CRYPTOGAMIE

Bryologie-Lichénologie

ANCIENNE REVUE BRYOLOGIQUE ET LICHENOLOGIQUE

Fondée par Т. Husnot en 1874

Directeur de la publication : Dr Héléne Bischler-Causse

Rédaction :

Bryophytes : Dr Héléne Bischler & M. Denis Lamy, Laboratoire de Cryptogamie, 12 rue

Buffon, F-75005 Paris. Tel. 01.40.79.31.84 — Fax : 01.40.79.35-94 Email : lamy

@ mnhn.fr

Lichens: Dr Chantal Van Haluwyn, Laboratoire de Botanique et de Cryptogamie, Faculté

de Pharmacie, В.Р. 83, F-59006 Lille Cedex. Tél 03.20.96.40.40 —

Fax : 03.20.95.90.09

Editeur : A.D.A.C. — 12 rue Buffon F-75005 Paris

COMITÉ DE LECTURE

Bryologie : J. Berthier (Clermont-Ferrand), B. Bodo (Paris), В.В. Buck (Bronx), J.L. De Sloover

(Namur), P. Geissler (Genéve), S.R. Gradstein (Utrecht), J.P. Hebrard (Marseille), S. Jovet-Ast

(Paris), A. Lecointe (Caen), М.С. Noailles (Paris), R. Ochyra (Kraków), C. Suire (Bordeaux),

B.C. Tan (Cambridge, USA).

Lichénologie : J, Asta (Grenoble), A. Bellemére (Paris), T. Bernard (Rennes), B. Bodo (Paris), J.C.

Boissière (Paris), P. Clerc (Genève), W.L. Culberson (Durham), 5. Déruelle (Paris), P. Diederich

(Luxembourg), M.C. Janex-Favre (Paris), J. Lambinon (Liege), M.A. Letrouit-Galinou (Paris), X.

Llimona (Barcelone), L. Nimis (Trieste), G. Rambold (Munich), Cl. Roux (Marseille), M.R.D.

Seaward (Bradford), A. Sérusiaux (Liege), H. Sipman (Berlin), E. Stocker-Wörgötter (Salzburg),

V. Wirth (Stuttgart).

MANUSCRITS

Les manuscrits doivent étre adressés (en 3 exemplaires) à la Rédaction de Cryptogamie,

Bryologie-Lichenologie. L'avis du Comité de Lecture sera requis avant accord pour publication.

Bien que Cryptogamie soit une revue de langue française, les manuscrits rédigés en français, en

anglais, en allemand, en espagnol et en italien, sont pris en considération dans la limite des

possibilités de la Rédaction à trouver des lecteurs compétents dans ces langues. Les disquettes de

micro-ordinateurs (IBM, IBM compatible et MacIntosh) sont vivement souhaitées. Les instruc-

tions aux auteurs sont publiées dans le fascicule 1 de chaque tome. Les auteurs recevront

25 tirés-à-part gratuits ; les exemplaires supplémentaires seront à leur charge.

TARIFS DES ABONNEMENTS Tome 18, 1997

CRYPTOGAMIE comprend trois sections : Algologie, Bryologie-Lichénologie, Mycologie.

Pour une section : France : (350 Е ht) 357,35 F ис Étranger: 380,00 Е

Pour les 3 sections : France : (950 Е ht) 970,00 Е ис Étranger : 1050,00 Е

VAT Registration number : FR29302655170

Paiement par chéque bancaire ou postal à l'ordre de : A.D.A.C. — CRYPTOGAMIE (CCP La

Source 34 764 05 S) ; adressé à : A.D.A.C. , 12 rue Buffon, F-75005 Paris. :

Cryptogamie, Bryologie-Lichénologie est indexé par Biological Abstracts, Current Contents,

Index of Fungi et Bibliography of Systematic Mycology, Publications bibliographiques du CNRS

(Pascal).

Couverture : Pore de Marchantia foliacea Mitt. (cliché H. Bischler)

Source : MNHN. Paris

CRYPTOGAMIE

BRYOLOGIE LICHENOLOGIE

TOME 18 FASCICULE 1 1997

CONTENTS

Harald KÜRSCHNER — An annotated, corrected, and updated list of the

bryological literature of Southwest Asia...

J. MARTÍNEZ-ABAIGAR, E. NÜNEZ-OLIVERA, A. GARCÍA-ÁLVARO

and N. BEAUCOURT — Additions to the bryophyte flora of la Rioja

and the Iberian System (Northern Spain) ........................... 47

Valeria SPAGNUOLO, Paolo CAPUTO, Salvatore COZZOLINO, Rosa

CASTALDO and Paolo DE LUCA — Length polymorphism in the

intragenic spacer І of the nuclear ribosomal DNA of some Pottiaceae

(Pottiales, Musci).. 55

Gregorio ARAGON & Isabel MARTINEZ — Contribution to the knowledge

of the Epiphytic Lichens of Toledo Mountains (Toledo, Spain)

(In Spanish) Ts 63

J. M. EGEA & P. TORRENTE — Three new species of lichenicolous fungi

from Cape province (South Africa) — Correction (In Spanish)...... 17

Book Reviews (in French) 79

Tastructions dux antes ee а Vr des 83

Instzuchaus to АН нь se... dmi dient ee Sin e on d ew a 85

U | Bibliothéque Centrale Muséum

Np 3 300190037910

3001, 000379,

00037511 BN Paris

Cryptogamie, Bryol. Lichenol. 1997, 18(1) : 1-46 1

AN ANNOTATED, CORRECTED, AND UPDATED LIST

OF THE BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA

Harald KURSCHNER

Institut fiir Systematische Botanik und Pflanzengeographie

der Freien Universitát Berlin, Altensteinstr. 6, D-14195 Berlin

ABSTRACT — An up-to-date list of 294 references on bryological literature of Southwest Asia

(Near and Middle East) is given. The references are annotated for the first time and provide

information on content, especially the new taxa published in the older literature, independant of

the bibliographic descriptions. In addition, a guide to national (Afghanistan, Bahrain, Iran, Iraq,

Israel, Jordan, Kuwait, Lebanon, Oman, Qatar, Saudi Arabia, Syria, Turkey, United Arab

Emirates, Yemen) and regional (e. g. Egypt/Sinai) literature is given which allows a quick

orientation and reflect the bryofloristic state of knowledge. Finally, an index of collectors of the

different countries with reference to author(s) and date is presented.

ZUSAMMENFASSUNG — Die Arbeit enthält eine auf dem neuesten Stand befindliche,

294 Publikationen umfassende Literaturübersicht über die bryologische Literatur Südwest-Asiens

(Naher — und Mittlerer Osten). Dabei werden die Arbeiten erstmals inhaltlich kurz kommentiert

und dabei vor allem die in der älteren Literatur neu beschriebenen Sippen aufgeführt. Zusätzlich

wird, zur schnelleren Orientierung, eine nach Ländern (Afghanistan, Bahrein, Iran, Irak, Israel,

Jordanien, Jemen, Katar, Kuweit, Libanon, Oman, Saudi Arabien, Syrien, Türkei, Vereinigte

Arabische Emirate) und Regionen (Ägypten/Sinai-Halbinsel) geordnete Übersicht gegeben, die

auch den bryofloristischen Kenntnisstand widerspiegelt. Den Abschluß bildet ein, nach Ländern

geordnetes Verzeichnis der Hauptsammler, mit Angabe der Quelle(n) in der ihre Aufsammlungen

veröffentlicht wurden.

INTRODUCTION

More than 15 years ago, a first compilation of bryological publications of

the Near and Middle East was published (Frey & Kürschner 1981), including

150 references. Aim of the study was, to give an overview of the state of bryofloristic

knowledge to that time, stimulating future investigations on the bryophyte flora and

vegetation. The area was considered again in the Conspectus of bryological taxonomic

literature, part 2 (guide to national and regional literature, Greene & Harrington 1989),

which provide a wide selection of up-to-date works and the “...тоге worthwhile of the

older publications...published in the last fifty years...” (Greene $: Harrington 1989), but

includes only literature of particular relevance to the identification of bryophytes.

Source > MNHN, Paris

2 H. KURSCHNER

No. of publications listed

1981 (cf. Frey & Kürschner

| 1981)

Actual no. of publications

Tab. 1 — Number of bryological publications of the Near and Middle East and the increase in

publications since the first bibliography (Frey & Kürschner 1981). * No records of bryophytes up

to now (cf. Frey & Kürschner 1991).

Publications on the bryophyte flora and vegetation of the area therefore are lacking.

Since the publication of the first reference list, intensive collecting activities, especially

on the Arabian Peninsula (Oman, Saudi Arabia, Yemen), in Jordan and in Turkey

took place, which led to an enormous increase of bryological publications (Tab. 1).

Various new country records were reported which complete our knowledge of the

distribution and occurrence of the different taxa. Beside these new country records

several new and endemic taxa has been described, indicating the evolutionary potential

of Southwest Asia. In addition to these floristic papers and updated checklists, studies

on the different bryophyte communities (e. g. terrestrial, epilithic and epiphytic

communties) of the area were published, which give an impression of the vegetation,

the community structures, the ecology and habitat requirements, the syntaxonomic and

synhierarchic affinities, the biodiversity, and the status and function of bryophytes in

the different ecosystems. Of special interest during the last five years were the

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 3

40%

30%

20°4

G SOUTHWEST

Г ASIA

f T

40* 50* d

Fig. 1 — Southwest Asia (Near and Middle East; Af Afghanistan, Ba Bahrain, Ir Iran, Iq Iraq,

Is Israel, Jo Jordan, Ku Kuwait, Le Lebanon, Om Oman, Qa Qatar, SA Saudi Arabia, Si

Egypt/Sinai Peninsula, So Yemen/Socotra, Sy Syria, Tu Turkey, UAE United Arab Emirates, Ye

Yemen).

morphological-anatomical adaptations and the life strategies (e. g. reproduction and

dispersal strategies) of xerophytic bryophytes which, beside some few exceptions, have

been more or less neglected in the older publications. All these publications, together

with the following corrected, updated and annotated list of references can be seen as

a step foreward to a more or less complete bryophyte flora of the Near and Middle

East, which leads to a better understanding of the distribution, phytogeographical,

ecological and evolutionary relationships of the different taxa and the biodiversity of

the area.

The up-to-date list of references now comprises 294 titles. It include also the

titles of the 1981 compilation which partly are corrected because of bibliographical

errors and now are annotated. The bibliography deal with bryological literature and

include floras, checklists, phytosociological, ecological and related works. Beside a

Source : MNHN. Paris

4 H. KURSCHNER

complete citation of each reference, arranged in alphabetical order of authors and then

chronologically following the first author, a short guide to national (Afghanistan,

Bahrain, Iran, Iraq, Israel, Jordan, Kuwait, Lebanon, Oman, Qatar, Saudi Arabia,

Syria, Turkey, United Arab Emirates, Yemen) and regional (e. g. Egypt/Sinai

Peninsula) literature is given, the data arranged on a geographical basis (Fig. 1). Where

an individual work is equally relevant to several countries or areas, it is cited under

each of these.

Finally, for the first time, an index of the main collectors of the different

countries is presented, with reference to author(s) and date, where the collections are

published. This attempt must be seen as preliminary, because not all collectors are

indicated in the different publications.

Inevitably some references will have been overlooked again. This may be true

especially for monographies and taxonomic revisions, which often include locality or

collection data of the Near and Middle East, but also for some few publications,

published in only local distributed journals. Nevertheless I hope, that the bibliography

is more or less complete and adequate to enable a user to locate quickly such titles as

seem appropriate to his or her requirements. Until a few exceptions the references are

annotated for the first time (the annotations are placed in square brackets [ ]) and

provide information on content (especially new described taxa), independent of the

bibliographic descriptions, in the hope, that they will assist the user to judge the

relevance of a particular work.

It is much to be hoped that the bibliography again act as a stimulus to the

investigation of the bryoflora and vegetation and the production of new publications

for those countries for which no or only a few adequate accounts exist.

LIST OF LITERATURE,

ARRANGED IN ALPHABETICAL ORDER OF AUTHORS

AGNEW S., 1973 A new Middle Eastern Grimmia. Journal of Bryology 7: 339-

342.

[Grimmia gibbosa from Iraq is described and illustrated].

AGNEW S. & TOWNSEND C. C., 1970 — Trichostomopsis Card., a moss genus new

to Asia. Israel Journal of Botany 19: 254-259.

[Discussion of the taxonomical status of Trichostomopsis; two species, Т. aaronis and

Т. haussknechtii are transferred].

AGNEW S. & VONDRACEK M., 1975 — A moss flora of Iraq. Feddes Repertorium

86: 341-489.

[Illustrated flora with keys to genera and species, synonymy, notes on habitats and

taxonomy, localities and distribution, based largely on the collections of S. Agnew and

E. Hadaé useful literature list].

AITCHISON J. E. T., 1881 — On the flora of the Kuram valley, Afghanistan. Journal

of the Linnean Society, Botany 18: 1-113.

[Musci on page 113; reports two species with collection data].

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 5

AL-GIFRI A. N. & KURSCHNER H., 1996 — First records of bryophytes from the

Hadramout and Abyan Governorate, Southern Yemen. Studies in Arabian

bryophytes 20. Nova Hedwigia 62: 137-146.

[List of 18 moss and 7 hepatic taxa with collection data based on the authors’

collections, including phytogeographical and ecological remarks and several distribu-

tion maps; first records of bryophytes from the mainland of southern Yemen (former

Peoples Democratic Republic of Yemen — PDRY)].

AL-GIFRI A. N., KURSCHNER H. & MIES B., 1995 — New records, additions and

a new species, Sematophyllum socotrense Buck (Sematophyllaceae, Musci) to

the bryophyte flora of Socotra (Yemen). Studies in Arabian bryophytes 19.

Nova Hedwigia 61: 467-480.

[Checklist of the bryophyte flora of Socotra based on the authors’ collections and the

literature, including one new species and nine taxa new to the flora].

ARNELL 5., 1957 — A new species of Riccia from Israel. Bulletin of the Research

Council of Israel, Section D, 6: 56-57.

[Description and illustration of R. palaestina and report of К. terracianoi, previously

known only from Abyssinia].

ARNELL S., 1963 — Liverworts (Hepaticae). In: STORMER P.: Iranian plants

collected by Per Wendelbo in 1959, Appendix p. 35. Arbok for Universitetet

i Bergen. Mat.-Naturv. Serie No. 11: 3-35.

[List of eight taxa with collection data].

ARZENI C. B., 1965 — Bryophytes of the cedar groves of Lebanon. The Bryologist

68: 109-113.

[List of 35 taxa with collection data and notes of world-wide distribution, based on the

author’s and C. George’s collections].

BALFOUR I. B., 1903 — The ferns, mosses and liverworts of Sokotra. Jn; FORBES

H. O. (ed.): The natural history of Sokotra and Abd-el Kuri. London,

Pp. 537-541.

[List of 16 taxa with collection data, based on the collections of the Balfour Expedition

(1. B. Balfour, Lt. Cockburn, A. Scott 1879-1880, B.C.S.); cf. also Mitten (1888)].

BARBEY C. & BARBEY W., 1882 — Herborisation au Levant. Egypte, Syrie et

Méditerranée. Lausanne, 183 p.

[List of 24 mosses (Musci pp. 170-171) and three hepatics (Hepaticae p. 171) with

collection data].

BARBEY W., 1890 — Lydie, Lycie, Carie 1842, 1883, 1887. Etudes Botaniques.

Lausanne, 82 p.

[Reports Prerogonium gracile collected by C. F. Forsyth-Major from Haghios Ilias

(Samsun Dag)].

BAUM B. & JOVET-AST S., 1962 — Riccia récoltés en Israel. Revue bryologique et

lichénologique 31: 103.

[List of eight species with collection data].

Source . MNHN, Paris

6 H. KURSCHNER

BAUMGARTNER J., 1939 — Musci. In: RECHINGER K. H.: Ergebnisse einer

botanischen Reise nach dem Iran, 1937, pp. 534-536. Annalen des Naturhis-

torischen Museums in Wien 50: 410-536.

[List of 33 taxa with collection data, based on the collections of K. H. Rechinger;

includes the description of Grimmia crassifolia var. cucullata var. nov.].

BECKER H., 1967 — Bryologische Studien in Steppengebieten und im Bereiche

angrenzender Waldzonen Anatoliens. Mitteilungen der Ostalpin-Dinarischen

Pflanzensoziologischen Arbeitsgemeinschaft 7: 47-48.

[Includes some more general ecological notes on mosses (Barbula spp., Syntrichia spp.)

from Inner Anatolia].

BILEWSKY F. & NACHMONY S., 1955 — A contribution to the bryophyte flora

of Palestine. Bulletin of the Research Council of Israel, Section D, 5: 47-

58.

[List of 131 mosses with collection data, 50 being new to the area, based on the

collections of the authors’, T, Kushnir and D. Zohary; including the descriptions of

Tortula israelis sp. nov., Ephemerum sessile var. kushniri var. nov. and Philonotis

laxissima var. gemmiclada var, nov.].

BILEWSKY F., 1959 — A further contribution to the bryophyte flora of Palestine.

Bulletin of the Research Council of Israel, Section D, 7: 56-64.

[List of 95 mosses with collection data, four being new to the area].

BILEWSKY F., 1963 — Introduction aux Muscinées d'Israel. Tel Aviv.

[Not seen].

BILEWSKY F., 1965 — Moss flora of Israel. Nova Hedwigia 9: 335-434.

[Illustrated flora with keys to genera and species, synonymy and distribution data;

plates separately].

BILEWSKY F., 1971 — Some recent bryological records for Israel. Revue bryologique

et lichénologique 37 (4): 963-965.

[List of 11 taxa with collection data].

BILEWSKY F., 1974 — Some notes on the distribution of mosses in Israel and

Palaestina. Revue bryologique et lichenologique 40: 245-261.

[Checklist with synonyms, distribution data and some ecological notes].

BILEWSKY F., 1977 — New records of mosses in Israel. Israel Journal of Botany 26:

[Adds five taxa to the known flora].

BISCHLER Н. & JOVET-AST S., 1975 — Récoltes d'Hépatiques de Jérusalem à

Nablus et 4 En Gedi. Revue bryologique et lichénologique 41: 17-26.

[Reports 16 species with collection data, details of localities and ecological notes],

BISCHLER Н., 1978 — Plagiochasma Lehm. et Lindenb. П. Les taxa européens et

africains. Revue bryologique et lichénologique 44 : 223-300.

[Illustrated revision of Plagiochasma with key to the species, notes on the synonymy,

description of the species and notes on the systematic affinities, ecology and

distribution; includes collection data from Afghanistan, Oman, Turkey and Yemen,

and distribution maps].

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 7

BISCHLER Н., 1979 — Plagiochasma Lehm. et Lindenb. Ш. Les taxa d'Asie et

d'Océanie. Journal of the Hattori Botanical Laboratory 45 : 25-79.

[Illustrated revision of Plagiochasma with key to the species, notes on the synonymy,

description of the species and notes on the systematic affinities, ecology and

distribution; includes collection data from Afghanistan, Iran, Oman, Turkey and

Yemen, and distribution maps].

BISCHLER H. & JOVET-AST S., 1986 — The hepatic flora of South-West Asia: a

survey. Proceedings of the Royal Society of Edinburgh 89 B: 229-241.

{General account of the phytogeographical relationships, distribution patterns of some

Marchantiales and habitat adaptations of the hepatic flora].

BISCHLER-CAUSSE H., 1989 — Marchantia L. The Asiatic and Oceanic taxa.

Bryophytorum Bibliotheca 38: 1-317.

[Taxonomic revision with keys to the species and subspecies, details of specimen,

illustrations, notes on synonymy, phylogeny, habitat and distribution; includes M.

papillata subsp. grossibarba from Afghanistan and country records for Afghanistan,

Iran, Israel, Lebanon, Syria, Turkey, and Yemen].

BIZOT М., 1942 — Contribution à la flore bryologique du Liban. Revue bryologique

et lichénologique 13 : 49-53.

[List of 67 moss and 13 hepatic taxa with collection data, including the description of

Grimmia crinita var. libani var. nov. Hepaticae identified by Ch. Meylan].

BIZOT M., 1945 — Quelques mousses de Palestine. Revue bryologique et lichénologique

15 : 68-69.

[List Of 21 mosses (Israel, Lebanon) with collection data].

BIZOT M., GAUME R. & POTIER DE LA VARDE R., 1952 — Une poignée de

Mousses libanaises. Revue bryologique et lichénologique 21 : 11-13.

[List of 32 mosses from Lebanon (collected by M. Gombault) and Syria (collected by

M. Pallary) with collection data; includes the description of Cinclidotus pachylomoides

sp. nov. and Pogonatum aloides var. gombaultii var. nov.].

BIZOT M., 1954 — Remarques sur Tortula papillosissima (Copp.) Broth. Revue

bryologique et lichénologique 23 : 268-270 (publ. Jan. 1955).

[Reports 7. papillosissima from Syria, including notes on the systematic affinities,

morphology and ecology].

BIZOT M., 1955 — Contribution à la flore bryologique d'Asie mineure et de l'Ile de

Chypre. Revue bryologique et lichénologique 24 : 69-72.

[List of 72 mosses (Israel, Lebanon, Syria) with collection data and some remarks to

floristic affinities].

BIZOT M., 1956 — Nouvelles remarques sur Tortula papillosissima (Copp.)

Broth. Revue bryologique et lichenologique 25 : 268-271 (publ. March

1957).

[Note on the systematic affinities and related taxa of T. papillosissima ].

BIZOT M., 1962 — Quelques Mousses d’Afghanistan. Revue bryologique et lichéno-

logique 31 : 90.

[List of 11 mosses with collection data, based on the collections of C. v. Regel (1958,

1960)].

Source : MNHN, Paris

8 H. KÜRSCHNER

BOISSIER E. & BUHSE F., 1860 — Aufzaehlung der auf einer Reise durch

Transkaukasien und Persien gesammelten Pflanzen. Nouveaux Memoires de la

Société Impériale des Naturalistes de Moscou 12: 1-264.

[Musci on pp. 234-237, Hepaticae on pp. 237-238; list of 43 moss and six hepatic taxa

with collection data (identified by W. Schimper)].

BORNMÚLLER J., 1908a — Florulae Lydiae. Mitteilungen des Thüringischen

botanischen Vereins, Neue Folge 24: 1-140.

[Hepaticae and Musci frondosi on pp. 136-137; list of 15 moss and nine hepatic taxa

(identified by V. Schiffner) with collection data; includes Antitrichia breidleriana sp.

nov. (nom. nud.) and Reboulia hemisphaerica var. microspora var. nov. (nom. nud.); for

description and validation cf. Schiffner (1908)].

BORNMULLER J., 1908b — Beitráge zur Flora des Elbursgebirge Nord-Persiens.

Bulletin de l'Herbier Boissier, série 2, 8 : 109-124, 545-560, 723-738, 821-836,

915-930.

[Hepaticae and Musci on pp. 833-836 and 915-917; list of seven hepatic and 67 moss

taxa (identified by V. Schiffner) with collection data (cf. also Schiffner 1897, 1908)].

BORNMULLER J., 1909 Ergebnisse einer im Juni des Jahres 1899 nach dem

Sultan-dahg in Phrygien unternommenen Reise nebst einigen anderen Beitra-

gen zur Kenntnis der Flora dieser Landschaft Inner-Anatoliens. Beihefte zum

botanischen Centralblatt 24, Abteilung 2: 440-503.

[Hepaticae and Musci оп р. 499; list of one hepatic and seven moss taxa (identified by

V. Schiffner) with collection data (cf. also Schiffner 1901)].

BORNMÜLLER J., 1910 — Bearbeitung der von J. A. Knapp im nordwestlichen

Persien gesammelten Pflanzen. Verhandlungen der zoologisch-botanischen

Gesellschaft in Wien 60: 61-194.

[Musci on pp. 193-194; list of nine mosses (identified by У. Schiffner) with collection

data (cf. also Schiffner 1908)].

BORNMULLER J., 1911 — Collectiones Straussianae novae. Weitere Beiträge zur

Kenntnis der Flora West-Persiens. Teil 3. Beihefte zum botanischen Central-

blatt 28, Abteilung 2: 458-533.

[Hepaticae on p. 522, Musci on p. 523; list of three hepatic and eight moss taxa with

collection data (cf. also Schiffner 1908)].

BORNMULLER J., 1914 — Zur Flora des Libanon und Antilibanon. Beihefte zum

botanischen Centralblatt 31, Abteilung 2: 177-280.

[Musci (incl. Hepaticae) on р. 280a; list of two hepatic and four moss taxa with

collection data identified by V. Schiffner].

BORNMULLER J., 1915 — Plantae Brunsianae. Aufzählung der yon F. Bruns im

nördlichen Persien gesammelten Pflanzen. Beihefte zum botanischen Central-

blatt 33, Abteilung 2: 270-324.

[Musci and Hepaticae on p. 324; reports Marchantia polymorpha and Leskea laxiramea

with collection data].

BORNMÜLLER J., 1931 — Zur Bryophyten-Flora Kleinasiens. Magyar Botanikai

Lapok 30: 1-21.

[Checklist of 123 moss and 22 hepatic taxa with collection data based on the author’s

collection and older data].

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 9

BREMER B., 1980 — A taxonomic revision of Schistidium (Grimmiaceae, Bryophyta).

Lindbergia 6: 89-117.

[Revision with key to the species, details of specimen, notes on taxonomy, habitat and

distribution, including collection data from the Near and Middle East].

BROTHERUS V. F., 1906 — Lieutenant Olufsen’s second Pamir expedition. Plants

collected in Asia-Media and Persia, Musci. Botanisk Tidsskrift 27: 203-

208.

{List of mosses with three records from Iran, including collection data].

BROWN М. 5., 1937 — Mosses from Syria. The Bryologist 40: 84-85.

[List of two hepatic and 42 moss taxa based on the collection of W. B. Evans

1930-1931; exact locality data are lacking].

BRUGGEMAN-NANNENGA M. A., 1987 — An annotated list of Fissidens species

from the Yemen Arab Republic and Sultanate of Oman, with F. laxetextu-

ratus nov. spec. Studies in Arabian bryophytes 7. Nova Hedwigia 45: 113-

117.

[List of seven species with collection data, illustration of the new species, taxonomic and

habitat notes; includes additional remarks on Fissidens arabicus (cf. Pursell &

Kirschner 1987)].

BRULLO 5., PRIVITERA М. & PUGLISI M., 1991 — Note sulla flora е vegetazione

briofitica di alcune aree desertiche di Israele. Candollea 46: 145-153.

[Phytosociological study, reporting 21 moss and one hepatic taxa with collection data

and ecological notes; includes the description of two new associations and one new

subassociation].

CANO М. 1., GUERRA J. « ROS В. M., 1993 — A revision of Ше moss genus

Crossidium (Pottiaceae) with the description of the new genus Microcrossi-

dium. Plant Systematics and Evolution 188: 213-235.

[A world-wide revision with key to the species, details of specimens, illustrations, notes

on synonymy, phylogeny, habitat and distribution].

CETIN B. & YURDAKULOL E., 1985 — The moss (Musci) flora of Gerede-Aktas

(Bolu) forest. Doga Bilim Dergisi Seri A 2, 9: 29-39. In Turkish, with English

summary.

[List of 133 species with collection data based on the authors’ collections, 38 being new

for square A 2].

GETIN B. & YURDAKULOL E., 1986 — The liverworts (Hepaticae) of Bolu district

(Gerede-Aktas forest — Yedigöller National Park). Doga, Türk Biyoloji

Dergisi Seri C 10: 53-56. In Turkish, with English summary.

[List of 12 species with collection data based on the authors’ collections, two being new

to Turkey].

GETIN B. & YURDAKULOL E., 1988 — The moss flora of Yedigöller National

Park. Doga, Türk Botanik Dergisi Seri C 12: 128-146. In Turkish, with

English summary.

[Key to 88 mosses from the area based on collections of the authors’, 12 being new for

square A 2 and one, Fissidens monguillonii, new to Turkey].

Source : MNHN, Paris

10 H. KURSCHNER

GETIN B., 1988a — Checklist of mosses of Turkey. Lindbergia 14: 15-23.

[Checklist of 654 taxa (73 new to Turkey) based on the literature and an examination

of herbarium specimens; includes a useful literature list].

CETIN B., 1988b — Checklist of the liverworts and hornworts of Turkey. Lindbergia

14: 12-14,

[Checklist of 145 taxa based on the literature and an examination of herbarium

specimens; includes a useful literature list].

CETIN B., 1988с — Cinclidotus nyholmiae, a new species from Köprüllü Canyon

National Park (Antalya) in Turkey. Journal of Bryology 15: 269-273.

[Includes description, illustrations and collection data of the new species].

CETIN B., 1988d — Some mosses (Musci) of Dilek Peninsula National Park. Doga,

Türk Botanik Dergisi Seri C 12: 207-214. In Turkish, with English summary.

[List of 29 mosses with collection data, based on the author’s collections; includes two

illustrated new records (Fissidens adianthoides, Encalypta rhabdocarpa var. leptodon )

for Turkey].

CETIN B., 1989a — A new moss (Musci) record for Turkey. Doga, Tiirk Botanik

Dergisi Seri C 13: 139-142. In Turkish, with English summary.

[Reports Cinclidotus fontinaloides from Antalya area (square С 12), together with

collection data and an illustration].

CETIN B., 1989b — A new moss (Musci) record for Turkey. Doga, Tiirk Botanik

Dergisi Seri C 13: 143-146. In Turkish, with English summary.

[Reports Plagiothecium cavifolium from Bolu area (square A 2) together with collection

data and an illustration].

CETIN B., 1989c — A new moss (Musci) record for Turkey. Fontinalis antipyretica ssp.

antipyretica var. gracilis (Lindb.) Schimp. Doga, Tiirk Botanik Dergisi Seri C

13: 452-455, In Turkish, with English summary.

[Includes collection data and an illustration].

CETIN B., 1989d — Pohlia wahlenbergii (Web. & Mohr) Andrews var. calcarea

(Warnst.) Warburg, a new moss (Musci) record for Turkey. Doga, Tiirk

Botanik Dergisi Seri C 13: 147-150. In Turkish, with English summary.

[Includes collection data and an illustration].

CETIN B., 1989e — Porella pinnata L., a new liverwort (Hepaticae) recorded from

Turkey. Doga, Tiirk Botanik Dergisi Seri C 13: 134-138. In Turkish, with

English summary.

[Includes collection data and an illustration].

ÇETIN B., 1989f — Some mosses (Musci) of Antalya district [Kôprüllü Canyon

National Park, Güllükdagi (Termessos) National Park and Kursunlu water-

falls]. Doga, Türk Botanik Dergisi Seri C 13: 456-469. In Turkish, with

English summary.

[Key to 90 mosses from Antalya area, 55 being new to square C 12].

GETIN B., 1989g — The liverworts (Hepaticae) of Antalya district (Kursunlu

waterfalls — Köprüllü Canyon National Park). Doga, Türk Botanik Dergisi

Seri C 13: 151-156. In Turkish, with English summary.

[List of 15 species with collection data based on the author’s collection, 10 being new

to square С 12, and one, Marchantia paleacea new to Turkey].

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 11

ÇETIN B., 1990 — A new record for the moss flora of Turkey. Turkish Journal of

Botany 14: 160-162,

[Reports Fissidens serrulatus from Artvin district (square A 4) together with collection

data].

GETIN B., 1991 — Hygrohypnum duriusculum (De Not.) Jamieson: a new record for

the moss flora of Turkey. Lindbergia 17: 3-4.

