sparattoides, Bonn., sternum x
11. Gonolabis suniatrana, Bonn., apex of abdomen,
ventral view, x
12. Gonolabis oblita, Burr, ^ x
13. Anisolabis maritima, Bor., sternum x
14. Gonolabis javana, Bonn., apex of abdomen, ventral
view, ^ X
6.
11.
10.
4.
10.
10.
6.
5.
4.
6.
10.
6.
5.
10.
Tr^s. Ent. Soc. Land., jpio, Plate XLVIL
E. Wilson del.
C. Hentschel.
DETAILS OF LABIDURIDAE.
Explanation of Plate XLVII.
Jjahidiiridac, a family of the Dermaptcra. 208
Kaiiada (Noixvelle Caledoiiie), et descriptions des qiielques
especes noiivelles ou peu connues. (Ann. Soc. Linn. Lyon, xi,
p. 222.)
Perkins, E. C. L. (1899.) Fauna Hawaiiensis, vol. ii, part i,
Orthoptera, Cambx’idge.
Rehn, James, A. G. (1903") Studies in American Forticulidae.
(Proc. Nat. Sci. Phil., p 299, 1903.)
ScuDDER, S. H. (1890.) The Fossil Insects of North America.
II, The Tertiary Insects. 1890, New York.
VERHOE.'fF, Karl W. (1901.) Ueber Dermapteren, II. (Sitz, Berlin,
Der Ges. Naturf. Freunde zu Berlin, No. 1, p. 7.)
- (1902) Ueber Dermapteren, I. (Zool. Auz. No. 665.)
■ - (1904) Ueber Dermapteren, V. (Arch. f. Naturg., 1904, p. 115.)
Explanation of Plates XLVI, XLVII.
[(See Explanation far.imj the Plates.]
( ^04 )
VIII. On the Qeo\niii\'\(\ix(i of the Arncntine liepiihlic. By
Louis B. Prout, F.E.S.
[Road April 6tli, 1910.J
Plate XLVIII.
Although a good foundation has been laid by Bunneister
and Berg for the study of certain Lepidopterous families
in the Argentine Bepublic, the Geometridae have hitherto
been greatly neglected. Burmeister’s work, “ Description
physique de la Republicpie Argentine ” (Paris and Buenos
Aires, 1876-80, 8vo), contained “ Lepidoptera, Part I,”
but this did not reach to the Geometridae, although a few
genera which were formerly classed as Lithosiid foimd
their way into it. Almost all that is known of the
Argentine representatives of the family in question is
therefore to be sought in scattered articles in periodicals,
such as those published in Buenos Aires, where Berg has
described a very few species, Rothschild’s “ Novitates
Zoologicae,” containing Warren’s descriptions of the new
species in the Tring Museum, and the “ Annales de la
Societe Entomologique de Belgique,” containing Dognin’s
of those in his own collection. By gathering these to¬
gether, and at the same time making known the numerous
new species which I have received through the generosity
of j\Ir. A. F. Bayne, F.E.S., of Buenos Aires, his brother,
Mr. W. M. Bayne, of Mendoza, and Dr. Eugenio Giacomelli,
of La Rioja, I hope to bring our knowledge of the
Geometi'idae of Argentina at least up to the level of that
of some of the other, better-worked families. In order to
make this memoir as complete as circumstances will allow,
I am including in it a large number of faunistic records
which have been courteously supplied to me by M. Paul
Dognin, the well-known French student of South American
Geometridae, to whom I am further indebted for his un¬
wearying kindness and co-operation in the quest of the
solution of knotty points. Further records have been
gathered from material kindly sent for my inspection by
Dr. M. Bastelberger, of Wurzburg, as well as from that
contained in the British IMuseum collection. It is much
TRANS, ENT. SOC. LOND. 1010. — PART III. (NOV.)
Mr. L. B. Pront on Argentine Geometridae. 205
to be regretted that the vast Tring collection of Geometridae
is not yet arranged in a manner which will allow of its
systematic study ; at the same time, I have to acknow¬
ledge with gratitude that every facility possible has been
granted me by the Hon, Walter Rothschild and Dr. Karl
Jordan, while Mr. W. Warren has been ever ready with
his assistance and advice.
Regarding the collection of unnamed moths referred
to by Dr. H. Weyenbergh, on pp. 181-2, of Napp’s “Die
Argentinische Republik ” (Buenos Aires, 187G, 8vo), Mr.
P. C. T, Snellen wrote me (in litt., February 9, 1907) as
follows: “Dr. Weyenbergh indeed handed to me, after
his return to Europe, a small collection of Heterocera.
Unfortunately, however, that collection was, shortly after,
destroyed by accident, with the exception of a few
Fyralidae that were taken out before. Otherwise I
Avould, according to the wishes of my deceased friend,
have worked it out long ago.” Weyenbergh {toe. cit.,
p. 181) commits himself to nothing more definite than
the following : “ The Geometridae appear to be less
numerous, at any rate thus far I know at most fifty
species. . . . The species I have met with represent,
as I believe, the following or related g-enera : Ennomos,
Boarmia, Gnerphos, Zerene ?, Hibernia, Cidetria, Acid alia
and Eninthecia. This group has as 3^et had little
attention.”
Burmeister’s “ Reise durch die La Plata-Staaten ”
(2 Bde., Halle, 18G1, 8vo), though containing much valu¬
able information on the natural history of the country,
and a good many records of Lepidoptera, does not include
any Geometridae. E. W. White, F Z.S., in his “ Cameos
from the Silver Land ” (2 vols., London, 1881-82, 8vo),
only touches the Lepidoptera in one or two places, and
has nothing on Geometridae. The same applies to W. H,
Hudson’s “The Naturalist in La Plata” (ed. 4, London,
1903, 8vo).
I have thought it best, with one trifling exception, to
accept the present political boundaries of the Ai^entine
Republic as the basis of my work. To include the few
scattered records from Argentine Patagonia has not added
materially to the size of my paper; and as the only
systematic surveys which touch Patagonia (Mabille in
“ Mission Scientihque du Cap Horn ” and Staudinger in
“Ergebnisse der Hamburger Magalhaensischen Sammel-
200
Lou is P), Front on the
reise”) confine themselves to the extreme south, it seemed
desirable to give myself the widest scope which my title
warranted. The single exception, to which I have alluded,
consists therefore in the exclusion of the part of Tierra
del Fuego which belongs to Argentina; this could not
have been logically included without the rest of the
island, and in any case I should have had scarcely
anything to add to the work of Staudinger.
The localities where Mr. W. M. Bayne has collected are
of the utmost interest, having been previously almost
untouched by entomologists. His collecting followed the
route of the Argentine Great Western Railway from Villa
Mercedes to Las Cuevas, i.e. right away into the Andes
as far as the Chilian frontier; lie has also taken a few
species at stations on some of the branch lines. Although
some of the geographical names quoted in the following
pages may probably be unfamiliar to some readers, I have
not thought it necessary to indicate their location, as they
are stations on the railway. It may, however, be of some
interest to quote a few of the principal altitudes from the
official prospectus of the railway : — -
Jletres.
Villa Mercedes . .513
San Luis . 719
La Paz . 500
San Rafael . 685
Mendoza . 753
San Juan . 633
Cacheuta . 1245
Puente del Inca . 2720
Las Cuevas . 3151
Puente del Inca is referred to by Mr. H. J. Elwes, in
his valuable paper on “ The Butterflies of Chile ” {vide
Tr. Ent, Soc. Bond., 1903, pp. 264-5), as a promising i
locality for the entomologist, and Mr. Bayne’s experience ■
has borne this out ; nearly all the species which he has ;
there obtained seem to be new to science, though natur- (
ally with some relation to the mountain fauna of Chili,
and, to some extent, to that of the highlands of Peru and
Bolivia.
In the fauna of so vast and varied a country as Argen¬
tina there is, as might be expected, great diversity ; and
although our knowledge of it, as regards the moths, is
Geometridae of the Aryenline Ik])')d>lic. 207
still almost in its infancy, I may venture on a few
general remarks. From the extreme north I have no
records at all, but M. Dognin has received interesting
collections from Metan, while the neighbourhood of
Tucuman has been a good deal worked entomologically.
In these localities there is a considerable element of the
subtropical fauna, though even here we realise that we
are in a different region from the strictly equatorial, the
gorgeous Braccinae” of Warren (Siosta, Sangalopsis, etc.)
and many other tropical genera {Ccdlipia, most “ Pahja-
dinaef etc.) being apparently unrepresented, others again
(e. g. ScordyUa) quite weakly represented. On the other
hand, there is hardly a trace (perhaps not a trace) of
the distinctively southern, or Chilian fauna; Tephrinopsis
siibumhrata, Dogn., may possibly be an exception from
this generalisation. From the north-east the British
Museum has obtained some small but valuable collections
made about Florenzia, Ocampo and Goya ; excepting,
perhaps, Atyria chihcha (Schaus), I have not noted any¬
thing characteristically tropical in these, the affinities of
the majority being distinctly with the Geometridae of
S. Brazil and of Paraguay. When we reach Buenos
Aires, we find nearly the same type of forms still prevail¬
ing; Mr. Druce has recorded Atyria durnfordi (Druce)
from this locality, and Mr. A. F. Bayne has once taken
Phrygionis p)aradoxata incolorata, but I am inclined to
view these as stragglers. Mr. Bayne thinks the Rio de
la Plata forms somewhat more of a natural boundary
than the geographical distance would have led one to
anticipate ; at any rate he has seen at Montevideo and
Tacuarembo several species, and even genera {Aplodes,
Melinoides, Bronchelia, etc.), which have not yet occurred on
the Argentine side. Of the fauna of the pampas country
(which will certainly be but meagre) practically nothing
is known ; from its frontal armature one might hazard
the conjecture that Hoplolygris cicatriculata (Berg) had
its origin here, but it has spread from Tucuman and Goya
in the north to the Rio Negro in the south.
The very few specimens which I have seen from
San Luis, however {Arizela didcis, Prout, Narragodes
gyda, Prout, etc.), seem to indicate the commencement
of the western, or Mendoza fauna. This fauna, as
shown by my material from Balde, La Paz, and even to
Mendoza city, and I think northward {e.g. Retamito),
208
Mi^jouis B. Front on the
is cliaracterised by the predominance of small, obscure
greyish species, particularly of the group of genera
Mimophyh, Narragodes, Psodopsis, Idialcis, though some
totally unrelated species, such as Litkostege pax, Front,
appear superficially to be in quite suitable company among
tliem. The insects from the high Andes are of quite a
different type, the very glossy scaling being a marked
feature, irrespective of generic relationship. Phopcilodes
otophora, Front, the two species of Chrismop)teryx, Teph-
rinop)sis suhscripta, Front, Salpis carneitincta, Front, S. ruheiifi,
Front, and Caiascia carnca, Front, are perhaps the chief
examples; in several of them a more or less pronouncedly
fleshy tone of colouring is observable. A few of these
Andean species, or at least their genera, are already
known to have some northward range, and no doubt
further material (for example, from the Bolivian Andes)
would more extensively exemplify these resemblances. I
may mention that Perizoma sordesmis, Dogn., was described
from Fern, Leucocliesias mcsargyrata, Mab. (under its
synonym of Aspilates niveipmnaria, from Bolivia;
while Salpis carneitincta, Front, has a very close Feruvian
relative in S. pucchi, Dogu., and I have seen in M.
Dognin’s collection a specimen, also from Feru, of a species
apparently indistinguishable from Gnophos (?) oreas.
Front.* I have already mentioned a Chilian element in
this western fauna ; the genus Salpis is best known in
Chili, Xanthorhoii edmondsii (Butl.) was originally described
from Chili, and my new genus Baynia certainly belongs
near some known Chilian genera. As we proceed further
southwards, this characteristic becomes still more marked ;
Berg’s Rio Santa Cruz species {Synneuria uniforrnata,
Odontothera (?) crucifer aria), Mabille’s Pachropliylla oculata,
from Aysen River, and the whole of the Magellan records
point strongly in this direction.
On commencing my work, I contemplated including
tabulations for the determination of genera, in the hope of
stimulating local research, as has been so admirably done
by Meyrick and Turner in their Australian revisions, or
by several others in faunistic works of the best type ; but
I found tliat a far-reaching study such as could have
rendered this effective (especially in the case of the chaotic
Boariniinae) would have delayed publication for an inde-
* Peru is also tlie habitat of the two nearest allies of this species,
rhanchani, Pogn., and insulHa, Warr.
Gcometridae of the Argentine Repuhlic. 209
finitely long period, or turned me aside from revisional
work in other sub-femilies, which is already claiming all
my available time ; I was therefore compelled reluctantly
to abandon the idea, and many of the generic names are
merely given according to the current usage of the few
workers at the Neotropical fauna. I have, however, given
much attention to the structural characters, particularly
in cases of doubtful generic location ; and where I have
transferred species from one genus to another, I have
generally assigned adequate reasons for so doing.
In almost every instance, even with the common species,
I have given information as to the sources of my records
and the collections in which specimens are to be found.
This will, I trust, be of assistance in the future elimination
of errors of determination, etc., of which a few must of
necessity creep into a pioneer work of this kind. The
types of all the new species are in my own collection or in
that of the British Museum, and I shall lose no available
opportunity of placing co-types, or reliably determined
specimens in the latter repository. Mr. A. F. Bayne has
been most generous in withholding nothing which I asked
of him, either in material or in notes, and without his aid,
and the labours of his brother in collecting and breeding,
this work would have been deprived of nine-tenths of its
value.
The measurements of wing-expanse in the descriptions
of the new species are the theoretical extremes (the added
length of the two forewings lAus width of thorax), not the
superficial expanse from tip to tip in set specimens.
S\.\h4a,m{\y—OENOCHROMINAE.
Oenochroininae, Ortliostixinae et Meeoceratinae (Warren).
1. “Almodes stellidaria (Guenee).”
? Polysemia stellidaria, Guenee, Spec. Gen. des Lep., x,
450 (1858).
? Boarmia (Deileptenia) sqnamigera, Felder, Reise Novara,
Lep. Het., tab. cxxvi, 11 (1875).
Tucuman, in coll. Dognin.
The determination of Guenee’s Polysemia stellidaria is
perliaps somewhat problematical, though it was certainly
a true Almodes. I did not study in M. Dognin’s collection
the species which he (or Mr. Warren) has thus identified,
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) P
210
MIFLouis B. Piout on the
and merely give the record on liis authority. Guenee’.s
type Avas from Haiti.
2. Almodes(?) benesignata (Dognin).
Almodes hcnesignata, Dognin, Ann. Soc. Ent. Belg., 1, 106
(1906).
Metan, Salta (type), in coll. Dognin.
An interesting species, in some measure intermediate
between Almodes and Ergavia ( = Polysemia, auctt.). The
typical Almodes forms seem to belong entirely to Mexico,
Central America and the West Indies, whereas Ergavia
ranges certainly from Mexico to Paraguay.
^\ih4a.m\\y—HEMITHEINAE.
Hemitheidac, Bruand ; Geometrinae, auctt.
3. Gel ASM A munda (Warren).
Gelasma munda, Warren, Nov. Zool., iv, 425 (1897).^
1 Gelasma stigmatica, Warren, Nov. Zool., xi, 20 (l904)-
(var. ?).
(^) La Plata City (type), in coll. Rothschild.
Apparently a Avidely-distributed species in South
America. The British Museum collection has it from
Panama and Venezuela, practically typical, and I am
strongly inclined to suspect that G. stigmatica, Warr.,
from Peru, is a large, strongly-marked aberration, or at
most a local race. I have the typical form from Rio.
4. Aplodes frondaria (Guenee).
Synchlora frondaria, Guenee, Spec. Gen. des Lep., ix, 875
(1858).
? Thalera minuata, Walker, List. Lep. Ins., xxxv, 1613
(1866).
Goya, Corientes, in coll. Br. Mus. ; ? Tucuinan and Los
Vasquez, in coll. Dognin.
Also widely distributed, from Vera Cruz (Mexico) to
Uruguay, but chiefly on the Atlantic side of South
America, and rather frequent in the West Indies. The
only western specimens known to me are from Callao. It
shows no appreciable geographical variation.
Geometridac of the Argentine Re-pnblie. 211
I have little doubt that the insect recorded by M. Dognin
(in litt.) as Aiolodes minuata, Walk., is this species.
Walker’s type is apparently lost, his type label having
been attached to a North American specimen, belonging
to Synchlora aerata, Fab. ; Walker’s description and data
(Santa Marta, collected by Bouchard) sliow that this
cannot possibly be right.
Guenee is in error in stating, under his generic diagnosis
of Synchlora, that the antenna is merely thickened
anteriorly in the ^ . The accepted determination of his
frondaria is evidently correct, or even if his description
can really apply to some other species, it will certainly be
a close ally (e. g. Pachycoims tridentata, Warr., Nov. Zooi.,
iv, 428), while there is no allied species in which the $
antenna is not bipectinate. One must suppose that he
had two females of Aplodes frondeevia before him, and
mistook the sex of one.
5. Lissochlora sanguinipunctata (Dognin).
Lissochlora sanguinipunctata, Dognin, Ann. Soc. Ent. Belg.,
1, 204 (1906).
Tucuman (type), in coll. Dognin.
6. Miantonota erina (Dognin).
Achleyi'a erina, Dognin, Ann. Soc. Ent. Belg., xl, 143 (1896).
Miantonota erina, ab. di^uncta, Warren, Nov. Zook, xvi, 81
(1909).
Tucuman (type of ab. disjuncta), in coll. Rothschild.
The typical form of the species is distributed from Mexico
to Rio Janeiro, and I believe stands in coll. E. Dukin-
field Jones (from Casti’O, Parana) as Geometra apiciata
(Schaus, MS.).
7. Auophylla multiplagiata (Warren).
Auophylla multiplagiata, Warren, Nov. Zook, iv, 424
(1897).
Tucuman, in my collection and those of M. Dognin and
Dr. Bastelberger.
o
The species was described from Paraguay, and I know of
no further localities than these two.
212
Louis B. Prout on the
8. Racheospila sigillaria (Guenee).
Eacheospila sigillaria, Guenee, Spec. Gen. des L^p., ix, 375
(1858).
Racheospila tenuimargo, Warren, Nov. Zool., xii, 319
(1905) (nov. syn.).
Tucuman, in coll. Bastelberger.
Guenee’s type was from Montevideo. Northwards the
species reaches as far as the West Indies and Mexico.
Warren’s B. tenuimargo, from Brazil, is the same species
(sec. specim. typ.).
9. Anomphax gnoma (Butler).
Omphax gnoma, Butler, Tr. Ent. Soc. Bond., 1882,
p. 367.1
Anomphao: gnoma, Warren, Nov. Zool., xvi, 74 (1909).^
(‘^) Salta, in coll. Rothschild.
Previously only known from Chili (i) ; the series from
which Butler described was collected at Las Zorras by
Mr. T. Edmonds ; Izquierdo, An. Univ. Chile, liii, 815,
says it ranges from Aconcagua to the interior of Araucania.
The species is interesting as being the only member of the
sub-family yet discovered in Chili. It does not seem to
be very closely related to any known species, yet the
following short but interesting account of the early stages,
translated from Izquierdo, shows clearly the Heinitheine
characteristic.
Larva on [Duvarct] Sehmiis dependens DC., reache.s, when full-
grown, a length of 18 mm., thickness H mm. Uniform green, of
the same colour as the leaves of the food-plant ; dorsal area reddish
in some examples. Bears an extraordinary resemblance to a little
twig. When taken between the fingers it feels decidedly hard, more
so than any other larva of like size known to the writer. In the
medio-dorsal line there is a series of intersegmental dots of a dark
coffee-colour. The anterior extremity of the larva terminates in two
small sharp horns with reddish spots, which simulate to perfection
the little buds of a twig. These, horns are appendages of the head,
and this latter, in repose, is bent downwards and closed up so tightly
with the legs that it is impossible to distinguish them. Laterally
the body has a very fine, conspicuous yellow line, which terminates
in the horns of the head. Pupates among leaves without making a
cocoon. The pupa, green at first, becomes nearly white, and two or
three days before emergence, the wing-cases assume a dark green
213
Geometridae of the Argentine Be^iblic.
colour. Duration of pupal stage about 14 days (6 to 20 February).
Imago on the wing from November to March. (Izquierdo, An.
Univ. Chile, liii, 815, tab. iii. 12 [larva].)
10. Hydata spilosata (Warren).
Hydata spilosata, Warren, Nov. Zool., xiv, 204 (1907).
Ciudad de Tucuman (type), in coll. Rothschild.
Sub-family— A CIDALIINAE.
Stcrrhinae et Cyllopodinae pars (Warren).
11. Asellodes laternaria (Guenee).
Asellodes laternaria, Guenee, Spec. Gen. des Lep., ix, 425,
tab. xxi, 3 (1858).
Gran Chaco, near Florenzia, October 1902, in coll. Br.
Mus.
Giienee’s type was from Brazil. That author described
the genus among his Acidalidae, and I have no hesitation
in accepting this as its correct position. Schaus (Tr.
Amer. Ent. Soc., xxvii, 194), on account of the point of
origin of the second radial, would advocate removing this
and the following genus to the Heinitheinae, but the
peculiar abortions of the hindwing and the transparency
of the wings suggest much more affinity with the present
sub-family, while the presence of a double areole in the
forewing is a still stronger argument for maintaining
Guenee’s arrangement.
12. PSEUDASELLODES LACUNATA (Dognin).
Pseudasel lodes lacunata, Dognin, Ann. Soc. Ent. Belg
1, 109 (1906).
Metan, Salta (type), in coll. Dognin.
13. PsEUDASELLODES CASSIOPEIA (Bastelberger).
Pseudascllodes cassiopeia, Bastelberger, Ent. Zeit. Stuttg
xxii, 59 (1908).
Tucuman (type), in coll. Bastelberger.
214
Louis B. Prout on the
14. Semaeopus oenopodiata (Gueiioe).
Acidalia oenopodiata, Guenee, Spec. Gen. ties Lep., ix, 509
(1858).
Tuciiman and Metan (Salta), in coll. Dognin ; La Rioja,
])r. E. Giacomelli.
This species was described from Brazil, and certainly
occurs about Rio, whence I suspect a great part of
Guenee’s Brazilian material was obtained. The placing of
the species in Acidalia was an error; the double areole
of the forewing and the highly developed hindtibial tuft of
the ^ point clearly to its relation to Semaeopus, H.-S.,
of which it may be regarded as forming Section III — $
antenna dentate-fasciculate, not pectinate as in the type
and the section Gnemodcs, Guen. (VVarr. restr.)
15. Semaeopus rubripuncta (Dognin).
Cneniodes rubripimcta, Dognin, Ann. Soc. Ent. Belg., xlvi,
344 (1902).
Tucuman (type), in coll. Dognin.
16. Anisodes subcarnearia (Warren).
Anisodes subcarnearia, Warren, Nov. Zool., vii, 145 (1900).
Buenos Aires, 1 taken October 12, 1906, by H.
Wilkinson, in coll. Br. Mus. Belongs to the ab. grisect of
Warren, loc. cit.
Warren’s type of the species, as well as that of ab. grisea,
came from Sao Paolo (E. D. Jones). It is by no means
certain that grisea is not a separate species.
17. Anisodes stollaria (Schaus).
Anisodes stollao'ia, Schaus, Tr. Amer. Ent. Soc., xxvii, 189
(1901).
Anisodes subpallida, ab. figurata, Warren, Nov. Zool., xii,
320 (1905).
Tucuman, in coll. Dognin.
Widely distributed from Mexico to S.E. Brazil,
agree with Schaus and Dognin in treating it as specifically
distinct from A. subpallida.
Geometridae of the Argentine Bopuhlic.
215
18. Anisodes excavaria (Schaus).
Anisodes excavaria, Schaus, Tr. Amer. Eut. Soc., xxvii, 190
(1901).
Buenos Aires, one ^ taken by Mr. A. F. Bayne, Novem¬
ber 9, 1903, ill coll. L. B. Front.
Tlie species was described from S.E. Brazil. The iden¬
tification of my specimen seems certain, although the ^ is
not so characteristic as the Mr. Bayne has met with
no second specimen as yet.
19. Euephyua serrulata (Packard).
Euephyra semdata, Packard, Fifth Rep. Peabody Ac. Sci.,
73 (1873).
Ephjjra obscura, Druce, Biol. Centr.-Amer., Lep. Het., ii,
536, tab. xeix, 6 (1898) (nov. syn.).
Buenos Aires, not rare from December to February
(A. F. Bayne) ; Tucuman and Los Vasquez, in coll. Dognin ;
Mendoza (W. M. Bayne).
A species of extremely wide range, occurring from the
southern United States to Buenos Aires; distinctly vari¬
able, but not geographically, unless the West Indian
nnnaria, Walk., be a form of the same, which I hold as at
present too doubtful to justify sinking serrulata.
20. Pleuroprucha insulsaria (Guenee).
Acidalia insulsaria, Guenee, Spec. Gen. des Lep., ix, 469
(1858).
Acidalia asthenaria. Walker, List Lep. Ins., xxii, 737
(1861).
Acidalia imparata. Walker, List Lep. Ins., xxvi, 1598
(1862).
Acidalia persimilata, Grote, Proc. Ent. Soc. Philad., i, 347,
tab. iii, 5 (1863).
Depitedm instdaria, Hulst, Tr. Amer. Ent. Soc., xxiii, 300
(1896) ; Dyar s List N. Amer. Lep., 291 (1902) (ex
err. pro insulsaria).
Tucuman, in coll. Dognin ; Gran Chaco, near Florenzia,
in coll. Br. Mus. ; Buenos Aires, one ^ in garden, February
11, 1903 (A. F. Bayne), in coll. L. B. Prout.
A species of extraordinarily wide range in eastern
America. Mr. Bayne’s specimen represents the southern-
21G
MtT Louis B. Prout on the
most limit yet recorded ; northwards I am not able to
indicate so precisely its known range, but it is certainly
frequent in the eastern United States, and Packard
(“ Monograph,” p. 335) gives several localities in Massachu¬
setts, etc., and one or two even in Maine. I formerly
attempted to keep the more desh-coloured and the more
greenish forms specifically distinct as msMfearm, Guen., and
asthenaria. Walk., respectively, but I cannot find that the
colour difference is even locally constant, except that I
think all the southern ones belong to the green-tinted
form (asthenaoHa). M. Dognin has furnished me with two
records for Tucuman, one as Deptalia asthenaria. Walk.,
the other as Ptychopoda insulsaria, Guen., but I have no
further note on the material in his collection.
21. Dichromatopodia sigillata (Walker).
Pyrinia sigillata. Walker, List Lep. Ins., xxvi, 1491
(1862).
Tucuman and Los Vasquez, in coll. Dognin.
Walker’s type was from Santarem.
22. Dichromatopodia concomitans (Warren).
Dichromatopodia concomitans, Warren, Nov. ZooL, xiv, 207
(1907).
Tucuman, in coll. Rothschild (type) and in coll. Bastel-
berger.
23. Ptychopoda piperata (Warren) (?).
Ptychopoda piperata, Warren, Nov. ZooL, viii, 456 (1901).
Gran Chaco, near Florenzia, October 1902, in coll. Br.
Mus.
Warren’s types (^ and $) were from Sao Paolo, S.E,
Brazil. They were missing from the boxes of types \vhen
I studied these a few years ago ; but the presence of topo-
types in the British Museum collection and in that of
Mr. E. Dukinfield Jones has enabled me to examine it,
and to confirm Warren’s reference of it to the genus
Ptychopoda. I am not absolutely certain that the Gran
Chaco insect is specifically identical ; its postmedian line
on the forewing is somewhat incurved near the inner
margin.
Geometridae of the Argentine Re^uhlic.
217
24. Ptychopoda spernata (Walker).
Acidalia spernata, Walker, List Lep. Ins., xxii, 730 (1861)
(nec var. /I).
Acidalia hotydaria, Walker, List Lep. Ins., xxvi, 1599 (1862)
(nov. syn.).
Ptychopoda limitata, Warren, Nov. Zool, iv, 443 (1897)
(nov. syn.).
Anteois pyginaeata, Warren, Nov. Zool., viii, 453 (1901)
(nov. syn.).
Goya (Corrientes) and Los Vasquez (Tucuman), in coll.
Br. Mus. ; Tucuman, in coll. Dognin ; Buenos Aires, not
rare (A. F. Bayne).*
I do not think the synonymy of this species, as given
above, has heretofore been published, though I called Mr.
Warren’s attention to it a long time ago. Walker’s type,
from Venezuela, is in very poor condition, but the species
varies little, and is very easily recognizable. His type of
hotydaria, from the same collection (Dyson’s), is better,
and would help to remove uncertainties of determination,
if such existed.
This species is to be regarded, so far as present knowledge
extends, as a typical Ptychopoda, though the areole (which
is simple) is rather narrow, and in one specimen which 1
have examined, but not denuded, is almost certainly “ open ”
distally ; i.e. SC^ does not anastomose with, though closely
approaching, the stalk of the other four subcostals. This
peculiarity occasionally occurs in the North American
species genimata. Pack, (an anomalous species, but nearer
to Ptychopoda than to Acidalia = Leptomeris, in which
Hulst places it), and deniissaria, Hiib. = ferrugcda. Pack. ;
see Packard’s “ Monograph,” pi. iii, fig. 16, and Explanation
of pi. iii, fig. 17 (“PJois ferruginata ”). With the last-named,
spernata is closely related — perhaps a geographical race
rather than a species.
25. Ptychopoda asceta (Prout), nov. sp.
9. 12 mm. Head and thorax bone-colour, slightly tinged with
reddish ; abdomen above deep purple fuscous, except anal end,
beneath pale. Forewing bone-colour, slightly tinged with reddi.sh,
* About Buenos Aires, Pt. spernata occurs from November to
February. It probably has a wide distribution in eastern South
America.
218
Mr. Louis B. Prout on the
the entire median area occupied by a rich, dark band, mingled with
ferruginous and bluish scales ; proximal margin of band from two-
lifths costa, angled below SC, thence slightly oblique basewards, but
making a slight curve distad in submedian area, reaching inner
margin at beyond two-fifths ; outer boundary of band (postmedian
line) from beyond two-thirds costa, forming a deep sinus distally to
cell, oblique basewards to submedian fold, thence again forming a
projection at SM^ reaching inner margin at beyond two-thirds ;
band shortly followed by a thick line parallel to its outer margin,
brownish tinted at costal end and not very distinct, overlaid with
slaty scales from E- onward, hence more conspicuous ; diffuse obscure
dark marks distally about R‘^ and submedian fold. Hind wing mostly
dark, having a dark, ferruginous-tinted patch at base, a median
band coloured as in forewing, a broad purple-bluish shade distally to
median band as far as subterminal, redder shading from subterminal
to terinen. Underside duller, the dark markings similarly placed,
but uniform fuscescent, diffuse and ill-defined.
Gran Chaco, near Florenzia, October 1902 (S. R. Wagner),
type in coll. Br. Mus. A pretty and distinct species, which
in its coloration and banded forewing may be very roughly
compared with Ptychopoda (?) rusticata (Hiib.) of Europe,
but is of much more diminutive size.
Note. — I have a further species of Ptf/chopoda, from La Paz,
Mendoza (VV. M. Bayne), presented by Mr. A. F. Bayne, which I
cannot at present identify, but which is not in sufficiently good con¬
dition to be satisfactorily made the type of a new species, and must
therefore wait for the discovery of supplementary material.
26. Ptychopoda (?) muscilineata (Dognin).
Pfychopoda muscilineata, Dognin, Ann. Soc. Ent. Belg., 1,
104 (1906).
Tucuman (type), in coll. Dognin.
27. Ptychopoda (?) longipedata (Warren).
Ptychopoda (?) lonqipedata, Warren, Nov. ZooL, vii, 163
(1900).
Parana, Entre Rios (type), in coll. Rothschild.
Mr. Warren, in erecting this species, raises a query as to
its generic position, presumably on account of its leg-
structure and the shape of the wings. He does not mention
the venation, but I am inclined to suspect it will prove to
have the areole double, and to be an aberrant member of
Geometridaa of the Argentine Bepuhlic.
219
111}' new genus Metasiop>ds, next to be described. It does
not seem to liave been observed, unless possibly by Moschle]-
in erecting his genus Leptostciles, how frequent is the double
areole in the smaller new-world Acidaliids, which super¬
ficially might have passed as congeneric with the Ptychopoda
or the Acidalia of the Palaearctic Region.
Metasiopsis (Prout), nov. gen.
Face normal (broad, flat, smooth-scaled) ; palpus minute in the
type-species, always small, shortly rough -scaled below; tongue pre¬
sent, slender ; antenna of ^ dentate, with single longish bristles and
fascicles of longish cilia,* of ^ subserrate, minutely ciliated. Hiiid-
tibia in short, spurless, somewhat hairy, hindtarsus wholly, or
almost wholly aborted ; hindtibia in $ with terminal spurs only,
and these usually not long. Forewing with co-sta subconcave or
nearly straight, apex (especially in 9 , which is narrow-winged) acute,
though not produced ; hindwing rather narrow, particularly in the
9- Forewing with cell somewhat over one-half ; SC2 arising from
cell well before its apex, anastomosing very shortly with SC' and
then more strongly with forming a double areole ; stalk of
SC®-® from just before apex of cell; radials normal; M' approxi¬
mated to R®. Hindwing with C anastomosing at a point with SC,
thence gradually diverging ; SC® long-stalked with R' ; M' closely-
approximated at its base to R®.
Type of the genus : Metasiopsis dcxicosta (Warten) =
Ptychopoda fiexicosta, Warren.
I can find no existing genus, with the possible exception
— if Moschler’s characterisation is defective — of Leptostales,
to which Warren’s fiexicosta ai d its nearest allies can be
referred. Its double areole, and its narrower hindwing,
with C closely approximated to cell for some distance
beyond the subbasal fusion, separate it from Ptychopoda.
Moreover Mr. A. F. Bayne tells me that its resting posture
is different, the wings being more closed. The sexual
* \o. plemyraria, Guen. ( = balisturia, Gnen. = peri rrumta, Pack.),
which is rather closely allied to typical Metasiopsis, the bristle is
long and strong, the cilia apparently ill-developed ; I presume the
bristles are the “ lames, longues, mais extremement minces ” of Guenee
(Spec. Gen. des Lep., ix. 453), who classes the antenna as “pectinee.”
It may well be that Mdschler’s troublesome Leptostales (Abh. Senck.
Ges., xvi. 238), which is also defined as possessing pectinated ^
antenna, comes under the same heading, and that Metasiopsis will
prove to be a development ot this ; but he gives the f hindtibia as
having two spurs.
220
Mi^jouis B. Prout on the
dimorphism in the type species is very pronounced, and it
is not surprising that Mr. Warren, with only museum
material available, described the two sexes as separate
species. Apparently near relatives, which I would at least
provisionally refer to Metasiopsis, are plemyraria, Guen.,
Spec. Gen. des Lep. ix, errata {Acidalia), ossularia, Hiib.,
Zutr., fig. 900-10 {Lcptomeris), trianyularis, Warr. {vide
infra), and no doubt ohliquaria, Warr., Nov. Zool., vii, 147
(Antcois) ; the last-named I have not seen since my atten¬
tion was directed to the generic distinctness of Metasiupsis,
but it appears to be closely allied to plemyraria, and, as
Warren says, it reminds also of trianyularis. Probably
further research will add others to the genus, perhaps
including longipedata, Warr., as suggested above. Hiibner’s
osmlaria has the palpus better developed than in Jlexicosta,
but most of its characters agree well with those of the
last-named, and the sexual dimorphism, though less pro¬
nounced, shows the same tendency ; plemyraria is even
nearer, at least in palpal structure.
On account of the confusion into which these genera
have fallen, and the little attention which has been paid
to the double areole, even by authors who, like Hulst, have
intended to use it as generic, it will not be out of place to
point out wherein the new genus differs from others pos¬
sessing the character. From Euacidalia, Pack., which
virtually agrees with it in leg-structure in both sexes,
MctasiopAs is distinguished by the shape of the wings and
by the point of origin of SC^ of the forewing ; in Euacidalia
this vein is stalked with SC^A From Pigia, H.-S., it also
differs in both these respects (in Pigia SC'^ is short-stalked,
or at least connate, with SC^'^), but here, in addition, the
$ hindleg has four well-developed spurs. From the large
and somewhat heterogeneous group which, pending fuller
revision, I unite as Pdamalia, Hiib., in which SC'^ arises
from the cell as in Metadopsis, the absence of the median
spurs in the $ affords the readiest distinction ; but I do
not know of any species of Hamalia with vein C of the
hindwing as in Metasiopsis. Scelolophia, Hulst, and Tri¬
centra, Warr., have SC‘^ and of the hindwing connate
or very short-stalked, apart from various “secondary sexual”
characters, ^emaeopus, H.-S., agrees with Metasiopsis in
forewing venation, but differs in almost everything else —
size, shape, leg-structure (both sexes), hindwing venation,
etc. The Old World genera are not at all likely to collide
221
Qeometridae of the Ao'rjentine Rejmhlic.
with the present one ; but it may be remarked that they
seldom have SC^ arising from the cell. Rhodostroiihia,
Hiib., Dithalama, Meyr., Dithccodes, Warn, Antanepsia,
Turn., “ Erythrolophus ” semiustus, Hmpsn., and one or two
species of Craspediopsis, Warn, alone occur to me, though
exhaustive research would no doubt reveal a few others.
None of those named bear any resemblance to Metasiopsis,
and it would be easy to point out valid structural distinctions
in each case.
28. Metasiopsis flexicosta (Warren).
$ Ptychopoda JlexicostA, Warren, Nov. Zook, vii, 101
(1900).
^ Ptychopoda mb^trigata, Warren, Nov. Zool., vii, 104
(1900).
Buenos Aires, December to March, flies out of “ Alfalfa”
(lucerne) (A. F. Bayne) ; San Luis (W. M. Bayne) ; La
Paz, Mendoza (W. M. Bayne) ; Parana, Entre Rios (both
Warren’s types, besides other ^s, in coll. Rothschild) ;
Tucuman (coll. Dognin et coll. Bastelberger) ; Goya, Cor-
rientes (coll. Bn Mus.) ; Ocampo, Santa Fe, August 1902,
and near Florenzia, Gran Chaco, October 1902 (S. R.
Wagner, in coll. Bn Mus.).
Apparently a common and widely-distributed species in
the Argentine Republic, but with the exception of a pair
from Sapucay, Paraguay, in coll. Bn Mus., I am not yet
acquainted with any taken outside its boundaries. The
Z is very variable in depth of colouring ; the $ somewhat
less variable, always pale, with much stronger and more
diffuse lines (bands) on the underside, somewhat recalling
those of our European Aspitates (Asjnlates).
29. Metasiopsis triangularis (Warren).
Ptychojioda triangularis, Warren, Nov. Zook, iv, 445
(1897).
Anteois triangularis, Warren, Nov. Zook, vii, 147 (1900).
Gran Chaco, near Florenzia, October 1902, in coll. Bn
Mus.
The typical form occurs in S.E. Brazil. The only two
Argentine examples which I have seen are of a slightly
paler colour although in perfect condition ; but I can
see no essential difference.
222
MiT Louis B. Front on the
Hamalia (Hiibtier).
Haemalea, Hiibiier, Zutr., ii, 11 (indescr.).
Hamalia, Hiibner, Verz. bek. Sclimett., 309 (1826 ?).
Acratodes, Guenee, Spec. Gen. des Lep., x, 57 (1858).
Lipomelia, Warren, pars, Nov. Zool., passim (1895, etc.)
(nec Proc. Zool. Soc., 1893, jx 359).
I include under this genus those Acidaliids with
normally-shaped wings, in which the hindleg structure
agrees with Acidalia ( = Leptomeris, Meyr. ), the $
leg being aborted, without spurs, the ^ with all spurs,
but in which the areole is double, SC" arising from the
cell, and anastomosing with SCh It will perhaps be prac¬
ticable to divide it, but the stalking of SC" and of the
hindwing, which gives such reliable distinction between
Acidalia and Ptychopoda, does not here seem very promis-
ing ; by far the most usually they are quite short-stalked
or connate, very rarely separate or long-stalked, and it is
just possible that their apparent congeners which show the
long stalking are not really so close to them as they super¬
ficially appear. Two or three South American groups
(? subgenera) which Warren catalogues as Lipomelia — the
ms«-group, the «6?e/a-group, etc. — clearly belong here rather
than to Lipomelia (suhusta, Warr.), which has the areole
simple.
30. Hamalia flexifascia (Front), nov. sp.
(Plate XLVIll, fig. 21.)
5 . 21 mm. Head, body and wings nearly concolorous, glossy
light ochreous-brown. Forewing with a somewhat curved ferrugin¬
ous-brown median band, which is narrow at costa, widens a little,
passing behind the upper part of cell, then bending basewards .so
that its proximal edge cuts across lower angle of cell, here forming
a somewhat darkened triangle ; a sinuate, somewhat interrupted
dark line margins the band proximally, while in the centre of the
band there are some patches of somewhat brighter brown ; a dark
subterminal line, following nearly the same course as in Hamalia
adela (Dogn.),* but forming a stronger tooth on and a somewhat
broader lobe in middle of wing, reaching very near the distal margin
below ; a series of dark dashes on distal margin between the
veins, but apparently .shorter and less strongly developed than in
* Acidalia adela, Dogn., Le Nat., 1890, p. 38 ; Lep. Loja (2), 62,
tab. vi, 20.
Geometridae of the Argentine Bcpuhlic. 223
adela, where the pale vein-ends only narrowly break the continuity
of the dark line. Hind wing slightly shaded with grey in basal part,
an elongate cell-spot, shortly followed by an indistinct grey median
line ; an indistinct outer dark line nearly parallel with termen, but
slightly wavy, and somewhat sinuate inwards distally to cell and
below' Mb Underside unmarked, forewing tinged with reddish,
especially at costa and on veins (much as in adela).
Buenos Aires, January 27, 1907 (A. F. Bayne). Type
in coll. L. B, Prout.
I fancy the species has been overlooked, or mixed with
some of its allies ; if it has ever been named, I have been
quite unable to trace it. It comes rather near some forms
of adela, Dogn., but, apart from certain superficial differ¬
ences noted above, the $ antenna apparently has rather
longer ciliations, and the hindwing has SC^ and shortly
stalked, whereas in all the adela which I have examined
they are connate or just separate.* Besides the type
specimen,! possess a worn $ trom Ciudad Bolivar, Orinoco,
Venezuela, dated August 17, 1898, which is quite clearly
conspecific, and the British Museum has a $ from Dominica
and another from Grenada which I refer here, the latter
darker-coloured and with the hindwing almost unmarked.
All these three are smaller than the type specimen, perhaps
on account of sex.
31. HaMALIA (?) PTYCHOPODA (PlOUt), nov. sp.
(Plate XLVIII, fig. 29.)
^ $. 17-18 mm. Face fuscous, vertex of head white; palpus
short, brown, 2nd joint dark-marked on the outer side ; antenna in
^ (?), in 5 minutely ciliated (more or less broken in all the speci¬
mens). Thorax and abdomen concolorous with wings ; ^ hindtibia
thickened and with pencil of hairs, hindtarsus very short and curved ;
$ hindtibia perhaps wdth one median spur absent.f Wings whitish
bone-colour, sprinkled with brown atoms. Forewing with extreme
costal edge fuscous ; discal dots grey -blackish, in the type very small,
* I have seen a single adela in which they might, by a stretch of
the term, be described as “ stalked,” their common base being slightly
prolonged beyond the mere point ; but even this differs materially
from the formation in jlexifascia.
t This is certainly the case with the only 9 hindleg available for
examination, and I cannot find any indication that a spur has been
broken off ; the type 9 has lost one hindleg, the other 9 both. Hence
there is sufficient evidence to justify a query as to the generic position
but not enough for placing it elsewhere.
224
Louis B. Front on the
in the other specimens distinct ; transverse lines vague, formed by-
increase of the dark sprinkling, tlie inner and outer, however, usually
rendered more distinct by dark dots on the veins ; inner line from
one-third costa to about two-fiftlis inner margin, oblique distad from
costa to there bent (almost angled), and thence somewhat in¬
curved ; outer line starting (oblique distad) fi’om a rather distinct
dark costal spot at about three-fourths, nearly parallel with termen
from R2 to inner margin, but somewhat sinuous ; median line (shade)
obsolescent in the type specimen, fairly distinct in the ^ , where it
is seen to be placed midway between the other lines, somewhat
curved outside discal spot, and faintly sinuate basewards in sub¬
median area. Hiiidwing in the type without discal dot, which,
however, is distinct in the other specimens ; two indistinct lines,
the inner crossing discal spot (when this is present), the outer mid¬
way between this and termen. Terminal line of both wings dark,
broadly interrupted at the vein-ends ; fringe pale, dark-spotted at
the vein-ends. Underside without markings.
Mendoza (foothills to Andes), type ($) and one ^ and $
in coll. L. B. Front, all taken February 8, 1903, by W. M.
Bayne. Others in coll. A. F. Bayne, who describes it (in
litt.) as “a fairly common species.” Not known from any
other locality.
In shape and general appearance, this species may be
likened to Haracdia lotydata and its allies ; but
its small size and the weakness of the markings give much
suggestion also of a species of Ptychopoda, with which it
further agrees in the long-stalking of SC^ and in hind¬
wing. The insect may be somewhat variable, but un¬
fortunately none of the specimens are in very fresh condition.
The general facies is pretty uniform, but in the $ the lines
seem rather stronger, and are rather further apart, than in
the ^s. In all the specimens there is some indication of
darker shading in the distal area, but this is never very
prominent, and perhaps sometimes tends to become more
irregular, or concentrated towards the tornus, as in certain
well-known species of both Kamalia and Ptychopoda.
32. Hamalia (?) PLATENSIS (Frout), nov. sp.
(Flate XLVIII, fig. 11.)
9 . 20-22 mm. Face brown, darker above ; vertex white, occiput
and patagia brown, the rest of the body white ; palpus rather short,
* Acidalia botijdata, Walk., List Lep. Ins., xxii, 727.
225
Geometridae of the Argentine Rejntblic.
brown, darker spotted, beneath white ; antenna basally white above,
otherwise brown, ciliation very short. Wings white. Forewing
above between costa and vein C fuscous to beyond one-half ; mark¬
ings light fuscous brown ; antemedian line scarcely indicated, a spot
on costa at nearly two-fifths and one on inner margin at nearly one-
half indicating its position ; a median line or shade indicated by a
spot on costa at about middle, and one on inner margin beyond
middle, and faintly traceable in submedian area in fi’esh .«pecimens ;
postmedian line at nearly three-fourths, indicated by a distinct dark
spot on costa, and a rather indistinct, curved series of vein dots ;
subterminal area with light fuscous clouds, the smaller being across
R*, the larger almost continuous, band-like, from above R® to inner
margin, but really divisible into two spots on R^ and M* and a small
band starting proximally to these, just above JVP, and reaching to
inner margin just before tornus ; some more indistinct terminal
clouding ; cell-spot blackish; marginal line blackish, interrupted at
vein-ends. Hindwing weakly marked, faint suggestions of post¬
median line and sub terminal clouding ; marginal line as in fore-
wing. Underside with costal margin tinged with brown, less darkly
than above, but more broadly (about to SC) and reaching to apex ;
cell-spot of forewing usually present ; otherwise almost without
markings ; marginal line of both wings nearly as above.
Buenos Aires, 3 collected by A. F. Bayne, February 10,
1903 (type), February 23, 1906, and one without date, all
in coll. L. B. Prout. A fourth from the same locality,
in coll. Br. Mus. (H. Wilkinson).
The ^ is still unknown. The $ has four well-developed
spurs and the double areole as in Hamcdia', but in the
hindwing- SC“ and are stalked for nearly one-half their
length.
SCELOLOPHIA (Hulst).
Scelolophia, Hulst, Tr. Amer. Ent. Soc., xxiii, 301 (1896).
Calyptocome, Warren, Nov. Zook, vii, 148 (1901) (nov. syn.).
The ^s of this genus can readily be distinguished from
Hamalia by the tufts or fringes of hair on the hindwing
beneath, the $s by having the outer median spur extremely
small, or often wanting. I have found variation in the
latter point between very close allies; thus it is present in
Scelolophia phor curia (Guen.), absent in>S'. roseo/wa (Warn),
which is sunk at the British Museum to phorcaria ; see
also Warren, Proc. U.S. Mus., xxx, 437.
Mr. Warren, in erecting Calyptocome, overlooked Hulst’s
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) Q
22G
AlrT Lonis B. Prout on the
older genus, which is founded on the same type, imiinaria,
Guen. = formosa, Hulst. I was in the first instance
indebted to my friend Mr. R. F. Pearsall, of Br ooklyn (in
lift., April 30, 1906), for this determination, and my own
study of the literature confirms it.
33. SCELOLOPHIA TURBATA (Walker).
Cambogia turhata, Walker, List Lep. Ins., xxvi, 1755
(1862).
Acidalia concolm'aria, Druce, Biol. Centr.-Amer., Lep.
Piet., ii, 121 (1892) (? Dognin, 1890).
Gran Chaco, near Florenzia, one in coll. Br. Mus.
The species apparently ranges from Mexico and the
W^est Indies to S.E. Brazil, and I cannot definitely differ¬
entiate it from the North American ‘pannaria, Guen.,
but forbear to sink it until I have been able to study more
North American material. The Gran Chaco specimen is
in rather poor condition, but seems to agree very exactly
with Walker’s type from Brazil.
34. PoLYGRAPHODES DENTICULATA (Warren).
FolygrciflwcUs dcnticulata, Warren, Nov. Zook, vii, 159
(1900).
Parana, Entre Rios, the type and “ a good series,”
and ^s, in coll. Rothschild; Goya, Corrientes. in coll. Br.
Mus.
35. Acidalia* eburneata (Guenee).
Acidalia eburneata, Guenee, Spec. Gen. des Lep., ix, 474
(1858).
Tucuman in coll. Dognin ; Goya and near Florenzia, in
coll. Br. Mus.
A common and widely-distributed species in South
America, reaching from Mexico to Brazil and Argentina.
36. Acidalia subquadrata (Guenee).
Acidalia subquadrata, Guenee, Spec. Gen. des Lep., ix,
459 (1858).!
Acidalia confertaria. Walker, List Lep. Ins., xxii, 733
(1861).2
* On the correct use of tlie name Acidalia (= the LqAomeris
of Meyrick, Graspedia of Ilampson, Emmiltis of Warren) vide
Entoni., xlii, 3. (1909).
227
Geometridae of the Argentine BepuUic.
Acidalia apparitaoda, Walker, List Lep. Ins xxii 733
(1861).3
Craspedia internexata, Warren, Nov. Zool. xi 35
(1904)^ (nov. syn.),
Tucuman, in coll. Dognin ; Gran Chaco, near Florenzia
in coll. Br. Mus. ’
Also known from (i) Brazil and French Guiana, C) (3)
Honduras, (^) Ecuador, and indeed most parts of Central
and South America, from Mexico to Argentina. Very
common and variable. The British Museum example
belongs to Guenee’s var. A. I suspect that Warren’s other
species of the group {atomaria, approbata, trias) will have
to sink ; possibly also Walker’s compensata from Florida.
37. Acidalia recusataeia (Walker).
Acidalitt, recuscLtOjria^ Walker, List Lep. Ins xxiii 788
(1861).
Los Vasquez, in coll. Dognin.
Walkei s type (a ^) was from Brazil, and is in worn
condition. A paler ^ labelled “ Argentina ” in coll. Br.
Mus. probably belongs here ; it is at least closely related,
agreeing in all essentials.
38. Acidalia defixaria (Walker) (?).
Acidalia defixaria, Walker, List Lep. Ins xxii 731
(1861).! ^ ’ ’
1 Acidalia martharia, Walker, List Lep. Ins., xxxv, 1627
(1866).
Goya, Corrientes, in coll. Br. Mus.
Walker’s type(i) came from Venezuela. The two speci¬
mens which are thus determined at the British Museum
are in damaged condition, but hardly seem to me to belontJ"
to this species, as their colour appears to be more ochreou?
and their central shade straighter.
39. Acidalia albidulata (Warren) (?).
Craspedia albidulata, Warren, Nov. Zool., iv, 432 (1897).
Goya, Corrientes, in coll. Br. Mus.
These examples are mostly in poor condition, but
evidently agree with a form from Paraguay which has
been determined for me by Mr. Warren as his alhididata,
described from S.E. Brazil. On the other hand, they are
Q 2
228
Louis B. Front on the
smaller, with the postmedian line of both wings more
dentate, than a topotype of alhidulata presented to the
British Museum by Mr. Schaus. Except in the presence
of the black dots on the post median line, or at least
of the characteristic one on IF, they come very close to
certain forms of A. cclipes, described below.
40. Acidalia qnicornata (Warren).
Craspedia nnicornata, Warren, Nov. Zool., vii, 151 (1900).
Tucuman and Los Vasquez, in coll. Dognin.
Warren’s types were from S.E. Brazil. I have not studied
M. Dognin’s specimens, but they have no doubt been
determined by Mr. Warren.
41. Acidalia eclipes (Front), nov. sp. (Flate XLVIII,
fig. 18.)
^ 9- 18-20 mm. Face dark fuscous ; vertex whitish ; {^antenna
moderately ciliated. Wings whitish, faintly tinged with ochreous,
sparsely sprinkled with minute blackish fuscous dots (discernible
only on magnification), and traversed by weak grey-brown lines and
shades ; an antemedian line, bent in cell ; a subdentate postmedian,
parallel with termen, but making a moderate angle distally on ;
an oblique median shade, somewhat angled on R* ; some indistinct
shading on either side of the sinuous pale subterminal ; discal dot
small, black ; a series of small black terminal dots, or short dashes,
between the veins. Hindwing similar, without the antemedian
line ; median shade passing on the proximal side of the cell-spot,
sometimes nearly straight, but usually somewhat bent round it.
Underside whitish, the markings usually weak excepting the post¬
median line, which is rather sharjj, and the terminal dots, which are
as above ; discal dots sometimes well expressed.
Buenos Aires, not uncommon ' from January to early
March (A. F. Bayne), the type ($, February 27, 1903), and
others (both sexes) in coll. L. B. Frout — also in coll. A. F.
Bayne ; ? Gran Chaco, in coll. Br. Mus.
Somewhat variable. The discal dots are usually very
distinct, but I have one specimen in which they are obso¬
lescent on the fore- and minute on the hindwing. In
another, the wings are more strongly powdered with dark
atoms, and the central shade is more strongly developed,
so that the whole insect appears darker.
A thoroughly typical Acidalia in all points of structure,
229
Geomctridae of the Argentine RepMic.
and reminding in its aspect of the Palaearctic A. ocliro-
leucata, H.-S., which in its turn belongs to a large and
difficult group, embracing the African minorata, Boisd.,
and others. From the two preceding species, to which it
also comes very near, it is distinguished by having the
postmedian line somewhat differently formed, and not
marked with blackish points ; from alhidulata also by its
whiter underside, and perhaps somewhat less elongate
wings ; from unieornata by the usually less dusted upper-
side and less ochreous vertex and costa. It may, however,
be a local form of the last-named, its already mentioned
dark aberration particularly approaching it ; or it may
prove to be a local race of one of the little-known species
from further north, such as A. discriminaria, Walk., List
Lep. Ins.,xxii, 732, from Venezuela. In any case it seemed
desirable to figure and call attention to the form, and
better not to refer it definitely to any known species than
to risk associating it with the wrong one, very little
material being available for detailed study.
The two rather small weakly-marked specimens from
Gran Chaco (near Florenzia, October 1902), in the British
Museum collection seem likely to belong here, but I
am not certain. To some extent they look intermediate
between this species and alhidulata.
42. Acidalia ATRiDiscATA (Warren).
Craspedia atridiscata, Warren, Nov. ZooL, xi, 33 (1904).
Tucuman, in coll. Dognin.
Warren’s type was from Bolivia. I know of no other
localities.
43. Atyria chibcha (Schaus).
Flavinia chibcha, Schaus, Proc. Zool. Soc., 287 (1892).
Cyllopoda tenuis, Warren, Nov. Zool., xii, 312 (1905) (ab.)
(nov. syn.).
'L’ucuman and Metan, in coll. Dognin ; Sante Fe
(Ocampo) and Goya, in coll. Br. Mus.
Schaus’ type was from Peru. The determination of the
British Museum specimens is, I think, by Mr. H. Druce.
Cyllopoda chibcha, oh. cuneifera,Vi axr., Nov. Zool., xi, 15,
does not belong here, but to the following species.
I refer provisionally to the genus Atyria (Hiib., Zutr.,
indescr.) H.-S., the species of the Cyllopoda group in which
230
w Louis B, Front on the
the areole is simple. Mr. Warren, on his favourite char¬
acter of the ^ antenna, recognises at least four genera —
Cyllopoda, Dahn., Atyria (Hiib.) H.-S., Atyriodes, Warr.,
and Paratyria, Warr.
44. Atyria semidivisa (Warren).
? Xanthyris basina, Boisduval, Lep. Gnat., 93 (1870).^
Cyllopoda semidivisa, Warren, Nov. Zool., iv, 420 (1897).^
Flavinia Druce, Ann. Mag. Nat. 'Hist., (7) iii, 296
(1899) (nov. syn.).^
Cyllopoda chihcha, ah. mTiet/cra, Warr., Nov. Zool., xi, 15
(1904) (nov. syn.).^
(2) Parana, Entre Rios (types), in coll. Rothschild.
Also recorded from Bolivia (^) and Paraguay (‘^). The
British Museum has examples from the latter locality.
Mr. Warren (in litt.) tells me he considers it to be the
same as Boisduval’s Xanthyris basina (^) from Guatemala
and Venezuela; but I suspect, on geographical grounds,
that the two may be rather close allies than synonyms.
Even at Santa Catharina, according to Boisduval, basioia
is replaced by another species (castina, Boisd., nom. nud.).
45. Atyria durnfordi (Druce).
Flavinia durnfordi, Druce, Ann. Mag. Nat. Hist., (7) iii,
296 (1899).
Buenos Aires (taken by Durnford), in coll. H. Druce.
Also occurs in Paraguay.
46. Atyria isis (Hiibner).
Atyria isis, Hiibner, Zutr., ii, 31, fig. 385-6 (1823).^
Fla.vinia dubia, Schaus, Proc. Zool. Soc., 286 (1892) (nov.
syn.).2
The northern provinces of Argentina, according to
Burmeister, Descr. Phys. Arg. Rep., v (1), 420.
Distribution : Brazil (^) (^) ; Paraguay (Burmeister, loe.
cit.).
Note. — Burmeister (/or. cf^.) also records Atyria osiris.
Cram.,” from the northern provinces and Parana, Entre
Rios. I am quite unable to say what species he had
before him. The true osiris, Cram. {Cyllopoda), is only
known to me from the Guianas, and the species which is
often confounded with it, Cyllopoda jatropharia (Linn.),
231
Geometridae of the Argentine Re'pnblie.
also belongs to more northerly latitudes. Besides, Bur-
meister implies that both his species have the simple
areole. Possibly his osiris was Atyria {Atyriodes) approxi-
mans, Walk., which agrees in some points (e.g. white face)
with his description, and occurs in Brazil. The only
species which I have yet definitely recorded from Parana
is A. semidivisa (Warn), which is too unlike Cramer’s
figure and Burmeister’s description to satisfy the needs of
the case at all.
^\xh4a,m{\j—LARENTIINAE.
Eudulinae, Heterusiinae, Eiicestiinae, Trichopteryginae, Te-
phroclystiinae, Hydriomeninae et Astheninae (Warren).
47. Eudule cupraria (Walker).
Crocota cupraria, Walker, List Lep. Ins., ii, 536 (1854).
Euphanessa unicolor, Robinson, Ann. N.Y. Lyc. Nat. Hist.,
ix, 153, tab. i, 2 (1869) (nov. syn.).
Eudule hesperina Burmeister, Descr. Phys. Arg. Rep.,
v(l), 428 (1878) (nov. syn.).
Buenos Aires, one February 10, 1903 (A. F. Bayne),
Mendoza, one $ , June 1902 (W. M. Bayne), both in coll.
L. B. Prout; Santa Fe, August 1902, and near Florenzia,
October 1902 (S. R. Wagner), in coll. Br. Mus. ; various
localities in the provinces of Cordova and Buenos Aires
(Burmeister).
The specimen which bears Walker’s type label was from
Peru. The species is of general distribution from the
southern United States to Buenos Aires. The North
American authors do not appear to have discovered the
identity of their species (pinicolor, Rob.) with Walker’s, as it
is still catalogued in Dyar’s List under its younger name.
48. Eudule hesperina (Burmeister).
Eudule hesperina $, Burmeister, Descr. Phys. Arg. Rep.,
v(l), 428 (1878).
Etcdule costata, Warren, Nov. Zook, iv, 456 (1897) (nov.
syn.).
Provinces of Cordova and Buenos Aires (Burmeister).
I have only seen this species from S.E. Brazil (whence
came Warren’s type) and Paraguay. Burmeister con¬
founded it with the preceding, giving the $ a black costa,
232
M l^^ouis B. Front on the
the ^ not. The distinction is not really sexual, and I
strongly incline to agree with Warren in treating it as
specific ; but it is more in accordance with common pre¬
cedent to restrict Burmeister’s name to his ^ than to his
^.especially as this course preserves an older name, dating
from 1878, and sinks the more recent (1897).
49. Eudule LOBiFORMis (Druce).
EuduJe lohiformis, Druce, Ann Mag. Nat. Hist., (7) iii, 294
(1899).
Los Vasquez, in coll. Dognin.
Druce’s type was from Maranham, Amazons. There is
also, as Druce mentions, a single specimen from Chiriqui,
Panama, in coll. Br. Mus. Its occurrence singly, in three
such widely distant localities, is curious, and one is half
inclined to wonder whether the remarkable costal con¬
formation, wdiich alone seems to distinguish it from
cupraria, is some recurrent abnormality rather than the
mark of a species.
50. Eudule w^eyenberghii (Snellen).
Eudvle weyenberejhii, Snellen, Bob Ac. Cienc. Cordoba, ii,
890 (i877).
Cordova (Weyenbergh), /fA Snellen.
The British Museum possesses two examples from
Bolivia, which I believe have been compared with Snellen’s
type, but which in any case agree very accurately with
his description of the undersurface and the body parts,
though their upperside agrees more nearly, in the smoky
suffusion, with their under than his description suggests.
Perhaps a local form.
51. Mennis limbata (Burmeister).
Eudule limhata, Burmeister, Descr. Phys. Arg. Rep., v (1),
518, tab. xxiv, 4 (1878).^
Mennis eytherea, Schaus, Proc. Zool. Soc. 287 (1892) (nov.
syn.).2
Oran, Gran Chaco (M. Ruscheweyh), Burmeister’s
type(^); La Rioja (Dr. E. Giacomelli), one ^ in coll. Br.
Mus. ; Tucuman, in coll. Dognin.
Schaus’ type (2) Avas from Peru.
Geometridac of the Argentine Bepuhlic. 233
52. Ameria invaria (Walker).
Ameria invaria, Walker, List Lep. Ins., ii, 555 (1854).^
Eudnle unicolor (Herrich-Sebaeffer, MS.), Moschler, Verb,
z.-b. Ges. Wien, x.Kvii, 660 (1877) (nec Robinson,
1869).2
Eudule aurora, Burmeister, Descr. Pbys. Arg. Rep. v (1),
427 (1878).3
Eudule invaria, Berg, An. Soc. Cient. Arg., xiii, 179
(1882).^
Eudulophasia invaria, Warren, Nov. Zook, iv, 456 (1897).
Ameria latiorata, Warren, Nov. Zool., xii, 341 (1905) (ab.)
(nov. syn.).®
(^) Corrientes, one There are also two examples in coll.
Br. Mus., labelled Argentina, O. W. Tbomas, but without
more precise indication of locality.
Range very extensive, from Mexico to S.E. Brazil,
Paraguay and Corrientes, but I have not seen it from
western South America. Walker’s types (^) were from
Venezuela.
Berg’s determination (^) of the names quoted in the above
synonymy (2) (^), is manifestly correct. He informs us that,
as regards Burmeister’s aurora, it is made on the strength
of specimens in coll. Staudinger, which have been compared
with Walker’s types.
The species varies a little in the width of the black
margins, which is somewhat increased in ab. Icdiorata,
Warr. f), described as a separate species.
53. ScoRDYLiA quadruplicaria (Hlibnei’-Geyer).
Heterusia (nom. indescr.) quadrufliearia, Hiibner-Geyer,
Zutr. Exot. Schmett., iv, 7, fig. 603-4 (1832).
Seordylia quadruiflicaria Guenee, Spec. Gen. des Lep.,
X, 384 (1858).
Seordylia qiiadrupliearia var. latior, Thierry-Mieg, Ann.
Soc. Ent. Fr., 1894, 56 (1894) (var.).
Metan, Salta, in. coll. Dognin ; La Rioja (Giacomelli),
in coll. Br. Mus.
Common in Brazil and Paraguay ; Hiibner’s type was
from Rio Janeiro. According to Thierry-Mieg, the form
latior occurs in Peru and N. Brazil, but I have only seen
284
MIFT. ouis B. Prout on the
it fiom Paraguay. S. latissima, Th.-Mieg,* from Peru, I
follow Warreu in treating as a distinct species.
54. SCORDYLIA CEUCIATA (Staudinger).
Seordylia cruciata, Staudinger, Iris, vii, 98 (1894).
Tucuinan, in coll. Dognin.
Staudinger’s types were from Bolivia.
55. ScoRDYLiA ZERiTis (Felder).
Heterusia zeritis, Felder, Reise Novara, Lep. Het., tab. cxxx,
1, la (1875).
Tucuman, in coll. Bastelberger.
Felder’s type was from Bogotd, Colombia. The species
is unrepresented in the British Museum collection.
56. Lithosteoe giacomellii (Prout), nov. sp.
(Plate XLVIIl, fig. 28.)
7 . 22 mm. Head and body grey, sliglitly tinged with brown,
thorax beneath somewhat ligliter than above. Forewing with basal
and central areas and distal margin light grey, finely speckled with
black, a band between basal and median areas ferruginous brown,
outer band (except extreme margin) of the same ferruginous brown ;
basal area bounded by a curved, somewhat irregular w’hitish line ;
median area bounded by whitish lines, tlie jiroximal gently curved,
the distal sinuate, bending outwards below costa, below cell and
above inner margin, which it reaches near tornus ; brown distal area
shading gradually into the greyer termen, a pale subterminal line
scarcely traceable excejjt at costa; terminal line black, interrupted
at vein-ends; fringe (imperfect) apparently pale, chequered with
brown-grey. Hindwing uniform smoky grey, fringe concolorous,
cell-spot extremely indistinct, terminal line less distinct than in fore¬
wing. Underside of forewing weakly marked, divided into two
areas by the indistinctly pale postmedian line, the proximal area
being more grey, the distal more brown ; of hind wing paler brown-
grey than above, the discal dot somewhat less indistinct.
La Rioja (Dr. E. Giacomelli). Type in coll. L. B. Prout.
Two other in my collection (Piclii Ciego, January 25,
1905, W. M. Bayne; La Paz, Mendoza, October 25, 1905,
W. M. Bayne) I provisionally refer to the same species, as
* Scxirdylia quadrnplicaria var. lidissima, Tli.-Mieg, Ann. Soc.
Ent. Fr., 1894, 57 = Heterusia amjdijicata, Warr., Nov. Zool., xi, 89
(1904) (nov. syn.).
Geometridae of the Argentine BepuUic. 235
they agree in structure and general facies ; but in both ,
the ferruginous areas are pale ocbreous, the basal patch
somewhat more angulated, the inner edge of the median
band more strongly concave, and the underside somewhat
lighter and more sharply marked. The La Paz example
is extreme in these respects and may be brietly diagnosed
thus :
ah. angidata, Prout (nov, ab.).
Basal, median and terminal areas much darker, the median
somewhat constricted, pale subterminal line moderately distinct,
conspicuous at costa ; basal area and inner edge of median angled
in cell ; hindwing beneath with a strong dark central line separating
the two areas.
So far as I am aware, this species and those which
follow are the first recorded representatives of the genus
Lithostege in the Neotropical Region, and their discovery
is vei’y interesting, I have much pleasure in dedicating
the present pretty little species to my kind friend Dr.
E. Giacomelli, from whom we may expect further valuable
discoveries in the little-worked district of La Rioja, not¬
withstanding that he finds it but poor in species.
57. Lithostege pax (Prout), nov. sp. (Plate XL VI II,
fig. 30.)
d $ . 20-22 mm. Head and body fuscous grey ; thorax paler
beneath. Forewing with costa and termen somewhat rounded, shape
therefore intermediate between Lithostege and Chesias, which have
not hitherto been very sharply defined ; basal area fuscous grey,
bordered by a darker line, which is bent outwards ; to this follows a
paler area (in the type and some others much paler and sometimes
more brownish tinted, in darker examples little paler and not tinged
with brown), traversed by one or more dusky lines, which are angled
in the cell ; median band fuscous grey, its proximal margin strongly
inbent, its distal margin much as in L. giacomelUi ; a dusky line
traverses this band parallel to its distal margin; a pale, sometimes
white line borders the median band distally ; outer area concolorous
with that between basal and median bands, traversed by some in¬
distinct or moderately distinct wavy lines (vcxriable in individual
specimens), sometimes with the distal margin a little darkened,
sometimes with faint traces of an oblique streak from apex, or of the
“ twin spot” so frequent in the Larentiinae, one with a conspicu¬
ous pale (browmish tinted) dash between SC^ and from end of
230
Mr^^ouis B. Prout on the
median band to subterminal region ; terminal line dark, interrupted
at the vein-ends ; fringe concolorous. Hindwing rather small, fuscous
grey, unicolorous. Underside somewhat paler, usually very w'eakly
marked, with faint traces of the postmedian line on both wings,
which, in one or two specimens, becomes a conspicuous dark
line dividing a darker proximal from a lighter distal area, as in
L. giacomellii, ab. angulata.
La Paz, Mendoza, 4 1 ^ (September to November
1905, at light, etc.), Balde, 1 $ (September 20, 1908), all
collected by Mr. W. M. Bayne, who states (in litt.) that
the moth sits with its wings rather closely folded to the
body. The type La Paz, October 22), in coll. L. B.
Prout, is a rather sharply marked form. The species is
variable, not only in the respects indicated above, but also
in the breadth of the median band, yet is easily recog¬
nised ; only a pale, rather worn $ (March 1903), with the
“twin spots” pronounced, and a slight difference in vena¬
tion, presents a rather distinct aspect, and may be a
separate, though closely allied species.
A noticeable point in the venation of the ^s (excepting
that last named), as also in the unique $ of L. tzaddi, is
the curvature of M^ of the forewing, which arises from M
rather steeply, and rather near M^, afterwards curving into
its normal position. This formation is foreshadowed in
the $ of L. pax, and in many species of the group (par¬
ticularly in Chesias), but seldom so pronounced ; it reaches
its extreme development in Neocliesias, described below.
Veins C and SC of hind wing are, in the ^ of L. pax,
rather far apart towards the base, gradually converge, and
anastomose at a point near the end of the cell ; this is a
slight exaggeration of the structure familiar in the ^ of
the genus Carsia. I do not consider this necessarily
generic; compare the variation in Anaitis.
58. Lithostege (?) tzaddi (Prout), nov. sp.
(Plate XLVIll, fig. 6.)
$ . 26 mm. Face somewhat jirominent, tufted ; palpus moderate,
strongly rough-scaled ; antenna nearly simple, pubescent ; (legs lost).
Head, thorax and palpus grey (white mingled with fuscous), whiter
beneath. Groundcolour of forewing whitish, densely irrorated with
fuscou.s, markings darker fuscous, mostly somewhat ill-defined : a
difi’use subbasal line, very oblique from costa, acutely angled on sub¬
median fold, thence oblique basewards, but obscured by the dark
Geometridae of the Anjentme Bepuhlic. 237
clouding which follows; a broad, diffuse, dark cloud from costa
before one-third, very oblique to middle of wing, thence bent back
to inner margin, but becoming narrower and more indistinct ; some
rather dark shading in the central area, more brown-tinted tlian the
other markings; marginal area broadly darkened, the pale sub¬
terminal line barely traceable; in addition, some characteristic
blackish-fuscous vein-marks, namely : an oblique line from cell-spot
along SC^ to apex of areole; and a figure slightly recalling, on left
wing, the Hebrew letter tzaddi (placed sideways), formed by con¬
nected markings at the origin of and the medians. Hind wing
and undersurface pale brownish grey without markings.
Cacheuta, January 12, 1904 (W. M. Bayne). Type in
coll. L. B. Front,
From the entire structure and aspect of this species, I
feel scarcely any doubt it is a Lithostege (probably rather
nearly related to L. pax), though it is unfortunate that the
legs are lost, and that the J is unknown. The double
areole is broad, stalked with SC^“^, and following
strongly the peculiar curvature described under the pre¬
ceding species ; in the hindvving SC‘^ and R^ are long-
stalked, from near and somewhat curved.
Neochesias (Prout), nov. gen.
Face with horny prominence, short-scaled; palpus moderate,
rather .stout, rough-scaled ; tongue present ; antenna ( d ) slightly
thickened, tapering, pubescent. Thorax scarcely hairy beneath ;
femora glabrous ; foretibia very short, with strong apical claw ;
hindtibia not dilated, with four well-developed spurs. Frenulum
present. Forewing long and narrow, costa slightly arched, apex
rounded, distal margin very oblique, tornus rounded ; cell over one-
half, discocellulars nearly vertical ; areole double, ample ; SC* from
middle of outer areole, SC^ stalked with SC^'^ from its apex, R*
from about two-thirds of its lower margin, M* from near R^,
from very near M*, strongly curved, running in direction of tornus for
about one-third, parallel with M* for about two-thirds ; R^, M* and
thus forming an irregular trident ; submedian area ample. Hind¬
wing ample, apex somewhat produced, yet rounded, distal margin
slightly gibbous between R^ and IVP, faintly sinuate inwards above
and below the gibbosity ; cell slightly over one-half, very broad,
DC parallel with termen for two-thirds, then oblique outwards ; C
and SC diverging widely near base, anastomosing from beyond one-
third to three-quarters of cell ; SC^ connate or short-stalked with
2:38
B. Prout on the
(varying in individual specimens) ; M* from very near or
even connate, curved almost as in forewing ; submedian area
ample, SM^ normal, running to termen.
Type of the genus : Neochesias opistlioleuca, Prout,
nov. sp.
In spite of an evidently close relationship to Chesias
and Lithostege, as shown by frons, antenna, foretibia, etc.,
the genus differs not only from these, but from the whole
of the group (Warren’s Eucestiinae) in the broad hindwing,
with submedian area (and therefore SRP) normally devel¬
oped, in the extreme curvature of of both wings, and
superficially in the general facies. Its allies, moreover,
have the cell of hindwing less (usually much less) than
one-half the wing-length, at least in its anterior portion.*
59. Neochesias opistholeuca (Prout), nov. sp.
(Plate XLVIII, fig. 38.)
. 30-32 mm. Head, face, thorax and legs fuscous, more or less
sprinkled with whitish; thorax whitish beneath; abdomen pale.
Forewing fuscous, irrorated with whitish, and slightly clouded with
darker fuscous ; basal area traversed by several obscure wavy lines,
sometimes in part obliterated by the dark clouding; antemedian
line from before one-half costa to one-half inner margin, rather
direct, but more waved in some specimens than in others ; post¬
median from just beyond two-thirds costa to beyond four-fifths inner
margin, more or less angulated outwards in middle ; the enclosed
median area not traversed by dark lines, but sometimes dark-clouded
distally to antemedian line ; postmedian line followed by a narrow,
bisected pale band, as in many Larentiid genera ; subterminal area
with some obscure wavy bands; traces of a short, ill-defined, oblique
dark mark at apex ; a terminal dark line or series of dashes, inter¬
rupted at the vein-end.s. Hindwing white, unmarked, with indi¬
cations of a dark terminal line, distinct only about apex. Underside
of furewing jiale, almost unmarked, excepting a dark mark at costa
representing the commencement of the postmedian line, sometimes
also a lighter, browner mark midway between this and apex ; of
hindwing white, unmarked.
Aristides Villanueva. September 12, 1905 (W. M. Bayne).
Type in coll. L. B. Prout, Another $ from La Paz,
* If the Stamnodes-Gallipia group really belongs to the Euces¬
tiinae,” it is necessary to add this last clause, as they have the
biangulate discocellulars, with consequent prolongation of the cell
in its lower (posterior) half.
Geometridae of the Argentine Be^niblic. 239
Mendoza, November 24, 1905, in coll. L. B. Proiit, A
third, from Balde, in coll. A. F. Bayne. $ unknown.
Synneuria (Mabille).
Synneuria, Mabille, Bull. Soc. Philom. (7) ix, 67 (1885) ;
Miss. Sci. Cap Horn, vi, Lep. 25 (1888).
Lisso2)sis, Warren, Nov. Zool., i, 398 (1894), (nov. syn.).
60. Synneuria uniformata (Berg).
Carsia uniformata, Berg, An. Soc. Cient. Argent., iv, 208
(1877).!
Synneuria virgellata, Mabille, Bull. Soc. Philom. (7) ix, 68
(1885) ; Miss. Sci. Cap Horn, vi, Lep. 25, tab. ii, 9
(1891).2
Synneuria uniformata, Staudinger, Hamb. Magalh.
Sammelr., ii, Lep. 85 (1899).®
(1) Rio Santa Cruz (F. P. Moreno). Berg states that
Senor Moreno collected about the newly discovered lakes
at the foot of the Andes.
Faunistically this species belongs more to Southern
Chili than to the Argentine Republic. Mabille (^) records
it from Patagonia without precise locality ; Staudinger (®)
from Tierra del Fuego. Mabille, Nouv. Arch. Mus. Paris
(3) i, 156, gives Punta Arenas (Chili), and I myself have
it from there and from Chilian.
The synonymy of the species, as indicated above, was
first made known by Berg in a short note in Bull. Soc.
Ent. Fr., 1889, p. ccxli. As regards the genus, I am
not sure that it is more than a subgenus of Stamnodes,
Guen.
61. Graphidipus subpisciata (Dognin).
Graphipidus (in err. pro Gra])hidipus) subpisciata, Dognin,
Ann. Soc. Ent. Belg., xlvii, 273 (1903).
Tucuman, in coll. Rothschild ; Metan, Salta, in coll.
Dognin; La 'Rioja (Dr. E. Giacomelli), in coll. Br. Mus. et
coll. L. B. Prout.
With the exception of Bolivia, whence the species was
originally described, I know of no other examples than
the Argentine ones here indicated. Hence its geographical
range would appear to be limited.
240
Mf^iOuis B. Front on the
62. Rhopalodes Argentina (Berg).
Bhopalodes arqentina, Berg, An. Soc. Cient. Argent., xv,
164 (1883).
Corrientes, 2 December 1876 (Berg).
I know this species only from Berg’s description, from
which it is clear that it is a true Rhopalodes. It is not
improbable that a closer acquaintance will show it to
supplant some better-known name, e. g. variegata. Warn
(Nov. ZooL, vii, l7l), ab. deriifata. Warn (Nov. Zool., xiv,
246), from Brazil and Paraguay.
63. Rhopalodes (?) muscosaria (Berg).
Rhopalodes muscosaria, Berg, An. Soc. jCient. Argent.,
xix, 273 (1885).
Province of Buenos Aires, one $ (Berg).
Also unknown to me. On account of the small size
(22 mm.), and the fact that Berg knew the $ only, I have
ventured to query the generic position. The insect is
described as having the forewing greenish (or grey-green),
with waved, denticulate black lines, white- or greyish-
margined, the hindwing white.
64. Rhopalodes otophora (Front), nov. sp.
(Plate XLVIII, fig. 37.)
^ 9- 28-30 mm. Forewing wliitisli fleshy, irrorated with
brown, the basal, median, and marginal areas marked with fuscous
greyish; lines blackish, slightly interrupted (more so in the 9 )>
namely : a very irregular line bounding basal area, throwing strong
projections into and below cell; an antemedian from about one-third
costa to one-half inner margin, oblique outwards to middle of cell,
thence approximately perpendicular to inner margin, but more or
less dentate ; a postmedian from beyond two-thirds costa to about
three-fourths inner margin, outangled between SC" and R' and
between R'* and Mh incurved between R^ and submedian fold ; a
grey-brown line between basal and antemedian, following the same
course as the latter ; a faint line dividing the pale area distally to
postmedian ; central area pale in the middle, especially in costal
part ; discal mark elongate, standing in an elongate, somewhat ear¬
shaped pale patch, which is more conspicuous in the $ than in the
type (cj); a pale subterminal line, an indistinct dark subapical
dash, dark tornal spot, and weak, much interrupted terminal line.
GcoTnctridae of the Argentine Reimllic. 241
Hindwing slightly paler, without markings, proximally paler than
distally. Underside nearly without markings, the forewing with
faint indications of the pattern of the upperside.
Puente del Inca (W. M. Bayne). Type $ (bred from
larva, January 5, 1906), and one $ (March 16, 1905), in
coll. L. B. Prout, one $ in coll. A. F. Bayne.
The antenna is pointed at apex, but not appreciably
dilated before it; this character, however, at which Guenee
so greatly marvelled, is not essential to the genus — it
does not occur, for instance, in the group of rosida, Dogn.,
Le Nat., 1892, 274 On the other hand, B. otojjhora is
slightly aberrant in having the lobes of the hindwing
rather smaller than in most of the species. In this respect
it comes nearer to some of the Chilian genera. The
venation, as in many of the allies, is slightly variable, but
the three which I have studied agree in having the double
areole ample, very short-stalked with SC^'^, slightly
nearer to R^ than to Rb disco ?ellulars hardly oblique. In
the hindwing the $ has C closely approximated to SC
to near end of cell, connected by the usual bar before
diverging, SC'^ separate from R^ ; both ^s have G ana¬
stomosing strongly with SC, SC“ and are either connate
with the radials or shortly (M^ very shortly) stalked.
“ Larva green, pupa sage-green ” (W. M. Bayne, in
litt.).
65. Pachrophylla oculata (Mabille).
Lohophora oculata, Mabille, Bull. Soc. Philom. (7) ix, 70
(1885) ; Miss. Sci. Cap Horn, vi, Lep. 28, tab. iii, 7
(1888).!
Lohophora ? {Pachrophylla ?) oculata, Staudinger, Hamb.
Magalh. Sammelr., ii, Lep. 88 (1899).^
(^) Aysen River, W. Patagonia.
Originally described (^) from Orange Bay, in the S. of
Patagonia, which belongs politically to Chili. Physically,
too, the genus is entirely Chilian. The species would
seem to be quite nearly related to some forms of varians,
Butl., Tr. Ent. Soc. Lond., 1882, 400. It has also, no
doubt, much in common with amoena, Philippi = ternata,
Ye\di. = jacintaria. Stand, (nov. syn.), the type of the genus
Tomopteryx. Whether Tomopteryx can really, as Butler
says (Tr. Ent. Soc. Lond., 1882, 402), be kept generically
separate from Pachrophylla, I have not yet been able to
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) R
242
M^f^ouis B. Prout 071 the
decide. The types of the two genera stand rather widely
apart in shape, etc., but they seem to be somewhat con¬
nected by intergrades.
Baynia (Prout), nov. gen.
Face smooth (?) ; palpus moderate, with rather long projecting
scales, 3rd joint pointed; tongue developed; antenna (d) rather
long, tapering at extremity, nearly simple, minutely ciliated. Hind-
tibia not dilated, all spurs present. Abdomen probably long (broken
in the unique type specimen). Frenulum present. Forewing with
margins entire, inner margin with triangular tooth of projecting
scales in middle ; cell over one-half, discocell ulars scarcely oblique,
from very slightly below centre ; areole double, ample, SC“
anastomosing rather strongly with SC^, and afterwards with SC^'^
from point of separation of SC® ; stalk of SC^'® connate with R' ;
M* from close to (almost connate). Hindwing {^) with angles
rounded, distal margin nearly straight between (feebly sinuate) ; cell
about one-half, extremely broad, discocell ulars somewhat oblique
inivards ; C anastomosing with SC for about the middle half of
cell ; SC“ well separate from R^ (as in d® of Rhopalodes, etc.) ; R^
from near R^, obliquely placed so as to reach margin nearly midway
between IR and R'^ ; R^ long-stalked with IVP, the former running to
near anal angle, the latter to the angle ; inner-marginal area folded
and somewhat aborted, a small lobe at its base, fringed with long
hairs, SM^ apparently absent; under-surface with a fringe of long
hairs along M and R®, becoming shorter towards termen. 9
unknown.
Type of the genus : Baynia odontota, Prout, nov. sp.
Evidently related to Triptila, Warn, Nov. Zool., i, 898
( = Phacelophora, Stgn, Hamb. Magalh. Sammeln, ii, Lep.
98 = Diphacelo2}ho')'a, Berg, Com. Mus. Buen., i, 311),
which also has teeth of projecting scales from inner
margin. But in the present genus the hindwing is more
ample, its lobes less developed, and the entire inner-
marginal structure different. Of characters which are
usually regarded as generic, the long stalking of with
is the most noteworthy. Staudinger also emphasises
the three thoracic dorsal tufts in 23^')'ornata, Mab. — Avhich,
if not (as I believe) a synonym oi' virescens, Philippi, is at
least an extremely close ally; in my specimen of Baynia
odontota the thorax is badly worn, so that I cannot say
positively that it may not have been tufted.
243
G-eometridae, of the Argentine Bepiiblic.
66. Baynia odontota (Front), nov. sp.
(Plate XLVIII, fig. 25.)
22 mill. Face pale grey (probably greenish in absolutely
fresh specimens) ; palpus grey, mixed with darker scales. Fore¬
wing greenish,* copiously marked with dark fuscous transverse lines,
which are mostly more or less interrupted : basal line complete,
double in its anterior half ; some interrupted lines between this and
the median area; anteniedian line complete, blackish, strongly
curved in cell, faintly sinuate posteriorly ; postmedian somewhat
interrupted, outcurved distally to cell, incurved in submedian area,
here followed by a whitish mark ; the anteniedian is duplicated
distally, the postmedian proximally, these duplicating lines meeting
at submedian fold, and enclosing a paler area which contains the
elongate cell-spot; otherwise the median area is somewhat darkened ;
some further interrupted lines in distal area, two of them crossed by
dark vein-streaks on and R^, producing5a subquadrate mark ;
dark vein-streaks on M and SM^ in median area ; various other
dark vein-marks and cloudings ; inner-marginal scale-tuft dark
fuscous. Hindwing white, the termen and inner margin shaded
with smoky brown. Undersurface whitish, without markings, the
forewing greyer tinged than the hindwing ; hair-tuft of hindwing
brownish.
Aristides Villanueva, December 26, 1905 (W. M. Bayne).
Type in coll. L. B. Front.
67. Rhopalista dismutata (Warren).
Rhopalista dismutata, Warren, Nov. Zool., xi, 75 (1904).
Tucuman, October 1901, 700 metres (Dinelli). Type
and another ^ in coll. Rothschild. Also in coll. Dognin
from the same locality and from Salta.
Not yet recorded elsewhere.
68. Hammaptera repandaria (Schaus).
Bhopalista repandaria, Schaus, Tr. Amer. Ent. Soc., xxvii,
267 (1901).
Metan, Salta, in coll. Dognin.
* Probably, like the European Acasis vtretata (Hiib.), of a fuller
green when freshly bred. My specimen, which is at present unique,
rather closely resembles in coloration examples of that species which
have stood a few years in the cabinet, or which have been captured
wild.
R 2
244
is B. Front on the
A common species in Brazil and Paraguay, extending
also to Venezuela.
69. Hammaptera combustaria (Herrich-Schaeffer).
Encosmia conilustaria, Herrich-Schaefifer, Samrnl. Aus-
sereur. Scbmett., i, fig. 355 (1855).
Larentia comhustaria, Herrich-Scliaefier, Samml. Ausserenr.
Schmett., i, 52, 66 (1856, 1858).
Cidaria jacintharia, Guenee, Spec. Gen. des Lep., x, 458,
tab. xii, 2 (1858).
Buenos Aires, September 25, 1903 (A. F. Bayne), in
coll. L. B. Front; a second from the same locality in coll.
A. F. Bayne.
Herrich-Schaeffer’s and Guenee’s types were from Brazil,
whence also it is represented in coll. Br. Mus. Mr. Bayne
tells me it occurs rather freely in Montevideo.
70. Hammaptera genurma (Schaus).
Hamma'ptera genurma, Schaus, Tr. Amer. Ent. Soc., xxvii,
270 (1901).
Tucumau, in coll. Dognin.
Schaus’ type was from Peru.
71. Hammaptera jugurtharia (Guenee) (?)
Cidaria jugurtharia, Guenee, Spec. Gen. des Lep., x, 460
(1858).
Tucuman, in coll. Dognin.
The species was originally described from Colombia.
Its exact range is somewhat uncertain, as it belongs to
a group of forms which have not yet been exhaustively
studied, and which may be either piizzlingly similar but
distinct species, or mere geographical forms and aberra¬
tions. They have been received rather freely of late in
some of the rich collections from Eastern Peru. Guen^e’s
was one of the largest forms — he gives 42 mm. expanse;
the Tucuman race (or relative), of which M. Dognin
has very kindly given me an example, is considerably
smaller, and lacks the white subterminal dots in the dark
apical cloud of the forewing beneath, which seem char¬
acteristic of the Peruvian forms. A comparison of Guen4e’s
types is much to be desired.
