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Zeitschrift für Zoologie

Supplement 29

Revision of the eastern Palearctic species

of Dusona Cameron

(Insecta, Hymenoptera, Ichneumonidae, Campopleginae)

Rolf Hinz (t) & Klaus Horstmann

Verlag Dr. Friedrich Pfeil » München, 31. Dezember 2004 +» ISSN 0177-7424 » ISBN 3-89937-057-0

Revision of the eastern Palearctic species of Dusona Cameron

(Insecta, Hymenoptera, Ichneumonidae, Campopleginae)

Rolf Hinz (t) & Klaus Horstmann

Hinz, R. (t) & K. Horstmann (2004): Revision of the eastern Palearctic species of

Dusona Cameron (Insecta, Hymenoptera, Ichneumonidae, Campopleginae). — Spix-

iana Suppl. 29: 1-183

The eastern Palearctic species of Dusona Cameron are revised. A key and descrip-

tions are provided for 173 species. 68 species are described as new. The following

new synonymies are indicated: Dusona americana (Ashmead, 1890), syn. D. oyamadai

Hinz, 1994; D. annexa (Förster, 1868), syn. Campoplex limnobius Thomson, 1887;

D. bicoloripes (Ashmead, 1906), syn. C. foersteri Roman, 1942; D. circumcinctus

(Förster, 1868), syn. Campoplegidea melanocerus Uchida, 1932, ? syn. Delopia nigri-

femur Sonan, 1929; D. glauca caliginosa (Walley, 1940), syn. D. latisulcor Hinz, 1994;

D. mercator (Fabricius, 1793), syn. Campoplex oxyacanthae Boie, 1855, syn. C. meso-

xanthus Förster, 1868; D. meridionator Aubert, 1960, syn. D. kyushui Hinz, 1994;

D. pugillator (Linnaeus, 1758), syn. D. amictor Hirz, 1979, syn. D. scutellatoides Hinz,

1985. A lectotype is designated for C. foersteri Roman.

Address of junior author: Dr. Klaus Horstmann, Lehrstuhl Zoologie II, Biozen-

trum, Am Hubland, D-97074 Würzburg, Germany.

Introduction (by the junior author)

Dusona Cameron is a species-rich genus, and it is

well represented in the eastern Palearctic region.

This region is defined as Russia east of the Ural

Mountains (Sverdlovsk = Yekaterinburg Province

included), Iran, Kazakhstan, the central Asian

states, Mongoliya, northern China (as defined by

Yu & Horstmann 1997: 7), Korea and Japan. No

material was available from Afghanistan. The 173

species treated here may represent half of the

species existing in that region. The fauna of south-

eastern Sibiria (from Chita Province to Primorye,

but only the southern parts of those districts), the

southern Kuril Islands (Kunashir, Shikotan) and

Japan is documented best, 80 per cent of the spe-

cies living there may well be covered in this pa-

per. The fauna of the central Asian states and of

Mongoliya is less well known, and very few spec-

imens were available from northern Sibiria (prob-

ably with a poor fauna anyway), Iran, northern

China and Korea.

We did not succeed in subdividing the genus

into species groups. It is easy to recognize a basal

group characterized by plesiomorphies (described

as Kartika by Gupta & Gupta 1976) or some spe-

cialized groups (as the D. mercator group defined

by Hinz 1977, which nevertheless might be poly-

phyletic), but it was impossible to sort all species

into well defined groups. There are too few corre-

lations between character states and too many

intermediate species with a high intraspecific var-

iability. This is also demonstrated by the many

exceptions and cross-references noted in the key.

Therefore the species are arranged alphabetically.

Because the determination of Dusona speci-

mens is difficult, the specimens should be careful-

ly mounted, and all parts of the body must be

clearly visible. The antennae have to be stretched

forwards or sidewards, so that the frons can be

studied, including the dorsal parts of the antennal

carina. The junction of the genal carina with the

oral carina must be visible at one side of the head,

in cases of doubt the head must be turned or a

palpus must be removed. The wings have to be

stretched upwards, and the legs downwards (es-

pecially the front legs), so that all parts of the

prepectal carina, amesopleurum, a metapleurum

and the propodeum can be studied. The gaster

should be visible from both sides, and the ovipos-

itor sheaths should be in their normal position.

Specimens without at least one complete antenna

and one complete hind leg are often indetermina-

ble. A high variability in all characters has to be

taken into account. In some frequent species this

variability is referred to in the key and the de-

scriptions, but in rare species it is unknown. As in

many difficult groups of Ichneumonidae, a deter-

mination may remain uncertain without the op-

portunity to compare specimens with determined

material.

Sexual dimorphism is not as important in Du-

sona as in some other genera of Campopleginae,

and usually the determination of males is not

much more difficult than that of females. The

number of flagellar segments is often slightly high-

er in males (by 5-10 per cent), and if the flagellum

is particularly stout or slender in the female, it

usually presents a more intermediate proportion

in the male. The propodeum is often more nar-

rowly and less deeply depressed in the male, with

the longitudinal carinae and the wrinkles more

strongly developed. The gaster is slenderer in

males. In some species, the colour pattern is dif-

ferent between the sexes: yellow marks may be

more extensive in the male (maxillary palpus,

mandible, tegula, coxae and trochanters of front

and mid legs), and if the hind tibia is black in the

female, it may be subbasally marked with whitish

yellow in the male (but not in all species, and

sometimes only in some specimens of a species).

In 1975 Rolf Hinz started working on the taxono-

my of eastern Palearctic Dusona. He first studied

the relevant types and published the results of

these studies rather quickly (Hinz 1979), but then

he had difficulties to complete his work, because

he could not get enough material for a compre-

hensive revision of the genus. He therefore decid-

ed to describe at first only those species which

show distinctive characters and which were rep-

resented in the material by females (Hinz 1979,

1985). Between 1985 and 1987 he sorted the avail-

able material to 180 items and produced a key to

all of these taxa. But he did not complete and

publish this work, probably because he would

have had to abandon his breeding experiments

with Ichneumon species for one or two years, and

he always was much more interested in working

with living animals than in taxonomic studies on

dead material.

He was further discouraged by a misfortune

with one of his papers. In 1979 he had sent a paper

with descriptions of 18 new species to a journal in

Stockholm, but the editors misplaced the type-

script, and after some time he gave it up as lost.

He described three of the species, of which he had

got additional material in the meantime, in 1985

(D. melanator, D. sachalini, D. yamanakai), and in

his notes he synonymized one species with an

already described one (D. oyamadai = D. americana).

He also removed some name labels and type la-

bels, and sent back some of the types to the collec-

tors and/or depositories, where a few of them

cannot be found now (D. melanator, D. oyamadai).

In the autumn of 1993, he received the typescript

back unpublished, with the remark that the jour-

nal had changed its field of interest in the mean-

time. Because he was already too ill at that time to

be able to revise the paper, he sent the unchanged

typescript to E. Diller in Munich, the latter pub-

lished it as quickly as possible, and it came out in

early 1994. For these reasons, this paper (Hinz

1994) represents a state of knowledge from 1979.

Three species are described in it for a second time,

which had already been described in 1985, with

the 1994 descriptions having been prepared be-

fore 1979, and from shorter type series.

When I received the Hinz collection of Dusona

and his notes from Zoologische Staatssammlung

Munich in 1995, I found the eastern Palearctic

material sorted to species (but many specimens of

described species already sent back to their own-

ers), including the types of most of the species

described by Hinz (1979, 1985, 1994), with the

types (named and labelled) of about 40 unde-

scribed new species, and with the material of a

further 30 unnamed species, the latter mostly rep-

resented by males. I also found a list with all

named species (described or undescribed), a pro-

visional key to all items, and lists of localities,

collecting dates, collectors and depositories for

the specimens already sent back, but almost no

descriptions (except those already published).

Because on the one hand it was urgent to send the

types of described species back to their owners,

and on the other hand the preparation of a com-

prehensive revision would not be possible with-

out those types being at hand, I decided to restrict

the work to material already present (except that

in the meantime I received additional material

from museums in Linz and Ottawa) while includ-

ing and describing all species, also those repre-

sented only by males. I revised the key according-

ly, drew up the descriptions, and added explana-

tions, figures and some remarks on the taxonomy.

Those new species already sorted out and named

by Hinz are published with Hinz & Horstmann as

authors. In species not known to or not accepted

and named by Hinz, Horstmann is the only au-

thor.

Hinz (1985: 301 £.) described the new species

D. filator in a paper on Sibirian Dusona species. But

the types originate from Khoper Reserve near

Varvarino/Voronezh Province in European Rus-

sia, therefore the species is not included in this

revision.

The recent studies were supported by many

colleagues: E. Diller (Zoologische Staatssammlung,

Munich) allowed me to remove the Dusona mate-

rial of the Hinz collection from Munich to my

home in Würzburg. R. W. Carlson (National Mu-

seum of Natural History, Washington, D. C.),

A. Freitag and S. Knispel (Musee Zoologique, Lau-

sanne), F. Gusenleitner (Biologiezentrum, Landes-

museum, Linz), J. T. Huber (Biosystematics Re-

search Institute, Agriculture Canada, Ottawa),

D.R. Kasparyan and A. Khalaim (Zoological

Institute, Russian Academy of Sciences, St. Peters-

burg), J. Sawoniewicz (Instytut Biologii, Bialys-

tok), M. R. Shaw (National Museums of Scotland,

Edinburgh), A. Taeger (Deutsches Entomolo-

gisches Institut, Müncheberg), C. Taylor (Natural

History Museum, London), D. B. Wahl (American

Entomological Institute, Gainesville) and K. Yoshi-

zawa and T. Yoshida (Systematic Entomology,

Faculty of Agriculture, Sapporo) sent types and

other material for comparison. D. R. Kasparyan,

A. Khalaim, J. Sawoniewicz, K. Yoshizawa, T. Yo-

shida and D. S. Yu (Biosystematic Information

System, Vancouver) helped with reading or inter-

preting locality labels. A. Hausmann (Zoologische

Staatssammlung, Munich) completed some names

of host species. U. Horstmann (my daughter) and

M. R. Shaw assisted me with the English lan-

guage. I am indebted to them all.

Explanations of terms

Special terms used in the descriptions of Ichneu-

monidae are defined and figured by Townes (1969:

36 ff.). Some additional explanations are given

here. Some definitions (e.g. that of the glymma)

differ slightly from those given by Gupta & Gupta

(1977:5 ff).

Antennal carina (Figs 3-7): Carina surround-

ing the antennal socket. The descriptions refer to

the dorsal part only.

Areolet (Figs 22-25): Its upper side is sessile,

pointed or stalked. It receives the second recur-

rent vein basad of, at, or distad of the middle.

Generally, the shape of the areolet has strong

infraspecific variability. Therefore this character

is not used in the key, and the descriptions do not

give the full variability, but usually refer to the

holotype of the respective species only.

Black lateral stripe (Figs 40-42): A structure

on either side of the 3" gastral tergite in most of

those species in which the epipleurum is not sep-

arated from its tergite (see explanation under epi-

pleurum). It is a horizontal carina on the inside of

the tergite, which is marked with black, the black

colour showing through the tergite. This charac-

ter must be checked on both sides of the tergite,

because the intensity of the black colour may vary,

or its visibility may be obscured by the pigmenta-

tion of the tergite.

Epipleurum (= laterotergite) (Figs 39-43): Lat-

eral part of the 3"% gastral tergite, which is com-

pletely or partly or not at all separated from the

remaining part ofthat tergiteby ahorizontal crease

or bulge (epipleura of other tergites not being

referred to in the descriptions; exception: D. mer-

cator). Usually the crease or bulge is marked with

black. If the epipleurum is separated from its

tergite, it is usually folded in and concealed on the

inside of the latter. If the epipleurum is separated

only on the anterior 0.3 or less of the tergite, it is

sometimes folded back secondarily and pendent

as a rectangular or triangular lobe. These struc-

tures have strong infraspecific variability. If the

epipleurum is not separated from its tergite, the

former horizontal crease is often replaced by a

black lateral stripe, which is well separated from

the ventrolateral edge of the tergite (see explana-

tion under black lateral stripe).

Glymma (Figs 35-36): A large or small groove

or pit on either side of the petiole. A large glymma

is often bordered by a longitudinal carina dorsal-

ly. A small glymma is often situated at the anteri-

or edge of a more or less distinct narrow lateral

area, this area being bordered by fine carinae dor-

sally and ventrally and/or defined by a change in

the sculpture of the surface (coriaceous instead of

smooth).

Mesopleurum: The descriptions usually refer

to the central part only. The speculum and the

depression in front of the speculum are mentioned

separately.

Nervellus (Figs 26-29): It is inclivous, vertical

or reclivous, and it is intercepted near the middle

1 22

Figs 1-2. Head, top view (a: tangential line on eye and temple). 1. Temples strongly constricted, 0.6%x as long as

an eye, tangential lines intersect on the middle of the mesoscutum, ocellar index 0.8 (D. admontina). 2. Temples

scarcely constricted, 0.9x as long as an eye, tangential lines intersect on the apex of the gaster, ocellar index 1.7

(D. capitator).

b a /

3 4

Figs 3-7. Antennal socket and antennal carina, anterolateral view (a: eye; b: frons; c: antennal socket; d: antennal

carina; e: radial wrinkles). 3. Antennal carina low and narrow (D. bellipes). 4. Antennal carina low, widened to

a crescent-shaped smooth plate dorsally (D. angustifrons). 5. Antennal carina raised, narrow (D. sobolicida).

6. Antennal carina raised, its rim bent upwards and widened to an crescent-shaped transversely striate plate (ear-

like) (D. crassiventris). 7. Antennal carina raised, with a transversely striate projection dorsally (nose-like)

(D. pineticola).

or below the middle. Rarely it is not intercepted.

Petiolar suture (Figs 35-38): Suture separat-

ing the 1“ gastral sternite from its tergite. The

suture is often partly or completely obliterated.

Pleural carina: Carina bordering the meta-

pleurum dorsally and posteriorly, thus separating

it from the propodeum.

Prepectal carina (=epicnemial carina) (Figs

16-21): The ventral part lies between its junctions

with the transverse carinae of the prepectus (= epi-

cnemium); the pleural parts extend between their

junctions with the transverse carinae and the an-

terior edges of the mesopleura. In some species

the ventral part of the prepectal carina and the

transverse carinae are raised. In other species ei-

ther the transverse carinae or the pleural parts of

the prepectal carina are more or less obliterated.

Propodeum (= 1“ abdominal tergite) (Figs 30-

34): The propodeum ranges from not depressed

to deeply depressed, the depression being a broad

sulcus or a rather narrow longitudinal furrow.

Independent of the depression, the propodeum is

very incompletely to almost completely areolated.

If very incompletely areolated, the anterior bound-

ary of the area superomedia and some posterior

stubs of the apical transverse carina and/or the

longitudinal carinae are present. These are not

mentioned in the descriptions. If completely areo-

lated, the median longitudinal carinae, parts of

the lateral longitudinal carinae, the basal trans-

verse carina and parts of the apical transverse

carina are present, and only the median part of the

apical transverse carina (the posterior boundary

of the area superomedia) is lacking (exception:

D. sachalini). In an intermediate condition, either

two longitudinal carinae (consisting of parts of

the median longitudinal carinae and the apical

transverse carina) are present posteriorly and me-

dially, or the costulae are complete and the longi-

tudinal carinae are almost completely lacking.

Propodeal spiracle: Measurements of the

length and width are made to the centre of the rim

surrounding the opening.

Punctation: In a dense punctation the punc-

8 95 10 11

13 14 15

Figs 8-12. Genal carina and oral carina, side view (a: mandible; b: gena; c: occiput; d: genal carina; e: oral

carina). 8. Genal carina low and straight ventrally, joining the scarcely raised oral carina distant from the base of

the mandible (D. montana). 9. Genal carina low ventrally, bent forwards, joining the strongly raised oral carina

distant from the base of the mandible (D. tenuis). 10. Genal carina low ventrally, bent forwards, joining the oral

carina at the base of the mandible (D. angustifrons). 11. Genal carina and oral carina both distinctly raised

ventrally, forming a lopsided triangular pyramid, the tip of the latter shifted ventrally and situated behind the

base of the mandible (D. aurita). 12. Genal carina and oral carina both strongly raised ventrally, forming a high

lopsided triangular pyramid, the tip of the latter shifted ventrally and situated behind the base of the mandible

(D. mercator).

Figs 13-15. Genal carina and oral carina, rear view (a: mandible; b: gena; c: occiput; d: genal carina; e: oral

carina). 13. Genal carina straight ventrally, joining the oral carina at an acute angle distant from the base of the

mandible (D. montana). 14. Genal carina bent sidewards ventrally, joining the oral carina at a right angle distant

from the base of the mandible (D. tenuis). 15. Genal carina bent sidewards ventrally, joining the oral carina at a

right angle at the base of the mandible (D. angustifrons).

tures are for the greater part wider than their

interspaces. In a disperse punctation the punc-

tures are for the greater part narrower than their

interspaces. In a rather dense punctation many

punctures are as wide as their interspaces.

Radial wrinkles (Figs 3-7): Wrinkles on the

ventral part of the frons, running upwards and

sidewards from the dorsal part of the antennal

carina.

Spiracular carina: Carina connecting the pro-

podeal spiracle with the pleural carina.

Tangential lines on eyes and temples (Figs

1-2): Imaginary lines defined to describe the slope

of the temples.

Tarsal claws: The hind tarsal claws are de-

scribed as short (about as long as the empodium)

or long (distinctly longer than the empodium).

Thyridium: The thyridium ıs described as cir-

cular (1.0-1.5x as long as wide), oval (about 2.0x

as long as wide) or elongate (more than 2.5x as

long as wide).

Transverse carina (Figs 16-21): Carina sepa-

rating the ventral from the pleural part of the

prepectus (see explanation under prepectal cari-

na).

Definitions of indices

Epipleural index: Ratio of the length of the crease

or bulge separating the epipleurum from the 3°“

gastral tergite to the lateral length of that tergite.

Genal index: Ratio of the length of the oral

carina between its junction with the genal carina

and the base of mandible to the basal width of a

mandible.

16 17 18 20 21

Figs 16-21. Prepectal carina and transverse carina, ventrolateral view (slanted) (a: pronotum; b: ventral part of

the prepectus; c: pleural part of the prepectus; d: ventral part of the prepectal carina; e: pleural part of the

prepectal carina; f: transverse carina). 16. Prepectal carina low, ventral part merging with the pleural part, the

latter complete, transverse carina fine and indistinct (D. montana). 17. Prepectal carina low, ventral part merging

with the pleural part, the latter obliterated dorsally, transverse carina obliterated (D. micrator). 18. Prepectal

carina low, ventral part merging with the pleural part, the latter obliterated dorsally, transverse carina distinct

anteriorly, obliterated posteriorly (D. opaca). 19. Ventral part of the prepectal carina merging with the transverse

carina, both carinae raised, pleural part of the prepectal carina low, obliterated dorsally (D. tenuis). 20. Ventral

part of the prepectal carina merging with the transverse carina, both carinae low, pleural part of the prepectal

carina present medially, obliterated ventrally and dorsally (D. petiolator). 21. Ventral part of the prepectal carina

merging with the transverse carina, both carinae low, pleural part of the prepectal carina obliterated (D. japonica).

22 23 24 25

Figs 22-25. Areolet (a: areolet; b: 2”! recurrent vein). 22. Areolet short-stalked, receiving the 2" recurrent vein

basad of the middle (D. circumcinctus). 23. Areolat pointed, receiving the 2" recurrent vein at the middle

(D. carınifrons). 24. Areolet narrowly sessile, receiving the 2" recurrent vein distad of the middle (D. variator).

25. Areolet stalked, receiving the 2” reccurent vein distinctly distad of the middle (D. admontina).

Figs 26-29. Nervellus (a: nervellus; b: mediella; c: submediella; d: discoidella). 26. Nervellus inclivous, dis-

tinctly intercepted, nervellar index 0.8 (D. infesta). 27. Nervellus vertical, not intercepted (D. angustifrons).

28. Nervellus reclivous, weakly intercepted, nervellar index 0.7 (D. tenuis). 29. Nervellus distinctly reclivous,

intercepted, nervellar index 0.5 (D. xenocampta).

Malar space index: Ratio of the shortest dis-

tance between the rim bordering the mandibular

grouve and the eye (= malar space) to the basal

width of a mandible.

Nervellar index: Ratio of the length of the

upper (anterior) branch of the nervellus (between

its junctions with the mediella and the discoidella)

to the total length of the nervellus.

Ocellar index: Ratio of the shortest distance

between the eye and the hind ocellus to the long-

est diameter of the hind ocellus.

Ovipositor index: Ratio of the visible length

ID)

)

))

33 34

Figs 30-34. Propodeum, top view (a: area basalis; b: area superomedia; c: area petiolaris; d: costula = part of the

anterior transverse carina; e: parts of the posterior transverse carina; f: parts of the median longitudinal carina;

g: parts of the lateral longitudinal carina). 30. Propodeum almost completely areolated anteriorly, carinae

indistinct posteriorly, area basalis stalked, area superomedia bordered by a carina posteriorly (D. sachalini).

31. Propodeum almost completely areolated anteriorly, carinae partly indistinct posteriorly (D. gastator). 32. Pro-

podeum broadly depressed, sulcus with long transverse striae, longitudinal carinae distinct medially and

posteriorly (D. carinifrons). 33. Propodeum scarcely depressed, transversely striate medially and posteriorly,

costulae complete (D. circumcinctus). 34. Propodeum narrowly depressed, with short transverse striae in the

longitudinal furrow, carinae almost completely obliterated (D. meridionator).

of the ovipositor sheaths (seen from the side, in

normal position) to the length of a hind basitar-

sus.

Spiracular index: Ratio of the length of the

spiracular carina (or, if that carina is obliterated,

the distance between the spiracle and the pleural

carina) to the length of the spiracle.

Thyridial index: Ratio of the shortest distance

between the thyridium and ihe anterior edge of

the 2”! gastral tergite to the length of the thyrid-

ium.

Tibial spur index: Ratio of the length of the

shorter to the length of the longer spur of the mid

tibia.

Figs 35-38. 1“ gastral segment, side view (a: glymma; b: narrow lateral area; c: a suture). 35. Gliymma

large, bordered by a longitudinal carina dorsally, a long row of transverse wrinkles in front of the glymma,

petiolar suture distinct (D. nidulator). 36. Glymma small, at the anterior edge of a narzow lateral area, petiolar

suture partly obliterated (D. angustifrons). 37. Glymma obliterated, a narrow lateral area present, petiolar suture

obliterated (D. aemula). 38. Petiole smooth laterally, without glymmae or lateral areas, petiolar suture indistinct

or obliterated (D. auriculator).

oe a

39 40

41 22

[€

une,

Figs 39-43. 2"! and 3"! gastral tergite, side view (a: ventrolateral crease of the 2” tergite; b: ventrolateral crease

of the 3" tergite; c: black lateral stripe of the 2" tergite; d: black lateral stripe of the 3" tergite; e: epipleurum of

the 3" tergite). 39. Epipleurum separated from the 3"4 tergite and folded in, epipleural index 0.8 (D. nidulator).

40. Epipleurum separated from the 3" tergite and folded in anteriorly or folded back and pendent, epipleural

index 0.3, a black lateral stripe parallel with the ventrolateral crease in a short distance (D. perditor). 41. Epipleurum

of the 3" tergite as a small triangular lobe anteriorly, pendent, a black lateral stripe well above the ventrolateral

edge (D. nigridorsum). 42. Epipleurum not separated from the 3" tergite, a black lateral stripe well above the ventro-

lateral edge (D. angustifrons). 43. 2" tergite with black lateral stripes parallel with the ventrolateral edges in a

short distance, epipleurum not separated from the 3" tergite, the latter without black lateral stripes (D. mercator).

AEI

DEI

EDI

HEL

KIS

LAU

LIN

MIN

MOS

NHM

PAR

BET

SAP

SAW

WAR

WAS

ZSM

Abbreviations of depositories

American Entomological Institute, Gaines-

ville 4

Deutsches Entomologisches Institut, Mün-

cheberg

National Museums of Scotland, Edinburgh

Zoological Museum, Helsinki

Institute of Biological Methods of Plant

Protection, Kishinev

Biosystematics Research Institute, Agricul-

ture Canada, Ottawa

Musee Zoologique, Lausanne

Biologiezentrum des Landesmuseums,

Linz

Institute of Zoology, Minsk

Zoological Museum, State University, Mos-

cow

Natural History Museum, London

Museum National d’Histoire Naturelle,

Paris

Zoological Institute, Russian Academy of

Sciences, St. Petersburg

Systematic Entomology, Faculty of Agri-

culture, Sapporo

Sawoniewicz collection, Instytut Biologii,

Bialystok

Museum and Institute of Zoology, Wars-

zawa

National Museum of Natural History,

Washington, D.C.

Zoologische Staatssammlung, Munich

Key to species

Epipleurum of the 3" gastral tergite separated

from the tergite by a crease and folded in, the

crease usually marked with black, epipleural

indexatlesst AI 339) 3 2

Epipleurum separated from the tergite at most

on its anterior 0.4, often not separated at all or

not folded in, the black lateral stripe of the 3"

tergite often situated above its ventrolateral

edge and divergent from the latter (Figs 40-43).

ee A RER IN LE REIHE 76.

Gaster black. Hind tibia yellowish white me-

SE RT er

Gaster more or less marked with red. .......... 7.

Epipleuwallindex SS. nn. senanesesestneeneneee 4.

Epipleural index 0.7-0.8. Glymma present. 5.

Mesopleurum with dense punctation. Propo-

deum with the longitudinal carinae obliterat-

ed medially, the sulcus irregularly wrinkled,

with longitudinal wrinkles posteriorly. Glym-

ma small. 38-43 flagellar segments. Body length

7-9 mm (See also couplets 16 and 168). ..........

RER NE SR ED m melanator Hinz, 1985 ?3

Mesopleurum with fine and disperse puncta-

tion. Propodeum with more or less complete

longitudinal carinae, the sulcus distinctly trans-

versely wrinkled medially and posteriorly.

Glymma lacking. 30-34 flagellar segments.

BodyAlenpth,6- Immer men ne kn

BER LE vidua (Gravenhorst, 1829) 23

31 flagellar segments. Body length 6 mm.

Speculum smooth and shining. Mesopleurum

with fine and disperse punctation. Propodeum

slightly depressed, the longitudinal furrow co-

riaceous anteriorly and medially, with some

wrinkles posteriorly, the longitudinal carinae

almost completely obliterated. d unknown. .

neh snstnansenaht: nigrina Horstmann, spec. nov. ?

35-46 flagellar segments. Body length 7-13 mm.

Mesopleurum with distinct and rather dense

or dense punetation..........nersseuen-caanszensnssrzenner. 6.

Speculum for the greater part smooth and

shining. Propodeum somewhat depressed, re-

ticulate-rugose, with a median longitudinal

keel posteriorly, almost completely areolated,

but the carinae covered with wrinkles. 45-46

flagellar segments. Body length 11-13 mm. ...

Ban re aiendelaen: myrtilla (Desvignes, 1856) ?3

Speculum for the greater part coriaceous. Pro-

podeum not or scarcely depressed, without

longitudinal carinae medially, with fine and

irregular wrinkles anteriorly and medially,

with longitudinal wrinkles posteriorly. 35-37

flagellar segments in the ?, 38-39 segments in

the S. Body length 79 mm. ........................

. viduator Hinz & Horstmann, spec. nov. ?d

Antennal carina more or less raised dorsally

and/or frons depressed above the carinae, the

carina therefore distinctly protruding (Figs 5-7).

ee ee ee ken 8.

Antennal carina not modified dorsally, nar-

row and low, not distinctly protruding (Fig. 3).

Rn 0 N En ende: 12.

Frons scarcely depressed, antennal carina

raised, its rim bent upwards (ear-like). Epipleu-

ral index 0.3-0.5 (If frons scarcely depressed

and epipleural index greater, see couplet 12).

ee A N ee ER 9.

Frons deeply depressed above the antennal

carinae, the latter distinctly raised. ............ 10.

10.

11.

12.

Antennal carina distinctly raised dorsally, its

rim distinctly bent upwards (ear-like), the frons

with distinct longitudinal wrinkles, which

more or less reach the anterior ocellus. Hind

femur completely black. 53-57 flagellar seg-

ments. Body length 15-16 mm. ................

En er infesta (Förster, 1868) 23

Antennal carina not much raised (variable),

the frons directly above the carina somewhat

depressed and with short longitudinal wrin-

kles, which do not reach the anterior ocellus.

Hind femur often marked with red apically.

52-58 flagellar segments. Body length 14-

17 mm (See also couplets 37 and 97). .............

RI pugillator (Linnaeus, 1758) 25

Epipleural index 0.4-0.5. Body length 15-19 mm.

BE EEE TEASER ER TER 94.

RR E E LOLE LER D ESEREH TE HEREHE TÖTET I}:

Frons very deeply depressed (funnel-like), the

depression distinctly rugose. Mesopleurum

with coarse punctation. Epipleural index 0.8.

Ventrolateral crease of the 3" gastral tergite

marked with black. Postpetiole marked with

red laterally. All gastral tergites marked with

black dorsally. 56 flagellar segments. Body

length 12mm unknown...

infundibulum Horstmann, spec. nov. d

Frons distinctly depressed (but not funnel-

like), the depression coriaceous. Mesopleu-

rum with fine punctation. Epipleural index

0.6. 3’! and 4" gastral tergites red. Black lateral

stripe of the 3"! tergite situated above its ven-

trolateral edge. 60 flagellar segments. Body

lensth4l5:mm. unknown. en.

ee linearis Horstmann, spec. nov. d

Temples straight, strongly constricted behind

the eyes, the tangential lines on eyes and tem-

ples intersect on the centre of mesoscutum.

Genal index about 0.9 (Both characters less

distinctive in the 8). Area basalis partly re-

placed by a longitudinal keel. Area superome-

dia bordered by a carina posteriorly (Fig. 30).

45-48 flagellar segments. Body length 9-10 mm.

ner sachalini Hinz, 1985 25

Temples less strongly constricted behind the

eyes and/or more strongly rounded. Genal

index smaller. Area basalis sessile, as long as

or broader than long, often indistinct or re-

duced. Area superomedia not bordered by a

CAriMa POSTENIONY- mean eeenseeeesnssennamnergenenen 13.

13.

14.

15:

16.

117,

18.

Only the 2"! and 3" gastral tergites marked

with red. Glymma present... 14.

Gaster more extensively marked with red, at

least the 4" tergite marked with red laterally

eo ger oa ee re ee 18.

53-58 flagellar segments. Body length 12-

16 mm. Hind femur usually red or marked

with red. Hind tibia yellowish red, sometimes

darkened basally. Propodeum deeply de-

pressed, the sulcus distinctly transversely stri-

ate. Epipleural index 05 0.8...

ee cultrator (Gravenhorst, 1829) 23

34-48 flagellar segments. Body length 5-11 mm.

Hind femur black. Propodeum slightly de-

pressed, with fine irregular wrinkles, often

with longitudinal wrinkles posteriorly. .... 15.

46-48 flagellar segments. Body length 10-

11mm. Hind tibia black. Propodeum almost

completely areolated. Epipleural index 0.9.

? unknown (See also couplet 54). ...................

Bene obscurator Horstmann, spec. nov. d

34-43 flagellar segments. Body length 5-9 mm.

Carinae of the propodeum for the greater part

lacking. Epipleural index 0.3-0.7. ............... 16.

38-43 flagellar segments. Body length 7-9.

Metapleurum distinctly punctate on a coria-

ceous background. Epipleural index 0.3-0.4.

Hind tibia yellow medially (See also couplets

Aramel6ß)e...e melanator Hinz, 1985 23

34-35 flagellar segments. Body length 5-7 mm.

Metapleurum coriaceous and with fine wrin-

kles, not distinctly punctate. Epipleural index

U ee eekkkerrcca 17.

Tegula yellow. Front femur yellowish red.

Hind tibia yellow medially. 34-35 flagellar seg-

ments. Body length 5-6 mm. d unknown. .....

En chechziri Hinz & Horstmann, spec. nov. ?

Tegula black. Front femur partly black. Hind

tibia black. 35 flagellar segments. Body length

7 mm. unknown. nennen

Lennnanesagee nigrotibialis Horstmann, spec. nov. ?

Genal carina obliterated ventrally close to its

junction with the oral carina. Propodeum

slightly depressed, almost completely areo-

lated anteriorly, carinae often indistinct poste-

riorly (Fig. 31). Epipleural index 0.9. Gaster

scarcely compressed posteriorly. 35-36 flagel-

lar segments. Body length 7-8 mm. d unknown.

VERLEIH En ER gastator Hinz, 1985 2

Genal carina complete ventrally. Gaster often

more compressed posterionly. een 19.

19.

20.

21.

22.

24.

Mesopleurum rugose centrally, without dis-

tinct punctation. Propodeum not depressed,

with distinct longitudinal carinae. Epipleural

index 0.9. 38-42 flagellar segments. Body length

7-8 mm. ........ leptogaster (Holmgren, 1860) 24

Mesopleurum distinctly punctate centrally,

somelmestügose-punttate. rn... 20.

Gaster from the apex of the 2”! tergite on-

wards red, the posterior tergites sometimes

brown dorsally, but not distinctly black. ..21.

Gaster more extensively darkened, usually at

least the posterior tergites distinctly black dor-

Sy nennen 24.

27 flagellar segments. Body length 7 mm. Flag-

ellum slender basally, clavate subapically, the

broadest segment 1.7x as wide as the apical

end of the 1* segment. Tegula black. Gliymma

present. Ovipositor index 1.1. d unknown. ...

ee iwatae Hinz, 1994 2

At least 34 flagellar segments. Flagellum at-

tenuate subapically. Ovipositor index 0.5-0.6.

neuen en aharegn 22.

58-63 flagellar segments, 2” segment 1.5x as

long as wide. Body length 11-15 mm. Meso-

pleurum with fine and rather dense puncta-

tion, the interspaces slightly coriaceous. Petio-

lar suture present only anteriorly. Glymma

very small or lacking. d of this variety un-

known (See also couplet 45). ...................0.00.

De frage similator Hinz, 1994 var. ?

Number of flagellar segments lower, 2" seg-

ment at least 1.9x as long as wide. Body length

Tl en Bl AMSSEN an 23)

Mesopleurum densely and finely rugose-punc-

tate, the interspaces coriaceous, rather dull.

Hind femur red or brown in the d, dark brown

orblack in the 2. Petiolar suture distinct. Glym-

ma lacking. Posterior gastral tergites marked

with brown dorsally. 34-39 flagellar segments.

Bodyienetn7 Im eeeeee

EEE SEINE admontina (Speiser, 1908) 25

Mesopleurum with fine and disperse punc-

tures and shining interspaces. Hind femur

black. Petiolar suture scarcely developed.

Glymma present. Posterior gastral tergites

completely red. At least 42 flagellar segments

(Both flagellar tips broken off in the only avail-

able specimen). Body length 11mm. ? un-

known. .... gastroides Horstmann, spec. nov. d

Frons on either side with a small raised point

between the antennal socket and the eye. .....

EURE 25,

29.

26.

27.

28.

29,

Frons without such raised points. ............. 27%

Hind femur black, marked with red apically.

Hind tibia yellowish red to red. Propodeum

somewhat areolated, broadly but not deeply

depressed between the longitudinal carinae,

the sulcus for the greater part transversely

wrinkled. Glymma small. 2”! gastral tergite

red on the posterior 0.4-0.6. 3" and 4" tergites

red, the 5" tergite red in the 9, for the greater

part black in the d. 54-55 flagellar segment.

Bodylenetn M-13 mme 2.2 nee

IE bellipes (Holmgren, 1872) ?3

Hind femur completely black. Hind tibia black

or marked with black. Gliymma large. 2" gas-

tral tergite red on the posterior 0.1-0.3. ..... 26.

Propodeum not depressed or scarcely de-

pressed, irregularly wrinkled, often with one

or more longitudinal wrinkles posteriorly.

Hind tibia yellowish brown to dark brown or

black medially. 5" gastral tergite often red or

marked with red. 45-55 flagellar segmentes.

Body»length IP mm. er art ne

ee bicoloripes (Ashmead, 1906) 24

Propodeum slightly depressed, for the greater

part irregularly wrinkled, with distinct trans-

verse wrinkles posteriorly. Hind tibia black.

5" gastral tergite black. 46-55 flagellar seg-

ments. Body length 9-13 mm. ....................

chabarowski Hinz & Horstmann, spec. nov. 93

Epipleural index 0.4-0.7. Epipleurum of the 3"4

gastral tergite folded in, rarely folded back

secondarily and pendent, in the latter case the

former ventrolateral crease present as bulge

close to the ventrolateral edge, the bulge usu-

ally marked with black (in this case similar to

Bie A). venseran. een ass ern enee 28.

Epipleural index 0.7-0.8. Epipleurum folded

INMEISS9). ea @neeneseererensuesalterenedesheheehessteertne 42.

32-43 flagellar segments. Body length 6-9 mm.

MR LEEREN RI EEUREUR ERRLL DES EEE EE EE BERE 29:

46-58 flagellar segments. Body length 9-17 mm.

ER AL nes Sl.

Propodeum distinctly depressed, distinctly

transversely wrinkled in parts of the sulcus.

Gaster red medially, black basally and apical-

ly, the 2”! tergite red on the posterior 0.3-0.5,

the red marks more extended laterally. Hind

tibia yellow or yellowish brown medially, black

basally, marked with brown apically. 41-43

flagellar segments. Body length 8-9 mm. ........

Be a contumax (Förster, 1868) 23

30.

Sl:

32.

33:

Propodeum shallowly or not at all depressed,

not distinctly transversely wrinkled. Gaster

darkened dorsally, red laterally, either the 3"

and 4" tergites marked with black dorsally or

the posterior tergites marked with red ventro-

laterally, the 2" tergite red at most on the

posterior 0.2 dorsally (Propodeum rather nar-

rowly depressed, with short transverse wrin-

kles in the narrow longitudinal furrow; gaster

black dorsally: see D. specularis, couplet 54)..

RE RE 30.

Flagellum almost filiform, the shortest seg-

ments 0.9x as long as wide. Hind tibia light

yellow medially, black basally and apically.

Gaster for the greater part black, 2" to 4" or 5"

tergites marked with red laterally. 37-41 flag-

ellar segments. Body length 7-8 mm. ..............

.... bellator Hinz & Horstmann, spec. nov. ?d

Flagellum distinctly fusiform, the shortest seg-

ments 0.8x as long as wide in the 2. Hind tibia

yellowish brown to dark brown medially, nar-

rowly black basally, scarcely darkened apical-

ly. Posterior gastral tergites black dorsally,

marked with red ventrolaterally. Posterior gas-

tral sternites red. 32-36 flagellar segments.

Body length 8-9 mm. d unknown. .............

ERENTEIER.. JENIE maruyamae Hinz, 1994 9

Frons with a distinct median longitudinal keel.

Pleural part of the prepectal carina lacking. 2”

gastral tergite with a black edge posteriorly.

Epipleural index 0.6-0.7. Hind tibia red, indis-

tinctly darkened basally. 49-51 flagellar seg-

ments. Body length 11 mm. d unknown........

2. m. Armee. dur ARE carinator Hinz, 1985 ?

Frons at most with a fine median longitudinal

carina. Pleural part of the prepectal carina at

least partlyspresent... een 32.

Mesopleurum smooth or scarcely coriaceous

between the punctures, strongly shining. Pleu-

ral part ofthe prepectal carina partly obliterat-

ed, not reaching the anterior edge of the meso-

pleurum. Hind tibia yellowish medially, nar-

rowly black basally, broadly black apically.

2"l gastral tergite black dorsally and posteri-

orly. Epipleural index 0.4-0.5. 54-58 flagellar

segments. Body length 10-13 mm. ...................

BER TEE Ahr yamanakai Hinz, 1985 25

Mesopleurum more or less strongly coriaceous

between the punctures. Pleural part of the

prepectal carina complete and/or epipleural

Index WOW LScereeersnsnenenennenessceresnensarsesrsesseneer 33.

Temples not much constricted behind the eyes,

tangential lines on eyes and temples intersect

34.

39.

36.

37.

on the 1* gastral segment (similar to Fig. 2).

Temple 0.8x as wide as an eye. Hind tibia

yellow with a red apex. Hind tarsus yellow.

Epipleural index 0.5-0.6. 48-51 flagellar seg-

ments. Body length 10-12 mm. ? unknown. .

N flavitarsis Horstmann, spec. nov. d

Temples shorter and more strongly constrict-

ed behind the eyes. Hind tarsus usually black.

BE RE. 34.

Epipleural index 0.3-0.5. Mesopleurum punc-

tured on a distinctly coriaceous background.

Longitudinal carinae of propodeum often

present medially....n......r..e... ee ee 35,

Epipleural index 0.607.222. ee 38.

2" gastral tergite dorsally black at least on the

posterior edge, sometimes tinged with red in

front of the edge. Hind tibia yellowish brown

medially, narrowly black basally and apically.

Epipleural index 0.2-0.3. 45-55 flagellar seg-

ments. Body length 9-12 mm (See also couplet

1 er vigilator (Förster, 1868) 23

2"! gastral tergite red posteriorly. Hind tibia

yellow medially. ............-........0.:...20m ee 36.

Depression in front of the speculum with short

striae, which merge with an narrow area with-

out punctation anteriorly. Propodeum with

complete costulae and longitudinal carinae,

sulcus between the longitudinal carinae finely

coriaceous and shining anteriorly and medial-

ly. Epipleural index 0.4. 50 flagellar segments.

Body length 10 mm. ? unknown (See also cou-

plet 174). 22... E RE

en rufovariata Horstmann, spec. nov. d

Depression in front of the speculum with rath-

er long striae, which merge with the puncta-

tion anteriorly. Costulae and longitudinal car-

inae often incomplete, sulcus between the lon-

gitudinal carinae rugose or transversely wrin-

kled anteriorly and medially. 51-58 flagellar

segments. Body length 12-17 mm. .............. 37.

Frons somewhat depressed above the anten-

nal carinae. Antennal carina often somewhat

raised and its rim bent upwards. Epipleural

index 0.2-0.4. 52-58 flagellar segments. Body

length 14-17 mm (See also couplets 9 and 97).

a. pugillator (Linnaeus, 1758) 23

Frons slightly depressed. Antennal carina low,

its rim scarcely bent upwards. Epipleural in-

dex 0.5-0.7. 51-57 flagellar segments. Body

length 12-14 mm (See also couplet 41). ..........

rugifer (Förster, 1868) ?4

38. Depression in front of the speculum at least

39.

40.

41.

dorsally with distinct striae or wrinkles, which

usually merge with the punctation anteriorly.

Hind tibia yellowish brown to brown medial-

Ü Vers Rennen sentesymnennternenandeenateenweensehennenneekdelh 3%

Depression in front of the speculum with fine

striae, which usually merge with a narrow

coriaceous area without punctation anterior-

ly.or,granulate-stuigose.......susueetsnk 44.

Speculum smooth or almost smooth, strongly

shining. Mesopleurum slightly coriaceous and

shining between the punctures. Metapleurum

with rather dense or disperse punctation,

slightly coriaceous and shining between the

punctures. Propodeum distinctly depressed,

the sulcus for the greater part transversely

wrinkled, the longitudinal carinae present

medially. Gaster distinctly shining posterior-

ly. 42-46 flagellar segments. Body length

10-12 mm (See also couplets 46 and 64). ........

RE LEE RER EN polita (Förster, 1868) ?3

Speculum slightly coriaceous, shining. Meso-

pleurum distinctly coriaceous and rather dull

between the punctures. Metapleurum with

dense or rather dense punctation, coriaceous

between the punctures, sometimes rugose-

punctate. 48-57 flagellar segments. Body length

Id Asa een ed 40.

Propodeum broadly and not deeply depressed,

sulcus for the greater part irregularly wrin-

kled, with transverse wrinkles only posterior-

Iy, longitudinal carinae completely lacking

medially. Tegula yellow. Hind tibia yellowish

medially, narrowly black basally, broadly black

apically. 51-52 flagellar segments. Body length

13-14 mm. d tentatively placed here. ..............

DE rugosa Horstmann, spec. nov. ? ?d

Propodeum rather broadly depressed, sulcus

for the greater part transversely wrinkled, lon-

gitudinal carinae present medially and poste-

riorly. Tegula brown or black in the 9, but

often yellow. inithe on nen 41.

Hind tibia broadly yellow medially, narrowly

brown basally and apically. 51-57 flagellar seg-

ments. Body length 12-14 mm (See also cou-

pPeiBaerre rugifer (Förster, 1868) 25

Hind tibia yellowish brown to dark brown

medially, narrowly black basally, brown to

black apically, the dark apical mark some-

times lacking. 48-52 flagellar segments. Body

leHErNE O1 anany 2.10. ee ieeseneeransnene

Be hrnchdnnne nen stragifex (Förster, 1868) 23

22.

43.

45.

46.

34-42 flagellar segments. Body length 6-7 mm.

Flagellum for the greater part light brown ven-

trally. 2" flagellar segment 2.0-2.4x as long as

wide. Propodeum scarcely depressed, for the

greater part coriaceous, with longitudinal

wrinkles posteriorly. Glymma distinct. ..........

N maruyamator Hinz 1979 28

At least 39 flagellar segments. Body usually

longer. In cases of doubt flagellum black and /

or propodeum with transverse wrinkles pos-

FeNIOn ya een Besen 43.

Depression in front ofthe speculum with dense

and fine striae, which usually merge with a

narrow coriaceous area without punctation

anteriorly, or granulate-strigose, ventrally of-

ten coriaceous without striae. ................ 44.

Depression in front of the speculum at least

dorsally with distinct striae or wrinkles, which

merge with the punctation anteriorly, also ven-

trallyroftenwithrstnaetmnene nr ar 47.

. Propodeum distinctly and rather narrowly

depressed, with longitudinal carinae border-

ing the longitudinal furrow, transversely wrin-

kled almost everywhere between the carinae,

area superomedia only coriaceous. Hind tibia

yellowish brown medially, broadly black ba-

sally and apically. 2" gastral tergite red on the

posterior 0.6. 3“! and 4" tergites red. 49 flagel-

lar segments. Body length 13 mm. $ unknown.

ne coriacea Horstmann, spec. nov. d

Propodeum broadly and shallowly depressed,

often without longitudinal carinae medially,

transversely wrinkled only in some parts of

the sulcus, in other parts irregularly wrinkled.

2"! gastral tergite red at most on the posterior

WB. rrunlerantenesn 2snasns mess geönen nes ensgBenmuraeosr Prsarmensrrane: 45.

58-63 flagellar segments. Body length 11-14 mm.

Mesopleurum with rather dense punctation.

Hind tibia yellowish brown to brown medial-

ly, narrowly blackish basally, broadly black-

ish apically (sometimes paler in the 4). Propo-

deal spiracle about 4x as long as wide. Glym-

ma very small or lacking (See also couplet 22).

SID RSETN. HIER similator Hinz, 1994 9&

42-55 flagellar segments. Body length 9-13 mm.

Mesopleurum with dense punctation. Propo-

deal spiracle about 3x as long as wide. Glym-

Ma ande, 0 enenteeakeernde.ö2nssnsenasesessenezenreeer 46.

Speculum smooth or almost smooth, strongly

shining. Mesopleurum slightly coriaceous and

shining between the punctures. Metapleurum

with rather dense or disperse punctation,

47.

48.

49.

50.

slightly coriaceous and shining between the

punctures. Propodeum distinctly depressed,

sulcus for the greater part transversely wrin-

kled, longitudinal carinae present medially.