[Includes collection data and ecological notes].

GETIN B., 1993 — An investigation of the Köygegiz-Dalyan specially protected area

as regards to bryophyte flora. Turkish Journal of Botany 17: 255-261.

List of 73 species (65 moss and eight hepatic taxa) from the area, with keys to genera

and species, and notes on habitat).

CHAROUK S., 1982 — Contribution à l'étude bryologique du Liban. Mousses et

Hepatiques de la vallé de Nahr Ibrahim, — These Doct. 3° cycle, Marseille,

Univ. Aix-Marseille III (Centre St. Jeröme), 84 p.

List of 65 moss, 20 hepatic and one anthocerote taxa based on the author’s collections,

with collection data, notes on the habitat, ecology and world-wide distribution].

CROSBY M. R., 1979 — The mosses of Israel: a field report. Bulletin of the Missouri

Botanical Garden 67: 1-3.

Report of a research project, adding three species (Antitrichia breidleriana, Gigasper-

mum mouretii, Pyramidula tetragona) to the moss flora of Israel].

CROSBY M. R., SCHULTZE-MOTEL U. & SCHULTZE-MOTEL W., 1983 —

Katalog der Laubmoose yon Madagaskar und den umliegenden Inseln.

Willdenowia 13: 187-255.

Systematically arranged checklist of all mosses, based on the literature, with synonyms

and note on the distribution; including Socotra island (Yemen) and an extensive

literature list].

CRUNDWELL A. C. & NYHOLM E., 1977 — Dicranella howei Ren. & Card. and

its relationships to D. varia (Hedw.) Schimp. Lindbergia 4: 35-38.

[Treatment of the systematic affinities of the two taxa, including collection data from

Afghanistan, Iran, and Turkey].

CRUNDWELL A. C., DURING H. J. & KOUTSTAAL B. P., 1978 — Tortula

grandiretis Broth. in the Netherlands and in Turkey. Lindbergia 4: 255-257.

[Description of the species with notes on the floristic affinities and ecology, including

one find from Turkey (SW Turkey, Egridir area)].

CRUNDWELL A. C. & NYHOLM E., 1979 — Some additions to the bryophyte flora

of Turkey. I. Hepaticae. Journal of Bryology 10: 479-489.

[List of one anthocerote and 60 hepatic taxa with collection data, based on the authors”

collections; includes 20 hepatics new to the bryoflora of Turkey].

CZECZOTT H., 1939 — A contribution to the knowledge of the flora and vegetation

of Turkey. Feddes Repertorium, Beiheft 107: 1-281.

[Musci on pp. 267-271; list of 22 mosses with collection data (mainly identified by J.

Thériot), four being new to Turkey].

Source : MNHN Paris

12 H. KURSCHNER

DANIN A. & BARBOUR M. G., 1982 Microsuccession of cryptogams and

phanerogams in the Dead Sea area, Israel. Flora 172: 173-179.

[Description of a cyclic microsuccession of blue-green algae, lichens and mosses with

ecological data; includes four mosses from the Jericho — Almoq area].

DECAISNE J., 1834 — Florula Sinaica. Enumération des plantes recueilles раг М.

Bové dans les deux Arabies, la Palestine, la Syrie et l'Egypte. Annales des

Sciences Naturelles, Г“ série, 2 : 5-18 (to be continued), Paris.

[Musci on pp. 10-11; lists five moss taxa from Syría and Sinai].

DELGADILLO C. M., 1973 — A new species, nomenclatural changes and generic

limits in Aloina, Aloinella and Crossidium (Musci). The Bryologist 76: 271-

277.

[Circumscription of the moss genera Aloina, Aloinella, and Crossidium with locality data

from Jordan].

DELGADILLO C. M., 1975 — Taxonomic revision of Aloina, Aloinella and

Crossidium (Musci). The Bryologist 78: 245-303.

[World-wide treatment of Aloina, Aloinella and Crossidium with key to the species,

descriptions and illustrations, notes on taxonomy, phylogeny, habitat and distribution:

includes locality data from Israel, Jordan, Lebanon, Sinai, Syria and Turkey].

DELGADILLO C. M., 1982 — Pseudaloina (Pottiaceae, Musci), a new genus from

Yemen. The Bryologist 85: 401-404.

[Description of Pseudaloina woodii gen. nov. and sp. nov. with illustrations and notes

on the phylogenetic relationships].

DINOLA L., MAYER A. M. & HEYN C. C., 1983 — Respiration, photosynthesis

and drought tolerance in mosses from various habitats in Israel. Israel Journal

of Botany 32: 189-202.

Physiological study of six mosses with notes on ecology and adaptations to

mediterranean and desert habitats].

DULL R., 1982 — Der Stand der Erforschung der Laubmoosflora im temperaten

Europa und benachbarten Asien. Beihefte zur Nova Hedwiga 71: 351-367.

|General account on the floristic knowledge; includes an extensive literature list].

EL-OQLAH A. А. & LAHHAM J. N., 1985 — Mosses from northern part of Jordan.

Candollea 40: 147-152.

[List of 25 moss taxa with collection data based on the authors’ collections].

EL-OQLAH A. A., FREY W. & KURSCHNER H., 1988a — The bryophyte flora of

Trans-Jordan. A catalogue of species and floristic elements. Willdenowia 18:

253-279.

[Catalogue of 12 hepatic and 110 moss taxa based on the literature and the authors’

collections, with synonyms, specimen data, details of habitats and distribution, and a

discussion of phytogeography and floristic elements; comprehensive literature list].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 13

EL-OQLAH A. А., FREY W. & KÜRSCHNER Н., 1988b — Tortula rigescens Broth.

et Geh. (Pottiaceae), a remarkable species new to the moss flora of Jordan.

Lindbergia 14: 27-29.

[First record since type (Kneucker 1902, Sinai); with illustrations and notes on floristic

relationships].

EL-SAADAWI W. E., 1976 — Some mosses from Kuwait. The Bryologist 79: 515-

518.

[List of 15 moss taxa with collection data, notes on habitat and ecology, based on the

author's collections; first records of mosses from Kuwait].

EL-SAADAWI W. E., 1978 — Observations on drought resistance in some Kuwaiti

desert mosses. Proceedings of the Egyptian Academy of Science 31: 265-

270.

[Reports 13 mosses with notes on ecology, habitat sites and adaptations to desert

conditions (e.g. rhizoid-borne gemmae)].

EL-SAADAWI W. E., 19792 — Contribution toward a moss flora of Kuwait. Journal

of the University of Kuwait 6: 125-152.

[Reports 19 species with descriptions, notes on habitats, specimens and world-wide

distribution; includes a key to 11 of the Pottiaceae recorded].

EL-SAADAWI W. E., 1979b — Peculiar aerial rhizoides in some mosses from Kuwait.

Journal of Bryology 10: 575.

[Reports four mosses which develop upward growing, branched, brown rhizoids from

the axils of the upper leaves}.

FAKHR М. S, 1977 — Contribution à l'étude de la flore Rhéto-Liasique de la

formation de Shemshak de l'Elbourz (Iran). Secr. Etat Univ., CTHS,

Mémoires de la Section des Sciences Paris 5 : 1-284.

[Includes geology, description of Bryophyta, Pteridophyta, and conifers, and the

definition of two floristic zones. Ricciopsis iranica sp. nov. is described. Published also

1975 as These Doct. Sci. Nat. Univ. P. & M. Curie, 421p.].

FEHLNER C., 1883 — Bryum elwendicum n. sp. Verhandlungen der zoologisch-

botanischen Gesellschaft in Wien 33: 435-436.

[Description of the new species with notes to the floristic affinities].

FEHLNER C., 1885 — Hepaticae, Musci. In: STAPF O.: Die botanischen Ergebnisse

der Polak’schen Expedition nach Persien im Jahre 1882, pp. 4-6. Denkschrif-

ten der Kaiserlichen Akademie der Wissenschaften Wien, Mathematisch-

naturwissenschaftliche Klasse 50, Abteilung 2: 1-120.

[List of one hepatic and eight moss taxa with collection data].

FORSSKAL P., 1775 — Flora aegyptiaco-arabica. Sive descriptiones plantarum, quas

per Aegyptum inferiorem et Arabium felicem detexit, illustravit Petrus

Forsskal. Post mortem auctoris edidit Carsten Niehbuhr. Accedit tabula

Arabiae felicis geographico-botanica. — Copenhagen, cxxvi, 219 p.

[Mentioned on p. 125 as no. 652 Mnium, as no. 653 Bryum, both from Yemen, but

without locality data].

Source : MNHN. Paris

14 H. KURSCHNER

FREY W., 1969 — Beiträge zur Moosflora Afghanistans I. Bartramiaceae. Nova

Hedwigia 17; 351-357.

[List with collection and distribution data; includes a key to the species and a brief

review of earlier work].

FREY W., 1972 — Beitrage zur Moosflora Afghanistans II. Die pleurokarpen

Laubmoose. The Bryologist 75: 125-135.

Checklist of 41 taxa with distribution data, a key to the taxa, and notes of the

bryogeographical affinities}.

FREY W. & PROBST W., 1973 — Die Polstermoosvegetation im Karaj-Tal

(Elbursgebirge, Iran). Botanische Jahrbücher für Systematik, Pflanzengeschi-

chte und Pflanzengeographie 93: 404-423.

Study of the epilithic bryophyte communities; includes a list of species, a statistical

analysis (coverage values) and ecological data].

FREY W., 1974 — Die Lebermoose des Iran und Afghanistans. The Bryologist 77:

48-56.

¡Checklist of 38 taxa (32 recorded from Iran and 16 from Afghanistan) with

distribution data and notes of the bryogeographical affinities).

FREY W. & PROBST W., 1974a — Vegetationsanalytische Untersuchungen im

Diinengebiet bei Babolsar. Botanische Jahrbiicher fiir Systematik, Pflanzen-

geschichte und Pflanzengeographie 94: 96-113.

Study of the dune vegetation near the Caspian coast (association analysis); includes

four mosses (Bryum bicolor, Pleurochaete squarrosa, Tortella humilis, Tortula ruralis)

with collection data].

FREY W. & PROBST W., 1974b — Hängeformen von Pseudoleskeella laxiramea

(Schiffn.) Broth. und Leucodon immersus Lindb. (Musci) im siidkaspischen

Waldgebiet (Iran). Ein Beitrag zur Morphologie und zu den Lebensbedin-

gungen. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflan-

zengeographie 94: 267-282.

[Morphological study, showing the relationships between epiphytic pendant-forms of

two Caspian endemics and the ecological conditions].

FREY W. & KURSCHNER H., 1977 Studies on the bryophyte flora and

vegetation of the Mohammad Reza Shah National Park, N. Iran. /ranian

Journal of Botany 1: 137-153.

[Lists 13 hepatic and 78 moss taxa, two hepatics and 21 mosses being new to Iran, based

on the authors’ collections; includes collection data and notes on the ecology and

habitat sites].

FREY W. & KURSCHNER H., 1979 — Die epiphytische Moosvegetation im

hyrkanischen Waldgebiet (Nordiran). Beihefte zum Tiibinger Atlas des

Vorderen Orients, Reihe A, Naturwissenschaften, Nr. 5, 99 S.

[Study of the epiphytic hepatic and moss flora of the Caspian lowland and montane

forests, including a list of species, a statistical analysis (community structures, coverage

values, frequence and dominance values, Fisher-test), phytogeographical remarks and

distribution maps].

FREY W. & KURSCHNER H., 1981 — The bryological literature of southwest Asia.

Journal of the Hattori Botanical Laboratory 50: 217-229.

[Reviews the state of bryofloristic knowledge of the Near and Middle East; extensive

literature list].

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 15

FREY W. & KURSCHNER H., 1982 — The first records of bryophytes from Saudi

Arabia. Studies in Arabian bryophytes 1. Lindbergia 8: 157-160.

[Reports six mosses and four thallose hepatics based on the authors’ collections, with

specimen and distribution data and a brief history of the bryological exploration].

FREY W. & KURSCHNER H., 1983a — Contributions towards a bryophyte flora of

Iran, — New records from Iran. /ranian Journal of Botany 2: 13-19.

[Lists 23 mosses and 17 liverworts new to Iran with specimen data].

FREY W. & KURSCHNER H., 1983b — New records of bryophytes from

Transjordan with remarks on phytogeography and endemism in SW Asiatic

mosses. Lindbergia 9: 121-132.

[List of 62 moss and five hepatic taxa with collection data and notes on the world-wide

distribution, 43 taxa being new to the flora; includes phytogeographical remarks on

endemism in the moss flora of Southwest Asia and the description of the circum-

Tethyan element].

FREY W. & KURSCHNER H., 1984 — Studies in Arabian bryophytes 3. Crossidium

asirense (Pottiaceae), a new species from Asir Mountains (Saudi Arabia).

Journal of Bryology 13: 25-31.

[One new species described and illustrated, including notes on floristic and phylogenetic

affinities].

FREY W., 1986 — Bryophyte flora and vegetation of South-West Asia. Proceedings

of the Royal Society of Edinburgh 89 B : 217-227.

{General account on the state of floristic knowledge and the phytogeographical

affinities (floristic elements and endemics) of the moss flora].

FREY W. & KURSCHNER H., 1987 — A desert bryophyte synusia from the Jabal

Tuwayq mountain systems (Central Saudi Arabia) with the description of two

new Crossidium species (Pottiaceae). Studies in Arabian bryophytes 8. Nova

Hedwigia 45: 119-136.

Reports, for the first time, a desert bryophyte synusium (community structure, ecology,

distribution); includes the descriptions and illustrations of Crossidium laxefilamentosum

sp. nov. and C. desertorum sp. nov. (= C. deserti nom. nov. pro C. desertorum, cf. Frey

& Kürschner 1988a)].

FREY W. & KÜRSCHNER H., 1988a — Bryophytes of the Arabian Peninsula and

Socotra. Floristics, phytogeography and definition of the Xerothermic Pan-

gaean element. Studies in Arabian bryophytes 12. Nova Hedwigia 46: 37-120.

Includes a catalogue of the bryophytes (one anthocerote, 44 hepatic and 143 moss

taxa) with synonyms, details of specimens, habitats and distribution, and distribution

maps of many taxa, based largely on recent collections; extensive literature list].

FREY W. & KÜRSCHNER H., 1988b — Pseudocrossidium replicatum (Tayl.) Zander

replaces Barbula acutata C. Müll. A note to its synonymy, distribution and

the xeropottioid life syndrome. Cryptogamie, Bryologie-Lichénologie 9: 95-102.

[Taxonomic treatment of P. replicatum from Saudi Arabia with notes on ecology,

distribution and its adaptations to desert habitats].

FREY W. & KÜRSCHNER H., 1988c — Re-evaluation of Crossidium geheebii

(Broth.) Broth. (Pottiaceae) from Sinai, a xerothermic Pangaean element.

Journal of Bryology 15: 123-126.

[Adds С. geheebii (reduced to synonymy with C. aberrans by Delgadillo 1975) again to

the flora of Sinai].

Source : MNHN. Paris

16 H. KURSCHNER

FREY W. & KURSCHNER H., 1988d — The Arabian Peninsula — An “Empty

Quarter” in Bryology? Bryological Times 45: 1, 3.

[Reviews the present state of bryofloristic knowledge; valuable literature list].

FREY W., 1989 — The bryophytes of Saudi Arabia and their importance of their

conservation. Studies in Arabian bryophytes 9. In: ABU-ZINADA A. H.,

GORIUP P. D. & NADER I. A. (eds.): Wildlife conservation and develop-

ment in Saudi Arabia, pp. 209-219. Riyad, National Commission for Wildlife

Conservation and Development, Publication No. 3.

[Summarizes the bryofloristical and bryovegetational state of knowledge].

FREY W., HERRNSTADT I. & KURSCHNER H., 1990a — Pterygoneurum

crossidioides (Pottiaceae, Musci), a new species to the desert flora of the Dead

Sea area. Nova Hedwigia 50: 239-244.

[One new species described and illustrated, including notes on the ecology and floristic

affinities].

FREY W., HERRNSTADT I. & KURSCHNER H., 1990b — Verbreitung und

Soziologie terrestrischer Bryophytengesellschaften in der Judäischen Wüste.

Phytocoenologia 19: 233-265.

[Phytosociological study (description of seven associations, three subassociations and

one community) with detailed remarks on syntaxonomy, community structures,

ecological conditions, habitat sites and distribution of the communities].

FREY W. & KURSCHNER H., 1990 — Das Fossombronio-Gigaspermetum mouretii

ass. nov. in der Judäischen Wüste. I. Verbreitung und Soziologie. Cryptoga-

mic Botany 2: 56-63.

[Phytosociological study with remarks on syntaxonomy, community structure, ecolo-

gical conditions, habitat sites and distribution].

FREY W., GROLLE R. & KÜRSCHNER Н., 1991 — Taxonomie und Verbreitung

von Asterella P. Beauv. (Hepaticae, Marchantiidae) im Vorderen Orient.

Studien an arabischen Bryophyten 16. Nova Hedwigia 52: 305-318.

[Floristic treatment of Asterella with key to the three species, detailed descriptions,

illustrations and notes on taxonomy and distribution; 4. elegans, previously reported

from Turkey is deleted from the flora].

FREY W. & KURSCHNER Н., 199la — Crossidium laevipilum Ther. et Trab.

(Pottiaceae, Musci), ein eigenständiges, morphologisch und standortökolo-

gisch deutlich unterscheidbares Taxon der Saharo-arabischen Florenregion.

Cryptogamie, Bryologie-Lichenologie 12: 441-450.

[Verifies the species rank of C. laevipilum (reduced to a variety of C. crassinerve

by Delgadillo 1975); includes detailed descriptions, illustrations and distributional

notes].

FREY W. & KÜRSCHNER H., 1991b — Conspectus Bryophytorum Orientalum et

Arabicorum. An annotated catalogue of the bryophytes of Southwest Asia.

Bryophytorum Bibliotheca 39: 1-181.

[First attempt of a complete checklist of the bryophytes from the Near and Middle East

(Afghanistan, Bahrain, Iran, Iraq, Israel, Jordan, Kuwait, Lebanon, Oman, Qatar,

Saudi Arabia, Syria, Turkey, United Arab Emirates, Yemen, including Cyprus and

Sinai) with notes on taxonomy; four anthocerote, 210 hepatic and 922 moss taxa are

reported; extensive literature list].

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 17

FREY W. & KÜRSCHNER H., 1991c — Das Fossombronio-Gigaspermetum mouretii

in der Judäischen Wüste. 2. Okosoziologie und Lebensstrategien. Cryptoga-

mic Botany 2: 73-84.

[Reports two new life-strategies in desert mosses (geophytes, “innovative” colonists)].

FREY W. & KURSCHNER H., 19914 — Lebensstrategien von terrestrischen

Bryophyten in der Judäischen Wüste. Botanica Acta 104: 172-182.

[First analysis of life-strategies of species found in terrestrial bryophyte communites

along a transect across the Judean Desert; seven life-strategies together with their

characters are described and the zonation and distribution patterns along the ecological

gradient demonstrated].

FREY W. & KURSCHNER H., 199le — Morphologische und anatomische

Anpassungen der Arten in terrestrischen Bryophytengesellschaften entlang

eines ökologischen Gradienten in der Judäischen Wüste. Botanische Jahrbü-

cher für Systematik, Pflanzengeschichte und Pflanzengeographie 112: 529-552.

[Life-forms, water conducting and water storing structures, protection structures

against light and temperature, as well as sporophyte characters (morphological and

anatomical adaptations) of species found in terrestrial bryophyte communites along a

transect across the Judean Desert are investigated; their relation to ecological

conditions and to distribution patterns of the communities is demonstrated].

FREY W. & KURSCHNER H., 1992 — Bryosoziologische Untersuchungen in

Jordanien: terrestrische und epilithische Gesellschaften. Nova Hedwigia 54:

355-378.

[Phytosociological study (description of six associations and two communities) with

detailed remarks on syntaxonomy, community structures, ecological conditions, habitat

sites and distribution of the communities; includes a synopsis of the bryophyte

communities of Israel and Jordan].

FREY W. & KURSCHNER H., 1993a — Targionia hypophylla L. ssp. linealis

(Marchantiidae, Hepaticae), eine neue Unterart aus Saudi Arabien sowie

weitere Neufunde. Studien an arabischen Bryophyten 17. Nova Hedwigia 57:

127-133.

[Includes the description and illustration of a new subspecies. Adds three hepatic and

seven moss taxa to the flora of Saudi Arabia, two hepatics and four mosses being new

to the Arabian Peninsula].

FREY W. & KURSCHNER H., 1993b — Trichostomopsis trivialis (С. Müll.) Robins.

(Pottiaceae, Musci) eine südafrikanisch-mediterran disjunkte Sippe, neu für

die Bryoflora Jordaniens. Cryptogamic Botany 3: 152-156,

{Reports a new record with illustrations, description, notes on habitat and distribution;

includes a key to the Trichostomopsis species of the Near and Middle East].

FREY W. & KURSCHNER H., 1994 — Bryosoziologische Untersuchungen in

Jordanien 2. Ergänzungen und weitere neue terrestrische und epilithische

Gesellschaften aus der östlichen Wüste und der Basaltregion. Nova Hedwigia

59: 365-378.

[Phytosociological study, adding three associations and one subassociation to the

vegetation of Jordan; includes remarks on syntaxonomy, community structures,

ecological conditions, habitat sites and distribution of the communities].

FREY W., KÜRSCHNER Н., ROS R.M., GUERRA J. & CANO М. J., 1994 —

Tortula mucronifera (Pottiaceae, Musci), a new xerophytic species of the

Source : MNHN. Paris

18 H. KURSCHNER

Arabian Peninsula and Jordan. Studies in Arabian bryophytes 18. Nova Hedwigia 59: 345-351.

[One new species described and illustrated, including notes on floristic affinities, ecology

and distribution).

FREY W. & KURSCHNER H., 1995a Bryosoziologische Untersuchungen in

Jordanien. 3. Lebensstrategienanalyse der terrestrischen und epilithischen

Moosgesellschaften. Fragmenta Floristica et Geobotanica 40: 491-511.

[Analysis of life-strategies of species found in terrestrial and epilithic bryophyte

communities; six life-strategies together with their characters are described and the

zonation and distribution pattern along the ecological gradient demonstrated].

FREY W. & KURSCHNER Н., 1995b — Soziologie und Lebensstrategien epiphy-

tischer Bryophyten in Israel und Jordanien. Nova Hedwigia 61: 211-232.

[Phytosociological study (description of four associations and two fragment commu-

nites) with notes on syntaxonomy, community structures, ecological conditions, habitat

sites and distribution of the communities; includes a life-strategy analysis, demonstrat-

ing the strong correlation of life-forms, life-strategies and adaptations to the ecological

conditions}.

FRITSCH К., 1899 — Beitrag zur Flora von Constantinopel. Bearbeitung der von J.

Nemetz in den Jahren 1894-1897 in den Umgebungen von Constantinopel

gesammelten Pflanzen. I. Kryptogamen. Denkschriften der Kaiserlichen Aka-

demie der Wissenschaften, Wien. Mathematisch-naturwissenschaftliche Klasse

68, Abteilung 2: 219-250.

[Hepaticae on pp. 243-244, Musci on pp. 245-246; list of seven hepatic (identified by F.

Matouschek, V. Schiffner) and 19 moss taxa (identified by F. Matouschek) with

collection data].

FROEHLICH J., 1950 — Bryophyten aus Iran (Rechingeri iter iranicum secundum —

No. 9). Annalen des naturhistorischen Museums in Wien 57; 37-41.

[List of four hepatic and 56 moss taxa with collection data, one hepatic and 17 mosses

being new to Iran; includes the descriptions of Didymodon planifolius sp. nov. (= D.

planatophaceus nom. nov., cf. Froehlich 1953), Philonotis laxitexta sp. nov. and

Mielichhoferia compacta vat. papillosula var. nov].

FROEHLICH J., 1953 — Berichtigung zu Bryophyten aus Iran (Rechingeri iter

iranicum secundum — No. 9). Annalen des naturhistorischen Museums in Wien

59: 118.

[Nomenclatorical note, concerning the state of Didymodon planifolius nom. hom. = D.

planatophaceus nom. nov.].

FROEHLICH J., 1955 — Die von Dr. A. Gilli in den Jahren 1949 bis 1951 in

Zentral-, Ost- und Nordost-Afghanistan gesammelten Bryophyten. Mitteilun-

gen der Thiiringischen botanischen Gesellschaft 1: 59-70.

[List of four hepatic and 84 moss taxa with collection data, 46 mosses being new to

Afghanistan; includes the descriptions of Bryum funckiioides sp. nov., B. purpureo-

lucidum sp. nov., Tortula cabulica sp. nov., Т. cucullifolia sp. nov., Т. subtranscaspica

sp. nov. and Didymodon recurvirostris var. obtusifolius var. nov.].

FROEHLICH J., 1959 — Bryophyten aus Vorderasien. Annalen des naturhistorischen

Museums in Wien 63: 31-32.

[List of three hepatic and 40 moss taxa with collection data (collected by K. H.

Rechinger in Iraq and Turkey); includes the descriptions of Barbula rigidula var.

desertorum var. nov. and Polytrichum alpinum var. piriforme var. nov.].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 19

FROEHLICH J., 1964 — Bryophyten aus Afghanistan und Nordwest-Pakistan.

Annalen des naturhistorischen Museums in Wien 67: 149-158.

[List of one hepatic and 77 moss taxa with collection data (collected by K. H. Rechinger

in Afghanistan); includes the descriptions of Anoectangium pleuroweisioides sp. nov.,

Barbula cucullata sp. поу., Bryoerythrophyllum compactum sp. nov., Hydrogonium

heterophyllum sp. nov., Н. patulifolium sp. nov., Hymenostylium diversifolium sp. nov.

Mniobryum tortifolium sp. nov., Tortula pseudo-desertorum sp. nov., and Т. pseudo-

handelii sp. nov.].

GEHEEB A., 1902 — Beitrag zur Moosflora von Syrien. Allgemeine botanische

Zeitschrift 8: 42-44.

{Adds two hepatic and 10 moss taxa to the known flora of the vicinity of Beirut

(Lebanon)].

GEHEEB A., 1903/1904 — Bryophyta. In: KNEUCKER A.: Botanische Ausbeute

einer Reise durch die Sinaihalbinsel vom 27. Márz bis 13. April 1902.

Allgemeine botanische Zeitschrift 9: 185-189, 203-204; Ibidem 10: 4-5.

[Adds 20 moss taxa to the known flora with collection data and notes to the

distribution, 14 being new to the Sinai Peninsula; includes the descriptions of Tortula

kneuckeri sp. nov. and T. rigescens sp. nov.].

GILLI A., 1941 — Ein Beitrag zur Flora des Elbursgebirges in Nordiran. Feddes

Repertorium 50: 263-283.

[Mosses on p. 283; lists five mosses with collection data].

GILLI A., 1969 — Afghanische Pflanzengesellschaften. Vegetatio 16: 307-375.

[Phytosociological study of the xerophytic plant-communities; includes a few mosses

(Bryum affine, B. subrotundum, Distichium capillaceum var. compactum, Tortula

cucullifolia, Т. desertorum ) in the phytosociological relevées].

GILLI A., 1971 — Afghanische Pflanzengesellschaften. II. Die mesophilen und

hygrophilen Pflanzengesellschaften im sommertrockenen Gebiet. Vegetatio

23: 199-234.

[Phytosociological study of the meso- and hygrophytic plant-communities; includes

several hygrophytic mosses and liverworts in the phytosociological relevées (e.g. р. 221.

Adiantetum capilli-veneris with one hepatic and 10 moss taxa; р. 222, Heracletum persici

with five mosses; p. 223 Ligularierum thomsonii with four mosses; p. 228 Poetum annuae

with six mosses].

GILLI A., 1975 — Afghanische Ruderal- und Segetalgesellschaften. Feddes Reperto-

rium 86: 171-197.

[Phytosociological study of the ruderal and segetal plant-communities; includes one

hepatic (Mannia androgyna ) and three mosses (Amblystegium varium, Barbula fallax,

Philonotis falcata ) in the phytosociological relevées].

GILLI A., 1977 — Die Waldgebiete im Osten Afghanistans. Feddes Repertorium 88:

375-387.

[Phytosociological study of the forests and woodlands; includes several terrestrial,

epilithic and epiphytic bryophytes in the phytosociological relevées].

Source : MNHN. Paris

20 H. KURSCHNER

GÖKLER І., INOUE Н. & ÖZTÜRK M., 1983/1984 — A new record for Turkey,

Pellia neesiana (Gott.) Limpr. Ege University Journal of Science Faculty, Serie

B 7: 85-89 (publ. April 1984).

[Reports a new record with description and collection data].

GÖKLER 1, ÖZTÜRK М. & KESERGIOGLU T., 1985/1986 — Checklist

of liverworts (Hepaticae) recorded from Turkey. Ege University Journal of

Science Faculty, Serie B 8: 1-10 (publ. April 1985).

[Checklist of 143 taxa based mainly on older literature, showing the distribution

within the country on a grid basis (turkish grid square system introduced by Davis

1965)].

GOKLER L, 1986 — Taxonomical investigations on some liverworts (Hepaticae)

distributed in Turkey. II. Marchantiales and Anthocerotales. Doga, Türk

Biyoloji Dergisi Seri C 10: 353-361. In Turkish, with English summary.

[Morphological and anatomical study of five common Marchantiidae (Conocephalum

conicum, Lunularia cruciata, Marchantia polymorpha, Plagiochasma rupestre, Targionia

hypophylla) and Anthoceros punctatus).

GOKLER I. & OZTURK M., 1986 — Taxonomical investigations on some liverworts

(Hepaticae) distributed in Turkey. 1. Jungermanniales Anacrogynae and J.

Acrogynae. Doga, Tiirk Biyoloji Dergisi Seri C 10: 163-170. In Turkish, with

English summary.

[Morphological and anatomical study of seven Jungermanniidae (Fossombronia pusilla,

Frullania dilatata, Lophocolea cuspidata, Metzgeria furcata, Pellia endiviifolia, Porella

cordeana, P. platyphylla) and distributional notes].

GOKLER I. & OZTURK M., 1987a — A new record for Turkey, Porella thuja

(Dicks.) C. Jens. Doga, Turkish Journal of Botany 11: 313-315.

[Reports a new record with description, illustration, collection data and world-wide

distribution].

GOKLER I. & ÖZTÜRK M., 1987b — New concepts concerning the methods used

in the identification of liverworts (Hepaticae). Doga, Türk Botanik Dergisi

Seri C 11: 306-312. In Turkish, with English summary.

[General account on the available methods used in bryophyte taxonomy].

GÖKLER I., 1989 — A new record for the hepatic flora of Turkey, Lejeunea

lamacerina (Steph.) Schiffn. Doga, Türk Botanik Dergisi Seri C 13: 470-473.

In Turkish, with English summary.

[Reports a new record with illustrations, description and locality data].

GOKLER I. & OZTURK M., 1989 — An investigation on the liverworts (Hepaticae)

of Black Sea region. Doga, Tiirk Botanik Dergisi Seri C 13: 242-248.

[List of 27 hepatic taxa based on the authors’ collections with collection data; Bazzania

trilobata is reported for the first time from Turkey].

GOKLER І. & ÖZTÜRK M., 1991 — Liverworts of Turkey and their position in

South-West Asia. Candollea 46: 359-366.