Geomctridae of the Argentine Hepublic. 245
72. Hammaptera emberizata (Guenee).
Cidaria emberizata, Guenee, Spec. Gen. des Lep., x, 461
(1858)4
Larentia fureiferata, Walker, List Lep. Ins., xxiv, 1188
(1862).2
Cidaria perspieuata, Walker, List Lep. Ins., xxv, 1396
(1862).3
? Cidaria eduetata, Walker, List Lep. Ins., xxv, 1397
(1862).^
1 Hammaptera eonstricta, Warren, Nov. Zoo!., viii, 466
(1901) (nov. syn.).®
Tucuman, in coll. Dognin ; (®) La Soledad, Entre Rios,
one $ in coll. Rothschild.
Range: (}) (^) (^) Brazil; (^) Venezuela ; Peru, in coll.
L. B. Prout. Probably widely distributed, but I confess
I have not fully mastered the variation of the species.
Warren’s suggestion (Nov. Zool., xii, 47) that griimata,
Feld., batis, Warr., and nireigutta, Schaus (= snbpulverata,
Warr., Nov. Zool., viii, 464, nov. syn.), may be forms of it,
does not seem to me tenable. On the other hand, eon-
strieta, Warr., is almost certainly identical with perspicuata,
Walk., which Schaus determines as emberizata, and which
fits well to Guenee’s description.
73. Hammaptera fringillata (Guenee).
Cidaria fringillata, Guenee, Spec. Gen. des Lep., x, 462
(1858).
Coremia lateraria. Walker, List Lep. Ins., xxv, 1310
(1862).
Cidaria interealata. Walker, List Lep. Ins., xxv, 1397
(1862) (nov. syn.).
Tucuman, in coll. Dognin ; Goya, Corrientes, in coll.
Br. Mus.
A very variable little species, akin to the preceding.
So far as I know, it does not extend very far north in
South America, but it is evidently common in S.E. Brazil
and in Paraguay. Although I have not cited them in the
synonymy, 1 have not much doubt that H apicata, Warr.,
Nov. Zool., xi, 51, and H. definita, Warr., Nov. Zool., xiv,
227 (which in any case is a synonym or small ab. of
apieata), will prove to be referable here.
246
Mr. Louis B, Prout on the
74. Hammaptera tenera (Warren).
Hammcoptera tenera, Warren, Nov. Zool., vii, 178 (1900).
Parana, Entre Rios (type and others), in coll. Roths¬
child ; Tucuman, in coll. Dognin, coll. Bastelberger et coll.
Br. Mus. ; Buenos Aires, one $ taken in a “plaza” (scjuare)
in the city, June 18, 1902 (A. F. Bayne), in coll. L. B.
Prout; La Plata City, a good $ taken July 4, 1903 (A. F.
Bayne), in coll. L. B. Prout.
I have also examples from Sapucay, Paraguay, July
1902, and have seen it from S.E. Brazil.
This and the following species, though well distinguish¬
able one from the other in the shape of their median
band, etc., are both very closely akin to the common
parinotata, Zell. (Verb. z.-b. Ges. Wien, xxii, 495 ; anoinala,
Butl., P. Z. S., 1878, 491) of North and Central America
and the West Indies, and either or both of them might
well form a local race of it. I formerly thought it was
tenera which should sink to parinotata, and it is certain
that the more northerly specimens quoted by Warren
{loc. eit.) with his Argentine tenera, really belong to Zeller’s
species ; but on a close comparison I find an even more
exact correspondence between (especially in its
Jamaican forms, anomala, Butl.) and nigrolineata, Warr.
It is evidently best for the present to keep the three
names separate.
75. Hammaptera nigrolineata (Warren).
Hammaptera niqrolineata, Warren, Nov. Zool., xi, 52
(1904).
? Hammaptera fumida, Warren, Nov. Zool, xii, 47
(1905).
Tucuman, in coll. Dognin et coll. Bastelberger.
Sapucay, Paraguay, is the only other locality known to
me for the typical form; I have seen many examples from
there. But Warren’s H. fumida, from Chanchamayo, Peru,
seems to be only a slight local modification of it, and I
have pointed out under the preceding species that it is
quite likely H. parinotata (Zell.) = anomala (Butl.) is
another local race, though it is slightly smaller and
perhaps more slender in build.
Geometridae of the Argentine Republic, 247
76. Hammaptera pontina (Druce).
Melanippe pontina, Druce, Biol. Centr.-Amer., Lep. Het.,
ii, 168, tab. Ivii, 2 (1893).
Tucuman, quoted in M. Dognin’s MS. list as iu liis
collection.
I know only Driice’s Central American specimens
(Mexico and Guatemala). Is the determination correct ?
77. Hammaptera subvinosa (Dognin).
Hammaptera subvinosa, Dognin, Ann. Soc. Ent. Belg., 1, 206
(1906).
Tucuman (type), in coll. Dognin.
78. Hammaptera nigrociliata (Front), nov. sp.
^ . 26 mm. Face blackisli grey ; palpus blackish grey, the joints
tipped with whitish ; pectus and coxae varied with blackish-grey
and whitish, forefemur blackish-grey anteriorly, foretibia and tarsus
blackish with whitish rings, midtibia and tarsus the same, but pale
on their inner side, hindleg pale. Ground colour of wings white,
that of fore wing almost entirely covered by the blackish -grey mark¬
ings, which are without a trace of the green tinge so prevalent in
species of this group. Forewing with dark basal patch, intermediate
band and median band, the three almost joined into one dark area
in one of the specimens (in the type, this part of the wing is much
rubbed, but evidently the space between basal and median areas is
somewhat lighter), the basal and median band.?, however, somewhat
the darkest ; median band occupying middle one-third of costa, and
almost equally broad on inner margin, hence with its outer boundary
here rather near tornus ; its inner boundary somewhat concave, its
outer indented on SC®, sending out a minute tooth anteriorly thereto,
bilobed from R® to M-, and slightly oblique distad from to inner
margin ; a somewhat blacker cell-spot ; a narrow w'hite band
traversed by a weak dark line separates the median from terminal
area, which is again dark, somewhat variegated with a rufous tinge
and with deeper black, and containing the irregular, interrupted,
pale subterminal line. Hindwing white, faintly shadowed with grey
basally, a dark cell-spot rather near base (cell being short), a terminal
black line from tornus nearly to fringe of this part of the wing
blackish ; inner marginal fold moderately developed. Undersurface
of forewing similarly marked to upper, but more’ weakly ; of hind-
248
Mr. Louis B. Prout on the
wing speckled with grey, with weak interrupted grey lines and
bauds, terminal black line reaching SC.^
La Rioja (E. Giacomelli), type and a second $ in coll.
L. B. Prout,
79. Hammaptera (?) ochribasis (Warren).
Entej)hria ochribasis, Warren, Nov. Zool., xii, 332 (1905).
Tucuman (type, $), in coll. Rothschild.
I cannot quote any further distribution Avith certainty,
I fully believe the insect is conspecific, or at least very
closely related, with Hammapteo'a ralaria (Dogn.),* and
Warren’s name may have to sink ; but I cannot assert
this positively in the face of serious questions which have
not yet been entirely set at rest. Typical ralaria is from
Loja, Ecuador, whence also comes a form that M. Dognin
considers conspecihe, yet “not quite typical”; and it is
this latter form — which also occurs in other parts of
Ecuador, in Peru, and I think in Paraguay — that I have
compared with Warren’s type of ochribasis. Again, the
last-mentioned is somewhat discoloured, being stained at
the base (so that the name is unhappily chosen), and this
does not make identification easier; and Mr. Warren, who
has been in constant correspondence with M. Dognin,
determines a different species as ralaria, and a different
species as ochribasis ! In any case the true ochribasis is
not an Entephria,, for it has not the biangulate disco-
cellulars of the hindwing. If it is not a Hammaptera,
it will no doubt prove a Euphyia. The $ is unknown
to me.
80. Hammaptera halisma (Schaus).
Hammaptera hedisma, Schaus, Tr. Amer. Ent. Soc., xxvii,
269 (1901).
Epirrho'e limitata, Warren, Nov. Zool., xi, 117 (1904).
Hammaptera obtusaria, Warren, Nov. Zool.,xiv, 227 (1907).
Tucuman, in coll. Dognin,
Schaus’ and both Warren’s types were from Peru. I
have compared the two last-named with a specimen pre¬
sented by Schaus to the British Museum as halisma.
Except that in obtusaria the median band is slightly
narrower, there is no appreciable difference.
^ Cidaria rolaria, Pogn., Ann, Soc, Ent. Belg., xliii, 252 (1899),
Geometridac of the Argentine RepvMic.
249
Tliis species and a few others which are referred by
Warren to Hammaptera may almost be regarded as tran¬
sitions between that genus and Euphyia, Hlib. (type, picata,
Hlib.), which is evidently the genus frequently referred to
by Warren in his papers as Epirrho'e. The true Epirrlio'e
(type alternata^ Miill. = soeiata, Bork.) has the areole simple,
and I am not aware of its occurrence in the Neotropical
Region.
81. “Euphyia eductata (Walker).”
Cidaria eductata, Walker, List Lep. Ins., xxv, 1397 (1862).
Tucuman, in coll. Dognin.
Quoted merely on M. Dognin’s authority. He records
it as a separate species in his MS. list, and under the
present genus. Schaus, on the other hand, refers it to
Hammaptera emherizata (Guen.), vide Proc. Zool. Soc.
Lond., 1896, 648, and this appears to me correct; Dognin’s
eductata will probably prove different.
82. Euphyia euficorpus (Dognin).
Anapalta riifieorpus, Dognin, Ann. Soc. Ent. Belg., 1, 205
(1906).
Tucuman (type), in coll. Dognin.
I have failed to discover any grounds for maintaining
Warren’s genus Anapalta (Nov. Zool., xi, 43) as distinct
from Euphyia,H\A). — unless possibly for a few species with
different palpi, such as aeerbata, Feld., for which Anapalta
was originally proposed. In any case, most of tlie species
which Warren now refers to his genus have the $ antenna
well ciliated, and the palpus, so far as I have yet investi¬
gated, not materially different from typical Euphyia.
83. Euphyia ruficoesia (Dognin).
Anapalta o'uficoesia, Dognin, Ann. Soc. Ent. Belg., xlviii,
359 (1904).
Los Vascj[uez (type), in coll. Dognin.
84. Euphyia corrivulata (Warren).
Epirrho'e corrivulata, Warren, Nov. Zool., vii, 173 (1900),
Parana, Entre Rios, ^ (type) in coll. Rothschild.
I have not seen any other examples certainly referable
to this — i.e. no others of Warren’s typical form. It is
250
Louis B, Front on the
however, very probable that the type in question is a
small and somewhat abnormal specimen of the following,
in which case corriinilata will be the oldest name for the
species.
85. Euphyia zona (Dognin) (praec. subsp. ?)
(Plate XLVIII, fig. 8.)
Scotosia zona, Dognin, Ann. Soc. Ent. Belg., xlv, 181 (1901).
Scotosia tetrica, Berg, MS. (in Mus. Buen. Aires).
Buenos Aires, two one dated December 1, (1902?),
the other January 24, 1909 — the latter here figured — both
presented by Mr. A. F. Bayne ; one or two others in coll.
A. F. Bayne; Tucuman, one ^ in coll. Dognin.
Described from two ^s and one ^ from Santa Cruz, S.
Brazil ; M. Dognin also possesses a $ from Rio Grande do
Sul (probably Porto Alegre), received through Staudinger.
Larger (usually 86 mm., both sexes, as against 30 mm.)
and darker than Warren’s corrivulata, apparently also
somewhat more glossy ; the basal patch better defined,
the distal margin of median band somewhat less outbent
in middle.
When first I compared this insect with Warren’s type,
I took it to be a distinct species, its size and aspect being
so decidedly different. Then Mr. Warren determined them
as couspecific, and I re-examined the question, with the
results indicated above. I had it figured as “ nov. subsp.”
under the name of tetrica, which, I was told by Mr. Bayne,
it bears in the Buenos Aires Museum. But fortunately I
have learned, in time to suppress that name, that it is
Dognin’s zona. Both he and Prof. Berg were trapped, by
its dark coloration, into calling it a Scotosia. The Buenos
Aires examples and those in coll. Dognin suggest that it is
not at all a variable insect as a rule ; on the other hand,
the existence of this form and corrivulata side by side
at Tucuman prevents my treating it wholly as a local race;
while the presence of a ^ in coll. Br. Mus. (simply labelled
Argentinn, O. W. Thomas), somewhat lighter than M.
Dognin’s and mine, yet agreeing with them in size and all
essentials, gives a further hint that possibly Mr. Warren
may be right in sinking zona.
Mr. Bayne tells me it is fond of coming into houses and
buildings, and is difficult to obtain in good condition.
Oidaria mutabilis, Mab., Bull. Soc. Philom., (7) ix, 69
Geometridae of the Argentine Reimblic. 251
( = Ypsipetes mutdbilis, Mab., Miss. Sci. Cap Horn, vi, Lep.
31, tab. iii, 2), from Orange Bay, Patagonia, stated to be
very variable, may possibly be allied to zona ; but the
lighter, brighter colour, differently formed basal line, and
less outangled postmedian, render a union altogether too
precarious, especially as I only know it from Mabille’s
figure and description.
86. Euphyia limbata (Prout), nov. sp.
(Plate XLVIII, fig. 14.)
(J. 31 mm. Face somewhat protuberant, with some whitish scales
(somewhat abraded) ; palpus red-brown, whitish below; antenna pale
ochreous, narrowly reddish in centre of shaft above, provided with
cilia at least as long as diameter of shaft. Forewing with distal margin
somewhat more oblique than in most of the genus ; brownish white,
irrorated with brown, and rather broadly shaded with brown proxim-
ally to subterminal, and distally to subterminal except from apex to
R* and between R^ and ; markings darker brown, namely : a band
close to base, from costa to middle of cell ; a subbasal curved band
from nearly one-sixth costa to inner margin ; a similar, somewhat
waved, antemedian from about one-third costa ; a postmedian from
about two-thirds costa, forming two small teeth directed distad on
either side of R^, two larger teeth or lobes distad on either side of
M', and becoming a mere line below middle of wing, with further
distad bends on either side of SM* ; traces of less distinct transverse
lines between subbasal and antemedian bands, and (almost losing
themselves below middle of wing) proximally and distally to post¬
median band ; an inconspicuous, extremely minute cell-dot in the
pale median space ; subterminal line pale, indistinct. Hindwing
brownish white, with a postmedian dark line from R' to inner margin,
outbent between R^ and and again below ; marginal area
broadly infuscated, except apically and in middle (as in forewing),
and containing indistinct and interrupted traces of a pale subterminal.
Underside of both wings pale, the forewing with the postmedian
band distinct, but narrowed, and losing itself shortly below M^, a
broad distal dark margin almost as above, but somewhat more distinct;
hindwdng marked as above.
Cacheuta, November 15, 1904 (VV. M. Bayne), type m
coll. L. B. Prout.
Apparently a true Euphyia, DC and of the hind¬
wing formed as in the type species. The hindtarsal spinules
seem stronger and more numerous, but this appearance
may be only due to the abrasion of some of the scales ; in
252
Louis B. Prout on the
any case it would not affect the generic position. The pale
central area and the form of the postmedian somewhat
recall Anticlea derivcda (Schiff.), etc.
CiiRiSMOPTERYX (Prout), nov. gen. (praec. subgen.?)
Face prominent, horny, with strongly appressed scales, not tufted ;
palpus rather strong, of moderate length, somewhat up-curved, with
.s/iorfZ|/ projecting scales* ; tongue strong ; antenna rather long, in
rather thick, with short, even ciliation, in 9 slender, minutely cili¬
ated. Legs rather slender ; hindtibia fully spurred. Forewing
with double fireole, the outer long, SC^ anastomosing with SC^'"* only
just beyond SC*, or almost at the point of their separation, stalked
(sometimes quite considerably) with SC**, DC* inbent, R* normal.
Hindwing rather long and narrow, particularly in 9 i being some¬
what produced apically, but with angles moderately rounded off ; C
anastomosing with SC nearly to end of cell, SC* stalked with R\ dis-
cocellulars not biangulate, R* normal, DC* first slightly inbent, then
oblique outwards. Wing-margins entire or very faintly crenulate,
scaling very glossy, fringes very long.
Type of the genus: Chrismeypteryx expolita, Prout,
nov. sp.
I can find no existing genus to Avhich the species about
to be described can satisfactorily be assigned. Yet it is
not without considerable reluctance that I erect a new one
for them, as their distinctive characters — face, palpus, wing-
shape and texture — seem comparatively slight, and they
may possibly be connected with normal species of Euphyia
by intergrades. In the shape of the hindwing, and in the
shiny, almost greasy appearance of the wings, they some¬
what remind of some of the species oi Larentia" (sens,
lat.) of the Alps of Europe, such as anstriacaria H.-S. ; but
these have biangulate discocellulars in the hindwing, and
normal face and palpus.
87. Chrismopteryx expolita (Prout), nov. sp.
(Plate XLVIII, fig. 34.)
(J. 34 mm. Forewing glossy brown, tinged with fuscous, and
finely irrorated with dark fuscous ; markings weak, consisting chiefly
* In one specimen tliere seems one little tuft of strong, coarse
scales, but in any case the general contrast with the longish-fringed
palpus of Euphyia (and indeed most Larentiids) seems fairly note¬
worthy.
Gcometridae of the Argentine Repuhlic. 253
of three series of dark vein-dots, representing the transverse lines,
and a moderately distinct cell-spot ; subbasal line curved, from about
one-sixth costa, followed by a faintly pale line ; antemedian line
from about two-fifths costa to before one-half inner margin, traceable
chiefly by a faintly pale line which precedes it and by the vein-dots,
somewhat dentate ; postmedian from about two-thirds costa, com¬
mencing obliquely distad, then parallel with termen, a blunt lobe
towards middle, the dots on and M' being placed further distad
than the rest ; the veins, especially R^ and the medians, somewhat
darkened across cell, and less distinctly distally ; no terminal line ;
outer half of fringe paler than inner. Hindwing rather paler than
forewing, unmarked. Underside unmarked. Head and thorax
above concolorous with forewing, abdomen with hindwing.
Puente del Inca (W. M. Bayne), type in coll. L. B.
Prout.
88. Chrismopteryx vicina. (Prout), nov. sp.
(Plate XLVIII, fig. 33.)
9 . 39-40 mm. Colour and general aspect of G. expolita, to which
it is very closely related, differing as follows. The lines somewliat
better defined, and with traces of additional ones between subbasal
and antemedian, and distally to postmedian ; subbasal line some¬
what more oblique ; antemedian more angled in cell ; postmedian
not appreciably inbent on R^ ; a slightly darkened terminal line,
consisting of a series of somewhat crescentic marks separated by the
vein-ends ; underside paler, a darkened cell-spot and postmedian
present on each wing.
Puente del Inca, April 1904 (W. M. Bayne), type in
coll. L. B. Prout. (An extremely worn, almost scaleless
^ taken in the same locality in April 1902, by Mr. A. F.
Bayne, seems to belong to this species, but may be
C. expolita.) Punta de las Vacas, 9> March 16, 1905
(W. M. Bayne), in coll. L. B. Prout. A third 2 in coll.
A. F. Bayne.
It is not impossible that this species is only the ^ of
the preceding, as unfortunately no 9 is known of that,
nor any reliable ^ of this ; but Mr. A. F. Bayne, who has
a very true eye for moths, considers the two distinct, and
the differences noted above, though small, are not wholly
unimportant, so that I am inclined to agree with his
view.
254
^ouis B. Prout on the
80. “ Glaucopteryx tepidata (Guenee).”
?? Tephrosia undularia, Blanchard, Gay’s Hist. Chile, vii,
93 (1854).!
?? Larentia tepidata, Guenee, Spec. Gen. des Lep., x, 281
(1858).!
Glaucopteryx tepidata, Berg, Exped. Bio Negro, 93, tab. ii,
12 (1881).2
(^) Nueva Roma, two specimens.
I have not been able to identify Berg’s species, but
quote it for bibliographical completeness. He gives no
description, and his figure (uncoloured) is only mediocre ;
it suggests a small glossy Larentiid, with very ordinary
forewing, and having, as the most noteworthy feature of
the hindwing, a double series of postmedian (or rather,
postmedian and subterminal) vein-dots. But Berg says
that in some specimens these are entirely wanting. He
evidently, therefore, knew other specimens which he
referred here, besides the two from the Rio Negro
expedition, but how many species he confounded together,
I am not prepared to say. In the British Museum there
is a specimen of Perizoma impromissata, Walk., from
Buenos Aires, which he himself presented as Glaucopteryx
tepidata. Altogether it is most improbable that his
tepidata, or any part of it, was correctly identified as that
of Guenee — nov. nom. for Tephrosia undidaria, Blanch.
This latter was from Chili. (^)
Hoplolygris (Prout), nov. gen.
Frons protuberant, with strong projecting horny plate above
(subconcave in horizontal plane), shorter plate below, and strong
projecting tufts of hairs laterally ; palpus quite moderate, porrect,
2nd joint shortly rough-scaled ; tongue developed ; antenna about
one-half the length of wing, in both sexes nearly simple, minutely
ciliated. Thorax nearly glabrous beneath ; femora glabrous ; fore¬
tibia with well-developed apical claw ; hindtibia not dilated, in both
sexes with all spurs well developed. Abdomen rather short,
moderately stout, the lateral tufts well developed. Wings not
broad, their margins entire ; scaling smooth ; frenulum developed.
Forewing with termen rounded, rather strongly oblique ; cell one-
half ; SC“ anastomosing with SCh and then (shortly, or often at a
mere point) with SC^'^ to form a double areole, of which both parts
are ample ; SC'* separating from SC^ shortly after SC^ ; K.^ short-
stalked or (quite exceptionally), connate with SCH Hindwing
255
Geometridae of the Argentine Republic.
with costa long, apex pronounced ; cell nearly one-half, DC“ very
oblique ; C anastomosing with SC to near end of cell ; stalked
with ; R2 central ; M* well separate from R^.
Type of the genus : Hoplolygris cicatriculata (Berg) =
Lygris cicatriculata, Berg.
A somewhat isolated genus. I have not chosen its
name to indicate any close relationship to Lygris, but to
maintain some nomenclatural association with the original
publication of its sole known species by Berg. I suppose
he referred it to Lygris on account of its type of coloration
and markings, and having thus referred it, deceived
himself into seeing, or imagining “ the wings of the ^
beneath with a tuft of hair at the base ” ; if a tuft exists,
even in the very freshest specimens, it must be so slight
as to be practicably negligible, for a most careful in¬
vestigation has failed to discover it, and Warren (Nov.
Zool., viii, 467) passes the same criticism. Warren (Nov.
ZooL, iv, 454), places the species in Pelurga, Hiib., and
describes its frontal armature, without mentioning how
radically this differs from the simple horny protuberance
of P. comitata, Hiib. He ^overlooks the other differences
(except the wing-outline), namely the apical claw of the
foretibia, the non-biangulate discocellulars of hind wing and
some points of detail in the forewing venation.
The presence of the sharp frontal plates is rare in the
Geometridae, though occurring in a few Australian genera,
such as Giampa, Walk. Its general association with sandy
desert country suggests that perhaps the genus Hoplolygris
will be found to inhabit the pampas, and may have had
its origin there.
90. Hoplolygris cicatriculata (Berg).
Ijygris cicatriculata, Berg, Act. Acad. Cienc. Cordoba, i, 85
(1875); Bull. Mosc., xlix (2), 223 (1876).^
Pelurga straminea, Warren, Nov. Zool., iv, 454 (1897).^
Pelurga cicatriculata, Warren, Nov. Zool., viii, 467 (1901).
(^) Rio Negro, in damp places, November 1874, rather
common, also at Buenos Aires and Cordova ; (f) La Gama,
one November 1893 (Warren’s type), in coll. Roths¬
child; Buenos Aires, a good number (A. F. Bayne and
P. Richards) in coll. L. B. Prout, mostly November, one
each in January and March, also from Berg, etc., in coll.
250
AJp^Louis B. Prout on the
Br. Mus. ; Tucuraan and Los Vasquez, in coll. Dognin ;
Corrientes and near Florenzia, in coll. Br. Mus.
Not yet recorded outside tlie confines of the Argentine
Republic.
91. OcHYRiA ARGENTINA (Berg, MS.) (Pi’out), nov. sp.
(Plate XLVIII, fig. 4).
d $ . 20-23 mm. Face rather strongly protuberant ; palpus
moderate, rather long-haired beneath, 3rd joint moderate, more
shortly rough-scaled ; tongue present ; antenna in d strongly
dentate, with fascicles of cilia, in 9 filiform. Abdomen short.
Forewing triangular, costa little arched, distal margin entire, slightly
convex, oblique (especially in the 9 )> longer than inner margin ;
areole double, the anastomosis of SC^ with SC^'^ very brief, SC®
separating from SC'* shortly after SC® ; hindwing rather narrow
(especially in the 9 ), costa long, distal margin entire, angles some¬
what rounded off, cell less than one-half, discocellulars strongly
oblique, nearly straight or faintly bisinuate, not angled, R® central.
Forewing brown-grey (whitish densely irrorated with brown),
slightly tinged with fleshy ochreous, the usual Larentiid pattern
clearly marked out by whitish transverse lines ; boundary of basal
area before one-fifth, forming an angle outwards in cell ; antemedian
whitish line from beyond one-third costa, gently curved or bent ;
postmedian at about two-thirds, incurved distally to cell, and more
weakly in lower half of wing, forming a rather strong projection at
R® ; subterminal whitish, dentate, especially distinct at costa, placed
on a rather .strongly brown marginal area, which is usually darkest
proximally to the subterminal, and especially over against cell and
towards inner margin ; median area darkest proximally and distally,
its proximal margin spotted with dark fuscous at costa, M, and
inner margin, its distal margin dark-spotted on veins, and dark-
marked from R* to R® ; terminal line dark fuscous, somewhat
variable, usually more or less interrupted at the vein-ends ; fringe
long, dark-chequered at vein-ends, proximal half somewhat darker
than distal. Hindwing brownish grey, usually with weakly paler
l)ostmedian line (course as in forewing), and subterminal ; terminal
line as in forewing. Under-side somewhat richer sandy brown,
forewing with antemedian and postmedian as above, but weaker
(sometimes, as in the type specimen, indicated only by dark costal
spots), and a weakly pale subterminal, preceded by a band of darker
tone tliiin the ground-colour; hindwing beneath dusted with fuscous,
and with the postmedian line distinct, being dark-shaded proximally.
Geometriclae of the Argentine Repuhlic. 257
besides a subterminal line and a darkened band preceding it, much
as in fore wing.
Buenos Aires, April 17, 1905 (A. F. Bayne), tyj^© {$)
coll. L. B. Front. I have two other Buenos Aires ,^s
from Mr. Bayne, one dated, I think (but the label is
nearly effaced), April 18, 1902, the other July 1901, and
noted as “flying on a bright winter day.” The British
Museum also possesses two ,^s from Buenos Aires, sent by
Prof. Berg under the name of Coremia argentina. I know
of no other locality in the Argentine Republic, ]3ut Mr.
Bayne informs me it is common in Montevideo, where he
met with it freely at the end of March 1904 ; the British
Museum has two of each sex labelled Colon, Montevideo,
May 3-7, 1896 (Oldfleld Thomas).
Whether the genus Ochijria has any permanent validity,
I am not at all sure. It is at present used for the species
of the Xanthorhoe group which have the ^ antenna fasci¬
culate, but I know of no other distinctive characters ;
and it is doubtful whether argentina has any close affinity
with them. The shape of wings reminds more of Cctrsia,
which, however, has quite different structure ; and the
strongly dentate, almost pectinate antenna is really not
very closely akin to the subdentate antenna of Ochyria
designata (Hufn.) and its allies.
A somewhat variable species, yet not .excessively, and
always y,»resenting a characteristic aspect. The median
area varies in width, and in the sharpness of its projection
at and M^ ; the subterminal is sometimes much less
distinct and less deeply dentate than in the type, and the
dark shading proximally to it is not always the same ; the
discal spots are sometimes large and distinct in all wings,
both above and below, sometimes ([uite minute.
92. Xanthorhoe suffocata (Dognin).
Xanthorhoe suffocata, Dognin, Ann. Soc. Ent. Belg., 1, 205
(1906).
Tucuman (type), in coll. Dognin.
93. Xanthorhoe edmondsii (Butler).
HypocJiroma edmondsii, Butler, Tr. Ent. Soc. Lond., 1882,
364.
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) S
258
Louis B. Prout on the
Mendoza (W. M. Bayne), three in coll. L. B. Prout; La
Rioja (E. Giacomelli).
Described from Chili, whence the British Museum has
a fair series, from dift'erent sources. Also occurs at Callao,
Peru (coll. Br. Mus.). I know of no other localities.
The larva has been described and figured by Izquierdo,
An. Univ. Chile, liii, 829, tab. iv, 3. It feeds on Boer-
haavia discolor, and reaches a length, when fully grown, of
27 mm. It has two varieties, one green, the other grey,
the latter being the less frequent. The green variety is
of a • pale, uniform citron-colour, the venter somewhat
paler ; head and legs whitish. The tubercles are small,
provided witli fine setae. Stigmata yellow, surrounded
by dark line. The grey variety is of an earthen colour,
or of that of the bark of a tree. It greatly resembles
the old sticks of the plant. The tubercles are more con¬
spicuous, usually margined with black. Ventrally there
is a pattern formed by some waved lines. The egg is
spherical, whitish, its surface smooth and shining. The
pupa is of a chestnut colour, the body darker than the
wings ; it is not enclosed in a cocoon, but the larva merely
joins the leaves together by a very few threads in order
to pupate. The pupal stage lasts 14 days.
Except in the pectinated ^ antenna, slightly broader
wings and lighter colour, this species reminds somewhat
of Cani'ptogramma (?) stellata (Guen.).
94. Eubolia meridionalts (Wallengren).
Euholia meridionalis, Wallengren, Wien Ent. Monats., iv,
176 (1860).
Patagonia, one $ (type) in the Stockholm Museum,
collectetl on the voyage of the “ Eugenie.”
I have been unable to make out this species from
Wallengren’s brief diagnosis, or even to guess at its
correct generic location. Its author says that its palpus
and wing- form remind of Euholia clicnopodiata (Linn.) =
mensuraria (Schiff.). Staudinger (Hamb. Magalh. Sum-
melr., ii, Lep. 84) surmises it is not from the Magellan
region.
95. Camptogramma (?) stellata (Guenee).
Scotosia stellata, Guenee, Spec. Gen. des Lep., x, 443
(1858).
259
Geometridac of the Argentine Repuhlic.
V. Scotosia haesitata, Guenee, Spec. Gen. des Lep., x, 444
(1858).
Phihalapteryx intestinata, pars, Walker, List Lep. Ins.
XXV, 1337 (1862) (nec Guenee).
Scotosia impauperata, Walker, List Lep. Ins., xxv, 1363
(1862).
Larentia halteolata, Herrich-SchaefFer, C.B. zool-min. Ver
Regensb., xxiv, 190 (1870).
Scotosia alhosignata, Packard, Fifth Peab. Rep., 61 (1873).
rhilereme alhosignata, Packard, Monogr. Geom. U.S., 174,
tab. ix, 14 (1876).
Phihalaptertyx intestinata, Druce, Biol. Cent.-Amer., Lep.
Het., ii, 156 (1893) (nec Guenee).
Gypsochroa haesitata, Hnlst, Tr. Amer. Ent. Soc., xxiii, 290
(1896).
Gamptogramma foedata, Warren, Nov. Zook, vii. 172
(1900) (nov. syn.).
Gypsochroa silellata, Hulst (ex err. pro stellata'), Dyar’s
List N. Amer. Lep., 286 (1902).
Tucuman, in coll. Dognin ; Gran Chaco, near Florenzia,
in coll. Br. Mus.
Very widely distributed and common from the southern
United States, the Bahamas and West Indies to Brazil,
Paraguay and Argentina. Varies very little except in size,
and I have not been able to discover any indication of
geographical variation. I suspect that Gnophos fuscovariata,
Wallengr., Wien. Ent. Monats., iv, 176 (1860) = Diselisio-
procta fuscovariata, Wallengr., Eugenies Resa, Ins., 379, tab.
vii, 6 (1861), is related to this, but from the figure and
description I cannot actually identify it. Some notes on
its structure, kindly sent me by Prof. Aurivillius,
confirm the suspicion that it belongs to the Larentiinae,
in spite of Wallengren’s erroneously placing it among
Boarmidae.
As I have not been able to satisfy myself concerning the
best generic position for the present species, I have allowed
it to retain provisionally that given to it by Warren and
the other workers at the South American fauna. In spite
of some resemblance to Philereme = Scotosia, it caniiot
remain there, on account of the non-biangulate discocel-
lulars of the hindwing; nor can it be referred to Ochyria,
Hiib. {Gypsochroa, Hulst, nec Hiibner), as it has not even
the true fasciculate $ antenna.
2 GO ilF Louis B. Prout on the
96. Camptogramma (?) euria (Prout), nov. sp.
9. 34 mm. Head, body and wings r. ddisb lirown, abdomen and
liindwing somewhat paler and greyer. Forewing very b'.oad, ape.x
acute, termen gently waved, slightly concave between ajiex and
middle ; traversed by numerous indistinct wavy lines, of a somewhat
olivaceous brown tint, the most distinct being three proximally and
three distally to a narrow, pale median band ; the pale band runs
from middle of costa to middle of inner margin, has an average width
of about 1 mm., is interrupted by a dark line on and widens
somewhat to inner margin ; the interspaces of the lines which bound
it are weakly darkened, so that there is a very faint suggestion of the
characteristic dark median band of the Larentiids, with pale centre ;
on inner margin near tornus, the beginnings of two further waved
lines are moderately distinct ; distal area slightly grey-shaded; ter¬
minal line continuous, brown, tinged with olive ; fringe divided by
a darkish line. Hindwing with angles well pronounced, distal
margin gently waved or subcrenulate ; greyish brown, with five feebly
expressed brownish wavy lines at approximately ecpial distances, the
innermost commencing at costa distally to the cell, but crossing its
oblique lower arm ; terminal line and fringe as in forewing.
Underside without markings, brown, densely irrorated with pale
scales.
Buenos Aires (H. Wilkinson), type in coll. Br. Mus.
I do not know any South American species with
which to compare this, unless possibly it be “ Cidaria”
ob.soletaria, Staml., Hamb. Magalh. Sammelr., ii, Lep. 102,
fig. 23, from Tierra del Fuego, with which it rather nearly
agrees in coloration ; but that has not quite such a broad
forewing, has the termen of the hindwing strongly cren-
ulate and the discoc^llulars biangulate, and is altogether
more of a Triphosa or Philcreme in appearance, so that
there is no probability of a really close relationship. The
European Larcnlia makata (Rbr.) also reminds a little of
the new species, but again only quite superficially ; the
more acute apex, the perhaps even broader forewing and
the venation of the hindwing are distinctive in this
comparison also.
According to imaginal characters, Camptogramma seems
hardly more than a subgenus of Euiihyia, with broader
wings, more crenulate margins, and some other very
minute distinctions. In C. (?) eiiria the face is rather
prominent, rough-scaled, the palpus moderate, rough-scaled,
tongue well developed, antenna laminate, shortly ciliated,
261
Geometridae of the Argentine BepMie.
hindleg with all spurs well developed, forewing with
double areole, hiinlwing with DC somewhat inangled,
from middle.
97. Orthonama obstipata (Fabricius).
Phataena olstipata, Fabricius, Ent. Syst., iii (2), 199
(1794).
^ Geometra Jlnviata, Hiibner, Samml. Eur. Schmett., v,
fig. 280, 281 (1796 ?).
7 Geometra gemmata, Hiibner, Samml. Eur. Schmett., v,
fig. 283 (1796 ?).
Camptogramma lapillata, Guenee, Spec. Gen. des Lep., x,
430 (1858).
Camptogramma haeeata, Guenee, Spec. Gen. des Lep., x,
430 (1858).
Camptogramma exagitata, Walker, List Lep. Ins., xxv, 1331
(1862).
Phihalapteryx intrusata, Walker, List Lep. Ins., xxv, 1339
(1862)!
Cido.ria peraciitata^ Walker, List Lep. Ins., xxv, 1421
(1862).
Coremia dbruptata, Walker, List Lop. Ins., xxvi, 1713
(1862).
Camptogramma signatarvi, Walker, List Lep. Ins., xxvi,
1718 (1862).
Coremia alternata, Walker, List Lep. Ins., xxxv, 1681
(1866).
Coremia pigrata, Walker, List Lep. Ins., xxxv, 1681
(1866).
Percnoptilota fiuviata, Hulst, Tr. Amer. Ent. Soc.,xxiii, 282
(1896). ‘
Plemyrio- mortuaria, Schaus, MS., fide Dyar, Proc. Ent. Soc.
Wash., X, 34 (1908).
Buenos Aires, fairly common (A. F. Bayne) ; Goya, in
coll. Br. Mus.
Range practically worldwide, with the exception of the
Arctic regions, Australia and New Zealand, and Chili. In
the last-named country it is replaced by a very close ally
Orthonama plemyrata (Feld.),* which I regard as specific¬
ally distinct, or at least a geographical race. Elsewhere I
cannot differentiate even geographical forms. The species
* Cidaria plemyrata, Feld., Reise Novara, Lep, Het., tab. cxxxii,
15,
2G2
Ml'. Louis B. Front on the
is suspected of being a migrant, and probably wanderers
from one locality, interbreeding with those which have
bred in jmother, serve to stamp out any attempts at local
variation.
It seems necessary to accejit Guenee’s suggested deter¬
mination of ohslipata, Fab., for this species ; vide Spec.
Gen. des Lep., x, 430, also Dyar, Froc. Ent. Soc. Wash., x,
34. I refer the species and its allies (including the ejfluata
group, etc.), to Orthonama, Hiib., Verz. bek. Schmett., 331
(type lignata, Hiib., Warr. seh). Warren (Nov. ZooL,
passim ; vide ii, 116) recognises two genera, Coenocalpe
and Orthonama, according to the build of the ^ antenna.
I doubt their validity in this sense, and hincy, from the
material I have placed under the microscope, that half-a-
dozen genera could be differentiated as easily as two. In
any case the name Coenocalpe is inapplicable, its type being
lapidata, Hiib., a species with biangulate discocellulars to
the hindwing. Ilorisme, Hiib. (type, tersata, Hiib.) differs
from tlie present group in the dorsal crests, etc.
98. Orthonama centrostrigaria (Wollaston).
Coremia centrostrigaria, Wollaston, Ann. Mag. Nat. Hist.
(3) i, 119 (1858).!
Phihalapteryx mediata, Walker, List Lep. Ins., xxv, 1338
(1862).2
rhihalajjteryx latirupta. Walker, List Lep. Ins., xxxv, 1684
(1866)>
Cidaria luscinata, Zell., Verb. z.-b. Ges. Wien, xxiii, 205
(1873).^
Cidaria interrwptata, Rebel, Ann. Hofmus. Wien, ix, 76
(1894).5
ricmyria paranensis, Schaus, Tr. Amer. Ent. Soc., xxvii,
273 (1901).6
Coenocalpe custodiata, Warren, Nov. Zook, xii, 442 (1905)
(in err., nec Guen.).'^
Buenos Aires, a I'ather worn which is clearly referable
to this widely-distribute.d species. In coll. L. B. Front,
presented by Mr. Fercy Richards.
Bange : Canaries (^) (^), Azores (^), the Atlantic States of
America (^) (^),* and as far south as Brazil (^) (*'). Frobably
Oi'curs pretty generally along the eastern coast of South
* Various localities recorded by Packard.
263
Geometridac of the Argentine Me-piiblic.
America, though I cannot speak positivel}’’ of this. I have a
specimen from Jamaica, and the British Museum another.
I published a note on the synonymy of this species a
couple of years ago (Entom., xli, 77), but bad not at
that time discovered that rnediata. Walk., belonged here.
Schaus, Proc. Zool. Soc., 1896, 646, refers it to jiuviata
but this is explained by the fact, which I learned from
Mr. E. Dukinfield Jones’ collection, that Mr. Schaus
formerly confoumied centrostrigaria and fturiata as sexes
of a single species.* I have now examined Walker’s type
of mediata.
99. Orthonama vittulata (Schaus).
Phihalapteryx vittulata, Schaus, Tr. Amer. Ent. Soc.
xxvii, 272 (1901).
? Orthonama dendlineata, Warren, Nov. Zool., xiv, 228
(1907).
Tucuman, in coll. Dognin.
Schaus’ type was from Bolivia. I have not seen it, but
I believe that Warren’s densilineata, from Peru, will prove
a synonym, or an aberration or variety, of the same.
100. Orthonama superjecta (Prout), nov. sp.
(Plate XLVIII, fig. 40.)
^ . .30 mm. Antenna nearly simple, minutely ciliated. Wings
with margins gently waved (in hindwing suhcrenulate), the terminal
line almost continuous (not broken at all into spots), only slightly
thickened between the veins and interrupted at the vein-ends.
Forewing grey, finely irrorated with brown, crossed by numerous
dark lines (mostly straight), and shaded with brown basally, and
in the median area with the exception of its central part ; edge of
basal area angled on SC, inner brown band of median area slightly
constricted on SC ; subterminal line nearly straight, very faintly
waved, dark shaded proximally ; discal spot distinct, black ; fringe
narrowly pale proximally and distally, more broadly dark medially.
Hind wing similarly, but weakly marked, slightly darkened distally
(especially at inner margin), with rather conspicuous pale sub¬
terminal line. Underside marked nearly like the upper, but much
more weakly.
* When this was written, I had overlooked Dr. Dyar’s note in
Proc. Ent. Soc. Wash., x, 34 (sub Hydriomena latir\tpta), which
confirms, from Mr. Schaus’ own collection, the deduction which I
made from Mr. Jones’.
264
MrTTjOuis B. Front on the
Buenos Aires, January 31, 1908 (A. F, Bnyne), type in
coll. L. B. Front.
General aspect of fittulata, Schaus, lyndilninda, Warr.,
Kov. Zool., xii, 834, etc., but distinct from all species
known to me. Differs in the subcrenulate margin, the
somewhat darker colour, arrangement of the lines distally
to the postmedian, and in the more unicolorous liindwing,
without a distinct postmedian. No oblique shade from
apex. Size somewhat larger than vittulata, Schau.s.
101. Ortiionama prouti (Bastelberger).
Coenocalpe prouti, Bastelberger, Ent. Wochenbl., xxiv, 207
(1907).
Tucuman (type), in coll. Bastelberger.
102. Fteroca'PHA umbrinata (Guenee).
Scotosia umbrinata, Guenee, Spec. Gen. des Lep., x, 443,
tab, X, 2 (1858).
Scotosia umbrinata ab. velutina, Oberthiir, Et. Ent., vii, 35,
tab. iii, 16 (1883) (ab.).
Tucuman, the type form and the dark-banded ab.
velutina, in coll. Dognin : I have also seen the former from
the same locality in coll. Bastelberger.
Widely distributed and common in South America.
Guenee’s types were from Brazil, Oberthiir’s from Feru.
103. Fterocypha floccosaria (Walker).
^ Scotosia floccosaria, Walker, List Lep. Ins., xxxv, 1685
(1866).!
? ^ Obila dispar. Walker, Char. Undescr. Lep., 51 (1869).
Pterocypha occidtaria, Schaus, MS., in coll. E. D. Jones.
Becorded by Druce, Biol. Centr.-Ainer., Lep. Het., ii,
156, as occurring in the Argentine Republic; no detail
given.
Walker’s type (^) was from Bogotfl. I suspect Obila
dispar. Walk., from Limas, is the $ of the same species.
Schaus (Froc. Zool. Soc., 1896, 646) records divulscda.
Walk., as such, but — according to the Walker type (or
cotype) at Oxford- — I do not feel able to accept this.
Ranges from Mexico to Faraguay and S.E. Brazil.
Geomdridae of the Argentine Rcptiblic. 265
104-. Pteeocypha SIMPLICIATA (Warren),
Pterocypha simpliciata, Warren, Nov. Zool., xii, 838
(1905).
Sunclia Corral, Santiago del Estero (type), in coll.
Rothschild.
105. Calocalpe affirm ata (Guenee).
Scotosia affirmata, Guenee. Spec. Gen. des Lep., x, 447
(1858).!
Scotosia proqj'cssata, Walker, List Lep. Ins., xxv, 1348
(1862).2 •
Scotosia duhiferata, Walker, List Lep. Ins., xxv, 1849
(1862).3
‘Scotosia nietagrainmata, Walker, List Lep. Ins., xxv, 1349
(1862).-^
Scotosia congoata. Walker, List Lep. Ins., xxv, 1351 (1862)
(ab.).
? Larentia suhccrtaria, Herrich-Schaeffer, C.B. zool. -min.
Ver. Regensb., xxiv, 189 (1870).
Ccdocalpe rosenhergi, Warren, Nov, Zool., vii, 172 (1900)
(nov. syn.).®
Ccdoccdpe furva, Warren, Nov. Zool., xii, 224 (1905) (nov.
syn.).6
Argentina (witliout exact locality), in coll. Br. Mus., a
single example, referable to the ab. congoata, Walk.
Distribution very general from Mexico southwards.
Q) Brazil, (^) Haiti, (^) Colombia, (^) Venezuela, (^) Ecuador,
(®) Paraguay. Probably the Cuban sicbceodaria, H.-S., is
the same.
Somewhat variable, especially in colour, which follows
somewhat nearly the same shades as in the European
Philercme transverscda (Hufn.) = rhamnata (Schiff.).
Walker’s congoata, erroneously supposed to come from
the Congo,* is the aberration with brighter brown, less
marked, submarginal area, the counterpart of an aberra¬
tion well known to students of Palaearctic Gcomctridac,
from its occurrence in Caloccdpc cerrincdis (Scop.) = ccrtata
(Hub.) and Philercme transverscda.
* Cf. Swiiilioe, Tr, Ent. Soc. Lond., 1904, 580.
206
Ml'. Louis B. Front on the
106. Calocalpe fimhriata (Front), nov. sp.
$ . 46 mm. Of about the size, shape and colour of the European
Calocal2)e cervinalis, Scoj). (= certata, Hub.), the crenulations of
the liindwing margin not quite so deep as in most of the allies.
Scheme of pattern as in the allies. Forewing with basal patch and
median band bounded by rather sharply defined blackish-fuscous
lines, subbasal line at 2^ mm., somewhat dentate, with a large pro¬
jecting s])ot on M ; antemedian from costa at 9 mm. to inner margin
at 6 mm., bent outward nearly to discal spot in cell ; postmedian
line from costa at 13 mm. to inner margin at about 9 mm., slightly
oblique outwards from costa to beyond SC^, parallel with termen to
between and IVP, then sinuate (or weakly angled) inwards,
finally making two rather strong projections outwards above and
below SM^ ; median band therefore rather narrow ; the pale band
which follows is traversed by a \veak line ; distal boundary line of
this pale band moderately distinct, and marked with some dark
vein-dots ; subterminal line (not thickening into a large tornal spot
as in the ajffirmata group) preceded by a conspicuous dark fuscous
baud, stronger and more regular than the dark shading in any
known form of affirmata and its allies, but interrupted on the veins ;
cell-spot distinct. Hindwing rather weakly marked ; postmedian
line and the succeeding one incficated from inner margin to about
R*, dark-dotted on the veins ; subterminal preceded by dark band,
though much less consjricuous than in forewing. Underside more
weakly marked, but with the cell-spots conspicuous (as in the
allies).
Cacheuta, end of September or beginning of October,
1904, taken by an inspector on the railway. Type in coll.
L. B. Front, presented by Mr. A. F. Bayne.
The specimen is unfortunately in poor condition. Mr.
Bayne tells me he has, or has seen, another example still
more damaged, but I have no further data concerning it.
The species agrees very closely in all points with typical
Calocalpe (= Eucosmia), but as the $ is unknown, I cannot
say positively that it may not belong, like cauqnenensis,
Butl., to the section (or genus) Trip)hosa, Steph., which
lacks the hair-tuft of hindwing beneath. The coarse of
the dark lines is not (piite the same as in affirmata,
Guen., and its small Chilian ally cxacta, Butl., Tr. Ent. 8oc.
Loud., 1882, 415, but its most striking distinction is in the
strong:, dark submarginal band. Otherwise it would seem
rather closely akin to them. I suspect that “ Larentia ”
Geomctridac of the Argentine Bepuhlie. 267
ehillanensis, Butl., Tr. Eiit. Soc. Lond., 1882, 395, is another
rather near relative, but it is certainly not conspecific.
107. Triphosa cauquenensis (Butler).
Scotosia eauquenensis, Butler, Tr. Ent. Soc. Bond., 1882,
415.
Puente del Inca, a fine series bred from pupae collected
under stones during the construction of the railway (W. M.
Bayne); also from Las Cuevas (W. M. Bayne). The first
batch of moths was bred in November 1903, others in
September and October 1905, and probably there are also
other dates which I have not expressly noted.
Hitherto only recorded from the mountains of
Cauquenes, Chili (Butler’s types).
The variation of the Puente del Inca and Las Cuevas
race can har dly be described as extreme, yet some examples
are of a somewhat warmer, browner tone than the others,
some have the transverse lines more strongly, others more
weakly expressed, while the breadth of the median area is
also liable to some variation. There are distinct indica¬
tions of a local race, differing from the Cauquenes types in
somewhat smaller size, warmer tone of colouring, and more
uniform markings — weaker differentiation of the median
band. In the whole of the long series which I have
examined, there is only one which I can call even nearly
typical. I therefore feel justified in describing the
Argentine form as Triphosa cauquenensis inca, Prout,
nov. var. (subsp. ?).
Mr. A. F. Bayne, who bred a large number from his
brother’s pupae, has kindly supplied me with a nice series,
besides presenting examples to the British Museum. He
further sent me some empty pupa-cases. In shape and
colour they greatly resemble that of T. dubitata, Linn.
The cremaster, however, shows distinct differences; in
dubitata this is more elongate, and the pair of spines are
fused for a considerable distance before diverging, i. e. may
be roughly compared to a letter Y with short arms; in
cauquenensis they are divergent from their base, as in the
letter V.
108. Perizoma impromissata (Walker).
Yqysipetes imptromissata, Walker, List Lep. Ins., xxiv, 1268
(1862).!
268
Mr. Li>nis B. Prout on the
Cam2otograriima corticeata, Walker, List Lep. Ins., xxvi, 1715
(1862^2
I'erizoraa fasciolata, Warren, Nov. Zook, iv, 454 (1807).^
Pcrizoma m uscosata, Warren, Nov. ZooL, vii, 180 (1900).^
? Pcrizoma ochritincta, Warren, Nov. Zool, xii, 336
(1905).^
YPonzomd] imellci, Tliieny-Mieg, MS., in coll. Dognin.
(•‘^) Goya, in some numbers (also from that locality in
coll. Br. Mus.) ; (■^) Parana, Entre Rios ; Buenos Aires
(A. F. Bayne), (also in coll. Br. Mus., from Berg — under
the name “ Glaueo'pteryx tegndatci” vide snjn'a — and from
H. Wilkinson); Mendoza (?) March 1905 (W. M. Bayne).
Geographical distribution: (^) (^) Montevideo ; (^) Para¬
guay (Warren’s type) ; (®) ? Huatuxco, Vera Cruz (Warren’s
type of ochritinda).
I liave very carefully compared Warren’s fasciolata with
Walker’s types, and though the latter are in damaged
condition there can be no doubt about their identification.
They happen to be slightly larger than the average, but
the British Museum also possesses specimens from Colon,
^Montevideo (Oldfield Thomas), which are typical in size.
As regards Warren’s type of ochritinda, its wings are, as
he says, somewhat narrower, or, as 1 have indicated in my
MS. notes, “ somewhat narrower than the average ” as com-
parecl with typical imgiromissata ; were it not for the
geographical difficulty, I should not hesitate to sink it.
Thanks to the generosity of Mr. A. F. Bayne, who has
allowed me absolutely unrestricted selection from among
the long series which he has collected, I have a picked lot
of thirty-seven Buenos Aires examples of this remarkably
variable species. A full analysis of their variation might
almost be made the subject of a separate i3aper; to a
British entomologist one may give some idea by saying
hat it is a veritable Dysstroma citrata (immanata) among
be Argentine Geomdridae. The commonest forms seem
to be the rather sombre ones with a moderately darkened
median band, or pretty nearly unicolorous ; not infrequently
the band is black, reminding of Xanthorhoe ferrugata ab.
unidentaria, especially when (as sometimes happens), the
paler areas are more or less coloured with ochre. Some¬
times tlie nearly unicolorous forms are strongly tinted with
mossy green (ab. muscosata, W^arr.), sometimes they are
nearly melanic. Very fre(piently the median band is more
269
Gcomctridae of the Argentine Rcpuhlie.
or less pale-centred, but this phase of variation gives rise
to an infinity of slightly ditferent forms ; the pale portion
may be broader or narrower, sometimes broken into large
costal and two small innermarginal spots, it may be only
slightly paler than the rest of the band, or even on the
other hand clear white — the most striking forms — or again
it may be tinged with flesh-colour or almost reddish,
slightly suggesting the type form of X. ferrugata, or with
a somewhat more ochreous shade. In one or two speci¬
mens the areas placed proximally and distally to the median
band are quite reddish. My single Mendoza specimen is
of the ab. muscosata.
Mr. Bayne tells me that this is the only Geometrid
common in the winter in Buenos Aires. I find the dates
range from March 31 to September 25, It is interesting
that the only three species for which I have noted winter
records from there — Hamviaptera tencra, Ochyria argentina
and the present species — all belong to the Larentiinae, the
sub-family which is so preponderant in Arctic and mountain
regions, so comparatively deficient in representatives in
tropical climes.
109. Perizoma sordescens (Dognin).
Perizoma sordescens, Dognin, Ann. Soc. Ent. Belg., lii, 18
(1908).
La Paz, Mendoza, September 18, 1903, one $ (VV. M.
Bayne); Paltnira, one ^ (W. M. Bayne); both in coll.
L. B. Prout; San Rafael, one in coll. A. F. Bayne.
Dognin’s type was from Peru ; I have compared my
specimens therewith.
A somewhat variable species in colour and in intensity
of markings, but the median band seems constant in shape.
My? is darker than my I have a worn specimen from
Palmira, September 13, 1903, which is apparently a close
relative, or likely even an aberration, of F. sordescens, but
I cannot decide definitely until similar specimens are met
with in better condition. It appears to have the median
band entirely absent — an extreme of variation not unknown
in some of the most variable of the Larentiinae {e.g. Perizoma
niveata, Steph. = albidata, Schiff., Oporinia dilutata, Schitf.,
etc.); but it is harder to reconcile the position of a rather
distinct subbasal band with the markings of P. sordescens ;
on the other hand, the black subapical marks are typical
270
B. Front on the
for tliose of that species. I forbear to name the form at
present. Its captor, Mr. W. ]\I. Bayne, obtained eggs
from it, but the larvae died before a food-plant could be
discovered.
From Maassen’s description and figure (Stiibel’s Reisen,
Lep., 1G8, tab. ix, 15) I should not be at all surprised if
Cidaria aneiuliferata, Maassen, from Bolivia, were another
form of this species, in which case his name would have
priority.
The build of the abdomen, which in the San Rafael $
is, in death, somewhat upcurved anally, and the presence
of an oblique apical dash on forewing, suggest that this
species may have some affinity with the genera Di/sstroma,
Eustroma, Lygris, etc., of the Palaearctic Region — the
genus Cidaria (pro parte) of Guenee’s .system. It is
hardly likely it will prove to bear any close larval re¬
lationship to the seed-feeding Perizoma (= Emmelesia) of
Europe.
110. Perizoma (?) iduna (Prout), nov. sp.
$ . 25 mm. Face slii'htly tufted, palpus rather long, antenna
almost simple, pultescent ; frenulum apparently wanting or very
weak ; discocellulars of hindwing rather weakly biangulate. Fore-
wing with basal and median areas brown, strongly tinged with pink,
traversed by very ill-defined, more fuscous brown waved lines ;
proximal edge of median band ill-defined, its limiting line can just
be made out to be strongly curved ; postmedian line (distal margin
of median band) from about two-thirds costa, dentate and slightly
oblique outwards to below then inbent to Mq where it projects a
sharp angle into median band, then dentate and nearly perpendicular
to inner margin ; distal area pale violet grey, almost whitish, but with
a large apical blotch (to below R") of varied pink, ferruginous and
fuscous, and a small, ill-defined tornal mark. Hindwing uniform
ferruginous brownish. Underside of forewing olivaceous-grey in cell
and submedian area, delicate pink in apical portion and at extreme
margins, the colours not shar])ly defined ; discal spot and postmedian
line well marked in dark olivaceous brown. Underside of hindwing
delicate pink, slightly shaded with olive-grey basally, a large dark
discal dot, and distinct, curved, somewhat diffuse and wavy post¬
median line.
S. Pablo, 1200 in., Villa Novoes (?), Tucuman, September
3, 1901, type in coll. Br. Mus.
Geometridae oj the Argentine Uejiuhlic.
271
111. Dysstroma citrata (Linne)(?).
Phalaena {Geomeira) citrata, Linne, Fauo. Suec., ed. 2, 332
(1761).
Phalaena immanata, Haworth, Lep. Brit. (2), 323 (1809).
Poli/phasia tncncata rufihrunnea, Warren, Nov. Zool., vii,
181 (1900).
“ Parana, Entre Rios,” in coll. Rothschild (types of
Warren’s truncata rujihritnnea).
I have very fully discussed the synonymy of this widely
distributed Holarctic species in Trans. City Bond. Ent.
Soc., xviii, 33-60, together with that of its congener
Dysstroma truncata, Hufn., to which Mr. Warren referred
his rufihrunnea. And I there expressed the doubt, which
I still entertain, whether the supposed Parana record were
not founded on some error in labelling, or — unlikely though
this latter explanation seem — on some European material
accidentally imported into Argentina. “ Not only the
species, but the entire group is absent from the vast
intervening area of tropical America, and the specimens
are so normal in appearance as would be almost incredible
on the assumption of an ancient geographical isolation ”
{loc. cit., p. 51).
112. Hypolepis prionogramma (Prout), nov. sp.
(Plate XLVIII, fig. 19.)
. 24 mm. Face and palpus light brown. Antenna brown, the
.scales of dorsal surface in alternate bands of brown and straw-colour.
Forewing whitish, dusted and clouded with brown, and with a
decided tinge of straw-colour, especially in basal and costal areas
and on median vein ; the markings brown ; basal line from inner
margin, outangled, bounded distally with white, but lost in the costal
clouding after vein M ; an irregular subbasal band, its distal margin
projecting at SM^ and more broadly below M, the band constricted
in cell, widening again costad ; median band much as in H. serratilinea
(Warr.),* containing distinct white spot at lower angle of cell ; sub¬
terminal line weakly indicated, but with distinct dark patches
pro.ximally to it at costa and (smaller) at inner margin, some vague
dark clouding between ; termen clouded with darkish brown, the
most prominent patch being a large irregular triangle between costa
and its acute apex placed on ^t about 2h mm. from termen ;
* PsaUodes serratilinea, Warr., Nov. Zook, xi, 70 = Hypolepis
serratilinea, Warr., ibid., 521.
272
AiPf Louis B. Prout on the
fringe pale straw-colour, cliequerecl with darkish brown. Hindwing
whitish, tinged with straw-colour distally ; weak beginnings of two
lines on inner margin ; the hair-scales of the inner-marginal area
somewhat browner. Underside of forewing brownish, weakly
marked, the costa broadly yellower and mottled with brown ; a
brown blotch close to distal margin, between and R^ and
(narrower and weaker) between SC^ and IR ; fringe as above.
Underside of hind wing with rather large, distinct dark cell-spots,
a zigzag dark postmedian line running into a large dark spot near
inner margin at about two-fifths, a much weaker dark mark near
inner margin between this and tornus.
Tucuman (type), iu coll. L. B. Prout ; also in coll. Dognin
from same locality.
Similar to H. serrutilinea, Warr., from Peru (under which
name M. Dognin records it in his MS. list), but dis-
tinguished, apart from its somewhat different tone of
colour, by the form of the basal and subbasal dark markings,
and, to a less extent, those of the distal margin. In the
former respect, indeed (and in the presence of the white
mark in the centre of the median band), it comes nearer to
IL completa, Warr., Nov. Zook, xi, 520, also from Peru.
The antenna is quite normal, rather thick — as in all the
males of the group, so far as my observation goes — but
neither “rasped above” nor with “short, subclavate teeth •
below.” According to Warren, Nov. ZooL, xi, 521, H.
serratilinca ^ should be abnormal iu these particulars; but
as I do not feel altogether satisfied that this $ was con-
specific with the original (^) type, I do not emphasise the
antennal characters as a difierential feature of H. priono-
gramma. In the new species the discocellulars are some¬
what bent on loth wings, weakly approaching the
“biangulate” form, and with from slightly below the
middle ; yet certainly not of the typical Fsaliodes form.
The hair-tuft on abdomen beneath is strongly developed.
113. Spargania flavisquamata (Dognin).
Eriop)ygidict fiavisquamata, Dognin, Ann. Soc. Ent. Belg.,
xlviii, 360 (1904).
Tucuman (type), in coll. Dognin, also from the same
locality in coll. Bastelberger et coll. Br. Mus.
I consider the genus Eriopygidia, Warr., Nov. Zook, vii,
175, inseparable from Sjmrgania, Guen., Spec. Gen. des
Lep., X, 454.
Geometridae of the Argentine Reggiiblic.
278
114. Larentia profugaria (Herrich-Scliaeffer).
Larentia profugaria, Herrich-Schaeffer, Samml. Aussereur.
Scbmett., i, fig. 410-11 (1855).
Aspilates (?) hemichlorcda, Walker, List Lep. Ins., xxiv,
1069 (1862).
Larentia omphacina, Dognin, Ann. Soc. Ent. Belg.,
xlv, 179 (1901).
Cidaria anteviridata, Thierry-Mieg, Le Nat., xxix, 259
(1907).
Tucuman, in coll. Bastelberger.
Tbe species is cbietly known as Brazilian. Tbence
came tbe types of hemiehlorafa, anteviridata, omphacina and
pvohahXy tbongb by an evident mistake Herricb-
Scbaeffer recorded it as from “ North America.” It was
partly tins error, no doubt, wbicb misled Dognin into
redescribing tbe South American insect as omphacina.
There is, however, a good deal of variation in tbe species,
though not such as to I'aise any doubts in my mind as to
tbe specific identity of tbe different forms.
The generic position of profugaria is a matter of some
perplexity, as it seems to present no salient features for
generic differentiation and yet tbe facies suggests that it
should be sni generis. Of course it is only a “ Ijarentia ”
in tbe comprehensive sense in which that name is used by
Hampson — areole double, discocellulars of hind wing bian-
gulate. The suggestian was made tentatively by Packard
(Monogr., p. 212), merely from Herricb-Schaeffer’s figure,
that it might belong to Chloraspilates, and this has been
followed, without even a query, by Hulst in Dyar’s List N.
Amer. Lep., 805 ; but it is of course altogether erroneous.
115. Sarracena xanthochlorata (Walker).
Sgharites xanthochlorata, Walker, List Lep. Ins., xxiv, 1427
(1862).
Tucuman, in coll. Dognin.
Walker’s type was from Venezuela. It is believed that
this form, in spite of tlie very differently coloured hindwing,
is merely the ^ to Sarracena chlamydaria, H.-S., Samml.
Aussereur. Scbmett., i, fig. 73 = Sarracena hrevilinea, Warr.,
Nov. Zook, xi, 524. But as there is no certainty about
this at present, I have accepted M. Dognin’s record, which
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) T
274 I'fff Louis B. Prout on the
will show wliat form (and sex) has as yet been observed in
Argentina.
116. Stenorrhoe longipenxis (Warren).
Stenorrhoe loiKjipennis, Warren, Nov. Zool., vii, 183 (1900).
Goya, Corrientes, in coll. Br. ^lus.
The species was described from S.E. Brazil. I know of
no other localities.
Eueupithecia (Berg, MS.) (Prout), nov. gen.
Face Hat, not tufted ; jJ^dpus very short, porrect, 2nd joint rough-
scaled, 3rd joint small ; tongue short ; antenna in ^ with strong
fascicles of cilia, a longer and a shorter pair of fascicles to each joint,
in 9 very shortly ciliated. Thorax glabrous beneath ; femora
glabrous ; hindleg short, hindtibia in ^ with hair-pencil, all spurs
wanting, in 9 'vith terminal spurs only. Abdomen long. Wings
long and narrow, smooth-scaled ; frenulum developed. Forewing
with cell long ; SC' from about two-thirds of cell ; SC‘^ out of SC'i
anastomo.sing with SC-'' to form a long, simple areole ; SC'*® from
welPbefore apex of cell ; radials normal ; M' a])proximated to ;
M” approximated to M', noticeably curved. Hindwing with cell
rather long ; C anastomosing with SC to beyond one-half of cell ;
SC‘^ stalked with R' ; R^ from much a1)ove middle of cell ; medians
nearly as in forewiug, M- less curved.
Type of tlie genus : Eueupithecia cis2)latcnsis, Prout,
nov. sp.
According to my invariable custom, I liave adopted an
available MS. name in preference to coining a new one ;
but it must be admitted that Berg’s name is by no means
elegantly formed.
This interesting genus bears much superficial resemblance
to some of the narrower-winged species of Eu]}'ithecia and
Encymatoge, except in its more elongate abdomen ; yet its
smooth, flatly -rounded face, minute palpus and aborted ^
hindleg, all recalling the Acidaliinae, remove it rather
widely therefrom. It is perhaps related to Stenorrhoe,
Warr., Nov. Zool., vii, 183, which shares with it the frontal
and hindtibial structure (I know only one 9 of Stenorrhoe,
with the legs lost), but which differs in having the areole
double, and in some other points.
Geometridae of the Argentine Re'puUic.
275
117. Eueupithecia cisplatensis (Berg, MS.)
(Prout), uov. sp. (Plate XLVIII, fig. 5.)
5 • 21-23 mm. Head, body and legs nearly uniform light grey,
tinged with brown. Forewing with antemedian and iiostinedian
lines scarcely indicated except by dark vein-dots, the former at about
one-third, bent outwards in cell, the latter at about two-thirds,
angled on R* ; cell-spot large, dark, roundish, the faintest possible
indications of a darkened median shade ; terinen with some rather
indistinct dark dots between the veins. Hindwing with no distinct
markings excepting a large, dark, roundish cell-spot and dark
terminal dots ; in addition, very weakly banded with alternately
greyer and browner shades or lines, which are sometimes slightly
better expressed at inner margin. Underside similar, but with
antemedian line obsolete on forewing, on the other hand with a
distinct postmedian line of small dark dots, angled on Rh
Buenos Aires, latter half of January, in coll. L. B. Prout
et coll. Br. Mus., the type ($) January JO, 1903, in coll.
L. B. Prout ; Gran Chaco, near Florenzia, October 1902,
in coll. Br. Mus.
I know of no other localities. Specimens were sent
from Buenos Aires to the British Museum by Prof. C. Berg,
as long ago as 1886, under the manuscript name of
Eueiipithecia cisplatensis \ but they have never yet been
described.
118. Sebastia maleformata (Warren).
Sebastia maleformata, W^arren, Nov. Zook, ii, 112 (1895).
Buenos Aires, one J, December 21, 1903 (A. F. Bayne),
in coll. L. B. Prout.
Warren’s type was from S.E. Brazil.
119. Eupithecia batida (Dognin) (?).
Eupithecia batida, Dognin, Ann. Soc. Ent. Belg., xliii, 140
(1899).
Buenos Aires, one January 5, 1907 (A. F. Bayne),
in coll. L, B. Prout.
My specimen is somewhat worn, and although it seems
to agree with Dugnin’s batida (from Colombia), I cannot be
certain of the identification, upon such slight material. It
is not impossible it belongs to the subgenus Stenopla,
Warn, Nov. Zool., vii, 166 ; it reminds of the type-species
T 2
27G ]\I^Louis B. Prout on the
of that genus, S. mrescens, Warr. {ibid.), of which 1 know
only the
120. Eupithecia semilugens (Dognin).
Tephroclystia semilugens, Dognin, Ann. Soc. Ent. Belg.,
1, 209 (1906).
Tucuman (types), in coll. Dognin.
121. Eupithecia chincha (Dognin).
Eupithecia chincha, Tfogwm, Ann. Soc. Ent. Belg., xliii, 138
(1899.)
Tucuman, in coll. Dognin.
Dognin’s types were from Ecuador.
122. Eupithecia furvipennis (Dognin).
Tephroclystia furvipennis, Dognin, Ann. Soc. Ent. Belg.,
1, 209 (1906).
Tucuman (type), in coll. Dognin.
123. Eupithecia indefinita (Snellen).
Eupithecia indefinita, Snellen, Tijd. Ent., xvii, 75, tab. v, 11
(1874).
Tucuman, in coll. Bastelberger.
Described by Snellen from Bogota.
124. Eupithecia bellimargo (Bastelberger).
Tephroclystia bellimargo, Bastelberger, Intern. Ent. Zeit.
Guben, i, 255 (1907).
Tucuman (type), in coll. Bastelberger.
125. Eupithecia i-actevirens (Dognin).
Tephroclystia lactevirens, Dognin, Ann. Soc. Ent. Belg., lii,
21 (1908).
Metan, Salta (type), in coll. Dognin.
126. Eupithecia truncatipennis (Warren) (?).
Tephroclystia truncatig)cnnis, Warren, Nov. Zooh, iv, 451
(1897).
Buenos Aires, one January 27, 1902 (A. F. Bayne),
in coll. L. B. Prout.
Geometridae of the Argentine Republic. 277
The species was described from S.E, Brazil. My
specimen is too worn to admit of any confidence in the
determination, but from its shape and general aspect, I
judge it likely to belong here. Many of the Buenos Aires
species occur also in S.E. Brazil.
127. Eupithecia argica (^Prout), nov, sp.
(Plate XLVIII, fig. 16.)
17 mm. Antennal ciliation minute. Face, palpus and body-
above concolorous with forewing ; thorax paler beneath. Forewing
pale brown-grey, irrorated with fuscous, and traversed by fuscous
lines ; three approximately equidistant lines proximally to discal
spot, rather distinct costally, angled in cell, and becoming more or
less indistinct or obliterated posteriorly ; cell-spot rather distinct,
seated on an acutely angulated (not very distinct) fuscous line, which
runs from costa near thirddine to M well before middle of wing ; post¬
median from costa at nearly two-thirds, accompanied by a dark
wedge-mark proximally at SC^ then oblique distad, rectangularly
bent at thence straight (and parallel with termen) to inner margin;
a very slightly paler band follows, bounded and intersected by two
very weak lines, which follow the same course as the postmedian ;
subterminal line lunulate-dentate, indistinct, shown only by the dark
dusting of the rest of the distal area ; a dark mark on submedian fold
proximally to the subterniinal, as in many neotropical species of the
genus ; marginal line dark, interrupted at the vein-ends. Hindwing
much paler, only the inner- marginal area concolorous with forewing,
and showing weak indications of beginnings of dark transverse lines ;
cell-spot scarcely indicated ; marginal line very weak. Forewing
beneath paler than above, somewhat darkened in costal and distal
areas ; proximal area without definite markings ; discal spot rather
large, distinct ; a curved (not angled) somewhat diffuse postmedian
line, weaker in lower half of wing ; a somewhat pale band following,
and bounded by a curved (not angled) line, to which succeeds the
darkened terminal area with rather indistinct subterminal line, as
above. Hind wing beneath pale, with faint suggestion of a subangulate
antemedian line ; cell-spot rather large, distinct ; postmedian line
fuscous-brown, distinct, strongly curved behind cell-spot ; a similar
line midway between this and distal margin ; marginal line distinct.
Temperlay, Buenos Aires, February 1901 (A. F. Bayne).
Type in coll. L. B. Prout.
An inconspicuous little species, of somewhat the tone
of colour of an average form of the European E. vulgata
278
Mr. Louis B. Prout on the
(Haw.), or intermediate between that and E. e.riguata
(Hiib.). In such a large and difficult genus, it is impossible
to say positively tliat it has not been already described, but
I have studied all Warren’s and Dognin’s types — which
cover by far the greater pro])ortion of those de.scribed —
without finding it, and think it therefore fairly safe to
publish it as new. The specimen I have determined as
truncatipennis, Warr., may be a malformed example of it,
but each looks normal to its own shape. The type of
argica is in good condition.
128. Eupithecia ammorrhoa (Prout), nov. sp.
(Plate XLVIII, fig. 12.)
9. 21 nun. Head, body and torewing light sandy -grey, irrorated
with darker sandy-ochreous ; thorax and base of abdomen, as well
as forewing, paler beneath. Antenna sandy, darker ringed. Wings
long and narrow. Forewing crossed by several sandy-ochreons lines,
the most distinct being : a curved antemedian from beyond two-fifths
of costa to middle of inner mai’gin ; a postmedian from about two-
thirds of costa, angled distad above (but here weak), distinct again,
and parallel with termen, from R*, slightly inbent from R^ to kP ;
and a finer line accompanying the postmedian distally at about 1 mm.,
following the same course, darkened from costa to the angulation on
SC’, and crossed by a short blackish mark on M" ; termen narrowly
clouded with sandy-ochreous, more broadly at apex ; cell-spot black,
distinct ; costal area somewhat dotted and striated with fuscous ;
marginal line blackish, interrupted at vein-ends ; fringe at base
narrowly white, then with a bar of light fuscous, then again light,
though somewhat mixed with pale fuscous. Hindwing whitish,
with a dark line almost parallel with and near to termen, distinct
from inner margin, but becoming faint before one-half and dis¬
appearing before costa ; the weak beginnings of other dark mark¬
ings from inner margin to M and LP, this area also somewhat less
white than rest of wing and dotted with fuscous ; cell-spot minute ;
marginal line and fringe as in forewing. Underside of forewing
very weakly marked, excepting a moderately dark cell-spot (which
is more elongate than above), and the fine line distally to the post¬
median, which is rather conspicuous in its darkened costal part, and
well traceable across the wing ; underside of hindwing nearly white,
almost unmarked, excepting a small cell -spot.
San Juan, November 15, 1904 (W. M. Bayne), type in
coll. L. B. Prout.
Geomctridae of the Argentine Republic.
279
The shape and the course of markings somewhat recall
Eupithecia sihylla, Butl., Tr. Ent. Soc. Lond., 1882, 405,
but the colour is entirely different. Except that the bases
of the forewings are somewhat rubbed (probably in setting)
the specimen is in beautiful condition. Besides the mark¬
ings enumerated, there are evidently some further lines in
the basal area, while the median area contains two faint,
angulated lines, the proximal of which crosses the cell-spot
and joins the antemedian at about the middle of the wing.
In structure, the species is a thoroughly typical Eupithecia,
the face perhaps somewhat more sloping (prominent below)
than in some species, the palpus rather stout and rough-
scaled, though not so extreme as is sometimes the case in
the genus.
Egg yellow, oval, of the usual Eupithecia form (W. M.
Bayne, in litt.).
Note. — I have also from Mr. A. F. Bayne a rather worn
$ of apparently another species of Eupithecia, taken in
Buenos Aires on January 16, 1905. I have thus far failed
to identify it, and although it should be quite recognizable,
it is not sufficiently good to justify its being made the
type of a new species, even should it prove to be hitherto
undescribed.
129. Amaurinia glauculata (Walker).
Thalassodes glauculata. Walker, List Lep. Ins., xxvi, 1560
(1862).
Tucuman, in coll. Dognin.
Walker’s type was from Brazil. I believe the same form
occurs in Paraguay, but its actual range yet remains to be
worked out.
180. Amaurinia coerulea (Warren).
Amaurinia coerulea, Warren, Nov. Zoo!., xii, 51 (1905).
Tucuman ( Warren’s type and others), in coll. Rothschild;
also one example, smaller and more unicolorous, from same
locality, in coll. Baste Iberger.
It is not unlikely that this will sink before some older
name, e. g. peruviensis, Schaus, Tr. Amer. Ent. Soc., xxvii,
262, but the species, or forms, run very close, and I have
not yet been able to examine sufficient material to justify
any definite conclusions. I should not be at all surprised
280
Af^Louis B. Prout on the
if the species recorded by M, Dognia as glauculata (vide
supra) proved to be the same as this.
131. Cambogia tegularia (Guenee) (?).
Cambogia tegularia, Guenee, Spec. Gen. des Lep., ix, 433
(1858).
Tucuman, in coll. Bastelberger ; Metan, Salta, in coll.
Dognin.
I am in some uncertainty which species is the true
tegidaria of Guenee, His types were from Brazil, and
will probably prove conspecific with the Argentine insect ;
but a considerable mixture of forms, from different parts
of Central and South America, stands in coll. Br. Mus.
under this name. Mr. WaiTen has named the Metan
specimens in coll. Dognin, as “ odatis, Druce ? ”, but I find
this is certainly not the odatis of Druce, Biol. Centr.-Amer.,
Lep. Het., ii, 113. Mr. Schaus, in coll. E. D. Jones, calls
it (the Metan species), or something extremely like it,
tegidaria, Guen.
Sub-family BOARMIINAE.
Fidoniinae, Onrapteryginae, Deiliniinae, Nepliodiinae,
Abraxinae, Semiothisinae, Selidoseminae, Ennominae,
Frosopolophinae, Seotopiteryginae, Ascotinae, et Bistoni-
nae, Warren.
Narkagodes (Warren),
Narragodes, Warren, Nov. Zool., vii, 200 (1900).
This interesting little genus, even if Narragodes laevis,
Warr., Pi •oc. U.S. Mus., XXX, 524 (Peru), and another un¬
described species from Peru really belong to it, evidently
has its head-quarters in S.E. Brazil and the Argentine
Republic. Four species, including the one here described,
are now known from these countries, and I believe some
unnamed Brazilian material in the British Museum col¬
lection will furnish one or two others. Eupileta, Warn,
Nov. Zool. xii, 362, also scarcely differs, so far as I can
see, except in secondary $ characters. Apart from Eupileta
and the doubtful Peruvian species, the genus divides
readily into two sections ; —
A. Palpus short, antennal pectinations moderate, SC^
Geometridae of the Argentine BejjvMic. 281
of forewing anastomosing strongly with C, well separate
from {fuscata, Warr. ; comminuta, Dogn.).
B. Palpus strong, ^ antennal pectinations long, SC^ of
forewing anastomosing at a point only with C, and (at
least mgyda) later at a point with {psychidia, Warr. ;
gyda, Prout).
Both sections agree in general build and scaling, in the
smooth face, dilated ^ hindtibia, strong anastomosis of C
of hindwing with SC, absence of SC^ of forewing, etc,
132. Narragodes comminuta (Dognin).
Marragodes (in err. pro Narragodes) comminuta, Dognin,
Ann. Soc. Eut. Belg., 1, 211 (1906).
Tucuman (type and others), in coll. Dognin.
I possess a ^ from Central Bolivia ; I know of no other
localities.
138. Narragodes psychidia (Warren).
Narragodes psychidia, Warren, Nov. ZooL, viii, 480 (1901).
Parana, Entre Bios (type), in coll. Rothschild ; Tucuman,
in coll. Dognin.
134. Narragodes gyda, Prout, nov. sp.
(Plate XL VIII, %. 15.)
(J. 15-16 mra. Fuscous, finely irroratecl with blackish fuscous,
and with the markings blackish fuscous. Forewing with subbasal
line at 1 mm., not very conspicuous, broadly dark shaded proximally,
at least in costal half of wing ; antemedian from before one- third
costa, strongly bent outward in cell and slightly so towards inner
margin, reaching inner margin at about two-fifths ; postmedian from
costa at just beyond two-thirds, sharply outangled at R^, sinuate
inwards distally to cell, and more deeply in submedian area, again
outbent on SM^, and reaching inner margin at about three-quarters ;
a central shade from costa at scarcely beyond one-half, somewhat
oblique outwards to cell-spot, then very strongly oblique inwards,
running quite near to antemedian line in the inner-marginal half of
wing ; cell-spot moderately large ; subterminal line only indicated
by some indistinct, interrupted dark shading proximally to it ; some
very indistinct marginal spots between the veins ; fringe very in¬
distinctly chequered, the darker and paler shades very little con¬
trasted. Hindwing without markings, a small dark cell-spot barely
indicated. Underside of both wings with cell-spot and indistinct
postmedian line, that of the forewing distinct at costa.
282
Louis B. Front on the
San Luis, December 23, 1905 (type), in coll. L. B. Front;
Meudoza, February 13, 1905, one ^ in coll. L. B, Front ;
La Faz, Mendoza, November 24 and December 22, 1905
two ^s in coll. A. F. Bayne (all these four collected by
W. M. Bayne) ; ? Gran Chaco, near Florenzia, October
1902 (S. R. Wagner), in coll. Br. Mus.
Nearly related to N. j^sychidia, Warr., with which it
agrees in structure. It is of a less brown colour (psychidin
might relatively be described as tinged with reddish) and
further differs in the deep inangnlation of the antemedian
line, the stronger expression of the median and postmedian,
etc. The Mendoza specimen is less strongly dark-irrorated
than the San Luis (type), hence its dark lines stand out yet
more clearly, removing it further in aspect from Warren’s
species. The Gran Chaco specimen is not precisely like
the western examples, but I think I am right in referring
it to this species as a dark variety; yet the tongue and
palpus seem slightly more yellow-tinged, the course of the
antemedian line does not look precisely the same, and
future study may possibly establish another closely-related
species.
135. Fsodopsis TORTILINEA (Frout), uov. sp.
(Flate XLVIII, fig. 13.)
^ 5-20 mm. Head, body and wings greyish fuscous, more or
less irrorated with whitish. Forewing with the markings dark
fuscous, only the broad median line conspicuous ; antemedian line
from al)Out one-fifth costa, throwing a very strong projection distad
in cell, and obtusely angled on submedian fold ; median line (shade)
from one-half costa to one-half inner margin, strongly outbent at
and making a sinus inwards between I\P and SM^ ; postmedian line
ill defined except in lower half, commencing beyond two-thirds of
costa, dentate, thickened and strongly sinuate inwards between kP
and SM-, narrowly i ale-margined distal ly ; marginal line slightly
darker than ground-colour ; fringe rather pale, weakly chequered
with darker. Hindwing unicolorous, somewhat paler than forewing.
Underside without markings.
Aristides Villanueva, September 25, 1905 (W. M. Bayne),
type in coll. L. B. Front ; Balde (W. M. Bayne), a ^ in
coll. A. F. Bayne.
I am indebted to Mr. Warren for the generic location
of this obscure little species, as I have not yet had the
opportunity of examining his genus Fsodopsis (Nov. Zook,
Geometridae of the Argentine Re'pvhlic. 283
xi, 560) structurally. It agrees in the small palpus, in the
strong anastomosis of C of hindwing with SC, and in the
general build. The face is slightly prominent, flattened,
smooth-scaled, the antenna in the ^ subdentate, with thin
fascicles of moderate cilia, in the ^ shortly ciliated ; SC^'^
are coincident, not stalked, and anastomose or are connected
with to form a rather long narrow areole ; the costal
area of hindwing is ample, and C, after anastomosis, diverges
from cell even more sharply than in Narragodes, to which
the genus is apparently related.
136. Mimophyle noctuata (Warren).
Mimophyle nochiata, Warren, Nov. ZooL, viii, 480 (1901).
Parana, Entre Rios (type), in coll. Rothschild ; Retamito,
one November 28, 1903 (W. M. Bayne), in coll. L. B.
Front; Palmira, one March 13, 1905 (W. M. Bayne), in
coll. L. B. Prout.
The ^ is still unknown. The three known ^s show no
appreciable variation.
137. Mimophyle sabulosa (Prout), nov. sp.
(Plate XLVIII, fig. 36.)
d' 22-28 Him. Face prominent, rounded, smooth-scaled ; build
more slender than in M. ')ioctnatn, Warr. ; wings elongate, of delicate
texture, hindwing with anal angle rounded. Head and antenna
pale greyish fuscous; thorax and forewing pale greyi.sh fuscous
above, paler beneath ; abdomen and hindwing dirty white, sparsely
irrorated with fuscous, especially the di.stal half of hindwing above.
Forewing with a dark discal spot and ill-defined, zigzag antemedian
and postmedian lines ; antemedian from about one-third costa,
strongly angled in cell, scarcely traceable below middle of wing ;
postmedian from fourth-fifths costa, throwing out a sharp, deep tooth
into median area between SC“ and R*, a broader projection into
median area between R* and R^, and another in submedian area,
but with a tendency to produce distad-directed teeth on all the
veins ; a dark mark on inner margin before one-fourth, probably
indicating a remnant of a subbasal line ; marginal area somewhat
paler than the rest of wing, especially just distally to the postmedian
line ; a row of dark marginal dots between the veins. Hindwing
with terinen subcrenulate, marginal dots as in forewing; otherwise
almost without markings, the cell-spot small and faint. Under¬
surface almost without markings ; each wing with a very small,
faint cell-spot ; marginal spots weaker than above, more inclined to
284
ouis B. Front on the
be prolonged into a line interrupted by the vein-ends; forewing in
addition with the costal extremity of the postmedian line weakly
indicated.
Retamito, November 28, 1903 (W. M. Bayne), type in
coll. L. B. Front,; La Faz, Mendoza, Angnst 25, 1905 (W.
M. Bayne), one ^ in coll. L. B. Front ; Falinira, September
13, 1903 (W. M. Bayne), one $ in coll. A. F. Bayne ;
Coronel Salas (W. M. Bayne), one ^ in coll. A. F. Bayne.
The ^ is still unknown. The species is probably vari¬
able, both in size and in depth of colour. My La Faz
besides being considerably larger than the type, is more
heavily dark-speckled, particularly in the distal area. The
shape of the wings, and the peculiar conrse of the jiost-
median, gives a very characteristic and unmistakable asjaect.
Whether sabnlosa can be allowed to remain permanently
in Mimophyle must depend on the degree of weight
attached to the frons (not protuberant in oioctuata), the
shape and textnre of the wings, or other characters not
yet specially studied. One curious feature, however,
remains to be noticed, the variability of vein C of tlie
hindwing, usually one of the most stable features in
Geometrid venation. In the type-species (noctuata) this
is very closely appressed to SC for some distance near the
base, with apparently a break in the cell wall atone point,
constituting point anastomosis (the scaling is too dense to
allow of absolute certainty without denudation). In one
example (La Faz) of srdndosa, the left hindwing shows this
point-contact, the right wing an appreciably longer anas¬
tomosis ; while in the type specimen the anastomosis is
still stronger, and in Loth wings. In palpus, antenna, legs,
forewing venation and fovea, sabulosa agrees well enough
with the generic characters of Mimophyle. This genus
(and perhaps the allied Eupileta, Warr., Nov. Zook, xii,
362 ; cf Froc. U.S. Mus., xxx, 521) may be regarded as
making a transition from those with the more perfect
anastomosis of C with SC {Narrayodes and Psodopsis) to
those of more normal Boanniid structure, commencing
with the following genus.
138. Idialcis metria (Front), nov. sp.
(Flate XL VIII, lig. 20.)
d 9 . 22-28 mm. Head and uppersurface of body and wings
brown irrorated wdth fuscous; undersurface much paler, greyish.
Geometridae of the Argentine Bcpnhlie. 285
Forewing with termen subcrenulate ; basal area with a few black
spots ; antemedian line black, bent or angled in cell, then oblique,
but somewhat waved, to inner margin at little beyond one-fourth,
sometimes quite indistinct; postmedian line black, from costa at
about five-sixths, very strongly toothed outwards between SC® and
Rl, then sinuate inwards, projecting again, but more bluntly, between
E,-'* and M', thence strongly oblique inwards to submedian fold,
finally slightly oblique or nearly perpendicular to inner margin at
about one-half ; antemedian accompanied proximally by a diffuse
brown or fuscous band (sometimes distinct, sometimes hardly darker
than the ground-colour), postmedian, accompanied distally by a fine
or thicker brown line ; cell-spot black, distinct ; subterminal line
strongly dentate, pale, sometimes quite ill-defined, sometimes
rendered distinct by being filled up with darkish shading proxim¬
ally ; marginal line black, continuous, thickened more or less into
spots between the veins. Hindwing with termen crenulate ; ante¬
median of forewing and its accompanying proximal band continued
as subbasal markings ; a black cell-spot ; a generally indistinct
median shade, usually nearer to post median than to cell -spot ; a
somewhat sinuous postmedian, but continuing approximately parallel
with termen, accompanied distally by a brown line ; subterminal
nearly as in forewing, sometimes still weaker ; marginal line con¬
tinuous, distinct, not thickened into spots between the veins.
Underside glossy, withoirt markings, excepting a cell-spot on each
wing.
Aristides Villanueva, February 24, 1905 (W. M. Bayne),
type (^) in coll. L. B. Front ; Bakle, February 21, 1904,
the only known $ , in coll. L. B. Front ; also six $s from
La Faz and Mendoza, February and March, 1903-1905, in
coll. L. B. Front et coll. A. F. Bayne : all collected by Mr.
W. M. Bayne.
Decidedly variable, not only in the strength of the
markings, but even in their position, especially in the
course of the postmedian of forewing ; in this the sub¬
costal tooth is constant, but the sinuosities from here to
inner margin seem alike in hardly two examples, varying
from very slight to very deep, the outer line sometimes
approaching close to the inner in submedian area, some¬
times keeping well away from it. Some examples have
distinct pale markings distally to the postmedian, espe¬
cially a patch near the inner margin.
Evidently quite closely related to jacintha, Butl., Tr.
Ent. Soc. Bond., 1882, 414, from Chili, with wdiich it
28G
Louis B. Front on the
entirely agrees structurally, unless possibly the face may
be very slightly more protuberant and the palpus slightly
more upeurved. It may be distinguished by its generally
smaller size and its darker, much more uniiorm colouring,
but may possibly prove a mere local race. The genus
Idialcis, Warr., Proc. U.S. Mus., xxx, 509, for which Mr.
