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FIELDIANA: BOTANY

A Continuation of the

BOTANICAL SERIES

of

FIELD MUSEUM OF NATURAL HISTORY

VOLUME 31

The Library of the

JAN 1 8 1979

FIELD MUSEUM OF NATURAL HISTORY

CHICAGO, U. S. A.

£20.5

TABLE OF CONTENTS

PAGE

1. Two New Species of Palms from Nicaragua. By S. F. Glassman ... 1

2. Tropical American Plants, VI. By Louis 0. Williams 11

3. Agriculture, Tehuacan Valley. By C. Earle Smith, Jr 49

4. Flora, Tehuacan Valley. By C. Earle Smith, Jr 101

5. Preliminary Studies in the Palm Genus Syagrus Mart, and Its Allies.

By S. F. Glassman 145

6. Tropical American Plants, VII. By Louis 0. Williams 165

7. Supplement to Orchids of Guatemala. By Donovan S. Correll .... 175

8. Preliminary Notes on Scrophulariaceae of Peru. By Gabriel Edwin . . 223

9. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 233

10. Tropical American Plants, VIII. By Louis O. Williams 247

11. Notes on the Flora of Costa Rica, I. By William C. Burger 273

12. A New Eurystyles from Nicaragua. By Alfonso H. Heller 279

13. New Species in the Palm Genus Syagrus Mart. By S. F. Glassman . . 285

14. A Revision of the Family Geastraceae. By Patricio Ponce de Leon . . 303

15. Studies in American Plants. By Dorothy N. Gibson 353

16. Two New Nicaraguan Juglandaceae. By Antonio Molino R 357

17. Studies in the Palm Genus Syagrus Mart. By S. F. Glassman .... 363

18. Tropical American Plants, IX. By Louis O. Williams 401

-31

STUDIES IN AMERICAN PLANTS

DOROTHY N. GIBSON

TWO NEW NICARAGUAN JUGLANDACEAE

ANTONIO MOLINA R.

STUDIES IN THE PALM GENUS SYAGRUS MART.

S. F. GLASSMAN

TROPICAL AMERICAN PLANTS, IX

LOUIS 0. WILLIAMS

FIELDIANA: BOTANY

VOLUME 31 NUMBERS 15, 16, 17, 18

Published by

FIELD MUSEUM OF NATURAL HISTORY

MAY 23, 1968

Imcratf if Rfinfe

JUL1H968

LAI n ii«**>

TWO NEW NICARAGUAN

JUGLANDACEAE

ANTONIO MOLINA R.

Field Associate

FIELDIANA: BOTANY

VOLUME 31, NUMBER 16

Published by

FIELD MUSEUM OF NATURAL HISTORY

MAY 23, 1968

PUBLICATION 1046

Library of Congress Catalog Card Number: 68-26378

PRINTED IN THE UNITED STATES OF AMERICA

BY FIELD MUSEUM PRESS

Two New Nicaraguan Juglandaceae

ANTONIO MOLINA R.

In our continuing explorations and botanical investigations in

Nicaragua which we are able to carry on with assistance from the

National Science Foundation, we have collected thousands of num-

bers of plant specimens which increase the usefulness of the herbaria

in Zamorano and in Chicago, as well as herbaria which receive dupli-

cates in exchange. There have been recognized a considerable num-

ber of undescribed plants in these collections, as was to be expected,

for Nicaragua is the least known biologically of the Central American

countries.

We have seen two trees of the Juglandaceae in the Cordillera

Central de Nicaragua which we did not recognize, one an Engel-

hardtia the other an Alfaroa. We have accumulated adequate ma-

terials of both of these trees and both appear to be new.

Alfaroa williamsii A. Molina, sp. nov.

Arbor valida usque ad 25 m. et 0.5 m. diam. Folia subopposita vel alterna

composita, foliola 3-5 paribus anguste elliptica vel elliptico-oblanceolata acuta

vel acuminata coriacea basi cuneata; inflorescentia masculina paniculata erecta

multiflora; inflorescentia foem. spica pauciflora; flores masc. tepala 2 oblonga vel

obovata; flores foem. tepala 4 oblanceolata vel spathulata; fructus rotundatus

vel obovoideus.

Trees 15-25 m. tall, the trunks 0.3-0.5 m. in diameter, the bark smooth or

becoming rugose, easily separated from the trunk in short, thin sheets, greenish-

yellow or pale cream colored. Branches cylindrical, glabrous, commonly striated

or verrucose and with many brownish elliptical or rounded lenticels; leaves sub-

opposite or alternate, compound, mostly 15-25 cm. long, the leaflets 3-5 alternate

or subopposite pairs, coriaceous, narrowly elliptic or elliptic-oblanceolate, acute or

acuminate, obliquely cuneate to the base, with 14-20 pairs of secondary nerves,

glabrous except obscurely lepidote below, 4-13.5 cm. long and 1.5-3.5 cm. broad,

petioles 1.4-4.5 cm. long, rachis canaliculate, 3-13 cm. long, petiolules short,

1-2 mm. long; staminate inflorescence terminal or axillary-terminal, panicles of

several erect many-flowered catkins; pistillate inflorescence a terminal or subaxil-

lary few-flowered spike; staminate flowers many, sessile, lepidote-glandular, bract

trilobate, 3-4 mm. long, lateral lobes 0.5 mm. long, the central lobe 1 mm. long,