[Checklist of 145 taxa based mainly on older literature, showing the distribution within

the country on a grid basis (turkish grid square system introduced by Davis 1965);

includes remarks on the phytogeographical relationships and a new record, Asterella

lindenbergiana which is only a Corsinia coriandrina (cf. Kürschner 1996b)].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 21

GÖKLER L, 1992 — An investigation on the liverworts of West Anatolia. Doga,

Turkish Journal of Botany 16: 1-8. In Turkish, with English summary.

[Reports two anthocerote and 32 hepatic taxa with collection data based on the

author’s collections, four being new to square C 11 and seven new to the western part

of Turkey (Aegean region)].

GRIFFITH W., 1849 — Notulae ad plantas asiaticas, part IL. On the higher

cryptogamous plants. Calcutta, pp. 257-628.

[Hepaticae on pp. 299-352, mosses on pp. 376-485; includes some hepatic and moss taxa

with collection data from Afghanistan].

GUERKE W. R., 1978 — A monograph of the genus Jubula Dumortier. Bryophytorum

Bibliotheca 17: 1-118.

[Floristic treatment of the genus Jubula with key to the species, notes on taxonomy,

habitat and distribution; includes locality data from Iran].

HALWAGY M. & EL-SAADAWI W. E., 1992 — Drought and changes in the

bryoflora and angiosperm flora in Kuwait in the years 1974-1990. Acta

Botanica Neerlandica 41: 183-195.

[Ecological study, indicating drastic changes in growth-rates and species diversity of 12

mosses due to successive years of drought].

HANDEL-MAZZETTI H. v., 1905 — Hepaticae. In: PENTHER A. & ZEDER-

BAUER E.: Ergebnisse einer naturwissenschaftlichen Reise zum Erdschias-

Dagh, S. 387-388. Annalen des naturhistorischen Museums in Wien 20:

359-464.

[List of one anthocerote and five hepatic taxa with collection data].

HANDEL-MAZZETTI Н. v., 1909 — Musci. Іп: HANDEL-MAZZETTI Н. v.:

Ergebnisse einer botanischen Reise in das Pontische Randgebirge im Sands-

chak Trapezunt, unternommen im Jahre 1907, S. 124-132. Annalen des

naturhistorischen Museums in Wien 23: 6-212.

[List of 138 mosses with collection data and details of localities, 14 being new to the

flora].

HANDEL-MAZZETTI H. v., 1914 — Die Vegetationsverhältnisse von Mesopota-

mien und Kurdistan. Annalen des naturhistorischen Museums in Wien 28:

48-111.

[General description of the main vegetation units of Syria and Iraq; includes also

mosses and liverworts in the vegetation tables and habitat notes].

HART H. C., 1891 — Some acount of the fauna and flora of Sinai, Petra and Wadi

Arabah. London, 255 p.

[Musci on pp. 118-119, 168-169, Hepaticae on pp. 119-120, 170-172; list of 27 moss and

four hepatic taxa with collection data and notes on distribution].

HATTORI 5., 1976 — Studies on the asiatic species of the genus Porella (Hepaticae).

VI. Journal of the Hattori Botanical Laboratory 40: 121-138.

[Descriptions and illustrations of asiatic Porella species; includes P. arboris-vitae, new

to Turkey with collection data].

Source :MNHN. Paris

22 H. KURSCHNER

HATTORI S., 1978a — Studies on the asiatic species of the genus Porella (Hepaticae).

VII. A synopsis of asiatic Porellaceae. Journal of the Hattori Botanical

Laboratory 44: 91-120.

[Floristic treatment of the asiatic Porellaceae with key to the genera, species,

descriptions and illustrations; includes phytogeographical data from Afghanistan]

HATTORI 5., 1978b — A note on asian Porella platyphylla. Miscellanea Bryologica et

Lichenologica 8 : 1-2.

Includes locality data from Afghanistan].

HEBRARD J. Р., 1982 — Mousses et hépatiques du bassin méditerranéen. Beihefte zur

Nova Hedwigia 71: 369-386.

(General account on the state of bryofloristic knowledge; useful literature list].

HENDERSON D. M. & MUIRHEAD C.W., 1955-1958a — Contributions to the

bryophyte flora of Turkey. Notes from the Royal Botanic Garden Edinburgh

22: 29-43 (publ. Oct. 1955).

[List of 109 moss and 20 hepatic taxa with collection data and some taxonomic notes,

based on the collections of P. H. Davis 1949 in western Turkey, and Р. Н. Davis & J.

Dodds 1952 in central and south-eastern Turkey and the Black Sea area; adds 42 moss

and 10 hepatic taxa to the known bryoflora].

HENDERSON D. M. & MUIRHEAD C.W., 1955-1958b — Eastern mediterranean

bryophytes. Notes from the Royal Botanic Garden Edinburgh 22: 121-129

(publ. Oct. 1956).

[List of 63 moss, seven hepatic and one anthocerote taxa with collection data, some

taxonomic and distribution data, based on the collections of P. H. Davis 1943 in

Lebanon and Syria; includes an illustration of Tortula papillosissima].

HENDERSON D. M., 1955-1958a — Contributions to the bryophyte flora of Turkey:

П. Notes from the Royal Botanic Garden Edinburgh 22: 189-193 (publ. Sept.

1957).

{List of 38 moss and one hepatic taxa with collection data and some taxonomic notes,

based on the collections of Р. Н. Davis 1954 in south-eastern Turkey; adds 10 taxa to

the known bryoflora].

HENDERSON Р. М., 1955-1958b — Contributions to the bryophyte flora of Turkey:

Ш. Notes from the Royal Botanic Garden Edinburgh 22: 611-620 (publ. Dec.

1958).

[List of 64 moss, four hepatic and two anthocerote taxa with collection data and some

taxonomical notes, based on the collections of P. H. Davis & O. Polunin 1956 in

western and south-western Turkey, and J. McNeill 1956 in central and south-eastern

Turkey; one new species, Grimmia cucullata is described and illustrated].

HENDERSON D. M., 1959-1961a — Contributions to the bryophyte flora of Turkey:

IV. Notes from the Royal Botanic Garden Edinburgh 23: 263-278 (publ. June

1961).

[List of 108 moss and eight hepatic taxa with collection data, based on the extensive

collections of P. Н. Davis & I. C. Hedge 1957 and a few records of L. F. Н. Merton

1958 from the vicinity of Ankara].

HENDERSON D. M., 1959-1961b — Contributions to the bryophyte flora of Turkey:

У. Summary of present knowledge. Notes from the Royal Botanic Garden

Edinburgh 23; 279-301 (publ. June 1961).

[Summarizes the present state of bryofloristic knowledge in tabular form, showing the

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 23

distribution within the country on a grid basis (turkish grid square system introduced

by Davis 1965); includes a short plantgeographical overview].

HENDERSON D. М. & PRENTICE Н. T., 1963 — Bryophyta. In: HEDGE I. С. &

WENDELBO P.: Studies in the flora of Afghanistan 1, pp. 29-31. Arbok for

Universitetet i Bergen. Mat.-Naturv. Serie No. 18: 1-56.

[List of 20 moss and one hepatic taxa with collection data, based on the collections of

I. C. Hedge & P. Wendelbo].

HENDERSON D. M., 1963-1964 — Contributions to the bryophyte flora of Turkey:

VI. Notes from the Royal Botanic Garden Edinburgh 25: 279-291 (publ. July

1964).

List of 103 moss and 15 hepatic taxa with collection data, based on the collections of

D. M. Henderson & 1. D. A. Stainton 1960 and a few records of L. Е. Н. Merton 1958

from the vicinity of Ankara].

HENDERSON D. M. & PRENTICE H. T., 1969 — Contributions to the bryophyte

flora of Turkey: УШ. Notes from the Royal Botanic Garden Edinburgh 29:

235-262.

Checklist in tabular form of mosses, anthocerotes and hepatics, showing the

distribution within the country on a grid square system; useful literature list].

HEPPER Е. N., 1977 — Outline of the vegetation of the Yemen Arabic Republic.

Publications from the Cairo University Herbarium 7/8: 307-322.

Short overview on the landscapes and vegetation of northern Yemen; reports Hypnum

cupressiforme, Pleurochaete squarrosa, Trichostomopsis sp. and Trichostomum brachy-

dontium from the rocky slopes of the Montane Plains (hills around Sumara, Yerim,

Yizla Pass, Haddah and Shibam/Kawkaban)].

HERRNSTADT І., HEYN С.С. & CROSBY М. R., 1980 — New data on the moss

genus Gigaspermum. The Bryologist 83: 536-541.

Reports a new record, G. mouretii for Israel, with illustrations, notes on taxonomy and

world-wide distribution].

HERRNSTADT І., HEYN С. С., BEN-SASSON R. & CROSBY М. R., 1982 — New

records of mosses from Israel. The Bryologist 85: 214-217.

[Reports 44 species new to Israel, with collection data and some taxonomic notes].

HERRNSTADT I. & HEYN C. C., 1987 — Bryology in Israel. Taxon 36: 772-

773.

[Reviews the history and state of bryological research].

HERRNSTADT І., HEYN С.С. & CROSBY М. R., 1991 — A checklist of the mosses

of Israel. The Bryologist 94: 168-178.

[Checklist of 232 mosses, 45 being new to the flora of Israel, with some taxonomic notes

and distribution records based on squares of 10 х 10 km; includes the descriptions of

Phascum galilaeum sp. nov., P. cuspidatum var. arcuatum var. nov. and Р. cuspidatum

var. marginatum var. nov.].

HERRNSTADT L, 1992 — Check-list of the bryophytes collected during Iter

Mediterraneum П. Bocconea 3: 217-222.

[List of 41 moss and two hepatic taxa with collection data and some taxonomic notes,

collected occasionally by some members of the 2nd OPTIMA excursion to Israel

(19.3.-10.4.1992)].

Source : MNHN. Paris

24 H. KURSCHNER

HERRNSTADT I. & HEYN C. C., 1993 — New species linking Phascum and Pottia

(Pottiaceae). Nova Hedwigia 57: 135-139.

[Presents a detailed description and illustration of the previously described species

Phascum galilaeum (cf. Herrnstadt et al. 1991)].

HERZOG T., 1938 — Botanische Ergebnisse der Deutschen Hindukusch-Expedition

1935. Bryophyta. Feddes Repertorium, Beiheft 108: 3-12.

[List of two hepatic and 31 moss taxa with collection data and distributional notes;

includes the descriptions of Brachythecium plumosum var. laxo-alare var. nov.,

Dicranoweisia brevifolia sp. nov. (with illustration), and Weisia viridula var. remotidens

var. nov.].

HILPERT F., 1933 — Studien zur Systematik der Trichostomataceae. Beihefte zum

Botanischen Centralblatt 50: 587-706.

[Floristic treatment of the Pottiaceae subfam. Trichostomoideae, including moss

records from Sinai]

IGNATOV M. S., 1995 — On the identity of Rhynchostegium haussknechtii (Musci,

Brachytheciaceae). Fragmenta Floristica et Geobotanica 40; 197-199.

[R. haussknechtii, described from eastern Turkey and known only from the type

collection appears to be identical to Fontinalis antipyretica var. heldreichii, with

illustration of the lectotype].

IMAM М. & GHABBOUR 5. I., 1972 — A contribution to the moss flora of Egypt.

Botaniska Notiser 125: 518-522.

[List of 60 mosses from Egypt with collection data, including one taxon, Brachythe-

cium rivulare from the Sinai Peninsula (collected by M. Hasib without exact locality

data)].

JONES E. W., 1969 — Hepaticae. In: TOWNSEND С. C.: Bryophytes from

Socotra, pp. 795-796. Transactions of the British Bryological Society 5:

790-796.

[List of three hepatic taxa with collection data based on the collections of A. R. Smith

1967 (Middle East Command Expedition); includes a fuller description of Fimbriaria

pusilla Mitt).

JONES E. W., 1985 — African hepatics XXXIV. — Some little known or new

Lejeuneaceae. Journal of Bryology 13: 385-398.

[Reports Lejeunea aethiopica, collected by D. G. Long & A. G. Miller (E 10150) from

Yemen].

JOVET-AST S., 1957 — Riccia frostii Aust. au Sahara et en Turquie. Revue

bryologique et lichénologique 26 : 67-68.

[Adds a further locality of R. frostii, collected Бу Н. de Lesse in the vicinity of Amasya,

to Turkey].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 25

JOVET-AST S., 1960 — Quelques Bryophytes d’Iran. Revue bryologique et lichénolo-

gique 29 : 181 (publ. march 1961).

[List of one anthocerote, four hepatic and eight moss taxa with collection data, based

on the collection of R. Heim 1960 along the Caspian Sea].

JOVET-AST S., 1961 — Targionia lorbeeriana К. M., trois localités nouvelles. Revue

bryologique et lichénologique 30 : 278.

[Adds a further locality of Т. lorbeeriana, collected by L. Kofler 1960 in the vicinity of

Jerusalem, to Israel].

JOVET-AST S., BISCHLER Н. & BAUM B., 1965 — Hépatiques récoltées en Israel.

Israel Journal of Botany 14 : 36-48.

[List of two anthocerote and 23 hepatic taxa, nine being new to Israel; with locality data

and notes on habitat sites].

JOVET-AST S. & BISCHLER H., 1966 — Les Hépatiques d'Israël : Enumération,

notes écologiques et biogéographiques. Revue bryologique et lichénologique 34 :

91-126.

[Checklist of 31 hepatic and two anthocerote taxa based on the literature and the

authors’ collections, with notes on taxonomy, distribution, habitats and specimens].

JOVET-AST S., 1970 — Cyathodium africanum Mitt. au Yémen et en Afrique. Revue

bryologique et lichénologique 37: 57-62.

[Reports a new record to the flora of Yemen collected obviously by G. Schweinfurth

(not Forsskal, as indicated, "Manod, oberes W. Madfar, bei Hodjela, 30. Jan. 1889")

and kept at the Herbarium Stephani at Geneva; includes morphological description,

illustrations and notes on distribution].

JOVET-AST S. & BISCHLER H., 1970 — Les Hépatiques d'Egypte et du Sinai.

Enumération, notes écologiques et biogéographiques. Revue bryologique et

lichénologique 37 : 265-290.

[Checklist of 13 species based on the authors' collections and records in the literature,

with notes on habitats and ecology].

JOVET-AST S., BISCHLER H. & BAUDOIN R., 1976 — Essai sur le peuplement

hépaticologique de la région méditerranéenne. Journal of the Hattori Botan-

ical Laboratory 41 : 87-94.

[Study on the ecological requirements of selected mediterranean liverworts, with notes

оп ecology and distribution maps (covering Israel and Turkey)].

JOVET-AST S., 1981 — У. Hepatiques. In: LEONARD J.: Contribution à l'étude de

la flore et de la vegetation des deserts d’Iran, Fasc. 1, pp. 26-27. Bruxelles.

[Reports Riccia cavernosa and К. frostii from southeastern Iran (Jaz Murian area)].

JOVET-AST S., 1983 — Riccia trichocarpa Howe et Riccia canescens Steph.

Cryptogamie, Bryologie-Lichénologie 4; 37-46.

[Taxonomic treatment; A. canescens is considered as a synonym of R. trichocarpa;

includes notes on synonymy, detailed description, illustrations, ecological data and

collection data from Iran].

JOVET-AST S., 1986 Les Riccia de la région méditerranéenne. Cryptogamie,

Bryologie-Lichénologie 7, supplément 3 : 287-431.

[Floristic treatment of the genus Riccia in the Mediterranean area with key to the

species, descriptions and illustrations and some general distribution remarks on Oman,

Saudi Arabia, and Yemen].

Source . MNHN, Paris

26 H. KÜRSCHNER

JOVET-AST S., 1989 — Un complexe de taxons dans le genre Riccia. Cryptogamie,

Bryologie-Lichénologie 10 : 95-117.

[Floristic treatment of six taxa of the genus Riccia, with key to the species, descriptions,

illustrations and distribution maps; includes general distribution remarks on the

Arabian Peninsula and Iran, and locality data from Oman (К. atromarginata var.

Jovet-astii )].

JURATZKA J. & MILDE J., 1870 — Beitrag zur Moosflora des Orientes. Kleinasien,

das westliche Persien und den Caucasus umfassend. Verhandlungen der

zoologisch-botanischen Gesellschaft in Wien 20: 589-602.

[List of с. 150 moss taxa with collection data, based on the collections of С.

Haussknecht 1865 and 1868; includes the descriptions of Atrichum haussknechtii sp.

nov., Barbula haussknechtii sp. nov., Brachythecium umbilicatum sp. nov., Bryum

dalachanicum sp. nov., В. juratzkae sp. nov., Encalypta intermedia sp. nov., Entosthodon

angustifolius sp. nov., Fissidens persicus sp. nov., Leucodon caucasicus sp. nov.,

Rhynchostegium haussknechtii sp. nov. (cf. Ignatov 1995), Trichostomum mildeanum sp.

nov., and T. persicum sp. nov.].

KILBERTUS С. & BIZOT M., 1971 — Contribution à la connaissance de la flore des

muscinées d'Afghanistan. Bulletin de l'Académie de la Société Lorraine des

Sciences 10 (2) : 67-75,

[List of 41 mosses with collection data based on the authors' collections; includes the

description and illustration of Hygrohypnum smithii var. denticulata var. nov.].

KILPPER K., 1964 — Über eine Rhät/Lias-Flora aus dem nördlichen Abfall des

Alburs-Gebirges in Nordiran. Teil 1: Bryophyta und Pteridophyta. Palaeon-

tographica, Abteilung B 114: 1-78.

[Report of a Rhaeto-Liassic flora from the Karmozd-Zirab area (Northern Iran);

includes one find of a thallose liverwort, Hepaticites cf. arcuatus, and remarks on the

systematic affinities].

KITAGAWA М., 1964 — Bryophytes of Afghanistan. In: KITAMURA S.: Results of

the Kyoto University scientific expedition to Karakorum and Hindu Kush

1955. 3. Plants of West Pakistan and Afghanistan, pp. 163-166. Kyoto.

[List of three hepatic and 27 moss taxa with collection data and notes of the world-wide

distribution, based on the collections of S. Kitamura].

KOPONEN T., 1980 — A synopsis of Mniaceae (Bryophyta). IV. Taxa in Europe,

Macaronesia, NW Africa and the Near East. Annales Botanici Ееппісі 17:

125-162.

[Floristic treatment of the Mniaceae (32 taxa) with key to the genera (Cinclidium,

Cyrtomnium, Mnium, Plagiomnium, Rhizomnium, Trachycystis) and species, notes on

synonymy and illustrations; includes collection data from Afghanistan, Iran, Iraq,

Israel, Lebanon, Syria, and Turkey].

KOPONEN T., 1982 — Miscellaneous notes on Mniaceae (Bryophyta). ХІ. Distribu-

tion of Plagiomnium rostratum. Memoranda Societatis pro Fauna et Flora

Fennica 58: 17-20.

[Mapping of the total distribution of Р. rostratum; includes collection data from

Afghanistan, Iraq, Turkey].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 27

KOPONEN T., 1993 — Miscellaneous notes on Mniaceae (Bryophyta). XVI.

Plagiomnium undulatum (Hedw.) T. Kop. in Ethiopia. Bulletin du Jardin

Botanique National de Belgique 62: 397-402.

[Mapping of the distribution of P. undulatum; includes the area of the Near and Middle

East].

KRAMER W., 1978 — Contribution to the taxonomy and distribution of

several taxa of Tortula Hedw. sect. Rurales De Not. (Musci) with bistratose

lamina. The Bryologist 81: 378-385.

[Taxonomic treatment of T. caninervis, T. handelii, T. handelii var. ferganensis (comb.

nov.) and T. pseudo-handelii; includes illustrations, notes on synonymy and collection

data from Afghanistan, Iran, Lebanon, Syria, and Turkey; 7. desertorum, T.

pseudo-desertorum, and Т. saharae are reduced to synonyms of Т. caninervis, and Т.

handelii is reported new to Iran, while T. pseudo-handelii is new to Iran and Turkey].

KRAMER W., 1980 — Tortula Hedw. sect. Rurales De Not. (Pottiaceae,

Musci) in der östlichen Holarktis. Bryophytorum Bibliotheca 21: 1-165.

[Systematic treatment of Tortula (Syntrichia) from the Near and Middle East, with key

to the species, illustrations, descriptions, locality data (Afghanistan, Iran, Iraq, Israel,

Jordan, Lebanon, Sinai, Syria, Turkey), and notes on taxonomy and distribution;

includes several new names, statuses and new combinations, and the descriptions and

illustrations of Tortula ruralis subsp. hirsuta var. submamillosa var. nov., T. ruralis

subsp. ruralis var. substereidosa var. nov., and T. virescens var. iranica var. nov.].

KÜRSCHNER H., 1984a — Epiphytic communities of the Asir Mountais (SW Saudi

Arabia). Studies in Arabian bryophytes 2. Nova Hedwigia 39: 177-199.

[Study of the unique epiphytic bryophyte and lichen communities of the monsoon-

influenced woodlands of south-western Arabia (coverage values, association analysis),

with notes on community structures, ecological conditions, habitat sites, distribution

and phytogeographical relationships; adds one hepatic and nine moss taxa to the flora

of Saudi Arabia].

KÜRSCHNER H., 1984b — An epilithic bryophyte community in the Asir Mountains

(SW Saudi Arabia). Studies in Arabian bryophytes 4. Nova Hedwigia 40:

423-434.

[Study of an epilithic bryophyte communitiy of the Arabian Peninsula (coverage values,

importance values) with notes on community structure, succession cycles (initial,

optimal, terminal phases), ecological conditions, habitat sites and distribution; adds one

hepatic and four moss taxa to the flora of Saudi Arabia].

KÜRSCHNER H., 1989 — Bryophytes from Saudi Arabia, collected by І. S.

Collenette. Studies in Arabian bryophytes 15. Nova Hedwigia 48: 73-83.

[List of nine hepatic and 41 moss taxa with collection data and notes on habitat and

distribution, three hepatics and 12 mosses being new to the flora of Saudi Arabia;

includes some bryogeographical remarks which indicate a well established Afromon-

tane exclave in the Arabian Peninsula].

KÜRSCHNER H., 1989 [1991] — Studies in the bryophyte vegetation of Saudi

Arabia: terrestrial, epilithic and epiphytic communities. Studies in Arabian

bryophytes 14. In: HERBEN T. & McQUEEN C. B.(eds.): Proceedings of the

Sixth Meeting of the Central and East European Bryological Working Group

(CEBWG), pp. 149-163. Prühonice.

[Reviews the state of knowledge in bryovegetation research; includes the epiphytic

Fabronietum socotranae prov. ass. nov., Hypno-Leucodontetum dracaenae prov. ass.

Source : MNHN. Paris

28 H. KURSCHNER

nov. and the description of a hydrophytic synusium (Gymnostomum aeruginosum-

synusium of the Adiantho-Primuletum verticillatae ) of rock cavities with dripping water

in the montane belt of south-western Arabia (Saudi Arabia, Yemen].

KÜRSCHNER H., 1995 — Weissia ovatifolia (Pottiaceae, Musci), eine neue Art aus

dem Edomitischen Bergland Jordaniens. Nova Hedwigia 60: 499-504.

[One new species described and illustrated; includes notes on the ecology and floristic

affinities].

KURSCHNER H., 1996a — Additions to the bryophyte flora of Yemen. New records

from the Taizz and Jiblah areas. Studies in Arabian bryophytes 21. Nova

Hedwigia 62: 233-247.

[Adds 16 mosses to the flora of northern Yemen with notes on collection data and

phytogeographical relationships, six being new to the Arabian Peninsula; includes a list

of all published taxa, a first bryogeographical analysis of Yemen (including Socotra)

and some distribution maps of typical paleotropical and Afromontane taxa].

KÜRSCHNER Н., 1996b — Towards a bryophyte flora of the Near and Middle East.

New records from Iran, Jordan, Kuwait, Lebanon, Oman, Saudi Arabia,

Syria, and Turkey. Nova Hedwigia 63: 261-271.

[List of 31 new records with collection data for various countries of the Near and

Middle East based on the author's collections and older collections (e.g. С. В. Arzeni

from Lebanon, W. N. Koeltz, L. Р. Н. Merton, and 5. Agnew from Iran, О. Volk and

S.-W. Breckle from Afghanistan); adds 15 taxa to the known flora of Iran, three to

Jordan, two to Kuwait, one to Oman, three to Syria, and three to Turkey). Includes

also the locality data of 39 previously reported taxa (ef. Frey & Kürschner 1991b) from

Afghanistan, Iran, Jordan, Lebanon, Saudi Arabia, Syria, and Turkey].

LIPKIN Y. & PROCTOR V. W., 1975 — Notes on the subgenus Trabutiella of the

aquatic liverwort Riella (Riellaceae, Sphaerocarpales). The Bryologist 78:

25-31.

[Study of Riella populations of Israel (crossing-experiments, spore morphology)

concerning the taxonomic significance of several characters; А. paulsenii is reduced to

a synonym of R. cossoniana, and R. cossoniana var. echinata should be regarded as a

synonym of R. helicophylla].

LONG D. G., 1979 — Some additions to the bryophyte flora of Iraq. Revue

bryologique et lichenologique 45: 103-105.

[Adds three moss and one hepatic taxa collected by W. B. Evans during late 1918 and

early 1919 to the known flora].

LONG D.G., 1982 — Collection and presentation of bryophytes in Arabia. Bulletin of

the Emirates Natural History Group 18: 18-19,

[Information how to collect and preserve bryophytes in arid countries].

LONG D.G., 1986 — Some Socotran mosses collected by Schweinfurth. Studies in

Arabian bryophytes 5. Nova Hedwigia 43: 443-445.

[Enumerates four mosses collected by G. Schweinfurth 1881 on Socotra and reported

by Müller (1901), which has not been mentioned again by recent authors; Edentella

schweinfurthii is shown to be invalid and a synonym of Pseudosymblepharis soco-

trana].

Source : MNHN., Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 29

LONG D.G., 1987 — Hepaticae and Anthocerotae of the Arabian Peninsula. Studies

in Arabian bryophytes 10. Nova Hedwigia 45: 175-195.

[List of one anthocerote and 31 hepatic taxa with synonyms, details of specimen,

habitats and world-wide distribution, based on literature and recent collections (mainly

1. 5. Collenette 1979-1984, D. б. Long & A. б. Miller 1981, A. б. Miller 1978, 1979,

1984, J. R. I, Wood 1976-1981); includes a preliminary phytogeographic assessment and

an extensive literature list].

LORENTZ P. G., 1868 — Über die Moose, die Hr. Ehrenberg in den Jahren

1820-1826 in Aegypten, der Sinaihalbinsel und Syrien gesammelt. Physikali-

sche Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin

1868: 1-57 (mit 15 Tafeln).

[List of 11 moss taxa from Sinai and 18 moss taxa from Syria with collection data;

includes the descriptions and illustrations of Brachythecium syriacum sp. nov., Bryum

syriacum sp. nov., Philonotis caespitosa var. syriaca var. noy., Polytrichum commune

var. ehrenbergii var. nov., Trichostomum aaronis sp. nov., Т. ehrenbergii sp. nov., Т.

mosis sp. nov., Webera sacra sp. nov., and W. sinaitica sp, nov.].

LORIA М. & HERRNSTADT І., 1980 — Moss capsules as food of the harvester ant,

Messor. The Bryologist 83: 524-525.

[Reports the feeding on moss capsules of Aloina aloides, Bryum bicolor and Crossidium

crassinerve by harvester ants in the northern Negev in winter 1980, when no other food

sources were available].

MATOUSCHEK F., 1905 — Muscineae. In: PENTHER A. & ZEDERBAUER E.:

Ergebnisse einer naturwissenschaftlichen Reise zum Erdschias Dagh, S.

385-387. Annalen des naturhistorischen Museums in Wien 20: 359-464.

[List of 41 mosses with collection data].

McADAM S. V., 1981 — The identity of a gathering of Eurhynchium speciosum (Brid.)

Jur. from Turkey. Journal of Bryology 11: 560-562.

[Note on the morphological and cytological characters of E. speciosum and E. swartzii,

to clarify the taxonomic affinities].

MENZEL M., 1992 — Nomenklatorische Notizen zu Eddys “Handbook of Malesian

Mosses” (Band 2). Willdenowia 22: 197-199.

[Antedates and replaces the genus Pseudosymblepharis by Chionoloma; includes С.

socotrana comb. nov. from Socotra (cf. also Long 1986 and Zander 1993)].

MIES B., 1994 — Checkliste der Gefäßpflanzen, Moose und Flechten und botanische

Bibliographie der Insel Sokotra und des Sokotrinischen Archipels (Jemen,

Indischer Ozean). Senckenbergiana Biologica 74: 213-258.

[Checkist of the known hepatic and moss taxa with notes on synonymy and former

reports in literature; extensive literature list].

MIFTAH N., 1969 — Botany. Cryptogams. Teheran University Publications No. 754.

In Farsi.

[Not seen].

Source : MNHN. Paris

30 H. KURSCHNER

MITTEN W., 1888 — Muscineae. Jn: BALFOUR J. B.: The botany of Socotra, pp.

330-336. Transactions of the Royal Society of Edinburgh 31: 1-446.

[List of 11 moss and five hepatic taxa with collection data and notes on the distribution,

based on the collections of the Balfour Expedition (I. B. Balfour, Lt. Cockburn, A.

Scott 1879-1880, B.C.S.); includes the descriptions of Fabronia socotrana sp. nov.,

Fimbriaria pusilla sp. nov., Frullania socotrana sp. nov., Schlotheimia balfourii sp. nov.,

Symblepharis socotrana sp. nov., Weisia punctulata sp. nov., and W. socotrana sp. nov].

MULLER C., 1901 — Genera Muscorum frondosorum. Leipzig, 474 S.

[Description of acrocarpous moss genera; includes four mosses from Socotra, collected

by С. Schweinfurth (Barbula riebecki sp. nov., B. schweinfurthiana sp. nov., Campylopus

polytrichoides, and Edentella schweinfurthi sp. nov., ef. also Long 1986) and a note on

Trichostomum ehrenbergii from Sinai].

NACHMONY S., 1961 — New additions to the moss flora of Palestine and the

neighbouring countries, Bulletin of the Research Council of Israel, Section D

10: 352-355.

[List of 28 mosses, five being new to the area].

NICHOLSON W. E., 1920 — Mosses from the Caspian and Black Sea regions. The

Bryologist 23: 90-91.

[List of two hepatic and 15 moss taxa from the south shore of the Caspian Sea and the

Elburz Mts. (without exact locality data), based on the collections of W. J. Parsons 1917].

NYHOLM E. & WIGH К., 1973 — Cytotaxonomical studies in some Turkish mosses.

Lindbergia 2; 105-113.

[Reports, partly for the first time, mitotic chromosome numbers of 16 mosses].

ОСНІ Н. & KURSCHNER H., 1988 — Bryum nanoapiculatum (Bryaceae, Musci), a

species new to the moss flora of the Arabian Peninsula. — Studies in Arabian

bryophytes 13. Nova Hedwigia 47: 359-361.

[One new species from northern Yemen described and illustrated].

OZTURK М. & GOKLER L, 1988 — Ecology of West-Anatolian liverworts.

Bryological Times 47: 1-3.

[General account on the bryofloristic knowledge of western Turkey (Aegean region),

with notes on some bryophyte communities, their distribution and ecology].

PAROLLY G., 1995 — Die Steinschuttfluren (Heldreichietea) des Westlichen und

Mittleren Taurus (Türkei). Pflanzensoziologische, floristische und ökologi-

sche Untersuchungen. Dissertationes Botanicae 247: 1-374.

