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THE
TRANSACTIONS
OF
THE LINNEAN SOCIETY OF LONDON.
SECOND SERIES—VOLUME XVI.
ZOOLOGY.
THE PERCY SLADEN TRUST EXPEDITION
TO
THE INDIAN OCEAN IN 1905,
UNDER THE LEADERSHIP OF
Mr J. STANLEY GARDINER, M.A.
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LONDON :
PRINTED BY JOHN CLAY, M.A. AT THE CAMBRIDGE UNIVERSITY PRESS.
SOLD AT THE SOCIETY’S APARTMENTS, BURLINGTON HOUSE;
AND BY LONGMANS, GREEN, AND CO., PATERNOSTER-ROW.
1913—1914.
REPORTS
OF THE
PERCY SLADEN TRUST EXPEDITION
THE INDIAN OCEAN IN 1905,
UNDER THE LEADERSHIP OF
Mr J. STANLEY GARDINER, M.A.
VOLUME THE FIFTH
[BEING THE SIXTEENTH VOLUME OF THE SECOND SERIES, ZOOLOGY, OF THE
TRANSACTIONS OF THE LINNEAN SOCIETY OF LONDON.]
LONDON :
PRINTED BY JOHN CLAY M.A., AT THE CAMBRIDGE UNIVERSITY PRESS.
SOLD AT THE SOCIETY’S APARTMENTS, BURLINGTON HOUSE;
AND BY LONGMANS, GREEN, AND CO., PATERNOSTER-ROW.
1913—1914.
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VIII.
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CONTENTS.
PART J.—Jutny, 1913.
Report on the Calcareous Sponges collected by H.M.S. “ Sealark” in the
Indian Ocean. By Arruur Denpy, D.Sc. F.RS., FL.S., Professor of Zoology
im the Unwersity of London (King’s College). (Plates 1-5.) . pages 1-29
Second Report on the Arachnida—the Scorpions, Pedipalpr, and Supplementary
Notes on the Opiliones and Pseudoscorpions. By 8. Hirst. (Communicated
by Prof. J. Stantey GARDINER, M.A., F.RS., F.L.S.) (Text-figures 1-7.) 31-87
On a new Land-Leech from the Seychelles. By W. A. Harpine, W.A., F.L.S.
(Plate 6 and 1 Text-figure.) . : : aa Ie ; : : . 39-43
On Gypsina plana, Carter, and the relations of the Genus. By Marsgorte
Linpsegy, B.Sc., The University, Manchester. (Communicated by Prof. J. StANLEY
GarpinerR, M.A., F.R.S., F.L.S.) (Text-figures 1-6.) . ‘ : . 45-51
Pelagic Crustacea Decapoda of the Percy Sladen Expedition in H.MLS.
“ Sealark.” By Stantey Kemp, B.A., Assistant Superintendent, Indian Museum,
Calcutta. (Communicated by Prof. J. Stantey GARDINER, V.A., F.RS., FL.S.)
(Plate 7 and 1 Text-figure.) . , : ; ; : : . 53-68
Polychaeta of the Indian Ocean, together with some species from the Cape
Verde Islands. The Serpulide, with a Classification of the Genera Hydroides
and Hupomatus. By Heten L. M. Pixens, B.Sc., £.Z.S., Demonstrator of
Zoology and Reid Fellow, Bedford College, University of London. (Communi-
cated by Prof, J. Srantey Garpiner, WV.A., F.RS., F.LS.) (Plates 8 and 9.)
69-92
Coleoptera: Nitidulide, Heteroceride. Par A. GROUVELLE. (Commumaqué par
le Prof. J. SranueyY Garpiner, V_A., F.R.S., PLS.) (Text-fig. 1-6.) . 93-116
PART I1].—SeEprempBer, 1913.
Coleoptera, Pselaphide, de lArchipel des Seychelles. Par A. RaArrray.
(Communiqué par le Prof. J. Sranuey Garpiner, M.A. F.RS, F.L.S.)
(Planche 10.) : ‘ : 4 : : : . : : . 117-138
Rhynchota. Part I: Suborder Heteroptera. By Ww. Lucas Distant.
(Communicated by Prof. J. Stanney Garpiner, M.A., F.RS., £L.S.) (With
Plates 11-13:) : : : : ; : : : : . 139-191
XI.
XII.
UNL,
XIV.
XV.
XVI.
XVII.
XVIII.
XIX.
[wl
Coleoptera; Hydrophilide, Histeride. By Hueu Scott, M.A. (Cantab.),
PLS. FES., Curator in Entomology in the University of Cambridge.
(Plate 14.). : : : ; ; ; . : . pages 193-235
Coleoptera, Chrysomelide: Hispine of the Seychelles. By S. MauuiK, F.E.S.,
University of Cambridge. (Communicated by Prof. J. STANLEY GARDINER,
M.A., FRS., FILS.) (Text-figures 1-5.) ; : 4 : 5 QB~—WA®
Neuroptera, Myrmeleondae from the Indian Ocean. By James G. NEEDHAM.
(Communicated by Prof. J. Stantey Garpiner, V.A., F.RS., ELS.) (Text-
figures 1-3.) : : : : 4 : : : : . 243-246
PART II1L—January, .1914.
Coleoptera: Anthribide. By K. Jorpan, Ph.D. (Communicated by Prof.
J. Srantey Garpiner, W.A., FBS, FDS.) (Plate 15.) . . 247-267
Marine Alge, Rhodophycee. By Mrs A. Weper-van Bosse, Ph.D. (Com-
municated by Prof. J. Srannuey Garpiner, W.A4., . RS. FLAS.) (Plates
16-18 and 1 Text-figure.) . , ; ; , : ; : . 269-306
PART IV:——Juny, 1914.
Diptera: Heteronewride, Ortalide, Trypetide, Sepside, Micropende, Droso-
philide, Geomyzde, Milichide. By C. G. Lams, M.A., B.Sc., Clare College,
Cambridge. (Communicated by Prof. J. StantEy Garpiner, W.A., ERS,
FLAS.) (Plates 19-21 and 48 Text-figures.) . : d : . 807-372
Diptera: Scatopside, Simulude. Von Dr GuNTHER ENDERLEIN, Stettin.
(Mitgetheilt von Prof. J. Stantey GarpINER, M.A., F.R.S., FLAS.) (Mit 1
Text-figur. ) : é : 5 : ; : é : : . 373-375
Coleoptera, Buprestidae. Par Cu. KerremMans. (Communiqué par le
Prof. J. Srantey Garpiner, M.A., RS. FLAS.) (Avec 1 figure dans le
texte.) ; ; ‘ : é : : : : . 877-378
Coleoptera; Platypodide and Ipide from the Seychelles Islands. By Lt.-Col.
Win Sampson, F.E.S. (Communicated by Prof. J. Srantey Garpiner, M.A.,
ERS., FLAS.) (Text-figures 1-5.) . ; : one ‘ . 379-391
Coleoptera, Curculiomde. By Grorce Cuaries Caampion, A.L.S., F.Z.S.
(Plates 22-24.) . } : : : : : : : . 393-497
PART V. DerEcEmMpBgErR, 1914.
Titlepages, Contents, and Index : : 3 ; : : . 499-517
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No. L—REPORT ON THE CALCAREOUS SPONGES COLLECTED BY
H.M.S. “SHEALARK” IN THE INDIAN OCEAN.
By ArtHur Denpy, D.Sc., F.R.S., F.L.S., Professor of Zoology in the
University of London (King’s College).
(With Plates 1—5.)
Read 3rd April, 1913.
THE collection of calcareous sponges comprises only thirteen recognizable species, viz. :
PAGE
1. Leucosolena gardineri n. sp. 2
2. Dendya prolifera n. sp. . > ae ‘ : E 6
3. Leucascus simplex Dendy . : : 9
4. Leucetta chagosensis n. sp. . : : : : 10
5. Leucetta pyriformis n. sp. . 5 11
6. Perrcharax heteroraphis Poléjaett . ; : é 13
7. Pericharaw peziza n. sp. A : 15
8. Leucaltis clathria (Haeckel) . : : : 16
9. Grantessa hastifera (Row) . : ; 19)
10. Grantia indica n. sp. . : ; : : ; 20
11. Leucandra echinata Schutiner ; 23
12. Leucandra wasinensis (Jenkin) . ; 24
13. Leucilla proteus n. sp. . : 65)
Tt will be seen from this list that, though small, the collection contains a number of
very interesting forms. Thus the genus Dendya has hitherto been known only from a
single Australian species, D. tripodifera. Leucascus, another Australian genus, has only
been recorded once since first described by myself in 1892. The genus Pericharax has
not been met with since it was obtained by the “Challenger” and described by Poléjaeff
in 1883, and the “Sealark” Expedition has not only rediscovered the type species but
has also added a new one. Grantessa hastifera is a remarkable species only recently
described by Row from the Red Sea under the name Grantilla hastifera, and its re-
discovery by the ‘“‘Sealark” has given the opportunity for reconsidering its systeniatic
position. The most conspicuous and abundant species in the collection is Leucaltis
clathria (Haeckel), hitherto generally known as Heteropegma nodus-gordi Poléjaeft, which
appears to enjoy a remarkably wide distribution and has been described by various authors
under different names.
SECOND SERIES—ZOOLOGY, VOL. XVI. 1
bo
PERCY SLADEN TRUST EXPEDITION
As might be expected from the locality a large proportion of the species are
apparently new, though some of these are very closely related to already known forms.
Of the six old species, Lewcascus simplex has previously been obtained in Australian
and New Zealand waters; Pericharax heteroraphis at Tristan da Cunha in the South
Atlantic ; Leucaltis clathria from the Amirantes, Ceylon, E. Africa, Australia, Florida, the
Bermudas and Portugal; Lewcandra echinata from the Amirantes and Mauritius, and
probably Australia; Leucandra wasinensis from the East Coast of Africa, and Grantessa
hastifera from the Red Sea.
The classification adopted in this paper is that elaborated by myself and my colleague,
Mr R. W. H. Row, in a systematic and phylogenetic revision of the Calearea which we
hope to publish shortly. The appearance of the memoir in question will, I hope, render
it unnecessary for me to discuss questions of taxonomy and phylogeny on the present
occasion, and I content myself with giving diagnoses of the families and genera represented
in the collection.
Family Homoccelide Dendy and Row MS.
The whole of the gastral cavity and its various outgrowths lined by collared cells
throughout the life of the sponge. Sponge colony rarely radiate, and if so the
central individual retains the primitive ascon structure with a lining of collared cells and
without a special gastral cortex. No subgastral sagittal radiates. No true dermal
membrane or true dermal cortex.
Genus LeucosotentaA Bowerbank (emend.).
Diverticula of the gastral cavity, if any, never radially arranged around a central
tube. Skeleton composed of triradiate or quadriradiate spicules, to which oxea may be
added. No uteoid dermal skeleton ever present. Nuclei of collared cells basal or apical.
Section B.
Without oxea. Sponge colony typically forming a reticulation of simple ascon-
tubes.
1. Leucosolena gardineri un. sp.
(Plate 1, figs. 1, 2; Plate 3, figs. 1, 2, 3.)
This species is represented in the collection by two specimens, which, although
coming from the same locality, differ considerably from one another in appearance. This
difference, however, may be partly (though not entirely) accounted for by a difference in
the condition of the specimens*, the one (cxx. 7; Plate 1, fig. 1) being expanded and
having more or less conspicuous and widely open vents, while in the other (cxx. 11;
Plate 1, fig. 2) the vents are difficult to recognize and almost if not completely closed.
Moreover the latter specimen is full of large maturing ova, and the mesoglcea is densely
* Cf. Minchin [1892].
DENDY—REPORT ON THE CALCAREOUS SPONGES 3
charged with peculiar yellow bodies, which doubtless accounts for the fact that it is much
more opaque in appearance than the former.
Both specimens are of the typical ‘“Clathrina” form, the ascon-tubes being very
* slender and forming a very close-meshed reticulation, especially on the surface, which
appears closely and minutely punctate from the presence of the very numerous pseudo-
pores. It will be desirable to describe the two specimens separately, and they might even
be considered as representing distinct varieties.
cxx. 7. The reticulation of the ascon-tubes (Plate 1, fig. 1) is so close that the
pseudopores are scarcely visible to the naked eye, and the whole colony appears at first
sight to be enveloped in a continuous dermal membrane (pseudoderm). The entire sponge
is about 25 mm. in greatest diameter by 15 mm. in height. It may be described as
proliferously lobose ; the lobes lamellar, ascending and folded, varying greatly in size and
shape, and occasionally bearing an open vent at the summit. The colour in alcohol is
white and the texture soft and delicate.
The canal system is of the reticulate type E*, without pseudogasters or pseudoscula,
but the ascon-tubes in the middle of the lamellze form very wide gastral cavities (lined by
collared cells) which open to the exterior through the vents. These central cavities are
surrounded by a rather thin wall composed of the reticulation of smaller tubes, which open
into them.
The specimen is not in a good condition for histological examination, but there
appears to have been a rather copious endogastric network of slender connective tissue
threads, as in Leucosolenia proaima Dendy. The collared cells are in a somewhat
elongated condition, with basal nuclei. Their transverse diameter at the base is about
0-004 mm. and they line the gastral surfaces in a perfectly normal manner.
The skeleton exhibits the normal arrangement and the spicules may be classified as
follows :—
(1) Hquiangular triradiates; with straight, fairly sharply pointed rays (Plate 3,
fig. 1,a). These are of two principal sizes, (a) with rays measuring about 0°14 mm. in
length by 0°012 mm. in thickness at the base, characteristic of the pseudoderm, (8) with
rays about 0°074 by 0:007 mm., characteristic of the deeper parts of the sponge.
(2) Quadriradiates (Plate 3, fig. 1, b), resembling the deep triradiates but with a
very sharply pointed, perfectly straight apical ray, projecting into the gastral cavity at
right angles to the three facial rays; length of apical ray about 0°05 mm., but variable.
cexx. 11. The entire colony, a portion of which is represented in Plate 1, fig. 2,
forms a dorsi-ventrally flattened crust, composed of low ridges and lobes and pierced by
a few large holes which give it a coarsely clathrous character. The reticulation of the
ascon-tubes on the surface is not quite so fine-meshed as in cxx. 7, so that the innumerable
pseudopores are easily visible to the naked eye. The vents are minute and difficult to
recognize, situated on low rounded eminences. ‘The entire sponge is some 40 mm. in
maximum diameter, but not more than 3 or 4 mm. in height (thickness). The colour in
spirit is light brown, apparently owing to the numerous yellow bodies which it contains.
* Cf. Dendy [1891].
1—2
4 PERCY SLADEN TRUST EXPEDITION
The canal system, except for the obliteration of the vents, due probably to temporary
contraction, agrees closely with that of cxx. 7, and here again there appear to be remnants
of an endogastric network.
The spiculation (Plate 3, fig. 2) differs from that of cxx. 7 only in the absence of
any distinction between deep and dermal triradiates, all bemg of about the same size as
the deep triradiates in the specimen above described.
The histology exhibits great peculiarities, which I am inclined to associate with
the contracted condition of the sponge and with the presence of the numerous large
ova. The mesoglea is much thickened, especially in the neighbourhood of the ova,
which lie separately in spherical cavities not lined by any special epithelium™. Frequently
the presence of an ovum (Plate 3, fig. 3, ov.) causes a conspicuous inward bulging of the
wall of the ascon-tube in which it lies. Most remarkable is the presence in the mesogloea
of enormous numbers of small spherical bodies of a yellowish colour and varying from
about 0°004 to 0°006 mm. in diameter. These bodies are abundant throughout the
mesogloea but especially so immediately beneath the gastral surfaces of the tubes and
around the large ova (Plate 3, fig. 8, yb). They lie in the meshes of a network formed
of delicate strands of mesogloea with here and there small nucleated amcebocytes or
stellate connective tissue cells (collencytes). They occasionally appear to adhere to the
surface of an ovum or even to become embedded in the peripheral cytoplasm of the latter.
Sometimes very similar bodies are actually found imbedded in the nucleus of the ovum,
but I do not think they can really be the same. These yellow bodies are so remarkable
that I append a detailed account and discussion of the subject to the description of this
species.
There are apparently no collared cells at all, the inner surfaces of the ascon-tubes
(easily recognizable both by their position and by the projecting apical rays of the
quadriradiates) being lined, at any rate to a large extent, by a flattened pavement
epithelium, similar to and quite as well developed as that which lines the outer surfaces.
This absence of collared cells from the gastral surfaces is highly remarkable, but is to
some extent in harmony with the observations of Minchin [1900] on contracted specimens
of Clathrina, and with those of Weltner [1907] on the disappearance of the collared
cells in Spongilla (Hphydatia) during the winter months. Whether or not, in the
present case, the collared cells are represented by the yellow bodies in the mesogloea will
be discussed presently.
The ova are very large, attaining a diameter of about 0°086 mm., although they have
evidently shrunk to some extent, so that they no longer fill the cavities in which they lie.
They nearly all seem to be in about the same stage of growth (Plate 3, fig. 3). The
cytoplasm is coarsely granular, or flocculent-looking ; there is no vitelline membrane and
no sharp outline to the cytoplasm. The nucleus is large and centrally placed, it usually
has an irregular outline and no nuclear membrane. It is differentiated from the cyto-
plasm by its denser and much less coarsely granular appearance and frequently contains
‘a small number (? from 1 to 3) of conspicuous spherical bodies (nucleoli ?), closely
* Such an epithelium when present is probably derived from amceboid wandering cells becoming flattened.
A few such may sometimes be detected around the ovum.
DENDY—REPORT ON THE CALCAREOUS SPONGES i)
resembling in appearance the yellow bodies of the mesogloea (but more variable in size).
In preparations stained first with borax carmine and subsequently with picro-indigo-
carmine the cytoplasm of the ovum stains indigo-green, the nucleus much the same and
the nucleolus (?) brighter green ; whereas, in the same sections, the nuclei of the epithelial
cells and those of the amocebocytes (? collencytes) are all stained bright red. As we shall
see presently the yellow bodies in the mesoglcea also become stained green by this
method.
Leucosolenia gardinert is evidently nearly related to L. depressa, Dendy [1891],
from Port Phillip, Australia, but differs in the proportions of the spicules. Both are
also evidently closely related to the common European species Leucosolenia (Clathrina)
coriacea, of which I have already [Dendy, 1905] described a variety from Ceylon
(Leucosolenia coriacea, var. ceylonensis). Possibly all should be regarded merely as
varieties of the same species.
Register Nos., Localities, de, cxx. 7, cxx. 11, both from Salomon (Chagos
Archipelago), 10—14 fathoms.
The Yellow Bodies of Leucosolenia gardinert (cxx. 11).
These bodies (Plate 8, fig. 3, yb) were subjected to very careful examination™ with
a view to determining their real nature. They appear to be insoluble in water, alcohol,
xylol, ether and hydrochloric acid (5°/,), and even boiling with dilute hydrochloric acid
for several minutes appears to have little or no effect upon them. Cold caustic potash
(5 °/,) has little effect upon them, except that they became very transparent and perhaps
somewhat swollen. When boiled with caustic potash of the same strength, however, they
slowly disappear.
Iodine, dissolved in potassium iodide, causes them to assume a deep orange colour.
On adding strong sulphuric acid the colour deepens to brown and then black, and the
bodies break up into minute black particles without showimg any blue coloration.
Schultze’s solution also gives no blue colour.
From these observations I conclude that the bodies are certainly organic but that
they contain neither starch nor cellulose and are not of vegetable origin.
Their staining reactions are very characteristic. They stain well with eosin but
rather faintly with hematoxylin (Ehrlich’s logwood) and borax carmine. With borax
carmine followed by picro-indigo-carmine they stain bright green, while the ordinary
tissue nuclei of course stain red.
Sometimes they stain uniformly and appear almost homogeneous in structure, but
more often they exhibit a distinctly vesicular character, showing a thin wall with
granular or reticulate contents apparently collected on its inner surface, as if coagulated
or precipitated there. Thus they often appear like spherical nuclei with a deep-staining
chromatin network and a very distinct nuclear membrane, but no surrounding cytoplasm.
Their staining reactions, however, are not those of typical nuclei.
The most probable explanation of these yellow bodies seems to be that they are
* All these observations were made upon spirit-preserved material.
6 j PERCY SLADEN TRUST EXPEDITION
metamorphosed collared cells. It is true that they are considerably larger than the latter
and differ much in structure, but their arrangement in the mesogloea and the fact that
there are no other collared cells in the sponge are very suggestive. It is possible that
they may be concerned with the nutrition of the large ova, whose staining reactions are
so curiously similar. I would suggest that the sponge having become full-fed, and the
reserve material being stored up in the collared cells, the latter migrate inwards and are
ingested by the probably amceboid ova. During this process the sponge becomes con-
tracted and its ordinary activities suspended. The collared cells are probably replaced
again later on by metamorphosis of amcebocytes, which at first line the gastral cavities as
a flattened epithelium.
The idea that the yellow bodies may be symbiotic alge also naturally suggests
itself, but their chemical reactions and the fact that they contain no nuclei appears to
me to be fatal to this view. I have also compared them carefully with the ‘‘ yellow
granules” of Leucosolenia (Clathrina) coriacea, but the latter are very much smaller
and obviously quite different.
Genus Drenpya Bidder.
Sponge colony consisting of a large central individual lined by collared cells, from
which radially arranged diverticula are given off. Skeleton composed of equiangular
triradiates, to which quadriradiates may be added. Nuclei of the collared cells probably
always basal.
This genus was proposed by Bidder [1898] for Carter’s Clathrina tripodifera, of
which I gave a full account in my Monograph of Victorian Sponges, Part I [1891].
Hitherto this has remained the only known species, but I have now to describe a new
and very interesting form.
2. Dendya prolifera n. sp.
(Plate 1, figs. 3, 4; Plate 3, figs. 4, 5.)
The single specimen of this very interesting species is unfortunately in a very poor
state of preservation, especially as regards histological characters. It has now the form
of a flattened sac (Plate 1, figs. 3, 4), but how far the flattening may be due to artificial
compression I am not prepared to say. The sac is much broader at one end than at the
other and the vent is situated at the narrow extremity. It is also slightly curved, with
one margin convex and the other concave. The broad end has been artificially truncated,
so that the central gastral cavity opens widely to the exterior at this end as well as at
the vent. A good deal of the wall of the sac has also been torn away from one side.
There is no indication of dorsiventral differentiation of the two flattened sides and the
sponge probably grew either erect or pendent.
The thickness of the wall of the sac is very irregular and the surface very uneven,
owing to the development of the very numerous proliferating radial tubes. These tubes
-are for the most part arranged in dense bunches, between which the thin, translucent
gastral membrane is in many places plainly visible from the outside, especially in the
neighbourhood of the vent, where it forms a thin oscular membrane or collar.
DENDY—REPORT ON THE CALCAREOUS SPONGES 7
The length (or height) of the specimen is 31 mm. and the maximum breadth about
14mm. The thickness of the wall of the sac near the broad end (made up chiefly of the
radial tubes) is usually about 2 mm., but very variable. (
The texture of the whole sponge is delicate and fragile, and the colour in alcohol pure
white.
Owing to the bad state of preservation it is impossible to determine the distribution
of the collared cells, but it is probable that they occur on the inner surface of the gastral
membrane as well as in the radial tubes. The canal system makes a close approach to
that of Dendya tripodifera, differing chiefly in the fact that the radial tubes are much
less regularly arranged, while their blind outer ends, lying at very various levels, do not
form an approximately even surface to the sponge, as they do in the Australian species.
The radial tubes appear first as minute hollow buds, growing outwards from the thin
wall of the central gastral cavity (gastral membrane). Such buds occur not only in the
neighbourhood of the osculum but also on the lower parts of the sponge wall between the
bunches of more fully developed radial tubes. They may be quite solitary but are usually
linked together in irregular networks. I have not been able to determine how this
“linked” arrangement of the young radial chambers originates. Their appearance, when
the sponge-wall is viewed from the outside, suggests that they arise from one another by
budding, like yeast-cells. Probably some grow out directly from the wall of the central
gastral cavity and then give rise to others by budding. When fully grown the tubes are
arranged in bushily branched bunches (Plate 3, fig. 4), the smaller ones coming off from
the larger ones and the largest only (i.e. the main stems of the bunches) opening direct
into the central gastral cavity. These main stems usually measure about 0°3 mm. in
diameter. heir branches become much narrower and closely crowded together, their walls
actually fusing where they come in contact with one another, but without lateral com-
munication of their gastral cavities. Different bunches of tubes, on the other hand,
remain to a large extent separate from one another. The openings of the main tubes of
the bunches are very conspicuous on the inner surface of the gastral membrane, usually
arranged in small groups and varying very much in size (Plate 1, fig. 4).
Sometimes a group of chambers forms an irregular, rounded nodule, attached by a
narrow stalk to the gastral membrane, and perfectly independent of its neighbours on all
sides. One such nodule measures as much as 4 mm. in diameter, and the opening of its
hollow stem into the gastral cavity, oval in outline, measures no less than 1°5 mm. in
longer diameter. Usually, however, the openings in the wall of the gastral cavity are
much smaller than this. .
The prosopyles cannot be distinguished. They probably occur both in the gastral
membrane and in the walls of the radial tubes.
The skeleton consists of triradiates and quadriradiates ; I have found no oxea. The
spicules all lie tangentially in the thin walls of the radial tubes or in the gastral membrane,
and when an apical ray is developed it projects as usual into the gastral cavity or one of
its outgrowths. In the gastral membrane (wall of the central cavity), as in the walls
of the radial tubes, the spicules may be said to form a single layer, but their rays overlap
one another copiously. Their arrangement is nowhere very regular, but in the gastral
‘LIBRARY |S
8 ' PERCY SLADEN TRUST EXPEDITION
membrane there is a more or less strongly marked tendency to orientation with one ray
pointing away from the osculum, and in the radial tubes there is a distinct orientation
with one ray pointing towards the blind end of the tube. In fact the radial tubes may
almost be said to have an articulate tubar skeleton, albeit of an ill-defined character, the
“joints” being very confused.
The spicules may be classified as follows :—
(1) Triradiates of the gastral membrane (Plate 3, fig. 5, w):—These vary in form
from regular, equiradiate and equiangular to markedly sagittal. The angles, however,
appear always to be equal. The rays are usually long and slender, usually straight but
sometimes a little crooked, varying greatly in length, up to about 0°27 mm. with a
diameter of about 0°01 mm. at the base. They are sharply and gradually pointed and
sometimes the diameter increases slightly before tapering off to the apex.
(2) Triradiates of the radial tubes (Plate 3, fig. 5, ¢):—These are usually more or
less markedly sagittal, owing to the greater length of the distal (basal) ray, but they
remain equiangular. The paired rays, as usual, follow the curvature of the wall of the
radial tube. At the blind distal ends of the radial tubes, where the spicules become much
smaller, the sagittal character tends to disappear. The spicules in the walls of the radial
tubes perhaps never become as large as the largest of those in the gastral membrane.
In a triradiate from about the middle of a tube the distal ray measured about 0:12 mm.
in length, and each of the oral rays about 0°085 mm., all with a diameter of about
00075 mm. at the base. A triradiate from the blind end of a tube was almost equiradiate,
with rays about 0°07 mm. in length.
(3) Quadriradiates:—These differ from the triradiates only in the development
of a slender and frequently crooked apical ray (Plate 3, fig. 5, e). They occur both in
the gastral membrane (Plate 8, fig. 5, b) and in the walls of the radial tubes (Plate 3,
fig. 5, d, e), but are much more conspicuous in the latter, where their apical rays measure
about 0°09 mm. in length.
There can, I think, be no question of the close relationship of this species to the
Australian Dendya tripodifera. It ditters chiefly in the much less regular arrangement
of the radial tubes and in the absence of the special “tripod” spicules from the distal ends
of these tubes.
Register No., Locality, dc. cvt. 2, Amirante Isles (Seychelles), 13.10.05, E. 16,
39 fathoms.
Family Leucascide Dendy (emend.).
Sponge typically forming a massive colony, usually with several or many oscula, but
may be integrated into a single individual with definite external form. With no large
central gastral cavity lined by collared cells, but with an exhalant canal system devoid
of collared cells. Flagellated chambers ranging from long, and possibly branched, with
a tendency to radial arrangement around the larger exhalant canals, to small, approxi-
mately spherical and irregularly scattered. With a distinct and independent dermal
membrane (or cortex), pierced by true dermal pores. Skeleton consisting mainly of
_DENDY—REPORT ON THE CALCAREOUS SPONGES 9
equiangular and equiradiate spicules, which may become sagittal at the oscular margins.
Radiates of the chamber layer with no definite arrangement, but irregularly scattered
in the walls of the large elongated chambers, or between the small spherical chambers.
No subgastral sagittal radiates. Nuclei of collared cells probably always basal.
Genus Leucascus Dendy.
Flagellated chambers greatly elongated, tubular, and sometimes copiously branched.
Section A.
No oxea present.
3. Leucascus simplex Dendy [1892].
(Plate 1, fig. 5; Plate 4, fig. 1.)
This species is represented in the collection by a considerable number of examples
encrusting the branches of several specimens of a small gorgonid.. The sponge forms
uregular, more or less lobulated, cushiony masses (Plate 1, fig. 5), with small vents
occurring singly here and there in prominent positions. Examination with a pocket lens
shows the thin, pore-bearing dermal membrane, the groups of inhalant pores being
especially conspicuous over the ends of the radially disposed inhalant channels. The
colour in spirit is light brownish yellow and the texture soft and compressible and
rather fragile.
The canal system agrees precisely with that which I described in the case of the
Australian specimens, and of which I gave an illustration in my “Studies on the
Comparative Anatomy of Sponges,’ Part V [1893, fig. 1]. The flagellated chambers
are very long, copiously branched, and with their blind, outer ends directed more or
less at right angles towards the dermal surface, where they are covered over by a thin
cortex formed by the spiculiferous, pore-bearing dermal membrane. Internally the main
stems of the branching systems of chambers open into wide exhalant canals which
converge towards the vents. The thin walls of these exhalant canals are not lined
by collared cells, but they are strengthened by numerous spicules.
The blind ends of the branches of the radial chambers form a sort of reticulate
pattern beneath the thin dermal cortex, in the meshes of which lie the outer ends of the
large inhalant channels, which run in between the groups of chamber-branches at right
angles to the surface.
The skeleton consists of triradiate and quadriradiate spicules of fairly uniform size,
irregularly scattered in the walls of the radial chambers and of the exhalant canals, and
likewise in the dermal cortex, but nowhere forming more than a thin layer, though of
course with overlapping rays.
The spicules (Plate 4, fig. 1) are all regular, with moderately stout, gradually sharp-
pointed rays, measuring about 0°1 mm. in length by 0:01 mm. in diameter at the base.
In the walls of the exhalant canals many of them have well-developed but slender,
straight or nearly straight, very sharp-pointed apical rays, projecting into the lumen
SECOND SERIES—ZOOLOGY, VOL. XVI. 2
10 PERCY SLADEN TRUST EXPEDITION
of the canal and attaining a length nearly, if not quite, equal to that of the facial rays.
The stronger development of these apical rays constitutes the only difference (and this
is probably not constant) which I can detect between the Providence specimens of this
species and those from Australia. As the spicules have not yet been properly figured
I take the opportunity of doing so on this occasion.
Previously known Distribution. Near Port Phillip Heads, Bass Straits; and
Watson’s Bay, Port Jackson [Dendy 1892]; New Zealand [Kirk 1897].
Register Nos., Locality, dc. X1x. 4, xxm. 2, a number of specimens, all from
Providence, 4.10.05, D. 4, 50—78 fathoms.
Genus Leucetta Haeckel (emend.).
Canal system leuconoid, with small, spherical or subspherical flagellated chambers
irregularly scattered through the chamber layer.
Section A.
No oxea present.
4. Leucetta chagosensis n. sp.
(Plate 1, fig. 6; Plate 4, fig. 2.)
There are five specimens of this sponge in the collection, but two of them (cx. 10)
are perhaps parts of the same.
The external form (Plate 1, fig. 6) is massive or encrusting, irregular, lobose, with
rather large, scattered vents with slightly prominent margins. The thin dermal membrane
covers over irregular, meandering subdermal cavities.
All the specimens are of much the same size; CxIx. 11, which may be regarded
as the type, measures about 27 mm. in length by 18 mm. in breadth and 12 mm. in
greatest thickness.
Texture fairly compact, firm and resilient, but friable.
Colour in alcohol ranging from white to rather dark brown; in the latter case
probably stained by other specimens.
The canal system is typically leuconoid. The flagellated chambers are oval and
measure about 0156 by 0°104 mm. in diameter. They are eurypylous and each one
probably has several prosopyles. The thin, pore-bearmg dermal membrane overlies a
system of subdermal cavities, which liein the rather thick gelatinous ectosome, but are not
to be sharply distinguished from the rest of the inhalant canal system, consisting of wide,
irregular canals or lacune. Similar wide exhalant canals or lacunz converge into the
still wider oscular tubes, which are surrounded by a moderately thick layer of gelatinous
mesoglcea resembling the ectosome. The oscula are variable in size, commonly from 1 to
2 mm. in diameter.
The main skeleton is confused, and consists of triradiate spicules densely and
irregularly scattered throughout ectosome and choanosome alike. In the dermal
DENDY—REPORT ON THE CALCAREOUS SPONGES 11
membrane the spicules lie tangentially, and a few very large triradiates are scattered here
and there amongst the small ones.
The spicules (Plate 4, fig. 2) are equiangular and approximately equiradiate tri-
radiates, of three kinds:—(1) Small, with straight, conical, gradually sharp-pointed
rays measuring about 0°19 by 0°0174 mm. (at base). These form the whole of the
skeleton of the choanosome and the greater part of the dermal skeleton. (2) Large,
with straight, conical, gradually sharp-pointed rays measuring about 0°59 by 0°046 mm.
(at base). These occur sparsely scattered in the dermal membrane only. (3) Small,
resembling (1) but rather smaller, and with two of the rays bent sharply back near their
bases until they come to extend nearly at right angles to the third ray. These occur
in the oscular margins, along or parallel to which their bent rays are extended.
As regards histology the condition of the specimens allows me to say very little, but
it is worth observing that the nuclei of the collared cells are distinctly basal.
This species is evidently nearly related to Leucetta microraphis Haeckel, but my
specimens of that species (from Port Phillip, Australia) have the large triradiates much
larger (and especially stouter) and scattered all through the sponge instead of being
confined to the dermal surface.
The specimens of L. chagosensis all came from the Chagos archipelago.
Register Nos., Localities, dc. uxtit., Lagoon, Diego-Garcia, 8.7.05 ; cx. 10, Egmont
Reef; oxi. 5, Egmont Lagoon; cx1x. 11, Salomon.
5. Leucetta pyriformis n. sp.
(Plate 1, fig. 7; Plate 4, fig. 3.)
This pretty little species was represented in the collection by two specimens which
closely resembled one another in external form, each being a single, small, pear-shaped
individual with a single terminal vent at the broad end.
Specimen xc. 5, A, which may be taken as the type, is represented in Plate 1, fig. 7.
It measured about 10 mm. in height by 5°5 mm. in greatest breadth. The texture was
harsh owing to the presence of the very large triradiate spicules, which could be clearly
seen on the surface with a lens, as shown in the photograph. The colour in spirit was
hight brown, but this may have been due to accidental staining by other specimens in the
same jar.
The canal system is typically leuconoid.. Wide inhalant canals commence beneath
the thin, pore-bearing dermal membrane and run inwards at right angles to the surface,
interdigitating with exhalant canals which open into the central gastral cavity. Both
Systems are of course more or less branched, and between them lie the rather large
flagellate chambers, closely packed together and without order. The chambers are
oval or nearly spherical, often polygonal from mutual pressure ; they are eurypylous and
have numerous prosopyles; they measure about 0-174 mm. in maximum diameter.
The dermal skeleton consists of several confused layers of large and small triradiates
placed tangentially. The skeleton of the chamber layer also consists of large and small
triradiates, thickly and irregularly scattered. There is a well-developed, though not very
9)
Aa—a4
12 PERCY SLADEN TRUST EXPEDITION
thick, gastral cortical skeleton, continued for some distance into the main exhalant canals
and consisting of small triradiates and quadriradiates with long, slender apical rays.
There are no subgastral nor subdermal sagittal or pseudosagittal radiates, but small
sagittal radiates* occur as usual in the thin and rather narrow oscular collar.
The spicules are all equiangular and approximately equiradiate, and there are two
chief forms :—
(1) Very large triradiates (Plate 4, fig. 3, a), with stout rays tapering gradually
to fairly sharp points and measuring about 1:0 by 01 mm.
(2) Small triradiates (Plate 4, fig. 3, b), with rather slender rays measuring about
0°17 by 0:0125 mm.
Both forms are abundant and they occur intermingled at the surface and in the
chamber layer, but the small ones are much more numerous than the large ones, while
intermediate sizes (Plate 4, fig. 3, c) occur in much smaller numbers.
The quadriradiates of the gastral cortex differ only from the small triradiates in the
development of the long, slender apical ray, while the sagittal triradiates of the oscular
collar are but shght modifications of the ordinary small form, with the two oral arms bent
back till they he approximately at right angles to the basal arm.
The specimens are not well preserved from the histological point of view, but the
nuclei of the collared cells are basal.
This species differs from Leucetta chagosensis in its well-defined external form, in the
presence of the large triradiates in the chamber layer, as well as in the dermal membrane,
in the spicular measurements and in the presence of the gastral quadriradiates. It comes
closer to Leucetta microraphis, as represented by my Australian specimens, but again
differs in the well-defined external form, consisting of only a single person of very
definite shape.
It may quite possibly be identical with one of the numerous forms included by
Haeckel [1872] under his Leucetta primigena, but I do not think it possible to dis-
entangle all these forms from one another.
Register Nos., Localities, dc. xc. 5, A and B, Cargados Carajos, 1.9.05, B. 29,
45 fathoms.
Genus PERICHARAX Poléjaeff (emend.).
Colony individualised, with large central cavity opening by a wide osculum and
surrounded by a very thick wall. Canal system leuconoid, with subspherical, scattered
chambers, and with subdermal cavities whose walls are supported by a special skeleton
derived partly from inturned rays of tangential dermal triradiates. Skeleton of chamber
layer confused, composed of equiangular triradiates of two very different sizes.
Poléjaeff [1883] proposed the genus Pericharax for the reception of two specimens
obtained by the “Challenger” at Tristan da Cunha, which he regarded as representing
_ two varieties of one and the same species, Pericharaz carteri. The one variety he named
* These have been dissolved out, but their spicule sheaths are still clearly recognizable and give the form
quite distinctly.
DENDY—REPORT ON THE CALCAREOUS SPONGES 13
homoraphis and the other heteroraphis. My examination of the type specimens in the
British Museum collection has led me to the conclusion that not only are these two
so-called varieties specifically distinct, but that var. heteroraphis alone belongs to the
genus Pericharax, the other being probably a Leucetta, without the characteristic dermal
spicules of Pericharac.
The genus has not been recorded since it was first described, but the rediscovery
by the “Sealark” of P. heteroraphis, and of a new species closely related to it, in the
Indian Ocean, and the careful study of these forms, has led me to accept the genus
as a valid one and to characterize it as above. The position assigned to the genus, in the
family Leucascidee, is justified not only by the character of the skeleton, composed mainly
of equiangular and irregularly scattered radiates, but also, as in the case of Leucetta, by
the basal position of the nuclei of the collared cells.
6. Pericharax heteroraphis Poléjaett.
(Plate 1, fig. 8; Plate 5, figs. 1, 2.)
Pericharax cartert var. heteroraphis Poléjaeff [1883 |.
The single specimen in the collection (Plate 1, fig. 8) agrees very well with the
description and figure of the external form of P. carteri given by Poléjaeff. It is almost
globular and about 20 mm. in diameter. There is a single vent about 4 mm. in diameter,
with a slightly prominent oscular margin; and the sponge has apparently been attached
by the lower pole, opposite to the vent. The surface is smooth, and covered by a thin,
pore-bearing dermal membrane which, under a pocket lens, exhibits a beautifully
reticulate appearance. The texture is firm and compact, and the colour in spirit pale
greyish yellow.
The large central cavity, which opens to the exterior through the osculum, is
surrounded by a wall about 4 mm. thick, but gradually diminishing in thickness towards
the oscular margin. The inner surface of this wall is pierced by numerous exhalant
apertures, for the most part grouped in depressions.
There is a well-defined, gelatinous ectosome, about 0°26 mm. thick. This layer
contains no flagellate chambers, but is broken up into trabecul, running vertically to
the surface, by the numerous small subdermal cavities, which are roofed over by the
thin, pore-bearing dermal membrane. The subdermal cavities open collectively into large,
irregular inhalant lacunz, which ramify inwards. Similar exhalant lacune, ramifying
in the opposite sense, open into the large central cavity. In the mesoglea between the
exhalant and inhalant lacunze lie the spherical or subspherical flagellate chambers, thickly
scattered, and each about 0°12 mm. in diameter.
The main skeleton consists of triradiate spicules, of two chief and very different
sizes, scattered thickly and without order throughout the chamber layer of the sponge,
the small spicules occupying the interstices between the large ones.
There is a well-developed dermal skeleton, composed of a dense reticulation of small,
tangentially placed triradiates, the rays of which frequently lie parallel and in juxta-
position with one another to form multispicular meshes, continued below into the
wo & v 2
PyOlCA7
Sv0P§ #5
i oO a 9
Bs = ®e@=> Cae
14 PERCY SLADEN TRUST EXPEDITION
supporting skeleton of the subdermal cavities. The latter is formed partly by small
triradiates lying parallel to but beneath the surface, partly by inturned rays of some
of the dermal triradiates, and partly by small triradiates which lie in planes more or
less at right angles to the surface, with one ray pointing outwards and lying parallel
to the inwardly-directed rays of the dermal triradiates in the trabecule between the
subdermal cavities.
The figure given by Poléjaeff of the skeleton of the subdermal cavities in
Pericharax carteri, var. heteroraphis, appears to be highly diagrammatic. He does not
seem to have recognized the nature of the inwardly turned rays of the dermal triradiates,
and figures the deep triradiates of the ectosome as markedly sagittal, and the whole
as far too regular in arrangement. I believe, however, that the deeper spicules of the
ectosome are sometimes sagittal, like many of those in the dermal membrane.
There is a thin gastral cortical skeleton, composed of quadriradiate spicules, which
lines not only the central gastral cavity but also the larger exhalant canals, the surface
of both of which bristles with the projecting apical rays. The spicules may be classified
as follows :—
(1) Large triradiates of the main skeleton (Plate 5, fig. 1, a); equiangular and
typically equiradiate; rays long, straight and gradually sharp-pointed, measuring, say,
about 1°55 mm. long by 0°1 mm. in diameter at the base.
(2) Small triradiates of the main skeleton (Plate 5, fig. 1, b; fig. 2, a); equiangular
and typically equiradiate ; rays straight, slender and gradually sharp-pointed, measuring,
say, about 0°18 by 0°015 mm. Intermediate sizes (Plate 5, fig. 1, c) between (1) and
(2) occur in comparatively small numbers.
(3) Triradiates of the dermal skeleton (Plate 5, fig. 2, b); irregular, but probably
really equiangular; with slender, irregularly curved rays, often’ unequal in length and
often blunt at the apex. One of the rays (z) is frequently bent inwards out of the tangential
plane, so as to take part in the formation of the skeleton of the subdermal cavities. Rays
measuring, say, about 0°13 by 0°008 mm.
(4) Quadriradiates of the gastral cortex (Plate 5, fig. 2, c); equiangular and
typically equiradiate, with long, straight, or nearly straight, slender facial rays, fairly
sharply pointed and measuring about 0°18 by 0°0125 mm.; with a sharp-pointed apical
ray, irregularly hooked* at the extremity in a characteristic fashion and about
0-1 mm. long.
(5) Quadriradiates of the larger exhalant canals (Plate 5, fig. 2, d); similar to
those of the gastral cortex, but with long, straight apical rays, measured up to 0°26 mm. in
length ; as usual, intermediate forms occur.
(6) Quadriradiates of the oscular collar (Plate 5, fig. 2, e); resembling those of the
gastral cortex further in, but conspicuously sagittal owing to the sharp backward bending
of the oral rays until they come to lie parallel to the oscular margin. These spicules are
* This hooking does not occur in the “Challenger” specimen, but I cannot regard it as constituting
a character of specific value.
DENDY—REPORT ON THE CALCAREOUS SPONGES 15
abundant in the narrow membranous collar which surrounds the vent, and by no means
confined to the actual margin.
The only histological feature that requires notice is the basal position of the nuclei
of the collared cells in the spirit-preserved material.
Previously-known Distribution. Tristan da Cunha (Poléjaeff).
Register No., Localities, dc. cxvut., Salomon (Chagos Archipelago), 15 fathoms,
10.6.05.
7. Pericharax peziza* n. sp.
(Plate 1, fig. 9; Plate 5, figs. 3, 4.)
The single specimen (Plate 1, fig. 9) has the form of a shallow, thick-walled cup, the
mouth of which is oval in outline. It has apparently been attached below to some
foreign object, but there is no indication that it ever possessed a stalk. The mouth of
the cup is obviously a greatly dilated osculum and is surrounded by a narrow, thin
oscular margin, or collar. The outer surface of the cup is smooth, but appears minutely
reticulate under a hand lens; the inner surface is perforated by the very numerous
openings of the exhalant canals, which show but little tendency to grouping.
The longer diameter of the sponge measures 37 mm., the shorter diameter, at right
angles to this, about 29 mm. The depth of the cup internally is about 14 mm. and the
maximum thickness of the wall about 7 mm. The thin oscular margin is under 2 mm.
in width.
The texture is firm but not very compact, and the colour in spirit pale wax yellow.
The canal system minutely resembles that of P. heteroraphis, excepting for the
enormous dilatation of the osculum, which finds a parallel, but in a still more exag-
gerated condition, in Grantia labyrinthica. The gelatinous ectosome is about 0°26 mm.
thick, and is broken up into trabecule by the small subdermal cavities, roofed over by the
thin, pore-bearing dermal membrane and opening below into the large inhalant lacune.
The ramifications of the inhalant lacunz roughly interdigitate with those of the exhalant
lacunze, which open into the central gastral cavity. In the mesoglea between the two
sets of lacunze are thickly scattered the spherical or subspherical flagellated chambers,
each about 0°1 mm. in diameter.
The arrangement of the skeleton is also identical with that found in P. heteroraphis,
and the spicules are very similar, though for the most part a little smaller in average
dimensions, as follows :—
(1) Large triradiates of the main skeleton (Plate 5, fig. 3, a); equiangular and
typically equiradiate; rays long, straight, slender and gradually sharp-pointed, measuring,
say, about 1-4 mm. in length by 0:07 mm. in diameter at the base.
(2) Small triradiates of the main skeleton (Plate 5, fig. 3, b; fig. 4, a); equiangular
and typically equiradiate ; rays straight, slender and gradually sharp-pointed, measuring,
say, about 0°18 by 0°015 mm. Intermediate sizes (Plate 5, fig. 3, c) between (1) and
(2) occur in comparatively small numbers.
* The specific name was suggested by the resemblance in external form to the fungus Peziza.
16 PERCY SLADEN TRUST EXPEDITION
(3) Small triradiates of the dermal skeleton (Plate 5, fig. 4, 6); irregular, but
probably really equiangular ; with slender, irregularly curved rays, often unequal in
length and sometimes blunt at the apex. One of the rays (x) is frequently bent inwards,
out of the tangential plane. Rays measuring, say, about 0°1 by 0°0055 mm.
(4) Quadriradiates of the gastral cortex; equiangular and typically equiradiate ;
with long, straight, slender facial rays, fairly sharply pointed and measuring about
0176 by 0°0125 mm.; with short, slightly crooked apical ray, not hooked and about
0075 mm. long.
(5) Quadriradiates of the larger exhalant canals. Similar to (4) but frequently
with very much longer, almost or quite straight apical rays, measured up to 0°29 mm.
in length.
(6) Quadriradiates of the oscular collar; sagittal, but otherwise resembling those of
the gastral cortex further in.
As in P. heteroraphis, the position of the nuclei of the collared cells is basal. The
ectosome contains large numbers of pigment cells filled with minute granules of a golden
brown colour. Similar, but less numerous pigment granules occur in the ectosome of
P. heteroraphis, probably also in special cells.
This species is distinguished from P. heteroraphis chiefly by its remarkable external
form.
Register No., Locality, dc. XxXvi., Cargados Carajos, 31.8.05, 12 fathoms.
Family Leucaltide Dendy and Row MS.
Sponge colony tubular and ramified, or even anastomosing, with many oscula; or
individualized, with large central cavity and single osculum. Wall of colony composed
of at least two distinct layers, viz. a dermal cortex with a strongly developed skeleton
of tangential radiates, and a chamber layer with skeleton greatly reduced or even
absent. A thin gastral cortex or membrane may or may not be present. Skeleton
composed, mainly at any rate, of equiangular radiates. No subgastral sagittal radiates.
Nuclei of collared cells probably always basal.
Genus Leucautis* Haeckel (emend.).
Sponge colony tubular, ramified and anastomosing. Flagellate chambers elongated
and branched, more or less radially arranged around the central gastral cavity.
I am now convinced that Bidder [1898] was quite right in considering this genus
to be closely related to the radiate Homoccela as represented by Dendya; its apparent
affinity with the Amphoriscidz is evidently due to convergence.
8. Leucaltis clathria Haeckel.
(Plate 2, figs. 1, 2.)
Leucaltis clathria Haeckel [1872].
Heteropegma nodus gordw Poléjaeff [1883].
* This genus is generally known under Poléjaeft’s name Heteropegma, but Leucaltis has priority.
DENDY—REPORT ON THE CALCAREOUS SPONGES 17
Leucaltis bathybia var. mascarenica Ridley [1884].
FHeteropegma nodus gordu von Lendenfeld [1885].
Clathrina latitubulata Carter [1886 ].
Heteropegma nodus gordi Dendy [1892].
Heteropegma latitubulata Dendy [1892].
Heteropegma nodus-gordit Dendy [1893].
Heteropegma nodus-gordw Hanitsch [1895 |.
Heteropegma nodus Gordw Bidder [1898].
Heteropegma nodus-gordi Dendy [1905].
Heteropegma nodus gordi Jenkin [1908].
This widely distributed species is well represented in the collection by a number of
fine specimens. It is easily recognizable by its external form, consisting of an irregular
mass of anastomosing tubes resembling a reticulate Lewcosolenia, on an enlarged scale.
This form has been well illustrated by Poléjaeff in the “Challenger” Report, and I need
only add that the diameter of the tubes varies considerably in different specimens and
is often much greater than represented in Poléjaeft’s figures.
It is unnecessary to figure the typical external form again, but I give photographs of
two specimens to show the range of variation in this respect (Plate 2, figs. 1, 2).
The wall of the tube always consists of three sharply defined layers, each with its
own characteristic spiculation. On the outside we have the well-developed dermal cortex,
supported by tangentially disposed triradiates and quadriradiates, of large but variable
size. The quadriradiates are commonly larger than the triradiates and the apical ray
projects inwards towards the gastral cavity, but the extent to which quadriradiates are
developed varies very greatly in different specimens and they may be very scarce.
The middle layer consists of the branching, radially arranged tubes, lined by collared
cells, whose walls are supported only by the very small, regular triradiates and quadri-
radiates, with very slender rays, and to some extent by the apical rays of the large
dermal quadriradiates.
The inner layer is a thin gastral membrane, extending between the exhalant openings
of the radial tubes and supported by a single layer of minute sagittal triradiates and
quadriradiates. The ends of the facial rays are often swollen, but to a very variable
extent, and the apical rays of the quadriradiates are often very long and thick in
proportion to the slender facial rays.
The general anatomy has already been figured by Poléjaeff and myself, and the
spicules have been figured more or less satisfactorily by Haeckel, Poléjaeff and Jenkin.
I am able to confirm Bidder’s statement as to the basal position of the nuclei of the
collared cells.
A careful examination of Haeckel’s type-specimen of Leucaltis clathria, fragments
of which were brought from Jena by my colleague, Mr R. W. H. Row, leaves no doubt
that it is a “ Heteropegma” and specifically indistinguishable from the species obtained
by the “Challenger” and named by Poléjaeff ““ Heteropegma nodus gordu.” Haeckel’s
figures of the large spicules are good. His fig. 3¢ is evidently a portion of the gastral
membrane with the characteristic minute sagittal tri- and quadriradiates. The shape of
SECOND SERTES—ZOOLOGY, VOL. XVI. 3
18 PERCY SLADEN TRUST EXPEDITION
these small spicules is, however, not well represented ; the sinuosity of the rays and the
swelling of their extremities being exaggerated. He does not figure the characteristic
small, regular tri- and quadriradiates of the chamber layer, though these are abundantly
present in his specimen. The canal system of the sponge, though no longer well
preserved, is evidently closely similar to that described by Poléjaeff and Dendy for
“ Heteropegma nodus gordi.”
Similarly an examination of a specimen of Ridley’s Leucaltis bathybia var. mas-
carenica, received from the British Museum, proves conclusively that this also is really
Leucaltis clathria.
In 1892 I showed that Carter’s Clathrina latitubulata is in reality a “ Heteropegma,”
but thought that it might be distinguished from H. nodus gordw of Poléjaeff by shght
spicular peculiarities. Further investigation has convinced me that such differences in
spiculation as exist cannot be regarded as of specific value.
Previously known Distribution. Coast of Florida (Haeckel); off Bermudas (Polé-
jaeff) ; Cape York, Torres Straits (Poléjaeff); near Port Phillip Heads (Carter, Dendy) ;
Ceylen (Dendy); West Coast of Portugal (Hanitsch); Amirante group, Seychelles
(Ridley), Wasin, E. Africa (Jenkin).
Register Nos., Localities, &c. XXx1x. 1 and xc. 3, Cargados Carajos, 45 fathoms ;
LXXVI. 3, xc. 2 and cv. 1, Amirantes, 28—29 fathoms; cx. 11 and cx. 2, Egmont
Reef.
Family Sycettide Dendy.
Flagellated chambers elongated, arranged radially round a central gastral cavity,
their ends projecting more or less on the dermal surface and not covered over by a
continuous dermal cortex strengthened by tangential dermal spicules. Tubar skeleton
articulate, with subgastral sagittal radiates. Collared cells usually confined to the
radial chambers in the adult, and probably always with apical nuclei.
Genus Sycon Risso (emend.).
Radial chambers usually more or less united in places where they come in contact
with one another, always crowned distally with tufts of oxeote spicules. Properly defined
inhalant canals usually present, which may be covered by a thin, pore-bearmg dermal
membrane without special skeleton.
Sycon? sp.
Three very minute Syconoid sponges were picked out from amongst other specimens
from Coetivy, but when I came to examine them microscopically, I found that all the
spicules had been completely dissolved out, presumably by some acid in the spirit in
which they were kept. They probably belong to the genus Sycon and are most likely
immature.
DENDY—REPORT ON THE CALCAREOUS SPONGES 19
Family Heteropiide Dendy.
Flagellated chambers varying from elongated and radially arranged to spherical and
irregularly scattered, with a distinct and continuous dermal cortex covering over the
chamber layer and pierced by inhalant pores. Subgastral sagittal radiates and subdermal
pseudosagittal triradiates present, with or without an articulate tubar skeleton. Nuclei
of collared cells probably always apical.
The term “pseudosagittal” is proposed by myself and Mr Row, in our forthcoming
paper on the classification and phylogeny of the Calcarea, for the subdermal spicules of
this family to distinguish them from ordinary sagittal triradiates, as we have come to the
conclusion that the “basal” ray is not homologous in the two cases. We have discussed
the nature of these spicules in the paper referred to.
Genus GRANTESSA von Lendenfeld (emend.).
Canal system syconoid. Dermal cortex without colossal longitudinally placed oxea.
Section A.
With large, usually radially arranged oxea, but without microxea.
9. Grantessa hastifera (Row).
(Plate 2, fig. 6.)
Grantilla hastifera Row [1909].
I have found three specimens of this interesting sponge associated with Leucascus
sinyplex amongst the branches of an Alcyonarian (Gorgonid) colony.
The species was first described by Row from a single specimen from the Red Sea, but
it is so well characterized that there can be no question of the specific identification.
I cannot, however, agree with Row in referring it to his genus Grantilla.
Row’s figure of the general anatomy as seen in transverse section is in the main
correct; but the exhalant canals of the radial flagellate chambers are represented much
too narrow, and the subgastral sagittal triradiates and subdermal pseudosagittal tri-
radiates are represented of about the same size, while the subgastral spicules are really,
on an average, a good deal stouter than the subdermal ones; moreover, the fine, hair-like
oxea (trichoxea) which project from the dermal surface are omitted.
The spiculation is as follows :—
(1) Dermal triradiates; lying tangentially in the dermal cortex, as described and
figured by Row.
(2) Subdermal pseudosagittal triradiates, with the two oral rays considerably
curved in a plane at right angles to the basal ray and thereby giving rise to deceptive
appearances of inequality according to the point of view.
(3) Subgastral sagittal triradiates, similar to, but typically stouter than the fore-
going. The characteristic bending of the oral rays, in a plane parallel to the gastral
surface, and the fact that they are of approximately equal length, are very clearly shown
when the sponge wall is examined from the gastral surface as a transparent object.
3—2
20 PERCY SLADEN TRUST EXPEDITION
(4) Gastral cortical triradiates, considerably more slender than the dermal tri-
radiates; as described and figured by Row (fig. 3, d; misprinted 3, a in the text).
(5) Large oxea; piercing the sponge wall at right angles and sometimes projecting
from both gastral and dermal surfaces. The outer end is very characteristically hastate,
much flattened and with a backwardly projecting tooth or barb on one side only,
something like a crochet needle. Row’s figures of this spicule are fairly characteristic,
but the barb in my specimens may be more strongly developed.
(6) ‘Trichoxea; slender hair-like oxea projecting from the dermal surface. These
spicules are not mentioned by Row but they occur in his preparations as well as in my
own. They do not appear to be true microxea.
In drawing up the above notes I have had the advantage not only of examining
Mr Row’s original preparations, but also of consulting him personally, and I am glad to
say that he entirely agrees with me on all points.
As so little is known of the external form of the species, I give a photograph of one
of the ‘‘Sealark” specimens (Plate 2, fig. 6).
Previously known Distribution. Red Sea.
iequster, Nos Uocahity, atc) xxi, ALB, (©) Providencey4 0:05) as) sae
78 fathoms.
Family Grantiide Dendy (emend.).
Flagellate chambers ranging from elongated and radially arranged to small,
spherical and irregularly scattered. With a distinct dermal cortex and a proper cortical
skeleton of tangential radiates, sometimes supplemented by, and occasionally replaced
by, oxea. Skeleton of chamber layer ranging from regularly articulate to irregularly
scattered. Subgastral sagittal radiates typically present. No subdermal pseudosagittal
triradiates. Subdermal quadriradiates, if present, always associated with a chamber-layer
skeleton of confused triradiates. Nuclei of collared cells probably always apical.
Genus GrantTIA Fleming (emend.).
Canal system syconoid. Colossal longitudinal oxea, if present, projecting from the
surface. Tubar skeleton articulate, composed of radiate spicules which may or may not
be supplemented by oxea.
Srction C.
With both large oxea and microxea.
10. Grantia indica n. sp.
(Plate 2, fig. 3; Plate 4, figs. 4, 5.)
This beautiful little sponge is represented in the collection by two specimens of
closely similar appearance. I select for description of the external form the one re-
presented in the photograph (Plate 2, fig. 3), which I have labelled B, the other, A,
having been used for microtome sections and spicule preparations.
DENDY—REPORT ON THE CALCAREOUS SPONGES 21
The sponge is a single Sycon person, which was evidently attached in life by a
contracted base, but there is no stalk. The body of the sponge has an inflated
appearance, especially on one side, and is surmounted by a well-developed oscular collar,
not a mere fringe of spicules but a thin membrane supported by a special skeleton. The
surface is coarsely bristly from the projection of large oxea, which point obliquely
upwards. The specimen figured measures about 10 mm. in total height, including the
oscular collar, and 4°8 mm. in maximum transverse diameter; while the height of the
collar is about 3mm. Specimen A was 9:25 mm. in total height, with a transverse
diameter of 5 mm. in the middle of the body, but the height of the oscular collar was only
1:75 mm. The colour in spirit is pale yellow.
The wall of the sponge, about 1°25 mm. thick in the middle of the body, surrounds a
wide gastral cavity. It gradually diminishes in thickness upwards to the base of the
oscular collar. The canal system is like that of Grantia, with long, unbranched (some-
times ? slightly branched) radial chambers, approximately circular in transverse section,
united laterally with one another throughout their length, and with narrow, irregularly
shaped inhalant canals in the interstices between them.
There is a thick dermal cortex (ectosome), with abundant mesogloea, overlying the
distal ends of the radial chambers. This cortex is excavated by large, irregular sub-
dermal cavities, which extend inwards, around and between the ends of the chambers,
and communicate with the inhalant canals. The small dermal pores are scattered over
the surface of the sponge.
There is also a thick gastral cortex, with abundant mesogloea which projects into the
gastral cavity around the bases of the apical rays of the gastral quadriradiates. The
gastral cortex is pierced by short exhalant canals, into which the radial chambers open
either singly or in groups. The junction of each chamber with the exhalant canal is
marked by a well-developed diaphragm.
The arrangement of the skeleton in most respects agrees with that of a typical
Grantia. The dermal skeleton is composed of stout, tangentially placed triradiates,
arranged in several layers, and the huge oxea project obliquely through it from deep down
in the chamber layer. The triradiates of the dermal cortex sometimes appear to be
invading the chamber layer. The gastral cortex is made up chiefly of the facial rays of
large gastral quadriradiates, whose strongly developed apical rays project into the gastral
cavity, accompanied by minute microxea. It also contains small quadriradiates which, in
several close-set tiers, surround the exhalant canal of each radial chamber, with their
apical rays projecting into the canal. The skeleton of the chamber layer is articulate and
many-jointed, the proximal joints being formed by characteristic subgastral sagittal quadri-
radiates with minute, rudimentary apical rays and long, straight, centrifugally directed
basal rays, often more or less grouped in bundles. The skeleton of the oscular collar is a
close interlacement of oxea and radiates. The former run lengthwise and parallel with
one another and with the basal rays of the radiates; the latter have very long, slender
basal rays and stouter paired rays extended almost at right angles to the basal ray, and
most of them have straight, sharp-pointed apical rays.
22 PERCY SLADEN TRUST EXPEDITION
The spicules (Plate 4, figs. 4, 5), may conveniently be grouped in eight categories, as
follows :—
(1) ‘Triradiates of the dermal cortex; typically sub-equiangular and equiradiate,
with centre of spicule slightly uplifted out of the facial plane ; rays fairly stout, gradually
sharp-pointed, measuring say about 0°22 by 0°02 mm. at the base, but variable.
(2) Triradiates of the articulate tubar skeleton; markedly sagittal, with widely
extended oral rays; rays nearly straight and gradually sharp-pointed. In a typical
example, from about the middle of the chamber layer, the paired (oral) rays measured
about 0°15 by 00146 mm. and the basal ray about 0°27 by 0°012 mm.
(3) Subgastral sagittal quadriradiates; markedly sagittal, with widely extended
oral rays, and long, slender, centrifugally directed basal ray, all gradually and sharply
pointed and remarkably straight, except for the usual bending of the oral rays out of the
facial plane in accommodation to the shape of the chambers or exhalant canals; with
minute, very slender, apical ray, vestigial or sometimes (?) absent ; paired rays measuring
about 0°146 by 0°01 mm.; basal ray about 0°345 by 0°01 mm.
(4) Quadriradiates of the gastral cortex; large and stout, with gradually sharp-
pointed facial rays which may or may not be differentiated sagittally, and with very
strongly developed apical ray, usually curved and gradually sharp-pointed, sometimes as
large or larger than the facial rays. In a fairly typical example all four rays measured
about 0°27 by 0°02 mm.
(5) Triradiates and quadriradiates of the oscular collar; very strongly sagittal ;
paired rays extended almost in a line with one another, nearly straight, gradually and
sharply pointed ; basal ray much longer and more slender than the paired rays, straight,
gradually and finely pointed. In a typical triradiate the paired rays measured about
0:25 by 0°01 mm., and the basal ray about 0°62 by 0:0083 mm. The orally directed
apical ray, when developed, is rather short, straight or nearly so, gradually and sharply
pointed, and measures about 0°075 by 0:0063 mm.
(6) Quadriradiates of the exhalant canals; these are much smaller than the gastral
quadriradiates ; their oral rays are usually considerably longer than the centrifugally
directed basal ray, and curved around the exhalant canal; the apical ray is very slender,
nearly straight, and all the rays are sharply pointed. The dimensions vary considerably ;
im a typical case the basal ray measures about 0°05 by 0:006 mm. ; the paired rays about
0:07 by 0:006 mm., and the apical ray about 0:025 by 0°003 mm.
(7) Large oxea; nearly straight, fusiform, gradually and sharply pointed, nearly
symmetrical but sometimes with a slight indication of hastate pointing at the outer end ;
occurring in the oscular collar and projecting from the dermal surface with their inner
portions deeply imbedded in the chamber layer; outer ends generally broken off; a
complete example of about average size measured 2°6 by 0°04 mm.
(8) Microxea; straight or slightly curved, slender, sharp-pointed at both ends,
hastate, with a slight enlargement about + of the total length from one end; size about
0:09 by 0-003 mm. (in the thickest part); observed only on the gastral surface, and there
only in small numbers.
DENDY—REPORT ON THE CALCAREOUS SPONGES 23
All the above descriptions and measurements (1—8) are taken from spicules in
sections, and not from boiled-out preparations, in which the isolated spicules cannot be
referred with absolute certainty to their proper positions.
Microxea closely similar to those of this species occur abundantly on both dermal and
gastral surfaces of Grantia labyrinthica [Dendy 1891, A].
Register Nos., Localities, de. xc. 4, A, B, Cargados Carajos, 1.9.05, B. 29,
45 fathoms.
Genus Leucanpra Haeckel (emend.).
Sponge usually a single person, or a colony of persons in which the component
individuals are readily recognizable. Canal system leuconoid. Skeleton of chamber layer
more or less confused, but frequently with vestiges of an articulate tubar skeleton in the
form of subgastral or other sagittal radiates. Dermal skeleton of tangentially placed
triradiates, which may sometimes develop an apical ray. Colossal longitudinally placed
oxea, when occurring in the dermal cortex, never forming a smooth layer, but always
projecting conspicuously from the surface.
Section A.
With large, usually radially arranged oxea, but without microxea.
11. Leucandra echinata Schuttner.
(Plate 2, fig. 4.)
Leucandra echinata Schuffner [1877].
Leucandra echinata Ridley [1884].
? Leucona echinata Carter [1886].
2 Leucandra echinata Dendy [1892].
Schuffner originally described his Lewcandra echinata from Mauritius, and I have no
hesitation in identifying two specimens in the “Sealark” collection with his species. The
spicules were figured by Schuffner, but as the external form has not yet been figured
I give a photograph of one of the “Sealark” specimens (Plate 2, fig. 4).
Carter, in describing his Lewconia echinata from near Port Phillip Heads, Australia,
makes no mention of Schuftner, and it is very doubtful whether he intended an identi-
fication of the Australian with the Indian Ocean species. The two are, however,
evidently very closely related and may be specifically identical, for my own observations
on Australian specimens indicate a wide range of variation. It is even possible that both
the Australian and Indian Ocean forms may ultimately be regarded as varieties of the
European Leucandra aspera (O. Schmidt), but a complete revision of the species of
Leucandra will be required to settle this and similar questions.
Previously known Distribution. Mauritius (Schuffner); Amirantes (Ridley) ;
? Australia (Carter and Dendy).
Register Nos., Locality, &c. uxxxix. 1, xc. 4, C, both from Cargados Carajos,
B. 29, 1.9.05, 45 fathoms.
24 PERCY SLADEN TRUST EXPEDITION
12. Leucandra wasinensis (Jenkin).
(Plate 2, fig. 5.)
Leucilla wasinensis Jenkin [1908].
A single small specimen (Plate 2, fig. 5), which agrees so closely with the description
given by Jenkin of his Leucilla wasimensis that I have no hesitation in making a specific
identification, was obtained from Saya de Malha. .
The specimen is a good deal smaller than that described by Jenkin, but the external
form calls for no special comment, being that of a typical single Leucon person; the
peristomial fringe of slender oxea is present, but inconspicuous, owing partly to erosion.
All the spicule forms described and figured by Jenkin are present and a direct comparison
with preparations of his sponge shows a close agreement between the two specimens in
this respect. I find that, in the Wasin specimen, the outer ends of some of the large
oxea are flattened, lancet-shaped, and this is apparently what Jenkin means by saying
that they are “snake-headed.” This does not appear to be a constant character, however,
even in the type; I cannot say whether or not it occurs in the “Sealark” specimen, as the
outer ends of nearly all the oxea are either broken off or more or less badly eroded. The
triradiates of the chamber layer are a good deal smaller in the “Sealark” specimen than
in the type, and this may be true to some extent also of the other spicules; but I do not
consider that there is any difference of specific value.
The most remarkable feature of the species is the presence of well-developed, centri-
petally directed apical rays on many of the radiates of the dermal cortex. Formerly,
I regarded this character as sufficient reason for the inclusion of a species possessing it in
the genus Leucilla, and such species as the present certainly seem to show how the
characteristic subdermal quadriradiates of the Amphoriscidz have arisen, but apical rays
so often appear on triradiates in all sorts of situations that I no longer regard their
presence in the dermal cortical radiates as sufficient by itself to justify the exclusion of a
species from the genus Leucandra. This point has been more fully discussed by Mr Row
and myself in our revision of the Calcarea.
Previously known Distribution. Wasin, East Africa (Jenkin).
Register No., Locality, dc. vit. 7, Saya de Malha, 6.9.05, C. 15, 55 fathoms.
Family Amphoriscide Dendy (emend.).
Flagellate chambers ranging from elongated and radially arranged to small,
spherical and irregularly scattered. With a distinct dermal cortex supported by a
skeleton of tangentially placed radiates, to which oxea may be added. Some or all of
the dermal radiates with large apical rays, which project inwards through the chamber
layer to a greater or less extent and form the principal part of its skeleton. No articulate
tubar skeleton, but subgastral sagittal radiates may be present and sometimes, in the
leuconoid forms, a confused skeleton of quadri-radiates in the chamber layer. Nuclei of
collared cells probably always apical.
DENDY—REPORT ON THE CALCAREOUS SPONGES 25
Genus Leucitia Haeckel (emend.).
Canal system leuconoid or sylleibid. Skeleton of chamber layer typically composed
of the centripetally and centrifugally directed apical rays of subdermal and subgastral
quadriradiates, but subgastral sagittal triradiates and confused chamber layer quadri-
radiates may be present, while the subgastral quadriradiates may be absent.
Section B.
With large oxea or trichoxea, but no microxea.
13. Leucilla proteus n. sp.
(Plate 2, fig. 7; Plate 5, fig. 5).
There are several very small specimens of a Leucilla, attached to an Avicula shell in
company with a number of other small organisms, for which it seems necessary to propose
a new specific name.
R. N. cvr. 1, B (Plate 2, fig. 7) may be taken as typical. The sponge is more or
less cylindrical in form, with a single vent at the free extremity. The length of the
specimen selected was about 7 mm. and the diameter 2°5:mm. The colour in spirit was
white.
The wall of the sponge, surrounding the central gastral cavity, is rather thin. The
canal system is sylleibid, almost if not quite syconoid towards the osculum, where the
flagellate chambers are thimble-shaped and more or less radially arranged.
The dermal skeleton consists mainly of the facial rays of large quadriradiates, with a
marked tendency to become sagittal, especially towards the osculum, and then also to be
oriented in the usual manner, with the unpaired ray pointing aborally.
The skeleton of the chamber layer consists mainly of the strong apical rays of these
quadriradiates, which commonly extend right through it and even project into the
gastral cavity. The comparatively slender basal rays of usually much less strongly
developed subgastral sagittal triradiates traverse the chamber layer in the opposite
direction. In one specimen at any rate there are irregularly scattered large radiates in
the older parts of the chamber layer. (In the same specimen I have also observed
numerous slender triradiates lying tangentially amongst the dermal quadriradiates.
I am inclined to regard this specimen as more nearly adult than the others; it is also
larger.)
The gastral skeleton is rather feebly developed, consisting of slender, tangentially
placed triradiates and quadriradiates, and of the oral rays of the subgastral sagittal
triradiates. Smaller quadriradiates may occur in the walls of the exhalant canals, towards
the gastral surface.
Tn some specimens, if not in all, there is a very feebly developed oscular fringe of
minute, slender oxea, and a few hair-like oxea may also occur, projecting here and there
from the dermal surface.
The spicules may be described under the following heads :—
SECOND SERIES—ZOOLOGY, VOL. XVI. 4
26 PERCY SLADEN TRUST EXPEDITION
(1) Quadriradiates of the dermal surface (Plate 5, fig. 5 a, b); with all four rays
strongly developed, stout and fairly sharply pointed ; facial rays lying tangentially to the
surface, commonly more or less sagittal; apical ray directed centripetally through the
chamber layer, either longer or shorter than facial rays; size variable, facial rays
measuring say about 0°39 by 0°035 mm.
(2) Quadriradiates of the gastral surface (Plate 5, fig. 5 d); much smaller than (1),
with slender, sharp-pointed facial rays and short, sharp-pointed apical ray projecting into
the gastral cavity ; facial rays measuring about 0°16 by 0:015 mm. These spicules become
strongly sagittal towards the oscular margin, with the recurved oral rays stouter than the
basal ray.
(3) Triradiates of the gastral surface (Plate 5, fig. 5); similar in all respects
to (2), except for the absence of the apical ray, and like them becoming strongly sagittal
at the oscular margin.
(4) Quadriradiates of the exhalant canals; much smaller than the gastral quadri-
radiates and with proportionally longer apical rays.
(5) Subgastral sagittal triradiates (Plate 5, fig. 5 c); with the two oral rays widely
extended in a line with one another beneath the gastral surface and the basal ray directed
centrifugally through the chamber layer. All rays gradually and sharply pointed. Size
very variable, averaging perhaps about 0°18 by 0°02 mm. for the oral rays and 0°33 by
0-02 mm. for the basal ray.
(6) Oxea; these occur chiefly in the oscular margin, where they form a very feebly
developed fringe. They are very slender, not very long, and very sharply pointed at each
end; almost hair-like, but sometimes with a distinct swelling at some distance from the
finely drawn out distal extremity, length say about 0125 mm. These spicules seem to be
intermediate between microxea and trichoxea.
This species is evidently very closely related to Leucilla australiensis (Carter)*, and
it may even prove to be specifically indistinguishable. The following differences may,
however, be indicated.
(1) The external form is cylindrical rather than ovoid.
(2) The sponge wall is thinner.
(3) The skeleton of the chamber layer is almost entirely composed of the apical rays
of the subdermal quadriradiates and the basal rays of the subgastral triradiates, whereas in
L. australiensis there is a well developed skeleton of confused radiates in the chamber layer.
(4) Oxea are present. These may or may not be constant and it is possible that
they may also occur in L. australiensis, though they have never been observed in that
species except in one doubtful case where a very few were found (possibly accidental).
It is highly probable that all the specimens of L. proteus obtained by the “ Sealark ”
are young and that some of the apparent differences between them and L. australiensis
may disappear in later life.
Register Nos., Localities, dc. cvi. 1, A, B, C, D, &c., Amirante, 13.10.05, E. 16,
39 fathoms.
* 11886, p. 133.]
DENDY—REPORT ON THE CALCAREOUS SPONGES 27
LIST OF LITERATURE REFERRED TO
1898. BrppErR, G. P. “The Skeleton and Classification of Calcareous Sponges” (Proc. Roy. Soc. Lond.
Vol. lxiv. pp. 61—76).
1886. Carter, H. J. “Descriptions of Sponges from the Neighbourhood of Port Phillip Heads, South
Australia” (Ann. and Mag. Nat. Hist., Ser. 5, Vols. xvii. pp. 40—53, 112—127, 431—441,
502—516, xvi. pp. 3455, 126—149, 271290, 369379, 445—466, pl. 10).
1891. Denpy, A. “Monograph of Victorian Sponges: Part I. The Organization and Classification of
the Calcarea Homoccela, with Descriptions of the Victorian Species” (Trans. Roy. Soc. Vict.
Vol. iii. pp. 1—82, pl. 19).
1891. Id. “Studies on the Comparative Anatomy of Sponges. II. On the Anatomy of Grantia
labyrinthica, etc.” (Quart. Journ. Micro. Sci. N.S. Vol. xxxii. pp. 1—48, pl. 1—5).
1892. Id. “Synopsis of the Australian Calcarea Heteroccela, with a proposed Classification of the
Group and Descriptions of some new Genera and Species” (Proc. Roy. Soc. Vict. N.S. Vol. v.
pp. 69—116).
1893. Id. “Studies on the Comparative Anatomy of Sponges. V. Observations on the Structure
and Classification of the Calcarea Heteroccela” (Quart. Journ. Micro. Sci. N.S. Vol. xxxv. pp.
159—257, pl. 10—14).
1905. Id. “Report on the Sponges collected by Professor Herdman at Ceylon in 1902” (Report Pearl
Oyster Fisheries. Roy. Soc. Lond. Part iii. pp. 57—246, pl. 1—16).
1872. HarcKEL, E. “Die Kalkschw&émme: eine Monographie.” 3 vols. 4to. Berlin.
1895. Hanirscu, R. “Notes on a Collection of Sponges from the west coast of Portugal” (Trans.
Liverpool Biol. Soc. Vol. ix. pp. 205—219, pl. 12, 13).
1908. JENKIN, C. F. “The Marine Fauna of Zanzibar and British East Africa, etc.—The Calcareous
Sponges” (Proc. Zool. Soc. Lond. 1908, 11. pp. 484—456).
1897. Kirk, H. B. “Notes on New Zealand Sponges; Fourth Paper” (Trans. Proc. N.Z. Institute,
Vol. xxx. pp. 313—816, pl. 31, 32).
1885. LENDENFELD, R. von. “A Monograph of the Australian Sponges. Part III. The Calcispongie ”
(Proc. Linn. Soc. N.S.W. Vol. ix. pp. 1083—1150, pl. 59—67).
1892. Mincuin, HK. A. “The Oscula and Anatomy of Leucosolenia elathrus, O.S.” (Quart. Journ.
Micro. Sci. N.S. Vol. xxxiii. pp. 477—495).
1900. Id. “Porifera” (Lankester’s Treatise on Zoology, Part II).
1883. Pontyarrr, N. DE. “Calcarea” (Report on the Scientific Results of the Voyage of H.MS.
Challenger, Zoology, Vol. viii. II. (Part 24) pp. 1—76, 1—9).
1884. Ripiuy, 8. O. “Spongiida” (Report on the Zoological Collections made in the Indo-Pacific
Ocean during the Voyage of H.M.S. Alert, 1881—2), Rep. Zool. Coll. Alert, pp. 366—482,
582—630, pl. 39—43, 53, 54.
1909. Row, R. W. H. “Report on the Sponges collected by Mr Cyril Crossland in 1904—5. Part I.
Calearea” (Journ. Linn. Soc. Zool. Vol. xxxi. pp. 287—400, pl. 35—41).
1877. ScHurrner, O. “Beschreibung einiger neuer Kalkschwimme” (Jenaische Zeitschrift, xi.
pp. 403—433, pl. 24—26).
1907. Werner, W. “Spongillenstudien V. etc.” (Arch. Naturges. Jahrg. 73, Bd. i. pp. 273—286).
28
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Mo oO 9 Po
GS Co SI GS Ox GS ©9 fo IP
5 3h
PERCY SLADEN TRUST EXPEDITION
DESCRIPTION OF PLATES
Pate 1.
(All the Figures in this Plate are from photographs.)
Leucosolenia gardinert n. sp. R.N. cxx. 7. x4.
Leucosolenia gardinert nv. sp. R.N. oxx. 11. x5.
Dendya prolifera n. sp. x 24.
Dendya prolifera n. sp. Upper portion of specimen, x 64.
Leucascus simplex Dendy. R.N. xx. 2,4. x 3.
Leucetta chagosensis n. sp. R.N. xix. 11. x 23.
Leucetta pyriformis n. sp. R.N. xc. 5,A. x5.
Pericharaz heteroraphis Poléjaeff. R.N. cxvi. x 24.
Pericharax peziza n. sp. R.N. XXVI. x 2.
PLATE 2.
(All the Figures in this Plate are from photographs.)
Leucaltis clathria Haeckel. R.N. oxi. 2. x 24.
Leucaltis clathria Haeckel. R.N. xxxix. 1. x 24.
Grantia indica n. sp. R.N. xc. 4,B. x 64
Leucandra echinata Schutfner. R.N. LXXXIxX. 1. x 4.
Leucandra wasinensis Jenkin sp. R.N. vu. 7. x6.
Grantessa hastifera Row sp. R.N. xxi 1. x4.
Leucilla proteus n. sp. R.N. ovi 1,8. x 6.
PLATE 3.
Leucosolenia gardineri n. sp. R.N. Cxx. 7. Spicules, x 370.
a. Triradiates; 6. quadriradiates.
Leucosolenia gardineri n. sp. R.N. cxx. 11. Spicules, x 370.
a. Triradiates; 6. quadriradiates.
Leucosolema gardineri nu. sp. R.N. cxx. 11. Part of a thin section stained with borax
carmine and picro-indigo carmine, x 666.
ep. flattened epithelial cells; nu. nucleus of ovum; ov. ovum (cytoplasm and yolk granules) ;
yb. yellow bodies.
4.
Dendya prolifera nu. sp. Section through two branching diverticula of the central gastral
cavity (radial tubes), x 30.
5.
C.
é.
RS)
2
c.g. central gastral cavity; b.d. branching diverticula (radial tubes).
Dendya prolifera n. sp. Spicules, x 278.
a. triradiates from the gastral membrane; 0. quadriradiate from the gastral membrane ;
triradiate from end of young radial tube; d. quadriradiate from middle of radial tube;
apical rays of quadriradiates from radial tube.
PLATE 4.
Leucascus simplex Dendy. R.N. xxi. 2, 4. Spicules, x 278.
a, triradiates; b. quadriradiates; a.r. apical ray.
Leucetta chagosensis u. sp. R.N. cxix. 11. Spicules, x 67.
Leucetta pyriformis n. sp. R.N. xc. 5,4. Spicules, x 107.
a. large triradiate from dermal surface; 6. small triradiates; c. intermediate triradiate ;
quadriradiate.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
DENDY—REPORT ON THE CALCAREOUS SPONGES 29
4. Grantia indica n. sp. R.N. xc. 4, A. Spicules, x 67.
a. triradiates; b. quadriradiates; c¢. oxea.
5. Grantia indica n. sp. R.N. xc. 4, A. Spicules, x 180.
a. equiangular triradiate, probably from dermal cortex; 0. sagittal quadriradiate, probably
subgastral; c. small quadriradiates, probably from exhalant canals; d. quadriradiates of gastral
cortex, side view (a.r. apical ray); ¢. microxea.
PLATE 5.
1. Pericharaz heteroraphis Poléjaeff. Spicules, x 36.
a, Large triradiates of main skeleton; b. small triradiates of main skeleton; c. triradiates of
main skeleton of intermediate size.
2. Pericharasz heteroraphis Poléjaeff. Spicules, x 180.
a. small triradiates of main skeleton; 6. triradiates of dermal skeleton (. bent rays, fore-
shortened); c. apical rays of gastral quadriradiates, side view; d. quadriradiates, probably from
larger exhalant canals (ar. apical ray); e. quadriradiate, probably from oscular collar.
3. Pericharax peziza n. sp. Spicules, x 36.
a. Large triradiate of main skeleton; b. small triradiates of main skeleton; c. triradiates
of main skeleton of intermediate size.
4. Pericharax peziza nu. sp. Spicules, x 180.
a. Small triradiates of main skeleton; 6. triradiates of dermal skeleton (a. bent rays,
foreshortened); c. quadriradiates (a.r. apical rays).
ig. 5. Leucilla proteus, n. sp. Spicules, x 106.
a. quadriradiates of dermal skeleton, side view (a.r. apical ray); b. quadriradiates of dermal
skeleton, facial views; c. subgastral sagittal triradiates; d. gastral quadriradiate; e. gastral
triradiate.
7 |
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Percy SLADEN Trust EXPEDITION.
(Denby)
TORVAINS, ILIUNIN, SOG, SIBIR, 2, ZOO'IL, WO)L, XW, PIL,
CALCAREOUS SPONGES
Percy Suapen Trust ExprDITIOoN.
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CALCAREOUS SPONGES
PERCY SLADEN TRUST EXPEDITION / TRANS. LINN. Soc. SER.2.Z00L.VOL.AVIPL. 3
(DENDY)
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Arthur Dendy, del. University Press, Cambridge
CALCAREOUS SPONGES.
PERCY SLADI
Arthur Dendy, del
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(DENDY)
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CALCAREOUS SPONGI
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Trans. Linn. Soc. Si
ER.2.Z00L.VOL.XVI Ph. 4.
University Press, Cambridge.
TRANS. Linn. Soc. Ser 2 Zoon Von. XVI Pu.5.
RCY SLADEN TRUST EXPEDITION.
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(DENDY)
, Cambridge
University Press
Arthur Dendy, del.
CALCAREOUS SPONGES.
No. II—SECOND REPORT ON THE ARACHNIDA—THE SCORPIONS,
PEDIPALPI, AND SUPPLEMENTARY NOTES ON THE OPILIONES
AND PSEUDOSCORPIONS.
By 8. Hirst.
(Text-figures 1—7.)
(Contributed by permission of the Trustees of the British Museum and communicated by
Professor J. STANLEY Garpiner, M.A., F.R.S., F.L.S.)
Read Ist May, 1913.
THE distribution of the Araneze was considered at some length in my first paper
on the results of this expedition, and therefore I will content myself in the present report
with making a few remarks on the other groups of Arachnida.
The species of scorpions occurring in the Seychelles are three in number, but one of
them is the cosmopolitan Isometrus maculatus De Geer. One of the two remaining
species—Lychas brauert Krpln.—is peculiar to these islands and its presence there is of
considerable interest, for the distribution of the genus to which it belongs is very
suggestive of the former existence of continuous land between the Oriental region and the
southern half of the African continent. The genus Lychas has four representatives in
Africa, two of them being tropical forms and the other two S. African. Another species is
only known to occur at Round Island, near Mauritius, whilst the genus is well represented
in the countries from India to Australia. Lastly there is the large black scorpion, Chiro-
machus ochropus C. L. Koch, which is also found at Round Island and at Zanzibar.
It is difficult to account for the distribution of this species except by former land
connections.
Four species of Pedipalpi inhabit the Seychelles, and three of them are restricted to
those islands, but these indigenous species are of small size, and our knowledge of the
groups to which they belong is still very incomplete; it would be useless, therefore, to
discuss their distribution. The remaining species, Tarantula scaber Gerv., has the same
distribution as the large black scorpion mentioned above, occurring at Round Island and
Zanzibar.
Tt is well known that Pseudoscorpions sometimes attach themselves to insects, and
this habit must help considerably in the dispersal of these little arachnids. They are most
often found clinging to the legs of flies, but occasionally also to those of other insects, and
sometimes they occur under the elytra of beetles*. Several Pseudoscorpions collected on
vertebrates have been forwarded to the British Museum during the last few years. The
* A very useful account of the pseudoparasitism of chelifers on other arthropods has been published by
Mr H. Wallis Kew (Naturalist, 1901, p. 193).
32 PERCY SLADEN TRUST EXPEDITION
data of these specimens is as follows:—1. A specimen found on a sheep at Tomotoku,
Gold Coast (H. Palmer). 2. One hundred and forty specimens captured by Dr S. A.
Neave on the skin of a mole-rat (Lachyoryctes audax Thos.), at Emba, east of Mount
Kenga. Some parasitic mites of the family Gamasidz were present in the same tube as
these chelifers. 8. Pseudoscorpion found on Anas undulata, Seziwa Swamp, Chagwe,
Uganda (S. A. Neave). 4. A specimen, Chelifer (Chernes*) sp., found on a mouse at
Jung-ting-shan, China (Capt. H. E. M. Douglas). These details may perhaps throw some
light on the distribution of Pseudoscorpions. Nos. 1—3 have been sent (together with
many other unnamed specimens of this order) to Mr E. Ellingsen for determination. On
receiving my list of them, Mr Ellingsen drew my attention to the fact that he had
suggested* that Chthonius tetrachelatus Preyssler—a European Pseudoscorpion recorded
by him as occurring in the Seychelles—might perhaps have been introduced into that
locality by birds. It seems quite possible that false scorpions may be carried from one
place to another by such means, but I think that it is more probable that this species has
been introduced into the Seychelles through human agency, viz. by trade. In his paper
on the Pseudoscorpions collected by the Danish Expedition to Siam, Mr C. J. With
discusses their geographical distribution, and mentions that several species of these
arachnids have been found at Copenhagen in rice on ships from India. Therefore this
means of transportation must not be overlooked.
It may, perhaps, be of interest to note that a species of Pseudoscorpion was collected
by Dr v. Willemoes Suhm on the isolated islets known as St Paul’s Rocks, during the
voyage of the “Challenger” (see Narrative of the Cruise, vol. 1. part 1, p. 205, 1885).
Several years ago I sent his specimens to Mr C. J. With, who identified them as Chelofer
garypordes Ell., a species discovered by Fea at Bolama, Portuguese Guinea. It seems to
me quite possible that this chelifer has been carried to St Paul’s Rocks whilst attached to
a bird ; otherwise it is extremely difficult to account for its occurrence there. Perhaps I
ought to add that the three species of birds which are found at St Paul’s Rocks are widely
distributed forms and occur also in Africa.
SCORPIONES.
1. Lychas brauert Krpln.
Archisometrus brauwert Kraepelin, Mt. Mus. Hamburg, 13, p. 123 (1896) ; Kraepelin,
Das Tierr., Scorp. &c. p. 46 (1899).
Localities. Mahé (in high jungle) ; Silhouette; Praslin. Prof. Kraepelin’s specimens
were captured on Praslin.
2. Isometrus maculatus De Geer.
Localities. Dennis Island and Bird Island, Seychelles. Astove. Specimens were
also obtained at Aldabra (Takamaka, Ile Esprit, and Ile Michel).
3. Chiromachus ochropus C. L. Koch.
Ischnurus ochropus, C. L. Koch, Die Arachn. iv. p. 69 (1838).
* Mitt. Zool. Mus. Berlin, iv. p. 402 (1910).
HIRST—REPORT ON THE ARACHNIDA OTHER THAN ARANEAD 33
Chiromachus ochropus, Pocock, Ann. and Mag. Nat. Hist. (6), xii. p. 320 (1893).
Ichnurus ochropus, Kraepelin, Das Tierr., Scorp. &. p. 153 (1899).
Chiromachus ochropus, Pocock, Ann. and Mag. Nat. Hist. (7), x. p. 364 (1902).
Localities. Three specimens from Ile au Récif and one from Praslin. Three
specimens were obtained by Mr H. P. Thomasset at Frigate Island, Seychelles, and
another specimen was collected on the same island by the naturalists on the Valhalla.
In the British Museum there are a number of specimens from Praslin (Madame la
Veuve F. Adam) and others from Round Island, near Mauritius (Sir H. Barkly), and from
Zanzibar.
PEDIPALPI.
1. Schizomus similis sp. n. (Text-figs. 1—3.)
Cephalothorax. Eyes apparently absent. Cephalic sternum longer than wide. Pale
stripe of second thoracic segment indistinct.
Fig. 1. Schizomus similis sp. n. Lateral Fig. 2. Schizomus similis sp. n. Lateral view of
view of palp. first leg.
Palp fairly robust. Anterior part of ventral surface of trochanter considerably pro-
duced so as to form a sharp angular projection. Femur very deep (high). Patella a little
more than twice as long as deep. Claw about half the length of the upper margin of the
tarsus (Fig. 1).
First leg much longer than the body ; its femur considerably longer than the tibia.
Foot very distinctly shorter than tibia, and about thirteen and a half times as long as
deep; second metatarsal segment almost as long as the united length of the first five
(proximal) segments of the tarsus; distal tarsal segment about equal in length to the sum
of the lengths of the three proximal segments and rather more than half the length of the
metatarsus (Fig. 2).
Femur of fourth leg a little less than two and a half times as long as its (greatest)
depth (Fig. 3).
Flagellum very long and slender, being about nine times as long as deep.
SECOND SERIES—ZOOLOGY, VOL. XVI. 5
34 PERCY SLADEN TRUST EXPEDITION
Length of body (not including flagellum) 4 mm. ; of first leg (from base of trochanter)
4°8 mm.
Colour rather dark brown.
Locality. Long Island (Mahé), Seychelles ; specimens from wood containing termites.
Remarks. As will be seen from the above description, this new species of Schizomus
differs considerably from S. latepes Hans., which was hitherto the only Tartarid known to
occur in the Seychelles. I have not seen any male specimens of it, and therefore will not
attempt to discuss its affinities with other species.
2. Tarantula scaber Gerv.
Localities. Mahé, Seychelles. In the British Museum Collection there are examples
of this species from Round Island, near Mauritius and also from Zanzibar.
3. Charinus seychellarum Krpln.
Charinus seychellarwum Kraepelin, Mt. Mus. Hamburg, vol. xv. p. 41 (1898) ;
Kraepelin, Das. Tierr., Scorpions, &. p. 250 (1899).
Localities. Mahé and Long Island; Silhouette ; Praslin ; Feélicité.
Fig. 3. Schizomus similis sp. n. Lateral view of femur of the fourth leg.
OPILIONES.
Family Phalangodide.
An important paper dealing with the Opiliones of this family (and also with the
family Assamiidee) has recently been published* by Dr C. Fr. Roewer. The presence or
absence of a scopula on the ventral surface of the tarsi of the third and fourth legs is one
of the chief characters which he uses to separate the Phalangodidee into subfamilies. This
character, however, does not appear to me to be a convenient one for the purpose, for this
scopula is sometimes very weak and indistinct in these Opiliones, and is then easily over-
looked ; moreover, in other species, the scopula is well developed only on the last two
(distal) segments of the tarsus. By using this character and also that of the number of
the segments of the terminal part of the tarsus of the first leg, Dr Roewer places, into
different subfamilies, genera which seem to me to be very closely allied to one another.
For instance the genera Sitalces KE. Simon, Podoctis Thorell, and Hrecanana Strand are
placed by him into the subfamilies Ibaloninze, Podoctinee and Erecananinze respectively,
whereas they resemble one another very closely in structure. Personally, I think that it
is probable that Hrecanana will ultimately prove to be a synonym of Stitalces. It is true
* Arch. Natg. A, 1912, heft 3, p. 111 &e.
HIRST—REPORT ON THE ARACHNIDA OTHER THAN ARANEAG 30
that the terminal part of the first tarsus is bisegmented in Sitalces and Podoctis, whilst it
is said to be undivided in Hrecanana, but, even if this is really the case, I doubt if it is of
generic importance.
It must be mentioned, however, that the number of the segments of the terminal
part of the tarsus of the first and second leg is constant in the family Cosmetidee, the
segments being always three in number (see Roewer, Arch. Natg. A, 1912, heft 10, p. 3),
but I do not think it follows that this character is of much systematic value in the family
Phalangodide. Personally, I agree with Dr Loman* and consider that the number of
tarsal segments of the legs is not a good generic character.
With regard to the employment of the presence (or absence) and number of thorns or
denticles on the scutum and free dorsal segments as a systematic character, it must be
remembered that there is often a considerable amount of variation in this respect—the
size and sometimes even the number of the spines varying in different individuals of the
same species. This feature must, therefore, be used with caution, even when determining
species. I am very doubtful about the use of the presence or absence of armature as
a means of distinguishing genera.
Fig. 4. Tbalonius inseriptus Loman. Lateral Fig. 5. Ibalonius inseriptus Loman.
view of palp of male. Lateral view of palp of female.
1. Ibalonius inseriptus Loman. (Text-figs. 4, 5.)
Tbalonius inscriptus +I. bimaculatus Loman, Zool. Jahrb. Syst. v. xvi. pp. 201 and 202
(1902).
Ibalonius inscriptus, Hirst, Trans. Linn. Soe. Ser. 2, Zool. xiv. p. 389, text-fig. 7 (1911).
Paribalonius inscriptus +P. bimaculatus Roewer, Arch. Naturg. (1912) heft 3,
je eo
Besides the sexual difference in the shape and armature of the cheliceree, adult
examples of this species also differ markedly from those of the opposite sex in the
armature of the palp, the proximal segments of this appendage being much more strongly
armed in the male than is the case in the female (see Figs. 4, 5); both of these sketches
are taken from specimens of the two-spotted form of the species (= 1. bmaculatus Loman),
but this sexual difference is also well-marked in examples of J. cnscryptus which are not
furnished with spots.
Locality. For localities see my previous report on the Aranev, Opiliones and Pseudo-
scorpions of the Seychelles, referred to above.
* Die Fauna Stidwest-Australiens, Vol. iii. pp. 127—134 (1910).
36 PERCY SLADEN TRUST EXPEDITION
Genus PHaLANGODES Tellkampf.
In his paper on the Phalangodidze, Dr Roewer states that the tarsus of the first leg
has three segments in Scotulemon and four in Phalangodes, and says that the former
-is restricted to Hurope and the Mediterranean region, Phalangodes occurring in North and
Central America. Unfortunately, however, he was not able to examine specimens of all
the known species. I take this opportunity of pointing out that the tarsus of the first
leg of one French species*—Phalangodes querilhaci, Lucas, has four distinct segments as
in the North American species which are referred by Roewer to the genus Phalangodes ;
the tarsus of the second leg of this species has ten segments, and its ocular tubercle is
well removed from the anterior margin of the cephalothorax.
10. Phalangodes? nanus sp. n. (Text-figs. 6, 7.)
Body. Cephalothoracic part of body much narrower than and not so high as the
abdominal part, the latter is widest at the posterior end of the scutum and is wider than
long.
Scutum slightly longer than the combined length of the patella and tibia of the fourth
leg. Transverse grooves five in number; the first one is deep and forms the boundary
between cephalothorax and abdomen ; it is not procurved but practically straight, and is
Fig. 6. Phalangodes? nanus sp. n. Palp Fig. 7. Phalangodes? nanus sp. n. Palp
from the inner side. from the outer side.
parallel with the four other grooves, which are very superficial. First area (cephalo-
thorax) much the largest, and it is only slightly shorter than the combined length of the
three first abdominal areas of the scutum. Surface of scutum unarmed ; there is a longi-
tudinal series of minute granules on each side of it, and some exceedingly minute granules
are also present on the abdominal part, but they are not numerous and are very difficult
to see. Ocular tubercle situated practically in the nuddle of the cephalothoracic area; it
is normal in appearance, being transversely elongated, low, and long oval in shape ;
neither processes nor granules are present on its surface. A very slight elevation is
situated in front of the ocular tubercle, between it and the anterior margin.
Free dorsal segments unarmed and without any distinct granules.
Cheliceree short, and, apparently, there are no distinct processes or granules either
on the first or second segments; the latter is not swollen.
Palp apparently more slender than is usually the case in the European species of the
genus. There are one or two obsolete granules on the lower surface of the trochanter,
and they bear fine sete. Femur armed ventrally with four spines on its outer side, the
* Our specimens of this species were presented to the Museum several years ago by M. Eugéne Simon.
Pp S g
HIRST—REPORT ON THE ARACHNIDA OTHER THAN ARANEA 37
two proximal ones being much the longest ; towards the distal end there is also an inner
row of three spines, the one which is situated nearest the distal end being much shorter
than the others. Patella with a long spine on each side, and these two spines are practi-
cally equal in length. Tibia with three very long spines and a little apical spine on the
outer side ; it has two long inner spines also. ‘Tarsus very short and its proximal end is
narrowed ; on each side it has two long spines (Figs. 6, 7).
Legs of moderate length ; there are some distinct granules on the posterior surface of
the coxa of the second leg and also a few minute granules on the anterior surface of the
coxa of the fourth. Femur of fourth leg not quite straight, its proximal end having
a slight inward curve, and the distal end an exceedingly slight outward curve, when
viewed from above. Tarsal segments 3, 5, 5, 5. Claws of posterior legs smooth (without
teeth).
Length 1:75 mm.
Colour. Body and appendages brownish ; patella, tibia and tarsus of palp slightly
infuscated ; the distal segments of the legs deeply infuscated, except for the distal ends
of the tarsi, which are usually brownish.
Locality. Mahé, Seychelles; a single specimen of the female sex.
y yy gle sp
The following species of harvest-man from the Seychelles has recently been described
by Dr Roewer :—
11. Metazalmouis ferruginea Roewer.
Arch. Natg., A (1912), heft 3, p. 135.
Seychelles (exact locality unknown).
PsEUDOSCORPIONS.
1. Feella affinis, Hirst, Trans. Linn. Soc. Ser. 2, Zool. xiv. p. 394, text-fig. 11
(1911).
Nore. Mr Ellingsen informs me that this species also occurs in India (Nagpur).
Us
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A
No. IJ.—ON A NEW LAND-LEECH FROM THE SEYCHELLES.
By W. A. Harprne, M.A., F.L.S.
(Plate 6 and 1 Text-figure.)
Read 16th January, 1913.
THE four specimens of a land-leech brought back by Professor Stanley Gardiner from
the Seychelles are the first examples of Hirudinea to be recorded from these Islands.
This species not only proves to be new, but possesses unusual features which, if not
singly, at least collectively justify the establishment of a new genus: it is worthy of note
also on account of its habitat, for although its restriction to such situations seems
improbable, the individuals so far obtained have to be added to the list of fauna
found in the moist humus which accumulates in the hollow leaf-bases of Pandanus
and palm~*.
Terrestrial leeches fall into two groups. Of one of these Zrocheta, a form not
unknown in England, may be taken as the type. It comprises several genera of
Herpobdellidee of carnivorous habit, with toothless, more or less vestigial jaws, which prey
for the most part upon earthworms.
It is another group however, the Hemadipsine, whose members are usually
indicated by the term “land-leech,” and to it the new form from the Seychelles is to
be referred.
This includes a number of voracious, blood-sucking parasites, typical of which is
Hemadipsa zeylanica, so familiar to the traveller in the hilly parts of India and Ceylon.
The Heemadipsine are essentially an Oriental group, although represented by several
more or less aberrant genera in other regions of the globe.
In all, seven well-established genera have been recognised hitherto; they all affect
considerable altitudes and are conspicuous by their absence from Madagascar and Africa.
Of these genera, Hamadipsa (Tennent, 1861) multiplies exceedingly throughout the
Oriental region ; it ranges as far north as Japan, as far east, it is said, as New Guinea
and the Pelew Islands, and probably requires further sub-division ; Xerobdella (von
Frauenfeld, 1868) occurs in the South Austrian Alps; Mesobdella, a form having affinities
with the Glossosiphonide, is found in Chile and was first described by Professor R.
Blanchard in 1893; the same authority has recorded (1894 and 1897) Phytobdella from
the Philippines and Moluccas and Planobdella (1897) from Celebes; Moquina
(R. Blanchard, 1888) and Philemon (R. Blanchard, 1897) are confined to Australia and
* Cf. R. C. Punnett, “On an Arboricolous Nemertean from the Seychelles.” Trans. Linn. Soc. Ser. 2, Zool.
xii. (1907), p. 57.
40 PERCY SLADEN TRUST EXPEDITION
Tasmania and differ from the rest in the possession of but two jaws, a peculiarity shared
by Idiobdella seychellensis.
The appearance of a new species possessing this remarkable characteristic of the
Australian land-leeches renders advisable the sub-division of the Hzemadipsine into the
following two series :
Series 1. Vrignathofere. With three jaws, one supero-median and two infero-
lateral. The bite inflicts the familiar y-shaped incision.
Including the genera Hamadipsa, Xerobdella, Phytobdella, Plano-
bdella and Mesobdella.
Series 2. Duognathofere. With two jaws, the supero-median member being
absent. The bite inflicts a nearly straight cut.
Including the genera Moquimia, Philemon, and Idiobdella (described
below).
It is to Moquima, with its typical or ‘“‘ complete
Idiobdella is most nearly related, since the ‘“ complete
ry)
somite formed of five rings that
” somite in Philemon is but
four-ringed.
[Moquinia is synonymous with the Geobdella of Whitman, 1886. The latter name
however, having been applied by de Blainville to Trocheta in 1828, justified the proposal
of the former name by R. Blanchard in 1888*. The first adequate description of the
Australian land-leeches was given by Miss Ada M. Lambert in 1897, 1898. ]
After the number of the jaws, the most striking character presented by Idiobdella
consists in the position of the genital orifices.
Apathy (1888) laid it down as an axiom that in all leeches the male opening occurs
in somite XI and the female in somite XII (somites X and XI, as counted by Whitman),
but it would have been more correct to have stated that these orifices occur within the
limits of these somites, since in Helobdella stagnalis the genital ducts open by a common
pore between them.
Even now, however, this statement admits of an exception, for in J. seychellensis the
male opening is situated in somite X and the female in somite XIII, an occurrence unique
among leeches.
The reproductive organs in the new species do not differ materially from those of
Philemon and Moquinia except in regard to the testes. Of these, only nine pairs exist
in I. seychellensis and the reduction has taken place anteriorly ; the anterior pairs show
a marked and progressive diminution in size, whilst the pair corresponding to the foremost
pair in allied species 1s absent.
The female apparatus as a whole, and also the much coiled epididymes, are bulky in
comparison with the size of the animal, and this probably bears some relation to the
unusual position of their respective external openings.
* Article “ Hirudinea” in Dictionnaire Encyclopédique des Sciences Médicales, Paris, 1888.
+ “The Structure of an Australian Land Leech” (Philemon), Proc. Royal Soc. of Victoria, Vol. x. (1897),
p- 211; also “ Description of two new Species of Australian Land Leeches” (Geobdella), loc. cit: Vol. xi. (1898),
p- 156.
HARDING—ON A NEW LAND-LEECH FROM THE SEYCHELLES 41
The “posterior organs” described by Miss Lambert as well developed in Philemon
and less conspicuous in Moquinia, were not observed in J. seychellensis, though it is not
impossible that they may be detected in fresh material more favourable for examination.
Somite VIT comprises four rings instead of three, as in Hemadipsa and Moquinia,
and the nephridiopores begin in the last ring of this somite and so differ as regards the
position of the first pair from the arrangement described by Miss Lambert in the
Australian land-leeches; again the ear-like lobes, termed “auricule” by Blanchard,
which overhang the posterior pair of these pores in most of the Heemadipsinze and form
a characteristic feature, are absent in Idiobdella.
I now proceed to give a brief diagnosis of this new genus’ and of the single species
upon which it has been founded.
IDICBDELLA, gen. nov.
[tSu0s strange, peculiar; Bd<AXa, leech. |
Hemadipsinee with two denticulate jaws. Complete somite formed of five rings.
Without auricule. No genital apertures in somites XI and XII.
Idiobdella seychellensis, sp. n.
The body of this little species has a rough or granular appearance due to the
presence of small, closely-set papillee disposed transversely on the rings. Posterior sucker
small and not wider than the widest part of the body.
Colour (after immersion in alcohol) in three individuals fawn, in the fourth brown,
uniform above and below. The dorsal surface is traversed by three deep brown
longitudinal stripes ; two paired rows of large papilla are conspicuous upon the ventral
surface, a pair of papillee occurring in each “complete” somite upon the third ring.
Five pairs of eyes situated respectively on rings 2, 3, 4, 5, and 8.
Rings, 97. Somites I—IV uniannulate, V and VI triannulate, VII with four rings,
the fifteen somites VIJI—XXII “complete” with five rings, XXIII with four rings,
XXIV—XXVII uniannulate.
Anus perforating the groove which separates the last ring from the posterior sucker.
Nephridia, 17 pairs. The nephridiopores occur, as is usual, in the marginal lines
of the body, and on the posterior edge of the rings which they pierce; the first pair open
on the posterior part of ring 14 (the last ring of somite VII); the last pair lie between
somites XXV and XXVI, whilst the 16th and 17th pairs are separated by six rings,
instead of five as in the case of the other pairs.
Genital apertures separated by eleven and a half rings, the male orifice being
situated between rings 28 and 29, that is to say, between the fourth and fifth rings
of somite X; and the female orifice lying upon ring 40, namely, the first ring of
somite XIII.
Size. The four examples (in alcohol) were in a fairly well extended condition and
were measured with the following results: No. 1, 13 mm. long and 2°25 mm. wide;
No. 2, 12°50 mm. long and 2°50 mm. wide; No. 3, 9 mm. long and 2°50 mm. wide ;
No. 4, 11°50 mm. long and 2:25 mm. wide.
SECOND SERIES—ZOOLOGY, VOL. XVI. 6
42 PERCY SLADEN TRUST EXPEDITION
Localities. Three specimens “from fallen and decaying leaf-base of palm in very
damp place, 1500 feet, Silhouette Island, Seychelles,” the fourth specimen “from a
Pandanus leaf-base, above Mare aux Cochons, Silhouette.”
Several issues, not wanting in interest, are raised by the discovery of Idiobdella
seychellensis.
XIV — XX
Tdiobdella seychellensis. Diagram of (A) dorsal and (B) part of ventral surface, showing eyes, annulation
and external openings. Somites numbered in Roman, and rings in ordinary figures. n.p., nephridiopores ;
an., anus.
In the absence of any evidence supporting the view that this species and the other
members of its Series are survivors of a type more primitive than that possessing three
jaws, we may assume, with more than a fair share of probability, that the Duognathoferee
are derived from some form not far removed from Hamadipsa, and are led to enquire by
what means the loss of the supero-median jaw was brought about.
The Gnathobdellide present a fairly complete series of forms ranging from typical
HARDING—ON A NEW LAND-LEECH FROM THE SEYCHELLES 43
blood-sucking species with fully armed jaws to those of carnivorous habit in which these
features are more or less vestigial or even absent.
Whatever interpretation be placed upon this scale of facts, whether it be read
upwards or downwards, one conclusion is the same: the change to which it bears witness
has been a gradual one.
But no intermediate stages are known to exist between the Trignathoferze and the
Duognathoferee, and it seems, therefore, not unreasonable to suppose that the dis-
appearance of the third jaw occurred suddenly; in short, that we have here an instance
of mutation.
Again it is difficult to understand how the presence of only two jaws, capable of
making a wound one-third less in size than that inflicted by three jaws, can be of any
benefit to a blood-sucking parasite and the doubt is increased by a consideration of the
comparative scarcity of the species provided with the lesser armament and the enormous
multiplication of the three-jawed forms.
The mutation, then, has not been to the advantage of the race and the Duognathoferze
are to be regarded as one of Nature’s unsuccessful experiments.
We know little, however, of the anatomy, still less of the distribution, and nothing
at all of the embryology of many of the known species of Haemadipsine ; other forms
probably await discovery, and further speculation seems idle in the present state of
our knowledge.
In conclusion, Idiobdella seychellensis is not merely at a disadvantage in respect of
its armament. In the only island in which it has so far been found there are no terrestrial
warm-blooded animals upon which it can prey and it probably sucks the blood and juices
of earthworms and other soft-bodied invertebrates.
It is the smallest of the Hemadipsine and its diminutive size, its reduced number of
testes, its lack of auricule, and its extreme scarcity substantiate the view that it is
a degenerating species in process of extermination.
EXPLANATION OF PLATE 6.
Fig. 1. Idiobdella seychellensis, from a specimen preserved in alcohol, dorsal view. x 4.
Fig. 2. The same, ventral view. x 4.
Fig. 3. The same, dorsal view of head region, showing the eyes. x 16.
Fig. 4. The same, dorsal view of posterior extremity. x 16.
Fig. 5. Ventral view of anterior region. The anterior sucker and part of the body posterior to it have
been slit up by a median incision exposing the two jaws. Greatly enlarged. ce. cut edge of body
wall; pe. entrance to pharynx; 7. jaw; s. interior of sucker.
Fig. 6. Reproductive organs. Greatly enlarged. The parts, which originally were tightly packed
together, have been opened out for the sake of clearness and the posterior pairs of testes are
omitted. ps. penis sac; ep. epididymis; v.d. vas deferens; ¢1, ¢2, £3, first, second and third pairs
of testes ; ov. ovary; p.ovd. and c.ovd. paired and common oviducts; v. vagina; vg.d. vaginal duct to
exterior.
, A Al Any
+ _ |
(a ae ie tite
is ; | Cate :
i ; 7 City : :
) Sethi,
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a
Percy SLADEN TRUST EXPEDITION. Trans. Linn. Soc. SzR.2.Z00L. VoL XVI. Pi.6.
(HARDING)
x76.
xl
Xlll
XIV
XV
WAH. & H.W. del. University Press Cambridge.
IDIOBDELLA SEYCHELLENSIS.
No. IV.—ON GYPSINA PLANA, CARTER, AND THE RELATIONS OF
THE GENUS.
By Marsorie Linpsey, B.Sc., The University, Manchester.
(Text-figures 1—6.)
(CommuntcateD BY Pror. J. Stanrey Garpiner, M.A., F.RS., F.LS.)
Read Ist May, 1913.
In the collections made by Prof. J. Stanley Gardiner in the Indian Ocean in 1905
specimens of a Foraminifer were obtained from Mauritius* (> 100 fathoms), Amirante
(25 fathoms) and Providence (50—78 fathoms) which have acquired a fixed mode of
existence to an extraordinary degree.
They form irregular spreading masses over any object on which they happen to grow,
generally old coral detritus, Polytrema, sponges and polyzoa—in short, on pretty nearly
any kind of decayed calcareous matter.
The most striking feature of this foraminifer, in addition to its encrusting habit, is
its extraordinary size, not only of the mass as a whole, but to a certain extent also of the
individual chambers. The specimens reach in some cases (e.g. those from Mauritius) three
or four inches diameter in their widest parts and they may spread until they completely
surround the decayed matter on which they are growing.
This incrustation is not simply a uniformly thin layer, but grows out into projections
forming an irregular series of knobs all over the surface.
The spirit material is of a pink colour, but it goes white on drying.
When magnified the surface+ is seen to consist of numbers of chambers arranged
more or less regularly with well defined walls separating them, giving it the appearance of
being divided into areolse, each roughly circular in shape and each perforated by a few
large foramina. These areole vary in size in the specimens from different localities, but
are largest in those from Mauritius, and these specimens, on account also of the larger
size to which the mass as a whole grows, have been mainly used for working on. Here
the areole vary from 90—233 », but an average chamber is somewhere about 120 in
diameter.
The areole are separated by grooves about 5» wide, which represent the walls.
The perforations are not very numerous and are from 10—20 in each areolz, each
being 5—7 p in diameter.
* See “Description of the Expedition,” Trans. Linn. Soc. Ser. 2, Zool., xii. (1907), p. 122.
tT For a figure of a similar surface see Challenger Report, ix. plate 102.
46 PERCY SLADEN TRUST EXPEDITION
The walls and foramina are of about the same size in the other specimens, but the
chambers as above stated are smaller, those from Providence varying between 80 and
150», with an average of 105» and from Amirante 49—115 p with an average of 89 p.
Fig. 1. Specimen of Gypsina plana, Carter, from 100—200 fathoms, oft Mauritius, nat. size.
A vertical section through the incrustation shows the chambers arranged in rows
parallel to the surface, the chambers of one row alternating more or less regularly with
those of the row above.
The typical form of the chambers as seen in such a section is more or less oblong, the
upper and lower walls being slightly convex outwards; the surface walls do not differ
from the others in any noticeable features, though on the whole they are somewhat flatter.
The chambers are seen communicating with one another by gaps in the wall
representing the foramina.
Fig. 2. Vertical section, magnified about 120 diameters.
On account of the presence of the irregular projections of more or less spherical
nature, such a true vertical section is rather difficult to obtain, and the most usual one
is a transverse section through one of these knobs.
In such a transverse section the cells are seen more or less concentrically arranged
round a common centre, especially in the outer layers, the inner ones being frequently
LINDSEY—ON GYPSINA PLANA CARTER, AND THE RELATIONS OF THE GENUS 47
irregular in arrangement, shape and size, and while those chambers towards the outside
of the knob are very similar in shape to those seen in a vertical section, the inner ones are
irregularly polygonal.
Fig. 3. Section through the centre of a knob, showing Fig. 4. Section showing the arrangement in the outer
the irregular arrangement, shape and size of the layers of a knob or in a section at right angles to the
chambers, also the perforated plates. x 96. surface in the ordinary thin part of the layer. x 75.
The foramina occur not only in the walls seen in section, but also in those in the
plane of the section, in which case the walls are seen as perforated plates.
Wirt
Fig. 5. Enlargement of portion of Fig. 4 to show the gaps in the walls of the chambers
as well as the perforated plates. x 212.
These perforations occur right up to the centre of the foraminifer, and not merely in
the superficial layers as in Homotrema and Sporadotrema, and they are all of the same
48 PERCY SLADEN TRUST EXPEDITION
kind, there being no evidence of the so-called pillar pores of Polytrema nor any other
large spaces or passages of any kind.
Compared with the foramina of the Polytrema group these are of relatively large size
(5p in this form and 1 pw in Polytrema) and relatively few in number.
The walls are seen in thin sections to consist of fibrous crystalline calcite; each
chamber has not its own individual wall, but is built up, so to speak, on the layer
below (Figs. 4 and 5).
No coiled initial chambers have been observed, but this is possibly due to the large
size of the specimens making search very difficult, and it is more than probable that the
mass did arise from some such coiled chambers as are found in other sedentary fora-
minifera. Once started on its encrusting mode of life it simply enveloped everything it
came across, as is well seen in some sections (unfortunately too thick to photograph) where
the foraminifer is seen surrounding Polytrema. Here the chambers immediately round the
Fig. 6. Section showing a shell of Polytrema in the centre surrounded by the chambers of Gypsina.
Polytrema are smaller and flatter than the ordinary ones, but do not differ otherwise, and
gradually merge into quite typical perforate chambers.
Some of the spirit material was decalcified and sections of the remaining soft parts
were obtaimed. These, however, were not very successful, as the material shrank
considerably on being embedded in wax, and they showed no further details except that
in one series there was a suggestion of an initial chamber, but this was unfortunately not
clear enough to make a definite statement about.
Briefly, then, the foraminifer may be described as follows: Sedentary, of large size,
the surface marked into clearly defined areolze about 120 in diameter and with the walls
of each chamber perforated by a few large foramina—5 p» in diameter. Below the surface
are seen rows of chambers communicating by perforations in the walls and not by large
open passages. There is no general aperture and no supplemental skeleton of spines and
tubercles as in Tinoporus.
A point worthy of notice in connection with this foraminifer is the occurrence of large
LINDSEY—ON GYPSINA PLANA CARTER, AND THE RELATIONS OF THE GENUS 49
numbers of sponge spicules embedded in it and penetrating it in all directions in exactly
the same manner as in Polytrema.
That these are foreign bodies incorporated in some way by the foraminifer and are
not part of its structure seems reasonably certain, as both the foraminifer and the spicules
are so typical of their kind. But it is nevertheless singular that these tetraxonid spicules
seem about the only kind of foreign matter picked up by the foraminifer, and they may
therefore be of some importance to the animal in the living condition; for it seems probable
that the spicules were picked up during growth, as they are too large and firmly imbedded
to have penetrated later by the foraminifer being overwhelmed by the sponge.
As to the affinities of this foraminifer, there are four genera all among the Rotaliidae
to which by its general appearance it might belong; these are Planorbulina, Cymbalopora,
Tinoporus and Gypsina. Planorbulina consists typically of a single layer of chambers,
spiral in the early stages but more irregular later, the chambers of successive whorls
alternating with one another and each having an opening at each end close to the line of
union with the previous whorl. In certain forms there are a number of small chambers
not arranged in any particular order which cover the surface, but as in the form under
discussion there are no specialised openings of any kind or any additional chambers on the
surface, its inclusion in this genus is not possible.
In Cymbalopora the successive chambers are not closely contiguous and are separated
by intervals which appear on the under surface as fissures, and each chamber opens
directly into a central cavity on this underside. Here again the presence of these
specialised openings renders it impossible to connect the two.
The original Tinoporus of Carpenter was characterised by the acervuline massing
of the chambers and by the absence of any specialised mouth or general aperture; in
some forms there was a supplemental skeleton of spines, &c., in others this was absent.
It was these latter which Carter in 1877 split off as his new genus Gypsina on
account of their greater simplicity of form, and it is to this genus that the foraminifer
under discussion really belongs, as shown by its general appearance, the relative size
of the foramina and cell walls, together with the absence of any supplemental
skeleton.
But it must be confessed that the original description of this genus was rather an
unsatisfactory one; it was, however, amended by Brady in the Challenger Reports, and
now reads as follows:
“Test free or attached, spheroidal or spreading, structure acervuline radiating or
laminated, chambers rounded or polyhedral; coarsely perforated. No supplemental
skeleton, no canal system and no general aperture.”
In the same paper in which this new genus was instituted, Carter points out that a
foraminifer previously described by him in a paper in 1876 as Polytrema planwm must
now be included in this new genus under the new name of Gypsina melobesiodes, as it
was merely a small specimen of a foraminifer with a melobesia-like growth; and the fact
that this specimen of Carter’s comes from Mauritius and was found coating old coral
detritus, &c., “like the white saccharine icing on bridecake,” to a very considerable extent
renders it probable that the material collected by Prof. J. Stanley Gardiner is really
SECOND SERIES—ZOOLOGY, VOL. XVI. 7
50 PERCY SLADEN TRUST EXPEDITION
the same thing. But neither under its old name of Polytrema plana nor its new one of
Gypsina melobesiodes is the description really clear.
As no figures were given in the later paper and those in the former were of specimens
41-inch in diameter, no real idea of the structure is obtaimable, and further, as the series of
canals shown in the original figure and described in the original description as the
distinguishing feature of the foraminifera were in the later paper shown to be non-
existent, the name G'ypsina plana (or melobesiodes) seems to be nothing more definite
than a name, and hence it is proposed to apply it to the specimens under discussion as
being in all probability the correct identification of at any rate the Mauritius specimens.
Of the two specific names, however, “plana” must take precedence over the more
suitable ‘“melobesiodes.” Except that the difference in size of the chambers in the
Amirante and Providence specimens makes them look more heavily built than the ones
from Mauritius, and that in the material under observation these latter incrustations were
much larger in size, there is no real difference in structure. Further, as both the size
of the chambers and the size of the mass vary considerably in the material from the same
districts, there seems no reason for making three species of them, and hence all must now
be known as Gypsina plana.
It also seems probable that an unidentified specimen from the West Indies in the
Halkyard Collection of Foraminifera in the Manchester Museum belongs to this species ;
the size of the chambers is rather smaller, being only 70—80 p, but it grows to the extent
of an inch or more over the surface of a shell of Strombus gigas, and is therefore
considerably larger than any other known Gypsina. If this specimen does really belong
to this species, it would seem to have a fairly wide distribution both in space and depth.
As reference has been made several times to the relatively large size of Gypsina
plana, it is perhaps advisable to have a few measurements of other forms of Gypsina
to compare with it. Carter gives no measurements at all of his type Gypsina
vesicularis, and only outside measurements (2°3 inches) of G. plana, nor are any to be
found elsewhere (except indirectly, as when the “Challenger” figures are said to be
magnified 30—40 times), but all are of small size, not bemg more than a millimeter
or so in outside diameter. No measurements of the areolz are to be found, but in the
specimens in the Halkyard Collection which were measured, the following seem to be
about the average sizes. (For convenience in comparison the measurements of the present
material are given also) :
G. plana, Mauritius as as ... 90—230 p
Re Providence aS: bee ... 80—150p
re Amirante site re .. 49—115 p
ie W. Indies Ae ae ... 70— 80p
G. unherens, Guernsey... a3 .. 70— 90p
G. vesicularis, Naples an a ... 380— 48p
G. globulus ... aye : 20— 28 p
So that, though the difference in size of the anoolae between G. globulus and
G. plana Mauritius is great, G. inherens leads up to it through the Amirante and
Providence specimens, the whole forming a gradual series.
LINDSEY—ON GYPSINA PLANA CARTER, AND THE RELATIONS OF THE GENUS 51
With the exception of the above-mentioned gradual increase in the size of the
chambers, there is very little difference of zoological importance on which to base the
species, and, indeed, by some all are grouped under the one species of G. globulus, and
even in those cases where the genus is sub-divided, the distinction lies mainly in the form
and habit and not in any real difference of structure.
The name Gypsina globulus is then used in the restricted sense in which Reuss first
used it—for the spherical forms, the subconical or convex forms constitute the species
“vesicularis "—the original type species of Carter (formerly Tinoporus vesicularis), while
the most usual encrusting forms of small size and fairly regular contour are grouped
under the name “ inheerens.”
But although these species are very artificial, it is perhaps the better way to let them
stand as being the most convenient way of describing the form and habit of any particular
specimen, and into this scheme of things Gypsina plana fits quite conveniently as being
the encrusting forms of far greater size than are covered by the designation Gypsina
imherens.
I am indebted to Prof. S. J. Hickson, F.R.S., for allowing me to examine this
material, and to him and Dr Hickling for assistance during that examination ; also to the
Manchester Museum for the use of the books and specimens of the Halkyard Collection.
‘LITERATURE.
BraDy, H. B. Foraminifera. Challenger Reports, Vol. ix. 1884.
CARPENTER, W. B., and Jones, Rupert. Introduction to the study of Foraminifera. Ray Soc. 1862.
Carter, H. J. On Polytrema. Annals and Mag. Nat. Hist. Ser. 4, Vol. xvi. 1876.
— On a Foraminifera with a Melobesia-like form of growth. Annals and Mag. Nat. Hist. Vol. xx.
1877.
Hickson, S. J. On Polytrema and allied Genera. Trans. Linn. Soc. Ser. 2, Zool. Vol. xiv. (1911),
pp. 443—462, pl. 30—32.
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No. V.—PELAGIC CRUSTACEA DECAPODA OF THE PERCY SLADEN
EXPEDITION IN H.MS. ‘“SEALARK.”
By Stanuey Kemp, B.A., Assistant Superintendent, Indian Museum, Calcutta.
(Plate 7 and 1 Text-figure.)
(CoMMUNICATED BY PRror. J. Stantey Garpiner, M.A., F.R.S., F.L.S.)
Read 19th June, 1913.
THE pelagic Decapod Crustaceans obtained by Mr J. Stanley Gardiner comprise only
thirteen determinable species represented by adult or sub-adult individuals. Possibly if
larger nets had been used the number would have been much greater ; for such actively
swimming animals as Decapods readily avoid capture, and even the net, 4 ft. square,
employed by Mr Gardiner was not large enough to obtain them in any numbers.
The collection is, however, a very interesting one, more particularly when comparison
is made between the species it contains and those found further to the north in the
Arabian Sea and the Bay of Bengal. Special attention has been paid recently to the
midwater fauna of these areas by the Royal Indian Marine Survey Ship “ Investigator”
and a large collection of pelagic Decapoda has been obtained. It is noteworthy that only
four of the thirteen species found by Mr Gardiner are common to the two collections. As
specific instances of this difference it may be mentioned that Acanthephyra sanguinea,
Alcock, the common species of the genus in the Arabian Sea and Bay of Bengal is replaced
in the present collection by the allied A. purpurea, A. M. Edw., and Lucifer acestra,
Dana, obtained by Mr Gardiner in great quantities, appears to be of the rarest occurrence
in Indian seas; the Sergestidee also of the two areas are wholly different. Gennadas
parvus, Bate, seems to be rare in the more northerly situations, and G. scutatus, Bouvier,
found in its typical form in the area surveyed by Mr Gardiner, is represented nearer the
coasts of India by a distinct race for which the name indicus is proposed.
I have to thank Mr Gardiner not only for entrusting me with this interesting
collection, but also for the patient manner in which he has waited for the completion
of my report, long delayed through pressure of other work.
54 PERCY SLADEN TRUST EXPEDITION
Tribe PENEIDEA.
Family Sergestidee.
SerceEstes, Milne Edwards.
1. Sergestes challengeri, Hansen (Plate 7, fig. 1).
Sergestes challengeri, Hansen, Proc. Zool. Soc. London, 1903, pp- 61 and 72, pl. xi, figs. 2, a—n; Kemp,
Proc. Zool. Soc. London, 1910, p. 641, pl. liii, figs. 2—4, pl. liv, figs. 2—5.
Sept. 27, 1905. 8. by E. of Farquhar; 10° 27’S., 51° 17’ E. p. 800 to 0 fms. ; one
male, 23 mm. q. 1000 to 0 fms. ; one young, 18 mm. and fragments of a larger female.
Oct. 16, 1905. 6 miles N.N.W. of Desroches Atoll. nn. 200 to 0 fms. one female,
29 mm.
This species is of particular interest in that it possesses a great number of luminous
organs. Hitherto it was known only from a single mutilated example obtained by the
“Challenger” expedition and described in detail by Hansen. The additional examples
have enabled me to supplement Hansen’s account of the structure of the photophores and
the results of this investigation have formed the subject of a separate paper”.
The specimens are unfortunately not in good condition, but their agreement with
Hansen’s account is so close as to leave no doubt of their specific identity with the
‘“‘Challenger” example found in the neighbourhood of the Fiji Islands. The only points
in which they differ from his description are that the rostrum (fig. 1) is directed forwards
and upwards, rather more horizontally than in Hansen’s figure and with a much more
acute apex; the hepatic spine of the carapace seems better defined and the penultimate
segment of the second maxillipede is distinctly longer than the antepenultimate. , In the
last feature they resemble the closely allied species S. gloriosus, Stebbing, a form which is
also provided with photophores. In the broad apex of the antennal scale, in the develop-
ment of the branchie and in all other details noted by Hansen there is complete
agreement.
The eyes, which were missing in the type specimen, extend a trifle beyond the middle
of the basal segment of the antennular peduncle and are a little shorter than the antennal
peduncle. In dorsal view the cornea occupies very nearly half the length of the whole
organ; it is scarcely at all depressed and, seen laterally, its greatest length is about
equal to its height.
I have been able, though not without considerable difficulty, to observe the photo-
phores in all the positions noted by Hansen and, in addition, have also found the
following, including those of the eyes and last two pairs of pereeopods which were missing
in the type: .
Eye: on the under surface of the stalk, one close to the cornea and one at the
proximal end.
Fourth perzeopod ; two on the ischium, one at the proximal and one at the distal end,
and one situated distally on both merus and carpus, the latter very minute.
Fifth perzeopod: same disposition as in the preceding, but the proximal photophore
on the ischium is missing.
* Kemp, Proc. Zool. Soc. London, 1910, p. 639.
KEMP—PELAGIC CRUSTACEA DECAPODA 595
Fourth and fifth abdominal somites: one additional photophore on each in the mid-
ventral line.
Sixth abdominal somite: an additional organ on each side of the infero-lateral margin
in the anterior quarter.
These twenty-two additional organs bring the total number known in the species up
to 139, and more will doubtless be found on the third maxillipedes and on the second
and third legs which have been wanting in all examples which have been examined.
A schematic view of the ventral surface of the cephalothorax and abdomen, showing the
disposition of the photophores, will be found in the accompanying text-figure™.
Mr Gardiner informs me that on each occasion when this species was taken he noted
the presence of phosphorescent prawns in the townet. This observation is
of importance, being, so far as I am aware, the only occasion on which light
has been seen to issue from a compound luminous organ or photophore in
a Decapod crustacean.
The closely allied Sergestes gloriosus, described by Stebbing from the
S. African coast, may be distinguished from the present species by the much
narrower apex of the antennal scale. It also possesses a far greater number
of photophores ; but, seeing that the example described measures 50 mm.,
whereas the largest known individual of S. challengeri is only 29 mm. in
length, this character may not prove reliable, for it is by no means im-
probable that the number of these organs increases with age. In this
connection it may be mentioned that in the male specimen of S. challengers,
23 mm. in length, the petasma is rudimentary.
In Stebbing’s figure of the ventral surface of the eye only one photo-
phore is shown, the proximal one, which is very distinct in the present
Species, being omitted.
2. Sergestes gardinert, sp. nov. (Plate 7, figs. 2—5).
SSonom (9004 Ss) by why von Harquhar; N07 272) a Zak:
p. 800 to 0 fms.; two young, badly damaged. gq. 1000 to 0 fms.; three
males, three females, 15—24 mm. sg, 750 to 500 fms. ; one female, 20 mm.
Oct. 6, 1905. Between Providence and Alphonse Ids.; 8° 16’ S.,
51° 26’ KE. aa. 900 to 0 fms. ; one male, 17 mm.
Oct. 16, 1905. Five miles off Desroches Atoll. Il. 750 to 0 fms. ; one
male, 21 mm.
The rostrum is high and projects obliquely upwards and forwards
(fig. 2). The apex is slender and acutely pointed and extends well beyond
the front. There is a prominent tooth in the middle of the dorsal edge.
The antero-inferior margin is concave, as is also the dorsal margin between
the tooth and the apex.
On the carapace there is no post-ocular spine and the hepatic is represented merely
* The black spots represent photophores and the circles the points of origin of the thoracic and abdominal
appendages. The divisions between the somites are shown by dotted lines.
56 PERCY SLADEN TRUST EXPEDITION
by a bluntly rounded lobe which, however, is conspicuous in dorsal view (fig. 3). The
gastro-hepatic groove is well marked laterally, but is feebly developed in the mid-dorsal
Ine. The cervical groove is distinct throughout.
The cornea of the eye is large, scarcely depressed, and is noticeably wider than the
stalk. Seen from above it is broader than long and its length is a trifle less than that of
the stalk. At the distal end of the stalk, on its inner and superior aspect, there is a small
round lobe, similar to that found in S. kroyert, Bate, but shorter and less cylindrical.
The basal segment of the antennular peduncle is about as long as the two following
taken together. The second and third segments are stout, the latter slightly longer than
the former. The breadth of the second is contained about two and a half times in the
length of its internal margin ; the third is scarcely three times longer than broad.
The antennal scale is not very broad at the apex (decidedly less so than in S. robustus
and S. bisulcatus); the lamellar part slopes directly away from the stout spine which
terminates the outer margin and does not project beyond its base as in several of the
allied species.
The basal segments of the third maxillipedes are not stouter than those of the third
pareeopods, and the two distal segments bear bristles on both margins. Above the base of
the third perzeopods are two well-formed pleurobranchs, the upper nearly half the length
of the lower. The upper branchia above the fourth pair of legs is a little longer than the
similarly situated branchia on the preceding segment ; it is about two-thirds the length of
the lower branchia.
The form of the petasma, shown in fig. 4, differs considerably in detail from that of
any species of the same group of which figures are extant.
The outer uropod (fig. 5) is four times as long as broad; the length of the ciliated
part at the distal end of its outer margin is exactly one-third of the total length of the
segment.
The specimens are not in very good condition, but the characters given above are,
I believe, amply sufficient for the recognition of the species. SS. gardineri belongs to the
robustus group of the genus Sergestes, which in Hansen’s amended account* comprises
eight species. It is readily separated from S. challengeri, Hansen, and S. gloriosus,
Stebbing, by the absence of photophores and from S. japonicus, Bate, and S. profundus,
Bate, by the large size of the eyes. From S. robustus, Smith, S. bisulcatus, Wood-Mason,
S. prehensilis, Bate, and S. kroyeri, Bate, it is distinguished by details in the structure of
the rostrum, eye, antennal scale and outer uropod. It is, apparently, in the last of these
species that it finds its nearest ally.
3. ?Sergestes edwardsi, Kroyer.
Sergestes edwardsi, Hansen, Proc. Zool. Soc. London, 1896, p. 961.
Sept. 27, 1905. S. by E. of Farquhar, 10° 27’ S., 51° 17’. r. 500—250 fms. ;
one male, ca. 18 mm.
June 30, 1905. 7 miles N.W. of Yéyé, Peros Atoll. N. 400—0 fms.; one male,
one female, ca. 13 mm.
* Hansen, Proc. Zool. Soc. London, 1896, p. 949, amended in Proc. Zool. Soc. 1903, p. 71.
KEMP—PELAGIC CRUSTACEA DECAPODA 57
Three specimens, in which the outer uropod is externally ciliated throughout its
length, appear to belong to this widely distributed species.
I have devoted considerable time to the remainder of the Sergestes in the collection,
but have been unable to satisfy myself of their identity. The specimens are not numerous,
and are, for the most part, in poor condition. Some appear to belong to the group
comprising S. halia, Faxon, and S. incertus, Hansen, while one male, 38 mm. in length,
seems to represent a new species allied to the Atlantic S. hensent, Ortmann. This specimen
is of considerable interest, but its damaged condition precludes a complete statement of its
specific characters. It agrees with S. henseni in the possession of two pleurobranchs at the
base of the third perzeopods, but differs in the absence of a supra-orbital spine and in the
armature and larger number of subsegments (apparently seven) in the terminal segment of
the third maxillipedes.
Lucrrer, Vaughan Thomson.
The members of this genus are very imperfectly known and, owing to the incomplete
accounts which most of the earlier authors have given, the synonymy of the various species
is very difficult to trace.
Vaughan Thompson*, when first describing the genus, omitted to give a specific
name to the form which he examined ; the figure probably represents ZL. acestra of Dana
(=L. reynaudiu of most recent authors) but of this there is no certainty.
I agree with Faxon that Milne Edwards’{ figure of L. reynaudi and his descriptions
of that species and of L. typus are quite insufficient for exact identification ; the account
given by Eydoux and Souleyet§, who regard these two forms as sexes of a single species,
is equally unsatisfying and it is only with the publication of Dana’s work in 1852|| that
a sound basis for the classification of the species is reached. Dana's treatment is fortu-
nately very good and surpasses that of nearly all subsequent writers; the identification
of all the forms which he has described should not be a matter of any considerable
difficulty.
The use of the nomenclature which Dana proposed has, however, one serious draw-
back: the name LZ. reynaudii must be employed in a different sense from that adopted by
most recent writers. The species which he recognised as that originally described under
this name by Milne Edwards—a species which must henceforth bear the name L. reynaudi,
M. Edw. (Dana)—is unquestionably distinct from the long-eyed form to which most
authors have assigned the name. The latter form, as Faxon has pointed out, is L. acestra,
Dana.
Dana did not refer any of the species he examined to JL. typus, M. Edw.; his
L. pacificus, according to Ortmann, is to be regarded as a synonym of that form, but no
* Vaughan Thompson, Zoological Researches, Cork, 1829?, Mem. iii, p. 58, pl. 7, fig. 2.
7 Faxon, Mem. Mus. Comp. Zool. Harvard, xviii. 1895, p. 214.
{ H. Milne Edwards, Hist. Nat. Crust. ii. 1837, p. 469, pl. 26, fig. 10.
§ Eydoux and Souleyet, Voy. de la “Bonite,” Zool. i. 1841, p. 249.
|| Dana, U.S. Explor. Exped. Crust. i. 1852, p. 668.
{| Ortmann, Zool. Jahrb. Syst. v. 1890, p. 454.
SECOND SERIES—ZOOLOGY, VOL. XVI. 8
58 PERCY SLADEN TRUST EXPEDITION
reasons for this course are given, and in view of the imperfection of Milne Edwards’
account its adoption cannot be recommended. L. pacificus, Dana, is without doubt distinct
from the species to which most authors have applied the name typus.
In Mr Gardiner’s collection there are two species of Lucifer. One, which was found
in great abundance, is L. acestra, Dana (= L. reynaudii, auct.), while the second, of which
only three specimens were obtained, is, I believe, Dana’s L. reynaudi, a species distinct
from the form common on the Indian coast, 1.e. the L. typus of most authorities.
4. Lnucvfer acestra, Dana.
Lucifer acestra, Dana, U.S. Explor. Exped., Crust., i, 1852, p. 671, pl. 44, fig. 9, and Faxon, Mem.
Mus. Comp. Zool. Harvard, xviii, 1895, p. 214 (synonymy).
May 16, 1905. N. of Chagos Archipelago; 3° 31'S., 72° 27’ BE. A, 25 to 0 fms. ;
two. 50 to 0 fms. ; five.
May 17—18, 1905. N. of Chagos Archipelago; 4° 16’ S., 71° 53’ KE. B, 25 to
0 fms.; one. CC. 1200 to 0 fms.; four. H. surface ; six.
June 30, 1905. 7 miles N.W. of Yéyé, Peros Atoll. M. 25 to 0 fms.; twelve.
75 to 0 fms.; one. N. 600 to 0 fms. ; seven.
June 30, 1905. 1—2 miles N.W. of passage into Salomon Atoll. O. 180 to
O fms.; one.
July 4—5, 1905. Off passage into Salomon Atoll. Q. surface; twenty-nine.
Aug. 22,1905. 5 miles W. of Black River, Mauritius. a, 25 to 0 fms. ; twenty.
Sept. 4, 1905. 8. of Saya de Malha Bank. m. 50 to 0 fms. ; three.
Sept. 7, 1905. Between Saya de Malha Banks; forty.
Sept. 9, 1905. N. of N. Saya de Malha Bank. n. surface; eleven.
Sept. 27, 1905. S. by E. of Farquhar, 10° 27’ S., 51° 17’ E. pp. 200 to 0 fms. ;
nine. 300 to 0 fms.; thirteen. 400 to 0 fms.; twenty-four. 500 to 0 fms.; eight.
600 to 0 fms.; seven. 800 to 0 fms.; seven. gq. 1000 to 0 fms.; many. s, 250 to
0 fms.; very many. ;
Oct. 1—2, 1905. N. of N. Farquhar. w. surface; two.
Oct. 2—3, 1905. Providence, off Cerf. x. surface; nine.
Oct. 6, 1905. Between Providence and Alphonse, 8° 16’ 8., 51° 26’ HE. aa. 900
to 0 fms.; twenty-five. bb. 140 to 0 fms. ; nineteen.
Oct. 7, 1905. S. of Alphonse. dd. surface; eleven.
Oct. 16, 1905. 38 miles N.W. of entrance into Desroches Atoll. kk. 50 to 0 fms. ;
four.
Oct. 16, 1905. 6 miles N.N.W. of entrance into Desroches Atoll. nn, 200 to
0 fms.; five.
5. Lucifer reynaudu, H. Milne Edwards (Dana).
Lucifer reynaudii, Dana, U.S. Explor. Exped., Crust., i, 1852, p. 672, pl. 45, figs. 1 a—d.
June 30, 1905. 7 miles N.W. of Yéyé, Peros Atoll. M. 25 fms.; one male.
July 5,1905. Off passage into Salomon Atoll. Q. surface; one female.
Oct. 7, 1905. S. of Alphonse Island. dd, surface ; one male.
KEMP—PELAGIC CRUSTACEA DECAPODA 59
The principal characters of the two male specimens are as follows :—
In the cephalothorax the neck is about one-quarter longer than the thorax. At the
junction of the two there is a prominent outstanding spine on either side, and at the
extreme anterior end there are five spines, one representing the rostrum, one behind each
eye, and one at each antero-lateral angle.
The eyes are club-shaped and, apparently, of somewhat variable size. In the two
specimens their length is contained respectively four and three-quarter and five and
a half times in the length of the cephalothorax. They are a little shorter than the basal
antennular segment.
The second segment of the antennular peduncle is about one and a half times the
length of the third; the antennal scale reaches to the middle of the former segment.
The last pair of pereeopods reaches to, or slightly beyond, the anterior extremity of the
cephalothorax.
On the first abdominal somite there is a pair of stout outstanding spines situated
subdorsally close to the cephalothorax, and there is a similar but more slender spine on
either side of the first five somites at the base of the pleopods. All these spines are
conspicuous in dorsal view. The sixth somite is a little shorter than the two preceding
taken together. It terminates dorsally in a sharp spine, and on its inferior margin bears
two large teeth, the first of which, situated in the middle of the ventral margin, is smaller
and more sharply pomted than the posterior. Behind the latter tooth there is a pair of
small and very sharp spinules.
The telson reaches to about half the length of the outer uropod and is provided with
a prominent rounded lobe on its inferior aspect, a little behind the middle point. The
telson bears two pairs of dorsal spinules, the anterior being situated immediately above
the proximal edge of the ventral lobe. The apex is slightly concave, with a long spine at
each outer angle and two shorter pairs between them, the median pair being the shortest.
There are minute prickles on the inner side of the outer spines and on both sides of the
two inner pairs.
The inner uropod reaches to a point midway between the apices of the outer uropod
and telson. The outer uropod is a little more than five times as long as broad, and
the spine which terminates the outer margin reaches well beyond the apex of the
lamella.
The specimens measure about 8°5 and 9 mm. respectively from the tip of the eye to
the apex of the outer uropod.
The single female is of considerably larger size, measuring 12 mm., and differs in
several details from the male. The eyes are decidedly smaller, their length being
contained six times in the length of the cephalothorax. The sixth abdominal somite
is unarmed below, except for a pair of fine spinules in the same position as those found in
the male. The distal armature of the telson exactly resembles that of the male; the
outer uropod has the same proportions, but the spine on the external margin does not
surpass the apex of the lamella.
The specimens differ from Dana’s description of individuals from the Sooloo Sea
in having the cephalothorax proportionately shorter; the pereeopods, moreover, are
8—2
60 PERCY SLADEN TRUST EXPEDITION
considerably longer than is indicated in his figure. I believe, none the less, that the
specimens may safely be referred to the same species: the presence of the spinules on the
inferior margin of the last abdominal somite in the male appears to constitute a valid
specific criterion.
Lucifer typus, auct., as typified by a number of specimens from the Ceylon coast,
is very closely allied to this species; but males may easily be distinguished from it
by the invariable absence of the pair of spinules behind the posterior tooth on the ventral
edge of the last abdominal somite. Both sexes differ from Dana's L. reynaudw in the
proportions of the outer uropod. These segments are broader in L. typus, not more than
four and a quarter times as long as wide, and the spime which terminates the outer
margin does not reach so far backwards.
Family Peneide.
GENNADAS, Spence Bate.
Considerable difficulties are encountered in the determination of the species of
Gennadas, and this seems more especially to be the case with the several closely allied
forms occurring in the Indian Ocean, the Bay of Bengal and the Arabian Sea. The
species found in these waters are, for the most part, very closely allied, and the sexual
characters are almost the only features of diagnostic value. These characters are, indeed,
well defined, and their use renders the recognition of any species which has been
adequately figured a comparatively simple matter: the difficulty lies in recognising the
two sexes of a single species. Gennadas parvus, Bate, and G. sordidus, Kemp, afford an
instance of this. Both species have been described from the male sex only, and though
the petasmata yield distinctions of a quite obvious nature, in other respects the characters
of the two are so closely similar that very little reliance can be placed upon them for
specific determination. In examining Mr Stanley Gardiner’s collection several females
were found which must certainly be referred to one of these two species, while others
which must also be identified with one of the two forms, but have a different type of
thelycum, are included in a fine collection of Gennadas recently made in Indian waters
by the R.I.M.S.S. “Investigator.” On consideration, I have decided that Mr Gardiner’s
specimens represent the female of G. parvus, while those obtained by the “ Investigator ”
are females of G. sordidus. In making these determinations I have, however, been
largely influenced by the fact that males of G. parvus, which are very scarce in the
“Investigator” material, are found to the exclusion of male examples of G. sordidus
in Mr Gardiner’s collection. The conclusions, therefore, are not based on evidence of
a really satisfactory nature and, should further investigations reveal the existence of a
third form of male exhibiting the same affinities in general structure that exist between
G. parvus and G. sordidus, the matter will be still further complicated.
6. Gennadas parvus, Spence Bate (Plate 7, figs. 6, 7).
Gennadas parvus, Kemp, Proc. Zool. Soc., 1909, p. 721, pl. 83, figs. 1—6, pl. 75, fig. 1.
Sept. 27, 1905. S. by E. of Farquhar, 10° 27’S., 51° 17’ EK. p, 800—0 fms. ; one
female. gq. 1000 to 0 fms. ; one male and four females. gs. 750 to 500 fms. ; two females.
KEMP—PELAGIC CRUSTACEA DECAPODA 61
Oct. 16, 1905. 4 miles N.W. of Desroches Atoll. Il. 750 to 0 fms.; one male.
The two male specimens agree closely with the description of the type published
in 1909. The mid-dorsal carina of the carapace is, however, continued as far as the
posterior margin, and the distance between the gastro-hepatic and cervical grooves”,
though somewhat variable, seems always to be less than one-quarter the distance from
the latter groove to the posterior edge. The description of the type specimen, which was
badly damaged, is probably unreliable in these respects.
Considerable variation exists in the form of the mandibular palp, and this is note-
worthy owing to the importance which Bouvier has assigned it as a specific determinant.
In the type the ultimate segment is longer than the greatest width of the basal segment ;
but in the two males in the present collection the proportions are different. In one the
last segment is in length about equal to the width of the basal segment, while in the
other it is even shorter.
The third segment of the endopod of the first maxillipedes is, in these specimens,
only one and a half times the length of the second. In the first pair of pareeopods, which
was missing in the type, the carpus is a little more than two-thirds the length of the
merus, and is about as long as the chela.
The petasmata of the two specimens are as nearly as possible in exact agreement
with that of the “Challenger” example (Joc. cit., pl. lxxv, fig. 1). The distal lobes are, as
usual, bent inwards and outwards in a very curious manner and, when seen from below,
present the appearance shown in fig. 6.
Six females, four of which were taken in the same net as one of the males
already mentioned, are also referred to this species, the female of which was previously
unknown. The evidence for this determination is, as I have outlined above, somewhat
meonclusive.
The form of the thelycum is illustrated in fig. 7. It bears a very close resemblance
to that of G. tunayrez, Bouvier, a species which in this respect exhibits a rather unusual
amount of variation and has hitherto been found only in the Atlantic. In the present
examples, however, the design is very constant and differs from that of G. tenayres
in the presence of an additional pair of tubercles between the legs of the third and
fourth pairs.
7. Gennadas scutatus, Bouvier.
Gennadas scutatus, Bouvier, Rés. Camp. Sci. Monaco, xxxiii. 1908, p. 42, pl. 8.
Oct. 16, 1905. Near Desroches Atoll. mm. 400 to 0 fms.; one male, one female.
The specimens differ from Bouvier’s description in a few unimportant details. The
endopod of the second maxilla bears from two to four curved spines on its dorsal aspect
near the apex. The third segment of the endopod of the first maxillipede is broadly oval
in the male specimen, while in the female it is narrow, as shown in Bouvier’s figure. The
widest part of the merus of the second maxillipede is, in one specimen, situated much
closer to the distal end of the segment than is indicated by Bouvier. The apex of the
* In previous papers I have called these grooves the cervical and post-cervical respectively; the terminology
here employed seems preferable.
62 PERCY SLADEN TRUST EXPEDITION
telson, which is only extant in the male example, is slightly convex and is furnished with
four pairs of plumose setze.
The petasma is quite typical and resembles that of the Atlantic specimen figured
by Bouvier* rather more closely than that of the “Challenger” specimen} from the
N. Pacific, and is noticeably different from that of the individuals recorded from the
Indian Ocean, 8S. of Cape Comorm. Several additional examples bearing the closest
resemblance to those from Cape Comorin have recently been obtained by the “ Investi-
gator.” The differences which are noted in Rec. Ind. Mus., 1910, p. 178, pl. xin, fig. 10,
are in my opinion sufficiently constant and well marked to entitle the form to subspecific
rank. It may be known as G. scutatus, subsp. indicus.
The thelycum of the female specimen collected by Mr Gardiner has the curious form
figured by Bouvier and, except that there is a small median notch in the distal or anterior
margin of the long plate lying between the last three pairs of legs, agrees exactly with
his account. The triangular biperforate plate between the legs of the third pair is
clearly seen.
The examples are unfortunately in very bad condition, and in both the third maxilli-
pedes and all five perzeopods are wanting. When complete neither specimen appears to
have exceeded 20 mm. in length.
8. Gennadas alcocki, Kemp (Plate 7, fig. 8).
Gennadas aléocki, Kemp, Rec. Ind. Mus., 1910, v. p. 174, pl. 13, figs. 5—8.
Sept. 27, 1905. S. by HE. of Farquhar; 10° 27’S., 51°17’ KE. p. 800 to 0 fms. ;
one female, 24 mm.
Oct. 16, 1905. 4 miles N.W. of Desroches Atoll. 1], 750 to 0 fms.; one female,
26 mm.
The basal segment of the endopod of the first maxillipede is, in one specimen,
provided with only two spines on the inner margin, whereas in the type five are found
in this situation. Bouvier, in his exhaustive memoir on the Atlantic species of this
genus, defines the proportions and spinulation of the oral appendages at considerable
length ; but my own experience leads me to believe that these details are very variable
and of little taxonomic value.
In the two examples of G’. alcock: contained in the present collection the thelycum
is closely similar in pattern to that of the type, but in both specimens the plate lying
between the legs of the third pair is shaped even more clearly in the form of a W than
in the original example (see fig. 8).
9. Gennadas sp.? (Plate 7, fig. 9).
Oct. 16, 1905. 4 miles N.W. of Desroches Atoll. Il, 750 to 0 fms.; one female,
ca. 24 mm.
The rostral crest is of the usual form, and is continued backwards as a carina which
extends to the posterior margin of the carapace. Both antennary and infra-antennary
* Bouvier, Bull. Mus. Océanog. Monaco, 1906, No. 80, text-fig. 13, p. 11.
+ Kemp, Proc. Zool. Soc. 1909, Pl. 75, fig. 2.
KEMP—PELAGIC CRUSTACEA DECAPODA 63
angles are sharply acute, and in dorsal view a small but distinct branchiostegal spine is
visible. The distance between the gastro-hepatic and cervical grooves, measured dorsally,
is about one-quarter the distance from the latter groove to the posterior margin of the
carapace.
The antennal scale appears to have been very strongly narrowed distally : its tips
are broken. The second segment of the antennular peduncle is long ; seen in lateral view
its dorsal measurement is more than three-quarters that of the ultimate segment.
_ The second segment of the mandibular palp is, in length, just equal to the greatest
width of the basal segment. The internal lobes of the second maxilla are widely sepa-
rated from one another, but the anterior lobe of the internal lacinia is not wider at the
apex than at the base as in Gennadas precox and G. calman. The posterior lobe of the
external lacinia is about one and a half times as wide as the adjacent lobe of the internal
lacinia. At its base the slender apex of the endopod bears on its external aspect two
curved spines. The basal segment of the endopod of the first maxillipede is furnished
with three spines ; the third segment is about one and a quarter times the length of the
second.
In the first pair of pereeopods the carpus is about equal in length to the chela and
the dactylus is shorter than the palm. In the second pair the chela is a trifle shorter than
the carpus and is a little more than two-thirds the length of the merus. The last three
pairs of legs are missing and the tip of the telson is broken off.
The thelycum is an elaborate structure quite unlike any which have hitherto been
observed. Fig. 9 will convey a better idea of its appearance than a long description.
The large cordiform plate between the legs of the last pair is free and unattached to the
sternum in its anterior third.
The female on which the above description is based represents, I believe, a species
hitherto undescribed. If I am right in my supposition that the undescribed females in
Mr Gardiner’s and in the “Investigator” collections belong respectively to Gennadas
parvus and to G. sordidus, the only species known from the Indian Ocean of which the
female remains unrecognised is G'. precox, and it seems clear from the very different shape
of the lobes of the second maxilla that the present specimen cannot be assigned to that
form. From male Gennadas parvus and G. sordidus it may be distinguished by the
comparatively greater length of the second segment of the antennular peduncle.
Two female specimens from the vicinity of Farquhar Island, 1000 to 0 fms., seem
to belong to the same species as that described above; but they are very badly
mutilated.
Tribe CARIDEA.
Family Hoplophoride.
HoptopHorus, H. Milne Edwards.
10. Hoplophorus gracilirostris, A. Milne Edwards.
Hophorus gracilirostris, A. Milne Edwards (Oplophorus), Ann. Sci. Nat. Zool. (Sér. 6), xi. 1881, art. 4, p. 6
and Recueil figs. Crust. nouv. ou peu connus, 1883, pl. 29; Alcock, Desc. Cat. Indian deep-sea Crust. Macrura
and Anomala, 1901, p. 73.
64 PERCY SLADEN TRUST EXPEDITION
Oct. 16, 1905. 3 miles N.W. of Desroches Atoll. kk. 250 to 0 fms.; one young,
27 mm.
I am unable to find any valid distinction between this small specimen and larger
examples of H. gracilirostris determined by Alcock. The rostrum is of great length, one
and a half times that of the carapace, and the spine on the third abdominal somite is of
enormous size, reaching as far as the posterior edge of the fifth somite and completely
overshadowing that of the fourth somite which has the same proportions as in the adult.
These characters, in my opinion, are correlated with post-larval growth and do not afford
specific distinctions.
The rostrum bears ten small teeth above and eight below, and there are twelve
spinules on the outer margin of the antennal scale. The carinze of the carapace and the
spines on its antero-lateral and postero-lateral edges are closely similar to those of adult
examples.
11. Hoplophorus foliaceus, Rathbun.
Oplophorus foliaceus, Rathbun, Bull. U. 8. Fish Commission for 1903 (1906), xxiii, pt. 11., p. 922,
text-figs. a, ),
Sept. 27, 1905. 8. by E. of Farquhar; 10° 27’S., 51°17’ E. p. 500 to 0 fms.; one
young, ca. 25 mm.
In this young example the apex of the rostrum is broken off. Except that the third
pareeopods are proportionately somewhat shorter it agrees exactly with Miss Rathbun’s
account. The long spines on either side of the finely crenulated apex of the telson
appear a little shorter than in Miss Rathbun’s figure.
The spine on the second abdominal segment—a feature which at once distinguishes
this species from all other members of the genus—is very strongly developed ; its apex
reaches beyond the middle of the fourth somite. The spine on the third somite reaches
the posterior margin of the fourth, while that on the latter somite is about half its length.
The branchiostegal spine is prominent, but does not form the termination of a carina as
in H. gracilirostris, and the lateral carinze of the rostrum posteriorly curve downwards
behind the orbit and are not continued backwards on either side of the median crest as in
that species. The two posterior rostral teeth are situated on the carapace behind the
longitude of the orbital notch.
Hoplophorus foliaceus was described by Miss Rathbun from specimens obtained in
337 and 442 fathoms in the vicinity of the Hawaiian Islands.
AcANTHEPHYRA, A. Milne Edwards.
12. Acanthephyra purpurea, A. Milne Edwards,
Acanthephyra purpurea, Kemp, Fisheries, Ireland, Sci. Invest. for 1905, p. 4 (synonymy) and Coutiére,
Bull. Mus. Océanog. Monaco, no. 70, 1906, p. 12.
May 17, 1905. N. of Chagos Archipelago, 4° 16’S., 71° 53’ E. ©, 1200 to 0 fms. ;
one male, three females, 52—72 mm.
Sept. 27, 1905. S. by E. of Farquhar, 10° 27’S., 51°17’ E. q, 1000 to 0 fms.; one
male, one female, 66 and 69 mm., and one damaged post-larval example.
KEMP—PELAGIC CRUSTACEA DECAPODA 65
Oct. 16, 1905. 4 miles N.W. of Desroches Atoll. Il. 750 to 0 fms.; two young,
22 and 36 mm.
Acanthephyra purpurea has not hitherto been recorded from the Indian Ocean.
Judging from the fact that no specimens have been obtained by the “Investigator,” it
seems probable that it does not occur in the more northerly portions of this region, but is
replaced in the Arabian Sea, the Bay of Bengal and the Andaman Sea by its near ally,
A. sanguinea. From this species, as Alcock has pointed out, it is readily distinguished
by the presence of a prominent branchiostegal spine which is buttressed by a short
carina. ‘
Little indication of the great variation which is known to exist in the species is
shown in the six adult examples contained in the collection. The rostrum is in all cases
longer than the carapace and its dorsal and ventral teeth are well spaced; the telson
extends noticeably beyond the uropods and bears from four to five dorso-lateral spinules.
In these features the specimens closely resemble the type (from the Atlantic) figured by
A. Milne Edwards, and differ from the form found most commonly, though not invariably,
in the N.E. Atlantic. Coutiére (Joc. cit., p. 18) recognises the form with numerous
spinules on the telson under the name, var. multispina, but considering the enormous
range of variation which is known I am not of the opinion that such details merit
recognition in the nomenclature.
The specimens in the collection exhibit the following rostral armature :
BEEGG4 B18
The eyes are in most cases damaged. In examples in which they exist the cornea
is of a dark reddish brown and is not jet-black as in all other examples that I have
examined. Differences in eye-colour seem to afford useful taxonomic indications; in the
present case the colour may have been altered by prolonged immersion in formalin, though
such treatment does not appear to have had this effect in Notostomus and Sergestes.
Notostomus, A. Milne Edwards.
In determining the solitary example of this genus contained in Mr Gardiner’s collection
I have had occasion to consider the known species as far as it is possible to do so from the
published descriptions, and, with a view to affording some slight assistance to those who
may subsequently be working at the genus, I have appended a partial synopsis of the
twelve that have been described.
The knowledge acquired within recent years of the considerable post-larval changes
that occur in an allied genus leads one to suppose that, with the acquisition of more
abundant material, the present taxonomy of Notostomus must be subjected to revision.
In species of Acanthephyra there is often a great range of variation in regard to the
length of the rostrum and the dorsal armature of the abdominal somites, and, in the case
of the former character, it has been shown that this variation is largely correlated with
growth. In very young post-larval examples of Acanthephyra purpurea the rostrum is
short ; at a later stage, before the fully adult phase is attained, it has a comparatively
great length, whereas in the adult it again becomes relatively short, but not so short as
SECOND SERIES—ZOOLOGY, VOL. XVI. 9
66 PERCY SLADEN TRUST EXPEDITION
in the early post-larval stages. That this extraordinary development also takes place in
Notostomus admits of very little doubt. Smith has, indeed, remarked that in an example
of NV. robustus, 53 mm. in length, the rostrum is very much longer than in two other
specimens measuring 135 and 150 mm. respectively. In the following synopsis the
characters employed are drawn for the most part from the carination of the carapace ;
there is reason to believe that this feature does not alter during the course of late post-
larval growth.
Partial synopsis of the species of Notostomus*.
I. ‘he lateral carina of the rostrum terminates near the post-orbital carina but is nut continuous with it.
A. In the posterior half of the carapace the customary short carina immediately below the post-orbital is
absent ; there is no post-antennal carina ... : aon sib ace oa Sragilis, Faxon
B. A short carina is present in the posterior half of the carapace immediately below the post-orbital ; a
post-antennal carina is distinct, at least in the hepatic region.
1. The post-antennal carina is indistinct behind the ridge separating the branchial and hepatic
regions ... sos as 06 : ie ot Bis abe vescus, Smith.
2. The post-antennal carina is aves ond divtitact throughout almost the whole length of the
carapace.
a. The post-orbital carma curves downwards in its extreme anterior part, but behind this is
practically straight.
i. The dorsal ridge of the carapace, viewed laterally, is straight over a considerable part of
its median portion... Sie “els ae ae: 300 ne japonicus, Bate.
ii. The dorsal ridge of the carapace is very strongly arched in lateral view,
patentissimus, Bate; westergreni, Faxon ; longirostrist, Bate.
b. The post-orbital carina curves downwards in its extreme anterior part, and behind this is
descendant to the upper limit of the branchio-hepatic ridge and ascendant from this point
to the posterior margin ... as ae 300 906 S60 go robustus, Smith.
II. The lateral carina of the rostrum is continuous with the post-orbital.
A, There is no carina in the posterior half of the carapace immediately below the post-orbital,
gibbosus, A. M. Edw. ; elegans}, A. M. Edw. ; murrayit, Bate.
&. A short carina is present in the posterior half of the carapace immediately below the post-orbital,
perlatus, Bate; brevirostris, Bate.
13. Notostomus perlatus, Spence Bate (Plate 7, fig. 10).
Notostomus perlatus, Spence Bate, Voy. H.M.S. “Challenger,” xxiy., Crustacea Macrura, 1888, p. 831,
pl. 124, fig. 2.
1 Notostomus brevirostris, Spence Bate, ibid., p. 832, pl. cxxxiv., fig. 4
May 17,1905. N. of Chagos Archipelago; 4° 16’S., 71° 53’ E. 1200—0 fms. ; one
male, 138 mm.
* For descriptions of the species the following papers should be consulted :
A. Milne Edwards, Ann. Sci. Nat. Zool. (Sér. 6), xi. 1881, art. 4, p. 7 and Recueil, figs. Crust. nouv.
ou peu connus, 1883, pls. 19, 40.
Smith, Ann. Rep. U.S. Fish Commission for 1882 (1884), p. 377 and id. for 1885 (1886), p. 72.
Bate, Voy. H.M.S. “Challenger,” xxiv. Crust. Macrura, 1888, p. 824.
Faxon, Mem. Mus. Comp. Zool. Harvard, xviii. 1895, p. 170.
Notostomus corallinus, A. M. Edw., does not seem to be correctly referred to this genus.
7 The position of this species is doubtful.
KEMP—PELAGIC CRUSTACEA DECAPODA 67
The rostrum is slender, slightly damaged at the extreme apex, and reaches to about
two-thirds the length of the antennal scale. On the upper margin there are four small
widely-spaced teeth, while a fifth, which appears to have existed near the apex, is broken
off. On the lower margin there are six similar but larger teeth.
The carapace is very strongly arched dorsally and its greatest depth is more than
half its extreme length including the rostrum. In the mid-dorsal line it is very sharply
carinate and is furnished with a series of small spinules about 70 in number, which extend
from the rostral base as far as the posterior margin. Laterally there are several pro-
minent longitudinal carinee, three of which extend throughout almost the whole length of
the carapace. The uppermost of these, the post-orbital carina, commences very close to
the posterior margin, defines the upper limit of the branchial region, and is continuous
anteriorly with the carima on the side of the rostrum. Before reaching the base of the
latter it gives off a small branch from its lower side which runs towards the back of
the orbital notch, but disappears before reaching the margin. On comparison with other
species, such as NV. armata, Smith, it will be seen that this branch is the real anterior
extremity of the post-orbital carina. Another short carina curves upwards and forwards
from the margin of the orbital notch and fuses with the lateral keel of the rostrum. In
the posterior half of the carapace, immediately below the orbital and parallel with it,
there is a short carina and there are also two long carinze near the inferior margin. The
uppermost of the latter, the post-antennal carina, begins very near the posterior margin
and terminates anteriorly, before reaching the margin, in a sharp outstanding spine which
projects over the base of the antenna. This carina is connected with the orbital by a
well-defined oblique ridge separating the branchial and hepatic regions. The other
earina, the marginal, runs close to the inferior margin, fading away shortly before reaching
the anterior end of the carapace; it is continuous behind with the fine keel that borders
the posterior edge. On the anterior margin a sharply rounded lobe defines the inferior
limit of the orbit and immediately below this lobe there is a short spine. The antero-
inferior angles are broadly rounded. The surface of the carapace is finely corrugated,
especially on its postero-lateral portions and on either side of the dorsal crest.
The eyes are well pigmented ; the cornea is broader than the stalk and is set very
obliquely on it. The small ocellus is distinct in dorsal view and is connected with the
cornea. The antennal scale is about two and a half times as long as wide; its distal
margin is convex and terminates in a spine which reaches a little beyond the narrow apex
of the lamellar part. The upper (and outer) ramus of the antennular peduncle is greatly
thickened at the base. This swollen portion comprises rather more than fifty joimts and
extends, in the single male examined, beyond the apex of the antennal scale by two-thirds
the length of that organ.
Tn the second maxilla the two lobes of the distal lacinia project beyond the proximal
lacinia, the whole appendage resembling that of Acanthephyra purpurea. The endopod
of the first maxillipedes differs from that of Hymendora, and resembles Acanthephyra in
being composed of three distinct segments. The third maxillipedes reach a little beyond
the apex of the antennal scale, as do also the pereeopods of the third and fourth pairs.
9—2
68 PERCY SLADEN TRUST EXPEDITION
The fifth pareeopods reach about to the apex of the scale; the dactylus is exceedingly
small and is hidden in a dense tuft of setze.
There are sharp carinz on all the abdominal somites, those of the last four
terminating in long spines. The telson is about twice the length of the sixth somite
and extends beyond the apices of the inner uropods. Dorsally it bears two carinze which
are connected anteriorly and join again shortly before reaching the very narrow apex.
There appear to have been two pairs of dorso-lateral spimules on the telson in its
posterior third.
The identity of this specimen with that described by Spence Bate from 2150 fms. in
the neighbourhood of Celebes is, I believe, certain. J. brevirostris from the Atlantic
coast of 8. America is synonymous with N. perlatus if it really differs from that form
only in the trivial distinctions mentioned by Bate.
EXPLANATION OF PLATE 7.
Fig. 1. Sergestes challengeri, Hansen, anterior part in lateral view.
Vig. 2. Sergestes gardineri, sp. nov., anterior part in lateral view.
Hig. 3 F 7 ss anterior part im dorsal view.
Fig. 4 - 5 ne petasma.
Fig. 5 a A fe outer uropod.
Fig. 6. Gennadas parvus, Bate, lobes of petasma seen from beneath.
Fig. 7 ss 5 » thelycum.
Fig. 8. Gennadas alcocki, Kemp, thelycum.
Fig. 9. Gennadas sp., thelycum.
Fig. 10. WNotostomus perlatus, Bate, lateral view.
Percy Sirapen Trust Exprpition.
(Kemp) TRANS. LINN. SOC. SER. 2, ZOOL. VOL. XVI. PIL, 7
S. Mondul. A. Chowdhary; del.
PELAGIC DECAPODA
No. VI.—POLYCHATA OF THE INDIAN OCEAN, TOGETHER WITH
SOME SPECIES FROM THE CAPE VERDE ISLANDS.
THe SERPULIDA, wiTH A CLASSIFICATION OF THE GENERA HyYDROIDES AND
HKuPoMATUS.
By Hexen L. M. Prxety, B.Sc., F.Z.S., Demonstrator of Zoology and Reid Fellow,
Bedford College, Unversity of London.
(Plates 8 and 9.)
(CoMMUNICATED BY Pror. J. StantEy GArpinER, M.A., F.R.S., F.LS.)
Read Ist May, 1913.
Tue Serpulids described here belong to five collections. Two of these were made by
Prof. J. Stanley Gardiner, F.R.S., one from the Maldive Archipelago in 1899 and the
other from the Seychelles and Chagos Groups in 1905.
The three other collections were made by Mr Cyril Crossland, M.A.: the first during
1901—2 in the neighbourhood of Zanzibar, the second during some parts of the years
1904—8 in the Red Sea and the third in the Cape Verde Islands in July and
August, 1904.
I take this opportunity ot thanking both Prof. Stanley Gardiner and Mr Crossland
for the opportunity of examining these specimens, most of which are beautifully preserved
and of considerable interest. I should also like to thank Dr Marett Tims for his help
and interest during the preparation of the paper.
The collections consist of 22 species and three varieties belonging to ten genera. To
include these forms one new genus has had to be made, viz. Pomatolezos but I think two
have been shown to be superfluous, viz. Hucarphus Mérch and Glossopsis Bush. Seven
of the species and two varieties seem to be entirely new. The genera Spirobranchus and
Hydroides ave especially well represented. The following is a list of the species showing
to which collection they belong—the letters M., SC., Z., R., V. representing Maldive,
Seychelles or Chagos, Zanzibar, Red Sea and Cape Verde Islands respectively. The
previously known distribution is indicated in brackets.
1. Serpula vermicularis, Lin. SC. Z. R. V. (very wide distribution).
2. Hydvrordes norwegica, Gunn. Mediterranean ? (Mediterranean, English Channel,
N. Sea). .
70 PERCY SLADEN TRUST EXPEDITION
3. Hydroides homoceros, n. sp. M. Z.
4 Xn heteroceros, Grube, Z. RK. (Red Sea, Ceylon).
5. - bifurcatus, n. sp. M.
6. tA monoceros, Gravier, Z. R. (Red Sea).
7. Eupomatus exaltatus, Marenz. Z. R. (Japan, Ceylon).
8 ms spinosus, n. sp. R.
9. Pomutostegus stellatus, Abildgaard, R. (W. Indies, Florida, Red Sea).
10. Spirobranchus giganteus, Pallas, M. Z. R. (W. Indies, Florida, Red Sea).
Lil, te ys var. turbinatus, nov. M.
12. i gardineri, n. sp. SC.
13. fe sempert, Morch, Z. R. (Philippine Is., Ceylon).
14. Fy 6 var. acroceros, Willey, Z. R. (Ceylon).
15. As i » aceros nov. SC.
16. £8 contieret, Gravier, Z. R. (Red Sea).
U7, a maldiviensis, n. sp. M.
Pomatoleios, ng.
18. s crosslandi, n. sp. Z.
19. Vermilopsis pygidialis, Willey, M. Z. R. (Ceylon). _
20. Salmacina dysteri, Huxley, Z. R. (Atlantic, Mediterranean, Red Sea, English
Channel, N. Sea).
21. Protula tubularia, Mont. V. (Atlantic, Mediterranean, English Channel,
N. Sea).
22. Protula intestenum, Lam. SC. Z. (Mediterranean).
23. Spirorbis papillatus, n. sp. Z.
2A. be levis, Quatrefages R. (Mediterranean).
25. Pa cornuarietis, Phil. R. (Mediterranean).
Details as to the localities from which specimens were collected have been recorded
at the beginning of the general description of each species. With regard to distribution
it will be noticed in the above list that several Serpulids common to the Red Sea and
Zanzibar have also been recorded by Prof. Willey from Ceylon. The two Mediterranean
species of Spirorbis, both collected at Suez, may very easily have been brought into the
Red Sea by way of the canal, a method of entry which seems quite certain in the case of
Hydroides norwegica, which was found growing on ships dry-docked in Suez on their
arrival from Alexandria. Hupomatus exaltatus also has an interesting distribution. This
species was founded by Marenzeller on one specimen from Japan in 1885, and was not
recorded again until twenty years later, when another single specimen was described
by Prof. Willey from the Gulf of Manaar, although previously to this Mr Crossland had
collected the three specimens recorded here from the neighbourhood of Zanzibar.
As usual in the Serpulidee, there is an enormous amount of variation; this is no doubt
accentuated by the prolonged action of preservatives, and emphasises the importance
of more systematic work being done on fresh material. which is sufficiently plentiful
to allow numerous specimens to be compared. In this paper which is of necessity almost
wholly systematic, I have endeavoured to tabulate at the beginning of the description of
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 71
each species the characteristics on which I have based my conclusions in identifying
specimens. Doubtless it will be found necessary to emend some of these characteristics
when more is known, for there can of course be no certainty with regard to the validity of
any one characteristic for systematic purposes.
Where not otherwise stated, the specimens are in Mr Crossland’s collections and the
quotations as to colour of living specimens, &c. are from his notes.
Genus Serpuna, Linné (81) 1767; Philippi (40) 1844.
Generic characteristics: 1. Collar setae bayonet-shaped, with spines at base of blade.
2. Opereulum funnel-shaped, with numerous radii ending in serrations on margin.
3. Uncini with only a few teeth.
1. Serpula vermicularis, Linné, 1767.
Tubus vermicularis, Ellis (17) 1755.
Serpula echinata, Gmelin (20) 1789; imbutiformis and infundibulum, Chiaje (10)
1825; contortriplicata, Savigny (47, p. 73) 1826; fascicularis, Lamarck (80) 1838 ;
aspera and pallida, Philippi (40) 1844; vermicularis and philippi, Mérch (387) 1863 ;
narconensis, Baird (2) 1864; octocostata, Quatrefages (42) 1865; crater, Claparéde
(12) 1870; patagonica, Grube (23) 1877; narconensis var. magellanica, M’Intosh
(32) 1885.
_ The above list includes the authors who originally gave the synonyms. For lists of
those who followed them consult Mérch, 1863 (87) and Saint-Joseph, 1894 (44).
Specific characteristics: 1. Collar setze with generally two large blunt processes at
base of blade. 2. Uncini generally have five teeth; there may be 4—-7. 3. Branchie
(20—32 pairs) long, with numerous pinne and bare filamentous extremities. 4. Serra-
tions on operculum vary very much in number ; there may be as many as 100. 5. Maxi-
mum length recorded is 50 mm. with 157 segments.
Previously known distribution: Atlantic, Mediterranean, English Channel, North Sea,
Magellan Strs., Marian Is., Orkney Is., Burdwood Bank, Table Bay.
Localities. Several specimens from Jambiani on the under sides of stones; a few
Specimens (2—3 cm. long) dredged in Wasin Channel, 7—10 fathoms; three small
specimens from the bottom of SS. Juba lying off Zanzibar; also a few specimens from the
Red Sea (Suakin?). Mr Crossland collected several other specimens from Porto Praya,
Cape Verde Islands.
The largest specimen of all, measuring slightly over 40 mm., came from Diamant
Island, Peros Banhos Atoll, Chagos Archipelago.
Nearly all possible variations in tube are to be met with, and there is also extreme
diversity in the arrangement of the red colour markings on operculum, branchiz, thoracic
membrane, &c. These variations have however already been described by Saint-Joseph
(44), Fauvel (18), &c., and it is unnecessary to mention them here. Some specimens are
undoubtedly in an immature state and show so few as 18 pairs of gills and 16 serrations
on the edge of the operculum—some of the uncini have only four teeth. In these and
72 PERCY SLADEN TRUST EXPEDITION
some other points there is agreement with the two specimens described by Gravier (21)
from the Gulf of Aden.
Only the one specimen from Diamant Island has as many as 50 serrations round the
margin of the opercular funnel, and this is the number given by Saint-Joseph (44) as the
minimum. The general number in these specimens is 30—40.
Genera Hyprormpes and EuPpomMATUus.
As so much confusion exists with regard to these genera, it has been necessary to
endeavour to draw up some scheme of classification. The definitions givén by the
authors who established them are so indefinite that it is almost impossible to decide into
which, if either, to put many of the recently described species with the characteristic
two-tiered opercula.
The operculum, in each case, consists of an outer more or. less funnel-shaped part as
in the genus Serpula with a varying number of radii ending generally in pointed teeth.
From the centre, or near the centre, of this arises a crown of spines—these may be all
alike (homoceros) or, as very frequently happens, one may be different from the rest
(heteroceros). Further, these spines very frequently have secondary processes. The
latter may occur internally and, in this case, especially if they are near the base of the
spine, be difficult to see from the outside, or they may be lateral and therefore quite
noticeable. Moérch points out that such characteristics of the spines are of sub-generic
value (87, p. 372), but his definitions are not sufficiently definite to distinguish some of
the more recently described specimens. Forms having lateral processes to their opercular
spines have been already referred to the genus Hydroides by Bush (7) and some other
authors. It seems best therefore to keep to this distinction and the genus Hydrordes will
then include H. norwegica, for which Gunnerus (25) established the genus (describing the
spines as “ bacillis teretiusculis, multispinosus”), and also many others. It would include
quite well the genus Hucarphus, established by Morch, which appears only to differ in
having the lateral processes terminal in position thus giving the condition “ bacillis
planis...apice arcuato..., utrinque cornuto,” described by Mérch (87, p. 372). In another
place (87, p. 378) Mérch draws attention to the fact that these lateral processes of the
spines may be sub-terminal (“extremitate utrinque aurita, interdum aculeo intermedio
terminali, unde uti cruciati”), and in such a position they are sometimes found in
H. heteroceros both in the specimens I examined from Zanzibar and the Red Sea and
in Grube’s specimens (22), though the more normal position is about half-way down.
Thus either the species H. heteroceros and many others should be put in the genus
Eucarphus, or else Eucarphus should be included in the genus Hydroides in which
the secondary processes (hooks) on the spines may be terminal or not. The latter
course, being more consistent with the laws of priority, is the one taken here.
The genus Glossopsis also seems to be quite unnecessary. 7. mina, for which it was
established by Bush (7), falls quite naturally into place in the Table given below, being in
fact closely related to Hl. monoceros, Grav.
The genus Hupomatus, Phil. “bacillis simplicibus apice leviter arcuato, sorta,” then
PIXELL—POLYCHATA OF THE INDIAN OCEAN 73
includes forms with opercular spines on which hooks are never present laterally and may
be absent altogether (figs. 4 and 5).
Except with regard to the shape of the operculum these genera resemble the genus
Serpula very closely and are for this reason by some authors regarded as sub-genera only.
It must be remembered however that the operculum, which is not generally a satisfactory
organ on which alone to base distinctions, is here developed as a highly specialised organ
of offence as well as of defence. Owing to the way in which the spines can be separated
and then brought together by the action of muscles (figs. 2a and 6), the central cavity
forms a highly efficient cage in which no doubt small animals are held prisoners and killed
by coming in contact with the sharp central spines. Small Crustacea were frequently
found imprisoned in this way, and one specimen of Hydroides heteroceros was found
grasping with its spines a small Serpulid in its tube which had evidently just been torn
from its substratum.
The general characteristics of the genera Hydroides and Hupomatus then are :
1. Collar setze of bayonet shape with two conical processes at base of blade.
2. Uneini with a few coarse teeth—the most anterior one being larger than
the rest.
”)
3. Abdominal setze trumpet-shaped (‘‘cornet comprimé” Gravier, |“ Spateln
Marenzeller).
The two genera can easily be distinguished by their opercula, and upon the
apparently constant characteristics of these opercula the annexed scheme of classification
Is suggested,
[#. norwegica*, Gunn
more than one pair of lateral processes H, multispinosa, Marenz.
la . parvus, Treadwell
[ H., homoceros, n. sp.
not terminal lz bispinosa, Bush
H
Alspunes alike . erucigera, Morch
(homoceros) |
one pair of lateral
oe H. lunulifera, Clpd.
eae ae with jpuogesses Cully terminal Hl ee |
lateral processes (Eucarphus Morch) ER MAE IO ear
1. dirampha, Morch
benzont, Morch
. heteroceros, Grube
bifurcatus, 2. sp.
. monoceros, Gray.
minax, Grube
. albiceps, Ehrb.
one spine only without lateral processes
Spines of 2 kinds
(heteroceros) |
only one spine with lateral processes
Sass SSSSISSUSSESSILSS
NN
. uncinatust, Phil.
. elegantulus, Bush
. protulicola, Bened.
. spongicola, Bened.
. dipoma, Schm.
. exaltatus, Marenz.
. Spinosus, N. sp.
Hupomatus Spines all alike (homoceros)
opercular spines with-
out lateral processes
SO
SESS SSESSSS SSIS]
Spines not all alike (heteroceros)
X
* According to Saint-Joseph H. pectinata and H. tripanon are only varieties of H. norwegica [46, p. 247],
so also apparently are H. (Hupomatus) elegans, Haswell [26 and 27] and H. abbreviata, Kr [29].
+ The spines vary from 6—11. Bush [8, p. 498] separates Ehler’s specimens [15] with 11 spines as
#. floridanus ; this seems to be unnecessary and the forms #. gracilis and H. humilis, described by this author,
are possibly also #. wneinatus. EH. (Hydroides) dianthus, Verrill, seems to be no more than a variety.