Gaster distinctly shining posteriorly. 42-46 flag-

ellar segments. Body length 10-12 mm (See

alsereouplets BI and/6A). 2... nee

u N polita (Förster, 1868) ?3

Speculum coriaceous, somewhat shining. Meso-

pleurum distinctly coriaceous and rather dull

between the punctures. Metapleurum rugose

or with dense or rather dense punctation, co-

riaceous between the punctures. 45-55 flagel-

lar segments. Body length 9-13 mm. .......... 26.

Petiole strongly sculptured laterally in front

of the glymmae, usually with a long row of

distinct transverse wrinkles, sometimes with

irregular wrinkles. Petiolar suture complete

and distinct. 1% gastral sternite coriaceous an-

teriorly and medially, with fine transverse stri-

ae posteriorly (Fig. 35). Hind tibia yellowish,

not darkened basally, distinctly darkened api-

cally. 50-52 flagellar segments. Body length

12-183 mm 2. nidulator (Fabricius, 1804) 23

Petiole smooth or with fine sculpture laterally

in front of the glymmae, at most with some

transverse wrinkles anteriorly. 1“ gastral ster-

nite often smooth medially. Number of flagel-

lar segments often lower...........uucunrsenesene. 48.

IM ENTE ee u 49.

kemalestw a Pen ee: 61.

Gaster from the 3" tergite onwards black dor-

sally (to a variable extent), red laterally, the

posterior tergites at least half red in lateral

view. Genital claspers for the greater part red.

ER N TER OHONCHEENDERIERTRENE AN 50.

Posterior gastral tergites for the greater part

or completely black. Genital claspers for the

ereaten part black... 2,

47-48 flagellar segments. Body length 11-12 mm.

2" flagellar segment 1.6x as long as wide.

Front and mid coxae yellow apically. Front

and mid trochanters yellow. Hind tibia yel-

lowish, narrowly black basally, not distinctly

darkened apically. Longitudinal carinae of

propodeum scarcely indicated, the propodeum

not depressed, wrinkled, with short longitu-

dinal wrinkles posteriorly. Glymma distinct.

Sankcouple 7. en er

Bauuı momoii Hinz & Horstmann, spec. nov. d

alle

32.

54.

43-44 flagellar segments. Body length 8-10 mm.

Atleast the mid coxa and mid trochanter black.

2"! flagellar segment 2.2x as long as wide.

Front coxa yellow apically. Front trochanter

yellow. Hind tibia yellowish brown, black ba-

sally. Propodeum scarcely depressed, with-

out longitudinal carinae, for the greater part

finely wrinkled, with fine transverse wrinkles

posteriorly. Glymma very small or lacking. 43

flagellar segments. Body length 9-10 mm. ? in

eouplet: 70. ..-....:.4-.24:2=4:22:52:2222008532202082220 0000er

ER, detritor Hinz & Horstmann, spec. nov. d

2"! flagellar segment 1.7x as long as wide.

Front coxa and front trochanter black. Hind

tibia blackish, sometimes tinged with yellow-

ish brown medially. Propodeum not depressed,

rather completely areolated, coarsely and ir-

regularly wrinkled, with distinct longitudinal

wrinkles posteriorly. Glymma distinct. 44 flag-

ellar segments. Body length 8mm. ? in cou-

plet:-7A. en... ee sa ee en

ine pictator Hinz & Horstmann, spec. nov. d

Pleural carina indistinct, covered with wrin-

kles medially. Propodeum shallowly depressed,

almost completely areolated, wrinkled, with

longitudinal wrinkles posteriorly. Hind tibia

black. Glymma present. 2" gastral tergite red

on the posterior 0.2-0.3. 3" and 4" tergites red.

46 flagellar segments. Body length 11 mm. ?

inlcouplel/635... ze... caudator Hinz, 1985 &

Pleural carina distinct and complete medially

(Pleural carina partly covered with wrinkles;

hind tibia yellowish red medially: see D. tere-

brator,.couplet,s9). „„.e.. a2 Se 53:

Gaster black, at most the 3" and 4" tergites red

laterally, the 4" at least half black. ............. 54.

Gaster more extensively marked with red. ...

A EEE co ter oe 39:

Mesopleurum with fine and dense punctation

on a coriaceous background. Speculum finely

granulate-strigose. Hind tibia black. Propo-

deum almost completely areolated, slightly

depressed, with fine irregular wrinkles. Glym-

ma distinct. 46-48 flagellar segments. Body

length 10-11 mm. ? unknown (See also cou-

plet lb). 2 See ee en

re obscurator Horstmann, spec. nov. d

Mesopleurum with distinct and partly dis-

perse punctation on a strongly shining back-

ground. Speculum smooth. Hind tibia broad-

Iy yellowish brown medially, narrowly black

39.

56.

58.

basally, rather broadly dark brown apically.

Propodeum coarsely rugose, the longitudinal

carinae indicated, with a shallowly depressed

longitudinal furrow, the latter with short trans-

verse wrinkles. Glymma small. 40 flagellar

segments. Body length 9 mm. ? unknown. ....

nn specularis Horstmann, spec. nov. d

Glymma lacking, at most a small and shallow

furrow present. 2” flagellar segment 2.2x as

long as wide. Hind tibia rather dark yellowish

brown, black basally, not distinctly darkened

apically. Propodeum not depressed, with or

without longitudinal carinae, in some parts

with fine transverse wrinkles, in particular

posteriorly. 2" gastral tergite red on the pos-

terior 0.2 dorsally, the red marks more extend-

ed laterally. 3“ and 4" tergites red. 43-44 flag-

ellar segments. Body length 11-12 mm. 2 in

eouplet’68:.:.....,..2: montana (Roman, 1929) &

Glymma present. Basal flagellar segments less

slender (Glymma as a very small pit; 58-63

flagellar segments: see D. similator, couplet 45).

ER. er Suse csineerzehuheetncensundslaisernkgsheneet 56.

Genal index 0.9. 2" gastral tergite black or red

on the posterior 0.1. 45-47 flagellar segments.

Body length 12-13 mm. ? in couplet 66. ........

ee Hinsal yezoensis (Uchida, 1928) 3

Genalımde< at mosH 0 ee 5%

2nd flagellar segment 2.0x as long as wide. ....

era a Re RE VRR 58.

2" flagellar segment at most 1.7x as long as

NG Ener a N 59.

Tegula black. Hind tibia black, to a variable

extent yellowish brown medially. Propodeum

elongated, reaching to the middle of the hind

coxa, more or less completely areolated, scarce-

ly depressed, partly transversely wrinkled.

Glymma small. 2" gastral tergite red on the

posterior 0.2. 3" tergite completely, 4" for the

greater partred. 42-45 flagellar segments. Body

length 9-10 mm. 2 in couplet 62. ................

A BE longicauda (Uchida, 1928) 3

Tegula yellow. Hind tibia yellow medially,

black basally, brownish apically. Propodeum

not elongated, more or less completely areo-

lated, somewhat depressed, partly transversely

wrinkled. Glymma distinct. 2” gastral tergite

red on the posterior 0.2. 3" tergite completely,

4" almost completely, 5% Jaterally red. 40-44

flagellar segments. Body length 9-11 mm. 2 in

EOUPIEFOT.N A nern A lbeannensdentanchnaens

ee spinator Hinz & Horstmann, spec. nov. d

59.

60.

61.

62.

63.

64.

Maxillary palpus black. Mandible for the great-

er partor completely black. Tegula black. Meso-

pleurum distinctly coriaceous and dull be-

tween the punctures. Metapleurum rugose,

without distinct punctures. 39-43 flagellar seg-

ments. Body length 9-11 mm. 2 in couplet 66.

EN terebrator (Förster, 1868) &

Maxillary palpus yellow. Mandible for the

greater part yellow. Tegula usually yellow.

Metapleurum often with distincet punctures

on a coriaceous background. ................... 60.

Propodeum not depressed or slightly de-

pressed, for the greater part irregularly wrin-

kled, without transverse wrinkles or with

transverse wrinkles only posteriorly, longitu-

dinal carinae usually obliterated anteriorly and

Meddally-sensx scene net sen ern ierektaenenetee 26.

Propodeum distinctly depressed, with trans-

verse wrinkles at least medially and posterior-

ly, longitudinal carinae present at least medi-

ally and Posteriorly........n..enesnzeersenae een 39

Ovipositor index at least 0.8. ................ 62.

Oyipositor. index 0:5-0.7 en nn 69.

Ovipositor index 1.4-2.2. Ovipositor bent up-

wards. Mesopleurum slightly coriaceous be-

tween the punctures, distinctly shining. ... 63.

Ovipositor index 0.8-1.1. Ovipositor straight.

Mesopleurum distinctly coriaceous between

the punctures, scarcely shining (exception:

Di. Molita)®.... nee eargseneneene: 64.

2rd flagellar segment 1.3x as long as wide, 5"

segment as long as wide. Pleural carina indis-

tinct or lacking. Depression in front of the

speculum with short wrinkles. Ovipositor in-

dex 2.2. 45 flagellar segments. Body length

11mm. Sin,couplet, 32. een eenattessgeronee

en rgengsen anregen caudator Hinz, 1985 ?

2"d flagellar segment 2.3x as long as wide, 14"

segment as long as wide. Pleural carina dis-

tinct. Depression in front of the speculum with

distinct striae. Ovipositor index 1.4. 40-48 flag-

ellar segments. Body length 9-12 mm. d in

couplet 58. ........... longicauda (Uchida, 1929) ?

2" flagellar segment 1.3x as long as wide.

Speculum smooth or almost smooth, strongly

shining. Metapleurum with rather dense or

disperse punctation, slightly coriaceous and

shining between the punctures. Gaster strongly

shining posteriorly. Ovipositor index 0.7-0.8.

42-46 flagellar segments. Body length 10-12 mm

(See also couplets 39 and 46). ..........eeeeeenesennenne.

EEE REN polita (Förster, 1868) ?

117

69.

66.

67.

68.

69.

2"! flagellar segment at least 1.6x as long as

wide ass HEINE 69.

2"4 flagellar segment 1.6-1.9x as long as wide.

Speculum coriaceous, dull. Metapleurum co-

riaceous and rugose. Propodeum completely

or almost completely areolated. ................. 66.

2nd flagellar segment at least 2.1x as long as

N 67.

39-43 flagellar segments. Body length 9-11 mm.

Genal index 0.5. Hind tibia broadly yellowish

brown medially. 2" gastral tergite red on the

posterior 0.5. Ovipositor index 1.0. d in cou-

pie og re terebrator (Förster, 1868) ?

45-47 flagellar segments. Body length 12-13 mm.

Genal index 0.9. Hind tibia black. 2" gastral

tergite black or red on the posterior 0.1. Ovi-

pesitoninde< 0.978 IMeouplei 56%... 22...

een yezoensis (Uchida, 1928) ?

40-41 flagellar segments, the shortest segments

0.8-0.9x as long as wide. Body length 10 mm.

Hind tibia yellowish brown medially, distinctly

darkened basally and apically. Glymma large.

SIMCOUpIEL 38.2... essen

Ber spinator Hinz & Horstmann, spec. nov. ?

43-48 flagellar segments, all segments at least

as long as wide. Hind tibia dark yellowish

brownitorblackt. Aare Area. 68.

The shortest flagellar segments as long as wide.

Hind tibia dark yellowish brown, darkened

only basally. Propodeum often incompletely

areolated. Glymma indicated as a small fur-

row or lacking. 43-45 flagellar segments. Body

lengtn.10-12. 781m eouplet 59 %.2.2..2.2.2ue2...

REISEN N el es montana (Roman, 1929) 2

All flagellar segments longer than wide. Hind

tibia black. Glymma small. 48 flagellar seg-

ments. Body length 13 mm. d unknown. .......

yezoensoides Hinz & Horstmann, spec. nov.

Gaster black dorsally, broadly red laterally, 3°“

tergite black dorsally. 6'" tergite only narrow-

ly black dorsally. Hind tibia yellowish brown,

black basally, scarcely darkened apically.......

EN 70.

Gaster red medially, black anteriorly and pos-

teriorly, 3° and/or 4" tergites usually com-

pletely red, 6" tergite at least half black in

lateral view (D. pictator with a black hind tibia,

which is intermediate in this character, is in-

cluded here). Hind tibia often more darkened,

In’partieular apieally.nenesenseeereeeeeeene IR

70.

Zar

72.

713.

44 flagellar segments. Body length 9-10. 2“

flagellar segment 2.3x as long as wide. Propo-

deum scarcely depressed, without longitudi-

nal carinae, partly with fine transverse wrin-

kles. Glymma very small. d in couplet 51. ....

RR detritor Hinz & Horstmann, spec. nov. ?

48-50 flagellar segments. Body length 11-13 mm.

2"! flagellar segment 1.5x as long as wide.

Propodeum not or scarcely depressed, with-

out longitudinal carinae, irregularly wrinkled,

with a short longitudinal keel posteriorly.

Glymma distinct. d in couplet 50. .................

ee momoil Hinz & Horstmann, spec. nov. ?

Ciypeus small, rounded, its apical edge round-

ed forwards medially. Malar space index 1.1.

Propodeum not depressed, for the greater part

irregularly wrinkled, longitudinal carinae part-

ly indistinct. Subgenital plate black. 40 flagel-

lar segments. Body length 7 mm. d unknown.

Es papator Hinz & Horstmann, spec. nov. ?

Clypeus large, flat, its apical edge truncate or

almost truncate medially. Malar space index

0.3-0.4. 41-60 flagellar segments. Body length

8-17. mm en nee 12.

Mesopleurum slightly coriaceous and shining

between the punctures. Metapleurum with

dense and fine punctation on a rather smooth

background. Hind tibia yellowish brown,

sometimes somewhat darkened basally and

apically. 6" gastral tergite red ventrolaterally.

Subgenital plate red. 56-60 flagellar segments.

Body length 13-17 mm. d unknown. ..............

..sasayamae Hinz & Horstmann, spec. nov.

Mesopleurum and metapleurum distinctly

coriaceous and dull between the punctures,

and/or 41-46 flagellar segments and body

length 8-12 mm. Subgenital plate often black

(58-63 flagellar segments; posterior gastral

tergites and sternites completely black: see

D"similator, couplet 5). ..... ne 73:

Propodeum not depressed or slightly de-

pressed, for the greater part irregularly wrin-

kled, without transverse wrinkles or with

transverse wrinkles only posteriorly, longitu-

dinal carinae usually obliterated anteriorly and

meckally. u: 2.2... 0 Sa ee 74.

Propodeum slightly or distinctly depressed,

with transverse wrinkles at least medially and

posteriorly, longitudinal carinae present at

least medially and posteriorly. Mesopleurum

and metapleurum distinctly coriaceous and

dull between the punctures (42-46 flagellar

74.

73.

7.6.

78.

segments; mesopleurum and metapleurum

slightly coriaceous and shining between the

punctures: see D. polita, couplet 64). .......... 73:

Genal index 0.2-0.3. Mesopleurum and meta-

pleurum more or less shining between the

punctures (variable). 6" gastral tergite red ven-

trolaterally. Subgenital plate red. Gaster not

much compressed posteriorly. 41-44 flagellar

segments. Body length 8-10 mm. 4 in couplet

a a E RNEUE INS NUINERRA FRE OORETERRERER

Genal index 0.5. Mesopleurum and metapleu-

rum distinctly coriaceous and dull between

the punctures. 6" gastral tergite black. Sub-

genital plate usually black. Gaster distinctly

compressed posteriorly. 45-55 flagellar seg-

ments. Body length 9-13 mm. ............ 26.

Hind tibia black. Propodeum slightly de-

pressed, with complete and distinct longitudi-

nal carinae. Subgenital plate red to brown.

45-46 flagellar segments. Body length 9-10 mm.

&unknowanz....... memorator Hinz, 1994 2

Hind tibia yellow to brown medially, dark-

ened basally and apically. Propodeum dis-

tinctly depressed, longitudinal carinae often

obliterated anteriorly. Subgenital plate often

black. 48-57 flagellar segments. Body length

POTT ze nennen anna ranseennennnt nenne 41.

Antennal carina more or less distinctly raised

dorsally, often also distinctly widened and/or

its rim bent upwards, the frons above the car-

inae often depressed (Figs 5-7).................. ZUR

Antennal carina not raised dorsally, formed

either as a narrow low carina or widened as a

low plate or projection, the frons above the

carinae not or only slightly depressed (Figs

a 98.

Genal carina joining the oral carina at or very

close to the base of the mandible, in the latter

case the genal index 0.1 or less (Figs 10, 15)..

re BE 78.

Genal carina joining the oral carina distant

from the base of the mandible, genal index at

leash. 0 24(Fies/8-9, I HA) rel 81.

26-29 flagellar segments. Body length 6 mm.

Shortest flagellar segment 0.7x as long as wide

in the 2, 0.9x as long as wide in the d. Hind

femur partly or completely red or reddish

brown. Ovipositor index 1.3. Ovipositor sheaths

distinetiy widened apically."...........e............

ee alticola (Gravenhorst, 1829) 23

79.

80.

81.

82.

39-50 flagellar segments. Body length 9-10 mm.

Flagellum slenderer. Hind femur black. Ovi-

POSHORBIMIER OB an 73.

50 flagellar segments. Body length 9 mm. Pos-

terior gastral tergites from the 3" tergite on-

wards for the greater part red. Posterior gas-

tral sternites red. d unknown (See also couplet

tod). chinensis Horstmann, spec. nov. $

39-43 flagellar segments. Body length 9-10 mm.

Posterior gastral tergites and sternites black.

ERS ESE RER RR NER EEE RR ENR 80.

Ocellar index about 0.5. Frons with a rather

low median longitudinal carina. Antennal ca-

rina distinctly raised and widened in the g,

less strongly modified in the 2. 2" gastral

tergite red on the posterior 0.1-0.2. 39-41 flag-

ellar segments. Body length 9 mm. % also in

couplet 1032... ucrainator Hinz, 1985 ?3

Ocellar index about 1.0. Frons with a strong

median longitudinal keel, which is distinctly

raised medially. Antennal carina distinctly

raised in both sexes. 2" gastral tergite red on

the posterior 0.2-0.3. 42-43 flagellar segments.

Body;leneth,9-10.mm. .......2.2.. 2...

RR ucrainica Hinz, 1972 23

Genal carina raised close to the junction with

the also raised oral carina, forming a lopsided

triangular pyramid, the tip of the latter shifted

ventrally and situated behind the base of the

mandible (Figs 11-12). Hind tibia yellowish

brown medially, narrowly black basally, broad-

ly black apically. Propodeum broadly and

deeply depressed, distinctly transversely wrin-

kled in almost all parts of the sulcus.......... 82.

Genal carina not distinctly raised, its junction

with the oral carina situated above the base of

the mandible (Figs 8-9, 13-14). .................. 83.

Frons distinctly depressed above the antennal

carinae, the latter distinctly widened dorsally

to a nose-like projection, with fine transverse

striae (Fig. 7). Ventral part of the prepectal

carina merging with the pleural part, the latter

joining the anterior edge of the mesopleurum.

52-54 flagellar segments. Body length 10-13 mm.

enden aurita (Kriechbaumer, 1883) ?3

Frons not much depressed above the antennal

carinae, the latter somewhat raised dorsally,

but not widened, the rim with fine transverse

striae (Fig. 5). Ventral part of the prepectal

carina merging with the transverse carina, both

somewhat raised, the pleural part of the pre-

pectal carina lower and obliterated dorsally.

83.

84.

80:

86.

87.

88.

45-50 flagellar segments. Body length 8-10 mm

(Seeralsoreouplei 128) NAEH R

ne sobolicida (Förster, 1868) 24

Epipleurum of the 3" gastral tergite complete-

ly fused with the tergite (Figs 42-43). ........ 84.

Epipleurum of the 3" gastral tergite to some

extent separated from the tergite by a horizon-

tal crease or bulge anteriorly (Fig. 40). ...... 92.

Antennal carina not much raised. Frons with a

fine median longitudinal carina. Hind femur

sometimes marked with red........................ 85.

Antennal carina strongly raised and/or frons

above the carinae deeply depressed. Hind fe-

Mur black en. .csesnmesonnelefereieesneetanesnsensueknnennu.n 87.

Hind femur completely or partly red. Posteri-

or gastral tergites red or marked with red.

Posterior gastral sternites red. 42-46 flagellar

segments. Body length 8-9 mm (See also cou-

pletilll).s2.. 0: auriculator Aubert, 1964 28

Hind femur black. Posterior gastral tergites

andksternites:blacer Mrs net 86

2"! flagellar segment 1.5x as long as wide.

Propodeum extending to 0.3 of the hind cox-

ae, rather broadly depressed. Petiole with

distinct glymmae. 49-50 flagellar segments.

Body length 11 mm. $ unknown (See also cou-

PlEB2UN er een

uihreneeguheee parvicavata Horstmann, spec. nov. d

2"! flagellar segment 2.5-2.7x as long as wide.

Propodeum extending to 0.5-0.7 of the hind

coxae, with a narrow longitudinal furrow.

Petiole without or with very small glymmae.

40-47 flagellar segments. Body length 9 mm.

Stentatively.placedlhere7..... un...

ehe hsssigregi propodeator Hinz, 1994 $ ?5

Pleural carina usually obliterated or replaced

by wrinkles medially. 2" gastral tergite red on

the posterior 0.1. Antennal carina distinctly

thickended dorsally and produced as a nose-

like projection. Frons distinctly depressed

above the antennal carina. 46-51 flagellar seg-

ments. Body length 11-14 mm. ...............

EUR SE NENEN pineticola sibirica Hinz, 1985 24

Pleural carina complete. 2" gastral tergite usu-

ally more extensively marked with red. ... 88.

Frons scarcely depressed above the antennal

carinae, with distinct radial wrinkles. Anten-

nal carina with a nose-like dorsal projection.

Hind tibia black in the 9, yellow with a black

apex in the d. 2"! gastral tergite red with a

small black dorsal mark anteriorly. 3" tergite

89.

ID.

92%

with black lateral stripes. 41-49 flagellar seg-

ments.Bodylength7- Immer

DER DE Re anceps (Holmgren, 1860) ?3

Frons distinctly depressed above the raised

antennal carinae, with a more or less distinct

longitudinal bulge medially.................... 89.

Frontal bulge with a cone-like projection ven-

trally. Rim of the antennal carina distinctly

bent upwards (ear-like) and transversely stri-

ate. Hind tibia yellow, narrowly darkened ba-

sally and apically. 3 gastral tergite with black

lateral stripes. 56-62 flagellar segments. Body

length 15-17 mm (See also couplet 93). ..........

ERS re ee. mactatoides Hinz, 1994 23

Frontal bulge without a cone-like projection

ventrally: .......... nn 90.

Hind tibia whitish yellow medially, narrowly

black basally, broadly black apically. Anten-

nal carina strongly widened dorsally, with a

transversely striate nose-like projection. 3°“

gastral tergite with black lateral stripes. 56-60

flagellar segments. Body length 11-13 mm. ...

ee REN okadai (Uchida, 1942) Pd

Hind tibia yellow to yellowish brown medial-

ly, narrowly black basally, broadly black api-

cally. 3"! gastral tergite without black lateral

stripes. 61-69 flagellar segments. Body length

16-17mmM. 2... Al,

Rim of the antennal carina distinctly bent up-

wards (ear-like) (Fig. 6). Frons not deeply de-

pressed above the antennal carinae and with

strong radial wrinkles. 1” gastral tergite 2.4x

as long as wide. 61 flagellar segments. Body

length 17 mm. 6 unknown... een

A crassiventris Horstmann, spec. nov. $

Frons deeply depressed above the antennal

carinae (funnel-like), with short wrinkles in

the depressions. 1" gastral tergite about 4.6x

as long as wide. 69 flagellar segments. Body

length 16-14 mm. ..:.....ecm1 208

rg matsumurae (Uchida, 1928) 23

Frons deeply depressed above the antennal

carinae, with a longitudinal bulge or keel me-

dially. Antennal carina strongly raised dorsal-

ly, either with a dorsal projection (nose-like)

(Fig.7) or its rim bent upwards (ear-like)

TS) 93.

Frons slightly depressed above the antennal

carinae, scarcely raised medially, sometimes

with a median longitudinal carina. Antennal

carina distinctly but not strongly raised dor-

Sally aussen snstindenteinsnsmehensnee nen gen rer are 95.

93. Frons between the antennal sockets with a

94.

95.

96.

cone-like projection, which merges with a me-

dian longitudinal bulge dorsally. Rim of the

antennal carina distinctly bent upwards (ear-

like) and transversely striate. Hind tibia yel-

low, narrowly darkened basally and apically.

Black lateral stripe of the 3'% gastral tergite

situated above the ventrolateral crease or

bulge. 56-62 flagellar segments. Body length

15-12 mm! (See also couplet 89)...

N N ER mactatoides Hinz, 1994 ?5

Frons with amedian longitudinal keel or bulge,

without a cone-like projection. Ventrolateral

crease of the 3'4 gastral tergite marked with

Blades... 2222. nennen 94.

Depressions of the frons slightly sculptured

and with few wrinkles. Rim of the antennal

carina somewhat bent upwards (ear-like) and

at the same time somewhat widened (nose-

like), with fine transverse striae on the dorsal

projection. Mesopleurum slightly coriaceous

between the punctures. Speculum slightly co-

riaceous. 2" gastral tergite for the greater part

black. 59 flagellar segments. Body length

II impressifrons Hinz, 1979 23

Depressions of the frons with coarse wrinkles.

Antennal carina distinctly widened dorsally

(nose-like), with fine transverse striae on the

dorsal projection. Mesopleurum smooth be-

tween the punctures. Speculum partly smooth.

2rd gastral tergite for the greater part red.

56 flagellar segments. Body length 15 mm.

Uno WT re eptsaendernngeennensahsredunrehensnee

Messuslakser rugifrons Horstmann, spec. nov. d

42-45 flagellar segments. Body length 8-10 mm.

Rim of the antennal carina shortly but dis-

tinctly bent upwards (ear-like), not widened

dorsally. Frons above the antennal carinae

somewhat depressed and with fine wrinkles.

Hind tibia yellowish brown medially, narrowly

black basally, broadly black apically. Epipleu-

ral index 0.2. Epipleurum of the 3'% gastral

tergite folded in, the black lateral stripe situat-

ed above the ventrolateral crease. d of this

variety unknown (See also couplet 176). .......

EN EEE recta (Thomson, 1887) var. ?

52-63 flagellar segments. Body length 14-

18 mm. Antennal carina more strongly modi-

edge 96.

Antennal carina distinctly raised and widened

dorsally, with a transversely striate nose-like

projection (as in Fig. 7). Frons distinctly de-

pressed above the antennal carinae. Hind tibia

97:

98.

39:

for the greatest part yellow, scarcely darkened

basally and apically. Epipleural index 0.2-0.3.

Epipleurum of the 3“ gastral tergite folded in,

sometimes folded back secondarily and pen-

dent, the black lateral stripe situated above

the ventrolateral crease or bulge. 59-63 flagel-

lar segments. Body length 16-18 mm. .............

a ERS mactator (Förster, 1868) ?3

Antennal carina distinctly raised, its rim some-

what bent upwards (ear-like), not much wid-

ened (variable) (Figs 5-6). Frons above the an-

tennal carinae not much depressed. Epipleu-

ral index 0.3-0.4. Epipleurum of the 3" gastral

tergite folded in, the ventrolateral crease

marked with\black ME Zu 97.

Frons with dense punctation and with long

radial wrinkles, which almost reach the medi-

an ocellus. Mesopleurum shining and almost

smooth between the punctures. Hind tibia

blackish, tinged with yellowish brown subba-

sally. 57-61 flagellar segments. Body length

EIS) NN rss celator Hinz, 1985 23

Frons coriaceous and with short radial wrin-

kles, which extend to 0.3 the distance to the

median ocellus. Mesopleurum distinctly co-

riaceous between the punctures. Hind tibia

yellow medially, narrowly brown or black

basally and apically. 52-58 flagellar segments.

Body length 14-17 mm (See also couplets 9

and SR)Rrnnk pugillator (Linnaeus, 1758) ?3

Antennal carina not much raised, but wid-

ened dorsally to a crescent-shaped plate or a

nose-like projection, this part either smooth or

transversely striate (Fig. 4). Frons not de-

pressed above the antennal carinae. At most

52Ktlagellar segments. Arne nennen 99:

Antennal carina not or scarcely modified dor-

sally, narrow and low, its rim at most some-

what bent upwards (Fig. 3) (58-66 flagellar

segments; rim of the antennal carina widened

to a narrow crescent-shaped plate: see D. scutel-

lator, couplet; 22A)..........rrtssapeansurnszansenzosnee 105.

Pleural part of the prepectal carina obliterated

(Fig. 21). Pleural part of the prepectus with

well separated punctures. 47-51 flagellar seg-

ments. klindilessiblacks.....re.2....% 152.

Pleural part of the prepectal carina at least

partly' present (Figs'16-20).................: 100.

100. Ventral part of the prepectal carina merging

with the transverse carina, the pleural part

lower (Figs 19-20). Hind tibia yellowish to

brown, darkened only basally. Glymma in-

101.

102.

103.

104.

105.

dicated as a small longitudinal furrow. 29-34

flagellar segments. Body length 6-7 mm......

En RER EEE 101.

Ventral part of the prepectal carina usually

merging with the pleural part (Figs 16-18).

Hind tibia black at least apically. 33-49 flagel-

lar segments. Body length 7-10 mm. ..... 102.

Mesopleurum with rather dense punctation.

Hind femur completely black. Propodeum

scarcely depressed, irregularly wrinkled. 32

flagellar segments. Body length 6 mm. 3 un-

known...... minutor Horstmann, spec. nov. ?

Mesopleurum with dense or very dense punc-

tation, partly rugose-punctate. Hind femur

red or brown on the apical 0.2. Propodeum

somewhat depressed, strongly transversely

wrinkled in the shallow sulcus. 29-34 flagel-

lar segments. Body length 6-7 mm (See also

COUPIEHTAI) Mar. nen. anne ianenenreregereransenetanenene seitens

spiracularis Hinz & Horstmann, spec. nov. 24

Genal carina joining the oral carina at the

base of the mandible (Figs 10, 15). Propo-

deum distinctly depressed, with distinct

transverse wrinkles at least in the posterior

half. 2" gastral tergite red on the posterior

GEN 2. 2 en enessegesregsreveetegeeeras szene 103.

Genal carina joining the oral carina distant

from the base of the mandible (Figs 8-9, 11-14).

2"d gastral tergite red on the posterior 0.2-0.6.

Ucsausssnedäcen AussdessnötngrsbassaerngetiHsengesnnTegensdesn gerne 104.

Hind tibia yellowish brown to dark brown or

black medially. 33-38 flagellar segments. Body

length Immer np 160.

Hind tibia whitish yellow medially. 39-40

flagellar segments. Body length 9 mm. 4 in

couplet 80.................. ucrainator Hinz, 1985 ?

Propodeum scarcely or not depressed, irreg-

ularly wrinkled, partly with longitudinal

wrinkles. Hind tibia black. 2” gastral tergite

red on the posterior 0.2-0.3. 48-52 flagellar

segments. Body length 9-10 mm (See also

couplet 234). „un... fundator Hinz, 1990 25

Propodeum somewhat depressed, with dis-

tinct transverse wrinkles at least in the poste-

rior half. 2”! gastral tergite red on the poste-

rior 0.3-0.6. 43-48 flagellar segments. Body

length, 8-1 mmr ta Seren 215:

Ventral part of the prepectal carina merging

with the transverse carina, these carinae of-

ten raised, the pleural part lower or obliterat-

ed at the junction (Figs 19-21). ................ 106.

106.

107.

108.

109.

Ventral part of the prepectal carina merging

with the pleural part, both of equal height at

the junction, the transverse carina at most as

high, often lower or obliterated at the junc-

tion, rarely raised at some distance from the

junction (Figs 16-18) (This character has strong

infraspecific variability, which could not al-

ways be taken into account in the key). .......

Re ER EL RER EN neeRoc-coh: 156.

Hind femur for the greater part or complete-

ly red or reddish brown. Propodeum elon-

gated, reaching to the middle of the hind

coxa (exception: D. carpathica). Glymma very

small’or lacking. ................0...0 107.

Hind femur for the greater part or complete-

ly black, at most lightened apically........ 112.

Genal carina joining the oral carina at the

base of the mandible (at a right angle), the

former somewhat bent sidewards and for-

wards (Figs 10, 15). 6'" gastral tergite black.

unge en ken vn entas gehen en ran nee 108.

Genal carina joining the oral carina distant

from the base of the mandible (at an acute

angle) (Figs 8, 13). 6" gastral tergite for the

greater part or completely red. ............... 110.

All flagellar segments at least as long as wide

in the 2. Depression in front of the speculum

rather irregularly wrinkled dorsally, striate

medially. Glymma completely lacking. 2"“

gastral tergite red on the posterior 0.7. Epi-

pleurum of the 3" gastral tergite separated

from the tergite by a crease anteriorly. 34

flagellar segments. Body length 7 mm. d un-

KNOWAM ie... 20028222000 25aneaennact nneezentneE \

en collaris Hinz & Horstmann, spec. nov. ?

The shortest flagellar segments about 0.8x as

long as wide in the ?. Depression in front of

the speculum usually distinctly striate dor-

sally and medially. Giymma often present as

a small pit or replaced by a lateral area. 2"“

gastral tergite red on the posterior 0.1-0.3.

Epipleurum of the 3“ gastral tergite com-

pletely fused with the tergite. ................. 109.

Ventral part of the prepectal carina slightly

raised submedially, scarcely emarginate me-

dially. Propodeum extending to 0.3-0.4 of the

hind coxae. Ovipositor index 0.6-0.7. 30-36

flagellar segments. Body length 6-8 mm (See

also couplet 122) 2.0. nn

ERETER carpathica (Szepligeti, 1916) 23

Ventral part of the prepectal carina strongly

raised submedially, deeply emarginate me-

110.

ala

12?

113.

dially. Propodeum extending to 0.5-0.7 of the

hind coxae. 26-32 flagellar segments. Body

lenetin Im Haren 118.

Genal index 0.5. Flagellum brown. All flagel-

lar segments longer than wide. Propodeum

not depressed, finely transversely wrinkled

in the posterior half. Glymma lacking. 41-43

flagellar segments. Body length 7 mm. d un-

known... kasparyani Hinz, 1985 2

Genal index at most 0.3. Flagellum black.

Some flagellar segments as long as wide. ...

IE ER ERLERNEN. > NEFFEN 111.

Temples strongly constricted, tangential lines

on eyes and temples intersect on the meso-

scutum. Depression in front of the speculum

with strong striae. Metapleurum densely

punctate on a coriaceous background. Pro-

podeum somewhat depressed, with short

transverse wrinkles in the narrow longitudi-

nal furrow. Glymma lacking. 42-46 flagellar

segments. Body length 8-9 mm (See also cou-

plet 85). u........... auriculator Aubert, 1964 93

Temples less strongly constricted, tangential

lines intersect on the prescutellar grouve. De-

pression in front of the speculum finely stri-

ate or partly with irregular wrinkles. Meta-

pleurum rugose-punctate. Propodeum scarce-

ly depressed, transversely striate posterior-

ly. Glymma very small or lacking (Both flag-

ellar tips broken off in the only available ?).

Body length 7-8mm. d tentatively placed

here ern rufator Hinz, 1994 ? ?5

Epipleurum of the 3" gastral tergite separat-

ed from the tergite anteriorly, the ventrola-

teral crease marked with black, or with a

black lateral stripe parallel with the crease in

a short distance (Fig. 40)... 115:

Epipleurum of the 3 gastral tergite com-

pletely fused with the tergite (Figs 42-43). ..

Dennerle nen Yansn oubges rn etc kguregherhegsagtt 115,

Frons with a strong median longitudinal keel,

the latter raised medially. Propodeum dis-

tinctly depressed, longitudinal carinae indi-

cated, sulcus in all parts with transverse wrin-

kles. 2" gastral tergite red on the posterior

0.2, posterior edge often brown. Epipleural

index 0.2-0.3, the epipleurum folded in. 58-62

flagellar segments. Body length 14-17 mm...

REN carinifrons (Holmgren, 1860) 25

Frons at most with a fine median longitudi-

nal carina. 2" gastral tergite black dorsally

and posteriorly. Epipleural index 0.1-0.3.....

EEE AN EON EE PUR SACHRRENE 114.

114.

115.

116.

197

118.

64 flagellar segments. Body length 18 mm.

Mesopleurum with disperse punctation on a

smooth background. Depression in front of

the speculum with distinct long striae. Spirac-

ular index 0.3. Epipleural index 0.2, the epi-

pleurum pendent as a small triangular lobe.

d Unknomn. ............. perditator Hinz, 1994 2

47-49 flagellar segments. Body length 12 mm.

Mesopleurum with dense punctation on a

slightly coriaceous and shining background.

Depression in front of the speculum finely

granulate-strigose. Spiracular index 1.0. Epi-

pleural index 0.2-0.3, the epipleurum folded

in or pendent as a narrow rather rectangular

lobez. an 222 perditor (Förster, 1868) 95

Genal carina joining the oral carina at the

base of the mandible, often at a right angle,

the genal carina often bent sidewards and

forwards close to the junction (Figs 10, 15).

BEE LIE EIERN EEE ROHR 116.

Genal carina joining the oral carina some

distance from the base of the mandible, often

at an acute angle, the genal carina often

straight close to the junction (Figs 8-9, 11-14)

(D. chinensis and D. thomsoni are included

here: Genal index 0.1. 2”! gastral tergite red

on the posterior 0.1 or completely black dor-

sally. Posterior gastral tergites red at least

ventrolaterally. Posterior gastral sternites red

ormarkediwithmed)we. ee 126.

55-56 flagellar segments. Body length 11-

12 mm. Hind tibia black. 2" gastral tergite

red at most on the posterior 0.1. ...................

ee ee ee genator Hinz, 1979 23

26-47 flagellar segments. Body length 6-9 mm.

OT 117.

26-32 flagellar segments. Ventral part of the

prepectal carina strongly raised submedial-

ly, deeply emarginate medially. Propodeum

extending to 0.5-0.7 of the hind coxae. 2"“

gastral tergite red on the posterior 0.1-0.2. In

D. juvenilis ovipositor index 1.3. ............. 118.

30-47 flagellar segments. Ventral part of the

prepectal carina less strongly raised subme-

dially and/or propodeum not extending to

the middle of the hind coxae. Ovipositor in-

IEWSENTERE nahe 119.

Pleural part of the prepectal carina complete

or almost complete. 3" gastral tergite red. 4"

tergite often for the greater part red, some-

times black or marked with black. Oviposi-

tor index 0.6. 26-30 flagellar segments. Body

length 7-8 mm. ..... aemula (Förster, 1868) ?3

19:

120.

12T.

122.

Pleural part of the prepectal carina incom-

plete, often partly replaced by wrinkles. 3"

gastral tergite often marked with black. 4"

tergite black. Ovipositor index 1.3. 26-32 flag-

ellar segments. Body length 7-9 mm. ...........

ee juvenilis (Förster, 1868) ?4

2"! gastral tergite completely black or red on

the posterior 0.1. Propodeum distinctly de-

pressed, without longitudinal carinae medi-

ally, distinctly transversely wrinkled in at

least in some parts of the sulcus. ............ 120.

2nd gastral tergite red on the posterior 0.3. ..

Fe EEE DUI EIS EHE NEELLTE NE LESETEEEREN 123.

Glymma lacking. Speculum and depression

in front of the speculum with distinct striae.

Hind tibia black, in some dd tinged with

white subbasally on the dorsal side. 38-42

flagellar segments. Body length 7-9 mm. ....

a 127.

Glymma present, sometimes rather small. 30-

36 flagellar segments. Body length 6-8 mm.

a 122,

Mesopleurum distinctly coriaceous between

the punctures. Pleural carina distinct. Propo-

deum broadly depressed. Mandible for the

greater part or completely black. Mid tibia

yellow. 38-42 flagellar segments. Body length

VS TAN auenezeeseeenenannearae liberator Hinz, 1985 ?5

Mesopleurum almost smooth between the

punctures. Pleural carina partly indistinct.

Propodeum narrowly depressed. Mandible

broadly yellow medially. Mid tibia black. 39

flagellar segments. Body length 9 mm. 2 un-

known. ee RE

Gahedsnfesnge obscuripes Horstmann, spec. nov. d

Hind corner of pronotum not much produced,

distinctly shorter than the tegula. Propodeum

for the greater part irregularly wrinkled,

sometimes with short transverse wrinkles in

the shallow sulcus. 30-36 flagellar segments.

Body length 6-8 mm (See also couplet 109).

an erta carpathica (Szepligeti, 1916) 23

Hind corner of pronotum produced to a

flange, which projects about as far as the

tegula and which is rounded dorsally and

excavated ventrally. Propodeum usually with

long transverse wrinkles in the sulcus at least

posteriorly. 32-36 flagellar segments. Body

length 6-7 mm (See also couplet 160). ..........

Denen minor (Provancher, 1879) ?3

123.

124.

125.

126.

127%

34-37 flagellar segments. Body length 6-7 mm.

Hind tibia yellowish brown medially, nar-

rowly black basally, broadliy black apically.

Glymma very small or lacking. ...................

aiännsedaess nenne nsbe serien scolator Hinz, 1985 ?3

39-47 flagellar segments. Body length 7-10 mm.

ER REEL ade 124.

Hind tibia black in the ?, at most tinged with

yellowish brown subbasally. Hind tibia yel-

lowish medially, narrowly black basally,

broadly black apically in the d, the apical

ring extending over 0.3 of the tibia. 2” gas-

tral tergite red on the posterior 0.3-0.4. 42-47

flagellar segments. Body length 7-9 mm (Va-

riety: 39-41 flagellar segments. Body length

Z mm. ee flagellator (Fabricius, 1793) 23

Hind tibia distinctly yellow or yellowish

brown medially in both sexes, the black api-

eal,ring,shorterindihe go... 0 125,

Hind tibia light yellow medially, narrowly

black basally and apically, the black apical

ring extending over 0.1-0.2 of the tibia in

both sexes. 2" gastral tergite red on the pos-

terior 0.4-0.6 in the ?, on the posterior 0.3-0.4

in the d. 42-46 flagellar segments. Body length

8-10. mm... inermis (Förster, 1868) ?3

Hind tibia yellowish brown medially, nar-

rowly black basally, broadly black apically in

the 9, the black apical ring extending over

0.3 of the tibia. 2" gastral tergite red on the

posterior 0.3-0.4. 41-44 flagellar segments.

Body length 8-9 mm. d unknown. ...............-

her petitor Hinz & Horstmann, spec. nov. 2

Genal carina raised close to the junction with

the also raised oral carina, forming a lopsid-

ed triangular pyramid, the tip of the latter

shifted ventrally and situated more or less

behind the base of the mandible (Figs 11-12).

REDE ER crerpscnecnnooe 12%

Genal carina not distinctly raised, its junction

with the oral carina situated above the base

of the mandible (Figs 8-9, 13-14). ........... 132.

Mesopleurum slightly coriaceous between the

punctures. Hind tibia yellow or yellowish

brown medially, narrowly black basally,

bzoadly black apieallyor nee 128.

Mesopleurum smooth between the punc-

tures. Hind tibia completely yellowish or only

narrowly darkened basally and apically .....

RE a seh ende 129.

128. Antennal carina low and narrow (Fig. 3). Pre-

129.

150;

131.

pectal carina and transverse carina low. Pro-

podeum broadly and shallowly depressed,

the sulcus for the greater part irregularly

wrinkled. Gaster from the 2" tergite onwards

narrowly black dorsally and broadly red lat-

erally. 39-41 flagellar segments. Body length

SL RN SURER AIR HRNUREEEEENENER

hbaglin rufiventrator Horstmann, spec. nov. 23

Antennal carina somewhat raised (Fig. 5).

Ventral part ofthe prepectal carina and trans-

verse carina somewhat raised. Pleural part of

the prepectal carina present. Propodeum

broadiy and deeply depressed, distinctly

transversely wrinkled in almost all parts of

the sulcus. Gaster red medially, black anteri-

orly and posteriorly. 45-50 flagellar segments.

Body length 8-10 mm (See also couplet 82).

a ulnunlescetlradanens sobolicida (Förster, 1868) ?3

Genal carina and oral carina very strongly

raised, the gena deeply depressed close to

the junction of both carinae, the depression

about as deep as the width of the 2": flagellar

segment. Sulcus of the propodeum irregular-

ly wrinkled anteriorly, transversely wrinkled

posteriorly. 2” gastral tergite completely

black dorsally. 49-51 flagellar segments. Body

lensihiehl 12 m. een. seen

RER rl lividariae Hinz, 1963 23

Genal carina and oral carina not very strong-

ly raised, the depression of the gena 0.5-0.8x

as deep as the width of the 2"‘ flagellar seg-

ment (Figs 11-12). 2"X gastral tergit dorsally

yellow or red on the posterior 0.1-0.3. .........

nn. ae aen den hauen änsparirerknenternnnagpinehengpenen 130.

Flagellum light brown on the apical 0.3. Pleu-

ral part of the prepectal carina almost com-

plete. Propodeum broadly but not deeply

depressed, with transverse wrinkles over al-

most the total breadth of the sulcus. 47-49

flagellar segments. Body length 9-11 mm (See

alSERCONBIEE AOL) REEL FE 2

Be dl ner), splenditor Hinz, 1985 ?3

Flagellum black apically........................ 131.

Pleural part of the prepectal carina more or

less complete ventrally and medially, the dor-

sal 0.2 replaced by several wrinkles. Propo-

deum distinctly depressed, with transverse

wrinkles over almost the total breadth of the

sulcus. 3° gastral tergite without black later-

al stripes (Fig. 43). 53-55 flagellar segments.

Bodyalenerh Tamm rennen

EIERN EEE mercator (Fabricius, 1793) 24

152.

133.

134.

193:

136.

Pleural part of the prepectal carina indicated

by an indistinct short wrinkle only. Propo-

deum distinctly depressed, with short trans-

verse wrinkles in the midline of the sulcus, in

other parts irregularly wrinkled. 3‘ gastral

tergite with distinct black lateral stripes (Fig.

42).47 flagellar segments. Body length 10 mm.

NUnkDOWDr ee nn

A tibiatoria Horstmann, spec. nov. d

Gaster from the 3" tergite onwards almost

completely red. Genal index 0.1. 50 flagellar

segments. Body length 9mm. d unknown

(Seeralso eouplebzyı er nenn

N chinensis Horstmann, spec. nov. 9

Posterior gastral tergites distinctly marked

with black or completely black. .............. 133.

Gaster black dorsally, red laterally, rarely the

3 tergite completely red, but in that case the

2"d tergite broadly red laterally. ............. 134.

Gaster red (or rarely yellow) medially, black

anteriorly and posteriorly, usually the 3'°

tergite completely red, often also the 4" tergite

for the greater part or completely red, the 2‘

tergite often black anteriorly, red posteriorly

(If the 3° tergite is black dorsally and red

laterally, then the 2”! and 4" tergites are al-

most completely or completely black). ........

EEE EN E 141.

58 flagellar segments. Body length about

15 mm. Hind tibia and hind tarsus complete-

ly light yellow. 2" to 7% gasteral tergites

narrowly black dorsally, broadly red lateral-

ya, 2-UNKnOWMS - nase zenesstenenansnresennstehennche

ER scalprata Horstmann, spec. nov. d

Number of flagellar segments and body

length often smaller. Hind tibia distinctly dark

brown or black at least basally................ 1.39.

Hind tibia dark brown to black. Ventral part

of the prepectal carina and transverse carina

not raised, pleural part indicated by wrinkles

or obliterated. 48-50 flagellar segments. Body

fenstheit 12 mama ee een er her

RER spinipes (Thomson, 1887) 25

Hind tibia broadly yellow to brown medial-

Tyan sun et ana hear uch anash nn neh rennen 136.

Propodeum irregularly wrinkled at least pos-

teriorly, sometimes with some longitudinal

wrinkles.Eostulae fine. ..-.............:..:::-- 137.