[Phytosociological study of the scree formations of the Taurus Mountains; includes a

few mosses (Bryum provinciale, В. rutilans, Tortella tortuosa, Tortula norvegica var.

norvegica, Т. princeps subsp. echinata in the phytosociological relevés].

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 31

POTIER DE LA VARDE R., 1956 — Contribution ä la flore bryologique d'Israel.

Revue bryologique et lichenologique 25: 120-123.

[List of five Fissidens species with collection data, collected by F. Bilewsky, with the

description and illustration of F. bilewskyi sp. nov.].

PROSKAUER J., 1953 — On a collection of liverworts from Israel. Palestine Journal

of Botany, Jerusalem Series 6 (2): 123-125.

[List of one anthocerote and 14 hepatic taxa with collection data, based on the

collections of D. Zohary 1949-1951].

PURSELL R. A. & KURSCHNER H., 1987 — Fissidens arabicus (Fissidentaceae), a

new species from the Asir Mountains (Saudi Arabia). Studies in Arabian

bryophytes 6. Nova Hedwigia 44: 101-103.

[One new species described and illustrated, with notes on the floristic affinities and

ecology].

RABINOWITZ-SERENI D., 1931 — Elenco dei Muschi e delle Epatiche Palestinesi.

Annali di Botanica 19: 333-339.

[List of six hepatic and 19 moss taxa, mainly collected in Israel 1926-1927].

RAEYMAEKERS G., 1983 — Philonotis rigida Brid. in Europe. Lindbergia 9: 29-33.

[Floristic study with locality data from Lebanon and Turkey].

REIMERS H., 1927 — Die von Prof. Dr. K. Krause in Kleinasien besonders im

Pontus, 1926 gesammelten Leber- und Laubmoose. Notizblat des Botanischen

Gartens und Museums zu Berlin-Dahlem 10: 27-42.

[List of one anthocerote, six hepatic and 29 moss taxa with collection data, some

taxonomic notes and remarks on the world-wide distribution].

REIMERS H., 1957 — Einige bemerkenswerte Moose des óstlichen Mediterrangebie-

tes. П. Bryum splachnoides (Нагу.) С. М. in Vorderasien. Willdenowia 1:

696-701.

[Reports Bryum splachnoides from Lebanon (collected by J. & F. Bornmiiller 1910) and

Israel (collected by F. Bilewsky 1953), with notes on the floristic affinities and

distribution].

RENAULD Е. & CARDOT J., 1894 — Mousses nouvelles de ГНегбіег Boissier.

Bulletin de l'Herbier Boissier, série 1, 2: 32-33.

[Includes the descriptions of Grimmia anodon var. sinaitica var. nov. from Sinai, and

Polytrichum autrani sp. nov from Lebanon].

ROBINSON H. & GODFREY R. K., 1960 — Contribution to the bryophyte flora

of Turkey. Revue bryologique et lichénologique 29: 244-253 (publ. march

1961).

[List of 14 hepatic and 96 moss taxa with collection data, based on the collections of

R. K. Godfrey 1952].

ROS ESPIN R. M., 1987 — Riella cossoniana Trab., nueva Hepática para la flora

Europea. Cryptogamie, Bryologie-Lichénologie 8: 227-233.

[Study of R. cossoniana with description, illustrations, and notes on taxonomy, ecology

and distribution; includes a distribution map and collection data from Israel (cf. also

Lipkin & Proctor 1975)].

Source : MNHN. Paris

32 H. KÜRSCHNER

RUNGBY S., 1959 — A contribution to the bryophyte flora of the Near and Middle

East, Botaniska Notiser 112: 80-84.

[List of three hepatic and 40 moss taxa based on the author's collections in early 1956

in Lebanon, Iraq, Israel and Syria; includes the description and illustrations of Barbula

bistrata sp. nov].

SALMON E. S., 1899 — А new moss from Afghanistan. Journal of Botany, British and

Foreign 37: 241-242.

[Tortula (Pterygoneurum) media sp. nov. described and illustrated, with notes on

floristic affinities]

SALMON E. S., 1900 — Bryological Notes. 1. Cinclidotus pachyloma sp. nov. Revue

bryologique et lichénologique 27: 59-65.

[One new species from Syria (ex Herb. Post) described and illustrated].

SAVULESCU T., 1928 — Contributions à la flore d'Arabie. Bulletin de l'Académie des

Sciences de Roumanie 11 : 14-24.

[Reports Plagiochasma rupestre from Yemen (Jabal Shibam, Menaka), collected by G.

Botez 1911].

SCHIFFNER V., 1896 — Ueber die von Sintenis in Türkisch-Armenien gesammelten

Kryptogamen. Österreichische Botanische Zeitschrift 46; 274-278.

List of two hepatic and 12 moss taxa with collection data, based on the collections of

P. Sintenis 1894; includes the descriptions of Amblystegium fallax var. crassicostatum

var. nov., Philonotis calcarea var. orthophylla var. nov., and Pseudoleskea atrovirens

var. revoluta var. nov.].

SCHIFFNER V., 1897 — Musci Bornmülleriani. Ein Beitrag zur Cryptogamenflora

des Orients. Österreichische Botanische Zeitschrift 47: 125-132.

List of four hepatic and 35 moss taxa with collection data, based on the collections of

1. Bornmüller from Iran and Turkey (“Нег Persico-turcicum 1892-1893”, “Plantae

Anatoliae orientalis 1889, 1890”); includes the descriptions of Bryum pseudotriquetrum

var. bornmülleri var. nov., Grimmia orbicularis yar. persica var. nov., and Tortula

bornmülleri sp. nov.].

SCHIFFNER V., 1901 — Einige Materialien zur Moosflora des Orients. Osterreichi-

sche Botanische Zeitschrift 51; 156-161.

[List of three hepatic and 16 moss taxa with collection data, based on the collections

of J. Bornmüller from Turkey ("Iter Anatolicum tertium 1899"), Iran (“Iter Persico-

tureicum 1892-1893”) and Syria (“Iter Syriacum 1897”); includes the description of

Philonotis calcarea var. seriatifolia var. nov.].

SCHIFFNER V., 1908 — Beiträge zur Kenntnis der Bryophyten von Persien und

Lydien. Österreichische Botanische Zeitschrift 58: 225-231, 304-318, 341-349

(mit 3 Tafeln).

List of one anthocerote, 16 hepatic and 82 moss taxa with collection data, based on

the collections of J, Bornmüller from Iran (“Iter Persicum alterum 1902") and Turkey

("Lydiae et Cariae 1906"), J. A. Knapp 1884 from Iran, and T. Strauss 1903, 1904 from

Iran; includes the descriptions and illustrations (three plates) of Antitrichia breidleriana

sp. nov., Fimbriaria silachorensis sp. nov., Grimmia caespiticia var. bornmiillerorum var.

nov., Hypnum decipiens var. napaeiforme var. nov., Leskea laxiramea sp. nov.

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 33

Philonotis seriata var. persica var. nov., Reboulia hemisphaerica var. microspora var.

nov., Timmiella grosseserrata sp. nov., Tortula азота sp. nov., Т. demawendica sp.

nov., and Webera pentasticha sp. nov.].

SCHIFFNER V., 1909 — Hepaticae. In: HANDEL-MAZZETTI Н. v.: Ergebnisse

einer botanischen Reise in das Pontische Randgebirge im Sandschak Trape-

zunt. Annalen des naturhistorischen Museums in Wien 23: 133-141 (mit 2

Tafeln).

[List of 50 hepatic taxa with collection data and details of localities; includes the

descriptions and illustrations of Nardia handelii sp. nov., N. handelii var. flaccida var.

nov., N. lignicola sp. nov., and N. subtilissima sp. nov.).

SCHIFFNER V., 1910 — Bryologische Fragmente. LXV. Leskea laxiramea. Oster-

reichische Botanische Zeitschrift 60: 436.

[Reports a new locality for this species, based on a collection of F. Bruns 1909 in

Iran].

SCHIFFNER V., 1913 — Bryophyta aus Mesopotamien und Kurdistan, Syrien,

Rhodos, Mytilini und Prinkipo. Gesammelt von Dr. Heinrich Frh. v.

Handel-Mazzetti (Wissenschaftliche Ergebnisse der Expedition nach Meso-

potamien, 1910). Annalen des naturhistorischen Museums in Wien 27: 472-

504.

[List of 113 moss and 12 hepatic taxa with collection data and some taxonomic notes,

based on the collections of H. v. Handel-Mazzetti 1910; includes the descriptions and

illustrations of Amblystegium kurdicum sp. nov., Anoectangium handelii sp. nov.

Fontinalis mesopotamica sp. nov., Funaria handelii sp. nov., Grimmia mesopotamica sp.

nov., б. singarensis sp. nov., G. subcaespiticia sp. nov., Mniobrywn latifolium sp. nov.,

Orthotrichum cupulatum var. bistratosum var. nov., O. rupestre var. kurdicum var. nov.,

Prerygoneurum cavifolium var. muticum var. nov., Riccia frostii var. crystallinoides var.

nov, R. frostii var, major var. nov., Tortula aestiva var. brevifolia var. nov., T.

brevissima sp. nov., and T. handelii sp. nov.].

SCHIFFNER V., 1939 — Hepaticae. In: RECHINGER K. H.: Ergebnisse einer

botanischen Reise nach dem Iran, 1937, 5. 533. Annalen des naturhistorischen

Museums in Wien 50: 410-536.

[List of four species with collection data, based on the collections of K. H. Rechin-

ger].

SEÇMEN Ó., LEBLEBICI E. & GÖKLER I., 1989 — A new liverwort for Turkey:

Ricciocarpus natans (L.) Corda. Doga, Türk Botanik Dergisi Seri С 13:

311-313. In Turkish, with English summary.

[Reports a new record with collection data, illustration and notes on taxonomy].

SHIRZADIAN 5., 1989 — Additions to the moss flora of Iran. Yushania 6: 17-20.

[Adds nine moss taxa to the known flora].

SHIRZADIAN S. & KUMAR S. S., 1994 — A report on the moss flora of Northern

Iran. Iranian Journal of Botany 6: 177-184.

[List of 29 moss taxa with collection data and notes of the world-wide distribution, 24

(not 29 as mentioned by the authors) being new to the flora].

STEPHANI F., 1894 — Hepaticarum species novae V. Hedwigia 33: 1-10.

[Includes the description of Fimbriaria persica sp. nov., based on the collection of C.

Haussknecht from Iran].

Source : MNHN. Paris

34 H. KURSCHNER

STORMER P., 1963 — Iranian plants collected by Per Wendelbo in 1959. VI. Mosses

(Musci). Arbok for Universitetet i Bergen. Mat.-Naturv. Serie No. 11: 3-34.

[List of 89 species with specimen data, notes on previous records and distribution;

includes a preliminary list of all mosses known from Iran with synonyms and the

description and illustration of Eurhynchium speciosum var. angustifolium var. nov.].

TCHIHATCHEFF Р. de, 1860 — Asie Mineure Vol. ПІ, Tome 2, Botanique. Paris,

676 p.

Musci and Hepaticae on pp. 658-661; list of 32 moss and four hepatic taxa with

collection data, based on the author's collections in Turkey].

THERIOT I., DIXON Н. N. & BUCH H., 1934 — Bryophyta nova (17-25). Annales

Bryologici 7: 157-162.

Includes the description of Splachnobryum arabicum sp. nov., collected by J. Fernandez

1925 in Oman].

TOWNSEND C. C., 1966a — Bryophytes from Azraq National Park, Jordan.

Transactions of the British Bryological Society 5: 136-141.

List of one hepatic and 11 moss taxa with collection data, based on the author's

collections 1965; includes one find of J. B. Gillett 1963 from Wadi Rum and some

remarks on the floristic affinities of selected taxa].

TOWNSEND C. C., 1966b — Mosses from Iran and Afghanistan. Transactions of the

British Bryological Society 5: 131-135.

List of 15 moss taxa from Afghanistan and 18 moss taxa from Iran with collection

data, based on the collections of P. Furse 1964; includes an illustration of Tortula

cabulica (cf. Froehlich 1955)].

TOWNSEND C. C., 1969a — Bryophytes from Socotra. Transactions of the British

Bryological Society 5: 790-796.

[List of nine moss taxa with collection data, descriptions and illustrations, based on the

collections of A. R. Smith 1967 (Middle East Command Expedition); includes the

decription and illustration of Tortella smithii sp. nov.].

TOWNSEND C. C., 1969b — Contributions to the bryophyte flora of Turkey VII.

Notes from the Royal Botanic Garden Edinburgh 29: 233-234.

[List of 19 moss taxa with collection data based on the collections of C. Barclay 1968].

TOWNSEND C. C., 1977 — Bryophytes from some Greek Islands. Revue bryologique

et lichénologique 43: 389-396.

[Includes a short note on the taxonomic relationships of Gymnostomum mosis, collected

by M. Grierson on the Sinai-Peninsula].

TOWNSEND С. С., 1978a — Notes on mosses from Ceylon and India, 1-5. Journal

of Bryology 10: 125-137.

[Includes some taxonomic notes on Tortula schmidii (C. M.) Broth. (Syn. T. fragilis

Tayl.) and collection data from Yemen].

TOWNSEND C. C., 1978b — Review of A. J. E. Smith “The moss flora of Britain and

Ireland”. Excerpta Botanica, Sectio A, 31: 398-400.

[Includes a note on a record of Funaria pulchella from Saudi Arabia, without locality

data].

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 35

TOWNSEND C. C., 1979 — A new combination in African Tortula. Journal of

Bryology 10: 576.

[Tortula porphyreoneura comb. nov. is recorded from Yemen (without exact locality

data)].

TOWNSEND C. C., 1989 — Grimmia (Musci): a variety new to the Lebanon and a

new species from Greece. In: TAN K. (ed.): The Davis and Hedge Festschrift.

Edinburgh, pp. 45-52.

[Description and illustration of Grimmia crassifolia var. cucullata based on the author's

collection from Lebanon with notes on the floristic affinities].

TREGUBOV A. & TREGUBOV V., 1969/1970 — Les Bryophytes des foréts

caspiennes du Nord de l'Iran. Bulletin de la Faculté Forestiére de l'Université

de Téhéran 17: 1-30.

[List of 11 hepatic and 73 moss taxa with collection data, based on the authors’

collections, partly identified by J. Froehlich (Wien) and М. 5. Grom (Ljubljana)].

TUTEL B., 1973 — Cryptogamic flora of Turkey. Verhandlungen des Internationalen

Symposiums über Abies equi-trojani und die türkische Flora. Istanbul,

pp. 233-238.

[Some general remarks on the state of floristic knowledge within the cryptogamic flora

of Turkey; includes some notes on bryophytes).

ай

VANA J., 1982 — The hepatics of well- and poorly- known areas in temperate

Europe and Asia. Beiheft zur Nova Hedwigia 71: 341-350.

[General account on the floristic knowledge (Afghanistan, Iran, Turkey); includes an

useful literature list].

VOLK O. H., 1988 Riccia crenatodentata (Marchantiales), sp. nov. aus Arabien.

Studien über Bryophyten aus Arabien 11. Nova Hedwigia 46: 27-35.

[One new species described and illustrated; includes collection data from Oman and

Saudi Arabia and a list of three further Riccia species].

VONDRACEK M., 1962 — New bryophytes for the Iraq flora. Bulletin of the Iraq

Natural History Institute 2: 9.

[Reports 26 moss taxa, based on a collection of E. Hadaë (without locality data)].

VONDRACEK M., 1965 — Some new mosses from Iraq, collected by В. Надаё.

Bulletin de la Société des Amis des Sciences et de Lettres de Рогпап, série D

6: 117-122,

[One new genus, Steppomitra gen. nov., two new species, Steppomitra hadacii sp. nov.,

Syntrichia hadacii sp. nov., and three new varieties, Grimmia ovalis var. simplex var.

nov., Physcomitrella patens var. marginata var. nov., Syntrichia handelii var. pseudo-

desertorum var. nov. described and illustrated].

Source : MNHN. Paris

36 H. KÜRSCHNER

WAGENITZ G., 1962 — Zur Kenntnis der Flora und Vegetation Anatoliens

(Ergebnisse einer Reise im Herbst 1957). Willdenowia 3: 221-288.

[List of flowering plants with some general remarks on the flora and vegetation of

Turkey; includes four hepatic and 10 moss taxa with collection data, identified by H.

Reimers].

WALTHER K., 1967 — Beiträge zur Moosflora Westanatoliens I. Mitteilungen aus

dem Staatsinstitut für Allgemeine Botanik in Hamburg 12: 129-186.

[List of two anthocerote, 55 hepatic and 311 moss taxa with collection data, based on

the author’s collections in Turkey 1964 and 1965, and collections of C. v. Regel; reports

131 taxa as new to the flora].

WALTHER K. & LEBLEBICI E., 1969 — Die Moosvegetation des Karagöl-Gebietes

im Yamanlar Dag nérdlich Izmir. Monographs of the Faculty of Science, Ege

University No. 10: 3-48.

[Phytosociological study of the epilithic and epiphytic bryophyte vegetation (description

of 20 associations) with notes on syntaxonomy, community structures, ecology and

habitat sites; includes a list of one anthocerote, 10 hepatic and 135 moss taxa with

collection data, 15 being new to the flora of Turkey].

WALTHER K., 1970 — Beiträge zur Moosflora Westanatoliens II. Mitteilungen aus

dem Staatsinstitut für Allgemeine Botanik in Hamburg 13: 167-180.

[List of 14 hepatic and 121 moss taxa with collection data, based on the author's

collections in Turkey 1967-1969; adds 54 taxa to the known flora].

WALTHER K., 1975 — Zur Moosvegetation der Liquidambar-Wälder Südwest-

Anatoliens. Phytocoenologia 2: 13-18.

[Phytosociological study of the epiphytic bryophyte vegetation (description of two

associations) with notes on syntaxonomy, community structures, ecology and habitat

sites],

WALTHER K., 1979 — Die epiphytischen Moosgesellschaften des Nif Dag bei Izmir,

Westanatolien. Documents Phytosociologiques 4: 943-950.

[Phytosociological study of the epiphytic bryophyte vegetation (description of four

associations, two of them described as new) with notes on syntaxonomy, community

structures, ecology and habitat sites].

WASHBOURN R. & JONES R. F., 1937 — Percy Sladen Expedition to Lake Huleh,

Palestine. Proceedings of the Linnean Society of London 1937: 97-99.

[Includes a find of Riella helicophylla, collected by E. M. Blackwell in 1923].

WEITZ S. & HEYN C. C., 1981 — Intra-specific differentiation within the

cosmopolitan moss species Funaria hygrometrica Hedw. The Bryologist 84:

315-334.

[Morphological and cytological study of various populations of F. hygrometrica from

different regions, showing a significant inter-population and inter-regional differentia-

tion; includes collection data from Israel].

WETTSTEIN R. v., 1889 — Beitrag zur Flora des Orientes. Bearbeitung der von Dr.

A. Heider im Jahre 1885 in Pisidien und Pamphylien gesammelten Pflanzen.

Sitzungsberichte der Akademie der Wissenschaften in Wien 98: 348-398.

[Muscineae on p. 364; reports two hepatic and three moss taxa with collection data].

Source : MNHN Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 37

WHITEHOUSE Н. І. К. & CRUNDWELL А. C., 1991 — Gymnostomum calcareum

Nees & Hornsch. and allied plants in Europe, North Africa and the Middle

East. Journal of Bryology 16: 561-579.

[Taxonomic study of Gymnostomum calcareum, G. mosis, and G. viridulum, on the basis

of their morphology and from culture studies; includes descriptions, illustrations, notes

on synonymy, collection data from Afghanistan, Iran, Iraq, Israel, Jordan, Oman and

Sinai, and distribution maps].

WHITEHOUSE H. L. K. & CRUNDWELL A. C., 1992 — Gymnostomum calcareum

Nees & Hornsch. and G. viridulum Brid. in Europe, North Africa and the

Middle East. Bulletin of the British Bryological Society 59: 35-50.

[List of locality data of G. calcareum and G. viridulum, which were examined and

confirmed by the authors’ during the course of their work (cf. Whitehouse & Crundwell

1991); includes locality data from Afghanistan, Iran, Iraq, Israel, Jordan, Lebanon,

Sinai, Syria, Turkey, and Yemen).

YAYINTAS A. & IWATSUKI Z., 1988 — Some moss records from western Turkey.

Hikobia 10: 209-213.

[List of 87 moss taxa in tabular form, with notes on altitudinal distribution and the

substrata].

YAYINTAS A., 1993 — New moss record for Türkiye, Plagiothecium curvifolium

Schlieph. ex Limpr. (Plagiotheciaceae). Journal of the Faculty of Science, Ege

University, Series B 15: 21-23.

[Reports a new record with illustration, description and notes on habitat and

distribution].

YAYINTAS A. & TONGUG Ó., 1993 — Platydictya confervoides (Brid.) Crum, a new

moss record for Türkiye. Journal of the Faculty of Science, Ege University,

Series B 15: 17-20.

[Reports a new record with illustration, description and notes on habitat and

distribution].

YAYINTAS A., 1994 — A new moss record for Tiirkiye, Plagiothecium denticulatum

(Hedw.) B.S.G. var. obtusifolium (Turn.) Moore (Plagiotheciacee). Journal of

the Faculty of Science, Ege University, Series B 16: 19-21.

[Reports a new record with illustration, description and notes on habitat and

distribution].

YAYINTAS A., AYSEL V., GÜNER Н. & TONGUC Ó., 1994 — The moss flora

of Bozcaada. Turkish Journal of Botany 18: 29-32. In Turkish, with English

summary.

[List of 36 taxa, eight being new for square В 6].

YAYINTAS А. & TONGUG Ӧ., 19944 — A new moss record for Turkey,

Plagiothecium succulentum (Wils.) Lindb. (Plagiotheciaceae). Turkish Journal

of Botany 18: 517-518.

[Reports a new record with illustration, description and notes on habitat and

distribution].

Source : MNHN, Paris

38 H. KURSCHNER

YAYINTAS A. & TONGUÇ Ó., 1994b — New moss records from Thrace for A 1.

Journal of the Faculty of Science, Ege University, Series В 16: 51-61.

{List of 54 taxa with collection data, new to turkish Thrace (square A 1)].

ZANDER R. H., 1993 -- Genera of the Pottiaceae: mosses of harsh environments.

Bulletin of the Buffalo Society of Natural Sciences 32: 1-378.

[Taxonomic treatment of the Pottiaceae with keys to the genera, descriptions, excellent

illustrations by P.M. Eckel, and notes on evolutionary relationships, taxonomy,

habitats and distribution; includes several new names, statuses and new combinations,

e.g. Crossidium woodii comb. nov. (Pseudaloina woodii Delg., cf. Delgadillo 1982),

Syntrichia virescens var. iranica comb. nov. (Tortula virescens var. iranica Kramer, cf.

Kramer 1980) or Weissia artocosana nom. nov. for Symblepharis socotrana Mitt. =

Pseudosymblepharis socotrana (Mitt.) Ther., cf. Mitten 1888].

GUIDE TO NATIONAL AND REGIONAL LITERATURE

Afghanistan

HEPATICAE — Bischler (1978, 1979), Bischler-Causse (1989), Frey (1974), Frey &

Kiirschner (1991b), Froehlich (1955, 1964), Gilli (1971, 1975, 1977), Griffith (1849),

Hattori (1978a, b), Henderson & Prentice (1963), Herzog (1938), Kitagawa (1964),

Vana (1982).

MUSCI — Agnew & Townsend (1970), Aitchison (1881), Bizot (1962), Cano, Guerra

& Ros (1993), Crundwell & Nyholm (1977), Diill (1982), Frey (1969, 1972, 1986), Frey

& Kiirschner (1991b), Froehlich (1955, 1964), Gilli (1969, 1971, 1975, 1977), Griffith

(1849), Henderson & Prentice (1963), Herzog (1938), Kilbertus & Bizot (1971),

Kitagawa (1964), Koponen (1980, 1982), Kramer (1978, 1980), Kürschner (1996b),

Salmon (1899), Townsend (1966b), Whitehouse & Crundwell (1991, 1992).

Bahrain

No records of bryophytes up to now (cf. Frey & Kiirschner 1991b).

Egypt/Sinai

HEPATICAE — Frey & Kiirschner (1991b), Hart (1891), Jovet-Ast & Bischler (1970).

MUSCI — Agnew & Townsend (1970), Bilewsky (1974), Decaisne (1834), Delgadillo

(1975), El-Oglah, Frey & Kürschner (1988b), Frey (1986), Frey & Kürschner (1988c,

1991b, 1993b), Geheeb (1903/1904), Hart (1891), Hilpert (1933), Imam & Ghabbour

(1972), Kramer (1980), Lorentz (1868), Müller (1901), Renauld & Cardot (1894),

Townsend (1977), Whitehouse & Crundwell (1991, 1992).

Iran

HEPATICAE — Arnell (1963), Bischler (1979), Bischler-Causse (1989), Boissier &

Buhse (1860), Bornmüller (1908b, 1911, 1915), Fehlner (1885), Frey (1974), Frey,

Source : MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 39

Grolle & Kürschner (1991), Frey & Kürschner (1977, 1979, 1983a, 1991b), Froehlich

(1950), Guerke (1978), Jovet-Ast (1981, 1983, 1989), Kürschner (1996b), Miftah (1969),

Nicholson (1920), Schiffner (1897, 1901, 1908, 1939), Stephani (1894), Tregubov &

Tregubov (1969/1970), Váña (1982).

MUSCI - Agnew & Townsend (1970), Baumgartner (1939), Boissier & Buhse (1860),

Bornmiiller (1908b, 1910, 1911, 1915), Brotherus (1906), Cano, Guerra & Ros (1993),

Crundwell & Nyholm (1977), Düll (1982), Fakhr (1977), Fehlner (1883, 1885), Frey

(1986), Frey & Kiirschner (1977, 1979, 1983a, 1991b, 1993b), Frey & Probst (1973,

1974a, b), Froehlich (1950, 1953), Gilli (1941), Jovet-Ast (1960), Juratzka & Milde

(1870), Koponen (1980, 1993), Kramer (1978, 1980), Kiirschner (1996b), Miftah (1969),

Nicholson (1920), Schiffner (1897, 1901, 1908, 1910), Shirzadian (1989), Shirzadian &

Kumar (1994), Stormer (1963), Townsend (1966b), Tregubov & Tregubov (1969/1970),

Whitehouse & Crundwell (1991, 1992).

Iraq

HEPATICAE — Froehlich (1959), Handel-Mazzetti (1914), Long (1979), Rungby

(1959), Schiffner (1913).

MUSCI — Agnew (1973), Agnew & Townsend (1970), Agnew & Vondrácek (1975),

Cano, Guerra & Ros (1993), Frey (1986), Frey & Kiirschner (1991b, 1993b), Froehlich

(1959), Handel-Mazzetti (1914), Koponen (1980, 1982), Kramer (1980), Long (1979),

Rungby (1959), Schiffner (1913), Vondraéek (1962, 1965), Whitehouse & Crundwell

(1991, 1992).

Israel (incl. Cis-Jordan, Golan)

HEPATICAE — Arnell (1957), Barbey & Barbey (1882), Baum & Jovet-Ast (1962),

Bischler-Causse (1989), Brullo, Privitera & Puglisi (1991), Frey, Herrnstadt &

Kürschner (1990b), Frey & Kürschner (1990, 1991b-e, 1992, 1995b), Hart (1891),

Herrnstadt (1992), Jovet-Ast (1961), Jovet-Ast & Bischler (1966), Jovet-Ast, Bischler &

Baudoin (1976), Jovet-Ast, Bischler & Baum (1965), Lipkin & Proctor (1975),

Proskauer (1953/1954), Rabinowitz-Sereni (1931), Ros Espin (1987), Rungby (1959),

Washbourn & Jones (1937).

MUSCI — Barbey & Barbey (1882), Bilewsky (1959, 1963, 1965, 1970, 1974, 1977),

Bilewsky & Nachmony (1955), Bizot (1954, 1956), Brullo, Privitera & Puglisi (1991),

Cano, Guerra & Ros (1993), Crosby (1979), Danin & Barbour (1982), Delgadillo

(1975), Dinola, Mayer & Heyn (1983), Frey (1986), Frey, Herrnstadt & Kürschner

(1990a, b), Егеу & Kürschner (1990, 1991а-е, 1992, 1993b, 1995b), Hart (1891),

Herrnstadt (1992), Herrnstadt & Heyn (1987, 1993), Herrnstadt, Heyn, Ben-Sasson &

Crosby (1982), Herrnstadt, Heyn & Crosby (1980, 1991), Koponen (1980, 1993),

Kramer (1980), Loria & Herrnstadt (1980), Nachmony (1961), Potier de la Varde

(1956), Rabinowitz-Sereni (1931), Reimers (1957), Rungby (1959), Weitz & Heyn

(1981), Whitehouse & Crundwell (1991, 1992).

Jordan

HEPATICAE — Barbey & Barbey (1882), Bischler & Jovet-Ast (1975), El-Oglah, Frey

& Kirschner (1988a), Frey & Kiirschner (1983b, 1991b, d, е, 1992, 1994, 1995a, b),

Hart (1891), Townsend (1966a).

MUSCI — Barbey & Barbey (1882), Bilewsky (1959, 1970, 1974), Bilewsky &

Source : MNHN, Paris

40 H. KURSCHNER

Nachmony (1955), Cano, Guerra & Ros (1993), Delgadillo (1973, 1975), El-Oqlah,

Frey & Kürschner (1988a, b), El-Oglah & Lahham (1985), Frey (1986), Frey &

Kürschner (1983b, 1991a, b, d, e, 1992, 1993b, 1994, 1995a, b), Frey, Kürschner, Ros,

Guerra & Cano (1994), Hart (1891), Kramer (1980), Kürschner (1995, 1996b),

Nachmony (1961), Townsend (1966a), Whitehouse & Crundwell (1991, 1992).

Kuwait

HEPATICAE — Frey & Kürschner (1988a, 1991b).

MUSCI — El-Saadawi (1976, 1978, 1979a, b), Frey (1986), Frey & Kürschner (1988a,

d, 1991b, 1993b), Halwagy & El-Saadawi (1992), Kürschner (1996b).

Lebanon

HEPATICAE — Arzeni (1965), Barbey & Barbey (1882), Bischler-Causse (1989), Bizot

(1942), Bornmüller (1914), Charouk (1982), Frey & Kürschner (1991b), Geheeb (1902),

Henderson & Muirhead (1955-1958b), Kiirschner (1996b), Rungby (1959).

MUSCI — Arzeni (1965), Barbey & Barbey (1882), Bizot (1942, 1945, 1955, 1956),

Bizot, Gaume & Potier de la Varde (1952), Bornmiiller (1914), Brown (1937), Cano,

Guerra & Ros (1993), Charouk (1982), Delgadillo (1975), Frey (1986), Frey &

Kiirschner (1991b), Geheeb (1902), Henderson & Muirhead (1955-1958b), Koponen

(1980, 1993), Kramer (1978, 1980), Kiirschner (1996b), Lorentz (1868), Nachmony

(1961), Raeymaekers (1983), Reimers (1957), Renauld & Cardot (1894), Rungby

(1959), Townsend (1989), Whitehouse & Crundwell (1992).

Oman

HEPATICAE — Bischler (1978, 1979), Frey & Kürschner (1988a, d, 1991b), Jovet-Ast

(1986, 1989), Long (1987), Volk (1988).