Warren specifies type jacintha, Butl., was evidently
characterised on some other species, for he gives the $
antenna as bipectinate, of fore wing as “ stalked or
coincident,” etc. According to the rules of nomenclature,
however,* Idialcis must stand io\' jacintha, and the generic
characters must be amended. For present purposes it is
sufficient to point out that the ^ antenna is nearly
siinple, not bipectinate, SC- of forewing arises separately
from cell, anastomosing with SC\ and C of hindwing is
approximated to SC to at most two-thirds of cell (which
can scarcely be described as “ nearly the whole length ”),
and does not appear to anastamose. Phihalapteryx edna,
Butl., Tr. Ent. Soc. Bond., 1882, 414, also belongs to this
genus. The essential characters differ little from those of
Mimopliyh, though the point of origin of SC‘^ is different.
139. Neotaxia plana (Dognin).
Neotaxia plana, Dognin, Ann. Soc. Ent. Belg., xlviii, 3G6
(1904).
Tucuman (type), in coll. Dognin.
140. Phrygionis paradoxata (Guenee).
Byssodes paradoxata, Guenee, Spec. Gen. des Lep., ix, 400
(1858).!
Eulepidotus paradoxata, Moschler, Abh. Senck. Ges., xvi,
245 (1890).2
Byssodes privignaria, Druce, Biol. Centr.-Amer., Lep. Het.,
ii, 98 (1892).3
Phrygionis incolorata, Warren, MS., in coll. Br. Mus. (var. ;
an sp. ?).
Buenos Aires, one ^ at light in house, April 1901
(A. F. Bayne), in coll. L. B. Pi’out ; a second, undated, in
coll. A. F. Bayne.
* “When in the original publication of a genus, one of the species
is definitely designated as type, this species shall be accepted as type
regardless of any other consideration.” (“ Science,” October 18, 1907,
p. 521, Art. .30, Rule 1 (a).)
Geometridae of the Argentine Repvhlic.
287
Guenee (^) gives the locality for his specimen as
“ Brazil (?).” The forms wliich appear to agree with his
description, and which I take it that Moschler (^) has
rightly identified from Porto Rico, are widely distributed
— West Indies, Central America (^), Para (in coll. Br. Mus.),
Rio Janeiro (coll. Br. Mus.), being localities known to me.
It seems to pass in some collections, as well as in the
“ Biologia,” as 'primgnaria, Guen. ;* but the true privig-
naria, from Martinique, was a “ cafe-au-lait ” coloured
species with the distal margin of forewing somewhat
elbowed. Further south (Sao Paolo, Porto Alegre, Monte¬
video, Buenos Aires) iyficdA par adoxata, is replaced, so far
as I can judge from meagre material, by a local race which
entirely lacks the orange shade behind the postmedian of
hindwing and has the red spot pro.ximally to the subter¬
minal metallic one above R^ wanting or much reduced.
Should this prove constant, as I expect, it will stand as
Phrygionis paradoxata incolorata (Warren, MS.), Prout, nov.
subsp.
141. Ratiaria platinata (Guenee).
Urapteryx platinata, Guenee, Spec. Gen. des Lep., ix, 33
(1858).!
Ratiaria argentilinea, Walker, List Lep. Ins., xxii, 827
(1861).2
Byssodes mollita, Doguin, Bull. Soc. Ent. Fr., 1891, clvii
(nov. syn.) (var. ?).^
Tucuman, in coll. Rothschild.
Range: (^) (‘^) Brazil; (^) Venezuela ; Colombia, in coll.
Rothschild.
142. Nematocampa arenosa (Butler).
Nernatocampa arenosa, Butler, Tr. Eiit. Soc. Loud., 1881,
323 (1881).
Tucuman and Salta, in coll. Dognin.
Butler’s type was from the Amazons. The species is
also represented in coll. Br. Mus. from Mexico and Rio
Janeiro, and in coll. Dognin from Costa Rica, Ecuador and
south Brazil, so that it evidently has a wide range. It
varies considerably, and may conceivably be the same as
vest'Uaria H.-S., Samml. Aussereur. Schmett., i, fig. 368 =
resistaria H.-S., ibid. p. 41, with which Butler compares it.
* Bijssodes prirkjnaria, Guen., Spec Gen. des Lep., ix, 400.
288
M^Louis B. Front on the,
The last-named species is said to be from Brazil; but
Herrich-Scbaeffer’s figure looks so much like a specimen
of the North American N. limhata (Haw.) = Jilamentaria,
Guen., that I suspect there may be a mistake as to the
locality, as we know is the case with a few of Herrich-
Schaefter’s records.
“ 143. Urepione quadrilineata (Walker).”
Ilijperetis (?) quadrilineata, Walker, List Lep. Ins., xxvi,
1502 (1862).^
? Azelina contorta, Thicrry-Mieg, Le Nat., xiv, 217 (1892).-
? Semiothisa (?) contorta. Druce, Biol. Centr.-Amer., Lep.
Hot., ii, 133, tab. liii, 19 (1893).=^
Urepione quadrilineata, Warren, Nov. Zook, ii, 158 (1895).
Tucuman, in coll. Dognin.
The above synonvmy is according to Schaus, Tr. Amer.
Ent. Soc., xxvii, 177, and the localities should be : (^)
Amazons ; (^) Peru ; (^) Central America, Colombia and
Ecuador. But according to M. Dognin (in litt.j Schaus’
mlloriui {Joe. cit.) is synonymous with contorta and qnad-
rilineata a little different. The three may all be one
species. Personally, I have been inclined to regard those
forms with the antemedian line close to (or touching) the
discal spot as distinct and as the true valloma ; this is the
form (or species) which occurs in Paraguay, and M. Dognin
informs me that the line is so placed in his Tucuman
“ quadrilineata.”
144. Melinoides apricaria (Herrich-Schaeffer).
Metrocampa apricaria, Herrich-Schaeffer, Samml. Aus-
sereur. Schmett., tab. Ixiv, 3G3 (1855).
Heterolocha apricaria, Guenee, Spec. Gen. des Lep., ix, 108
(1858).
Salta, in coll. Dognin.
Widely distributed from Mexico to Uruguay, originally
described from Venezuela. Strongly variable.
145. Melinoides fulvitincta (Warren).
Melinoides fulvitincta, Warren, Nov. Zook, xii, 69 (1905).
Tucuman (type), in coll. Rothschild.
I have only seen the type specimen, a It is quite
G-eometridae of the Argentine Republic. 280
possible it is only a very extreme aberration of M.
apriearia.
146. Melinoides albarita (Dognin).
Iteterolocha albarita, Dognin, Ann. Soc. Ent. Belg., xxxvii,
158 (1893).
Tucuman, in coll. Dognin.
Described from Loja. I have seen a large number from
Bogotd, Colombia.
147. Numia deceptrix (Warren).
Nuiiiia deceptrix, Warren, Nov. Zool., xii, 69 (1905).
Tucuman, in coll. Rothschild (type) et Bastelberger.
148. Perusia citrinata (Snellen).
Perusia citrinata, Snellen, Tijd. Ent., xvii, 25, tab. ii, 1
(1874).
Salta, in coll. Dognin.
Snellen’s type was from Bogotd. I believe the species
(or form) is widely distributed.
149. Microxydia orsitaria (Guenee).
Stegania{T) orsitaria, Guenee, Spec. Gen. des Lep., x, 49
(1858).
? Rumia defixata. Walker, List Lep. Ins., xxvi, 1499
(1862).
? Goi'ycia sulphurata, Maassen, Stlibel’s Reisen, Lep. 161,
tab. viii, 18 (1890).
? Microxydia rufifinibriata, Warren, Nov. Zool., xi, 574
(1904).
Tucuman, in coll. Dognin et coll. Bastelberger.
Widely distributed throughout a great part of the
Neotropical Region, though I am not aware of its occurrence
further south than Tucuman and S.E. Brazil.
“Varies excessively,” as Guenee gathered even from the
three specimens known to him. It is probable that the
names cited in the synonymy above should be sunk to it,
and not unlikely some of the others which have been
brought forward as new species of Microxydia.
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) U
290
Mr. Louis B. Prout on the
150. Lomographa bicineta (Dogniu).
Coricia hicincta, Doguin, Le Nat., xiv, 144 (1892).
Tucuman, in coll. Dogiiin.
Dognin’s types were from Ecuador. Occurs also in
Peru.
151. Nipteria CURVIFASCIA (Warren).
Nii^teria curvifascia, Warren, Nov. Zool., viii, 476 (1901).
Rosario, two one $ (type $ and cotypes), in coll.
Rothschild ; Buenos Aires, in coll. Br. Mus.
No doubt the specimens from Castro, Parana, mentioned
by Warren (loc. cit.), belong to a nearly allied species which
stands in coll. E. D. Jones as caintaria, Schaus (MS.), and
which has also been presented to the British Museum by
Mr. Schaus, under the name of curvifascia. I have seen
no other example of true curvifascia than the Argentine
ones.
152. Nipteria (?) fumida (Burmeister).
Upcnora fumida, Burmeister, Descr. Phys. Arg. Rep.,
V (1), 414 (1878).!
Nephodia fumida, Berg, An. Soc. Cient. Argent., xiii, 181
(1882).2
(^) Buenos Aires ; (f) Oran and Salta.
Burmeister’s genus Upcnora was erected for this species.
He figures only the venation (Atlas, tab. xvii, 15) and this
somewhat incorrectly. But Berg’s careful notes and cor¬
rections (^) make it certain that the species belong to the
Nephodiinac of Warren. Its actual identity has not been
made out. The only species of the group which I yet
know from Buenos Aires is curvifascia) but had the
authors had this species before them, they could hardly
have overlooked the distinct band. It is in the highest
degree probable that Berg’s Salta fumida will prove to be
N. marginata, Warr. ; but there is no certainty that Berg
correctly identified this with the Buenos Aires insect.
153. Nipteria marginata (Warren)
(praec. syn. ?).
Nipteria marginata, Warren, Proc. U.S. Mus., xxx, 502
(1906).
Salta, type in coll. Rothschild ; Tucuman, in coll. Dognin.
Geometriclae of the Argentine BeimUic.
291
154. Nipteria saturata (Dognin).
Nipteria saturata, Dognin, Ann. Soc. Ent. Belg., 1, 117
(1906).
Metan, Salta, type (i^) in coll. Dognin.
155. Pantherodes pardalaria (Hiibner).
Panthera imrdalaria, Hiibner, Zutr., ii, 25, fig. 8o5-6
(1823).
Pantherodes 'pardalaria, Guenee, Spec. Gen. des Lep., x,
200 (1858).
Panthera pardalaria, ab. obliterata, Warren, Nov. Zool., xvi,
100 (1909) (ab.).
Argentina (O. W. Thomas), without exact locality,
in coll. Br. Mus. ; Tucuman, in coll. Rothschild et coll.
Bastelberger. I believe Mr. A. F. Bayne has taken one or
two in Buenos Aires, but have no precise notes; a small
$ from thence (November 7, 1908, C. Gi'ant), in coll. Br.
Mus., has the abdomen entirely dark, and if this is really
a reliable distinction for P. eolubraria, Guen., Avhich Mr.
E. Dukinfield Jones assures me is structurally a distinct
species, it will have to be referred there.
156. Cataspilates plurilineata (Warren).
Aspilates plurilineata, Warren, Nov. Zool., iv, 473 (1897).
Mendoza, January 1903, one $ (W. M. Bayne), in coll.
L. B. Prout; Cacheuta (W. M. Bayne), one J in coll. A.
F. Bayne.
Warren’s type was from Bolivia. The species occurs
also in Brazil and in Peru.
The single Mendoza example is small and narrow¬
winged, even for a $, but it is of course impossible to .say
whether this indicates the existence of a local race.
157. Cataspilates pseud aluma (Dognin).
Cataspilates pseudaluma, Dognin, Ann. Soc. Ent. Belg., li,
18 (1907).
Tucuman (type), in coll. Dognin.
u 2
292
Mr. Louis B. Front on the
158. Macaria (?) REPETITA (Prout), nov. sp.
(Plate XLVIII, fig. 3.)
1^. 29 min. Head, body and wings pale reddish brown, irrorated
with darker — nearly as in the European Lithina jMraria, Linn.,
but slightly duller ; hindwing paler abov'e, especially in costal
region. Forewing with the lines dark, mostly straight, only the
po.stmedian well expressed throughout; subbasal line weak ; ante-
median oblique, weakly sinuate, from costa at one-third to inner
margin at scarcely beyond one-fourth, most distinct at inner margin ;
a faint median shade at somewhat before one-half ; postmedian from
costa at nearly four-fifths to inner margin before two-thirds, some¬
what darkened on the veins, and accompanied distally by a weaker
line. Hindwing with a moderately distinct dark discal spot, a post¬
median line from inner margin at about three-fifths, nearly straight,
but faintly curved towards termen, becoming indistinct, and vanish¬
ing about R' ; distal margin from tornus to rather broadly shaded
with 2)ale brownish (colour of forewing). Undersurface rather
thickly scaled (much as in Lithina), inner margin of forewing the
palest ; markings nearly as above.
La Paz, Mendoza, August 23, 1905 (W. M. Bayne).
Type in coll. L. B. Prout.
Recalls in shape, and in some other points, Thamnonomct
imitata, Druce, Biol. Centr.-Amer., Lep. Het., ii, 134, tab.
liii, 20, 21, but differs, inter alia, in having non-pectinate
antenna and in wanting the long frontal cone of hairs.
Unfortunately, both antennae are broken, but stumps
remain, one of them to the length of about ten segments,
and these show a somewhat dentate, ciliated structure,
perhaps not so strongly serrate-dentate as in the group of
Maearia gigantata, Guen., Spec. Gen. des Lep., x, 78. I
am not satisfied with the existent divisions of Macaria on
secondary male characters, nor, certainly, with Meyrick’s
impossible sinking of the genus to Opisthograptis, Htib.
The present species has not the facies of a Macaria, but
rather that of a Lithina ( = Lozogranima). It may probably
be near, or possibly even a form of Lozogranima (?) setaria,
Schaus, J. N. Y. Ent. Soc., vi, 146, from S.E. Brazil, with
which I am unacquainted, but the brief description gives
no information as to wing-shape or structure, and the
generic query is Mr. Schaus’ own. In Lithina vein SC^
is present; in M. repetita the structure is as in Macaria
and Thamnonoma, SC^ absent, SC^ closely approaching G
Gcometridae of the Argentine Rcfublic.
293
(possibly connected at a point) and afterwards connected
by a short bar with SC^ The fovea is strongly developed.
The hindlegs are wanting.
I have assumed Macaria, Curt. (September 1, 1826), to
be an older name than Hiib. (Zutr., indescr. ;
Verz., 298, 1826 ?). The exact date of publication of the
latter is still unascertained, but as Meyrick and Aurivillius
recommend quoting 1826, and Hampson 1827, it seems
undesirable to substitute the Hiibnerian name for Curtis’ ;
according to a recommendation of the German Zoological
Society, a genus with a type fixed should, in cases of
doubt, be preferred to one without. Curtis fixed his type,
Macaria litnrata (Clerck), and gave a generic diagnosis far
in advance of anything of Hiibner’s.
159. Macaria peltigerata tGuenee).
Macaria peltigerata, Guenee, Spec. Gen. des Lep., x, 79,
(1858).
Tucuinan, in coll. Dognin.
Guenee’s type was from the Amazons.
160. Macaria transvisata (Guenee).
Macaria transvisata, Guenee, Spec. Gen. des Lep., x, 7l
(1858).
Paraseinia distans, Butler, Tr. Ent. Soc. Lond., 1881, 343
{partinx).
Buenos Aires, February 1901, February 16, and
December 16, 1903 (A. F. Bayne), three ^s in coll. L. B.
Front. I believe Mr. Bayne has taken one or two sub¬
sequently, and there is one from the same locality
(H. Wilkinson) in coll. Br. Mus.
This is the most southerly locality known for this very
widely-distributed eastern Neotropical species (or form),
which ranges northwards to Central America. It is
doubtful whether it is really anything more than an
aberration or variety of M. regulata (Fabr.) = notata
(Cram., nec Linn.) = enotata (Guen.), but the Buenos Aires
examples belong to the transvisata form. Butler’s distans
was properly “ nov. nom.” for notata. Cram. (fig. G), nec
Linn., in other words, an absolute synonym of enotata,
Guen. ; but his Amazonian specimen is a somewhat inter¬
mediate form, though clearly transvisata if that be a
separate species.
294
AF^^jouis B. Prout on the
161. Macaria arenisca (Dognin).
$ Macaria arenisca, Dognin, Ann. Soc. Ent. Belg., xl, 145
(1896).
$ Semiothisa inexcisa, Warren, Nov. Zool., iv, 480 (1897)
(nov. syn.).
IVIetan, Salta, in coll. Dognin.
Very generally distributed in the Neotropical Region,
from the West Indies and Colombia to nortliern Argentina.
Dognin described from nine ^s from Ecuador, Warren from
a $ from Trinidad. The two are certainly synonyms,
though there may be some slight sexual differences.
162. Macaria ramparia (Schaus).
Semiothisa ramparia, Schaus, Tr. Amer. Ent. Soc., xxvii,
168 (1901).
Tucuman, in coll. Dognin.
Schaus’ type was from S.E. Brazil, and I have not seen
specimens from any other country, excepting M. Dognin’s.
163. Macaria subclathrata (Warren).
Semiothisa subclcdhrata, Warren, Nov. Zool., iv, 432 (1897).
Tucuman, in coll. Dognin.
Likewise described from S.E. Brazil. Known to me
also from Paraguay and British Guiana.
164. Tephrinopsis rectilineata (Warren).
Tephrinopsis rectilineata, Warren, Nov. Zool., vii, 206
(1900).
Parana, Entre Rios (type and a series of both sexes), in
coll. Rothschild ; Buenos Aires, fairly common, especially
at light, from February to April (A. F. Bayne); San Juan,
one December 21, 1903, in coll. A. F. Bayne ; ? Tucuman
and Los Vasquez, in coll. Dognin.
Also occurs at Sapucay, Paraguay.
A decidedly variable species in the strength of the lines,
the presence or absence of a duplicating line or shade,
wider or narrower, distally to the postmedian, etc. It is
not impossible that the specimens which I refer to the
following species really represent another form of this,
in which case the range of variation is still wider.
M, Dognin’s Tucuman and Los Vasquez specimens, no
Geometridae of the Argentine JRepuhlie. 295
doubt identified by Mr. Warren himself as rectilmeata, will
almost certainly prove to belong to the fragilis form. On
the other hand, a nice aberration taken by Mr. Bayne in
Buenos Aires, April 24', 1903, seems to me undoubtedly
referable to reetilineata — perhaps to a second brood. Mr.
Bayne at one time thought it a good species, and pointed
out that in that season reetilineata was observed from
February 27 to March 13 and not subsequently, until this
straggler appeared, in fine condition. Its antemedian line
is curved instead of straight, the median obsolescent,
median area a little narrowed, postmedian line slightly
angled at R^, etc. But other specimens to a large extent
intergrade between this and the typical forms.
1G5. Tephrinopsis fragilis (Warren) (?).
Tephrinopsis fragilis,W oxYQn, Nov. Zooh, xi, 562 (1904),
Tucuman, 450 m., January 1902, in coll. L. B. Prout;
also in coll. Bastelberger from same locality;’^ La Rioja
(Dr. E. Giacomelli), $ and $ in coll. L, B. Prout ; Tunuyan,
IMarch 1902 (A. F. Bayne), one $ in coll. L, B. Prout ; La
Paz, Mendoza, January 1903 (W. M. Bayne), one $ in coll.
L. B. Prout ; Mendoza City (W. M. Bayne), one February
23, 1903, in coll. A. F. Bayne, and one ^ in coll. L, B. Prout;
Villa Mercedes, April 15, 1904, one ^ in coll. A. F. Bayne.
Warren’s Tephrinopsis fragilis was described from Pisco,
Peru, unfortunately from poor specimens ; the British
Museum has apparently the same thing from Callao and
from Paraguay. I repeatedly and carefully compared also
the western Argentine forms (here enumerated) with the
types, and noted that they belonged “ almost certainly ” to
the same species, though somewhat larger, and apparently
with some other very slight differences, I cannot say
positively that the insect which I am recording as fragilis
is not a race of reetilineata, but I have not yet seen any
specimen which I have hesitated to refer definitely either
to the one or the other, and I am glad to know that Mr.
A. F. Bayne also supports me in keeping them distinct.
The fragilis forms, though as variable as the reetilineata,
are always more strongly dark-dusted, the lines weaker
and somewhat less straight, the postmedian of forewing
marked with dark vein-spots or dashes, the underside
* And I think in coll. Dognin from Tucuman and Los Vasquez ;
see under T. reetilineata.
296
JMi^Louis B. Prout on the
less clearly marked, etc. ; they are also a little larger
{rectilineata ; 20-22 mm. ; fragilis 24-25 mm.), but as the
typical fragilis from Peru measure only 20-22 mm., I do
not overpress this. If the two are geographical races of
one species, their range is a little curious ; for although
rectilineata can roughly be regarded as eastern and fragilis
as western, I have recorded a specimen of the former from
San Juan, 68° 57' W. long., and one of the latter from
Villa Mercedes, 55° 20' W. long.
The example from Villa Mercedes is the most interest¬
ing aberration; it is rather large and very well marked,
the postmedian line replaced by the row of dark vein-
dashes, which are very strong, and the shade distally to
postmedian also rather conspicuous. Perhaps, taken in
conjuction with the date (April 15), this indicates a more
sharply-marked second brood, as I have suggested regard¬
ing the previous species.
166. Tephrinopsis subumbrata (Dognin).
Tephrinopsis siihumlrata, Dognin, Ann. Soc. Ent. Belg.,
1, 119 (1906).
Buenos Aires, one Tucuman, a pair (the three speci¬
mens from which the original description was made), in
coll. Dognin ; also others from Tucuman, in coll. Dognin et
coll. Bastelberger ; Salta, in coll. Dognin ; La Rioja (Dr.
E. Giacomelli), in coll. L. B. Prout ; La Paz, Mendoza, ^
at light, November 22, 1903 (W. M. Bayne), in coll. L. B.
Prout ; Mendoza City, February 2, 1905 (W. M. Bayne),
in coll. L. B. Prout; Palmira (W. M. Bayne), in coll. A. F.
Bayne et coll. L. B. Prout (one or two bred ex ovo, W. M.
Bayne). Mr. A. F. Bayne has also two or three undated
specimens from La Paz, etc.
Not yet known outside the Argentine Republic. A
single example of an undescribed species from Araucania,
Chili, closely related to this, stands in the British Museum
collection.
Another very variable species. The upper surface is
generally quite weakly marked, and comparatively uni-
colorous, though much darker in some examples than in
others ; but one interesting aberration, from Mendoza, has
both wings quite pale proximally to the postmedian line,
and strongly dark-shaded distally thereto. On the under¬
side of the hindwing there is still greater variation ; the
Geometridae of the Argentine Meggnblic. 297
postmedian may be almost continuous, or broken up into
spots, both lines vary much in the acuteness of their angu¬
lation, some specimens are more clouded or banded than
others distally to the postmedian, while the degree of
dark dusting varies just as materially as above. Finally,
the form from Salta, as I learn from an example kindly
sent for my inspection by M. Dognin, is decidedly larger
than the typical forms, being in this respect intermediate
between the present species and the following, though
clearly referable to the present one.
This species, with that about to be described, and the
Chilian insect mentioned above, will form a very compact
group, quite distinct in aspect from normal Teplirinopsis,
and which will possibly in time be found susceptible of
generic separation ; but at the moment I can point to
nothing more tangible than their quite different, more
glossy scaling and their different scheme of markings.
The forewing venation has been examined in most of
them, and yields nothing abnormal; SC^ is absent, SC^
usually free from (though close to) C, anastomosing at a
point or connected with SC^h Even the $ antennal
structure is much like that of the generic type, T. paral-
lelaria (Walk.), being subdentate, the moderate or longish
cilia being arranged in anterior and posterior pairs of
small compact fascicles on each segment.
167. Tephrinopsis sijbscripta (Prout), nov. sp.
(Plate XLVIII, fig. 2.)
. 33-34 mm. Extremely close to T. suhmnbrata, Dogn., dis¬
tinguished by its considerably larger size, longer antennal cilia (in
subumbrata scarcely longer than diameter of shaft, in subscripta
about twice as long), slightly narrower and more extremely glossy
wings, usually shorter cell of the hindwing, with consequent longer
stalking of SC^ with Rh* and by the position and course of the lines
on the underside of the hindwing. These seem to be always strongly
expressed, whereas in sKbimibrata they are oftenest comparatively
weak and interrupted. Antemedian line from costa at about 6 mm.
to inner margin at about 5 mm. (in the type specimen obsolescent at
its extremities), oblique outwards from C to bifurcation of SC^-R^
thence diffuse, running nearly straight to inner margin, or slightly
* Both species, however, are slightly variable in venation ; in
subtimbrata SC^ and Ri may be either virtually connate or quite
appreciably stalked.
298
m Louis B. Front on the
concave, accompanied by dark marks along the veins ; postmedian
line obsolescent at costa, very oblique outwards from SC^ to R*,
obsolescent distally to cell, very strongly concave from R^ to anal
angle, dark-marked on R^, and
Puente del Inca, type and two co-types (one dated
January 14, 1904) in coll. L. B, Front, one $ in coll. A. F.
Bayne; all collected by W. M. Bayne.
Mr. A. F. Bayne is strongly of opinion that this species
is distinct from T. mihmnbrcda. of the lower altitudes, and
the structural differences favour this view. The great
variability of Dognin’s species, however, has already been
commented upon, and must not be lost sight of, and even
the position of its lines sometimes shows an approach to
T. subsc7'i2}ta ; so that possibly some systematists will
prefer to regard the latter as a local, mountain race. On
the other hand, it is just possible tliat there are tw'o oi
three very closely allied species mixed up under S7(huinhrata.
168. Tephrinopsis mesoscia (Front), uov. sp.
(Plate XLVIII, fig. 26.)
d. 20 mm. Face fuscous, lighter above. Palpus fuscous. Antenua
much as in the preceding species, with rather thin, sessile fascicles
of moderately long cilia. Legs light brown-grey, spotted with
fuscous ; fore- and midtibiae and tarsi fuscous anteriorly, the latter
ringed Avith light brown-grey ; hindtibia much dilated. Body fuscous
above, light brown-grey beneath. Wings somewhat tawny browm,
the ground-colour being formed by a mixture of brown-grey, tawny
and whitish scales, irrorated with dark fuscous (more sparsely on
hindwing). Forewing with lines blackish fuscous ; a subbasal line
from one-fourth costa, strongly outcurved in cell, weaker and
more slightly outcurved in posterior half ; antemedian from
two-fifths costa to just beyond one-half inner margin, entangled
on M and SM2, accompanied distally by a diffuse dark grey
shade ; postmedian line reduced to a thick, oblique streak from
two-thirds costa as far as R* ; subterminal commencing as a Avhite
streak from costa, obliterated beyond R', where it meets a longi¬
tudinal white streak running to the end of postmedian ; an ill-
defined taAvny blotch in median area distally to cell ; some ill-defined
dark markings in sub terminal region, the most distinct being a dark
spot on each side of R^, like the “ twin spots ” of many Larentiids ;
marginal line dark, thickened between the veins ; fringe intersected
by a weak dark line. Hindwing with an indistinct dark antemedian,
which traverses the extreme distal margin of cell, and is angled at
Geometridae of the Argentine Be'puhlic. 299
lower extremity ot cell ; very faint suggestions of median and post-
median lines. Underside light brown-grey ; fore wing with the lines
indicated at costa and the central shade across the wing ; hindwing
with the markings rather stronger than above.
La Paz, Mendoza, October 25, 1905 (W. M. Bayne),
type in coll. L. B. Prout.
Fovea wanting ; SC^ connected with C, and anasto¬
mosing at a point with SC^'^.
169, Xenoecista pallidata (Warren).
Xenoecista 'pcdlidata, Warren, Nov. Zool., iv, 484' (1897).
La Plata, in coll. Rothschild.
Warren’s type was from the Amazons ; he states {loc. cit.)
that the La Plata specimen is rather larger and more
darkly marked. Other localities are Colombia (coll.
Rothschild), British and French Guiana and Paraguay
(coll. Br. Mus.). .
170. Thamnonoma tripartita (Prout), nov. sp.
(Plate XLVIII, fig. 35.)
(J. 24 mm. Forewing light clay-colour, dusted with fuscous grey,
the dusting being the most dense in the median area and terminally ;
proximal margin of central area weakly defined, from about one-
third of costa, curved, then oblique basewards ; weak indications of
a dark median shade near proximal margin of central area, nearly
vertical, becoming more distinct at inner margin ; distal margin of
central area weakly defined, but with a row of vein-dots indicating
postmedian line, somewhat incurved distally to cell and bent distad
at inner margin, otherwise parallel with termen, from costa at
beyond two-thirds to inner margin at about two-thirds. Hindwing
with basal and median areas rather strongly dusted with fuscous-
grey, distal area somewhat lighter, more shaded with clay-colour ;
a dark mark on inner margin indicates probably the point of origin
of an obsolete antemedian. Underside more weakly marked, faint
indications of the colour arrangement of the uppersurface. Head,
body and legs more or less concolorous with wings.
Aristides Villanueva, November 3, 1905 (W. M. Bayne)
type in coll. L. B. Prout; La Paz, one ^ in coll. A. F.
Bayne.
In the coloration and in the weakness of the markings
this species reminds a little of T. ochrifciscia, Warr., Nov,
iOiiis B. Prout on the
;}0()
Zool., iv, 494, from the Bermudas; but it is different in
shape (hindwing not toothed at R^, etc.), and is hardly a
typical Thamnonoma — fovea undeveloped, face without
hair- tuft. The antenna shows the Thamnonoma structure ;
the hindtibia is rather short and thick, with hair-pencil ;
in the forewing SC^ is absent, SC^ connected by short
bars successively with C and h
171. Cyclomia ocellata (Warren),
Gyclomia costipancta, ab, ocellata, Warren, Nov. Zool., vii,
213 (1900).
Santa Fe, Ocampo, August 1902, and Gran Chaco, near
Florenzia, October 1902 (S. K. Wagner), in coll. Br. Mus.
Described from Venezuela. Warren also mentions
similar forms from Bolivia.
172. Cyclomia vinos a (Dognin).
Fidonia vinosa, Dognin, Le Nat., xii, 19 (1890); Lep.
Loja, 62, tab. vi, 17 (1891).
Metan, Salta, in coll. Dognin.
Dognin’s type was from Ecuador. The species also
occurs in Colombia, Bolivia, and I think Peru. The head¬
quarters of the genus Cyclomia, would seem to be about
the equator, and northward to the West Indies. The
present species and the preceding are the only stragglers
yet detected in Argentina, and even they have not been
found very far south.
173. Dyscia SERENA (Dognin).
Dyscia serena, Dognin, Ann. Soc. Ent. Belg., 1, 118 (1906).
Buenos Aires, type in coll. Dognin.
174. Thysanopyga strigata (Warren).
Thysanopyga drigata, Warren, Nov. Zool., xiv, 293 (1907).
Ciudad de Tucuman, April 1903, type (^) in coll.
Rothschild ; La Rioja, one ^ (Dr. E. Giacomelli), in coll.
L. B. Prout.
I know of no specimens but these two. My ^ is less
strongly dusted with rust-colour than Warren s type, but
of course it is impossible to say from a couple of specimens
whether the variation is sexual or local or individual.
301
Geometridae of the Argentine Reijuhlic.
The forewing venation is as in the type species {apici-
truncaria, H.-S.) and carfinia, Druce (= nigricosta, Warn,
Nov. Zoo]., xii, 61, nov. syn.), SC^ being free,* whereas
in ail the examples of divisaria, Walk., which I have
examined, SC^ anastomoses with C.
175. Oenothalia vestita (Warren).
Oenothalia vestita, Warren, Nov. Zool., xvi, 105 (1909).
Tucuman, 1100 m., January or February, 1905, type
{$) in coll. Rothschild ; Salta, one $ (“ belongs apparently
to the same species,” Warren, loc. citi) in coll. Rothschild.
176. Sphacelodes vulneraria (Hiibner).
Brotis vulneraria, Hiibner, Zutr. Exot. Schmett., ii, 23,
fig. 319, 20 (1823) ; Verz. bek. Schmett., 803
(1826 ?).
Sphacelodes vulneraria, Guenee, Spec. Gen. des Lep., x,
117, tab. xxii, 9 (1858).
? Sphacelodes micacearia, Guenee, Spec. Gen. des Lep., x,
117 (1858) (ab.?).
Brotis fusilineata. Walker, List Lep. Inst, xx, 264 (1860).
Sphacelodes Jloridensis, Holland, Papilio, iv, 72 (1884).
Sphacelodes hrunneata, Warr., Nov. Zook, xiv, 292 (1907)
(ab. ?) (nov. syn.).
Tucuman, in coll. Dognin.
A widely distributed and often abundant species,
ranging from the eastern United States to Brazil. It
has not yet, however, been much observed in Argentina,
and probably reaches its southernmost limit at about the
latitude of Tucuman.
177. Drepanodes plebejata (Snellen).
Apicia plehejcda, Snellen, Tijd. Ent., xvii, 19, tab. i, 7
(1874).
Tucuman, in coll. Dognin.
I am not able to indicate many localities for this
species, though they suggest a wide distribution. The
* Warren, Nov. Zool., iv, 477, says that in Thysanopyga SC^
anastomoses strongly with C ; he must have made this note from
divisaria, or in any case not from the type species, for in the latter
these two veins are quite far apart, so that no error of observation
would have been possible.
302
Af^Louis B. Prout on the
British Museum possesses examples from Paraguay and
Montevideo. Snellen’s type was 'pi'dbably from Colombia.
178. Drepanodes asina (Druce).
Drepanodes asina, Druce, Biol. Centr.-Amer., Lep. Het.
ii, 31, tab. xliv, 4 (1892).
Tucuman, in coll. Dognin.
Druce’s type was from Guatemala. The species occurs
also in Peru, and I strongly suspect that D. fidvilinea,
Warr., Nov. Zool., xi, 138, is a mere aberration of it.
179. Gynopteryx gladiaria (Guenee).
Gynoidcryx gladiaria, Guenee, Spec. Gen. des Lep., ix, 79
(1858).
Gynopteryx seriaria, Guenee, Spec. Gen. des Lep., ix, 79
(1858).
? Gynopteryx rhomhao'ia, Guenee, Spec. Gen. des Lep., ix,
79, tab. X, 4 (1858).
Gynopteryx telysaria, Walker, List Le^). Ins., xx, 94
(I860).
Apicia calhisaria, Walker, List Lep. Ins., xx, 99 (1860).
Apicia nazadaria, Walker, List Lep. Ins., xx, 109 (1860).
Apicia uxiaria. Walker, List Lep. Ins., xx, 110 (1860).
Gynopteryx ennomaria, Warren, Nov. Zool., iv, 492 (1897).
Apicia maldama, Scbaus, Tr. Amer. Ent. Soc., xxvii, 175
(1901).
? Gynopteryx liyulifera, Warren, Nov. Zool., xii, 370
' (1905).
Hyperythra heterolocha, Stand inger and Bang- Haas, MS.
Buenos Aires (A. F. Bayne), in coll. L. B. Prout; also
in coll. Br. Mus. (H. Wilkinson), and mentioned from
there by Guenee (his seriaria, var. A) ; San Luis, La
Paz, Palmira, Mendoza and San Juan (W. M. Bayne),
in coll. L. B. Prout, coll. A. F. Bayne, etc. ; La Rioja
(Giacomelli) ; Tucuman, in coll. Dognin (recorded as
cnnomariai) et coll. Bastelberger ; Los Vasquez (as
ennomaria), in coll. Dognin ; Santa Fe, Ocampo, in coll.
Br. Mus. ; Gran. Chaco, near Florenzia, in coll. Br. Mus. ;
also a few specimens in coll. Br. Mus. without exact
localities (O. W. Thomas).
A common species in the southern parts of Brazil, in
Paraguay and in Argentina. I have not seen specimens
from the eastern part of the continent further north
303
Geometriclae of the Argentine Repuhlie.
than Rio Janeiro. As regards the west it seems to occur
more sparingly in Bolivia and S.E, Peru, and is repre¬
sented in E. Peru by a local race or subspecies which I
propose to describe elsewhere. In Paraguay, about Sapucay,
there occurs also a very interesting twin species, differing
structurally in the shorter, more strongly ciliated pectina¬
tions of the $ antenna, but otherwise excessively similar,
and showing a parallel range of variation, though usually
of a very pale fawn-colour, rarely of the bright yellow
which is commonest mgladiaria, and further distinguishable
in the smaller subapical patch, which is usually broken
up into three black spots, and the sharper postmedian line
of the undersurface. This is Gynopteryx nitida (Warr.),*
? = inapicata (Warr.).t It is not a geographical repre¬
sentative, for true gladiaria also occurs at Sapucay.
For the study of the true Gynopteryx gladiaria and its
aberrations, the brothers Bayne have provided excellent
material. I have already recorded (Proc. Ent. Soc. Bond.,
1906, pp. XV, xvi) that they have bred the species rather
freely ex ovo, and have established the identity of most of
the named forms, which were earlier believed to rank as
specifically distinct. It is unnecessary to reproduce the
statistics there given, but I may add that the ^ form
recorded from San Juan, and which I now take to be that
indicated by Mr. W. M. Bayne in his note on the parent $
of “ brood 1,” seems to be identical with that which Mr.
Warren has named ennomaria. It appeared also among
“ brood 4 ” and “ brood 8,” and is therefore clearly not con¬
fined to any one season of the year. On the other hand,
I have not yet seen any more of the nearly uniform,
purplish brown form (gnaldama, Schaus) which occurs in
both sexes, and which I still think may be seasonal.
The range of variation cannot here be exhaustively
analysed, but it deserves some attention. Guenee’s
gladiaria (the type form) is yellow, in the $ weakly
marked, in the $ (as in all ^ forms) with the lines dis-
* Bruchystichia nitida, AVarr., Nov. Zool., xi, 1.34.
t Gynopteryx nazadaria, ab. hiapicata, AVarr., Nov. Zool.,
xiv, 307. Certainly not referable to nazadaria, Walk., which
has the normal pectinations of gladiaria. Seems to agree accur¬
ately with a form of nitida that occurs at Sapucay, but as it
was described from Peru, I think it safer to quote tlie synonymy
with a query, until confirmatory material comes to hand. In any
case the short pectinations separate it from gladiaria,, so that it need
not be fui’ther considered here.
304
jN^^Louis B. Prout on the
tinct ; Guenee”, says that in the latter the ground-colour,
proximally to the postmedian line and as far as to the base,
is washed with reddish, and this is nearly always the case,
though the depth of the reddish shading varies greatly.
I formerly called those in which it was strongly developed
ah. rhomharia, Guen., in this following some examples so
determined in the British Museum and Oxford University
collections ; but I find Guende founds his rhomharia on
the shape of the forewing — apex less falcate, elbow at end
of more pronounced — hence probably it should rather
supplant (ah. ?) lignlifera, Warn* The ab. seriaria,
Guen., properly speaking, occurs only in the $ ; it is
characterised by the presence of dark spots on the lines
of the forewing, which may be smaller (Guenee’s type
form) or larger (his var. A) ; it is connected with gladiaria
by intermediates, minute spots appearing, in every degree
of development, on the postmedian. Walker’s nxiaria is
synonymous with ab. seriaria ; his telysaria is determined
by Schaus (Proc. Zool. Soc., 1896, 644) as rhomharia,
Guen., but I would rather refer it to the extreme form
of gladiaria ^ which, as noticed above, I formerly called
ab. rhomharia. Walker’s nazadaria is a pale, weakly-
marked ^ aberration, the ground-colour much less yellow
than in the type ; ealhisaria, Walk., is another aberration,
fawn-colour, not yellow, the markings as in ab. seriaria,
to which therefore it bears nearly the same relation as
does the form maldama to typical gladiaria. The form
ennomaria. Warn, may possibly become in some localities
(e. g. San Juan) a local race of the but, as I have
already pointed out, it occurs among typical forms at Pal¬
mira. Schaus’ maldama is the brownish form (? seasonal) of
gladiaria, the lines rather weak in the strong in the
but the two sexes agreeing in colour. As heterolocha, the
firm of Staudinger sent me a ^ from Bolivia, intermediate
in colour between typical gladiaria and the form maldama,
the postmedian line rather nearer the termen than usual,
both lines of forewing with very small dark dots on the
veins. As regards lignlifera, Warr. (? = rhomharia,
Guen.), I have seen only the type and hesitated to
sink it to gladiaria, on account of its less falcate apex and
its slightly ditferent aspect. Yet it is clear that Mr. E. D.
* I find, after writing this, that M, Dognin (in litt.) also siiggests
this synonymy.
Geomeiridae of the Argentine Re'pnblic. 305
Jones, from whom Mr. Rothschild obtained it, must have
taken it for gladiaria, or he would not have parted with
it ; for he has none like it left in his own collection. M.
Dognin, in his MS. list, records ligulifera for Buenos Aires
(Warren determ.) ; I have only ^s from that locality, but
a single ^ in the British Museum collection is quite normal
for gladiaria.
Further variation occurs in the distance which separates
the lines of the forewing. I have an interesting $ of the
maldaina form in which they are strongly convergent,
being only 1 mm. apart at inner margin; at the other
extreme of position (shown in one maldama ^ and in my ^
heteroloeha) they reach a distance of nearly 6 mm. at inner
margin.
From Buenos Aires I have seen the type and ab.
seriaria ; from different parts of Mendoza nearly all the
forms known to me ; from La Rioja and Gran Chaco ab.
seriaria’, from Santa Fe ab. ennomaria.
My Buenos Aires specimens are dated February and
March, but there is evidently a succession of broods. From
eggs obtained in July, Mr. Bayne has reared moths in
November, if not in September ; from larvae which
hatched on September 24, 1903, moths appeared in
November, and from pairings of these another brood
was already emerging on January 11, 1904.
Mr. Bayne tells me the eggs are oval, green, changing
to dark brown or blackish before hatching. As to the larva
I have no information. Mr. Bayne has sent me the empty
pupa case, which reminds somewhat in form, texture and
colour of those of the pseudo-A?momos of Europe {alniaria,
Linn., etc.), but has a series of large, blackish spiracular
spots ; anal extremity with eight hooks, the strong central
pair and six more slender.
180. Apicia fusilinea (Warren).
Apieia fusilinea, Warren, Nov. ZooL, xiv, 298 (1907).
Salta (type), in coll. Rothschild.
181. Apicia olivata (Dognin).
Apieia olivata, Dognin, Ann. Soc. Ent. Belg., li, 18 (1907).
Tucuman (type), in coll. Dognin.
TRANS. ENT. SOC. LOND. 1910 — PART III. (NOV.) X
30G
uuis B. Prout on the
182. Apicia laevipennis (Dognin).
Apicia laevipennis, Bognin, Ann. Soc. Ent. Belg., Hi, 25
(1908).
Tucuman (type), in coll. Dognin.
183. Apiciopsis stefania (Dognin).
Apicia defania, Dognin, Ann. Soc. Ent. Belg., xxxviii, 130
(1891-).i
Apiciopsiii ohliqimria, Warren, Nov. Zool., xi, 131
(1904).2
Tucuman, in coll. Dognin.
Dognin’s originals (^) were described from Ecuador,
Warren’s type (^) from Bolivia. Occurs also in Peru.
A variable species. A pale, weakly-marked $ from
Tucuman, in coll. Br. Mus., may possibly be a distinct
but closely related species, but I believe it is merely a
form of stefania.
184. Tetracis(?) erosinata (Dognin).
Tetrads if) erosinata, Dognin, Ann. Soc. Ent. Belg., li, 19
(1907).
Tucuman, one $ (type), in coll. Dognin.
185. Anisoperas UNDILINEA (Warren).
Anisoperas undilinea, Warren, Nov. Zook, xi, 565 (1904).
Tucuman, one April 1902 (type), in coll. Rothschild ;
also one from the same locality in coll. Bastelberger ; La
Rioja (E. Giacomelli) in coll. L. B. Prout ; Mendoza and
Palmira, a fine series bred by Mr. W. M. Bayne, in coll.
L. B. Prout, coll. A. F. Bayne, etc.
No other localities are known.
There is a moderate range of variation. The colour
varies appreciably, according to the density of the dark
dusting on the light-brown ground-colour ; the ante-
median and postmedian lines vary a little in their distance
one from the other, and the latter in the depth of the
sinus in submedian area ; a brown median shade (near
the postmedian) is usually well developed, sometimes weak,
very occasionally almost obsolete ; the white vein-dots dis-
tally to the postmedian are occasionally almost entirely
wanting. The $s are on the average larger than the d's.
307
Geometridae of the Argentine RepuUic.
Mr. W. M. Bayne, who has found the larva of this
and several other species {Gynopteryoz gladiaria, Microgonia
mundata, etc.) to take kindly to a garden chrysanthemum,
and has been very successful in rearing them, has bred A.
undilinea, ex ovo, on more than one occasion. The dates
of emergence indicated on my specimens are August 31 to
September 13, and November 23 to December 7, so I
suppose there ai’e two generations in the year.
186. Anisoperas cervinicolor (Warren).
Anisoperas cervinicolor, Warren, Nov. Zook, xiv, 297
(1907).
Ciudad de Tucuman, January 1902, type ($) in coll.
Rothschild ; a J from same locality in coll. Dognin.
187. Geitonia gabraria (Walker).
AzelinciiT) gabraria, Walker, List Lep. Ins., xx, 256
(1860).!
Anisoperas vesta, Druce, Biol. Centr.-Amer., Lep. Het., ii,
532, tab. xcviii, 18 (1898) (nov. syn.).^
Geitonia gracililinea, Warren, Nov. Zook, xii, 370 (1905)
(nov. syn.).^
Salta and Tucuman, in coll. Dognin.
Range : (^) {^) Central America ; (^) Venezuela ; Colombia,
in coll. L. B. Front.
The types of the three names cited in the above
synonymy have been compared, and their identity is quite
certain.
Possibly I ought to have used the generic name of
Xylolocha, Warr., Nov. Zook, ii, 158, for this species,
as Warren cited “ Type : Xylolocha gabraria. Walk.
{Azelina)!' But I learn from Mr. Warren that he en¬
tirely misidentified that species ; and I accept as type of
Xylolocha the species capucina, Th.-Mieg, Le Nat., 1892,
217 = gahraria, Warr., nec Walk. (? = linearis, Warr.,
Nov. Zook, iv. 507).
188. Sabulodes exhonorata (Guenee).
Sabulodes exhonorata, Guenee, Spec. Gen. des Lep., ix, 45
(1858).
Clysia suhopcdaria. Walker, List Lep. Ins., xx, 46, xxi, 493
(1860) (nov. syn.).
X 2
808 B. Front on the
Cli.ocrodes snhc/armna, Walker, List Lep. Ins., xx, 221 (1800)
(iiov. syn.).
Tucuman, in coll. Dognin.
Widely distributed, and common from Mexico and the
West Indies to Brazil and Paraguay, perhaps especially so
in the more southerly parts of its range. Guenee’s type
was from Brazil ; Walker’s were both from Haiti, and he
discovered their identity one with the other, but failed to
recognise in them the exhonorata of Guenee. The species
is not particularly variable.
189. Sabulodes atropesaria (Walker).
Choerodes (?) atr02xsaria, Walker, List Lep. Ins., xx, 222
(1860).
t^abulodes arenularia, Snellen, Tijd. Ent., xvii, 12, tab. i, 1
(1874) (nov. syn.).
Los Vasquez, in coll. Dognin.
The types of both the names (which are absolutely
synonymous) come from Bogotfi*. I have only seen the
species from Colombia and Ecuador.
190. Sabulodes polydora (Thierry-Mieg).
Epione ij) polydora Thierry-Mieg, Le Nat., xvi, 235 (1892).
Tucuman, in coll. Dognin et coll. Bastelberger.
Originally described from Peru. Seems to be common
in Ecuador, Peru and Bolivia.
Epione hogotata, Snell., Tijd. Ent., xvii, 13, tab. i, 8,
presumably from Colombia, is apparently related, but
distinct.
191. Spododes adumbrata (Warren).
Spododes adumbrata, Warren, Nov. Zoo)., ii, 156 (1895).
Tucuman, in coll. Dognin.
Warren’s type was from Brazil.
192. Bassania amethystata (Walker).
Bassania amethystata, Walker, List Lep. Ins., xx, 269
(I860).
Erosina meropia, Druce, Biol. Centr.-Amer., Lep. Het., ii,
55, tab. xlvi, 10 (1892) (nov. syn.).
Geometridae of the Argentine Republic. 309
Tucumaa, in coll. Dognin.
Walker’s type was from Venezuela, Druce’s from Central
America. The species is widely distributed in South
America.
193. Erosina hyberniata (Guenee).
Erosina hyberniata, Guenee, Spec. Gen. des Lep., ix, 149,
tab. vi, 7 (1858).^
Sabulodes rusticata, Maassen, StiibeTs Reisen, Lep. 153,
tab. viii, 1 (1890).^
Tucuman, 450 rn., in coll. L. B. Front ; Tucuman and
Los Vasquez, in coll. Dognin.
Described from Brazil (^) and Ecuador (“). Another com¬
mon and widely-distributed species, reaching northward
to Mexico and westward to Callao.
194. Herbita asinana (Front), nov. sp.
(Flate XLVIII, fig. 22.)
42-46 mm. Antenna with very short j^ectinations, or long
projecting teeth (hardly longer than width of shaft), well ciliated.
Hindtibia moderately dilated. Forewing with apex acute, termen
elbowed at R ^ ; hindwing with termen less even than in typical
Herbita, etc., being very faintly waved, and minutely produced at
R 3. Forewing greyish fawn-colour, finely irrorated with dark
scales, the irroration weakest in terminal area ; discal dot small,
blackish ; a small, irregular whitish costal mark at or just beyond
four-fifths, bounded proximally by some dark scales ; a faint grey
costal mark between the whitish mark and the apex. Hindwing
somewhat paler, unmarked, the presence of a discal dot faintly
indicated by a few darkened scales. Undersurface of forewing
paler than upper, with costal and distal margins narrowly, and apex
more broadly, shaded with a slightly fuller colour — approaching
that of uppersurface — but with the two shades not sharply differ¬
entiated. Undersurface of hindwing slightly more freckled than
uppersurface, discal spot small, distinct, blackish. Face concolorous
with forewing; thorax above and beneath concolorous wdth upper-
and underside of forewing, abdomen with hindwing.
Cacheuta, June 11, 1904 (W. M. Bayue), type in coll.
L. B. Front ; two other Js from same locality (coll. L. B.
Front et coll. A. F. Bayue), in less fine condition, taken by
an inspector on the railway, end of September or beginning
of October, 1904.
310
Mi^Rouis B. Prout on the
Probably related to Herhita capona, Dognin, Ann. Soc.
Ent. Belg., xliv, 228, with which it agrees in antennal and
leg-structure and in the character of the subapical mark,
and very nearly in wing-shape. The antennal form is quite
distinct from that of typical Herhita, the “ pectinations ”
being here mere lateral prolongations of the strong ventral
lamellae. Hindtibia much less dilated, with the hair-
pencil quite small. Hindtarsus not shortened (nearly as
long as hindtibia). The elbow at slightly more promi¬
nent than in capona. Subcostal venation as in most
Herhita, Ira, Erosina, etc. (SC^ from cell), whereas in
capona the cell is a little shorter, and SC^ arises well
down the stalk of SC^~^.
195. Hekbita aemula (Warren).
Herhita aeimda, Warren, Nov. Zooh, xi, 140 (1904).
Tucuman, in coll. Dognin.
Warren’s type was from Mexico. I know of no inter¬
mediate localities.
196. Ira albirenata (Warren).
Ira albirenata, Warren, Proc. U.S. Mus., xxx, 543 (1906).
Ira decurtaria, Warren, loc. cit. (nec Herrich-Schaeffer).
Warren’s type was from S.E. Peru. I cannot separate
that form definitely from the one which Warren mis-
identifies as decurtaria, H.-S. — recognisable by the elbowed
termen of forewing. M. Dognin records this “decurtaria"
from Tucuman. If there be two species, this one will
require a new name.
197. Microgonia nimbata (Guenee).
Oxyd-ia nimhata, Guenee, Spec. Gen. des Lep., ix, 59
(1858).
Tucuman, coll. Dognin.
Appears to be widely distributed from Colombia to
Paraguay and Brazil. I have not attempted to cite any
synonymy, being too uncertain which of the numerous
very similar forms, sometimes cited as species, really
belong here.
Geometridae of the Argentine Mejynhlic.
811
198. Microgonia mundata (Guenee).
Oxydia nmindata, Guenee, Spec. Gen. des Lep., ix, 58, tab.
X, i (1858).^
Microgonia nmindata ab. crassior, Warren, Nov. Zool., xiv,
312 (1907) (ab.).2
Mendoza (W. M. Bayne), in coll. L. B. Prout, et coll.
A. F. Bayne ; La Rioja (E. Giacomelli), in coll. L. B. Prout
et coll. Br. Mus.
Range : Brazil (^), Paraguay, in coll. L. B. Prout, etc. ;
Peru, Ecuador, etc. 0. Mr. Warren inclines to think that
these latter form a local race, distinguishable by their
larger size, deeper colour, and stronger markings ; yet he
only erects it as an “ aberration.” I have insufficient
material from the more northerly localities to allow of my
forming a definite opinion, but it seems doubtful whether
the distinctions are really racial ; at any rate I possess the
strongly-marked forms from Sapucay and Porto Alegre,
and my Mendoza series shows a wide range of colour-
variation.
Mr. W. M. Bayne has bred a fine lot exovo, from Palmira
and perhaps other localities in Mendoza. One generation
appeared (perhaps more or less forced ?) in June and July
1903, another in December of the same year. The latter
brood seems the lighter-coloured, though both vary. None
are quite so light as some specimens which I possess from
other localities, and the first brood consists of more or less
reddish-brown forms, one or two of the most extreme
becoming fuscous brown. The zigzag subterminal shade
of the hindwing is weak in the entire series of both
broods, often barely traceable. My three examples from
Dr. Giacomelli, La Rioja, also vary a good deal ; all three
belong to the lighter forms, but one is richer (more
ochreous) and strongly marked, one greyer and weaker-
marked.
The pupa reminds of that of Gyno'pteryx gladiaria,
having nearly the same colour and markings, but it is of
course larger. The anal extremity has only two supple¬
mentary hooks, not six, as in gladiaria.
Pero (Herrich-Schaeffer).
Pero, Herrich-Schaeffer, Samml. Aussereur. Schmett., i, 28
(1855).
312
Mr. Louis B. Prout on the
Peryama, Herrich-Schaeffer, Samml. Aussereur. Schrnett.,
i, 28 (1855).
Azelina, Guenee, Spec. Gen. des Lep., ix, 156 (1858).
Si/ncmia, Guenee, Spec. Gen. des Lep., ix, 163 (1858).
1 Stenas2nlates, Packard, Monogr. Geom., 212 (1876).
Mcmnarea, Hulst, Tr. Amer. Ent. Soc., xxiii, 379 (1896).
I am generally loth to merge together genera which are
difterentiable on any structural character, but the group
which I am citing under the oldest name of Few, H.-S.,
seems — with the possible exception of Stenaspilates, Pack.
— so entirely homogeneous in all respects save the $
antennal structure, and a division upon that character
sometimes separates such evidently close allies, that I
feel constrained to regard the distinction as at most sub¬
generic. With the type of Stenaspilates {meskearia, Pack.)
I am not personally acquainted ; but Mr. J. A. Grossbeck
writes me that it differs little from Pew except in the ^
antenna. Stenodonta, Warr., Nov. Zook, xii, 379, probably
forms another section, unless equivalent to Stenaspilates ;
but as the ^ is unknown, and I have no material for study,
I leave it provisionally separate. It may be added that
the application of the names Pew and Azelina was long
reversed; vide Warren, Nov. Zook, xii, 367. The name
Pew belongs typically, as is there shown, to the species
with simply ciliated ^ antennae, while Butler, Tr. Ent.
Soc. Lond., 1882, 356-7, virtually restricted Azelina to the
pectinated species.
The Argentine representatives of this genus are mostly
of small size, in comparison with many of those from the
more northern parts of the continent.
Suhgenus I. — Pew, Herrich-Schaeffer. ^ antenna simply
ciliated.
199. Pero hoedularia (Guenee).
Azelina hoedularia, Guenee, Spec. Gen. des Lep., ix, 158
(1858).
Tucuman, in coll. Dognin.
A widely-distributed species, ranging from Central
America and the West Indies to southern Brazil. Guenee
described it from Brazil. I cannot see any appreciable
geographical variation, nor, indeed, any really noteworthy
individual variation.
Geometridac of the Argentine Bcimhlic.
813
200. Pero jonesaria (Schaus).
Azelina jonesaria, Schaus, J. N. Y. Eat. Soc., v, 162
(1897).
Buenos Aires, February 25, 1902, end of November to
December (A. F. Bayne), several in coll. L. B. Prout et
coll. A. F. Bayne ; Mendoza, captured and bred ex ovo,
one generation in December 1903, and another in April,
1904 (\V. M. Bayne); La Soledad, Entre Rios, in coll.
Rothschild.
Schaus’ type was from S.E. Brazil, and I have seen
topotypes in coll. E. D. Jones.
One might almost suspect, in this variable little species,
a much smaller southern representative of Pero variaria
(Walk.) = jctniaieensis (Schaus), but its range of variation
seems even wider than in that species. Besides the
habenaria-\ike forms (costa of forewing yellowish between
the lines) which occur in both sexes and apparently in all
localities with the type, I may mention that I have a
very red-brown $ from Buenos Aires, and a very weakly-
marked $ (the large black discal mark of forewing obsolete)
bred from Mendoza.
201. Pero fusca (Warren).
Azelina fusca, Warren, Nov. Zool., viii, 483 (1901).
Parana, Entre Rios (type $), in coll. Rothschild.
I know of no other examples, but am inclined to suspect
that the discovery of the $ will lead to the removal of the
species to the subgenus Azelina, which is the richer in
species in the Argentine Republic. P. fusca would seem to
have much affinity with the new species, P. amnicincta and
norma, described below. The type is in poor condition,
which might possibly account for its lacking some of the
characteristic markings of my new species ; but it shows
larger discal marks on forewing, and a somewhat differ¬
ently formed postmedian, and presents a sufficiently differ¬
ent aspect to disallow of their union.
Subgenus II. — Azelina, Guenee. ^ antenna bipectinate.
202. Pero gonopteraria (Guenee).
Azelina gonopteraria, Guenee, Spec. Gen. des Lep., ix, 160,
tab. vi, 5 (1858).
314
iNlfT Louis B. Front on the
]\Ietan, Salta, in coll. Dognin.
Common in some parts of Brazil, whence it was described
by Guende.
203. Peeo MATHILDA (Butler).
Azelina mathilda, Butler, Ann. Mag. Nat. Hist., (5) viii,
36 (1881).
Tucuman, in coll. Dognin.
Butler’s type was from Ecuador, and the species is also
very common in Peru. By some mistake, it has been
siudc to variaria, Walk. Walker’s type, a $ and without
locality, is certainly = jamaicensis, Schaus.
204. Pero eicurvata (Warren).
Peo'o hicurvata, Warren, Nov. Zook, viii, 490 (1901).
Tucuman (type), in coll. Rothschild ; Tucuman and
Los Vasquez, in coll. Dognin.
205. Pero palmirensis (Prout), nov. sp.
(Plate XLVIII, fig. 39.)
Pero {Azelina) ‘pahnirens.is, Prout, nov. sp.
36 mm. Antenna with moderately long, well ciliated pectina¬
tions. Hind tibia with the terminal spurs rather short. Forewing
with costa slightly arched at base, then straight or faintly concave
nearly to apex ; apex acutely pointed ; distal margin roundly
excised below apex, somewhat projecting at E' and sharply elbowed
at ; light brown, finely irrorated with pale and dusky scales
which give it a dull, slightly greyish tone ; lines brown ; ante-
median rather weak, from costa at 5 mm. to inner margin at 4 mm.,
outbent beneath costa, above M and in submedian area, curved
basewards at inner margin ; postmedian from costa at 14 mm. to
inner margin at 9 mm., rather straight, but faintly curved base-
wards at its extremities ; median area rather dark brown from the
cell-spot (which is small and rather weak) to the postmedian line ;
terminal area paler than basal. Hindwing with distal margin
slightly crenulate to Mk here produced, bluntly angled, thence
faintly sinuate ; pale, brownish grey, becoming slightly browner
terminally, especially in tornal region ; a dark brown outer line
from inner margin at about 2 mm. above tornus, pale-margined dis-
tally, distinct and rather thick (dark-shaded proximally) at first,
becoming finer and more indistinct, almost vanishing beyond IE ;
marginal line faintly darkened. Forewing beneath paler than above.
315
Geometridae of the Argentine BegyiMic.
antemedian line obsolete, postmedian rather distinct at costa, be¬
coming weaker, and vanishing after no other markings ; hind¬
wing beneath somewhat as above, but rather browner-shaded
costally, jjostmedian line reaching to costa, but nowhere very
strongly-marked or thickened, an indistinct diffuse discal spot.
Head (with palpus and antenna), body and legs concolorous with
wings.
Palmira, September 13, 1903 (W. M. Bayne). Type in
coll. L. B. Front.
Not very close to any species known to me. May best
be compared with Pero hicurvata, Warr, Nov. Zool., viii,
490, but is greyer in colour, the median area of fore¬
wing broader, postmedian line less incurved to inner
margin, marginal area unmarked, etc.
206. Pero olivacea (Warren).
Pero olivacea, Warren, Nov. Zook, xi, 578 (1904).
Tucuman (type), in coll. Rothschild ; Los Vasquez, in
coll. Dognin.
Quoted in M. Dognin’s MS. as a Pergama, from which
it may be assumed that the antennal pectinations are very
short, and surmounted with cilia, as in the following
species ; but I have no note on the point. Pectinations
certainly exist, being mentioned in Warren’s description.
207. Pero amnicincta (Warren, MS.) (Prout),
nov. sp. (Plate XLVIII, fig. 32.)
Pero (Azelina) amnicincta, Prout, nov. sp.
32-34 mm. Head and thorax greyish fuscous, abdomen
slightly paler. Antenna with short pectinations, surmounted with
fascicles of cilia (intermediate in structure between the present
subgenus and Pergama). Fore wing with produced apex, strong
tooth at end of and an angle at M^, below which the margin is
sinuate inwards ; greyish fuscous, densely sprinkled with blackish
scales ; lines blackish fuscous ; antemedian from costa at about
4 mm. (beyond one-fourth) to inner margin at nearly 4 mm.,
strongly oblique outwards from costa, forming a deep projection
into median area in cell, approximately perpendicular from M to
■inner margin, but with rather strong curves on submedian fold and
just below SM2, as in many of the genus ; postmedian from costa
at nearly four-fifths to inner margin at or before two-tbirds, deeply
incurved between and submedian fold, and again close to inner
316
r. Louis B. Prout on the
margin, dark-shaded proximally ; discal spot minute, black ; mar¬
ginal line fine, blackish, slightly interrupted. Hindwing rather
strongly crenulate, with the longest projection at ; paler than
forewing, except inner margin, which is strongly dark shaded, the
dusting concentrating into a small irregular blotch close to tornus ;
a postmedian line beginning on inner margin near tornus, but
becoming faint, and almost disappearing after middle of wing ;
traces of a fine black marginal line, a series of distinct black dots
placed on it between the veins, those between and and on
either side of the submedian fold the strongest. Underside of fore¬
wing paler than upper, but somewhat darkened and more dusted
along costa and in apical region ; an elongate blackish discal spot,
and the beginning of a tine dark postmedian line from costa.
Underside of hind wing rather strongly dark-speckled, with large
black cell-spot, which is nearly bisected with pale at the fold
(position of R^), and with a strongly sinuate fuscous postmedian
line.
Buenos Aires, October 10, 1905 (A. F. Bayne). Type
in coll. L. B. Prout. A second Mendoza, March 1908,
is clearly of the same species, though rather worn, and
with the median area of forewing considerably narrower —
antemedian making, from costa, a stronger bend towards
postmedian, and thenceforward further from base, while
the postmedian bends slightly more basad towards inner
margin. I also saw a specimen of the species in coll.
Rothschild some years ago — I think from Tucuman —
bearing the MS. name of Pero amnicincta, Warr., which I
have adopted. I have seen almost all the described species
of this extensive genus, and have studied the descriptions
of the few others, and cannot find any agreeing with the
present insect or the following, which may conceivably be
a form of it.
208. Pero norma (Prout), nov. sp.
(Plate XLVIII, fig. 1.)
Peo'o {Azelina) no^’ma, Prout, nov. sp.
33 mm. Head and body nearly unicolorous light brownish
grey, the face in part and the palpus darker. Antenna as in
P. amnicincta. Forewing divided into three contrasted areas by
the transverse lines, the basal and terminal greyish-fawn colour,
sprinkled with blackish, the median tawny, very weakly sprinkled
with darker, and darker shaded distally ; antemedian line almost
Geometridm of the Argentine Bepuhlie. 317
as in amnicincta, the projection in cell not quite so deep ; post¬
median from about four-fifths costa to two-thirds inner margin,
forming a slight projection between and R^, irregularly incurved
between R® and submedian fold, and again at SM^ ; discal dot
small, black, with a few dark scales behind it ; marginal line not
appreciably darkened. Hindwing pale grey tinged with brownish,
a tawny brown postmedian line beginning from a blackish spot on
inner margin near tornus, but becoming faint, and almost dis¬
appearing after middle of wing ; a small fuscous cloud on termen
from tornus to M^, and three blackish marginal spots, one between
AH and AP, and one on either side of submedian fold. Underside
uniform pale grey, tinged with brownish ; forewing with faint
traces of darker postmedian line, which is pale-shaded distally, and
of an elongate cell-spot ; hindwing with a very weak, wavy post-
median line, and a somewhat crescentic (or angulated), but rather
indefinite, blackish cell-spot.
Rodeo del Medio, May 20, 1904 (W. M. Bayne). Type
in coll. L. B. Front.
The above description has been drawn up from the type
^ alone, because I felt uncertain whether a couple of ^s
in coll. Br. Mus. belonged to the same species ; but a
further ^ received, during the progress of this work, from
my kind correspondent Dr. E. Giacomelli, satisfies me
that this is the case, and I am able to add that the species
is evidently a variable one. If it be further conspecific
with F. anmicincta, it is still more variable, for that has
an altogether darker coloration, and a very much deeper
sinus in the postmedian line of the forewing.* The ^s
lack the tawny median area of the type, have the ante-
median line of forewing rounded rather than angled in
the cell, the postmedian rather straighter, the underside
of the hindwing more strongly dusted with fuscous, and
w'ith its discal spot better expressed. But my specimen
from Dr. Giacomelli has a weakly tawny shade over the
basal and median areas, darkening a little towards the
postmedian line, and in one or two other respects serves
to connect my ^ with the $s which I had previously seen.
Accepting, then, all the four examples as belonging
together, the following localities are to be added : —
Gran Chaco, near Florenzia, October 1902 (S. R.
* That members of the present genus can vary considerably in
this latter particular is proved by P. Vignata, Warr. ; but the
divergence between amnicincta and norma is even more extreme.
318
i^^^^JOuis B. Prout on the
Wagner), ^ in coll. Br. Mus. ; Goya (collected by Perrins,
presented by Lord Walsingham), ^ in coll. Br. Mus. ; La
Rioja (E. Giacomelli), ^ in coll. L. B. Prout. The species
therefore has a wide distribution in the Argentine
Republic.
Subgenus III. — Fergama, Herrich-Schaeffer. $ antenna
dentate-fasciculate.
209. Pero lignata (Warren).
Fergmna lignata, Warren, Nov. ZooL, iv, 499 (1897).
Fergama lignata ab. obscurior, Warren, Nov. Zooh, iv, 500
(1897).
? Tucuman, etc.
Specimens in coll. Br. Mus. (“Argentina, O. W. Thomas ”)
and in coll. Bastelberger (Tucuman) have been determined
as belonging to this species, and seem to agree well enough
with it ; but they may also be referable to the following.
I am by no means clear that the two are not geographical
races of one widely-distributed species.
Warren described his types and other examples from
Venezuela, Trinidad, Colombia and the Amazons. The
species seems pretty generally distributed in Central
America, Ecuador, Peru, the Guianas, etc., and the British
Museum possesses one from Paraguay.
210. Pero uniformis (Warren).
Fergama uniformis, Warren, Nov. ZooL, iv, 500 (1897).
Los Vasquez, in coll. Dognin ; Argentina ( J. H. Leech),
without nearer indication of locality, in coll. Br. Mus.
Warren’s type was from S.E. Brazil.
211. Stenodonta incurvata (Warren).
Stenodonta incurvata, Warren, Nov. ZooL, xii, 379 (1905).
Suncha Corral, Santiago del Estero (type, ^), in coll.
Rothschild.
212. Cnephora griseata (Maassen).
Azelina griseata, Maassen, Stiibel’s Reisen, 156, tab. viii, 7
(1890).
Metan, Tucuman and Los Vasquez, in coll. Dognin.
Maassen’s type was from Colombia. Also known to me
from Peru.
Geometridae of the Argentine Bepuhlic. 319
Salpis (Mabille).
Honorana, Butler, Tr. Ent. Soc. Loud., 1882, 365 (1882),
riec Blanchard.
Salpis, Mabille, Bull. Soc. Philom. (7) ix, 65 (1885) ;
Miss. Sci. Cap Horn, vi, Lep. 22 (1888) (nov. syn.).
Lasiops, Warren, Nov. Zool., ii, 143 (1895) (noin. praeocc.).
Pseudosalpis, Staudinger, Hamb. Magalh. Sammelr., ii, Lep.
82 (1899) (nov, syn,).
An interesting genus of rather stoutly-built moths, with
densely hairy clothing, occurring chiefly in Chili, the
southern Andes and the extreme south of South America,
though one species is known from Peru. Except in the
shape of the wings, it seems to come very near Macro-
lyrcea, Butl., Tr, Ent. Soc. Bond., 1882, 349, but I do think
it would be desirable to unite the two. On the contrary,
the present genus will divide readily into at least three
subgenera, which it is by no means improbable may later
take full generic rank.
That l)r. Butler misidentified Blanchard’s figure of
Honorana notaturia,* the type of the genus Honorana
seems to me certain, and Sir George Hampson and Mr.
Warren are entirely in accord with me. What species
M. Blanchard had before him remains at present a mystery,
as nothing like it seems to be known; perhaps it is not
even a Geometrid. The Honorana notaturia of Butler
(vix descr.) will require a new name and an adequate
diagnosis, 'but the present is not the occasion; both it and
Honorana aenea, Butl., Tr. Ent. Soc. Bond., 1882, 365,
must be transferred to the genus Salpis.
As I have given some attention to the genus, and have
new species to introduce in each of its subgenera, a full
generic diagnosis will not be out of place.
Salpis, Mabille. — Face somewhat protuberant, densely
rough-scaled ; palpus I’ather long, densely haired (long¬
haired in most species), 3rd joint elongate, cylindrical or
somewhat fusiform, with appressed scales ; tongue strong ;
antenna long, in the typical section nearly as long as
forewing. Thorax usually robust, pectus densely hairy.
Femora hairy, hind tibia with all spurs present, $ hindtibia
much dilated, with strong hair-pencil ; tarsi strongly spinu-
lose. Abdomen long, in $ stout. Wings densely scaled,
* Blancliai’d, Gay’s Hist. Chile, vii, 92, tab. vii, 8.
320
\i^Loiiis B. Prout on the
Fieiinlum present. Forewing with cell more than one-
half, C free, SC^ free, SC^ from cell anastomosing with
gQ3-5 fi-0,1) before apex of cell. Hindwing with C
approximated to SC to one-half of cell, SC^ separate from
R^, AH separate from R^. The following subgenera are
differentiable : —
Suhgenns I. — Saljns, Mabille. Antenna extremely long,
in ^ simple or simply ciliated. Terminal joint of palpus
long. Forewing rather broad, apex not produced : einten-
oiata, Mab., aenea, Butl., infelix, Butl.,* p^iechi, Dogn.,j‘
carneitincta, Prout, ruhens, Prout.
Suhgenns II. — {Lasioi^s, Warren, nom. praeocc.) Micro-
dontopera (Warren, MS.), Prout, nov. subgen. Antenna
moderately long, in ^ shortly or moderately bipectinate.
Terminal joint of palpus long, P^orewing not broad, its
apex acutely produced : J chilenaria, Felder,| felderi,
Butl., II elentilinenta, Butl.,^ eiidora, Prout.
Suhgenns III. — Antygophanes, Prout, nov. subgen. ^
antenna very shortly bipectinate. Terminal joint of palpus
moderate. Abdomen very long. Forewing narrow : orhi-
fera, Prout.
Note. — I have, unfortunately, not seen Salpis antenncita,
Mab., the type of the genus, but Mabille’s figure and de¬
scription leave no doubt that it belongs to the aenea group.
Staudinger, Hamb. Magalh. Sammelr., ii, 81, 82, evidently
unacquainted with this group of moths, rashly accuses
Mabille of error in referrino' it to the Geometridae, and
erects a new genus, PseiidosaJpis, on the type Salp>is alhi-
pxinctaria, Mab., Nouv. Arch. Miis. Paris (3), i, 155. This
species, which also I do not know in nature, seems quite
rightly placed in the genus, but as I am not quite sure as
to the subgenus, I have not cited it in the above survey.
Salpis scodionoeata , Mab., Nouv. Arch. Mus. Paris (3), i,
* Sahvludeii infelix, Butl., Tr. Ent. Soc. Loud., 1882, 350.
t Lasiops {!) puechi, Dogn., Ann. Soc. Ent. Belg., xlviii, 368.
X Other slighter and less constant differences exist in the less
dense scaling (less mixed with hair-scales), less crenulate margin of
hindwing, somewhat more slender terminal joint of palpus, occasional
approximation or even anastomosis of SC of forewing with C, closer
approximation of to in hindwing, etc.
§ Colotois (?) chilenaria, Felder, Reise Novara, Lep. Het., tab.
cxxiv, 6.
II Azelina felderi, Butler, Tr. Ent. Soc. Bond., 1882, 355, tab.
xvi, 5.
IF Monoctenia dentilineata, Butler, Tr. Ent. Soc. Bond., 1882, 358,
tab. .xvi, 12.
Gcometridae of the Argentine RepuUie. 821
15G, tab. xi, 5, also referred by Staudinger to Pseudosalijis,
seems more doubtful generically, though it may quite
conceivably be a more slenderly-built Salpis.
213. Salpis carneitincta (Front), nov. sp.
d $• 37-40 mm. Very similar to o,enea, Butl. (Honorana),
from Chili, but smaller, costa of forewing faintly concave, ground¬
colour tinged with fleshy, the large black discal spots of both wings
and the black terminal line absent, other lines less distinct, some¬
what diffuse, the white vein-dots of postmedian line distinct, a
characteristic whitish line traversing the base of the very long
fringes ; underside nearly concolorous with up2)er (in aenea whitish,
especially on hindwing). Antenna very long, in d almost simple,
minutely pubescent.
Puente del Inca (W. M. Bayne), 2 d, ¥ in coll. L. B.
Front, 2 dj 2 ¥ in coll. A. F. Bayne. The type d (in coll.
L. B. Front) was picked up dead and stiff, yet in good
condition, in March 1902 ; one d and one ¥ are dated
February 17, 1904; another d March 16, 1905.
One pair in Mr. Bayne’s collection may be regarded as
an aberration, or possibly a very closely related species,
showing distinct dark discal spots and moderate dark
shading to the transverse lines, in these respects making
an approach to S. aenea.
Some of the distinctive characters given above for S.
carneitineta are shared by S. infelix, Butl. {Sahidodes), but
that is a darker insect, with the distal margin of hind¬
wing more deeply toothed, postmedian line nearer termen
and differently formed, and numerous other differences.
M. puechi, Dogn. {Lasiops ?), from Peru, is also very close
to carneitincta. I have seen M. Dognin’s unique type,
although I cannot now compare it ; but it should be dis¬
tinguishable by its greenish grey tone, distinct postmedian
line on upper surface of hind wing, absence of whitish line
on the base of the fringes, etc.
214. Salpis rubens (Prout), nov. sp.
(Plate XLVIII, fig. 31.)
(J. 37-44 mm. Exceedingly like the preceding species, but with
the forewing somewhat more produced,* the ground-colour somewhat
* Five specimens have been carefully measured, and have given
the following result, reduced to an assumed costal length of
TRANS. ENT. SOO. LOND. 1910 — PART III. (noV.) Y
322
M^Louis B. Prout on the
brighter, the transverse lines slightly better expressed, sometimes
Ijlackish-marked on the veins, the fringe less extremely long and
more variegated with paler and darker, its basal line pinkish, not
whitish. The hindtibia is perhaps less strongly thickened, but the
surest structural distinction is in the antenna, which is well
ciliated (the cilia in fine fascicles, their length nearly equal to the
width of the shaft), whereas that of carneitincta is virtually simple.
Zaiijon Amarilla (W. M. Bayne), type and another ^ in
coll. L. B. Prout, both dated November 16, 1905 — one, at
least, at light; Puente del Inca (W. M. Bayne), a slightly
worn $ in coll. A. F. Bayne. The dates, so far as available,
suggest tliat S. ruhens occurs much earlier in the season
than S. carneitincta (the former November, the latter
February and March).
215. Salpis Seudora, Prout, nov. sp.
(Plate XLVIII, fig. 17.)
Salpis (Microclontopera) cudora, Prout, nov. sp.
^ . 38 mm. Pafiius with 3rd joint long (about 1 mm.), fusiform ;
antenna with short, strongly clavate pectinations to beyond two-
thirds, then serrate-dentate. Face grey ; palpus whitish-grey,
marked with fuscous, more strongly above ; antenna mostly ferru¬
ginous, its dorsal scaling grey ; legs whitish grey mixed with fuscous,
the tarsi mainly fuscous, each joint tipped with whitish grey ; thorax
and abdomen nearly concolorous with fore- and hindwing. Fore¬
wing with costa nearly straight, apex acute, distal margin weakly
crenulate ; brownish grey, coarsely irrorated with dark fuscous,
browner on the veins and slightly darker-clouded in distal part ; the
lines fuscous ; antemedian thick, from nearly one-third costa to
beyond two-fifths inner margin, making strong outward curves in
cell and in submedian area, the angles on M and SM^ accentuated by
blackish dots ; postmedian from four-fifths costa, rather weak, but
rendered prominent by a series of blackish vein-dots, each accom¬
panied distally by a pale vein-dot ; the dots on and on are
placed more proxiniad than the rest of the series, and the line itself
is lunulate between the dots ; a terminal series of small blackish
dots between the veins; fringe nearly concolorous with wing,
20 mm., which is the actual measurement of the largest specimen
{rubens) : —
Costa. Inner margin. Distal margin.
(S', rnbens . . 20 13J- to 14 13 to 13|
S. carneitincta 20 15 „ 10 13| ,, 15
Geometridae of the Argentine EejpuUie. 323
weakly paler at its base. Hindwing witli costa ratlier long, apex
moderately pronounced, distal margin weakly crenulate ; white,
with a few fuscous scales in basal part and moderately dusted with
fuscous in distal half; cell-spot rather large, dark fuscous ; a fuscous
postmedian line from inner margin close to tornus to costa at about
three-fourths, sinuate basewards in submedian area and very weakly
between radials, slightly projecting on M* and R^, darker-spotted
on veins ; terminal line very fine, greyish fuscous, with dark dots
between the veins as in forewing; fringe concolorous with wing.
Forewing beneath whitish grey, more brownish costally and distally,
costal region speckled with fuscous ; a small cell-spot and a
moderately distinct postmedian. Hindwing beneath nearly as
above.
Puente del Inca, bred November 12, 1903 (W. M.
Bayne), type in coll. L. B. Prout; a second $ from the
same locality, December 15, 1905, in coll. L. B, Prout;
others, both sexes, in coll. A. F. Bayne ; Balde, one $ in
coll. A. F. Bayne.
The $ seems to have the hindwing less light, and
perhaps shows other slight differences ; unfortunately
I omitted to make detailed notes on Mr. Bayne’s $s. I
possess one myself, which I assumed to be referable to this
species, but which may rather be a very close ally, as I
find, in addition to the more superficial distinctions, that
it has a shorter 3rd joint to the palpus than have my ^s.
The specimen is larger (44 mm.), browner, more strongly
dark-dusted, particularly in the hindwing, the lines are
rather weak, the postmedian in both wings straighter, the
cell-spot of hindwing wanting above. Mr. W. M. Bayne
supplied upon it the following note : “ kil. 168, Transandine
Railway, three pupae under stones. Two have emerged,*
the other remains ; I have kept these, hoping the other
might be a but have had to kill them.” I suspect the
second of the $s must be a specimen in coll. A. F. Bayne
on which I made the note that it was rather larger (^. e.
than typical specimens), “ with hindwing a little darker and
with other slight differences, 2^ossiblg a distinct species;
but I cannot at the moment follow the matter up. To
judge from the few examples which I have seen, the
Chilian group to which S. eudora is the most nearly
related (/S', ehilenaria, Feld., etc.) show a good deal of sexual
dimorphism, as well as of individual variation, and it is
* Mine is dated as bred November 3, 1905. — L. B. P.
Y 2
324 Al^TjOuis B. Front on the
difficult to do iimcli sound revisional work without larger
material.
My type $ Avas bred from a cocoon which Mr, W. M.
Bayne found attached to the underside of a stone, and I
believe a few of the other examples were obtained in the
same manner. He notes that the moth sits Avith the
Avings closed in a more or less Deltoid or Noctuid posture.
210. Salpis orbifera, Front, nov. sp.
(Flate XLVIII, fig. 23.)
Salpis {Antygophancs') orhifera, Front, nov. sp.
g. 30 mm. Palpus moderate, densely hairy, without the very
elongate, fusiform terminal joint which characterizes true Salpis.
Antenna Avith very .short subclavate pectinations. Third joint of
palpus whitish beneath. Head and body fuscous, mixed Avith paler
and blacker scales. Fore wing with costa nearly straight, ape.x
pronounced, but not produced, distal margin oblique, gently boAved,
almost smooth ; fuscous, irrorated with darker fuscous and with
whitish, the markings blacker ; antemedian line almost obsolete,
apparently sharp-angled in cell, thence very oblique and more or
less denate to one-third inner margin ; postmedian sinuous and
oblique, from near apex to two-thirds innej- margin, the pale scales
rather more numerous di.stally to it than proximally; a circular pale
spot, AAuth fine blacki.sh circumscription and some dark scales in
centre, at end of cell, bisected by tlie discocellulars ; fringe con-
colorous with Aving, chequered with slightly paler patches between
the veins. Hindwing with costa long and nearly straight, distal
margin nearly smooth, paler fuscous, with rather large dark discal
•spot. Underside jjale fuscous, costa of forewing and whole of hind¬
wing irrorated with dark scales ; both Avings with faint traces of a
postmedian line ; hindwing with discal spot as above.
Cacbeuta, January 12, 1904 (W. M. Bayne). Type in
coll. L. B. Front.
A very distinct species. Frobably its palpus and other
characters (wing-shape, etc.) are enough to indicate its
generic distinctness from Salpis, yet it has also very much
in common therewith, and especially Avith the subgenus
Microdontop)era.
Arizela (Front), nov. gen.
Face not protuberant, smoothly scaled ; palpus rather .short, por-
rect, shortly rough-scaled ; tongue apparently Avanting ; antenna
Geomctridac of the Argentine Ecjnchlic. 325
scarcely one-half the length of forewing, in hotli sexes shortly
bipectinate nearly to apex, the pectinations very short in the 9,
apical segments nearly simple, with projecting scales and short
ciliation. Thorax hairy beneath ; femora somewhat hairy ; hind-
tibia not dilated, in both sexes with four rather approximated spurs ;
abdomen rather robust, not crested. Wings rather long and narrow,
thickly but smoothly scaled, frenulum present. Forewing with
costa nearly straight, apex not acute, distal margin entire, gently
rounded, oblique ; cell little over one-half, discocellulars incurved,
SCi'2 long-stalked, their stalk anastomosing with C, SC^ later ap-
pressed to SC^'^, but not anastomosing, SC^'^ from before apex of
cell, R* and R^ curved, M* from rather near R^, from beyond
two-thirds of cell. Hind wing with C anastomosing strongly with
cell, SC^ connate with R^, medians as in forewing.
Type of the genus ; Arizeki dnlcis, Prout, nov. sp.
Apparently related to Hasodima, Butl., Erilophodes,
Warr. and, Leueolithodes, Warr., but differing markedly in
the strong anastomosis of the costal and subcostal of hind¬
wing, in various other points of venation, head-structure
(with antenna), etc., so that the relationship can hardly be
very close,
217. Arizela dulcis (Prout), nov. sp.
(Plate XLVIII, fig. 27.)
. 28 mm. Face and vertex brown ; palpus brown, tipped with
white ; antennal shaft white above, brown spotted, brown beneath, pec¬
tinations brown. Thorax white, narrowdy brown anteriorly, legs white-
and brown-banded. Abdomen whitish, with single dorsal and double
ventral series of brown spots. Wings silky white, the markings
brown, more or less mixed with grey. Forewing narrowly dark at
base ; antemedian line before one-third, somewhat curved, tei’minat-
ing abruptly below M, reappearing only as a large spot on SM^; post¬
median broad, waved, from three-fourths costa to two-thirds inner
margin, parallel with termen, interrupted between R^ and R^ ; a
costal blotch closely preceding antemedian, a second (grey-centred)
in middle, abf5orbing the cell-spot ; an angulated mark from sub¬
median fold to inner margin, closely preceding postmedian line ; a
few subterminal .spots; some dots in costal area and proximally to
postmedian line ; fringe white, broadly chequered with brown from
apex to M*, thence unicolorous. Hind wing with a few costal and
terminal spots, the most con.spicuous being at termen between R* and
R^ ; a very small discal spot, slightly above middle of discocellulars ;
remnants of a postmedian band from costa to R^ (just distally to the
82G
]\I^Louis B. Front on the
cell) and again from before to subincdian fold ; fringe white,
unchequered. Underside the same.
San Luis, February 21, 1904 (W. M. Bayne). Type ($)
in coll. L. B. Front. A ^ precisely similar from Palmira
in coll. A. F. Bayne.
218. Ehilopjiodes colorata (Warren).
Eriloiilwdcs colorata, Warren, Nov. Zool., i, 466 (1894).
Mascora variegata, Schaus, MS., in coll. F. D. Jones.
“Argentina” (J. H. Leech), in coll. Br. Mus., an aberra¬
tion ($) with the ground-colour of both wings strongly
dark-dusted. Described from S.E. Brazil.
219. Leucolithodes arana (Dognin).
GariiJcta arana, Dognin, Ann. Soc. Ent. Belg., xxxix, 117
(1895).!
Erilo2diodcs arana, Warren, Nov. Zool., xi, 582 (1904).^
Tucuman, in coll. Dognin.
Dognin (^) described the species from Ecuador. It
occurs also in Colombia.
Warren (^) treats it as an Erilophodcs', though com¬
menting on its close “superficial” resemblance to Leuco-
lithodes pantcata (Feld.),*^ he overlooks the fact that it
virtually lacks the abdominal tufts of Erilophodcs, and also
agrees in venation with Leucolithodes. It is therefore best
treated as forming a second section of that genus, with the
J antenna bipectinate.
220. Leucochesias mesargyrata (Mabille).
Lcncochcsias nusarqyrata, Mabille, Nouv. Arch. Mus. Paris,
(3) i, 158 (1889).t
Aspilates (?) nimipennaria, Maasseu, Stiibel’s Beisen, Lep.
98, 162, tab. ix, 16 (post August 2, 1890) (nov. syn.).
Port Santa Cruz, E. coast of Patagonia (Mabille) ;
Uspallata Pass, 7000 ft. (Geo. Boag), in coll. Br. Mus.
* Bryoptera panteata, Feld., Reise Novara, Lep. Het., tab. cxxvi,
V = Leucolithodes pantherata, Warr. (in err. jiro panteata), Nov.
Zool., xi, 582.
t Possibly not published till 1890, as it is indexed in the
“Zoological Record” for that year, and the copy at our Natural
History Museum was not received until May 13, 1890. But even
this, the latest possible date, gives some months’ priority over
Maassen.
327
Geomctridcie of the Argentine Rcpuhlic.
Mabille also records the species from Punta Arenas,
which belongs to Chili. Maassen’s type was from the
highlands of Bolivia.
221. Odontothera (?) cruciferaria (Berg).
Aspilates cruciferaria, Berg, An. Soc. Cient. Argent., iv,
206 (1877).!
Spartopteryx (?) denticulata, Butler, Tr. Ent. Soc. Bond.,
1882, 387 (1882) (nov. syn.).^
Aspilates glyphicarms, Mabille, Bull. Soc. Pliilom. (7) ix,
66 (1885) ; Miss. Sci. Cap Horn, vi, Lep. 23, tab. ii,
10 (1888).^
(^) Rio Santa Cruz, Patagonia (F. P. Moreno).
Also occurs in Chili and Tierra del Fuego (^).
Apparently variable, but I have only seen a very few
specimens.
I know of no existing genus to which this species seems
satisfactorily referable. It differs from typical Odontothera,
Butl.* — which, by the way, was not intelligibly diagnosed,
being compared to certain Larentiid genera — not only in
having the ^ antenna bipectinate, but also in some points
of venation, etc. ; nevertheless it comes rather near it in
most respects, and as it is necessary to assign it some
provisional generic position, I consider Odontothera the
most satisfactory that I can find. I imagine both may
be related, though probably not very closely, to the
Ischnopteryx — Cidariophanes group.