obtuse or acute, tepals 2, oblong or obovate, about 1 mm. long, stamens 7-9,

anthers two, sessile, oblong, 1-1.5 mm. long and about 1 mm. broad; pistillate

357

358 FIELDIANA: BOTANY, VOLUME 31

flowers sessile, 5-7 mm. long, the tepals 4, thick and fleshy, glabrous or usually

lepidote, oblanceolate or spathulate, 2-4 mm. long and 0.5-1.5 mm. broad, stigma

bifid, about 1-1.5 mm. long, the style about 2 mm. long, ovary 2 mm. long, sub-

tended by a trilobate bract 1-2 mm. long and with lobes 0.5-2 mm. long; fruit a

sessile, woody, hard, glabrous, 8-10 celled with as many as 13 septae, drupaceous

nut, rounded or shortly obovoid, 14-17 mm. long and 13-16 mm. in diameter, with

8-9 raised ribs, the tepals and style persistent at the apex.

NICARAGUA: tree 15-20 m. tall, in old cut-over cloud forest area

between Disparate de Potter and Aranjuez, Cordillera Central de

Nicaragua, Dept. Matagalpa, alt. 1,300 meters, Jan. 12, 1963, Wil-

liams, Molina and Williams 23717 (type, F; EAP) ; tree to 15 meters,

wide spreading, cut-over cloud forest area, Cordillera Central de

Nicaragua, road to Aranjuez, Dept. Jinotega, alt. 1,400 meters,

Jan. 8, 1967, Williams and Molina 20143; "Areno bianco," tree 20-

25 meters tall, 30-50 cm. in diameter, cut-over cloud forest area,

El Arenal, between Aranjuez and Santa Martha, Dept. Matagalpa,

alt. 1,400 meters, March 7, 1967, Molina 203 40.

There have been three species of Alfaroa previously described

from Central America from which A. williamsii is easily distinguished

by its small nearly round, glabrous and ribbed fruits. It is at the

same time the third member of the Juglandaceae to be found in

Nicaragua within recent years where none was previously known,—

Juglans olanchana Standl. & L. Wms. and the two species described

in this paper.

It is a pleasure to name this distinctive Nicaraguan tree for my

friend and field companion of many years, Louis O. Williams, who

has spent his life working with the flora of Central America.

Engelhardtia nicaraguensis A. Molina, sp. nov.

Arbor valida usque ad 25 m. vel ultra. Folia pinnata, usque ad 12 cm. longa;

foliola 4-6 pares, lamina coriacea, lanceolata, lanceolato-oblonga vel anguste

oblonga, acuminata vel acuta, basi uni- vel biauriculata; fructa obovata bractea

trilobata in racima elongata.

Large forest trees, 10-25 m. tall, the trunk cylindrical, 25-50 cm. in diameter

or perhaps more, bark yellowish or grayish-green; leaves pinnate, opposite, the

petioles 2.5-3.5 cm. long, glabrous or minutely lepidote, the rachis subterete or

obscurely canaliculate, 4-9 cm. long, slender; leaflets 4-6 pairs, opposite or sub-

opposite, short petiolulate, coriaceous, lanceolate, lanceolate-oblong to narrowly

oblong, 3-8.5 cm. long and 1.5-2.3 cm. broad, sometimes with slightly revolute

margins (in dry specimens), glabrous or lightly punctate-lepidote, somewhat

glaucous below, acuminate or acute, the base obtuse or more commonly attenuate

and oblique with a very small revolute auricle or usually biauriculate; central

vein prominent on both surfaces of the leaflet, secondary veins several to many

MOLINA: NEW NICARAGUAN JUGLANDACEAE 359

and less prominent; the lepidote petiolules 1-2 mm. long; female inflorescence

cymosely branched (one seen), about 15 cm. long, each branch with 5 or perhaps

more short pedunculate flowers; nuts obovoid or ovoid, 6-8 mm. long and 6-7 mm.

in diameter, subtended by a rigidly membranaceous, reticulately nerved, lepidote

trilobate bract, central lobe of bract oblong-linear or spathulate, about 2-3.8 cm.

long and 0.6-0.8 cm. broad, emarginate, the lateral lobes narrowly oblong, obtuse,

about 1.5 cm. long and 0.5-0.8 cm. broad, fruiting peduncles 0.2-0.5 cm. long.

NICARAGUA: Arbol 25 m., tallo 25-50 cm. diametro, bosque

nebuloso vecindades de Santa Maria de Ostuma, departamento de

Matagalpa, alt. 1,300 m., Mayo 1957, Juan B. Sales 2967 (F, type;

EAP) ; arbol 10-15 m. en bosque humedo cerca Aranjuez, departa-

mento de Jinotega, alt. 1,160 m., Agosto 1957, Juan B. Sales y B. J.

Taylor 2967a.

The lack of really good herbarium material makes it difficult to

establish clearly the relationship of this species, although it must

be closest to Engelhardtia mexicana Standl. The Mexican species

has leaves with 3-4 pairs of leaflets and a petiole 1.5-2 cm. long and

the rachis 2-6 cm. long, the petiolules of the leaflets 2-3 mm. long;

the leaflets in that are linear-lanceolate, 5-8.5 cm. long, with the

apex long attenuate, entire or sometimes with tooth-like undulations

near the apex, the auricles at the base obtusely deltoid or rounded ;

the lobes of the bracts are oblong or linear-oblong; the nut broadly

obovoid and 6-7 mm. long.

Publications 1045, 1046, 1047, and 1048

UNIVERSITY OF ILLINOIS-URBANA