MUSCI — Bruggeman-Nannenga (1987), Cano, Guerra & Ros (1993), Frey 1986),

Frey & Kürschner (1988a, d, 1991b, 1993b), Kürschner (1996b), Thériot, Dixon &

Buch (1934), Whitehouse & Crundwell (1991).

Qatar

No records of bryophytes up to now (cf. Frey & Kürschner 1991b).

Saudi Arabia

HEPATICAE — Frey, Grolle & Kürschner (1991), Frey & Kirschner (1982, 1987,

1988a, d, 1991b, 1993a), Jovet-Ast (1986), Kürschner (1984a, b, 1989, 1989[1991]),

Long (1987), Volk (1988).

MUSCI — Cano, Guerra & Ros (1993), Frey (1986), Frey & Kürschner (1982, 1984,

1987, 1988a, b, d, 1991b, 1993a, b), Frey, Kürschner, Ros, Guerra & Cano (1994),

Kiirschner (1984a, b, 1989, 1989[1991], 1996b), Pursell & Kürschner (1987), Townsend

(19785).

Syria

HEPATICAE — Barbey & Barbey (1882), Bischler-Causse (1989), Brown (1937), Frey

& Kiirschner (1991b), Handel-Mazzetti (1914), Henderson & Muirhead (1955-1958b),

Rungby (1959), Schiffner (1913).

Source - MNHN. Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 41

MUSCI — Agnew & Townsend (1970), Barbey & Barbey (1882), Bilewsky (1970, 1974,

1977), Bizot (1954, 1955, 1956), Bizot, Gaume & Potier de la Varde (1952), Brown

(1937), Cano, Guerra & Ros (1993), Decaisne (1834), Delgadillo (1975), Frey (1986),

Frey & Kiirschner (1991b, 1993b), Handel-Mazzetti (1914), Henderson & Muirhead

(1955-1958b), Koponen (1980), Kramer (1978, 1980), Kürschner (1996b), Lorentz

(1868), Nachmony (1961), Rungby (1959), Salmon (1900), Schiffner (1913), White-

house & Crundwell (1992).

Turkey

HEPATICAE — Bischler (1978, 1979), Bischler-Causse (1989), Bornmúller (1908a,

1909, 1931), Cetin (1988b, 1989e, g, 1993), Cetin & Yurdakulol (1986), Crundwell &

Nyholm (1979), Frey, Grolle & Kiirschner (1991), Frey & Kiirschner (1991b), Fritsch

(1899), Froehlich (1959), Gökler (1986, 1989, 1992), Gökler, Inoue & Öztürk

(1983/1984), Gökler & Öztürk (1986, 1987a, b, 1989, 1991), Gökler, Öztürk &

Kesergioglu (1985/1986), Handel-Mazzetti (1905), Hattori (1976), Henderson (1955-

1958a, b, 1959-1961a, b, 1963-1964), Henderson & Muirhead (1955-1958a), Henderson

& Prentice (1969), Jovet-Ast (1957), Jovet-Ast, Bischler & Baudoin (1976), Öztürk &

Gökler (1988), Reimers (1927), Robinson & Godfrey (1960), Schiffner (1896, 1897,

1901, 1908, 1909, 1913), Segmen, Leblebici & Gökler (1989), Tchihatcheff (1860), Tutel

(1973), Váňa (1982), Wagenitz (1962), Walther (1967, 1970, 1975, 1979), Walther &

Leblebici (1969).

MUSCI — Barbey (1890), Becker (1967), Bornmiiller (1908a, 1909, 1931), Cano,

Guerra & Ros (1993), Cetin (1988a-d, 1989a-g, 1990, 1991, 1993), Cetin & Yurdakulol

(1985, 1988), Crundwell, During & Koutstaal (1978), Crundwell & Nyholm (1977),

Czeczott (1939), Delgadillo (1975), Düll (1982), Frey 1986), Frey & Kiirschner (1991b),

Fritsch (1899), Froehlich (1959), Handel-Mazzetti (1909), Henderson (1955-1958a, b,

1959-1961а, b, 1963-1964), Henderson & Muirhead (1955-1958a), Henderson &

Prentice (1969), Ignatov (1995), Juratzka & Milde (1870), Koponen (1980, 1982, 1993),

Kramer (1978, 1980), Kiirschner (1996b), Matouschek (1905), McAdam (1981),

Nyholm & Wigh (1973), Parolly (1995), Raeymaekers (1983), Reimers (1927),

Robinson & Godfrey (1960), Schiffner (1896, 1897, 1901, 1908, 1913), Tchihatcheff

(1860), Townsend (1969b), Tutel (1973), Wagenitz (1962), Walther (1967, 1970, 1975,

1979), Walther & Leblebici (1969), Wettstein (1889), Whitehouse & Crundwell (1992),

Yayintas (1993, 1994), Yayintas, Aysel, Giiner & Tongug (1994), Yayintas & Iwatsuki

(1988), Yayintas & Tongug (1993, 1994a, b).

United Arab Emirates

НЕРАТІСАЕ — Frey & Kürschner (1988a, 4, 1991b), Long (1982, 1987).

MUSCI — Frey & Kiirschner (1988a, d, 1991b), Long (1982).

Yemen

HEPATICAE — Al-Gifri & Kürschner (1996), Al-Gifri, Kürschner & Mies (1995),

Balfour (1903), Bischler 1978, 1979), Bischler-Causse (1989), Frey, Grolle & Kürschner

(1991), Frey & Kürschner (1988a, d, 1991b), Jones (1969, 1985), Jovet-Ast (1970, 1986),

Kürschner (1989[1991], 1996a), Long (1987), Mies (1994), Savulescu (1928), Townsend

(1969).

MUSCI — Al-Gifri & Kürschner (1996), Al-Gifri, Kürschner & Mies (1995), Balfour

Source : MNHN. Paris

42 H. KURSCHNER

(1903), Bruggeman-Nannenga (1987), Cano, Guerra & Ros (1993), Crosby, Schultze-

Motel & Schultze-Motel (1983), Delgadillo (1982), Forsskal (1775), Frey (1986), Frey

& Kirschner (1988a, d, 1991b), Hepper (1977), Kürschner (1989[1991], 1996a), Long

(1986), Menzel (1992), Mies (1994), Mitten (1888), Müller (1901), Ochi & Kirschner

(1988), Townsend (1969, 1978a, 1979), Whitehouse & Crundwell (1992), Zander (1993).

INDEX OF COLLECTORS WITH REFERENCE TO AUTHOR(S) AND DATE

Afghanistan

Aitchison, J. E. T. (Aitchison 1881); Breckle, S.-W. (Frey 1969, 1972, 1974, Frey &

Kürschner 1991b, Kürschner 1996b); Frey, W. (Frey 1969, 1972, 1974, Frey &

Kürschner 1988a, 1991b, Kramer 1978, 1980, Kürschner 1996b); Furse, Р. (Townsend

1966b); Gilli, A. (Froehlich 1955, Gilli 1969, 1971, 1975, 1977); Hedge, І. C.

(Henderson & Prentice 1963, Kiirschner 1996b); Herzog, T. (Herzog 1938); Kilbertus,

G. (Kilbertus & Bizot 1971); Kitamura, S. (Kitagawa 1964); Probst, W. (Frey 1969,

1972, 1974, Frey & Kirschner 1988a, 1991b, Kramer 1978, 1980, Kiirschner 1996b);

Rechinger, K. H. (Froehlich 1964); Regel, C. v. (Bizot 1962); Volk, O. (Frey 1972, 1974,

Frey & Kiirschner 1991b, Kiirschner 1996b); Wendelbo, P. (Henderson & Prentice

1963, Kiirschner 1996b).

Egypt/Sinai

Bischler, H. (Jovet-Ast & Bischler 1970); Bové, N. (Decaisne 1834); Ehrenberg, G. C.

(Lorentz 1868); Frahm, J.-P. (Frey & Kiirschner 1991b); Frey, W. (Frey & Kiirschner

1988a, 1991b); Grierson, M. (Townsend 1977); Hart, H. C. (Hart 1891); Hasib, M.

(Imam & Ghabbour 1972); Jovet-Ast, S. (Jovet-Ast & Bischler 1970); Kneucker, A.

(El-Oglah, Frey & Kürschner 1988b, Geheeb 1903/1904); Kürschner, H. (Frey &

Kürschner 1988a, 1991b).

Iran

Agnew, S. (Kürschner 1996b); Boissier, E. (Boissier & Buhse 1860); Bornmüller, J.

(Bornmüller 1908b, Schiffner 1897, 1901, 1908); Bruns, F. (Bornmüller 1915, Schiffner

1910); Buhse, F. (Boissier & Buhse 1860); Frey, W. (Frey 1974, Frey & Kürschner 1977,

1979, 1983a, 1988a, 1991b, Kramer 1978, 1980, Kürschner 1996b); Furse, Р.

(Townsend 19665); Gilli, A. (Gilli 1941); Haussknecht, C. (Frey, Grolle & Kürschner

1991, Juratzka & Milde 1870, Kürschner 1996b, Stephani 1894); Heim, R. (Jovet-Ast

1960); Knapp, J. A. (Bornmüller 1910, Schiffner 1908); Koeltz, W. N. (Kürschner

1996b); Kramer, W. (Frey & Kürschner 1991b, Kramer 1978, 1980, Kürschner 1996b);

Kürschner, H. (Frey & Kürschner 1977, 1979, 1983a, 1988a, 1991b, Kramer 1978, 1980,

Kürschner 1996b); Léonard, J. (Jovet-Ast 1981); Merton, L. F. H. (Kürschner 1996b);

Olufsen, Lt. (Brotherus 1906); Parsons, W. J. (Nicholson 1920); Probst, W. (Frey &

Kürschner 1977, 1979, 1983a, 1991b, Frey & Probst 1973, 1974a, b, Kramer 1978,

1980); Rechinger, K. H. (Baumgartner 1939, Froehlich 1950, 1953, Schiffner 1939);

Shirzadian, S. (Shirzadian 1989, Shirzadian & Kumar 1994); Stapf, O. (Fehlner 1885,

Frey, Grolle & Kürschner 1991); Strauss, T. (Bornmüller 1911, Frey, Grolle &

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 43

Kiirschner 1991, Schiffner 1908); Tregubov, A. (Frey & Kürschner 1979, 1991b,

Tregubov & Tregubov 1969/1970); Tregubov, V. (Frey & Kürschner 1979, 1991b,

Tregubov & Tregubov 1969/1970); Wendelbo, P. (Arnell 1963, Stormer 1963).

Iraq

Agnew, S. (Agnew 1973, Agnew & Vondráček 1975); Evans, W. В. (Long 1979); Hadat,

E. (Agnew & Vondráček 1975, Vondráček 1962, 1965); Handel-Mazzetti, Н. v.

(Handel-Mazzetti 1914, Schiffner 1913); Rechinger, К. Н. (Froehlich 1959); Rungby, S.

(Rungby 1959).

Israel (incl. Cis-Jordan, Golan)

Baum, B. (Baum & Jovet-Ast 1962, Jovet-Ast, Bischler & Baum 1965); Bilewsky, F.

(Bilewsky 1959, 1965, 1971, 1974, 1977, Frey & Kürschner 1991b, Potier de la Varde

1956, Reimers 1957); Bischler, H. (Bischler & Jovet-Ast 1975, Jovet-Ast & Bischler

1966, Jovet-Ast, Bischler & Baum 1965); Bizot, M. (Bizot 1945, 1955); Blackwell, E. M.

(Washbourn & Jones 1937); Crosby, M. R. (Crosby 1979, Herrnstadt, Heyn,

Ben-Sasson & Crosby 1982, Herrnstadt, Heyn & Crosby 1980, 1991); Danin, A. (Danin

& Barbour 1982, Frey, Herrnstadt & Kiirschner 1990a, Herrnstadt, Heyn & Crosby

1991); Frey, W. (Frey, Herrnstadt & Kiirschner 1990a, b, Frey & Kiirschner 1990,

1991а-е, 1995b); Herrnstadt, I. (Frey, Herrnstadt & Kürschner 1990a, b, Herrnstadt

1992, Herrnstadt, Heyn, Ben-Sasson & Crosby 1982, Herrnstadt, Heyn & Crosby 1980,

1991); Jovet-Ast, S. (Baum & Jovet-Ast 1962, Bischler & Jovet-Ast 1975, Jovet-Ast &

Bischler 1966, Jovet-Ast, Bischler & Baum 1965); Kofler, L. (Jovet-Ast 1961);

Kiirschner, H. (Frey, Herrnstadt & Kiirschner 1990a, b, Frey & Kiirschner 1990,

1991a-e, 1995b); Kushnir, Т. (Bilewsky & Nachmony 1955); Nachmony, S. (Bilewsky &

Nachmony 1955, Nachmony 1961); Rabinowitz-Sereni, D. (Rabinowitz-Sereni 1931);

Rungby 5. (Rungby 1959); Zohary, D. (Bilewsky & Nachmony 1955, Proskauer 1953).

Jordan

El-Oglah, A. A. (El-Oglah, Frey & Kürschner 1988a, b, El-Oqlah & Lahham 1985,

Kiirschner 1996b); Frey, W. (El-Oqlah, Frey & Kürschner 1988a, b, Frey & Kürschner

1983b, 1988a, 1991a, b, 1992, 1993b, 1994, 1995a, b, Frey, Kürschner, Ros, Guerra &

Cano 1994, Kramer 1978, 1980); Gillet, J. B. (Townsend 1966a); Hart, H. C. (Hart

1891); Kürschner, Н. (El-Oglah, Frey & Kürschner 1988a, b, Frey & Kiirschner 1983b,

1988a, 1991a, b, 1992, 1993b, 1994, 1995a, b, Frey, Kürschner, Ros, Guerra & Cano

1994, Kramer 1978, 1980, Kürschner 1995, 1996b); Lahham, J. N. (El-Oglah &

Lahham 1985); Townsend, C. C. (Townsend 1966a).

Kuwait

Brown, С. (Kürschner 1996b); El-Saadawi, W. E. (El-Saadawai 1976, 1978, 1979a, b,

Halwagy & El-Saadawi 1992).

Lebanon

Arzeni, C. B. (Arzeni 1965, Kürschner 1996b); Barbey, C. (Barbey & Barbey 1882);

Barbey, W. (Barbey & Barbey 1882); Bizot, M. (Bizot 1942, 1945, 1955); Bornmüller,

Source : MNHN, Paris

= H. KÜRSCHNER

J. (Bornmüller 1914); Charouk, S. (Charouk 1982); Davis, P. H. (Henderson &

Muirhead 1955-1958b, Kürschner 1996b); George, C. (Arzeni 1965); Gombault, M.

(Bizot, Gaume & Potier de la Varde 1952); Rungby, S. (Rungby 1959); Townsend, C.

C. (Townsend 1989).

Oman

Fernandez, J. (Thériot, Dixon & Buch 1934); Frey, W. (Frey & Kürschner 1988a,

1991b, Jovet-Ast 1989, Kürschner 1996b); Gallagher, M. (Bruggeman-Nannenga 1987,

Frey & Kürschner 1988a, Long 1987); Kürschner, H. (Frey & Kürschner 1988a, 1991b,

Jovet-Ast 1989, Kürschner 1996b); Long, D. G. (Bruggeman-Nannenga 1987, Long

1987); Miller, А. С. (Bruggeman-Nannenga 1987, Long 1987); Radcliffe-Smith, А.

(Frey & Kürschner 1991b, Long 1987).

Saudi Arabia

Al-Shehri, A. (Kürschner 1996b); Baierle, H. U. (Frey & Kürschner 1988a, 1991b,

Kürschner 1996b); Collenette, S. (Frey, Grolle & Kürschner 1991, Kürschner 1989,

1996b, Long 1987); Frey, W. (Frey 1989, Frey, Grolle & Kürschner 1991, Frey &

Kürschner 1982, 1984, 1987, 1988a, b, 1991b, 1993a, Frey, Kürschner, Ros, Guerra &

Cano 1994, Kürschner 1996b, Pursell & Kürschner 1987, Volk 1988); König, P. (Frey

& Kürschner 1988a, 1991b, Kürschner 1996b); Kürschner, H. (Frey, Grolle &

Kürschner 1991, Frey & Kürschner 1982, 1984, 1987, 1988a, b, 1991b, 1993a, Frey,

Kürschner, Ros, Guerra & Cano 1994, Kürschner 1984a, b, 1989 [1991], 1996b, Pursell

& Kürschner 1987, Volk 1988).

Syria

Barbey, C. (Barbey & Barbey 1882); Barbey, W. (Barbey & Barbey 1882); Bizot, M.

(Bizot 1954); Bornmüller, J. (Bornmüller 1914, Schiffner 1901); Bové, N. (Decaisne

1834); Davis, P. H. (Henderson & Muirhead 1955-1958b); Ehrenberg, G. C. (Lorentz

1868); Evans, W. B. (Brown 1937); Frey, W. (Frey & Kürschner 1991b, Kürschner

1996b); Handel-Mazzetti, H. v. (Handel-Mazzetti 1914, Schiffner 1913); Kürschner, H.

(Frey & Kürschner 1991b, Kürschner 1996b); Pallary, M. (Bizot, Gaume & Potier de

la Varde 1952); Rungby, S. (Rungby 1959).

Turkey

Barclay, C. (Townsend 1969b); Becker, H. (Becker 1967); Bornmüller, J. (Bornmüller

1908a, 1909, 1931, Reimers 1957, Schiffner 1897, 1901, 1908); Çetin, B. (Çetin 1988a-d,

1989a-g, 1990, 1991, 1993); Crundwell, A. C. (Crundwell & Nyholm 1977, 1979,

Whitehouse & Crundwell 1991, 1992); Czeczott, H. (Czeczott 1939); Davis, P. H.

(Henderson 1955-1958a, b, 1959-1961a, Henderson & Muirhead 1955-1958a); Dodds, J.

(Henderson & Muirhead 1955-1958a); Forsyth-Major, C. F. (Barbey 1890); Godfrey, R.

K. (Robinson & Godfrey 1960); Gökler, I. (Gökler 1986, 1989, 1992, Gökler, Inoue &

Öztürk 1983/1984, Gökler & Öztürk 1986, 1987a, b, 1989, 1991, Gökler, Öztürk &

Keserçioglu 1985/1986, Kürschner 1996b, Seçmen, Leblebici & Gökler 1989); Handel-

Mazzetti, H. v. (Handel-Mazzetti 1909, Schiffner 1909, 1913); Haussknecht, C.

(Juratzka & Milde 1870, Ignatov 1995); Hedge, I. C. (Henderson 1959-1961a); Heider,

A. (Wettstein 1889); Hein, P. (Kürschner 1996b); Henderson, D. M. (Henderson

Source : MNHN, Paris

ANNOTATED BRYOLOGICAL LITERATURE OF SOUTHWEST ASIA 45

1963-1964); Krause, K. (Reimers 1927); Kiirschner, H. (Kiirschner 1996b); Lesse, H. de

(Jovet-Ast 1957); MeNeill, J. (Henderson 1955-1958b); Merton, L. F. H. (Henderson

1959-1961a, 1963-1964); Nemetz, J. (Fritsch 1899); Nyholm, E. (Crundwell & Nyholm

1977, 1979, Çetin 1988a, b); Parolly, С. (Parolly 1995, Kürschner 1996b); Penther, A.

(Handel-Mazzetti 1905, Matouschek 1905); Polunin, O. (Henderson 1955-1958);

Rechinger, К. Н. (Froehlich 1959); Regel, С. v. (Walther 1967); Sintenis, P. (Schiffner

1896); Stainton, J. D. A. (Henderson 1963-1964); Tchihatcheff, P. de (Tchihatcheff

1860); Tongug, Ö. (Yayintas, Aysel, Güner & Tongug 1994, Yayintas & Tonguç 1993,

1994a, b); Wagenitz, G. (Wagenitz 1962); Walther, K. (Frey, Grolle & Kiirschner 1991,

Frey & Kürschner 1991b, Walther 1967, 1970, 1975, 1979, Walther & Leblebici 1969);

Yayintas, A. (Yayintas 1993, 1994, Yayintas, Aysel, Güner & Tongug 1994, Yayintas

& Iwatsuki 1988, Yayintag & Tongug 1993, 1994a, b); Yurdakulol, E. (Getin &

Yurdakulol 1985, 1986, 1988); Zederbauer, E. (Handel-Mazzetti 1905, Matouschek

1905).

United Arab Emirates

Deil, U. (Frey & Kürschner 1988a, 1991b, Kürschner 1989 [1991]; Edmondson, J. (Frey

& Kürschner 1988a, Long 1987).

Yemen

Al-Gifri, A. N. (Al-Gifri & Kürschner 1996, Al-Gifri, Kürschner & Mies 1995,

Kürschner 1996a); Ammar, M. Y. (Kürschner 1996a); Balfour, I. B. (Balfour 1903,

Mitten 1888); Botez, С. (Savulescu-1928); Champagne, P. (Frey & Kürschner 1988a,

Ochi & Kürschner 1988); Cockburn, Lt. (Balfour 1903, Mitten 1888); Deil, U. (Frey &

Kürschner 1991b, Kürschner 1989 [1991]); Forsskal, Р. (Forsskäl 1775); Grierson, A.

(Al-Gifri & Kürschner 1996); Henchie, S. (Kürschner 1996a); Hepper, F. N. (Frey &

Kürschner 1988a, 1991b, Hepper 1977, Kürschner 1996a); King, R. (Frey & Kürschner

1988a, Long 1987); Kürschner, Н. (Al-Gifri & Kirschner 1996, Kürschner 1996a);

Long, D. G. (Bruggeman-Nannenga 1987, Frey & Kürschner 1988a, Jones 1985,

Kürschner 1996a, Long 1987); Mies, B. (Al-Gifri, Kürschner & Mies 1995); Miller, A.

G. (Bruggeman-Nannenga 1987, Frey & Kürschner 1988a, Jones 1985, Kürschner

1996a, Long 1987); Radcliffe-Smith, A. (Kürschner 1996a); Schweinfurth, G. (Al-Gifri,

Kürschner & Mies 1995, Frey, Grolle & Kürschner 1991, Frey & Kürschner 1991b,

Jovet-Ast 1970, Kürschner 1996a, Long 1986, Müller 1901); Scott, A. (Balfour 1903,

Mitten 1888); Smith, А. В. (Al-Gifri, Kürschner & Mies 1995, Frey, Grolle &

Kürschner 1991, Kürschner 1996a, Jones 1969, Townsend 1969a); Wood, J. R. I.

(Bruggeman-Nannenga 1987, Delgadillo 1982, Kürschner 1996a, b, Long 1987).

Acknowledgement — I am grateful to D. Lamy (Paris) for helpful bibliographical remarks and

valuable additions to improve this list of literature.

REFERENCES

DAVIS P. H., 1965 — Flora of Turkey and the East Aegean Islands, vol. 1. Edinburgh, 567 p.

FREY W. & KÜRSCHNER H., 1981 — The bryological literature of Southwest Asia. Journal

of the Hattori Botanical Laboratory 50: 217-229.

Source : MNHN. Paris

46 H. KURSCHNER

FREY W. & KURSCHNER H., 1991 — Conspectus Bryophytorum Orientalum et Arabicorum.

An annotated catalogue of the bryophytes of Southwest Asia. Bryophytorum Biblio-

theca 39: 1-181.

GREENE S. W. & HARRINGTON A. J., 1989 — The Conspectus of Bryological Taxonomic

Literature. Part 2. Guide to national and regional literature. Bryophytorum Bibliotheca

37: 1-321.

Source : MNHN. Paris

Cryptogamie, Bryol. Lichenol. 1997, 18(1) : 47-54 47

ADDITIONS TO THE BRYOPHYTE FLORA

OF LA RIOJA AND THE IBERIAN SYSTEM

(NORTHERN SPAIN)

J. MARTINEZ-ABAIGAR, E. NUNEZ-OLIVERA, A. GARCIA-ALVARO

and N. BEAUCOURT

Área de Biología Vegetal, Universidad de La Rioja, Luis de Ulloa 20, E-26004 Logroño, España.

Fax-Number: 34 + 41 — 299274; e-mail:

javier martinez@daa unirioja.es — Correspondence to: J. Martinez-Abaigar

RESUME — Quatorze taxons de bryophytes, récoltés pour la plupart dans les ceintures

bioclimatiques Oroméditerranéenne et Crioroméditerranéenne du Système ibérique, sont signalés

pour la premiére fois à La Rioja (Espagne). Six d'entre eux sont nouveaux pour l'ensemble du

Systéme iberique : Lophozia longiflora, Nardia geoscyphus, Bryum schleicheri var. latifolium,

Grimmia caespiticia, Pohlia obtusifolia et Seligeria recurvata. Des données chorologiques et

écologiques, leur présence sur la Liste touge des bryophytes de la Péninsule iberique, et leur

importance bryologique dans le Systéme ibérique sont ajoutées.

ABSTRACT — Fourteen taxa of bryophytes, mostly collected in the Oromediterranean and the

Crioromediterranean bioclimatic belts of the Iberian System, are reported as new for La Rioja

(Spain). Six of them are new for the entire Iberian System: Lophozia longiflora, Nardia geoscyphus,

Bryum schleicheri var. latifolium, Grimmia caespiticia, Pohlia obtusifolia and Seligeria recurvata.

Chorological, ecological and conservation data are added and the bryological importance of the

Iberian System is emphasized.

Key WORDS: La Rioja, Iberian System, Spain, bryophytes, flora, new records.

INTRODUCTION

La Rioja is the less extensive Autonomous Region of peninsular Spain.

However, the 5000 km? that it occupies show an outstanding ecological variability,

mainly due to its wide altitudinal range, from 280 m in the northeasternmost part to

2271 m in the peak of San Lorenzo (Sierra de la Demanda), in the southwestern area

of the region. The geological materials are also quite varied, dominated by Tertiary,

Jurassic-Cretaceous and Cambrian formations of diverse chemical nature (IGME,

1980). The climate combines three marked influences: continental, atlantic and

mediterranean (Núñez-Olivera & Martinez-Abaigar, 1991). La Rioja belongs entirely

to the mediterranean bioclimatic region and participates in four bioclimatic belts:

Mesomediterranean, Supramediterranean, Oromediterranean and an incipient Crioro-

Source : MNHN, Paris

48 J. MARTINEZ-ABAIGAR ег al.

MEDITERRANEAN,

Fig. 1 — Location map of the Iberian System in the Iberian Peninsula. The ranges of the Iberian

System where bryological sampling has been achieved are marked with consecutive numbers:

1, Páramos de La Lora; 2, Neila; 3, Demanda; 4, Cameros; 5, Moncayo; 6, Albarracin and

Monterde; 7, Javalambre and Gúdar; 8, Palomita; 9, Peñagolosa; 10, Titaguas; 11, Espadán.

mediterranean (Rivas-Martínez, 1987). Nevertheless, the montane zones represent an

atlantic island within the mediterranean region. The southern half of La Rioja is

formed by the foothills of the Iberian System, a 400 km-long range that extends from

the Cantabrian range up to nearly the Mediterranean sea, constituting the natural

boundary between the Ebro Valley and the Meseta (Fig. 1).

The bryological knowledge of La Rioja is irregular (Martinez-Abaigar et al.,

1995). The Supramediterranean bioclimatic belt, where the great majority of the forests

is found, is reasonably well-known. By contrast, the Mesomediterranean belt, the more

thickly populated and also the most perturbed zone, remains underexplored (Martinez-

Abaigar & Núñez-Olivera, 1996). Also, relatively few bryological data are available for

the Oromediterranean and the Crioromediterranean belts.

Bryological sampling of the Iberian System began early in the century,

although some scattered pioneer records were already given by Lagasca et al. (1802).

However, the most intense research, mainly promoted by Dr. Cruz Casas and

Source : MNHN, Paris

ADDITIONS TO THE BRYOPHYTE FLORA OF LA RIOJA 49

co-workers, has been carried out since the seventies. A significant number of ranges

with notably different ecological characteristics have been sampled (Table 1), but it

must be taken into account that sampling was not homogeneous in the different zones.

The bryological importance of the Iberian System lies in the biogeographical link that

it establishes between the northern ranges of Spain (Galician Massif, Cantabrian range,

Basque mountains, Pyrenees) and the central (Central System) and southern (Sierra

Nevada, for instance) ones (Casas de Puig, 1975; Casas, 1990b). Thus, many

bryophytes that were previously known from traditionally explored montane zones,

such as the Pyrenees, the Central System and Sierra Nevada (see Allorge, 1928, and the

references therein), finish up by being found also in the Iberian System (Casas de Puig,

1975; Casas et al., 1981; Martinez-Abaigar, 1986; Martinez-Abaigar & Ederra, 1992).

The 14 bryophytes listed in this paper were collected in more or less remote

places of La Rioja, mostly in the Oromediterranean and the Crioromediterranean

bioclimatic belts of the Iberian System. Therefore, subarctic-subalpine and montane

species prevail in our list. Hardly accessible places in montane zones of La Rioja are

interesting bryological refuges, since all of the quoted taxa are new for this province

and six of them, marked by an asterisk, are new for the entire Iberian System.

ANNOTATED LIST OF BRYOPHYTES

The nomenclature of the liverworts follows Grolle (1983), that of the mosses

Casas (1991). Voucher specimens are located in the personal herbarium of J.

Martinez-Abaigar and, when mentioned, also in BCB.

Hepaticae

Diplophyllum taxifolium (Wahlenb.) Dum. — La Rioja: Viniegra de Abajo, peak of

Urbión (30T WM1051, 2100 m), J. Martínez-Abaigar & E. Núñez-Olivera, 7/95.

Protosoils on quartzites, with Pohlia elongata. This subarctic-alpine species (Duell,

1983) was previously known in the Iberian System from the Moncayo (Casas er al.,

1984).

* Lophozia longiflora (Nees) Schiffn. — La Rioja: Anguiano, Regatillo stream (30T

WM2276, 1250 m), J. Martinez-Abaigar, 4/85. Humid acidic bank of a stream, among

Polytrichum debris, with Lophocolea bidentata and Plagiochila porelloides, under a

Fagus sylvatica canopy. The range of the species in Spain includes the provinces of

Alava, Asturias, Gerona, Guipúzcoa, Huesca, León and Lérida (Casas, in litt.). This is

the first record for the Iberian System and also the first one to the south of the river

Ebro, consequently outside from the Spanish northern mountains (Pyrenees, Basque

mountains, Cantabrian range).

* Nardia geoscyphus (De Not.) Lindb. — La Rioja: locality and habitat as those of

Lophozia longiflora. Nardia geoscyphus was previously known in Spain from the

provinces of Asturias, Baleares, Barcelona, Gerona and Huesca (Simó & Vigón, 1977;

Casas, in litt.). As L. longiflora, it is new for the Iberian System, being also the first

Source - MNHN. Paris

50 J. MARTINEZ-ABAIGAR et al.

peninsular record outside the northern mountains. N. geoscyphus is a “rare” species in

the Red List of the Iberian bryophytes (Sergio er al., 1994a).

Musci

Bryum radiculosum Brid. — La Rioja: Alberite (30T WM4594, 465 m), J. Martinez-

Abaigar, E. Núñez-Olivera & N. Beaucourt, 4/95. Wall of an irrigation ditch.