Propodeum distinctly transversely wrinkled

laterally and posteriorly. Costulae usually

distinct, almost reaching the propodeal spir-

BCl. <utssnsagensnaasnensnunaknsannenauedingenserandaraneneange 138.

137. Genal index 0.5. Mandible broadly yellow

138.

139:

140.

medially. Tegula yellow to brown. Hind tib-

ia yellowish brown medially, narrowly black

basally, broadly dark brown to black apical-

ly. Propodeum scarcely depressed, for the

greater part irregularly wrinkled, often with

some longitudinal wrinkles posteriorly. 39-46

flagellar segments. Body length 7-9 mm (See

alsoreouplets22land 23a)... een

en neegrensnesuncase subimpressa (Förster, 1868) 95

Genal index 0.25. Mandible and tegula black.

Hind tibia black basally and on the ventral

side, dark brown apically and on the dorsal

side. Propodeum slightly depressed, trans-

versely wrinkled anteriorly and medially, ir-

regularly wrinkled posteriorly. 41 flagellar

segments. Body length 9 mm. d unknown.

gs: tritor Hinz & Horstmann, spec. nov. ?

39-42 flagellar segments. Body length 7-8 mm.

Genal index 0.1. Hind tibia yellowish medi-

ally, narrowly black basally, broadly black

apically, the apical ring extending over 0.3 of

the tibia. 2" gastral tergite almost complete-

black nr.2..... thomsoni Hinz, 1963 23

44-58 flagellar segments. Body length 8-14 mm.

Genal index 0.4-0.5. Hind tibia yellowish

brown medially, narrowly brown or black

basally, broadly brown or black apically, the

apical dark mark sometimes lacking. ...........

Be ee ee euch erde 139.

Pleural part of the prepectal carina obliterat-

ed (Fig. 21). Nervellus distinctly reclivous.

Flagellum brown apically. Posterior gastral

sternites black. 45-47 flagellar segments. Body

lensthes me. meer ee

PSERÄTE RER BE constantineanui Hinz, 1977 25

Pleural part of the prepectal carina present at

least medially (Figs 19-20). Nervellus verti-

cal. Flagellum black. Posterior gastral ster-

nites red or reddish brown...................... 140.

44-48 flagellar segments. Body length 9-11 mm.

Speculum often with fine striae or granulate-

strigose also ventrally. Metapleurum rugose,

punctures usually not discernible..................

NE circumcinctus (Förster, 1868) 23

49-58 flagellar segments. Body length 10-

14 mm. Speculum shining, often smooth at

least ventrally. Metapleurum often with dis-

tinct punctures (punctate or rugose-punctate).

ER SE NERRE signator (Brauns, 1895) 25

141.

142.

143.

144.

145.

Nervellus reclivous, nervellar index 0.4-0.6

(Fig. 29). Hind tibia whitish yellow or yel-

lowish brown medially, narrowly black ba-

sally, broadly black apically (Pleural part of

the prepectal carina obliterated; flagellum

brown apically: see D. constantineanui, cou-

plee 19). rar 180.

Nervellus inclivous, vertical or reclivous, in-

tercepted more or less far below the middle,

or not intercepted at all (Figs 26-28). ..... 142.

Genal carina bent sidewards and forwards

and more or less obliterated close to its junc-

tion with the oral carina, the gena somewhat

depressed, the oral carina raised (Figs 9, 14).

Ventral part ofthe prepectal carina and trans-

verse carina strongly raised (Fig. 19). Hind

tibia black in the ?, whitish yellow medially,

narrowly black basally and broadly black

apically in the d. 2"! gastral tergite red on the

posterior 0.3. 3"! and 4" tergites red, the 4"

tergite sometimes marked with black poste-

riorly. 44-48 flagellar segments. Body length

Simmern tenuis (Förster, 1868) 243

Genal carina sometimes bent sidewards close

to its junction with the oral carina, but not

obliterated. Prepectal carina often not much

raised. Gaster often less extensively marked

with. red... LEER 143.

65-67 flagellar segments. Body length 18-

20 mm. Hind tibia completely yellow. Gaster

yellow medially. 3" tergite without black lat-

eral’stripes (See also eouplet 195),

RE falcator (Fabricius, 1775) 23

At most 56 flagellar segments. Body length at

MOST 13 mImn euren nennen ven segan anne 144.

Mesopleurum distinctly coriaceous and dull

between the punctures. 2" gastral tergite red

on the posterior 0.2-0.4 (exception: D. amura-

oe 145.

Mesopleurum shining and slightly or not at

all coriaceous between the punctures. .. 148.

50-51 flagellar segments. Body length 9-10 mm.

Genal index 0.6. Hind tibia black, marked

with brown ventrally. 2"’ gastral tergite red

on the posterior 0.2-0.4. 3" tergite red. 4"

tergite red, marked with black posteriorly.

2 unknown. „... en... Sagen

sogen varıpes Horstmann, spec. nov. d

36-47 flagellar segments. Body length 6-

10 mm. Genal index 0.1-0.4. Hind tibia often

yellowish brown to brown medially. ...........

Eelldun nee Kasbtann£yenth ana swienehtggeehaensesnerb rer ee 146.

146.

147.

150.

Genal index 0.3-0.4. Mesopleurum with dense

punctation. 2” gastral tergite red on the pos-

terior 0.1-0.2. 3° tergite red, more or less

marked with black, sometimes almost com-

pletely black. 4" tergite black, at most nar-

rowly marked with red anteriorly. 36-41 flag-

ellar segments. Body length 6-10 mm (See

alsescouplets227 and 235). .%.u namen

TER amuratör Hinz, 1985 23

Genal index 0.1-0.3. 2" gastral tergite red on

the posterior 0.2-0.5. 3° and 4" tergites red,

sometimes marked with black. 42-47 flagel-

[ar Sepments. eretranehatraraenneneeaensenennaennganenen 147.

Genal index 0.1-0.2. Temple with very fine

and disperse punctation medially. Mesopleu-

rum with disperse or rather dense puncta-

tion. Petiole with narrow lateral areas. 2‘

gastral tergite red on the posterior 0.2-0.3.

42-47 flagellar segments. Body length 8-9 mm.

Seegalsoreouplet 210), 2...

BE N ER annexa (Förster, 1868) ?d

Genal index 0.2-0.3. Temple with fine and

rather dense punctation medially. Mesopleu-

rum with dense punctation. Petiole without

glymmae or narrow lateral areas. 2" gastral

tergite red on the posterior 0.3-0.5. 43-46 flag-

ellar segment. Body length 8-9 mm (See also

couplers 185, 206 and 215)... nenn

BE lade meridionator Aubert, 1960 25

. 29-37 flagellar segments. Body length 6-9 mm.

ELSE Ran este sosesatnsusrn er plnhtnnatir lets henfenesedekgesensen 149.

RE ERNST EEE 150.

. Hind tibia black, at most tinged with dark

brown medially. Propodeum slightly or not

at all depressed, at most with short trans-

verse wrinkles. 2” and 4" gastral tergites

black. 35-37 flagellar segments. Body length

SINN OAUNKNOWEN een kekeehteemnaneene

EINE An affinitor Hinz, 1979 2

Hind tibia yellowish brown, narrowly brown

basally, not darkend apically. Propodeum

somewhat depressed, strongly transversely

wrinkled in the shallow sulcus, in other parts

strongly irregularly wrinkled. 2" gastral

tergite red on the posterior 0.4. 3’ and 4"

tergites red. 29-34 flagellar segments. Body

length 6-7 mm (See also couplet 101). ..........

spiracularis Hinz & Horstmann, spec. nov. 23

Mesopleurum slightly coriaceous between the

punctures. Mid tibia light yellow dorsally.

Hind tibia blackish, dorsally marked with

yellowish brown at base. 44 flagellar seg-

151!

192.

153.

154.

ments, the 2"! segment 2.1x as long as wide.

Body length 8 mm. d unknown. .............

. belokobyli Hinz & Horstmann, spec. nov. 2

Mesopleurum at least partly smooth between

the punctures (exception: some specimens of

D. petiolator). Mid tibia yellowish brown to

blackish brown dorsally. Hind tibia black ba-

sally. Base of flagellum usually stouter. ......

ann sasdn feunerun came sunwansennnennssßcncntlensvsskersnsntsgtrtessee 151.

Pleural part of the prepectal carina lacking

(Fig. 21). Pleural part of the prepectus with

well’ separated punetures.........n.... 152%

Pleural part of the prepectal carina partly

present (Figs 19-20) and/or pleural part of

the prepectus distinctly rugose-punctate and

witkne"wrinklester Amen en! 153

Temple as long as the width of an eye. Malar

space index 0.7. Propodeum not depressed,

irregularly wrinkled. 47-48 flagellar segments.

Body length 13 mm. $ unknown. ...........

ENRERR TEN longigena Horstmann, spec. nov. d

Temple 0.7x as long as the width of an eye.

Malar space index 0.4. Propodeum distinctly

depressed, for the greater part transversely

wrinkled. 47-51 flagellar segments. Body

lemetke 9-1 0"nm te.

temnator Hinz & Horstmann, spec. nov. ?4

Temples 0.9-1.0x as long as the width of an

eye, not much constricted behind the eyes,

tangential lines on eyes and temples inter-

sect on the propodeum (Fig. 2). Pleural part

of the prepectal carina at least for the greater

part present. 40-45 flagellar segments. Body

length 9-11 mm (See also couplet 188). ........

a BE bucculenta (Holmgren, 1860) ?3

Temples shorter and more constricted be-

hind the eyes and/or pleural part of the pre-

pectal carina obliterated and replaced by

dense punctation or by short wrinkles. .......

nee rer: 154.

Pleural part of the prepectal carina obliterat-

ed and replaced by dense punctation or by

short wrinkles (Fig. 21). Black lateral stripes

of the gastral 3" tergite distinct. Ovipositor

index 0.5. 44-52 flagellar segments. Body

lenSthia1 O1 ne an ee dene Aachen ken

ERROR japonica (Cameron, 1906) ?3

Pleural part of the prepectal carina usually

present, sometimes shortened (Figs 19-20)

(in the d of D. petiolator rarely replaced by

irregular wrinkles). Black lateral stripes of

the 3% gastral tergite sometimes reduced...

IR REIERCHEN L O SERIES RRRERR TE TER BEER DE 155.

27,

155. 42-45 flagellar segments. Body length 8-10 mm.

156.

159.

3" gastral tergite without black lateral stripes

or these stripes indistinct. Ovipositor index

EAN een petiolator (Fabricius, 1804) 23

53-56 flagellar segments. Body length 10-

11 mm. 3 gastral tergite with distinct black

lateral stripes. Ovipositor index 0.5. .............

petiolatoides Hinz & Horstmann, spec. nov. ?d

Genal carina joining the oral carina at the

base of the mandible (Figs 10, 15). ......... 157.

Genal carina joining the oral carina some

distance from the base of the mandible (Figs

SEITE AI 161.

. 2"! gastral tergite red on the posterior 0.2-0.4.

ORSaREnwapeHranEE ra nse Lungsadeten Dh dngnnesuneissnkgarnenseheee ter 158.

2"d gastral tergite red at most on the posterior

N ee ee ee Mr 159:

. Temples scarcely constricted behind the eyes,

tangential lines on eyes and temples inter-

sect on the apex of the gaster or not at all.

Temple with distinet and dense punctation

medially. Metapleurum coriaceous and with

dense punctation (sculpture almost as in the

adjoining parts of the mesopleurum). Hind

femur marked with red apically. 43-44 flagel-

lar segments. Body length 8 mm. .................-

BEN temporalis Horstmann, spec. nov. 23

Temples more strongly constricted behind

the eyes. Temple with fine and disperse punc-

tation medially. Metapleurum rugose-punc-

tate or rugose (sculpture as in the adjoining

parts of the propodeum). Hind femur black.

RN re Bern A a RER EERCHESRSRERRCRRES 123.

Genal carina joining the oral carina short dis-

tance from the base of the mandible, genal

index 0.1 or less (sometimes hardly visible).

Propodeum slightly depressed, in most parts

finely and irregularly wrinkled, with short

transverse wrinkles in the narrow longitudi-

nal furrow, without costulae. Gliymma lack-

ing. 36-37 flagellar segments. Body length

7 mm. d unknown (See also couplet 209). ...

.... radiator Hinz & Horstmann, spec. nov. 9

Genal carina joining the oral carina distinctly

at the base of the mandible. Propodeum rath-

er shallowly and broadly depressed, usually

with long transverse wrinkles posteriorly,

with complete costulae. Glymma small, some-

times developed as a shallow furrow. .. 160.

160.

161.

162.

163.

164.

Hind corner of pronotum not much produced,

distinctly shorter than the tegula. 33-38 flag-

ellar segments. Body length 7-9 mm. ...........

angustifrons (Förster, 1868) 23

Hind corner of pronotum produced to a

flange, which projects about as far as the

tegula and which is rounded dorsally and

excavated ventrally. 32-36 flagellar segments.

Body length 6-7 mm (See also couplet 122).

EA ES. minor (Provancher, 1879) 23

Genal carina raised close to the junction with

the also raised oral carina, forming a lopsid-

ed triangular pyramid, the tip of the latter

shifted ventrally and situated more or less

behind the base of the mandible (Figs 11-12).

2"! gastral tergite red laterally on the posteri-

CO) 1 Io BR Eee 162.

Genal carina and oral carina not or not much

taised(FIgS'8, 19). we 164.

Flagellum light brown on the apical 0.3. Meso-

pleurum scarcely coriaceous or smooth be-

tween the punctures. Hind tibia light yellow,

very narrowly darkened basally and apically

or not at all darkened. 47-49 flagellar seg-

ments. Body length 9-11 mm (See also cou-

plet 130). 22 splenditor Hinz, 1985 23

Flagellum dark brown to black to its apex.

Mesopleurum distinctly coriaceous between

the punctures. Hind tibia whitish yellow me-

dially, narrowly black basally, broadly black

apically. 37-39 flagellar segments. Body length

Sn ee... 163.

Genal carina and oral carina not very strong-

ly raised, the depression of the gena about

half as deep as the width of the 2" flagellar

segment (Fig. 11). Propodeum somewhat de-

pressed, in most parts irregularly wrinkled.

3"! gastral tergite without black lateral stripes.

37-39 flagellar segments. Body length 8 mm.

Sunknownzerr...ne dictor Hinz, 1979 2

Genal carina and oral carina very strongly

raised, the gena deeply depressed close to

the junction of both carinae, the depression

about as deep as the width of the 2" flagellar

segment. 3"4 gastral tergite with black lateral

stripes. 38 flagellar segments. Body length

8 MMS. unknown... ee

ER N Le. lamellifer Hinz, 1994 ?

Epipleural index 0.1-0.4. Epipleurum of the

3“! gastral tergite either folded in or folded

back secondarily and pendent, in the latter

case the former crease present as a bulge and

163

166.

167.

168.

the ventrolateral edge of the epipleurum run-

ning parallel with this bulge in a short dis-

tance. Black lateral stripe, if present, on the

bulge or close to the latter (Fig. 40). ...... 165.

Epipleurum of the 3° gastral tergite fused

with the tergite, rarely discernible as a small

triangular lobe anteriorly. Black lateral stripe,

if present, distinctly distant from the ventro-

lateral edge and divergent from the latter

Ersileie) ei A ee 179.

Petiole on either side with a long row of

distinct transverse wrinkles in a long longi-

tudinal furrow in front of the glymma (as in

Fig. 35). 46-48 flagellar segments. Body length

IILmmM. nr insignita (Förster, 1868) ?J

Petiole on either side smooth or coriaceous in

front ofthe glymma, at most with some trans-

Merse wrinklesianferiorly: -.:..2.........2.0; 166.

2d gastral tergite dorsally black posteriorly,

often laterally red in front of the posterior

edge, rarely also dorsally tinged with red in

front of the posterior edge....................... 167.

2"d gastral tergite dorsally red at least on the

posterior 0.2, rarely the posterior edge brown-

ish, but not black, often the red marks more

extended laterally. Usually the 3" and 4*

tergites completely or almost completely red

(EXTEPFION: DTECIOTAES)).er.2022020202242er een 1722

Hind tibia black, rarely somewhat lightened

medially (dark reddish brown). Propodeum

slightly depressed, distinctly and rather ir-

regularly wrinkled, the longitudinal carinae

sometimes indicated. 3“ and 4" tergites red,

rarely marked with black dorsally. 42-49 flag-

ellar segments. Body length 8-12 mm .........

norikurae Hinz & Horstmann, spec. nov. ?d

Hind tibia yellowish brown to light brown

medially, dark brown to black basally and

apically, and/or 4" gastral tergite almost com-

pleiely blacksu nr elle sehn, 168.

38-43 flagellar segments. Body length 7-9 mm.

Propodeum somewhat depressed, distinctly

irregularly wrinkled, with short and strong

longitudinal wrinkles posteriorly. Area su-

peromedia sometimes delimited laterally, for

the greater part coriaceous and not wrinkled.

Gaster usually for the greater part or com-

pletely black dorsally, 2” and 3" tergites to a

variable extent marked with red laterally,

rarely the posterior tergites narrowly red ven-

trally (See also couplets 4 and 16). ................

NEN unse: melanator Hinz, 1985 23

169.

170.

TA:

44-55 flagellar segments. Body length 9-14 mm.

3“ gastral tergite usually for the greater part

zedhdarsalyna ine 169.

Temples not much constricted behind the eyes,

tangential lines on eyes and temples inter-

sect on the propodeum (similar to Fig. 2).

Propodeum slightly depressed, irregularly

wrinkled, dull, somewhat transversely wrin-

kled posteriorly. 2” to 7" gastral tergites

broadly marked with red laterally and to a

variable extent darkened dorsally. 53-55 flag-

ellar segments. Body length 14mm. d un-

known ns een

Rn peptor Hinz & Horstmann, spec. nov. ?

Temples more strongly constricted behind

the eyes. At least the 6" and 7" tergites black

dorsally and laterally. Body often shorter. ..

ER RE ARE 170.

Metapleurum coriaceous and with dense

and fine wrinkles, punctation hardly visible

(sculpture as in the adjoining parts of the

propodeum). Speculum finely coriaceous, dis-

tinctly shining. Depression in front of the

speculum almost completely striate. Propo-

deum somewhat depressed, with distinct lon-

gitudinal carinae, distinctly and for the great-

er part irregularly wrinkled, dull, distinctly

transversely striate posteriorly. 3" to 5"" gas-

tral tergites red, 5" tergite black dorsally.

50 flagellar segments. Body length 12 mm.

&unknowna.n. nubilator Hinz, 1994 2

Metapleurum with fine punctation on a co-

riaceous background, scarcely rugose (sculp-

ture as in the adjoining parts of the meso-

pleurum). Speculum more distinctly coria-

ceous. Depression in front of the speculum

for the greater part granulate-strigose, with

distinct striae or wrinkles only dorsally (this

character being variabile) (Depression in front

of the speculum with distinct striae ventral-

ly; propodeum without transverse wrinkles:

see D. yamanakai, couplet 32)................... 1WAlQ

Depression in front of the speculum granu-

late-strigose, with some fine striae dorsally.

Propodeum scarcely depressed in the area

superomedia, distinctly and narrowly de-

pressed in the area petiolaris, with short trans-

verse wrinkles in the longitudinal furrow, in

other parts irregularly wrinkled. 2" gastral

tergite black. 4" tergite for the greater part

black, marked with red only laterally. 44 flag-

ellar segments. Body length 9 mm. 4 of this

variety unknown (See also couplet 178). .....

ER EN graptor Hinz, 1985 var. ?

11722,

173:

174.

Depression in front of the speculum with

distinct striae or wrinkles dorsally. Propo-

deum rather broadly depressed, for the great-

er part with fine irregular wrinkles, often

with strong transverse wrinkles posteriorly.

2"! gastral tergite often laterally and rarely

also dorsally marked with red in front of the

dark posterior edge. 4" tergite often com-

pletely or almost completely red. 48-55 flag-

ellar segments (Variety: 45-47 segments).

Body length 9-12 mm (See also couplet 35).

BE vigilator (Förster, 1868) 23

Metapleurum distinctly and densely rugose-

punctate, not discernibly coriaceous. Hind

tibia yellow medially, narrowly black basal-

ly, broadly black apically. Propodeum scarce-

ly depressed, coarsely and irregularly wrin-

kled, almost completely areolated, but the

carinae scarcely discernible among the wrin-

kles. 3“ gastral tergite without black lateral

stripes. 43 flagellar segments. Body length

10 mm. $ unknown (See also couplet 233)..

ren inconspicua Horstmann, spec. nov. d

Metapleurum usually distinctly coriaceous

or finely wrinkled between the punctures.

Propodeum not coarsely wrinkled all over.

3"! gastral tergite with black lateral stripes.

EEE EN NE A 173.

Speculum and depression in front of the

speculum densely and finely striate, the stri-

ae merging with dense punctation anterior-

ly. Metapleurum densely rugose-punctate,

not coriaceous at least dorsally and centrally.

Hind tibia black. Propodeum scarcely de-

pressed, irregularly wrinkled in the shallow

depression, somewhat shining. 48-49 flagel-

lar segments. Body length 10 mm. d un-

KINO Wan a ee ER,

a venitor Hinz & Horstmann, spec. nov. ?

Speculum coriaceous. Depression in front of

the speculum finely granulate-strigose, the

striae merging with a coriaceous area with-

out punctation anteriorly (exception: D. nigri-

dens). Metapleurum coriaceous between the

punctures. Hind tibia usually yellowish me-

dially (exception: D. rectoides). ................ 174.

Propodeum somewhat depressed, with com-

plete costulae and longitudinal carinae, shin-

ing in the depression. Area superomedia co-

riaceous. Area petiolaris with fine irregular

or transverse wrinkles. Hind tibia whitish

yellow medially, narrowly black basally, rath-

er broadly black apically. 50 flagellar seg-

173.

176.

177.

178.

ments. Body length 10 mm. ? unknown (See

alsolieouplEEIO EEE ERRER

ee rufovariata Horstmann, spec. nov. d

Propodeum more strongly wrinkled and/or

not distinctly shining in the depression (ex-

ception: D. graptor), carinae more reduced..

ER ER EEE 175:

Propodeum distinctly depressed, distinctly

transversely wrinkled over the total breadth

of the sulcus at least posteriorly. ............. 176.

Propodeum more or less distinctly depressed,

for the greater part irregularly wrinkled, at

most with some short transverse wrinkles

not extending over the total breadth of the

SUlCUS nee ER 177.

Hind tibia whitish yellow medially, narrow-

Iy black basally, broadly black apically. Pro-

podeum with a distinct sulcus, with short or

indistinct transverse wrinkles anteriorly,

more transversely wrinkled medially, strong-

ly transversely wrinkled over the total breadth

of the sulcus posteriorly. 42-45 flagellar seg-

ments. Body length 8-10 mm (See also cou-

pletgay er recta (Thomson, 1887) 23

Hind tibia yellowish brown medially, nar-

rowly black basally, broadly black apically.

Propodeum finely transversely striate over

the total breadth anteriorly and posteriorly,

more irregularly wrinkled medially (Both

flagellar tips broken off in the only available

specimen). Body length 11 mm. d unknown.

.. rectator Hinz & Horstmann, spec. nov.. ?

37-41 flagellar segments. Body length 6-9 mm.

Metapleurum with very fine punctation on a

distinctly coriaceous background. Hind tibia

blackish in the $, yellowish brown to brown

medially, black basally and apicaliy in the 4.

Propodeum slightly depressed, with partly

developed longitudinal carinae, for the great-

er part irregularly wrinkled, with some lon-

gitudinal wrinkles posteriorly. ...............

rectoides Hinz & Horstmann, spec. nov. ?d

44-52 flagellar segments. Body length 8-12 mm.

Metapleurum with distinct punctation on a

coriaceous background. Hind tibia lighter

medially. Longitudinal carinae of propodeum

Scareely, developed... ...... ee 178.

44-50 flagellar segments. Body length 8-10 mm.

Tegula yellow to brown. Depression in front

of the speculum granulate-strigose, with a

narrow coriaceous area without punctation

in front of the depression. Hind tibia medial-

129:

180.

181.

ly yellowish brown in the $, yellow in the g,

narrowly black basally, broadly black apical-

ly. Propodeum not depressed in the area su-

peromedia, the latter coriaceous, distinctly

depressed in the area petiolaris, with short

and fine transverse wrinkles in the rather

narrow longitudinal furrow, coriaceous and

with fine irregular wrinkles in areas beside

the furrow (See also couplet 171). .................

lu graptor Hinz, 1985 23

52 flagellar segments. Body length 12 mm.

Tegula black. Depression in front of the specu-

lum with fine striae, which merge with the

punctation anteriorly. Hind tibia yellowish

brown medially, narrowly brown basally and

apically. Propodeum broadly and shallowly

depressed, for the greater part irregularly

wrinkled, with some longitudinal wrinkles

Posteniorly. 2 unknown. -:-...s.2.2..-0240.

a nigridens Horstmann, spec. nov. d

Nervellus reclivous, nervellar index 0.4-0.6

(Fig. 29). Flagellum and hind femur black. .

ee re ER Rs Sa 180.

Nervellus inclivous, vertical or reclivous, in-

tercepted more or less far below the middle,

or not intercepted at all (Figs 26-28) (D. con-

tumator and D. rubator with a nervellar index

of 0.6 are included here. D. contumator: Flag-

ellum brown apically. D. rubator: Hind fe-

BRRETENN Eu ne lee eh ieh 181.

Propodeum not depressed, for the greater

part with very fine irregular wrinkles, with

some short transverse wrinkles posteriorly.

2"d gastral tergite red on the posterior 0.2.

3" tergite red. 4" tergite broadly red anteri-

orly. 52 flagellar segments. Body length

12 men Pu KMOWEL. en eieeseredenhnssunnene

DEN TREN nervellator Horstmann, spec. nov. d

Propodeum deeply depressed, with distinct

transverse wrinkles in the sulcus medially

and posteriorly. 2” gastral tergite red on the

posterior 0.1-0.4. 3”! and 4" tergites red. 45-53

flagellar segments. Body length 8-12 mm (See

alseotpler20 N Bern ine

ef xenocampta (Förster, 1868) 93

Temples at least 0.9 (rarely 0.8) x as long as

the width of an eye, not much constricted

behind the eyes, tangential lines on eyes and

temples intersect on the posterior end of the

propodeum or on the gaster or not at all

Le a NE 182.

182.

183.

184.

185.

Temples shorter and/or more strongly con-

stricted behind the eyes (D. fundator and

D. pulmentariae, which are intermediate in this

character, are included here. D. fundator: Hind

tibia black; propodeum irregularly wrinkled.

D. pulmentariae: Hind tibia light yellow ba-

sally and medially, distinctly black apically).

ee 189.

Hind tibia yellowish brown to brown, often

narrowly darkened basally and/or apically.

2rd gastral tergite red on the posterior 0.7,

rarely black dorsally and marked with red

only laterally. 3°“ and 4" tergites red. .... 183.

Hind tibia blackish, in some species with a

brownish mark medially on the dorsal side

(with a whitish mark in D. meridionator).......

EEE RRTSO TH EHrereDR Er EEReE 184.

Hind femur red. Mesopleurum distinctly co-

riaceous between the punctures. Propodeum

scarcely depressed, irregularly wrinkled. 46

flagellar segments. Body length 10 mm (See

AlSOreoupletHl U) ER ER N.

alpinator Hinz & Horstmann, spec. nov. 24

Hind femur black. Mesopleurum smooth be-

tween the punctures. Propodeum distinctly

depressed, with short and strong transverse

wrinkles in the longitudinal furrow, with very

dense punctation in the areas beside the fur-

row. 40-45 flagellar segments. Body length

9:10: mm. ER capitator Hinz, 1985 25

Mesopleurum distinctly coriaceous between

tbezpuneluresn. 0 ee 185.

Mesopleurum smooth between the punc-

tures. Hind tibia completely black or tinged

with brown medially on the dorsal side.......

aka s nem asna has uannd tee rensneun des DIsgslkesttelänheegtine 186.

Hind tibia white medially on the dorsal side.

Propodeum deeply depressed, distinctly

transversely wrinkled in the narrow longi-

tudinal furrow. Pleural carina and spiracular

carina distinct. Spiracular index 0.3-0.5. 47

flagellar segments. Body length 8-9 mm. 2 of

this variety unknown (See also couplets 147,

206. and AS le TR ter

BAM IR IMEN meridionator Aubert, 1960 var. &

Hind tibia completely black or tinged with

brown medially. Propodeum shallowly and

broadly depressed, with long transverse wrin-

kles posteriorly. Pleural carina often partly

indistinct. Spiracular carina obliterated. Spirac-

ular index 0.9-1.1. 44-48 flagellar segments.

Body length 10-11 mm (See also couplet 216).

BEER RER stygia (Förster, 1868) 95

186. 33-35 flagellar segments. Body length 8 mm.

187.

188.

Propodeum scarcely depressed and partly

with fine transverse wrinkles in the 9, not

depressed and irregularly wrinkled in the d.

2nd gastral tergite red on the posterior 0.6-0.8

in the 9, on the posterior 0.2 in the 6. Usually

the 3"! and 4" tergites red. Glymma and black

lateral stripes of the 3" gastral tergite lack-

ine-Ovipesitorinde 132. nassen

I setator Hinz, 1985 23

40-46 flagellar segments. Body length 10-

12mm. Oyipositor index 0.6. ....-...2.. 187.

Pronotum with punctures ventrolaterally.

Propodeum scarcely depressed, irregularly

wrinkled. Glymma distinct. 2" gastral tergite

red on the posterior 0.3. 3" tergite red. 4"

tergite red laterally. 43 flagellar segments.

Body length 11 mm. d unknown. ...........

er pseudobucculenta Hinz & Horstmann,

spec. nov. ?

Pronotum distinctly wrinkled ventrolateral-

ly. Propodeum slightly depressed, with long

and fine transverse wrinkles almost over the

total breadth of the sulcus at least posterior-

ly. Petiole without glymmae, at most with

very small lateral furrows. 2” gastral tergite

red on'the posterior 0.1. ..*.................. 188.

Flagellum black. 4" gastral tergite red. Ven-

tral part of the prepectal carina often dis-

tinctly raised, usually twice as high as the

width of the apical segment of a maxillary

palpus. 40-45 flagellar segments. Body length

IA mm" (Seealsoreouplet 153)... ne...

EN bucculenta (Holmgren, 1860) ?5

Flagellum light brown apically. 4" gastral

tergite black. Ventral part of the prepectal

carina not distinctly raised. 43-46 flagellar

segments. Body length 10-12 mm. ................

Ehen bucculentoides Hinz & Horstmann,

spec. nov. $d

. Allfemora and tıbiae red... 190.

Hind femur black and/or hind tibia distinct-

lyamarked withrblack. er... 191.

. Flagellum, tegula, trochanters, trochantelli,

postpetiole and anterior 0.3-0.4 of the 2" gas-

tral tergite black. Remaining part of 2! and

3“ and 4" tergites red. Mesopleurum coria-

ceous and dull between the punctures. Pro-

podeum broadly but not deeply depressed,

partly transversely wrinkled, partly irregu-

larly wrinkled. 44-46 flagellar segments. Body

length 9-10 mm (See also couplet 183). ........

alpinator Hinz & Horstmann, spec. nov. ?d

191:

192:

193.

194.

195.

Flagellum marked with brown apically. Tegu-

la yellow. Trochanters, trochantelli, postpet-

iole and 2”! to 4" gastral tergites light red.

Mesopleurum smooth between the punc-

tures. Propodeum distinctly depressed, trans-

versely striate in the longitudinal furrow,

irregularly wrinkled in areas beside the fur-

row. 39-40 flagellar segments. Body length

8 mm. d unknown........ rubator Hinz, 1985 2

Front and mid femora and all tibiae and tarsi

yellow, the tarsi marked with brown apical-

ly. 2" to 4" gastralttergites light red, 2" tergite

black on the anterior 0.2. Mesopleurum, meta-

pleurum and propodeum for the greater parts

distinctly and densely rugose-punctate. Pro-

podeum slightly depressed. 40-42 flagellar

segments. Body length 9-11 mm. .................

confusator Hinz & Horstmann, spec. nov. 25

Legs not as extensively marked with yellow.

2" gastral tergite not as extensively marked

withired. Wr RER 192.

Hind tibia yellow basally and medially, black

on the apical 0.3. Flagellum light brown api-

cally. 36-40 flagellar segments. Body length

6-Imm pulmentariae Hinz, 1963 23

Hind tibia with a different colour pattern. ..

ee ER 193.

Pleural part of the prepectal carina distinctly

joining the anterior edge of the mesopleu-

rum (on either side) (Fig. 16). ................. 194.

Pleural part of the prepectal carina not join-

ing the anterior edge of the mesopleurum,

either obliterated dorsally or replaced by sev-

eral wrinkles (Figs 17-18) (This character has

strong infraspecific variability). .............. 203.

59-67 flagellar segments. Body length 14-

20 mm. Hind tibia yellow, sometimes marked

with red apically. 3“ gastral tergite without

black-lateral’stripes: nn... ee 198:

At most 53 flagellar segments. Body length at

most 12 mm. Hind tibia marked with black.

Depression in front of the speculum for the

greater part with distinct longitudinal striae.

Sulcus of the propodeum for the greater part

transversely wrinkled. Hind tibia yellow api-

cally. Gaster yellow medially. 65-67 flagellar

segments. Body length 18-20 mm (See also

couplet 143). ..... falcator (Fabricius, 1775) 23

Depression in front of the speculum granu-

late-strigose in the ventral half. Sulcus of the

propodeum for the greater part coriaceous

196.

197:

198.

199:

or irregularly wrinkled, with transverse wrin-

kles posteriorly. Hind tibia reddish apically.

Gaster yellowish red medially. 59-66 flagel-

lar segments. Body length 14-17 mm (See also

COUPIER ZDF) Em nn ee ntenehitnanenae

RT obliterata (Holmgren, 1872) 23

Mesopleurum slightly coriaceous and dis-

tinctly shining between the punctures. Meta-

pleurum slightly sculptured and distinctly

shining centrally. Hind tibia black. Propo-

deum slightly depressed, in most parts irreg-

ularly wrinkled, with transverse wrinkles

posteriorly. 2" gastral tergite red on the pos-

terior 0.3. 3" and 4" tergites red (Both flagel-

lar tips broken off in the only available spec-

imen). Body length 9 mm. d unknown (See

Blsesenupler >30)... nu rsgennneeer

nearfehunae nitidipleuris Horstmann, spec. nov. ?

Mesopleurum distinctly coriaceous and rather

dull between the punctures. Metapleurum

distinctly punctured and/or distinctly coria-

ceous and rather dull centrally. .............. 197.

34-44 flagellar segments. Body length 6-9 mm.

TE UNE NER ORT ANE MERLTERERR 198.

45-53 flagellar segments. Body length 8-12 mm.

Propodeum with long or short transverse

wrinkles at least posteriorly. ............... 200.

Glymma lacking. 3" gastral tergite often with-

out black lateral stripes. Depression in front

of the speculum with distinct striae, which

merge with the punctation anteriorly. Hind

tibia black, marked with whitish yellow sub-

basally. Propodeum narrowly depressed,

with short transverse wrinkles in the longi-

tudinal furrow, with long transverse wrin-

kles posteriorly. 36-42 flagellar segments.

Body length 6-9 mm. d tentatively placed

here (See also couplets 208 and 222).............

re a ee schikotani Hinz, 1994 2 ?3

Glymma distinct. 3" gastral tergite with dis-

tinct black lateral stripes. Propodeum broad-

ly depressed, sulcus for the greater part with

IrePMlarnwrIn les. ee ernuntenneressasnene 199.

Genal index 0.3. Depression in front of the

speculum granulate-strigose or with fine stri-

ae, which merge with a coriaceous area with-

out punctation anteriorly. Hind tibia dark

yellowish brown medially, narrowly black

basally, broadly black apically. 34-40 flagel-

lar segments. Body length 6-8 mm................

ET IR A humilis (Förster, 1868) ?3

200.

201.

202.

Genal index 0.6. Depression in front of the

speculum with wrinkles, which merge with

the punctation anteriorly. Hind tibia com-

pletely black or tinged with brown medially.

42-44 flagellar segments. Body length 7-9 mm

(seeralsoreouplet 238)... nennen

ES Re a variator (Hinz, 1990) 23

2"d gastral tergite dorsally red on the posteri-

or 0.2-0.3, the red marks more extended lat-

erally. 3° and 4% tergites.red................... 201.

2nd gastral tergite red on the posterior 0.1. 4*

tersiteiblack 2 ar 22 202.

Propodeum distinctly depressed, transversely

wrinkled over almost its total breadth, with-

out longitudinal carinae. 50-53 flagellar seg-

ments. Body length 11-12 mm. .......................

N le, indistinctor Hinz, 1979 23

Propodeum distinctly depressed, with short

transverse wrinkles in the midline of the sul-

cus, in other parts irregularly wrinkled, lon-

gitudinal carinae indicated medially. 45-53

flagellar segments. Body length 8-12 mm (See

alsozeouplet 180). nz Se une

en ee xenocampta (Förster, 1868) 25

Metapleurum densely punctate on a coria-

ceous background, dull. Flagellum brown

apically in the $, dark brown to black in the

d. Hind tibia black, with whitish yellow marks

subbasally (these marks rarely reduced). Pro-

podeum distinctly depressed, with transverse

wrinkles in the longitudinal furrow, densely

rugose-punctate or transversely wrinkled be-

side the furrow. Glymma very small or lack-

ing. 3" gastral tergite usually completely red.

Epipleurum of 3" tergite completely fused

with the tergite. 45-51 flagellar segments.

Body lensth 9-10 mm. u... enssannees

ee contumator Hinz, 1979 25

Metapleurum finely punctate on.a very slight-

ly sculptured background, shining. Flagel-

lum black. Hind tibia black, tinged with

brown medially on the dorsal side. Propo-

deum not strongly depressed, longitudinal

carinae indicated medially, strongly trans-

versely wrinkled posteriorly, in other parts

coriaceous and with fine wrinkles. Glymma

distinct. 3" gasteral tergite red only laterally.

Epipleurum of the 3’ tergite indicated ante-

riorly, the black lateral stripes distinctly dis-

tant from the ventrolateral edge of the tergite.

48-49 flagellar segments. Body length 8 mm

(Seeralso eouplek >30) nn en ee

N nigridorsum Horstmann, spec. nov. ?d

203.

204.

209:

206.

Genal index smaller than 0.3. Mesopleurum

distinctly coriaceous between the punctures

(exception I DIamericani) ernennen 204.

Genalındex atleast 0a ee DA

49-50 flagellar segments. Body length 11 mm.

Hind tibia yellow medially, narrowly black

basally, broadly black apically. Propodeum

slightly depressed, partly transversely wrin-

kled, partly irregularly wrinkled in the broad

longitudinal sulcus. Glymma present, but

sometimes very small. 2" gastral tergite black

or dark brown posteriorly, marked with red

in front of the dark posterior edge. 3" tergite

with black lateral stripes. ? unknown (See

also’cOUuplet 86)... we annsessnnsnnsensnessenaun:

ren ans: parvicavata Horstmann, spec. nov. E

36-47 flagellar segments. Body length 6-9 mm.

Glymma often lacking. 2" gastral tergite to a

variable extent red posteriorly, rarely com-

Pletely black. 2. .......22u22.22222B20222021 2022220 rewean- 203.

Front and mid coxae whitish yellow, nar-

rowly darkened basally (possibly only in

the 4). Hind tibia yellow to yellowish brown,

narrowly black basally, slightly marked with

brown apically. Propodeum slightly de-

pressed, longitudinal carinae indicated me-

dially, transversely wrinkled in the broad

sulcus. Glymma« very small or lacking. 2”

gastral tergite red on the posterior 0.1-0.2. 3"

tergite with black lateral stripes. 40-44 flagel-

lar segments. Body length 7-8mm. ? un-

ET Tr EEE

ee flavopicta Horstmann, spec. nov. d

Front and mid coxae black, at most the tip of

the front coxa narrowly yellow. Hind tibia

marked with dark brown or black at least

apically (Genal index about 0.3; hind tibia

scarcely darkened apically: see D. opaca, cou-

Plet 220). ..2...2..0.2...2.200020eaucssennzasnsrazsasensananne 206.

Metapleurum coriaceous and with fine punc-

tures, the punctures distinctly discernible

(sculpture almost as in the adjoining parts of

the mesopleurum). Hind tibia black, some-

times tinged with dark brown subbasally in

the $, marked with white subbasally on the

dorsal side in the d. Glymma lacking. 2"

gastral tergite red on the posterior 0.3-0.5. 3"

tergite with black lateral stripes. 43-46 flagel-

lar segments. Body length 8-9 mm (See also

eoupletsl47, 185:and 215) 22 nn inc

LEN meridionator Aubert 1960 25

207.

208.

209.

Metapleurum coriaceous and finely rugose,

the punctures scarcely discernible among the

wrinkles (sculpture almost as in the adjoin-

ing parts of the propodeum). ............... 207.

Hind tibia black, sometimes tinged with dark

brown subbasally. Mandible black, tinged

with dark brown medially. Propodeum broad-

ly depressed, for the greater part irregularly

wrinkled, with short transverse wrinkles in

the shallow sulcus. Glymma lacking. 2” gas-

tral tergite red on the posterior 0.1-0.2. 3“

tergite with black lateral stripes. 40-44 flagel-

lar segments. Body length 7-8 mm. d un-

known. micrator Hinz, 1979 2

Hind tibia distinctly yellow to yellowish

brown medially (In D. schikotani the hind tib-

ia is often only narrowly marked with whit-

ish yellow ’subbasally). .... ee. ee 208.

Mandible black, at most tinged with brown

medially. Propodeum slightly depressed,

without longitudinal carinae medially, trans-

versely wrinkled in the narrow longitudinal

furrow at least posteriorly, rather coarsely

irregularly wrinkled beside the furrow. 24

gastral tergite red on the posterior 0.1-0.3. 3“

gastral tergite often without black lateral

stripes. 36-42 flagellar segments. Body length

6-9 mm. d tentatively placed here (See also

couplets.198 and 222). ....22.... nn re

ee: schikotani Hinz, 1994 2 ?&

Mandible distinctly marked with yellow me-

dially. 3’ gastral tergite with black lateral

SEHIPES. na ee 209.

Mesopleurum densely punctate on a distinct-

ly coriaceous background. Propodeum slight-

ly depressed, the narrow longitudinal fur-

row with short transverse wrinkles. Glymma

lacking. 2”! gastral tergite red on the posteri-

or 0.1. 3"! and 4" tergites red, both marked

with dark brown medially. 36-37 flagellar

segments. Body length 7 mm. d unknown

(See alsoreouplet 159). ne ee

... radiator Hinz & Horstmann, spec. nov. ?

Propodeum distinctly depressed and strong-

ly transversely wrinkled in the d, less dis-

tinctly depressed and slightly transversely

wrinkled in the ®, the sulcus rather broad in

both sexes. Glymma lacking or present as a

small shallow furrow. 2” gastral tergite usu-

ally red on the posterior 0.2-0.3. 3 and 4"

tergites red. In some specimens the gaster is

More darkened.........a nennen 210.

210. Mesopleurum densely punctate (rarely less

24.

212.

densely punctate) on a slightly coriaceous

and shining background, the background

sometimes smooth. Genal carina always

straight ventrally. Transverse carina obliter-

ated posteriorly. 36-43 flagellar segments.

Body length 6-9:mm.......22...sus:H:s23ese5@ni8 sone

ER americana (Ashmead, 1890) 23

Mesopleurum with disperse or rather dense

punctation on a distinctly coriaceous back-

ground. Genal carina sometimes bent for-

wards ventrally close to its junction with the

oral carina. Transverse carina complete. 42-47

flagellar segments. Body length 8-9 mm (See

also couple AZ)... n..0c2.:i020nniesisssserscgsasssangen

N LS N nn annexa (Förster, 1868) 25

Petiole without glymmae, in their place some-

times a change in the sculpture of the surface

or very fine longitudinal carinae or very shal-

low pits (Figs 37-38) (This character has strong

mftaspecitie varlability).'............r... 219:

Petiole with distinct small or large glymmae

ae SS en 223.

Hind tibia completely black, at most slightly

tinged with brown medially. 3“ gastral tergite

with black lateral stripes. ................ 213.

Hind tibia white to yellowish brown medial-

ly, to a variable extent darkened basally and

EICHE 217.

. 2X sastral tergite black dorsally, at most red

onstheyposterion 0. 4.2... u.2.2. 214.

2"d sastral tergite dorsally red on the posteri-

SEO ON er enter 245.

. Propodeum scarcely depressed, without lon-

gitudinal carinae, with short transverse wrin-

kles in a weakly indicated narrow longitudi-

nal furrow, irregularly wrinkled beside the

furrow. 2" gastral tergite with a black poste-

rior edge, red laterally. 3“ and 4" tergites

red, the 3”! marked with black dorsally. Pos-

terior sternites red. 42-43 flagellar segments.

Body length 9 mm. d unknown. ............

BE meritor Hinz & Horstmann, spec. nov. ?

Propodeum somewhat depressed, with lon-

gitudinal carinae medially, transversely wrin-

kled between the carinae over the total

breadth of the sulcus. 2" gastral tergite usu-

ally red on the posterior 0.1, sometimes com-

pletely black dorsally. 3 tergite red, some-

times marked with black dorsally. 4" tergite

for the greater part black. Posterior sternites

black. 41-43 flagellar segments. Body length

21».

216.

218.

9-10 mm (Variety: 39 flagellar segments. Body

lenethizimm) ar obtutor Hinz, 1994 23

Propodeum distinctly depressed, with short

transverse wrinkles over the total length of

the narrow longitudinal furrow, irregularly

wrinkled beside the furrow (Fig. 34). 2" gas-

tral tergite dorsally red on the posterior

0.3-0.5. 43-46 flagellar segments. Body length

8-9 mm (See also couplets 147, 185 and 206).

EN EEE meridionator Aubert, 1960 ?3

Propodeum slightly depressed, without a nar-

row longitudinal furrow. 2" gastral tergite

red on the posterior 0.4-0.6.........e.. 216.

Temples less strongly constricted behind the

eyes, tangential lines on eyes and temples

intersect on the propodeum. Antennal carina

with a short dorsal projection (nose-like). Pro-

podeum with long transverse wrinkles pos-

teriorly. 44-48 flagellar segments. Body length

10-1: mm (Seealso’ couple #189). rn...

TEN EEE stygia (Förster, 1868) ?5

Temples distinctly constricted behind the

eyes, tangential lines on eyes and temples

usually intersect on the scutellum. Antennal

carina without a dorsal projection, but with

distinct radial wrinkles instead. Propodeum

irregularly wrinkled posteriorly. 43-45 flag-

ellar segments (Variety: 48 segments). Body

lensth I mamnn 2 nase nee srasnpaengennes

templator Hinz & Horstmann, spec. nov. ?d

34 flagellar segments. Body length 5 mm.

Hind tibia whitish yellow medially, narrow-

ly dark brown basally, broadly dark brown

apically. Propodeum slightly depressed, lon-

gitudinal carinae indicated beside the broad

and shallow sulcus, for the greater part

coarsely and irregularly wrinkled. 2" gastral

tergite dorsally red on the posterior 0.1. 3“

and 4 tergites red, marked with black dor-

sally:>? unknewm: zurstsesssesenesessesmsennee

Ga EN 0 nanus Horstmann, spec. nov. d

38-48 flagellar segments. Body length 7-11 mm.

RR EEE ERENEN NE IL EN 218:

Speculum and depression in front of the

speculum granulate-strigose, the striae merg-

ing with a coriaceous area without puncta-

tion anteriorly. Hind tibia yellowish brown

medially, narrowly black basally, marked

with dark brown apically. Propodeum some-

what depressed, transversely striate posteri-

orly in the broad sulcus, in other parts irreg-

ularly wrinkled. 2" gastral tergite red on the

219.