222. Callipseustes trisecta (Warren).
Callipseustes trisecta, Warren, Nov. ZooL, xiv, 287 (1907).
Ciudad de Tucuman (type), in coll. Rothschild ; also
from the same locality in coll. Dognin.
223. Amblurodes chloroclystata (Guenee).
Syrtodes chloroclystata, Guenee, Spec. Gen. des Lep., x,
452 (1858).
Gidaria cldorophaearia. Walker, List Lep. Ins., xxxv, 1690
(1866).
Syrtodes hryifera, Felder, Reise Novara, Lep. Het., tab.
cxxxi, 28 (1875).
* Tr. Ent. Soc. Lond., 1882, 409,
328
MiTLouis B. Front 07i the
Los Vasquez, in coll. Dognin.
Guenee’s types were from Brazil. I am not at all sure
that the forms from the more northern parts of the
continent really belong with it. The synonymy is given
as it has been supplied to me by Mr. Warren (in litt.),
since I have not been able personally to investigate it.
Walker’s type of chlorophaearia was from Bogota; Felder’s
hryifei'a (a from French Guiana.
224. Ischnopteryx xylinata (Guenee).
Syi'todcs xylinata, Guenee, Spec. Gen. des Lcp., x, 452
(1858).
Tucuman, in coll. Dognin.
Also described from Brazil, where it seems to be a rather
frequent species.
PSEUDOBREPHOS (Prout), nov. gen.
Face protuberant, with a bluntly-pointed, horny projection partly
concealed by rough hairs ; palpus moderate, densely rough-scaled,
3rd joint blunt, smoother-scaled, almost concealed by long hair-
scales of 2nd joint ; tongue apparently short ; antenna moderate, in
^ bipectinate almost to apex with rather short, stout, subclavate
pectinations. Thorax rough-haired above, somewhat crested pos¬
teriorly ; pectus hairy ; (fore- and hindlegs wanting) ; abdomen
moderately robust, not crested. Wings densely scaled, distal margins
nearly smooth, faintly waved; frenulum developed. Forewing
triangular, inner margin rather long, termen little oblique ; cell
rather long ; SC^ free, SC^ from cell, anastomosing at a point with
point of departure of SC^ from which is well removed from
cell ; Ml from close to R^, from rather near end of cell. Hind¬
wing with cell scarcely over one-half, C approximated to cell to near
one-half, thence gradually diverging, SC^ connate with R', medians
as in forewing.
Type of the genus : Tf^euddh^'cphos, haynei, Prout,
nov. sp.
Related to Catophocnissa, Warr., Nov. Zook, i, 464, from
Chili, but distinguished by the longer and sharper frontal
prominence, the absence of abdominal crests and the non-
* This would probably vary somewhat in other specimens, and
need not be insisted upon.
Geometridac of the Argentine Reimhlic. 329
crenulated distal margins of the wings. The venation is
practically identical, for the point of anastomosis of SC^
is no doubt individual ; in the Catophoenissa which I have
examined, this vein anastomoses normally with SC^‘‘
heyond SC^, or remains free.
225. PSEUDOBREPHOS BAYNEI (Prout), nov. sp.
(Plate XLVIII, fig. 10.)
27 mm. Forewing whitish, strongl}’’ irrorated and marked
with fuscous and dark fuscous, leaving only narrow spaces proxi-
mally to antemediau line and distally to postmedian white, the
latter, however, widening into a patch costally, its limit on costa
being near apex ; lines black ; a basal, angulated in cell, reaching
only to submedian fold ; an antemediau, oblique outwards from
costa, making strong projections into median band in cell and in
submedian area ; a postmedian line from beyond two-thirds costa,
strongly inbent distally to cell, sending three nearly equal teeth
outward on R^, IVP and M^, deeply incurved in submedian area, and
acutely angled outward on SAP ; the area between the lines rather
uniformly dark-shaded, suggesting a median band; termen with
black spots between the veins. Hindwing orange, fuscous-clouded
at base and along margins as far as median band, and rather broadly
bordered with dark fuscous ; a narrow dark fuscous median band,
outangled at posterior angle of cell, and bent towards anal angle
posteriorly; a fine black terminal line. Underside of forewing
whitish, somewhat sprinkled with fuscous, fuscous-shaded along
costa, inner margin and in terminal one-fourth ; a broad dark fuscous
tranverse bar at just beyond one-half (crossing the cell near end), and
faint indications of a second bar margining the terminal shading
proximally ; an orange suffusion in basal half of wing. Underside
of hindwing whitish, sprinkled with fuscous, an angulated transverse
band and dark border nearly as above, the latter weaker and more
diffuse ; no orange suffusion.
Buenos Aires, April 1, 1907 (A. F. Bayne), type in coll.
L. B. Prout.
I know of no second example of this pretty and interest¬
ing little species, which, in its general scheme of colouring,
reminds a good deal of the Chilian Catophoenissa dihapha,
Feld.,* in miniature.
* Epimecis dihapha, Feld., Reise Novara, Lep. Het., tab. cxxv,
10, 10a.
330
Mi^jouis B. Prout on the
226. Catascia carnea (Prout), nov. sp.
(Plate XL VIII, fig. 24.)
46-50 ram. Face roumled-promiiient; palpus quite moderate,
rather stout; tongue well developed ; antenna rather long, with
short ciliation. Legs and abdomen slender. Head, body, wings,
etc., rather uniform flesh-colour, wings irrorated witli darker scales,
slightly paler beneath. Forewing with termen rather more oblique
than is typical for the genus (shaped much as in narrow-winged s
of C. andereggaria, Lah., of Europe) ; lines grey, indicated very
faintly except on veins, where they are made rather conspicuous by
dark spots ; antemedian from costa at about one-fourth, inbent about
SM2 ; postmeJian from costa at about 3 mm. from apex, slightly
oblique inwards to E^, outbent to R^, deeply incurved between
and SM^, and reaching inner margin at about two-thirds ; cell-spot
distinct, but small ; no terminal line. Ilindwing somewhat paler
in cell and costally, without markings. Underside without markings.
Puente del Inca, April 12, 1904 (W. M. Bayne), type in
coll. L. B. Prout; a second ^ from same locality, April
1902 (A. F. Bayne), in coll. L. B. Prout ; two from same
locality in coll. A. F. Bayne.
This interesting insect is probably a true Catascia, by
which name I understand Lederer’s section C of Gnophos
{mde Ent. Kec., xvi, 121). The contour of the distal
margin of both wings is very closely that of this group, the
frons, though less strongly protuberant than in some of
them, agrees fairly well with others, while several other
characters, together with the general facies and the
mountain habitat, lend some further support to the generic
position here suggested ; it would fall into Lederer’s first
subsection, with C. myrtillata, Thunb., on account of the
simply ciliated ^ antenna; the ^ is still unknown. Only
the venation of the forewing is not that of typical Catascia ;
SC^ anastomoses shortly with C, SC^ (arising from cell)
shortly with SC\ and subsequently at a point (or in one
specimen by a short connective bar) with SC^‘^ ; in all the
European Catascia which I have examined, SC“ is free
from SC^,* and except in opcraria, Hiib., ancj andereggaria,
Lah., SC^ is also free from C. The inconstancy of the sub-
costals in many Boarmiids is, however, matter of common
knowledge.
* But I find that Meyrick, Tr. Ent. Soc. Lond., 1892, 136, has
recorded eight myrtilldta {ohf'dscaria) with these veins not tree, and
(page 138) one iveni witli triple anastomosis as in carnea.
331
Gcomctridae of the Argentine Repuhlic.
Mr. A. F. Bayne discovered Cataseia carnea on his first
visit to Puente del Inca, of which some account is given
in Trans. City Lond. Ent. Soc., xii, 70-72. It was flying
over some low, thorny bushes at dusk, in company — as was
subsequently made out — with a species of Chrismoj^teryx
and perhaps some other Geometrids.
227. Gnophos (?) OREAS (Prout), nov, sp.
(Plate XLVIII, fig. 9.)
5 . 30 mm. Head, body and wings grey, upper surface of wings
coarsely speckled with brownish fuscous, the markings somewhat
ill-defined and mostly diffuse, formed by accumulations of the fuscous
speckling. Forewing with a fine, curved line from one-fifth costa to
one-fourth inner margin ; an ill-defined median band from before
one-half costa to about one-half inner margin, narrowed at costa,
absorbing in its distal part the large but very ill-defined dark cell-
spot ; a second, somewhat narrower band proximally to the pale
dentate subterniinal ; the ill-defined beginning of a postmedian at
costa midway between the bands ; a terminal dark line ; fringe dark-
marked at the vein-ends. Hind wing with a somewhat dentate
median line, accompanied proximally by an ill-defined band con¬
tinuing that of forewing and absorbing the cell-spot ; subterminal
line and its accompanying band nearly as in forewing. Underside
more weakly dark-speckled, both wings with large, roundish dark
cell-spot and dark submarginal band ; terminal dark line distinct.
Cacheuta, January 12, 1904 (W. M. Bayne). Type in
coll. L. B. Prout.
Evidently rather closely related to Gnophos insolita
(Warr.) * and chanehani (Dogn.) f, forming with them a
compact group which may provisionally — after Warren —
be treated as a section of Gnophos, Tr. = Scotopterix, Hiib.,
with veins SC^-^ coincident, as in Sienalcidia, etc. The
termen is less crenulate than is typical, the $ antenna
bipectinate with long pectinations, but the wing-form and
pattern and the absence of fovea in $ distinguish them
from Stenaleidia. The frons is less protuberant than in the
European species which I have studied, but if the con¬
stitution of the genus as given in Hampson’s “ Moths of
* Scotopterix insolita, Warr., Nov. Zool., viii, 479 = Cyniatopliora
{!) fumigrisea, Dogn., Ann. Soc. Ent. Belg., 1, 211 (nov. syn.).
f Cymatophora Ci) chanehani, Dogn., Ann. Soc. Ent. Belg., xlviii,
364.
382
M^^iouis B. Front on the
India” can be inaintainetl, still more extreme instances of
flattened Irons can be found.
228. Odysia molaria (Guenee).
^ Odysia molaria, Guenee, Spec. Gen. des Lep., x, 4,
tab. xvi, 6 (1858).
Boarmia dcnticularia. Walker, List Lep. Ins., xxi. 362
(I860).
Tephrosia inefectaria, Walker, List Lep. Ins., xxi, 412
(1860).
Tephrina siqncdaria, Walker, List Lep. Ins., xxvi, 1657
(1862) (ab.).
Tephrina responsaria, Walker, List Lep. Ins., xxvi,
1658 (1862).
Argentina, a pair without exact locality (0. W. Thomas),
in coll. Br. Mus.
Common in S. Brazil, whence came all the types.
Occurs also in Paraguay.
I have examined Walker’s types, as well as a specimen
in M. Dognin’s collection, which has been compared with
Guenee’s. The sexes differ somewhat, both sexes further
vary in size, and Walker’s signataria is a light aberration;
but the species is easily recognised, and the variation is
not such as to justify the list of synonyms or the confusion
that has hung over it. It has been erroneously believed
to be the Tephrosia hyherniaria of Guenee (Spec. Gen. des
Lep., ix. 264), but I am not aware that the error has
found its way into print.
It is quite possible that the following genus of Warren’s
will have to sink to Odysia. So far as I have checked
them, the characters of the two genera agree quite closely,
excepting the curious specialisation of the $ antenna in
Odysia.
229. Stenalcidia (?) ustifumosa (Warren).
Bryoptera (?) ustifumosa, Warren, Nov. Zook, iv, 466
(1897).^
Cymatophora ustifumosa, Warren, Nov. Zook, vii, 196
(1902).2
(2) Parana, Entre Rios, a series in coll. Rothschild ; Metan,
Salta, in coll. Dognin ; Gran Chaco, near Florenzia, 2 $
in coll. Br. Mus. ; La Paz, Mendoza, September 22, 1905
(W. M. Bayne), 1 in coll. L. B. Front.
Geomctridae of the Argentine Rejniblie.
333
Warren’s type (^) was a ^ from Paraguay. I have not
seen the Parana series, and am not absolutely certain that
all the others are correctly referred. Warren (^) says that
the sexes differ considerably, but it would seem that the
^s are also very variable inter se.
I suppose, from Warren’s reference of the ^ to “ Cyma-
tophoi'a ” {Melanolophia, infra) that it has no fovea, but I
am not quite clear about his use of that name. It has
certainly no close affinity with the commotaria group.
The venation, etc., are typical for Stenalcidia, but if fovea is
really wanting, it will form a new section, or allied genus.
230. Stenalcidia dimidiaria (Guenee).
Tephrosia dimidiaria, Guenee, Spec. Gen. des Lep., ix, 261
(1858).
Buenos Aires, common, sitting on trees, posts, etc. (A. F.
Bayne); La Plata City, June 9, 1896 (Oldfield, Thomas),
a worn ^ in coll. Br. Mus.
Apparently distributed in the southern part of Brazil ;
I have also seen it from Paraguay and (a local race ?) from
Ecuador.
M. Doguin, in his MS. list, also records “ Stenalcidia
dimidiaria, Walk.” from Tucuman. Walker’s Boarmia
dimidiaria (List Lep. Ins., xxi, 355), from Bogotd, was
erected quite independently of Guenee’s Tephrosia, but
(sec. specim. typ.) happens to be also a typical Stenalcidia,
so that his name becomes invalid. I believe, however, that
the Tucuman species has proved to be entirely distinct
from the Colombian ; hence the record must be allowed to
lapse until further elucidation is forthcoming.
231. Stenalcidia fusca (Warren).
Stenalcidia fusca, Warren, Nov. Zook, iv, 472 (1897).
Tucuman, 3 $ in coll, Dognin.
Described from Paraguay ; also occurs in S.E. Brazil.
232. Stenalcidia differens (Warren).
Stenalcidia differens, Warren, Nov, Zook, iv, 471 (1897).^
(^) La Plata City, one in coll. Rothschild,
The species was described from Paraguay, whence also
it is represented in coll. Br. Mus, Warren {loc. cit.) says
that the La Plata example is smaller.
334
MrllK Duis B. Front on the
233. Stenalcidia dukinfieldia (Schaus).
Boarmia duhinjicldia, Schaus, J. N. Y. Ent. Soc., v, 104
(1897).
Buenos Aires, common excepting in midwinter (A. F.
Bayne), in coll. L. B. Front et coll. A. F. Bayne ; also
from the same locality (H. Wilkinson) in coll. Br. Mus.
Schaus’ types were from S.E. Brazil. The species has
apparently a rather wide distribution, yet is not at present
known from further north than Bolivia. I have seen
examples from that country and from Faraguay, as well as
from Brazil and Argentina.
Varies considerably in depth of colour and somewhat in
intensity of markings. Mr. Bayne has given me one or
two very pretty, light-coloured forms, and others, again,
of quite a dull fuscous. The ^s are, on the average, more
obscurely marked than the ^s.
234. Stenalcidia invenusta (Dognin).
Stenalcidia invenusta, Bognin, Ann. Soc. Ent. Belg., xlviii,
121 (1904).
Tucuman (type) in coll. Dognin ; La Rioja (Giacomelli),
in coll. L. B. Front.
M. Dognin described this species from only. Dr.
Giacomelli has kindly sent me a ^ which agrees exactly
with these, and also a somewhat smaller, with the lines
of the forewing rather closer together and stronger, the
outer more deeply elbowed, which I have hardly any doubt
is the other sex of the same species.
235. Stenalcidia illineata, Dognin.
Stenalcidia illineata, Dognin, Ann. Soc. Ent. Belg., 1, 210
(1906).
Tucuman (type), in coll. Dognin.
236. Stenalcidia grisea (Warren) (?).
Stenalcidia grisea, Warren, Nov. Zook, vii, 198 (1900).
Physocleora martena, Schaus, MS., in coll. E. D. Jones.
Dr. Bastelberger records this species from Tucuman. I
saw his specimen some years ago, and noted it as a small
Hymenouiima which I could not definitely determine, pos¬
sibly a small specimen of ne2Jhalia, Druce (see infra), but
Gcomctridac of the Argentine Bc^ublie. 335
with the postmedian line of the hindwing placed further
distad and without a dark border beneath. The specimen
has since been determined for Dr. Bastelberger (I do not
know by whom) as Stenaleidia go'isea, but it scarcely seems
to me possible that this can be correct.
Typical S. grisea is from S.E. Brazil, and it is very
likely that the species will be discovered in the Argentine
Republic.
237. Stenalcidia celosoides (Dognin).
Boarmia celosoides, Dognin, Ann. Soc. Ent. Belg., xxxix,
111 (1895).
Tucuman, in coll. Bastelberger.
Dognin’s type was from Ecuador.
238. Stenalcidia junctilinea (Warren).
Aids jttnetilinea, Warren, Nov. Zool., viii, 477 (1901).
Tucuman (type), in coll. Rothschild.
Occurs also in Paraguay (coll. Br. Mus.).
Excepting their generally somewhat larger size, I can
find absolutely no difference between the group to which
this species belongs and typical Stenaleidia. Unless the
whole of that genus were suppressed as structurally in¬
separable from the old-world Aids, it would seem impossible
to xeiei junctilinea to the last-named. Provisionally I can
point to nothing more definite by way of distinction than
the constant coincidence of SC^'^ in Stenalcidia, which I
have not yet found in Aids.
239. Stenalcidia despecta (Prout), nov. sp.
(Plate XLVIII, fig. 7.)
d $ . 24-30 mm. Face fuscous ; palpus fuscous, paler beneath ;
d antennal pectinations rather long. Thorax concolorous with fore¬
wing, tarsi pale-marked at the ends of the joints. Wings somewhat
narrower than in most of the genus, the crenulations of termen of
hindwing pronounced. Forewing pale greyish ochreous, usually
dusted with fuscous, very variably in intensity, the least dusted
part being placed distally to the postmedian ; lines very ill defined,
usually obsolete except at inner margin ; antemedian line (when
discernible) from costa at beyond two-fifths, outcurved in cell,
thence oblique basewards to inner margin at from one-third to two-
fifths (variable) ; postmedian from costa at beyond two-thirds.
336
j\M^jOuis B. Prout on the
somewhat projecting about lb'* and M’, then oblique basewards to
inner margin at scarcely beyond one-half (thus near antemedian),
sometimes accompanied distally by a weak and ill-defined brownish
or greyish shade, which occasionally terminates on inner margin as
a distinct spot ; in a few well-marked examples some dark vein-
dashes on this shade, somewhat as in S. quisquiliaria, Quen. ; cell-
spot minute, indistinct, sometimes obsolete ; a pale subterminal
indicated in a few of the darkest examples ; terminal dark line fine,
thickened into blackish dots between the veins. Hindwing usually
slightly paler, especially in costal region ; terminal area usually con-
colorous with that of forewing ; inner margin usually dark-shaded,
especially in its middle, where there are nearly always the beginnings
(often quite distinct) of two or three dark lines, which disajjpear at or
before the cell ; cell-spot sometimes rather large and distinct, some¬
times small and inconspicuous, occasionally obsolete ; pale sub¬
terminal indicated in the darkest examples ; terminal line with the
interneural dots weaker than in forewdng. Undersurface nearly
unicolorous fuscous greyish, hindwing somewhat paler and rougher-
scaled than forewing, cell-spot usually present on both wings or at
least on hindwing, sometimes quite conspicuous on the latter.
Mendoza, La Paz, Palmira, etc. — mostly bred (W. M.
Bayne), a large number in coll. L. B. Prout, coll. A. F.
Bayne, etc. ; type $ (excl. SeiAember 8, 1905) in coll.
L. B. Prout.
Mr. W. M. Bayne lias been very successful in breeding
this species ex ovo on chrysanthemum ; the broods emerged
May 1904, and August (end) to September 1905; 1 have
also wild specimens from La Paz, March 1903, and
Palmira, December 1903, and suppose there is a succes¬
sion of broods. Extremely variable, though always an
obscure-looking little insect. One or two of the lightest
specimens are virtually free from dusting, and marking¬
less; sometimes three approximated dark spots on inner
margin (ends of antemedian, postmedian, and shade
distally to the latter) are the only distinct markings.
A typical Stenalcidia in structure, but with ^ hindtibia
not dilated. Perhaps nearest to S. fusea, Warr.
240. Physocleora obscura (Schaus).
Physoeleora obscura, Schaus, J. N. Y. Ent. Soc., vi, 148
(1898).
Buenos Aires, one $, December 9, 1902 (A. F. Bayne),
in coll. L. B. Prout.
Gcometridae of the Argentine Republic. 837
Described from S.E. Brazil. I have seen co-types in
the collection of Mr. E. Dukinfield Jones, and a specimen
presented by Mr. Schaus in that of the Bi'itish Museum,
and believe the identification is correct ; but I cannot say
that I am absolutely certain. Mr. Warren inclined to
refer my specimen to enana, Dognin.
241. Physocleora punctilla (Schaus).
Physocleora punctilla, Schaus, J. N. Y. Ent. Soc., vi, 148
(1898).
Tucuman, in coll. Dognin.
Schaus’ type was from S.E. Brazil ; the species occurs also
in Paraguay.
242. Physocleora enana (Dognin).
Boarniia enana, Dognin, Ann. Soc. Ent. Belg., xxxix, 112
(1895).
Los Vasquez and Tucuman, in coll. Dognin. Dognin’s
type was from Ecuador.
243. Physocleora nigrescens (Prout), nov. sp.
9 . 20 mm. Light brown, densely speckled with blackish, only
the median area of forewing remaining light (somewhat speckled
with darker brown), a very broad distal area of forewing and almost
the whole of hindwing particularly blackened. Forewing with base
of costa blackish ; a suhbasal, irregularly curved dark line at beyond
one-fifth ; an antemedian, incurved below cell, at two-fifths ; a post¬
median, strongly inbent below cell, at nearly three-fifths, followed
by a dark ferruginous shade, to which again follows the blackish
marginal area ; cell-spot black, distinct ; subterminal line in¬
distinctly pale, strongly dentate, filled in proximally with black ;
terminal black spots between the veins. Hindwing with black
cell-spot and terminal spots distinct, black antemedian and post¬
median lines (slightly inhent below cell), and pale subterminal dis¬
cernible, a ferruginous tinge distally to po.stmedian. Undersurface
similar, basal area of forewing more obscure, no ferruginous shading
di.stally to postmedian.
Gran Chaco, near Florenzia, October 1902 (S. R.
Wagner). Type and another quite similar ^ in coll.
Br. Mus. A worn example from Tucuman, in coll.
Bastelberger, seems to be the same species. Possibly
TRANS. ENT. ,SOC. LOND. 1910. — PART III. TnOV.) Z
338
I mis B. Front on the
an extremely darkened form of S. 2'>U7ictiUff, Schaus, of
which I can only compare very poor specimens.
Venation normal for the group, SC^ ‘‘^ of forewing co¬
incident, from very slightly above middle of disco-
cellulars. In the type specimen, the stalk of is
almost connate with SC^‘^. It is very doubtful, as I
understand Mr. Warren now recognises, whether Fhyso-
cleora is structurally differentiable from Stenaleiclia\ it
seems to be merely a “ genus of convenience,” for the
accommodation of the smaller species, which have rather
a different facies from the more typical Stenalcidia.
244. Hymenomima extersaria (Warren).
Ilymenominut extersaria, Warren, Nov. Zook, iv, 4G7
(1897).!
Hyrienoriima densata, Warren, Nov. Zook, xiv, 27G (1907)
(ab.) (nov. syn.).-
Tucuman, in coll. Dognin.
Described from Colombia Q) and Peru (2). I have seen
a good many specimens from Peru, but do not know of
any more southerly records beyond that t)f M. Dognin.
The species varies in depth of colour and strength of the
pale distal blotch of forewing, which is almost absent in
the ab. densata-, the undersurface seems very constant.
245. Hymenomima nephalia (Druce)(?).
Boai'mia oicijhalia, Druce, Biol. Centr.-Amer., Lep. Het., ii,
75, tab. xlviii, 10 (1892).
Tucuman, in coll. Dognin ; also (?) in coll. Bastelberger.
The true nephalia is from Mexico, and I have seen only
Druce’s type specimen. There stand with it, in the British
Museum, two other examples from Paraguay, which may
possibly represent a local race, with rather longer hind¬
wing, its postmedian line further from the cell-spot and
some other differences, but which I should personally have
been disposed to regard as a separate, though nearly allied
species. I believe it is to this latter that M. Dognin’s
“ nejphalia ” should be referred ; and I have already (see
Stenalcidia grisea, Warr.) recorded that Dr. Bastelberger
has a specimen which might be an extreme aberration of
the same. Here, as in many other cases, more material is
the great desideratum.
Geometridac of the Argentine Republic. 331)
246. Hymenomima amberia (Schaus).
Boarmia amberia, Schaus, Tr. Anier. Ent. Soc,. xxvii, 183
(Feb. 1901V
Hymenomima subsordida, Warren, Nov. Zool., viii, 478
(Dec. 1901V
(“) Parana, Entre Rios (type), in coll. Rothschild ; Tucu-
man, in coll. Dognin ; ? Gran Chaco, near Florenzia, in
coll. Br. Mas. ; Pirapb (Bastelberger, J. B. Nass. Ver.
Nat,, Ixi, 75).
Schaus’ type (^) was from S.E. Brazil ; I have also seen
the typical form from Paraguay.
The British Museum example from Gran Chaco is con¬
siderably smaller and of a somewhat different aspect ; it
may represent a local race or allied species, though possibly
it is a mere aberration.
I have accepted the synonymy as communicated to me
by M. Dognin, and this may possibly prove to be correct,
although on going into the question together last year, we
decided that it was by no means a certainty.
247. Hymenomima perfuscimargo (Prout), nov. sp.
d. 29 mm. Of the size and somewhat the aspect of B. exangulata,
Warn, Tr. Amer. Ent. Soc., xsx, 508, from British Guiana ; fovea
extremely develoj^ed, much as in that species. Wings bone-colour
tinged with ochreous, speckled with fuscous, and with almost the
entire distal area broadly fuscous, though with the apex of forewing
paler, the subterminal pale line faintly traceable and the space
between this and the termen irregularly mixed with the lighter
groundcolour, especially on the hindwing ; proximal boundary of
this area (= postmedian line) on forewing from two-thirds of costa
to three-fifths of inner margin, incurved below cell, on hindwing
from near three-fourths of costa, outangled at SC^, somewhat out-
curved from Bb then inbent below cell, finally reaching inner
margin at beyond two-thirds, by a bend distad, as in H. exangulata.
Forewing in addition with basal area more or less fuscous, a fuscous
discal mark, preceded by a dark mark on costa which represents the
beginning of a faintly traceable median shade, angled below SC,
crossing discal spot, running obliquely towards inner margin and
continued as an antemedian line on hind wing. Hindwing in
addition with a rather large comma-shaped cell-spot, placed rather
near postmedian line, the cell being longer than in most of the allies.
Undersurface much more weakly marked, the terminal dark baml
Z 2
340
Mi'^iduis B, Proub vn the
of forewing ill-defined but complete, that of hind wing still weaker,
and only present in costal half of wing, vanishing beyond R3.
Head, body and legs of groundcolour, face and palpus rather more
strongly fuscous-speckled, foreleg dark-marked anteriorly.
Gran Chaco, near Florenzia, October 1902 (S. R.
Wagner). Type in coll. Br. Mus.
248. Iridopsis fulvitincta (Warren).
Chora fulvitincta, Warren, Nov. Zool., iv, 460 (1897).^
1 IridojJsis commixtata, Dognin, Ann. Soc. Ent. Belg., xlviii,
121 (1904) (nov. syn.).^
Irido'psis ohliquata, Dognin, Ann. Soc. Ent. Belg., xlviii,
364 (1904) (nov. syn.).^
Iridopsis pallescens, Warren, Nov. Zool., xiv, 277 (1907)
(nov. syn.).^
(^) La Plata City (type), in coll. Rothschild ; )2) (^)
Tucuman (types), in coll. Dognin, also from same locality
in coll. Bastelberger ; Buenos Aires (A. F. Bayne) in coll.
L. B. Prout, also from same locality in coll. Dognin ;
Metan and Los Vasquez, in coll. Dognin (recorded as
ohliquata) ; La Rioja (Giacomelli), in coll. L. B. Prout ;
Argentina (O. W. Thomas), without more precise locality,
in coll. Br. Mus.
Evidently a very widely distributed species in the
Republic. Elsewhere, the only specimens which I can
with any confidence refer here are Warren’s type of
pallescens {*), from Paraguay, and three examples from
Callao in coll. Br. Mus. The species is. an exceedingly
variable one, and will give some trouble, being very closely
allied to several others (such as syrniaria, Guen., delin-
quaria., Walk., etc.), which are nevertheless evidently
distinct. I think the synonymy proposed above, with the
possible exception of commixtata, Dogn., will prove correct ;
but unfortunately Warren’s type of fulvitincta is in
exceedingly bad condition, and it will be impossible to settle
its identification positively until a long and representative
series from La Plata is available for study.
249. Iridopsis rectura (Dognin).
Iridoqjsis rectura, Dognin, Ann. Soc. Ent. Belg., xlviii, 363
(1904).
Tucuman (type), in coll. Dognin; I also saw in coll.
Bastelberger, from the same locality, an example of an
Gcometridac of the Argentine Republic. 341
Iridopsis, which I noted as perhaps a form of the same
species.
250. Melanolophia apicalis (Warren).
Cymatophora apicalis, Warren, Nov. Zool., vii, 195 (1900).
Buenos Aires (H. Wilkinson), 2 1 ^ in coll. Br. Mus.
Warren’s type was from S.E. Brazil; I know of uo
further distribution.
The Buenos Aires $ is larger and of a warmer, darker
brown than the ^s.
The name of Cymatophora (Hiibner, Tent., indescr. ;
Packard, nom. praeocc. — nec. Treitschke) is clearly unten¬
able for this group, which indues argilaria, Guen., coin-
motaria, Mssn., and many others. On the other hand, it
is certainly referable to Melanolophia, Hulst, Tr. Amer.
Ent. Soc., xxiii, 357 ; the agreement of the South American
representatives with canadaria, Guen., the type of that
genus, is very close structurally, and even superficially.
251. Melanolophia (?) granillosa (Dognin).
Cymatophora (?) granillosa, Dognin, Ann. Soc. Ent. Belg.,
xlvi, 349 (1902).
Tucuman, in coll. Dognin.
Dognin’s type description does not give any definite clue
as to the generic affinities.
252. Melanolophia (?) eobusta (Warren).
Cymatophora robusta, Warren, Nov. Zool., viii, 477 (1901).
Parana, Entre Rios, type in coll. Rothschild.
From the figure in the British Museum Collection (I
cannot now compare the type), I do not think this can
really be a Melanolophia. The shape, and the style of
pattern, much rather suggest that it is related to usti-
fnmosa, Warren, which I have provisionally placed with
Stenalcidia.
253. Melanolophia (?) gnamptographa (Prout),
nov. sp.
7 . 38 aim. Head, body and legs brown, more or less tinged
with reddish. Forewing reddish brown, irrorated with dark fuscous ;
lines brighter brown, with a slight olivaceous tinge ; anteraedian ill-
defined, from before one-fourth costa to before one-third inner
842
Mr. Louis B. Front on the
margin, outbeiit in cell, in submedian area and close to inner
margin; postmedian distinct, concise, from about two-thirds costa
to nearly two-thirds inner margin, forming a sinus in cell and a
second in submedian area, and accompanied distally by some ill-
defined grey shading, which is densest in costal half of wing, and
somewhat obscures the postmedian itself on costa ; an ill-defined,
greyer (less ferruginous-brown) median shade at about two-fifths
(well before discal spot), slightly outenrved in middle of wing ;
subterminal white, much interruj)ted, especially in posterior half of
wing ; distinct between SC^ and SC^ and between SC^ and R* as
elongate white spots and from Ri to as an outcurved line, or
lunule, accompanied distally (and in anterior half of wing proxi-
mally) by some blackish-fuscous marks ; cell-spot small, blackish ;
termen with distinct blackish dots between the veins ; fringe clouded
with fuscous, with a narrow ■i)ale line at its base. Hindwing paler,
especially in basal part, nearl}' unicolorous, cell-dot indistinct,
terminal dots almost obsolete, fringe with narrow pale line at base.
Underside almost unmarked, cell -dots weak, the faintest possible
suggestion of a paler, proximally darker-shaded postmedian on hind¬
wing. Shape rather near M. (?) limosa (Dogn.),* but slightly nar¬
rower, termen of forewdng slightly more elbowed at IF, hindwing
slightly more produced apically.
Tucuman, 450 in., type in colL Br. Mus.
254. Eotropis semicana (Bastelberger).
Ectroins semicana, Bastelberger, Soc. Ent., xxii, 138 (1907).
Tucuman (type), in coll. Bastelberger.
255. Bronchelia pudicaria (Guenee).
Bronchelia ijudicaria, Guenee, Spec. Gen. des Lep, ix, 287
(1858).!
Bronchelia detexta, Walker, List Lep. Ins., xxi, 452
(1860).2
Tucuman, in coll. Dognin et coll. Bastelberger.
A widely distributed species, described by Guenee (^)
from Brazil, by Walker (‘^) from the West Indies. On
Dr. Bastelberger’s Tucuman examples see J. B. Nass. Ver.
Nat., lx, 90. I saw a ^ from his collection which I was
inclined to refer to 'puellaria, but I may have been mis¬
taken ; I imagine it is the ''pudicaria of his note.
* Tephrosia limosa, Dogn., Ann. Soc. Ent. Belg., xxxix, 116.
Gcomctridac of the Argentine Bcimhlic.
343
256. Bronchelia puellaria (Guenee).
Bronehelia puellaria, Guenee, Spec. Gen. des Lep., ix, 287
(1858).
Bronchelia mundaria, Walker, List Lep. Ins., xxi, 452
(1860).
Bronchelia nigriplena, Warren, Nov. Zook, xiv, 272 (1907)
(nov. syn.).
Bronchelia puellaria, ab. confertistriga, Bastelberger, J. B.
Nas. Ver. Nat., lx, 89 (1907) (ab. ?).
? Tucuman (see under B. pudicaria, supra).
Widely distributed from Mexico (?) to Brazil.
257. “Bronchelia transitaria (Walker).”
?? Bronchelia transitaria. Walker, List Lep. Ins., xxi, 453
(1860).
Tucuman, in coll. Dognin.
This name is included in M. Dognin’s MS. list, but I
suspect some error of determination. Walker’s transitaria
was from Haiti, and is (sec. specim. typ.) a synonym of the
common Bronchelia scolopaiea (Drury) of Central America
and the West Indies, of which I have never seen specimens
from far south.
258. Bronchelia curvilinea (Warren).
Bronchelia curvilinea, Warren, Nov. ZooL, xiv, 271 (1907).
Ciudad de Tucuman (type), in coll. Rothschild.
259. Thyrinteina arnobia (Stoll).
Phalaena Geometra arnobia, Stoll, Cramer’s Pap. Exot., iv,
188, tab. ccclxxxiii, I (1782).
Amphidasys arnobia, Guenee, Spec. Gen. des Lep., ix, 211,
(1858).
Boarmia oppositaria. Walker, List Lep. Ins., xxi, 361,
(1860).
Boarmia quadricostaria, Herrich-Schaeffer, C. B. zool.-min.
Ver. Regensb., xxiv, 52 (1870).
Biston immissus, Felder, Reise Novara, Lep. Het., tab.
cxxxiii, 24 (1875) (ab.).
Thyrinteina quadricostaria, Moschler, Abb. Senck. Ges.,
xvi, 269, fig. 12 (1890).
Tucuman, in coll. Dognin.
344
M^Louis B. Prout on the
Common and widely distributed from Central America
and the West Indies to Brazil. Decidedly variable, I am
not sure whether the synonyms here given have ever
before been published, though they have long been known
to w'orkers at the South American Geometridae.
260. Devarodes interlineata (Berg, MS.) (Prout),
nov. sp.
Cymopsis (?) interlineata, Berg, MS., in coll. Br. Mus.
Devarodes interlineata, Warren, Nov. Zook, xvi, 100
(indescr.)
28 mm. Body and wings dull black, slightly tinged with
brownish. Basal joint of j^alpus, collar, breast and tail orange-
yellow. Forewing above with a long whitish streak from base to
near termen, margined anteriorly by cell and posteriorly reach¬
ing well beyond submedian fold, which appears as a dark line
bisecting it ; an elongate whitish patch from DC to near termen,
margined by and R^, bisected by R^ as a dark line. Hindwing
above unicoloroi;s. Forewing beneath with the streak obsolescent,
but remaining distinct as a larger or smaller spot at its distal end ;
subajiical blotch broadened, so as to reach to SC^, but rounding off
distally. Hindwing beneath with elongate whitish marks distally
to cell and between this and costa.
Missiones (C. Berg), in coll. Br. Mus., the type of the
species.
A second almost identical, though with slightly more
white beneath, from Sapucay, Paraguay, stands in coll. Br.
Mus. ; Warren mentions this, or another, from the last-
named locality.
I do not know what are the actual affinities of this
species, which Warren — overlooking the fact that Berg’s
name was unpublished — does not describe in the note in
which he advocates removing it to Devarodes. Berg pre¬
sented it to the British Museum with the MS. name of
Cymopsis (1) interlineata attached. Warren communicates
nothing further than that “ the hindwing has no radial ”
{i. e. no R^). The following particulars may aid in the
recognition of the species. Face somewhat prominent ;
eye small; palpus moderate, porrect, rather slender, 2nd
joint shortly scaled; ^ antenna bipectinate, with rather
long, strong branches ; hindleg {$) rather short, tibia
dilated, with terminal spurs only ; retinaculum long and
strong, C of forewing well removed from subcostals, SC^‘^
Trnn.s. Ent. Soc. Lond., 19J0. Pl.XLVIIf.
ARGENTINE GEOMETRIDAE
Explanation of Plate XLVIIl.
Fig. 1. Pero (Azelina) norma, Front, uov. sj)., ^
2. Tephrinopsis suhscripta, Front, nov. sp., ^
3. Macaria (?) repetita, Front, nov. sp., ^
4. Ochi/ria argentma, Front, nov. sp., ^
5. Eneitpiithecia cisplatensis, Front, nov. sp., ^
6. Lithostege (?) tzaddi. Front, nov. sp., 9
7. Ste7ialcidia despecta, Front, nov. sp., ^
8. Pnphyia zona (Dognin) (corrividata, Warr. var. ?),
9. Gnophos (?) omts. Front, nov. sp., 9
10. Pseudobrephos haynei, Front, nov. sp., ^
11. Hamaha {T) platensis, Front, nov. sp., ^
12. Papifhecia aiyimorrhoa, Front, nov. sp., $
13. Psodopsis tortilinea, Front, nov. sp., ^
14. Pnphyia limbata, Front, nov. sp., ^
15. Narragodes gyda, Front, nov. sp., ^
16. Eupithec.ia argica, Front, nov. sp., ^
17. tz>alpis (Microdontopera) eudoi'a, Front, nov. sp., ^
18. Acidalia ecUpes, Front, nov. sp., ^
19. Hypolepis prionogramma, Front, nov. sp., ^
20. Idialcis metria. Front, nov. sp., 9
21. Hanadia flexifascia, Front, nov. sj)., $
22. llerbita asinana, Front, nov. sp., ^
23. Salpis (Antygophayies) orbifera, Front, nov. sp., ^
24. Catascia carnea, Front, nov. sp., ^
25. Baynia odontota, Front, nov. sp., ^
26. Tephrinopsis mesoscia, Front, nov. sp., ^
27. Arizela dulcis, Front, nov. sp., 9
28. Lithostege giacomellii, Front, nov. sp., 9
29. HamaMa (?) ptychopoda, Front, nov. sp., 9
30. Lithostege pax, Front, nov. sp.,
31. Salpis rubens. Front, nov. sp., ^
32. Pero {Azelina) aynnicincta. Front, nov. sp., ^
33. Chrismopteryx vicina. Front, nov. sp., 9
34. Chrmnopteryx expolita, Front, nov. sp.,
35. Thamnonotna tripartita. Front, nov. sp., ^
36. Mimophyle sabulosa. Front, nov. sp.,
37. Rhopalodes otophora. Front, nov. sp., ^
38. Neochesias opistholeuca. Front, nov. sp.,
39. Pero {Azelina) palmirensis. Front, nov. sp., ^
40. Orthonama superjecta. Front, nov. sp., 9
Geometridac of the Argentine Republic. 345
long-stalked, SC^ running into SC^'^ (or, SC^ absent, SC^
connected by SC^ '^ by a bar), discocell ular, nearly vertical ;
hindwing with cell rather long, C free, nearly parallel to
SC (not very close) to fully one-half cell, thence gradually
diverging.
261. Melanchroia chephise (Stoll).
\Plialaena Geometra] chephise, Stoll, Cramer’s Pap. Exot.,
iv, 182, tab. ccclxxxi, E (1782) {Phalaena Geometra
cephise, ibid.. Index, 248).
Phalaena eorraria, Fabricius, Mant. Ins., ii, 881 (1787).
Melanchroia cephise, Hiibner, Verz. bek. Schmett., 173
(1826 ?).
Callimorpha cephise, Verloren, Cat. Lep. Crameri, 142
(1837).
Tanagraif) expositata,WQ\keY, List Lep. Ins., xxv, 1461
(1862).
Tucuman (Burmeister).
Widely distributed from the southern United States,
Mexico and the West Indies to Paraguay and southern
Brazil, but I have at present no other records for Argentina
than that contained in Burmeister’s Descr. Phys. Rep.
Argent.
Explanation of Plate XLVIII.
[See Explanation facing the Plate.]
( S4(j )
IX. Third Paper on the Tetriginae (Orthoptera) in the
Oxford University Museum. By J. L. Hancock,
M.D., F.E.S.
[Read March 2nJ, 1910.]
Plate XLIX.
The present studies are mainly founded on a series of
forty-seven specimens of Tetriginae (Orthoptera) recently
acquired by the Oxford University Museum. I am indebted
to Professor E. B. Poulton and Mr. R. Shelford for the
privilege of examining them. They have enabled me to add
supplemental data to my two former papers published in
the Transactions of the Entomological Society of London.
These studies are made still more complete by the
inclusion of notes founded on material in my private
collection.
The section Cladonotae is represented by ten known
genera in Africa ; the following diagnostic table based on
their distinctive characters may prove useful in separating
them. Bolivar* recognised in his “Essai” (1887) species
belonging to three genera, namely : Xcrophyllum, Trachy-
tettix, and Pantelia. Later Karschf (1890) added species
of Acmo-phyllum, Trypophylhim, Hippodes, Hypsaeus, and
still later Piezotettix,^ making eight genera recognised up
to that time. I have recently added another genus, Clado-
ramus,^ and in the present paper supplemented a new one,
Medinophylluin, bringing the whole number up to ten now
known and differentiated in the table herewith presented.
In each genus the type is given. The genus Hypsaeus
and Piezotettix also occur in the Oriental region, while the
other eight genera are confined to Africa.
* Ann. Soc. Entorn. Belg. xxxi, 1887.
I Entoniol. Nachricht. 1890.
J Berl. Entom. Zeit., xxvi, p. 196.
§ Trans. Entom. Soc. London, p. 217, 1907.
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.)
Dr. J. L. Hancock’s Third Paiicr on the Tetriginac. 347
SYNOPTICAL TABLE OF AFRICAN GENERA
OF THE SECTION CLADONOTAE.
1. Anterior and middle femora slender,
elongate.
2. Pronotum compressed, foliaceous.
3. Dorsum of pronotum strongly com¬
pressed, highly foliaceous, angu-
late, pronotum in profile triangu¬
late ; first articles of posterior
tarsi nearly twice the length of
the third. Type westivoodi, Boh Hypsaeus, Boh*
3.3. Dorsum of pronotum subfolia-
ceous, more or less compressed,
tectiform- cristate, in profile
above straight or arcuate ; first
articles of posterior tarsi nearly
three times the length of the
third. Type cultratus, Stal . Piezotettix, Bol.f
2.2. Pronotum not compresso-foliaceous,
low obtuse tectiform, apex of pos¬
terior process abbreviated, widely
rounded ; elytra and wings
wanting ; first articles of pos¬
terior tarsi much longer than the
third. Type vicarius, Karsch . Hippodes, Karsch.
1.1. Anterior and middle femora with
both margins more or less foliaceo-
dilate, carinae either entire, un¬
dulate, lobate, serrate or lobato-
dentate.
4. Facial frontal scutellum more or less
compressed, and either crenulate
or spinulose, but not glabrous.
5. Crest of pronotum highly foliaceous
above, more or less semicircular
and regularly arcuate, the border
smoothly longitudinally silicate
at the anterior two-thirds. Type
servillei, Fairm . Xerophyllum, Fairm.
* The African species are aetpialis, Karsch, inaeipudis, Karsch, and
(ingulatus, sp. nov.
t The African species are clypeatus, Karsch, and karschi, Bolivar.
848
Dr. J. L. T^ncock’s Tldrd Paper on the
5.5. Crest of pronotum low, com-
presso-cristate, siibfoliaceous,
scabrous-spinose, 2)rocluced for¬
ward, border above somewhat
distinctly sulcate longitudi¬
nally and crenulate or erose
sjiinulose posteriorly. Type
crenulatus, Hanc . Cladoramub, Haiic.
6. Pronotum dejiressed.
7. Humeral angles outwardly pro¬
duced, amjiliate, laminate and
serrate ; eyes substylate ; vertex
very wide, and armed with dis¬
tinct acute sj^ine on each side.
Type hnfo, Costa . Trachytettix, Stal.
7.7. Humeral angles wanting, dorsum
of j^ronotum acute tectiform,
anteriorly acute ; vertex
narrow ; body apterous. Type
cristidata, Bol . Pantelia, BoL
6.6. Pronotum largely comjDressed and
foliaceous, border above sharp, not
at all longitudinally sulcate.
4.4. Facial frontal costa glabrous.
8. Margins of anterior and middle
femora entire or below sub¬
undulate.
9. Foliaceous crest of 2:)ronotum per¬
forate with foramina, border
subsemicircular but little
obtuse angirlate above about
the middle. Type (jlabrifrons,
Karsch . Trypophylluji, Karsch.
9.9. Foliaceous crest of j^ronotum
arcuate before the middle,
not abruptly cristate, rounded
angulate above at about the
middle and dellexed back¬
ward, aj^ex subobtuse or
lengthily extended backward
beyond the hind femoral
apices, acuminate ; inferior
external carinae of posterior
Tetriginae in the Oxford University Museum. 349
femora entire. Type xero- [nov.*
j)hylloides, Bol . Medinophyllum, gen.
8.8. Margins of anterior and middle
femora below tri- or quadrilo-
bate ; foliaceous crest abruptly
highly elevated cristate forward,
superior margin above undu¬
late or sinuate ; inferior ex¬
ternal carinae of posterior
femora incrassate bearing lobes.
Type undulatum, Karsch. . . Acmophyllum, Karsch.
Genus Hypsaeus, Bol.
Ann. Soc. Entom. Belgique, xxxi, p. 200, 1887.
H. angidatus, sp. nov.
Face in profile subrounded, little sinuate below the frontal scu-
tellura ; scutellum elongate, margins entire, above curvate divergent,
but subparallel below toward the median ocellus ; antennae inserted
between the lower fourth of the eyes. Pronotum triangulate, strongly
compresso-foliaceous, highly elevated and angulate above the middle,
the summit of the angle rounded, anteriorly extended beyond the
head, the inferior margin of process convex, the frontal apex obtuse-
rounded ; margin above at the anterior half of crest arcuate toward
the process in front, but flattened toward the elevated summit ;
behind the middle deflexed backward and toward the hind apex very
slightly undate-sinuate ; anterior and posterior halves of the upper
margin subequal in length, the posterior process scarcely extended
backward to the knees of the hind femora ; lateral lobes of pronotum
posteriorly unisinuate, posterior angles obliquely excised behind.
Elytra and wings wanting. Anterior femora elongate, little com¬
pressed, superior carinae entire, below slightly bilobate ; middle
femora slender, margins entire ; posterior femora above ampliate,
the superior carina strongly quadrilobate, the apical or fourth lobe
triangular and acute; the superior external carina incrassate pos¬
teriorly ; ventral carina substraight at the apical three-fourths, and
here bearing five indistinct small tubercles or lobuli ; first articles
of the posterior tarsi much longer than the third, the first and
second pulvilli nearly equal, the third much longer than the rest
* The genus Medinophyllum also includes the species Acmophyllum
conrudti, Bol.
850 Dr. J. L. I^ncock’s Third Pcqm' on the
in length. Colour fusco-rufescent, the crest of pronotum wholly
opaque, margin above marked with fuscous.
Length of pronotum, ^ 12.5 nun. ; height at the middle, 9 mm. ;
posterior femora length, 7.6 mm.
Locality : Bitje Ja River, S. Cameroons (Rosenberg).
One example in the author’s collection. This species
resembles H. acquedis, Karsch, but is distinguished by the
smaller stature and by the more distinct right angle
summit above.
Genus Xerophyllum, Fairm.
Ann. Soc. Entom. France, p. 24G, 1846.
The following table is a revision of the species of the
African genus Xeroqdiylluin, and includes three new
species.
TABLE OF XEROPHYLLUM SPECIES.
1. AVings entirely explicate and extended
more or less beyond the pronotal
apex ; foliaceous crest curvate, an¬
teriorly substraight deflexed back-
Avard, i^osterior process lengthily
extended backward beyond the
apices of hind femora; length of
pronotum, (J 14 mm. ; 5 15 mm. . extensum, s'p. nov.
1.1. Wings not entii’ely explicate, rarely
reaching to and not beyond the
pronotal apex ; foliaceous crest
semicircular or subreniform.
2. Anterior process of pronotum obtuse
angulate, the crest regularly arcu¬
ate ; anterior femora above nearly
entire or somewhat sinuate, below
undulate ; lateral carinae of frontal
scutellum compressed crenulate ;
wings extended nearly to the prcj-
notal apex ; length 19 mm. . . . servillei, Fairm.
2.2. Anterior process of pronotum sub¬
acute, crest largely elevated
behind the middle ; anterior
femora above serrate, below
Tetriginac in the Oxford University Museum. 851
lobato-dentate ; wings cov^ered
by the sides of pronotum ; length
of pronotum, cj 12 mm. . . .
3. Pronotum behind at the apex
incised, crest highly arched-
cycloidal ; inferior margin of pos¬
terior femora strongly denticulate,
inferior external carina bearing
denticles : length of pronotum,
12 mm .
3.3. Pronotal apex obtuse-rounded,
margin barely undate or sub-
sinuate ; crest subreniform ;
frontal scutellura cristate pro¬
duced, entire or very slightly
serrulate : vertex carina bearing
a median projecting denticle ;
posterior femoral carinae ven-
trally strongly quinquelobate,
inferior external carina trilobate ;
length of pronotum, ^ 10.5 mm.;
5 11.5 mm .
4. Dorsal crest very highly compresso-
elevated, semicircular, margin
behind near the apex crenulate ;
length of pronotum, 13 mm. .
4.4. Dorsal crest very highly com-
presso-elevated, semicircular,
but very little subangulate-
rounded above tlie middle ;
lateral carinae of frontal scu-
tellum minutely spinulose ;
length of pronotum, ^ 11.5
mm. ; $ 11.8 ram .
simile, Bob
platycorys, Westw.
cristifrons, sp. nov.
galeatnm, Karsch,
fusewn, Bol. (larva) 1
neavei, sp. nov.
X. extensum, sp. nov. (Plate XLIX, fig. 1.)
(See preceding synoptical table.) Lateral carinae of frontal
scutellum minutely spinulose or denticulate. Pronotum highly
elevated and strongly compresso-foliaceous, crest very thinly trans¬
lucent bearing radiating veins; superior margin forward of the
middle of dorsum regularly smoothly curvate, highly and roundly
elevated subangulate behind the middle and extended deflexed back¬
ward substraight to the acute apex ; behind more than a third of
852
Dr. J. L. cock’s Third Paper on the
the posterior margin above minutely crenulate barely undate ; the
posterior process lengthily extended backward beyond the hind
femora, a distance nearly equal to their length, the lateral margins
below convex, anterior process in front curvate, apex bidentate, sub-
obtuse, the inferior margin strongly ol)lique, substraight ; lateral
lobes posteriorly bisinuate. Elytra oblong ; wings fully explicate,
freely exposed at the sides and ends, and more or less distinctly ex¬
tended beyond the pronotal apex, and beyond the hind femora a
distance equal to their length. Femora subscabrous granulate, an¬
terior femora foliaceous-dilate, superior carina serrulate, terminating
in a small denticle, ventral margin .serrulate-sublobulate ; middle
femora above serrulate-sublobate, inferior carina ampliate backwards
toward the apex, serrate and incised bilobate ; posterior femora
externally scabrous, superior carina compresso-ampliate and serrate
backwards terminating acutely, superior surface before the knees
bearing two distinct fuscous spines, the inner one erect ; external
femoral carina thin, inconspicuous and minutely serrulate, the in¬
ferior external armed with two obscure minute denticles ; ventral
carina crenulate and somewhat distinctly quinquelobate. Colour
greyish or greyish rufescent, the crest sometimes cinerous, subhya¬
line, and rarely marked with fuscous along the margin.
Length of pronotum, 14 mm. ; posterior femora, 5.3 mm.
9 pronotum, 15.5 mm. ; posterior femora, 6 mm.
Seven examples; three males and four females from
N.E. Rhodesia in the Oxford University Museum, col¬
lected by S. A. Neave in the following localities: — Two
from Lofu River, 3500 ft., No. 2129, August 13, 1908 ;
one from Chisinga Plateau, Kalangwini district, 4500 ft.,
September 25, 1908 ; three from Upper Kalangwisi
Valley, 4200 ft.. No. 2125, September 3, 1908, and one
from Alala Plateau, Ndola district, 4000 ft., October 12,
1905.
This species may be a dimorphic long-wing form of
X. neavei.
X. crisiifrons, sp. nov.
Frontal scutellum cristate produced, the margins between the an¬
tennae subentire or barely minutely serrulate, above slightly denticu¬
late, median carina of vertex slightly crenulate, and bearing a dis¬
tinct denticle projecting between the middle of the eyes. Pronotum
strongly and thinly compressed foliaceous, bearing radiating veins ;
superior margin subreniforin, forward before the middle flattened
Tetriginae in the Oxford Unirersity Museum. 353
arcuate, in front strongly roundly produced, behind the middle more
largely elevated curvate, backward the margin at the apical fourth in¬
distinctly quadri- or quinque-undate or subsinuate at the obtuse apex ;
superior margin rather widely longitudinally sulcate ; anterior pro¬
cess subobtuse, the apex strongly bidentate, inferior lateral margins
viewed from above strongly serrate, viewed from the side strongly
oblique, substraight or little curvate produced over the head ; pro-
notum posteriorly extended distinctly beyond the femoral ajjices ;
lateral lobes posteriorly bisinuate, the upper sinus very shallow,
posterior angles strongly oblique, subobtuse and obliquely excised
behind. Elytra oblong ; wings not visible. Anterior femora com-
pressed-foliaceous, superior carina serrulate-undate, below subquadri-
lobate, middle femoral carinae above comjiressed, serrulate-undulate,
below strongly quadrilobate, or the two middle lobes fused in one,
then trilobate ; posterior femora rugose, superior cai'ina crenulate
(in the female bearing four tubercles), and the apex triangularly
elevated; superior area before the knee bispinose, external pagina sub-
tuberculate, as viewed from above the inferior external carina strongly
trilobate, the first very small, ventral carina strongly quinque-lobate
and finely serrate between the lobes. Colour rufescent, infuscate on
the pronotum, the rest of body lighter.
Length of pronotum, $ 11'5 mm. ; posterior femora, 5 mm.
pronotum, 10'5 mm. ; posterior femora, 4'3 mm.
Two examples from Abuthsi, River Niger, Africa
(Rosenberg), in the author’s collection.
X. neavei, sp. nov. (Plate XLIX, fig. 2.)
(See preceding table.) Lateral carinae of frontal scutellum
minutely spinulose or denticulate. Pronotum subscabrous-granu-
late, highly elevated, very thinly compresso-foliaceous, sides
bearing radiating veins, conspicuous when held against the light,
superior margin of crest nearly semicircular, but very little sub-
angulate-rounded above the middle, the anterior half regularly
smoothly curvate, the edge above somewhat widely longitudinally
sulcate, little planate, posterior half curvate, but the margin at the
apical third part minutely crenulate, and here toward the apex
slightly undate and little sinuate above the apex (more distinct in
the female), anterior process subobtuse, the front border arcuate
and the apex bidentate, and just above in front little denticulate ;
inferior lateral borders of anterior process strongly oblique, produced
beyond the head, margin substraight, but viewed from above later¬
ally serrate ; posterior j^rocess extended backward little beyond the
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) A A
354
Dr. J. L. ^.ncock’s Third Paper on the
apices of the hind femora ; lateral lobes posteriorly bisinuate.
Elytra oblong ; wings very short, nearly completely covered by the
sides of the pronotum, not reaching to apex of abdomen. Femora
scabrous-granulate ; anterior femora foliaceous dilate, dorsal margin
.serrulate, terminating in minute denticle; ventral margin serrulate,
sublobate ; middle femora above serrulate, subundulate, sinuate at
the apex, below distinctly amplicate backward and serrate-lobate,
being deeply bi-incised near the apex and distinctly lobate ; pos-
terioi- femora rugose-scabrous on the external pagina ; superior
margin strongly serrate, terminating in a denticle ; superior areas
before the knee bearing two fuscous acute denticles, the outer one
less erect ; ventral margin crenate quinque-lobate ; the inferior ex¬
ternal Carina inconspicuous and minutely serrulate, often bearing
minute tubercle near the middle. Colour greyish-fuscous or
ferruginous, the superior margin of crest marked with fuscous.
Length of pronotum, cJ 11'5 mm. ; posterior femora, 5-5*5 mm.
$ 1 1 *8 mm. ; posterior femora, b mm.
Four examples from N.E. Rhodesia in the Oxford Uni¬
versity Museum, collected by S. A. Neave at the follow¬
ing points : — Three from Upper Kalungwisi Valley,
4200 ft.. No. 2140, September 8, 1908 ; one from Chisinga
Plateau, Kalungwisi district, 4500 ft., No. 2139, Sep¬
tember 25, 1908.
One specimen has a singular abnormality in presenting
a deep angulate incision excavated from the front part of
the pronotal crest near the frontal apex. This individual
seemingly was born with this peculiarity, and is possibly a
mutation.
I take pleasure in dedicating this interesting species to
S. A. Neave, who contributed the specimens to the Oxford
University Museum.
Genus Cladoramus, Hancock.
Trans. Entom. Soc. London, p. 217, 1907.
As shown in the preceding synoptical table, this genus
occupies a position midway between Xero2yhyIlmi, Fairm.,
and Trachytettix, Still.
C. crenulatus, Hanc.
Six examples are represented in the present material
collected and presented to the Oxford University Museum
by S. A. Neave. They were taken at the following
Tetriginae in the Oxford, University Museum. 355
points in N.E. Rhodesia : — Five from Chisinga Plateau,
Kalungwisi, 4500 ft., No, 2149-2153, September 17, 1908,
and one from Mporokoso, 4500 ft., No. 2154, Septem¬
ber 2, 1908. The type female example from which the
original description was drawn came from East Loangwa
Rhodesia.
Genus Acmophyllum, Karsch.
Entom. Nachrichten, p. 21, 1890.
In the preceding synoptical table of the genera of
African Cladonotae I have classed both Acmophyllum
xerophylloides and A. conradti, described by Bolivar, under
the new genus Medinophyllum. The following table
therefore is confined to the three remaining species
enumerated below : —
TABLE OF SPECIES.
1. Inferior external carina of posterior fe¬
mora quadrilobate ; anterior margin
of pronotal crest oblique, above before
the middle summit flattened-undu-
late ; length of pronotum, $
16 '3 mm .
1.1. Inferior external carina of posterior
femora distinctly unilobate at the
middle.
2. Anterior margin of pronotum ver¬
tical, above before the middle
summit sinuate ; length of prono¬
tum, 9 17 mm .
2.2. Anterior half of pronotal margin
above before the middle regu¬
larly rounded ; length of prono¬
tum, 18 mm .
A. excavatum, sp. nov.
Facial costa barely scutelliform, the lateral carinae entire, acute
angulate above gradually divergent to the median ocellus, com-
presso-elevated between the antennae yet in profile flattened ; face
distinctly sinuate below at the ocellus ; antennae inserted far below
the eyes, equal in distance to one-half their height ; vertex on each
side next to the eyes bearing a small elevated tubercle. Pronotum
- A A 2
undulatum, Karsch.
excavatum, sp. nor.
nigro-punctatum, Bol.
356 Dr. J. L. IHBcock’s Third Paper on the
strongly compresso-foliaceous almost wholly cristate, more highly
elevated above at the middle summit ; the superior margin of ante¬
rior half abruptly elevated, in front vertical undate and above
strongly unisinuate-subundate ; the posterior half backward gently
deflexed undulate and near the apex concave, posteriorly lengthily
extended beyond the apices of the hind femora and acute ; superior
margin of crest marked with fuscous, viewed from above sinuous ;
anterior process below arcuate irroduced above the head, the apex
little straight excised ; lateral lobes of pronotum posteriorly
bisinuate, posterior angles elongate, obliquely extended outward
and rounded. Elytra oblong subacuminate ; wings fully explicate,
extended backwards to the pronotal apex. Anterior femora com¬
presso-foliaceous, superior margin subtrilobate, below quadrilobate,
or the two middle lobes fused in one ; middle femoral carinae above
slightly undulate, below compresso-ampliate, sinuate, and bearing a
small apical lobe ; posterior femoral carinae above subentire,
minutely serrulate, marked by small fuscous dots, and near the
apices terminating in acute denticle ; inferior carina of external
pagina strongly compressed-incrassate, viewed from above little
undulate and bearing a strong projecting median lobe ; ventral
margin entire or indistinctly undulate and marked with fuscous ;
first articles of posterior tarsi equal in length to the third, the first
and second pulvilli minute, the third distinctly longer. Colour,
greyish-rufescent, sparingly speckled with fuscous, the tibiae in-
fuscated toward the apices, joints and apices of tar.si black.
Length of pronotum, 5 17‘2 mm. ; height at middle, 7 mm. ;
posterior femora length, 7 mm.
Locality : Bitje Ja River, S. Cameroons (Rosenberg), in
the author’s collection.
Genus Rhopalotettix, nov.
Body smoothly granulose ; eyes globose ; face strongly retreating ;
vertex strongly obliquely produced in an elongate cephalic process,
cuspidate at the apex, viewed from above longitudinally sulcate,
sides entire ; vertex, between the eyes forward broadly fossulate,
middle not carinate. Antennae long, longer than the head, inserted
below the eyes ; frontal costa narrowly sulcate, superior paired
ocelli placed between the lower third of the eyes ; median ocellus
placed far below the eyes. Pronotum anteriorly truncate, posteriorly
lengthily acuminate-subulate, extended beyond the femoral apices ;
dorsum narrow between the shoulders, deplanate ; median carina
low, scarcely elevated ; anterior carinae .short parallel ; humeral
Tctriginae in the Oxford University Museum. 357
angles nearly straight ; posterior angles of the lateral lobes turned
down obtuse, yet obliquely excised. Elytra small and widened
posteriorly, apex widely rounded ; wings perfectly explicate.
Anterior femora strongly elongate, superior carinae basally com¬
pressed, acute and minutely serrulate ; middle femoral carinae above
compressed, terminating in an apical denticle ; posterior femoral
carinae above terminating in a denticle, below longitudinally
curvate ; genicular denticle produced ; posterior tibiae narrow, the
canthi minutely serrulate and bearing many small feeble denticles ;
first and third articles of posterior tarsi equal in length, the third
joint distinctly clavate.
This genus resembles Bhynchotettix, Hancock,* but
differs in the cuspidate apex of the produced cephalic
process, in the absence of a median carina on vertex, in
the prominent globose eyes, in the presence of elytra and
wings, in the absence of a spine arming the posterior
angles of the lateral lobes of pronotum, in the equal
length of the first and third tarsal articles, and in the
distinctly clavate form of the third joint of the posterior
tarsi,
B. clavipes, sp. nov.
Body smooth granulate ; eyes prominent and strongly globose ;
face strongly oblique ; vertex strongly obliquely produced in an
elongate cephalic process, in profile extended beyond the eyes more
than the greatest length of one of them, in width equal to nearly
two-thirds the height of one of the eyes, and terminating in a
deflexed acute spine; process ventrally compressed, the middle
forming the facial median carina above, basally sinuate between the
lower part of the eyes; the process extension viewed from above
narrower than the vertex backward, longitudinally sulcate, sides
parallel entire but cuspidate at the apex ; vertex between the eyes
little narrowed forward, broadly transversely fossulate, nearly equal
in width to one of the eyes, middle not carinate; superior paired
ocelli placed between the lower third of the eyes ; median ocellus
situated far below the eyes, the distance from the median ocellus
to the eyes much greater than the distance between the eyes ; frontal
costa narrowly sulcate, forked between the paired ocelli above,
downwards compressed scarcely elevated; antennae long and
slender, longer than the head, consisting of fourteen articles, the
first basal articles crassate, second smaller globose, the next four
* Trans, Entom. Soc. London, p. 228, 1907.
358
Dr. J. L. Hancock’s Third Paper on the
short and indistinct, but the rest from the seventh to the terminal
apical article strongly elongate. Pronotum anteriorly truncate,
posteriorly lengthily acuminate-subulate e.xtended beyond the hind
femoral apices; dorsum deplanate, between the shoulders scarcely
widened ; humeral angles substraight, transversely subrounded ;
median carina percurrent but low barely elevated ; anterior prozonal
carinae short and j^arallel ; lateral lobes of pronotum somewhat
small, margin anteriorly arcuate; posterior angles turned down
and obliquely excised. Elytra small, widened backward, the apices
rounded, the central external two-thirds marked with black ; wings
perfectly explicate, extended barely beyond the pronotal apex.
Anterior femora strongly elongate, superior margins little com¬
pressed basallj' and serrulate, below entire ; middle femora elongate,
superior carinae comi^ressed minutely serrulate, and terminating in
an apical spine, ventral carinae little compre.ssed subentire or barely
undulate ; posterior femora slender, superior carinae forward arcuate,
posteriorly terminating in distinct antegenicular denticle, very
minutely serrulate, genicular apical lobe distinctly produced ; ventral
carinae slightly curvate, carinae of external paginae distinct and
between them bearing oblique granulate rugae; posterior tibiae
narrow, the canthi minutely serrulate and bearing many very small
denticles ; first and third articles of posterior tarsi equal in length ;
the third tarsal article distinctly clavate, the three pulvilli of the
first tarsal article equal in length and obtuse below. Colour dark
cinero-griseous, the hind tibiae infuscate, but light annulate behind
the knees.
Entire length of body, $ , 21‘2 mm. ; pronotum, 13‘8 mm. ;
posterior femora, 6 mm. ; hind tibiae, 5 mm. ; antennae, 6 mm. One
example in the author’s collection.
Locality: Padang Pandjaiig, West Sumatra (Rolle).
This species is remarkable in the form of the long
antennae, produced vertex, globose eyes, and clavate form
of the third article of the posterior tarsi. Type in the
author’s collection.
Genus Thoradonta, Hancock.
Trans. Entom. Soc. Loudon, p. 407, 1908.
This genus is represented by small forms with sub¬
quadrate vertex; elongate antennae inserted below the
eyes ; the pronotum anteriorly truncate, posteriorly abbre¬
viated acuminate ; with the median carina of pronotum
sinuate or bicristate ; dorsum widened between the obtuse
Tetriginae in the Oxford University Museum. 359
carinate humeral angles, and above bearing a super¬
numerary carinula on each side ; the lateral lobes out¬
wardly laminate, the posterior angle dentate or spined ;
elytra ovate and wings present; anterior and middle
femora elongate, margins slightly compressed, above sub¬
entire, below undulate ; the first and third articles of the
posterior tarsi subequal or the first a little longer than the
third, the first two pulvilli often acute subspiculate.
T. dentata, Hancock.
One female in the Oxford University, No. 269, from
Selangor, Kuala Lumpur, Malay Peninsula, collected and
presented by H. C. Pratt. The type in the Oxford
University Museum is from Penang or Prince of Wales
Island.
Genus Dasyleurotettix, Helm.
Proc. Acad. Nat. Sci. Philadelphia, p. 68, 1904.
D. eurriei, Rehn. (Plate XLIX, fig. 3.)
Ten examples, Nos. 2142-2148, from N.E. Rhodesia, in
Oxford University Museum, collected and presented by
S. A. Neave; nine from the Upper Kalungwisi Valley,
4200 ft., September 11, 1908; one from Chinsali, 4300 ft.,
April 11, 1908.
Genus Tetrix, Latreille.
Hist. Nat. Crust. Ins., vol. hi, p. 284, 1802.
T. hipunctata, Linne.
Two examples (varieties) in Oxford University Museum
from Persian Gulf, (?) Bussorah.
T. suhulata, Linne.
Two long-wing and one short-wing (varieties) in the
Oxford University Museum from Persian Gulf, (?) Bussorah.
Genus Euparatettix, Hancock.
Spolia Zeylanica, vol. ii, p. 145, 1904.
E. interruptus, Brunner.
One ? example. No. 1106, from Malay Peninsula,
S.W. Johore, Kukub, collected and presented by H. N.
360 Dr. J. L. ^ncock’s Third Paper on the
Ridley to the Oxford University Museum, This specimen
is nearly Avholly black, the tirst and second legs banded with
fuscous, while the black hind tibiae present a white
annulation behind the knees.
E. pulvillus, sp, nov. (Plate XLIX, figs, 4 and 4a.)
A slender species allied to Paratettix histrieus, Bol. Head ex-
serted ; vertex equal in width to one of the eyes, truncate, middle
carinate, frontal costal in profile above little subangulate produced
beyond the eyes, and between the antennae more subarcuate pro¬
duced, rather narrowly sulcate. Pronotuin punctate-granulate, con¬
stricted before the shoulders, and between the shoulders only slightly
widened ; anterior pronotal carinae short and parallel ; median
Carina percurrent, compressed, slightly undulate forward and
behind the shoulders, posterior pi’ocess extended far beyond the
femoral ajrices ; posterior angles of the lateral lobes subacute. Elytra
subovate and externally strongly reticulate ; wings caudate extend¬
ing beyond the pronotal apex two millimeters. Anterior and middle
femora elongate, nearly entire, but the inferior carinae of middle
femora scarcely undulate ; posterior femora slender, superior carina
serrulate terminating in a denticle before the knee, the external
area at the middle plainly bituberculate ; posterior tibiae ventrally
infuscate and the apical fourth fuscous ; first articles of posterior
tarsi equal in length to the third, the first two basal pul villi acute-
spiculate, the third nearly as long as the first and second together,
and flat below. Body below sparingly hirsute. Colour of body
testaceous marked with black on the disk of the pronotiim.
Entire length of body, J 15 mm.; pronotum, 12 mm.; posterior
femora, 5 '5 mm.
One example from Malay Peninsula, Selangor, River¬
side Estate, collected and presented to the Oxford Univers¬
ity Museum by H, C. Pratt.
E. angustivertex, Bol.
Three examples from N.E. Rhodesia in the Oxford
University Museum — two from Upper Kalungwisi Valley,
4200 ft., Nos. 2159 and 2163, September 11,1908; one
from Lofu River, 3500 ft.. No. 2155, August 17, 1908,
pollected and presented by S. A. Neaye.
Tetriginae in the Oxford University Museum, 3G1
Genus Paratettix, Bolivar.
Ann. Soc. Entoin. Belgique, xxxi, p. 270, 1887.
P. pictibs, sp. nov. (Plate XLIX, figs. 5 and 5«.)
Body small, coloured fuscous and light variegated. Head not ex-
serted, dorsum above little rugose-granulate ; vertex little narrower
than one of the eyes, little narrowed toward the front, viewed from
above not produced beyond the eyes ; frontal costa in profile slightly
projecting beyond the eyes above, but between the antennae subar-
cuate ; between the eyes indistinctly sinuate, the lateral rami
moderately separated ; antennae inserted between the lower angles
of the eyes ; eyes in profile subconoidal, the occiput covered. Pro-
notum anteriorly truncate, between the shoulders convex, little
widened, posterior process subulate, extended beyond the femoral
apices behind ; anterior prozonal carinae very short, quite widely
separated and parallel ; median carina marked with fuscous, little
compressed percurrent, subgibbulose near the front margin ; posterior
angles of the lateral lobes rather obliquely excised. Elytra ovate;
wings caudate and black. Femora elongate, little compressed,
margins entire and sparingly pilose ; middle femora in male nar¬
rowed toward the apices ; posterior femora elongate, margins minutely
serrulate, antegenicular and genicular denticles prominent ; posterior
tibiae black with white annulation behind the knees and marked
with white at the distal third ; second and third posterior tarsal
pulvilli subequal in length.
Entire length of body, ^ 9-6 mm ; pronotum, 8 mm. ; posterior
femora, 4 mm.
One example, No. 2158, from N.E. Rhodesia, Mporokosa,
4500 ft., August 31, 1908, collected and presented to Oxford
University Museum by S. A. Neave. This species is
nearly allied to wilverthi, Bolivar, differing in being more
rugose, smaller in stature, in not being depressed behind
the shoulders on the dorsum, and in the less depressed
median carina behind the anterior border. It also differs
in the equal length of the second and third posterior tarsal
articles.
Genus Prototettix, Bolivar.
Ann. Soc. Entom. Belgique, xxxi, p. 255, 1887.
As I interpret the African genus Prototettix, Boh, there
are three representatives, namely, iimpressus. Boh, afri-
362 Dr, J. L. cock’s Third Paper on the
canus, Hanc., and hticculentus, sp. nov., described below.
The species foss^datiis, Boh, and lohdatus, Still, inhabit¬
ing South America, and formerly classed by Bolivar
in this genus, are quite different from the African forms.
As a result of a revised study of these insects the two
latter species were recently transferred to my genus Rytina-
tettix, as noted in the Transactions of the Entomological
Society of London, p. 416, 1908.
P. hucculentibs, sp. nov. (Plate XLIX., figs. 6 and im.)
Colour shining, brownish fuscous, obscurely light variegated,
sparingly sprinkled with various large tubercles. Body incrassate,
head not at all exserted, occiput covered ; vertex transverse, rugose,
twice the width of one of the eyes, frontal carinulae wanting ; in
profile the distinct median crassate carina of vertex fused with the
facial costa above, forming a rounded contour elevated above the
eyes and strongly arcuately jjroduced beyond them ; in front view
widely sulcate, little divergent downward toward the median ocellus ;
eyes conoidal in profile, the face below on each side tumid, antennae
inserted barely between the low^er angles of the eyes. Pronotum
above rugose sprinkled with large tubercles, anteriorly convex ;
dorsum very obtuse tectiform, barely impressed behind the
shoirlders ; anterior prozonal carinae low, little convergent back¬
ward ; humeral angles obtuse and indistinctly lineate carinate but
the lateral carinae behind on the process more distinct, entire ; median
carina acute, subpercurrent, arcuate anteriorly, depressed at the middle
and subconvex backward toward the apex, posterior process abbre¬
viated, subacute, not reaching to the knees of the hind femora ;
lateral lobes rugose, inferior margin nearly straight, little sinuate,
slightly reflexed outwards, posterior angles obliquely excised ; pos¬
terior superior sinus very small, nearly obsolete. Elytra small,
narrow, subacuminate towards the apices ; wings not visible or
wanting. Anterior femora elongate, above entire, below indis¬
tinctly unilobulate ; middle femora bicarinate, compressed, mar¬
ginal carinae above and below subtrilobulate ; posterior femora
elongate, the outer area incrassate, marginal carinae above and below
curvate, adorned with fuscous and minute flavous markings, minutely
serrulate-granulate ; antegenicular denticle acute, genicular denticle
wanting ; posterior tibiae incrassate, ampliate towards the apices,
external canthi bearing many strong spines, and about seven spines
arm the inner canthi ; tibiae black with small light annulation behind
the knees ; the first hind tarsal articles incrassate, longer than the
Tetriginae in the Oxford University Museum. 363
thii'd, all three pulvilli nearly equal in length and flat below, not at
all acute.
Entire length of the body, 5 H'® mni- ; pronotum, 9 mm. ;
posterior femora, 6 ’5 mm.
One example, No. 2162, from N.E. Rhodesia, Upper
Kalungwisi, 4200 ft., September 7, 1908; collected and
presented to Oxford University Museum by S. A. Neave.
Genus Coptotettix, Bolivar.
Ann. Soc. Entom. Belgique, p. 289, 1887.
Owing to the incomplete descriptions of some of the
species of African Coptotettix, the separation of the species
is made difficult. I have attempted to give a revised
table of the African species in the order in which they
seem to be more readily recognized.
TABLE OF SPECIES.
1. Antero-dorsal margin of pronotum
angulate
2. Body moderately crassate.
3. Antennae sliort, inserted between
and towards the middle of the
eyes, lateral carinae behind the
anterior margin of dorsum
obsolete ; vertex obtuse-angu-
late, rounded produced before
the eyes ; third pulvilli of pos¬
terior tarsi longer than the
second . awjulatus, Bol.
3.3. Antennae elongate, inserted be¬
tween the lower part of the
eyes ; lateral carinae near the
anterior margin of dorsum pre¬
sent, parallel ; vertex rounded
before the eyes ; third and
second pulvilli of posterior
tarsi subequal in length . . . mi'iiutus, Bol.
2.2. Body narrower, rugose, legs
shorter, posterior process
extended far beyond the
apices of posterior femora ;
wings barely passing pronotal
apex ; dorsum anteriorly covered
with elongate rugae and bi-
364
Dr. J. L. ^ncock’s Third Paper on the
rugate between the shoulders ;
elytra oblong, narrow, apices
narrowly rounded . discolor, Bol.
1.1. Antero-dorsal margin of pronotum
very slightly convex or trun¬
cate.
4. Antero-dorsal margin slightly con¬
vex ; elytra rather wide, elong¬
ate, apices acutely angulate,
externally lightly punctate im¬
pressed ; first articles of the
posterior tarsi little longer than
the third, the third pulvilli
longer than the first ; pulvilli
flattened below not at all
spiculate . convexus, S2). nov.
4.4. Antero-dorsal margin truncate.
5. Posterior pronotal j^rocess not
abbreviated ; wings extended to
pronotal apex or caudate.
6. Median carina of pronotum j^er-
current, scarcely elevated, un¬
dulate, gibbulose anteriorly
before the shoulders ; dorsum
sparingly rugulose and between
the shoxilders bearing elevated
lines ; posterior angles of lateral
lobes widely rounded ; vertex
behind the fossae bearing dis¬
tinct transverse jslaca ; first pul¬
villi of jiosterior tarsi spinose . bilineatus, Bol.
6.6. Median carina of pronotum pluri-
interruj)ted, scarcely elevated,
dorsum covered with shining
tubercles and short irregular
lines ; posterior process not or
scarcely jjassing the apices of
posterior femora ; wings often
caudate . annulipes, Karsch.
5.5. Posterior process of jironotum
not reaching to the apices of
hind femora ; wings shortened,
hardly reaching to the ab¬
dominal apex.
Trans. Tiit. Soc.Lonci..J!JlO. Pl.XLIX.
H.Kniglit del.
West,!Newjnas.ii imp.
NEW SPECIES OF TETRIGINAE.
Explanation of Plate XLIX.
Fig. 1. Xerophyllum extensum, sp. nov. ; side view of body.
2. Xerophyllum neavei, sp. nov. ; side view of body.
3. Dasyleurotettix curriei, Rehn ; side view of body.
4. Euparatettix 2}ulmllus, sp. nov. ; side view of body ; 4a,
vertex of head from above.
5. Paratettix jnctus, sp. nov. ; side view of body ; 5a, vertex
of head from above.
6. Prototettix hucculentui^, sp. nov. ; side view of body ; 6a
vertex of head from above.
7. Coptotettix convexus, sp. nov. ; side view of body ; 7a,
vertex of head from above.
Tetriginae in the Oxford University Mnsenm. 365
7. Elytra oblong and black, anterior
legs rufous . rufipes, Bol.
7.7. Elytra ovate ; length of male pro-
notum 6 mm . parvulus, Hanc.
? mfuscata, Walk, (not
able to classify).
C. convexus, sp. nov. (Plate XLIX, figs. 7 and 7a.)
(See preceding table). Colour fusco-testaceous, hind femora of
female often testaceous, of male variegated ; vertex narrowed for¬
ward, scarcely wider than one of the eyes, middle carinate, on each
side oblong fossulate, and bearing a transverse ruga behind ; frontal
carinulae roundly flexed backward each side of mid-carina, not
projecting beyond the eyes; frontal costa arcuate produced before
the eyes, viewed in front widest between the antennae ; antennae
inserted between the lower third of the eyes. Pronotum anteriorly
barely convex, posteriorly lengthily subulate extended beyond the
hind femoral apices ; dorsum coarsely granulose, bearing small
shining elongate tubercles, and bi-rugose, convex between the
shoulders but behind the shoulders little subdepressed ; subconcave
backward on the process ; median carina indistinctly undulate,
scarcely elevated, subgibbulose between the sulci forward ; lateral
prozonal carinae forward low, granulate and parallel ; humeral
angles widely obtuse and scarcely distinct ; posterior angles of
lateral lobes oblique, apex subtruncate. Elytra rather wide, elongate,
apices acutely angulate, externally lightly punctate impressed ; wings
fuscous or little testaceous, extended backward barely beyond the
pronotal apex. Anterior and middle femora elongate, margins entire
(more compressed in the male) ; posterior femora incrassate, extern¬
ally rounded, granose, carinae above and below entire and sub-
granose ; first articles of posterior tarsi little longer than the third,
the third pulvilli longer than the first, the pulvilli flattened below
not at all spinose.
Entire length, $ 13 mm.; pronotum, 11 '8 mm; posterior
femora, 6 mm. ^ 12 mm. ; j^ronotum, 10'5 mm. ; posterior femora,
5‘5 mm.
Four examples, Nos. 2156, 2157, 2160, and 2161, in the
Oxford University Museum, from N.E. Rhodesia, Upper
Kalungwisi Valley, 4200 ft., September 1908, collected
and presented by S. A. Neave.
Explanation of Plate XLIX.
[See Explanation facing the Plate.]
( 30)6 )
X. Descriptions of Micro-Lepidoptera from Mauritius and
the Chagos Isles. By E. Meyrick, B.A., F.R.S.
[Read May 4th, 1910.]
The specimens from Mauritius now described were col¬
lected by Lieut.-Colonel N. Manders during bis residence
in the island some years ago. They are of much interest,
owing to the highly specialised character of the fauna, of
which little has been made known hitherto, so far as the
Micro-Lepidoptera are concerned. I believe that most of
the native forest has now been destroyed, and the land
placed under cultivation ; but probably if a skilled collector
would carefully examine the limited portions of forest,
difficult of access, which still exist, and also the higher
regions of the mountains, many species might still be
recovered before extermination.
The Chagos Islands, though geographically not very
remote, have no zoological affinity to Mauritius, but belong
to the Indian region. The two species described here
were collected by Mr. T. Bainbrigge Fletcher.
By the kindness of the above gentlemen the types are
in my collection.
PYRAUSTIDAE.
ScoPARiA, Haw.
Scop aria lenigna, n. sp.
$ $. 15-16 mm. Head and thorax fuscous mixed with whitish.
Palpi 2, dark fuscous, apex mixed with whitish, basal joint white.
Antennal ciliations of ^ minute. Abdomen grey-whitish. Fore¬
wings elongate-triangular, costa slightly arched, faintly sinuate
in middle, apex obtuse, termen gently rounded, oblique ; white,
irregularly irrorated with fuscous ; an irregular dark fuscous
almost basal fascia ; lines thick, white, first slightly curved, rather
oblique, edged posteriorly with dark fuscous suffusion, second
somewhat irregular, slightly curved, nearly direct, subterminal
entire, almost touching second in middle ; orbicular and elavi-
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.)
Mr. E. Meyrick’s Descri'ptions of Micro-Lepicloptera. 367
form small, round, cloudy, dark fuscous, vertically placed ; discal
obscurely 8-shaped, outlined with dark fuscous, upper half suf¬
fused with fuscous and connected with a dark fuscous spot on
costa, lower half incomplete and suffused with white ; cilia white,
basal third barred with fuscous. Hindwings pale brassy-whitish-
ochreous ; cilia white.
Mauritius ; four specimens. A distinct species, rather
of European type.
PTEROPHORIDAE.
Trichoptilus, Wals.
Trichoptihis loaJilbergi, Zell.
Mauritius ; two specimens. Widely distributed, prob¬
ably artificially introduced.
Platyptilia, Hb.
Platyptilia hrachymorpha, Meyr.
Mauritius ; two specimens. Also very widely spread.
Platyptilia censoria, n. sp.
^ . 17 mm. Head and thorax pale brownish-ochreous mixed with
whitish (damaged). Abdomen pale ochreous, marked with dark
fuscous suffusion on sides towards middle. Forewings cleft from
beyond f, segments broad, ternien of first concave, of second some¬
what rounded, oblique ; whitish-ochreous mixed with whitish and
sprinkled with brownish : an acute-triangular dark fuscous spot
before cleft, apex directed anteriorly ; a straight whitish subterminal
line obscurely indicated, followed by stronger fuscous irroration :
cilia whitish, on termen with basal third fuscous limited by a
blackish line, with dark fuscous patches beneath lower angles of
both segments and above upper angle of second, and two fuscous
black-tipped scaleteeth on dorsum before and beyond cleft, with a
few scattered black scales. Hindwings with segments moderately
broad, termen of second sinuate ; grey; cilia pale greyish, with a
rather large triangular darker grey black-tipped scaletooth on
middle of dorsum, and a few scattered black scales.
Mauritius ; one specimen. Not closely approaching
any other.
368
Mr. E. Meyrick’s Descriptions of
EUCOSMIDAE.
Spilonota, Stepli.
Spilonota rhothia, n, sp.
^ . 15-16 mm. Head and thorax fuscous or rather dark fuscous,
slightly sprinkled with whitish. Antennae with notch at Ab¬
domen fuscous, anal tuft fuscous-whitish. Forewings elongate,
somewhat dilated posteriorly, costa gently arched, with strong fold
reaching from base to |, apex obtuse, termen slightly rounded,
rather oblique ; brown, strigulated or mixed or almost wholly
suffused with dark fuscous ; costa posteriorly with four faint pairs
of strigulae of whitish irroration ; a broad irregular streak of whitish
suffusion extending along dorsum from near base to tornus, before
middle with a projection upwards formed by a tuft of whitish scales,
before tornus interrupted by a spot of groundcolour ; ocellus indi¬
cated by indistinct leaden-grey margins terminated beneath in the
whitish suffusion, and sometimes containing two or three undefined
longitudinal blackish marks ; three or four small undefined blackish
marks separated by whitish irroration on termen : cilia fuscous more
or less suffused with dark fuscous, with rows of whitish points.
Hindwings wdth 3 and 4 stalked ; grey, rather thinly scaled, darker
posteriorly, veins dark fuscous ; cilia pale fuscous, more whitish
towards tips.
Mauritius, one specimen ; also three from Pusa, Bengal
(Lefroy), and Maskeliya, Ceylon (Pole), in February and
March ; bred from larvae feeding on leaves of Psidium
guava, with which tree the insect has doubtless been intro¬
duced into Mauritius. S. Wals., from the West
Indies, must be nearly allied to this, but is described as
having two or three dark fuscous tufts about the sub¬
median fold instead of the whitish tuft of this species, and
a pale costal patch at end of costal fold.
Rhopobota, Led.
Rhopolota, physalodes, n. sp.
^ 9 • 13-14 mm. Head whiti.sh-ochreous, sometimes tinged with
grey. Palpi pale ochreous, sometimes mixed with blackish. Thorax
pale ochreous mixed with greyish, shoulders suffused with blackish.
Abdomen grey. Forewings elongate-oblong, costa gently arched,
in (4 with dense erectile fringe of hairs lying on upper surface
from base to f, apex obtuse, termen vertical, somewhat rounded-
Micro- Lepidoptera from Mauritius and Ghagos Isles. 369
prominent on lower half; pale oclireous, sometimes mixed with
light grey-greenish and sprinkled with whitish, strewn with un¬
defined strigulae of fuscous and dark fuscous irroration ; quad¬
rate spots outlined or suffused with dark fuscous on costa at base,
J, and middle ; in an oval fovea in disc from near base to
filled with rough scales, edged posteriorly with dark fuscous suf¬
fusion ; cilia dark grey, sprinkled with whitish points, towards
base suffused with pale ochreous. Hind wings grey; veins dark
grey ; cilia grey.
Chagos Islands, I. du Coin, in June, two specimens;
also one from Galle, Ceylon, in April (Fletcher).
GELECHIADAE.
Brachmia, Hb.
Brachmia aiUonoma, n. sf).
$ . 12 mm. Head pale ochreous. Palpi whitish-ochreous, ter¬
minal joint with a few dark fuscous scales indicating basal and
supramedian rings. Antennae pale ochreous suffusedly ringed
with fuscous. Thorax light brownish-ochreous. Abdomen whitish-
ochreous, tinged with grey. Fore wings elongate, costa gently
arched, apex obtuse, termen very obliquely rounded ; 7 to apex, 8
and 9 out of 7 ; pale yellow-ochreous suffusedly mixed with light
brownish ; sligmata dark fuscous, plical somewhat before first discal ;
a small cloudy fuscous spot on dorsum beneath second discal ; a row
of dark fuscous dots round posterior part of costa and termen : cilia
whitish-ochreous tinged with brownish. Hindwings pale greyish ;
cilia ochreous-grey-whitish.