SECOND SERIES—ZOOLOGY, VOL. XVI. 10
74 PERCY SLADEN TRUST EXPEDITION
2. Hydroides norwegica, Gunn (25) 1768; Mérch (87) 1863; Marenzeller (84) 1893;
Saint-Joseph (45) 1898.
Specific characteristics: the radii of the operculum form about 22 to 36 rounded
lobes on the edge of the funnel: in the centre of this is the crown of (14—20) generally
17 spines with sharp lateral processes.
Locality. Numerous specimens from the bottoms of SS. Thyra and Silda which were
dry-docked in Suez after coming through the Canal from Alexandria. This species,
common in the Mediterranean and English Channel, has not been recorded before from
the Red Sea and it was no doubt in this case brought from the Mediterranean attached to
the ships.
Tubes white and delicate with conspicuous lines of growth, more or less adherent
but distal end generally erect the whole tube being very much larger than contained
animals. The length of these is generally about 20 mm. of which the long branchiz and
thorax together make up about a half; the width varies from 1—2 mm.
The functional double funnel shaped operculum seems more often to be carried on the
right the small club-shaped one being on the left, but this is of course subject to reversal
after loss (Zeleny 51), and both fully developed on their very long pedicles is not an
uncommon occurrence. ‘The horny transparent spines round the inner funnel have sharp
lateral processes which appear to collect debris so that the whole top of the operculum is
frequently hidden by it. The lower funnel of the operculum is colourless and opaque,
just below it on the slender stem occurs a bright crimson spot.
The collar is entire and the very large thoracic membrane is colourless and trans-
parent. The branchize of which there are about 15 pairs are variously “coloured red,
violet, yellow with white transverse markings or may be colourless” ; there are numerous
opposite pinnee extending almost to their extremities.
A full description of this species is given by Saint Joseph (45, pp. 440—443).
3. Hydroides homoceros, n. sp. (Plate 8, fig. 1).
Specific characteristics: the opercular funnel has about 17 teeth with lateral
processes (fig. 1a) and the central crown consists of seven slender spines, each having
a pair of lateral hooks about half-way up and a median basal internal one.
Localities. Two specimens from Miladumadulu Atoll and one from Minikoi in
Prof. Stanley Gardiner’s Maldive Archipelago Collection ; also two from 10—15 fathoms
off Ras Osowarmembe and four from the bottom of SS. Juba off Zanzibar in Mr Crossland’s
collection.
Tube “slightly ribbed, not much bent, mouth simple.”
The specimens are from 10—20 mm. long without the branchial crown which is
3—4 mm. high, and composed of 12—17 pairs of gills with only short terminal filaments.
The inter-branchial membrane is also very short. The spines of the very beautiful
operculum (fig. 1@) are transparent and yellowish. So also are the teeth with their
lateral processes into which the margin of the lower funnel is produced. One pair of
these processes is terminal, the other about half-way down. The pedicle varies in height;
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 795
it is always, however, longer than the other branchiz. There is a small rudimentary
operculum on the side opposite the functional one.
The thorax has the usual seven segments, the collar fascicle being composed of about
a dozen of the typical sete (fig. 1b) with an equal number of smaller ones with narrow
blades. The collar itself is deep, the ventral lobe generally entire, though sometimes
notched and variously folded or crumpled. The thoracic membrane is well developed ; in
two or three of the specimens the posterior ventral free part is nearly as long as the
whole thorax.
The abdomen has about 80 segments ; its greatest width is 2 mm.
The uncini have the usual shape and five or six teeth ; the abdominal setze (fig. 1c)
are in groups of eight.
This species has apparently rather close affinities with H. bispinosa, Bush (8) from
Bermuda. ‘There are, however, no figures nor sizes given of this species and from the
description there seem to be several points of difference (8, p. 496).
4. Hydroides heteroceros, Grube (22) 1868 (Plate 8, fig. 2).
Serpula (Hydroides) uncinata, Gravier (21) 1908 ; non Philippi (40) 1844.
Eupomatus heteroceros, Willey (50) 1905.
Specific characteristics: the opercular funnel has about 30 well-marked serrations
and the central crown nearly always consists of seven long stiff spines; six have lateral
hooks; the seventh is generally larger and curved over the others and has no lateral
processes (figs. 2a and b).
Localities. Several specimens from wall of Suez Quay; one from Suakin Harbour
obtained by divers; and two from a depth of 10 fathoms off Zanzibar.
Tubes thick, flattened on side of attachment, often coiled, marked by faint longi-
tudinal lines, aperture circular.
Length variable ; the largest specimens were about 40 mm. long and 4 mm. wide, but
most are only three-fourths of this size.
“The body is generally of a dull yellowish colour; the branchize have dark crimson
bases but are light yellow distally. There is, however, a great deal of variation in colour,
the gills occasionally appearing purple and orange. The pedicle of the operculum is
banded with crimson and white alternately.” The functional operculum is not necessarily
on the right as given by Gravier. In two specimens it appeared on the left and is no
doubt subject to reversal after loss (Zeleny 51).
The rudimentary operculum is short with a terminal knob showing six or seven radial
constrictions.
The serrations round the outer opercular funnel have laterally elongated terminal
processes sometimes produced into sharp teeth at the sides. In two out of the seven
specimens examined there were only five spines with the lateral hooks in addition to the
large curved plain one, instead of the usual number six—there is also a great deal of
variation in the position of these lateral hooks. In some, they are about half-way down
(figs. 2a and b) as figured also by Gravier* (21, fig. 286) and described by Willey (50),
* There can be no doubt, I think, that Gravier’s figure and descriptions of 1. wncinata refer to this
Species; they differ considerably from the true Hupomatus wneinatus.
10—2
76 PERCY SLADEN TRUST EXPEDITION
but in others they are quite near the distal end as figured by Grube (22), ze. they,
are sub-terminal and similar to those described by Morch in his genus Hucarphus
(87, p. 378).
All the spines are provided with interior median basal spines. There is a large
thoracic membrane and collar, the latter being very ample and much folded ; the ventral
lobe seems generally to be entire. The branchize, of which there are 16—18 pairs, have
very long terminal filiform processes.
The setze are of the usual shape and resemble Gravier’s figures (15, text-figs. 463—
466), the uncini having 5—7 teeth. The abdomen consists of about 150 segments.
A thin calcareous tube with conspicuous growth lines containing a very young
animal of presumably the same species was found firmly grasped by the opercular spines
of the largest specimen and the scar on the tube looked very much as though it had been
torn away from the substratum on which it had been growing. The total length of this
young form is barely 3 mm. and it still retains its original functional operculum with
a single row of serrations on the left side—the shorter one which was going to develop
into the first adult functional one with the characteristic shape (Zeleny, 51, p. 934) being
on the right. It has the ordinary seven thoracic segments but only about 24 abdominal
ones; the latter have tori containing 12 uncini of the usual shape. Each fascicle of
ventral setz: consists of three trumpet-shaped ones and one very long fine capillary one.
The collar setze (fig. 2c) are remarkable in having a coarsely serrated fin-like process at
the base of the blade instead of the ordinary two processes. I do not know whether this
could be a stage in the development of such sete. There are five or six in each fascicle
and the same number of fine capillary ones.
5. Hydroides bifurcatus, n. sp. (Plate 8, fig. 3).
Specific characteristics: the opercular funnel has 25 sharp pointed teeth round its
edge and a central crown of seven curved spines; six of these are bifid at the distal end
giving two sharp outwardly directed lateral teeth; each has also a median inner hook
a short distance below these lateral ones. The seventh spine is much curved and has
only the inner basal upwardly directed median tooth which is also present on all the
others (fig. 3).
Locality. A single specimen from Minikoi, Prof. Stanley Gardiner’s Collection, no
tube. It measures 20 mm. without its branchial crown and its greatest width is 2 mm.
The gills, of which there are 20 pairs, are 3 mm. long and have large terminal filiform
processes. The opercular pedicle is a little longer than the branchiz so that the operculum
projects just above them. Owing to the condition of six of the spines this species might
have been referred to the genus Hucarphus though it does not resemble any other species
of this group so nearly as it does H. heteroceros.
The setee are as usual in the genus—the uncini have seven teeth in the thorax and
five in the abdomen, but are everywhere of the characteristic pattern.
6. Hydroides monoceros, Gravier (21) 1908.
Specific characteristics: opercular funnel asymmetrical and with about 16 serrations
PIXELL—POLYCHATA OF THE INDIAN OCEAN 7
having enlarged extremities ; central crown has six small spines and a very large one with
a lateral hook on either side of the strong curved terminal spine.
Localities. Three specimens about 15 mm. long from the bottom of SS. Juba off
Zanzibar; one much larger specimen (54 mm. long and 3 mm. maximum width) from
Suakin, Red Sea.
The tubes are thick and more or less curved, with longitudinal and transverse ridges
and entire margins to apertures. The functional operculum with its pedicle is 4—8 mm.
long. It is quite asymmetrical, the large spine arises from the dorsal side and often lies
almost flat over the rest of it. This seems to be the chief point of difference from
H, minax, Grube, which has a symmetrically placed central crown.
The large spine with its strong terminal and pair of lateral hooks varies much in
length sometimes being as long as 3 mm. and thus able to extend some distance beyond
the ventral boundary of the lower funnel when lying closely over it, a position in which it
is often found. The edge of the lower funnel is produced into 16 or 17 processes with
terminal enlargements, the funnel is asymmetrical the radii being much larger on the
ventral than on the dorsal side.
The rudimentary operculum on the opposite side of the body is club-shaped.
The branchize (15—-20 pairs) are of a brownish colour banded with white. There is
a short interbranchial membrane, and the numerous rather small pinnze extend nearly to
the end leaving only a very short terminal filament.
The setze and uncini agree with those figured by Gravier (21).
7. Eupomatus exaltatus, Marenz. (83) 1885; Willey (50) 1905 (Plate 8, fig. 4).
Specific characteristics: the inner funnel of the operculum is raised on a short
column and has eight or nine strong hook-like spines, without secondary processes except at
base ; the dorsal one is twice as large as the others and bends suddenly at a right angle
over the top of them (fig. 4).
Localities. Two specimens from Wasin Channel, 7—10 fathoms; one from the
bottom of SS. Juba, Zanzibar, and one (operculum only) from Dongonab Harbour, Red
Sea.
Tubes round and almost smooth, being only slightly marked with growth lines and
three low dorsal ridges. Large operculum on very long pedicle projecting some distance
beyond gills when these are expanded.
Body of a dull green colour, branchize 13—15 on each side, greyish violet colour with
two clear broken black lines on the outer side of each ; terminal filament without pinnze
“3 mm, only.
After preservation the slightly larger specimen from Wasin has a total length of
19mm., the operculum with its pedicle being 433mm. The abdomen consists of about
90 segments, the terminal ones provided with long fine sete.
The smaller specimen from Wasin has nine central spines to its operculum which
occurs on the right side; that is, it is exactly the same as the one described by
Marenzeller (83); the larger specimen, however, has its functional operculum on the left
side, and it only has eight central spines as in Willey’s (50) specimen. The outer funnel of
78 PERCY SLADEN TRUST EXPEDITION
the operculum has 24 teeth round its edge. The secondary operculum is in both cases
quite small and rudimentary.
The specimen from the bottom of SS. Juba is smaller than the other two and has
its functional operculum (with eight spines) on the right ; the smaller one on the left is
also almost fully developed and shows the same number of spines.
Another operculum with eight central spines was found with some material from
Dongonab, but no trace of the rest of the animal could be found.
The uncini have six or seven teeth, in this being intermediate between Willey’s and
Marenzeller’s specimens ; otherwise the uncini and setee agree very closely with those
figured by the latter author.
This species is interesting in having a wide distribution considering its apparent
rarity. The species was established by Marenzeller in 1884 on one specimen from the
east coast of Eno-sima Island (S. Japan). Willey (50) records another single specimen
without a tube from the Gulf of Manaar in 11 fathoms in 1905.
8. Hupomatus spinosus, n. sp. (Plate 8, fig. 5).
Specific characteristics: opercular lower funnel with about 32 long pointed teeth,
the upper or central one with 10 strong tall spines, each with distinct terminal hook
pointing outwards and a nearly terminal inner one pointing downwards and inwards and
also an inner median one near the base. The eleventh spine is much longer and curved
over these and has only a basal hook (fig. 5).
Locality. Five specimens from Suez.
The tubes are thick with longitudinal ridges and somewhat irregular growth lines—
one has sand grains adhering to it. The length of the thorax and abdomen combined
varies from 10—23mm. with a maximum width of 3mm.
The branchize, of which there are about 11 pairs, have rather short terminal filaments
and pinnze. The pedicle, which is generally rather longer than the gills, is 3mm. in one
of the larger specimens. The spines of the operculum are almost black and are much less
raised than in HL. exaltatus. There are also dark brown bands anterior to the tori, these
being especially marked in the thorax, giving a very distinctive appearance.
The collar is entire and not very high nor ample, the ventral lobe thus forming a
straight band. The collar fascicle consists of large bayonet sete together with some very
fine capillary ones. The thoracic uncini have seven teeth, the abdominal six; both these
and the setze have the usual shape. The abdomen consists of about 100 segments in the
large specimens.
Genus Pomatostecus (Schmarda).
Generic characteristics: 1. Several horny disc-like opercula united by a central
vertical column. Pedicle with broad lateral wings. 2. Collar sete bayonet-shaped
and covered with fine hair-like processes, generally with simple bladed forms also.
3. Abdominal setze sickle-shaped (setee of Salmacina). Ehlers (15, p. 299) considers
this point of special importance in distinguishing Pomatostegus from Spirobranchus.
PIXELL—POLYCHATA OF THE INDIAN OCEAN 79
4. Uncini with about nine teeth, the anterior one larger and hollowed out underneath
like a gouge (cf. Spirobranchus). 5. Uncinigerous tori of right and left side almost
meeting on the ventral side of thorax, leaving only a narrow straight depression along the
median line (Ehlers, 15, p. 297).
9. Pomatostegus stellatus (Abildgaard) (1) 1789.
Terebella stellata, Abildgaard (1) 1789.
Pomatostegus stellatus, Mérch (37) 1863; Benedict (8) 1886; Ehlers (15) 1887;
Gravier (21) 1908.
The distinguishing characteristics of the several species of this genus that have been
described do not seem to be at all clear. Bush (8, p. 499) states that P. stellatus has
a smaller operculum and more elongated body than P. brachysoma, Schmarda, of which
she gives a long description, mentioning 45 mm. as the length (but giving no width) of
her one specimen. As a matter of fact, the one specimen of P. stellatus described in
detail by Ehlers (15) is said to have a length of 35 mm. only, and to be composed of 140
segments. P. macrosoma, Schmarda (48), has been said to differ from both of these in
having (1) a divided ventral lobe to its collar and (2) a greater number of turns to the
branchial spirals; however, the latter is said by Ehlers to be an unsatisfactory specific
characteristic as is also frequently the special form of collar or thoracic membrane.
Therefore P. macrosoma is probably identical with P. stellatus and possibly P. brachysoma
also.
Locality. Five specimens from “ the side of a coral block,” Suakin, Red Sea.
These specimens vary in total length from 21°5—52 mm. and have 70—96 abdominal
segments. They agree as to colour with the description given by Gravier, but I have
only been able to find very few bladed sickles (setze of Apomatus) among the thoracic
setae, the majority being rather short, stiff, simple and bladed forms. The abdominal
setae have the rounded teeth described by this author, but have nearly lost their sickle-
hike curve, this is, no doubt, due to their lengthened immersion in preservatives (ef.
Saint-Joseph, 44, p. 334). The tubes are thick and irregular, overgrown with Polyzoa,
Spirorbis tubes, &c. The cavity is, however, smooth and cylindrical, measuring 83—4 mm.
in diameter. There is nothing else that need be added to Ehlers’ excellent description
(15, pp. 296—300) of one specimen without a tube from Hast Key.
Genus SPIROBRANCHUS, Blainville (6) 1817.
Generic characteristics: 1. Operculum with a calcareous plate generally bearing
a group of branched spines. Pedicle with broad lateral wings. 2. Collar sete as in
Pomatostegus. 3. Abdominal sete trumpet-shaped, the edges compressed and toothed
and produced at one place into a long fine point. 4. Uncini the same shape as in
Pomatostegus, but with rather more numerous teeth (9—25). 5. Uncinigerous tori
of the two sides widely separated ventrally in front and gradually approaching one
another towards the end of the thorax, thus leaving a triangular depression.
80 PERCY SLADEN TRUST EXPEDITION
10. Spirobranchus giganteus (Pallas) (Plate 8, fig. 6).
Serpula gigantea, Pallas (89) 1766.
Cymospira gigantea, Blainville (6) 1817; Schmarda (48) 1859; Quatrefages (42)
1865.
Spirobranchus giyanteus, Mirch (37) 1863; Benedict (3) 1886; Ehlers (15) 1887 ;
Gravier (21) 1908.
Specific characteristics : 1. Opercular plate with only two antler-like processes which
sometimes however branch close to their bases. 2. Abdomen about 11 times as long as
its greatest breadth and with numerous (200—300) segments.
After studying numerous specimens which vary a great deal among themselves in
unimportant details, these seem to be the only constant characteristics not mentioned
among the generic ones. Ehlers (15) gives these with two other characteristics as
distinguishing S. gigantea from S. incrassatus and S. tricornis, both of which have three
processes, differently branched on their opercula, and from the measurements given their
abdominal regions are only 4—7 times as long as broad. Ehler’s specimen of S. incrassatus
seems however to have been a very small one. He gives the condition of the ventral lobe
of the collar as another specific characteristic, pointing out that his specimens (? how
many) of S. gigantea and S. incrassatus have this lobe divided into two triangular lappets,
while in S. tricornis it is undivided and very elongated. At the same time he suggests
that the condition with two lappets may be caused by damage.
In the specimens that I have examined there seems to be no constancy at all with
regard to the collar. The ventral lobe is in several cases (fig. 6a) folded inwards in the
middle and gives a striking resemblance to two triangular lappets, but in only the one
large specimen from Hulule, Male, are there two distinct triangular lappets which show
no sign of having been possibly caused by the original lobe being torn. In a great many
the whole ventral lobe is much shrunken and folded back as mentioned by Ehlers, and
when unrolled no appearance of triangular lobes could be distinguished, though the collar
appeared notched in the centre.
The very numerous specimens of Pomatoceros triqueter from Plymouth that I have
examined also show great variety as to their collars, and I am inclined to think that these
are never of much use from a systematic point of view.
In spite of Ehlers’ statement to the contrary, I have not found in any of the
numerous specimens examined that all the collar setae were the simple blades which are
constantly present to some extent in the collar fascicle of Serpulids. In some specimens
however (fig. 6c) the characteristic ones approach that condition more nearly than in
others (fig. 6b) where the hairy projection is larger. The one figured by Benedict
(3, fig. 43) is very similar to the former. Gravier (21) does not mention the collar sete
of his Red Sea specimens.
Localities. Numerous specimens were obtained from the Zanzibar region in 10
fathoms. One specimen came from Khor Dongonab, and several others also came from
the Red Sea, having been obtained by divers in Suakin Harbour.
In Prof. Stanley Gardiner’s collection there is a large and beautiful specimen from
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 81
Hulule, Male, and two (one incomplete) from Mamaduwari, 8. Mahlos, Maldive Islands.
These are about double the size of the Zanzibar and Red Sea specimens: one contracted
specimen is nearly 100mm. long with maximum abdominal width of 9 mm., another
expanded specimen is 116mm. long and only 6—7 mm. wide.
The tubes in most cases are completely covered with coral; there is the usual blue
colour on the inside round the aperture, which is overhung by the characteristic pointed
process.
The branchize, forming a spiral of four or five turns on each side, are generally
“white or yellow at their bases, but of a brilliant crimson or other shade of red towards
their distal ends.” The massive pedicle of the operculum arises on the left side just dorsal
to the branchiz and has a “greenish colour.” The collar, also “brightly coloured,
wrapped over the edge of the tube when the worm was extended.”
In every case the ventral lobe of the collar is produced at the side interior to the
lateral lobe into a process which is sometimes more or less fimbriated, and suggests the
languet of Pomatoceros. I have not seen this condition described before for the genus
Spirobranchus, but it seems to be of constant occurrence. Considerable variation was
found among the abdominal sets; two of the commonest forms are shown in figs. 6,d
and e. There is nothing else, I think, to add to the detailed description given by Ehlers.
Most of the specimens agree rather with those of Gravier from the Red Sea in having the
processes on the operculum shorter than in Ehlers’ and Schmarda’s figures. The opercular
plate is in every case distinctly elongated dorso-ventrally and is broader ventrally.
11. Spirobranchus guganteus, var. turbinatus nov.
Locality. Hulule, Male Atoll.
This beautifully preserved specimen resembles the above as to set, operculum, &c.,
but has rather fewer branchize with longer free extremities and the ample collar
enormously developed on the right dorsal side and rolled up like a scroll. In size it
resembles the other specimens from the Maldive Archipelago.
12. Spirobranchus gardineri, n. sp. (Plate 8, fig. 7).
Specific characteristics: 1. Opercular plate nearly circular with one very long process
arising from near its centre—the process being trifid near its distal extremity only
(fig. 7a). 2. Thoracic uncini very large and a somewhat different shape from the
abdominal ones (figs. 7c, d, e).
Three specimens from Providence Reef (N. of Madagascar) collected by Professor
Stanley Gardiner.
The three specimens are almost exactly the same size, and in each the posterior part
of the abdomen is cut off. In only one case is the latter region present ; it belongs to a
specimen which gives the following measurements : Pedicle of operculum, proximal region
4mm. long and 2mm. broad—this is attached almost exactly in the middle of the dorsal
surface between the two sides of the branchial crown. The more flattened distal part of
the pedicle is 13 mm. long, and with its narrow wings measures 5 mm. in width.
The opercular plate narrows slightly towards its point of attachment to the pedicle,
ze. towards the dorsal side. It has a bright orange-coloured edge in spirit specimens ;
SECOND SERIES—ZOOLOGY, VOL. XVI. ul
82 PERCY SLADEN TRUST EXPEDITION
slightly nearer the dorsal than the ventral side, arises the process which is 10mm. high
and ends in the three branches which sometimes themselves give rise to short secondary
branches. In one specimen the operculum is overgrown with a branching Bryozoon, but
in the other two cases it is quite smooth.
The branchial crown is five-spiral and 15 mm. high.
The thorax to the front of the collar measures 9 mm. long and about 7 mm. wide, and
is composed of the usual seven segments. The abdomen, with nearly 200 segments, is
36mm. long and 6 mm. across its widest part. The ventral surface is very distinctly
segmented and has a deep feecal groove.
The collar is entire ventrally, but folded inwards in the centre so as to appear at first
sight to have a median fissure (fig. 7a). Two specimens thus appear to have somewhat
rounded ventral lobes, while the other has two triangular lobes very much like those of
Spirobranchus giganteus shown in fig. 6 a.
The collar setze are as usual in the genus (fig. 7 5).
There were about 12 of these and 12 plain ones with a fringe on one side instead
of a distinct blade.
The thoracic uncini are enormous (about 300, long) and have 26 teeth, the most
anterior one being long and hollowed out underneath like a gouge (figs. 7¢ andd). The
abdominal ones are much smaller, with only 14 teeth (fig. 7 e), in front of the anterior
gouged one is a small process. The abdominal setae are as usual (fig. 7 f) in the genus.
13. Spirobranchus semperi, Mérch (87) 1863; Willey (50) 1905.
Serpula quadricornis Grube (24) 1878.
Specific characteristics: 1. Operculum flat, with four distinct spiny horns. 2. Uneini
with 9—13 teeth. 38. Branchial crown forms only one turn of a spiral.
Localities. Zanzibar and Red Sea.
14. Spirobranchus senvperi, Morch, var. acroceros, Willey (50).
Specific characteristics : this only differs from the above in having a conical opercular
dise with the four horns, reduced in size, at the top.
Localities. Numerous specimens from Suakin Harbour and Agig Bay; also three
from Dongonab in the Red Sea and several from Zanzibar in Mr Crossland’s collections ;
one specimen without its operculum from Suvadiva, Maldive Group, in Prof. Stanley
Gardiner’s collection.
From the four first places both varieties are represented. The tubes are generally
rough with longitudinal ridges and much overgrown with Bryozoa, &c. One which
contained the var. a@croceros and is not much overgrown is triangular, with a distinct
notched median keel ending in a projection over the aperture. Collar very ample, entire
and much folded ventrally and with small processes internal to the dorso-lateral lobes.
The opercular plate resembles Grube’s figure (24 Taf. xv. fig. 6) very closely. The
two dorsal horns divide into two main tines quite close to their bases, whereas the two
ventral generally only give rise to short processes. These horns are often hidden by
Hydrozoa, debris, &c., and in one they are united and altogether hidden by a colony of
PIX ELL—POLYCHATA OF THE INDIAN OCEAN 83
caleareous polyzoa. The pedicle of the operculum is very wide and has fringed lateral
wings. Many of the specimens are larger than those described by Willey: one of the
largest has the abdomen 17 mm. long, containmg 84 segments of greatest width 4mm.
and the thorax 5mm. long and nearly the same in width. The branchiz (24 pairs) are
4mm. high when extended, and the operculum very slightly higher.
15. Spirobranchus semperi Mérch, var. aceros nov.
This only seems to differ in having a flat opercular disc without horns, but very
much overgrown with Hydrozoa—the yellow chitinous perisares forming branching
processes 1mm. or more high in some places. Amongst these were found some small
crustacea. The one specimen present has about the same dimensions as the specimen
of Spirobranchus semperi quoted above, but is slightly more elongated owing to its
having been preserved in its tube.
Locality. Amirante Islands, F.I., 29 fathoms.
This has been placed as a variety only because it seems possible for opercular spines
not to develop in some cases. In one specimen of Spirobranchus contiert described below
three-branched spines were present on the dorsal half of the plate, while the ventral
was quite free from them.
16. Sprrobranchus contier: (Gravier) (Plate 9, fig. 8).
Pomatoceropsis contiert, Gravier (21) 1908.
Specific characteristics: 1. Interbranchial membrane more or less fimbriated along
its free edge. 2, Operculum with five (or six) much branched processes; pedicle winged.
3. Uncini with nine or ten teeth only.
This species seems to resemble closely Mérch’s S. dendropoma (87), for which,
however, Benedict (8) describes processes on the outer part of the radioles themselves
at the level of the interbranchial membrane.
Localities. Numerous specimens “commensal with two species of coral from entrance
to dock, Suez,” others from sandbanks, Chaki-chaki Bay, Pemba; also from Zanzibar and
from Suakin Harbour, Red Sea, from divers.
Tubes covered with layer of coral to near mouth.
“ Branchiz yellow, green and white with long filiform tips, interbranchial membrane
extends about half-way up. Body dark chocolate red, velvety in appearance, the thoracic
membrane being dark green and yellowish. After being in alcohol some time a pink
colour dissolves out and the bodies then appear dark blue.”
The process inside the lateral lobe of the collar that Gravier refers to as the languet
is distinctly a small lateral projection of the ventral lobe of the collar. It varies a good
deal in shape and is very similar to that generally met with in the genus Spiro-
branchus, being less definite in shape as a rule than the typical languet in Pomatoceros
triqueter.
The variation in these specimens is very marked. In most the five or six branching
processes on the operculum are hidden by a more or less cone-shaped or spherical mass
of debris which seems to be held together partly by these processes and partly by the
11—2
84 PERCY SLADEN TRUST EXPEDITION
presence of numerous colourless scalariform filaments. I have not seen any specimens,
having an operculum with spines springing from the top of a truncated cone, as described
by Gravier from Météore, but in more than one case specimens of Spirobranchus semperi
and its variety acroceros have been collected and placed with this species.
The wings on the peduncle are fringed on their upper free ends or quite plain as
pointed out by Gravier (21) or there may be only a single filamentous process or the
whole wing may be so reduced as to appear at first sight to be absent. The operculum
is always carried on the left side and there is no secondary one. The interbranchial
membrane may have a single cirrus between two adjacent gills or a much divided
process.
Length of specimens varies from 15 to 40 mm. and the width from 2—3 mm.
The sete agree with those so beautifully figured by Gravier (21, text-figs. 482—487).
The uncini, while much resembling Gravier’s figure in some directions, show a distinctly
gouge-shaped anterior tooth (figs. 8@ and b); this is particularly clear in front view
(fig. 8c), therefore I have ventured to include Gravier’s species, which I cannot imagine
can be different from this, in the genus Spirobranchus.
17. Spirobranchus maldivensis, n. sp. (Plate 9, fig. 9).
Specific characteristics: 1. Operculum a thin calcareous plate, without processes,
supported by a tall pedicle with thin lateral wings (fig. 9a). 2. Collar setee with a short
wide finely striated fin-like process at the base of the narrow anterior blade (fig. 90).
3. Branchize about 32 pairs with numerous long pinne except at their distal ends, which
are bare and filamentous. 4. Thoracic uncini have about 15 teeth im addition to the
large gouge-shaped one (figs. 9d and e) and the abdominal 13. 5. Abdominal setze
narrow compressed trumpets with one side produced into a long process (fig. 9 /).
Characteristics 1 and 2 separate this species widely from any other in the genus, in
fact, 1b may at some later time be found advisable to establish a new genus for it. The
collar setze are somewhat similar to those of Omphalopoma, Lang, and some other genera
and the operculum is unusual in having no spines.
Locality. ‘Three specimens in their tubes from Mulaku, Maldive Archipelago, Prof.
Stanley Gardiner’s collection.
Three tubes are fused together longitudinally and a fourth is firmly fused with a
Gastropod (Yurritella?) shell. In the latter the small end of the shell at first sight
appears to be a large pointed process overhanging the mouth of the Serpulid tube. Hach
tube has one coarsely serrated ridge. Two of the specimens are 20 mm. long and 2 or
3 mm. in their widest part; the other is 30 mm. long but not more than 2 mm. wide.
The branchize are 7—8 mm. high and not spirally arranged. The collar is high,
folded inwards along the median ventral line and interior to the dorso-lateral lobes are
small folded lateral processes of the ventral lobe as in Spzrobranchus, &c. >
The fascicles of collar setze are rather long and slender containing about 10 of the
characteristic sete (9b) and about the same number of simple narrow blades (fig. 9c).
All the other thoracic sete are simple blades.
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 85
There are 6 uncinigerous tori on each side leaving between their ventral ends a
triangular space with its apex towards the posterior.
The thoracic uncini are 86 » long and have about 15 pointed teeth in addition to
the large anterior gouge-shaped one (figs. 9d and e).
POMATOLEIOS, n. g.
Generic characteristics: 1. Collar sete and eyespots absent. 2. Uncini with
fairly numerous teeth, the most anterior being larger and gouged underneath (fig. 10d).
3. Abdominal setze trumpet-shaped with one side produced into a long spine (fig. 10¢).
4. Operculum flat with winged pedicle. 5. Tube with a flap over the entrance.
18. Pomatoleios crosslandi, n. sp. (Plate 9, fig. 10).
Specific characteristics: 1. All thoracie sete simple striated blades (fig. 100).
2. Uncini with 10 or 11 teeth in both thorax and abdomen. 3. Branchize with very
high inter-branchial membrane and long bare terminal filaments (fig. 10@).
Localities. Ras Shangani and Chwaka. Numerous specimens but no tube present.
Mr Crossland, however, gives the following notes. ‘Rough coiled tube, blue coloured
especially inside and having a flap over the entrance: occurring many together not far
below high water mark, e.g. on old boiler at Ras Shangani and at Chwaka near high tide
mark down to half tide, but easiest to collect at high neap tide.”
Mr Crossland also draws attention to the remarkable vitality of these specimens
which he had some difficulty in narcotising. The tube appears to be quite exceptional,
but unfortunately there are none present in the collection: the total absence of collar
sete is also rare. Though numerous specimens, of total lengths varying from 4 to
_ 14mms., have been carefully searched no sign of these sete has been found. Neither
are there apparently any eyespots as in the genus Placostegus ; the specimens also differ
in other obvious ways from this genus. Should still younger specimens than those that I
have been able to examine be found to have collar sete, then this genus may be able
to be included in the genus Pomatoceros. In Pomatoceros triqueter the collar sete are
sometimes much reduced in the adult (Soulier 49), but according to Saint-Joseph (44)
they are always present.
The six fascicles of thoracic sete present are all alike, being composed of simple
striated blades (fig. 106).
The collar varies a great deal; the ventral lobes seem generally to be high and
entire but folded as in fig. 10a, and they give rise to a small dorsal lobe or languet
which lies just inside the dorso-lateral lobe of the collar.
The branchize, 13—16 pairs, are remarkable for the height of their interbranchial
membrane (fig. 10a); this joins them together for about half their height and nearly half
of the free part is bare of pinnze forming a long tapering filament.
The opercular pedicle is short and has thick lateral wings which end distally in
straight edges close under the operculum. It arises almost in the middle line of the
dorsal surface slightly, if anything, to the left and is attached towards the dorsal side of
86 PERCY SLADEN TRUST EXPEDITION
the flat opercular plate. The latter is circular, rather thick and quite soft ; this condition
may I think be due to the fixative though I am not sure whether the corrosive sublimate
used contained any acid with it or not. At any rate, the appearance is much more that
of a softened calcareous plate than of a chitinous one.
Genus VERMILIOPSIS, Saint-Joseph (44) 1894.
Generic characteristics: 1. Collar setz simple blades. 2. Uncini with fairly
numerous teeth, the most anterior are larger and blunter than the rest. 3. Abdominal
setee geniculate. 4. Some thoracic sete are bladed sickles (setee of Apomatus), thus
differing from genus Vermilia with ordinary bladed setz only. 5. Operculum with a
horny somewhat cylindrical or conical cap.
19. Vermaliopsis pygidialis (Willey) (Plate 9, fig. 11).
Vermilia pygidialis, Willey (50) 1905.
Specific characteristics: 1. Branchize with ocelli and elongated often much swollen
ends free from pinne. 2. Operculum with a conical (sometimes truncated) chitinous
cap (fig. 116). 3. Uncini with 13 or 14 teeth. 4. Terminal dorsal gland generally
present.
Localities. One large specimen from Ras Osowarmembe, Zanzibar, in 10—15
fathoms; others from Suez and Suakin in Mr Crossland’s collections. One specimen
from Suvadiva, Maldive Group and a portion from Funafuti in Prof. Stanley Gardiner’s
collection. Tubes ridged longitudinally and attached to the under sides of stones, &c.
Two specimens are about 25 mm. long but the others only 15 mm. or less without
their gills, z.e. about the size of Willey’s specimen—one small specimen is regenerating its
collar, branchize and anterior thoracic segments. The general colour during life is “ bright
pink or red, the gills being banded with white.” There are from 10—15 pairs of bran.
chize ; their swollen ends appear to be glandular, though they are not so much developed
as in Salmacina dystert. The pyriform ocelli are arranged in groups along the outer side
of the rachises.
The thoracic membrane is short and between the ventral borders of the thoracic
uncinigerous tori is a depressed shield-shaped space with its broad end forwards. The
terminal dorsal gland in some preserved specimens appears brownish and chitinous, in
others whitish. po
The operculum is often covered with a reddish or white calcareous incrustation
(fig. 11a). When this was removed the translucent, light brown, horny cap was
revealed. This is in most somewhat truncated (fig. 11b) and has several deep brown
rings round it. In the specimens collected in August at Suez there is no incrustation
and the horny cap is quite conical and dark in colour. The incrustation is specially
noticed in specimens collected in November and February. The specimen from Suvadiva
Atoll is without any incrustation.
This species differs from Vermiliopsis glandigerius, Grav., in its branchize and oper-
culum; its pedicle too is not ribbon-like, though it is certainly much wrinkled and
swollen below the vertically placed horny cap.
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 87
The Vermiliopsis infundibulum, Lang (fully described by Saint-Joseph, 46), has a
terminal tooth to its operculum, a greater number (15—16) of teeth to its uncini and
apparently no swollen ends to its branchiz. Both Gravier and Saint-Joseph record that
in these species the operculum occurs on the left—they seem to have plenty of specimens
for examination, so presumably the position is constant in this genus. In all the
specimens of Vermiliopsis pygidialis that I have been able to examine it occurs on the
right, as also in Willey’s specimen from South-West Cheval Paar, Ceylon.
Genus FinocrRaNa (Oken) (88) 1815.
Characteristics were originally given by Berkeley (5) to correspond with his
previously described Filograna implexa (4).
Saint-Joseph (44, p. 335) takes the genus Filograna to include the sub-genus
Filograna s. str. with two opercula as in F. implexa, and also the sub-genus Salmacina
including S. imerustans, Claparéde (12) and S. edificatrix, also S. (Filograna) hualeyr,
Ehlers (51), as well as S. dyster1, Huxley (28).
According to this arrangement the characteristics of the genus would be: 1. Tubes
very slender, filiform, colonial. 2. Branchize 8. 3. Thorax with 7—9 segments.
4. Collar setee with a large fin-like expansion at base of blade. 5. Other thoracic
setze sickle-shaped (setze of Salmacina) or ordinary bladed forms. 6. Abdominal setze
more or less geniculate and serrated. 7. Hermaphrodite.
Sub-genus Satmacina, Claparéde.
Generic characteristics: 1. No operculum. 2. The ends of the eight branchiz may
or may not have spathulate enlargements due to the presence of large granular cells.
Prof. McIntosh, in a paper communicated to the British Association in Sept. 1912,
still upholds the view that Filograna implexa and Salmacina dysteri are identical,
stating that both are to be found in the same colony in some places.
20. Salmacina dyster: (Huxley).
Protula dystert, Huxley (28) 1855; Claparede (11) 1863.
Filopora filograna, Dalyell (18) 1853.
Salmacina dysteri, Giard (19) 1876; Saint-Joseph (44) 1894; Gravier (21) 1908;
Pixell (41a) 1913.
Specific characteristics: 1. Branchizw 8 with spathulate enlargements containing
granular masses at their ends. Similar granules occur at the end of the pinne and just
in front of their bases along the gill rachises. 2. Spermatozoa developed in segments
anterior to those producing ova.
Localities. Large quantities from the bottom of SS. Juba and from other places off
Zanzibar ; also from Wasin. Masses were also growing on a small buoy in the entrance
to Suez Canal and on “SS. Fayoun, a pilgrim ship which had been stationary for about
8 months; one colony covered a square foot, was 8 or 9 inches high, and was apparently
rapidly growing.”
88 PERCY SLADEN TRUST EXPEDITION
Some specimens from 8S. Vincent, Cape Verde Islands, are also present but in a poor
state of preservation.
This species has already been fully described by Huxley (28) 1855 and Saint-Joseph
(44) 1894.
It has been recorded from the Gulf of Aden by Gravier (21), who obtained a
colony to the north of Ambouli in the Gulf of Tadjourah. He, however, failed to find
any in process of budding, whereas all four stages recorded by Saint-Joseph (44, p. 343)
are present in the material collected near Zanzibar in the spring and summer of 1901
and 1902.
The number of thoracic segments was very variable between 7 and 9.
Genus Protuta, Risso (43) 1826.
Generic characteristics: 1. No operculum. 2. Collar setz simple tapered blades.
3. Terminal dorsal gland present.
21. Protula tubularia, Montague (36) 1803.
For synonyms see Saint-Joseph (44) and Fauvel (18).
Specific characteristics after Fauvel (18, p. 693): 1. Ventral lobe of collar entire.
2. Abdominal setze sickle-shaped.
Locality. Three specimens from Porto Praya, Cape Verde Islands.
The largest specimen is 30 mm. long without its branchial crown, which is missing.
I could only find simple bladed sete in the thorax so that these specimens seem to
resemble those from Courseulles described by Fauvel (18, p. 694) in this particular.
The bodies are long, narrow and almost cylindrical with the usual wide folded
thoracic membrane. Saint-Joseph (44) has given a detailed account of this species.
22. Protula intestinum, Lam.
Specific characteristics after Fauvel (18, p. 694): 1. Ventral lobe of collar notched
in median line. 2. Abdominal sete geniculate.
Localities. One specimen dredged off Zanzibar, another in 10 fathoms off Wasin.
A portion of a thin, smooth tube 5 mm. in diam. from same region.
The specimens are small forms less than 3 cm. with distinctly notched colourless
collars. During life the numerous gills were also “colourless with bands of orange spots
and the body was slightly tinged with scarlet.” The abdomen was wide and somewhat
fusiform.
Prof. Stanley Gardiner’s collection contained one large orange specimen from
Cargados Carajos, 30 fathoms; this, without its branchial crown, was 140 mm. long
and the greatest width of the abdomen was 21 mm.
Genus Srrrorsis, Daudin (14) 1800.
Characteristics of genera and sub-genera have already been given (41).
PIXELL—POLYCH ATA OF THE INDIAN OCEAN 89
Sub-genus Laosprra, Caullery and Mesnil (9), emended Pixell (41).
23. Spirorbis papillatus, n. sp. (Plate 9, fig. 12).
Specific characteristics: 1. Large collar setze with coarsely serrated blades and fin-
like expansions (fig. 12¢). 2. Operculum with brood pouch, terminal cap flattened and
surrounded with a wide phalange covered with papillee (figs. 12a and b). 8. Some set
of second fascicle ordinary sickles (fig. 12d). 4. Some setze of third fascicle bladed
sickles (fig. 12e).
Locality. Chuaka, Mr Crossland’s Zanzibar collection. Numerous specimens
“closely crowded together on the base of Sertularia colonies which are abundant in
the sand not far below high tide level.”
Tubes smooth, their bases overgrown with a red calcareous encrusting alga. Other
specimens from a Pinna shell in Suakin Harbour, Red Sea, encrusted with a polyzoon.
In both the terminal part of the tube ascends, being in some nearly erect; the
aperture is circular and entire, about 1°5 mm. in diameter.
Animals bright scarlet 2—3 mm. long; deep entire collar and wide thoracic
membrane.
The operculum varies a good deal in appearance. Fig. 12a shows a form containing
only a few ova. The edge of the phalange is provided with long papille. In fig. 12
these are much shorter and more irregular and the flexible side of the operculum is much
distended with well advanced embryos which already have three or four setigerous seg-
ments. In other specimens the papille are much smaller and sometimes only appear
to be scattered on the upper flattened side of the phalange.
Branchiz generally 11, sometimes only 10. They have numerous long thin pinnee
starting some distance below the distal end, which tapers to a long thin filament. The
operculum displacing the second gill on the left reaches about the same height as the
functional ones.
In the long setigerous region of the abdomen, the stomach is very evident in nearly
all specimens owing to its dark granular walls. This is followed by eight to twenty
setigerous segments, the geniculate setee having long deeply serrate blades.
24. Spirorbis levis, Quatrefages (42) 1865; Claparéde (12a) 1868; Saint-Joseph
(44) 1894.
Specific characteristics: 1. Collar sete simple blades with coarse striations.
2. Other thoracic seteze simple blades. 3. Ova incubated in operculum.
Locality. A few specimens on a tube of a Sabellid from the North Harbour, Suez.
Tube with two or three regular coils and a circular aperture: the surface is generally
marked with three longitudinal ridges and fine scalariform markings.
The specimens obtained complete from their tubes were young. Only two specimens
with ova in their opercula were found.
25. Sprrorbis cornuariatis, Phil. (40) 1844; Mérch (37) 1863; Marion and Bo-
bretzki (85) 1875; Caullery and Mesnil (9) 1897.
SECOND SERIES—ZOOLOGY, VOL. XVI. ‘ 12
90 PERCY SLADEN TRUST EXPEDITION
Specific characteristics: 1. Large collar sete with coarsely serrated blades and
fin-like expansions. 2. Ova incubated in tube. 3. Talon of operculum with large,
more or less hook-like projections.
Locality. Several specimens attached to the tubes of Spirobranchus contieri from
entrance to dock, Suez.
Some of the setee of the third thoracic segment are, as Caullery and Mesnil (9) point
out, bladed sickles, not simple capillary ones as stated by Marion and Bobretzki (86).
There are 9—12 segments in abdomen, each torus consisting of about 20 uncini.
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PERCY SLADEN TRUST EXPEDITION
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EXPLANATION OF PLATES 8 AND 9.
PLATE 8.
. 1. Hydroides homoceros, u. sp. 1a, Operculum, x 24; 16, collar seta, x 300; 1c, abdominal
seta, x 375.
. 2. Hydroides heteroceros, Grube. 2a, Operculum with spines open, x17; 2b, operculum with
spines closed, x 17; 2c, collar seta of a young specimen, ? same species, x 300.
. 3. Hydroides bifurcatus, n. sp., operculum, x 24.
. 4. Hupomatus exaltatus, Marenz., operculum, x 24.
. 5. Hupomatus spinosus, n. sp., operculum, x 24.
. 6. Spirobranchus giganteus, Pallas. 6a, Ventral lobe of collar, x19; 6b and c, two extreme
forms of collar sete, x 300; 6d and e, two forms of abdominal set, x 300.
. 7. Spirobranchus gurdineri, n. sp. 7a, Anterior end photographed from ventral side, showing
collar, operculum, &., x 3; 76, collar seta, x 300; 8c, thoracic uncinus, x 300; Sd, front view of
anterior end of same to show the gouge-shaped tooth, x 400; 8e, abdominal uncinus, x 300;
8 f, abdominal seta.
PLATE 9.
8. Spirobranchus contiert, Gravier. 8a, Thoracic uncinus, side view, x 220; 8b and ¢, the same,
x 400—B, side view, c, front view.
g. 9. Spirobranchus maldivensis, n. sp. 9a, Operculum and pedicle, ventral view, x 8°5; 9b, charac-
teristic collar seta, x 300; 9c, simple collar seta, x 300; 9d ande, lateral and front views
respectively of anterior end of thoracic uncinus, x 300; 97, abdominal seta, x 300.
10. Pomatoleios crosslandi, n. sp. 10a, Anterior half of animal, x 17; 10, thoracic seta, x 220;
10c, abdominal seta, x 220; 10d, anterior end of thoracic uncinus, front view of, x 220.
ll. Vermiliopsis pygidialis (Willey). Operculum, x17; lla, with calcareous incrustation ;
116, with incrustation removed.
g. 12. Spirorbis papillatus, n. sp. 12a, Operculum with large ova, x 48; 126, operculum with
embryos, side view, x 48; 12¢, collar seta, x 375; 12d and e, sete from second and third thoracic
fascicles respectively, x 375.
Trans. Linn. Soc. SER.2.ZooL Von. XVI PL. 8.
PERCY SLADEN TRUST EXPEDITION.
(Giana)
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Percy SLADEN TRUST EXPEDITION.
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No. VII.—COLEOPTERA: NITIDULIDA, HETEROCERID.
Par A. GROUVELLE.
(CoMMUNIQUE PAR LE PROFESSEUR J. STANLEY GARDINER, M.A., F.R.S., F.L.S.)
(Text-ric. 1—6.)
Lu le 1 mai 1913.
Cerre étude du matérial ramassé par la Percy Sladen Trust Expedition note 27
espéces de Nitidulidee réparties entre 8 genres: Pria, Brachypeplus, Cilleus, Carpophilus,
Haptoncus, Stelidota, Lasvodactylus. et Cybocephalus. Sur ces 27 espéces 14 sont décrites
pour la premiére fois. Une seule espéce d’Heteroceridz est notée (voir p. 116).
Pria. Les insectes de ce genre se rencontrent dans presque tout l’ancien continent.
Les deux espéces récoltées aux Seychelles sont nouvelles; l'une P. nitidior est voisine des
espéces décrites, la deuxiéme P. scott: se rapproche jusqu’d un certain point des Mystrops.
Il est probable qu'elle est localisée dans la région des Seychelles, tandis qu’on peut penser
retrouver P. nitidior dans d'autres parties de la région Malgache.
Brachypeplus. Les deux espéces rapportées 4 ce genre semblaient localisées jusqu’’
ce jour; l'une 4 Ceylan et dans I’Inde méridionale, l'autre 4 Ceylan et dans |’ Archipel
Malais. Ces deux espéces n’ont rien d’intéressant au point de vue de la caractéristique
d'une faune.
Cilleus. Les Cillews sont répandus dans toutes les régions tropicales et sub-
tropicales; sur les 5 espéces recueillies aux Seychelles quatre n'ont pas encore été
décrites, la cinquiéme est identique & une espéce de Madagascar, C. megacephalus. Trois
des espéces nouvelles se rapprochent des formes connues de Madagascar et la quatriéme
dune espéce décrite de Ceylan. Il faut toutefois constater que les Cilleus sont des
insectes souvent rares et que de nouvelles découvertes peuvent modifier les idées qui
peuvent exister sous la distribution géographique de leurs diverses formes.
Carpophilus. Ce genre comprend un certain nombre d’espéces cosmopolites ou
& aire de dispersion considérable. Sur les 6 espéces signalées dans ce mémoire, deux
(C. dimidiatus Fabr. et C. hemipterus Linn.) sont entiérement cosmopolites, une
(C. humeralis Fabr.) est répandue dans toutes les régions tropicales et subtropicales
de l’ancien continent, une (C. angustatus Murr.) semble spéciale & la faune Malgache
et les deux derniéres n’ont pas encore été décrites. Elles semblent rares; peut-étre seront
elles retrouvées dans d’autres tles de la région Malgache.
94 PERCY SLADEN TRUST EXPEDITION
Haptoncus.
especes 4 aire de dispersion considérable.
(HZ. luteolus Er.) est un cosmopolite des régions tropicales et subtropicales, deux autres
(ZZ. ocularis Fairm. et H. minutus Reitt.) se rencontrent dans presque toutes les régions
tropicales et subtropicales de l’ancien continent : la premiére décrite de la Polynésie, puis
de l’'Inde, ete. ete., a été retrouvée sur la céte occidentale de | Afrique ; la deuxiéme décrite
de Java a été retrouvée dans toute la Malaisie, la Cochinchine, |’Inde, 4 Madagascar et sur
Comme le genre Carpophilus, le genre Haptoncus comprend .des
L’une des espéces recueillies par l’expédition
la cote orientale du continent A fricain.
Seychelles.
Aucun individu répondant al! Haptoncus sobrinus (Haptoncura) Grouv. (1894, Ann. :
Soc. Ent. Fr. p. 15) n’a été trouvé parmi les nombreux spécimens récoltés par l’expédition.
Stelidota.
Genre également cosmopolite.
moment, particuliére aux Seychelles.
Distribution géographique des espeéces.
x
Une seule espéce parait spéciale a la région des
L’unique espéce semble, au moins pour le
Groupe Groupe
Farquhar | Aldabra (incl.|
aoe, Suoie (Camps | (Conelb fs AMEE Provenance des espéces déja décrites
Chagos | Amirantes| Pierre et Astove, P ya
Seychelles 4 2
Provi- Assomption
dence) jet Cosmelédo)
‘
Brachypeplus notatus
Brachypeplus zequalis
Cilleeus opaculus
Cilleeus micros
Cilleus subplagiatus
Cilleeus confusus
Cilleeus megacephalus
Carpophilus humeralis......
var. foveicollis
| Ceylan
Ceylan, Archip. Malais
Madagascar
Ancien Continent, Rég. trop. et subtrop. |
Asie, Rég. trop. et subtrop. i
Carpophilus hemipterus ... — — | Cosmopolite
Carpophilus fusciceps ......
Carpophilus scotti ......... |
Carpophilus dimidiatus Cosmopolite
var. contingems...............
var. ochropterus
Carpophilus angustatus
Haptonceus luteolus ....... j
Haptoncus ocularis
Haptoncus minutus
Haptoncus sobrinus
Haptoncus obscurus.........
Stelidota explanata
Lasiodactylus pictus
Lasiodactylus breviusculus
Cybocephalus brevis
Cybocephalus minimus
Cybocephalus minutus......
Cybocephalus
Cybocephalus
Heterocerus vulpes
subpallipes
tantillus......
Ancien Continent, Rég. trop. et subtrop.
Afrique aust., Madagascar
Madagascar
Cosmopolite, Rég. trop. et subtrop. |
Ancien Continent, Rég. trop. et subtrop. |_
Ancien Continent, Rég. trop. et subtrop.
Iles Mascareignes
Asie mér., Arch. Malais, Madagascar
Madagascar
Madagascar
GROUVELLE—COLEOPTERA: NITIDULIDA, HETEROCERID At 95
Lasiodactylus. Ce genre est représenté dans toutes les régions tropicales et sub-
tropicales, mais les formes spéciales 4 l’ancien continent s’écartent sensiblement de celles de
Amérique. Les espéces récoltées par l’expédition appartiennent au L. pictus Macl.
répandu depuis l’Australie et le Japon jusqu’a la cdte orientale de Afrique; et & une
forme (L. breviusculus Fairm.) considérée jusqu’a ce jour comme particuliére 4 Madagascar.
Cybocephalus. Ce genre est également représenté dans toutes les parties du
monde. Les cinq especes décrites dans ce mémoire paraissent toutes spéciales 4 la
faune des Seychelles.
Meligethini.
Pria Stephens, 1830, Ill. Brit. Ent. im. p. 30, 49.
il, (72@ S@0, i, Fo (Asie, il)
Ovata, convexa, nitida, tenuissime fulvo-testaceo pubescens, nigra; antennis clava
excepta, pedibus, capite prothoraceque rufo-testaceis. Caput transversum, fronte con-
Fig. 1. Pria scotti, n. sp., ¢, x 12.
vexum, dense punctulatum ; margine antico inter oculos subexplanato, antice subtruncato;
oculis prominulis. Prothorax transversus, antice angustus, lateribus arcuatus, juxta
basin subparallelus, antice truncatus, basi modice arcuatim emarginatus, dense punctu-
latus ; marginibus lateralibus stricte marginatis, ad posticos angulos sat late subexplanatis.
Scutellum modice transversum, subtriangulare, quam prothorax validius et paulo parcius
punctulatum. Hlytra apicem versus attenuata, 1 et 4 longiora quam simul latiora, apud
marem separatim rotundata, apud feminam separatim acuminata, dense punctulata.
Clava antennarum maris elongata, feminae brevior, in utroque sexu paulatim in-
erassata. Long. 1,5—1,8 mill.
Ovale, convexe, brillant, couvert d'une pubescence roux testacé, trés fine, médiocre-
ment visible, roux testacé avec la massue des antennes et les élytres noirs. Antennes
tres allongées chez les males bien développés, plus courtes chez la femelle; 1° article
x
droit, 2™° a 8m allongés chez le male, 6™° & 8™* suballongés, progressivement plus
courts chez la femelle, 9™° 4 11™® formant une massue progressive, plus longue chez
le male. Téte transversale ; front assez convexe, densément pointillé; marge antérieure
subexplanée et subtronquée en avant des naissances des antennes; bords latéraux
entre le prothorax et la base des antennes, convergents; yeux échancrant légérement
les cdtés de la téte, présentant une saillie transversale, presqu’égale & la longueur
96 PERCY SLADEN TRUST EXPEDITION
de l’orbite. Prothorax rétréci en avant, arrondi sur les cétés, subparalléle contre la
base, assez nettement plus de deux fois plus large 4 la base que long, densément
pointillé, mais un peu plus faiblement que la téte; bord antérieur tronqué, angles
antérieurs vus de dessus obtus, un peu émoussés, vus de face plus arrondis; cdtés bordés
par une marge tres étroite, s’élargissant sur les angles postérieurs ; ceux-ci vus de dessus,
un peu aigus, émoussés, saillants en arriére, vus de face droits, largement émoussés; base
largement et faiblement échancrée. Kcusson en forme de triangle curviligne, plus large
que long, un peu plus grand a la base que le cinquiéme de la longueur de la base
du prothorax, a peine plus fortement ponctué que la téte. Elytres, & la base, de la largeur
du prothorax, arrondis aux épaules, atténués vers le sommet, faiblement arqués sur les
cdtés, sauf vers lextrémité, contimuant presque la courbure des cétés du prothorax,
arrondis séparément a l’extrémité chez le male, acuminés séparément et trés nettement
chez la femelle, environ une fois et un tiers plus longs que larges ensemble & la base,
densément et un peu plus fortement pointillés que le prothorax ; marges latérales assez
fortement infléchies dans la moitié basilaire, trés finement rebordées ; stries suturales
marquées sur la moitié apicale de l’élytre. Pattes larges; tibias postérieurs subparalléles,
environ quatre fois plus longs que larges; bord latéral des tibias antérieurs 4 peine
visiblement crénelé. Lignes fémorales des hanches intermédiaires rebordant la hanche
et sinfléchissant 4 Pextrémité trés prés de l’épisterne. Premier segment de l’abdomen
subégal au métasternum. Segment complémentaire de l’abdomen du male terminé en
saillie aigue; dernier segment présentant au sommet une légére élévation longitudinale.
Loc. “Seychelles. Mahé: country above Port Glaud, 500—1000 feet, XI. 1908,
19; Cascade Estate, ca. 1000 feet, 1909, 1 gf, 2 9.”
2. Pria mitidior, n. sp.
Oblonga, subparallela, convexa, nitidissima, vix perspicue aeneo-micans, tenuissime
flavo-cinereo pubescens, nigro-picea; capite, antennis clava excepta pedibusque subpiceo-
rufis; elytris lata plaga communi, latera attingente, antice transversim, postice angulose
terminata et stricto apicali margine subpiceo-ochraceis notatis. Caput transversum, fronte
convexum, dense punctulatum; margine antico subexplanato, antice subtruncato ; oculis
modice prominulis. Prothorax transversus, antice angustus, lateribus arcuatus, juxta
basin parallelus, antice truncatus, ad extremitates basis retrorsum inflexus, dense punctu-
latus; lateralibus marginibus stricte marginatis. Scutellum subtriangulare, modice
transversum, dense punctulatum. Elytra lateribus subparallela, juxta apicem attenuata,
apice separatim late rotundata, paulo longiora quam simul latiora, dense punctulata.
Clava antennarum apud feminam elongata, subpiriformis. Long. 1,5 mill.
Oblong, moins de deux fois plus long que large, convexe, trés brillant, présentant un
trés faible reflet bronzé-cuivreux, couvert d’une pubescence flave cendré, trés fine, médio-
crement visible; téte, antennes sauf la massue et pattes roux de poix clair; prothorax
brun de poix ; élytres jaundtres, trés légérement teintés de nuance de poix, marqués sur
la région scutellaire d’une large tache triangulaire de la couleur du prothorax et vers le
sommet d’une bande transversale, de la méme couleur; dessous du corps roux de poix,
plus foncé sur le métasternum. Antennes modérément allongées chez la femelle; article
GROU VELLE—COLEOPTERA: NITIDULIDA, HETEROCERID A 97
6me & 8™° suballongés, progressivement plus courts, 9™° & 11™° formant une massue pro-
gressive, environ une fois et deux tiers plus longue que large. Téte transversale, front
assez convexe, densément pointillé; marge antérieure subexplanée, subtronquée; bords
latéraux, entre le prothorax et la base des antennes, convergents, fortement échancrés par
les yeux; ceux-ci présentant une saillie transversale, subégale & la moitié de lorbite.
Prothorax rétréci en avant, arrondi sur les cotés, paralléle contre la base, trés nettement
plus de deux fois plus large a la base que long, densément et & peine plus faiblement
pointillé que le front; bord antérieur tronqué ; angles antérieurs vus de dessus arrondis,
vus de face presque droits; cdtés bordés par une marge tres étroite, faiblement élargie
vers la base; angles postérieurs vus de dessus aigus, saillants en arriére, vus de face
presque droits, émoussés; base largement tronquée entre les saillies des angles postérieures.
Eeusson en forme de triangle curviligne, environ deux fois plus large que long, égale & la
base, environ au quart de la longueur de la base du prothorax, densément et & peine plus
fortement ponctué que celui-ci. Elytres a la base, de la largeur du prothorax, arrondis
aux épaules, faiblement arqués sur les cotés, atténués vers le sommet dans la partie
apicale, largement et séparément arrondis 4 lextrémité, 4 peine plus longs que larges
ensemble dans leur plus grande largeur, densément pointillés; cétés continuant presque
la courbure des cdtés du prothorax; marges latérales fortement infléchies surtout & la
base, trés étroitement rebordées. Stries suturales 4 peine marquées au sommet. Pattes
larges ; tibias postérieurs subtriangulaires, environ quatre fois plus longs que larges dans
leur plus grande largeur; bords latéraux des tibias antérieurs 4 peine visiblement
erénelés. Lignes fémorales des hanches intermédiaires arquées avant l’extrémité et ré-
joignant l’épisterne prés de la hanche postérieure. Premier segment de l’abdomen plus
court que le métasternum.
Loc. Seychelles. Silhouette, VIII. 1908, 1 8.
Carpophilini.
BracHyPEPLus Erichson, 1842, in Wiegm. Arch. i. p. 148.
3. Brachypeplus notatus, Murray.
Brachypeplus notatus Murray, 1864, Monogr. Nitid. p. 300, 309; Grouvelle, 1908,
Ann. Soc. Ent. Fr. Ixxvii. p. 328.
Brachypeplus patruclis Alluaud (nec Murray), Liste Coléopt. p. 114.
Loc. “Seychelles. Silhouette: from decaying fruit, VIII. 1908, 3 specimens.
Mahé: Baie Lazare, 1908, 1 specimen; Cascade Estate, ca. 1000 feet, 1909, 1 specimen ;
1892 (Alluaud). Praslin: Cétes d’Or Estate, XI. 1908, 2 specimens; 1892 (Alluaud).
La Digue, 1892 (Alluaud). lle Marie Anne, XII. 1908, 1 specimen.”
La description originale de cette espece a été faite sur des insectes provenant de
Ceylan; depuis elle a été retrouvée 4 Sumatra.
4. Brachypeplus equalis (Walker).
Nitidulopsis equals Walker, 1858, Ann. Mag. Nat. Hist. ser. 3, i. p. 206.
SECOND SERIES—ZOOLOGY, VOL. XVI. 13
98 PERCY SLADEN TRUST EXPEDITION
Brachypeplus equalis Olliff, 1885, Proc. Linn. Soc. N. 8. Wales, p. 70; Grouvelle,
1908, Ann. Soc. Ent. Fr. Ixxvii. p. 328.
Brachypeplus omalinus Murray, 1864, Monogr. Nitid. p. 299, 309, pl. 34, fig. 7.
Une série de 230 exemplaires.
Loc. Seychelles. ‘Silhouette: various localities, VIII.—IX. 1908; near coast ;
high forest near Mont Pot-i-eau, ca. 1500 feet; Mare aux Cochons and forest above.
Mahé, 1908—9: country above Port Glaud, 500—1000 feet ; near Morne Blanc; Cascade
Estate, 800—1000 feet, and forest above; Baie Lazare; Port Victoria (sea-level), from
rotting mangoes. Praslin: Cétes d’Or Estate, XI. 1908. Tle Marie Anne, XII. 1908.
This species is very abundant everywhere in rotting fruit, especially oranges and jak-fruit
(Artocarpus integrifolia), where it is found very often in company with Haptoncus
ocularis, certain Hydrophilide (Dactylosternum insulare, Cercyon uniformis Sharp and
Cercyon fructicola Scott), and certain Staphylinide.”
Cette espéce a été décrite sur des individus venant de Ceylan.
Cittaus Castelnau, 1835, Etud. Entom. i. p. 133.
TABLEAU DES CILLZUS DES SEYCHELLES.
1. Insecte glabre fe eis Hf 66 aS ep, bes Biss Sel ee
B, x pubescent... ae. nas ik ae ae ae am Hage}
2. Large; élytres un peu plus longs que larges ensemble __... subplagiatus, n. sp.
» Etroit; élytres presque deux fois plus longs que larges ensemble... MACYOS, N. Sp.
3. Stries des élytres trés fines, séparées par des intervalles beaucoup plus larges
que les stries a tis fa ie 250 ote .. opaculus, n. sp.
» Stries des élytres trés serrées cae a ie 500 oe Fee ees
4. Pubescence longue; base des élytres rougedtre ... bn megacephalus Cast.
. He tres courte; élytres concolores a4 wa mae confusus, 1. Sp.
5. Cilleus opaculus, n. sp. (Fig. 2.)
Subdepressus, sat latus, opaculus, tenue flavo-pubescens, nigricans; antennis clava
excepta pedibusque subfusco-rufis ; abdominis segmentis apice stricte ochraceo-marginatis.
Prothorax elytraque tenuissime asperuli, ille transverso, haec sesquilongiora quam simul
latiora, lineis pubescentibus, tenuibus, sat approximatis, apicem versus subconfusis ornata.
Long. 3,5—4,2 mill.
Allongé, paralléle, subdéprimé, médiocrement large, couvert d’une pubescence flave,
fine, couchée ; coloration générale noiratre, antennes sauf la massue et pattes rougedtres,
tres légerement enfumées ; segments de l’abdomen tres étroitement bordés au sommet de
jaune testacé. Massue des antennes oblongue, égale environ au tiers de la longueur
totale de lantenne. Téte densément pointillée. Prothorax et élytres tres finement
chagrinés. Prothorax environ une fois et un tiers plus large que long, 4 peine plus étroit
au sommet qu’a la base, tres faiblement arrondi sur les cétés; angles antérieurs obtus,
GROU VELLE—COLEOPTERA: NITIDULIDA, HETEROCERID A 99
émoussés, postérieurs arrondis; marges latérales étroitement rebordées en gouttiere ;
base rebordée ; région des angles postérieurs un peu plus claire que le disque. Ecusson
transversal, subtriangulaire. Elytres environ une fois et demie plus longs que larges
ensemble, largement arrondis aux angles postérieurs externes, finement chagrinés, garnis
d'une pubescence fine, couchée, disposée en lignes assez rapprochées, devenant confuses
vers Textrémité. Segments apparents de l’abdomen plus finement chagrinés que les
élytres, subdensément pointillés. Sommet des trois avant-derniers sternites frangés de poils
jaunes. Dernier sternite du male subtronqué, sternite supplémentaire petit, invisible
de dessus.
Loc. Seychelles. ‘Silhouette, 7 specimens, VIII.—IX. 1908; from Mare aux
Cochons, &c. Mahé; Cascade Estate, ca. 1000 feet, 1908—9, 3 specimens.”
6. Cilleus micros, n. sp. (Fig. 3.)
Subconvexus, angustus, nitidus, glaber, rufo-piceus; antennis pedibusque dilutioribus,
elytrorum apice late infuscato, basi plus minusve ochracea ; abdominis segmentis apicem
[
Fig. 2. Culleus opaculus, n. sp., x 6. Fig. 3. Cillews micros, n. sp., x 12.
versus plus minusve infuscatis, stricte testaceo-marginatis. Caput subdense punctulatum.
Prothorax vix paulo longior quam latior, in disco subparce, ad latera dense punctulatus ;
punctis subelongatis. Elytra circiter sesquilongiora quam simul latiora, striato-punctata ;
striis ad latera et ad apicem attenuatis ; intervallis latis, fere planis. Long. 3,2 mm.
-Trés allongé, paralléle, subconvexe, glabre, roux de poix; antennes et pattes plus
claires ; base des élytres plus ou moins jaundtre, sommet largement rembruni; segments de
Yabdomen étroitement bordés de testacé au sommet, devenant plus foncés vers l’extrémité
de ’abdomen. Massue des antennes briévement oblongue, moins longue que le tiers de la
longueur totale de l’antenne. Téte presque densément pointillée; yeux séparés du
sommet du prothorax par une distance subégale & leur diamétre longitudinal. Prothorax
un peu plus long que large, assez éparsément et finement pointillé sur le disque, plus
densément et plus fortement vers les bords latéraux ; marges latérales déclives, bordés
dans la moitié basilaire par une strie assez bien marquée, visible de dessus ; cdtés 4 peine
arqués ; angles antérieurs étroitement arrondis, postérieurs plus largement ; base finement
13—2
100 PERCY SLADEN TRUST EXPEDITION
rebordée aux extrémités. Hcusson transversal, subtriangulaire. Elytres environ une
fois et demie plus longs que larges ensemble, assez largement arrondis aux angles
postérieurs externes, ponctués-striés; stries effacées sur les marges latérales, atténuées
au sommet, se perdant dans une marge apicale finement alutacée; intervalles larges,
& peine convexes, presque ponctués en hone. Segments apparents de l’abdomen assez
densément et fortement ponctués sur les cétés. Avant-derniers sternites bordés de
jaune testacé. Métasternum 4 peime sillonné. Bord externe des tibias armé de dents
épineuses peu serrées, plus accentuées vers l’extrémité.
Loc. Seychelles. “Silhouette; 4 specimens, VIII.—IX. 1908; 1 specimen being
recorded as found between leaf-bases of a growing Pandanus Horner in forest above Mare
aux Cochons, over 1000 feet.”
7. Cilleus subplagiatus, n. sp.
Subdepressus, sat latus, nitidus, glaber, ater; antennis pedibusque plus minusve
rufo-piceis ; singulo elytro ad latera et juxta humerum vage rufo-maculato. Caput in
disco subparce subtenuiterque punctatum. Prothorax transversus, subparce punctulatus.
Elytra 1 et 4 longiora quam simul latiora, tenuiter punctato-striata ; tmtervallis latis.
Abdominis segmenta apice stricte testaceo-marginata Long. 6 mill.
Allongé, paralléle, presque déprimé, modérément large, glabre, noir ; antennes sauf la
massue roux de poix; pattes sauf les tarses plus foncées que les antennes; sur chaque
élytre une vague tache longitudinale, rougedtre, placée vers le cété, au-dessous de |’épaule;
sommet des segments de l'abdomen étroitement bordés de testacé. Massue des antennes
briévement oblongue, égale environ au quart de la longueur totale de l’antenne. Téte
éparsément pointillée sur le front, plus densément en avant et sur les cdtés. Prothorax
presqu'aussi long que large, plus finement et plus éparsément ponctué que la téte ; angles
antérieurs obtus, émoussés; postérieurs plus largement arrondis; marges latérales déclives,
bords latéraux finement rebordés, cachés, sauf & la base, lorsque l’insecte est examiné
de dessus; base finement rebordée. Ecusson transversal, subtriangulaire. Hlytres une
fois et un tiers plus longs que larges ensemble, assez largement arrondis aux angles
postérieurs, finement ponctués en liones s'atténuant un peu vers le sommet et s’effacant
un peu avant de l’atteindre. Segments apparents de l’abdomen densément pointillés.
Avant-derniers sternites finement bordés de testacé. Dernier sternite du male sinué,
pygidium tronqué; pygidium de la femelle largement arrondi.
Voisin de C. simont Grouy. (1893, Ann. Soe. Ent. Fr. lx. p. 383); distinct par son
prothorax moins transversal, plus rectangulaire, ses élytres plus finement striés et
marqués vers les épaules d'une vague tache ochracée.
Loc. Seychelles. ‘“ Mahé: high forest of Morne Blanc and Morne Pilot, XI. 1908,
1A toe
8. Cilleus confusus, n. sp.
Subdepressus, angustus, nitidulus, nigro-piceus, tenue flavo-aureo pubescens; pilis
elongatis, erectis, ad latera intermixtis; antennis pedibusque rufo-piceis; segmentis
abdominis apice stricte testaceo-marginatis. Caput subparce punctulatum. Prothorax
GROU VELLE—COLEOPTERA: NITIDULIDA, HETEROCERIDA 101
vix longior quam latior, angulis praecipue posticis sat late rotundatus, plus minusve
subparce punctatus; punctis subelongatis. Elytra ad apicem vix ampliata, circiter
sesquilongiora quam simul latiora, dense tenuiter lineato-punctata ; punctis elongatis, ad
latera et juxta apicalem marginem confusis. Long. 3,5 mill.
Trés allongé, paralléle, subdéprimé, couvert d’une pubescence flave doré, trés courte et
serrée, entrémélée sur les cotés du prothorax et des élytres de poils plus longs, dressés ;
brun de poix avec les antennes et les pattes rougedtres et les segments de l’abdomen
étroitement bordés de testacé. Massue des antennes oblongue, plus longue que le tiers de
la longueur totale de l’antenne. ‘Téte trés légérement convexe, environ aussi longue que
large, trés densément pointillée sur le disque; yeux séparés du sommet du prothorax
par un intervalle inférieur & leur diamétre longitudinal. Prothorax subcarré, moins
densément et moins fortement ponctué que la téte, un peu plus fortement ponctué sur les
marges latérales ; celles-ci infléchies, trés finement bordées sauf vers les angles antérieurs ;
bord antérieur tronqué; angles antérieurs arrondis, postérieurs plus largement; base
arquée, rebordée comme les cdtés et les angles postérieurs. Ecusson subtriangulaire,
environ deux fois plus large que long, densément ponctué. Elytres un peu plus d'une fois
et demie plus longs que larges ensemble, assez largement arrondis aux angles apicaux
externes, arqués séparément au sommet, un peu moins brillants que le prothorax,
densément couverts de points allongés, disposés en lignes serrées presque jusqu’au sommet.
Segments apparents de l’abdomen trés finement pointillés, le dernier un peu plus fortement,
Iégérement infléchi au sommet, tous présentant des poils dressés, insérés sur leurs cdtés.
Avant-derniers sternites étroitement bordés de testacé. Métasternum longuement
sillonné au sommet. Bord externe des tibias simple.
Toc. Seychelles. “Mahé: Cascade Estate, ca. 800 feet, 1 specimen.”
9. Cilleus megacephalus Castelnau.
Cilleus megacephalus Castelnau, 1835, Etud. Ent. i. p. 134; Alluaud, Liste Coléopt.
a 1A
Cilleus scapularis Fairmaire, 1869, Ann. Soc. Ent. Fr. xxxvii. p. 201.
Loc. Seychelles. “Silhouette ; from several localities near Mont Pot-i-eau, and
from Mare aux Cochons, 8 specimens, VIII.—IX. 1908. Mahé: Long Island, VII. 1908,
1 specimen.”
Cette espéce a été décrite sur des insectes provenant de Madagascar ; les exemplaires
des Seychelles ne different pas sensiblement du type.
CARPOPHILUS Stephens, 1830, Ill. Brit. Ent. ii. p. 30, 50.
Subg. UropHorus Murray, 1864, Monogr. Nitid. p. 342.
\
10. Carpophilus humeralis (Fabricius).
Nitidula humeralis Fabricius, 1798, Suppl. Ent. Syst. p. 74.
Carpophilus humeralis Murray, 1854, Monogr. Nitid. p. 343; Alluaud, Liste Coldopt.
p- 115; Grouvelle, 1908, Ann. Soc. Ent. Fr. Ixxvii. p. 330, 336.
36 exemplaires.
102 PERCY SLADEN TRUST EXPEDITION
Loc. “Seychelles: Silhouette, VIII. 1908: Mahé; Cascade Estate, ca. 1000 feet,
1909; Long Island, VII. 1908; 1905 (Gardiner) : Praslin; Cotes d’Or Estate, XI. 1908 :
Félicité, XII. 1908. Farquhar Atoll, 1905. Providence, Cerf Island, 1905.”
Se rencontre dans les régions tropicales et subtropicales de I Afrique, de |’Asie, et de
lArchipel Indo-Malais.
Var. foveicollis (Murray).
Carpophilus foveicollis Murray, 1864, Monogr. Nitid. p. 344; Alluaud, Liste Coléopt.
p- 115; Grouvelle, 1908, Ann. Soc. Ent. Fr. xxvii. p. 330.
La variété fovercollis Murr. n’a pas encore été signalée } Madagascar. Les exemplaires
des Seychelles sont nettement différentiés des exemplaires types de C. humeralis Fabr.
par labsence de la tache rouge humérale et par la ponctuation moins accentuée. Par
contre les fossettes latérales du prothorax ne sont pas toujours marquées.
21 exemplaires.
Loc. “Seychelles: Mahé; Cascade Estate, ca. 800—1000 feet, and forest above,
1908—9 ; forest on summit of Morne Blane, from rotting fruits of Pandanus seychellarum,
X. 1908 ; 1905 (Gardiner). Amirantes: Poivre Island, 1905.”
Subg. CarPoPHILUs, sen. str.
11. Carpophilus hemipterus (Linné).
Dermestes hemipterus Linné, 1767, Syst. Nat. ed. 12, 1. 11. p. 565.
Carpophilus hemipterus Murray, 1864, Monogr. Nitid. p. 362; Horn, 1879, ‘Trans.
Am. Ent. Soc. vi. p. 277; Ganglbauer, 1899, Kifer Mitteleur. i. p. 460; Alluaud, Liste
Coléopt. p. 115; Grouvelle, 1908, Ann. Soc. Ent. Fr. xxvii. p. 330, 337.
27 exemplaires.
Loc. ‘Seychelles: Mahé; Cascade Estate, ca. 800—1000 feet, 1908—9 : Silhouette;
high forest above Mare aux Cochons, IX. 1908, 1 specimen. Chagos Islands; Diego
Garcia, ‘in water-butt, East Point, 23. VII. 1905’ (T. B. Fletcher). Coetivy, 1905.
Farquhar Atoll, 1905. Providence; Cerf Island, 1905.”
Cosmopolite.
12. Carpophilus fusciceps, n. sp.
Suboblongus, sat convexus, nitidus, flavo-pubescens, piceo-testaceus, antennarum clava
capiteque subinfuscatis. Antennae subgraciles ; clava piriformi. Caput convexiusculum,
subparce punctulatum, inter antennarum bases bimpressum, antice subsinuatum ; oculis
submagnis, subprominulis. Prothorax transversus, antice quam postice angustior,
lateribus rotundatus, sat parce punctulatus; margine antico truncato; angulis anticis
obtusis, hebetatis; lateribus tenuiter marginatis; angulis posticis breviter rotundatis ;
basi subtruncata, tenuiter marginata. Scutellum transversissimum, subpentagonale.
Elytra paulo breviora quam simul latiora apice vix separatim subtruncata, sat parce
punctulata. Abdominis duo aperta segmenta, subparce punctulata, subrugosula.
Long. 1,8 mill.
GROUVELLE—COLEOPTERA: NITIDULIDA, HETEROCERIDA 103
Suboblong, environ deux fois plus long que large, assez convexe, brillant, couvert d’une
pubescence flave, fine, couchée, médiocrement courte, assez dense surtout sur les cotés, ne
masquant nullement la couleur du tégument, testacé, teinté de nuance de poix; massue
des antennes et téte trés légerement enfumées. Antennes relativement gréles; 3™°
article environ trois fois plus long que large, 4™° un peu allongé, 5™° & 8™° progressivement
un peu plus épais, 8™° amorcant la massue; celle-ci subpiriforme, environ une fois et
demie plus longue que large, et subégale environ au cinquiéme de la longueur totale de
Yantenne, dernier article un peu plus court et plus étroit que le précédent, terminé
en bouton trés surbaissé. Téte moins de deux fois plus large que longue, convexe,
rétrécie brusquement 4 la naissance des antennes, peu densément pointillée, biimpres-
sionnée entre les naissances des antennes ; tempes nulles ; bords latéraux trés convergents
en avant, échanerés par les yeux : saillie transversale de ces derniers subégale & la moitié
de la longueur de l’orbite. Prothorax un peu plus rétréci en avant qu’d la base, arrondi
sur les cotés, un peu plus de deux fois plus large dans sa plus grande largeur que long,
assez éparsément pointillé, tres finement rebordé sur les cétés et & la base; bord antérieur
subtronqué; angles antérieurs obtus, émoussés, postérieurs briévement arrondis; base
subtronquée. EKeusson subtriangulaire, presque subpentagonal, plus de deux fois plus large
a la base que long, subégal environ au quart de la largeur du prothorax & la base,
subdensément pointillé. Elytres, a la base, de la largeur du prothorax, arrondis aux
épaules, formant avec les cdtés du prothorax un angle obtus trés marqué, arqués sur les
cdtés, 4 peine plus larges dans leur plus grande largeur que le prothorax dans la sienne,
briévement arrondis aux angles apicaux externes, presque subtronqués ensemble au
sommet, environ aussi longs que larges ensemble dans leur plus grande largeur, sub-
éparsément et un peu plus fortement pointillés & la base que le prothorax ; marges
latérales assez fortement infléchies, étroitement rebordées. Segments visibles de
Pabdomen subéparsément pointillés, suabrugueux ; avant-dernier segment en partie caché.
Mésosternum sans caréne. Métasternum presque sans strie longitudinale. Lignes
fémorales des hanches intermédiaires recourbées 4 l’extrémité, lignes des hanches
postérieures rebordant la hanche.
Loc. “Seychelles. Silhouette: near Mont Pot-a-eau, VIII. 1908, 2 ?.”
13. Carpophilus scotti, n. sp.
Suboblongus, convexus, nitidus, flavo-pubescens, capite prothoraceque piceus, elytris
sordido-testaceus, antennis pedibusque dilutior. Antennae subgraciles, clava piriformi.
Caput subdepressum, parce punctulatum, inter antennarum bases fere arcuatim impressum,
antice deflexum et subtruncatum; oculis submagnis, vix prominulis. Prothorax trans-
versus, antice quam postice angustior, lateribus rotundatus, parce punctulatus; margine
antico antrorsum subarcuato, utrinque subsinuato ; anticis angulis obtusis, apice breviter
rotundatis, posticis obtusis, vix hebetatis; lateribus basique tenuiter marginatis, hac
subtruncata. Scutellum transversum, subtriangulare. Elytra longiora quam simul
latiora, apice vix separatim subtruneata. Abdominis duo aperta segmenta, parce
punctulata, subrugosula. Long. 1,2 mill. 7
Suboblong, environ deux fois et demie plus long que large, convexe, brillant, couvert
104 7 PERCY SLADEN TRUST EXPEDITION
dune pubescence flave, fine, couchée, courte, assez dense, ne masquant nullement la
couleur du tégument ; téte et prothorax brunatres, élytres d’un testacé sale, antennes et
pattes plus claires. Antennes relativement gréles ; 3™° article nettement moins de trois
fois plus long que large, 4™° carré, 5™° & 8™° progressivement un peu plus épais, 8™°
amoreant la massue ; celle-ci subpiriforme, environ une fois et demie plus longue que large,
et subégale environ au quart de la longueur totale de l’antenne ; dernier article plus court
et plus étroit que le précédent, terminé en bouton convexe, déprimé. Téte environ deux
fois plus large que longue, faiblement convexe, assez brusquement rétrécie 4 la naissance
des antennes, éparsément pointillée, biimpressionnée, presquimpressionnée en are entre
les naissances des antennes, infléchie en avant de cette impression, subtronquée au bord
antérieur ; tempes nulles; bords latéraux vus de dessus tres convergents en avant, & peine
échancrés par les yeux ; ceux-ci présentant une saillie transversale inférieure 4 la moitié de
la longueur de l’orbite. Prothorax un peu plus rétréci en avant qu’a la base, arrondi sur
les cétés, un peu moins de deux fois plus large dans sa plus grande largeur que long,
éparsément pointillé, trés finement rebordé sur les cdtés et & la base ; bord antérieur trés
faiblement arqué en avant dans le milieu, subsinué vers les extrémités; angles antérieurs
vus de dessus, obtus, britvement arrondis, postérieurs obtus, émoussés ; base subtronquée.
Ecusson subtriangulaire, environ deux fois plus large & la base que long, moins large que
ree
Fig. 4. Carpophilus scotti, n. sp., x 16.
le quart de la base du prothorax, pointillé, tres légerement convexe. Hlytres & peu pres
de la largeur du prothorax, en angle obtus, émoussé aux épaules, formant avec les cétés
du prothorax un angle obtus trés marqué, arqués sur les cétés, 4 peine plus larges dans
leur plus grande largeur que le prothorax dans la sienne, trés brievement arrondis aux
angles apicaux externes, presque subtronqués ensemble au sommet, plus longs que larges
ensemble dans leur plus grande largeur, éparsément et plus fortement pointillés que le
prothorax ; convexité transversale tres marquée; marges latérales fortement infléchies,
finement rebordées. Segments visibles de l’abdomen éparsément pointillés ; avant-dernier
segment en partie caché. Mésosternum sans caréne longitudinale ; métasternum longi-
tudinalement strié. Lignes fémorales des hanches intermédiaires et postérieures bordant
les hanches.
Loc. ‘Seychelles. Silhouette : near Mont Pot-a-eau, VIII. 1908, 1 specimen.”
Subg. Myornorax Murray, 1864, Monogr. Nitid. p. 372.
14. Carpophilus dinidiatus (Fabricius).
Nitidula dimidiata Fabricius, 1792, Ent. Syst. 1. p. 261.
GROUV ELLE—COLEOPTERA: NITIDULID®, HETEROCERID A 105
Var. contengens (Walker).
Nitidula contingens Walker, 1858, Ann. Mag. Nat. Hist. 3 ser. ii. p. 206.
Carpophilus contingens Olliff, 1885, Proc. Linn. Soc. N. S. Wales, x. jo OY 3
Grouvelle, 1908, Ann. Soc. Ent. Fr. Ixxvii. p. 332.
Carpophilus biguttatus Murray, 1864, Monogr. Nitid. p. 364; Alluaud, Liste
Coléopt. p. 115.
Loc. Seychelles: Félicité, XII. 1908, 8 exemplaires : 1892 (Alluaud).
Var. ochropterus Klug.
Carpophilus ochropterus Klug, 1862, Peters Reis. p. 209, pl. 12, fig. 9; Murray,
1864, Monogr. Nitid. p. 377; Fairmaire, 1869, Ann. Soc. Ent. Fr. sér. 4, ix. jo AOO s
Alluaud, Liste Coléopt. p. 116; Grouvelle, 1908, Ann. Soc. Ent. Fr. xxvii. p. 332.
42 exemplaires.
Loc. ‘Seychelles: Silhouette, VIII. 1908: Mahé; country above Port Glaud,
500—1000 feet, XI. 1908; from rotting fruit of Pandanus seychellarum, summit of
Morne Blanc, X. 1908; Cascade Estate, ca. 1000 feet, 1908—9: Praslin, Cotes d’Or
Estate, XI. 1908: fle Marie Anne, XII. 1908. F arquhar Atoll, 1905. Providence, Cerf
Island, 1905.”
La variété contingens tend & se répandre dans toutes les régions tropicales et
subtropicales ; la variété ochropterus semble localisée, jusqu’’ ce jour, sur la cdte orientale
de l'Afrique, 4 Madagascar, aux Mascareignes, aux Comores et aux Seychelles.
15. Carpophilus angustatus Murray.
Carpophilus angustatus Murray, 1864, Monogr. Nitid. p. 376 ; Fairmaire, 1869, Ann.
Soc. Ent. Fr. sér. 4, ix. p. 199; Alluaud, Liste Coléopt. p. 115.
Loc. “Farquhar Atoll, 1905, 8 specimens. Seychelles: Silhouette, VIII. 1908,
1 specimen; 1892 (Alluaud). Madagascar.”
Nitidulini.
Hapronous Murray, 1864, Monogr. Nitid. p. 401.
Haptoncura Reitter, 1875, Verhandl. Nat. Ver. Briinn, xii. p. 61, 64, sep. p. 11, 14.
16. Haptoncus ocularis (Fairmaire).
Epurea ocularis Fairmaire, 1849, Rev. Zool. sér. 2, 1. p. 363, sep. p. 28; Reitter,
1873, Verhandl. Nat. Ver. Briinn, xii. p. 27.
Haptoncura ocularis Reitter, 1875, Verhandl. Nat. Ver. Briinn, xiii. p. 62 (sep. p. 12);
Blackburn, 1902, Trans. R. Soc. S. Austr. xxvi. p. 306; xxvii. 1903, p. 116.
Haptoncus ocularis Fauvel, 1908, Rev. d’Ent. Caen, xxii. p. 301; Grouvelle, 1908,
Ann. Soc. Ent. Fr. xxvii. p. 345.
Nitidula significans Walker, 1858, Ann. Mag. Nat. Hist. ser. 3, ii. p. 206. °
SECOND SERIES—ZOOLOGY, VOL. XVL. 14
106 PERCY SLADEN TRUST EXPEDITION
Haptoncus tetragonus Murray, 1864, Monogr. Nitid. p. 401, pl. 33, fig. 7; Reitter,
1884, Wien. Ent. Zeitg. ii. p. 259; Sharp, 1878, Trans. Ent. Soc. Lond. p. 139; 1908,
Fn. Hawaiiensis, i. 5, p. 508.
Epurea theme: Reitter, 1873, Verhandl. Nat. Ver. Briinn, xu. 1873, p. 28, 41.
Haptoncus decoratus Reitter, 1873, l.c. p. 28, 41; Fauvel, 1903, Rev. d’Ent. Caen,
xxi. p. 801; Alluaud, Liste Coléopt. p. 116; Grouvelle, 1906, Ann. Soc. Ent. Fr. Ixxv.
jos Bo
Epurea bifasciata Krantz, 1895, Deutsche Ent. Zeitschr. p. 148.
L’Haptoneus decoratus Reitt. se sépare d’H. ocularis Fairm. par les marges explanées
du prothorax beaucoup plus larges, par la ponctuation générale plus forte, par la
troncature apicale des élytres moins marquée et par la tendance 4 la coloration noire plus
accentuée ; mais ces divers caractéres n’ont pas une fixité absolue, et comme d’autre part
H. ocularis Fairm. joint 4 son aire d’expansion trés étendue, une variabilité considérable,
il faut considérer H. decoratus Reitter comme une variété d’H. ocularis plus spéciale & la
région Malgache.
A-peu-prés 350 exemplaires.
Loc. “Seychelles. Silhouette, VITI.—IX. 1908 ; various localities, near coast, and
in the high forests near Mont Pot-a-eau and Mare aux Cochons. Mahé, 1908—9 ; various
localities ; high forest of Morne Blanc and Morne Pilot, including some found in rotting
fruits of Pandanus seychellarum in company with Carpophilus humeralis var. foveicollis,
X. 1908; Cascade Estate, ca. 1000 feet; Baie Lazare, &c. Praslin: Cétes d’Or Estate,
XI. 1908; 1892 (Alluaud). La Digue, 1892 (Alluaud). Very abundant everywhere in
rotting fruit, especially oranges and jak-fruit, almost always in company with Brachy-
peplus equalis and certain Hydrophilide and Staphylinide (see under B. equalis,
p- 98).”
Ancien continent ; régions tropicales et subtropicales.
17. Haptoncus luteolus (Erichson).
Epurea luteola Erichson, 1843, in Germ. Zeitschr. iv. p. 272; Horn, 1879, Trans.
Am. Ent. Soc. vii. p. 301.
Haptoncura luteola Reitter, 1875, Verhandl. Nat. Ver. Briinn, xiii. p. 62, sep. p. 12.
Hoaptoncus luteolus Alluaud, Liste Coléopt. p. 116; Sharp, 1890, Godm. and Salvin,
Biol. Centr.-Amer. Col. 11. 1, p. 305; Grouvelle, 1906, Ann. Soc. Ent. Fr. lxxv. p. 75;
1908, Ixxvii. p. 346 (note).
Nitidula intendens Walker, 1858, Ann. Mag. Nat. Hist. ser. 3, 1. p. 206.
Haptoncus pauperculus Reitter, 1873, Syst. Eintheil. Nitid. p. 179.
Haptoncus floreolus Sharp, 1890, Godm. and Salvin, Biol. Centr.-Amer. Col. ii. 1,
pl. 9, fig. 24, 2.
Haptoncus pubescens Murray, 1864, Monogr. Nitid. p. 403.
Haptoncus subquadratas Reitter, 1877, Mittheil. Miinch. Ent. Ver. i. p. 22;
Fauvel, 1903, Rev. d’Ent. Caen, xxii. p. 301; Grouvelle, 1906, Ann. Soc. Ent. Fr.
boxy, jo 75.
GROUVELLE—COLEOPTERA: NITIDULIDA, HETEROCERIDA 107
52 exemplaires.
Loc. ‘Seychelles. Silhouette, WL 1908 ; near coast, and Mare aux Co-
chons, over 1000 feet. Mahé: country above Port Glaud, 500—1000 feet, XI. 1908;
Cascade Estate, ca. 1000 feet, 1908—9; 1892 (Alluaud). Praslin: Cotes d’Or Estate,
XI. 1908; 1892 (Alluaud). fle Marie Anne, XII. 1908.”
Cosmopolite ; régions tropicales et subtropicales.
18. Haptoncus minutus (Reitter).
Epurea nanuta Reitter, 1873, Syst. Hintheil. Nitid. p. 27, 40.
Haptoncura minuta Reitter, 1875, Verhandl. Nat. Ver. Briinn, xiii. p. 62, sep. p. 12.
Haptoncus minutus Grouvelle, 1905, Mém. Soc. Esp. Hist.-Nat. i. p. 242; 1906,
Ann. Soc. Ent. Fr. Ixxv. p. 73; Ixxvii. 1908, p. 346.
Haptoncura nitescens Grouvelle, 1877, Ann. Mus. civ. Genova, xxxviii. p. 362.
9 exemplaires.
Loc. ‘Seychelles. Silhouette; Mare aux ; Cochons, over 1000 feet, IX. 1908.
Mahé: Cascade Estate, 800—1000 feet, 1908—9.”
Ancien continent ; régions tropicales et subtropicales.
19. Haptoncus sobrinus Grouvelle.
Haptoncus sobrinus Grouvelle, 1894, Ann. Soc. Ent. France, p. 15 (fig.); Alluaud,
Liste Colépt. p. 116.
Loc. Seychelles: Mahé, 1892 (Alluaud). Mascareignes: La Réunion; Maurice
(Alluaud).
Cette espéce ne se trouve pas parmi les spécimens recoltés par la Percy Sladen Trust
Expedition.
20. Haptoncus obscurus, 0. sp.
Oblongus, modice convexus, nitidulus, tenuissime flavo-griseo pubescens, piceus,
antennis clava excepta, pedibus, corpore subtus ex parte dilutior. Antennarum clava
sesquilongior quam latior. Caput transversum, convexiusculum, subdense punctulatum,
inter antennarum bases biimpressum; lateralibus marginibus, convergentibus ; oculis
magnis. Prothorax transversus, antice subvalde, postice vix angustus, lateribus arcuatus,
subdense punctulatus ; angulis anticis obtusis; marginibus lateralibus modice explanatis,
laevibus ; angulis posticis obtusis, hebetatis; basi subtruncata. Scutellum triangulare,
dense punctulatum. Elytra humeris rotundata, lateribus parum arcuata, apice latissime
separatim arcuata, 1 et 4 longiora quam simul latiora, quam prothorax minus dense sed
magis valide punctulata. Long. 1,2—1,4 mill.
Oblong, environ deux fois et un quart plus long que large dans sa plus grande
largeur, modérément brillant et convexe, couvert d’une pubescence trés fine, flave cendrée,
brun de poix avec les antennes sauf la massue, les pattes, la partie antérieure du dessous
du corps plus claires, un peu rougedtres. Antennes relativement longues; 3™° article
plus d'une fois et demie plus long que large, 4™°, 5™° et 6™° subégaux, subcarrés, 7™° et
surtout 8™° trés transversaux, 9™° 4 11™° subégaux, formant une massue brusque, un peu
plus longue que large, dont le dernier article légérement plus étroit que les précédents est
14—2
108 PERCY SLADEN TRUST EXPEDITION
terminé par un bouton en forme de céne trés surbaissé. Téte moins de deux fois plus
large que longue, un peu convexe, densément pointillée, biimpressionnée entre les nais-
sances des antennes; bord antérieur tronqué; cdtés subrectilignes, convergents entre la
base et la naissance des antennes, brusquement rétrécis aprés les yeux a la naissance des
antennes ; angles postérieurs nuls; yeux gros, i facettes relativement fortes, présentant
une saillie latérale inférieure 4 la moitié de Vorbite. Prothorax assez rétréci en avant,
trés faiblement a la base, arqué ‘sur les c6tés, environ deux fois plus large dans sa plus
erande largeur que long, plus ou moins densément pointillé; bord antérieur subtronqué ;
angles antérieurs obtus ; marges latérales modérément explanées, lisses ; angles postérieurs
obtus, un peu émoussés; base subtronquée. Ecusson triangulaire, environ deux fois plus
large 4 la base que long, densément et plus fortement ponctué que le prothorax. Elytres de
la largeur du prothorax 4 la base, arrondis aux épaules, arqués sur les cotés et trés faiblement
élargis, tres largement et séparément arrondis au sommet, presque subtronqués, environ
une fois et un quart plus longs que larges dans leur plus grande largeur, moins densément,
mais plus fortement pointillés que le prothorax ; marges latérales fortement infléchies, assez
étroitement rebordées explanées ; ponctuation atténuée vers le sommet des élytres ; stries
suturales nulles. Yeux séparés en dessous, par un intervalle subégal 4 leur diamétre trans-
versal. Tibias terminés 4 langle apical externe en angle aigu, bien marqué.
Hspéce voisine des H. piconus Grouv. et dispersus Grouv.; colorée sensiblement
comme la premiére, mais ayant les élytres notablement plus courts, beaucoup plus foncée
que la deuxiéme et en plus nettement convexe, plus parallele.
22 exemplaires.
Loc. “Seychelles. Mahé: Cascade Estate, ca. 1000 feet, I. 1909; from leaf-bases of
growing Stevensonza-palm, near Morne Blanc, 1908, 13 specimens; coastal plain at Anse
aux Pins, I. 1909; Port Victoria, XII. 1908; Long Island, VII. 1908.”
Srenipota Hrichson, 1843, in Germ. Zeitschr. iv. p. 300.
21. Stelidota explanata, n. sp. (Figs. 5, 6.)
Stelidotarum latarum facies, sed prothoracis lateralibus marginibus praecipue antice
posticeque latissime concavo-explanatis, medio intus arcuatim terminatis ; coxarum posti-
carum axillari spatio apice acuto-anguloso.
Mas. Tibiae intermediae apice intus breviter incrassatae, posticae arcuatae, apice
haud longe intus dilatatae et truncatae. Femina. Tibiae posticae rectae, apice intus
modice dilatatae. Long. 2,5—8,5 mill.
Ovale, large, convexe, brillant, couvert d’une pubescence flave-cendrée, trés courte,
éparse sur la téte et le prothorax, dessinant sur les élytres des lignes fines alternative-
ment mieux marquées. Antennes gréles; 3™° article environ quatre fois plus long que
large. Téte densément ponctuée, présentant en avant, entre les antennes, une forte
dépression qui s‘étend sur l’epistome. Prothorax fortement transversal, arrondi sur les
cdtés, subparalleéle a la base, largement échancré au bord antérieur; angles antérieurs un
peu saillants en avant, arrondis, postérieurs presque droits, 4 peine émoussés; base sinuée
et obliquement tronquée de chaque cdté & l’extrémité, finement et étroitement rebordée ;
GROUVELLE—COLEOPTERA : NITIDULIDA, HETEROCERID i 109
marge antérieure infléchie par rapport au disque; marges latérales trés largement ex-
planées-concaves surtout en avant et en arriére; partie concave limitée en dedans, au
milieu, par une partie arquée; ponctuation médiocre, assez écartée sur le disque, plus dense
et plus forte vers les cdtés. Ecusson transversal, subtriangulaire, arrondi au sommet, trés
légérement convexe, éparsément ponctué. Elytres ovales, acuminés vers le sommet, con-
tinuant presqu’d la base la courbure du prothorax, briévement et séparément arrondis au
sommet, environ aussi longs que larges ensemble 4 la base, striés-ponctués ; intervalles
alternes un peu élevés surtout vers le sommet, les autres devenant convexes dans la
a
ia
Fig. 5. Stelidota explanata, n. sp., 3, x 10. Fig. 6. Stelidota explanata, n. sp., lignes
fémorales des hanches postérieurs.
partie apicale; marges latérales assez largement explanées 4 la base, se rétrécissant pro-
gressivement vers le sommet. Lignes fémorales des hanches postérieurs fermées, formant
un angle aigu, acuminé au sommet, atteignant presque le sommet du premier segment de
Vabdomen (fig. 6). Métasternum presque densément ponctué; premier segment de l’ab-
domen plus fortement ponctué 4 la base que le métasternum.
Male. ‘Tibias intermédiaires bri¢vement élargis en dedans vers l’extrémité; pos-
térieurs arqués, presque brusquement élargis et tronqués en dedans avant l’extrémité.
Femelle. Tibias postérieurs droits, modérément dilatés en dedans dans le dernier tiers de
la longueur.
5 exemplaires.
Loc. ‘Seychelles. Silhouette, VIII. 1908. Mahé: forest on summit of Morne
Blane; Cascade Estate, ca. 1000 feet, and forest above, 1908—9.”
Lastopactytus Perty, 1830—1834, Del. Anim. p. 34.
Lordites Krichson, 1843, in Germ. Zeitschr. iv. p. 316.
22. Lasiodactylus pictus (MacLeay).
Nitidula picta Macleay, 1825, Annulosa Javanica, p. 40, ed. Lequien, 1833, p. 147.
Lasiodactylus pictus Arrow, 1909, Ann. Mag. Nat. Hist. ser. 8, iv. p. 191.
Lordites glabricola Candeze, 1861, Mém. Liége, xvi. p. 340.
Lasiodactylus glabricola Reitter, 1873, Syst. Eintheil. Nitid. p. 88; 1884, Wien.
Ent. Zeitg. ii. p. 266; Grouvelle, 1908, Ann. Soc. Ent. Fr. lxxvii. p. 368.
Lordites costulatus Fairmaire, 1868, Ann. Soc. Ent. Fr. sér. 4, vill. p. 775.
110 PERCY SLADEN TRUST EXPEDITION
Lasiodactylus costulatus Reitter, 1873, Syst. Eimtheil. Nitid. p. 88; Riinckel, in
Grandidier, Hist. Madag. Col. pl. xxx. fig. 5; Grouvelle, 1908, Ann. Soc. Ent. Fr.
Ixxvil. p. 365; Alluaud, Liste Coléopt. p. 120.
Lasiodactylus testudinarius Reitter, 1873, Syst. Hintheil. Nitid. p. 87; Grouvelle,
1895, Bull. Soc. Ent. Fr. p. cclvii.; 1908, Ann. Soc. Ent. Fr. Ixxvil. p. 365.
Loc. “Seychelles. Mahé: from near Morne Blanc, X. 1908, 12; from rotting
mangoes at Port Victoria, XII. 1908, 24 specimens.”
Cette espéce se trouve répandue dans toutes les régions tropicales et subtropicales
depuis Madagascar jusqu’a l’extrémité de |’ Archipel Malais et jusque sur la partie sud
du Japon.
23. Lasiodactylus breviusculus Fairmaire.
Lasiodactylus breviusculus Fairmaire, Ann. Soc. Ent. Fr. sér. 4, viii. 1868 (1869),
p. 775; Riincekel, in Grandid. Hist. Madag. Col. pl. 32, fig. 14; Alluaud, Liste Coléopt. p. 119.
7 exemplaires.
Loc. ‘Seychelles. Mahé: Cascade Estate, ca. 1000 feet, 1908—9; Baie Lazare,
1908; 1907 (Thomasset) ; 1905 (Gardiner).”
Madagascar. [les Mascareignes.
Cybocephalini.
CypocrPHaLus Erichson, 1844, in Germ. Zeitschr. v. p. 441.
Phantazomerus Jacquelin du Val, 1854, Bull. Soc. Ent. Fr. p. 307.
Stagnomorpha Wollaston, 1854, Ins. Mader. p. 482.
Le genre Cybocephalus est largement représenté dans la faune des [es Seychelles ;
cing espéces nouvelles sont décrites dans ce mémoire, toutes au moins jusqu’a ce jour sont
spéciales & cette faune et toutes sont établies sur un nombre d’individus important,
souvent méme considérable. Aucune faune ne nous a jamais donné une semblable
caractéristique™.
Toutes les especes décrites dans ce travail ont la massue des antennes dilatée en
dedans, le dessous de la téte coupé transversalement par un sillon qui doit probablement
recevoir le bord antérieur du sternum dans le mouvement dinflexion de la téte, le sommet
du métasternum creusé de fossettes recevant les pattes dans la position de repos et la
région basilaire des élytres marquée d’un pli plus ou moins subcaréniforme et plus ou
moins long qui limite la partie de la marge basilaire des élytres cachée au repos par le
pronotum.
* The data of capture of the Cybocephalus show the following facts: four out of the five species were found
in more than one island of the Seychelles archipelago, and one of these was also found in a coral island of the
Amirantes group: all the five species were found in a number of different localities within each island, and often
on several different occasions in the same locality; they are distributed in all parts of the islands, each of the
five having been found both in cultivated country near sea-level and also at considerable elevations in the
endemic forests. This last statement applies also to several other of the Nitidulide, but it is the reverse of the
way in which many of the Seychelles beetles are distributed, many species being found either exclusively in the
endemic forests, or exclusively in the cultivated country.—H. Scorr.
GROUV ELLE—COLEOPTERA: NITIDULIDA, HETEROCERID Ai 111
Enfin l'étude d’un grand nombre d’exemplaires montre que les colorations métalliques
si fréquentes chez les Cybocephalus ne conduisent pas 4 une précision absolue quant & la
fixation des espéces et des sexes. Les bordures métalliques de la marge antérieure du
prothorax sont plus ou moins développées, elles peuvent méme disparaitre.
TABLEAU DES CYBOCEPHALUS DES SEYCHELLES.
1. Pattes presque entierement testacées 68 ae Bs at ie Bey ee
f & noires ...
2. ‘Trés convexe, subhémisphérique ; élytres moins longs que larges ensemble
subpallipes, n. sp.
, Moins convexe, profil faisant ressortir le maximum de courbure vers l’avant-corps de
Vinsecte ; élytres plus longs que larges ensemble ; taille trés petite .
tantillus, n. sp.
3. Noir dans les deux sexes ; élytres 4 peine aussi longs que larges ensemble
minutus, n. sp.
,, Au moins la marge antérieure du prothorax métallique... ae site ey nd
4. Profil subhémisphérique ; élytres suboblongs, au plus aussi longs que larges ensemble
i brevis, n. sp.
, Profil faisant ressortir le maximum de courbure vers l’avant-corps de linsecte : élytres
ovales, plus longs que larges ensemble... sue Es 1. MUNIMUS, N. Sp.
24. Cybocephalus tantillus, n. sp.
Ovatus, convexus, nitidissimus, ater; corpore subtus nigro-brunneus, prosterno
abdomineque dilutioribus, antennis pedibusque fulvo-testaceis. Caput transversissimum
antice vix productum, truncatum, vix perspicue punctulatum. Prothorax laevis; angulis
anticis subrectis hebetatis, posticis obtusis. Elytra basi modice arcuata et humeris
breviter rotundata, apice apud marem conjunctim rotundata et circiter tam elongata
quam simul lata, apud feminam conjunctim subacuminata et paulo longiora quam simul
lata. Metasternum subparce punctulatum, glabrum. Abdomen vix pubescens. Long.
0,5 mill.
Faiblement ovale, trés nettement plus d’une fois et demie plus long que large dans sa
plus grande largeur, convexe, brillant, lisse ou presque lisse, noir; dessous du corps brun
noiratre, prosternum et abdomen plus clairs; antennes et pattes fauve testacé, fémurs
postérieurs un peu enfumés; marges latérales du prothorax et extrémité des élytres
souvent peu foncés. Massue des antennes environ aussi longue que le quart de la longueur
totale de l'antenne, dernier article plus court et un peu plus étroit que le précédent. Téte
& peine saillante en avant des naissances des antennes, subtronquée au bord antérieur.
Prothorax largement échancré au bord antérieur; angles antérieurs presque droits
émoussés, postérieurs obtus ; cdtés faiblement arrondis, & peine visiblement rebordés.
Keusson triangulaire, plus de deux fois plus large & la base que long. Elytres modérément
arqués & la base lorsquils sont vus de dessus, bri¢vement arrondis aux épaules lorsque
112 PERCY SLADEN TRUST EXPEDITION
celles-ci sont vus de face, arrondis ensemble au sommet et environ aussi longs que larges
4 la base chez le male, subacuminés ensemble au sommet et un peu plus longs que larges
ensemble chez la femelle, trés étroitement rebordés sur les marges latérales; de chaque
cdté dela base un pli oblique, assez allongé, mais n’atteignant pas le bord latéral, limitant
en dehors une marge brusquement infléchie par rapport 4 la surface de l’élytre surtout
contre la base. Sillons antennaires faiblement convergents. Métasternum presqu’éparsé-
ment ponctué. Segments de l’abdomen se développant dans un plan assez incliné par
rapport au plan du métasternum. Pubescence de l’abdomen peu accentuée. Lignes
fémorales des hanches postérieures divergentes, rejoignant tangentiellement le sommet
du premier segment. Profil longitudinal gibbeux en avant.
36 exemplaires.
Loc. ‘‘Seychelles. Silhouette: Mare aux Cochons, plateau and near by, IX. 1908.
Mahé: Cascade Estate, ca. 800—1000 feet, 1908—9; forest near Mount Harrison, ca.
1700 feet, III. 1909; forest between Trois Fréres and Morne Seychellois, 1500—2000 feet,
XII. 1908; Mare aux Cochons district, ca. 1500 feet, I. 1909; Anonyme Island, I. 1909.
Praslin: Cotes d’Or Estate, XI. 1908, 3 specimens, one of which is recorded as found
between leaf-bases of a growing ¢ ‘ Coco-de-Mer’ (Lodoicea seychellarum) in the Vallée
de Mai.”
25. Cybocephalus minutus, n. sp.
Ovatus, convexus, nitidissimus, ater; antennis, anticis et intermediis pedibus
prosternoque subsordido-testaceis. Antennarum clava intus dilatata. Caput vix
perspicue punctulatum. Prothorax antice utrinque anguloso-excisus, angulis anticis
obtusis, late hebetatis, posticis rotundatis, laevis. Elytra basi humerisque rotundata,
apud marem apice conjunctim rotundata, apud feminam apice conjunctim subacuminata,
circiter tam elongata quam simul lata; prope humeros et juxta basin breviter
subearinata, in exteriora carinae inflexa. Metasternum vix perspicue alutaceum.
Abdominis segmenta vix perspicue pubescentia. Long. 0,5—0,8 mill.
Ovale, environ une fois et un tiers plus long que large dans sa plus grande largeur,
trés convexe surtout dans la partie antérieure, trés brillant, & peine visiblement pointillé
méme avec un fort grossissement, noir; antennes, pattes antérieures et intermédiaires et
prosternum d’un testacé un peu enfumé, parfois marges latérales du prothorax et
extrémité des élytres plus claires. Antennes terminées par une massue de trois articles
assez bien marquée, dilatée en dedans, 3™° article allongé, plus de deux fois plus long que
large, 5™° plus long que 4™° et 6™°, 8™° amorgant la massue, 11™° plus court et plus étroit
que 10™*. Téte peu saillante en avant des naissances des antennes, brievement tronquée
au bord antérieur. Prothorax fortement sinué aux extrémités du bord antérieur, trés
finement rebordé aux angles antérieurs et postérieurs et sur les cotés, fortement émoussé
aux angles antérieurs, largement arrondi aux angles postérieurs. Hcusson triangulaire,
transversal. Hlytres arrondis & la base et aux épaules, largement arrondis au sommet
chez le male, plus étroitement chez la femelle, environ aussi longs que larges ensemble
dans leur plus grande largeur ; marges latérales étroitement rebordées ; de chaque coté
contre la base un pli oblique, n’atteignant pas le bord latéral, limitant en dedans, une
GROU V ELLE—COLEOPTERA: NITIDULIDA, HETEROCERIDA 113
marge fortement infléchie par rapport a la surface de lélytre, surtout contre la base.
Sillons antennaires légérement convergents. Métasternum 4 peime visiblement alutacé,
segments de l’'abdomen se développant dans un plan un peu incliné par rapport au plan
dumétasternum. Pubescence del’abdomen trés courte, couchée, orientée dans la longueur,
disposée en ligne transversale sur lessegments. Lignes fémorales des hanches postérieures
trés divergents formant un angle obtus, a peine émoussé, avec le sommet du 1° segment
abdominal. Profil longitudinal plitot gibbeux.
8 exemplaires.
Loc. ‘Seychelles. Silhouette: Mare aux Cochons, IX. 1908. Mahé: above Port
Glaud, 500—1000 feet, XI. 1908; Cascade Estate, ca. 1000 feet, I. 1909; Long Island,
VII. 1908; Anonyme Island, I. 1909. Praslin: from between leaf-bases of growing
$ ‘Coco-de-Mer’ (Lodoicea), XI. 1908, 1 specimen (cf. C. tantillus).”
26. Cybocephalus mimmus, n. sp.
Breviter ovatus, convexissimus, nitidissimus, ater; apud marem capite et prothoracis
antico margine viridi-aeneis ; primi segmenti abdominis medio et posticis femoribus plus
minusve aeneis ; apud feminam capite plus minusve fusco-aeneo; prothoracis antico margine
plus minusve stricte viridi-aeneo. Caput antice modice productum, breviter truncatum,
subdense et vix perspicue punctulatum. Prothorax sublaevis; anticis angulis rectis,
hebetatis, posticis late rotundatis. Elytra basi modice rotundata, humeris obtusa et
hebetata, apice conjunctim rotundata, sublaevia, ad latera vix perspicue punctulata, prope
humeros oblique vix carinata et inter carinam et lateralem marginem subabrupte inflexa.
Metasternum dense tenueque punctulatum. Abdomen pubescens. Long. 0,6—0,9 mill.
Ovale, environ une fois et demie plus long que large dans la plus grande largeur, trés
convexe, tres brillant, noir; antennes testacées, 4 peine rembrunies & l’extrémité ; téte,
marge antérieure du prothorax vert métallique, parfois bleuatre chez le male ; téte noire
plus ou moins bronzée et marge antérieure du prothorax plus ou moins étroitement vert
métallique chez la femelle ; dessous du corps noir, milieu du premier segment de l'abdomen
et souvent fémurs intermédiaires et postérieurs vert métallique chez le male. Massue des
antennes assez bien marquée, dilatée en dedans, égale environ au tiers de la longueur de
Yantenne, dernier article plus étroit et moins long que le précédent. Téte modérément
saillante en avant des naissances des antennes, bri¢vement subtronquée au bord antérieur,
assez densément, mais 4 peine visiblement pointillée. Prothorax arrondi au bord antérieur,
fortement sinué vers les extrémités, faiblement arqué; sur les cdtés, fortement arrondi
aux angles postérieurs ; angles antérieurs presque droits ; ponctuation encore plus fine et
beaucoup plus espacée que celle de la téte; cdtés et angles antérieurs et postérieurs trés
finement rebordés. Ecusson triangulaire, plus de deux fois plus large 4 la base que long,
également & peine visiblement pointillé. Elytres largement arrondis aux épaules, presque
subtronqués, 4 peine visiblement ponctués sur le disque, plus densément et plus visible-
ment vers les cOtés, arrondis ensemble au sommet & peu pres de la méme maniére dans les
deux sexes, environ aussi longs que larges ensemble a la base, trés finement rebordés sur
les cotés ; régions humérales coupées par un pli oblique et assez brusquement infléchies
entre ce pli et le bord latéral. Sillons antennaires faiblement convergents. Métasternum
SECOND SERIES—ZOOLOGY, VOL. XVI. 15
114 PERCY SLADEN TRUST EXPEDITION
et premier segment de l’'abdomen densément et trés finement pointillés; segments de
labdomen se développant sensiblement dans le méme plan que le métasternum, celui-ci
marqué, au sommet, de chaque cOté par une impression concave, lisse, recevant la patte
lorsqu’elle s'eflace contre le métasternum. Pubescence plus marquée sur l’abdomen.
Lignes fémorales des hanches postérieurs rejoignant tangentiellement le sommet du
premier segment. Profil longitudinal tres convexe, gibbeux vers l’avant.
La coloration métallique ne semble pas présenter une régularité absolue chez les
miles et les femelles du C. minimus. On rencontre des femelles 4 téte plus ou moins
bronzée et d’autres presque dépourvues de la bordure vert métallique de la marge
antérieure du prothorax.
66 exemplaires.
Loc. ‘Seychelles. Mahé: from near Morne Blane; country above Port Glaud,
500—1000 feet, XI. 1908; Cascade Hstate, ca. 800—1000 feet, 1908—9; Port Victoria,
XII. 1908; Long Island (swept from long grass) and Round Island, VIL. 1908 ; Anonyme
Island, I. 1909.”
27. Cybocephalus brevis, n. sp.
Oblongus, convexus, nitidissimus, ater ; antennis tarsisque rufo-testaceis, clava vix
infuscata ; capite apud marem et prothoracis antico margine viridi-aeneis, apud feminam
prothoracis antico margine stricte viridi-aeneo. Caput antice parum productum, breviter
truncatum, subparce tenuissimeque punctulatum. Prothorax parcissime vix perspicue
punctulatum; anticis angulis breviter, posticis late rotundatis. Elytra basi arcuata,
humeris obtusa et hebetata, apice apud marem conjunctim rotundata, apud feminam
conjunctim subacuminata, paulo longiora quam simul latiora, in disco vix perspicue ad
latera densius et validius punctulata, prope humeros oblique subcarinata et inter carinam
et lateralem marginem abrupte subinflexa. Metasternum dense punctulatum. Abdomen
pubescens. Long. 0,1—1,1 mill.
Oblong, plus d'une fois et demie plus long que large, convexe, trés brillant, noir ;
antennes et tarses roux testacé, massue des antennes a peine enfumée ; téte, bouche et
marge antérieure du prothorax vert métallique chez le male, marge antérieure du
prothorax plus ou moins étroitement bordée de vert métallique chez la femelle. Massue
des antennes assez bien marquée, subégale environ au tiers de la longueur totale de
l’antenne, dernier article un peu plus étroit et presqu’aussi long que le précédent. Téte trés
faiblement saillante en avant des naissances des antennes, britvement subtronquée au bord
antérieur, assez densément, mais & peine visiblement pointillée. Prothorax faiblement
arrondi au bord antérieur, fortement sinué vers ses extrémités, cdtés arqués, angles
antérieurs assez briévement arrondis, postérieurs tres largement; ponctuation trés espacée,
i peine visible sur le disque, plus serrée et un peu plus marquée sur les marges latérales ;
angles antérieurs et postérieurs et cdtés trés finement rebordés. Kcusson triangulaire,
environ deux fois plus large & la base que long. Elytres arrondis a la base lorsquils sont
vus de dessus, en angle obtus émoussé aux épaules lorsque celles-ci sont vues de face; arrondis
ensemble au sommet chez le male, subacuminés ensemble chez la femelle, un peu plus
longs que larges dans leur plus grande largeur, 4 peine visiblement pointillés sur le disque,
GROU VELLE—COLEOPTERA: NITIDULIDA, HETEROCERIDA 115
plus densément et plus visiblement sur les cdtés, trés finement rebordés sur les cOtés ;
régions humérales coupées obliquement par une faible caréne, plutot un pli se raccordant
en arc avec la base de lélytre, rejoignant presque le bord latéral et enfermant avec
eelui-ci un espace assez brusquement infléchi par rapport 4 la surface de l’élytre. Sillons
antennaires un peu convergents. Métasternum densément pointillé. Segments de
VYabdomen se développant dans un plan trés peu incliné par rapport au plan du
métasternum. Pubescence assez marquée sur le métasternum et l’abdomen. Lignes
fémorales des hanches postérieures divergentes rejoignant le sommet du premier segment
de l’abdomen en formant un angle obtus. Pattes relativement larges. Profil longitudinal
arqué, non gibbeux.
24 exemplaires,
Loc. “Seychelles: Mahé; from grass in cultivated country, about 1000 feet, XII.
1908 ; Cascade Estate, ca. 1000 feet, I. 1909; coastal plain at Anse aux Pins, I. 1909;
Anonyme Island, I. 1909 : Silhouette; Mare aux Cochons, [X.1908. Amirantes: Poivre
Island, on Scaevola koenagu, 10. X. 1905.”
28. Cybocephalus subpallipes, n. sp.
Breviter oblongus, convexissimus, nitidissimus, ater, vix cyaneo-micans; antennis
dilute testaceis; prothoracis lateribus strictissime, elytrorum humeris breve stricteque
ochraceo-marginatis; pedibus fulvo-testaceis, posticis femoribus plus minusve infuscatis.
Caput antice parum productum, subtruncatum, vix perspicue punctulatum. Prothorax
vix perspicue punctulatus; punctis ad latera paulo validioribus; angulis anticis sub-
breviter, posticis late rotundatis. Elytra basi arcuata, humeris obtusa et hebetata, apice
apud marem conjunctim rotundata, apud feminam conjunctim subacuminata, circiter tam
simul lata quam elongata, in disco vix perspicua, ad latera densius et validius punctulata,
prope humeros oblique vix carinato et inter carinam et lateralem marginem subabrupte
inflexo. Metasternum medio parce punctulatum. Abdomen parum pubescens. Long.
0,7—0,8 mill,
Oblong, presqu’une fois et demie plus long que large dans sa plus grande largeur, tres
convexe, trés brillant, noir avec un léger reflet bleudtre; antennes d’un testacé clair ;
marges latérales du prothorax et angles huméraux des élytres bordés de jaune testacé, les
premiers trés étroitement, les seconds étroitement et briévement ; dessous du corps noir
sauf la base de l’abdomen, celle-ci jaune testacé ; pattes d’un testacé un peu roussatre avec
les fémurs postérieurs plus ou moins enfumés. Massue des antennes égale environ au
tiers de la longueur totale de l’antenne, dernier article plus étroit que le précédent. Tete
médiocrement saillante en avant, subtronquée au bord antérieur, 4 peine visiblement
pointillée. Prothorax arrondi au bord antérieur, assez fortement sinué aux extrémités ;
cdtés faiblement arqués; angles antérieurs assez briévement arrondis, postérieurs trés
largement ; ponctuation presque nulle sur le disque, un peu plus nette sur la région des
angles antérieurs; angles antérieurs et postérieurs et cOtés tres finement rebordés.
Keusson triangulaire, plus de trois fois plus large & la base que long. Elytres arrondis a la
base lorsqwils sont vus de dessus, en angle obtus émoussé aux épaules lorsque celles-ci
sont vues de face, & peine moins longs que larges ensemble dans leur plus grande largeur,
116 PERCY SLADEN TRUST EXPEDITION
arrondis ensemble au sommet chez le male; 4 peine plus longs que larges ensemble et
subacuminés ensemble au sommet chez la femelle; & peine visiblement pointillés sur
le disque, un peu plus nettement sur les marges latérales ; région humérale coupée par un
pli arqué, se raccordant au bord latéral presqu’en angle droit 4 l’extrémité de la troncature
humérale, enfermant entre lui et le bord latéral un espace brusquement infléchi
)
par rapport 4 la surface de lélytre. Sillons antennaires modérément convergents.
Métasternum assez densément et finement pointillé. Segments de l’abdomen se
c=)
développant dans un plan assez incliné par rapport au plan du métasternum. Pattes
postérieures s’effagant au repos dans une fossette creusée sur le sommet du métasternum.
Pubescence de l’abdomen plus, développée que celle du métasternum. Lignes fémorales des
hanches postérieures divergentes rejoignant tangentiellement le sommet du premier
segment de l’abdomen. Profil longitudinal subdemicirculaire.
47 exemplaires.
Loc. “Seychelles. Silhouette: near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908;
Mare aux Cochons, from the plateau and forest near by, [X. 1908; low coconut-planted
country near coast, Pointe Etienne, IX. 1908. Mahé: high forest of Morne Blane and
Morne Pilot, 1908; country above Port Glaud, 500—1000 feet, XI. 1908; summit of
Mount Sebert, ca. 2000 feet, I. 1909; slopes of Morne Seychellois, 1500—2000 feet, II.
1909; Mare aux Cochons district, ca. 1500 feet, 1909; Cascade Estate, ca. 1000 feet;
coastal plain at Anse aux Pins, I. 1909 ; Anonyme Island, 1909 ; Long Island, VII. 1908.
le Marie Anne, XII. 1908.”
29. Cybocephalus, n. sp. ?
Un exemplaire & coloration probablement incomplete, a la téte et le dessous du corps
roux testacé. Il semble appartenir 4 une espéce nouvelle.
Loc. ‘Seychelles. Mahé: Mare aux Cochons district, 1000—2000 feet, I. 1909.”
Heteroceridee.
HETEROCERUS Fabricius, 1792, Ent. Syst. 1. p. 262.
Une seule espéce représente la famille des Heteroceridae; elle provient d’Aldabra ;
elle est identique & une espéce de Madagascar (H. vulpes Grouvelle). Il est probable que
de nouvelles découverts feront retrouver dans les nombreuses tiles de l’Océan Indien d’autres
formes soit de Inde, soit du continent Africain.
30. Heterocerus vulpes Grouvelle.
Heterocerus vulpes Grouvelle, 1906, Ann. Soc. Ent. Fr. xxv. p. 165, 168, t. 7, f. 11.
Loc. Aldabra: Takamaka, X. 1908 (Fryer). Madagascar.
No. VIII.—COLEOPTERA, PSELAPHIDA,
DE L ARCHIPEL DES SEYCHELLES.
Par A. RAFFRAY.
(Planche 10.)
(Communrqué PAR M. Le Proresseur J. STANLEY Garpiner, M.A., F.RS., F.LS.)
Lu le 1 mai 19138.
Monsieur Hugh Scott, Curator in Entomology 4 |’'Université de Cambridge, a bien
voulu me communiquer une série de Psélaphides qu'il a recueillis, au cours de la “ Percy
Sladen Trust Expedition to the Indian Ocean,” dont il faisait partie, dans ce petit archipel
des Seychelles perdu en plein Océan indien, & environ 3000 kilometres du Cap Comorin,
extréme pointe Sud du Bengale, 1900 kilométres de la céte d'Afrique et 1100 kilometres
~ du Cap d’Ambre, derniére terre septentrionale de la grande ile de Madagascar.
La faune entomologique de ces petites iles montagneuses, dont les sommets, de for-
mation granitique, sont encore couverts de foréts primitives, oti croit l’étrange palmier
Coco-de-Mer, doit nécessairement présenter un intérét que ne dément pas l’étude des
Psélaphides rapportés de cette exploration et qui sont assez nombreux pour révéler des
recherches habiles et patientes. Ce sont, du reste, les premiers Psélaphides qui aient
jamais été recueillis aux Seychelles.
Ces petits étres adéphages qui, par leur genre de vie le plus souvent endogée ou,
tout au moins, abritrée dans des repaires ombreux et humides, échappent, plus que tous
autres, 4 linfluence de homme, méritent un examen spécial qui permette sinon de
déterminer d'une facon précise, du moins de pressentir et d’esquisser leurs affinités
biogéographiques et fauniques.
On peut les diviser en trois catégories :
I. Les genres et sous-genre nouveaux et autochthones représentés par des especes
nouvelles.
II. Les genres déja connus et représentés par des espéces nouvelles.
III. Les genres déji connus et représentés par des especes également déja connues
d’autres pays.
I. les genres et sous-genre nouveaux et autochthones sont au nombre de 5,
représentés par 10 espéces.
SECOND SERIES—ZOOLOGY, VOL. XVI. 16
(L IBRARY'=c
W® Ge
118 PERCY SLADEN TRUST EXPEDITION
1°. Genre Hughia, de la tribu des Pyaxidicerint; 2 especes (phantasma, cari-
nata). Ce genre qui a, en exagération, les traits distinctifs de la tribu & laquelle il
appartient, possede, en méme temps, des caracteéres qui lui sont propres et en font un type
relativement isolé. C’est un élément tout a fait autochthone, qui n’est pas plus voisin des
formes asiatiques que des formes africaines.
2°, Genre Cerennea, de la méme tribu que le précédent: une seule espéce (brevi-
pennis) qui, 4 premiere vue, a tout a fait le facies étrange de Hughia phantasma, mais en
réalité, en est trés différente et se rapproche, au contraire, davantage de Pyaidicerus qui
est un genre exclusivement asiatique. Mais, en dépit de cette affinité morphologique, c’est
une forme isolée et nettement autochthone.
3°, Genre Apheloplectus, une seule espece (longicollis).
4°, Genre Omobathus, une seule espéce (elegans).
Tous les deux de la tribu des Huplectina et du groupe dont les Trimium européens
sont le type, avec, comme caractéristique commune, la massue antennaire formée d’un seul
article qui est trés gros.
Apheloplectus est une forme isolée, présentant des analogies lointaines avee Philiopsis
de Malaisie et de Madagascar et Trimiomorphus de Singapore.
Omobathus est extrémement voisin dAphilia qui comprend quelques espéces
spéciales aux Iles Malaises, de Sumatra & la Nouvelle-Guinée. En réalité la nouvelle
forme des Seychelles n’est qu'une Aphilia légerement transformée, mais autochthone.
5°. Sous-genre Sunorfoides nouveau sous-genre dérivé du genre Sunorfa dont il
nest qu'une légére modification. Le genre Sunorfu a 6 sternites seulement chez les ¢, et
le sous-genre Sunorfoides en a 7. Crest 1a, & mon avis, une modification morphologique
qui est plus que spécifique; mais, comme d’autre part, il y a identité compléte avec
Sunorfa, jestime qu'il est préférable de n’en faire qu’un sous-genre qui renferme 4 espéces
nouvelles (bicolor, mgripennis, picea, punctipenns). Ce qui prouve diailleurs que
Sunorfoides nest quune forme légerement modifiée par lisolement du genre Sunorfa, et
que cette transformation est peut-étre encore en voie de progression, c’est que le genre
Sunorfa lui-méme existe aux Seychelles ot il n’est représenté que par une seule espéce.
Nous retrouverons le genre Swnorfa dans la section suivante, et ce sera le moment de
discuter, pour le genre et son sous-genre, ses origines et ses affinités.
II. Genres déja connus, représentés par des espéces nouvelles.
1°. Genre Octonmucrus, de la tribu des Huplectini, une seule espéce : semupunctatus.
Ce genre est presque également réparti entre |'Indo-Malaisie (Siam, Sumatra, Bornéo,
Célébes) et l'Afrique tropicale (cétes occidentale et orientale, Zanzibar et Mont Kénia).
Semipunctatus est une forme légérement isolée et, par conséquent, autochthone.
2°. Genre Batrisodes, de la tribu des Batrisini, une seule espece : caudatus.
Ce genre qui compte plus de 180 espéces, est distribué trés inégalement entre
Europe (12 espéces), Amérique du Nord (45), Australie (19), Asie continentale et
RAFFRA Y—COLEOPTERA, PSELAPHIDAi DES SEYCHELLES 119
insulaire plus de 100 espéces, et enfin l'Afrique ot l’existence de ce genre n’a été constatée
que tout récemment, sur la céte orientale d'Afrique (3 especes).
Ce genre est trop nombreux et trop disséminé, pour étre trés homogéne, sans
cependant que des caractéres sérieux permettent de le diviser sans le pulvériser.
Lespéce des Seychelles appartient aux formes asiatiques, mais avec une modification
assez importante qui lisole de toutes les autres et en fait un élément décidément
autochthone.
3°. Genre Batramis, de la tribu des Brachyglutini, 2 especes : egregua, insularis.
Cest un genre presque exclusivement asiatique (32 espéces) répandu de Ceylan au
Nord de la Chine et a la Nouvelle-Guinée orientale, sauf 2 especes d’Australie et 1 espéce
qui semble égarée en Gréce, au Mont Olympe.
Ce genre, en tout cas, est non seulement inconnu en Afrique et 4 Madagascar, mais
on n’y connait, jusqu’a ce jour, aucune forme affine.
Les deux espéces des Seychelles, bien que spécifiquement autochthones, sont, en
réalité, des types asiatiques.
4°. Genre Triomicrus, également de la tribu des Brachyglutini, une seule espéce
(Seychellarum).
Ce genre trés tranché et essentiellement asiatique, ne comptait que 5 espéces de
la Chine orientale et du Japon. L’espéce des Seychelles n’a rien d’anormal, c'est un type
aslatique. .
5°. Genre Sunorfa, de la tribu des Tychini, avec une seule espéce nouvelle
( fasciculata).
Le genre Sunorfa, qui comptait déja 19 espéces, dont 16 de la Nouvelle-Guinée, 1 de
Ile Vanikoro, 1 de Singapore, 1 de Sumatra, est un type Océano-Malais, dautant plus que
sur 16 espéces de Nouvelle-Guinée, 15 viennent de la Cote Sud-est qui se relie & l’Océanie
mélanésienne et aux Nouvelles Hébrides ot l’on rencontre l’espéce la plus orientale de
Sunorfa.
Le genre Sunorfa et son sous-genre Swnorfoides, que nous venons de voir dans la
section précédente, dans laquelle rentrent les formes génériques et sous-génériques
autochthones, sont done, en réalité, tous les deux des types malais & tendance océanienne
prépondérante.
La présence simultanée, dans ce petit archipel des Seychelles, de ces deux formes
génériques qui ne different entre elles que la présence ou l’absence, dans le seul sexe mile,
dun septiéme sternite (ou segment ventral) et dérivant évidemment l'une de lautre,
semblent en voie de transformation inachevée, est un cas curieux pour l'étude de la
biogéographie entomologique.
Sunorfa et Sunorfordes appartiennent & la premiére section de la tribu des Tychini
assez pauvrement représentée dans les régions chaudes de I Amérique et de |’Asie, mais fait
complétement défaut 4 Madagascar et n’a été constatée que tout récemment en Afrique,
sur la cote orientale, 4 Shimoni, au Sud de Mombasa, par MM. Alluaud et Jeannel qui y
ont découvert un genre nouveau peu éloigné de Sunorfa et qui, en tout cas, est une forme
16—2
120 PERCY SLADEN TRUST EXPEDITION —
essentiellement asiatique, ses affinités immédiates étant avec le genre Tanypleurus qui est,
jusqu’’ présent, localisé a la région du détroit de Malacca.
Ces deux découvertes, en quelque sorte simultanées, sur la cédte orientale d'Afrique
et dans l’archipel des Seychelles, qui lui fait face, de types purement malais, voisins les
uns des autres, qui étaient, jusqu’alors, complétement inconnus dans cette partie du
monde, constituent un fait 4 enregistrer.
6°. Genre Apoplectus, de la 1° section de la tribu des Tychini: une seule espéce
(obesus).
Ce genre, qui n'est pas trés lointain de Sunorfa, ne comptait encore que 5 espéces,
toutes de la Nouvelle-Guinée septentrionale, mais surtout orientale; ¢est donc une forme
presque plus océanienne qu’asiatique, et l’espece des Seychelles n’est séparée des autres
que par des caractéres normalement spécifiques.
III. Genre et espéce déja connus.
Un seul genre Thesiastes, de la tribu des Huplectini, et une seule espéce, cordicollis
Raffr.
Le genre Thesvastes qui comprend une quinzaine d’espéces un peu hétérogénes, qu'il
me semble, cependant, difficile de dissocier génériquement, est disséminé depuis |’Amérique
du Nord et le Chili, en passant par [Afrique orientale et Singapore, jusqu’a la Nouvelle-
Guinée.
Lrespéce qui a été trouvée, en assez grand nombre, dans les trois tiles de Mahé,
Silhouette et Praslin, est absolument identique 4 celle que j’ai découverte, autrefois,
Zanzibar. C'est une espéce africaine, mais qui appartient & un type cosmopolite.
En résumé les Psélaphides connus de l’Archipel des Seychelles se repartissent comme
suit au point de vue biogéographique.
Genres exclusivement autochthones (Hughia, Cerennea, Apheloplectus) ds sok WS.
Genre autochthone, mais dérivant de formes asiatiques légérement modifiées (Omo-
bathus) BAO es ae ws he She bbs sof 1
Genre 4 trés grande prédominance asiatique (Batraavs) : a!
Genre exclusivement asiatique (Zriomucrus) ... : 1
Genres 4 trés grande prédominance océano-malaise (Sunorfa, Sunorfoides) ... 2
Genre exclusivement océano-malais (A poplectus) — oe “3 ne seen yal
Genre indo-africain (Octomicrus) ... sie ae she site ae ee, Tal
Genres plus ou moins cosmopolites (Thesvastes, Batrisodes) ... ze Ap Sey oe
Quant aux 17 espéces, elles se repartissent comme suit: Espéces exclusivement
autochthones (Hughia phantasma et carimata, Cerennea brevipennis, Aphelo-
plectus longicollis) ... ee we ‘ee ane au sie nee Sse
Espéces & type asiatique plus ou moins pur ou modifié (Omobathus elegans, Batri-
sodes caudatus, Batraais egregia, Triomacrus Seychellensis) Ht Uae
Espéces & type océano-malais plus ou moins pur ou modifié (Batrams imsularis,
Sunorfa fasciculata, Sunorfoides nigripennis, bicolor, picea, punctipennis,
Apoplectus obesus) ... eh se a ee Am Bae h//
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 121
Espéce & type indo-africaine (Octomicrus semipunctatus) She ase ae ae vy Dh
Espéce africaine (Thesvastes cordicollas) ... abe sat san Be the Str Aaa
C’est done vers les terres les plus orientales de |’Asie continentale et insulaire et
jusqu’aux rives de locéan pacifique, qu’avee 6 genres et sous-genre sur 12, et 11 espéces
sur 17, dérivant plus ou moins immédiatement de types asiatiques et océano-malais, il
faut aller chercher les affinités morphologiques des Psélaphides qui habitent le petit
Archipel des Seychelles.
D’aprés le récit trés intéressant de ses chasses entomologiques aux Seychelles, que
Mr Hugh Scott a publié dans les “Transactions of the Linnean Society of London ”
(vol. xiv. 1910, p. 21), il a surtout exploré les hautes foréts indigenes et, d’apreés les
notes manuscrites qu'il a bien voulu me donner, tous les Psélaphides des Seychelles ont
été récoltés dans ces foréts indigénes, entre 800 et 2400 pieds daltitude; méme le
Thesiastes cordicollis que j’avais découvert dans l'ile de Zanzibar, au niveau de la mer, a
été rencontré, aux Seychelles, jusqu’a 1500 pieds daltitude.
Il faut savoir grand gré & Mr Hugh Scott qui s’occupait d’Entomologie générale,
d’avoir recueilli ces coléoptéres liliputiens, d’autant plus qu'il les recherchait directement,
sans employer la méthode plus rapide et plus commode du tamisage des détritus végétaux
et de la terre quils recouvrent, procédé qui lui edt fourni d’ailleurs une récolte bien plus
abondante des insectes endogés.
Aprés avoir passé en revue les éléments constitutifs de la faune psélaphique des
Seychelles et constaté quils sont, pour les deux tiers, intimement liés 4 des types
asiatiques, il est nécessaire d’en faire ressortir les lacunes, constatation négative qui
contribue, avec les formes autochthones, 4 donner 4 cette faune psélaphique un caractére
tres spécial.
Le grand bloe des anciens Bryaxis, qui constitue la majorité de la tribu des
Brachyglutini, répandu dans le monde entier avec les modifications génériques des
Brachygluta, Reichenbachia, Rybaxis et leurs nombreux dérivés, fait complétement défaut
aux Seychelles qui est, je crois, le seul pays ou, la présence de Psélaphides étant constatée,
il ne soit pas représenté; car les genres Batraais et Triomicrus, qui appartiennent a la
méme tribu, sont des formes excentriques et de transition avec les tribus qui précédent ou
qui suivent.
La tribu des Pselaphini, qui est répandue dans toutes les parties du globe, excepté a
Madagascar, manque également aux Seychelles.
Il en est de méme de celle des Ctenistini qui est également trés cosmopolite; mais
cela est moins surprenant, les Ctenzstini affectionnant, en général, les régions basses et, en
montagne, les prairies humides; tous les Psélaphides connus des Seychelles proviennent
des foréts plus ou moins élevées.
La tribu des Tyrini qui, dans toutes les régions chaudes du globe, jusqu’en Europe
tempérée et en Nouvelle-Zélande, renferme tant de belles especes, manque totalement
aux Seychelles.
Enfin la sous-famille des Clavigeride, dont le centre géographique est certainement
122 PERCY SLADEN TRUST EXPEDITION
Madagascar et qui tous, plus ou moins, sont myrmécobies, n’ont pas de représentants aux
Seychelles, bien qu’on y ait trouvé des Psélaphides dans les Fourmiliéres.
On ne peut malheureusement établir de point de comparaison ni avec les iles Comores
ni avec les iles Mascareignes d’ou, en dépit de leurs cimes parfois trés élevées et boisées,
comme 2 la Réunion, on ne connait aucun Psélaphide.
I] nen est pas de méme avec l’ile de Madagascar qui est d’ailleurs la grande terre la
plus voisine de l’Archipel de Seychelles. Bien que la faune psélaphique malgache soit
trés incompléetement connue, quelques Psélaphides y ont été recueillis par Goudot et
Coquerel, il y a longtemps déja, puis par moi-méme, par MM. Sikora et Perrier et enfin
par mon ami M. Alluaud. On y compte maintenant 35 genres, dont 28 autochthones, et
64 espéces, toutes autochthones. C'est bien peu si l’on songe a |’étendue de cette ile,
mais cest déja presque suffisant pour pouvoir présumer que la grande ile Malgache a une
faune psélaphique trés spéciale.
Or il n’y a pas un seul genre qui soit commun aux Seychelles et 4 Madagascar. Au
point de vue des Psélaphides, ces deux terres quoique isolées, aussi bien l’une que l'autre,
fauniquement de |’Afrique, continent le plus voisin qui semblerait devoir étre leur grand
pourvoyeur, n’ont entre elles aucun point de contact. En ce qui concerne les Psélaphides,
Madagascar apparait isolé, les Seychelles au contraire étroitement asiatiques.
Tribu Pyxidicerini Raffray, Ann. Soc. Ent. France, Ixxii. 1903, p. 504.
Genre Hucuta, nov. gen. (PI. 10, figs. 1, 2).
Oblonga, parum convexa. Caput magnum, transversum, prothorace latius, antice
truncatum et in fossa palpari totum excavatum, trilobatum, lobo mediano angusto,
antennas ferente. Oculi mediocres, parum prominentes. Antenne: basi approximate,
11-articulate ; articulis duobus primis magnis, sequentibus minoribus, monilibus, clava
magna, biarticulata. Prothorax plus minusve cordatus, foveatus et sulcatus. Elytra
brevia, subtriangularia vel subquadrata, plus minusve carinata. Abdomen late mar-
ginatum, segmentis dorsalibus subzequalibus, ventralibus ¢ sex, ¢ septem, septimo trans-
versim rhomboidali et longitudinaliter carinato. Pedes mediocres, sat elongati; coxis
omnibus approximatis; femoribus leviter incrassatis; tarsis elongatis et gracilibus,
articulis 1° minuto, 2 supra inconspicuo, infra occulto, 3 elongato, subcylindrico, leviter
arcuato ; ungue unico.
Ce nouveau genre est certainement un des plus étranges dans une tribu qui présente
elle-méme une conformation extraordinaire.
La téte est grande, transversale, composée de trois parties, la médiane étroite, un peu
relevée, séparée des deux latérales par un sillon allant du cou jusqu’a l’extrémité du front
et formant, en avant, un tubercule sur lequel sinsérent les antennes, contigués 4 leur
base; les parties latérales, représentant les joues, sont beaucoup plus grandes, un peu
surbaissées, brusquement et sinueusement tronquées en avant, creusées d'une profonde
fossette dans laquelle disparaissent les palpes dont, au repos, les articles sont repliés les
uns sur les autres; l’angle antéro-externe de ces joues est trés pointu, langle postérieur,
c'est & dire la tempe, est au contraire trés arrondi. Ces insectes, fort rares, étant presque
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 123
tous des exemplaires uniques, je n’ai pu en sacrifier pour faire une préparation dans le
baume, sans laquelle il est impossible de voir les palpes dépliés, 4 moins de les examiner
vivants. Mais je pense que ces palpes sont, avec de légéres variantes, construits comme
dans toute cette tribu. La face inférieure de la téte est grande, un peu convexe et ne
laisse pas apercevoir les organes buccaux qu'elle recouvre. Les antennes sont relative-
ment grandes, les deux premiers articles robustes, la massue de deux articles, beaucoup
plus forte chez les $ que chez les 9, les articles intermédiaires petits, moniliformes.
Le prothorax est plus ou moins cordiforme, avec trois fossettes et un sillon variables.
Les élytres sont trés variables, courts, presque triangulaires (phantasma), presque carrés
(carmmata), ils portent toujours des carénes et les bords sont planés et plus ou moins
tranchants. L’abdomen est trés largement rebordé sur les cétés; les 3 ont 7 segments
ventraux, le 5° trés étroit, le 6°™° un peu plus grand, fortement et anguleusement
entaillé au milieu, pour enchdsser le 7°™® qui est en losange transversal, avec une fine
earéne longitudinale. Les pieds sont moyens et normalement conformés ; les tarses ont
3 articles, mais, comme pour tous les genres de cette tribu, les 1°7 et 3°™° articles sont
seuls visibles, sans préparation transparente, le 2°™® étant trés petit et occulté en
dessous, entre le 1% et le 3°™°, ce dernier est tres long et semble, & lui seul, former
tout le tarse.
Dans tous les autres genres de cette tribu, la fossette palpaire, qui est toujours
creusée aux dépens de la joue supérieure, est & découvert et plus ou moins libre ou
rebordée seulement sur les cétés. Dans Hughia, au contraire, la partie antérieure de la
fossette palpaire est bien & découvert, mais elle s’enfonce, comme dans une caverne creusée
dans la joue supérieure qui la recouvre en partie; ce qui rend du reste l’extraction du palpe
extrémement difficile.
Ce nouveau genre devra venir en téte de la tribu, avant le genre Pyazdicerus qui en
est le type et qui habite exclusivement les iles malaises.
Je suis heureux de dédier ce genre remarquable & Mr Hugh Scott, Curator in
Entomology 4 l’université de Cambridge, auquel la science doit cette belle découverte.
1. Hughia phantasma, nov. sp. (Pl. 10, fig. 1).
Castanea, antennis pedibusque dilute ferrugineis, parce et brevissime pubescens.
Capitis lobo mediano punctato. Antennarum articulis 1° magno, elongato, 2 minore,
quadrato, sequentibus minutis, monilibus, clava in utroque sexu diversa. Prothorax
subcordatus, valde punctatus, foveis tribus quarum lateralibus majoribus, sulco longi-
tudinali obsoletissimo. Elytra brevia, antice angustata, minus punctata, basi bifoveata,
obtuse bicarinata, lateribus deplanata. Abdomen elytris longius, postice leviter amplia-
tum, apice obtusum. Pedes sat elongati; femoribus leviter incrassatis; tibiis anticis
perparum sinuatis, intermediis subrectis, extus leviter ampliatis, posticis leviter arcuatis
et extus paululum ampliatis.
g. Antennarum articulis 1° crassiore, 10 maximo, globoso, intus basi dentato et
medio penicillato, 11 multo minore, subovato, obtuse acuminato. Segmentis ventralibus
5 angusto, arcuato, 6 majore, medio margine postica valde angulatim emarginata, 7 magno,
124 PERCY SLADEN TRUST EXPEDITION
rhomboidali, tenuiter carinato. Femoribus intermediis juxta basin infra spina recurva
armatis; tibiis posticis apice calcaratis.
g. Antennarum articulis 1 graciliore, cylindrico, 10 magno, ovato-truncato, 11 majore,
ovato, apice obtuse acuminato. Elytra perparum breviora.
Long. 1,80 mill.
Le lobe médian de la téte est séparé des lobes latéraux (les joues) par un profond
sillon marqué, un peu en arriére, d’une grande fossette. Les cdtés des élytres sont assez
fortement obliques de l’épaule au-dela du milieu, ot ils forment un angle excessivement
obtus et deviennent ensuite droits, jusqu’a lextrémité. La pubescence est rare et courte,
mais trés légérement écailleuse. Les fossettes 4 la base des élytres sont profondes et vont
en sallongeant, pour former, entre les carénes, un sillon large et un peu vague qui va,
comme les carénes, s’atténuant vers l’extrémité.
Loc. Mahé; 4, forét indigéne au sommet de la Montagne Anse Major, environ 2000
pieds, II. 1909; 9, forét haute et tres humide, entre Trois Fréres et Morne Seychellois,
environ 1500 a 2000 pieds, XII. 1908.
2. Hughia carimata, nov. sp. (Pl. 10, fig. 2).
Preecedenti valde affinis, picea, antennis, elytris pedibusque obscure rubris, pube
dispersa, vix perspicua. Caput valde transversum, temporibus minus rotundatis, magis
obliquis, totum obsolete et disperse punctatum; lobo mediano breviore, latiore, leviter
triangulari.
Antennarum articulis 10 paululum minore, globoso et simplice, 11 eadem longitudine,
longiore, ovato, apice acuminato. Prothorax paulo brevior, lateribus minus rotundatis,
punctatus, lateribus late foveatus, medio, ante basin, late depressus et fundo bifoveatus,
sulco dorsali obsoleto. Elytra quadrata, disperse punctulata, lateribus leviter rotundata,
fortiter bicarinata. Abdomen elytris vix longius, postice rotundatum ; segmentis dorsali-
bus disco perparum deplanatis, 1° basi transversim impresso ; ventralibus 3 et 4 margine
postica triangulatim impressis, 5 angusto, 6 medio angusto et margine postica angulatim
emarginato, 7 valde transverso, rhomboidali, carinula longitudinali leviter asymetrica
(Pl. 10, fig. 2). Femoribus sat crassis; tibiis omnibus, pone medium, extus, leviter
ampliatis ; posticis apice breviter calcaratis g. 9. Incognita. Long. 1,90 mill.
Cette espece, dont le ¢ seul est connu, différe de la précédente par la téte encore
plus transversale, avec les tempes moins arrondies, plus obliques, le lobe médian plus court,
plus large au sommet qu’a la base; le 10°™® article des antennes est bri¢vement ovalaire,
simple, le 11°™° est aussi gros que lui, plus long et acuminé ; le prothorax est moins long,
les cdtés sont d’abord & peu prés droits jusqu’au milieu, ott ils forment un angle extréme-
ment obtus et, de 14, obliques jusqu’’ la base ; sur les cotés il y a une immense fossette et,
au milieu, une grande dépression presque bilobée en avant, avec une double fossette peu
marquée dans le fond; les élytres sont & peu pres carrés, avec les cétés légérement
arrondis et moins planés, les deux carénes sont beaucoup plus accentuées, sans étre
tranchantes, et atteignent l’extrémité ; les segments dorsaux de l’'abdomen sont convexes,
trés légérement aplatis sur le disque et aussi sur les cotés.
Loc. Silhouette: forét au-dessus de la Mare aux Cochons, VIII-—IX. 1908.
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 125
Genre CERENNEA, nov. gen. (PI. 10, fig. 3).
Oblonga. Caput magnum, valde transversum, trilobatum, prothorace latius ; antice
in fossa palpali excavatum, ista fossa insuper, juxta lobum medianum angustum et
antennas ferentem, aperta. Oculi mediocres. Antennze novem-articulatee; articulis
duobus primis magnis, sequentibus monilibus, clava magna, biarticulata. Prothorax
latitudine sua longior, subcordatus, quinque-impressus. Elytra brevia, basi attenuata,
humeris élevata et dentata, bicarinata. Abdomen magnum, late marginatum ; segmentis
dorsalibus subzequalibus, ventralibus 2, 3, 4 magnis, 5 multo minore, 6 (ultimo) transverso
triangulari $? Pedes sat crassi et minus elongati; tarsis crassioribus.
Ce nouveau genre qui appartient 4 la méme tribu que Hughia, a surtout le facies de
Hughia phantasma ; mais il en est trés distinct, d’abord par le nombre des articles des
antennes qui est de 9 seulement, au lieu de 11, et ensuite par la téte: la cavité palpaire,
qui est encore située sur la partie supérieure de la téte, n’est plus fermée et recouverte
que sur les cdtés, elle est ouverte en dessus, le long du lobe médian ; en d’autres termes
cette fossette palpaire chez Cerennea n’est recouverte que partiellement par un rebord
du cdoté de la joue, tandis que, dans Hughia, elle est recouverte entitrement par
toute la joue.
Cette conformation, malgré une analogie trompeuse de facies, éloigne Cerennea de
Hughia, pour le rapprocher, au contraire, de Py«idicerus qui a également la cavité
palpaire partiellement découverte; mais chez Pyaidicerus, le rebord de la joue est
réduit presque & une simple caréne, en sorte que la cavité palpaire, au lieu d’étre
réellement recouverte, est plutdt simplement limitée, extérieurement, par ce rebord
de la joue.
Au point de vue morphologique Hughia et Cerennea sont fort imtéressants, car
ils représentent les deux premiers termes de la série: cavité palpaire entiérement
recouverte, Hughia; cavité palpaire partiellement recouverte, Cerennea; le troisiéme
terme sera Pyxidicerus, cavité palpaire simplement limitée, en dehors, par un rebord
de la joue, et le quatriéme et dernier terme: cavité palpaire entiérement ouverte et sans
rebord, sera représenté par tous les autres genres connus de la tribu.
Cerennea différe, en outre, de Pyxidicerus par le nombre des articles des antennes,
les élytres beaucoup plus courts, carénés, les épaules saillantes et dentées. Les palpes
repliés dans la fossette, n’ont pu étre examinés ; l’exemplaire unique était déja dans un
état trop défectueux pour que je courre le risque de le briser complétement, en tentant
Yextraction du palpe.
3. Cerennea brevipennis, nov. sp. (Pl. 10, fig. 3).
Obscure fulva, vix perspicue pubescens. Caput minute et disperse punctatum ;
lateribus obliquis, angulis anticis acutis, posticis leviter obtusis; lobo mediano antice
rotundato, longitudinaliter bisulcato, postice angustiore et, juxta collum, sulcis duobus
obliquis angulato. Oculi parum conspicui, medio siti. Antennz breves et crasse,
articulis 1 magno, elongato, intus leviter rotundato, angulo externo acuto, 2 magno,
globoso, 3—6 minutis, monilibus, 7 minuto, transverso, 8 magno, transversim trapezoidali,
SECOND SERIES—ZOOLOGY, VOL. XVI. 17
126 PERCY SLADEN TRUST EXPEDITION
9 (ultimo) magno, breviter conico. Prothorax fortius punctatus, capite angustior et
latitudine sua longior, subcordatus, utrinque lateribus depressione magna, subovata, disco
fovea magna, oblonga, ante basin foveis duabus subovatis approximatis. Elytra brevia, ad
basin valde attenuata, lateribus postice rotundata et deplanata, basi profunde bifoveata,
earinis duabus rotundatis, humeris obliquis, prominulis et dentatis. Abdomen magnum,
segmentis dorsalibus duobus primis basi media transversim, minute impressis ; ventralibus
2, 3, 4, medio tuberculo minuto ornatis. Pedes sat crassi; femoribus anticis et intermediis
erassioribus; tibiis omnibus extus medio rotundato-incrassatis, anticis et intermediis
perparum sinuatis, posticis: leviter arcuatis.
Long. 1,20 millim.
Le rebord latéral qui recouvre partiellement la cavité palpaire est plus large que
le lobe médian, et l’entaille qui découvre cette cavité ne dépasse que de trés peu le milieu,
car la cavité ne s’étend pas sur toute la téte. Le prothorax est arrondi en avant, avec les
angles antérieurs marqués, mais obtus et émoussés; les cOtés sont droits Jusqu’au milieu
et, ensuite, obliques jusqu’d la base. Les cdtés des élytres sont déprimés; les carénes
soblitérent vers l’extrémité, l’externe part de la dent humérale, l interne atteint & peine
lextrémité et s’arrondit un peu en dedans.
Tl est impossible de déterminer exactement le sexe de lunique exemplaire; il n’y
a que 6 seoments ventraux, caractére commun aux deux sexes dans la plupart des genres
de la tribu des Pyaidicerim. Les tubercules des 2°™*, 3°™e et 4°™° seoments ne sont pas
un indice sexuel de grande valeur et les tibias sont tous mutiques 4 l’extrémité, ce qui
permet de croive que cet exemplaire est une femelle.
Loc. Mahé: forét haute et trés humide, par environ 1500 & 2000 pieds, entre Trois
Freres et Morne Seychellois, XII. 1908.
Tribu Euplectini Rafiray, Rev. d’Ent., 1890, pp. 82, 91.
Genre APHELOPLECTUS, nov. gen. (PI. 10, figs. 4, 5).
Elongatus, subparallelus, parum convexus. Caput antice valde attenuatum ; fronte
recte truncata, impressa. Oculi medio siti, prominentes. Palpi mediocres; articulis
2 elongato, gracili, apice abrupte clavato, 3 minuto, globoso, 4 magno, pyriformi, extus
leviter sinuato, acuminato. Antenne parum elongate, articulis 2 primo majore, 3—9
minutis, monilibus, 10 transverso, 11 maximo, subconico, acuminato. Prothorax capite
latior et multo longior, suboblongo-ovatus, postice attenuatus et lateribus, ante basin,
fovea laterali leviter sinuatus, trifoveatus. Hlytra elongata, subparallela, basi bifoveata ;
sulco dorsali abbreviato. Abdomen marginatum; segmentis dorsalibus zqualibus,
ventralibus g et 2 sex, quorum 4 et 5 minoribus, 6 transverso, leviter triangulari. Pedes
validi; femoribus crassis; tibiis extus medio crassioribus; tarsorum articulis 2 majore,
leviter clavato, 3 minore, cylindrico; ungue unico.
La téte est large, en ovale transversal et convexe en arriére, avec les tempes
arrondies, assez brusquement rétrécie en avant des yeux; le front est tronqué carrément,
avec les angles antérieurs un peu noueux ; toute la partie antérieure et rétrécie de la téte
est impressionnée, et les cdtés sont un peu carénés. Les yeux sont assez proéminents,
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 127
situés vers le milieu, en avant du vertex. Le 2°™° article des palpes (PI. 10, fig. 5) est
relativement long, tres gréle et subitement en massue arrondie 4 l’extrémité, le 3°™° est de
méme grosseur que la massue du 2°™*, le 4°™° est gros, britvement ovoide, 4 coté interne
réguliérement arrondi, le cété externe un peu sinué, il est trés acuminé au sommet avec
un petit appendice terminal. Les antennes sont relativement courtes, les deux premiers
articles robustes, le 2°™¢ plus gros que le premier, les suivants sont petits, moniliformes et
serrés les uns contre les autres, le 10°™° transversal, sans étre plus grand que les précédents,
le dernier trés gros. Le prothorax est grand, bien plus long que large, arrondi en avant,
rétréci en arriére et légérement sinué sur les cétés & la hauteur de la fossette latérale ; il
y a trois fossettes libres. Les élytres sont assez longs, en quadrilatére assez régulier,
avec deux fossettes basales et un sillon dorsal raccourci. L'abdomen est un peu plus
étroit que les élytres, rebordé, les tergites sont égaux entre eux; les sternites, au nombre
de six dans les deux sexes, vont en décroissant du 2°™* au 4&™e, le 5¢me est trés étroit, le
6eme et dernier trés transversal, faiblement triangulaire. Les pieds sont robustes, les
fémurs plus ou moins renflés, les tibias épaissis au milieu en dehors. Le 2®™° article des
tarses est long et légérement en massue, le 3°™¢ plus court, cylindrique.
Ce nouveau genre qui ne renferme qu'une seule espéce, présente des affinités plus ou
moins Jointaines avec plusieurs genres: la téte ressemble & celle de Philiopsis, les
antennes dont le 2°™¢ article est plus gros que le 1°, le rapprocheraient de Trimiomorphus
dont il a la forme allongée et le prothorax & fossettes libres; mais le prothorax est
beaucoup plus grand, plus allongé que dans aucun autre genre voisin. Crest en réalité
une forme assez isolée.
4. Apheloplectus longicollis, nov. sp. (PI. 10, figs. 4, 5).
Totus rufus, vix perspicue pubescens. Caput postice transversim ovatum, antice
attenuatum et impressum. Antenne pubescentes, articulis 1 subquadrato, 2 majore,
subgloboso, 3—9 minutis, monilibus, 10 minuto, transverso, 11 magno, subconico.
Prothorax magnus, elongatus, antice rotundatus, postice attenuatus, lateribus pone
medium leviter sinuatus, disco longitudinaliter perparum deplanatus; foveis lateralibus
magnis, mediana minore rotundata. Elytra elongata, lateribus parallela, basi bifoveata,
sulco dorsali ante medium abbreviato. Femoribus leviter incrassatis, tibiis extus medio
crassioribus.
g. Femoribus anticis multo crassioribus et tibiis extus medio magis incrassatis.
Long. 0,80 millim.
Loc. Tle Félicité: dans la forét, XII. 1908.
Genre OmopatHus, nov. gen. (Pl. 10, fig. 6).
Elongatus, antice attenuatus, convexus. Caput sat elongatum, postice transversum,
convexum, temporibus rotundatum, antice attenuatum, impressum ; fronte recte truncata,
angulis anticis leviter nodosis et lateribus obtuse carinatis. Oculi medio siti, prominentes.
Palpi mediocres ; articulis 2 parum elongato, apice clavato, 3 minuto, globoso, 4 mediocri,
ovato. Antenne sat elongate, articulis duobus primis majoribus, eequalibus, 1 cylindrico,
17—2
128 PERCY SLADEN TRUST EXPEDITION
2 leviter ovato, 3—9 minutis, monilibus, 9 leviter transverso, 10 paululum mayore,
transverso, 11 magno, ovato, acuminato. Prothorax elongato-cordatus, foveis tribus
liberis. Elytra magna, subovata, humeris elevatis et angulatis, basin versus oblique
deplanatis et paululum concavis ; fovea suturali magna; stria suturali tenuissima, dorsali
nulla. Abdomen elytris brevius, marginatum, basi angustius, postice rotundatum ;
segmento dorsali 1° majore; segmentis ventralibus 1° deplanato, coxis posticis breviore,
2 magno, 3, 4, 5 brevissimis, 6 (ultimo) magno, subtriangulari, apice obtuso. Metasternum
magnum, convexum. Pedes elongati, parum crassi; tarsorum articulo 2 longiore, leviter
clavato, 3 minore, cylindrico ; ungue unico.
Ce nouveau genre est curieux. la téte ressemble beaucoup a celle du genre
précédent, mais elle est plus allongée. Les palpes sont plus petits et le dernier article
moins gros. Les antennes sont plus longues, les deux premiers articles gros et égaux
entre eux, les suivants trés petits, les 9°™° et 10°™° vont en s’élargissant graduellement et
sont transversaux, en sorte que la massue n’est plus aussi exclusivement formée par le seul
dernier article qui est, dailleurs, encore trés gros. Le prothorax est également allongé,
cordiforme, avec trois fossettes libres. Les élytres sont grands, un peu ovoides, & peine
plus rétrécis en avant qu’en arriére ; les épaules sont proéminentes, formant un petit cone
pointu au sommet, trés évasé a la base, et dont le cdté tourné vers la base des élytres forme
un pan coupé triangulaire, qui est non seulement aplati, mais un peu concave ; assez loin
de la base, le long de la suture, il y a, de chaque cété, une grande fossette de laquelle part
une trés fine strie suturale ; il n’y a pas de strie dorsale. L’abdomen est plus court que
les élytres, et plus étroit & sa base, il sélargit en arriére et s'arrondit, sa marge est
rebordée et le premier segment dorsal est beaucoup plus grand. II n’y a que 6 segments
ventraux au seul exemplaire connu qui me parait étre certaimement une femelle ; le 1°" est
plat, visible, mais plus court que les hanches postérieures qui sont trés coniques et
contigués, le 2°™° est tres grand, les 3, 4, 5 extrémement courts, le 6°™° (dernier) grand,
triangulaire, obtus 4 l’extrémité. Les pieds sont longs, avec les fémurs et tibias peu
renflés.
Ce genre est trés voisin d’Alphilia Reitt., dont il a le faciés; chez Aphilia, les
épaules, si on les examine avec soin et sous différents angles, montrent une tendance
a devenir saillantes et coniques. Omobathus differe par la téte plus grande, plus atténuée
en avant, avec le front plus largement impressionné; le prothorax plus cordiforme et
trifovéolé ; la conformation insolite des épaules et Vabdomen arrondi, au lieu d’étre
acuminé en arriere.
5. Omobathus elegans, nov. sp. (Pl. 10, fig. 6).
Totus rufus, nitidus, glaber, setis attamen aliquot elongatis in elytris, abdomine et
pedibus. Antennarum articulo ultimo pubescente. Prothoracis foveis lateralibus
majoribus, mediana minuta, punctiformi. Tibiis rectis, leviter extus crassioribus, ?.
Long. 0,90 millim.
Loc. Mahé: Cascade Estate, 800 pieds et au-dessus.
RAFFRAY—COLEOPTERA, PSELAPHIDA! DES SEYCHELLES 129
Genre THESIASTES Casey, Col. Not. in Ann. N. York Se. Ac., 1893, pp. 444, 457.
6. Thesiastes cordicollis, Rattray.
Thesvastes cordicollis Raftray, Rev. d’Ent., 1. 1882, p. 81.
Les 21 exemplaires de |’Archipel des Seychelles sont absolument identiques 4 ceux
que jai découverts, autrefois, 4 Zanzibar et qui constituent le type de l’espéce.
Loc. Silhouette: foréts indigénes prés de Mont Pot-i-eau, environ 1500 pieds, VIII.
1908. Mahé: Cascade Estate, environ 800 pieds ou au-dessus; dans la méme localité,
dans une souche d’arbre pourrie, dans une fourmiliére de Camponotus foraminosus Forel,
subsp. grandidiert Forel (déterm. par A. Forel) (un seul exemplaire qui ne se trouvait
probablement qu’accidentellement parmi des fourmis, car cette espéce ne semble pas
ordinairement myrmécophile ni 4 Zanzibar, ni aux Seychelles) ; forét haute et trés humide,
1500 & 2000 pieds environ, entre Trois Fréres et Morne Seychellois, XII. 1908 ; Long
Island, petit lot cultivé prés de la Grande Ile, VII. 1908. Praslin : Cdtes d’Or Estate,
forét de palmier coco-de-mer (Lodoicea Seychellarwm), dans la Vallée de Mai, XI. 1908.
Cet insecte semble répandu dans |’Archipel depuis les cultures du petit tlot de Long
Island, jusqu’aux hautes foréts autochthones.
Genre Octomicrus Schaufuss, Psel. Siams, 1877, p. 14.
7. Octomicrus semapunctatus, nov. spec. (PI. 10, fig. 7).
Elongatus, parallelus, deplanatus, totus rufus, pube inconspicua ; capite et
prothorace minute, elytris et abdomine vix perspicue punctatis. Caput subquadratum,
antice leviter attenuatum ; fronte recte truncata et profunde transversim foveata; inter
oculos foveis duabus sulcis parallelis, obsoletis cum fovea frontali junctis. Antenne parum
elongatze, crassx ; articulis duobus primis majoribus, 3—9 minutis, monilibus, 10 paululum
majore, transverso, 11 mediocri, breviter ovato. Prothorax capiti longitudine et latitudine
subzequalis, cordatus, lateribus, pone medium sinuatus et obtuse dentatus; foveis
lateralibus magnis, mediana rotundata et dorsali oblonga, plus minusve sulco longitudinali
connexis. LHlytra elongata parallela; basi foveis duabus quarum externa magna, paululum
sulciformi. Abdomen elongatum; segmentis dorsalibus tribus primis basi impressis,
4 majore. Metasternum convexum et punctatum. Pedes crassi, breves; femoribus,
preesertim anticis et intermediis, crassis; tibiis extus medio crassioribus.
g. Segmentis ventralibus (Pl. 10, fig. 7) 5 minore, arcuato, medio transversim
impresso et utrinque tuberculato, 6 valde arcuato, medio impresso, 7 rhomboidalli,
latitudine sua longiore, tenuiter carinato.
¢. Segmentis ventralibus simplicibus, 6 (ultimo) magno, triangulavi.
Long. 1,00—1,10 millim.
Le genre Octomicrus renterme des éléments de facies un peu hétérogéne, mais qui ont
tous, pour liens communs, la forme du corps tres allongée, paralléle, la massue antennaire
peu accentuée et formée de deux articles, le prothorax avec les cdtés dentés en arriére
de la fossette latérale et sinués en avant et en arriére ; quant 4 la sculpture de la téte et
du prothorax elle est assez variable, mais il n’y a jamais de sillon transversal au prothorax
130 PERCY SLADEN TRUST EXPEDITION
Cette espece se distingue de toutes les autres africaines et asiatiques, par la présence, sur
le prothorax, d’une fossette discoidale plus ou moins reliée & la fossette basale par un
sillon longitudmal un peu variable. La forme la plus voisine est une espéce nouvelle
(angustatus Raftray, i. 1.), qui sera trés prochainement publiée et qui a été découverte par
MM. Alluaud et Jeannel dans les foréts du Mont Kénia, Afrique orientale anglaise.
Loc. Mahé: foréts de la Mare aux Cochons, 1500 & 2000 pieds, I—II. 1909.
Tribu Batrisini Raffray, Rev. d’Ent., 1890, pp. 82, 108.
Genre BatrisopEs Reitter, Verh. Nat. Ver. Briinn, xx. 1881, p. 205.
8. Batrisodes caudatus, nov. sp. (Pl. 10, figs. 8, 9). Oblongus, sat crassus, parum
convexus, castaneus, totus setis flavis, depressis vestitus ; capite et prothorace fortiter,
elytris et abdomine tenuiter rugoso-tuberculatis. Caput transversum; fronte medio
depressa et sulcata, angulis anticis obtusis sed elevatis; temporibus obliquis et leviter
rotundatis ; vertice toto carinato, foveis duabus sulco arcuato et antice angulato junctis.
Oculi prominentes, medio siti. Antenne crasse, clava parum conspicua; articulis
1° paululum majore, 3—7 quadratis, 8 breviore, leviter transverso, 9, 10 crescentibus,
transversis, 11 ovato, acuminato. Prothorax capite longior et paululum latior, pone
medium dentatus et valde coarctatus; sulcis longitudinalibus tribus, dorsali postice
in foveam desinente, sulco transverso nullo ; ante basin utrinque tuberculo valido, acuto ;
basi ipsa quadrifoveata. Elytra latitudine sua longiora; humeris obliquis et dentatis ;
basi bifoveata, sulco dorsali brevissimo. Abdomen elytris vix longius, postice attenuatum ;
seomentis dorsalibus tribus primis subeequalibus, 1° basi trifoveato, 4 postice plus minusve
porrecto, 5 (ultimo) supra inconspicuo, infra declivi et incurvo; segmentis ventralibus
2 majore, 3, 4, 5 minoribus, 6 magno. Pedes validi, crassi; femoribus inflatis; tibiis
subrectis, anticis et intermediis medio leviter incrassatis.
g. Segmento 4 dorsali in cornu valido, insuper recurvo, apice obtuso, porrecto (Pl. 10,
fig. 9). Metasternum valde impressum. Segmentis ventralibus 3—5 leviter medio
depressis, 6 magno, transverso, apice bisinuato, medio valde impresso. ‘Tibiis intermediis
apice calcaratis.
Long. 3,60 millim.
?. Segmento 4 dorsali multo minus porrecto et apice valde obtuso ; ultimo ventrali
transverso, apice vix sinuato, simplice. Metasternum leviter impressum.
Long. 2,80 millim.
Par la forme de la téte, du prothorax et des antennes, la sculpture et la pubescence
du corps, cette espéce se rapproche du groupe xxx. (Vid. Gén. et Cat. Psélaphides, Ann.
Soc. Ent. France, 1904, pp. 28, 88) dont toutes les espéces habitent les iles de Java,
Sumatra, Singapore et Penang; mais elle s’en distingue, comme d’ailleurs de toutes les
espéces connues, par le singulier développement du 4°™° tergite qui, chez le g surtout,
sallonge, en arriére, en une large corne relevée en dessus et obtuse au sommet; quant au
5&me tergite, caché par la proéminence du 4°™¢ et invisible quand linsecte est regardé par
en dessus, il est perpendiculaire 4 l’axe du corps et méme un peu recourbé en dessous, pour
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 131
venir s'appliquer contre le 6°™° sternite. C'est en réalité une forme asiatique & modification
autochthone assez importante.
Tl y a deux couples dont les ¢ et les $ different entre eux, non seulement par le plus
ou moins grand développement de la corne du 4°™* tergite, les caractéres sexuels du
6°me sternite et l’éperon des tibias intermédiaires, modifications normales, mais encore par
la taille qui est considérablement moindre chez la $ que chez le f; d’autre part les deux
sexes sont identiques.
Mais ce qui est singulier cest que les deux # ont été trouvés ensemble A Vile
Silhouette, dans une fourmilitre, et les deux 9 3 I’lle Praslin et, pour ces dernieres, les
notes que Mr Hugh Scott a bien voulu me communiquer, n’indiquent pas qu’elles aient
été rencontrées avec des fourmis. Y a-t-il 14 une seule espéce ou deux races insulaires 4
I] faudrait pour en décider avoir f et 2 de la méme localité.
Loc. Silhouette: forét au-dessus de la Mare aux Cochons, dans une fourmiliére de
Vollenhovia lavithorax Emery, subspecies alluaudi Emery (déterm. A. Forel), habitant
une souche d’arbre, 6. IX. 1908, ¢. Praslin: Cdtes d’Or Estate, forét de palmier
coco-de-mer, dans la Vallée de Mai, XI. 1908, 2.
Tribu Brachglutini Raffray, Ann. Soc. Ent. France, Ixxili. 1904, p. 108.
Genre Barraxis Reitter, Verh. zool.-bot. Ges. Wien, xxxi. 1881, p. 464.
9. Batraxis egregia, nov. sp. (Pl. 10, fig. 10).
Oblonga, convexa, nitidissima, glabra, nigropicea, elytris obscure rubescentibus,
palpis, pedibus et antennis rufis, articulis duobus ultimis infuscatis exceptis. Caput
antice attenuatum; fronte medio depressa, angulis anticis quadratis, elevatis; sulcis
duobus obliquis antice connexis ; vertice convexo; temporibus obliquis. Oculi prominentes.
Antennz elongate, articulis duobus primis majoribus, 3—9 cylindricis, 3—7 latitudine
sua duplo, 8, 9 dimidio longioribus, 10 trapezoidali, transverso, 11 magno, triangulari et
margine externa sulcato. Prothorax capite paululum latior, antice attenuatus, angulo
antico rotundato, lateribus postice obliquis, angulis posticis acutis, ante basin medio sulco
transverso utrinque abbreviato, et basi ipsa carinata. Elytra latitudine sua equilonga,
humeris et lateribus rotundata, angulis externis posticis leviter auriculatis, foveis basalibus
et stria dorsali deficientibus. Abdomen elytris paululum brevius, postice rotundatum,
basi lateribus emarginatum; segmento 1° dorsali magno, basi transversim triimpresso,
impressione media latiore; margine laterali triangulari, intus carinula obliqua et sinuata
limitata. Metasternum impressum. Pedes elongati ; femoribus, preesertim anticis et
intermediis, clavatis ; tibiis anticis leviter sinuatis, medio crassioribus, intermediis rectis,
simplicibus, apice vix incrassatis, posticis longioribus, perparum arcuatis, apice clavatis et
caleare tenui, longo, filiformi preeditis.
g. Tibiis anticis apice intus valde calcaratis.
Long. 1,70 millim.
Cette espéce se distingue de toutes les autres par sa coloration spéciale et aussi par
Yabdomen un peu plus large & sa base que les élytres et ot il porte une entaille assez
132 PERCY SLADEN TRUST EXPEDITION
profonde qui semble destinée 4 recevoir l’angle postéro-externe de |’élytre, qui est un peu
saillant en forme d’oreillette. La marge externe du premier segment dorsal est grande, en
triangle trés allongé, et limitée, 4 l’intérieur, par une caréne qui va en s'atténuant de la
base au sommet. .
Cette espéce devrait former un groupe & part, prés du groupe vi. (Vid. Ann. Soe. Ent.
France, 73, 1904, pp. 119, 199) qui a déja lVabdomen brusquement rétréci, mais non
encore entaillé, 4 la base, et le dernier article des antennes triangulaire ; elle est particu-
liérement voisine de indica Raffr. de l'Inde, qui est uniformément ferrugineuse, avec des
antennes plus épaisses et dont les tibias postérieurs n’ont pas ce long éperon filiforme qui,
chez egregua, est lapanage des ? aussi bien que des 3.
Loc. Mr Hugh Scott en a recueilli une dizaine d’exemplaires qui proviennent presque
tous de |’Ile Praslin : Cotes d’Or Estate, forét de palmier coco-de-mer. Ile de Mahé, un
seul exemplaire; Cascade Estate, 800 pieds ou au-dessus. Ile Silhouette, un seul
exemplaire: forét au-dessus de la Mare aux Cochons, plus de 1200 pieds, IX. 1908.
10. Batraxis msularis, nov. sp.
Oblonga, convexa, dilute castanea, nitida, pedibus antennisque rufis, pube tenui, sat
elongata, dispersa. Caput transversum, lateribus leviter rotundatum; fronte truncata,
utrinque triangulatim valde foveata ; vertice magno, convexo. Oculi mediocres, paululum
ante medium siti. Antennze elongate ; articulus duobus primis paululum majoribus, 3—9
ceylindricis, 3 latitudine sua triplo, 5, 6 duplo, 4, 7 dimidio longioribus, 8, 9 fere quadratis,
10 majore, quadrato, 11 magno, irregulariter ovato, intus subrecto, extus convexo.
Prothorax antice plus et postice minus attenuatus, ante basin medio fovea minuta, basi ipsa
carinata. Elytra latitudine sua paululum longiora, humeris rotundata, lateribus perparum
rotundata ; angulis posticis simplicibus. Abdomen elytris paulo brevius, postice
rotundatum, basi haud coarctatum; segmento 1 magno, basi haud impresso, margine
laterali angusta. Metasternum foveatum. Pedes elongati; femoribus clavatis; tibiis
anticis et intermediis rectis, ad apicem gradatim incrassatis, posticis perparum incurvis et
apice sat abrupte clavatis.
Long. 1,50 millim.
Cette espéce est trés différente de la précédente ; la téte transversale est arrondie sur
les cdtés; de chaque cdté, en avant, il y a une fossette un peu triangulaire qui fait
ressortir les angles antérieurs; le dernier article des antennes est arrondi et un peu
oblique en dehors, mais sans étre triangulaire et sans avoir de sillon ; les cdtés des élytres
sont presque paralléles ; enfin l’abdomen n’est ni entaillé ui méme étranglé & la base.
Cette espéce appartient au groupe 11. (Vid. Ann. Soc. Ent. France, 73, 1904, pp. 118,
198) qui ne renfermait encore qu'une seule espéce (hartella, Raftr.) de la Nouvelle-Guinée
Allemande (Céte Sud-est), mais elle en différe par la forme du corps beaucoup plus
allongée, la téte plus transversale, les antennes plus longues, plus gréles, avec le dernier
article plus allongé et plus oblique, la coloration générale est plus foncée.
Loc. Mahé: Cascade Estate, III. 1909: un seul exemplaire en assez mauvyais état.
RAFFRAY—COLEOPTERA, PSELAPHIDA! DES SEYCHELLES 133
Genre Triomicrus Sharp, Trans. Ent. Soc. London, 1883, p:. 325.
il. Triomicrus seychellensis, nov. sp. (Pl. 10, fig. 11).
Oblongus, antice attenuatus, rufus, glaber, antennis pedibusque testaceis. Caput
leviter transversum, antice valde attenuatum; fronte medio leviter depressa et angulis
anticis perparum nodosis ; foveis tribus liberis. Oculi medio siti, sat prominentes. Palpi
sat elongati; articulis 2 elongato, gracili, apice abrupte clavato, 3 globoso, 4 secundo
paululum longiore, oblongo, apice obtuse acuminato. Antenne sat elongate, graciles ;
articulis duobus primis multo majoribus et latitudine sua fere duplo longioribus, 3 leviter
obconico, latitudine sua fere triplo, 5 duplo, 4, 6 dimidio longioribus, 7 fere quadrato,
8 monili, 9 paululum majore et transverso, 10 plus quam duplo majore, transverso, 11
magno, ovato, acuminato. Prothorax capite major, antice multo plus quam postice
attenuatus, latitudine sua maxima ante medium, foveis tribus minutis, liberis. Elytra
latitudine sua paululum longiora, ad basin attenuata; humeris nullis, lateribus leviter
obliquis ; foveis basalibus et stria dorsali deficientibus. Abdomen elytris brevius, postice
rotundatum ; segmento 1° dorsali majore. Metasternum breve, concavum. Segmento 1°
ventrali brevi, sed conspicuo, 2 magno, medio triangulatim deplanato, 3—5 brevissimis,
6 magno, toto transversim impresso. Pedes elongati; femoribus, preesertim anticis et
intermediis, infra nodosis ; tibiis anticis gracilibus, subrectis, apice intus vix perspicue
sinuatis, intermediis subrectis, crassioribus, intus apice calcare valido et obliquo armatis ;
posticis leviter incurvis, ad apicem incrassatis, intus apice brevissime calcaratis 3.
Long. 1,30 millim.
Le genre Triomicrus qui, Jusqu’a ce jour, ne comprenait que quelques especes
spéciales & la Chine orientale et au Japon, a le premier sternite court mais déji apparent,
et il constitue ainsi une forme de transition entre les Brachyglutini dont le premier
sternite est invisible et les Tychini dont ce méme sternite est le plus grand et dépasse les
hanches postérieures; mais il rentre dans la tribu des Brachyglutini parceque la face
inférieure de la téte porte cette carene longitudinale qui est caractérisque de cette tribu
et dont les Tychini n’ont pas méme un vestige.
Le Triomacrus seychellensis qui présente les caractéres primordiaux de ce genre est
une forme cependant assez isolée. Chez tous les Triomicrus asiatiques le 3°™° article
des palpes est obconique, le 4°™° fusiforme et les élytres ont une strie dorsale. Chez le
seychellensis (Pl. 10, fig. 11) le 3°™° article des palpes est globuleux, le 4°™° est & peine
rétréci 4 la base, en ovale tres allongé, obtusément acuminé au sommet et les élytres n'ont
pas de strie dorsale. Je ne crois pas que ces différences puissent légitimer la création
dun nouveau genre, d’autant plus que les quatre espéces asiatiques ne présentent pas,
entre elles, une homogénéité absolue et, qu’en s’engageant dans cette voie, on serait
fatalement conduit & pulvériser les genres, sans que ce procédé présentat, & mon avis,
aucun avantage pour la science. Les caracteres fondamentaux du genre Triomicrus
sont l’allongement et la gracilité plus grande des palpes, la briéveté du métasternum, le
plus grand écartement des hanches postérieures et la visibilité du 1° sternite qui, cepen-
dant, ne dépasse pas les hanches.
SECOND SERIES—ZOOLOGY, VOL. XVI. 18
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134 PERCY SLADEN TRUST EXPEDITION
Triomicrus seychellensis présente tous ces caractéres.
Loc. Silhouette: forét, environ 1500 pieds, 15. VIII. 1908: trouvé sur la terre,
parmi les feuilles mortes, particulitrement 4 la base des feuilles tombées de palmiers,
dans un endroit tres ombreux et humide. Deux exemplaires.