220.

221.

222.

posterior 0.1. 3"! and 4" tergites red. 46-47

flagellar segments. Body length 8 mm. 2 un-

known. ..... nebulosa Horstmann, spec. nov. d

Depression in front of the speculum with

distinct striae and wrinkles, which merge

with the punctation anteriorly, and/or hind

tibia marked with white medially........... 219:

43-48 flagellar segments. Body length 9-11 mm.

Hind tibia dark yellowish brown medially,

narrowly and indistinctly darkened basally

and apically (43-46 flagellar segments; hind

tibia black or marked with white subbasally

on the dorsal side: see D. meridionator, cou-

PIEB21S)....0. een 220.

38-42 flagellar segments. Body length 7-9 mm.

Hind tibia broadly dark brown or black api-

Gallllıyn. nn A nn een 227.

Ocellar index about 1.0. 2" gastral tergite red

on the posterior 0.3. 3" and 4" tergites red,

rarely marked with black dorsally. Hind gas-

tral sternites black, 43-47 flagellar segments.

Body length 9-10 mm (See also couplet 240).

IHR IH opaca (Thomson, 1887) ?3

Ocellar index 0.5. 2" gastral tergite black

dorsally, marked with red laterally. 3 and

4" tergites red. Hind gastral sternites reddish

brown. 48 flagellar segments. Body length

Immo unknoWme...eenernremuersknne:

Bar: ternata Hinz & Horstmann, spec. nov. 9

Hind tibia yellowish brown to brown medi-

ally, narrowly dark brown to black basally,

broadly dark brown to black apically. Propo-

deum not depressed, finely and irregularly

wrinkled, often with longitudinal wrinkles

posteriorly, without longitudinal carinae me-

dially. 2” gastral tergite dorsally black or red

on the posterior 0.1, marked with red lateral-

ly. 3" and 4" tergites for the greater part red,

to a variable extent marked with black dor-

sally. 39-46 flagellar segments. Body length

7-9 mm (See also couplets 137 and 234). ......

EEE RT subimpressa (Förster, 1868) ?3

Hind tibia whitish medially, narrowly black

basally, broadly black apically. Propodeum

distinctly depressed, distinctly transversely

wrinkled in the shallow sulcus (Hind tibia

yellowish brown medially; propodeum dis-

tinctly depressed: see D. americana). ...... 222.

Malar space index 0.3-0.4. Glymma indicated

as a very small pit. 1* gastral sternite with

distinct transverse striae at least in the poste-

rior half. 2” gastral tergite red on the poste-

223.

224.

225:

226.

rior 0.1. 3” tergite red only laterally. 4" tergite

completely black. 3" tergite with black later-

al stripes. 36-41 flagellar segments. Body

length 6-10 mm (See also couplets 146 and

DIE A amurator Hinz 1985 23

Malar space index 0.6. Glymma lacking. 1*

gastral sternite for the greater part smooth.

2"d gastral tergite red on the posterior 0.1-0.2.

3" tergite red. 4" tergite broadly red anteri-

orly. 3"! tergite often without black lateral

stripes. 36-42 flagellar segments. Body length

6-9 mm. d tentatively placed here (See also

coupletss198 and 208) rn ee

San ERROR schikotani Hinz, 1994 2 ?&

58-66 flagellar segments. Body length 15-17 mm.

Mesopleurum slightly coriaceous and shin-

ing between the punctures. Hind tibia yel-

low or yellowish brown medially, narrowly

black basally, at most slightly darkened api-

cally. 3" gastral tergite without black lateral

STEIPES. ne cennec e e E 224.

35-52 flagellar segments. Body length 6-12 mm.

Mesopleurum distinctly coriaceous between

the punctures. Hind tibia dark brown or black

atleast apically (exceptions: D. opaca, D. opaco-

I een 226.

Metapleurum densely rugose-punctate on a

smooth background. 2" gastral tergite black

dorsally, red laterally. 3 to 7" tergites al-

most completely red. 58-66 flagellar segments.

Body length 15-16 mm. d unknown. ............

BR EEE scutellator Hinz, 1979 ?

Metapleurum rugose or punctate on a coria-

ceous background. At least the 6" and 7'"

gasteral tergites for the greater part black. ..

Ban Saba Suse bene neun ans Tre EEE 225.

Hind femur marked with red apically. Hind

tibia yellow, scarcely darkened apically. 2”

gastral tergite black dorsally, marked with

red laterally. 3" to 4" or 5" tergites red. 59-66

flagellar segments. Body length 15-17 mm

(See also'couplet Tb) "een

BEER EEN AL in obliterata (Holmgren, 1872) 23

Hind femur black. Hind tibia dark yellowish

brown. 2" to 7" gastral tergites black dorsal-

ly, to a variable extent red laterally. 61 flagel-

lar segments. Body length 15 mm. d un-

known. ae RE EE

.... scriptor Hinz & Horstmann, spec. nov.

35 flagellar segments. Body length 6-7 mm.

2"d gastral tergite red on the posterior 0.5-0.7.

Dass besessen trans ee 227.

227,

228.

229.

230.

37-52 flagellar segments. Body length 7-12 mm.

2"! gastral tergite less extensively marked

with red at least dorsally. ............. 228.

Flagellum with all segments distinctly long-

er than wide. Hind femur marked with red.

Hind tibia dark brown to black. Propodeum

with a narrow longitudinal furrow, with short

transverse wrinkles in the furrow. 1* gastral

tergite 5.2x as long as wide. 2" tergite red on

the posterior 0.7. 35 flagellar segments. Body

lenefh 2. mm. unknown. ......22...ut2:2

onosseseiten longistilus Horstmann, spec. nov. &

Flagellum with the shortest segments wider

than long. Hind femur black. Hind tibia

broadly whitish medially, black basally and

apically. Propodeum with a broad shallow

sulcus, irregularly wrinkled, more longitudi-

nally wrinkled posteriorly. 1" gastral tergite

3.4x as long as wide. 2"4 tergite red on the

posterior 0.5. 35 flagellar segments. Body

leneth.61mm.°2 unknown: rennen

ee, parva Horstmann, spec. nov. d

Metapleurum with fine but distinct and dense

punctation on a coriaceous background (sculp-

ture about as in the adjoining parts of tie

mesopleurum). 49-52 flagellar segments. ....

Un nes aspscznunnne ragen ieheze zen asranehsänsnene 229:

Metapleurum finely and densely rugose-

punctate, the interspaces not coriaceous

(sculpture about as in the adjoining part of

the propodeum) (In D. variator with 42-44

flagellar segments the interspaces are coria-

eous) re anal Eh 232:

Posterior gastral tergites from the 3" tergite

onwards red, at most tinged with brown dor-

sally. Hind tibia light yellowish brown medi-

ally, narrowly black basally, broadly black

apically. Propodeum with a broad shallow

sulcus, with long transverse wrinkles poste-

riorly. 51 flagellar segments. Body length

U mMMES. unkmawm...n.eeeneneeeennn

Bra rufoapicalis Horstmann, spec. nov. 2

Posterior gastral tergites completely black or

distinctly marked with black dorsally... 230.

Hind tibia dark brown medially, black basal-

Iy and apically. Propodeum slightly de-

pressed, the longitudinal carinae indicated

medially, for the greater part irregularly wrin-

kled, somewhat transversely wrinkled pos-

teriorly. All gastral tergites black dorsally, 3"

and 4" tergites laterally red in the ?, 2" to 5"

tergites laterally red in the 4. Genital clasper

231.

232.

233:

of the d yellowish brown. 48-49 flagellar seg-

ments. Body length 8 mm (See also couplet

DD Ar ee ne ee ea

et nigridorsum Horstmann, spec. nov. ?d

Hind tibia yellow to yellowish brown medi-

ally. At least the 3" gastral tergite completely

idee Le RE ER PSAIENEBRELE. 231:

Propodeum deeply depressed, the longitu-

dinal carinae indicated only posteriorly, dis-

tinctly transversely striate in the longitudi-

nal sulcus, area superomedia only coriaceous.

2"d gastral tergite dorsally red on the posteri-

or 0.2, almost completely red laterally in front

ofthe brown posterior edge. 3"! and 4" tergites

red. Posterior tergites and sternites black.

Genital clasper black. 52 flagellar segments.

Body length 10 mm. ? unknown. ..............

ER levibasis Horstmann, spec. nov. d

Propodeum slightly depressed, the longitu-

dinal carinae indicated medially, irregularly

wrinkled, with some longitudinal wrinkles

posteriorly. 2" gastral tergite red laterally

(including the posterior edge). 3X tergite red.

4" tergite red with a black dorsal mark. Pos-

terior tergites black dorsally and red lateral-

ly. Posterior sternites red. 49 flagellar seg-

ments. Body length 11 mm. d unknown. ....

.. ventrator Hinz & Horstmann, spec. nov.

Propodeum not or almost not depressed, ir-

regularly wrinkled, often with one or more

longitudinal wrinkles posteriorly. ......... 233.

Propodeum distinctly depressed, often trans-

versely wrinkled in some parts of the longi-

tudinal furrow, sometimes with a short lon-

gitudinal keel posteriorly (D. nitidipleuris, in

which the propodeum is scarcely depressed,

but distinctly transversely wrinkled posteri-

only, isameluded.here)n.n.22rs0 a... 233.

Propodeum coarsely wrinkled, almost com-

pletely areolated, but the carinae hardly dis-

cernible among the wrinkles. Metapleurum

with distinct and very dense punctures, not

coriaceous. Hind tibia yellow medially, nar-

rowly black basally, broadly black apically.

3" gastral tergite without black lateral stripes.

43 flagellar segments. Body length 10 mm. ?

unknown (See also couplet 172). ....................

en inconspicua Horstmann, spec. nov. d

Propodeum finely wrinkled. 3“ gastral tergite

with black lateral stripes. .................... 234.

234. Hind tibia black. 2" gastral tergite red on the

238:

236.

posterior 0.2-0.3. 3’! and 4" tergites red, some-

times 4" tergite marked with black posterior-

ly. 48-52 flagellar segments. Body length

9-10 mm (See also couplet 104).....................

EEE ERDE fundator Hinz, 1990 ?5

Hind tibia yellowish brown to brown medi-

ally, black basally and apically. 2" gastral

tergite completely black dorsally or red on

the posterior 0.1, marked with red laterally.

34 and 4" tergites for the greater part red, to

a variable extent marked with black dorsally.

39-46 flagellar segments. Body length 7-9 mm

(see also’eouplets 137 and 22). ................

lehren subimpressa (Förster, 1868) 23

Depression in front of the speculum granu-

late-strigose, the striae merging with a nar-

row coriaceous area without punctation an-

teriorly. Hind tibia whitish medially on the

dorsal side, black on the ventral side and

basally and apically (sometimes whitish me-

dially also on the ventral side). Propodeum

broadly and distinctly depressed, for the

greater part finely and irregularly wrinkled

and with some short transverse wrinkles in

the 2, more deeply depressed, more strongly

wrinkled and with the longitudinal carinae

indicated in the d. 2" gastral tergite red on

the posterior 0.1. 3" tergite red only laterally.

36-41 flagellar segments. Body length 6-10 mm

(see also couplets 146 and’ 222).................

AREA amurator Hinz, 1985 23

Depression in front of the speculum with

fine or distinct striae, which merge with the

punctation anteriorly. Hind tibia yellowish

brown to dark brown medially, brown or

black basally and apically, or completely

black. Gaster usually marked with red to a

STEAFEL TORTEN: esecdersnsnne-sSensnknenncessgrsgesnessenren 236.

2"d gastral tergite with a dark brown to black

posterior edge, to a variable extent marked

with red in front of the posterior edge, or

completely black dorsally. Hind tibia yel-

lowish brown to dark brown medially or

completely black. Longitudinal carinae of pro-

podeum usually distinct medially, sometimes

also anteriorly, the sulcus between the longi-

tudinal carinae irregularly wrinkled or trans-

versely wrinkled. Glymma large. 3“ and 4"

gastral tergites red, to a variable extent

marked with black. 37-48 flagellar segments.

Bodyıleneih 712mm Mena

Aagemenatasunier glauca caliginosa (Walley, 1940) ?4

237

238.

239.

2"! gastral tergite red on the posterior 0.1-0.3,

the posterior edge rarely slightly darkened

(Glymma very small or lacking, 2" gastral

tergite red on the posterior 0.1 or completely

black dorsally: see D. obtutor, couplet 214)..

RE NEO en. 237.

Hind tibia black, at most tinged with dark

brownmedialle 22.272222 238.

Hind tibia broadly yellowish brown medial-

ly, narrowly and often indistinctly darkened

basally and apically. 3“ and 4" tergites for

the greater part or completely red. ........ PSEN

Mesopleurum distinctly coriaceous and dull

between the punctures. Metapleurum coria-

ceous and rugose-punctate, rather dull. Pro-

podeum slightly depressed, finely coriaceous

and rugose, usually with a short longitudi-

nal keel posteriorly and with transverse wrin-

kles beside the keel, in the d longitudinal

carinae sometimes indicated. Glymma large.

2"! gastral tergite red on the posterior 0.1-0.3.

3" tergite red. 4" tergite marked with red or

completely black. 42-44 flagellar segments.

Body length 7-9 mm (See also couplet 199).

KRUSE variator (Hinz, 1990) 23

Mesopleurum slightly coriaceous and shin-

ing between the punctures. Metapleurum

slightly rugose, distinctly shining centrally.

Propodeum broadly and shallowly depressed,

without alongitudinal keel posteriorly. Glym-

ma rather large. 2" gastral tergite red on the

posterior 0.3. 3"? and 4" tergites red. At least

42 flagellar segments (Both flagellar tips bro-

ken off in the only available specimen). Body

length 9 mm. d unknown (See also couplet

196). ... nitidipleuris Horstmann, spec. nov. ?

Penultimate flagellar segment 1.0-1.1x as long

as wide. Temple with very fine and disperse

punctation medially. Hind tibia broadly yel-

low to yellowish red medially, distinctly dark

brown to black basally and apically. Hind

basitarsus in lateral view for the greater part

dark brown to black. Propodeum slightly

and rather broadly depressed, for the greater

part transversely wrinkled and with weak

longitudinal carinae in the ?, more deeply

depressed, coarsely transversely wrinkled

and with strong longitudinal carinae in the

d. Glymma rather large. 2”! gastral tergite

dorsally red on the posterior 0.2-0.3, some-

times the posterior edge slightly marked with

brown. 39-41 flagellar segments. Body length

a remota (Förster, 1868) ?3

- Penultimate flagellar segment 1.2-1.4x aslong

as wide. Hind tibia broadly yellowish brown

to reddish brown medially, narrowly medi-

um brown basally and apically, the apical

mark sometimes indistinct. Hind basitarsus

in lateral view usually for the greater part

light brown. Propodeum somewhat de-

pressed, with short transverse wrinkles in

the rather narrow longitudinal furrow, irreg-

ularly wrinkled beside the furrow, without

longitudinal carinae medially. 2” gastral

tergite red on the posterior 0.2-0.3. ........ 240.

240. Temple with fine and rather dense puncta-

tion medially. Propodeal spiracle rather large,

4.0x as long as wide. Glymma very small,

sometimes lacking. 43-47 flagellar segments.

Body length 9-10 mm (See also couplet 220).

BEN Leni e.ar opaca (Thomson, 1887) ?3

- Temple with very fine and disperse puncta-

tion medially. Propodeal spiracle small, 2.5x

as long as wide. Glymma small. 39-40 flagel-

lar segments. Body length 7 mm. d unknown.

Mesautelichsseleuscndsstsentgbessen opacoides Hinz, 1985 2

Descriptions of species

Dusona admontina (Speiser, 1908)

Description. ?: Body length 7-9 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.8 (Fig. 1). Frons slightly de-

pressed, coriaceous, very finely and rather dense-

ly punctate, with a fine median longitudinal cari-

na, with rather long radial wrinkles, which extend

to 0.4 the distance to the median ocellus. Antennal

carina low and narrow. Face coriaceous, finely

and densely rugose and rugose-punctate. Clypeus

flat, coriaceous, with fine rather dense or disperse

punctation, its apex truncate medially. Malar space

index 0.4. Genal carina low and almost straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.3. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 37-39 segments, filiform,

the apical 0.2 attenuate, 2" segment 3.0x, shortest

segment 1.0x, penultimate segment 1.0x as long

as wide. Pronotum densely rugose laterally on the

dorsal 0.1-0.2, with longitudinal wrinkles ventral-

ly. Mesoscutum coriaceous and densely rugose-

punctate. Scutellum coriaceous, finely and rather

densely punctate, with lateral carinae on the ante-

rior 0.4. Prescutellar groove for the greater part

with fine punctation on a smooth background,

with longitudinal striae laterally. Prepectal carina

low, ventral part merging with the pleural part,

the latter complete, joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum coriaceous, dull, densely rugose-

punctate. Speculum coriaceous and dull. Depres-

sion in front of the speculum with rather long

longitudinal wrinkles, which merge with the punc-

tation anteriorly. Metapleurum coriaceous and

finely and densely rugose. Pleural carina distinct.

Areolet small, stalked, receiving the 2" recurrent

vein distinctly distad of the middle (Fig. 25).

Nervellus vertical, weakly intercepted. Nervellar

index 0.7. Tibial spur index 0.6. Hind femur 4.8x

as long as high. Hind tarsal claws short, with

distinct pectinate teeth basally and medially. Pro-

podeum extending to 0.4 ofthe hind coxae, scarcely

depressed, completely areolated, for the greater

part irregularly wrinkled, with distinct longitudi-

nal wrinkles posteriorly. Propodeal spiracle small,

2.6x as long as wide. Spiracular carina narrow.

Spiracular index 0.9. 1* gastral tergite 4.0x, 2"

tergite 1.9x as long as wide. Petiole without glym-

mae, with indistinct narrow lateral areas, smooth

in front of these areas. Petiolar suture fine but

complete. 1* gastral sternite coriaceous. Thyrid-

ium elongate. Thyridial index 1.3. Epipleurum

separated from the 3" gastral tergite, the crease

marked with black. Epipleural index 0.8. Gaster

distinctly but not strongly compressed posterior-

ly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible almost completely yellow (teeth black). Tegu-

la yellow or light brown. Wings hyaline. Stigma

dark brown. Front leg from the trochanter or the

tip of the coxa onwards yellowish brown. Mid

femur completely yellowish brown or yellowish

brown on the apical 0.2. Mid tibia and tarsus

yellowish brown. Hind tibia yellowish brown,

sometimes narrowly black basally. Hind tarsus

yellowish brown to brown. 2" gastral tergite red

on the posterior 0.2-0.3. Tergites from the 3“ on-

wards red, more or less indistinctly marked with

brown dorsally. Gastral sternites red. Pilosity

whitish.

d: Flagellum with 34-35 segments. Front and

mid legs yellow, mid coxa marked with black

basally. Hind femur reddish brown to dark brown.

Genital clasper red. In other characters similar to

the 2.

Material (2??, 238): Russia: Khabarovsk Terr.: Khekh-

tsir near Khabarovsk; Primorye: Chuguyevka, Khasan;

Kuril Isl.: Shikotan (PET).

Dusona aemula (Förster, 1868)

Description. 2: Body length 7-8mm. Temples

slightly constricted behind the eyes, 0.7-0.8x as

long as the width of an eye, tangential lines on

eyes and temples intersect on the propodeum.

Ocellar index 0.8. Frons slightly depressed, coria-

ceous, with fine dense wrinkles, with a very fine

median longitudinal carina, without radial wrin-

kles. Antennal carina low and narrow. Face and

dorsal half of the clypeus densely and finely ru-

gose-punctate. Clypeus flat, coriaceous and with

rather dense punctation ventrally, its apex dis-

tinctly convex medially. Malar space index 0.6.

Genal carina low ventrally, distinctly bent for-

wards and joining the oral carina at the base of the

mandible at a right angle. Temple finely coria-

ceous and with fine disperse punctation medially.

Flagellum with 26-29 segments, slightly but dis-

tinctly fusiform, 2” segment 1.9x, shortest seg-

ment 0.9x, penultimate segment 1.0x as long as

wide. Pronotum with dense punctation and longi-

tudinal wrinkles laterally on the dorsal 0.3, with

longitudinal wrinkles without punctation ventral-

ly. Mesoscutum and scutellum coriaceous, dense-

ly and finely rugose-punctate. Scutellum with lat-

eral carinae on the anterior 0.4. Prescutellar groove

with short longitudinal striae. Ventral part of the

prepectal carina merging with the transverse car-

ina, both distinctly raised, the former deeply emar-

ginate medially. Pleural part of the prepectal car-

ina low, almost joining the anterior edge of the

mesopleurum. Mesopleurum finely coriaceous and

densely punctate. Speculum with fine oblique stri-

ae. Depression in front of the speculum with fine

and long longitudinal striae, which merge with

the punctation anteriorly. Metapleurum rugose-

punctate, with fine and very dense punctation.

Pleural carina distinct. Areolet rather small, short-

stalked, receiving the 2" recurrent vein distad of

the middle. Nervellus slightly inclivous, intercept-

ed. Nervellar index 0.8. Tibial spur index 0.6. Hind

femur 4.5x as long as high. Hind tarsal claws

rather long, with short pectinate teeth basally.

Propodeum extending to the middle of the hind

coxae, broadly and rather deeply depressed, trans-

versely wrinkled almost all over the broad sulcus,

longitudinal carinae and costulae almost complete,

but partly covered with wrinkles. Propodeal spir-

acle small, 2.4x as long as wide. Spiracular carina

strong, partly widened. Spiracular index 0.5-0.8.

1% gastral tergite 4.3x, 2"! tergite 1.5x as long as

wide. Petiole with small and somewhat elongate

glymmae at the anterior edge of narrow lateral

areas, smooth in front of the glymmae. Sometimes

only the narrow lateral areas present (Fig. 37).

Petiolar suture for the greater part obliterated.

1% gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3 gastral tergite, the latter with short

and indistinct black lateral stripes. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly yellowish brown dorsally and

medially. Tegula yellowish brown. Wings hya-

line. Stigma yellowish medially. Femora, tibiae

and tarsi of the front and mid legs yellowish red,

the tarsi darkened apically. Hind tibia dark yel-

lowish brown medially, narrowly black basally

and apically. Hind femur and hind tibia some-

times completely red. 2" gastral tergite red on the

posterior 0.2. 3’ and 4" tergites red, the 4" dorsal-

ly marked with black apically or completely black.

Pilosity whitish.

d: Flagellum with 28-30 segments. Propodeum

with stronger carinae and wrinkles. Mandible

broadly yellowish brown medially. In other char-

acters similar to the 2.

Material (1°): Kazakhstan: Dzhabagly River ESE of

Chimkent, 1700 m (PET).

Dusona affinitor Hinz, 1979

Description. 2: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.8. Frons not depressed, coriaceous, dense-

ly rugose and rugose-punctate, without a median

longitudinal carina and without distinct radial

wrinkles. Antennal sockets rather close to each

other. Antennal carina low and narrow. Face and

dorsal half of the clypeus coriaceous, densely ru-

gose and rugose-punctate. Clypeus slightly round-

ed, coriaceous and with fine rather dense puncta-

tion ventrally, its apex truncate medially. Malar

space index 0.3. Genal carina low and straight

ventrally, joining the scarcely raised oral carina at

an acute angle. Genal index 0.3. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 35-37 segments, slightly

fusiform, 2” segment 1.5x, shortest segment 0.9x,

penultimate segment 1.0x as long as wide. Prono-

tum with longitudinal wrinkles laterally. Meso-

scutum and scutellum coriaceous, with dense or

very dense punctation. Scutellum with lateral car-

inae on the anterior 0.4. Prescutellar groove al-

most completely smooth. Ventral part of the pre-

pectal carina merging with the transverse carina,

both carinae slightly raised, the former not emar-

ginate medially. Pleural part of the prepectal car-

inae present as a wrinkle medially, obliterated or

covered with wrinkles dorsally and ventrally.

Mesopleurum distinctly and very densely punc-

tate on an almost smooth background. Speculum

finely granulate-strigose, shining. Depression in

front of the speculum with fine and long longitu-

dinal striae, which merge with the punctation an-

teriorly. Metapleurum very finely and densely

rugose or rugose-punctate, the wrinkles smooth.

Pleural carina distinct. Areolet large, narrowly

sessile or pointed, receiving the 2” recurrent vein

atthe middle. Nervellus distinctly inclivous, weak-

ly intercepted. Nervellar index 0.7-0.8. Tibial spur

index 0.7. Hind femur 4.8x as long as high. Hind

tarsal claws short, with few short pectinate teeth

basally. Propodeum extending to 0.4 of the hind

coxae, a narrow longitudinal furrow weakly indi-

cated, for the greater part irregularly wrinkled,

with some transverse wrinkles in the longitudinal

furrow, sometimes with a short longitudinal keel

posteriorly, carinae almost completely obliterat-

ed. Propodeal spiracle small, 1.7-2.0x as long as

wide. Spiracular carina narrow. Spiracular index

0.9. 1% gastral tergite 4.2x, 2" tergite 1.7% as long

as wide. Petiole without glymmae, with narrow

lateral areas, smooth in front of these areas. Petio-

lar suture almost completely obliterated. 1% gas-

tral sternite for the greater part smooth, with few

transverse striae posteriorly. Thyridium elongate.

Thyridial index about 2.0. Epipleurum not sepa-

rated from the 3" gastral tergite, the latter without

distinct black lateral stripes. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.7.

Colour: Black. Mandible tinged with brown

subapically. Wings hyaline. Stigma dark brown

medially. Front femur yellowish brown dorsally

and apically. Front tiba and tarsus yellowish

brown. Mid femur yellowish brown on the apical

0.2. Mid tibia yellowish brown. Mid tarsus dark

brown. Hind tibia tinged with dark brown medi-

ally. 3:4 gastral tergite red, narrowly tinged with

black dorsally. Pilosity whitish.

d unknown.

Material (2??): Russia: Kuril Isl.: Tretyakovo on Ku-

nashir (PET). Japan: Hokkaido: Sapporo (SAP).

Dusona alpinator Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Kirgiziya, Alayskiy Mts.,

2800 m, 30.5.1956 (MOS). — Paratypes: 1? Kaza-

khstan, Alma-Ata, 1400-1800 m, 3.-12.6.1951 (Hinz /

ZSM); 13 Kirgiziya, Kirgizskiy Mts., Issykata, 1800

m, 19.5. 1951 (MOS).

Description. 2: Body length 9-10 mm. Temples

distinctly but not strongly constricted behind the

eyes, 0.9x as long as the width of an eye, tangen-

tial lines on eyes and temples intersect on the

anterior part of the propodeum. Ocellar index 1.0.

Frons scarcely depressed, coriaceous, finely ru-

gose-punctate, with a fine median longitudinal

carina, without distinct radial wrinkles. Antennal

carina low and narrow. Face and dorsal half of

the clypeus coriaceous, very densely punctate.

Clypeus flat, coriaceous and densely punctate ven-

trally, its apex almost truncate medially. Malar

space index 0.4. Genal carina low, almost straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.4. Temple coria-

ceous and with fine disperse punctation medially.

Flagellum with 44 segments, stout, slightly fusi-

form 22d segment 1.3x, shortest segment 0.9x,

penultimate segment 1.0x as long as wide. Prono-

tum with very dense punctation laterally on the

dorsal 0.4, for the greater part with irregular wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous and with dense punctation. Scutellum with

lateral carinae on the anterior 0.7. Prescutellar

groove smooth medially, with short longitudinal

wrinkles laterally. Prepectal carina low, its ventral

part merging with the pleural part, the latter al-

most joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

coriaceous, strongly and densely punctate. Specu-

lum coriaceous and granulate-strigose. Depres-

sion in front of the speculum with short longitudi-

nal wrinkles, which merge with the punctation

anteriorly. Metapleurum coriaceous, with fine and

dense irregular wrinkles. Pleural carina fine, part-

ly indistinet and covered with wrinkles. Areolet

large, pointed, receiving the 2" recurrent vein at

the middle. Nervellus somewhat inclivous, inter-

cepted. Nervellar index 0.7-0.8. Tibial spur index

0.7. Hind femur 5.1x as long as high. Hind tarsal

claws long, with short pectinate teeth basally. Pro-

podeum extending to the basal 0.4 of the hind

coxae, slightly depressed, almost without carinae,

with short transverse wrinkles medially in the

shallow broad sulcus, irregularly wrinkled in oth-

er parts. Propodeal spiracle small, 2.0x as long as

wide. Spiracular carina obliterated. Spiracular in-

dex 1.1. 1* gastral tergite 3.9x, 2" tergite 1.4x as

long as wide. Petiole smooth laterally, without

glymmae and longitudinal areas. Petiolar suture

partly indicated. 1* gastral sternite for the greater

part smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum not separated from the 3°“

gastral tergite, sometimes discernible as a small

lobe anteriorly, 3" tergite with distinct black later-

al stripes well above the ventrolateral edge. Gaster

distinctly but not strongly compressed posterior-

ly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Wings slightly darkened. Stigma light brown me-

dially. All femora and tibiae red. Front and mid

tarsi dark brown, hind tarsus black. 2" gastral

tergite red on the posterior 0.6-0.7. 3" and 4"

tergites red. 5" tergite narrowly red anteriorly.

Pilosity on head and thorax brown.

d: Temples broad and not much constricted

behind the eyes, 1.1x as long as the width of an

eye, tangential lines on eyes and temples intersect

on the 1" gastral segment. Flagellum with 46 seg-

ments. In other characters very similar to the ?

and therefore placed to the same species, despite

the different shape of the temples.

Variety: Body length 7 mm. Flagellum with 42

segments. Mandible, tip of the front coxa and

front trochanter yellow. Front and mid tarsi yel-

lowish red. Pilosity whitish. Material: 19 Russia,

Altaysk Terr., Biysk (PET). Probably this 8 be-

longs to another species.

Dusona alticola (Gravenhorst, 1829)

Description. 9: Body length 6 mm. Temples slight-

ly constricted behind the eyes, 0.9x as long as the

width of an eye, tangential lines on eyes and tem-

ples intersect on the propodeum. Ocellar index

0.9. Frons slightly depressed, coriaceous, with fine

disperse punctation, almost not wrinkled, with a

fine and shortened median longitudinal carina,

without radial wrinkles. Antennal carina slightly

raised, the rim slightly widened, with with fine

transverse striae. Face and dorsal 0.3 ofthe clypeus

coriaceous, for the greater part densely rugose-

punctate, face almost not punctate medially.

Clypeus flat, coriaceous and with rather dense

punctation ventrally, its apex slightly convex me-

dially. Malar space index 0.5. Genal carina low

ventrally, slightly bent forwards and sidewards,

joining the low oral carina almost at a right angle.

Genal index 0.1. Temple finely coriaceous, with

very fine disperse punctation medially. Flagellum

with 26-27 segments, distinctly fusiform, 2"4 seg-

ment 2.0x, penultimate segment (which is the

shortest segment) 0.7%x as long as wide. Pronotum

coriaceous, punctate and wrinkled laterally on the

dorsal 0.3, with longitudinal wrinkles ventrally.

Mesoscutum coriaceous, finely and densely punc-

tate. Scutellum finely coriaceous, for the greater

part with fine and disperse punctation, with later-

al carinae on the anterior 0.4. Prescutellar groove

smooth medially, with longitudinal striae lateral-

ly. Ventral part of the prepectal carina merging

with the transverse carina, both distinctly raised,

the former distinctly emarginate medially. Pleu-

ral part of the prepectal carina low, complete or

almost complete. Mesopleurum coriaceous, with

disperse or rather dense punctation. Speculum

very finely coriaceous, shining. Depression in front

of the speculum with rather long longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum shining, finely rugose-punctate

medially. Pleural carina distinct. Areolet rather

small, pointed, receiving the 2" recurrent vein

distinctly distad of the middle. Nervellus slightly

inclivous, weakly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 4.9x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, rather broadly and deeply de-

pressed, sulcus with long transverse wrinkles, car-

inae complete or almost complete. Propodeal spir-

acle small, 2.4x as long as wide. Spiracular carina

narrow. Spiracular index 0.7. 1% gastral tergite

3.7%x, 2"d tergite 1.4x as long as wide. Petiole with

distinct glymmae, which are bordered by longitu-

dinal carinae dorsally, smooth in front of the glym-

mae. Petiolar suture almost complete. 1* gastral

sternite finely coriaceous and rugose. Thyridium

oval. Thyridial index 0.8. Epipleurum not separat-

ed from the 3" gastral tergite, the latter with dis-

tinct black lateral stripes well above the ventrola-

teral edge. Gaster distinctly compressed posteri-

orly. Ovipositor index 1.3, ovipositor sheaths

distinctly widened apically (in lateral view).

Colour: Black. Tegula light brown. Wings hy-

aline. Stigma yellowish brown medially. Front

and mid legs from the femur onwaräs light red,

the femora sometimes tinged with brown ventral-

ly. Hind femur reddish brown to brown. Hind

tibia broadly yellowish brown medially, scarcely

marked with brown basally, rather narrowly

marked with brown apically. Hind tarsus dark

brown, marked with yellowish brown basally. 2“

gastral tergite red on the posterior 0.3, sometimes

completely black dorsally and marked with red

only laterally. 3 tergite red. 4" tergite completely

black or narrowly marked with red anteriorly.

Pilosity whitish.

d: Flagellum with 29 segments, filiform, the

apical 0.8 attenuate, shortest segment 0.9x, penul-

timate segment 1.0x as long as wide. Hind femur

brown to dark brown. Hind tibia distinctly marked

with brown also basally. Genital claspers black. In

other characters similar to the ®.

Material (1°): Russia: Sverdlovsk Prov.: Nizhniy

Tagil (PET).

Dusona americana (Ashmead, 1890)

Dusona oyamadai Hinz, 1994 (syn. nov.)

The type of Casinaria americana Ashmead (WAS)

which is much damaged (head, greater parts of

the legs, one front wing and gaster lacking), has

recently been studied by the junior author. It agrees

well with European material of the species. D. oya-

madai was synonymized with D. americanaby Hinz

(unpublished notes; see introduction). The holo-

type of D. oyamadai is lost, but two paratypes

(12, 18) from Novoaleksandrovsk/Sakhalin / Rus-

sia (PET), which belong to D. americana, have re-

cently been studied by the junior author (see be-

low). D. americana is similar to D. annexa (see key,

couplet 210).

Description. 9: Body length 7-9 mm (rarely 6 mm;

see below). Temples distinctly constricted behind

the eyes, 0.7x as long as the width of an eye,

tangential lines on eyes and temples intersect on

the scutellum. Ocellar index 0.8. Frons scarcely

depressed, coriaceous, densely rugose-punctate,

with a fine median longitudinal carina, with short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal half of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, coriaceous

and with rather dense punctation ventrally, its

apex truncate medially. Malar space index 0.4.

Genal carina low and straight ventrally, joining

the scarcely raised oral carina at an acute angle.

Genal index 0.2. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 37-43 segments (rarely 36 segments;

see below), filiform, the apical 0.2 attenuate, 2"‘

segment 2.2x, shortest segment 1.1x, penultimate

segment 1.4x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.2, the

wrinkles smooth, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous,

densely rugose-punctate. Scutellum with lateral

carinae on the anterior 0.8. Prescutellar groove

with some longitudinal striae. Prepectal carina

low, ventral part merging with the pleural part,

the dorsal 0.2-0.3 of the latter obliterated or cov-

ered with wrinkles. Transverse carina obliterated

or present anteriorly. Mesopleurum densely punc-

tate (rarely dispersely or rather densely punctate;

see below) on a finely coriaceous and shining

background. Speculum granulate-strigose, some-

what shining. Depression in front of the speculum

with distinct longitudinal striae, which merge with

the punctation anteriorly. Metapleurum densely

rugose. Pleural carina complete. Areolet rather

large, stalked, receiving the 2” recurrent vein al-

most at the middle. Nervellus vertical or slightly

reclivous, weakly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 5.1x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.4 of the

hind coxae, slightly and rather narrowly de-

pressed, with short transverse wrinkled in the

shallow longitudinal furrow, in other parts irreg-

ularly wrinkled, carinae almost completely oblit-

erated. Propodeal spiracle rather small, 2.9x as

long as wide. Spiracular carina narrow. Spiracular

index 0.6. 1* gastral tergite 4.6x, 2" tergite 1.9x as

long as wide. Petiole without glymmae, with dis-

tinct narrow lateral areas, smooth in front of these

areas. Petiolar suture fine. 1* gastral sternite fine-

ly sculptured, partly smooth. Thyridium elongate.

Thyridial index about 2.0. Epipleurum not sepa-

rated from the 3"% gastral tergite, the latter with

distinct black lateral stripes well above the vent-

rolateral edge. Gaster distinctly compressed pos-

teriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible narrowly marked with yellow dorsally and

medially. Tegula yellowish brown. Wings hya-

line. Stigma dark brown medially. Front and mid

legs from the trochanter onwards yellowish red,

tibiae and tarsi yellow. Hind femur red or black

(apparently often for the greater part red in Euro-

pean material, but black in eastern Palearctic ma-

terial). Hind tibia yellowish brown medially, nar-

rowly black basally, broadly dark brown or black

apically. Hind tarsus yellowish brown to dark

brown. 2"d gastral tergite red on the posterior 0.2-

0.3. 3’ and 4" tergites red, the latter sometimes

marked with black dorsally or laterally. Pilosity

whitish.

d: Propodeum distinctly depressed, longitu-

dinal carinae present medially and posteriorly,

with long transverse wrinkles between these car-

inae. Mandible broadly marked with yellow me-

dially. Tegula yellow. Front coxa yellow apically.

In other characters similar to the ®.

Variability: Those ?2 from Sakhalin which have

recently been studied by the junior author (in-

cluding a paratype of D. oyamadai) are smaller

than characteristic D. americana (body length 6 mm,

36 flagellar segments), and the mesopleurum is

less densely punctate centrally (on a shining back-

ground, in that respect differing from D. annexa).

Probably this variation led Hinz (1994: 36) to de-

scribe a new species. dd collected at the same

localities do not differ from D. americana.

Material (7??, 958): Russia: Amur Prov.: Novorossi-

yka (PET); Sakhalin: Aniva (Hinz/ZSM, PET), Nova-

leksandrovsk; Kamchatka: Dalkiy, Kozyrevsk (PET).

Kazakhstan: Zailynskiy Alatau (MOS). China: Liao-

ning Prov.: Kaigen (= Kaiyuan) (SAP). North Korea:

Chongjin (SAW).

Dusona amurator Hinz, 1985

Description. 2: Body length 8-10 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.6. Frons scarcely depressed, coriaceous

and densely rugose, with a fine median longitudi-

nal carina, with distinct radial wrinkles, which

extend to half the distance to the median ocellus.

Antennal carina low and narrow. Face and dorsal

0.7 of the clypeus densely and rather coarsely

rugose. Clypeus flat, finely coriaceous and with

rather dense punctation ventrally, its apex almost

truncate medially. Malar space index 0.3-0.4. Ge-

nal carina low and straight ventrally, joining the

slightly raised oral carina at an acute angle. Genal

index 0.3-0.4. Temple finely coriaceous and with

fine disperse punctation medially. Flagellum with

38-41 segments, filiform, the apical 0.2 distinctly

attenuate, 2" segment 1.8x, shortest segment 0.9x,

penultimate segment 1.2x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.4, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous and densely

punctate. Scutellum with lateral carinae on the

anterior 0.6. Prescutellar groove with longitudinal

striae or almost smooth. Ventral part of the pre-

pectal carina merging with the pleural part or

with the transverse carina, in the latter case the

transverse carina slightly raised. Pleural part of

the prepectal carina low, its dorsal 0.3 obliterated.

Mesopleurum distinctly coriaceous, distinctly and

densely punctate. Speculum coriaceous or granu-

late-strigose. Depression in front of the speculum

with fine or distinct longitudinal striae, which

merge with a narrow area without punctation

anteriorly. Metapleurum finely and densely ru-

gose. Pleural carina distinct. Areolet rather large,

Bei

short-stalked, receiving the 2" recurrent vein

slightly distad of the middle. Nervellus slightly

inclivous, intercepted. Nervellar index 0.7-0.8. Tib-

ial spur index 0.6. Hind femur 4.5x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.4 of the

hind coxae, broadly and deeply depressed, sulcus

with short transverse wrinkles in the midline, with

long transverse wrinkles posteriorly, in other parts

irregularly wrinkled, costulae partly present, lon-

gitudinal carinae present only posteriorly. Propo-

deal spiracle rather small, 2.2x as long as wide.

Spiracular carina narrow or slightly widened.

Spiracular index 0.3-0.5. 1% gastral tergite 4.0x, 2"<

tergite 1.9x as long as wide. Petiole with very

small glymmae at the anterior edge of narrow

lateral areas, smooth in front of the glymmae.

Petiolar suture distinct. 1“ gastral sternite smooth

or with fine sculpture anteriorly, with distinct

transverse striae on the posterior half. Thyridium

elongate. Thyridial index about 1.3. Epipleurum

not separated from the 3'“ gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with reddish brown

dorsally and medially. Tegula dark brown or black.

Wings hyaline. Stigma brown medially. Front leg

from the femur onwards yellowish red, the femur

sometimes marked with brown. Mid femur yel-

lowish red on the apical 0.1-0.2. Mid tibia yellow-

ish red. Mid tarsus brown, marked with yellow

basally. Hind tibia whitish yellow medially (some-

times only on the dorsal side), narrowly black

basally, broadly black apically. 2" gastral tergite

red on the posterior 0.1-0.2. 3" tergite completely

red or narrowly marked with black posteriorly.

4" tergite completely black or narrowly marked

with red anteriorly. Pilosity whitish.

d: Longitudinal carinae of propodeum some-

times present medially. Gliymmae sometimes ob-

literated. 3" gastral tergite sometimes marked with

red only laterally. In other characters similar to

the 2.

Material (4??, 255): Russia: Buryatiya: Khamar-Da-

ban Mts. (MOS); Amur Prov.: Novorossiyka; Yevreys-

kaya A.O.: Amurzet (PET); Kuril Isl.: Dubovoye on

Kunashir (Hinz/ZSM), Shikotan (PET).

Variety: Body length 6 mm. Flagellum with 36

segments. Ventral part of the prepectal carina

merging with the transverse carina (as in some

specimens of D. amurator). Material: 1? Russia,

Irkutsk Prov., Dachnaya (PET). This ? may be a

small specimen of D. amurator.

Dusona anceps (Holmgren, 1860)

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.9. Frons slightly depressed, coriaceous,

finely and densely punctate, with a fine median

longitudinal carina, with distinct radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina distinctly raised, the rim

distinctly bent upwards (ear-like), with fine trans-

verse striae on the crescent-shaped surface. Face

and dorsal 0.4 ofthe clypeus coriaceous and dense-

ly rugose. Clypeus slightly concave, coriaceous

and with rather dense punctation ventrally, its

apex rather thick and truncate medially. Malar

space index 0.4. Genal carina low and straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.2-0.3. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 41-46 segments, filiform,

the apical 0.3 attenuate, 2”! segment 1.8x, shortest

segment 0.9x, penultimate segment 1.4x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.6, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum finely co-

riaceous and densely punctate. Scutellum with

lateral carinae on the anterior 0.8. Prescutellar

groove smooth. Prepectal carina low, ventral part

merging with the pleural part, the dorsal 0.3 of the

latter obliterated. Transverse carina obliterated.

Mesopleurum finely coriaceous, distinctly and

densely punctate. Speculum punctate posteriorly,

very finely coriaceous and shining anteriorly. De-

pression in front of the speculum with distinct

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum finely coriaceous,

distincetly and densely punctate. Pleural carina

distinct. Areolet rather large, short-stalked, re-

ceiving the 2” recurrent vein at or slightly basad

of the middle. Nervellus vertical, weakly inter-

cepted. Nervellar index 0.7. Tibial spur index 0.5.

Hind femur 4.5x as long as high. Hind tarsal claws

rather long, with distinct pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

rather broadly and not deeply depressed, sulcus

with long transverse wrinkles, costulae partly

present, longitudinal carinae almost completely

obliterated. Propodeal spiracle rather small, 4.3x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.6. 1* gastral tergite 4.0x, 2"4 tergite

1.6x as long as wide. Petiole with small glymmae

atthe anterior edge of narrow lateral areas, smooth

in front of the glymmae. Petiolar suture fine, com-

plete. 1% gastral sternite smooth. Thyridium oval.

Thyridial index about 1.0. Epipleurum not sepa-

rated from the 3"! gastral tergite, the latter with

distinct black lateral stripes well above the ventro-

lateral edge. Gaster distinctly compressed poste-

riorly. Ovipositor index 0.6.

Colour: Black. Mandible narrowly tinged with

reddish brown dorsally. Wings hyaline. Stigma

yellowish brown medially. Apical half of the front

femur and front tibia and tarsus yellow, the tarsus

marked with brown apically. Apical 0.2 of the mid

femur and basal 0.1 of the mid tibia yellowish red.

Remaining part of the mid tibia dark brown. 2“

gastral tergite red on the posterior 0.7. 3" and 4"

tergites red. Pilosity whitish.

d: Flagellum with 44-49 segments. Transverse

carina complete. Maxillary palpus yellow. Mandi-

ble broadly marked with yellow medially. Tegula

yellow. Front and mid legs yellow, the coxa black

basally. Hind tibia yellow medially, narrowly black

basally, broadly black apically. In other characters

similar to the ®.

Material (15?2, 1036): Russia: Krasnoyarsk Terr.:

Maklakovo, River Yenisey; Primorye: Kamen Rybolov

(PET), Lazo (MOS), Novokatsalinsk, Posyet, Yakovlev-

ka, Vladivostok; Kuril Isl.: Kunashir (PET). Turkmeni-

ya: Ashadag (MOS). Kazakhstan: Kustanay. Mongoli-

ya: Selenge (PET). Japan: Hokkaido: Rebun-dake

(SAP).

Dusona angustifrons (Förster, 1868)

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.9. Frons not depressed, coriaceous, densely ru-

gose-punctate, with a fine median longitudinal

carina, with rather long radial wrinkles, which

extend to half the distance to the median ocellus.

Antennal carina low, but the rim more or less bent

upwards, sometimes widened to a smooth nar-

row crescent-shaped plate (Fig. 4). Face and dor-

sal 0.4 of the clypeus coriaceous, very densely

rugose-punctate. Clypeus almost flat, coriaceous

and with rather dense punctation ventrally, its

apex truncate medially. Malar space index 0.5.

Genal carina low, bent forwards and sidewards,

joining the oral carina at the base of the mandible

at a right angle (Fig. 10 and 15). Temple finely

coriaceous and with very fine disperse punctation

medialiy. Flagellum with 33-37 segments, slightly

fusiform, 2" segment 2.5x, shortest segment 0.9x,

penultimate segment 1.1x as long as wide. Prono-

tum coriaceous and densely rugose-punctate lat-

erally on the dorsal 0.2, with longitudinal wrin-

kles ventrally. Mesoscutum coriaceous, with fine-

ly and densely punctate. Scutellum coriaceous,

rather densely punctate, with lateral carinae on

the anterior 0.9. Prescutellar groove for the great-

er part smooth, with few longitudinal striae later-

ally. Prepectal carina low, ventral part merging

with the pleural part, the latter complete, joining

the anterior edge of the mesopleurum. Transverse

carina complete, low posteriorly, somewhat raised

anteriorly. Mesopleurum distinctly coriaceous and

densely punctate. Speculum coriaceous, for the

greater part rather dull, a small ventral part al-

most smooth and shining. Depression in front of

the speculum with distinct longitudinal striae,

which merge with the punctation anteriorly. Meta-

pleurum densely rugose. Pleural carina distinct.