Chagos Islands ; one specimen.
METACHANDIDAE.
Metachanda, Meyr.
Metachanda Jimhriata, n. sp.
$. 15-16 mm. Head and thorax dark purple-fuscous, with an
ochreous-yellow line above eyes. Palpi ochreous-yellowish mixed
with dark fuscous. Fore wings elongate, costa gently arched, apex
obtuse, termen very obliquely rounded ; dark purple-fuscous ; an
irregular transverse somewhat oblique blackish bar representing first
TRANS. ENT. SOC. LOND. 1910. — PART III. (NOV.) B B
870
Mr. E. IMeyrick’s Dmcrijitions oj
(liscal and plical stigmata, edged posteriorly with light ochreous-
yellowish, not reaching costa or dorsum ; an undefined oval blackish
spot representing second discal, enclosing an incompletely indicated
8-shaped mark of light ochreous-yellowish suffusion; a light yel¬
lowish spot on costa at whence an outwards-curved series of pale
yellowish dots preceded by some blackish suffusion runs to tornus :
cilia grey, basal third dark fuscous obscui’ely barred with pale
yellowish. Hind wings whitish, posterior half grey; cilia grey,
becoming ochreous-whitish towards tornus and dorsum.
Mauritius, Les Mares (a swampy piece of original
forest), in December ; two specimens.
Taragmarcha, u. g.
Head with appre.ssed scales ; ocelli present ; tongue developed.
Antennae 1, ba-sal joint long, slender, without pecten. Labial palpi
very long, recurved, second joint expanded with long rough pro¬
jecting scales above and towards apex beneath, tei'ininal joint much
longer than second, slender, acute. Maxillary palpi very short,
filiform, appressed to tongue. Posterior tibiae clothed with rough
scales above. Forewings with 2 from angle, 3 absent, 7 absent, 11
from middle. Hindwings f, elongate, apex obtuse, termen rounded,
cilia 2 ; 3 and 4 connate, 5 nearly parallel, 6 absent.
Taragmurcha laqueata, n. sp.
9 . 17 mm. Head and thorax rather dark glossy fuscous, face
mixed with white. Palpi white, second joint mixed with dark
fuscous, anterior edge of terminal joint dark fuscous. Antennae
whitish ringed with dark fuscous. Abdomen grey. Forewings
elongate, co.sta gently arched, apex obtuse, termen very obliquely
rounded ; rather dark glossy 'fuscous ; a white streak along basal
third of costa, thence slightly outwards-curved to dorsal beyond
middle ; an inwards-curved white line from f of costa to dorsum
before tornus, dilated on costal half into an oblique patch ; a series
of four elongate confluent whitish spots crossing wing close before
apex : cilia grey-whitish, towards tornus fuscous, basal third white
round apex and termen to near tornus, and with white marks on
extremities of postmedian line. Hindwings rather dark grey ; cilia
grey.
Mauritius, Les Mares, in December, one specimen ;
also a second much damaged, apparently without the
costal extension of the white anterior streak, but probably
identical.
Micro- Lepido'j)tera from Mauritius and Gliagos Isles. 371
Tanychastis, n. g.
Head with appressed scales; ocelli absent; tongue developed.
Antennae over 1, in ^ simple, basal joint moderate, without pecten.
Labial palpi long, recurved, second joint with appressed scales,
terminal joint shorter than second, slender, acute. Maxillary palpi
very short, filiform, appressed to tongue. Posterior tibiae smooth-
scaled, with some rough scales towards base above. Forewings with
2 from towards angle, 7 absent, 11 from before middle. Hindwings
£-, elongate-ovate, cilia 1 J; 3 and 4 connate, 5 somewhat approximated,-
6 absent.
Tanychastis lysiyama, n. sp.
^ . 9 mm. Head and thorax dark bronzy-fuscous. Palpi, antennae,
and abdomen dark fuscous. Fore wings elongate, rather narrow,
costa anteriorly straight, po.steriorly gently arched, apex rounded-
obtuse, termen rounded, rather strongly oblique ; dark fuscous ; a
somewhat irregular white transverse line from costa at reaching
across wing : a fine white inwards-curved line from f of costa to
tornus, interrupted beneath costa and above toimus : cilia dark
fuscous, tips whitish at apex. Hindwings and cilia dark fuscous.
Mauritius; one specimen.
Ancylometis, Meyr.
Ancylomctis trigonodcs, Meyr.
Mauritius; three specimens. In the hindwings veins
3 and 4 are sometimes coincident, but 5 remains separate.
Ancylometis dilucida, n. sp.
^ . 16 mm. Head, palpi, antennae, and thorax dark fuscous.
Abdomen light fuscous, anal tuft ochreous. Forewings elongate,
costa gently arched, apex obtuse, termen obliquely rounded ; dark
purplish-fuscous, with a few scattered whitish-ochreous scales : a
transverse ochreous-yellowish streak about i, suffused posteriori}^
angulated near dorsum, sending from its angle an undefined streak
of whitish-ochreous suffusion to dorsum beyond middle ; an un¬
defined patch of blackish suffusion in disc before middle, a smaller
spot about f (these represent the stigmata), a spot between and
beneath these, and a spot on dorsum at |, all with some adjacent
groups of whitish-ochreous or light yellowish scales ; a cloudy
whitish-ochreous transverse mark on costa at |, whence an abruptly
curved series of scattered whitish-ochreous scales, preceded by some
blackish suffusion, runs to tornus : cilia fuscous, on basal half dark
372
Mr, E. Meyrick’s Descriptions oj
fuscous with some whitish-ochreous scales at intervals. Hinclwings
with ridge and groove along lower margin of cell ; pale fuscous,
becoming fuscous on posterior half ; cilia whitish-fuscous, becoming
whitish-ochreous on lower part of termen and dorsum, with fuscous
subbasal shade.
Mauritius ; one specimen.
COSMOPTERYGIDAE.
Gosmopteryx, Hb,
Cosmopteryx dacryodes, n. sp.
7 mm. Head dark bronzy-fuscous, crown with three fine
white lines, face pale bronzy. Palpi white lined with black.
Antennae blackish, towards base lined with white, four apical joints
white, then five black, two white, one black, two white. Thorax
dark bronzy-fuscous, with three fine white lines. Abdomen greyish-
ochreous, anal tuft black on sides. Posterior tibiae black, with white
median and apical rings, and a silvery ring between these. Fore-
wings narrowly lanceolate, apex slenderly very long-produced, cau¬
date ; blackish ; dorsal edge finely white towards base ; subcostal,
median, and subdorsal very short snow-white dashes arranged in an
oblique series about I, widely remote from base and band, subcostal
fine, others thicker than usual ; a broad orange postmedian trans¬
verse band, anteriorly edged by a vertical golden-metallic fascia
followed above middle by a minute black speck, and posteriorly on
lower half by a golden-metallic spot edged with black anteriorly,
above posterior edge of which is a blackish dot on costa and some
whitish sufl’usion in costal cilia, between these a narrow orange
projection extends to termen, and is continued as a white streak
along termen to apex : cilia dark violet-grey. Hindwings and cilia
dark violet-grey.
Mauritius ; one specimen. Probably nearest to the
African C. cognita, Wals.
Cosmoptcryx mimetis, Meyr.
Mauritius ; two specimens. A very widely distributed
species, probably attached to some plant of cultivation.
Stagmatophora, HS.
Stagmatophora coriaccUa, Snell.
Mauritius ; four specimens, bred from larvae feeding
in the pods of the cotton-plant, with which the species has
been introduced from India. - ... -
Micro- Lepidoptera from Mauritius and Chagos Isles. 373
Idioglossa, Wals.
Idioglossa higemma, Wals.
]\Iauritius; one specimen.
OECOPHORIDAE.
Epiphractis, Meyr.
Epiphractis ampliitriclia, n. sp.
22 mm. Head and thorax rosy -grey. Palpi dull flesh-coloiu
irrorated with grey. Antennae simple, grey. Abdomen light grey,
anal tuft grey-whitish. Forewings suboblong, costa rather strongly
arched, apex obtuse, termen nearly straight, little oblique : on lower
surface with a fringe of extremely long hairs directed downwards
from vein 12 except towards extremities; dull light brownish-
crimson, with some scattered fuscous scales ; an inwardly oblique
purplish-fuscous streak from dorsum beyond middle, reaching J
across wing ; a cloudy purplish -fuscous dot in disc beyond f, and
a short inwardly oblique streak from tornus : cilia light greyish -
crimson. Hindwings ochreous-whitish, slightly crimson-tinged to¬
wards termen ; a long subcostal pencil of ochreous-whitish hairs
lying along upper margin of cell beneath forewings ; cilia ochreous-
whitish.
Mauritius, Les Mares, in December; one specimen.
GRACILARIADAE.
Acrocercops, Wallgr.
Acrocercops macrochalca, n. sp.
. 10 mm. Head white. Palpi white, second joint dark fuscous
towards apex, with a short apical projection of scales beneath.
Thorax white, patagia golden-fulvous. Abdomen light ochreous-
grey. Forewings very narrowly elongate, apex rather short-pointed ;
golden-orange-fulvous ; costal third occupied by a suffused shining
brassy-golden streak from base to near apex ; three narrow rounded
snow-white spots lying along dorsum, strongly black-edged above,
connected on dorsal edge, first short, from base, second longest, third
reaching tornus, a fourth similar disconnected spot along termen ; a
white striga along posterior part of costa into apex : cilia light
greyish-ochreous, base suffused with orange-fulvous, at apex with
374
Mr. Fl^^ey rick’s DesrriptAons of
a fine white strigula edged with blackish, on costa with a blackish
basal line. Hindwings grey ; cilia greyish-ochreous.
Mauritius ; one specimen.
GLYPHIPTERYGIDAE.
Chokeutis, Hb.
(yhorcvtis hjerkandrcUa, Tlinb.
Mauritius; two specimens.
Glyphipteryx, Hb.
GJyphi'fttcryx ditiorana, Walk.
Mauritius; two specimens.
TINEIDAE.
Bedellia, Stt.
Bcdellia sommdenidla, Z.
Mauritius ; one specimen.
Lyonetia, Hb.
Lyonctia carcinota, n. sp.
$ . 10 min. Head, palpi, antennae, and thorax silvery-white,
scales of crown slightly rough behind hllet. (Abdomen broken).
Forewings very narrowly elongate, short-pointed, apex slightly
produced, acute ; light glossy ochreous ; basal third silvery-white ;
an oblique white wedgeshaped mark from dorsum beyond middle
of wing, anteriorly edged wnth dark grey, reaching f across wing,
and a similar smaller mark from termen beyond tornus ; five very
small white spots on posterior third of costa, last preceding a small
round black apical spot mostly in cilia ; cilia pale grey, round apex
suffused with whitish, with two diverging black apical hooks.
Hindwings rather dark grey; cilia grey.
Mauritius; one specimen. Allied to L. 'proholactis
from the Seychelles, and slightly more primitive in
character.
Micro- Lepidojptera from Mauritius and Ghagos Isles. 375
Hieroxestis, Meyr,
Hieroxestis 'phaeoclialca, Meyr.
Mauritius ; three specimens. Also in Reunion, the
Seychelles, and South Africa.
Hieroxestis irraematura, Meyr.
Mauritius ; four specimens. Also in South Africa.
Hieroxestis subcervinella, Walk.
Mauritius ; one specimen. Also in the Seychelles..
OiNOPHILA, Stph.
In some of the following species the frontal and occipital
tufts are drawn together so as to conceal the fillet, but the
structure of the head remains peculiar and characteristic,
the lower part of face and back of crown being smooth,
and the two tufts being really separate at origin. Besides
the type-species, which is now common to Europe and
Africa but probably originally African, and a second
described form probably scarcely distinct, there are only
three other species known, all from the Seychelles; the
five now added are therefore in themselves a majority of
the genus.
OinopMla siccata, n. sp.
. 9-10 mm. Head grey-whitish, occipital tuft with a grey bar.
Palpi grey-whitish, externally with a dark fuscous line. Antennae
whitish. Thorax whitish, with shoulders blackish, a black anterior
bar between patagia, and three posterior black dots arranged in a
triangle. (Abdomen broken.) Forewings elongate-lanceolate, ajiex
produced, acute, downturned ; pale grey sprinkled with whitish
points ; costa marked with irregularly placed short somewhat
oblique black strigulae ; a small round black spot on base of fold ;
a group of three or four black dots near beyond this, two above and
below fold at and two rather larger in disc at | and | ; a short
blackish dash towards costa at | ; apex suffused with whitish and
marked with black : cilia ochreous-grey-whitish, on costa towards
apex with a black basal line. Hindwings dark violet-grey, with a
whitish patch occupying basal | ; a long grey bairpencil from base
lying along costa beneath forewings ; cilia grey-whitish.
Mauritius ; two specimens.
37G
Mr. l^^Leyrick’s Descriptions of
Oinopliila trixysta, n. sp.
9 . 9 mm. Crown wliitisli-ochreous with a dark brownish central
strijie, occipital tuft whiti.sh-ochreous, dark fuscous in centre beneath,
fillet and face shining whitish, frontal tuft dark fuscous. Palpi
whitish. Thorax dark lirownish, with two whitish-ochreous stripes.
Alxlomen light fuscous. Forewings elongate-lanceolate, apex
slenderly long-produced, acute, dovvnturned ; rather dark bronzy-
brown ; three rather suffused whitish-ochreous longitudinal stripes,
viz. one costal from base to middle, one rather above middle from
base to J, and one dorsal somewhat mixed with groundcolour from
base to tornus; costal edge ochreous-whitish from about J to f,
where it forms a short tine very oblicpie strigula ; an elongate-
triangular whitish-ochreous spot along lower portion of termen ; an
ochreous-whitish dot on termen beneath apex : cilia pale whitish-
ochreous. Hindwings grey ; cilia whitish-ochreous-grey.
Mauritius ; one specimen.
Oinophila pcntacarpa, n. sp.
$ . 10 mm. Head bronzy-fuscous, tufts ochi’eous mixed centrally
with dark fuscous, face whitish-bronzy. Palpi fuscous, internally
whitish. Antennae greyish, towards base white sharply lined with
black. Thorax dark fuscous. (Abdomen broken.) Forewings elon¬
gate-lanceolate, apex somewhat produced, acute hardly downturned ;
dark purple-fuscous, strewn with scattered whitish-ochreous long
hairscales; five roundish whitish-ochreous spots, viz. three beneath
costa on anterior half, one beneath costa towards apex, and one on
tornus : cilia pale greyish-ochreous, base mixed with dark fuscous.
Hindwings light grey ; cilia pale greyish-ochreous.
Mauritius ; one specimen.
Oinophila fragosa^ n. sp.
^ 9 • 9-10 mm. Crown whitish-ochreous, sides whitish, tufts
united to conceal fillet, whitish-ochreous infuscated in middle, face
whitish. Palpi whitish, second joint with a fuscous lateral line,
terminal joint infuscated. Thorax whitish-ochreous with two white
stripes, patagia dark bronzy-fuscous. Abdomen greyish. Forewings
elongate-lanceolate, apex produced, acute, downturned ; dark bronz}'-
fuscous ; a white streak partially tinged with yellowish running
along dorsum and termen from base to near apex, broadest towards
posterior part of dorsum, upper edge somewhat pi'ominent at ^ and
forming an angular prominence at f, where it reaches half across
wing, beyond this very narrow ; two very oblique white strigae
Micro-Lepidoptera from Mauritius and Chagos Isles. 377
from costa posteriorly, not reaching termen ; extreme apex suffused
with blackish : cilia whitish, beneath apex suffused with fulvous at
base, opposite apex with a blackish subbasal line. Hindwings light
grey ; cilia gi’ey-whitish, greyer towards base.
Mauritius ; three specimens.
Oinophila syntricha, n. sp.
^ . 10 mm. Head ochreous-whitish, crown with a central yellow-
ochreous stripe, tufts united to conceal fillet, mixed with pale
ochreous. Palpi whitish-ochreous. Thorax fulvous-ochreous, with
two whitish stripes. Abdomen whitish-ochreous. Forewings
elongate-lanceolate, apex acute, slightly downtnrned ; fulvous-
ochreous, irregularly and suffusedly strigulated with ochreous-
whitish, especially along costa ; a more or less complete line of
black scales along submedian fold : cilia whitish-ochreous, suffused
with fulvous-ochreous towards base, round apex with a darker median
line. Hindwings pale grey ; cilia ochreous-whitish tinged with grey
towards base.
Mauritius ; two specimens.
Sapheneutis, Meyr.
SapJieneutis cubitalis, n. sp.
^ . 15 mm. Head and thorax pale greyish-ochreous tinged with
brownish and sprinkled with fuscous, shoulders suffused with dark
fuscous. Antennal ciliations minute. Palpi moderately long, pale
greyish-ochreous irregularly sprinkled with dai'k fuscous. Abdomen
pale gi’eyish, anal tuft pale ochreous. Forewings elongate, costa
gently arched, apex obtuse, termen slightly rounded, oblique ; all
veins separate, 7 to apex ; pale greyish-ochreous suffused with light
brownish ; costa dotted wdth dark fuscous on anterior half, and with
five small blackish spots on posterior half ; some scattered dark
fuscous strigulae towards costa anteriorly ; an irregular dark fuscous
streak from base above middle to J ; an irregular dark fuscous streak
from J of costa to before middle of dorsum, obtusely angulated out¬
wards below middle of wing ; a terminal series of blackish dots :
cilia ochreous-whitish tinged with brownish and sprinkled with
fuscous and dark fuscous, at apex with a dark fuscous spot. Hind¬
wings light grey with brassy reflections ; cilia ochreous-whitish
tinged with grey.
Mauritius ; one specimen.
November 8, 1910.
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( 879 )
XL The Behaviour of Coleoptera in time of Floods. By
Norman H. Joy, M.R.C.S., F.E.S.
[Read April 6th, 1910.]
In the spring of 1905 when I first saw Bianous coemdescens
alive in Soutli Devon, I was much struck by the extremely
rapid way it moved over the surface of a pool of water
near which I found it. An examination of its mode of
progression soon convinced me that it was not produced
by any movement of its legs, and I was much puzzled
until Commander Walker referred me to a paper which
had shortly before been published in France. As this
paper is of no great length, I think it best to quote it
in full. It appeared in “ Comptes Rendus Hebdomadaires
des Seances et Memoires de la Societe de Biologie,” 1905,
tome second, pp. 102-3, and is entitled “ Sur un mode
particulier de locomotion de certains Stenus,” par MM. G.
Billard et G. Bruyant.
“ Nous avons observe au bord des ruisselets all mentis
par les sources pures et froides de la montagne deux especes
de Stenus {S. tarsalis, Lj., et S. cicindeloides, Schell.) qui
presentent un mode particulier de locomotion k la surface
de I’eau.
“Habituellement accroches ils sont exposes k tomber sur
la nappe liquide qui coule au-dessous d’eux, parfois avec
une grande rapidite. Comme beaucoup d’autres insectes
ils peuvent marcher k la surface de I’eau, mais leur vitesse
de progression est alors trfes faible, et ils risqueraient
ainsi d’etre entrain^s au loin. Pour lutter contre la
rapidite du courant, ils usent du proced4 suivant. Ils
expulsent par I’extremite anale une substance dont le
contact avec la surface de I’eau produit une reaction qui les
chasse tres vivement en avant. L’animal peut en incurvant
son abdomen orienter sa course vers le point qu’il veut
atteindre. Nous avons cherch4 Texplication de cette
locomotion speciale : elle reside pour nous dans les vari¬
ations brusques de tension superficielle que provoque sur
I’eau la substance expulsee par ranimal.
“Lorsqu’on place I’insecte k la surface d’une nappe d’eau
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) C C
380
Mr. Norman H. Joy on the
pure, sur laquelle on a projete de la poudre de Lycopode,
on voit les spores fuir en arriere de I’aniinal et celui-ci
laisser un large sillage, lorsqu’il vent fuir rapidement :
aucune reaction ne se produit au contraire, lorsque I’insecte
inarche simplement sur la surface. Si Ton d^tache
I’extr^mit^ de I’abdoinen, on voit au bout de quelques
instants celle-ci expulser par intermittence la sub.stance 4
tension superficielle tres basse, et se mouvoir rapidement
comme un morceau de camphre ou de thymol.”
* * * *
I have confirmed most of these observations with D.
coerulescens and several species of the genus Stenus. My
studies soon inevitably led me to the general study of the
behaviour of Coleoptera during flood-time, as it must be
chiefly at such a time that these powers of locomotion
are brought into play. I have from time to time written
notes on my observations which I hoped to publish when
I had made a more complete study of the matter. How¬
ever, from want of time, little progress has been made
during the last two years, so I think it best to publish
them in the hope that they may stimulate others to con¬
tinue what is undoubtedly a very interesting subject.
When flat land or a river valley is flooded by heavy rains
it is well known that a very large number of beetles and
other insects is to be found in the “ flood rubbish ” washed
or blown up at the sides of the water. It is also notorious
how very local are some of the beetles inhabiting the
banks of streams, and one wonders how it is these species
are not washed away at every flood, or at any rate
scattered all down the valley. I believe that as a fact
most beetles soon find their way to laud by various methods,
and comparatively few, and those mostly of the larger
species, get washed down the middle of the stream.
This can be proved by examining flood refuse caught
on bridges, etc., in mid stream, and that blown up at the
sides of the river. In the former large beetles predomin¬
ate, very few small ones being found, whereas the latter
generally teems with small insects. In some places the
smaller beetles might be washed out of the mid-stream
rubbish by the force of the current, but I have noticed
that in the rubbish which collects over the arch of a bridge,
when the whole arch is submerged, large beetles are found
in greater numbers. It is chiefly due to the wind that
the fine rubbish and small beetles so soon find the edge
Behaviour of Coleoptera in time of Floods. 381
of the flood. It is worth noting here how very soon many
of the beetles leave this rubbish when it touches the ground
on the subsidence of the flood. I make a rule now, when
collecting flood refuse, of always taking it at the height
of the flood, while it is floating on the water. From
some rubbish fished out of the Kennet in 1908 I took
about thirty specimens of Ilyobates forticornis. On col¬
lecting about the same quantity two days after, when the
water had subsided and left it resting on the ground, only
five or six more specimens were obtained. No doubt there
is a large death-rate at every flood, some beetles soon
succumbing to their wetting. It is difficult to account for
the absence of certain species one would expect to find
in the rubbish, particularly Anehomenus albipes and Oodes
helopioides. The former is a very abundant riverside
species, yet it quite seldom turns up in flood refuse.
The latter I have never taken in this way, although it is
a common species in the Kennet valley, and Commander
Walker remarks on this (Ent. Mo, Mag., Vol. xliv, p. 135)
with regard to the Thames.
But apart from the help of the wind many waterside
beetles have other methods of reaching the shore quickly,
viz. by swimming in various ways, walking on the surface
of the water, and by this curious method described by
MM, G. Billard and G. Bruyant, which, for want of abetter
term, I will call “skimming.”
Now let us return to Dianous coeruleseens. It is in this
species that this method of progression is perhaps most
easily watched, as it so readily performs it. When a piece
of wet moss from near a waterfall is entirely immersed in
water this beetle may sometimes be seen, covered with a
glittering silvery layer of air, walking quite easily on the
moss. At other times the beetle crawls to the outside of
the moss and floats to the surface of the water, back upper¬
most, doubled up in the form of an inverted n> with the
legs folded in front of the abdomen. The elytra first reach
the surface and are soon free of the water. The thorax
and head then clear the water and dry ; the body is then
straightened, with the apex pointing slightly upwards, the
base of the abdomen sometimes taking some time to dry.
Now the whole of the upper surface is above water and
dry, the anal end being just at the surface, and the whole
of the imderparts and legs being under water. The beetle
then generally starts off “skimming” at a most remark-
c c 2
382
Mr, Norman II. Joy on the
able speed, sometimes almost straight, sometimes in a
series of curves. During the skimming the front legs are
held either doubled up, or spread out, as in the position
of a well “set” insect. The middle and posterior legs are
held close to the sides. When the beetle gets to shore it
dries itself carefully, curling its abdomen upwards, and
using its middle and back legs. I have seen it frequently
when on a leaf or some other floating raft enter the water
and deliberately swim to the bank. At times, especially if a
beetle has been kept for some time in a wet tube, it appears
to become waterlogged and remains at the bottom of the
water with its legs doubled up, and probably drowns. I
have confirmed the experiments of the above observers by
cutting off the end of the abdomen. A careful study of the
glands and the fluid they secrete would be of great interest.
I have little doubt the glands are analogous to those of
Bemhidium and Myrmedonia.
I have experimented with a fair number of other
species of Coleoptera, especially of the genus Stemts. It
seems probable that most of the latter are able to “skim,”
but some do it far more readily than others. It is import¬
ant to experiment on these insects directly they have been
captured, and with as little handling as possible, as a
negative result would be of no value if they have been
“ bottled ” for some time. Of the species with which I
have experimented perhaps S. guttula is the most expert
“ skimmer ” ; it moves so rapidly on the surface of the
water that it is really difficult to follow with the eye.
S. hipundatus also skims very readily, especially on a sunny
day. By repeatedly and quickly throwing a specimen out
as soon as it had got to shore I computed that it had
skimmed quite twenty feet before I lost it, when, however,
I think it was almost exhausted. This species when washed
out by throwing water on to the bank at the side of a pond
took deliberately to the water and “skimmed” to another
part.
Of other members of the genus Stemis I have experi¬
mented with the following : —
S. ossmm, S. Jlavipes, S. picipes and S. tarscdis skimmed
more or less strongly on several occasions; the last very
easily freed itself of the water and floated on the surface.
These species did not generally start skimming at once,
and often required some slight stimulation. I failed to get
S. juno to skim, but when the last segment of the ab-
Behavimr of Coleo'jptera in time of Floods. 388
domen was cut off this moved very rapidly, as was also the
case with S. hrunnipes. I did not induce any of the follow¬
ing to skim : S. dedaratus, S. lustrator, S. hiiphthalinus,
S. himacidatus, S. providus and S. opticus. I was unable to
experiment with enough specimens of some of these species
(only one in the case of S. ojdicus) to definitely conclude
that they cannot skim, except in the case of S. biqjhthahnus.
On many occasions I tested this species under the most
natural conditions and in bright sunshine with negative
results. Among other genera I found one species which
possesses this special power of locomotion, viz. Anch-
omenus alhipes. When thrown on the water this species
swims very strongly with the fore and middle legs, using
the back ones very slightly or not at all. By this method
it progresses at a good speed, but if carefully watched it will
be occasionally seen to give a distinct but short spurt
forwards, which however is sometimes rather more pro¬
longed. The legs are kept moving all the time, but the
spurt forward is undoubtedly due to skimming, and when
the apex of the abdomen is cut off it is found to skim
for a short time. The beetle does not seem able to skim
for long at a time, and those kept in a bottle for a short
time refused to do so at all. There are many other closely
allied forms and waterside species which might prove to
possess the power of skimming if they were tested.
A great number of beetles gain land when flooded out
by swimming, and some land beetles are very strong
swimmers. Here again there is much to be learnt as
to the exact method of swimming each species adopts.
The members of the genus Bemhidium upon which I have
experimented {B. assimile, B. articulatum, B. nitidulum)
swim very well. When B. assimile was flooded out by
gradually immersing a piece of turf it took to the
water quite naturally, and swam at a wonderful speed,
moving the legs very rapidly as in running, the front legs
which do nearly all the work being well spread out, the
back legs being sometimes held still and crossed behind.
Badister bipushdatus also swims well, using all its legs,
the back ones often very strongly.
Most of the Steni are good swimmers, even those that
also skim; D. coerulescens, S. guttula and S. bipunctatus
progress almost entirely by skimming, however. S. bimacu-
latus moves quite rapidly by swimming and uses its front
legs well spread out in a horizontal plane, as in Bembidium.
384
Mr. Norman
H. Joy on the
It also wriggles its body, which no doubt helps it along
to some extent. S. 'providus swims much like the last.
*8. juno, S. tarsrd.is, and 8. lustrator swim well, but 8.
declarahis, 8. fiamptcs and 8. huphthcdmus were more sluggish
in their actions. Chilopora longitarsis swims very readily,
using its back legs strongly.
These are, of course, all more or less waterside species,
and one would expect them to be good swimmers. Some
waterside species, however, are almost helpless when on
the surface of the water, and this is specially the case
with the 8ta'phylinidac. Quedius manrorufm hardly pro¬
gresses at all by ineffectual wriggling of its body, yet it
is commonly found in company with D. coerulescens in
moss at the sides of waterfalls. Evaesthetus is quite help¬
less, but all very small species are much affected by
capillary attraction, being drawn to small pieces of stick,
etc., and are quite unable to get a start off. Dyschirins
globosus has great difficulty in overcoming this attraction,
and then only swims feebly, using all the legs alike as
in walking. Tachypori move their legs, which are well
spread out, rapidly, but make quite slow progress.
Trogophloei and Homcdotae also swim, but are much
affected by capillary attraction. Oxypoda vittata only in¬
effectually wriggles its body.
Quedius vexans, which often gets washed out of moles’
nests in low-lying districts and is not uncommonly found
in flood rubbish, struggles violently with a great deal of
the body, and gets along at a fair rate, chiefly by using
the forelegs very rapidly ; the middle legs are not moved
so fast, and the back legs only occasionally. The front
legs are moved in a vertical plane, as is the case with a
dog, only they are never lifted out of the water. Q.
fvliginosus, Hctcrothops nigra and 8tilicus ajfinis behave
like Q. vexans.
Besides swimming there is yet another method of pro¬
gress, viz. — walking on the water. 8tenus juno on several
occasions deliberately cleared itself of the water, and
walked with its tarsi on the surface of that element.
It first raised the front part of its body on the anterior
and middle tarsi ; the abdomen being afterwards cleared
with the help of the back legs. 8. opticus very easily cleared
itself of the water and walked on the surface with the
abdomen high in the air. Occasionally it allowed the
abdomen to touch the water, but then always stopped for
Behaviour of Goleoptera in time of Floods. 385
a moment to dry it. When Gnypeta labilis was flooded off
a piece of grass, it remained standing on the water with
the abdomen curled up over the body. In this attitude it
was blown along at a surprising speed by a very slight
breeze. It also took flight directly from the water.
As might be expected a great many species of Coleoptera,
which must only very accidentally find themselves in
water, are quite helpless when they get there; among
them I may mention, Bhizohius litura, Hypera nigriros-
tris, Apion, Oxytelus, etc. Drusilla canaliculata lay
motionless for some time, then after some trouble cleared
itself of the water enough to swim, keeping the antennae
raised and constantly moving. Myrmedonia collaris be¬
haved in much the same way, floating to the top in the
form of an S with the underside of the abdomen upper¬
most, and remaining in this position for some minutes.
Bradfield, Berks, February 5, 1910.
( 386 )
XIl. On the Position of the Rhopalosomidae, with the
Description of a Second Species. By Claude
Morley, F.Z.S., F.E.S.
[Read June 1st, 1910.]
Plate L.
Among the various boxes of Ichncnmonidae received by me
in connection with the “ Fauna of India,” one from the
Indian Museum at Calcutta and another from Colombo
contained the sexes of an insect I entirely failed to place.
Mr. Rowland Turner has, however, been so good as to point
out to me its very close relationship with the anomalous
genus lihopaloso7na, Cresson (Proc. Ent. Soc. Philad., iv,
1865, p. 58, pi. v\ = Sibpllma, Westw., Trans. Ent. Soc.
1868, p. 329). Though probably generically distinct there¬
from, it is so extremely unlike any other known Hymen-
opteron that I am strongly averse to erecting a temporarily
unnecessary genus for its reception ; and I shall, conse¬
quently, describe it as
Rhopalosoma abnormis, sp. n. (Plate L, fig. 15.)
An entirely testaceous species with only the flagellum and eyes
black, the mandibles and anus infuscate. Head short but not abruptly
constricted behind the internally subacutely einarginate eyes ; occiput
abruptly declived, bordered below and not emarginate ; vertex neither
broad nor punctate, laterally impressed behind the outer ocelli, which
are prominent and enclose a triangular black mark, with the apical
ocellus below the level of the upper orbits ; frons convex and very
obsoletely punctate, finely carinate longitudinally in the centre ; face
extremely short, strongly transverse and obsoletely transaciculate ;
clypeus obsoletely discreted, large, transverse and truncate at both
extremities ; labrum slightly exserted and broadly rounded, not
trilobed, apically ; mandibles strong and subtridentate with a large
black apical tooth, surmounted by one half its length and a third
minute excrescence ; palpi flavous and elongate. Antennae inserted
at centre of eyes and hardly longer than half body, black and
strongly attenuate throughout with the apically truncate and
biarticulate scape alone pale; flagellum eleven-jointed in ten-
jointed in 5 , with the joints pilose, elongate, cylindrical, and the
basal seven or eight each internally emitting two distinct setae from
its apex ; terminal joint coriaceous, thrice longer than broad and
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
Mr. Claude Morley on the Ehopalosomidae. 387
apically acuminate. Thorax subcylindrical, obsoletely pilose and
hardly punctate throughout ; prothorax normal and extending to
base of tegulae ; mesonotum gradually declived anteriorly, finely
sulcate above tegulae but with no notauli ; metathorax evenly declived
with no trace of carinae, its spiracles oblique, impressed and sub-
linear : mesosternum very short, convex throughout, pilose and not
laterally sulcate. Scutellum immarginate, subdeplanate, triangular
and apically obtuse, obsoletely punctate and pilose with its basal
fovea simple. Abdomen subnitidulous, shagreened and shortly pilose ;
basal segment subsessile, basally half the length of its apical
breadth, which is hardly half (or in $ a third) of its total length,
convex and with no sculpture, gradually explanate to near its
constricted apex with distinct spiracles at its basal third and a
determinate discal fovea before its apex ; second segment as long as
the first and about half as broad again, gradually explanate basally
and not strongly constricted at its apex, with its base both above and
below longitudinally niultistrigose and thyridii linear; third and
fourth segments together about as long as second and extremely
closely connected though not connate, the fourth becoming narrower
apically with its dorsal margin subemarginate ; fifth as long as
fourth ; sixth and in ^ seventh much shorter and deflexed ; venter
convex throughout and not plicate, with four well-developed segments
and two very short ones preceding the ( ^ ) stout and black genital
valvulae, which extend only to the anus, or ( $ ) the elongate
hypopygium, which extends to anus and covers base of the minute
terebra. Legs very slemler but not elongate, hind ones of ^ hardly
extending beyond the abdomen ; anterior trochanters appear uni-
(though probably no more than telescoped), the hind ones bi-articulate ;
front tibiae hardly longer than the metatarsi, and sparsely spinose
internally, each with a single elongate and basally sinuate calcar ;
tarsi long and in slender with the claws small and pectinate,
basally not lobed though shortly pulvillate in 5 > the four apical
joints strongly spatuliform and dilated with their apex setiferous on
either side and base constricted, claws simple and shorter than the
bilobed pulvilli ; posterior femora apically subspinate internally with
their tibiae very elongately bicalcarate ; front and hind metatarsi
basally sinuate, the latter with the inner calcar basally strongly
pectinate in Wings neither broad nor ample, hyaline with the
stigma narrow, inconspicuous and centrally subconstricted ; all the
nervures testaceous ; upper wing with nervures disposed as in
Ichneunionidae excepting the extreme pellucescence of the second
recurrent which is, however, distinctly traceable and the aberrantly
bifurcate first intercubital or submarginal nervure ; upper basal
nervure emitted from the median much further from base than the
3SH
Mi^'laude Morley o??. the
lower ; lower basal cell with an elongate, free anti simple nervure.
Lower wing with the neuration of Ichneuraonidae excepting an
elongate and basally pellucid additional nervure, between the median
and radial; nervellus straight, opposite and not intercepted, with
the wing margin cleft to near its junction with the posterior nervure
and another very distinct incision at the humeral nervure, rendering
the anal angle entirely lobate. Length, 10 mm. ^ $ .
A single example of each sex is all I have seen of this
remarkable insect. The ^ (type) is in my collection, and
was most kindly given me by Mr. O. S. Wickwar, of
Colombo, who took it at Mankulam, in Northern Ceylon,
during November 1908. The $ is in the Calcutta Museum,
under Dr. Annandale’s care ; it was captured at that city
on August 9, 1907.
My insect differs to an appreciable extent from the only
previously described species of theRhopalosomidae R. poeyi,
which was brought forward by Cresson (loc. cit. p. 59) in
his paper “ On the Hymenoptera of Cuba,” and three years
later by Westwood under the synonymous name 8. aenig-
matica in his paper {Joe. cit.) “Descriptions of New Genera
and Species of Exotic Hymenoptera.” Therefrom the male
of B. dbnen'mis differs in its posterior tarsi which bear no
combing apparatus, and in having the calcaria equally long
in both ; the hypopygium is retracted, and does not entirely
conceal the base of the non-exserted valvulae, as is the case
in the female R. poeyi, which also has the terebra slightly
exserted, the spicula reflexed and exceeding the valvulae
in length ; the basal segment is less elongate in B. ahnormis
with the second not abruptly constricted and arcuate,
though similarly (as in Tiphia) trans-strigose, basally ;
the discal segments are equally emarginate, and the
conformation of the metathoracic spiracles is alike in
both, but I fail to discover any trace of distinct petiolar
spiracles in R. poeyi.
These points of divergence are drawn from the three
female specimens of Cresson’s species in the British
Museum, two of which were captured in St. Domingo by
Mr. Tweedie (not Tweedy, as given by Westwood) and
acquired in 1855 ; and the third, from Jamaica, ex coll.
Gosse, was obtained in 1847 ; it is also known to occur in
Hayti. The localities of the new species are consequently
of great interest and show it to be widely distributed in the
Oriental region.
#
Position of the Rhopalosoviidac. 389
The affinities of the present genus are so uncertain that
it is well to regard it as a distinct family. To prove this it
is only necessary to remark that Nylander thought it an
ant, Haliday a Fossor, Westwood a social wasp, Smith
an Ophionid Ichneumon, and Cresson a Braconid ! The
last places it unhesitatingly in “ Braconides,” but “ Divi¬
sion - ?” Yet in his very rough figure he dots the
second recurrent nervure as being pellucid and at least
traceable ; he thought it “ a connecting link between the
Ichnmmones genuini and the aclsciti ; from the former it
differs by the paucity of the antennal joints, and from the
latter by the anterior wings having a faint indication of
a second recurrent nervure. Its structure places it, beyond
doubt, in the family Ichneunionidae ” {sensu lato), “ while
its general appearance, together with the arrangement of
the wing veins, seems to place it among the Adsciti, where
I will allow it to remain for the present.” Fred Smith
“On the Affinities of the Genus Sibyllina, of Westwood”
(Proc. Ent. Soc. 1868, p. 1), is chiefly concerned in showing —
as he conclusively does — that this insect cannot appertain
to the Vespidae, as placed by Westwood, who however was
so uncertain as to add quoad afinitates animum excrucians” !
But the position assigned it by Smith, with some assurance,
near the IcJmeumonidous genus Anomalon is equally
untenable in these days of fuller knowledge of that family,
especially since the contrary sexes possess simple tarsi
{cf. also Meeting of Ent. Soc., November 16, 1868).
Many pertinent characters of its relationship are set
forth by Westwood in his Thesaur. Ent. Oxon. (1874,
pp. 130-31), and the male is beautifully figured at lib. cit.
p. xxiv, fig. 9 ; but, still inclining to place it in the
Aculeata, he is unable to suggest any natural position for
the genus. Dalla Torre, so far from assisting us in the
matter, places Bhopalosoma among the genera sedis incertae
at the tail of the Braconidae (Cat. Hym. iv, 1898, 307)
and Sibyllina — incorrectly rendering Westwood’s name
Sybillina — almost at the beginning of the Vespidae {1. e. ix,
1894, 113). The synonymy has never been questioned,
and is, I consider, sufficiently apparent.
For my own part, I am entirely satisfied that the
Bhopalosomidae can be placed nowhere among the Para¬
sitica or Terebrant Hymenoptera, a favourite dumping
ground for aberrant genera in days when they were less
studied than is now the case. Fred Smith appears equally
390 Mr. Claude Corley on the HhoiKilosoniidae.
satisfied that it has no place among the Aculeates ; but
Haliday wrote, “ I consider it a Sphegid, with smooth
legs, Pelopaens” and he was rarely incorrect : in 1900,
Ashmead (Canadian Entom., xxxii, p. 148 ; cf. also Proc.
Ent. Soc, Wash., iii, p. 303) erected a family for the sole
reception of this genus of a single species and placed it
between the Cosilidae axxd Thynnidae, with neither of which
it is at all closely connected. Sharp (Camb. Nat. Hist., ii,
101) “can expre.ss no opinion as to whether it is allied
to the Scoliidae or to the Bphegidae * ” ; he takes it for
granted that it should stand near one or the other, and in
that case the lobation of the hindwings renders it closer to
the former.
The facts of the antennae being thirteen-jointed in ^
and twelve-jointed in and of the dorsal abdominal seg¬
ments being seven in $ and six in certainly prove it
to be Aculeate ; though the neuration is that of the
ICHNEUMONIDAE (probably most closely resembling that
of Forster’s genus Barylypa among the Anomalides), with
a few additional nervures.
Granting it an Aculeate, related to the Scoliidae, we can
do little more than say with Fred Smith, “Place Sibyllina
in any group of the Hymenoptera, and it will, as it were,
stand alone ; it has little affinity that I can discover,
certainly it has no strong affinity, with any other known
insect.”
Have we here the “ ancestral type ” of Hymenopteron,
emitting Fossores according to its body and Parasites
according to its wings ? Or more probably this anomalous
combination is brought about by some process (though
hardly cross-breeding of Pelopaeus and Aiiomalon]) of
very remarkable specialisation.
* I am afraid the name Splierddae is too well established to ever
be universally corrected now-a-days. In May 1900, the late Eev.
T. A. Marshall wrote to me, “ I enter a protest against the spelling
of the Ichneumon genus Sphegophaija. 2<?)7j|, genitive
“ wasp ” in Greek, should be rendered in our nomenclature Sphex,
isphecis, etc., as the older authors well knew. The mistake of sphegis,
Sphegidae, is a new corruption, which, if sanctioned, will be forth¬
with adopted by everybody. I fear it is almost too late now. Such
blunders are trifling, of course, but I know no reason why a mistake
of spelling should be allowed in Greek and Latin, when it would be
hooted in English.”
Monk Soham House,
Suffolk,
March 1910.
( 391 )
XIII. Notes on the Scoliidae. By Rowland E. Turner,
F.Z.S.
[Read June 1st, 1910.]
Plate L.
The new species which furnish the material for this
paper are mostly in the collection of the Berlin Museum,
though a few are from other sources. The material shows
that the genus Anthohosca ranges all up the east of Africa
into Arabia, and also adds a new genus to the small family
Sapygidae, the type of the genus having been placed in
the British Museum by the late Colonel Bingham. Ash-
mead has divided the Scoliidae into several families, which
I retain as sub-families, but some of the genera he includes
must be placed elsewhere.
Family SCOLIIDAE.
Sub-family MYZININAE.
Myzine nigrita, sp. nov. (Plate L, fig. 6.)
^ . Clypeus produced and rather broadly truncate at the apex,
coarsely punctured. Head rugose, broader than the pronotuni, the
posterior ocelli more than half as far again from the eyes as from each
other ; the antennal tubercles large, the scape short, no longer than
the second joint of the flagellum, antennae short, scarcely longer than
the thorax and median segment combined, very stout, the second
joint of the flagellum more than half as long again as the first, a
little shorter than the third, joints 5-10 strongly arcuate beneath.
Head and thorax clothed rather sparsely with long cinereous grey
pubescence, the abdomen very sparsely clothed with short greyish -
white pubescence. Thoi’ax closely punctured, the pronotum much
shorter than the mesonotum, the median segment rounded, not trun¬
cate. Abdomen closely and finely punctured, the petiole about equal
in length to the third joint of the posterior tarsi, the first segment,
excluding the petiole, a little longer than broad, not constricted
at the apex, the whole abdomen of an elongate fusiform shape ;
seventh dorsal segment with a longitudinal depression on each side
on the apical half, the apex narrowly and not deeply emarginate, the
depth of the emargination scarcely as great as the width at the apex.
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
392 Mr. Rowland I^Turuer’s Notes on the Scoliidae.
Aculeus moderately recurved, short, the apex only reaching the
emargination of the dorsal segment. Second abscissa of the radius a
little longer than the third, first recurrent nervure received just be¬
yond the middle of the second cubital cell, second at one-fifth from
the base of the third cubital cell.
Black ; the calcaria whitish. Wings hyaline, nervures black.
Length 15 mm.
Hah. Cape Colony.
Type in Berlin Museum.
The arcuate joints of the antennae, the short aculeus
and the more flattened form of the first abdominal segment
distinguish this species at once.
Myzine continua, Cam.
Flesia contimta, Cam., Rec. Albany Mus., i, 5, p. 299, 1905.
I think Cameron has very much underestimated the
variability in the males of this group, and that his continua
and interruyta are probably the same species. I think
they probably belong to Myzine rather than to Flesia, but
Cameron does not mention the shape of the first abdominal
segment. The deep slit in the apical dorsal segment of
the abdomen which is mentioned in the description is
alway present in Myzine, but is often absent in Flesia. If
it is considered impossible to distinguish between the
males of the two genera, it seems there is no excuse for de¬
scribing them in Flesia, the name Myzine having priority,
and its use for the males being recommended by Saussure.
Some of the males described by Cameron from Baluchistan
as Flesia have proved beyond doubt to belong to Myzine.
The descriptions of continua and of interrupta apply fairly
well to a species which has a wide range in South Africa,
and shows considerable variation both in colour and the
smaller details of neuration. I think it probable that it
is the male of Myzine rufifrons, Fabr. ; the range of both
extends from the Zambesi to Cape Colony. In the males
of this genus, and in both sexes of Tiphia, there is a fine
field for the species monger, which fortunately has not
yet been exploited.
Myzine rujinodis, sp. nov. (Plate L, fig. 5.)
^ . Clypeus short and broad, the apical margin almost transverse,
shallowly emarginate in the middle, strongly punctured at the base,
almost smooth at the apex. Head, thorax, and median segment
Mr. Rowland E. Turner’s Notes on the Scoliidae. 393
closely and rather coarsely punctured, the middle of the scutelluni
narrowly smooth and shining. Antennae very stout, the first joint
of the flagellum half as long as the second, the third distinctly
longer than the second, the scape short and coarsely punctured ; the
front between the antennae densely clothed with long cinereous-grey
pubescence, which is also sparsely distributed over the whole of the
head and thorax. Pronotum much shorter than the mesonotum, very
slightly narrowed anteriorly, the anterior margin almost transverse.
Abdomen rather slender, shining, finely and shallowly punctured
with a short petiole which is about equal in length to the fourth
joint of the posterior tarsi, the first segment beyond the petiole
swollen and globular, broader than long. Apical segment more
sparsely punctured, the emargination for the aculeus rather shallow,
broader at the apex than deep, the surface of the segment, before the
emargination, not flattened or depressed. Aculeus long and slender,
strongly recurved. Basal joint of the posterior tarsi about three-
quarters of the length of the tibia. Second abscissa of the radius more
than half as long again as the third, first recurrent nervure received
at two-thirds from the base of the second cubital cell, second before
one-sixth from the base of the third cubital cell.
Black ; mandibles, anterior margin of the clypeus, margins of the
pronotum, tegulae, a narrow transverse band at the apex of the first
dorsal segment of the abdomen, a transverse spot on the middle of
the apical margin of segments 2-6, with a smaller transverse spot on
each side, and similar, but much smaller marks on ventral segments
2-5 pale yellow ; first dorsal segment, except the petiole and the
yellow apical band, ferruginous red ; legs dull ferruginous. Length
15 mm.
Hah. Cape Colony {DNge).
Type in Berlin Museum.
I think that I am right in placing this species in Myzine
rather than in Plesia, the males of the two genera are
however not always to be distinguished with certainty.
The group of South African species characterised by an
elongate basal segment of the abdomen and the absence of
an emargination of the apical dorsal segment are certainly
males of Plesia. I suspect however that some species show¬
ing neither of these characters, but of a rather slender form,
and with a long pronotum narrowed anteriorly, may also
prove to have females of the Plesia type of neuration.
The present species is easily distinguished by the red
colour of the first dorsal segment, but unless there is some
marked point of distinction it is better to leave the males
304 Mr. Rowland E. Turner’s Notes on the Seoliidae
of this group undescribed until larger collections of both
sexes are available and local variations better understood.
riesia {Mesa) saussurei, sp. nov.
$ . Clypeus very short, punctured, the apex depressed and smooth.
Head coarsely punctured, more closely on the front than on the
vertex ; the scape shining and finely punctured ; flagellum opaque,
the three basal joints shining, the first joint short, almost concealed
in the apex of the scape, the third longer and thicker than the
second. Pronotum rather shorter than the mesonotum and scutellum
combined, the sides almost parallel, coarsely punctured-reticulate ;
mesonotum and scutellum coarsely but more sparsely punctured,
propleurae indistinctly striated, mesopleurae coarsely punctured,
metapleurae striated. Median segment as long as the pronotum,
deeply but irregularly punctured, nearly twice as broad as long,
distinctly margined, with two low, median, longitudinal carinae,
the posterior truncation steep, the surface finely rugulose. Abdo¬
men stout, about equal in length to the head, thorax and median
segment combined, rather closely punctured ; petiolate, the petiole
linear, equal in length to rather more than half of the dorsal surface
of the first segment, which is nearly as long as the second; sixth
dorsal segment longitudinally striated, broadly rounded at the apex.
Tarsal ungues bifid. Second abscissa of the radius scarcely half as
long as the third, and less than two-thirds of the length of the first,
second recurrent nervure received beyond the middle of the third
cubital cell, the third transverse cubital nervure oblique, the radial
cell rounded at the apex, scarcely separated from the costa.
Black ; the mandibles, clypeus, scape, three basal joints of the
flagellum, tarsi and tlie ajDex of the pygidium fusco-ferruginous.
Wings fuscous. Length 10 mm.
Hah. Madagascar, Antananarivo. Two specimens.
Type in the Zoological Museum at Berlin.
Sub-family TIPHIINAE.
Tijihia conradti, sp. nov.
$. Clypeus short, almost transverse at the apex. Front round the
base of the antennae tranversely depressed, slightly concave and very
finely rugulose ; the remainder of the head shining and sparsely
punctured, smooth round the ocelli. Posterior ocelli very near to¬
gether, nearly three times as far from the eyes as from each other.
Scape closely punctured, with sparse greyish pubescence, the apex
Mr. Rowland E. Turner’s Notes on the Scoliidae. 395
smooth and shining. Pronotvun sparsely punctured, the posterior
margin broadly smooth and shining ; mesonotum and scutellum
shining, with a few very large punctures, propleurae almost smooth,
mesopleurae rather sparsely punctured, the sides of the median seg¬
ment finely obliquely striated. Median segment almost as long as
the pronotum, mesonotum and scutellum combined, subopaque,
almost smooth, with the three usual carinae in the middle, the median
one not reaching the ape-x, the two lateral ones half as far again from
each other at the base as at the apex, the margins of the segment
distinctly raised both at the apex and the sides, a fine longitudinal
Carina on each side narrowly separated from the raised lateral mar¬
gin. Abdomen shining, finely punctured, the two basal segments
sparsely, the others more finely and closely, the apical half of the
pygidium smooth ; the basal segment broad, without a transverse
Carina at the base. Stigma very small, almost absent ; first recurrent
nervure received at the middle of the first cubital cell, second at
three-quarters from the base of the second cubital cell ; second
transverse cubital nervure strongly curved inwardly.
Black ; mandibles, antennae and anterior tibiae beneath fusco-
ferruginous, the apical half of the pygidium pale testaceous ; spines
of the tibiae and tarsi pale ferruginous ; the pubescence on the
abdomen and legs whitish. Wings hyaline, tinged with fuscous,
nervures black. Length TO mm.
Hah. N. Cameroon {Conradt).
Type in Berlin Museum.
The median segment is unusually long in this species.
Tiphia ahriqjta, Turn.
Tiphia ahrupta, Turn., Ann. and Mag. Nat. Hist. (8), ii, p.
127, 1908, ?.
The type is from Salisbury, Mashonaland. A specimen
in the Berlin Museum from Langenburg, Lake Nyassa,
has the median segment distinctly longer and the pygidium
rather broader. The species of TipMa are subject to local
variation, the length of the median segment seeming to be
especially influenced. If new species are founded on such
characters before long series are available and both sexes
known, much confusion may result. I therefore prefer to
treat the Nyassa form merely as a local race.
There are at least five species of Tipthia in the collection
from Langenburg, which may be distinguished by the
following key to the females : —
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) D D
896 Mr. Rowland TT Turner’s Notes on the Scoliidae.
A. First dor.sal segment of the abdomen
with a transverse carina at the base.
a. Head and pronotum punctured. . T. abrnpta, Turn., var.
b. Head and pronotum rugose . . . T. scabrosa, Gerst.
B. First dorsal segment without a carina
at the base.
a. Opaque, wings subhyaline, antennae
black or fuscous . T. monumatajM, Turn.
b. Shining, antennae orange or ferru¬
ginous.
Antennae orange wings fusco-
violaceous, legs black, stigma
very small . T. fulvicornis, Turn.
b~ Antennae and legs ferruginous,
wings subhyaline, stigma
large . T. pedestris, Gerst.
Tiphia nataleusis, Siii.
Tipliia natalensis, Sm., Descr. n. sp. Hymen., p. 184,
1879, ?.
In the type specimen tlie second recurrent nervure is
received only just beyond the middle of the second cubital
cell, and the carina at the base of the first dorsal abdo¬
minal segment is very indistinct. There is a specimen
from Cape Colony in the Berlin Museum which I refer
with doubt to this species in which the second recurrent
nervure is received much nearer to the apex of the cell,
the carina of the first segment is more distinct, and the
punctures are rather coarser, the insect is also much
larger. But it is not advisable to found a new species
in this diflficult genus on a single specimen in which the
differences are so slight, especially as a specimen in
the British Museum from Howick, Natal, is somewhat
intermediate.
Large collections of the species of Tiphia with full data
are much needed. My impression, derived from a season’s
collecting in Assam, is that seasonal forms may occur as
in Lepidoptera.
Tiphia siinlaensis, Cam., was plentiful in April and May,
but none were taken later, whereas Tiphia himalayensis,
Cam., was taken in August and September, but not in the
spring months. T. himalayensis is twice as large as sim-
laensis, the median segment is a little longer, the carina on
Mr. Rowland E. Turner’s Notes on the Scoliidae. 397
the median segment more distinct, the median one ex¬
tending to the apex, the head and thorax more coarsely
punctured and the stigma on the forewing a little smaller.
The male of himalayensis (= ccinaliculata, Cam.) has the
radial cell shorter than in swilaensis, the sculpture of the
median segment much coarser, the wings darker and the
clypeus without the feeble emargination on the anterior
margin. But Bingham took himalayensis in Sikkim in
April at 4000 ft. elevation. My Assam specimens were
all taken at an elevation of from 5000 to 6000 ft., near
Shillong.
My suggestion of seasonal dimorphism in this genus is
merely tentative, and is put forward in the hope that it
may lead to further observation on the subject, and to more
importance being attached to accurate dates of capture by
Hymenopterists.
Sub-family COSILINAE.
Anthohosca arahica, sp. nov. (Plate L, fig. 3.)
5 Clypeus almost triangular, triincate at the apex, smooth and
shining. Mandibles stout, without teeth, rather blunt at the apex.
Head scarcely broader than the thorax, shining, rather sparsely
punctured and thinly clothed with long cinereous haiis, the punc¬
tures closer on the front than on the vertex. Ocelli in a broad tri¬
angle, the posterior ocelli not quite as far from each other as from
the eyes. Antennae short and stout, the scape shining and smooth,
the second joint of the flagellum shorter than the third. Thorax
and abdomen shining, sparsely punctured ; median segment very
minutely and rather closely punctured; pronotum much longer than
the mesonotum, very slightly narrowed anteriorly, scutellum very
broadly rounded at the apex ; propleurae shining, very sparsely
punctured, mesopleurae more closely punctured, metapleurae smooth
and shining. Median segment shorter than the pronotum, gradually
sloped to the ape.x. First abdominal segment nearly as broad at the
apex as the second segment, obliquely sloped anteriorly, with a sharp
Carina at the base beneath, without a groove between the first and
second ventral segments ; apical dorsal segment punctured rugose,
with close pale fulvous pubescence at the base, more finely punc¬
tured and narrowly rounded at the apex. Posterior tibiae much
broadened, anterior tarsi with a comb, tarsal ungues bifid. Wings
not reaching the apex of the abdomen, radial cell rounded at the
apex, not reaching beyond the third cubital cell ; second abscissa of
the radius longer than the first, but a little shorter than the third,
D D 2
398 Mr. Rowland Tl. Turner’s Notes on the ScoHidae.
tlie tliird cubital cell longer on tlie radial than on the cubital
uervure, receiving the second recurrent nervure before one-third
from the base.
Black ; a small spot below the anterior ocellus, the inner orbits of
the eyes continued and uniting on the vertex, a line on the posterior
margin of the head behind the summit of the eyes, the posterior
margin of the pronotum and a spot at each of the anterior angles, a
spot on the mesopleurae, a curved band on the scutellum, a spot at
tlie base of the median segment and a larger one on each side at the
posterior angles and a narrow transverse band on each side at the
base of dorsal abdominal segments 1-4 pale yellow ; antennae and
tibiae fuscous, the anterior tibiae with a yellow line above ; man¬
dibles, tegulae, tarsi and pygidium testaceous brown. Wings
hyaline, nervures testaceous. Length 9 mm.
Ildh. Lahej, near Aden, May.
The genus has been recorded by Fox from Somaliland,
but not previously from Arabia. In colour this is very
unlike the Australian species of the genus, also in the
bifid ungues.
Anthohosca minima, sp. nov.
9 . Mandibles falcate, acute at the apex ; clypeus a little produced,
truncate at the apex. Head rounded, scarcely broader than the
thorax, convex, shining, finely and sparsely punctured ; antennae
inserted rather nearer to each other than to the eyes, the second joint
of the flagellum as long as the third. Eyes touching the base of the
mandibles, the posterior ocelli much further from the eyes than from
each other. Thorax sparsely and shallowly punctured, the propleurae
smooth and shining ; pronotum distinctly narrowed anteriorly,
median segment no longer than the scutellum, obliquely slojjed
posteriorly. Abdomen finely punctured, the apical dorsal segment
coarsely punctured and covered with stiff fulvous pubescence, the
first segment narrowed to the Itase. Second abscissa of the radius
more than twice as long as the first, but much shorter than the
third ; the third cubital cell much longer than high, receiving the
second recurrent nervure at one-third from the base. Eadial cell
very narrowly truncate at the apex. Posterior tibiae strongly
serrate.
Black ; mandibles, antennae and legs testaceous brown ; abdomen
dark reddish-brown ; clypeus, inner margin of the eyes, a spot be¬
hind the eyes near the summit, the posterior margin of the prono¬
tum, a spot on the scutellum, a spot on the middle of the median
segment, a narrow transverse band on each side of dorsal abdo-
Mr. Rowland E. Turner’s Notes on the Seoliidac 399
minal segments 2-4, the tegulaeand a line on the anterior tibiae pale
yellow. Length 5 mm.
Hab. Mombassa (Hildehranclt).
Type in Berlin Museum,
Very near arahiea, but differs in the more elongate
radial cell, in the greater length of the third cubital cell,
the smaller size and the lesser breadth of the tibiae.
Anthohosea Jiavojneta, sp. nov. (Plate L, fig. 4.)
^ . Clypeus sparsely punctured, much broader than long, subtrun¬
cate at the apex. Head and thorax minutely and closely punctured ;
the head covered sparsely with short, upright, cinereous pubescence.
Antennae stout, a little shorter than the thorax and median segment
combined, the third joint of the flagellum as long as the first and
second combined. Scutellum subtriangular, rather narrowly trun¬
cate at the apex ; the mesonotum with two shallow longitudinal
furrows on each side. Median segment as long as the mesonotum,
rounded, very finely rugulose. Abdomen finely shagreened, elon¬
gate, tapering to the extremities, the hypopygium small, rounded
and ciliated. Posterior tibiae rather strongly serrate, all the tarsal
ungues bifid. Third abscissa of the radius longer than the first and
second combined ; first recurrent nervure received a little beyond
the middle of the second cubital cell, second a little before the middle
of the third cubital cell. First transverse cubital nervure straight at
the base, sharply bent before the middle.
Black ; the clypeus, except a small, elongate, black spot on the
centre, the inner margin of the eyes narrowly, not reaching the
summit, a small spot behind the summit, a broad band on the pos¬
terior margin of the pronotum, the tegulae, a large spot on the
middle of the scutellum, another on the postscutellum, the apex of
the anterior femora and the anterior and intermediate tibiae and
tarsi above pale yellow ; tibiae and tarsi ferruginous brown. Wings
hyaline, nervures black. Length 11 mm.
Hab. Zanzibar.
Type iu the Zoological Museum at Berlin.
The posterior tibiae are more strongly serrate than in
most species of the genus, but this character is not
generic, there being much difference in the development
of the serration in the different species. Cameron’s
genus Odontothynnus comes near this species, but differs
in the bidentate clypeus and the simple ungues of the
posterior tarsi. I have not seen specimens, but Cameron
400 ]\Ir. Rowland E. Turner’s Notes on the Scoliidne.
was evidently not acquainted with Anthobosca at the time
of writing his description, and I doubt if the points of
distinction will prove to be of generic value.
Sub-family SCOLIINAE.
Scolia {Diliacos) violacca, Lepel.
Campsomeris violacea, Lepel., Hist. Nat. Insect. Hym., iii,
p. 502, 184.5,
Scolia insvlaris, Sm., Journ. Proc. Linn. Soc., Zool. iii,
p. 153, 1858,
Liacos mstilaris, Turn., Ann. Mag. Nat. Hist. (8), iv, p. 168,
1909.
Saussure places violacea, Lepel., in Diliacos. It is un¬
doubtedly the species I referred to insularis., Sm. My
identification is certainly correct if the species so named
in the British Museum by Smith is identical with the
type, which I have no reason to doubt. Smith, how¬
ever, does not put insularis in the Liacos group in his
description.
Scolia (Trielis) techowi, sp. nov.
9 . Clypeus raised, flattened in the middle and irregularly longi¬
tudinally striated, the anterior margin depressed and rounded, the
striated area as long as broad, rounded at the base. Head shining,
sparsely punctured, almost smooth round the ocelli, closely punc¬
tured between the antennae ; the posterior ocelli a little further
from the eyes than from each other. There is a very obscure, im¬
pressed, transverse line on the front half-way between the base of the
antennae and the anterior ocellus. Thorax rather sparsely, but
very deeply punctured, the scutellum very sparsely punctured,
pleurae finely punctured, the sides of the median segment shining
with a few very small and shallow punctures. Median segment and
fourth and fifth abdominal segments finely and closely punctured,
the three basal segments of the abdomen more sparsely punctured,
the second and third smooth in the middle. Sixth dorsal segment
rounded at the apex, with a blunt spine on each side, longitudinally
rugose-striate. Third cubital cell rather variable in shape, usually
a little more than half as long on the cubital as on the radial nervure,
the second transverse cubital nervure often incomplete, when com¬
plete almost interstitial with tlie inner margin of the radial cell; the
stigma large.
Black ; mandibles and flagellum beneath fusco-ferruginous ;
Mr. Rowland E. Turner’s Notes on the Scoliidae. 401
pygidium and tarsi ferruginous brown, the spines of the tarsi
whitish, one of the spines of the posterior tibiae long and spatuli-
form. Head and thorax sparsely clothed with grey pubescence,
dense and long between the antennae and on the posterior margin of
the head, the abdominal segments with an apical fascia of hairs
grey on the three basal segments, bright fulvous on the fourth and
fifth, the pygidium densely covered with stiff golden setae. Wings
hyaline, faintly clouded with fuscous at the apex, nervures
ferruginous. Length 17 mm.
Hah. S.W. Africa, Windhoek (Techoxo).
Type in Berlin Museum.
Very nearly allied to Elis 'punctnm, Sauss., of which it
may be a western form, but the clypeus is much less
coarsely striated, the front between the antennae less
prominent and the colour of the pubescence and colour
and sculpture of the apical dorsal segment are very dif¬
ferent. Trielis 'punchtm is in the Berlin Museum collec¬
tion from Lake Nyassa. The present species is in some
respects intermediate between punctxim and stigma ; the
latter species also occurs at Windhoek.
The probable male of iechoivi has the clypeus yellow with a
large triangular black mark in the middle, the pronotum, base of the
scutellum, po.stscutellum, tegulae, femora, tibiae, tarsi, a broad,
straight band occupying the apical two-thirds of dorsal segments
1-4, and a similar band narrowed on the sides of the fifth and sixth
segments yellow ; the apical half of the seventh dorsal segment
ferruginous. The yellow abdominal bands are continued rather
more narrowly on ventral segments 2-6, those on segments 2-3
narrowly emarginate anteriorly on each side. Wings hyaline, ner¬
vures pale ferruginous. Radial cell truncate at the apex, the third
cubital cell complete, more than twice as long on the radial as on the
cubital nervure. First abdominal segment nearly as broad at the
apex as long. Length 16 mm.
Scolia (Trielis) stigma, Sauss.
The probable male of this species differs from that of
techowi in the rather slender form of the first dorsal abdo¬
minal segment and in the shape of the yellow bands on
dorsal segments 2-4, which are more or less deeply emar¬
ginate in the middle anteriorly and contain a transverse
black spot on each side. The flagellum is often more or
less ferruginous. Length 14 mm.
O O
402 Mr. Rowlan^^^l Turner’s Notes on the Scoliidae.
Scolia {Diclis) qibinqucfasciata, Fabr.
Seoliei qidnquefasciata, Fabr., Spec. Insect., i, p. 453,
1781,
Seolia sirjnata, Sin., Cat. Hyni. B.M., iii, p. 105, 1855,
Hah. West Africa.
Fcolia mansueta, Gerst., Monatsber. Akad. Wiss. Berlin,
p. 512, 1857.
Hal). Mozambique.
These seem to me to be merely geographical races of the
same species. Dielis soleata, Gerst., which is common in
Mashonaland and the adjacent districts on the Zambesi,
is undoubtedly a race of quinquefasciata, differing only in
colour.
Soclia {Dielis) socotrana Kirby. (Plate L, fig. 7.)
Dielis socotrana, Kirby, Bull. Liverp. Mus., iii, p. 14, 1900.
5 • Clypeus rounded at the apex, slightly convex, smooth and
shining in the middle, with a short longitudinal sulcus at the apex.
Front closely punctured above the antennae, vertex smooth and
shining. Thorax coarsely punctured, the disc of the mesonotum
sparsely, the scutellum, postscutellum and base of the median
segment at the sides shining and almost smooth. Pleurae sparsely
punctured ; abdomen with a few scattered piliferous punctures, a
row of similar punctures before the apex of dorsal segments 1-4.
Apex of the radial cell broadly truncate, very slightly oblique.
Black ; the mandibles fusco-ferruginous, spines of the tibiae whitish
Wings hyaline, tinged with yellow, clouded with pale fuscous beyond
the radial cell ; nervures testaceous. Dense white pubescence on
the sides of the clypeus, on the sides of the first abdominal segment,
a band of the same on the apical margin of the second and third
dorsal and ventral segments and of the fourth ventral segment ;
dull sparse fulvous brown pubescence on the front, thorax and
median segment ; dense bright golden fulvous pubescence on the
posterior margin of the head and pronotum, and golden pubescence
on the apical margin of the fourth dorsal segment and on the two
apical segments. Tarsal ungues ferruginous. Length 16 mm.
Hob. Somaliland, Salable {von Erlanger). Berlin
Museum.
Scolia {Dielis) marshalli, sp. nov.
$ . Clypeus coarsely and sparsely punctured, the centre shining
and almost smooth. Front very coarsely and closely punctured
Mr. Rowland E. Turner’s Notes on the Scoliidae. 403
densely clothed with erect, long, black hairs ; the space round the
ocelli very sparsely punctured, vertex smooth and shining, an arched
depressed line above the posterior ocelli, the margins of the head
fringed with long black hairs ; scape very closely punctured. Thorax
deeply punctured, the scutellum, postscutellum and median segment
very sparsely, the disc of the mesonotum and posterior margin of
the pronotum smooth, the sides of the pronotum covered with long
erect black pubescence, the posterior truncation of the median
segment smooth. Abdomen with a few scattered piligerous punc¬
tures, the segments above and below fringed at the apex with long
black hairs, a transverse row of punctures some way before the apex
of each segment. Pygidium densely covered with black setae.
Tibiae and tarsi spinose and hairy.
Black ; the apex of the pygidium fusco-ferruginous. Wings
fuscous, fusco-hyaline near the apex, the whole glossed with green¬
ish blue. In some lights there are faint iridescent reflections on
the abdomen. Length 26 mm.
Hah. Gaza Land, Chirinda Forest (Odcndaal). Ex
col]. Marshall. March.
This is a very distinct species, having the wings more
obscure at the apex and hind margin than in Discolia
fasciatipennis, Sm. (alaris, Sauss.), the pubescence is also
longer and denser tlian in that species. It may possibl}’^
prove to be the female of Dielis hrachicera, Grib., and
seems to be more nearly allied to dimidiatipcnnis, Sauss.,
than to any other species.
Scolia {Dielis) crinita, Sauss.
Elis crinita, Sauss., Mem. soc. phys. et hist. nat. Geneve,
xiv, p. 65, 1854,
? Campsomeris princeps, Kirby, Trans. Ent. Soc. London,
p. 451, 1889, $.
I consider that there can be little doubt that these
forms are sexes of one species.
Scolia {Dielis) radula, Fabr.
Tiphia radula, Fabr., Syst. ent., p. 854, 1775,
Dielis formosa, var. maculiceps. Cam., Tijdsch. v. Ent.,
xlix, p. 218, 1906,
Cameron’s identification of his insect as a variety of
formosa, Guer., is quite mistaken ; there can be no doubt
that his description refers to radula. The male described
404' Mr, Rowla^ E. Turner’s Notes on the Scoliidae.
on the same page as septemcincta , Fabr., does not appear
to be that species, but the description is not clear.
The species of Dielis described by Mantero (Ann. Mus.
Civ. Storia Nat. Genova, xl, p. 592, 1900), and Schulz
(Berlin Ent. Zeitsch., xlix, p. 212, 1904), as D. formosa,
Guer., from New Guinea seems also to be wrongly identi¬
fied, for in true formosa the second recurrent nervure is
always complete. The shortening of that nervure is how¬
ever characteristic of D, subopaca, Turn., which is very
closely allied to Discolia cnlta, Sm., from New Guinea.
Scolia {Dielis) subopaca, Turn.
Campsomei'is {Dielis) subopaca, Turn., Ann. Mag. Nat. Hist.,
(8). iv, p. 175, 190<^, $
This is probably the Australian subspecies of Dielis
aurulenta, Sm., which has a wide range in the Malay
Archipelago. Dielis agilis, Sm., from Celebes seems to
me to be the male of aurulenta, not of lindenii as Saussure
suggests. The sexes of the Australian form being known,
it is unlikely that tlie male of aurulenta is very different
to that of subopaca. In aurulenta the second recurrent
nervure, though present, is often indistinct, in agilis it is
well marked ; in subopaca it is incomplete in both sexes,
and in Scolia culta, Sm., from New Guinea wholly absent
in the only specimen I have seen.
Scolia {Dielis) humboldti, Cam.
Dielis humboldti, Cam., Nova Guinea, v, 1, p. 53, 1903,
Hab. Bismarck Archipelago, Ralum.
Females from Ralum agree well with Cameron’s de¬
scription. There are two male species of Dielis in the
Berlin Museum from Ralum, one of which agrees with
the remarks of Cameron on a male Dielis, sp. 4, 1. c. p. 53,
which is almost certainly the male of huinboldti ; the
other, which is probably the male of ferrugineipes, Mant.,
has the clypeus wholly yellow, also the pronotum, base of
the scutellum, a transverse line on the postscutellum, the
four basal abdominal segments at the apex and the outer
side of the tibiae. The pubescence on the median seg¬
ment is grey in humboldti pale fulvous on the other
male, and in the latter the first abdominal segment is
distinctly narrower in proportion to the length. But this
male can only be assigned to ferrugineipes with doubt,
Mr. Rowland E. Turner’s Notes on the Scoliiclac. 405
Scotia {Diclis) fernigineipcs, Mant.
Elis {Eielis) o.nmdata, Fabr., var. ferrugineijns, Mant.,
vVnn. Mus. Civ. Storia Nat. Genova, xl, p. 591,
1900, $.
I see no sufficient reason for regarding this as a variety
of annulata, seeing that two or three very similar species
occur in New Guinea. There is a specimen in the Berlin
Museum from Ralura, in the Bismarck Archipelago.
Family SAPYGIDAE.
Parasapyga, gen. nov.
Differs from Sapyga in the antennae, which are
rather slender, thicker at the base than towards the apex ;
the much more robust build, the vertical truncation of the
median segment and the almost conical abdomen. The
second cubital cell is longer on the radial nervure than
high, as in Polochrum, and the third cubital cell is not
much narrowed on the radial nervure, but the second
recurrent nervure is received at about one quarter from
the base of the third cubital cell, somewhat further from
the base than in Sapyga and not interstitial with the
second transverse cubital nervure as in Polochrum.
Parasapyga mdlleri, sp. nov. (Plate L, fig. 8.)
9 . Clypeus broadened from the base to the apex, the apical margin
feebly sinuate on each side, coarsely reticulate, with a low carina
from the base not reaching the apex. Maxillary palpi small, six-
jointed, the basal joint the longest. Head scarcely broader than the
pronotum, coarsely punctured ; the posterior ocelli further from the
eyes than from each other, the eyes deeply emarginate, not quite
reaching the base of the mandibles ; the interantennal prominence
broadly rounded at the apex, the sides raised and forming low carinae
close to the base of the antennae. Mandibles strongly bidentate at
the apex. Antennae inserted much further from each other than
from the eyes, gradually tapering towards the apex, the scape finely
punctured, the first joint of the flagellum short, the second and third
about equal in length. Thorax, median segment and pleurae coarsely
and closely punctured, the anterior margin of the pronotum almost
straight, very broad, the median segment vertically truncate from
just behind the postscutellum. Abdomen almost conical, shining,
finely and rather sparsely punctured, the first segment very little
406 Mr. Rowlaf^ E. Turner’s Notes on the Scoliidae.
narrower than the second, concavely truncate at tlie base, the apical
segment acute. Legs without spines, except at the apex of the tibiae,
the basal joint of the tarsi as long as all the otliers combined. Second
abscissa of the radius nearly three times as long as the first and more
tlian half as long as the third.
Black ; the inner margins of the eyes as high as the emargination,
the outer margins, the apex of the interantennal prominence, a
transverse band on each side of the clypeus near the apical margin,
the anterior margin of the pronotum narrowly interrupted in the
middle, and a small spot on each side at the basal angles of the
scutellum and postscutellum pale yellow ; abdomen ferruginous tarsi
fuscoferruginous. Wings fusco-hyaline, clear hyaline at the extreme
base, nervures black. Length 15 mm., length of costa 10 mm.
Hah. Sikkim, Tukvar, 4000 ft. {Mdllcr).
Type in B. M.
( 407 )
XIV. New Fossorial Hymenoptera from Australia. By
Rowland E, Turner, F.Z.S.
[Read June 1st, 1910.]
Plate L.
Most of the new species described in this pajier are from
the collection of Dr. R. C. L. Perkins, who has kindly
handed them to me for identification and description.
Most of the species from the Cairns district were taken
by the well-known collector, Mr. F. P. Dodd. They
include representatives of two genera, Spilomena and
Piagetia, not hitherto recorded from Australia, also further
species of the rare Australian genus Sphodrotes.
Family THYNNIDAE.
Rhagigaster unicolor, Guer., st. mutatus, st. nov.
Differs from the typical Sydney form in the following
details : —
The head is a little broader than long, the median
segment much more strongly broadened from the base,
the vertex and front more closely and coarsely punctured
and the sixth dorsal segment narrower in the middle, not
triangular as in unicolor, but more sharply narrowed from
the base.
The triangular space at the apex of the clypeus is
much narrower, and is rugose instead of smooth.
Hah. S. Queensland, Tambourine Mts., Toowoomba.
The differences in the local races of the male are very
slight, but the females differ much more especially in the
shape of the head and median segment. In these points
the S. Queensland form approaches the Victorian form
lyelli, Turn., much more nearly than the Sydney form.
Tachyiiomyia adusta, Sni.
Thynnus adushcs, Sm., Cat. Hym. B.M.,vii, p. 43, 1859, $.
Aelitrus inlosidus, Sm., Cat. Hym. B.M., vii, p. 56, 1859,
A male of this species from the Tambourine Mts., S.
Queensland, has the legs black, the wings a little more
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
408
iVfF Rowland E. Turner on
clouded with fuscous and the hypopygiuin with only one
lateral notch near the base. For this form I propose the
name
Tachynomyia adusta, vSm., st. imiformis, st. nov.
Agriomyia hermanni, sp. nov.
9. Head very thin, large, scarcely convex, more than half as
broad again anteriorly as long, rounded at the sides, nearly as broad
on the posterior margin as long, the front closely and microscopically
punctured, with a median longitudinal sulcus, an obscure oblique
depression on each side in the ocellar region, the vertex sparsely
punctured. Clypeus with a short carina not extending to the apex.
Eyes small, touching the base of the mandibles, the antennae
inserted as far from each other as from the eyes. Pronotuin much
narrower than the head, more than half as broad again anteriorly
as long, slightly narrowed posteriorly, slightly convex, the anterior
angles depressed and produced into short spines, shining, with
scattered punctures, a very short sulcus from the middle of the
anterior margin, the projjleurae finely aciculate. Scutellum narrow,
a little longer than the breadth at the base, narrowed to the apex,
sparsely punctured. Median segment closely punctured, thinly
covered with cinereous pubescence, no longer than the scutellum,
broadened from the base and obliquely truncate posteriorly. Abdo¬
men shining, with a few scattered punctures, the first segment, with
a broad depression before the feebly raised apical margin, the
depression scarcely arched in the middle, second segment with four
strong transverse carinae, including the raised apical margin, the
fifth segment longitudinally subcarinate in the middle ; pygidium
elongate ovate, with a carina from the base to the middle.
Black ; the mandibles, clypeus, antennae, legs, sides of the
abdominal segments, carinae of the second dorsal segment and
base of the pygidium fusco-ferruginous. Length 10 mm.
Clypeus produced and narrowly truncate at the apex,
sparsely punctured. Head rounded, finely and shallowly punctured,
with a shining longitudinal line on the front, the antennae inserted
a little nearer to the eyes than to each other. Thorax closely
punctured ; the scutellum shining, more sparsely punctured and
rather strongly convex. Median segment obliquely depressed.
Abdomen fusiform, shining, sparsely and shallowly punctured ;
hypopygiuin very broadly triangular, slightly rounded at the
apex, with a short apical spine; the first ventral segment truncate
at the apex and subtuberculate.
Black ; the clypeus, the inner margins of the eyes broadly as high
New Fossorial Hymeno2:)tera from Australia. 409
as the base of the antennae, a short oblique line on each side
between the antennae, a small spot on the vertex, the margins of
the pronotum, tegulae, basal angles of the scutellum, a spot on the
mesopleurae, postscutellum, a broad transverse band emarginate
posteriorly on the median segment, a narrow transverse band
narrowly interrupted in the middle on dorsal abdominal segments
2-6 and on ventral segments 2-4 pale yellow. Wings pale dull
flavo- hyaline, nervures pule ferruginous. Length 13 mm.
Hah. Darnley Island {Elgner).
The female is somewhat intermediate between rotundi-
ceps, Sm., and luctuosus, Sm., differing from the former in
the much flatter pronotum, from the latter in the different
shape and more convex surface of the pronotum, and from
both in the rather broader pygidium and the shallow
depressions on the head. The male has the hypopygium
much broader and more triangular than in either of the
two species mentioned above, and the colour of the wings
and median segment differs. Small colour variations in
the males of Agriomyia seem often to be associated with
structural differences in the females, and should not be
treated as of no value. In some genera of the family,
however, the yellow markings of the male vary very much
in the same species.
Agriomyia manifesta, sp. nov.
9 . Head thin, large, not flattened but slightly convex, more than
half as broad again anteriorly as long, rounded at the posterior
angles, subopaque and without punctures, with a very short frontal
sulcus ; eyes small, oval, almost touching the base of the mandi¬
bles. Thorax shining and almost smooth, the median segment very
sparsely punctured, pronotum a little broader anteriorly than long,
narrowed posteriorly, raised in the middle of the anterior margin
into an indistinct hump-like prominence which is divided at the
apex by a very faint longitudinal sulcus, the anterior angles pointed.
First dorsal abdominal segment transversely depressed at the apex,
second segment with five transverse carinae including the raised
apical margin, the basal carina low and very near the second.
Pygidium elongate ovate, narrowed at the base, with a small tuft of
bright fulvous hairs on each side.
Black ; antennae, legs, second dorsal segment and pygidium fusco-
ferruginous. Length 9 mm.
(J . Clypeus produced and narrowly truncate at the apex, shining,
410
Mr. Rowland E. Turner on
very sparsely punctured. Antennae inserted as far from each otlier
as from the eyes, the fi’ont between tliem not raised, and divided by
a well-marked, short, longitudinal sulcus. Head rounded, not large,
closely punctured. Thorax closely punctured, the median segment
almost smooth, oblique from the postscutellum. Abdomen very
sparsel}' and .shallowly punctured, the transverse line at the base
of the segments not very deeply impressed. Hypopygium very
broadly rounded at the apex, with a short apical spine.
Black ; clypeus, front between the antennae, inner margins of the
eyes very broadly as high as the base of the antennae, a small spot
on the vertex, the anterior margin on the pronotum broadly,
tegulae, a quadrate spot on the mesonotum, a small spot on the
mesopleurae below the anterior wings, a small spot on each side
at the anterior angles of the scutellum, postscutellum, the whole
of the oblique slope of the median segment, a broad continuous
band narrowly emarginate in the middle posteriorly on the second
and third dorsal segments of the abdomen, a narrow band scarcely
interrupted on the first segment, an interrupted band on the fourth
and fifth segments, the apex of the tubercle on the first ventral
segment and transverse bands on ventral segments 2-4 yellow.
Wings hyaline, nervures fuscous. Legs and the undersurface of
the scape ferruginous. Length 11 mm.
Hah. Queensland, Bundaberg (ex coll. Perkins) ; ^ ^ in
CO'p.
The female closely resembles rotiindiceps, Sm., but the
head is a little smaller and slightly convex, not flattened
as in rotundiceps. The hump on the anterior margin of
the pronotum is more strongly developed in rotundiceps.
The male closely resembles A. maculata, Gu^r., var.
variegata, Klug., which may prove to be a good species;
the clypeus in the present species is a little more convex,
the bands on the second and third segments are con¬
tinuous, and the depressed line on the base of the
segments less distinct, the yellow mark on the median
segment is not trilobed posteriorly, the scape is ferruginous
beneath and the hypopygium is much more broadly
rounded at the apex. The males of this genus are very
difficult to distinguish, but colour differences appear to
be more constant than in some genera of the Thynnidae.
The female of variegata appears to be quite different, but
I have not seen specimens Avhich can be referred to that
species with absolute certainty.
Neio Fossorial HymenoptcQ'a from Australia. 411
Zdcboria longicornis, Turn.
Thynnus {Zeleboria) longicornis. Turn., Proc. Linn. Soc.
N.S. Wales, xxxiii, p. 108, 1908, $ .
5 . Head shining, very sparsely punctured, half as broad again
anteriorly as long, strongly rounded posteriorly, a short, deep, longi¬
tudinal sulcus on the front ; the clypeus with a carina from the base
to the apex. Thorax finely and not very closely punctured, the
pronotum much narrower than the head, a little longer than broad,
the sides almost parallel, a deep longitudinal sulcus from the anterior
margin not reaching half-way to the middle. Median segment
long, not much shorter than the pronotum, the dorsal surface flat,
broadened from the base and almost vertically truncate posteriorly.
First abdominal segment closely and shallowly punctured, the apical
margin transversely depressed ; second segment with six well-marked
transverse carinae, including the raised apical margin; segments
3-5 shining, shallowly punctured; the fifth ventral segment more
coarsely punctured. Pygidium deflexed, almost twice as long as the
greatest breadth, sparsely punctured and rounded at the apex.
Black ; the antennae, mandibles and spines of the tarsi fusco-
ferruginous. Length 12 mm.
Hah. South Queensland, Tambourine Mts.
The female is nearly related to other species of Zeleboria.
The locality, North-West Australia, given for the type
male may have been erroneous, as it is not likely that the
species ranges across the continent.
Asihenotliynnus Icnrandcnsis, sp. nov.
d . Clypeus produced and narrowly truncate at the apex, trans¬
versely depressed before the apex. Head shining, with a fe^v
scattered punctures, the interantennal prominence almost transverse
at the apex, raised above the base of the scape and divided by a
shallow sulcus. Antennae slender, as long as the head, thorax and
median segment combined, the joints moderately arcuate beneath.
Posterior ocelli nearly twice as far from the eyes as from each other.
Thorax shining and sparsely punctured, the pronotum much narrower
than the head, narrowed anteriorly and broadly emarginate on the
anterior margin, scutellum broadly rounded at the apex. Median
segment a little longer than the breadth at the base, finely aciculate.
Abdomen flattened, elongate and slender, the sides nearly parallel,
narrowed at the base ; segments 1-4 depressed on the apical margin,
most broadly in the middle ; the first segment longer than the second,
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) E E
412
Mi\Rowland E. Turner on
more than twice as broad at the apex as at the base, shining, with a
sulcus from the base to the middle. Hypopygium produced into an
apical spine, with a very short spine on each side near the base;
claspers rather broad at the base, narrowly produced and pointed at
the apex.
Black ; mandibles, clypeus, margins of the eyes broadly, narrowly
interrupted on the summit by a ferruginous line, and continued
across the vertex, the margins of the pronotum, a quadrate spot on
the mesonotum, a spot on the scutellum, the tegulae, a broad band on
the mesopleurae in front and a spot before tlie intermediate coxae, a
spot in the middle of the median segment and a curved band on each
side, a spot on each side near the apex of abdominal segments 1-5,
the coxae and a line on the femora and tibiae bright yellow ; an
oblique line on each side on the vertex, a round spot on the yelhjw
band behind the eyes, the abdomen and the legs ferruginous. Wings
hyaline, slightly iridescent, nervures fuscous, stigma pale testaceous
brown. The second abscissa of the radius is equal in length to the
third, the tirst recurrent nervure is received at two-thirds from the
base of the second cubital cell, the second at one quarter from the
base of the third cubital cell. Length 6 mm.
$ . Clypeus short, with a carina. Head smooth and shining,
almost semicircular, not quite twice as broad anteriorly as long, very
slightly convex, the front concave on each side above the base of the
antennae, the concave spaces much broader than long. Eyes small,
situated close to the base of the mandibles ; antennae no longer than
the anterior breadth of the head. Thorax and median segment
smooth and shining, the pronotum much narrower than the head,
nearly twice as broad as long, the anterior angles produced acutely,
an indistinct longitudinal carina in the middle, the sides almost
parallel ; scutellum much broader than long, rounded posteriorly ;
median segment shorter than the pronotum, obliquely truncate
posteriorly, produced anteriorly into a pointed tubercle. Abdomen
smooth with a few scattered punctures, the segments broadly de¬
pressed at the apex, the raised portion before the depression strongly
bilobed, second segment with two transverse carinae in addition to
the raised apical margin. Fifth ventral segment almost smooth.
Pygidium deflexed, narrow, lanceolate, very slightly widened and
rounded at the apex.
Light castaneous brown, the abdominal segments pale luteous on
the sides and apical margins. Length 3 mm.
Hob. Queensland, Cairns.
A variety of the male has the abdomen black with
yellow markings and the segments more or less fusco-
New tossorial Hymenoptera from Australia. 413
ferruginous at the apex, but I can find no structural
difference. The tubercle on the median segment of the
female is remarkable.
AsthenothynmLS ruh'omacidatus^ Turn.
Thynnus {Aeolotkynnns) rubromaculatus, Turn., Proc.
■■ ■ Tiinn. Soc. N.S. Wales, xxxiii, p. 118, 1908,
9 , Head and thorax smooth and shining ; the head nearly half
as broad again as long, slightly convex, very strongly rounded
posteriorly, the clypeus without a carina ; no sulcus on the front ;
the antennae very short, not equal in length to the distance between
the eyes, which are very small and situated near the base of the
mandibles. Thorax very narrow, not more than one-third as broad
as the anterior margin of the head, the pronotum distinctly longer
than broad, the sides parallel, without a sulcus, the anterior angles
not produced ; scutellum a little narrower than the pronotum ;
median segment slightly broadened from the base, obliquely sloped
posteriorly, the dorsal surface before the slope only a little longer
than the scutellum. Abdomen as broad as tlie head, shining, with
a few scattered punctures at the base of the apical segments ; second
segment with a transverse carina close to the base, another, lower,
just before the apex, the apical margin feebly raised ; segments 3-5
with a strongly-raised mark rounded posteriorly on each side at the
base. Fifth ventral segment punctured. Pygidium long and very
narrow, almost linear, strongly compressed on the sides. Inter¬
mediate tibiae scarcely broader than the posterior pair, the inter¬
mediate tarsi slender.