Tribu Tychini Raftray, Ann. Soc. Ent. France, Ixxil. 1903, p. 490; 1904, p. 254.
Genre SuNoRFA Raffray, Rev. d’Ent., 1. 1882, p. 28.
12. Sunorfa fasciculata, nov. sp.
Oblongo-ovata, convexa, tota rufa. Caput leviter transversum; fronte antice
arcuatim carinata, angulis anticis obtusis et leviter nodosis, postice late excavata; vertice
transversim convexo ; temporibus leviter rotundatis; margine postica integra. Antenne
parum elongatze, articulis duobus primis majoribus, 1° latitudine sua longiore, 2 quadrato,
3—8 minutis, monilibus, 9, 10 transversis, latitudine crescentibus, 11 magno, ovato. Pro-
thorax capite longior et paulo latior, postice attenuatus, angulis anticis rotundatis, lateribus
leviter arcuatis ; juxta basin sulco transverso recto et medio interrupto. Elytra minute
sed confertim punctata, latitudine sua longiora, subovata; humeris elevatis et carinatis ;
foveis quatuor basalibus; stria suturali tenuissima, dorsali deficiente. Abdomen breve
et postice rotundatum; margine laterali angusta ; segmentis dorsalibus zequalibus ;
ventralibus duobus primis magnis, zequalibus, 83—5 minutis et decrescentibus, 6 magno
transverso. Metasternum magnum, deplanatum. Pedes validi; coxis posticis valde
distantibus; femoribus clavatis; tibus apice leviter clavatis, posticis incurvis.
g. Carina frontali magis rotundata et angulis anticis magis nodosis ; vertice antice
abrupte truncato et fasciculato. Segmento sexto ventrali (ultimo) transversim depresso.
Long. 1,00—1,10 millim.
Le front forme, en avant, un bourrelet arrondi et assez mince et, en arriére, il y a
une grande excavation; chez le male cette excavation est limitée, en arritre, par une
troncature abrupte et caréniforme du vertex, cette caréne émet un faisceau de poils dorés
au-dessus de la cavité frontale. Chez la femelle le vertex est simplement fortement et
transversalement convexe.
Par la forme et la position du sillon transversal du prothorax, cette espéce se
rapproche dexsculpta Shaufuss de Sumatra, sur laquelle l’entomologiste dresdois avait
fondé le genre Bythinimorpha qui est synonyme de Sunorfa Raffr. qui lui est antérieur.
Mais exsculpta est beaucoup moins allongée et les élytres sont a peine ponctués.
Loc. Ile Félicité: forét, XII. 1908. Mahé: Cascade Estate, III. 1909. Praslin:
Cotes d’Or Estate, forét de palmiers coco-de-mer, dans la Vallée de Mai, XI. 1908.
Genre SunorFA Raftray.
Sous-genre SUNORFOIDES, nov. subgen. (Pl. 10, figs. 12—14).
Ce nouveau sous-genre ne différe du genre Sunorfa que par la présence, chez le 3,
d'un septiéme sternite qui fait toujours défaut chez Sunorfa.
RAFFRAY—COLEOPTERA, PSHELAPHIDAi DES SEYCHELLES 135
Le corps est plus ou moins ovoide; la téte toujours transversale et diversement
impressionée surtout chez les ¢; le 2°™° article des palpes est légerement renflé au sommet,
le 3°™e petit, subtriangulaire, 4°™° grand, allongé, plus ou moins sécuriforme. Prothorax
plus ou moins transversal et rétréci en arriére ou briévement cordiforme, avec un sillon
transversal sinué au milieu. LElytres assez grands, 4 épaules marquées, avec 4 fossettes
basales. Abdomen rebordé & segments dorsaux (tergites) égaux; segments ventraux
(sternites) 1 trés grand, dépassant de beaucoup les hanches postérieures qui sont écartées,
2 2 5 allant en diminuant, ¢ 6 plus grand, 7 petit, en ovale transversal, ? 6 (dernier)
grand, triangulaire obtus au sommet. Pieds robustes; tibias antérieurs et intermédiaires
un peu renflés au milieu, les postérieurs renflés & l’extrémité et-toujours plus ou moins
arqueés.
La téte est diversement impressionnée, mais n’est pas armée, comme cela arrive
fréquemment dans le genre Sunorfa. Le front forme généralement, en avant, un
bourrelet plus ou moins arrondi qui limite des impressions ou sillons variables ; la marge
postérieure a toujours un sillon qui s’avance sur le vertex. Mais le seul caractére
important qui oblige 4 séparer Sunorfordes de Sunorfa est la présence, chez le ¢ des
premiers, d’un septieme sternite qui manque chez les seconds.
Mr Hugh Scott m’a communiqué une préparation trés réussie dune Sunorfoides @
dont il est cependant difficile de déterminer l’espéce, parceque les reliefs sont peu
apparents dans le baume. I] m’a été possible de donner une figure des organes buccaux
(PI. 10, fig. 12); il y manque les paraglosses que je n’ai pu apercevoir, la préparation de
Yinsecte entier ayant trop d’épaisseur, pour permettre lusage d’objectifs assez forts pour
discerner ces organes trés diaphanes, surtout quand les différentes parties de la bouche ne
sont pas dissociées.
Cette bouche ne présente d’ailleurs aucun caractére spécial, seulement les mandibules
sont plus allongées et moins recourbées que dans la grande majorité des Psélaphides.
Ce sous-genre semble assez répandu aux Seychelles. Mr Hugh Scott en a découvert
quatre espéces.
Un tableau comparatif permettra d’abréger les descriptions tout en rendant leur
détermination plus facile.
A 2. Corps roux. Elytres noirs ou enfumés.
B 2. Elytres noirs.
C2. Taille plus grande (1,20—1,30). Elytres fortement pontués. Téte surtout g plus
transversale, fortement triimpressionnée Be vat a4 nigripennis.
C1. Taille plus petite (1,10). Elytres moins fortement ponctués. Téte moins trans-
versale, un large sillon arqué sur le front... |... sige soe WRGOUNDIP.
B1. Elytres plus ou moins enfumés, fortement ponctués. Téte avec un tres large sillon
transversal, front tronqué carrément g, plus arrondi — punctipennis.
A 1. Corps entiérement noir de poix, antennes et pieds testacés. Elytres fortement
ponctués. Téte a sillon frontal circulaire, front légérement arrondi priced.
18—2
136 PERCY SLADEN TRUST EXPEDITION
13. Sunorfoides ngripennis, nov. sp. (Pl. 10, fig. 13).
Obscure rufa, elytris nigropiceis, pube grisea tenui et dispersa. Caput validum ;
fronte antrorsum carinata et leviter arcuata, angulis externis rotundatis, incisis et
irregulariter punctatis ; temporibus quadratis sed angulo postico rotundato; antice foveis
tribus magnis, frontali transversa, lateralibus obliquis; vertice transversim convexo ;
margine posteriore, juxta collum, suleata. Antennarum articulis 1 magno, subcylindrico,
2 dimidio minore, sequentibus minutis, monilibus, 9—10 transversis, latitudine cres-
centibus, 11 breviter ovato. Prothorax capite longior et paululum latior, cordatus,
latitudine sua maxima ante medium et angulis anticis valde rotundatis, lateribus ad
basin obliquis; sulco transverso leviter sinuato, medio obtuse angulato et perparum
ampliato. Elytra magna, sat forte, sed paululum irregulariter et parum confertim
punctata, humeris nodosa ; basi quadrifoveata 9.
Long. 1,20—1,30 millim.
Le g de cette espéce reste inconnu.
Loc. Mahé: prés du Morne Blanc, environ 800 pieds, X—XI. 1908; forét haute
et trés humide, 1500 & 2000 pieds environ, entre Trois Fréres et Morne Seychellois,
XII. 1908.
14. Sunorfoides bicolor, nov. sp. (Pl. 10, fig. 14).
Méme coloration que nigripennis a laquelle elle ressemble beaucoup, mais un peu plus
petite. La téte est un peu moins transversale; le bourrelet antérieur du front est tres
obtusément angulé au milieu, les angles antérieurs sont arrondis, entaillés et ponctués ;
sur le vertex 4 la hauteur des yeux et tres pres d’eux, il y a, de chaque cété, une fossette
d’ot' part un sillon large et diffus qui suit le contour du bourrelet frontal; les tempes sont
plus obliques. Les antennes sont semblables, mais cependant le premier article est moins
grand. Prothorax un peu plus court, avec les angles antérieurs moins accusés et les cdtés
moins rectilignes. Elytres plus courts, ponctuation moins forte et encore moins serrée.
g. Segments ventraux (Pl. 10, fig. 14); 1 trés grand, 2, 3, 4 allant en diminuant
légérement, 5 notablement plus court, 6 plus grand que le 4°™°, avec une dépression
médiane transversale, 7 en ovale tres transversal, avec un petit tubercule.
Long. 1,10 millim.
Loc. Silhouette: cette espéce a été prise en assez grand nombre dans la forét,
au-dessus de la Mare aux Cochons, & la base des feuilles d’un palmier (Stevensonia),
24, IX. 1908; un exemplaire fut trouvé le méme jour sur le sommet le plus élevé de
lTle & 2400 pieds, sur la terre entre des feuilles mortes tombées et humides.
15. Sunorfoides punctipennis, nov. sp.
Rufa, paululum pallidior, elytris plus minusve infuscatis. Caput in utroque sexu
diversum. Antennee et prothorax sicut in preecedente (bicolor); prothoracis attamen
sulco transverso medio magis angulato et basi ipsa tenuiter punctulata. Elytra magna,
fortiter et sat confertim punctata, humeris minus nodosa; basi foveis quatuor.
RAFFRAY—COLEOPTERA, PSELAPHIDA DES SEYCHELLES 137
g. Caput quadrato-transversum ; fronte antice recte truncata et carinata, medio
leviter amplata et triangular, angulis anticis obtusis, leviter elevatis; in fronte sulco
transverso, lato, tenuiter punctato, medio postice leviter ampliato et utrinque, juxta
oculos, desinente; vertice maxime convexo; margine posteriore, juxta collum, valde
emarginata et suleata. Metasternum depressum. Segmentis ventralibus 5 brevissimo,
6 quarto multo majore, medio depresso, 7 minuto, transversim oblongo-ovato. Femoribus
magis inflatis.
?. Caput sicut in “ bzcolore,” sulco attamen frontali latiore, magis punctato, angulis
anticis magis elevatis et prominentibus. Segmento ventrali 6 breviter triangulari, apice
rotundato.
Long. 1,10—1,20 millim.
Loc. Mahé: Cascade Estate, 800 pieds et au-dessus; prés du Morne Blanc, environ
800 pieds, 10. XI. 1908. Silhouette: forét prés du Mont Pot-a-eau, environ 1500 pieds,
VIII. 1908: quelques exemplaires seulement dans chaque localité.
16. Sunorfoides picea, nov. sp.
Tota nigro-picea, palpis, antennis pedibusque rufo-testaceis, pube pallida tenui et
dispersa. Caput valde transversum ; carinula anteriore frontali medio exili et rotundata,
lateribus crassiore, angulis anticis parum notatis, obliquis et vix incisis; sulco frontali,
lato, profundo, medio transverso et lateribus obliquo; vertice magno et convexo; tem-
poribus quadratis sed angulo postico hebetato; margine postica, juxta collum, minus
emarginata. Antennarum articulis 1° magno, elongato, 2 subquadrato, ceterum sicut in
precedentibus speciebus. Elytra fortiter et sat confertim punctata.
g. Segmentis ventralibus 5 brevissimo, 6 quarto multo majore, basi preesertim late
depresso, 7 minuto, valde transverso.
?. Segmento ventrali 6 (ultimo) magno, triangulari, apice obtuso.
Long. 1,10—1,20 millim.
Loc. Silhouette: il en a été pris quelques exemplaires seulement, dans la forét,
au-dessus de la Mare aux Cochons, entre les bases des feuilles du Pandanus Sechellarum,
22. IX. 1908.
Genre ApopLectus Raffray, Rev. d’Ent., ix. 1890, pp. 96, 102.
Glapharsenus Raftray, Ann. Mus. Nat. Hung., 1903, p. 36.
17. Apoplectus obesus, nov. sp.
Brevis crassus et convexus, antice attenuatus, dilute ferrugineus, tenuiter pubescens.
Caput latitudine sua longius, antice attenuatum; lateribus obliquis; angulo postico
obtuso; fronte leviter arcuata et medio depressa; angulis anticis parum elevatis ; sulco
frontali transverso, integro; inter oculos foveis duabus inter se et ab oculis subsequaliter
distantibus, cum sulco frontali sulcis duobus obliquis junctis ; margine postica sinuata et
medio, juxta collum, sulcata. Mandibule magnze et prominentes. Oculi sat magni sed
parum prominentes. Antennze breves et crassz; articulis 1 elongato, 2 quadrato, 3—8
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138 PERCY SLADEN TRUST EXPEDITION
minoribus, leviter transversis, 9 magis et 10 valde transversis, 11 magno, breviter ovato,
apice obtuso. Prothorax capite latior et brevior, transversus, postice multo plus quam
antice attenuatus, lateribus rotundatus et pone medium obsolete unidentatus et bisinu-
atus; foveis tribus liberis quarum mediana minore. Elytra minute et disperse rugoso-
punctata, leviter transversa, ad basin attenuata, lateribus rotundata; humeris elevatis et
obtuse carinatis; basi bifoveata, fovea externa majore. Abdomen elytris subzequale,
apice rotundatum; segmentis dorsalibus equalibus; ventralibus 1 conspicuo, coxas
posteriores haud superante, 2, 3 paululum majoribus, 4, 5 decrescentibus, 6 magno, trans-
verso, basi medio impresso. Metasternum planum, tenuiter sulcatum. Pedes validi;
femoribus crassis; tibiis subrectis, ad apicem leviter incrassatis @.
Long. 1,60 millim.
Cette espéce, dont on ne connait malheureusement que la 9, se rapproche de latus
Raffray; mais sa téte est imponctuée; le prothorax est plus transversal, la double
sinuosité et la dent placée entre elles, en avant de la base, sont & peime marquées; la
fossette humérale des élytres n’est pas sulciforme ; l’abdomen n’est pas plus long que les
élytres et son premier segment dorsal n’a pas d'impressions basales.
Ce genre ne comprenait, jusqu’’ ce jour, que six espéces toutes de la Nouvelle-
Guinée et surtout des possessions allemandes sur la cdte sud-est.
Loc. WUn’y en a que deux exemplaires, l’un de lle Praslin: Cotes d’Or Estate, de
la forét de palmiers coco-de-mer, dans la Vallée de Mai, XI. 1908. Le second, mal-
heureusement sans téte, de l’ile de Mahé: Cascade Estate, 800 pieds et au-dessus.
EXPLICATION DE LA PLANCHE.
Fico. 1. Hughia phantasma, n. g. et n. sp., ff.
Hig, 2 Pe carinata, n. g. et n. sp., §/, derniers sternites.
Mig. 38. Cerennea brevipennis, n. g. et un. sp.
Fig. 4. Apheloplectus longicollis, n. g. et n. sp, ff.
Iie is x palpe maxillaire.
Fig. 6. Omobathus elegans, n. g. et n. sp. 2.
Fig. 7. Octomicrus semipunctatus, n. sp., g, derniers sternites.
Fig. §&. Batrisodes caudatus, n. sp., -
Iie, @), ie a Jf. arriére corps vu de profil: H, hanche postérieure: F, fémur:
S, sternites: 7’, tergites: H, élytres.
Fig. 10. Batraxis egregia, nu. sp., .
Fig. 11. Triomicrus seychellensis, n. sp., palpe maxillaire.
g. 12. Subgenus Swnorfoides, nov., organes buccaux: ZL, labre: M, mandibule: PL, palpes labiaux:
LA, langnette: MH, menton: PWM, palpe maxillaire: MA, machoire.
Fig. 13. Sunorfoides nigripennis, n. sp., ¢.
Fig. 14. FA bicolor, n. sp., &; abdomen, face ventrale.
TRANS. LINN. Soc. SER. 2.Z00L. Vou.XVI. PL.10.
PERCY SLADEN TRUST EXPEDITION.
(RAFFRAY)
A
University Press, Cambridge
D.Raffray del.
COLEOPTERA-PSHLAPHIDAE.
No. IX—RHYNCHOTA. PART J: SUBORDER HETEROPTERA.
By Wm. Lucas Distant.
(CoMMUNICATED BY J. STANLEY GARpDINER, M.A., F.RS., F.1.8.)
(With Plates 11—13.)
Read 5th June, 1913.
Smvce the Linnzean Society in 1909 did me the honour to publish a previous
enumeration of the Rhynchota of the Seychelles Islands*, based on the collections made
by Prof. Gardiner during the first expedition to those islands, a second zoological visit
under the same leadership has been concluded. Mr Hugh Scott, a member of the
expedition, who remained in the Seychelles for eight months and devoted himself to
entomological research, has made important collections, especially in the Order with which
I have had the pleasure of acquiring some familiarity. I am now enabled in the suborder
Heteroptera alone to enumerate 139 species, of which no fewer than 73 have been
regarded as nondescript, while 16 new genera have been proposed.
The distributional affinities of these insects are of intense interest. Many migrants
have arrived and remained with their specific characters unmodified; many genera
hitherto regarded as Neotropical and Oriental in character claim constituents in this
fauna; Aithiopian and Palearctic relationships are pronounced. A more detailed analysis
is deferred to the second portion of this contribution, enumerating the members of the
suborder Homoptera, when the whole of the Rhynchotal aflinities will be detailed; this
will also include a few species previously recorded but not contained in the present
collections. Meanwhile some notes have been added to the Family divisions.
I have affixed an asterisk to the numbers of those species which have been recorded
previously, and to these I have only added the new localities and not repeated the whole
geographical distribution.
Order RHYNCHOTA.
Suborder HETEROPTERA.
Family Pentatomide.
In the previous enumeration of this Family from the Seychelles (Trans. Linn. Soc.
Lond., 1909), I could only record 7 species, which with one listed elsewhere brought the
number to 8 only. This collection comprises 19 species and only misses one (Mahea
sexualis) described in my previous contribution. The number of Pentatomidze found in
the Seychelles up to this record therefore includes 20 species.
* Trans. Linn. Soc. London, ser. 2, Zool., vol. xiii. part 1, Oct. 1909, p. 29.
140 PERCY SLADEN TRUST EXPEDITION
Subfamily Cydnine.
1*. Cydnus pallidipenns, Reut.
Cydnus pallidipennis Reut., Ofv. Finn. Vetensk. Soc., xxv. p. 3 (1882).
Aithus pallidipennis Bergr., Rev. d’Ent., xii. p. 198 (1893).
Loc. Seychelles. Mahé; 1906 (Thomasset); Long Island, VI. 1908.
2. Macroscytus fryert, sp. n.
Black, shining, posterior margin of pronotum and the corium dark castaneous ;
antenne and rostrum (the latter imperfectly seen in carded specimen) ochraceous ; body
beneath shining piceous; femora dark castaneous; tibize ochraceous minutely spotted
with black and with the spinules black; tarsi ochraceous; head rounded in front, a few
coarse punctures on anterior margin, central lobe prominent; antennze with the second,
third and fourth joints subequal in length, third and fourth stoutest (fifth mutilated in
typical specimen) ; pronotum sparsely and irregularly but distinctly punctate, the surface
irregular, two distinct punctate indentations on each lateral marginal area, one before
middle, the other near posterior angle, margins of both head and pronotum rather longly
sparsely ciliate; scutellum sparsely but distinctly punctate, its apex subacute; corium
sparsely and very finely punctate, the basal area of lateral margin ciliate ; membrane pale
brownish ochraceous, passing abdominal apex; tibize strongly spinulose, the spinules
black.
Long. 74 mm.
Loc. Aldabra, 1908—9 (Fryer).
Allied to M. lobatus Sign., but differmg in the distinctly coloured tibize, different
punctuation of the pronotum and scutellum, broader apex of scutellum ete.
A single specimen only of this species has been received.
3*. Geotomus proximus, Sign.
Geotomus proximus Sign., Ann. Soc. Ent. France, 1883, p. 54, t. 4, fig. 163.
Loc. Seychelles. Silhouette, low country, VIII. 1908. Mahé: country above Port
Glaud, 500—1000 feet, XI. 1908: Cascade Estate, 800—1000 feet: marshes on coastal
plain, Anse aux Pins and Anse Royale, I. 1909: Bird and Dennis Islands, 1908 (Fryer).
Previously recorded from Praslin and La Digue.
4. (feotomus sp.
A single carded specimen from Mahé is closely allied to G. proaimus but the colora-
tion of body and legs is bright, pale castaneous with the antennze ochraceous.
Loc. Seychelles. Mahé: high forest near Morne Blane, XI. 1908.
CHILOCORISTOIDES, gen. nov.
General characters of Chilocoris but differing in the structure of the scutellum which
is distinctly shorter than broad and with the lateral margims distinctly sinuate before apex
which is rounded.
DISTANT-—-RHYNCHOTA. PART I: SUBORDER HETEROPTERA 141
The two species described by Breddin from India, as C. assmuthi and O. solenopsides,
and which I included in the Indian fauna as Chilocoris? will also come under Chilo-
coristoides.
Type C. felicitatis Dist.
Chilocoris found in India, Java and Eastern Siberia is also, according to Bergroth,
found in Madagascar, he having described a species, C. madagascariensis from that
island. From Bergroth’s description of the scutellum of this species he is evidently
correct in his generic fixation. Chilocoris probably remains to be discovered on the
Seychelles.
5. Chilocoristordes felicitatis, sp. n. (Plate 11, figs. 1, 1a).
Body above and beneath castaneous; membrane brownish ochraceous ; legs and
rostrum ochraceous ; head rounded in front, lobes about equal in length, anterior margin
with erect spinules and a few slender hairs; ocelli near base and nearer to eyes than to
each other; antennze with the first and second joints ochraceous, remaining joints
brownish-ochraceous, second joint shortest, third, fourth and fifth incrassated and almost
subequal in length ; pronotum with a transverse impression near middle which is coarsely
punctate, more sparingly so in its central area, a few scattered punctures in the posterior
angular areas, anterior margin distinctly concavely sinuate behind head; scutellum
shorter than broad, lateral margins sinuate before apex which is rounded, anterior and
lateral margins closely punctate, the latter impunctate on the apical margins, a few
scattered punctures on disk ; corium with a darkly punctate curved submarginal line from
base to about one third before apex, and dark punctures on the claval margin, the inner
disk of corium sparingly punctate; membrane passing abdominal apex ; rostrum reaching
the posterior coxze ; tibize longly spinose.
Long. 3 mm. :
Loc. Seychelles. Félicité Island, from forest, XII. 1908.
6. Chilocoristoides mahensis, sp. n. (Plate 11, fig. 2).
Above dark shining piceous, corium a little more castaneous ; membrane subhyaline ;
head rounded in front, with erect spinules and a few slender hairs; antennz brownish
ochraceous, second joint small, third, fourth and fifth incrassate and subequal in length ;
pronotum glabrous, without transverse impression and with a few scattered punctures on
each side of posterior disk; scutellum with a few scattered punctures on disk, anterior
and lateral margins closely punctate, the latter impunctate on the apical margins, shorter
than broad, lateral margins sinuate before apex which is rounded ; corium with a darkly
punctate curved submarginal line from base to about one fourth before apex, and dark
punctures on the claval margin, the upper inner disk of corium sparingly punctate ;
membrane slightly passing abdominal apex ; tibiee longly spinulose.
Long. 2 mm.
Loc. Seychelles. Mahé: high forest of Morne Blane and Pilot ; Cascade Estate,
about 800—1500 feet.
SECOND SERIES—ZOOLOGY, VOL. XVI. 19
142 PERCY SLADEN TRUST EXPEDITION
Allied to the preceding species C. felicitatis, principally differing in the punctuation,
absence of the transverse impression to the pronotum, smaller size etc.
Subfamily Pentatomine.
7. Husarcocoris imperator, sp. n. (Plate 11, fig. 3.)
Head brassy-black, coarsely punctate, the central lobe ochraceous, blackly punctate ;
pronotum ochraceous, coarsely darkly punctate, anterior marginal area black, the lateral
and anterior margins and two spots connected with the anterior margin pale flavescent,
posterior angles obtusely subprominent and there slightly black ; scutellum brassy-black,
with an oblique fascia on each side commencing near basal angles, joined beyond middle
and thence straightly and medially continued to apex where it is broadened, and a small
central spot on basal margin, flavescent; corium ochraceous, darkly punctate, base of
costal margin ochraceous; membrane hyaline, slightly passing abdominal apex; body
beneath brassy-black, lateral margins of sternum flavescent, basal marginal areas of
prosternum, and broad lateral margins to abdomen, pale ochraceous, darkly punctate ;
legs pale ochraceous with dark speckles; antennze dark ochraceous, the apical joint
piceous, second, third and fourth joints subequal in length ; head about as long as breadth
between eyes, the lateral margins distinctly sinuate; scutellum coarsely punctate, the
flavescent fascia impunctate, except on apical area, its lateral margins moderately sinuate,
its apex rounded and just passing base of membrane; rostrum imperfectly seen in the
carded unique type.
Long. 44 mm. Lat. 34 mm.
Loc, Aldabra, 1908 (Fryer).
Allied to H. V—flavum Horv. from Madagascar.
8. Carbula marginella (Thunb.).
Cimex marginellus Thunb., Hem. Rostr. Cap., ii. 4 (1822).
Carbula litigatri« Kirk., Cat. Hem. (Het.), i. p. 88 (1909).
Loc. Cosmoledo, 1907 (Thomasset). Aldabra, 1907 (Thomasset).
This species is widely distributed in South Africa and has been recorded from the
Congo area.
9. Stenozygum aldabranum, sp. n. (Plate 11, fig. 4).
Above, shining metallic, olivaceous green with ochraceous and greyish markings ;
head with two transverse ochraceous spots on each side of central lobe and before the
eyes, the central lobe longitudinally streaked with the same colour, greyishly so towards
base, a small greyish spot near inner margin of each eye; pronotum with the lateral
margins and a central longitudinal irregular fascia (the lateral margins continued inwardly
behind each eye), ochraceous, anterior and basal margins and discal spots on anterior lobe,
more or less greyish; scutellum with three large basal spots, the apex and a large
subapical spot, ochraceous, the central and apical spots connected by a narrow ochraceous
line; corium with the basal third of apical margin and a large subapical spot, ochraceous,
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 143
the latter inwardly and outwardly margined with greyish, and with a small greyish discal
spot, membrane blackish, the apical area greyish-white ; connexivum metallic-green with
greyish spots ; body beneath and legs greyish, the lateral areas of sternum and abdomen
more or less ochraceous, the lateral with marginal and submarginal dark spots; legs more
or less streaked with black ; antennze black, first joint pale at base, second shorter than
third, third and fifth subequal, fourth a little longest; head with the lateral margins
sinuate, their edges a little reflexed, practically impunctate ; pronotum with the posterior
lobe and the transverse depression dividing the lobes somewhat coarsely punctate ;
scutellum coarsely punctate, the ochraceous macular markings impunctate or nearly so;
corium thickly punctate; membrane passing the abdominal apex.
Long. 7 mm.
Loc. Aldabra, 1908—9 (Fryer).
10. Stenozygum insularum, sp. n.
Allied to the preceding species S. aldabranum, but smaller, the colour dark indigo-
blue, the markings pale flavescent, not ochraceous, basal spots to the scutellum much
smaller; third joint of antennz longer in proportion to the second joint than in S. alda-
branum; apex of scutellum a little narrower; legs and antenne darker in hue; inner
edge of lateral margins to anterior pronotal lobe with a distinct longitudinal series of
punctures.
Long. 55 mm.
Loc. Cosmoledo, 1907 (Thomasset).
11*. Nezara viridula (Linn. ).
Cimex viridula Linn., Syst. Nat., ed. 10, p. 444 (1758).
Nezara viridula Dist., Biol. Cent.-Am., Rhynch., i. p. 78 (1880) for synonymy.
Loc. Seychelles. Mahé: Cascade Estate, ca. 1000 feet, and other localities. Bird
and Dennis Islands (Fryer). Previously taken in Cargados Carajos Islands.
12. Nezara spicata, sp. n.
Nezara acuta Bergr. (nec Dall.), Rev. d’Ent., xii. p. 198 (1893).
Green ; lateral margins of head, lateral margins and lateral angles of pronotum, five
small spots on basal margin and the apex of scutellum, basal lateral margin of corium
and the outer margins of connexivum ochraceous, the segmental angles to connexivum
»black ; membrane hyaline; body beneath and legs green, a little paler than above ; coxee
and apices of femora, tibize (more or less), lateral margins of prosternum, abdominal
spiracles and rostrum, ochraceous, apex of rostrum black; body above thickly punctate,
the pronotum and base of scutellum somewhat rugulose ; antennze green, apex of third
joint and apical halves of fourth and fifth joints, black; second and third joints subequal
in length, shorter than fourth and fifth, fourth slightly longer than fifth ; lateral angles of
pronotum shortly spinously produced, almost laterally directed.
Long. 11 to 13 mm. ; exp. pronot. ang]. 7 to 9 mm.
19—.2
144 PERCY SLADEN TRUST EXPEDITION
Loc. Seychelles. Mahé, various localities. Silhouette: low country near sea,
Grande Barbe, VIII. 1908.
Allied to N. acuta Dall. but differing in the pronotal spinous angles which are
shorter and more laterally directed, in acuta they are much longer, testaceous and
forwardly directed ; the anterior lateral pronotal angles in spicata are a little longer and
more prominent than in the corresponding angles in acuta.
Bergroth in his enumeration of the Seychelles heteropterous fauna mistook this
species for N. acuta Dall. and his record must therefore be deleted from faunistic
catalogues.
13. Nezara heegeri (Fieb.).
Acrosternum heegeri Fieb., Eur. Hem., p. 331 (1861).
Loc. Seychelles. Silhouette, near Mont Pot-d-eau, ca. 1500 feet, and Mare aux
Cochons, over 1000 feet, VIII.—IX. 1908. Mahé, Cascade Estate, ca. 1000 feet, 1908—9.
Aldabra, 1908—9 (Fryer). Assumption, 1908—9 (Fryer); 1910 (Dupont).
A widely distributed species. Found in the Southern Paleearctic region and Canary
Islands. In the Ethiopian region it is recorded from the Gold Coast, Congo area, and
Madagascar. I also found it in Mansell Weale’s collection made in Natal.
The Nezara emmerez described by Schouteden from Mauritius (Wien. ent. Zeit.,
xxiv. p. 52, 1905) from the description given also appears to be Fieber’s species.
14. Menida transversa (Sign.) ?
Rhaphigaster transversus Sign., Ann. Soc. Ent. France (3), vill. p. 936 (1861).
A single specimen taken in Aldabra by Mr Fryer appears to generally conform to the
description of the species described from Madagascar by Signoret, and which I have not
seen.
15*. Bathycelia prelongirostris, Bergr.
Bathycela prelongirostris Bergr., Rev. d’Ent., xii. p. 200 (1893).
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, [X. 1908. Mahé:
Cascade Estate, about 1000 feet or over. Previously taken in Coetivy.
16*. Amirantea gardineri, Dist.
Amirantea gardinert Dist., Trans. Linn. Soe. London, ser. 2, Zool., xii. p. 31,
Pl, 4), 16 ep (909),
Loe. Seychelles. Mahé: near Morne Blanc, about 1000 feet, many specimens, 13.
XI. 1908. Silhouette: from several localities in forest, at or above 1000 feet; on one
occasion several specimens beaten from “ Bois merle” (Schnmudelia racemosa Linn.).
Subfamily Asopine.
17. Cantheconidea migratoria, sp. n.
Testaceous, mottled with black; antennz with the first, second and third joints
testaceous, apical half of third black, fourth and fifth black with their bases pale
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 145
flavescent, first jot short, thick, not nearly reaching apex of head, second and fourth
longest, almost subequal in length, third and fifth subequal ; head coarsely punctate, many
of the punctures black, lateral margins concavely sinuate in front of eyes, lobes about equal
in length, apical margin moderately rounded ; pronotum with the lateral margins sinuate,
the anterior portion finely crenulate, the anterior angles very shortly palely spinous,
posterior angles moderately strongly laterally produced, their apices bifid, the anterior
spine much the longest, the whole surface closely coarsely punctate with a central pale
longitudinal carinate line, the lateral margins and the anterior area a little more palely
testaceous; scutellum longer than broad at base, its apex rounded, thickly punctate,
somewhat more coarsely so on basal area, three small spots on basal margin and the
apical margin pale flavescent, a dark foveate spot at each basal angle ; corium somewhat
opaque, thickly finely punctate; membrane black, only slightly passing the abdominal
apex ; connexivum ochraceous, with a dark olivaceous spot on each side of the segmental
incisures ; body beneath pale testaceous, mottled and marked with black; sternum and
abdomen with a submarginal blackish irregular fascia which on the abdomen is connected
with the lateral margin by a series of distinct spots; femora black with their bases pale
testaceous, anterior femora with a prominent spine before apex ; tibize black, more or less
centrally flavescent, anterior narrowly, intermediate and posterior broadly, anterior tibiz
moderately dilated ; rostrum about reaching the posterior coxe.
Long. 8 to 9 mm.
Loc. Aldabra, 1907 (Thomasset).
The first and only representative of the Asopime yet recorded from the islands
visited by this Expedition.
Subfamily Tessaratomine.
18*. Sepina seychellensis, Dist.
_ Sepina seychellensis Dist., Trans. Linn. Soc. London, ser. 2, Zool., xiii. p. 31, Pl. 4,
f 3 a (1909).
Loc. Seychelles. Mahé: near Morne Blanc, about 1000 feet; Cascade Estate,
800—1000 feet (many specimens were found together in the decayed outer leaf-bases
of a Stevensonia palm growing in the forest, 29. XII. 1908). Silhouette: Mare aux
Cochons.
19. Sepia antennata, sp. n.
Fuscous-brown ; lateral margins of head and margins of central lobe, lateral and
anterior margins and a discal longitudinal spot to pronotum, basal angles and a central
longitudinal fascia to scutellum (neither reaching base nor apex) and a central discal spot
to corlum, ochraceous or testaceous; membrane piceous; connexivum ochraceous or
testaceous with dark spots at segmental incisures ; body beneath ochraceous or testaceous,
sternum and abdomen with a black submarginal fascia; legs brownish-ochraceous, apices
of femora, tibise and tarsi infuscated ; antennze with the first and second joints brownish
or piceous, remaining joints black, apices of third and fourth joints broadly annulated with
146 PERCY SLADEN TRUST EXPEDITION
pale ochraceous, first joint incrassate, scarcely or very slightly passing apex of head,
second longest, about as long as pronotum, third very short, about as long as first, fourth
longer than fifth ; head with the lobes subequal in length, the apices of the lateral lobes
inwardly a little prominent, lateral margins strongly sinuate, a spot at inner margins
of eyes either ochraceous or testaceous; pronotum with the lateral margins moderately
sinuate, slightly transversely wrinkled, distinctly punctate, anterior area slightly callose ;
scutellum, except on the macular markings distinctly punctate, moderately wrinkled ;
corium thickly finely punctate; membrane slightly passing abdominal apex in @, not
reaching the same in &.
Long. ¢ 7, 2? 85 mm., Lat. 34 to 4 mm.
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet. fle aux Récifs, 1908
(Thomasset).
The collection contains two carded specimens of this species, one of each sex from
each of the above localities. The salient features of S. antennata are found in its slender
form and antennal characters.
Family Coreidee.
In my previous communication on this fauna the Coreidz were only represented by
six species, one of which was recorded elsewhere. These are all included in this collection,
with others, now bringing up the species in the family to twelve in number.
Subfamily Coreine.
20. Anoplocnemis curvipes (Fabr.).
Cimeax curvipes Fabr., Mant. Ins., ii. p. 288, 96 (1787).
Loc. Aldabra and Cosmoledo, 1907 (Thomasset).
A widely distributed species found nearly or quite over the Continental area of the
Ethiopian region.
21*. Leptoglossus membranaceus (Fabr.).
Cimex membranaceus Fabr., Spec., ii. p. 351, 79 (1781).
Loc. Seychelles. Mahé: near Morne Blane, about 1000 feet ; Cascade Estate, about
1000 feet ; Long Island.
Previously taken in Praslin and in the Amirante Islands.
Subfamily Pseudophleine.
22*. Acanthomia horrida (Germ.).
Syromastes horrida Germ., in Silberm. Rev., v. p. 145 (1837).
Loc. Seychelles. Silhouette: near Mont Pot-a-eau, all over 1000 feet ; low country
near sea, Grande Barbe; marshy plateau of Mare aux Cochons, about 1000 feet ; cultivated
country and coast. Mahé: Cascade Estate, about 1000 feet; Long Island.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 147
23*, Stenocephalus punctipes, Stal.
Stenocephalus punctipes Stal, En. Hem., i. p. 85, 1873.
Loc. Seychelles. Mahé: Cascade Estate, about 1000 feet; Long Island; Baie
Lazare. Praslin. Bird and Dennis Islands (Fryer).
24. Cletus clavatus (Sign.).
Gonocerus clavatus Sign., Ann. Soc. Ent. France, 1860, p. 943.
Loc. Aldabra: Takamaka or Ie Michel (Fryer).
This species was originally described from Madagascar.
25. Hydara tenuicorns (Westw.).
Coreus tenuicornis Westw., in Hope Cat. i. p. 24 (1842).
Loc. Seychelles. Mahé: Port Victoria, XII. 1908.
Widely distributed throughout the Ethiopian region, including Madagascar.
Subfamily Alydine.
26*,. Leptocorisa apicalis Westw.
Leptocorisa apicalis Westw., in Hope Cat. u. p. 18 (1842).
Loc. Seychelles. Silhouette: cultivated country near coast; near Mont Pot-a-eau,
over 1000 feet; Mare aux Cochons, over 1000 feet. Mahé: near Morne Blanc, Cascade
Estate, and other localities. This species is often swept from grass in clearings in the
mountain-forests, at about 1000 feet.
Genus TENOSIUS.
Tenosius Stal, Ofv. Vetensk.-Akad. Forh., 1859, p. 460.
Sjdstedtina Schout.
Dr Schouteden has recently proposed a new genus (supra) for the species he
originally denoted as Tenosius sjdstedti in the results of Dr Sjéstedt’s Kilimandjaro-
Meru-Expedition. In subsequently describing that species elsewhere Schouteden also
proposed the new genus, and also lately proposed the location therein of my Tenosius
robustus, from the Transvaal. I cannot concur in the validity of this subdivision, one
which would alter the conception of the distribution of Tenosius in this enumeration, and
which therefore prompts these comments.
27. Tenosius proletarius (Schaum).
Alydus proletarius Schaum, Ber. Akad. Berl., 1853, p. 358.
Loc. Aldabra: Takamaka and Picard Island, XII. 1908—I. 1909 (Fryer).
Found also in 8. Africa, Mozambique, and again in Brit. India.
148 PERCY SLADEN TRUST EXPEDITION
28*. Hypselopus intermedius, Dist.
Hypselopus intermedius Dist., Trans. Linn. Soc. London, ser. 2, Zool., xiii. p. 34
(1909).
Loc. Seychelles. Mahé: Cascade Estate, about 800 feet.
The type was previously described from the same locality.
Subfamily Corizine.
29. Serinetha toricollis, Bergr.
Serinetha torricollis Bergr., Rev. d’Ent., xii. p. 200 (1893).
Loc. Seychelles. Silhouette: Mare aux Cochons. Feélicité Island, from forest,
XII. 1908.
Bergroth gives the sexual dimensions of this species as ranging from 10°5 to 13°5 mm.
The five specimens now before me measure 133 to 15 mm. Bergroth describes the species
as “SS. fraterne Westw. affinis,” but it is really more closely allied to S. ethiopis Dist., a
name I proposed for the species recognised by Dallas and Stal as Westwood’s fraterna,
the type of which I have elsewhere figured (Proc. Zool. Soc., 1901, Pl. 30, fig. 6).
30. Serinetha hamatica (Germ.).
Leptocoris hematica Germ., in Silberm. Rev., v. p. 144 (1837).
Loc. Aldabra: Ile Michel, X. 1908 (Fryer). Cosmoledo, 1907 (Thomasset). Found
also in tropical Africa, S. Africa, Madagascar and Mauritius.
31. Corizus scottr, sp. n. (Plate 11, fig. 11).
Head, pronotum and scutellum black or piceous; head with a central longitudinal
line, a transverse basal line, and the eyes, ochraceous ; pronotum with the lateral margins
(not including the posterior angles), and a more or less distinct central longitudinal line,
ochraceous; scutellum with the apex and frequently a short marginal line on each
side of base, ochraceous; corium hyaline, the basal angle piceous, the apical angle,
inner posterior marginal cell, and the venation more or less black or piceous, in some
specimens the apical angle is dull sanguineous, and in others most of the veins are
flavescate, membrane hyaline; abdomen above black, its lateral margins narrowly
ochraceous, connexivum spotted with ochraceous; head and body beneath ochraceous ;
coxee black; legs ochraceous, apical areas of femora, bases and apices (narrowly) of tibize
piceous or black; head more or less coarsely granulose; antennze brownish-ochraceous,
basal joint black, incrassate, about reaching or very slightly passing apex of head,
second and third joints subequal in length, fourth longest, thickened, in some specimens,
as in the one figured, distinctly thickened and compressed before middle, thus giving
an appearance of an extra or fifth joint ; eyes prominent, distinctly passing the anterior
pronotal angles; pronotum coarsely punctate, the anterior margin convexly prominent
and black, posterior angles subglobosely prominent and black ; scutellum coarsely punctate ;
membrane passing the abdominal apex.
Long. 3 6 to 7 mm., ? 74 to 85 mm.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 149
Loc. Seychelles. Silhouette: Mare aux Cochons ; swept from low vegetation, almost
all on plateau of Mare aux Cochons, about 1000 feet (H. Scott).
Family Lygeide.
In my last enumeration of this family I was able to record 14 species from the
Seychelles, of which only six had been received directly and eight recorded elsewhere.
The present enumeration amounts to 23 species, including all those previously obtained
by Prof. Gardiner with the exception of Nysiws cargadosensis, and those listed by
Bergroth, excluding his Stilbocoris solivagus of which no example has been captured.
Of the eight recorded elsewhere one has now been reduced as a synonym.
Subfamily Lygeine.
32. Nysius euphorbre, Horv.
Nysius euphorbie Hory., Ann. Mus. Hung., viii. p. 11 (1910).
Loc. Seychelles. Mahé: Cascade Estate, about 1000 feet. Bird and Dennis Islands
(Fryer).
Originally described from Mauritius:
33. Nysius albipennis, sp. n.
Head black, finely speckled with dull ochraceous; pronotum dull ochraceous with
somewhat oblique longitudinal black fascize and a transverse black fascia just before
anterior margin; scutellum dull ochraceous with a more or less well defined central
black carination; corium greyish-white and talc-like, two linear black spots on the
apical margin; membrane pale hyaline; body beneath black, legs ochraceous, posterior
femora speckled with fuscous on their apical halves, apices of tarsi black ; head about as
long as breadth between eyes, more or less finely granulate; antennz ochraceous, basal
joint black, apical joint fuscous, basal joint moderately incrassate, scarcely passing apex of
head, second joint subequal to or very slightly longer than third, third and fourth sub-
equal, the latter moderately thickened; pronotum coarsely punctate, slightly thickened ;
scutellum somewhat gibbous, coarsely punctate ; membrane passing abdominal apex.
Long. 4 to 44 mm.
Loc. Seychelles. Silhouette: high forest near Mont Pot-a-eau, ca. 1500 feet ;
Mare aux Cochons, over 1000 feet. Mahé: country above Port Glaud, 500—1000 feet,
XI. 1908; near Morne Blanc; Cascade Estate, about 1000 feet; Mare aux Cochons
district, 1000—2000 feet, I. 1909; Long Island and various localities in low country.
Aldabra: Takamaka, 1908 (Fryer).
Allied to NV. euphorbie Hory., antenne longer, first joint more incrassate, different
colour of the posterior femora; corium paler &c.
SECOND SERIES—ZOOLOGY, VOL. XVI. 20
150 PERCY SLADEN TRUST EXPEDITION
Subfamily Cymine.
34*, Ninus sechellensis, Bergr.
Ninus sechellensis Bergr., Rev. d’Ent., xi. p. 201, 1893.
Loc. Seychelles. Silhouette, low coconut-planted country near coast, IX. 1908.
Mahé: Cascade Estate, about 1000 feet, and cultivated country near sea-level; marshes
on coastal plain of Anse aux Pins and Anse Royale; Port Victoria.
Subfamily Heterogastrine.
HYGINELLUS, gen. nov.
Allied to Hyginus Stal, but differing by the long rostrum which considerably passes
the posterior coxee, and by the short and rather robust spine to the anterior femora.
A single and somewhat mutilated specimen is alone contained in the collection, but
in this faunistic enumeration it cannot be ignored. In the specific description of the type
the secondary structural characters will be detailed.
35. Hyginellus gayer, sp. n. (Plate 11, figs. 6, 6a).
?. Head, pronotum and scutellum black ; apex of scutellum and about basal half of
corium greyish-white, the latter with a short sublateral black streak, apical area of
corium black; membrane pale bronzy, the base black and its apical area piceous;
connexivum black with an elongate ochraceous lateral spot before apex; body beneath
legs and rostrum black, apex of first joint of rostrum and bases of intermediate femora
flavescent (posterior legs mutilated in type); head thickly coarsely punctate, a little
longer than length between eyes; antennze piceous, first joint shorter than head, second
about twice as long as first, remaining joints mutilated in type, second joint shortly
thickly pilose, first joint slightly thickened ; ocelli near base, nearer to eyes than to each
other; pronotum closely, coarsely punctate, transversely impressed a little before middle,
with a central longitudinal carinate line not extending beyond middle of posterior lobe,
anterior margin almost truncate, posterior margin moderately concave, lateral margins,
especially on anterior lobe slightly subampliate, posterior angles subnodulose, scutellum
with a cruciform elevation, excluding apex, closely coarsely punctate; clavus and corium
longitudinally distinctly punctate ; membrane reaching abdominal apex and with distinct
basal cells ; rostrum extending considerably beyond the posterior coxe, first joint slightly
passing base of head, second joint passing anterior coxee, third joint slightly passing
intermediate coxze ; head beneath, and sternum thickly coarsely punctate.
Long. 9 mm.
Loc. Seychelles (J. A. de Gaye).
Subfamily Pachygronthine.
36*. Pachygrontha bipunctata, Stal.
Pachygrontha bipunctata Stal, Hem. Afr., 11. p. 149 (1865).
Loc. Seychelles. Mahé: from grass in plantations &¢. near coast, not m mountains.
Originally described from Mauritius.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 151
37. Pachygrontha confusa, sp. n.
Closely allied to the preceding species P. bipunctata but differing by the longer
antenne, much longer than the body, first joint as long as head, pronotum and scutellum
together, second joint distinctly longer than third, shghtly longer than head and pronotum
together ; rostrum slightly passing the anterior cox ; in some specimens the small black
spot at about middle of apical margin to corium is distinct, in others it is absent, while
the submarginal black or piceous fascia to the abdomen beneath as in P. bipunctata is
much paler and-more testaceous in hue, or entirely absent ; sometimes the lateral margins —
of the pronotum are distinctly darkly punctate, but this is a varietal character found in
both species.
Long. 7 to 8 mm.
Loc. Seychelles. Mahé: near Morne Blanc, about 1000 feet; Port Victoria ;
Cascade Estate, 800—1000 feet; Long Island. From grass in plantations &c., mostly
near coast, not in mountains.
Bergroth recorded P. bipunctata from Mahé, and possibly overlooked the fact that
two species exist there; P. confusa in this collection is the more abundant.
Subfamily Oxycarenine.
Genus MARUTHAS.
Maruthas Dist., Faun. Brit. Ind., Rhynch., v. p. 44 (1910).
Oxycarenus Horv. (nec Fieb.), Ann. Mus. Hung., x. p. 609 (1912).
The type of this genus was received from Ceylon. The structural differences
between Maruthas and Oxycarenus were detailed (supra), and these Horvath seems to
have ignored.
38. Maruthas fumigatus, sp. n. (Plate 12, fig. 1).
Head black ; pronotum pitchy-brown with a broad transverse piceous fascia on the
anterior half and near the anterior margin, which is dull ochraceous; scutellum piceous ;
corium dull ochraceous, the lateral margins paler, membrane very pale fuliginous ; body
beneath (as best seen in unique carded type) and legs castaneous ; tibize ochraceous, their
bases and apices more or less castaneous ; antennze with the first joint black, not reaching
apex of head, second longest, ochraceous with its apex black, third and fourth black, their
bases more or less ochraceous ; head longer than broad, attenuate and obtusely angulate
anteriorly ; thickly finely granulate and pilose ; pronotum longer than broad, transversely
impressed near middle, before which it is transversely subglobose and centrally depressed,
thickly coarsely punctate, the lateral margins longly pilose ; scutellum coarsely punctate ;
membrane not quite reaching abdominal apex ; anterior femora robust and strongly spined
beneath.
Long. 4 mm.
Loc. Cosmoledo, 1907 (Thomasset).
152 PERCY SLADEN TRUST EXPEDITION
Subfamily Aphanine.
Genus CLERADA.
Clerada Sign., in Maillard, Notes sur I’Ie de la Réunion, Ins., p. 28 (1862).
Mahisa Horv. (nec Dist.), Ann. Mus. Hung., ix. p. 337 (1911).
Horvath (supra) has stated that Clerada= Mahisa. These genera, apart from other
characters, may be separated by the spined anterior femora.of the latter. Clerada, as
well known, and as long since pointed out by Stal, have ‘‘femoribus inermibus.”
39*, Clerada apicicornis, Sign.
Clerada apicicornis Sign., in Maillard, Notes sur V'lle de la Réunion, Ins., p. 28
(1862).
Loc. Seychelles. Mahé: Port Victoria.—In my previous enumeration of the
Rhynchota of the Seychelles, this species was recorded from Le Craze. Originally
described from the Island of Réunion ; recorded also from Brit. Ind. and Ceylon; Samoa ;
Mexico ; Venezuela, and West Indies.
40*. Paromius seychellesus (Walk.).
Plocwomerus seychellesus Walk., Cat. Het., v. p. 120 (1872).
Loc. Seychelles. Silhouette: low country near sea, Grande Barbe; high country
near Mont Pot-a-eau. Mahé: near Morne Blanc, about 1000 feet ; Cascade Estate, about
1000 feet ; Long Island. Also found in Ceylon, Burma and Japan.
41. Paromius apicatus (Stal).
Rhyparochromus apicatus Stal, Ofv. Vetensk.-Akad. Forh., 1855, p. 34.
Loc. Seychelles. Mahé: near Morne Blane; Long Island. From grass in
cultivated places at low elevations (Hugh Scott). Originally described from Natal and
also recorded from Mauritius.
42*, Pamera vincta.
Pamera vincta Say, New Harm. Ind., 1831.
Plociomerus reductus Walk., Cat. Het., v. p. 120.
Loc. Seychelles. Silhouette: Mare aux Cochons &c. Mahé: Cascade Estate,
Long Island &c. Mr Scott usually swept the species from grass in open or cultivated
places. I have in previous enumeration (Trans. Linn. Soc. Lond., 1909, p. 39) given the
distribution of this widely dispersed species. I now add the P. reductus Walk. to the
synonymy. Bergroth correctly surmised that this species=P. parvula Dall., which
however he treated as a distinct species and not as a synonym of P. vincta Say (Rev.
d’Ent., xii. p. 197, 1893).
43. Pamera sladeni, sp. n. (Plate 11, fig. 12).
Head, anterior lobe of pronotum and scutellum black ; pronotal collar and posterior
lobe of pronotum castaneous-brown, the latter with a more or less well marked central
DISTANT—RHYNCHOTA. PART IT: SUBORDER HETEROPTERA 153
longitudinal fascia, the anterior area and the posterior angles black; scutellum with two
discal obscure dull greyish spots and its apex pale ochraceous ; corium pale ochraceous,
claval streaks, a small sublateral spot near base, another beyond middle, a third near apex,
and a patch (containing a small pale spot) near inner angle, piceous ; membrane bronzy,
basal area black ; lateral margin pale with a brown spot; body beneath black; legs pale
ochraceous, anterior femora, excluding extreme bases and apices, more than apical halves
of intermediate and posterior femora and extreme apices of tibize and tarsi black, extreme
apices of intermediate and posterior femora, pale ochraceous; antennee brownish-
ochraceous, apices of second and third joints and fourth joint, excluding base, black, first
Joint a little shorter than head, second and fourth subequal in length, each slightly longer
than third ; head with a fine central longitudinal incision between the eyes ; lateral and
posterior margins, and an indistinct central line to anterior pronotal lobe greyishly pilose,
the anterior lobe very slightly longer than the posterior lobe; clavus longitudinally,
rather coarsely punctate ; rostrum reaching the intermediate coxee, ochraceous, basal joint
and apex piceous, first joint not reaching base of head, second about reaching anterior
cox ; anterior femora longly spined beneath.
Long. 5 to 6 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons. Mahé: country above Port
Glaud, about 500—1000 feet; near Morne Blanc, about 1000 feet; Port Victoria ;
Cascade Estate, about 1000 feet. Usually swept from grasses in open or cultivated places
(H. Scott).
44. Lachnophorus singalensis (Dohrn).
Rhyparochromus singalensis Dohrn, Stett. ent. Zeit., xxi. p. 404 (1860).
Loc. Seychelles. Mahé, from grass in cultivated country, about 1000 feet. Also
recorded from Madagascar, Bombay and Ceylon.
45. Lachnophorus albidomaculatus, sp. n. (Plate 11, fig. 9).
Head, anterior lobe of pronotum and scutellum black, lateral margins and posterior
area of pronotum pale testaceous, the latter thickly blackly punctate, a small black oblique
spot near the posterior angles; corium brownish-ochraceous with piceous mottlings, an
elongate spot in clavus, three spots outside clavus, the lowermost near inner angle with
two in transverse series beyond it, greyish-white ; membrane piceous with a spot beneath
apical area of corium and some apical macular markings greyish-white; body beneath
and legs black or dark castaneous, apices of the femora and basal joints of the tarsi
ochraceous; antennee black, first joint slightly passing apex of head, second a little longest,
third and fourth subequal and distinctly incrassate.
Long. 5 to 6 mm.
Loc. Seychelles. Silhouette: from near Mont Pot-d-eau. Mahé, above Port Glaud,
500—1000 feet. Bird Island (Fryer). Aldabra, 1907 (Thomasset).
46. Cligenes gardineri, sp. n. (Plate 11, fig. 10).
Head black, shortly greyishly pilose, apex testaceous; anterior lobe of pronotum
castaneous, posterior lobe ochraceous, darkly punctate, posterior angles castaneous ;
154 PERCY SLADEN TRUST EXPEDITION
scutellum pale castaneous, its apex ochraceous; corium pale shining ochraceous, a spot on
lateral margin near middle, the apical angle, and a spot near inner angle, black; membrane
subhyaline; body beneath castaneous, legs and rostrum ochraceous; antennee with the
first and fourth joints ochraceous, second and third joints and extreme base of fourth,
black or piceous, first joint passing apex of head, second and fourth subequal, each slightly
longer than third; rostrum slightly passing the anterior cox, first joint not reaching
base of head ; pronotum with the transverse impression slightly nearer to the posterior
than to the anterior margin ; apical margins of corium straight.
Long. 2 mm.
Loc. Seychelles. Silhouette: near Mont Pot-d-eau, ca. 1500 feet; Mare aux
Cochons. Mahé: country above Port Glaud, about 500—1000 feet ; near Morne Blane ;
Cascade Estate, 800—1000 feet ; Mare aux Cochons district, 1000—2000 feet. Praslin :
Cotes d’Or Estate. Félicité.
47*. Dimella nitida (Reut.).
Dinia nitida Reut., Ofv. Finska Vetensk. Soc. Férh., xxv. p. 24 (1832).
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet ; marshes on coastal plain
at Anse aux Pins and Anse Royale. Félicité Island. Recorded by Bergroth from La Digue.
Originally described from West Coast of Africa.
A variable species in depth of coloration. Mr Scott reports it as
of marshy places where it is swept from low vegetation.”
cé
characteristic
48. Aphanus consocialis, sp. n. (Plate 11, fig. 7).
Head black, greyishly pilose; pronotum with the anterior lobe—excluding lateral
margins—castaneous, posterior lobe ochraceous, thickly darkly punctate, lateral margins
pale flavescent with a dark spot near anterior angles and an ocellated dark spot at
posterior angles ; scutellum castaneous, with two contiguous elongate ochraceous spots on
apical area; corium ochraceous, darkly punctate, two dark spots on lateral margin, one
before and the other at apical angle ; body beneath castaneous ; legs ochraceous; antennze
brownish-ochraceous, apex of third joimt and the whole of fourth, castaneous, first joint
shghtly passing apex of head, second slightly longer than third or fourth, which are
subequal ; posterior pronotal lobe coarsely darkly punctate with a central pale impunctate
line ; scutellum slightly convex on basal area ; membrane pale brownish, the veins paler
and with a whitish spot at apex ; anterior femora spined beneath near apex, intermediate
and posterior tibize finely spinulose.
Long. 44 to 5 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons. Mahé: Long Island.
Allied to A. lineosws found in Minikoi, Ceylon and Burma.
49*. Dieuches annulatus (Sign. ).
Rhyparochromus annulatus Sign., Ann. Soc. Ent. France, 1860, p. 949.
Loc. Seychelles. Silhouette, Mahé, Dennis Island, Bird Island. Previously
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 155
received from Mahé, Coetivy, Farquhar Atoll, Amirantes. Originally described from
Madagascar.
50*. Dieuches placidus (Stal).
Beosus placidus Stal, Hem. Afr., 11. p. 171 (1865).
Loc. Seychelles. Silhouette, from near Mont Pot-a-eau, ca. 1500 feet, and Mare
aux Cochons, over 1000 feet. Mahé, high damp forest between Trois Fréres and Morne
Seychellois, 1500—2000 feet; also Cascade Estate, &c. Praslin: Cotes d'Or Estate.
Dennis Island (Fryer). Aldabra: Takamaka (Fryer). Originally described by Stal from
Bourbon.
51. Dieuches cardwu, sp. n. (Plate 11, fig. 13).
Head black, its apex ochraceous; pronotum anteriorly castaneous, posteriorly
ochraceous, thickly coarsely darkly punctate, anterior margin (centrally broken), lateral
and posterior margins, and a central pale line on posterior half, ochraceous, a central
longitudinal line on anterior half black ; scutellum castaneous, with two elongate spots on
apical area and the apex ochraceous ; corium brownish-ochraceous, the lateral margins
bright pale ochraceous, with a castaneous spot behind middle and the apex and apical
margin black or piceous; membrane pale bronzy-brown, the apical margin paler with
greyish spots ; head beneath and sternum black, sternal margins, coxee, trochanters and
legs ochraceous; abdomen beneath pale castaneous, the lateral margins ochraceous;
antennee ochraceous, apices of first, second and third joints, and apical half of fourth joint,
black or piceous, first joint about as long as head, remaining joints almost subequal in
leneth, each about as long as pronotum; scutellum very finely punctate on basal area ;
corium (except on lateral margins) longitudinally darkly punctate; membrane passing
abdominal apex; rostrum ochraceous, almost reaching the intermediate coxee, first jot
reaching base of head ; anterior femora finely spined beneath; pronotum constricted near
middle.
Long. 7 to 8 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons plateau, over 1000 feet. Mahé:
from near Morne Blanc; Cascade Estate, about 1000 feet; Mare aux Cochons district,
1000—2000 feet. Praslin.
52*,. Letheus punctus, Bergr.
Letheus punctus Bergr., Rev. d’Ent., xii. p. 203 (1893).
Loc. Seychelles. Mahé: from near Morne Blane, about 1000 feet ; Cascade Estate,
about 1000 feet ; Long Island. Previously received from Praslin.
53. Letheus stellatus, sp. n. (Plate 11, fig. 8).
Black ; two small contiguous spots at each posterior pronotal angle, two small spots
on inner margin of clavus (one before middle, the other at apex), apices of femora, tibize
and tarsi ochraceous; lateral margins of corium very narrowly ochraceous ; membrane
greyish-brown, paler at base; antennze piceous-brown, the basal joint piceous, second joint
156 PERCY SLADEN TRUST EXPEDITION
longest, third and fourth subequal in length, first shortest ; head—excluding basal
margin—thickly finely punctate, apex reaching middle of first antennal joint ; pronotum
with the lateral margins moderately ampliate, anteriorly punctate behind head, posterior
lobe very coarsely punctate; scutellum centrally depressed, thickly rather coarsely
punctate; corium thickly finely punctate, clavus longitudinally punctate ; membrane
about reaching abdominal apex; rostrum reaching the intermediate coxe.
Var. Two small rounded spots on corium almost in line with those on clavus, and a
small spot on apical margin of corium, ochraceous. In some specimens the ochraceous spots
are almost entirely missing.
Long. 6 to 7 mm.
Loc. Seychelles. Mahé: high forest of Morne Blanc. Aldabra: Takamaka (Fryer).
Cosmoledo, 1907 (Thomasset). Assumption, 1910 (Dupont).
Allied to L. punctus Bergr., but a more elongate species, posterior lobe of pronotum
less coarsely punctate ; scutellum more finely punctate; rostrum not passing the inter-
mediate coxee ; corium darker, less irrorated with ochraceous, and pale margin much
narrower.
ORBELLIS, gen. nov.
Head somewhat elongate, immersed to eyes which scarcely pass the anterior angles
of the pronotum, acuminate in front where the central lobe is elevated, ocelli near eyes
and a little removed from base; antennze only a little shorter than the body, first joint
about as long as head but considerably passing its apex, third joint a little shorter than
second, subequal to fourth, first moderately and gradually thickened from near base to
apex ; pronotum somewhat flat, oblique, posterior margin about twice as broad as anterior,
the latter slightly concave, the former truncate, lateral margins almost straightly oblique
and slightly ampliate, obscurely transversely impressed before middle, where it has the
appearance of a slightly convex lobe; scutellum about as long as broad, a little foveately
depressed at base; corium with the lateral margins subparallel and slightly but distinctly
reflexed, apical margins straight ; membrane scarcely passing the abdominal apex, venation
forming a small transverse cell near base; legs somewhat long, femora moderately
thickened, anterior femora with a few short spines near apex, tibiee finely spinulose, first
joint of tarsi longer than remaining joints together ; rostrum imperfectly seen in unique
carded type.
Allied to Letheus, but differing in the longer antennze and pronotum &e.
54. Orbellis typicus, sp. n. (Plate 11, fig. 5).
Head brassy-black, its apex testaceous; pronotum pale castaneous, the lateral
margins, anterior margin (centrally broken and not reaching anterior angles), and basal
margin (inwardly angulate), ochraceous, posterior area more or less suffused with black ;
scutellum pale castaneous ; corium pale bronzy-brown, the lateral margins narrowly pale
ochraceous, with some irregular darker spots, the most prominent of which is an elongate
discal black spot behind middle; membrane pale bronzy-brown ; body beneath and femora
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 157
castaneous, tibize and tarsi ochraceous ; antennee with the first and second joints brownish-
ochraceous, their apices distinctly darker, third black with nearly apical half pale
ochraceous, fourth black ; scutellum thickly punctate, the punctures a little darker on the
basal foveate impression ; clavus thickly longitudinally punctate ; corium thickly punctate
except on the pale lateral margins ; other structural characters as in generic diagnosis.
Long. 6 mm.
Loc. Seychelles. Mahé: Cascade Estate, forest above 1000 feet, I 1909.
[55*. Budeus corcumcinctus Walk. (Rhyparochromus), Cat. Het., v. p. 97 (1872).
I am still unable to discover the fully developed form of this immature species. |
Family Pyrrhocoride.
Subfamily Pyrrhocorine.
Four species belonging to this family were enumerated in my previous list, two
collected by Prof. Gardiner and two recorded elsewhere. All these were collected by
Mr Scott and one other which requires generic distinction.
56*. Scantius forsterr (Fabr.).
Cimex forstert Fabr., Spec. Ins., i. p. 368 (1781).
Loc. Cosmoledo and Astove, 1907 (Thomasset). Assumption, 1908 (Fryer).
In the previous collection, as recorded, received from Farquhar Atoll.
JOURDAINANA, gen. nov.
_ Head about as long as breadth between eyes, elongated anteriorly, the central lobe
robust and prominent ; antennze robust, first and fourth joints more robust and almost
subequal, first slightly passing apex of head, third shortest, second longest; rostrum
reaching the intermediate coxz, first joint not quite reaching base of head ; pronotum
nearly twice as broad at base as long, the lateral margins ampliate and reflexed, very
coarsely and rugosely punctate, crossed by a transverse raised impunctate fascia a little
before middle ; scutellum small, its basal area coarsely punctate, the apical area laterally
and apically raised and impunctate ; corium moderately broad, its lateral margins sub-
parallel and a little acutely reflexed, thickly punctate, apical margins almost obliquely
straight, membrane reticulately veined, not passing abdominal apex, legs moderate in
length, anterior femora thickened, very shortly spinous beneath, anterior tibize apically
thickened, basal tarsal joint longest, on posterior tarsi as long as remaining joints
together.
This genus is named after John Jourdain, who sighted Silhouette on January 19,
1609.
57. Jourdaimana rugifer, sp. n. (Plate 11, fig. 14).
Head and antennz black; pronotum piceous, the discal transverse raised fascia black,
the lateral margins ochraceous ; scutellum black, the apical area piceous ; corium pitchy-
SECOND SERIES—ZOOLOGY, VOL. XVI. 21
158 PERCY SLADEN TRUST EXPEDITION
brown, thickly finely punctate, the lateral margins ochraceous and very sparsely punctate;
membrane black; sternum black; abdomen beneath testaceous, the apical segmental
joint and anal appendages black ; legs dark castaneous or piceous, extreme apices of the
anterior and intermediate femora, ochraceous; head very finely and obscurely wrinkled
with a slight central discal longitudinal incision, lateral margins of the pronotum
impunctate. Other structural characters as in generic diagnosis.
Long. 6 mm.
Loc. Seychelles. Mahé, 1907 (Thomasset). A single example.
58*. Dysdercus ortus, Dist.
Dysdercus ortus Dist., Trans. Linn. Soc. London, ser. 2, Zool., xiii. p. 36, Pl. 4, f. 13
(1909).
Loc. Seychelles. Mahé; Long Island, VII.1908. Bird and Dennis Islands, 1908
(Fryer). In my previous enumeration recorded from Praslin and Cousin Island.
59*. Dysdercus fasciatus, Sign.
Dysdercus fasciatus Sign., Ann. Soe. Ent. France (8), viii. p. 954 (1861).
Loc. Aldabra, 1908 (Fryer). Cosmoledo, 1907 (Thomasset).
60*. Dysdercus mgrofasciatus, Stal.
Dysdercus nigrofasciatus Stal, Ofv. Vetensk.-Akad. Forh., 1855, p. 36.
Loc. Seychelles. Dennis Island, 1908 (Fryer).
Family Tingidide.
The two species here recorded are the first received from the Seychelles; one belongs
to a genus found in Ceylon, the other constitutes a new genus with neotropical affinities.
61. Dulinius nigrolineatus, sp. n. (Plate 11, fig. 16).
Pale hyaline; venation pale fuscous; extreme edges of the reflexed concave
membranous margins to the pronotum somewhat suffused with ochraceous ; margins and
two central lines to pronotum black ; antennze very finely pilose, first joint moderately
thickened and extending to about apex of head, apical joint distinctly darker; lateral
margins of the pronotum laminately produced, reflexed and concave, the extreme edge
recurved ; hood large, globular, covering the head ; elytra a little more than twice as long
as abdomen, obliquely widening from near base, the apical margins rounded.
Long. body 2, to apex of elytra 34 mm.
Loc. Low coconut-planted country near the coast, Pointe Etienne.
A single specimen from Silhouette forms a second species of the genus which was
founded to contain a Ceylonese species.
CANTINONA, gen. nov.
Elongate ; pronotum without an anterior vesicle or hood; antennz long, finely
pilose, first and second joints short, a little thickened, second a little shorter than first,
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 159
third very long, fourth distinctly longer than first and second together; head sub-
quadrate, three short porrect spines between the antennal bases and a blunt robust spine
outside the antennal bases, eyes close to the anterior margin of the pronotum ; pronotum
with the anterior margin subtruncate, the lateral margins laminately convexly dilated,
the posterior margins longly, angularly, posteriorly developed, the anterior area tricarinate,
the central carination continued to the posterior angle, the lateral marginal areas sub-
hyaline, bi-areolate ; elytra with the discoidal area long and moderately slender, closely
finely areolate, subcostal area small and ill-defined, costal area broad, somewhat obliquely
widened at base, the lateral margins subparallel, apical margins rounded, areolate ; sutural
area ill-defined ; legs long and slender.
Allied to Gelchossa Kirk, n. n. = Leptostyla Stal preoce.
This genus is named after Alberto Cantino, on whose charts the Seychelles appeared
in 1502.
62. Cantinona precellens, sp. n. (Plate 11, fig. 15).
Head black, apex of anterior central spine ochraceous; antenne black, first joint
flavescent, apical area of third and the whole of fourth obscure ochraceous ; pronotum
black, the anterior margin ochraceous, longitudinal carinations brownish-ochraceous, lateral
marginal areas subhyaline with the margins of the areolets black; elytra with the
discoidal area black, narrow subcostal area obscure ochraceous, costal area subhyaline, the
margins of the areolets ochraceous, extreme costal margin black, from the apex of
the discoidal area a curved fasciate series of black margined areolets to apical margin
of costal area; legs pale ochraceous, tarsi black. Structural characters as in generic
diagnosis.
Long. 33 mm.
Loc. Seychelles. Mahé: Cascade Estate, ca. 1000 feet, 1909 ; Mare aux Cochons
district, 1500—2000 feet, I. 1909. Félicité, 1. XII. 1908.
Three specimens examined, two of which are typical, agreeing with the figure, the
third is a pale variety in which the black coloration is replaced by pale brownish-
ochraceous.
Family Aradide.
Four species of Aradidz are now enumerated from Seychelles, only one having been
previously recorded.
63*. Prctinus mvalidus, Bergr.
Pictinus mvalidus Bergr., Entomol. Tidskr., xv. p. 109 (1894).
_ toe. Seychelles. Silhouette: forest near Mont Pot-d-eau, ca. 1500 feet; Mare aux
Cochons and forest above; low coconut-planted country near coast, Pointe Etienne.
Mahé: high forest of Morne Blanc and Pilot; high damp forest between Trois Fréres and
Morne Seychellois, 1500—2000 feet ; slopes of Morne Seychellois, 1500—2000 feet; Mare
aux Cochons district, 1500—2000 feet; Cascade Estate. Praslin: Cotes d’Or Estate.
21—_2
160 PERCY SLADEN TRUST EXPEDITION
Félicité, from forest. Originally recorded from La Digue. Mr Scott collected a large
series, almost entirely from the endemic forests. The majority of specimens were found
under bark, but some were found between the leaf-bases of a growing Stevensonia palm
near Morne Blane, and several others in a similar situation on the summit of Mount
Sebert, 28. XII. 1908.
64. Pictinus varicolor, sp. n. (Plate 11, fig. 17).
Black; base of membrane irregularly, broadly pale flavescent; connexivum
testaceous; legs castaneous; sternum more or less piceous; abdomen beneath casta-
neous; antennze piceous, first joint strongly incrassate, shorter than head, second joint
moderately incrassate, attenuate at base, a little shorter than first, third longest and
more slender, fourth short, thick, pyriform, about as long as second; pronotum broader
than long, the lateral margins obliquely sinuate, obtusely angularly toothed before
middle, the posterior angles subprominent and subnodulose, anterior lobe with four
prominent tubercles arranged in pairs, between which is a longitudinal ridge, posterior
margin prominent; scutellum strongly ridged laterally and centrally; corium shorter
than scutellum; membrane with venation indiscernible ; body beneath imperfectly seen
in the carded unique type.
Long. 3 mm.
Loc. Seychelles. Mahé: Cascade Estate 800—1000 feet.
65. Newroctenus caffer (Stal).
Brachyrhynchus caffer Stal, Ofv. Vetensk.-Akad. Forh., 1855, p. 38.
Loc. Seychelles. Silhouette, Mare aux Cochons. Mahé, Cascade Estate, ca. 1000
feet. Not uncommon in South Africa and recorded from Madagascar and Rodriguez.
66. Aneurus bicolor, sp. n. (Plate 11, fig. 18).
Black ; central lobe to head, elevations to anterior lobe of pronotum, hemelytra,
connexivum, abdomen beneath, tibize and tarsi castaneous; antenne black, first joint
short, strongly incrassate, not reaching apex of head, second and third joints less
incrassate than first, short, subequal in length, fourth longest, cylindrical; head rugosely
punctate, the separation of the central lobe incised, pronotum broader than long, anterior
lobe with a flattened transverse elevation on each side, lateral margins moderately
sinuate ; scutellum a little broader than long, its apex broadly rounded ; corium a little
longer than scutellum; membrane not reaching abdominal apex; connexivum broad,
moderately upturned; membrane with indiscernible venation, reflecting the castaneous
coloration beneath.
Long. 34 to 4 mm.
Loc. Seychelles. Mahé: from near Morne Blane. Praslin: from Coco-de-mer
forest in the Vallée de Mai, Cétes d’Or Estate, XI. 1908.
A species of this genus, A. brevisculatus Bergr., has been described from Madagascar,
but the structure of the scutellum alone is sufficient to differentiate the species. It is
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 161
more nearly allied to A. green? Dist. from Ceylon, but from this species it differs in the
shorter antennze, narrow pronotum «ec.
Family Hydrometride.
The one species brought home by Prof. Gardiner, and recorded in my previous paper,
has again been found by Mr Hugh Scott.
Subfamily Hydrometrine.
67*. Hydrometra ambulator, Stal.
Hydrometra ambulator Stal, Ofv. Vetensk.-Akad. Forh., xii. p. 45 (1855).
Loc. Seychelles. Silhouette: from marshy plateau of Mare aux Cochons, over
1000 feet, and from the lower part of a stream near the coast at Anse Moudon. Mahé:
from near Morne Blanc, about 1000 feet ; marshes on coastal plain of Anse aux Pins and
Anse Royale; Mare aux Cochons district, 1500—2000 feet. Previously recorded from
Mahé. Found in 8. Africa.
Subfamily Veliine.
Three species of this subfamily are now enumerated, including the one listed
previously.
68*. Rhagovelia nigricans (Burm.).
Velia nigricans Burm., Handb. xi. p. 213 (1835).
Loc. Seychelles. Silhouette: marshy plateau of Mare aux Cochons, over 1000 feet,
and from rivers near coast. Mahé: from near Morne Blanc; Mare aux Cochons district,
1000—2000 feet; Port Victoria. Formerly recorded from Praslin and then stated to be
a very widely distributed species.
PICAULTIA*, gen. nov.
Apterous form, °. Head almost as long as breadth, including eyes, which are
prominent, distinctly separated from base of head and slightly projecting beyond the
anterior angles of the pronotum; antennz four-jointed, hirsute, first and second joints
a little stoutest, first passing apex of head and a little longer than second ; rostrum
passing the anterior, but not reaching the intermediate cox; pronotum strongly trans-
versely impressed near middle creating somewhat the appearance of an anterior lobe,
which is a little convex and possesses a slender curved spine near each end of the incision,
the lateral margins hirsutely pilose, the posterior margin rounded, an oblique piece of the
mesonotum exposed on each side and near its apex a slender curved lateral spine is
also exposed ; connexivum very long and broad, upturned, converging posteriorly ; legs
moderately long, femora straight, intermediate and posterior tarsi long, subequal and
two-jointed. The spines and general structure of the pronotum are the salient
characters of this genus.
* Named after Captain Lazare Picault who commanded an expedition to these islands, 1742.
162 PERCY SLADEN TRUST EXPEDITION
69. Picaultia pronotalis, sp. n. (Plate 11, fig. 19).
Apterous form, ?. Black; anterior transverse fascia to pronotum, margins of con-
nexivum, and legs ochraceous; apices of femora, the tibize and tarsi more or less
infuscate ; antennze brownish-ochraceous, longly pilose, first joint a little longer than
second, third and fourth almost subequal in length; pronotum centrally longitudinally
carinate, the whole body more or less pilose.
Long. 2 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons plateau, over 1000 feet,
VIII.—IX. 1908. All the specimens received are apterous and the mature form has
yet to be described. A female specimen is figured.
70. Microvelia repentina, Dist. (?) (Plate 11, figs. 20, 21).
Microvelia repentina Dist., Faun. Brit. Ind., Rhynch., xi. p. 174, fig. 127 (1904).
Apterous form, ¢. Black; pronotum castaneous, with a large black spot on each
anterior lateral margin, and a transverse ochraceous fascia near anterior margin; legs
ochraceous, the apices of the femora and the tibiz more or less piceous (Plate 11,
fig. 20).
Apterous form, §. Head and pronotum castaneous, posterior pronotal area ochra-
ceous ; abdomen above black, connexivum and legs ochraceous; apices of femora and the
tibize more or less infuscate (Plate 11, fig. 21). .
Long. 15 mm.
Loc. Seychelles. Mahé: country above Port Glaud, about 500—1000 feet.
Silhouette : Mare aux Cochons plateau, over 1000 feet. Calcutta, Madras.
This species, which I originally described from Calcutta, I have since received from
Madras, collected by Capt. W. S. Patton, I.ML.S., in all its stages of development.
Although I have not seen the imago from the Seychelles, this collection still contains
numerous immature specimens which I cannot differentiate from those which I received
from Madras and which are here figured. The mature form which I described and
figured had the legs nearly uniformly ochraceous, but I have since seen many specimens
from Calcutta and Madras in which the apices of the femora and the tibize were infuscate
or piceous.
Subfamily Gerrine.
Division Gerrinaria.
Mr Scott's collection only contains the two species previously recorded.
71*. Gerris (Limnogonus) cerevventris, Sign.
Gerris cereiventris Sign., in Maillard, Notes sur Vile de la Réunion, Ins., p- 30
(1862).
Loc. Seychelles. Silhouette: marshy plateau of Mare aux Cochons, over 1000 feet,
and rivers near coast. Mahé: from near Morne Blanc; marshes on coastal plain of Anse
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 163
aux Pins and Anse Royale; Mare aux Cochons district, 1000—2000 feet. Previously
taken in Praslin.
72*,. Gerris (Limnogonus) dolosa (Bergr.).
Limnogonus dolosus Bergr., Rev. d’Ent., 1893, p. 204.
Loc. Seychelles. Mahé: from near Morne Blanc; marshes on coastal plains at
Anse aux Pins and Anse Royale; Long Island; Cascade Estate. Silhouette: Mare aux
Cochons. Previously recorded from La Digue and Coetivy.
Division Halobatinaria.
73*. Halobates alluaudi, Bergr.
Halobates alluaudi Bergr., Rev. d’Ent., 1893, p. 204.
Loc. Seychelles. Mahé: Long Island. Aldabra (Fryer). In addition to the other
localities (Chagos Islands, Amirantes, Coetivy &c.), which I previously enumerated (Tr.
Linn. Soc., ser. 2, Zool., xiii. p. 37), where this species had been obtained, it has also
been taken in the Red Sea, Port Sudan. At Long Island, Mahé, Mr Scott took this
species at “low water, small ones (immature) together near shore. Big ones (mature)
farther out.”
Family Henicocephalide.
74. Henicocephalus, sp.
A single immature example of this genus was contained in this collection, and,
though not enabling identification to be made, it must be included in the enumeration.
Unfortunately this specimen, in a subsequent attempt to remount it, was
irretrievably injured.
Loc. unrecorded : the specimen was almost certainly from one of the islands of the
Seychelles group.
Family Reduviide.
The total number of species belonging to the Reduviide as previously recorded was
six only, the present enumeration brings the number to 18; one species however,
Mascaregnasa typica Dist., previously found by Prof. Gardiner, is not contained in
Mr Scott’s collection.
Subfamily Emesine.
75. Ploiariola scotti, sp. n. (Plate 12, fig. 2).
Head anteriorly black, posteriorly fuscous-brown with the lateral margins black ;
antennee creamy-white with numerous black spots or annulations; pronotum with the
anterior lobe piceous with two longitudinal ochraceous spots, the segmental margins
distinet, the posterior lobe fuscous-brown, with longitudinal spots and the basal margin
} ochraceous ; abdomen above castaneous, darker at base, anterior legs black, spotted and
164 PERCY SLADEN TRUST EXPEDITION
annulated with ochraceous, the coxee broadly ochraceous at base; intermediate and
posterior legs creamy-white with black spots or annulations; hemelytra fuscous-brown,
with the venation and numerous spots creamy-white, these spots are larger and more
prominent on the costal margin ; wings greyish-white, the venation brownish ; first joint
of antennee a little shorter than intermediate femora, second almost as long as first ;
anterior area of pronotum very much shorter than posterior area; pronotal lateral
margins nearly obliquely straight.
Long. incl. hemelytra 54 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908.
Mahé: Long Island, VII. 1908; Cascade Estate, ca. 1000 feet.
From the description this species seems to be allied to P. morstalli Schum. from
German East Africa. It is certainly allied to P. mexta Dist. from Ceylon, from which it
differs in the shorter hemelytra &c.
76. Calphurma reticulata, Dist.
Calphurma reticulata Dist., Ann. & Mag. Nat. Hist., (8) iii. p. 503 (1909).
Loc. Seychelles. Mahé: Cascade Estate, ca. 1000 feet. A single specimen
agreeing perfectly with the typical specimens received from Calcutta and Ceylon.
CALPHURNIOIDES, gen. nov.
Allied to Calphurnia but differmg by the longer antenne and different proportional
length of joimts, the longer anterior pronotal lobe, and the non-spinulose legs. Cal-
phurma (2) aberrans Dist. from Ceylon will also be included in this genus. I pointed
out the structural differences (Faun. Brit. India, Rhynch., v. p. 176) and stated that
I had “ provisionally included it in Calphurnia.”
Another character is a long erect spine at the apex of the scutellum, which is
probably present also in Calphurnia. All my specimens of that genus were and are
pinned through the scutellum, the spine being thus completely mutilated ; carded
specimens in this collection enable a better diagnosis to be made. Calphurnia aberrans
Dist. was also similarly mutilated.
77. Calphurmoides elongatus, sp. n. (Plate 12, fig. 15a).
Body and legs dull pale ochraceous; anterior femora annulated with pale fuscous-
brown ; eyes black ; hemelytra spotted and speckled with brownish-ochraceous, a distinct
black spot on disk of membrane ; antennze with the first joint a little longer than the
intermediate femora, second shorter than first but much longer than third; head
narrowed at base but not distinctly pedunculate, strongly transversely impressed
between eyes, postocular much longer than anteocular area; pronotum elongate, the
lateral margins piceous, anterior lobe more than half the length of posterior lobe, the first
with the lateral margins convex, the anterior angles moderately prominent, its disk
longitudinally excavate, posterior lobe widened posteriorly, centrally longitudinally
suleate, darkly transversely ridged near posterior margin; scutellum strongly erectly
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 165
spined near apex; intermediate femora with their apices moderately infuscated; the
transverse veins to the corium distinct and darker in hue; membrane palely mottled with
brownish-ochraceous, the venation distinct ; legs not spinulose.
Long. 7 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908.
78. Stenolemus madagascariensis (Westw.) ?
Plearia madagascariensis Westw., Proc. Ent. Soc. London, 1844, p. 113.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet.
A single immature specimen of Stenolemus appears to represent’ Westwood’s species.
79. Luteva malayana, Dist.
Luteva malayana Dist., Faun. Malay Zool., i. p. 258, Pl. 16, fig. 2 @ (1908).
Loc. Seychelles. Silhouette: from near Mont Pot-a-eau, ca. 1500 feet. Mahé:
Cascade Estate, ca. 800 feet; Long Island. Feélicité. Originally described from the
Siamese Malay States, and since received from Ceylon. Only one mature specimen is
contained in the collection.
80. Gardena seychellensis, sp. n. (Plate 12, fig. 5).
Pitchy-brown head in front of eyes and apical area of abdomen above more piceous ;
anterior femora (excluding apices) and basal areas of anterior tibize paler and more
ochraceous in hue; antennee with the first joint as long as abdomen and about as long as
second joint; anterior femora (including bases and apices) finely and somewhat longly
spined beneath; anterior tibia about half the length of the femora; head strongly
transversely impressed between eyes, posterior lobe longer than anterior and strongly
attenuated at base; pronotum about as long as anterior tibiz, attenuated at base,
anteriorly broadened and moderately dilated on each side; posterior legs longer than
intermediate legs, apices of posterior femora and bases of posterior tibiz annulated with
pale ochraceous.
Long. excl. legs 15 to 16 mm.
Loc. Seychelles. Mahé: 2 specimens obtained by shaking fallen dead leaves of
Stevensonia palm in forest at foot of Morne Blanc, ca. 800 feet, 16. IX. 1908; Cascade
Estate, ca. 1000 feet, 1909, 1 specimen.
RosLania*.
Head not or very slightly narrowed at base, very slightly transversely impressed
between eyes, anteocular and postocular areas nearly subequal in length; rostrum
reaching the anterior coxe, first joint not reaching eyes; antennz long, slender, first
joint nearly as long as intermediate femora, subequal to second; pronotum a little shorter
than anterior coxee, very slightly broader than base of head and attenuated posteriorly,
mesonotum very strongly attenuated anteriorly, discally depressed from a little behind
* Named after M. du Roslan who headed an expedition to the Seychelles in 1769.
SECOND SERIES—ZOOLOGY, VOL. XVI.
iS)
bo
166 PERCY SLADEN TRUST EXPEDITION
apex ; apterous ; abdomen above with the lateral margin recurved and ridged ; anterior
femora distinctly longer than the anterior coxze, spined beneath on their anterior halves ;
anterior tibize less than half the length of the femora; anterior tarsi single-jointed, only
moderately shorter than the tibiz; intermediate and posterior legs long and slender,
tibize a little longer than the femora, posterior legs longer than the intermediate legs ;
posterior femora about as long as the abdomen ; intermediate and posterior tarsi two-
jointed, first jomt moderately globose.
Allied to Ischnonyctes Stal.
81. Roslania insularis, sp. n. (Plate 12, fig. 7).
Head and thorax pale ochraceous; eyes black, lateral margins of head and thorax
narrowly blackish ; antennze fuscous; abdomen above darker ochraceous, the segmental
margins distinct ; anterior legs pale ochraceous, a broken subapical annulation to femora,
annulations to tibiz, and tarsi—excluding bases—black ; intermediate and posterior legs
fuscous, more or less annulated with ochraceous; structural characters as in generic
diagnosis.
Long. excl. legs 18 mm.
Loc. Seychelles. Praslin, XI. 1908.
Subfamily Saicinee.
82. Polytowus modestus, sp. n. (Plate 12, fig. 4).
Pale ochraceous; head pale testaceous; antennee fuscous; the tarsi and apices of
tibize piceous; head transversely impressed behind eyes, posterior lobe with the lateral
margins distinctly ridged and convexly rounded; antennze longly pilose, first jomt a
little longer than head and pronotum together; pronotum with the anterior lobe longer
than the posterior, discally sulcate, lateral spines to posterior lobe shorter than pro-
notum and a little upwardly and forwardly directed, anterior angles distinctly
prominent; hemelytra scarcely passing the abdominal apex; scutellum armed with two
long slightly curved spines, the longer at base, the shorter at apex; rostrum with the
first joint slightly passing eyes; legs longly pilose, the anterior femora beneath more
rigidly setose; anterior tibize distinctly curved and apically very slightly dilated.
Long. 54 mm.
Loc. Seychelles. Silhouette; near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908.
Subfamily Stenopodine.
83*. Oncocephalus angulatus, Reut.
Oncocephalus angulatus Reut., Act. Soc. Sc. Fenn., xii. p. 696, t. 1, f. 11 (1883).
Loc. Seychelles. Mahé; 1908—9, 1 specimen; 1888, 1 specimen (J. J. Lister).
Also recorded from Mauritius and Johanna Island, Comoro Group.
84*, Oncocephalus sordidus, Stal.
Oncocephalus sordidus Stal, Ofv. Vetensk.-Akad. Férh., 1855, p. 44.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA Loe
Loc. Seychelles. Silhouette, VIII. 1908. Previously recorded from Mahé. Found
also in South and East Africa, Madagascar, and Réunion.
Subfamily Acanthaspidine.
85*. Conorhinus rubrofasciatus (de Geer).
Cimex rubrofasciatus de Geer, Mém., ili. p. 349, t. 35, fig. 12 (1778).
Loc. Seychelles (no particular locality given), R. P. Dupont and J. A. de Gaye.
As previously stated, outside the Palearctic region this species is very widely
distributed.
Subfamily Echtrichodiine.
QUINSSYANA, gen. nov.*
Subelongate; head broad, scarcely extending beyond anterior margins of eyes,
anterior margin with a short central porrect spinous prolongation, sharply transversely
constricted behind eyes, from which to base it is moderately convexly gibbous both above
and beneath, the base distinctly constricted; antennee five-jointed, longly pilose, first jomt
much longer than head; rostrum short, first and second joints subequal in length; pro-
notum twice broader at base than the anterior margin, transversely constricted before
middle, the anterior lobe narrow, convexly raised, centrally foveate, the posterior lobe
longer than anterior, strongly convexly raised, strongly foveate near its anterior margin,
the posterior angles rounded and subprominent inwardly determinated by a sharp longitu-
dinal constriction; scutellum with the lateral and posterior margins sharply ridged, the
apical spinous angulations prominent and very slightly curved; corium considerably shorter
than membrane which is long and apically a little widened, its apical margin convexly
rounded; legs moderately slender, femora more or less thickened, anterior tibize distinctly
dilated at apices. Structure of abdomen beneath concealed in the carded specimens from
which this diagnosis is taken.
Allied to Antiopula Bergr. (Antiopa Stal n. preoce.), differmg in the five-jointed
antennee, first joint longer than head which is more truncate anteriorly.
86. Quinssyana typrcalis, sp. n. (Plate 12, fig. 6).
Head, pronotum and scutellum shining black; basal constriction to head, anterior
margin of pronotum, posterior margin and apical spinous angulations to scutellum,
pale ochraceous; corium brownish-ochraceous, paler at base; membrane pale piceous;
legs and rostrum ochraceous, femora moderately infuscated at apices; antennz with
the first, second and third joints piceous, fourth and fifth pale ochraceous, first and
second subequal in length, third, fourth and fifth much shorter and also subequal ; femora
slightly thickened, anterior tibize moderately inwardly dilated at apices. Other structural
characters as in generic diagnosis.
* Named after M. Le Queau de Quinssy, an old administrator of the islands, whose tomb still stands in
the Government House garden at Mahé.
22—2
168 PERCY SLADEN TRUST EXPEDITION
)
Long. 3 mm.
Loc. Seychelles. Mahé: from forest of rather stunted “ Capucin” (Northea) trees,
summit of “Montagne Anse Major,” ca. 2000 feet, 1. II. 1909, 2 specimens.
87. Quinssyana varicolor, sp. n.
Head and posterior lobe of pronotum shining black; basal constriction to head,
anterior lobe of pronotum and scutellum ochraceous; corium and membrane piceous;
corium outwardly margined with brownish-ochraceous; legs ochraceous, femora more or
less apically infuscated; antennee with the first three joints obscure ochraceous, remainder
mutilated in type, first and second joints subequal in length.
Long. 3 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, VIII.—IX. 1908,
2 specimens.
This species apart from its distinct coloration differs from @. typicalis by the
proportionately longer antenne, the first and second joints being distinctly longer than
those in the typical species; the pronotum is also a little broader and the body generally
more robust.
88. Qumnssyana funeralis, sp. n.
Black ; trochanters obscure, ochraceous ; antennz with the first, second and third
joints black, fourth and fifth joints more or less ochraceous, base of fourth infuscate,
basal area of first more or less brownish-ochraceous, first and second joints subequal
in length, first a little but distinctly thickened at apex, all the joints strongly pilose,
third a little longer than fourth or fifth; rostrum brownish-ochraceous; apical spimous
angulations to scutellum ochraceous, a little obliquely raised.
Long. 4 mm.
Loc. Seychelles. Silhouette: near Mont Pot-a-eau, VIII. 1908, 1 specimen.
RocHoniA™, gen. nov.
Head about as’ long as pronotum, robustly elongate, the lateral lobes appearing as
covering flaps and extending to antennal bases, their inner margins almost meeting,
their apical margins roundly oblique, beyond these the central lobe is considerably
exserted and strongly centrally longitudinally ridged, its lateral margins sinuately
oblique, its apex subacute, anteocular area much longer than postocular, the latter behind
eyes distinctly gibbous both above and beneath; eyes large but not prominent, ocelli
behind eyes, base moderately constricted; rostrum robust, first joint slightly shorter than
the second, first joint passing insertion of antennz but not reaching eyes, second joint
almost reaching base of head; antennze longly strongly pilose, first joint about as long as
pronotum, about equal to second, third short, remainder mutilated in type; pronotum
with the anterior lobe shorter than the posterior, rugosely wrinkled, anterior margin
truncate, anterior angles subprominent, lateral margins convex, its posterior disk with
* Named after the Abbé Rochon who accompanied M. du Roslan in his expedition to the Seychelles (1769).
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 169
a very strong central sulcation which extends through half the length of posterior lobe
which has its lateral margins rounded and gibbous; scutellum with two apical spinous
angulations which are moderately curved; corium short, membrane about as long as head
and pronotum together, not quite reaching abdominal apex; connexivum prominent and
upwardly reflexed; legs moderately robust, femora thickened, anterior and intermediate
femora more strongly so; anterior tibiz distinctly dilated at apices.
By the very distinct structure of the head this genus has some affinity with
Vilius Stal; further comparison is however precluded by the mutilated antennze in the
unique type.
89. Rochonia galeatus, sp. n. (Plate 12, fig. 3a).
Head, pronotum and scutellum black, somewhat shining ; posterior pronotal lobe
with a small ochraceous spot near its anterior angles; antennze castaneous, antenniferous
tubercles ochraceous ; corium dull piceous, membrane black, opaque ; connexivum
ochraceous with large black spots; body beneath shining black, lateral margins of
abdomen with ochraceous spots; legs black, anterior and intermediate legs with the
apices of the femora and nearly basal halves of tibize ochraceous; posterior legs with
apices of femora and the whole of tibize more or less ochraceous; bases of intermediate
and posterior femora ochraceous; structural characters as in generic diagnosis.
Long. 13 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, VITI—IX. 1908.
Subfamily Harpactorine.
Genus NAaGusta.
Nagusta Stal, Ofv. Vetensk.-Akad. Férh., 1859, p. 374; id., Hem. Aft., ii. p. 59
(1865); id, Kn. Hem., iv. p. 7 (1874).
This genus, well and adequately described by Stal, is found in the Palearctic,
Ethiopian and Oriental regions. Its distribution has been rather obscured by some
recent and perhaps careless criticism. Thus Schouteden (Rev. Zool. Afric., i. p. 113
(1912)) has recently complained of the short description I have given to a species which
he refers to as N. pallida Dist. I know nothing of such a species, nor have I described
such an one.
90. Nagusta maura, sp. n. (Plate 12, fig. 8).
Head and anterior lobe of pronotum pitchy-brown, posterior lobe of pronotum,
scutellum and corium purplish-brown; membrane greyish with a metallic sheen, the
venation darker; body beneath brownish, more or less thickly cretaceously tomentose ;
legs testaceous, anterior femora obscurely darkly annulate, intermediate and posterior
femora darker at apices; head with the postocular about twice the length of the ante-
ocular area with a slender oblique spine on each side of apex; antennze testaceous,
sparsely and shortly pilose, first joint about as long as anterior femora, second a little
shorter than first; rostrum with the first joint about as long as the other two joints
170 PERCY SLADEN TRUST EXPEDITION
together and distinctly passing the eyes; pronotum about as long as the head, anterior
lobe shorter than the posterior lobe which has two obtuse tubercles on disk a little
in front of the hind angles which are somewhat longly straightly spinous; membrane
slightly passing the abdominal apex ; legs shortly finely pilose.
ono enrnes Oealls ermnns
Loc. Seychelles. Silhouette: from near Mont Pot-d-eau, and from Mare aux
Cochons, over 1000 feet, VIII. 1908. Mahé: near Morne Blane, about 1000 feet; Cascade
Estate &c.
Subfamily Nabidine.
91*. Acanthobrachys elegantula (Stal).
Arbela elegantula Stal, Hem. Afr., iii. p. 42 (1865).
Loc. Seychelles. Silhouette : near Mont Pot-d-eau, ca. 1500 feet; Mare aux
Cochons and forest above, over 1000 feet; cultivated country near coast. Mahé: near
Morne Blane, ca. 1000 feet; Cascade Estate, ca. 1000 feet, and forest above; also low
country. Swept from grasses; characteristic of more open places in lower part of
mountain-forests, about 1000 feet (H. Scott). Originally described from Bourbon.
92*, Nabis capsiformis, Germ.
Nabis capsiformes Germ., in Silberm. Rev., v. p. 132 (1837).
Loc. Seychelles. Mahé: from grass in cultivated country, about 1000 feet;
Caseade Estate, about 1000 feet. Almost universally distributed.
Family Saldide.
93. Salda msignis, sp. n. (Plate 12, fig. 9).
Head, pronotum, scutellum and corium black ; lateral margin, outer cellular space,
and four small spots—two outside each claval area and one on each side of inner angle—
to corium, and membrane—excluding black venation—dull greyish-white ; body beneath
black ; legs ochraceous ; antennze black, basal joint ochraceous, fourth fuscous-brown,
third distinctly thickened, about as long as fourth; pronotum subtruncately emarginate
before scutellum, the lateral margins oblique, very slightly sinuate, the anterior margin
less than half the breadth between posterior angles; corium moderately convexly rounded
at lateral margins; posterior tibize distinctly spinulose, anterior and intermediate tibise
apparently non-spinulose, in unique carded type.
Long. 24 mm.
Loc. Seychelles. Mahé: from cultivated country near sea-level at Cascade, 20. IL.
1909, 1 specimen.
Family Ceratocombide.
Ceratocombide Fieb., Eur. Hem., p. 25 (1861).
Dipsocoride Kirk., Canad. Entomol., 1906, p. 38.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 171
In the previous enumeration only one species had been described from the Seychelles.
This is contained in the present collection with five other species, making six in all. CeicAa
All these species appear to be endemic to the Seychelles. 3yh0°* "ONT,
1d wow “a\Z)\
94*, Ceratocombus imsularis, Reut. ) LIBRARY) =o!
Ceratocombus insularis Reut., Rev. d’Ent., xii. p. 208 (1893). a =e |
ae Mass Qy/
Loc. Seychelles. Mahé: from near Morne Blanc, about 1000 feet, some specimens ‘a @ ee
being recorded as from leaves of Pandanus Horne: ; Cascade Estate, about 1000 feet and
above, some specimens being recorded as found among damp dead leaves (mostly of palms)
on the ground in a valley containing many Stevensonia palms; Mare aux Cochons district,
1000—2000 feet; Morne Seychellois, ca. 1500 feet. Silhouette: near Mont Pot-a-eau,
ea. 1500 feet (2 specimens found between leaf-bases of growing Roscheria palm, several
found under the bark of a stump); Mare aux Cochons, over 1000 feet; from long grass
in cultivated country, VIII. 1908, 1 specimen. Previously recorded from La Digue.
95. Gen. ¢ alboclavatus, sp. n. (Plate 12, fig. 17).
Head, pronotum and scutellum black; hemelytra pitchy-brown, the venation of
the corium prominent and black, the membranal venation also darker; clavus creamy-
white ; legs ochraceous ; antennze mutilated in typical specimens; head not quite as long
as breadth between eyes, obtusely narrowed in front, the lateral margins oblique; eyes
pearly-white ; pronotum a little less than half as long as breadth at base, the anterior
margin half the width of posterior margin which is slightly concave; scutellum small,
triangular, its length about half the width at base; corium and membrane with the
venation as clearly shown in the enlarged figure ; femora moderately thickened; posterior
tibie about as long as corium.
Long. 14 mm.
Loc. Seychelles. Silhouette : Mare aux Cochons, over 1000 feet ; near Mont
Pot-a-eau, ca. 1500 feet. Mahé: country above Port Glaud, 500—1000 feet.
The absence of the antennze prevents the exact determination or description of the
genus in which this species should be placed. Its distinct markings make its recog-
nition easy, and its generic position must remain in abeyance until perfect specimens are
procured.
SEYCHELLESANUS*, gen. nov.
Body moderately short and broad; head large, subtriangular, in length more than
half the width between eyes which reach the anterior angles of the pronotum ; rostrum
reaching the posterior cox, first joint reaching the anterior margin and second the
posterior margin of the anterior coxee which are large and globose; antenne about as
long as the body, longly setose, with the first and second joints incrassate, first joint short,
not reaching apex of head, second longer than first, remaining slender; pronotum
transverse, in length half the width of base, slightly convex, the lateral margins
* In honour of Moreau de Séchelles after whom these islands were named.
172 PERCY SLADEN TRUST EXPEDITION
moderately rounded and longitudinally submarginally impressed, anterior margin about
three-fourths the breadth of posterior margin, and moderately concave ; scutellum about
as long as broad at base, subtriangular, transversely impressed before middle ; hemelytra
convex, lateral margins rounded, apices roundly truncate, venation as shown in Plate 12,
fig. 10; femora moderately thickened, tibize setose, posterior tibize about as long as head,
pronotum and scutellum together.
96. Seychellesanus typicus, sp. n. (Plate 12, fig. 10).
Head, pronotum and scutellum rosy-red ; hemelytra dull ochraceous, the apical cells
paler in hue; antennze with the first and second joints pale ochraceous, remainder a little
darker in hue; legs ochraceous ; lateral margins of hemelytra rosy-red; body above
pilose ; hemelytra posteriorly depressed and finely but distinctly punctate ; other
structural characters as in generic diagnosis.
Long. 1 mm.
Loc. Seychelles. Mahé: 6 specimens, all found among damp dead leaves (mostly
of palms) on ground, in valley contaming many Stevensonia palms, above Cascade,
ca. 1000 feet (cf. Ceratocombus insularis).
In these specific descriptions I have not attempted to describe in words the venation
of the hemelytra. This feature is accurately and better demonstrated by the enlarged
figures taken under the microscope by the artist, Mr H. Knight.
97. Seychellesanus picturatus, sp. n. (Plate 12, fig. 11).
Body above castaneous-red, hemelytra a little paler and transversely crossed at base
by a broad white fascia which contains anteriorly a dull red spot on each side and also is
laterally margined with the same colour; antenne with the first and second joints
ochraceous, remainder a little darker in hue; legs ochraceous; head longly and hemelytra
more shortly pilose; hemelytra less distinctly veined than in the preceding species
(S. typicus), and more finely punctate.
Long. 1 mm.
Loc. Seychelles. Mahé: Cascade Estate, about 800 feet, 1 specimen.
98. Seychellesanus variegatus, sp. n. (Plate 12, figs. 12, 13, 14).
Typical form (fig. 12).
Head testaceous; pronotum, scutellum and hemelytra, black; hemelytra with a
broad central whitish fascia separated by the tegminal suture and with an attached white
spot on each side, three apical greyish-white spots on each side of apex; legs and antennze
pale ochraceous.
Var. a (fig. 14). Differing from typical form in having the central white fascia to
the hemelytra without the attached white spots on each side, the apical spots darker and
with some longitudinal slightly paler discal fasciee.
Var. b (fig. 18). Hemelytra with a pale narrow submarginal fascia on each side
of scutellum, some pale lines on each side of the central white fascia, eight linear white
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 173
spots in curved transverse series before the apical rounded spots. Hemelytra distinctly
and somewhat coarsely punctate ; head longly pilose ; antenn longly setose.
Long. 1 mm.
Loc. Seychelles. Mahé: Mare aux Cochons district, 1000—2000 feet, I. 1909,
1 specimen (the type); damp forest between Trois Fréres and Morne Seychellois,
1500—2000 feet, XII. 1908, 1 specimen (var. a); Cascade Estate, ca. 800 feet or over,
2 specimens (var. 0).
OGERIA*, gen. nov.
Body oblong ; bead short, transverse, strongly deflected, twice as broad between eyes
as long, eyes moderately inclined backward and touching the anterior pronotal angles ;
antennee with the first and second joints incrassate, first joint shorter than head though
passing its apex and inserted just beneath the inner margins of eyes, second a little longer
than first, remainder slender, the whole strongly setose ; rostrum unseen in unique carded
type; pronotum about half as long as broad at base, anterior margin a little more than
half as broad as posterior margin which is moderately convex; scutellum small, about as
long as broad; hemelytra elongate, venation as figured in type O. insularis (Plate 12,
fig. 16); femora moderately incrassated, posterior tibiee scarcely longer than head,
pronotum and scutellum together.
Type O. insularis Dist.
99. Ogeria insularis, sp. n. (Plate 12, fig. 16).
Head, pronotum and scutellum black; hemelytra with about basal half white with
an anterior rounded black spot on each side of scutellum, about apical half piceous with
the venation black; antennez with the first and second joints ochraceous, remainder a
little darker; legs ochraceous, bases of tibizee narrowly infuscated, head and pronotum
thickly finely punctate ; other structural characters as in generic diagnosis.
Long. 14 mm.
Loc. Seychelles. Mahé: one specimen, found (together with Seychellesanus typicus
and some Ceratocombus insularis) among damp dead leaves, mostly of palms, on the
ground in a valley containing many Stevensonia palms, above Cascade, ca. 1000 feet.
Family Cimicide.
100. Cimex rotundatus, Sign.
Cimex rotundatus Sign., Ann. Soc. Ent. France, 1852, p. EY Ca 2) a se
Loc. Aldabra (Fryer). Previously recorded from Mauritius ; Réunion; W. Africa ;
W. Indies; Socotra Isld.; Ceylon; Brit. India; Formosa; China.
* Named after Charles Oger, one of the officers in the Roslan expedition of 1769.
SECOND SERIES—ZOOLOGY, VOL. XVI. 23
174 PERCY SLADEN TRUST EXPEDITION
Family Capside.
In my previous enumeration of the Capsidee in this fauna only three species could be
recorded ; the number is here increased to twenty-four. Two species however recorded
by Reuter (Trigonotylus ruficornis var. tenuis and Cyrthorrhinus parviceps) from the
Seychelles are not found in this collection.
The Capside from the Seychelles have very wide and diverse generic aftinities.
Aretas, Felusacus and Hermotinus were founded for the reception of Oriental species,
while Vannius, Cylapus and Fulvius were originally regarded as neotropical genera. One
species, Callicratides rama Kirby, an Oriental species, is found in the Seychelles. Some of
the other genera are of world-wide distribution ; others are described as new.
Genus CALLICRATIDES.
Callicratides Dist., Faun. Brit. Ind., Rhynch., ii. p. 417 (1904).
Hyalopeplus Reut. (nec Stal), Acta Soc. Sci. Fenn., 1910, p. 158.
101. Calhicratides rama (Kirby).
Capsus rama Kirby, Journ. Linn. Soc. London, Zool., xxiv. p. 106 (1891).
Loc. Seychelles. Mahé: Cascade Estate, about 1000 feet, II. 1909, one specimen.
Originally described from Ceylon where it is not uncommon. Only a single specimen
contained in this collection, the species probably introduced.
102. Megacelum flagellatum, sp. n. (Plate 18, fig. 14).
Above fuscous-brown, more or less irrorated with dark ochraceous; anterior and
posterior margins of pronotum and lateral margins of corium, narrowly pale testaceous ;
pronotum with two anterior black spots between which the colour is paler and ochraceous ;
cuneus castaneous, the anterior and lateral margins pale testaceous ; membrane fuliginous
with opaline lustre; body beneath (imperfectly seen in carded type) and legs dull
ochraceous, lateral areas of the sternum black; bases and apices of tibize black, posterior
femora with the apical halves castaneous annulated with ochraceous ; antennee with the first
joint a little thickened, ochraceous, apically black, nearly as long as head and pronotum
together, second joint ochraceous with three black annulations, more than twice as long as
first, third joint castaneous, basally ochraceous followed by black, a little shorter than
second, fourth piceous with its base pale ochraceous ; head finely, centrally, longitudinally
incised ; pronotum distinctly punctate, the two anterior black spots callose; apex of
seutellum darkly subnodulose ; tibize finely spinulose.
Long. 8 mm.
Loc. Seychelles. Silhouette: marshy plateau of Mare aux Cochons, ca. 1000 feet.
Mahé: Cascade Estate, from forest 1000—2000 feet.
Allied to M. antennatum Kirby from Ceylon and Nilgiris, but with the first and
second joints of antennz longer, and with different colour and markings, pronotum also
distinct.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 175
103. Megacelum (Creontiades) mimicum, sp. n. (Plate 18, fig. 15).
Head ochraceous; pronotum pale testaceous with darker speckles, and with a black
subbasal marginal line; scutellum pale testaceous, the basal area speckled with black and
with the apex black and subnodulose; corium ochraceous; cuneus ochraceous, a black
spot at inner angle and the posterior margin more or less sanguineous; membrane
ochraceous, the cellular margins sanguineous; body beneath (imperfectly seen in carded
type) pale testaceous; legs ochraceous, posterior femora (excluding basal areas) castaneous;
antennze ochraceous, first joint a little thickened and about as long as pronotum and
sparingly, shortly hirsute, second joint about twice as long as first, slightly curved, third
a little shorter than second, fourth less than half the length. of the third ; head moderately
and obliquely deflexed before eyes, and with a distinct narrow central longitudinal
sulcation ; pronotum with the anterior margin about half the width of the posterior ;
seutellum with the disk somewhat longitudinally raised; tibize distinctly spinulose ;
corium distinctly pilose.
Long. 74 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet; near Mont Pot-
d-eau, 1500 feet; also cultivated country near coast. Mahé: near Morne Blanc, about
1000 feet; Morne Seychellois, ca. 1500—2000 feet; Long Island. Praslin; Cotes d’Or
Estate. Bird and Dennis Islands (Fryer).
Allied to M. rubrinerve Stal, from Central America, but structurally differing from
that species by the shorter first and second joints of the antenne, different colour
markings &c. For M. rubrinerve Stal, I had (1883) founded a genus (Creontiades), but
subsequently (1904) considered it as only a sectional division of Megacalum. Poppius
however is now using Creontiades as a distinct genus.
104*. Collaria improvisa, Reut.
Collaria improvisa Reut., Rev. d’Ent., xi. p. 208 (1893).
Loc. Seychelles. Silhouette; near Mont Pot-a-eau, 1500 feet ; Mare aux Cochons,
over 1000 feet. Mahé: from near Morne Blanc, under and over 1000 feet; Cascade
Estate and forest above ; also places in the low country.
Swept from grass &c. ; characteristic of more open places and clearings in lower part
of mountain-forests (H. Scott). Also (teste Poppius) found in Madagascar, Kilimandjaro
and 8. Africa.
The genus Collaria was till recently regarded as a neotropical and nearctic one, and
its presence in the Seychelles is probably due to accidental introduction. Poppius
however has recently described a species collected during Sjéstedt’s “ Kilimandjaro-Meru
Exp.”
105. Aretas sanguinarius, sp. n. (Plate 18, fig. 12).
Ochraceous ; eyes black ; apex of head, lateral margins of pronotum, about basal half
of corium posteriorly oblique, apex of clavus, posterior margin of cuneus, and cellular
margins to membrane, sanguineous; body beneath and legs ochraceous; antennze with
23—2
176 PERCY SLADEN TRUST EXPEDITION
the first joint sanguineous, incrassate, spinulose, about as long as head, second joint
ochraceous, about three times as long as first, third ochraceous and about as long as first,
fourth mutilated in type; head between eyes as broad as long, finely, centrally, longi-
tudinally suleate ; pronotum broader than long, the lateral margins oblique, transversely
impressed at about one-fourth before anterior margin ; scutellum broad, triangular (in the
figure as in the type, the scutellum is somewhat detached from the pronotum) ; posterior
femora somewhat strongly incrassate, tibize finely spinulose, posterior tarsi with the first
and second joints subequal in length.
Long. 4 mm.
Loc. Seychelles. Mahé: Long Island, VII. 1908, one specimen.
The genus Aretas Dist. was founded for the reception of a Ceylonese species.
106. Vannius mahensis, sp. n. (Plate 18, fig. 8).
Greyish-white ; basal joint of antennze, eyes, lateral margins of pronotum and two
spots at base of same, lateral margins of scutellum, an oblique fascia at outer margins of
clavus, two spots beyond apex of clavus, a cuneal spot, and intermediate and posterior
femora sanguineous; intermediate and posterior femora annulated with greyish-white ;
intermediate tibize moderately tinged with sanguineous; head slightly and convexly
produced in front of eyes and with a slender central longitudinal sulcation ; antennze with
the basal joint moderately incrassate, sanguineous, its extreme base pale ochraceous, about
as long as head, second palely sanguineous, nearly four times as long as first, third and
fourth more ochraceous in hue; pronotum with an anterior transverse constriction which
defines an anterior collar; scutellum with a central longitudinal carination ; cuneus longer
than broad.
Long. 3 mm.
Loc. Seychelles. Mahé: Cascade Estate, III. 1909, one specimen.
Poppius has recently (1910) described a species of this genus from Madagascar
(V. annulicornis) but apart from other characters this may be distinguished from the one
I have here described, by a character he gives in a key, viz. “ Die Seiten des Habsschildes
gelb.” I originally founded the genus for the reception of a Central American species.
107. Cylapus mugratorius, sp. n. (Plate 18, fig. 6).
Head ochraceous, eyes greyish, inwardly margined with black ; pronotum, scutellum
and corium dark castaneous, more or less spotted with dull greyish, the spots more
numerous on the corium and the spot on the scutellum central and longitudinal ; cuneus
ochraceous with a black apical spot; membrane piceous with dark greyish mottlings;
body beneath and legs castaneous, basal areas of femora ochraceous, rostrum castaneous,
its apex black; antennz slender, castaneous, first joint moderately thickened, its base
ochraceous, a little longer than head, second joint nearly as long as head, pronotum and
scutellum together, its apex slightly thickened, third about one and a half times as long
as second and a little longer than fourth, head short, transverse, in length considerably
shorter than breadth between eyes where it is moderately centrally longitudinally
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 177
depressed ; pronotum about half as long as broad at base; membrane passing abdominal
apex.
Long. 4 mm.
Loe. Seychelles. Silhouette: near Mont Pot-a-eau, ca. 1500 feet, two specimens,
one of them from between leaf-bases of a Stevensonia palm; forest above Mare aux
Cochons, over 1000 feet. Mahé: near Morne Blanc, 500—1000 feet ; high forest, Morne
Blane district ; Morne Seychellois, over 1500 feet.
Cylapus is another nearctic and neotropical genus represented in the Seychelles.
108. Felisacus auritulus, sp. n. (Plate 18, fig. 7).
Head, pronotum and scutellum bright golden-yellow, eyes and antennz black, base of
first joint yellow ; corium subhyaline, the costal margin and cuneus yellow; membrane
subhyaline with a violaceous lustre, the cellular veins distinct and darker ; body beneath
and legs golden-yellow ; head long and broad, constricted behind eyes; antennze with the
first joint a little shorter than the pronotum, second about one and a half times as long as
the first (remainder mutilated in unique type); rostrum reaching the posterior coxe ;
pronotum posteriorly convexly tumid, strongly constricted before middle, and with a very
narrow anterior collar, posterior angles subnodulose, posterior lobe very finely and
obsoletely punctate and with a transverse series of coarser punctures at the anterior
constriction.
Long. 4 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, VIII. 1908, one
specimen.
The genus Felisacus has been previously regarded as an Oriental one, and this species
is allied to F. glabratus Motsch. from Ceylon.
Genus EuRYSTYLUS.
Eurystylus Stal, Ofv. Vetensk.-Akad. Forh., 1870, p. 671; Reut., Act. Soc. Sci.
inarmim., GO, jo, WD.
Paracalocoris Dist. (part.), Biol. Centr. Am., Rhynch., i. p. 263 (1883).
Eurycyrtus Reut., Ofv. Finsk. Vetensk. Soc. Férh., xxi. p. 34 (1879).
Olynvprocapsus Kirk., Trans. Ent. Soc. London, 1902, p. 255.
Reuter having stated that his genus Hurycyrtus was available for and should include
the Oriental and Ethiopian species which I had included in my genus Paracalocoris and
that the latter should be restricted to the Neotropical species for which I founded it, I
followed his ruling (Faun. Brit. Ind., Rhynch., v. p. 251). But he has since proposed
another alteration and is of opinion that his Hurycyrtus is only a synonym of Hurystylus
Stal. In this opinion he is in all probability correct as Poppius has seen Stal’s type. He
has however somewhat added to the complexity of this involved synonymy by writing in
his “ Verzeichnis der Synonymen ’”—Paracalocoris Dist. = Hurystylus Stal, whereas, as he
has elsewhere stated, it is only a section of Paracalocoris which = Hurycyrtus Reut. =
Eurystylus Stal.
178 PERCY SLADEN TRUST EXPEDITION
109. Hurystylus alboplagiatus, sp. n. (Plate 12, fig. 18).
Above dull black with scattered patches of white pile; head between eyes, narrow
lateral and posterior margins to pronotum, extreme lateral margins to corium, and
scutellum, ochraceous or testaceous; cuneus castaneous with the apex black; pronotum
with two black spots behind middle; antennze with the first joint piceous, shortly pilose,
about as long’as the pronotum, second joint pale castaneous-brown, extreme base greyish,
apical third piceous where it is distinctly thickened, about twice as long as first, third and
fourth piceous, nearly basal half of third greyish; head nearly as long as breadth between
eyes; pronotum with an anterior submarginal transverse incision ; scutellum moderately
tumid, the basal area testaceous; membrane pale piceous, the venation darkly distinct ;
legs castaneous-brown, tibize palely annulated at base and near middle, tarsi ochraceous,
apex piceous; body beneath more or less testaceous or ochraceous, abdomen with a
sublateral series of dark spots on each side, a distinct black spot on mesosternum but
indistinctly seen in carded unique type.
Long. 5 mm.
Loc. Seychelles. Mahé: Cascade Estate, about 1000 feet, II. 1909, one specimen.
110. Lygus cinnamomeus, sp. n. (Plate 13, fig. 16).
Head, pronotum and scutellum dull ochraceous; basal area of pronotum and scutellum
(the latter with three dull ochraceous spots, two basal and one apical) and corium blackish ;
apex of clavus, lateral margin and a large marginal spot near middle, dull ochraceous ;
euneus ochraceous, the apex blackish; body beneath and legs ochraceous, apical areas of
posterior femora blackly annulate; antennze ochraceous, second jot more than twice as
long as first with the apex black, remaining joints more or less fuscous; eyes black
marked with ochraceous; pronotum finely transversely striate and slightly granulose ;
membrane dark fuliginous, the apex paler; posterior femora incrassate, tibiz finely
setose.
Long. 4 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons and forest above, over 1000 feet.
Mahé: near Morne Blane, ca. 800 feet (many specimens, numbered 68, were beaten from
cinnamon-trees, Cinnamomum zeylanicum, XI. 1908); grass in cultivated country, about
1000 feet; Cascade Estate,.about 1000 feet; Mare aux Cochons district, 1000—2000
feet ; Morne Seychellois, 1500—2000 feet.
111. Lygus sanguineosignatus, sp. n. (Plate 13, fig. 17).
Fuscous-brown; head between eyes, disk of pronotum and an oblong spot near
middle of lateral margin to corium ochraceous, a lateral marginal spot before base of
cuneus, sanguineous ; cuneus greyish-white, its apex fuscous; membrane fuliginous with a
greyish-white spot on the lateral margins; body beneath and legs ochraceous ; antenne
dull ochraceous, second joint more than twice as long as first, third joint fuscous; eyes
black, transverse; pronotum finely transversely wrinkled and subgranulose; corium
shortly, finely pilose.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 179
_ Long. 4 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, VIII. 1908, one
specimen.
112. Lygus silhouettensis, sp. n. (Plate 13, fig. 18).
Ochraceous ; basal and sublateral areas of pronotum, scutellum and clavus, brownish-
ochraceous; scutellum with three obscure paler spots, two on basal margin and one at
apex; cuneus and membrane greyish-white, the latter with opaline lustre ; body beneath
more or less testaceous, legs ochraceous, apices of posterior femora and bases of posterior
tibiz annulated with fuscous; antennze ochraceous, second joint more than twice as long
as first, its apex fuscous, third fuscous, pale at base; eyes black, inwardly testaceous ;
pronotum transversely striate and subgranulose ; scutellum and corium distinctly, some-
what longly, palely pilose. |
Long. 4 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908, one
specimen.
113. Proboscidocoris pluto, sp. n. (Plate 18, fig. 4).
Body above black; sides of head level with insertion of antennae to apex, two spots
between and near eyes, narrow lateral margins to pronotum, apex of scutellum, lateral and
apical margins of corium and margins of cuneus, ochraceous; body beneath dull dark
castaneous; femora castaneous, base of intermediate and posterior femora ochraceous,
tibiee and tarsi pale ochraceous, tibial bases and apex of anterior tibize black; antennz
with the first joint castaneous, second ochraceous with its apex black, third and fourth
fuscous, base of third pale ochraceous, first joint about as long as head, second more than
twice as long as first, third and fourth together as long as second; pronotum thickly,
finely punctate; corium finely, obscurely pilose; scutellum moderately tumid, finely,
transversely striate ; membrane fuliginous, passing abdominal apex, cellular areas darker
with their margins pale; rostrum ochraceous, its apex black.
Long. 43 to 5 mm.
Loc. Seychelles. Silhouette: low coconut-planted country near the coast, Pointe
Etienne. Mahé: marshes on coastal plain at Anse aux Pins and Anse Royale; marshy
ground near sea-level at Cascade.
I have placed this species in Reuter’s genus Proboscidocoris which is very near
Peciloscytus excepting for the more porrect head. I have seen an undescribed species
from Uganda, in bad condition, labelled Proboscidocoris intermedius Popp., which appears
to be nearest to the species above described.
114. Hermotinus villosulus, sp. n. (Plate 18, fig. 5).
Head and pronotum longly pilose; head and anterior area of pronotum greyish,
remaining pronotal area pale brownish with darker spots, two transverse blackish spots
anteriorly, and four darker brown spots posteriorly ; scutellum pale brownish, with two
rounded black spots near base and the apex white; corium mottled brown, the clavus and
180 PERCY SLADEN TRUST EXPEDITION
apical area more piceous, the lateral margins largely spotted with white; cuneus pale
ochraceous, with two small spots and the apex brownish ; membrane piceous mottled with
greyish; body beneath and legs pale ochraceous, the first with some minute brownish
spots, femora and tibize annulated with brownish ; antennze piceous, first joint and central
annulation and apex of second joint ochraceous, first joint longer than head, second nearly
three times the length of first, third and fourth together nearly as long as second ;
pronotum very longly pilose on the lateral margins; rostrum reaching the intermediate
coxee ; membrane passing abdominal apex.
Long. 44 mm.
Loc. Seychelles. Mahé: Cascade Estate, ca. 1000 feet, one specimen.
The genus Hermotinus Dist. was founded for the reception of a species from
Teusasserim, since received also from Ceylon and India. The above is the second species
I have seen.
115. Cyrtopeltus atricornis, sp. n. (Plate 18, fig. 13).
Head, pronotum and scutellum ochraceous; eyes black ; corium dull greyish, cuneus
greyish with a fuscous spot at base and apex; membrane greyish, subhyaline with opaline
lustre ; antenne black, first joint passing apex of head, second more than twice as long as
first, third about as long as second; head considerably longer than broad between eyes,
the apex produced; pronotum with the anterior margin about half the breadth of
posterior margin, transversely constricted near middle, behind which it is finely but
distinctly punctate; scutellum with a central longitudinal ridge; cuneus elongate, much
longer than broad, its apex acute; body beneath brownish-ochraceous, the sternum more
or less ochraceous ; legs stramineous, finely setose.
Long. 44 mm.
Loc. Seychelles. Mahé: Cascade Estate, ca. 1000 feet.
One varietal specimen has the antennz fuscous, with the basal joint almost
ochraceous.
Cyrtopeltus is both a Palearctic and Ethiopian genus.
116. Dereocoris seychellensis, sp. n. (Plate 12, fig. 19).
Pronotum and corium shining black; head testaceous, eyes black; margins of
pronotum narrowly ochraceous; scutellum ochraceous; membrane pale fuliginous,
excepting at apex reflecting the dark abdomen beneath; sternum black; abdomen
beneath piceous ; rostrum, coxee, and legs pale ochraceous, apices of the tibiz black;
antennee ochraceous, apex of first jot and about apical third of second joint black, first
joint nearly as long as head, second more than twice as long as first, about apical third
distinctly thickened and pilose, the last two joints pilose and together shorter than the
second ; eyes touching the anterior margin of pronotum ; pronotum and corium distinctly
punctate ; scutellum slightly tumid; membrane distinctly passing the abdominal apex ;
tibiee finely setose.
Long. 4 mm.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 181
Loc. Seychelles. Mahé: Anonyme Island; marshes on coastal plain at Anse aux
Pins and Anse Royale; Long Island.
This species is allied to the 8S. and C. African D. ostentans Stal. Poppius in
referring to this species (Sjéstedt’s Kilimand.-Meru Exped., Pt. 12, p. 50) transferred
it to the genus Camptobrochis, but has since acknowledged the correctness of Stal’s
generic location, by returning a specimen to the Entomol. Research Com. of Trop. Africa
labelled Deraocoris ostentans.
117. Dereocoris? cardut, sp. n. (Plate 18, fig. 1).
Purplish-red ; eyes black ; legs purplish-red, bases of femora and central annulations
to intermediate and posterior tibie, ochraceous ; antenne with the first and second joints
purplish-red, base of first pale ochraceous, more than apical third of second black, fourth
and fifth black, their bases pale ochraceous, first joint longer than head, second a little
more than twice as long as first, its apical third moderately thickened and finely pilose,
third and fourth together considerably shorter than second; eyes touching anterior
margin of pronotum; pronotum and corium finely punctate, scutellum about as long as
breadth at base, a little tumid; membrane passing abdominal apex; pale fuliginous, the
venation darkly distinct ; tibize finely setose.
Long. 45 mm.
Loc. Seychelles. Mahé: beaten from a tree in highest forest on summit of Morne
Pilot, over 2000 feet, 22. XI. 1908, one specimen.
I have provisionally placed this species in Derwocoris but with some hesitation ; the
characters found in the antennz and scutellum making it scarcely congeneric.
118. Fulvius dolobratus, sp. n. (Plate 13, fig. 10).
Above brownish-ochraceous; eyes black; lateral margins of the corium pale
ochraceous; cuneus piceous, its anterior margin ochraceous; membrane fuliginous ;
body beneath dark testaceous; legs ochraceous; antennze pale piceous, about apical
third of the second joint pale ochraceous, first joint passing apex of head and moderately
thickened, second about twice as long as first; eyes not touching the anterior pronotal
angles; pronotum and scutellum distinctly thickly finely punctate; coritum obscurely
finely punctate; posterior femora thickened, shorter than posterior tibie; rostrum
ochraceous, apically fuscous, first joint about or nearly reaching base of head.
Long. 3} mm.
Loc. Seychelles. Praslin: Cétes d’Or Estate, XI. 1908, one specimen.
Fulvius Stal, was founded as a neotropical genus, and is also found in the Palearctic
region.
119. Fulvius pictus, sp. n. (Plate 18, fig. 11).
Head, pronotum and scutellum black ; apex of head, three longitudinal spots at base,
and two spots (sometimes almost obsolete) at base of pronotum, one near each posterior
angle ; corium piceous, basal area and base of costal margin, brownish-ochraceous, cuneus
piceous, its base and apex pale ochraceous, apex of clavus ochraceous; membrane fuli-
SECOND SERIES—ZOOLOGY, VOL. XVI. 24
182 PERCY SLADEN TRUST EXPEDITION
ginous ; femora black, extreme femoral apices, tibize and tarsi pale ochraceous ; rostrum
ochraceous ; antennze with the first joint black, its apex ochraceous and passing apex of
head, moderately thickened, second jot ochraceous with its apex black, about twice as
long as first, third and fourth together about as long as second ; head longer than breadth
between eyes, its apex subacute; corium finely, palely, pilose; membrane passing the
abdominal apex ; body beneath black, the coxze ochraceous.
Long. 3 to 34 mm.
Loc. Seychelles. Silhouette: near Mont Pot-a-eau, about 1500 feet; Mare aux
Cochons, over 1000 feet ; low coconut-planted country near the coast, Pointe Etienne.
Mahé: from between leaf-bases of a growing Stevensonia palm near Morne Blane,
ca. 800 feet, one specimen; also one specimen from low country.
120. Fulvius? niger, sp. n. (Plate 18, fig. 9).
Black ; head in front of eyes dull ochraceous; antennze with the first joint black,
remaining joints ochraceous and finely pilose, first joint scarcely passing apex of head,
second about or nearly twice as long as first, third and fourth together about as long as
second; head longer than breadth including eyes, which are, rather long and adpressed ;
pronotum transversely constricted near middle, the anterior angles rounded, lateral
margins roundly oblique, very slightly sinuate, the surface distinctly finely punctate ;
scutellum and corium distinctly finely punctate, membrane fuliginous, the basal area
reflecting the dark abdomen beneath; legs black, tarsi pale ochraceous, posterior tibiz
scarcely longer than the posterior femora.
Long. 3 mm.
Loc. Seychelles. Silhouette: from Mare aux Cochons, over 1000 feet. Mahé:
precise locality uncertain.
Apart from the length of the posterior tibiee, the general characters of this species
appertain to the genus Mulviws in which I have provisionally placed it, the carded
condition of the type making more precise generic identification very difficult. It is also
allied to Fulvidius Poppius.
MaARALAUDA™, gen. nov.
Subelongate, moderately broadened posteriorly ; head longer than breadth between
eyes, anteriorly narrowed, the central lobe prominent; antennz with the first joint
shorter than head, second about twice as long as first and a little thickened on apical
third, third and fourth together about as long as second ; rostrum passing the posterior
coxee, first joint slightly passing base of head ; pronotum convex on basal area, deflected
and narrowed anteriorly, lateral margins obliquely rounded, about twice broader at base
than at anterior margin ; scutellum subtriangular, as long as broad at base, transversely
striate; corium (including cuneus) a little longer than head, pronotum and scutellum
together, laterally deflected, cuneus broader than long, cuneal fracture profound ;
* Named after the “Sealark” which carried the ‘“‘ Percy Sladen Trust Expedition” to the Indian Ocean in
1905,
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 183
membrane passing abdominal apex ; femora moderately thickened, posterior coxze some-
what contiguous, tibize finely setose.
121. Maralauda lania, sp. n. (Plate 18, fig. 3).
Body above dull dark sanguineous, pronotum and more distinctly the corium speckled
with white pile, all the margins a little paler sanguineous; membrane fuliginous ; body
beneath and rostrum dark sanguineous, femora pale ochraceous, their apices sanguineous,
tibize reddish-ochraceous, darkly setose, tarsi pale ochraceous, their apices black ; antennze
with the first and second joints testaceous, bases and apex of the second black, third and
fourth black, basal half of third pale ochraceous; pronotum and more especially the
corium shortly finely pilose ; scutellum transversely striate ; other structural characters as
in generic diagnosis.
Long. 4 mm.
Loc. Seychelles. Mahé: Cascade Estate, forest, 1000—2000 feet; Mare aux
Cochons district 1000—2000 feet; Morne Seychellois, over 1500 feet. Only from the
high endemic mountain-forests.
122. Sthenarus leucophilus, Reut.
Sthenarus leucophilus Reut., Ofv. Finsk. Vetensk. Soc. Forh., xlvii. no. 22, p. 88
(1904—1905).
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet ; marshes on coastal plain
at Anse aux Pins and Anse Royale: characteristic of marshy places, where it was swept
from low vegetation. Bird Island (Fryer).
This species was originally described by Reuter from Bourbon, and has since been
recorded by Poppius from Kilimandjaro.
STHENA RUSOIDES, gen, Nov.
Subelongate ; head transverse, shorter than broad, eyes somewhat prominent but not
touching the anterior angles of the pronotum ; antennze with the first jomt very slightly
longer than head, second a little more than twice as long as first and moderately thickened
on its apical area, third and fourth finely setose and together about as long as second ;
rostrum short, robust, not extending beyond the anterior coxe; pronotum about half as
long as broad at base which is one-third broader than anterior margin, a distinct narrow
anterior collar, lateral margins moderately rounded ; scutellum subtriangular, considerably
shorter than broad at base; corium (excluding cuneus) about as long as head, pronotum
and scutellum together; cuneus much broader than long, cuneal fracture profound ;
membrane passing abdominal apex; posterior coxe contiguous; tibiz shortly finely
Setose.
123. Sthenarusoides montanus, sp. n. (Plate 13, fig. 2).
Black, shining ; head, rostrum and legs pale ochraceous, apices of the femora black ;
antennze ochraceous, extreme apex of first joint, about apical half of second, and the third
and fourth jomts black; head with a few scattered punctures; pronotum and corium
24__9
184 PERCY SLADEN TRUST EXPEDITION
thickly punctate, scutellum more sparingly and finely punctate; membrane pale fuli-
ginous, on basal area reflecting the dark abdomen beneath ; other structural characters as
in generic diagnosis.
Long. 3 mm.
Loc. Seychelles. Silhouette: near Mont Pot-i-eau, about 1500 feet. Mahé: high
forest of Morne Blane and Pilot; Mare aux Cochons district, 1000—2000 feet; Morne
Seychellois, 1500—2000 feet ; Cascade Estate, ca. 1000 feet ; forest near Mount Harrison,
about 1700 feet; also one specimen from Port Victoria.
This species is found in fern and other undergrowth in high damp mountain-forests
(H. Scott).
124, Psallus? mahensis, sp. n. (Plate 18, fig. 19).
Dark olivaceous, lateral marginal areas of corium, and cuneus more or less pale
ochraceous ; body beneath more or less castaneous, legs ochraceous, intermediate and
posterior femora castaneous; antennze with the basal joint castaneous, a little passing
apex of head, second joint ochraceous, more than twice as long as first, third and fourth
Joints fuscous ; corium shortly but distinctly pilose ; membrane pale fuliginous.
Long. 2 mm.
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet ; marshes on coastal plain
at Anse aux Pins and Anse Royale; Cascade, cultivated country near sea-level. This
species is characteristic of marshy places, both on the coast and at an elevation of about
1000 feet: a number of specimens were swept (with specimens of Sthenarus leucophilus)
from a little marsh containing much of a wide-spread Onagraceous plant (Jussi@a sp.) on
Cascade Estate at about 800 feet.
Family Anthocoride.
In the previous enumeration only one species of this family could be recorded, the
Lasiochilus alluaudi Reut. This species is found in the present collection with ten
others, making eleven species in all. The Ceylonese species Seselliws parallelus Motsch.
is found in the Seychelles, while the genus Lasvochilus, in which six species are located, is
a genus well represented in the neotropical region.
125*. Lasiochilus alluaudi, Reut.
Lasiochilus alluaudi Reut., Rev. d’Ent., xii. p. 209 (1898).
Loc. Seychelles. Silhouette: near Mont Pot-ad-eau, ca. 1500 feet; Mare aux
Cochons and forest above, over 1000 feet. Mahé: high forest of Morne Blane and Pilot ;
Cascade Estate, about 1000 feet; also from low country. Previously recorded from La
Digue.
126. Lasvochilus scotti, sp. n. (Plate 13, fig. 20).
Ochraceous ; head about as long as broad ; antennee pilose, first joint scarcely passing
apex of head, second about twice as long as first, longer than third or fourth which are
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 185
about subequal in length; pronotum transversely constricted behind middle, after which
the surface to base is distinctly punctate, before the constriction transversely globose,
base before scutellum obliquely deflected; scutellum punctate; corium finely pilose ;
cuneus broad; membrane fuliginous, the margins paler; legs ochraceous, apices of the
tarsi fuscous.
Long. 2 mm.
Loc. Seychelles. Silhouette: near Mont Pot-a-eau, about 1500 feet. Mahé:
Morne Seychellois, over 1500 feet.
Allied to Z. alluaudi Reut.
127. Lasiochilus gardineri, sp. n. (Plate 18, fig. 21).
Head, pronotum and scutellum black; apex of head and corium pale ochraceous ;
cuneus piceous ; membrane fuliginous, the margins paler; body beneath imperfectly seen
in unique carded type—head and sternum black, abdomen piceous, its base ochraceous ;
legs ochraceous; antennze pilose, ochraceous, first joint scarcely passing apex of head,
second nearly twice as long as first and longer than either third or fourth which are
almost subequal in length; head longer than broad at base; pronotum strongly trans-
versely constricted behind middle, thence to base distinctly punctate, before the constriction
transversely globose, base deflected to scutellum which is sparingly punctate; corium
shortly finely pilose, cuneus more distinctly pilose ; apices of the tarsi fuscous. j
Long. 2 mm.
Loc. Seychelles. Mahé: country above Port Glaud, about 500—1000 feet, 5. XI.
1908, one specimen.
128. Lasiochilus sladent, sp. n. (Plate 18, fig. 22).
Ochraceous ; eyes black ; a fascia between eyes and broad apical margin of cuneus,
piceous ; membrane dull ochraceous; body beneath and legs ochraceous ; antennze pilose,
ochraceous, basal joint scarcely passing apex of head, second joint about twice as long as
first, third almost as long as second, fourth considerably shorter than third; head
considerably longer than broad at base ; pronotum obscurely pilose and punctate, strongly
transversely constricted near middle, behind which the surface is concavely depressed, and
before which it is transversely globose; scutellum punctate; corium finely pilose, the
cuneus more strongly pilose.
Long. 25 mm.
Loc. Seychelles. Silhouette: Mare aux Cochons and forest immediately above,
over 1000 feet; low coconut-planted country near the coast, Pointe Ktienne.
129. Lasiochilus seychellensis, sp. n. (Plate 18, fig. 23).
Head and pronotum black ; scutellum piceous ; apex of head and corium ochraceous ;
cuneus piceous; membrane dull ochraceous; body beneath (imperfectly seen) and legs,
ochraceous ; antennze somewhat longly pilose, ochraceous, apex of second joint piceous,
first joint scarcely passing apex of head, second joint scarcely twice the length of first,
186 PERCY SLADEN TRUST EXPEDITION
and only slightly longer than third, the first and second moderately thickened very little
longer than breadth at base; pronotum pilose, strongly transversely constricted near middle,
behind which it is obscurely, finely punctate, and before which it is distinctly globose ;
scutellum distinctly punctate ; corium pilose, the cuneus more strongly pilose.
Long. 25 mm.
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet.
130. Lasiochilus praslinensis, sp. n. (Plate 18, fig. 24).
Head, pronotum and scutellum testaceous-red ; corium and cuneus piceous, basal
lateral area of corium dull ochraceous ; membrane fuliginous; body beneath testaceous-
red ; legs fuscous ; antennze with the first joint testaceous, remaining joints ochraceous,
first scarcely passing apex of head, second twice as long as first, its extreme apex a little
darkened ; head longer than broad at base; pronotum moderately transversely impressed
near middle, behind which it is distinctly and rather coarsely punctate and before which
it is transversely globose ; scutellum distinctly punctate, moderately concavely impressed ;
corium shortly pilose, the cuneus more strongly pilose.
Long. 25 mm.
Loc. Seychelles. Praslin : Cétes d’Or Estate, from Coco-de-mer forest in the Vallée
de Mai, XI. 1908.
PARALASIOCOLPUS, gen. nov.
Subelongate, moderately depressed; head longer than broad, somewhat longly
produced in front of eyes, the anteocular more than twice the length of the postocular
area ; antennze pilose, with the first joint scarcely passing the apex of head, second about
twice the leneth of first; pronotum with the apical margin less than half the breadth
of posterior margin, transversely constricted near middle, the basal margin concave ;
scutellum transversely ridged near base, subtriangular, the disk more or less broadly
foveate ; corium about as long as from region of eyes to apex of clavus; cuneus large,
broad, triangular, almost equilateral; membrane with three longitudinal veins; rostrum
short, reaching the anterior trochanters.
Type P. piceus Dist.
Allied to the neotropical genus Lasiocolpus Reut., from which it principally differs in
the short rostrum.
131. Paralasiocolpus piceus, sp. n. (Plate 18, fig. 25).
Head, pronotum and scutellum piceous, corium black, an obscure pale spot at base of
lateral margin to cuneus; membrane dark fuliginous; body beneath and legs black or
piceous ; antennze ochraceous, pilose, first joint and apex of second black, first scarcely
passing apex of head, second twice as long as first; pronotum and scutellum coarsely
punctate ; corium very obscurely finely pilose ; tarsi pale ochraceous, other characters as
in generic diagnosis.
Long. 3 mm.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 187
Loc. Seychelles. Silhouette: near Mont Pot-a-eau, ca. 1500 feet; Mare aux
Cochons, over 1000 feet, and high forest above. Mahé: high forest of Morne Blanc and
Pilot; Cascade Estate, forest over 1000 feet; Mare aux Cochons district, 1000—2000
feet.
132. Paralasiocolpus marginatus, sp. n. (Plate 18, fig. 26).
Black; scutellum testaceous-red; outer margin of clavus, basal marginal area
(broadly) and posterior margin (narrowly) of embolium and posterior margin (narrowly) of
cuneus dull, pale stramineous ; antennz pilose with the first joint black, not quite reaching
apex of head, second black with its apex ochraceous, about twice as long as first, first and
second moderately thickened, remaining joints ochraceous ; head considerably longer than
breadth at base; pronotum transversely constricted near middle, behind which it is
thickly, rather coarsely punctate, before which it is transversely globose and rugose ;
scutellum moderately foveate, transversely ridged at base and with a central longitudinal
ridge ; corium distinctly pilose; legs piceous, tarsi ochraceous ; body beneath imperfectly
seen in the unique typical carded specimen.
Long. 2 mm.
Loc. Seychelles. Mahé: Cascade Estate, forest above 1000 feet, I. 1909, one
specimen.
OSTORODIASOIDES, gen. nov.
Elongate, subdepressed ; head anteriorly narrowed and produced in front of eyes,
distinctly longer than broad; antennze setose with the first jomt not reaching apex of
head, second joint more than twice as long as first, first and second moderately thickened,
third and fourth joints each shorter than second ; rostrum passing the anterior coxe, first
joint about reaching base of head; pronotum with an anterior collar, transversely
constricted behind middle, before the constriction a central and lateral foveation, anterior
margin nearly half the length of posterior margin, which is concavely sinuate ; scutellum
subtriangular, apically depressed ; corium with the margins subparallel, the cuneus large,
almost equilateral; anterior femora broadly thickened, shortly spinose beneath ; tarsi
three-jointed.
Type O. typicus Dist.
133. Ostorodiasoides typicus, sp. n. (Plate 18, fig. 29).
Ochraceous ; eyes and a spot behind apex of clavus, black; antennz ochraceous,
setose, first joint not reaching apex of head, second more than twice as long as first, both
moderately thickened; pronotum behind the transverse constriction distinctly punctate,
before constriction with three foveations, one central and one near each lateral margin ;
scutellum abruptly deflected from base to about middle, where it is transversely
constricted ; corium finely pilose, cuneus more strongly pilose and a little darker in hue ;
membrane dull ochraceous; body beneath and legs ochraceous ; anterior femora broadly
thickened, shortly spinose beneath; other structural characters as in generic diagnosis.
Long. 24 to 3 mm.
188 PERCY SLADEN TRUST EXPEDITION
Loc. Seychelles. Silhouette: from Mare aux Cochons and forest above, over 1000
feet. Over 50 specimens were beaten from palm-leaf thatch of hut at Mare aux Cochons,
IX. 1908 (H. Scott).
134. Ostorodiasoides signatus, sp. n. (Plate 18, fig. 28).
Ochraceous ; eyes and space between them, posterior margin of pronotum, base of
scutellum, claval suture, and cuneus piceous or black; head beneath, sternum (excluding
prosternum), and abdomen beneath, piceous or black; legs ochraceous; antenne
ochraceous, setose, first and second joints moderately thickened, second more than twice
as long as first and black at apex; pronotum transversely constricted behind middle,
behind constriction distinctly punctate, before constriction with three foveations, one
central and one near each lateral margin; scutellum abruptly depressed near middle;
corium distinctly pilose ; anterior femora broadly thickened, shortly spinose beneath.
Long. 25 to 3 mm.
Loe. Seychelles. Silhouette: Mare aux Cochons and forest immediately above,
over 1000 feet ; low coconut-planted country near the coast.
Genus SESELLIUS.
Sesellius Dist., Faun. Brit. Ind., Rhynch., ui. p. 6 (1906).
Scoloposcelis Popp. nec Fieb.
135. Sesellius parallelus (Motsch.) (Plate 18, fig. 27).
Anthocoris parallelus Motsch., Bull. Soc. Mose., xxxvi. (3), p. 89 (18638).
Sesellius parallelus Dist., Faun. Brit. Ind., Rhynch., iii. p. 7, fig. 6 (1906).
Scoloposcelis picicornis Popp., Acta Soc. Sci. Fenn., xxxvii. no. 9, p. 26 (1909).
Scoloposcelis parallelus Popp., Wien. ent. Zeit., xxix. p. 140 (1910).
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908, one
specimen. Mahé: Mare aux Cochons* district, 1000—2000 feet, I. 1909, one specimen.
Originally described from Ceylon. I am now able to give a better figure of this species,
the Ceylonese specimen I figured in Faun. Brit. Ind. series being in a somewhat indifferent
condition.
Family Nepide.
136*. Ranatra grandocula, Berger.
Ranatra grandocula Bergr., Rev. d’Ent., xii. p. 207 (1893).
Loc. Seychelles. Mahé: Cascade Estate. Silhouette: from a stream on the Mare
aux Cochons plateau, over 1000 feet.
* The name “ Mare aux Cochons” is given to two distinct localities, one in the island of Mahé, and the
other in Silhouette.
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA 189
Family Notonectide.
Subfamily Notonectine.
137*. Anisops varius, Fieb.
Anisops varius Fieb., Abh. béhm. Ges. Wiss., (5) vil. p. 483 (1851).
Loc. Seychelles. Silhouette: from a stream on the marshy plateau of Mare aux
Cochons, over 1000 feet ; from near Mont Pot-a-eau, about 1500 feet, and from a stream
near sea-level at Anse Mondon. Previously taken in Praslin, Mahé and Coetivy.
Also recorded from Travancore, Philippines, Eastern Palearctic region and North
Africa.
138. Anisops aldabrana, sp. n. (Plate 18, fig. 30).
Vertex between eyes and pronotum, silvery-grey ; eyes black ; scutellum ochraceous
with a black spot at each basal angle; hemelytra blackish-grey, paler on basal area ;
sternum and legs dull stramineous, the latter more or less streaked with black ; abdomen
beneath fuscous ; vertex of head distinctly narrowed at base ; pronotum with the anterior
margin angularly produced between bases of eyes, posterior margin a little concave before
scutellum, about as long as vertex; scutellum shorter than broad, obliquely depressed on
each side, the basal angles depressed.
Long. 5 mm.
Loc. Aldabra: fle Michel, X. 1908; Takamaka, 1908 (Fryer).
Subfamily Pleinge.
139. Plea letourneuaz, Sign.
Plea Letourneuxi Sign., Bull. Soc. Ent. France, 1880, p. xxxiv.
Loc. Seychelles. Mahé: marshes on coastal plain at Anse aux Pins and Anse
Royale, I. 1909. Originally described from Hgypt.
EXPLANATION OF PLATES 11—13.
Puate 11.
Fig. 1. Chilocoristoides felicitatis, gen. et sp. n. a, odoriferous aperture.
Wig. 2. Fe mahensis, gen. et sp. n.
Fig. 3. Husarcocoris imperator, sp. u.
Fig. 4. Stenozygum aldabranum, sp. n.
Fig. 5. Orbellis typicus, gen. et sp. nu.
Fig. 6. Hyginellus gayei, gen. et sp. n. a, rostrum and body beneath.
Fig. 7. Aphanus consocialis, sp. n.
Fig. 8. Letheus stellatus, sp. n.
Fig. 9. Lachnophorus albidomaculatus, sp. n.
Fig. 10. Cligenes gardineri, sp. n.
SECOND SERIES—ZOOLOGY, VOL. XVI. 25
190 PERCY SLADEN TRUST EXPEDITION
Fig. 11. Corizus scotti, sp. n.
Fig. 12. Pamera sladent, sp. n.
Fig. 13. Dieuches cardwi, sp. n.
Fig. 14. Jourdainana rugifer, gen. et sp. n.
Fig. 15. Cantinona precellens, gen. et sp. n.
Fig. 16. Dulinius nigrolineatus, sp. n.
Mig. 17. Pictinws varicolor, sp. n.
Fig. 18. Aneuwrus bicolor, sp. n.
Fig. 19. Prcaultia pronotalis, gen. et sp. n.
Fig. 20. Microvelia repentina Dist.
Fig. 21. ms “9 a
Fig. 22. Yanga seychellensis Dist. a, opercula ¥. 0b, face*.
PuatE 12.
Fig. 1. Maruthas fumigatus, sp. n.
Fig. 2. Ploiariola scotti, sp. n.
Fig. 3. Rochonia galeatus, gen. et sp. n. a, head showing rostrum.
Fig. 4. Polytozus modestus, sp. n. a, spines to scutellum.
Fig. 5. Gardena seychellensis, sp. n.
Fig. 6. Quinssyana typicalis, gen. et sp. n. a, antenna.
Fig. 7. Roslania insularis, gen. et sp. n.
Fig. 8. Nagusta maura, sp. n.
Fig. 9. Salda imsignis, sp. n.
Fig. 10. Seychellesanus typicus, gen. et sp. n.
Fig. 11 ‘i pucturatus, gen. et sp. n.
Hig. 12. i variegatus, gen. et sp. n., typical form.
Fig. 13. ‘a s Re e var. b.
ig. 14. 5 5 Fe i. var. @.
=
ox
Calphurnioides elongatus, sp. n. a, spine to scutellum.
7
ga de
16. Ogeria insularis, gen. et sp. n.
Gen.? alboclavatus, sp. n.
e
eo
Eurystylus alboplagiatus, sp. n.
EoMcocolcoilcoiico
Cio)
pA
JT
ga ©
—_
No)
Derewocoris seychellensis, sp. un.
PLATE 13.
Fig. 1. Dercwocoris? cardui, sp. n.
Fig. 2. Sthenarusoides montanus, gen. et sp. n.
Mig. 3. Maralauda lana, gen. et sp. n.
fo)
Fig. 4. Proboscidocoris pluto, sp. n.
Fig. Hermotinus villosulus, sp. 0.
Fig. Cylapus migratorius, sp. n.
5
6.
Fig. 7. Felisacus auritulus, sp. uv.
Fig. 8. Vannius mahensis, sp. n.
Fig. 9. Fulvius? niger, sp. n.
* This species belongs to the Homoptera and will be enumerated in a subsequent portion of this
contribution relating to that Suborder.
Percy SLtapEN Trust EXPEDITION.
aan) TRANS. LINN. SOC. SER. 2, ZOOL. VOL. XVI. PL 11
H. K. del.
RHYNCHOTA FROM SEYCHELLES
Percy Suapen Trust EXxPeEpItTion.
(Distawt) TOMAINS, JILIN IN, SOC, SIE. 2 ZOOL, WO)L, XWil, IIL, 12
H. K. del.
RHYNCHOTA FROM SEYCHELLES
TRAINS, ILIUNIN, SOC. SIEIR. 2, ZOOIL, WOlL, OWI, IPL. Is
Percy SLapEN Trust EXPEDITION.
(Distant)
H. K. del.
RHYNCHOTA FROM SEYCHELLES
(> . ¢ : : 7
DISTANT—RHYNCHOTA. PART I: SUBORDER HETEROPTERA
10,
, 1,
ig. 12.
ig. 13.
. 14.
ig. 15.
. 16.
Fulvius dolobratus, sp. n.
» pictus, sp. n.
Aretas sangwnarius, sp. n.
Cyrtopeltis atricornis, sp. n.
Megacelum flagellatum, sp. n.
# (Creontiades) mimicum, sp. n.
Lygus cinnamomeus, sp. v.
» sanguineosignatus, sp. n.
» sihouettensis, sp. n.
Psallus? mahensis, sp. n.
Lasiochilus scott, sp. nu.
gardineri, sp. n.
sladent, sp. un.
5 seychellensis, sp. 0.
5 praslinensis, sp. n.
Paralasiocolpus piceus, gen. et sp. n.
"3 marginatus, gen. et sp. n.
Sesellius parallelus (Motsch.).
Ostorodiasoides signatus, gen. et sp. n.
p typicus, gen. et sp. n.
Anisops aldabrana, sp. n.
bo
bo
191
No. X.—COLEOPTERA; HYDROPHILIDA, HISTERIDA.
By Huew Scorr, M.A. (Canras.), F.L.S., F.E.S., Curator in Entomology in the
University of Cambridge.
(Plate 14.)
Read 5th June, 1913.
THIS paper is a report on the Hydrophilide and Histeride of the islands in the
western Indian Ocean visited by the Percy Sladen Trust Expedition in 1905 and
1908—9: that is, the Seychelles, the Farquhar Group, Aldabra and the neighbouring
islands, &c. It is the result of a study of the collection formed by this Expedition,
but species previously recorded from these islands but not collected by the Expedition
are also included in their proper systematic position, so that the paper.may provide a
complete enumeration of the Hydrophilidee and Histeridee known from these groups
of islands. 23 species of Hydrophilide and 16 of Histeridse are here enumerated,
while previously only 6 Hydrophilide and 8 Histeridze had been recorded. 14 new
species and 3 new genera are described.
As regards general considerations of affinities, distribution, &c., the two families
are considered separately, a brief general account being prefixed to the systematic account
of each on pages 194 and 222 respectively.
Types. A first set of all the material, including the types of all new species and
genera, will be given to the British Museum. A second set will be kept in the Cambridge
University Museum.
Localities. Under each species is given a locality-list including the fullest possible
data as to where and when it was found. The dates “1908” or “1908—9” standing
alone (z.e., not followed by a collector's name) after any data, always indicate that the
specimens were collected by myself, or by some of those who helped me while I was
in the Seychelles. Similarly “1905” standing alone indicates that the material was
obtained by the earlier Perey Sladen Trust Expedition (that is, the Expedition of
H.M.S. “Sealark”). In all other cases the collector's name is placed in brackets after
the data. When a species is found outside the area of the Expedition, a brief summary
of its external distribution is given at the end of the locality-list.
Literature. In the literature-references the abbreviation “ Alluauid, Liste Coléopt.”
stands for Alluaud’s “Liste des Insectes Coléoptéres de la Région Malgache” (vol. xxi.
of Grandidier's “Histoire Physique, Naturelle et Politique de Madagascar,” 1900). I
have frequently had to refer to papers by Régimbart, Ann. Soe. ent. France, vol. 1xxii.
1903, and vol. Ixxv. 1906. The Histeridz collected in the Seychelles by Alluaud in
1892 were reported on by J. Schmidt in Bull. Soc. ent. France, 1893, p. 99. Certain
* Almost entirely the same system is followed as in my paper on the Lamellicornia and Adephaga, in
vol. xv. of these Transactions, 1912, p. 215.
194 PERCY SLADEN TRUST EXPEDITION
other matters connected with the literature are mentioned in the writer’s paper in
vol. xv. of these Transactions, 1912, p. 215.
It is a pleasure to express my thanks for the generous help which I have
received from several entomologists in dealing with these families of Coleoptera. Monsieur
Charles Alluaud has very greatly assisted me by lending a number of specimens for
purposes of comparison. I am specially indebted to Monsieur P. Lesne for his kindness
in lending me valuable specimens from the Paris Museum, and for comparing specimens
which I have sent to him with species in the Paris Collections. Monsieur A. d’Orchymont
has examined a number of the Hydrophilide ; I am indebted to him for several deter-
minations and for allowing me to make absolutely unrestricted use of his very full and
accurate manuscript-notes
receive certain minute forms among them.
on them. Lastly I have to thank Mr George Lewis for
determining certain of the Histeridze and for the gift of several valuable specimens.
Hydrophilide.
23 species, representing 12 genera, are here enumerated from the islands visited
by the Expedition. Only 6 species had previously been recorded. Of the 17 additions,
9 are described as new to science, and it has been necessary to erect 3 new genera to
22 species are actually represented in the
collection made by the Expedition : the remaining one (C@lostoma punctulatum) has been
recorded as found by a previous collector.
The distribution of the species, so far as known, is shown in the following table:
Before the names of the species:
| * indicates that the species is new to science,
+ indicates that the species is recorded from
these islands for the first time, but not
new to science.
Distribution among the islands
visited by the Expedition
Seychelles| Coetivy
Group Island aallatoronra
External Distribution
* Hydrena mahensis............ |
7 Berosus acutispina............
+ Berosus prolongatus .........
Berosus bergrothi ............
Volvulus stagnicola ......... |
* Paracymus alluaudianus
Helochares melanophthalmus
Helochares melanophthalmus )|
var. sechellensis {|
8. Philydrus parvulus .........
9. 7 Philydrus abnormalis.........
10. *Paromicrus carinatus.........
11. *Paromicrus thomasseti ......
12. *Paromicrus atomus............
13. Dactylosternum insulare ...
14. 7 Dactylosternum pygmeum
15. fCclostoma rufitarse .........
16. Celostoma punctulatum
17. *Bourdonnaisia mahensis ...
18. *Bourdonnaisia silhouette ...
19. *Cercyon fructicola ............
20. +Cercyon laticollis ............
| 21, +Cereyon uniformis ............
Ri Sd Sty G9 GS =
22, CROC Ds scasooocesecodoenac00
23. *Paroosternum degayanum...
Madagascar
Madagascar
Madagascar
Trop. Africa
Madagascar, Africa, Trop. Asia, S. Europe
| Madagascar, Africa, India, ce.
Madagascar, British E. Africa, Ceylon, [? Sumatra]
All warmer regions of Old World
Mauritius
Madagascar, E. and 8. Africa
Madagascar, Africa
| Madagascar
Ceylon, India
Y
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 195
It is seen that no Hydrophilidze were collected in the Chagos, Amirantes, Farquhar,
or Cargados Carajos groups of coral-islands, and none have ever to my knowledge been
recorded from those groups.
The table shows that 18 species were found in the Seychelles: 15 of these were
found only in that archipelago (and, in the case of two of them, in the outlying
coral-island of Coetivy, close to the South); 3 species were found also in Aldabra.
9 species in all were found in Aldabra, that is the three (all widely distributed species)
which occur also in the Seychelles, and six others. In discussing affinities it will be
best to consider the Seychelles and Aldabra separately.
Seychelles. Of the 18 species, 13 are terrestrial and 5 are aquatic. Among the
terrestrial forms there is almost certainly an endemic element in the fauna, consisting
of the completely wingless new genus Bowrdonnaisia, represented by two distinct
species each confined to a single island: the extreme restriction of their habitat to
the very summits of the highest peaks, in the heart of the endemic forests, is dealt
with on p. 215. In affinities this genus appears to be isolated.
The second new genus, Paromicrus, is fully winged, and represented by three
minute species, not confined to single islands nor so restricted in habitat as Bowrdon-
nosia, but still almost entirely confined to the endemic forests, where they are
amongst the most abundant beetles under the bark of decaying wood (see p. 207).
This genus is allied to the Hawaiian Omicrus: at first I thought it was endemic to
the Seychelles, but M. d’Orchymont has informed me (in /itt.) that he has two other
species, one from Engano, a small island south-west of Sumatra, the other from the
Abor country (Assam), Neither of these is identical with any of the Seychelles species,
specimens of all of which were submitted to M. d’Orchymont: so the Seychelles species
may perhaps be confined to those islands, though the genus is not.
The third new genus, Paroosternwm (also fully winged), is represented by only two
specimens: it is allied to a genus known from Central America and Japan. Such
affinities may seem remarkable, but until Hydrophilide so minute as these have been
more thoroughly collected throughout the world, it would be unwise to attach too
much weight to them.
Among the remaining land-inhabiting Hydrophilide it is doubtful whether there
is any endemic element at all. Dactylosternwm insulare (found also in Aldabra) is
very wide-spread: Dactylosternum pygmeum is known from Mauritius: Celostoma
punctulatum is wide-spread African and Madagascan: Cercyon fructicola may perhaps
be allied to a form known from Mauritius : Cercyon laticollis is known from Madagascar ;
Cercyon uniformis from Ceylon and India. These species are all found in rotting fruit.
—very often in the fruit of imported trees—at various elevations (see locality-lists),
and, with the possible exception of Cercyon fructicola, are probably immigrant or
imported species.
Turning now to the aquatic forms: the writer has several times mentioned* that
the very numerous mountain-streams of the Seychelles have only a small aquatic insect-
* Trans. Linn. Soc. London, Ser. 2, Zool., xiv. 1910, p. 28: ibid. xv. 1912, pp. 242, 258.
196 PERCY SLADEN TRUST EXPEDITION
fauna, and that only one species of water-beetle (the Dytiscid Copelatus gardinert) was
found in them in any numbers. This is fully borne out by the aquatic Hydrophilide :
only two specimens (one of Phalydrus abnormalis and one of Philydrus parvulus var.)
were found at any considerable elevation, both of them in the Mare aux Cochons plateau
(Silhouette), over 1000 feet above sea-level. All the other aquatic Hydrophilidee
which I collected in the Seychelles were found quite near sea-level, either in the
coastal swamps or estuary at Anse Royale, Mahé, or in a pond near the beach in
Félicité Island. Hydrena mahensis, though described here as a new species, is very
closely allied to or possibly only a form of a Madagascar species: Paracymus alluaudianus
(found in Coetivy), also described as new, is allied to a species found in Madagascar
and in Kast Africa and Eritrea: Helochares melanophthalmus var. sechellensis is a local
form of a very wide-spread species: Philydrus parvulus is very widely distributed in
the warmer regions of the Old World, and occurred in Coetivy and Aldabra as well
as in the Seychelles: Philydrus abnormalis occurs in Madagascar, British East Africa,
Seychelles (Silhouette), Ceylon, and possibly also in Sumatra.
Aldabra. 7 of the 9 species are aquatic and 2 terrestrial. The three species which
occur also in the Seychelles (Philydrus parvulus, Philydrus abnormals, Dactylosternum
imsulare) are all very wide-spread, as stated above: a fourth species Helochares melanoph-
thalmus (represented in the Seychelles by a special form) is also very wide-spread, Of
the 5 remaining species, the 3 species of Berosus are all known from Madagascar (one
of them, Berosus acutispina, having very close allies in Africa and apparently also in
Tonkin) ; Celostoma rufitarse is Madagascan and African: the Volvulus is somewhat
doubtfully referred to an African species. The Aldabran Hydrophilid fauna therefore
contains Madagascan and African elements, and an element consisting of species very
widely spread in the warmer parts of the Old World. Its general nature thus appears
similar to that of the Lamellicorn and Adephagous fauna of Aldabra (see vol. xv. of these
Transactions, 1912, pp. 220 and 242).
Reduction of wings. As mentioned above, the new genus Bowrdonnaisia is
completely apterous (see p. 214). No other case of reduction of the alar organs was
found in the Hydrophilide. [An interesting case was found in the Histeride, see
pp. 224, 230.]
Hydreenini.
Hyprana, Kugelann.
Subgenus Hyprana, s. str., Ganglbauer, Kif. Mitteleur., iv. 1, 1904, p. 197.
1. Hydrena mahensis, sp. nov.
Minuta, sat nitida, capite nigricante, pronoto medio infuscato marginibus sat late
ferrugineo-testaceis, elytris fusco-testaceis, antennis palpisque pedibusque testaceis : capite
sat dense punctato: palpis valde elongatis, articulo 2 fere zque longo ac 3 + 4, extrorsum
arcuato, gracillimo, apice parum incrassato: prothorace transverso, ante medium parum
angustato, angulis anticis rotundatis, pone medium fortius angustato et lateribus leviter
SCOTT—COLEOPTERA ; HYDROPHILIDA, HISTERID”% 197
sinuatis, angulis posticis argutis fere rectis, margino antico zque lato ac basi, marginibus
lateralibus tenuiter crenulatis, disco fortiter dense punctato, utrinque ad latera, antice
fortius, impresso: elytris sat curtis, in parte media quam prothorace latioribus, apice
separatim rotundatis et sutura dente minuta instructa, marginibus lateralibus explanatis,
arcuatis ; sat fortiter dense seriatim punctatis, intervallis angustis (zeque latis ac seriebus),
planis, nullo modo elevatis. Long. corp. circa 14 mm.
General colour brownish-testaceous: head darker, almost black; pronotum with a
roughly rectangular transverse median area darkly infuscate, and the margins rather
widely ferrugino-testaceous ; elytra brownish-testaceous ; antennee, palps, legs and feet
testaceous. Head: pre-frons finely punctured, nearly smooth in the middle, post-frons
more strongly punctured: fronto-antennal suture nearly straight: labrum large and
bilobed : maxillary palpi very long, jomt 2 very long, about as long as 3 and 4 together,
curved outwards, very slender from the base to nearly ? of its length, slightly swollen
in its apical portion ; joint 3 slender at the base, in its distal half about as swollen as
the apex of 2; joint 4 about 14 times as long as 3, more swollen, convex on the inner
side and nearly straight outwardly : antennze with joint 2 as broad as the apex of 1,
joint 3 very minute, almost invisible, 4 produced inwardly into a finely-pointed process,
club 5-jointed and pubescent. Pronotwm transverse, having its greatest breadth at
a point about the middle, with the sides slightly narrowed in front of this poimt to
the anterior angles, which are rounded ; with the sides behind the widest point more
strongly narrowed, and sinuate in such a way that the posterior angles are definite
and nearly right angles: width of anterior and posterior margins about equal, that
of the anterior greater than that of the head and eyes: lateral margins finely crenu-
late: disc closely and strongly punctate, much more strongly than the head; with
an impression on either side quite near the lateral margin, broader and more marked
in front, gradually narrowing towards the posterior angles. Scutellum visible but
minute. Hlytra about 24 times as long as the pronotum, with lateral margins curv-
ing outwards from the base, so that a little behind the shoulder the elytra become
considerably broader than the pronotum even at its broadest point: lateral margins
explanate, very finely crenulate near the shoulder and near the apex, the crenulations
much less close than on the thorax: apices separately rounded, with a minute tooth
present at the apex of the sutural margin: each elytron bears about 15 or 16 series
of punctures, 9 of which lie between the humeral callus and the suture: the punctures
are about the same size as those on the thorax, in each series separated from one another
by about their own diameter; while the intervals between the series are about the
same width as the series themselves and not at all raised or convex. From each
puncture rises a minute pale procumbent hair, not easily visible: some such hairs
are also visible on the head and thorax. The nature of the punctuation is such that
its serlate arrangement is not always very apparent.
Beneath, the body is covered with pale pubescence. There are no raised ridges on
the metasternum; but in the one specimen examined the pubescence is thinner on two
longitudinal areas, one on either side of the middle line of the metasternum. In this
specimen also the 5th abdominal segment and part of the 6th appeared nearly bare
SECOND SERIES—ZOOLOGY, VOL. XVI. 26
198 PERCY SLADEN TRUST EXPEDITION
of pubescence, but I could not quite determine whether this was natural or due to
abrasion; lack of material prevented my determining the sex of the specimens.
This species is very closely allied to the Madagascan Hydrena marginicollis
Régimbart (Ann. Soe. ent. France, Ixxi. 1903, p. 51), and may possibly prove to be
only a variety or small local race of that species. I have been able to study a typical
specimen of margimicollis through the kindness of M. Alluaud: it is considerably
larger ; has the prothorax slightly more narrowed in front of the middle, and a little
more narrowed and with sides more strongly sinuate behind the middle; the palps
are formed in the same way, but the apex of the long 2nd joint, and the terminal
joint, appear slightly less incrassate than in mahensis. Also marginicollis has the
elytra a little less strongly separately rounded at the apex, and the angle between
them (at the end of the suture) a little less deep: but the form of the elytra at the
apex might depend on the sex, which smallness and fragility of material have prevented
my determining either in mahensis or marginicollis. [Beneath marginicollis resembles
mahensis : the apical part of the 5th segment of the specimen before me is nearly
bare of pubescence, the basal part of the 6th is widely bare and smooth, while the
apical part of the 6th has scanty pubescence. |
Loc. Seychelles. Mahé: marshes on coastal plain at Anse aux Pins and Anse
Royale, 19—21. I. 1909, 2 specimens.
Hydrophilini.
Berosus, Leach.
Subgenus ENopturus, Hope, Col. Man., ii. 1838, p. 128.
2. Berosus (Hnoplurus) acutispina, Fairmaire.
Berosus acutispina Fairmaire, Ann. Soc. ent. France (sér. 4), vill. 1868, p. 196;
Régimbart, op. cat., lxxil. 1903, p. 36; Alluaud, Liste Coléopt., p. 228.
29, 4 9, from Aldabra. There is much variation even in this small series, specimens
varying in length from 443—64 mm. The colour is also variable; the dark marblings
on the posterior part of the elytra vary, being reduced in some specimens while in
others they form a double oblique dark band with a much paler portion between its
two parts™.
The determination was confirmed by Monsieur Lesne, who states that he can find
no differences between specimens submitted to him and Fairmaire’s type. Monsieur
d’Orchymont, to whom I had previously submitted specimens, considered them to be
the East African Berosus gracilispina Régimbart (Ann. Soc. ent. France, Ixxv.
1906, p. 264), which may possibly be the same as Erichson’s Berosus cuspidatus from
* Monsieur d’Orchymont called my attention to variation in the manner in which the striz meet one another
at the apex of the elytron: he informs me that in the subgenus Hnoplurus they usually meet in the following
pairs: 1 and 2, 3 and 8, 4 and 7, 5 and 6, 9 and 10. But in 1 ¢ and 2 9 of the Aldabra B. acutispina, the
pairs are: 1 and 2, 3 and 4, 5 and 6, 7 and 8,9 and 10. Monsieur d’Orchymont has observed this variation also
in specimens of B. (Hnoplurus) indicus Motsch.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 199
Angola. Monsieur d’Orchymont compared the Aldabra specimens with a 2 from Tonkin
determined as Berosus acutispina Fairmaire: he found certain differences in punctuation
and form, especially in the form of the tarsal claws, which are much shorter in the
Aldabra specimens, and in the front tarsus more arcuate, than in the Tonkin specimen.
In view of Monsieur Lesne’s determination it appears possible that the Tonkin specimen
is not of the same species as the true acutispina described by Fairmaire from
Madagascar.
I am unable to say if the Madagascan acuwtispina Fairmaire is the same as the
African gracilispina Régimbart, but at least they seem to be extremely closely allied.
Loc. Aldabra: from “ Wilson’s Well,” Takamaka, X.—XI. 1908 (Fryer). Mada-
gascar.
3. Berosus (Hnoplurus) prolongatus, Fairmaire.
Berosus prolongatus Fairmaire, Ann. Soc. ent. France (sér. 4), viii. 1868, p. 196;
Régimbart, op. cit., lxxii. 1903, p. 36; Alluaud, Liste Coléopt., p. 229.
3 g, 1 2 from Aldabra: compared with a specimen (from Madagascar) kindly lent
from the Paris Museum by Monsieur Lesne, who informs me that it appears quite similar to
Fairmaire’s type. The Aldabra specimens are very much paler in colour, being almost
entirely yellow: only the posterior part of the head, the extreme apex of the terminal
joint of the maxillary and labial palps, the two longitudinal patches on the pronotum, the
strie* and punctures on the interstices of the elytra, and some vague markings on the
anterior and posterior parts of the elytra, are darker. In three of the Aldabra specimens
the prothorax is a little shorter in proportion to its width than in the specimens lent from
Paris.
The punctuation on the pree-frons is finer and considerably less dense than on the
post-frons, where it is coarser and closer, especially behind (at the extreme anterior margin
of the pree-frons it is very fine and rather close). The intra-ocular systematic punctures
form a line of confluent punctures nearly parallel to the margin of the eye and larger than
the general punctuation. The scutellum is an elongated triangle with sharp apex, with
a rather wide border raised a little above its general surface, and a line of punctures along
either side against the border, the median part of the surface being impunctate. The
setiferous punctures on the elytra are well marked but few, being most numerous on the
3rd, 4th, 5th and 11th interstices. The one ? has the apex of its elytra a little more
narrowly and deeply truncate-emarginate than the 3 ¢.
Beneath, the prosternum and anterior coxee are pale yellowish; remainder of the
underside dark, almost black, the middle coxze and mesosternum being slightly more
dilute, very dark reddish-black : basal part of the femora scarcely infuscate in the anterior
pair, almost black in the middle and posterior pairs; remainder of the femora, tibize and
tarsi reddish-yellow. Under-surface of body, cox and basal portions of femora dull,
* Tn all 4 specimens of B. (Enoplurus) prolongatus the manner in which the strize meet one another at the
apex of the elytron is almost that which d’Orchymont informs me is normal in this subgenus: cf. footnote on
B. acutispina, p. 198.
26—2
200 PERCY SLADEN TRUST EXPEDITION
very minutely and closely rugose-punctate, covered with very fine, short, pale pubescence,
except the mesosternum.
Loc. Aldabra: from “ Wilson’s Well,’ Takamaka, X.—XI. 1908 (Fryer).
Madagascar.
Subgenus BERosus, s. str., Ganglbauer, Kaif. Mitteleur., iv. 1, p. 220.
4. Berosus bergrothi, Régimbart.
Berosus bergrothi Régimbart, Bull. Soc. ent. France, 1900, p. 51, and Ann. Soe. ent.
France, lxxii. 1903, p. 40; Alluaud, Liste Coléopt., p. 229; Kolbe, Abh. Senckenb. Ges.,
xxvi. 1902, p. 574.
A large series of about 260 specimens, agreeing well with Régimbart’s description.
But there are certain variations. Régimbart stated that the dark metallic patch on the
disc of the prothorax is not divided by a pale median line: in a number of the specimens,
however, it is narrowly divided, completely or almost completely. Some specimens have
the anterior margin of the prothorax dark in the middle part, behind the head ; but this
dark colour is not united to the front of the median patch, which never reaches either the
front or hind margins.
Three specimens exhibit more striking variation. In one of them, a @, the dark
thoracic patch is reduced to two very small patches, completely separated and situated in
the posterior part of the disc. In the two others, ??, the dark mark is entirely absent,
the prothorax being uniformly pale: these also have the dark elytral marks much reduced,
in one specimen practically absent. They show no signs of immaturity, the head being
entirely dark bronze with metallic reflections, as in all other specimens.
Régimbart also stated that the thoracic punctuation is not interrupted by a smooth
median space: this is quite true in many specimens, but in some a slight indication of a
median longitudinal smooth line can be seen with a high power.
Judging by Régimbart’s descriptions (/.c.), this species must be closely allied to
Berosus villosulus Rég. (Madagascar) and B. viticollis Boh. (S. Africa and Madagascar).
Loc. Aldabra: Takamaka, X.—XI. 1908 (Fryer); 1895 (Voeltzkow). Madagascar.
Votvuuus, Brullé*.
5. Volvulus stagnicola (Mulsant).
Brachygaster stagnicola Mulsant, Ann. Soc. Linn. Lyon (sér. 2), 1. 1853, p. 385,
and Opuse. Ent., u. 1853, p. 177; Alluaud, Liste Coléopt., p. 229.
Volvulus stagnicola Régimbart, Ann. Soc. ent. France, xxv. 1906, p. 267.
Brachygaster denticulata Régimbart (nec Muls.), Ann. Soc. ent. France, lxxi. 1903,
p. 41.
Volvulus cupreus Régimbart, Bull. Soc. ent. France, 1900, p. 51; Alluaud, Lc. ;
Kolbe, Abh. Senckenb. Ges., xxvi. 1902, p. 574.
* The name Regimbartia is proposed for this genus, and the synonymy dealt with, by Zaitzev in his Cat.
Hydrophil., Horze Soc. Ent. Ross., xxxviii. 1908, p. 362.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 201
Régimbart first described specimens from Aldabra as cupreus: in 1903 (U.c.) he
stated that he considered this a synonym of denticulata: and again in 1906 (l.c.) he
considered that this was a mistake, and that the Aldabra specimens were referable not to
denticulata, but to stagnicola. The name cupreus then became a synonym of stagnicola,
and as such it is given by Zaitzev in his Catalogue (1908, p. 362).
In the absence of types wherewith to compare it, I refer a single 3, which agrees with
Régimbart’s descriptions, to this species. Length, including the extended head, 5mm. ;
palps reddish, legs black, slightly ferruginous : head fairly closely punctured : thorax less
closely punctured in the middle, nearly as closely as the head at the sides: elytra with
the two strize nearest the suture effaced towards the base over about + of their length,
the 3rd and 4th faint towards the base, the 5th reaching almost to the base, the 6th and
7th abbreviated a little before the base (the 6th ending in the region of the callus, in
which region its punctures become widely spaced out, and the 7th ending a trifle further
from the base than the 6th), and the 8th, 9th and 10th (ze. the outermost) reaching
practically to the base: striz much deeper behind, crenulate-punctate : the punctures of
the intervals much finer than those of the strize, and the hairs which they bear extremely
short and fine and hard to see (possibly the specimen is worn; but the pubescence is less
conspicuous than in several specimens determined as V. enews Brullé which I have before
me).
Loc. Aldabra: Takamaka, 1908, 1 g¢ (Fryer); 1895 (Voeltzkow). Volvulus
stagnicola is widely spread in Tropical Africa (see Régimbart, 1906, l.c.: V. denticulatus
is known from Madagascar).
Paracymus, Thomson.
Subgenus ParacyMus, s. str.
6. Paracymus alluaudianus, sp. nov.
Supra nigro-zeneus, prothoracis lateribus et elytrorum apicibus obscure rufescentibus,
antennis palpisque rufo-ferrugineis, antennarum clavis atque articuli 4' palporum apice
infuscatis ; subtus capite, meso- et metasternis, abdomineque nigris, prosterno pedibusque
rufo-ferrugineis : capite pronoto elytrisque dense fortiter punctatis: mesosterno pone
dentem anteriorem carinula longitudinali instructo ; femoribus mediis pubescentid haud
ultra medium attingente. Long. corp. 2 mm.
Shining, with a strong brassy lustre; antennze, palps and legs rather dark reddish ;
beneath, the head, meso- and metasterna and abdomen are black, while the prosternum is
reddish. The species is characterised by the strong and dense punctuation of the upper
surface : on the head the punctures are very small near the anterior margin, larger behind,
the largest being on either side near the eye ; on the pronotum the punctures are a little
finer and further apart than those on the elytra: on the elytra they are densest and
largest, often not separated by much more than their own diameter, and especially close
in the scutellar region. The elytra have no trace of seriate punctuation: the sutural
stria is well marked, abbreviated in front. Under high power the elytral punctures
appear simple and circular.
202 PERCY SLADEN TRUST EXPEDITION
Paracymus alluaudianus is closely allied to the Madagascan and Hast African
Paracymus chalceus Régimbart (Ann. Soc. ent. France, Ixxi. 1903, p. 32), a cotype of
which I have been able to study through the kindness of M. Lesne. Its upper and under
sides are coloured similarly to those of allwaudianus, but it is a trifle larger, and at once
distinguished by its much finer and less dense punctuation. This difference is very
noticeable when the two are viewed side by side. In chalceus (as in allwaudianus) the
elytral punctures are simple and circular.
Among the European species, w@neus Germar is separated from scutellaris Rosenh. and
punctillatus Rey by having an elevated longitudinal lamina on the mesosternum behind
the tooth and transverse crest, and by having the pubescence on the middle femora
scarcely reaching to the middle: seuwtellaris and punctillatus having no longitudinal
elevated lamina on the mesosternum, and the pubescence reaching beyond the middle of
the middle femora (Ganglbauer, Kif: Mitteleur., iv. 1, p. 241). In these respects
alluaudianus and chalceus approach eneus Germ., having a small but distinct elevated
lamina, and the pubescence scarcely reaching to, and certainly not beyond, the middle of
the middle femora.
Paracymus evanescens (Sharp) from Ceylon and India (of which a typical specimen
from the British Museum is before me) agrees with alluaudianus and chalceus in
possessing a small elevated mesosternal lamina and in having the pubescence not reaching
beyond the middle of the middle femora. But it is much smaller than either of those
species (the specimen before me measures only 1} mm.) and quite distinct from them in
punctuation (see below): also the underside (in this specimen at any rate) is, with the
exception of the head, much lighter-coloured, being yellowish.
Paracymus minor Régimbart (Ann. Soe. ent. France, Ixxii. 1903, p. 33) has a minute
longitudinal elevation on the mesosternum, but in it the pubescence extends beyond the
middle, to about ? the length of the middle femora: in this latter respect it differs from
alluaudianus, chalceus, evanescens, &c., and approaches scutellaris Rosenh. &e. But
though minor differs thus from evanescens, yet in their upper surfaces these two species
are rather closely alike. They are the same size, and in both of them the elytral
punctures, instead of being simple and circular, have the form of minute figures-of-8, as
though each puncture were formed by the coalescence of two”: in this point therefore
they differ widely from alluaudianus and chalceus. In evanescens the cephalic and
thoracic punctures are larger and subobsolete, in minor finer and clearer cut. Paracymus
evanescens and minor are therefore quite distinct from one another, and from alluawdianus
or chalceus: minor is known from Madagascar and perhaps occurs also in British Kast
Africa; I have been able to study the type of the species through the kindness of
M. Alluaud.
Paracymus alluaudvanus is named in honour of Monsieur Charles Alluaud.
Loc. Seychelles: Mahé; marshes on coastal plain at Anse Royale, I. 1909, 66
specimens. Coetivy; 1905, 2 specimens.
* Note, this only applies to the elytral punctures: cephalic and thoracic punctures are simple.
SCOTT—COLEOPTERA; HYDROPHILID®, HISTERIDAL 203
Hetocuares, Mulsant.
7. Helochares melanophthalmus, Mulsant.
Helochares melanophthalmus Mulsant, Hist. nat. Col. France, Palpicornes, 1844,
p. 1387; Alluaud, Liste Coléopt., p. 231; Régimbart, Ann. Soe. ent. France, Ixxu. 1903,
p- 27; Kolbe, Abh. Senckenb. Ges., xxvi. 1902, p. 574; id. Mitt. Zool. Mus. Berlin,
Wo UQWO, jos Zt.
Helochares nigrifrons Régimbart, Bull. Soc. ent. France, 1900, p. 50.