Areolet rather large, short-stalked, receiving the

2"d recurrent vein at or slightly distad of the mid-

dle. Nervellus vertical, not intercepted (Fig. 27) or

weakly intercepted. In the latter case nervellar

index 0.8. Tibial spur index 0.6. Hind femur 5.1x

as long as high. Hind tarsal claws short, with short

pectinate teeth basally. Propodeum extending to

0.4 of the hind coxae, rather broadly and distinctly

but not deeply depressed, sulcus with short trans-

verse wrinkles anteriorly and medially in the mid-

line, with long transverse wrinkles posteriorly,

irregularly wrinkled in other parts, costulae com-

plete, longitudinal carinae almost completely ob-

literated. Propodeal spiracle small, 2.7x as long as

wide. Spiracular carina often widened. Spiracular

index 0.5. 1“ gastral tergite 4.0x, 2" tergite 1.9x as

long as wide. Petiole with small glymmae at the

anterior edge of narrow lateral areas (Fig. 36),

glymmae sometimes almost obliterated, petiole

smooth in front of the glymmae. Petiolar suture

almost completely obliterated. 1* gastral sternite

smooth. Thyridium oval. Thyridial index about

1.5. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge (Fig. 42). Gaster

distinctly compressed posteriorly. Ovipositor in-

dSS0)5:

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown apically. Mandible nar-

rowly marked with yellowish brown dorsally.

Tegula dark brown or black. Wings hyaline. Stig-

ma light brown medially. Front leg from the fe-

mur onwards yellowish red. Mid femur yellowish

brown on the apical 0.2. Mid tibia yellowish brown.

Mid tarsus dark brown. Hind tibia yellowish

brown to dark brown medially, narrowly marked

with dark brown basally, broadly marked with

dark brown apically (apparently rather dark and

sometimes completely black in eastern Palearctic

material). Hind tarsus sometimes marked with

yellowish brown basally. 2" gastral tergite red on

the posterior 0.1. 3° and 4" tergites red, the 4"

tergite sometimes marked with black posteriorly

or completely black. Pilosity whitish.

d: Flagellum with 36-38 segments. Petiolar

suture complete. Mandible broadly yellow medi-

ally. Tegula yellowish brown. Front and mid legs

from the tip of the coxa onwards yellowish red. In

other characters similar to the ®.

Material (31?2?, 1188): Russia: Irkutsk Prov.: Dach-

naya; Chita Prov.: Adrianovka, Chita, Klichka, Ner-

chinskiy Zavod; Amur Prov.: Novorossiyka; Yevrey-

skiy A.O.: Londoko; Primorye: Barabash-Levada,

Khualaza, Ussuriysk (PET); Sakhalin: Novikovo (KIS),

Novoaleksandrovsk, Ozerskiy, Tymovskoye; Kuril

Isl.: Tretyakovo on Kunashir (PET). Kazakhstan: Zai-

lynskiy Alatau Mts. (MOS). Mongoliya: Hentiy Prov.

(PET). Japan: Hokkaido: Sapporo (AEI); Kyushu:

Hiko-san (SAP).

Dusona annexa (Förster, 1868)

Campoplex limnobius Thomson, 1887 (syn. nov.)

Bachmaier (1979: 73) selected as lectotype for Cam-

poplex annexus Förster a specimen which had been

determined as D. limnobia (Thomson) by Hinz

(1962: 3), but did not formally synonymize both

taxa. The synonymy was also overlooked by Yu &

Horstmann (1997: 143 and 149).

Description. ?: Body length 8-9 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.8. Frons scarcely de-

pressed, coriaceous, finey and densely rugose-

punctate, with a fine median longitudinal carina,

with few radial wrinkles, which extend to half the

distance to the median ocellus. Antennal carina

low or slightly raised, the rim sometimes slightly

bent upwards. Face and dorsal half of the clypeus

coriaceous and densely rugose. Clypeus flat, co-

riaceous and dispersely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina low ventrally, bent forwards and sidewards,

joining the slightly raised oral carina at a right

angle. Genal index 0.1-0.2. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 42-47 segments, filiform,

the apical 0.2 attenuate, 2” segment 2.4x, shortest

segment 1.1x, penultimate segment 1.4x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous

and densely punctate. Scutellum with lateral car-

inae on the anterior 0.8. Prescutellar groove al-

most smooth, with few longitudinal striae lateral-

ly. Prepectal carina low, ventral part merging with

the pleural part or with the transverse carina.

Pleural part almost complete or its dorsal 0.1-0.4

obliterated or covered with wrinkles. Transverse

carina complete, low posteriorly, somewhat raised

anteriorly. Mesopleurum distinctly coriaceous and

with disperse or rather dense punctation. Specu-

lum coriaceous and finely granulate-strigose. De-

pression in front of the speculum with distinct

longitudinal wrinkles and striae, which merge with

the punctation anteriorly. Metapleurum rugose

or rugose-punctate. Pleural carina distinct. Areo-

let rather large, stalked, receiving the 2" recurrent

vein distad of the middle. Nervellus reclivous,

weakly intercepted. Nervellar index 0.7. Tibial

spur index 0.6. Hind femur 5.0x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to the middle of

the hind coxae, rather broadly and shallowly de-

pressed, sulcus for the greater part irregularly

wrinkled, usually with transverse wrinkles poste-

riorly, rarely with some longitudinal wrinkles,

costulae partly present, longitudinal carinae for

the greater part obliterated. Propodeal spiracle

rather small, 2.4x as long as wide. Spiracular car-

ina narrow. Spiracular index 0.8. 1 gastral tergite

4.3x, 2" tergite 2.2x as long as wide. Petiole usu-

ally without glymmae, with distinct narrow later-

al areas, sometimes with very small glymmae at

the anterior edge of these areas, petiole smooth in

front of these areas. Petiolar suture fine, partly

obliterated. 1* gastral sternite for the greater part

smooth, with few transverse striae posteriorly.

Thyridium elongate. Thyridial index about 1.0.

Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow

dorsally and medially. Tegula brownish black.

Wings hyaline. Stigma light brown medially. Front

leg from the trochanter or the femur onwards

yellowish red. Mid femur yellowish red on the

apical 0.3. Mid tibia and tarsus yellowish red.

Hind femur red or black. Hind tibia dark yellow-

ish brown medially, narrowly black basally, broad-

Iy black apically. 24 gastral tergite red on the

posterior 0.2-0.3. 3”! and 4" tergites red. 5" tergite

sometimes narrowly marked with red anteriorly.

Pilosity whitish.

d: Mesopleurum with rather dense or dense

punctation. Propodeum sometimes with longitu-

dinal carinae medially and posteriorly and with

distinct transverse wrinkles between these cari-

nae. Maxillary palpus yellow. Mandible broadly

yellow medially. Tegula brown. Tip of front coxa

and front trochanter yellow. 3" gastral tergite

sometimes marked with red posteriorly. 4" tergite

sometimes partly or completely black. In other

characters similar to the 2.

Material (6??, 1738): Russia: Buryatiya; Chita Prov.:

Ivan Lake, Kalga, Nerchinskiy Zavod; Yevreyskiy

A.O.: Amurzet; Khabarovsk Terr.: Khekhtsir near

Khabarovsk; Primorye: Pozhiga SO of Dalnerechensk

(PET), Sikhote-Alin (MOS), Spassk Dalniy (MOS,

PET); Sakhalin (SAP); Kamchatka: Milkovo (PET). Ka-

zakhstan: Alma-Ata (MOS, PET). Mongoliya: Selenge

Prov. (PET). Japan: Hokkaido: Sapporo (AEI).

Dusona auriculator Aubert, 1964

The type of this species (Aubert/LAU) has recent-

ly been studied by the junior author.

Description. 2: Body length 8mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

finely coriaceous, densely punctate, with a fine

median longitudinal carina, with indistinct radial

wrinkles, sometimes smooth ventrally. Antennal

carina in most specimens slightly but distinctly

raised, the rim widened to a smooth bulge, in few

specimens rather low and narrow. Face and dor-

sal 0.7 of the clypeus coriaceous and densely ru-

gose. Clypeus almost flat, coriaceous and with

rather dense punctation ventrally, its apex slight-

ly convex medially. Malar space index 0.3. Genal

carina slightly raised ventrally, almost straight or

slightly bent forwards and sidewards, joining the

slightly raised oral carina at an acute angle. Genal

index 0.2. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

43-46 segments, filiform, the apical 0.2 attenuate,

2"d segment 2.7x, shortest segment 1.3x, penulti-

mate segment 1.5x as long as wide. Pronotum

very densely punctate on a smooth background

laterally on the dorsal 0.1-0.2, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous and densely punctate. Scutellum part-

ly punctate on a shining background, with lateral

carinae on the anterior half. Prescutellar groove

with longitudinal striae. Prepectal carina low ven-

trally, somewhat raised ventrolaterally, merging

with the raised transverse carina. Pleural part of

the prepectal carina low, its dorsal 0.2 obliterated.

Mesopleurum with rather dense punctation on a

very finely coriaceous and shining background.

Speculum very finely coriaceous and granulate-

strigose. Depression in front of the speculum with

distinct long longitudinal striae, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, finely and densely rugose-punctate. Pleu-

ral carina complete. Areolet rather small, long-

stalked, receiving the 2” recurrent vein distad of

the middle. Nervellus distinctly reclivous, weakly

intercepted. Nervellar index 0.7. Tibial spur index

0.4. Hind femur 4.9x as long as high. Hind tarsal

claws short, with short pectinate teeth basally.

Propodeum extending to 0.7 of the hind coxae,

distinctly but narrowly and not deeply depressed,

with short transverse wrinkles in the longitudinal

furrow, in other parts irregularly wrinkled, costu-

lae fine but complete, longitudinal carinae almost

completely obliterated. Propodeal spiracle rather

small, 3.2x as long as wide. Spiracular carina nar-

row. Spiracular index 0.5. 1” gastral tergite 4.9x,

2"! tergite 2.9x as long as wide. Petiole smooth

laterally, without glymmae or lateral areas (Fig.

38). Petiolar suture fine, partly obliterated. 1“ gas-

tral sternite smooth. Thyridium elongate. Thyrid-

ial index about 1.5. Epipleurum not separated

from the 3"4 gastral tergite, the latter with or with-

out black lateral stripes. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellowish red medially. Tegula yel-

low. Wings hyaline. Stigma dark brown medially.

Front and mid legs from the tip of the coxa on-

wards yellow. Hind trochantellus and femur light

red. Hind tibia yellowish brown medially, nar-

rowly marked with dark brown basally, rather

broadly marked with dark brown apically. Hind

tarsus sometimes for the greater part yellowish

brown basally. 2"‘ gastral tergite dorsally red on

the posterior 0.1, the red marks more extended

laterally. 3 and 4" tergites light red, sometimes

narrowly marked with black dorsally. 5" to 7"

tergites broadly black dorsally, marked with red

laterally, or for the greater part or completely red.

Gastral sternites red. Pilosity whitish.

d: Propodeum extending to 0.5-0.6 of the hind

coxae. Petiole sometimes with very small and shal-

low glymmae at the anterior edge of indistinct

narrow lateral areas. Genital clasper usually red,

rarely black. In other characters similar to the 2.

Material (4229, 1588): Russia: Khabarovsk Terr.:

Khekhtsir near Khabarovsk; Primorye: Khasan, Pozhi-

ga SO of Dalnerechensk (PET). Japan: Hokkaido:

Sapporo (AEI); Honshu: Fukuoka, Nagano (OTT),

Saitama (OTT, SAP), Sasayama (SAP); Kyushu: Hiko-

san (Hinz/ZSM).

Dusona aurita (Kriechbaumer, 1883)

Description. 2: Body length 10-13 mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.8. Frons coriaceous and

densely rugose-punctate dorsally, strongly de-

pressed with long radial wrinkles ventrally, the

latter extending to 0.8 the distance to the median

ocellus, with a distinct median longitudinal carina

on the top a slightly raised median longitudinal

bulge. Antennal carina strongly raised and wid-

ened to a nose-like dorsal projection, the latter

with fine transverse striae. Face and the greater

part of the clypeus coriaceous, densely rugose

and rugose-punctate. Clypeus flat, a small medi-

an area coriaceous and rather densely punctate

ventrally, its apex truncate medially. Malar space

index 0.3. Genal carina straight ventrally, distinct-

ly raised close to the junction with the also raised

oral carina, forming a lopsided triangular pyra-

mid, the tip of the latter shifted ventrally and

situated behind the base of the mandible (Fig. 11).

Gena depressed close to the junction, the depres-

sion about 0.7% as deep as the width of the 2"

flagellar segment. Genal index 0,4. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 52-54 segments, filiform,

the apical 0.3 distinctly attenuate, 2” segment

1.5x, shortest segment 0.9x, penultimate segment

1.3x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.4, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous and very densely punctate. Scutel-

lum with lateral carinae on the anterior 0.7.

Prescutellar groove smooth medially, with fine

longitudinal striae laterally. Prepectal carina low,

ventral part merging with the pleural part, the

latter complete, joining the anterior edge of the

mesopleurum. Transverse carina present. Meso-

pleurum strongly and densely punctate on a fine-

ly coriaceous and shining background. Speculum

finely granulate-strigose, shining. Depression in

front of the speculum with distinct rather long

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum densely rugose or

rugose-punctate. Pleural carina strong. Areolet

large, stalked, receiving the 2” recurrent vein

slightly distad of the middle. Nervellus inclivous,

strongly intercepted. Nervellar index 0.7-0.8. Tib-

ial spur index 0.6. Hind femur 5.2x as long as

high. Hind tarsal claws rather short, with distinct

pectinate teeth basally. Propodeum extending to

the middle of the hind coxae, broadly and deeply

depressed, sulcus with long transverse wrinkles,

sometimes with a short median longitudinal keel

posteriorly, carinae almost completely obliterat-

ed. Propodeal spiracle large, 3.4x as long as wide.

Spiracular carina narrow. Spiracular index 0.3.

1* gastral tergite 4.9x, 2" tergite 2.0x as long as

wide. Petiole with small but deep glymmae at the

anterior edge of narrow lateral areas, with very

fine longitudinal striae in front of the glymmae.

Petiolar suture almost completely obliterated.

1“ gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster strongly compressed posteriorly. Ovi-

positor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow

dorsally and medially. Tegula yellowish brown to

dark brown. Wings hyaline. Stigma yellowish

brown medially. Front trochanter brown. Front

femur, tibia and tarsus yellowish red. Mid femur

yellowish brown on the apical 0.2. Mid tibia yel-

lowish. Mid tarsus dark brown. Hind tibia yel-

lowish brown medially, narrowly marked with

dark brown basally, broadly marked with dark

brown apically. 2" gastral tergite red on the pos-

terior 0.3. 3! and 4" tergites red, sometimes tinged

with black. Pilosity whitish.

d: Propodeum less strongly depressed, longi-

tudinal carinae almost complete. Mandible broad-

ly yellow medially. Tegula yellowish brown. Front

leg from the tip of the coxa onwards yellowish

red. Mid tibia yellow. Hind tibia yellow medially.

Genital clasper black. In other characters similar

to the 2.

Host: Hypomecis punctinalis (Scopoli, 1763) (Ge-

ometridae) (MIN).

Material (1422, 788): Russia: Novosibirsk Prov.:

Akademgorodok (PET); Altaysk Terr.: Gorno-Al-

taysk, Volchikha (MIN); Krasnoyarsk Terr.: Shushens-

koye (MOS); Khabarovsk Terr.: Khabarovsk; Primor-

ye: Khasan (PET), Vladivostok (KIS, PET). Mongoliya:

Dornod Prov.: Choybalsan (PET). Japan: Hokkaido:

Sapporo (SAP).

Dusona bellator Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarovsk, 1.7.1983 (PET). -

Paratypes: 1? same locality, 12.6.1983 (Hinz /

ZSM); 13 same locality, 13.6.1985 (PET); 1? Rus-

sia, Primorye, Vladivostok, 1.6.1962 (MOS); 233

Japan, Honshu, Nagano, 400-700 m, 8.-20.5.1963

(OTT); 18 Japan, Honshu, Nagano, Kinasa, 600-

1000 m, 13.5.1962 (OTT); 15 Japan, Honshu, Na-

gano, Yamada, 1200 m, 11.5.1962 (OTT); 13 Japan,

Honshu, Kamitakai, 800 m, 5.5.1962 (Hinz/ZSM).

Description. ?: Body length 7-8 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons scarcely depressed, coriaceous, dull,

finely rugose-punctate, with a very fine median

longitudinal carina, with distinct radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina low and narrow. Face

and dorsal half of the clypeus coriaceous and dull,

densely rugose-punctate. Clypeus flat, coriaceous

and dull, finely and rather densely punctate ven-

trally, its apex smooth, slightly convex medially.

Malar space index 0.4. Genal carina low, almost

straight ventrally, joining the low oral carina at an

acute angle. Genal index 0.6. Temple coriaceous

and with fine disperse punctation medially. Flag-

ellım with 37-41 segments, slightly fusiform, 2“

segment 1.8x, shortest segment 0.9x, penultimate

segment 0.9-1.0x as long as wide. Pronotum ru-

gose-punctate laterally on the dorsal 0.2, with

dense punctation and longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous and

rugose-punctate. Scutellum with lateral carinae

on the anterior 0.4. Prescutellar groove smooth.

Prepectal carina low, its ventral part merging with

the pleural part, the latter almost joining the ante-

rior edge of the mesopleurum. Transverse carina

obliterated. Mesopleurum distinctly coriaceous,

with rather dense or dense punctation. Speculum

finely coriaceous or almost smooth. Depression in

front of the speculum with fine longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum coriaceous, densely rugose or ru-

gose-punctate. Pleural carina strong. Areolet rath-

er small, short-stalked, receiving the 2" recurrent

vein distad of the middle. Nervellus distinctly

inclivous, intercepted. Nervellar index 0.7-0.8. Tib-

ial spur index 0.7. Hind femur 5.1x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to the basal 0.3 of the hind coxae, slightly

and broadly depressed, almost without carinae,

for the greater part coriaceous and with fine irreg-

ular wrinkles, with some transverse and longitu-

dinal wrinkles posteriorly in the shallow sulcus.

Propodeal spiracle small, 2.5x as long as wide,

Spiracular carina distinct, sometimes widened.

Spiracular index 0.5-0.7. 1“ gastral tergite 3.0x, 2“

tergite 1.0x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal

carinae dorsally and ventrally, smooth in front of

the glymmae. Petiolar suture indicated. 1“ gastral

sternite for the greater part smooth, with fine

transverse striae posteriorly. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum partly separat-

ed from the 3" gastral tergite, black lateral stripe

ina short distance to the horizontal crease. Epipleu-

ral index 0.4. Gaster distinctly but not strongly

compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus, mandible me-

dially (broadly) and tegula yellow. Wings hya-

line. Stigma medium brown medially. Front leg

from the trochanter onwards yellow, the femur

yellowish red. Mid leg from the apical 0.3 of the

femur onwards yellow. Hind tibia light yellow

medially, narrowly black basally, broadly black

apically. Hind tarsus yellowish brown to brown.

2X gastral tergite dorsally red on the posterior 0.1,

the red marks more extended laterally. 3" tergite

for the greater part red, marked with black dorsal-

ly. 4" tergite marked with red on the ventrolateral

half. 5'" tergite sometimes with narrow red marks

ventrolaterally. Pilosity whitish.

d: Front and mid coxae with yellow tips. Mid

trochanter yellow. Mid femur yellowish red. Lon-

gitudinal carinae of the propodeum indicated.

Epipleural index 0.6. In other characters similar to

the 2.

Dusona bellipes (Holmgren, 1872)

Description. 2: Body length 11-13 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons slightly depressed,

coriaceous, finely and dispersely punctate, a fine

median longitudinal carina indicated only medi-

ally, without radial wrinkles, with a small raised

point on either side between the antennal socket

and the eye. Antennal carina low and narrow

(Fig. 3). Face and dorsal 0.6 of the clypeus coria-

ceous and densely rugose-punctate. Clypeus flat,

coriaceous and densely punctate ventrally, its apex

truncate medially. Malar space index 0.3. Genal

carina low and almost straight ventrally, joining

the low oral carina at an acute angle. Genal index

0.6. Temple coriaceous and with very fine dis-

perse punctation medially. Flagellum with 54-55

segments, filiform, the apical 0.3 attenuate, 2"

segment 1.8x, shortest segment 0.9x, penultimate

segment 1.2x as long as wide. Pronotum coria-

ceous and densely rugose-punctate laterally on

the dorsal 0.3, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous and very

densely punctate. Scutellum with lateral carinae

on the anterior 0.3. Prescutellar groove smooth

medially, with longitudinal striae laterally. Pre-

pectal carina low, the ventral part merging with

the pleural part, the latter complete, joining with

the anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum finely coriaceous,

with rather dense or dense punctation. Speculum

very finely coriaceous, shining. Depression in front

of the speculum dorsally with short longitudinal

striae, which merge with a coriaceous area with-

out punctation anteriorly, granulate-strigose and

shining on the ventral 0.3. Metapleurum densely

and rather coarsely rugose. Pleural carina distinct.

Areolet large, narrowly sessile, receiving the 2”

recurrent vein at the middle. Nervellus slightly

inclivous, distinctly intercepted, its upper branch

bent inwards. Nervellar index 0.7. Tibial spur in-

dex 0.6. Hind femur 6.5x as long as high. Hind

tarsal claws short, with distinct pectinate teeth

basally. Propodeum extending to the middle of

the hind coxae, broadly and not deeply depressed,

sulcus with long transverse wrinkles, costulae and

longitudinal carinae almost complete. Propodeal

spiracle small, 2.7x as long as wide. Spiracular

carina narrow. Spiracular index 0.8. 1‘ gastral

tergite 5.1x, 2" tergite 2.4x as long as wide. Peti-

ole with small but distinct glymmae at the anteri-

or edge of narrow lateral areas, partly with very

fine longitudinal striae in front of the glymmae.

Petiolar suture fine, partly obliterated. 1“ gastral

sternite for the greater part smooth, with fine

transverse striae posteriorly. Thyridium oval. Thy-

ridial index about 1.5. Epipleurum separated from

the 3" gastral tergite, the crease marked with black.

Epipleural index 0.6-0.7. Gaster strongly com-

pressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown medially and apically.

Mandible narrowly marked with yellow dorsally

and medially. Tegula yellowish brown to dark

brown. Wings hyaline. Stigma medium brown

medially. Front femur yellowish red dorsally, black

ventrally. Front tibia and tarsus yellow. Mid fe-

mur yellowish red on the apical 0.3. Mid tibia and

tarsus yellowish red. Hind femur yellowish brown

on the apical 0.2. Hind tibia yellowish brown,

without dark marks. 2"! gastral tergite red on the

posterior half. 3° and 4" tergites red. 5" tergite

marked with red ventrolaterally. Pilosity whitish.

d: Mandible rather broadly yellow medially.

Tegula yellow. Front coxa and trochanter marked

with yellow. Front and mid femur yellowish dor-

sally, brownish ventrally. Front and mid tibia and

tarsus yellow. 5% gastral tergite completely red. In

other characters similar to the ?.

Material (1?): Japan: Honshu: Sasayama (SAP).

Dusona belokobyli Hinz & Horstmann,

spec. nov.

Holotype (2): Russia, Primorye, Chuguyevka,

3:0.11985. (PET):

Description. 9: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.6x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.6. Frons slightly depressed, coriaceous,

densely rugose-punctate, without a median longi-

tudinal carina, with short radial wrinkles. Anten-

nal carina slightly raised and its rim slightly bent

upwards (inconspicuous). Face and dorsal 0.6 of

the clypeus distinctiy and densely rugose-punc-

tate. Clypeus flat, finely coriaceous and rather

densely punctate ventrally, its apex slightly con-

cave medially. Malar space index 0.3. Genal cari-

na low, almost straight ventrally, joining the low

oral carina at an acute angle. Genal index 0.1.

Temple coriaceous and with distinct rather dense

or dense punctation medially. Flagellum with 44

segments, filiform, the apical 0.2 attenuate, 2

segment 2.1x, shortest segment 0.9x, penultimate

segment 1.0x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.3, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, with very dense puncta-

tion. Scutellum with lateral carinae on the anterior

0.3. Prescutellar groove with distinct longitudinal

striae. Ventral part of the prepectal carina merg-

ing with the transverse carina, both slightly raised,

the former not emarginate medially. Pleural part

of the prepectal carina indicated medially, obliter-

ated dorsally and ventrally. Mesopleurum finely

coriaceous and with dense punctation. Speculum

finely coriaceous or almost smooth, shining. De-

pression in front of the speculum with distinct

short longitudinal striae, which merge with the

punctation anteriorly. Metapleurum densely ru-

gose-punctate. Pleural carina strong. Areolet rath-

er small, short-stalked, receiving the 2" recurrent

vein at the middle. Nervellus slightly reclivous,

slightly intercepted. Nervellar index 0.8. Tibial

index 0.6. Hind femur 4.9x as long as high. Hind

claws short, with short pectinate teeth basally and

medially. Propodeum extending to the middle of

the hind coxae, shallowly and rather narrowly

depressed, transversely wrinkled in the longitu-

dinal furrow, in other parts irregularly wrinkled,

costulae complete, longitudinal carinae almost

completely obliterated. Propodeal spiracle 3.4x as

long as wide. Spiracular carina distinct. Spiracular

index 0.7. 1“ gastral tergite 4.4x, 2" tergite 2.3x as

long as wide. Petiole without glymmae, with in-

distinct narrow lateral areas, these areas coria-

ceous, petiole smooth in front of the areas. Petio-

lar suture almost completely obliterated. 1“ gas-

tral sternite for the greater part smooth, with fine

transverse striae posteriorly. Thryridium oval.

Thyridial index 1.8. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellow. Man-

dible narrowly reddish brown dorsally and medi-

ally. Tegula dark brown. Wings hyaline. Stigma

dark brown medially. Front femur reddish brown

dorsally, dark brown ventrally. Front tibia and

tarsus yellow. Mid femur yellow on the apical 0.1.

Mid tibia yellow dorsally, medium brown ven-

trally. Mid tarsus medium brown, basitarsus yel-

lowish. Hind tibia marked with medium brown

basally on the dorsal side, in other parts dark

brown to black. 2" gastral tergite dorsally red on

the posterior 0.1, the red marks slightly more ex-

tended laterally. 3° and 4" tergites red, the 3'*

narrowly marked with medium brown dorsally.

Pilosity whitish, pilosity of the face tinged with

light brown.

d unknown.

Dusona bicoloripes (Ashmead, 1906)

Campoplex foersteri Roman, 1942 (syn. nov.)

The type (8) of Campoplex bicoloripes Ashmead

from Hakone near Tokyo/Honshu/Japan (WAS)

has recently been studied by the junior author; it

had been misinterpreted by Hinz (1979: 216). Lec-

totype (?) of Campoplex foersteri Roman here des-

ignated: “10/6 38.” (from Lousberg/ Aachen/D;

after Förster 1868: 818), “Pugillator Foerst.”, coll.

Förster/ZSM. The specimens in coll. Förster are

syntypes of this taxon, because Roman (1942: 18)

referred to Campoplex pugillator sensu Förster nec

Linnaeus. The name given by Roman is not a

nomen novum in the definition of the Code (arti-

cle 60), but a new description validated by an

indication (article 31.1.2 of the Code), because

Förster (1868: 816) did not describe a new species,

but referred to Gravenhorst (1829: 606 ff.) as the

author, and the latter referred to Linnaeus. The

name Campoplex foersteri Teunissen is considered

here not as a separate taxon, but as a citation of

Campoplex foersteri Roman, because Teunissen

(1947: 266) expressly referred to Roman as the

author of the name. This was overlooked by Yu &

Horstmann (1997: 146).

D. bicoloripes is a variable and poorly defined

species. It differs from D. chabarowski (= D. bicolo-

ripes sensu Hinz 1979: 216) in the sculpture of the

propodeum (see key, couplet 26). Specimens

without raised points on the frons are similar to

D. stragifex and also differ from the latter mainly

in the sculpture of the propodeum. As far as

known, D. bicoloripes is thelytokous in Europa

(and probably in central Asia), but arrhenotokous

in Japan. Populations in eastern Russia (Khaba-

rovsk Terr., Primorye) may represent an interme-

diate position (19 from Khekhtsir among a long

series of ?%)

Description. 2: Body length 9-12 mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyesand

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons slightly depressed,

coriaceous, finely and rather densely punctate,

with a fine median longitudinal carina, with few

short radial wrinkles (which may be lacking), usu-

ally on either side with a distinct or indistinct

raised point between the antennal socket and the

eye, which may be completely obliterated. Anten-

nal carina low and narrow. Face and dorsal 0.7 of

the clypeus coriaceous, densely rugose-punctate.

Clypeus flat, coriaceous and with rather dense

punctation ventrally, its apex truncate medially.

Malar space index 0.3. Genal carina low and rath-

er straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with very fine dis-

perse punctation medially. Flagellum with 45-55

segments, filiform, the apical 0.7 attenuate, 2"

segment 1.5x, shortest segment 0.9x, penultimate

segment 1.1x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, very densely punctate.

Scutellum with lateral carinae on the anterior 0.6.

Prescutellar groove smooth. Prepectal carina low,

ventral part merging with the pleural part, the

letter complete or indistinct and covered with wrin-

kles dorsally. Transverse carina present. Meso-

pleurum distinctly coriaceous and densely punc-

tate. Speculum punctate posteriorly, finely coria-

ceous or granulate-strigose and shining anterior-

ly, sometimes smooth ventrally. Depression in

front of the speculum dorsally with rather short

longitudinal striae or wrinkles, which merge with

the punctation anteriorly, ventrally often granu-

late-strigose or with fine striae, which may merge

with a narrow area without punctation anteriorly.

Metapleurum coriaceous, finely and densely ru-

gose-punctate. Pleural carina distinct. Areolet

large, stalked, receiving the 2" recurrent vein at

or basad of the middle. Nervellus verticai or slight-

ly reclivous, distinctly intercepted. Nervellar in-

dex 0.7. Tibial spur index 0.6. Hind femur 5.1x as

long as high. Hind tarsal claws short, with short

pectinate teeth basally. Propodeum extending to

0.3 of the hind coxae, scarcely or not at all de-

pressed, irregularly wrinkled, often with a long

median keel posteriorly, sometimes with some

longitudinal wrinkles, rarely with short transverse

wrinkles beside that keel, costulae shortened or

absent, longitudinal carinae present posteriorly,

rarely also present medially. Propodeal spiracle

large, 2.9x as long as wide. Spiracular carina nar-

row or more or less covered with wrinkles. Spirac-

ular index 0.7. 1% gastral tergite 3.6x, 2"! tergite

1.8x as long as wide. Petiole with large glymmae,

which are bordered by longitudinal carinae dor-

sally, with very fine sculpture in front of the glym-

mae. Petiolar suture fine. 1 gastral sternite coria-

ceous or finely rugose, sometimes smooth medial-

ly. Thyridium oval. Thyridial index about 1.5.

Epipleurum separated from the 3" gastral tergite,

the crease marked with black. Epipleural index

0.7-0.8. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellow medially. Tegu-

la yellowish brown to dark brown. Wings hyaline.

Stigma yellowish brown medially. Front leg from

the trochanter onwards yellowish red. Mid femur

yellowish red on the apical 0.3. Mid tibia yellow-

ish red. Mid tarsus dark brown. Hind tibia brown

to dark brown, narrowly black basally, sometimes

also marked with black apically, sometimes com-

pletely black. 2" gastral tergite red on the posteri-

or 0.3. 3" to 5" tergites red. Posterior gastral ster-

nites sometimes reddish brown. Pilosity whitish.

Specimens from central Asia and eastern Russia

are often darker: Mid leg black, marked with dark

brown. Hind leg black. 2" gastral tergite complete-

ly black or red on the posterior 0.1. 4" tergite for

the greater part black. 5" tergite completely black.

d: Similar to the 9.

Material (52??, 3838): Russia: Yevreyskiy A.O.: Bi-

robidzhan; Khabarovsk Terr.: Khekhtsir near Kha-

barovsk, Troitskoye; Primorye: Kamenushka, Lazo,

Roshchino on River Iman, Shkotovo (PET), Spassk

Dalniy (MOS, PET), Ussuriysk; Kuril Isl.: Tretyakovo

on Kunashir (PET). Kazakhstan: Alma-Ata (LIN, PET),

Talgar (PET). Kirgiziya: Dzhangi-Dzhol (LIN). Japan:

Hokkaido: Jozankei (AEI), Sapporo (AEI, SAP), Toka-

chi-dake (SAP); Honshu: Hakone near Tokyo (WAS),

Hiroshima (AEI), Nagano (OTT), Sasayama (SAP),

Yokohama (AEI).

Dusona bucculenta (Holmgren, 1860)

Description. 2: Body length 9-11 mm. Temples

slightly constricted behind the eyes, 0.9-1.0x as

long as the width of an eye, tangential lines on

eyes and temples intersect on the propodeum or

the 1% gastral segment. Ocellar index 1.0. Frons

not depressed, densely rugose and rugose-punc-

tate, with short radial wrinkles, without a median

longitudinal carina. Antennal sockets close to each

other. Antennal carina low and narrow. Face and

dorsal half of the clypeus very densely punctate

on a finely coriaceous or smooth background.

Clypeus slightly concave, densely punctate on a

finely coriaceous or smooth background ventral-

ly, its apex slightly convex medially. Malar space

index 0.5. Genal carina low ventrally, bent side-

wards, joining the slightly raised oral carina at a

right angle. Genal index 0.3. Temple with rather

dense punctation on a very finely coriaceous or

almost smooth background medially. Flagellum

with 40-45 segments, filiform, the apical 0.2 slight-

ly attenuate, 2" segment 1.4x, shortest segment

0.8x, penultimate segment 1.2x as long as wide.

Pronotum densely rugose-punctate on a smooth

background laterally on the dorsal 0.3, with longi-

tudinal wrinkles ventrally. Mesoscutum and

scutellum strongly and densely punctate on a

smooth background. Scutellum with lateral cari-

nae on the anterior 0.6. Prescutellar groove smooth

medially, with longitudinal striae laterally. Ven-

tral part ofthe prepectal carina more or less strong-

ly raised, merging with the raised transverse car-

ina or with the pleural part, the latter low, its

dorsal 0.3 obliterated. Sometimes the pleural part

is obliterated also ventrally and present only me-

dially. Mesopleurum with strong and rather dense

or dense punctation on a smooth background.

Speculum punctate posteriorly, finely coriaceous

and shining anteriorly. Depression in front of the

speculum with distinct longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum densely rugose-punctate on a smooth back-

ground. Pleural carina distinct. Areolet large, short-

stalked, receiving the 2" recurrent vein slightly

distad of the middle. Nervellus vertical, weakly

intercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 4.8x as long as high. Hind tarsal

claws rather long, with distinct pectinate teeth

over their total length. Propodeum extending to

0.4 of the hind coxae, broadly and not deeply

depressed, sulcus with transverse wrinkles medi-

ally and posteriorly, sometimes also anteriorly,

costulae partly present, longitudinal carinae present

posteriorly and sometimes also medially. Propo-

deal spiracle rather large, 2.9x as long as wide.

Spiracular carina narrow, sometimes obliterated.

Spiracular index 0.5. 1* gastral tergite 4.8x, 2"4

tergite 1.7x as long as wide. Petiole without or

with very small glymmae, the latter at the anterior

edge of indistinct narrow lateral areas, petiole

smooth on front of these areas, sometimes with

very fine striae. Petiolar suture fine, sometimes

partly obliterated. 1“ gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.5. Epipleu-

rum not separated from the 3"! gastral tergite, the

latter with short or indistinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Mandible narrowly marked

with reddish brown medially. Wings hyaline. Stig-

ma medium brown to dark brown medially. Front

femur marked with yellowish brown dorsally.

Front tibia and tarsus yellowish brown. Mid tibia

yellowish brown to dark brown. 2‘ gastral tergite

red on the posterior 0.1. 3” and 4" tergites red.

Pilosity whitish.

d: Similar to the 2.

Variability: Apparently in European material the

ventral part of the prepectal carina is merging

with the transverse carina, and both are raised,

whereas in eastern Palearctic material the ventral

part is merging with the pleural part, and both are

rather low. The significance of this difference is

unknown.

Host: Heliothis viriplaca (Hufnagel, 1767) (Noctu-

idae) (PET).

Material (53??, 2639): Russia: Altaysk Terr.: Lake

Teletskoye (Hinz/ZSM, MIN); Yakutsk A.S.S.R.: Be-

tyakh na Amgu; Amur Prov.: Novorossiyka; Primor-

ye: Chernigovka, Novokachalinsk, Pogranichnyy (PET),

Vladivostok (HEL, KIS); Magadan Prov.: Tunguska

(PET). Kazakhstan: Alma-Ata (MOS), Yanvaruyevo

on Ural River (PET). Tadzhikistan: locality unreadable

(MOS). Kirgiziya: Kashkasi in Kirgyzskiy Mts., 1800 m,

Kyzyl-Oy, 1800 m (LIN). Mongoliya: Ara Hangay

Prov., Hentiy Prov., Selenge Prov. (PET). China: Hei-

longjiang Prov.: Charbin (= Harbin) (AEI), Maoershan

(WAR); Xinjiang Prov.: Houxia, 2000-3000 m (LIN).

Dusona bucculentoides Hinz & Horstmann,

spec. nov.

Types. Holotype (2): Russia, Primorye, Ussuri-

ysk, 2.9.1962 (MOS). — Paratypes: 1? Russia, Pri-

morye, Spassk Dalniy, 2.7.1961 (Hinz/ZSM);

13 Russia, Primorye, Khasan, 9.9.1982 (PET).

Description. 2: Body length 11-12 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.0. Frons not depressed, coriaceous and dense-

ly rugose-punctate, without a median longitudi-

nal carina and without radial wrinkles. Antennal

sockets close to each other. Face and dorsal 0.7 of

the clypeus densely rugose-punctate. Clypeus flat,

with a rather dense punctation on an almost

smooth background ventrally, the apex truncate

medially. Malar space index 0.5. Genal carina low

ventrally, slightly bent sidewards, joining the low

oral carina at an acute angle. Genal index 0.6.

Temple with fine rather dense punctation on a

smooth background medially. Flagellum with

43-46 segments, filiform, the apical 0.2 attenuate,

2X segment 1.5x, shortest segment 0.9x, penulti-

mate segment 1.3x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum distinctly and densely punc-

tate on an almost smooth or smooth background.

Scutellum with lateral carinae on the anterior 0.6.

Prescutellar groove smooth or with fine longitu-

dinal wrinkles. Prepectal carina low, its ventral

part merging with the pleural part, the dorsal 0.4

of the latter obliterated. Transverse carina obliter-

ated. Mesopleurum strongly and densely punc-

tate on a smooth background. Speculum with fine

oblique striae. Depression in front of the specu-

lum with distinct longitudinal striae, which merge

with the punctation anteriorly. Metapleurum dis-

tinctly and densely rugose-punctate. Pleural cari-

na distinct. Areolet large, pointed or short-stalked,

receiving the 2" recurrent vein distad of the mid-

dle. Nervellus almost vertical, weakly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 4.6x as long as high. Hind tarsal claws

rather long, with distinct pectinate teeth almost

over their total length. Propodeum extending to

the middle of the hind coxae, broadly and not

deeply depressed, with transverse wrinkles al-

most all over the broad sulcus, costulae and longi-

tudinal carinae partly indicated. Propodeal spira-

cle large, 3.1x as long as wide. Spiracular carina

distinct, narrow. Spiracular index 0.4. 1 gastral

tergite 4.3x, 2" tergite 1.7x as long as wide. Peti-

ole without glymmae, with narrow lateral areas,

with very fine sculpture or smooth in front of the

lateral areas. Petiolar suture partly obliterated. 1*

gastral sternite smooth. Thyridium elongate. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly reddish brown medially. Flag-

ellum medium brown apically. Wings hyaline.

Stigma brownish black medially. Front femur with

a yellow stripe anteriorly. Front tibia yellowish

red. Front tarsus brown. Mid tibia dark brown. 2“

gastral tergite red on the posterior 0.1. 3" tergite

red. Pilosity whitish.

d: Temples 0.8x as long as the width of an

eye. Propodeum almost not depressed and with

transverse wrinkles only medially. In other char-

acters similar to the 2.

Dusona capitator Hinz, 1985

Description. ?: Body length 9-10 mm. Temples

scarcely constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the apex of the gaster. Ocellar

index 1.7 (Fig. 2). Frons not depressed, very densely

punctate on a smooth background, with a fine

median longitudinal carina, with fine radial wrin-

kles, which extend to half the distance to the me-

dian ocellus. Antennal carina low and narrow.

Face and dorsal half of the clypeus densely rugose

on a smooth background. Clypeus slightly con-

cave, rather densely punctate on a finely coria-

ceous or smooth background ventrally, its apex

slightly convex medially. Malar space index 0.4.

Genal carina low ventrally, slightly bentsidewards,

joining the low oral carina at an acute angle. Genal

index 0.2. Temple finely coriaceous and with fine

rather dense punctation medially. Flagellum with

40-43 segments, slightly fusiform, 2"! segment 1.7x,

shortest segment 0.9-1.0x, penultimate segment

1.3x as long as wide. Pronotum densely punctate

on a smooth background laterally on the dorsal

half, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum with rather dense or dense

punctation on a smooth background. Scutellum

with lateral carinae on the anterior half. Prescutel-

lar groove für the greater part smooth. Prepectal

carina low, ventral part merging with the pleural

part, the dorsal 0.4 of the latter obliterated. Trans-

verse carina obliterated. Mesopleurum densely

punctate on a smooth background. Speculum

punctate posteriorly, smooth anteriorly. Depres-

sion in front of the speculum with distinct longi-

tudinal wrinkles, which merge with the puncta-

tion anteriorly. Metapleurum very densely punc-

tate on a smooth background. Pleural carina

complete. Areolet large, narrowly sessile or short-

stalked, receiving the 2” recurrent vein slightly

distad of the middle. Nervellus slightly reclivous,

weakly intercepted. Nervellar index 0.7-0.8. Tibial

spur index 0.5. Hind femur 3.8x as long as high.

Hind tarsal claws long, with short pectinate teeth

basally and medially. Propodeum extending to

0.4 of the hind coxae, distinctly and rather nar-

rowly depressed, with short transverse striae in

the longitudinal furrow, very densely punctate or

rugose-punctate on a smooth background beside

the furrow, carinae almost completely obliterated.

Propodeal spiracle rather large, 2.9x as long as

wide. Spiracular carina indistinct or replaced by

several wrinkles. Spiracular index 0.5. 1” gastral

tergite 3.6x, 2" tergite 1.3x as long as wide. Peti-

ole without glymmae, with slightly indicated nar-

row lateral areas, smooth in front of these areas.

Petiolar suture fine, sometimes partly obliterated.

1" gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3" gastral tergite, but a narrow lobe may

be present anteriorly, the tergite with indistinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible yellowish red medially. Wings

hyaline. Stigma medium brown medially. Front

leg from the femur onwards yellowish red. Mid

femur yellowish brown on the apical 0.7. Mid

tibia and tarsus yellowish brown. Hind tibia yel-

lowish brown or rarely dark brown medially, nar-

rowly marked with dark brown or black basally

and apically, sometimes not marked with brown

or black. 2" gastral tergite red on the posterior 0.7,

rarely more or less completely black dorsally. 3"

and 4" tergites red. Pilosity whitish.

d: Similar to the ®.

Material (782?, 5334): Turkmeniya: Bol-Balkhan

Mts. (Hinz/ZSM, PET). Uzbekistan: Samarkand. Ka-

zakhstan: Alma-Ata (MOS), Arys, Dzhambul. Kirgizi-

ya: Aravan near Osh (LIN), River Chatkal (PET), Ta-

lasskiy Alatau Mts., 1800-1900 m (LIN). Mongoliya:

Töv Prov. (= Central Prov.): Kerulen (PET).

Dusona carinator Hinz, 1979

Description. ?: Body length 11mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous and with very fine irregular wrinkles,

with a distinct and slightly raised median longitu-

dinal carina, without radial wrinkles. Antennal

carina low, slightly widened to a very narrow

crescent-shaped plate, the latter with fine trans-

verse striae. Face and dorsal half of the clypeus

coriaceous and densely punctate. Clypeus flat,

finely coriaceous and with rather dense puncta-

tion ventrally, its apex slightly convex medially.

Malar space index 0.4. Genal carina low and

straight ventrally, joining the low oral carina at an

acute angle. Genal index 0.7. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 49-51 segments, filiform,

the apical 0.3 attenuate, 2”! segment 1.8x, shortest

segment 0.9x, penultimate segment 1.2x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous

and very densely punctate. Scutellum with lateral

carinae on the anterior 0.2. Prescutellar groove

almost completely smooth. Ventral part of the

prepectal carina merging with the transverse car-

ina, both carinae low. Pleural part of the prepectal

carina completely obliterated. Pleural part of the

prepectus densely rugose-punctate. Mesopleurum

distinctly coriaceous and with rather dense or

dense punctation. Speculum punctate posteriorly,

coriaceous and dull anteriorly. Depression in front

of the speculum dull, dorsally with short longitu-

dinal wrinkles, which merge with the punctation

anteriorly, ventrally coriaceous and granulate-stri-

gose. Metapleurum finely and densely rugose-

punctate or rugose. Pleural carina distinct. Areo-

let large, short-stalked, receiving the 2" recurrent

vein at or basad of the middle. Nervellus slightly

inclivous, distinctly intercepted. Nervellar index

0.8. Tibial spur index 0.7. Hind femur 5.5x as long

as high. Hind tarsal claws rather short, with short

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, with a narrow

shallow longitudinal furrow, with long transverse

wrinkles in and beside the furrow, carinae almost

completely obliterated. Propodeal spiracle large,

3.3x as long as wide. Spiracular carina narrow.

Spiracular index 0.6. 1% gastral tergite 4.8x, 2”

tergite 1.7x as long as wide. Petiole without glym-

mae, with narrow lateral areas, smooth in front of

these areas. Petiolar suture almost completely ob-

literated. 1* gastral sternite for the greater part

smooth, with few transverse striae posteriorly.

Thyridium elongate. Thyridial index about 1.0.

Epipleurum separated from the 3° gastral tergite,

the crease marked with black. Epipleural index

0.6-0.7. Gaster distinctly compressed posteriorly.

Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown.

Mandible broadly yellow medially. Wings hya-

line. Stigma dark brown medially. Front leg from

the tip of the coxa onwards yellowish red. Mid

femur yellowish brown on the apical 0.2-0.5. Mid

tibia and tarsus yellowish brown. Hind tibia yel-

lowish brown, narrowly marked with dark brown

basally. Hind tarsus dark brown. 2" gastral tergite

black dorsally and posteriorly, laterally marked

with red in front of the posterior edge. 3"! and 4"

tergites red, sometimes narrowly marked witn

black dorsally. 5" tergite completely black or

marked with red anteriorly and laterally. Posteri-

or gastral sternites red or brown. Pilosity whitish.

d unknown.

Material (2?2): Russia: Kuril Isl.: Kunashir (SAP). Ja-

pan: Honshu: Kamikochi near Norikura-dake (AEI).

Dusona carinifrons (Holmgren, 1860)

Description. 2: Body length 14-17 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.8. Frons scarcely de-

pressed, criaceous, densely rugose-punctate, with

a distinct median longitudinal carina, which is

raised medially, without radial wrinkles. Anten-

nal carina low and narrow. Face and dorsal 0.7 of

the clypeus coriaceous, densely rugose-punctate.