* Bryum schleicheri Lam. & DC. var. latifolium (Schwaegr.) Schimp. — La Rioja:

Viniegra de Abajo, Urbiön valley (30T WM1051, 1800 m), A. Garcia- Alvaro, 7/95,

duplicate in BCB. Wet grassland, bog margin. Previously known from the Pyrenees, the

Cantabrian range and Sierra Nevada (Casas, 1990b), this subarctic-subalpine species

(Duell, 1985) is new for the Iberian System.

Climacium dendroides (Hedw.) Web. & Mohr — La Rioja: Brieva de Cameros, river

Brieva (30T WM1764, 1450 m), A. Garcia-Älvaro, 6/95. Wet soil. In the Iberian

System, it was already found in the Sierra de Albarracin (Casas de Puig et al., 1977)

and in the Moncayo (Casas et al., 1984). A distribution map of this species in the

Tberian Peninsula was published by Casas er al. (1985).

Dicranella palustris (Dicks.) Crundw. ex E. Warb. — La Rioja: Villoslada de Cameros,

Hoyo Mayor (30T WM2650, 1800 m), J. Martinez-Abaigar, E. Núñez-Olivera & A.

Garcia-Alvaro, 7/94. Bogs and wet stream beds. It was known from most of the high

ranges of Spain, including the Central System and Sierra Nevada (Casas et al., 1985).

In the Iberian System it was previously collected in Sierra de la Demanda (Gil &

Guerra, 1985).

Dryptodon patens (Hedw.) Brid. — La Rioja: Viniegra de Abajo, peak of Urbiön (30T

WM1051, 2100 m), J. Martinez-Abaigar & E. Nünez-Olivera, 7/95. Protosoils on

quartzites. It is widely distributed in montane zones of the Iberian Peninsula, including

several localities in the Iberian System: Piqueras, Picos de Urbión (Casas de Puig, 1975)

and Sierra de Neila (Casas et al., 1995).

* Grimmia caespiticia (Brid.) Jur. — La Rioja: Viniegra de Abajo, peak of Urbión (30T

WM1051, 2100 m), J. Martinez-Abaigar 4 E. Nünez-Olivera, 7/95. Protosoils on

quartzites, with Dryptodon patens, Cynodontium bruntonii and Racomitrium sudeticum.

Muñoz (1995) recently summarized the distribution of G. caespiticia in Spain and

validated the records for the Pyrenees (provinces of Gerona, Huesca and Lérida), the

Cantabrian range (Cantabria) and the Central System (Sierra de Gredos, province of

Ávila). It was also recently discovered in Portugal (Serra da Estrela: Sergio er al.,

1994b). Our record is new for the Iberian System. This subarctic-subalpine species

(Duell, 1984) was included into the category “rare” in the Red List of the bryophytes

of the Iberian Peninsula (Sergio ег al., 1994a).

Plagiothecium denticulatum (Hedw.) B., S. & G. var. obtusifolium (Turn.) Moore — La

Rioja: Viniegra de Abajo, peak of Urbión (30T WM1051, 2100 m), J. Martinez-

Abaigar & E. Nünez-Olivera, 7/95. Sheltered soil on quartzite crevices. Although the

variety was only considered as a morphotype by Ireland (1986), it was retained by

Casas (1991). This taxon was already known in the Iberian System from neighbouring

localities to ours (Casas, 1990a ; Casas er al., 1995).

Source : MNHN, Paris

ADDITIONS TO THE BRYOPHYTE FLORA OF LA RIOJA 51

RANGE AND MAXIMUM ALTITUDE REFERENCE NUMBER OF SPECIES

1. Páramos de La Lora (1205 m) Casas et al. (19924) 171

2. Neila (2049 m) Casas (19904) 9

Casas et al. (1995) 146

3. Demanda (2271 m) Casas et al. (1981) 165

Casas de Brugués (1981) 188

Gil & Guerra (1985)" 140

4. Cameros (1766 m) Tonglet (1906) т

Casas de Puig (1975) 133

Martinez-Abaigar et al. (1985) 136

Martinez-Abaigar de Ederra (1992 123

5. Moncayo (2316 m) Tonglet (1903) 37

Casas et al. (1984) 172

6. Albarracin and Monterde (1920 m) Casas de Puig et al. (1977) 143

7. Javalambre and Güdar (2020 m) Casas de Brugués (1981) 121

Casas de Puig et al. (1982) 142

8. Palomita (1781 m) Casas & Puche (1985) 133

9. Peñagolosa (1813 m) Puche (1986) 184

10. Titaguas (840 m) Casas et al. (1976) 69

11. Espadán (1103 m) Puche ef al. (1993) 120

* This paper includes the records by Casas de Puig (1975), but only mosses are considered.

** This paper includes also records from Picos de Urbión.

*** This paper includes also records from Sierra Cebollera.

**** This paper only makes reference to mosses.

Tab. 1 — Ranges of the Iberian System sampled for bryophytes, arranged from the northwestern

extreme to the southeastern one. Order numbers are those shown in Fig. 1. The most significant

bibliographic references and the total number of the collected species for each range are given.

Source : MNHN, Paris

52 J. MARTINEZ-ABAIGAR et al.

Plagiothecium laetum B., S. & G. — La Rioja: Viniegra de Abajo, peak of Urbiön (30T

WM1051, 2100 m), J. Martinez-Abaigar & E. Nünez-Olivera, 7/95. Soil on quartzites.

It was previously recorded in the Iberian System from the Sierra de Neila (Casas,

1990a).

* Pohlia obtusifolia (Brid.) L. Koch — La Rioja: Viniegra de Abajo, peak of Urbion

(30T WM1051, 2100 m), J. Martinez-Abaigar & E. Nünez-Olivera, 7/95, duplicate in

BCB. Soil on quartzites. This arctic-alpine species (Duell, 1985) was only known in the

Iberian Peninsula from the Pyrenees (Casas Sicart, 1986). It is an “endangered” species

in Spain (Sérgio er al., 1994a).

Racomitrium sudeticum (Funck) В. & 5. — La Rioja: Viniegra de Abajo, peak of

Urbiön (30T WM1051, 2100 m), J. Martinez-Abaigar & E. Núñez-Olivera, 7/95. Soil on

quartzites. This subarctic-subalpine species (Duell, 1992) occurs in the Pyrenees,

Cantabrian range, Basque mountains and the Iberian System (Sierra de Neila: Casas

et al., 1992b).

* Seligeria recurvata (Hedw.) B., S. & G. — La Rioja: San Millan de la Cogolla,

Cardenas valley (30T WM0678, 1300 m), J. Martinez-Abaigar 4 E. Nünez-Olivera,

9/93. Protosoils in fissures of Cambrian limestones and schists. Puche (1989) gave a

distribution map of the species in the Iberian Peninsula, including the provinces of

Burgos, Gerona, Guipúzcoa, Huesca, Navarra and Vizcaya. Our record is new for the

Iberian System and the first one to the south of the river Ebro, outside from the

Pyrenees, the Basque mountains and the Cantabrian range.

ACKNOWLEDGEMENTS — We are very grateful to Drs. P. Boudier, Cruz Casas, L. Hedenäs,

N.G. Hodgetts, J. Muñoz, J.A. Shaw, J. Váña and H.L.K. Whitehouse for help in the

identification of some critical specimens and for informations about the chorology of the taxa.

This work is a part of the results of the project AMB95-0468, financially supported by the

Comisión Interministerial de Ciencia y Tecnologia (CICYT) of Spain.

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Cryptogamie, Bryol. Lichénol. 1997, 18 (1) : 55-61 55

LENGTH POLYMORPHISM IN THE INTRAGENIC SPACER 1

OF THE NUCLEAR RIBOSOMAL DNA

OF SOME POTTIACEAE (POTTIALES, MUSCI)

Valeria SPAGNUOLO ', Paolo CAPUTO ', Salvatore COZZOLINO ?,

Rosa CASTALDO ! and Paolo DE LUCA!

! Dipartimento di Biologia vegetale, Università degli Studi di Napoli Federico II,

Via Foria, 223, 180139 Napoli (Italy).

? Orto botanico, Università degli Studi di Napoli Federico II,

Via Foria, 223, 1-80139 Napoli (Italy).

ABSTRACT — Authors report on a length polymorphism in the intragenic spacer 1 (ITS1) of the

nuclear ribosomal DNA repeat. Species of the genera Barbula, Bryoerythrophyllum, Didymodon,

Microbryum, Pleurochaete, Syntrichiu, Timmiella, Tortella, Tortula, and Weissia were studied. The

differences in length of the ITS! match to a great extent recent evidence on the circumscription of

genera based on morphology.

INTRODUCTION

Pottiaceae are the largest family of mosses, both in species and genus number;

they have a widespread distribution, with various genera represented in Italy.

Several morphological characters of the Pottiaceae are related to the high

tolerance to desiccation (Proctor, 1979; Giordano et al.. 1993). Adaptation to such severe

selective pressures makes group delimitation and inference of phylogenetic relationships.

complicate when relying only on morphology. Morphological characters, in fact, may

develop in a parallel or convergent fashion in response to the same environmental stimuli.

As a result, taxonomy of Pottiaceae, which presently relies only on morphological traits, is

much controversial (in the ensuing sections of this paper we will follow Zander’s nomen-

clature), both in monographic and floristic works (for example, Corley et al., 1982; Cortini

Pedrotti, 1992; Saito, 1975; Smith, 1978; Zander, 1993).

For these reasons, an independent source of evidence would indeed be useful in

elucidating boundaries of taxa and historical propinquity within Pottiaceae.

Novel sources of evidence can easily be found in macromolecular analysis, and

especially in DNA studies. Molecular approaches, however, have been extensively used in

seed plants, and there is a dearth of such studies for cryptogams, and especially for

bryophytes. Possibly, one of the reasons for this depends upon the comparatively large

amounts of tissue which were required until a short time ago. Nowadays, the advent of

Source : MNHN, Paris

56 V. SPAGNUOLO et al.

PCR technique allows the usage of very reduced amounts of material, and makes any

study virtually independent from large amounts of collected or cultured material.

One of the most proficuously exploited systems in embryophytes are the nuclear

ribosomal repeated genes (n-rDNA), as they possess properties which turn them into

excellent tools to delimitate taxa and to infer their relationships across a wide range of

taxonomic circumscriptions. In fact, n-rDNA consists of three highly conserved coding

units (185, 5.85 and 265), which can be employed across very far removed taxa (Mishler et

al. 1994, and references therein); these coding regions are separated by two variable

internal transcribed spacers, ITS1 and 2, which can be used in comparisons among species

or closely related genera (Suh et al., 1993 ; Kim & Jansen, 1994 and references therein ;

Baldwin er al., 1995).

In this paper we report information on the length of the ITSI regions in some

Pottiaceae, comparing the data with the classifications by Corley er al. (1982) and Zander

(1993).

MATERIALS AND METHODS

The mosses analyzed and their acronyms are listed in Table 1. They were partly

collected in the Botanical Garden of Naples (TIM, DIDINS and DIDLUR) and partly in

the Natural Reserve of Castelvolturno (CE) (MIC, PLE, TERFLA, BAR and SYN) or in

the Natural Reserve of Valle delle Ferriere (SA) (BRY, TERNIT, TERTOR, TULSUB,

TULMUR, WED). Voucher specimens of all taxa examined are deposited at NAP.

DNA was extracted from approximately 200-300 mg of fresh gametophytic

tissue, by using the protocol by Caputo et al. (1991) with the following modifications:

30mM f-mercaptoethanol (instead of 15mM) in the extraction buffer and DNA precipi-

tation carried out in PEG 8000 10% and NaCl 5M 0.12 vols. (instead of 0.7 vol.

isopropanol). Extraction was carried out also on dried material, previously rehydrated.

For some species, a single fresh gametophytic tip was directly PCR amplified after

grinding on liquid N».

Amplification of ITS1 was carried out by using a pair of primers, annealing in

the 3 region of the 185 (5'-GGAGAAGTCGTAACAAGGTTTCCG-3) and in the

5 region of the 5.85 (5 -CCAAACAACCCGACTCGTAGACAGC-3’). Typical reaction

mixtures (100 pl) contained approximately 5 ng genomic DNA or 100 u fresh tissue. Blotto

to 2% final volume (De Boer ег al., 1995) was added to the mixture. PCR reactions were

conducted in a thermal cycler (Perkin Elmer Cetus 9600) for 30 cycles. Initial conditions

were as follows: 1 min denaturation at 94°C, | min annealing at 55°C, 45 sec extension at

72°C. Samples were denatured for 5 min at 94°C before the beginning of the first cycle;

extension time was increased of 3 sec/cycle; extension was further prolonged for 7 min at

the end of the last cycle. Samples were then removed from the thermocycler and stored at

-20°C until analyzed.

PCR samples were resolved electrophoretically on an agarose gel (0.2 ul of the

raw PCR product per lane, 2% agarose) and visualized by ethidium bromide fluorescence.

Length of the amplified fragments were estimated by comparison with a 100 bp DNA

ladder (Promega) and a PCR marker (Promega). All PCR fragments were tested for

homology via hybridization against a clone containing a complete sunflower ribosomal

repeat obtained through the courtesy of M. Arnold (University of Gainsville, Florida),

acknowledged here.

Source : ММНМ Paris

ITS 1 LENGTH IN POTTIACEAE

Table 1 — Taxa of Pottiaceae analyzed in the present study, their acronyms, length (bp) of the raw

ITSI fragments and subfamilial attribution according to Zander (1993) and Corley et al. 1982). The

synonyms which are relevant to the discussion, as desumed by Zander, are reported.

Taxon Acronym | ITS! Length Subfamilial attribution

(bp) | Zander 1993 Corley et al. 1982

Barbula Hedw. Merceyoideae | Pottioideae

Barbula convoluta Hedw. BAR 530+

Bryoerythrophyllum Chen Merceyoideae | Pottioideae

Bryoerythrophyllum recurvirostrum (Hedw.) | BRY 440

Chen

Didymodon Hedw. Merceyoideae | Pottioideae

Didymodon insulanus (De Not.) Hill DIDINS 410

Didymodon luridus Homsch. in Spreng. DIDLUR 410

Microbryum Schimp. Pottioideae Pottioideae

Microbryum starckeanum (Hedw.) Zand, | MIC 450

= Portia starckeana (Hedw.) С. Mall.

= Weissia starckeana Hedw.

Syntrichia Brid. Pottioideae Pottioideae

Syntrichia ruralis (Hedw.) Web. et Mohr. — | SYN 530

var. arenicola

= Tortula ruraliformis Hedw.

Tortula Hedw. Pottioideae Poitioideae

Tortula muralis Hedw. TULMUR 380

Tortula subulata Hedw. TULSUB 380

Weissia Hedw. * Pottioideae Trichostomoideae

Weissia controversa Hedw. WEL 450

Pleurochaete Lindb. Trichostomoideae | Trichostomoideae

Pleurochaete squarrosa (Brid.) Lindb. PLE 470

Tortella (Lindb.) Limpr. Trichostomoideae | Trichostomoideae

Tortella flavovirens (Bruch) Broth TERFLA 470

Tortella nitida (Lindb.) Broth. TERNIT 470

Tortella tortuosa (Hedw.) Limpr. TERTOR 470

Timmiella (De Not.) Limpr. Timmielloideae | Trichostomoideae

Timmiella cf. barbuloides (Brid.) Monk. TIM 580

In order to avoid length estimation errors due to mistermination of PCR

products, PCR fragments were all cut with Bin I and EcoR V. From preliminary observa-

tion of the available 18S and 5.8S sequences in the literature, in fact, these are among the

restriction endonucleases which have sites in the 3’ region of the 18S (downstream to our

5” primer) and in the 5’ region of the 5.85 (upstream to our 3’ primer).

RESULTS

The length of the raw PCR fragments is reported in Table | for all taxa. Raw

1751 fragments show length polymorphisms in the taxa examined here. In particular,

within Pottioideae (Fig. 1), SYN has the longest raw ITS1 fragment (530 bp); MIC and

WEI share a shorter fragment (450 bp), but longer than those of TULMUR and

TULSUB, which are in turn identical to each other (380 bp).

Source : MNHN. Paris

Fig. | — Electropherogram of the raw ITSI

fragments for the genera of Pottioideae in

study. From left to right: molecular weight

standard (PCR marker: 1000, 750, 500, 300,

150 bp), SYN, TULMUR, TULSUB, MIC,

WEI, molecular weight standard (100 bp DNA

ladder, the brightest fragment is 500 bp).

Fig. 2 — Electropherogram of the rz

fragments for the genera of Merc

study. From left to right: molecular weight

standard (PCR marker), BAR, DIDINS,

DIDLUR, BRY, molecular weight standard

(100 bp DNA ladder). Details of the length in

bp of the various fragments of the two stan-

dards are reported in the caption to Fig. 1

Fig. 3 — Electropherogram of the raw ITSI

fragments for the genera of Trichostomoideae

and Timmielloideae in study, From left to

right: molecular weight standard (PCR mar-

ker), PLE, TERFLA, TERNIT, TERTOR,

TIM, molecular weight standard (100 bp

DNA ladder). Details of the length in bp of the

various fragments of the two standards are

reported in the caption to Fig. 1.

Source : MNHN. Paris

ITS 1 LENGTH IN POTTIACEAE 59

Within Merceyoideae (Fig. 2), BAR has the longest raw ITS] (530 bp); a shorter

fragment characterizes BRY (440 bp); DIDINS and DIDLUR have the shortest fragment

(410 bp). Within Trichostomoideae (Fig. 3), PLE, TERFLA, TERNIT, and TERTOR

have an identical raw ITS1 fragment (470 bp). Within Timmielloideae (Fig. 3), TIM has a

raw ITS] 580 bp long.

Digestion with Bin I reduced the length of all our fragments of about 10-20 bp,

although the very small fragment resulted was very difficult to observe in our gel system;

Eco ВУ digestions yielded a fragment of about 100 bp plus fragments of different length in

the various samples. This fragment approximately corresponds in length to the distance

between the EcoRV site and the 3’ terminus of our minus primer in all the sequences.

examined in the literature. In all these sequences, the Eco RV site is approx. 20 bp

downstream the 3’ end of ITS1. Therefore, differences in length of our fragments are

attributable to polymorphisms within ITS1.

DISCUSSION AND CONCLUSIONS,

In the ensuing discussion we will comment our data by comparing them with the

conservative taxonomic treatment suggested by Corley er al. (1982) and the novel propo-

sals made by Zander (1993). Given the intricacies of pottiaceous nomenclature at all

taxonomic hierarchies an account of the differences in treatment between Corley er al.

(1982) and Zander (1993) follows (see also Table 1).

Following the former authors, the taxa examined here belong to two different

subfamilies: the Pottioideae, with Barbula, Bryoerythrophyllum, Didymodon, Pottia (inclu-

ding Microbryum starckeanum = Pottia starckeana), and Tortula (including Syntrichia)

and the Trichostomoideae, with Pleurochaete, Timmiella, Tortella, and Weissia. The more

splitted classification of Zander (1993), on the contrary, indicates four subfamilies:

Merceyoideae, with Barbula, Bryoerythrophyllum, and Didymodon; Pottioideae, h

Microbryum, Syntrichia, Tortula, and Weissia; Trichostomoideae, with Pleurochaete and

Tortella, and Timmielloideae, with Timmiella.

Asa whole, 1751 fragments of Pottioideae and Merceyoideae are more variable

than those of the Trichostomoideae and, of course, of the monotytpic Timmielloideae

(Figs. 1, 2, 3).

Within Pottioideae (Fig. 1), Microbryum starckeanum has an ITS1 of the same

length as Weissia controversa. Notably, M. starckeanum, recently segregated from Pottia (=

Pottia starckeana (Hedw.) C. Müller), was first described as W. starckeana Hedw. In this

case, the length of the ITSI does not provide support for the separation.

The two species of Tortula analyzed here are both placed in sect. Tortula by

Zander (1993). However, Corley et al. (1982) included T. muralis in sect. Cuneifoliae (В. &

S.) Spruce, and T. subulata in sect. Tortula. Our data, which indicate identity in length of

the ITS1 fragments in both species (Fig. 1), would concur to a greater extent with the

treatment of Zander. The same authors include Т. ruraliformis within the sect. Rurales De

Not. of the same genus. This section has been recently synonymized with genus Syntrichia.

Since ITSI of Syntrichia ruralis var. arenicola (= Tortula ruraliformis) is longer than those

of T. muralis and T. subulata (Fig. 1), a separation between Tortula sensu stricto and

Tortula sect. Rurales (=Syntrichia sensu Zander) finds support in our data.

Within Merceyoideae, the two species of Didymodon have and ITS1 of identical

length (Fig. 2). The latter genus is different in length when compared to both the other

Source : MNHN. Paris

60 У. SPAGNUOLO et al.

genera of the subfamily present in this analysis, Barbula and Bryoerythrophyllum. In

particular, Barbula and Didymodon, which were regarded as belonging to the same genus

by some authors (Smith, 1978), appear well distinct (Fig. 2).

As to the Trichostomoideae, our data indicate an identical length of the ITS1 in

both Pleurochaete and three species of Tortella analyzed here (Fig. 3). The length of the

ITS] of the latter taxa differs from that of Timmiella (Fig. 3). This genus, placed by Zander

in its own subfamily on the grounds of its autapomorphic bistratose laminal cells not

aligned near the costa, had been formerly included by Corley er al. (1982) in Trichosto-

moideae. Founding on our data, the separation suggested by Zander seems appropriate.

Of course, our data, which concur to a greater extent with Zander's classification

(1993) than with that by Corley et al. (1982), are indicative of separation only among the

polymorphic taxa listed above, and no inference can be drawn on the phylogeny of the

genera and species. Further studies, including sequencing the ITS] of a wider representa-

tion of taxa, are needed to rtain the mutual relationships within the family, as well as

the possible biological significance of the length variations in ITS1.

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Buffalo Society of Natural Sciences 32: 1-3

Source : MNHN. Paris

Cryptogamie, Bryol. Lichénol. 1997, 18 (1) : 63-75 63

CONTRIBUCIÓN AL CONOCIMIENTO

DE LOS LÍQUENES EPIFÍTICOS 4

DE LOS MONTES DE TOLEDO (TOLEDO, ESPANA)

Gregorio ARAGON ! & Isabel MARTINEZ ?

1 Dpto. Biologia Vegetal П, Facultad de Farmacia,

Universidad Complutense, 28040-Madrid, España.

2 Dpto. Biología Vegetal I, Facultad de Biologia,

Universidad Complutense, 28040-Madrid, España

RESUMEN — Se presentan los resultados floristicos de un estudio sobre los liquenes epifiticos de los

Montes de Toledo (España). En total 133 taxones, de los cuales 60 son primera cita provincial. De

ellos destacamos Calicium glaucellum, Chaenothecopsis pusilla, Koerberia biformis, Lecidea botryosa,

Lecidella pulveracea, Megalaria laureri, Micarea bauschiana, M. denigrata, Ochrolechia balcanica,

Pannaria olivacea, Rinodina conradii y Waynea adscendens por constituir importantes ampliaciones

de área.

ABSTRACT — The floristic results of a survey devoted to the epiphytic lichens from Montes de

Toledo are given. We include 133 taxa, 60 of them first provincial records. Important extensions of the

areal range in the Iberian Peninsula are afforded for Calicium glaucellum, Chaenothecopsis pusilla,

Koerberia biformis, Lecidea botryosa, Lecidella pulveracea, Megalaria laureri, Micarea bauschiana, M.

denigrata, Ochrolechia balcanica, Pannaria olivacea, Rinodina conradii and Waynea adscendens.

RÉSUMÉ — Dans les Monts de Toléde (Espagne), 133 taxons de lichens épiphytes ont été recensés,

parmi ceux-ci 60 sont cités pour la premiere fois pour la Province de Toléde. Les résultats obtenus

contribuent à une meilleure connaissance de la chorologie des taxons suivants pour la Péninsule

Ibérique: Calicium glaucellum, Chaenothecopsis pusilla, Koerberia biformis, Lecidea botryosa, Leci-

della pulveracea, Megalaria laurei, Micarea bauschiana, M. denigrata, Ochrolechia balcanica, Pannaria

olivacea, Rinodina conradi et Waynea adscendens.

INTRODUCCIÓN

Los Montes de Toledo presentan una longitud de 150 Km y una anchura de unos

50 Km, y su superficie queda repartida entre las provincias de Toledo y Ciudad Real. Están

constituidos por una serie de sierras cortas, próximas entre sí, cuyas altitudes oscilan entre

los 1150 m del Collado Marchés y los 1447 m del Rocigalgo. Los materiales dominantes

son areniscas y pizarras del Acadiense superior (Cámbrico) y cuarcitas armoricanas y

pizarras arcillosas con fuerte esquistosidad (Ordovicico). La red hidrográfica está com-

puesta por una serie de cortos ríos que vierten sus aguas al Tajo y al Guadiana (Fig. 1).

Source : MNHN, Paris

64 G. ARAGON & I. MARTINEZ

TOLEDO

“өз

ж 2

Fig. 1. — Localización del área de estudio: límites provinciales (+-+-+). Relación de las localidades

muestreadas: 1 — San Pablo de los Montes, arroyo del Marchés, UJ8177, 950 m. 2 — Los Navalu-

cillos, arroyo del Chorro, UJ5879, 1000 m. 3 — Hontanar, arroyo del Gatillo, 077384, 1100 m. 4 —

Hontanar, río Estena, UJ6380, 875 m. -Todas la localidades están situadas en el huso 30S de la

cuadrícula U.T.M.

Fig. 1. — Situation of the study area: provincial limits (+-+-+). Sample localities: 1 — San Pablo de

los Montes, arroyo del Marchés, UJ8177, 950 m. 2 — Los Navalucillos, arroyo del Chorro, UJ5879,

1000 m. 3 — Hontanar, arroyo del Gatillo, UJ7384, 1100 m. 4 — Hontanar, río Estena, UJ638,

875 m. -АП the listed sample sites are located in the spindle and zone 30S of the U.T.M. grid.

Source : MNHN. Paris

LÍQUINES EPIFÍTICOS DE LOS MONTES DE TOLEDO (ESPANA) 65

El clima es subhümedo templado o fresco de caräcter continental, con sequia

estival atenuada y superávit de agua en invierno y primavera (Estación meteorológica de

San Pablo de los Montes, Tm = 12,5%C, m = 8,44°C, T = 24°C, Pm= 798,8 mm). La zona

de estudio se encuentra enclavada en los pisos mesomediterráneo y supramediterráneo

(Rivas-Martínez, 1987).

La vegetación vascular está formada por encinares (Pyro bourgeanae-Quercetum

rotundifoliae Rivas-Martínez 1987), alcornocales (Sanguisorbo agrimonioides-Quercetum

suberis Rivas Goday 1959) y melojares (Arbuto unedonis-Quercetum pyrenaicae Rivas

Goday ex Rivas-Martínez 1987), todos ellos situados en el piso mesomediterráneo. En el

piso supramediterráneo aparecen melojares (Sorbo torminalis-Quercetum pyrenaicae

(Rivas Goday 1964) Rivas-Martinez 1987 y jarales (Genisto hirsutae-Cistetum ladaniferi

Rivas Goday 1955). Biogeográficamente, la zona de estudio pertenece al subsector ore-

tano, sector toledano-tagano, provincia luso-extremadurense, región mediterránea

(Rivas-Martínez, 1987).

El grado de conocimiento que existe sobre la flora de líquenes epifíticos en la

provincia de Toledo es escaso, limitándose a trabajos antiguos (Maheu & Gillet, 1922; Des

Abbayes, 1946) o parciales (Crespo, 1979; Crespo & Atienza, 1989; Fuertes & Burgaz,

1989; Burgaz et al., 1994; Sarrión & Aragón, 1995 y Sarrión & Burgaz, 1995). Unicamente,

Martínez et al. (1993) aportan un catálogo exclusivamente de la Sierra de San Pablo

(Montes de Toledo). La escasez de datos liquénicos, unido a las peculiares características

que presentan estos montes justifican este trabajo. Hasta ahora, las localidades estudiadas

corresponden únicamente al sector occidental de los Montes de Toledo, la zona que

presenta formaciones arbóreas mejor conservadas y con mayor influencia atlántica,

MATERIAL Y MÉTODO

Las recolecciones incluyen material epifítico recolectado sobre diferentes foró-

fitos: Castanea sativa, Cistus ladanifer, Crataegus monogyna, Fraxinus angustifolia, Ilex

aquifolium, Juniperus oxycedrus, Quercus faginea, Quercus pyrenaica, Quercus ilex, Quer-

cus suber, Sorbus torminalis y Taxus baccata, asi como tocones de fagäceas y madera

muerta de enebro.

La abundancia relativa de las especies en la zona se señala con las abreviaturas:

CC (muy común), C (común), R (rara) y RR (muy rara). Se señalan con número arábigo

las localidades de recolección (Fig. 1) y el número de recolección. El material se encuentra

depositado en el herbario MA (Real Jardín Botánico de Madrid) y en el herbario personal

del segundo de los autores (HIMM). En cada taxon se incluye el número de recolección del

primero de los autores, indicado por Ar., número de orden y año de recolección. Para las

especies más interesantes se incluyen algunos comentarios sobre sus características eco-

lógicas y corológicas más relevantes y, para las especies menos conocidas se señala alguna

referencia bibliográfica donde exista una descripción, foto o dibujo que la caracterice (Ic.).

En la identificación de los táxones se ha seguido a Clauzade & Roux (1985),

Purvis er al. (1992), Giralt (1994) en el género Rinodina, Jorgensen (1978) en el género

Pannaria y Tibell (1987) en el orden Caliciales. Para la nomenclatura se ha seguido el

criterio de Nimis (1993) en la mayoría de los casos y el criterio de Wirth (1980) en lo

referente a la distribución europea de las especies.

Source : MNHN. Paris

66 G. ARAGON & I. MARTINEZ

RESULTADOS

Se han identificado un total de 133 taxones, 60 de los cuales son primeras citas

para la provincia de Toledo (signo *).

* Amandinea punctata (Hoffm.) Coppins & Scheid. —R, 4, Ar. 1008/95, Taxus baccata.

Anaptychia ciliaris (L.) Massal. — С, 3, Ar. 847/95, Quercus pyrenaica; 4, Ar. 907/95,

Quercus ilex ; 4, Ar. 1001/95, Fraxinus angustifolia.

* Bacidia naegelii (Hepp) Zahlbr. — En España existen pocas citas y muy dispersas por

todo el territorio. La más cercana es de Ciudad Real (Sarrión et а/., 1993). — RR, 2, Ar.

1040/95, Quercus ilex.

* Bacidia rosella (Pers.) de Not. — CC, 2, Ar, 499/95, 567/95, Quercus ilex ; 4, Ar. 1025/95,

Quercus faginea.

* Bacidia rubella (Hoffm.) Massal. — СС, 1, Ar, 78/95, Quercus pyrenaica; 2, Ar. 426/95,

Quercus ilex ; 4, Ar. 908/95, Quercus faginea.

Buellia cedricola Werner — (Ic.: Burgaz & Sarrión, 1995). De distribución mediterranea,

se desarrolla habitualmente sobre madera muerta de enebro en comunidades del Buellie-

tum cedricolae (Sarrión $ Burgaz, 1995). En la Península Ibérica ha sido citada reciente-

mente en Ciudad Real, Jaén y Toledo (Sarrión & Burgaz, 1995). — С, 2, Ar. 464/95, Ar.

469/95, madera muerta de Juniperus oxycedrus; 4, Ar, 913/95, madera muerta de Juniperus

oxycedrus.

* Buellia disciformis (Fr.) Mudd — С, 2, Ar. 425/95, 496/95, Quercus ilex.