Régimbart first determined (1900, /.c.) specimens from Aldabra as H. mgrifrons
Branesik, stating that he was inclined to consider it as simply a form of melanophthalmus :
in his later paper (1903, J.c.) he still refers the Aldabra specimens to it, but treats it
as var. ngrifrons of melanophthalmus ; Kolbe states (1902, l.c.) that mgrifrons Brancsik
cannot be separated from melanophthalmus. But Zaitzev in his Catalogue (p. 381, 1908)
treats the two as distinct species. I have been unable to investigate the matter and
follow Régimbart and Kolbe in referring the form found in Aldabra to melanophthalmus.
Loc. Aldabra: Takamaka, 1908, 12 specimens (Fryer): 1895 (Voeltzkow).
Madagascar, Africa, 8. Kurope.
Helochares melanophthalmus Mulsant var. sechellensis, Régimbart.
Var. sechellensis Régimbart, Ann. Soc. ent. France, Ixxii. 1903, p. 27; Kolbe, Mitt.
Zool. Mus. Berlin, v. 1910, p. 24.
This form is clearly distinguished from that found in Aldabra by its finer and less
dense punctuation, and consequently more shining appearance, and by its generally larger
size. ‘The specimens before me agree well with Régimbart’s description.
Loc. Seychelles. Félicité: from a pond in a plantation at Grande Anse, XII. 1908,
7 specimens. Mahé: marshes on coastal plain at Anse Royale, I. 1909, 5 specimens; 1896
(Brauer). La Digue: 1892 (Alluaud).
PuityprRus, Solier.
Subgenus Merayprus, Rey; Ganglbauer, Kif. Mitteleur., iv. 1, p. 243.
8. Philydrus (Methydrus) parvulus, Reiche.
Philydrus parvulus Reiche, Ann. Soc. ent. France, sér. 3, iv. 1856, p. 359;
Régimbart, Bull. Soc. ent. France, 1900, p. 50; id., Ann. Soe. ent. France, Ixxu. 1903,
p- 31; Alluaud, Liste Coléopt., p. 231; Kolbe, Abh. Senckenb. Ges., xxvi. 1902, p. 574;
id., Mitt. Zool. Mus. Berlin, v. 1910, p. 24.
Régimbart’s remarks (J.c.) about the extreme variability of this species in colour, size,
&c., are amply borne out by the large series of over 140 specimens before me. Some
specimens are small and very pale, others much larger and dark, and there are various
intermediate gradations. This variation occurs, to a large extent at any rate, in-
dependently of sex. The triangular yellow marks on the head in front of the eyes appear
to be constant in both sexes.
204 PERCY SLADEN TRUST EXPEDITION
In both sexes there is a blunt tooth at the base of both claws of all the feet. In
the ¢ the claws of the anterior tarsi are much more sharply bent, and the teeth more
prominent, than in the 2. In the claws of the middle and posterior tarsi I have not been
able to make out any appreciable difference in the two sexes, neither does there appear to
be any appreciable difference between the outer and inner claws in either sex. This does
not agree with Kuwert’s statement (Verh. naturf. Ver. Briinn, xxviii. 1889, p. 56),
““* Aeussere Krallen simmtlicher Fiisse mit einem langen scharfen Zahn.” The exact form
of the claws is not easy to determine, even with a high power.
Both sexes have a clear-cut, rather deep, rounded emargination, set with a series of
yellowish seta, in the middle of the posterior margin of the 5th ventral segment. The
whole segment is pubescent, and the pubescence is sometimes rather denser at the hind
margin immediately on either side of the emargination: but the sete fringing the
emargination are of quite a different nature, being much stouter at the base and tapering
to the apex, set in a single regular series close together. Kuwert (/.c.) stated of this
species: “ Fiinftes Segment mit einem kleinen gelbhaarigen Griibchen am Ende.” But
in the series before me the emargination is present in both g and ?, and apparently of
the same form in both sexes. The segment is sometimes a little impressed near the
emargination, and when the setze are stuck together they render its nature less apparent :
this might possibly account for Kuwert’s use of the word “ Griibchen”
“* Ausschnitt.”
Loc. Seychelles: Mahé; coastal plain at Anse Royale, I. 1909; 1896 (Brauer) :
Félicité, from a pond at Grande Anse (sea-level), XII. 1908. Coetivy, 1905. Aldabra:
Takamaka, 1908 (Fryer); 1895 (Voeltzkow). Madagascar, Tropical Africa, Egypt, Syria,
India.
instead of
Philydrus parvulus, Reiche, var. 4
A single ? has the entire prze-frons yellow, in contrast to the black posterior part of
the head. It appears as though the triangular yellow patches have extended across the
head and united: in the other specimens they seem to be constant, though Régimbart
states (/.c.) that they are variable in this species. The specimen also has the 3rd joint of
the maxillary palpi a trifle shorter in proportion to the 2nd, and the thoracic and elytral
punctuation slightly finer and less dense, than in other females of the series.
In having the preze-frontal region entirely yellow and the punctuation finer, this
specimen resembles Philydrus fragilis Sharp (Ceylon: Trans. Ent. Soc. London, 1890,
p. 350). But it agrees with the other specimens of parvulus in size and general form and
in the stronger punctuation of the head. Phalydrus fragilis is larger than the largest of
the series of Philydrus parvulus, and slightly more oval in outline, owing to the side-
margins of the elytra being more curved: its punctuation, especially on head and thorax,
is finer and more obsolete, even than that of the var. of parvulus under consideration.
The specimen of fragilis before me has on either side of the head, in the region of the
intra-ocular systematic punctures, a slight impression which I have not seen in parvulus :
its yellow pree-frons is obscured in the middle, whereas that of the Philydrus parvulus
var. from Silhouette is entirely yellow.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 205
Loc. Seychelles. Silhouette: Mare aux Cochons plateau, over 1000 feet, IX. 1908.
It is the only specimen from Silhouette, and the only one from the Seychelles mountains,
all the other Seychelles specimens having been found almost at sea-level.
Subgenus Puityprus*, s. str., Horn; Ganglbauer, Kaf. Mitteleur., iv. 1, p. 243.
9. Philydrus abnormalis, Sharp.
Philydrus abnormalis Sharp, Trans. Ent. Soc. London, 1890, p. 351.
Enochrus abnormicollis} Zaitzev, Cat. Hydrophilidee, Horze Soc. Ent. Ross., xxxviil.
1908, p. 385.
Philydrus mollis Régimbart, Ann. Soc. ent. France, Ixxii. 1903, p. 32; id., op. cit.,
Ixxv. 1906, p. 263.
(2) Philydrus rubricollis Régimbart, t.c., p. 58.
Four specimens : determined by comparison with a typical specimen of abnormalis
in the British Museum (Mr George Lewis’ Collection), with which they closely agree.
By the kindness of Monsieur Lesne I have also been able to compare them with a
cotype of Régimbart’s mollis: this is immature (Régimbart stated that almost all the
specimens on which his description was based were immature), and consequently
appears very different in some ways, being light yellow instead of the normal dark
blackish-brown ; but a close examination has convinced me that it is identical with
Philydrus abnormalis. Also Monsieur d’Orchymont has informed me that the Seychelles
specimen submitted to him is identical with one from Tamatave in his collection.
Philydrus abnormalis and Philydrus mollis are therefore the same: possibly the
Sumatran Philydrus rubricollis Régimbart is also identical, as its description agrees
and Régimbart stated that it might be so. Two other species, known from both
Indo-China and Sumatra, appear to be closely allied, having the palps very greatly
elongated in the same manner, &c.; they are Philydrus ferrugatus Régimbart and
P. mgritulus Régimbart, described in the same paper with rubricollis (t.c., pp. 57, 58).
The four specimens are dark brownish-black, with the labrum and the sides of
the head in front of the eyes, the four margins of the pronotum narrowly, and the
outer margins of the elytra more broadly, dark reddish; antennze and the greatly-
elongated palpi testaceous. Beneath, dark reddish-brown or pitchy-reddish, the
abdominal segments blackish with posterior margins dark reddish: surface dull,
the metasternum, femora, and abdominal segments covered with very short, fine, pale
pubescence: legs dark reddish-brown, femora somewhat infuscate, tarsi paler, testaceous.
On the disc of the elytra two series of larger, setiferous punctures can be distinguished,
the outer less complete and more irregular than the inner. ‘The anterior margin of
the pree-frons 1s widely, very shallowly, and subangularly, emarginate.
* Zaitzev proposes the name Lwmetus for this subgenus in his Catalogue, l.c.
7 The name abnormalis is changed into abnormicollis in Zaitzev’s Catalogue: this must be by error, as no
reason is given, and abnormalis is not preoccupied in the genus. Zaitzev’s reference to Sharp’s description is
otherwise correct.
SECOND SERIES—ZOOLOGY, VOL. XVI. 27
206 PERCY SLADEN TRUST EXPEDITION
Two structural points appear to have escaped notice hitherto :
(1) The mentum, which is transverse, is deeply excavated in the middle in front:
the front margin runs obliquely forwards on either side of the excavation: most of
the surface (including the excavated part) is strongly rugose, near the posterior angles
it is punctate.
(II) The /ast ventral segment is almost truncate or perceptibly sinuate behind, and
in the middle the hind margin is set with a series of minute setze rather like the teeth of
a comb. They are pale in colour, and quite different from the ordinary pubescence of
the segment, bemmg much stouter than the ordinary hairs: the sete in the middle
of the series are shorter than those on either side. Insufficiency of material has
prevented my determining whether this structure is confined to one sex, or present
and of the same form in both sexes; in Philydrus parvulus sete of a somewhat
similar kind are present in both sexes: see p. 204.
Loc. Aldabra: Takamaka, 1908, 3 specimens (Fryer). Seychelles: Silhouette ;
from a small pool on the plateau, Mare aux Cochons, over 1000 feet, 25. IX. 1908,
1 specimen. Madagascar, British East Africa, Ceylon, [Sumatra ? |.
Paromicrus, gen. nov. (Plate 14, figs. 1—11).
Antenne 9-articulatee, articulis 7—9 clavam compactam formantibus. Labrum haud
occultum, transversum. Palpi maxillares articulo 2° sat incrassato, 3° quam 2° breviore,
4° quam 3° longiore. Mentum trapezoidale, transversum. Elytra seriebus regularibus
punctorum munita. Ale perfecte. Prosternum breve, antice in medio elevatum sed
haud carinatum. Mesosternum in medio elevatum; parte elevata haud carinata sed
aream marginatam, antice in apicem ventraliter directum productam, formante. Meta-
sternum in medio leviter elevatum, antice inter coxas intermedias productum, haud
carinatum, cum mesosterno sutura transversd conjunctum. Segmentum abdominis basale
in medio longitudinaliter carinatum. ‘Tarsi omnes 5-articulati, articulis 1—4 brevibus,
subzequalibus, articulo 5 longiore.
Genus Omicro affine, sed valde differt in forma mesosterni; metasterno in medio
minus elevato, articulo 2° palporum maxillarium minus incrassato.
Form oblong-oval (in Paromicrus atomus slightly narrowed behind), moderately
convex, the upper surface bearing an extremely fine and scanty pubescence which is not
easily visible without a compound microscope. Head short and broad, depressed in front
of the eyes, which project subangularly, truncate in front. Antenne (Pl. 14, fig. 2)
9-jointed in both sexes, the Ist joint elongate, 2nd short and broad, 3rd narrow, joints
4—6 transverse and very short, 7—9 forming a compact club. Labrum (Pl. 14, figs. 3, 4)
exposed, very short and transverse ; its front margin either shallowly smuate (P. carinatus
and P. thomasseti) or angularly emarginate (P. atomus) on either side, and bearing a
group of ciliz in the middle, and one on either side outside the sinuation. Mandibles
(Pl. 14, fig. 5) rather delicate, without apical or other teeth. Mamille (Pl. 14, fig. 6)
with the lobes membranous, the upper one with several regular rows of setze hooked at
their extremities, the lower one with a minute ciliate process on its upper margin near
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERID At 207
the outer angle, and with the outer margin shortly ciliate above: maxillary palpi with
the 2nd joint incrassate distally, 3rd joint shorter than 2nd, 4th joint longer than 3rd.
Labium (PI. 14, fig. 7) with the mentum trapezoidal, transverse and slightly impressed in
the middle in front; lobes of the ligula projecting and densely ciliate ; palpi short and
minute, densely ciliate, the basal joint very short, 3rd joint narrower and a little shorter
than the 2nd.
Elytra with regular series of punctures, without strize except in P. atomus, in which
a very fine sutural stria is present. Wings present and fully developed in all the three
species, over 14 times as long as the elytron. Prosternum short, elevated in the middle
in front, the elevated portion sloping somewhat on either side, but not carinate: behind
the coxee it does not end in a free process. Mesosternum (PI. 14, figs. 9—11) elevated
in the middle ; the elevated portion does not form a narrow keeled lamina, but a margined
area varying in form and shape in the different species, but always ending anteriorly in a
ventrally-projecting point (see Pl. 14, fig. 9, a@ and b): behind, the mesosternum is divided
from the metasternum between the coxze by a transverse suture. Metasternwm elevated
in the middle, not sharply or suddenly, projecting forward between the cox but not
keeled. Abdomen composed of 5 ventral segments, the basal one with a median longi-
tudinal carina. Tursi (Pl. 14, fig. 8) 5-jointed, joints 1—4 short and subequal in length,
joint 5 considerably longer; the short basal joint is not always easily visible.
Type of the genus: Paromicrus carinatus, sp. nov.
Paromicrus is akin to Omicrus Sharp (Trans. Ent. Soc. London, 1879, pp. 81, 82),
from the Hawaiian Islands. I have a specimen of the type of that genus, Omicrus
brevipes Sharp (l.c.) before me. The great difference between it and Paromicrus lies in
the form of the mesosternum ; in Omicrus this forms in the middle a sharply-elevated,
narrow, linear lamina, continuous with the elevated and forward-projecting part of the
metasternum, whereas in Paromicrus it forms a broad margined area differing in form in
the different species, and produced into a ventrally-directed point in front. The middle
part of the metasternum is much more sharply and strongly elevated in Omucrus. The
maxillary palps of Omicrus are formed (as stated by Dr Sharp, /.c.) much as in Mega-
sternum : their 2nd joint is much more incrassate than in Paromicrus, and the terminal
joint appears narrower and more tapering towards the apex. Moreover, Onucrus brevipes
is completely different in appearance from any of the species of Paromicrus: its shape is
not the same, its elytra are not explanate at the sides and have not the series of
punctures, &c.
Three species of this genus were found: none of them is confined to one island, but
I found each of them in Mahé, Silhouette and Praslin, and P. atomus also in Félicité.
P. carinatus and P. thomasseti were obtained only in the high endemic-forest districts,
but P. atomus was also found in some places in the lower country. They are found either
under the bark of decaying wood, or (in the case of P. carinatus and P. thomasseti)
between the leaf-bases of growing precinctive palms*. They are sometimes very
abundant, and on some occasions I obtained a large number of specimens from a single
* See vol. xiv. (1910) of these Transactions, p. 24; and cf. vol. xv. (1912) pp. 225, 226.
27—2
208 PERCY SLADEN TRUST EXPEDITION
decaying log or a single growing palm-tree. Two other species of the genus are known,
from Engano (an island near Sumatra) and from the Abor country (see p. 195). The
three Seychelles species may be distinguished as follows :—
Larger : each elytron with three raised longitudinal caring: sides of thorax
3): 0) EN (CRU Meno dic conn Gods adoriaucnGeea id eee aA Reirson 1. P. carimatus.
Larger: elytra without carine, but with intervals between series of
punctures convex ; sides of thorax explanate in front......... 2. BP. thomasseti.
Smaller: elytra without carinze and with interstices flat; sides of thorax _
NOt yeRplan ater... eee Mea eee ERE RE Sey ol Si'blveinn acta vnanion 3. P. atomus.
10. Paromicrus carmatus, sp. nov. (Plate 14, figs. 1—3, 5—9).
Oblongo-ovalis, prothoracis elytrorumque marginibus sat valde explanatis, subopacus,
fusco-ferrugineus vel fuscus, marginibus pedibusque dilutioribus, palpis antennisque
testaceis, his clav& interdum infuscaté; prothorace minutissime reticulato, disco postice
utrinque parum elevato-tumido; elytris suturé atque carinis tribus in disco obtusis
elevatis, sat grosse haud profunde seriatim punctatis, seriebus duabus in intervallo
utroque inter carinas, prope medium basis elevatione tumid& obtusi’; mesosterni parte
elevatd aream fere oblongam, superficie parum concava, formante. Long. corp. ca. 14 mm.
Form oblong-oval, sometimes very slightly narrowed behind, with the sides of the
thorax and elytra explanate. Colour more or less dark brownish ferruginous with margins
and legs lighter, antennze and palpi testaceous. Subopaque, this being due to extremely
fine reticulation or rugulosity of the surface; with a compound microscope the head is
seen to be closely covered with a sculpture of extremely fine transverse lines, and the
prothorax with a reticulation consisting of excessively fine punctures and scratches, while
the surface of the elytra is very minutely rugulose-reticulate : a few fine hairs can also be
seen on the thorax and elytra. The disc of the thoraw is slightly elevated or tumid on
either side behind, which causes the sloping sides of the disc to appear vaguely impressed
on either side in front of this elevated portion. The elytra are without any sutural stria ;
they have regular series of rather large but very shallow punctures; the space between
the first series and the suture is strongly elevated, and the 3rd, 5th, and 7th interstices
are also elevated into blunt but perfectly well-marked carinze running the length of the
elytron nearly to its posterior extremity : thus the dorsal part of the elytron has blunt
longitudinal carime, and two series of punctures in each interval between them; the
intermediate interstices are only very slightly convex; in the sloping lateral part, the
alternate interstices are not elevated into carinze; both the carinze and the other
interstices bear lines of very minute punctures; at the base of each elytron, a little
outside the middle and in the region of the outermost carina, is an obtuse but quite
distinct boss, or elevation. The elevated part of the mesosternum (Pl. 14, fig. 9) forms
an area longer than broad, with sides nearly parallel and strongly margined, its surface
slightly concave and with only a minute trace of a median elevation behind. The under-
surface of the body is beautifully reticulated, and the femora, the elevated central part of
the metasternum, and the abdominal segments, bear very sparse short hairs.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERID Ai 209
The series consists of nearly 300 specimens ; some of the smallest are only 1 mm. long;
some immature specimens are yellow.
This. species is at once distinguished by the remarkable sculpture of the elytra and
their basal boss, by the conspicuously explanate side-margins of thorax and elytra,
and the form of the mesosternum. The plan of the elytral sculpture—alternate interstices
being raised into carinee—recalls that of Oxyomus palmaruwm (see vol. xv. of these
Transactions, p. 224, and Pl. 12, fig. 1).
Loc. Seychelles. Silhouette: high forest near Mont Pot-a-eau, circa 1500 feet,
VIII. 1908; many specimens, including one recorded as found between leaf-bases of a
Roscheria palm, and one as found in a decaying fallen ‘‘ Bois Rouge” (Wormia). Mahé:
high forest of Morne Blane and Morne Pilot, 1000—2000 feet (some specimens found
between leaf-bases of a growing Stevensonia palm) ; high forest between Trois Fréres and
Morne Seychellois, and slopes of Morne Seychellois, 1500—2000 feet ; Mare aux Cochons
district, 1500—2000 feet; forest above Cascade Estate, 800—2000 feet. Praslin:
Coco-de-Mer forest in Vallée de Mai, Cotes d’Or Estate, XI. 1908.
11. Paronicrus thomasseti, sp. nov. (Plate 14, fig. 10).
Oblongo-ovalis, nitidus, fusco-ferrugineus, marginibus suturdque pedibusque diluti-
oribus; capite prothoraceque dense minutissime punctulatis, hoc lateribus solum ad
angulos anticos explanatis; elytris lateribus minus late explanatis, haud carinatis, sat
fortiter seriatim punctatis, intervallo quoque parum convexo et linea irregulari punctu-
lorum minutorum munito, elevatione basali obsoleta; mesosterni parte elevaté aream
latiorem, polygonam vel fere circularem, superficie in medio postice longitudinaliter
carinato-elevata, formante. Long. corp. ca. 14 mm.
Head very finely and closely punctulate, with a rather vague transverse impression,
more or less developed, on either side between the eyes. Thorax with the lateral margins
explanate only in front, and even there not broadly explanate, the disc very closely and
finely punctulate, with a slight impression on either side of the middle at the base.
Elytra with the side-margins explanate, more broadly behind, and a little less con-
spicuously so than in P. carinatus; with regular series of fairly strong punctures, the
interstices being slightly convex and bearing each an irregular line of minute punctures :
carinze are quite absent, the alternate interstices being not more elevated than the rest,
and a basal boss corresponding to that in P. carinatus is either absent altogether, or only
a very slight trace of an elevation is visible. The elevated part of the mesosternwm
(Pl. 14, fig. 10) is very differently formed from that in P. carinatus; it is broader in
proportion to its length, polygonal, with the sides convergent behind, and with a sharply-
elevated median longitudinal carnation on the posterior part of its surface. The scanty
hairs on the under-surface appear a little longer than in P. carinatus, and the inter-
mediate and posterior tarsi are shorter and stouter than in that species.
Distinguished by its shiny surface, by the sides of the thorax being explanate only
in front, by the absence of carinze on the elytra, and by the different form of the
mesosternum.
210 PERCY SLADEN TRUST EXPEDITION
The series contains about 170 specimens, the majority of which were found between
the leaf-bases of various wild palms.
This species is dedicated to Mr H. P. Thomasset.
Loc. Seychelles. Silhouette: high forest near Mont Pot-a-eau, cirea 1500 feet,
a number of the specimens having been found between the leaf-bases of two growing palms
(a Stevensonia and another, genus not recorded) on the 2nd and 7th of August 1908 ;
forest above Mare aux Cochons, including 26 specimens from leaf-bases of one growing
Stevensonia and six from those of another, both on Sept. 22nd. Mahé: forest above
Cascade Estate, including over 40 specimens from leaf-bases of a single growing
Stevensonia; summit of Mount Sebert, over 2000 feet, 11 specimens from leaf-bases
of a growing Stevensonia; from near Morne Blanc, some from leaf-bases of a growing
Verschaffeliia palm, others from those of a Stevensonia. Praslin : Coco-de-Mer forest in
the Vallée de Mai, Cotes d’Or Estate, 11 of the specimens being from between the leaf-
bases of a growing $ Lodoicea palm (‘‘ Coco-de-Mer ”).
12, Paronucrus atomus, sp. nov. (Plate 14, figs. 4 and 11).
Minutus, oblongo-ovalis, postice parum angustatus, nitidus, ferrugineus plus minusve
infuscatus; capite prothoraceque dense minutissime punctulatis, hoe lateribus haud
explanatis ; elytris strié suturali tenui, antice obsoleta, munitis, lateribus postice anguste
explanatis, fortiter seriatim punctatis, seriebus parum impressis, intervallis fere planis,
intervallo quoque lined punctulorum minutorum munito ; mesosterni parte elevata aream
rhomboidalem, elevatione transversé tenui instructam, formante. Long. corp. ca. 1 mm.
Head and prothorax appearing smooth, but under a compound microscope seen
to be closely covered with an exceedingly fine, semi-reticulate punctuation. Prothorax
practically not explanate at all at the sides, only under a high power is a very shght trace
of flattening of the margin visible in front. Hlytra with the side-margins only very
narrowly explanate, with a very fine sutural stria, often difficult to see and becoming
obsolete in front, with regular series of large strong punctures, the series slightly
impressed, the interstices almost flat and each bearing a series of minute punctures.
Elevated part of the mesosternum (Pl. 14, fig. 11) forming a roughly diamond-shaped
four-sided area, which bears a fine transverse elevated ridge, and behind this (towards
its posterior extremity) a median elevation. The elevation of the median part of the
metasternum is more marked than in the preceding species.
The labrum (Pl. 14, fig. 4) is differently formed from that of P. carinatus or
P. thomasseti, the sinuations of its anterior margin being deeper and more angular.
The distinguishing features are the smaller size, shiny surface, the side-margins of
the thorax not being explanate and those of the elytra only very slightly so, the inter-
stices on the elytra being nearly flat, and the form of the mesosternum.
The series consists of over 100 specimens. It is not confined to the high mountain-
forests, as some specimens were found in lower country. I have no record of finding any
specimen between the leaf-bases of a palm.
Loc. Seychelles. Silhouette: high forest near Mont Pot-a-eau, circa 1500 feet;
forest near Mare aux Cochons plateau, many specimens, including some recorded as found
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 211
under bark of a partly-decayed Dracena stem; low coconut-planted country near the
coast at Pointe Etienne. Mahé: forest of Morne Blanc ; high forest between Trois Fréres
and Morne Seychellois, 1500—2000 feet ; forest above Cascade Estate ; Mare aux Cochons
district, 1500—2000 feet; Long Island. Praslin: Coco-de-Mer forest in Vallée de Mai,
Cotes d’Or Estate. Félicité: from forest.
Spheridiini.
DAcTYLOSTERNUM, Wollaston.
13. Dactylosternum insulare (Castelnau).
Dactylosternum insulare Castelnau, Hist. Nat. Col., u. 1840, p. 59.
Dactylosternum imsulare Alluaud, Liste Coléopt., p. 233; Régimbart, Ann. Soe.
ent. France, Ixxu. 1903, p. 46; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 24.
A series of about 150 specimens of this subcosmopolitan species.
Loc. Seychelles. Silhouette: from near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908 ;
Mare aux Cochons and forest above, 1908. Mahé: from near Morne Blane, ca. 500—
1000 feet, 1908; high forest between Trois Freres and Morne Seychellois, 1500—
2000 feet, 1908; Cascade Estate, ca. 1000 feet, 1908—9; Long Island, VII. 1908;
1906 (Thomasset); 1905; 1896 (Brauer). Praslin: Cotes d’Or Estate, 1908; 1905.
Also found in the Seychelles by Alluaud (1892). Very common in rotting fruit, especially
oranges and “jak,” where it is often found together with Cercyon uniformis, Cercyon
fructicola, certain Nitidulide (especially Haptoncus ocularis Fairm. and Brachypeplus
equalis Walker), and certain Staphylinide. As can be seen from the above list, it
is found in many localities, from cultivated country near sea-level to the endemic
mountain-forests.
Aldabra: Picard Island (Fryer).
Described from Mauritius ; distributed over all the warmer regions of the Old World.
14. Dactylosternum pygmeum, Régimbart.
Dactylosternum pygmeum Régimbart, Ann. Soc. ent. France, Ixxii. 1903, p. 45.
16 specimens. This determination was first made for me by Monsieur d’Orchymont,
and has since been confirmed by Monsieur Lesne, who found no difference between the
specimen submitted to him and Régimbart’s type. The specimens agree with Régimbart’s
description: he lays stress on the prothorax being perceptibly wider at the base than the
elytra, but this is not a very perceptible character ; in contracted specimens the posterior
angles of the thorax slightly overlap the shoulders of the elytra, and in extended speci-
mens the thorax does sometimes appear a trifle wider at the base.
This species is entirely distinct from even the smallest specimens of Dactylosternum
insulare not only by its much shorter and more convex form, but by a number of other
characters also. The punctuation is very much finer and much less dense, and quite
different in nature; in D. insulare it consists of fairly strong simple punctures, but in
pygmeum each puncture is very fine and has two minute fine scratches or striole radiating
212 PERCY SLADEN TRUST EXPEDITION
from it, usually at an obtuse angle to one another ; on the head the punctuation is closer,
and the striolze almost form a network, but on the thorax and elytra (especially) they are
much more separated; the striole le in various directions, but usually more or less
transverse. In D. pygmeum the front margin of the eye is only so shallowly emarginate
that the emargination is hard to discern, not deeply and sharply emarginate as in
D. insulare. In D. pygmeum also the club of the antennz is less compact, and joints
2—6 are proportionately longer, the apex of the scutellum is more bluntly rounded, and
the first jomt of the hind tarsus is much shorter in proportion to the second ; the series of
punctures on the elytra are less deeply canaliculate behind, and their arrangement differs
in various details the 2nd—5th interstices being proportionately rather wide, &c.
I have also had for comparison a specimen from the Hawaiian Islands of the species
determined by Dr Sharp as D. subquadratum Fairm.* It closely resembles D. pygmeum,
having the same short and convex form, and the same kind of punctuation consisting of
very fine punctures, each with two radiating short striole. It is, however, much larger
and the punctuation is very much closer; also its elytra do not narrow slightly from the
shoulder to the point where the margin curves round to the apex, as they do in
D. pygmeum. It agrees with pygmeum in having the eye not emarginate in front, the
antenna somewhat less compact than that of D. imsulare, and the apex of the scutellum
blunt: in the undersides I have been unable to detect any difference; in both the posterior
femora are very finely and sparsely, the middle femora much more strongly and closely
punctate. The two forms are at any rate very closely allied.
Loc. Seychelles. Mahé: near Morne Blanc, XI. 1908, 2 specimens ; Cascade Estate,
800—1000 feet, 1908—9, 14 specimens. Mauritius T.
Ca@tostoma, Brullé.
Cyclonotum, Exichson.
15. Calostoma rufitarse (Boheman).
Cyclonotum rufitarse Boheman, Ins. Caffrar., i. 1848, pode
Celostoma rufitarse Régimbart, Ann. Soc. ent. France, Ixxu. 1903, p. 45.
Cyclonotum rufitarse Régimbart, op. cit., xxv. 1906, p. 269.
A series of 31 specimens from Aldabra is referred to this species. For comparison
I have had a specimen determined as C. rufitarse Boh. from Madagascar (ex coll.
Régimbart, Paris Museum), and two determined as C. punctulatum Klug, from Mada-
gascar (ex coll. Sharp, British Museum). The Aldabra specimens agree closely with
Régimbart’s rufitarse: both they and it are decidedly smaller and considerably less broad
in proportion than the specimens of punctulatwm, which is in accordance with Régimbart’s
* Sharp, Fauna Hawaiiensis, vol. iii. part 5, 1908, p. 578, and Trans. R. Dublin Soc., ser. 2, vol. iii.
1885, p. 218. D. subquadratwm was described by Fairmaire as a Cyclonotwm (Rev. Mag. Zool. (2), i. 1849,
p- 412) and is included by Zaitzev in the genus Celostoma (= Cyclonotum) (Cat. Hydrophil. 1908, p. 404). It
was described from Tahiti.
+ Zaitzev in his Catalogue, 1908, p. 402, gives ‘‘ Madagascar” as the only record of distribution for this
species. Régimbart after his description gives only “ Tle Maurice: Curepipe (Carié).” I am not aware that it
has ever been recorded from Madagascar, or anywhere but Mauritius.
SCOTT—COLEOPTERA ; HYDROPHILIDA, HISTERID A 213
statement of the difference between the two species (Ann. Soc. ent. France, xxii. 1903,
p. 45). Otherwise I have failed to detect any appreciable difference in either upper- or
under-sides. In both the punctuation above is rather fine, very close, and nearly even on
head, thorax and elytra; the posterior femora are very finely and sparsely punctate, while
the intermediate femora have numerous moderately strong punctures ; and the palpi are of
the same reddish-yellow colour in both. The Aldabra specimens are mostly 5 mm. long,
or with head extended 6 mm. : a few are smaller”.
A single specimen from the Seychelles (Mahé ; low country, 1908) is a little shorter
and perceptibly broader in proportion than the Aldabra specimens, and a trifle more
strongly punctured, especially on the elytra. I am uncertain to which species this should
be referred [see footnote ; C. punctulatum Klug has been recorded from the Seychelles ;
see below |.
Loc. Aldabra: Takamaka, X.—XI. 1908 (Fryer) ; Picard Island, 1907. Madagascar ;
East and South Africa.
16. Celostoma punctulatum (Klug).
Spheridium punctulatum Klug, Ins. Madag., 1833, p. 161 (73).
Celostoma punctulatum Alluaud, Liste Coléopt., p. 232; Régimbart, Ann. Soe. ent.
France, lxxu. 1903, p. 45; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 24.
Cyclonotum punctulatum Régimbart, Ann. Soc. ent. France, Ixxv. 1906, p. 269.
This species is recorded from the Seychelles by Kolbe (/.c.). I only obtained one
specimen of the genus Celostoma in the Seychelles, and am uncertain to which species it
should be referred : it is mentioned above, under C. rufitarse.
Loc. Seychelles: Mahé (Brauer). Madagascar; East Africa, West Africa (Gabon,
&e., teste Régimbart, /.c.).
BouRDONNAISIA, gen. nov. (Plate 14, figs. 12—21).
Corpus ovale, haud fortiter convexum, supra nitidissimum et glabrum. Oculi antice
haud emarginati. Antennee in ? 9-, in ¢ 8- vel 9-articulatze, articulis tribus ultimis
* Tt is impossible not to feel some doubt as to the identity of these specimens. The types of rufitarse and
punctulatum have not been accessible, and it is not absolutely certain that the specimens used for comparison
really belong to those species, though they probably do so. Moreover, beyond the difference in size and general
form, there do not appear to be real differentiating characters between the two species. I hoped the form of
the scutellum might possibly prove a differentiating character, since in a specimen from Brit. Hast Africa
(ex coll. Régimbart, Paris Mus.) the scutellum is much narrower at the base, proportionately longer and
consequently more acute at the apex [this specimen is labelled “punctwlatum Klug ?,” but its identity with that
species is very doubtful, owing to the relatively narrow form of the insect as a whole and of its scutellum;
Régimbart admitted the possibility of the presence of a third species among some rufitarse from Hast Africa,
see Ann. Soe. ent. France, Ixxv. 1906, p. 269]: but Régimbart’s specimen of rufitarse, the Aldabra specimens,
and one at least of the specimens of punctwlatum, all agree in having the scutellum broad at the base and
relatively short. Neither does the form of the head appear to differentiate the two, though in specimens before
me of a third species (C'. simplex Sharp, Ceylon) its relative narrowness is at once apparent. The characters
Separating species of this genus seem in several cases very unsatisfactory, and probably a thorough revision,
with examination of genitalia, &c., is needed.
SECOND SERIES—ZOOLOGY, VOL. XVI. 28
214 PERCY SLADEN TRUST EXPEDITION
clavam laxam formantibus. Palpi maxillares articulo secundo sat elongato ad apicem
parum incrassato, articulo tertio quam secundo breviore, articulo ultimo quam tertio parum
longiore. Mentum transversum, antice impressum ; palpi labiales breves, articulo secundo
elongato, incrassato, dense ciliato, articulo tertio parvo, quam secundo multo breviore.
Elytra sine stria aliqua. Ale omnino desunt. Prosternum breve, medio antice haud
carinatum. Mesosterni pars elevata haud elongata, antice utrinque marginata, postice cum
processu metasterni perfecte conjuncta, sutura transvers’ vix visibili. Metasternum in
medio elevatum, antice inter coxas intermedias productum. Segmentum basale abdominis
haud carinatum. Tarsi intermedi et postici hirsuti, articulo basali sat incrassato, quam
secundo longiore.
Body oval, not very convex, shining and glabrous above: under-surface subopaque,
sterna and abdominal segments minutely reticulate, the latter with a very fine short
pubescence. Hyes not emarginate in front. Antenne (Pl. 14, fig. 13) 9-jointed in 2,
8- or 9-jointed in ¢ (see below, under B. mahensis), the last three joints forming a loose
club ; basal jomt elongate, about equal in length to those intervening between it and the
club taken together, second joint about as broad as basal joint, slightly pyriform, third
shorter and much more slender, the following two (or one in B. mahensis 2) very short, the
sixth (or fifth in B. mahensis 2) very short and transverse. Labrum (PL. 14, fig. 14) hidden
under the front of the head, short and transverse, rather deeply sinuate in the middle of
the front margin and fringed with hairs on either side. Mandibles (Pl. 14, fig. 15) with
an elongate slender apical tooth, the inner margin below this membranous and fringed
with hairs. Mazxille (PI. 14, fig. 16) with the lobes membranous, the upper one densely
setose along its upper margin, the lower one with two setiferous processes at its upper
angle and a fringe of setze on the margin below these; palpi (Pl. 14, figs. 16, 17) with
second joint elongate and somewhat swollen apically, third joint shorter than second,
fourth joint a little longer than third and blunt at the apex. Mentuwm (Pl. 14, fig. 18)
transverse, impressed and concave in middle in front, with front margin slightly sinuate
in middle; lobes of the ligula densely ciliate, projecting beyond the mentum; labial
palpi short, second joint elongate, densely ciliate and slightly swollen, third joint very
small, only about half as long as second, narrow, blunt at the apex. Hlytra without
a sutural or any other stria. Wings entirely absent: the most careful examination of
a number of specimens of both sexes, in both species, has failed to reveal any trace of these
organs. Prosternwm very short, not carinate in the middle in front; a narrow lamina
extends backwards between the front coxee, and the ventral part of it projects slightly
behind (apparently slightly more in B. mahensis than in B. silhouette), but scarcely
enough to be called a free process. Mesosternum (Pl. 14, figs. 20, 21) forming a rather
short elevated ridge, strongly margined on either side in front, and very closely united to
the forward-projecting portion of the metasternum, the transverse suture between the two
being sometimes scarcely visible: the median portion of the metasternum is elevated and
projects some way forward between the middle coxee to meet the mesosternum. [In
B. silhouette the elevation of the mesosternum forms a sharp ridge, which continues as
a carina without interruption along the metasternum to its posterior extremity (Pl. 14,
fig. 21): in B. mahensis the elevated portions of mesosternum and metasternum are broader
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 215
and much blunter, so that there is no sharp central carina (Pl. 14, fig. 20).] Basal
abdominal segment without a carina. Tarsi (Pl. 14, fig. 19) nearly as long as the tibie,
hirsute, with the basal jomt somewhat incrassate and considerably longer than the second ;
second, third, and fourth joints becoming gradually shorter, fifth joint about as long as
third and fourth together.
Type of the genus: Bourdonnaisia mahensis, sp. nov.
It is difficult to speak of the affinities of this genus. In several points it resembles
Celostoma, but differs from that genus in its less convex form and general outline, in the
absence of a sutural stria and the presence of a seriate arrangement in the punctuation on
the elytra, &c. It appears to be isolated, and is very remarkable in its complete wing-
lessness, and in the sexual dimorphism in the antennee of one of its species: I am unaware
that any other case is known among the Hydrophilidz of the ¢ antenna having a different
number of joints from that of the ?.
Habitat. A long series of the two species was obtained, and the specimens were only
found in extremely circumscribed areas on the very summits of the highest peaks of Mahé
and Silhouette, at elevations of over 2000 feet, in the highest and dampest forest-zone.
These peaks are covered with forest of the endemic “capucin” tree, Northea seychellarum,
which has big smooth leathery leaves. These leaves, when fallen and dead, collect in
hollows on the ground, lying closely on one another and (owing to the extreme humidity)
having films of moisture between them. In this moisture the beetles live: when the
leaves are pulled apart and turned over the little creatures are seen crawling on the wet
and slimy surfaces, their shape causing them to fit closely down on to the leaf, and their
somewhat depressed form being doubtless a convenience in the narrow spaces between the
leaves. They were found nowhere but in this particular habitat, and are further remark-
able in that one species is confined to Mahé and the other to Silhouette* (an arrangement
which, together with the restriction of areas, recalls the distribution of many Hawaiian
Coleoptera).
The name Bourdonnaisia is taken from the Governor of Mauritius, Mahé de la
Bourdonnais ; the largest island of the Seychelles was named “ Mahé” after him in 1744,
and the whole group was for some time known as “Les Iles de la Bourdonnais t.”
17. Bourdonnaisia mahensis, sp. nov. (Plate 14, fig. 12).
é¢. Sat breviter ovalis, corpore supra nigro-piceo, subtus piceo-ferrugineo, palpis
antennisque flavis, harum clavis nigrescentibus, tarsis flavo-ferrugineis; capite levi, ad
marginem anteriorem rarius tenuissime punctulato; prothorace scutelloque levibus ;
elytris marginibus externis postice parum explanatis, sat fortiter, crebre, et regulariter
serlatim punctatis, punctis serierum alternatarum parum fortioribus. Long. corp. ca.
132 mm.
a, antennis 8-articulatis; 2, antennis 9-articulatis.
* The finding of these beetles is referred to in the writer’s introductory paper, Trans. Linn. Soc. London,
ser. 2, Zool., vol. xiv. 1910, pp. 25, 29, 33.
t See J. Stanley Gardiner, in the Geographical Journal for February, 1907, p. 152.
216 PERCY SLADEN TRUST EXPEDITION
Rather shortly and broadly oval, shining pitchy-black above, the base of the thorax
sometimes narrowly testaceous and the suture of the elytra sometimes vaguely ferruginous;
under-side dark pitchy ferruginous ; palps and antennee yellow, clubs of the latter blackish.
Submentum bearing a transverse series of long hairs (absent in B. silhouette), standing in
a shehtly curved line. Head with very fine scanty punctuation near the anterior margin ;
rest of head and prothorax quite smooth and impunctate (only under a compound micro-
scope is an extremely scanty and obsolete punctuation visible). Scutellumsmooth. Llytra
with the outer margins slightly flattened towards the apex, rather strongly and regularly
punctured, the punctures arranged in a number of closely-approximated series ; in each
series the punctures are well spaced out, at some distance from one another, and those in
each alternate series are slightly stronger. The seriate nature of the punctuation is not
always quite evident at first sight, but is very clearly visible if an elytron is removed and
examined separately. Certain peculiarities of the sterna are mentioned in the description
of the genus (see Pl. 14, fig. 20).
Antenne (Pl. 14, fig. 13): in the ¢ the antenna consists of only eight joints, as
opposed to nine in the ¢; it has only two short transverse joints between the slender
elongated third jot and the club, instead of three as in the 2. When balsam-
preparations of the g¢ antenna are viewed as transparencies under a high power
(4-inch objective), there is an indication that the proximal of these three transverse
joints, z.e. that which corresponds to the 4th joint in the ?, may have fused with the
third joint, thereby reducing the number in the $ from 9 to 8. Usually the number of
joints cannot be surely determined without making balsam-preparations, and there is
no other external sexual differentiation: but when several transparent balsam-mounts
were made, the sexes were easily distinguished by the genital structures within the apex
of the abdomen, and it was then seen in each case that the ? antenna has 8 joints,
the 2 9.
The series consists of about 100 specimens.
Loc. Seychelles. Mahé: summit of Morne Pilot, over 2000 feet, X.—XI. 1908;
high forest between Trois Fréres and Morne Seychellois, about 2000 feet, 9 and 10.
XII. 1908; slopes of Morne Seychellois, about 2000 feet, 4. II. 1909; high forest above
Cascade Estate, I. 1909; summit of “ Montagne Anse Major” (Mare aux Cochons district),
about 2000 feet, 1. IT. 1909.
18. Bourdonnaisia silhouette, sp. nov. (Plate 14, figs. 16 and 21).
22. Major, magis elongata, elytris parum subtilius punctatis; mesosterno et meta-
=) Sate, Cry Cesar
sterno in medio argute longitudinaliter carinatis. Long. corp. ca. 24 mm.
¢ et 2, antennis 9-articulatis.
Much larger ; form more elongate-oval as contrasted with the short broadly-oval form
of B. mahensis. Elytra with the margins a trifle less explanate behind, and with the
punctuation slightly finer. Maxillary palpi darker in colour. Thorax sometimes testaceous
on either side at the base. Peculiarities in the form of the sterna (see Pl. 14, fig. 21), as
compared with that of B. mahensis, are mentioned in the description of the genus.
SCOTT—COLEOPTERA ; HYDROPHILIDA, HISTERID At 217
Antenne : these organs are 9-jointed in f and ¢, and I have been unable to detect
any difference in form between those of ¢ and those of 9. Several balsam-preparations
were made, and the sexes distinguished by the genital structures, as in the case of the
preceding species.
The series consists of 32 specimens.
Loc. Seychelles. Silhouette: highest peak, about 2400 feet, IX. 1908.
Crercyon, Leach.
All the four species of this genus found in the Seychelles belong to the subgenus
Cercyon s. str. The ventral surface of the mesosternal lamella is very narrow (in C. fructi-
cola and C. laticollis linear, in C. uniformis very slightly broadened in the middle and
tapering to a point before and behind) and the front of the metasternum is not cut out to
receive the posterior end of this lamella. The front tibize are simple (not emarginate out-
wardly before the apex): the sides of the prothorax are not scalloped before the base: the
convexity of thorax and elytra forms a single curve (except when the specimens are
unnaturally extended).
19. Cercyon fructicola, sp. nov.
Ovalis, depressus, angustus, lateribus subparallelis, nitidus, parcissime tenuissimeque
pubescens ; supra omnino piceo-niger, palpis antennisque rufo-testaceis, harum clavis in-
fuscatis, corpore subtus pedibusque omnino rufo-ferrugineis: capite pronotoque modice
fortiter et crebre punctatis: elytris sat profunde punctato-striatis, intervallis ad basin fere
planis, ad apicem parum convexis, sat tenuiter crebre punctatis. Long. corp. (incl. cap.)
12—23 mm. ,
Form oval, depressed, rather narrow, with sides subparallel: in some specimens the
thorax appears very slightly broader at the base than the elytra: from the base to about
half the length the sides of the elytra are nearly parallel or very slightly curved, beyond
that point they curve inwards to the not very blunt apex. Body above entirely pitchy-
black, shining, only the pronotum and scutellum sometimes a trifle more dilute, and the
front margin of the head and front and hind margins of the pronotum narrowly reddish ;
antennze and palps reddish-testaceous, clubs of the antennze infuscate ; body beneath and
legs entirely ferruginous. Very fine short hairs rise sparsely from some of the punctures
on the head, pronotum, and from the intervallar punctures of the elytra. Head with
front margin subtruncate, very slightly rounded ; moderately, strongly and fairly closely
punctured, intraocular systematic punctures not very distinct. Pronotwm with punctua-
tion about equal to that of the head, with sides gradually narrowing from the base
forwards and slightly curved. Scutellum considerably longer than broad at base, smooth,
with only a few punctures. Hlytra each with nine rather deep punctate striz, the
punctures close together near the base, separated by twice their own diameter or more in
the posterior part, strongest in the outermost strize: striz 1—5 reach to the base, 1 (the
sutural) bending outwards on either side of the scutellum; 6 is abbreviated before the
218 PERCY SLADEN TRUST EXPEDITION
base by the humeral callus; 7 reaches to the base (5 and 7 curving very slightly towards
one another, on either side of the callus, at the base); 8 and 9 are abbreviated a little
before the base; at the apical end, 1—3 run to the apex, 4 unites with 9, 5 with 8,
enclosing 6 and 7: intervals nearly flat, the outermost ones (and the inner ones towards
the apex) slightly convex, with rather numerous fine punctures, and with a not quite
smooth appearance in the general surface. Mesosternum with the raised lamella very
narrow, linear.
This species appears to resemble Cercyon nigerrimus Régimbart (Ann. Soe. ent. France,
Ixxii. 1903, p. 49; known from Mauritius), judging from the description of that species.
But M. Lesne, to whom I submitted a specimen of Cercyon fructicola for comparison,
informs me that the unique type of Cercyon nigerrimus is larger, decidedly shorter and
more convex, with the pronotum much less clearly punctuated, &c.
74 specimens.
Loc. Seychelles. Silhouette: near Mont Pot-d-eau, ca. 1500 feet, VIII. 1908;
Mare aux Cochons plateau and forest above, ca. 1000 feet and over, VIII.—IX. 1908.
Mahé: high forest of Morne Blanc, X. 1908; high damp forest between Trois Fréres and
Morne Seychellois, 1500—2000 feet, XII. 1908 ; Cascade Hstate, ca. 1000 feet, and forest
above, 1908—9; Mare aux Cochons district, 1500—2000 feet, I—II. 1909. Praslin :
Cotes d’Or Estate, XI. 1908.
The above list shows that in Mahé and Silhouette this species was taken principally
if not entirely in the mountain-forests. It was often found in rotting fruit, especially
oranges and “jak,” often in company with Cercyon unifornus, Dactylosternum msulare, &e.
(see under those two species).
20. Cercyon laticollis, Régimbart.
Cercyon laticollis Régimbart, Ann. Soe. ent. France, lxxii. 1903, p. 48.
13 specimens: the determination was first made by Monsieur d’Orchymont, and con-
firmed by Monsieur Lesne, to whom a specimen was submitted. The specimens agree well
with Régimbart’s description. The lateral margins of the prothorax, when viewed from
the side, are seen to be slightly sinuate behind the anterior angles. On the elytra the
Ist (sutural) and 2nd strize do not reach quite to the base, 3, 4 and 5 almost reach the
base, 6 is abbreviated a little before the base, the callus being situated between its front
end and the base, 7 reaches further forward than 6, 8 and 9 end some distance from the
base, but 9 reaches further forward than 8: at the apical end, in at any rate most of the
specimens, stria 4 unites with 9, and 5 with 8, enclosing 6 and 7: the large punctures
of the strize lie in each case on the outer side of the stria : the intervals between the inner
striz are nearly flat, the outer ones convex, the intervallar punctures being extremely fme
and not close and bearing exceedingly fine hairs : the suture is darkened near the apex.
I have distinguished a g and a ¢ by dissecting out the genitalia, but can see no
external difference between them.
Loc. Seychelles. Silhouette: Mare aux Cochons, VIII. 1908. Mahé: Long Island,
VII. 1908; low country on the main island. Praslin: Cotes d’Or Estate, XI. 1908.
Madagascar.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 219
Cercyon laticollis Régimbart, var. ?
A single specimen, apparently immature, differs from the preceding series. The sides
of the elytra are more rounded and their greatest width is behind the base, instead of their
being widest at the base and tapering gradually ; the inner strize are strongly impressed
right up to the base, instead of becoming faint in the region of the scutellum, and the
intervals appear slightly less smooth. The sides of the pronotum are not sinuate behind
the anterior angles. The scutellum is narrower at the base and proportionately more
elongated. At first it seemed probable that this might be the 2 of C. laticollis: but as
stated above dissection has revealed the presence in the series of that species of 3 and 2 not
differing externally, and the true position of this specimen must therefore remain doubtful.
Loc. Seychelles. Silhouette: Mare aux Cochons, IX. 1908.
21. Cercyon uniformis, Sharp.
Cercyon uniformis Sharp, Trans. Ent. Soc. London, 1890, p. 357.
Determined by comparison with the type. A series of over 160 specimens, showing
considerable variation in size, but apparently constant in coloration, &c.
Loc. Seychelles. Mahé: high forest of Mornes Blane and Pilot, ca. 2000 feet,
including some from strong-smelling rotting fruits of Pandanus seychellarum on summit
of Morne Blane, X. 1908; country above Port Glaud, 500—1000 feet ; Cascade Estate,
ea. 1000 feet, and forest above, large numbers, 1908—9; Mare aux Cochons district, 1000
—2000 feet, including some from forest on summit of ‘“‘ Montagne Anse Major,” 1909.
Silhouette: near Mont Pot-ai-eau, ca. 1500 feet, VIII. 1908; Mare aux Cochons, over
1000 feet, 1908. |
I found this species very common in rotting fruit, especially oranges and “jak
(Artocarpus integrifolia), where it is frequently obtained in company with Dactylosternum
msulare, Cercyon fructicola, &c. It can be seen from the above locality-list that it is also
found in the dampest endemic forests and at the highest elevations, which is remarkable,
”
as it is probably an introduced species. It is recorded also from Ceylon and India.
22. Cercyon sp.
Length (not incl. extended head), 14 mm. Oval, with the greatest width a little
before the middle of the elytra, narrowed before and behind with the apex of the
elytra bluntly rounded, convex ; head and pronotum shining, elytra subopaque. Head
black, with the extreme anterior margin of the epistome very narrowly testaceous ;
palps and antennz pale testaceous, the latter with blackish clubs: pronotum pitchy-
blackish, with anterior margin narrowly and lateral margins more broadly testaceous,
and posterior margin obscurely reddish. Elytra with the basal quarter obscurely reddish,
the extreme base and scutellum being darker (blackish, especially in the middle): from
about + to about 2 the length is occupied by a broad black band extending from
about the region of the 8th stria to the suture, and stretching obliquely inwards and
backwards ; in front this black band is vaguely bounded, shading gradually into the
basal reddish portion, but behind it is more sharply bounded; behind it the whole
apical portion of the elytron is pale yellowish testaceous, excepting the interval between
220 PERCY SLADEN TRUST EXPEDITION
suture and ist stria, which is darker; laterally the testaceous colour extends along
the side of the elytron between the black band and the side margin, joiming in front
with the outer extremity of the basal reddish area, but not quite reaching the humeral
angle, which is darker. Legs and feet reddish-testaceous.
Head and pronotum finely and rather closely punctate, the pronotum with anterior
and posterior angles obtuse, the latter very widely so: the fine posterior margin of
the pronotum is seen under a high power to be interrupted by punctures. Sceutellum
minute, rather narrow, acute at apex, with a few punctures. Hlytra with 10 punctate
strie, the punctures larger and nearer together near the base, separated by several
times their own diameter behind; stria 1 (the sutural) and strie 2—5 reach nearly
to the base; stria 6 ends in front further from the base; striz 7—9 end in front
about + the length of the elytron from the base (the 7th being just traceable further
forward by a few punctures); stria 10 reaches about as far forward as 6: there is a
slight callosity at the base just outside the anterior extremity of the 5th, and im front
of that of the 6th, stria. Behind, the 10th (outermost) stria ends at about the middle
of the elytron; 9 and 8 can be traced, running parallel to the outer margin, right
round the apex to the suture, and they thereby prevent strie 1—5 from reaching
the apex, while 6 and 7 are abbreviated some way further forward. The intervals
are scarcely convex near the base, but moderately convex behind; they bear very
fine widely-separated punctures, much smaller and finer than those on the striz.
Under a very high power the subopacity of the general surface of the elytra is seen
to be due to the presence of an excessively fine, minute, and close dotting.
This may well be a new species. I have found nothing agreeing with it in any
of the collections in the British Museum. Monsieur d’Orchymont suggested that it
might possibly be allied closely to crenulatus Régimbart (India, Java, &¢.) or crenato-
striatus Régimbart (Mauritius); but Monsieur Lesne, to whom I submitted it, informs
me that both those species are much larger and differently sculptured. As I have
only a single specimen, and have been unable to see a number of species, I have
thought it best to describe but not to name it.
Loc. Seychelles. Mahé: Anonyme Island, 9. I. 1909, 1 specimen.
PaROOsTERNUM, gen. nov. (Plate 14, fig. 22).
Oosterno peraftine, sed differt in forma sternorum. Pars media prosterni haud
carinata, sed tabulam oblongam marginatam, superficie fere plana (perparum concava)
formans. Mesosternum (ut in Oosterno) aream ovatam elevatam formans. Meta-
sternum in medio valde elevatum, sed impressione longitudinali distincta in medio partis
elevatz: lineis femoralibus ad angulos anteriores continuatis.
Body somewhat elongate-oval, moderately convex, with lateral margins of thorax
and elytra not in the least explanate; above shining, with very scanty fine hairs on
the elytra; beneath, central part of metasternum shining, its sides and the abdominal
segments subopaque and bearing fine short pubescence. Head deflexed in front,
labrum hidden. Antenne 9-jointed: basal joint elongate, longer than joints 2—6
SCOTT— COLEOPTERA; HYDROPHILIDA, HISTERID A 221
taken together, joint 2 short but longer than broad, joint 3 narrower, short but also
longer than its breadth, 4 and 5 very short, 6 very short and transverse, applied to
the base of the club, joints 7—9 forming a very compact obovate club*. Mallary
palpi with second joint considerably swollen distally, third jot very slightly swollen
distally, fourth joint slightly swollen in the middle, and subequal to, or slightly longer
than the third (owing to insufficiency of material it is impossible to make a preparation
of the mouth-parts, hence the relative lengths of joints cannot be exactly determined).
Mentum narrowed in front, scarcely impressed in the middle of the front margin.
Wings examined in one specimen and found fully developed, about twice as long as
elytron. Prosternum not carinate in the middle, but with its median portion forming
an elevated oblong area, finely-margined on either side, with surface shghtly concave
in front, and with hind margin angularly excised (between the anterior coxze) to
receive the apex of the elevated part of the mesosternum (see Pl. 14, fig. 22).
Mesosternum forming an elevated, narrowly ovate, margined area, acuminate in front,
with surface concave. Metasternwm strongly elevated in the middle, with a marked
longitudinal impression in the middle of the elevated portion; in front it is only
slightly produced between the middle coxee, and is depressed, so that the middle of
the front margin is hidden under the posterior extremity of the elevated mesosternal
area: the “femoral lines” + are very clearly marked and uninterrupted, reaching to
the anterior corner of the metasternum: in their front part they are fine raised lines,
while their posterior portion is formed by the edge of the elevated median part of the
metasternum. Basal abdominal segment with a median longitudinal carina. Anterior
tabie with the outer margin bearing about 5 short spines distally, and rounded off
at the apex. Tarsz 5-jointed, scarcely hirsute, with the basal joint distinctly longer
than the second.
Type of the genus: Paroosternum degayanum, sp. nov.
This minute insect is very closely allied generically to Oostermum Sharp (Biol.
Centr.-Am., Col. i. 2, p. 112, Pl. 3, fig. 16), but differs from it in the form of the
sterna. Two species of Ovsternum are known, O. costatwm Sharp (/. ¢.) from Central
America, and O. sorex (Sharp) from Japan (described as Cercyon sorex, Trans. Ent.
Soc. London, 1874, p. 418): I have examined specimens of both these and compared
them closely side by side with Paroosternum. In both genera the middle part of the
prosternum is distinctly differentiated from the lateral parts, but im Oosternwm this
middle part has an elevated central longitudinal keel and sloping sides, while in
Paroosternum it forms an oblong elevated area without a trace of a median keel:
this is the principal difference and is alone sufficient to separate the genera. The
oblong elevated table of Paroosternum recalls the prosternum of some Histeridze: in
the one known species its surface is very slightly concave. The genera are closely
similar in the form of the mesosternum (the surface of the ovoid area being however
* As in the Cercyon spp. which I have examined, a kind of transverse ridge near the apex of the terminal
joint of the club makes it appear as if there were a small fourth joint at the apex.
+ “Schenkellinien”: see Ganglbauer, Die Kafer von Mitteleuropa, iv. 1, p. 273 (1904).
SECOND SERIES—ZOOLOGY, VOL. XVI. 29
222 PERCY SLADEN TRUST EXPEDITION
slightly concave in Paroosternum degayanum but not at all concave in Oosternum
costatum), and in having the front of the metasternum depressed and sunk beneath
the posterior margin of the elevated mesosternal area, instead of united with it as in,
e.g., Oryptopleurum. Paroosternum degayanum has a marked median longitudinal
impression on the metasternum: in the specimens of Oosternum costatwm examined
there is no trace of impression on the metasternum, which is slightly convex; in the
one specimen of Oosternuwm sorexw which I have seen the elevated median part of the
metasternum is slightly concave in the middle, but this concavity is quite different
from the marked longitudinal impression of Paroosternum degayanum. The latter
has its femoral lines continued to the anterior corners of the metasternum, while in
Oosternum costatuwm they become obsolete in front.
23. Puroosternum degayanum, sp. nov. (Plate 14, fig. 22).
Elongato-ovalis, sat convexus, nitidus, fuscus, palpis antennisque testaceis, harum
clavis infuscatis, pedibus fusco-testaceis ; capite prothoraceque distincte sat rare punc-
tatis, hoe serie basali punctorum fortiorum; elytris striis impressis et sat fortiter
punctatis, interstitiis haud elevatis et tenuiter sat rare punctatis. Long. corp. circa 1 mm.
The surface is shining, but a very fine reticulation is visible under a compound
microscope. The pubescence of the upper surface is not easy to see, consisting of
extremely scanty and fine pale hairs arising from the punctures on the interstices of
the elytra (in these specimens there are no hairs on head or thorax, but possibly they
are broken off). The punctures on head and thorax are very distinct but not close:
the thorax has a regular series along its base, a feature which is also present im
Oosternum costatum Sharp. The elytral striz are strongly impressed and strongly
punctate; the interstices finely and not closely punctured, not in the least raised into
costee or carinze, their median parts being almost flat.
Oosternum costatum Sharp is much less shining and more strongly reticulated :
its pubescence is much more conspicuous: its head and thorax are more strongly and
closely punctured: and the alternate interstices of the elytra are raised into costee.
Oosternum sorex (Sharp) is much larger than Paroosternum degayanum and quite
different from it in general appearance.
This species is dedicated to Mr J. A. de Gaye, F.L.S., who during the time that
he lived in the Seychelles was a keen student of the entomology of the islands.
Loc. Seychelles. Mahé: from near Morne Blanc, X.—XI. 1908, 2 specimens.
Histeridee.
16 species are here enumerated as occurring in the islands visited by the Expedition.
The number previously recorded was 8, and of the 8 species now added 5 are new to
science. The 16 species represent 6 genera (in three of which two subgenera are repre-
sented). 13 out of the 16 species are actually represented in the collection made by
the Expedition, while the remaining 3 (Platysoma richtert, Carcinops 14-striata,
Saprinus erichson) were found only by previous collectors.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDAI 228
The following table shows the distribution of the species so far as known:
Before the names of the species : Distribution among the groups of |
* indicates that the species is new to science, islands visited by the Expedition
+ indicates that the species is recorded from | —— Sa ~ External Distribution
these islands for the first time, but not | Seychelles | Cargados Providence
new to science. Group Group | (Farquhar Gr.) |
24. Platysoma tenuimargo ...... -— |
25. Platysoma richteri............ a ngacoome || ial aeaaetee Madagascar
26. Carcinops 14-striata ......... Sey ain csscceeay lei c cmieee Subcosmopolitan (?)
27. Paromalus alluaudi ......... ——
28. *Paromalus gardineri ......... ——
29. *Paromalus calciger............ —
30. Bacanius ambiguus .........
31. Bacanius inopinatus ......... —
32. +Bacanius atomarius ......... aE riences Wh tal yare Hawaiian Islands
33. Acritus (Halacritus)algarum | ——, | ...... | ws... Ceylon ;_ Perim (Red Sea)
34, * Acritus (Afletes) daubani ...
35. *Acritus (A#letes) davidsoni —
36. *Acritus (Ailetes) fryeri...... waaa
SP Saprinus) erichsomierseee een |i =| eect | cesar Madagascar
38. jSaprinus disjunctus ......... | ...... ee ee Madagascar
39. Saprinus apricarius ......... —— | sees — Warmer regions of Old World
It will be seen that no Histerid was collected by the Expedition in the Chagos
or Amirantes groups, nor was any found by Mr Fryer during his 4 months’ stay in
Aldabra or his visits to the neighbouring islands: nor have any species ever been recorded
from these groups.
The table shows that 15 species have been found in the Seychelles, 14 of them
only in that archipelago, and 1 widely-distributed species both in that archipelago and
also in the coral-island of Providence: while the remaining 1 species (Saprimus
disjunctus) was found only in the coral-islands of the Cargados Carajos group.
Of the 15 species found in the Seychelles, 9 are known only from those islands,
and most of these are probably endemic. Of these 9 species, Platysoma tenwimargo
has been stated to be closely allied to a New Guinea species (see p. 224): Paromalus
alluaudi is said to be closely allied to an Eastern species (see p. 225): Paromalus
gardinert (very possibly not endemic) in some ways resembles a Ceylon species.
Nothing can be said about the affinities of Paromalus calciger, the two species of
Bacanius, and the three species belonging to the subgenus diletes of Acritus. As
stated on p. 230, it appears that no member of the whole great genus Acritus is
known from Africa, and only two species from Madagascar; and until such minute
creatures as Bacanius and Acritus are more collected and known, it is almost idle
to speculate about the Seychelles species.
Of the 6 species found in the Seychelles but known also from elsewhere, two
(Platysoma richteri, Saprinus erichsoni) are Madagascar species: one (Bacanius ato-
marius) is known from the Hawaiian Islands: one (Acritus algarum, a coast species)
from Ceylon and Perim Island: while two are very wide-spread (Carcinops 14-striata
and Saprinus apricarvus, the latter of which was also found in Providence).
The one species, Saprinus disjunctus, found in the Cargados group is a Madagascar
species.
29—2
224 PERCY SLADEN TRUST EXPEDITION
Local distribution within the Seychelles group. As far as my own experience
goes, 8 out of the 9 species known only from these islands were found in the endemic
mountain-forests, but several of these were found also in lower, cultivated places: it
is noteworthy that one of them (Bacanius ambiguus), which I found under the bark
of decaying wood in several islands, both in the forests and in cultivated places, was
originally found by Alluaud under seaweed on a sandy beach, in company with the
seaweed-species Acritus algarum. The remaining peculiar species (Paromalus gar-
dinert) was only found on a cultivated islet (Long Island). On the other hand, of
the non-peculiar species, Acritus algarum is a coast species inhabiting seaweed, and
the wide-spread Saprinus apricarivus was obtained in an outlying coral-island (Bird
Island) which possesses none of the endemic vegetation or other special characteristics
of the Seychelles. The species known from the Hawaiian Islands was found on Cascade
Estate, but only two specimens were collected, and I have no record of the exact spot,
whether forest or otherwise. The other 3 non-peculiar species were not collected by
this Expedition. The material is in some cases too small to judge of the local
distribution: but I think there is some indication of a division between endemic
forms inhabiting the endemic mountain-forests, and non-endemic forms inhabiting
cultivated places and the coasts (cf. vol. xv. of these Transactions, 1912, bottom of
Os SIL} eyevel jo, Mil'9)))
Reduction of wings. A remarkable case of reduction of these organs to minute
vestiges has been discovered in Acritus (Halacritus) algarum: details are given on
p- 230. As mentioned there, reduced wings had been found in two Hawaiian species
of Acritus, but species belonging to a different subgenus, iletes: lack of material
prevented my examining the wings in the di/etes from the Seychelles. In Bacanis
ambiguus and imopinatus I found the wings fully developed. They were not examined
in any of the other genera or species.
Puatrysoma, Leach.
Subgenus PLATYSOMA, s. str.
24. Platysoma tenuimargo, Schmidt.
Platysoma tenuimargo Schmidt, Bull. Soc. ent. France, 1893, p. 100; Alluaud, Liste
Coléopt., p. 108 ; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 21.
I am indebted to Mr George Lewis for the determination of this species. It is
stated by Schmidt to be closely allied to P. conditum Marseul, from New Guinea. The
collection contains 12 specimens.
Loc. Seychelles. Silhouette: forest near Mont Pot-d-eau, ca. 1500 feet, VIII.
1908; forest above Mare aux Cochons, over 1000 feet, IX. 1908 (one specimen recorded
as found in the decayed head of a felled Verschaffeltia palm); low coconut-planted
country near coast at Pointe Etienne, IX. 1908. Mahé: Cascade Estate, over 800 feet.
La Digue: 1892 (Alluaud).
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 225
Subgenus Cyxisrosoma, Lewis, Ann. Mag. Nat. Hist., ser. 7, xv. 1905, p. 302.
25. Platysoma (Cylistosoma) richteri, Schmidt.
Platysoma richtert Schmidt, Ent. Nachr., xv. 1889, p. 335; id., Bull. Soc. ent.
France, 1893, p. 99; Alluaud, Liste Coléopt., p. 108 ; Kolbe, Mitt. Zool. Mus. Berlin,
Valo UO} p. 21:
Not obtained by the Percy Sladen Trust Expedition.
Loc. Seychelles: La Digue, 1892 (Alluaud). Madagascar.
Carcinoes, Marseul.
26. Carcinops quattuordecimstriata (Stephens).
Carcinops quattuordecimstriata (Stephens) ; Lewis, Syst. Cat. Histeride, p. 38,
1905; Bickhardt, Coleopt. Cat., Histeridze, p. 60, 1910.
Carcinops pumalio (Erichson) ; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 21.
Not obtained by the Percy Sladen Trust Expedition. The Catalogues of Lewis
and Bickhardt, referred to above, are followed in placing punuilio Er. as a synonym
of quattuordecimstriata Steph. Lewis gives as the distribution ‘‘ Cosmopolitan” ; while
Bickhardt gives South and West Europe, and North America.
Loc. Seychelles. Mahé: cultivated country (Brauer).
Paromauus, Erichson.
Subgenus Paromatus, Lewis, Ann. Mag. Nat. Hist., ser. 7, xix. 1907, p. 316.
All three of the species found in the Seychelles belong to this subgenus: that
is to say, they have no sutural stria on the elytra, and the keel of the prosternum
has definite strize on either side and is not narrowed in front of the coxz (see Lewis,
Opmcits ps sls, tie) 4):
27. Paromalus alluaudi, Schmidt.
Paromalus alluaudi Schmidt, Bull. Soc. ent. France, 1893, p. 100; Alluaud, Liste
Coléopt., p. 109; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 21.
1 ¢ and 1 9, determined by comparison with Schmidt’s actual type ¢ and ? and
with another typical pair, all sent by Monsieur Alluaud. I have closely compared
upper and under surfaces of both sexes side by side; the ? obtained by me has the
prosternal strize a little wider apart, but in all other respects, including the remarkable
sculpture, it agrees with the specimens received from Alluaud: the ¢ agrees closely.
Schmidt states that this species has the facies of certain Indian species, and is
very closely allied to Paromalus mendicus Lewis from Japan and Java.
Loc. Seychelles. Silhouette: forest above Mare aux Cochons, well over 1000 feet,
IX. 1908, 1 g and 19. La Digue, in numbers under bark, 1892 (Alluaud).
226 PERCY SLADEN TRUST EXPEDITION
28. Paromalus gardineri, sp. noy. (Plate 14, fig. 23).
$2. Sat latus, ovalis, niger, nitidus, antennis pedibusque rufo-piceis; capite dense
subtiliter punctato, strié marginali integré; prothorace dense subtiliter punctato, striad
marginali anteriore integra; elytris fortiter dense punctatis, basi extus parum impresso
fortiusque punctato, strié dorsali externa und sat obsolet’, ad medium abbreviata, sub-
humerali tenui, vix visibili, suturali null&; propygidio dense sat fortiter punctato ;
pygidio subtilissime punctulato, ad basin fortius punctato; prosterno inter strias angusto,
basi parum latiore, striis subparallelis, inter coxas leviter sinuatis; mesosterno strid
marginali antice late interrupta, strid transversd arcuata impressd, ad latera abbreviata ;
meso- et metasterno et segmento primo abdominis sparse subtilissime sat obsolete
punctulatis. Long. corp. ca. 2 mm.
Form oval, not oblong, rather broad, slightly convex ; head and prothoraw densely
and very finely punctured, the former with marginal stria complete, the latter with the
stria along the front margin fine but complete. Hlytra densely and strongly punctured,
punctures finer towards the inner basal angle; the base outside the middle with a
very slight transverse impression (more marked in some specimens than in others) and
with conspicuously strong punctures, which continue also to some extent along the basal
part of the dorsal stria; with one dorsal stria, outside the middle, rather broad at the
base, somewhat obsolete and difficult to see from certain aspects, terminating at about the
middle of the length of the elytron; there is an extremely fine subhumeral stria, only
visible with ditficulty ; the suture is slightly elevated behind. Propygidiwm and
pygidium devoid of peculiar sculpture in either sex, and the same in both sexes, the
former fairly densely and strongly punctate, the latter with fairly strong punctures
just along the base and with the rest of the surface to the apex extremely finely
punctulate. Prosternum with the anterior lobe very finely punctulate, the portion
between the striz narrow, very slightly broader at the base, the striz subparallel,
very slightly sinuate between the coxe. Mesosternum with the marginal stria widely
interrupted in front, just turning the angle but not reaching nearly to the emargination
in the middle of the front margin; transverse stria impressed, in the form of an arch
with the convex side directed forwards, not reaching to the sides. Meso- and
metasternum and 1st abdominal segment very finely, not closely, and rather obsoletely
punctulate, the punctuation becoming a little stronger at the posterior and outer parts
of the Ist abdominal segment. Anterior tibie with four teeth externally (PI. 14,
fig. 23). I have found no means of distinguishing the sexes without dissection of the
genitalia.
Specimens were submitted to Mr George Lewis, who failed to identify this species
with any in his collection, and considers that it is almost certainly new. He sent me for
comparison a specimen of Paromalus biarculus Marseul from Ceylon. This closely
resembles Paromalus gardineri in some respects, especially in the form of the
mesosternal transverse stria and of the under surface as a whole; but is abundantly
distinguished by its greater breadth, more oblong form, much finer punctuation of the
upper surface, by having the anterior marginal stria of the prothorax widely interrupted
SCOTT—COLEOPTERA; HYDROPHILIDAI, HISTERID A 227
in the middle, and by the special sculpture distinguishing the pygidium of the ¢ from
that of the ¢. Though Paromalus gardineri appears to be new, it is doubtful whether
it is truly endemic to the Seychelles, since (unlike the other two Seychelles species)
it was not found in the forests, but on a cultivated islet. It is dedicated to Professor
J. Stanley Gardiner.
Loc. Seychelles. Mahé: Long Island, VII. 1908, 10 specimens, 5 of which were
recorded to have been found in a decaying stem of a coconut-palm.
29. Paromalus calciger, sp. nov. (Plate 14, figs. 24, 25).
g. Angustus, oblongus, subconvexus, rufo-piceus, nitidus; capite sat dense punctato,
strid marginali haud ante oculos continuata’; prothorace dense punctato, utrinque pone
oculos parum impresso, antice angustato lateribus sat rotundatis, strié marginali pone
caput integra; elytris dense fortiter punctatis, striis nullis ; propygidio pygidioque sat
dense subtiliter punctatis; prosterno inter strias angusto, basi parum latiore, striis
subparallelis, inter coxas parum sinuatis ; mesosterno ante striam transversam punctato,
strid marginali antice late interrupti, strié transvers4 integra, biangulata, parte media
lat&é parumque arcuatd, partibus lateralibus oblique retrorsum currentibus; metasterno
segmentoque primo abdominis dense sat fortiter punctatis, punctis ad latera fortioribus ;
tibiis anticis apice subito conspicueque incurvatis, extus 4-dentatis. Long. corp. (incl.
cap.) 2 mm.
Form narrow, neither strongly depressed nor strongly convex. Head rather densely
and strongly punctured, a very high power showing finer punctures among the larger
ones: the stria is present on either side only as far as a point just above the front margin
of the eye: the clypeus (in one specimen at least) is obsoletely impressed on either
side. Prothorax with disc somewhat convex, narrowed in front with sides slightly
rounded, densely punctured, with a slight impression in front on either side behind
the eye, with the marginal stria fine but continuous along the front margin. Hlytra
densely and strongly punctate, the punctures near the suture becoming fine towards the
base; there is a slight obsolete impression (in some aspects hardly visible) at the base
just outside the middle. Propyyidiwm and pygidium finely and fairly densely punctured.
Prosternum with the space between the striz narrow, slightly broader at the base ; strize
subparallel, slightly sinuate between the cox, converging slightly (but not meeting) at
the extreme apex: anterior lobe finely punctate. Mesosternum (Pl. 14, fig. 25) with
marginal stria present at the sides, widely interrupted along the front margin; it just
turns the corner of the front margin, but does not reach the median emargination :
transverse stria complete and distinct, biangulate, the middle part wide (about as wide
as the two lateral parts together) and slightly arcuate (with convex side of the curve
directed backwards), the lateral parts running obliquely outwards and backwards and also
slightly curved: meso-metasternal suture just visible from certain aspects with a very
high power: in front of the transverse stria the mesosternum is punctate, but behind,
between it and the suture, almost impunctate. Metasternwm and 1st abdominal segment
densely and rather strongly punctured, punctures stronger at the sides.
The anterior tibia are bent strongly inwards at the apex, forming a conspicuous
228 PERCY SLADEN TRUST EXPEDITION
blunt heel-like projection behind the base of the tarsus (whence the name calczger, “ heel-
bearer”): anterior and posterior aspects are figured, Pl. 14, fig. 24, a and b (the anterior
tibia of Paromalus gardineri is also figured, Pl. 14, fig. 23, for comparison, to show the
more normal form). Externally the tibie are 4-toothed.
In a single specimen the remarkable form of the tibize might well be considered to be
an abnormality or monstrosity: but since there are three specimens all having them
formed in exactly the same way, it must be taken to be normal. All three specimens,
however, have been shown by dissection to be $, so the form of the tibize may be a sexual
character. In addition to this, the species is characterised by its narrow subconvex form
and dense punctuation, by having the frontal stria widely interrupted and the anterior
thoracic stria complete, and by the form of the mesosternal transverse stria. It is
absolutely distinct from either of the other Seychelles species. As to its affinities I can
say nothing, not having sufficient knowledge of the genus: but I have seen no species
with such tibize, nor closely resembling it in other ways, in the British Museum, nor have
I read any description closely agreeing with it.
Loc. Seychelles. Silhouette: endemic forest near Mont Pot-a-eau, ca. 1500 feet,
VIII. 1908, 1 g (Type). Mahé: high forest of Morne Blanc, X. 1908, 1 ¢; Cascade
Kstate, 800 feet or more, 1 ¢.
Bacantus, Leconte.
30. Bacanius ambiguus, Schmidt.
Bacanius ambiguus Schmidt, Ann. Soc. ent. France, 1893, Bulletin, p. 101; Alluaud,
Liste Coléopt., p. 110; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 21.
A large series, determined by comparison with cotypes kindly lent by Monsieur
Alluaud. There is considerable variation in size, length (not including the head) varying
from less than 0°8 to 0°9 mm.
Wings. I have examined these organs in 5 specimens (4 from Silhouette and
1 from Félicité), and find them fully developed, about twice the length of the elytra.
Loc.* Seychelles. Silhouette: from under bark of felled dicotyledonous trees
among coconut-plantations near coast, Pointe Etienne, 17. IX. 1908, over 100 specimens ;
from forest near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908, 4 specimens; found under
bark of partly decayed stem of a Dracaena (together with one Acritus daubani), Mare aux
Cochons, IX. 1908, 5 specimens. Mahé: from a felled coconut-stem on Long Island, VII.
1908, 2 specimens. Praslin: Cétes d’Or Estate, XI. 1908, 2 specimens. TF élicité: from
forest, XII. 1908, 7 specimens. La Digue: taken in numbers in company with Acritus
(Halacritus) algarum under seaweed on the sandy beach, 1892 (Alluaud).
* Bickhardt, in Junk and Schenkling’s Coleopt. Cat., Part 24, Histeride, 1910, p. 85, gives ‘‘ Malacca” as
the only locality of Bacanius ambiguus. I have before me a copy of Schmidt’s paper, referred to above, in
which he describes the species from the Seychelles, and I know of no subsequent record of its occurring
elsewhere. Bickhardt’s reference to Schmidt’s description is otherwise correct.
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERID A 229
31. Bacanius mopinatus, Schmidt.
Bacanius inopinatus Schmidt, Ann. Soe. ent. France, 1893, Bulletin, p. 102; Alluaud,
Liste Coléopt., p. 110; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 21.
9 specimens, determined by comparison with a cotype kindly lent by Monsieur
Alluaud.
Wings: examined in | specimen, fully developed, twice as long as the elytron.
Loc. Seychelles. Mahé: Cascade Estate, 800 feet or over, 1908—9; 1892
(Alluaud).
32. Bacanius atomarius, Sharp.
Bacanius atomarius Sharp, Trans. R. Dublin Soc., iii, 1885, p. 128; Scott, Fauna
Hawaiiensis, i. p. 510, Pl. 15, fig. 37, 1908. ;
Two specimens from the Seychelles are smaller than the Hawaiian examples, but
after close comparison of both upper and under surfaces, side by side under the microscope,
I am unable to find any distinguishing characters.
Loc. Seychelles. Mahé: Cascade Estate, 800 feet or over, 2 specimens. Hawaiian
Islands (Oahu).
Acritus, Leconte.
Subgenus Haxicrirus Schmidt, Bull. Soc. ent. France, 1893, p. 103.
This subgenus is distinguished from A/etes and Acritus s. str. by the mesosternum
being produced forward into a blunt point, and by the anterior tibize being dilated,
emarginate at the extremity, and furnished with short spines along their outer edge. In
the specimens before me (Halacritus algarum) there is a small visible scutellum, as in
Acritus s. str. Also the diverging striz on the metasternum become obsolete behind :
they diverge rather widely, and then end blindly without curving at all: under a very
high power there can be faintly seen a trace of a stria running down from the anterior
corner of the metasternum to meet the diverging stria, but the two do not unite, and
their course is such that, even if they did, they would form an angular or V-shaped stria,
instead of the normal stria curving gradually round behind the middle coxa. Moreover,
the striz on the 1st abdominal segment, although they curve normally, appear to become
obsolete and not to reach the side edge of the segment. A somewhat analogous condition
is seen in the Hawaiian Acritus (dletes) subrotundus (Fauna Hawaiiensis, iil. pp. 533—4,
Pl. 15, fig. 33): in this, however, the striz on both metasternum and 1st abdominal
seoment diverge very little and become obliterated behind after a very short course: this
species belongs to a different subgenus and differs entirely from Halacritus in other ways.
This character of the diverging striz in Halacritus appears not to have been noted by
Schmidt.
The nature of the wings is discussed below, under Halacritus algarum.
33. Acritus (Halacritus) algarum, Schmidt (Plate 14, fig. 26).
Acritus (Halacritus) algarum Schmidt, Ann. Soc. ent. France, 1893, Bulletin,
p- 103; Alluaud, Liste Coléopt., p. 111; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p. 22.
SECOND SERIES—ZOOLOGY, VOL. XVI. 30
230 PERCY SLADEN TRUST EXPEDITION
The specimens before me consist of 18 from Mahé, 4 from Ceylon (named and given
by Mr George Lewis), and 8 from Perim Island. Many of the Seychelles specimens are
much larger and darker than the others. Those from Perim are especially small and
some of them light in colour; but I have seen no characters indicating the presence of
more than one species.
Wings: examined in 5 specimens from Mahé, 2 from Ceylon and 2 from Perim ;
in all these found to be reduced to minute vestiges, scarcely more than 4 the
length of the elytron. Micrometer measurements of wing- and elytron-lengths have
been made in 2 Perim specimens, 2 Ceylon specimens, and 2 Seychelles specimens.
Allowance must be made for error due to crumpling of the wing, &c. ‘The size of the
organs of course varies with that of the individual. In the Perim specimens the wing is
0719 mm., the elytron 0°51 mm. long: in a large Seychelles specimen, wing 0°23 mm.,
elytron 0°6 mm. The ratios between wing-length and elytron-length are 9:26 in the
Perim specimens, 10:27 in the Ceylon and one of the Seychelles specimens, 11:29 in
the other Seychelles specimen. Relative sizes of wing and elytron are shown in PI. 14,
fig. 26.
Reduced wings have also been found in Acritus (diletes) longipes Sharp and its var.
haleakale Scott and in Acritus (diletes) subalatus Scott, all from the Hawaiian Islands (see
Fauna Hawaiiensis, iii. pp. 529, 530, and Pl. 15, figs. 25, 26).
Loc. Seychelles. Mahé: Long Island, from seaweed on beach, VII. 1908,
8 specimens; Anonyme Island, from seaweed, I. 1909, 10 specimens. La Digue and
Marie Anne Islands, under seaweed, 1892 (Alluaud). Ceylon. Perim Island, Red Sea
(J. J. Walker, voyage of H.M.S. “ Penguin”; specimens in British Museum).
Subgenus AiLerEs, Horn, Proc. Am. Phil. Soc., xii. 18738, p. 356.
Apart from Acritus (Halacritus) algarum, no member of the genus Acritus was
previously known from the Seychelles. 3 species of the subgenus //etes have now been
found there, and are described below. Each is represented by 3 specimens, and all were
found in the mountain-forests. All 3 species are minute forms, small even for the genus
Acritus. The complete absence of a visible scutellum is the only character separating
Afletes from subgenus Acritus s. str. According to Bickhardt’s Catalogue of Histeridz,
1910, no Aletes is recorded from Madagascar or S. Asia: no member of the whole genus
Acritus is known from continental Africa: and the only two species known from
Madagascar (Acritus alluaudi Schmidt and madagascariensis Schmidt, both of which
I have seen through the kindness of Monsieur Alluaud) belong to the subgenus Acrztus
s. str. It may be noted that of 33 species of Acritus known from the Hawaiian
Islands, 82 are dletes and only 1 belongs to Acritus s. str. (see Fauna Hawaiiensis, i.
p. 514, &c.).
34. Acritus (Aletes) daubam, sp. nov. (Plate 14, figs. 27, 28).
Ovalis, convexus, nitidus, ferrugineo-piceus, antennis pedibusque fusco-testaceis ;
capite rare tenuissime punctulato; prothorace haud dense punctato, ante basin in medio
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERID Ai 231
linea transversi crenulato-punctata parum arcuaté, spatio inter hane lineam et basin
impunctato ; elytris tenuiter haud dense punctatis; pygidio fere impunctato; prosterno
inter strias circa dimidio longiore quam basi lato, striis antice divergentibus ; mesosterno
strid marginali integri; lined transvers’ punctorum elongatorum inter meso- et
metasternum ; metasterno in medio fere impunctato, ad latera sat fortiter haud dense
punctato ; segmento primo abdominis fere impunctato. Long. corp. { mm.
Rather narrowly oval, moderately convex, shining, pitchy-ferruginous with legs and
antennze paler. Head appearing impunctate, but under a high power a few very fine
punctures are visible; no suture between frons and clypeus can be seen. The punctures
on the prothorax are spaced out, separated usually by about twice their own diameter : in
front of the base in the middle is a slightly arcuate crenate-punctate line, the space
between it and the base being impunctate and not impressed; this line is not continued
laterally to the posterior angles of the thorax. The elytral punctures are slightly less
strong than those on the thorax, shallow and spaced out, so that except under a high
power the elytra sometimes appear impunctate. I have had difficulty in satisfactorily
making out the pygidial punctuation: the propygidiwm appears to be excessively finely
punctulate, the pygidiwm almost impunctate. The prosternum (PI. 14, fig. 28) between
the striz is about 14 times as long as its breadth at the base ; the strize are very slightly
nearer together in the middle than at the base, in front they diverge rather widely.
Mesosternum (Pl. 14, fig. 28) with marginal stria entire, with the surface bearing a few
excessively fine punctures only visible under a high power: separated from the
metasternum by a chain of strong longitudinally-elongated depressions. Metasternum
almost impunctate in the middle, rather strongly but not closely punctate at the sides.
First abdominal segment almost impunctate, with a few punctures visible under a high
power.
Characterised by its rather narrowly oval and convex form, numerous but not dense
punctures, the presence of the arcuate punctate line at the base of the thorax, and of the
transverse series of elongate punctures between meso- and metasternum.
This species is dedicated to Monsieur E. Dauban, owner of the island of Silhouette,
in which all 3 specimens were found.
Loc. Seychelles. Silhouette: found, together with many Bacanius ambiguus,
under bark of a felled dicotyledonous tree among coconut-plantations near coast, Pointe
Etienne, 17. IX. 1908, 2 specimens; found under bark of partly decayed stem of a
Dracena, together with several Bacanius ambiguus, in forest near Mare aux Cochons, IX.
1908, 1 specimen.
35. Acritus (Ailetes) davidsoni, sp. nov. (Plate 14, figs. 29, 30).
Sat late ovalis, parum depressus, nitidus, piceo-ferrugineus, antennis pedibusque
dilutioribus ; capite subtilissime haud dense punctulato; prothorace elytrisque dense
striolato-punctatis, striolis ad mediam basin prothoracis et ad angulos internos basales
elytrorum convergentibus; prothorace serie basali punctorum munito; pygidio dense
subtiliter striolato-punctulato, striolis transversis; prosterno inter strias subtilissime
punctulato, parum longiore quam basi lato, antice et postice fere wque lato, im medio
30—2
232 PERCY SLADEN TRUST EXPEDITION
parum angustiore ; mesosterno strid marginali in medio interrupta ; linea inter meso- et
metasternum tenui, haud punctata; meso- et metasterno et parte anteriore segmenti
abdominalis primi dense sat fortiter striolato-punctatis. Long. corp. $ mm.
Form rather broadly oval and a little depressed ; colour reddish-pitchy. I have been
unable to see any suture between clypeus and frons. The thorax and elytra are fairly
closely punctured, the punctures being drawn out before and behind into extremely fine
scratches, producing a dense striolation: the striole curve and bend slightly, and their
courses follow different directions in different parts; but there is a general convergence
towards the middle of the base of the thorax and towards the inner basal angles of the
elytra, towards which points also the actual punctures become stronger and closer
together; towards the shoulders of the elytra the punctures become obsolete, but the
striole are still present. In addition, the prothorax bears a series of punctures closely
following the basal margin. Even on the head traces of this striolation are just visible.
The propygidium and pygidium are also closely and finely striolate-punctate, the general
direction of the striolz being transverse.
The prosternum (Pl. 14, fig. 30) between the striz is a little longer than broad at
the base, narrowest in the middle, with front and hind margins of nearly equal breadth.
The mesosternum (Pl. 14, fig. 30) has the marginal stria interrupted in the middle ; just
within this stria on either side is a series of very fine punctures. The division between
meso- and metasternum is visible as a fine line, but without any special sculpture. Meso-
and metasternwm strongly punctured, the punctures closer in the middle of the
metasternum than at the sides, drawn out into short striole longitudinal in direction.
First abdominal segment with sculpture of a similar nature, but much sparser, almost
absent towards the posterior margin. Even the prosternum bears a few very fine
punctures and a minute trace of striolation.
The above description is made from two specimens. There is a third which
resembles them in form, structure, and colour, but differs in being totally devoid of the
characteristic striolation. The punctures are present, and distributed in the same
manner, but are not drawn out into striole at all. I am inclined to think the difference
may be sexual, but paucity of material prevents my attempting to prove this by
dissection.
This species is dedicated to His Excellency Mr W. E. Davidson, Governor of the
Seychelles at the time of my visit.
Loc. Seychelles. Silhouette: high damp forest near Mont Pot-a-eau, ca. 1500 feet,
VIII. 1908, two specimens (one of the striolate specimens, and that devoid of striolation).
Mahé: from near Morne Blanc, X. 1908, one specimen.
36. Acritus (Aletes) fryert, sp. nov. (Plate 14, figs. 31, 32).
Minutissimus, oblongo-ovalis, depressus, ferrugineus, antennis pedibusque dilutioribus ;
capite impunctato; prothorace elytrisque dense subtiliter striolato-punctatis, punctis
subtilissimis, in basi prothoracis parum fortioribus ; prosterno inter strias in medio parum
angustiore quam in basi, striis antice parum divergentibus; lined inter meso- et meta-
SCOTT—COLEOPTERA; HYDROPHILIDA, HISTERIDA 233
sternum in medio punctis aliquibus majoribus; meso- et metasterno et segmento
abdominali primo subtiliter striolato-punctulatis. Long. corp. $ mm.
Form oblong-oval and depressed. Head impunctate, without visible suture between
frons and clypeus. Thorax and elytra covered with a dense fine striolation. The
punctures themselves are extremely fine and some way apart, but the fine striole into
which they are drawn out before and behind form a dense sculpture: towards the base of
the thorax the punctures are a little stronger, and they form a somewhat irregular series,
not very close, along the actual base. The general direction of the striole is much more
longitudinal than in the preceding species: there is no general convergence towards
middle of base of thorax or towards inner basal angles of elytra: at the apex of the elytra
the inner striole curve inwards towards the sutural angle. Sculpture of the pygidiwm
not satisfactorily made out; it appears to have extremely fine transverse striolations, but
to be smooth and unsculptured at the apex.
Prosternum (Pl. 14, fig. 32) between the striz a little more than 14 times as long
as broad at the base, slightly narrower in the middle than at the base, with striz
diverging somewhat in front ; surface slightly convex with a few fine striolations. It is
not possible to see whether the mesosternal marginal stria is entire or not, since its
median portion is hidden under the base of the prosternum. The division between meso-
and metasternum is visible: it has no definite transverse series of punctures, but in its
median part there are about six rather larger punctures placed transversely, not very
close together. Meso- and metasterna and 1st abdominal segment finely striolate-
punctulate, the striole being short and generally longitudinal in direction, and the
punctures very fine; this sculpture becomes obsolete at the hind margin of the Ist
abdominal segment.
Distinguished from the preceding by its much smaller size, oblong and flattened
form, by its very fine punctures, the different arrangement of the striole, &c.
This species is dedicated to Mr J. C. F. Fryer.
Loc. Seychelles. Mahé: Cascade Estate, over 800 feet, 1909, 1 specimen; Mare
aux Cochons district, 1500 feet or over, 1909, 1 specimen. Praslin: Cétes d’Or Estate,
1908, 1 specimen.
Saprinus, Erichson.
Subgenus SAPRINUS, s. str.
37. Saprinus erichsoni, Marseul.
Saprinus erichsoni Marseul, Ann. Soc. ent. France, sér. 3, ill. 1855, p. 363, Pl. 16,
fig. 8; Alluaud, Liste Coléopt., p. 110; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, 1D Bil.
Not obtained by the Percy Sladen Trust Expedition. The single specimen found by
Alluaud was stated by J. Schmidt (Bull. Soc. ent. France, 1893, p. 101) to be perfectly
identical with specimens from Madagascar.
Loc. Seychelles: Mahé, 1892 (Alluaud). Madagascar.
234 PERCY SLADEN TRUST EXPEDITION
Subgenus Hypocaccus Thomson; Lewis, Ann. Mag. Nat. Hist., ser. 7, iv. pp. 1—8,
figs. 6, 7.
38. Saprinus (Hypocaccus) disjunctus, Marseul.
Saprinus disjunctus Marseul, Ann. Soc. ent. France, sér. 3, 11. 1855, p. 731, Pl. 20,
fig. 163; Alluaud, Liste Coléopt., p. 109.
22 specimens, determined by comparison with a named specimen from Madagascar
(kindly given by Mr George Lewis).
Loc. Cargados Carajos Islands: Establishment Island, 28. VIII. 1905, common in
dead birds. Madagascar. Comoros (teste Alluaud).
39. Saprinus (Hypocaccus) apricarius, Erichson.
Saprinus apricarius Erichson, in Klug, Jahrb. Ins., 1. 1834, p. 194.
4 specimens from Seychelles and 1 from Providence: specimens submitted to
Mr George Lewis, by whom the determination was confirmed. In all the specimens the
elytral punctuation is reduced in area, the punctures not extending forward at all along
the interstices between the 2nd, 3rd, and 4th dorsal strize: the species is very variable m
this respect. The Seychelles specimens are smaller, 24—22 mm. long (without head),
and have the elytral punctures stronger; they have two irregular rows of strong
punctures along the base of the thorax; they fairly closely resemble some South African
specimens in the British Museum.
The specimen from Providence is larger, about 34 mm. long (without head); it has
the punctures on thorax and elytra much finer, those along the base of the thorax being
reduced to a single series of fine punctures: in punctuation it somewhat resembles some
specimens from Zanzibar in the British Museum, but these are not so large.
Loc. Seychelles: Bird Island, VII. 1908 (Fryer). Providence: Cerf Island, on
rotten fish, 3. X. 1905. South Europe, Africa, Southern Asia and Malay Archipelago.
DESCRIPTION OF PLATE 14
Fig. 1. Paromicrus carinatus, gen. et sp. nov., x 30.
His, 2 % 55 antenna.
Fig. 3 _ ‘5 labrum.
Fig. 4. = atomus, labrum.
Fig. 5 55 carinatus, mandible (perhaps slightly distorted).
Fig. 6 e A’ maxilla.
Fig. 7 i x labium.
iti, &. i m middle tibia and tarsus.
Ing, 5 ‘ elevated part of mesosternum: a, ventral view; 0, lateral view ;
S, meso-metasternal suture; ©, middle coxal cavity.
Fig. 10. Paromicrus thomasseti, elevated part of mesosternum, ventral view.
Fig. 11. - atomus, e 9 * 0
hale
TRANS. LINN. Soc. SER. 2.Z00L Vou. XV]. PL 14.
PERCY SLADEN TRUST EXPEDITION.
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University Press, Cambridge.
H.Scott & E.Wilson del.
COLEOPTERA:HYDROPHILIDA, HISTERIDA.
, 12,
, 118
ig. 14.
ig. 15.
ig. 16.
ig. 17.
18s
. 19.
, 20;
SCOTT—COLEOPTERA ; HYDROPHILIDA, HISTERID AG 235
Bourdonnaisia mahensis, gen. et sp. nov., g, x 24.
. 55 antenna ~ and @.
A 5 labrum.
“ mandible.
silhouette, maxilla.
i mahensis, last 3 joints of maxillary palp.
f ye labium.
A: e hind tibia and tarsus.
elevated part of mesosternum, and metasternum; middle coxe
»
22 By)
removed: S, meso-metasternal suture.
Fig. 21. Bourdonnaisia silhouette, same parts as in preceding figure.
Fig. 22. Paroosternum degayanwm, gen. et sp. nov.; under-surface, showing pro-, meso-, and
metasterna. ‘
Fig. 23. Paromalus gardineri, sp. nov., g’, front tibia, seen from in front.
Fig. 24. . calciger, sp. nov., ~, front tibia: a, seen from in front; 6, seen from behind.
Fig. 25. 5 x gf, mesosternum.
Fig. 26. Acritus (Halacritus) algarum Schmidt: outline of elytron and wing to same scale.
Fig. 27. 5 (diletes) daubani, sp. nov.
Fig. 28. - Ee 5 pro- and mesosterna.
Fig. 29. A = davidsont, sp. nov.
Fig. 30. 3 x 5 pro- and mesosterna.
Fig. 31. - s Sryert, sp. nov.
Fig. 32. “a 3 - prosternum.
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No. XI.—COLEOPTERA, CHRYSOMELIDA:: HISPINAD OF THE
SEYCHELLES.
By 8S. Mautik, F.E.S., University of Cambridge.
(CommunicaTeD BY Pror. J. STANLEY GARDINER, M.A., F.R.S., F.L.S.)
(Text-figures 1—5.)
Read Ist May, 1913.
Tue Hispine collected by the Percy Sladen Trust Expedition consist of two
interesting species. It is necessary to erect two new genera for them. Nesohispa,
n.g., resembles Xiphispa Chap., which occurs in Madagascar and in New Guinea.
Brontispa Sharp, which has also certain affinities with Nesohispa, occurs in Rodriguez
Island, Mauritius, Luzon, New Guinea, the Aru Islands, and New Pomerania (New
Britain). Nesohispa seems to be an intermediate form between Xiphispa and
Brontispa.
Rhabdotohispa, n.g., has a close resemblance to Celenomenodera Blanch., which
is distributed in Madagascar and West Africa. In Rhabdotohispa the apical joint
of the antenna seems to be formed by the fusion of several joints which are distinct
in Celenomenodera. The sensory pit under the last jomt of the antenna of Rhab-
dotohispa is a very remarkable character. Both species were taken only in the high
endemic mountain-forests*. In preparing this paper I have received great assistance
from Mr Hugh Scott, who has very kindly checked my work and has made the drawings
of the antenne.
Nesonispa, n.g. (Figs. 1 and 2).
Body elongate, parallel-sided, not very strongly depressed. Head longer than
broad, produced between the antenneze into a blunt process; near the middle of the
head there commences a longitudinal groove which widens out at the base of the
process and continues along this to its apex. Hyes viewed dorsally not so prominent
as in Brontispa Sharp, inferior, large, convex. Antenne (Fig. 2) 11-jointed, scarcely
reaching the base of the thorax; joint 1 longest, almost as long as 2 and 3 together,
almost cylindrical, constricted at the apex (the constricted portion somewhat resembling
a very short separate joint); joints 2—6 subequal in length, constricted at the base,
* The locality-lists would seem to indicate that these Hispine are in some way connected with certain of
the endemic palms.—H. Scorv.
SECOND SERIES—ZOOLOGY, VOL. XVI. 31
238 PERCY SLADEN TRUST EXPEDITION
dilated at the apex, the apical dilatation of the 2nd being less pronounced than that
of the others; these joints have somewhat the appearance of a string of beads, a very
distinctive character, and are seen under a high power to be distinctly but rather
sparsely punctate; joints 7—11 forming a slender acuminate club, joint 7 longer,
8—10 equal in length and each shorter than 7, apical joint shghtly longer and sharply
acuminate; under a high power joint 7 is seen to have a smooth surface, sparsely
punctate as in the preceding joints, while joints 8—11 are minutely rugulose and
covered with short pubescence. Mouwth-parts placed in a hollow surrounded by a
Fig. 1. Mesohispa lambagiras, x 8. Fig. 2. Wesohispa lambagiras, antenna from above.
raised edge, which is met in front by the mterantennal process, the under surface of
which is concave. Labial palpi 3-jointed, proximal joint short and subglobular,
2nd joint shorter than the apical, which is more incrassate than the 2nd and rounded
at apex. Mazillary palpi 4-jointed, joint 2 incrassate, long, joint 3 very short, apical
joint longest, slightly incrassate, bluntly pointed. Prothorax quadrate, slightly narrowed
in front with sides widely sinuate, with anterior angles obliquely truncate, forming a
sharp projection internally and an obtuse prominence externally; from this obtuse
prominence on either side a ridge runs obliquely backwards on the disc. Scutellum
MAULIK—COLEOPTERA, CHRYSOMELIDA): HISPINA OF THE SEYCHELLES 239
with the base straight and the apex rounded. Hlytra parallel-sided, slightly narrowed
behind, truncate at apex, projecting a little beyond the abdomen ; with eight straight
and regular series of punctures and an abbreviated scutellar series; interstices smooth
and flat, not raised at all into ribs or ridges. Prosternwm with a raised edge in
front which meets the anterior angles of the pronotum, truncate at the base, which
is closely applied to the nearly straight front edge of the mesosternum ; metasternum
with an impressed median longitudinal line. Front coxew prominent, hind coxze not
prominent. Two first ventral abdominal segments not soldered together. Legs short,
femora incrassate; in the tarsus the claw-joint projects beyond the third joint, the
claws not being partially hidden in the pubescence of the under side.
This genus resembles Brontispa Sharp in shape and general appearance, but differs
from it in having the head longer than broad, and the eyes not prominent, the antennze
scarcely reaching the base of the thorax, the Ist antennal joint relatively shorter, the
apical joint sharply acuminate, and the prothorax narrowed in front. From Xiphispa
Chap. it also differs in the length and structure of the antennz, the relative length
of the 1st and three following joints, in the form of the prothorax (which is conical in
Xiphispa), and in having the claw-joint of the tarsus projecting beyond the 3rd joint.
1. Nesohispa lambaguras*, n. sp. (Figs. 1 and 2).
Upper side black ; under side, legs, first six joints and a little of the basal portion
of the 7th jomt of the antennze dark reddish-brown: near the truncate apex of the
elytra the colour is a little diluted with dark red-brown. Surface of head strongly
punctate ; behind there is an almost impunctate portion in the occipital region, which
under a high power is seen to have transverse striations. The sinuate sides of the
prothorax have three indentations: its surface is convex and strongly punctate, except
in the posterior central portion, which is almost impunctate, but is seen under a high
power to have very fine punctures. Base of elytra bi-sinuate with the anterior angles
rounded.
Length: the type measures 81 mm.; a larger specimen is 94 mm. without the
head, which it has unfortunately lost.
Described from three examples. Type in the British Museum.
Loc. Seychelles. Mahé: from damp forest on summit of Morne Pilot, over
2000 feet, X. or XI. 1908 (same loc. and one of same dates as Rhabdotohispa scotti,
q.v.), 1 specimen (the type); Cascade Estate, 800—1500 feet, 1909, 1 specimen ;
found dead between leaf-bases of growing Stevensonia palm, summit of Mt. Sebert,
nearly 2000 feet, 28. XIT. 1908, 1 specimen.
RHABDOTOHISPA, n. g. (Figs. 3—5).
Body nearly parallel-sided, slightly broader behind. Head strongly transverse,
very slightly produced into a blunt prominence between the antenne. Hyes strongly
convex. Antenne (Fig. 4) short, 6-jointed ; joint 1 short, slightly incrassate, obliquely
* The specific name is derived from Sanskrit lamba =long, ciras = head, the word meaning “long-headed.”
S, ¢ 5) 5 §
31—2
240 PERCY SLADEN TRUST EXPEDITION
truncate on the inner side in the basal portion; joint 2 longer than 1, slightly incrassate,
convex on the inner side and nearly straight on the outer side, constricted at the
base, which causes the swollen articulating portion sometimes to appear as a small
separate joint ; the antenna is slightly elbowed at the base of joint 3, jomts 3—6
being directed shghtly outwards; jomts 3—5 subequal in length, subcylindric, broader
at apex than base, gradually broadening towards the apex of the antenna; joint 6
longest, in g nearly as long as 83—5 together, in $ about as long as 4 and 5 together,
gradually broadening towards the bluntly-rounded apex. The apical portion of this
6th joimt bears beneath a very remarkable excavation, with clean-cut and slightly
overhanging edge; this excavation is larger in f than in ? and differently shaped
Fig. 3. Rhabdotohispa scotti, 9, x 8. Fig. 4. Rhabdotohispa scotti, Vig. 5. Rhabdotohispa scottr; last joint
right antenna of 2 from of right antenna of @ and ¢, viewed
above. from beneath, showing the outline of
the sensory pit (the two figures to
same scale).
(Fig. 5), but in both sexes it is roughly U-shaped, the limb on the inner side of the
antenna being much broader than that on the outer side. It has not been possible
to examine satisfactorily the minute structure of the surface of the excavation, but
a balsam preparation under a high power shows the presence of small groups of
papilliform structures situated in pits which are surrounded by a dense mass of a
special form of short sete. Mouth-parts in a slight hollow situated transversely
between the eyes, the lower margin of which it reaches on either side, surrounded
by a slightly raised margin which is interrupted behind, and at either side in front.
Labrum pale and membranous, bearing on its front (upper) surface setee placed in a
sinuate transverse line some way behind the front margin. Mandibles subtriangular.
MAULIK—COLEOPTERA, CHRYSOMELIDA: HISPINA OF THE SEYCHELLES 241
Maaille with the outer lobe 2-joimted, the basal jomt being rather short and stout,
the apical joint subconical, pointing slightly inwards; inner lobe membranous; palpi
short, 4-jointed, joints 1—3 short and transverse, joint 4 incrassate, bluntly rounded
apically, more convex on the inner side, about as long as 1—3 together. Mentum
subtrapezoidal, broader in front than behind, with anterior angles rounded; ligula
membranous, projecting in front; labial palpi short, joints 1 and 2 short, joint 3
incrassate, blunt at apex, more convex on the inner side, about as long as 1 and 2
together. _Prothorax cylindrical, only slightly produced in front, not margined at
the sides, much narrower at the base than the base of the elytra, with an almost
impunctate longitudinal line reaching nearly from apex to base. Scutellwm shield-like,
base straight, apex widely rounded, surface seen under a high power minutely reticu-
late. Hlytra with sides nearly straight, slightly broadening behind, with anterior
angles rounded ; towards the apex the lateral margin bends inwards, forming a widely
rounded obtuse angle; suture raised into a ridge which commences a little behind
the scutellum; also with three other longitudinal ridges on each elytron, converging
somewhat at the base, and united by somewhat irregular transverse ridges: there
are two rows of very rugose punctures between each pair of ridges; these punctures
become so wide on the lateral surface as to run into each other. The surface generally
presents a very rugose and rough appearance. Under side finely and sparsely punctate.
First and second ventral abdominal segments not soldered. Legs short, femora mode-
rately incrassate.
This genus resembles Celanomenodera Blanch. in general shape and form, but
the prothorax is not produced so far over-the head in front. It has a cylindrical
prothorax like that of Promecotheca Blanch., but without the posterior constriction.
2. Rhabdotohispa scotti, n. sp. (Figs. 3—5).
?. Black; with two longitudinal lines on the prothorax, with irregular patches
on the disc and the extreme apex of the elytra, with apices of femora and bases of
tibize inwardly, and usually with the 4th, 5th and basal half of the 6th antennal joints,
ochreous.
These pale markings are variable. In the hind and middle femora and tibize
they are very much obscured with black, sometimes completely absent. They are
constant at the extreme apices of the elytra, and fairly constant on the fore femora
and tibie. In the antennee the pale colour sometimes appears in the second joint
also; on the other hand it may be very much obscured in joints 4, 5, and basal
half of 6.
g. Smaller; the extreme apices of elytra are never ochreous, and the whole
insect may be completely black except the apices of the fore femora and the bases of
the tibize, and the 5th and basal fourth of the 6th joints of antennee, which are tinged
with ochreous.
From the gradual variation observable in the examples before me it may be
fairly concluded that some specimens may be completely black without a trace of
ochreous.
31—3
242 PERCY SLADEN TRUST EXPEDITION
Length : 33—4} mm.
Described from 9 examples, 5 g and 4 3. Type in the British Museum.
Loc. Seychelles. Mahé: beaten from Roscheria palm-trees in damp forest on
summit of Morne Pilot, over 2000 feet, 29. X. and 15. XI. 1908, 4 2, 3 2; found sitting
on leaves of Stevensonia palm, in a piece of secondary native forest, Mare aux Cochons
district about 1500 feet, in the late afternoons of 30. I. and 2. II. 1909, 1 g 1 @.
No. XIL—NEUROPTERA, MYRMELEONIDA FROM THE INDIAN OCEAN.
By James G. NEEDHAM.
(CommunicateD By Pror. J. Sranney Garpiner, M.A., F.R.S., F.LS.)
(Text-Figures 1—3.)
Read 5th June, 1913.
Four species of antlions, collected in the Seychelles and other islands of the south-
western part of the Indian Ocean by the Percy Sladen Trust Expedition, were sent to me
by Mr Hugh Scott for study, and are here reported upon. One of the species is the well-
known Myrmeleon obscurus Rambur which is widely spread in that part of the world.
Rambur and Walker reported it from Mauritius, and in the present collection it is
represented by specimens from nearly all the islands that were visited. A new species
of Formicaleo, belonging to the subgenus Distoleon, is represented by a single specimen
from Mahé, Seychelles Islands. The other two species, singularly enough, were collected
only from the coral atoll of Aldabra. One of these, a fine Acanthaclisis, 1s new to science.
The list, with localities and other data, together with descriptions of the new species
follows.
1. Myrmeleon obscurus, Rambur (Fig. 1).
Myrmeleon obscurus Rambur, Hist. Nat. Ins. Neur., 1842, p. 403; Walker, Cat.
Neur., Part Il, 1853, p. 370.
. Deane ae
: CLL. ke SS j
be | oe ioral 4 ES =
& s : i H = LES Le 3 SS SS
Fig. 1. Myrmeleon obscurus Rambur, wings.
This species is represented by fifteen specimens from a number of different islands.
Length to wing tips 27 mm., expanse of wings 50 mm. The specimens agree well with
Walker’s description, which is ample for recognition.
244 PERCY SLADEN TRUST EXPEDITION
Loc. Aldabra, 1908 (Fryer). Farquhar Atoll, 1905. Amirantes: Poivre and
d’Arros Islands, X. 1905. Coetivy, 1905. Seychelles: Bird Island, 1908 (Fryer) ;
Dennis Island, 1908 (Fryer); Silhouette, near coast*, 1908; Mahé, 1905 and 1908.
Described from Mauritius: widely distributed in Africa.
2. Acanthaclisis maritimus, sp. nov. (Fig. 2).
Length 30 mm., male appendages 5 mm. additional; female 33 mm. exclusive of
appendages. Antenne 10 mm.; fore wing 41 mm., hind wing 37 mm. Expanse of fore
wings 86 mm.
A rather elongate, long-winged species. Colours brown and yellow. Face yellow:
head above from the base of the antennee upward and backward fuscous, with three
transverse rows of yellow spots, the spots of the middle row being smaller than on those
of the other rows. Antenne fuscous, annulate with paler brown ; regularly increasing in
diameter from the second segment to near the end of the club, and mucronate on the very
Wa LR
Fig. 2. Acanthaclisis maritimus, sp. nov., wings; cu, cubital vein; r, radial planate; «, median nexus.
tip. Prothorax one half longer than wide, grayish above, its sides yellow, with a pair of
rather broad dorsal stripes that are made up of vermiculate fuscous markings. The mid-
dorsal grayish tract is sprinkled with minute fuscous dots, that are sometimes confluent
in three pairs of indistinct dashes. Meso- and metathorax above grayish, marked
irregularly with fuscous, and suffused with fuscous at the junction of the segments above.
The rather long and sparse hairs clothing the thorax are fuscous in front and above, and
white at the sides and in the rear. Legs yellow, thinly clothed with long yellow hairs,
and the femora beset with numerous short blackish spines. The tibize of the fore and
middle legs are doubly ringed with fuscous, one ring being apical and the other near the
base, the latter on the fore tibia being more extensive and sometimes divided. Tarsi
yellow, with the base of the first segment, all of the second, third and fourth segments
* Tn 1908—9 I never observed WM. obscurus in the endemic mountain-forests of Mahé and Silhouette, but
obtained specimens only from the lower parts of the islands.—H. Scorr.
NEEDHAM—NEUROPTERA, MYRMELEONIDA FROM THE INDIAN OCEAN 245
and the tip of the fifth segment, fuscous : spurs and claws brown: spurs gently curved, as
long as the three basal joints of the tarsus.
Wings grayish with a yellowish stigma, veins alternately fuscous and yellow, as are
also some of the terminal forks. Im the basal part of the subcostal cell are numerous
clustered dots, with three groups of such dots farther out beyond the base of the radial
sector. There is a brown spot, usually conspicuous (in one specimen apparently absent)
on the apex of the cubito-anal loop, sometimes double and very variable. There are some
indistinct brownish touches along the middle of the median vein and on the nexus near
the tip of that vein, on a few cross-veins immediately before this nexus and about the
middle of the hind border. The hind wing is unmarked behind the radial vein save for
faint touches of brown at the median nexus. The two cross-veins in the space beyond
the stigma, behind the vein #,, are tinged with brown. The veins in the costal area are
forked from the level of the base of the radial sector to the stigma, and connected
together by a rather irregular series of gradates. The radial planate is well detined
(although forming a broken line) in both wings, but the cubital planate is little developed,
its course being hardly traceable. The wings are elongate, obtuse at the tips and parallel-
sided from the middle to the level of the stigma.
Abdomen slender, inconspicuously clothed with a very short smooth brown pubescence.
Colour yellowish, with large subapical dorsal spots on all the segments, overspreading
segments 2, 7, and 8, transversely divided on segment 3, and divided lengthwise by a
narrow yellow median line on segment 6. Male appendages yellow, thick and appressed-
parallel at base, forcipate beyond, slender and held horizontally, and thickly clothed with
fuscous recurved hairs. |
Of the four specimens two are males and two are females. One of the males is
almost destitute of the larger brownish wing markings, but is in all other respects
apparently identical with the other specimens.
Loc. Aldabra. This fine species is represented by four specimens collected by
Mr J. C. F. Fryer (1908).
3. Hormicaleo ornatus, sp. noy. (Fig. 3).
A slender pale fulvous species, closely allied to F. bistrigatus (Rambur) from Tahiti
and Australia.
Length about 27 mm. Expanse of wings 78 mm. Antennze long (9 mm. of one are
present, with the clubbed end entirely lacking), yellowish-brown, slightly darker at the
ends, and with black half rings about the basal segments on the dorsal side. Face yellow
below. A broad fuscous band between the eyes surrounds the bases of the antennze
posteriorly, and there is another paler brownish band extending transversely between the
eyes across the rear of the head above. Wings long and narrow with delicate venation
and elegant markings and iridescence. Fore and hind wings of equal length, both are
widest just before the stigma, which is indistinct. The veins are brownish. There is a
quadrate brown spot on the first cross-vein beyond the stigma, behind vem #,, and there
is more or less of brown on the cross-vein next following. There is a similar brown spot
in the fore wing on the median nexus and there are paler spots on three or four of the
246 PERCY SLADEN TRUST EXPEDITION
gradate cross-veins just behind the nexus. There are also slight touches of brown along
the basal portion of the cubital vein where joined by cross-veins in the fore wing, and
there are two conspicuous oblique brown streaks ; one covering the cubital are and being
half a dozen or more cells long, and the other traversing the base of the terminal forks of
the radial sector, and being parallel to the hind margin. In the hind wing there is but
one streak, corresponding in position to the one last mentioned, but being a little larger,
filling the space between the marginal forks and the last gradate series of cross-veins and
extending to the apex of the wing.
Legs slender, yellow, sprinkled with fuscous dots that are more or less confluent
about the apices of femora and tibiz. The colour is paler on each succeeding pair, the
hind femora being wholly yellow, except a spot at the tip. The spines that thinly clothe
all the legs are blackish, and the spurs and claws are brown. The spurs are gently
curved, but the claws are straight to very near their tips. The terminal tarsal seement
is clothed beneath with a thick brush of deep black clavate hairs. Abdomen wholly
3. Formicaleo ornatus, sp. nov., wings.
obscure pale brownish, with a suggestion of two dark brownish areas toward the ends of
each segment dorsally. The first tarsal segment is slightly shorter than the fifth, which
nearly equals the second, third and fourth taken together.
This species lacks the prothoracic stripes of Ff’. bistrigatus and also the brown dashes
along the cubital vein, and it has, along with lesser markings, the distinct conspicuous
brown streak along the cubital are in the fore wing, which that species lacks. It falls in
the subgenus Distoleon Banks.
Loc. Seychelles. Mahé: Cascade Estate, one specimen labelled “at light,” 1. XI.
1905 (Gardiner).
4, Creagris pervigil (Walker).
Myrmeleon pervigil Walker, Cat. Neur., Part II. 1853, p. 354.
I refer two specimens from Aldabra with considerable hesitancy to Walker's species,
the type of which I have not seen. They agree fairly well in colour and markings but are
smaller in size than the measurements given by Walker.
Loc. Aldabra (Fryer). Walker’s species is described from Natal.
opt ie
No. XIII.—COLEOPTERA: ANTHRIBID AL.
By K. Jorpan, Ph.D.
(ComMUNICATED BY Pror. J. STANLEY GaRpINER, M.A., F.R.S., F.L.S.)
(Plate 15.)
Read Ist May, 1913.
ONLY two species of Anthribidze have hitherto been recorded from the Seychelles.
The present collection contains no less than 21 species, the majority being minute and
many represented by a long series of specimens, testifying to the diligent search bestowed
by Mr Hugh Scott on small Coleoptera.
The greater number of the Seychelles Anthribidze are of small size, a feature also met
with in the Anthribidee of St Helena, New Zealand, and the northern temperate regions,
and a large percentage is indigenous, belonging to genera not known to be represented
outside the Seychelles.
As must be expected, the affinities of the species are with Madagascan and African
forms on the one hand and Indian species on the other. The majority of the species
belong to that division of the family in which the antennz are inserted on the dorsal
surface of the rostrum and which we call Anocering Lac. It is worthy of note that all
the Anthribide known from St Helena also belong to this division, which is likewise more
abundantly represented in Madagascar than in Africa, the Oriental Region and America.
While, however, the Mascarene Anocerinz belong to a large extent to Caranistes and
allied genera, this genus has not been found in the Seychelles. Nor has a species of
Tophoderes been met with, a genus abundant in species on Madagascar. Xenotropis
rugicollis, Fairm. (1895), described from La Digue, but not contained in the present
collection, is of Mascarene affinity; Phlaobius gugas cervinus, Klug (1833), as well as a
new species of Hpitaphius, also point to Madagascar, and Phla@obius pustulosus, Gerst. (1871)
is found, besides the Seychelles, in Madagascar and in Kast Africa. On the other hand,
Contexta murina, Jord. (1901) was only known from Ceylon, and the new genus
Hormiscops comes nearest to the Oriental genus Phaulimaa.
A feature not without interest is the great similarity which some of the minute
Anthribidz bear to species of other families of beetles, such as Clerids, Anthicids and
Scaphidiids, the resemblance being so close that I only detected the strangers among the
Seychelles Anthribidze when I began to work out the collection.
SECOND SERIES—ZOOLOGY, VOL. XVI. 32
248 PERCY SLADEN TRUST EXPEDITION
XeENoTRoPIS, Fairm. (1895).
Xenotropis, Fairm., Bull. Soc. Ent. France, (1895) p. 281. Allied to the Madagascan
genus Diastatotropis Lac. (1866).
1. Xenotropis rugicollis, Fairm. (1895).
Xenotropis rugicollis, Fairm., l.c.; Kolbe, Mitt. Zool. Mus. Berlin, 5. p. 39 (1910).
This fairly large species (length 13 mm.) was not met with during the Seychelles
Expedition. Fairmaire described it from La Digue.
SINTOROPS, gen. nov.
$¢. Elongatus. Rostrum cylindricum vel subcylindricum, apice valde dilatatum, in
ft multo longius quam in ¢. Oculi laterales, fortiter granulosi elliptici. Antenne sat
longee tenues, clava valde compressa angusta laxissima. Antennarum fovez sublunatze
haud sulciformes. Labiophorus (=pedunculus gularis) tridentatus (¢) vel unidentatus
(¢). Tarsorum articulus tertius latus lobis rotundatis; unguiculi dentibus magnis
instructi.—Genotypus: S. alleus, spec. nov.
In facies and the position and shape of the eyes the sexually dimorphic species for
which this new genus is proposed bears a rather close resemblance to Sintor Schonh. (1839).
The coarse granulation of the eyes recalls the African Anthribid known as Ischnocerus
nigellus Sparrm. (1785), and the very loose club of the antennz the Oriental genus
Nessiara Pase. (1860). For the present Sintorops is best placed near Ischnocerus.
The new genus can at once be recognised by the anterior margin of the false mentum
or gular plate (for which I propose to employ the term labiophore) being produced in the
dé into three large teeth and in the ? into a single median one, which is rounded like the
corresponding tooth of the ¢.
The sexual difference in the length of the rostrum in Sintorops is a feature not yet
met with among the Anthribide in such a marked degree, the difference being even more
pronounced than in the American Systaltocerus platyrhinus Labr. and Imh. (1842). The
rostrum, in the ?, is approximately as long as it is broad at the apex, while it is three
times as long in the ¢, the structure of the rostrum also not being the same in the sexes.
In the ¢ the narrow proximal portion is cylindrical and bears a distinct median carina, and
the three teeth of the labiophore are large and prominent, being directed obliquely down-
ward, the apex of the rostrum, inclusive of the mouth-parts, in a lateral aspect resembling
the snout of some small carnivorous mammal with the mouth open. In the ? the rostrum
is more flattened; the median carina of the upper surface is at most vestigial, and the
lateral teeth of the labiophore are obsolete, while the median tooth is horizontal and
therefore hardly visible in a frontal or lateral view of the snout. The apical widening of
the rostrum is quite abrupt in both sexes, a distinctly projecting angle being formed
proximally to the base of the mandibles some distance below the insertion of the
antenna.
The apical margin of the rostrum is depressed and feebly emarginate, the strongly
convex upper lip being exserted. The mandibles are very flat and bear a blunt tooth near
JORDAN—COLEOPTERA: ANTHRIBIDA 249
the base and a small excision on the distal side of this tooth. The labium is narrow, being
divided into two slender divergent lobes from the apex halfway down to the insertion of
the palpi.
The antennee resemble to some extent those of Sintor and Ischnocerus ; they reach
about to the centre of the elytra in the $ and to the basal fourth of the same in the ?,
being clothed with very thin hairs, which form a longer pile on the three segments of the
club than in the allied genera, the club and the distal segments of the shaft bearing also
some bristles. Segments 1 and 2 are short and somewhat incrassate, 3—8 long and
slender, 3 being the longest, 9—11 flat, elongate-ovate, nearly the same in length, 9 rather
more triangular, 9 and 10 slightly truncate at the apex, 10 and 11 joined to the previous
segment with a very narrow base. The antennal groove is situated close to and partly
upon the dilated apical portion of the rostrum, being halfmoon-shaped.
The frons narrows considerably towards the rostrum, but remains nevertheless very
broad in both sexes.
The prothorax is slightly broader than it is long; the anterior carina is slightly
undulate and extends in an almost even curve to the middle of the sides; the basal
carina is distinct, but there is no subbasal one, whereas the lateral basal longitudinal
carina is indicated by a line composed of a few granules, the line forming an acute angle
with the lateral carina.
The scutellum is small and semicircular. The elytra are only a very little wider
than the prothorax, slightly narrowing from the shoulders, being twice as long as they are
broad together. The pygidium has the apex obtusely pointed and upturned in the 3, and
subtruncate in the 2.
The pro- and mesosternal intercoxal processes are narrow. The metasternum bears
posteriorly a carina at each side of the median line, and the antecoxal sclerite of this
sternite is completely divided in the centre.
The legs are nearly the same in. the sexes, excepting the rather greater lengths of the
d-legs. The tarsi are somewhat over half the lengths of the tibize, the anterior tibia and
tarsus being the longest. The first segment is longer than the claw-segment in the fore
tarsus and as long as that segment in the hind tarsus. The second and third tarsal
segments are rather broader than they are long in both sexes, the lobes of the third being
rounded-ovate and very conspicuous. The teeth at the inside of the claws are large, flat
and convergent.
The ovipositor of the ? is very remarkable for its structure. The two halves are very
flat and thin (in a dorso-ventral sense) and diverge distally, each tapering to a sharp
point, the free apical portions being a little longer than the two gonapophyses are broad at
the point where they commence to diverge and each bearing at the outer edge a tooth
pointing frontad and the vestige of a second tooth. The eighth abdominal tergite of the
d (concealed under the pygidium) is deeply cleft. There is only one median vein in the
hind wing between the cell and the submedian (or anal veins), this median vein not
reaching to the short hook-like vein emanating from the cell.
32—2
SAGAS
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250 PERCY SLADEN TRUST EXPEDITION
2. Sintorops alleus, spec. nov. (Plate 15, figs. 1, 2).
$%. Sordide ochraceus vel nigro-brunneus, pube sat sparsa grisea parum maculatim
vestitus, elytrorum interstitiis alternis fusco-tessellatis, antennis pedibusque pallidioribus, -
antennarum articulis 9° et 10° apicibus exceptis brunneo-nigris, 11° luteo; omnino
grossissime ac dense punctatus.
Long. (cap. excl.) 3°5—5 mm.
A considerable series of both sexes was obtained. The colour varies from a dirty pale
ochreous to blackish brown, the suture being blackish in most pale $3, while many 22 bear
a blackish patch or transverse band at the commencement of the apical declivity of the
elytra. The grey pubescence does not conceal the colour of the derm and forms ill-defined
spots on the pronotum and elytra, a spot at the shoulders being especially noticeable.
There are no tufts or tubercles, the basal callosities of the elytra also being but feebly
developed.
The prothorax is rather abruptly constricted at the apex, the sides slightly slanting
from the dorsal carina forward in the ?, while they are parallel in the @.
Segment 3 of the g-antenna is half as long again as 1 and 2 together and one-third
longer than 8, 9 is one-fifth shorter than 8 and one-fifth longer than 10, 11 being a trifle
longer than 10. In the ? segment 3 is one-fourth or one-fifth longer than 8, and 8 to 11
are equal in length, only 10 being a very little shorter than the others.
Loc. Mahé: high forest of Morne Blanc, X. XI. 1908; high damp forest between
Trois Fréres and Morne Seychellois, about 1500—2000 ft., XII. 1908; Cascade Estate,
forest, about 1000 ft., I.—III. 1909, and near Mt. Harrison, 1700 ft., III. 1909 ; Mare aux
Cochons district, 1000—2000 ft., I. II. 1909. Silhouette: high forest above 2000 ft., VIII.
1908, also from 1500 ft.; Mare aux Cochons, IX.; high forest above Mare aux Cochons,
IX. 1908. Type from Mahé.
The specimens were beaten from dead leaves, etc. One has been “ bred from a larva
found in a fallen and rotting palm-leaf” above Mare aux Cochons on Silhouette.
HorMIscops, gen. nov.
$9. Phaulimie Pasc. (1860) affinis. Rostrum brevissimum, longitudine duplo latius,
ad apicem ampliatum, margine apicali recto. Oculi magni, obliqui, subdorsales, sat fortiter
granulosi, subintegri vel subtiliter sinuati. Frons capitis antrorsum angustata, antice
dimidio rostri angustior. Antenne breviter pilose, clava compressa articulis remotis.
Labium sat profunde divisum. Palpi maxillares crassi. Carina pronoti basalis, utrinque
antrorsum continuata laterum medium non attingens. Prosternum breve ; metasternum
inter coxas medias haud reflexo-marginatum, sclerito transverso antecoxali in medio
interrupto. Tarsorum articulus primus ceteris simul sumptis longior vel equalis ;
unguiculi fortiter dentati—Genotypus: H. tibealis, spec. nov.
There are four species in the collection. They are small Anthribids which recall by
their short and convex body the American genera Husphyrus Lec. (1876) and Ormiscus
Waterh. (1845), but are most nearly related to the Oriental genus Phaulunua, differing
JORDAN—COLEOPTERA: ANTHRIBID At 251
from it especially in the carina of the pronotum being basal and the club of the antenna
more loose. The eye is either feebly emarginate, or bears a distinct though small sinus.
The lobes of the labiophore are long and rounded. The labium is deeply excised, but the
sinus does not extend down to the insertion of the palpi. The maxillary palpi are
unusually stout. The buccal fissures are accompanied on the outer side, from the bases
of the mandibles to the apex of the fissures, by a rather sharply defined depression. The
antennee agree best with those of Nessiara Pasc. (1860). They are clothed with short
hairs and a few longer bristles. Segments 1 and 2 are incrassate, 3 is slender and about
as long as 9, 8 at most twice as long as it is broad, being only half the length of 9, 9—11
are flattened, 9 is triangular with the sides rounded distally, about half as long again as
broad, 10 as broad as it is long, 11 ovate, the apex of 9 and 10 truncate and the base of
10 and 11 thin, the club being loose. f
The pronotum is much broader than it is long, being conical, and bears rather large,
but dispersed and shallow, punctures. It is convex with a distinct flattening before the
carina. The latter is almost evenly incurved dorsally, straightening very faintly towards
the sides, and laterally curves forward in an obtuse angle with the point rounded off. The
true basal cariniform edge of the pronotum gradually curves forward laterally and joins
the main carina, forming with it a very acute angle. ‘The elytra are coarsely punctate-
striate. The prosternum is short. The intercoxal process of the mesosternum is truncate
with the angles somewhat rounded, the mid coxee not being notched.
3. Hormascops tibialis, spec. nov. (Plate 15, figs. 4, 4 a, 4D).
$?. Rufo-brunneus, luteo-griseo-pubescens, antennarum segmentis duobus primis
luteis, femoribus tibisque sordide luteis; pronoto duabus maculis magnis brunneis una
utrinque in disco posita, his maculis in medio disci conjunctis vel separatis ; elytris quoque
brunneo-signatis: macula subbasali dorsali, altera elongata fere lineiformi mediana, atque
fascia lata parum sinuosa limbali aut sublimbali ab humero ad apicem extensa. In ¢ tibiee
anticee crassze, tarsi antici tenues.
Long. (cap. excl.) 2°2 mm.
Several specimens of both sexes.
The broad fore tibia and slender fore tarsus of the ¢ are characteristics which are not
found in the following species. The ground-colour varies, some specimens being very
much paler than others. The frons is one-fourth the width of the rostrum in the g and
one-third in the ?. The brown markings of the elytra are sometimes so enlarged as
to render the elytra brown with grey spots. The pronotum has often a small brown
lateral median spot.
Loc. Mahé: high forest of Morne Blane, X. 1908; above Port Glaud, 500—
1000 ft., 5. XI. 1908; Mare aux Cochons, 1000—2000 ft., I. II. 1909. Silhouette:
Mare aux Cochons, IX. 1908. Praslin: Cétes d’Or Estate, XI. 1908. Félicité: forest,
XII. 1908. Type from Mahé.
4. Hormascops sobrinus, spec. nov. (Plate 15, fig. 5).
é?. Niger, griseo-pubescens, antennis pedibusque luteis vel luteo-rufis, pronoto et
252 PERCY SLADEN TRUST EXPEDITION
utroque elytro duabus maculis fusco-pubescentibus notatis ; oculis sinu parvo instructis ;
tibiis anticis maris non incrassatis.
Long. (cap. excl.) 1°5—2°5 mm.
A good series of both sexes.
The pronotum and elytra are rather more convex than in the previous species. The
two fuscous patches on the pronotum and the two patches on each elytrum are placed as
in H. tibialis, but are very inconspicuous on account of the black or brownish black colour
of the derm. A median stripe on the pronotum and the suture are often more densely
pubescent-grey. The fore tibia is very slightly broader in the ¢ than in the @.
Loc. Mahé: above Port Glaud, 500—1000 ft., XI. 1908; Morne Blanc, 1000 ft.,
XI. 1908; high damp forest at summit of Pilot, above 2000 ft., X. 1908; high damp
forest between Trois Fréres and Morne Seychellois, 1500—2000 ft., XII. 1908; Cascade
Estate, 800—1000 ft., I Hl. 1909; summit of Montagne Anse Major, in forest of
Capucin trees (Northea), 2000 ft., IT. 1909. Silhouette: Mont Pot-d-eau, VIII. 1908 ;
Mare aux Cochons, 1000 ft., IX. 1908 ; forest above Mare aux Cochons, IX. 1908. Marie
Anne : forest of “Takamaka” trees, “‘ Bois de Natte,” XII. 1908. Type from Mahé.
5. Hormiscops letus, spec. nov.
?. Nigro-brunneus, griseo-pubescens, antennarum basi, ore tibiisque luteis, pronoto
elytrisque luteis nigro-plagiatis, oculo simu parvo instructo, articulo 1° tarsorum quatuor
posticorum reliquis segmentis simul sumptis breviore.
Long. (cap. excl.) 2°5 mm.
Two specimens, one of them minus head and prothorax.
The centre of the head is blackish. The frons is broader than in the other species.
The sides of the pronotum are occupied by a large brownish black patch which reaches
neither the carina nor the apical margin, is longitudinally divided by an irregular line of
continuous spots and meets the patch of the other side in the centre of the pronotum,
separating a bell-shaped posterior median luteous patch from the luteous apical border.
The lateral carina is rather longer than in the two previous species here described and
the angle replaced by a nearly even curve. The structure of the surface of the pronotum
and elytra agrees with that of H. tbialis, only being a trifle finer. The elytra bear each
two blackish patches, a rounded one on the subbasal callosity, and a very large lateral one
which occupies the limbal margin from the shoulder to the apical declivity, narrows
dorsally, and extends to the second or third stripe.
The pygidium is semicircular, with the basal groove continued as far as the centre or
beyond.
Loc. Praslin: Cétes d’Or Estate, XI. 1908. Silhouette: Mare aux Cochons, VIII.
1908. Type from Praslin.
6. Hormiscops (?) tessellatus, spec. nov.
g. A speciebus antea descriptis differt oculis profundius sinuatis, carina pronoti
basali lateribus valde arcuata haud angulata ad laterum medium continuata.
JORDAN—COLEOPTERA: ANTHRIBIDA 253
Brunneo-rufus, luteo-griseo-pubescens, brunneo-maculatus, antennarum basi, pedibus
atque ore plus minusve luteis ; antennis pronoti basin superantibus, articulo 3'° secundo
fere duplo longiore, 8° longitudine secundi, 9°—11° conicis, parum compressis, 11° apice
breviter acuminato ; fronte rostri dimidio basali latiore; pronoto confertissime ruguloso-
punctato, latitudine paululo breviore; elytris subparallelis, punctato-striatis, pronoto
triente longioribus, striis luteo-griseis fasciis transversis brunneis interruptis; subtus
griseus.
Long. (cap. excl.) 2°2 mm.
One ¢, which is not a true Hormascops.
The upper lobe of the very distinctly sinuate eye is much narrower than the lower
portion of the eye. The segments of the club of the antenna are nearly alike except for
the eleventh segment being acuminate at the tip, while the other two are truncate; they
appear regularly reversed conical in a view of the broader side, 9 being longer than 3.
The occiput bears two rufous patches separated by a luteous grey pubescent median
line, these markings being continued on to the pronotum. The latter is a very little
broader than it is long, being widest posterior to the apex of the lateral carina; the sides
are rounded. The carina is almost evenly curved dorsally, and flexed forward in a wide
are at the sides, reaching nearly to the middle, with the lateral angle hardly at all
indicated. The two rufous apical patches mentioned before are united posteriorly ; there
is a very large brown central patch in front of the carina, enclosing a small grey spot; the
sides bear two pairs of brown spots which nearly form two longitudinal lines.
The elytra are very slightly wider than the pronotum, being one-third longer than it,
their length also surpassing their breadth by nearly one-third. They are luteous grey,
the large spot on the basal callosity of each elytrum being brown; the interspaces 1, 2, 4,
6, 8 are rather more extended rufous than the others, thus luteous grey stripes being
formed which, however, are interrupted by 6 or 7 irregular transverse bands ; the sides of
the elytra are slightly blackish.
The first segment is longer in all the tarsi than the fourth.
Toc. Mahé: Mare aux Cochons district, I. II. 1909.
Epiraputus, Fairm. (1898).
Epitaphius, Fairm., Ann. Soc. Ent. Belg., (1898) xli. p. 493.
The genus is only known from Madagascar and the Continent of Africa.
7. Epitaphius lucheneus, spec. nov. (Plate 15, fig. 3).
a2. Brevis, brunneo-niger, griseo-albo-pubescens nigro-variegatus, antennis pedi-
busque rufis, illis clava, his tibiarum apicibus atque tarsorum articulis 2°—4° nigris.
Pronotum postice parum depressum, carina dorsali recta in medio levissime angulata,
lateribus in angulum obtusum rotundatum sed distinctum antrorsum flexa.
Long. (cap. excl.) 6—7°3 mm.
The black markings are irregular, not sharply defined, more or less confluent, and
somewhat variable. On the pronotum there is a median spot close to the apical margin,
254 PERCY SLADEN TRUST EXPEDITION
and several confluent spots further back at each side of the centre, an irregular stripe
running down to the carima on each side of the disc bounding a posterior grey-white
median patch; a subapical black lateral patch in front of the lateral carina. On the
elytra the black markings form a very irregular net-work, the grey-white pubescence
being predominant ; there is a somewhat larger spot on each elytrum between stripes 1
and 3 in the centre, another below the shoulder and a third before the centre, the last
two being limbal; the subbasal, very feebly convex, callosity bears some streaks or
a patch.
In the g-antenna segment 3 is the longest, 4 to 7 are slightly shorter than 3,
8 is a little shorter than 7 and as long as 9; 9 and 10 bear some longer pubescence
beneath at the apex, but are not pilose. In the 9-antenna segments 3 to 5 slightly
decrease in length, 6 equals 7, and 8 is shorter, being also shorter than 9.
The prothorax is one-third shorter than it is broad and rather strongly convex, being
planate in front of the carina but not impressed. ‘The dorsal carina is practically straight,
not being obviously undulate, and its lateral angle, though being strongly rounded, is
more distinct than usual in this genus. The elytra are less than twice as long as they
are broad, the proportion of length to breadth being 8 : 5 in the type-specimen (2).
Three $f and two 22.
Loc. Silhouette: Mare aux Cochons, plateau and forest above it, IX. 1908. Mahé:
Cascade Estate, 800—1500 ft. Type from Silhouette.
Puiaosius, Schénh. (1826).
Phleobius, Schénh., Disp. Meth., p. 36 (1826).
An Old World genus, not represented in America and most abundantly developed in
the Ethiopian Region. With the exception of a few forms with conspicuous markings
the species are difficult to distinguish. The pattern being in many instances almost the
same, one has to rely on the details of structure in discriminating between the different
species. However, the width and structure of the frons, the size of the eyes, the shape
of the antenna, pronotum, pygidium and prosternum, the shape and the structure of
the elytra and legs frequently offer easily recognised distinguishing characters. Some
of the earlier species are still a puzzle to me, the descriptions not being sufficiently precise,
and there exist in collections quite a number of undescribed species. The genus can con-
veniently be divided into the following sections :
A. Frons with median groove bounded by two low ridges, or rugate-punctate
without groove :
(a) Third segment of tarsi large-——Here belong P. alternans, Wied. (1819),
P. albomaculatus, Allard (1895), P. albescens, Jord. (1895), and some undescribed forms.
(b) Third segment of tarsi small, posterior angle of pleura of posterior abdominal
segments projecting. —Here belong most species: P. gigas, Fabr. (1775), P. nigroungulatus,
Gylh. (1833), P. ceylonicus, Jord. (1902), P. pallipes, Jord. (1895), P. pilipes, Jord. (1895),
P. variegatus, Kolbe (1895), P. cordiger, Fahrs. (1839), P. humilis, Kolbe (1895), etc.
JORDAN—COLEOPTERA: ANTHRIBIDA 255
(c) Third segment of tarsi small, posterior angle of the ante-pygidial pleura not
projecting as a tubercle-—Here belong P. papuanus, Jord. (1904), P. podicalis, Kolbe
(1895), P. affinis, Kolbe (1895), P. hypoxanthus, Jord. (1911), P. catenatus, Kolbe (1895),
P. compressicorms, Gylh. (1833), P. quadricommatus, Qued. (1886), P. striga Jord.
(1904), ete.
B. Frons with narrow median carina, which is sometimes divided longitudinally.—
Here belong P. pustulosus, Gerst. (1871), and allies.
The collection from the Seychelles contains two species, one of which has already
been recorded from these islands.
8. Phleobius gigas cervinus (Klug) (1833) (Plate 15, fig. 14).
Anthribus cervinus, Klug, Abh. Ak. Berlin, 1, p. 188 (1833) (Madagascar).
Anthribus ngroungulatus, Fahrs. (nec Gylh. 1833), in Schénh., Gen. Curc., 5, p. 241,
no. 2 (1839) (partim).
Phieobius nigroungulatus, Lacord., Gen. Col., vii. p. 577 (1866); Gemm. & Har.,
Cat. Col., 9, p. 274 (1872); Fairm., Bull. Soc. Ent. France, p. 324 (1893) (Seychelles) ;
Kolbe, Mitt. Zool. Mus. Berlin, 5, p. 39 (1910) (Mahé).
Several specimens of both sexes.
Loc. Mahé: Cascade Estate, 800—1000 ft., 1908—9. likewise collected in the
Seychelles by J. de Gaye and J. J. Lister, no special localities being given with the
specimens.
Also known to me from Mauritius, Bourbon and Madagascar.
This form is not the same as P. nigroungulatus Gylh. (1833). Through the kindness
of Professor Y. Sjéstedt, I have been enabled to examine the type-specimen of that insect,
a 2 from Canton, as well as a ¢ from Madagascar referred by Fahreens in 1839 to
nigroungulatus. The ? is an immature specimen. Its legs are very pale, but there is
nevertheless a vestige of a fuscous patch at the base and beyond the centre of the tibiz.
The elytrum bears on the basal swelling four black spots, of which the anterior one of the
inner pair is the largest, its pubescence being slightly prolonged so as almost to form
a tuft. The black spots along the suture and in the apical half of the third, fifth and
seventh interspaces are also conspicuous, while the declivous apical area is uniformly
buffish grey and sharply defined. We have specimens from Japan which agree with the
type of nigroungulatus except in the derm being darker (mature examples). As far as
I can judge from the rather scanty material I have seen of true negroungulatus, I am
inclined to regard this insect as a western form of Phlaobius gigas, and cervinus as the
Mascarene subspecies of the same species.
True gigas, better known under the later name of griseus Fabr. (1792), inhabits the
Papuan Subregion, occurring from Celebes to Australia and New Guinea (inclusive of the
adjacent islands), and is characterised by bolder white markings and by the apex of the
claw-segments of the tarsi and the claws themselves being rufescent instead of black.
The Mascarene subspecies, cervinus Klug, ditters from negrowngulatus in the white
dots on the pronotum as well as the black spots on the elytra being less distinct, in the
SECOND SERIES—ZOOLOGY, VOL. XVI. ; 33
256 PERCY SLADEN TRUST EXPEDITION
pubescence on the subbasal black spots of the elytra not forming a slight tuft, and in the
longitudinal groove of the frons being better defined. The above-mentioned ¢ recorded by
Fahrens belongs to cervinus.
9. Phlaobius pustulosus, Gerst. (1871).
Phleobius pustulosus, Gerst., Arch. Naturg., (1871) p. 76; and in Decken’s Reise
OPAtire mie ps 250) tb. L2tioa salsa):
One small ?, which is purer white and slenderer than any other specimens I have
seen of this species. P. pustulosus bears a raised median line on the frons ; the prothorax
is but slightly narrowed at the apex, the anterior angles being very prominent, and has a
transverse row of five tufts. The elytra also bear tufts. The tibize have three black or
brown spots, one of which is placed at the base.
Loc. Aldabra, 1908 (Fryer).
Occurs also in Hast Africa and in Madagascar.
Contexta, Jord. (1902).
Contexta, Jord., Deutsche Ent. Zeitschr., (1902) p. 78.
An Oriental genus.
10. Contexta murina, Jord. (1901) (Plate 15, figs. 11, 11 a, 110).
Contexta murina, Jord., l.c. (Ceylon).
The specimens from the Seychelles only differ from the few I have from Ceylon in the
club of the antenna being darker.
Ten specimens were obtained on Mahé, which vary in the ground-colour from
rufescent brown to pale rufous.
Loc. Mahé: Cascade Estate, 800—1000 ft. ; Long Island, VII. 1908.
CLERANTHRIBUS, gen. Nov.
$9. Elongatus, nitidus. Rostrum brevissimum. Antennarum foveze apertee. Oculi
ovati integri laterales grosse granulosi. Antenne prothoracis basin haud superantes, clava
distincta, articulis 9° et 10° late triangularibus. Mandibule prope apicem dente acuto
instructee. Prothoracis carina a basi remota, arcuata, lateribus tenuissima ad medium
continuata, angulo laterali distincto. Elytra basi emarginata. Prosternum longum ;
processus intercoxalis mesosternalis angustus. Abdomen cylindricum. Pygidium con-
vexum verticale. Tibize posticze apice supra setis brevibus instructe ; tarsorum articulus
tertius lobis longis angustis.—Genotypus: C. colydiopsis, spec. nov.
The genus has no very near relative. It belongs to the Anocerine, and may be
placed near Choragus. The species here described recall Anthicids and Colydiids by their
facies, one of them also closely resembling a small Clerid which occurs with it.
The rostrum is at least four times as broad as it is long, the sides being entirely
occupied by the mandibles and the antennal grooves; the apical margin is straight with
the angles rounded off. The eyes are quite lateral, prominent but not large, coarsely
JORDAN—COLEOPTERA: ANTHRIBIDA 257
granulated, transversely ovate, bemg widest above. The antennal fovea is smaller than
the eye, which it touches, triangular, rounded dorsally, the dorsal edge being rather more
dorsal than the inner edge of the eye. The labiophore, on the under side, is flat, apically
only one-fifth wider than proximally, with the lobes rounded off. The mandibles bear
a tooth near the apex. The distal part of the labium is broad, non-divided, being broader
than the mentum; the latter is exposed and on a level with the under surface of the
labiophore.
The antenna is slightly longer in the g than in the ?, and does not reach the base of
the prothorax in either sex; segments 1 and 2 are incrassate and of about the same
length, 2 being somewhat claviform, 3 is slender, slightly shorter than 2 and as long as 5,
4 being a little shorter, 5—8 are thicker, 8 being the shortest of them; the club is
moderately compressed, 9 and 10 triangular, asymmetrical, 9 surpassing 3 somewhat in
length, 10 broader than long, 11 rounded.
The prothorax is much longer in the $ than in the $, cylindrical above, with the base
constricted, the dise:swollen and the apical margin projecting as a hood on to the head.
The carina is remote from the base at the sides and gradually incurved, being nearest to
the base in the centre; its lateral angle is distinct and projects a little as a tooth in
a dorsal aspect ; below this projection the carina is bent backwards a very little and then
runs forward almost to the centre of the sides, this lateral branch being less elevated than
the dorsal carina. The short basal longitudinal carina is vestigial and joins the dorsal
carina in an acute angle. The elytra are emarginate from shoulder to shoulder, and bear
rows of minute punctures, not stripes, the punctures being more distinct in the basal area;
the suture is not depressed, but the basal callosities and the depression behind them are
marked. The pygidium reminds one of Basitropzis, inclining forward in the ¢, and being
vertical in the ?.
The prosternum is about as long as the head and rostrum together, the antecoxal
portion being remarkably long. The mesosternal process is narrower than the intercoxal
process of the cylindrical abdomen. The legs do not differ much in size among themselves,
and also do not exhibit any obvious sexual difference. The hind tibia bears on the
slanting dorsal apical surface a number of bristles. The first tarsal segment is as long as
the claw-segment or a little longer ; the teeth of the claws are long.
The underwings (of C. colydiopsis) have some characteristics which are apparently
only found in the Anocerinz. The portion from the apex of the cell outward is half as
long again as the proximal portion, 7.e. the cell reaches to two-fifths of the wing. The
anal area is strongly reduced, as are also the anal veins, and there are no veins between
the cell and the anterior anal vein (which usually reaches down to the base in Anthribide,
but does not do so in the present genus). Moreover, the recurrent vein starting from the
lower cell-angle is very short and feebly developed. The edge of the wing bears a fringe
consisting of bristles placed at regular intervals and being at least six times as long as the
hairs situated on the surface of the wing; the fringe-bristles of the anal lobe are three
times as long as those of the apical portion of the outer margin.
33—2
258 PERCY SLADEN TRUST EXPEDITION
11. Cleranthribus colydiopsis, spec. nov. (Plate 15, fig. 6).
é?. Niger, nitidus, pilis sparsis vestitus, pube alba signatus, ex parte rufus, capite
dense rugato-punctato, pronoto grosse punctato, fascia subapicali atque stria brevi
mediana subbasali albis notato, elytris pone basin rufis fascia transversa alba ornatis.
Long. (cap. excl.) 1°3—2°5 mm.
The specimens vary considerably in size and also to some extent in markings, but the
long series of examples obtained contains individuals which are intermediate. Moreover,
the specimens which differ in size or colour, or in both, are apparently alike in structure.
The whole body is glossy black, the mouth-parts, the antennee (usually with the
exception of the club), the bases of the pronotum and elytra, the postbasal depression and
frequently a postmedian sublateral patch of the elytra, and the legs are more or less bright
rufous. A subapical transverse band on the pronotum and a band situated in the
depression of the elytra as well as a short median line in front of the carina, a lateral spot
touching the carina, a sutural spot behind the scutellum, the epimerum of the mesosternite
and the episternum of the metasternite (as well as part of the hind edge of the meta-
sternum) are pubescent-white, the head and the rufescent posterior portion of the elytra
also bearing white pubescence in well-preserved specimens. This pubescence consists of
broad flat hairs.
Sometimes the greater portion of the elytra is rufous, only a sutural stripe or patch
remaining blackish brown. In specimens with a distinct white subapical band on the
pronotum, this portion of the derm is also rufescent (e.g. in the type-specimen).
The prothorax is one-third ($) or one-sixth (¢) longer than it is broad, being widest a
little before the centre, the sides being very slightly rounded. The dise bears dispersed
punctures, which are rather large, and the sides, particularly ventrally, are densely and
coarsely punctured. The elytra are three-fifths as broad as they are long, being slightly
broader than the prothorax. They are sharply marginate at the base and around the
shoulders. In one of the $3 the apical area of the elytra is somewhat flattened, the centre
of each elytrum being in this place longitudinally swollen, an obtuse smooth crease being
formed.
Loc. Mahé: Mare aux Cochons, I.—II. 1909; country above Port Glaud, 500—
1000 ft., XI. 1908 ; high forest of Morne Blanc and Pilot, XI. 1908 ; near Morne Blane ; high
damp forest between Trois Fréres and Morne Seychellois, about 1500—2000 ft., XII. 1908 ;
from grass in cultivated country, about 1000 ft., XII. 1908; forest above Cascade Estate,
I. and IL. 1909; also in low country ; top of Mount Sebert, nearly 2000 ft., I. 1909;
forest of stunted Capucin trees (Northea) on summit of “ Montagne Anse Major,” 2000 ft.
or over, II. 1909; Cascade Estate, about 1000 ft., II. and III. 1909; jungle between
Trois Fréres and Morne Seychellois, 1500—2000 ft., XII. 1908. Silhouette: high country
near Mont Pot-a-eau, VIII. 1908; Mare aux Cochons plateau and jungle near by, VIII.
and IX. 1908.
Many of the specimens were obtained in the forests by beating. Along with these
occurred frequently a Clerid beetle of the same size and colour.
JORDAN—COLEOPTERA: ANTHRIBIDA 259
12. Cleranthribus anthicopsis, spec. nov. (Plate 15, figs. 7, 7 a, 7 b).
g. Rufus, prothoracis medio elytrorumque macula transversa antemediana suturam
non attingente et plaga apicali nigro-brunneis, rostro carina mediana instructo, prothorace
magis convexo quam in C. colydiopsi, omnino sparsis pilis vestitus, absque fasciis et maculis
albis, lateribus meso- et metasterni exceptis.
Long. (cap. excl.) 2°6 mm.
Only 1 ¢, which has the facies of an Anthicus on account of the prothorax being more
swollen than in C. colydiopsis.
The antenna, which is entirely pale rufous, is rather longer than in the f¢ of the
previous species, particularly segments 9 and 10, 9 being about one-third longer than it is
broad. The pronotum is strongly convex about the centre, rounded at the sides, gradually
narrowing from the middle to the apex. The lateral carima is somewhat longer than in
colydiopsis. The elytra also are rather more rounded and somewhat broader than in the
other species. There are no white markings on the upper side, and the dispersed hairs are
rather longer than in colydiopsis, particularly in the neighbourhood of the scutellum. The
postbasal depression, which in colydiopsis is rufous and bears a white band, is partly
occupied in anthicopsis by a large brown spot, which is widest laterally, but reaches neither
the suture nor the lateral margin; the brown apical area occupies about two-fifths of the
elytra, but is not sharply defined and presumably varies individually. The legs are longer
than in colydiopsis, the first tarsal segment being longer than the claw-segment.
Loc. Mahé: Mare aux Cochons district, I.—II. 1909.
ScIRTETINUS, gen. nov.
a2. Generibus Choragus Kirby (1818), Acarodes Woll. (1879) atque Holostilpna
Jord. (1907) dictis affinis. Nitens, absque pubescentia. Antennze setis longis vestite,
clavee articulis valde remotis. Pronoti carina antebasalis basi parallela, lateribus ad
medium usque continuata. Tibiz quatuor posticze supra ad apicem excise vel oblique
truncate ac ibi setis brevibus vestitee.—Genotypus: S. ewmelas, spec. nov.
The body is as strongly convex as in the allied genera, particularly Acarodes and
Holostilpna, the dorsal outline from the apex of the pronotum to the tip of the pygidium
being almost evenly curved except for a small interval between the antebasal carina of the
pronotum and the basal edge of the elytra. The species are glossy black or brown without
any pubescence. The head can be retracted into the prothorax, in fact, it is only exserted
in a few specimens of the collection, as a rule only the antennze and the tip of the
mandibles projecting from the cavity of the thorax. The skeleton is very firm, especially
the under surface.
The mandible bears a tooth near the apex. The fourth segment of the maxillary
palpus is as long as the preceding segments taken together, but much slenderer, being
elongate-conical with the tip truncate. The labial palpi have a very slender end-segment,
while the other two segments are rather stout. The labium is entire, its apex only being
feebly sinuate.
260 PERCY SLADEN TRUST EXPEDITION
As in the allied genera segments 1 and 2 of the antenne are incrassate, 2 being
clavate, whereas 3—8 are slender. 2 is about as long as 3 and 4 together. The club is
flattened and very distinct, 9 and 10 being almost alike, asymmetrical, with one side more
rounded than the other and the apex as well as the base so narrowed as to form a pedicel ;
segment 11, which is elliptical, also has a narrow base, the club being very loose, much
more so than in Acarodes and Holostilpna. The g-antenna is somewhat longer than that
of the $ and reaches beyond the base of the pronotum.
The antebasal carina of the prothorax is parallel to the base, and is curved forward at
the sides in a slightly obtuse angle, the lateral branch reaching to the middle of the sides.
The short basal longitudinal carina connects the angle of the antebasal carina with the
true basal angle of the prothorax. The tip of the lateral carina is slightly curved upwards,
which recalls Acaromimus Jord. (1907).
The elytra are rather strongly seriate-punctate, the interspaces bearing a row of very
minute punctures.
The legs are short and stout, particularly the femora. The hind tibia, and to a lesser
extent also the mid tibia, is deplanate or excised on the upper side at the apex, this space
being studded with short erect bristles. The fore tibia, especially in the ¢, is somewhat
longer than the other tibiz. The first tarsal segment is about as long as the fourth, or at
any rate not essentially longer. The third segment is broad and the tooth of the claws
distinct.
The species, which have a considerable power of jumping as in the allied genera, differ
from Choragus, particularly in the absence of pubescence from the body, the glossy derm,
the much smoother pronotum, the tooth near the apex of the mandibles, ete. Holostilpna
has two lateral longitudinal carinee separated by a groove, and in Acarodes, Acaromumus
and Xenorohestes, which contain glossy black species, the prothoracic carina is basal, not
antebasal as in Choragus, Scirtetenes and Holostilpna.
18. Scirtetinus eumelas, spec. nov. (Plate 15, fig. 8).
g¢. Ater, nitidus, ore, antennarum basi, tarsis atque tibiarum dimidio apicali plus
minusve luteis, capite pronotoque lineis impressis tenuissimis rete regulare formantibus
instructis, angulo laterali carinee subrecto non rotundato, elytris seriatim punctatis.
Long. (cap. excl.) 0°9—1'7 mm.
A series. The specimens vary in the extent of the luteous tint, particularly of the
tibiee, which are sometimes entirely of that colour. The outline of the insect in a dorsal
aspect is elliptical, but the contour is bent in at the juncture of prothorax and elytra.
In a lateral view the pronotum and elytra are about double the height of the sterna and
abdomen.
The most striking feature of the species is the regular network of engraved lines
which cover the head and pronotum, the lines forming rather large hexagonal meshes, the
sides of which, however, are not of equal length, some meshes being even pentagonal. As
each mesh has a single distinct puncture, the net bears a remarkable likeness to a layer
of cells, the punctures representing the nuclei.
The carina is very feebly curved backward in the centre; the lateral angle is not
JORDAN—COLEOPTERA: ANTHRIBIDA 261
rounded and only a little larger than 90°. The elytra are slightly more than half as long
again as the pronotum. They bear rows of rather large punctures, but only the sutural
row is distinctly impressed as a stripe.
The under surface of the thorax bears large dispersed punctures.
The underwing is reduced, being very narrow, but variable in length. It bears only
two veins, which are vestigial and the hind margin is dilated beyond the middle.
Loc. Mahé: country above Port Glaud, 500—1000 ft., XI. 1908; Cascade Estate,
1000 ft., IL—III. 1909; Mare aux Cochons, 1000—2000 ft., I.—IT. 1908. Silhouette:
Mont Pot-a-eau, 1500 ft., VIII. 1908; Mare aux Cochons, VIII.—IX. 1908. Praslin :
Cotes d’Or Estate, XI. 1908. Félicité: from forest, XII. 1908. Type from Mahé.
14. Scirtetinus dinudiatus, spec. nov. (Plate 15, fig. 12).
d¢. Ater, nitidus, subtus picescens, ore et antennarum basi luteis, tarsis pices-
centibus ; pronoto cum capite densissime coriaceo, punctis sat magnis leviter impressis
numerosis omnino instructo, elytris pronoto paululo longioribus et angustioribus, grosse
punctato-striatis, tibiis posticis compressis latis.
Long. (cap. excl.) 1—1°2 mm.
A small series.
The very dense leathery structure of the surface of the head and pronotum renders
these parts less shiny black than are the elytra ; both bear numerous evenly distributed,
rather large but shallow punctures, which are much smaller than the punctures of
the under side of the thorax. The carina forms a slightly obtuse angle, which is not
rounded. The thorax is about one-tenth shorter and also one-tenth broader than the
elytra, as well as one-tenth broader than it is long. The elytra are less convex
than in S. ewmelas, their vertical width in a lateral aspect not surpassing that of the
sterna; they are coarsely punctate-striate and very glossy. The hind tibia equals in
width about one and a half interspaces of the elytra (measured near base).
Underwings very small.
Loc. Mahé: slopes of Morne Seychellois, 1500—2000 ft., Il. 1909; forest behind
Trois Fréres, 2000 ft., I. 1909; Cascade Estate, 800—1500 ft., III. 1909; high damp
forest between Trois Freres and Morne Seychellois, 1500—2000 ft., XII. 1908.
15. Scortetinus luteipes, spec. nov.
39. S. dimidiato valde affinis, colore luteo magis extenso, tiblis pro maxima
parte et totis tarsis luteis, tibuis posticis minus dilatatis,
A small series.
The proportions of the thorax and elytra are the same as in S. dimidiatus. The
antennz are pale yellow with brown club, the entire fore and mid tibie, the apex of
the hind tibia and all the tarsi are luteous. The punctures of the pronotum are rather
larger towards the sides, and therefore do not contrast so much in the neighbourhood
of the lateral carina with the large deep punctures of the under surface. The hind tibia
is not nearly so broad as in dimediatus.
262 PERCY SLADEN TRUST EXPEDITION
Loc. Mahé: Cascade Estate, 800—1500 ft.; forest near Mt. Harrison, 1700 ft.,
UL, USO2)
16. Scirtetinus piceus, spec. nov.
g. Duabus speciebus przecedentibus simillimus, antennis pedibusque luteis, femori-
bus plus minusve piceis, pronoto grossius punctato, elytris profundius striatis, pronoto
parum latioribus atque dimidio longioribus.
Long. (cap. excl.) 1°2 mm.
Two #¢.
The hind tibia bears a small tubercle in front of the apical setose excision. The
proportions of the thorax and elytra are quite different from those observed in the
two previous species, the elytra being distinctly though slightly broader than, and
half as long again as, the pronotum. I have examined one of the specimens as to
the underwings and found them to be more than twice as long as the whole specimen,
The proximal area of the wing to the main knee (apex of cell) is narrow and only
half the length of the much broader outer portion; the anal lobes are very strongly
reduced and there are no distinct anal veins.
Toc. Praslin: jungle on the Cédtes d'Or Estate, XI. 1908, from leaf-bases of
Pandanus. Mahé: Cascade Estate, 800 ft.—Type from Praslin.
A much larger 9, length 1:7 mm., may also belong here. It was found in the
head of a Pandanus Horner in Silhouette, 22. IX. 1908.
Cuoracus, Kirby (1818).
Choragus, Kirby, Trans. Linn. Soc. London, xii. (1818) p. 447 (type: C. sheppardt).
A cosmopolitan genus. A number of species have been placed into this genus
which do not belong there. In true Choragus the prothorax and elytra bear at least
some vestiges of pubescence; the eyes are lateral, the prothoracic carina antebasal and
laterally curved forward, but does not reach beyond the centre of the sides. The
head and pronotum are coriaceous and punctate.
17. Choragus ornatus, spec. nov. (Plate 15, fig. 13).
Nigro-brunneus, picescens, antennarum basi pedibusque luteo-rufis, utroque elytro
in dorso maculis duabus magnis luteis una basali altera subapicali ornatis ; cylindricus,
Sparsissume pubescens.
Long. (cap. excl.) 2 mm.
One specimen, probably a ¢.
The second segment of the antenna is the longest of all, being clavate and
more than twice as long as segment 3, 5 to 8 decrease in length, 8 being less than
twice as long as it is broad; the club is broad, hairy, 9 triangular, as broad as it 1S
long, 10 rounded proximally, subemarginate distally, twice as broad as long, 11 rounded,
narrowing at the apex, slightly longer than broad.
JORDAN—COLEOPTERA: ANTHRIBIDA 263
The prothorax and elytra are almost perfectly cylindrical. The former is slightly
wider than long and rounded-angustate from the middle to the apex, the whole upper
surface very densely reticulate-punctate; the carina is dorsally almost straight, being
very faintly incurved thrice, its lateral angle is obtuse, but not rounded off, and the
lateral carina does not extend to the centre of the sides.
The elytra are not broader than the prothorax, but more than half as long again.
They are clothed, like the pronotum, with a very sparse pubescence which is only
visible under a strong lens, and regularly punctate-striate, the punctures being large ;
the base of the sutural interspace is much widened and bears a short stripe of punctures.
The anterior luteous patch expands between stripes 2 and 8 and reaches the basal
margin between 4 and 8, whereas the subapical patch expands between stripes 1 and 8,
the suture being here pitchy.
The prosternum is as convex as the pronotum and has the same surface-structure.
Loc. Mahé: Cascade Estate, 1000 ft., IJ.—IIT. 1909.
18. Choragus bolus, spec. nov.
$%. Piceo-brunneus, sparsissime pubescens, antennarum basi tibiis tarsisque
pallidioribus, capite et pronoto confertissime rugato-punctatis, subreticulatis, angulo
carine rotundato, elytris pronoto duplo fere longioribus, illo quoque latioribus, ad
apicem usque grosse striato-punctatis, pygidio basi fovea media instructo.
Long. (cap. excl) 2—2°5 mm.
One ¢# and one 3.
The present insect is clothed with short stiff appressed hairs, which, however, do
not conceal the derm anywhere and are not condensed into markings. The head is
coarsely rugate-punctate. The pronotum is minutely coriaceous and very densely
covered with shallow punctiform pits which form a net; it is a little broader than
it is long, with the sides rotundate in the centre, narrowed towards the apex and
faintly incurved in front of the carina; the disc is slightly depressed before the carina.
The angle of the carina is rounded, and the lateral carina does not reach to the
middle of the sides. The elytra are more strongly convex than the pronotum, and
almost twice as long, being broadest before the centre and thence to apex nearly
evenly rotundate-angustate. The punctures of the stripes are very large and deep.
The pygidium bears a smooth groove at the base, as a continuation of the central
groove of the propygidium. The sterna are studded with large deep punctures and
the abdominal segments bear a row of similar punctures at their bases. The tarsal
segments 2 and 3 are narrower than in any species of Scortetinus.
A second specimen, a f with the antennz missing except for the basal segments,
doubtless belongs to this species. It is larger than the 3, being 2°5 mm. long. The
pygidium, which is granulose in the ?, is more rugose in the §. The abdomen of
this ¢ is medianly flattened and the last sternite distinctly swollen at each side.
The underwings are large in both specimens. The anal area is narrower than
the apex of the cell; there is a small basal anal lobe separated from the anal area
SECOND SERIES—ZOOLOGY, VOL. XVI. 34
264 PERCY SLADEN TRUST EXPEDITION
by a sinus as in the allied Anthribide. The main anal vein and the veins forming
the cell are well developed. As in the species of Scirtetinus there are also here no
veins between the cell and the apex of the anal vein.
Loc. Silhouette: high forest above Mare aux Cochons, IX. 1908; near Mont
Pot-a-eau, VIII. 1908.
CoRYNACIA, gen. nov.
g. Cylindricus; antennarum fovea subdorsali prope oculum sita; antennis brevibus,
articulis 1° et 2° incrassatis, 3!°—9° brevibus, 3° longiore, 9° octavo parum crassiore,
10° nono multo latiore, 11° decimo latiore et 2°—5° simul sumptis longitudine eequali ;
oculis rotundis lateralibus ; carina pronoti antebasali lateribus ad apicem continuata,
elytrorum basi emarginata ; pedibus brevibus, articulo 1° quarto breviore, 3'° angusto.—
Genotypus: C. scotti, spec. nov.
As the collection contains only a single specimen of this interesting genus, I have
abstained from relaxing it for more minute examination. The structure of the antenna
is so peculiar that there will be no difficulty in recognising the insect.
The first two antennal segments are incrassate as usual in this group of genera,
segment 3 is less than half the length of 2, but distinctly longer than 4, 4 to 6 are
nearly alike, 7 and 8 are shorter, a little thicker and about as long as they are broad,
9 resembles 8 except for being thicker, the club being formed by 10 and 11, which
are rather hairy and somewhat flattened; 10 is rounded-triangular, bemg as broad
as it is long, with the base nearly as broad as segment 9, 11 is three times as long
as 10 and equals in length segments 2 to 5 taken together, being distinctly curved
and of nearly even width throughout with the apex rotundate-angustate.
The rostrum is more than four times as long as it is broad, being flat, with a
feeble median depression ; its apical margin is slightly incurved. The antennal fovea
is transverse and touches the eye. The latter is coarsely granulose and circular, with
traces of two angles below. ‘The frons is very broad, being posteriorly wider than
anteriorly.
The prothorax and elytra are long and cylindrical, and bear dispersed short ap-
pressed hairs, which do not conceal the colour of the derm to the slightest extent.
19. Corynecia scottr, spec. nov. (Plate 15, figs. 15, 15a).
Piceo-ater, subnitidus, densissime coriaceus, confeJissime rugato-punctatus, ore,
antennarum funiculo, tibiis tarsisque luteo-rufis, carina prothoracis lateribus valde
arcuata absque angulo laterali, elytris prothoracis latitudine, illo plus dimidio longioribus,
grosse atque densissime striato-punctatis.
Long. (cap. excl.) 2 mm.
One specimen, apparently a @.
The antenna only reaches a little beyond the middle of the prothorax; segments
1 to 5 are pale rufous, 6 to 8 rufous brown, 9 to 11 black. The pronotum is as wide
as it is long, being widest at the apical fourth. It is minutely coriaceous and exceedingly
JORDAN—COLEOPTERA: ANTHRIBIDA 265
densely pitted all over with grooves, which give it a reticulated appearance. The carina
is antebasal, but not very remote from the base, being straight above with a faint
curve backwards in the centre; laterally it is flexed forward in a large curve and
reaches the apical margin at some distance below the eye. The elytra are as broad
as the prothorax and more than half as long again. They are cylindrical, coarsely
rugate and punctate-striate, and bear distinct humeral callosities. In spite of the
coarse structure of the derm, both the upper and under surfaces of the species are
glossy. The first segment is about half the length of the fourth segment in all the tarsi;
the claws have a distinct tooth. ;
Loc. Silhouette: Mare aux Cochons or forest above it, IX. 1908.
ACHORAGUS, gen. nov.
é?. Chorago valde affinis, carina prothoracis basali.—Genotypus : A. tener, spec. nov.
One elongate species.
The mandible bears an acute tooth near the apex. ‘The eye is large, lateral,
coarsely granulose and round. The antenne are slightly longer in the g¢ than in
the ?, segments 1 and 2 are incrassate as usual in this group of genera, 3 to 8 are
almost alike in length, the club is but very slightly compressed, the segments being
separate but not much stalked, 9 and 10 obliquely truncate at the apex, 11 subglobular.
The pronotum is less convex than the elytra; the carina is basal and is continued to
centre of the sides, forming an obtuse lateral angle, this lateral carina is less elevated
than the dorsal one. The elytra are elongate and cylindrical; the suture is not
depressed, the basal margin very slightly imcurved from shoulder to shoulder, being
almost straight. Legs as in Choragus; hind tibia with some stiff bristles dorsally at
the apex. The teeth of the claws long and slender. The prosternum is short; the
antecoxal portion being scarcely as long as the diameter of the eye.
20. Achoragus tener, spec. nov. (Plate 15, fig. 9).
é?. Rufo-brunneus, ore, antennarum funiculo, tibiis tarsisque luteis, pube alba
vestitus, sutura nigricante, maculis rufo-brunneis seminudis notatus; prothorace den-
sissime punctato-reticulato, elytris sat dense punctulatis.
Long. (cap. excl.) 1°6—2-2 mm.
A small series.
The white pubescence consisting of rather broad flat appressed hairs is not very
dense anywhere, but nevertheless rather conspicuous on account of its colour. The
brown patches, which vary in extent, are as follows: a very large central patch on
the pronotum, reaching from near the apical margin to the carina and divided medianly
by a white ill-defined stripe, which is interrupted in some specimens, at the sides
often a smaller patch; a very large dorsal area on the two elytra together, rounded,
ill-defined, enclosing on each elytrum a white postmedian spot ; moreover, each elytrum
34—2
266 PERCY SLADEN TRUST EXPEDITION
bears a dorsal basal brown patch often connected with the central area, and frequently
also a lateral spot (cf fig. 9). All these markings are much more in evidence when
examined with a low power than under a strong lens.
The prothorax is as long as it is broad, being widest at the angle of the carina,
which projects a little laterally, and is rounded-angustate forward. It bears the very
dense network of shallow punctures also found in Choragus Kirby (1818). The elytra
are one-sixth broader than and twice as long as the pronotum, and rough with irregular
puncturation ; some larger punctures along the suture form a row which is slightly
impressed, there being no other stripes. The base of the elytra is somewhat depressed.
The pygidium is different in the sexes. It is almost square in the @, large, one-third
broader than long, vertical, with the angles rounded off; in the ¢ it is proximally
twice as broad as it is long, triangular, with the sides less rounded than the apical
margin.
The prosternum is punctured, whereas the rest of the under side is minutely
coriaceous. The eighth tergite of the g (beneath the pygidium) is divided into two
narrow convergent lobes of nearly even width and with rounded tips.
Loc. Mahé: country above Port Glaud, 500—1000 ft., XI. 1908 ; top of Mt. Sebert,
2000 ft., I. 1909; Mare aux Cochons, I.—II. 1909. Silhouette: near Mont Pot-d-eau,
1000 ft., VIII. 1908; Mare aux Cochons, IX. 1908.
Dysnos, Pasc. (1859).
Dysnos, Pase., Ann. Mag. Nat. Hist., ser. 3, iv. (1859) p. 488.
The eyes are large and converge posteriorly, the frons being narrower behind
than anteriorly. The antennze are slender, bear long bristles and have a very loose
club. The basal angles of the pronotum are produced backwards, the short longitudinal
basal lateral carina forming an obtuse angle with the dorsal carina.
The genus is apparently cosmopolitan in the tropics, being known from the
Oriental and Nearctic Regions as well as the Seychelles.
21. Dysnos ethiops, spec. nov. (Plate 15, fig. 10).
$¢. Supra niger, opacus, subtus rufescens, ore, antennarum basi, coxis, tarsorum
apicibus plus minusve rufis, pube griseo-aurea sparsissime vestitus ; pronoto creberrime
ruguloso-punctato, elytris antice valde convexis, confertissime granulosis punctato-striatis,
callo subbasali sat distincto.
Long. (cap. excl.) 3°6 mm.
Four specimens.
The frons is posteriorly half as wide as the rostrum. The prothorax is half as
broad again as wide, slightly narrowing from the base to the centre and thence much
more strongly to the apex, which is barely more than half the width of the base.
The elytra are half as long again as, but not broader than, the prothorax, the sub-
basal dorsal and humeral callosities being distinct. The pygidium is triangular, with
the apex rounded and the sides nearly straight in g, and one-third broader than it
PERCY SLADEN TRUST EXPEDITION. TRANS. LINN. Soc. SER. 2.Z00L Vou. XVI Pu 15.
(JORDAN)
E.Wilson del. University Press, Cambridge.
COLEOPTERA : ANTHRIBIDA,.
JORDAN—COLEOPTERA: ANTHRIBIDA 267
is long. The under side is coarsely punctured ; the puncturation is much sparser on
the sides of the first three abdominal segments than on the remainder of the abdomen ;
the last sternite bears a short apical carina in one of the examples (type). The first
segment of the tarsi is only a trifle longer than the fourth.
The golden pubescence of the upper side is condensed in small ill-defined spots.
Loc. Mahé: Mare aux Cochons, from marsh at top of pass, 2000 ft., I.—IT. 1909 ;
Cascade Estate, 800—1500 ft., from a fungus. Silhouette: high country near Mont
Pot-a-eau, VIII. 1908; from fungus. Type from Mahé.
Armcerrus, Schénh. (1833).
Anthribus (subgen. Arecerus), Schonh., Disp. Meth., p. 40 (1827) (type: A. fascicu-
latus, Degeer).
Arecerus, Schénh., Gen. Curc., i. p. 173 (1833).
Areocerus, id., l.c., 5, p. 273 (1839).
An essentially Oriental genus.
22. Arecerus fasciculatus (Degeer) (1775).
Curculio fasciculatus, Degeer, Mém. Ins., v. p. 276, t. 16, f. 2 (1775).
Throughout the tropics.
Loc. Seychelles: Mahé; Cascade Estate, 800—1000 ft., II. 1909 ; Anonyme Island,
I. 1909; forest above Cascade Estate, I. 1909; Long Island, VII. 1908. Silhouette ;
near Mont Pot-a-eau, VIII. 1908 ; Mare aux Cochons, VIIJ.—IX. 1908. Praslin; Cotes
dOr Estate, XI. 1908. Cargados Carajos Islands: Siren Island, 27. VIII. 1905;
Establishment Island, 28. VIII. 1905.
The four specimens from the Cargados Carajos Islands have no brown spots on
the tibize.
EXPLANATION OF PLATE 165.
Fig. Te Sintorops alleus, spec. nov., §/, p. 250.
Mig. 2. BS ie spec. nov., 2, p. 250.
Fig. 3. Epitaphius licheneus, spec. nov., p. 253.
Fig. 4. Hormiscops tibialis, spec. nov., a, antenna; 6, head, p. 251.
Fig. 5. 5 sobrinus, spec. nov., p. 251.
Fig. 6. Cleranthribus colydiopsis, spec. nov., p. 258.
Fig. 7. : anthicopsis, spec. nov., a, head; b, antenna, p. 259.
Fig. 8. Scirtetinus ewmelas, spec. nov., p. 260.
9.
Achoragus tener, spec. nov., p. 265.
Fig. 10. Dysnos cethiops, spec. nov., p. 266.
Fig. 11. Conteata murina, Jord., a, antenna; b, head, p. 256.
Fig. 12. Scirtetinus dimidiatus, spec. nov., p. 261.
Fig. 13. Choragus ornatus, spec. nov., p. 262.
Fig. 14. Phiewobius gigas cervinus, Klug, p. 255.
Fig. 15. Coryncecia scotti, spec. nov., a, antenna, p. 264.
‘
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No. XIV.—MARINE ALGA, RHODOPHYCEZ.
By Mrs. A. WEBER-VAN Bossz, Pu.D.
(Plates 16-18 and 1 text-figure.)
(CoMMUNICATED BY PROFESSOR STANLEY J. GarpinER, M.A., F.R.S., F.L.S.)
Read 5th June, 1913.
[Reprinted from Trans. Linn. Soc., Ser. 2. Botany, Vol. viii. pp. 105-142, pls. 12-14. ]
I. Introduction.
THE algze treated of in the following paper were collected by Mr. J. Stanley Gardiner
during the Perey Sladen Trust Expedition to the Indian Ocean in 1905. It was at first
intended that Mrs. A. Gepp should work out the collection, but she handed it over
to me, and illness prevented her joining in the work, which we would only have been too
glad to carry out together. She had, however, sorted the material before sending it
to me.
The collection does not contain all the Rhodophycee collected by Mr. Stanley Gardiner ;
the Lithothamnia have been worked out by the much lamented Mr. Foslie. The literature
on the algze of this part of the Indian Ocean is scarce, for besides the paper by Mr. Foslie
and the Report on the Chlorophyceze and Pheophyceee of the ‘‘Sealark”’ Expedition by
A. Gepp, M.A., F.L.S., and Mrs. E. Gepp, I only know of one paper in which more than
a few algze of this region are mentioned. Mr. Reinbold, in “ Meeresalgen der Deutschen
Tiefsee-Expedition,” mentions 26 species of Rhodophycez from Diego Garcia and Mahé,
10 of which have been also collected by Mr. Stanley Gardiner. Besides this paper of
Mr. Reinbold, I have found two other short references to algee from Diego Garcia
and the Seychelles. Mr. W. B. Hemsley adds a note on Alge to his “ Report on the
Vegetation of Diego Garcia.” Of the alge named only Dasya indica, J. Ag., has any
interest for us at present; it is the only red alga mentioned. A note by “J. B.” in
“ Knowledge” treats only of Myxophycese, and may therefore be passed over in silence.
The “Sealark ’”? collection had special interest for me because I am just publishing,
with the aid of Mr. Reinbold, the first part of a list of alge collected during the
“ Siboga” Expedition in the Malay Archipelago. This part contains the Myxophycee,
Chlorophyceze, and the Phzeophyceze; the second part, containing the Rhodophyce,
will, I hope, follow soon. The study of a region so near to the Malay Archipelago
afforded me great interest from a phyto-geographical standpoint. It had, however,
one drawback ; some species found by me in the “Siboga” collecticn and recognized as
new are also found in the Stanley Gardiner collection, and the fact that the paper on
the algze of the Indian Ocean will appear before the ‘“Siboga” paper, will diminish
the number of novelties in the latter. But this could not be avoided, and I have
described the new species as fully as I could; only in one case, when the barren,
small fragment could not possibly have been recognized apart from the fertile
specimens of the ‘‘Siboga” Expedition, have I referred to the forthcoming paper.
SECOND SERIES—ZOOLOGY, VOL. XVI. 35
270 PERCY SLADEN TRUST EXPEDITION
Amongst the alge of the Indian Ocean J found species of two new genera, Tapeino-
dasya and Oligocladus, described for the first time from the Malay Archipelago in ‘‘ Les
Annales du Jardin botanique de Buitenzorg,” vol.ix.1910. It afforded me great pleasure
to describe these species, which confirm the validity of my new genera. One genus,
Pseudendosiphonia, is new in the present collection; it is most closely allied to Hndo-
siphonia, and yet is so different that it can be easily distinguished ; one species, Dasya
indica, had to be described as a new genus—Amphisbetema.
Many of the algze are barren, and can therefore not be specifically named; others are
only fragments. I have given the generic names of all, because I think that, for our
knowledge of the geographical distribution, even a generic name may have some value.
The collection is rich in novelties; I often hesitated before describing a given alga as
new, but when, after careful investigation, I could not -identify it with any existing
diagnosis, I felt I had no alternative. Of the novelties, which include one genus,
17 species, one variety and one form, 11 are from deep water, 2 from reef and deep
water, 7 from reef alone. It may be well to recall here that Mr. and Mrs. Gepp
found Avraimvillea amadelpha in material from reef and deep water. Many algze appear
to accommodate themselves to both modes of living. The ‘“Sealark” Expedition had,
like the ‘“ Siboga” Expedition, the advantage of the use of a dredge, and several hauls
from deep water, where many Rhodophycez flourish, have given a rich harvest. The
nature of the bottom surrounding the reefs and islands, whence these algee come, has
been treated in detail in the above-mentioned paper by Mr. Foslie. I think it therefore
unnecessary to do this again.
The geographical distribution shows that a great resemblance exists between the
algological flora of the Indian Ocean and that of the Malay Archipelago as well as that
of the east coast of Africa. Of the 26 species found at Mahé and Diego Garcia by the
“* Deutsche Tiefsee-Expedition,” 7 occur also at Dar-es-Salaam, and of the 8 other
species collected at Dar-es-Salaam by that expedition 5 are found in the Stanley
Gardiner collection *. But it is a puzzle to know how Cladhymenia oblongifolia and
Calliblepharis prolifera, only known hitherto from New Zealand, and Chondria pusilla,
only known from the West Indies, came to be inhabitants of the present region. It
emphasizes the fact that the geographical distribution of algze is a factor that should
always be very carefully considered. The finding of Peyssonnelia Harveyana, known
only from the Atlantic Ocean at Brest, puzzled me less, because it may be possible that
the determination of the barren plant is erroneous. I have therefore queried it.