Clypeus flat, coriaceous and densely punctate ven-

trally, its apex slightly convex medially. Malar

space index 0.5. Genal carina low and rather

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.6. Temple

coriaceous and with fine disperse punctation me-

dially. Flagellum with 58-62 segments, filiform,

the apical 0.3 distinctly attenuate, 2"! segment

1.4x, shortest segment 0.8x, penultimate segment

1.2x as long as wide. Pronotum densely rugose-

punctate on a smooth background laterally on the

dorsal half, with longitudinal wrinkles ventrally.

Mesoscutum finely coriaceous, very densely punc-

tate. Scutellum with rather dense or disperse punc-

tation on an almost smooth background, with lat-

eral carinae on the anterior 0.4. Prescutellar groove

smooth medially, with distinct longitudinal striae

laterally. Ventral part of the prepectal carina merg-

ing with the transverse carina, both carinae slight-

ly raised, the former not emarginate medially.

Pleural part of the prepectal carina completely

obliterated. Pleural part of the prepectus densely

rugose-punctate on a smooth background. Meso-

pleurum with rather dense or dense punctation

on a very finely coriaceous and shining back-

ground. Speculum punctate and with strong lon-

gitudinal striae posteriorly, finely coriaceous or

granulate-strigose and rather shining anteriorly.

Depression in front of the speculum with distinct

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum densely rugose-

punctate on a smooth background. Pleural carina

distinct. Areolet large, pointed, receiving the 2”*

recurrent vein at or slightly basad of the middle

(Fig. 23). Nervellus almost vertical, weakly inter-

cepted. Nervellar index 0.6. Tibial spur index 0.7.

Hind femur 6.0x as long as high. Hind tarsal claws

rather short, with distinct short pectinate teeth

almost over their total length. Propodeum extend-

ing to 0.4 of the hind coxae, broadly and rather

deeply depressed, sulcus with transverse wrin-

kles, costulae partly present, longitudinal carinae

distinct medially and posteriorly (Fig. 32). Propo-

deal spiracle rather large, 3.2x as long as wide.

Spiracular carina narrow. Spiracular index 0.5.

1* gastral tergite 4.8x, 2"X tergite 2.5x as long as

wide. Petiole without glymmae or with very small

and shallow glymmae at the anterior edge of nar-

row lateral areas, smooth in front of these areas.

Petiolar suture complete. 1” gastral sternite coria-

ceous anteriorly, rugose medially, transversely

striate posteriorly. Thyridium oval. Thyridial in-

dex 1.5-2.0. Epipleurum separated from the 34

gastral tergite anteriorly, folded in or pendent as a

very narrow rectangular lobe, the crease marked

with black, a black lateral stripe divergent from

the ventrolateral edge of the tergite posteriorly.

Epipleural index 0.2-0.3. Gaster strongly com-

pressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus medium brown.

Mandible broadly yellow medially. Tegula yel-

lowish brown. Wings hyaline. Stigma medium

brown medially. Front leg from the tip of the coxa

onwards yellowish red. Mid femur yellowish red

on the apical 0.3. Mid tibia yellowish red. Mid

tarsus yellowish brown. Hind femur yellowish

red on the apical 0.1. Hind tibia yellowish red.

Hind tarsus medium brown. 2" gastral tergite

dorsally red on the posterior 0.1-0.2, laterally al-

most completely red. 3"! and 4" tergites red, some-

times the 34 tergite marked with brown dorsally.

5" tergite red, sometimes marked with black pos-

teriorly. Posterior tergites often narrowly marked

with red ventrolaterally, posterior sternites often

red or marked with red. Pilosity whitish.

d: Carinae ofthe propodeum somewhat strong-

er, costulae sometimes complete. Mid femur for

the greater part yellowish- red. Posterior tergites

and genital clasper black. In other characters sim-

ilar to the ®.

Material (11??, 539): Russia: Altaysk Terr.: Gorno-

Altaysk (MIN); Krasnoyarsk Terr.: Shushenskoye

(MOS); Irkutsk Prov.: Irkutsk, Marituy on Lake Bay-

kal; Chita Prov.: Adrianovka, Nerchinskiy Zavod;

Yevreyskiy A.O.: Amurzet; Khabarovsk Terr.: Khekh-

tsir near Khabarovsk (PET). Kazakhstan: Alma-Ata

(MOS), Borovoye near Kokchetav (PET). Kirgiziya:

Terskey-Alatau Mts. (MOS).

Dusona carpathica (Szepligeti, 1916)

This species is very similar to D. angustifrons, it

differs from the latter in: Body slightly smaller.

Flagellum slightly stouter. Ventral part of the pre-

pectal carina merging with the transverse carina,

both carinae slightly raised.

Description. ?: Body length 6-8 mm. Temples

distinctly constricted behind the eyes, 0.9%x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.9. Frons not depressed, coriaceous, finely

and densely rugose-punctate, with a fine median

longitudinal carina, with short radial wrinkles,

which extend to 0.3 the distance to the median

ocellus. Antennal sockets rather close to each oth-

er. Antennal carina low and narrow, its rim slight-

ly bent upwards. Face and dorsal half of the

clypeus coriaceous and densely rugose-punctate.

Clypeus flat, finely coriaceous and with rather

dense punctation ventrally, its apex truncate me-

dially. Malar space index 0.4. Genal carina low

ventrally, slightly bent sidewards and distinctly

bent forwards, joining the oral carina at the base

of the mandible at an almost right angle. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 30-36 segments,

slightly fusiform, 2" segment 2.0x, shortest seg-

ment 0.8-0.9x, penultimate segment 0.9-1.0x as

long as wide. Pronotum densely and finely ru-

gose and punctate laterally on the dorsal 0.2, with

longitudinal wrinkles ventrally. Mesoscutum co-

riaceous and very densely punctate. Scutellum

coriaceous and densely punctate, with lateral car-

inae on the anterior 0.8. Prescutellar groove smooth

medially, with longitudinal striae laterally. Ven-

tral part of the prepectal carina merging with the

transverse carina, both carinae slightly raised, the

former scarcely emarginate medially. Pleural part

of the prepectal carina low, its dorsal 0.4 obliterat-

ed. Mesopleurum coriaceous, dull, densely punc-

tate. Speculum coriaceous and granulate-strigose,

with a small almost smooth and shining area ven-

trally. Depression in front of the speculum with

distinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum coriaceous,

finely and densely rugose. Pleural carina com-

plete. Areolet rather small, stalked, receiving the

2"d recurrent vein distinctly distad of the middle.

Nervellus vertical or slightly reclivous, weakly

intercepted. Nervellar index 0.7-0.8. Tibial spur

index 0.7. Hind femur 4.8x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to 0.4 of the hind

coxae, broadly and shallowly depressed, sulcus

for the greater part irregularly wrinkled, partly

with short transverse wrinkles, costulae partly

present, longitudinal carinae for the greater part

obliterated. Propodeal spiracle rather small, 2.7%

as long as wide. Spiracular carina sometimes wid-

ened. Spiracular index 0.3. 1“ gastral tergite 4.2x,

2"! tergite 1.8x as long as wide. Petiole with small

glymmae at the anterior edge of narrow lateral

areas, smooth in front of these areas. Petiolar su-

ture partly obliterated. 1* gastral sternite smooth.

Thyridium elongate. Thyridial index about 1.0.

Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown

to black. Mandible narrowly marked with yellow

dorsally. Tegula yellowish brown to dark brown.

Wings hyaline. Stigma yellowish brown medially.

Front leg from the femur onwards yellowish red.

Mid femur yellowish red on the apical 0.3 or com-

pletely yellowish red. Mid tibia yellowish red.

Mid tarsus brown. Hind femur usually black,

sometimes red or marked with red. Hind tibia

yellowish red or yellowish brown medially, nar-

rowly marked with brown or black basally, broadly

marked with brown or black apically. Hind tarsus

brown or black. 2" gastral tergite red on the pos-

terior 0.1-0.2 (rarely 0.3). 3" tergite red, sometimes

marked with brown dorsally. 4" tergite some-

times marked with red anteriorly. Pilosity whit-

ish.

d: Mandible broadly yellow medially. Front

and mid legs from the trochanter onwards yellow

or yellowish red, sometimes the tips of the coxae

yellow. In other characters similar to the 2 (also in

the variability of the colour pattern).

Material (36??, 33595): Russia: Tomsk Prov.: Tomsk;

Irkutsk Prov.: Irkutsk; Chita Prov.: Chita, Nerchinskiy

Zavod; Yakutsk A.S.S.R.: Olekminsk; Amur Prov.:

Novorossiyka; Primorye: Roshchino on River Iman

(PET); Magadan Prov.: Magadan, Debin (MOS); Kam-

chatka: Kozyrevsk. Kirgiziya: Tokmak (PET).

Dusona caudator Hinz, 1985

Description. 2: Body length 11mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.3. Frons slightly depressed,

coriaceous, finely and densely rugose punctate,

with a distinct median longitudinal carina, with

short radial wrinkles. Antennal carina low and

narrow. Face and dorsal 0.4 of the clypeus coria-

ceous and densely punctate. Clypeus flat, coria-

ceous and dispersely or rather densely punctate

ventrally, its apex truncate medially. Malar space

index 0.5. Genal carina low ventrally, slightly bent

sidewards, joining the slightly raised oral carina

at an acute angle. Genal index 0.8. Temple finely

coriaceous and with very fine and disperse punc-

tation medially. Flagellum with 45-46 segments,

filiform, the apical 0.2 attenuate, 2”! segment 1.3x,

shortest segment 0.7x, penultimate segment 1.0x

as long as wide. Pronotum coriaceous and dense-

Iy punctate laterally on the dorsal 0.3, with longi-

tudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, finely and densely punc-

tate. Scutellum with lateral carinae on the anterior

0.2. Prescutellar groove smooth. Prepectal carina

low, ventral part merging with the pleural part,

the latter complete, joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum with rather dense or dense puncta-

tion on a slightly coriaceous and shining back-

ground. Speculum punctate posteriorly, slightly

granulate-strigose and shining anteriorly. Depres-

sion in front of the speculum dorsally with dis-

tinct rather short longitudinal wrinkles, ventrally

with fine longitudinal striae and wrinkles, which

merge with the punctation anteriorly. Metapleu-

rum finely and densely rugose, sometimes partly

coriaceous and finely punctate. Pleural carina fine,

indistinct and covered with wrinkles medially.

Areolet large, short-stalked, receiving the 2”! re-

current vein at or basad of the middle. Nervellus

slightly reclivous, distinctly intercepted. Nervel-

lar index 0.7. Tibial spur index 0.7. Hind femur

4.6x as long as high. Hind tarsal claws long, with

short pectinate teeth basally. Propodeum extend-

ing to 0.3 of the hind coxae, broadly and rather

deeply depressed, sulcus for the greater part ir-

regularly wrinkled, with some short transverse

wrinkles anteriorly, with some longitudinal wrin-

kles posteriorly, costulae and longitudinal carinae

complete. Propodeal spiracle large, 2.2x as long as

wide. Spiracular carina narrow and indistinct.

Spiracular index 0.5. 1 gastral tergite 2.9x, 2nd

tergite 1.5x as long as wide. Petiole with rather

large glymmae, smooth in front of the glymmae.

Petiolar suture fine, complete. 1* gastral sternite

coriaceous. Thyridium almost circular. Thyridial

index about 1.5. Epipleurum separated from the

3 gastral tergite, the crease marked with black.

Epipleural index 0.8. Gaster slightly compressed

posteriorly. Ovipositor index 2.2.

Colour: Black. Mandible tinged with reddish

brown medially. Wings hyaline. Stigma dark

brown medially. Front femur reddish brown on

the apical 0.3. Front tibia and tarsus reddish brown.

Mid tibia dark reddish brown. 2" gastral tergite

red on the posterior 0.4. 3" tergite red. 4" tergite

marked with red anteriorly.Pilosity whitish.

d: Frons with a rather fine median longitudi-

nal carina. Flagellum less stout, 2° segment 1.7x,

shortest segment 0.9x as long as wide. Metapleu-

rum coriaceous, finely and densely rugose and

rugose-punctate. Propodeum with the sulcus rath-

er narrowly and not deeply depressed, costulae

and longitudinal carinae complete or almost com-

plete. Glymmae rather small, but distinct. 2" gas-

tral tergite red on the posterior 0.2. 3 and 4

tergites red, the 4" tergite with a small black mark

dorsally. In other characters similar to the 2.

Material (17, 234): Russia: Primorye: Roshchino on

River Iman (Hinz/ZSM, PET).

Dusona celator Hinz, 1985

Description. 2: Body length 14-16 mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons slightly but dis-

tinctly depressed above the antennal carina, coria-

ceous and densely punctate dorsally, with a dis-

tinct median longitudinal carina, with long radial

wrinkles, which almost reach the median ocellus.

Antennal carina slightly but distinctly raised, its

rim distinctly bent upwards (ear-like), forming a

narrow crescent-shaped plate, the latter with fine

transverse striae. Face and dorsal 0.7 ofthe clypeus

coarsely and densely rugose-punctate. Clypeus

flat, strongly and densely punctate on a finely

coriaceous background ventrally, its apex trun-

cate medially. Malar space index 0.4. Genal carina

low ventrally, straight or slightly bent sidewards,

joining the slightly raised oral carina at an acute or

a right angle. Genal index 0.4. Temple finely coria-

ceous and with fine disperse punctation medially.

Flagellum with 57-61 segments, filiform, the api-

cal 0.3 attenuate, 2” segment 1.4x, shortest seg-

ment 1.0x, penultimate segment 1.5x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal half, with longitudinal wrinkles

ventrally. Mesoscutum coriaceous, finely and

densely punctate. Scutellum densely punctate on

an almost smooth background, with lateral cari-

nae on the anterior 0.3. Prescutellar groove with

fine longitudinal striae. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

distinct, almost complete, the dorsal 0.2 replaced

by several wrinkles. Transverse carina obliterat-

ed. Mesopleurum strongly and densely punctate

ona very finely coriaceous or smooth background.

Speculum punctate posteriorly, coriaceous or gran-

ulate-strigose anteriorly. Depression in front of

the speculum dorsally with distrinct longitudinal

striae, which more or less merge with the puncta-

tion anteriorly, ventrally finely coriaceous and

shining. Metapleurum densely rugose and rugose-

punctate, not coriaceous. Pleural carina distinct.

Areolet large, short-stalked, receiving the 2"’ re-

current vein slightly basad of the middle. Nervel-

lus vertical, distinctly intercepted. Nervellar in-

dex 0.7. Tibial spur index 0.5. Hind femur 5.2x as

long as high. Hind tarsal claws rather long, with

short pectinate teeth basally. Propodeum extend-

ing to 0.3 of the hind coxae, broadly and deeply

depressed, sulcus for the greater part or complete-

ly transversely wrinkled, costulae shortened, lon-

gitudinal carinae distinct posteriorly, more or less

complete medially. Propodeal spiracle large, 3.2x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.4. 1% gastral tergite 4.0x, 2"d tergite

1.8x as long as wide. Petiole with distinct glym-

mae, smooth in front of the glymmae. Petiolar

suture fine, partly obliterated. 1“ gastral sternite

smooth. Thyridium oval. Thyridial index about

1.0. Epipleurum separated from the 3" gastral

tergite anteriorly, the crease marked with black.

Epipleural index 0.3. Gaster distinctly compressed

posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible broadly yellowish red medially. Tegula

dark brown. Wings slightly infuscate. Stigma light

brown medially. Front trochanter and femur yel-

lowish red. Front tibia and tarsus yellow. Mid

femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. Hind tibia black, rather

narrowly marked with dark yellowish brown sub-

basally. 2" gastral tergite narrowly tinged with

red in front of the posterior edge. 3" and 4" tergites

red, the 4" marked with black posteriorly. Pilosity

whitish.

d: Tip of the front coxa yellowish red. In other

characters similar to the 9.

Material (4?%, 238): Russia: Chita Prov.: Adrian-

ovka, Karymskaya (Hinz/ZSM), Urulyunguy E of

Borzya; Primorye: Kiyevka (PET). North Korea: Kyong-

song (SAW).

Dusona chabarowski Hinz & Horstmann,

spec. nov.

This species was characterized by Hinz (1979: 216)

under the name D. bicoloripes. It is similar to

D. bicoloripes (see under this name) and D. stragifex.

Specimens without raised points on the frons dif-

fer from D. stragifex in the less strongly striate

depression in front of the speculum and the less

strongly sculptured propodeum, but the differen-

tiation may be difficult. Whereas D. bicoloripes and

D. stragifex both occur from Europe to Japan,

D. chabarowski is only known from eastern Russia

so far.

Types. Holotype (2): Russia, Khabarovsk Terr.,

Khekhtsir near Khabarovsk, 20.6.1983 (PET). —

Paratypes: 5?2, 385 same locality, 12.6.-4.7.1983

(322, 288 PET, 229,15 Hinz/ZSM); 19, 13 Rus-

sia, Khabarovsk Terr., Troitskoye, 24.-28.6.1983

(PET); 2?2 Russia, Primorye, Anisimovka near

Shkotovo, 17.-24.6.1979 (PET); 12 Russia, Primor-

ye, Khasan, 3.7.1981 (PET); 12, 339 Russia, Pri-

morye, Roshchino on River Iman, 29.5.-5.6.1966

(PET); 259 Russia, Primorye, Spassk Dalniy,

28.5.1985 (KIS).

Description. 2: Body length 9-13 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 1.2. Frons slightly depressed, coriaceous,

partly with fine and dense punctation, with a fine

median longitudinal carina, with short radial wrin-

kles, usually on either side with a distinct or indis-

tinct raised point between the antennal socket and

the eye, which may be completely obliterated.

Antennal carina low and narrow. Face and dorsal

0.7 of the clypeus coriaceous and densely rugose-

punctate. Clypeus flat, coriaceous and with rather

dense punctation ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 46-55 seg-

ments, filiform, the apical 0.3 attenuate, 2" seg-

ment 1.8x, shortest segment 0.9x, penultimate seg-

ment 1.2x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, very densely punctate.

Scutellum with lateral carinae on the anterior 0.7.

Prescutellar groove for the greater part smooth.

Prepectal carina low, the ventral part merging

with the pleural part, the latter almost joining the

anterior edge of the mesopleurum. Transverse

carina present anteriorly. Mesopleurum distinctly

coriaceous, with rather dense or dense puncta-

tion. Speculum granulate-strigose, rather dull.

Depression in front of the speculum granulate-

strigose and with fine longitudinal striae, which

often merge with a narrow coriaceous area with-

out punctation anteriorly. Metapleurum coria-

ceous, finely and densely punctate or rugose-punc-

tate. Pleural carina distinct. Areolet large, stalked,

usually receiving the 2” recurrent vein at the mid-

dle. Nervellus vertical or slightly inclivous, dis-

tinctly intercepted. Nervellar index 0.6-0.8. Tibial

spur index 0.6. Hind femur 4.4x as long as high.

Hind tarsal claws short, with distinct pectinate

teeth basally. Propodeum extending to 0.3 of the

hind coxae, broadly and not deeply depressed,

sulcus for the greater part irregularly wrinkled,

with a short median longitudinal bulge and at the

same time with distinct long transverse striae pos-

teriorly, costulae shortened, longitudinal carinae

present posteriorly, obliterated medially. Propo-

deal spiracle large, 3.2x as long as wide. Spiracu-

lar carina narrow. Spiracular index 0.7. 1“ gastral

tergite 3.1x, 2" tergite 1.7% as long as wide. Peti-

ole with large glymmae, which are bordered by

longitudinal carinae dorsally, petiole for the greater

part smooth in front of the glymmae. Petiolar

suture fine. 1* gastral sternite smooth medially,

finely coriaceous anteriorly and posteriorly. Thy-

ridium oval. Thyridial index about 1.5. Epipleu-

rum separated from the 3" gastral tergite, the

crease marked with black. Epipleural index 0.7-0.8.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.6.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly tinged with reddish brown

medially. Wings hyaline. Stigma medium brown

to dark brown medially. Front femur marked with

yellowish red dorsally and apically. Front tibia

and tarsus yellow to yellowish brown. Mid femur

yellowish brown on the apical 0.2. Mid tibia brown.

2" gastral tergite red on the posterior 0.1-0.3.

34 and 4" tergites red, often marked with black

dorsally (to a variable extend). Pilosity whitish.

d: Longitudinal carinae of the propodeum

sometimes present also medially. Maxillary pal-

pus yellowish brown. Mandible broadly yellow

medially. Front femur for the greater part yellow-

ish red. 2"! gastral sometimes completely black

dorsally, with narrow red marks laterally. 3° and

4" tergites often black dorsally, broadly marked

with red laterally. In other characters similar to

the 2.

Dusona chechziri Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarovsk, 14.6.1985 (PET). —

Paratype: 1? same locality, 13.6.1985 (Hinz /

ZSM).

Description. ?: Body length 5-6 mm. Temples not

much constricted behind the eyes, 0.9x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the posterior edge of the pro-

podeum. Ocellar index 1.1. Frons scarcely de-

pressed, coriaceous, dull, with very fine disperse

punctation, with a fine median longitudinal cari-

na, with short radial wrinkles, which extend to 0.3

the distance to the median ocellus. Antennal cari-

na low and narrow. Face and clypeus coriaceous,

finely and densely punctate. Clypeus flat, its apex

slightly convex medially. Malar space index 0.4.

Genal carina low and straight ventrally, joining

the slightly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 34-35 segments, filiform, the apical 0.1 slight-

ly attenuate, 2" segment 2.0x, shortest segment

0.9x, penultimate segment 0.9-1.0x as long as wide.

Pronotum with fine punctation and fine longitu-

dinal wrinkles laterally. Mesoscutum and scutel-

lum coriaceous, finely and densely punctate.

Scutellum with lateral carinae on the anterior 0.4.

Prescutellar groove smooth or with some fine lon-

gitudinal striae. Prepectal carina low, the ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the meso-

pleurum. Transverse carina obliterated. Mesopleu-

rum with rather dense or dense punctation on a

coriaceous and dull background. Speculum finely

coriaceous and granulate-strigose. Depression in

front of the speculum with rather long longitudi-

nal wrinkles, which merge with the punctation

anteriorly. Metapleurum coriaceous and with fine

irregular wrinkles. Pleural carina distinct. Areolet

rather small, long-stalked, receiving the 2"4 recur-

rent vein distad of the middle. Nervellus slightly

inclivous, intercepted. Nervellar index 0.8. Tibial

spur index 0.7. Hind femur 5.3x as long as high.

Hind tarsal claws short, with long pectinate teeth

basally and medially. Propodeum extending to

the basal 0.3 of the hind coxae, slightly and broad-

ly depressed, coriaceous and with fine irregular

wrinkles in the sulcus, with some longitudinal

wrinkles posteriorly. Propodeal spiracle small, 2.0x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.7. 1% gastral tergite 3.0x, 2” tergite

1.1x as long as wide. Petiole with distinct glym-

mae, smooth in front of the glymmae. Petiolar

suture partly obliterated. 1“ gastral sternite for the

greater part smooth, with fine transverse striae

posteriorly. Thyridium oval. Thyridial index about

1.0. Epipleurum separated from the 3" gastral

tergite, the horizontal crease marked with black.

Epipleural index 0.6-0.7. Gaster distinctly but not

strongly compressed posteriorly. Ovipositor in-

dex 0.7.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible broad-

ly yellow medially. Tegula yellowish. Wings hya-

line. Stigma medium brown medially. Front leg

from the trochanter onwards yellowish, the tro-

chanter and femur tinged with yellowish brown.

Mid leg from the apical 0.2 of the femur onwards

yellowish. Hind tibia broadly light yellow medi-

ally, narrowly black basally, broadly black apical-

ly. Hind tarsus yellowish to yellowish brown. 2“

gastral tergite marked with red laterally. 3" tergite

broadly marked with red laterally. Pilosity whitish.

d unknown.

Dusona chinensis Horstmann, spec. nov.

Holotype (?): China, Shaanxi Prov., Qinling

Mts., Xunyangba, 1000-1300 m, 23.5.-13.6.1998

(LIN).

Description. 2: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.6x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.6. Frons slightly depressed, coriaceous,

with fine and rather dense or dense punctation,

with narrow smooth areas ventrally, with a fine

and partly obliterated median longitudinal cari-

na, without radial wrinkles. Antennal carina nar-

row, slightly but distinctly raised, its rim bent

upwards. Face and dorsal 0.6 of the clypeus dense-

ly rugose-punctate and rugose. Clypeus flat, fine-

ly coriaceous and with fine and rather dense punc-

tation ventrally, its apex truncate medially. Malar

space index 0.3. Genal carina low ventrally, slight-

ly bent forwards, joining the slightly raised oral

carina close to the base of the mandible at an acute

angle. Genal index 0.1. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

elluım with 50 segments, filiform, the apical 0.3

attenuate, 2”! segment 2.2x, shortest segment 1.2x,

penultimate segment 1.4x as long as wide. Prono-

tum densely rugose-punctate and rugose laterally

on the dorsal 0.2, with longitudinal wrinkles ven-

trally. Mesoscutum coriaceous, very densely punc-

tate. Scutellum densely rugose-punctate, with lat-

eral carinae on its anterior 0.6. Prescutellar groove

with fine longitudinal striae. Ventral part of the

prepectal carina merging with the transverse car-

ina, both carinae low, the former not emarginate

medially. Pleural part of the prepectal carina very

low, its ventral part complete but indistinct, its

dorsal 0.3 covered with wrinkles. Mesopleurum

finely coriaceous, distinctly and densely punctate.

Speculum finely coriaceous, partly almost smooth.

Depression in front of the speculum with rather

long longitudinal striae, which merge with the

punctation anteriorly. Metapleurum densely ru-

gose. Pleural carina fine but distinct. Areolat rath-

er rather small, long-stalked, receiving the 2" re-

currrent vein slightly basad of the middle. Nervel-

lus slightly reclivous, not intercepted. Tibial spur

index 0.5. Hind femur 5.8x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally and medially. Propodeum extending to the

middle of the hind coxae, narrowly and not deep-

ly depressed, almost without carinae, with trans-

verse wrinkles almost over its total width. Propo-

deal spiracle rather large, 3.3x as long as wide.

Spiracular carina narrow. Spiracular index 0.5.

1" gastral tergite 5.3x, 2” tergite 2.9x as long as

wide. Petiole with small and rather shallow glym-

mae at the anterior edge of indistinct narrow lat-

eral areas, smooth in front of the glymmae. Petio-

lar suture for the greater part obliterated. 1“ gas-

tral sternite smooth. Thyridium elongate. Thyridial

index about 1.5. Epipleurum not separated from

the 3"4 gastral tergite, the latter with distinct black

lateral stripes, which are situated well above the

ventrolateral edge of the tergite. Gaster distinctly

compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Mandible

narrowly yellow medially and dorsally. Tegula

yellowish brown to brown. Wings hyaline. Stig-

ma blackish medially. Front leg from the trochant-

er onwards yellow, femur yellowish red. Mid fe-

mur reddish brown. Mid tibia and tarsus yellow.

Hind tibia broadly yellowish brown medially,

narrowly black basally and apically. 2"“ gastral

tergite red on the posterior 0.1. Tergites from the

3"! onwards red, 3'% tergite distinctly tinged with

brown dorsally, the posterior tergites indistinctly

tinged with brown dorsally. Posterior gastral ster-

nites red. Pilosity whitish.

d unknown.

Dusona circumcinctus (Förster, 1868)

Delopia nigrifemur Sonan, 1929 (? syn.)

Campoplegidea melanocerus Uchida, 1932 (syn. nov.)

Townes (1984: 193 f.) studied the holotype (4) of

Delopia nigrifemur Sonan (from Taipei/ Taiwan)

and compared it with a d in his collection from

Kuandouchi/Taiwan (AEI). The identification of

the taxon is based on this specimen. This synony-

my is not beyond doubt, however, because the

type of D. nigrifemur is published as having been

bred from Orgyia postica (Walker, 1855) (Lyman-

triidae) (Sonan 1929: 418), whereas Dusona circum-

cinctus (Förster) was bred in northern Germany

from Cabera exanthemata (Scopoli, 1763) and C. pu-

saria (Linnaeus, 1758) (Geometridae) (1%, 738, leg.

and coll. Hinz/ZSM). The holotype (3) of Campo-

plegidea melanocerus Uchida from Anping/ Taiwan

(DEI) has recently been studied by the junior au-

thor.

Description. $: Body length 9-11 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely punctate, with a fine median

longitudinal carina, with short and indistinct ra-

dial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.7 of the clypeus coriaceous and

densely rugose-punctate. Clypeus flat, finely co-

riaceous and with rather densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.3. Genal carina low ventrally, slightly bent

sidewards, joining the slightly raised oral carina

at a right angle. Genal index 0.5. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 45-50 segments, filiform,

the apical 0.2 attenuate, 2" segment 2.0x, shortest

segment 1.0x, penultimate segment 1.2x as long

as wide. Pronotum with longitudinal wrinkles,

the dorsal 0.2 with additional punctures. Meso-

scutum coriaceous and densely rugose-punctate.

Scutellum coriaceous and rather densely punc-

tate, with lateral carinae on the anterior 0.4-0.6.

Prescutellar groove with distinct longitudinal stri-

ae. Ventral part of the prepectal carina merging

with the transverse carina, the former low, the

latter low or somewhat raised, its rim sometimes

bent upwards. Pleural part of the prepectal carina

indistinct ventrally and distinct medially, its dor-

sal 0.4 obliterated. Mesopleurum distinctly and

densely punctate on a finely coriaceous and shin-

ing background. Speculum with fine oblique stri-

ae. Depression in front of the speculum with long

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum densely rugose or

rugose-punctate. Pleural carina distinct. Areolet

large, pointed or short-stalked, receiving the 2"«

recurrent vein basad of the middle (Fig. 22).

Nervellus vertical, distinctly intercepted. Nervel-

lar index 0.7. Tibial spur index 0.6. Hind femur

5.0x as long as high. Hind tarsal claws short, with

rather long pectinate teeth basally and medially.

Propodeum extending to the middle of the hind

coxae, rather broadly and rather shallowly de-

pressed, sulcus coriaceous or with irregular wrin-

kles anteriorly, with long transverse wrinkles pos-

teriorly, costulae complete, longitudinal carinae

obliterated medially and anteriorly, present pos-

teriorly (Fig. 33). Propodeal spiracle large, 4.0x as

long as wide. Spiracular carina narrow. Spiracular

index 0.2-0.4. 1" gastral tergite 5.3x, 2”! tergite

2.6x as long as wide. Petiole without glymmae or

with very small glymmae at the anterior edge of

narrow lateral areas, smooth in front of these are-

as. Petiolar suture fine, partly obliterated. 1* gas-

tral sternite for the greater part smooth, with some

transverse striae posteriorly. Thyridium elongate.

Thyridial index about 1.0. Epipleurum not sepa-

rated from the 34 gastral tergite, the latter with

distinct black lateral stripes well above the vent-

rolateral edge. Gaster strongly compressed poste-

riorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellow to yel-

lowish brown. Mandible narrowly or broadly yel-

low medially. Tegula yellow or yellowish brown.

Wings hyaline. Stigma dark brown medially. Tip

of the front coxa sometimes marked with yellow.

Front trochanter and femur yellowish red. Front

tibia and tarsus yellow. Mid femur yellowish red

on the apical 0.2. Mid tibia yellow. Mid tarsus

brown. Hind tibia broadly yellowish brown or

brown medially, narrowly black basally, some-

times narrowly marked with brown apically. 2"4

gastral tergite black dorsally, to a variable extend

marked with red laterally. 3 and 4" tergites for

the greater part red, narrowly marked with black

dorsally. Posterior tergites for the greater part

black, narrowly marked with red ventrolaterally.

Posterior gastral sternites red or brown. Pilosity

whitish.

d: Similar to the ®.

Material (1322, 988): Russia: Chita Prov.: Chita,

Kozlovo; Amur Prov.: Novorossiyka; Khabarovsk

Terr.: Khekhtsir near Khabarovsk (PET); Primorye:

Khasan (KIS), Spassk Dalniy (Hinz/ZSM, KIS, PET));

Kuril Isl.: Dubovoye on Kunashir, Shikotan (PET).

China: Shaanxi Prov.: Qinling Mts. (LIN). Japan:

Hokkaido: Sapporo (AEI); Honshu: Sasayama (SAP).

Dusona collaris Hinz & Horstmann, spec. nov.

Holotype (?): Russia, Yakutsk (HEL).

Description. 2: Body length 7 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons scarcely depressed, coriaceous and

slightly wrinkled, without radial wrinkles, a me-

dian longitudinal carina slightly indicated. An-

tennal carina low and narrow. Face and dorsal

half of the clypeus coriaceous, distinctly and very

densely punctate. Clypeus flat, the ventral half

finely coriaceous and finely and rather densely

punctate, the apex slightly convex medially. Ma-

lar space index 0.5. Genal carina low ventrally,

slightly bent sidewards and forwards, joining the

oral carina at the base of the mandible at a right

angle. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 34 seg-

ments, filiform, the apical 0.2 attenuate, 2"X seg-

ment 2.3x, shortest segment 1.0x, penultimate seg-

ment 1.3x as long as wide. Pronotum with very

dense punctation laterally on the dorsal 0.2, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum finely coriaceous and densely punctate.

Scutellum with lateral carinae on its anterior half.

Prescutellar groove smooth (partly destroyed).

Ventral part of the prepectal carina joining the

transverse carina, both carinae strongly raised,

the former deeply emarginate medially. Pleural

part of the prepectal carina low, the dorsal 0.3

obliterated. Mesopleurum finely coriaceous, dis-

tinctly and densely punctate. Speculum with fine

oblique striae, a small ventral area smooth. De-

pression in front of the speculum with rather long

longitudinal wrinkles, which merge with the punc-

tation anteriorly. Metapleurum densely rugose-

punctate. Pleural carina distinct. Areolet short-

stalked, receiving the 2” recurrent vein slightly

basad of the middle. Nervellus slightly reclivous,

indistinctly intercepted. Nervellar index 0.6. Tibi-

al spur index 0.6. Hind femur 5.3x as long as high.

Hind tarsal claws short, whith short pectinate teeth

basally. Propodeum extending to the middle of

the hind coxae, broadly and rather deeply de-

pressed, almost without carinae, with short trans-

verse wrinkles in the sulcus, in other parts with

irregular wrinkles. Propodeal spiracle rather large,

2.7x as long as wide. Spiracular carina distinct,

partly widened. Spiracular index 0.5. 1" gastral

tergite 3.8x, 2" tergite 1.6x as long as wide. Peti-

ole without glymmae and lateral areas, smooth

laterally. Petiolar suture almost completely oblit-

erated. 1“ gastral sternite for the greater part

smooth, with fine transverse striae posteriorly.

Thyridium oval. Thyridial index about 1.0. Epi-

pleurum partly separated from the 3" gastral

tergite, the crease marked with black, black lateral

stripes present, merging with the crease anterior-

ly, divergent from the ventrolateral edge posteri-

orly. Epipleural index 0.3. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly yellowish medially. Wings

hyaline. Stigma yellowish medially. Front leg from

the trochanter onwards yellowish red. Mid femur

and tibia yellowish red. Mid tarsus light brown.

Hind femur reddish brown, indistinctly marked

with brown basally. Hind tibia reddish brown,

narrowly dark brown basally and apically. Hind

tarsus dark brown. 2” gastral tergite red on the

posterior 0.7. 3'% and 4" tergites red. 5" tergite red

anteriorly. Pilosity whitish.

d unknown.

Dusona confusator Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Kazakhstan, Alma-Ata,

1500 m, 12.8.1957 (MOS). — Paratypes: 1? Kaza-

khstan, 15 km S Alma-Ata, 27.5.1994 (LIN); 18

Kirgiziya, Lake Issyk-Kul, 20 km W Cholpon-Ata,

22.5.1994 (LIN); 18 Kirgiziya, Alayskiy Mts., Gul-

cha, 2500 m, 15.7.1996 (Hinz/ ZSM).

Description. 2: Body length 9-11 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

asthe width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.7. Frons not depressed, coriaceous,

densely rugose-punctate, with a fine median lon-

gitudinal carina, with long radial wrinkles, which

almost reach the median ocellus. Antennal carina

low and narrow. Face and dorsal 0.8 of the clypeus

very densely punctate on an almost smooth back-

ground. Clypeus flat, rather densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.4. Genal carina low and straight ventrally,

joining the distinctly raised oral carina at an acute

angle. Genal index 0.5. Temple coriaceous and

with rather dense punctation medially. Flagellum

with 40-42 segments, filiform, the apical 0.8 slight-

ly attenuate, 2" segment 1.6x, shortest segment

1.0x, penultimate segment 1.1x as long as wide.

Pronotum densely rugose-punctate laterally. Meso-

scutum and scutellum finely coriaceous and dense-

ly punctate. Scutellum with lateral carinae on the

anterior half. Prescutellar groove smooth or with

some fine longitudinal striae. Prepectal carina low,

the ventral part merging with the pleural part, the

latter almost joining the anterior edge of the meso-

pleurum, its dorsal end covered with wrinkles.

Transverse carina obliterated. Mesopleurum with

very dense punctation on a finely coriaceous or

almost smooth background, sometimes partly ru-

gose. Speculum rugose-punctate and with longi-

tudinal wrinkles. Depression in front of the specu-

lum with short longitudinal wrinkles, which merge

with the punctation anteriorly. Metapleurum with

coarse and dense irregular wrinkles. Pleural cari-

na fine, partly obliterated and covered with wrin-

kles. Areolet large, short-stalked, receiving the 2"“

recurrent vein slightly basad of the middle. Nervel-

lus slightly reclivous, weakly intercepted. Nervel-

lar index 0.7. Tibial spur index 0.7. Hind femur

5.5x as long as high. Hind tarsal claws long, with

distincty pectinate teeth basally and medially. Pro-

podeum extending to the middle of the hind cox-

ae, slightly and broadly depressed, sulcus for the

greater part irregularly wrinkled, sometimes with

some short transverse wrinkles in the midline,

costulae complete, longitudinal carinae partly in-

dicated. Propodeal spiracle small, 3.0x as long as

wide. Spiracular carina obliterated. Spiracular in-

dex 1.0. 1* gastral tergite 5.1x, 2" tergite 2.3x as

long as wide. Petiole without glymmae, with nar-

row lateral areas, which are bordered by carinae

dorsally and ventrally, petiole with very fine sculp-

ture or almost smooth in front of the lateral areas.

Petiolar suture partly obliterated. 1% gastral ster-

nite with fine transverse wrinkles, sometimes al-

most smooth medially. Thyridium elongate. Thy-

ridial index about 1.5. Epipleurum not separated

from the 3"4 gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellow. Man-

dible marked with yellow dorsally and medially.

Tegula yellow. Wings slightly yellowish. Stigma

yellowish brown medially. Front leg from the tip

of the coxa onwards yellow. Mid leg from the

trochanter onwards yellow. Hind tibia and tarsus

yellow. Mid and hind tarsi marked with brown

apically. Postpetiole sometimes marked with red

posteriorly. 2" gastral tergite yellowish red on the

posterior 0.8. 3’ and 4" tergites yellowish red.

Pilosity whitish.

d: Some parts of the mesopleurum densely

rugose-punctate or rugose. Longitudinal carinae

of the propodeum present medially and posteri-

orly, obliterated anteriorly. Mandible yellow with

black teeth. Tip of the mid coxa yellow. In other

characters similar to the 2.

Dusona constantineanui Hinz, 1977

Description. 2: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons scarcely depressed, coriaceous, distinct-

ly and densely punctate, with a fine median longi-

tudinal carina, without radial wrinkles. Antennal

carina low and narrow. Face and dorsal 0.7 of the

clypeus coriaceous, densely rugose-punctate and

rugose. Clypeus flat, finely coriaceous and dis-

tinctly and rather densely punctate ventrally, its

apex almost truncate medially. Malar space index

0.5. Genal carina low and almost straight ventral-

ly, joining the slightly but distinctly raised oral

carina at an acute angle. Genal index 0.5. Temple

finely coriaceous and with fine disperse puncta-

tion medially. Flagellum with 45-47 segments, fil-

iform, the apical 0.3 distinctly attenuate, 24 seg-

ment 2.2x, shortest segment 1.1x, penultimate seg-

ment 1.4x as long as wide. Pronotum densely

rugose-punctate on a smooth background lateral-

ly on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum coriaceous, very densely

punctate. Scutellum finely coriaceous, rather

densely punctate, with lateral carinae on the ante-

rior half. Prescutellar groove smooth medially,

with longitudinal striae laterally. Ventral part of

the prepectal carina merging with the transverse

carina, both carinae low. Pleural part of the pre-

pectal carina obliterated. Pleural part of the pre-

pectus densely punctate and rugose-punctate.

Mesopleurum with distinct and rather dense or

dense punctation on asmooth background. Specu-

lum punctate posteriorly, smooth anteriorly. De-

pression in front of the speculum with strong

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum very densely punc-

tate or densely rugose-punctate on a smooth back-

ground. Pleural carina complete. Areolet rather

large, short-stalked, receiving the 2”! recurrent

vein distad of the middle. Nervellus distinctly

reclivous, weakly intercepted. Nervellar indext

0.5-0.6. Tibial spur index 0.6. Hind femur 4.5x as

long as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.4 of the hind coxae, not depressed

anteriorly and posteriorly, slightly and narrowly

depressed medially, almost smooth in the midline

anteriorly and medially, with short transverse or

irregular wrinkles beside the shining midline, with

long transverse striae posteriorly, carinae almost

completely obliterated. Propodeal spiracle rather

large, 2.7x as long as wide. Spiracular carina nar-

row. Spiracular index 0.3. 1” gastral tergite 3.7x,

2" tergite 1.9x as long as wide. Petiole without

glymmae, with weakly indicated narrow lateral

areas, smooth in front of these areas. Petiolar su-

ture for the greater part obliterated. 1* gastral

sternite smooth. Thyridium elongate. Thyridial

index about 1.0. Epipleurum not separated from

the 3" gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellow medially. Flag-

ellum light brown medially and apically. Tegula

yellowish brown. Wings hyaline. Stigma yellow-

ish brown to medium brown medially. Front tro-

chanter sometimes marked with reddish brown.

Front femur yellowish red. Front tibia and tarsus

yellow, the tarsus marked with black apically.

Mid femur yellowish red on the apical 0.3. Mid

tibia yellow. Mid tarsus medium brown to dark

brown. Hind tibia medially yellow to yellowish

brown on the dorsal side, brown on the ventral

side, narrowly black basally, broadly black apical-

ly. 2" gastral tergite dorsally red on the posterior

0.1-0.2, laterally red on the posterior 0.4. 3" tergite

red, marked with black dorsally. 4" tergite red,

sometimes tinged with brown or black dorsally.

Pilosity whitish.

d: Flagellum medium brown medially and

apically. Front trochanter yellowish brown to red-

dish brown. Hind tibia medially broadly whitish

yellow on the dorsal side, light brown on the

ventral side, narrowly black basally, rather broad-

ly black apically. Genital clasper black. In other

characters similar to the 9.

Material (4??, 385): Russia: Yevreyskiy A.O.: Amur-

zet; Primorye: Dalnerechensk, Khasan (PET), Sikhote-

Alin (MOS), Yakovlevka (MOS, PET). China: Hei-

longjiang Prov.: Charbin (= Harbin) (AEI).

Dusona contumator Hinz, 1979

Description. 2: Body length 9-10 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons slightly depressed,

coriaceous, with very fine punctation, with a fine

median longitudinal carina, with short radial wrin-

kles. Antennal carina low and narrow, the rim

slightly bent upwards. Face and dorsal 0.6 of the

clypeus coriaceous, densely rugose-punctate.

Clypeus slightly convex, coriaceous and rather

densely punctate ventrally, its apex slightly con-

vex medially. Malar space index 0.4. Genal carina

low and straight ventrally, joining the low oral

carina at an acute angle. Genal index 0.3. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 45-51 segments,

filiform, the apical half attenuate, 2” segment 1.7x,

shortest segment 0.9x, penultimate segment 1.5x

as long as wide. Pronotum with longitudinal wrin-

kles, the dorsal 0.2 with additional punctures.

Mesoscutum and scutellum coriaceous, densely

punctate. Scutellum with lateral carinae on the

anterior 0.6. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter complete, joining the anterior

edge of the mesopleurum. Transverse carina ob-

literated. Mesopleurum distinctly coriaceous,

densely punctate. Speculum coriaceous and punc-

tate posteriorly, finely granulate-strigose anteri-

orly. Depression in front of the speculum granu-

late-strigose and with fine longitudinal striae,

which merge with the punctation or with a nar-

row coriaceous area without punctation anterior-

ly. Metapleurum coriaceous, very densely punc-

tate. Pleural carina distinct. Areolet rather large,

short-stalked, receiving the 2" recurrent vein at

or slightly distad of the middle. Nervellus re-

clivous, distinctly intercepted. Nervellar index 0.6.

Tibial spur index 0.6. Hind femur 4.5x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, broadly and distinctly depressed,

with short transverse wrinkles anteriorly, some-

times with a coriaceous area without wrinkles

anteriorly, with long transverse wrinkles posteri-

orly, carinae almost completely obliterated, some-

times the longitudinal carinae indicated medially

and posteriorly. Propodeal spiracle large, 3.4x as

long as wide. Spiracular carina widened. Spiracu-

lar index 0.2. 1“ gastral tergite 3.7x, 2" tergite 1.8x

as long as wide. Petiole without glymmae or with

very small glymmae, without narrow lateral are-

as, smooth in front of the glymmae. Petiolar su-

ture almost completely obliterated. 1“ gastral ster-

nite smooth. Thyridium oval. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter with somewhat indistinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible dark reddish brown medially. Flagel-

lum medium brown apically. Wings hyaline. Stig-

ma light brown medially. Front femur reddish

brown dorsally, black ventrally. Front tibia and

tarsus yellow. Mid femur yellow on the apical 0.1.

Mid tibia yellow. Hind tibia marked with whitish

yellow subbasally on the dorsal side. 2" gastral

tergite dorsally black or marked with red on the

posterior 0.2, the red marks more extended later-

ally. 3"! tergite red, sometimes tinged with brown.

Pilosity whitish.

d: Flagellum medium brown to dark brown

apically. In other characters similar to the ®.

Material (922, 7388): Russia: Amur Prov. (HEL, PET);

Yevreyskiy A.O.: Amurzet (PET); Primorye: Khasan

(Hinz/ZSM, MOS, PET). Japan: Hokkaido: Ohnuma

(AEN).

Dusona conturıax (Förster, 1868)

Description. 2: Body length 8-9 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 1.1. Frons slightly depressed, coriaceous,

finely and dispersely punctate, with a fine median

longitudinal carina, with short and indistinct ra-

dial wrinkles, the latter sometimes lacking. An-

tennal carina low and narrow. Face and dorsal

half of the clypeus coriaceous and densely punc-

tate. Clypeus flat, coriaceous and rather densely

punctate ventrally, its apex slightly convex medi-

ally. Malar space index 0.3. Genal carina low ven-

trally, slightly bent sidewards, joining the low

oral carina at an acute angle. Genal index 0.1-0.2.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 41-43 seg-

ments, filiform, the apical 0.2 attenuate, 2” seg-

ment 1.8x, shortest segment 0.9x, penultimate seg-

ment 1.0x as long as wide. Pronotum with punc-

tures and fine longitudinal wrinkles laterally on

the dorsal 0.2, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous, dull,

very densely punctate. Scutellum with lateral car-

inae on the anterior half. Prescutellar groove with

fine longitudinal striae. Prepectal carina low, the

ventral part merging with the pleural part, the

latter complete or almost complete, joining the

anterior edge of the mesopleurum or its dorsal 0.2

replaced by several wrinkles. Transverse carina

more or less obliterated. Mesopleurum with rath-

er dense or dense punctation on a coriaceous and

somewhat shining background. Speculum finely

coriaceous, shining. Depression in front of the

speculum with fine longitudinal wrinkles, which

merge with the punctation anteriorly. Metapleu-

rum coriaceous, finely and densely punctate, partly

rugose. Pleural carina distinct. Areolet large,

stalked, receiving the 2" recurrent vein basad of

the middle. Nervellus vertical, weakly intercept-

ed. Nervellar index 0.8. Tibial spur index 0.5. Hind

femur 4.6x as long as high. Hind tarsal claws

short, with long pectinate teeth basally and medi-

ally. Propodeum extending to 0.4 of the hind cox-

ae, rather broadly and deeply depressed, with

short transverse wrinkles in the midline of the

sulcus, in other parts irregularly wrinkled, cari-

nae almost completely obliterated. Propodeal spir-

acle large, 3.0x as long as wide. Spiracular carina

narrow. Spiracular index 0.3. 1* gastral tergite

3.9x, 2" tergite 1.3x as long as wide. Petiole with

distinct glymmae, which are bordered by longitu-

dinal carinae dorsally, smooth in front of the glym-

mae. Petiolar suture partly obliterated. 1* gastral

sternite coriaceous anteriorly, smooth medially,

transversely striate posteriorly. Thyridium oval.