Dark reddish brown, the sides of the abdomen and the apical
portion of the dorsal segments pale luteous ; the head beneath dull
ferruginous. Length 4 mm.
Hah. South Australia, Adelaide {Perkins)-, in
cop.
This species differs considerably from others of the genus
in the sculpture of the second dorsal segment of the
abdomen and the absence of a frontal sulcus. The male
also differs in the blunt hypopygium from most other
species of the genus.
Asthenothynnus minutissimus, sp. nov.
$ . Clypeus produced and narrowly truncate at the apex, shining,
finely punctured at the base, smooth at the apex. Head shining,
EE 2
414
r. Rowland E. Turner on
linely and sparsely punctured, with a short fronted sulcus, the
interantennal prominence low and transverse at the apex. Antennae
rather shorter than the thorax and median segment combined, the third
joint of the flagellum much longer than the second, the apical joints as
thick as the basal. Thorax shining, sparsely punctured, the median
segment very minutely punctured; the pronotum narrower than
the head, very feebly and widely emarginate anteriorly, scutellum
convex, broadly truncate at the ajjex. Abdomen flattened, elongate
fusiform, smooth and shining, the segment narrowly depressed on
the apical margin. Hypopygium narrow, much longer than broad,
the sides almost parallel, blunt at the apex and without a .spine.
Third abscissa of the radius longer than the second, the second
recurrent nervure received at one-third from the base of the third
cubital cell.
Black ; the mandibles, the margins of the eyes narrowly (except
at the summit), the margins of the pronotum, a spot on the
mesonotum, the postscutellum, a small spot on each side at the
apical angles of the median segment, the apex of the tegulae, a
minute spot on the mesopleurae below the anterior wings, a
lateral spot on each side of abdominal segments 2-6, narrowly pro¬
duced on the apical margin of segments 5-6 and the anterior tibiae
beneath pale yellow ; legs testaceous brown. Wings hyaline,
slightly iridescent, nervures pale testaceous brown. Length 3 mm.
Hah. S. Australia, Adelaide {Perkins).
Nearly allied to pygmaciis, Turn., but is a smaller
species and much more finely punctured.
Asthcnothynnus ? 2J^rkinsi, sp. nov.
(J. Clypeus produced and narrowly truncate at the apex. Head,
thorax and median segment finely and very closely punctured ; the
head a little broader than the thorax ; antennae filiform, a little
longer than the thorax and median segment combined, the apical
joints very feebly arcuate beneath ; the interantennal carina trans¬
verse. Posterior ocelli half as far again from the eyes as from each
other. Pronotum a little narrowed anteriorly ; the scutellum very
broadly rounded at the apex ; median segment rounded. Abdomen
elongate, tapering slightly at the extremities, flattened, the segments,
very slightly depressed at the base, shining, finely and shallowly
punctured, the four basal segments narrowly depressed on the apical
margin. Hypopygium broad, the sides parallel, the apical angles
produced into short spine.s, with a strong median spine. Third
abscissa of the radius a little longer than the second ; first recurrent
nervure received at two-thirds from the base of the second cubital
Neiv Fossoricd Hymenoptcra from Australia. 415
cell. A very deep longitudinal sulcus reaches from the base to the
middle of the first dorsal segment of the abdomen.
Black, with short whitish pubescence on the head and pleurae ;
the anterior margin of the clypeus, mandibles, the apex of the in-
terantennal prominence, the margins of the pronotuni, tegulae, a
longitudinal line on the scutellum and the pustscutellum yellow;
an obscure spot on each side on the vertex, the second and third
abdominal segments, the apex of the first and the base of the fourth
dark ferruginous ; the tibiae, tarsi and apex of the femora testaceous
brown. Wings hyaline, nervures fuscous. Length 8-10 mm.
Hah. Queensland, Bundaberg {Perldns) \ two specimens.
This is not a typical Asthcnothynnus^ but is better
placed in that genus than in any other.
Eimdiotliynnns arcid, sp. nov.
5 . Head flattened, twice as broad anteriorly as posteriorly, not
rounded at the sides, slightly raised in the middle and depressed
on the sides, subopa([Ue, with a few punctures in longitudinal
rows on the front, and a very short frontal sulcus. Eyes oval,
touching the base of the mandibles. Pronotum longer than broad,
flattened, subopaque, with an obscure median carina, the sides
almost parallel. Scutellum rounded at the apex ; the median
segment shorter than the scutellum, obliquely truncate posteriorly,
shining, very sparsely punctured ; the pronotum about half as long
again as the scutellum and median segment combined. Abdomen
broader than the thorax, shining, the segments very finely aciculate
at the base, with a few deep punctures before the apex ; the first
segment with a transverse groove just before the apex, the second
with three strong transverse carinae in addition to the less strongly-
raised apical margin ; the third and fourth segments narrowly
depressed on the apical margin. Pygidium almost vertically
truncate, very narrowly elongate ovate, with a tuft of golden hairs
on each side. Fifth ventral segment coarsely punctured. Inter¬
mediate and posterior tibiae broad and spinose, tarsi slender.
Black, the abdominal segments on the sides and at the apex
very pale luteous ; tibiae and tarsi luteous brown ; antennae and
mandibles fusco-ferrugiiious ; the posterior slope of the median
segment thinly clothed with long white pubescence. Length 6 mm.
Mandibles bidentate, the inner tooth very short and blunt.
Clypeus produced and narrowly truncate at the apex, with a low
carina from the base to the middle, clothed with long white
pubescence, finely and closely punctured, more closely at the base
than at the apex. Head, thorax and median segment finely and
416
T. Rowland E. Turner on
closely punctured ; with sparse grey pubescence, closer and longer
on the head than on the thorax ; antennae no longer than the
thorax and median segment combined, scarcely narrowed to the
apex, the joints slightly arcuate beneath, the third joint of the
flagellum more than half as long again as the second, the carinae
between the antennae only slightly oblique ; the posterior ocelli a
little further from the eyes than from each other. Pronotum
short, strongly narrowed anteriorly ; scutellum strongly convex ;
median segment rounded. Abdomen elongate, tapering slightly
at the extremities, shining and shallowly and finely punctured,
segments 2-5 strongly constricted at the base, the first segment
with a longitudinal sulcus from the base to the middle.
Hypopygium lanceolate, produced into a long acute spine on each
side at the base. Second recurrent nervure received very near
the base of the third cubital cell ; second abscissa of the radius
longer than the third.
Black ; the apical half of the clypeus, marked with black on
the centre, the base of the mandibles, the margins of the
pronotum, the interantennal carinae, a spot on the tegulae, a small
longitudinal spot on the posterior portion of the mesonotum, a
small spot on the scutellum and a minute one at each of the
anterior angles, a transverse line on the postscutellum, a spot on
each side of the first abdominal segment and a short narrow band
on each side of segments 2-5 near the apical margin pale yellow ;
legs fuscous brown, lighter on the anterior pair, calcaria white.
Wings hyaline, nervures fuscous, the stigma pale testaceous.
Length 10 mm.
Hah. Queensland, Cairns (ex coll. Perkins) ; 2$.
Very near hipao'titus. Turn., especially in the male sex,
but is a smaller and more slender species. In hipartitus
the female has a strong tubercle on the pronotum. The
colour of the legs in the male seems to be a constant
difference in the limited number of specimens I have seen,
though in some species of Thynnidae this character is
quite unreliable.
Acolothynnus InchrymosuSy sp. nov.
d- Clypeus slightly advanced and narrowly truncate at the
apex, finely and closely punctured and thinly clothed with whitish
pubescence. Head and thorax closely punctured rugose ; antennae
as long as the thorax and median segment combined, of almost even
thickness throughout, the third joint of the flagellum nearly half
as long again as the second, but slightly shorter than the fourth.
Neio Fussorial Hymenoptera from Australia. 417
the interantennal carina depressed in the middle, almost transverse.
Pronotum almost as broad as the head, scarcely narrowed anteriorly,
the anterior margin strongly and broadly emarginate ; scutellum
closely punctured, with an obscure median carina, broadly rounded
at the apex. Median segment closely punctured, rounded. Abdomen
elongate, the sides almost parallel, the segments punctured and very
strongly constricted at the base, the sixth segment with an acute
spine on each side at the apical angles beneath ; hypopygium rather
short, very broad, not narrowed at the apex, ending in three long
spines, the central spine the longest. Second abscissa of the radius
more than twice as long as the third, second recurrent nervure
received at about one-eighth from the base of the third cubital cell.
Black ; the base of the mandibles, a narrow transverse line close
to the apex of the clypeus, a small transverse spot on each side above
the base of the antennae, the anterior margin of the pronotum very
narrowly and the postscutellum whitish yellow; calcaria whitish.
Wings hyaline ; nervures fuscous. Length 9 mm.
Hah. South Australia, Adelaide.
Differs from dimidiatus, Westw., in tlie narrower clypeus,
tlie broader hypopygium and the colour; from armiger.
Turn., in the much broader hypopygium and the broadly
emarginate pronotum. The very broad hypopygium
separates it from umhrip)ennis, Sm., as well as the smaller
size and colour differences.
Zaspilothynnus rufolutius, Tinn. (Plate L, fig. 1 fig. 2 ^.)
Thynnus rufoluteus, Turn., Ann. Mus. Nat. Hung., 1910,
p. 114.
d . Clypeus longer than In-oad, convex at the base, deflexed
towards the apex, the apical margin broadly truncate, finely longi¬
tudinally striated ; the labrum produced far beyond the clypeus,
broad and rounded at the apex, without an eniargination. Mandibles
slender, with a very feeble tooth on the inner margin near the apex.
Head finely and closely punctured, the interantennal prominence
narrowly rounded at the apex, almost touching the base of the
clypeus. Antennae shorter than the thorax and median segment
combined, of about equal thickness throughout, inserted as far
from each other as from the eyes. Thorax very finely and closely
punctured, pronotum not as broad as the head, narrowed anteriorly,
the margin raised ; scutellum convex, broadly rounded at the apex.
Median segment minutely punctured, oblique. Abdomen elongate,
longer than the head, thorax and median segment combined, tapering
slightly towards the apex, minutely and sparsely punctured, the
418
Ml^( )wland E. Turner mi
seventh dorsal segment produced into a flattened plate which is
narrowly rounded at the apex and closely longitudinally striated ;
a spine on each side at the apical angles of the sixth ventral segment ;
hypopygium with a blunt spine on each side near the base, thence
sharply narrowed and elongate to the base of the acute apical spine.
Black ; base of the mandibles ; clypeus, a V-shaped mark between
the antennae, the orbits of the eyes narrowly interrupted on the
summit, a narrow transverse band on the vertex, a broad band on
tbe posterior margin of the pronotum, the anterior angles of the
pronotum, tegulae, a large quadrate spot on the mesonotum, a
vertical band on the inesopleurae, the posterior half of the scutellum
and a small spot at each of the anterior angles, the postscutellum,
a broad band narrowed towards the apex on the median segment, a
transverse band on dorsal abdominal segments 1-6, interrupted in
the middle on segments 2-5, a transverse band interrupted in the
middle on ventral segments 2-4, the intermediate and posterior
coxae beneath and the process of the sternum between the inter¬
mediate coxae bright yellow, femora at the apex, tibiae and tarsi
light reddish brown. Wings hyaline, nervures black, the stigma
pale testaceous browm. Length 17 mm.
$ . A female from the same locality differs from the type in having
the ground colour almost black instead of ferruginous.
Hah. Cape York, Q. {Elgner).
The type female in the Hungarian National Museum is
from Cooktown.
Family SPHEGIDAE.
Sinlomena australis, sp. nov. (Plate L, fig. 9.)
9 . Clypeus raised into a median carina, which broadens at the
apex and forms a smooth triangular surface. Head very finely
rugose, a carina from the anterior ocellus to the base of the clypeus,
the inner orbits of the eyes raised and forming distinct carinae.
Antennae inserted low down on the sides of the clypeus, much
further from each other than from the eyes, the flagellum a little
more than twice as long as the scape. Eyes at the base of the
clypeus separated by a distance nearly equal to twice the length
of the scape, on the vertex by less than three-quarters of that
distance ; the posterior ocelli a little nearer to each other than to
the eyes. Thorax narrower than the head, very minutely punctured,
the pronotum depressed, much narrower than the mesonotum, the
angles reaching the tegulae, mesonotum and scutellum broader than
long; a well-defined broadly triangular area almost covering the
dorsal surface of the median segment, rugose and longitudinally
Neiu Fossoricd Hymenoptera from Australia. 419
striated, the two median striae more distinct than the lateral ; the
posterior truncation of the segment finely transversely striated.
Abdomen smooth and shining, fusiform, not petiolate. The second
cubital cell is much higher than long, the first abscissa of the radius
longer than the second, the recurrent nervure received just before
the apex of the first cubital cell. Stigma nearly twice as long on the
costa as its greatest breadth.
Testaceous ; the head black, antennae testaceous. Wings hyaline,
strongly iridescent, nervures and stigma pale testaceous. Length
3 '5 mm.
Hah. N, Queensland, Kuranda, August (ex coll.
Ferhins).
This is the first species of Spilomena recorded from
Australia; it is much larger than the European species,
but only differs slightly in neuration and structure. In
colour there is a close resemblance to Fison melanocephalum,
Turn., and to a species of Halictus, both found in the
same locality. It is remarkable that this plan of colouring
should be prevalent at Cairns, whereas at Mackay several
degrees further south none of these species occur. It is
possible that the colouring may have some connection
with the conditions which prevail locally, Cairns being a
district of dense tropical jungle, and open grass country
largely predominating at Mackay.
Sphex mimulus, sp. nov.
9 . Mandibles with three teeth, the outer one long and pointed,
the second, on the inner margin, strong and broad, the third small.
Clypeus almost transverse on the apical margin, produced in the
middle into two small teeth which are separated from each other
by a shallow emargination. Head subopaque, very shallowly
punctured, the front and clypeus clothed with very fine silver
pubescence, with sparse, long, black hairs. Posterior ocelli a little
further from the eyes than from each other ; second joint of the
flagellum half as long again as the third joint. Scutellum flat, post-
scutellum without a groove or tubercles, median segment coriaceous,
with a few indistinct striae on the sides. Abdomen shining, with¬
out punctures, the petiole of the basal segment as long as the third
joint of the posterior tarsi. Second cubital cell rhomboidal, the
third very short on the radial nervure ; first recurrent nervure
^ received close to the apex of the second cubital cell, almost inter¬
stitial with the second transverse cubital nervure ; radial cell
on
420 Mr^iovvland E. Turner
truncate at the apex. Tarsal ungues with two teeth near the
base.
Black ; the fifth and sixth abdominal segments and the base and
apex of the fourth bright chestnut brown. Wings fulvo-hyaline,
broadly fusco-hyaline at the apex, nervures fuscous. Length 23 mm.
Hah. Queensland, Cairns.
This species closely resembles Zaspilothynnus excavatus,
Turn., in size and colour. It is closely allied to Sphex
confrater, Kohl., from New Britain, but the shape of the
clypeus is different. As far as I know no Thynnus of
similar colouring occurs in New Britain, nor is the plan
of colouring one that is at all prevalent among Queensland
Hymenoptera. The present species must be a rarity as
I have never seen another specimen. Z. eoxavatus is a
fairly commcn insect in the tropical scrubs of the Cairns
district, and also occurs at Cooktown. '
Key to the Australian Species of
THE Genus Cerceris.
? ?
A. Clypeus with a prominent lamina
projecting from near the base.
a. The lamina arched, much longer than
broad. Orange, me.sonotum and
fourth abdominal segment black . G. aurantiaca, Sm.
b. The lamina short. Black with yellow
marks on front and thorax, second,
fourth and fifth segments of abdo¬
men with narrow yellow apical
bands . C. opposita, Sm.
B. Clypeus without a lamina.
a. Triangular space at the base of the
medium segment smooth and shin¬
ing.
a^. Clypeus with a small tubercle on
each side near the middle, de¬
pressed from the middle to the
apex.
a-^ Reddish orange ; vertex, nieso-
thorax, third and fourth abdo¬
minal segments black . . . C. victrix, Turn,
b’*. Black ; three apical segments of
abdomen more or less cas-
taneous
G. perkinsi, Turn,
Ncio Fossorial Hymmo'pUra from Australia. 421
b^. Clypeus without tubercles.
a-\ Clypeus porrect, widely and
deeply emarginate ; abdomen
smooth and shining .... 0. euchroma, Turn,
bb Clypeus less strongly porrect,
very shallowly emarginate or
transverse at apex ; abdomen
closely punctured,
ah Clypeus shallowly emargin¬
ate ; triangular space at
base of median segment
with two yellow spots ;
third abdominal segment
only black entirely . . . G. multiguttata, Turn
bh Clypeus transverse ; triangular
space unspotted ; third and
fourth abdominal sf-gnients
black . 0. inexpectata, Turn.
b. Triangular space at base of median
segment not smooth,
a^. Triangular space almost wholly
striated.
a^ Triangular space finely trans¬
versely striated . C. Jluvialis, Sm.
b^. Triangular space longitudinally
or obliquely striated,
a^. The striae close and almost
longitudinal. Black, second
and fifth abdominal seg¬
ments orange . G. varipes, Sm.
b'*. The striae oblique, not reach¬
ing the apex. Black,
marked with dull yellow
and ferruginous . ... G. labeculata, Tura
b^. The triangular space striated in the
corners only or entirely punctured,
a^ Clypeus porrect and broadly
emarginate . G. vennsta, Sm.
b^. Clypeus not porrect.
First abdominal segment
shorter than the breadth
at the base.
a^ Black, two basal and fifth
abdominal segments, legs
422
Mr. Rowland E. Turner on
and marks on head and
thorax red orange . .
b". Black, marked with dull
yellow and ferruginous .
b^. First abdominal segment as
long as broad or longer,
a^ Abdomen, except basal seg¬
ment, sparsely punctured ;
ochreous yellow, first and
third segments black . .
b’’. Three basal abdominal seg¬
ments at least deeply and
closely punctured, abdo¬
men marked with yellow
or yellow and ferruginous,
a®. Fourth and fifth segments
more sparsely punctured
than the basal ones,
thorax wdiolly black
b'’. Fourth and fifth segments
as closely punctured as
the basal ones, post-
scutellum and a spot on
each side of the pro-
notum yellow,
ah Scutellum black,
abdomen marked
with yellow and
ferruginous . .
IF. Scutellum yellow,
abdomen with
yellow markings
0. gilesi, Turn.
G. ant'q^odes, Sm.
C. australis, Sauss.
G. saeva, Sm.
G. minuscula, Turn.
G. praedura, Turn.
Cerceris victrix, sp. nov. (Plate L, fig. 10.)
$ . Mandibles acute at the apex, notched near the middle of the
inner margin, with a blunt tooth on the basal side of the notch.
Clypeus broad, almost transverse at the apex, with four blunt teeth
on the apical margin of the middle lobe, which is subconcavely
depressed from near the middle, with a small tubercle on each side
at the base of the depression, the apical half smooth and shining, the
basal sparsely punctured. Front longitudinally rugose-striate, vertex
punctured-rugose ; antennae inserted fully half as far again from the
eyes as from each other, with an elevated carina between them, the
second joint of the flagellum as long as the first and third combined,
Neiv Fossorial Hymenoptera from A%istralia. 423
the apical joint equal in lengtli to the tenth and rounded at the
apex ; the base of the antennae separated from the base of the
clypeus by a distance equal to the length of the third joint of the
flagellum. Ocelli in a very broad triangle, the posterior pair fully
half as far again from the eyes as from each other ; the eyes
separated from the posterior margin of the head by a space nearly
equal to their greatest breadth. Thorax much narrower than the
head, longitudinally rugose-striate on the mesonotum and scutellum,
pleurae rugosely punctured. Median segment and abdomen strongly
punctured, the triangular space at the base of the median segment
smooth. First abdominal segment short, more than half as broad
again as long ; pygidial area rather broadly truncate at the apex, half
as broad again at the base as at the apex, finely granulate.
Reddish orange ; the vertex, mandibles, mesonotum, pleurae,
thorax beneath, the apex of the median segment in the middle, the
third and fourth abdominal segments and the apex of the pygidial
area black. Wings flavo-hyaline, with a fuscous cloud along the costa
near the apex, nervures ferruginous. Length 16 mm.
^ . Differs from the female in the following points : front,
mesonotum and scutellum closely punctured, the triangular space at
the base of the median segment transversely striated, the middle lobe
of the clypeus much longer than broad, without tubercles in the
middle and not depressed at the apex, the antennae separated from
the base of the clypeus by a distance equal to the length of the
first and second joints of the flagellum combined, and the extent of
the black area on the vertex and median segment is much less.
Length 14 mm.
Hah. Queensland, Cairns ex coll. Ferhins) ; Moreton
Bay ($ ex coll. F. Smith).
I think there can be no doubt that these are sexes of
the same species.
Cerceris perhinsi, sp. nov. (Plate L, fig. 12.)
9 . Clypeus short, strongly porrect and broadly emarginate at
the apex, coarsely punctured, smooth and shining at the base.
Mandibles shining, with a broad triangular tooth on the inner
margin near the middle. Head much broader than the thorax,
closely and deeply punctured, the punctures on the front confluent
longitudinally. Antennae inserted near together on the front, fully
twice as far from the eyes as from each other and half as far again
from the anterior ocellus as from the base of the clypeus ; the
second joint of the flagellum a little longer than the third. The
frontal carina is broad at the base of the clypeus, narrow and
424
Mr. lowland E. Turner on
strongly raised between the antennae and obscurely continued to the
anterior ocellus. Eyes diverging towards the clypeus, the ocelli in
a wide triangle, the posterior pair further from the eyes than from
each other. Thorax subopaque, shallowly and sparsely puncturetl,
the scutellum almost smooth. Median segment very sparsely and
finely punctured, the triangular space at the base smooth and shining,
with a faint median sulcus. Pleurae and abdomen deeply but not
very closely punctured, the petiole about twice as broad as long ;
pygidial area rugulose, about twice as long as broad, truncate at the
apex, the sides almost parallel, slightly narrowed at the apex.
Black ; a large spot on each side of the front below the base of the
antennae, the scape beneath and the tibiae beneath yellow ; the fifth
dorsal segment, the sides of the fourth more obscurely and a trans¬
verse spot on each side at the base of the second dark castaneons
brown ; a dull ochraceous spot on the tegulae. Wings hyaline, the
costa as far as the apex of the radial cell rather broadly fuscous ;
nervures black. Length 12 mm.
Hal. Queensland, Cairns {U. C. L. Ferhins).
Cerceris euchroma, sp. nov. (Plate L, fig. 11.)
9 . Clypeus short and broad, the middle lobe twice as broad as
long, porrect and deeply and widely emarginate at the apex, sparsely
and shallowly punctured. Head much broader than the thorax^
closely but not very coarsely piinctured, shallowly on the front, more
deeply on the vertex ; antennae inserted twice as far from the eyes
as from each other, with a sharp longitudinal carina between them,
the second joint of the flagellum as long as tbe first and third
combined. Posterior ocelli fully twice as far from the eyes as from
each other, the cheeks behind the eyes rather broader than the eyes.
Thorax and median segment finelj' and rather sparsely punctured,
shining, scutellum very sparsely punctured, the postscutellum and
the triangular area at the base of the median segment smooth and
shining. Abdomen smooth and shining, the basal segment more
than half as broad again as long ; pygidial area finely granulate,
widest in the middle, a little broader at the base than at the apex ;
sixth ventral segment deeply emarginate at the apex, the angles pro¬
duced and showing as blunt tubercles beyond the dorsal segment.
Pleurae striated, the upper portion of the mesopleui’ae punctured,
the lower portion rugosely striated.
Black ; the base of the mandibles, clypeus, front, scape beneath,
cheeks, a spot on each side on the vertex, prunotum, tegulae,
scutellum, postscutellum, mesopleurae, legs, a broad curved band on •
each side of the median segment almost enclosing a large black spot,
the third and fifth abdominal segments, the basal half of the second
Neiu Fossorial Hymeno'ptera from Australia. 425
dorsal segment and the whole of the two basal ventral segments
orange yellow ; antennae and pygidium dull ferruginous, the apex
of the flagellum fuscous above. Wings hyaline, fusco-hyaline along
the costa, nervures fuscous, the stigma testaceous. Length 11 mm.
Hah. Queensland, Cairns (ex coll. Perkins).
Liris (hicalis, Sm.
Larrada ducalis, Sm., Journ. Linn. Soc. Zool., p. 84, 1861.
Hah. Queensland, Cairns (coll. Perkins).
This wide-ranging species has not previously been
recorded from Australia. It ranges westward as far as
N. India. The Australian specimens are rather small,
about 17 mm. in length, the wings a little paler, and the
anterior margin of the clypeus less deeply notched in the
middle.
Tachysphex adelaidae, sp. nov.
9 . Clypeus shining, very sparsely punctured, the apical margin
very slightly rounded, almost transverse. Antennae inserted a little
nearer to each other than to the eyes, slender, the second joint of the
flagellum about twice as long as the first, but a little shorter than
the third. Eyes separated on the vertex by a distance equal to the
length of the first and second joints of the flagellum combined.
Head smooth and shining, with a very obscure frontal sulcus, with
a large shallow depression behind the posterior ocelli, the eyes
almost touching the posterior margin. Thorax smooth and shining,
the pronotum depressed below the mesonotum, but not very steeply
sloped, the mesonotum without lateral furrows. Median segment
opaque, a little shorter than the mesonotum, the posterior truncation
very finely transversely striated. Abdomen highly polished beneath,
subopaque above, the four basal segments very feebly depressed at
the apex, more broadly in the middle than at the sides ; the pygidial
area shining, very sparsely punctured, narrow and pointed. Comb
of the anterior tarsi fairly long, the spines feeble. Tliird ab.scissa of
the radius longer than the second, equal in length to the fir.st ; the
first recurrent nervure is received nearer to the second than to the
base of the second cubital cell.
Black ; tegulae testaceous brown, mandibles and apex of the
scape beneath fusco-ferruginous ; legs bright ferruginous, coxae and
trochanters black. Wings hyaline, faintly tinged with fuscous,
nervures fuscous. Length 8 mm.
Hob. South Australia, Aclelaide (ex coll. Perkins).
Near T. pugnator, Turn., from which it may be dis¬
tinguished by the colour of the legs, the narrower space
42G
Mr. Rowland E. Turner on
between the eyes on the vertex, the slenderer antennae
and the absence of parapsidal furrows on the mesonotum,
Piagctia intaminata, sp. nov. (Plate L, fig. 14.)
9 . Clypeus short, slightl}’- convex in tlie middle, the apical
margin feebly emarginate, clothed with short whitish pubescence
which extends on to the front. Mandibles notched on the outer
margin. Antennae inserted at the base of the clypeus, much nearer
to each other than to the eyes, the second joint of the flagellum
more than twice as long as the first, the third as long as the second,
the scape nearly as long as the second and third combined. Eyes
converging towards the vertex, separated on the vertex by a distance
exceeding the length of the second joint of the flagellum. Head
opaque, almost smooth, a very delicate frontal sulcus almost reaching
the anterior ocellus ; the space round the ocelli feebly depressed, the
depression almost reaching the eyes. Posterior ocelli close together,
very small. Thorax and abdomen opaque, without visible punc¬
tures, sparsely covered with very fine greyish pubescence, which
is thicker and paler at the apex of the first abdominal segment ;
pronotum much narrower than the mesonotum, faintly transversely
striated ; median segment longer than broad, transversely rugose-
striate. First abdominal segment more than half as long again as
the breadth at the apex, very narrow at the base ; pygidial area
triangular, sparsely punctured, bare. The second abscissa of the
radius is very short, a little more than half as long as the third, the
first as long as the second and third combined ; the first recurrent
nervure is received nearer to the base of the second cubital cell than
to the second recurrent nervure, which is received before the middle
of the same cell.
Black ; the anterior and intermediate tarsi fuscous above, testa¬
ceous beneath ; the tegulae and the fourth and fifth abdominal
segments on the extreme apical margin testaceous brown. Wings
hyaline, with a pale blue flush in some lights, a pale fuscous band
from the base of the radial cell not quite crossing the wing ;
nervures fuscous. Length 10 mm.
Hob. Queensland, Cairns (ex coll, Perkins).
No species of the genus has been previously recorded
from Australia. There is no tubercle on the posterior
femora in this species.
Sphodrotes ruhricaUis, sp. nov. (Plate L, fig. 13.)
9 . Clypeus short, very broadly rounded at the apex, clothed
with silver-white pubescence, the extreme apical margin bare.
Nm Fossorial Hymenoptcra from Australia. 427
Mandibles acute at tlie apex, without a tooth on the inner margin,
deeply notched near the base on the outer margin. Antennae
inserted a little nearer to each other than to the eyes, the second
joint of the flagellum nearly twice as long as the first and a little
longer than the third. Head coarsely and closely, but shallowly,
punctured ; the posterior ocelli a little further from the eyes than
from each other ; the inner margins of the eyes parallel, the front
slightly concave. Thorax closely and rather strongly punctured,
the pronotum transverse, very slightly depressed below the
mesonotum, mesopleurae strongly convex. Median segment very
coarsely longitudinally rugose-striate ; much broader than long,
narrowed posteriorly and vertically truncated, the surface of the
truncation coarsely transversely rugose-striate, .with a deep median
sulcus, the sides of the segment obliquely striated. Abdomen closely
and rather strongly punctured, ovate, broad at the base, the apical
segment pointed, without a pygidial area. Radial cell rounded at
the apex, both recurrent nervures received by the second cubital
cell, the first near the base, the second just before three-quarters
from the base.
Black ; the mandibles, tegulae, legs and abdomen ferruginous
red ; antennae ferruginous, more or less shaded with fuscous. Wings
hyaline, tinged with fuscous, nervures fuscous, the stigma and costa
testaceous. Length 8 mm.
The male differs in having the anterior margin of the clypeus
almost transverse ; the joints of the flagellum shorter, the second no
longer than tlie third ; the whole insect more coarsely punctured
and the apical segments of the abdomen clouded with black.
Hah. S Australia, Adelaide (ex coll. Perldns) ;
8?, 1
Superficially this species closely resembles Pison
imfico^'ne, Sm.
Sphodrotes piloselhis, sp. nov.
Clypeus short and broad, broadly rounded at the apex, finely
punctured and clothed with very short pale golden pubescence which
extends on to the front. Mandibles deeply notched on the outer
margin. Eyes parallel on their inner margin, separated at the base
of the clypeus by a distance equal to twice the length of the scape ;
posterior ocelli as far from each other as from the eyes. Antennae
inserted almost as far from each other as from the eyes, the second
joint of the flagellum no longer than the third. Head, thorax, and
abdomen coarsely and closely punctured ; the dorsal surfaoe of the
median segment coarsely rugose, with a carina from the base not
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) F F
428
Mi^^vowlaad E. Turner on
reaching tlie apex, the sides of the segment striated, the surface of
tlie posterior truncation transversely rugose. The neuration differs
from Unit of <S. jnmctuosus, Kohl, in the position of the first recurrent
nervure which is interstitial with the first transverse cubital nervure.
Black; the legs (except the coxae), tegulae, posterior angles of the
IJronotum, scape and flagellum beneath ferruginous brown ; the first
dorsal segment of the abdomen and the apical margin of the other
segments fusco-ferruginous. Wings hyaline tinged with fuscous,
nervures fuscous, the stigma testaceous. Length 7 mm.
Hab. Queensland, Cairns (ex coll. Perkins).
Extremely near punctitosus, Kohl, from which it may be
separated by the difference in the position of tlie first
recurrent nervure. The relationship of Sphodrotes to
Acanthostethus, Sm., is very close, but in that genus the
first transverse cubital nervure is missing, the angles of
the median segment are produced into stout spines and
the eyes converge towards the clypeus. The relationship
to Sericopliorus, Sm., does not seem to be nearly so close.
Nitcla nigricans, sp. nov.
$ . Clypeus convex in the middle and subcarinate ; antennae
inserted low down on the sides of the cly^jeus, further from each other
than from the eyes, the second joint of the flagellum very little
longer than the first and a little shorter than the third. Posterior
ocelli nearly twice as far from each other as from the eyes, which are
separated on the vertex by a distance equal to the length of the
scape and first joint of the flagellum combined. Head finely
coriaceous, an obscure groove from the anterior ocellus reaching half¬
way to the base of the clypeus. Pronotum transverse, the angles not
prominent, the margins raised, leaving a narrow transverse groove
interrupted in the middle between them. Mesonotum opaque,
without punctures, the scutellum smooth and shining. Median
.segment longer than broad, narrowed towards the apex, longitudi¬
nally striated, with finer and irregular transverse striae forming
reticulations ; the face of the posterior truncation transversely
striated, with a deep median sulcus. Abdomen shorter than the
thorax and median segment combined, smooth and shining, the
segments broadly depressed at the apex. Hindwing almost entirely
without nervures.
Black ; the mandibles fuscous in the middle, spines of the tibiae
pale testaceous. Wings hyaline, nervures fuscous. Length 5 mm.
Hab. Queensland, Bundaberg (ex coll. Perkins).
Very near N. kurandae,T\xxn., but differs in the colour
Trans.ETtt.Soc.Lond.,1910. Plate P.
Horace Knight del.etlith.
West, Newman chr.
NEW FOSSORIAL HYMENOPTERA.
Explanation of Plate L.
Fig. 1. Zasjyilofhynnus ru/ohde'iis. Turn. p. 41
2. Zaspilothy'imus ntfolutens, Turn. $ , p. 41
3. Anthohosca arabica, sp. n., p. 397.
4. Anthohosca jiavopicta, sp. n., p. 399.
.5. Myzine rujinodis, sp. n., p. 392.
6. Myzine nifjrita, sp. n., p. 391.
7. Dielis socotrana, Kirby, p. 402.
8. Parasapycja mdlleri, sp. n., p. 405.
9. Spilomena australis, .sp. n., p. 418.
10. Cerceris victrix, sp. n. , p. 422.
11. Cerceris euchroma, sp. n., p. 424.
12. Cerceris perhinsi, sp. n., p. 423.
13. Sphodrotes rubricatns, sp. n., p. 426.
14. Piacjetm intaminata, sp. n., p. 426.
15. Rhopalosoma abnormis, sp. n., p. 386.
Ncio Fossorial HymcTwptcra from Amtralia. 429
of the legs, antennae and wings and in the coarser
sculpture of the median segment. There is a specimen of
N. kurandae from Bundaberg in the same collection.
Crahro erythrogaster, sp. nov.
^ . Mandibles bidentate at the apex ; clypens broadly rounded at
the apex, with a delicate median carina, clothed with fine silver
pubescence. Eyes separated at the base of the clypeus by a distance
equal to about three-quarters of the length of the scape ; antennae
inserted further from each other than from the eyes, 13 jointed, the
second joint of the flagellum as long as the third and fourth com¬
bined ; the posterior ocelli a little further from each other than the
eyes. Head massive, as broad as the thorax flnely punctured-rugose,
broadly emarginate posteriorly, the eyes separated from the posterior
margin of the head by a distance equal to half the length of the
scape. Thorax rather more coarsely punctured-rngose, the pronotum
very shallowly emarginate anteriorly, the margins raised, the narrow
groove between them coarsely rugose ; mesoplenrae with a vertical
sulcus below the anterior wing, w'ith a shallow striated groove for the
intermediate femora ; median segment rounded at the apex, the
dorsal surface very coarsely rugose-striate, the sides slightly concave
and obliquely striated, the posterior truncation transversely striated.
Abdomen oval, sessile, very closely and minutely punctured. The
recurrent nervure is received just beyond two-thirds from the base
of the cubital cell, the transverse cubital nervure before the middle
of the radial cell.
Black ; the base of the mandibles, scape, base of the flagellum
apex of the femora, tibiae and tarsi brownish yellow ; abdomen
ferruginous red. Wings hyaline, nervures testaceous. Length
6 mm.
Hah. Queensland, Bundaberg (ex coll. Perkins).
Allied to C. doddii. Turn. The groove for the inter¬
mediate femora is very shallow, but the edges are raised,
forming marginal carinae, which give the space between
them an appearance of being depressed more than is really
the case. The structure is similar in C. doddii.
Explanation of Plate L.
[Ncc Explunation facing the Plate.^
F F 2
( 430 )
XV. Descri'ptions of Malayan Micro-Lepidoptera.
By E. Meyrick, B.A., F.R.S., F.Z S.
[Read June 1st, 1910.]
In the' following paper 10 genera and 103 species of
Too'tricina and Tineina from the Malayan region are
described as new. Except where otherwise specified, the
types are in my own collection.
CARPOSINIDAE.
Mcridarchis lembula, n. sp.
$ . 20 mm. Head and thorax whitish-brown sprinkled with dark
fuscous. Palpi very long, porrected, second joint expanded with
rough scales above and beneath, terminal joint thick ; pale brownish
irrorated with dark fuscous, suffused with white above. Abdomen
pale greyish -ochreons. Forewings elongate, rather narrow, some¬
what dilated posteriorly, costa slightly arched, apex round-pointed,
termen sinuate, oblique ; light brownish-ochreous suffusedly mixed
with whitish, with scattered dark fuscous scales ; a suffused brown
patch extending on dorsum from near base to near middle ; six small
fuscous spots on costa between \ and apex ; an irregularly inwards-
curved line of brown and blackish scales running from fourth of
these spots at J of costa to dorsum before tornus : cilia whitish-grey
sprinkled witli fuscous. Hindwings with apex unusually pointed
and prominent ; grey-whitish ; cilia whitisli.
Java, Bandong ; one specimen.
Mcridarchis agycrata, n. sp.
J 9 ■ 22-23. mm. Head, palpi, and thorax brownish mixed with
white and dark fuscous ; palpi in moderately long, curved,
ascending, second joint with appressed scales, terminal joint shorter,
slender, pointed, in 9 very long, arched, porrected, second joint with
scales somewhat roughly expanded, terminal joint moderate. An¬
tennal ciliations in J 3. ' Abdomen whitish-ochreous. Forewings
elongate, narrow, slightly dilated posteriorly, costa hardly arched,
apex obtuse, termen straight, rather oblique ; brown, irregularly
mixed wdth whitish suffusion and sprinkled with black ; six small
spots of blackish irroration on costa between | and apex ; the black
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
Mr. E. Meyrick’s Descriptions of Micro- Lepidoptera. 431
scales appear to form also a very undefined patch in middle of disc
and subterminal fascia, the adjoining areas being more whitish-mixed :
cilia pale brownish, sprinkled wfth whitish points. Hindwings and
cilia whitish.
Java, Bandong; two specimens.
Paramorpha gyritis, n. sp.
(^ . 16-17 mm. Head and thorax ochreous-white. Palpi 2, white,
basal half dark fuscous. Antennal ciliations 5. Abdomen whitish.
Forewings elongate, costa moderately arched, apex obtuse, termen
slightly rounded, oblique ; white, sprinkled with dark grey, towards
dorsum and posteriorly more or less tinged or suffused with light
grey ; a blackish line enclosing an oval spot of ground-colour at base
of costa and continued to dorsum near base ; four large blackish dots
arranged in a curved transverse series beyond J from costa to beneath
submedian fold, three others in a curved series beyond middle from
costa to above fold, and two on transverse vein ; a small dark fuscous
spot on costa at two less marked towards apex, one towards dorsum
at 4) and an undefined cloudy subterminal shade : cilia light grey
irrorated with whitish points. Hindwings and cilia whitish.
Malay States, Gunong Ijan ; two specimens.
Carposina crypsicliola, n. sp.
^ 5 • 15-18 mm. Head and thorax pale fuscous sprinkled with
darker and whitish. Palpi 2f, white sprinkled with grey, basal and
lower longitudinal half dark fuscous. Antennal ciliations of 2.
Abdomen light grey. Fore wings elongate, rather narrow, costa
gently arched, apex obtuse, termen slightly rounded, very oblique ;
light fuscous or brownish, finely and irregularly sprinkled with
whitish and dark fuscous ; costa with about seven small oblique
darker marks ; normal tufts in disc brownish, partially obscurely
edged with whitish, but hardly defined or differing from ground¬
colour : cilia pale fuscous, variably mixed with darker fuscous and
whitish. Hindwings and cilia light grey.
Sumatra, Pura ; four specimens.
TORTRICIDAE.
Procalyptis clielopliora, u. sp.
(J. 19-20 mm. Head and thorax brownish-ochreous. Palpi rather
long, expanded with dense rough projecting scales above, pale ochre-
ous irrorated with brownish. Antennae with a fine horny claw on
432
Mr. E. Meyrick’s Descriptions of
inner side of stalk near base, ciliations 1. Abdomen grey, towards
base and at apex pale oclireous. Forewings suboblong, costa ante¬
riorly gently arched, posteriorly nearly straight, with strong rounded
costal fold from base to near apex obtuse, termen slightly sinuate,
nearly vertical ; brownish-ochreous sutfusedly irrorated with red-
brownish, with some scattered dots or strigulae of blackish scales ;
oblique central fascia and triangular apical patch very faintly suffused
with darker red-brownish, 'hardly traceable ; cilia reddish-brown,
tips ochreous-grey-whitish, beneath tornus with a dark grey spot.
Hindwings pale oclireous, towards apex obscurely suffused with
reddish-fuscous ; cilia ochreous-whitish, with grey subbasal line,
on dorsum and tornus elongated and suffused with pale fulvous-
uchreous.
Malay States, Loeboe Rajah ; in May, three specimens.
Hoinona meneianciy Walk.
(Pandemis menciana, Walk. Cat. xxviii, 310; Godana
simulana, ib. xxxv, 1801 ; Tortrix pullatana, Snell. Tijd. v.
Ent., xliv, 68, pi. v, 3.)
Occurs in India, Java, China, and Japan.
Cacoecia epicyrta, Meyr.
Java, Bandong ; seven specimens, rather smaller and
darker than the Indian form.
Cacoecia secura, n. sp.
5 . ,32 mm. Head, palpi, and thorax orange-fulvous suffused
with lilac-purplish. Abdomen ochreous-yellow. Forewings oblong,
rather broad, costa anteriorly strongly arched, posteriorly hardly
sinuate, apex obtuse, termen concave below apex, rounded-prominent
beneath ; fulvous-orange suffused wdth lilac-purplish. Hindwings
ochreous yellow.
Flores ; one specimen.
POLEMOGRAPTIS, n. g.
Palpi moderate, 'erect, second joint with appressed scales rather
expanded towards apex, terminal joint short. Thorax without crest.
Forewings with 7 separate, to termen. Hindwings wdth 3 and 4
connate, 5 absent, 6 and 7 closely approximated towards base.
Polcmoyraptis miltocosma, n. sp.
9 . 12 mm. Head and thorax ochreous (damaged). Palpi ochre-
ous-orange. Abdomen grey. Forewdngs suboblong, costa moderately
Malayan Micro-Lepidoptera. 433
arched, apex obtuse, termen nearly straight, little oblique ; slaty-
grey; an orange- red stripe along costa throughout, marked with
several dots and groups of black scales and a small slaty -grey spot
near base, low'er edge very irregularly indented, continued as an
irregular streak along termen to tornus ; six oblique transverse red
bars, first almost basal, first five reaching dorsum but not to costal
stripe, sixth confluent with projection of costal stripe, but not reach¬
ing tornal extremity of terminal streak : cilia orange, with a dark
grey spot beneath tornus. Hindwings dark grey ; cilia grey, darker
towards base.
Borneo, Kuching (Hewitt); in February, one specimen,
Attcria stephanitis, n. sp.
^ $ . 30-32 mm. Head, thorax, and abdomen metallic blue-
blackish, face in ^ whitish. Palpi blue-blackish, in ^ wdiitish
internally. Antennae blackish, basal joint in $ whitish anteriorly.
Forewings oblong, costa anteriorly strongly, posteriorly gently
arched, apex obtuse, termen rounded, little oblique ; 7 to apex or
termen ; bright orange ; base narrowly blue-blackish, produced
shortly along costa ; apical area beyond a nearly straight line from
middle of costa to f of dorsum wholly purple-blackish, except a
large transverse-oval orange spot before apex : cilia deep purple.
Hindwings with 3 and 4 connate ; bright orange ; a purple-blackish
streak along dorsum ; apical fifth purple-blackish ; anterior edge
straight ; cilia deep purple, between tornus and apical patch orange.
Flores, in November ; two specimens.
Attcria liclianla, n. sp.
28-29 mm. Head and thorax light orange, face and palpi
purple-blackish. Abdomen indigo-blackish, anal tuft pale greyish-
ochreous. Forewings oblong, costa anteriorly moderately, posteriorly
slightly arched, with strong costal fold reaching from base to |, apex
rounded -obtuse, termen rounded, hardly oblique ; orange, towards
apex broadly suffused with reddish : cilia reddish -orange. Hind¬
wings purple-blackish ; an orange apical patch, occupying nearly
half of wing; cilia dark grey with purple-blackish basal line, round
apical patch orange.
Philippines, Mindoro, at 5000 feet, in November; two
specimens.
Scliocnotenes paraptera, n. sp.
^ 9 • 19-20 mm. Head and tliorax whitish-fuscous or whitish.
Abdomen ochreous- whitish. Fore wings suboblong, rather dilated
434
Mr. E. Meyrick’s Dcscrifjtions of
posteriorly, costa gently arched, apex rounded-obtuse, termen slightly
ro'nnded, somewhat oblique ; cell considerably narrowed posteriorly,
8 to apex, 8-11 very long, curved, parallel ; ochreous-white or
fuscous-whitish, thinly scaled between veins ; costa dotted with
dark fuscous; markings indefinite, tinged with fuscous and edged
with dark fuscous strigulae, viz. traces of edge of basal patch, patches
on middle of costa and dorsum beyond middle indicating central
fascia, a roundish patch in disc at f, a roundish costal patch beyond
I, and transverse streaks from before and beyond tornus : cilia white.
Hindwings thinly scaled, with blue-violet reflections, in whitish,
in $ fuscous-whitish ; cilia concolorous.
Malay States, Gunong Ijan ; India, Khasi Hills ; in
April, two specimens.
EUCOSMIDAE.
S'pilonota thalassitis, n. sp.
g. 15-16 mm. Head white. Antennae with notch at J. Thorax
white, shoulders blackish. Abdomen grey. Forewings elongate,
costa slightly arched, with very broad fold extending from base to
f and reaching half across wing, filled with dense whitish hairs,
apex obtuse, termen slightly sinuate, somewhat oblique ; ochreous-
white; markings fuscous suffusedly mixed with blackish-fuscous;
a large blotch covering costal fold, posteriorly extended to | of costa
and to beneath submedian fold, irregular-edged, connected with a
spot on base of dorsum, a quadrate spot on dorsum before middle,
and a triangular blotch resting on termen from apex to near tornus ;
a triangular spot on dorsum towards tornus, its apex also touching
terminal blotch. Hindwings with 3 and 4 stalked ; fuscous ; cilia
whitish-fuscous, becoming fuscous towards base.
Java, Bandong ; two specimens.
Ilho2)obota 'pythonias, n. sp.
g. 17 mm. Head and thorax pale ochreous, slightly tinged with
reddish and greenish. Palpi pale brownish-ochreous, dense projecting
scales of second joint mixed with dark fuscous suffusion. Abdomen
fuscous. Forewings elongate, co.sta moderately arched, with strong
fold reaching from base to beyond middle, apex obtuse, termen
sinuate, almost vertical ; dark olive-green suffusedly mixed with
black ; costal fold spotted with black ; a pale olive-green sh streak
along dorsum from 1, beyond middle abruptly dilated so as to cover
dorsal half of wing, including before tornus two superposed black
spots surrounded with some darker green suffusion ; a broad bi-
435
Malayan Micro-Lepido'ptera.
sinuate streak of black suffusion running from above middle of
disc to apex, above which is a pale greenish suhoval spot at f ; costa
posteriorly with four pairs of whitish strigulae ; a leaden-metallic
mark on middle of termen : cilia greyish with rows of whitish points,
above apex with a black patch. Hindwings rather dark grey; cilia
grey.
Java, Bandong^ one specimen.
Argyroplocc hranchiodcs, n, sp.
d . 20 mm. Head pale ochreous, lower part of face whitish.
Palpi ascending, pale ochreous, marked with dark fuscous and brown-
reddish, base white. Thorax light brownish-ochreous, collar tinged
with reddish. Abdomen elongate, grey, at base with expansible
lateral tufts of greyish-ochreous hair-scales. Posterior tibiae clothed
above with long rough dense whitish-ochreous scales, with expansible
tuft of very long whitish-ochreous hairs from base above and beneath.
Forewings suboblong, somewhat dilated posteriorly, costa gently
arched, apex rounded, termen rounded, somewhat oblique ; light
brownish-ochreous, more brownish posteriorly, mixed with dark
fuscous except towards costa, disc longitudinally streaked with dark
fuscous; costal edge dark fuscous, with ten pairs of pale strigulae ;
a cloudy whitish-ochreous streak beneath costa from ^ to near apex,
marked posteriorly with four black interneural dashes, sending a
short angulated ochreous-whitish line downwards from its extremity,
preceded by a patch of pale greyish suffusion ; an undefined patch
of pale suffusion towards dorsum beyond middle, whence a cloudy
pale upwards-curved streak runs beneath middle of disc to near
termen, divided by an indistinct dark line; margins of ocellus in¬
dicated by some slight blue-greyish suffusion; termen suffused with
dark fuscous and strigulated with ochreous-whitish : cilia pale
brownish-ochreous barred with brown-reddish mixed with dark
fuscous, with a pale apical patch. Hindwings rather small, dark
fuscous, darker posteriorly ; dorsal area folded, and clothed above
with long pale ochreous hairs ; cilia grey-whitish, with dark grey
subbasal line.
Borneo, Pulo Laut; one specimen.
Argyroploce halixanta, n. sp.
. 18 mm. Head light brownish, back of crown dark fuscous.
Palpi rather short, porrected, liglit brownish. Antennae and thorax
blackish. Abdomen elongate, grey. Posterior tibiae clothed with
long rough whitish hairs above and beneath. Fore wings subtriangular,
436
Mr. E. ]\Ieyrick’s Descriptions of
rather broad, costa strongly arched, apex obtuse, termen rounded,
somewhat oblique ; whitish-ochreous posteriorly suffused with
yellow-brownish ; basal patch large, blackish, its outer edge rather
curved, running from ^ of costa to of dorsum, followed by two
silvery-whitish striae separated by some blackish strigulae ; costa
beyond this with several pairs of short whitish strigulae and some
blackish scales ; a longitudinal linear upwards-angulated leaden grey
mark in disc about f, surrounded with some silvery-whitish suffusion;
a terminal fascia of mixed blue-leaden and blackish marks and
strigulae, broader towards costa, including an oblique 8-shaped
blackish spot above middle, partially edged with whitish : cilia
grey sprinkled with whitish points and barred with blackish.
Hind wings rather small, short, triangular ; rather dark fuscous, base
whitish, subhyaline ; a streak of whitish suffusion along dorsum ;
cilia white, with fuscous subbasal line.
Borneo, Pulo Laut, in June; one specimen.
Arfjyroploce cscharotciy n. sp.
. 21 min. Head and thorax pale ochreous. Palpi rather short,
ascending, pale ochreous, with a few dark fuscous scales. Abdomen
elongate, dark grey. Posterior tibiae clothed above with very long
dense expansible fine white hairs. Forewings elongate, moderately
broad, posteriorly dilated, costa moderately arched, apex obtuse,
termen slightly rounded, nearly vertical ; yellow-ochreous, towards
costa orange-tinged, irregularly marked with interrupted pale
bluish-leaden striae and a few scattered black strigulae ; costa
marked with very small black spots and strigulae, with a larger
semicircular black spot in middle ; basal patch indicated by two
irregular black longitudinal streaks in disc ; a large rounded blackish
pale-edged blotch extending on dorsum from middle to tornus, and
reaching more than half across wing ; an oblique elongate-oval
blackish pale-edged blotch running from f of disc above middle to
middle of termen : cilia ochreous-yellow, basal third irregularly
barred wdth blackish except towards tornus. Hindwings dark
fuscous ; dorsal area somewhat grooved and clothed with long grey
hairs towards tornus ; cilia grey -whitish, with dark grey basal line.
Celebes, in September; one specimen.
Argyroploce tectigera, n. sp.
$ . 16 mm. Head and palpi dark fuscous, palpi moderate,
ascending. Abdomen rather dark fuscous. Posterior tibiae clothed
witli appressed pale greyish-ochreous hairs above, with an expansible
Malayan Micro-LefidofUra. 437
pencil of whitish-ochreous hairs from before middle. Forewings
elongate, posteriorly dilated, costa gently arched, apex obtuse, termen
slightly rounded, hardly oblique ; fuscous, irregularly sprinkled
with w'hitish ; a large elongate- triangular dark fuscous blotch
extending along costa from base to edged with whitish suffusion,
reaching more than half across wing, its apex sending a short pro¬
jection mixed with some black scales to near dorsum at f ; a group
of black scales towards dorsum at I ; costa posteriorly strigulated
with dark fuscous and whitish ; a very small fulvous-ochreous spot
at apex ; margins of ocellus faintly indicated with blue- greyish,
anterior followed by a curved series of four undefined dark brown
dots, posterior followed by an oclireous-fulvous mark on termen :
cilia fuscous mixed with darker, and sprinkled with whitish points.
Hindwings fuscous ; dorsum thickened and clothed with pale
greyish-ochreous hairs ; cilia whitish-fuscous, wdth fuscous subbasal
line.
Java, Bandong ; one specimen.
GELEGHIADAE.
Tisis, Walk,
Head smooth-scaled ; ocelli absent ; tongue developed. Antennae
over 1, in with more or less marked sinuation and thickening of
dense scales near base, in $ more or less roughened with scales
near base, basal joint long, without pecten. Labial palpi very long,
recurved, second joint with appressed scales, terminal joint as long
as second, slender, acute, or in ^ sometimes clothed with dense
rough scales to apex. Maxillary palpi very short, filiform, appressed
to tongue. Posterior tibiae clothed wuth rough hairs, basal joint of
tarsi long, rough-scaled above. Forewings with 2 from angle, 3
absent, 5 absent, 7 and 8 stalked, 7 to apex, 9 sometimes out of 7
11 from middle. Hindwings 1 or over 1, elongate-trapezoidal, apex
obtuse, termen more or less rounded, cilia J-f ; 4 and 5 out of 3, or
4 absent, 6 and 7 stalked.
The generic names Tingentem, Walk., Tipha, Walk.,
Tirallis, Walk., and Cacogamia, Snell., are synonyms of
this. To this genus are referable hicolorella. Walk., melio-
rella. Walk., ^alyhaeella. Walk. (= latifcbsciella. Walk).,
luteella, Snell., elegans, Snell., and helioclina, Meyr., but
the sixteen other species from Ceylon and S. India which
I have referred to Tipha (or Tingentera), Walk., must be
placed in a new genus, for which I propose the name
438 Mr. E. Mey rick’s B&scri^tiom of
Mnesteria, distinguished by the different position of vein 2
of forewings, which rises widely before angle of cell, and
presence of vein 4 ; I make 'pharotrata, Meyr., the type of
this.
Tisis hemixijsta, n. sp.
^ 2 . 18-20 mm. Head and thorax rather dark fuscous, more or
less mixed with light ochreous-yellowish. Palj^i in dark fuscous,
terminal joint clothed throughout with long rough projecting hairs,
in 9 ochreous-yellow, anterior edge of terminal joint dark fuscous.
Antennae dark fuscous, with a white anteapical band, in ^ sinuate
and thickened with dense scales above base. Abdomen dark
fuscous, anal tuft of ^ ochreous-yellowish. Posterior tibiae
ochreous-yellowish, apical half dark fuscous, tarsi white, basal
joint dark fuscous. Forewings elongate, narrow, costa gently
arched, slightly sinuate in middle, apex obtuse, termen very
obliquely rounded ; 9 separate ; dark fuscous, with a bronzy-
purplish tinge ; a transverse orange-streak at I, enlarged on costa
and extended along it to near base ; a transverse orange streak
from dorsum about middle, reaching f across wing ; space between
these two streaks mixed with bright silvery-metallic ; an orange
streak along costa from middle almost to apex ; terminal area tinged
with coppery-metallic : cilia rather dark shining coppery-fuscous.
Hindwings in with costal third thinly clothed with long fine
expansible hairs except towards apex ; 4 absent ; dark fuscous ;
cilia rather dark fuscous. Fore wings in ^ beneath with dorsal on
anterior half surrounded with a fringe of long fine loose hairs.
Borneo, Kuching (Hewitt), in September ; two
specimens.
Tisis eurylampis, n. sp.
$. 17 mm. Head, thorax, and abdomen fuscous. Palpi light
yellowish, anterior edge of terminal joint fuscous. Antennae
bronzy-fuscous, apex (about yV) white. Forewings elongate, rather
narrow, costa gently arched, apex obtuse, termen obliquely rounded ;
2 and 4 stalked, 9 out of 7 near base ; orange ; basal sixth silvery-
metallic-fuscous ; an irregular silvery-metallic submedian spot before
middle ; apical f rather dark shining coppery-fuscous, anterior edge
convex : cilia coppery-fuscous. Hindwings with 4 present ; rather
dark fuscous ; cilia light fuscous.
Borneo, Kuching (Hewitt), in December ; one
specimen.
Malay an Micro- LcpMoptera.
439
Tisis aryyropliam, n. sp.
9. 21 mm. Head, thorax, and abdomen fuscous. Palpi light
yellowish -fuscous (terminal joint broken). Antennae dark purplish-
fuscous, apical half whitish. Forewings elongate, rather narrow',
costa anteriorly moderately, posteriorly slightly arched, apex
rounded-obUise, termen rounded, rather strongly oblique ; 9 sepa¬
rate ; orange ; base narrowly fuscous, tow'ards costa silvery-metallic ;
a silvery-metallic violet-fuscous spot on costa at J, and one on
dorsum opposite ; a large violet-fuscous jjatch occupying apical of
wing except along costa, where ground-colour forms a streak reaching
to near apex, margin suffused : cilia fuscous. Hindwings and cilia
fuscous, slightly ijurple-tinged ; 4 present.
Borneo, Kuching (Hewitt) ; one specimen.
Tisis cliaradraea, n. sp.
^ . 20 mm. Head and thorax dark fuscous mixed with ochreous-
yellowish, face fuscous, sides pale yellowish. Palpi ochreous-yellow,
terminal joint shorter than second, slender, yellow-whitish. An¬
tennae pale yellowish suffused above wdthdark fuscous on basal half,
apical half whitish, towards base sinuate and thickened with scales
above this. Abdomen dark fuscous, beneath ochreous-yellow. Fore¬
wings elongate, rather narrow', costa anteriorly rather strongly,
posteriorly hardly arched, apex obtuse, termen rounded, rather
oblique ; 2 and 4 approximated at base ; deep orange ; a leaden-
metallic streak edged beneath by a black streak extending along
costa almost from base to | ; from extremity of this tw'o broad purple-
blackish streaks run to beyond middle of dorsum and tornus respec¬
tively, confluent above but separated on low'er | by a curved streak
of ground-colour, anterior streak marked with a silvery-metallic line ;
basal area as far as these marked above middle with a black longi¬
tudinal streak, and on dorsal half irregularly mixed with blackish ;
beyond these streaks dorsal | somewhat mixed w'ith dark purple-
fuscous ; a coppery-purple-blue streak along termen : cilia shining
coppery ; undersurface with dorsal t on anterior half stirrounded
with long hairs. Hindwings with 4 present, 5 running to near apex,
costal area reduced ; dark fuscous ; cilia purplish-fuscous.
Malay States, Padang Bengas; one specimen.
Tisis hyacintliina, n. sp.
$ . 13-15 mm. Head deep shining prismatic-blue, collar pos¬
teriorly ochreous-yellowish. Palpi ochreous-yellow, terminal joint
dark fuscous anteriorly. Antennae light yellowish, towards apex
440
Mr. E. Mey rick’s Descriptions of
more wliitish, towards base suffused above with dark fuscous.
Thorax shining indigo-blue, apex of patagia ochreous-yellow.
Abdomen dark fuscoms, apex ochreous-yellow. Posterior tibiae
ochreous-yellow. Forewings elongate, rather narrow, costa sliglitly
arched, apex obtuse, termen very obliquely rounded ; 9 out of 7 ;
deep indigo-blue ; apical | orange, towards apex suffinsed with dark
firscous : cilia pale bronzy-grey. Hindwings with 4 present ; dark
grey ; costa in ^ fringed with fine long hairs towards middle ; cilia
grey.
Borneo, Kuching (Hewitt) ; in February, three speci¬
mens. The second specimen attributed by Walker to his
hicolorella is also an example of this species.
Tisis impcrat'i'ix, n. sp.
9 . 20 mm. Head and thorax shining blue-blackish. Palpi
blackish, second joint externally with purple-blue reflection.s.
Antennae thickened above with rough scales on basal half, deep
purple-blue white from middle to above |, apical portion dark
fuscous. Abdomen purple-blacki.sh. Posterior legs purple-blackish,
three apical joints of tansi white. Forewings elongate, rather narrow,
costa gently arched towards base and apex, slightly sinuate in
middle, apex rounded-obtu.se, termen obliquely rounded ; 9 separate :
bright deep coppery-jmrple, wuth strong indigo-blue reflections ;
cilia coppery-purple. Hindwings and cilia dark purple-fuscous ; 4
present.
Borneo, Kuching (Hewitt) ; one specimen.
Ptilosticha, n. g.
Head smooth-scaled ; ocelli present; tongue developed. Antennae
t, compressed, above clothed throughout with long rough projecting
scales, ba.sal joint moderate. Labial palpi very long, recurved, second
joint with appres.sed scales, terminal joint as long as second, slender,
acute. Maxillary palpi rudimentary. Posterior tibiae clothed with
long hairs above, basal joint of tarsi long, rough-scaled above. Fore¬
wings with 2 and 3 stalked or coincident, 7 and 8 stalked, 7 to costa,
11 from middle. Hindwings rather over or under 1, elongate-
trapezoidal or elongate-ovate, cilia 5- ; 3 and 4 short-stalked, 5 from
above middle of transverse vein, parallel, 6 and 7 long-stalked.
Type cyanop)laca. A curious genus, in some respects
abnormal here, but apparently best regarded as an excep¬
tional form of this group. Snellenia himacidata, Wals., from
Sandakan probably belongs to this genus, and Dasycera (?)
441
Malayan M icro-Lepido2>te7'a.
hernstcmiella, Snell., from Salawatti is also an allied form,
if not generically identical.
PtilosticTia cyanoplaca, n. sp.
9 . 22 mm. Head and thorax indigo-black, sides of collar whitish-
ochreous with a blue gloss. Palpi orange, anterior edge of terminal
joint blackish. Antennae purple-blackish, on apical half with rough
scales ochreons-whitish. Abdomen blacki.sh, with basal and post¬
median orange bands, apex pale ochreous. Posterior tibiae orange.
Forewings elongate, rather narrow, co.sta faintly sinuate in middle,
posteriorly slightly arched, apex obtuse, termen very obliquely
rounded; 2 and 3 stalked; deep orange; an indigo-blackish basal
fascia ; stigmata indigo-blue edged with black, discal forming two
moderately large round approximated .spots, plical forming an elon¬
gate streak from near basal fascia to beneath first discal ; veins 2-8
marked with strong partially confluent purple-blackish streaks,
those on 2-5 suffused together towards base and connected with
second discal stigma, others not reaching cell. Hindwings rather
o\'er 1, blackish ; a very irregular interrupted light ochreous-
yellowish fascia almost at base ; cilia pale ochreous-yellowish,
towards base blackish, round apex and tornus wholly blackish.
Borneo, Kuching (Hewitt) ; in October, one specimen.
Ftilosticlia incandescens, n. sp.
9. 15-17 mm. Head and thorax dark purple- fuscous, shoulders
bright deep purple. Palpi orange, anterior edge of terminal joint
blackish. Antennae dark fuscous, apical third white except tip.
Abdomen bright orange, two apical segments dark fuscou.s. Posterior
tibiae with strong median and apical tuffs above, bright orange,
apical tuft dark fuscous, median spurs white. Forewings elongate,
narrow, costa sinuate, apex obtuse, termen very obliquely rounded ;
3 absent (coincident with 2) ; bright orange ; a black ba.sal patch,
with a deep purple spot on base of costa ; discal stigmata repre¬
sented by two bright blue-pui'ple black-edged spots, second larger ;
a deep purple blackish -edged band extending along termen from
apex to beneath second discal stigma, wdth which it coalesces, crossed
in lower portion by one or two indistinct suffused narrow streaks
of ground-colour : cilia dark fuscous. Hindwings rather under 1 ;
blacki.sh ; a broad orange fascia almost at base, extending to middle
of costa and to near beyond tornus ; cilia dark grey, round dor.sum
and tornus orange.
Borneo, Kuching (Hewitt); in January and February,
four specimens.
442
Mr. K. ]\Ieyrick’s Descriptions of
Gonaepa, Walk.
Distinguished from Crocanthes by the shorter antennae,
not longer than forewings, with very long ciliations in
and presence of ocelli. To this genus is referable heliarcha,
Meyr., described as a Crocanthes, though the differences
are correctly noted.
Gonaepa pyrocliorela, n. sp.
12-14. mm. Head and thorax dark fuscous. Palpi light
ochreous-oraiige, terminal joint dark fuscous anteriorly. Antennae
under 1, fuscous ringed with black, ciliations 5. Abdomen rather
dark fuscous, apex, light ochreous-orange. Forewings elongate,
rather narrow, po.steriorly dilated, costa gently arched, apex obtuse,
termen obliquely rounded ; 2 and 4 short-stalked, 5 approximated,
7 absent, 8 and 9 stalked ; deep reddish-orange ; costal and dorsal
edge and all veins marked by fine black lines, all terminated by a
rather curved black transverse line from costa before J to dorsum
before tornus; discal stigmata represented by small black spots; a
black marginal line round posterior fourth of costa and termen to
near tornus ; cilia dark grey, tips whitish. Hindwings deep reddish-
orange ; all veins mai ked throughout with fine black lines ; a black
discal spot on end of cell; a rather irregular black line from | of
costa to below middle of termen, indented in middle ; a slender
irregular black streak along upper half of termen ; cilia dark grey,
tips whitish.
New Guinea, St. Aignan I., Sudest I., Trobriand Is.,
Kei Is. ; from January to April, seven specimens.
Crocanthes, IMeyr.
I now think it more correct to regard the absent veins
of fore wings as being 3 and 7, instead of 2 and 9.
Crocanthes sidonia, n. sp.
13-14 mm. Head ochreous-whitish, face crimson-tinged. Palpi
crimson, strongly compressed laterally throughout, flat, terminal
joint shorter than second, rather obtiise. Antennae ochreous-whitish,
basal joint crimson above. Thorax bright crimson-rose, anteriorly
ochreous-whitish dor.sally. Abdomen whitish -ochreous. Forewings
elongate, rather narrow, costa gently arched, apex round-pointed,
termen somewhat sinuate, oblique ; 2 and 4 long-stalked, 5 connate,
9 and 10 out of 8 ; pale whitish-ochreous ; markings bright crimson
443
Malayan Micro- Lepicloptera.
rose ; a rather broad costal streak throughout, costal edge posteriorly
and apex yellow ; four irregular transverse fasciae, first basal, second
before middle, connected with first on dorsum by a narrow streak'
third beyond middle, fourth terminal, broader, dilated towards costa,
connected with third by a streak on dorsum ; a streak of dark grey
suffusion from * of costa to middle of termen, sometimes reduced to
a small costal sjrot or wholly obsolete ; cilia deep yellow, sometimes
grey beneath tornus. Hindwings pale yellowish, sometimes more
or less tinged with grey, especially towards apex, termen tinged
with rosy below middle ; cilia light ochreous-yellow.
5. 14-15 mm. Palpi normal, terminal joint whitish. Forewings
almost wholly suffused with crimson, except yellow costal and aj^ical
area ; discal stigmata sometimes indicated by spots of purple suffu¬
sion ; a fascia of grey suffusion from f of costa to tornus, attenuated
towards costa : cilia deep yellow, beneath tornus dark grey. Hind-
wings grey, darker towards apex, sometimes yellowish-tinged ante¬
riorly ; cilia pale yellowish, more or less suffused with grey.
New Guinea, St. Aignan I., Suclest I., Sariba I. (Meok);
Queensland, Cairns (Dodd) ; in April, seven specimens.
Crocanthes teiiieraria, n. sp.
$ . 18 mm. Head whitish, lower part of face crimson-tinged.
Palpi whitish, second joint tinged with crimson. Antennae ochreous-
whitish. Thorax whitish, shoulders crimson. Forewings elongate,
rather narrow, costa gently arched, apex round-pointed, termen
somewhat sinuate, oblique ; 2 and 4 stalked, 5 connate, 9 and 10 out
of 8 ; crimson, dorsal | from i to termen suffu.sed with purplish-grey,
extended indefinitely to costa at I and * ; an irregular yellow- whitish
spot towards dorsum before 1 ; a moderate subquadrate yellow-
whitish spot in middle of disc ; costal edge yellow from | to J, and
more widely suffused w ith deep yellow from | to apex ; cilia deep
yellow'. Hindwings pale yellowish tinged throughout with crimson;
cilia yellow'.
New Guinea, Area R. (Meek); one specimen.
Crocanthes scioxantha, n. sp.
9. 17 mm. Head and thorax ochreous-yellow, face w'hitish.
Palpi very long, ochreou.s-yellow, terminal joint whitish. Abdomen
whitish, marked laterally with ochreous-yellow'. Forewings elon¬
gate, rather narrow, costa gently arched, apex pointed, termen rather
concave, oblique ; 2 and 4 stalked, 5 connate, 9 and 10 out of 8 ;
light yellow, becoming whitish towards costa and termen ; four
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) G G
444
Mr. E. Mey rick’s iJcsciiplions of
irregular deep oclireous-yellow fasciae parallel to termen, occupying
most of wing, first basal, narrowest, fourth subterminal, broadest,
connected with termen by four suffused bars : cilia oclireous-yellow.
Hindwiiigs whitish; three broad deep oclireous-yellow fasciae similar
to last three of forewings, and appearing as continuations of them ;
cilia whitish-yellowish.
New Guinea, Fakfak, 1700 feet, in December; one
specimen.
Crocanthcs thrasydora, n. sp.
^ . 23 mm. Head and thora.\ blackish. Palpi with second and
terminal joints evenly scaled, articulation imperceptible, ochreous-
yellowish, terminal joint sufi'used with blackish, acute. Antennae
blackish, shortly fasciculate-ciliated. Abdomen blackish, segments 2
and 3 orange, next four segments with orange margins. Forewings
elongate, narrow towards base, posteriorly dilated, costa slightly
arched, apex obtuse, termen almost straight, rather oblique ; 2 and
4 short-stalked, 5 approximated, 9 and 10 out of 8 ; purplish -black ;
a broad orange fascia rising from dorsum beyond middle and running
towards f of costa but not reaching it : cilia blackish. Hindwings
purplish-black ; a broad orange fascia from middle of costa to outer
half of dorsum, occupying median third of wing ; cilia blackish.
New Guinea, Kapaur, in January; one specimen.
Crocanthcs carcharias, n. sp.
25 mm. Head and thorax blue-black, face and front of collar
whitish-ochreous. Pal}ii "with second and terminal joints evenly
scaled, articulation imi)erceptible, yellowish, terminal joint mostly
sutl’used with blackish, acute. Antennae moderately ciliated, purple-
blackish, with broad white band above middle. Abdomen orange,
last four segments with purple black-edged margins, anal tuft black¬
ish. Forewings elongate, narrow, somewhat dilated posteriorly, costa
slightly arched, apex obtuse, termen obliquely rounded ; 2 and 4
short-stalked, 9 and 10 out of 8 ; purple-black ; a large irregular-
edged orange patch crossing wing beyond middle, suffusedly con¬
nected with costa but separated from dorsum by a slender streak of
ground-colour, containing a small transverse-oval purple-black spot
(second discal stigma) near its anterior edge in middle ; cilia purple-
blackish. Hindwings orange ; base narrowly black ; an irregular
purple black streak along termen, forming a triangular spot at apex,
a long triangular projection above middle reaching middle of disc,
and a shorter subtriangular projection below middle; cilia blackish.
New Guinea, Fergussou I. (Meek) ; one specimen.
Malayan Micro- Lcpidoptcra.
445
Crocanthes scdetopa, n. sp.
cj. 9-10 mm. Head whitisli-oclireou.s. Palpi whitish-ochreous,
laterally comjjressed, terminal joint thickened with slightly rough
scales anteriorly, jjointed. Antennae twice as long as forewings,
ochreous-whitish. Thorax and abdomen dark fuscous. Forewings
elongate, rather narrow, costa gently arched, apex obtuse, termen
somewhat sinuate, rather oblique ; 4 and 5 out of 2, 9 and 10 out of
8 ; dark fuscous ; two rather oblique wedgeshaped whitisli-ochreous
spots from costa posteriorly, reaching half across wing ; cilia fuscous,
basal half dark fuscous. Hindwings blackish-fuscous ; cloudy light
fuscous suffused transverse streaks before and beyond middle, second
indented in middle ; a sharply-defined sub-metallic prismatic-fuscous
streak from 4 of costa to below apex and continued along termen to
middle; cilia fuscous.
Moluccas, Amboina, in February ; three specimens.
Sarisophoo’a tamiodcs, n. sp.
^ $ . 13-14 mm. Head and thorax yellow-ochreous. Palpi
yellow-ochreous, second joint infuscated or sprinkled with dark
fuscous, terminal joint in ^ longer than second, abruptly bent back,
thickened posteriorly with rather rough scales. Antennae pale
yellowish indistinctly dotted with fuscous, basal third and an apical
band bhackish. Abdomen pale ochreous. Forewings elongate, rather
narrow, costa slightly arched, apex obtuse, termen nearly straight,
rather oblique; 2 and 3 stalked, 7 and 8 stalked, 7 to apex; deep
ochreous-yellow, with some scattered dark fuscous scales, especially
posteriorly ; discal stigmata dark fuscous, from second a more or
less marked dark fuscous line runs directly towards dorsum but not
reaching it : cilia ochreous-yellow. Hindwings in ^ pale ochreous-
yellowi.sh, deeper towards apex, slightly fuscous- tinged on tornal
area ; in 9 grey, apex yellow-tinged ; cilia whitish-yellowish.
New Guinea, Sudest L, St. Aignan I. (Meek) ; four
specimens.
Par elliptic lihrata, n. sp.
9 . 24 mm. Head and thorax whitish-ochreous. Palpi whitish-
ochreous, second joint externally suffused with fuscous, scales some¬
what projecting atajjex beneath, anterior edge of terminal joint dark
fuscou.s. Antennae OA er 1, whitish-ochreous. Abdomen pale greyish-
ochreoms. Forewings elongate, rather narrow, costa moderately
arched, apex obtuse, termen slightly rounded, oblique; 4 and 5
G G 2
446
]\[r. E. Mey rick’s Dcscriptiom^ of
stalked ; wliitish-ochreous, suffusedly ami irregularly irrorated with
fuscous; discal stigmata round, black; an undefined streak of dark
fuscous irroration running from second discal stigma to apex : cilia
whitish-ochreous tinged and sprinkled with fuscous, at apex with a
bar of dark fuscous suffusion. Hindwings with 3 and 5 stalked;
whitish*fuscous, rather darker towards apex ; cilia whitish-fuscous,
with two faint darker lines.
Java, Bandong ; one specimen.
Homaloxestis perichlora, n. sp.
14 mm. Head pale whitish-yellowish, crown centrally suf¬
fused with light fuscous. Palpi yellow-whitish, second joint
externally suffused with fuscous except at apex, anterior edge of
terminal joint dark fuscous. Antennae yellow-whitish. Thorax
fuscous, shoulders pale whitish-yellowish. Abdomen grey, anal tuft
whitish-yellowish. Posterior tibiae whitish-yellowish. Forewings
elongate, slightly dilated posteriorly, costa slightly arched, apex
obtuse, ternien slightly sinuate, oblique ; 4 and .5 approximated,
8 and 9 out of 7, 7 to termen ; rather dark fuscous, faintly purplish-
tinged ; a rather broad whitish-yellowish stri])e along costa from
base to apex, narrowed posteriorly : cilia pale yellow, beneath tornus
fuscous. Hindwings grey ; cilia grey, round apex suffused with
ochreous-whitish.
Borneo, Kuching (Hewitt) ; in December, one speci¬
men.
Frisilia melannrdis, n. sp.
^ $. 21-23 mm. Head and thorax pale yellow-ochreous. Palpi
yellow-ochreous, in d extremely long, mixed with dark grey at
apex, in $ with exterior edge of terminal joint dark fuscous.
Antennae pale yellow-ochreous, with dark fuscous almost apical
band, basal joint tufted above in Abdomen light yellow-
ochreous. Forewings elongate, rather narrow, costa gently arched,
apex obtuse, termen straight, little oblique ; in 2 and 4 stalked,
3 absent, in 9 3 and 4 out of 2, 7 and 8 stalked, 9 absent ;
ochreo US-yellow with some scattered dark fuscous specks; discal
stigmata round, dot-like, blackish ; in ^ an oval glandular swelling
between dorsum and subdonsal groove in middle; a suffused dark
fuscous line along upper part of termen : cilia ochreous-yellow, at
apex with a dark fuscous spot. Hindwings in ^ with large and
long expansible pencil of brownish hairs from base of costa lying
447
Ma lay an M icro-Leindo'jjtri 'a .
beneath forewings; grey; cilia whitish-yellowish, with two faint
fuscous shades, towards tornus more or less suffused with fuscous.
Borneo, Kuching (Hewitt) ; in October, February, and
April, five specimens.
Lecithocera nodosa, n. sp.
18 mm. Head and thorax fuscous mixed with pale ochreous.
Antennae ochreous-whitish. Abdomen pale ochreous-yellowi.sh mixed
with fuscous. Forewings elongate, costa moderately arched, apex
round-pointed, termen concave, little oblique ; 3 and 4 out of 2, 7
absent, 8 and 9 stalked ; dark fuscous ; .some pale ochreons-yellowish
suffusion towards costa near ba.se and before apex ; a pale ochreons-
yellowish median fascia, lower half narrow, upper half enlarged into
a quadrate blotch, lower portion of this blotch marked with a dark
fuscous dot and small transver.se posterior spot ; cilia whitish-
yellowish, on costa barred with dark fuscous, at apex with a dark
fuscous patch, on termen mostly occupied by two broad fuscous
shades. Hindwings dark fuscous; cilia whitish-yellowish , mostly
occupied by two broad fuscous shades.
Malay States, Padang Rengas ; one specimen.
Lecithocera octonias, n. sp.
. 19 mm. Head pale ochreous. Palpi whitish-ochreous, second
joint thickened with somewhat rough scales beneath towards apex,
basal I fuscous, apex more yellowish preceded by a curved fuscous
line, anterior edge of terminal joint dark fuscous. Antennae
fuscous, ciliations 1. Thorax pale ochreous, tinged and sprinkled
with brownish. Abdomen fuscous, anal tuft ochreous-yellowish.
Forewings elongate, rather narrow, costa gently arched, faintly
sinuate in middle, apex obtuse, termen sinuate, little oblique ; 2 and
3 stalked, 4 and 5 closely approximated at base, 8 and 9 out of 7,
7 to termen; rather dark fuscous, anterior g snffusedly mixed
throughout with whitish-ochreous; a straight narrow dark fuscous
fascia at •?, edged posteriorly with light ochreous-yellow ; an 8-
shaped ochreous-yellow discal blotch on end of cell, filled w'ith dark
fuscous ; a slightly incurved indistinct pale ochreous-yellowish sub-
terininal line, enlarged and distinct on co.sta, -where it is preceded by
a dark fuscous spot ; termen suffused with dark fuscous : cilia
ochreous-yellow, at apex with a patch of dark fuscous suffusion.
Hindwings rather dark fuscous; cilia pale ochreous-yellowish, with
two fuscous shades.
Borneo, Kucliing (Hewitt) ; in November, one specimen.
448
Mr. E. Meyrick’s Descriptions^ of
Lecitliucera clcmsfrata, n. sp.
19 nan. Head and thorax fuscous with a faint purplish
gloss. Palpi dark fuscous, second joint smooth-scaled, extreme
apex whitish-ochreous. Antennae dark fuscous, with a narrow whitish
band at apical f white except extreme aj^ex. Abdomen dark
fuscous, with an inflated lobe on each side. Posterior tibiae dark
fuscous, apex ochreoius-whitish. Forewings elongate, rather narrow,
costa gently arched, apex obtuse, termen sinuate, little oblique ;
2 and 3 stalked, 4 and 5 apj^roximated, 7 and 9 stalked, 7 to apex,
8 aV)sent; dark glossy slaty-fuscous; narrow blackish-fuscous fasciae
at f and |, slightly whitish-edged posteriorly ; a transverse-oval
blackish-fuscous spot in disc at f, slightly whiti.sh-edged, containing
some ochreous-yellowish scales towards centre : cilia dark fuscous,
on termen pale yellow-ochreous spotted with dark fuscous suffusion
on basal third. Hindwings and cilia dark fuscous.
Borneo, Kuching (Hewitt) ; in November, one speci¬
men.
Lecithoceni Icucochlura, n. sp.
9 • 16-18 mm. Head, palpi, and thorax white mixed with pale
yellowish. Antennae whitish. Abdomen yellow-whitish. Forewings
elongate, costa gently ai’ched, apex obtuse, termen hardly sinuate,
rather oblique ; 2 and 3 long-stalked, 4 and 5 connate, 8 and 9 out
of 7, 7 to apex ; light oclireous-yellow ; extreme base white ; a
straight white fascia near base ; a straight white fascia before middle,
preceded by a fascia of light brownish suffusion : cilia light ochreous-
yellow. Hindwings pale whitish-yellowish, more whitish towards
base ; cilia pale whitish-yellowish.
Borneo, Kuching (Hewitt) ; in October and November,
two specimens.
Lecithocera hemiacma, n. sp.
(^. 12 mm. Head light brownish, face pale ochreous, sides of
crown ochreous-yellow. Palpi light ochreous-yellowish, terminal
joint little more than half second, anterior edge fuscous. Antennae
pale ochreous-yellowish. Thorax fuscous. Abdomen pale fuscous,
anal tuft ochreous-yellowish. Forewings elungate, posteriorly slightly
dilated, costa slightly arched, apex obtuse, termen slightly rounded,
oblique ; 2 and 3 stalked, 8 and 9 out of 7, 7 to apex ; light bronzy-
fuscous, sprinkled with dark fuscous; discal stigmata cloudy, dark
fuscous, first dot like, second represented by a transverse mark ;
449
Malayan Micro- Lepidoptera.
cilia wliitish-bronzy, with fuscous subbasal shade. Hindwings
'light grey; cilia whitish-grey.
Borneo, Kuching (Hewitt) ; in October, one .specimen.
Specially characterised by the short terminal joint of
palpi.
Lecithocera fausta, n. sp.
26 mm. Head and thorax pale yellow-ochreous, faintly
brownish-tinged. Anten''ae pale ochreous-yellovvish, faintly ringed
with brownish. Fnrewings elongate, posteriorly <lilated, costa
gently arched, faintly sinuate in middle, apex obtuse, termen
sinuate, rather oblique ; 2 and 3 stalked, 8 and 9 out of 7, 7 to
termen; pale ochreous-yellowish, slightly sprinkled with fu.scous
and dark fuscous specks; discal stigmata small, dark fuscous; a
slight rather oblique .shade of dark fuscous irroration between second
and dorsum ; apical margin fuscous : cilia pale ochreous-yellowish,
with traces of two faint brownish shades. Hindwings and cilia
whitish-yellowish ; 4 absent.
Philippines, Luzon, 5000 feet; one specimen.
Lecithocera qiienda, n. sp.
17 mm. Head light yellow-ochreous, centrally tinged with
fuscous. Palpi light yellowish, terminal joint f of second, anterior
edge fuscous. Antennae whitish-ochreous, near base suffu.sed with
dark fuscous above. Thorax light yellow-ochreous tinged with
fuscous. Abdomen whitish-ochreous. Forewings elongate, rather
dilated posteriorly, costa moderately arched, apex obtu.se, termen
straight, rather oblique ; 2 and 3 stalked, 8 and 9 out of 7, 7 to
termen ; pale yellow-ochreous tinged with brownish, basal and
terminal areas and dorsal half irrorated with fuscous and dark
fuscous; discal stigmata moderate, dark fuscous; a slender somewhat
oblique streak of darker irroration from second to dorsum : cilia
light ochreous-yellowish. Hindwings grey ; a loo.se dorsal pencil of
pale ochreous-yellowish hairs from base; cilia whitish-yellowish.
Java, Bandong ; one specimen.
Onehcda inter gnttella, Walk,
(Coydtdia interynttella, Walk. Cat. xxx, 1038.)
Borneo, Kuching, from October to February (Hewitt).
This species, which is the type of Walkers genus Coy-
dalla, agrees in all respects with Onehcda except in having
450
]\Ir. E. Meyrick’s Bmriptimu of
the antennae slightly longer than forewings, whereas in
Onehala they are usually about as long; it cannot he
properly separated from Onehihi.
Braclmiia orhata, n. sp.
5 . 18 mm. Head pale glossy fuscous mixed with ochreous-
wliitisli. Palpi ochveous-wliitisli, basal | of second joint and
anterior edge of terminal joint dark fuscous. Antennae 1, ochret)U.s-
white, towards base spotted with grey. Thorax fuscous. Abdomen
rather dark fuscous, apex mixed with ochreous-yellowish. Posterior
tibiae grey. Forewiiigs elongate, costa gently arched, ajiex round-
jiointed, termen sinuate, obli(|ue; 3 absent, 2 and 4 .stalked, 8
absent, 7 and 9 stalked, 7 to apex ; rather dark fuscous, slightly
purplish- tinged ; terminal edge dark fuscous : cilia fuscous, between
apex and tornus whitish-ochreous becoming white at base. Hind-
Avings rather dark grey ; cilia fuscous-whitish Avith tAvo fuscous
shades.
Borneo, Kuching (HeAvitt) ; one specimen.
Brachmia striyosa, n, sp.
^ 9- 9-11 mm. Head and thorax Avhitish-ochreous tinged with
fuscous. Paljn Avhitish-ochreous sprinkled with dark fuscous,
second joint Avith a subapical ring of dark fuscous irroration, in $
with scales roughly projecting above toAvards apex, terminal joint
in ^ shorter than second, in $ as long. Antennae i, Avhitish-
ochreous, faintly dotted Avith fu.scous. Abdomen greyish, .segmental
margins ochreous-whitish. Posterior tibiae ochreous-whitish. Fore-
Avings elongate, costa gently arched, apex obtuse, termen oblicpiely
rounded ; 2 and 3 stalked, 7 and 8 stalked, 7 to apex, 9 sometimes
out of 7 ; pale yellow-ochreous, sprinkled with ferruginous points,
especially on dorsal half ; a black dot near ba.se toAvards costa,
sometimes one on base of fold, and a small spot on base of dorsum ;
stigmata black, plical someAvhat obliquely before first di.scal ; a
cloudy blackish dot on dorsum beneath second discal ; a roAv of
submarginal black dots round posterior half of costa and termen ;
cilia pale ochreous, sprinkled Avith ferruginous points. llindAvings
and cilia grey.
Borneo, Kuching (Hewitt) ; iu November, two
specimens.
Hylograptis, u. g.
Head Avith appressed scales; ocelli present; tongue developed.
Antennae |, ba.sal joint moderately elongate, Avithout pecten. Labial
451
Malayan Micro-Lepidcyptera.
palpi long, recurved, second joint thickened with dense scales,
forming a long expansible tuft towards apex above, terminal joint
much shorter than second, slender, acute. Maxillary palpi very
short, filiform, appressed to tongue. Posterior tibiae clothed with
rough scales above. Poi’ewings with 2 and 3 stalked, 8 and 9 out
of 7, 7 to apex, 11 from middle. Hind wings over 1, trapezoidal,
termen sinuate, cilia i ; 3 and 4 stalked, .') iiarallel, 6 and 7 stalked.
Allied to Brachmia.
Hylograptis thryptica, n. sp.
9. 18-19 mm. Head and thorax fuscous. Palpi dark fuscous,
terminal joint whitish towards apex. Abdomen grey. Forewings
elongate, rather narrow, costa gently arched towards base, thence
nearly straight, ajiex obtuse, termen nearly straight, somewhat
oblique ; rather light brown ; a dark purplish-fuscous streak along
costa from base to near apex, suffused beneath with indigo-bluish ;
beneath this a deep ferruginous suprainedian streak from ba.se to |,
and a similar rather sliorter submedian streak ; terminal fourth
more or less suffused with purplish and bluish, crossed by a slightly
curved deep ferruginous transverse streak at about f, mixed with
black at costal extremity and on lower half : cilia ferruginous-
ochreous, on tornus suffused with dark fuscous. Hindwings and
cilia rather dark fuscous.
New Guinea, Woodlark I., Sariba I. (Meek) ; two
specimens.
Nothris malacodes, 11, sp.
c? 9 . 10-12 mm. Head and thorax pale ochreous, shoulders dark
fuscous. Palpi w’ith second joint brownish-ochreous, basal half
irrorated with black, apex w'hiti.sh, tuft moderate, triangular, ter¬
minal joint ochreous-whitish with anterior edge dark fuscous.
Antennae ochreous-whitish ringed with dark fuscous, ciliations in
^ 1. Abdomen grey, anal tuft whitish-ochreous. Forewings
elongate, narrow, costa slightly arched, apex round-pointed, termen
very obliquely rounded ; yellow-ochreous, variably spotted and
blotched with fuscous except towards costa anteriorly, e.specially
round stigmata, along dorsum and termen, and towards costa beyond
middle, but these markings are sometimes little developed ; costa
on anterior half dotted with dark fuscous, edge black towards base ;
stigmata black, distinct, plical obliquely before first discal : cilia
yellow-ochreous. Hindwings with G and 7 approximated towards
base ; light grey ; cilia pale ochreous-grey.