I have given a list of the localities with the algze found at each, and a general list of
all the Rhodophycez collected by Mr. Stanley Gardiner. The systematic observations
on the different genera and species conclude the paper.
I wish here to express my sincere thanks to Mr. Stanley Gardiner for entrusting me
with his collection, and to Messrs. Gepp and Cotton for the kind help they have given
me; and I may perhaps add a word of deep-felt regret that Mrs. Gepp was unabie to do
her share of the work.
* The number of species common to both regions is probably far greater, for in the recently published paper by
Br. Schroder, ‘“* Zellpflanzen von Ost-Afrika,” Hedwigia, Bd. lii., four more species are mentioned, also found in the
present collection.
WEBER-VAN BOSSE—MARINE ALG 271
IT. List of Localities and Names of the Algz found at each of them.
Cuacos ArcHIPELAGO: DireGo Garcia, EGMont, SaLomon, PEROS (ILE pu Cory).
1. Gelidium crinale (Turn.), Lamx.; 2. Gelidiopsis rigida (Vahl), Web. v. B.;
3. Hypnea pannosa, J. Ag.; 4. Laurencia obtusa (Huds.), Lamx.; 5. L. spinulifera,
Kiitz.; 6. Z. pygmea, nu. sp.; 7. Chondria pumila, Vick.; 8. Herposiphonia prorepens
(Harv.), Schm.; 9. Dasyopsis Geppii, n. sp.; 10. D. palmatifida, n. sp.; 11. Dictyurus
purpurascens, Bory ; 12. Antithamnion adnatum, J. Ag.; 13. Ceramium cinnabarimum
(Gratel.), Hauck; 14. Reinboldiella Schmitziana (Reinb.), de Toni; 15. Peyssonnelia
calcea, Heydr.; 16. Melobesia callithamnioides, Falk.; 17. Amphiroa fragilissima
(Linn.), Lamx.; 18. Jana tenella, Kiitz.
COETIVY, REEF.
1. Chantransia Liagore,u.sp.; 2. Lnagora orientalis, J. Ag.; 3. L. hawatiana, Butt. ;
4. Gelidium pannosum, Grun.; 5. Champia compressa, Harv.; 6. Laurencia obtusa
(Huds.), Lamx.; 7. LZ. papillosa (Forsk.), Grev., f. australica in herb. Kiitz.; 8. L. cey-
lanica, J. Ag.; 9. Oligocladus Prainiwi, n. sp.; 10. O. pusillus, n. sp.; 11. Dasyopsis
aperta, n. sp.; 12. D. Geppii, n. sp.; 18. D. palmatifida, n. sp.; 14. Heterosiphonia
spec., fragment; 15. Dictyurus purpurascens, Bory; 16. Jania tenella, Kiitz.
SEYCHELLES ARCHIPELAGO (various localities).
1. Sarconema furcellatum, Zan.; 2. Gelidiopsis variabilis (Grev.), Sclim.; 3. Coral-
lopsis Cacalia, J. Ag.; 4. Hypnea Valentie, Turn. ; 5. H. pannosa, J. Ag.; 6. Champia
spec. ; 7. Lawrencia papillosa (Forsk.), Grev., f. australicain herb. Kiitz.; 8. Acanthophora
orientalis, J. Ag.; 9. Roschera (Tolypiocladia) glomerulata (Ag.), n. comb., fragment ;
10. Peyssonnelia coccinea, J. Ag.; 11. P. biradiata, n. sp.; 12. Jania tenella, Kitz.
Mauer, Cap TERNE, REEF.
1. Dermonema dichotomum, Harv.; 2. Galaxaura fastigiata, Decne. ; 3. Gelidiopsis
rigida (Vahl), Web. v. B.; 4. Corallopsis Cacalia, J. Ag.; 5. Hypnea pannosa, J. Ag.;
6. Acanthophora orientalis, J. Ag.; 7. Amphiroa fragilissima (Linn.), Lamx.; 8. Jania
tenella, Kitz.
PRASLIN, REEF.
1. Galavaura Liebmanni (Aresch.), Kjellm.; 2. Gelidiopsis rigida (Vahl), Web. v. B. ;
3. Hypnea pannosa, J. Ag.; 4. Polysiphonia spec., fragment; 5. Amphiroa foliacea,
Lamx.; 6. Jania tenella, Kiitz.
AMIRANTH.
1. Galaxaura veprecula, Kjellm.; 2. Iridea spec. ; 3. Polycelia van Hoevellii, n. sp.,
fragment; 4. Gracilaria compressa (Ag.), Grev.; 5. Calhidlepharis prolifera (Harv.),
J. Ag.; 6. Hypnea spec.; 7. Champia compressa, Harv.; 8. Calarthrum Albertisii
(Pice.), Borg.; 9. Hypoglossum spec. (spathulatwm, Kutz.?); 10. Nitophyllum spec.,
35—2
272 PERCY SLADEN TRUST EXPEDITION
fragment ; 11. Laurencia ceylanica, J. Ag.; 12. Pseudendosiphonia Gardineri,
n. g., n. sp.; 13. Cladhymenia oblongifolia, Hook. et Harv.; 14. Dasyopsis Geppii,
n. sp.; 15. D. palmatifida, n. sp.; 16. Tapeinodasya Ethele, n. sp.; 17. Dictyurus pur-
purascens, Bory; 18. Griffithsia spec., fragment; 19. Cryptonemia seminervis, J. Ag.;
20. Peyssonnelia polymorpha (Zan.), Schm., var. Gardineri, n. var.
AupaBRa (collected by J. C. F. Fryer).
1. Galaxaura hawaiiana, Butt.; 2. Hypnea divaricata (), Grev.; 3. H. pannosa,
J. Ag.; 4. Laurencia nidifica, J. Ag.; 5. L. obtusa (Huds.), Lamx.; 6. L. papiliosa,
(Forsk.), Grev., f. awstralica in herb. Kitz.; 7. Chondria simpliciuscula, nu. sp.;
5. Amphisbetema indica (J. Ag.), n. n.; 9. Spyridia filamentosa (Wulf.), Harv., frag-
ment; 10. Halymenia polyclada, A. & K. 8S. Gepp, var. aldabradensis, n. var.
SAYA DE Manna.
1. Galaxaura fastigiata, Deene., fragment; 2. G. obtusata (Sol.), Lamx., fragment ;
3. Eucheuma Cottonii, n. sp.; 4. Hauchea microspora, Born.; 5. Gloioderma(?) expansa,
n. sp.; 6. Chylocladia perpusilla, n. sp.; 7. Dictyurus purpurascens, Bory; 8. Crypto-
nemia seminervis, J. Ag.; 9. Cryptonemia spec.; 10. Peyssonnelia coccinea, J. Ag.;
11. P. Harveyana(?), Crouan; 12. Cruoriopsis cruciata, Dut.
CARGADOS CARAJOS.
1. Gloiophlea articulata, n. sp.; 2. Iridea spec., fragment; 3. Hucheuma Cottonii,
n. sp.; 4. Dasyopsis Stanleyi, n. sp.; 5. D. Geppw, n. sp.; 6. Heterosiphonia Rendilec,
n. sp.; 7. Dictyurus purpurascens, Bory; 8. Haloplegma Preissia, Sond. ; 9. Ceramium
spec.
III. Systematic Survey of all the Alge collected by Mr. J. Stanley Gardiner
in the Indian Ocean.
REEF OR
FAMILy. LocauLity. ThAGOOR. DEPTH. DISTRIBUTION.
HELMINTHOCLADIACER.
1. Chantransia Liagore, n. sp. ............ Coetivy. Reef.
1. Liagora orientalis, J. Ag. ....| Coetivy. INOS |i Badbng Indian Ocean, Ceylon.
2. Liagora hawatiana, Butt. ............... Coetivy. NRCC rm i, atestets Pacific Ocean, Laie Point, Koolanloa,
Oahu.
1. Dermonema dichotomum, Harv. .........) Mahé, Terné. Recta) Me S| keds Ceylon, Malay Archipelago, Formosa,
CHETANGIACER.
1. Gloiophlea (?) articulata,n. sp. ...... Cargados Carajos. | sw... a 30-45 fms,
1. Galawawra Tiebmanni (Ax.), Kjellm....| Praslin. TOBE = p0000 Gulf of Mexico.
2. Galarawra hawaiiana, Butt. ............ Aldabra. Passe Houa- | ...... Pacific Ocean, French Frigate Shoal,
reau, outside. Bird Island.
3. Galaxaura fastigiata, Decne. ............ Mahé, Terné. Tee, SR eens Pacific Ocean, Philippines ; Malay Archi-
Saya de Malha, | s..eea sae Over 26 fms.| pelago.
4, Galuxaura veprecula, Kjellm. ......... Airis sp 000 20-25 fms. | Indian Ocean, Madagascar.
5. Galaxaura obtusata (Sol.), Lamx. ...... Saya de Malha, | ......... 26 fms. Warm Atlantic, West Indies; Pacific
Ocean, Loo Choo Isl., Japan.
WEBER-VAN BOSSE—MARINE ALG
273
. Chondria simpliciuscula, n. sp. ......... |
REEF OR
FamMiny. LocauLity. TLAGOOR. DEPTH. DISTRIBUTION.
GELIDIACEZ.
1. Gelidiwm crinale (Turn.), Lamx. ...... Diego Garcia. Mmeethayee - li” uals Mediterranean ; Red Sea.
2. Gelidiwm pannosum, Grun. .........:.. Coetivy. INO = aon 000 Ceylon, Friendly Islands, Fiji, Tonga,
and Samoa Islands.
GIGARTINACE.
i Cargados Carajos. 22 fms.
Il, TipHEe@ BIWEGoy D scoscodaccevesose0c000 { eaten 20-25 fms.
1, Polycelia van Hoevellii, n. sp. ......... Amirante. 30-100 fms. | Malay Archipelago.
RHODOPHYLLIDACER.
1. Sarconema furcellatum, Zan. ............ Seychelles, Long Isl.) Reef. 8 | ...... Red Sea; Indian Ocean.
1. Hucheuwma Cottonit, n. sp. ...........005- Cargados Carajos. 26 fms Many species of Huchewma occur on
Saya de Malha. 25 fms the east coast of Africa.
SPH#ROCOCCACER,
. Gelidiopsis variabilis (Grev.),Schm....| Seychelles, Long Isl.) Reef. = | ...... Indian Ocean, east coast of Africa.
Seychelles, Praslin
2. Gelidiopsis rigida (Vahl), Web. v. B.+ | Diego Garcia. Reef i esis Tropical seas.
Mahé, Terné.
1. Gracilaria compressa (Ag.), Grey. ...... Amber, || so e00000 25 fms Atlantic Ocean; Mediterranean; Gulf of
Mexico.
1. Corallopsis Cacalia, J. Ag. ........005- Mahé, Terné. Rieehwa yy iy eee Red Sea, east coast of Africa.
Seychelles, Long Isl.| Reef.
1. Calliblepharis prolifera (Harv.),J. Ag.) Amirante. = | ......... 25 fms New Zealand.
1. Hypnea divaricata (?), Grev. ...........- Aldabra. lla, == Sab c00 Gulf of Mexico; Pacific, Tongatabu.
2. Hypnea Valentiz (Turn.), Mont. ...... Seychelles, Long Isl.) Reef. 8 | ...... Red Sea; Indian Ocean ; Port Denison.
3. Hypnea pannosa, J. Ag. .......eseeeee eee Chagos Archipelago.) Lagoon, = | ...... Coast of Mexico; Mauritius; Ceylon;
Aldabra. Reef. Tonga Archipelago; New Caledonia.
Seychelles. Reef
Praslin. Reef
Mahé, Terné. Reef.
AN Hy pied SPCC. .......-.cneceereenee sce smecee Amirante, | wc... 25 fms.
RHODYMENIACEZ,
1. Fauchea microspora, Born. ............... Saya de Malha, | ......... 53 fms. Mediterranean.
1. Gloioderma (?) expansa,n. sp. ......... Saya de Malha. | ......... 53-55 fms.
1. Champia compressa, Harv. ............ Ane, «|| senc00000 20-25 fms. | Cape of Good Hope; Malay Archipelago ;
Coetivy. Reef Pacific Ocean.
Mm OMONMUPUG SPCCe vatijeacsefecccenseceeseemecee Seychelles, Long Isl.) Reef
1. Chylocladia perpusilla, n. sp. ......... Saya de Malha. | ......... 29 fms.
1. Celarthrum Albertisii (Picc.), Borg. ...) Amirante. =| ss see 20 fms. Canary Islands; Bermudas; Malay
Archipelago.
DELESSERIACER.
1. Hypoglosswm spec. .........06e see seeeeeees Amirante. 25 fms
1. Nitophyllwm spec. Amirante. 25 fms
RHODOMELACER.
1. Laurencia nidifica, J. Ag. ...............| Aldabra. Ree == 000 Sandwich Islands; Malay Archipelago.
Aldabra. : . qRpe aeyo
2, Lawrencia obtusa (Huds.), Lamx. ...4 | Coetivy. WEG || goo000 { ore 5 HIE Hlbelss Aitoliayy
Egmont. eee
3. Laurencia papillosa f. australica ...... Aldabra. REGn = T0000 New Caledonia.
Coetivy. Reef
Seychelles, Long Isl.| Reef. ;
4, Laurencia spinulifera, Kiitz. ............ Chagos Arch.,Diego| Reef. | seven Indian Ocean.
Garcia.
5. Laurencia pygmexa, N. Sp. .........0.+-+- Chagos Arch., Diego | Reef
Garcia. f
6. Lawrencia ceulanica, J. Ag. ......-..0-- Amami ——||_—gawoo0000 20 fms Ceylon; Malay Archipelago.
i Coetivy. Reef 5 : i
1. Acanthophora orientalis, J. Ag.......... Mahé, Terné. echelons Zanzibar ; Malay Archipelago; North
Seychelles, Long Isl.| Reef. Australia ; Tonga Archipelago ;
Marianne Islands.
1. Pseudendosiphonia Gardineri,n.g.n.sp.| Amirante. 3 | ws.ereee 25 fms
1. Cladhymenia oblongifolia,Hook.etHary.. Amirante. = | ........ 25 fms New Zealand.
1. Chondria pumila, Vick. ..........06.00.-+ Chagos Arch.,Diego| Reef. 8 | see Barbados.
Garcia.
2 Aldabra. Reef.
274
PERCY SLADEN TRUST EXPEDITION
= . REEF OR
FamMizy. Locantry, TEAC GOR, DEPTH. DISTRIBUTION.
RHODOMELACE® (con.).
Ue JENS NAO OAG) (NEO) Soocaossesonesaos connor Praslin. Reef [pelago.
1. Roschera glomerulata (Ag.), n.comb....| Seychelles, Long Isl.) Reef. § | .... Zanzibar ; Indian Ocean; Malay Archi-
1. Herposiphonia prorepens (Harv.),Schm.) Chagos Arch, |} eaeeee | wee Algoa Bay; Western Australia.
1. Oligocladus Prainii, n. sp. .............- Coetivy. Reef.
2. Oligocladus pusillus, nD. sp. .........-.- | Coetivy. Reef
1. Dasyopsis Stanleyi, n. sp. ..........- ..|Cargados Carajos. | ........ 30and47 fms.
|
5 Coetivy. Reef.
2 SINaO GHOE
2. Dasyopsis aperta, 0. Sp. ..........06-- { Ghaaniies Canaan, || cescccere 47 fms.
Armingya ===} Gagongoon 20-25, fms.
Ree: : | Cargados Carajos. | .......6 25 fms.
3. Dasyopsis Geppii, 0. Sp. ........0.060- Coston, Reef.
Chagos Arch. :
Ascatenti®, == |) asnaonaeo 20-25 fms. | Malay Archipelago.
4, Dasyopsis palmatifida, n. sp. ......... Coetivy. Reef.
Chagos Arch.
1. Tapeinodasya Ethele, n. sp. Asani, = = = || coesonona 20-25 fms.
1. Heterosiphonia Rendlei, n. sp. .| Cargados Carajos. | ......... 47 fms.
2. Heterosiphonia spec. ............4.- ....| Coetivy. Reef.
1. Dictyuwrus purpurascens, Bory ...... Saya de Malha. 55 fms. Cape Comorin ; Indian Ocean, Mauritius,
Coetivy. Reef. | Ceylon; Malay Archipelago; Diego
Amirante. 30 fms. Garcia.
Cargados Carajos. 30 and 47fms.
| | Chagos Arch., 22 fms.
(| Salomon.
1. Amphisbetema indica (J. Ag.) ......... Aldabra. Reckless Diego Garcia.
CERAMIACER.
il, CrraiiB0a FIDEO> socooceconcoenacsoadansnceu JNO, =H Seam nd | 20-25 fms.
1. Haloplegma Preissvi, Sond. ............ Cargados Carajos. | ......... 20, 30, and} Western Australia, Tasmania; Malay
47 fms. Archipelago.
1. Antithamnion adnatum, J. Ag. ......... Salomon. Riectmie ll) \ actes | New Zealand; Malay Archipelago.
1. Spyridia filamentosa (Wulf.), Harv. ...| Aldabra. Re ¢ fa | eect ee Mediterranean; Red Sea; Dar-es-Salaam;
Malay Archipelago ; West Indies.
1. Ceramium cinnabarinum(Gratel.),Hck.) Salomon, 2] eee | tees | Mediterranean ; Malay Archipelago.
2. Cerqmiwm Spec. \......-steeseen eee ens er sere Cargados Carajos. | ......... 47 fms |
1. Reinboldiella Schmitziana (Rbd.), | Chagos Arch., Eg-| Reef. | ...... Japan; Malay Archipelago.
de Toni.) mont, Salomon.
GRATELOUPIACES.
1. Halymenia polyclada, A. & E.S.Gepp,) Aldabra.
var. aldabradensis, n. v.
4 +3 eho Amirante. 20-25 fms. | Atlantic; Mediterranean; Red Sea.
1. Cryptonemia seminervis, J. Ag. ...... See dla Melina, 55 fms.
2. Cryptonemna Spec. .........-eecee-eeeeeees Saya de Malha. 55 fms.
SQUAMARIACER,
1. Peyssonnelia coccinea, J. Ag. ............| ‘Sey.chell essai uu uieteeeer ce 31 fms West coast of Australia; Malay
Saya de Malha. 55 fms Archipelago.
2. Peyssonnelia Harveyana (?), Crouan ...| Saya de Malha. 55 fms Coast of France near Brest.
3. Peyssonnelia calcea, Heydr. ............ leno, = IRE nc Tami, near German New Guinea; Malay
Peros, Coin. Archipelago.
4. Peyssonnelia polymorphat. Gardinert,| Amirante. 4 424 2 | ceveevene 30, 45-60
n. f. fms.
5. Peyssonnelia biradiata, n. Sp............. Seychelles, | — ...coveoe 31 fms. | Malay Archipelago,
6. Peyssonmelid SPC. ..c.ececscesecssaeeenees Saya de Malha.
1. Cruoriopsis cruciata (?), Duf. ............ Saiyancew Vial] na) aa | ester 29 fms.
a
CoRALLINACER.
. Melobesia callithamnioides, Falk. ......
. Amphiroa fragilissima, Lamx.
. Amphiroa foliacea, Lamx. ..............-
. Jama tenella, Kiitz.
Egmont.
Chagos Arch.
Mahé, Cap Terné.
Praslin.
Coetivy.
Seychelles.
Peros, Coin.
Praslin.
Mahé, Cap Terné.
Adriatic; Malay Archipelago.
Atlantic (?) ; Mediterranean.
West Indies; Pacific; Malay Archi-
pelago; Dar-es-Salaam.
Marianne Islands; Malay Archipelago.
Malay Archipelago ; Gulf of Mexico.
WEBER-VAN BOSSE—MARINE ALG A 275
LV. Systematic Account*.
Fam. HELMINTHOCLADIACEA.
CHANTRANSIA (DC.), Schmitz.
1. CHANTRANSIA LIAGORA, 0. sp.
Thallo endophytico et epiphytico, nano, czespitibus globosis sparsis Liagorze insidente,
constante e filis hospitem intrantibus et filis erectis. Filis erectis parce ramosis,
ramis post monosporangia terminalia delapsa egredientibus; pilis non visis,
verisimiliter raris. Monosporangiis aut terminalibus aut lateralibus, pedicello
ramoso suffultis, ramis denuo monosporangia ferentibus. Filis latis 6-8-12 u,
articulis 16-20 » longis. Monosporangiis non maturis; antheridiis et carpogoniis
non Visis.
Coetivy, reef, on Liagora hawaiiana; in alcohol.
This little Chantransia forms a small tuft, which penetrates between the horizontal
filaments of the Ziagora and attaches itself at their base. I could detect no creeping
filaments in the host plant giving rise to new tufts. It has a height of 1 mm., the axes
are sparingly and irregularly branched and bear monospores both at the apex and
laterally. After the monospore has fallen off, or perhaps while still i situ, the filament
or branch that carries it grows out sideways. I also noted clusters of carpospores ;
these are not arranged in series as in C. efflorescens. The filaments have a breadth of
6-8, rarely of 12 », and their cells a length of up to 20n. On account of the alge
havin® been first preserved in formalin and afterwards in alcohol, it is difficult to judge
the shape of the chromatophore.
I believe that this alga is a new species; there are, so far as I know, only two other
species of Chantransia on Liagora, namely, Ch. Barbadensis and Ch. Nemalionis, which
were found on Liagora elongata by Miss Vickers. Our alga differs from both species
in the smaller size of its cells and filaments.
Liacora, Lamouroux.
1. Liacora ortenTaAuis, J. Ag.
J. Agardh, Analecta algolcgica, 1896, p. 99.
Coetivy, reef; in alcohol.
Distribution. Indian Ocean, Ceylon.
2. LIAGORA HAWATIANA, Butt.
Butters, “ Liagora and Galaxaura,” Minnesota Bot. Studies, vol. iv. part 2, 1911, p. 169.
Coetivy, reef; in alcohol.
Distribution. Laie Point, Koolauloa, Oahu, Pacific Ocean.
Liagora hawaiiana belongs to the group of Liagore with dichotomously branched
fronds. It differs from all known species of this group in having an articulate frond ;
at the top of the thallus the articulations are inconspicuous, but lower down they are
* [Unless otherwise stated the specimens are from surface reefs. J. S. G.]
276 PERCY SLADEN TRUST EXPEDITION
very distinct. The articulations are at the base of each dichotomy and the internodes
are as a rule long.
DERMONEMA, Greville.
1. DERMONEMA DICHOTOMUM, Harv.
Harvey, Ceylon Algee, n. 98.
Schmitz u. Hauptfleisch in Engler u. Prantl, Natiirl. Pflanzenfam., Teil 1, Abt. 2, 1896, p. 335.
De Toni, Syll. Alg. vol. iv. sect. 1. 1897, p. 102.
Mahé, Cap Terné; dry specimens.
Distribution. Ceylon; Formosa; Malay Archipelago.
Fam. CHATANGIACEH A.
GuoropHia@a, J. Agardh.
1. GLOIOPHL@A (?) ARTICULATA, n. sp. (Plate 16. fig. 1; Plate 18. figs. 26, 27.)
Frondibus ‘dichctomis, articulatis, usque ad 18 cm. altis, ramulos adventitios ex nodis
orientes gerentibus; internodiis cylindraceis, subecompressis, ad basin contractis;
internodio terminali apice rotundato. Jrondibus constantibus e cylindro centrali
filis hyalinis, tenuibus, peripheriam versus excurrentibus contexto. Filis per
dichotomiam divisis, stratum periphericum cellularum minimarum moniliformium
arete conjunctarum et cuticula crassa coopertarum formantibus. Organis
generationis ignotis.
Cargados Carajos, dry specimen, 30 fms.; in alcohol, 45 fms.
The alge from Cargados Carajos are barren, the determination is therefore not beyond
doubt, but the anatomical structure of these plants is so typical and so like that of the
family Cheetangiace (Pl. 18. figs. 26, 27) that I should be very much surprised if the
organs of fructification should prove these algze to belong to another family. Amongst
the Cheetangiaceze they stand nearest to Gloiophlea. ‘They are distinguished, however,
from Glotiophlea scinaioides—the only known species of the genus—by their articulate
frond; the internodes are usually—though not always—contracted into a short pedicel-
like node, and from this node adventitious branches may spring, but as a rule the
branching is dichotomous. The peripheral layer consists of small cortical cells, covered
by a rather thick cuticle; these small tightly adhering cortical cells distinguish the
genus Gloiophlea from the genus Scinaia.
GALAXAURA, Lamouroux.
1. GALAXAURA LIEBMANNI (Aresch.), Kjellm.
Areschoug, “ Phycez nove,” in Act. Reg. Soc. Scient. Upsal. ser. 3, vol. i. 1854, p. 356.
Kjellman, “ Floridé Sligtet Galavaura,” in Kongl. Sv. Vet.-Akad. Handl. Bd. xxxiii. 1900, p. 46.
Praslin, reef; in alcohol.
Distribution. Mexico; Vera Cruz.
Though Galaxaura LIiebmanni is only known from the West Indian tropical seas, the
peculiar structure of the short cortical filaments and the compact outward appearance
WEBER-VAN BOSSE—MARINE ALG 277
of the algze from Praslin seem to indicate that they belong to G. Liebmanni rather
than to G. collabens from Australia, described by Kjellman as ‘“‘solitaria, erecta et
robusta.” The plants from Praslin are short, intricate, and grow close together; the
short cortical filaments consist, as a rule, of four cells only, and of these the lower
one is by far the largest.
2. GALAXAURA HAWATIANA, Butt. (Plate 17. fig. 25.)
Butters, “ Liagora and Galaxaura,” Minnesota Bot. Studies, vol. iv. part 2, 1911, p. 176.
Aldabra, reef, outside Passe Houareau.
Distribution. Pacific Ocean, French Frigate Shoal, Bird Island.
The alge from Aldabra are very much like Galaxaura hawaiiana, Butt., and the
differences between the description of G. hawatiana (I have seen no type specimen) and
the alge from Aldabra are not, I think, sufficiently great to warrant the description of a
new species. The algze from Aldabra are a little longer (they have a length of 7 cm.,
a breadth of 2 mm.), and the long internodes are not tapering, but rounded or
truncate. The frond is slightly but distinetly annulate, and in the dried specimens of a
remarkable dark brown-red colour; the anatomical structure is like that of G. hawaiiana,
Butt. Butters records G. hawaiiana from depths of 14-16 and 26 fathoms; the
specimens from Aldabra were found at the outside of Passe Houareau. This difference
of locality may perhaps account for the small differences observable in the two plants.
I have given a figure of the Aldabra plant, which shows very well the long terminal
internodes.
3. GALAXAURA FASTIGIATA, Decne.
Decaisne, Sur les Corallines, 1842, p. 16.
Kjellman, “ Floridé Sligtet Galavaura,” in Kongl. Sy. Vet.-Akad. Hand]. Bd. xxxiii. 1900, p- 64.
Mahé, Cap Terné ; dry specimen.
Saya de Malha, 26 fms. or over; fragment only.
Distribution. Pacific Ocean; Philippines ; Malay Archipelago.
4. GALAXAURA VEPRECULA, Kjellm.
Kjellman, “ Floridé Sligtet Galaxaura,” in Kong]. Sv. Vet.-Akad. Handl. Bd. xxxiii, 1900, ». 80.
Amirante, 20-25 fms.; in alcohol.
Distribution. Indian Ocean, Madagascar.
5. GALAXAURA OBTUSATA (Sol.), Lamx.
Solander in Ellis, Zoophytes, 1786, p. 113.
Lamouroux, Polypiers coralligénes, 1816, p. 262.
Agardh, Epicrisis systematis Floridearum, 1876, p. 525.
Saya de Malha, 26 fms.; only a small specimen.
Distribution. Warm Atlantic, West Indies; Pacific Ocean, Loo Choo Islands,
Japan.
SECOND SERIES—ZOOLOGY, VOL. XVI. 36
278 ! PERCY SLADEN TRUST EXPEDITION
Fam. GELIDIACE.
GELIDIUM, Lamouroux.
1. GELIDIUM CRINALE (Turn.), Lamx.
De Toni, Syl. Alg. vol. iv. sect. 1. 1897, p. 146.
Fucus crinalis, Turner, Hist. Fuc. tab. 198.
Chagos Archipelago, Diego Garcia, in alcohol ; reef.
Distribution. Mediterranean; Red Sea.
2. GELIDIUM PANNOSUM, Grun.
Grunow, Algen der Fiji, Tonga u. Samoa Inseln, 1874, p. 40.
Coetivy, reef, on a piece of coral; in alcohol.
Distribution. Upolu, Samoa Archipelago; Ceylon; Friendly Islands.
Fam. GIGARTINACEA.
Inipma, Bory.
1. IRIDmMA spec.
Cargados Carajos, fragment with cystocarps, in alcohol, 22 fms.
Amirante, fragment with tetraspores, in alcohol, 20-25 fms.
The specimens from both localities are too fragmentary to allow of a specific
determination, though the tetraspores being scattered among the peripheral cells
indicate that the plant belongs to the subgenus Rhodoglossum, J. Ag. Both specimens
are so alike in anatomical structure and outward appearance that, as far as can be
judged from the small fragments, I feel almost certain that they belong to the same
species.
Ponyceenia, J. Agardh.
1. Potyoe@1Lia vAN HOEVELLI, n. sp.
Amirante, 30-100 fms. ; in alcohol.
Distribution. Malay Archipelago. .
The species is represented in the collection by such a poor and barren fragment that
I should have been unable to name it, had it not been for my studies on the rich
material collected during the ‘“‘ Siboga” Expedition in the Malay Archipelago. This alga
will therefore be described and figured in the forthcoming paper on the ‘‘Siboga”’ algee.
Fam. RHODOPHYLLIDACEA.
SarconEMA, Zanardini.
1. SARCONEMA FURCELLATUM, Zan.
Zanardini, Plante Mar. Rubro Enum. p. 264.
De Toni, Syll. Alg. vol. iv. sect. 1. 1897, p. 367.
WEBER-VAN BOSSE—MARINE ALGAE 279
Seychelles, Long Island.
Distribution. Red Sea; Indian Ocean.
Evucueuma, J. Agardh.
1. EvcHrumaA Corrontt, n. sp. (Plate 16. fig. 2.)
Thallo crasso, carnoso, depresso, decumbente, lateraliter et subdichotomice ramoso,
latere superiore verrucis et tuberculis aggregatis, latere inferiore tuberculis remoti-
oribus obsito. Verrucis nonnunquam forma irregulari; tuberculis vulgo parvis et
rotundis, nonnunquam apiculatis. Regione mediana thalli ad apices e cellulis satis
magnis hyphis circumdatis constante, hyphis infra item auctis et cellulas satis
magnas formantibus; peripheriam versus cellulis diminutis et ibidem series breves
formantibus. Organis fructificationis ignotis.
Saya de Malha, 25 fms. ; in alcohol.
Cargados Carajos, 26 fms.; dry specimen.
#. Oottonii is unfortunately a barren plant, and shows, in a dry state, a great likeness
to species of Gracilaria. I moistened it, to restore as much as possible its natural
appearance, and was then struck by its great resemblance to Hucheuma. 'The figures
are made from these moistened specimens, and show well the horizontally spread,
laterally branched thallus, covered with warts and more or less pointed tubercles.
The anatomical structure at the apex is also entirely different from that of Gracilaria,
though, in a cross-section at some distance from the apex, only round, rather large, and
somewhat thick-walled cells are to be seen. But these cells are of different origin from
the primary large cells, for at the apex the latter are seen to be surrounded by narrow
hyphee, which grow out lower down into large cells and are then no longer discernible
as hyphe.
The genus Hucheuma is described by Schmitz in Engler u. Prantl, ‘‘ Pllanzen-Familien,”
as having a central axis of slender elongated hyphe. ‘This axis is found, for instance,
in EF. spinosum. Agardh has, however, called attention to the fact that LH. Schrammi
has a structure in which “fila strati interioris sparsim inter cellulas strati intermedii
quasi introducta obveniunt.” Schmitz, again, supposes that EL. Schrammi, Crouan, is
not an Hucheuma but an Huryomma.
My studies on the Hucheumata of the Malay Archipelago seem to indicate that there
are species of Hucheuma without a central axis of longitudinal hyphe. The investi-
gations are not complete, but it appears to me not improbable that the cylindrical erect
species have an axis of narrow cells, and that in the flat prostrate species this is absent.
I did not find an axis in any prostrate specimens of Huchewma of which I was able to
study the cystocarp. The whole question needs to be further investigated, but for
the present it seems highly probable that the algze from Saya de Malha and Cargados
Carajos belong to the genus Hucheuma, and the name I propose for it is in kind
remembrance of Mr. Cotton’s friendly help.
36—2
280 PERCY SLADEN TRUST EXPEDITION
Fam. SPH HROCOCCACE.
GELIDIoPsSIS, Schmitz.
1. GELIDIOPSIS VARIABILIS (Grev.), Schmitz.
Schmitz, ‘Marine Florideen von Deutsch Ost-Afrika,” in Engler’s Bot. Jahrb. Bd. xxi. H. 1, 2,
1895, p. 148.
Seychelles, Long Island, reef.
Distribution. Indian Ocean; East coast of Africa.
2. GELIDIOPSIS RIGIDA (Vahl), Web. v. B.
Recueil des Travaux botaniques Néerlandais, no. 1, 1905, p. 9.
Fucus rigidus, Vahl, in Naturh. Sellskabets Skrifter, v. p. 46.
Seychelles, reef; dry specimen. '
Praslin, reef; dry specimen and in alcohol.
Diego Garcia, reef; dry specimen.
Mahé, Cap Terné, reef ; dry specimen.
Distribution. Common in tropical seas.
In 1904 I wrote in the above-mentioned paper that it appeared to me that the well-
known Gelidiwn rigidum of the tropical seas was not a Gelidiuwm, but belonged to the
genus G'elidiopsis. In 1909 Prof. Okamura gave, in his ‘“ Icones of Japanese Alge,”
pl. 59, a figure of this alga and also of its large apical cell, and named it Gelidiwm
rigidum. JI had failed to detect this large apical cell, and its absence was one of
the reasons why I thought that the alga could not be a Gelidiwm; I had seen no
cystocarps. Quite recently, in “cones of Japanese Algew,” vol. ii. No. 10, p. 188,
Prof. Okamura has recalled his statement, made before he had seen my paper, and
agrees with me in naming Gelidium rigidum, Gelidiopsis rigida. This alga varies in
its ramification, the lateral branches being short or long, or sometimes unilaterally
inserted. I believe, however, that these variations are only a form of growth.
Gelidiopsis being a feminine substantive in the Greek language, the adjective should
have the feminine ending, and I made an error when I called the alga Geliddopsis
rigidum in 1904.
GRACILARIA, Greville.
1, GRACILARIA COMPRESSA (Ag.), Grev.
Agardh, Species Algarum, 1823, p. 308.
Greville, Algze Britann. 1830, p. 125.
De Toni, Syll. Alg. vol. iv. sect. 11. 1900, p. 438.
Aymirante, 25 fms.
Distribution. Atlantic Ocean ; Mediterranean ; Gulf of Mexico.
The specimen is rather slender for G. compressa and scantily branched, but I assume
WEBER-VAN BOSSE—MARINE ALG 281
that the great depth at which it was gathered may in some measure account for
this. It bears cystocarps, and these have a little funnel at the top, exactly as in
G. compress«.
CoraLuopsis, Greville.
1. CoRALLOPsIs CAcALtA, J. Ag.
J. Agardh, Epicrisis systematis Floridearum, 1876, p. 4.09.
De Toni, Syll. Alg. vol. iv. sect. 11. 1900, p. 459.
Mahé, Cap Terné; dry specimens.
Seychelles, Long Island; dry specimens.
Distribution. Red Sea. -
The specimens of the Seychelles are more branched than those of Mahé, Terné,
still I believe that they belong to the same species; some branches recall those of
C. minor.
CALLIBLEPHARIS, Kiitzing.
1. CALLIBLEPHARIS PROLIFERA (Harv.), J. Ag. (Plate 16. fig. 3.)
Epicrisis systematis Floridearum, 1876, p. 432.
Rhodymenia prolifera, Harvey, in Flora Nova-Zeelandia, p. 249.
Amirante, 25 fms.; dry specimens.
Distribution. New Zealand. .
Harvey described this alga under the name of Rhodymenia prolifera, but J. Agardh
pointed out that the structure of the cystocarp was unlike that of Rhodymenia and
similar to that of Calliblepharis. The alga differs, however, from the other species of
this genus in bearing the cystocarps on the broad part of the thallus and not on special
leaflets ; Agardh called it therefore Calliblepharis ? prolifera. As the specimens from
Amirante were fertile, I studied the ripe cystocarps; they have the same structure as
those of Calliblepharis, and I do not think that the absence of special leaflets should be
a reason to maintain J. Agardh’s note of interrogation.
It is remarkable to find Calliblepharis prolifera in Mr. Stanley Gardiner’s collection,
since it had previously only been found in New Zealand.
Hypnea, Lamouroux.
1. Hypnra (DIVARICATA P), Grev.
Greville, Alg. Brit. p. lix.
De Toni, Syll. Alg. vol. iv. sect. 11. 1900, p. 478.
Aldabra, fragment only, lagoon; dry specimen.
Distribution. Gulf of Mexico; Mascarenes (?); Tongatabu.
As far as I could judge by the small fragment, the specimen belonged to H. divaricata,
Grey. It differed from H. Valentig in the absence of small stellate spines.
282 PERCY SLADEN TRUST EXPEDITION
2. Hypnea VALENTI® (Turn.), Mont.
Cryptog. Canar. p. 161.
De Toni, Syll. Algarum, vol. iv. sect. 11. 1900, p. 479.
Fucus Valentie, Turner, Historia Fucorum, 1809, tab. 78.
Seychelles, Long Island; dry specimen.
Distribution. Red Sea, Indian Ocean; Port Denison.
3. HypnrA PANNOSA, J. Ag.
J. Agardh, Alg. Liebman. p. 14.
Grunow, Algen der Fiji, Tonga u. Samoa Inseln, 1874, p. 39.
Aldabra; dry specimen.
Seychelles, reef; dry specimen,
Praslin; in alcohol.
Chagos Archipelago, Peros, Tle du Coin, lagoon ; in alcohol.
Mahé, Cap Terné, reef; dry specimen.
Distribution. Coast of Mexico; Mauritius; Ceylon; Tonga Archipelago; New
Caledonia.
Among the specimens from Praslin were some overgrown by a sponge; the branches
thus hemmed in were obviously less branched and more cylindrical than the free-growing
ones. The specimens from Mahé, Terné, are exactly like those from Mexico in Kitzing’s
herbarium and figured in Tab. Phye., Band xvili. tab. 27. The specimens from the
other localities are like those gathered by Harvey at the Friendly Islands, No. 44, and
by Schmarda on the west coast of Ceylon. There is a difference between Harvey’s and
Schmarda’s specimens, but both being barren I abstained from further examination.
4. HYPNEA spec.
Amirante, 25 firs.
An indeterminable specimen, altered perhaps a little in consequence of the great
depth at which it grew.
Fam. RHODYMENIACE.
Faucuea, Montagne et Bory.
1, FAUCHEA MICROSPORA, Born.
Bornet in Rodriguez, Algas de las Baleares, 1889, p, 253, et in Bull. Soc. Bot. t. xxxvii. 1890,
p. 142.
Saya de Malha, 53 fms.; dry specimens.
Distribution. Mediterranean.
The plants are all barren, but the whole outward appearance of these algze corresponds
so well with the description given by Bornet that I feel no doubt that they belong to
this species.
WEBER-VAN BOSSE—MARINE ALG 283
GLoropERMA, J. Agardh.
1. GLOIODERMA (?) EXPANSA, n. sp. (Plate 18. figs. 28, 29.)
Thallo membranaceo, plano, verisimiliter horizontaliter expanso, margine prolifera-
tionibus in thallos juveniles excrescentibus. Thallo seepe anastomosante, constante
e strato unico cellularum maximarum centralium, utroque latere duobus stratis
cellularum parvarum et uno strata cellularum minimarum periphericarum cincto.
Cystocarpiis et antheridiis ignotis ; tetrasporangiis in strato corticali parum mutato
dispersis, cruciatis divisis.
Saya de Malha, 53-55 fms. ; dry specimens.
This alea seems to be new to science, and I have referred it to the family of the
Rhodymeniacez on account of its large central cells placed in one series only. Its
outward appearance recalls Rhodophyllis peltata, Grun., of the Tonga Archipelago, but
it differs from that alga in having only one series of large central cells, whilst all species
of Rhodophyllis have two series. Amongst the Rhodymeniacee it belongs, I think, to
the genus Gloioderma, because it has very small peripheral cells and cruciate
tetrasporangia scattered in the cortical layer, which is, nevertheless, very little
changed.
Unfortunately I have seen neither procarps, cystocarps, nor antheridia, and therefore
the generic position of the alga remains uncertain. It needs to be further investigated,
for it was gathered at a great depth (53-55 fms.), from which few Rhodophycee are
known.
By its broad, membranous, unbranched frond, which bears excrescences along its
margin, and sometimes, though rarely, on the frond itself, this alga is easily distinguished
from all other species of Gloioderma, as these have mostly a dichotomous or pinnate
ramification. The entirely different mode of growth of the alga from Saya de Malha,
and the great depth at which it was gathered, almost suggest that it may be the type of
anew genus.
CuHamptia, Desvaux.
1. CHAMPIA COMPRESSA, Harv.
Harvey, Genera South African Plants, ed. 1, p. 402,
De Toni, Syll. Aig. vol. iv. sect. 11. p. 561.
Amirante, 20-25 fms. ; small fragment, in alcohol.
Coetivy, reef; dry specimen.
Distribution. Cape of Good Hope; Ceylon; Malay Archipelago; Friendly Islands; |
Australia (?) ; New Caledonia.
2. CHAMPIA spec.
Seychelles, Long Island, reef; dry specimen.
A small specimen intermingled with Gelidiopsis variabilis. The simple main axis
bears a few short branches, irregularly placed. Several main branches have grown
284 PERCY SLADEN TRUST EXPEDITION
together, and cannot be separated without damage. The plant has distinct inflated
articulations blunt at both sides, and is certainly different from Champia compressa. It
strongly recalls Ch. parvula, especially the figure given by Kiitzing in his. Tab. Phyc.,
Band xv. tab. 89, where Ch. parvula is figured under the name of Lomentaria intertexta.
The plant from the Seychelles is, hewever, very small, having a height of only 1 cm., and
it is barren.
CHYLOCLADIA, Greville.
1. CHYLOCLADIA PERPUSILLA, n. sp. (Plate 16. figs. 4, 5.)
Thallo minusculo, decumbente, ramuloso, cavo, sine diaphragmate; ramo_pri-
mario interne multis cellulis magnis ; ramulis constantibus e strato unico peripherico
cellularum magnarum et parvarum secundum quarum latus interne fila ramosa
decurrunt, nonnunquam glandulas ferentia. Sporangiis in ramulorum dilatatorum,
cavitatem prominentibus, nec in cavernis propriis sitis. Cystocarpiis et antheridiis
ignotis. hallo lato 0°5-0°6 m.
Saya de Malha, creeping on Udotea, 29 fms.
This tiny plant was found creeping on Udotea, though it did not appear to possess
rhizoids. It has a main axis, from which spring slender ramuli, sometimes opposite, but
mostly irregular; the specimen is, however, so small that it is impossible to say much
as to its ramification. The main axis has a length of 1 em. and a breadth of 600 n, the
branches have a length of 2-5 mm. The branches are entirely hollow, but the down-
growing filaments that are proper to the genus bear glands (?); the main axis is filled
with large cells, loosely attached to one another. I could see no diaphragms, neither in
the branches nor at the base of the branches; the tetraspores are scattered over the
inflated top of some branches ; they protrude into the cavity of the branch, but they are
not situated in a special hollow, as they are in Lomentaria.
On account of the position of the tetraspores and the absence of diaphragms, I believe
that this little plant belongs to the genus Chylocladia. Its small size characterizes it as
a new species.
C@LARTHRUM, Borgesen.
1. C@LARTHRUM (ALBERTISII ?) (Picc.), Borg.
Lomentaria Albertisii, Piccone, Alghe in ‘“ Crociera del Corsaro,” 1884, p. 37.
Amirante, 20 fms.; dry specimen.
Distribution. Canary Islands; Guadeloupe and Bermudas; West Indies; Malay
Archipelago.
I think it very probable that the alga from Amirante is Celarthrum Albertisii, because
I found that species amongst the “Siboga” material. I cannot deny, however, that
the specimen has smaller articulations than the type and that I did not see in my
WEBER-VAN BOSSE—MARINE ALG 285
preparations the peculiar glands described by Mr. Borgesen. To what extent, however,
these differences depend upon the depth at which the plant was gathered, I do not
venture to say. The specimen—and it is only a fragment—gives the impression of being
still very young and therefore I wish to be doubly cautious in my opinion as to it.
Fam. DELESSERIACE A.
Hypociossum, Kitzing.
1. HYPoGLossuM spec.
Amirante, 25 fms.; dry specimen.
A fragment; probably Hypoglossum spathulatum (Wiitz.), J. Ag. (Delesseria hypo-
glossoides, Harvey, Austral. Alg.), but the specimen is too badly preserved to be
identified with certainty.
NitopHyLium, Greville.
1. NrvopHYLLUM spec.
Amirante, 25 fms.
Only a poor fragment, attached to other alge and indeterminable, but with a.
_tetraspore-bearing sorus typical of this genus.
Fam. RHODOMELACEA,
Subfam. LAURENCIE&.
LAURENCIA, Lamouroux.
1. Lavrencia nipiFica, J. Ag.
J. Agardh, Spec. Alg. ii. 1863, p. 749.
De Toni, Syll. Alg. vol. iv. sect. 11. 1903, p. 785.
Aldabra, reef.
Distribution. Sandwich Islands; Malay Archipelago.
9. LAURENCIA OBTUSA (Huds.), Lamx.
Hssai de classification, p. 42.
De Toni, Syll. Alg. vol. iv. sect. 111. 1903, p. 791.
Fucus obtusus, Hudson, Flora Anglica, ed. 3, p. 586.
Aldabra, reef; dry specimens.
Coetivy, reef; dry specimen.
Egmont, reef and shoal; in alcohol.
Distribution. Mediterranean ; Fiji Islands; Malay Archipelago.
Laurencia obtusa is represented by two forms in the Stanley Gardiner collection ; from
Aldabra there are three packets (nos. 4, 5, 8). No. 4 is the stoutest plant, no. 5 and
SECOND SERIES—ZOOLOGY, VOL. XVI. 37
286 PERCY SLADEN TRUST EXPEDITION
no. 8 are more slender, while the alga from Coetivy Reef is very much like specimens in
Kiitzing’s herbarium and named L. obtusa var. racemosa. The specimens from Egmont
Reef are rather short, but they probably represent young plants. They show the charac-
teristic branching of the species, and all the branches have blunt truncated apices.
3. LAURENCIA PAPILLOSA (Forsk.), Grev., f. AUSTRALICA, in herb. Kiitzing.
Aldabra, reef; dry specimen.
Seychelles, Long Island ; dry specimen.
Coetivy, reef.
Distribution. New Caledonia.
I have been in doubt about these algze for they differ from typical specimens of
LL. papillosa. They come near to two barren specimens from New Caledonia in
Kiitzing’s herbarium. Mr. Stanley Gardiner’s specimens being also barren, I abstained
from giving a diagnosis.
4, LAURENCIA SPINULIFERA, Kitz.
Kiitzing, Tab. Phye. Bd. xv. 1865, tab. 61, p. 22.
Chagos Archipelago, Diego Garcia, reef; in alcohol.
Distribution. Indian Ocean.
The specimens come near to some forms of L. obtusa, but they are distinguished by
their small size. I think that until all the forms of ZL. obtusa from the Hast Indies are
better known, we may retain Kiitzing’s species.
5. LAURENCIA PYGMHA, n. sp. (Plate 16. fig. 6.)
Fronde tereti, filiformi, tenui, circa 1 cm. alta, pulvinatim ceespitosa, intricata ; ramis
primariis decumbentibus, secundariis erectiusculis; ramulis clavatis, truncatis.
Ramis primariis usque ad 250 n» erassis, ramis secundariis et ramulis tetrasporiferis
100-150 u crassis. Cystocarpiis et antheridiis non visis.
Chagos Archipelago, Diego Garcia, reef; in alcohol.
Laurencia pygmea is distinguished by its size, which is indeed dwarf for a Lawrencia.
It grows in tufts, and the alcohol material looks as if it had grown on stones or shells
and had been cut off with a knife. The thallus consists of decumbent branches—I have
seen no rhizoids—which give off suberect ascending ones; these are branched irregularly.
Sometimes the ramuli are subopposite, sometimes subverticillate or even single. They
are always slender, and not particularly short as are the ramuli of LZ. pannosa, Zan.
From this plant, Z. pygm@a may also be known by its thalli, which, though intricate,
are easily teased out under a pocket-lens, whereas in the case of L. pannosa this cannot
be done, according to Zanardini, without damaging the frond. The mode of branching
reminds me of L. obtusa, but the thallus is very slender, the main branches having a
diameter of 200-250 » and the side-branches and tetrasporiferous ramuli from 100-150 p.
Characteristic of this small alga are the swollen light-reflecting lateral membranes of
the central cells. From Z. indica, Hauck, it is easily known by its small size.
WEBER-VAN BOSSE—MARINE ALGAE 287
6. LAURENCIA CEYLANICA, J. Ag.
J. Agardh, Epicrisis Syst. 1876, p. 660.
De Toni, Syll. Alg. vol. iv. sect. 111. 1903, p. 805.
Amirante, at 20 fms.; dried specimen and in alcohol.
Coetivy, reef; in alcohol.
Distribution. Ceylon.
Subfam. CHONDRIES.
AcANTHOPHORA, Lamouroux.
1. ACANTHOPHORA ORIENTALIS, J. Ag.
J. Agardh, Species Algarum, ii. 1863, p. 820.
De Toni, Syll. Alg. vol. iv. sect. m1. 1903, p. 822.
Borgesen, “Some new or little-known West Indian Floridee, 11.,” in Bot. Tidsskr. Bd. xxx.
1910, p. 201.
Mahé, Cap Terné; dry specimen.
Seychelles, Long Island; dry specimen.
Distribution. Zanzibar; Malay Archipelago; North Australia; Tonga Archipelago ;
Marianne Islands.
Both specimens are full grown and large, but both are barren, and thus a study of
the stichidia is out of the question. Ina recent paper Mr. Borgesen has called atten-
tion to the fact that the name of A. Thierii ought to be changed to A. spicifera and that
the antheridia of A. orientalis are very much like those of A. spicifera.
PSEUDENDOSIPHONIA, 0. g.
Thallus radicibus parvis substrato affixus et verisimiliter horizontaliter expansus,
constat ex axi principali, ramis lateralibus incremento infinito et ramis dorsalibus
ventralibusque incremento finito. Ad apicem vegetationis axis principalis radiatim
se findit ; axis principalis postea etiam inferius bilateralis. Axes laterales eodem
modo quo axis principalis nascuntur. Thallus formatus e cellula centrali et quatuor
cellulis pericentralibus pluribus stratis cellularum corticalium peripheriam versus
diminutarum cinctis. Cystocarpia et antheridia ignota. Stichidia in apice
pedicelli monosiphonii, sporam unam in singulis segmentis continentia. Pedicellus
monosiphonius ramificatus; stichidia nuda.
1. PSEUDENDOSIPHONIA GARDINERT, n. sp. (Plate 17. figs. 13-15; Plate 18. fig. 30.)
Stichidiis sparsis extra axillaribus; ceteroque confer diagnosin generis.
Amirante, 20-25 fms.; in alcohol.
Pseudendosiphonia Gardineri recalls, by its short thorny branches and its stichidia
borne on a monosiphonous stalk, the genus Hndosiphonia. It has, however, a different
37—2
288 PERCY SLADEN TRUST EXPEDITION
habit, having a well defined stem and regularly placed lateral branches, whereas Hndo-
siphonia has no definite stem nor regular ramification. Anatomically the genus is
characterized by its four big pericentral cells (fig. 30, p.c.), surrounded by several layers
of cortical cells which are always much smaller. In Endosiphonia the four pericentral
cells are surrounded by cells of the same size.
The stem of Pseudendosiphonia is attached to small flat stones or particles of rock
by means of small rhizoids and around it stand four branches with a divergence of 7;
they take their origin from the central cell and are therefore endogenous; the lateral
ones are of unlimited growth (Langtriebe), the dorsal and ventral ones remain short
(Kurztriebe). The lateral branches have the same development as the principal stem,
but their side-branches remain at first short and give the plant a thorny appearance
that strongly reminds one of Endosiphonia. At the top of some branches there are
luxuriously developed trichoblasts (“ leaves,” Falkenberg), but on most they had fallen
off, probably because they have so many side-branches and such large cells.
The stichidia bear a great resemblance to those of Hndosiphonia, particularly to
E. clavigera, for they are placed irregularly on the stem and branches (Pl. 1%. fig. 14)
and not in the axils of the latter. They spring from peripheral cells and have a mono-
siphonous pedicel, but this bears branches, which is not the case with #. clavigera. The
stichidium itself was bare in all the specimens examined, but I think it likely that the
branches of the pedicel may develop further stichidia. I noted at the top of these
branches some divided cells that I thought would develop into stichidia, but no later
stages were observed. The cells covering the tetraspores in the stichidium are not
horizontally disposed as in Hndosiphonia but are arranged more or less obliquely. It is
not certain, however, how far this may change as the sporangium attains maturity.
This alga differs from the genus Hndosiphonia by its habit and anatomical structure,
and though nearly related to it, I believe I am justified, on account of these differences,
in describing it as a new genus.
I have great pleasure in dedicating this interesting species to Mr. J. Stanley
‘Gardiner.
CLADHYMENIA, Hooker et Harvey.
1. CLADHYMENIA OBLONGIFOLIA, Hook. et Harv. (Plate 16. fig. 7.)
Hooker et Harvey, Alg. Nov. Zel. no. 70.
De Toni, Syll. Alg. vol. iv. sect. m1. 1903, p. 850.
Amirante, 25 fms.; dry specimens and in alcohol.
Distribution. New Zealand.
The specimens from Amirante are small (Pl. 16. fig. 7) but identical with some that I
received from Prof. J. Agardh from New Zealand under the name of Cladhymenia
oblongifolia, var. I think that the small size of the Amirante specimens may be
accounted for by the great depth from which they come.
WEBER-VAN BOSSE—MARINE ALG 289
Cuonpria, C. Agardh.
1. CHonpRIA PUMILA, Vick. (Plate 16. fig. 8.)
Vickers, “Liste des algues marines de la Barbade,’ in Ann. d. Sc. Nat. 9™¢ série, Bot. vol. i.
p. 45.
Chagos Archipelago, Diego Garcia, reef; in alcohol.
Distribution. Barbados.
On comparing the Chondria from the Chagos Archipelago with Ch. pumila, Vick.
from Barbados, I could detect no character whereby to distinguish it. I conclude,
therefore, that they belong to one species even though the distance between the two
localities is very great; but perhaps this minute plant will be detected later in inter-
vening localities. Miss Vickers intended to give a figure, but her untimely death
prevented her from carrying out this intention. I have therefore given an illustration
of the Chagos plant. .
2. CHONDRIA SIMPLICIUSCULA, n. sp. (Plate 16. figs. 9, 10.)
Thallo filiformi, repente, intricato, pauci ramoso ; ramis interdum oppositis, plerumque
singulis; apice filorum truncato, cellulis apicalibus in fovea crateriformi
immersis. Cellula centrali et quinque cellulis pericentralibus uno aut duobus
stratis cellularum corticalium circumdatis. Cellulis periphericis desuper visis ad
apicem hexagonalibus, in parte inferiore cum pariete undulata et ea re moniliformi.
Organis fractificationis non visis.
Aldabra, reef, on Laurencia papillosa.
This small species is characterized by its filaments being mostly horizontally disposed,
sparingly branched, and bearing the tetrasporangia not in special stichidia but at the
apex of ordinary branches, which ultimately may grow out into long vegetative shoots
when the formation of tetraspores has ceased.
After having been dried Ch. simpliciuscula adhered firmly to the alga on which it had
spread itself, but I detected no union between the two plants. It belongs to the section
Celochondria, Falk., as is clearly shown by fig. 10 on PI. 16.
PoLyYsIPHONTA, Greville.
1. PoLYSIPHONIA spec.
Praslin, reer, on Hypnea pannosa.
A fragmentary specimen that 1 was unable to name.
RoscHera, Sonder.
1. RoscHERA GLOMERULATA (Ag.), n. comb.
Tolypiocladia glomerulata, Schmitz, Marine Florid. von Deutsch Ost-Afrika, 1895, p. 160.
Seychelles, Long Island ; dry specimen, fragment.
Distribution. Zanzibar; Indian Ocean; Malay Archipelago.
%
290 PERCY SLADEN TRUST EXPEDITION
A small fragment, attached to Gelidiopsis variabilis. In the ‘Siboga” paper I hope to
point out that Schmitz was perfectly right, when he supposed that Roschera africana,
Sonder, and Tolypiocladia glomerulata belonged to the same genus. I have therefore
made use in this paper of Sonder’s older generic name.
Herposrpnonta, Naegeli.
1. HerPosiPHONIA PROREPENS (Harv.), Schm.
Harvey, Phycol. Australica, tab. 185 (B).
Chagos Archipelago.
Distribution. Algoa Bay ; Western Australia.
After describing the present plant, Harvey adds: ‘“‘ This alga was first described from
Algoa Bay, where it occurs on corallines. The Australian plant is more slender, with
fewer tubes, but otherwise the same.” The Herposiphonia on Dasyopsis Geppii has
8 pericentral tubes; it stands therefore nearest to Harvey’s Australian form. My
material is too scanty to make out whether these forms are specifically distinct from
the Algoa plant, as some authors have thought probable.
OxigocLtADuS, Weber-van Bosse.
1. Oxtcoctapus PRarinit*, n. sp. (Plate 18. fig. 31.)
Thallo filiformi repente, ramis normaliter dorsaliter exeuntibus, erectis, endogenis,
ramulos unilaterales ferentibus. Filo constante ex uno ordine cellularum centrali
et ex quatuor ordinibus cellularum pericentralibus. Trichoblasti desunt. Filis
latis 120-200 pu, ramulis 12-80 ». Organis fructificationis ignotis. Rhizoidis
unicellularibus.
Coetivy, reef, on Dasyopsis aperta; in alcohol.
The alga that I have named Oligocladus Prainii belongs to the subfamily of the
Herposiphonez, and is very nearly allied to the new genus Oligocladus, provisionally
described in ‘Annales du Jardin botanique de Buitenzorg,” 1910, p.31. The specimens
were found growing among the penicilli of Dasyopsis aperta and are barren; the
determination is therefore not quite certain, but in anatomical structure these algze so
closely resemble O. Boldinghii, Web. v. B., that I feel almost sure that they belong
to this genus. I may also add, that in a few long branches I have seen a con-
densation of protoplasm in two cells of each segment. This reminded me of the
two sporangia in each segment in Oligocladus Boldinghi, but in the present case I
could not detect the slightest trace of division in the protoplasm. They were, however,
too young, and I must leave it to future investigators to settle the question.
O. Praini is a creeping filamentous alga with four pericentral cells, dorsally placed,
endogenous branches, and unicellular exogenous ventral rhizoids. Thus far it is lke
O. Goldinghw. It differs from this alga in having no trichoblasts, but a naked, straight
* Named in honour of Sir David Prain, in remembrance of a lovely walk in the Botanical Gardens at
Calcutta.
WEBER-VAN BOSSE—MARINE ALG A 291
apical cell, and a much denser ramification. The absence of trichoblasts is no reason
to separate O. Prainit from the genus Oligocladus, for in the genus Polysiphonia we
find species with and without these organs.
The main axis of the plant was attached by means of its long rhizoids to the
branches of Dasyopsis, and, as may be seen in fig. 31, the youngest part raises itself
from its support, grows vertically upwards, though its apex shows a slight curve. The
apical cell is rather long and cuts off the segments at its base by a transverse wall.
These segments remain undivided for a long time—I have counted eight of them under
the apical cell before the pericentral cells are begun to be cut off.
The dorsally placed branches arise irregularly with a distance of 3, 4, 6, 7, 8, 9, and
even more segments between them; they bear branches of the second order on the side
turned towards the main axis. In some cases I observed near the apex of the main axis
long ramified branches alternating with short-simple ones, but in later stages I could
detect no further difference. Perhaps the long ones bear the tetrasporangia and the
short ones remain sterile, but this is a mere suggestion. The first segment of the
branches issuing from the main axis has only short pericentral cells; the second
segment has pericentral cells of the normal size; all the branches of the first and
second orders arise from the central tube. ‘The size of the branches and of the main
axis differs considerably according to the place where the measurement is taken. The
main axis has a breadth of 120-2004; the branches 40-80 in the lower parts, but at
the top only 12-20; the segments are 13 times as long as the diameter, and at the
top as long as broad.
At the top of the main axis all the branches are arranged on the dorsal side of the
filament, but at some distance from the apex their position is sometimes altered ; this
is the result of a slight twisting of the cells of the main axis.
2. OLIGOCLADUS PUSILLUS, 0. sp.
Thallo filiformi, repente ; ramis erectis, endogenis, simplicibus, normaliter dorsaliter
exeuntibus. Filisconstantibus ex uno ordine cellularum centrali et quatuor ordinibus
cellularum pericentralibus. Trichoblastis numerosis, filorum apices cingentibus.
Filis latis 40-100 ». Organis fructificationis ignotis. Rhizoidis unicellularibus.
Coetivy, reef, on Dasyopsis aperta; im alcohol.
Oligocladus pusillus, on account of the numerous trichoblasts that surround the
slightly curved top, stands nearer to O. Boldinghii than O. Pram. It is a smaller
plant, though its filaments are broader. Its creeping filaments give off side-branches
on the dorsal side at a distance of 5, 7,12, 20 segments and even more. ‘These
ascending side-branches remain simple; when, however, they develop into creeping
filaments, as occasionally happens, they likewise give off branches on their dorsal
side. The creeping branches have a diameter of 100», the side-branches a little less,
and at the top of both the diameter sinks to 40. The segments have a height of
80-100 p, but this sinks at the top to 20 » and even less.
Organs of fructification have not been observed.
292 PERCY SLADEN TRUST EXPEDITION
Dasyopsts, Zanardini.
1. Dasyopsis STANLEYI, n. sp. (Plate 17. fig. 16.)
Thallo elongato, subdichotomo, pauce ramoso, penicillis longis, gracilibus cincto;
ramificatione radiata, constante e penicillis basim versus deciduis, spinas non
relinquentibus. Stichidiis oblongis et cystocarpiis ex penicillis orientibus. An-
theridiis non visis. Thallo alto 16cm. MHabitu Dasye elegantis.
Cargados Carajos, 30 fms.; in alcohol and dry specimens.
To rightly understand the new species of Dasyopsis, it may be worth while to give
a short account of the characters of the genus. Dasyopsis has a sympodial mode of
growth; the stem or axis being composed of short branches with definite growth ;
these are pushed aside by their own first branch, and the distance between the base of
the mother-branch and the point of insertion of its first branch varies in different
species.
But this is a feature Dasyopsis has in common with all Dasyeze; its special character
consists in the absence of true pericentral cells. The cells that surround the central
tube are not originally segments of this tube, as in the other Rhodomelacee, but they
are downward-growing hyphe, the mother-cell of which is cut off from the branch that
is pushed aside when sympodial growth sets in (Pl. 18. fig. 32, m). The hyphze may be
of the same diameter as the central cells, in which case they resemble true pericentral
cells, but they have quite a different origin. The displaced branches continue to
develop, they are shorter or longer, naked or corticated, according to the species, and
carry several monosiphonous filaments, the so-called penicillus, at the apex.
Dasyopsis Stanleyi consists of a subdichotomously divided principal axis or stem,
carrying at indefinite intervals branches of unlimited growth, which may ramify again
and are surrounded at the top by monosiphonous penicilli. This pretty alga so much
resembles Dasya elegans that I felt at first inclined to take it for that species; after
careful examination, however, I discovered that the plant has the characteristic
structure of Dasyopsis; the branches of definite growth are separated by only one
short segment from each other, and grow out in any direction from the main axis. As
every branch or so-called penicillus gives off a downward-growing hypha, the central
axis soon increases in thickness ; true pericentral cells are of course absent.
The penicilli are composed of cells having a breadth of 6-13 ,; their length varies
from being almost square at the base to 72 at the top. They are very delicate and
soon fall off, unless they happen to bear cystocarps or stichidia at their base. ‘The
stichidia were very rare in my preparations; I observed only one but unfortunately
lost it; I saw, however, that it was long and cuspidate. The cystocarps were numerous
in fertile specimens.
2. DASYOPSIS APERTA, n. sp. (Plate 17. fig. 17; Plate 18. fig. 32.)
Thallo ex axibus principalibus primo surgentibus, postea decumbentibus constante, ende
denuo rami adscendentes nascuntur. Apice axis principalis et ramorum penicillis
ramulosis, monosiphoniis cincto, penicillorum pilis postea decidentibus, basi
WEBER-VAN BOSSE—MARINE ALG A 293
monosiphonio brevi tantummodo restante. hallo sympodiali, radiato; ramis
cum divergentia {. Primo segmento libero ramorum ad basim satis magnam
cellulam incremento basipetal disjungente. Cellulis centralibus magnis, hyphis
numerosis angustis cinctis. Stichidiis ramulis penicillorum sessilibus; 8-10 sporas
im unoquoque segmento ferentibus. Cystocarpiis et antheridiis ignotis.
Coetivy, reef ; in alcohol.
Cargados Carajos, 47 fms.; dry specimen.
This new Dasyopsis attains a height of 2°5 cm., and consists of a small tuft of primary
ascending cylindrical axes or stems, that finally become horizontal and give rise to new
ascending ones. The tufts, on growing older, become more or less intricate, but I
never saw the fronds anastomose. Dasyopsis aperta is a good species for the study of
the peculiar Dasyopsis structure, because the cells are large, even at the apex of the
shoot. It is sufficient to spread the alga on the slide and to tear off carefully with
a needle the surrounding penicilli to see plainly the row of central cells, each carrying
a pushed aside apex and its first branch whose lowest cell builds up the sympodium.
These branches stand spirally around the main axis, so far as I could ascertain, with
a divergence of 4, and carry long, incurved filaments at their top, which form together
the peuicillus. The first cell, after the branch has been pushed aside, cuts off a cell at
its base (Pl. 18. fig. 32, m); this is the mother-cell of the downward-growing hyphex,
which are at first almost as large as the cells of the central axis or tube. At the top ofa
stem or axis, where the mutual position of the different cells is still unaltered, the central
tube appears to consist of a central and four pericentral cells, but on following the
pseudo-pericentral cells to their origin, it will be clearly seen that they are downward-
growing hyphe, arising from the first free cell of the pushed aside branch. The sym-
podial axis is afterwards covered by a thick layer of smaller hyphz, which by growing
between them alter the original position of the cells.
The stem or main axis is cylindrical, though tapering towards its apex, which is
densely covered with penicilli; these, however, soon fall off, and the base of older stems
is almost denuded, though not entirely, for remnants of the basal portion of the penicilli
often remain.
The sessile stichidia are borne on the side-branches of the penicilli; they are
cylindrical and, when the tetrasporangia have acquired their full size, a little torulose.
I counted from 6-8 sporangia in different segments; and the whole is crowned by one
or two barren cells. Neither cystocarps nor antheridia were seen.
There are, according to Falkenberg*, two other species of Dasyopsis, which, like
D. Stanleyi and D. aperta, have penicilli facing every way, namely Dasyopsis spinella
and D. cervicornis. From the first both D. Stanleyi and D. aperta differ in the absence
of the little spines that have given D. spinella its specific name, and from D. cervicornis
in the shape of their stem, cylindrical throughout, not angular at the insertion-point
of the penicilli.
* Falkenberg, “ Die Rhodomelaceen des Golfes yon Neapel,” 1901, p. 667.
SECOND SERIES—ZOOLOGY, VOL. XVI. 38
294 PERCY SLADEN TRUST EXPEDITION
3. Dasyopsis GEppit, n. sp. (Plate 17. figs. 18-20; Plate 18. fig. 33.)
Thallo carnoso, plano, palmato, inciso, horizontaliter expanso, sympodialiter con-
structo, ad apicem radioso ; divergentia ignota. Penicillis lateralibus tantum bene
evolutis, dorsalibus et ventralibus aliquamdiu conspicuis, postea evanescentibus.
Ramis lateralibus pro parte cum axi principali coalitis. Strato corticali cellularum
rotundarum aut angulosarum contiguarum. Antheridiis cylindricis in turmis in
facie inferiori thalli sitis. Cystocarpiis et stichidiis ignotis.
Amirante, 20-25 fms.; in alcohol.
Cargados Carajos, 25 fms.
Chagos Archipelago, Salomon Atoll, reef; in alcohol.
Coetivy, reef.
Dasyopsis Geppit has a broad, flat, fleshy frond, slightly palmatifid and attached to
the substratum by a thick pedicel (Pl. 17. figs. 18, 19). Its outward appearance
differs entirely from any known species of Dasyopsis, and it is only by the study of
the youngest segments of the frond that an insight into its anatomical structure is
obtained. This is in principle like that of Dasyopsis, but differs in so far as the
displaced branches, instead of turning in every direction, as they do in D. Stanleyi,
spread themselves principally in a bilateral direction and grow congenitally into a
leaf-like frond. The penicilli are situated on the margin of this frond, but also on its
upper and lower surface; the cells of the filaments have a breadth of 16-28 , and
a length of 32-88, 100, and 140 pz.
At the top of the different segments the penicilli are so closely packed together, in
consequence of the smallness of the cells, that it is impossible to trace the central
tube down into the frond or to observe the divergence of the penicilli around its apex
(Pl. 18. fig. 33).
Dorsi-ventral organization does exist, though I could not trace it at the apex of the
frond. The ventral side, that turned towards the substratum, bears antheridia at
the base of old penicilli. The antheridia stand in groups of five and six together
(Pl. 17. figs. 19, 20), and are borne on a monosiphonous pedicel. The dorsal side of the
frond is barren in all my specimens. The position of the antheridia is interesting,
for D. Geppii is both a reef and a deep-water plant. If it were only collected on the
reef, one might infer that the antheridia kept to the ventral side to ensure ‘protection
from the sun, but since these alge have been found with ventrally placed antheridia,
not only on the reef but also in deep water, this explanation does not hold good.
Dorsi-ventral organization of any kind was hitherto unknown in Dasyopsis,
I have great pleasure in dedicating this alga to my kind friend, Mr. Gepp, of the
British Museum.
4. DASYOPSIS PALMATIFIDA, n. sp. (Plate 17. fig. 21.)
Thallo carnoso, plano, profunde palmatifido, segmentis aut laciniis seepe terminan-
tibus in filamento cylindrico, substrato bulbo affixo. Segmentis ad apicem radiatis,
postea horizontaliter expansis. Penicillis circum apicem thalli bene eyolutis,
infimis evanescentibus. Penicillis constantibus e filis multo crassioribus quam
WEBER-VAN BOSSE—MARINE ALGA 295
penicillis Dasyopsis Geppii; filis stichidia in apicibus pedicellorum brevium
monosiphoniorum circumdantibus. Stichidiis sporis numerosis. Cystocarpiis et
-antheridiis ignotis.
Amirante, 20-25 fms.; in alcohol.
Chagos Archipelago, Salomon Atoll; in alcohol.
Coetivy, reef; in alcohol.
Dasyopsis palmatifida resembles in essential points D. Geppiz, but its frond is far
more deeply palmatifid and the different segments often terminate in a cylindrical
branch. ‘These fasten themselves to any hard object and produce at that spot a little
tubercle, which much resembles the thick pedicel of D. Geppii. I have considered
the question whether D. palmatifida might not be the stichidia-bearing form of
D. Geppii, but have been unable to solve it. In addition to its deeply palmatifid frond,
D. palmatifida differs also from D. Geppi in the size of the filaments forming the
penicilli; these cells having a length of 104-106 and a breadth of 12-20, 36-40 w.
Stichidia with tetrasporangia are borne at the base of the penicilli on monosiphonous
pedicels ; they have a blunt apex.
TAPEINODASYA, Weber-van Bosse.
1. TAPEINODASYA ETHEL, n. sp. (Plate 17. figs. 22, 23.)
Thallo procumbente, constante e sympodio bilaterali; tertio quoque segmento aat
ramulum lateralem spinosum incremento finito ferente, aut ramulum lateralem
spinosum incremento sympodiali infinito. Ramulis adventitiis preeterea ex axillis
ramulorum spinosorum emergentibus sed parvis etiam adultis. Axi constante
e cellula centrali cum quatuor cellulis pericentralibus ; quo numero verumtamen
cellularum pericentralium in segmentis spe majore ob concrescentiam ramulorum.
Axi centrali cellulis corticalibus, peripheriam versus diminutis, cincto. Hyphis
numerosis in parietibus inter cellulas pericentrales nascentibus. Organis fructi-
ficationis ignotis.
Amirante, 20-25 fms.; dry and in alcohol.
Tapeinodasya Ethele is the second known species of this genus; the type, 7. Bornetii,
being described in the “Recueil des Travaux botaniques Néerlandais,’ No. 1, 1904. The
plant from Amirante differs in having a less dense ramification, which is, as a rule,
bilateral. The branches have the outward appearance of little spines. At the top of
the main axis they point outwards in three directions, but immediately below the
erowing-point the branches stand in two rows. In my preparations the distance
between the succeeding branches in the sympodium is three cells, but I should not be
surprised if in parts of the frond the distance amounted to only two cells, as in
the other species, Z. Boruetii, the number is known to vary. ‘The branch that is
displaced ends in a spine; it is either of definite growth and develops no further, or of
indefinite growth, in which case it may give rise toa lateral sympodial branch. The
branches grow congenitally with the main axis for one or two segments, and in a
cross-section through such a segment a greater number of pericentral cells is visible.
38—2
296 PERCY SLADEN TRUST EXPEDITION
The central cell is surrounded normally by four pericentral cells, but between these
other large cells soon appear and numerous hyphe develop in the membranes. I have
not seen them enter the pericentral cells as they do in 7. Bornetit, but it may be that
this takes place in the basal part of the frond. A layer of small cortical cells surrounds
the main axis. The latter has a winged appearance on account of the pericentral cells
of the spine-like lateral branches growing sideways together with it for a few segments.
Adventitious branches appear occasionally on the dorsal side of the frond in the axil
of lateral branches, but they remain short; in 7. Bornetii these branches attain full
development.
The characteristic dorsi-ventral organization of 7. Bornetii is far less conspicuous in
T. Ethele, and shows itself in barren specimens only in the rudimentary axillary
branches. No organs of fructification were present, and it was only because I was
familiar with the genus that I was able to name the specimens from Amirante.
HETEROSIPHONIA, Montagne.
1. HerrrostpHontIa RENDLEI, n. sp. (Plate 16. figs. 11, 12.)
Thallo fasciculato, pluribus ramis principalibus e callo basali surgentibus; ramis
principalibus totis ecorticatis, pauce ramosis, penicillos ad segmentum quodque
secundum, distichos et alternantes gerentibus. Cellulis pericentralibus 7—S-10.
Penicillis ramulos unilaterales et stichidia potius obtusa quam acuta ferentibus
tetrasporis instructis. Antheridiis et cystocarpiis ignotis. Thallo alto 8 cm.
Cargados Carajos, 47 fms.; dry specimen.
I believe Heterosiphonia Kendlei* to be a new species because none exists amongst
the Heterosiphonie ecorticate that is exactly like it. It belongs tothe small group with
more than four pericentral cells, the branches or penicilli of which arise at a distance
of two or three segments from each other and are turned regularly to the right and
left. H. Rendlei stands nearest to H. subsecundata (Suhr), Falk., but differs in the
pericentral cells not being transversely divided.
The principal axis has a height of about 8 em.; it is erect and carries small horizontally
spread penicilli which have unilateral monosiphonous branchlets incurved at the top.
The number of pericentral cells in the main axis varies from 6-10, and the segments
are three times as broad as long. In the penicilli the few basal polysiphonous segments
are almost as long as broad or a little broader.
I have seen neither cystocarps nor antheridia, only stichidia, which are borne on the
side-branches of the penicilli; they are elongated with a blunt apex.
2. HETEROSIPHONIA spec.
Coetivy, reef; dry specimen.
This is a small specimen, 1 cm. high; it grows on Codiwm. I failed to recognize it
amongst the described species and yet do not dare to describe it as new.
* Named in honour of Dr. Rendle, Keeper of the Botanical Department of the British Museum, renowned for
the kind hospitality that it offers to every visitor.
WEBER-VAN BOSSE—MARINE ALG 297
AMPHISBETEMA, 0. g.
Thallus constat e pede crasso rhizomatoideo, unde nascuntur axes erecti, corticati,
regulariter sympodii modo compositi e ramis incremento finito conspicue mono-
podialibus dorso-ventraliter crescentibus. Axis constat ex una cellula centrali et
sex cellulis pericentralibus, tribus aut quatuor ordinibus cellularum longitu-
dine cellule primariz pericentralium cinctis; quos ordines stratum corticale
erassum e cellulis parenchymaticis fissis compositum cingit. Cystocarpia et
-antheridia ignota. Stichidia sporis verticillatis, eque ac in Dasya non omnino
cellulis corticalibus obtectis.
1. AMPHISBETEMA INDICA (J. Ag.). (Plate 17. fig. 24; Plate 18. fig. 34, and text-fig.)
Dasya indica, J. Agardh, Till Alg. Syst. xi. 1890, p. 111.
Diagnosis the same as that of genus.
Aldabra, reef, Passe Houareau, outside; dry specimens.
Distribution. Diego Garcia.
The alga for which I propose the generic name of Amphisbetema was described in
1890 by J. Agardh under the name of Dasya indica; the structure of the stichidia,
almost identical with that of Dasya, probably induced this eminent algologist to
describe the plant under that name, for he was quite cognisant of the differences in
ramification between his alga and other Dasye. In De Toni’s ‘“‘Sylloge Algarum,”
iv. p. 1146, 1908, Dasya indica has received the name of Wilsonea (?) indica.
I found this alga in the collection of Mr. Stanley Gardiner, and through the kindness
of Sir David Prain I was able to compare it with authentic material from Diego Garcia,
now preserved in the Herbarium at Kew, and I wish to express my best thanks to him
for so liberally placing this material at my disposal. The study of the tvpe-specimens
enabled me to identify my barren material from Aldabra. This differs only from the
type in being a little stronger; the alge from Diego Garcia bear stichidia with
tetrasporangia, a fact which may account for their being more slender.
Amphisbetema has a strong rhizome-like base from which arise erect main axes,
attaining a height of 8-9 cm., with distichous bilateral branches. These main axes
are composed of branches or shoots of definite growth, succeeding each other at a
distance of six segments to be then displaced by their own first branch. The branches
have, so far as I could make out, monopodial growth; sympodial growth sets in only
when each in turn is pushed aside. All the branches have a dorsi-ventral organization ;
their first branch of the second order is always inserted at a distance of six segments
from the base, and all the succeeding ones at a distance of two segments; they stand
in two rows on the ventral side of the branch, and only by subsequent growth do they
ultimately take their bilateral position. The branches of the second order have the
same ramification as the primary ones and are always turned to one side; each
principal branch carries as a rule six pairs of branches of the second order; but the
nearer to the apex the less do they develop.
After the formation of these six pairs of branches, the vigour of each mother-
branch seems to be exhausted. At a distance of three or four cells from the apex,
998 PERCY SLADEN TRUST EXPEDITION
a segment remains undivided : its contents, balled together, look like a large drop of oil
and have a reddish-brown colour. The first segment succeeding this undivided cell may
show a division into a central and pericentral cells, the next is undivided, and the
ultimate one has usually the shape of a spine. Sometimes the top of a branch is
crowned by a simple or ramified monosiphonous filament. I could find no explanation
for this difference in development; the monosiphonous filaments are not trichoblasts,
nor could any trace of these organs be detected. The large undivided cell, with the cells
above it, falls off after a shorter or longer period, and the branch that carried them is
blunt in consequence.
Schematic figure of apex of Amphis-
betema indica, showing the sym-
podial growth of main axis and the
monopodial growth of the displaced
shoots. After 6 segments shoot I
is pushed aside by its first branch,
shoot II is also displaced after
6 segments by its first branch, and
the other shoots follow in like
manner. The displaced shoots
carry side-branches at every second
segment.
The axes have in each segment a central and six pericentral cells; in the last-formed
side-branches this number may sink to four, but as a rule it is six. The pericentral cells,
in common with those of Endosiphonia and Melanothamnus, cut off to the outside cells
quite as long as themselves; and this may be repeated several times. In the branches
the cells on the dorsal side are a little longer than those on the ventral, but this difference
disappears with subsequent growth. The whole central strand is surrounded bya rather
thick layer of parenchymatous cells. The peripheral cells are of equal size and elongated
longitudinally, not radially as in Melanothamnus. Though the cortical layer is pretty
WEBER-VAN BOSSE—MARINE ALG A 299
thick, the transverse membranes of the succeeding segments within are plainly visible,
owing to the fact that the central strand consists of a large number of cells of equal
length. The segments are short, and this fact gives, whenever the apex of a shoot is
pushed aside, the impression that the main axis is divided into three branches. Careful
examination will, however, show that this impression is erroneous, being brought about
by the displaced shoot, its first branch—the new shoot—and its second branch standing
very close together, owing to the short segments and the congenital growth of the first
two segments.
My plants are barren: J. Agardh, however, described the stichidia, which he stated
were like those of Dasya and bore verticillately developed tetraspores partially uncovered
on the outside.
Amphisbetema bas received its name* on account of the difficulty experienced in
assigning to it its true systematic position. It has characters in common with severai
genera of Rhodomelaceze and also with Melanothamnus, Born. et Falk., a genus of
uncertain position so long as the organs of fructification remain unknown.
With Streblocladia, Amphisbetema agrees in the regular alternating mode of mono-
podial and sympodial growth; with Melanothamnus and Endosiphonia in the numerous
secondary pericentral cells of equal length; with Heterosiphonia in the dorsi-ventral
organization of its branches ; and with Dasya in the structure of the stichidia.
The association of all these characters in one genus makes me inclined to consider
Amphisbetema as an old form of Rhodomelaceze which existed before the genera that
have now developed one or more of the characters—in our new genus still blended
together—in their own special way and to the exclusion cf the other characters.
Dicryurws, Bory.
1. DicTyURUS PURPURASCENS, Bory.
Bory de St. Vincent, in Bélanger, Voy. Indes orient. p. 170.
Falkenberg, Die Rhodomelaceen, 1901, p. 675.
De Toni, Syll. Alg. vol. iv. sect. 111. 1903, p. 1173.
Saya de Malha, 55 fms.; dry specimen.
Coetivy, reef; drv specimen and in alcohol (fragment).
Cargados Carajos, 30 and 47 fms.; dry specimen and in alcohol.
Ayirante, 30 fms. ; in alcohol.
Chagos Archipelago, Salomon Atoll, 22 fms.; in alcohol.
Distribution. Cape Comorin ; Coast of India; Mauritius; Ceyion; Malay Archi-
pelago.
Fam. CERAMIACE.
GrirFirusia, C. Agardh.
1. GRIFFITHSIA spec.
Anmirante, 20-25 fms.; in alcohol.
Only a small fragment, but with the characteristic tetraspores of the genus.
* apudeoPirnia, controversy, a point in dispute.
300 PERCY SLADEN TRUST EXPEDITION
k HALopLeGMA, Montagne.
1. HatnorpreGmMa PREeEtsstiI, Sond.
Sonder, Alg. Preiss. p. 24.
Harvey, Phye. Austr. 1859, tab. 79.
De Toni, Syll. Alg. vol. iv. sect. 111. 1903, p. 1366.
Cargados Carajos, at 22, 30, and 47 fms.; both dry and in alcohol.
Distribution. Western Australia ; Tasmania, in the River Tamar above Georgetown ;
Malay Archipelago.
The specimens of H. Preissvi from Cargados Carajos are shorter, less branched, and
more flabellate than most plants found in the herbaria under that name. I have not
ventured, however, to describe these specimens as new, for I have found no microscopic
character to distinguish them from the type, the size of the main branches and the
peripheral filaments being essentially the same in the different plants. The specimens
from Cargados Carajos bear tetraspores; I have not seen cystocarps or antheridia, and
it is possible that these may show differences in structure. Dr. Harvey, in his ‘“ Phye.
Australica,” however, calls attention to the fact that the external form of H. Preissz is
very variable. He found it growing in the River Tamar, a considerable distance above
Georgetown, where it appeared at first to be a distinct species ; but he was able to trace
it down to the Heads of Port Dalrymple, and found that it blended into the usual variety.
I feel very much inclined to think that the specimens from Cargados Carajos owe their
small size in great measure to the depth at which they were gathered.
ANTITHAMNION, Naegeli.
1. ANTITHAMNION ADNATUM, J. Ag.
J. Agardh, Analecta algologica, 1892, p. 12.
Chagos Archipelago, Salomon Atoll.
Distribution. New Zealand; Malay Archipelago.
A small specimen with tetraspores, attached to Herposiphonia.
Spyripia, Harvey.
1. SpyRIDIA FILAMENTOSA (Wulfen), Harv.
Harvey in Hooker, Brit. Flora, vol. ii. p. 337 ; Phycol. Brit. 1871, pl. xlvi.
Aldabra reef, outside Passe Houareau.
Distribution. West Indies; Mediterranean; Red Sea; Indian Ocean; Malay
Archipelago.
Only a small fragment attached to Amphisbetema indica.
CrrAmtum (Roth), Lyngbye.
1. CERAMIUM CINNABARINUM (Gratel.), Hauck.
Hauck, Die Meeresalgen von Deutschland, 1885, p. 112.
Chagos Archipelago, Salomon Atoll.
Distribution. Mediterranean; Malay Archipelago.
A small specimen amongst other alge.
WEBER-VAN BOSSHE—MARINE ALG Ad 301
2. CERAMIUM spec.
Cargados Carajos, 47 fms.
On Heterosiphonia Rendlei; barren’ specimens.
REINBOLDIELLA, De Toni.
1. REINBOLDIELLA SCHMITZIANA (Reinb.), De Toni.
Phycez japon. nove, 1895, p. 35.
Gloiothamnion Schmitzianum, Reinbold, in Hedwigia, 1895, p. 205.
Chagos Archipelago; Egmont, reef, in alcohol, on Gelidiopsis: Salomon Atoll, on
Gelidiun. :
Distribution. Japan; Malay Archipelago.
Fam. GRATELOUPIACEA.
Hatymenta (C. Ag.), J. Agardh.
1. Hatymenta PonyciaDa, A. & EH. 8S. Gepp.
Var. ALDABRADENSTS, n. var.—Thallo e callo basali subito cuneatim expanso 10 cm.
alto, 4-5 em. lato deinde palmati-partito constante ; ramis usque ad 6 em. longis,
05-1 cm. latis, ramulis lateralibus iterum iterumque subdichotomicis pinnatis,
angustis.
Aldabra, outer reef; dry specimens.
Distribution. Christmas Island.
Halymenia polyclada belongs to Agardh’s Acanthymenia section of Halymenia.
According to Schmitz, it is highly probable that this group is identical with the
genus Gelinaria, Sond. Schmitz is further of opinion that the differences existing
in anatomical structure are sufficient to separate Acanthymenia from the genus Haly-
menia. So long, however, as we do not know the cystocarps of the genus Gelinaria
I think it more expedient to follow Agardh, although ultimately we will probably have
to follow Schmitz and call the species of the Acanthymenia group by the generic name
of Gelinaria.
The specimens from Aldabra are very similar to H. polyclada, which is nearly allied
to H. formosa. It has a subdichotomous ramification, but lateral ramuli are plentiful
along the margins of the upper branches. In the lower part of the fronds the margins
are almost entire. It is a stronger plant than H. polyclada, but has the same short stalk
expanding suddenly into a broad frond. How far H. formosa, Durvillei, and polyclada
are really good species, is a question that future investigations must decide.
SECOND SERIES—ZOOLOGY, VOL. XVI. 39
302 PERCY SLADEN TRUST EXPEDITION
CryPTonemta, J. Ag.
1. CRYPTONEMIA SEMINERVIS (Ag.), J. Ag.
J. Agardh, Algee Liebman. p. 11 in not.
De Toni, Syll. Alg. vol. iv. sect. 1v. 1905, p. 1610.
Saya de Malha, 55 fms.; dry specimens.
Amirante ; dry specimen.
Distribution. Mediterranean; Atlantic ; Red Sea.
in a dry condition the plants appear to have no midrib, but ou moistening a delicate
midrib became distinctly visible, and this extended upwards to about the middle of the
leaf. The plants are barren.
2. CRYPTONEMIA spec.
Saya de Malha, 55 fms.; dry specimen.
A poor fragment, but differing from Cryptonemia seminervis by its much thicker frond
and dark red colour.
Fam. SQUAMARIACE A.
Tt is much to be regretted that so many of the specimens of Squamariaceze collected
by Mr. J. Stanley Gardiner are sterile, for without fruit it is almost impossible to name
the genera and species of this puzzling family. I happened to have studied a large
collection of Indian Peyssonnelie before I undertook to name the present collection, and
this has helped me to recognize some of the species mentioned hereafter.
In my “ List of the ‘ Siboga’ Algze” I will treat in detail of the Peyssonnelie, and only
mention here that I have found it of great help to keep up a distinction already hinted at
by the late Dr. F. Schmitz*, namely, the distinction based on the differences in arrange-
ment of the filaments of the hypothallus. These horizontal filaments, which run over
the substratum, give off the ascending vertical filaments of the perithallus, and Schmitz
observed that some hypothalli consisted of straight filaments running close to one
another (Peyssonnelia), and that others consisted of curved fan-shaped groups of filaments,
as in the basal layer of Cruoriella. Tio these two subgenera, if I may distinguish them
provisionally by such a name, I wish to add a third, differing in having no hypothallus
proper, but a mesothallus, 7. e., a layer of cells occupying the middle of the thallus (as is
the case, for instance, in Ralfsia expansa), which gives off branches both downward and
upward. My investigations do not yet allow me to decide whether this division of the
genus Peyssonnelia, based as it is upon anatomical characters, is supported by differences
in the structure of the fruit. Sometimes this is the case, but my specimens are mostly
barren.
For this third subgenus I should like to propose the name of Héhelia, in honour of my
dear friend, Mrs. E. 8. Gepp.
* Schmitz, ‘‘ Marine Florideen von Deutsch Ost-Afrika,” in Engler’s Bot. Jahrbuch, Bd. xxi. Heft 1, 2, 1895,
p. 173.
WEBER-VAN BOSSE—MARINE ALG
PryssonNELiA, Decaisne.
Subgen. HuPryssonneELra, n. subg.
Hypothallus filis arcte junctis, flabelli modo fusis, maxime erectis,
1. PEYSSONNELIA COCCINEA, J. Ag.
J. Agardh, Epicrisis Floridearum, 1896, p. 385.
De Toni, Syll. Alg. vol. iv. sect. 1v. 1905, p. 1695.
Seychelles, 31 fms. ; dry specimen.
Saya de Malha, 55 fms.
Distribution. West coast of Australia; Malay Archipelago.
303
The specimen is small and sterile; it was only by comparing it with “ Siboga ” material
that I was able to name it.
2. PEYSSONNELIA HARVEYANA(?), Crouan. (Plate 18. fig. 35.)
Crouan in J. Agardh, Spec. Algarum, 11. 1852, p. 501.
De Toni, Syll. Alg. vol. iv. sect. rv. 1905, p. 1694.
Saya de Malha, 55 fms., on several dry, calcareous, little pebbles.
Distribution. Coast of France, near Brest.
The specimens are of a bright red colour and adhere firmly to the substratum; they
approach both P. conchicola and P. Harveyana, but on account of the large cells of the
hypothallus I feel inclined to regard them as P. Harveyana. With regard to distribution,
it is more likely that they belong to P. conchicola, an inhabitant of the Red Sea.
The specimens are all barren, and it is impossible to name them with certainty.
3. PEYSSONNELIA CaLCcEA, Heydr.
Heydrich, ‘‘ Neue Kalkalgen von Deutsch Neu-Guinea” in Biblioth. Bot. von Frank u. Luerssen
1897, p. 10.
Egmont, reef; in alcohol.
Peros, Tle du Coin ; in alcohol.
Distribution. Tami, near German New Guinea; Malay Archipelago.
The specimens bear nemathecia with tetraspores.
4, PEYSSONNELIA POLYMORPHA (Zan.), Schmitz.
7
Form GarpIneEri, n. f.—Thalio primo adherente, deinde a substrato soluto, tubera
forma diversa, sed superficie levi formante, colore parum rubro cum siccatus est.
Hypothallo cellulis longis 36-44-60 n, latis 8-10-16 ». Perithallo constante e
seriebus cellularum sursum directarum, cellulis infimis majoribus, cellulis superiori-
bus minoribus, longioribus aut brevioribus quam latis; latis 16-20-24 p.
fructificationis ignotis.
Amirante, 30, 45-60 fms. ; dry specimens.
Organis
The specimens from Amirante are nearly allied to P. polymorpha, but the loose thalli
are smaller and the crusts are thinner than in the typical specimens of that species. The
anatomical structure of the two algze is much alike, as far as the vegetative thallus is
39—2
304 PERCY SLADEN TRUST EXPEDITION
concerned, and reproductive organs were not to be found. ‘The cells of the f. Gardineri
are perhaps, on the whole, somewhat smaller than those of the type, and seen from above
f. Gardineri is covered by a smooth peripheral layer, which P. polymorpha had lost in
all the specimens I have seen.
I conclude that f. Gardineri is a deep-water form of P. polymorpha.
Subgen. Ernpnia, n. subg.
Mesothallus filis rectis aut ramosis, sinuatis, utroque latere fila perithalli emittens.
5. PEYSSONNELIA BIRADIATA, 0. sp.
Thallo per substratum extento, tantum hic illic adheerente, coriaceo, rubro, rotundo,
diametro 2 ad 3. centimetrum, constante e mesothallo et perithallo. Mesothallo
constante e filis ramosis, sinuatis, parva flabella efficientibus. Perithallo constante
e filis sursum et deorsum directis. Perithallo summo diviso in partem inferiorem
obliquam, cellulis magnis, et in partem superiorem erectam, hic illic zonatam,
cellulis parvis. Organis fructificationis ignotis.
Seychelles, 31 fms. ; dry specimen.
Distribution. Malay Archipelago.
A small but characteristic specimen.
Among the algze from Saya de Malha there are several specimens that I believe belong
to the genus Peyssonnelia, but they are unfortunately barren. They have much in
common with the Peyssonnelie of the Hthelia-group, for they have a mesothallus that
gives off branches both in a downward and upward direction (PI. 18. fig. 36). They are
furthermore much calcified, hard as stone, and attached to bits of coral.
But besides these features found among the “thelia-group, the Peyssonnelie from
Saya de Malha are characterized by long filaments (PI. 18. fig. 37) almost free from each
other and covered by a layer of pectic substance, which colours red with ruthenium.
The iilaments are like those we find in the sori of other Peyssonnelie, but in the new
plants they are not confined to sori alone, but cover the whole upper surface. Although
I made sections through all parts of the thallus, I could find no trace of fructification.
We know that the fronds of Stragularia species amongst the Ralfsiacezee are covered
by filaments, and it appears probable that we have here something analogous amongst
the Peyssonnelie ; but until the reproductive organs are discovered it is difficult to
speculate as to these structures. A doubt stiil lurks in my mind whether, after all,
these filaments may not belong to an encrusting parasitic alga; but it is well worth
while to call attention to these specimens.
Cruortopsis, Dufour.
1. Cruortiopsis crucIATA, Dufour.
Dufour, Elenc. Alg. Lig. p. 35, after De Toni.
De Toni, Syll, Alg. vol. iv. sect. rv. 1905, p. 1689.
Saya de Malha; dry specimen on Udotea.
WEBER-VAN BOSSE—MARINE ALG A 305
Distribution. Adriatic; Malay Archipelago.
The alga is barren, but its resemblance to specimens from Nias, Sumatra, bearing
cystocarps and antheridia, is so great that I believe it belongs to the same species.
Fam. CORALLINACEA.
Me topesta, Lamouroux.
1. MELOBESIA CALLITHAMNIOIDES, Falk.
Falkenberg, Die Meeresalg. d. Golfes v. Neapel, 1878, p. 265.
Egmont, reef, on different algze ; in alcohol.
Distribution. Atlantic, Brest? ; Mediterranean.
AmpuHtroa, Lamouroux.
1. AMPHIROA FRAGILISSIMA (Linn.), Lamx.
Lamouroux, Pol. flexibles, p. 300.
A. Weber-van Bosse and M. Foslie, “The Corallinacez of the ‘ Siboga’ Exped.,” Monogr. Ixi.
1904, p. 89.
Chagos Archipelago, Salomon Atoll; in alcohol.
Mahé, Cap Terné; dry specimens.
Distribution. West Indies; Pacific; Malay Archipelago; Dar-es-Salaam.
2. AMPHIROA FOLIACEA, Lamx.
Lamouroux, Voyage de l’Uranie par Freycinet, Zoologie par Quoy et Gaimard, p. 628.
A. Weber-van Bosse and M. Foslie, “The Corallinaceze of the ‘ Siboga’ Exped.,’” Monogr. 1x1.
1904, p. 92.
Praslin ; in alcohol.
Distribution. Marianne Islands ; Malay Archipelago.
JANIA, Lamouroux.
1. JANIA TENELLA, Kitz.
Kiitzing, Tab. Phyc. Bd. viii. 1858, tab. 85, p. 41.
Coetivy, reef ; in alcohol.
Peros, Tle du Coin ; in aleohol.
Praslin ; in alcohol.
Seychelles; dry specimens.
Mahé, Cap Terné; dry specimens.
Distribution. Malay Archipelago; Gulf of Mexico.
306
PERCY SLADEN TRUST EXPEDITION
EXPLANATION OF PLATES 16-18.
Puate 16.
Vig. 3. Gloiophlwa articulata, n.sp. xX &.
2. Hucheuma Cotton, n. sp. Nat. size. Two forms.
3. Calliblepharis prolifera, J. Agardh. x 3.
4. Chylocladia perpusilla, n. sp. x 2.
5. Fertile portion of a swollen ramulus of Chylocladia perpusilla. x 68.
6. Laurencia pygmea, 0. sp. ' X 2.
7. Cladhymenia oblongifolia. x #.
&. Chondria pumila, Vick. x 3.
9. Chondria simpliciuscula, n. sp. X 3.
10. a Pa Top of frond, with regular rows of hexagonal cells. x 80.
11. Heterosiphonia Rendlei, n. sp. x 3.
12. Penicillus of Heterosiphonia Rendlet, with stichidia. x 24.
PLATE 17.
Fig. 13. Pseudendosiphonia Gardineri, nu. gen. et sp. X 3.
14. Top of frond of Ps. Gardineri. x 12. st.=stichidia.
15. Stichidium of Ps. Gardinert. x 100.
16. Dasyopsis Stanleyi,n.sp. x &.
17. Dasyopsis aperta, n. sp. x 3.
18. Dasyopsis Geppii, un. sp. X 3.
19. if = Underside of thallus, with clusters of antheridia. x 4.
20. os 3 Antheridium.
21. Dasyopsis palmatifida, nv. sp. xX 3.
22. Tapeinodasya Kthele, n. sp. x 3.
23. e. Part of frond. x 11.
24. Amphisbetema indica (J. Ag.). Nat. size.
25. Galaxaura hawatiana, Butt. Nat. size.
PuaTeE 18.
Fig. 26. Gloiophiea articulata, u. sp. Section through apex of frond. x 82.
27. i Peripheral filament, showing small cortical cells. x 340.
28. Gloioderma? expansa. Nat. size.
29. 55 a Transverse section, showing large central cells. x 40.
30. Pseudendosiphonia Gardinert. Section through thallus. x 16. c.=central tube; p.c.=peri-
central cells.
31. Oligocladus Prainti, n. sp. x 32.
32. Dasyopsis aperta, u. sp. Part of penicillus. x 46. m. = mother-cell of downward-growing
hypha.
33. Dasyopsis Geppii, nu. sp. Young displaced branch. x 180.
34. Amphisbetema indica. Apex of frond. X 24.
35. Peyssonnelia Harveyana. Transverse section. X 128.
36. Peyssonnelia ? subgen. Ethelia. Section through thallus ; highly magnified.
37. - Young plant, section through upper part of thallus ; highly magnified.
Percy StapEen Trust Expeprrion.
TRANS. LINN. SOC. SHR. 2. ZOOL. VOL. XVI. PL. 16.
(WEBER-VAN Bosse)
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MARINE ALGA FROM THE INDIAN OCEAN
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7
No. XV.—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA,
SEPSIDA, MICROPEZIDA, DROSOPHILIDA, GEOMYZIDA, MILICHID Ai.
By C. G. Lamp, M.A., B.Se., Clare College, Cambridge.
(CoMMUNICATED BY PROFESSOR J. STANLEY Garpiner, M.A., F.R.S., F.LS.)
(Plates 19—21 and 48 Text-figures.) .
Read 4th March, 1914.
THIS paper concludes the description of the Acalyptrate muscids of the Seychelles
expedition. Among the families treated of there are several forms which, in the rather
fluid state of the classification, must be regarded as possibly needing relegation to other
or even to new groupings when the world forms are more fully known. Hence it has
been thought desirable to refer fairly fully to the chetotaxy even in individuals which
are placed in genera already founded. To save space the following abbreviations are
used :
f. 0. fronto-orbital bristles. prescut. prescutellar bristles.
1. v. inner vertical bristles. p. a. post-alar bristles.
o. v. outer vertical bristles. s. a. supra-alar bristles.
p. v. post-vertical bristles. i.a. intra-alar bristles.
oc. ocellar bristles. st. p. sternopleural bristles.
post orb. the row of post-orbital bristles. mesopl. mesopleural bristles.
d.c. dorso-central bristles. acr. the acrostichal row.
h. humeral bristles. div. divergent.
n. p. the two notopleural bristles. conv. convergent.
presut. presutural bristles.
In describing the head a systematic method has been adopted: the insect is
adjusted in the field of the binocular microscope by means of a gimbal-like stand which
enables all possible rotations and translations to be made. It is first adjusted to view
the vertex perpendicularly, this is called the Top View; it is then twisted about to
view the face perpendicularly, and this is called the Front View; finally it is adjusted to
look at the side of the head, and this is the Side View. ‘These views correspond to the
Plan and Elevations of the engineer's drawings. The structures seen in each view are
then described in detail and in order. This method obviates the troublesome process of
continually shifting the position of the insect when comparing it with a given
description.
SECOND SERIES—ZOOLOGY, VOL. XVI. 40
308 PERCY SLADEN TRUST EXPEDITION
The detailed figures in the plates have been reduced from large scale camera lucida
drawings; the insect was in all cases carefully set in the proper plane by means of
sighting the orbital bristles and the vibrissee etc. till they just came over one another.
Most of the drawings are thus exact views of the individual insect’s head ; in some cases
the insertions only of the bristles are shown, giving a more diagrammatic view.
Most of the wings were photographed from dry slide specimens; a large scale was
adopted from which the figures were reduced: in the case of a unique specimen, or a very
small set in which one specimen could not be spared for the mount, a large scale camera
lucida drawing was made as for the heads.
Since nearly every species has its wings figured, no description of the venation is (in
general) given in the text. In some cases the wings of species that have already been
described by others are figured where no such figure exists in the original papers*.
Heteroneuride.
HETEROMERINGIA Ozerny, Wien. Ent. Zeit., xxi. (1903), p. 72.
1. Heteromeringia plumicornis, n. sp. (Fig. 1 and Plate 19, fig. 1).
This species is a very pale form, and from the description and figures of Hendelia
Beckeri, Cz. (Czerny, t. c., p. 84 and Taf. 2, figs. 1, 2, 3) must have a considerable super-
ficial resemblance to that species.
Head. Eyes bare. Top view :—frons entirely orange and shining, paler towards
the front: the small central area between the three ocelli is shiny black. Hind margin
somewhat concave. Chetotaxy normal; front f. 0. b. touching at tip; next pair parallel;
hind pair somewhat divergent; 1. v. parallel; 0. v. divergent; p. v. finer, long, much
divergent, inserted just on vertex very close together; cc. fine, divergent, inserted
between front and side ocelli; post oc. row longest just at back of head. All the bristles
somewhat orange. Front view :—antennz well separated at base, face about square ;
usual two convergent vibrissee with minute following row: face all pale yellow, slightly
hollowed. Side view (Plate 19, fig. 1) :—likewise all yellow in colour as are the palpi
and the thickened hairy tongue. Antenne yellow, except that the upper basal part of
third joint is black: the second joint has an elegant calyx-like form and a well-marked
spine ; the arista is very stout, black, and profusely haired all round with stout black hairs.
Thorax. Orange, shining, covered with fine bristles, and with two brown stripes
extending from near the front to the scutellum. Pleuree shining, darkened from below
humeri to abdomen, scutellum a little darkened laterally on the disk. Two d. «. b. on
each side along the inner margins of the brown stripes; one h., two n. p., one i. a., one
s. a. bristle. Scutellum with two small dorsal bristles and two large subparallel terminal
ones.
Wings, as Fig. 1, glassy with brown veins; a little darkened at tips. Halters orange.
Legs, pale yellow, all the tibiz and tarsi somewhat infuscate. Front femora with a
regular row of spines on lower face, middle femora with a similar strong row beneath ;
* A first set of the material, including the Typxs of all new genera and species, will be placed in the
British Museum. A second set will be retained in the Cambridge University Museum.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDAL, TRYPETID AM, SEPSIDA, ETC. 309
the pre-apical tibial bristles are small except on the middle pair of legs which have also a
stout terminal tibial spine ; the legs all clothed with small black bristles.
Abdomen. Dark shining brown, covered with long bristly yellowish hairs. The
male has a well-marked knobbed hypopygium with the usual pair of scale-like
appendages.
Size, including antenne, 2°8 mm.; wing, 2°1 mm.
Localities. Seychelles. Silhouette: Mare aux Cochons, IX. 1908; low coconut-
planted country near Pointe Etienne, 17. [X. 1908. Mahé: Anonyme Island, I. 1909.
Fig. 1. Heteromeringia plumicornis, n. sp. x 35.
2. Heteromeringia mgrifrons, nu. sp. (Fig. 2 and Pl. 19, fig. 2).
This is a black somewhat shining species and is more like the normal forms of the
genus than is the last one.
Head. Eyes bare. Top view :—hind margin distinctly concave; frons uniformly
and entirely shining black; cheetotaxy normal; front f. o. b. touching, next two pairs
parallel; i. v. converging; o. v. fairly divergent; p. v. very divergent, inserted just on
vertex between hind ocelli; these bristles are black; oc. b. very minute and pale. Front
view :—antennz nearly touching at base; vibrissee touching, a row of fine following
bristles; face all yellow-orange. Side view (Plate 19, fig. 2):—jowls orange in front
getting blacker to hind of head which is shining black behind the eyes. Post-ocular
bristles extending along top of eye. Hind jowls with bristles, a few long ones on hind
margin of head. Antennz all clear orange yellow, except the flagellum of the arista
which is darkened; the 2nd joint has one upper and two lower bristles and a row of
minute marginal bristles; the pubescence of the arista follows the aristal colour and is
normal. The finely bristled palpi are much flattened in the horizontal plane; the distal
third is orange the rest black ; tongue orange.
Thorax. Entirely black, the dorsum and scutellum very slightly roughened, pleura
and metanotum very shining. The chtotaxy is normal: two principal d.c. b., a third
rather smaller just in front of the front pair, h., two n. p., strong s. a. and p. a.
Dorsum covered with fine bristles, except behind. Scutellum with two long terminal b.,
and two small dorsal near them. A stout mes. p. and st. p. b. Legs yellow except the
end of the front femora, all the front tibize and tarsi, and the hind tibize (except tips)
which are black ; all the femora with rows of bristles: front femora with a close set lower
row and a row of 4 or 5 longer ones behind; middle femora with two rows below, front
40—2
310 PERCY SLADEN TRUST EXPEDITION
short and stout, back long and stout, hind femora with lower row on distal third; very
stout spur on middle tibia. Legs all bristly ; front tarsi distinctly flattened.
Wings, as Fig. 2, slightly infuscate, especially round the distal 2ths of the 2nd vein.
Halters with infuscate heads and yellow stalks.
Abdomen similar to the thorax, clothed with black bristly hairs, a small pair of
scale-like appendages are visible, somewhat remote (vertically) from the tip.
Males only were present.
Size 3 mm.; wing, 24 mm.
Localities. Seychelles. Silhouette: Mare aux Cochons plateau, over 1000 feet, IX.
1908. Mahé: Cascade Estate, ca. 800 feet, 1908—9.
Fig. 2. Heteromeringia nigrifrons, n. sp. x 30.
3. LHeteromeringia mgriceps, n. sp.
Two specimens, one ¢ and one ?, were present, which were practically identical
in their principal structural details with the previous species. The differences
in colour are, however, very marked, and in default of evidence of dimorphism
in the males it is thought best to describe them as a distinct species, at least
provisionally.
$ Head. The face is entirely black instead of yellow, so that there is no pale colour
at all on any part of the head, except that the lower eye-margins are slightly orange in ©
some lights. The tip of the third antennal joint is blackened: only the extreme tip of
the palpi is orange.
Legs. The whole of the front femora, except the extreme tips, is black : the mid and
hind femora have, basally, a slightly darkened ring.
The ? has the same differences in the head colours but the front femora are only
blackened at the tip, and the others have no rings.
Size as last species.
Locality. Seychelles. Mahé: Cascade Estate, 800 feet or over, 1908—9.
ALLoMETOPON Kertész, Ann. Mus. Nat. Hung., iv. (1906), p. 320.
A single specimen, in a rather battered condition, can be referred with practical
certainty to this genus. The genus was described from a New Guinea species.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA@, SEPSID@, ETC. 311
4. Allometopon flavum, n. sp. (Fig. 3 and Plate 19, fig. 3).
The species is entirely shining yellow except for slightly browner lines on the thorax,
brownish tarsi, and the deep black third antennal joint and arista. The bristles are all
distinctly yellow or brownish.
The cheetotaxy agrees fairly with Kertész’s generic description, except that the ocellar
bristles are not strikingly long.
Head. Eyes bare. Top view :—frons orange, finely punctulate, darker and smoother
on vertex round ocellar area: ocellar patch shining, eye-margin rather shining. Four
f. o. b., all subparallel; nearly parallel i. v., divergent o. v., all these bristles arise from
small papillee: p. v. small, divergent, inserted just behind middle of hind ocelli ; oc. small,
divergent, inserted just on ocellar patch. The crossed bristles on front of frons are
comparatively stout.
In front view the antennz are seen to touch at base; the face is very narrow and
short. In this view the stout subcylindrical yellow palpi are easily seen. Side view
(Plate 19, fig. 3) :—the antennz much as Kertész’s description and figure l.c., p. 331. The
orange 2nd joint is beautifully calyx-like, with bordering spines, one at the top being very
long. The rounded rather flattened black third joint has a long blackish, distinctly
pubescent arista. A stout vibrissa, followed by a row of marginal mouth bristles ; orange
jowls, bristly behind, with one very long yellow bristle quite behind: six large post-orbit.
bristles on top.
Fig. 3. Allometopon flavum, n. sp. x 20.
Thorax. Shining orange, with three narrow brown-orange stripes, evanescent
behind: surface all covered with small bristles. The chzetotaxy agrees fairly with
- Kertész’s description: two hind d. c. b., a much smaller one in front; two prescut. ; h.
and n. p.are visible. The rounded orange scutellum has very fine pale pubescence : divergent
terminal bristles, basal side bristles (scars only visible), 4 discal (on one side an extra
one in the actual specimen).
Wings, Fig. 3: yellowish with brown veins ; costa with strong double rowed bristles.
Halters, orange.
Legs. The front femora are strongly spined beneath from apex to base: the
middle femora are also strongly spined in a row behind, the spines becoming progressively
shorter from base to apex; the hind femora are unarmed ; the middle tibia has two very
stout black end-spurs.
Abdomen. Orange with rows of short discal, and long marginal bristles.
312 PEROY SLADEN TRUST EXPEDITION
Size, 34 mm.; wing, 3} mm.
Locality. Seychelles. Praslin; Cétes d’Or Estate, XI. 1908.
Ortalide.
This family has been revised recently by Dr Hendel, and in the course of his work
the material from the British Museum was placed at his disposal; this included several
new species from the Seychelles. Only one further undescribed species is therefore
dealt with here, namely a member of the genus Acrosticta (Loew): this genus is an
interesting occurrence, since it has hitherto been known only from North and South
America, with the exception that one species (A. pallipes Grimshaw) is common both
to Bolivia and Peru and Hawaii (see Hendel, Wien. Ent. Zeit., xxviii. (1909), p. 249).
Platystomine.
PLAGIOsTENOPTERINA Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 3.
5. Plagiostenopterina ruficeps Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 3.
Wing, Fig. 4.
9 specimens.
Fig. 4. Plagiostenopterina ruficeps, Hend. x 15.
Localities. Seychelles: Silhouette, coast (La Passe), 1908: Mahé, VII. 1908 (Long
Island). Aldabra: 1908—9 (Fryer). Assumption: 1909 (R. P. Dupont).
prone : Ea RARE SIN ate mes SE gO
Fig. 5. Plagiostenopterma cyanosoma, Hend. «x 15.
LAMB—DIPTERA: HETERONEURID, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 313
6. Plagiostenopterina cyanosoma Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 4.
Wing, Fig. 5.
13 specimens.
Localities. Seychelles: Silhouette, plateau of Mare aux Cochons, over 1000 feet,
IX. 1908: Mahé; Cascade Estate, 800—1500 feet, 1908—9 ; Forét Noire district, 1908 ; GILA/ y,
slopes of Morne Seychellois, over 1500 feet, 4. II. 1909; 1907 (Thomasset) ; 1905. Ave® NOx
Aldabra: 1907 (Thomasset). ie eee
7. Plagvostenopterima submetallica Loew, Bericht. Akad. Berl., 1852, 660. 29.
Wing, Fig. 6.
Locality. Seychelles. Mahé: Cascade Estate, 800—1500 feet, 1908—9.
Also known from various parts of Africa.
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Fig. 6. Plagiostenopterina submetallica, Loew, x 15.
Scuotastes Loew, Mon. Dipt. N. Amer., vol. iii. (1873), p. 38.
8. Scholastes cinctus Guérin, Voyage Coq., Zool., 299.
Localities, Seychelles: Mahé; Cascade Estate, ca. 800 feet, 1908—9 ; Long Island
(near sea-level), VII. 1908. Chagos Islands: Salomon Atoll and Diego Garcia, 1905.
9. Scholastes vicarius Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 4.
Locality. Seychelles. Mahé: Anonyme Island (near sea-level), I. 1909.
Pararpewio Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 1.
10. Parardelio pilosa Hendel, l.c., p. 2. Wing, Fig. 7.
Fig. 7. Parardelio pilosa, Hend. x 20.
Locality. Seychelles. Mahé: 1906, a number of specimens (Thomasset); from
near Morne Blanc, X.—XI. 1908; Cascade Estate, 800—1500 feet, 1908—9.
314 PERCY SLADEN TRUST EXPEDITION
Ulidine.
PseuDEUXESTA Hendel, Genera Insectorum, 106, p. 30.
11. Pseudeuxesta prima Osten-Sacken, Ann. Mus. Gen., xvi. p. 470. Described
from Celebes and New Guinea. Wing, Fig. 8.
Localities. Seychelles. Mahé: from beach, Long Island, VII. 1908, 10 specimens ;
from near Morne Blanc, X.—XI. 1908, 1 specimen.
Fig. 8. Pseudeuxesta prima, Ost.Sack. «x 15.
Acrosticta Loew, Berlin. Ent. Zeitschr., xi. (1867), p. 293.
See Hendel, Wien. Ent. Zeit., xxviii. (1909), p. 247.
12. Acrosticta exsul, n. sp. (Fig. 9 and Plate 19, figs. 4, 5).
This species approximates to A. scrobiculata Loew.
Head. Eyesbare. Top view:—hind boundary flat; vertex rather shining greenish-
black, finely punctate ; broad triangular eye-margins the same; frons with parallel sides,
from tip of ocelli to forehead rather shining brownish-red, paler in front. The forward
eye-margins, beyond f. o. b., are narrowly silvery. Frons pitted all over, except just in
front of ocelli: each pit carries a fine bristly hair, the centre ones converging. Front
eye-margins bordered by a regular outward bent row of such bristles. The bristles
on paler part in front bend forwards, the others are irregularly oriented.
Fig. 9. Acrosticta exsul, n. sp. x 20.
There are two upper f. o. b. bristles on the shiny upper eye-margins, equal sized
verticals, i. v. bent towards one another, the o. v. outwards; divergent oc. between
front and back ocelli, divergent p. v. well below vertex; a double row of post-ocular
bristles on upper half of the hind eye-margins.
S|
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDH, ETC. 315
Front view (Plate 19, fig. 4) :—antenne well separate at base, face orange in that
place, eye-margins well marked, densely pollinated: upper lip shining greenish-black,
as is the protuberant clypeus. Side view (Plate 19, fig. 5) :—face deeply excavate, lower
edge of broad lower eye-margins with a row of bristles: the bristly hind jowls and
the back of head all grey pollinated. The bristly tongue is retracted in the specimen,
but it can be seen that the palpi are orange: there is a hind lobe to the tongue which
is chitinised like the clypeus and is also greenish-black.
The antenne are darkish orange, the second joint singly spined above, the almost
bare flagellum of the arista alone being blackish.
Thorax. Somewhat shining greenish-black with regular rows of fine hairs. The
pleura are shining bluish-black, very slightly scrobiculate. The specimens are rather
damaged, but the following bristles can be made out: hind d. ¢., hum., n. p., 2 s. a., p. a.
Scutellum bare, subtriangular, coloured as thorax; divergent terminal and smaller
basal bristles.
Wings, Fig. 9, slightly suffused, with brown veins, darkened at tip, on stigma
and below it to 2nd vein. Halters, knobs yellow, stalks darkened at the base. Legs
entirely dark reddish-black except about the distal third of the femora and the knees
which are orange, the under surface of the femora (when legs are stretched horizontally)
shows signs of a purplish chitinisation. The legs are covered with small spines, and on the
front femora beneath the spines are longer and form a regular row. All the femora (in
the dried specimens) are remarkably flattened sideways. There is a strong black spine
at the end of the middle tibia.
Abdomen and metasternum coloured as the thorax, the 2nd and 3rd segments
making up most of the length, the surface with sparsely scattered black bristles.
Two °.
Size, 34 mm.; wing, 3} mm.
Locality. Seychelles. Mahé: from near Morne Blanc, X.—XI. 1908.
Curysomyza Fallen, Scenop. 1. (1817).
13. Chrysomyza enea Fabr., Ent. Syst., iv. (1794), p. 335; Hendel, Zool. Anz.,
xxxlv. (1909), p. 614.
Localities. Seychelles. Mahé: Cascade Estate, ca. 800 feet, 1908—9: very common
in the plantations, sitting on leaves of bushes, &c.
14. Chrysomyza azwrea Hendel, Wien. Ent. Zeit., xxxi. (1912), p. 5.
Localities. Seychelles: Silhouette; from near Mont Pot-a-eau, ca. 1500 feet, and
Mare aux Cochons plateau, over 1000 feet, VIII.—IX. 1908: Mahé; from near sea-level
(Long Island), VII. 1908; Forét Noire district, X.—XI. 1908; Cascade Estate,
ea. 800 feet, 1908—9. Chagos Islands; Salomon Atoll, 1905.
In the Ann. Soc. Ent. France, lxiv. (1895), p. 366, the species Chrysomyza demandata
Macq. is recorded by Gigho-Tos.
SECOND SERIES—ZOOLOGY, VOL. XVI. 4]
316 PERCY SLADEN TRUST EXPEDITION
Trypetide.
Crratitis MacLeay, Zool. Journal, xvi. (1829), p. 482.
15. Ceratitis capitata Wied., Anal. Entom. (1824), 55 (Trypeta).
Localities. Seychelles: Mahé; Cascade Estate, and Mare aux Cochons district
(1000 feet or over), 1908—9: Dennis Island, VIII. 1908: Praslm, a number of
specimens from an introduced creeper (Passiflora fetida), 30. XI. 1908. Amirantes :
Poivre Island, 10. X. 1905, ¢ and 2 i covrtu.
Oxyna Robineau-Desvoidy, Myodaires, 755, i. (1830).
16. Oxyna sororcula, Wied., Aussereur. Zweifl. Ins., ii. (1830), 509. 52.
Described by Wollaston from Madeira as Ensina vacillans, Ins. Dipt. Mad.,
A.M.N.HL, i. (1858), 117.
Localities. Seychelles. Mahé: Port Victoria, from grass in Botanic Gardens, XII.
1908; Cascade Estate, ca. 800 feet, 1908—9; country above Port Glaud, XI. 1908;
Mare aux Cochons district, 1000 feet or more, 1909. Silhouette: plateau of Mare
aux Cochons, and coconut-planted country near coast (Pointe Etienne), IX. 1908.
Acrp1a Robineau-Desvoidy, Myodaires, 720. 14 (1830).
17. Acidia seychellensis, n. sp. (Fig. 10 and Plate 19, fig. 6).
Fig. 10. Acidia seychellensis, n. sp. ~ 20.
A single female specimen of this genus was collected ; generically it agrees well with
the British species but the eyes occupy a little more of the side of the head.
Head. Eyes with a very few almost invisible hairs. Top view :—frons slightly
widening from vertex to forehead, chestnut brown, slightly shining, the short strips
on which upper f. o. stand and the black ocellar area together with a small depressed
triangle in front of it are quite shining. The duller part is very minutely shagreened
and has scattered tiny hairs in front. Three pairs crossed front f. 0., the corresponding
absolute eye-margins with a row of tiny hairs along them ; two pairs parallel but backward-
bent upper f. 0. Long slightly conv. i, v. just at eye-angle on vertex; div. o. v.
just on hind eye-margin; roughly forming a triangle with these is a small bristle
behind the head: in this view the long post-orb. row can be seen running up to the
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDM, ETC. 317
outer v.b. ‘The p. v. parallel and rather bent backwards, inserted on the sloping back
of vertex, the distance between each and the next 1.v.is the same. Oc. fine, subparallel,
inserted each side of the large front ocellus. Front view :—face brown, same width
as frons to just below 2nd antennal joint, then widening uniformly to jowls. The
lunular crescentic furrow continued round and demarcating eye-margin from face. A
very shallow broad triangular keel forms two shallow fovee. Antenne almost touching
at base, divergent. Side view (Plate 19, fig. 6) :—antennze orange, the 2nd joint is very
bristly inside, outside smooth, neat bordering row of bristles below, a spine about midway
above ; 8rd joint long, straight upper and curved lower profile; arista with pale pubescent
lower joints, darker flagellum with short hairs all round in no definite rows. The
narrow front jowls with short bristle below, orange; hind ones deeper, universally
bristly with one long spine: hind head below black and bristly. Palpi flattened
laterally, very bristly below, orange: tongue the same. The shining clypeus is visible.
Thorax. Dorsum, pleura, scutellum and metanotum all shming black, the dorsum
covered with fine hairs: the chetotaxy as Acidia cognita though possibly not relatively
so strongly developed: there is a pair of d.c. rather forward, being roughly in a line
with wings; long prescut., h., n. p., prescut., a large post-a.; the pleura has fine sparse
bristles, but the pinning renders the macrochetes doubtful though a sterno-pl. can be
seen. Scutellum a blunt shallow triangle, the terminal bristles crossed at tip, the large
side pair ending off nearly level with them.
Wings, as Fig. 10, with brown-black coloured marks, Ist and 3rd veins bristly above.
Halters with black knobs and orange stalks.
Legs. All the coxee and femora black ; the rest yellow-orange, except that the basal
half of the hind tibia is blackened. Front femora with the two hind rows of spines
and the long lower row: middle with long spinous hairs; hind with similar hairs,
more conspicuous below at base, and one or two at tip on top. Hind tibie with
longish spiny hairs, a somewhat regular row dorsally.
Abdomen. Black, pointed, with adpressed hairs and long marginal bristles,
ovipositor not extruded.
Size, 4mm.; wing, 3? mm.
Locality. Seychelles. Silhouette: plateau of Mare aux Cochons, IX. 1908.
18. Acidia(?) ocellata, n. sp. (Fig. 11 and Plate 19, figs. 7, 8).
There is a single specimen of a fine Trypetid from Marie Anne Island which does
not seem to agree well in its characters with any of the well-known genera. Its general
affinities seem nearest Acidia, and hence it is provisionally referred to that genus
pending a revision of the whole family. The description will therefore be given in full.
Head. Eyes under high power with a very few fine scattered hairs: frons
widening a little from back to front. Top view :—frons, vertex (as well as hind head)
entirely pale orange, dull except for the ocellar area and the upper and lower eye-
margins, which are shining: the latter have about 3 irregular rows of fine bristles.
On them stand the three lower crossed f. 0. pairs, the middle one being crossed almost
41—2
318 PERCY SLADEN TRUST EXPEDITION
half-way ; the two upper f. 0. pairs are sub-parallel and bent backward. V. b. on distinct
pits, the i. v. very long, sub-parallel and bent a little backward, inserted just behind upper
eye angle, the o. v. long and div., inserted in a line with post-orb. row, a little remote
from eye: the latter row is seen to be stout and consists of bristles alternately longer
and shorter. Well below on back of head is a smaller bristle forming a triangle
with the v. b.; the p. v. are shortish, stout, and well below vertex, they diverge. The
oc. are stout, div. and are inserted one each side of the front ocellus. There are small
scattered bristles on the frons. The lunular arch is demarcated from the frons by a
red shallow suture. Front view (Plate 19, fig. 7):—much as last species, but the
central ‘“‘keel” is merely a broader triangular swelling merging below with the lip.
Side view (Plate 19, fig. 8):—the pale orange antennz have the second joint bristled
inside, bare outside, with marginal bristle row there and below, a long spine on base at
tip; third joint parallel-sided; the arista quite bare under a high power. The
jowls are pale orange, hairy and bristly below with a long stout spime just
behind lowest point of eye. The head immediately behind eyes is bare, but bristly
behind. Palpi orange, flattened, bristly below. The tongue is retracted in the
specimen.
Fig. 11. <Acidia (?) ocellata, n. sp. x 13.
Note. The specimen has an extra f. o. b. on the left side in front of the normal
front bristle.
Thorax. Orange, and covered with very many irregularly arranged dorsal bristles ;
the pleuree, &c. yellow. A pair of d. ¢. further behind than last species, large prescut.,
very strong h., n. p., presut. low down, three strong alar (two on edge of disc, one in
line with prescut.). Scutellum with strong basal bristles near angle, the terminal pair is
broken off, metanotum yellow-orange. There is a strong st. pl. br.
Wings, Fig. 11. The striking feature about the wings, as seen without a lens, is
the two large blackish ocelli occupying a large part of the wing, and bordered almost all
round by a whitish border ; the rest of the wing is suffused with honey coloured pigment,
which borders the costa and forms a rough V-shaped mark behind the black spot.
Under the microscope the costa is seen to be very bristly; a strong spur occurs at
the breach by the auxiliary branch; the Ist vein is stoutly bristled, as is the third
from its junction with the second to a point about 2 the distance between the
cross vein and the end. ‘The anal cell is well poimted. The small calypters are
beautifully bordered with dark margins and silky hairs. The whole hind wing-margin
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 319
is also very neatly ciliated in black. Halters orange. Legs all orange: front femora
with the usual two back rows of bristles and a very stout row of 6 bristles below:
the coxae have two very stout spines and a few others: tibize spurred. The middle
femora are hairy; the tibie have a row of 5 stout though comparatively strong
spines in the middle part (somewhat nearer base) on tip, a crown of orange spines
with one excessively strong black one below; all the tarsal joints have the curious
stout spines at the basal sides of each joint that are found in A. cesio (Harris). The
hind femora have 3 or 4 stout spines on the apex in front, tibize with a central row
of some 7 spines in front: the long Ist tarsal joint has beautiful golden hairs on it:
each joint has two basal spines, smaller than those on the middle legs.
Abdomen. Orange at base merging to black towards the end of 2nd segment : 2nd
segment very long, next two about equal, last about as long as previous two ; triangular
in profile. The globular hypopygium is orange: it is complex, and in profile can be seen
a pair of upper scales and a pair of lower more pointed ones.
The last segment of the abdomen bears a border of very strong downward pointing
spines; the rest of the abdomen is covered with short bristly orange hairs, the marginal
ones being scarcely longer.
Size, about 7 mm.; wing, about 64 mm.
Locality. Seychelles. Marie Anne Island, from forest, 2. XII. 1908.
TepHRitTis Latreille, Hist. Nat. Crust. Ins., xiv. (1804), 389.
19. Tephritis aldabrensis, n. sp. (Fig. 12 and Plate 19, fig. 9).
Fig. 12. Vephritis aldabrensis, n. sp. x 20.
Head. Eyes bare. Top view :—frons orange-yellow, rather shining but
minutely granulate: margins and ocellar area and vertex a little greyer. Lunule
demarcated by a red arc. Frons quite parallel-sided: in this view the Ist and
2nd joints of antenna are seen to be very bristly, the former with 3 or 4 longish
pale bristles, the latter with many small black ones, chetotaxy normal, viz. 3 pairs
rather con. front f. 0., two pairs parallel and backwardly bent hind f. o., the pair nearer
vertex being smaller and crossing the very stout parallel upright 1. v. ; 0. v. smaller, white,
p- v. the same, parallel and well down head; between the o. v. and p. v. is another
similar bristle, and these bristles (viz. o. v., p. v., extra pair) are continuous with the
very stout post-orbital row (extending half-way down eyes) and like them are white.
320 PERCY SLADEN TRUST EXPEDITION
The oc. are very long and arched, div. and inserted one each side of front ocellus. There
are practically no tiny bristles on the head. Front view :—face yellowish-white, no
foveze or keel ; antenne orange, the overlap of 2nd joint swollen inside, the 3rd joint long,
oval and rounded at tip, the arista with pale base and darker flagellum, the latter
just perceptibly pubescent. The antenne are just separated at the base by a narrow
projection from the lunule over the antennz. Side view (Plate 19, fig. 9) :—mouth margin
projecting somewhat, the sides bristly as is edge of jowls and the swollen hind jowls;
the latter have a long bristle on them. The post-orb. row can be seen to have small
bristles between the large ones. Palpi flattened, pale yellow and hairy.
Thorax. Ground colour dark and covered with grey pollen in a good specimen, and
also dorsally with tiny white bristles, which are, however, long just over the scutellum,
cheetotaxy normal, being all very long: two d. c., one just behind cross suture, one
a little further than half-way from front pair to end of thorax. A very stout bristle
and a clump of small stubbly ones on the orange humeri, large and small h., presut.,
two s.a., one long and smaller ones by the mesopl. suture, one st. pl.; the pleurz all with
bristly hairs except in front. The bare pollinated scutellum has a rather small
terminal pair, crossed half-way, and excessively long parallel side pair. The wings
are maculated as in Fig. 12, the darkening being bright brown, paler towards the
hind margin. Halters yellow with orange stalks. Legs all orange; the lower row
of bristles on front femora very stout and long.
Abdomen. Brown, slightly shining, covered with pale hairs. The ante-penultimate
segment in the male is nearly as long as the three previous ones: in top view it is
bordered by some 5 or 6 long pale bristles. The hypopygium below the last segment
is shining chestnut brown. The terminal segments of the female are shining black
as is the long dorsally flattened ovipositor sheath, which is longer than the rest of
the abdomen.
Size, 35 mm.; wing, 3 mm.
Locality. Aldabra, 1908 (Fryer).
Ruaspocuata de Meijere, Bydr. t. d. Dierkunde, xvii. (1904), p. 109.
20. Rhabdocheta spinosa, n. sp. (Fig. 13 and Plate 19, fig. 10).
A considerable number of an insect occurred which seems to fit in best with de Meijere’s
genus, though it will be seen that the characters do not agree absolutely with his
definition of the genus.
Head. Eyes bare. Frons about as broad as long, nearly parallel-sided, the ridge
of the vertex from eye to eye very sharp; orange in front, greyish-yellow round
ocellar area and eye-margins: lunular are with flattish ridge and linearly darkened :
ocellar spot black. The bristles are of two characters, (a) long darkish ones coming
gradually to a point (viz. the two main front f. o. and the inner y.): the others (@) are
white, long or short, but roughly cylindrical up to the tip where they attenuate much
more suddenly, or are even greatest in diameter about half-way. There are two long
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 321
true (a) front f. o., upright and conv. to centre; in front (above a black side spot on
the face) is an auxiliary bristle (type 8). Two very slightly div. and backward bent
hind f. 0. : the distal one is practically of type a and is inserted inside the proximal front
f. o. so that the distal hind f. 0. and proximal front f. 0. are nearly in the same transverse
line: the proximal pair of hind f. 0. is of type 8. I. v. of type a, very long, upright though
slight conv. and bent back, inserted close to eye and absolutely on the vertical ridge.
What represent the outer v. are of type 6 and are a little down the eye orbit pointing
back and separated from the stout post-orb. row (of type 8) by a few tiny black
bristles. P. v. of type 8, below vertex, almost touching at tip, bent up. Between
each of these and the o. v. is another bristle of type 8. Oc. very long, of type B
though more pointed, straight, div., inserted practically in a line with front ocellus
but with bases a little further from it than hind ocelli are from one another: the oc.
are so long that they cross the hind f. 0. at the ends. On the frons about 3 down
from front ocellus is a parallel pair of type 8 projecting over the forehead. Apart
from the macrochetes the frons is bare. Front view:—face very gently arched,
the eyes approach nearest about 3 down from antennal base to mouth margin.
Antenne touch at base and in this view the 2nd joint is seen to have a small internal
finely bristled flap over the base of 3rd. The mouth margin is much curved in outline
with the upper lip just projecting bluntly from the curve: face all yellow except the
side margins of mouth which are brown lined. In this view the black spots between
Fig. 13. ARhabdocheta spinosa, n. sp. x 25.
the eyes and antennal bases on each side below f. 0. are well seen, as are the blackened
compressed bristly palpi. Side view (Plate 19, fig. 10) :—the triangular yellow projection
of frons, with the above mentioned spot, is very prominent: the profile is concave
and the projection of the lip well seen. At lower angle of jowl is a bristle of type B
followed by a pointed one of type a. The jowls are yellow merging into the blackened
back of head, the lower part of latter with pale bristles. The antennee are orange,
2nd joint a little greyer: below, the 2nd joint has a fringe of pale marginal bristles
but is practically bare on the outside (see however above, front view); 3rd joint very
long, roughly pyramidal with pointed tip, pubescent and darkened at tip: arista with
orange basal joints and paler flagellum: it is closely covered with moderately long pale
hairs. Palpi as above, tongue orange, hairy at tip.
322 PERCY SLADEN TRUST EXPEDITION
Thorax. In good specimens the colour is pale grey all over except that the sides are
a little darker. When this grey pollination is rubbed off or otherwise spoiled, the ground
colour is black with the humeri and a patch between them and the wing bases distinctly —
orange. The dorsum is rather irregularly clothed with small bristles of type § arising
from scattered tiny irregularly distributed black dots. These small bristles are larger on
the prothorax and a pair just on edge behind head is very distinct. The ground colour
shows to a more or less extent through the orangish grey pollen, and one may often dis-
tinguish it to form indistinct thoracic lines, e.g. one from the above mentioned pair of small
bristles in front down to the level of the first d. c. pair. The macrochetes stand on dark
spots; these are: 2 pair d. c., the front just behind cross suture, h., n. p., presut., 3s. a., also
st. p.andmesop. All these are of type a. Scutellum as thorax with two large div. discal
bristles on blackened spots, a small black crossed terminal pair not on spots: sometimes
there are in addition one or two small scattered bristles of type 8. The outline of the
scutellum is a shallow rounded triangle. The metanotum is large, two ridged, grey on
top and blackened beneath.
Wings as in Fig. 13. They have the remarkable “bulle” characteristic of
this and the related genera, the position of these being shown in the figure. The
general scheme of colour is as shown in the figure, and is black. The venation
is not typical. In de Meijere’s description and figure the distances on the costa
between the end of the second vein and that of the 3rd is about equal to that between
the 3rd and 4th. In this species, the second vein meets the costa roughly midway
between 1 and 3, and is curved up; also, the 4th vein is not straight, but is remarkably
undulate as shown. The dotted line in Fig. 13 just below the third vein shows where
a fold occurs (the “sulcus cubitalis” of Girschner, see Kertész, Ann. Mus. Nat. Hung.,
1903, p. 358); when viewed from beneath this appears as a well chitinized extra vein.
Unlike de Meijere’s figure and description (where no discal cross vein is shown or
mentioned) that vein is here well developed. The coxal nick and its two bristles are
very well developed. The rest of the venation is readily seen from the figure. No bristles
occur on any veins. The halters are clear orange-yellow.
The legs are yellow except for two incomplete darkened rings at the ends of front
femora, the whole of mid and hind femora are darkened except at the tips, and there is
a dark ring near the base of the middle and hind tibize. Two upper rows of bristles on
front femora all weakly developed, black, and with 4 bristles each: a lower row of 3
stouter bristles, the first black, the next two white ; terminal spine to mid-tibize, no other
well-marked bristles.
Abdomen. ¢ darkish grey with silvery hairs; the hypopygium very projecting
from below base: last segment bordered with stout black bristles. ¢, last segment
forming a flattened sheath about as long as rest of abdomen.
Size g, over 25 mm. ; wing, about 3 mm.
Localities. Seychelles. Silhouette; from near Mont Pot-i-eau, ca. 1500 feet, VIII.
1908. Mahé: from near Morne Blanc, X.—XI. 1908; Cascade Estate, ca. 800 feet,
1908—9. This species was swept from grass in forest-clearings &c.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 323
Sepside.
Sepsis Fallen, Dipt. Suec. Ortal., 20 (1820).
21. Sepsis rufa Macq., Dipt. Ex., Suppl., iv. (1850), 296. 3; Becker, Mitt. Zool.
Mus. Berlin, u. (1903), p. 144.
There is a long range of a variable species which all agree very closely in the
cheetotaxy of head, thorax, abdomen and legs: they divide roughly into a large pale
form (typical rufa) and a smaller dark form, but a few intermediates occur. The sole
difference appears to be that the middle femora in the pale form have three stout bristles
underneath at middle, while the dark one has only one or two there, and all the bristles
are weaker in the smaller form. The front femora and tibiz agree with the description
of Becker (/.c.) and with the description given by de Meijere, Ann. Mus. Nat. Hung., iv.
(1906), p. 179, and with his figure, /.c. tab. 11, fig. 11. De Meijere in that paper draws
attention to the same fact, namely that S. rwfa seems very variable in body, leg, and head
colour, and also in size, which he finds varies from 24 to 4 mm.: in all his specimens he
also finds the cheetotaxy similar.
As far as can be seen without detailed dissection, the genitalia are the same in all
the male specimens. On the whole the females are darker than the males.
Localities. Seychelles: from a number of localities, both near sea-level and in the
high mountain-forests. Silhouette: near Mont Pot-i-eau, ca. 1500 feet, VIII. 1908; Mare
aux Cochons, IX. 1908. Mahé: Cascade Estate, ca. 1000 feet; from near Morne Blanc,
X.—XI. 1908; Anonyme Island, I. 1909.
22. Sepsis albopunctata, n. sp. (Plate 19, fig. 11).
The collection includes three specimens of a male with pale white spots at the tips of
the wings, and indistinct black spots by them, which do not agree with any of the
published descriptions.
Head. Bright shining yellow; on vertex, upper eye-margins, space round each ocellus,
a broad line below and above, brown orange. Antennz yellow with usual dorsal bristle
on 2nd joint, and orange arista. Cheetotaxy normal, viz. oc., post-v., inner and outer
v.; there is a group of bristles each side on back of head between neck and eye-margin,
and the usual clump just over neck. Three well-marked vibrissze followed by a row of
equal hairs on the mouth-margin behind them.
Thorax. Shining orange yellow, pleuree somewhat lighter, a silvery patch on upper
half of sternopleura. Two d. ¢., the line of these continued to the front by a line of
bristles, which are much more distinct than usual: acrostichals in two irregular rows
of the same character as last bristles; other bristles normal. There is a notable dark
orange line from humeri to wing base along the suture.
The wings have quite normal venation: 2nd vein slightly turned up at tip, 3rd and
4th parallel: a terminal white spot (involving the tip of 3rd vein) between 4 and 2, a palish
dark spot just over the tip of 2, bounded by the pale spot in front, and extending down-
wards to rds the distance between 2 and 3. The discal cross vein somewhat beyond the
SECOND SERIES—ZOOLOGY, VOL, XVI. 42
324 PERCY SLADEN TRUST EXPEDITION
middle of cell; the distance between cross veins on 4 slightly longer than abbreviated end of 5
beyond the last cross vein. Discal cross vein about half length of last one. Halters yellow,
brown stalks. Legs all yellow except the darkened terminal two or three tarsal joints.
Front pair (Plate 19, fig. 11):—femur with a hump on middle carrying two large and several
small thorns, proximally two stout spines and smaller one, a few hairs behind these ; bristly
on tip, one bristle near distal one-third is bigger: tibia bent and distorted, proximally three
bristles, then a small hump with four bristles on it. When viewed in profile, the ventral
side of tibia is seen to be excavated, principally near the hump, so that it can receive the
femoral hump to a greater or less extent; a minute preapical bristle: tarsi with first joint
strongly pectinate below, especially at base. Middle pair:—femur with one bristle on outer
side towards middle ; tibia with three bristles on hind edge (as stretched out) from middle
towards tip, preapical, two small terminal, one beneath about middle ; tarsi, first two joints
with double row of side bristles pointing backwards. Hinds legs; femur with one outside
bristle two-thirds from base ; tibia clubbed in distal half, a ring of three bristles at middle ;
tarsi as middle legs.
Abdomen with very humped and knobbed shining yellow second segment, the others
purplish shining black, with beginning of petiole and end cavity more orange: second
bristled all over, stronger at side: others with smaller sparser general bristles and large
bordermg macrochetes.
Size, about 3 mm. ; wing, about 2 mm.
Locality. Seychelles. Mahé: Cascade Estate, 800 feet or over, 1908—9; marshes
on coastal plain of Anse aux Pins and Anse Royale, I. 1909.
In addition to the above there are a few specimens of the female of a dark species
with white and black spots on the wing.
Localittes. Seychelles. Mahé: Cascade Estate, and marshes on coastal plains at
Anse aux Pins and Anse Royale, 1908—9.
Micropezide.
Neretus Fabricius, Syst. Antl., 264 (1805).
23. Nereius alluaudi Giglio-Tos, Ann. Soc. Ent. France, lxiv. (1895), p. 366.
There is a long series of a species of Nereiws which agrees well with the description of
this Seychelles species. Giglio-Tos says that one of his specimens had three longitudinal
lines of which the middle was much narrower. All the present specimens possess the three
lines, the two outer and broader ones of which extend from the calli to the scutellum: the
middle one is much narrower and only extends about half-way down the dorsum. The
breadth of the lines varies somewhat in the different specimens.
Localities. Seychelles. Silhouette: near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908;
Mare aux Cochons and forest above, over 1000 feet, IX. 1908. Mahé: Forét Noire
district, and high forest of Morne Blane and Pilot, X.—XI. 1908; Cascade Estate and
forest above, ca. 1000 feet and over, 1908—9; Mare aux Cochons district, 1000—2000
feet, L—II. 1909; 1905 (Gardiner) ; 1892 (Alluaud).
This species is very characteristic of the damp endemic mountain-forests, throughout
which it is found.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 325
Drosophilide.
DrosopHita Fallen, Dipt. Suec., Geomyz., 4 (1823).
Many species of this genus were collected, only two of which can be referred to known
species from published descriptions.
The genus is represented by a large variety of forms, some of which might well be
considered subgenera or even genera, but as these are represented by single species it is
thought best to merely indicate such possible divisions.
For convenience of reference to what follows the species may be grouped as follows :—
I. Species with costa to third vein (Leucophenga Mik.).
sericea, p. 326.
grossipalpis, p. 328. :
II. Species with remarkably constricted waist and short wings.
*notabilis, p. 329.
*III. Species with a remarkable slit to the costa, the end provided with spines and
bristles.
fracticosta, p. 329.
mgerrima, p. 331.
nigrobrunnea, p. 332.
IV. Species like Chymomyza :
*punctiscutellata, p. 333.
V. Species with remarkable transverse eyes :
*aberrans, p. 334.
VI. Species with remarkable spines on front tarsus :
*spinipes, p. 336.
VII. Species with curious curled hairs on front legs :
*hirtipes, p. 337.
VIII. Species with marmorated thorax.
punctulata Loew, p. 339.
jucunda, p. 339.
*TX. Species with a distinct shining vertical triangle.
Jimtima, p. 340.
caliginosa, p. 341.
pallipes, p. 342.
triangulifer, p. 343.
rufuloventer, p. 344.
X. Species of ordinary pale form recalling melanogaster.
nasuta, p. 346.
melanogaster Mg., p. 347.
similis, p. 347.
It is highly probable that the species and groups marked thus * should be raised to
generic or at least subgeneric rank, but as so few specimens are available, this course is
not adopted.
42—2
326 PERCY SLADEN TRUST EXPEDITION
24. Drosophila sericea, n. sp. (Fig. 14 and Plate 19, fig. 12).
A very handsome species in which the male has a very silvery thorax.
$ Head. Eyes bare, red. Top view :—Frons nearly parallel-sided from the vertex
downwards, a little over one-fifth of maximum axial breadth from eye to eye: creamy
white; the area within the ocelli is grey with silvery reflections: ocelli bright red. The
pair of lower f. 0. are inserted nearly midway on the frons and point forward: the first
pair of outer backwards-pointing f. o. are inserted nearly level with the former, the second
pair roughly midway between them and the vertical bristles. These f. 0. bristles are lower
down the eye-margins than in the European species L. maculata: approximated 1. v., diy.
o. v.; a small pair of deeply crossed p. v. b., stout oc. ; a row of minute crossed bristles on the
centre of the frons; a row of similar bristles below front f. 0., so that it can be looked on as
a continuous row of fine orbitals. Front view :—Face coloured as frons, slightly diverging
to insertion of vibrissze, practically flat in section, a strong pair of vibrisse. Side view
(Plate 19, fig. 12) :—Back of head darkish grey ; the post orb. continuous round mouth-
margin; a stout bristle on lower hind angle of jowl. Antenne with the just visible
first joint and the second joint coloured like the frons, slightly bristled, second with one
longer outer dorsal b.; third joint parallel-sided with rounded tip, finely pubescent,
slightly darkened ; arista pale for first half, then darker: six upper rays, three lower rays
(1st near middle), terminal ray: on the under face of the plane of the arista is a row
of tiny rays, denser for the proximal third and then extending more sparsely to nearly
the end*.
Palpi blackened and shining, tongue yellow, rugose at the tip: the mouth opening in
vertical view is long and narrow, and the margin is blackened laterally.
The thorax and scutellum are both brilliantly silvery in good specimens, the
ground colour beneath being orange; the dorsal surface is covered with small bristles.
There are two d.c., the front pair the smaller, small prescut., one h. and a few smaller
bristles on humeri, two n. p., three bristles over wing, the p. a. large. The small
thoracic bristles can be looked on as forming 8 or 10 acr. rows between the d. «.
lines. Scutellum slightly arched, almost circular in profile, quite bare, with two large
side bristles on large grey ill-defined spots, and two stout crossed terminals. Pleura
somewhat orange-yellow with a darkened irregular stripe from the metanotum to the
sternopleura. Two large st. p. Wings as Fig. 14, veins brown, the costa darker: the latter
extends to 38rd: a fine dense black short bristly ciliation extends to about half-way
between the ends of 2 and 3; it here becomes sparser, and between that termination
and the end of 3 are four or five minute black tubercles inserted just on the lower
edge of the costa. In macroscopic view the costal margin is distinctly infuscate.
Halters with white knobs, and yellowed stalks. Legs entirely pale yellow-white
except that the mid and hind knees are narrowly darkened. The usual preapicals
* This inner row of small rays is a useful character: it varies in strength of ray, and in number, and
is occasionally quite absent. It is plainly visible in Z. maculata. It will likewise be found in other insects:
for example, in Lonchea plumata (see last paper, Trans. Linn. Soc., Vol. xv, Part 3, 1912, p. 303) it is
almost as strong and evident as the upper and lower rows and is complete from end to end.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 327
on hind and mid tibie, front femora with a few longer fine bristles beneath and
outside.
Abdomen. The ground colour is yellow, the second segment is bristly only on
its upper angles and margins and is silvery; third pale yellow, rather silvery, with
long marginal and a few scattered discal bristles ; fourth with three large black marks,
the side ones extending right over the edges and from front to back, the middle
also extending from front to back, hence the orange-yellow ground only shows as two
rather irregular dark orange lines; the three black bars are nearly equal in width:
the fifth segment has also three marks, the middle extending right across, the side
ones sometimes only reaching over the side margins and not front and back: sixth
segment with a small dark central spot. These last three segments have long scattered
dorsal and longer marginal bristles.
@ The female differs in its colour scheme as follows. The thorax is more or
less silvery in side view on tip, but always quite visibly so on the sides of the thorax,
and on the upper part of the pleura. The dorsum is brownish orange, darkish, with
two dim darkenings forming side stripes along the d. c. lines, and darkened again
before scutellum. The latter is darkened except at the tip which is white.
Fig. 14. Drosophila sericea, n. sp. x 25.
The abdomen has its second segment creamy with large dark triangular spots
occupying the sides and most of the lower margins, but well separated distally on the
centre line. The next two segments have the orange ground much covered with black
somewhat shining colour forming two large side and one large middle mark, both
extending from margin to margin: these segments could also be collectively described
as being shining black with two orange stripes separating three broader black ones.
The fifth segment has three spots like the male, as has the sixth segment, the last has
two large side spots with a narrow line between.
A certain amount of variation occurs, the blackening being variable in extent in
both sexes.
Size, 25 mm. ; wing, 24 mm.
Localities. Seychelles. Silhouette: forest near Mont Pot-d-eau, ca. 1500 feet,
VIII. 1908 ; Mare aux Cochons, IX. 1908. Mahé: Cascade Estate, ca. 800 feet and
over, 1908—9; Morne Seychellois, over 1500 feet, II. 1909; high forest of Morne
328 PERCY SLADEN TRUST EXPEDITION
Blane and Pilot, 1908 ; Mare aux Cochons district, over 1000 feet, 1909. Throughout
the damp endemic mountain-forests.
25. Drosophila grossipalpis, n. sp. (Fig. 15 and Plate 19, fig. 13).
? Head (Plate 19, fig. 13) :—very much like last species in all essentials of
bristles and structure ; frons all bright orange-yellow, finely rugose longitudinally for
the proximal two-thirds: the ocelli red with a grey area inside. Face and orbits
the same as frons; back of head rather darker. The antennz are also yellow-orange.
Palpi orange, very stout, flattened in vertical plane. Tongue orange.
Thorax. AJl rather dull orange, cheetotaxy as last species. Pleura rather lighter
orange including the humeri: a brown ill-bounded spot below the sternopleural bristles.
Scutellum, etc., coloured as thorax, the same in shape as the last species.
Wings as Fig. 15, blackened on costa. Halters orange. Legs all yellow-orange.
Fig. 15. Drosophila grossipalpis, n. sp. x 20.
Abdomen. Ist segment orange, 2nd all black (except just at front margin in
centre) extending well over sides; 3rd all orange with darker spot on centre line
near distal margin and a much smaller brown spot near the proximal margin; 4th,
mid line yellow-orange with similar central spots as the 3rd, then entirely black
well over the sides, then orange and finally (on belly side) black; 5th orange with
two small oval black side spots; 6th orange. The whole is covered with black discal
bristles and long marginal bristles.
The central spots on 3rd and 4th segments are often almost absent or very
small.
There are three imperfect specimens of what may be the male of this species: it
agrees in all respects with the females described above, but the abdomen is entirely
black, the narrow margins being a little paler: the specimens are in no way silvery
or sericeous.
Size as last species.
Localities. Seychelles. Silhouette: near Mont Pot-d-eau, ca. 1500 feet; Mare
aux Cochons, highest forest, ca. 2000 feet. Mahé: near sea-level, Long Island, VII.
1908; also throughout the endemic mountain-forests (Cascade Estate ; Morne Seychellois,
over 1500 feet; Mare aux Cochons district, over 1000 feet). Félicité: from drier
type of forest near sea-level, XII. 1908. Praslin: XI. 1908, one of the three ¢.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 329
26. Drosophila notabilis, n. sp. (Fig. 16 and Plate 19, figs. 14, 15).
A remarkable form with constricted waist, small wings, and a very peculiar
frontal marking.
Head. yes finely and sparsely pilose. Top view (Plate 19, fig. 14) :—frons
very broad, shining, with broad eye-margins and a remarkably demarcated central
area as figured. This central area is translucent greyish bordered by brown-orange,
the broad eye-margins are similar in colour. The front f. 0. points forward, the hind
one outwards and backwards, i. v. convergent, o. v. divergent, p. v. absent. The oc.
stand nearly upright, parallel and bending forward at the tip. Front view :—the face
is shining orange with an elongate knob between antenne.
Side view (Plate 19, fig. 15):—the eyes are rounded in front; |
the mouth projects somewhat ; the antennee are orange, 3rd
joint paler: 2nd joimt swollen and nearly as big as the
3rd with a fine short bristle in front; the 3rd is pubescent,
Fig. 16. Drosophila notabilis,
especially so at tip, the arista with six upper, three lower a. x BO
and a terminal ray, the inner row is evident though sparse.
Fine single vibrissee, a few bristles on the hinder jowl; orange hind head, with fine
post-orb. row. The clypeus above the palpi is very large indeed; palpi darkened,
club-shaped, hairy ; tongue darkened and hairy.
Thorax darkened orange. Two d. ¢. with fine small rows extending in a line to
the front. The acr. bristles indistinct, two rowed. No sign of humeral bristles, one
behind the humeri which may be a single notopleural; two close together on the
side above and just before base of wing. Scutellum tawny-orange, crossed terminal
bristles, a very minute basal pair.
Wings as Fig. 16, translucent with brown veins. Halters with longish white
heavy and bituberculated darker stalks.
Legs. Yellow, bristles very few; one inner outstanding pre-apical on front
femora; pre-apical and 2 or 3 small crowning bristles on the mid tibia: the rest
are not clearly visible owing to the state of the specimen.
Abdomen with much constricted waist, the first 3 segments dull orange, the last
segments shining black, all sparsely bristled, the marginal bristles the longer.
Size, over 2 mm.; wing, under 14 mm.
Locality. Seychelles. Mahé: marshes on coastal plain, Anse aux Pins and Anse
Royale, I. 1909; Anonyme Island, I. 1909.
27. Drosophila fracticosta, n. sp. (Fig. 17 and Plate 19, fig. 16).
This (and the next two species) have a remarkable break at the end of the first
section of costa; this ends in a swollen projection of the costa which is somewhat
spear-shaped, black, and very bristly.
The present species must be near to D. gratiosa, de Meiyere (Tijdschr. v. Ent.,
54, 1911, p. 404) with which it agrees fairly closely in general facies and in head and
thoracic characters, but the abdominal characters are quite different.
330 PERCY SLADEN TRUST EXPEDITION
? Head. Eyes bare. Top view :—frons slightly narrowing in front, about half width
of head at back, dull black from the vertex forward, brown-orange from the forehead back-
wards, the colours merging in two indistinct boundary lines from ocellar area to outer edges
of lunular boundary. Eye-margins broad and shining black from front f. 0. to vertex and
back of head. Front f. 0. sub-parallel and forwardly bent, hind f. o. divergent and back-
wardly bent, inserted rather close together ; a row of fine bristles extends from latter to
forehead, the first, between the two f. 0., being the longest. Convergent i. v., divergent
0. v., p. v. crossed at tip, oc. inserted between basal and front ocelli, long and divergent.
Post-oc. rows evident. Front view :—antennz touching basally ; a well-marked snout
or keel extending from the base of the antennz to two-thirds down face, widening to
the end which has a rounded margin : face black, with sparse pale orange pollen, the
snout much more orange, clypeus prominent. Side view (Plate 19, fig. 16) :—antennze
darkened orange, 2nd joint with a large basal bristle and a smaller bristle in front ;
3rd joint silky, arista orange with darker rays, the top row with 5 progressively
smaller rays; bottom row with one ray between 3 and 4 on top; one end ray, a few
fine rays of the inner row. There is a stout vibrissa close to the eye-margin followed
by a row of smaller ones on the darkened jowl; hind jowls orange with 2 or 3 bristles;
palpi clubbed, flattened, black, with a few short bristles: tongue more orange.
Fig. 17. Drosophila fracticosta, n. sp. x 20.
Thorax. Dorsum, scutellum, and pleura above a line from. the wing base to
below the humeral calli and the metanotum, all intensely black; the dorsum very
shining, other parts duller, humeri slightly orange in tone, duller ; scutellum minutely
granulate: lower pleura and sternum entirely pale yellowish-white. Only the last ~
pair of d. c., two h., n. p., three bristles over wing base, one large and one small
st. p.; the dorsum all covered with fine bristles; the acrostichals form about eight
rows between d. c., but are entirely the same in character as the rest of the small
bristles. The scutellum is rounded in outline and swollen in profile; the terminal
bristles just cross at tip, small side bristles inserted somewhat nearer the terminal
bristles than to the base.
Wings, Fig. 17, yellow with yellow veins: the costal slit is very marked, and
ends in a black callosity provided with spines, and turned over so as to lie roughly
perpendicular to the plane of the wings. Costa to 4th vein. Halters with black knobs
and pale stalks. Legs entirely whitish-yellow: the front femoral rows weak, one
spine at end of middle tibia.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDAL, TRYPETIDA, SEPSIDA, ETC. 331
Abdomen. 2nd segment whitish-yellow with black margins wider at sides ;
3rd black with two faintly defined transverse elongate marks each side of the centre
line; 4th black with two large roughly semicircular pale spots with their diameters
on the base of the segment, distal margin narrow and pale; 5th similar but with
smaller elongate spots; 6th black with a central pale line which is widest proximally;
genital segments pale. In side view the whole of the abdomen appears black. The
belly is pale on the centre Ime. There is a small amount of variation in the exact
shape of the spots and amount of orange on the 38rd segment.
Size, about 24 mm.; wing, about 24 mm.
Localities. Seychelles. Mahé: ‘Cascade Estate, at or above 800 feet, 1908—9.
Praslin : Cotes d’Or Estate, XI. 1908.
28. Drosophila nigerrima, n. sp. (Fig. 18 and Plate 19, fig. 17).
This is close to the last in many structural points, but is remarkable for its
almost entirely black colour and its coriaceous texture.
Head. Eyes bare, colour entirely black, except for the brownish antennz and
lighter face. Top view :—frons dull black and finely granulate, except for the ocellar area,
a line from it to forehead, and the broad eye-margins, which are shining; the frons
narrows slightly from behind to the front ; sub-parallel front f o., slightly divergent
back f. 0., the usual fine row of bristles continuing the latter to forehead, the first
(between the f. 0.) being the longest of them; strongly convergent 1. v. nearly touching
at tips, divergent o. v.; p. v. directed backwards, nearly meeting at the tips, inserted
just below the vertex; divergent oc. on the ocellar triangle. Front view :—antennee
just separated at base by the beginning of the facial keel which widens downwards
in a uniform manner to about three-fifths of the depth of face. The face is all dark
except that the snout formed by the keel is paler than the rest especially on its
side surfaces. The black prelabrum is very evident. The palpi are included, so cannot
be described. Side view (Plate 19, fig. 17):—the 3rd antennal joimt is brownish and
pubescent; 2nd joint with a well marked spine on tip and a much smaller curved one
in front; arista with 4 top, 2 bottom, one terminal ray: some 3 or 4 sparse rays of
the inner row are visible. Convergent vibrissee with a following row of fine bristles ;
hind jowls with a few longer bristles. The post-orb. row is present, but the bristles are
not quite so evident as in most cases.
Thorax entirely black, slightly shining, dorsum finely punctulate and uniformly
covered with very short adpressed hairs: the last pair of d. ¢., two smallish h., n. p.,
small presut., p. a., one larger and one or two smaller bristles over the wing, no acr. ;
pleural bristling not visible owing to the position of the single specimen. Scutellum
more velvety black, rounded in outline and profile ; crossed terminal, and two smaller
side bristles, nearer the terminal than base.
Wings as Fig. 18, smoky with dark-orange veins; costal break well marked,
black, spiny, and turned up perpendicular to the wing-plane. Costa to 4th. Halters
dull brown-orange with suffused tips. Legs black except for somewhat orange knees
SECOND SERIES—ZOOLOGY, VOL. XVI. 43
332 PERCY SLADEN TRUST EXPEDITION
and tarsi, a moderately developed lower front femoral row of bristles, stout terminal spine
on the middle tibia.
Abdomen. Black, m dorsal view the following spots can be seen: on the second
segment, two indistinct silvery patches at the upper angles, which are rather bristly on
the sides; 4, 5, and 6 have each two pale silvery spots, transversely elongate, with the
front border coincident with the proximal margin, the curved lower boundaries extending
to about one-third the segmental breadth ; the spot on the 6th extends more over the side —
margins, and that segment is more greyish black than the others. AIl the segments
except Ist are bristled dorsally and marginally, the latter bristles being the longer. In
side view two inferior silvery spots can be seen on edges of 2nd just below the upper
angles and two inferior round silvery patches on 3rd (which is immaculate above), these are
nearly as big im diameter as the segment’s breadth. A further silvery patch can be
glimpsed on the belly of the 4th segment.
Fig. 18. Drosophila nigerrima, n. sp. x 30.
Size, 3 mm.; wing, 3 mm.
Locality. Seychelles. Mahé: a single specimen, Cascade Estate, at or above
800 feet, 1908—9.
29. Drosophila nigrobrunnea, n. sp. (Fig. 19 and Plate 19, fig. 18).
This is the 3rd species with a well-marked costal slit and swelling.
There are but three specimens, one headless, and one gummed which has conse-
quently lost its proper colouring.
Head. Eyes bare. Top view :—frons dull brown-black behind merging into dull
orange over the forehead ; ocellar area slightly shining; eye-margins very shining dark
brown (there is a slight silvery sheen on the frons in side view). The eyes project
much backward, so that the hind margin is very concave. Parallel forward pointing f. o.,
slightly divergent hind f. o.; the continuing row of bristles from the latter is absent,
except for the Ist small bristle between the two f. o. ; strongly convergent 1. v., smaller
straight and divergent o. v.; p. v. behind the vextex, crossed about two-thirds from
the tip; rather long divergent oc., inserted between back and front ocelli. Front
view :—face dullish grey-orange ; antennze with the large 2nd joints touching at the base;
a very well-marked keel between the 3rd joints, which widens gradually till it meets the
rather swollen upper lip, being demarcated therefrom merely by a sulcus: two deep
antennal foveee are thus formed. Clypeus evident; darker. Side view (Plate 19,
fig. 18) :—the head bristles seem rather more adpressed than usual ; 2nd joint of antenne
LAMB—DIPTERA: HETERONEURIDA), ORTALIDA, TRYPETIDA, SEPSIDA), ETC. 333
with a well-marked outwardly divergent tip bristle, and a smaller one outside towards
the end: it is orange in colour; 3rd joint suffused darker orange, pubescent ; arista with
five upper, two lower, one terminal, and two or three sparse inner rays. Vibrissa con-
vergent, followed by an extra fine hair row ; a few bristles on the hind jowl: post-orb. row
well behind the eyes. Palpi small, very slightly clubbed, orange. Tongue orange.
Thorax. Shining brown-black up to a line extending from below the humeri to the
abdominal insertion, paler on the humeri; the pleura and sternum yellowish white.
Dorsum covered with fine pale hairs. One d. c. pair, two h., n. p., small presut., three
over wing (one being the post-alar), two strong sternopl. Scutellum minutely pubescent,
rounded in outline, and swollen and rounded in profile; terminal bristles crossed at tip,
smaller side ones fairly near base. The metanotum is well developed in breadth and is
shining brown.
Wings as Fig. 19. Yellowish with brownish veins; a little more darkened just
below the costal swelling: the slit is well-marked, and
ends in a rather rounded upwardly bent black hairy
process. Costa to 4. Halters pale yellow.
Legs entirely yellowish white, lower and hind
rows of bristles on the front femora present, the
bristles weak: mid tibial spur strong.
Abdomen. Somewhat shining brown-black except: ee
that (1) the base is paler and (2) the fifth segment has
two large pale yellow spots which occupy the proximal margins of the segment except for
about one-third its breadth, and extend in a roughly triangular form over the segment
nearly to the distal margin, (3) the sixth segment has its sides yellow: it is more shining
than the others. The second, third and fourth segments are about equal in width, the
fifth and sixth are nearly twice as wide; genital segments orange-yellow.
19. Drosophila nigrobrunnea,
n. sp. x 30.
Size, just under 2 mm.; wing, just under 2 mm.
Localities. Seychelles. Mahé: Cascade Estate, 800 feet or over, Praslin: Cotes
d'Or Estate, XI. 1908.
30. Drosophila punctiscutata, n. sp. (Fig. 20 and Plate 20, fig. 19).
This is a single @ specimen of a narrow form which has a strong superficial
resemblance to Chymomyza. Oc. bristles on front of ocellar triangle.
Head. Eyes densely though shortly pubescent. Top view :—frons entirely shining
black ; with a side light the eye-margins and a long triangle nearly reaching the forehead
can be seen to be differentiated by being rather greyer in tone. The frons is about as broad
in front as its axial length, and widens in a curved line along the eyes to the vertex. It is
coneave behind. The ocelli stand out brilliantly as three silvery spots. The front f. o.
parallel and forward bent, hind f. 0. inserted close behind them, bent back and continued by
a row of equal fine bristles to the front. I. v. just off vertex, much cony. ; 0. v. just at
angle of vertex close to i. v., div.; p. v. just on curve of vertex, crossed well below tips ;
oc. long, div., inserted on frons in front of front ocellus and with bases further apart than
4133)
334 PERCY SLADEN TRUST EXPEDITION
hind pair. <A few tiny bristles on ocellar area. Post-orb. row well-developed especially
behind v. b. Front view :—the face is whitish brown, deeply hollowed, the antennse
contiguous basally, no central keel. Side view (Plate 20, fig. 19):—the 2nd antennal
joimt swollen and hairy, bearing a longer dorsal and a front spme; 3rd joint small and
pubescent, both joints dark brown; the arista dark with six upper, three or four lower,
one end ray and a sparsely scattered inner row. The black cheeks have a long vibrissa
at the angle followed by some finer hairs. The post-orb. bristles evident on the black
hind head, but rather remote from the orbits. Tongue orange. Palpi black, knobbed, with
bristles.
Thorax. Dorsum orange with indistinct lines a bit darker along d. c. line, covered
with sparse bristles. In front where it touches the head it is darker. There are two
pairs d. c., the first just about the level of the cross suture: h., n. p., twos. a., presut. :
pleural bristles destroyed by pin. Scutellum orange in mid line with two large black
spots on sides up from base: these bear basally two long bristles, and on their extreme
tips are the crossed terminal pair.
'
,
t :
\
|
!
1
Fig. 20. Drosophila punctiscutata, n. sp. x 50.
Wings as Fig. 20; glassy with orange veins; costa well-marked to the end of 8
with fine black ciliation, less marked to 4 with paler ciliation. Halters with rather
elongate orange heads and darker stalks. Legs entirely yellow, 2 or 3 longer ‘bristles
distally and behind on the front femora.
“Abdomen. Entirely shining black except for the orange ovipositor and the centre
line of the belly, bristled discally with longer marginal bristles.
Size, about 2 mm. ; wing, about 13 mm.
Locality. Seychelles. Mahé: Mare aux Cochons district, 1000—2000 feet. I.
1909.
31. Drosophila aberrans, n. sp. (Fig. 21 and Plate 20, figs. 20, 21).
This species is aberrant in general facies; the eyes are very oblique, and project
backwards behind the vertex: the oc. b. are inserted well outside the ocellar area.
¢ Head. Eyes strongly pubescent, red. Top view (Plate 20, fig. 20) :—frons
mainly orange, eye-margins slightly greyish yellow, a fine silvery border to the eyes can
be seen in some lights extending forward from the hind f. 0. The actual vertex, as well
as the back of head, are like the eye-margins. The area surrounding the ocelli down to
the oc. b. is blackened. The rest of the frons is (as stated above) orange, fairly shining,
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA), SEPSIDA, ETC. 335
and faintly ridged in a more or less longitudinal direction. Head bristles black and
strong; front f. 0. pointing forward and outward and more remote from eye boundary
than usual, the distance between their insertions being about half the frontal breadth at
the corresponding cross line; hind f. o. about half-way between the front f. 0. and the
i. v., pointing backwards; a continuing row of 6 or 7 small bristles to the forehead.
Convergent i. v., divergent o. v.; p. v. on vertex behind ocelli, crossed half-way ; oc.
inserted well beyond front ocellus (much in the position shown in Plate 20, fig. 22),
divergent and stout. In this view the hind angles of eyes can be seen to project behind
the head. Front view :—the head is very broad, about 24 times as broad as deep, owing
to the projecting eyes. The face to the ridged mouth margin is pale with a definite though
narrow central keel forming two antennal foveze: these are slightly darkened at the
bottom. The clypeal edge is blackened ; palpi clubbed, black, and bristled ; tongue yellow
and hairy. Side view (Plate 20, fig. 21):—antennze darkened grey, paler inside, the
lighter third joint with white pubescence; second joint with four or five stout spines and
smaller hairs. Arista stout, with the inner row of rays very marked and strong; upper
row with about four straight rays at base, then three curved ones; lower row with one
long straight ray below the upper, two or three curved ones, a terminal one. A pair of
very stout vibrisse followed along the mouth edges by finer bristles. The hind jowls with
three or four longer bristles. A well-marked post-oc. row. In this view the eyes are
- much elongate, the hind facets are perceptibly smaller than those in front.
Fig. 21. Drosophila aberrans, n. sp. x 30.
Thorax. Brown with orange tone, dull: some sign of sparse paler pollination in
front; two ill-defined narrow paler lines along the d. c. lines. The dorsum is covered
with small bristles arranged in regular longitudinal lines. Only the last pair of true
large d. c.: the fine bristles between the d. c. lines form 6-rowed acrostichals. The
humeri bear some 3 or 4 somewhat prominent bristles round the edge. ‘Two n. p., presut.,
two b. over wing-base, two on alar callus, the inner smaller. The pleurze below the line
from wing-base to point below humeral callus is more or less darkened: the sternopl. has
one stout lower and two smaller upper bristles. Scutellum rounded, bare, dull, con-
colorous with the thorax except for a slightly lighter tip and margins ; two stout basal,
and two stout crossed terminal bristles. Wings, as Fig. 21, suffused, with brownish
veins, the two cross veins being somewhat blackened; costa to 4th, no costal break, but
1 or 2 spines there. Halters orange. Legs yellowish-white, bristled in rows, the under
336 PERCY SLADEN TRUST EXPEDITION
row on the front femora uniform from base to tip; the last joint of the front tarsi with
rather unusually well-developed pads and claws, and a few long hairs.
Abdomen somewhat variable in colour; usually rather blackened with yellowish
margins at the sides only of segments 2 to 4, 5 with a broader continuous yellow
proximal margin. The extent of dark and light areas varies a little. There is a large
hypopygial knob, black and shining above; just beneath it is yellow and pointed, and has
an opposing yellow appendage springing from a previous ventral segment.
? The female differs as follows: the third antennal joint is generally much darker,
the oe. b. are not inserted as in the 2, but outside the line joining ocelli (Plate 20, fig. 20):
the thorax has a somewhat lighter ground colour, especially in front, so that in one
or two specimens we may distinguish a darkened middle line between the inner acrostichal
row and another pair of lighter lines followed by darker ones parallel to them. Some
specimens were immature. Front feet normal.
Size, about 1? mm.; wing, 2 mm.
Localities. Seychelles. Silhouette: Mare aux Cochons, IX. 1908. Mahé: near
Morne Blanc, X. 1908; Cascade Estate, 800 feet or above, 1908—9.
32. Drosophila spinipes, n. sp. (Fig. 22 and Plate 20, figs. 22, 23, 24).
This is a remarkable species represented only by 5 rather imperfect female speci-
mens; unlike nearly all the known species, these females have specialised characters on
the front tarsi. The oc. are in front of ocellar area.
Fig. 22. Drosophila spinipes, n. sp. x 35.
? Head. Eyes quite bare. Top view (Plate 20, fig. 22):—slightly concave behind;
behind the front f. 0. b., dull orange, the upper eye-margins more shining, beyond these
bristles merging to yellow: the ocellar area is not demarcated from the rest of frons, it is
clothed with bristly hairs, which are exceptionally long: many similar (but shorter)
small convergent bristles exist on the front part of head beyond front f. 0. The front f. o.
are parallel, there are two true hind f. o. b., shghtly divergent, with the usual continuing
row of fine bristles down to the forehead ; these are very well marked and regular. I. v.
nearly meeting at tips; o. v. much divergent; p. v. nearly meeting at the tips; the oc.
are quite parallel, and are inserted in front of the front ocellus. All the bristles are
brown. Front view:—face rather narrow, about twice as high as broad, fairly parallel-
sided; yellow: the large 2nd antennal joint approximated at base, no true central keel,
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETID A, SEPSIDA, ETC. 337
but a shallow depression for the antennz. Side view (Plate 20, fig. 23):—antenne ;
2nd joint large, orange, covered with pale bristles with one spine at top; small darkened
3rd joint, the arista with basal jomts very distinct ; three upper, one lower, one terminal
ray, the ner row is absent. Short stout vibrissa with usual following row of fine
bristles. Palpi orange, clubbed; tongue with remarkable file-like surface. The post-
orbital row is strongly developed just behind the v. b.
Thorax. Orange, entirely covered with pale brown bristles, last pair d.c. very close
to scutellum ; humeri with two large and a few other bristles, 2 n. p., presut., 3 over
wing-base, one small and one large sternopl. Scutellum orange, bare, flattened, shaped
like a triangle with much-rounded apex ; terminal bristles crossed about 4 from tip, a long
bristle on side of scutellum, slightly convergent to centre line. Wings, Fig. 22, slightly
yellowish ; costal slit present, but no thickening; costa to 4. Halters orange. Legs
all orange, femoral rows of bristles weak; front tarsi with remarkable spines beneath
(Plate 20, fig. 23), black claws and a few curious long hairs on the last jomts. The
other tarsi have no spines, but possess the black claws and the hairs. All the other
spines are very weak, though there is a moderate terminal mid-tibial spur.
Abdomen broad, browner on disc with scattered discal and marginal bristles. The
genital segments are large, forming in profile a blunt triangle whose length is roughly
equal to that of the rest of the abdomen ; this triangle’s axis is about perpendicular to the
body-axis. There is the usual hairy hump behind, and the tip of the triangle is formed
by the, ovipositor, which has a crown of stout short black spines.
Size, about 13 mm. ; wing, a little over 12 mm.
Localities. Seychelles. Mahé: near Morne Blanc, X. 1908. Silhouette: Mare aux
Cochons, IX. 1908. Praslin: Cotes d’Or Estate, XI. 1908.
33. Drosophila hirtipes, n. sp. (Fig. 23 and Plate 20, fig. 25).
There are two specimens of this species, both males: the dorsal sides of the front
tibia and tarsus are provided with remarkable long curved hairs.
é Head. LEyes minutely and closely pubescent, the hairs very short, and rather
fine. Top view :—eye-margins from vertex to front f. o., dullish brown-orange ; the
ocellar area the same, with a rounded front margin; between these the frons is more
shining orange: the ocellar area from in front to the vertex as well as the front of the
frons is clothed with distinct bristly hairs. Front f. 0. sub-parallel, hind diverging; the
continuing bristle row of the latter well marked down to forehead, its first bristle (just
behind and outside the lower f. 0.) much the largest. I. v. nearly touching at the
tip, o. v. divergent; behind these (just below the top eye-margins) the last 2 or 3
bristles of the strongly marked post-orbital border are especially prominent, and converge
to the centre. P. v. inserted on the vertical angles of the ocellar area, somewhat con-
vergent. Oc. long, much divergent, inserted just outside and behind the front ocellus.
Front view:—the large 2nd joints of the antenna are separated by a distinct space, in
the middle of which is the upper narrow part of the facial keel; this broadens uniformly
to the thickened upper lip with which it is almost confluent. The mouth-opening
338 PERCY SLADEN TRUST EXPEDITION
is unusually large: the face is all blackish with minute pollination, except for the
space between the antennal bases. Owing to the broad face, the antennal fovez are
rather shallow, the bottom being but little below the level of the eye-plane. Side
view (Plate 20, fig. 25):—the large 2nd antennal joint is orange, its inner face with long
hairs; there is an especially conspicuous top bristle, and two near the front margin, a
fourth lying between the top one and the others; the smallish dark 3rd joint is slightly
silky ; the arista has 6 straight upper, 2 lower and a terminal ray, the inner row has
some 5 or 6 rays. On each side of the face there is a clump of about 4 stout vibrissz,
followed by a strong row along the lower eye-margin : hind jowls, &c., with many bristles,
3 or 4 being especially long. Palpi dark, clubbed, strongly bristled at the end; tongue
rather paler, a few long bristles behind, and very bristly at the tip.
Thorax. Moderately shining black with minute sparse golden pollen: the surface
is covered with exceptionally stout and long discal bristles, which form acrostichals
which are irregularly 6-rowed between the two pairs of approximated d. c.; the last
discal bristles might be considered to form 4 true (though small) prescutellars. The
humeri are somewhat orange, and bear two bristles; n., presut., alar and p. a. as usual,
but very strong. Two very long sternopl.: the pleuree more orange. Scutellum as thorax,
but entirely bare: rather spade-shaped in profile, and flattened; two crossed terminal
and two long slightly convergent side bristles, inserted at about one-third the distance
from the base to the terminal pair. Wings, Fig. 23, slightly yellowish with brown veins:
eee es
Fig. 23. Drosophila hirtipes, n. sp. ~x 30.
costa more bristly than usual, extending to 4, but the bristles ending before 3: no true
costal slit. The alule have quite long ciliation. Halters orange. Legs all orange, the
front legs with an inferior femoral row of three very stout bristles, the back row of 3 or 4
is much less strong: strong spur and preapical to the middle tibia. All the legs with
neatly arranged rows of general bristles. The most remarkable character is in the front
legs. The tibia has two irregular rows of long hairs which begin a little before half-way ;
they number some 8 or 9: each is bent upwards in side view and inwards in front view.
The first tarsal joint is similarly provided with 8 to 10 hairs, and a few more occur at the
ends of the next 2 or 3 joints.
Abdomen. All brownish-black with numerous discal, and long marginal, bristles.
The whole insect is remarkably bristly for a Drosophila.
Size, about 3 mm.; wing, about 25 mm.
Localities. Seychelles. Mahé: Cascade Estate, 800 feet or above; marshes on
coastal plain, Anse aux Pins and Anse Royale, I. 1909.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETID A, SEPSIDA, ETC. 339
34. Drosophila punctulata Loew, Cent. i. 100, Berlin. Ent. Zeitschr., vi. (1862).
Becker, in his paper on the Diptera of Madeira, says that Wollaston’s D. repleta
(Ann. Mag. Nat. Hist., Ser. 3. 1. (1858), 117), is the same species. It seems fairly
cosmopolitan, being known from Madeira, Cuba, &c.
The specimens agree with the description, although there is some variability in
respect to the spotting of the thoracic dorsum. It may be noted that the costa is
distinctly slit, spined and swollen at the end of Ist section. Two pairs d. ¢. are present.
The inner row of aristal rays are present, though but 5 in number.
Localities. Seychelles. Mahé: Cascade Estate, 800 feet and over, 1908—9.
35. Drosophila jucunda, n. sp. (Fig. 24 and Plate 20, fig. 26).
An elegant species with beautifully marked thorax, &c.
Head. Eyes bare. Top view :—frons yellow, darkened and slightly curvilinearly
ridged between eye-margin and pale centre line. The ocellar area is also darkened. Hye-
margins rather shining. Front f. 0. nearly parallel, inserted on inner edge of eye-margins ;
hind f. 0. somewhat divergent: a row of fine bristles from there forward to frons. The
div. i. v. and conv. o. v. are situated on two black triangles on the vertex; these
are continuous with the black back of head; crossed p. v.; div. oc. inserted rather
close to front ocellus. Front view :—2nd joints of antennee touching at base ; a sharp,
rather narrow keel most prominent in centre. Clypeus black margined, rest of face dusky
Cas eS re er SA eS
Fig. 24. Drosophila jucunda, n. sp. x 40.
orange. Side view (Plate 20, fig. 26):—the black top of vertex is very prominent ;
antennee ; basal joints yellow, one evident spine on top; 3rd joint suffused and pubescent ;
arista pale to branching of lower ray except where rays leave it, 4 upper rays, the first
close to base, two lower and one terminal, no sign of any mner rays. Vibrissa stout
with even row of rather stout small bordering bristles. Palpi rounded stick shaped,
black, tongue orange. A few long bristles on hind jowl. Post-orb. b. strongest above.
Thorax. Dull black with small discal bristles in rows and well-defined orange
pollinated marks as follows: two longitudinal slightly interrupted stripes from front to
about level of front d.c. b.: the black band between carries 4 acr. rows. In front of the
cross suture is a short transverse line each side. An ill-defined sparse pollination on the
front of the thorax lies between these and the front of the central pale line. Just
outside the end of the longitudinal lines is a short oval line, outside which again are
SECOND SERIES—ZOOLOGY, VOL. XVI. 44,
340 PERCY SLADEN TRUST EXPEDITION
two small spots above wing base: a similar very small spot behind the last d. c. b.
Humeri orange, as is an irregular spot above the n. p. b. The pleure are dusky orange
in varying degrees of intensity. The bristles are two d. ¢., one h., n. p., presut. (small),
two over wing, p.a. Scutellum as thorax, with two pollinated basal side spots and a
terminal one. Terminal b. crossed at tip, side-basal ones somewhat convergent. The
wings, as Fig. 24, pellucid with orange veins, costa strong to third, weaker to fourth ;
no slit at end of 1st segment, but bristles there. Halters orange. Legs pale with
darkened front femora, and faint indistinct darkened rings proximally and distally
on other femora and proximally on the tibis, the middle one of which has a long
terminal spur.