Thyridial index about 1.0. Epipleurum separated

from the 3" gastral tergite anteriorly, the crease

marked with black. Epipleural index 0.5. Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.7.

Colour: Black. Maxillary palpus yellowish

brown. Mandible yellowish brown or dark brown

medially. Tegula brownish black. Wings hyaline.

Stigma yellowish medially. Front leg from the

femur onwards yellowish red. Mid femur yellow-

ish red on the apical 0.3. Mid tibia yellowish red.

Mid tarsus brown. Hind tibia broadly yellowish

brown medially, narrowly black basally, indis-

tinctly tinged with brown apically. Hind tarsus

dark brown. 2"! gastral tergite dorsally red on the

posterior 0.3-0.5, the red marks more extended

laterally. 3" tergite red. 4" tergite marked with

red anteriorly and/or ventrolaterally. Pilosity

whitish.

d: Propodeum not so broadly and not so deep-

ly depressed, longitudinal carinae sometimes

present medially and posteriorly. Maxillary pal-

pus yellowish. Mandible broadly yellow medial-

ly. Tegula yellow or yellowish brown. Front tro-

chanter yellowish red. Mid femur yellowish red.

Front and mid tibiae and tarsi yellow. 4'" gastral

tergite sometimes completely red. In other charac-

ters similar to the 9.

Material (722, 238): Russia: Irkutsk Prov.; Chita

Prov.: Klichka, Urulyunguy E of Borzya, Zabaykalsk

(PET).

Dusona coriacea Horstmann, spec. nov.

Holotype (3): Japan, Honshu, Norikura-dake,

2000 m, 30.7.1954 (AEI).

Description. d: Body length 13 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed

above the antennal carina, coriaceous, very finely

punctate, with a fine median longitudinal carina,

with fine and short radial wrinkles. Antennal car-

ina low and narrow. Face and dorsal 0.6 of the

clypeus coriaceous and very densely punctate.

Clypeus flat, coriaceous and rather densely punc-

tate ventrally, its apex truncate medially. Malar

space index 0.4. Genal carina low and straight

ventrally, joining the low oral carina at an acute

angle. Genal index 0.4. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 49 segments, filiform, the apical

0.2 slightly attenuate, 2" segment 1.7x, shortest

segment 1.0x, penultimate segment 1.2x as long

as wide. Pronotum coriaceous and densely rugose-

punctate laterally on the dorsal 0.2, with longitu-

dinal wrinkles ventrally. Mesoscutum coriaceous,

finely and densely punctate. Scutellum coriaceous,

finely and rather densely punctate, with lateral

carinae on the anterior 0.6. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter almost joining

the anterior edge of the mesopleurum. Transverse

carina partly indicated, low. Mesopleurum dis-

tinctly coriaceous, rather densely punctate. Specu-

lum and depression in front of the speculum co-

riaceous, dull, in small parts granulate-strigose.

Metapleurum coriaceous, finely and very densely

punctate. Pleural carina distinct. Areolet large,

long-stalked, receiving the 2" recurrent vein slight-

ly basad of the middle. Nervellus slightly inclivous,

distinctly intercepted. Nervellar index 0.8. Tibial

spur index 0.4. Hind femur 4.8x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to the basal 0.3 of

the hind coxae, distincly and rather narrowly de-

pressed, the depression coriaceous and without

wrinkles anteriorly, transversely wrinkled medi-

ally and posteriorly, costulae partly present, lon-

gitudinal carinae distinct medially and posterior-

ly, covered with wrinkles anteriorly. Propodeal

spiracle rather large, 3.6x as long as wide. Spirac-

ular carina indistinct and covered with wrinkles.

Spiracular index about 0.3. 1” gastral tergite 4.6x,

2"d tergite 2.7x as long as wide. Petiole with deep

and rather long glymmae, the latter bordered by a

carina dorsally, petiole smooth in front of the

glymmae. Petiolar suture fine, partly obliterated.

1* gastral sternite smooth. Thyridium elongate.

Thyridial index about 1.0. Epipleurum separated

from the 3"! gastral tergite, the crease slightly

marked with black. Epipleural index 0.7. Gaster

distinctly compressed posteriorly.

Colour: Black. Maxillary palpus brown basal-

ly, yellow apically. Mandible narrowly yellow dor-

sally and medially. Tegula yellowish brown. Wings

hyaline. Stigma yellowish brown medially. Front

trochanter and femur yellowish red. Front tibia

and tarsus yellow. Mid leg from the apical 0.2 of

the femur onwards yellow. Hind tibia dark yel-

lowish brown medially, broadly black basally, very

broadly black apically. 2" gastral tergite red on

the posterior 0.6. 3"! and 4" tergites red. 5" tergite

narrowly marked with red ventrolaterally. Geni-

tal clasper black. Pilosity whitish.

? unknown.

Dusona crassiventris Horstmann, spec. nov.

Holotype (2): North Korea, Wonsan, 29.8.1987

(SAW).

Description. 2: Body length 17 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons distinctly de-

pressed (but not funnel-like), coriaceous and

densely punctate dorsally, with strong radial wrin-

kles in the depression ventrally, these wrinkles

extending to half the distance to the median ocel-

lus, without a median longitudinal carina. Anten-

nal carina distinctly raised, its rim distinctly bent

upwards and widened to a crescent-shaped trans-

versely striate plate (ear-like) (Fig. 6). Face and

dorsal 0.8 of the clypeus coarsely and very dense-

ly punctate and rugose-punctate. Clypeus flat,

finely coriaceous and with rather fine punctures

and wrinkles ventrally, its apex truncate medial-

ly. Malar space index 0.3. Genal carina low ven-

trally, slightly bent sidewards, joining the slightly

raised oral carina at an acute angle. Genal index

0.4. Temple with distinct and disperse or rather

dense punctation on a finely coriaceous and part-

ly smooth background medially. Flagellum with

61 segments, filiform, the apical 0.4 distinctly at-

tenuate, 2"! segment 1.5x, shortest segment 1.0x,

penultimate segment 1.3x as long as wide. Prono-

tum with very dense punctation on a smooth back-

ground laterally on the dorsal 0.6, with wrinkles

ventrally. Mesoscutum and scutellum strongly and

densely punctate on a finely coriaceous back-

ground. Scutellum with lateral carinae on the an-

terior 0.3. Prescutellar groove with fine longitudi-

nal striae. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter almost joining

the anterior edge of the mesopleurum. Transverse

carina indicated anteriorly. Mesopleurum distinct-

ly and very densely punctate on a smooth back-

ground. Speculum slightly granulate-strigose,

shining. Depression in front of the speculum for

the greater part with rather long longitudinal stri-

ae, which merge partly with the punctation, part-

ly with a narrow area without punctation anteri-

orly. Metapleurum distinctly and very densely

punctate on a smooth background. Pleural carina

distinct. Areolet large, narrowly sessile, receiving

the 2" recurrent vein slightly basad of the middle.

Nervellus slightly inclivous, distinctly intercept-

ed. Nervellar index 0.75. Tibial spur index 0.5.

Hind femur 4.7x as long as high. Hind tarsal claws

short, with dense and rather long pectinate teeth

almost over their total length. Propodeum extend-

ing to 0.3 of the hind coxae, broadly and distinctly

but not deeply depressed, sulcus with long trans-

verse wrinkles posteriorly, short transverse wrin-

kles medially, coriaceous and punctate without

wrinkles anteriorly, costulae partly present, lon-

gitudinal carinae distinct, almost reaching the cos-

tulae. Propodeal spiracle large, 2.4x as long as

wide. Spiracular carina narrow. Spiracular index

0.3. 1% gastral tergite 2.4x, 2" tergite 1.7x as long

as wide. Petiole with large and deep glymmae,

smooth in front of the glymmae. Postpetiole 0.9x

as long as wide, conspicuously rounded laterally.

Petiolar suture fine, partly obliterated. 1“ gastral

sternite smooth. Thyridium oval. Thyridial index

about 1.0. Epipleurum not separated from the 3'“

gastral tergite, the latter without black lateral

stripes. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus brown basal-

ly, yellowish red medially and apically. Mandible

broadly yellow medially. Wings slightly tinged

with brown. Stigma dark yellowish brown medi-

ally. Front femur yellowish red dorsally and api-

cally. Front tibia and tarsus yellow. Mid femur

yellowish red on the apical 0.2. Mid tibia yellow-

ish red. Mid tarsus blackish. Hind tibia blackish,

tinged with dark brown medially. 2" gastral tergite

dorsally red on the posterior 0.1, almost complete-

ly red laterally, the posterior edge black sublater-

ally. 3"! and 4" tergite narrowly black dorsally,

broadly red laterally. Pilosity whitish.

d unknown.

Dusona cultrator (Gravenhorst, 1829)

Description. 2: Body length 12-16 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.1. Frons slightly depressed, coriaceous, finely

and densely punctate, with a distinct median lon-

gitudinal carina, with distinct radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina low and narrow. Face

and clypeus coriaceous, densely rugose-punctate.

Clypeus flat, its apex truncate medially. Malar

space index 0.4. Genal carina almost straight ven-

trally, slightly raised, joining the slightly raised

oral carina at an acute angle. Genal index 0.2-0.3.

Temple coriaceous and with fine disperse puncta-

tion medially. Flagellum with 53-58 segments,

filiform, the apical 0.3 distinctly attenuate, 2" seg-

ment 1.5x, shortest segment 0.9x, penultimate seg-

ment 1.4x as long as wide. Pronotum coriaceous

and densely rugose-punctate laterally on the dor-

sal 0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, dull, very dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.3. Prescutellar groove smooth medially,

with longitudinal striae laterally. Prepectal carina

low, the ventral part merging with the pleural

part, the latter complete, joining the anterior edge

of the mesopleurum, sometimes the dorsal 0.1

indistinct. Transverse carina obliterated. Meso-

pleurum finely coriaceous, with fine rather dense

or dense punctation. Speculum finely coriaceous

and granulate-strigose, shining. Depression in

front of the speculum with rather short longitudi-

nal striae, which merge with the punctation ante-

riorly. Metapleurum coriaceous, finely and densely

punctate. Pleural carina distinct. Areolet large,

pointed or short-stalked, receiving the 2” recur-

rent vein slightly basad of the middle. Nervellus

vertical, distinctly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 4.8x as long as

high. Hind tarsal claws rather long, with rather

long pectinate teeth almost over their total length.

Propodeum extending to 0.3 of the hind coxae,

broadly and deeply depressed, sulcus transverse-

ly wrinkled, costulae almost complete, longitudi-

nal carinae distinct posteriorly, sometimes indi-

cated medially. Propodeal spiracle rather large,

3.3x as long as wide. Spiracular carina narrow.

Spiracular index 0.5. 1 gastral tergite 3.4x, 24

tergite 1.2x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal car-

inae dorsally, smooth or with very fine longitudi-

nal striae in front of the glymmae. Petiolar suture

complete. 1“ gastral sternite with very fine sculp-

ture, shining, sometimes with transverse striae

posteriorly. Thyridium oval. Thyridial index about

1.5. Epipleurum separated from the 3" gastral

tergite, the crease marked with black. Epipleural

index 0.5-0.8. Gaster distinctly compressed poste-

riorly. Ovipositor index 0.7-0.8.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellowish red medially. Wings

hyaline. Stigma yellowish brown medially. Front

leg from the tip of the coxa onwards yellowish

red. Mid leg from the femur onwards yellowish

red, the tarsus sometimes darkened. Hind femur

red, sometimes marked with brown or black ba-

sally (to a variable extend). Hind tibia yellowish

brown, sometimes narrowly marked with brown

basally. Hind tarsus yellowish brown to brown.

2" gastralttergite red on the posterior 0.3. 3" tergite

red on the anterior half or completely red. Pilosity

whitish.

8: Propodeum with complete costulae, often

the longitudinal carinae present medially and pos-

teriorly. Tegula yellowish brown to dark brown.

Mid leg from the tip of the coxa onwards yellow-

ish red. Hind tarsus for the greater part yellowish

red, marked with dark brown apically. In other

characters similar to the 2.

Material (1622, 438): Russia: Tomsk Prov.: Tomsk;

Irkutsk Prov.: Irkutsk; Chita Prov.: Kalga; Khabarovk

Terr.: Khekhtsir near Khabarovsk, Troitskoye; Primo-

rye: Pogranichnyy, Roshchino, Vladivostok. Kaza-

khstan: Akmolinskiy, Chimkent, Yanvartsevo (PET).

Kirgiziya: Chatkal (MOS), River Tyup (LIN). Japan:

Hokkaido: Sapporo (SAP); Honshu: Kamikochi near

Norikura-dake (AEI).

Dusona detritor Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Kuril Isl., Kunashir,

Mendeleyevo, 2.8.1981 (PET). — Paratypes: 18

Russia, Kuril Isl., Kunashir, Yuzhno-Kurilsk,

11.7.1973 (PET); 15 Japan, Honshu, Kamikochi,

near Norikura-dake, 23.7.1954 (AEI).

Description. ?: Body length 9-10 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed,

coriaceous, finely punctate, with a very fine medi-

an longitudinal carina, with short radial wrinkles.

Antennal carina low and narrow. Face and dorsal

half of the clypeus densely rugose-punctate.

Clypeus flat, finely coriaceous and with fine rath-

er dense punctation ventrally, its apex slightly

convex medially. Malar space index 0.4. Genal

carina low and rather straight ventrally, joining

the slightly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 43-44 segments, filiform, the apical 0.2 atten-

uate, 2“ segment 2.3x, shortest segment 1.0x, pe-

nultimate segment 1.3x as long as wide. Prono-

tum with dense punctation laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, very densely

punctate. Scutellum with lateral carinae on the

anterior half. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum finely coriaceous, distinctly and

densely punctate. Speculum finely coriaceous,

shining. Depression in front of the speculum with

distinct longitudinal stiae, which merge with the

punctation anteriorly. Metapleurum densely ru-

gose. Pleural carina distinct. Areolet rather small,

short-stalked, receiving the 2" recurrent vein dis-

tad of the middle. Nervellus vertical, intercepted.

Nervellar index 0.6. Tibial spur index 0.5. Hind

femur 6.2x as long as high. Hind tarsal claws

rather long, with distinct pectinate teeth almost

over their total length. Propodeum extending to

the middle of the hind coxae, scarcely depressed,

slightly rugose-punctate and with some fine trans-

verse wrinkles, Costulae complete, longitudinal

carinae obliterated. Propodeal spiracle large, 4.3x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.3. 1% gastral tergite 5.4x, 2"d tergite

1.7x as long as wide (but the latter slightly de-

formed in the only available 2). Petiole with small

and shallow glymmae at the anterior edge of in-

distinct narrow lateral areas, smooth in front of

the glymmae. Petiolar suture very fine. 1% gastral

sternite smooth. Thyridium oval. Thyridial index

about 3.0. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.6. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly marked with yellow medially. Tegu-

la dark brown. Wings hyaline. Stigma dark brown

medially. Front leg from the tip of the coxa on-

wards yellow to yellowish red. Mid leg from the

apical 0.2 of the femur onwards yellow. Hind tibia

dark yellowish brown, distinctly marked with

black basally. 2" gastral tergite marked with red

laterally. 3° gastral tergite black anteriorly, red

posteriorly. Posterior tergites marked with black

dorsally, broadly red laterally. Posterior sternites

red. Pilosity whitish.

d: Glymma partly obliterated. Petiolar suture

distinct. Tegula yellowish brown to brown. Mid

femur yellowish on the apical 0.4. Genital clasper

red. In other characters similar to the 2.

Dusona dictor Hinz, 1979

Description. 2: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons slightly depressed, coriaceous and fine-

ly rugose, with a fine median longitudinal carina,

with short radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.8 of the clypeus

finely coriaceous and densely rugose-punctate.

Clypeus slightly convex, coriaceous and with rath-

er dense punctation ventrally, its apex truncate

medially. Malar space index 0.3. Genal carina

raised close to the junction with the also raised

oral carina, forming a lopsided triangular pyra-

mid, the tip of the latter shifted ventrally and

situated behind the base of the mandible. Gena

depressed close to the junction, the depression

about half as deep as the width of the 2"4 flagellar

segment. Genal index about 0.5. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 37-39 segments, filiform,

the apical 0.2 slightly attenuate, 2” segment 1.5x,

shortest segment 0.8x, penultimate segment 0.9x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, very densely punctate. Scutellum with

lateral carinae on the anterior 0.6. Prescutellar

groove smooth medially, with few longitudinal

striae laterally. Prepectal carina low, ventral part

merging with the pleural part, the dorsal half of

the latter obliterated. Transverse carina partly

present. Mesopleurum distinctly coriaceous, dull,

densely punctate. Speculum coriaceous, dull. De-

pression in front of the speculum with fine rather

long longitudinal striae, which merge with the

punctation anteriorly. Metapleurum finely and

densely rugose and rugose-punctate, the wrinkles

coriaceous. Pleural carina distinct. Areolet rather

small, short-stalked, receiving the 2"! recurrent

vein distad of the middle. Nervellus slightly in-

clivous, distinctly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 5.2x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.4 of the

hind coxae, rather broadly and not deeply de-

pressed, sulcus for the greater part irregularly

wrinkled, with long transverse wrinkles posteri-

orly, costulae indistinct, longitudinal carinae indi-

cated medially and posteriorly. Propodeal spira-

cle rather large, 3.2x as long as wide. Spiracular

carina narrow. Spiracular index 0.4. 1” gastral

tergite 3.6x, 2X tergite 1.6x as long as wide. Peti-

ole with distinct glymmae, which are bordered by

longitudinal carinae dorsally, smooth in front of

the glymmae. Petiolar suture for the greater part

obliterated. 1“ gastral sternite for the greater part

smooth, with few transverse striae posteriorly.

Thyridium elongate. Thyridial index about 1.0.

Epipleurum not separated from the 3% gastral

tergite, 3° tergite without black lateral stripes.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.8.

Colour: Black. Maxillary palpus brown basal-

ly, yellowish red medially and apically. Mandible

narrowly tinged with dark brown subapically.

Tegula medium brown. Wings hyaline. Stigma

medium brown medially. Front femur black ven-

trally and basally, yellowish red dorsally and api-

cally. Front tibia and tarsus yellow. Mid femur

narrowly marked with yellow apically. Mid tibia

yellow. Mid tarsus yellowish brown. Hind tibia

light yellow medially, narrowly black basally,

broadly black apically. Hind tarsus dark brown.

2"! gastral tergite dorsally red on the posterior 0.2,

laterally red on the posterior 0.4. 3" tergite red. 4"

tergite marked with red anteriorly. Pilosity whit-

ish.

d unknown.

Material (3??): Russia: Yevreyskiy A.O.: Amurzet;

Kuril Isl.: Dubovoye on Kunashir (PET). Japan: Hok-

kaido: Jozankei (SAP).

Dusona falcator (Fabricius, 1775)

Description. 2: Body length 18-20 mm. Temples

slightly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous,

densely rugose-punctate, without a median longi-

tudinal carina, with distinct radial wrinkles, which

extend to 0.3 the distance to the median ocellus.

Antennal carina low and narrow, the rim slightly

and indistinctly bent upwards. Face and dorsal

0.8 of the clypeus finely coriaceous, coarsely and

densely rugose-punctate. Clypeus flat, finely co-

riaceous and rather densely punctate ventrally, its

apex truncate medially. Malar space index 0.3.

Genal carina straight ventrally, distinctly raised,

joining the distinctly raised oral carina at an acute

angle, the junction of both carinae not shifted

ventrally. Gena slightly depressed close to that

junction. Genal index 0.4. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 65-67 segments, filiform,

the apical 0.3 distinctly attenuate, 2"’ segment

1.6x, shortest segment 1.0x, penultimate segment

1.5x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.4, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, rather densely or densely punc-

tate. Scutellum with lateral carinae on the anterior

0.8. Prescutellar groove with fine longitudinal stri-

ae. Prepectal carina low, ventral part merging with

the pleural part, the latter complete, joining the

anterior edge of the mesopleurum. Transverse

carina complete. Rarely the ventral part of the

prepectal carina merging with the transverse car-

ina, and the pleural part somewhat lower. Meso-

pleurum finely coriaceous, strongly and densely

punctate. Speculum finely coriaceous, granulate-

strigose, shining. Depression in front of the specu-

lum with rather long longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum densely rugose, the wrinkles coriaceous. Pleu-

ral carina complete. Areolet large, short-stalked,

receiving the 2" recurrent vein slightly basad of

the middle. Nervellus distinctly inclivous, dis-

tinctly intercepted. Nervellar index 0.7. Tibial spur

index 0.5. Hind femur 5.8x as long as high. Hind

tarsal claws rather long, with very long pectinate

teeth almost over their total length. Propodeum

extending to 0.4 of the hind coxae, broadly and

not deeply depressed, sulcus almost completely

transversely wrinkled, costulae partly present,

longitudinal carinae complete or almost complete.

Propodeal spiracle large, 3.3x as long as wide.

Spiracular carina narrow. Spiracular index 0.4.

1" gastral tergite 4.5x, 2"X tergite 2.5x as long as

wide. Petiole with rather large glymmae, smooth

in front of the glymmae. Petiolar suture almost

completely obliterated. 1* gastral sternite smooth.

Thyridium oval. Thyridial index 1.3. Epipleurum

not separated from the 3" gastral tergite, the latter

without black lateral stripes. Gaster strongly com-

pressed posteriorly. Ovipositor index 0.4.

Colour: Black. Maxillary palpus dark brown

or black. Mandible marked with reddish brown

medially. Tegula yellowish brown. Wings tinged

with yellow. Stigma yellowish brown medially.

Front trochanter and femur yellowish red. Front

tibia and tarsus yellow. Sometimes also the front

coxa marked with yellow apically. Mid femur

marked with yellow on the apical 0.3. Mid tibia

yellow. Mid tarsus dark brown. Hind femur nar-

rowly marked with yellow apically. Hind tibia

completely yellow. Hind tarsus brownish black.

2" gastral tergite dorsally yellow on the posterior

0.2, almost completely yellow laterally. 3 and 4"

tergites yellow. 5" tergite marked with yellow

ventrolaterally. Pilosity whitish.

d: Similar to the 2.

Material (4??, 15): Russia: Altaysk Terr. (MIN). Ka-

zakhstan: Borovoye near Kokchetav (PET).

Dusona flagellator (Fabricius, 1793)

This species is very similar to D. inermis, particu-

larly in the shape of the genal carina. It differs

from the latter in: Hind tibia less extensively

marked with yellow. 2” gastral tergite less exten-

sively marked with red.

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons not depressed, densely ru-

gose, with a distinct median longitudinal carina,

with distinct radial wrinkles, which extend to half

the distance to the median ocellus. Antennal cari-

na low and narrow. Face and dorsal 0.3 of the

clypeus coriaceous, densely rugose-punctate and

rugose. Clypeus flat, coriaceous and densely punc-

tate ventrally, its apex slightly convex medially.

Malar space index 0.4. Genal carina slightly but

distinctly raised ventrally, distinctly bent forwards

and sidewards, joining the oral carina at the base

of the mandible at a right angle. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 42-47 segments, filiform,

the apical 0.2 attenuate, 2”! segment 2.4x, shortest

segment 1.3x, penultimate segment 1.7x as long

as wide. Pronotum coriaceous and densely rugose

laterally on the dorsal 0.2, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, dull, very densely punctate. Scutellum with

lateral carinae on the anterior 0.7. Prescutellar

groove smooth. Ventral part of the prepectal car-

ina merging with the transverse carina, both cari-

nae strongly raised, the former deeply emarginate

medially. Pleural part of the prepectal carina low-

er, its dorsal 0.2 replaced by several wrinkles.

Mesopleurum distinctly coriaceous, with rather

dense or dense (rarely disperse) punctation. Specu-

lum with fine oblique striae or granulate-strigose,

rather dull. Depression in front of the speculum

with distinct longitudinal striae, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, finely and densely rugose-punctate. Pleu-

ral carina distinct. Areolet rather large, short-

stalked, receiving the 2” recurrent vein almost at

the middle. Nervellus distinctly reclivous, not in-

tercepted or weakly intercepted. In the latter case

nervellar index 0.7. Tibial spur index 0.6. Hind

femur 5.1x as long as high. Hind tarsal claws

short, with short pectinate teeth basally. Propo-

deum extending to the middle of the hind coxae,

broadly and deeply depressed, sulcus with short

transverse wrinkles in the midline, irregularly

wrinkled in other parts, costulae obliterated, lon-

gitudinal carinae almost completely obliterated or

slightly indicated medially and posteriorly. Pro-

podeal spiracle rather small, 3.6x as long as wide.

Spiracular carina narrow or partly widened. Spirac-

ular index 0.3. 1% gastral tergite 4.5x, 2" tergite

1.8x as long as wide. Petiole with small glymmae

at the anterior edge of narrow lateral areas, glym-

mae sometimes obliterated, petiole smooth in front

of these areas. Petiolar suture obliterated. 1* gas-

tral sternite smooth. Thyridium oval. Thyridial

index about 1.5. Epipleurum not separated from

the 3'4 gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.7.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with reddish

brown dorsally. Wings hyaline. Stigma yellowish

medially. Front leg from the femur onwards yel-

lowish red. Apical half of mid femur and mid tibia

yellowish red. Hind tibia black, sometimes tinged

with yellowish brown subbasally on the dorsal

side. 2" gastral tergite red on the posterior 0.3-0.4.

34 and 4" tergites red, the 4" tergite sometimes

marked with black dorsally. Pilosity whitish.

d: Maxillary palpus yellow. Mandible broad-

ly yellow medially. Tegula yellow. Front and mid

legs from the tip of the coxa onwards yellowish

red, sometimes the mid coxa completely black.

Hind tibia whitish yellow medially, narrowly black

basally, very broadly black apically (apical black

ring extending over 0.3 of the tibia). In other char-

acters similar to the 9.

Material (1477, 933): Russia: Altaysk Terr.: Gorno-

Altaysk (MIN); Yakutsk A.S.S.R.: Bestyak; Amur

Prov.: Svobodnyy; Primorye: Spassk Dalniy; Kam-

chatka: Kozyrevsk (PET). Kazakhstan: Alma-Ata,

Chimkent (MOS, PET). Kirgiziya: Kirgizskiy Mtes.

(LIN).

Variety: Body length 7 mm. Genal carina low ven-

trally. Flagellum with 39-41 segments. Material:

12,18 Kazakhstan: Aksu-Dzhabagly, ESE of Chim-

kent (MOS, Hinz/ZSM). Probably these are small

specimens of D. flagellator.

Dusona flavitarsis Horstmann, spec. nov.

The species is similar to D. rossica Hinz (described

from European Russia). The latter differs in: Body

length 13-14 mm. 56 flagellar segments. 4" gastral

tergite red.

Types. Holotype (3): Mongoliya, Sühbaatar Prov.,

Shiliin-Bogd uul Mts., 10.7.1971 (PET). — Para-

types: 13 same locality and date (Hinz/ZSM);

13 Russia, Chita Prov., Zabaykalsk, 21.6.1925

(PET).

Description. d: Body length 10-12 mm. Temples

not much constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the 1* gastral tergite. Ocellar

index 1.2. Frons slightly depressed, densely ru-

gose-punctate, with a fine and partly obliterated

median longitudinal carina, with fine radial wrin-

kles, which extend to 0.3 the distance to the medi-

an ocellus. Antennal carina low and narrow. Face

and dorsal 0.7 of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely punctate ventrally, its apex slightly

convex medially. Malar space index 0.5. Genal

carina low and straight ventrally, joining the slight-

ly raised oral carina at an acute angle. Genal index

0.5. Temple finely coriaceous and with fine rather

dense punctation medially. Flagellum with 48-51

segments, filiform, the apical 0.3 attenuate, 2"

segment 1.5x, shortest segment 0.9x, penultimate

segment 1.1x as long as wide. Pronotum very

densely punctate on a smooth background later-

ally on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum finely coriaceous, very

densely punctate. Scutellum very densely punc-

tate on a smooth background, with lateral carinae

on the anterior 0.4. Prescutellar groove with longi-

tudinal striae. Ventral part of the prepectal carina

low, merging with either the pleural part or the

transverse carina, pleural part almost joining the

anterior edge of the mesopleurum. Transverse

carina complete or obliterated posteriorly. Meso-

pleurum finely coriaceous, rather densely or dense-

ly punctate. Speculum slightly granulate-strigose

anteriorly, distinctly punctate posteriorly. Depres-

sion in front of the speculum granulate-strigose

and with fine longitudinal striae, which merge

with the punctation anteriorly. Metapleurum finely

coriaceous and densely punctate. Pleural carina

distinct. Areolet rather large, short-stalked or

pointed, receiving the 2” recurrent vein at the

middle. Nervellus vertical, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 4.3x as long as high. Hind tarsal claws long,

with short pectinate teeth basally. Propodeum

extending to 0.3 of the hind coxae, broadly and

not deeply depressed, transversely wrinkled in

almost all parts of the sulcus, costulae and longi-

tudinal carinae indicated, the latter obliterated

anteriorly. Propodeal spiracle large, 2.9x as long

as wide. Spiracular carina narrow. Spiracular in-

dex 0.5. 1* gastral tergite 3.1x, 2" tergite 1.5x as

long as wide. Petiole with rather large glymmae,

smooth in front of the glymmae. Petiolar suture

fine and partly obliterated. 1“ gastral sternite

smooth. Thyridium oval. Thyridial index about

1.0. Epipleurum separated from the 3% gastral

tergite anteriorly, the crease marked with black.

Epipleural index 0.4-0.5. Gaster distinctly com-

pressed posteriorly.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly marked with yellow medially.

Tegula yellowish brown. Wings hyaline. Stigma

light yellowish brown. Front leg from the tip of

the coxa onwards light yellowish red. Mid leg

from the femur onwards light yellowish red. Hind

tibia and hind tarsus light yellowish red, the tibia

marked with light red apically. 2n‘ gastral tergite

red on the posterior 0.2-0.4, the red marks more

extended laterally. 3° tergite red. 4" tergite nar-

rowly red anteriorly.

? unknown.

Dusona flavopicta Horstmann, spec. nov.

Types. Holotype (S): Russia, Kuril Isl., Kunashir,

Tretyakovo, 13.6.1968 (PET). — Paratypes: 18 Rus-

sia, Primorye, Pozhiga SO of Dalnerechensk,

20.7.1983 (PET); 439 Japan, Hokkaido, Sounkyo,

18.6.1938 (AEI, 18 Hinz/ZSM); 18 Japan, Shiko-

ku, Mt. Ishizuchi, 14.7.1933 (SAP).

Description. d: Body length 7-8 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.9. Frons slightly depressed, coriaceous,

finely rugose-punctate, without a distinct median

longitudinal carina, without radial wrinkles. An-

tennal carina low and narrow. Face and dorsal 0.7

of the clypeus densely and finely rugose-punc-

tate. Clypeus slightly concave, coriaceous and rath-

er densely punctate ventrally, its apex truncate

medially. Malar space index 0.4. Genal carina low

ventrally, straight or slightly bent sidewards, join-

ing the slightly raised oral carina at an acute an-

gle. Genal index 0.1-0.25. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 40-44 segments, filiform, the api-

cal 0.7 attenuate, 2" segment 1.4x, shortest seg-

ment 1.0x, penultimate segment 1.3x as long as

wide. Pronotum densely and finely rugose-punc-

tate laterally on the dorsal 0.2, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous and densely punctate. Scutellum with

lateral carinae on the anterior 0.8. Prescutellar

groove smooth medially, with fine longitudinal

striae laterally. Prepectal carina low, ventral part

merging with the pleural part, the latter covered

with wrinkles on its dorsal 0.3. Transverse carina

indicated anteriorly. Mesopleurum coriaceous,

dull, distinctly and densely punctate. Speculum

granulate-strigose, somewhat shining. Depression

in front of the speculum with fine and long longi-

tudinal wrinkles, which merge with the puncta-

tion anteriorly. Metapleurum densely and finely

rugose. Pleural carina complete. Areolet rather

small, long-stalked, receiving the 2" recurrent vein

distad of the middle. Nervellus vertical, weakly

intercepted. Nervellar index 0.8-0.9. Tibial spur

index 0.6. Hind femur 4.8x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to 0.3 of the hind

coxae, slightly and rather narrowly depressed,

costulae partly present, longitudinal carinae indi-

cated medially and posteriorly, the area between

the longitudinal carinae almost completely trans-

versely wrinkled. Propodeal spiracle rather small,

3.6x as long as wide. Spiracular carina narrow.

Spiracular index 0.5. 1* gastral tergite 4.5x, 2"4

tergite 2.0x as long as wide. Petiole with small

glymmae, these sometimes obliterated, smooth in

front of the glymmae. Petiolar suture fine. 1% gas-

tral sternite smooth anteriorly, with fine trans-

verse striae posteriorly. Thyridium oval. Thyridi-

al index about 2.0. Epipleurum not separated from

the 3" gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible yellow with black teeth. Tegula yellow.

Wings hyaline. Stigma light yellowish brown me-

dially. Front and mid legs for the greater part

yellow, coxae narrowly marked with black basal-

ly, femora yellowish red. Hind tibia yellowish

brown, narrowly dark brown basally, narrowly

marked with brown apically. Hind tarsus medi-

um brown to dark brown. 2” gastral tergite red

on the posterior 0.1-0.2, sometimes the red marks

more extended laterally. 3 and 4" tergite red,

partly marked with black dorsally. Pilosity whit-

ish, face with very dense pilosity.

? unknown.

Dusona fundator (Hinz, 1990)

This species was mixed up with D. stygia by Hinz

(1990: 262 ff.), one of the paratypes of D. fundator

from Escorial/Spain (Hinz/ZSM) belonging to

D. stygia. Both species differ from each other main-

ly in the sculpture of the propodeum (irregularly

wrinkled in D. fundator, distinctly transversely

wrinkled in D. stygia).

Description. @: Body length 9-10 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons scarcely depressed, coriaceous, fine-

ly and densely rugose-punctate, without or with

an incomplete median longitudinal carina, with

some radial wrinkles, which extend to half the

distance to the median ocellus. Antennal carina

low, its rim widened to a narrow and sometimes

indistinct crescent-shaped plate dorsally, the lat-

ter with fine transverse striae. Face and dorsal 0.4

of the clypeus coriaceous, densely rugose-punc-

tate. Clypeus flat, finely coriaceous and with rath-

er dense punctation ventrally, its apex slightly

convex medially. Malar space index 0.3. Genal

carina low and straight ventrally, joining the slight-

ly raised oral carina at an acute angle. Genal index

0.5. Temple finely coriaceous and with fine rather

dense punctation medially. Flagellum with 48-52

segments, filiform, the apical 0.3 attenuate, 2"

segment 1.4x, shortest segment 0.9x, penultimate

segment 1.1x as long as wide. Pronotum coria-

ceous and densely rugose-punctate laterally on

the dorsal 0.4, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous, very

densely punctate. Scutellum with lateral carinae

on the anterior 0.6. Prescutellar groove smooth

medially, with fine longitudinal striae laterally.

Prepectal carina low, ventral part merging with

the pleural part, the dorsal 0.2-0.4 of the latter

obliterated and replaced by wrinkles. Transverse

carina obliterated. Mesopleurum densely or very

densely punctate on a finely coriaceous and shin-

ing background. Speculum punctate posteriorly,

finely coriaceous or granulate-strigose and shin-

ing anteriorly. Depression in front of the specu-

lum with longitudinal wrinkles and fine striae,

which merge with the punctation anteriorly. Meta-

pleurum densely rugose and rugose-punctate.

Pleural carina distinct. Areolet rather large, point-

ed, receiving the 2" recurrent vein basad of the

middle. Nervellus slightly reclivous, weakly in-

tercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 4.9x as long as high. Hind tarsal

claws short, with short pectinate teeth basally and

medially. Propodeum extending to 0.3 of the hind

coxae, shallowly or not at all depressed, irregular-

ly wrinkled, with a short median longitudinal

keel and/or some longitudinal wrinkles posteri-

orly, carinae almost completely obliterated. Pro-

podeal spiracle large, 3.3x as long as wide. Spirac-

ular carina narrow, sometimes indistinct. Spiracu-

lar index 0.5-0.7. 1% gastral tergite 3.3x, 2" tergite

1.6x as long as wide. Petiole with rather large or

sometimes with small glymmae, wich are bor-

dered by longitudinal carinae dorsally, smooth in

front of the glymmae. Petiolar suture fine, partly

obliterated. 1“ gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum of the 3" gastral tergite present as asmall and

sometimes indistinct triangular lobe anteriorly,

the latter pendent and not marked with black, the

tergite with distinct black lateral stripes well above

the ventrolateral edge. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Wings hyaline. Stigma brown

medially. Front femur sometimes marked with

red dorsally. Front tibia brownish. Mid tibia some-

times narrowly marked with yellowish brown

basally. 2" gastral tergite red on the posterior 0.2-

0.3. 3" tergite red. 4" tergite marked with red

anteriorly or completely red. Pilosity whitish.

d: Mandible narrowly marked with yellow

dorsally. In other characters similar to the 2.

Material (1?, 13): Russia: Khabarovsk Terr.: Vys-

okogornyy (PET). Japan: Hokkaido: Tokachi-dake

(SAP).

Dusona gastator Hinz, 1985

Description. 2: Body length 7-8mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.2. Frons scarcely depressed, coriaceous and fine-

ly rugose-punctate, with a very fine median longi-

tudinal carina, without radial wrinkles. Antennal

carina low and narrow. Face coriaceous, densely

rugose and rugose-punctate. Clypeus slightly con-

vex, coriaceous, finely and densely punctate, its

apex convex medially. Malar space index 0.8. Ge-

nal carina obliterated ventrally, not joining the

oral carina, the latter slightly raised. Temple fine-

ly coriaceous and with fine disperse punctation

medially. Flagellum with 35-36 segments, filiform,

the apical 0.2 attenuate, 2"! segment 1.7x, shortest

segment 1.1x, penultimate segment 1.6x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous

and very densely punctate. Scutellum with lateral

carinae on the anterior half. Prescutellar groove

with distinct longitudinal striae. Prepectal carina

low, ventral part merging with the pleural part,

the latter complete, joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum coriaceous, densely rugose and ru-

gose-punctate. Speculum and depression in front

of the speculum with long longitudinal striae,

which merge with the punctation anteriorly. Meta-

pleurum finely and densely rugose. Pleural carina

distinct. Areolet small, stalked, receiving the 2"‘

recurrent vein slightly distad of the middle. Nervel-

lus slightly reclivous, not intercepted. Tibial spur

index 0.6. Hind femur 4.4x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to 0.3 of the hind

coxae, shallowly and rather narrowly depressed,

with short transverse wrinkles in the longitudinal

furrow, irregularly wrinkled beside the furrow,

area basalis and area superomedia bordered by

carinae, the latter not bordered by a carina poste-

riorly, costulae partly present, longitudinal cari-

nae present anteriorly and medially, partly indis-

tinct or obliterated posteriorly (Fig. 31). Propode-

al spiracle small, 1.7x as long as wide. Spiracular

carina somewhat widened. Spiracular index 0.8.

1% gastral tergite 2.8x, 2"X tergite 1.5x as long as

wide. Petiole without glymmae, with indistinct

narrow lateral areas, smooth in front of these are-

as. Petiolar suture distinct. 1" gastral sternite

smooth. Thyridium almost circular. Thyridial in-

dex about 2.0. Epipleurum separated from the 3"«

gastral tergite, the crease marked with black. Epi-

pleural index 0.9. Gaster slightly compressed pos-

teriorly. Ovipositor index 0.5.

Colour: Black. Wings hyaline. Stigma dark

brown medially. Front femur reddish brown on

the apical 0.7. Front tibia and tarsus reddish brown.

Mid tibia reddish brown. Mid tarsus brown. Hind

tibia reddish brown, narrowly black basally and

apically. 2” gastral tergite red on the posterior 0.6.

3" and 4"tergites red. 5'" tergite sometimes marked

with red anteriorly. Pilosity whitish.

d unknown.

Material (5??): Russia: Irkutsk Prov.: Irkutsk (Hinz /

ZSM); Kamchatka: Milkovo, Kozyrevsk. Mongolia:

Hövsgöl Prov.: Tosontsegal (PET).

Dusona gastroides Horstmann, spec. nov.

Holotype (JS): Japan, Honshu, Kamikochi, near

Norikura-dake, 24.7.1954 (AEI).

Description. d: Body length 11 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.8. Frons slightly depressed

above the antennal carinae, coriaceous, finely and

dispersely punctate, with a fine median longitudi-

nal carina on top of a slightly raised bulge, with

short radial wrinkles. Antennal carina low, dor-

sally narrow, medially widened to a crescent-

shaped plate. Face and dorsal half of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low and straight ventrally, joining

the slightly raised oral carina at an acute angle.

Genal index 0.5. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with at least 42 segments (the apical segments

broken off in the only available specimen), fili-

form, the apical 0.3 attenuate, 2"! segment 1.9x,

shortest segment 1.0x as long as wide. Pronotum

coriaceous and densely rugose-punctate laterally

on the dorsal half, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous, dull,

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove for the

greater part smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

finely coriaceous, dispersely punctate. Speculum

coriaceous and granulate-strigose, shining. De-

pression in front of the speculum with fine long

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum finely coriaceous and

dispersely punctate centrally, rugose in other parts.

Pleural carina complete. Areolet large, pointed,

receiving the 2” recurrent vein at the middle.

Nervellus reclivous, intercepted. Nervellar index

0.6. Tibial spur index 0.6. Hind femur 5.3x as long

as high. Hind tarsal claws rather long, with short

pectinate teeth basally. Propodeum extending to

the middle of the hind coxae, scarcely depressed,

with complete costulae and almost complete lon-

gitudinal carinae, the rather narrow area between

the longitudinal carinae for the greater part irreg-

ularly wrinkled, with some longitudinal wrinkles

posteriorly. Propodeal spiracle rather large, 3.0x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.5. 1% gastral tergite 5.1x, 2" tergite

3.4x as long as wide. Petiole with rather large

glymmae, which are bordered by longitudinal

carinae dorsally, with very fine longitudinal striae

in front of the glymmae. Petiolar suture partly

obliterated. 1" gastral sternite partly coriaceous.

Thyridium elongate. Thyridial index about 2.0.

Epipleurum separated from the 3" gastral tergite,

the crease marked with black. Epipleural index

0.8. Gaster distinctly compressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly light yellow medially. Wings hya-

line. Stigma medium brown medially. Front tro-

chanter and femur yellowish red. Mid femur yel-

lowish red on the apical 0.2. Front and mid tibiae

and tarsi yellow. Hind tibia yellowish brown me-

dially, narrowly black basally, broadly black api-

cally. 2"! gastral tergite red on the posterior 0.3.

Tergites and sternites from the 3" segment on-

wards red. Genital clasper red. Pilosity whitish.

? unknown.

Dusona genator Hinz, 1979

The type material of this species is not homoge-

nous: The types from southern China (including

the holotype) and from Japan are described in

detail below. One paratype from Khasan/ Primo-

rye/Russia (MOS) belongs to D. petiolatoides. One

paratype from River Amba-bira/ Primorye/Rus-

sia (PET) differs from the holotype of D. genator in

some details and is mentioned as a variety below.

Description. 2: Body length 10-12 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons not depressed, coria-

ceous, finely rugose, with distinct radial wrinkles,

which extend to 0.3 the distance to the median

ocellus, without a median longitudinal carina.

Antennal sockets large and close to each other.

Antennal carina low and narrow. Face and dorsal

0.8 of the clypeus densely rugose and rugose-

punctate. Clypeus slightly convex, scarcely coria-

ceous and rather densely punctate ventrally, its

apex truncate or slightly convex medially. Malar

space index 0.3. Genal carina slightly raised ven-

trally, straight or slightly bent forwards and side-

wards, joining the oral carina at the base of the

mandible at an acute angle. Temple finely coria-

ceous and with fine rather dense or dense puncta-

tion medially. Flagellum with 55-56 segments, fil-

iform, the apical 0.3 attenuate, 2" segment 1.7x,

shortest segment 1.1x, penultimate segment 1.2x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

finely coriaceous, very densely punctate. Scutel-

lum with lateral carinae on the anterior 0.4. Pres-

cutellar groove smooth medially, with fine longi-

tudinal striae laterally. Ventral part of the prepec-

tal carina merging with the transverse carina, both

carinae distinctly raised, the former scarcely emar-

ginate medially, the rim of the latter bent up-

wards. Median 0.3 of the pleural part of the pre-

pectal carina present, the pleural part obliterated

or replaced by several wrinkles dorsally and ven-

trally. Pleural part of the prepectus slightly de-

pressed and with strong dorsoventral wrinkles.

Mesopleurum very densely punctate on a smooth

background, partly with wrinkles, especially in

the lower posterior corner. Speculum very finely

coriaceous, almost smooth, shining. Depression

in front of the speculum with distinct longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum densely rugose-punctate or ru-

gose. Pleural carina distinct. Areolet rather large,

pointed, receiving the 2" recurrent vein almost at

the middle. Nervellus almost vertical, not inter-

cepted or weakly intercepted. In the latter case

nervellar index 0.7-0.8. Tibial spur index 0.7. Hind

femur 5,6x as long as high. Hind tarsal claws

rather short, with few distinct pectinate teeth ba-

sally and medially. Propodeum extending to 0.4

of the hind coxae, not or almost not depressed, for

the greater part irregularly wrinkled, often with

transverse wrinkles posteriorly, costulae almost

complete, longitudinal carinae present only pos-

teriorly. Propodeal spiracle rather large, 3.4x as

long as wide. Spiracular carina fine and narrow or

indistinct. Spiracular index 0.5-0.7. 1" gastral tergite

5.3x, 2"d tergite 3.5x as long as wide. Petiole with-

out glymmae, sometimes with indistinct narrow

lateral areas, smooth laterally. Petiolar suture part-

Iy obliterated. 1“ gastral sternite for the greater

part smooth, with few transverse striae posterior-

ly. Thyridium elongate. Thyridial index 1.5-2.0.

Epipleurum not separated from the 3 gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster strongly

compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with reddish brown

medially. Wings hyaline. Stigma brownish black.

Front femur black basally and ventrally, vellow-

ish dorsally and apically. Front tibia and tarsus

yellowish brown. Mid femur yellowish on the

apical 0.2. Mid tibia yellowish brown. Mid tarsus

dark brown. 2" gastral tergite completely black or

red on the posterior 0.1. 3" and 4" tergites red. 5'*

tergite sometimes narrowly marked with red an-

teriorly. Pilosity whitish.

d: Maxillary palpus yellowish brown. Man-

dible broadly yellow medially. Front leg from the

tip of the coxa onwards yellow, the femur brown

ventrally. In other characters similar to the 2.