Buellia lauricassiae (Fée) Müll. Arg. —R, 2, Ar. 374/95, Quercus ilex.

*Calicium abietinum Pers, — В. 1, Ar. 90/95, tocón de Quercus pyrenaica; 2, Ar. 559/95,

tocón de Quercus ilex; 3, Ar. 561/95, tocón de Quercus pyrenaica.

*Calicium glaucellum Ach. — (Ic.: Tibell, 1987). Taxon bastante frecuente en la zona de

estudio y siempre sobre áreas descortezadas de tocones de Quercus pyrenaica. Aparece

junto a Calicium abietinum, Calicium salicinum y Pertusaria flavida. En España sólamente

ha sido citado en Álava (Etayo, 1991), Ciudad Real (Sarrión er al.,1993), Lugo (Valcárcel

et al., 1993), Navarra (Etayo, 1989) y Segovia (Martinez & Aragón, 1996). — CC, 1, Ar.

80/95, tocón de Quercus pyrenaica; 2, Ar. 180/95, 456/95, tocón de Quercus pyrenaica; 3,

Ar, 841/95, tocón de Quercus pyrenaica.

*Calicium salicinum Pers. — Coloniza zonas descortezadas de los troncos y tocones de

fagaceas. Aparece de forma dispersa por la Península Ibérica: Ciudad Real (Sarrión et al,

1993), Lérida (Hladún, 1984), Málaga (Seaward, 1983) y Navarra (Etayo & Gómez-Bolea,

1992). — R, 1, Ar. 72/95, tocón de Quercus pyrenaica.

Calicium sp. — L. Tibell (in litt.) nos confirma que se trata de una nueva especie que

conoce desde hace 20 años y se encuentra ampliamente distribuida por Europa. Desar-

rolla un talo grueso y de aspecto verrucoso. Es frecuente observarla sobre madera muerta

y descortezada de Juniperus oxycedrus. — C, 2, Ar. 463/95, madera muerta de Juniperus

oxycedrus; 4, Ar. 1007/95, madera muerta de Juniperus oxycedrus.

Source : MNHN, Paris

LÍQUINES EPIFÍTICOS DE LOS MONTES DE TOLEDO (ESPANA) 67

*Caloplaca cerina (Hedwig) Th. Fr. — R, 2, Ar. 532/95, Quercus ilex.

Caloplaca ferruginea (Hudson) Th. Fr. — C, 2, Ar. 498/95, 521/95, Quercus ilex ; 3, Ar.

865/95, Quercus pyrenaica.

*Caloplaca haematites (Saint-Amans) Zwackh — RR, 4, Ar. 912/95, Quercus ilex.

*Caloplaca herbidella (Hue) Magnusson — C, 4, Ar, 903/95, 1015/95, Juniperus oxycedrus;

4, Ar. 909/95, Quercus ilex.

*Candelariella vitellina (Hoffm.) Müll. Arg. — R, 3, Ar. 842/95, Quercus pyrenaica.

Candelariella xanthostigma (Ach.) Lettau — R, 2, Ar. 206/95, tocón de Quercus pyrenaica.

Cetraria merillii Du Rietz — Especie pionera que coloniza las ramas de Cistus ladanifer

junto con Evernia prunastri. — C, 2, Ar. 198/95, Cistus ladanifer.

* Chaenothecopsis pusilla (Ach.) A. Schmidt — (Ic.: Tibell 1987). Su área de distribución se

amplia notablemente, ya que las ünicas citas en la Peninsula Iberica son de Minho

(Sampaio, 1970) y Navarra (Etayo, 1990). — RR, 2, madera muerta de Arbutus unedo.

*Cladonia chlorophaea (Sommerf.) Sprengel — R, 3, Ar. 895/95, base de Quercus pyre-

naica,

Cladonia cyathomorpha Stirt. ex Walt. Watson — Habita en los tocones de fagaceas que se

descomponen con facilidad, junto a Cladonia fimbriata у С. subulata. —R, 4, Ar. 1046/95,

tocón de Quercus ilex.

Cladonia fimbriata (L.) Fr.— CC, 2, Ar. 173/95, 382/95, 531/95, Quercus ilex; 2, Ar. 481/95,

Quercus faginea; 3, Ar. 896/95, Quercus ругепаіса; 4, Ar. 1045/95, tocón de Quercus

faginea.

Cladonia pyxidata (L.) Hoffm. — R, 3, Ar. 897/95, base de Quercus pyrenaica.

Cladonia rangiformis Hoffm. — R, 3, Ar. 898/95, base de Quercus pyrenaica.

*Cladonia squamosa Hoffm. — R, 3, Ar. 899/95, base de Quercus pyrenaica.

*Cladonia subulata (L.) Wigg. — R, 2, 438/95, tocón de Quercus pyrenaica.

*Coelocaulon aculeatum (Schreber) Link — De carácter terricola, suele colonizar fisuras

en la madera de enebro, donde se acumula algo de suelo. — R, 2, Ar. 468/95, base de

Juniperus oxycedrus; 4, Ar. 1029/95, base de Juniperus oxycedrus.

*Coelocaulon crespoae Barreno & Vázquez— R, 1, Ar. 76/95, tocón de Quercus pyrenaica.

*Collema fasciculare (L.) Wigg. — CC, 2, Ar. 427/95, 537/95, Quercus ilex ; 4, Ar. 1041/95,

Quercus ilex.

Collema furfuraceum (Arnold) Du Rietz — CC, 2, Ar. 410/95, Quercus ilex ; 4, Ar. 1042/95,

Quercus ilex.

Collema nigrescens (Hudson) DC. — C, 2, Ar. 156/95, Quercus ilex : 3, Ar. 873/95, Quercus

pyrenaica.

Collema subflaccidum Degel. — C, 2, Ar. 543/95, Quercus ilex; 4, Ar. 1039/95, Quercus ilex.

Source . MNHN. Paris

68 G. ARAGON & I. MARTINEZ

*Collema subnigrescens Degel. — C, 2, Ar. 392/95, Quercus ilex; 4, Ar. 1023/95, Quercus

ilex.

Cyphelium tigillare (Ach.) Ach. — (Ic.: Tibell 1971). Se desarrolla habitualmente sobre

madera de gran dureza junto con Buellia cedricola y Pyrrhospora elabens. Existen muy

pocas referencias de este taxon en la peninsula Ibérica: Ciudad Real (Sarrión & Burgaz,

1995), Granada (Amo y Mora, 1870), Navarra (Etayo, 1990) y Toledo (Sarrión & Aragón,

1995). — RR, 2, Ar. 477/95, madera muerta de Juniperus oxycedrus.

Degelia atlantica (Degel.) P.M. Jorg. & P. James — C, 2, Ar. 399/95, Quercus ilex; 4, Ar.

1004/95, Quercus ilex.

Degelia plumbea (Lightf.) P.M. Jørg. & P. James — CC, 2, Ar. 155/95, 167/95, Quercus ilex;

4, Ar. 904/95, Quercus ilex; 4, Ar. 900/95, Juniperus oxycedrus; 4, Ar. 1027/95, Fraxinus

angustifolia; 4, Ar. 1026/95, Quercus faginea.

Dendriscocaulon umhausense (Auersw.) Degel. — CC, 2, Ar. 482/95, Quercus ilex; 2, At.

446/95, Quercus faginea; 3, Ar. 849/95, Quercus pyrenaica; 4, Ar. 1014/95, Juniperus

oxycedrus; 4, Ar. 1010/95, Fraxinus angustifolia.

Diploschistes muscorum (Scop.) Santesson CC, 2, Ar. 170/95, Quercus ilex; 3, Ar.

848/95, Quercus pyrenaica; 3, Ar. 853/95, tocón de Quercus pyrenaica.

Diplotomma populorum Massal. — C, 2, Ar. 124/95, 205/95, Quercus pyrenaica.

Evernia prunastri (L.) Ach. — CC, 3, Ar. 887/95, Quercus pyrenaica; 4, Ar. 913/95, Quercus

ilex .

Hypocenomyce scalaris (Liljeblad) M. Choisy — C, 2, Ar. 466/95, madera muerta de

Juniperus oxycedrus; 4, Ar. 902/95, Juniperus oxycedrus.

Hypogymnia physodes (L.) Nyl. — CC, 2, Ar. 197/95, Cistus ladanifer; 2, Ar. 459/95,

madera muerta de Juniperus oxycedrus.

Hypogymnia tubulosa (Schaerer) Havaas — CC, 2, Ar. 171/95, Quercus ilex; 3, Ar. 854/95,

Quercus ilex; 3, Ar. 888/95, Quercus pyrenaica.

* Koerberia biformis Massal. — (Ic.: Henssen, 1963). Especie mediterránea y poco fre-

cuente en la zona de estudio, recolectada siempre sobre troncos de encina centenarios y en

zonas húmedas. Acompaña a Leptogium lichenoides, Polychidium muscicola y Waynea

adscendens. En la Península ha sido citada en Cádiz (Arvidsson, 1979), Málaga (Seaward,

1983), Jaén (Bricaud & Roux, 1993), Salamanca (Marcos Laso, 1985) y Portugal (Sam-

paio, 1924; Тауагеѕ, 1950). — RR, 2, Ar. 480/95, Quercus ilex.

Lecanora chlarotera Nyl. — CC, 2, Ar. 189/95, Cistus ladanifer, 2, Ar. 141/95, 216/95,

Quercus ilex; З, Ar. 880/95, Quercus ilex; 3, Ar. 877/95, Quercus pyrenaica; 4, Ar. 1036/95,

Quercus ilex.

*Lecanora intumescens (Rebent.) Rabenh. — CC, 2. Ar. 512/95, Quercus ilex; 4, Ar.

1035/95, Quercus ilex.

Lecanora pallida (Schreber) Rabenh. — C, 3, Ar. 876/95, Quercus pyrenaica.

*Lecanora pulicaris (Pers.) Ach. — C, 3, Ar. 882/95, Quercus ilex; 3, Ar. 879/95, Quercus

pyrenaica.

Source . MNHN. Paris

LIQUINES EPIFITICOS DE LOS MONTES DE TOLEDO (ESPANA) 69

*Lecanora rugosella Zahlbr. — CC, 2. Ar. 186/95, Cistus ladanifer; 3, Ar. 881/95, Quercus

ilex; 3, Ar. 878/95, Quercus pyrenaica.

* Lecanora symmicta (Ach.) Ach. — Б, 1, Ar. 83/95, tocón de Quercus pyrenaica.

* Lecanora varia (Hottm.) Ach. — Aparece en tocones de fagaceas, iluminados, en zonas

descortezadas y duras. — CC, 2, Ar. 200/95, tocön de Quercus pyrenaica; 3, Ar. 875/95,

tocön de Quercus pyrenaica; 4, Ar. 1033/95, madera muerta de Juniperus oxycedrus.

*Lecidea botryosa (Fr.) Th. Fr. — En los Montes de Toledo se desarrolla habitualmente

sobre madera muerta de fagäceas. En España presenta una distribución incierta, siendo la

cita mas cercana de Madrid (Crespo & Bueno, 1982). — CC, 1, Ar. 75/95, tocön de

Quercus pyrenaica; 2, Ar. 430/95, 440/95, tocön de Quercus pyrenaica; 2, Ar. 529/95,

madera muerta de Juniperus oxycedrus.

Lecidella euphorea (Flórke) Hertel — CC, 2, Ar, 551/95, Quercus ilex; 2, Ar. 433/95, tocón

de Quercus pyrenaica; 3, Ar. 843/95, Quercus pyrenaica.

*Lecidella pulveracea (Th. Fr.) Sydow — La mayoria de las citas son del norte de la

Peninsula; la mas cercana es de Madrid (Crespo & Bueno, 1982). — RR, 4, Ar. 1037/95,

Пех aquifolium.

Lepraria incana (L.) Ach. — R, 4, Ar. 1039/95, madera muerta de Juniperus oxycedrus.

* Leprocaulon microscopicum (Vill.) D. Hawksw. — CC, 1, Ar. 95/95 base de Quercus ilex;

2, Ar. 177/95, base de Quercus pyrenaica; 2, Ar. 553/95, base de Quercus ilex.

Leptogium lichenoides (L.) Zahlbr. — CC, 2, Ar. 405/95, 505/95, Quercus ilex; 4, Ar.

1020/95, Quercus ilex; 4, Ar. 1017/95, Fraxinus angustifolia; 4, Ar. 1032/95, Taxus baccata.

Leptogium saturninum (Dickson) Nyl. — RR, 2, Ar. 420/95, Quercus ilex .

Lobaria amplissima (Scop.) Forss. — Aparece epifítica en fresnos cercanos a cursos de

agua. — R, 2, Ar. 525/95, Fraxinus angustifolia; 4, Ar. 1011/95, Fraxinus angustifolia.

Lobaria pulmonaria (L.) Hoffm. — CC, 2, Ar, 520/95, Fraxinus angustifolia; 2, Ar. 185/95,

Quercus pyrenaica; 4, Ar. 1002/95, Fraxinus angustifolia.

Lobaria scrobiculata (Scop.) DC. — CC, 2, Ar. 184/95, Quercus pyrenaica: 4, Ar. 1013/95,

Juniperus oxycedrus; 4, Аг. 1012/95, Fraxinus angustifolia,

* Megalaria laureri (Th. Fr.) Hafellner — Taxon frecuente, recogido sobre corteza de Пех

aquifolium en zonas umbrosas y hümedas. En la Península Ibérica ha sido citado en

Cataluña (Llimona et al., 1987) y Navarra (Etayo & Gómez-Bolea, 1992). — RR, 4, Ar.

1038/95, Ilex aquifolium.

* Megaspora verrucosa var. mutabilis (Ach.) Nimis & Roux — RR, 2, Ar. 563/95, Quercus

faginea.

* Micarea bauschiana (Körber) V. Wirth & Vezda — (Ic.: Coppins, 1983). Su distribución

conocida en España se restringe a Galicia (Carballal et al., 1995) y Jaén (Sarrión & Burgaz,

1995). — RR, 4, Ar. 1034/95, tocón de Quercus faginea.

* Micarea denigrata (Fr.) Hed]. — (Ic.: Coppins, 1983). Nuestra cita constituye la localidad

más meridional de este taxon en España. — RR, 2, Ar. 533/95, Quercus pyrenaica.

Source : MNHN., Paris

70 G. ARAGON & I. MARTINEZ

*Mycobilimbia berengeriana (Massal.) Haf. & V. Wirth — R, 2, Ar. 129/95, base de Quercus

pyrenaica (sobre briöfitos); 3, Ar. 858/95, base de Quercus pyrenaica.

*Mycobilimbia hypnorum (Libert) Kalb & Haf. R, 1, Ar. 77/95, base de Quercus

pyrenaica (sobre briófitos); 3, Ar. 859/95, Quercus pyrenaica (sobre briófitos).

*Mycocalicium subtile (Pers.) Szat. — В, 2, Ar. 565/95, tocón de Quercus faginea.

Nephroma laevigatum Ach. — CC, 2, Ar. 172/95, 383/95, 400/95, Quercus ilex; 2, Ar.

182/95, Quercus pyrenaica; 4, Ar. 1016/95, Fraxinus angustifolia.

Normandina pulchella (Borrer) Nyl. — C, 2, Ar. 424/95, 503/95, Quercus ilex.

*Ochrolechia balcanica Vers. — La cita más cercana es de Ciudad Real (Sarrión et al.,

1993). —RR, 2, Ar. 502/95, Quercus ilex.

Ochrolechia pallescens (L.) Massal. —R, 4, Ar. 1021/95, Quercus ilex.

*Pannaria ignobilis Anzi — (Ic. Jorgensen, 1978). Abundante en la zona de estudio.

Aparece en zonas poco alteradas donde existe una elevada humedad ambiental, siempre

sobre cortezas de pH neutro-básico. — CC, 1, Ar. 82/95, Quercus ругепаіса: 2, Ar. | 54/95,

174/95, 549/95, 453/95, Quercus ilex; 2, Ar. 454/95, Quercus faginea; 2, Ar. 178/95, Quercus

pyrenaica; 4, Ar. 910/95, Quercus ilex; 4, Ar. 1009/95, Fraxinus angustifolia.

Pannaria mediterranea Tavares — CC, 2, Ar, 179/95, 412/95, Quercus ilex; 2, Ar. 457/95,

Quercus faginea; 3, Ar. 857/95, Quercus pyrenaica; 4, Ar. 906/95, Quercus ilex.

*Pannaria olivacea P.M, Jorg. — (Ic.: Jorgensen, 1978). Muy escaso en la zona de estudio,

recolectado sobre corteza de Juniperus oxycedrus. Aparece junto a Calicium sp., Parmelia

conspersa y P. pulla, De distribución mediterránea, en la Península Ibérica, se conoce de

Cádiz y Málaga (Arvidsson, 1979) y Portugal (Jorgensen, 1978). — RR, 2, Ar. 166/95,

Juniperus oxycedrus.

Parmelia caperata (L.) Ach. — C, 2, Ar. 375/95, Quercus ilex; 3, Ar. 840/95, Quercus ilex.

*Parmelia conspersa (Ach.) Ach. — C, 2, Ar. 476/95, Juniperus oxycedrus; 4, Ar. 911/95,

madera muerta de Juniperus oxycedrus.

Parmelia exasperata de Not. — C, 2, Ar. 160/95, Quercus ilex; 3, Ar. 867/95, Quercus

ругепаіса.

*Parmelia fuliginosa (Fr. ex Duby) Nyl. — С, 2, Ar. 445/95, Juniperus oxycedrus; 2, Ar.

158/95, Quercus ilex; 2, Ar. 153/95, Quercus pyrenaica.

Parmelia glabra (Schaerer) Nyl. — CC, 2, Ar. 157/95, 390/95, 418/95, 513/95, Quercus ilex;

3, Ar. 844/95, Quercus pyrenaica; 4, Ar. 1022/95, Quercus ilex.

Parmelia pulla Ach. — C, 2, Ar. 473/95, Juniperus oxycedrus; 4, Ar. 1018/95, madera

muerta de Juniperus oxycedrus.

Parmelia quercina (Willd.) Vainio — R, 2, Ar. 162/95, Quercus ilex.

Parmelia saxatilis (L.) Ach. — CC, 2, Ar. 443/95, 479/95, madera muerta de Juniperus

oxycedrus; 3, Ar. 870/95, Quercus pyrenaica.

Parmelia subaurifera Nyl. —R, 3, Ar. 866/95, Quercus pyrenaica.

Source : MNHN, Paris

LIQUINES EPIFITICOS DE LOS MONTES DE TOLEDO (ESPANA) 71

Parmelia sulcata Taylor — CC, 2, Ar. 514/95, Quercus ilex; 2, Ar. 150/95, Quercus

pyrenaica; 3, Ar. 869/95, Quercus pyrenaica.

Parmelia tiliacea (Hofim.) Ach. — CC, 2, Ar. 143/95, 161/95, 376/95, Quercus ilex; 2, Ar.

151/95, Quercus pyrenaica; 2, Ar. 523/95, Taxus baccata; 3, Ar. 856/95, 871/95, Quercus

pyrenaica; 4, Ar. 1019/95, Quercus pyrenaica.

* Parmelia tinctina Maheu & A, Gillet — De carácter silicicola, en estas sierras coloniza la

madera de encbro, de gran dureza y resistencia a la descomposición. Aparece junto con

Parmelia conspersa, P. pulla y Р. saxatilis. — R, 2, Ar. 460/95, madera muerta de Juniperus

oxycedrus,

Parmotrema chinense (Osbeck) Hale & Ahti — R, 3, Ar. 845/95, Quercus pyrenaica.

Peltigera canina (L.) Willd. — CC, 2, HIMM 555, Quercus pyrenaica; 3, HIMM 413, tocön

de Quercus pyrenaica.

Peltigera collina (Ach.) Schrad. — CC, 2, HIMM 549, Quercus ilex; 3, HIMM 418, tocón

de Quercus pyrenaica; 4, Ar. 1000/95, Fraxinus angustifolia.

Peltigera horizontalis (Hudson) Baumg. — R, 2, HIMM 664, base de Quercus ilex.

*Peltigera hymenina (Ach.) Delise — La cita mas cercana es de Segovia (Martinez &

Aragón, 1996). — RR, 2, HIMM 558, base de Quercus ilex.

Peltigera neckeri Hepp ex Müll. Arg. — CC, 2, HIMM 552, Quercus ilex; 2, HIMM 556,

Quercus pyrenaica; 3, HIMM 673, Quercus pyrenaica.

*Peltigera ponojensis Gyelnik — R, 2, HIMM 553, base de Quercus ilex.

Peltigera praetextata (Flörke ex Sommerf.) Zopf — CC, 2, HIMM 559, Quercus ilex: 2,

HIMM 557, Quercus pyrenaica; 3, HIMM 672, Quercus pyrenaica.

Pertusaria albescens (Hudson) M. Choisy & Werner — CC, 2, Ar. 144/95, 379/95, Quercus

ilex; 3, Ar. 863/95, Quercus pyrenaica.

Pertusaria amara (Ach.) Nyl. — CC, 2, Ar. 528/95, madera muerta de Juniperus oxycedrus;

2, Ar. 147/95, Quercus pyrenaica; 3, At. 872/95, Quercus pyrenaica.

*Pertusaria coccodes (Ach.) Nyl. — R, 3, Aragon 864/95, Quercus pyrenaica.

*Pertusaria coronata (Ach.) Th. Fr. — С, 2, Ar. 491/95, Quercus ilex; 2, Ar. 380/95, Quercus

pyrenaica.

Pertusaria flavida (DC.) Laundon — CC, 2, Ar. 492/95, 522/95, Quercus ilex; 2, Ar. 470/95,

madera muerta de Juniperus oxycedrus; 3, Ar. 862/95, Quercus pyrenaica; 4, Ar. 912/95,

madera muerta de Juniperus oxycedrus.

*Pertusaria hemisphaerica (Flörke) Erichsen — C, 2, Ar. 493/95, Quercus ilex; 3, Ar.

874/95, Quercus pyrenaica.

Pertusaria pertusa auct. — CC, 2, Ar. 490/95, 544/95, Quercus ilex; 2, Ar. 452/95, Quercus

faginea; 3, Ar. 861/95, Quercus pyrenaica; 4, Ar. 1030/95, Quercus ilex; 4, Ar. 1031/95,

madera muerta de Juniperus oxycedrus.

* Phaeophyscia orbicularis (Necker) Moberg — R, 3, Ar. 846/95, Quercus pyrenaica.

Source : MNHN, Paris

72 G. ARAGÓN & I. MARTINEZ

Phlyctis argena (Sprengel) Flotow — С, 2, Ar. 218/95, Quercus ilex; 3, Ar. 860/95, Quercus

pyrenaica.

*Physcia adscendens (Fr.) H. Olivier — C, 2, Ar. 163/95, Quercus ilex; 3, Ar. 852/95,

Quercus pyrenaica.

Physcia aipolia (Humb.) Hampe — C, 3, Ar. 889/95, Quercus pyrenaica.

*Physcia biziana (Massal.) Zahlbr. — C, 2, Ar. 214/95, Quercus ilex.

Physcia semipinnata (Gmelin) Moberg — CC, 2, Ar. 165/95, 213/95, 394/95, Quercus ilex;

3, Ar. 855/95, Quercus ilex; 3, Ar. 850/95, Quercus pyrenaica.

Physconia distorta (With.) Laundon — CC, 2, Ar. 552/95, Quercus ilex; 3, Ar. 894/95,

Quercus pyrenaica; 4, Ar. 1044/95, Juniperus oxycedrus.

Physconia enteroxantha (Nyl.) Poelt — CC, 2, Ar. 159/95, 385/95, Quercus ilex; 3, Ar.

892/95, Quercus pyrenaica.

Physconia perisidiosa (Erichsen) Moberg — C, 2, Ar. 398/95, Quercus ilex; 3, Ar. 891/95,

Quercus pyrenaica.

Physconia subpulverulenta (Szat.) Poelt — R, 3, Ar. 890/95, Quercus pyrenaica.

Physconia venusta (Ach.) Poelt. — CC, 2, Ar. 448/95, Quercus faginea; 4, Ar. 1043/95,

Juniperus oxycedrus.

* Polychidium muscicola (Swartz) Gray — Saxícola y asociado a briöfitos, a veces, coloniza

la base de las encinas donde existe acúmulo de humedad. — R, 4, Ar. 1003/95, en la base

de Quercus ilex.

Pseudevernia furfuracea (L.) Zopf — CC, 2, Ar. 403/95, Quercus ilex; 3, Ar. 886/95,

Quercus pyrenaica; 4, Ar. 1024/95, Sorbus torminalis.

Pyrrhospora elabens (Ет.) Haf. — C, 2, Ar. 524/95, madera muerta de Juniperus oxycedrus;

4, Ar. 1006/95, madera muerta de Juniperus oxycedrus.

Ramalina calicaris (L.) Fr. — В. 2, Ar. 123/95, Castanea sativa.

Ramalina farinacea (L.) Ach. — CC, 2, Ar. 181/95, 414/95, Quercus ilex; 3, Ar. 883/95,

Quercus pyrenaica.

Ramalina fastigiata (Pers.) Ach. — CC, 2, Ar. 415/95, Quercus ilex; 3, Ar. 885/95, Quercus

pyrenaica.

Ramalina fraxinea (L.) Ach. — CC, 2, Ar. 122/95, Castanea sativa; 2, Ar. 389195, 535/95,

Quercus ilex; З, Ar. 884/95, Quercus pyrenaica.

*Rhizocarpon obscuratum (Ach.) Massal. De preferencias saxicolas, en la zona de

estudio sorprende su apariciön en las raices descubiertas de algunas encinas. — C, 2, Ar.

1304/95, Quercus ilex.

*Rinodina capensis Hampe in Massal. — RR, 2, Ar. 536/95, en ramas de Quercus ilex.

*Rinodina conradii Körber — (Ic.: Magnusson, 1947). Recolectado sobre briöfitos que

crecen sobre madera en descomposición de Quercus pyrenaica, por lo general, cercanos a

un curso de agua. Su ärea de distribuciön se amplia notablemente, ya que las citas

Source . MNHN, Paris

LIQUINES EPIFITICOS DE LOS MONTES DE TOLEDO (ESPANA) 73

anteriores en la peninsula quedan restringidas al norte de la misma: Navarra (Etayo,

1987), Vizcaya (Renobales & Barreno, 1990) y Portugal (Magnusson, 1947). — RR, 3, Ar.

851/95, Quercus pyrenaica (sobre briöfitos).

*Rinodina exigua (Ach.) Gray — C, 2, Ar. 500/95, Quercus ilex.

Rinodina sophodes (Ach.) Massal. — C, 2, Ar, 203/95, tocön Quercus pyrenaica.

* Saccomorpha icmalea (Ach.) Clauz. & Roux — Coloniza las raices de encina junto con

Rhizocarpon obscuratum. — RR, 2, Ar. 1305/95, en raices de Quercus ilex.

* Trapeliopsis flexuosa (Fr.) Coppins & Р. James — С, 2, Ar. 471/95, madera muerta de

Juniperus oxycedrus.

*Usnea glabrescens (Vainio) Vainio — В. 2, Ar. 417/95, Quercus ilex.

Usnea subfloridana Stirton — C, 2, Ar. 541/95, Quercus ilex.

*Usnea wasmuthii Räsänen — С, 1, Ar. 1289/95, Quercus pyrenaica.

* Waynea adscendens Rico — (Іс.: Rico, 1991). Taxon muy frecuente, recolectado siempre

sobre troncos de encina centenarios, у en zonas húmedas (bordes de rios, valles cerrados),

orientados al norte, donde existe una mayor influencia oceánica. Siempre se ha recogido

en estado estéril. En la Península Ibérica sólamente ha sido citado de Jaén (Rico, 1991). —

CC, 2, Ar. 423/95, 495/95, Quercus ilex; 4, Ar. 905/95, 1005/95, Quercus ilex.

Xanthoria parietina (L.) Th. Fr. — C, 2, Ar. 152/95, 483/95, Quercus ilex.

DISCUSIÓN

Del análisis de los resultados, hay que señalar que la mayoría de las especies

recolectadas son de amplia distribución. El elemento mejor caracterizado es el oceánico-

suboceánico, representado por Collema nigrescens, C. subflaccidum, C. subnigrescens,

Degelia plumbea, Dendriscocaulon umhausense, Lobaria amplissima, L. pulmonaria, L.

scrobiculata, Nephroma laevigatum, Normandina pulchella y Peltigera collina. Todas ellas

ligadas a la presencia de una elevada humedad atmosférica. Dentro de este grupo

distinguimos algunos táxones de distribución atlántico-mediterránea: Degelia atlantica,

Pannaria ignobilis y Р. mediterranea. La gran abundancia de estas especies, refleja la

existencia de una apreciable influencia atlántica sobre la zona de estudio.

El elemento europeo-montano aparece representado por numerosas especies

que, hasta ahora, sólo se conocían del norte de la Península:Calicium abietinum, C.

glaucellum, C. salicinum, Megalaria laureri, Micarea bauschiana, M. denigrata y Rinodina

conradii. Sin embargo, su presencia en la zona es escasa.

El elemento submediterráneo está representado por Lecanora intumescens,

Ochrolechia balcanica, Parmelia glabra, P. quercina, P. tiliacea, Physcia biziana, Ph.

semipinnata, Physconia subpulverulenta y Ph. venusta.

El elemento mediterráneo presente en la zona está representado por especies

como: Buellia cedricola, Coelocaulon crespoae, Koerberia biformis, Pannaria olivacea у

Waynea adscendens, que tienen en comün exigir una elevada humedad ambiental.

Se puede observar la existencia de un gradiente creciente de humedad, desde las

localidades más orientales hasta las más occidentales. Esto se traduce en un incremento de

la abundancia de ciertos liquenes, desde la localidad 1 a la 4 (Fig. 1), especialmente de

Source : MNHN, Paris

74 G. ARAGON & I. MARTINEZ

aquellos que poseen unos requerimientos hidricos mäs acusados (eg. Degelia, Lobaria,

Nephroma).

Debido a las particulares condiciones climaticas e hidricas de los Montes de

Toledo, nos encontramos ante una gran riqueza floristica, tanto de flora vascular como

liquénica. Esta flora está constituida por una mezcla de especies tipicamente mediter-

räneas y de otras de procedencia centroeuropea y atlantica, adaptadas a ambientes de

mayor humedad. Esta riqueza y diversidad floristica, hacen de los Montes de Toledo un

lugar con un gran valor biolögico, que los hacen merecedores de medidas adecuadas de

protección.

AGRADECIMIENTOS -Queremos agradecer a los Drs. A. R. Burgaz (Madrid, España), J. Etayo

(Navarra, España), E. Timdal (Oslo, Noruega), L. Tibell (Uppsala, Suecia) y a Dña. Inés Ibáñez

(Madrid, España) la revisión de algunas de las especies más conflictivas. Agradecemos también la

financiación de este trabajo a la Junta de Castilla-La Mancha, Consejería de Agricultura y Medio

Ambiente.

BIBLIOGRAFÍA

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CARBALLAL R., LOPEZ DE SILANES M.E., BAHILLO L. & ALVAREZ J., 1995 — Recopila-

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nica 13 (1): 279-294.

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ETAYO J., 1987 — Algunos líquenes epífitos de la montaña de Navarra. Anales del Jardin botánico de

Madrid 44 (1): 23-29.

ETAYO J., 1988 — Liquenes epífitos y hongos liquenicolas interesantes de Navarra (España).