Abdomen. Dorsal view: 2nd segment yellow with black distal margin, interrupted
in middle third; 3rd segment yellow with similar interrupted margin, but can best be
described as black with three yellow spots, two lateral ones extending to proximal margin,
mid one sub-triangular and extending across segment ; 4th black with three yellow spots
attached to proximal edge; 5th with three similar but much smaller spots; rest all black.
All the distal edges are narrowly yellow. In side view, 2, 3, 4 have distinct yellow
spots: genitalia yellow. All segments with fairly stout bristles, both discal and
marginal.
Size, 15 mm.; wing, about 1°6 mm.
Localities. Seychelles. Praslin: Cotes d’Or Estate, XI. 1908, 7 specimens. Mahé:
Mare aux Cochons district, 1000—2000 feet, I. 1909.
36. Drosophila finitima, n. sp. (Fig. 25 and Plate 20, fig. 27).
Head. Eyes closely and finely pubescent, vertex somewhat concave behind. Top
view :—frons matt orange; the eye-margins, and a rather small though definite triangle
surrounding and including the ocellar area, shining. Nearly parallel front f. o., slightly
div. hind f. 0.; the continuing row of bristles present, the first large and opposite
front f. 0. Conv. i. v.; div. 0. v.; p. v. crossed about one-third from tip, inserted below
vertex ; div. ocellar based between the large front ocellus and the side ones. Ocellar area
with small bristles; 3 or 4 small crossed bristles on front of frons and a few more over the
antennal bases. Front view :—face slightly greyish-yellow: large 2nd antennal joints
touching at base: a central keel starts just below them and runs down to level of third
joint, widening a little, and cut off from upper lip by a groove. Side view (Plate 20,
fiz. 27):—orange 2nd antennal joint, bristled, with two longer ones: 3rd joint a little
suffused ; arista long-rayed, five top, two bottom and a terminal ray, inner row continuous
but of fine bristles. One vibrissa, with stoutish bristle-row along edge of orange lower eye-
margin, orange hind jowls with bristles; post-orb. row especially strong behind v. b.
Palpi and tongue orange, former slightly clubbed and bristled at end.
Thorax. Brown-orange, somewhat shining, bristled dorsally to a little beyond level
of first d.c., then bare. Two pairs d. ¢. with little bristles forming six-rowed acrostichals
between them, and eight-rowed beyond: 2 smallish h., n. p., pr. sut., 2s. a, p. a; 3
nearly equal st. pl. Scutellum as thorax, bare, crossed terminal and equal side bristles
LAMB—DIPTERA: HETERONEURIDA, ORTALID A, TRYPETIDA, SEPSIDA, ETC. 341
inserted fairly near base. Wings, Fig. 25, slightly yellowish with brownish veins ; costa
as usual, dark ciliated to between 3 and 4, fine and rather long cilize to fourth. Halters
pale orange. Legs: all yellow-orange, a rather weak distal hind row of 3 or 4 front
femoral bristles: usual mid-tibial spur.
Abdomen. Brownish-black to black in mature specimens, sometimes paler at base ;
third segment shining black at base except in middle, fourth with basal half shining,
last almost all shining. In the female the bands are usually indistinctly dull orange :
there is a certain amount of variability in the abdominal colour: in one or two females
there is no sign of marks, and the abdomen is uniformly black.
Fig. 25. Drosophila finituma, n. sp. x 40.
Size, about 13 mm. ; wing, about 12 mm.
Localitws. Seychelles. Silhouette: Mare aux Cochons and forest above, IX. 1908 ;
highest forest, ca. 2000 feet, 1908. Mahé: high forest of Morne Blanc and Pilot,
X.—XI. 1908; slopes of Morne Seychellois, over 1500 feet, II. 1909; Cascade Estate,
800 feet or above.
37. Drosophila caliginosa, n. sp. (Fig. 26 and Plate 20, fig. 28).
This is a small dark species which should be near D. nigra (de Meijere, Tijdschr.
Ent., li. (1908), p. 153) but the legs differ from his description.
Head. Eyes with dense, short, stubby pubescence. Top view :—frons entirely
black, eye-margins shiny, extending beyond front f. 0. in a blunt point, a well-marked
shining frontal triangle extending two-thirds towards forehead, with very sharp bounding
lines; the rest of the frons (forming a Y-shaped area with broad stalk on forehead, and
arms narrowing to vertex) is dull black. Front f. 0. sub-parallel and bent forward ; hind
bent outwards and backwards, the row of small bristles from this to frons are present,
the first being quite large. I. and o. v. as usual; p. v. rather small, just crossed;
oc. almost parallel, inserted one outside each side of, and nearly in line with the front
ocellus : the ocellar area has a few minute bristles. Front view :—the lunular edge over
the antennze is somewhat orange: the antennz are separated at the base by the median
keel, which widens out along the line of the third antennal joint to form a well-marked
snout extending down to the somewhat swollen mouth-margin, but well separated there-
from. The face is all dark, but slightly pollinated. Side view (Plate 20, fig. 28) :—second
449
342 , PERCY SLADEN TRUST EXPEDITION
antennal joint rather swollen, with spines, two in front being long; small oval third joint,
pollinated, but hardly pubescent; both joints dark, but sometimes with slight orange
tone ; arista with five rays on top, two below and one terminal; inner row short, but quite
evident. Stout pair vibrissze, followed by mouth bristles which are moderately strong,
approximating to small extra vibrissee; hind jowl with a few bristles. Post-orb. row well
developed, the last one or two (behind outer v. b.) being quite stout and converging to
mid-plane of head. Palpi clubbed, bristly, black; tongue blackened with paler tip.
Thorax. Entirely black, moderately shiny, dorsum entirely covered with regular
rows of small dark bristles: two pairs d. c., inserted rather close together, the front pair
smaller ; the small discal bristles form 8 acrostichal rows just in front of the d.c., and 6
between them: small but evident prescut. pair; two h., usual n. p., presut. and wing
bristles ; three sternopl. and several much smaller ones. Scutellum bare, duller black,
very minutely granulate, almost dull brassy; terminal bristles just touching at tips,
small side bristles inserted midway along sides. Wings, Fig. 26, translucent with
yellow veins: costa to 4th, the fine black bristling ending a little before 3. There
is a distinct costal slit, but no callosity. Halters dusky-brown. Legs: except tarsi,
entirely brownish-black, tarsi yellowish-white, but the dark bristles render this less
conspicuous. Front femora with only three moderate bristles of lower row; no con-
spicuous hind row. Mid-tibize with moderate spine and few other small crowning bristles;
preapicals small.
Fig. 26. Drosophila caliginosa, n. sp. x 45.
Abdomen. Same colour as thorax, about two irregular cross rows of discal bristles,
and a marginal row of somewhat longer ones: these bristles appear black, but when
reflecting light are paler than the ground colour.
Size, a little over 15 mm. ; wing, a little under 15 mm.
Localities. Seychelles. Mahé: Cascade Estate, 800 feet or above; marshes on
coastal plain at Anse aux Pins and Anse Royale, I. 1909.
38. Drosophila pallipes, n sp. (Fig. 27 and Plate 20, fig. 29).
There is a single male specimen of this species.
Head. Eyes with profuse short pubescence. Top view :—frons with broad black
shining eye-margins and a well-marked large black shining frontal triangle, between the
eye-margin and the triangle is a long triangular stripe which narrows to practically zero
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 343
at the vertex; it is dark dull black-brown, merging in front with the rest of the frons,
which is orange. The ocelli form conspicuous silvery spots on the triangle. Parallel
front f. 0. ; subparallel hind f. 0. ; the row of bordering bristles starts with the usual large
one inserted just outside front f. o. and extends to forehead ; long subparallel i.v., diver-
gent o. v.; p. v. inserted below the vertex and crossed ; oc. inserted between basal and
front ocellus, somewhat divergent; the ocellar area has some 6 or 8 minute but stout
bristles on it ; there are two rows of minute pale bristles extending to the forehead along
the centre lines of the narrow triangles and over the frons. Front view :—antennee just
separated basally, a central keel which widens but very little extends nearly to the upper
lip, but does not meet it. The face is orange. Side view (Plate 20, fig. 29):—second
antennal joint pale yellow, with upper and outer spines and a few small bristles; 3rd
jot brown, silky; arista pale at base with long feathers, five upper, two lower and
a terminal: the inner row is represented by only one or two hairs. Stout vibrissa
followed by usual row of fairly strong bristles ; jowls orange and hind head black. Post-
orb. row well-marked, three or four behind the v. b. being very stout. Tongue and palpi
yellow, the latter rather small, slightly thickened at end.
Thorax. Shining black with rows of small dorsal bristles except behind; these
bristles are brown in most lights; two d..c.: twos. a. The other bristles cannot be seen
owing to the method of pinning. Pleurz black, two st. p. Scutellum, black, but polli-
nated with minute palish pollen: long crossed terminal, long slightly divergent side
bristles. Wings, Fig. 27, very slightly darkened with brown veins; costal ciliation black
and strong to before 3 then finer to 4. Halters with
pure white knobs and slightly yellow stalks. Legs all very
pale yellow including the coxze: all the bristles weak, and
but little sign of the femoral rows.
Abdomen blackish, except for the two basal segments:
the fourth segment has two very shiny black spaces along — Fig. 27. Drosophila pallipes,
the proximal margin towards the sides, the last segments HSDs a H:
are largely shiny.
Size, just under 15 mm. ; wing, about 1} mm.
Locality. Seychelles. Mahé: Mare aux Cochons district, 1000—2000 feet, I.
1909.
39. Drosophila triangulifer, n. sp. (Fig. 28 and Plate 20, fig. 30).
This species must be closely related to D. pallipes, but its legs are all darkened.
Head. Eyes densely pubescent with fine stout hairs. Top view :—the well-marked
frontal triangle extends about three-fourths down to forehead ; both it and the broad eye-
margins are shining black and meet on the vertex. The front of frons is orange, and is
prolonged backwards in two triangular stripes between vertical triangle and eye-margins.
These narrow uniformly to their ends, and are more brown orange, darkening as
they narrow. Front f. 0. nearly parallel as are the hind; the small bristles beyond the
344 PERCY SLADEN TRUST EXPEDITION
former extend down to forehead, the first one stout, just opposite the base of front f o.
Considerably convergent i. v., divergent o. v., inserted rather close together; p. v. just
below the vertex, crossed at one-third from tip, oc. divergent, inserted midway between
the end and side ocelli. A few fine bristles on front of frons and on ocellar area. Front
view :—antenne just not touching at base of large second joints; a well-marked facial
knob widening below, most prominent just about level of end of third joint ; much concave
mouth margin: face all orange. Side view (Plate 20, fig. 30) :—second antennal joint yellow
orange, bristly, with two spines ; third joint orange, sparsely silky, arista with long curved
rays, five on top, two below, one terminal: inner row well-marked, and somewhat longer
than usual. Vibrissa, and row of following bristles on orange eye-margins, which form
the jowls: hind jowls with two or three long bristles: the post.-orb. row longer below
and behind v. b. Clypeus very evident; palpi and tongue dark orange, the former bristled.
Thorax. Entirely shining black including the pleurze and scutellum: dorsum very
sparsely (almost imperceptibly) pollinated and covered with moderately long lines of
bristles ; scutellum much more densely pollinated. Two d. ¢., the dorsal bristles forming
six rows of acrostichals; two h., n. p., presut., s. a. bristles, large lower and two smaller
upper st. pl, scutellar terminal b. crossed just beyond half-way; the equal side bristles
near base, just divergent. Wings, Fig. 28, very slightly smoky with brown veins: usual costal
ciliation to between 3 and 4 and then on to 4.
~— 955) Halters with almost white heads, stalks dark-
Oe | ened. Legs mostly dusky white, front coxze
| black, femora darkened except at the tips (in
| mature specimens), front femora only moderately
spined behind.
Abdomen. Shining black, more shining
3 a than rest on proximal half of the third segment
Fig. 28. Drosophila triangulifer, n. sp. x40. (except in centre) and on most of the proximal
part of the next segment except in centre.
The dorsum with rather long irregularly arranged bristles, longer on the margins.
Size, a little over 14 mm. ; wing, about 14 mm.
Localities. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908.
Mahé: high forest of Morne Blanc and Pilot, X.—XI. 1908; slopes of Morne Seychellois,
over 1500 feet, II. 1909; Mare aux Cochons district, 1000—2000 feet, II. 1909; Cascade
Estate, at or above 800 feet, 1908—9.
40. Drosophila rufuloventer, n. sp. (Fig. 29 and Plate 20, fig. 31).
A dark species with black thorax and a dark russet-brown abdomen.
Head. Eyes densely but shortly pilose. Top view:—vertex and frons entirely
rather shining intense black, with the large frontal triangle and the eye-margins still more
shining ; slightly concave behind. Parallel front and hind f. o., the continuing row of
small bristles to frons evident, the first bristle of these being large and just outside the
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 345
front f. 0. Convergent 1. v., divergent o. v.: p. v. convergent, inserted just below vertex.
Oc. divergent, inserted nearer front ocellus than hind. Ocellar area and front of frons
with a few minute bristles. The head bristles are all proportionately rather stout for
their length. Front view :—face shining black with a little sparse pale pollen; large
second antennal joints separated at base by the beginning of the middle keel, which
broadens into a distinct elongated knob reaching a little below the level of the third
antennal joint. The forehead forms an arched recess in which the second joints of antenna
are partly hidden; mouth-margin much convex. Side view (Plate 20, fig. 31) :—second
antennal joint with two spines and a few bristles, rather ochreous; third small, also
ochreous, arista with five top, two bottom and one end ray: inner row of five or six small
and equal rays fairly evident. Vibrissa each side, with a row of fine bristles on margin of
the broad, somewhat shining ochreous lower eye-margins. Hind jowls black and bristled.
The post-orbital row strongest behind the verticals. Palpi dark ochreous and bristly as
is the tongue. The shining clypeus is very evident in profile.
|
Se re NN
Fig. 29. Drosophila rufuloventer, n. sp. x 40.
Thorax. Entirely rather shining black, including the pleuree and the somewhat
duller scutellum: the latter and the dorsum are covered with minute sparse pale pollen
very regularly and beautifully spaced. The dorsum is also covered with regular bristles,
pale in side illumination, which extend nearly to the back of the same. Two d. c., the
discal bristles forming six-rowed acrostichals : the middle row of these ends in two small
true presc. b. Two h., two n. p., presut., two sup. a., small post a. The scutellum is
flattened on its posterior half: terminal bristles crossed about middle, equal side bristles
subparallel. Wings, Fig. 29, with brownish veins. Halters darkened orange. Legs much
darkened orange; front femora rather blackened, front tibize darker than rest ; inferior
row of four distally placed front femoral bristles.
Abdomen. Darkish russet brown, paler towards the tip. About two lines of discal
bristles, and slightly longer marginal ones.
Size, about 14 mm. ; wing, about 14 mm.
Locality. Seychelles. Mahé: Caseade Estate, at or above 800 feet, IT. 1909.
346 PERCY SLADEN TRUST EXPEDITION
41. Drosophila nasuta, n. sp. (Fig. 30 and Plate 20, fig. 32).
A somewhat large pale orange form with well-marked facial knob.
Head. [yes densely covered with short bristly hairs: hind eye-angles projecting a
little behind plane of head. Top view :—frons light-orange, not shining (see below “front
view ”), eye-margins very slightly greyer in tone. No triangle; the areas between eye-
margins and ocellar area ending off in rounded margins on the vertex. A very narrow
and shallow groove runs from the front ocellus to the forehead. Ocelli red. Front f. o.
a little diverging, hind the same; the small bristle row from latter begins with a stout
bristle nearly between the two f. o. (instead of as usual nearly opposite the front), the
continuing small bristles only number four or five. Slightly convergent 1. v., divergent
0. v., p. v. crossed at tips inserted just on curve of vertex. Oc. long and divergent,
inserted outside the line joining the ocelli and just behind the front one. A few small
bristles on the ocellar area and behind it; two tiny crossed pairs by the central furrow
just over the forehead. Front view :—the large second joints just separate at base: just
below them starts the snout which widens gradually till it joms the swollen upper lip ;
two deep fovese formed for antenne by the frons and snout. Face all orange: in this
Fig. 30. Drosophila nasuta, n. sp. x 30.
view (with a side light) the frons of the male is seen to be quite silvery. Side view
(Plate 20, fig. 32) —second antennal joint orange with fine bristles, especially side ; one
upper and one front spine. Third joint somewhat darkened distally and inside ; arista
with long straight rays, seven on top, one terminal, three below; the inner row exceptionally
well-developed, of some eight to ten rays. A stout vibrissa, with following row of
diminishing bristles on lower eye-margin; three or four bristles on hind jowl; post-
orbital row well seen, strongest and bent upward behind v. b. Palpi rather clubbed at
end, orange and bristly as is the tongue. Be.
Thorax. Dorsum non-shining orange, very uniformly covered with small bristles,
forming eight acrostichal rows before the d. c. lines and six between the two pairs of d. c.
Humeri much paler, as is a stripe from them to the wing base; this is succeeded by a dark
orange stripe from prothorax to.halters: the rest is pale yellow orange. Two h., n. p.,
presut., three over wing and another (post a.) just above scutellar base; large and two
small st. pl. The scutellum is flattened with well-marked margin ; terminal b. crossed
about two-thirds down, smaller side ones inserted on the edge and slightly convergent.
Wings, Fig. 30, somewhat yellowish with brownish veins, Halters orange. Legs all
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETID®, SEPSIDA, ETC. 347
orange, covered with very regular rows of fine bristles. Three distal bristles in front
inferior femoral row : large spur to middle tibia.
Abdomen. All orange, about 2% irregular lines of discal and a line of slightly longer
marginal bristles.
The female has little or no silvery sheen on the head; she has a short stout pointed
ovipositor, the hump is rather conical and hairy : the abdominal segments are sometimes
a little darkened distally.
Size, 24 mm. ; wing, just under 25 mm.
Localities. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908;
near Mont Pot-d-eau, ca. 1500 feet, VIII. 1908. Mahé: Cascade Estate, at or above
800 feet ; Mare aux Cochons district, 1000—2000 feet, II. 1909. Félicité: XII. 1908.
42. Drosophila melanogaster Meig., System Beschr., vi. (1830), 85. 9.
This insect under the above name and the synonym D. ampelophila Loew (see
Austen, Ent. Mag., 1905, p. 276) has been recorded from most parts of the world.
There are many specimens in the collection that can be referred without doubt to the
males of this species.
Localities. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet. Mahé:
Cascade Estate. Félicité: XII. 1908.
43. Drosophila similis, n. sp. (Fig. 31 and Plate 20, figs. 33, 34).
g This is a small species which is exceedingly like D. melanogaster. Its front legs,
however, are devoid of the extraordinary combs that are found in the latter species.
Fig. 31. Drosophila similis, n. sp. x 40.
Instead of these concentrated combs, under a high power (about 60) one can see on the
under side of the first two joints of the tarsus a row of smaller combs each of about half
a dozen teeth (Plate 20, fig. 33): the first joint has 4 such tiny combs, the next has
about 3. All the joints are covered with stout bristly short hairs as well. In all the
characters of colour, cheetotaxy, &c., the two species appear to be almost absolutely in-
distinguishable. The hind f. 0. is rather differently inserted, being slightly nearer the
front one and more remote from the i. v. than in D. melanogaster, and the auxiliary bristle
between the f. 0. is a little more in line. There is also some difference in the front and
SECOND SERIES—ZOOLOGY, VOL. XVI. 45
348 PERCY SLADEN TRUST EXPEDITION
side views (Plate 20, fig. 34). It is, however, impossible to be absolutely certain of the
constancy of the various minute differences that are apparent on prolonged examination,
nor can they be exactly expressed in a manner suitable for description.
The venation of the two species is practically identical (Fig. 31).
Size and wings as last.
Localities. Seychelles. Silhouette : Mare aux Cochons, over 1000 feet, IX. 1908.
Mahé: Cascade Estate, at or above 800 feet, 1908—9.
?¢ Corresponding to the above are many specimens of females, but it is practically
impossible to sort them out, as apparently none were caught paired with males.
Localities. These ?s were collected in Silhouette, Mahé (Cascade Estate and Mare
aux Cochons district), and Félicité Islands.
44. Drosophila, sp.
? There are 5 specimens of a female of the melanogaster group which can be
separated from the last two species by the second vein meeting the costa further down.
In all other respects they are practically identical. In the absence of a male, the species
is left undescribed.
Localities. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet. Félicité.
ZAPRIONUS Coquillett, Proc. U. 8S. Nat. Mus., xxiv. (1902), 31.
45. Zaprionus vittiger Coq., l.c. 32.
Four specimens of this African species were collected.
Localities. Seychelles. Silhouette: near Mont Pot-a-eau, ca. 1500 feet, VIII.
1908 ; Mare aux Cochons, IX. 1908. Mahé: Cascade Estate, 800 feet or over, 1908—9.
Cuymomyza Czerny, Zeitschr. fiir Syst. Hym. u. Dipt., i. (1908), p. 199.
A single specimen of a new species of this genus was present.
46. Chymomyza bicolor, n. sp. (Fig. 32 and Plate 20, fig. 35).
Head. Eyes, very faintly and sparsely pubescent under high power. Top view:—
frons shining, yellow, slightly lined; a very shallow narrow central depression from the
ocelli to the large dull orange lunular arc: eye-margins broad and shining yellowish
grey, extending forward to near the forehead. There are present the front f. o. parallel
and bent backwards; the middle f. o. inserted fairly near the former, nearly meeting in
centre of head and slightly bent forward ; hind f. 0. bent outwards and backwards, the
tiny bristle between front and mid o. b. is present ; conv. i. v., div. 0. v., both long ;
p. v. very minute, inserted well below vertex ; div. oc. inserted between ocelli, but a little
inside line joining their centres. A few tiny bristles on orange lunular patch and
on edge of a little triangular piece between the antennal bases, also a small divergent
pair on ocellar area behind ocelli. Front view:—face shining yellowish, very
shghtly concave; large 2nd antennal joints quite touching at base, and covered with
small bristles. Side view (Plate 20, fig. 35):—the 2nd joint is yellow with 3 or 4
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDAM, ETC. 349
‘exceptionally large bristles ; 3rd joint small, darkened, somewhat silky ; arista darkened
except for basal one-fourth; 3 or 4 upper, 3 lower, terminal, and a few fine inner rays.
The yellow face is very slightly concave; large vibrissa, the following bristles just on
mouth margin, they are 4 in number and very stout, nearly as big as the vibrisse ; hind
jowls yellow with two spines: a long one on back of mouth. Post-orb. row evident, two
long ones behind v. b.
Thorax. Shining orange, a slightly darkened spot just behind humeri, entirely
covered with fine bristles which form some 8 irregular acr. rows. Two d.c., the front the
shorter, h., n. p., presut., 3 over wings. Scutelluim as thorax but bare, tip rather obtusely
rounded ; two sub-basal bristles, two long terminal, slightly converging towards tips.
Wings, Fig. 32, tip white, costa darkened up to this tip; all the veins pale. Halters
whitish yellow. Legs: front coxee pale, femora and tibiz black, -except at base, Ist
tarsal joint black, the others yellow. Mid and hind legs all yellow.
Fig. 32. Chymomyza bicolor, n. sp. x 50.
Abdomen. Entirely shining black, covered with short dark bristles, which show up
light in oblique illumination.
Size, over 2 mm. ; wings, about 13 mm.
Locality. Seychelles. Mahé: Cascade Estate, about 1000 feet, I. 1909.
é
DETTOPSOMYIA, n. g.
There are 5 somewhat badly preserved specimens of a very remarkable Drosophilid
of quite unusual facies, which must be referred to a new genus (Plate 20, fig. 36).
Head. Top view (Plate 20, fig. 37) :—head wider across eyes than long, the eyes are
covered. with short bristly hairs, they project backwards. F rons broader than wide,
roughly rectangular in form with a well-marked sutural fold over the antenne. Eye-
margins very wide. Ocelli nearly in the middle of the frons. The cheetotaxy is abnormal :
in front over antennze are two f. o., the smaller forward inner one being slightly bent
forward, the neighbouring backward outer one being longer, and slightly bent back. On
the swollen hind eye-margins are two more bristles, which can best be looked on as the
vertical pairs: the front bristle (corresponding to inner vertical) is nearly upright, the hind
(or outer vertical) bends outwards. The p. v. pair are strongly crossed and are low down
on back of head; the oc. are divergent and are situated almost on the ocellar area, but
the bases are outside the line joining the front to the side ocellus. Front view (Plate 20,
45—2
390 PERCY SLADEN TRUST EXPEDITION
fig. 38) :—the face is almost entirely occupied by a large broad knob or snout extending
from the frons to the mouth-margin: this knob separates the antennee widely at the
base. Side view (Plate 20, fig. 39):—sub-quadrate in form with a projecting frons on
the edge of which the antennz are inserted. Jowls deep, the irregularly oval eyes
projecting backwards over the head. Mouth-margin projecting, so that the face is
concave in profile. Antennze with spine and hairs on second joimt, third jomt oval with
a strongly pectinate arista bearing upper and lower rays, and a few short rays of the
inner row. Vibrissa on side of snout with a row of long bristles above mouth-margin.
Palpi flattened, paddle-shaped.
Thorax. Fairly normal in shape: as far as the rather damaged state of the speci-
mens permits, there can be made out the following bristles:—a hind pair of d. ¢., and
another forward pair just behind the cross-sutural line: just across the suture are two
more small bristles on the same d.c. line: acrostichals in two rather sparse rows. Two h.,
presut. just above them., n. p.; alar bristles not to be seen clearly, two long st. jo Wine
scutellum has long terminal bristles crossed about two-thirds down, and long side bristles:
the scutellum is evidently swollen. The wings are peculiar, Fig. 33: there is a very well-
Fig. 33. Dettopsomyia formosa, n. sp. x 50.
marked Milichid-like nick in the costa with two spines (such as in D. fracticosta, &c.),
the end is pointed, the costa to fourth, all the veins much undulated, the second bends
sharply up to the costa half-way between first and third.
The abdomen and legs show no specially noteworthy characters. The whole
insect (except the abdomen) is diversely and beautifully variegated.
Type. The following species.
47. Dettopsomyia formosa, n. sp. (Fig. 33 and Plate 20, figs. 36, 37, 38, 39).
Head. Vertex; broad eye-margins shiny greyish-white, swollen behind ; bordering
the eye-margin inside is a black line joining the black vertex behind, broader than the
margin in front, narrower behind, in front each of these lines is divided into a short fork
by two silvery-orange spots over the lunule ; the central area of frons orange-silvery with
an elongate oval ocellar area which is black : it may also be described by saying that the
frons is all black between the eye-margins except for an elongated horseshoe-shaped area
from the antennze to the vertex, which 1s orange-silvery, as well as the above two orange
spots. The facial knob is dark brown on top half, with a silver-orange dorsal spot,
LAMB—DIPTERA: HETERONEURIDAL, ORTALIDA, TRYPETID A, SEPSIDA, ETC. 351
lower part and mouth-margin silvery-orange; the jowls like upper eye-margins, but
in side view more shiny brown on mouth-margin. Hind head dark, except where the
swollen upper eye-margins extend downwards. Antenne dark brown, except the outer
margin of second joint, which is paler; arista pale with four or five upper, three lower,
and one terminal stout hair. Palpi dark blackish brown; tongue yellow. Cheetotaxy
and eyes as in generic description.
Thorax. The dorsum is elegantly variegated in light grey and darkish-brown.
A brown-black central stripe from the front to back, which carries the acr. row; on
each side is a broad grey stripe which is divided longitudinally by an interrupted
brown stripe forming a brown elongate spot on the grey just before suture, and an
irregular brown mid-line thence to scutellum: this line is just joined to the middle
brown line by a little bar about midway between the two pairs of d. c.’s; from humerus
to wing bases is another brown line, attenuated in front. Humeri grey, a very fine
line of grey along the suture to the wing base; below this the pleure are brown, followed
by a grey line parallel to the brown one and starting from the front cox; the sterno-
pleura brown. Scutellum, brownish-black, two indistinct white basal spots, and a dis-
tinct terminal one. Wings with venation and mottling as shown in Fig. 33. Halters
darkish orange-grey, paler stalks. Legs: cox black-brown, femora same except for
white tips; tibia with 5 sub-equal rings, beginning basally these are white, brown, white,
brown, white; tibia orange. Front femora with two or three long bristles below, and
outside.
Abdomen. All dark shining brownish black.
Size, just under 14 mm.; wing, about 14mm.
Localities. Seychelles. Mahé: Cascade Estate, 800 feet and over, 1908—9; near
sea-level (Anonyme Island), I. 1909.
ECHIDNOCEPHALUS, n. g.
There are 5 specimens of a species which is allied by its venation to Liomyza. In
general macroscopic appearance it is much like the new Geomyzid genus Amygdalops
(p. 357), and it is remarkable that such a close superficial similarity in facies exists
(Plate 20, fig. 40).
Head. Eyes entirely bare. Top view (Plate 21, fig. 41):—head more than twice
as broad as long, excessively excavate behind, so that plan is like a crescent with
rounded horns: the frons widens gradually from the antennze to just beyond last f. o., it
then widens suddenly to the vertex, which is about twice as wide (along the margin) as
the breadth of frons. There are three f. 0., a small one just on the edge of the frons, a
stouter one behind, and a still stouter about the level of the front ocellus; all the f. 0. are
slightly bent backwards and very slightly outwards: i. v. sub-parallel, o. v. slightly
diverging ; beyond these, and a little below the vertex is a pair of much smaller divergent
bristles; there is a row of small post-orbital bristles. Tiny p. v. down behind vertex.
Long sub-parallel oc. inserted outside the line joining ocelli, but equidistant from them.
Front view (Plate 21, fig. 42) :—the face is very small, broader than high; the antennze
392 PERCY SLADEN TRUST EXPEDITION
nearly reach the eye-margins, the overall breadth from eye to eye is nearly five times the
depth of the face; if the mouth is viewed somewhat from below, the opening is seen to
be excessively wide. Side view (Plate 21, fig. 43):—a long flattened oval, eyes much
projecting backwards, and hiding the face in front; the jowls are not seen, being
apparently bent in sideways; hind head visible below. The antennz have the second
joint with three longish spines and a few small bristles; third joint irregular oval with
elegant recurved hairs at the tip, arista bare and inserted rather close to second joint.
On the lower angle of the facial mouth-margin is a long stout vibrissa, converging and
pointing downwards; it is followed by four downward-pointing bristles just below the
eyes; the hind head is bristly. Thorax: dorsum smooth and bare; cheetotaxy: two
evident d.c. behind the suture, continued in front by about three much finer bristles,
diverging to the humeral callus. The acr. bristles are represented by two or three
longish bristles in a single line from the front of the thorax up to the cross suture: just
in front are a few fine bristles on the front margin of the thorax. The hind notopleural
is large, and above it is an evident pre-alar, between the line joining these and the humeral
knob are many much smaller bristles, but it is scarcely possible to specify which is the
second notopleural, &c. ; the humeri and below are covered with fine bristles. Scutellum
with short divergent end pair, and behind these and little further apart is a much smaller
sublateral pair; two small sternopleurals, and one above, are all that are very evident.
The metanotum is much developed: when the insect’s thorax is viewed perpendicularly
to its horizontal plane, the metanotum’s outline projects beyond the scutellar boundary.
Venation as Fig. 34, rather like Ziomyza. No special leg characters.
Type. The following species.
48. Hchidnocephalus barbatus, n. sp. (Fig. 84 and Plate 20, fig. 40, Plate 21,
figs. 41, 42, 43).
Head. Structure as described and figured for the genus. Frons and vertex pale
shining orange, darkened inside the ocelli, and somewhat darkened in a roughly U-shaped
mark above the antennz: the mouth-margin
(viewed from below) is darkened, especially
on the front lip.
Thorax. Orange on dorsum and
on pleura down to a rather narrow line
starting just below the humeri and extend-
ing below the wing insertion to the
abdomen ; this line is darker brown-orange:
below this the thorax is all yellow. Wings
slightly smoky, more intensely so between the costa and the long second vein: all the
veins brown. Halters with long brownish knobs and lighter stalks. Legs all yellow:
front femora with row of four or five long bristles on top and about five behind on
the distal half; small terminal spine on the mid-tibia ; distinct preapical spine on the
hind femur.
Fig
s:
34. Echidnocephalus barbatus, n. sp. x 30.
LAMB—DIPTERA: HETERONEURIDA, ORTALID®, TRYPETIDA, SEPSIDA, ETC. 353
Abdomen. Dark shining orange-brown, sparsely bristled; the narrow margins and
the belly, paler.
Size, 25 mm. ; wing, about 24 mm.
Localities. Seychelles. Félicité, XII. 1908. Mahé: Cascade Estate, ca. 800 feet ;
marshy ground near sea-level at Cascade, II. 1909.
AstEIA Meigen, System Beschr., vi. (1830), 88. 209.
49. Asteia mgra, n. sp.
A single specimen of an exceedingly dark form, quite distinct from any other ; it is
all shiny black except the abdomen and parts of the legs: the wings are rather shorter
in proportion than usual.
Head. Entirely shiny black; face rather translucent, a little silvery beneath
antenne ; antenne black, slightly rufous centrally inside and outside, about four upper and
four lower pectinations to black arista; one pair f. 0., inner and outer v., ocellar, stout
long vibrissee. Palpi black, tongue rather orange.
Thorax. Entirely bare shiny black including all the pleura, extremely indistinctly
and distantly pollinated, Ist and 2nd d.c., other bristles rather damaged. Wings slightly
yellowed with yellow veins; costal fringe very fine, distinct, especially on anal angle.
Halters black. Legs: front, coxa and femur black, tibia black at base, tarsi yellow.
Middle cox pale, femur black except at ends, tibia black at tip, tarsi yellow ; hind coxee
pale, femur black-ringed on distal third with pale knees, tibia with a narrow. black
ring near base, tarsi pale.
Abdomen. Entirely orange.
Size, 1 mm.; wing, 14 mm.
Locality. Seychelles. Mahé: from near Morne Blanc, X.—XI. 1908.
Geomyzide.
CurtroMYIA Robineau-Desvoidy, Myodaires, 621 (1830).
There is a rather unsatisfactory set of some 17 specimens that can be referred to this
genus. They all (with one exception) possess the crossed frontal bristles, and that
specimen agrees absolutely in other characters with several others. The facies of the
species is unlike that of the European ones: the acrostichal rows are indistinguishable
from the general thoracic dorsal bristles which are all small and dark instead of long and
pale. The large hind d. c. are present, but the forward row of smaller ones is absent. All
the macrochetes are shorter and darker. In view of the known diversity of the cheetotaxy
in this genus (see the specific descriptions in Becker's paper on Peletophila, Zeit. fiir Syst.
Hym. und Dipt., iv. pp. 131—133) it is not thought desirable to make a new genus for
the present species.
50. Chiromyra dubia, n. sp. (Fig. 35 and Plate 21, fig. 44).
There are 11 specimens of two sexes which appear to be best treated as one species,
although they differ in abdominal colour. There are no perceptible differences in
structure.
? Entirely orange and yellow.
354 PERCY SLADEN TRUST EXPEDITION
Head. Top view :—yellow, dull except for a small triangular projection in front,
and a narrow edge surrounding the red ocellar area, and the extreme edges of the eye-
margins; these are rather shining. The basal ocelli nearer than distance from front to basal.
Conv. front f. 0. pair, two backwardly pointing hind f. o. pairs (in one specimen the former
are missing: in another there are 3 hind f. o. on one side). Between the 3 f. 0. pairs
are a few very minute bristles. The surface of the frons has also a few small scattered
bristles. I. v. conyv., almost in line with the f o., o. v. div. ‘Tiny crossed p. v. below
vertex. Much div. oc. inserted between ocelli. There are about 4 small evident bristles
behind the v. b.: these are possibly the last bristles of the post-orb. row, but the true
p. 0. row 1s somewhat separated from this group and its bristles are much smaller than the
above 4 and are directed sideways and not upward. Front view :—the yellow antennz are
closely touching and half le in a deep pit in the white face. The orange palpi are best
seen in this view: the eyes are nearest just below base of antennze. Side view (Plate 21,
fig. 44):—the eyes are somewhat oval (though nothing like C. minima, Beck.), the long
axis being about 30° to the horizontal head axis. Jowls broad (see fig.) all covered with
palish hairs. Crossed vibrissal pair followed by another nearly as stout and a few similar
ones behind them. The black flagellum of the pale yellow antennz is very conspicuous.
Thorax. The dorsum is orange and is covered with small black bristles, the
humeri pale yellow as is the area behind them and all the pleura. The last d. c. pair is
large and fairly close to back of thorax, it is followed by a distinct row of bristles, the first
2 or 8 being longer than the others and diminishing to the size of the ordinary discal
bristles, which form 6 scattered and irregular rows of acr. b. Humeri with a large hind
and small front b.; usual 2 n. p., a presut., the alar bristles are not very distinguishable
from the dorsal bristling. There is a distinct mesopl. and a sternopl. bristle. The yellow
scutellum is bare with a pair of approximated terminal bristles and two smaller side ones:
the top is somewhat flattened. Wings,
i Fig. 35, normal, glassy with pale yellow
veins. Halters with conspicuous pale
yellow knob. The legs are all yellow with
i flattened femora and black claws.
Abdomen. Orange, with small dark
bristles. The last segment is knobbed
with longer bristles and two small dark
Fig. 35. Chiromyia dubia, n. sp. x 40. ;
side spots.
¢ Two perfect and two headless specimens are possibly the males of the above. In
any case they cannot be separated without much more material.
The head characters are almost the same as above, but the bristles are rather
stronger. The thorax is also the same, but again the bristles are stronger ; the bristle in
front of the d. c. is stronger, and there is a very strong prescut. pair, which is quite absent
from the above.
The abdomen is dark brown with narrow pale margins.
Size, over 1? mm. ; wing, over 1? mm.
Locality. Seychelles. Mahé: Cascade Estate, 800 feet or over, 1908—9.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSID4, ETC. 355
51. Chiromyia sexspinosa, n. sp. (Fig. 36 and Plate 21, fig. 45).
Four males of a very distinct form.
Head. Top view :—colour and cheetotaxy as last species except as follows : ocelli
in an equilateral triangle, post-orb. row much longer. Front view :—the eyes are nearest
just above antennal bases. Side view (Plate 21, fig. 45):—jowls a little deeper and
hairy, but the broad true eye-margins are bare. Antennze rather more orange. Palpi &c.
as last.
Thorax. Very much as last but all the bristles much stouter, the p. a. especially so;
small prescut., only one h. bristle that is of any
size. Colour the same.
Scutellum bare, in no way flattened; sub-
parallel terminal, two side, and two small discal
bristles behind the terminal. Wings, Fig. 36, much
as last, as are the halters and legs.
Abdomen. Dark orange, basal segment with
Fig. 36. Chiromyia seaspinosa,
extensively blackened upper corners, the others with
a distinct round spot each side and an indistinct ees
darkening in centre line.
Size, 2 mm.; wing, about 13 mm.
Locality. Seychelles. Mahé: Cascade Estate, 800 feet or over, 1908—9.
52. Chiromyia hirtiscutellata, n. sp. (Fig. 37 and Plate 21, fig. 46).
Two males: this species has bristles nearly as pale as the European forms.
Head. Top view :—pale yellow with very slight shininess. The basal ocelli nearer
together than distance from basal to front. The cheetotaxy is quite normal, though slightly
longer than the other species. The small upward pointing bristles behind the v. b. are very
regular and well developed; the post-orb. group behind the eye is composed of much longer
bristles than usual and overlaps the true p. 0. row: p. v. crossed half-way. Front view :—
the eyes are nearest below the bases of the touching antenne: face white. Side view
(Plate 21, fig. 46):—the eyes are rather rounded, antennee yellow with browner flagellum,
and a yellow hair on 2nd joint above. The jowls are very broad behind, covered with
yellow bristles except on eye-margins ; 3 distinct vibrissal pairs meeting or just crossing.
Thorax. Coloured as usual; the d. c. pair preceded by a smaller auxiliary pair:
dorsal bristling profuse but irregular. A
very large and strong h. b.; n. p., and p. a.
are also very long and strong, a smaller
presut. and s. a. Single mesop. and st. pl.
Scutellum slightly flattened, long touching
terminal, shorter side bristles, and some 6
or 8 small discal bristles. Wings, Fig. 37,
quite normal. Halters with bright yellow
knobs. Legs normal. Fig. 37. Chiromyia hirtiscutellata, n. sp. x 40.
SECOND SERIES—ZOOLOGY, VOL. XVI. 46
356 PERCY SLADEN TRUST EXPEDITION
Abdomen. Ist segment orange: others with distal two-thirds darker or even
brown-orange, the margins yellow, bristled all over, bristles longer on genital segment.
Size, about 12 mm. ; wing, about 12 mm.
Localities. Seychelles: Mahé, Cascade Estate, at or above 800 feet. Amirantes:
Eagle Island, X. 1905.
Apuaniosoma Becker, Mitt. Zool. Mus. Berlin, uu. (1903), Dipt. Aig., p. 186.
See also Czerny and Strobl, Verh. Zool.-Bot. Ges. Wien, lix. (1909), p. 283.
There are a few specimens of a small fly which appear more nearly placeable in this
genus than elsewhere, though not agreeing entirely with the generic characters as given
by Becker or as interpreted by Czerny. In both the species A. approximatum Beck. and
A. quadrinotatum Beck. as noted by Czerny (l.c.) the front f. 0. b. are absent, and not
present and crossed as in Chiromyia. The present insect agrees in that respect. But in
the above two species (as also noted by Czerny) there are but two true o. b. placed rather
far down the frons and a few more forward bristles much smaller in size. The present
insect has a third upper o. b. between the normal two of the genus, and the succeeding
row of 3 or 4 bristles not very small, though they are distinctly smaller and paler.
Again, in both the above species the single d. c. are succeeded by a row of fair sized
bristles with two well-marked moderate acr. rows between. The present species has
the dorsum covered with rows of quite small bristles which give about 6 acr. rows.
The facies, head structure and venation &c. agree well with the ordinary species; it is
hence thought undesirable to erect a new genus on the above points.
53. Aphaniosoma sexvittata, n. sp. (Fig. 38 and Plate 21, fig. 47).
Head. Eyes bare, head hollowed behind: frons yellow, narrowing in front to about
half the width just at back of frons: round each ocellus is a small reddish ring : the central
area of the frons is very slightly depressed and is a little rufous in tone. The three f. o.
mentioned above are followed by 3 or 4 progressively smaller bristles more nearly on the
side of the eye. Div. o. v., conv. 1. v.; much conv. p. v. inserted just below vertex
opposite each of the hind ocelli; oc. div., rather upright, inserted between ocelli: a pair of
small bristles on ocellar area between hind pair; some 5 pairs of small bristles on frontal
area, and a few overhanging forehead. Front view:
pale 2nd antennal joints touching,
face small, a deepish central pit: all pale yellow. Side view (Plate 21, fig. 47) :—2nd
joint hidden in pit; 3rd rounded finely silky, yellow; arista darker on flagellum, quite
bare. Convergent vibrisse, the large jowls with many stout bristles on side and below.
Post-orb. row stout on upper hind orbits, especially behind v. b. Tongue and palpi yellow.
Thorax. Orange with six appreciably darkened stripes, two from post-alar calli to
humeri, practically vanishing about the cross suture; two starting from scutellum
extending about three-fifths over thorax, one between each of above stripes, starting at
back of thorax, and ending about the cross suture. The narrow line between mid-pair is
pale yellow. The prothorax (where head touches) is darkened brown. The disc is
covered with rows of small bristles. Pleurze pale yellow. Scutellum bare, rounded and
arched, the disc rather darker, Last pair d. c. followed by smaller bristles in a row, the
LAMB—DIPTERA: HETERONEURID@, ORTALIDA, TRYPETID®, SEPSIDA, ETC. 357
pair next to the d.c.’s are little bigger than others which are like the other general
bristles of the dorsum. The fine dorsal bristles form six approximately regular acy.
rows; h., two n. p., presut., rather large p. a., small prescut., two basal, two terminal ;
scutellars just crossed at tip. Metapleura dark grey brown. Wings as Fig. 38: veins
pale brown. Halters with almost white head and orange
stalks. Legs pale yellow, a row of small spines on top of front
femora.
Abdomen. Second to fourth segments grey with narrow
pale margins; next with two dark spots confluent proximally
on fifth segment, discrete and smaller on the next. Fig. 38. Aphaniosoma
seavittata, n. sp. x 30.
Size, about 1-4 mm.; wing, about 14 mm.
Locality. Seychelles. Mahé: Cascade Estate, at or above 800 feet, 1908—9.
AMYGDALOPS, n. ¢.
See Plate 21, fig. 48.
Head. Eyes sparsely pubescent under high power. Top view (Plate 21, fig. 50) :—
frons about as wide as eye-breadth : somewhat convergent in front. Hind head concave,
but little excavate. Front and back pair of f. 0. parallel to axis, bent a little backwards:
front pair near antennze with small ancillary bristle in front: hind mid-way between hind
end eye-margin and the front one. I. v. conv. (only the insertions shown in fig.), 0. y. div.,
inserted right on edge of vertex and rather close together. P. v. crossed at tip, inserted
below vertical ridge and each opposite a hind ocellus. Long fine oc. inserted just outside
middle of line joining front to back ocellus. Front view (Plate 21, fig. 49):—face narrowest
about level of end of third antennal joint, no keel, but two fine furrows demarcating the
front eye-margins. The antennze touch at the base. Side view (Plate 21, fig. 51, the axis
of head is much inclined to horizontal in the fig.):—head much longer than deep, the eyes
much pointed at the projecting hind upper angle. Forehead very visible in front. Antenne
erect at base, second joint with an upwardly-directed bristle above at end; third sub-
triangular, with the rounded tip very elegantly haired. Arista very like a Drosophilid :
upper set of rays, longer and closer at base, lower set and an evident inner row (see p. 326).
The jowls deepen from front to back ; there is a pair of strong vibrissee each side and hairs
along margin of jowl. The post-orb. row is confined to the upper hind angle, and is small.
The palpi are rather long, slightly clubbed and flattened in vertical plane.
Thorax. Normal in form; scutellum somewhat arched, roughly triangular in plan,
with rounded tip. Last pair d. c., four rowed
aer. conv. in front, h., n. p., two small b. over
wing, div. somewhat upwardly-arched long
terminal scutellars and two small sub-discal
bristles. Two rather small st. pl. Wings, Fig.
39: there is a fine regular black costal bristling Se Sane i
between ends of 1 and 2, bey ond it is a pale Fig. 39. Amygdalops thomasseti, n. sp. x 30.
eiliation. The end of 5 is remote from the
margin. Legs normal, a few long bristles in a row below front and hind femora.
46—2
358 PERCY SLADEN TRUST EXPEDITION
Note. This form has many Drosophilid-like characters.
Type. The following species.
54. Amygdalops thomasseti, n. sp. (Fig. 89 and Plate 21, figs. 48, 49, 50, 51).
The cheetotaxy is as given in the generic description ; the present species has a fine
row of bristles extending from the d. c., the first (in front of d. ¢.) being somewhat longer
than the rest: the acrostichals form four well-defined rows in front, less well defined
behind, the mid-rows diverge at base of thorax. Other small bristles (like the acrosti-
chals) are on the general surface of the dorsum. The following details refer to colour
characters principally.
Head. Vertex and frons shining orange, with a long darkened triangle from the
vertex to about the mid-frons. The base of this vertical triangle is about half the distance
between eyes in middle of frons: forwardly it is bounded by fine darker lines which
do not meet. Hye-margins very broad, orange, silvery especially towards front. Hind
head hollowed and darkened orange. Face orange with narrow silvery eye-margins
broadening out sideways into the broadish somewhat silvery-orange jowls. Antennze
orange, arista a little darker: upper hairs are, dorsally five or six very long ones on basal
half, three or four shorter ones beyond; below some four fine hairs, and a very evident
inner row; palpi and tongue yellow with a few hairs.
Thorax. Dullish dark brown, two narrow orange stripes from the neck to the cross
suture. A pale side stripe from right over humeri along line of bristles and base of wing
to the entirely yellow halters. Below this a sharply defined parallel brown stripe from
the neck to the abdominal insertion. All the rest of the thorax below the pale yellow is
orange. The rounded arched seutellum dark brown. Wings, as Fig. 39, suffused from
just before whole length of third vein to just beyond fourth and fifth veins: this
darkening is very strong (almost forming a terminal spot) from just before where the second
vein ends to the tip of wing: rest of wing pale, veins brown on the dark part, yellow on
the uncoloured part. Legs entirely palish yellow. Front femora with two roughly
complete rows of fine long bristles, one row inside, one out, both pointing downwards and
outwards. Mid-tibia with terminal spine: the fine ciliation below front and hind first
tarsal joints form two fairly conspicuous clumps just at beginning of these joints.
Abdomen. Dark brown on top and sides, yellow below. Sub-marginal bristles
small: a few others scattered on the dorsum. The male hypopygium is in the form of a
small hemispherical knob below the last segment.
Size, 2°1 mm.; wing, under 2 mm.
Localities. Seychelles. Silhouette; Mare aux Cochons, IX. 1908. Mahé: near
Morne Blane, X.—XI. 1908; marshes on coastal plain, Anse aux Pins and Anse Royale,
I. 1909. Praslin: Cétes d’Or Estate, XI. 1908.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 359
Milichide.
Mrticuta Meigen, System. Beschr., vi. (1830), 131.
55. Milichia superba, n. sp. (Fig. 40 and Plate 21, fig. 52).
This species is one with entirely silvery abdomen.
¢ Head. Entirely black. Top view :—frons entirely covered with dark somewhat
olivaceous pollination: lunular arch shining black. The area surrounding the brown
ocelli is not quite so pollinated, but is a little shining, as are two narrow triangular
eye-margins down to front f. 0. Two f. 0. very slightly divergent from centre line: front
forward, hind backward bent; below front one a row of smaller orbital bristles
alternately smaller and larger. Parallel 1. v., div. o. v.; sub-parallel p. v.; oc. stout,
inserted between ocelli; on ocellar area and just behind it are two small div. pairs; row
of small bristles along orbital stripe from inside v. to top f. 0. Scattered frontal bristles
two pairs of which are rather striking, a forward crossed pair over frons, a backward
crossed pair about middle; between these is a third pair and others. Front view :—
antenne separated at base by a projection of the black lunule: two smooth black long-
oval fover with narrow ridge between; mouth emarginate, the centrally projecting part
of the lip is dark orange pollinated. Side view (Plate 21, fig. 52) :—second joint of
antenne rather elongate with front bristles, pollinated with grey ; third joint black, the
arista brown with just perceptible pubescence under high power (x 45), the basal joints
abruptly thicker than flagellum ; three stout vibrissee with stout following row and similar
bristles on hind jowl. Palpi black, stout and somewhat flattened laterally, spiny.
Tongue black and spiny, recurved tip just perceptibly orange toned. The eyes occupy
nearly the whole view, the margins below mid-line being wider than above.
Thorax. Black, somewhat suffused with ochreous pollen. Dorsum covered with
bristles which are somewhat variable in length, so that it is impossible to discriminate
exactly between any possible front dorso-centrals and the other bristles: the hindmost
pair of these is however very large, and just behind them is a smaller pair of prescutellars ;
h., n. p. and presut., three stout st. pl., but the supra-alars have been destroyed by the side
pinning. Scutellum as thorax with two stout basal and two stout crossed terminal
bristles. Metanotum rather more pollinated. Wings
as Fig. 40. The costal slit well marked with two
short terminal spines: the costa is black bristled along
margin to just beyond the second vein; thickened to
fourth but only ciliated. Halters all shining black.
Legs all black, femora all with two inferiorly-pointing
rows of bristles, those on the hind legs the longest.
Fig. 40. Milichia superba, n. sp. x 20.
Abdomen. All covered densely with silver ex-
cept first segment: the second segment nearly as long as the next two, these and the
last sub-equal. The margins edged with minute black bristles, the terminal segment with
long bristles. The silver extends well over the sides on to the belly, but less so on the
base of the second segment.
LIBRARY)
} 2
= 2
360 PERCY SLADEN TRUST EXPEDITION
¢ The frons is somewhat wider and has a few more, but rather smaller, scattered
bristles ; the lunular area has dark orange pubescence.
Abdomen. Concolorous with thorax, covered with fine bristles except on basal
part of second segment, and about the basal third of the next two which are smooth.
The legs are altogether stouter than in the male, the bristles less conspicuous, and the
first tarsal joint of hind legs is swollen.
Note. It is just possible that this is the ¢ of another species, though the two
occurred in the same locality.
Size, 34 mm.; wing, 22 mm.
2 ai
Locality. Seychelles. Mahé: Cascade Estate, at or about 800 feet, 1908—9.
Minicute.ia Giglio-Tos, Ann. Soc. Ent. Fr., lxiv. (1895), p. 367, Dipt. des Seychelles.
56. Milichiella lacterpennis Loew, Berlin. Ent. Zeitschr., ix. (1865), Cent. vi. 185, 97.
Several specimens of this species occurred: it is apparently very widely spread over
the world.
Localities. Seychelles. Mahé: from near Morne Blanc, X.—XI. 1908; Cascade
Estate, at or above 800 feet, 1908—9.
57. Milichiella tristis, n. sp. (Fig. 41 and Plate 21, fig. 53).
This species has no silvery sheen in the male, but is in both sexes entirely slightly
shining brown-black, none of the abdominal segments being in any way shining or even
brassy.
Head, black. Top view :—dull black, the ocellar area and upper eye-margins rather
shining, narrow lower ones pollinated. On these lower ones are two inwardly bent f. o.
with 5 or 6 smaller bristles; two upper f. o., the first outwardly bent, the other back-
ward, a small bristle between, another behind. Conv. i. v., div. o. v. on the shining
triangular ends of upper eye-margins. Two nearly parallel backward bent p. v. inserted
just behind hind ocelli. Stout oc. inserted between the ocelli: tiny bristles on ocellar
area, and a small div. pair just on vertex between ocelli. From front of ocellar triangle
to near the frons are two rows of about 4 pairs of converging bristles, and there are
scattered bristles just over the forehead. Front view :—antennz separated at base by
projection of shining lunule. Two long oval black foveze with moderate separating ridge.
Mouth edge arched, upper lip but little projecting and orange in tone. Side view
(Plate 21, fig. 53) :—in this aspect the frons is seen to be greyish in colour. Antenne
black with one stout spine and smaller bristles on 2nd joint: 3rd orbicular with black
arista suddenly thinning to flagellum which is slightly though evidently pubescent.
Vibrissee 4 or 5 each side, the first much the longest; a following row of small mouth
marginal bristles. Palpi stout, black and bristly. Tongue orange at the tip. The eye
nick is subtriangular with apex nearer top than bottom.
In the female the shining parts of frons are more evident, and the small bristles
rather smaller than in the male.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDAL, SEPSIDA, ETC. 361
Thorax. Black, slightly shining, very finely covered with minute lighter pollination,
dise covered uniformly with small bristles. The last pair of d. c. present and almost in
the same cross line with them are two small prescutellars. The specimens are not in
very good condition, but one can make out p., n. p., presut., and two on the post-alar
calli. The scutellum is coloured like thorax, bare with two basal and two slightly
converging terminal bristles. Metanotum, shining, somewhat pollinated in one specimen.
Wings, Fig 41, glassy with quite pale veins: costa with usual two-spined break,
minutely bristled to just beyond the junction with
the second vein which is bent along costa in the
usual way ; it is then slightly thickened to the end
of 4. Halters with orange knobs and darkened
stalks. Legs black except for the orange tarsi, the
last 4 tarsal joints are suffused in part, giving a
spotted appearance to the feet: there are no ‘ A, oe rh
s : : : Fig. 41. MMilichiella tristis, n. sp. x 30.
specially outstanding bristles on the legs, the bristle
clothing being as usual more or less in lines, except for a terminal spine on the mid-
tibie.
Abdomen. Somewhat shining black brown. In oblique bright light, one might say
it was the colour of dull oxidized copper. It is dorsally covered with bristles except for the
basal half of the 2nd segment ; this is about the same length as the 5th, each is about as
long as the 3rd and 4th together. Terminal segment with a few long bordering bristles.
There is no sign of any silvery or truly shining parts at all.
The female is much like the male; the relative sizes of 2nd to 5th segment of
abdomen is about the same in one of the two @, but in the other the mid segments are
relatively longer; this is probably due to the extension of the body with eggs; in both a
fleshy annulate ovipositor is visible.
Size, about 14 mm.; wing, about 1°7 mm.
Locality. Seychelles, Mahé: Cascade Estate, at or above 800 feet, 1908—9.
Another species of the genus was recorded from the Seychelles by Giglio Tos (Ann.
Soe. ent. France, lxiv., 1895, p. 367) as M. argentea Fabr.: this has been identified as
being distinct from that species and is now called M. tos: Becker (Ann. Mus. Nat. Hung.,
v. (1907), p. 536).
Eccopromma Becker, Ann. Mus. Nat. Hung., v. (1907), p. 540.
This genus was founded by Becker for certain Chilian species which differed from
Milichiella mainly in the very profuse and stout bristles on the thorax. Two males from
Mahé can be referred to a new species of this genus.
58. EHccoptomma solitarium, n. sp. (Fig. 42 and Plate 21, figs. 54, 59).
¢ Head, black. Top view:—frons narrowing much from vertex to forehead,
being about 4 times as broad behind as in front. The eye-margins and minute ocellar
triangle are shining, the rest is most elegantly and minutely rugulose, two sets of the
minute pits forming lines from outside the i. v. to the lunule; there is a minute furrow
362 PERCY SLADEN TRUST EXPEDITION
from the tip of the ocellar triangle. The head bristles are long. There are two pairs
lower f. o., the first just above the lunular level the other behind, the first pair is crossed
half-way, the other at tip: between them and down to forehead are several small bristles.
Right down on tips of lunular arch (at side of base of Ist antennal joint) is another pair of
orbital bristles bent backwards. The two pairs of upper f. o. are roughly parallel to axis
of head, the first forwardly, the second backwardly bent, inserted rather close with a fine
bristle between. The mid area of frons (between the pitted lines mentioned above) has
some few longish thin bristle-pairs, two pairs about one-third way down are bent back
and are more conspicuous. I. v. nearly parallel, o. v. div., between is a third extra some-
what smaller bristle; true p. v. well on hind head, slightly div., between and a little
above is asomewhat smaller div. pair; oc. inserted between ocelli on the small ocellar area,
rather upright, slightly div.; between hind ocelli is another smaller div. pair. Front
view :—forehead below lunule dark chestnut, face black with greyish pollen, narrowest
midway from forehead to mouth edge at point where vibrissal row starts ; it is furrowed ;
antennze touching at base. Side view (Plate 21, fig. 54) :—antennee black, 2nd joint with
3 bristles, top one longest ; 3rd joint with orange silky pubescence ; arista brownish, lower
joints thickest, flagellum diminishing slowly in diameter from them, finely pubescent. Main
pair of vibrissee crossed, the 7 or so following ones stout, straight. Mouth strongly bristled
below. Small regular post-orb. row ; well-marked nick at hind head as figured.
Thorax. Black with sparse ochreous pollen. The dorsum is very bristly, some of
the general bristles being nearly as stout as the macrochetes. One can readily distinguish
three dorso-central pairs behind the suture which are very long: there is also a fourth
bristle almost in the line of these before the suture: this bristle, with the humeral and
side presuturals form an evident cross line of larger bristles from one humerus to the
other: this row is partaken in by certain of the general bristles which lie in its direction
and are larger than the rest. This is a distinct, though smaller, prescutellar pair, which
form another evident cross row with the last pair of dorso-centrals, an extra bristle each
side between the prescutellar and the dorso-centrals, and the bristles over the wing bases. _
Pleurze bare between humerus and cross suture, with a stout bristle just behind and below
the humeral bristle; three stout sternopleural bristles. Scutellum rounded in profile
and section, with two basal and two
> _| terminal bristles, apparently crossed (only
SS one is present). Wings, as Fig. 42 : glassy
with brown veins; costal slit present as
ya usual with two short stout spines: costa
black bristled and bordered to just beyond
2nd vein, the thickening going to the 4th
vein. Halters black. Legs black, the tarsi
—— slightly paler in parts, especially the front
Fig. 42. Hecoptomma solitariwm, n. sp. x 30, Ist tarsal joint. The front femora with
abundant stout hairs on hind side
for about the distal half. The other femora with downwardly pointing long bristly
hairs.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 363
Abdomen (Plate 21, fig. 55). Silvery except for the Ist segment and the central
area and middle of the base of the second segment. The sides are much bent under and
the silvery shine is carried over the edges on the lower half of second segment and all the
others: the first segment is swollen at the upper corners and is there covered with black
hairs on its outer edge: it is pointed in the centre as usual and is all very shiny black.
The second segment bears a tuft of black hairs at each of its upper corners: its dorsum
has small bristly black hairs, some of which form two moderately regular lines parallel to
the posterior edge: the 3rd and 4th segments have two similar rows: the last segment
has the usual long bordering bristles. The relative sizes will be seen from Fig. 55.
Size, 31 mm. ; wing, 22 mm.
Locality. Seychelles. Mahé: Cascade Estate, at or above 800 feet, 1908—9.
DesmomeEtopa Loéw, Berlin. Ent. Zeitschr., ix. (1865), 185, vi. Cent.
59. Desmometopa M-mgrum Zett., Dipt. Scand., vil. 2743, 10.
Five specimens are in no way distinguishable from forms of this species.
Localities. Seychelles. Mahé: near Morne Blanc; Cascade Estate, at or above
800 feet ; marshes on coastal plain, Anse aux Pins and Anse Royale.
60. Desmometopa inauratum, n. sp. (Fig. 43 and Plate 21, fig. 56).
This species is very near to D. M-ngrum in most of its characters, including the
scheme of frontal marks, but the thorax is covered (in good specimens) with a very dense
golden pollination, and the jowls are less deep.
Head. Top view:—the frons is deep dullish black down to the antennze except
that there is a large equilateral sharp-pointed ocellar triangle, the upper eye-margins
joined to this triangle, and two narrow parallel stripes on the frons (carrying the usual
two rows of mid-frontal bristles) which are all bright grey: a very distinct black M
is thus formed on the frons. Lower two pairs f. o. slightly conv., with fine bristles
between, and a rather larger small bristle on extreme front of eye-stripes. Two pairs much
overhanging hind o. b., small bristles between them, and a rather large upright small
bristle between last and the i. v. I. v. bent a little back and a little conv., o. v. bent
much out; p. v. on vertex just outside and behind hind ocelli, slightly conv. ; two small
bristles between 0. v. and p. v. and small pair behind and between hind ocelli. Oc.
inserted between the bright-brown ocelli, rather upright, div. The rows of bristles on the
light frontal stripes end in a rather larger forward bent pair. Front view :—facial pit
deep and black with a shallow dividing ridge which is greyer: with a side light the pits
are also grey. Side view (Plate 21, fig. 56):—hind and lower eye-margins black, the
jowls greyer beneath these. Antennal second joint black bristled, the top bristle long,
third joint slightly pollinate, in the $ conspicuously reddened beneath, sometimes nearly
all red, in the ? black. Stout vibrissee with a row of small bristles along the absolute
lower edge of jowl; a large bristle on hind jowl. Palpi stout and bristled at the tip, in
the g much reddened on the proximal one-third to two-thirds; in the ? usually only
SECOND SERIES—ZOOLOGY, VOL. XVI. 47
364 PERCY SLADEN TRUST EXPEDITION
reddened at the base, though sometimes more extensively so. The long tip of tongue is
ochreous.
Thorax. The ground colour is black, but in good specimens it is densely covered
with a fine close granular ochreous-yellow pollination, which is best developed on the hind
part of the thorax and on the bare scutellum : this is demarcated from the thorax by an
extremely shallow line which is greyer. The dorsum (except the basal part) is uniformly
clothed with short black bristly hairs. Last two pairs of d. c, the hind pair largest,
prescut., two h., n. p., presut., three small bristles above wing, a very long one on post-
alar callus, with a shorter one inside it. No distinct pleural bristle except a stout sternopl.
over middle coxze. Scutellum with two basal and two long apical bristles crossed at the
tips. Wings, Fig. 43, glassy with pale, almost white, veins. Costal bristles long at
the base, bristly as usual on first segment with two spines at nick: black bristles as far
as second vein, the thickening extending, but attenuating, to the fourth. Halters clear
yellow, the stalks slightly darkened. Legs brownish black, middle and hind tarsi tawny
orange, knees narrowly the same.
Fig. 43. Desmometopa mauratum, n.sp. x 30.
Abdomen. Black, slightly shining, covered with bristly hairs, the submarginal row
being the longer, and especially so on the last segment, which is obtusely pointed, longer
than the preceding one, which is again longer than the next.
Size, 2mm.; fa little smaller; wing, 2 mm.
Localities, Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, from vege-
tation on the plateau, VIII.—IX. 1908. Mahé: Cascade Estate, at or above 800 feet,
1908—9 ; marshes on coastal plain, Anse aux Pins and Anse Royale, I. 1909.
In addition to the above species, which are represented by a fair number of indi-
viduals, there are two- more, one with but two specimens, the other unique: all are
unfortunately in rather bad condition, but the species are so marked in character that
short descriptions can be given.
61. Desmometopa microps, n. sp. (Fig. 44 and Plate 21, fig. 57).
There are two specimens, both rather damaged.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETID@, SEPSIDA, ETC. 365
Head. Top view :—frons, &c., much as last species: as far as can be seen the
bristles are practically the same. Front view :—face black, rather less deep than normal.
Side view (Plate 21, fig. 57) :—this is the most abnormal. The antennz are normal, but
the jowls are very large and swollen, and the hind eye-margins are also very wide : vibrissz
very large, followed by a few small bristles. The palpi are black and the tongue is pale
and horny.
Fig. 44. Desmometopa microps, n. sp. ~ 50.
Thorax. Much as last but the pollen has mostly disappeared ; cheetotaxy the same
as far as can be seen. The venation, Fig. 44, calls for no remark, except that the last
cross vein is a little nearer the wing margin.
In other respects, including the size, the species agrees with the last, except that the
abdominal segments are more nearly equal.
Locality. Seychelles. Mahé: Cascade Estate, at or above 800 feet, 1908—9.
62. Desmometopa approximatonervis, n. sp. (Fig. 45 and Plate 21, fig. 58).
This is a single female of a small species which has its cross veins extremely close
together. It is in rather a battered condition ; much stress cannot be laid on the degrees
of ‘‘shininess” of the head, &c., as the specimens may be somewhat “ greasy.”
Head. Top view :—frons black with browner line over forehead. Two lower f. o.,
two upper f. o. bent as usual, a finer bristle between each of above. I. v. and o. v.
normal ; crossed p. v., small parallel pair between these ; oc. div., between ocelli. The two
frontal rows of three pairs of bristles are quite long, there are a few over forehead. The
post-orb. row is long behind o. v. Front view :—the large antenne take up nearly all the
face, the pit is deep. Side view (Plate 21, fig. 58) :—jowls very narrow, pale; hind head
narrow black. Antennz black, second joint spined above, third large, round, brownish
black, arista moderately pubescent. Long vibrissa with inferior small bristles on jowl
edge, hind jowl with one or two bristles. Palpi pale orange, longer and less clubbed than
usual, bristly at end. Tongue long and thin, chitinous.
472
366 PERCY SLADEN TRUST EXPEDITION
Thorax. Shiny black with sparse yellow pollen, covered with fine bristles except
just at base. Stout last d.c., very fine prescut. A stout bristle on p. a. callus, other
bristles destroyed by pinning. Scutellum very minutely
; rugulose, two moderate basal, two long terminal bristles,
crossed at tip. Wings, as Fig. 45, moderately strong ciliation
; in first section costa, there are none of the strongish black
bristles visible on costa to end second vein, but the stout
Fig. 45. Desmometopa ciliation is uniform in character up to the fourth vein. Halters
approwimatonervis,n. sp. x30. brown, a little darker on tip. All the legs brownish black with
the tarsi paler tawny.
Abdomen. Shiny black, with brownish tone in parts, covered with black hairs, the
last segment is nearly twice as long as the previous one; the next are roughly equal in
length.
Size, about 1°4 mm.; wing, about 1'4 mm.
Locality. Seychelles. Mahé: Cascade Estate, at or above 800 feet.
Manpiza Fallen, Dipt. Suec., Oscinid., 8 (1820).
63. Madiza albipennis, n. sp. (Fig. 46 and Plate 21, fig. 59).
This is an almost entirely shining black insect with markedly milky wings.
Head. Top view :—frons black and shining, narrowest about opposite front ocellus,
widening just perceptibly in front: it is paler just over the forehead. The first of the
front f. 0. is close to the forehead, the second just in front of the hind pair: between are
three small bristles; both converge a little. The narrow eye-margins extend to the
front of the hind pair of f. 0., they are more shining than the rest in side illumination, as
is an evident ocellar triangle. The hind f. o. are fairly close, they diverge over eyes, have
“a small bristle between, and then two or three small ones up to thei. v.: these converge
somewhat; o. v. bent out, a small bristle between the i. and o. v. bent inwards: this is
really the last of the regular post-orb. row. P. v. almost exactly on vertex, pointing back
and sub-parallel. Oc. inserted rather nearer front ocellus than side one: a few small
bristles on ocellar area; four in a row over the forehead, a small pair on middle of frons.
Front view :—antenne widely separated at the base by an orange triangular projection of
the lunular arch which nearly meets a more obtuse orange upward projection of the lower
lip just about the level of the end of the antenna. The face has a roughly circular
outline almost cut into two deep pits by these two projections. Side view (Plate 21,
fie. 59) :—second antennal joint with a spine; third almost quite spherical, blackish with
pale silky pollen, arista slender, just perceptibly pubescent under a high power (x50). A
large upper vibrissa is apparently present, but its end is broken off, below it a smaller
one: jowl-margin with fine bristles. The jowls are very deep, shining black, and in good
light can be seen to be minutely striate longitudinally. Hind angle of jowl with a few
extra long bristles. When looked at from below the mouth is seen to be orange. Palpi
rather flattened and strap-like: rather orange dorsally, black below, sparsely bristled at
end and below. Tongue long and horny with a pointed tip.
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETID A, SEPSIDA, ETC. 367
Thorax. Quite smooth shining black clothed with fine black bristly hairs.
Cheetotaxy normal, except that a small pair of d. c. bristles are visible in front of the
usual last pair, and the tiny prescutellars are more visible. Scutellum somewhat brassy,
its surface minutely granulate; the usual four bristles,
terminal crossed. Wings, as Fig. 46, milky-white with
pure white veins: the costa black bristled to just beyond
the second vein, then slightly thickened to the fourth.
Halters with long yellowish-white heads and _ slightly
suffused stalks. The legs are black except the end of
front coxze, front and middle trochanter, and the middle Fig. 46. Madiza albipennis,
and last tarsi which are paler orange coloured except at eye
the distal ends of each jomt. The hind femora are distinctly flattened: middle tibize
with terminal spine.
Abdomen. Shining black, last segment a little longer than others, all clothed with
small moderately sparse black hairs: the long fleshy ovipositor is nearly half as long as
the abdomen.
Size, about 2°1 mm. without ovipositor ; wing, 12 mm.
Locality. Seychelles. Mahé: Cascade Estate, at or above 800 feet.
Rurcnorssa Loew, Wien. Ent. Monatschr., vi. (1862), p. 174.
64. Rhicnoessa ferruginea, n. sp. (Fig. 47 and Plate 21, fig. 60).
Head, distinctly concave behind in top view. Top view :—frons orange, slightly
shining: ocellar triangle grey ; eye-margins and two fine frontal stripes more yellow, one
outstanding lower f. 0. bent out, two upper f. 0. bent out and last somewhat back ; tiny
bristles between the f. 0., and an inner row of small bristles from level of middle f. 0. to
forehead. I. v. and o. v. on the back of the somewhat swollen vertex, finer convergent
p- v. well down on back of head; two ocellar inserted nearer front than side ocellus, two
small bristles between the hind ocelli, also between them and a few other bristles on the
ocellar area; the yellow frontal lines bristled from the level of front of the ocellar triangle
to lunule ; three pairs of these bristles are very much larger than others, and form three
well-marked macrocheetal crossed pairs: a similar pair overhang the forehead. Front
view :—the antennal bases touch, so that the yellow depression below lunule forms but a
single shallow triangular depression, in which is a pair of small bristles; the yellow lip is
arched and the clypeus very visible. Side view (Plate 21, fig. 60) :—antennee orange ;
2nd joint bristled above, one being very long; 3rd a little darkened, disc-shaped ; arista
with lower joint stout, orange, second less stout, black pubescent flagellum. Strong con-
vergent true vibrissa on upper angle of mouth, followed by two large supplementary
vibrissee and a few stoutish mouth bristles. Hind jowls bristly all over, especially long
below mouth. Post-orb. row regular from middle of eye upwards, a few strong bristles
behind v. b. Palpi orange, somewhat flattened laterally, slender and black haired :
tongue orange.
°
368 PERCY SLADEN TRUST EXPEDITION
Thorax. Ferruginous-ochreous, faintly pollinated with grey, only just perceptibly
greyer in a narrow central line; covered dorsally with uniform small black bristles;
between the lines of the d. c. these form six somewhat irregular rows of acrostichal bristles.
(1+8) d.c., the hind pair not in line with the other, but further apart ; prescutellars ;
a few small bristles on humeri, n. p., presut., 3 alar. There are also two small bristles
over the front coxee, one sternopleural, one large and one smaller one above along the side
of the mesopleural suture, almost in line with last notopleural. Scutellum bare, same colour
as thorax, flattened on disc, rounded, two basal and two crossed terminal bristles. Wings,
Fig. 47, slightly yellowish with yellow veins; 1st segment of costa rather bristly, the
small nick with few bristles, costa attenuating to 4th vein, with similar ciliation all
along. Halters all orange. Legs all yellow orange: a few longer bristles on front femora,
terminal spine to mid and last femora.
Licey en z B BAS eS Se
Fig. 47. Rhicnoessa ferruginea, n. sp. x 35.
Abdomen. Darker orange, almost brown-black in many cases, but much lighter on
the margins; covered with black bristly hairs, which are longer on the margins and sides
of the segments and very long on margins of the last three segments.
Size, about 24 mm.; wing, about 13 mm.
Localities. Seychelles: Mahé; a large series from seaweed on beach, Anonyme
Island, I. 1909; Long Island, VII. 1908, 1 specimen. Aldabra: 1908—9 (Fryer).
65. Rhicnoessa aurisetulosa, n. sp. (Fig. 48 and Plate 21, fig. 61).
This is an almost entirely greyish-yellow species with small eyes. All the bristles
are golden brown in side light, the macrochetes being a little darker, the small ones quite
pale.
Head. The cheetotaxy, is very like the last species, but will be repeated here for
convenience. Frons pale orange, ocellar area grey inside, pale pollinated grey just round
it as is the vertex, the narrowish eye-margin, and the paler frontal stripes. 3 f. o. as last,
the small row on inside eye-margins rather longer than last. I. v., 0. v., p. v. &c¢. much
as last. Front view :—the antennze quite touch at base, face pale yellow. Side view
(Plate 21, fig. 61) :—eyes much smaller, antenne all paler, 2nd joint similarly bristled but
rather more stoutly, arista very slightly slenderer. Jowls similarly bristled, but with
pale bristles: palpi quite yellow, almost pale, as is the tongue. The true approximating
LAMB—DIPTERA: HETERONEURIDA, ORTALIDA, TRYPETIDA, SEPSIDA, ETC. 369
vibrissa on the mouth angle is followed along the edge by 4 or 5 extra ones progressively
diminishing in size, and pointing away from vertical plane of head.
Thorax. Orange with a minute orange-grey granulation, dorsum all bristly with
fine bristles which form very irregular acrostichal rows of some 6 bristles wide between
the d. c.’s. There are (1+8) d.c., the last somewhat outside the line of the first three ;
prescutellars, one large humeral and two other smaller ones on the humeral callus ; two
not. ; presut., mesopleura with bristles, of which three are practically macrochetes, a stout
bristle and smaller one on prothorax over cox. Scutellum flattened, same colour as
thorax, the dorsum with a few scattered bristly hairs, a basal and crossed terminal pair.
Wings, as Fig. 48, glassy with pale yellow veins, no distinct variation in costal bristles,
though they are stouter from 2nd vein to base than from 4th to 2nd. Halters all yellow.
Legs all yellow, no very outstanding bristles, except upper and lower row on front femora ;
the mid tibia with terminal spines.
Fig. 48. Rhicnoessa aurisetulosa, n. sp. x 30.
Abdomen. In male each segment (2nd to last) has its basal half orange, sometimes
getting black from internal discolouration, and the distal half greyish orange: the line of
demarcation is very sharp ; bristles on dorsum, longer along hind margins, somewhat tufted
bristles (in top view) at upper side of 2nd segment. There is a rounded knobby genital
segment at end, not bent beneath, with small scale-like appendages ; the ? abdomen is all
orange, segments subequal, with a soft ovipositor.
Size, just over 24 mm. ; wing, just over 2 mm.
Localities. Seychelles. Mahé: from seaweed on beach, Anonyme Island, I. 1909,
a considerable number ; from seaweed, Long Island, VII. 1908, 6 specimens.
370 PERCY SLADEN TRUST EXPEDITION
Addenda and corrigenda to the author’s report on Sapromyzide, Chloropide &c.
(Vol. xv of these Transactions, Part 3, 1912, pp. 303—348, and Plates 15, 16).
Omission. , Family Sapromyzide.
In the paper on the Sapromyzide (op cit., pp. 307—316) the condition of the
terminal scutellar bristles of the genus Sapromyza was unfortunately omitted. They are
crossed or much convergent except on S. nudiuscula where they diverge strongly.
Nomenciature. Family Chloropide (op. c7t. pp. 329—343).
Just previously to the publication of the above an important paper by Herr T. Becker
appeared in the Ann. Mus. Nat. Hung. for 1912. In that paper the names oculata and
longipennis were used for two species of Oscinis. Hence those names are preoccupied.
On communicating with Herr Becker he was good enough (at the author's request) to
suggest names for the two Seychelles species whose names are preoccupied ; these names
are complicata for oculata and colorata for longipennis.
Further, on p. 249 of the same paper Herr Becker describes an Oscinella acuticornis
from Abyssinia which is the same as the Oscinis acuticornis of the above Seychelles
paper. Herr Becker also states that he considers this species and a similar one to be
representative of a new genus Paroscinella. The synonymy must therefore read :—
Os. oculata Lamb (op. cit., p. 340) = Os. complicata Beck., nom. nov.
Os. longipennis Lamb (op. cit., p. 342) = Os. colorata Beck., nom. nov.
Os. acuticorms Lamb (op. cit., p. 340) = Oscinella acuticorms Beck.
Note on Geographical Distribution.
Owing to the little knowledge there is of the enormous number of acalyptrate forms
that must exist in the world fauna, it is very difficult to draw any strong conclusions
from the insects recorded in this and the previous paper. This matter is rendered still
more inconclusive by reason of the fact that so many Dipterous genera have representatives
over almost all the world. The general aspect of the Seychelles acalyptrates, however, is
that of the African fauna. Several species are common to both (as for example Llythea
inwenata, Anatricus arenaceus, Zaprionus vittiger, Plagiostenopterina submetallica),
while many others are manifestly related to African forms (eg. Sapromyza striata,
Meroscins, Rhabdocheta).
The occurrence of the typical 8S. American genus Acrosticta is of much interest,
as it occurs also in Hawaii. A similar case is that of the Chilian genus Hccoptomma.
The New Guinea genus Allometopon is worth noting.
It is probable that further research will show that several of the new species are really
very widely spread, but the collecting of small flies is so much neglected, that it will be
long before any adequate information is available on the question.
The author desires to express his gratitude to Mr J. E. Collin, F.E.S., of Newmarket,
and to Herr F. Hendel of Vienna; both gentlemen freely placed their unrivalled knowledge
and judgment at his service.
LAMB—DIPTERA: HETERONEURIDAL, ORTALIDA, TRYPETIDAL, SEPSIDA, ETC.
PLATE 19.
Fig. 1. Heteromeringia plumicornis, n. sp. side view of head, x 40.
Fig. 2. - nigrifrons, n. sp., side view of head, x 40.
Fig. 3. Allometopon flavum, n. sp., side view of head, x 25.
F ig. 4. Acrosticta exsul, n. sp., front view of head, x 36.
Fig. 5. e » 0. sp., side view of head, x 40.
Fig. 6. Acidia seychellensis, n. sp., side view of head, x 32.
Fig. 7. Acidia (?)ocellata, n. sp., front view of head, x 20.
Fig. 8. 3 s n. sp., side view of head, x 20.
Fig. 9. Tephritis aldabrensis, n. sp., side view of head, x 36.
Fig. 10. Rhabdocheta spinosa, n. sp., side view of head, x 36.
Fig. 11. Sepsis albopunctata, n. sp., front leg of ¢, x 38.
Fig. 12. Drosophila sericea, n. sp., side view of head, x 36.
Fig. 13. y grossipalpis, n. sp., side view of head, x 36.
Fig. 14. , notabilis, n. sp., diagrammatic top view of head, x 36.
Fig. 15. 3 _ n. sp., side view of head, x 36.
Fig. 16. a fracticosta, n. sp., side view of head, x 36.
Fig. 17. ae nigerrima, 0. sp., side view of head, x 36.
Fig. 18. s nigrobrunnea, n. sp., side view of head, x 36.
PLATE 20.
Fig. 19. Drosophila punctiscutata, n. sp., side view of head, x 45.
Fig. 20. 3 aberrans, 0. sp., top view of head, x 45.
Fig. 21. re zs n. sp., side view of head, x 45.
Fig. 22. i spinipes, n. sp., top view of head (diagrammatic), x 45.
Fig. 23. a rf n. sp., side view of head, x 45.
Fig. 24. * - n. sp., front tarsus, x 50.
Fig. 25. * hirtipes, nu. sp., side view of head, x 45.
Fig. 26. 5 jucunda, nu. sp., side view of head, x 45.
Fig. 27. - rs finitima, u. sp., side view of head, x 45.
Fig. 28. a caliginosa, nu. sp., side view of head, x 45.
Fig. 29. * pallipes, nu. sp., side view of head, x 405.
Fig. 30. G triangulifer, n. sp., side view of head, x 45.
Fig. 31. a rufuloventer, v. sp., side view of head, x 45.
Fig. 32. si nasuta, u. sp., side view of head, x 45.
Fig. 33. i similis, n. sp., front leg of §, x 40.
Fig. 34. os 3 n. sp., side view of head, x 45.
Fig. 35. Chymomyza bicolor, n. sp., side view of head, x 45.
Fig. 36. Dettopsomyia formosa, n. sp., side view of insect, x 30.
Fig. 37. 3 2 n. sp., top view of head, x 45.
Fig. 38. i 2 n. sp., front view of head, x 45.
Fig. 39. ‘ 4s n. sp., side view of head, x 45.
Fig. 40. Echidnocephalus barbatus, n. g. and sp., side view of insect, x 20.
EXPLANATION OF PLATES 19—21.
SECOND SERIES—ZOOLOGY, VOL. XVI.
48
37 1
372
PERCY SLADEN TRUST EXPEDITION
PLATE 21.
Echidnocephalus barbatus, n. sp., top view of head, x 45.
n. sp., front view of head, x 45.
» »”
5 % n. sp. side view of head, x 45.
Chiromyia dubia, n. sp., side view of head, x 45.
seaspinosa, n. sp., side view of head, x 45.
hirtiscutellata, n. sp., side view of head, x 45.
Aphaniosoma sexvittata, side view of head, x 45.
Amygdalops thomasseti, n. g. and sp., side view of insect, x 25.
n. sp., diagrammatic top view of head, x 45.
” ”
n. sp., diagrammatic front view of head, x 45.
5 Ny n. sp., side view of head, x 45.
Milichia superba, u. sp., side view of head, x 32:5.
Milichiella tristis, n. sp. side view of head, x 45.
Eccoptomma solitariwm, n. sp. side view of head, x 32:5.
n. sp., top view of abdomen, x 15.
» ”?
Desmometopa inauratum, n. sp., side view of head, x 45.
macrops, nu. sp., side view of head, x 45.
2?
is approximatonervis, n. sp. side view of head, x 45.
Madiza albipennis, n. sp., side view of head, x 45.
Rhicnoessa ferruginea, n. sp., side view of head, x 45.
aurisetulosa, n. sp., side view of head, x 45.
22
Prrcy SitapEN Trust EXPEDITION.
(lisse) TRANS. LINN. SOC. SER. 2, ZOOL. VOL. XVI. PL. 19
C. G. Lamb et E. Wilson del.
Percy SLADEN Trust EXPEDITION.
(Lamp) SIS SVAUINES He SINGIN Eats © Gans eae © ©) ani) ae GV/plewl eae 2.0)
DIPTERA
C. G. Lamb et E. Wilson del.
Percy SuApEN Trust EXPEDITION.
(Lams)
TRAINS, IUNIN, SOC: SBIR. 2, ZOOIL, WOIL, 2OWIL, PIL, Zi
DIPTERA
C. G. Lamb et EH. Wilson del.
No. XVI.—DIPTERA : SCATOPSIDA, SIMULIIDA.
Von Dr. GUNTHER ENDERLEIN, STETTIN.
(Mit 1 Textfigur.)
(MirceTHEILT von Pror. J. Stanuey Garpiner, M.A., F.R.S., F.L.S.)
Gelesen den 5. Mirz, 1914.
Fam. Scatopside.
PsectroscraRA Kiefter*.
Psectrosciara Kieffer, Trans. Linn. Soc. London, ser. 2, Zool., vol. xv. 1912, p. 192,
Plate 9, figs. 12—14; Enderlein, ¢. ¢. p. 192; Arch. Naturg., xxvii. 1, Suppl. 3, 1911,
pp. 188, 192; Zool. Anz., xl. 1912, p. 280, figs. 12—14.
1. Psectrosciara mahensis Kiefter, /.c.
Unter dem neu erhaltenen Material findet sich noch ein weiteres Exemplar dieser
Form. Die basale Unterbrechung von m, ist sehr kurz.
Loc. Seychellen. Mahé: Cascade Estate, ca. 800 Fuss, 1908—9, 19. (Die Typen
sind im Museum von London und Stettin.)
RuHEGMOCLEMA Enderlein.
Rhegmoclema Enderlein, Zool. Anz., xl. 1912, p. 276, fig. 8.
Typus: Rhegmoclema rufithorax Enderlein 1912, Seychellen.
m, an der Basis nicht unterbrochen. cu, ist wellig, und zwar zwei—bis dreimal stark
gebogen. 1, und 77 laufen bei allen Arten wenig getrennt von einander. Der kurze
Radialramus scheint fiir die Gattung charakteristisch zu sein.
2. Rhegmoclema rufithorax Enderlein, /.c. (Fig. 1.)
?. Kopf rétlichbraun, Augen schwarz. Fiihler kurz, so lang wie der Kopf,
12-gliedrig; das 1. Glied kurz, das 2. dick und so lang wie am Ende breit, das 3.
* This genus was described by Kieffer (/.c.) from specimens accidentally sent to him among a number
of Chironomide: it was placed by him in Sciarine. Meanwhile Enderlein had also seen a specimen, and
referred it in manuscript to a new genus of the Lestremiine group of Sciaridz. On receiving later the
specimens described by Kieffer, he referred the genus, under Kieffer’s name Psectrosciara, to the Lestremiine,
in a paper in Arch. Naturg., U.c. This paper appeared before Kieffer’s description of the genus was published.
Finally, in a paper in Zool. Anz., l.c., Enderlein removed Psectrosciara to the Scatopside.—H. Scorr.
48—2
374 PERCY SLADEN TRUST EXPEDITION
schmal und so lang wie breit, das 4.—11. sehr breit und sehr kurz, das 12. kriftig, so lang
wie an der Basis breit, am Ende abgerundet und mit Spuren von 2 Einschnitten, schwarz-
braun, 3. Glied blass gelblich.
Thorax ziemlich glatt, rostgelb, ebenso das kurze und breite Scutellum, das mit einer
Anzahl schwarzer Haare besetzt ist. Riickenschild mit sehr feimer, aber kurzer und wenig
dichter schwarzer Pubescenz. Abdomen schwarz, ziemlich glatt, flach und breit, nach
hinten zu etwas verbreitert ; hinten abgestutzt abgerundet. Beine rostgelb, Schenkel
und Schienen leicht gebraunt. 4. und 5. Tarsenglied sehr schwach gebriiunt. Halteren
braun mit gelbem Stiel.
Eligel mit Spuren eines braunlichen Hauches; Costalzelle, Zelle R, und Zelle F
schwach gebriiunt. ¢, 7, und rr gelbbraun, die
tbrigen Adern sehr schwach briiunlich. 7, endet
etwas proximal vom Ende des ersten Fliigeldrittels,
vy sehr wenig proximal von der Fligelmitte.
Mediangabel lang, etwa 44 des ziemlich steil auf-
gerichteten Stieles; am Ende des zweiten Drittels
stark eingeschniirt und dann ziemlich stark nach
Fig. 1. Rhegmoclema rufithorax. x 40.
aussen divergierend. cv, gerade, den Fliigelrand
nicht erreichend. cw, etwa in der Mitte doppelt gebrochen und den Fliigelrand nicht
erreichend. Membran stark in allen Farben irisierend.
Korperlinge: 1,6—1,9 mm.
Fliigelliinge: 1,4—1,8 mm.
Toc. Seychellen. Mahé: Hochwald von Morne Blane und Pilot, Okt. bis Nov.
1908, 1 3, 6 2; Cascade Estate, 800—1500 Fuss, 1909, 2 3, 3 8.
Typen im Museum von London, Stettin und Cambridge.
Fam. Simuliide.
Smmutium Latreille, 1813.
3. Simulium speculiventre, nov. spec.
?. Kopf schwarz, Scheitel und Stirn etwas glatt. Fiihler 0°7 mm. lang, matt
schwarz mit weisslichgrauem Reif, 1. und 2. Glied meist ockergelb, zuweilen aber stark
geschwiirzt. Palpen matt schwarz.
Thorax schwarz, ziemlich matt, Seiten mit weisslichgrauem Reif. Riickenschild mit
wenig dichter gelber anliegender Pubescenz, ebenso das Scutellum. Abdomen matt
schwarz, der Kranz der langen Haare auf dem 1. Segment ockergelb; ein nach hinten
verbreitetes Mittelstreifen tiber die Tergite ist hinten poliert glatt, vorn wenig glatt. Die
nach oben etwas herumgreifenden Sternite sind an den Seiten mit gelben anliegenden
Haaren wenig dicht besetzt. Die Grenzen zwischen Tergiten und Sterniten sind eine
Spur heller und ein wenig erhaben. Coxen schwarz und etwas matt grau. Trochanter
rostgelb. Schenkel rostgelb, mit goldgelber anliegender Behaarung ; Hinterschenkel oben
am Ende mit einem graubraunem Fleck. Schienen rostgelb, am Ende schwarzlich grau,
ENDERLEIN—DIPTERA; SCATOPSIDA, SIMULIIDAL 375
Hinterschiene am Ende des ersten Viertels mit einem miissig scharfen schwiirzlichgrauen
Ring. Tarsen schwarz, mittlerer Metatarsus mit Ausnahme der Endhilfte, hinterer mit
Ausnahme der Endspitze, hell rostgelb. Metatarsus der Vorder- und Mittelbeie diinn.
Halteren-rostfarben, Stiel dunkler.
Fliigel hyalin, Adern rostgelb. 7, und 7, divergieren gleich von der Basis ab stark.
Korperlinge: 1,8—2,3 mm.
Fliigellinge : 2;—2,4 mm.
Loc. Seychellen. Mahé: Cascade Estate, 800—1500 Fuss, 1908—9, 5 2; Hoch-
wald hinter Trois Fréres, 1500—2000 Fuss, XII. 1908, 1 2; in der Ni’he von Morne Blanc,
C—O 1908, 42
Zu dieser Art gehért vermutlich das in Folgendem beschriebene ¢:
g. Augen schwirzlich. Fiihler gelblich graubraun. Thorax tief sammetschwarz,
Unterseite, Seiten, Hinterrandsaum, Scutellum und Seitensaum des Riickenschildes mit
weisslichorauem Hauch; auch der Vorderrandsaum des Riickenschildes ein wenig’ weiss-
lichgrau. Bei starker Vergrésserung erkennt man in der hinteren Hialfte des Riickenschildes
eine Anzahl anliegende braungelbe Haare. Abdomen tief sammetschwarz, der Kranz der
langen Haare auf dem 1. Segment gelbbraun. Zwei der mittelsten Tergite mit weiss-
lichgrauem Hauch. Beine gelbbraun, Metatarsus der Hinterbeine mit Ausnahme der
Endspitze mehr braungelb. Haltere dunkel rostfarben, Stiel braun. Fliigel hyalin, Adern
hell rostgelb.
Korperlinge : 2,2 mm.
Fligellinge: 2,2 mm.
Loc. Seychellen. Mahé: in der Nahe von Morne Blanc, X.—XI. 1908, 1 3.
Typen im Museum von London, Stettin und Cambridge (England).
Simulium ruficorne Macq. 1838 von der Insel Bourbon zeichnet sich durch rostgelbe
Fiihler und durch weisse Metatarsen der Mittel- und Hinterbeine aus; die Fliigel sind
weiss. Simulium griseicollis Beck. 1903 (Agypten) ist im miinnlichen Geschlecht mit
vorzugsweise auf der vorderen Hilfte des Thoraxriickens goldgelben filzigen Behaarung ;
Schwinger sind weiss; Thorax mit ihnlichem grauem Reif, der aber vorn durch 3 feine
schwarze Liingslinien unterbrochen wird. Simuliwm damnosum Theob. 1903 aus Ostafrika
hat das Riickenschild mit dichter hell goldener Pubescenz, Vorder- und Mittelbeine
schwarz, den Metatarsus der Hinterbeine mit einem breiten blassgelben Mittelband und
den Metatarsus der Vorderbeine angeschwollen.
No. XVII—COLEOPTERA, BUPRESTIDA.
Par Cu. KERREMANS.
(Avec 1 figure dans le texte.)
(CoMMUNIQUE PAR LE ProressEuR J. Srantey Garpiner, M.A., F.RBS., F.L.S.)
Lu le 5 mars, 1914.
1. Pseudocastalia fryeri, nov. sp.—Long. 9; larg. 3 millim.—Oblonga, antice
attenuata, deplanata, capite thoraceque zeneis, punctatis; fronte planata; thorace medio
sulcato, lateribus angulatim dilatatis; elytris testaceis, sutura costisque quinque nigris ;
subtus nigra, sparse griseo-pilosa.
Allongé, atténué en avant, aplani; téte et pronotum bronzés, élytres testacés avec la
suture et quatre cOtes noires; dessous noir, recouvert d’une pubescence
espacée et grise.
Téte plane, subrugueuse, grossiérement et inégalement ponctuée.
Pronotum plus large que long et plus étroit en avant qu’en arriére; la
marge antérieure tronquée et finement rebordée ; les cdtés obliquement
atténués depuis le tiers postérieur jusqu’au sommet, anguleusement
dilatés au tiers postérieur, brusquement convergents ensuite vers la base ;
le milieu du disque profondément creusé en arriére, l’impression prolongée
en sillon vers l’avant, interrompue avant le sommet ; la surface couverte
de gros points inégalement espacés. Hcusson trés petit. Elytres allongés, Fig. 1. ae aes
a cotés subparalléles, conjointement arrondis et multidentés au sommet, talia fryeri, nov. sp.
les deux dents externes plus fortes et plus espacées que les terminales ;
la surface présentant cing cdtes lisses: la premiére, pres de la suture, interrompue un peu
apres le milieu, les trois discales entiéres, la cinquiéme, prémarginale, interrompue un
peu avant le sommet, ainsi que la quatriéme, par le calus huméral ; les espaces intercostaux
avec deux séries de points épais et trés réguliers. Dessous assez grossitrement ponctué ;
pattes robustes, d’un noir bleuatre.
Loc. Aldabra: Ile Michel, X. 1908 (Fryer).
2. Inidotenia mahena Fairm., Bull. Soc. Ent. France, 1891, p. 46; Kerrem.,
Monogr. Bupr., t. iii. (1909), pl. 22, fig. 1.
Loc. Seychelles. Mahé: (Philibert); 1905 (Gardiner); 1908—9 (Scott). La Digue;
1892 (Alluaud).
378 PERCY SLADEN TRUST EXPEDITION
3. Chrysobothris dorsata Fabr., Mant. Ins., t. 1. (1787), p. 179.
Loc. Seychelles: Mahé; 1905 (Gardiner); 1909, “several specimens, including
larvee and pupe from dead ‘Agati’ wood (Adenanthera pavonina Linn.), given by R. P.
Dupont.” Tles Mascareignes, Afrique australe.
4. Dicercomorpha alluaudi Kerrem., Bull. Soc. Ent. France, 18938, p. 103.
Loc. Seychelles: La Digue, 1892 (Alluaud). Cette espéce n’a pas été retrouvee.
5. Belionota prasina Thunb., Nov. Ins. Sp., pars v. (1789), p. 90.
Toc. Seychelles: Mahé, 1908—9, “2 specimens, and larve and pup from dead
wood of the mango-tree.” Iles Mascareignes; Indo-Malaisie ; Afrique orientale.
6. Agrilus owas Cast. et Gory, Monogr. Bupr., t. u. (1837), p. 65, Pl. 14,
fig. 86.
Loc. Seychelles: Mahé; Cascade Estate, circa 800 pieds, 1 exemplaire. Mada-
gascar.
7. Sponsor pilosellus, nov. sp—hLong. 4,5; larg. 1,7 millim.—Oblongus, parum
convexus, antice posticeque rotundatus, totus niger, griseo-pilosus ; elytris profunde
arcuatim striatis, apicibus separatim rotundatis.
Ecourté, élargi, peu convexe, atténué et arrondi en avant et en arriére, les cotés
paralléles, entiérement d’un noir intensé et brillant, le pronotum couvert d'une courte
pubescence dressée, les élytres couverts d'une plus longue pubescence couchée, grises.
Téte convexe, finement pointillée; antennes gréles, dentées & partir du quatriéme
article. Pronotum assez grand et assez convexe, faiblement bisinué et finement rebordé
en avant, tronqué en arriére, largement arrondi sur les cotés avec une fine caréne latérale
oblique, les angles postérieurs légérement rentrants; une fossette préscutellaire ; la
surface couverte d'une fine ponctuation d’ot émerge une pubescence gris jaunatre et
dressée. Ecusson petit, inséré entre le rebord des élytres. Ceux-ci rebordés et sillonnés
le long de la base, presque droits sur les cétés, largement et séparément arrondis en
arriere, le sommet dentelé; la surface couverte de stries profondes et ponctuées, paralléles
et arquées; les interstices unisérialement ponctués. Dessous finement ponctué et
pubescent.
Loc. Aldabra; Takamaka, XI. 1908 (Fryer).
No. XVIIL—COLEOPTERA ; PLATYPODIDA AND IPIDA FROM THE
SEYCHELLES ISLANDS.
By Lr.-Cot. Winn Sampson, F.E.S.
(Text-figures 1—5.)
(CommuNIcaTED BY Pror. J. Stantey Garpiner, M.A., F.R.S., F.L.S.)
Read 5th March, 1914.
Havine been requested by Mr Hugh Scott, the Curator in Entomology in the
University of Cambridge, to determine the Platypodide and Ipide collected during
the Percy Sladen Trust Expeditions to the Indian Ocean in 1905 and 1908—9, the
following pages show the results of my examination of the material handed to me.
The 1905 collection consists of only one specimen without antennee, but appears to
be a species of the genus Triarmocerus, of which T. birmanus Eichh., has been described
from Burma, and 7. cryphaloides Hichh., from Madagascar.
In the 1908—9 collection, the Platypodide are represented by a single genus and
species, and the Ipidee by 9 genera, comprising 22 species, of which 8 are new (as also is
one variety of an old species). In addition, references are given to 3 other species (and
a somewhat doubtful fourth) recorded as found in these islands by previous collectors, but
not found by this Expedition, so that in all 25 (or possibly 26) species of Ipidz are
enumerated from the Seychelles.
It has been necessary to make a new genus, Sciatrophus, for a very curious beetle
found only at Praslin in the Vallée de Mai, and taken there solely from the leaf-bases of
growing gf coco-de-mer ; nothing at all resembling this insect was found in any of the
other islands, and for the reasons given below, the genus must be placed on the debatable
land between the Curculionide and Ipide, at any rate for the present.
There are a few species of the genus Cryphalus which are left undetermined ;
Hichhoff described a good many single specimens from India and Burma, but as his types
are lost or at any rate not available, no identification on his descriptions alone would be
satisfactory.
In this paper, Stephanoderes and Hypothenemus are not treated as subgenera of
Cryphalus. i
Types. A first set of the material, including types of all new forms, will be placed
in the British Museum: a second set will be retained at the Cambridge University
Museum.
SECOND SERIES—ZOOLOGY, VOL. XVI. 49
380 PERCY SLADEN TRUST EXPEDITION
Genus incertz sedis.
ScCIATROPHUS, gen. nov. (Figs. 1—5).
Caput elongatum, depressum ; submentum valde in medio productum, maxillee lobo
intus setis rigidis ciliato, mentum latum, ovale, antrorsum subangustatum lateribus fortiter
rotundatis ; antennee scapo plano triangulo ; funiculo 5-articulato, hujus articulo 1° trun-
cato pyriformi, 2° minore, sequentibus transversis, latitudine vix crescentibus, capitulo
oblongo, acuminato, solido sed transversis pilis notato. Pronotum transversum, a prosterno
carina seyunctum ; coxe anteriores leviter distantes, tibiee anticee versus apicem leviter dila-
tate, apice rotundatz, extus non serrate, tarsorum articulus 3 bilobus. Scutellum parvum.
Head flattened and broad, visible beyond the thorax from above; the labial and
maxillary palpi three-jointed (Figs. 3, 4); labrum distinct, submentum prolonged medianly
upwards into a long peduncle, the mentum being attached to its inner side towards the
apex (Fig. 3); the lateral arms of the hypostoma very long; the antenne (Fig. 2)
elbowed, with a depressed triangular scape and a funiculus of fine joints, the first truncate
pyriform, the second rather shorter, and the remainder hardly increasing in size towards
the club, which is narrow and cone-shaped with two transverse rows of hairs, one near the
base and the other round the centre, the apex having a tuft of pale hairs with 2—3
indistinct rows of hairs just below. The eyes prominent and nearly round, the antennz
inserted below the dorsal margin of the head beneath the eyes, whilst the pronotum
is divided from the prosternum by a ridge; the anterior coxe are placed moderately close
together, the tibize being somewhat dilated towards the apex, and not having any serration
on the outer edge; the tarsal joints short and the third bilobed (Fig. 5).
1. Sciatrophus fuscus, sp. n. (Figs. 1—5).
Oblongo-ovalis, fuscus, pubescens, thorace subquadrato sed lateribus leviter rotundatis,
zqualiter minutissime granulato, pubescente ; elytris convexis, setosis, subtilissime lineato-
punctatis, lateribus subparallelis, apice rotundato, interstitiis planis, punctatis piliferis.
Long. 1°5 mm.
Head and prothorax black, elytra varying from light brown to black, but covered
with longish pale yellow hairs ; the head is produced anteriorly,
minutely rugose and pubescent, with a short polished longi-
tudinal carina in the centre over the mouth, and several rugose
longitudinal lines on the considerably raised surface of the
anterior margin below the eyes which are somewhat prominent
and round, but so placed on the lateral edges of the head as to
be raised longitudinally down the centre. The antennez
(Fig. 2), which are geniculate, have the scape as long as the
funiculus together with half the club, and it is basally very thin,
but becomes somewhat abruptly widened on the outside into a
large triangular surface ; the club is acuminate, not compressed,
and only slightly broader than the first funicular joint, it has no
sutures, but there are two circles of hairs, one near the base
*Hic. 1. Sciatrophus fuscus.
oO b)
gen. et sp. nov. 9, x 24.
* Unfortunately the specimen figured was not in a good position. Its head is so much depressed that the
elongation of the front is not visible, and the thorax is too much drawn away from the elytra, so that its base
appears too curved.
SAMPSON—COLEOPTERA ; PLATYPODIDAI AND IPIDA 381
and the other about the middle, with three other very minute ones close to the apex,
which has, in addition, a tuft of whitish hair on the extremity. Mouth-parts (Figs. 3, 4).
The mandibles are short and stout, with a sharply bent apical point and a large blunt
tooth below. The submentum is produced anteriorly in the centre beyond the junction of
the mentum, which is consequently hidden but is heart-shaped; the ligula is very small,
the palpi three-jointed, the two first joints beige transverse and the third longer than
broad, as well as being slightly larger at the apex, which is uniformly rounded. The
maxillze are armed with a few strong spines on the inside, the apical ones being curved
inwards, the palpi are three-jointed, the two first transverse and the third cylindrical ;
the labrum is bisinuate with some longish hairs near the apex.
The prothorax is one-third broader than long, minutely granulate over its whole
surface and covered (in perfect specimens) with yellow hairs; there is.a very faint median
longitudinal line, the sides are slightly rounded and narrowed anteriorly, the anterior and
posterior edges being straight and the latter faintly margined. The anterior legs
Sciatrophus fuscus, gen. et sp. nov.
Fig. 2. Antenna. Fig. 3. Mouth-parts from beneath (labrum not shown).
Fig. 4. Maxilla. Fig. 5. Anterior tarsus.
(Fig. 5) have the coxee nearly spherical, the trochanters very small, the femora largely
‘dilated and furnished with a small lobe on the inner apical surface similar to that found
in Trypodendron and other genera; the tibie are gradually enlarged apically, with strong
bristles and one very minute spine or tooth on the inner side, otherwise they are quite
plain; the two first tarsal joints are very short, the first nearly quadrate, the second
transverse, the third deeply bilobed and two-thirds the length of the first two joints com-
bined ; the fourth joint is very minute, the fifth nearly as long as the first three taken
together.
49—2,
382 PERCY SLADEN TRUST EXPEDITION
The scutellum is very minute; elytra nearly twice as long as the prothorax and
slightly wider than the latter at their middle, lmeate punctate and covered with longish
yellow hairs which are finer than those on the prothorax, the interstices very faintly
punctate and furnished with similar hairs, the apex rounded; viewed laterally the
declivity is very abrupt; the under surface is transversely shagreened, especially the
centre portion of the mesosternum; the anterior cox are moderately close together,
the middle ones being farther apart, whilst the posterior are still more so. The first and
second exposed abdominal segments are in some specimens obscurely separated, the
third and fourth very narrow and the fifth equal to the two preceding ones in breadth.
Loc. Seychelles. Praslin: 40 specimens were found between the leaf-bases of a
& Coco-de-mer (Lodoicea seychellarwm) which was felled for collecting purposes in the
Vallée de Mai, Cétes d’Or Estate, 28. XI. 1908*.
Platypodidee.
Puatypus Herbst.
2. Platypus lepidus Chap., Mon. Platypides, 1866, p. 282, fig. 171.
In the series of both sexes of this species from Seychelles there is considerable
difference shown in size, some being larger and others smaller than the type; there are
also variations in the number and arrangement of the prothoracic punctures on the
females, but these differences are not sufficient to constitute definite varieties. M.Chapuis
pointed out (op. cit., p. 283), “Les exemplaires des différentes localités que nous avons
signalées different quelque peu entre eux; ainsi, les deux petits groupes de points du
corselet peuvent manquer chez la femelle; la forme est un peu plus étroite dans individu
provenant des iles Philippines.”
Loc. Seychelles. Silhouette: near Mont Pot-d-eau, ca. 1500 ft, WII. 1908;
Mare aux Cochons, over 1000 ft. Praslin: Cétes d’Or Estate, XI. 1908. 36 specimens.
In Silhouette (VIII.—IX. 1908) this beetle frequently came to the lamp at night in the
huts near Pot-a-eau and at Mare aux Cochons; the hut-walls were built of freshly split
lathes of the palm Verschaffeltia, but whether the Platypus had any connection with this
wood or was merely attracted from the surrounding forest is uncertaint. Celebes, Molucca
and Philippine Islands.
Ipide.
CRYPHALUS Er.
3. Cryphalus pallidus Hichh., Berlin. Ent. Zeitschr., xv. 1871, p. 181.
5 specimens.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908. Mahé:
Cascade Estate, ca. 1000 feet ; Long Island, VII. 1908. Madagascar.
* Scratrophus fuscus is not the only species of beetle which was found only between the bases of the
leaves of this remarkable tree. Atenius lodoicew was almost exclusively so found (see Vol. xv. of these
Transactions, p. 226, 1912), also a fine Scydmsenid which has not yet been studied.—H. Scorr.
7 One or more of the species of Xyleborus also came sometimes to the light, but unfortunately there
is no certain record to show which species.
SAMPSON—COLEOPTERA; PLATYPODIDA AND IPIDA 383
4. Cryphalus trypanus, sp. n.
g. Oblongo-ovalis, fusco-testaceus ; caput globosum insertum, fronte fusca, medio
transverse carinata; mentum longissime versus basin angustatum, apice utrinque oblique
truncato, ligula prope menti medium inserta, maxillze mala oblonga, setis rigidis ciliata ;
palpi labiales articulis 1° et 2° fere quadratis, 3° cylindrico, apice oblique truncato, parvis
tuberculis ornato. Antennze funiculo 4-articulato, articulo 1° crasso, subgloboso, 2° sub-
quadrato, sequentibus transversis, latitudine sensim crescentibus ; capitulo infuscato,
articulo 1° ovali, ceteris lunatis, suturis incurvatis. Prothorax transversus, margine apicali
tuberculis ordinatis, antrorsum angustatus, dorso anterius scabrato, posterius squamulis
ornato. Elytra cylindrica lineato-punctata, squamulis setisque seriatis vestita, latitudine
thoracis et illo paululum longiora, lateraliter a basi ad medium subrecta, deinde arcu
subzequali ad apicem obtuse rotundata. . Tibize angustze sublineares, versus apicem leniter
dilatatze, subrectze, extus spinuloso-dentatze, valde pilosee, apice oblique truncatee ; tarsi
articulo 3° cordato.
Long. 2 mm.
?. Mari similis sed prothorace antrorsum minus angustato ; tibiis minoribus capillis
ornatis, fronte medio longitudinaliter carinata. Long. 2 mm.
Head black, rugose, with a slight median longitudinal line meeting a much more
prominent and shiny transverse ridge near the vertex of the head ; the antennal club has,
on its upper side, three very curved sutures, whilst those on the under side are much
straighter and four m number; the eyes are emarginate. The prothorax is transverse
and considerably narrowed towards the apex, dark-coloured, with transverse rows of rather
scattered elevations anteriorly, which decrease somewhat in size up to the gibbous portion
of the dorsum, which is situated behind the centre near the posterior margin; at this
point the prothorax is yellowish in colour, being covered with recumbent scales; the
anterior edge of the prothorax is furnished with 6—8 somewhat prominent tubercles, and
the whole of the darker part is covered with semi-erect hairs; the posterior edge is
margined, as are also the sides, almost half-way to the front. The anterior tibiz are
furnished with long and coarse hairs on the inner edge, whilst in the females these hairs
are either absent altogether, or are very fine and short. The scutellum is very minute
and dark. The elytra are very slightly longer than broad, and about one-fifth longer
than the prothorax, having longitudinal lines of shallow punctures, the interstices flat,
broad, and covered with small pale scales with a single row of longish squamulate hairs
down the centre ; the small interstitial scales and some of those on the prothorax are of a
rather peculiar shape.
Not unlike C. vestetus and C. fuliginosus Blandf. but differs outwardly from the
former in not possessing a dense pubescence, from the latter by the structure of the
sutures of the antennal club, and from both these species by the prominent tubercles
on the prothorax. The obliquely truncate labial palpi with their minute terminal
tubercles are probably common to many other species of the genus, they are seen in
C’. tiie Panz., in which species the tubercles are rather longer; the cordate third tarsal
384 PERCY SLADEN TRUST EXPEDITION
joint is very similar to that of C. picee Ratz., and the lateral margination of the prothorax
is like that of C. abietis and C. picew Ratz.
Dissection shows that the anterior inner surface of the prothorax in the male is
provided with a transverse carina similar to that on the head, but having on its edge and
front side (which is somewhat convex) a series of minute longitudinal ridges, forming what
is probably a stridulating area, whilst the strongly developed carina on the head, which is
certainly capable of being withdrawn under the prothorax by an upward movement of the
head, may act as a scraper.
Mr Charles J. Gahan, in his very interesting paper on “Stridulating Organs in
Coleoptera” (Trans. Ent. Soc. London, 1900, Pt. IIL, p. 4386), describes a somewhat
similar apparatus in Spilispa imperialis Baly, but the position of the file and scraper are
there reversed, and the shape of both parts are very different to those of C. trypanus ; in
the female of C. trypanus the inner surface of the prothorax is quite smooth.
50 specimens.
Loc. Seychelles. Silhouette ; from near Mont Pot-d-eau, ca. 1500 feet, and
Mare aux Cochons and high forest above, VIII.—1IX. 1908. Mahé: high forest of Morne
Blane and Morne Pilot, up to ca. 2000 feet, X.—XI. 1908; high forest between Trois
Freres and Morne Seychellois, 1500—2000 feet, XII. 1908; Mare aux Cochons district,
1000—2000 feet, I.—II. 1909, including some specimens from stunted “ capucin ”
(Northea) forest on summit of “ Montagne Anse Major,” over 2000 feet. Marie Anne
Island: XII. 1908, one specimen.
On one occasion a number of specimens were captured, one after the other, flymg in
broad daylight in a small space among the trees in the high mountain-forest of Mahé.
STEPHANODERES Hichh.
5. Stephanoderes communis Schauf., Tijdschr. Ent., xxxiv. 1891, p. 11.
For the same reason as that given regarding Xyleborus madagascariensis, these —
insects are placed provisionally in the above species ; they slightly exceed the length
given in the description (14 mm.) and the interstices are not costate.
Loc. Seychelles. Mahé: Mare aux Cochons district, over 1000 feet, I.—II. 1909,
one specimen; Long Island, VII. 1908, two specimens. Madagascar.
6. Stephanoderes vulgaris Schauf., Tijdschr. Ent., xl. 1897, p. 209; Kolbe, Mitt.
Zool. Mus. Berlin, v. 1910, p. 40.
Not obtained by the Percy Sladen Trust Expedition.
Loc. Seychelles: La Digue, 1892 (Alluaud).
Nors. There are several undetermined specimens of Cryphalus from Silhouette and
Mahé, and there is one undetermined specimen of Stephanoderes from Silhouette.
SAMPSON—COLEOPTERA ; PLATYPODIDA AND IPIDA 385
HypotHENEMUS Westw.
7. Hypothenemus eruditus Westw., Trans. Ent. Soc. London, 1836, p. 34.
Stephanoderes arecce Kichh., Rat. Tom., 1879, p. 165.
4 specimens.
Loc. Seychelles. Silhouette : low coconut-planted country near the coast at Pointe
Etienne, IX. 1908. Mahé: Cascade Estate, ca. 1000 feet ; Long Island, VII. 1908.
Praslin: Cotes d’Or Estate, XI. 1908. Very widespread, probably cosmopolitan.
8. Hypothenemus validus Blandf., var. valens, noy. var.
Hypothenemus validus Blandf., Biol. Centr.-Am., Col. iv., pt. 6, 1904, p. 228.
H, validi similis sed angustior, tuberculis prothoracis minoribus ; plaga fusco-rufa.
There is nothing to separate this insect from Mr Blandford’s species, as far as can be
seen in the single specimen before me, except the slight differences mentioned above.
Loc. Seychelles. Silhouette: Mare aux Cochons, over 1000 feet, IX. 1908, one
specimen.
The typical form of H. validus was described from Mexico: the very closely-allied
H. opacus Hichh. is known from South America.
CRYPHALOMORPHUS Schauf.
9. Cryphalomorphus crenatus, sp. n.
g. Ellpticus, piceo-brunneus, subnitidus, tenue pubescens, capite globoso sparse
punctato, antice breviter pallide-ciliato, linea media longitudinali, thorace latitudine longiore,
apicem versus rotundato-angustato, dorso valde convexo, margine apicali medio tuberculis
6—8 prominulis notato, anterius valde scabrato minutissimis pilis ornato, postice dense
asperato, medio utrinque oblique impresso ; elytris thoracem latitudine prope zequantibus,
et illo vix sesqui longioribus, convexis, apice rotundato, declivi, interstitiis rugulis multis
transversis asperatis sed ante apicem granulatis, seriatim pilis longioribus et brevioribus
adspersis ; stria suturali impressa, ad scutellum depressa.
Long. vix 1°75 mm.
?. Mari similis sed tuberculis margine apicali medio minus conspicuis aliterque
ordinatis.
Long. 2 mm.
In the males, the tubercles on the prothoraciec anterior edge are divided into two
groups by a central narrow longitudinal space which can be traced some distance
posteriorly ; the anterior half of the prothorax is more depressed than in the females and
is more acutely rounded apically, the male elytra also terminate more acutely ; when
viewed dorsally the male is slightly shorter and narrower than the female with the sides
somewhat curved, and laterally the declivity of the elytra is more abrupt than in the
latter sex, but these differences vary considerably in individuals of the series and form no
386 PERCY SLADEN TRUST EXPEDITION
very certain indication of the sex in isolated specimens. The antennze which I have been
able to examine microscopically, after mounting in balsam, all exhibit, without exception,
the diagonal sloping suture in the club mentioned by H. Schaufuss.
This species is easily distinguished from C. commumus by the different elytral
sculpture, the longitudinal line on the head, and by the anterior thoracic tubercles ;
it differs from the following species by the lesser number and smaller size of the tubercles
before the declivity, whereas in the latter the granulations are continuous from base to
apex.
Loc. Seychelles. Mahé: 43 specimens, all from high damp forest between Trois
Fréres and Morne Seychellois, 1500—2000 feet, 9 and 10. XII. 1908; probably from bark
or wood of the endemic Wormia ferruginea, the “Bois Rouge,” though this is not quite
certain.
10. Cryphalomorphus confragosus, sp. n.
Ellipticus, piceo-brunneus, tenue pubescens, capite globoso, minutissime granulato, in
margine antico pilis griseis ciliata, linea media longitudinali, thorace latitudine longiore,
apicem versus rotundato-angustato, dorso valde convexo, margine apicali tuberculis com-
pluribus obsoletis notato, anterius valde scabrato, minutissimis pilis ornato, postice dense
scabrato, medio utrinque oblique impresso ; elytris thoracem latitude prope zquantibus,
et illo vix sesqui longioribus, convexis, apice rotundato, declivi, interstitiis granulatis
a basi ad apicem.
Long. 1°5 mm.
In this species, the prothorax is narrower medianly than in the preceding, thus
conforming more closely to C. communis, but the elytral sculpture is very different from
either of the other species. In both the new species the antennz are exactly similar in
shape and structure to those described by H. Schaufuss.
Loc. Seychelles. Mahé: Cascade Estate, 800—1000 feet, I. 1909, one specimen.
XyLEBORUS Hichh.
11. Xyleborus semigranosus Blandf., Trans. Ent. Soc. London, 1896, p. 211.
13 specimens.
Loc. Seychelles. Silhouette: from near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908,
and Mare aux Cochons and forest above, IX. 1908; some specimens from fungus. Mahé:
Cascade Estate, about 1000 feet, and from forest above, II. 1909. Sumatra, from
tobacco.
12. NXyleborus madagascariensis Schaut., Tijdschr. Ent., xxxiv. 1891, p. 23.
I place these insects provisionally in the above species, as | am unable to obtain the
loan of any specimens from Herr Schaufuss for comparison, but they do not differ
materially from the description. About 20 specimens.
Loc. Seychelles. Silhouette : found boring in the stem of an introduced species of
Dracena, at the edge of the Mare aux Cochons plateau, over 1000 feet, IX. 1908.
Madagascar.
——
ee
SAMPSON—COLEOPTERA ; PLATYPODIDA AND IPIDA 387
13. Xyleborus andrewesi Blandf., Trans. Ent. Soc. London, 1896, p. 227.
Loc. Seychelles: Silhouette, Mare aux Cochons, IX. 1908, one specimen. India,
Belgaum (Andrewes).
14. Xyleborus perforans (Woll.).
Tonicus perforans Woll., Cat. Col. Mader., 1857, p. 96.
Xyleborus kraatz Kichh., Berlin. Ent. Zeitschr., xii. 1868, p. 152; Kolbe, Mitt.
Zool. Mus. Berlin, v. 1910, p. 40.
Unfortunately, in this series of over 40 specimens there is not a single male, so that
determination is very difficult ; nearly all the specimens have much more the appearance
of a typical X. perforans Woll. (=kraatz Hichh.) than of X. affinis Eichh., which latter,
Prof. Kolbe (/.c.) states to have been taken in Mahé.
Loc. Seychelles. Silhouette, VITI.—IX. 1908: near Mont Pot-i-eau, ca. 1500 feet;
Mare aux Cochons and the forest above, up to the highest parts; also some specimens from
low cultivated country near Pointe Etienne. Mahé: high forest between Trois Fréres and
Morne Seychellois, 1500—2000 feet, XII. 1908 ; Cascade Estate, ca. 1000 feet; Anonyme
Island, I. 1909. Praslin: Cotes d’Or Estate, XI. 1908. X. kraatzi has been previously
recorded from Mahé and La Digue, 1892 (Alluaud).
In the Seychelles this species was taken in many localities and at various elevations.
Tt is very widely distributed. In my collection there is a single specimen that I took in
Old Calabar, S. Nigeria, some years ago.
15. Xyleborus torquatus Kichh., Berlin. Ent. Zeitschr., xii. 1868, p. 146.
’ Xyleborus badwus Hichh., t.c., p. 280; Kolbe, Mitt. Zool. Mus. Berlin, v. 1910, p- 40.
Loc. Seychelles. Silhouette: from near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908,
2 specimens. (X. badius is recorded from La Digue, 1892 (Alluaud).) Central and South
America, Madagascar, Mauritius, Japan, Tahiti.
16. Xyleborus confusus Hichh., Berlin. Ent. Zeitschr., xi. 1867, p. 401; Kolbe, Mitt.
Zool. Mus. Berlin, v. 1910, p. 41.
Xyleborus insularis Sharp, Trans. R. Dubl. Soc., iii. 1885, p. 193; Faun. Hawaiiens.,
ii. pt. 3, Col. 1, 1900—1903, p. 177.
Bostrichus ferrugineus Bohem., Voyage de |’ Kugénie, Ent., 1858, p- 88.
Loc. Seychelles. Silhouette: from near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908;
Mare aux Cochons and forest above, up to highest parts, some specimens being from
fungus. Mahé: Long Island, VII. 1908; forest above Cascade Estate, 1000—2000 feet,
Il. 1909. Praslin: Cotes d’Or Estate, XI. 1908. Previously recorded from Seychelles by
Boheman (/.c.). Madagascar, German East Africa, Sandwich Islands, South America.
17. Xyleborus dilatatus Eichh., Rat. Tom., p. 393; Kolbe, Mitt. Zool. Mus. Berlin,
v. 1910, p. 41. |
Not obtained by the Percy Sladen Trust Expedition.
Loc. Seychelles: Mahé and La Digue, 1892 (Alluaud). Mauritius.
SECOND SERIES—ZOOLOGY, VOL. XVI. 50
388 PERCY SLADEN TRUST EXPEDITION
18. Xyleborus bucco Schauf., Tijdschr. Ent., xl. 1897, p. 212; Kolbe, Mitt. Zool.
Mus. Berlin, v. 1910, p. 41.
Not obtained by the Percy Sladen Trust Expedition.
Loc. Seychelles: La Digue, 1892 (Alluaud).
[ Xyleborus affinis Eichh., Berlin. Ent. Zeitschr., xi. 1867, p. 401; Kolbe, Mitt. Zool.
Mus. Berlin, v. 1910, p. 41.
This species is stated by Kolbe (/.c.) to have been taken by Brauer in Mahé. It was
not obtained by the Perey Sladen Trust Expedition. It has been referred to under
Xyleborus perforans. |
19. Xyleborus abruptus, sp. n.
Cylindricus, subnitidus, fusco-testaceus ; prothorace subgloboso, margine antico sub-
crenato, dorso medio valde transverse elevato, postice punctato et in margine basali medio
densius pubescente ; elytris quam prothorace paululum longioribus, indistincte lineato-
punctatis, interstitiis confuse punctatis, apice oblique truncato-retuso, margine per totum
acuto, subcirculari, fundo utrinque ad suturam gradatim elevato et medio impresso, nitido,
interstitiis perspicue irregulariter punctatis.
Long. 3°5 mm.
Head convex, coarsely roughened between the eyes, the rest of the front finely
shagreened ; the eyes are placed near together and are large, emarginate, with coarse
facets, there is a transverse row of pale hairs over the mouth and a faint short longi-
tudinal line in the centre just above; the prothorax is large and globose, with the
posterior angles obtuse and the sides rounded, anteriorly asperate and pubescent, slightly
depressed behind the strongly marked transverse dorsal ridge, the depression covered
with a patch of piliferous punctures intersected longitudinally by a sub-elevated median
line; the elytra are very slightly longer than the prothorax, expanding somewhat towards
the declivity, but nowhere exceeding the greatest breadth of the prothorax, the sides are
straight until the declivity, when they become abruptly truncate, each ending apically in
‘a blunt spine (as seen from above) caused by the sudden thickening of the sutural edges ;
the elytra become very dark apically, the apical truncate surface is nearly vertical, shiny,
with four impressed and curved striz on each side of the suture; the upper edge of the
declivity is clothed with a fringe of pale hairs, and the fundus has an irregular surface,
depressed on each side of the suture near the apex ; the irregular interstitial punctures
on the fundus are very evident.
Closely allied to X. amputatus and X. mancus Blandt., especially to the latter, from
which the present species only differs in the shape of the prothorax, the number of rows
of punctures on the fundus, and the punctuation of the interstices. Three specimens.
Loc. Seychelles. Mahé: high forest of Morne Blanc, X. 1908, and from near
Morne Blanc, XI. 1908; Cascade Estate, ca. 1000 feet, I. 1909.
SAMPSON—COLEOPTERA; PLATYPODIDA AND IPIDA 389
Eccorrorpterus Motsch.
20. Hecoptopterus sexspinosus Motsch., Bull. Mose., xxxvi. 1863, I. p. 515.
Xyleborus abnormis EKichh., Berlin. Ent. Zeitschr., xii. 1868, p. 282; Rat. Tom.,
1879, p. 341.
Xyleborus sexspinosus Haged., Coleop. Catalog., Ipidee, 1910, p. 110.
Loc. Seychelles. Silhouette: Mare aux Cochons and forest near by, IX. 1908,
6 specimens. Burma, Ceylon, Java, Manila, Cameroons, German East Africa, and a
variety X. multispinosus Haged. from Sumatra. I have specimens from Borneo, Celebes,
India (Nilgiri Hills) and New Guinea.
TRIARMOCERUS Hichh.
21. Triarmocerus, sp.?
As stated above, the absence of the antennze in the single specimen before me,
prevents any trustworthy determination ; the tibize are similar to the generic description,
as given by Eichhoff, although the specimen does not agree in other points with either of
his described species.
Loc. Coetivy, 24. TX. 1905.
Dryocares Eichh.
22. Dryocetes furvus, sp. n.
Oblongus, subfuscus, piceo-niger, antennis pedibusque testaceis ; prothorace oblongo-
ovali anterius rugulis transversis exasperato, posterius rugulose punctato, elytris ad
apicem oblique declivibus, lineato-punctatis, interstitiis post medium uniseriatim pilis
erectis obtusis ornatis.
Long. 2°25 mm.
Differing from D. nubilus Blandf. in being slightly narrower, the prothorax rather
less rounded basally and the median smooth line less distinct ; the setze more obtuse and
the general colour of the insect less shiny.
Loc. Seychelles. Silhouette: from near Mont Pot-d-eau, ca. 1500 feet, VIII. 1908,
one specimen.
Coccotryrees Eichh.
23. Coccotrypes pygmeus Hichh., Rat. Tom., 1879, p. 310.
Loc. Seychelles. Mahé: high forest of Morne Blane and Pilot, XI. 1908, one
specimen. Madagascar; recorded also from Manila, West Africa, and Santo Domingo.
KEY TO THE THREE NEW SPECIES OF COCCOTRYPES.
globular, with apical prothoracic tubercles 008 ae parvus, sp. n.
1. Prothorax
longer than broad 2
strongly oblique, no anterior tubercles on thorax ce bicolor, sp. n.
2. Hlytral declivity
le oblique, with anterior tubercles on thorax ... one declivis, sp. n.
50—2
390 PERCY SLADEN TRUST EXPEDITION
24. Coccotrypes bicolor, sp. n.
Oblongus, thorace nigro, semiovato, toto crebre exasperato; elytris ferrugineis,
lineato-punctatis, alternatim longius et brevius seriatim pilosis, apice fortiter obliquo,
utrinque subimpresso-retuso.
Long. 3 mm.
Head black, longitudinally strigose especially laterally, with a short shiny carina
posteriorly in the centre, and a transverse row of yellow hairs over the mouth. Prothorax
black and semi-ovate, with the base truncate, rounded at the angles and faintly margined,
the anterior edge without any prominent tubercles; dorsally depressed from base to apex,
the surface closely rugose and sparsely hairy, and having a slight central longitudinal
ridge posteriorly. Scutellum small, black and shiny, Anterior coxe large, very hairy
and placed very nearly but not quite touching each other; the femora large and flattened;
the tibize broadened apically, abruptly truncate, with a few strong spines and some hairs
on the outer lateral margin. Elytra reddish-brown, half as long again as the prothorax,
parallel-sided for the basal three-fourths and thence rounded to the apex, the declivity
commences abruptly at the apical fourth and is slightly depressed on each side of the
suture; the elytra are strongly punctate, the punctures appearing slightly larger at
the base than at the declivity, each being provided with a very minute hair, whilst the
interstices are flat and furnished with a single row of longish hairs. The metasternum
is much raised along its centre and deeply grooved longitudinally at the basal portion, the
first visible abdominal segment has a bisinuate basal edge and is somewhat raised
along the centre and rather broader than the second, the third and fourth narrow, the
fifth of equal breadth to the preceding two together; all the segments are coarsely
punctured and hairy.
This large species resembles C. graniceps Kichh. in many ways, but has a much more
abrupt declivity, alternate rows of long and short hairs on the elytra, no carina above the
mouth, the prothorax longer than broad, ete.
Loc. Seychelles. Mahé: slopes of Morne Seychellois, 1500—2000 feet, 4. IT. 1909.
2; specimens.
25. Coccotrypes declivis, sp. n.
Breviter ovalis, subdepressus, piceo-brunneus, pilosus, subnitidus, thorace antrorsum
angustato, anterius granulis exasperato, posterius granulato-punctato; elytris lineato-
punctatis, alternatim longius et brevius seriato-pilosis, apice subtiliter subimpresso-retuso.
Long. 2 mm.
Head black, front closely rugulose strigose with a transverse row of yellow hairs
over the mouth ; prothorax ovate and slightly longer than broad, laterally curved from
base to apex, with a transverse dorsal elevation just behind the centre, the whole surface
slightly roughened and having scattered granulations anterior to the dorsal elevation ;
the anterior edge is furnished with numerous blunt tubercles; the scutellum small and
shiny. The elytra are the same breadth as the prothorax and nearly one-half longer,
and posteriorly the surface on each side of the suture is depressed, whilst laterally they
SAMPSON—COLEOPTERA; PLATYPODIDA AND IPIDAi 391
are parallel to beyond the middle and then rounded to the apex; the apical declivity
commences beyond the middle and is obtuse and slightly depressed centrally.
Near C. graniceps but smaller, and has no frontal carina, the thorax is longer than
broad and not equally granulate over the whole surface, the elytral hairs are similarly
arranged to those of C. bicolor, &e. A slight depression extends the whole length of the
elytra, causing the suture to appear raised when viewed in certain lights; in some of
the specimens no such depression is discernible from the commencement of the declivity to
near the apex. Ten specimens.
Loc. Seychelles. Silhouette: Mare aux Cochons, [X. 1908. Mahé: high forest of
Morne Blane and Pilot, X.—XI. 1908; country above Port Glaud, 500—1000 feet, XI.
1908; from between leaf-bases of growing Stevensonia-palm, near Morne Blane, one
specimen ; forest above Cascade Estate, 1000—2000 feet, II. 1909; Mare aux Cochons
district, 1000—2000 feet, I.—II. 1909.
26. Coccotrypes parvus, sp. n.
Breviter ovalis, piceo-brunneus, pilosus, subnitidus, thorace globoso, anterius granulis
exasperato, posterius granulato-punctato ; elytris lineato-punctatis, alternatim longius et
brevius seriatim pilosis, apice obliquo, utrinque subtiliter subimpresso-retuso.
Long. 1°8—2 mm.
Head pitchy-brown, with roughened surface and a few hairs placed transversely over
the mouth ; the thorax equal in length and breadth with the sides rounded from base to
apex, the anterior edge having several coarse tubercles, of which the middle six are more
prominent than the rest, the whole surface roughened but less so posteriorly, with a few
scattered tubercles above the anterior edge and having some longish upstanding hairs,
especially laterally. The elytra are half as long again as the thorax, but not quite equal
to it in breadth at its widest part ; the interstices flat with rather widely placed piliferous
punctures ; laterally the elytra are subparallel for the posterior two-thirds and rounded to
the apex with a very slight depression of the apical third.
Loc. Seychelles. Mahé: high forest of Morne Blanc and Pilot, XI. 1908, one
specimen. Praslin: Cotes d’Or Estate, Coco-de-mer forest, XI. 1908, one specimen.
27. Coccotrypes, sp.?
There are two specimens, without antennze and otherwise in bad condition, of a
species which probably belongs to this genus. They are characterised by the base of the
declivity bearing a transverse stria of brilliant golden hairs; and are very distinct from
the other species found in the Seychelles.
Loc. Seychelles. Silhouette: Mare aux Cochons, and high forest above Mare aux
Cochons, IX. 1908.
No. XIX.—COLEOPTERA, CURCULIONID&.
By GrorGE CHARLES CHaAmpton, A.L.S., F.Z.S.
(Plates 22—24.)
Read 7th May, 1914.
THE Curculionids here enumerated from the Seychelles, Aldabra, Chagos, and the
adjacent islands number 134 species, 27 only of which can at present be identified with
described forms. The presence of one entirely new group, the Phcenicobatina, including
three genera and 28 species, shows that these islands have a comparatively rich weevil-
fauna, now brought to light by Mr Hugh Scott's exhaustive methods of collecting in the
mountain forests. Probably he is almost the first entomologist who has set to work to
examine properly the leaf-bases of palms (more especially of the Coco-de-Mer) and screw-
pines in these or any other tropical regions*. Hence, a very interesting series of minute
forms has been discovered, and, as might be anticipated, they are nearly all new. The
Cossonina, as usual in various Oceanic Islands (even in cooler regions like St Helena and
New Zealand), are abundantly represented, and include 50 species, with ten new genera.
The Trypetina and Antliarrhina (like most of the Phcenicobatina) are probably attached
to palms, and doubtless endemic. Amongst the Cryptorrhynchina there is a singular
dearth of apterous forms, one species only (an Acalles) being represented. The con-
spicuous winged Otiorrhynchids, Cratopus spp., are, on the other hand, abundant, im
individuals, both on the coast and in the mountains, though but few species are present.
The apterous Hylobiina and Anchonina, insects living amongst dead leaves on the ground,
are also characteristic weevils in the Seychelles, several species being common in the
mountain forests. The three apterous Otiorrhynchids belong to as many genera, one of
which seems to occur also in Madagascar. The Attelabine, Barina, Cyladina, and
Calandrinze are either introduced forms, or belong to genera occurring elsewhere, in
Ceylon, Madagascar, 8. Africa, &e. Out of a total of 134 species, at least 100 are probably
endemic, and the entire group Pheenicobatina, so far as at present known, 1s peculiar
to the Seychelles. Kolbe’s list (1910) includes 17 species only for the same groups of the
Rhynchophora.
The Hawaiian Islands possess almost the same number of Curculionidee [vide
Perkins, Introd. Fauna Hawaiiensis, p. xxii. (1913) }-132 species, in 12 genera, endemic,
and 14 species, in 9 genera, introduced.
* See the general account of the collecting in Vol. xiv. of these Transactions, 1910, pp. 24, 25.
394 PERCY SLADEN TRUST EXPEDITION
The insects enumerated in the following pages were all captured by Mr Hugh Scott,
excepting those obtained in 1905, which were collected by the H.M.S. “Sealark”
Expedition, and some obtained by other collectors, whose names are given: he is also
responsible for the notes on habits and data of capture in this paper.
Types.
A first set, including the types of all new forms, will be placed in the
British Museum, and a second set retained in the Cambridge University Museum.
Inst of Species.
Sub-fam. Attelabine.
1. Huops viriditinctus, n. sp.
Sub-fam. Otiorrhynehine.
2. Lepydnus (n. gen.) nepenthacola, un. sp.
3. Brachycyrtus minor, n. sp.
4. Epirrhamphus (n. gen.) litoralis, n. sp.
5. Cratopus adspersus, C. O. Wat.
6. 3 griseovestitus, Linell.
, A viridisparsus, Fairm.
8. 5 aurostriatus, Fairm.
§). 5 segregatus, 0. Sp.
FP var. subcinctus, n.
10. 5) muticus, N. sp.
Sub-fam. Curculionine.
Group Hylobiina.
1l. Cycloterodes sechellarum, Kolbe.
12. Hucycloterodes (n. gen.) terreus, n. sp.
13. Gonotrachelus (n. gen.) 5-carinatus, n. sp.
Group Anchonina.
14. Tanyomus (n. gen.) palmicola, n. sp.
15. Cycloterinus humeroalatus, Kolbe.
16. * ampliatus, Kolbe.
17. - foveatus, Kolbe.
18. és carinifer, 1. sp.
19. ys canaliculatus, n. sp.
20. nt unicristatus, Nn. Sp.
21. e erosus, D. Sp.
22. ni spheropterus, 1. sp.
23. - microphthalmus, n. sp.
24, B5 ceecus, N. sp.
Group Phoenicobatina, n.
25. Phenicobates (n. gen.) vittatus, n. sp.
26. Fs flexirostris, n. sp.
27. 55 cylindricus, n. sp.
28. 5 parallelus, n. sp.
29. 59 nigrolumbatus, n. sp.
30. A albosetosus, 0. sp.
3l. i fortorostris, n. sp.
32.
33.
34,
35.
36.
37.
38.
39.
40.
4],
42.
43.
44.
45.
46.
47.
48.
49.
50.
ol.
52.
Phenicobates duplovestitus, n. sp.
o praslinensis, n. sp.
depressirostris, n. Sp.
fovewentris, n. sp.
y curvupes, . Sp.
simplex, N. Sp.
tenuis, n. sp.
‘s peropacus, n. sp.
gibbirostris, n. sp.
o stricticollis, n. sp.
x stevensoniw, 1. sp.
+s hispidulus, n. sp.
cuspidatus, n. sp.
5 lodoicece, nu. sp.
5 pandanicola, n. sp.
4 alatus, 0. sp.
5 rufitarsis, D. sp.
23 sebertensis, n. sp.
Phenicobatopsis (n. gen.) echinatus, n. sp.
9 setuceps, N. Sp.
Phenicogenus (nu. gen.) mermis, 2. sp.
Group Anthonomina.
Endeopsis (n. gen.) curvimanus, n. sp.
33 anthonomoides, nu. sp.
An delicatus, n. sp.
Rhynchenus spissus, n. sp.
Group Cryptorrhynchina.
Acalles seychellensis, n. sp.
58. *Camptorrhinus brullei, Boh.
59.
60.
61.
62.
63.
64.
65.
66.
67
Cryptorrhynchus mangifere, F.
A erratus, D. Sp.
Cryptorrhynchidius (n. gen.) graniger, n. sp.
Rhetogenes (n. gen.) sexcristatus, n. sp.
5 spurcus, 2. Sp.
Baridiomorpha (n. gen.) triplaris, n. sp.
Rhombosoma (n. gen.) acuminatum, n. sp.
Scytha (un. gen.) aldabrana, un. sp.
Group Zygopina.
. *Sphadasmus granocostatus, Kairm.
The species marked with an asterisk are not represented in the collections made by the expedition.
CHAMPION—COLEOPTERA ; CURCULIONIDA 395
Group Barina. 99. Stenotrupis polita, n. sp.
68. Baris multivaga, un. sp. 100. ” sericata, nS Sp:
69. 5, seychellensis, n. sp. 101. ” nitidula, n. sp.
; 102. ce silvicola, n. sp.
Group Cyladina. 103. = dumetorum, n. sp.
70. Cylas formicarius, F. 104. Microtrupis (n. gen.) longipennis, n. sp.
Group Antliarrhina. 105. 5 puncticeps, n. sp.
aL ae ; ee 106. 3 piligera, n. sp.
. Microplatymerus (n. gen.) lodoiceivorus, n. sp. 10%, Gastrmaypis (a, gpa) lcdgene, wm, a.
Group Trypetina. 108. Iicrohimatium (n. gen.) pubescens, n. sp.
72. Homalorrhynchus (n. gen.) serripes, n. sp. 109. Himatinum confluum, n. sp.
73. 3 rubricatus, ND. sp. 110. ” PUGGAVERMG 15 IYO.
7A. i verschaffeltie, n. sp. 111. Fe breviusculum, n. sp.
75. Sphodrias (n. gen.) magdaloides, n. sp. 112. Melarrhinus incrustatus, n. sp.
76. Sycites (n. gen). pilicornis, n. sp. 113. Phleophagosoma conicicolle, n. sp.
: 114. # aldabranum, n. sp.
Group Cossonina. 115. Oxydema fusiforme, Woll.
77. Dryophthorus lymexylon, ¥. 116. Orthotemnus filiformis, n. sp.
78. » ecarinatus, 0. Sp. 117. Cossonus incivilis, Fahr.
79. Cherorrhinodes (n. gen.) tenwiculus, n. sp. 118. re suturalis, Boh.
80. Pentarthrum longicolle, n. sp. 119. Rhyncolosoma (n. gen.) dubiwm, Gahan.
81. » punctatissimum, n. sp. 120. Tetragonorrhamphus (n. gen.) tuberculirostris,
82. FS seychellarwm, n. sp. n. sp. :
83. BS scotti, n. sp. 121. Coptus latiusculus, n. sp.
84. TLemnorrhamphus (n. gen.) latirostris, n. sp. 122. Proéces compressicollis, n. sp.
85. Hypopentarthrum (n. gen.) microcephalum, n. sp. 133, » silvestris, Kolbe.
86. Stenopentarthrum (n. gen.) pandane, n. sp. 124, Stenomimus orientalis, n. sp.
87. Trapezirrhynchus (un. gen.) silhouettensis, n. sp. 125, Stenoscelis crassifrons, Woll.
88. Stenotrupis lodorcercola, n. sp. 126. Dendroctonomorphus mwriceus, n. sp.
89. 55 rufipes (Fairm.).
90. 5 convexiuscula, n. sp. Sub-fam. Calandrinz.
Sl. % crassupes, D. Sp. 127. Hugnoristus braueri, Kolbe.
92. ij parallela, n. sp. 128. Trochorrhopalus strangulatus, Gyll.
93. “3 jilum, un. sp. 129. Cosmopolites sordidus, Germ.
94. o biformis, n. sp. 130. Polytus mellerborgi, Boh.
95. Fe tarsalis, D. sp. 131. Calandra oryze, L.
96. 4 caliginosa, n. sp. 132. * linearis, Herbst.
97. a conicicephala, n. sp. 133. ¥ exarata, Boh.
98. - nemoralis, n. sp. 134. 5 stigmaticollis, Gyll.
Sub-fam. Attelabinee.
Kuops.
Euops Schonherr, Gen. Cure. v. p. 318 (1839); Jekel, Ins. Saund., Coleopt., p. 219
(1860); Lacordaire, Gen. Col. vi. p. 550.
The single species of this genus found by Mr Scott in Silhouette is probably endemic.
It belongs to the section Synaptops, Jekel, and comes near £. metnert and EH. bowringi,
Jekel, and EH. (Rhynchites) suffundens, Walk., all of which are from India or Ceylon.
1. Huops viriditinctus, n. sp. (Pl. 22, figs. 1, 1a, ¢.)
Shining, eeneo-piceous, the upper surface often reddish-brown with an zneous lustre,
in mature examples greenish-reneous above and beneath, the legs (a broad space on the
SECOND SERIES—ZOOLOGY, VOL. XVI. . 51
396 PERCY SLADEN TRUST EXPEDITION
femora excepted) and antennze rufo-testaceous or reddish-brown ; almost glabrous above,
the punctures on the pygidium, under surface, and legs each bearing a minute hair-like
scale, the tibize very finely ciliate within, the hairs longer in . Rostrum stout, widened
outwards, finely punctate, very little longer than the enormously developed coalescent
eyes ; head with the basal portion subglobose, moderately exserted, and sparsely minutely |
punctate. Prothorax transverse, very convex, campanulate, bisinuate and sharply mar-
gined at the base; transversely strigose and faintly punctate, deeply sulcate within the
basal margin and shallowly grooved within the anterior margin. Elytra broad, sub-
triangular, transversely depressed on the disc below the base, separately rounded at the
apex ; sharply punctate-striate, the seriate punctures (except at the base) extending
outwards across the broad, subangular interstices, which thus appear strongly transversely
rugose. Legs elongated, the anterior pair especially, in ¢, shorter in ?; tibie finely
serrulate within, the anterior and intermediate pairs bowed, elongate, and strongly
unguiculate in ¢, the anterior pair short and deeply sinuate within in 9.
Length 3—4 mm. (¢9).
Loc. Seychelles: Silhouette.
Fourteen specimens, varying a little in colour. ‘‘ Beaten from bushes or small trees,
on or near the plateau of Mare aux Cochons, at an altitude of about 1000 feet, Sept.
1908.” The smoother head, eneo-piceous colour, &c., separate this species from
EH. suffundens.
Sub-fam. Otiorrhynchine.
Sect. Otiorrhynchine Aptere.
LEPYDNUS, n. gen.
Rostrum broad, short, not longer than the head, truncate at the tip, suleate, the
median groove terminating in a smooth fovea in front, the scrobes lateral, very, large,
deep, angular, extending backwards to above the eyes; antennze long, inserted at the
apex of the rostrum, rather slender, the scape curved, joint 1 of the funiculus obovate,
2—7 small and submoniliform, the club large, ovate; eyes rounded, convex, lateral,
coarsely facetted; mandibular scars small; mentum transverse, covering the maxillee ;
prothorax constricted at the base and towards the apex, truncate in front and behind ;
scutellum wanting; elytra oblongo-cordate, broadly mflexed at the sides beneath, the
humeri extending forwards; anterior and intermediate coxee narrowly, the posterior coxz
very widely, separated; metasternum very short, the episterna not visible ; abdomen
greatly narrowed posteriorly, segments 1 and 2 equal in length, 3 and 4 short,
3—5 narrow, the intercoxal process broadly truncate in front, the first suture deeply
sinuate ; legs rather slender; femora clavate; anterior and intermediate tibiz sharply,
the posterior tibize feebly, unguiculate, the latter widened at the tip and with the
cavernous articular space narrow and apical; tarsi with joint 3 bilobed, the claws long,
stout, and divergent ; body oval, squamose and setose throughout, apterous.
Type, L. nepenthicola,
CHAMPION—COLEOPTERA ; CURCULIONID& 397
This peculiar little apterous Otiorrhynchid may be known by its short, sulcate,
apically foveate rostrum, with large, deep, lateral scrobes, the apically inserted antenne,
the convex eyes, the subcordate, basally emarginate elytra, the equal length of the first
two ventral segments, the powerful tarsal claws, and the strongly setose body, antennee,
and legs. The insect appears to inhabit the forests on the highest peaks of Silhouette and
Mahé. Some of the specimens captured by Mr Scott were found in the pitchers of
Nepenthes. Lepydnus belongs to Lacordaire’s “‘ Otiorhynchides vrais.”
2. Lepydnus nepenthicola, n. sp. (Pl. 22, figs. 2, 2a.)
Moderately elongate, somewhat flattened above, piceous or reddish-brown ; somewhat
thickly clothed with minute cupreous or golden scales, which are in great part hidden by
a dark brown exudation, and also, the antennz and legs included, set with blunt, dark
setee, the sete often long and erect on the elytra; the surface closely punctate. Rostrum
shallowly trisuleate, the median groove terminating in a deep fovea. Prothorax nearly as
long as broad, abruptly constricted and much narrowed anteriorly, and also constricted
just before the base, the disc somewhat uneven. Elytra much wider than the prothorax,
the humeri rounded and anteriorly produced; with rows of closely placed coarse
punctures, the interstices densely punctate and feebly convex. Beneath with coarse
scattered punctures intermixed with the fine punctuation ; metasternum slightly hollowed
down the middle in 2.
Length (includ. rostr.) 375—44, breadth 14—2 mm. (f9).
Loc. Seychelles: Silhouette, Mahé. Silhouette: near Mont Pot-a-eau, ca. 1500 feet,
VIII. 1908, 3 specimens; found floating in pitchers of Nepenthes on highest peaks, over
2000 feet, VIII. 1908, 2 specimens. Mahé: Mare aux Cochons district, 1500—2000 feet,
I—IL. 1909; from forest of stunted “‘Capucin” (Northea) trees on summit of “Montagne
Anse Major,” 2000 feet, II. 1909; Cascade Estate.
Ten specimens, varying in size, in the development of the humeri, and in the length
of the elytral setze. The setze on the tarsi extend on to the claw-joint, and those on the
antennal scape are very conspicuous. The cupreous scales are scarcely visible in dirty
examples. In one of the two Silhouette specimens labelled as found in Nepenthes the
elytral setze are short and wholly decumbent, like those on the prothorax.
BRACHYCYRTUS.
Brachycyrtus Fairmaire, Ann. Soc. Ent. France, 1886, p. 76, pl. 2, fig. 10.
This genus is based upon a single species from Madagascar. An insect found by
Fryer on Aldabra agrees so nearly with Fairmaire’s description and figure of B. seto-
jasciatus* that it can be treated as congeneric for the present.
3. Brachycyrtus minor, n. sp. (Pl. 22, figs. 3, 3a.)
Obovate, rufo-piceous, the antennal club infuscate; densely clothed with brownish-
white scales, the elytra each with an interrupted, oblique, angulate fascia towards the
apex and a small patch at the base fuscous, the prothorax and elytra also set with rather
* This insect is given nine joints to the funiculus in the otherwise excellent figure.
51—2
398 PERCY SLADEN TRUST EXPEDITION
long, curled, decumbent, whitish setze, the legs and antennze closely setulose ; the surface
densely, finely punctate. Rostrum concave and canaliculate, deeply triangularly excised
at the tip, the scrobes narrow, parallel. Antennz stout, joint 1 of the funiculus much
stouter than those following, 3—7 transverse, the club moderately large. Eyes finely
facetted, not prominent. Prothorax strongly transverse, feebly rounded at the sides,
scarcely narrower at the apex than at the base. Elytra ovate, very convex, obliquely
narrowed anteriorly, and at the base not wider than the prothorax ; with regular rows of
conspicuous, rather coarse punctures, the interstices somewhat convex. Legs moderately
stout, the tarsi broad.
Length (includ. rostr.) 24, breadth 14 mm.
Loc. Aldabra: Takamaka (Fryer).
One specimen, found in November or December 1908, possibly an introduction from
Madagascar? Much smaller than B. setofasciatus, Fairm. (length 5—6 mm.), the pro-
thorax and elytra more even, the second joint of the funiculus smaller, the legs not so
stout. The prothorax is apparently vittate down the middle, due to abrasion of the
scales.
EPIRRHAMPHUS, n. gen.
Rostrum excessively stout, short, about as long as the head, a little narrowed out-
wards (with the head appearing subconical as seen from above), feebly emarginate at the
apex, Sulcate, the scrobes superior, distant from the margin, deep, as long as the eyes,
subparallel on their inner edge and curved externally ; antennze moderately long, stout,
inserted: near the tip of the rostrum, joints 3—7 of the funiculus transverse, the club
ovate ; eyes finely facetted, moderately large, superior, rounded (oval as seen from above),
not prominent; mentum very small, leaving the maxillee and palpi exposed ; mandibular
scars small; prothorax transverse, truncate at the base and apex, narrow; scutellum
wanting ; elytra oval, broad ; anterior and middle cox very narrowly, the posterior coxee
widely, separated ; metasternum short, the episterna not visible ; ventral segment 2 much
longer than 3 and 4 united, the latter very short; legs stout; femora unarmed ; tibiz
sinuous within, the anterior and intermediate pairs unguiculate, the posterior pair widened
at the tip, ciliate along their apical margin, and with the cavernous articular surface
glabrous ; tarsi with joint 3 bilobed, pubescent beneath, the claws small, connate at the
base; body obovate, densely squamose throughout, apterous.
Type, LE. litoralis.
This genus appears to be not uncommon on the coasts of Aldabra and Farquhar, and
may be expected to occur further south in Madagascar. The type is not unlike a
Strophosomus or Cneorrhinus. The superiorly placed scrobes and eyes (the head and
rostrum together appearing subconical as seen from above, and the outer portion of the
former distinctly extending beyond the eyes) separate Hpirrhamphus from Ellimenistes
and other described African forms. It may be placed near Hadrorhinus, Schénh.
4. Epirrhamphus litoralis, n. sp. (Pl. 22, figs. 4, 4a.)
Piceous, densely clothed with small cinereous or brownish scales, the prothorax
CHAMPION—COLEOPTERA ; CURCULIONIDA 399
and elytra often darker on the disc, the prothorax sometimes fusco-trivittate and the
elytra with a very large, common, laterally angulate space mottled with the same colour,
the outer limits of this space being traceable or indicated by a few streaks in several
of the light-coloured individuals ; the surface also thickly set with short, curled, adpressed
sete, which are very conspicuous on the legs and antennze; above and beneath very
densely, finely punctate. Head and rostrum hollowed and sharply grooved down the
middle, the inter-ocular space thus appearing raised on each side, the outer margins of
the former narrowly extended beyond the eyes when the insect is viewed from above.
Prothorax rounded at the sides, a little narrower at the apex than at the base. Elytra
convex, with nine regular rows of conspicuous, moderately coarse punctures, the interstices
broad and almost flat. Intermediate tibize in 3 strongly, in 2 more feebly, unguiculate.
Length (includ. rostr.) 35—5, breadth 13—2? mm.
Loc. Aldabra: 1907 (Thomasset); XII. 1908 (Fryer). Cosmoledo: 1907 (Thomasset).
Farquhar Atoll: 1905 (Gardiner).
Numerous examples, varying in the colour of the vestiture, the dark patch on the
elytra and the prothoracic vittze sometimes wholly wanting and the scales uniformly
cinereous. A single specimen only was found on Cosmoledo. The individuals with
strongly unguiculate middle tibize are assumed to be males.
Sect. Otiorrhynchine Alatz.
CRATOPUS.
Cratopus Schénherr, Cure. Disp. Meth., p. 120 (1826); Gen. Cure., 11. p. 46.
A large number of species of this genus have been described, mostly from Mauritius
(Ile de France), Bourbon (Réunion), the Seychelles and adjacent islands, 8. Africa, and
India. The abundant material obtained by Mr Hugh Scott, Mr Gardiner, and other
collectors, in the islands under investigation, shows that there are two common variable
types present—one peculiar to the coasts, the other confined to the mountain forests,—
and, in addition, a rare, peculiar endemic form in the mountains of Mahé. The former have
been described under various names, and it is perhaps convenient to give some of them
specific rank, if localized on particular islands. It may be stated that the genitalia of each
of them have been examined and no structural differences found, even between such
dissimilar species as C. adspersus and C. muticus. Several of these local forms are
represented in the British Museum by specimens received in 1870, as from “ Round
Island,” which is assumed in their register to be the island of that name a little to the
north of Mauritius, whereas the Round Island of the Seychelles is probably the correct
locality for these insects*.
In addition to the species here enumerated, Fairmaire (Bull. Soc. Ent. Fr. 1891,
p. xlvi) mentions one other, C. humeralis, Boh., from the Seychelles, probably in error.
* The very fine Tenebrionid, Dysceladus tuberculatus, C. O. Waterh., from “Round I., Mauritius,” has
been found in abundance in 1905 on Frigate I. in the Seychelles.
400 PERCY SLADEN TRUST EXPEDITION
The forms here dealt with may be distinguished thus :
a. Anterior femora dentate.
1
av. Scutellum squamose.
a. Eyes depressed, oval; surface-vestiture fine, hair-like, at most condensed
into a few small spots on the elytra
adspersus, C. O. Wat. [Amirantes, &c. ].
b°. Hyes moderately prominent, oval ; surface-vestiture coarser, the scales less
elongate, oblong-oval on the elytra, and condensed into numerous
patches on the latter... ... griseovestitus, Linell [Seychelles].
c. Eyes very prominent, rounded, coarsely facetted ; surface-vestiture much as
in C. griseovestitus es che viridisparsus, Fairm. [ Aldabra}.
b. Scutellum bare and shining.
d’. Femora and tibiz rufescent ; the depressions of the upper surface, including
the coarse seriate punctures on the elytra, thickly clothed with metallic
golden-green, golden, or blue scales
aurostriatus, Fairm. [Seychelles].
e". Femora and tibize black ; the depressions of the upper surface very sparsely
squamose, the scales less metallic, at most clustered into a few scattered
patches.
a
Elytra without submarginal line of white scales
segregatus, n. sp. [Silhouette ].
b’. Elytra with a sharply-defined, interrupted, submarginal line of white
scales... ie She segregatus, var. subcinctus n. | Praslin].
b. Anterior femora unarmed ; scutellum bare; surface-vestiture dense, except along
the bare raised elytral interstices Sue sia muticus, n. sp. [Mahé].
5. Cratopus adspersus. (Pl. 22, fic. 5, 2.)
Cratopus adspersus C. O. Waterhouse, Rep. Zool. Coll. “ Alert,” p. 576 (1884).
Loc. Amirantes: Eagle Island (Mus. Brit.) ; Desroches, Poivre and D’Arros Islands,
1905. Chagos Islands, 1905; Egmont, Salomon, and Peros Banhos Atolls. Coetivy,
1905. Seychelles: Bird and Dennis Islands, 1908 (Fryer). Farquhar Atoll, 1905.
Cargados Carajos: Establishment Island, 1905. Aldabra: 1907 (Thomasset), 1908
(Fryer). Astove: 1907 (Thomasset). Cosmoledo: 1907 (Thomasset). Assumption :
1909 (Dupont).
A very long series of this insect has been obtained by Messrs Gardiner, Thomasset,
Dupont, and Fryer in the coral-islands and atolls above quoted. None were obtained
from the coasts of the mountainous granitic islands of the Seychelles group. The type
was from Eagle Island, where it has also been found by Mr Gardiner. C. adspersus
is distinguishable from the other very similar coast-forms by the oval, depressed eyes, the
closer and finer granulation of the prothorax and elytra, and the more elongate and
hair-like scales on the upper surface, this being particularly noticeable on the head
and prothorax. The vestiture, too, is more uniformly distributed, at most condensed into
CHAMPION—COLEOPTERA ; CURCULIONID 2 401
very small scattered spots on the elytra. The scales are often cupreous, golden, greenish,
or bluish-grey, sometimes cinereous, as in the type. The length varies from 6—11, and
the breadth from 24—44 mm.
6. Cratopus griseovestitus. (Pl. 22, fig. 6, 3.)
Cratopus griseovestitus Linell, Proc. U. 8. Nat. Mus., xix. p. 697 (1897); Kolbe,
Mitteil. Zool. Mus. Berlin, v. p. 41 (1910).
Cratopus sparsutus Fairm., Ann. Soc. Ent. Belg., xli. p. 485 (1898).
Loc. Seychelles: Mahé (1905 and 1908—9), Silhouette (VIII. 1908), Praslin (1905) ;
Round Island (Mus. Brit.).
This is exclusively a coast-species, living upon bushes such as Tournefortia argentea
which grow just behind the beach*. It was found at several places in Mahé (Baie
Lazare, and the coast near Cascade), and a number of specimens were obtained in two
islets (Long Island, VII. 1908, and Anonyme Island, I. 1909) lying off the coast of the
main island. . In Silhouette it was met with at various parts of the coast.
The types of both Linell and Fairmaire were from Mahé, whence I have seen
upwards of 60 examples collected by Messrs Gardiner and Scott, as well as 19 from
Silhouette, three from Praslin, and four from Round Island. C. griseovestitus is a form of
C. adspersus with more convex eyes, coarser and more scattered granules on the prothorax
and elytra, and less elongate, broader scales on the upper surface, these being usually
condensed into numerous small definite patches on the elytra. The insect seems to be a
little more elongate than C. adspersus, especially the male, and to have a relatively longer
basal joint to the funiculus. The surface-vestiture is variable in colour, often more or less
metallic. Kolbe also records C. griseovestitus from Silhouette and Praslin.
7. Cratopus viridisparsus.
Cratopus viridisparsus Fairm., Bull. Soc. Ent. France, 1896, p- 222; Linell, Proce.
SS: Nat. Mus., xix. p. 702) (1897).
Loc. Aldabra: Takamaka, Picard Island, &c., 1908 (Fryer); 1907 (Thomasset).
The description of C. viridisparsus would apply to similarly-coloured varieties of
either of the two forms obtained by Messrs Thomasset and Fryer in Aldabra: one with
oval, depressed eyes (C. adspersus), the other (C. viridisparsus, Fairm.) with the eyes
coarsely facetted, more rounded, and strongly convex. M. Lesne, however, who has been
kind enough to compare one of our examples with the latter, assures me that it agrees
with Fairmaire’s type. The eyes in the present species are even more prominent than
they are in C. griseovestitus. The vestiture is equally variable in colour, and condensed
into numerous small patches on the elytra; but it is nearly as fine and hair-like as
in C. adspersus. Of the 93 examples before me from Aldabra, 21 have depressed, and
72 prominent, eyes. Linell says Dr Abbott obtained two examples there.
* There is little doubt that the two species found in the coral-islands (C. adspersus and vuridisparsus)
are obtained in the same way, though I have no personal experience in their cases.—H. Scorv,
402 PERCY SLADEN TRUST EXPEDITION
8. Cratopus aurostriatus.
Cratopus aurostriatus Fairm., Bull. Soc. Ent. France, 1892, p. cli; Kolbe, Mitteil.
Zool. Mus. Berlin, v. p. 41 (1910).
Cratopus parcesquamosus Fairm., Bull. Soc. Ent. France, 18938, p. ecexxiv ; Kolbe,
loc. cit.
Cratopus abbott: Linell, Proc. U. 8. Nat. Mus., xix. p. 704 (1907).
Femora and tibize rufous; the depressions of the upper surface each with a small
dense cluster of rounded, golden-green, blue, or golden scales, the elytra sometimes with
a dense irregular or interrupted submarginal stripe of carneous or whitish scales similar
to that along the sides of the body beneath; the elytra coarsely and regularly striate-
punctate, the interstices feebly transversely plicate.
Loc. Seychelles: Mahé; 1888 (Lister) ; (Brauer) ; 1905; 1906 (Thomasset) ;
1908—9: Cerf Island, 1906 (Meade-Waldo): Praslin, 1905: Félicité, XII. 1908: Round
Island (Mus. Brit.). Glorioso Island (Abbott). Madagascar.
Var. Femora and tibize piceous or rufo-piceous, the femora sometimes sparsely
viridi-squamose between the scattered hairs; the prothorax closely granulate, and the
elytra strongly transversely plicate, the irregular depressions of the surface clothed with
golden-green or blue scales, sometimes with golden or carneous scales intermixed.
Loc. Seychelles: Mahé, 1908—9; Marie Anne Island, 1908; Round Island (Mus.
Brit.).
This species’ was found by Mr Scott to be abundant in a number of localities in the
mountain forests of Mahé, where it was beaten from various dicotyledonous trees. C. awro-
striatus occurred up to the highest parts of the mountains, and has also been taken nearer
sea-level in two islets (Cerf Island and Long Island) lying off the coast of Mahé. It has
also been met with in several more distant islands of the Seychelles, z.e. Praslin, Félicité,
and Marie Anne, but it is noteworthy that no specimen is recorded from Silhouette. It
is considered by some people to be an introduced insect, possibly responsible in part for
the destruction of endemic “Capucin” (Northea) trees. Fairmaire gives simply “Sey-
chelles” for it. His C. parcesquamosus is based on an abraded example of the same
species. The tarsi in fresh specimens are often clothed above between the long white hairs
with narrow blue scales, but no reliance can be placed on this character. The male has
the prothorax more rounded at the sides, and relatively broader, than the female; the
anterior legs more elongated, and the femora still more robust; and the anterior coxe
thickly clothed with long white hairs. The two specimens from Félicité Island have the
scales of the upper surface golden-cupreous and the whitish submarginal stripe well
developed. The variety with strongly plicate elytra and irregularly distributed confluent
patches of scales is represented by five examples only. We are indebted to M. Lesne of
the Paris Museum for the loan of a co-type of each of Fairmaire’s species.
9. Cratopus segregatus, n. sp.
Elongate, subfusiform, shining, black, the antennee piceous or ferruginous ; above
CHAMPION—COLEOPTERA ; CURCULIONID& 403
clothed with minute, widely scattered, whitish or pale green scales, which are sometimes
here and there condensed into a small patch on the depressions, especially towards
the sides, the lower surface with scattered whitish hairs; the elytra with a transverse
space on each side of the scutellum, the intermediate and posterior femora with a patch
towards the apex in front, and an interrupted stripe along the sides of the body beneath
(sometimes reduced to one or two spots), clothed with larger carneous or whitish scales ;
the tarsi with narrow blue scales intermixed with the long white hairs above. Head very
sparsely punctate, narrowly foveate between the eyes, the latter very large, oval, de-
pressed, finely facetted ; rostrum rather narrow, flattened, feebly convex at the apex,
sparsely punctate, obliquely carinate on each side anteriorly; joint 2 of the funiculus
a little longer than 1. Prothorax transverse, constricted and much narrowed in front,
strongly rounded at the sides in ¢, narrower, subconical, and with the sides subparallel at
the base in 2; coarsely, confusedly punctate, the interspaces becoming granulate towards
the sides, the disc obsoletely canaliculate posteriorly. Scutellum small, smooth, bare.
Elytra long, very much wider than the prothorax, acuminate at the apex, granulate at the
sides below the humeri; the outer margin serrulate towards the tip, the apices mucronate,
the humeri obliquely truncate; coarsely striato-punctate, the interstices broad, feebly
transversely plicate, and also sparsely punctate, the punctures each preceded by a minute
granuliform prominence. Beneath closely transversely strigose and sparsely, finely,
granulato-punctate, the flanks of the prothorax coarsely granulate. Anterior coxz
thickly clothed with long white hairs in g. Anterior femora strongly clavate, enormously
thickened in fully-developed ¢, and armed with a small tooth. Anterior tibize sparsely
denticulate.
Length (includ. rostr.) 9—15, breadth 3—53 mm. (#2).
Loc. Seychelles: Silhouette, 1908.
Var. subcinctus, n. (Pl. 22, fig. 7, g.)
The elytra with an interrupted submarginal stripe (similar to that along the sides of
the body beneath), and a small patch at the base of each dorsal interstice, densely clothed
with rounded carneous or white scales, the disc sometimes with a few small patches
of similar scales; the intermediate and posterior femora with a few intermixed narrow
blue scales towards the apex, the patch of larger white scales wanting.
Loc. Seychelles: Praslin; 1905; 1906 (Meade-Waldo); XI—XII. 1908: Round
Island (Mus. Brit.). .
This insect and its variety are forms of C. aurostriatus that require distinctive
names. The one selected as typical of C. segregatus is abundant in Silhouette, in the
same way that C. aurostriatus is in Mahé; it was beaten from bushes and trees in the
mountain-forests, and some specimens also were obtained at lower levels: its variety
subcinctus inhabits similar places in Praslin, where typical C. auwrostriatus also occurs.
C. segregatus simply differs from C. aurostriatus in having black legs, and the vestiture
of the upper surface greatly reduced, even in fresh examples, the minute scales being
clustered, at most, into a few patches in the depressions of the elytral surface and
extending over the interstices.
SECOND SERIES—ZOOLOGY, VOL. XVI. 52
404 PERCY SLADEN TRUST EXPEDITION
10. Cratopus muticus, n. sp. (Pl. 22, fig. 8, 2.)
Elongate, rather narrow, fusiform ; nigro-piceous or piceous, the antennze, femora,
and tibie rufo-piceous or ferruginous ; thickly clothed, except upon the raised portions of
the upper surface, with small, rounded or oval, pale brown, carneous, or whitish scales,
sometimes with pale greenish scales intermixed on the humeri, or along the suture and
outer interstices of the elytra, on the coxee, and along a broad space down the middle of
the under surface; the legs and a space down the middle beneath sparsely clothed with
long white hairs, the tarsi with intermixed narrow bluish or green scales above, and the
femora also with a few oval greenish scales. Head very sparsely punctate, narrowly
foveate between the eyes, the latter very large, oval, moderately prominent, and finely
facetted ; rostrum a little longer than broad, rather narrow, without marginal carina.
Prothorax transverse, rounded at the sides, narrowed and constricted anteriorly ; con-
fusedly punctate and coarsely granulate, the vestiture becoming longer, denser, and
transversely arranged along the sides. Scutellum small, shiming, bare. Elytra very much
wider than the prothorax, elongate, widened posteriorly in $, compressed and strongly
acuminate at the apex, granulate at the sides below the humeri, the latter obliquely
truncate, the outer margin serrulate towards the tip, the apices mucronate; with rows of
coarse punctures, separated by raised, subgranulate or interrupted, glabrous interstices,
the intervening depressed spaces being densely squamose. Beneath closely transversely
strigose and sparsely granulato-punctate. Legs very elongate, rather slender; anterior
femora unarmed ; anterior and intermediate tibize very strongly, and the posterior tibiz
feebly, unguiculate in $, the uncus on the middle pair short, and that on the posterior
pair wanting, in §.
Length (includ. rostr.) 10—114, breadth 3—33 mm. (¢9).
Loc. Seychelles : Mahé.
Ten specimens were obtained ; one by Gardiner in 1905; one by Thomasset in 1906 ;
and eight by Scott in 1908—9, at the following localities: near Morne Blane, ca.
1000 feet, XI. 1908; Cascade Estate, II. 1909; forest on summit of Montagne Anse
Major, ca. 2000 feet, Il. 1909; Mare aux Cochons district, over 1000 feet, I. 1909. It
was found but rarely, by beating, and it appeared to be usually, if not always, beaten from
”
the endemic “‘Capucin” tree (Northea seychellarum).
C. muticus differs from all the species described by Boheman by the unarmed anterior
femora. It is a rather slender, elongate form, with densely squamose body, the elytral
interstices being raised and bare.
Group Hylobiina.
Sub-fam. Curculionine.
CYCLOTERODES.
Cycloterodes Kolbe, Mitteil. Zool. Mus. Berl., v. p. 44 (1910).
The following additions to Kolbe’s diagnosis of this genus are required: ‘“‘ Anterior
coxee subcontiguous; prosternum very deeply emarginate at the apex, and broadly
CHAMPION—COLEOPTERA ; CURCULIONID Ai 405
depressed down the middle; eyes large.” The excavate prosternum of Cycloterodes is
suggestive of the Cryptorrhynchina, but the rostrum is free and does not even rest upon
the subcontiguous anterior coxee in repose. There is an insect from Ceylon in the British
Museum, standing under the MS. name Cycloteres rugulosus, Dohrn, that belongs to the
same genus. The type is a large apterous insect, with the surface densely coated with
minute earthy-looking scales, and also set with short erect setze.
11. Cycloterodes sechellarum. (Pl. 22, figs. 9, 9a, 3.)
Cycloterodes sechellarum Kolbe, Mitteil. Zool. Mus. Berl., v. p. 44 (1910).
Loc. Seychelles: Silhouette, Mahe.
Described from a single example from Silhouette. Mr Scott found a long series of it
in Silhouette, and one in Mahé. In the former island it was found in numbers at the
Mare aux Cochons plateau, at an elevation of 1000 feet, at night, crawling on the palm-
lath hut-walls, or on the trunks of some Jack-fruit trees near by, often in wet weather.
Kolbe’s description must have been made from a dirty specimen, as he does not mention
the numerous tuberculiform prominences scattered along the elytral interstices, some of
which are clothed with black scales, two at the base of each elytron (like the two on the
anterior portion of the prothorax) being conspicuous; the elytra, too, have each two or
three dirty-white transversely-placed spots at about one-third from the tip. The rostrum
of the female is bare at the apex, and less convex and more feebly carinate above than in
the male. The first ventral segment of the male is very broadly, feebly depressed.
KUCYCLOTERES, n. gen.
Rostrum strongly deflexed, resting on the anterior coxae in repose, short, very stout,
slightly widened outwards, the scrobes very deep, superiorly placed, extending forwards to
the apex above and running obliquely downwards to beneath the eyes posteriorly, the
antennee inserted near the tip; mandibles dentate within, without scar, visible beyond the
tip of the rostrum when closed; mentum very small, transverse; eyes oval, transverse,
moderately large, separated by the width of the rostrum, incompletely hidden in repose ;
antennz with 7-jointed funiculus, joints 1 and 2 elongate, the others short and moniliform,
the club ovate, the scape reaching the eyes; prothorax with its broadly rounded base
accurately fitted to that of the elytra; scutellum wanting; prosternum deeply arcuate-
emarginate in front, unimpressed, the anterior portion extremely short; anterior coxz
subcontiguous, the intermediate coxee moderately, the posterior coxze very widely, separated ;
metasternum extremely short, without visible episterna ; ventral segments 1 and 2 connate,
2 less than half the length of 1, 3 and 4 extremely short, the first suture sinuate, the
others straight; legs robust; femora dentate; tibiee angularly dilated at the middle
within, unguiculate at the inner apical angle, and with a short tooth on the inner side
near the apex; tarsi stout, spongy-pubescent beneath, joint 1 triangular, 2 transverse,
3 broadly bilobed, narrowly emarginate for the reception of the claw-joint, the claws free;
body robust, subrotundate, Erodiiform, densely coated with minute earthy-looking scales,
apterous.
52—2
406 PERCY SLADEN TRUST EXPEDITION
Type, #. terreus.
The insect forming the type of this genus has the general appearance of such
aberrant Cryptorrhynchids as Hybomorphus and Ocladius, but it is in reality related to
Cycloteres. The regularly rotundate shape is suggestive of the Tenebrionid genus Hrodius.
The anterior portion of the prosternum is extremely short, thus allowing the rostrum to
rest on the anterior coxze in repose. Achynius, Fairm., from Madagascar, may be an
allied form.
12. Hucycloteres terreus, n. sp. (Pl. 22, figs. 10, 10 a.)
Very broad, rotundate-ovate, opaque, black, the antennze and tarsi rufo-piceous ; the
entire surface covered with minute brown or reddish-brown scales and also minutely
setulose, the legs and antennze with more conspicuous sete. Rostrum bare at the tip, .
when denuded of scales shining, closely, finely punctate, and 4-carinate from the base
to near the apex. Head, prothorax, and elytra densely, extremely minutely punctulate,
the prothorax with intermixed scattered moderately coarse punctures. The elytra with
regular rows of foveze placed in shallow striz, the interstices broad and very sparsely,
obsoletely granulate. Beneath coarsely punctate.
Length (exclud, head) 5—63?, breadth 33—51 mm.
Loc. Seychelles: Silhouette, Mahé, Praslin.
Six examples, sex not ascertained. In Silhouette this species was found in the forest
above Mare aux Cochons, IX. 1908: in Praslin, in the ‘‘ Coco-de-mer” forest in the Vallée
de Mai, Cotes d’Or Estate, XI. 1908: in Mahé, in the Mare aux Cochons district, ca. 1500
feet, I.— II. 1909.
GONOTRACHELUS, n. gen.
Rostrum very stout, curved, as long as the prothorax, the apical portion slightly
widened and parallel-sided, the antennze inserted beyond the middle, the scrobes deep and
obliquely descending to the lower surface ; head globose, constricted behind the eyes, the
latter small, superior, coarsely facetted, and placed at the base of the rostrum; antennze
(fig. 11) with a short 5-jointed funiculus and a small, ovate, pubescent club; prothorax
subquadrate, sharply carinate; scutellum wanting; elytra broad, oval, produced at the
apex; anterior coxee contiguous; posterior coxee very widely separated, the intercoxal
process truncate in front; metasternum short, the episterna covered by the elytra ;
ventral segments 1 and 2 long, connate at the middle, 3 and 4 short, together as long as 5,
the first suture sinuate and the others straight; tibie sinuate within, armed with a
slender claw at the outer apical angle; tarsi very short, spongy-pubescent beneath,
joint 3 bilobed, the claws minute, free; body oblong, convex, subglabrous, apterous.
Type, G. quinquecarimatus.
The insect from Long Island, Mahé, from which the above characters are taken may
be described as a Gononotus (Lec.) with the antennal structure, 7.e. a 5-jointed funiculus,
of Dryotribus Horn ( = Thalattodora, Perkins). Both these genera have been referred to
Cossonina, but Gonotrachelus (and Gononotus also) seems to me to be better placed
CHAMPION—COLEOPTERA ; CURCULIONIDA 407
amongst the Hylobiina, near Styphloderes. All these insects occur on the sea-shore, and
appear to be easily introduced in drift-wood, &c. Gononotus has been recorded from
Florida and Cuba, and Dryotribus from Florida, the Antilles, the Hawaiian Is., Adéle
Island (N.W. Australia), and Nyew-tew Island, China. Gonotrachelus, therefore, may
have been introduced into the Seychelles, but from whence it is impossible to say at
present.
13. Gonotrachelus quinquecarinatus, n. sp. (PI. 22, figs. 11, 11a, 6.)
Oblong-ovate, opaque, piceous, the antennze ferruginous. Head and rostrum rugosely
punctate; antennz short, the joints of the funiculus comparatively stout, 2—5 transverse.
Prothorax subquadrate, about as broad as long, abruptly constricted anteriorly ; very
coarsely, confluently punctate, and sharply 5-carinate, the two outer carinze on each side
sinuous, becoming divergent anteriorly, the outer one marginal and forming a blunt tooth
at its point of termination. Elytra much wider than the prothorax, oval, truncate at the
‘base, the humeri obtuse; coarsely punctate-striate, the interstices narrow and more
or less costate, 3, 5, and 7 sharply raised. Beneath coarsely, closely punctate, the
imterspaces alutaceous.
Length 2—2}4 mm.
Loc. Seychelles: Long Island, near Mahé, VII. 1908.
Two specimens, probably male and female.
Group Anchonina.
Anchonina Champion, Biol. Centr.-Am., Coleopt., iv. 4, p. 66 (1902).
Faust (Deutsche ent. Zeit. 1892, p. 19) referred numerous American genera to
this group and others from the same continent were subsequently added by myself.
Cycloterinus, Kolbe, has the head, &c., formed exactly as in these American forms—
Kolbe, in fact, notes its resemblance to Oncorrhinus,—and the genus is much better
placed in Anchonina than in Hylobiina. The group is represented in the Seychelles
by two genera. All the species are apterous, and in one, Cycloterinus cecus, the eyes
are wanting. C’. foveatus is not unlike the American genus Theognete, which has the eyes
placed on the sides of the rostrum, instead of on the smooth, globose, retractile head.
Mr Scott notes that he obtained all his material, with the exception of one species
(ce; carinifer), from the forest-clad mountains, from amongst dead leaves on the
ground. The largest form, Tanyomus palnucola, was found exclusively on the concave
inner surfaces of the bases of damp decaying palm-leaves. The larger forms of Cyclo-
terinus (such as C. foveatus and C. humeroalatus) were also often found in similar
situations on dead palm-leaves, but some of them may also have been on leaves of
dicotyledonous trees. The smaller Cycloterinus (e.g. C. wnicristatus) were found on
leaves of the “Bois Rouge” (Wormia ferruginea) and of the ‘Capucin” (Northea
Sechellarum), both endemic trees. The one exception mentioned above (C. carinifer)
was not found on the ground at all, but in decaying outer leaf-bases of a growing
Lodoicea-palm.
408 PERCY SLADEN TRUST EXPEDITION
The small forms of Cycloterinus are very difficult to see on the dead Worma
and Northea leaves, owing to their flattened form and cryptic coloration. They are
frequently found sitting on the sides of the raised midrib and other veins of the
“Bois Rouge” (Wormia) leaves. They did not appear to be abundant in Silhouette
in the season of the S.E. Trade-winds (July—September 1908). Several species were
found in that island, but only in small numbers: [a specimen of one of them
(C. ampliatus) was bred from a larva found in the rotting substance of the base of
a fallen palm-leaf|, These insects were abundant in the forests of Mahé in November
and December 1908, and it was then that the great series of some species (eg.
C. unicristatus) were principally accumulated. Examples were frequently found in coitu
on the dead leaves. They were much scarcer again in the forests behind Cascade
in January and February 1909. It was several times observed that there were, in
the sides of the midribs of the leaves, little holes each surrounded by a tiny pile of
leaf-substance reduced to dust; and it is possible that these were the work of the
small forms of Cycloterinus, which are probably connected with the dead leaves
throughout their life-cycle. (Cf. Scott, “Eight Months Entomological Collecting m
the Seychelles Islands,” Vol. xiv. of these Transactions, 1910, pp. 29—30.)
TANYOMUS, n. gen.
Head globose, polished, the eyes small, depressed, transverse, placed low down imme-
diately behind the base of the rostrum; rostrum arcuate, long in 3, shorter in ?, constricted
at the base, the antenns inserted near the tip, the scrobes inferior, deep, extending
to beneath the eyes; funiculus 7-jointed, joint 1 elongate, the others decreasing
in length; prothorax subglobose, constricted near apex; scutellum wanting; elytra
subscutiform, the humeri acutely produced anteriorly ; prosternum bituberculate between
the rather narrowly separated anterior coxe, the anterior portion long; metasternum
very short, without visible episterna; ventral segments 1 and 2 long, connate, 3 and 4
together longer than 5, the sutures straight; femora unarmed; tibiz straight on
their outer edge, each with a curved hook at the apex arising from near the outer
angle, the anterior pair broadly produced inwards at the apex and with a short tooth
at the inner angle in both sexes; tarsi clothed with long projecting hairs, spongy-
pubescent beneath, joint 3 with two narrow divergent lobes of unequal length, the
claws long and divergent ; body apterous, setose, elongate, subfusiform.