452
Mr. E. Moyrick’s DcseriiMons of
Borneo, Kuching (Hewitt); India, Niigiri Hills, 3500
feet (Andrewes), N. Cooig, 3500 feet (Newcome), Cuttack
(F'letcher); Ceylon, Galle (Fletcher), Puttalarn (Pole);
from May to July, and October to December, ten speci¬
mens. Resembles Yi^oloiihits ianthes, but smaller, and
plical stigma before first discal instead of beyond it.
COSMOPTERYGIDAE.
Cosmo-ptcryx toranla, n. sp.
. 8 null. Head lironzy-grey, crown with a central whitisli line.
Palpi white lined with dark fu.scous. Antennae dark fu.scou.s,
tovvard.s ha.se lined with whiti.sh, four apical joints whitish, four
next dai’k fuscoivs, then several alternately white and dark fuscous.
Thorax dark fiiscoins, with three fine whitish lines. Abdomen grey,
anal tuft whitish. Forewings narrowly lanceolate, apex slenderly
long-produced, acute, caudate ; dark fuscous ; three slender white
longitudinal streaks anteriorly, subcostal from base to b not nearly
reaching band, median from ba.se to band, subdorsal from j almost
to band ; costal edge slenderly white from j to band ; a broad orange
transverse band beyond middle, edged with two silvery-white
fasciae slightly converging downwards, anterior followed by a black
dot above middle, posterior preceded by some dark fuscous sca'es
towards extremities ; from middle of posterior fascia a white sinuate
streak, yellow at base, runs along termen to apex : cilia ochreous-
grej'-whitish, Itecomhig grey towards base round apex. Hindwings
grey ; cilia grey-whiti.sh.
Borneo, Kuching (Hewitt) ; one specimen.
Com)iopUryx xuthogastra, n. sp.
,^.8-9 rnm. Head dark bronzy-fuscous with fine white lines
above eyes and in centre of crown. Palpi white lined with dark
fuscous. Antennae black, towards base lined with white, four
apical joints white, then four black, one white, one black, two white.
Thorax blackish, with three fine white lines. Abdomen fulvous-
ochreou.s, segmental margins grey, apex silvery-whitish. Po.sterior
tibiae dark fuscous with white ba.sal, median, and ajiical rings, and
silvery-metallic subapical ring. h'orewings elongate-lanceolate,
apex slenderly long-produced, acute, caudate ; blacki-h ; an oblique
white subcostal dash about and short white median and subdonsal
dashes directly beneath its posterior portion, all widely remote from
base and band ; a broad orange transverse liand beyond middle,
margined anteriorly by a thick golden-metallic vertical fascia, and
458
Malaymi MicTO-Leimlrq)tera.
posteriorly by two golden-metallic spots preceded by a few black
scales, lower spot larger and wholly anterior to upper ; a short fine
violet-white dash in apex, and one between this and band : cilia
grey, suffused with blackish round apex, with a white spot on costal
metallic spot. Hiiidwings dark grey ; cilia grey.
Borneo, Kuching (Hewitt) ; in January, two specimens.
Stagmatophora rotalis, n. sp.
(J. 12 mm. Head, palpi, and antennae whitish. Thorax rather
dark fuscous. Abdomen grey, towards base ochreous tinged. Fore¬
wings elongate-lanceolate, acute ; rather dark fuscous ; a semicircular
white spot on dorsum before middle, reaching nearly half across
wing ; a pale ochreous-yellow semioval blotch extending along
termen from tornus to near apex, marked on tornus with a violet-
golden-metallic dot followed by a black speck, and a smaller golden-
metallic dot beyond this, and receiving on upper edge an oblique
white line from near costa in middle, two pairs of short white direct
streaks from costa posteriorly, and a white line from apex : cilia
whitish-yellowish, beneath tornus grey. Hindwings grey ; cilia
pale greyish.
Borneo, Kuching (Hewitt); in October, one specimen.
Placoptila, Meyr.
Ill the geneiic characters it .should have been stated
that 7 and 8 of forewings are stalked ; G and 7 of hind-
wings are sometimes approximated at base. The real
affinity of the genus is with Limnoccia, of which it is a
development.
Placoptila eyctnolyclina, n. sp.
1^ 9" 11-12 mm. Head and thorax dark glossy leaden-grey; eyes
crimson. Palpi whitish-grey, anterior edge dark fuscous. Antennae
grey. Abdomen dark grey, in ^ with very long dense posteriorly
projecting expansible fuscous hairpencil from above anal segment.
Fore wings elongate-lanceolate, pointed ; black ; base narrowly
leaden-grey ; three narrow transverse prismatic blue-metallic fasciae,
white on costa, at J and before and beyond f respectively, third in
^ interrupted in middle : cilia blackish, round apex white towards
tips, in 9 more broadly. Hindwings with 6 and 7 approximated
at base; in $ blackish above and beneath, in $ dark fuscous;
cilia dark grey, in $ towards base with dense copjiery -blackish
scales, towards middle of costa with an expansible tuft of long
454
Mr. E. Meyrick’s BeMri'ptions of
whitish scale.s, with a very long expansible pencil of brownish hairs
from base of dorsum. Forewings in ^ on undersurface with a
small projecting pencil of hairscales from near below lower angle
of cell.
Bornp:o, Kuching (Hewitt); in February, three speci¬
mens. Very like electrica, but easily distinguished by the
secondary sexual structures of abdomen and hindwings ;
the fasciae are also wider, the second not interrupted, the
white patch in apical cilia more developed, and vein 7 of
l)ind wings separate.
Limnoecia iostrota, n. sp.
$ 9. 14-16 mm. Head and thorax dark jJurple -fuscous. Palpi
grey-whitish, terminal joint much longer than second, dark purple-
fuscous anteriorly. Antennae dark fuscous, with a white band at
about |. Abdomen dark fuscous. Posterior legs shining dark
purple-fuscous, tibiae smooth-scaled, with bristly tufts on origin of
spurs, spurs rough-scaled, two apical joints of tarsi white. Fore¬
wings elongate-lanceolate, pointed ; dark bronzy-fuscous ; a deep
blue-purple basal patch and median fascia ; a slender white fascia
at ^ ; a rather incurved inwardly oblique silvery-white streak from
costa at I, terminating in a deep bine-purple spot on tornus, pre¬
ceding which is a similar spot sometimes surmounted by a white
mark : cilia dark bronzy-fuscous, becoming white towards tips.
Hindwings and cilia dark fuscous.
Borneo, Kuching, from November to January (Hewitt);
India, N. Coorg, 3500 feet, in May (Newcome) ; five
specimens.
Stathviopoda plinthiota, n. sp.
8 mm. Head and palpi white, crown pale ochreous. Thorax
rather dark fuscous marked with white, shoulders pale ochreous.
Forewings narrow, widest near base, thence narrowed to acute
apex ; dark brown ; a shining white fascia about b c>n costa
moderate, widely dilated downwards so as to extend on dorsum from
near base to near middle of wing, anterior edge with a prominence
in middle ; a rather broad light yellowish fascia about f, narrowed
downwards ; apex suffused with light yellowish : cilia light greyish-
ochreous. Hindwings grey ; cilia light ochreoiis-grey.
Borneo, Kuching (Hewitt) ; in April, one specimen.
ErctmoceTci cyanosoma, n. sp.
12 mm. Head, palpi, antennae, and thorax purple-blackish.
Aljdomen deei^ iudigo-blue, anal tuft blackish, beneath with yellow
455
Malayan Micro-L&pido'pUra.
subbasal and postiuedian bands. Forewings elongate-lanceolate,
pointed ; indigo-blackish ; a pale yellow spot in disc at one on
dorsum towards tornus, and one on costa at f ; cilia dark fuscous.
Hindwings dark purple-fuscous ; cilia dark fuscous.
SuMBA ; in February, one specimen.
Eretmocera xanthonota, n. sp.
$ . 16-17 mm. Head, palpi, antennae, and thora.x: purple-blackish,
base of palpi yellowish, thorax with two small orange spots pos¬
teriorly. Abdomen orange, with subdorsal purple-blackish stripes
not extended over tw'o praeanal segments, anal tuft blackish, beneath
orange. Forewings elongate-lanceolate, pointed ; very deep purple ;
four ochreous-yellow or orange spots, first on dorsum at i of wing,
small, others moderately large, roundish, in disc at |, on tornus, and
on costa at * resjDectively ; cilia dark fuscous, base purplish. Hind-
wings dark purple-fuscous ; cilia dark fuscoirs.
SuMBA ; in February, three specimens.
OECOPHORIDAE.
Ncphogenes thctopa, n. sp.
^ $ . 16-18 mm. Head whitish-ochreous. Palpi ochreous- whitish,
irrorated with dark fuscous except ajjex of second joint. Thorax
whitish-ochreous sprinkled with fuscous. Abdomen grey, segmental
margins whitish-ochreous. Forewings elongate, rather narrow,
costa gently arched, apex obtuse, termen very obliquely rounded ;
whitish-ochreous sprinkled with fuscous and dark fuscous ; stigmata
blackish, plical obliquely beyond first discal, second discal connected
with tornus by an oblique streak of dark fuscous irroration; a
curved line of dark fuscous suffusion from | of costa to tornus,
sharply indented above middle ; cilia whitish-ochreous, sprinkled
with fuscous points. Hindwings grey ; cilia whitish-ochreous more
or less tinged with grey.
Celebes, Bonthain Peak; three specimens.
Periacma macra, n. sp.
$ . 10-12 mm. Head and thorax whitish-ochreous, more or
less sprinkled irregularly with dark fuscous. Palpi whitish-
ochreous, second joint with three dark fuscous bands, terminal joint
with dark fuscous median band. Abdomen grey. Fore wings elon¬
gate, narrow, costa slightly arched, apex acute, termen extremely
oblique ; whitish-ochreous, with some scattered fuscous scales ;
•ioO Mr. E. Mey rick’s Desco'iptions of
seven small black spots or marks on costa, last two sometimes ob.so-
lete ; some irregular fuscous suffitftion forming an undefined broad
fascia about and costal jDatch about middle ; stigmata black, plical
forming a dash beneath first discal ; a series of irregular suffused
blackish dots along termen ; cilia dark grey, on costa whitish-
ochreous. Hindwings ovate-lanceolate, acute, grey ; cilia grey.
Borneo, Kuching, in February (Hewitt); Ceylon,
Maskeliya, Matale, in August and October (Pole, Alston) ;
three specimens.
Cvyptolecliia aridula, n. sp.
d- 10-11 mm. Head and thorax pale ochreous sprinkled with
fuscous, shoulders dark fuscous. Palpi whitish-ochreous, second
joint mixed with dark fuscous. Abdomen grey, anal tuft whitish-
ochreous. Forewings elongate, rather narrow, costa gently arched,
apex round-pointed, termen extremely obliquely rounded ; 7 to
costa ; pale ochreous sprinkled with fuscous and dark fuscous ; a
small dark fuscous spot on base of costa ; a small dark fuscous spot
on middle of costa, and a dark fuscous spot at f , connected by fuscous
sufl'usion ; stigmata dark fuscous, plical rather obliquely before first
discal ; a moderate dark fuscous terminal fascia from apex to tornus,
anterior edge rather excavated in middle : cilia fuscous mixed
with dark fuscous. Plindwings narrowly pointed, grey ; cilia pale
grey.
Borneo, Kuching, in February and April (Hewitt);
two specimens.
Cryptolechia laica, n. sp.
9 . 16 mm. Head and thorax browuish-ochreous, shoulders dark
fuscous. Palpi light ochreous-yellowish, second joint sprinkled with
dark fuscous. Abdomen grey. Forewings elongate, costa moder¬
ately arched, apex obtuse, termen slightly rounded, rather strongly
oblique ; 7 to apex ; brownish-ochreous, sprinkled with dark fuscous ;
base of costa sufl’used with dark fuscous ; stigmata dark fuscous,
first discal rather large, plical rather obliquely beyond first discal,
an additional dot beneath second discal ; a series of dark fuscous
dots along posterior half of costa and termen; cilia pale ochreous.
Hindwings fuscous ; cilia pale fuscous.
Borneo, Kuching, in November (Hewitt); one
specimen.
2'onica scncscens, n. sp.
^ . 20-22 mm. Head and thorax wliitish-ochreous mixed with
browmish. Palpi ochreous-whitish, secOnd joint sprinkled with dark
457
Malayan Micro- Lcpido2')tera.
fuscous, terminal joint with two dark fuscous rings. Ab loinen grey.
Forewings suboblong, moderate, costa gently arched, concave be¬
tween I and f, apex obtuse, termen nearly straight, somewhat
oblique ; wliitish-ochreous mixed or suffused with light brownish,
more or less strewn with blackish scales tending to be arranged in
longitudinal lines; stigmata represented by tufts of mixed brownish
and whitish-ochreous scales ; prominences of costa marked with
similar tufts ; an angulated transverse series of small tufts about i, and
others along termen : cilia whitish-ochreous, round apex curled and
suffused with brownish. Hindwings grey ; cilia pale grey.
New Guinea, Sariba L, Kei Is., in October; two
specimens.
XYLORYCTIDAE.
Cryptophasa megalorma, n. sp.
5. 49 mm. Head, palpi, and thorax ochreous-wliitish ; palpi
wdth lower ^ of second joint externally dark fuscous ; thorax with
anterior and posterior spots of fuscous suffusion. Abdomen ochre-
ous-whilish, on dorsum with broad almost confluent bands of dark
grey suffusion, with almost concealed subbasal dark red-brown
band. Forewings elongate, moderately broad, costa gently arched,
apex obtuse, termen somewhat obliquely rounded ; ochreous-w’hitish,
suffusedly irrorated with fuscous except tow’ards costa and in middle
of disc ; base of costa blackish ; stigmata forming rather large black
spots, approximated, first discal oblique-oval, second transverse-oval,
plical between these two, round ; two or three streaks of blackish
suffusion on veins about | ; four small round black spots on jjosterior
half of costa, and a row- of large black dots along termen, becoming
larger round tornus : cilia fuscous-whitish, suffused wdth fuscous
on a subbasal shade and posterior half. Hindwings fuscous, paler
towards costa ; a suffused blackish dot in middle of disc : a series of
small round suffused blackish spots round apex and termen ; cilia as
in forewings.
New Guinea, Ron Island, in July (Doherty); one
specimen. Type in Coll. R)t!ischild.
Athrypsiastis, n. g.
Head with loosely appressed hairs ; ocelli present ; tongue de¬
veloped. Antennae f, in ^ bipectinated, towards apex simple,
basal joint moderate, without pecten. Labial palpi very long,
recurved, second joint with appressed scales, terminal joint somewhat
shorter than second, slender, acute. Maxillary palpi very short
458
Mr. E. IMeyrick’s DescriitfAons of
filiform, appressed to tongue. Forewings with 2 from 3 and 4
stalked, 8 and 9 out of 7, 9 to apex, 11 from beyond middle.
Hindwings over 1, ovate, cilia \ ; 3 and 4 short-stalked, 5-7 nearly
parallel, 7 to termen.
Allied to Ptochoryctis, but more primitive in character ;
the termination of vein 9 of forewings in apex is very
exceptional.
Athrypsiastis lAiaeoleuca, n. sp.
1^. 17 mm. Head and thorax whitish-brownish. Palpi and an¬
tennae pale brownish. Abdomen pale ochreous. Forewings rather
elongate-triangular, costa gently arched, apex obtuse, termen slightly
rounded, rather oblique ; pale ochreous-brown ; cilia pale ochreous.
Hindwings and cilia white.
New Guinea, Humboldt Bay, in October ; one
specimen.
Linoclostis musicocles, n. sp.
^ 9 • 11-13 mm. Head, palpi, thorax, and abdomen white. An¬
tennae in ^ black, in 5 grey. Forewings elongate, moderate, costa
moderately arched, apex obtuse, termen obliquely rounded ; 4 absent
(coincident with 3) ; white ; a fuscous line from | of costa to tornus,
right-angled in middle, dark fuscous towards extremities ; space
between this and termen more or less tinged with fuscous ; a dark
fuscous line round posterior sixth of costa and termen to near
tornus : cilia white, round apex with a fine dark fuscous ante-
median line, beyond this beneath apex with two fuscous bars,
between which the line is interrupted. Hindwings whitish-grey ;
cilia white.
Borneo, Kuching (Hewitt), in Apiil and August ; two
specimens.
Epimactis mffiisella, Walk.
{Gryptolcchia suffmella, Walk. Cat. xxix, 748.)
Borneo, Kuching; India, Khasi Hills. This species is
very similar to Odites paracyrta, Meyr., from Ceylon.
Ep)imactis crainhalea, n. sp.
16 mm. Head pale ochreous. Palpi ochreoiis-whitish,
basal I of second joint dark fuscous. Antennae ochreous-whitish,
simple. Thorax pale ochreous, dorsally suffused with fuscous.
459
Malayan Micro-Lepidoptcra.
Abdomen grey, anal tuft pale ochreous. Forewings elongate, pos¬
teriorly slightly dilated, costa gently arched, apex obtuse, termen
obliquely rounded ; 7 to termen ; brownish-ochreous, along costa
pale ochreous, towards dorsum infuscated, especially anteriorly ;
second discal stigma rather large, dark fuscous ; cilia pale ochreous.
Hind wings with 3 and 5 short-stalked ; grey, thinly scaled towards
base ; a subdorsal groove filled with long hairs ; cilia pale grey,
round tornus strongly elongated.
Java, Bandong ; one specimen.
STENOMIDAE.
Bassarodes, n. g.
Head with appressed scales, side-tufts raised, rough, expanded
behind ; ocelli absent ; tongue developed. Antennae f , in ^ very
shortly ciliated, basal joint moderately elongate, without pecten.
Labial palpi very long, recurved, second joint thickened with dense
scales, somewhat rough beneath, especially in (J, terminal joint as
long as second, moderate, acute. Maxillary palpi very short, fili¬
form, appressed to tongue. Posterior tibiae with dense rough hairs
above. Forewings with 2 from towards angle, 3 and 4 approxi¬
mated, 8 to apex, 11 from before middle. Hind wings 1, oblong-
ovate, cilia 4 ; 3 and 4 stalked, 5 connate, 6 and 7 parallel, 7 to
apex.
Bassarodes siriaca, n. sp.
^ 9 • 23-30 mm. Head light ochreous-yellowish, tufts of crown
suffused with dark brown. Palpi ochreous suffusedly mixed with
dark brown. Thorax pale ochreous-yellowish, in ^ with central
stripe of brown suffusion, in 5 wholly suffused with brownish.
Abdomen pale ochreous-yellowish, in $ suffused with brownish.
Forewings elongate, moderate, posteriorly dilated, costa gently
arched, apex rounded- obtuse, termen slightly rounded, little oblique ;
in pale yellowish, sometimes suffused with pale brownish on veins
and margins, with .second discal stigma dark brown, an oblique
patch of brown suffusion between this and apex, and in one speci¬
men a streak of dark brown suffusion along fold curved upwards
beneath middle of wing ; in 9 ochreous suffusedly irrorated
with dark brown, with a broad longitudinal median dark fuscous
band, suffused and undefined towards costa, lower margin well-
marked and pale-edged, curved downwards on anterior half and up¬
wards beyond middle, terminating in tornus : cilia pale yellowish,
in sometimes suffused with light brownish, in ^ suffused with
TRANS. ENT. SOC. LOND. ] 910. — PART IV. (DEC.) H H
460
Mr. E. Meyrick’s Descriptions of
fuscous. Hinclwings in $ light yellowish, sometimes sprinkled
with fuscous towards tornus, cilia concolorous ; in 5 fuscous, cilia
whitish-ochreous suffused with fuscous.
Solomon Is., Cboiseul, New Georgia; three specimens.
Stcnoma 2)hasriiatopa, n. sp.
1^. 18-20 mm. Head and thorax ochreous-white. Palpi rather
stout, white, lower half of second joint dark fuscous. Antennal
ciliations 4. Abdomen hairy above towards base, white, more or
less tinged with light brownish. Forewings elongate, moderate,
somewhat dilated posteriorly, costa gently arched, apex rounded,
termen somewhat rounded, hardly oblique; ochreous-white; mark¬
ings light fuscous irrorated with dark fuscous ; a small spot on base
of costa, and two oblique strigulae on costa towards base ; an
elongate mark on fold near base ; plical and first discal stigmata
small, plical very obliquely beyond first discal ; second discal repre¬
sented by an irregular spot, connected with middle of costa by an
oblique streak ; an oblique strigula from costa at f ; a curved line
near before termen, sometimes interrupted into three segmenis :
cilia whitish, basal half faintly barred with brownish. Hindwings
and cilia ochreous white.
Solomon Is., New Georgia, Gizo (Meek); three
specimens.
ETHMIADAE.
Ethmia praeclar a, n. sp.
$ . 33-37 mm. Head grey, with three black dots on base of
collar. Palpi rather short, especially in grey-whitish, second
and terminal joints each with black subbasal and subapical bands,
subapical of second incomplete anteriorly. Antennal ciliations in
J. Thorax grey with six black dots. Abdomen orange, with
round black spot on each segment (seven), beneath with two series
of similar spots. Forewings elongate, rather narrow, costa gently
arched, apex obtuse, termen faintly sinuate, slightly oblique ; grey ;
about nineteen moderate black dots, viz. one in middle of base, two
subcostal and median near base, one subcostal at i, one towards
dorsum beneath this, one on fold beyond these, two discal and sub¬
dorsal before middle, one on fold beyond middle, one towards costa
at I, one below middle beyond this, one towards costa at I, one in
middle beneath this, three approximated in a transver.se series
below middle beyond this (two upper sometimes confluent), and
three in a curved transverse series before apex ; eleven black dots
461
Malayan Micro- Lepidoytera.
round posterior part of costa and termen : cilia dark grey, becoming
whitish towards tips. Hindwings orange ; a blackish triangular
apical patch covering \ of wing, lower angle slightly produced on
termen; cilia yellow, base orange, round apical patch dark grey
with whitish tips.
Lombok, 1 Queensland (Mackay), 1 exactly
similar and I think certainly identical.
Etlimia ij ullata, n. sp.
9 . 30-34 mm. Differs from praeclara only as follows ; palpi
longer; abdomen with dorsal spots absent on two basal segments,
and very small or absent on third ; forewings rather shorter and
broader, marginal dots ten in number and more quadrate, tips of
cilia not whitish ; hindwings with apical patch much larger, occu¬
pying ^ of wing, triangularly prominent near costa, extended on
termen as a narrow streak to middle, on undersurface with a dark
fuscous costal band throughout.
Solomon Is., Guadalcanal' (Meek) ; four specimens.
This species is at once distinguished from all the similar
species by the dark fuscous costal band of undersurface
of hindwings, this area being usually orange.
Ethmia argopa, n. sp.
9 . 32-34 mm. Differs from praeclara only as follows : subbasal
band of second joint of palpi incomplete anteriorly : abdomen with
dorsal spot absent on basal segment, very small or absent on second ;
forewings somewhat broader, black dots larger, lowest of three
approximated dots towards termen below middle absent, two lower
of praeapical dots coalesced, marginal dots nine in number, cilia
paler grey ; hindwings ochreous-yellow, cilia pale yellow, round
apical patch pale grey.
Malay States, Padang Rengas; two specimens. It
seems convenient to describe here also the following
closely-allied Australian species.
Ethmia thoraea, n. sp.
. 27 mm. Differs from praeclara only as follows : head whitish-
grey with black spot behind each antenna, base of collar with four
subconfluent black dots ; palpi longer ; antennal ciliations | ; thorax
whitish-grey, black dots larger : forewings rather shorter, costa more
arched, termen more oblique, whitish-grey, black dots larger, post¬
median subcostal placed at § instead of |, lowest of three approxi-
H H 2
462
Mr. E. Meyrick’s Dcscrij^tionfi of
mated dots towards termen below middle absent, two lower of
praeapical dots coalesced, marginal dots ten in number.
Queensland (Turner) ; one specimen.
Ethmia colonella, Wals.
Distinguished by its small size, relatively long antennal ciliations
of ^ (1), second joint of palpi with subapical ring obsolete, and form
of apical patch of hindwings, which sends a triangular projection
inwards to end of cell.
New Guinea, Kei Is.
GLYPHIPTERYGIDAE.
Tortyra heryllitis, n. sp.
(J. 17 mm. Head blackish, face metallic blue-green. Palpi
metallic blue-green, towards base pale j’ellowish, towards apex black.
Abdomen dark fuscous. Forewings rather elongate-triangular, costa
slightly arched, apex obtuse, termen rather obliquely rounded ; dark
fuscous-bronze ; a bright metallic -green basal patch, outer edge
running from \ of costa to f of dorsum, straight ; a slender bright
metallic-green fascia near beyond and parellel to this ; near beyond
this a blackish transverse line edged anteriorly with blue-purple ;
whole posterior area beyond this strewn with coppery-golden and
blackish scales, towards preceding line tinged with bright purple :
cilia silver-grey, with blackish basal line. Hindwings with tornus
not produced ; dark fuscous ; a slender indistinct whitish-ochreous
streak beneath lower margin of cell ; cilia grey, with dark fuscous
basal line, outer half white.
Nicobar Is. ; one specimen.
Tortyra cypridia, n. sp.
9 . 15 mm. Head and palpi shining dark prismatic-bronze.
Antennae prismatic purple-blackish, white above on a subapical
band. Thorax dark shining bronze, with three metallic green-blue
lines. Abdomen dark fuscous. Forewings elongate-triangular, costa
posteriorly gently arched, apex obtuse, termen rounded, rather ob¬
lique ; dark fuscous-bronze ; metallic green-blue subcostal, median,
and subdorsal streaks from base to beyond ^ ; a transverse metallic
green-blue line before middle, interriipted in middle ; a narrow
blackish fascia beyond middle, edged anteriorly by a crimson-purple
line, and posteriorly by bright prismatic crimson-purple costal,
discal, and dorsal patches, costal partly suffused with metallic-green ;
463
Ma I ay an Micro-Le23id oiJtera .
a bright prismatic crimson-purple submarginal line round posterior
part of costa and termen, area between this and preceding markings
mixed with dark fuscous and irrorated with ochreous-whitish ; cilia
silvery-grey-whitish, with blackish basal line. Hindwings dark
fuscous ; cilia grey, with blackish basal line, outer half whitish.
Borneo, Kuching (Hewitt) ; in February, one specimen.
Tortyra mitrifnga, n. sp.
(^. 16-17 mm. Head shining blue-blackish, upper half of face
metallic blue-green. Palpi greenish-bronze, becoming black towards
apex. Antennae purple-black, above with a white band above
middle. Thorax blackish, with three metallic-green stripes. Abdo¬
men dark fuscous. Forewings elongate-triangular, costa posteriorly
moderately arched, apex obtuse, termen rounded, somewhat oblique ;
blackish ; a metallic-green subcostal streak from base to I, and a
triangular patch formed by confluence of usual median and sub¬
dorsal streaks ; a metallic-green ti'ansverse streak at | ; a pale
metallic green-blue transverse line before middle, edged with dark
purple-fuscous suffusion ; beyond this a large circular patch of
suffused ochreous-whitish irroration enclosing a suffused dark purple-
fuscous spot marked with some golden-metallic scales, rest of wing
posteriorly sprinkled with ochreous-whitish ; a golden-metallic
streak along upper part of termen ; cilia submetallic-grey, with
blackish subbasal line. Hindwings with tornus not produced ; dark
fuscous ; a suffused narrow ochreous-whitish or pale yellowish streak
beneath lower margin of cell ; a hyaline dorsal streak ; cilia whitish,
with dark fuscous subbasal line.
Celebes (Doherty) ; India, Khasi Hills, in November
and December; three specimens.
Tortyra lihanota, n. sp.
19 mm. Head metallic-green. Palpi metallic-green, becoming
silvery-white towards base and black towards apex. Antennae
purple-black, above with a narrow white band about |, five apical
joints ochreous. Thorax blackish, with three metallic-green stripes.
Abdomen dark fuscous. Forewings rather elongate-triangular, costa
gently arched, apex obtuse, termen obliquely rounded ; deep golden-
bronze ; a purple-blackish basal patch reaching to marked with
metallic-green subcostal, median, and subdorsal streaks, two latter
broader and confluent posteriorly ; a metallic-green transverse streak
at I ; a black slightly sinuate transverse median streak, broader
towards costa, edged anteriorly with a blue-leaden line and pos-
4C4 A[r. E. Meyrick’s Bescrvptions of
teriorly with a golden-metallic line ; whole posterior area beyond
this irregularly mixed with purple-blackish and strewn with scattered
golden-metallic specks, which i)Osteriorly tend to be arranged in
longitudinal lines : cilia light shining bronze, with black basal line.
Hindwings broader and shorter than usual, termen rather promi¬
nently rounded towards middle and sinuate above this ; dark
fuscous; a slender suffused pale ochreous-yellowish streak beneath
lower margin of cell ; a subhyaline dorsal streak ; cilia grey, tips
whitish.
New Guinea, Woodlark I.; in March, one specimen.
Tortyra 'prodigella. Walk.
Java; Celebes. Snellen’s siriana is a synonym of
this, and his basalis is a synonym of pretiosa, Walk. ; but
the Australian species described by me under the name
of ijrodigella. Walk., is really a distinct species renamed
below.
Tortyra exanthista, n. sp.
(Tortyra vrodiqella, Meyr. (nec Walk.), Proc. Linn. Soc.
N.S. Wales, 1907, 100.)
Queensland, Cairns. Sufficiently described.
Imma denticulata, n. sp.
^ $ . 18-19 mm. Head, palpi, antennae, thorax, and abdomen
fuscous (palpi in ^ broken). Forewings elongate, posteriorly dilated,
costa slightly arched, apex obtuse, termen rounded, slightly oblique ;
7 and 8 stalked, 8 to termen ; rather dark fuscous ; extreme costal
edge ochreous-whitish ; a suffused indistinct darker fuscous mark
on transverse vein ; a very small ochreous-whitish triangular spot
on costa near apex : cilia fuscous, tips whitish. Hindwings in ^
fuscous, darker posteriorly, towards base thinly scaled, with broad
hyaline median and subdorsal streaks from base to about | ; in 5
dark fuscous, on basal half thinly scaled with broad median and
subdorsal streaks subhyaline and mixed with whitish ; cilia in ^
fuscous becoming whitish towards tips, in $ whitish with dark
fuscous basal line.
Timor, Dilli, in May, 1 J (type) ; Borneo, Sandakan, in
February, 1 Malay States, Penang I., in November*
1 ?■
Malayan Micro- Lepido'ptera.
465
Imma uranitis, n. sp.
^ . 25-26 mm. Head light fuscous mixed with whitish-ochreous,
lower part of face whitish-yellowish, collar with series of broad
erectible prismatic scales. Palpi with second int excurved, with
expansible hairs towards apex above, light ochreous-yellowish with
broad anterior and short narrow posterior dark purple-fuscous
stripes, terminal joint dark fuscous. Antennal ciliations 1. Thorax
lilac-fuscous, beneath forewings with a long expansible lateral pencil
of whitish-ochreous hairs, covered at base with broad pearly-white
scales. Abdomen light bluish-lilac-fuscous, rough-scaled above. Fore¬
wings elongate, rather broad, posteriorly dilated, costa gently arched,
apex rounded-obtuse, termen rounded, little oblique; 7 and 8 stalked,
8 to termen ; dark fuscous, slightly purplish-tinged ; a moderate
ochreous-yellow rather oblique median fascia ; a triangular ochreous-
yellow spot on costa before apex ; a very fine irregular ochreous-
whitish line along upper part of termen : cilia fuscous, becoming
whitish towards tips, or mostly ochreous-whitish. Hindwings dark
fuscous ; a large undefined lighter patch occupying most of disc,
with strong bright blue reflections ; cilia fuscous, becoming whitish
towards tips, with dark fuscous subbasal line.
Celebes, Tawaya, Dongala ; in August and September,
two specimens.
Imma monastica, n. sp.
^ $. 17-21 mm. Head and thorax fuscous. Palpi fuscous, in
^ normal. Antennal ciliations 1. Abdomen dark fuscous, in ^
laterally clothed with rough hairs on basal half. Posterior tibiae
tufted with long dense expansible ochreous-whitish hairs on basal
half beneath. Forewings rather elongate-triangular, costa in ^
nearly straight, in $ slightly arched, apex rounded, termen rounded,
somewhat oblique ; 7 and 8 stalked, 8 to termen ; dark fuscous,
faintly purplish-tinged ; a wedge-shaped ochreous-white mark from
costa near apex ; a fine white denticulate terminal line : cilia fuscous,
tips whitish, above tornus with a suffused whitish spot. Hindwings
dark fuscous, lighter anteriorly ; broad median and dorsal prismatic-
violet-tinged hyaline longitudinal patches, confluent towards base,
upper extending to about f, lower nearly to termen ; cilia fuscous,
outer half whitish.
Moluccas, Amboina, in August and February (Doherty);
twelve specimens. Near airosignata, Feld., but without
the characteristic black patch of hindwings in and the
cilia of forewings in that species are white on upper part
of termen (I), or almost wholly ($).
400
Mr. E. ]\Ieyrick’s Dearriptions of
Imma lyrifera^ ii. sp.
9 . 18-19 mm. Head orange-yellow. Palpi yellow, with a fiiscons
anterior stripe on terminal joint and upper half of second. Thorax
fuscous-purple, with four orange stripes on anterior half, and two
posterior orange marks. Abdomen rather dark grey. Forewings
elongate, moderately l)road, posteriorly dilated, costa gently arched,
apex rounded-obtuse, termen rounded, hardly oblique ; 7 and 8
stalked, 8 to apex ; fuscous-purple, suffused with blue towards
costa; markings orange; short costal and median streaks from
base ; an irregular rather outwards-curved streak from middle of
costa to f of dorsum, dilated on costa, interrupted in middle and
near dorsum ; a series of eight or nine interneural dashes between
this and following streak, but mostly not reaching either ; a terminal
streak, wide on costa and attenuated to tornus, anterior edge concave,
enclosing a blacki.sh striga from costa : cilia whitish, in one specimen
suffused with grey towards base, in the other with a blackish basal
line, Hindwings hyaline, veins blackish-grey ; a moderately broad
rather dark grey band along costa ; a moderate blackish-grey termi¬
nal fascia, becoming abruptly very narrow near tornus ; cilia fuscous,
outer half whitish, with blackish basal line.
New Guinea, Kei Is., Woodlark I. ; in December and
March, two specimens.
Imma peolploca, n. sp.
9. 21mm. Head, palpi, and thorax ochreous yellow. Abdomen
dark fuscous. Forewings elongate, rather broad, posteriorly dilated,
costa moderately arched, apex rounded -obtuse, termen nearly
straight, hardly oblique; 7 and 8 stalked, 8 to termen ; dark fuscous-
purple ; an ochreous-yellow streak running round base, costa, and
termen, forming a narrow basal patch which extends on costa and
dorsum so that edge is concave, very slender through middle of
costa, at apex forming a triangular patch, and again very finely
attenuated towards tornus : cilia yellow, beneath tornus fuscous.
Hindwings and cilia blackish-grey.
New Guinea, Jobi I., in May; one .specimen.
Imma nuhigena, n. sp.
1^. 27-29 mm. Head white or whitish, .scales of crown and
collar long, dense, erectible, partially suffused with fuscous. Palpi
ochreous-whitish or whitish-ochreovrs, second joint rather excurved,
with expansible scales towards apex above, with fuscous lateral
stripe or sometimes wholly fuscous externally, terminal joint
M %layan Micro- Lcpiclo'j^Ura .
467
with dark fuscous posterior streak. Antennal ciliations very sliort.
Tliora.x purplish -fuscous. Abdomen rather elongate, fuscous, with
long hairs on sides. Posterior tibiae witli whitish-ochreous hairs
beneath on basal half. Forewings elongate, rather narrow towards
base, considerably dilated posteriorly, costa anteriorly straight,
posteriorly gently arched, apex rounded, termen rounded, some¬
what oblique ; 7 and 8 stalked, 8 to termen ; dark fuscous ; basal
f darker -with a strong purple gloss, limited by a cloudy irregularly
waved pale greyish-ochreoustransver.se line, sometimes little marked;
a large rounded purple-blackish apical blotch, limited by a similar
less marked line from i of costa to dorsum before tornus, approxi¬
mated to first above middle but nearly ob.solete there, space between
these two lines .somewhat irrorated with pale greyish -ochreous :
cilia fuscous, with dark fuscous subbasal line. Hindwings fuscous,
lighter towards apex, ba.sal half suffused with blackish-fuscous ; cilia
fuscous, on lower half of termen whitish towards tips.
Bali, in April ; three specimens. Very near strepsizoiia
(of which the type specimen, described as a is really
therefore the differences are not sexual), but differs by
white and fuscous head and palpi (yellow in strejjsizonct),
less developed pale transverse lines, and peculiarly coloured
hindwings (in streiosizona uniform dark fuscous), with cilia
not white externally on upper part of termen.
Imma mormopa, n. sp.
5 ■ 14-15 mm. Head, thorax, and abdomen dark fuscous.
Palpi slender, whitish-fuscous. Antennae in pubescent-ciiiated.
Forewings triangular, costa strongly arched, apex obtuse, termen
somewhat rounded, slightly oblique ; 7 and 8 separate, 8 to costa ;
purple-blackish ; four thick transverse suffused whitish-fuscous
streaks, first two straight, third curved, sharply indented towards
dorsum, fourth from f of costa to tornus, curved, narrower ; a
whitish-fuscous discal spot between second and third : cilia fuscous,
basal third dark fuscous, at apex with a small w'hitish spot.
Hindwings dark fuscous ; .straight whitish-fuscous postmedian and
subterminal transverse streaks, former suffirsed with white on upper
half ; cilia rather dark fuscous, base whitish, at apex suffused with
whitish.
Moluccas, Banda, Amboina ; in August and February,
two specimens. This is a very interesting species, being
the most primitive form of the genus known to me, and
clearly showing the affinity with Simaethis and Brenthia ;
by assuming it as the starting-point, it becomes possible
468 Mr. E. Meyrick’s DcMriptions of
to trace the development of the rest of the species, hitherto
obscure.
Brcntliia caelicola, n. sp.
^ 9- 11-12 imn. Head dark kronze, face edged with white.
Pal^ii white, obliquely barred with dark fuscous. Autenuae white
spotted with dark fuscous, ciliations iu very short. Thorax dark
bronze, witli five fine white lines on anterior half. Abdomen dark
fuscous. Forewings elongate-triangular, costa slightly arched, apex
rounded-obtuse, termen nearly straight, little oblique ; dark bronzy-
fuscous ; a straight thick transverse streak of whitish irroratiou
at I ; a similar thicker streak at |, but terminated above by an
oblique streak of violet-golden-metallic irroration from middle of
costa ; space between these irregularly marked with whitish irrora¬
tion, and with a transverse-oval discal spot outlined with white ;
a moderately broad blackish terminal fascia, edged with whitish
irroration anteriorly, nearly divided into three spots by streaks of
ground-colour from anterior edge, lowest spot largest, and marked
just before termen with an irregular series of seven partially-
connected violet-golden-metallic dots : cilia bronzy-fuscous, with a
darker fuscous subbasal shade, tow^ards tips whitish. Hindwings
dark fuscous; an oval white spot in middle of disc ; a brilliant deep
blue elongate spot beneath costa beyond middle, and a similar
transverse streak before apex ; a white transverse streak before
central third of termen, its lower extremity resting on termen ; cilia
fuscous with dark fuscous subbasal line, with a patch of whitish
sulfusion at apex, and oblique patches of white suffusion above and
below middle of termen.
New Guinea, Kei Is. ; from February to May, four
specimens.
Brcntliia sriJaconia, n. sp.
(J. 9-11 mm. Head bronzy-fuscous, margins of face white. Palpi
white, with four dark fuscous rings. Antennae white spotted with
dark fuscous, ciliations moderate. Thorax rather dark bronzy-
firscous. Abdomen dark fuscous. Forewings elongate-triangular,
costa slightly arched, apex rounded-obtuse, termen somewhat
rounded, little oblique ; dark fuscous ; an irregular straight whitish
transverse streak before ; a transverse-oval discal spot outlined
with white, surrounding which is an irregular whitish ring almost
or quite touching costa and dorsum; a moderately broad blackish
terminal fascia, edged anteriorly with whitish and preceded by a
parallel whitish line, upper extremity cut off so as to form two small
469
Malayan Micro -Lepicloptera.
sjiots, marked near posterior edge with five violet-golden-metallic
dots, of which two are in the two upper spots, and upper spot also
preceded by a small golden-violet spot : cilia fuscous, with darker
subbasal line, at apex with whitish patch. Hindwings dark fuscous ;
a small obscure whitish spot in middle of disc ; a transverse shining
violet mark before apex ; on lower f of wing a shining violet- white
subterminal streak and suffused orange terminal streak : cilia dark
fuscous, with oblique wliitish patches in middle of termen and at
tornus.
New Guinea, Kei Is.; in January and April, two
specimens.
Brenthia spintheritis, n. sp.
d?. 11-12 mm. Head, thorax, and abdomen dark fuscous.
Palpi white with four dark fuscous rings. Antennae white ringed
with dark fuscous, ciliations in $ long. Forewings elongate-
triangular, costa slightly arched, apex rounded-obtuse, termen
somewhat rounded, little oblique ; dark fuscous ; an obscure cloudy
straight thick transverse streak of whitish irroration at some¬
times little marked ; a large undefined discal patch of pale violet-
golden-metallic irroration above middle reaching to costa ; a
moderately broad praeterminal blackish fascia obscurely edged all
round with whitish suffusion, upper end cut off to form a separate
spot, marked near posterior edge with five pale violet-golden-metallic
dots, of which one is in upper spot : cilia fuscous, with dark fuscous
subbasal line, at apex with a whitish patch. Hindwings dark
fuscous ; a small cloudy whitish spot in disc before middle ; a
thick whitish transverse streak at f abruptly constricted near each
extremity ; a whitish submarginal line, touching termen in middle,
towards upper extremity attenuated and suffused with golden-violet,
not quite reaching costa ; cilia dark fuscous, with white patch in
middle of termen, and less marked whitish patches at apex and
tornus.
New Guinea, Kei Is. ; in April and May, three
specimens.
SOBAREUTIS, n. g.
Head smooth ; ocelli present ; tongue apparently absent. An¬
tennae in d' simple, basal joint moderate, without pecten. Labial
palpi rather long, recurved, diverging, second joint with scales rather
rough beneath, with two or three short bristles at apex laterally,
terminal joint as long as second, moderate, pointed. Maxillary palpi
very short, loosely scaled, porrected. Posterior tibiae smooth-scaled,
with long projecting bristly scales at origin of spurs. Forewings
470
i\fr. E. Mey rick’s Deseo-^jfions of
with 2 from near angle, 7 to termen, 11 from before middle.
Hindwings t, elongate-ovate, cilia 1 ; 3 and 4 connate, 5-7 nearly
parallel.
Apparently allied to G1 yphipteryx or its ancestors.
Goharcutis conchophancs, n. sp.
12 mm. Head and i)a]pi white. Thorax dark fuscous, pos¬
terior extremity white. Abdomen dark fuscous, segmental margins
partially white. Posterior tibiae dark purple-fuscous, median band
and apical bristles white. Forewings elongate, rather narrow, slightly
dilated posteriorly, costa slightly sinuate, posteriorly moderately
arched, apex obtuse, termen obliquely rounded ; dark i)urplish-
fuscous ; two rather narrow shining w’hite fasciae, first near base,
not reaching costa, second beyond middle, rather inwards-oblique,
contracted above middle : cilia dark fuscous. Hindwings and cilia
dark fu.scous.
Borneo, Kuching (Hewitt) ; in November, one speci¬
men.
Thriambeutis, n. g.
Head with appressed scales ; ocelli present ; tongue short. An¬
tennae nearly 1, above with fringe of long rough projecting scales
except towards apex, basal joint short, without pecten. Labial palpi
short, porrected, loosely scaled, tolerably pointed. Maxillary palpi
obsolete. Forewings with 2 from §, 7 to termen, 11 from middle.
Hindwings under 1, elongate-ovate, cilia | ; 2-7 separate, tolerably
parallel.
Thriamheutis hemicausta, n. sp.
9 . 21 mm. Head, palpi, and thorax reddish-orange. Antennae
black, apical fourth whitish. Abdomen reddish-orange, apical half
blackisli. Forewings elongate, narrow, slightly dilated posteriorly,
costa slightly sinuate, apex obtuse, termen obliquely rounded ;
reddish-orange : a purple-blackish apical patch occupying rather
more than half wing, anterior edge irregular: cilia purple-blackish.
Hindwings reddish-orange ; apical half blackish, division irregular;
cilia reddish-orange, round apical half blackish.
Solomon Is., Isabel I. (Meek) ; one specimen.
PLUTELLIDAE.
Atteva monoplaneiis, n. sp.
9. 31-32 mm. Head dark grey, with white spots on sides of
face, betw'een antennae, and behind eyes, collar and back of crown
471
Malayan Micro-Lcpidoiytera.
orange. Palpi dark grey. Antennae grey, suffu.sed with whitish
towards apex. Thorax and abdomen bright deep orange. Fore¬
wings elongate, narrow towards base, posteriorly dilated, costa
gently arched, apex rounded-obtuse, termen somewhat rounded,
little oblique ; bright deep orange ; a deep purple patch occujjying
rather more than apical half, anterior edge suffused ; an oval white
spot in disc at rather below middle: cilia dark fuscous, white
towards tips at apex. Hindwings bright deep oi’ange ; a blackish
apical patch occupying rather less than half wing, anterior edge
suffused; cilia orange, round apical patch blackish.
Bismarck Is., New Hanover; in March, two specimens.
Atteva heliodoxa, n, sp.
^ . 34 mm. Head white, sides of crown marked with dark grey.
Palpi white, externally grey. Antennae grey. Thorax and abdomen
bright orange, anal valves very elongate, suffused with grey towards
apex. Posterior tibiae rather short, slender, white, with loose hairs
above, tarsi long. Fore wings elongate, narrow towards base, poste-
I'iorly dilateil, costa slightly arched, apex rounded-obtuse, termen
somewhat rounded, little oblique ; bright deep orange, apical half
deep fuscous-purple, division suffused ; a round white spot in disc
at i ; a transverse-oval large white spot in middle of disc, and
another still larger at J below middle ; five or six small round white
spots in an irregular longitudinal series beneath posterior half of
costa: cilia dark fuscous. Hindwings bright deep orange; a dark
fuscous patch occupying apical fourth of wing, and narrowly ex¬
tended along termen to below middle, anterior edge suffused ; cilia
orange, round apical patch dark fuscous.
New Guinea, Jobi I., in May; one specimen. Nearest
conspicua, Wals., but in that species the dark apical patch
of hindwings occupies more than half wing.
Atteva halanuta, n. sp.
5. 31 mm. Head white, with two large black spots on crown.
Palpi white, with a fuscous anterior streak. Antennae dark grey,
basal joint white. Thorax orange-fulvous, collar, a spot on each
side, and two posterior spots white. Abdomen bright orange. Fore¬
wings elongate, narrow, somewhat dilated posteriorly, costa slightly
arched, faintly sinuate in middle, apex obtuse, termen faintly sinuate,
somewhat oblique ; fulvous-orange, with 18 or 19 white spots or
marks, viz. six or seven small spots along anterior half of costa, a
moderate round subdorsal spot near base, two others discal and
472
Mr. E. Mey rick’s Descriptions of
dorsal near beyond this, a longitudinal-oblong spot on fold before
middle and one in disc at^, two moderate spots in disc above middle
(anterior transverse) and a larger round sirot on dorsum beneath these,
a subcostal streak from beyond middle to |, its extremities touching
costa, a small spot beneath posterior extremity of this, a larger round
spot almost at apex, and an irregular transverse mark towards
termen on lower half ; cilia fulvous-orange, tips paler. Hindwings
bright orange, thinly scaled and subhyaline towards base ; cilia
orange.
Philippines, Manila ; one specimen.
Anaplicmtis protona, n. sp.
^ 9- 18-19 mm. Head, palpi, and thorax blackish, collar orange-
yellow ; terminal joint of palpi in ^ as long as second, in 9 much
shorter. Antennae blackish, apical third whitish. Abdomen orange-
yellow, apical third blackish. Forewings elongate, rather dilated
posteriorly, costa moderately arched, apex rounded, termen obliquely
rounded ; purplish-black ; a rather broad orange-yellow transverse
band, extending from J to beyond middle, posterior edge with an
irregular projection below middle : cilia dark fuscous. Hindwings
orange-yellow ; a black basal spot ; a black patch occupying apical
f or rather more, and extended as a narrow streak along termen to
near tornus ; cilia blackish, on dorsum and tornus yellow.
New Guinea, Woodlark I. (Meek), in March and April ;
six specimens. Differs from isochrysa by wider black basal
area of forewings, posterior projection of yellow band, black
basal spot of hindwings, and larger apical patch.
Melodryas, n. g.
Head with appressed scales ; ocelli present ; tongue absent. An¬
tennae 1, rather stout, somevdiat compressed, basal joint moderate,
without pecten. Labial palpi moderately long, curved, ascending,
second joint thickened with scales, terminal joint as long as second,
moderate, acute. Maxillary palpi short, distinct, porrected. Poste¬
rior tibiae shortly rough-scaled. Forewings with 2 from angle, 3
absent, 7 to apex, 11 from middle. Hindwings 1, elongate-ovate,
cilia ^ ; 2-7 tolerably parallel.
Allied to Anapliantis.
Melodryas doris, n. sp.
9 . 20-21 mm. Head and thorax blackish, face and sides of
crown yellow. Palpi yellow, second joint with blackish lateral line.
473
Malayan Micro- Lepidoi^tera.
Antennae whitish-ochreous more or less tinged with grey, towards
base suffused with fuscous. Abdomen orange, apical half purple-
black. Forewings elongate, narrow, slightly dilated posteriorly,
costa gently arched, apex rounded, termen obliquely rounded ;
purplish-black ; a broad orange transverse fascia extending from
about ^ to middle : cilia dark fuscous. Hindwings orange ; extreme
base black ; a black apical patch occupying nearly half of wing ;
cilia orange, round apical patch blackish.
Solomon Is., Rendova, New Georgia (Meek); two
specimens.
TINEIDAE.
Dccadarchis discreta, n. sp.
9. 17-18 mm. Head and palpi white. Thorax ochreous-white,
shoulders blackish. Abdomen light grey. Forewings elongate,
costa moderately arched, apex pointed, upturned, termen very
obliquely rounded ; bronzy -blackish ; markings ochreous-white or
whitish-ochreous ; a moderate somewhat curved fascia at J, dilated
towards dorsum ; a semicircular spot on costa at and another on
dorsum before tornus ; a spot crossing wing just before apex : cilia
ochreous-white or whitish-ochreous, with dark fuscous patches at
apex and on middle of termen. Hindwings grey, thinly scaled
towards base ; cilia whitish-grey.
New Guinea, Kei Is.; Solomon Is., Guadalcanal-;
two specimens.
Monopis thiantha, n. sp.
d- 17 mm. Head pale yellow. Palpi dark fuscous. Antennal
ciliations 2. Thorax dark fuscous. Abdomen pale ochreous-
yellowish. Forewings elongate, rather narrow, costa moderately
arched, apex round-pointed, termen very obliquely rounded ; 3 and
4 stalked out of 2, 7 and 8 stalked ; very dark brown, with some
scattered dark fuscous scales ; a pale yellow semioval spot extending
on dorsum from i to beyond middle, containing several small tufts
of raised scales ; a larger light yellow trapezoidal blotch extending
on costa from 5 to f and reaching half across wing, narrowed down¬
wards, with a short projection from lower anterior angle, and a
round hyaline sjjot within this angle : cilia pale brownish sprinkled
with dark fuscous. Hindwings light purple-brownish, on basal half
grey-whitish and semihyaline ; cilia pale yellowish, with a palo
brownish shade.
New Guinea, Kei Is., in March ; one specimen.
474< Mr. E. Mey rick’s Dcscrii^tionn of
Monoijis callichalca, n. sp.
16 mm. Head light ochreous-yellow. Palpi dark fuscous,
apex yellowish. Antennae simple. Thorax dark fuscous. Ab¬
domen fuscous, apex pale ochreous-yellowish. Forewings elongate,
costa moderately arched, apex obtuse, termen obliquely rounded ; 3
and 4 stalked, 9 and 10 stalked ; dark purple-fuscous, strigulated
with ochreous-whitish on median area, and on costa and dorsum
posteriorly ; a broad bronzy-yellow transverse fascia extending from
near base to | ; a brassy-yellow postmedian fascia, broad on costa
and dorsum, posterior margin angularly emarginate, so that it
becomes very narrow in middle ; an oval ochreous-whitish semi¬
hyaline spot in disc preceding this ; a small ochreous-whitish spot
on costa near apex : cilia dark fuscous mixed with ochreous-whitish.
Hindwings dark grey, with bronzy reflections; cilia grey-whitish.
New Guinea, Kei Is., in March ; one specimen.
Monopis syharita, n. sp.
18. Head yellow. Palpi dark fuscous, apex yellow-whitish.
Thorax and abdomen dark fuscous. Forewings elongate, costa
moderately arched, apex obtuse, termen obliquely rounded ; 3 and
4 connate, 9 and 10 stalked ; dark purple-fuscous ; a large brassy-
yellow rounded blotch extending on dorsum from towards base to
beyond middle, and reacliing to near costa ; an inwardly oblique
brasfsy-yellow spot from | of costa, reaching half across wing, its
lower extremity preceded by a small whitish subhyaline spot : cilia
dark fuscous, irregularly mixed with whitish-ochreous. Hindwings
dark fuscous, with bronzy reflections ; cilia grey, with dark fuscous
subbasal shade.
Solomon Is., Choiseul (Meek) ; one specimen.
Tinea calycodes, n. sp.
9 . 18-19 mm. Head and thorax white. Antennae 1. Abdomen
light grey. Forewings elongate, costa moderately arched, apex
obtuse, termen extremely obliquely rounded ; all veins separate ;
rather light yellowish-ferruginous-brown ; base narrowly white ; a
broad white transverse fascia occiq^ying from } to beyond middle,
edged with a few black scales, broader towards costa, posterior edge
with projection just beneath costa and above middle ; an irregular
white apical spot, preceded by a few blackish scales : cilia light
ochreous-brownish, with rows of dark fuscous points, with a
white patch beneath apex. Hindwings brassy-grey ; cilia whitisli-
ochreous sprinkled witli grey points.
Solomon Is., Bougainville (Meek); two specimens.
475
Malrujan Micro-Lcpido2')tera.
OSPHRETICA, n. g.
Head densely rough-haired ; ocelli present ; tongue developed.
Antennae in ^ ciliated Avith long fascicles, basal joint stout,
densely scaled. Labial paljji long, subascending, second joint u ith
dense rough projecting scales beneath, terminal joint somewhat
shorter, slender, cylindrical, obtuse, rough-scaled anteriorly. Max¬
illary palpi long, filiform, folded. (Posterior tibiae defective.) Fore¬
wings with 2 from towards angle, 4 and 5 somewhat approximated,
7 to costa, 8 and 9 stalked, 11 from before middle. Hind wings 1,
elongate-ovate ; 2-7 tolerably parallel.
Osphretica chomatias, n. sp.
1^. 12 mm. Head and palju blackish-fuscous. Thorax ochreous-
whitish, anterior half blackish-fuscous. Abdomen grey. Forewings
elongate, rather narrow, costa moderately arched, apex rounded-
obtuse, termen slightly rounded, oblique ; whitish-ochreous ; a
blackish-fuscous longitudinal spot on base of costa ; a semioval
blackish-fuscous blotch extending on dorsum from before to near
tornus, reaching half across wing ; a narrow blackish-fuscous patch
extending along costa from f to apex, cut across by three oblique
whitish strigulae. Hindwings rather dark grey.
Borneo, Kuching (Heivitt), in October; one specimen.
CORYPTILUM, Z.
I find this genus possesses folded maxillary palpi, and
must therefore be referred to the neighbourhood of Tinea.
Corxjptilnm atrescens, n. sp.
g $. 28-36 mm. Head and palpi black. Antennae, thorax, and
abdomen blackish-fuscous. Forewings elongate, narrow towards
base, posteriorly dilated, costa moderately arched, apex rounded,
termen obliquely rounded ; dark ashy-fuscous irrorated with
blackish, with some scattered blue-metallic scales on A'eins ; a broad
subcostal streak of coppery-red suffusion from base to an oblique
transverse whitish blotch crossing disc beyond middle : cilia dark
purplish-fuscous with two blackish lines. Hindwings blackish ; an
elongate blotch of orange-reddish suffusion extending in disc from
base to beyond middle, sometimes much obscured or nearly obsolete
cilia blackish.
Java, Western district; four specimens. Smaller
and narrower-winged than rutildlum, from which it is
distinguished by the mainly blackish hindwings.
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) I I
47G Mr. E. Meyrick’s DescrijJtions of
Themeliotis, n. g.
Head with short rough hairs ; ocelli present ; tongue absent.
Antennae f, in ^ dentate, fasciculate-ciliated, basal joint with
flap of dense projecting scales anteriorly. Labial palpi moderate,
ascending, second joint with dense projecting tuft of scales beneath,
terminal joint shorter, cylindrical, obtuse. Maxillary palpi obso¬
lete. Thorax with posterior crest. Posterior tibiae with appressed
scales. Forewings with 2 from towards angle, 7 to termen, 8 and 9
stalked, 11 from before middle. Hindwings under 1, elongate-ovate,
cilia f ; 3 and 4 connate, 5-7 parallel.
Allied to Ardiosteres.
Themeliotis stereodes, n. sp.
12 mm. Head white. Palpi whitish, second joint blackish-
fuscous, tuft fuscous. Thorax white, shoulders dark fuscous.
Abdomen pale greyish-ochreous. Forewings elongate, moderate,
costa anteriorly strongly, posteriorly moderately arched, apex
obtuse, termen obliquely rounded ; white ; a moderate blackish-
fuscous basal fascia, enclosing a small white spot on base of dorsum ;
a juoderate blackish-fuscous median fascia, considerably dilated
towards costa ; a very small dark fuscous spot on costa towards
apex : cilia pale whitish-yellowish. Hindwings pale whitish-
yellowish tinged with grey ; cilia pale whitish-yellowish.
New Guinea, Sariba I. (Meek); one specimen.
Tinissa cinerascens, n. sp.
14-16 mm., $. 20-22 mm. Head and thorax ochreous-
whitish. Palpi white, second joint somewhat marked with fuscous
suffusion, terminal joint with dark fuscous median ring. Antennal
ciliations of 3. Abdomen ochreous-whitish, in $ more or less
suffused with grey. Posterior tibiae fuscous-whitish, posterior half
with dense rough expanded fuscous scales. Forewings elongate,
narrow towards base, posteriorly somewhat dilated, costa gently
arched, apex obtuse, termen obliquely rounded ; all veins separate ;
whitish-ochreous more or less infuscated, with a violet gloss, and
strigulated throughout with dark fuscous ; posterior half of costa
ochreous-whitish, marked with a small dark fuscous spot on middle
of costa and four others posteriorly ; a small obscure dark fuscous
mark on end of cell, sometimes followed by an obscure pale spot ; a
suffused dark fuscous terminal line : cilia whitish-ochreous, towards
tips somewhat infuscated. Hindwings in whitish-ochreous more
Malayan Micro- Lqnd,optera. 477
or less tinged with grey and snffusedly spotted with darker grey, in
$ dark grey ; cilia ochreons-Avhitish, in $ tinged with grey.
New Guinea, Kossel L, Sudest I. (Meek); six speci¬
mens. The species of this genus are very similar in
general appearance, but good characters are found in the
neuration, and markings of terminal joint of palpi.
Tinissa 'plirictodcs, n. sp.
. 16 mm., $. 22 mm. Head and thorax ochreons-whitish.
Palpi ochreons-whitish, second joint obscurely barred with rather
dark fuscous suffusion, terminal joint with dark fuscous basal and
subapical rings. Antennal ciliations of ^ 3. Abdomen ochreons-
whitish more or less infuscated. Posterior tibiae ochreous-whitish,
posterior half with dense rough expanded rather dark fuscous scales.
Forew'ings elongate, rather narrow' tow'ards base, somew'hat dilated
posteriorly, apex obtuse, termen hardly rounded, oblique ; 8 and 9
stalked ; whitish-ochreous, suffusedly strigulated throughout with
dark fuscous, strigulae more confluent in J ; posterior half of costa
dark fuscous, with a small whitish-ochreous spot at |, another
beyond it, and several irregular dots on or beneath costa pos¬
teriorly ; a suffused dark fuscous terminal line : cilia fuscous, base
whitish-ochreous. Hind wings in ^ whitish- fuscous, in 9 rather
dark fuscous ; cilia in ^ whitish-ochreous tinged with fuscous, in ^
suffused with fuscoiis.
Solomon Is., Choiseul (Meek); two specimens.
Tinissa rigida, n. sp.
18 mm. Head white, crown centrally suffused wdth brownish.
Palpi white, second joint tinged with fuscous suffusion, with tw'o in¬
complete dark fuscous bars, terminal joint wdth dark fuscous supra-
median ring. Antennal ciliations H. Thorax whitish spotted with
fuscous suffusion. Abdomen whitish-fuscous. Posterior tibiae
whitish, posterior half with dense roughly tufted brownish scales.
Forewings elongate, rather narrow, costa gently arched, apex obtuse,
termen obliquely rounded ; 7 and 8 stalked ; white, with pairs of
irregular partly confluent dark brown striae rising from small
spots on costa, these striae partially obsolete at j and 4 of wing.
Hindwings and cilia whitish-fuscous.
New Guinea, Kei Is. ; one specimen.
Scardia satyr odes, n. sp.
. 19 mm. Head, palpi, thorax, and abdomen dark fuscous ;
second joint of palpi with long dense triangular tuft beneath, ter-
I I 2
478 Mr. E. Meyrick on Malayan Micro-Le.'pido'ptcTa.
minal joint short, slender, pointed. Antennae fasciculate-cili¬
ated (2). Forewings elongate, costa moderately arched, apex
obtuse, tennen nearly straight, rather strongly oblique; all veins
separate, 7 to termen ; dark brown ; very obscure dark fuscous
markings, viz. a suffu.sed slightly oblique fascia before middle, a
transverse spot from costa at |, several small scattered spots pos¬
teriorly, especially a round spot towards apex and some small spots
on posterior part of costa and termen; two dark fuscous tufts of
scales on antemedian fascia, and one subcostal on spot at f , beneath
which is a small round pale ochreous spot : cilia dark fuscous.
Hindwings blackish ; cilia dark fuscous.
New Guinea, Rossel I. (Meek) ; one specimen. Allied
to the Australian ‘primaeva.
( 479 )
XVI. On Zizeeria (Chapmaoi), Zizera (Moore), a yroup of
Lycaenid Butterjiies. By T. A, Chapman, M.D.
[Read March 2nd, 1910.]
Plates LI — LX.
Moore instituted the genus Zizera for a group of butter¬
flies, of which he placed three, that occur in Ceylon, viz.
I'.arsandra, imlica and gaika, in it. His diagnosis of the
genus accords with liarsandra (Igsimon) as the type.* He
appends to the diagnosis, however, the words “ Type,
Z. alms” Alstis, however, belongs to a totally different
group of “ blues,” and does not conform to Moore’s diagnosis
of Zizera. One can only suppose that Moore, without
really examining alsns, thought to make his work more
acceptable to British readers by placing a British species
as type. To appeal to an unintelligent public in this way
is clearly to be regretted in a matter of science, whatever
it may be in other fields of human activity.
The question arises. Does Zizera belong as a generic
name to alms or to karsandra ?
It may be noted that both de Niceville and Bingham,
in dealing with the genus, cite alsus as the type ; and
Butler, in his very useful list of the species (Proc. Zool.
Soc., 1900, p. 104), includes alsns, and adds the alter ego
of alsiis, lorquinii. The rule apparently governing the
point is clearly expressed thus: ‘'When, in' the original
publication of a genus, one of the species is definitely
designated as type, this species shall be accepted as type
regardless of any other consideration.”
At first view this seems to make alsus unquestionably
the type. But one may view the matter thus : alsus is
not a species of the genus, and so cannot by any one (even
Moore) be made the type. It is further clear that Moore
made the genus for and on karsandra, and having done
that, the law of priority, valid now and here for him, if
for no one else before publication, it was impossible to
assert anything outside it as type. The case is certainly a
puzzling and unusual one. Another consideration also
* “Lepidoptera of Ceylon,” vol. i, p. 78, 1880.
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
480
Dr. T. A. Chapman m Zizeeo'ia,
arises. Zizera has been used for tlie harsandra group
over and over again, but, I think, never for alsus, till we
come to Butler’s list (1900). Short as twenty years may
be, it is perhaps sufficiently long, in this unusual case, to
add weight to the view that harsandra is the type, on the
recently strongly advanced contention that use and wont
shall be duly considered in this question.
If all this, however, is mere prejudiced special pleading,
and this view is finally held to be erroneous and that
harsandra (with its allies) has to be provided with a new
genus, I would suggest that the new name be Zizeeria, as
making the change least troublesome.
Possibly my best course is to replace Zizera by Zizeeria
(for harsandra), it will make matters clearer, and only
add another synonym if my alternative contention holds
good. To do this is, of course, rather to sit on the fence.
If Zizera adheres to als^is {minimus) I fancy it becomes a
synonym, but that is a matter that does not now con¬
cern me. For details of C. minimus 1 may refer to Tutt’s
“ British Lepidoptera,” vol. x (Butts., vol. iii).
My interest in the Zizeerias originated in the resem¬
blance of one or two species in some of their forms to
Lyeaeno'psis, and in the question whether the genus is a
genus of Lycaenopsids. I am now quite satisfied that
Zizeeria and Lyeaeno'psis belong to quite separate groups.
I have no very clear ideas as to the limits and contents of
the group to which Zizeeria belongs, nor as to what its
name ought to be. As it has not yet got one, we may for
convenience call this section of it the Zizecriidi; we may
define the whole group as containing those species with
the dorsal armature in two moderately separated portions,
i. e. with a narrow and (comparatively) structureless medio-
dorsal portion, and each lateral portion carrying a more or
less long curved hook articulated to it. In Zizeeria this
hook is remarkably long and slender. In the structure
of the girdle or ring there are one or two interesting
points. One of these is the central dorsal portion
which I have described as narrow ; this consists of two
portions, the anterior and posterior; in most of the species
these are fairly close together, and look like what, no
doubt, they are, the anterior and posterior margins of
the dorsal bridge more fully chitinised, as margins often
are ; but in ossa, less in malia, but very markedly in
lysimon, these two portions are separate. So that in
481
a Group of Lycaenid Butterflies.
lysimon there appears to be the usual narrow dorsal bridge,
but further back is another transverse strip, that is at first
glance very puzzling, as the membrane that attached it to
the other portion is lost in maceration. This lower bridge
is seen in figs. 18, 19 and 20. Another item is a distal,
weakly-chitinised, finger-like proces.s, best seen in ossa, but
also very evident in other species; see fig. 13, malm. It
is also very plain in indica (fig. 29). 1 have also noted
(under ossa) another projection (see fig. 19).
It is noteworthy that both these processes occur (some¬
what modified) in Antizcra, especially atrigemmata and
lucida. The Indian Zizeerias were those in which I first
was interested, and a critical examination of these affords
a solution of nearly all difficulties.
De Niceville (“ Butterflies of India,” 1890) gave thirteen
species, but stated definitely his opinion that there were
really only four species. Bingham (“ Fauna of British
India,” 1907) adopted this view and gives only four
species.
The fact is that there are six species of Zizeeria {sensu
lato) in British India. In Butler’s inclusive list of the species
in the Proc. Zool. Soc. (1900) these six species appear
under nine names, or eleven if we include names applied
to the same species in other regions.
The six Indian species are malm, ossa, karsandra, gaika,
otis and sangra. The two last names represent two dis¬
tinct species, one of which, sangra, is a form of lahradus ;
if the name otis belongs really to the same species, then
the other species is indica (Murray).
In Butler’s list, antanossa is a distinct (African) species
allied to otis {indica), and atrigemmata probably belongs
to the Zizeeriid group, but to a genus very distinct from
Zizera.
Gaika is certainly related to the Zizeeriidi, but its neu-
ration and the dorsal armature of the appendages make
its exact position very doubtful. It certainly does not
belong to the genus Zizeeria (type kccrsandra).
We have then six (or seven if we make lysimon and kar-
saiulra distinct) species of true Zizeeriids, i.e. that conform
to my definition of them as possessing very long slender
curved and sharp-pointed hooks (or articulated spines) to
the dorsal armature of the male appendages (making with
gaika, seven (or eight) ).
When we consider the structure of the male clasps,
482
]3r. 'IV A. Chapman on Zizeeria,
these seven species divide themselves into two groups, viz.
1st, maha, ossa and lysimon with l-arsandra, and 2nd,
lahradus, indica, and antanossa.
If we regard the neuration, they again divide into two
groups, fjaika forms one and all the other species the
other.
Rough camera sketches of the relations to each other of veins
11 and 12 of forewing, in A, Alsus {minim'^is), L, lysimon, G, Gaila,
At., atrujemmatu,'^ c ~ costa, veins as numbered — enlargement x 16.
The atrigemmata * group differs much in the append¬
ages, especially the dorsal hooks are shorter and have
rounded and blunt points. The neuration is also different
from the true Zizeeriids.
We find, then, that we have in the ZiZEERliDl (Zizera,
Moore) (type harsandra) 7 species — maha, ossa, lysimon,
harsandra, lahradus^ antanossa, indica. These species fall
into two divisions, which ought properly perhaps to be
distinct genera, but may be called divisions or sub-genera
if any one so prefers.
1. ZiZEERiA (restricted). Moore’s definition of Zizera,
but with these additions : a spot in cell underside fore¬
wing, beneath hindwing the first three spots (from costa)
are in line ; clasps with broad extremity and bold teeth,
and soldered to each other for some distance basally. Type
harsandra (with lysimon, maha and ossa).
2. ZiziNA, n. g. — Moore’s definition of Zizera, with the
addition of there being no spot in cell beneath forewing,
the spot beneath hindwing in space between veins 6 and
* Placed in Zizera by Butler, 1. c.
483
a Ormif of I/ijcaenid Butterflies.
7, advanced basally so as to be quite out of line with
those above and below it. Clasps only slightly soldered to¬
gether and with minute terminal teeth, but with a solitary
hair * longer than the clasp, thick and clubbed, arising
close to its base. Type, labradns (with indica and
antanossa).
3. ZizULA, n.g. — does not conform in neuration to
Zizera, and requires a separate generic heading, vein 11
of forewing joining vein 12, and not again separating,
i.e. it forms merely a bar from cell to vein 12. Like
Zizeeria (restricted), the hiiidwing spots are in a con¬
tinuous row, but like Zizina, it has no spots in cell of
forewing. The apj^endages possess a long clubbed hair
just as in Zizina, so that one doubts whether the neura¬
tion has the importance that custom compels one to attach
to it.
The wings are rather more elongated than in Zizeeria
and the spots beneath fall into a long oval on each wing,
rather than into the usual “ blue ” pattern as in Zizeeria.
I suggest
Actizera (Aktin f Zizera)
as a generic name for atrigemmata, with which Iv.cida,
stellata and imuagaea appear to be congeneric.
Veins 11 and 12 of forewing approach one another as
in Plebeiid blues, but are far from touching each other.
The appendages are similar to those of Zizeeriidi as to
the dorsa except for the comparative shortness of the
hooks, which terminate with a rounded end with very
little tapering. The clasps are bent, of quite a different
pattern to those of Zizeeriidi proper, and have teeth (when
present) not terminally, but along the inner lower margin.
The facies, at least of atrigemmata, is very Zizeeriid, and
on its superficial characters one might suspect it to be a
geographical race of lysimon.
When we come to the individual species my interest
has largely been in the Indian forms, of which de Nice-
ville describes thirteen species, but states very distinctly
his opinion that there are only four.
* This hair may be regarded as homologous with the lowest of a
continuous row that exi.sts on the clasp of lysimon, or more probably
with a still lower one of the same series, absent in lysimon.
f In reference to the white ray beneath the hind wing, present in
atrigemmata, usually very marked in litcida.
484
Dr. T. A. Chapman on Zizeeria,
In examining the genitalia of this group, as a possible
outlier of the Lycaenopsids I found that there were cer¬
tainly more than four Indian species. Accordipgly I felt
it desirable to examine the whole group as far as possible.
The Indian species are really six, which I will call pro¬
visionally ossa, harsandra, indica (Murray = otisV),
sangra (Moore = lahradus) and gaiJca.
I further concluded that the central group of true Zizee-
riids contained only these six, with the possible exceptions
of antanossa, which may be held to be distinct, but which
may perhaps be as easily regarded as a geographical race of
indica, and lysimon, which I incline to accept as distinct
from harsandra.
Zizeeria. — 1. Karsandra is usually smaller than rnaha
or ossa, but is sometimes *24 mm. in expanse, whilst ossa is
often only 22, and malm is sometimes as small ; indeed, I
have one specimen of only 20 mm.
Beneath the fore-wing the post-discal row of spots is
bolder than in nudia, and instead of being in a slightly
curved line is much arched, bringing the spot between
veins 2 and 8 much nearer the angle of the cell. Some
specimens of malm and ossa, however, vary in this direc¬
tion. A more constant difference is that harsandra has a
spot between veins 9 and 10, which much increases the
curved appearance of the row of spots. I am not sure
that this spot is ever absent in harsandra or lysimon ; in
malm it is present as a not very rare aberration.
I have a specimen from the Moore collection labelled
sangra (Mergui), and placed in it with real sangra, that
apparently was taken in the same locality as and with sangra,
and that differs from the type by having no spot in the cell
beneath the forewing, and in the row of spots being
rather small and more equal as in sangra, and not large
and bold and of graduated size. It can nevertheless be
recognised as harsandra by tlie position of the second spot
in the row beneath the hindwing. For myself, I must
confess I had exaiqined the appendages before noting these
several details.
The appendages are shown in figs. 20 and 24.
Fig. 20 shows how the clasps resent the attempt to
flatten them by lateral pressure, thereby agreeing with
maha and ossa (figs. 18 and 19).
The Mergui specimen noted above is clearly an aberra¬
tion of harsandra, as wanting sundry spots, and though
485
a Group of Lycaenid Butterflies.
the spots in the row are small like lysimon, they are
graduated as in harsandra, and the appendages are those
of karsandra. Fig. 4 is abnormal as showing a spot
beneath that in the cell.
Figs. 20 and 24 (karsandm), show how closely the clasp
is related in form to those of maha and ossa.
2. Lysimon. — I incline to agree with Butler in treating
lysimon and karsandra as distinct species; at any rate, it
is the most convenient way of dealing with them, though
I am not disposed to quarrel with any one who asserts
them to be only si;b-species or marked geographical races
of a single species, as it would be a quarrel rather about
words than fact.
The points of difference between lysimon (fig. 1) and kar¬
sandra (fig. 4) are ; 1st, Underside spotting, the pattern is
nearly identical, but the spots of upper wing are small, of
equal size, and in a slightly curved row in lysimon, large,
of varying si2!e and in a much bent row in karsandra.
This crisp statement may somewhat exaggerate the differ¬
ence, but it serves, in my experience, to separate the two
forms, with few exceptions. 2ud, The appendages differ in
the two forms ; lysimon has the form shown in figs. 22 and
23. Figs. 22 and 23 are practically of the same form, the
attitude of the clasp being rather different in the two
specimens. This I take to be the usual form of the append¬
ages in the species. Fig. 23 is from a specimen from
Teneriffe, and does perhaps differ a little from the more
typical form shown in fig. 22 in the greater width just
below the crown of teeth, and by the less prominence
of the terminal teeth. Figs. 20 and 24 show the form
in karsandra. Here the terminal teeth project more
markedly by a narrowing of the neck below and by a
depression in the middle of the toothed margin, so that
what one may call the heel is more prominent than is the
toe, as in fig. 22. 3rd, The third point is the geogra¬
phical distribution ; lysimon is African (with a small
subsidiary European range) ; karsandra is Asiatic.
It is of some importance to note that there seems to be
no intermediate, no gradation of one form with the other.
Yet in the Red Sea region they apparently meet. I have
lysimon from Aden and karsandra from Egypt and South
France. The latter (from Lang collection) is doubtless
erroneous. Whether those from Egypt are so or not 1
cannot say.
48G
Dr. T. A. Chapman on Zizecria,
Knysna is a synonym of lysimon ; at least, I have ex¬
amined no hnysna that are other than lyaimon. A large
specimen is illustrated (fig. 18).
]\raha. — Chandda, diluta, sqiialida, oqmlina, marginata,
((Ibococrvlevs, argia,japonica,al()pe. I enter all these as syn¬
onyms of maha, as I do not profess to be able to discriminate
whether any of them really belong to ossa, and possibly
hold priority over that name. I take opalma, Poujade, by
description to be a form of viaha, and it is very unlikely
to be anytluTig else, but as I have not seen a specimen,
I cannot assert that it certainly is maha. Specimens
labelled opalma in B. M. are from China, and are maha,
opalina, Poujade, is from Thibet. Butler ])laces mar¬
ginata, Poujade, and alhocoernleus, Riiber, as synonyms
of opalma. By Poujade’s description marginata is either
sangra or indica, probabl}^ the former. Alhocoernleus (from
Hong Kong), according to Bober’s figure, is a form of
maha. It is not a Thibetan form, and is probably a variety
of argia (= maha), the only species mentioned by Kershaw
(“ Butterflies of Hong Kong ”) to which it can possibly be
referred.
In Kershaw’s “ Butterflies of Hong Kong,” Plate VIII,
fig. 26, called Zizera maha, is Moore’s sangra, a form of
Z. lahradus; and Plate IX, fig. 8, called Z. argia, is Z.
maha, of which argia. is a local form, rather than perhaps
simply a synonym. In the maha group, de Niceville
(1890) describes five species, but says he does so merely for
convenience, and that he is satisfied that there is really
only one species.
Butler (1900), gives four species that he places before
lysimon, and we may assume that these belong to the
maha group. These four species afford ten names and
synonyms. I have little doubt they belong to the same
five species that de Niceville gives.
Bingham (1907), gives only maha, and adds various
other names as synonyms.
An examination of the genitalia shows that we have in
this group two species, one of which is of course maha.
The other I shall call ossa, without expressing any definite
opinion that it may not be entitled to be called dilnta, or
some other of the list of names quoted above. None of
tlie descriptions of any of the synonyms attributable (more
or less) to malm are definite enough to enable me to say
that they refer to ossa and not to maha. Diluta comes the
a Go'oup of Lycaenid Butterflies.
487
nearest. I select ossa, because it is Swinhoe’s name, and
he more confidently than any one else persists in saying
there are two species (and only two) in this group,
though his description of ossa is not more distinctive than
are those attributed to other names. I have specimens of
chandala^ apparently so labelled by Moore, that have very
much the look of ossa, but are really maha. Possibly
Moore meant the name chandala^ to apply to ossa, and was
deceived by these specimens of maha into thinking them
identical.
A critical examination {i. e. of the genitalia) of the
type specimens to which all these names apply is the only
way of settling whether ossa is or is not a synonym of one of
them. Certainly such names as diluta Siud squalida appear
to point to one of the most common distinctions that
enable ossa to be (with some probability) recognised as
differing from maha.
The genitalia afford an unmistakable distinction between
these two forms, and each is quite constant in the con¬
siderable number of specimens I have examined. But
the superficial characters are much less to be depended on.
Some seasonal forms of maha are I fancy all but in¬
distinguishable from ossa, on the upper surface. The
difference where it exists is that maha always has at least
some blue of the same character as we (in England)
regard as belonging to “ Blues ” (such as icarus, thetis,
aegon, etc.). This may be slight and evanescent, the
specimen may look quite like ossa, until held at some
angle that enables the trace of blue to come into view.
Ossa is without this, and is of a bony whiteness tinted with
a grey or green tone, that does notvary with the incidence
of light, or at least never yields a definite blue.
Maha averages larger than ossa, and in size as in other
characters appears to be more variable; maha (in my
series) varies from 20 mm. to 82 mm., ossa from 22 to
28 mm.
The seasonal and geographical variation of maha is
very great, that of ossa apparently much less.
On the underside the markings are of the same pattern
in both species, and vary in much the same way. All
the spots may be markedly black on a pale, even, almost
white ground, or may be pale, almost identical in tint
with a pale sienna brown ground-colour, to be made out
by the slightly paler rings surrounding them.
488
Dr. T, A. Chapman on Zizecria,
This tendency to obsolescence of the spots is rare in the
forewing, common in the hind, and apparently more
frequent in ossa.
The row of spots in the disc of the foi’ewing may be in
a fairly regular curve, or by a variation common in many
blues with this row of spots, the row may be angled by
the spot above vein 2, approximating to the discal spoh
The former is the rule in maha (fig. 2), the latter in ossa
(fig. 3), but the exceptions are frequent. On the hind¬
wing the condition is reversed. The same spot approxi¬
mates the base most in maha, in ossa it is often quite in
line with those on either side, making a nearly straight
row of five spots. But again there are many exceptions,
so that one cannot confidently separate the species by
these characters. I have no doubt that any one who is
familiar with both species in the field, could separate them
easily, and in the cabinet, I entertain any doubt only over
perhaps 10 per cent.
The male appendages are of very similar structure. A
reference to photographs of them {maha, figs. 17, 18,
ossa, figs. 19, 21) will save any long description, but it
may be pointed out that the clasps are much longer and
more slender in maha — length 1'05 mm. in maha, 0'84 in
ossa, but the width is equal at the point where the two clasps
become free from each other. This point is, however, a
two-fifth of the clasp from its base in ossa, and less than a
third in maha. The result is tliat the clasp of ossa looks
short and squat beside that of maha. The end or head of
the clasp is also different. Regarding it, as its outline
suggests, as a foot with a heel and toe, the heel in maha is
square, in ossa produced backwards, and tending to have
an acute point. The toe consists in maha of two spines,
very separate and distinct, and usually of about equal
length ; in ossa there is only one spine, terminating in a
conical process, on the upper margin of which a notch
marks what is probably the homologue of the other spine
as developed in maha. The process is, however, propor¬
tionally larger than in maha. The sole of the foot is fiat
in maha, but in ossa has a hollow, due to the projecting
heel.
ZiziNA. — In the sub-genus or genus we have two Indian
species {sangra and indica) and one African {antanossa).
I say sangra and indica, because these names, though
possibly both merely synonyms, have each a definite sig-
a Group of Lycaenid Butterflies. 489
nification, which their real names, if I knew them, have
not.
Sangra is certainly a form of lahradics, but then Idbradus
is a name up to the present understood to signify the
quite Southern Polynesian and Australian forms, and its
occurrence in Java, Sumatra, and India has not been
recognised.
Then the name otis has been applied to a supposed
species consisting of both sangra and indica. My real
difficulty in regard to names is in fact that I have no idea
what otis is. If otis = sangra, then the two Indian forms
are lahradus and indica, but if otis = indica, then the real
names are lahradus and otis. I don’t know that this
difficulty can be solved without reference to the type
specimen of otis, if it exist.
I am, no doubt, to blame for not following this point
up, since it is not a valid excuse that I interest myself
more in the facts than the names; it is, however, an
explanation.
I shall call the species (1) lahradus, with its vars. sangra
and dryina ; and (2) indica, the latter purely for con¬
venience and clearness until the real value of otis as a
name is decided.
If we may attach any value to Bingham’s remark that
sangra and dccreta are slight varieties of otis, but that
indica may be separated by the larger size of the spots,
we must conclude that otis = lahradus and indica stands
good as the name of the other species. The Fabrician
description also rather favours sangra than indica, and the
habitat China seems to exclude indica, which happens to
be very descriptively named, hardly occurring out of
India.
De Niceville and Bingham recognised only one species,
and this they agreed must be otis, of which therefore
sangra and indica were both synonyms. Butler referred
sangra to otis, retaining indica as a separate species. In
making two species Butler was here unquestionably right.
In working the matter out, I found that Moore’s sangra
was rarer in India than indica, and was represented in his
collection by only a few specimens; I concluded, therefore,
that otis was the common (Indian) species indica.
This is contrary to Butler’s conclusion, and I fancy to
the belief of others, if those can be said to have a belief
who recognise otis only. Mr. Butler’s distinction between
490
])r. T. A. Chapman on Zizccrin,
otis and indica is precisely that wliich I draw between
sangra and indica. The B. M. material which he so dis¬
tributes, is, as judged by its labelling and localities, simply
otis = sangra — indica distinct ; since sangra is a form of
labradus, the distinction between these two forms that
Mr. Butler draws, confirms my view that otis is a synonym
of labradus. The same conclusion is led to by the evidence
given by Kershaw (which I refer to under maha) as to the
species found at Hong Kong.
Taking tcdjradiis as the first species, we have to enlarge
our conception of that insect by adding most of the forms
known usually as otis, as sangra, and also my recently
described dryina. Its range must also be extended north¬
ward to the Himalayas, and to Hong Kong (Kershaw)
(again a suggestion that it is otis), and in fact to include
the habitats recorded for otis.
The Indian (or sangra form) seems to be rare con-
tinentally. I have a specimen labelled by Moore “ N. Ind., ”
and one from Burmah. My other specimens are from Ungu,
Mergui (Moore Coll.), Andamans, Nicobars, Key Islands,
Celebes, and in the form dryina from Java and Sumatra.
Indica, on the other hand, has a comparatively very
restricted habitat, India, south and west, including Ceylon,
north-east as far as Lucknow. Labradus has an immense
range of variation geographical, seasonal, and aberrational.
The size varies from {lahradus) 32 mm., (dryina) 34 mm.,
down to 20 or even less in the sangra forms. The spots
beneath, especially beneath the hindwing, vary, as in
maha, from very distinct black spots in Avhite circles, to
mere ghosts in which the spots and circles are almost of
the ground-colour. How far this variation is seasonal
only, I don’t know.
Sangra may be distinguished from indica by the com¬
parative straightness of the row of spots under the fore¬
wings and their smaller and more uniform size, and by
the position of the second spot (from the costa) in the row
beneath the hindwing. In labradus {sangra) it is placed
so that the line it forms wdth the first spot would pass
close to the discal line and, if continued, would strike the
sixth spot. In indica it is less basal, so that a similar
line would strike the fourth spot. Both these points are
well brought out in figs. 5 {sangra) and 6 {indica).
The form dryina (Proc. Ent. Soc., 1908, p. Ixxxii) is
very large, superficially very like Lycacnopsis argiolus
491
a Group of Lycaenid Butterflies.
var. sikkima. Mr. Druce has a specimen from Java, whence
also Col. Swinhoe has specimens somewhat smaller ; I
have also one from Sumatra. This specimen was in Moore’s
series of Z. maha, which it much resembles, but is very
pale, almost white beneath. When I described dryina as
a new species, I had not worked out the Zizeeriids, and
supposed so large a form that was not maha must be new
(as indeed it was, but not as a species) ; it did not occur
to me that otis (i. e. sangra) could be so large, or labradus
occur so far north. The male appendages prove clearly,
however, that it is merely a local form of labradus.
Having figured these it could not remain as a puzzle
to any one else, had I failed to trace, as I now do, its real
position myself
The appendages of labradus will be better understood
by a reference to figs. 25, 26, 27, 28 and 34, and to
Plate B, fig. 2 in Proceedings for 1908, than by long
description. The figures (when reproduced) will probably
fail to show the minute teeth at the end of the clasp ;
on the end of the clasp to the right of fig. 26 nine
minute teeth can be counted on the distal margin.
These figures, with those of indica and gaika, show that
the clasps of these are nearly as well seen on lateral com¬
pression of the appendages, as when trouble (with risk of
damage) is taken to display them by antero-posterior
compression after dividing the ring of the appendages.
This contrasts with the facts concerning Zizeeria {maha,
etc.).
Indica. — As my account of sangra had to be somewhat
comparative with indica I have already really dealt with
indica. It very closely resembles karsandra beneath, differ¬
ing by the want of the cellular spot on the forewing, and
the position of the second on the hindwing.
One may remark here on the curious way in which
these species run in pairs.
Indica differs from sangra in the spotting, much as ossa
differs from media, and as again karsandra diffei's from
lysimon. One can hardly help theorising that lysimon
is dividing, unless one admits that it has already done so,
into these two species on precisely the same lines as the
other two pairs have already done.
The clasps in indica are not unlike those of labradus,
but the larger process being comparatively short and
simple in structure gives it at first glance an aspect of
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) K K
492 Dr. ^ A. Chapman on Zizccria,
considerable difference. Different aspects are shown in
figs. 29, 30 and 31.
Antanossa appears to vary in the size and distinctness
of its underside spotting in much the same way that
lalmulns {sangra, etc.), does, but as I have seen fewer
specimens, I cannot say that its range is quite as great.
I have a specimen of dryina (var. of lahradus) from
Sumatra, from the Moore collection (wherein it was ranged
with maha), that I could not define from antanossa by
spots.
Antanossa (fig. 32) agrees with indica in liaving the
spot in the interspace 4-5 of upper wing, usually some¬
what elongated obliquely, but this occurs sometimes in
lahradus and is wanting in some specimens of indica.
The genitalia point to antanossa being merely a geo¬
graphical variety of indica. There is a difference, but
hardly enough to carry much weight if the species were
not clearly distinct otherwise, and occurred on the same
ground.