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Source : MNHN. Paris

raid dl su din at

іклаў? quina

Cryptogamie, Bryol. Lichénol. 1997, 18 (1) : 77 т

TRES NUEVAS ESPECIES DE HONGOS LIQUENIZADOS

DE LA PROVINCIA DEL CABO (SUDAFRICA)

Correccion

J. M. EGEA & Р TORRENTE

Departamento de Biologia Vegetal (Botanica), Facultatd de Biologia,

Universidad de Murcia, Campus de Espinardo, Е-30071 Murcia, España

El nombre Arthonia capensis Egea & Torrente es un homonimo posterior de

Arthonia capensis Stizenb. (Lichenaea africana 1891, 2 : 212), por esto, debe ser rechazado

y renombrado como Arthonia brussei.

Arthonia brussei Egea & Torrente nom. nov.

Arthonia capensis Egea & Torrente, Cryptogamie, Bryol. Lichénol. 17 : 306 (1996), nom.

illeg. Typus : South Africa, Province del Cabo, west Coast national Park, Postberg, 50 m,

17.2.1994, J. M. Egea (PRE holotypus ; MUB isotypus).

El nombre lo hemos escogido en honor a Brusse (Pretoria, South Africa).

Source : ММНМ Paris

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Зе З нан

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Cryptogamie, Bryol. Lichénol. 1997, 18 (1) : 79-81 79

ANALYSES BIBLIOGRAPHIQUES

Denis LAMY

Lab. Cryptogamie, 12 rue Buffon, F-75005 Paris

CHURCHILL S.P, LINARES C.E.L. — Prodromus bryologiae novo-granatensis —

Introducción a la flora de musgos de Colombia. Santafé de Bogota. Museo de Historia

Natural, 1995, 2 vol. 1-453, i-xxvi ; 455-954[926]. [Instituto de Ciencias naturales. Museo

de Historia natural biblioteca «José Jeronimo Triana» n? 12] (éd. : Instituto de Ciencias

naturales, Museo de Historia natural, Universidad Nacional de Colombia ; ISBN 958-

608-115-X).

Afin de mieux situer leur ouvrage les auteurs présentent un historique des recherches

bryologiques en Colombie depuis J.C. Mutis au хут? s., suivi de données sur les herbiers colombiens

et de généralités sur les mousses (diversité, phytogéographie, écologie ; clé aux familles). Le traitement

taxonomique proprement dit comprend pour chaque taxon : description et illustration, écologie,

distribution, références bibliographiques. Afin d'aider au maximum les débutants en bryologie, les

auteurs donnent un glossaire illustré (pp. 844-863), la liste des collecteurs en Colombie avec informa-

tions biographiques et lieux de récoltes (pp. 865-878), la liste des auteurs des noms de genres (pp.

879-887), une bibliographie (pp. 889-897), et un index (pp. 899-924).

TIBELL L. — Caliciales. Flora Neotropica Monographs 1996, 69 : 1-78 [79], ill. (aut. :

Dept. Systematic Botany, Uppsala University, Villavagen 6, S-75236 Uppsala ; éd. : New

York Botanical Garden, Bronx, New York 10458, USA, ISBN 0-89327-393-7).

L'ordre des Caliciales est représenté par 51 especes (réparties en 17 genres) dans les

néotropiques. Ce sont principalement des espéces lichénisées, mais quelques espéces sont parasym-

biontiques, d'autres sont saprophytes ou parasites de lichens ou de plantes vasculaires. La plupart des

especes ont une dispersion passive des spores et un mazaedium bien développé, c'est-à-dire que les

spores müres s'accumulent sur ou dans les organes de la fructification. Le thalle des espéces

concernées varie de bien développé et dactyliforme, comme celui d' Acroscyphus, à inclus dans le

substrat chez quelques genres crustacés. L'ordre est considéré comme non monophylétique.Chaque

genre et espéce est décrit. La chimie, l'écologie et la distribution sont précisées pour chaque espèce.

BLOM H.H. — A revision of the Schistidium apocarpum complex in Norway and Sweden.

Bryophytorum Bibliotheca 1996, 49 : 1-333, ill. (aut. : Dept. Bot., Norwegian Univ. Science

and Technology, N-7055 Dragvoll ; éd. : Borntraeger Verl., Johannesstr. 3A, D-70176

Stuttgart, ISSBN 3-443-62021-3, prix : 140 DM).

L'auteur a révisé les espèces du complexe Schistidium apocarpum en Norvège et en Suède sur

la seule base de la morphologie du gamétophyte et du sporophyte. Il considère comme discriminants

les caractères suivants: structure de la cellule de la lame foliaire, enroulement de la marge de la feuille,

structure du mucron, forme de Гигпе, modèle de la cellule exotheciale et péristome. La réduction de

la structure, notamment dans le sporophyte, apparaît comme une tendance majeure dans l'évolution

Source : MNHN, Paris

80 ANALYSES BIBLIOGRAPHIQUES

de ce complexe. L'auteur considére que dans une méme touffe, deux formes différentes peuvent étre

des espéces distinctes bien que Іа proximité des pieds puissent faire envisager des croisements ou des

formes transitoires. Aussi, il décrit 31 espéces, 2 sous-espéces et 2 variétés appartenant au complexe

Schistidium apocarpum en Norvége et en Suede, là ou Bremer en 1981 n'en distinguait que quatre! Les

35 taxons sont répartis en 5 groupes informels. Des clés précèdent le traitement taxonomique dans

lequel il est donné pour chaque taxon : description et illustration, distribution et liste des spécimens

examinés. Les nouveautés taxonomiques sont les suivantes : Schistidium grandirete, S. lancifolium

(Kindb.) comb. et stat. nov. (75. pseudo-rivularis subsp. |.), S. trichodon var. nutans, S. confuscum, S.

frisvollianum, S. poeltii, 5. pulchrum, S. umbrosum (Zett.) comb. et stat nov. (= Grimmia apocarpum Г.

и.), S. recurvum, S. robustum (Nees & Hornsch.) comb. nov. (=Grimmia r.), S. scandicum, S.

subjulaceum, S. frigidum, S. fr. var. havaasii, S. venetum, S. crassipilum, 5. elegantulum, 5. e. subsp.

wilsonii, S. submuticum, S. s. subsp. arcticum. Bibliographie (pp. 320-326), Index (pp. 327-333).

WIGGINTON M.J. and GROLLE R. (supplemented by GYARMATI A.) — Catalogue

of the Hepaticae and Anthocerotae of Sub-Saharan Africa. Bryophytorum Bibliotheca

1996, 50 : 1-267, ill. (aut. : 36, Big Green, Warmington, Petersborough PE8 6TU, UK ;

éd. : Borntraeger Verl., Johannesstr. 3A, D-70176 Stuttgart, ISSBN 3-443-62022-1, prix :

110 DM).

Les auteurs présentent dans ce catalogue la liste des 39 anthocérotes et 706 hépatiques

présentes en Afrique hors Maroc, Algérie, Tunisie, Libye et Egypte (les iles de ГОсёап atlantique du

Cap Vert à Sainte-Héléne ont été incluses dans l'étude, mais les iles du groupe de Tristan da Cunha et

celles du Sud et du Sud-Est de l'Afrique du Sud en ont été exclues en raison de Гайтие de leur flore

avec celle des Antipodes et des régions subantarctiques). La synonymie compléte de chaque taxon a

été indiquée ; ainsi 2228 taxons sont cités dans le catalogue. Pour chaque taxon sont donnés le nom

adopté et au moins une référence pour chaque pays ой il a été récolté. Trois nouveaux synonymes sont

proposés : Radula multiramea Gola syn. de R. voluta Tayl., Prionolejeunea fissistipula Steph. syn. de

Drepanolejeunea vesiculosa (Mitt.) Steph., et Pycnolejeunea decurrens Steph. syn. de Cheilolejeunea

surrepens (Mitt.) E.W. Jones. La liste des abbréviations utilisées pour les pays ainsi qu'un synopsis des

familles et des genres précédent une importante bibliographie (pp. 233-262).

BERGPOR JOHANSSON — Íslenskir mosar. Rödaulmosaztt, tildurmosaztt, glitmo-

satt, faxmosaztt, breytingar of tegundaskrá. Fjörlrit nattirufradistofnunar 1996, 29 :

1-127, ill. (Náttúrufredistofnun Islands, Hlemmi 3, Posthölf 5320, 125 Reykjavik, Island).

L'auteur continue la publication de la flore des mousses d’Islande en donnant l'illustration,

la description, la distribution 4/1 espèce d'Entodontaceae, 6 espèces d’Hylocomiaceae, 11 espèces de

Plagiotheciaceae, et 12 espèce d’Hypnaceae. Des additions et un index des 354 taxons d tudies

complétent ce fascicule.

GRADSTEIN S.R. and MELICK H.M.H. van — De Nederlandse Levermossen en

Hauwmossen — Flora en verspreidingsatlas van de Nederlandse Hepaticae еп Anthocerotae.

Utrecht, Stiching Uitgeverij van de Koninklijke Nederlanse Natuurhistorische Verenigun.

1996, 1-368 p., 140 pl., 126 cartes (aut. : Systematisch-Geobotanisches Institut, Univ.

Göttingen, D-73073 Göttingen ; éd. : Dutch Bryological and Lichenological Society ;

ISBN 90-501 1-089-4, Prix : D.Fl. 75,-).

Aprés la publication de la De Nederlandse Bladmossen en 1989, voici la flore des hépatiques

des Pays-Bas. Fruit d'une collaboration de bryologues (A. Aptroot, A.C. Bouman, H.J. During, H.C.

Greven, A.C.A.M. van der Pluym, УУ. Rubers, Н.М. Siebel, Ph. Soliman, J.W. van Vliet, E.J. Weeda

et В.О. Van Zanten), cet ouvrage contient des clés aux familles, genres et espéces, ainsi que la

description, l'illustration, la distribution de 50 hépatiques et 2 anthocérotes, répartis en 31 familles.

Source : MNHN. Paris

ANALYSES BIBLIOGRAPHIQUES 81

Une introduction avec des données sur la morphologie, l'anatomie, la classification, la distribution et

la conservation des hépatiques en Hollande, ainsi qu'un glossaire illustré (pp. 29-42), rendent cet

ouvrage accessible aux chercheurs avertis comme aux débutants. Bibliographie (1p.), index (6p).

REDFEARN PL. Jr., TAN B.C. and SI HE — A newly updated and annotated checklist of

Chinese mosses. Journal of the Hattori Botanical Laboratory 1996, 79 : 163-357 (Dept.

Biol. Southwest Miss. State Univ., Springfield, MO 6584, USA).

Liste de 4832 noms de mousses publiés (parmi lesquels 2457 sont reconnus) et récoltés en

Chine avec les provinces concernées, jusqu'en 1994. Brachythecium propinnatum est proposé pour Br.

pinnatum Tak. hom. Шер. et, Pylaisa curviramea Dix. et Р. subimbricata Broth, et Par. sont transférés

sous Pylaisiella. Une bibliographie de 497 titres complete cette liste.

ANDREEV M., KOTLOV Y. and MAKAROVA I. — Check list of lichens and lichenico-

lous fungi of the Russian Arctic. The Bryologist 1996, 99(2) : 137-169 (Dept. Lichenology

and Bryology, Komarov Botanical Institute, Prof. Popov Str. 2, 197 376 St.Petersburg,

Russia).

И Les auteurs présentent la liste de 1102 taxons (1078 espèces, 7 sous espèces, 17 variétés)

répartis en 192 genres et présents dans la région arctique russe, sur la base de leurs récoltes et de

Panalyse de la bibliographie (300 references).

MATZER М. — Lichenicolous Ascomycetes with Fissitunicate asci on foliicolous lichens.

Mycological Papers 1996, 171 : i-x, 1-202, 97 fig. (Institute for Botany, Karl-Franzens-

Universität Graz, Holteigasse 6, A-8010 Graz ; éd. : CAB International, Wallingford,

Oxon OX10 8DE, UK, ISBN 0-85198-993-4, ISSN 0027-5522).

Une étude taxonomique des ascomycetes lichénicoles non lichénisés, ayant des asques

fissituniquées et se développant obligatoirement sur des lichens foliicoles, a été menée par l'auteur.

Ces champignons occupent une niche spécialisée requérant une adaptation physiologique et métabo-

lique aussi bien que des morphologies spéciales. Les taxons décrits appartiennent à quinze genres

dont huit sont exclusivement lichénicoles. 49 espèces et 2 sous-espèces sont décrites tandis que 17

espéces sont exclues. Onze stades anamorphes de téléomorphes sont connus. Description de deux

genres nouveaux (Lanatosphaera (Dothideales) et Paradoxomyces (Arthoniales, Arthoniaceae)) et de

36 espéces nouvelles. Cing nouvelles combinaisons sont proposées, et Trichophyma est réinstallé

comme genre autonome. Il faut remarquer la description d'espéces lichénicoles non lichénisées

appartenant aux genres Enterographa et Mazozia. L'auteur fait aussi un survol des espéces lichéni-

coles connues avec des catathécies. Une liste des lichens hótes avec le(s) champignon(s) lichénicole(s)

associé(s) complète cette étude. Bibliographie (pp. 187-197), index (pp. 198-202).

Source : MNHN. Paris

amend

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INSTRUCTIONS AUX AUTEURS

Cryptogamie, Algologie, Cryptogamie, Bryologie-Lichénologie et Cryptogamie, Mycologie publient

des articles originaux ou des articles de synthése en systématique, biologie et écologie des

eryptogames (algues, bryophytes et lichens, champignons, respectivement). Les manuscrits rédigés

en francais, en anglais, en allemand, en espagnol et en italien, sont pris en considération dans la

limite des possibilités de la Rédaction à trouver des lecteurs compétents dans ces langues. Les

auteurs n'écrivant pas dans leur langue doivent, avant soumission, faire relire leur texte par une

personne dont la langue du manuscrit est la langue maternelle. П n'y a pas de limite au nombre

de pages. La publication d'un article sera facilitée si les auteurs suivent soigneusement les

instructions ci-après.

Les articles sont lus et critiqués par des rapporteurs spécialistes des domaines concernés. Ces

rapporteurs sont choisis par la rédaction. La publication définitive des articles dépend de la

rédaction. Les manuscrits non conformes aux instructions qui suivent sont retournés aux auteurs

pour étre corrigés, avant leur soumission au Comité de Lecture. Dés leur acceptation, les droits

de reproduction des articles deviennent la propriété de la revue.

TEXTE. — Les manuscrits doivent étre fournis en triple exemplaire (l'original et deux copies),

dactylographiés sur une seule face, en double interligne (y compris les références et les légendes),

sans rature ni surcharge, sans mots coupés et avec une marge gauche de 4 cm. Pour raccourcir

les délais de parution, il est vivement recommandé d'envoyer á la rédaction la version finale de

l'article enregistrée sur disquette utilisable sous DOS (IBM) ou Macintosh ; le logiciel MS WORD

est conseillé.

Chaque manuscrit, paginé, devra comporter, dans l'ordre :

— une page comportant le titre de l'article et son éventuelle traduction en anglais comprendra les

noms, prénoms et adresses des auteurs (y compris, dans la mesure du possible, les numéros de

télécopie et de courrier électronique), l'auteur à qui la correspondance doit étre adressée devant

¿tre précisé ; le titre courant (haut de page) de 50 signes au maximum ; une liste de mots-clés ;

— deux résumés, l'un en français (en cas de nécessité, la rédaction peut aider à sa traduction),

l'autre en anglais, chacun d'eux d'environ 180 mots ou 15 lignes (non limitatif) et, éventuellement,

un troisiéme dans la langue de l'article ; ils doivent faire ressortir les résultats essentiels exposés

dans le texte ;

— le texte proprement dit, suivi des légendes des figures, des planches et des tableaux, dans cet

ordre.

La présentation du texte devra faire apparaitre clairement les subdivisions appropriées

à la nature de l'article (par ex. Introduction, Matériels et Méthodes, Résultats, Discussion.

Remerciements, Références), leur hiérarchie, ainsi que le début des paragraphes. Les symboles,

unités et la nomenclature doivent suivre les normes internationales : le systéme SI, en particulier,

doit être utilisé. La première mention du nom latin d'une espèce doit être accompagnée des noms

des autorités taxinomiques. Un fascicule du journal pourra étre consulté à titre d'exemple pour

une plus ample information sur les conventions suivies par la revue. Les légendes des figures et

des tableaux (dans la langue du manuscrit et en anglais) doivent étre explicites.

RÉFÉRENCES. — La liste bibliographique devra se faire par ordre alphabétique des auteurs et

chronologique par auteurs sans tenir compte des auteurs secondaires. Les titres des périodiques

devront étre cités en entier ; les ouvrages seront cités selon F.A. Stafleu & R.S. Cowan, 1976...,

Taxonomic literature. Ed. 2. Utrecht/Antwerpen : Bohn, Scheltema & Holkema. Les références

devront étre présentées selon les modéles suivants (noter que les noms des auteurs Chinois sont

écrits en entier et dans le style du changement national effectué en Chine en 1987 ; par ex. Chang

С.Е. devient Zhang Jungfu) :

Source - MNHN. Paris

AJISAKA Т., NORO Т., TRONO Jr. G.C., YOUNG-MENG CHIANG & YOSHIDA T., 1994 — Several

Sargassum species (subgenus Sargassum) in East Asia with fureately branching leaves. In: Abbott LA. (ed.),

Taxonomy of Economic Seaweeds. IV. La Jolla, California Sea Grant College, University of California, pp. 9-22.

ALBRECHT А. & REISE К., 1994 — Effects of Fucus vesiculosus covering intertidal mussel beds in the

Wadden Sea, Helgoländer Meeresuntersuchungen 48 (2-3): 243-256.

MAGGS СА. & HOMMERSAND M.H., 1993 — Seaweeds of the British Isles. 1 Rhodophyta, Part ЗА

Ceramiales. London, HMSO Books, The Natural History Museum, 464 p.

MONTAGNE C., 1838 — Centurie des plantes cellulaires exotiques nouvelles, Annales des Sciences Naturelles,

Botanique, ser. 2, 9: 38-57

Les renvois à la liste bibliographique se feront par le nom de l'auteur et l'année de publication ;

utiliser “er al.” lorsque l'article est signé par plus de deux auteurs et “&" entre deux auteurs (par

ex. Montagne, 1838 ; Maggs & Hommersand, 1993; Ajisaka er al., 1994).

ILLUSTRATIONS. — Toutes les illustrations doivent étre montées sur papier blanc rigide ; les

noms des auteurs et le numéro des figures doivent étre indiqués au verso de chacune d'elles. Toutes

les illustrations au trait doivent étre originales, à l'encre de Chine, les numéros et les lettres doivent

être effectués à l'aide de lettres de transfert. Les dimensions des originaux ne devront pas excéder

le triple de celle de leur reproduction définitive (justification de la revue : 12,5 x 18 cm). Les

illustrations au trait et les tableaux, doivent étre des originaux de qualité suffisante pour la

reproduction directe en offset. Les auteurs choisiront l'épaisseur des traits et la taille des caractères

en fonction de la réduction éventuelle des illustrations. Les illustrations photographiques doivent

étre fournies en trois exemplaires originaux (ou un original et deux photocopies laser). Les

documents photographiques doivent étre montés par planches, à la taille de leur reproduction

finale (justification maximale : 12,5 x 18 cm). Toutes les figures devront comporter les échelles

(les grandissements x... sont prohibés), les symboles nécessaires à leur compréhension, et étre

numérotées dans l'ordre d'appel dans le texte. La publication de planches en couleurs est à la

charge des auteurs.

ÉPREUVES. — Les épreuves devront étre vérifiées et retournées à la rédaction dans les 48 heures

suivant leur réception, par courrier et, si nécessaire, par fax. A ce stade, les possibilités de

correction sont limitées aux fautes de frappe. Si des modifications aux illustrations s'avéraient

nécessaires, le coüt du clichage des nouveaux originaux serait à la charge des auteurs. Si les

épreuves ne sont pas retournées dans les délais, l'article paraitra sans les corrections des auteurs.

TIRÉS-À-PART : limités à 150, dont 25 gratuits. Un bon de commande pour les tirés-à-part est

joint à l'envoi des épreuves.

Cryptogamie

Association des Amis des Cryptogames

(ADAC), Muséum National d’Histoire Natu-

relle, Laboratoire de Cryptogamie (PC), 12, rue

Buffon 75005 Paris France

Directeur de publication : Helene Causse-

Bischler

Cryptogamie, Algologie - Rédacteur : Bruno de

Reviers

Tél. (33) 1 40 79 31 98 - Fax (33) 1 40 79 35 94

- E-mail reviers@mnhn.fr

Cryptogamie, Bryologie-Lichénologie - Rédac-

teur : Denis Lamy

Tél. (33) 1 40 79 31 84 - Fax (33) 1 40 79 35 94

- E-mail lamy@mnhn.fr

Cryptogamie, Mycologie - Redacteur : Bruno

Dennetière

Tél. (33) 1 40 79 31 87 - Fax (33) 1 40 79 35 94

- E-mail eryplich@mnhn.fr

Source : MNHN. Paris

INSTRUCTIONS TO AUTHORS

Cryptogamie, Algologie; Cryptogamie, Bryologie-Lichénologie; and Cryptogamie, Mycologie pub-

lish original papers, and reviews, on the systematics, biology and ecology of cryptogams (algae;

bryophytes and lichens; and fungi, respectively). Manuscripts written in French, English, German,

Italian and Spanish are considered providing suitable referees fluent in the language of the

manuscript are available. Authors not writing in their first language should have manuscripts

checked for grammar and syntax by a suitable person before submission. There are no page limits

for papers. Publication will be facilitated if authors check carefully that the manuscript and

illustrations meet the requirements outlined below.

Papers are reviewed by referees in whose field the paper lies. The choice of reviewers is at the

discretion of the Editor. Final responsibility for the publication of papers rests with the Editor.

Manuscripts that do not conform to instructions which follow will be returned for correction prior

to review. On acceptance, papers become the copyright of the journal.

TEXT. — Manuscripts should be submitted to the Editor in triplicate, including the original copy.

All parts of the manuscript must be typed double-spaced (including references and legends), on

one side of the paper, with left margin of 4 cm. To minimize delays in publication, authors should

send the final version of their manuscripts on floppy disks (5 1/4, 3 1/2) using the DOS (IBM) or

Macintosh format, In addition, the use of MS WORD would be appreciated.

Each manuscript should include, in order:

A title page containing the title of the paper and, if necessary, its English translation; the

complete name and address of each author (including, as much as possible, fax number and e-mail

address), and author to whom correspondence should be sent; a running title of less than

50 letters, and a list of key words;

— Two abstracts, the first in French (if necessary, some help can be provided by the Editorial

Office), the second in English (a third one in the language of the text is accepted), each of no more

than 180 words or 15 lines, summarizing the major results of the paper;

The main text, followed by references, legends for figures, and tables, in that order.

The text should normally be divided into sections (e.g. Introduction, Materials and

Methods, Results, Discussion, Acknowledgements, References) appropriate to the nature of the

paper, Indent the first line of all paragraphs. Symbols, units and nomenclature should conform

to international usage. The SI system should be used throughout. The first mention of the Latin

name of a species in the text should be accompanied by the nomenclatural authorities, Consult

the current issue of the Journal for style of headings, sub-headings and other conventions.

Legends for figures, plates and tables should be self-explanatory, and written in the language of

the text and in English if it is not the language used for the text.

REFERENCES. — References should be arranged alphabetically and then chronologically by

author. Journals titles should be cited in full; and books, cited according to Ғ.А. Stafleu & R.S.

Cowan, 1976..., Taxonomic literature. Ed. 2. Utrecht/Antwerpen: Bohn, Scheltema & Holkema.

Conventions of style are provided in the following examples (note that names of Chinese authors

are written in full and listed in the style in accordance with the national change made in China

in 1987; e.g. Chang C.F. becomes Zhang Junfu):

AJISAKA T., NORO T., TRONO Jr. G.C., YOUNG-MENG CHIANG & YOSHIDA T., 1994 — Several

Sargassum species (subgenus Sargassum) in East Asia with furcately branching leaves. In: Abbott LA. (ed.),

Taxonomy of Economic Seaweeds. IV. La Jolla, California Sea Grant College, University of California, pp. 9-

ALBRECHT А. & REISE K., 1994 — Effects of Fucus vesiculosus covering intertidal mussel beds іп the

Wadden Sea. Helgoldnder Meeresuntersuchungen 48 (2-3): 243-256.

Source : MNHN. Paris

MAGGS С.А. £ HOMMERSAND М.Н., 1993 — Seaweeds of the British Isles. 1 Rhodophyta. Part ЗА

Ceramiales. London, HMSO Books, The Natural History Museum, 464 p.

MONTAGNE С., 1838

Botanique, ser. 2, 9: 38-57.

Centurie des plantes cellulaires exotiques nouvelles. Annales des Sciences Naturelles,

In the text, references should be cited by the name of the author and the year of publication; use

“et al”, for more than two authors, and use "&" between two authors (e.g. Montagne, 1838;

Maggs & Hommersand, 1993; Ajisaka el al., 1994).

ILLUSTRATIONS, — All illustrations should be mounted on white light-weight carbocard, with

author's names and figure numbers on the verso. Each line drawing should be original, clearly

drawn with black India ink, and of good quality; letters and numerals must be made with a

lettering device (not with a type writer), Originals should not be more than three times the size

of the final reduction (12.5 х 18 cm). Both drawings and tables should be ready for direct

reproduction by offset. Authors should choose very carefully the thickness of lines, and character

sizes, corresponding to final reduction, . Three copies of all photographic illustrations (or an

original and to laser photocopies) are required. Photographic figures should be ready for

same-size reproduction: maximum page size is 12.5 х 18 cm. АП figures should include scale bars

and symbols necessary for their understanding, and they should be numbered consecutively,

according to the order cited in the text. Color plates will be published only at the authors expense.

PROOFS. — Proofs should be checked and returned by airmail (if necessary, by fax) to the Editor

within 48 hours of receipt. At this stage, corrections should be restricted to those arising from

typesetting errors. If changes to illustrations are necessary at proof stage, new originals would

have to be supplied and the cost for re-photographing will be charged to author at the discretion

of the Editor. If proofs are not returned promptly, the article will be published without the

author's corrections,

REPRINTS: not more than 150, of which 25 are free copies. Reprint orders are enclosed with

proofs.

Cryptogamie

Association des Amis des Cryptogames

(ADAC), Muséum National d'Histoire Natu-

relle, Laboratoire de Cryptogamie (PC), 12, rue

Buffon 75005 Paris France

Director of the journal: Héléne Causse-Bischler

Cryptogamie, Algologie - Editor: Bruno de

Reviers

Tél, (33) 1 40 79 31 98 - Fax (33) 1 40 79 35 94

= E-mail reviers@mnhn.fr

Cryptogamie, Bryologie-Lichenologie - Editor:

Denis Lamy

ТӘ. (33) 1 40 79 31 84 - Fax (33) 1 40 79 35 94

= E-mail lamy@mnhn.fr

Cryptogamie, Mycologie - Editor: Bruno Den-

netiere

Tél. (33) 1 40 79 31 87 - Fax (33) 1 40 79 35 94

= E-mail cryplich@mnhn.fr

Commission paritaire 15-9-1981 - N° 58611 - Dépôt légal 1* trimestre 1997 - Imprimerie Е. Paillart

Sortie des presses le 21 février 1997 - Imprimé en France

liteur

President :

Trésorier

A.D.A.C. (Association des Amis des Cryptogames)

D. Lamy; Secrétaire : В. Dennetiére

В. de Reviers; Directeur de la publication :

Н. Causse

Source - MNHN. Paris

Société Francaise de Systématique <a

La Société Frangaise de Systematique réunit les systématiciens ou les personnes

intéressées par la Systématique et les informe en publiant un Bulletin. Elle convie ses

membres a des colloques annuels transdisciplinaires, au cours desquels les systematiciens

et d'autres scientifiques peuvent s’exprimer et débattre.

Cotisation annuelle: 100F

Demande d'adhésion à adresser au:

Secrétariat de la Société Française de Systématique, 45 rue Buffon, F-75005 Paris.

CCP 7-367-80 D PARIS

La Société édite aussi la série Biosystema.

Prix TTC du Biosystema (France, Etranger): 150 FF, membre SFS : 100 FF.

Biosystema 1 - Introduction à la systématique zoologique - (Concepts, Principes, Méthodes) par

L. Matile, P. Tassy & D. Goujet. 1987.

Biosystema 2 - Systématique Cladistique - Quelques textes fondamentaux. Glossaire. Traduction

et adaptation de D. Goujet, L. Matile, P. Janvier & J.P. Hugot. 1988

Biosystema 3 - La systématique et l'évolution de Lamarck aux théoriciens modernes. par

S. Lovirup. 1988.

Biosystema 4 - L'analyse cladistique: probléme et solutions heuristiques informatisées, par

M. d'Udekem-Gevers. 1990.

Biosystema 5 - Les introuvables de J.B. Lamarck- Discours d'ouverture du cours de zoologie et

articles du Dictionnaire d' Histoire naturelle. Edition préparée par D. Goujet. 1990,

Biosystema 6 - Systématique et Ecologie, par R. Barbault, Cl. Combes, F. Renaud, N. Le Brun

& A. Dubois, Edition coordonnée par J.P. Hugot. 1991.

Biosystema 7 - Systématique et Biogéographie Historique. Textes historiques et

méthodologiques. Traduction et adaptation de P. Janvier, L. Matile & Th. Bourgoin.

1991.

Biosystema 8 - Systématique et Société. Edition coordonnée par G: Pasteur. 1993,

Biosystema 9 - Les Monocotylédones, par J. Mathez. 1993.

Biosystema 10 - Systématique botanique : problémes actuels. Edition coordonnée par O. Poncy.

1993.

Biosystema 11 - Systématique et Phylogénie: modéles d'évolution biologique. Edition

coordonnée par P. Tassy et H. Lelievre. 1994.

Biosystema 12 - Phylsyst: logiciel de reconstruction phylogénétique, par [. Bichindaritz,

5. Potter & В. Sigwalt +. 1994.

Biosystema 13 - Systématique et Biodiversité. Edition coordonnée par Th. Bourgoin. 1995.

Biosystema 14 - Systématique et Informatique. Edition coordonnée par J. Lebbe, en preparation.

Le Conseil de la SFS, XII 1995

Source : MNHN Paris

SOMMAIRE

Harald KURSCHNER — An annotated, corrected, and updated list of the

bryo-logical literature of Southwest Asia

J, MARTÍNEZ-ABAIGAR, E. NUNEZ-OLIVERA, A. GARCIA-ALVARO

and N. BEAUCOURT — Additions to the bryophyte flora of la сар

and the Iberian System (Northern Spain)

Valeria SPAGNUOLO, Paolo CAPUTO, Salvatore COZZOLINO, Rosa

CASTALDO and Paolo DE LUCA — Length polymorphism in the

intragenic spacer 1 of the nuclear ribosomal DNA of some Pottiaceae

(Pottiales, Musci).

Gregorio ARAGON & Isabel MARTINEZ — Contribuciön al conocimiento

de los líquenes epifiticos de los montes de Toledo (Toledo, España) .

J. M. EGEA & P. TORRENTE — Tres nuevas especies de hongos liqueni-

zados de la provincia del Cabo (Sudafrica) — Correccion...........

Analyses bibliographiques ...

Instructions aux auteurs

Instructions to the authors ...............................212 110 еее.

Cryptogamie, Bryologie-Lichénologie 1997, 18 (1) : 1-86

TÉ

МИН.