Type, 7. palmicola.
The remarkable insect forming the type of this genus is not unlike a large black
Erirrhinus. The anteriorly produced spiniform humeri make the elytra appear conjointly
arcuate-emarginate at the base. The rostrum is longer, stouter, and more rugose
in the male than in the female. The bituberculate prosternum and the asymmetrically
lobed third tarsal jomt are also characteristic, the last-mentioned character, however,
is also to be found in two species of Cycloterinus, to which the present genus is
nearly related.
CHAMPION—COLEOPTERA ; CURCULIONIDA 409
14. Tanyomus palmicola, n. sp. (Pl. 22, figs. 12, 12a, ¢.)
Rather broad, moderately shining, black or pitchy-black, the tarsi and antenns
ferruginous or obscure ferruginous; the prothorax, rostrum, and legs setulose, the
elytra with scattered, long, erect, stiff, fulvous sete intermixed towards the sides
and apex with minute, short, adpressed hairs, the antennze sparsely pilose, the pro-
jecting hairs on the tarsi fulvous. Rostrum (f) stout, strongly arcuate, longer than
the prothorax, dull and shallowly punctate above, and with two small tubercles at
the apex beneath, (?) shorter and more slender, shining, closely punctate; the lateral -
portions suleate and carimate in both sexes. Prothorax about as broad as long, the
short anterior lobe narrow; coarsely, closely punctate, the punctures coalescing into
oblique or sinuous rugee towards the sides and base. LElytra much wider than the
prothorax, moderately elongate, transversely convex, depressed at the base, the acutely
produced humeri raised; with rows of rather scattered coarse punctures placed in
almost obsolete strize, the interstices feebly convex and with the setigerous punctures
extremely minute. Beneath shining, coarsely, closely punctate, the punctures becoming
finer towards the apex, the second ventral segment transversely tumid at the base
in both sexes.
Length 54—61, breadth 243—3 mm. ($9).
Loc. Seychelles: Mahé.
Twelve specimens, found in the high forests of Morne Blane and Pilot (XI. 1908),
and between Trois Fréres and Morne Seychellois (1500—2000 feet, I. 1909), on the
concave inner sides of the bases of damp decaying palm-leaves on the ground.
CYCLOTERINUS.
Cycloterinus Kolbe, Mitteil. Zool. Mus Berl., v. p. 41 (1910).
Three species from the Seychelles were referred to Cycloterinus by Kolbe, of which
C. foveatus may be taken as the type. These forms and various others were found by
Mr Scott in the same islands, some of them in great abundance. They vary greatly
mter se in the development of the rostrum in the male (it bemg much longer in this sex
than in the female in two of the species described by Kolbe), the emargination of the
apex of the prosternum, the extent of separation of the anterior coxee, the width of
the third tarsal joint, and the size of the tarsal claws. They agree, however, in
having the head small, globose, and shining; the eyes transverse, oval, depressed,
and placed low down immediately behind the base of the rostrum, showing a gradual
diminution in size till in one species (C. cecus) they are wanting; the tibiz unguiculate
at the apex; the ventral segments 1 and 2 long and connate, 3 and 4 extremely short,
the sutures straight; the metasternum very short, without visible episterna; the
elytra connate, in some of the species with the humeri dentiform or abruptly laminato-
explanate; the wings wanting.
410 PERCY SLADEN TRUST EXPEDITION
a. Third tarsal joint with asymmetrical lobes; rostrum longer in ¢ than in $;
prothorax subquadrate, 4-tuberculate; elytra tuberculate and with
abruptly dilated laminiform humeri; eyes present; body setulose.
a. Prothorax not granulate; elytra not margined laterally, humeri angulate
or subangulate in front .. a ; ee ... humeroalatus.
b*. Prothorax granulate; elytra margined bitsy, humeri rounded
ampliatus.
b. Third tarsal joint with symmetrical lobes, simply excavate above in the smaller
forms.
Gc}
Kyes present. :
a*, Body glabrous; prothorax subquadrate ; elytral humeri not prominent ;
rostrum about equal in length in the two sexes.
3
a’. Prothorax suleate, 4-tuberculate; elytra irregularly foveate; third tarsal
joint bilobed ; eyes rather large ; second ventral segment with a stout
tubercle in g es ee be 43 : ... foveatus.
b*. Prothorax 5-costate ; alternate Seal interstices snseenapeedlly costate ;
eyes small; third tarsal joint feebly bilobed ... .. carmifenr:
c’. Prothorax canaliculate; alternate elytral interstices uninterruptedly
costate; eyes small ... sce ... canaliculatus.
‘
b*. Body setulose; third tarsal jomt small, font lets or excavate.
d*. Prothorax subquadrate; elytral humeri dentiform; eyes small.
4
a’. Hlytra subcordate or subglobose.
Alternate elytral interstices simply raised, the first (sutural) with a
a’.
setigerous tubercle beyond middle in ¢ .. = 6uneristatus,
b’. Alternate elytral interstices interruptedly costate or tuberculate
er Osus.
b*. Elytra convex, subglobose, alternate interstices interruptedly costate
spheropterus.
e
Prothorax more rounded at sides; elytra short-oval, convex, humeri not
prominent: species very small ... ie ... nucrophthalmus.
d'. Eyes wanting; prothorax elongate, with long, cucullate, anterior lobe;
elytra oval, humeri not prominent ... Aes 206 .. CHCUS:
15. Cycloterinus humeroalatus. (Pl. 22, figs. 138, 13a, 3.)
Cycloterinus humeroalatus Kolbe, Mitteil. Zool. Mus. Berl., v. p. 42 (1910).
Loc. Seychelles: Mahé, Silhouette. Mahé: a large number of specimens ; high damp
forest of Morne Blane and Pilot, X.—XI. 1908; high damp forest between Trois Fréres
and Morne Seychellois, 1500—2000 feet, XII. 1908 and I. 1909; forest above Cascade
Estate, I. 1909; Mare aux Cochons district, 1000—2000 feet, I.—II. 1909; forest on
summit of “Montagne Anse Major,” ca. 2000 feet, II. 1909. Silhouette: only two
specimens, from forest near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908.
The types of this species were found in Mahé by Herr A. Brauer, beneath leaves
on the ground im the high forests, at an elevation of 600—750 metres. Mr Scott
CHAMPION—COLEOPTERA ; CURCULIONIDA 411
has found it in abundance, in similar situations, in the same island. Fully developed
males have the elytral tubercles developed into long conical processes at the commence-
ment of the apical declivity (the two on each elytron forming with those on the opposite
wing-case a transverse row), these tubercles in undeveloped specimens of the same
sex being reduced to small oblong prominences, as in the female. The humeral laminiform
prominences of the elytra vary in development in a similar way. The rostrum of the
male is long, carinate, much swollen above beyond the middle, and opaque to near
the tip; that of the female is shorter, simply curved, carinate in its basal half above, and
shining throughout. The first ventral segment of the male is transversely excavate in the
middle, and the second in fully developed examples is tumid in the centre behind.
The upper surface of the body is somewhat thickly clothed with short stiff erect sete.
The length varies from 3—4, and the breadth (at shoulders) from 13—22 mm.
16. Cycloterinus ampliatus. (Pl. 22, figs. 14, 14a, 3.)
Cycloterinus ampliatus Kolbe, Mitteil. Zool. Mus. Berl., v. p. 43 (1910).
Loc. Seychelles: Silhouette.
Found by Herr A. Brauer in the forests of Silhouette, at an elevation of 400—500
metres. Mr Scott met with five examples of it in the same island, amongst dead leaves
in the high jungle at about 1500 feet, and one was bred by him from a larva found
in damp decaying leaf-bases of a palm on the ground; these are all males. This
insect differs from feebly tuberculate examples of C. humeroalatus in having the rostrum
of the male much smoother and without median carina, and tumid above and beneath ;
the prothorax more rounded at the sides, more closely and less coarsely punctate,
with the interspaces more or less granulate, and the four transversely-placed tubercles
less prominent, the two inner ones arising further backward ; the elytra sharply margined
laterally, and with the laminiform humeral prominences rounded in front ; and the legs
reddish. The second ventral segment of the male is simple. The upper surface is
setulose as in C. humeroalatus.
ie Chiclotontimes youeotus. (Pl. 22, figs. 15, 15a, 6, g:)
Cycloterinus foveatus Kolbe, Mitteil. Zool. Mus. Berl., v. p. 42 (1910).
Loc. Seychelles: Mahé, Praslin. Mahé; high forest of Morne Blanc, X. 1908;
forest above Cascade Estate up to 1700 feet; Mare aux Cochons district, 1000—2000 feet,
L—Il. 1909.
The types of this insect were found in Mahé, on damp forest-clad slopes, at an
elevation of 400—750 metres, and Herr A. Brauer is also stated to have taken it
beneath leaves on the ground on the high land in Praslin. Mr Scott secured a
very long series of it, especially in the forest on Cascade Estate in Mahé, but he did
not meet with it in Praslin. C. foveatus is readily distinguishable by its glabrous
body, the very coarsely irregularly foveate elytra, and the prominent, posteriorly-directed,
laminiform tubercle (fig. 15) on the second ventral segment of the male. The rostrum
of the male is much swollen above at about one-third from the apex, and carinate down
SECOND SERIES—ZOOLOGY, VOL. XVI. 53
412 PERCY SLADEN TRUST EXPEDITION
the middle and opaque thence to the base; that of the female is simply curved, shining,
closely, finely punctate, and without median ridge. The length varies from 24—4 mm.
18. Cycloterinus carinifer, n. sp. (Pl. 22, figs. 16, 164, 2.)
Oblong-ovate, robust, dull, glabrous; nigro-piceous or black, the under surface
piceous, the rostrum, antennze, and tarsi, and sometimes the tibize and the base of the
femora also, ferrugimous or obscure ferruginous. Rostrum very stout in both sexes;
(¢) with the lower portion dilated and shovel-shaped from about the basal third, and
the upper portion opaque, carinate, and rugosely punctate to near the tip; (?) simply
arcuate, coarsely punctate, and more shining towards the apex. Eyes very small.
Prothorax about as long as broad, subquadrate, the posterior portion transverse, feebly
rounded at the sides and more or less angulate at each anterior angle, the tubulate
anterior portion abruptly separated from the rest; coarsely, confluently punctate, with
a sharp carina running down the middle and a short stout ridge midway between it
and a similar ridge above the anterior angle, the interspaces between these elevations
sulciform. Elytra considerably wider than the prothorax, somewhat scutiform, the humeri
obtuse ; with rows of oblong deep foveze, the interstices alutaceous, 3—6 strongly, the others
more feebly, costate, the ridge on 3 interrupted at some distance before the apex, and that
on 5 abruptly terminating at the commencement of the apical declivity. Beneath very
coarsely, closely punctate. Third tarsal joint moderately dilated, excavate above.
Length 24—3 mm. (9).
Loc. Seychelles: Praslin.
Two females and one male, found in the decaying outer leaf-bases of a growing
“ Coco-de-mer ” (Lodoicea sechellarum) in the Vallée de Mai, Nov. 28th, 1908. The
carimate, quadrisulcate prothorax and the costate interstices of the elytra readily dis-
tinguish C. carinifer. The rostrum of the male is similar to that of CL unicristatus.
The prosternum is arcuate-emarginate at the apex, and all the coxe are widely separated.
19. Cycloterinus canaliculatus, n. sp. (Pl. 28, figs. 17, 17a, 2.)
Oblong-ovate, convex, opaque; glabrous; nigro-piceous or black, the under surface,
rostrum, femora, and tibize rufo-piceous or obscure ferruginous, the antennze and _ tarsi
ferruginous. Rostrum in both sexes very stout, simply arcuate, finely punctate, and
feebly striate laterally, opaque in ¢, shining in @. Eyes very small. Prothorax longer
than broad, the posterior portion subquadrate, with the sides rounded and more or
less sinuate before the obtusely subangulate anterior angles, the tubulate anterior
portion rather long and abruptly separated from the rest; coarsely, confluently punctate,
and with a shallow median groove. EHlytra oval, much wider than the prothorax,
convex, abruptly declivous behind; seriato-foveate, the interstices alutaceous, 3 and 5
strongly, and 4 and 6 more feebly, costate. Beneath coarsely punctate; ventral segment 1
broadly excavate down the middle in ¢. Prosternum deeply arcuate-emarginate at
the apex. Anterior coxze moderately separated. Third tarsal joint moderately dilated,
excavate above.
Length 3—3+ mm. (9).
CHAMPION—COLEOPTERA ; CURCULIONIDA 413
Loc. Seychelles: Mahé.
Five specimens from the high damp forest:—Morne Blane and Pilot, XI. 1908;
between Trois Fréres and Morne Seychellois, 1500—2000 feet, XII. 1908; slopes of
Morne Seychellois, 1500—2000 feet, II. 1909. Differs from C. carinifer in having the
rostrum similarly shaped im the two sexes, the prothorax longer and simply canaliculate
on the disc, the elytra more inflated and with the costz uninterrupted posteriorly, the
prosternum more deeply emargimate at the apex, and the anterior and intermediate
coxee much less widely separated.
20. Cycloterinus wueristatus, n. sp. (Pl. 23. figs. 18, 18a, ¢.)
Short, broad, shining; varying in colour from black to ferrugimous, the tarsi and
the base of the antennee always rufo-testaceous; clothed above with short, curled,
semi-erect, fulvous or cinereous setz, which are condensed on the elytra into a conspicuous
line along the rather prominent alternate interstices, the tubercle on each side of the
suture in ¢ with a common tuft of longer erect hairs. Rostrum (g) very stout, arcuate,
nearly as long as the prothorax, the lower portion dilated and shovel-shaped from near
the base, the upper portion opaque, 7-carinate, and rugosely punctate to near the apex ;
(?) arcuate, cylindrical, shining, finely punctate, and carinate at the sides only. Hyes
very small. Prothorax transversely subquadrate, the tubulate anterior portion narrow,
the posterior portion with the sides subparallel or feebly rounded and usually with
a small tubercle at each anterior angle; densely, rather coarsely punctate, without
trace of median sulcus or carina. Elytra very broad, short, scutiform or subcordate
somewhat flattened on the disc anteriorly in the larger specimens, the humeri obliquely
truncate in front and more or less angulate externally; with rows of very coarse,
closely-placed, transverse punctures, separated by shining, sparsely punctulate, raised
interstices, the alternate interstices a little more prominent that the rest, the first
(sutural) with a setigerous tubercle at the commencement of the apical declivity in ¢.
Beneath closely, coarsely punctate; first ventral segment depressed in the middle at
the apex in g. Third tarsal joint feebly bilobed.
Length 14—2, breadth ~—14 mm. (f2).
Loc. Seychelles: Mahé.
This is an abundant insect in the high mountain-forests of Mahé, where it is
found on damp decaying leaves of Wormia ferruginea and Northea seychellarum on
the ground. It was obtained in the high forest of Morne Blane and Pilot up to
2000 feet, X.—XI. 1908; forest between Trois Freres. and Morne Seychellois, ca.
1500—2000 feet, XII. 1908 and I. 1909; forest above Cascade Estate, up to 2000 feet,
I.—III. 1909; Mare aux Cochons district, 1000—2000 feet, I.—II. 1909; forest on
Summit of “Montagne Anse Major,” ca. 2000 feet, I]. 1909. The very large number
of specimens captured by Mr Scott vary greatly in size and colour. The male is
easily recognizable by the tuft of erect hairs on the suture beyond the middle and
the inferiorly dilated shovel-shaped rostrum. The larger castaneous examples are some-
what thickly fulvo-setose, the smaller, more convex, blacker individuals having a shorter
cinereous setosity, which, however, is partly hidden by an earthy exudation.
53—2
414 PERCY SLADEN TRUST EXPEDITION
21. Cycloterinus erosus, n. sp. (Pl. 28, figs. 19, 194, 2.)
Short, opaque, glabrous; piceous or nigro-piceous, the antenne and tarsi, and
sometimes the rostrum, femora, and tibiz also, ferruginous. Rostrum in both sexes
very stout, arcuate, about as long as the prothorax, carinate along the middle and
sides, seriato-punctate, and opaque to near the tip, in ? a little less curved and more
shining towards the apex. Eyes very small. Prothorax nearly or quite as broad
as long, the posterior portion transversely quadrate, slightly widened forwards to the
obtusely angulate anterior angles, the tubulate anterior portion abruptly separated
from the rest; coarsely, confluently punctate. Elytra much wider than the prothorax,
subtriangular, the humeri obliquely truncate in front; seriato-foveolate, the interstices
3, 5, 7 each with a series of oblong, truncate, or subconical tubercles, which are sometimes
partly confluent on the disc, those on 7 projecting beyond the margins of the elytra
as seen from above, 4 and 6 also slightly raised. Beneath coarsely, closely punctate.
Third tarsal jomt moderately dilated, excavate above.
Length 14—2 mm. (¢9).
Loc. Seychelles: Mahé.
Thirteen specimens, picked up one or two at a time, in the high forests of
Morne Blane and Pilot, and also found in the Mare aux Cochons district, and on
the summit of “Montagne Anse Major.” This insect has the general facies of a minute
Brachycerus, the subconical or truncated tubercles on the elytra, which are sometimes
partly coalescent on the disc (forming interrupted ridges), projecting laterally beyond
the margins. The prosternum is unemarginate at the apex, and the anterior coxe
are moderately separated.
22. Cycloterinus spheropterus, n. sp.
Short, very convex, shining; rufo-piceous, the antennz, rostrum, and legs ferruginous,
set with minute, widely scattered, short, curled setz. Rostrum stout, arcuate, about
as long as the prothorax, opaque, rugulosely punctate, and carinate to beyond the
middle, smoother and shining at the tip. Eyes reduced to a few facets. Prothorax
slightly broader than long, gradually widened to beyond the middle and then abruptly
narrowed to the short tubulate anterior portion; coarsely, confluently punctate, and
with a shallow median groove. Elytra much wider than the prothorax, globose, the
humeri obliquely truncated in front and produced into an obtuse tooth externally ;
with closely packed rows of very coarse transverse punctures separated by narrow,
raised, shining interstices, 3 and 5 interruptedly costate, 5 with a conspicuous tubercle
at the commencement of the apical declivity. Third tarsal joint feebly lobed.
Length 2mm. (8).
Loc. Seychelles: Silhouette.
One specimen from the forest immediately above Mare aux Cochons, over 1000 feet,
IX. 1908. Ditters from small C. uncristatus by its globose, interruptedly costate elytra,
the anteriorly widened, rugosely punctate, canaliculate prothorax, and the simply
arcuate rostrum and non-cristate elytra of the male. The elytral costz are broken up
CHAMPION—COLEOPTERA ; CURCULIONIDA . 415
into several tubercles. The humeral protuberance is not quite symmetrical on the two
wing-cases. C spheropterus has the general facies of a small Acalles,
23. Cycloterimus microphthalmus, n. sp.
Oblong, convex, shining’; castaneous, the legs and the base of the antenne ferru-
ginous; sparsely clothed with minute, curled, suberect setz. Rostrum very stout,
arcuate, as long as the prothorax, rugulosely punctate, carinate, and opaque to near
the tip, the apex shining. Eyes reduced to a few facets. Prothorax barely as long
as broad, the posterior portion widened forwards to the rounded anterior angles and
then abruptly narrowed to the short tubulate anterior portion; densely, coarsely punctate.
Elytra short-oval, at the middle considerably wider than the prothorax; with rows of
closely placed, coarse, transverse punctures separated by very narrow, shining, raised
interstices. Beneath coarsely, closely punctate. Anterior coxze moderately distant.
Third tarsal joint small, feebly excavate above.
Length 14—14 mm. (6).
Loc. Seychelles: Silhouette.
Two specimens, from near Mont Pot-d-eau, ca. 1500 feet, VIII. 1908; one is
recorded as from a fallen “Bois Rouge” tree (Wormia ferruginea). Much smaller
than C. spheropterus, the prothorax more rounded at the sides and with the punctures
on the disc separate one from another, the elytra without tubercles and rounded at the
sides from the truncated base.
24. Cycloterinus cecus, n. sp.
Oblong, convex, opaque; reddish-brown, the rostrum and legs ferruginous; clothed
with minute, semierect, curled, ochreous set, the sete on the elytra condensed into a
compact line down the alternate interstices. Rostrum very stout, arcuate, nearly as
long as the prothorax, separated from the head by a deep transverse groove, minutely
punctate, rugulose and opaque towards the base, the long parallel-sided apical portion
shining, and wider than the rest. Eyes wanting. Prothorax elongate, subcylindrical,
the posterior portion subquadrate, as long as broad, and rounded at the sides anteriorly,
the narrower tubulate anterior portion extending forwards over the head ; densely
punctate, the punctures longitudinally confluent. Elytra oval, at the middle considerably
wider than the prothorax ; seriato-foveolate, the alternate interstices costate, the others
convex. Beneath very coarsely punctate. Prosternum long, arcuate-emarginate in front.
Anterior coxze very narrowly separated. Third tarsal joint small, feebly excavate above.
Length 2 mm. (?).
Loe. Seychelles: Mahé.
One specimen, found in the high forest of Morne Blanc or Pilot, in November, 1908.
The elongate, strongly cucullate prothorax gives this insect a very different appearance
from the forms described by Kolbe; nevertheless, it can quite well be placed in Cyclo-
termus for the present, some of the additional species now known being almost
intermediate. The antennee are wanting in the type.
416 PERCY SLADEN TRUST EXPEDITION
Group Pheenicobatina, n.
This group includes three genera agreeing in the following characters :—Tarsi with
joints 2 and 3 short, very broadly and equally dilated in the two sexes, the claws minute
or wanting ; rostrum free, deflexed, short and broad, or longer and dissimilarly formed, in
the two sexes; prothorax more or less cucullate in front; ventral segments 1 and 2 long
and connate, 3 and 4 very short, the sutures straight; prosternum unimpressed ; anterior
coxze more or less separated ; wings fully developed or wanting ; body oblong or elongate,
squamose or setose, rarely subglabrous above.
The numerous species obtained by Mr Scott, some of which seem to be very abundant,
appear to be absolutely confined to the endemic forests of the Seychelles: and as they
cannot be referred to any of the “tribes” of Lacordaire, a new group is required for their
reception. The peculiar tarsal structure of both sexes of these insects doubtless enables
them to cling tightly to the leaves of the endemic palms, &c., on which many of them
were observed in profusion. In steep places, where it was possible to look on to the
tops of palms (Stevensonia, &c.) growing below, certain species (e.g. Phanicobates vittatus)
were seen in large numbers sitting exposed to the light on top of the palm leaves ; they
presented a characteristic appearance as little dark linear objects resting lengthwise in the
crinkles of the leaves*. Various forms were continually beaten from palms, both from the
living and dead leaves: some small kinds appear to have been beaten principally, if not
entirely, from still hanging or fallen dead leaves. Whether any were beaten from
dicotyledonous plants or not, is uncertain. Several were found in the heads of felled
Verschaffeltia splendida (palms) in Silhouette. On a number of occasions certain species
were found sitting on the leaves of the endemic Pandani and of a low-growing mono-
cotyledonous plant (Curculigo seychellensis). Moreover, when endemic palms and
Pandani were cut down to investigate the leaf-base fauna, one or more forms of this
group were often found down between the leaf-bases: thus, in the case of four species
of Phanicobates, specimens are recorded as having been taken from between leaf-bases
of Pandanus, while in the case of two others specimens were found in a similar
situation in the palm Stevensonia. It may therefore fairly be assumed that they feed on
the palms and screw-pines.
The group “ Phcenicobatina” may be placed near the “ Erirrhina” (Hrirhinides) of
Lacordaire. The Family Proterrhinide of the Hawaiian Islands affords a somewhat
analogous instance of an isolated insular group of Coleoptera, all the species of which
(136) are endemic.
The genera may be tabulated thus:
Rostrum longer, dissimilarly formed in the two sexes, always smoother and more
slender in $; wings present or wanting Me Kae ... Pheenicobates.
Rostrum short and broad, similarly formed in the two sexes; wings wanting.
Tibize hooked at the apex; elytra oval; rostrum widened outwards
Phaenicobatopsis.
Tibiz unarmed at the apex; elytra oblong; rostrum not widened outwards
-Pheenicogenus.
* See general account of collecting in Vol. xiv. of these Transactions, 1910, p. 25.
CHAMPION—COLEOPTERA ; CURCULIONIDA ALT
PH@NICOBATES*, n. gen.
Rostrum nearly or quite as long as the prothorax, deflexed, dissimilar in the two
sexes, slender, cylindrical, and comparatively smooth in $, rugose, stouter, and often
gibbous above or angulate beneath (as seen in profile), and with the lower apical portion
shovel-shaped in the more typical forms in , the scrobes obliquely descending to beneath
the eyes; mandibles short, without scar, when closed just visible beyond the tip of the
rostrum ; antenne in f usually inserted nearer the tip of the rostrum than in ?, slender, the
scape abruptly clavate at the apex, the funiculus seven-jointed, joint 1 stout and obconic,
the club acuminate-ovate, annulate, pubescent; eyes moderately large, separated by less than
the width of the rostrum, inserted in the anterior portion of the head immediately behind
the base of the rostrum, the post-ocular portion of the head globose and punctate ;
prothorax long, more or less cucullate in front; scutellum minute or invisible; elytra
oblong, covering the abdomen at the tip, regularly punctate-striate ; prosternum unim-
pressed, arcuate-emarginate in front, the anterior cox inserted at about the middle,
narrowly or moderately separated (distant in P. pandanicola); mesothoracic epimera
small, not ascending; metasternum variable in length, the episterna narrow; ventral
segments 1 and 2 long, connate, 3 and 4 short, the sutures 2—4 straight and deep;
femora toothed or unarmed, the anterior pair only strongly toothed in some of the
species; tibiee armed with a short hook at the middle of the apical margin; tarsi (fig. 20 })
with joint 1 small, triangular, 2 and 3 short, very broadly and equally dilated, 3 emarginate
for the reception of the slender or oval fourth joint, 2 and 3 and the apex of 1 densely
spongy-pubescent beneath, the claws minute, divergent, sometimes wanting; body elongate,
subcylindrical, or oblong, sometimes fusiform in g, more or less squamose or setose, rarely
subglabrous above; wings fully developed or wanting.
Type, P. vittatus.
Twenty-five species are referred to this genus, many of them varying greatly in size.
The larger typical forms resemble a small Lexus, others have the general facies of a
Mecinus or Smacronyx. Four are without wings, and one of these species and two others
want the tarsal claws. Two of the large forms have the anterior femora only strongly
dentate ; and one, which seems to be attached to Pandanus, has the anterior coxee widely
separated.
a. Tarsal claws present, the claw-joint extending beyond the third.
1
a. Anterior coxee narrowly or moderately separated.
a. Wings fully developed.
ae
Anterior femora with a sharp tooth, the others unarmed: species large,
elongate.
Rostrum of ¢ flattened above, not dilated beneath; elytra usually
4
a.
vittate ; anterior coxz narrowly separated ... sae vittatus.
* The name Phenicobius has been used by Mérch in Mollusca (1852) and by Leconte in Anthribid
Coleoptera (1876); but Phenicobates is sufficiently different to be used for a genus of palm-frequenting
Curculionide.
418 PERCY SLADEN TRUST EXPEDITION
6b. Rostrum of ¢ abruptly bent and with lower apical portion shovel-
shaped ; elytra with two small patches of scales, and transversely
tumid before the middle, in $; anterior coxee more widely separated
flexirostris.
6%. All the femora with a more or less distinct tooth.
c’. Body long, cylindrical, ferruginous or brown above, usually mfuscate
or black along the sides ; rostrum of ¢ stout, of ? slender.
a. Vestiture short and wholly decumbent.
a’. Vestiture fine.
a’. Rostrum almost straight in both sexes... ... cylindricus.
b’. Rostrum strongly curved in 3 ie ae ... parallelus.
6°. Vestiture coarse; rostrum feebly curved in g ... mgrolimbatus.
b°. Vestiture very long, erect, and setiform towards apex... albosetosus.
d‘. Body moderately elongate, black, vestiture coarse and decumbent ;
rostrum very stout in ¢, comparatively stout in ...fortirostris.
e'. Body less elongate, the elytra in part ferruginous, vestiture consisting
of intermixed adpressed scales and erect setze ; rostrum slender in
both sexes.
c. Prothorax moderately rounded at sides; elytra longer and less
convex, the setz longer : ue --- duplovestitus.
d’. Prothorax strongly rounded at eee elytra shorter and more
convex, the setz short ... ae ... praslinensis.
Ce.
All the femora unarmed; rostrum of 2 stout, of 2 slender.
fi. Prothorax feebly constricted anteriorly; body oblong or elongate,
black : species small.
e. Vestiture of upper surface decumbent, moderately coarse.
ce. Rostrum of g (seen laterally) depressed above and arcuate beneath,
that of ? opaque: ... 506 ... depressirostris.
d°®. Rostrum of ¢ (seen laterally) ‘eae above and beneath, thus
appearing strongly sinuate, that of 9 shining fovewentris.
Rostrum of g gibbous above, that of 2 opaque --- CUrvupes.
f*. Rostrum of ¢ flattened or feebly curved above, that of 9 opaque.
‘. Eyes not prominent; elytra broader than prothorax symplex.
Gh
d’. Eyes rather prominent; elytra not wider than prothorax:
eé
species minute ae sao | UMS:
f°. Vestiture of upper surface wanting ody o opaque aoe and beneath ;
elytra broad, convex w+ peropacus.
4
g'. Prothorax strongly constricted Acuity: ood abloners in great part
ferruginous ; vestiture very coarse, decumbent.
g. Rostrum of g gibbous above... = ... gibbirostris.
h’. Rostrum of ¢ not gibbous above, Filial eed beneath.
g’. Prothorax and elytra fasciate ue Bs ... stricticollis.
h’. Prothorax and elytra not fasciate ... sas ... stevensonie.
b°. Wings rudimentary or wanting : species small.
CHAMPION—COLEOPTERA; CURCULIONIDA 419
d’. All the femora, or at least the anterior pair, dentate: body in part
ferruginous, oblong, more or less setose.
i‘. Prothorax and elytra with erect sete, the elytra not mucronate in ?
hispidulus.
7“. Elytra with erect sete towards apex only, usually mucronate in ?
cuspidatus.
ée’.
Femora unarmed: body black, convex, shining, vestiture wholly decumbent
lodoicee.
b.. Anterior coxee widely separated ; wings fully developed ; body elongate, cylin-
drical, wholly or in part ferruginous: vestiture very fine, decumbent ; ros-
trum slender in both sexes; all the femora feebly dentate... pandanicola,
b. Tarsal claws wanting, the claw-joint not, or very slightly, extending beyond the
third joint; anterior coxee narrowly separated ; femora unarmed ; body oblong,
black, almost bare above ; rostrum stouter in ¢ than in ?: species small.
c. Wings fully developed; elytra subcylindrical.
c. Rostrum of ? abruptly bent and moderately slender... ve. Glarus.
d®. Rostrum of ? arcuate and extremely slender ... Rs ... Trufitarsis.
d'. Wings wanting; elytra widened posteriorly _... ae ... sebertensis.
25. Phenicobates vittatus, n. sp. (Pl. 28, figs. 20, 20a, ¢; 200, anterior tarsus.)
Very elongate, narrow, subfusiform, feebly shining; black, the antennz (the club
excepted), the anterior half of the prothorax, a broad stripe running down the disc of each
elytron, the tibiee, the tarsi in part, and sometimes the intermediate and posterior femora
also, ferruginous, the prothorax and elytra occasionally entirely black, the elytral vittee in
immature examples ochreous ; sparsely clothed with minute, adpressed, hair-like, cmereous
scales. Rostrum opaque in both sexes, shorter than the prothorax ; ($) moderately broad,
flattened, somewhat depressed, rugulosely punctate, and squamose above, the shovel-shaped
lower apical portion not wider than the upper portion, feebly carinate above, and with a
sharp polished medium ridge beneath; (?) straight, slender, finely striate-punctate, and
almost bare. Head rugulosely punctate, squamose around the eyes. Prothorax much
longer than broad, narrowed and slightly constricted in front, feebly bisinuate at the base ;
densely, coarsely punctate, carinate down the middle anteriorly in ?, strongly cucullate in
front in g. Elytra very elongate, a little wider than the prothorax, gradually narrowing
from the base in ¢, subparallel in their anterior half in 2, conjointly rounded at the apex ;
with rows of closely-packed coarse punctures, the interstices narrow and rugulose.
Beneath coarsely, densely, the ventral segments 2—5 more finely and sparsely, punctate,
5 transversely depressed at apex in ¢. Anterior coxe narrowly separated. Anterior
femora armed with a sharp tooth.
Length (exclud. head) 24—44 mm.
Loc. Seychelles: Mahé, Silhouette.
Mahé, found in abundance; high forest of Morne Blane and Pilot, X.—XI. 1908 ;
high forest between Trois Fréres and Morne Seychellois, XII. 1908 ; forests above Cascade
SECOND SERIES—ZOOLOGY, VOL. XVI. 54
420 PERCY SLADEN TRUST EXPEDITION
Estate, up to 2000 feet ; Mare aux Cochons district, 1000—2000 feet, I.—IT. 1909 ; Morne
Seychellois, II. 1909. Silhouette : only two or three specimens were found, VIII.—IX. 1908.
A large, elongate form, superficially resembling a small Lixus, with the anterior
femora strougly dentate, the elytra usually vittate. Both this species and the next
vary greatly in size.
26. Phaenicobates flexirostris, n. sp. (PI. 23, figs. 21, 21a, g; 22, 9.)
Elongate, subfusiform (%), subcylindrical (2), opaque, the elytra and under surface
feebly shining; varying in colour from nigro-piceous with the front of the prothorax,
a broad dorsal stripe on each elytron, and the last three ventral segments ferruginous to
almost entirely black (the tarsi and antennee excepted) or ferruginous, the antennee (the
club excepted) always of that colour, the legs usually obscure ferruginous, with the
anterior femora more or less infuscate; sparsely clothed with rather coarse, adpressed,
setiform, cinereous or ochreous scales, which are condensed into four faint lines on the
prothorax and a line down each elytral interstice, the scales on the third interstice
clustered into an oblong spot at the base and another at about the basal fourth in ¢.
Rostrum opaque in both sexes, shorter than the prothorax; (¢) broad, abruptly bent
downwards at the middle, rugosely punctate and squamose above, the shovel-shaped,
horizontal, lower apical portion dilated posteriorly, extending outwards beyond the upper
portion, and with a polished lanciform median ridge beneath ; (?) straight, slender, sparsely,
finely striate-punctate, and almost bare. Head rugosely punctate, squamose. Prothorax
longer than broad, feebly constricted near the apex, rounded at the sides anteriorly ;
densely, rugosely punctate, carinate down the middle anteriorly in ?. LElytra a little
wider than the prothorax, narrowing from the base in g, subparallel in their anterior
half in 2, transversely depressed on the disc at the middle and below the base, the
intervening space appearing tumid in ¢; coarsely, closely punctate-striate, the interstices
narrow and transversely rugose. Beneath coarsely, densely, the ventral segments 2—5
more finely and sparsely, punctate, the first segment hollowed down the middle, and the
fifth transversely depressed at the apex, in ¢. Anterior coxee well separated. Anterior
femora armed with a sharp oblique tooth.
Length 2—4 mm. (29).
Loc. Seychelles: Mahé, Silhouette.
Found in large numbers in both islands: in Mahé, in much the same localities as the
preceding species; in Silhouette, both near Mont Pot-i-eau and the Mare aux Cochons.
The abruptly bent, more strongly spatulate rostrum of the male, the coarser vestiture,
which is condensed into two short streaks or spots on the third elytral interstice in the
same sex, and the more oblique femoral tooth, distinguish this species in all its varieties
from P. vittatus. Viewed in profile, the elytra of the male are transversely tumid before
the middle and have at this place a condensed patch of scales on the third interstice, which
is almost or quite wanting in the female. Small feebly-developed males have the rostrum
less abruptly bent downwards and less dilated.
27. Phenicobates cylindricus, n. sp.
Elongate, subcylindrical, opaque; ferruginous, the flanks of the prothorax, the sides
CHAM PION—COLEOPTERA ; CURCULIONIDA A421
of the elytra broadly, the base of the femora, the under surface in great part, and the
antennal club, black or piceous, the femora sometimes wholly ferruginous; sparsely
clothed with short, fine, decumbent, setiform, whitish scales, those on the elytra uni-
seriately arranged down each interstice. Rostrum (¢) opaque, moderately stout, almost
straight, much shorter than the prothorax, rugulosely punctate, squamose; (?) slender,
shining, finely striate. Prothorax about as long as broad, rounded at the sides, narrowed
and constricted towards the apex ; densely, finely, rugulosely punctate. Elytra elongate,
scarcely wider than the prothorax, subcylindrical in both sexes ; coarsely punctate-striate,
the interstices rugulose, narrow, and somewhat convex; the apices in @ armed with a
small dentiform prominence. Beneath shining, closely, moderately coarsely punctate ;
ventral segments 1 and 2 broadly depressed down the middle, and 5 transversely
depressed near apex, in g. Legs stout; femora each armed with.a short, sharp tooth.
Length (exclud. head) 223—3 mm. (79).
Loc. Seychelles: Mahé, Silhouette.
Mahé: high forest of Morne Blane and Pilot, X.—XI. 1908; forest between Trois
Freres and Morne Seychellois, 1500—2000 feet, XII. 1908; Cascade Estate and forest
above; Mare aux Cochons district, 1000—2000 feet, I.—II. 1909. Silhouette, near
Mont Pot-a-eau, ca. 1500 feet, VIIT. 1908.
Eleven specimens, varying very little in size. Distinguishable by its ferruginous
general coloration, with the sides of the prothorax and elytra black, the somewhat robust
build, the parallel-sided elytra, the fine vestiture, the uniformly unidentate femora, the
straighter non-spatulate rostrum of the male, and the shining, striate rostrum and
mucronate apices of the elytra of the female. The rostrum of the male is formed as
in the same sex of P. albosetosus.
28. Phenicobates parallelus, n. sp.
Elongate, cylindrical, somewhat depressed, opaque; ferruginous or brown, the flanks
of the prothorax and elytra, the head, rostrum, antennal club, femora, and under surface
in part, usually infuscate or black; sparsely clothed with minute, adpressed, hair-like,
cinereous or flavo-cinereous scales, those on the elytra uniseriately arranged down each
interstice. Rostrum (¢) stout, comparatively short, abruptly curved from near the base,
opaque, squamulose, and rugulosely punctate to the tip, and with the basal portion somewhat
gibbous (as seen in profile); (?) slender, cylindrical, almost straight, nearly as long as the
prothorax, shining, finely striate-punctate ; the antennze inserted near the tip in f, and at
about the middle in $. Prothorax about as long as broad, rounded at the sides, narrowed
and feebly constricted anteriorly, densely, finely, rugulosely punctate. Hlytra elongate,
cylindrical, scarcely wider than the prothorax; closely punctate-striate, the interstices
rugulose. Beneath shining, moderately coarsely, closely punctate; ventral segments 1
and 2 hollowed down the middle in ¢. Femora each with a small tooth. Tarsal joints 2
and 3 very broad.
Length (exclud. head) 13—21 mm. (¢?).
Loc. Seychelles: Mahé.
54—2
422 PERCY SLADEN TRUST EXPEDITION
The special localities for P. parallelus are, the high forest of Morne Blane and Pilot,
the Mare aux Cochons district, and the forest above Cascade Estate.
Five males and one female, varying greatly in size and colour, one large unicolorous
male having much stouter limbs than the smaller examples of the same sex. This insect
has the vestiture of the upper surface very fine and wholly decumbent, as in P. cylindricus,
from which it may be separated by its smaller size and narrower form, the less constricted
prothorax, the stouter, abruptly curved rostrum of the male, and the non-mucronate apices
of the elytra of the female.
29. Phencobates ngrolimbatus, n. sp.
Elongate, narrow, subcylindrical, feebly shining; ferruginous or obscure ferruginous,
the antennal club, the sides of the prothorax and elytra, and the under surface in part,
infuscate or black ; sparsely. clothed with short, curled, decumbent, rather coarse, whitish,
setiform scales, those on the elytra uniseriately arranged down each interstice. Rostrum
($) much shorter than the prothorax, feebly curved, almost parallel-sided, moderately stout,
rugulosely punctate, squamose; (?) longer and more slender, cylindrical, shining, finely
striate. Prothorax a little longer than broad, somewhat rounded at the sides, narrowed
and feebly constricted anteriorly, densely, finely punctate. Elytra elongate, subcylindrical,
scarcely wider than the prothorax; rather finely punctate-striate, the interstices narrow
and rugulose. Beneath shining ; closely, moderately coarsely punctate ; ventral segments 1
and 2 broadly depressed down the middle in g. Femora each with an indication of an
excessively minute tooth.
Length (exclud. head) 2;4,—22 mm. (29).
Loc. Seychelles: Mahé, Silhouette, Praslin.
A long series: found in Silhouette on a number of occasions in the forest near Mont
Pot-d-eau and above Mare aux Cochons: in Mahé, in the high forest of Morne Blanc, and
in the Mare aux Cochons district, 1000—2000 feet : in Praslin, in the Coco-de-mer forest,
Vallée de Mai, Cétes d’Or Estate. Narrower and less robust than P. cylindricus; the
vestiture coarser; the prothorax less constricted anteriorly; the elytra more finely
punctate-striate, the apices unarmed in the female; the femoral tooth almost obliterated
the rostrum feebly curved in the male.
30. Phenicobates albosetosus, n. sp.
Elongate, subcylindrical, feebly shining ; ferruginous, the elytra brown, the sides
of the latter, the prothorax, femora, and under surface more or less suffused with black,
the dark border to the elytra angularly extending imwards at a little before the
middle and the markings on the prothorax sometimes reduced to three longitudinal
streaks; sparsely clothed with short, coarse, decumbent, whitish or pale yellow,
decumbent, setiform scales, those on the elytra uniseriately arranged down each interstice
and abruptly becoming long, erect, and clubbed on the apical declivity. Rostrum (¢)
much shorter than the prothorax, almost straight, rather slender, slightly widened
apically, rugulosely punctate and squamose ; (?) slender and longer, shining, finely striate.
Prothorax longer than broad, rounded at the sides, narrowed and feebly constricted in
CHAMPION—COLEOPTERA ; CURCULIONIDA 423
front, densely, finely, rugulosely punctate. Elytra elongate, very little wider than the
prothorax, rather finely punctate-striate, the interstices rugulosely punctured ; the apices
not or feebly mucronate in ?. Beneath shining, closely, moderately coarsely punctate ;
ventral segments 1 and 2 broadly depressed down the middle in g. Femora each with a
small tooth.
Length (exclud. head) 22—22 mm. (f?).
Loc. Seychelles: Mahé, Silhouette.
Found in Mahé in the high forest of Morne Blanc and Pilot, in the forest above
Cascade Estate, and in the Mare aux Cochons district: and in Silhouette, in the forest
above Mare aux Cochons up to the highest parts. Five males and four females, one of the
latter having the elytra rather sharply mucronate at the apex. This is a form of
P. nigrolimbatus, with long, erect, whitish, clubbed setze on the apical declivity, and the
black border of the elytra angularly extending inwards before the middle. In one of the
Mahé females the elytra are mucronate at the apex, at some distance from the sutural
angle; it may belong to a different species.
31. Phenicobates fortirostris, n. sp.
Elongate, somewhat convex, opaque, the elytra shining; nigro-piceous or black,
the antennee (the club excepted) and tarsi ferruginous ; sparsely clothed with coarse,
whitish, adpressed, setiform scales, those on the elytra uniseriately arranged down
each interstice. Rostrum shorter than the prothorax, the antenne inserted beyond the
middle in both sexes, the apical portion slightly widened, parallel-sided in $, appearing
somewhat thickened; (¢) stout, rugulosely punctate to the tip, and squamose ; (?) more
slender, cylindrical, bare, and faintly punctate. Prothorax a little longer than broad,
rounded at the sides, narrowed anteriorly, and constricted just before the apex ; densely,
finely, rugulosely punctate. Elytra moderately long, very little wider than the prothorax,
parallel in their basal half; closely punctate-striate, the interstices rugose. Beneath
shining, closely punctate in 2, the ventral segments and the middle of the metasternum
sparsely punctured in 3, the first segment, at most, faintly hollowed down the centre, and
the fifth transversely depressed below the base, in this sex. Femora each with a small
tooth. Tarsal jomts 2 and 3 moderately dilated.
Length (exclud. head) 2—22 mm. (¢?).
Loc. Seychelles: Mahé.
A long series, varying a good deal in size, males predominating: they were found in
the high forests of Morne Blanc and Pilot, of Trois Fréres and Morne Seychellois, and of
the Mare aux Cochons district, up to ca. 2000 feet. This insect has the facies of a small,
narrow, elongate Barid. The vestiture is coarse, setiform, whitish, and entirely decum-
bent. Large males have the first two ventral segments somewhat swollen and the median
groove wanting, and the rostrum very stout. The females have a relatively thicker
rostrum than the same sex of the allied forms.
32. Phancobates duplovestitus, n. sp.
Moderately elongate, subopaque; ferruginous, the head and prothorax, the base of
424 PERCY SLADEN TRUST EXPEDITION
the rostrum, the sides of the elytra, the femora, tibiz, and under surface more or less
suffused with piceous or black; clothed with small, scattered, narrow, white scales inter-
mixed with numerous erect, blunt, pallid sete, the setze on the elytra uniseriately arranged
down each interstice and becoming much longer towards the apex, the legs also albo-
setose. Rostrum subcylindrical, rather slender, much shorter than the prothorax,
rugulose and dull in its basal half in %, striate-punctate and shining from the base in 8,
the antennz inserted at about the middle in both sexes. Prothorax as long as broad,
rounded at the sides, narrowed and feebly constricted anteriorly ; densely, finely,
rugulosely punctate. Elytra considerably wider than the prothorax, moderately long,
parallel at the base, gradually widened to about the middle, conjointly rounded at
the apex in both sexes, the humeri subrectangular ; finely punctate-striate, the inter-
stices rather broad and rugose. Beneath shining, coarsely, closely punctate; ventral
segment 1 broadly depressed down the middle in ¢. Femora each with a small sharp
tooth.
Length (exclud. head) 1}—2 mm. (@9).
Loc. Seychelles: Silhouette, Mahé.
Found in abundance: in Mahé, in the forests of Morne Blanc and Pilot (one specimen
being recorded as found between leaf-bases of a growing Pandanus), in the country above
Port Glaud, ca. 1000 feet, in the forests above Cascade Estate, and in the Mare aux
Cochons district; in Silhouette, both near Mont Pot-d-eau and in the forests above
Mare aux Cochons. A small form, clothed above with intermixed adpressed white scales
and erect setz, the latter becoming longer towards the apex. The wings, which are
extended in one of the smaller examples, are cousiderably longer than the body (rostrum
included), measuring 24 mm. in expanse.
33. Phanicobates praslinensis, n. sp.
Oblong, convex, subopaque; piceous, the antennz, rostrum, anterior portion of
prothorax, tarsi, and apex of abdomen ferruginous, the elytra reddish-brown, sometimes
darker along the sides; sparsely clothed with small whitish scales intermixed with short,
fine, suberect setee, the legs also setose. Rostrum slender, polished, shining, shorter than
the prothorax, slightly widened at the tip, finely striate-punctate, the antennz inserted
behind the middle. Prothorax about as long as broad, strongly rounded at the sides,
narrowed anteriorly, very densely, finely, rugulosely punctate. Elytra comparatively
short, a little wider than the prothorax, subparallel at the base, obliquely narrowed
posteriorly; coarsely punctate-striate, the interstices narrow and rugose. Beneath
shining, coarsely, closely punctate. Femora each with a small sharp tooth. Wings fully
developed.
Length (exclud. head) 14—14 mm. (@).
Loc. Seychelles: Praslin.
Seven specimens, apparently all females. This species agrees with P. duplovestitus
in having the upper surface clothed with intermixed scales and setze, but the setz in the
Praslin insect are short throughout. It also differs from the above-mentioned form in
=
CHAMPION—COLEOPTERA ; CURCULIONIDA 425
having the rostrum (?) more slender, the prothorax more rounded at the sides, the elytra
relatively shorter, more convex, and more coarsely punctate-striate. The examples
described were found in the Coco-de-mer forest, in the Vallée de Mai, on the Cétes d’Or
Estate, on Nov. 28th and 29th, 1908. The insect may be attached to Lodoicea, the
Coco-de-mer palm.
34. Pheenrcobates depressirostris, n. sp.
Moderately elongate, rather convex, somewhat shining; black, the antennz (the
club excepted) and tarsi ferruginous; sparsely clothed with coarse, scattered, adpressed,
setiform, white scales, those on the elytra uniseriately arranged down each interstice.
Rostrum (¢) stout, about as long as the prothorax, depressed towards the middle, sinuate
and strongly arcuate beneath (as seen in profile), and rugosely punctate to the tip ; (?)
slender, cylindrical, nearly straight, opaque, closely striate-punctate, shining at the tip ;
antennze inserted near the apex in fg, and at about the apical third in ?. Prothorax as
long as broad, rounded at the sides, narrowed and constricted anteriorly ; densely punctate,
sometimes with an indication of an abbreviated smooth median line. Elytra slightly
wider than the prothorax, subparallel in their basal half; closely, rather coarsely punctate-
striate, the interstices rugulose. Beneath shining, very sparsely punctate; ventral
segment 1 slightly hollowed down the middle, and 5 with a transverse depression, in ¢.
Femora unarmed, the anterior pair much thickened in ¢. Tarsal joints 2 and 3 moderately
dilated.
Length (exclud. head) 14—2 mm. (#9).
Loc. Seychelles: Mahé. )
Twenty specimens, including a pair mounted on the same piece of card, all from the
Cascade Estate, from an altitude of about 1000 feet. In this insect the rostrum of the
male is depressed at about the middle, and strongly arcuate and sinuate beneath (as seen
in profile), much as in the same sex of the very different P. flexirostris. The females
are separable from those of P. curvipes by the less curved tibiz, the slightly longer
rostrum, the smoother ventral surface and the coarser vestiture, which shows no sign
‘of condensation into a definite line on the second elytral interstice towards the apex.
35. Phanicobates foveiventris, n. sp.
Moderately elongate, subopaque ; black or piceous, the antennze and tarsi ferruginous;
sparsely clothed with small, white, adpressed, setiform scales, those on the elytra uniseri-
ately arranged down each interstice. Rostrum (%) about as long as the prothorax, very
stout, sinuate, strongly tumid near the base above and with the lower apical portion
dilated into a shovel-shaped process (appearing broad and strongly sinuate when viewed
in profile), rugosely punctate and squamose to the tip; (?) slender, cylindrical, almost
straight, shining throughout, at most feebly striate-punctate. Prothorax slightly longer
than broad, somewhat rounded at the sides, narrowed and constricted anteriorly in 2, less
distinctly so in 2; densely punctate. Elytra moderately long, subparallel in their basal
half in both sexes, very little wider than the prothorax in ¢, and of about the same width
in 2; closely punctate-striate, the interstices very narrow and rugulose. Beneath shining,
426 PERCY SLADEN TRUST EXPEDITION
rather coarsely punctate; ventral segment 1 broadly excavate down the middle, and
5 with a deep rounded fovea at the apex, in ¢. Femora unarmed; anterior pair of
very stout (appearing strongly arcuate, or even gibbous, in front, when the insect
is viewed in profile, with the legs unextended). ‘Tarsal joints 2 and 3 moderately
widened.
Length (exclud. head) 15—13 mm. (¢9).
Loc. Seychelles: Mahé, Silhouette.
Found in Mahé in the high forest of Morne Blanc and Pilot, the country above Port
Glaud ca. 1000 feet, the forests above Cascade Estate, and the Mare aux Cochons district :
also one specimen from the Silhouette Mare aux Cochons. Eleven specimens, nine of
which are males, showing various degrees of development of the rostrum. The two
females referred to this species (one of which is mounted on the same card with
a typical male) may be known from the corresponding sex of all the nearly allied
forms by the shining smoother rostrum, the latter being almost impunctate in one
example.
36. Phanicobates curvipes, n. sp.
Moderately elongate, robust, dull; black, the antennee (the club excepted) and tarsi,
and sometimes the tip of the rostrum (?) and the anterior margin of the prothorax also,
ferruginous or rufo-testaceous ; sparsely clothed with rather coarse, adpressed, setiform,
white scales, those on the elytra arranged in an irregular line down each interstice,
usually becoming closer and regularly placed on the second towards the apex. Rostrum
(¢) stout, a little shorter than the prothorax, curved, more or less tumid above behind
the middle (so as to appear constricted at the base), rugosely punctate and squamose to
the tip; (¢) slender, cylindrical, almost straight, bare, closely striate and opaque, shining
at the tip only ; the antennz inserted near the apex in ¢, and considerably beyond the
middle in ¢. Eyes depressed. Prothorax slightly longer than broad, feebly rounded
at the sides, constricted towards the apex, more narrowed anteriorly in $ than in ;
densely punctate. Elytra moderately long, wider than the prothorax, gradually narrowed
from the base in ¢, subparallel in their basal half in 3; closely, rather coarsely punctate-
striate, the interstices rugulose. Beneath shining, rather sparsely punctate ; first’ two
ventral segments broadly depressed down the. middle in g. Legs short, stout ; femora
unarmed, the anterior pair greatly thickened ; anterior and intermediate tibie strongly
bowed at the base in both sexes, and arcuately dilated externally ; tarsal] jomts 2 and 3
very broadly dilated.
Length (execlud. head) 145—14% mm. (¢).
Loc. Seychelles: Mahé, Silhouette.
Mahé, found in the high forest of Morne Blanc and Pilot, the country above Port
Gland, the high forests above Cascade Estate, and the Mare aux Cochons district
1000—2000 feet ; Silhouette, found in the forests above Mare aux Cochons.
A long series, varying greatly in size and in the development of the tumid upper
portion of the rostrum in the male. Viewed in profile, the rostrum in this sex is also
CHAMPION—COLEOPTERA ; CURCULIONIDA A427
slightly arcuate at the middle beneath, and narrower towards the base and apex. Small
females can be separated from the extremely closely allied P. tenuis by their relatively
broader elytra, less prominent eyes, and rather stouter anterior femora. Compared with
P. gibbirostris, the legs and rostrum (f) are shorter and stouter, the prothorax is
much less constricted anteriorly, the scales on the upper surface are smaller, and the
body is almost wholly black.
37. Phenicobates simplex, n. sp.
Moderately elongate, robust, dull (shining when denuded); black, the suture and
anterior margin of the prothorax sometimes piceous or reddish, the antennz (the club
excepted) ferruginous, the tarsi piceous or obscure ferruginous; sparsely clothed with
minute, adpressed, cinereous or whitish setiform scales, those on the elytra uniseriately
arranged down each interstice. Rostrum (¢) stout, shorter than the prothorax, flattened
above and slightly arcuate beneath (as seen in profile), rugose and squamose ; (?) slender,
cylindrical, opaque, closely striate-punctate. Eyes not prominent. Prothorax slightly
longer than broad, subeylindrical, feebly constricted and a little narrowed anteriorly ;
closely, coarsely punctate, sometimes with a short smooth median line on the disc.
Elytra moderately long, convex, a little wider than the prothorax, subparallel at the
base in both sexes; rather coarsely punctate-striate, the interstices rugulose. Ventral
segments 1 and 2 broadly, shallowly depressed down the middle in f. Legs stout ;
femora unarmed; anterior and intermediate tibie arcuate externally; tarsal joints
2 and 3 moderately dilated.
Length (exclud. head) 14—14 mm. (79).
Loc. Seychelles: Mahé, Silhouette.
Found in Mahé in the forests above Cascade Estate, the country above Port Glaud,
and the Mare aux Cochons district, at 1000—2000 feet: in Silhouette, at the Mare
aux Cochons and in the forest above. Nine specimens, all males but two. This insect
is very closely related to P. curvipes, from which it differs in having the rostrum of
the male flattened and not in the least tumid above, the elytra subparallel at the base
in both sexes, and the anterior tibiee less curved. The non-sinuate, shorter rostrum of
the male separates the present species from P. depressirostris. The tarsi are usually
infuseate. Abraded examples are shining and coarsely punctate above.
38. Phenicobates tenuis, n. sp.
Moderately elongate, narrow, somewhat convex, opaque (shining when denuded
of the vestiture) ; black or piceous, the antenne, tip of the rostrum, and tarsi ferruginous,
the tarsi often infuscate ; thickly set with cinereous or whitish, rather coarse, adpressed,
setiform scales, those on the elytra uniseriately arranged down each interstice. Rostrum
(¢) stout, shorter than the prothorax, feebly curved above (as seen in profile), rugosely
punctate and squamose to the tip; (?) slender, cylindrical, almost straight, opaque,
closely striate-punctate ; the antennz inserted near the tip in 2g, and considerably beyond
the middle in ¢. Eyes rather prominent. Prothorax longer than broad, feebly con-
stricted and narrowed anteriorly; closely punctate. Elytra long, not or very little
SECOND SERIES—ZOOLOGY, VOL. XVI. 55
428 PERCY SLADEN TRUST EXPEDITION
wider than the prothorax, parallel in their basal half in 9, and very gradually narrowing
from about the basal third in ¢; closely punctate-striate, the interstices rugulose.
Beneath rather sparsely punctate; first ventral segment shallowly depressed down the
middle in g. Femora unarmed. Anterior tibiz arcuate on their outer edge. Tarsal
joints 2 and 8 moderately dilated.
Length (exclud. head) 14—13} mm. (29).
Loc. Seychelles: Mahé, Silhouette, Praslin.
Mahé: high forest of Morne Blane and Pilot, country above Port Glaud, forests
above Cascade Estate (including the stunted forest on the summit of Mount Sebert,
ca. 2000 feet), high forest behind Trois Fréres, and the Mare aux Cochons district.
Silhouette: found both near Mont Pot-a-eau, ca. 1500 feet, and at the Mare aux Cochons
and in the forests above. Praslin: one specimen from the Coco-de-mer forest in the Vallée
de Mai, Cétes d’Or Estate. This is evidently a common insect in the islands. A minute,
narrow form, thickly set with small whitish setiform scales, the femora unarmed, the
rostrum of the male simply curved above (as seen in profile), the eyes more prominent
than in the allied forms, the elytra not or very little wider than the prothorax in either
sex. When fresh the entire insect is covered with an exudation which almost hides
the sculpture, and makes the surface appear opaque. The tarsi are often infuscate. In one
male the fifth ventral segment has an oblong shining prominence, no doubt accidental,
as it is not exactly in the centre. The female, as usual, has the elytra more parallel-sided
than the male.
39. Pheenicobates peropacus, n. sp.
Comparatively short, convex, opaque, and alutaceous above and beneath ; black or
piceous, the antennze (the club excepted) and tarsi, and sometimes the rostrum, the anterior
margin of the prothorax, and tibiz also, ferruginous; the prothorax and elytra almost
glabrous, the head, rostrum (¢), and legs clothed with small, adpressed, white, setiform
scales. Rostrum (¢) stout, considerably shorter than the prothorax, thickened towards
the base beneath (appearing arcuately dilated inferiorly when viewed in profile), rugulosely
punctate; (?) subeylindrical, more slender, and striate-punctate; the antennz inserted
far beyond the middle in both sexes, at near the tip in g. Prothorax about as long as
broad, narrowed and feebly constricted anteriorly; densely, finely punctate. Elytra
much wider than the prothorax, comparatively short, somewhat rounded at the sides,
subparallel at the base, the humeri obtuse; rather coarsely punctate-striate, the imter-
stices densely alutaceous. Beneath densely alutaceous, the metasternum and first ventral
segment with a few widely scattered fine punctures. Legs short, stout, the femora
unarmed, the second and third tarsal joints moderately dilated.
Length (exclud. head) 12—12 mm. (?2).
Loc. Seychelles: Silhouette, Mahé.
Mahé, found in the high forest of Morne Blanc and Pilot, the high forest between
Trois Freres and Morne Seychellois, 1500—2000 feet, the forests above Cascade Hstate,
and the Mare aux Cochons district. Silhouette, from the forest near Mont Pot-a-eau
(ca. 1500 feet) and above Mare aux Cochons (over 1000 feet).
CHAMPION—COLEOPTERA ; CURCULIONIDAI 429
Thirty-eight specimens. A small, convex, black or piceous form, almost glabrous
and opaque above, and comparatively smooth, opaque, and densely alutaceous beneath,
the femora unarmed, the tarsal joints 2 and 3 moderately dilated. The first ventral
segment is almost unimpressed in the male.
40. Phenicobates gibbirostris, n. sp. (Pl. 28, figs. 23, 23a, 3g.)
Moderately elongate, opaque ; ferruginous, rarely piceous with the antennee (the club
excepted) and tarsi ferruginous; clothed with irregularly distributed, coarse, curled,
decumbent, yellowish, setiform scales. Rostrum (f%) stout, curved, shorter than the
prothorax, gibbous and inflated towards the base above, rugosely punctate and squamose
to the tip; (2) straight, slender, flattened-cylindrical, closely striate-punctate. Prothorax
as long as broad, strongly constricted beyond the middle, rounded at the sides, the basal
portion in fully developed males as wide as the elytra, much narrower in smaller males
and in 9, the anterior portion somewhat tubulate; densely, finely punctate. Elytra
moderately long, subparallel at the base; rather coarsely punctate-striate, the interstices
narrow and rugulose. Beneath alutaceous, sparsely punctate; first ventral segment
obsoletely foveate at the apex in #2. Femora unarmed. ‘Tarsal joints 2 and 3 moderately
dilated.
Length (exclud. head) 13—24 mm. (?9).
Loc. Seychelles: Silhouette, Mahé.
Found in abundance in the same localities as the preceding species, the specimens as
usual varying much in size, the males, however, always having the rostrum conspicuously
inflated and gibbous above, instead of being angularly dilated beneath as in the same
sex of P. stevensonie and P. stricticollis. The coarse elytral vestiture is irregularly
distributed in the present species, showing very little tendency to a linear arrangement,
and there is no trace of bare transverse fascize.
41. Phanicobates stricticollis, n. sp. (Pl. 28, figs. 24, 24a, 3g.)
Moderately elongate, narrow, opaque; ferruginous or obscure ferruginous, the
antennal club infuscate, the base of the prothorax and a common transverse subapical
fascia on the elytra—sometimes extending forwards along the sides or reduced to a
marginal spot on each elytron—black ; somewhat thickly clothed with coarse, curled,
decumbent, setiform, yellowish scales, which are condensed into a small patch between
the eyes, an interrupted transverse median fascia on the prothorax, and a common basal,
median, and apical fascia on the elytra (leaving two almost bare transverse spaces on
the disc). Rostrum (g) moderately stout, flattened above, angularly dilated beneath,
squamose and rugosely punctate to the tip; (?) more slender, flattened-cylindrical, bare,
closely striate-punctate. Prothorax as long as broad, strongly constricted beyond the
middle, the apical portion much narrower than the basal portion ; densely, finely, rugu-
losely punctate, and with an indication of a smooth raised median line in 3. Elytra
moderately long, closely, rather coarsely punctate-striate, the interstices very narrow and
rugulose. First ventral segment slightly depressed in the middle at the apex in 2.
Femora unarmed.
p=)
430 PERCY SLADEN TRUST EXPEDITION
Length (exclud. head) 13—2 mm. ($2).
Loc. Seychelles: Silhouette, Mahé, Praslin.
Nine specimens. Mahé, from the Mare aux Cochons district (1000—2000 feet), and
from between leaf-bases of a growing Stevensonia-palm near Morne Blanc. Silhouette,
from the forest above Mare aux Cochons, including some specimens taken from between
leaf-bases of a growing Stevensonia at 2000 feet. Praslin, from the Coco-de-mer forest in
the Vallée de Mai. In this insect the coarse elytral vestiture, which is dense at the base
and sublinearly arranged, is interrupted by two more or less distinct bare transverse
fascize, the ground-colour of the subapical fascia being usually black. The curled setz on
the prothorax are shorter than in P. stevensonie, and do not extend over the head as in
that insect. The coarse, curled scales on the legs give a dilated appearance to the tibize.
The females have an indication of a smooth median ridge on the prothorax.
42. Phanicobates stevenson, n. sp.
Moderately elongate, subopaque ; piceous or reddish brown, the antenne and tarsi,
and sometimes the femora and tibize also, ferruginous or obscure ferruginous, the elytra
sometimes with an oblong black patch at the sides; clothed, the legs included, with
scattered, coarse, curled, setiform, decumbent, yellowish scales (those on the elytra
elongate-oval in shape), which are condensed into a small patch between the eyes, an
apical and a median fascia on the prothorax (the curled scales of the apical fascia pro-
jecting forwards over the head), and several interrupted lines on the elytra. Rostrum (¢)
a little shorter than the prothorax, moderately stout, flattened above and angularly
dilated beneath, squamose and rugosely punctate to the tip; (?) longer and more slender,
flattened-cylindrical, bare, closely striate-punctate ; the antennz inserted towards the tip
in g, and at about the middle in ?. Prothorax strongly constricted beyond the middle,
the basal portion in fully-developed males arcuately dilated and wider than the elytra,
narrower in smaller males and in ?; densely, finely punctate, the disc sometimes with an
indication of a smooth median line. Elytra moderately long, very gradually narrowing
from near the base in ¢, subparallel in their basal half in 2; rather coarsely, closely
punctate-striate, the interstices very narrow and rugulose. Beneath closely, moderately
coarsely punctate, smoother in @; ventral segment 1 with a deep, transverse, foveiform
depression at the apex in ¢ and a smaller fovea in ?. Legs rather stout, coarsely setose ;
femora unarmed ; tarsal joints 2 and 3 moderately broad.
Length (exclud. head) 14—22 mm. (69).
Loc. Seychelles: Silhouette, Mahé.
Silhouette: from near Mont Pot-a-eau, ca. 1500 feet ; from the forests above Mare
aux Cochons, including some specimens from decayed heads of felled Verschaffeltia-palms
just above the plateau, and some taken from between leaf-bases of a growing Stevensonia
at about 2000 feet. Mahé: from the high forests of Morne Blane and Pilot, from the
forests above Cascade Estate (including some specimens taken from between leaf-bases of
a growing Stevensonia at about 1000 feet), and from the Mare aux Cochons district.
Thirty-five examples, varying greatly in size and in the development of the prothorax
CHAMPION—COLEOPTERA ; CURCULIONIDA 431
in the male, one specimen of this sex having the basal portion arcuately dilated and wider
than the elytra. Amongst the species with a strongly constricted prothorax and unarmed
femora P. stevensonie may be known by the very coarse vestiture of the upper surface
and legs, the curled projecting setiform scales along the anterior margin of the prothorax,
and the sublineate, uninterrupted arrangement of the elongate-oval scales on the elytra.
Small males might easily be mistaken for P. stricticollis, in which the lines of scales on
the elytra are interrupted by two bare transverse fascize.
43. Phenicobates hispidulus, n. sp. (Pl. 28, figs. 25, 25a, ¢.)
Moderately elongate, subopaque ; ferruginous, the head and prothorax, the antennal
club, the base of the rostrum, the suture and sides of the elytra, the femora, tibize, and under
surface more or less suffused with piceous or black; sparsely clothed with minute adpressed
whitish scales intermixed with curled, suberect, fine, blunt setz, the setz on the elytra
uniseriately arranged down each interstice and becoming very long towards the apex, the
legs also setose. Rostrum subcylindrical, a little shorter than the prothorax, rugulosely
punctate and opaque to near the tip in ¢, more shining and striate in 9, the antennze
inserted at (2) or slightly beyond (¢) the middle. Prothorax about as long as broad,
rounded at the sides, narrowed and feebly constricted anteriorly; densely, finely,
rugulosely punctate. Elytra connate, a little wider than the prothorax, rather convex,
rounded at the sides, gradually widened to about the middle; rather coarsely punctate-
striate, the interstices narrow and rugose. Beneath shining, coarsely, closely punctate ;
ventral segment 1 feebly depressed down the middle in g. Femora each with a small
tooth. Wings wanting.
Length (exclud. head) 14—2 mm. (f9).
Loc. Seychelles: Silhouette, Mahé.
Silhouette, from high forest above Mare aux Cochons. Mahé, from forests of Morne
Blane and Pilot, from high forest between Trois Fréres and Morne Seychellois (1500—
2000 feet), and from the forests above Cascade Estate.
Twelve examples. Larger and more robust than P. duplovestitus, the elytra rounded
at the sides, the setze less erect on the prothorax and becoming (as in P. albosetosus) very
long on the apical portion of the elytra, the minute intermixed adpressed scales fewer in
number on the elytral surface, the seriate punctures on the latter coarser. In one of the
Silhouette examples the elytral setze are very long throughout. The longer and more
abundant setosity of the upper surface and the less elongate shape separate P. hispidulus
from P. cuspidatus.
44. Phenicobates cuspidatus, n. sp. (Pl. 28, fig. 26, 9.)
Elongate, narrow, somewhat depressed, more or less widened posteriorly, subopaque ;
ferruginous or brownish-ferruginous, with the head, base of the rostrum, antennal club,
flanks of the prothorax, sides of the elytra, femora, and under surface piceous or black,
sometimes almost wholly black above, or entirely ferruginous; very sparsely clothed with
minute, narrow, setiform, adpressed, pale yellowish scales, those on the elytra uniseriately
arranged down each interstice and becoming much longer and erect towards the apex.
432 PERCY SLADEN TRUST EXPEDITION
Rostrum cylindrical, rather slender, a little shorter than the prothorax, rugosely punctate,
squamulose, and opaque to near the tip in 2, shining, bare, and finely striate-punctate
in 2, the antennze inserted at about the apical third im ¢ and at the middle in 9.
Prothorax rather longer than broad, rounded at the sides, narrowed and feebly constricted
anteriorly ; densely, finely, rugulosely punctate. Elytra connate, long, gradually widened to
beyond the middle, rounded at the sides, usually mucronate at the apex m ?; very closely
punctate-striate, the interstices rugulose and not wider than the striz. Beneath shining,
coarsely punctured ; ventral segment 1 hollowed down the middle in g. Anterior femora
with a minute tooth, the others sometimes obsoletely dentate. Tarsal joimts 2 and 3 very
broadly dilated. Wings wanting.
Length (exclud. head) 13—24 mm. (29).
Loc. Seychelles: Mahé.
Found in abundance, males predominating: from the forests of Morne Blanc and
Pilot, including one specimen from between leaf-bases of a growing Pandanus ; from high
forests behind Trois Fréres and the slopes of Morne Seychellois, 1500—2000 feet ; from
the forests above Cascade, and the Mare aux Cochons district. Some specimens, irre-
spective of sex, have the elytra considerably widened posteriorly, others again have the
prothorax nearly as wide as the broadest portion of the elytra. A small, elongate,
posteriorly-widened form, with the upper surface usually ferruginous or brown, bordered
laterally with black ; the elytra conspicuously setose towards the apex, and in the female
usually mucronate at the tip; and the rostrum cylindrical in both sexes. The dentiform
prominence at the apex of each elytron of the female is usually well developed, but in
two examples it is minute and inconspicuous, and is sometimes entirely wanting.
45. Phaenicobates lodoicee, n. sp.
Oblong, convex, shining; black, the antennz, tarsi, and tip of the rostrum rufo-
testaceous ; clothed with small, hair-like, adpressed, whitish scales, those on the elytra
uniseriately arranged down each interstice. Rostrum slender, cylindrical, as long as the
prothorax, almost straight, faintly striate laterally. Prothorax transverse, strongly
rounded at the sides, narrowed and feebly constricted anteriorly ; closely punctate, the
punctures well separated on the disc. Elytra connate, comparatively short, not wider
than the prothorax, subparallel in their basal half, the humeri obtuse ; rather coarsely
punctate-striate, the interstices narrow, transversely wrinkled. Beneath sparsely punc-
tate; ventral segments 3 and 4 extremely short. Legs comparatively slender ; femora
unarmed ; tarsal joints 2 and 3 moderately dilated.
Length (exclud. head) 1}—14 mm. ().
Loc. Seychelles: Mahé, Praslin.
Three females: one labelled as having been taken from the leaf-bases of a growing ¢
“Coco-de-mer” (Lodoicea seychellarum) in the Vallée de Mai, Praslin, on Nov. 28th,
1908; one from the Cotes d’Or Estate, from the Coco-de-mer forest, Praslin, same date ;
one from the high damp forest between Trois Freres and Morne Seychellois, Mahé, in
Dec. 1908. A small, shining, convex form not very closely related to any of the other
species here described. The connate elytra denote absence of wings.
CHAMPION—COLEOPTERA ; CURCULIONID A A433
46. Phemcobates pandanicola, n. sp. (Pl. 28, fig. 27, 3.)
Elongate, moderately convex, subeylindrical, shining; varying in colour from
brownish-ferruginous, or ferruginous with the sides of the prothorax and elytra infuscate
or black, to black, with the antenne, tibiz, tarsi, tip of rostrum, and last three ventral
seoments ferruginous, the elytra often with a broad sordid ochreous stripe on the disc
of each in dark examples; very sparsely clothed with minute, adpressed, hair-like,
whitish scales, those on the elytra uniseriately arranged down each interstice and
becoming coarser along the basal margin ; the head with two lines of narrow white scales
between the eyes. Rostrum much shorter than the prothorax, cylindrical, slender, very
little stouter in f than in ?, the basal portion rugose and opaque in 4, and closely striate
in 2, the apical portion shining and almost smooth in both sexes, the antennee inserted
at () or just beyond (¢) the middle. Prothorax much longer than broad, suboval and as
broad as the elytra in fully developed $; narrower in 9, feebly constricted and narrowed
anteriorly in both sexes; closely, rather coarsely punctate, the interspaces alutaceous.
Elytra moderately elongate, subparallel in their basal half, rather coarsely punctate-
striate; the interstices rugulose. Beneath closely, coarsely punctate; ventral segments
1 and 2 hollowed down the middle in g. Femora each with a small tooth, which is
almost obsolete in small examples. Tarsi with joints 2 and 3 extremely broad, 4 elongate-
oval (as seen from above), the claws very minute. Coxee widely separated.
Length (exclud. head) 13—22 mm. (69).
Loc. Seychelles: Mahé, Silhouette, Praslin.
A long series, varying greatly in size and colour, and also in the lateral expansion of
the prothorax in the males. The more widely separated coxe separate this species from
all the allied forms. In Mahé this species was found near Morne Blanc, several of the
specimens having been taken from between the bases of the leaves of growing Pandanus ;
also in the high forest behind Trois Fréres, and above Cascade Estate. In Silhouette it
was found near Mont Pot-a-eau, ca. 1500 feet, and between the leaf-bases of growing
Pandanus hornev in the forest above Mare aux Cochons. In Praslin several examples
were found between leaf-bases of growing Pandanus on Cotes d'Or Estate. Most of the
specimens therefore are known to have been found in Pandanus, which the insect
doubtless attacks.
47. Phenicobates alatus, n. sp.
Elongate, narrow, rather convex, somewhat acuminate at apex, subopaque, almost
glabrous ; black above, rufo-piceous beneath, the antennze (the club excepted) ferruginous,
the tarsi piceous or obscure ferruginous. Rostrum (%) shorter than the prothorax,
moderately stout, feebly arcuate above (as seen in profile), rugosely punctate ; (2) slender,
cylindrical, curved, abruptly deflexed from about the middle, the apical portion shining
and finely punctate. Prothorax longer than broad, gradually narrowed from about the
middle forwards, feebly constricted towards the apex; closely, finely punctate, the
interspaces densely alutaceous. Elytra long, slightly wider than the prothorax, sub-
parallel in their basal half, more or less produced at the apex, the apices narrow and
434 PERCY SLADEN TRUST EXPEDITION
sometimes subcaudate; closely, rather coarsely punctate-striate, the interstices very
narrow and rugulose. Beneath shining, closely, rather coarsely punctate; ventral
segments 1 and 2 shallowly suleate down the middle in g. Legs short ; femora unarmed ;
tarsal joints 2 and 3 moderately broad. Wings fully developed.
Length, (exclud. head) 14—2 mm. ($@).
Loc. Seychelles: Silhouette, Mahé.
Twenty-five specimens, varying greatly in size: found in Silhouette, near Mont
Pot-a-eau and in the forests above Mare aux Cochons: in Mahé, in the forests of
Morne Blane and Pilot (including a specimen found between leaf-bases of a growing
Pandanus), in the stunted forest on the summit of Mount Sebert, in the Mare aux
Cochons district and the forest on the summit of “ Montagne Anse Major” (ca. 2000
feet) : and on two small islets lying off the coast of Mahé (Long and Round Islands, VII.
1908, 11 ¢ and $). Compared with P. sebertensis this species has the elytra a little more
convex, not widened posteriorly, and more acuminate at the apex; the rostrum of the
male is not so stout, and less dilated towards the base beneath, that of the female being
abruptly deflexed from about the middle ; and the tarsi are usually more or less infuscate.
One male example from Silhouette has the apices of the elytra considerably produced.
The Long Island series are taken as the types.
48. Phanicobates rufitarsis, n. sp.
Elongate, cylindrical, convex, opaque, subglabrous; black, the antennee and tarsi
rufo-testaceous. Rostrum (?) shorter than the prothorax, very slender, cylindrical,
strongly arcuate, shining, almost smooth. Eyes large. Prothorax longer than broad,
cylindrical, feebly constricted towards the apex and slightly narrowed anteriorly ;
densely, finely, rugulosely punctate, except along a very narrow alutaceous median
space. Elytra elongate, parallel in their basal half, slightly wider than the prothorax,
conjointly rounded at the apex; closely punctate-striate, the imterstices flat, a little
wider than the striz, densely alutaceous. Legs stout; femora unarmed; tarsal joints
2 and 3 very broadly dilated.
Length (exclud. head) 25 mm. (?).
Loc. Seychelles : Mahé.
One specimen, found in the high forest of Morne Blanc. A remarkably distinct form
related to P. alatus, characterized by its extremely slender, shining, arcuate rostrum (9),
large head, elongate, cylindrical body, and very broadly dilated rufo-testaceous tarsi.
Under the microscope scattered minute cinereous scales are visible on the body and legs.
The insect doubtless has fully developed wings, but the elytra have not been opened for
examination.
49. Phenicobates sebertensis, n. sp.
Elongate, narrow, somewhat widened posteriorly, opaque, subglabrous; black or
pitchy black above, rufo-piceous beneath, the antennze and tarsi ferruginous or ruto-
testaceous, the rostrum sometimes rufescent in §. Rostrum (¢) much shorter than the
prothorax, stout, curved above and tapering towards the apex (when viewed in profile),
CHAMPION—COLEOPTERA ; CURCULIONID At 435
rugosely punctate ; (?) slender, curved, cylindrical, the apical portion shining and finely
punctate. Prothorax longer than broad, somewhat rounded at the sides, gradually
narrowed from the middle forwards, feebly constricted towards the apex; closely, finely
punctate. Elytra long, at the base very little broader than the prothorax, gradually
widened to beyond the middle and obliquely narrowing thence to the apex; closely,
finely punctate-striate, the interstices very narrow and rugulose. Beneath shining,
closely, rather coarsely punctate ; ventral segments 1 and 2 deeply excavate down the
middle in ¢. Legs short; femora unarmed; tarsal joints 2 and 3 very broad. Wings
wanting.
Length (exclud. head) 14—1% mm. ($9).
Loc. Seychelles: Mahé.
Fourteen specimens, found on Jan. 16th, 1909, in the dwarf forest-vegetation
(containing many Pandani) on the top of Mount Sebert, at an elevation of about
2000 feet. P. sebertensis may be described as an apterous form of P. alatus. The
scutellum is not visible in either of them, and the humeri are similarly shaped.
PHGNICOBATOPSIS, n. gen.
Rostrum deflexed, broad and widened outwards in both sexes, longer than wide, the
scrobes obliquely descending before the eyes, the latter somewhat prominent, the antennz
inserted at about the middle; mandibles broad, minutely dentate at tip; antennze slender,
the scape and joint 1 of the funiculus strongly clavate at the tip, joints 3—7 of the
funiculus short, 2 longer, the club acuminate-ovate, annulate; head not very deeply
inserted into the prothorax; prothorax long, oval, cucullate at the apex above and
arcuate-emarginate beneath; scutellum not visible; elytra oval, connate; anterior and
intermediate coxee moderately, the posterior coxe widely, separated ; ante-coxal portion
of the prosternum short; metasternum very short, the episterna almost covered by the
elytral margins; femora unarmed ; tibize armed with a short hook at the apex; tarsal
joints 2 and 3 very broad, the claws minute and divergent ; body oblong-obovate, setose ;
wings wanting.
Type, P. echinatus.
Two species are referred to this genus, both of which appear to be fairly common in
the forests on the mountains of Mahé. The peculiar form of the rostrum, which is short,
broad, and widened outwards in both sexes, the very short metasternum, and the oval
elytra, separate the present genus from Phanicobates, some of the species of which are
apterous. These insects may be attached to Northea, a dicotyledonous tree.
50. Pheenicobatopsis echinatus, n. sp. (Pl. 28, figs. 28, 28 a.)
Oblong-ovate, rather convex, subopaque ; ferruginous or obscure ferruginous, with
the sides of the prothorax and elytra, and the under surface in great part, piceous or
black, the femora and tibiz, the apex of the antennal scape, and the funiculus usually
more or less infuscate ; the upper surface, femora, and tibize clothed with scattered long,
erect, rigid, blackish setee. Rostrum stout, considerably longer than the width of the
SECOND SERIES—ZOOLOGY, VOL. XVI. 56
436 PERCY SLADEN TRUST EXPEDITION
head behind the eyes, rugosely punctate. Prothorax longer than broad, slightly rounded
at the sides, constricted beyond the middle, a little narrower at the apex than at the
base; densely, finely, asperate-punctate. Elytra much broader than the prothorax,
oval; closely punctate-striate, the interstices rather convex, asperate-punctate. Beneath
shining, closely, coarsely punctate; first ventral segment obsoletely depressed down the
middle in ?. Tarsal joints 2 and 3 exceedingly broad.
Length (exclud. head) 14;—2 mm. (99).
Loc. Seychelles: Mahé, Silhouette.
Sixty specimens, all but one from Mahé. The special localities for it on that
island are: summit of “Montagne Anse Major,” the forest between Trois Fréres and
Morne Seychellois, and the Cascade Estate, at elevations between 1500 and 2000 feet.
The one specimen from Silhouette was found at the Mare aux Cochons.
The males are apparently a little narrower than the females. Dark examples
have the funiculus and the thickened portion of the antennal scape infuscate, the club
remaining pale. The oval elytra and the numerous long, erect, stiff, blackish setze, which
arise from minute asperities of the surface, readily distinguish the present species.
51. Phanicobatopsis seticeps, n. sp.
Oblong-ovate, narrow, opaque; ferruginous, the sides of the prothorax and elytra
piceous or black, the antennal club, head, femora, and the under surface in part, often
more or less infuscate ; clothed with a few minute, widely scattered, hair-like, decumbent,
whitish scales, these becoming longer, coarser, and setiform towards the sides and apex of
the elytra, the head, rostrum, anterior portion of the prothorax, femora, and tibize set with
fine, scattered, erect sete. Rostrum stout, short, not longer than the width of the head
behind the eyes, rugosely punctate. Prothorax longer than broad, convex, rounded at
the sides, constricted beyond the middle, a little narrower at the apex than at the base ;
densely, finely punctate. Elytra wider than the prothorax, oblong-oval; closely
punctate-striate, the interstices somewhat convex and rugulose. Beneath shining,
coarsely punctate, the punctures finer and more scattered in ?; first ventral segment
slightly depressed down the middle in ¢.
Length (exclud. head) 14—15 mm. (72).
Loc. Seychelles: Mahé.
Twenty-eight specimens. This insect, at first sight, would appear to be a small,
narrow, abraded form of P. echinatus, but such is not the case. The erect setz, when
present, are confined to the head and the front of the prothorax ; the elytra are narrower
and less ovate, and the short sete towards the apex are white and decumbent; the
rostrum is shorter; and the minute asperities of the upper surface are wanting. The
general coloration is similar in the two species. The localities for P. seticeps are: Mare
aux Cochons district, and the forest of rather stunted Capucin trees (Vorthea) on the
summit of the adjacent ‘“‘ Montagne Anse Major” (ca. 2000 feet); one specimen is from
the forest above Cascade Estate.
CHAMPION—COLEOPTERA ; CURCULIONIDA A437
PHG@NICOGENUS, h. gen.
Rostrum deflexed, short, broad, not longer than wide, a little narrowed towards the
tip, the scrobes obliquely descending to beneath the small eyes, the antennz inserted near
the apex ; antennze slender, the scape and joint 1 of the funiculus strongly clavate at the
tip, joints 3—7 of the funiculus short, equal, 2 a little longer, the club acuminate-ovate,
annulate, nearly as long as the scape ; head deeply inserted into the prothorax ; prothorax
and elytra truncate at the base, the prothorax cucullate at the apex above and with the
cephalic cavity beneath extending backwards to very near the anterior cox; scutellum
not visible; anterior and intermediate coxee narrowly separated ; metasternum barely as
long as first ventral segment, the narrow episterna exposed ; legs short, femora unarmed,
tibiee without apical hook, tarsi with joints 2 and 3 very broad, the claws minute and
divergent ; body oblong-ovate, variegated with patches of adpressed scales.
LIBRARY|=s0
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Type, P. inermis.
The minute insect forming the type of this genus is, so far as at present known,
peculiar to the Island of Praslin. The stout, broad, subparallel-sided rostrum, the oblong-
ovate body, the variegate upper surface, and the unarmed apices of the tibize, distinguish
it at once from all the other members of the group. The species is doubtless apterous.
No sexual characters are apparent in the three specimens before me.
52. Pheenicogenus inermis, n. sp. (Pl. 28, figs. 29, 29 a.)
Rather convex, shining (when denuded), ferruginous or obscure ferruginous, the
antennee rufo-testaceous ; variegated, the rostrum included, with dense, irregular, subcon-
fluent patches of narrow, adpressed, minute, flavo-cinereous scales, which become longer
and curled on the anterior portion of the prothorax. Rostrum bare at the tip, and, like
the head, densely punctate. Prothorax as broad as long, rounded at the sides, truncate at
the base, much narrowed anteriorly ; densely, finely punctate. Elytra connate, a little
wider than the prothorax, moderately long, subparallel at the base, somewhat produced at
the tip, the humeri obtuse; coarsely punctate-striate, the interstices rugulose. Beneath
closely, somewhat coarsely punctate.
Length (exclud. head) 14, 1,9 mm.
Loc. Seychelles: Praslin.
Three specimens, found on Noy. 28th and 29th, 1908, on the Cotes d’Or Estate, in
the Coco-de-mer forest in the Vallée de Mai.
Group Anthonomina.
ENDOPSIS, n. gen.
Rostrum curved, moderately stout, larger than the head and prothorax, the antennze
inserted beyond or at the middle, the scrobes deep and obliquely descending to beneath
the eyes; head large, inserted into the prothorax to near the eyes, the latter rounded,
prominent, and separated by less than the width of the rostrum; antenne with an
elongate-ovate, 3-jointed, pubescent club and a 6-jointed funiculus, jot 1 of latter stout,
56—2
438 PERCY SLADEN TRUST EXPEDITION
obovate, the others small; prothorax transverse, small; scutellum very small; elytra broad,
oblong or oval; anterior coxee contiguous, strongly exserted ; metasternum moderately
long, the episterna exposed ; ventral segments free, 2 intermediate in length between 1 and
3, 3—5 subequal, the pygidium ventrally exposed, the sutures straight ; femora strongly
clavate, 1 usually stouter than the others, all sharply dentate; tibize armed with a small
hook at the outer apical angle, the anterior pa bowed, more or less sinuate within ;
tarsi with joint 3 bilobed, spongy-pubescent beneath, the claws appendiculate ; body
oblong or oval, rather convex, pubescent or squamose, winged.
Type, EL. curvimanus.
The three species placed under this genus must come near Hndaus, Schénh., the
type of which was from Sierra Leone, and which, according to Lacordaire, also has
a 6-jointed funiculus. The Australian genera Diapelmus, Orpha, and Myossita, the New
Zealand Hugnomus (as identified by Pascoe), and the Central American Udeus, all have
similarly clavate, sharply dentate femora; but in these forms the head is more exserted
and the eyes, in consequence, are more distant from the prothorax, and the tarsal claws
(except in Diapelmus) are simple. Hnd@opsis superficially resembles Rhynchenus (Orchestes),
the type, H. curvimanus, being so like the immaculate form of R. crassipes that the
two insects were placed together in the Seychelles collections sent me for study.
53. Endeopsis curvimanus, n. sp. (Pl. 28, fig. 30, 2.)
Oblong, rather convex, shining, ferruginous or obscure ferruginous, darker beneath,
the antennee (the scape excepted) infuscate, the eyes black ; finely pubescent, the elytral
interstices each with a row of longer decumbent hairs. Head densely punctulate ; eyes
large, prominent, separated by about half the width of the rostrum; rostrum moderately
stout, curved, slightly longer than the prothorax, rugulosely punctate, in ? longer,
smoother, and sparsely, finely punctate, the antennze inserted considerably beyond the
middle in g, and further from the apex in ?. Prothorax small, transverse, rounded at the
sides, feebly constricted and much narrowed anteriorly ; densely punctate. Elytra broad,
subparallel at the base, slightly widened to about the middle, conjointly rounded at the
apex ; punctate-striate, the interstices closely punctulate and feebly convex. Beneath very
sparsely, finely punctate. Anterior femora more strongly clavate, and also more strongly
dentate, than the others, and finely ciliate between the tooth and the apex. Anterior
tibize bowed, strongly sinuate within, and arcuately dilated on the imner side towards
the tip.
Length (includ. head) 24—31 mm. (#2).
Loc. Seychelles: Silhouette, Mahé.
Nearly 40 specimens, beaten from certain dicotyledonous trees and bushes in the
mountain-forests. Silhouette; from near Mont Pot-d-eau and from the Mare aux
Cochons. Mahé; high forests of Morne Blanc and Pilot, high forest between Trois Freres
and Morne Seychellois, forest above Cascade, and the Mare aux Cochons district.
54. Hndeopsis anthonomoides, n. sp.
Oblong, convex, shining, ferruginous, the eyes black ; clothed with narrow, hair-like,
yellowish: scales, those on the elytra condensed into two irregular rows down each interstice.
CHAMPION—COLEOPTERA ; CURCULIONIDA 439
Head densely punctulate; rostrum curved, about as long as the head and prothorax,
rugulosely punctate, the antenne inserted at the middle ; eyes moderately large, separated
by a little less than the width of the rostrum. Prothorax transverse, rounded at the
sides, narrowed and very feebly constricted anteriorly, densely punctate. Elytra broad,
subparallel in their basal half, conjointly rounded at the apex; punctate-striate, the
interstices convex and punctulate. Legs comparatively short; anterior femora more
strongly clavate, and also more sharply dentate, than the others, and finely ciliate beyond
the median tooth; anterior tibiae widened towards the apex, thus appearing sinuate
within.
Length 13 mm. (¢ 7%).
Loc. Seychelles: Silhouette.
One specimen, taken at the Mare aux Cochons, Sept. 1908: Smaller and more
convex than H. curvimanus, the eyes not so large and less approximate above, the anterior
tibize less curved, the vestiture squamiform. The insect has exactly the appearance of
a small Anthonomus.
55. Hndaopsis delicatus, n. sp.
Oval, shining, black, the legs and antenne ferruginous ; clothed with an extremely
fine cinereous pubescence, and also set with scattered, longer, decumbent hairs, these
latter condensed into a single line down each elytral interstice. Head, prothorax, and
elytra very densely punctulate ; rostrum arcuate, slightly longer than the prothorax,
stout, widening a little outwards, rugulosely punctate and carinate, the antennz inserted
beyond the middle; eyes moderately large, separated by considerably less than the width
of the rostrum; prothorax transverse, rounded at the sides, narrowed and very feebly
constricted anteriorly; elytra broad, oval, finely punctate-striate, the interstices broad
and almost flat. Beneath very sparsely, finely punctate. Legs short ; femora strongly
clavate and sharply dentate; anterior tibiz feebly sinuate.
Length 2% mm. (? ?).
Loc. Seychelles: Silhouette.
One specimen, from Mare aux Cochons, taken in Aug. 1908. The oval, black body,
the very delicate cinereous pubescence, with a row of conspicuous longer hairs down each
elytral interstice, the basally narrowed rostrum, and the relatively larger prothorax, give
this insect a different appearance from its two allies.
RHAYNCH ZNUS.
Rhynchenus (Clairville], Ent. Helv., i. p. 70 (1798).
Orchestes Illiger, Mag., ui. p. 105 (1804).
A holarctic genus, but with a few representatives in warmer climes. The larvee are
leaf-miners. One species (Orchestes basipictus, Fairm., 1897) has been described from
Madagascar.
440 PERCY SLADEN TRUST EXPEDITION
56. Rhynchenus spissus, n. sp. (Pl. 28, figs. 31, 31 a, 9.)
Robust, dull; varying in colour from ferruginous to piceous, the rostrum in part or
entirely, the antennze and legs (one or two dark annuli on the posterior femora excepted)
ferruginous, the elytra in light-coloured examples more or less angulate-fasciate or spotted
with black (the dark markings sometimes reduced to a small oblong spot before the middle
of the suture) ; variegated with a dense clothing of flavo-cinereous and blackish (or dark
brown) pubescence, the light and dark hairs usually condensed into one or more interrupted
angulate fasciz on the elytra, the cinereous hairs in dark individuals reduced to a few
small patches; the prothorax and base of the elytra also set with scattered, long, pro-
jecting sete. Head densely punctulate; eyes very large, contiguous above; rostrum
strongly deflexed, moderately stout, curved, in $ reaching beyond the intermediate coxee,
shorter in 2, shining, punctate-striate, rugose at the base; antennz inserted far behind
the middle, the funiculus setose, 6-jointed, joints 2 and 3 elongate, slender, 4—6 shorter,
the club loosely 3-jointed. Prothorax short, the sides rounded to the abruptly separated
narrow anterior lobe, densely punctulate. Elytra broad, somewhat rounded at the sides
and blunt at the tip; narrowly punctate-sulcate, the mterstices broad, somewhat convex,
densely punctulate. Beneath shining, closely, finely punctate. Legs stout; femora
unarmed, the posterior pair greatly incrassate.
Length 22—23 mm. (f9).
Loc. Seychelles: Silhouette, Mahé.
Silhouette, from Mare aux Cochons, ix. 1908. Mahé, from near Morne Blane,
ca. 800—1000 feet, and from Cascade Estate and the forests above: also several
specimens from an islet near Mahé, Anonyme Island, 1. 1909. A long series, varying
greatly in colour and in the development of the black angulate fascize on the elytra, which
may be obsolete in light-coloured individuals or completely lost in darker examples.
R. spissus bears some resemblance to the European fF. loncere ; but it is a broader and
more robust insect with stouter legs. The specimens with the longer rostrum are assumed
to be females.
Group Cryptorrhynchina.
The nine Cryptorrhynchids contained in the collection under examination belong
to nearly as many genera, several of which are probably endemic. More species of the
apterous genus Acalles (so well represented in many Oceanic islands) might have reason-
ably been expected to occur in the Seychelles. In addition to the forms before me, an
introduced Camptorrhinus has been recorded from Mahé.
ACALLES.
Acalles Schénherr, Cure. Disp. Meth., p. 295 (1826); Gen. Cure., iv. p. 325 (1837).
A genus of world-wide distribution. The single remarkably distinct species now
described from the Seychelles is certainly endemic.
CHAMPION—COLEOPTERA ; CURCULIONIDA A441
57. Acalles seychellensis, n. sp. (PI. 23, figs. 32, 32a, ¢.)
Acuminate-ovate, very convex, opaque, the summits of the tubercles, the under
surface, and the rostrum of 9, shining; black, the antenne and tarsi obscure ferruginous,
the depressed portions of the surface bearing minute, scattered, yellowish, setiform scales,
the legs also setulose. Head confluently foveolate ; rostrum of ¢ similarly sculptured, and
also tricarinate, broad, widened outwards, and about as long as the posterior portion of
the prothorax, that of ? much smoother in its outer half, the antenne inserted at (3%) or
behind (?) the middle; eyes large. Prothorax broader than long, rounded at the sides,
which are parallel at the extreme base (as seen from above), the short anterior lobe
obliquely narrowing forwards; closely studded with intermixed conical and oblique
tubercles. Elytra subcordate, acuminate, broader than the prothorax, transversely gibbous ;
with rows of inconspicuous scattered punctures, the interstices convex and each studded
with a series of rather large oblong or conical tubercles, these latter becoming smaller and
more distant on the steep apical declivity. Beneath confluently foveate, the transverse
arcuate ridge on the mesosternum very prominent ; ventral segments 3 and 4 extremely
short. Legs rugosely punctate, the tibiz carinate.
Length 3—34, breadth 12—24 mm. (9).
Loc. Seychelles: Mahé, Silhouette.
Fourteen specimens, found in the forests of the higher country, amongst leaves on
the ground: Mahé, high forest of Morne Blanc and forest above Cascade Estate:
Silhouette, from near Mont Pot-a-eau and from the forest above Mare aux Cochons.
A highly specialized form, black, closely studded above with conical or oblong tubercles,
which are polished at their summits. The presence of this sluggish apterous insect on two
separate islands shows that they must have been formerly connected.
CAMPTORRHINUS.
Camptorhinus Schénherr, Cure. Disp. Meth., p. 283 (1826).
A genus represented in 8. Europe, Africa, Madagascar, Ceylon, Java, Sumatra, &c.
58. Camptorrhinus brullet.
Camptorhinus brullet Boh., in Schénh., Gen. Cure., iv. p. 174; Kolbe, Mitteil. Zool.
Mus. Berlin, v. p. 45 (1910).
Camptorhinus dorsiger Fairm., Ann. Soc. Ent. France, 1871, p. 53.
Loc. Seychelles: Mahé (A. Brauer). Madagascar.
An introduced insect in Mahé. C. dorsiger, Fairm., a co-type of which belonging to
the Paris Museum is before me, is a form of C. brullei with the tubercles on the alternate
elytral interstices larger, the terminal one on the fifth being especially conspicuous. Both
insects inhabit Madagascar.
442, PERCY SLADEN TRUST EXPEDITION
CRYPTORRHYNCHUS.
Cryptorhynchus Uliger, Mag., vi. p. 330 (1807).
An immense number of heterogeneous forms from all parts of the world are at present
referred to this genus. The two included in it in this paper are certainly not endemic, and
it is therefore unnecessary to say much about them.
59. Cryptorrhynchus mangifere.
Curculio mangifere Fabr., Syst. Ent., p. 139 (1774).
Cryptorhynchus mangifere Boh., in Schonh., Gen. Cure., iv. p. 91; Van Dine, Proc.
Hawai. Ent. Soc., 1. pp. 79—82 (1907).
Cryptorhynchus ineffectus Walk., Ann. Mag. Nat. Hist., (3) ii. (1859), p. 264.
Loc. Chagos Islands. Madagascar. Mauritius. Réunion. Java. India. Ceylon.
Africa; Zanzibar, Natal, &e. Hawaiian Islands.
One specimen, labelled ‘‘ Chagos Islands, flew on board, 3. vu. 05.” A cosmopolitan
insect, known as the “‘mango-weevil,” and doubtless often transported to distant places in
the fruit. The type of C. mangifere was found in a mango-nut. Walker's C. ineffectus
came from Ceylon. The insects in the British Museum representing C. humeralis, Gyll.,
and C. australis, Boisd., belong to the same species, but their identification is doubtful.
Two allied forms were described by Fairmaire in 1897 from Madagascar, C. transversus
and C. angustior. According to Emmerez (Revue Agricole, 30 Juillet, 1898), C. mangifere
is a destructive insect in Mauritius. The habits of the insect in India have been described
by Simmons in 1888, and by Cotes in 1889.
60. Cryptorrhynchus erratus, n. sp. (PI. 24, figs. 33, 33 a, 9.)
Oblong-ovate, convex, black, the antennze and the apices of the tarsi ferruginous ,
variegated with a dense clothing of intermixed broad, adpressed, and imbricate and
erect, narrower, brown, whitish, and black scales—the whitish adpressed scales condensed
into a subquadrate or transverse patch on the disc of the elytra before the middle
(mainly formed by a streak on the fourth and fifth interstices) and various small scattered
spots, and the black scales into some small spots on the prothorax and various short
streaks on the elytral interstices 2, 3, and 5 (the streak at the base of 3 and 5 conspicuous,
that on 2 sometimes wanting), the erect dark scales on the prothorax somewhat crowded
at the apex and across the middle, the legs faintly fusco-annulate. Head densely, rugosely
punctate above, transversely strigose beneath, the vertex with a short ridge (forming
a continuation of the rostral carina); eyes large, coarsely facetted ; rostrum long, stout,
strongly arcuate, reaching to between the middle coxz, carinate in its basal half, coarsely
punctate to the tip in ¢, the apical half much smoother in 2; antennal club about as long
as the four preceding joints united. Prothorax transverse, strongly rounded at the sides,
narrow and constricted at the “apex, bisinuate at the base; very coarsely, confluently
punctate. Scutellum transverse. Elytra slightly wider than the prothorax, moderately
elongate, narrowing from a little below the base ; punctate-striate, the interstices densely
punctate, 3 and 5 somewhat raised. Beneath densely, coarsely punctate; rostral canal
CHAMPION—COLEOPTERA ; CURCULIONIDA 443
very deep, terminating in the greatly developed, U-shaped, horizontal mesosternal process ;
ventral segments 2—4 subequal in length ; femora feebly bidentate, suleate along their
entire length beneath ; tibize broad, somewhat curved.
Length 51—8, breadth 24—3# mm. (f?).
Loc. Aldabra: Takamaka, 1908 (Fryer). Port Natal (Plant, in Mus. Brit.). Guinea,
River Niger (Mus. Brit.).
Described from a pair from Guinea and a female from each of the other localities ;
the one from Natal, in the Fry collection, found by Mr Plant, is taken as the type. This
insect must have been introduced in some way into Aldabra, it being evidently a very
widely distributed African form, possibly already described? C. erratus will doubtless
have to be removed from Cryptorrhynchus when the African Cryptorrhynchids are properly
studied. The W. African specimens are larger than the others and have some additional
black spots on the elytra. The Aldabra example is figured.
CRYPTORRHYNCHIDIUS, n. gen.
Rostrum moderately stout, arcuate, in repose reaching as far as the middle of the
intermediate coxee; rostral canal very deep, closed behind by the greatly developed,
U-shaped mesosternal process, which is on the same plane as the anteriorly emarginate
metasternum, the latter short and with broad episterna; antennz inserted near the
middle of the rostrum, the club oblong; scutellum prominent, polished; elytra deeply
sinuate at the base, the sides forming an almost continuous outline with those of the
basally widened prothorax; ventral segment 2 nearly as long as 3 and 4 united, 3 and 4
short, 1 elongate; intermediate and posterior coxee somewhat widely separated ; femora
feebly dentate, sublinear, the anterior pair longer than the others ; third tarsal joint
broadly bilobed; tarsal claws simple, divergent; body robust, very convex, laterally
compressed, acuminate-ovate, densely squamose.
Type, C. graniger.
Amongst the vast number of Cryptorrhynchids known to me I have not come across
any species with which C. graniger could be satisfactorily compared, and it must there-
fore be placed under a separate generic name for the present. Cryptorrhynchus, type
C. lapathi L., has the head transversely strigose beneath, the ventral segments 2—4
subequal in length, the elytra much wider than the prothorax, &c.
61. Cryptorrhynchidius graniger, n. sp. (Pl. 24, figs. 34, 34a, ?.)
Sphadasmus granocostatus Kolbe* (nec Fairm.), Mitt. Zool. Mus. Berlin, v. p. 45
(1910).
Black, the extreme base of the antennz red; variegated with a dense clothing
of rounded, imbricate, brown (or greyish) and white scales, the latter tending to form
irregular fascize on the elytra, the scattered granules along each elytral interstice bare
* T have examined one of the specimens determined by Kolbe as S. granocostatus Fairm., and find
that it is not a Sphadasmus, but is identical with C. graniger.
SECOND SERIES—ZOOLOGY, VOL. XVI. 57
444 PERCY SLADEN TRUST EXPEDITION
and polished, the large shallow punctures on the prothorax each bearing a forwardly-
directed, adpressed, setiform scale and the granules on the elytra each followed by a
similar posteriorly-directed scale, the legs also with intermixed setiform scales. Head
sparsely punctate beneath; eyes large ; rostrum curved, moderately stout, slightly
widened towards the base, more strongly so in ?, somewhat flattened, in ¢ a little longer
than the prothorax and sparsely, finely punctate, in $ almost smooth to near the base and
with the apical portion much more elongate, the antenne in $ inserted at a little before,
and in @ considerably behind, the middle; antennz sparsely setose. Eyes separated by
less than the width of the narrowest portion of the rostrum. Prothorax transverse,
gradually, arcuately narrowing from the base to the abrupt deep subapical constriction,
the sides angulate anteriorly and posteriorly, forming an almost continuous outline with
those of the elytra, the base deeply bisinuate ; the surface impressed with large shallow
foveiform punctures, which are hidden by the vestiture. Elytra rounded at the sides and
acuminate at the tip ; deeply punctate-striate, the interstices broad, convex, each bearing
a scattered series of polished granular elevations. Beneath very coarsely, closely punctate,
the first ventral segment unimpressed in both sexes. Femora hollowed towards the apex
beneath, so as to appear feebly unidentate. Anterior tibize densely ciliate within in ¢.
Length 6—10 mm. (¢9).
Loc. Seychelles: Mahé, Silhouette, Praslin.
One male and five females, apparently picked up singly, varying greatly in size and
in the colour of the scales, which in one example are almost wholly brown and in another
in great part white. In Mahé Mr Scott met with the insect near Morne Blanc and on the
Cascade Estate, and in Silhouette he found one specimen on the marshy plateau of
Mare aux Cochons; several of these were obtained by beating trees. My Gardiner had
previously, in 1905, captured an example on Praslin. This species agrees very nearly with
the description of Sphadasmus granocostatus, Fairm., from the Seychelles ; but as
Fairmaire definitely states that the femora are unarmed and the prosternum is not
eanaliculate, it is obviously not that insect. Kolbe’s specimen was found by A. Brauer
in Mahé.
RHETOGENES, n. gen.
Rostrum arcuate, rather stout, subcylindrical, in repose reaching as far as the
posterior margin of the middle coxe; rostral canal deep, squamose within, closed behind
by the metasternum, which is hollowed beneath the angularly notched anterior margin ;
prosternum very deeply emarginate in front; mesosternum depressed ; antennze inserted
at or beyond the middle of the rostrum, slender, the club ovate; prothorax much
narrower than the elytra; scutellum depressed; elytra broad, oblong; ventral segment 1
much longer than the metasternum, 2 a little shorter, 3 and 4 very short, the first suture
strongly sinuate; posterior coxee widely, the other coxee more narrowly, separated ;
femora dentate or unarmed beneath; tibiee sinuate; third tarsal joint bilobed ; tarsal
claws simple; body oblong, squamose.
Type, R. sexcristatus.
CHAMPION—COLEOPTERA ; CURCULIONIDA 445
The insect selected as the type of this genus is not unlike Cryptorrhynchus lapathi,
but differs from it in the strongly depressed mesosternum (the V-shaped mesosternal process
being entirely absent, and the rostral canal closed by the metasternum), the relatively long
second ventral segment, &c. The smaller second species referred to Rhetogenes has similar
sternal and abdominal characters.
62. hetogenes sexcristatus, n. sp. (Pl. 24, figs. 35, 35a, 3g.)
Rather broad, robust, black or piceous, the antennee, rostrum, and tarsi sometimes
rufescent ; variegated with a dense clothing of pale brown, brownish-black, and whitish
imbricate scales, the dark scales on the prothorax condensed into a triangular patch on the
basal half of the disc, and the whitish scales on the elytra clustered into various curved
or sinuate transverse streaks on the apical declivity, the prothorax and elytra also with
various fascicles of stout, erect, setiform, black and brown scales-—the prothorax with four
in a transverse row across the middle (the inner two black) and two at the apex, and the
elytra with several along each alternate interstice, the fascicles becoming larger towards
the base and smaller towards the apex, the legs annulate and setose. Head transversely
strigose beneath ; eyes large, coarsely facetted ; rostrum closely punctate, the apical
portion a little smoother in ?, the antennz inserted at a little beyond the middle.
Prothorax transversely subquadrate, constricted and abruptly narrowed anteriorly,
quadrinodose across the middle, the sides parallel in their basal half; densely punctate.
Elytra broad, oblong-subquadrate, broadly produced at the apex; punctate-striate, the
interstices broad, densely, finely punctate, 3, 5, and 7 with scattered setigerous promi-
nences. Beneath dull, coarsely, closely punctate; metasternum short, flattened, not
suleate ; first ventral segment unimpressed in both sexes. Femora dentate beneath.
Length 43—51, breadth 21,—22 mm. (79).
Loc. Seychelles: Silhouette, Long Island.
Three specimens from the Mare aux Cochons, Silhouette, 1X. 1908, and one from
Long Island (a small islet close to Mahé), VII. 1908, scarcely varying inter se.
63. Rhetogenes spurcus, n. sp.
?. Oblong, nigro-piceous, the antennze and the tips of the tarsi ferruginous ;
variegated with a dense clothing of brown and blackish scales, the brown scales condensed
into an interrupted median line on the prothorax, a patch on the scutellum, and various
faint spots along the alternate elytral interstices, the upper surface and legs also set with
numerous, intermixed, erect, stout, blunt, setiform scales. Rostrum about as long as the
prothorax, densely punctate, the apical half smoother and shining, the antennze inserted
at the middle. Head densely punctured above, strigose beneath ; eyes rather small, finely
facetted. Prothorax a little broader than long, abruptly narrowed from about the middle,
densely punctate, without definite prominences. Elytra much wider than the prothorax,
subparallel in their basal half; finely punctate-striate, the strize sinuate on the disc, the
interstices broad, densely punctate, the alternate ones here and there faintly nodulose.
Beneath densely punctate. Femora without definite tooth.
Length 34, breadth 14 mm.
57—2
446 PERCY SLADEN TRUST EXPEDITION
Loc. Seychelles: Silhouette, VIIT. 1908.
One specimen. A worn male from the Cascade Estate, Mahé, with more slender
setee, may belong here. A. small form, with depressed mesosternum, fine, sinuous elytral
strize, nodulose alternate interstices, and stout, blunt setze on the upper surface and legs.
The prothorax is without definite prominences.
BARIDIOMORPHA, nh. gen.
Rostrum parallel-sided, feebly curved, moderately stout, in repose reaching as far as the
posterior margin of the anterior cox ; rostral canal very deep, closed behind by the arcuate
mesosternal process, the latter raised above the level of the metasternum and narrowly
extending backwards between the middle cox ; metasternum long, the intercoxal process
narrow, truncate in front; antennee inserted at above the basal third or fourth of the
rostrum, slender, short, the club ovate; mandibles bifid at the tip; prothorax as long as
broad, narrower than the elytra, cucullate in front, deeply bisinuate at the base; scutellum
small, depressed ; elytra elongate, subparallel; ventral segment 2 nearly as long as 3 and
4 united ; intermediate coxe narrowly, the posterior coxee somewhat widely, separated ;
legs rather short, stout; femora unarmed, clavate; tibize short, sinuate within, the
intermediate pair toothed at the outer apical angle; tarsi narrow, joint 3 bilobed,
pubescent beneath, the claws simple, slender, free; body elongate, narrow, densely
squamose.
Type, B. triplaris.
The type of this Cryptorrhynchid genus has the general facies of an elongate, narrow,
densely squamose Barid. It is unknown to both Dr Heller and myself, and I cannot
suggest any nearly allied form.
64. Baridiomorpha triplaris, n. sp. (Pl. 24, figs. 36, 36a, 3.)
Rather convex, black, the tarsi piceous; variegated with a dense clothing of brown,
white, and black imbricate scales—the black scales condensed into various irregular
coalescent markings on the prothorax and elytra, and the white scales into a large patch
on each side of the prothorax at the base and numerous small scattered spots—intermixed
with semi-erect, curled, blunt setze, the setze on the elytra uniseriately arranged down each
interstice. Head closely punctate beneath; eyes large; rostrum (%) closely punctate to
the apex, (?) a little longer and smoother, densely squamose at the base. Prothorax
somewhat rounded at the sides to the narrower, subtubulate, anterior portion, densely
punctate. Elytra subparallel, broadly produced at the apex; conspicuously punctate-
striate, the interstices flat and densely punctate. Beneath densely, finely punctate, the
metasternum and first two ventral segments with coarse punctures intermixed; meta-
sternum sulcate down the middle posteriorly, broadly depressed down the middle in ¢.
Length 34—4 mm. (9).
Loc. Seychelles: Mahé, Silhouette.
Twelve examples, all males but two, showing very little variation. Found in
Silhouette in the high country near Mont Pot-a-eau and at the Mare aux Cochons, and
in Mahé in the high forest of Morne Blane and Pilot.
CHAMPION—COLEOPTERA ; CURCULIONIDA 447
RHOMBOSOMA, n. gen.
Rostrum arcuate, widened at the base, moderately long, the antennze inserted at
about the middle; eyes very large, lateral, coarsely facetted; rostral canal very deep,
closed by the prominent, horizontal, U-shaped mesosternal process; metasternum short,
the episterna broad, angularly dilated anteriorly; ventral segment 2 nearly as long as
3 and 4 united, the sutures shallow, the first arcuate; prothorax cucullate in front,
subtriangular ; scutellum small, prominent ; elytra subtriangular ; femora linear, hollowed
towards the apex; tibize rounded at the outer apical angle; third tarsal joint broadly
bilobed, the claws free; body rhomboidal, convex, laterally compressed, squamose,
winged.
Type, R. acuminatum.
The type of this genus approaches the American Pseudomus and Pseudomopsis,
differing from them in having broadly exposed metathoracic episterna, the wings fully
developed, &c. It has nothing to do with the somewhat similarly shaped Lobotrachelus
from 8. Africa, &e., which have approximate exposed eyes, these insects being nearly
related to the Zygopina. &. acwminatwm may be an introduced American form in
Silhouette ?
65. Rhombosoma acuminatum, n. sp.
Moderately elongate, piceous, the antenne and tips of the tarsi ferruginous ;
variegated with a rather dense clothing of brown, fuscous, and (a few) white scales, the
pale brown scales on the elytra condensed into an oblique fascia on the disc before the
middle and the white scales into a small spot on the second interstice towards the apex,
the surface also set with a few short, curled, adpressed setze, the legs annulate. Head
densely punctate above, sparsely so beneath; rostrum closely punctate and squamose at
the base, bare, shining, and much smoother in its apical half. Prothorax broader than
- long, obliquely narrowed forwards, slightly constricted near the apex, bisinuate at the base;
closely punctate, much smoother and alutaceous on the middle of the disc. Elytra a little
wider than the prothorax, rapidly, obliquely narrowing from a little below the base, the
humeri rounded; deeply punctate-striate, the interstices rather convex and minutely
punctate. Beneath coarsely punctured, the punctures on the ventral segments 3 and 4
reduced to a single transverse row. Legs rather long and stout, the anterior femora only
with a minute tooth.
Length 44 mm. (@ 2).
Loc. Seychelles: Silhouette.
One specimen, found near Mont Pot-a-eau, at about 1000 feet, VIII. 1908. It has
the scales on the prothorax somewhat abraded, the dise thus appearing to have a large
triangular dark patch.
ScYTHA, n. gen.
Rostrum cylindrical, feebly curved, deflexed, received in repose between the anterior
and middle coxee, the antennz inserted near the middle, jomts 2—7 of the funiculus short,
widening outwards, the club large and nearly as long as the funiculus; rostral canal
448 PERCY SLADEN TRUST EXPEDITION
almost obsolete; prosternum broad, feebly hollowed down the middle, very deeply
arcuate-emarginate in front, and with the ante-coxal portion reduced to a narrow strip ;
mesosternum flattened, large, the meso-metasternal suture transverse, placed midway
between the intermediate coxe ; ventral segment 2 a little longer than 3 or 4; prothorax
cucullate at the apex; scutellum small; elytra oblong; all the coxz widely separated ;
legs long, stout ; femora toothed; tibize strongly unguiculate ; tarsi with the third joint
bilobed, the first jot of anterior pair elongated, the claws stout, free ; body oblong,
squamose.
Type, S. aldabrana.
The minute insect from which the above definition has been taken may be described
as an aberrant Cryptorrhynchid with the rostral canal almost obliterated, the coxze widely
separated, the antennal club large, and the anterior tarsi elongated. The unique type
may be a male, as it has the anterior tibize angularly dilated within. It is possible that
the species is not a native of Aldabra.
66. Scytha aldabrana, n. sp.
Narrow, moderately convex, piceous, the antennee and the apices of the tarsi
ferruginous ; variegated with a dense clothing of brownish-cinereous, fuscous, and whitish
scales, the dark scales condensed into two vittee on the disc of the prothorax and various
small spots on the alternate elytral interstices, and the whitish scales into a short streak
on the third interstice at about the middle, the prothorax and the small prominences on
the alternate interstices also set with a few very short decumbent blackish setz, the legs
fusco-annulate. Rostrum shorter than the prothorax, densely punctate and squamose at
the base, for the rest sparsely punctured, bare, and shining. Prothorax convex, nearly as
long as broad, rounded at the sides, narrowed and constricted anteriorly, bisinuate at the
base ; densely punctate. Elytra wider than the prothorax, oblong, parallel at the base,
slightly produced at the tip; punctate-striate, the interstices densely punctate, 3, 5, and
7 somewhat raised and here and there subfasciculate with small dark sete. Beneath
densely punctate ; fifth ventral segment arcuate-emarginate at the apex, leaving the sixth
exposed. Anterior tibiz sinuate, angularly dilated towards the middle within. Anterior
tarsi elongated, the first joint longer than that of the other legs.
Length 24, breadth 1 mm, (¢ %).
Loc. Aldabra: Takamaka (Fryer).
One specimen, found in November or December, 1908.
Group Zygopina.
SPHADASMUS.
Sphadasmus Schénherr, Gen. Col., vii. 1, p. 290 (1844); Heller, Abhandl. Zool. Mus.
Dresden, 1892—93, p. 47.
Eight species of this genus are known from Africa or India. S. granocostatus Fairm.,
is probably not really congeneric.
CHAMPION—COLEOPTERA ; CURCULIONIDA 449
67. Sphadasmus granocostatus.
Sphadasmus granocostatus Fairm. (nec Kolbe*), Bull. Soc. Ent. France, 1893,
Pp: Cccxxv.
Loc. Seychelles.
Not represented in the collections before me. The type cannot now be found in
the Fairmaire collection in the Paris Museum.
Group Barina.
Two Barids are contained in the Seychelles collection before me; one of them is
probably an introduced form, but I am unable to identify it, if described; the other
may be endemic.
~ Baris.
Baris Germar, Ins. Spec. Nov., p. 197 (1824) (part.).
Baridius Schénherr, Gen. Cure., iii. p. 643 (1836) (part.).
A genus of world-wide distribution, including a very large number of species, some of
which have been accidentally introduced and established in countries distant from their
original home.
68. Baris multivaga, n. sp.
Oblong, robust, flattened above, shining, nigro-piceous or black, the antennze and legs
in part or entirely, and sometimes the rostrum also, ferruginous or fusco-ferruginous ; the
sides of the prothorax, a common, elongate, X-shaped patch on the basal half of the elytra
(extending outwards to the third stria), a small spot on the shoulders, the mesothoracic
epimera, the prosternum, and metathoracic episterna, somewhat thickly clothed with
coarse, oblong, ochraceous scales, the rest of the under surface, the rostrum, and legs set
with narrower pallid scales, the elytra also with scattered longer, setiform, whitish, and
smaller, inconspicuous, darker scales along each interstice. Rostrum strongly arcuate,
very stout in g, a little more slender in $, not much shorter than the prothorax, coarsely
punctate to the tip, the punctures placed in grooves along the sides, the antennz inserted
at about the apical third; eyes large; antennze stout. Prothorax transverse, the sides
strongly rounded anteriorly and parallel towards the base, the latter deeply sinuate ;
coarsely, closely punctate, except along a narrow smooth space down the middle, the
punctures becoming confluent laterally and the scales on this portion of the surface
transversely arranged. Llytra oblong, a little wider than the prothorax ; with sharply-
cut deep striz, and broad, coarsely, rugosely punctate interstices, the sculpture becoming
more granulate towards the sides. Pygidium coarsely punctured. Beneath coarsely,
closely punctate, the ventral segments more sparsely and more finely punctured down
the middle, broadly and deeply sulcate in 2.
Length 34—33 mm. ($9).
* The specimens determined by Kolbe as S. granocostatus Fairm. are really Cryptorrhynchidius
granuger, q.v. (anted, p. 443). '
450 PERCY SLADEN TRUST EXPEDITION
Loc. Seychelles: Bird Island, 1908 (Fryer). Amirantes: Desroches Island and
Eagle Island, 1905.
Eleven specimens: one found on Eagle Island, on October 17th, 1905, under stones ;
another on Desroches, in the same month; and nine on Bird Island, in July, 1908.
I have seen allied unnamed forms from Amboyna and Mauritius, and B. multwaga,
therefore, if introduced, is doubtless a native of the Malayan islands or adjacent
regions. Certain 8. European members of the genus, too, come rather near it, such as
B. spoliata, Boh.
69. Baris seychellensis, n. sp.
Oblong, subopaque, black; sparsely clothed with minute, narrow, cinereous scales,
which tend to form a faint median vitta on the prothorax, the elytra with a double row
of larger white scales at the base of the third interstice. Rostrum stout, about as long as
the prothorax, abruptly arcuate from the base, separated from the head by a deep
transverse groove, rugosely punctate to the tip, the antenne inserted at the apical third ;
head rugulosely punctured. Prothorax subquadrate, sinuate at the sides towards the
base, abruptly constricted before the apex, tubulate in front, the base deeply bisinuate ;
coarsely, closely punctate. Elytra wider than the prothorax, comparatively short, the
humeri obliquely truncate, becoming tumid laterally ; finely, narrowly striate, the inter-
stices broad, flat, irregularly uniseriate-punctate, and transversely rugose, the sculpture
becoming denser towards the sides. Pygidium narrowly exposed. Beneath shining,
very coarsely, the ventral segments finely and sparsely, punctate ; prosternum arcuate-
emarginate at the apex, and with a deep transverse sulcus in the middle anteriorly ;
anterior cox separated by a little less than their own width; first ventral segment
unimpressed. Legs rugosely punctate.
Length 25 mm. (@ ?).
Loc. Seychelles: Mahé.
One specimen, found on the Cascade Estate, at an elevation of about 1000 feet, in
January, 1909. This species will probably have to be removed from Baris, but it can
remain under that genus for the present. . seychellensis bears a certain resemblance to
some of the numerous American Pseudobaris, but the prosternum is not sulcate for the
reception of the antenne, &c.
Group Cyladina.
CYLAS.
Cylas Latreille, Hist. Nat. Crust. et Ins., 11. p. 196 (1802).
One species of this remarkable genus, the Sweet-potato weevil, has been found in the
Seychelles, where it has certainly been introduced, as in Hawaii, Guiana, &c.
70. Cylas formicarius.
Brentus formicarius Fabr., Ent. Syst., Suppl., p. 174 (1798).
Cylas formicarius Maskew, Monthly Bull. State Comm. Hort., Sacramento, Calif., 11.
pp. 535—587, figs. (1913).
CHAMPION—COLEOPTERA ; CURCULIONIDA 451
Cylas turcipennis Boh., in Schénh., Gen. Cure., i. p. 369 ; Perkins, Fauna Hawaiiensis,
il. p. 131 (1900). .
Loc. Seychelles: Mahé, Praslin. Known from Madagascar, India, Burma, Java,
Philippines, Bouru, Hawaiian Islands (Oahu and Maui), China, N. Australia, Southern
United States, Antilles, Guiana. The type of C. formicarius was from Tranquebar, that
of C. turcipennis from Java. In the Seychelles, a specimen was found in Mahé in 1905 ;
several were found on sweet-potatoes on Cascade Estate, about 800 feet, im 1909; others
from various low-country localities in Mahé, 1908—9; one from the cultivated islet,
Anonyme Island, near Mahé, 1909; and one from Cotes d'Or Estate, Praslin, XI. 1908.
Group Antliarrhina.
MICROPLATYMERUS, h. gen.
Rostrum short, stout, conical, convex, the antennze inserted low down near the base,
the scrobes short, shallow, curved, reaching the lower margin of the eyes; eyes large, oval,
transverse, lateral, not prominent, placed at the base of the rostrum, the head constricted
behind them, polished, globose; mandibles small, scarcely visible; antennee (fig. 37 6)
geniculate, short, the scape stout, reaching the posterior margin of the eyes, the funiculus
with five closely articulated joints, 3—5 strongly transverse, the club small, oblong-ovate,
shining, feebly annulate; prothorax greatly developed, rounded-trapezoidal ; scutellum
very small, transverse ; elytra oblong, comparatively short, somewhat obliquely cut off at
the base, covering the pygidium, the humeri subrectangular ; coxze very widely separated,
the anterior pair distant from the base of the prosternum, small, deeply inserted ; pro-,
meso-, and metasternum on the same plane, the last two fused; mesothoracic epimera
large, ascending, but not impinging on the humeri; metathoracic episterna rather broad,
polished, fused with the sternum; ventral segments 1 and 2 connate, 3 and 4 extremely
short, the sutures 1 and 2 subarcuate, the others straight; legs short ; femora very stout,
clavate, compressed ; tibiee short, subtriangular, unarmed at the tip, angulate at the outer
apical angle, rounded within, the anterior pair (fig. 37 c) narrowly produced at the mner
apical angle ; tarsi with joints 1 and 2 small, 3 strongly bilobed and clothed with long
hairs beneath, 4 as long as 1—3 united, the claws long and divergent; body oblong-oval,
depressed, polished, winged.
Type, M. lodoiceworus.
The minute insect from which the above-mentioned characters are taken is related to
the 8. African genus Platymerus, from which it differs in having the antennz geniculate,
with a short, closely-articulated, 5-jointed funiculus, and a small oblong club, and the
anterior coxee small and very widely separated. Microplatymerus has the general facies
of a small Calandrid, and the broad flattened sterna and ascending mesothoracic epimera
are suggestive of certain Madarides of the group Barina. No external sexual characters
have been detected. Platymerus and Antharrhinus live on African Cycads, the present
genus on Lodoicea, the Coco-de-mer palm.
SECOND SERIES—ZOOLOGY, VOL. XVI. 58
A452 PERCY SLADEN TRUST EXPEDITION
71. Microplatymerus lodoiceivorus, n. sp. (Pl. 24, figs. 37, 37 a—c.)
Rufo-testaceous, the eyes black, shining, finely flavo-pubescent. Head smooth ;
rostrum about half the length of the prothorax, sparsely, finely punctate. Prothorax
large, broader than long, gradually narrowing from a little before the base to the
extremely short collar-like apical portion, the sides rounded towards the base and apex,
the hind angles obtuse, the base subtruncate ; sparsely, minutely punctate, the punctures
becoming coarser and more closely placed towards the sides, the dise with a narrow
smooth space down the middle. Elytra barely one-half longer than the prothorax,
conjointly rounded at the apex ; sharply striate, the striz feebly punctate, the interstices
broad, flat, each with an irregular row of extremely minute punctures. Beneath very
sparsely, minutely, the lateral portions closely and somewhat coarsely, punctate, the
metathoracic episterna smooth, Intermediate and posterior tibiz each with two minute
teeth at the outer apical angle.
Length 13—2 mm.
Loc. Seychelles: Praslin.
Nearly one hundred specimens of this insect were obtained by Mr Scott from the
leaf-bases of a growing $ Coco-de-mer palm (Lodoicea seychellarum), in the Vallée de Mai,
on Nov. 28th, 1908.
Group Trypetina.
The three genera placed in this group, like some of the American Trypetina, probably
attack palms. Certain New Zealand forms at present referred to Cossonide (e.g. Pogo-
norrhinus and Arecophaga) would doubtless be better included here.
HoMALORRHYNCHUS, n. gen.
Rostrum long, slender, dissimilarly formed in the two sexes—nearly straight,
flattened, and rugose in ¢, cylindrical, arcuate, and polished in $,—the antenne inserted
at (¢) or considerably beyond (2) the middle, the scrobes inferior, extending to beneath
the eyes; antennz slender, the funiculus 7-jointed, the club oval, annulate, shining,
sparsely pubescent ; head large, somewhat deeply inserted, the eyes oval, transverse,
moderately large, placed on the anterior portion immediately behind the base of the
rostrum ; prothorax rounded laterally, sometimes broader and subglobose in g, truncate or
feebly sinuate at the base, with a short, narrow anterior lobe; scutellum small; elytra
oblong ; coxee widely separated ; mesothoracic epimera not ascending ; metasternum long
the episterna narrowly exposed; ventral segments 1 and 2 long, connate at the middle,
3 and 4 extremely short, the sutures straight; legs elongate; femora clavate, slender at
the base, unarmed ; tibize almost straight, feebly unguiculate at the outer apical angle ;
tarsi sparsely pilose, spongy-pubescent beneath, joint 1 small, 2 triangular, 3 bilobed,
the lobes long and divergent, densely ciliate at the tip, the claw-joint long, the claws
simple, free or subconnate ; body elongate, rather convex, finely pubescent, in one species
with small patches of scales, winged.
Type, H. serripes,
CHAMPION—COLEOPTERA ; CURCULIONID A 453
The three species referred to Homalorrhynchus, one of which has been found in the
leaf-bases of the Verschaffeltia-palm, are somewhat nearly related to the Tropical
American genera Nanus, Schonh. (= Homaloxenus, Woll.) and Hoplorrhinoides, Champ.,
one or both of which have been found on palms. The widely separated coxze, the
dissimilarly formed rostrum in the two sexes, the long legs, the clavate femora, and the
divergent, apically ciliate lobes of the third tarsal joint, are its chief characters.
The shining antennal club is suggestive of Calandrine, and the rostral structure somewhat
approaches that of the palm-feeding Derelomus; but the Seychelles insects are not other-
wise very closely allied to either of them. JLeptobaris, Gerst., from Mozambique, to
Judge from the published figure in Peters’ “Reise,” is not unlike the present genus, but
it is probably a true Barid. The generic name Homalorrhynchus is taken from the
flattened rostrum of the male.
72. Homalorrhynchus serripes, n. sp. (Pl. 24, fig. 38, 2.)
Elongate, moderately shining; ferruginous or reddish-brown, the head, prothorax,
and under surface, and the thickened portion of the femora, sometimes piceous or nigro-
piceous ; sparsely clothed with fine yellowish adpressed hairs, the hairs on the prothorax
long, those on the elytra uniseriately arranged down each interstice. Head densely
punctate ; rostrum (¢) about as long as the prothorax, straight, flattened above, rugosely
punctate to the tip, (?) cylindrical, much more slender, arcuate, shining, very sparsely,
minutely punctate; joint 2 of the funiculus about as long as 3 and 4 united, 3—7 short,
widening outwards. Prothorax convex, in fully-developed f nearly as long as broad, sub-
globose, and a little wider than the elytra, in ? transverse and less dilated; densely
punctate, the interspaces alutaceous. Elytra moderately long, parallel in their basal half,
in the larger males very gradually widened forwards; punctate-striate, the interstices
rugulose. Beneath shining, closely, rather finely punctate; ventral segments 1 and 2
deeply excavate down the middle in ¢. Anterior tibize of ¢ ciliate and finely or obsoletely
serrulate within, and also toothed at the inner apical angle. Tarsal claws free.
Length 24—37 mm. ($?).
Loc. Seychelles: Mahé.
Hight specimens, varying greatly in size, the head and prothorax piceous in one
of the larger males. The cilia on the anterior tibize of the male are so fine that they are
soon abraded. The insect occurs in the forests near Morne Blanc, the high damp forest at
the summit of Pilot (2000 feet), the forest behind Trois Fréres (1500—2000 feet), the
forest above Cascade, and the Mare aux Cochons district.
73. Homalorrhynchus rubricatus, n. sp.
Moderately elongate, convex, subcylindrical, somewhat shining; rufescent, the femora
and antennal club piceous; obsoletely pubescent. Head closely punctate; rostrum (g)
almost straight, as long as the prothorax, slender, with the parallel-sided apical portion a
little wider, flattened and rugosely punctate to the tip, (¢) very slender, strongly arcuate,
cylindrical, shining, faintly punctate. Prothorax nearly or quite as long as broad, rounded
at the sides, narrowed and feebly constricted anteriorly ; densely punctate, the interspaces
58—2
454 PERCY SLADEN TRUST EXPEDITION
dull and alutaceous. Elytra parallel in their basal half, of the same width as the
prothorax ; punctate-striate, the interstices not wider than the striz, rugulose. Beneath
shining, closely, coarsely punctate ; ventral segments 1 and 2 deeply excavate down the
middle in g. Anterior tibize unarmed on their inner edge. ‘Tarsal claws free.
Length 14—2 mm. ($9).
Loc. Seychelles: Mahé.
Three males and two females, apparently more or less abraded: found near Morne
Blane, in the high forest of Morne Blanc and Pilot, and in the forest behind Trois Fréres
(1500—2000 feet). Narrower and more convex than H. serripes, and of about the same
size as the smaller examples of that species, the rostrum more slender in both sexes, and
more strongly bowed in the female, the under surface more coarsely punctate, the cox
less widely separated, the anterior tibie unarmed on their inner edge in both sexes. The
insect was found in the same localities as H. serripes.
74. Homalorrhynchus verschaffeltie, n. sp. (Pl. 24, figs. 39, 39 a, 2.)
Elongate, convex, shining ; obscure testaceous, the prothorax more or less suffused
with piceous, the elytra each with a rather broad transverse black fascia beyond the
middle, not quite reaching the suture; clothed with scattered, minute, hair-like, pallid
seales, those on the elytra being uniseriately arranged down each interstice, and also with
patches of coarser whitish seales—one on each flank of the prothorax, as well as a trans-
verse row at the base, and three small streaks along the third elytral interstice (one at
the base, and another preceding and following the black fascia). Head densely punctate ;
rostrum (¢) a little longer than the prothorax, straight, rugosely punctate to the tip,
(?) very slender, slightly thickened at the base, arcuate, shining, obsoletely punctate.
Prothorax broader than long, strongly rounded at the sides to the narrow apical portion,
feebly bisinuate at the base ; closely punctate, the interspaces shining. Elytra moderately
long, very gradually narrowed from the base, as wide as the prothorax ; punctate-striate,
the interstices punctulate. Beneath sparsely punctate; ventral segments 1 and 2 broadly
excavate down the middle in ¢. Anterior tibizee unarmed on their inner edge. Tarsal
claws subconnate at the base.
Length 24—23 mm. ($9).
Loc. Seychelles: Mahé.
One male and two females. More shining than H. serripes, the prothorax relatively
shorter in the male, the elytra nigro-fasciate, both with small patches of whitish scales,
the tarsal claws subconnate at the base, the anterior tibize unarmed on their inner edge.
The coxee are all widely separated. The three specimens were taken near Morne Blane,
from between the leaf-bases of a growing Verschaffeltia-palm.
SPHODRIAS, n. gen.
Rostrum (¢), moderately stout, shorter than the prothorax, almost straight, flattened
above, the antenne inserted near the tip; antennz slender, the funiculus 7-jointed, the
club elongate, acuminate, rather slender, pubescent, annulate ; head inserted to near the
CHAMPION—COLEOPTERA; CURCULIONIDA 455
eyes, the latter oval, transverse, large, not prominent; prothorax large, oval, bisinuate
and closely fitting to the elytra at the base; scutellum rounded ; elytra elongate; coxee
widely separated ; metasternum long, the episterna exposed; ventral segments 1 and 2
moderately long, 3 and 4 together shorter than 2, the sutures straight, the intercoxal
process angulate in front; femora clavate, slender at the base, unarmed ; tibie strongly
unguiculate at the inner apical angle; tarsi spongy-pubescent beneath, joint 1 small,
2 large, transverse, 3 broader than 2, strongly bilobed, the two lobes rounded, and ciliate
externally, the claw-joint not much longer than 3, the claws small, divergent ; body
subcylindrical, elongate, squamose, winged.
Type, S. magdaloides.
This genus, the essential characters of which are taken from the male sex only, is
closely related to Homalorrhynchus, differing from it in the more elongate antennal club,
the strongly unguiculate tibie, the broader tarsi (the third joint of which has rounded
lobes and the claw-joint is comparatively short), the longer third and fourth ventral
segments, and the subcylindrical squamose body. The female doubtless has a slender arcuate
rostrum. The type has the general shape of a Magdalis. It may live upon some palm ?
75. Sphodrias magdaloides, n. sp. (Pl. 24, fig. 40, ¢.)
g. Subcylindrical, robust, moderately convex, opaque; obscure ferruginous, varie-
gated, the femora included, with black, the antennz ferruginous, with the club black ;
the upper surface maculated with dense patches of small adpressed ochreous scales, the
scales on the prothorax condensed into about ten confluent spots and those on the elytra
into numerous spots and streaks, which are mainly placed on the alternate interstices, the
head, rostrum, under surface, and legs with small scattered ochreous scales. Head densely
punctate ; rostrum considerably shorter than the prothorax, slightly widened outwards,
rugosely punctate to the tip; antennal club about as long as the last four joints of the
funiculus, joints 1 and 2 of the latter elongate, the others short. Prothorax about as long
as broad, rounded at the sides, narrowing from about the middle to the apical constriction ;
densely punctate. Elytra elongate, parallel in their basal half, very little wider than the
prothorax ; punctate-striate, the interstices flat, densely, rugosely punctate. Beneath
shining, densely, coarsely punctate; ventral segments 1 and 2 deeply excavate down the
middle. Anterior tibize closely serrulate along their inner edge, bowed inwards at the
apex, the apical uncus thus appearing very prominent.
Length 43, breadth 12 mm.
Loc. Seychelles: Silhouette.
One male, captured at Mare aux Cochons or in the forest immediately above,
Sept. 1908.
SYCITES, n. gen.
Rostrum (2) rather slender, long, feebly curved, flattened above, the antennze inserted
in the dilated median portion, the scrobes lateral, obliquely descending to beneath the
eyes, the buccal cavity transverse; mandibles short, feebly bidentate at the tip; head
rather small, the eyes oval, small, depressed, placed immediately behind the base of the
456 PERCY SLADEN TRUST EXPEDITION ©
rostrum ; antennee rather stout, the funiculus 7-jointed, joint 3 transverse, 4—7 cylindrical,
the club elongate, annulate, its basal half very little stouter than the funiculus, both
closely pilose ; prothorax long and broad, rounded laterally ; scutellum small, rounded ;
elytra oblong, covering the abdomen; cox very widely separated, the pro-, meso-, and
metasternum on the same plane, the mesosternum greatly developed, the anterior coxe
large and deeply inserted, the metathoracie episterna narrow; ventral segments 1 and 2
connate at the middle, 3 and 4 together about as long as 2, the sutures straight; legs
stout; femora feebly clavate, unarmed; tibize sinuous within, straight on their outer
edge, not carinate, the short uncus arising from about the middle of the apical margin ;
tarsi short, stout, joints 1—3 spongy-pubescent beneath, 2 transverse, much wider than 1,
3 broad, bilobed, the lobes rounded externally, the claws short, free ; body elongate, broad,
depressed, squamose and pilose, winged.
Type, S. pilicorms.
The type of this genus, the male only of which has come to hand, has exactly the
facies of a true Calandrid, from which it differs in having the pygidium completely covered
by the elytra, and the antennal club pilose and annulate. The antennz (f) are peculiarly
formed, the third joint of the funiculus being extremely short and those following sub-
cylindrical, the club, again, having a long cylindrical rugulose basal portion. Sycites
must be placed in the Trypetina of Lacordaire, not far from Nanus. Arthrotomus, type
A. depressus, Klug, from Madagascar, also comes near the present genus. The differently
formed mandibles separate Sycites from Rhina.
76. Sycrtes pilicornis, n. sp. (Pl. 24, fig. 41, 3.)
g. Elongate oval, broad, depressed, opaque above, shining beneath; dark brown,
the rostrum, antennee, legs, and under surface obscure ferruginous; the elytra and the
sides of the prothorax variegated with dense patches of small ochreous scales, those on
the elytra reduced to spots or streaks at the base, along the sides, and on the apical
declivity, the inter-ocular portion of the head also ochreo-squamose, the very minute
darker scales on the rest of the upper surface inconspicuous, the under surface and legs
finely pubescent. Head densely punctate; rostrum nearly as long as the prothorax,
slightly widened at the base, middle, and apex, asperato-punctate and subcarinate, armed
at the tip above with a small erect tooth on each side; antennee with the funiculus and
club closely, shortly pilose, the former with joint 2 triangular, 3 strongly transverse,
4—7 subcylindrical, short, together about as long as the club, 7 slightly longer than
broad, the widened portion of the club very shining, the base rugulose. Prothorax large,
slightly broader than long, distinctly wider than the elytra, bisinuate at the base, rapidly
arcuately narrowing from the middle to the shallow apical constriction ; densely, finely
punctate. Elytra very gradually narrowimg from the base, conjointly rounded at the
apex; finely punctate-striate, the interstices broad, flat, densely rugulose. Beneath
densely, moderately coarsely punctate; ventral segment 1 broadly excavate down the
middle. i
Length 43, breadth (prothorax) 12 mm.
Loc. Seychelles: Mahé.
CHAMPION—COLEOPTERA ; CURCULIONIDA A57
Two males, from the forest of Morne Blanc. Probably attached to some kind of
palm, though no record of this is attached to the specimens.
Group Cossonina.
The Cossonina, as in many oceanic islands, are abundantly represented in the
Seychelles. The collection before me contains 49 species, 29 of which belong to the
Pentarthrides, some of them being very minute, and numerous new genera, if the
characters used by Wollaston in his classification are to be considered of sufficient value,
are required for their reception. The 18 Cossonides, on the contrary, with two exceptions,
belong to known generic types, and are therefore much less specialized in the islands, some
of them, in fact, are introductions. The two Dryophthborides call for no special comment.
Many of the Pentarthrides are noted as having been found in the leaf-bases of endemic
palms, or of screw-pine, hence it is not surprising to find many new forms amongst them.
One species of Stenotrupis (= Proéces rufipes Fairm.), from the island of La Digue, is not
represented in Mr Scott’s collection, this making 50 in all from the Seychelles group.
Sect. Dryophthorides.
DRYOPHTHORUS.
Dryophthorus Schénherr, Curc. Disp. Meth., p. 332 (1826); Wollaston, Trans. Ent.
Soe. Lond., 1873, p. 442.
This genus is referred by most writers to the Cossonina, but its proper place is
probably in the Calandrinz, near the Sipalides.
A cosmopolitan genus, with many endemic species in the Hawaiian Is., possibly
introduced in Europe.
77. Dryophthorus lymeaylon.
Curculio lymexylon Fabr., Ent. Syst., i. 2, p. 420 (1792).
Curculio corticalis Payk., Mon. Cure., p. 41 (1792).
Dryophthorus corticalis Say, Descr. N.Am. Cure., p. 24 (1831); Complete Writings, i.
p. 292.
Dryophthorus assimilis Gahan, Monogr. Christmas Is., p. 116 (1900).
Loc. Seychelles: Mahé.
A common insect in the high forest of Morne Blane, Trois Fréres, and Cascade
Estate. I can see no difference between specimens from Europe, N. America, Christmas
Island, and Mahé, when clean examples are compared. They all have a well-defined
subapical carina to the elytra formed by the united, more or less costate, fifth and
seventh interstices. The elytra, themselves, vary in length, and in some examples the
alternate interstices towards the apex are distinctly more raised than the rest.
78. Dryophthorus ecarinatus, n. sp.
Elongate, subfusiform, opaque, when fresh plumbeo-pruinose ; nigro-piceous or black,
the antennz (the club excepted) and tarsi ferruginous, the femora and tibiz sometimes
458 PERCY SLADEN TRUST EXPEDITION
obscurely rufescent. Rostrum very stout, moderately long, coarsely punctate; head
similarly punctured ; eyes small, transverse, depressed. Prothorax long, subcylindrical,
deeply constricted beyond the middle; coarsely, closely punctate. Elytra comparatively
short, wider than the prothorax, obliquely narrowed from about the basal third, and
produced at the apex, the humeri oblique; coarsely punctate-striate, the interstices
uniformly convex throughout, the fifth and seventh interstices not carinate at the apex.
Length (exclud. rostr.) 14%—275 mm.
Loc. Seychelles : Mahé.
Twenty specimens, found in the same localities as the preceding species (and also
in the Mare aux Cochons District), in company with which it was very possibly taken.
Smaller than the smallest D. lymeaxylon, and separable from it by the non-carinate apices
of the elytra. No sexual differences have been detected.
Sect. Pentarthrides.
CHG@RORRHINODES, nN. gen.
Rostrum stout, curved, parallel-sided, about as long as the exposed portion of the
head, the antennz inserted at the middle, the scrobes oblique and rapidly descending ;
head long, exserted, the eyes small, lateral, somewhat prominent, distant from the
prothorax ; antennze moderately long, the funiculus 5-jomted, joint 1 stout, 2—4 widening
outwards, 2 small, triangular, 3—5 transverse, the club small, oval, annulate, pubescent ;
prothorax subcylindrical, constricted before the apex; scutellum scarcely visible; elytra
elongate, parallel-sided to beyond the middle, the apices simple; anterior coxze somewhat
narrowly separated ; metasternum long, unimpressed ; ventral segments 1 and 2 connate,
2 a little longer than 3 and 4 united, the latter short, the sutures straight ; legs short ;
tibize with a slender uncus at the outer apical angle; tarsi with joints 1 and 2 very short,
narrow, 3 wider, feebly lobed, the claws small; body very narrow, elongate-fusiform,
opaque, rugose and incrustate above.
Type, C. tenuiculus.
The type of this genus may be described as a very narrow, small, feebly-developed
Cherorrhinus (= Pentacoptus, Woll.), with a strongly exserted head, a curved, moderately
long, stout, subeylindrical rostrum, rather long antennze, and the stout, spiniform
prolongation of the outer angle of the tibiee reduced to a slender uncus. The general
facies is suggestive of that of Melarhinus. The type of Cherorrhinus is European, that
of Pentacoptus is from Japan. The 5-jointed funiculus separates the present genus
from Pholidoforus. C. tenwculus is probably winged.
79. Cherorrhinodes tenwculus, n. sp.
g. Very elongate, narrow, fusiform, feebly convex, opaque ; piceous, the antennee
and legs obscure ferruginous, the dense rugose punctuation of the upper surface visible
through the earthy inerustation. Rostrum stout, less rugosely punctured than the
exposed portion of the head, the two together much longer than the prothorax, the
head narrow, less than twice the width of the rostrum. Prothorax about as long as
——
CHAM PION—COLEOPTERA ; CURCULIONID # 459
broad, the strong subapical constriction leaving a collar-like anterior margin. Elytra
rather more than three times the length of, and considerably wider than, the prothorax,
attenuate at the apex ; with closely-packed rows of coarse, oblong punctures placed in
rather deep strive, the very narrow rugose interstices somewhat convex, 2 and 3 becoming
more prominent on the apical declivity. Beneath somewhat shining, closely punctate ;
ventral segment 1 shallowly sulcate down the middle.
Length 2 mm.
Loc. Seychelles: Silhouette.
One example, found on Sept. 6th, 1908, at Mare aux Cochons. The insect super-
ficially resembles a narrow Monotoma.
PENTARTHRUM.
Pentarthrum Wollaston, Ann. Mag. Nat. Hist., (2) xiv. p. 129 (1854); Trans. Ent.
Soc. Lond., 1873, pp. 13, 446, 513.
Four species from the Seychelles are referred to this genus, type P. huttoni (which
is probably an exotic form introduced into Europe), the numerous other ‘‘ Pentarthrides ”
from these islands being better placed elsewhere. The head-quarters of Pentarthrum
would appear to be in New Zealand or the Hawaiian Islands. One of the new species
now added, P. punctatissimum, is remarkable in having an unusually slender rostrum
in the female. Some of the known forms have an extraordinary wide distribution.
80. Pentarthrum longicolle, n. sp. (Pl. 24, fig. 42, 3.)
g. Very elongate, subcylindrical, rather convex, shining, black, the antennee and
tarsi nigro-piceous. Rostrum very stout, moderately long, widened towards the base,
coarsely, closely punctate, the punctuation becoming much finer on the apical half,
shallowly suleate at the base; eyes prominent, the head constricted behind them ;
antennee stout, joints 2—5 of the funiculus strongly transverse, the club small. Prothorax
about one-fourth longer than broad, rounded at the sides posteriorly, gradually narrowing
forwards, and deeply constricted before the apex; coarsely, closely punctate, except
along a narrow smooth median space. Scutellum minute. LElytra moderately long,
of the same width as the prothorax, subparallel in their basal half, broadly produced
at the apex, and with the apical portion narrowly explanate externally ; coarsely crenate-
striate, the very narrow convex interstices uniseriate-punctate. Beneath very coarsely,
the prosternum rugosely, punctate; metasternum depressed in the centre posteriorly;
ventral segment 1 excavate down the middle, 5 with a compressed, smooth, cariniform,
median ridge. Anterior coxee large, narrowly separated. Tibiz sharply mucronate
at the inner apical angle, and ciliate on the inner margin near the tip.
Length 3i1—3# mm.
Loc. Seychelles: Mahé.
Two males, one from the Mare aux Cochons district, the other from the forest
above Cascade. A very distinct form, with an unusually elongate prothorax, a basally
widened rostrum, and stout antennee. P. longicolle is not unlike the Tasmanian P. nigrum,
SECOND. SERIES—ZOOLOGY, VOL. XVI. 59
460 PERCY SLADEN TRUST EXPEDITION
Woll., but it is much more elongate. The 5-jointed funiculus distinguishes it from
Rhyncolus.
81. Pentarthrum punctatissimum, n. sp.
Elongate, subfusiform, robust, feebly shining; piceous, the tarsi, the base of the
antenne, and the antennal club, often ferruginous. Rostrum (¢) stout, thickened towards
the base, feebly curved, moderately long, densely, rugosely punctate, (2) more curved,
longer, and much more slender, subcylindrical. thickened towards the base and slightly
widened at the tip, shining, finely punctate, rugose at the base; antennee in ? inserted
near the base of the rostrum, in $ at about the basal third, the club small; eyes
moderately prominent; head, the exposed part of the basal portion included, very
rugosely punctate. Prothorax much longer than broad, subconical, strongly constricted
before the apex, and rounded at the sides posteriorly; densely, coarsely punctate.
Scutellum minute. Elytra as wide as the prothorax, moderately long, very gradually
narrowing from the base in ?, subparallel in their basal third in gf, conjointly rounded and
narrowly explanate at the apex; coarsely crenate-striate, the interstices very narrow,
somewhat convex, and rugulose, the first (sutural) closely uniseriate-punctate. Beneath
coarsely, densely punctate; metasternum flattened and sulcate posteriorly; ventral
seoments 1 and 2 broadly excavate down the middle in ~. Anterior coxe narrowly
separated.
Length 24—3}! mm. (62).
Loc. Seychelles: Praslin.
Forty-two specimens, found in the leaf-bases of a growing ¢ Coco-de-mer, in the
Vallée de Mai, on Noy. 28th, 1908. In this species the rostrum is dissimilarly formed in
the two sexes, the antennal club is quite small, and the entire surface, above and beneath
(including the exposed basal portion of the head), is densely, coarsely punctate.
82. Pentarthrum seychellarum, n. sp.
g. Elongate, subfusiform, convex, shining; piceous, the tarsi and antennze obscure
ferruginous. Rostrum stout, feebly curved, parallel-sided, about as long as the width of
the head with the eyes, much more finely punctured than the inter-ocular space, slightly
depressed at the base; eyes prominent, rather large; antennee with joints 3—5 of the
funiculus strongly transverse, the club small. Prothorax a little longer than broad,
oblongo-conic, rounded at the sides posteriorly and feebly constricted just before the apex;
densely, moderately coarsely punctate, except along the smooth abbreviated median line.
Scutellum minute. Elytra moderately long, slightly wider than the prothorax, sub-
parallel in their basal third and gradually narrowing thence to the apex, the apices not
explanate; crenate-striate, the interstices transversely rugulose and closely uniseriate-
punctate. Beneath closely punctate; ventral segment 1 excavate down the middle.
Anterior tibiz sinuate, and ciliate towards the apex, within.
Length 23 mm.
Loc. Seychelles: Mahé.
One specimen, from the Cascade Estate, 800—1500 feet. The parallel-sided rostrum
and finer sculpture separate P. seychellarwm from P. punctatissimus and P. longicolle.
CHAMPION—COLEOPTERA ; CURCULIONIDA 461
83. Pentarthrum scott, n. sp.
Moderately elongate, narrow, rather convex, shining; rufo-piceous, with a faint
brassy tinge. Head and rostrum closely, very finely punctate; rostrum stout, sub-
cylindrical, feebly curved, rather more than half the length of the prothorax; head
constricted behind the eyes, the latter prominent and moderately large; antennze short,
the club rather small, ovate. Prothorax a little longer than broad, somewhat oval, narrowed
and constricted towards the apex ; closely, finely punctate, except along a narrow space
down the middle. Scutellum extremely minute. Elytra moderately long, very gradually
narrowing from the base, conjointly rounded at the apex ; punctato-sulcate, the interstices
narrow, faintly punctulate. Beneath sparsely, very finely punctate, the metasternum
suleate. Anterior coxee separated by the width of one of them. Tarsi short, the third
joint feebly lobed.
Length 24 mm. (9).
Loc. Seychelles: Mahé.
One specimen, assumed to be female, the first ventral segment being unimpressed.
Smaller, narrower, and less convex than P. seychellarum, the prothorax very finely
punctate, the elytra punctato-sulcate and with much smoother, narrower interstices.
P. scotti was found in the high forest of Morne Blane.
TEMNORRHAMPHUS, h. gen.
Rostrum broad, transverse, parallel-sided, grooved above, truncate at the tip, leaving
the apices of the mandibles exposed, the scrobes very deep, transverse, the antennze
inserted near the apex, the buccal cavity large; head broad, feebly constricted behind
the eyes, the latter not very prominent; coxee large, the anterior pair separated
by the width of the coxa; antenne stout, with a 5-jointed funiculus and a small
club, the scape extending to far beyond the eyes; tibize with the outer apical angle
produced into a long curved hook, the inner angle mucronate; body elongate, robust,
subcylindrical, glabrous; the other characters as in Pentarthrum (type P. hutton,
Woll.).
Type, 7. laterostras.
The single species referred to Temnorrhamphus may be described as a robust
Pentarthrum with the head and rostrum formed very much as in the Rhyncolid-genus
Eremotes. It cannot be included in Pentarthrum as defined by Wollaston. The type is
a strongly-built, convex, subcylindrical insect, with the apices of the elytra more narrowly
explanate than in the widely distributed P. apicale, Broun, and the large anterior coxe
separated by a space equalling the width of one of them.
84. Temnorrhamphus latirostris, n. sp.
Moderately elongate, rather convex, subcylindrical, feebly shining ; piceous or nigro-
piceous, the elytral humeri rufescent, the antennze and tarsi obscure ferruginous. Head
and rostrum closely, finely punctate, the rostrum shallowly suleate down the middle, the
eroove extending backwards to the small inter-ocular fovea; antennze moderately stout,
59—2
462 PERCY SLADEN TRUST EXPEDITION
joints 2—5 of the funiculus strongly transverse. Prothorax about as long as broad,
convex, subtrapezoidal, constricted immediately before the apex and also at the
base (as seen from above), the base distinctly margined; densely, moderately coarsely
punctate, except along a narrow, smooth, abbreviated median line. Scutellum transverse,
small. Elytra barely twice the length of the prothorax, and of the same width, very
gradually narrowing from the base, the apex produced and narrowly explanate, the
seventh interstice also subangularly dilated externally towards the tip ; crenate-striate, the
interstices convex, transversely rugulose, and irregularly uniseriate-punctate. Beneath
alutaceous, rather finely punctate; metasternum sulcate. Anterior tibiz strongly
sinuate. Tarsi with joints 1—3 rather stout, 3 wider than 2 and feebly bilobed.
Length 24—3,, mm. (¢ %).
Loc. Seychelles: Mahé.
Four specimens, sex not definitely ascertained. These were found on Cascade Estate,
at about 800—1000 feet.
HyYPorENTARTHRUM, n. gen.
Rostrum curved, subcylindrical, subangularly dilated between the points of insertion
of the antennse, more slender in ?, the antennz inserted at (¢) or behind (2) the middle,
the scrobes inferior, short, not reaching the eyes; head small, short, sharply constricted
behind the eyes, the latter small and moderately prominent ; antennze with a 5-jointed
funiculus and a small club, the scape thickened in ¢; prothorax oval, greatly developed,
wider than the elytra in both sexes, constricted immediately before the apex ; scutellum
small; elytra moderately elongate, punctato-suleate; cox: small, widely separated,
anterior pair more distant than intermediate pair; prosternum broadly flattened ;
metasternum long ; first ventral suture distinct, smuate; tibize feebly unguiculate ; tarsi
short, rather stout, third joint feebly lobed, the claws long; body subfusiform,
subdepressed, glabrous: species small.
Type, H. microcephalum.
The single species included in this genus is nearly related to Pentarthrum, differmg
from it in the unusually small head, the long, subcylindrical, moderately stout rostrum, the
greatly developed prothorax, the broadly flattened prosternum, the very widely separated
anterior coxe, and the feebly unguiculate tibize. The head is so small that the eyes, at
first sight, appear to be inserted at the sides of the rostrum. The insect is attached to
the Coco-de-mer palm (Lodozcea).
85. Hypopentarthrum microcephalum, n. sp. (Pl. 24, fig. 438, 2.)
Moderately elongate, rather broad, subfusiform, somewhat convex above, broadly
flattened down the middle beneath, shining ; nigro-piceous, the antennze and legs piceous,
the base and club of the former and the tarsi ferruginous. Head and rostrum densely
punctate, the latter smoother in ?; rostrum (¢) about two-thirds the length of the
prothorax, moderately stout, (?) much longer and rather slender ; antennze with a stouter
scape in g than in ¢. Prothorax longer than broad, a little wider than the elytra ;
CHAMPION—COLEOPTERA ; CURCULIONIDA 463
closely, rather coarsely punctate. Scutellum smooth. Elytra moderately long, very
gradually narrowing from the base; closely punctato-sulcate, the mterstices flat. Beneath
closely, rather coarsely punctate; metasternum obsoletely canaliculate; ventral segment 1
hollowed down the middle in @.
Length 24—22 mm. (¢9).
Loc. Seychelles: Praslin.
Three males and one female, taken from the leaf-bases of a growing s Coco-de-mer,
in the Vallée de Mai, Nov. 28, 1908.
STENOPENTARTHRUM, nh. gen.
Rostrum cylindrical, rather stout, the antennz (¢) inserted towards the base; head
small, narrow, feebly constricted at a short distance behind the eyes, the latter small and
depressed ; antennz stout, with a five-jointed funiculus and a rather large abrupt club,
the scape reaching considerably beyond the eyes; prothorax oval, strongly constricted
before the apex ; scutellum small; elytra subeylindrical ; anterior coxze narrowly separated ;
legs short ; tibize feebly unguiculate ; tarsi stout, the claw-joint included, jomt 3 bilobed,
the claws small; body elongate, linear, somewhat depressed, delicately sculptured, glabrous.
Type, S. pandane.
The very small Pentarthrid from which the above characters are taken is not unlike
Stenotrupis gracilis (a species doubtfully included in Stenotrupis on account of its small,
non-elongated, conical head); but differs from it in having the rostrum and antennze
shorter and stouter, the tarsi widened and with a thicker claw-joint, and the prothorax
abruptly constricted before the apex. From Microtrupis the present genus may be
separated by the structure of the tarsi, the longer head, and the abruptly constricted
prothorax. Compared with the small, elongate species of Pentarthrum, such as P. angus-
tissimum, Woll., from Japan, the short limbs, delicate sculpture, relatively larger antennal
club, stouter claw-joint to the tarsi, &c., sufficiently distinguish Stenopentarthrum.
S. pandane is doubtless attached to Pandanus.
86. Stenopentarthrum pandane, n. sp.
g. Shining, nigro-piceous, the antenne (the club excepted) and tarsi ferruginous.
Rostrum feebly curved, nearly twice the length of the exposed anterior portion of the
head, sparsely punctate; head finely punctate; antennse with joints 83—5 of the funiculus —
strongly transverse. Prothorax longer than broad, oval, much narrowed anteriorly, and
abruptly constricted before the apex; closely, finely punctate. Elytra elongate, parallel,
of the same width as the prothorax; very finely punctate-striate, the interstices flat.
Beneath finely punctate; ventral segments 1 and 2 sulcate down the middle, 5 shallowly
foveate.
Length 13—1#4 mm.
Loc. Seychelles: Silhouette, Mahé.
Two specimens, both from the high forest, the one from Silhouette indicated as
having been taken from between the leaf-bases of a growing Pandanus Hornet.
464 PERCY SLADEN TRUST EXPEDITION
TRAPEZIRRHYNCHUS, n. gen.
Rostrum short, stout, widening outwards in both sexes, in g trapezoidal, as broad as
long, and rounded at the apex, in § more convex above, longer, and narrower, the antenne
inserted behind the middle, the scrobes transverse, shallow ; eyes small, lateral, somewhat
distant from the basal constriction of the head; head exserted, broad in 2, narrow in ?;
antennze rather slender, the funiculus five-jointed, joints 2—5 small, 3—5 transverse, the
club small, ovate, narrow, the scape sinuate and about reaching the posterior margin of the
eyes; prothorax oval; scutellum minute; elytra moderately long, punctate-striate ; coxee
small, widely separated ; metasternum lone; ventral segments 1 and 2 connate, 3 and
4 short, the sutures straight ; legs short ;\ tibiae somewhat feebly unguiculate at the outer
apical angle ; tarsi short, joints 1—3 small, subequal in length, 3 a little wider than 2, the
claw-joint about as long as the others united, the claws small; body narrow, subfusiform,
shining, glabrous, winged: species minute.
Type, ZT. silhouettensis.
The single species referred to this Pentarthrid-genus is nearly related to Proconus,
Broun (type, Pentarthrum asperirostre, Broun), from New Zealand, differmg from it in
having the head and rostrum dissimilarly formed in the two sexes, the antennal club much
smaller, the basal joints of the tarsi less widened, and the coxze not so widely separated.
The male of the New Zealand Pentarthrum helmsianum, Sharp (perhaps the finest known
Pentarthrid), has a somewhat similarly shaped rostrum.
87. Trapezirrhynchus sithouettensis, n. sp. (Pl. 24, fig. 44, g.)
Somewhat depressed, piceous or nigro-piceous, the rostrum, antennee, and legs
ferruginous or obscure ferruginous, shining, glabrous. Head and rostrum closely, minutely
punctate, together about as long as the prothorax; rostrum (¢) as broad as the head with
the eyes, transversely depressed at the base, (?) much narrower, transversely convex,
about twice as long as its width at the base, and more shining; eyes slightly prominent.
Prothorax about as long as broad, oval, narrowing forwards and moderately constricted
before the apex ; closely, finely punctate. Hlytra moderately long, not or very little wider
than the prothorax, subparallel in their basal half; closely punctate-striate, the interstices
very narrow and somewhat convex. Beneath closely punctate; metasternum obsoletely
eanaliculate; ventral segments 1 and 2 excavate down the middle in ¢.
Length 14—1% mm. (22).
Loc. Seychelles: Silhouette.
Three males and two females found at the Mare aux Cochons or in the forest above,
the males varying a little in the development of the head and prothorax.
STENOTRUPIS.
Stenotrupis Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 447, 515; Champion, Biol.
Centr.-Am., Coleopt., iv. 7, p. 13 (1909).
Dioédimorpha Broun, New Zealand Journ. Sci., i. p. 489 (1883).
Stenotrupis would appear to have its head-quarters in the Seychelles, whence 15 species
_CHAMPION—COLEOPTERA ; CURCULIONIDA 465
are now known, some of these being closely related to the type, S. crassifrons, Woll., from
Makian, in the Malayan Archipelago. They are all elongate forms, with the exposed
anterior portion of the head prolonged or dilated, and the small, depressed or inconspicuous,
eyes well separated or distant from the basal constriction; and the rostrum more or less
thickened at the tip, stouter in ¢ than in 2, sometimes much widened in the former, and
sometimes extremely slender in the latter, sex. In the remarkable series obtained by
Mr Scott there is a complete gradation from the robust S. crassipes to the minute slender
S. dumetorum. The males have the first one or two ventral segments more or less grooved
down the middle, and the fifth sometimes foveate. The genus also includes various forms
from Tropical America and New Zealand, and is doubtless represented in other regions.
One of the New Zealand species, S. (Pentarthrum) wollastoniana, Sharp, the type of
Dioédimorpha, Broun, is closely related to S. parallela; and Pentarthrum debile, Sharp,
from New Zealand, and P. prolzum, Sharp, from the Hawaiian Islands, also belong here.
a. Head with the exposed anterior portion oblong, dilated, or subconical, sometimes
considerably shorter or transverse in ?, and constricted at a greater or less
distance behind the eyes, the latter small; prothorax elongate.
aw
Tibize (or at least the anterior pair) distinctly mucronate at inner apical
angle, the outer uncus well developed.
av. Rostrum very long and subcylindrical in both sexes, at most feebly widened
at tip; body very elongate ; under surface coarsely punctate.
a. Body fusiform; prothorax more finely punctate; rostrum stout in 4%,
slender in ¢; anterior portion of head oblong in g, shorter in ?
lodorceicola.
b°. Body subeylindrical; prothorax coarsely punctate; rostrum stout in ?
[¢ unknown]; anterior portion of head transverse (2) ... rufipes.
b’. Rostrum much shorter; elytra less elongate: species small conveaxiuscula.
c’. Rostrum widened outwards, stout in 2, slender in @.
ce. Anterior portion of the head oblong ; body elongate, depressed.
a.
Upper surface densely, coarsely punctate ; legs stout ; third tarsal joint
broad, bilobed ; under surface closely, coarsely punctate: species
large and robust.. ae se oak we. Crassipes.
b'. Upper surface more anes punctate ; diel fae joint smaller ; under
surface sparsely, finely punctate: species more slender parallela.
d’. Anterior portion of the head transverse ; body subeylindrical ... ilum.
b'. Tibize not or obsoletely mucronate at inner apical angle, the outer uncus feebly
developed ; body depressed, narrow ; rostrum more or less thickened at
tip: species small or very small.
d*. Rostrum broad and much widened outwards in ¢, slender and more elongate
in ? ; apex of elytra sparsely pilose ... i d ... biformis.
e. Rostrum slender in ¢, stouter in ¢, slightly Pilea at hie
e. Upper surface finely pubescent throughout.
4
ce. Third tarsal joint broader, bilobed ; upper surface subopaque tarsalzs.
d‘. Third tarsal jomt narrower.
466 PERCY SLADEN TRUST EXPEDITION
a. Upper surface opaque, very densely punctate ; rostrum rather stout
Woe sen a at Be ee calaginosa.
b°. Upper surface subopaque, more sparsely punctate; rostrum more
slender in ¢ ae Bee ah ae conicicephala.
f°. Upper surface pubescent at apex only, shining.
e’. Prothorax closely punctate... ie uh a8: ... nemoralis.
jf’. Prothorax sparsely punctate... se AS Lae -.- poltia.
3
g. Upper surface glabrous or subglabrous.
g'. Upper surface subopaque, sericeous; rostrum rather stout in g
sericata.
ht. Upper surface shining.
ce. Body more coarsely punctate above: species larger... nitidula.
d’. Body finely punctate above: species small ... ... silvicola.
b. Head with the exposed anterior portion shorter, conical, very feebly constricted
behind the eyes, the latter depressed ; prothorax less elongate ; third tarsal
joint narrow: species minute ... ie ae S86 ... dumetorum.
88. Stenotrupis lodorceicola, n. sp.
Very elongate, narrow, somewhat fusiform, feebly shining ; black or pitchy-black, the
antennee and tarsi ferruginous, the rostrum, femora, and tibiz rufo-piceous. Head and
base of rostrum densely and somewhat rugosely, the rest of the latter finely, punctate, the
apical portion of the rostrum a little smoother in ?; head in ¢ elongated in front of the
deep basal constriction and with the post-ocular portion nearly twice the length of the eyes,
in ? much sborter, and with the post-ocular portion less than the length of the eyes, the
latter feebly convex; rostrum ($2) elongate, curved, thickened at the base, and slightly
widened at the apex, in g stout, in $ more slender; antennze moderately long, the club
rather small, the joints of the funiculus somewhat loosely articulated. Prothorax
elongate, obliquely narrowing from about the middle to the deep subapical constriction ;
closely punctate. Elytra elongate, as wide as the prothorax, gradually narrowing from
the base, the apices somewhat separately rounded and very narrowly explanate ; closely
punctate-striate, the interstices flat, faintly punctulate, and transversely rugulose.
Beneath closely, coarsely punctate; metasternum feebly sulcate posteriorly ; ventral
seoments 1 and 2 slightly depressed down the middle, and 5 transversely impressed, in
§. Tibiz mucronate at the inner apical angle. Third tarsal joint bilobed.
Length 32—4% mm. (92).
Loc. Seychelles: Praslin.
Twenty-two specimens, found in the leaf-bases of a growing g Coco-de-mer (Lodozcea
seychellarum), in the Vallée de Mai, Noy. 28th, 1908. In this insect the rostrum is
elongate in both sexes (the exposed portion of the head together with the rostrum being
about as long as the prothorax), and the head is much longer in the male than in the
female. S. lodoiceicola approaches S. parallela and S. crassipes. The tibize are mucronate
at the inner apical angle in both sexes, as in the Malayan S. crassifrons, Woll.
CHAMPION—COLEOPTERA ; CURCULIONIDA 467
89. Stenotrupis rufipes.
Proéces rufipes Fairm., Ann. Soc. Ent. France, 1902, p. 381; Kolbe, Mitteil. Mus.
Berlin, v. p. 45 (1910).
?. Very elongate, narrow, robust, flattened on the disc, somewhat shining ; piceous,
the antennee, apical half of the rostrum, and legs ferruginous. Head with the post-ocular
portion (as seen from above) a little longer than the somewhat depressed eyes, closely,
coarsely punctate ; rostrum feebly curved, stout, cylindrical, together with the head about
as long as the prothorax, slightly widened towards the base and apex, finely punctate in
its outer half, the punctuation of the basal portion as coarse as that of the head, the
antennee inserted at about the basal third. Prothorax elongate, one-half longer than broad,
gradually narrowed from the basal third to the deep apical constriction ; coarsely, closely
punctate. Elytra elongate, of the same width as the prothorax, subparallel in their
basal half; closely punctate-striate, the interstices narrow, flat on the disc, feebly
punctulate. Beneath subopaque, coarsely, closely punctate. Tibize mucronate at the
inner apical angle and strongly unguiculate at the outer apical angle.
Length 32 mm. (?).
Loc. Seychelles: La Digue.
The above description is taken from the type belonging to the Paris Museum.
Fairmaire included this species under Proéces, which has a 7-jointed funiculus, not
noticing, apparently, that five joints only were present in his P. rufipes. Amongst
the Seychelles forms here described, S. rufipes is perhaps nearest allied to S. parallela,
differing from it in the transverse, coarsely punctured exposed portion of the head ;
the stouter, more coarsely punctate rostrum ; the less elongate, more coarsely punctate
prothorax ; the larger punctures of the elytral striz; the coarsely punctate, subopaque
under surface; and the ferruginous legs, antenne, and rostrum. From S. lodoiceicola,
which has a similarly shaped head and coarsely punctate, dull under surface, it may
be distinguished by its narrower and more cylindrical form (the prothorax being less
narrowed anteriorly), the stouter rostrum in the female, &c.
90. Stenotrupis convexiuscula, n. sp.
&. Very elongate, narrow, subfusiform, somewhat convex, feebly shining ; black,
the antennee and tarsi ferruginous, the femora and tibiz piceous, the rostrum rufescent.
Head and base of rostrum densely, rugosely punctate, the rest of the rostrum shining and
faintly striate-punctate ; rostrum moderately stout, feebly curved, barely twice the length
of the head, distinctly widened at and before the base, and also at the tip; head oblong,
constricted far behind the eyes, the latter depressed; antenne rather slender, the club
abrupt. Prothorax elongate, subconical, rounded at the sides posteriorly, constricted
before the apex; densely punctate, except along the incomplete smooth median line.
Elytra moderately elongate, subparallel in their basal half, and narrowing thence to
the apex; very finely punctate-striate, the interstices flat and transversely rugulose.
Beneath more shining than above, closely punctate; metasternum sulcate; ventral
segments broadly depressed down the middle, 5 densely punctured and pubescent.
SECOND SERIES—ZOOLOGY, VOL. XVI. 60
468 PERCY SLADEN TRUST EXPEDITION
Anterior cox separated by the width of one of them. Anterior tibis mucronate at the
inner apical angle. Third tarsal joint moderately dilated.
Length 24 mm.
Loc. Seychelles: Silhouette.
One male, found in forest at the Mare aux Cochons, Sept. 6th, 1908. This insect is
more convex than most of its allies, and fusiform in shape; the head and prothorax are
densely punctured; the elytra, as compared with the elongate prothorax, are com-
paratively short, narrowed from the middle, and very finely punctate-striate ; and the
inter-antennal portion of the rostrum is conspicuously widened. The elongate head
brings the species into Stenotrupis.
91. Stenotrupis crassypes, n. sp.
¢. Very elongate, robust, filiform, moderately depressed, feebly shining, rufo-piceous.
Head and rostrum together about as long as the prothorax, closely, rather coarsely punc-
tate, the punctures becoming much finer on the apical portion of the rostrum, the latter
stout, curved, much widened towards the apex; head greatly developed, oblong, the sides
becoming parallel before the abrupt, deep, basal constriction, the post-ocular space about
three times the length of the eyes, the inter-ocular fovea oblong, the eyes small and feebly
convex ; antennze rather stout, the club moderately large. Prothorax elongate, gradually
narrowed from about the middle to the subapical constriction ; densely, rather coarsely
punctate, with an indication of a smooth median line. Scutellum transverse. Elytra
very elongate, subparallel, sightly compressed at the sides before the middle, conjointly
rounded at the apex ; closely, shallowly punctate-striate, the interstices not wider than
the striz, faintly punctulate. Beneath densely, rather coarsely punctate, the head
transversely rugose, the sides of the metasternum depressed and rugulosely punctured ;
ventral segments slightly hollowed down the middle, 5 with a very large rounded fovea.
Legs very stout; tibiz mucronate at the inner apical angle, the anterior pair sinuate
and ciliate within; third tarsal joint strongly bilobed, the claws small.
Length 6 mm.
Loc. Seychelles: Mahé.
One specimen, from the high forest of Morne Blane or Pilot. Larger, duller,
more coarsely punctured, and much more robust than S. parallela, the legs very stout,
the third tarsal joint strongly bilobed, the eyes a little more prominent, the head with a
conspicuous oblong fovea. S. crassipes is of about the same size as large S. (Pentarthrum)
prohaa, Sharp, from the Hawaiian Islands.
92. Stenotrupis parallela, n. sp.
Very elongate, narrow, filiform, depressed, shining; black, the antennze and tarsi, and
sometimes the rostrum also, ferruginous or obscure ferruginous. Head and rostrum rather
closely, conspicuously punctate, the latter a little smoother in ?;. rostrum curved,
widened towards the apex, in ¢ moderately stout and barely two-thirds the length of
the prothorax, in ? considerably longer and more slender; head oblong-oval, stout,
ab
CHAMPION—COLEOPTERA ; CURCULIONIDA 469
becoming subparallel-sided immediately before the abrupt basal constriction, the post-
ocular portion twice or thrice the length of the eyes, the latter depressed ; antennze
rather stout, joints 3—5 of the funiculus transverse, the club short-ovate, moderately
large. Prothorax about twice as long as broad, gradually narrowing forwards to the
subapical constriction, feebly rounded at the sides posteriorly; rather closely punctate,
except along a narrow space down the middle. Elytra very elongate, scarcely so wide as
the prothorax in ¢, parallel to near the apex, and there bluntly rounded and narrowly
explanate ; closely punctate-striate, the striz shallow, the interstices flat and faintly trans-
versely rugulose. Beneath very shining, sparsely, minutely punctate ; the head transversely
rugose, the lateral portions of the metasternum and first two ventral segments depressed
and densely alutaceous, the metasternum sharply canaliculate, the fifth segment densely
punctured ; the second segment obsoletely sulcate, and the fifth broadly depressed and
densely rugose, in $. Tibize distinctly mucronate at the inner apical angle, the anterior
pair feebly sinuate and ciliate within. ‘Tarsi short, stout, joint 3 bilobed.
Length 3—42 mm. (¢9).
Loc. Seychelles: Mahé, Silhouette, Praslin.
Fifty specimens, varying greatly in size and in the development of the rostrum in
the male. Apparently one of the commoner Cossonids of the Seychelles. Silhouette,
found near Mont Pot-a-eau, ca. 1500 feet, and in the forests above Mare aux Cochons,
including one specimen taken from the decayed head of a felled Verschaffeltia-palm.
Mahé, from the high forest behind Trois Fréres (1500—2000 feet), the Mare aux Cochons
district (1000—2000 feet), and the forest above Cascade Estate. Praslin, from the
Coco-de-mer forest in the Vallée de Mai, Cotes d’Or Estate. The present species is very
like the New Zealand S. (Pentarthrum) wollastonana, Sharp, but has a much longer
prothorax, a more coarsely punctured head, a smaller antennal club, and a broader third
tarsal joint. From the Malayan S. crassifrons, Woll., the type of the genus, S. parallela,
may be known by its more elongate prothorax, the stouter antennz, with less acuminate
club, and the more parallel elytra.
93. Stenotrupis filum, n. sp.
g. Very elongate, narrow, subcylindrical, rather convex, shining; black, the antennze
and tarsi ferruginous, the femora and tibize piceous. Head and rostrum closely punctate,
together about as long as the prothorax, the rostrum stout, curved, widened behind the
middle and at the apex, the head oblong, faintly sulcate between the rather small eyes,
the space between the latter and the basal constriction very little longer than the
eyes; antenne rather stout, the club short-ovate, moderately large. Prothorax very
elongate, gradually narrowing from about the basal third to the subapical constriction ;
closely punctate, except along a narrow space down the middle. Elytra elongate, at the
base as wide as the prothorax, very gradually narrowing to the conjointly rounded
apices ; shallowly punctate-striate, the interstices flat and transversely rugulose. Beneath
closely, finely punctate, the metasternum depressed laterally; ventral segments un-
impressed, 5 densely punctate, convex. ‘Tibiee mucronate at the inner apical angle.
Third tarsal joint bilobed.
60—2
470 PERCY SLADEN TRUST EXPEDITION
Length 3 mm.
Loc. Seychelles: Silhouette.
One male, found in the forest above Mare aux Cochons. This insect is of the size of
the smaller examples of S. parallela, differmg from that species in its more convex,
subeylindrical shape, and in having the head shorter and less inflated, the elytra relatively
shorter, the ventral segments 2 and 5 unimpressed (2), and the sides of the metasternum
less depressed and smoother.
94. Stenotrupis biformis, n. sp.
Very elongate, narrow, subfusiform, somewhat depressed, shining, ferruginous, the
apex of the elytra sparsely pilose. Head and rostrum closely, minutely punctate, the
latter much smoother and more shining in ?; rostrum (¢) feebly curved, very stout,
rapidly widened outwards, barely twice the length of the greatly-developed, exposed
portion of the head, (?) very slender, widened at the apex, and nearly four times the
length of the head, the antennze inserted at basal third or fourth; head constricted far
behind the eyes, broad and somewhat oval in 7, smaller, transverse, and more narrowed
anteriorly in ?, the post-ocular space twice the length of the eyes in g, the latter
depressed; antennze with a rather large abrupt club, comparatively stout in 3%, slender
in ?. Prothorax much longer than broad, subconical, rounded at the sides posteriorly,
constricted just before the apex; densely, minutely punctate, with an indication of a
smooth median line. Scutellum transverse. EHlytra elongate, subparallel to near the
apex; closely, very finely punctate-striate, the interstices flat, faintly transversely
rugulose. Beneath closely, minutely punctate ; ventral segments 1 and 2 deeply sulcate,
and 5 foveate, in g. Third tarsal joint feebly lobed.
Length 1,%—275 mm. ($$).
Loc. Seychelles: Silhouette.
One male and two females, found in the forest above Mare aux Cochons. In this
insect the head and rostrum are so dissimilarly formed in the two sexes that the male
could easily be taken for a different species; the female, too, has the anterior portion of
the rostrum unusually prolonged. The puncturing of the prothorax is extremely fine and
close. The general shape of the body is very like that of a small Rhizophagqus.
95. Stenotrupis tarsalis, n. sp.
Very elongate, narrow, subfusiform, depressed, alutaceous and opaque above, shining
beneath ; ferruginous, the eyes black, clothed when fresh with excessively minute hairs.
Head and base of rostrum densely, the rest of the latter more finely, punctate, the apical
portion of the rostrum smoother in 2; head in ¢ prolonged in front of the basal con-
striction, in ? shorter, the post-ocular portion a little longer than the eyes in ¢ and of
about the same length in 9, the inter-ocular fovea distinct, the eyes small and depressed ;
rostrum together with the head about as long as the prothorax, subcylindrical, slightly
widened at the tip, moderately stout in 3g, slender and more curved in §, the antennz
slender, inserted near the base in ?, and at the basal third in ¢, the club abrupt.
CHAMPION—COLEOPTERA ; CURCULIONIDA 471
Prothorax elongate, narrowing from about the basal third to the deep subapical con-
striction, rounded at the sides posteriorly; closely, finely punctate. Elytra elongate,
subparallel in their basal half; finely, shallowly punctate-striate, the mterstices flat and
densely rugulose. Beneath sparsely, finely punctate, the sides of the metasternum and
first ventral segment depressed and rugulose, the metasternum canaliculate; ventral
segments 1 and 2 excavate down the middle in g. Third tarsal joint greatly dilated,
bilobed.
Length 243—3 mm. (9).
Loc. Seychelles: Mahé.