The differences consist in the long basal hairs in
antanossa being distinctly longer and not terminating in
so finely produced a point, they are about twice the
length of the clasp ; in indica they are little more than
half as long again. In both species there are towards
the end of the clasp, about half a dozen long and strong
hairs, whose extremities range very much with those of
the basal hairs. In antanossa these hairs are on and close
to the end of the clasp, those further back being smaller,
whilst in indica the strong hairs are ranged along the
margin of the clasp, those on the extremity being small.
The upperside colouring of the width of border, etc.,
appears to vary in much the same way in all three species,
as it docs, in fact, throughout the group.
ZiZULA.
I propose this as the generic name for gaiha. Gaika is
obviously not a true Zizccria generically ; that it belongs
to the same tribe may be supported by the presence of the
remarkable hair on the clasps, that exists also in Zizina
{lahradus and indica). But it differs from true Zizecriac
in the neuration (vein 1 1 of forewing), in the form of the
dorsal hooks of the appendages, and in the pattern of
the marginal markings of the wings, these tending to fall
into straight lines, instead of the sagittate character in
a Group of Lycaenid Butterflies. 493
the other species of the group. (See fig. 8, compared
with wing figures of other species.)
The appendages are shown in Figs. 33 and 35 (the
latter more magnified). The hooks remind one of those
of Plebeiids, but it may be noted that short, thick and
curved as they are, they are without the definite angular
end of Plebeiids.
The most notable structure, perhaps, as marking ZizuLA
off from ZiZEERiA is the renmrkable form of the oedeagus,
which is very large, short and thick, with two immense
terminal, slightly curved sj^ines, very different from the
single fine, straight spine of the other species.
In all the rest of the tribe {Zizeeria, Zizina), the
oedeagus is large, flask-shaped, with a dilated base and
a narrower neck, terminating on one side in a fine spine.
This may be seen in several of the figures, where also may
be noted the very long portion of the eversible membrane
(vesica of Pierce), occupying the whole neck of the “ flask ”
(in the position it assumes in nearly all my mounts), which is
armed throughout its whole circumference by fine spicules,
looking very like a scaled surface; well seen in several
of the figures, especially perhaps, fig. 28.
In Zizida gaika, there appears to be no such armature
of the eversible membrane, which in truth I have not
definitely seen, but in one of the large terminal spines of
the oed^eagus, the upper one in both my figures, there
is centrally a curious little triangular, flange-like process
with one margin notched. As this occupies exactly the
same place in all the specimens I have examined, I take
it to be attached to the spine and not to a contained
membrane. This spine is smooth, polished, and seems to
taper to a point, the other looks very similar, but its lower
margin is serrated, and it has the appearance of being
articulated at its base ; it may, perhaps, belong to the
eversible membrane.
The clasps have much the aspect of those of indica
if very shortened and reduced, and the upper hairs
strengthened. It is further to be noticed that the great
basal hairs have a companion, are much larger and
stronger proportionally than in Zizina and are not simply
clubbed by a spindle-shaped expansion at the end, but
appear to be fluted or grooved for some distance, so that
the end is broadest at the very end, and then divided by
the grooves into several portions.
K K 2
494 Dr. TTA, Chapman on Zizecria,
Antizera,
Atrigemmata can hardly be included in Zizecria, it
belongs to a group which appears to include also lucida,
stellata and imnagaea. This group has the neuration of
ordinary Lycaenicls {Flcbeius, etc.). The dorsal armature of
the appendages has less long and slender hooks, and the
clasps are of a pattern quite different from either branch
of the Zizecrins. I suggest as a generic name for the
group Actizera, in reference to its resemblance to Zizecria,
and to the white ray so obvious in lucida and indicated in
atrigemmata. and others. A white streak found also, of
course, in Plebeiid and other Lyeacnas, but of which
Z^ 'zceria has no trace. The underside markings of
erschojji suggests strongly an alliance with them, but
the appendages show it to be a Plebeiid.
Atrigemmata- (figs. 36 and 39) may be taken as the
type of the genus, the clasps are without spines. The
appendages of Ineida (fig. 40) are curiously similar to
those of atrigemmata, differing in little except the posses¬
sion of spines on the convex margin of the clasp.
Fig. 7 shows the underside of lucida, and fig. 11 the
neuration.
The appendages of stellata (fig. 38) are much larger
and bolder, especially the dorsal half of the ring (or
girdle) is very broad and heavy, but the dorsal hooks
and form of the clasp, leave no doubt that it belongs
to the atrigemmata group.
As to imnagaea (fig. 37), there may be some doubt as
to its claim to being in the same genus with atrigemmata,
but it is certainly somewhere very near. The neuration
is the same and the appendages are to the same pattern.
The clasps notwithstanding their large size and great
breadth are really very similar to those of lucida, the
dorsal hooks are more Zizeeriid than those of the other
species of the genus. The peculiar short, broad oedcagus
is characteristic of the genus.
I have examined several other Palaearctic species that
had some appearance of alliance with panagaea, and find
on examining the appendages that some are so, but not
at all closely. Anisop)hthalma and cytis are the nearest;
adraea remotely, if at all. Korlana, though also an Everid
and closely related to prosecusa, appears to be quite distinct,
the facies of the underside is even more Zizeeriid than
495
a Group of Lycaenid Butterflies.
that of prosecusa. Apart from the appendages the two
species differ in the position of the second spot from the
costa, precisely as Icarsrmdra and indica do.
Suhcoerideus, Holland, Nov. Zool., vol. vii, p. 69 (1900),
I have not seen, but from the very careful description, 1
have very little doubt that it is a form of the very
polymorphic Z. labradus.
Delospila, Waterhouse, Proc. L. Soc. N.S.W,, 1903, p. 211,
Plate II, seems not to be a ZiZEERiA.
belongs to some distinct section of Lycaenids accord¬
ing to the evidence of the appendages.
Messaims and mahalloTcoaena belong to some group far
from Zizeeria, the latter shows a beginniog of the extra¬
ordinarily elaborate development of the dorsal armature
found in harherae, the appendages are also very suggestive
of alliance with Castcdius and Neopithccops.
ZIZEERIIDI.
Zizeeria.
Icarsandra.
hjsimon.
rnaha.
osna.
ZiZINA.
labradus (with, vars., otis (= sangra) and
dryina).
indica.
antanossa.
ZiZULA.
gaika.
Actizera.
atrigemmata.
Incida.
stellata.
2mnagaea.
etc.
Explanation of Plates
Prate LI. Fig. 1. Z. lysimon, underside.
2. Z. nudia, underside.
3. Z. ossa, underside, compare greater curvature of
row of spots in forewing, and that in fig. 2
49G
Dr. T. A. Chapman cm Zixceria,
on the liindvving, spots 3, 4, 5 (from costa)
are in line ; in fig. 3 the three spots in
line are 4, 5, and 6. The additional spot
below spot in cell is a not rare aberration
in Zkeeria. See also fig. 4.
4. Kdrxandm, in typical lymnon (European
and African) the spots differ in size and
boldness, as seen in fig. 1.
Plate LI I. Fig. 5. Z. lahnulus (var. sanyra).
6. Z. indica, line of spots in forewing compara¬
tively straight in 5, Ijent in 6. Note
different direction of line joining spots
1 and 2 (from costa) in the two figures.
7. A. lucida, note coalescence of spots below
vein 3 on forewing. These spots and some¬
times the next are wanting in atrigemmata.
Comparing also dellcda and panagaea, one
notes much plasticity in the development
of the spots.
8. Z. gaika, note continuity of inner marginal
line, and a spot above vein 10 in forewing.
Plate LIT I, showing neuration of —
Fig. 9. Z. onsa.
10. C. alsvs (minimus).
11. A. Incida.
12. Z. gaika.
Plate LTV. Fig. 13. Z. lysimon, var. knysna.
14. Z. karsandra (to show neuration).
15. Z. indica (to show neuration).
16. Z. maha (to show neuration).
The remaining figures all show appendages, all magnified
X 35, except fig. 35, which is x 45.
Plate LV. Fig. 17. Z. maha from a specimen of the Godman
collection (now at B. M.), labelled argia.
18. Z. maha, lateral view.
19. Z. ossa, lateral view (Poonah).
20. Z. karsandra, lateral view.
These four figures show well the long slender dorsal hooks, the
projection on the ring or girdle, below the dorsal processes and
separated from them by a narrow neck, the fine sharp spine ter¬
minating the oedeagus and the spines lining the eversible mem¬
brane. Note also, in fig. 19, ossa, the sharp spine on the ring just
opposite the projection, not apparently present in any other species
of the group.
Trans. Ent. Soc. Lend.., igio, Plate LI.
Pkot^, A. E. Tenge.
Zizeeria: 2. maha.
I 4. kanandra.
!
I. lysimon.
3. ossa.
1
Trans. Ent. Soc. Land.., iQio, Plate LII.
Photo, A. E. Touge.
i;. Zizina labradiis (var. sangra). 7. Antizera lucida.
6. Z. indica. 8. Zizula gaika.
Trans. Ent. Soc. Loud., igro, Plate LIII.
9
11
Photo, A. E. Tonge.
Neuration of —
9. Z. ossa. lo- C. alsus (minimus).
II. A. lucida. 12. Z. gaika.
Trans. Eni. Soc. Lond., IQIO, Plate LIV.
Photo, A. E. Tonge.
Neuration of —
13. Zizeeria lysimon ; 14. karsandia ; 15. Zizina indica ; 16. maha.
r
.'•J
:i
.i
: i
■ aAjfl
a
I
1
■ ■ ■ ■ ■(
i
.1
Trans. Ent. Soc. Land., ipio, Plate LVI.
ZiZEEKiA : 22. lysinion. 21. ossa.
23. lysinion. 24. karsanclra.
r
% *
f -
( ^
’f'
t
>^i
25, 26. Zizina labradus. 27, 28. var. sangra.
29, 30, 31. Zizina indica. 32. Z. antanossa.
f
f
" 4
i:
i
9
4
k
■ -ti
y.*-/fi
■■r‘
•vrir 9nv^ 'oi6t ’••vuot 'fti-j 'suo.tj.
;, 35. Zkula gaika. 34. Zizina labradus. 36. Antizera atrigcmmata.
1
; <
4
i
00
CO
o
"it
Antizera : 37. panagaea. 38. stellata.
39. atrigemmata. 40. lucida.
497
a Group of Lycaenid Butterflies.
Plate LVI. Fig. 21. Z. ossa, mounted in same manner as fig. 17 so
as to be comparable.
22. Z. lysimon, from Natal.
23. „ „ from Teneriffe.
24. Z. Mrsandra, Egypt (?).
Plate LVII. Fig. 25. Z. lahradns, lateral view.
26. „ „ , spread.
27. „ „ var. sangra (Celebes).
28. „ „ „ „ (Andamans).
Plate LVIII. Fig. 29. Z. indica, dorsal view.
30. „ „ (Bombay).
31. „ )>
32. „ antanossa.
Note in figs. 31 and 32 how the dorsal hooks tend to break short
off.
Plate LIX. Fig. 33. Z. gaika (Kutch).
34. Z. lahradus, var. sangra (Nicobars).
35. Z. gaika (Ceylon) (enlarged x 45, all others
are x 35).
Note the different form of dorsal hooks from all preceding figures
and the very remarkable oedeagus with two spines, the upper (in
plate) containing a very remarkable serrate process, apparently leally
in a groove of its underside, the lower looks as if articulated, and is
probably a spine of the eversible membrane, or its articulation may
enable it to close on the upper one, if that is grooved, and so form a
single sharp point. I hazard this as the best guess as to the real
structure of this remarkable organ that I can make.
Fig. 36. A. atrigemmata (specimen in B. M.).
Plate LX. Fig. 37. A. panagaea.
38. A. stellata (B.E. Africa).
Note the great size and breadth of the dorsal and lateral portion of
the girdle ; their overlapping somewhat confuses the preparation.
Fig. 39. A. atrigemmata.
Compare figs. 36 and 39 with fig. 19 as regards the distal finger-
like process of the ring and the sharp angle on its proximal edge.
The latter is even more notable in
Fig. 40. A. hicida.
( 498 )
XVII. Mr. Alfred D. Millar’s E-xim'imental Breeding from
the Ova of the Natalian forms of the Nymphaline
Gemis Euralia. By Roland Trimen, M.A., F.R.S.,
etc.
[Read June 1st, 1910.]
Plates LXI— LXV.
In a preliminary note communicated to the Society through
the good offices of my friend Professor Poulton on March
2, 1910 (see Proc. Ent. Soc., 1910, p. xiv), I had the
pleasure of making known the highly interesting results
of my friend Mr. Alfred D. Millar’s experiments in breed¬
ing from the ova of the three forms of Euralia known to
inhabit the coast-belt of Natal, and hitherto recognised
as distinct species, viz. — E. wnlilhergi, Wallengr., E. mima.
Trim., and E. deceptor, Trim. I now propose to give the
promised full account of these experiments, as illustrated
by the specimens, photographs, and carefully recorded
observations furnished by Mr. Millar.
Euralia is nearly allied to Diadema (= Hypolimnas),
and was founded by Westwood in 1850 on two West
African species (dubia, Palisot, and anthedon, Doubl.)
having the discoidal cell of the hindwings open. Other
distinctions of Euralia are the narrower head, considerably
longer and gradually clavate antennae, and the anal angle
of the hindwings more prominent in the male. The larva,
as Mr. Millar observes, differs from that of Diadema
misippus in its considerably longer spines generally, and
still longer pair of spines or horns springing from the top
of the head.
Euralia, in the advanced development of its mimicry of
the Danaine genus Amauris, almost equals the eurytus-
group of another Nymphaline genus, Eseudacraca, with its
similar wealth of close imitation of the Acraeine genus
Planema. I have recently commented on the latter mimi¬
cries in my appendix to the Rev. St. Aubyn Rogers’
“ Bionomic Notes on British East African Butterflies,”*
pointing out the persistency, exactness, and completeness
with which these Fseudacraeae reproduce the pattern and
* Trans. Ent. Soc. Lond., 1909, pp. 551-2.
TRANS. ENT. SOC, LOND. 1910. — PART IV. (DEC.)
Mr. R. Trimen on the Nymphaline Genus Euralia. 499
colouring of their models, the very variable Plcmemae —
every variation in both sexes appearing to be faithfully
copied throughout tropical and sub-tropical Africa wher¬
ever the genus Planema prevails. I mentioned that of the
very limited number (about 20) of provisionally recognised
species and sub-species of Plancmci, 10 were already known
to be thus deceptively mimicked by corresponding forms
of Pseudacraea, and expressed the opinion that other
parallel cases probably remained undiscovered. Similarly,
among about 26 provisionally admitted species and sub¬
species of Amauris, I find 8 cases of unquestionable mimi¬
cry by forms of Euralia.
E. mima and E. toaJilbergi are the representatives in
the east and south-east sub-region of the West African
E. dubia, Palis.,- and E. anthedon, Doubl.,* and like these
latter mimic very accurately species of the Danaine genus
Amam'is widely differing from each other in appearance.
The very variable E. dubia seems to follow in its variations
A. egialca. Cram., A. psyttalea, Plbtz, A. hecate, Butl., and
A. tartarea, Mab., inhabiting the same country ; but E.
anthedon seems to be constant in mimicking A. niavius,
Linn. In the same manner the eastern and south¬
eastern E. mima imitates closely A. cdbimaeidata, Butl, and
— very rarely — A. eeheria, Stoll ; while E. wahlbergi copies
A. niavius, sub-sp. dominicanus, Trim.f The third Natalian
form of Euralia — E. deceptor, Trim. — is more prevalent on
the tropical East African coast up to the equator, and
imitates in its turn, with equal accuracy, A. oehlea, Boisd.,
which has the same geographical range. Since its dis-
* Aurivillius (Ent. Tidskr., 1894, p. 282) note.s a Camaroon ^
Anthedon and a Kaflrarian ^ wahlbergi whicli nearly approximated
in the width of the hindmarginal border of the hindwings, and ex¬
presses the opinion that this shows them to be local races of the
same species. He also mentions {1. c., p. 281) that a $ A. dubia var.
c.erberus, Auriv., from Camaroon has the spots of the forewings quite
like those represented in my figure of E. mima from Natal in Trans.
Linn. Soc. Lond., xxvi, pi. xliii, f. 7 (1869).
t In Madagascar, a variation of E. wahlbergi, named mada-
gascariensis by Mabille, appears to be modified in simulation of
Amauris nossima, Ward, inhabiting the same island ; and a form
intermediate between E. dubia and E. mima, the drucei of Butler,
prevails — although there is no known Amauris from Madagascar
that could be its model. It seems possible that the dimorphism now
represented by madagascariensis and drucei had been developed and
established in the continental-African ancestors of these forms before
their range had extended to Madagascar,
500 Air. R. Trimen on Mr. Millar's Exiicrimental
covery in Natal in the year 1873, E. deceptor has hitherto
remained very much rarer tliere tlian wahlhergi and
mini a.
The obviously intimate affinity of. these three forms of
Enralia in Natal was brought to notice by me as far back
as 1873 in the “Transactions” of the Society; together
with a note of the fact that E. minia $ and E. wahlhergi ^
had once been taken paired. Although, in the long interval
since that date. Air. Alillar, Air. C. N. Barker, Air. Guy
A. K, Alarshall, and other entomological observers in
Natal, have called attention to a few other cases of the
pairing of the latter two forms, as well as to various
details of habits, distribution, etc., which supported the
view of their species-identity — no advance of importance
had been made towards settling the question until the
earlier part of last year, when Air. Alillar, having ascer¬
tained their common food-plant, resolved to test the case
by breeding from the ova of all three forms.
First Experiment. — Offspring reared from ova laid hy
two examples of Euralia deceptor.
The food-plant — a species of stinging-nettle identified
by Air. J, Aledley Wood as belonging to the genus Fleicrya
— was potted and enclosed with netting, so as to be at
hand for the safe rearing of the anticipated larvae. On
Alarch 21, 1909, at Alount Edgcumbe, near Durban, two
$ ^ A', deceptor were captured in the act of ovipositing,
and placed on the net-covered food-plant, where they
together during the 24th and 25th laid 139 eggs, from all
of which the larvae Avere hatched by Alarch 29. On the
31st the young larv^ae were dark-brown, except towards each
extremity, where the colouring was pale semi-transparent
greenish-yellow, and they Avere closely covered throughout
Avith minute dark spines. On April 5, after moulting, the
body generally had changed to shining-black, and the
strong spines toAvards the front of each segment, as Avell
as the pair of long spines or horns on the head, Avere
developed ; but not until the 15th, after the final moult,
Avas the full colouring and development attained, in which
the body Avas velvety-black Avith segmental very narroAv
anterior elevated pale-yellowish transverse bands or half¬
rings, bearing numerous acute spinulose brownish-yelloAv
spines, and the head AA^as shining yellowish-brown, with
the two long divergent spinulose spines or horns black.
The change to pupa began on April 19; 60 pupae were
Breeding of the Nymphaline Genus Euraha.
501
disclosed by the 24th ; and all the remaining larvae had
pupated by the 29th— with the exception of six taken tor
preservation as inflated dried specimens. tvt o ;i
The appearance of the imagines began on May 2. and
continued for 13 days, ending on the 14th. The ^ ^
took the lead, 29 of that sex only emerging during 2nd to
4th • the 2 2 came out in smaller number than the £ 6
a to 17) on the 5th and 6th, but in larger inimber than
the £ £ (45 to 24) from the 7th to the 11th ; while on the
last three days, 12tli to 14th, ? ? only emerged to the
number of 5. The total number of imagines bred trom
the 139 ova deposited by the two dcceptor mothers was
thus 127 (70 ^ ^ and 57 ? ?) ; 4 ova appear not to have
hatched, 2 larvae died, and 6 larvae were preserved.
Only 1 imago emerged with crippled or deformed wings.
The illustration (Plate LXI), reproducing a photograph
taken by Mr. J. H. Burn of Durban, represents in minia¬
ture the whole imago progeny of the two deceptor motheis,
with the exception of the deformed example just men¬
tioned and three others which were damaged in setting.
The very small percentage of loss in rearing this larp
decevtor progeny in confinement would seem to point, in tlie
case of a butterfly so rarely taken in Natal, to the existence
of severe persecution or other adverse conditions attecting
the insect when at liberty in that country. The mothers
under notice were fairly prolific, and the offspring reared in
confinement under protection were, with o^nly one excep¬
tion, fully developed and richly coloured. Mr. Millar
writes that deceptor is one of the rarest ot our boutn
African butterflies, and during my collecting (now extend-
ino- over 35 years) I have not taken a dozen good speci¬
mens.” I have of late years inclined to the beliet that,
like several other species of butterflies now found in Nata ,
deceptor is a recent immigrant into that country that has
extended its range along the coast from the northwaid
It is— especially as regards the so pronounced a mimic
of Amauris ochlea, that in its imago state this form should
be shielded from excessive persecution by insectivores ; and
one is led to suppose that its scarcity in Natal— where its
food-plant does not seem to be rare--may be due to
enemies attacking the larva or pupa. A butterfly of this
size so very conspicuous and striking in its pattern and
colo’ration— and with the unusual character of the con¬
siderably larger $ being more brightly tinted than the £
502 ]\Tr. R. Trimen on Mr. Millars Ex'pcrinimtal
on the upperside of the wings — could not escape the
notice of collectors in Natal, if it were anywhere numerous
or frecpient in its appearance.
On examining with a lens the photographed series — 68
^ ^ and 55 $ ^ — of the offspring of the two ^ ^ dcccptor,
one is struck with the very slight amount of variation
exhibited by either sex. Tlie specimens are so crowded
in the photograph that the hind-marginal area of the hind-
wings is more or less concealed except in the last example
in each of the nine rows, and in the three next lower
examples in the fourth row ; but it happens to be on that
part of the upperside that most of what little variation
there is occurs, viz. as regards the development or sup¬
pression of the incomplete hind-marginal series of thin
white lunules, and sub-marginal series of 4-5 small white
spots — of which the former alone sometimes appear in
the while the latter accompany the former in many
examples of the $. This variation is well shown in the six
^ ^ and in six ^ ^ from the decc’ptor progeny forwarded to
me by Mr. Millar. This constancy to type in so consider¬
able a number of offspring from two mothers is opposed
to my long-held opinion that E. deceptor would prove to
be conspecific with E. ivalilhergi and E. minia. Moreover,
the full-grown larva of deceptor — as shown by the dried
specimens and numerous cast skins just before pupation
received from Mr. Millar — differs from those of the two
other forms in having the cephalic horns black instead of
yellowish-brown and abruptly clavate at the tip, and also
in having the pale-yellowish segmental elevated half-rings
much narrower and in places discontinuous between the
expanded bases of the spines springing from them. Mr.
Millar notes that these differences were constant in the
whole number of dcccptor larvae reared. It is, however,
worth recording that the first of these distinctions, viz.
the blackness of the cephalic horns, has been observed by
Mr. Millar to mark also an earlier stage of the larva of
'walilbergi and mima, both head and horns as well as the
body spines then being black. This is shown by a younger
inflated larva-skin sent to me by Mr. Millar.
Whether the living pupa of deceptor presents any notice¬
able difference from tliat of loahlhergi or mima I cannot
say ; Mr. Millar notes none ; and a dead example and six
skins of the former exhibit no distinctions that I can
detect from three pupa-skins of wahlbergi and three of
Breeding of the Nynvphaline Genus Euralia. 503
mima — all received with the imagines in Mr. Millar’s
consignment.
Second Experiment. — Offspring reared, from ova laid
hy one example of Euralia wahlbergi.
On the same date (March 21, 1909) as that on which
the two g ? deceptor were captured, Mr. Millar observed a $
wahlbergi ovipositing on and in the immediate proximity
of Fleurya plants. He at once followed up this example,
and carefully collected ten eggs laid by her, securing them
consecutively as they were deposited. All these eggs
hatched on the 25th ; and on the 31st the young larvae
were noted as being dark-blackish, with short black spines.
On April 5, after moulting, the larvae were quite black,
with on each segment, placed anteriorly, a very thin
ti'ansverse yellowish band or half-ring ; the spines, spring¬
ing from these half-rings, were more developed, but still
black ; and the two long divergent cephalic horns were
fully formed and also black. The moult of April 10 was
followed by a great change in the aspect of the larvae, the
segmental yellowish half-rings being wider and more con¬
spicuous, and the head and its horns, together with the
spines, the legs and the pro-legs, being all brownish-yellow.
On the 14th, five of these larvae pupated, and the remain¬
ing five on the l7th. One of the pupae died ; but all the
rest produced the imago in the following order, viz. : —
April 24 — %vahlhergi-ioxmy 1
„ 25 — mima-ioxm, S $
„ 26 — mima-iorm, 1 1
„ 27 — ^ucthlbergi-ioTxn, 1 1
,, 29 — wahlbergi-iorm, 1 [deformed].
Total offspring of a single example of the ^ of the
wahlbergi-ioxxn : — 4 wahlbergi (2 ^ 1 1 sex unde¬
termined), 5 mima (4 ^ 1 ^).
This extremely interesting result, clearly demonstrating
the species-identity of the forms wahlbergi and mima, is
the more striking because of the small number of the
brood being not far from equally divided between the two
forms, and at the same time including both sexes of each
form. Mr. J. H. Burn’s photograph (reproduced on Plate
LXII) gives in miniature all the examples of this brood,
with the exception of the deformed one — the sex of which
was not noted — and also dried specimens of two full-grown
504 Mr. II. Trimen on Mr. Millars Experimental
and one half-grown larvae, and of one dried pupa and
eight pupa-skins.
Third Experiment. — Offspring reared from ora laid by
one example of hiirralia mima.
In remarkable contrast to the result just described was
that obtained by Mr. Millar from the ova (39) laid, about"
March 26, 1909, by a single ^ of the mirna-iorm. captured
a day or two previously. These ova all hatched by April 3,
and the larvae presented the same appearance, and went
throimh the same changes after their moults as those
above noted in the case of the larvae hatched from ova
laid by a $ of the ^oahlbergi form ; Mr. Millar being
unable to detect any difference whatever between the
larvae of the two batches. With the exception of two
killed for specimens, the whole of the larvae from the
mima ova pupated on April 29, and of the 37 pupae 4
died. The emergences of the perfect insects were as
follows, viz. : —
May 6 — 3 $ $ — mima-iorm.
„ 7-2 do.
„ 8-8 ^ 2 ? ? do.
„ 9— 4 ^ 3 ? ? do.
„ 10-1 $ — do.
„ 11 - 2?? do.
„ 12-1 ^ — do.
„ 13—1 ^ — do.
14-3 ^ 1 ? do.
„ 17—1 f
Total, 24 ^ 9 ? mima-iorm..
The entire series (33) of this brood consisted of the
mother’s form {mima) only ; not a single example of either
sex — as may be seen from the collective view of them in
Mr. Burn’s photograph, reproduced in Plate LXIII —
showed any divergence towards or indication of the
characters proper to the ivahlbeopi-form..
In view of the almost equal emergences of both
waldbergi and mima in the brood from eleven ova of a
single wahlbergi, Mr. Millar exjaerienced surprise and dis¬
appointment at this unexpected result. On receipt of his
accidentally very long-delayed letter of July 6, describing
these breeding experiments, I wrote (December 31, 1909)
Breeding of the Ny'ni'phalinc Genus Euralia. 505
to express the hope that he would repeat, if possible, a
breeding from mima ova, as I thought it highly probable
a second result would prove to be quite different from the
first one ; but I learned subsequently that he had already
anticipated my suggestion.
Fourth Experiment. — Offspring reared from ova laid
hy a second example of Euralia mima.
On November 21, 1909, Mr. Millar met with a ^ mima
ovipositing at Mount Edgeumbe, and secured consecutively
eleven eggs one by one as soon as laid. These were care¬
fully placed on the food-plant and kept under regular
observation ; they all hatched, and the larvae duly fed up
and pupated. Without exception the pupae all yielded
the imago, the butterflies emerging in the following order,
viz. ; —
December 19 — mima4oxm, 1
„ 20 — mima-iorm, S $
„ 20 — wahlbergi-form, 2 $
„ 22— 7?ima-form, 1 3 ^
„ 22 — tvahlhergi-ioxm, 1
Total offspring of a single example of the ^ of the
mima-ioxxxi : 8 mima {b $ f 3 ^ ^), 3 icalilhergi $).
This second result of breeding from the ova (11) of a
single mima — in such strange contrast to the earlier one
in which exclusively mima was obtained from between
three and four times as numerous ova (39) of a single
example of the same form — conclusively confirms the
result of the converse experiment described above (p. 503),
in which 4 (2 ^ 1 1 sex undetermined) of the waldhergi-
form, and 5 (4 ^ 1 $) of the mima-ioxxa were produced
from 9 of the 10 eggs laid by one wahlbergi. It places
beyond question the fact so long suspected but until 1909
never proved, of the absolute species-identity of the two
forms under notice ; * and it incidentally illustrates the
* Evidence exists to show that my friend and correspondent, Mr.
William D. Gooch, who collected largely in Natal in the years
1873-78, and attended specially to the larvae and pupae of Lepi-
doptera, was close upon discovering the species identity of the two
Euraliae in question. As I mentioned in “ S.-Afr. Butt.,” I, p. 281
(1887), in Mr. Gooch’s extensive series of drawings and notes, there
are two pencil outlines and written details of a larva stated to have
resulted in “ Eiiralia mima or dubia,” and one of the figures of the
larva is represented on the food-plant, which is unmistakably a
stinging-netlle.
506 Mr. R. Trimen on Mr. Millar's Experimental
wisdom of not resting content with one test experiment
only, when its result runs counter to that of a previous
experiment in the same field. In this case, when the
numerous progeny of the mima taken in March 1909
turned out to be without exception of the same form in
both sexes as the mother, a very imperfect knowledge of
the actual state of things would have been gained if that
single experiment of breeding from ova of miina had been
accepted as conclusive in its result, and so remained
unchecked by repetition.*
As indicated briefly in my preliminary note (/. c., p. xv),
the case of E. wahlhcrgi-mima. agrees with that of another
Nymphaline butterfly in Natal — Gharaxes zoolina-ne-
anthcs,\ but at the same time differs very strikingly by
exhibiting in addition close mimicry of two Avholly dis¬
tinct and very differently marked and coloured species of
tlie protected genus Amauris, and thus presents the first
recorded instance of hi-sexual mimetic dimorphism occur¬
ring within the limits of one and the same species. In the
Gharaxes mentioned there is no approach to mimetism,
the colouring and marking in both the zoolina and neanthes
forms being inconspicuous — especially in the latter — and
on the underside cryptic, and this makes it difficult to
discern what the Gharaxes gains by its bi-sexual dimor¬
phism ; whereas in the Enralia the advantage of that
condition in combination with close mimicry of two such
dominant distasteful Danaines as Amauris niavius domini-
camis and A. alhimaeulata is manifest.
In considering the very differing proportion of the two
forms of Enralia in the respective progenies reared from
the wahlbergi mother and the two mima mothers by Mr.
Millar, it is well to bear in mind that in no one of the
three cases in question is it known to which form the
* Mr. Millar has presented the progeny resulting from this fourth
breeding experiment, viz. 8 mima and 3 wahlhergi from ova of a
single mima, to the unrivalled bionomic series in the Hope Depart¬
ment of the Oxford University Museum ; and Prof. Poulton has
most kindly contributed the excellent photograph of them by Mr.
Alfred Robinson, which is reproduced in Plate LXIV.
t See Mr. G. F. Leigh’s interesting accounts in Proc. Ent. Soc.
Loud., 1908, pp. lxiA% Ixv (with comments by Mr. G. A. K. Marshall,
Prol. Poulton, and Dr. K. Jordan), recording the breeding ol both
neanthes and zoolina from ova of a single neanthes, and in op. cit.,
1909, pp. xlix, 1, the converse breeding of both forms from ova of a
single zoolina.
Breeding of the Nymphaline Genus Eurcdia. 507
male parent belonged — pairing having ended before cap¬
ture or observation of the female. Moreover, it is seldom
indeed that all the ova normally laid are obtained ; a wild
female watched or captured, unless immediately or very
soon after pairing, having usually already oviposited to a
greater or less extent ; so that, in a dimorphic or poly¬
morphic offspring, the full relative numbers of the sexes
and of the respective forms born from one and the same
mother cannot be strictly ascertained. The possible pair¬
ings within the limits of the sub-species Euvalia wahl-
bergi-mima are four, viz. : —
1. ^ X $ loahlbergi-iovm.
2. ^ X ^ mima-loxva.
3. ^ ivahlhergi-iovvcv x $ mima-iorm.
4. ^ mima-iovm X $ unhlhergi-iovro..
Of pairing No. 1 (both sexes ot loahlhergi-iovra), Mr.
G. F. Leigh (Proc. Ent. Soc. Lond., 1906, pp. Iv and Ivi)
has recorded two cases that came under his personal
notice, viz. one on December 16, 1904, and the other on
January 23, 1905.
Of pairing No. 2 (both sexes of mma-form), there is
the instance of the very closely-similar sexes taken by the
late Col. N. Bowker (and published in my “ S.-Afr. Butt.,”
I, p. 285, 1887); and also Mr. Leigh’s record (/.r.) of
seeing two mima paired on January 27, 1905.
Of pairing No. 3. (J loahlbergi-iovvo. with $ mwia-ioxnii)^,
I have no definite instance ; but in the “ Entomologist
for 1882 (p. 9), Mr. A. J. Spiller, a keen collector and
observer, mentions that in Natal he had taken the ^
anthedon ( = luahlbergi) mated with the ^ mima, and vice
versa,.
Of pairing No. 4 (J mima-iovva with ^ wahlbergi-^oxwij,
there are tAvo authenticated cases, viz. Capt. H. C. Har¬
ford’s (published by me in Trans. Ent. Soc., 1873, p. 107,
note, and in “ S.-Afr. Butt.,” I, p. 283), and Mr. H. M.
Millar’s in July 1890.*
The occurrence ot certainly three, and most probably
* Mr. A. D. Millar, wlio communicated tins case to me at the time,
now writes : — I have seen initncL and wcthlber^i three times paired,
but did not note and cannot remember which form was the ^ and
which the 9 .” Mr. Millar sent me the 9 wahlbergi of the 1890
case, with the information that his brother, Mr. H. M. Millar, had
captured it paired with a d mima.
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) L L
508 Mr. R. Trimen on Mr. Millar s Experimental
all four, of these possible pairings in nature suggests the
idea that an important determining influence on the
discrepant results of rearing from ova to imagines the
offspring of a ^ of the wahlhergi-^oxm, and (at different
times, between four and five months apart) the offspring
of two ^ $ of the mima-iorxn, may perhaps be found in
similarity or dissimilarity in each case of the two parents ;
that the crossing of the forms may yield a progeny com¬
posed of the two different forms in varying proportions ;
while such a result as the thirty-three mima (24 $ ^ and
9 ^ exclusively from one mother of the mwia-form may
be traceable to the mating of similar parents.
At the same time, it should not be forgotten that, in
the case of PajMio dardanus sub.-sp. eenca, where the $ is
non-mimetic and, though varying considerably within
moderate limits, constant to one coloration and pattern,
the female offspring of two individuals of each form of
the trimorphic and tri-mimetic $ has been proved, by Mr.
G. F. Leigh’s breeding experiments in Natal,* to present
very great discrepancy in the proportional representation
of the three forms. In the two cases of the predominant
cenea-form of the ^ progeny reared was respectively 24
ce7iea and 3 hippocoon in one case, and 15 cenea and 1
hippocoon in the other ; in two cases of the rarer liippocoon-
form of $ the ^ progeny was re.spectively 8 cenea, 3
hippocooii, and 3 troplio^ims in one case, and 13 cenea only
in the other ; while, in two cases of the scarce trophonins-
form, the $ progeny was in one case 2 cenea only, and in
the other 6 cenea and 1 trophonius.
With all this discrepancy, however, there stands out — as
has been well demonstrated by Prof. Poulton {1. c., p. 430)
— the constant feature of the immense preponderance of
the cenea-iovm in these six results ; and with this may be
compared the preponderance of the mima-iorm (like the
cenea-^oxm of the Papilio itself a close mimic of Ainauris
alhmiacidatd) in the three cases of Euralia wahlhergi-^nmia
progeny reared by Mr. Millar.
Since I became aware of the occasional pairing of the
Natalian Euralia wahlbergi and mima, I have been sur¬
prised at not finding in collections any examples more or
less combining the features of the two forms ; indeed,
* See Prof. Ponlton’s “ Heredity in six families of Papilio Dar¬
danus, Brown, sub-species cenea, Stoll, bred at Durban, by Mr.
(j. F. Leigb, F.E.S.” (Trans. Ent. Soc. Lond., 1908, p. 429).
Breeding of the Nynvphaline Geoius Eitrcdia. 509
even an ordinary amount of variation seemed scarcely to
occur in either of them to any noticeable degree. The
only marked instances of variation I came across were
those recorded by me (as “ Var. A”) in “ S.-Afr. Butt.,” I,
p. 285 and footnote (1887) as of a ^ and a ^ of mima,
in which the enlarged white spots of the forewings and the
almost white merely yellowish-edged patch of the hind-
wings showed an approach in the direction not of wahl-
hergi, but of the West- African chibid, Palis., and of the
Madagascar E. drucei, Butl., considered by Aurivillius *
to be a variety of dulha. But Mr. A. D. Millar, having
lately written to me that he possessed an example uniting
the characters of wahlhergi and mima, I replied pointing
out the interest attaching to it ; and he has now forwarded
an excellent photograph by Mr. D. James (see Plate LXV)
of this intermediate individual — apparently a ^ — side by
side with bred examples of the loahlhergi and mima
forms. It will be seen that while in the forewings this
specimen in the main agrees as to its chief markings with
wahlhergi, the inner-marginal white patch is reduced by
its lower half being much clouded with fuscous scales (the
submedian nervure being broadly clouded with black); and
that in the hindwings the white area is greatly reduced
to the size and shape of that in mima. Besides this, in
both fore and hindwings appears the sub-marginal series of
rounded white spots so characteristic of mima, but never
present in \oahlhergi. Moreover, Mr. Millar points out
that while the blue iridescence characteristic of wahlhergi
is retained in the forewings, there is a slight tinge of
the ochrey-yellow of mima on the reduced white patch of
the hindwings. The rarity of examples partaking of the
characters of both mimetic forms of this sub-species of
* Anrivillius (Ehop. Aetliiop., p. 150, 1899) queries whether E.
diffusa, Butl., of Madagascar (figured by Mabille in Grandidier
Madag. Lep. I, pi. 18a, f. 4, 1885-7) be not a “hybrid” between
E. drucei and E. madaffascariensis,^La.h. (u form close to im/dtergi);
and also whether E. daemona, Stand., from Camaroon (“Iris,” iX,
pi. 2, f. 1, 1896) be not similarly a “hybrid” between E. dnhia and
E. anthedon. Of the latter case I can give no opinion, not having
seen Staudinger’s figure ; but Mabille’s figure of the Madagascar
diffusa does certainly much resemble the photograph of Mr. Millar’s
e.xample uniting the features of mima and wahlhergi — the forewings
being almost identical, but the hindwings ditfering conspicuously in
the whitish space being very much larger — scarcely less than in
madagascariensis or wahlhergi.
L h 2
510 Mr. R. Trimeu on Mr. Millar's Experimental
Euralia is in contrast to the comparative frequency of
linking intergrades among the polymorphic mimetic forms
of the ^ Papilio dardanus throughout all the sub-species
of that extraordinary butterfly ; and it is the more notice¬
able because in the cenea sub-species of P. dardanus,
inhabiting the same districts as the Euralia under notice,
the two more numerous mimetic forms shown by the ^
{cenea and liip)pocoon) simulate respectively the very same
species of Amauris — viz. A. alhimaculata and A. niamus
sub-sp. dominicanus — that are the models of the Euralia.
As already indicated above (p. 499) E. wahlbergi and
E. viima are very closely-allied south-eastern representa¬
tives of the western and equatorial E. anthedon and
E. diibia respectively, and there can be little if any doubt
that the latter will be proved, as the former have been, to
be two co-existing forms of one species, usually if not
always present in the offspring of a single mother of either
form. Valuable evidence in this direction has already
been furnished by three West-African examples — kindly
brought to my notice by Prof. Poulton in the collections of
the Hope Department of the Oxford Universit}’’ Museum
— intermediate between anthedon and duhia. The first of
these from the Lower Niger (W. A. Forbes) is much closer
to duhia than to anthedon, the median Avhite macular band
of the fore wings being situate as in duhia but much
enlarged and with its component spots confluent ; some of
the white spots of the submarginal series are absent. The
second, from the Gaboon (Mark L. Sykes, circa 1888) is
nearer anthedon, but in the forewings the large inner-
marginal pale space is obscured with dull violaceous-blue
except for a good-sized elongate white spot between 1st
and 2nd median nervules, while thesub-apical white bar is as
clearly defined and fully developed as in normal anthedon ;
and in the hindwings the white area is somewhat restricted
and externally tinged not with the usual blue gloss but
with ochrey-yellow (as sometimes in duhia) : in both wings
the submarginal white spots are only partly represented,
but those present though obscured with dusky scaling are
enlarged. The third, from Camaroon, is in pattern gener¬
ally like the second, but the colouring is very mixed : in
the forewings the sub-apical bar is interiorly much obscured
with black irroration ; the upper part of the inner-marginal
patch is larger than in the second example and of a purer
white, but all its lower part is clouded with obscure ochrey-
Breeding of the Nymjih aline Genus Euralia. 511
yellow ; while in the hindwings the pale area is of a clear
ochrey-yellow (as in E. mima) but radiates strongly out¬
wardly, more especially along the inner-marginal border.
The figures (uncoloured) of the larva and pupa of
E. dubia, var. cerberus, Auriv., from Camaroon, published
by Aurivillius (Ent. Tidskr., 1894, t 4, ff 4, 4a, and 4b)
are smaller than the full-grown specimens of those of
E. mima and E. waJdbergi that I have received from
Mr. Millar, but they and the brief description (p. 282)
agree pretty fairly with the specimens in question. As,
however, the account of arrangement of the larval spines
(which the author shows to be singularly identical with
that exhibited by the larva of Salamis anacardii, L.) is
not quite complete enough for comparison, I will add
here what I have found to be the spinous armature both
in E. %oahlbergi-mima and in E. dece'ptor larvae. Head
with a pair of two long, stout, rather clavate and expanded
(more distinctly so in dece'ptor'), divergent horns on vertex;
these horns are moderately spinulose and also sparsely
bristly, and bear three or four spinules at the tip. The
first thoracic segment bears dorsally, one on each side of
median line, two very short slender processes, each ter¬
minating in a long fine bristle, and also four rather short
rigid spines finely spinulose throughout and tapering to a
sharp point — two (upper and lower) on each side ; and the
second and third thoracic segments each bear in addition
(in place of the slender processes on the first segment) a
pair of longer dorsal spinulose spines. The abdominal
segments each bear an additional acute similar spine on
median dorsal line, making seven in all on each segment,
except the last, which — more or less injured in the inflated
dried specimens — appears to bear only five, but has the
medio-dorsal spine placed far forward of the rest, close to
the front edge of the segment (as Aurivillius mentions in
his description of the Salamis anacardii larva). Besides
the conspicuous spines above-named, there are on each
segment quite low down, at base of legs and pro-legs
where these occur, two very short blunt spinulose spines,
one in advance of the other, both pointing downward.
The colour of the larva in final moult is velvety-black ;
each segment bears anteriorly a narrow elevated or
thickened pale-yellowish transverse band or half-ring,
from which .spring the from four to seven brownish-yellow
spines. The head, legs, and pro-legs are all brownish-
512 Mr. R. Triiiieu on Mr. Millar'' s Exitcrimcnial
yellow ; on the head the horns on the vertex are of the
same colour, but the prominent simple eyes are conspicu¬
ously black with large white centres. This description
applies also to E. deceptor, except that in that form the
cephalic horns of the last stage are black, and the elevated
pale-yellowish segmental half-rings are less regular and
defined, being partly internipted atid discontinuous
between the expanded bases of the spines.
All naturalists will value the very interesting and im¬
portant contribution that has been made by Mr. Millar to
insect bionomics by his persevering and well-directed
experiments, and will join me in congratulating him, and
in wishing him continued success in prosecuting his
intended further researches into the life-history and
genetic conditions of Euralia, and also in extending them
to other kindred problems that await solution among the
Lepidoptera of Africa.
Explanation of the Plates.
Plate LX I.
Euralia deceptor, Trim. — Offspring (68 ^ ^ and 55 $ ^ ) hred —
2nd to ]4tli May, 1909— from eggs laid by two “small, apparently
dry-season ” $ 5 ) captured while ovipositing at Mount Edgcumbe,
near Durban on 21st March, 1909.
[From a photograph by Mr. J. H. Burn.]
Plate LXII.
Euralia wahlbergi, Wallengr. — Offspring (3 tcahlhergi-iorm
— 2 (J d , 1 ? ; and 5 mima-form — 4 $ $ , 1 $ ) bred — 24th to
29th April, 1909 — from eggs laid by a single $ wahlhercji captured
while ovipositing near Durban on 21st March, 1909.
[From a photograph by Mr. J. H. Burn.]
Plate LXIII.
Euralia mima, Trim. (= second form of A. Wallengr.)
— Offspring (24 ^ and 9 9 ?» ‘hi of true mima-fovm) bred —
6th to 17th May, 1909 — from eggs laid by a single 9 mima captured
near Durban about 2Gth March, 1909.
[From a photograph by Mr. J. II. Burn.]
Plate LXIV.
Euralia mima., Trim. ( = second form of E. wahlbergi, Wallengr.)
— Offspring (8 mfma-form— 5 S 6 3 wahlbergi-ionn
MR. A. D. MILLAR’S EXPERIMENTAL BREEDING FROM THE OVA OF THE NATALIAN FORMS OF EURALIA.
Offspring (68 S d and 55 ? ? ) from ova of two Euralia deceptor laid on 24th and 25th March, 1909.
Trans. Ent. Soc. Lo?id., igio, Plate LXIl.
MR. A. D. MILLAR’S EXPERIMENTAL BREEDING
FROM THE OVA OF THE NATALIAN FORMS
OF EURALIA.
Photo, J. H. Burn.
Offspring (3 Euralia ivahlbergi — 2 S S , i 9 ^ E. mima—
4 cJ (J , I ? ) from ova of one E. wahlbergi laid on 21st
March, 1909.
Trans. Ent. Soc. Land., igio, Plate LX III
MR. A.
D.
MILLARS
experimental breeding from the ova of the
NATALIAN FORMS OF EURALIA.
Phoio, J. H. Burn.
Offspring (24 i 4 , 9 ! 9 ) from ova of one Euralia mima laid about 26th March, .909.
Trans. Ent. Soc. Lond., igio, Elate LXIV.
MR A D MILLAR’S EXPERIMENTAL BREEDING FROM THE OVA OF THE
NATALIAN FORMS OF EURALIA.
C. Henfschci,
7to, Alfred Robinson.
All figures are about § of the natural size.
Three wahlbergi ( ^ ) and eight mima (5 , 3 ? ) bred in 1909 from a mima 9 ot
Euralia wahlbergi, near Durban, Natal.
Ltd.
Breeding of the Nymphaline Gemts Euralia. 513
— c? c?) bred — 19th to 22nd December, 1909 — from eggs laid by a
single $ mima captured near Durban on 21st November, 1909.
[From a photograph by Mr. Alfred Eobinson, kindly contributed
by Prof. E. B. Poulton.]
Plate LXV.
Evralia wahlbergi, Wallengr. — Fig. 1, ^ of wahlbergi-lovm,
fig. 2, of mima-form ; both examples belonging to the offspring
from eggs laid by a single wahlbergi included in Plate LXII. These
are here given for comparison with fig. 3, representing a remarkable
example which combines the characters of both forms. The latter
example, in Mr. A. D. Millar’s collection, is without special record,
but was taken near Durban.
[From a photograph by Mr. D, James.]
( 514 )
XVIII. On a neio Genus of Stylopidae fo'om Australia.
By Arthur M. Lea, F.E.S,, Government Ento¬
mologist, Tasmania.
[Read October 5th, 1910.]
Plate LXVI.
In 1895, whilst on a collecting trip to Bridgetown, in
W. Australia, I obtained a number of specimens of a species
of Stylopidae in the greasy oil of a lamp, massed together
with other insects that had been attracted by its light.*
They were naturally somewhat damaged ; to remove the oil
they were placed for some time in chloroform, afterwards
being mounted on cardboard. Not having the necessary
literature to work them up, they were left untouched till
recently, when Mr. W. Dwight Pierce’s fine monograph of
the family was received, this enabling me to at last deal
with them.
Stylopidae have previously been recorded from Austra¬
lia (N.S. Wales and Queensland) in a rather out-of-the-
way publication so far as Australian workers are concerned.f
But all the species there noted belong to different genera
(sub-families according to Pierce) to the one now under
consideration, as they have the tarsi three or four jointed
and without terminal claws. Mr. Aug. Simson also informs
me that he sent a female in the abdomen of a Tasmanian
wasp to Mr. Saunders, but nothing appears to have been
published regarding it. Westwood mentions that “R. H.
Lewis informs me in a recent letter that he has captured
a Stylopized bee in Van Diemen’s Land.”
Austrostylops, n. g.
Head strongly transverse. Eyes coarsely facetted, occupying about
two-thirds the width of the head. Antennae six-jointed, third,
fourth and fifth joints ramose, the sixth elongate. Pronotum tri-
lobed, the median lobe somewhat depressed and produced on to head.
* I have seen no reference to other species being attracted to light ;
and, in fact, the males of most species appear to emerge in daylight,
and to die before sunset.
I Report of Work of the Experiment Station of the Hawaiian
Sugar Planters’ Association, Bulletin No. 1, Part 3, Honolulu, 1905.
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.)
Mr. A. M. Lea o% Genus of Stylopidae from Australia, 515
Mesonotum very narrow in middle, the sides suddenly dilated.
Scutellum transversely suboblong. Metanotum in about four equal
parts, of which three are placed transversely, the two outer ones sub-
triangular with the narrow part apical, the median part moderately
narrow, but not. pointed in front, and evenly emarginate behind ;
hind portion separated from the median one by a hollow membranous
space, the front angles acute and hind margin rounded. Abdomen
apparently composed of six segments. Elytra about once and one-
fourth as long as metanotum is wide. Wings large. Legs long
and thin ; tarsi five-jointed, with two thin, equal claws.
The tarsi are distinctly five-jointed, and with two claws, characters
which, according to Pierce’s table (page 70 of his monograph), would
place it in the Mengeuidea ; to which accordingly I refer it. Only
two genera (one fossil) are placed in the sub-family by Pierce, and
both have the antennae seven-jointed. In the Halictophagidue the
antennae at first appear to be similar, but they have one joint more
to the flabellum, and their tarsi are three-jointed only and without
claws.
The buccal cavity is irregularly rounded, and on one specimen
appears to have five processes, one on the upper part, round and
hollow ; and four in pairs, of which the upper ones are triangular in
shape, wide at the base, and pointed at the tip, and not quite touching
each other ; the lower ones are narrower and less triangular, with the
apices more obtuse and also not meeting. On another specimen two
long processes cross each other over the mouth, and two somewhat
shorter ones can be seen. These probably are broken off on the first-
mentioned specimen, allowing parts of the mouth to be more clearly
visible.
The abdomen is so badly twisted on all the specimens that it is
impossible to state what it is like on fresh ones.
Austrostylops GRACILIPES, n. sp.
. Of a pale but somewhat dingy flavous ; antennae, elytra and
wings lightly infuscated. Extremely finely pubescent.
Head notched in front and ridged close to each eye. Antennae
with first joint of moderate size, second short, third to fifth short,
but each with a long ramus, sixth elongate, not much shorter than
rami. Elytra rather narrow at base, with a thin strip membranous,
apical half wider. Wings with about seven (including the outer
one) long veins, and two short ones. Legs long and thin; front
coxae femora and tibiae of about equal lengths, but the coxae slightly
stouter than femora, and these than tibiae ; trochanters about one-
third the length of femora, to which they are closely applied ;
olG Mr. A. M. Lea on Genus of Styloindae from Australia.
first joint of tarsi about once and one-half the length of second,
and about the width of tibiae, second joint much thinner, third,
fourth and fifth slightly decreasing in length; middle legs appar¬
ently much as front ones ; hind coxae obliquely placed, about four
times as wide as long, touching in middle, femora, tibiae and tarsi
rather longer than in front legs, but otherwise much the same. Length
2 1 mm., expanse across wings 6 mm.
Hah. W. Australia, Bridgetown.
Explanation of Plate LXVI.
Austrostylops gracilipes, Lea.
Trans. Ent. Soc. Lend., ipio, Plate LX VI.
( 517 )
XIX. Hymenoptera Aculeata collected in Algeria hy the
Rev. Alfred Edwin Eaton, M.A., F.E.S., and
the Rev. Francis David Morice, M.A., F.E.S.
Part IV. Descriptions of new Sphegidae. By the
late Edward Saunders, F.R.S.
[Read March 2nd, 1910.]
[The following descriptions were intended for incorpora¬
tion in what would have been the concluding part
(dealing with the Sphegidae) of a Paper commenced
under the above title in these Transactions in 1901 and
continued in 1905 and 1908.
The author had before and during his last illness made
some progress with other portions of the intended work ;
but these are in too incomplete and fragmentary a state
to be published as they were left. The descriptions,
however, though evidently a mere rough draft which the
author would have revised and copied afresh before
sending it to press, do not seem to require any sub¬
stantial alteration in order to make them useful to
Hymenopterists. They are accordingly reproduced prac¬
tically as written, only a few clerical and grammatical
slips (due to haste) have been silently emended, and
here and there a blank space left by the writer for
localities, dates, measurements, etc,, to be added later,
has been filled by an insertion for which the author is
not to be considered responsible, such insertions being
always indicated by the employment of square brackets [ ].
Mr, Eaton presented all his captures to Mr. Saunders,
and they are now — except a few given or exchanged away
by the latter in his lifetime as duplicates — in the Natural
History Museum at South Kensington.
The other materials gathered by Mr. Saunders for his
paper have been handed over to a colleague whom he
had invited to co-operate in preparing it, and it is hoped
that in some form or other at least their main substance
may be made available to Hymenopterists by communi¬
cation to the Society before the end of the present
year.]
TRANS. ENT. SOC, LOND. 1910. — PART IV. (DEC.)
518 Mr. E. Saunders on Hyimnoptera Acvleata:
1. Sphex [Haiipactopus] eatoni, n. sp.
Niger, capite pro- et mesonoto (scutello excepto) pilis adpressis
argenteis dense vestitis. Ad S. higentem Kolil proxime accedit, is
tamen rugis validis mesonoti facile distinguitur. Etiam S. tyranno
Smith colore similis, is autem differt pedibus setis rigidis instriictis.
Ab S. stschurowskii Rad. propter scutellum rugosum etc. mox
distinguendus.
Black clothed with black hairs. 9 with the face (except a
bright glabrous spot at the apex of the clypeus), the vertex, the
pro- and mesonotum (excluding the scutellum), and the base of the
tegulae covered with a dense silvery pubescence which entirely
hides the sculpture beneath. ^ resembling the $ but with only
the face and mesonotum silvery. Clypeus covered with somewhat
remote black bristles which project beyond the silvery surface, its
apical margin rounded, raised and slightly sinuate in the centre,
more truncate in the ^ than in the 5 — ^just above the apex is a
shining shallow impression. IMandibles in the § large, the apical
tooth very elongate, more so than in stschurowskii var. hyalipennis.
Kohl. Head with black hairs in the ^ , the cheeks behind the eyes
with a long beard. (In the ^ tbe only long black hairs are on the
cheeks beneath, the mandibles, clypeus, and labrum ; a few very
short ones show through the silvery hairs of the vertex and back
of the head.) The pronotum propleurae and prosternum are
obliquely striate where the surface is not hidden by the silvery
pubescence (as it is, partly, in the $ ). jMesopleurae and sternum
rugose above, the rugosities [ ? of the former] taking the form of
striae as they approach the sternum in the $ ; entirely rugose (as
well as the sternum) in the ^ . Tegulae clothed at the base with
silvery hairs in the 5? with greyish-black ones in the
Wings nearly black, with violet reflections : the hind-wings in
the $ clear only beyond the cells ; the 2nd submarginal cell
narrowed to its apex where it is scarcely more than half as wide
as its base ; the 3rd a little less narrowed towards its apex than
is the 2nd in the $, narrowed in about the same proportion as
the 2nd in the ^ ; (the bases of the two cells are of about equal
length).
Scutellum moderately convex, rugose with signs of longitudinal
striae ; postscutellum transversely strigose. Propodeum with a penta¬
gonal area which is clothed with rather short semi-decumbent
hairs and rugose in both sexes— its sides are rugosely diagonally
strigose, and in the ^ the rugosities towards the base of the abdomen
are larger, and tend to become clathrate. In one of my 5 ^ the
area tends to be rufescent in certain lights. Legs entirelj^ black,
Descri'ptions of Neio Sphegidae. 519
9 witli the intermediate coxae trochanters and femora in front,
and the hind coxae, clothed with silvery pubescence : the front
metatarsi externally bear a comb of eight fine long curved bristly
spines with some shorter ones between them, on the side towards
the body are three rather thicker spines and a dense brush of very
short dark hairs. Front coxae ( 5 ) transversely strigose with their
central portion polished and punctured, femora shining and
punctured. In the ^ there is no sign of silvery pubescence, and
the anterior coxae are rugose and dull. The tibiae in both sexes
are simply spinose, not densely bristly as in tyranni(s, etc. Posterior
metatarsi slightly curved at the extreme base in both sexes.
Abdomen polished, its petiole slightly curved (its ventral length
subeqi;al to that of the hind coxae in the 5 > longer in the ^ ).
The rest of the body short and oval in the 9 1 longer propor¬
tionally in the i^. Long. 28-32 mm.
This fine species is allied to higms, Kohl. But of his
species Kohl observes that the mesonotum is so rugose
that the rugosities show through the silvery hairs. In
this species the silvery pubescence lies so flat on the
surface, that its rugosities, if any, must quite clearly be
very slight.
Biskra, v, 97. (Coll. Saunders.)
2. Astatus fumipennis [n. sp.].
Caput nigrum, nitidum, facie et genis postice albo-hirsutis.
Thorax nigernitidissimus,remotissimepunctatus,pleurisalbo-hirsutis;
alae anticae fuscae, posticae dilutiores. Abdomen rufo-testaceum,
valvulae dorsalis lateribus valde reflexis. [ 9 ]
Head and tiiorax shining black, mandibles red towards the
base; face and head posteriorly and thorax beneath clothed with white
hairs. Antennae with the 2nd joint of the flagellum 3 times at
least as long as the 1st and distinctly longer than the 3rd : the
scape clothed outwardly with white hairs, inwardly with thick
black bristles, 1st and 2nd joints of the flagellixm clothed with
stiff black hair.s. Vertex and region above the ocelli shining, largely
and remotely punctured, that between the ocelli and below them
closely punctured. Mesonotum (including the scutellum) very shiny
and very remotely punctured ; mesopleurae punctured. Anterior wings
dark smoky brown : radial cell rather more than twice as long as
wide, its apex truncate ; posterior wings clearer ; both pairs darkened
towards the apex. Legs black, clothed with black bristly hairs ;
the anterior tarsi pitchy-testaceous, the others more or less
520 Mr. E. Saunders on Hynienoptcra Acv.lmta :
piceoiis. Propodeuni finely reticulate above, its sides clothed
■with white hairs and diagonally stiigose. Abdomen entirely red ;
apical dorsal valve with the sides strongly reflexed, and with
black decumbent spine-like hairs as in hoops, etc.
Long. 12 mill.
A very distinct-looking species, but unfortunately only
a single $ was captured.
[Biskra, 25. v, 97. A. E. E. Type in Coll. Saunders.]
3. Astatus laetus, n. sp.
9 . Nitidus, niger; palpis,mandibularum basibus, tegulis,abdomine
toto pedibus que (coxis trochanteribus que exceptis), ferrugineo-
riifis ; alis plus minus infuscatis, cellula radiali brevi, latitudine
sua vix duplo longiore. Capitis vertex remote, facies dense
punctata. Thorax nitidissimus, mesopleuris crebre punctatis et
strigosis, propodei lateribns clatlirato-rugosis.
9 . Head and thorax black and very shining. Palpi and base of
the mandibles red. Antennae with the second joint of the flagellum
about three times as long as the first. Vertex and region above the
ocelli very remotely punctured ; face below them dull and closely
punctured, clothed sparingly at the sides with whitish hairs.
Cheeks behind the eyes also thinly clothed with white hairs.
Mesonotum including the scutellum very shining, and exceedingly
remotely punctured. Mesopleurae closely punctured and diagonally
strigose. Wings slightly smoky, darker towards the apex ; radial
cell very short, scarcely twice as long as wide, its apex truncate.
Tegulae red. Legs with the exception of the coxae and trochanters
entirely clear red; the anterior femora clothed with long whitish
hairs, intermediate and hind femora with brownish-black, tibiae and
tarsi with black S2)ines. Propodeum finely reticulate above, its sides
coarsely clathrate posteriorly, with a rather narrow almost smooth
(but very finely punctured) region anteriorly. Abdomen entirely
red, its apical valve with a very slightly raised margin, and with
adpressed pale spine-like hairs.
Long. 10 min.
A very distinct species, its bright red legs and very
bright thorax and head giving it a very peculiar appear¬
ance. From rtifipes it differs in the absence of black hairs
on the frons, the closely-punctured face, and the sculpture
of the sides of the propodeum.
[Biskra, 19. V, 97. A. E. E. Type in Coll. Saunders.]
Descri'ptions of New S2Jhegidae.
521
4. Astatus radialts, n. sp.
$ . Niger ; mandibularuni basibus, abdoniine toto, parte femoruni,
tibiis tarsis que rufis ; antennarmu articulo tertio secundo plus
quam duplo longiore ; eapite et thorace griseo hirtulis, crebre
punctatis ; alis subfumatis, apicibus harum valde infuscatis, cellula
radiali brevissima, latitudine sua vix duplo longiore, apice abrupte
truncate, cellula tertia submarginali angustissima ; ceteris ut in
boope.
5 . Easily distinguished from hoops or minor by the red legs, and
from rufipes by the short radial cell, as well as by the rather
unusually dense grey pubescence of the head and thorax. The
dark band of the wings also gives it an unusual appearance, this
being more pronounced than in the other species and not quite
reaching to the apex of the wing. The very narrow third sub-
marginal cell is another characteristic, but this varies slightly in my
examples. In the short radial cell the species approaches costae,
but of that species (which I do not know) Kohl says that the wings
are “ wasserhell,” and my specimens seem [too (?)] large for that
species, being 7^ mm. in length. The sculpture of the propodeum
is rather finer than in 6oops and minor.
Long. 8 mm.
$ (?). I have a single male, caught at Le Tarf, on the
same day and on the same flower {Foenicitlum vulgare) as
the female, which probably belongs to the same species.
It has the radial cell shorter and more truncate than iti
hoo2)S, and the third submargiual with a short nerve
emitted from about its centre (this is identical in both
wings, but may not be a constant character). The
antennae have the intermediate joints simple, not sinuate
as in hoo2)s, etc. It is, however, larger than one would
have expected the ^ of this species to be, viz. 10 mm.
But the species of Astahis are very variable in this
respect.
[A $ and 5 ^ $ answering to this description are placed
together (unnamed) in Coll. Saunders. Two are from
Bone, the rest from Le Tarf, 24. and 26. vii, 96. A. E. E.]
5. TaCHYTES BISKREx\SIS, n. sp.
d 9 • T. freygessneri, Kohl, valde affinis, differt tameu, pedibus
maxima parte rufo-testaceis; eapite et thorace pilis argenteis dense
vestitis ; alis hyalinis ; abdomine toto rufo-testaceo ; 9 segmentis
abdominalibus fascia lata apicali argeutea ornatis.
522 Mr. E. Saunders on Hymenoptcm AcuUata :
Very closely allied to freyf/essneri, Kolil, with which it seems
to agree in nearly all structural characters except that the vertex is
slightly wider in proportion : but in colour and ]mbescence it
differs so strikingly that I am describing it as new.
^ 9 • Head and scape of the antennae entirely clothed with
shining silvery hairs. Mandibles more or less testaceous-red,
clothed with silvery hairs at the base ; vertex between the eyes
about as wide as the second and half the first joint of the flagellum
taken together. Thorax and propodeum entirely clothed with
silvery hairs (these are more brilliant in the 9 than in the d)-
The disc of the mesonoturn in my only 9 is more or less rubbed,
and the surface is dull and exceedingly closely punctured ; wings
very clear and hyaline, tegulae and nervures very pale testaceous.
Legs in the black, with all the tarsi, the entire posterior tibiae,
the intermediate and anterior tibiae, i^osteriorly testaceous, anteriorly
piceous ; in the 9 S'!! the legs are clear testaceous and clothed with
silvery hair.s, the front femora anteriorly and the front and inter¬
mediate coxae black ; abdomen clear testaceous. Each segment in
the 9 bears a wide aj^ical band of silvery hairs covering nearly half
of the 1st and 2nd segments, | of the 3rd and the whole of the 4th
and 5th ; the apical dorsal valve is clothed with slightly golden
hairs; in the the disposition of the hairs is very similar, but they
are less vividly silvery, and the bands are narrower, the hairs of the
5th and 6th segments are golden, and the dorsal valve is silvery.
Beneath shining and glabrous in the 9 > iu tlie S sparsely clothed
with adpressed golden hairs.
Long. 12-13 mm.
[Biskra, 22. v, 27. A. E. E. Types ^ and 9 Coll.
Saunders.]
6. Tachytes aenea, n. sp.
(J. Aeneus argenteo-aureopilosus, tegulis tarsis que testaceis,
alarum nervis piceis, thorace pilis erectis densissime vestito, pro-
podeo linea central! nitida impres.so, abdomine pubescentia argentea-
aurea dense vestito.
[Here is added in the margin a pencil annotation “ changing in
different lights.”]
(J. Bronzy, densely clothed with golden-greyish hairs, tarsi tes¬
taceous, vertex between the eyes about as wide as the 2nd and 3rd
joints of the flagellum together, antennae rather short, joints of
the flagellum not twice as long as wide, face so densely covered
with hairs that the sculpture is invisible— these hairs in certain
523
Descriptions of Nevj Sphegidae
lights are bright silvery with a slight golden tinge. Vertex very
closely and exceedingly finely punctured. Thorax very densely
clothed all over with erect hairs, its sculpture hidden except on
the mesopleurae, which are rugosely punctured. Wings slightly
dusky, nervures piceous, tegulae pale testaceous, front metatarsus
with four pale short spines along its outer edge, and two at its apex,
tibiae with a few feeble spines, calcaria pale. Propodeum with a
smooth shining impressed central line ; punctured above and at
the sides, but so densely clothed with hairs that the sculpture is
mostly hidden. Abdomen clothed with adpressed sericeous pube¬
scence, in certain (lateral) lights forming wide apical silvery-golden
bands, viewed from in front and in other lights, apparently covered
entirely with silvery-golden hairs, each change of position causing
a difference in the apparent arrangement of lighter and darker
hairs, apical valve rufescent. Segments beneath shining, black,
punctured, their apices testaceous.
Long. 6-9 mm.
[Biskra, 25. iv, 97. A. E. E. Coll. Saunders.]
7. Tachytes maculicornis, n. sp.
Nigra argenteo-pubescens, mandibulis piceis, palpis alarumque
nervis laete testaceis, tegulis pallidis, valvula anali pilis argenteis
tecta. Maris antennae articulis 4-6 dilatatis et pallidis, 7-8
pallido-notatis : metatarsi intermedii apice valde spinoso producti.
^ 9 • Black clothed with silvery pubescence ; mandibles dai’k
piceous ; palpi tarsi and wing-nervures bright testaceous ; tegulae
pale. Antennae in the ^ with joints 3 to 8 of the flagellum beneath
each with a conspicuous pale ochreous spot (that of the 8th only
occupying its extreme base). Apical dorsal valve in both sexes
clothed with silvery hairs.
This species agrees closely with freygessneri in the
proportions of the vertex and antennae, and in the $ is
hardly to be distinguished from the latter except by its very
different coloration and more abundant silvery pubescence.
The ^ however presents most unusual characters. Its
antennae have joints 4, 5, 6 of the flagellum dilated — the
dilatation being widest at the base of each joint, which
therefore projects beyond the apex of the preceding one.
These three dilated joints are almost entirely pale ochreous
beneath, their apices only being narrowly black. The
third joint, though not dilated, is a trifle thickened and
pale beneath. Joints 7 and 8 are noi'inal except for their
TRANS. ENT. SOC. LOND. 1910. — PART IV. (DEC.) M M
524 Mr. E. Sai^lers on Hymcnoptcra Acidcata :
pale spots. Another striking character is the form of the
intermediate metatarsus. This is slightly bent and pro¬
duced at the apex anteriorly into a spine-like process
which bears at its extremity a short white spine. The
posterior metatarsi also are slightly bent at the base. In
disposition of pubescence this $ closely resembles its
[Biskra. Several and A. E. E. and F. D. M.] •
8. TaCHYSPHEX PHILIPPI, n. sp.
5 T. nitido affinis : puncturatione minutiore, tibiis anticis antice
palliclis tegulis que testaceis distinctus.
9 Black, clothed with a very short, inconspicuous grey pubes¬
cence ; front tibiae on their inner side, and the apical joints of all
the tarsi, testaceous ; mandibles piceous-red near the base, tegulae
testaceous. Head and thorax slightly shining ; the former (with
the face) closely punctured on a finely rugulose surface ; the latter
finely and rather closely punctured on an almost smooth surface.
Vertex between the eyes about equal to the 1st and 2nd joints of the
tlagelluin together ; wings slightly dusky ; mesopleurae punctured
on a slightly and very finely rugulose surface ; propodeum rather
unusually finely rugose, its area indefinite, its posterior declivity
transversely strigose, and its sides diagonally so. Comb of anterior
metatarsus composed of thin testaceous bristles, abdomen dull,
exceedingly finely and microscopically rugulose, without evident
puncturation. Segments 1, 2 and .3 with a lateral apical band of
silvery pubescence [visible] only in certain positions. Apical
dorsal valve elongate (as in iKctinipes., etc.) and irregularly
punctured.
Long. 8 mm.
Easily separated from nitidus, etc., by its fine punctura¬
tion, (inwardly) pale front tibiae, and testaceous tegulae,
combined with the elongate form of the dorsal valve
(this last character distinguishing it at once from the black
form of lativalvis). The specimen bears a ticket in F, F.
Kohl’s handwriting “ sp. ? a T’. nitido diversa.”
[1 9 Philippeville, 21. vi, 98. F. D. M. Type in Coll.
F. D. Morice.]
9. Tachysphex piagetioides, n. sp.
[Mr. Saunders appears to have described this species in
English only, leaving a blank space for a Latin diagnosis
to be added afterwards.]
525
DescH^ptions of Neiv Sphegidae.
9 Black, shining, in form something like a Crahro of the Coelo-
crahro group, mandibles towards the apex, tegulae and wing veins,
apices of the femora, and the tibiae (except a dark stain on their
inner side) testaceous. Head punctured. Width of vertex between
the eyes equal to the combined length of the first four joints of the
flagellum ; orbits parallel sided ; the face on each side more or less
clothed w’ith silvery hairs ; clypeus finely punctured, its centre
produced into a short angular process wdiich is carinated on the
centre, and slightly raised on its margins. Antennae with the third
joint of the flagellum considei’ably longer than the second and
slightly longer than the fourth. Mesonotum (including the scutel-
lum) shiny, clearly and rather remotely punctured, clothed with
inconspicuous greyish hairs. Wings hyaline, their nervures pale
testaceous, radial cell truncate at the apex, third submarginal
shortly ligulate. Mesopleurae transversely rugose, and sparingly
clothed with silvery hairs ; propodeum without a definite dorsal area,
transversely rugose, its posterior declivity with a well-marked central
fovea, the sides diagonally strigose and clothed with silvery hairs.
Legs w’ithi the tibiae spinose, front metatarsi with a comb of six
moderately long pale spines ; abdomen shining, with scarcely per¬
ceptible sculpture, the apices of the segments narrowly discoloured
(the discoloration widening along the sides of the segments) ; basal
segment about as long as wide ; dorsal area of apical segment very
largely punctured, its margins raised and testaceous. Beneath
shining.
Long, [about 7 mm.].
This insect was submitted by Mr. Morice to Herr F. F.
Kohl some years ago, and bears a label in the latter’s hand¬
writing. “ Tachysphex n. sp. transiens ad rarapiagetia,
Kohl.” Had it not been for this I should scarcely have
dared to place it in this genus.
[Biskra. F. D. M. Type in Coll. F. D. Morice.]
10. Tachysphex eatoni, n. sp.
9 Niger. Pro- meso- et metanota nitidissima, remotissime
punctata ; mesopleurae opacae, minute et crebre punctatae ; alae
hyalinae ; abdomen nitidum sed non politum, segmentis basalibus
minute punctatis, valvula dorsali elongata.
9 Black ; mandibles reddish except at the base ; tegulae piceous.
Vertex of head shining, finely and remotely punctured ; face more
or less dull,’’punctured closely and more deeply [than the vertex ?];
clypeus narrowly impressed at the apex, and widely truncate;
52(j ]\Ir. E. Saui^rs on Hymeno'ptera Aculeata:
vertex between the eyes scarcely so wide as the second and third
flagellar joints taken together ; pro- meso- and metanota very shining
and polished, with only a few very scattered punctures; ineso-
pleurae dull, exceedingly closely and finely punctured ; wings nearly
hyaline; propodeum above dull, finely and longitudinally rugulose,
clothed with very short and inconspicuous greyish hairs, its sides
diagonally striate ; anterior tarsi with a comb of slender and not
very long brownish hairs. Abdomen shining, but not polished like
the thorax ; finely punctured on the basal segments, those towards
the apex being almost impunctate ; apical dorsal valve polished and
elongate, its shape much as that of pectinipes.
Long. 7 mm.
The polished, almost impunctate, mesonotum, etc., of
this species will distinguish it at a glance.
[Biskra, 5. iv, 97. A. E. E. Type in Coll. Saunders.]
11. Tachysphex capitalis, n. sp.
Species insignis ; nigei’, mandibulis,tegulis, abdomine pedibusque
testaceis. Caput maximum, vertice latissimo, articulis 1, 2, 3, 4
antennarum flagelli simul sumptis aequali ; pronotum oblique-
strigosum ; propodeum transverse-striatum; abdomen impunctatum,
valvula dorsali lata punctis remotis impressa.
9 Head and thorax black, clothed with silvery white hairs.
Mandibles, tegulae, wing-veins, legs, and abdomen testaceous, wings
with a yellowish tinge. Head unusually produced posteriorly (from
a sideways view the cheeks behind the eyes are as wide as the eyes
themselves) ; vertex between the eyes exceedingly wide and convex
(as wide or even wider than the fir-st four joints of the flagellum).
This great width gives the face a most unusual appearance for that
of a species of this genus. Clypeus flat, its apex slightly elevated,
shining and bituberculate. Antennae with the second joint of the
flagellum nearly once and a half as long as the third. Pronotum
diagonally strigose ; mesonotum and mesopleurae thickly clothed
with silvery hairs ; propodeum clothed with long silvery hair.^,
transversely rugose above, diagonally so at the sides. Legs rather
thick ; front metatarsi with seven rather stout jjale spines. Abdomen
almost smooth, and without evident puncturation. The sides of
the second and following segments are clothed with silvery pubes¬
cence, visible only in certain lights, meeting almost in the centre
of the segment ; apical dorsal valve widely triangular, and with
large scattered punctures.
Long. 13 mm.
9 1. Biskra, 11. vi, 98. F. D. M.
[Type ill Coll. F. I). Morice.]
527
Descriptions of Neto Sphegidae.
12. Tachysphex [gracilitarsis], n. sp.*
[* In the original MS. a blank space is left for a
specific name to be afterwards inserted. The name ofi’ered
above is founded on the character by which the author
distinguishes his species from panzeri, var. oo'ciniensis.']
$ . T. panzeri var. oraniensi simillima. Tarsis posticis gracilio-
ribus horuin articulo quarto vix usque ad medium exciso.
5 . Very closely allied to large specimen.s of T. panzeri var.
oraniensis, and in colour, hairing and general appearance hard to
separate from the latter. The following structural character will,
however, easily distinguish it. The fourth joint of the posterior
tarsi is divided in the middle not quite to its centre, and its longer
apical spine (regarded laterally) does not extend to the apex of the
fifth joint. (In panzeri the fourth joint is divided up far beyond the
middle, almost to the base; and the longer apical spine, regarded
laterally, extends quite to the apex of the fifth joint.) In fact the
tarsi are altogether thinner and less strongly spined than they are in
panzeri, and the claws are less developed. Another good character,
though not quite so easy to appreciate, lies in the form of the dorsal
area of the sixth segment. In this species it is distinctly shorter and
wider than in panzeri, and more polished.
Long, [about 13 mm.].
Biskra, F. D. M.
[4 $ one of which must have been the “ type,” are in
Coll. F. D. Morice.]
1.3. Tachysphex [eduardi, n. sp.].
[In describing this species the author has again left a
blank for the subsequent insertion of a specific name:
and to fill this void the name eduardi is now suggested.]
Panzeri et pygidiali affinis, sedalis anterioribus fuscis sat distinctus ;
alae posteriores multo dilutiores. Tarsi antici ( d ) spinis brevibus
( 5 ) spinis longis curvatis nigris armati.
d . Allied to panzeri and pygidialis in the finely rugulose sides of
the propodeum, but at once distinguished by the dark anterior wings.
Head and thorax black ; face clothed with golden pubescence between
the ocelli and the antennae, with silvery on the sides of the face below
them and on the clypeus. Clypeus very convex longitudinally, its
anterior portion shining, strongly punctured, rounded in front.
Width of vertex between the eyes scarcely equal to the length of the
first and second joints of the flagellum taken together. Mandibles
52(S Mr. E. Saunders on irymcnopiera Aculeata:
reddish in the middle. The antennae appear to be rather thicker
and longer than in pyyklialis. Mesonotnm finely and very closely
punctured, clothed with pale decunibent .scarcely conspicuous pube¬
scence which gives it in certain lights a greyish appearance. Anterior
wings strongly tinged with smoky brown, posterior pair much
clearer ; tegulae testaceous. Propodeum very finely rugulose both
above and at the sides, clothed with a fine and very short grey
pubescence. Legs black with the exception of the tarsi which are
more or less piceous ; front tarsi without any regular comb ; the
spines of the metatarsi not longer than the middle of the joint itself.
Abdomen dull, exceedingly finely punctured ; the first three segments
red, the rest black.
$ (1). A single 5 fi’om Bone agreeing in sculpture, colour of wings,
etc., with the above, is closely allied to pygidialis, but has a slightly
narrower dorsal valve ; its front tarsi have a comb of long dark
curved spines, and the .sides of the propodeum have distinct striae
.amongst the rugulosities.
Long. 11 mm.
Had this female occurred at Biskra with the males I
should have had no doubt of its being the other sex of the
same species ; but coming as it does from Bone, a much
more northern locality, I feel it is wise to suggest the
possibility of its being distinct.
[Types in Coll. Saunders.]
14. Prosopigastra insignis [n. sp.].
$ . Species insignis et omnium generis huius maxima. Caput
nigrum vertice crebre punctato, tuberculo faciali magno (nitidissimo
et punctato), mandibulis, clypeique apice testaceis. Thorax niger,
opacus, crebre punctatus (disco nitido, remote-punctato, excepto).
Abdomen testaceum, segnientis 1-4 crebre, 5 parce punctatis, valvula
dorsali nitida, remote sed fortiter punctata, apice ipso truncato
Subtus, segmento secundo crebre, reliquis remote punctatis.
$ . Head and thorax black ; mandibles, apex of clypeus, tegulae and
veins of the wings, legs, and abdomen testaceous. Head dull, closely
punctured on the vertex ; face below the ocelli clothed with silvery
hairs, facial tubercle large, polished, punctured ; clypeus produced
anteriorly into a semicircular testaceous plate, which is somewhat
elevated along its edge ; cheeks posteriorly and the front of the
mesonotum clothed with silvery hairs. Mesonotum dull, strongly
and closely punctured except just on the disc, where, as well as on
the scutellum, it is shining and remotely punctured. Mesopleurae
529
Descriptions of Neio Sfihegidae.
densely clothed with silvery hairs ; wings with a yellow tinge ;
anterior tarsi with a comb of long black bristle-like hairs. Meta-
notum densely clothed with greyish hairs ; propodeum (dorsally)
finely reticulate ; its sides diagonally strigose and clothed with silvery
hairs. Abdomen densely and strongly punctured on the first four
segments, sparsely on the fifth ; apical dorsal valve narrowly trun¬
cate, shining, largely and remotely punctured ; the segments slightly
discoloured at their ajiices, and with a row of fine golden ciliae.
Beneath shining ; second segment closely punctured, the rest
remotely so.
Long. 10 mm.
$ 1. Biskra (11. vi, 98. F. D. M.).
[Type in Coll. F. D. Morice.]
This fine species is nearly double the size of any of the
other Palaearctic species. It somewhat resembles an
exceedingly large punctatissima, but the puncturation of
the mesonotum is finer, and only large and remote just on
the disc, whereas in punctatissima the large punctures
extend over nearly the whole surface. The silvery sides
of the thorax and propodeum and the grey-haired post-
scutellum are also characters of the new species, as well as
the wholly testaceous abdomen.
15. Gastrosericus moricei, n. sp.
(J. Caput atque thorax nigra pilis sericeis aureo-argenteis den-
sissime vestita. Antennarum scapus, tegulae, tuberculi [humerales],
abdomen et pedes testacei ; abdominis segmenta plus minus pilis
argenteis tecta, apicibus fere pellucidis et argenteo-fasciatis ; antennae
articulis 4-7 subtus sinuatis, articulo apicali subtus excavato
leniterque curvato.
. Black, with mandibles, scape of antennae, tubercles, tegulae,
and wing veins, legs and abdomen testaceous. Head and thorax,
including the propodeum, densely clothed with silvery hairs, which
in some lights have a more or less golden tint ; antennae with the
4th, 5th, 6th and 7th joints more or less sinuate, the apical joint flat¬
tened, somewhat excavated beneath and slightly curved. The silvery
clothing of the thorax is very dense, and is divided down the centre
by a sort of “parting.” Wings hyaline, anterior pair with the radial
cell very short and sharply truncate at the apex. Second submarginal
triangular, its base about as long as the radial. Front metatarsi prob¬
ably with a comb of fine transparent spine-like hairs, but of these
I can only see two on the only remaining front metatarsus in the
present specimen. Propodeum with its pubescence divided, down
530 Mr. E. Saunders on Hymenoptcra Aculenta :
the centre by a distinct parting ; its sculpture hidden, as is that
of the thorax ; abdomen pale testaceous. The apices of the segments
almost colourless and subdiaphanous. All the segments are more
or less clothed with silvery adpressed pubescence, and there is
a band of similar pubescence at the apex of each ; according to the
position the insect is held in, these silvery patches of pubescence
form different patterns ; beneath with the 2nd, 3rd and 4th segments
densely clothed with silvery hairs.
Long, [about 5 or 6 mm.].
[Biskra, 7. vi, 98. F. D. M. Type in Coll. F. D.
Morice.]
16. Dinetus simplicipes, n. sp.
^ . D. picto simillimus, minus flavopictus, differt ab eo metatarsis
anticis armatis 6 spinulis solum longitudine inaequalibus (harnm
quatuor plus minus dilatatis) femoribus anticis minus dilatatis
metatarsis intermediis gracilioribus et minus curvatis.
$ latet.
Closely allied in general aspect to pictus, F., but with the
abdomen (at any rate in the specimens before me) entirely rufo-
testaceous, showing in one specimen only any signs of flavous mark¬
ings, and having the apex concolorous (not black as h\ pictus). The
most important character, however, by which it may be known from
the above-mentioned species is the form and number of the spines on
the front metatarsi. These are inconspicuous, only six in number,
fine, irregular in size and length, and four only of them more or less
dilated, whereas in pictus they are seven in number, all thick and flat,
and form a very regular and conspicuous comb. The front femora
are less dilated, and the intermediate metatarsi are less curved and
more slender than in pictus. The propodeal area is much more finely
rugulose, and its sides more finely and densely strigose. The first
segment of the abdomen also, though finely rugulose, lacks the
distinct puncturation which exists on that segment in pictus.
Long. 6-6i mm.
Unfortunately Mr. Eaton met with males only of this
species.
[Five $ $ in Coll. Saiind. from Bone and Le Tarf (vi.
’96).]
17. Dinetus dentipes, n. sp.
8 9 . picto minor et gracilior, pilis argenteis micantibus ves-
titus ; trochanteribus anticis dente angulari instructis ; his in 9
subelongatis et paruni curvatis ; femoribus anticis in 9 subtus
ciliatis.
Descri'ptions of New Sphegidae. 531
^ . Smaller and paler than pictns, and more or less clothed with
fine silvery hairs. Face and antennae pale testaceous, clothed with
brilliant silvery pubescence ; vertex black, cheeks behind the eyes
testaceous, clothed with rather long silvery hairs. Prothorax pale
testaceous, clothed with silvery hairs, with a black band across the
central depression of its notum ; mesonotum black, clothed with
silvery hairs and punctured, its anterior angles pale ; scutellum, and
a spot on each side of it, polished, shining, and very largely and
remotely punctured ; mesopleurae black above, beneath (as well as
the sterna) shining and very remotely punctured. Wings hyaline,
their nervures pale testaceous, metanotum black, postscutellum pale
and shining ; propodeum finely rugulose, black, its “ area ” (the sides
of which are testaceous) finely rugulose ; the sides beyond the area
are clothed with brilliant silvery hairs. Legs pale testaceous with
the front trochanters produced into a large angular tooth. Front
metatarsi with four very fine, nearly white spines. Abdomen testa¬
ceous, very finely rugulose, first, second and third segment each with
a broad pale sub-apical band, the actual apex submerabranous and
colourless, clothed with silvery hairs. Beneath shining.
9. Very like the (J, but with the dark colour more extensive.
The front trochanters are not dentate, but unusually elongate and
slightly^ curved ; the front femora with a row of long white hairs on
their underside ; the metatarsi with a comb of seven fine whitish spines,
of these the two nearest the base are simple, the others more or less
flattened ; three spring from the apex of the joint. Abdomen with
the pale bands less in evidence than in the but the silvery apical
bands are more pronounced. Pygidial area shining.
Long. 5-5i- mm.
A very distinct species recognisable at once by the
silvery pubescence, and the peculiar structure of the front
trochanters.
[Types of ^ and $ are in Coll. Saunders, taken near Biskra
by Mr. Eaton in April.]
( 832 )
XX. Dcscl'iptions of two neiv species of Luciola in the
collection of Mr. H. E. Andrewes. By Ernest
Olivier, F.E.S., Member of the Entomological
Societies of France and Belgium, Correspondent of
the Museum of Natural History of Paris.
[Read December 7th, 1910. ]
I AM indebted to Mr. H. E. Andrewes for the privilege of
examining a fair collection of Lampyridcie from Ceylon
and the Nilghirri Hills. Among these, I have found two
interesting new species of which I give here the descrip¬
tions.
Luciola impressa, n. sp.
Oblonga, pubescens, capite et antennis nigris ; prothorace trans-
verso, antice rotundato, in medio antico angulose, basi recte truncato,
angulis sat prominulis, crebre punctato, flavo, duabus maculis
piceis discoidalibus ornato ; scutello triangulari, flavo, punctato ;
elytris oblongis, nigosis, bicostatis, pallide flavidis, macula parva
basili picea ; pectore flavo, tribus primis segmentis nigris, in medio
piceis; femoribus et tibiorum l)asi flavis, borum apice ac tar.sis nigris.
$ , tribus ultimis ventris segmentis cereis, lucidis, quinto utrinque
plaga cerea munito, ultimo supra incurvato, pygidii apice operto ;
9 , tribis ultimis ventris segmentis flavidis.
Long. 11-12 mill.
Ceylon ; Kandy, Peradeniya.
This species is allied to L. var'ia, Ern. Oliv., but it may
easily be distinguished by the black colour of the head,
the confused puncturing of the elytra, the structure of the
pygidium, and of the last ventral segments of the ^ ; the
fifth ventral segment has two lateral shells, the sixth is
produced into a narrow quadrangular lobe recurved upward
and hid by the apex of the pygidium recurved beneath.
It differs from L. humeralis, Walk., by the black head,
the pitchy spots of the prothorax, and by the apex of the
elytra being entirely yellow.
One 9 specimen has the prothorax entirely yellow.
TRANS. ENT. SOC. LOND. (1910). — PART IV. (DEC.)
Mr. E. Olivier’s Descriptions of New Species of Luciola. 533
Luciola auritula, n. sp.
Elongata, subparallela, pubescens, fusca ; capite flavo ; prothorace
transverso, antice vix anguloso, postice recte truncate, margine
lateraliter expanse et rotundato, tenue punctato, longitudinaliter
siilcatulo, flavo, angulis anticis fuscis ; scutello triangular!, flavo ;
elytris elongatis, parallelis, punctatis, obsolete costulatis, fuscis, basi
indeterminate flavidis ; pedibus infuscatis, coxis, femorum basi et
genubus flavidis. ventre nigro, ultimus segmentis cereis ; $ ignota.
Long. 8 mill.
Nilghirri Hills.
This is well distinguished by the lateral margins of the
prothorax depressed, expanded and rounded in front.
Head, scutellum, coxae, base of the thighs and knees
yellow. Prothorax yellow with the front angles fuscous.
Elytra fuscous with a yellow basal portion more or less
continued until the third of their length.
December 21, 1910.
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