Upwards of fifty examples of this species were obtained by Mr Scott in the forest
above Cascade and in the Mare aux Cochons district. S. tarsalis is easily distinguishable
from the allied forms by its long head, the greatly developed third tarsal joint, and the
uniformly ferruginous body, the upper surface of which is dull, alutaceous, and micro-
scopically pubescent, and the lower surface shining. It bears some resemblance to the
much larger S. (Pentarthrum) prolixa, Sharp, of the Hawaiian Islands.
96. Stenotrupis caliginosa, n. sp.
Very elongate, narrow, subparallel-sided, strongly depressed, opaque above, shining
beneath; piceous or nigro-piceous, the tip of the rostrum obscurely rufescent, the
antennz and tarsi rufo-testaceous; clothed with an excessively fine pubescence, which
becomes longer and more conspicuous on the apical portion of the elytra. Head and
rostrum very densely, rugosely punctate, the apex of the latter smoother and shining
in $; head strongly exserted, oblong, constricted at the base, the post-ocular space from
two to three times the length of the eyes, the latter small and depressed; rostrum (f)
rather long, feebly curved, moderately stout, widened towards the base and apex,
(?) longer and more slender; antennz slender, the club abrupt, ovate. Prothorax much
longer than broad, subconical, rounded at the sides posteriorly, strongly constricted before
the apex; very densely, minutely, rugosely punctate, sometimes with an indication of a
smooth. median line. Elytra elongate, subparallel in their basal half; closely, finely
punctate-striate, the interstices flat, densely rugose. Beneath closely, finely punctate ;
metasternum canaliculate ; ventral segments 1 and 2 suleate down the middle, and 5 with
a large rounded fovea, in ¢. Anterior coxe narrowly separated. Legs short; intermediate
and posterior tibize subtriangular ; third tarsal joint feebly lobed.
Length 24—24 mm. (¢9).
Loc. Seychelles: Mahé, Silhouette.
Seven specimens, mostly from the high forest: Silhouette, near Mont Pot-a-eau,
ca. 1500 feet; Mahé, high forest of Morne Blanc, forest above Cascade Estate, and
Mare aux Cochons district. A very depressed, parallel-sided form, opaque and extremely
finely pubescent above (when viewed under a powerful lens) and shining beneath, the
head elongated and strongly exserted. This species bears some resemblance to the
Malayan Leptomimus fragilis, Woll.*, an insect living in Bambusa; but it is much less
* The sexes of this insect were described by Wollaston as different species, Z. delicatulus being the ?.
472 ’ PERCY SLADEN TRUST EXPEDITION
rugose, the head, rostrum, and antenneze are differently formed, &c. The conspicuously
pubescent apex of the elytra is a character common to various members of the genera
Stenotrupis and Rhinanisus.
97. Stenotrupis conicicephala, n. sp.
Very elongate, depressed, filiform, subopaque; piceous, the humeri rufescent, the
antennee and tarsi rufo-testaceous, the femora and tibize obscure ferruginous ; clothed with
excessively minute hairs, which become longer at the apex of the elytra. Head and base
of the rostrum very densely and rugulosely, the rest of the latter more finely, punctate,
the rostrum (except at the base) much smoother in 2; rostrum with the base, apex, and
inter-antennal portion distinctly thickened, long, slender, and feebly curved, in the ¢ very
slender and as long as the prothorax, the antenne inserted near the base; head strongly
exserted, subconical, constricted far behind the eyes, the latter small and depressed ;
antennze slender, the club abrupt. Prothorax much longer than broad, subconical,
rounded at the sides posteriorly, and constricted immediately before the apex ; densely,
very finely punctate, with an indication of a smooth median line. Scutellum transverse,
minute. Elytra elongate, as wide as the prothorax, subparallel to near the apex; closely
and finely punctate-striate, the interstices very narrow, flat, and rugulose. Beneath
closely, minutely punctate; ventral segments 1 and 2 sulcate, and 5 foveate, in ¢. Third
tarsal jomt narrow, feebly lobed.
Length 2—2,, mm. (29).
Loc. Seychelles: Silhouette, Mahé.
Five specimens: found in Silhouette in the forest near Mare aux Cochons; in Mahé
in the forest above Cascade Estate and in the Mare aux Cochons district. Narrower and
more shining than S. caliginosa, the prothorax less rugose (the punctures separate one
from another), the elytral interstices smoother, the rostrum more slender in both sexes (in
the male not thicker than in the female of S. caliginosa). The extremely minute hairs
give a sericeous appearance to the upper surface. S. conicicephala was found in the high
forest in both islands.
98. Stenotrupis nemoralis, n. sp.
Very elongate, narrow, somewhat fusiform, depressed, shining; varying in colour
from piceous to ferruginous, the head and prothorax often darker than the elytra, the
rostrum, antenne, and legs ferruginous or rufo-testaceous ; the elytra finely pubescent at
the apex. Head closely, the rostrum minutely, punctate, the latter smoother in ?;
rostrum feebly curved, long, and slender, thickened at the tip, i very slender and
about as long as the prothorax; head subconical, constricted far behind the eyes, the
latter depressed ; antennze slender, the club abrupt. Prothorax much longer than broad,
a little wider than the elytra, subconical, feebly rounded at the sides posteriorly, con-
stricted just before the apex; closely punctate. Elytra elongate, very gradually
narrowing from the base; closely, finely punctate-striate, the interstices flat. Beneath
closely punctate; metasternum obsoletely or feebly canaliculate; ventral segments 1 and 2
CHAMPION—COLEOPTERA; CURCULIONIDA 473
slightly hollowed down the middle, and 5 transversely excavate, in ¢. Third tarsal joint
very little wider than the second.
Length 2—24 mm. ($9).
Loc. Seychelles: Silhouette.
Seventeen specimens, from the Mare aux Cochons plateau or the forest near by,
including two from decayed heads of felled Verschaffeltva-palms. Closely related to
S. polita, differing from it in the more closely punctured head and prothorax, the somewhat
attenuate elytra, which are less distinctly pubescent at the apex, and the obsoletely cana-
liculate metasternum. The more elongate, less coarsely punctured prothorax, the straighter
rostrum, and the narrower third tarsal joint, distinguish S. nemoralis from S. silvicola.
99. Stenotrupis polita, n. sp. ;
Very elongate, depressed, filiform, shining ; nigro-piceous (ferruginous or brown when
not fully coloured), the humeri, rostrum, and legs brown or rufescent, the antennee and
tarsi rufo-testaceous ; the elytra pilose at the apex. Head rather sparsely, the rostrum
very finely, punctate, the latter smoother in ?; rostrum thickened at the tip, feebly
curved, long, and slender, in the ¢ very slender and as long as the prothorax; head
oblongo-conic, constricted far behind the eyes, the latter depressed; antennze slender, the
club abrupt. Prothorax elongate, subconical, rounded at the sides posteriorly, the apical
collar sharply defined ; sparsely, finely punctate, except along a narrow smooth space
down the middle. Elytra elongate, parallel to near the tip; closely, finely punctate-
striate, the interstices flat and rugulose. Beneath closely, finely punctate; metasternum
suleate, the lateral portions, and those of the first ventral segment also, depressed,
opaque, and rugulose; ventral segments 1 and 2 suleate, and 5 transversely excavate,
in g. ‘Third tarsal jot narrow, feebly lobed.
Length 2—22 mm. (9).
Loc. Seychelles: Silhouette, Mahé.
Seventeen specimens, found in Silhouette in the forest above Mare aux Cochons, and
in Mahé in that above Cascade. The chief characters of this species are: the shining,
depressed body, the long, slender rostrum (especially in ¢), the sparsely punctured head
and prothorax, the elongate prothorax, and the long parallel-sided elytra, with the apex
conspicuously pubescent. The examples before me vary in size, one female from Mahé
being much smaller than the rest. The Malayan S. eailis, Pasc. (2, not $*), is an allied
form, with much coarser sculpture.
100. Stenotrupis sericata, n. sp.
Very elongate, narrow, depressed, subfusiform, subopaque, alutaceous, the rostrum
shining ; piceous, the tarsi and the base of the antenne ferruginous. Head and base of
rostrum densely, the rest of the latter more finely, punctate, the outer portion of the
rostrum smoother in ?; head in $ moderately prolonged in front of the basal constriction,
a little shorter in ?, the post-ocular portion about as long as the small depressed eyes, the
inter-ocular fovea very small; rostrum curved, subcylindrical, together with the head as
* The insect described as the male belongs to a different genus, Coptus, Woll.
ATA PERCY SLADEN TRUST EXPEDITION
long as the prothorax, slightly widened at the tip, moderately stout in g, slender in ?;
antennee slender, the club ovate and rather large. Prothorax considerably longer than
broad, narrowing from about the basal third to the subapical constriction, rounded at the
sides posteriorly ; closely punctate. Elytra very elongate, subparallel in their basal half,
conjointly rounded at the apex; rather coarsely punctate-striate, the interstices narrow,
rugulose, 3 becoming convex on the apical declivity. Beneath dull, closely, finely
punctate, the sides of the metasternum depressed and rugulose; ventral segments 1 and 2
suleate, and 5 transversely depressed, in g. Anterior coxze separated by less than the
width of the coxa. Third tarsal joint slightly wider than the second.
Length 3—32 mm. (¢$).
Loc. Seychelles: Silhouette.
One pair, from the forest near Mont Pot-a-eau, found in August, 1908. Duller than
S. lodoiceicola, the rostrum shorter and less widened towards the base, the anterior
portion of the head shorter in the male, the antenne shorter, with the joints of the
funiculus more closely articulated, the prothorax less elongate (the elytra thus appearing
relatively longer), the third elytral mterstice more convex before the apex. The aluta-
ceous sculpture gives a sericeous appearance to the surface, and when the insect is viewed
under the microscope a few excessively minute hairs are visible on the apical portion of
the elytra.
101. Stenotrupis nitidula, n. sp.
?. Very elongate, subfiliform, somewhat robust, moderately flattened above, shining;
nigro-piceous, the antennze and tarsi rufo-testaceous. Head and base of rostrum closely,
minutely, the rest of the latter sparsely, punctate; rostrum moderately long, curved,
thickened at the tip, not very slender; head subconical, constricted at the base, the
post-ocular space as long as the eyes, the latter depressed; antennz slender, the club
large. Prothorax elongate, subconical, rounded at the sides posteriorly, constricted
before the apex; closely, rather coarsely punctate. Scutellum small. Elytra elongate,
subparallel to near the apex; rather coarsely punctate-striate, the interstices flat and
transversely rugulose. Beneath closely, the sides of the metasternum rugosely, punctate.
Third tarsal jomt moderately dilated.
Length 3 mm.
Loc. Seychelles: Mahé.
One specimen, from the high damp forest at the summit of Pilot, at an elevation of
over 2000 feet. Near S. silvicola, but more robust and less depressed, the rostrum (?) as
stout as in the males of many of the allied forms, the antennze wholly rufo-testaceous,
with a large club, the puncturing of the prothorax and elytra coarser, the prothorax more
elongate, the sides of the metasternum more rugose, the third tarsal joint relatively less
dilated, &c. Narrower, smoother, and more shining than S. sericata, the rostrum (@)
stouter, &c. More depressed than S. filum, from Silhouette, the male only of which is
known, the antennze more slender and with a larger club, the prothorax much more
narrowed anteriorly, &c. The example described has the first ventral segment un-
impressed down the middle, and it is, therefore, undoubtedly a female.
CHAMPION—COLEOPTERA ; CURCULIONIDA 475
102. Stenotrupis silvicola, n. sp.
Very elongate, narrow, subfusiform, depressed, shining, almost bare; black or
pitchy-black, the elytra often piceous and with the humeri rufescent, the tarsi and the
base of the antennz rufo-testaceous, the rest of the legs and the tip of the rostrum
piceous or rufo-piceous. Head rather sparsely, the rostrum very finely, punctate, the
latter smoother in $; rostrum curved, long, and slender, thickened at the tip, in 2 very
slender and about as long as the prothorax ; head oblongo-conic, constricted far behind
the eyes, the latter depressed; antennze slender, the club abrupt. Prothorax a little
longer than broad, oval, constricted before the apex ; closely, uniformly punctate. Elytra
elongate, very gradually narrowed from the base; closely, finely punctate-striate, the
interstices flat and transversely rugulose. Beneath finely, the sides of the metasternum
and of the first two ventral segments more coarsely, punctate ; metasternum canaliculate
posteriorly ; ventral segments 1 and 2 sulcate, and 5 transversely excavate, in g. Third
tarsal joint rather broad.
Length 2—24 mm. (99).
Loc. Seychelles: Silhouette, Mahé.
Eighteen specimens, all from the high forest : found in Silhouette in the forest above
Mare aux Cochons, in Mahé in the high forest of Morne Blane and Pilot, the forest behind
Trois Fréres and on Morne Seychellois, the forest above Cascade Estate, and in that on
the summit of “ Montagne Anse Major,” ca. 2000 feet. This species is nearest allied to
S. polita, differing from it in the more curved rostrum in both sexes, the relatively
shorter, uniformly, and distinctly more coarsely punctured prothorax, the less parallel,
subglabrous elytra, and the broader third tarsal joint. Two of the females are less shining
than the rest, and one of them has the disc of the prothorax broadly depressed, but they
evidently belong to S. szlvicola, as shown by the tarsal structure.
103. Stenotrupis dumetorum, n. sp.
Very elongate, narrow, subfusiform, moderately depressed, shining; ferruginous,
with the eyes black and the space between them infuscate, the body in one example
nigro-piceous. Rostrum curved, slender, slightly thickened at the tip, in nearly as long
as the prothorax, in ¢ distinctly stouter and shorter, faintly punctulate; head small,
conical, sparsely, finely punctate, constricted at a little behind the eyes, the latter small
and depressed ; antenne slender, long, the club rather large, acuminate-ovate. Prothorax
slightly longer than broad, subconical, rounded at the sides posteriorly, constricted just
before the apex ; closely, finely punctate. Elytra elongate, subparallel in their basal half;
finely punctate-striate, the interstices flat. Beneath very finely punctate; metasternum
not or feebly canaliculate posteriorly; ventral segments 1 and 2 hollowed down the
middle, and 5 shallowly foveate, in ¢. Legs slender; tibie very feebly unguiculate ;
third tarsal joint narrow, very little wider than the second.
Length 13—2 mm. (f9).
Loc. Seychelles: Mahé, Silhouette, Praslin.
SECOND SERIES—ZOOLOGY, VOL. XVI. 61
476 PERCY SLADEN TRUST EXPEDITION
Twenty specimens, eleven of which are noted as having been taken from the leaf-
bases of growing Stevensonia-palms. Silhouette: several specimens were found in the
forest above Mare aux Cochons, one was taken from the leaf-bases of a growing palm in
the high forest, and one from fungus at an elevation of over 2000 feet. Mahe: a number
of specimens were taken from between leaf-bases of a growing Stevensonia-palm on summit
of Mount Sebert, one was found in a similar situation near Morne Blanc, and one was
found on Cascade Estate. In Praslin the species was found in the Coco-de-mer forest in
the Vallée de Mai.
This slender, minute Cossonid agrees with S. sz/vicola in having a narrow conical
head, &c., but differs from that species in its smaller size and more slender build, the less
elongate, more finely punctured prothorax, and the much narrower third tarsal jomt; the
head, too, is shorter and the eyes smaller. The Madagascar insect in the British Museum
doing duty for Proéces mgrifrons, Boh., superficially resembles the light-coloured examples
of S. gracilis, but it has a 7-jointed funiculus.
MIcROTRUPIS, n. gen.
Rostrum cylindrical, curved, not thickened at the tip, the scrobes inferior, the
antennee inserted at or behind the middle; antenne slender, with a 5-jointed funiculus
and a small club; head short, small, constricted immediately behind the eyes, the latter
small, depressed or prominent ; prothorax subquadrate or subcylindrical, about as long as
broad, feebly constricted before the apex; elytra elongate; coxee narrowly separated ;
metasternum elongate; legs short; tibiz feebly unguiculate; tarsi short, third jomt
small, not much wider than second; body linear or subfusiform, depressed, shining,
glabrous or finely pilose: species very small.
Type, WM. longipennas.
The three minute insects placed under Microtrupis are nearly related to some of the
smaller species included in Stenotrupis ; but they have a small head, a cylindrical rostrum,
a less elongate prothorax, &c. The basally constricted head, the depressed body, and the
shorter tarsi distinguish Microtrupis from the New Zealand Hntiwm, Sharp; and the
cylindrical rostrum, feebly constricted prothorax, &c., from Rhinanisus (type Pentarthrum
fulvicorne, Broun), even as amended to include many minute Tropical American forms.
M. longipennis and M. puncticeps agree well inter se; the third species, M. piligera, has
prominent eyes, confusedly punctate-striate elytra, and a sparsely pilose body, but it can
remain under ./icrotrupis for the present.
104. Microtrupis longipennis, n. sp.
Elongate, filiform, depressed, shining; varying in colour from piceous with the
rostrum, antennee, and legs ferruginous to entirely ferruginous. Rostrum feebly curved,
cylindrical, rather slender, much shorter than the prothorax, minutely punctate, the
antenne. inserted at the middle; head constricted at a little behind the small non-
prominent eyes, and closely punctured between them, the short anterior portion narrow ;
antennze with a moderately large club. Prothorax very little longer than broad, sub-
a ——
CHAMPION—COLEOPTERA; CURCULIONIDA AT7
quadrate, rounded at the sides posteriorly, constricted before the apex; finely punctate.
Scutellum minute. Elytra very elongate, scarcely wider than the prothorax, gradually
narrowing from about the basal fourth; finely punctate-striate, the interstices flat.
Beneath finely, rather sparsely punctate; metasternum elongate, finely canaliculate
posteriorly ; ventral segments 1 and 2 sulcate, and the metasternum broadly depressed
down the middle, in g. Legs short, slender.
Length 15—12 mm. (¢9).
Loc. Seychelles: Silhouette, Mahé.
Five specimens, found in Silhouette in the forest near Mare aux Cochons, and in
Mahé at Cascade Estate and in the Mare aux Cochons district. A minute, linear,
depressed form, with a small, short head, a subquadrate prothorax, and relatively very
elongate elytra. :
105. Microtrupis puncticeps, n. sp.
Moderately elongate, linear, slightly depressed, shining, obscure ferruginous, glabrous.
Rostrum curved, cylindrical, moderately stout, much shorter than the prothorax, the
basal portion coarsely, the apical portion finely, punctate; head short, narrow, coarsely
punctured between the eyes, and constricted immediately behind them, the eyes small
and slightly prominent; antennze with a rather large abrupt club. Prothorax sub-
quadrate, convex, about as long as broad, rounded at the sides posteriorly, and constricted
immediately before the apex; closely, somewhat coarsely punctate. Scutellum minute.
Elytra moderately long, subparallel to near the tip, finely punctate-striate. Beneath
rather sparsely, finely punctate; metasternum obsoletely canaliculate posteriorly.
Length 13 mm. (@).
Loc. Seychelles: Silhouette.
Two specimens, found in the forest at Mare aux Cochons. More robust and a little
larger than M. longipennis, the rostrum (?) stouter, the elytra parallel-sided and relatively
less elongate, the prothorax subquadrate and almost as coarsely punctured as in WM.
piliyera. The ventral segments are unimpressed, and the examples described are therefore
assumed to be females.
106. Microtrupis piligera, n. sp.
Moderately elongate, somewhat fusiform, feebly convex, shining, ferruginous, the
eyes black; clothed with very fine, scattered, erect, pallid hairs. Rostrum cylindrical,
curved, nearly as long as the prothorax, slender, very finely punctate, the antenns
inserted just behind the middle; head short, small, closely punctate between the eyes,
and constricted immediately behind them, the eyes small, prominent ; antennz slender,
the club abrupt and rather large. Prothorax about as long as broad, the sides rounded
and converging anteriorly to the faint subapical constriction; somewhat coarsely, closely
punctate. Scutellum transverse, very small. Hlytra moderately long, subparallel in their
basal half, a little produced at the apex; closely, subseriately punctate, the interstices flat
and here and there transversely rugulose.
61—2
478 PERCY SLADEN TRUST EXPEDITION
Length 14 mm. (¢ ?).
Loc. Seychelles: Silhouette.
One specimen, found at Mare aux Cochons on Sept. 6th, 1908. Recognizable by its
comparatively long rostrum, prominent eyes, feebly constricted, rather coarsely punctured
prothorax, and confusedly seriato-punctate elytra, and also by the fine erect pilosity of the
upper surface. The under surface has not been examined, for fear of damage to the
vestiture. Lasiotrupis clavigera is similarly pilose.
LASIOTRUPIS, n. gen.
Rostrum subcylindrical, curved, the antenne inserted towards the apex in ¢ and at
a little beyond the middle in 9, the scrobes inferior and extending backward to between
the eyes, the latter lateral, in part visible from above, the head feebly constricted immedi-
ately behind them ; antennze long, slender, the funiculus 5-jointed, the club large, ovate,
annulate; prothorax subquadrate; scutellum minute; elytra wider than the prothorax,
narrowing from near the base, moderately long, punctato-sulcate ; anterior coxze small,
separated by about their own width; intermediate and posterior rather widely separated ;
metasternum moderately long; ventral segments 1 and 2 long, connate, 3 and 4 very
short ; legs short, slender; tibize without visible uncus at the apex ; tarsi short, slender,
joint 3 simple, not wider than 2, the claw-joint rather stout, the claws divergent ; body
subfusiform, shining, sparsely pilose: species minute.
Type, L. clavigera.
The minute insect from which the above diagnosis has been drawn up has the general
facies of a Stenomimus or Tytthomimus; but the funiculus is 5- (not 7-) jointed ; the
antenne are long and slender, with an unusually large club; the tibie (as in Micro-
himatium) are apparently unarmed at the apex; and the third tarsal joint is simple.
The absence of ventral depression or fovea in the male is an unusual character amongst the
Cossonids. The hairs on the upper surface are not easily seen, except on the elytra,
and easily abraded. The specimens with somewhat gibbous rostrum and the antennze
inserted towards the tip are assumed to be males.
107. Lasiotrupis clavigera, n. sp.
Moderately elongate, feebly convex, shining ; ferruginous or obscure ferruginous, the
eyes black ; sparsely clothed above and beneath with rather long, fine, stiff, pallid hairs,
those on the elytra setiform and arising from the punctures of the strize. Rostrum (9)
shorter than the prothorax, depressed at the base and somewhat gibbous between the
points of insertion of the antennz, sparsely, faintly punctulate, the short apical portion
a little wider than the rest, (?) a little longer, simply curved, and with the apical portion
relatively more elongate ; antennze with jomts 3—5 of the funiculus transverse, the club
nearly as long as the funiculus. Prothorax subquadrate, about as long as broad, abruptly
narrowed just before the feeble apical constriction; closely, rather coarsely punctate.
Elytra considerably wider than the prothorax, parallel at the base, narrowing from about
the basal fourth ; punctato-sulcate, the interstices narrow, convex. Beneath alutaceous,
CHAMPION—COLEOPTERA ; CURCULIONIDA A79
very sparsely, the sides of the metasternum closely, punctate; metasternum canaliculate
down the apical half; ventral segments unimpressed in both sexes.
Length 14—14 mm. ($9).
Loc. Seychelles: Mahé, Silhouette.
Eight specimens, six of which were found in the Mare aux Cochons district of Mahé,
while one was found on Cascade Estate (Mahé), and one at the Mare aux Cochons of
Silhouette.
MIcROHIMATIUM, n. gen.
Rostrum feebly curved, moderately stout, parallel-sided, convex above, as long as the
prothorax in ?, shorter and a little thicker in 3, the antenne inserted at about the middle,
the serobes oblique, inferiorly placed, descending to the eyes beneath; eyes lateral, not
prominent ; head small; antennze with a 5-jointed funiculus, joint 1 stout, 2—5 small,
short, the club small, ovate, annulate, the scape longer in ? than in 3; prothorax oval ;
scutellum small; elytra oblong ; coxze small, widely separated; metasternum long, unim-
pressed, the narrow episterna almost covered by the elytra; ventral segments free, 3 and
4 extremely short, the first suture curved, the others straight; legs short : tibize rounded
and unarmed at the outer apical angle, and with a very minute uncus arising from the
apical margin near the inner angle; tarsi with joints 1 and 2 short, 3 bilobed, pilose
beneath, the claws long; body moderately elongate, depressed, rugulose, opaque, finely
pubescent, winged: species minute.
Type, MW. pubescens.
The type of this genus is a minute insect not very much larger than the European
Alaocyba carinulata, a species placed by Wollaston in Cossonina, but rejected from that
group by recent writers. It is very different from any of the “ Pentarthrides” described
by that author. The elytral vestiture is suggestive of that of the genus Himatium, Woll.
The insect was found in the leaf-bases of a growing ¢ Coco-de-mer palm, Lodoicea
seychellarum.
108. Microhimatium pubescens, n. sp. (Pl. 24, figs. 45 ¢, 45a; antenna.)
Moderately elongate, rather broad, depressed ; reddish-brown, the prothorax darker,
the antenne and legs ferruginous; above and beneath opaque, densely rugulose, clothed with
rather long, fine, adpressed, yellowish pubescence. Rostrum rugulose to the tip, consider-
ably longer in $ than in g. Prothorax about as long as broad, rounded at the sides,
gradually narrowed anteriorly, closely, mimutely punctate. Elytra a little wider than,
and scarcely twice the length of, the prothorax, subparallel at the base, conjointly rounded
at the apex ; shallowly, minutely striato-punctate, the interstices flat. Beneath minutely
punctate.
Length 1—1 4 mm. (f9).
Loc. Seychelles: Praslin.
Two females and one male, found in the Vallée de Mai, Nov. 28th, 1908, in the
leaf-bases of a growing g Lodoicea-palm.
A480 PERCY SLADEN TRUST EXPEDITION
Sect. Cossonides.
HIMATINUM.
Himatium Wollaston, Trans. Ent. Soc. Lond., 1878, p. 461 [nec Imatium, Clark,
1860 (emend. Himatiwm, 1871), Halticidee].
Himatinum Cockerell, Ent. News, xvii. p. 243 (1906).
The few described species of this genus are from India, 8. Africa, and North and
Central America. The three Seychelles forms are related to the one known from South
Africa. It is perhaps convenient to adopt Cockerell’s emendation of Wollaston’s name*;
Clark, however, wrote Imatiwm.
109. Himatinum confluum, n. sp.
Elongate, depressed, moderately shining ; nigro-piceous, piceous, or ferruginous, clothed
with short, stiff, decumbent, yellowish pubescence, the hairs on the elytra uniseriately
arranged down each interstice. Rostrum (¢) stout, barely half the length of the prothorax,
coarsely, rugosely punctured at the base and along the sides, the punctures longitudmally
confluent, the apical portion smooth along the middle from the transverse inter-antennal
fovea to the tip, (?) narrower and slightly longer. Prothorax oblongo-conic, longer than
broad, constricted towards the apex, rounded at the sides posteriorly ; coarsely, densely
punctate, the punctures becoming longitudinally confluent towards the outer margin.
Elytra elongate, subparallel in their basal half, in fully-developed males barely as wide as
the prothorax ; narrowly punctato-sulcate, the interstices flat, rugose, and irregularly
uniseriate-punctate. Beneath coarsely, densely punctate; ventral segments 1 and 2
broadly flattened down the middle, 1 depressed in the centre behind in both sexes.
Femora and tibiz stout; intermediate and posterior tibiee subangularly dilated on the
inner side before the middle in g, simply triangular in 9.
Length 24—3 mm. (29).
Loc. Seychelles: Mahé.
Over 70 specimens, almost all from the forest near Morne Blanc (some of them most
probably from under bark of a dead “ Bois montagne” tree, Campnosperma seychellarum
March.) : one was found in the forest above Cascade Estate. The specimens vary in size ~
and colour. The more shining surface, the sparser pubescence, the coarsely sculptured
prothorax, and the larger size and more robust build, readily distinguish H. conflwwm from
the other Seychelles members of the genus. Its smaller size, less rugose, shining upper
surface, and shorter rostrum (especially in ?), separate H. confluwm from the S. African
H. variolosum, Woll., and the less convex body from the Indian H. pubescens, Woll.
110. Himatinum rugipenne, n. sp.
Elongate, depressed, feebly shining, the elytra dull; obscure ferruginous or ferruginous,
thickly clothed with short, stiff, decumbent, ochreous pubescence, the hairs on the elytra
uniseriately arranged down each interstice. Rostrum (¢) short, broad, coarsely, rugosely
* The substitution of Wollastoniella for Wollastonia, Horn (nec Heer), is not required, the latter name
being a synonym of Zomolips, Woll.
CHAMPION—COLEOPTERA ; CURCULIONIDA: 481
punctate, the punctures longitudinally confluent, smoother along the middle from the
inter-antennal fovea to the tip. Prothorax much longer than broad, oblongo-conic, con-
stricted near the apex, and rounded at the sides posteriorly; densely, finely punctate.
Elytra subparallel in their basal half, of about the same width as the prothorax ; narrowly
punctato-suleate, the interstices densely rugose and closely uniseriate-punctate. Beneath
shining, densely, moderately coarsely punctate; ventral segment 1 broadly excavate down
the middle in 2, foveate at the apex in 2. Tibiee subtriangular in both sexes.
Length 13—24 mm. (99).
Loc. Seychelles: Mahé, Silhouette, Praslin.
Sixty-five specimens, twenty-nine of which are from Praslin, from the Coco-de-mer
forest in the Vallée de Mai: only one is from Silhouette, from under bark of a fallen twig
of the endemic “Bois Rouge” (Wormia ferruginea) in the forest-just above Mare aux
Cochons : in Mahé the majority of specimens are from under the bark of fallen and rotting
trunks of “ Bois Rouge” in the high forest between Trois Fréres and Morne Seychellois,
1500—2000 feet ; a few also are from the forest above Cascade Estate.
111. Mimatinum breviusculum, n. sp.
Moderately elongate, rather convex, opaque; obscure ferruginous, thickly clothed
_ with short, curled, stiff, ochreous pubescence, the hairs on the elytra semi-erect and uniseri-
ately arranged down each interstice. Rostrum short, broad, about twice as long as wide,
somewhat flattened above, densely, rugulosely punctate to the tip. Prothorax longer
than broad, gradually narrowing to the apical constriction, rounded at the sides posteriorly ;
very densely, minutely punctate. Elytra a little wider than the prothorax, comparatively
short, subparallel in their basal half; deeply punctate-striate, the interstices densely
rugulose. Beneath moderately shining, densely punctate.
Length 12 mm. (¢ 2).
Loc. Seychelles: Silhouette.
One specimen, found in the forest near Mare aux Cochons, Sept. 1908. Separable at
once from H. confluum and H. rugipenne by its very small size, shorter, more convex
form, opaque upper surface, more erect vestiture, the less rugose, somewhat flattened
rostrum, and the very dense minute punctuation of the prothorax. H. breviusculum has
much the facies of a small Monotoma. The joints of the funiculus are so closely articulated
that it is difficult to make out (as in some of the small American forms) whether there are
six or seven present, apparently the latter.
MELARRHINUS.
' Melarhinus Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 466, 547.
The single known species of this genus is from Madagascar. A closely allied form
has been found in Mahé.
112. Melarrhinus incrustatus, n. sp.
fg. Elongate-fusiform, somewhat depressed, opaque above ; nigro-piceous, the base
of the antennze and the fourth tarsal joint ferruginous; the upper surface very densely,
482 PERCY SLADEN TRUST EXPEDITION
rugosely punctate, the depressions filled with an earthy-looking incrustation, which
hides the minute, yellowish, hair-like scales arising from the punctures. Rostrum very
stout, a little longer than the exposed portion of the head, obsoletely canaliculate at
the base; eyes large and prominent ; antennze stout, joints 2—7 of the funiculus strongly
transverse, the club short-ovate. Prothorax longer than broad, narrowed anteriorly,
abruptly constricted before the apex, subtruncate at the base, obsoletely carinate down
the middle. Elytra elongate, much wider than the prothorax, subparallel in their basal
half, produced at the apex, the humeri rounded; shallowly punctato-suleate, the densely
rugose interstices appearing slightly raised. Beneath moderately shining, closely punctate;
metasternum suleate down the middle posteriorly ; ventral segment 1 deeply excavate,
5 densely punctate and clothed with moderately long hairs. Legs stout; intermediate
and posterior tibize bimucronate at the inner apical angle.
Length 35 mm.
Loc. Seychelles: Mahé.
One specimen, from the Mare aux Cochons district, captured in January or
February, 1909. Smaller and much less robust than M. nigritus, Woll., the conspicuous
median sulcus of the rostrum only just indicated, the prothorax and elytra subtruncate
at the base, the seriate punctures on the latter not so coarse, the intermediate and
posterior tibize bimucronate at the tip. Wollaston’s type is probably of the same sex,
male.
PHLG@OPHAGOSOMA.
Phleophagosoma Wollaston, Trans. Ent. Soc. Lond., 18738, pp. 28, 465, 545.
Two species were originally included in this genus, both from Japan, and P. minutum
given as the type, Wollaston subsequently adding, under a subgenus Amorphorhynchus,
many others from the Malayan Is., Ceylon, and New Zealand. Two forms from the
Seychelles are related to the latter. P. dubiwm, Gahan, from Christmas Island and
the Seychelles, is here placed under a new generic name, Khyncolosoma.
113. Phle@ophagosoma comceicolle, n. sp.
Elongate, fusiform, feebly convex, shining; nigro-piceous, the legs and antennz
obscure ferruginous. Rostrum stout, curved, convex, almost parallel-sided (the apical
portion, if anything, narrower than the basal portion), considerably longer than the
width of the head with the eyes, more finely punctured than the inter-ocular portion
of the head; eyes depressed, rather large; head narrow, parallel-sided for a short
distance behind the eyes, and then abruptly constricted ; antennez with joints 2—7 of
the funiculus closely articulated, transverse, the club abrupt, ovate. Prothorax much
longer than broad, conical, constricted immediately before the apex and at the sides
before the base; rather sparsely punctate, except along a narrow smooth median space. .
Scutellum very small. Elytra moderately elongate, a little wider than the prothorax,
gradually narrowing from the base, conjointly rounded at the apex; crenate-striate,
the interstices faintly uniseriate-punctate. Beneath coarsely, closely, the ventral segments
CHAMPION—COLEOPTERA ; CURCULIONID AI 483
more sparsely, punctate; metasternum sulcate posteriorly. Tibize mucronate at the
inner apical angle, the anterior pair sinuate, and also ciliate towards the apex, within.
Length 3 mm. (? ?).
Loc. Seychelles: Silhouette.
One specimen, from the forest near Mont Pot-a-eau, ca. 1500 feet, VIII. 1908.
Near P. angustulum Woll., from Batchian, differing from that insect in the more coarsely
punctured rostrum, the depressed eyes, and the more sparsely punctured conical prothorax.
The rostrum has the apical portion scarcely so wide as the basal portion, the contrary
being the case in P. aldabranum. The Hawaiian P. tenue has a similarly shaped
prothorax, but it is a larger and more robust insect, with a longer rostrum and prominent
eyes.
114. Phleophayosoma aldabranum, n. sp.
Elongate, fusiform, shining, piceous. Rostrum curved, stout, convex, the apical
portion widened and much longer than parallel-sided basal portion, closely punctate ;
head convex, similarly punctured, with a small fovea between the prominent eyes, and
constricted at a short distance behind them. Prothorax longer than broad, somewhat
oval, the subapical constriction sharply defined laterally, shallow on the disc; coarsely,
closely punctate. Scutellum minute. Elytra much wider than the prothorax, elongate-
subtriangular, the apices subconjointly rounded ; coarsely punctate-striate, the interstices
transversely rugulose and closely uniseriate-punctate. Beneath coarsely, the prosternum
more rugosely, punctate ; metasternum sulcate along the posterior half, the first ventral
segment also hollowed down the middle. Anterior coxe distant. Tibiz mucronate
at the inner apical angle, the anterior pair strongly sinuate within. .
Length 44 mm. (8).
Loc. Aldabra: Takamaka, 1908 (Fryer).
One specimen. Very near the Malayan P. proximum and P. puncticolle, Woll.,
differing from both of them in having the rostrum less densely punctate, the prothorax
more feebly constricted before the apex (this being especially noticeable on the disc),
and more coarsely and not so closely punctate; the elytra are narrowed from the base,
as in P. puncticolle and not parallel as in P. prozimum. These three species will
doubtless have to be removed from Phlaophagosoma and its subgenus Amorphorhynchus,
and included under Notiosomus or Oxydema, Woll., which have a similarly formed
rostrum.
OxXYDEMA.
Oxydema Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 487, 488, 574.
The five described species of this genus are from the Malayan Islands and Ceylon.
The Ceylon insect has been found on the Cargados Islands, between the Mascarenes
and the Seychelles.
115. Oxydema fusiforme.
Oxydema fusifornus Woll., loc. cit., p. 632.
Loc. Cargados Carajos Islands. Ceylon.
One example (¢), found on Aug. 28th, 1905, on Establishment Island. This insect
SECOND SERIES—ZOOLOGY, VOL. XVI. 62
484 PERCY SLADEN TRUST EXPEDITION
has the widened apical portion of the rostrum much longer than the parallel-sided
basal portion; the head long, depressed and strongly foveate between the prominent
eyes, and constricted at some distance behind them ; the prothorax abruptly constricted
before the apex ; and the first ventral segment sulcate down the middle. Length 53mm.
There are two specimens of O. fusiforme from Ceylon in the British Museum, and
O. elongatum, Pasc., from Singalang, seems to me to be scarcely separable from them.
ORTHOTEMNUS.
Orthotemnus Wallaston, Trans. Ent. Soc. Lond., 1878, pp. 489, 575.
The type-species of this genus, O. reflexus, is widely spread in the Malayan Islands.
A second, from Silhouette, in the Seychelles, has been captured by Mr Scott. Both
insects have the apical margin of the elytra recurved, so as to appear deeply sulcate
within.
116. Orthotemnus filiformis, n. sp.
Very elongate, narrow, somewhat convex, flattened on the disc, moderately shining,
black, the antennz and legs piceous. Head and rostrum closely, finely punctate; rostrum
stout, curved, about half the length of the prothorax, convex, the convexity continued
backwards to the sharply-defined post-ocular constriction of the head, the head feebly
foveate between the eyes, the latter large and prominent. Prothorax much longer than
broad, subconical, strongly constricted before the apex; closely, somewhat coarsely
punctate, except along an abbreviated narrow space down the middle, the interspaces
finely alutaceous. Scutellum minute, narrow. Elytra elongate, of the same width as
the base of the prothorax, parallel to near the apex, the apices conjointly round and
with the margin sharply reftlexed; punctate-striate, the sutural stria sulciform, the
interstices flat and obsoletely uniseriate-punctate. Beneath feebly shining, closely,
finely punctate; anterior coxee widely separated; metasternum compressed laterally,
broadly flattened and canaliculate down the middle; first ventral segment similarly
flattened. Tibiz mucronate at the inner apical angle, the anterior pair sinuate and
ciliate ‘within.
Length 35 mm. (2).
Loc. Seychelles: Silhouette.
One specimen, from the forest near Mont Pot-a-eau, VIII. 1908. Smaller and
narrower than O. reflexus, the rostrum and antenne shorter, the prothorax conical, the
elytra parallel to near the apex.
Cossonus.
Cossonus [Clairville], Ent. Helv., i. p. 58 (1798).
A genus of world-wide distribution, particularly numerous in species in Tropical
America. Two species only are represented in the Seychelles collection before me,
both known §.-African forms.
ee ee en
oa
CHAMPION—COLEOPTERA ; CURCULIONIDAI 485
117. Cossonus mewilis.
Cossonus incivilis Fahr., Ofv. Vet.-Ak. Stockh. Foérh., 1871, p. 285.
Loc. Seychelles: Mahé. 8. Africa: Caffraria.
A single specimen, agreeing with one from Natal in the British Museum. This
insect has an elongate-triangular, deep, rugosely punctate, carinate depression on the
dise of the prothorax and a broad space on each side of it almost smooth. It was found
in the high damp forest between Morne Seychellois and Trois Fréres, 1500—2000 feet,
XII. 1908.
118. Cossonus suturalis.
Cossonus suturalis Boh., in Schénh., Gen. Cure., iv. p. 1033.
Loc. Seychelles: Silhouette. S. Africa.
One example, also agreeing with Natal specimens in the British Museum (compared
by Mr Marshall with the type), found in more open secondary jungle-country in
Silhouette, 3. VIII. 1908. This insect has a rufo-testaceous patch extending down
the basal half of each elytron, and the deep excavation on the disc of the prothorax
(which is bordered laterally by a polished space) extending forwards to the subapical
constriction, and bordered on each side by a double series of confluent foveze. C. varw-
pennis, Gahan, from Christmas Island, is a smoother, more depressed, allied form.
RHYNCOLOSOMA, n. gen.
Rostrum stout, subcylindrical, short, curved, the antennz inserted at the middle,
the scrobes deep, oblique, descending to beneath the eyes ; eyes lateral, oval, prominent, the
head constricted behind them, the latter large, globose, and deeply inserted ; antennze
short, rather stout, the scape reaching to the middle of the eyes, the funiculus 7-jointed,
joint 1 stout, 2—7 closely articulated, rapidly widened outwards, strongly transverse,
the club small, ovate, annulate, sparsely pilose, the basal half polished ; prothorax oblongo-
conic; scutellum small, prominent; elytra oblong, coarsely punctate-striate ; anterior
coxze large, moderately distant; posterior coxee very widely separated; metasternum
long, not sulcate; legs short ; tibiee moderately broad, produced into a long curved
hook at the outer apical angle, and sharply mucronate at the inner apical angle; tarsi
short, joint 3 transverse, strongly bilobed, pilose beneath, the claws minute; body
moderately elongate, rather convex, shining, subglabrous, coarsely punctured above
and beneath, winged.
Type, Phleophagosoma dubiuwm, Gahan.
The type of P. dubiwm was from Christmas Island, and the insect proves to be
common in the Seychelles. It differs in various respects from all the heterogeneous
forms included by Wollaston under Phiwophagosoma, more especially in the conspicuously
bilobed third tarsal joint, the small antennal club, which has its basal half polished,
and the sharply mucronate inner apical angle of the tibiz; and a new generic name
is therefore required for the species. Rhyncolus (type FR. ater, L.) has a differently
formed head, &c.
62—2
486 PERCY SLADEN TRUST EXPEDITION
119. Rhyncolosoma dubsum.
Phleophagosoma dubiwm Gahan, Monogr. Christmas Isl., p. 114 (1900).
Somewhat fusiform, convex, shining; nigro-piceous, reddish brown, or obscure
ferruginous. Head sparsely, the rostrum closely, finely, punctate, the punctures becoming
much coarser between the eyes, the inter-ocular space depressed, the rostrum parallel-
sided, about twice as long as broad, pilose beneath. Prothorax oblongo-conic, a little
longer than broad, rounded at the sides posteriorly, feebly constricted near the apex,
coarsely, closely punctate. Elytra moderately long, about as wide as the prothorax,
subparallel in their basal half; very coarsely punctate-striate, the interstices narrow,
convex, rugulosely punctate. Beneath very coarsely, closely punctate, the punctures
on the prosternum more or less confluent, those on the ventral segments sometimes more
scattered, the prosternum with a curved ridge in front of each anterior coxa; first
ventral segment broadly depressed down the middle in ¢.
Length 24—38 mm. (49).
Loc. Seychelles: Mahé, Silhouette, Félicité. Christmas Island (Indian Ocean).
The above description, taken from the long variable series captured by Mr Scott,
will supplement that of Mr Gahan. In the Seychelles the insect was found freely
under bark in the mountain-forests. In Silhouette a number were found at 1500 feet
or more in the forest near Mont Pot-a-eau, and one also in the forest near Mare aux
Cochons: in Mahé a number were taken in the forests near Morne Blanc, and specimens
were also found in the high forests behind Trois Fréres and above Cascade Estate: in
Félicité several were found in the forest.
TETRAGONORRHAMPHUS, nN. gen.
Rostrum flattened, nearly as wide as the head, broad, subquadrate, obsoletely
bituberculate at base (¢), feebly arcuate-emarginate at the apex, leaving the tips of the
mandibles exposed, the antenne inserted at the middle, the scrobes lateral, oblique,
reaching to beneath the eyes; eyes small, lateral, oval, prominent, the head somewhat
exserted and feebly constricted behind them; antenne short, the scape reaching to the
posterior margin of the eyes, the funiculus seven-jointed, joint 1 stout, 2—7 transverse,
closely articulated, widening outwards, the club small, ovate, sparsely pilose, annulate, the
basal half polished ; prothorax oblongo-conic ; scutellum flattened, subquadrate, polished ;
elytra oblong, crenato-sulcate ; coxee widely separated; prosternum with a large, oval,
sharply-defined, flattened space extending forwards to the apex ; legs short ; femora stout,
clavate; tibiee flattened, subtriangular, produced into a long curved hook at the outer
apical angle, the anterior pair acutely rectangular or with a short tooth at the inner
apical angle and armed with a short tooth on the inner edge towards the apex ; tarsi
short, jot 3 rounded-triangular, much wider than 2, not lobed, pilose at the apex, the
claw-joint slender, moderately long, the claws minute; body moderately elongate,
depressed, shining, glabrous, winged.
Type, T. tuberculirostris.
The small insect from which the above diagnosis has been drawn up is one of the
a
CHAMPION—-COLEOPTERA ; CURCULIONIDA 487
commonest Cossonids in the mountain forests of the Seychelles. The broad, flattened,
subquadrate, alutaceous rostrum, the laterally-placed, rather prominent eyes, the short,
seven-jointed funiculus, the small antennal club, the strongly sculptured elytra, the widely
separated cox, the broadly flattened prosternum, &c., are characteristic. It does not
seem to come very near any of the genera described by Wollaston. The flattened, still
broader rostrum, the small eyes and antennal club, &c., separate Tetragonorrhamphus
from Coptus, near which the genus can be placed.
120. Tetragonorrhamphus tuberculirostris, n. sp. (Pl. 24, figs. 46, 46a, 3.)
Varying in colour from piceous to ferruginous, the elytra often rufescent in dark
individuals, the antennz and tarsi always ferruginous, moderately shining. Head and
rostrum densely alutaceous, rather dull, closely, very minutely punctate, the rostrum as
broad as long, a little narrower and more shining in ?, rounded at the anterior angles, the
head triangularly depressed or flattened, or shallowly foveate, between the eyes, the
depression in the larger examples preceded by two transversely-placed minute tubercles
(fig. 46a). Prothorax a little longer than broad, oblongo-conie, rapidly narrowing forwards
and rounded at the sides posteriorly, the apical constriction feeble, lateral, leaving a short
collar-like margin on each side ; the surface with an excessively minute dense punctuation
intermixed with scattered more conspicuous punctures. Elytra moderately long, depressed,
about as wide as the prothorax, subparallel in their basal half; crenato-sulcate, the
interstices narrow, convex, rugulose. Beneath with an excessively minute punctuation
intermixed with scattered coarser impressions; the prosternal depression distinctly
margined laterally, transversely-strigose.
Length 2—3 mm. |
Loc. Seychelles: Mahé, Silhouette, Praslin, Félicité, Marie Anne.
Upwards of 100 examples of this insect were captured by Mr Scott, showin
considerable variation in size and colour. The broader, more robust individuals, with
a subopaque, minutely bituberculate rostrum, are assumed to be males. No other
Cossonid is known to me possessing the last-mentioned character.
Mahé: a number of specimens were found under bark of fallen and rotting trunks
of “Bois Rouge” (Wornuia ferruginea) in the high forest behind Trois Fréres, 1500—
2000 feet ; it was also found near Morne Blanc, in the Mare aux Cochons district, and the
forests above Cascade (including summit of Mount Sebert). Silhouette: it was found in
the forests near Mont Pot-d-eau, ca. 1500 feet, and above Mare aux Cochons (two
specimens being recorded as from fallen “Bois Rouge”). Praslin: several were taken
in the Coco-de-mer forest in the Vallée de Mai. In Félicité and Marie Anne specimens
were found in a drier type of forest near sea-level, composed of Calophyllum Inophyllum
(“Takamaka”), Terminalia catappa and other trees.
Coptus.
Coptus Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 492, 578.
Wollaston included two small Malayan forms under this genus. A single species has
now to be added from Mahé.
488 PERCY SLADEN TRUST EXPEDITION
121. Coptus latiusculus, n. sp.
Moderately elongate, somewhat fusiform, slightly depressed, shiming; nigro-piceous,
the elytra obscurely rufescent. Rostrum very stout, curved, convex, parallel-sided, not
longer than the width of the head with the eyes, densely, rather finely punctate, the
convex inter-ocular space similarly punctate; eyes large, prominent, the head constricted
behind them ; antennze with joints 2—7 of the funiculus very closely articulated, strongly
transverse, the club abrupt, large, somewhat globose. Prothorax oblong-oval, a little
longer than broad, gradually narrowed forwards to the feeble subapical constriction, which
is more deeply impressed laterally ; closely, moderately coarsely punctate. Scutellum very
small. Elytra moderately long, very little wider than the prothorax, parallel at the base ;
crenate-striate, the narrow interstices transversely rugulose and irregularly uniseriate-
punctate. Beneath closely, rather finely punctate; metasternum narrowly sulcate.
Anterior coxee separated by about the width of one of them. ‘Tibize unarmed at the
inner apical angle.
Length 25 mm. (? 2).
Loc. Seychelles: Mahé.
One specimen, probably , the first ventral segment being unimpressed. It was
found on the Cascade Estate at 800—1000 feet. Near C. oculatus, Woll., from Sula,
but broader and more depressed, the prothorax more finely punctured.
PROKCES.
Proéces Schénherr, Gen. Cure., iv. p. 1080 (1838) ; Lacordaire, Gen. Col., vii. p. 339.
This genus was based upon two small Cossonids from Madagascar, and others have
since been added from Tahiti and the Seychelles*. Two species are represented in the
collection under examination. Proéces, type P. macer, Boh., was unknown to Wollaston.
The Seychelles forms obtained by Mr Scott agree in the following characters :—
‘Rostrum moderately long, curved, cylindrical, the antennze inserted behind the middle ;
eyes rounded, large, prominent, the head sharply constricted immediately behind
them; antennse with a seven-jomted funiculus, and an abrupt, annulate, ovate club;
prothorax long, conical, basally dilated ; elytra elongate, depressed, parallel or subparallel ;
anterior and intermediate coxee widely, the posterior coxze more narrowly, separated ;
anterior cox large; metasternum flattened down the middle, narrowly sulcate; outer
apical angle of the tibize produced into a long, curved hook, the inner angle acute; third
tarsal joint simple.”
The specimens of P. silvestris before me include both sexes.
122. Proéces compressicollis, n. sp.
Elongate, depressed, shining, ferruginous, the eyes black. Rostrum cylindrical, .
slightly widened at the tip, slender, curved, as long as the prothorax, sparsely, minutely
punctate, finely striate along the sides, the antennz inserted far behind the middle; head
* P. rufipes, Fairm., as already stated, is a Stenotrupis.
CHAMPION—COLEOPTERA ; CURCULIONID A 489
feebly foveate and sparsely, finely punctate between the eyes, constricted immediately
behind them, the eyes large and prominent; antennz with a rather large abrupt club,
joint 1 of the funiculus stout, 2—7 short and closely articulated. Prothorax longer than
broad, subconical, compressed and much narrowed anteriorly, broadly arcuato-explanate
posteriorly, the apical constriction well-defined ; finely, rather sparsely punctate. Scutellum
rounded, polished. Elytra moderately long, not so wide as the prothorax, very gradually
narrowed from the base; shallowly punctato-sulcate, the interstices flat on the dise,
becoming narrower outwards, minutely uniseriate-punctate. Beneath sparsely, finely
punctate; metasternum narrowly suleate. Tibize with their inner apical angle acute,
the anterior pair sinuate and ciliate within. Tarsi with joints 2 and 3 short,
3 simple.
Length 3—3,5 mm. (¢ 2).
Loc. Seychelles: Silhouette.
Three examples from the Mare aux Cochons, September, 1908, probably obtained
from stems of an imported Dracena.
P. ngrifrons, Boh., from Madagascar*, doubtless comes near P. compressicollis.
123. Proéces silvestris.
Catolethrus silvestris Kolbe, Mittheil. Zool. Mus. Berlin, v. p. 45 (1910).
Elongate, somewhat convex, shining, obscure ferruginous, the rostrum, antenne, legs,
and under surface ferruginous, the head infuscate, the eyes black. Head and rostrum
closely, finely punctate, the head faintly foveate between the eyes and constricted
immediately behind them, the latter prominent, the rostrum curved, cylindrical, rather
stout, much shorter than the prothorax, the antenne inserted at about the basal third ;
antennze rather slender, with a large, abrupt, annulate club. Prothorax considerably
longer than broad, conical, compressed at the sides anteriorly, the latter rounded towards
the base, the subapical constriction well-defined laterally; densely, finely punctate.
Scutellum small, narrow. Elytra moderately long, a little narrower than the prothorax,
parallel in their basal half; shallowly punctato-sulcate, the interstices flat, closely,
minutely uniseriate-punctate. Beneath finely punctate; metasternum sulcate; ventral
segments 1 and 2 deeply excavate down the middle in ¢. Tibiz with their inner
apical angle acute, the anterior pair sinuate and ciliate within. Tarsi with joint 1 about
as long as 2 and 3 united, 3 simple, scarcely wider than 2.
Length 3—34+ mm. (29).
Loc. Seychelles: Mahé.
One specimen, ?, from the Cascade Estate, agreeing with a co-type (f) from Mt Harrison
kindly communicated by Professor Kolbe. Less depressed and a little larger than
P. compressicollis ; the rostrum shorter and stouter, and closely punctate; the prothorax
* The minute insect representing this species in the British Museum (labelled ex Jekel) is wrongly
named, and belongs to a different genus.
490 PERCY SLADEN TRUST EXPEDITION
less compressed anteriorly, and much more densely punctate; the elytra parallel-sided,
and with the minute interstitial punctures more numerous.
The description now given will supplement that of Kolbe. Proéces silvestris cannot
possibly be included in Calotethrus, the type of which is the common Central American
C. longulus, Boh.
STENOMIMUS.
Stenonwmus Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 480, 564.
Stilbocara Broun, Ann. Mag. Nat. Hist. (6), xii. p. 387 (1898).
The species hitherto referred to Stenomimus (type, S. fry, Woll., from Brazil) are all
American. One from the Seychelles is so closely related to S. guatemalensis and some
other Central-American forms described by myself that it is here referred to the same
genus. Stilbocara (Phleophagosoma) constricticollis, Broun, from New Zealand, is
certainly congeneric; S. nitida, Broun, I have not seen.
124. Stenomimus orientalis, n. sp.
Subfusiform, moderately elongate, somewhat depressed, shining, ferruginous, the body
sometimes piceous. Rostrum curved, cylindrical, moderately stout, a little more than half
the length of the prothorax, closely, very finely punctate, the antennze inserted at the
middle; head flattened, punctate, and shallowly foveate between the eyes, and constricted
behind them, the eyes rather prominent ; antennze with an abrupt, pubescent, rather large,
ovate club. Prothorax slightly longer than broad, rounded at the sides posteriorly,
narrowing from about the basal fourth, and constricted before the apex; finely, closely
punctate. Scutellum polished. Elytra moderately long, wider than the prothorax,
narrowing from about the middle; punctato-sulcate, the sutural stria deeply impressed,
the interstices faintly punctulate. Beneath closely, finely punctate; metasternum sulcate ;
first ventral segment transversely depressed in the middle posteriorly in ¢. Coxze small,
widely separated. Tibize with their inner apical angle acute, the anterior pair very feebly
sinuate within. Tarsi slender, joint 3 a little wider than 2, simple.
Length 1Z—2,, mm. (¢9).
Loc. Seychelles: Mahé, Silhouette.
In Silhouette a number of specimens were found in the forests near Mont Pot-a-eau,
ca. 1500 feet, and above Mare aux Cochons. In Mahé examples were found near Morne
Blanc, also in the high forest between Trois Fréres and Morne Seychellois, 1500—
2000 feet.
The thirty-three specimens before me vary a little in colour; those with a feebly
depressed first ventral segment and a rather more closely punctured rostrum are assumed
to be males. This insect is very much smaller than the species here referred to Proéces,
and has a much shorter rostrum, smaller eyes, a shorter, less dilated prothorax, smaller
coxee, &c. S. orzentalis is apparently restricted to the mountain forests of the Seychelles.
It is very like S. constricticollis, but has a smoother and less constricted prothorax, and a
shorter rostrum.
a oes
ee
— wn.
ee
CHAMPION—COLEOPTERA ; CURCULIONID A 491
STENOSCELIS.
Stenoscelis Wollaston, Journ. Ent., i. p. 141 (1861); Trans. Ent. Soc. Lond., 1873,
pp. 504, 9594.
The described species of this genus are from 8. Africa, Japan, and St Helena, and one
is now before me from Aldabra. They are doubtless carried about in logs or timber.
125. Stenoscelis crassifrons.
Stenoscelis crassifrons Woll., Trans. Ent. Soc. Lond., 1873, p. 651.
Loc. Aldabra, 1908 (Fryer). S. Africa.
A single specimen from Takamaka, Aldabra I., agrees with the unique type of
S. crassifrons from the Cape of Good Hope, except that the head and rostrum are
narrower (probably a sexual character) and less rugosely punctate, and the sides of the
prothorax are more strongly sinuate. From S. hylastoides, from St Helena and 8. Africa,
the present species may be separated by the definite, coarse, transverse, seriate punctures
of the elytra (a character not definitely mentioned by Wollaston) and the setulose apical
declivity.
DENDROCTONOMORPHUS.
Dendroctonomorphus Wollaston, Trans. Ent. Soc. Lond., 1873, pp. 502, 591.
The two known members of this genus are from Ceylon and Mexico respectively, and
a third from Malabar is mentioned (but not described) by Wollaston. The Ceylon insect,
D. muricatus, is given as the type and an allied form is now added from the Seychelles,
possibly not really endemic, as there is a specimen from Java in the British Museum that
evidently belongs to the same species.
126. Dendroctonomorphus muriceus, n. sp.
Moderately elongate, convex, cylindrical, rather broad, shining, black, the tarsi and
antennee obscure ferruginous. Head and rostrum closely, finely punctate; rostrum very
broad, transverse, widened towards the base, obsoletely canaliculate, the junction with the
head indicated by a faint transverse groove, which is sometimes shallowly foveate in the
middle; eyes large, depressed ; antennze with a large, abrupt, rounded club. Prothorax
transverse, gradually narrowing forwards, the sides rounded posteriorly and slightly
compressed or sinuate at the middle; coarsely, closely punctate. Scutellum minute,
depressed. Elytra cylindrical, moderately long, bluntly rounded at the apex; coarsely
erenato-sulcate, the interstices not wider than the sulci, asperate and confusedly punctate,
the asperities becoming more distinct on the apical half. Beneath closely punctate, the
metasternum sulcate. Tarsi long and slender, the claws elongate.
Length 34—3# mm.
Loc. Seychelles: Mahé. Java.
A long series. This species was found in dead wood: the majority of specimens are
from near Morne Blanc (almost certainly from the dead trunk of a “ Bois montagne” tree),
SECOND SERIES—ZOOLOGY, VOL. XVI. _ 63
492 PERCY SLADEN TRUST EXPEDITION
and some were also found on Cascade Estate at about 1000 feet. D. muriceus is closely
related to D. muricatus, Woll., from Ceylon, differing from it in having the prothorax less
elongate and, at most, very feebly sinuate at the sides, and the head less densely punctate.
The specimen from Java in the British Museum, separated from D. muricatus as a distinct
species, is considerably larger than Seychelles examples.
Sub-fam. Calandrine.
Some authors treat the Calandrids as a separate sub-family of the Curculionide,
others include them under the Curculionine. Of the eight species examined from the
Seychelles, one only, Hugnoristus braueri, appears to be endemic.
. EUGNORISTUS.
Eugnoristus Schénherr, Gen. Cure., iv. p. 848 (1838).
The type of this genus, Calandra monacha Oliv., is from Madagascar, and various
other species have since been described from the same country, the Comoro Is., and the
Seychelles. They are probably all palm-feeders.
127. Eugnoristus brauerr. (Pl. 24, fig. 47, 2.)
Eugnoristus brauert Kolbe, Mitteil. Zool. Mus. Berlin, v. p. 47 (1910).
Loc. Seychelles: Mahé, Silhouette.
E. braueri was described from a single specimen from Mahé. There are 70 examples
of it in the collection before me (and also others in the British Museum). The majority
of the 70 specimens are from Silhouette, where they were found in the high forests near
Mont Pot-a-eau and above Mare aux Cochons in VIII. and IX. 1908: all (or almost all)
were found in the decayed outer leaf-bases of felled heads of an endemic palm
(Verschaffeltia splendida); on several occasions larvee were also found in the leaf-bases,
e.g. on Aug. 31 larvee, pupee, and adults were all found together. The insect pupates in
a cocoon made of the fibres of the palm, similar in nature to that constructed by some
other Calandrids. Several specimens were found in Mahé on Cascade Estate, in the
Mare aux Cochons district, and in various other localities, but are without special record
of habitat: and the collection also contains one specimen found in Mahé by Mr J. J.
Lister in 1888. The variation in length is from 6—114 mm. (rostrum excluded), and the
rostrum also is sometimes greatly elongated, especially in the female. The sexes are very
similar, but the males may be identified by the deeply excavate first ventral seement and
the laterally angulated basal portion of the rostrum. The posterior tibiz are closely
fulvo-ciliate along their inner edge for more than half their length in both sexes.
TROCHORRHOPALUS.
Trochorhopalus Kirsch, Mittheil. Dresd. Mus., 1877, p. 156; Faust, Ann. Mus. Civ.
Stor. Nat. Genova, xxxiv. p. 342 (1894).
A single species has been referred to this genus, the type of which came from Java.
CHAMPION—COLEOPTERA; CURCULIONIDA 493
128. Trochorrhopalus strangulatus.
Sphenophorus strangulatus Gyll., in Schonh., Gen. Cure., iv. p. 763.
Trochorhopalus strangulatus Kirsch, Mittheil. Dresd. Mus., 1877, p. 156; Gahan,
Monogr. Christmas Isl., p. 113.
Loc. Seychelles: Mahé. Java, Siam, Malacca, Philippines, Borneo, Bouru, New
Guinea, Mauritius, Christmas Island.
Eight specimens of 7. strangulatus were obtained by Messrs Gardiner and Scott in
Mahé. One was found in 1905; one is from near Morne Blane, ca. 1000 feet, XI. 1908;
and six are from Cascade Estate, 800—1000 feet, I. 1909. The males may be known by
the broadly depressed. metasternum and first ventral segment, the long fulvous cilia on
the inner edge of the posterior tibize towards the apex, and the stouter and more velvety
rostrum.
CosMOPOLITES.
Cosmopolites Chevrolat, Ann. Soc. Ent. France, (6) v. p. 289 (1885); Faust, Ann.
Mus. Civ. Stor. Nat. Genova, xxxiv. p. 342 (1894).
The single species referred to Cosmopolites, like Metamasius sericeus and its allies in
Tropical America, is a destructive insect to Musaceous plants in the east. It appears to
be known in the Seychelles as the banana-weevil. Chevrolat also includes Brazil in its
distribution, possibly in error.
129. Cosmopolites sordidus:
Calandra sordida Germ., Ins. Spec. Nov., i. p. 299.
Sphenophorus sordidus Gyll., in Schénh., Gen. Curc., iv. p. 925; Kolbe, Mitteil.
Zool. Mus. Berlin, v. p. 47 (1910).
Cosmopolites sordidus Chevr., Ann. Soc. Ent. France, 1885, p. 290; Faust, Ann.
Mus. Civ. Stor. Nat. Genova, xxxiv. p. 340 (1894).
Sphenophorus cribricollis Walk., Ann. Mag. Nat. Hist., (3) iv. p. 218 (1859).
Loc. Seychelles: Praslin, Mahé. Réunion, Java, Burma, India, Ceylon, Andaman
Islands, Malacca, Saigon, China, Sunda, Fiji.
The majority of the Seychelles specimens are from Praslin, XI. 1908. Examples
were also found in Mahé at various places in 1905 and 1908—9. Many were found
in fallen and rotting stems of banana-trees. The long series before me from the Seychelles
vary from 7$—11 mm. in length, exclusive of the rostrum. The sexes are very similar,
but the male may be identified by the broadly hollowed first ventral segment.
PoLytus.
Polytus Faust, Ann. Mus. Civ. Stor. Nat. Genova, xxxiv. p. 353 (1894)*.
A monotypic genus, the type of which is Sctophilus mellerborgii, Boh. (= Calandra
remota, Sharp), from Java, the Hawaiian Is., &c. The insect has doubtless been intro-
duced into the Seychelles, as well as into the Hawaiian Islands.
* Erratum.—tIn the second line of Faust’s diagnosis for “septimi” read “sexti.”
63—2
AQA PERCY SLADEN TRUST EXPEDITION
130. Polytus mellerborgi.
Sitophilus mellerborgw Boh., in Schénh., Gen. Cure., iv. p. 976.
Calandra mellenborgi (sic) Gemm. et Harold, Cat. Col., vill. p. 2653.
Polytus mellenborgi Faust, Ann. Mus. Civ. Stor. Nat. Genova, xxxiv. p. 353 (1894).
Calandra remota Sharp, Scient. Trans. Roy. Dubl. Soc., (2) mi. pp. 183, 254 (1885) ;
Perkins, Fauna Hawaiiensis, 1. p. 139.
Loc. Seychelles: Mahé, Silhouette, Praslin. Réunion, Burma (Rangoon), Java, Nias,
Hawaiian Islands (near Honolulu).
Twenty-six specimens from the Seychelles: the majority were found near Morne
Blanc in Mahé in’ XI. 1908, and one had previously been captured in Mahé in 1905.
Several were found near Mont Pot-4-eau in Silhouette in VIII. 1908, and one in Praslin,
XI. 1908. The male has the metasternum more broadly and deeply excavate posteriorly,
and the apical portion of the rostrum less elongate and more closely punctured, than the
female. We are indebted to Dr Heller for comparing a Seychelles example with others
from which Faust’s description of the genus was taken. Boheman describes the prothorax
as “obsoletissime carinato”: in the examples before me there is usually a narrow smooth
space down the middle, but there is no trace of a carina. P. mellerborg: has been found
commonly in the stems of banana and prickly pear near Honolulu.
CALANDRA.
Calandra [Clairville], Ent. Helv., i. p. 62 (1798).
Sitophilus Schénherr, Gen. Cure., iv. p. 967 (1837).
This genus, as now restricted, includes several widely distributed well known forms
which are carried everywhere in commercial products, and various typical species, such as
C. exarata and C. stegmaticollis in the east, and C. setulosa in the west, the last-
mentioned being less widely dispersed at present.
131. Calandra oryze.
Curculio oryze Linn., Amcen. Acad., vi. p. 395 (1763).
Cossonus quadrimacula Walk., Ann. Mag. Nat. Hist., (3) iv. p. 219 (1859).
It is not necessary to quote the synonymy of this insect, except to add that of
Walker, whose type was from Ceylon.
Loc. Seychelles: Silhouette, Mahé, 1908—9. Coetivy, 1905. Providence; Cerf
Island, 1905. Aldabra: Takamaka, 1908 (Fryer). Cosmopolitan.
In Mahé this species was found on Cascade Estate and in the Mare aux Cochons
district. A number of specimens collected by the 1905 Expedition are labelled
‘H.M.S. Sealark” and were presumably found on the ship.
132. Calandra linearis.
Rhynchophorus linearis Herbst, Kifer, vil. p. 5, pl. 100, fig. 1 (1797).
Cordyle striatus Thunb., in Nova Acta Ups., vil. p. 112 (1815).
Calandra linearis, var. striata Perkins, Fauna Hawaiiensis, i. p. 139.
CHAMPION—COLEOPTERA ; CURCULIONIDA 495
Calandra tamarind Christy, Trans. Ent. Soe. Lond., i. p. 36 (1834).
Loc. Seychelles : Long Island. Comoro Islands, Hawaiian Islands, North and South
America, Antilles, &e.
One specimen was found in the cultivated islet, Long Island, off Mahé, in July, 1908.
133. Calandra exarata.
Stitophilus exaratus Boh., in Schénh., Gen. Cure., iv. p. 970.
Sphenophorus exquisitus Walk., Ann. Mag. Nat. Hist., (3) iv. p. 218 (1859).
Calandra porcata Pasc., Ann. Mus. Civ. Stor. Nat. Genova, xxii. p. 306 (1885).
Loc. Seychelles: Mahé: Réunion, Ceylon, Singapore, Sumatra, Aru, &e.
Five specimens of this species were captured by Mr J. S. Gardiner in Mahé in 1905.
It is readily separable from C. stigmaticollis by its straight rostrum in both sexes, the
rougher, opaque surface, the coarser setosity, the relatively shorter elytra, and the
non-geminate arrangement of the strize, the elytral interstices thus being subequal in
width. The ochreous elytral markings are sometimes coalescent, forming a common
X-shaped patch. The female has the apical portion of the rostrum comparatively slender,
smoother, and more shining than in the male, that of the male being opaque and coarsely
punctate to the tip. Boheman’s type was from Bourbon. There is an example of this
species from Singapore in the British Museum labelled “found destroying sugar cane.”
134. Calandra stigmaticollis.
Sitophilus stugmaticollis Gyll., in Schénh., Gen. Cure., iv. p. 972.
Calandra stigmaticollis Kolbe, Mitteil. Zool. Mus. Berlin, v. p. 46 (1910).
Sitophilus subfasciatus Boh., in Schénh., Gen. Curc., iv. p. 971.
Sitophilus subsignatus Gyll., loc. cit., p. 973.
Calandra punctigera Pasc., Ann. Mus. Civ. Stor. Nat. Genova, xxii. p. 305 (1885).
Calandra sechellarum Kolbe, Mitteil. Zool. Mus. Berlin, v. p. 46 (1910).
Sphenophorus cruciger Motsch., in litt.
Loc. Seychelles: Silhouette, Mahé. Madagascar, Mauritius, India, Ceylon, Java,
Sumatra, New Guinea, Celebes, Ceram, Borneo, Batchian, Aru, Flores, Tonga Tabou,
Wallis, &c.
Seven specimens were found in the Seychelles, three near Mont Pot-d-eau in
Silhouette (VIII. 1908), three from various localities in Mahé (1908—9), and one from
a cultivated islet, Anonyme, off Mahé (I. 1909): they include both forms of the insect.
A very variable species that has been described under many different names. C. sechel-
larum, from Mahé ( =subsignatus, Gyll.), was based upon a single large female example,
reddish-brown in colour, with some dark streaks on the prothorax and elytra, and a rather
elongate second joint to the funiculus, characters that are of no value when a series is
examined. The name stigmaticollis seems preferable to adopt, rather than that of
subfasciatus. C. taitensis, Guér., from Tahiti, Fiji, &c., is a closely allied form, varying in
colour in a similar way; it may be separated from C. stigmaticollis by the more equal
width of the elytral interstices and the strongly carinate pygidium.
496
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
fo)
gw © Sy ge SY fe fo bo iS
ee
Dw Fw wy ES
Wy.
13.
. 19.
. 20.
ig. 21.
ig. 22.
ig. 23.
. 24,
. 25.
. 26.
. OF.
. 28.
2)
. 80.
yaoi
ig. 32.
PERCY SLADEN TRUST EXPEDITION
EXPLANATION OF PLATES 22—24.
PLATE 22.
Euops viriditinctus, n. sp., f ; a, profile of head.
Lepydnus nepenthicola, n. gen. et n. sp.; a, profile of head.
Brachycyrtus minor, n. sp.; a, profile of head.
Epirrhamphus litoralis, n. gen. et n. sp.; a, profile of head.
Cratopus adspersus, C. O. Waterhouse, ?.
Cratopus griseovestitus, Linell, #.
Cratopus segregatus, n. sp., var. subcinctus, n. var., i.
Cratopus muticus, n. sp. ¢.
Cycloterodes sechellarum, Kolbe, f ; a, profile.
Eucycloteres terreus, n. gen, et n. sp.; a, profile.
Gonotrachelus 5-carinatus, n. gen. et n. sp.; a, profile; 6, antenna.
Tanyomus palmicola, n. gen. et n. sp., §; a, profile of head and prothorax.
Cycloterinus humeroalatus, Kolbe, §; a, profile of head.
Cycloterinus anvpliatus, Kolbe, £ ; a, profile of head.
Cycloterinus foveatus, Kolbe, §; a, profile of head; 6, second ventral segment.
Cycloterinus carinifer, n. sp., 2; a, profile of head.
PLATE 23.
Cycloterinus canaliculatus, n. sp., f ; 4, profile of head.
Cycloterinus unicristatus, n. sp., f ; a, profile.
Cycloterinus erosus, n. sp., 2; a, profile of head.
Phenicobates vittatus, n. gen. et n. sp. ¢; a, profile of head; 6, anterior tarsus.
Phenicobates flexirostris, n. sp., §; a, profile of head.
Phenicobates flexirostris, nu. sp., 2; profile of head.
Pheenicobates gibbirostris, n. sp., f; 4, profile of head.
Phenicobates stricticollis, n. sp., f ; a, profile of head.
Pheenicobates hispidulus, n. sp., f; a, profile of head.
Pheenicobates cuspidatus, n. sp. ¢.
Phenicobates pandanicola, n. sp., J.
Phenicobatopsis echinatus, n. gen. et n. sp.; a, profile.
Phenicogenus inermis, n. gen. et n. sp.; a, profile of head.
Endcopsis curvimanus, n. gen. et n. sp., F.
Rhynchenus spissus, n. sp., 2 ; a, profile.
Acalles seychellensis, n. sp., J; a, profile.
Percy SLADEN TRUST EXPEDITION. ~ TRANS. LINN. Soc. SER.2. Zoot VoL.XVLPL. 22.
(CHAMPION)
E Wilson del
Cambridge University Press
COLEOPTERA, CURCULICNIDA.
Percy SLADEN TRUST EXPEDITION. TRANS. Linn. Soc. SER.2.Z00L VOL.XVIPL. 23,
(CHAMPION)
E.Wilson del. Cambridge University Press.
COLEOPTERA, CURCULIONIDA.
Percy SLADEN TRUST EXPEDITION. TRANS. LINN. Soc. SER.2.Z00L. VOL.XVIPL. 24.
(CHAMPION)
E.Wilson del Cambridge University Press
COLEOPTERA, CURCULIONIDA.
Fig.
Fig. +
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig
5
Fig.
Fig
g.
Fig.
Fig.
Fig.
CHAMPION—COLEOPTERA ; CURCULIONIDA A497
PLATE 24,
Cryptorrhynchus erratus, n. sp.. 2 ; a, profile.
Cryptorrhynchidius graniger, n. gen. et n. sp., 2 ; a, profile.
Rhetogenes sexcristatus, n. gen. et n. sp., f ; a, profile.
Baridiomorpha triplaris, n. gen. et n. sp., #3; a, profile of head.
Microplatymerus lodoicewvorus, n. gen. et n. sp.; a, profile of head; 6, antenna; c¢, anterior leg.
Homalorrhynchus serripes, n. gen. et un. sp., of.
Homalorrhynchus verschaffeltiw, n. sp., § ; a, profile of head.
Sphodrias magdaloides, n. gen. et n. sp., ¢.
Sycites pilicormis, n. gen. et n. sp. oi.
Pentarthrum longicolle, n. sp. &.
Hypopentarthrum microcephalum, n. gen. et n. sp., .
Trapezirrhynchus silhouettensis, n. gen. et n. sp., f.
Microhimatium pubescens, n. gen. et n. sp., f; a, antenna.
Tetragonorrhamphus tuberculirostris, nu. gen. et n. sp., #; a, head and rostrum from above.
Hugnoristus brauert, Kolbe, ¥.
‘
'
HN DX
[Norz.—Synonyms and Native Names are printed im italics, A star is added to names which
appear to be used for the first time.]
Acalles, Schinh., 440; mentioned,
393, 415.
seychellensis, Champ.*, 441 ;
mentioned, 394, 496.
Acanthaclisis, Ramb., mentioned,
243.
maritimus, Weedham*, 244.
Acanthaspidine, 167.
Acanthephyra, 4. MW. Hdw., 64;
mentioned, 65, 67.
purpurea, A. UM. Hdw., 64;
mentioned, 53, 65, 67.
var. multispina, Coutiere,
mentioned, 65.
sanguinea, Alcock, mentioned,
53, 65.
Acanthobrachys elegantula (Sta),
170.
Acanthomia horrida (Germ.), 146.
Acanthophora orientalis, J. Ag.,
287; mentioned, 271, 273.
spicifera (Vahl), Borges., men-
tioned, 287.
Thierii, ZLamour., mentioned,
287.
Acanthymenia, Ag., mentioned,
301.
Acarodes, Woll., mentioned, 259,
260.
Acaromimus, Jord., mentioned,
260.
Achoragus, Jord.*, 265.
tener, Jord.*, 265 ; mentioned,
267.
Achynius, Fairm., mentioned,
406.
Acidia, Rob.-Desv., 316.
cesio (Harris), mentioned,
319.
Acidia cognita, Wied., mentioned,
317.
ocellata, Lamb*, 317; men-
tioned, 318, 371.
seychellensis, Zamb*, 316;
mentioned, 371.
Acritus, Leconte, 229.
alluaudi, Schmidt, mentioned,
230.
madagascariensis,
mentioned, 230.
(ABletes) daubani, Scott*, 230;
mentioned, 223, 228, 235.
) davidsoni, Scott*, 231;
mentioned, 223, 235.
Scott*, 232;
?
Schmidt,
(
(——)_ fryeri,
mentioned, 223, 235.
) longipes, Sharp, men-
tioned, 230.
)
Scott, mentioned, 230.
) subalatus, Scott, men-
tioned, 230.
) subrotundus,
tioned, 229.
(Halacritus) algarum, Schmidt,
229; mentioned, 223, 224,
228, 230, 235.
Acrosternum heegeri, Fieb., 144.
Acrosticta, Loew, 314; mentioned,
312, 370.
exsul, Zamb*, 314; mentioned,
371.
pallipes, Grimshaw, mentioned,
312.
scrobiculata, Zoew, mentioned,
314.
Adenanthera pavonina, Linn., men-
tioned, 378.
(
(
(
(
var. haleakale,
men-
SECOND SERIES—ZOOLOGY, VOL. XVI.
Afletes, Horn, 230; mentioned,
223, 224, 229.
Hthus pallidipennis, Bergr., 140.
Agrilus owas, Cast. et Gory, 378.
Alaocyba carinulata, Woll., men-
tioned, 479.
Algx, Marine, Rhodophycee, by
A. Weber-van Bosse, 269-306.
List of Localities and Names
of, found at them, 271—272.
—— Systematic Survey of all
the, collected by Mr J. Stanley
Gardiner in the Indian Ocean,
272-274.
Allometopon, Kert., 310; men-
tioned, 370.
flavum, Zamb*, 311; men-
tioned, 371.
Alphilia, /e7tt., mentioned, 128.
Alydine, 147.
Alydus proletarius, Schaum., 147.
Amirantea gardineri, Distant, 144.
Amorphorhynchus, Woll., men-
tioned, 482, 483.
Amphiroa, Lama., 305.
foliacea, Lama, 305; men-
tioned, 271, 274.
fragilissima (Linn.), Lamex.,
305; mentioned, 271, 274.
Amphisbetema, Web. v. B.*, 297;
mentioned, 270, 299.
indica (J. Ag.), 297; men-
tioned, 272, 274, 298, 300,
306.
Amphoriscide, 24.
Amygdalops, Lamb*, 357; men-
tioned, 351.
thomasseti, Lamb*, 358; men-
tioned, 357, 372.
64
500
Anas undulata, Dubois, Pseudo-
scorpion found on, mentioned,
32.
Anatricus arenaceus, Loew, men-
tioned, 370.
Anchonina, Champ., 407.
Aneurus bicolor, Distant*, 160;
mentioned, 190.
brevisculatus, Sergr., men-
tioned, 160.
greeni, Distant, mentioned,
161.
Anisops aldabrana, Distant*, 189;
mentioned, 191.
varius, [%eb., 189.
Anocerine, mentioned, 247.
Anoplocnemis curvipes
146.
Anthicus, Payk., mentioned, 259.
Anthocoride, 184.
Anthocoris parallelus, Motsch., 188.
Anthonomina, 437.
Anthonomus, Germ., mentioned,
439.
Anthribide, 247-267.
Anthribus (Arecerus), Schénh.,
267.
cervinus, Klug, 255.
fasciculatus, De Geer, men-
tioned, 267.
nigroungulatus, Fahrs., 255.
Antiopa, Sta/, mentioned, 167.
Antiopula, Bergr., mentioned,
167.
Antithamnion, WVaegeli, 300.
adnatum, J. Ag., 300; men-
tioned, 271, 274.
Antliarrhina, 451.
Aphanine, 152.
Aphaniosoma, Beck., 356.
approximatum, eck., men-
tioned, 356.
quadrinotatum, eck., men-
tioned, 356.
sexvittata, Lamb*, 356; men-
tioned, 357, 372.
Aphanus consocialis, Distant*,
154; mentioned, 189.
lineosus, Distant, mentioned,
154.
Apheloplectus, Raffr.*, 126; men-
tioned, 118, 120.
longicollis, Raffr.*, 127; men-
tioned, 118, 120, 188.
Reitt., mentioned,
Aphilia, 118,
128.
(Fabr.),
INDEX
Apoplectus, afr, 137; men-
tioned, 120.
obesus, Raffr.*, 137; men-
tioned, 120.
Arachnida, Second report on the,
—the Scorpions, Pedipalpi, and
supplementary notes on the
Opiliones and Pseudoscorpions,
by 8. Hirst, 31-37.
Aradide, 159.
Arecerus, Schinh., 2677.
fasciculatus (De Geer), 267.
Areocerus, Schénh., 267.
Arbela elegantula, Stal, 170.
Archisometrus braweri, Kraep., 32.
Arecophaga, Broun, mentioned,
452.
Aretas, Distant, mentioned, 174,
176.
sanguinarius, Distant*, 175;
mentioned, 191.
Arthrotomus, Klug, mentioned,
456.
depressus, Klug, mentioned,
456.
Artocarpus integrifolia, Linn.,
mentioned, 98, 219.
Asopine, 144.
Asteia, Meigen, 353.
nigra, Lamb*, 353.
Atenius lodoicex, Scott, mentioned,
382.
Attelabine, 395.
Avrainville amadelpha, A. et H. S.
Gepp, mentioned, 270.
Bacanius, Leconte, 228.
ambiguus, Schmidt, 228; men-
tioned, 223, 224, 228, 231.
atomarius, Sharp, 229; men-
tioned, 223.
inopinatus, Schmidt, 229 ; men-
tioned, 223, 224.
Bambusa, Schreb., mentioned, 471.
Baridiomorpha, Champ.*, 446.
triplaris, Ohamp.*, 446; men-
tioned, 394, 497.
Baridius, Schonh., 449.
Barina, 449.
Baris, Germ., 449.
multivaga, Champ.*,
mentioned, 395, 450.
seychellensis, Champ.*, 450;
mentioned, 395.
spoliata, Boh., mentioned, 450.
Basitropis, Jek, mentioned, 257.
449 ;
d
Bathycelia prelongirostris, Bergr.,
144,
Batraxis, feité., 131; mentioned,
119, 120, 121.
egregia, Faffr.*, 131; men-
tioned, 119, 120, 132, 138.
hirtella, Raffr., mentioned,
132.
indica, Raffr., mentioned, 132.
insularis, Raffr.*, 132; men-
tioned, 119, 120.
Batrisini, 130; mentioned, 118.
Batrisodes, Reztt., 130 ; mentioned,
118, 120.
caudatus, Raffr.*, 130; men-
tioned, 118, 120, 138.
Belionota prasina, Thwnb., 378.
Berosus, Leach, 198.
acutispina, Mairmaire, 198;
mentioned, 194, 196, 199.
bergrothi, Régimbari, 200;
mentioned, 194.
cuspidatus, Hrichson, men-
tioned, 198.
gracilispina, Régimbaré, men-
tioned, 198, 199.
placidus, Stal, 155.
prolongatus, Fairmaire, 199;
mentioned, 194.
villosulus, égimbart, men-
tioned, 200.
vitticollis, oh., mentioned,
200.
(Enoplurus) acutispina, Fair-
maire, 198.
( ) indicus, Motsch., men-
tioned, 198.
(——) prolongatus, Fairmaire,
199.
merle”
“ Bows (Schmidelia race-
mosa, Linn.), mentioned, 144.
“ Bows montagne” (Campnosperma
seychellarum, MJarch.), men-
tioned, 480.
“ Bows Rouge” (Wormia ferruginea,
Baill.), mentioned, 386, 408, 413,
481, 487.
Bostrichus
387.
Bourdonnaisia, Scott*, 213; men-
tioned, 195.
mahensis, Scott*, 215; men-
tioned, 194, 214, 216, 235.
silhouette, Scuté*, 216; men-
tioned, 194, 214, 235.
Brachycerus, Yabr., mentioned, 414.
Bohem.,
Jerrugineus,
Brachycyrtus, Fairm., 397.
minor, Champ.*, 397; men-
tioned, 394, 496.
setofasciatus, Mairm., men-
tioned, 397, 398.
Brachygaster denticulata, Reégim-
bart (nec Muls.), 200; men-
tioned, 201.
stagnicola, Mulsant, 200; men-
tioned, 201.
Brachygluta, Thoms., mentioned,
121.
Brachyglutini, 131; mentioned,
119, 121, 133.
Brachypeplus, Hrichson, 97 ; men-
tioned, 93.
equalis (Walker), 97; men-
tioned, 94, 98, 106, 211.
notatus, Murray, 97; men-
tioned, 94.
omalinus, Murray, 98.
patruelis, Alluaud (nec Mur-
ray), 97.
Brachyrhynchus caffer, Stal, 160.
Brentus formicarius, Fabr., 450.
Brontispa, Sharp, mentioned, 237,
239.
Bryaxis, Klug, mentioned, 121.
Budzus (Rhyparochromus) cireum-
cinctus, Walk., 157.
Buprestidae, 377.
Bythinimorpha, Schauf:, men-
tioned, 134.
Calandra [Olairv.], 494.
exarata, Boh. 495; men-
tioned, 395, 494.
linearis, Herbst., 494; men-
tioned, 395.
var. striata, Perk.,
494.
mellenborgt (sic), Gemm. et
Harold, 494.
monacha, Oliv., mentioned,
492.
oryze, Linn., 494 ; mentioned,
395.
porcata, Pasc., 495.
punctigera, Pasc., 495.
remota, Sharp, 494.
sechellarum, Kolbe, 495.
setulosa, Gylh., mentioned,
494,
sordida, Germ., 493.
stigmaticollis, Gyll., 495; men-
tioned, 395, 494.
INDEX
Calandra taitensis, Guér., mentioned,
495.
tamarindi, Christy, 495.
Calandrine, 492.
Calcareous Sponges of the Indian
Ocean, by A. Dendy, 1-29.
Calliblepharis, A7itz., 281.
prolifera (Harv.), J. Ag., 281;
mentioned, 270, 271, 273,
306.
Callicratides, Distant, 174.
rama (Kirby), 174.
Calophyllum Inophyllum, Zinn.
(“Takamaka”), mentioned, 487.
Calphurnia aberrans, Distant, men-
tioned, 164.
reticulata, Distant, 164.
Calphurnioides, Distant*, 164.
elongatus, Distant*, 164,
mentioned, 190.
Campnosperma seychellarum,
March., mentioned, 480.
Camponotus foraminosus — subsp.
grandidier, Morel, mentioned,
129:
Camptobrochis, /%eb., mentioned,
181.
Camptorrhinus, Schénh., 441; men-
tioned, 440.
brullei, Boh., 441; mentioned,
394.
dorsiger, Fairm., 441.
| Cantheconidea migratoria, Dis-
tant*, 144.
Cantinona, Distant*, 158.
precellens, Distant*, 159;
mentioned, 190.
Cape Verde Islands. Polycheta-
Serpulide from, by H. L. M.
Pixell, 69.
Capsidee, 174.
Capsus rama, Kirby, 174.
“Capucin” tree (Northea seychel-
larum, Hook.), mentioned, 215,
404, 407.
Caranistes,
247.
Carbula litigatrix, Kirk, 142.
marginella (Zhwnb.), 142.
Carcinops, Varseul, 225.
pumilio (Erichson), 225.
quattuordecimstriata (Stevens),
225 ; mentioned, 222,
223.
Caridea, 63.
Carpophilini, 97.
Erichs., mentioned,
501
Carpophilus, Steph., 101; men-
tioned, 93, 94.
angustatus, A. Murray, 105;
mentioned, 93, 94.
biguttatus, A. Murray, 105.
contingens, Olliff, 105.
dimidiatus (/abr.), mentioned,
93, 94.
var. contingens ( Walker),
105; mentioned, 94.
—— var. ochropterus, Klug,
105 ; mentioned, 94.
fovercollis, A. Murray, 102.
fusciceps, Growv.*, 102; men-
tioned, 94.
hemipterus (Linn.), 102; men-
tioned, 93, 94.
humeralis (/abr.), 101; men-
tioned, 93, 94, 102.
var. foveicollis (A. Mur-
ray), 102; mentioned, 94,
106.
ochropterus, Klug, 105.
scotti, Grouv.*, 103; men-
tioned, 94, 104.
Catolethrus, Schénh., mentioned,
490.
longulus, Boh., mentioned, 490.
silvestris, Kolbe, 489.
Ceramiacer, 299; mentioned, 274.
Ceramium (oth), Lyngbye, 300.
sp., 301; mentioned, 272, 274.
cinnabarinum (Gratel.), Hauck.,
300 ; mentioned, 271, 274.
Ceratitis, McLeay, 316.
capitata, Wied., 316.
Ceratocombide, 170.
“Gen.?” alboclavatus, Dis-
tant*, 171; mentioned, 190.
Ceratocombus insularis, Reut., 171;
mentioned, 172, 173.
Cereyon, Leach, 217.
sp., 219; mentioned, 194, 221.
crenatostriatus, Régumbart,
mentioned, 220.
crenulatus, égimbart, men-
tioned, 220.
fructicola, Scott*, 217;
tioned, 98, 194, 195,
218; 219:
laticollis,
mentioned,
219.
var.?, 219.
nigerrimus, égimbart,
tioned, 218.
men;
211,
Régimbart, 218
194, 195, 217,
men-
64—2
502
Cercyon uniformis, Sharp,
mentioned, 98, 194,
211, 217, 218.
Cerennea, Raffr.*, 125; mentioned,
118, 120.
219;
195,
brevipennis*, Raffr., 125;
mentioned, 118, 120, 138.
Cheetangiacew, 276; mentioned,
272.
Champia, Desv., 283.
sp., 283; mentioned, 271, 273.
compressa, Harv., 283; men-
tioned, 271, 273, 284.
parvula, dg., mentioned, 284,
Champion, G. C., Coleoptera,
Curculionide, 393-497.
Chantransia (DC.), Schmitz, 275.
barbadensis, Kickers,
tioned, 275.
efflorescens (J. Ag.), mentioned,
275.
Liagore, Web. v. b.%,
mentioned, 271, 272.
Nemalionis, Ardiss. et Straff.,
mentioned, 275.
Charinus seychellarum, Araep., 34.
Chelifer garypoides, HII.,
tioned, 32.
(Chernes ?) sp., mentioned, 32.
Chernes? sp., mentioned, 32.
Chilocoris, Mayr., mentioned, 140,
assmutht, Breddin, mentioned,
141.
madagascariensis, Berg., men-
tioned, 141.
solenopsides, ‘Breddin,
tioned, 141.
Chilocoristoides, Distant*, 140.
felicitatis, Distant*, 141;
mentioned, 142, 189.
mahensis, Distant*, 141; men-
tioned, 189.
Chiromachus ochropus (C. L. Koch),
32; mentioned, 31, 33.
Chiromyia, #ob.-Des., 353; men-
tioned, 356.
dubia, Zamb*, 353; mentioned,
354, 372.
hirtiscutellata,
men-
2D);
men-
men-
Lamb*,
mentioned, 372.
minima, Beck., mentioned, 354.
355;
sexspinosa, Zamb*, 355; men-
tioned, 372.
Chloropide, &c.,
Corrigenda, 370.
Cherorrhinodes, Champ.*, 458
Addenda and
‘Chrysomelide :
INDEX
Cheerorrhinodes tenuiculus, Champ.*,
458; mentioned, 395.
Chondria, C. Ag., 289.
pumila, Vick., 289; mentioned,
270, 271, 273, 306.
simpliciuscula, Web. v. B.*,
289; mentioned, 272, 273,
306.
Chondriex, 287.
Choragus, Kirby, 262; mentioned,
256, 259, 260.
bolus, Jord.*, 263 ; mentioned,
265, 266.
ornatus, Jord.™,
tioned, 267.
Chrysobothris dorsata, Fabr., 378.
yas:
262 ;
5 men-
Hispine,
Maulik, 237-242.
Chrysomyza, Mallen, 315.
zenea, Mabr., 315.
azurea, Hendel, 315.
demandata, Macq., mentioned,
315.
Chthonius tetrachelatus, Preyssler,
mentioned, 32.
Chylocladia, Grev., 284.
perpusilla, Web. v. B.*, 284;
mentioned, 272, 273, 306.
Chymomyza, Czerny, 348; men-
tioned, 325, 333.
bicolor, ZLamb*, 348; men-
tioned, 349, 371.
Cilleus, Castelnau, 98 ; mentioned,
93.
confusus, Growv.*, 100; men-
tioned, 94, 98.
megacephalus, Castelnau, 101 ;
mentioned, 93, 94, 98.
micros, Growv.*, 99;
tioned, 94, 98.
opaculus, Growv.*, 98; men-
tioned, 94, 99.
scapularis, Fairm., 101.
men-
simoni, Growv., mentioned, 100.
subplagiatus, Growv.*, 100;
mentioned, 94, 98.
Cimex curvipes, Fabr., 146.
forsteri, Fabr., 157.
marginellus, Thunb., 142.
membranaceus, Fabr., 146.
rotundatus, Sign., 173.
rubrofasciatus, De Geer, 167.
owridula, Linn., 143.
Cimicide, 173.
Cinnamomum
zeylanicum, ees,
mentioned, 178.
Cladhymenia, Hook. et Harv., 288.
oblongifolia, Hook. et Harv.,
288; mentioned, 270, 272,
273, 306.
var., mentioned, 288.
Clathrina, Gray, mentioned, 4.
coriacea, Dendy, mentioned,
5, 6.
latitubulata, Carter, 17; men-
tioned, 18.
tripodifera, Carter, mentioned,
6.
Clavigeride, mentioned, 121.
Clerada, Stgn., 152.
apicicornis, Sign., 152.
Cleranthribus, Jord.*, 256.
anthicopsis, Jord.*, 259; men-
tioned, 267.
colydiopsis, Jord.*, 258; men-
tioned, 256, 257, 259, 267.
Cletus clavatus (Sign.), 147.
Cligenes gardineri, Distant*, 153,
mentioned, 189.
Cneorrhinus, Schénh., mentioned,
398.
Coccotrypes, Hichh., 389.
sp., 391.
bicolor, Sampson*, 390; men-
tioned, 389, 391.
declivis, Sampson*, 390; men-
tioned, 389.
graniceps, Hichh., mentioned,
390, 391.
parvus, Sampson*, 391; men-
tioned, 389.
pygmeeus, Hichh., 389.
“¢ Coco-de-mer” (Lodoicea seychel-
larum, Zabdill.), mentioned, 382,
412, 452, 479.
Codium, Berth., mentioned, 296.
Ceelenomenodera, Slanch., men-
tioned, 237, 241.
Celarthrum, Bérges., 284.
Albertisii (Picc.), Borges., 284 ;
mentioned, 271, 273.
Ceelochondria, Falk,, mentioned, 289.
Celostoma, Brullé, 212; mentioned,
215.
punctulatum (A7ug), 213; men-
tioned, 194, 195, 212.
rufitarse (Boheman), 212; men-
tioned, 194, 196, 213.
simplex, Sharp, mentioned,
213.
Coleoptera: Anthribide, by K.
Jordan, 247-267.
Coleoptera: Buprestide, by C.
Kerremans, 377-378.
Chrysomelide : Hispinz, by 8.
Maulik, 237-242.
Cureulionide, by G C.
Champion, 393-497.
Hydrophilide, Histeride, by
H. Scott, 193-235.
Nitidulide, Heteroceride, by
A. Grouvelle, 93-116.
Platypodide and Ipide, by W.
Sampson, 379-391,
Pselaphide, by A. Raffray,
117-138.
Collaria improvisa, eut., 175.
Conorhinus rubrofasciatus (De
Geer), 167.
Contexta, Jord., 256.
murina, Jord., 256; mentioned,
247, 267.
Copelatus gardineri, Scott, men-
tioned, 196.
Coptus, Woll., 487; mentioned,
473.
latiusculus, Champ.*, 488;
mentioned, 395.
oculatus, Woll., mentioned, 488.
Corallinacex, 305; mentioned, 274.
Corallopsis, Girev., 281.
Cacalia, J. Ag., 281;
tioned, 271, 273.
minor (Sond.), mentioned, 281.
Cordyle striatus, Thunb., 494.
Coreide, 146.
Coreine, 146.
Coreus tenwicornis, Westw., 147.
Corizine, 148.
Corizus scotti, Distant*, 148; men-
tioned, 190.
Corynecia, Jord.*, 264.
scotti, Jord.*, 264 ; mentioned,
267.
Cosmetidz, mentioned, 35.
Cosmopolites, Chevr., 493.
sordidus, Chevr., 493;
tioned, 395.
Cossonides, 480.
Cossonina, 457.
Cossonus [Clairv.], 484.
incivilis, Fahr., 485;
tioned, 395.
quadrimacula, Walk., 494.
suturalis, Boh., 485;
tioned, 395.
variipennis, Gahan, mentioned,
485,
men-
men-
men-
men-
INDEX
Cratopus, Schénh., 399 ; mentioned,
393.
abbotti, Linell, 402.
adspersus, C. O. Waterh., 400 ;
mentioned, 394, 399, 401,
496.
aurostriatus, Yairm., 402; men-
tioned, 394, 400, 403.
var., 402.
griseovestitus, Lnell, 401;
mentioned, 394, 400, 496.
Boh.,
humeralis,
399.
muticus, Champ.*, 404; men-
tioned, 394, 399, 400, 496.
parcesquamosus, Fairm., 402.
segregatus, Ohamp.*, 402;
mentioned, 394, 400, 403.
var. subcinctus, Champ.*,
403; mentioned, 394, 400,
mentioned,
496.
sparsutus, Fairm., 401.
viridisparsus, Jairm., 401;
mentioned, 394, 400.
Creagris pervigil (Walker), 246.
Creontiades, Distant,
175.
Cruoriella, Crowan, mentioned, 302.
Cruoriopsis, Du/., 304.
cruciata, Duf., 304 ; mentioned,
272, 274.
Crustacea Decapoda, Pelagic, by S.
Kemp, 53-68.
Cryphalomorphus, Schauf., 385.
communis, Schauwf., mentioned,
386.
confragosus, Sampson*, 386.
crenatus, Sampson*, 385.
Cryphalus, Hr., 382; mentioned,
379.
abietis, Ratz., mentioned, 384.
mentioned,
fuliginosus, Llandf., mentioned,
383.
pallidus, Hichh., 382.
piceze, Matz., mentioned, 384.
tilie, Panz., mentioned, 383.
trypanus, Sampson*, 383;
mentioned, 384.
vestitus, Blandf., mentioned,
383.
Cryptonemia, J. Ag., 302.
sp., 302; mentioned, 272, 274.
seminervis, J. Ag., 302; men-
tioned, 272, 274.
Cryptopleurum, J/wls., mentioned,
222.
503
Cryptorrhynchidus, Champ.*, 443.
graniger, Champ.*, 443; men-
tioned, 394, 449, 497.
Cryptorrhynchina, 440.
Cryptorrhynchus, J/liger, 442; men-
tioned, 443.
angustior, /airm., mentioned,
449,
australis,
442,
erratus*, Champ., 442; men-
tioned, 394, 443, 497.
Gyll.,
Boisd., mentioned,
humeralis,
449,
imepfectus, Walk., 442.
lapathi, Zinn., mentioned, 443,
445.
mangiferee (fabr.), 442; men-
tioned, 394.
transversus,
tioned, 442.
Ctenistini, mentioned, 121.
Curculigo seychellensis, £oj., men-
tioned, 416.
Curculio corticalis, Payk., 457.
Jasciculatus, De Geer, 267.
lymexylon, Fabr., 457.
mangifere, EFabr., 442.
oryze, Linn., 494.
Curculionide, 393-497.
Curculionine, 404.
Cybocephalini, 110.
Cybocephalus, Hrichson, 110; men-
tioned, 93, 95.
sp. n.% 116.
brevis, Grouww.*,
tioned, 94, 111.
mentioned,
Fairm., men-
114; men-
minimus, G'rowv.*, 113; men-
tioned, 94, 111, 114.
minutus, Growv.*, 112; men-
tioned, 94, 111.
subpallipes, Growv.*, 115;
mentioned, 94, 111.
tantillus, Growv.*, 111; men-
tioned, 94, 113.
Cyclonotum, Erichson, 212.
punctulatwm, Régimbart, 213.
rufitarse, Boheman, 212.
Cycloteres, Schiénh., mentioned,
406.
rugulosus, Dohrn., mentioned,
405.
Cycloterinus, Kolbe, 409 ; mentioned,
407.
ampliatus, Kolbe, 411; men-
tioned, 394, 408, 410, 496.
504
Cycloterinus cxecus, Champ.*, 415;
mentioned, 394, 407, 409, 410.
canaliculatus, Champ.*, 412;
mentioned, 394, 410, 496.
carinifer, Champ.*, 412; men-
tioned, 394, 407, 410, 413,
496.
erosus, Champ.*, 414; men-
tioned, 394, 410, 496.
foveatus, Kolbe, 411; men-
tioned, 394, 407, 409, 410,
496.
humeroalatus, Kolbe, 410;
mentioned, 394, 407, 411.
microphthalmus, Champ.*, 415 ;
mentioned, 394, 410.
spheropterus, Champ.*, 414;
mentioned, 394, 410, 415.
unicristatus, Champ.*, 413;
mentioned, 394, 407, 408,
410, 412, 414, 496.
Cycloterodes, Kolbe, 404.
sechellarum, Aolbe, 405; men-
tioned, 394, 496.
Cydnine, 140.
Cydnus pallidipennis, Keut., 140.
Cyladina, 450.
Cylapus, Say, mentioned, 174, 177.
migratorius, Distant*, 176 ;
mentioned, 190.
Cylas, Latr., 450.
formicarius, Fabr., 450; men-
tioned, 395.
turcipennis, Boh., 451.
Cylistosoma, Lewis, 225.
Cymbalopora, Hag., mentioned, 49.
Cymine, 150.
Cymospira gigantea, Blainville, 80.
Cyrthorrhinus parviceps, Leut.,
mentioned, 174.
Cyrtop:ltus atricornis, Dzstant*,
180; mentioned, 191.
Dactylosternum, Wollaston, 211.
insulare, Castelnau, 211; men-
tioned, 98, 194, 195, 196,
212, 218, 219.
pygmeum, Légimbart, 211;
mentioned, 194, 195, 212.
subquadratum, Mairm., men-
tioned, 212.
Dasya, C. Ag., mentioned, 299.
elegans (Martens), mentioned,
292.
indica, J. Ag., 297; mentioned,
269, 270.
INDEX
Dasyopsis, Zan., 292; mentioned,
291).
aperta, Web. v. B.*, 292; men-
tioned, 271, 274, 290, 291,
293, 306.
cervicornis, J. Ag., mentioned,
293.
Geppii, Web. v. B.*, 294;
mentioned, 271, 272, 274,
290, 295, 306.
palmatifida, Web. v. B.*, 294;
mentioned, 271, 272, 274,
295, 306.
spinella, Zanard., mentioned,
293.
Stanleyi, Web. v. B.*, 292;
mentioned, 272, 274, 293,
294, 306.
Decapoda, Pelagic, by 8S. Kemp,
53-68.
Delesseriahypoglossoides, Harv.,285.
Delesseriacex, 285; mentioned, 273.
Dendroctonomorphus, Woll., 491.
muricatus, Woll., mentioned,
491, 492.
muriceus, Champ.*, 491; men-
tioned, 395.
Dendy, A., Report on the Cal-
careous Sponges collected by
H.M.S. “Sealark” in the Indian
Ocean, 1—29.
Dendya, Bidder, 6; mentioned, 1.
prolifera, Dendy*, 6; men-
tioned, 1, 28.
tripodifera (Carter), mentioned,
1, &:
Dentalium subulatum,
mentioned, 90.
Derzocoris cardui, Distant*, 181 ;
mentioned, 190.
ostentans, Stal, mentioned, 18.
seychellensis, Distant*, 180,
mentioned, 190.
Derelomus, Schénh., mentioned, 453.
Dermestes hemipterus, Linn., 102.
Dermonema, Grev., 276.
dichotomum, Harv., 276; men-
tioned, 271, 272.
Desmometopa, Loew, 363.
approximatonervis, Lamb*,
365; mentioned, 366, 372.
inauratum, Lamb*, 363; men-
tioned, 364, 372.
microps, Lamb*, 364;
tioned, 365, 372.
M-nigrum, Zeit., 363.
Deshayes,
men-
Dettopsomyia, Zamb*, 349,
formosa, Lamb*, 350; men-
tioned, 371.
Diapelmus, Hrichs., mentioned, 438.
Diastatotropis, mentioned,
248,
Dicercomorpha alluaudi, Kerrem.,
378.
Dictyurus, Bory, 299.
purpurascens, Lory, 299 ; men-
tioned, 271, 272, 274.
Dieuches annulatus (Sign.), 154.
cardui, Distant*, 155; men-
tioned, 190.
placidus (Stal), 155.
Dinia mtida, Reut., 154,
Diniella nitida (feut.), 154.
Dioédimorpha, Broun, 464, 465.
Dipsocoride, 170.
Diptera: Heteroneuride, Ortalide,
Trypetide, Sepsidee, Micropezide,
Drosophilide, Geomyzide, Mili-
chide, by C. G. Lamb, 307-372.
Scatopside, Simuliide, by G.
Enderlein, 373-375.
Distant, W. L., Rhynchota. PartI:
Suborder Heteroptera, 139-191.
Distoleon, Banks, mentioned, 243,
246.
Dracena, Vand., mentioned, 211,
228, 231, 386, 489.
Drosophila, Fallen, 325; mentioned,
338.
sp., 348.
aberrans, Lamb*, 334; men-
tioned, 325, 335, 371.
ampelophila, Loew, mentioned,
347.
caliginosa, Lamb*, 341; men-
tioned, 325, 342, 371.
finitima, Lamb*, 340; men-
tioned, 325, 341, 371.
fracticosta, Lamb*, 329; men-
tioned, 325, 330, 350, 371.
gratiosa, De Mer., mentioned,
329.
grossipalpis, Zamb*, 328 ; men-
tioned, 325, 371.
hirtipes, ZLamb*, 337; men-
tioned, 325, 338, 371.
jucunda, Lamb*, 339; men-
tioned, 325, 371.
Lac.,
melanogaster, Meig., 347;
mentioned, 325, 348.
nasuta, Zamb*, 346; men-
tioned, 325, 371:
Drosophila nigerrima, Zamb*, 331 ;
mentioned, 325, 332, 371.
nigra, De Merj., mentioned,
341.
nigrobrunnea, Lamb*, 332;
mentioned, 325, 333, 371.
notabilis, Lamb*, 329; men-
tioned, 325, 371.
pallipes, ZLamb*, 342; men-
tioned, 325, 343, 371.
_ punctiscutata, Lamb*, 333;
mentioned, 325, 334, 371.
punctulata, Loew, 339; men-
tioned, 325.
repleta, Wol/., mentioned, 339.
rufuloventer, Lamb*, 344;
mentioned, 325, 371.
sericea, Zamb*, 326; men-
tioned, 325, 327, 371.
similis, Zamb*, 347; men-
tioned, 325, 371.
spinipes, Zamb*, 336; men-
tioned, 325, 371.
triangulifer, Lamb*, 343 ; men-
tioned, 325, 344, 371.
Drosophilide, 325.
Dryocetes, Hichh., 389.
furvus, Sampson*, 389.
nubilus, Blandf:, 389.
Dryophthorides, 457.
Dryophthorus, Schénh., 457.
assumilis, Gahan, 457.
corticalis, Say, 457.
ecarinatus, Champ.*, 457;
mentioned, 395.
lymexylon (fabr.), 457 ; men-
tioned, 395, 458.
Dryotribus, Horn, mentioned, 406,
407.
Dulinius nigrolineatus, Distant*,
158; mentioned, 190.
Duognathoferz, 40.
Dysceladus tuberculatus,
Waterh., mentioned, 399.
Dysdereus fasciatus, Sagn., 158.
nigrofasciatus, Stal, 158.
ortus, Distant, 158. —
Dysnos, Pasc., 266.
ethiops, Jord.*, 266;
tioned, 267.
CAO}
men-
Eecoptomma, Geck., 361; mentioned,
370.
solitarium, Lamb*, 361; men-
tioned, 362, 372.
Kccoptopterus, Wotsch., 389.
INDEX
Eccoptopterus sexspinosus, Motsch.,
389.
Echidnocephalus, Lamb*, 351.
barbatus, ZLamb*, 352; men-
tioned, 371, 372.
Echtrichodiinz, 167.
Ellimenistes, Schdnh., mentioned,
398.
Emesine, 163.
Endeopsis, Champ.*, 437.
anthonomoides, Champ.*, 438;
mentioned, 394.
curvimanus, Champ.*, 438;
mentioned, 394, 439, 496.
delicatus, Champ.*, 439; men-
tioned, 394,
Endeus, Schénh., mentioned, 438.
Enderlein, G., Diptera: Scatopside,
Simuliide, 373-375.
Endosiphonia, Zan., mentioned, 270,
287, 298, 299.
clavigera (Wollny), mentioned,
288.
Enochrus abnormicollis, Zaitzev,205.
Enoplurus, Hope, 198.
Ensina vacillans, Wol/., mentioned,
316.
Entium, Sharp, mentioned, 476.
Ephydatia, Zam., mentioned, 4.
Epirrhamphus, Champ.*, 398.
litoralis, Champ.*, 398; men-
tioned, 394, 496.
Hpitaphius, Yawm., 253;
tioned, 247.
licheneus, Jord.*, 253; men-
tioned, 267.
Epurea bifasciata, Krantz, 106.
luéeola, Erichson, 106.
minuta, Reitt., 107.
ocularis, Fairm., 105.
thiemei, Reitt., 106.
Erecanana, Strand, mentioned, 34,
35.
Eremotes, Woll., mentioned, 461.
men-
Erirrhinus, J/ontr., mentioned, 408.
Erodius, Fabr., mentioned, 406.
Kthelia, Web. v. B.*, sub-gen., 304 ;
mentioned, 302, 306.
Hucarphus, Wérch, mentioned, 69,
72, 76.
Kucheuma, J. Ag., 279.
Cottonii, Web. v. B.*, 279;
mentioned, 272, 273, 306.
Schrammi, Crowan, mentioned,
279.
spinosum, dg., mentioned, 279.
905
Hucycloteres, Champ.*, 405.
terreus, Champ.*, 406; men-
tioned, 394, 496.
Eugnomus, Pase., mentioned, 438.
Kugnoristus, Schénh., 492.
braueri, Kolbe, 492;
tioned, 395, 497.
Kuops, Schénh., 395.
Jekel,
men-
bowringi,
395.
_nietneri, Jekel, mentioned, 395.
395 ;
mentioned,
viriditinctus, Champ.*,
mentioned, 394, 496.
(Rhynchites) suffundens, Walk.,
mentioned, 395, 396.
Eupeyssonnelia, Web. v. B.*, 303.
Euplectini, 126; mentioned, 118,
120. :
Kupomatus, Phil., 72, 73.
dipoma, Schm., 73.
elegantulus, Bush, 73.
exaltatus, Marenz., 77; men-
tioned, 70, 73, 78, 92.
floridanus, Bush, mentioned,
73.
gracilis, Bush, mentioned, 73.
heteroceros, Willey, 75.
humilis, Bush, mentioned, 73.
protulicola, Bened., 73.
spinosus, Pixell*, 78; men-
tioned, 70, 73, 92.
spongicola, Bened., 73.
uncinatus, Phil, 73; men-
tioned, 75.
(Hydroides) dianthus, Verrii/,
mentioned, 73.
Eurycyrtus, Reut., 177.
Kuryomma, Stein, mentioned, 279.
Kurystylus, Stal, 177.
alboplagiatus, Distant*, 178;
mentioned, 190.
Eusarcocoris imperator, Dzstant*,
142; mentioned, 189.
V—flavum, Horv., mentioned,
142.
Kusphyrus, Zec., mentioned, 250.
Fauchea, Mont. et Bory, 282.
microspora, Born., 282; men-
tioned, 272, 273.
Feella affinis, Hirst, 37.
Felisacus, Distant, mentioned, 174.
auritulus, Distant*, 177; men-
tioned, 190.
glabratus, .Votsch., mentioned,
177.
506
Filograna (Oken), 87.
implexa, Berkeley, mentioned,
87.
Filopora filigrana, Dalyell, 87.
Formicaleo, Geoffr., mentioned, 243.
bistrigatus (Rambur),
tioned, 245, 246.
ornatus, Weedham*, 245,
Fucus crinalis, Turn., 278.
obtusus, Huds., 285.
rigidus, Vahl, 280.
Valentie, Turn., 282.
Fulvidius, Poppiws, mentioned, 182.
Fulvius, Séa/, mentioned, 174, 181.
men-
dolabratus, Distant*, 181;
mentioned, 191.
niger, Dastant*, 182; men-
tioned, 190.
pictus, Distant*, 181; men-
tioned, 191.
Galaxaura, Lamzx., 276.
collabens, Kjellm., mentioned,
277.
fastigiata, Decne., 277; men-
tioned, 271, 272.
hawaiiana, Butt, 277; men-
tioned, 272, 306.
Liebmanni (Aresch.), Kjellm.,
276; mentioned, 271, 272,
277.
obtusata (Sol.), Lamax., 277;
mentioned, 272.
veprecula, Ajellm., 277; men-
tioned, 271, 272.
Gardena seychellensis,
165; mentioned, 190.
Gelchossa, Kirk = Leptostyla, Stal,
mentioned, 159.
Distant*,
Gelidiacexe, 278; mentioned, 273.
Gelidiopsis, Schmitz, 280; men-
tioned, 301.
rigida (Vahl), Web. v. B., 280;
mentioned, 271, 273.
variabilis (Gerv.), Schmitz, 280 ;
mentioned, 271, 273, 283,
290.
Gelidium, Lamz., 278; mentioned,
301.
erinale (Zurn.), Lamex., 278 ;
mentioned, 271, 273.
pannosum, Grwn., 278; men-
tioned, 271, 273.
rigidum (Vahl), mentioned,
280.
Gelinaria, Sond., mentioned, 301.
INDEX
Gennadas, Bate, 60.
sp., 62; mentioned, 68.
alcocki, Kemp, 62 ;
68.
calmani, Kemp, mentioned, 63.
mentioned,
indicus, Kemp, mentioned, 53.
parvus, Bate, 60; mentioned,
53, 63, 68.
precox, Kemp, mentioned, 63.
scutatus, Bouvier, 61; men-
tioned, 53, 62.
subsp. indicus, Kemp*,
62; mentioned, 53.
sordidus, Kemp, mentioned, 60,
63.
tinayrei, Bouvier, mentioned,
61.
(Genus ignotum) alboclavatus,
Distant*, 171; mentioned, 190.
Geobdella, Whitman, mentioned, 40.
Geographical Distribution, Note
on, 370.
Geomyzidee, 353.
Geotomus sp., 140.
proximus, Sign., 140.
Gerrine, 162.
Gerrinaria, 162,
Gerris (Limnogonus) cereiventris,
Sign., 162.
(——) dolosa (Bergr.), 163.
Gigartinacex, 278 ; mentioned, 273.
Glapharsenus, Raftr., 137.
Gloioderma, J. Ag., 283.
expansa, Web. v. B.*, 283;
mentioned, 272, 273, 306.
Gloiophlea, J. Ag. 276.
articulata, Web. v. B.*, 276;
mentioned, 272, 306.
scinaioides, dAg., mentioned,
276.
Gloiothamnion Schmaitzianwm,
Reinb., 301.
Glossopsis, Bush, mentioned, 69, 72.
Gonocerus clavatus, Sign., 147.
Gononotus (Zec.), mentioned, 406.
Gonotrachelus, Champ.*, 406.
quinquecarinatus, Champ.*,
407; mentioned, 394, 406,
496.
Gracilaria, Grev., 280; mentioned,
279.
compressa (4g.), Grev., 280;
mentioned, 271, 273.
Grantessa, v. Lendenfeld, 19.
hastifera (fow), 19;
tioned, 1, 2, 28.
men-
Grantia, Fleming, 20.
indica, Dendy*, 20; men-
tioned, 1, 28, 29.
labyrinthica, Dendy, men-
tioned, 15, 23.
Grantiide, Dendy, 20.
Grantilla hastifera, Row, 19; men-
tioned, 1.
Grateloupiacee, 301; mentioned,
274.
Griffithsia, C. Ag., 299.
sp., 299; mentioned, 272,
274.
Grouvelle, A. Coleoptera: Niti-
dulide, Heteroceride, 93-116.
Gypsina, Carter, 49; mentioned,
48.
globulus, Rewss, mentioned, 50,
51.
inherens, mentioned, 50, 51.
melobesiodes, Carter, men-
tioned, 49, 50.
plana, Carter, and the Rela-
tions of the Genus, by M.
Lindsay, 45-51.
vesicularis (Carter), mentioned,
50, 51.
Hadrorhinus, Schénh., mentioned,
398.
Hemadipsa, Tennant, mentioned,
39, 40, 41, 42.
Hemadipsine, mentioned, 39.
Halacritus, Schmidt, 229.
algarum, Schmidt, 229; men-
tioned, 230.
Halobates alluaudi, Bergr., 163.
Halobatinaria, 163.
Haloplegma, JMont., 300.
Preissii, Sond., 300;
tioned, 272, 274.
Halymenia (C. Ag.), J. Ag., 301.
men-
Durvillei, Bory, mentioned,
301.
formosa, Harvey, mentioned,
301.
polyclada, A. and #. S. Gepp,
301; mentioned, 274.
—— yar. aldabradensis, Web.
v. B.*, 301 ; mentioned, 272,
274.
Haptoncura, Reitt., 105.
luteola, Reitt., 106.
minuta, Reitt., 107.
nitescens, Grouv., 107.
ocularis, Reitt., 105.
Haptoncus, A. Murray, 105; men-
tioned, 93, 94.
decoratus, Reitt., 106.
dispersus, Growv., mentioned, |
108. |
jloreolus, Sharp, 106. |
luteolus (Zrichson), 106; men- |
tioned, 94. |
minutus (Lvertt.), 107;
tioned, 94.
obscurus, Growv.*, 107 ;
tioned, 94, 106.
ocularis (fairm.), 105 ;
tioned, 94, 98, 211.
pauperculus, Reitt., 106.
Growv., mentioned,
men-
men- |
men-
picinus,
108. ;
pubescens, A. Murray, 106.
sobrinus, Growv., 107; men-
tioned, 94.
subquadratas, Reitt., 106.
tetragonus, A. Murray, 106. |
Harding, W. A., New Land- |
Leech from the Seychelles, 39—_
43.
Harpactorine, 169.
Helminthocladiacez,
tioned, 272.
Helobdella stagnalis (Linn.), men-
tioned, 40.
Helochares, Mulsant, 203.
melanophthalmus, Mulsant,
203; mentioned, 194, 196.
—— var. ngrifrons, Régim-
bart, mentioned, 203.
var. sechellensis, Régim-
bart, 203; mentioned, 194,
196.
nigrifrons, Régimbart, 203.
Hendelia Beckeri,
tioned, 308.
Henicocephalide, 163.
Henicocephalus, sp., 163.
Hermotinus, Distant, mentioned,
174, 180.
villosulus, Distant*, 179; men-
tioned, 190.
Herposiphonia, Vaegeli, 290; men-
tioned, 300.
275; men-
Czerny, men-
prorepens (Harv.), Schmitz,
290; mentioned, 271, 274.
Heteroceride, 116.
Heterocerus, Yabr., 116.
vulpes, Grouv., 116; men-
tioned, 94.
Heterogastrine, 150.
INDEX
Heteromeringia, Czerny, 308.
nigriceps, Lamb*, 310.
nigrifrons, Zamb*, 309; men-
tioned, 310, 371.
plumicornis, Lamb, 308; men-
tioned, 309, 371.
Heteroneuride, 308.
Heteropegma, Poléjaetf, mentioned,
16.
latitubulata (Dendy), 17.
nodus-gordi, Poléjaeff, 16, 17 ;
mentioned, ess
Heteroptide, 19.
| Heteroptera, by W. L. Distant,
139-191.
Heterosiphonia, Mont., 296; men-
tioned, 299.
sp., 296 ; mentioned, 271, 274.
Rendlei, Web. v. B.*, 296;
mentioned, 272, 274, 301,
306.
subsecundata (Suhr), men-
tioned, 296.
Himatinum, Cockerell, 480.
breviusculum, Champ.*, 481 ;
mentioned, 395.
confluum, Champ.*, 480; men-
tioned, 395, 481.
rugipenne, Champ.*, 480; men-
tioned, 395, 481.
variolosum, Woll., mentioned,
480.
Himatium, Woll., 480; mentioned,
479.
Hirst, §., Arachnida, Second Re- |
port.—Scorpions, Pedipalpi, and |
supplementary notes on Opiliones
and Pseudoscorpions, 31-37.
Hispine, 237.
Histeridee, 222.
Holostilpna, Jord., mentioned, 259,
260,
Homalorrhynchus, Champ.*, 452 ;
mentioned, 455.
rubricatus, Champ.*, 453; men-
tioned, 395.
serripes, Champ.*, 453; men- —
tioned, 395, 452, 454.
verschaffeltiz, Champ.*, 454 ;
mentioned, 395, 497.
Homoceelide, 2.
Homotrema, Hickson,
47.
Hophorus gracilirostris, A. M. Edw.,
63.
Hoplophoride, 63.
mentioned,
SECOND SERIES—ZOOLOGY, VOL. XVI.
507
| Hoplophorus, H. MW. Hdw., 63.
foliaceus (Rathbun), 64.
gracilirostris, 4. I. Edw., 63,
64,
Hoplorrhinoides, Champ., men-
tioned, 453.
Hormiscops, Jord.*, 250; men-
tioned, 247, 253.
letus, Jord.*, 252.
sobrinus, Jord.*, 251; men-
tioned, 267.
tessellatus, Jord.*, 252.
tibialis, Jord.*, 251; men-
tioned, 250, 252, 267.
Hughia, Raffr.*, 122; mentioned,
118,120, 125.
carinata, Raffr.*, 124; men-
tioned, 118, 120, 123, 138.
phantasma, Raffr.*, 123; men-
tioned, 118, 120, 125, 138.
Hyalopeplus, Reut. (nec Stal), 174.
Hybomorphus, Sawnd., mentioned,
406.
Hydara tenuicornis (Westw.), 147.
Hydrena, Kugelann, 196.
mahensis, Scott*, 196; men-
tioned, 194, 198.
marginicollis, Légimbart, men-
tioned, 198.
Hydrenini, 196.
Hydroides, Gunn., 72; mentioned,
69, 73.
abbreviata, Ardéyer, mentioned,
73.
albiceps, Lhrenb., 73.
benzoni, Worch, 73.
bifureatus, Pixell*, 76; men-
tioned, 70, 73, 92.
bispinosa, Bush, 73.
crucigera, Mérch, 73.
cunning, Moérch, 73.
dianthus, Verril/, mentioned, 92.
dirampha, Mérch, 73.
heteroceros, Grube, 75; men-
tioned, 70, 72, 73, 76, 92.
homoceros, Pixell*, 74; men-
tioned, 70, 73, 92.
lunulifera, Clap., 73.
minax, Grube, 73.
monoceros, Grav., 76; men-
tioned, 70, 72, 73.
multispinosa, Marenz., 73.
norwegica, Gunn., 74; men-
tioned, 69, 70, 72, 73.
parvus, Treadwell, 73.
pectinata, Phil., mentioned, 73.
65
508
Hydroides serratus, Bush, 73. |
tripanon, Clap., mentioned, 73.
(Eupomatus) elegans, Haswell,
mentioned, 73.
Hydrometra ambulator, Stal, 161.
Hydrometride, 161.
Hydrometrine, 161. |
Hydrophilide, 193. |
Hydrophilini, 198.
Hyginellus, Distant*, 150.
gayei, Distant*, 150;
tioned, 189.
Hyginus, Std, mentioned, 150.
Hylobiina, 404.
Hymendora, Sars, mentioned, 67.
Hypnea, Lama., 281.
sp., 282; mentioned, 271, 273.
divaricata (?), Grev., 281; men-
tioned, 272, 273.
pannosa, J. Ag., 282; men-
tioned, 271, 272, 273, 289.
Valentie (Zurn.), Mont., 282;
mentioned, 271, 273, 281.
Hypocaceus, Thomson, 234.
Hypoglossum, Kiitz., 285. |
sp., 285 ; mentioned, 271, 273. |
spathulatum (Kiitz.), J. Ag.,
285. |
Hypopentarthrum, Champ.*, 462.
mierocephalum, Champ.*, 462; |
mentioned, 395, 497.
Hypothenemus, Westw., 385; men-
tioned, 379.
arecce, Hichh., 385.
eruditus, Westw., 385.
opacus, Hichh., mentioned, 385.
validus, Blandf., 385.
var.valens, Sampson *,385.
Hypselopus intermedius, Distant,
148.
men-
Tbalonius bimaculatus, Loman, +
inscriptus, Loman, 35.
inscriptus, Loman, 35.
+ bimaculatws, Loman, 35.
Tdiobdella, Harding*,
tioned, 40.
seychellensis, Harding*, 41 ;
mentioned, 40, 41, 43.
Ilythea invenata, Lamb, mentioned,
370.
Imatium, Clark, 480.
Ipide, 382.
Tridea, Bory, 278.
sp-, 278; mentioned, 271, 272,
273.
41; men-
INDEX
Tridotenia mahena, Fairm., 377.
Ischnocerus nigellus, Sparrm., men-
tioned, 248, 249.
Ischnonyctes, Sta/, mentioned, 166.
Ischnurus ochropus, C. Li. Koch, 32.
Tsometrus maculatus, De Geer, 32; |
mentioned, 31.
“Jak” (Artocarpus
Innn.), mentioned, 219.
integrifolia,
| Jania, Lamz., 305.
tenella, Avitz., 305; mentioned, |
271, 274.
Jordan, K., Coleoptera :
bide, 247-267.
Jourdainana, Distant*, 157.
rugifer, Distant*,
tioned, 190.
Jussiza, sp., mentioned, 184.
157; men-
Kemp, 8., Pelagic Crustacea Deca-
poda, 53-68.
Kerremans, C., Coleoptera ; Bupres-
tide, 377-378.
Lachnophorus albidomaculatus,
Distant*, 153; mentioned, 189.
singalensis (Dohri), 153.
Leospira, Caullery «& Mesnil, 89.
Lamb, C. G., Diptera: Hetero-
neuride, Ortalide, Trypetidee,
Sepsidee, Micropezide, Drosophi-
lide, Geomyzide, Milichide,
307-372.
Lasiochilus alluaudi, Reuwt., 184 ;
mentioned, 185.
gardineri, Dastant*, 185; men- |
tioned, 191.
praslinensis, Distant*, 186 ;
mentioned, 191.
scotti, Distant*, 184; men-
tioned, 191.
seychellensis, Distant*, 185:
mentioned, 191.
sladeni, Distant *,
tioned, 191.
185 ; men-
Lasiocolpus, Rewt., mentioned, 186. |
Lasiodactylus, Perty, 109; men- |
tioned, 93, 95.
breviusculus, Fairm., 110;
)
mentioned, 94, 95.
costulatus, Reitt., 110.
glabricola, Reitt., 109.
pictus (MacLeay), 109; men-
tioned, 94, 95.
testudinarius, Reitt., 110.
Anthri- |
Lasiotrupis, Champ.*, 478.
clavigera, Champ.*,
mentioned, 395.
Laurencia, Zama., 285.
478 ;
ceylanica, J. Ag., 287; men-
tioned, 271, 272, 273.
indica, Hauck, mentioned, 286.
nidifica, J. <Ag.,
tioned, 272, 273.
obtusa (Huds.), Lame., 285 ;
mentioned, 271,272, 273, 286.
Kiitz.,
285; men-
— var. racemosa,
286.
pannosa, Zanard., mentioned,
286.
papillosa (/orsk.), mentioned,
286, 289.
f. australica, Aiitz., 286;
mentioned, 271, 272, 273.
pygmea, Web. v. B.*, 286;
mentioned, 271, 273, 306.
spinulifera, Avitz., 286; men-
tioned, 271, 273.
Laurenciex, 285.
mentioned,
Leech, Land-, new from the Sey-
chelles, by W. A. Harding, 39—43.
Leptobaris, Gersé., mentioned, 453.
Leptocoris hematica, Germ., 148.
Leptocorisa apicalis, Westw., 147.
Leptoglossus membranaceus(Fabr.),
146. °
Leptomimus delicatulus, Woll., men-
tioned, 471.
fragilis, Woll., mentioned, 471.
Leptostyla, Stal, mentioned, 159.
Lepydnus, Champ.*, 396.
nepenthicola, Champ.*, 397 ;
mentioned, 394, 396, 496.
Lethzeus, Dall, mentioned, 156.
punctus, Bergr., 155;
tioned, 156.
stellatus, Distant*, 155; men-
tioned, 189.
Leucaltide, 16.
Leucaltis bathybia var.
renica, Ridley, 17, 18.
clathria (Haeckel), 16; men-
tioned, 1, 2, 17, 18, 28.
Leucandra, Haeckel, 23; mentioned,
24.
aspera (0. Schmidt),
tioned, 23.
echinata, Schuffner, 23; men-
tioned, 1, 2, 28.
wasinensis (Jenkin), 24 ; men-
tioned, 1, 2, 28.
men-
Masca-
men-
Leucascide, 8.
Leucascus, Dendy, 9; mentioned, 1.
simplex, Dendy, 9; mentioned,
IL, B WO, Be,
Leucetta, Haeckel, 10; mentioned,
13.
chagosensis, Dendy*, 10-11;
mentioned, 1, 12, 28.
microraphis, Haeckel,
tioned, 11, 12.
primigenia, Haeckel, mentioned,
12!
pytiformis, Dendy*, 11 ; men-
tioned, 1, 28.
Leucilla, Haeckel, 25; mentioned, 24.
men-
australiensis, Carter, men-
tioned, 26.
proteus, Dendy*, 25; men-
tioned, 1, 28, 29.
wasinensis, Jenkin, 24.
Leuconia echinata, Carter, 23.
Leucophenga, Mik., mentioned, 325.
maculata (Duf.), mentioned,
326.
Leucosolenia, Bowerbank, 2; men-
tioned, 17.
coriacea var. _ceylonensis,
Dendy, mentioned, 5.
depressa, Dendy, mentioned, 5.
elathrus, 0. Schmidt, mentioned,
27.
gardineri, Dendy*, 2; men-
tioned, 1, 28.
proxima, Dendy, mentioned, 3.
(Clathrina) coriacea,
Dendy, mentioned, 5, 6.
Liagora, Lame., 275.
elongata, J. Ag., mentioned, 275.
hawaiiana, Butt., 275; men-
tioned, 271, 272.
orientalis, J. Ag., 275; men-
tioned, 271, 272.
Limnogonus dolosus, Bergr., 163.
Lindsey, M., on Gypsina plana,
Carter, and the Relations of the
Genus, 45—51.
Liomyza, Zett., mentioned, 351, 352.
Lithothamnia, mentioned, 269.
Lixus, Fabr., mentioned, 417, 420.
Lobotrachelus, Schénh., mentioned,
447.
Lodoicea, Labill., mentioned, 113,
210, 407, 425, 451, 462.
seychellarum, JLabill., men-
tioned, 112, 129, 382, 412,
413, 432, 452, 466, 479.
INDEX
Lomentaria Albertisii, Pice., 284.
intertexta, Chawvin, mentioned,
284.
Lonchza plumata, Zamb, mentioned,
326.
Lordites, Evichson, 109.
costulatus, Fairmaire, 109.
glabricola, Candeze, 109.
Lucifer, V. Thomson, 57.
acestra, Dana (=reynaudii, H.
M. Edw.), mentioned, 53, 57,
58.
pacificus, Dana, mentioned, 57,
58.
reynaudii, (Dana), H. M. Edw.
58-60 ; mentioned, 57, 58.
typus, H. WM. Hdw., mentioned,
57, 58, 60.
Lumetus, Zaitzev, mentioned, 205.
Luteva malayana, Distant, 165.
Lychas braueri (Krep.), 32; men-
tioned, 31.
Lygeide, 149.
Lygus cinnamomeus, Distant*, 178;
mentioned, 191.
sanguineosignatus, Distant*,
178; mentioned, 191.
silhouettensis, Distant*, 179;
mentioned, 191.
Macroscytus fryeri, Distant*, 140.
lobatus, Sign., mentioned,
140.
Madiza, Fallen, 366.
albipennis, Lamb*, 366 ; men-
tioned, 367, 372.
Magdalis, Germ., mentioned, 455.
Mahea sexualis, Distant,
tioned, 139.
Mahisa, Horv. (mec Distant), 152.
Maralauda, Distant*, 182.
lania, Distant*, 183; men-
tioned, 190.
Maruthas, Distant, 151.
fumigatus, Distant*, 151; men-
tioned, 190.
Mascaregnasa typica, Distant, men-
tioned, 163.
Maulik, S., Coleoptera, Chrysome-
lide : Hispinze of the Seychelles,
237-242.
Mecinus, Germ., mentioned, 417.
men-
Megacelum antennatum, Kirby,
mentioned, 174.
flagellatum, Distant*, 174;
mentioned, 191.
509
Megacelum rubrinerve, Std, men-
tioned, 175.
(Creotiades) mimicum, Dis.
tant™, 175 ; mentioned, 191.
Megasternum, Muls., mentioned,
207.
Melanothamnus, Born. et Falk.,
mentioned, 298, 299.
Melarrhinus, Woll., 481; men-
tioned, 458.
incrustatus, OChamp.*, 481 ;
mentioned, 395.
nigritus, Woll., mentioned, 482.
Meligethini, 95.
Melobesia, Zamza., 305.
callithamnioides, Falk., 305 ;
mentioned, 271, 274.
Menida transversa (Sign.)?, 144.
Meroscinis, Meij., mentioned, 370.
Mesobdella, Blanch., mentioned,
39, 40.
Metamasius sericeus, mentioned,
493.
Metazalmoxis ferruginea, Roewer,
37.
Methydrus, Rey, 203.
Microhimatium, Champ.*,
mentioned, 478.
479 ;
pubescens, Champ.*, 479; men-
tioned, 395, 497.
Micropezide, 324.
Microplatymerus, Champ.*, 451.
lodoiceivorus, Champ.*, 452;
mentioned, 395; 451, 497.
Microtrupis, Champ.*, 476 ; men-
tioned, 463.
longipennis, Champ.*,
mentioned, 395, 477,
piligera, Champ.*, 477; men-
tioned, 395, 476.
puncticeps, Champ.*,
mentioned, 395, 476.
Microvelia repentina, Distant, 162;
mentioned, 190.
Milichia, Meigen, 359.
superba, Lamb*,
tioned, 372.
Milichide, 359.
Milichiella, Gzglio-Tos, 360; men-
tioned, 361.
argentea,
361.
lacteipennis, Loew, 360.
tosi, Geck., mentioned, 361.
tristis, ZLamb*, 360; men-
tioned, 361, 372.
65—2
476 ;
477 ;
359 ; men-
Fabr., mentioned,
510
INDEX
Monotoma, Herbst, mentioned, 459, | Northea, Hook. 7, mentioned, 168,
481,
Moquinia, Blanch., mentioned, 39,
40, 41.
Myossita, Pasc., mentioned, 438.
Myothorax, A. Murray, 104.
252, 258, 384, 397, 402, 408, 435,
436.
seychellarum, Hook. 7., men-
| tioned, 215, 404, 408.
| Notiosomus, Wo//., mentioned, 483.
Myrmeleon obscurus, Rambur, 243 ; | Notonectidee, 189.
mentioned, 244.
pervigil, Walker, 246.
Myrmeleonide, 243.
Mystrops, Hrichs., mentioned, 93.
Nabidine, 170.
Nabis capsiformis, Germ., 170.
Nagusta, Stal, 169.
maura, Distant*,
tioned, 190.
pallida, Distant?, mentioned,
169.
Nanus, Schinh. (= Homaloxenus,
Woll.), mentioned, 453, 456.
Needham, J. G., Neuroptera,
Myrmeleonide from the Indian
Ocean, 243-246.
Nepenthes, Zinn., mentioned, 397.
Nepide, 188.
Nereius, Yabr., 324.
alluaudi, Giglio-Tos, 324.
Nesohispa, Maulik*, 237.
lambaciras, Maulik*,
mentioned, 238.
Nessiara, Pasc., mentioned, 248,
251.
Neuroctenus caffer (Stal), 160.
Neuroptera, Myrmeleonide, by J.G.
Needham, 243-246.
Nezara acuta, Bergr. (nec Dall),
143; mentioned, 144.
emmerezi, Schouteden,
tioned, 144.
heegeri (fieb.), 144.
spicata, Distant*, 143; men-
tioned, 144.
viridula (Linn.), 143.
Ninus sechellensis, Bergr., 150.
Nitidula contingens, Walker, 105.
dimidiata, Fabricius, 104.
humeralis, Fabricius, 101.
intendens, Walker, 106.
picta, Macleay, 109.
significans, Walker, 105.
Nitidulids, 93; mentioned, 211.
Nitidulini, 105.
Nitidulopsis equalis, Walker, 97.
Nitophyllum, Grev., 285.
sp., 285; mentioned, 271, 273.
239 ;
men-
169 ; men-
|
Notonectine, 189.
| Notostomus, 4. MW. Hdw., 65, 66.
armata, Smith, mentioned, 66.
brevirostris, Bate, 66; men-
tioned, 68.
corallinus, A. M. Hdw., men-
tioned, 66.
elegans, A. WM. Hdw., 66.
fragilis, axon, 66.
gibbosus, A. MW. Hdw., 66.
japonicus, Bate, 66.
longirostris, Bate, 66.
murrayi, Bate, 66.
patentissimus, Bate, 66.
perlatus, Bate, 66—68.
robustus, Smith, 66.
vescus, Smith, 66.
westergreni, Hazon, 66.
Nysius albipennis, Distant*
Distant,
149.
men-
d
cargadosensis,
tioned, 149.
euphorbie, Horv., 149.
Ocladius, Schénh., mentioned, 406.
Octomicrus, Schaufuss, 129; men-
tioned, 118, 120.
angustatus, /affr., mentioned,
130.
semipunctatus, Haffr.*, 129 ;
mentioned, 118, 121, 138.
Ogeria, Distant*, 173.
insularis, Distant*, 173 ; men-
tioned, 190.
Oligocladus, Web. v. B., 290; men-
tioned, 270.
Boldinghii, Web. v. B., men-
tioned, 290, 291.
Prainii, Web. v. B.*, 290;
mentioned, 271, 274, 291,
306.
pusillus, Web. v. B.*, 291;
mentioned, 271, 274.
Olympiocapsus, Kirk, 177.
Omicrus, Sharp, mentioned, 195,
206, 207.
brevipes, Sharp, mentioned,
207.
Omobathus, Raffr.*, 127; men-
tioned, 118, 120, 128.
Omobathus elegans, Raffr.*, 128;
mentioned, 118, 120, 138.
Omphalopoma, Lang,
84.
Oncocephalus angulatus, Zeut., 166.
sordidus, Stal, 166.
Oncorrhinus,
407.
Oosternum, Sharp, mentioned, 220,
221.
costatum, Sharp, mentioned,
221, 222.
mentioned,
Schinh., mentioned,
sorex (Sharp), mentioned, 221,
222,
Opiliones, Suppl. notes, by 8. Hirst,
34-37.
Oplophorus, A. 17. Hdw., mentioned,
63.
foliaceus, Rathbun, 64.
Orbellis, Distant*, 156.
typicus, Distant*, 156; men-
tioned, 189.
Orchestes, Illiger, 439.
basipictus, Yazrm., mentioned,
439.
Ormiscus, Waterh., mentioned, 250.
Orpha, Pasc., mentioned, 438.
Ortalide, 312.
Orthotemnus, Woll., 484.
filiformis, Champ.*, 484 ; men-
tioned, 395.
reflexus, Woll., mentioned, 484.
Oscinella acuticornis, Lamb (=O.
aucticornis, Beck.), 370.
longipennis, Lamb (=O. colo-
vata, Beck.), 370.
oculata, Lamb (=O. complicata,
Beck.), 370.
Ostorodiasoides, Distant*, 187.
signatus, Distant*, 188 ; men-
tioned, 191.
typicus, Distant*, 187; men-
tioned, 191.
Otiorrhynchine Alatz, 399.
Apter, 396.
Oxycarenine, 151.
Oxycarenus, Horv. (nec Fieb.), 151.
Oxydema, Woll., 483.
elongatum, Pasc., mentioned
484.
fusiforme, Woll., 483; men-
tioned, 395, 484.
Oxyna, £fob.-Desv., 316.
sororcula, Wied., 316.
Oxyomus palmarum, Scott, men-
tioned, 209.
Pachygrontha bipunctata, Stal, 150;
mentioned, 151.
confusa, Distant*, 151.
Pachygronthine, 150.
Pamera sladeni, Distant*, 152;
mentioned, 190.
vineta, Say, 152.
Pandanus, Rumph., mentioned, 39,
42, 416, 417, 423, 432-435.
Hornei, Balf. 7, mentioned,
100, 171, 262, 433, 463.
seychellarum, Balf. 7, men-
tioned, 102, 105, 106.
Paracalocoris, Distant, 177.
Paracymus, Thomson, 201.
zenus, Germar, mentioned, 202.
alluaudianus, Scoté*, 201; men-
tioned, 194, 196, 202.
chalceus, Régimbart,
tioned, 202.
evanescens (Sharp), mentioned,
men-
202.
minor, fégimbari, mentioned,
202.
punctillatus, Mey, mentioned,
202.
scutellaris, Rosenh., mentioned,
202.
Paralasiocolpus, Distant*, 186.
marginatus, Distant*, 187;
mentioned, 191.
piceus, Distant*, 186; men-
tioned, 191.
Parardelio, Hendel, 313.
pilosa, Hendel, 313.
Paribalonius bimaculatus (Loman)
+ inseriptus (Loman), 35.
inscriptus (Loman) + bimacu-
latus (Loman), 35.
Paromalus, Hrichson, 225.
alluaudi, Schmidt, 225; men-
tioned, 223.
biarculus, Marseul, mentioned,
226.
calciger, Scott*, 227; men-
tioned, 223, 235.
gardineri, Scott*, 226; men-
tioned, 223, 224, 227, 228,
235.
mendicus,
225.
Paromicrus, Scott*, 206 ; mentioned,
195, 207.
Lewis, mentioned,
atomus, Scott*, 210; men-
tioned, 194, 206, 207, 208,
234.
INDEX
Paromicrus carinatus, Scott*, 208 ;
mentioned, 194, 206, 207,
209, 210, 234.
thomasseti, Scott*, 209; men-
tioned, 194, 206, 207, 208,
210, 234.
Paromius apicatus (Séa/), 152.
seychellesus (Walk.), 152.
Paroosternum, Scoté*, 220; men-
tioned, 195.
degayanuin, Scott*, 222; men-
tioned, 194, 221, 235.
Paroscinella, Beck., mentioned, 370.
Passiflora feetida, Zinn., mentioned,
316.
Pedipalpi, by S. Hirst, 33.
Peletophila, Becker, mentioned, 353.
Peneide, 60.
Pentacoptus, Woll., mentioned, 458.
Pentarthrides, 458.
Pentarthrum, Woll., 459.
angustissimum, Woll., men-
tioned, 463.
apicale, Brown, mentioned,
461.
asperirostre, rown, men-
tioned, 464.
debile, Sharp, mentioned, 465.
fulvicorne, Brown, mentioned,
476.
helmsianum, Sharp, mentioned,
464,
huttoni, Woll., mentioned, 461.
longicolle, Champ.*, 459 ; men-
tioned, 395, 460, 497.
nigrum, Woll., mentioned, 459.
prolixum, Sharp, mentioned,
465.
punctatissimum, Champ.*, 460;
mentioned, 395, 459.
scotti, Champ.*, 461;
tioned, 395.
seychellarum, Ohamp.*, 460;
mentioned, 395, 461.
Pentatomide, 139.
Pentatomine, 142.
men-
Pericharax, Poléjaeff, 12; men-
tioned, 1, 13.
carteri, Poléjaeff; mentioned,
12, 13.
var. heteroraphis, Polé-
jaeff, 13; mentioned, 14.
var. homoraphis, Poléjaeff,
mentioned, 13.
heteroraphis, Poléjaeff; 13; men-
tioned, 1, 2, 15, 16, 28, 29.
511
Pericharax peziza, Dendy*, 15;
mentioned, 1, 28, 29.
Peyssonnelia, Decaisne, 303; men-
tioned, 302.
sp., 304; mentioned, 274.
biradiata, Web. v. B.*, 304;
mentioned, 271, 274.
calcea, Heydr., 303 ; mentioned,
271, 274.
coccinea, J. Ag., 303; men-
tioned, 271, 272, 274.
conchicola, Picc. et Grum.,
mentioned, 303.
Harveyana, Crouan, 303;
mentioned, 270, 272, 274,
- 306.
polymorpha (Zan.), Schmitz,
303 ; mentioned, 304.
—— f. Gardineri, Wed. v. B.*,
303; mentioned, 272, 274,
304.
Phalangodes, Vellkampf, 36.
nanus, Hirst*, 36.
querilhaci, Lucas, mentioned,
36.
Phalangodide, 34.
Phantazomerus, Jacquelin du Val,
110.
Phaulimia, Pasc., mentioned, 247,
250.
Philemon, Blanch., mentioned, 39,
40, 41.
Philiopsis, Hafr., mentioned, 118,
127.
Philydrus, Solier, 203, 205.
abnormalis, Sharp, 205; men-
tioned, 194, 196.
ferrugatus, Régimbart,
tioned, 205.
fragilis, Sharp, mentioned, 204.
mollis, Régimbart, 205.
nigritulus, Légimbart,
tioned, 205.
parvulus, Jezche, 203; men-
tioned, 194, 196, 204, 206.
204 ;
men-
men-
var., mentioned,
196.
rubricollis, Régimbart, 205.
(Methyarus) parvulus, Reiche,
203.
Phleobius, Schénh., 254.
affinis, Kolbe, mentioned, 255.
albescens, Jord., mentioned,
254,
albomaculatus, Allard, men-
tioned, 254.
65—3
512
Phieobius alternans, Wred., men-
tioned, 254.
catenatus, Kolbe, mentioned,
255.
ceylonicus, Jord., mentioned,
254,
compressicornis, Gylh., men-
tioned, 255.
cordiger, Fahrs., mentioned,
254,
gigas, Fabr., mentioned, 254.
cervinus (Klug), 255;
mentioned, 267.
griseus, Fabr., mentioned,
255.
humilis, Kolbe, mentioned, 254.
hypoxanthus, Jord., mentioned,
255.
nigroungulatus, G'ylh.,
tioned, 254, 255.
nigroungulatus, Lac., 255.
pallipes, Jord., mentioned, 254.
men-
papuanus, Jord., mentioned,
255.
pilipes, Jord., mentioned, 254.
podicalis, Kolbe, mentioned,
255.
pustulosus, Gerst., 256; men-
tioned, 247, 255.
quadricommatus, Qued., men-
tioned, 255.
striata, Jord., mentioned, 255.
variegatus, Kolbe, mentioned,
254,
Phleophagosoma, Woll., 482.
aldabranum, Champ.*, 483;
mentioned, 395.
angustulum, Woll., mentioned,
483.
conicicolle, Champ.*, 482; men-
tioned, 395.
dubium, Gahan, 486; men-
tioned, 482, 485.
minutum, Woll., mentioned,
482.
proximum, Woll., mentioned,
483.
puncticolle, Woll., mentioned,
483.
tenue, Woll., mentioned, 483.
Pheenicobates, Champ.*, 417 ; men-
tioned, 394, 416, 435.
alatus, Champ.*, 433; men-
tioned, 394, 419, 434, 435.
albosetosus, Champ.*, 422;
mentioned, 394, 418, 421,431.
INDEX
Pheenicobates curvipes, Champ.*,
426; mentioned, 394, 418,
425, 427.
cuspidatus, Ohamp.*, 431;
mentioned, 394, 419, 496.
cylindricus, Champ.*, 420,
mentioned, 394, 418, 422.
depressirostris, Champ.*, 425 ;
mentioned, 394, 418, 427.
duplovestitus, Champ.*, 423;
mentioned, 394, 418, 424,
431.
flexirostris, Champ.*, 420;
mentioned, 394, 418, 425,
496.
fortirostris, Champ.*, 423,
mentioned, 394, 418.
foveiventris, Champ.*, 425;
mentioned, 394, 418.
gibbirostris, Champ.*, 429;
mentioned, 394, 418, 427,
496.
hispidulus, Champ.*, 431,
mentioned, 394, 419, 496.
lodoicer, Champ.*, 432; men-
tioned, 394, 419.
nigrolimbatus, Champ.*, 422 ;
mentioned, 394, 418,
423.
pandanicola, Champ.*, 433;
mentioned, 394, 417, 419,
496.
parallelus, Champ.*, 421;
mentioned, 394, 418.
peropacus, Champ.*, 428;
mentioned, 394, 418.
praslinensis, Champ.*, 424;
mentioned, 394, 418.
rufitarsis, Champ.*, 434; men-
tioned, 394, 419.
sebertensis, Champ.*, 434;
mentioned, 394, 419.
simplex, Champ.*, 427; men-
tioned, 394, 418.
stevensonie, Champ.*, 430;
mentioned, 394, 418, 429,
431.
stricticollis, Champ.*, 429;
mentioned, 394, 418, 431,
496.
tenuis, Champ.*, 427; men-
tioned, 394, 418.
vittatus, Champ.*, 419; men-
tioned, 394, 416, 417, 420,
496.
Pheenicobatina*, 416.
Pheenicobatopsis, Champ.*, 435;
mentioned, 416.
echinatus, Champ.*, 435;
mentioned, 394, 436, 496.
seticeps, Champ.*, 436; men-
tioned, 394.
Phenicobius, érch,
417.
Pheenicogenus, Champ.*, 437 ; men-
tioned, 416.
inermis, Champ.*, 437; men-
tioned, 394, 496.
Pholidoforus, Woll., mentioned, 458.
Phytobella, Blanch., mentioned, 39,
40.
Picaultia, Distant*, 161.
pronotalis, Distant*, 162; men-
tioned, 190.
Pictinus invalidus, Bergr., 159.
varicolor, Distant*, 160; men-
tioned, 190.
Pinna, Zinn., mentioned, 89.
Pixell, H. L. M., Polycheta of
Indian Ocean, together with some
species from the Cape Verde
Islands. The Serpulidz, 69—92.
Placostegus, Phil., mentioned, 85.
Plearia madagascariensis, Westw.,
165.
Plagiostenopterina, Hendel, 312.
cyanosoma, Hendel, 313 ; men-
tioned, 312.
ruticeps, Hendel, 312.
submetallica, Loew, 313; men-
tioned, 370.
mentioned,
Planobdella, Blanch., mentioned,
39, 40.
Planorbulina, D’Orb., mentioned,
49.
Platymerus, Schénh., mentioned,
451,
Platypodide, 382.
Platypus, Herbst, 382.
lepidus, Chap., 382.
Platysoma, Leach, 224.
conditum, Marseuwl,.mentioned,
224,
richteri, Schmidt, 225; men-
tioned, 222, 223.
Schmidt,
mentioned, 223.
(Cylistosoma) richteri, Schmidt,
225. :
Platystomine, 312.
Plea letourneuxi, Szgn., 189.
Pleine, 189.
224 ;
tenuimargo, 5
Plociomerus parvula,
tioned, 152.
reductus, Walk., 152.
seychellesus, Walk., 152.
vineta, Say, mentioned, 152.
Dall, men-
Ploiariola mixta, Distant, men-
tioned, 164.
morstalli, Schwm., mentioned,
164.
scotti, Distant*, 163; men-
tioned, 190.
Podoctis, Thorell, mentioned, 34,
35.
Peeciloscytus,
179.
Pogonorrhinus, Hutton, mentioned,
452.
Polychzeta—Serpulide, by H. L. M.
Pixeli, 69-92.
Polycelia, J. Ag., 278.
van Hoevellii, Web. v. B.*, 278;
mentioned, 271, 273.
Polysiphonia, G'rev., 289 ;
tioned, 291.
sp., 289; mentioned, 271, 274.
Polytoxus modestus, Distant*, 166;
mentioned, 190.
Polytrema, sso, mentioned, 45,
48, 49, 51.
planum,
49, 50.
Polytus, Faust, 493.
mellerborgi, Boh., 494; men-
tioned, 395.
Pomatoceropsis contiert, Gravier, 83.
Pomatoceros, Phil., mentioned, 81.
Fieber, mentioned,
men-
Carter, mentioned,
triqueter, Zinn., mentioned, 80,
83, 85.
Pomatoleios, Prxell*, 85;
tioned, 69, 70.
crosslandi, Pixell*, 85; men-
tioned, 70, 92.
Pomatostegus (Schmarda), 78.
brachysoma, Schmarda, men-
men-
tioned, 79.
macrosoma, Schmarda, men-
tioned, 79.
stellatus (Abildgaard), 79;
mentioned, 70.
Pria, Steph., 95; mentioned, 93.
nitidor, Grouwv.*, 96;
tioned, 93, 94.
scotti, Grouv.*, 95 ; mentioned,
93, 94.
Proboscidocoris, eut., mentioned,
179.
men-
INDEX
Proboscidocoris intermedius, Popp.,
mentioned, 179.
pluto, Distant*, 179; men-
tioned, 190.
Proconus, brown, mentioned,
464.
Proéces, Schdnh., 488.
* compressicollis, Champ.*, 488 ;
mentioned, 395.
macer, Boh., mentioned, 488.
nigrifrons, Boh., mentioned,
476, 489.
rufipes, Mairm., 467; men-
tioned, 457, 488.
silvestris (Kolbe), 489; men-
tioned, 395, 488, 490.
Promecotheca, Blanch., mentioned,
241.
Protula, isso, 88.
dystert, Huxley, 87.
intestinum, Zam, 88; men-
tioned, 70.
tubularia, MJont., 88; men-
tioned, 70, 90.
Psallus mahensis, Distant*, 184;
mentioned, 191.
Psectrosciara, Keifer, 373.
mahensis, Keifer, 373.
Pselaphide, 117-138.
Pseudendosiphonia, Web. v. B.*,
287; mentioned, 270, 288.
Gardineri, Web. v. B.*, 287;
mentioned, 272, 273, 306.
Pseudeuxesta, Hendel, 314.
prima, Ost.-Sack., 314.
Pseudobaris, Le Conte, mentioned,
450.
Pseudocastalia fryeri, Kerrem.*,
377.
Pseudomus, Schénh., mentioned,
447.
Pseudophleinz, 146.
Pseudoscorpions, 31, 32;
note on, by 8. Hirst, 37.
Pyrrhocoride, 157.
Pyrrhocorine, 157.
Pyxidicerini, 122; mentioned, 118,
126.
Pyxidicerus, Motsch.,
118; 123) 125.
Suppl.
mentioned,
Quinssyana, Distant*, 167.
funeralis, Distant*, 168.
typicalis, Distant*, 167 ; men-
tioned, 168, 190.
varicolor, Distant*, 168.
513
Raffray, A., Coleoptera, Pselaphidee
de VAvchipel des
117-138.
Ralfsia expansa, J. Ag., mentioned,
302.
Ranatra grandocula, Bergr., 188.
Reduviide, 163.
Regimbartia, Zaitzev, 200.
Reichenbachia, mentioned,
121.
Reinboldiella, De Toni, 301.
Schmitziana (Kdd.), De Toni,
301 ; mentioned, 271, 274.
Rhabdocheta, De Mevj., 320; men-
tioned, 370.
spinosa, Lamb*, 320;
tioned, 321, 371.
Rhabdotohispa, Maulik*, 239
mentioned, 237.
Maulik*®, 241;
239, 240.
Rhagovelia nigricans (Burm.), 161.
Seychelles,
Spr.,
men-
scotti,
tioned,
men-
Rhaphigaster transversa, Sign., 144.
Rhegmoclema, Hnderl., 373.
rufithorax, Hnderl., 373 ; men-
tioned, 374.
Rhetogenes, Champ.*, 444.
sexcristatus, Champ.*, 445;
mentioned, 394, 444, 497.
spurcus, Champ.*, 445 ; men-
tioned, 394.
Rhicnoessa, Loew, 367.
aurisetulosa, Lamb*, 368;
mentioned, 369, 372.
ferruginea, Lamb*, 367 ; men-
tioned, 368, 372.
Rhina, Oliv., mentioned, 456.
Rhinanisus, Brown, mentioned,
472, 476.
Rhizophagus, Herbst, mentioned,
470.
Rhodoglossum, J. Ag., 278.
Rhodomelacex, 285; mentioned,
273, 274.
Rhodophycee, 269-306.
Rhodophyllidacez, 278.
Rhodophyllis peltata, Grwn., men-
tioned, 283.
Rhodymenia prolifera, Harv., 281.
Rhodymeniacee, 282 ;
273.
Rhombosoma, Champ.*, 447.
acuminatum, Champ.*,
mentioned, 394.
Rhynchenus [Clairv.], 439 ; men-
tioned, 438.
447 ;
mentioned,
/,
peo 2 wVQem |
pes LIBRARY}sp
Wg BO-e Ups
Mase OD
%
514
Rhynchzonus lonicerze, Herbst, men-
tioned, 440.
spissus, Champ.*, 440; men-
tioned, 394, 496.
Rhynchophorus linearis, Herbst,
494.
Rhynchota, Part I. Heteroptera,
by W. L. Distant, 139-191.
Rhyncolosoma, Champ.*, 485; men-
tioned, 482.
dubium, Gahan, 486; men-
tioned, 395.
Rhyncolus, Crewtz., mentioned,
460.
ater (Linn.), mentioned, 485.
Rhyparochromus annulatus, Sign.,
154.
apicatus, Stal, 152.
circumeinctus, Walk., 157.
singalensis, Dohrn., 153.
Rochonia, Distant*, 168,
galeatus, Distant *, 169 ; men-
tioned, 190.
Roschera, Sonder, 289; mentioned,
171, 242.
africana,
290.
glomerulata (Ag.), n. comb.,
289 ; mentioned, 274.
(Tolypiocladia) glomerulata
(Ag.), mentioned, 271.
Roslania, Distont*, 165.
insularis, Distant*, 166; men-
tioned, 190.
Rybaxis, Sawley, mentioned, 121.
Sonder, mentioned,
Saicinze, 166.
Salda insignis, Distant*, 170; men-
tioned, 190.
Saldidee, 170.
Salmacina, Clap., 87; mentioned,
78.
edificatrix, mentioned, 87.
dysteri (Hualey), 87; men-
tioned, 70, 86, 90.
dystert, Giard, 87.
incrustans, Clap., mentioned,
87.
(Filograna) huxleyi,
mentioned, 87.
Sampson, W., Coleoptera; Platy-
podidze and Ipidz from the Sey-
chelles Islands, 379-391.
Saprinus, Hrichson, 233.
apricarius, Hrichson, 234; men-
tioned, 223, 224.
Ehlers,
INDEX
Saprinus disjunctus, Marseul, 234;
mentioned, 223.
erichsoni, Marseul, 233; men-
tioned, 222, 223.
(Hypocaccus) apricarius, Hrich-
son, 234.
( ) disjunctus, Marsewl, 234.
Sapromyza, Fall., 370.
nudiuscula, Lamb, 370.
striata, Lamb, mentioned, 370.
Sapromyzide, Addenda and corri-
genda, 370.
Sarconema, Zan., 278.
furcellatum, Zan., 278; men-
tioned, 271, 273.
Scaevola Koenigii, Vahl, mentioned,
Halley.
Scantius forsteri (fabr.), 157.
Scatopside, 373.
Schizomus latipes,
tioned, 34.
similis, Hirst*, 33-34.
Schmidelia racemosa, Linn., men-
tioned, 144.
Scholastes, Loew, 313.
cinctus, Guérin, 313.
Hams., men-
vicarius, Hendel, 313.
Sciatrophus, Sampson*, 380; men-
tioned, 379.
fuscus, Sampson*, 380; men-
tioned, 381, 382.
Scinaia, Bivona, mentioned, 276.
Scirtetinus, Jord.*, 259;
tioned, 263.
dimidiatus, Jord.*, 261; men-
tioned, 267.
eumelas, Jord.*, 260; men-
tioned, 259, 261, 267.
luteipes, Jord.*, 261.
piceus, Jord.*, 262.
Scoloposcelis, Popp. nec Fieb, 188.
parallelus, Popp., 188.
men-
picicornis, Popp., 188.
Scorpions, by 8. Hirst, 32.
Scotolemon, Luc., mentioned, 36.
Scott, H., Coleoptera; Hydro-
philide, Histeride, 193-235.
Scytha, Champ.*, 447.
aldabrana, Champ.*, 448; men-
tioned, 394.
Sepina antennata, Distant*, 145.
seychellensis, Distant, 145.
Sepside, 323.
Sepsis, (allen, 323.
albopunctata, Lamb*, 323;
mentioned, 371.
Sepsis rufa, Macg., 323.
Sergestes, 1. Hdw., 54; mentioned,
56, 65.
bisuleatus, Wood-Mason, men-
tioned, 56.
challengeri, Hansen,
mentioned, 56, 68.
edwardsi, Aréyer, 56, 57.
gardineri, Kemp*, 55-56;
mentioned, 68.
gloriosus, Stebbing, mentioned,
54, 55, 56.
halia, Yaxon, mentioned, 57.
54-55 ;
henseni, Ortmann, mentioned,
Sie
incertus, AMansen, mentioned,
57.
japonicus, Gate, mentioned, 56.
kroyeri, Bate, mentioned, 56.
prehensilis, ate, mentioned,
56.
profundus, Sate, mentioned,
56.
robustus, Smzth, mentioned, 56.
Sergestide, 54.
Serinetha ethiopis, Distant, men-
tioned, 148.
fraterna,
148.
hematica (Germ.), 148.
toricollis, Bergr., 148.
Serpula, Zinn., 71 ; mentioned, 72,
73, 91.
aspera, Philippi, 71.
contortriplicata, Savigny, 71.
crater, Clap., 71.
Westw., mentioned,
echinata, Gmelin, 71.
JSascicularis, Lamarck, 71.
gigantea, Pallas, 80.
imbutiformis, Chiaje, 71.
infundibulum, Chiaje, 71.
narconensis, Baird, 71.
var. magellanica, McIn-
tosh, 71.
octocostata, Quatrefages, 71.
pallida, Philippi, 71.
patagonica, Grube, 71.
philippi, Méorch, 71.
quadricornis, Grube, 82.
vermicularis, Zinn., 71 ; men-
tioned, 69, 90.
(Hydroides) uncinata, Grav.,
Ua
Serpulide, by H. L. M. Pixell,
69-92.
Sertularia, Zinn., mentioned, 89.
Sesellus, Distant, 188.
parallelus (Motsch.), 188; men-
tioned, 184, 191.
Seychelles, Coleoptera, Chryso-
melide: Hispinee from the, by
S. Maulik, 237-242.
Platypodide and Ipide
from the, by W. Sampson,
379-391.
Pselaphide de ]’Archipel
des, by A. Raffray, 117-138.
New Land-Leech from the, by
W. A. Harding, 39-43.
Seychellesanus, Distant*, 171.
picturatus, Distant*, 172;
mentioned, 190.
typicus, Dastant*, 172; men-
tioned, 173, 190.
variegatus, Distant*, 172;
mentioned, 190.
var., 172; mentioned,
190.
Simuliide, 374.
Simulium, Latredlle, 374.
damnosum, Zheob., mentioned,
375.
griseicollis, Beck., mentioned,
375.
ruficorne, JMacq., mentioned,
375.
speculiventre, Hnderl.*, 374.
Sintor, Schénh., mentioned, 248,
249.
Sintorops, Jord.*, 248.
alleeus, Jord.*, 250;
tioned, 248, 267.
Sitalces, #. Simon, mentioned, 34,
35.
Sitophilus, Schonh., 494.
exaratus, Boh., 495.
men-
mellerborgu, Boh., 494; men-
tioned, 493.
stigmaticollis, Gyll., 495.
subfasciatus, Boh., 495.
subsignatus, Gyll., 495.
Sjéstedtina, Schout., 147.
Smicronyx, Schénh., mentioned, 417.
Sphadasmus, Schénh., 448.
granocostatus, Paum.
Kolbe), 449 ;
394, 448.
Spheridiini, 211.
Spheridium
213.
Spherococcacez, 280; mentioned,
273.
(nec
mentioned,
punctulatum, Klug,
INDEX
Sphenophorus cribricolls, Walk.,
493.
eruciger, Motsch., 495.
exquisitus, Walk., 495.
sordidus, Gyll., 493.
strangulatus, Gyll., 49
Sphodrias, Champ.*, 454.
magdaloides, Champ.*,
mentioned, 395, 497.
Baly,
2
wv.
455 ;
Spilispa imperialis, men-
tioned, 384.
Spirobranchus, Blainville, 79; men-
tioned, 69, 78, 81.
contierei, Gravier, 83; men-
tioned, 70, 90, 92.
dendropoma, M/dérch, mentioned,
83.
gardineri, Pixell*, 81; men-
tioned, 70, 92.
giganteus, Pallas, 80; men-
tioned, 70, 82, 92.
var. turbinatus, Prxell*,
81; mentioned, 70.
incrassatus, Hh/ers, mentioned,
80.
maldiviensis, Pimell*, 84; men-
tioned, 70, 92.
semperi, Morch, 82; mentioned,
70, 83, 84.
var. aceros, Pixell*, 83 ;
mentioned, 70.
—— var: Willey,
82; mentioned, 70, 84.
tricornis (Grube), mentioned,
80.
Spirorbis, Daudin, 88; mentioned,
7, Vile
cornuarietis, Phil., 89; men-
tioned, 70.
levis, Quatrefages, 89; men-
tioned, 70.
papillatus, Pixell*, 89; men-
tioned, 70, 92.
Sponges, Calcareous, by A. Dendy,
1-29.
Sponsor pilosellus, Kerrem.*, 378.
Sporadotrema, Hickson, mentioned,
47.
Spyridia, Harv., 300.
filamentosa (Wulf), Harv.,
300; mentioned, 272, 274.
302; mentioned,
acroceros,
Squamariacee,
274.
Stagnomorpha, Wollaston, 110.
Stelidota, Hrichson, 108 ;
tioned, 93, 94.
men-
515
Stelidota explanata, Growv.*, 108 ;
mentioned, 94, 109.
Stenocephalus punctipes, Stal, 147.
Stenolemus madagascariensis
(Westw.), 165.
Stenomimus, Woll., 490; mentioned,
478.
fryi, Woll., mentioned, 490.
guatemalensis, Champ., men-
tioned, 490.
nitida, Brown, mentioned, 490.
orientalis, Champ.*, 490; men-
tioned, 395.
Stenopentarthrum, Champ.*, 463.
pandane, Champ.*, 463; men-
' tioned, 395.
Stenopodinz, 166.
Stenoscelis, Woll., 491.
crassifrons, Woll., 491; men-
tioned, 395.
hylastoides, Woll., mentioned,
491.
Stenotrupis, Woll., 464; mentioned,
457, 488.
biformis, Champ.*, 470; men-
tioned, 395, 465.
caliginosa, Champ.*, 471; men-
tioned, 395, 466, 472.
conicicephala, Champ.*, 472;
mentioned, 395, 466.
convexiuscula, Champ.*, 467 ;
mentioned, 395, 465.
crassifrons, JWoll., mentioned,
466, 469.
crassipes, Champ.*, 468 ; men-
tioned, 395, 465, 466.
dumetorum, Champ.*, 475;
mentioned, 395, 465, 466.
exilis, Pasc., mentioned, 473.
filum, Champ.*, 469 ;
tioned, 395, 465, 474.
gracilis, Woll., mentioned, 463,
476.
lodoiceicola, Champ.*, 466;
mentioned, 395, 465,467,474.
nemoralis, Champ.*, 472; men-
tioned, 395, 466.
nitidula, Champ.*, 474; men-
tioned, 395, 466.
parallela, Champ.*, 468; men-
tioned, 395, 465, 466, 467,
470.
polita, Champ.*, 473; men-
tioned, 395, 466, 475.
rufipes (Hairm.), 467; men-
tioned, 395, 465.
men-
516
Stenotrupis sericata, Champ.*, 473;
mentioned, 395, 466, 474.
silvicola, Champ.*, 475; men-
tioned, 395, 468, 473, 474,
477.
tarsalis, Champ.*, 470; men-
tioned, 395, 465.
(Pentarthrum) prolixa, Sharp,
mentioned, 465, 468, 471.
) wollastoniana, Sharp,
mentioned, 465, 469.
Stenozygum aldabranum, Distant™,
142; mentioned, 143, 189.
insularum, Distant*, 143.
Stephanoderes, Hichh., 384; men-
tioned, 379.
arecce, Hichh., 385.
communis, Schauf., 384.
vulgaris, Schauf:, 384.
Stevensonia, Dwnc., mentioned, 108,
160, 165, 171-173, 177, 209, 210,
239, 242, 391, 416, 430, 476.
Sthenarus leucophilus, Rewt., 183;
mentioned, 184.
Sthenarusoides, Distant*, 183.
montanus, Distant*, 183; men-
tioned, 190.
Stilbocara, Broun, 490.
nitida, Broun, mentioned, 490.
solivagus, Bergr., mentioned,
(
149.
(Phleophagosoma) constricti-
collis, Brown, mentioned,
490.
Stragularia, Stromf., mentioned,
304.
Streblocladia, Schmitz, mentioned,
299.
Strombus gigas, Zinn., mentioned,
50.
Strophosomus,
398.
Styphloderes, Woll., mentioned, 407.
Sunorfa, Raffr., 134; mentioned,
118, 120, 135.
exsculpta, Schauf., mentioned,
134.
fasciculata, Raffr.*, 134; men-
tioned, 119, 120.
Sunorfoides, Raffr.*, 134; men-
tioned, 118, 119, 120, 135, 138.
bicolor, Rafr.*, 136; men-
tioned, 118, 120, 135, 137,
138.
nigripennis, Rafr.*, 136; men-
tioned, 118, 120, 135, 138.
Billb., mentioned,
INDEX
Sunorfoides picea, Raffr.*, 137 ;
mentioned, 118, 120, 135.
Raffr.*, 136;
118, 120, 135.
punctipennis,
mentioned,
Sycettide, 18.
Sycites, Champ.*, 455.
pilicornis, Champ.*, 456; men-
tioned, 395, 497.
Sycon, fzsso, 18.
sp., 18.
Synaptops, Jekel, mentioned, 395.
Syromastes horrida, Germ., 146.
Systaltocerus platyrhinus, abr.
and Imh., mentioned, 248.
Tachyoryetes audax, Zhos., men-
‘tioned, 32.
“Takamaka” (Calophyllum Ino-
phyllum, Zinm.), mentioned, 487.
Tanyomus, Champ.*, 408.
palmicola, Champ.*, 409; men-
tioned, 394, 407, 408, 496.
Tanypleurus, Steenstr., mentioned,
120.
Tapeinodasya, Web. v. B., 295;
mentioned, 270.
Bornetii, Web. v. B., men-
tioned, 295, 296.
Ethele, Web. v. B.*, 295;
mentioned, 272, 274, 296,
306.
Tarantula scaber, Gerv., 34; men-
tioned, 31.
Temnorrhamphus, Champ.*, 461.
latirostris, Champ.*, 461; men-
tioned, 395.
Tenosius, Stal, 147.
proletarius, Schawm., 147.
robustus, Distant, mentioned,
147.
sjéstedti, Schowt., mentioned,
147.
Tephritis, Latr., 319.
aldabrensis, Zamb*, 319; men-
tioned, 371.
Terebella stellata, Abildgaard, 79.
Terminalia Catappa, Zinn., men-
tioned, 487.
Tessaratomine, 145.
Tetragonorrhamphus, Champ.*, 486.
tuberculirostris, Champ.*, 487 ;
mentioned, 395, 486, 497.
Thalattodora, Perkins, mentioned,
406.
Theognete,
407.
Champ., mentioned,
Thesiastes, Casey, 129 ; mentioned,
120.
cordicollis, Raffr., 129; men-
tioned, 120, 121.
Tingidide, 158.
Tinoporus, Carpenter, mentioned,
48, 49.
versicularis, Carter, mentioned,
51.
Tolypiocladia glomerulata, Schmidt,
289, 290.
Tomicus perforans, Woll., 387.
Tomolips, Woll., mentioned, 480.
Tophoderes, Dej.,
247.
Tournefortia argentea, Zinn., men-
tioned, 401.
Trapezirrhynchus, Champ.*, 464.
silhouettensis, Champ.*, 464 ;
mentioned, 395, 497.
Triarmocerus, Hichh., 389; men-
tioned, 397.
mentioned,
sp.4, 389.
birmanus, Hichh., mentioned,
379.
cryphaloides, Hichh., men-
tioned, 379.
Trignathofere, 40.
Trigonotylus ruficornis var. tenuis,
Reuter, mentioned, 174.
Trimiomorphus, Aafr., mentioned,
118, 127.
Trimium, Awbé, mentioned, 118.
Triomicrus, Sharp, 133; mentioned,
119, 120, 121.
seychellensis, Raffr.*, 133 ;
mentioned, 119, 120, 134,
138.
Trocheta, Blainv., mentioned, 39,
40.
Trochorhopalus, Kirsch, 492.
strangulatus, Gyll., 493; men-
tioned, 395.
Trypeta capitata, Wied., mentioned,
316.
Trypetide, 316.
Trypetina, Lacord., 452.
Trypodendron, Steph., mentioned,
381.
Tubus vermicularis, Ellis, 71.
Turritella?, Zam., mentioned, 84.
Tychini, 134; mentioned, 119, 120,
133.
Tyrini, mentioned, 121.
Tytthomimus, Champ., mentioned,
478.
Udeus, Champ., mentioned, 438.
Udotea, Zama., mentioned, 284,
304.
Ulidine, 314.
Urophorus, Murray, 101.
Vannius, Distant, mentioned, 174.
annulicornis, Poppius,
tioned, 176.
mahensis*, Distant, 176; men-
tioned, 190.
Velia magricans, Burm., 161.
Veliine, 161.
Vermilia pygidialis, Willey, 86.
Vermiliopsis, St-Joseph, 86.
glandigerius, Grav., mentioned,
86.
infundibulum,
tioned, 87.
pygidialis, Wolley, 86; men-
tioned, 70, 87, 92.
Verschaffeltia, H. Wendl., mentioned,
210, 224, 382, 430, 453, 454,
469, 473.
splendida, H. Wendl.,
tioned, 416, 492.
Vilius, Sta, mentioned, 169.
men-
Lang, men-
men-
Vollenhovia levithorax subsp. al- |
luaudi, Hmery, mentioned, 131.
Volvulus, Brwilé, 200; mentioned,
196.
zneus, Brullé, mentioned, 201.
| Wollastonia,
INDEX
| Volvulus cupreus, Régimbart, 200;
mentioned, 201.
denticulatus, Régimbart, men- |
tioned, 201.
stagnicola
mentioned, 194, 201.
Weber-van Bosse, <A., Marine
Alge, Rhodophycee, 269-306.
Wilsonea «indica, De Toni, men-
tioned, 297.
Horn
mentioned, 480.
(nee
Wollastoniella, Horn (nec Heer),
mentioned, 480.
Wormia, fottb., mentioned, 209,
408.
ferruginea, Baill., mentioned,
386, 407, 413, 415, 481,
487.
Xenorohestes, mentioned, 260.
Xenotropis, fairm., 248.
rugicollis, Havrm., 248 ; men-
tioned, 247.
Xerobdella,, v. Lrauenfeld,
tioned, 39, 40.
Xiphispa, Chap., mentioned, 237,
239.
Xyleborus, Hichh., 386.
sp., mentioned, 382.
abnormis, Eichh., 389.
men-
(Mulsant), 200 ;
Heer), |
517
Xyleborus abruptus, Sampson*,
388.
affinis, Hichh., 388; men-
tioned, 387.
amputatus, Blandf:, men-
tioned, 388.
andrewesi, Blandf:, 387.
badius, Kichh., 387.
bucco, Schauf., 388.
confusus, Hichh., 387.
dilatatus, Hichh., 387.
insularis, Sharp, 387.
kraatzi, Kichb., 387.
madagascariensis, Schawf., 386 ;
mentioned, 384.
amancus, Blandf:, mentioned,
388.
multispinosus, Haged., men-
tioned, 389.
perforans (Woll.), 387 ;
tioned, 388.
semigranosus, Blandf:, 386.
seaspinosus, Haged., 389.
torquatus, Hichh., 387.
men-
Yanga seychellensis, istant, men-
tioned, 190.
Zaprionus, Cog., 348.
vittiger, Cog., 348; mentioned,
370.
| Zygopina, 448.
CAMBRIDGE - PRINTED BY JOHN CLAY, M.A. AT THE UNIVERSITY PRESS
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2nd Ser. ZOOLOGY. | (VOL. XVI. PART 1.
THE
TRANSACTIONS
OF
THE LINNEAN SOCIETY OF LONDON.
THE PERCY SLADEN TRUST EXPEDITION
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UNDER THE LEADERSHIP OF
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ree
os ie
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mS ne £) su hdk 29. hh a5) Ba Gls
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a nr
ee wae) fs
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:
|
i
LINNEAN SOCIETY OF LONDON.
MEMORANDA CONCERNING TRANSACTIONS.
The First Series of the Transactions, containing both Botanical and Zoological contributions, has been
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When Price to the Price to When Price to the Price to
Volume Published Public Fellows Volume Published Public Fellows
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2nd Ser. ZOOLOGY. | [VOL. XVI. PART 3.
THE
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OF su LIBRARY)|>9
z
ea 7
THE LINNEAN SOCIETY OF LONDON.
THE PERCY SLADEN TRUST EXPEDITION
TO
THE INDIAN OCEAN IN 1905,
UNDER THE LEADERSHIP OF
Mr J. STANLEY GARDINER, M.A.
VOLUME V.
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If. Parts I. XVIII. 1879-88 ...7 17 0...
Jil. Parts I. VI. 1884-88...5 18 0...
IV. Parts I.-IIJ. 1886-88...3 8 0...
VY. Parts J.-XI. 1888-94...6 10 6...
VI. Parts [—VIII. 1894-99...6 17 6...
VII. Parts I.-XI. 1896-1900...6 7 9...
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Qnd Ser. ZOOLOGY.| - [VOL. XVI. PART 4.
THE
TRANSACTIONS
OF
THE LINNEAN SOCIETY OF LONDON.
?
THE PERCY SLADEN TRUST EXPEDITION
TO
THE INDIAN OCEAN IN 1905,
UNDER THE LEADERSHIP OF
Mr J. STANLEY GARDINER, M.A.
VOLUME V.
Reports Nos. 15—19 of this volume; Nos. 101—105 of the whole series.
LONDON:
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II. Parts I. XVIII. 1879-88 ...°7 1'7
Ill. Parts I. VI. 1884-88... 5 18
IV. Parts) T—i 1886=88 =. 3 8
V. Parts [.-XI. 1888-94... 6 10
VI. Parts I.—VIII. 1894-99... 6 17
VII. Parts 1.—XI. 1896-1900...6 7
VIII. Parts J.—XIII. 1900-03 ...6 14
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SECOND SERIES.—ZooLoey (continued).
When Price to the Price to
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Qnd Ser. ZOOLOGY. ] [VOL. XVI. PART 5.
THE
TRANSACTIONS
OF
THE LINNEAN SOCIETY OF LONDON.
TITLEPAGE, CONTENTS, AND INDEX.
LONDON:
PRINTED FOR THE LINNEAN SOCIETY
BY JOHN CLAY, M.A. AT THE UNIVERSITY PRESS, CAMBRIDGE.
SOLD AT THE SOCIETY’S APARTMENTS, BURLINGTON HOUSE, PICCADILLY, W.
AND BY LONGMANS, GREEN, AND CO., PATERNOSTER-ROW.
December, 1914.
LINNBAN SOCIE
TY OF LONDON.
| MEMORANDA CONCERNING TRAN SACTION 8.
‘Nhe First Series: of the ‘Transactions, containing both Botanical and Zoological contributions, has been
completed in 30 Vols., and a few entire sets are still for sale.
complete sets, may be obtamed at the original prices.
Only certain single volumes, or parts to
The price of the Index to Vols. 1-25 is 8s. to
the public, and 6s. to Fellows; to Vols. 26-30, 4s. to the public, and 3s. to Fellows.
The Second Series of the ‘Transactions is divided into Zoological and Botanical sections.
of the Zoological parts of those which have been published are as undermentioned :—
Second SERIES.—ZooLoey.
When Price to the
Volume Published Public
a Ga lo
1 Parts EVIL
Th Parts I. XVIII 1879-88)... 7.17
IM. Pants* I. Vi. 1884-88: ... 518
_ IV.” Parts* L=IM, 1886-88...53 8:
V. + Parts L-Xb 1888-94"... 6-10
VI. Parts IVIL. 1894-99... 6.17.
Parts [.—XI. 1896-1900 ...
VII. G7
VIII. Parts I.-XfII. 1900-08... 6 14
Xe) Parts Ie 1903 «.0 9
Part ID. 1903... 0 8
Pambpe Wl agease oe WOO! sera lee 4:
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Part, }V.: 1904 ...0 6
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Part VIIL, -... 1904 -'....0 10
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‘ Part XI. 1907 .... 0 12
Part, XER eo 2 OT. 0, 63
Party Xs ios HOOF: % 22-0: * 6
Part XIV. Index 1907... 0 3
XG arena |e 1904 ...,0 38
Part Tk 1904 ...0 8
Part III. 1905) 2650) 9
Part IY. 1905 ... 0 10
Part V._ 1906 ...0 7
Part VI. 1906... 0 3
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