Material (4??, 18): Japan: Honshu: Kyoto (SAP). Chi-

na: Fujian Prov. (= Fukien Prov.): Shaowu (AEI, Hinz /

ZSM). The locality in southern China belongs to the

Oriental region.

Variety: Ventral part of the prepectal carina and

transverse carina rather low. Pleural part of the

prepectus not depressed, densely rugose-punc-

tate and without strong dorsoventral wrinkles.

Petiole with narrow lateral areas. 4" gastral tergite

for the greater part black, narrowly marked with

red anteriorly. 5" tergite black. Material: 1? Rus-

sia, Primorye, mouth of River Amba-bira (PET).

This ® differs remarkably from the material from

southern China and Japan in the prepectus and

the prepectal carina, but agrees well in most other

characters. Its status is unknown.

Dusona glauca caliginosa (Walley, 1940)

Dusona latisulcor Hinz, 1994 (syn. nov.)

In the eastern Palearctic Region, this is a variable

and poorly defined taxon. It is similar to D. remota,

but differs from the latter in: Hind tibia yellowish

brown to black medially. 2" gastral tergite with a

dark brown or black posterior edge. 3" and 4"

tergites often marked with black.

Description. 2: Body length 8-12 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6-0.8. Frons slightly de-

pressed, coriaceous, finely rugose-punctate, with

a fine and often incomplete median longitudinal

carina, with short and indistinct radial wrinkles.

Antennal carina low and narrow, the rim slightly

bent upwards (inconspicuous). Face and dorsal

0.7 of the clypeus coriaceous, densely rugose and

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely punctate ventrally, its apex truncate

medially. Malar space index 0.4. Genal carina

slightly raised ventrally, joining the distinctly

raised oral carina at an acute angle. Genal index

0.6. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 40-48

segments, filiform, the apical 0.3 attenuate, 2"4

segment 1.8x, shortest segment 1.0x, penultimate

segment 1.1x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.2-0.4,

with longitudinal wrinkles ventrally. Mesoscu-

tum and scutellum coriaceous, dull, very densely

punctate. Scutellum with lateral carinae on the

anterior 0.5-0.8. Prescutellar groove smooth medi-

ally, with distinct longitudinal striae laterally. Pre-

pectal carina low, ventral part merging with the

pleural part, the dorsal 0.2 of the latter obliterated.

Transverse carina obliterated or partly present.

Mesopleurum distinctly coriaceous, densely punc-

tate. Speculum coriaceous and granulate-strigose,

rather dull. Depression in front of the speculum

dorsally with long longitudinal striae, ventrally

granulate-strigose and with few fine striae (varia-

ble), the striae merging with the punctation ante-

riorly. Metapleurum coriaceous, finely and densely

rugose. Pleural carina strong. Areolet rather small,

pointed or short-stalked, receiving the 2" recur-

rent vein distinctly distad of the middle. Nervel-

lus slightly inclivous, distinctly intercepted. Ner-

vellar index 0.7. Tibial spur index 0.6. Hind femur

5.4x as long as high. Hind tarsal claws short, with

short pectinate teeth basally and medially. Propo-

deum extending to 0.4 of the hind coxae, broadly

and rather deeply depressed, with short trans-

verse wrinkles anteriorly, with long transverse

wrinkles and sometimes with a short median lon-

gitudinal keel posteriorly, in other parts irregular-

ly wrinkled, costulae complete, longitudinal cari-

nae distinct medially and posteriorly, almost reach-

ing the costulae. Propodeal spiracle rather large,

3.0x as long as wide. Spiracular carina slightly

widened. Spiracular index 0.3. 1 gastral tergite

4.4x, 2"d tergite 1.9x as long as wide. Petiole with

distinct glymmae, smooth in front of the glym-

mae. Petiolar suture obliterated. 1* gastral sternite

smooth. Thyridium oval. Thyridial index about

1.0. Epipleurum not separated from the 3“ gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellow dorsally,

often also medially. Wings hyaline. Stigma brown

medially. Front leg from the femur onwards yel-

lowish brown. Mid femur marked with yellowish

brown on the apical 0.2. Mid tibia yellowish brown.

Mid tarsus dark brown. Hind tibia yellowish

brown to dark brown medially, narrowly black

basally, broadly black apically. 2” gastral tergite

with a dark brown or black posterior edge, to a

variable extend marked with red in front of the

posterior edge, sometimes completely black dor-

sally and marked with red only laterally. 3° and

4" tergites red, to a variable extend marked with

black, the 4" tergite sometimes completely black.

5' tergite sometimes narrowly marked with red

ventrolaterally. Pilosity whitish.

d: Propodeum not so broadly and rather shal-

lowly depressed. Variability of the colour pattern

similar to that of the 2.

Material (16%?, 1839): Russia: Irkutsk Prov.: Dach-

naya; Chita Prov.: Bukuka, Chita; Khabarovsk Terr.:

Khekhtsir near Khabarovsk, Troitskoye; Primorye:

Khasan (PET), Ussuriysk (KIS), Vladivostok (Hinz /

ZSM); Magadan Prov.: Magadan (MOS); Kuril Isl.:

Yuzhno-Kurilsk on Kunashir (PET). North Korea: Hy-

angsan (SAW). Japan: Hokkaido: Daisetu-zan (SAP),

Maruyama (Hinz/ZSM, SAP); Honshu: Mt. Minoo in

Osaka-fu (SAP), Nagano (OTT), Norikura-dake (AEI),

Sasayama; Kyushu: Hiko-san (SAP).

Variety: Body length 7-10 mm. 37-44 flagellar seg-

ments. Longitudinal carinae of the propodeum

often obliterated medially. Mandible black. Hind

tibia black or tinged with dark brown medially on

the dorsal side. Material (18??, 238): Russia:

Irkutsk Prov.: Lake Baykal; Khabarovsk Terr.:

Vysokogornyy (PET); Magadan Prov.: Magadan

(MOS); Kamchatka: Atlasovo (PET), Kozyrevsk

(Hinz/ZSM, PET), Milkovo; Kuril Isl.: Kurilsk on

Iturup, Sernovodsk and Yuzhno-Kurilsk on Ku-

nashir (PET). Japan: Hokkaido: Daisetu-zan (SAP),

Sapporo (AEI). This variety might include small

and dark specimens of D. glauca caliginosa.

Dusona graptor Hinz, 1985

This species is rather variable, and its differentia-

tion from D. melanator, D. norikurae or D. rufovari-

ata is difficult (see below). The association of the

one d with the ?? is not beyond all doubt.

Description. 2: Body length 8-10 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, with fine irregular wrinkles, with a

fine median longitudinal carina, without or with

short radial wrinkles. Antennal carina low and

narrow. Face and dorsal 0.7 of the clypeus coria-

ceous and very densely punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.25.

Genal carina low and almost straight ventrally,

joining the low oral carina at an acute angle. Genal

index 0.4. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

44-47 segments, filiform, the apical 0.3 attenuate,

2"! segment 2.0x, shortest segment 0.9x, penulti-

mate segment 1.0x as long as wide. Pronotum

coriaceous and finely and densely rugose-punc-

tate laterally on the dorsal 0.1, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous and densely punctate. Scutellum with

lateral carinae on the anterior 0.8. Prescutellar

groove with few fine longitudinal striae. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter complete, joining the anterior

edge of the mesopleurum. Transverse carina ob-

literated. Mesopleurum distinctly coriaceous, with

rather fine and rather dense punctation. Specu-

lum coriaceous. Depression in front of the specu-

lum granulate-strigose or with some fine longitu-

dinal striae, which merge with a coriaceous area

without punctation anteriorly. Metapleurum dis-

tinctly coriaceous, rather densely punctate. Pleu-

ral carina distinct. Areolet rather large, short-

stalked, receiving the 2" recurrent vein basad of

the middle. Nervellus almost vertical, weakly in-

tercepted. Nervellar index 0.8. Tibial spur index

0.5. Hind femur 4.8x as long as high. Hind tarsal

claws short, with short pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

coriaceous, rather shining and almost not de-

pressed anteriorly, slightly and rather narrowly

depressed posteriorly, with fine transverse wrin-

kles in the longitudinal furrow, coriaceous and

with irregular wrinkles beside the furrow, carinae

almost completely obliterated. Propodeal spiracle

rather large, 3.3x as long as wide. Spiracular cari-

na narrow. Spiracular index 0.3. 1" gastral tergite

3.6%x, 2" tergite 1.6%x as long as wide. Petiole with

small but distinct glymmae, smooth in front of the

glymmae. Petiolar suture fine. 1* gastral sternite

smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum separated from the 3" gas-

tral tergite anteriorly, folded in or pendent as a

narrow rectangular lobe, the crease (or bulge)

marked with black, a black lateral stripe parallel

with the crease in a very short distance anteriorly

and divergent from the ventrolateral edge of the

tergite posteriorly. Epipleural index 0.3. Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Tegula yel-

lowish brown. Wings hyaline. Stigma light brown

medially. Front leg from the tip of the coxa on-

wards yellowish brown. Sometimes front coxa

black and front trochanter marked with black.

Mid femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. Mid tarsus dark brown.

Hind tibia dark yellowish brown medially, nar-

rowly black basally, broadliy black apically. 2"“

gastral tergite dorsally red on the posterior 0.3,

laterally red on the posterior 0.4-0.5. 3” and 4"

tergites red. 4" tergite sometimes narrowly marked

with black dorsally. 5'" tergite sometimes narrow-

ly marked with red anteriorly. Pilosity whitish.

d: Flagellum with 50 segments. Propodeum

with the costulae and the longitudinal carinae

partly present, the latter obliterated anteriorly.

Tegula yellow. Front and mid legs from the tip of

the coxa onwards yellow. Hind tibia light yellow

medially. Postpetiole with small red marks later-

ally. 2"@ gastral tergite almost completely red lat-

erally. 4" tergite narrowly black dorsally (MOS).

In other characters similar to the ?. This 4 is

similar to D. rufovariata, but differs in: Longitudi-

nal carinae of propodeum partly obliterated. 3°“

gastral tergite completely red.

Material (8??, 18): Russia: Yevreyskiy A.O.: Biro-

bidzhan; Khabarovsk Terr.: Khabarovsk; Primorye:

Pozhiga SO of Dalnerechensk (PET), Spassk Dalniy

(Hinz/ZSM), Ussuriysk (PET), Vladivostok (MOS)

Variety: Tegula dark brown. 2"‘ gastral tergite

black. 4" tergite marked with red only laterally.

Material: 1? Russia, Khabarovsk Terr., Khekhtsir

near Khabarovsk (PET). Probably this 2 repre-

sents a dark variety of D. graptor. It differs from

D. norikurae in: Metapleurum coriaceous and with

fine punctation. Propodeum coriaceous and rath-

er shining anteriorly. On the other hand, it differs

from D. melanator in: Depression in front of the

speculum granulate-strigose, with a coriaceous

area without striae or punctation in front the de-

pression. 3" gastral tergite almost completely red.

Dusona humilis (Förster, 1868)

Description. 2: Body length 6-8 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.9. Frons slightly depressed, coriaceous, a

punctation hardly visible, with a fine and incom-

plete median longitudinal carina, with short radi-

al wrinkles. Antennal carina low and narrow. Face

coriaceous, dull, finely and rather densely punc-

tate, with some very fine wrinkles. Clypeus flat,

coriaceous, dull, finely and dispersely punctate,

its apex truncate medially. Malar space index 0.4.

Genal carina almost straight and slightly raised

ventrally, joining the distinctly raised oral carina

at an acute angle, the gena slightly depressed

close to the junction. Genal index 0.3. Temple

coriaceous and with very fine disperse punctation

medially. Flagellum with 34-40 segments, filiform,

the apical 0.2 slightly attenuate, 2" segment 1.9x,

shortest segment 0.9x, penultimate segment 1.1x

as long as wide. Pronotum coriaceous and dense-

ly rugose-punctate laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum co-

riaceous, dull, finely and rather densely punctate.

Scutellum coriaceous, finely and dispersely punc-

tate, with lateral carinae on the anterior half. Pres-

cutellar groove smooth medially, with fine longi-

tudinal striae laterally. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

complete, joining the anterior edge of the meso-

pleurum. Transverse carina low, often obliterated

posteriorly. Mesopleurum coriaceous, dull, rather

densely punctate. Speculum coriaceous, somewhat

shining. Depression in front of the speculum gran-

ulate-strigose or with very fine longitudinal stri-

ae, which merge with a coariaceous area without

punctation anteriorly. Metapleurum coriaceous,

finely and densely punctate or rugose-punctate.

Pleural carina complete. Areolet large, short-

stalked, receiving the 2" recurrent vein slightly

distad of the middle. Nervellus slightly inclivous,

weakly intercepted, its upper branch slightly bent

inwards. Nervellar index 0.8-0.9. Tibial spur in-

dex 0.7. Hind femur 4.5x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to 0.3 of the hind

coxae, broadly and not deeply depressed, sulcus

coriaceous anteriorly, irregularly wrinkled and

with a short longitudinal keel posteriorly, costu-

lae partly present, longitudinal carinae almost com-

pletely obliterated. Propodeal spiracle rather large,

3.0x as long as wide. Spiracular carina narrow.

Spiracular index 0.3. 1% gastral tergite 3.4x, 2"

tergite 1.3x as long as wide. Petiole with distinct

glymmae, which are bordered by longitudinal

carinae dorsally, smooth in front of the glymmae.

Petiolar suture partly obliterated. 1* gastral ster-

nite usually smooth, rarely with fine sculpture.

Thyridium oval. Thyridial index about 2.0. Epi-

pleurum not separated from the 3" gastral tergite,

the latter with distinct black lateral stripes well

above the ventrolaterai edge. Gaster distinctly

compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus brownish

black. Mandible narrowly marked with reddish

brown dorsally and medially, sometimes com-

pletely black. Tegula dark brown. Wings hyaline.

Stigma dark brown medially. Front leg from the

femur onwards yellowish red, the femur marked

with black ventrally. Mid femur yellowish red on

the apical 0.2. Mid tibia yellowish red. Mid tarsus

dark brown. Hind tibia broadly dark yellowish

brown medially, narrowly black basally, broadly

black apically. 2” gastral tergite dorsally red on

the posterior 0.3, laterally red on the posterior 0.5.

34 and 4" tergites red, the 4" tergite sometimes

marked with black or completely black. Pilosity

whitish.

d: Front trochanter yellowish red. Front fe-

mur often completely red. In other characters sim-

ilar to the 2.

Material (10%, 18): Russia: Chita Prov.: Karymskoye;

Primorye: Ussuriysk (PET). Kirgiziya: Terskey Alatau

Mts., 2000-2400 m (LIN). Japan: Hokkaido: Jozankei

(AEI).

Dusona impressifrons Hinz, 1979

Description. 2: Body length 17-19 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 1.1. Frons coriaceous and

densely punctate dorsally, deeply depressed ven-

trally above the antennal carinae, coriaceous and

finely and densely rugose in the depressions, with

few distinct radial wrinkles, with a median longi-

tudinal bulge between these depressions and with

a fine median longitudinal carina on top of this

bulge. Antennal carina strongly raised, the rim

bent upwards (ear-like), at the same time with a

dorsal projection (nose-like), with fine transverse

striae on that projection. Face and dorsal half of

the clypeus coriaceous, coarsely and densely punc-

tate, partly rugose-punctate. Clypeus flat, coria-

ceous and finely and rather densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.3. Genal carina slightly raised and slightly

bent forwards ventrally, joining the slightly raised

oral carina at an acute angle. Gena narrowly de-

pressed close to the junction. Genal index 0.3.

Temple coriaceous and with fine rather dense

punctation medially. Flagellum with 59 segments,

filiform, the apical 0.4 distinctly attenuate, 2" seg-

ment 1.6x, shortest segment 1.0x, penultimate seg-

ment 1.3x as long as wide. Pronotum finely and

densely rugose-punctate laterally on the dorsal

0.4, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely or very

densely punctate. Scutellum with strong lateral

carinae on the anterior 0.2. Prescutellar groove

with longitudinal striae. Ventral part of the pre-

pectal carina merging with the transverse carina,

both carinae slightly raised. Pleural part of the

prepectal carina low, its dorsal 0.3 obliterated.

Mesopleurum finely coriaceous, distinctly and

densely punctate. Speculum finely coriaceous and

granulate-strigose. Depression in front of the

speculum dorsally with fine longitudinal striae,

which merge with the punctation anteriorly, the

ventral 0.3 granulate-strigose. Metapleurum co-

riaceous, finely and densely rugose-punctate. Pleu-

ral carina strong. Areolet large, pointed, receiving

the 2” recurrent vein slightly distad of the mid-

dle. Nervellus slightly inclivous, distinctly inter-

cepted. Nervellar index 0.7. Tibial spur index 0.5.

Hind femur 5.4x as long as high. Hind tarsal claws

short, with few distinct pectinate teeth. Propo-

deum extending to 0.3 ofthe hind coxae, distinct-

ly and broadly depressed, sulcus with short trans-

verse wrinkles anteriorly and medially, with long

transverse wrinkles posteriorly, costulae partly

present, longitudinal carinae present medially and

posteriorly. Propodeal spiracle large, 3.5x as long

as wide. Spiracular carina partly widened. Spirac-

ular index 0.1-0.2. 1“ gastral tergite 4.3x, 2" tergite

1.6x as long as wide. Petiole with distinct glym-

mae, which are bordered by longitudinal carinae

dorsally, with very fine longitudinal striae in front

of the glymmae. Petiolar suture for the greater

part obliterated. 1“ gastral sternite for the greater

part smooth, coriaceous anteriorly, with few trans-

verse striae posteriorly. Thyridium oval. Thyridi-

al index 1.5-2.0. Epipleurum separated from the

34 gastral tergite anteriorly, the crease marked

with black. Epipleural index 0.4-0.5. Gaster strong-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Tegula yel-

lowish brown. Wings slightly infuscate. Stigma

yellowish brown medially. Front leg from the tip

of the coxa onwards yellowish brown, the femur

marked with brown ventrally. Mid femur yellow-

ish red apically or completely black. Mid tibia

yellowish red. Mid tarsus brown. Hind tibia broad-

ly yellow medially, narrowly black basally, broadly

black apically. 2" gastral tergite dorsally red on

the posterior 0.1, laterally red on the posterior 0.3.

3"d and 4" tergites red, sometimes indistinctly

tinged with black. Pilosity whitish.

d: Spiracular index 0.4. Front femur yellow-

ish brown. Mid femur yellowish brown dorsally,

dark brown ventrally. In other characters similar

to the 2.

Material (22%, 18): Japan: Hokkaido: Apoi-dake,

Maruyama, Sapporo (SAP).

Dusona inconspicua Horstmann, spec. nov.

Holotype (J): Japan, Hokkaido, locality unreadable,

21.11.1928 (SAP). i

Description. d: Body length 10 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.7. Frons slightly depressed, coriaceous,

densely rugose-punctate, with a fine median lon-

gitudinal carina, with distinct radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina low and narrow. Face and

dorsal 0.6 of the clypeus very densely wrinkled,

not visibly coriaceous. Clypeus flat, densely punc-

tate on a smooth background ventrally, its apex

truncate medially. Malar space index 0.5. Genal

carina low and straight ventrally, joining the dis-

tinctly raised oral carina at an acute angle. Genal

index 0.4. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

43 segments, filiform, the apical 0.3 attenuate, 2"*

segment 1.7x, shortest segment 1.0x, penultimate

segment 1.1x as long as wide. Pronotum coria-

ceous and densely rugose-punctate laterally on

the dorsal 0.3, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous, dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.6. Prescutellar groove with longitudinal

striae. Prepectal carina low, ventral part merging

with the pleural part, the dorsal 0.3 of the latter

obliterated. Transverse carina indicated anterior-

ly. Mesopleurum finely coriacerous, distinctly and

densely punctate. Speculum coriaceous and fine-

ly granulate-strigose. Depression in front of the

speculum with fine longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum densely rugose-punctate, not visibly coria-

ceous. Pleural carina strong. Areolet large, point-

ed, receiving the 2” recurrent vein distinctly ba-

sad of the middle. Nervellus slightly inclivous,

intercepted. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 5.8x as long as high. Hind tarsal

claws short, with short but distinct pectinate teeth

basally and medially. Propodeum extending to

0.4 of the hind coxae, slightly and broadly de-

pressed, the broad shallow sulcus for the greater

part with strong irregular wrinkles, with some

short transverse wrinkles anteriorly and a median

longitudinal keel posteriorly, carinae almost com-

plete, but partly covered with strong wrinkles.

Propodeal spiracle small, 2.6x as long as wide.

Spiracular carina narrow. Spiracular index 0.8.

1" gastral tergite 4.8x, 2” tergite 2.9x as long as

wide. Petiole with small and rather shallow glym-

mae, smooth in front of the glymmae. Petiolar

suture obliterated. 1“ gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum of the 3"! gastral apparently tergite folded in

anteriorly, the crease not marked with black, black

lateral stripes not visible. Epipleural index about

0.2. Gaster strongly compressed posteriorly.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible nar-

rowly marked with yellowish red dorsally and

medially. Tegula yellowish red. Wings hyaline.

Stigma medium brown medially. Front trochant-

er and femur yellowish red. Front tibia and tarsus

yellow. Mid femur yellowish red on the apical 0.4.

Mid tibia yellow. Mid tarsus yellowish brown.

Hind tibia yellow medially, narrowly black basal-

ly, very broadly black apically. 2" gastral tergite

dorsally red on the posterior 0.2, broadly red lat-

erally, the posterior edge slightly tinged with

brown. 3°! and 4" tergites red. Pilosity whitish.

? unknown.

Dusona indistinctor Hinz, 1979

One paratype of this taxon from Tretyakovo on

Kunashir/Kuril Isl./ Russia (PET) belongs to D. xe-

nocampta.

Description. 2: Body length 11-12 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and rather densely punctate,

with a fine and incomplete median longitudinal

carina, with short radial wrinkles. Antennal cari-

na low and narrow. Face and dorsal 0.7 of the

clypeus coriaceous, densely and rather coarsely

rugose. Clypeus flat, finely coriaceous and finely

and dispersely punctate ventrally, its apex trun-

cate medially. Malar space index 0.3. Genal carina

low and straight ventrally, joining the slightly

raised oral carina at an acute angle. Genal index

0.3. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with 50-53

segments, filiform, the apical 0.4 distinctly attenu-

aten22d segment 1.9x, shortest segment 0.9x, pe-

nultimate segment 1.3x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.1, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, dull, very

densely punctate. Scutellum with lateral carinae

on the anterior 0.7. Prescutellar groove almost

completely smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

coriaceous, rather dull, distinctly and rather dense-

Iy punctate. Speculum punctate posteriorly, co-

riaceous anteriorly. Depression in front of the

speculum coriaceous, with fine and short longitu-

dinal striae, which merge with an area without

punctation anteriorly. Metapleurum coriaceous,

finely and densely punctate. Pleural carina dis-

tinct. Areolet large, short-stalked, receiving the

2"d recurrent vein at the middle. Nervellus slight-

ly reclivous, distinctly intercepted. Nervellar in-

dex 0.7. Tibial spur index 0.6. Hind femur 5.0x as

long as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, broadly and

deeply depressed, sulcus with long transverse

wrinkles over its total breadth, carinae almost

completely obliterated. Propodeal spiracle large,

3.1x as long as wide. Spiracular carina narrow.

Spiracular index 0.2. 1* gastral tergite 4.0x, 2"

tergite 1.5% as long as wide. Petiole with rather

small but distinct glymmae at the anterior edge of

indistinct narrow lateral areas, smooth in front of

these areas. Petiolar suture for the greater part

obliterated. 1* gastral sternite smooth. Thyridium

elongate. Thyridial index about 1.0. Epipleurum

not separated from the 3" gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellow, some-

times brown basally. Mandible broadly yellow

medially. Flagellum medium brown to dark brown

apically. Tegula dark brown. Wings hyaline. Stig-

ma yellowish brown medially. Front leg from the

tip of the coxa onwards yellow, trochanter and

femur marked with brown ventrally, sometimes

coxa black and trochanter completely brown. Mid

femur yellowish on the apical 0.2. Mid tibia yel-

lowish brown. Mid tarsus dark brown. Hind tibia

yellowish brown medially, narrowly black basal-

ly, broadly black apically. 2" gastral tergite dor-

sally red on the posterior 0.2, almost completely

red laterally. 3° and 4" tergites red. 5" tergite

sometimes marked with red ventrolaterally. Pi-

losity whitish.

d: Longitudinal carinae of propodeum present

medially and posteriorly. Tegula yellow. Flagel-

lum black to its apex. Greater part of the front

coxa and tip of the mid coxa yellow. Mid trochant-

er yellowish brown. Mid femur yellowish brown

basally and ventrally, yellow dorsally and apical-

ly. Mid tibia and tarsus yellow. 5'" gastral tergite

for the greater part red. In other characters similar

to the 9.

Material (72%, 234): Russia: Primorye: Nakhodka

(PET), Ussuriysk (LIN, PET), Vladivostok (PET). Ja-

pan: Hokkaido: Jozankei (AEI, Hinz/ZSM); Honshu:

Norikura-dake (AEI); Shikoku: Susaki (SAP). Hinz

(1979: 224) mentions additional localities in Japan.

Dusona inermis (Förster, 1868)

Description. ?: Body length 8-10 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.5. Frons scarcely depressed, coriaceous,

rugose-punctate, with a fine median longitudinal

carina, with short radial’wrinkles, which extend

to 0.4 the distance to the median ocellus. Antennal

carina low and narrow. Face and dorsal half of the

clypeus coriaceous, densely rugose-punctate.

Clypeus flat, finely coriaceous and rather densely

punctate ventrally, its apex truncate medially.

Malar space index 0.4. Genal carina slightly raised

ventrally, slightly bent sidewards and often also

forwards, joining the oral carina at the base of the

mandible almost at a right angle. Temple coria-

ceous and with fine disperse punctation medially.

Flagellum with 42-43 segments, filiform, the api-

cal 0.3 attenuate, 2"! segment 2.0x, shortest seg-

ment 0.9x, penultimate segment 1.1x as long as

wide. Pronotum with longitudinal wrinkles later-

ally, with additional punctures on its dorsal 0.2.

Mesoscutum and scutellum coriaceous, very dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.6. Prescutellar groove for the greater

part smooth. Ventral part of the prepectal carina

merging with the transverse carina, both carinae

distinctly but not very strongly raised, the former

distinctly emarginate medially. Pleural part of the

prepectal carina low, its dorsal 0.3 obliterated.

Mesopleurum distinctly coriaceous, rather dense-

ly or densely punctate. Speculum coriaceous and

granulate-strigose, scarcely shining. Depression in

front of the speculum with fine longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum coriaceous, finely and densely ru-

gose. Pleural carina distinct. Areolet large, stalked,

receiving the 2" recurrent vein at or distad of the

middle. Nervellus slightly reclivous, weakly in-

tercepted. Nervellar index 0.8. Tibial spur index

0.7. Hind femur 5.3x as long as high. Hind tarsal

claws short, with short pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

broadly and distinctly but not deeply depressed,

sulcus for the greater part with long transverse

wrinkles, carinae obliterated, the longitudinal car-

inae sometimes indicated medially and posterior-

ly. Propodeal spiracle rather large, 3.6x as long as

wide. Spiracular carina narrow. Spiracular index

0.3. 1% gastral tergite 4.4x, 2" tergite 2.1x as long

as wide. Petiole with rather small glymmae at the

anterior edge of narrow lateral areas, which may

be obliterated, smooth in front of the glymmae.

Petiolar suture partly obliterated. 1* gastral ster-

nite smooth. Thyridium oval. Thyridial index about

2.0. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly marked with yellowish red

dorsally and medially. Tegula yellowish. Wings

hyaline. Stigma medium brown medially. Front

trochanter marked with yellow and brown. Front

femur yellowish red. Front tibia and tarsus yel-

low. Mid femur yellow on the apical 0.3. Mid tibia

yellow. Mid tarsus brown. Hind tibia light yellow

medially, narrowly black basally, rather narrowly

black apically (apical black ring extending over

0.1-0.2 of the tibia). 2” gastral tergite red on the

posterior 0.4-0.6. 3"! and 4" tergites red. 5" tergite

sometimes marked with red anteriorly. Pilosity

whitish.

d: Flagellum with 44-46 segments. Longitu-

dinal carinae of the propodeum often present me-

dially. Maxillary palpus yellow. Mandible almost

completely yellow. Front and mid legs from the

tip of the coxa onwards yellow, the femora yel-

lowish red. Colour pattern of the hind tibia as in

the 2. 2" gastral tergite red on the posterior 0.3-0.4.

In other characters similar to the ?.

Host: Semiothisa liturata (Clerck, 1759) (Geometri-

dae) (MIN).

Material (7??, 483): Russia: Novosibirsk Prov.: Aka-

demgorodok (PET); Altaysk Terr.: Volchikha (MIN);

Irkutsk Prov.: Irkutsk; Chita Prov.: Chita, Karymskoye

(PET).

Dusona infesta (Förster, 1868)

Description. 2: Body length 15-16 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons slightly depressed,

coriaceous, densely rugose and rugose-punctate

dorsally, with a fine and sometimes indistinct

median longitudinal carina, with long and dis-

tinct radial wrinkles, which reach the median ocel-

lus. Antennal carina slightly but distinctly raised,

the rim distinctly bent upwards (ear-like), form-

ing a narrow crescent-shaped plate, the latter with

fine transverse striae. Face and dorsal 0.6 of the

clypeus coriaceous, densely rugose and rugose

punctate. Clypeus flat, coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.4. Genal carina slightly

raised ventrally, slightly bent sidewards, joining

the slightly raised oral carina almost at a right

angle. Genal index 0.3. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

ellum with 53-57 segments, filiform, the apical 0.3

distinctly attenuate, 2° segment 1.9x, shortest seg-

ment 1.0x, penultimate segment 1.3x as long as

wide. Pronotum densely rugose-punctate and with

oblique and longitudinal wrinkles laterally. Meso-

scutum and scutellum coriaceous, dull, very dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.7. Prescutellar groove with longitudinal

striae. Prepectal carina low, ventral part merging

with the pleural part, the latter almost joining the

anterior edge of the mesopleurum, the dorsal

0.1-0.2 replaced by several wrinkles. Transverse

carina present anteriorly. Mesopleurum coria-

ceous, densely punctate. Speculum punctate pos-

teriorly, coriaceous anteriorly, rather dull. Depres-

sion in front of the speculum granulate-strigose,

with short longitudinal wrinkles on the dorsal

half, which merge with the punctation or with a

narrow coriaceous area without punctation ante-

riorly. Metapleurum coriaceous, densely rugose-

punctate. Pleural carina distinct. Areolet large,

short-stalked, receiving the 2” recurrent vein at

or basad of the middle. Nervellus distinctly in-

clivous, distinctly intercepted. Nervellar index

0.7-0.8 (Fig. 26). Tibial spur index 0.7. Hind femur

5.9x as long as high. Hind tarsal claws short, with

distinct pectinate teeth basally and medially. Pro-

podeum extending to 0.4 ofthe hind coxae, broad-

ly and deeply depressed, sulcus almost complete-

ly with long transverse wrinkles, costulae partly

present, longitudinal carinae strong medially and

posteriorly. Propodeal spiracle rather large, 3.5x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.6. 1* gastral tergite 4.5x, 2"’ tergite

1.9x as long as wide. Petiole with large glymmae,

which are bordered by longitudinal carinae dor-

sally, almost smooth in front of the glymmae,

sometimes with some short transverse striae ante-

riorly. Petiolar suture obliterated or partly present.

1* gastral sternite for the greater part smooth,

sometimes with transverse striae posteriorly. Thy-

ridium oval. Thyridial index about 1.5. Epipleu-

rum separated from the 3"! gastral tergite anteri-

orly, the crease marked with black. Epipleural

index 0.4-0.5. Gaster strongly compressed posteri-

orly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellow medially. Tegu-

la yellow. Wings hyaline. Stigma light brown me-

dially. Front leg from the apical half of the coxa

onwards yellow. Mid femur yellow on the apical

0.2-0.5. Mid tibia yellow. Mid tarsus yellowish

brown. Hind tibia broadly yellow medially, nar-

rowly black basally and apically. 2" gastral tergite

red on the posterior 0.4. 3’ and 4" tergites red.

Pilosity whitish.

d: Similar to the ?. 13 from Hiko-san/Ky-

ushu/Japan (SAP) is darker: Mandibel narrowly

marked with reddish brown medially. Tegula yel-

lowish brown. Front coxa black.

Material (5?®2, 18): Russia: Irkutsk Prov.: Irkutsk;

Chita Prov.: Nerchinskiy Zavod (PET). Japan: Hokkai-

do: Sapporo (SAP); Honshu: Kamikochi near Noriku-

ra-dake, Nagano (AEI); Kyushu: Hiko-san (SAP).

Dusona infundibulum Horstmann, spec. nov.

Holotype (S): Japan, Shikoku, Kajigamori, 20.8.

1952 (SAD):

Description. d: Body length 14 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons coriaceous and densely punctate dor-

sally, with two very deep funnel-like depressions

ventrally, distinctly rugose in these depressions,

with a median longitudinal bulge, without a cari-

na on top of this bulge. Antennal carina distinctly

raised, its rim distinctly bent upwards (ear-like),

with fine transverse striae on the rim. Face and

dorsal 0.6 of the clypeus finely coriaceous, dense-

ly rugose-punctate. Clypeus flat, finely coriaceous

and rather densely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina straight ventrally, distinctly raised, joining

the raised oral carina at an acute angle, the tip of

the junction not shifted ventrally. Genal index 0.3.

Temple with distinct and rather dense or dense

punctaion on a very slightly coriaceous and partly

smooth background medially. Flagellum with 56

segments, filiform, the apical 0.3 attenuate, 2“

segment 1.5x, shortest segment 1.0x, penultimate

segment 1.3x as long as wide. Pronotum with

very dense punctation on an almost smooth back-

ground laterally on the dorsal 0.6, with rather

irregular wrinkles ventrally. Mesoscutum and

scutellum very finely coriaceous, densely punc-

tate. Scutellum with lateral carinae on the anterior

0.2. Prescutellar groove with distinct longitudinal

striae. Prepectal carina low, ventral part merging

with the pleural part, the latter complete, joining

the anterior edge of the mesopleurum. Transverse

carina indicated anteriorly. Mesopleurum strong-

ly and densely punctate on a smooth background.

Speculum finely granulate-strigose, shining. De-

pression in front of the speculum with fine longi-

tudinal striae, which merge with the punctation

anteriorly. Metapleurum densely rugose-punctate

on a smoth or almost smooth background. Pleural

carina complete. Areolet large, short-stalked, re-

ceiving the 2" recurrent vein distad of the middle.

Nervellus distinctly inclivous, strongly intercept-

ed. Nervellar index 0.8. Tibial spur index 0.7. Hind

femur 5.9x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to 0.4 of the hind

coxae, rather broadly and not deeply depressed,

sulcus with short transverse wrinkles anteriorly

and medially, with long transverse wrinkles pos-

teriorly, costulae partly present, longitudinal car-

inae slightly indicated medially, strongly devel-

oped posteriorly. Propodeal spiracle rather small,

3.1x as long as wide. Spiracular carina narrow.

Spiracular index 0.4. 1% gastral tergite 4.4x, 2"‘

tergite 2.3x as long as wide. Petiole with large and

somewhat elongated glymmae, for the greater part

smooth in front of the glymmae, with some very

fine longitudinal striae. Petiolar suture very fine,

partly obliterated. 1* gastral sternite coriaceous

anteriorly, smooth medially, transversely striate

posteriorly. Thyridium elongate. Thyridial index

about 1.5. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.8. Gaster distinctly compressed

posteriorly.

Colour: Black. Maxillary palpus blackish.

Wings distinctly tinged with brown. Stigma me-

dium brown medially. Front femur reddish brown

dorsally, black ventrally. Front tibia and tarsus

reddish brown. Mid femur dark reddish brown

dorsally, black ventrally. Mid tibia and tarsus dark

reddish brown. Hind tibia dark reddish brown

subbasally, narrowly black basally, very broadly

black apically. Postpetiole narrowly marked with

red laterally. 2" gastral tergite black dorsally, red

laterally. 34 to 5" tergites narrowly black dorsally,

broadly red laterally. Pilosity whitish.

® unknown.

Dusona insignita (Förster, 1868)

Description. 2: Body length 10-11 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous and

with fine rather dense punctation dorsally, coria-

ceous and almost without punctures ventrally,

with a fine and partly obliterated median longitu-

dinal carina, without radial wrinkles. Antennal

carina low and narrow. Face and dorsal half of the

clypeus coriaceous, densely rugose-punctate.

Clypeus flat, coriaceous and dull, with fine dis-

perse punctation ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.4.

Temple coriaceous and with very fine disperse

punctation medially. Flagellum with 46-48 seg-

ments, filiform, the apical 0.3 distinctly attenuate,

2"! segment 1.6x, shortest segment 1.0x, penulti-

mate segment 1.2x as long as wide. Pronotum

finely and densely rugose laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum coriaceous, densely punctate. Scutellum

finely coriaceous, rather densely or densely punc-

tate, with lateral carinae on the anterior 0.5-0.7.

Prescutellar groove for the greater part smooth.

Prepectal carina low, the ventral part merging

with the pleural part, the latter complete or almost

complete, the dorsal 0.1 sometimes replaced by

several wrinkles. Transverse carina present, low.

Mesopleurum coriaceous, dull, rather densely or

densely punctate. Speculum finely coriaceous,

shining. Depression in front of the speculum gran-

ulate-strigose and with fine longitudinal striae,

which merge with the punctation or with a nar-

row area without punctation anteriorly. Meta-

pleurum coriaceous, dull, finely and rather dense-

Iy punctate. Pleural carina complete. Areolet large,

pointed or short-stalked, receiving the 2"4 recur-

rent vein slightly distad of the middle. Nervellus

inclivous, distinctly intercepted. Nervellar index

0.8. Tibial spur index 0.6. Hind femur 5.0x as long

as high. Hind tarsal claws short, with short pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, deeply but not broadly depressed,

the longitudinal furrow almost smooth anteriorly,

and medially, transversely wrinkled posteriorly,

costulae almost completely obliterated, longitudi-

nal carinae present posteriorly, sometimes indi-

cated medially. Propodeal spiracle rather large,

2.8x as long as wide. Spiracular carina narrow.

Spiracular index 0.4. 1* gastral tergite 3.6x, 2"4

tergite 1.4x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal

carinae dorsally, with a long row of short trans-

verse striae or small pits in a long longitudinal

furrow in front of the glymmae, often with some

longitudinal striae ventrolaterally. Petiolar suture

complete. 1“ gastral sternite for the greater part

smooth or with very fine sculpture. Thyridium

oval. Thyridial index about 1.0. Epipleurum sepa-

rated from the 3" gastral tergite anteriorly, usual-

ly folded in, the crease marked with black, a black

lateral stripe parallel with the crease in a short

distance and divergent from the ventrolateral edge

of the tergite posteriorly. Epipleural index 0.2.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.7.

Colour: Black. Maxillary palpus dark brown.

Mandible marked with yellowish red to reddish

brown medially. Tegula dark brown. Wings hya-

line. Stigma yellowish brown medially. Front leg

from the femur onwards yellowish red. Mid fe-

mur black basally and ventrally, yellowish red

dorsally and apically. Mid tibia yellowish red.

Mid tarsus dark brown. Hind tibia yellowish

brown medially, narrowly black basally, broadly

black apically. 2" gastral tergite dorsally com-

pletely black or red on the posterior 0.1-0.2, later-

ally red on the posterior 0.3-0.5. 3" tergite red,

sometimes tinged with black dorsally. 4" tergite

completely black or marked with red anteriorly.

Pilosity whitish.

d: Propodeum with the longitudinal furrow

sometimes for the greater part irregularly wrin-

kled, with transverse wrinkles posteriorly. Maxil-

lary palpus yellowish red to brown. Mandible

broadly yellowish red medially. Tegula yellowish

brown. Front leg from the tip of the coxa onwards

yellowish. Mid leg from the tip of the coxa or from

the trochanter onwards yellowish. 2"4 gastral

tergite sometimes dorsally red on the posterior

0.3. 3X and 4" tergites sometimes completely red

or marked with black dorsally. In other characters

similar to the 2.

Material (3??, 13): Russia: Primorye: Roshchino on

River Iman. Kazakhstan: Alma-Ata (PET).

Dusona iwatae Hinz, 1994

Description. 2: Body length 7 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons scarcely depressed, coriaceous, finely

and densely rugose-punctate, with a very fine and

indistinet median longitudinal carina, with few

and indistinct radial wrinkles. Antennal carina

low and narrow. Face and dorsal 0.3 of the clypeus

coriaceous, finely and densely rugose-punctate.

Clypeus slightly convex, finely and rather dense-

ly punctate ventrally, its apex slightly convex

medially. Malar space index 0.4. Genal carina low

and straight ventrally, joining the low oral carina

at an acute angle. Genal index 0.4. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 27 segments, clavate sub-

apically, broadest segment 1.7x as wide as the

apical end of the 1* segment, 2” segment 2.3x,

shortest segment 0.8x, penultimate segment 0.9x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.1, with longitudinal

wrinkles ventrally. Mesoscutum coriaceous, fine-

ly and densely punctate. Scutellum finely coria-

ceous, with fine disperse punctation, with lateral

carinae on the anterior 0.3. Prescutellar groove

smooth medially, with longitudinal striae lateral-

lv. Ventral part of the prepectal carina merging

with the transverse carina, both carinae rather

low. Pleural part of the prepectal carina low, com-

plete, joining the anterior edge of the mesopleu-

rum. Mesopleurum distinctly coriaceous, rather

dull, finely and densely punctate. Speculum al-

most smooth, shining. Depression in front of the

speculum for the greater part with distinct longi-

tudinal striae, which merge with the punctation

anteriorly, the ventral 0.2 almost smooth. Meta-

pleurum coriaceous, finely and densely rugose-

punctate. Pleural carina fine but complete. Areo-

let rather small, stalked, receiving the 2" recur-

rent vein distinctly distad of the middle. Nervellus

slightly reclivous, intercepted. Nervellar index 0.4.

Tibial spur index 0.6. Hind femur 5.1x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, slightly and broadliy depressed,

sulcus with long transverse wrinkles, costulae fine

but complete, longitudinal carinae almost com-

pletely obliterated. Propodeal spiracle small, 2.0x

as long as wide. Spiracular carina partly widened.

Spiracular index 1.0. 1% gastral tergite 4.1x, 2"4

tergite 1.5x as long as wide. Petiole with distinct

long and narrow glymmae, which are bordered

by longitudinal carinae dorsally, smooth in front

of the glymmae. Petiolar suture complete. 1% gas-

tral sternite for the greater part smooth, with few

transverse striae posteriorly. Thyridium elongate.

Thyridial index about 1.0. Epipleurum separated

from the 3" gastral tergite, the crease marked with

black. Epipleural index 0.8. Gaster slightly com-

pressed posteriorly. Ovipositor index 1.1.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible nar-

rowly marked with reddish brown medially.

Wings hyaline. Stigma dark brown medially. Front

leg from the apical end of the trochanter onwards

yellow. Mid leg from the apical 0.3 of the femur

onwards yellow. Hind tibia yellowish brown, nar-

rowly black basally, not darkened apically. Hind

tarsus dark brown dorsally, yellowish brown ven-

trally. 2° gastral tergite dorsally red on the poste-

rior 0.3, laterally red on the posterior 0.4. 3% tergite

red. 4" to 6" tergites broadly medium brown dor-

sally, marked with red ventrolaterally. 7" tergite

red. Gastral sternites red. Pilosity whitish.

d unknown.

Material (19): Japan: Honshu: Sasayama (SAP).

Dusona japonica (Cameron, 1906)

The holotype (2) of Campoplex japonicus Cameron

from Kobe/Honshu/Japan (NHM) has recently

been studied by the junior author.

Description. 2: Body length 10-13 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons not depressed, coriaceous, rugose, with-

out a median longitudinal carina, without or with

rather short radial wrinkles, which extend to 0.2-0.4

the distance to the median ocellus. Antennal sock-

ets close to each other. Antennal carina low and

narrow. Face and dorsal 0.4 of the clypeus coria-

ceous, densely rugose-punctate. Clypeus flat or

slightly concave, finely coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.6. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.6.

Temple coriaceous and with fine dense puncta-

tion medially. Flagellum with 44-52 segments, fil-

iform, the apical 0.2 slightly attenuate, 2”! seg-

ment 1.5x, shortest segment 0.9x, penultimate seg-

ment 1.3x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.2, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum finely coriaceous, very densely punc-

tate. Scutellum with lateral carinae on the anterior

half. Prescutellar groove smooth. Ventral part of

prepectal carina merging with the transverse car-

ina, both carinae low. Pleural part of the prepectal

carina completely obliterated (Fig. 21). Pleural part

of the prepectus densely rugose-punctate. Meso-

pleurum distinctly and densely punctate on a

smooth background, sometimes with additional

wrinkles or densely rugose-punctate. Speculum

finely granulate-strigose. Depression in front of

the speculum with distinct rather short longitudi-

nal striae, which merge with the punctation ante-

riorly. Metapleurum densely rugose. Pleural cari-

na fine, sometimes covered with wrinkles. Areo-

let large, short-stalked, receiving the 2" recurrent

vein distinctly basad of the middle. Nervellus

slightly inclivous or vertical, weakly intercepted.

Nervellar index 0.7-0.8. Tibial spur index 0.7. Hind

femur 5.8x as long as high. Hind tarsal claws long,

with rather long pectinate teeth basally and medi-

ally. Propodeum extending to 0.4 of the hind cox-

ae, not depressed or with a shallow and narrow

longitudinal furrow, for the greater part irregu-

larly wrinkled, with few short transverse wrin-

kles in the furrow (if the latter is present), with

some longitudinal wrinkles posteriorly, carinae

almost completely obliterated. Propodeal spiracle

small, 3.0x as long as wide. Spiracular carina re-

placed by several wrinkles. Spiracular index 1.1.

1% gastral tergite 4.6x, 2"! tergite 1.9x as long as

wide. Petiole without or with very small glymmae

atthe anterior edge of narrow lateral areas, smooth

or with very fine sculpture in front of these areas.

Petiolar suture fine, partly obliterated. 1* gastral

sternite for the greater part smooth, with few trans-

verse striae posteriorly. Thyridium oval. Thyridi-

al index about 2.0. Epipleurum not separated from

the 3X gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.5.

Colour: Black. Wings hyaline. Front femur

dark brown on the apical 0.2. Front tibia dark

brown. 2"! gastral tergite completely black or red

on the posterior 0.1-0.2. 3’ and 4" tergites red. 5"

tergite sometimes marked with red anteriorly. Pi-

losity whitish.

d: Mandible broadly yellow medially. Front

leg from the coxa onwards with a longitudinal

yellow stripe anteriorly. Mid tibia sometimes

tinged with brown. In other characters similar to

the 2.

Material (26??, 1833): Russia: Irkutsk Prov.: Dach-

naya (Hinz/ZSM, PET), Lake Baykal (ZSM); Sakhalin:

Tymovskoye. Uzbekistan: Kamashi (PET). Kazakh-

stan: Chimkent (MOS). Tadzhikistan: Dushanbe. Kir-

giziya: Frunze (= Bishkek), Iski-Naukat (PET), Kir-

gizskiy Mts. (LIN, MOS, PET), Przhevalsk, River

Chatkal (PET), Terskey Alatau Mts. (LIN, MOS), Yan-

gibazar (LIN). Japan: Honshu: Nagano (OTT), Kobe

(NHM), Sasayama (SAP), Yokohama (AEI, Hinz/

ZSM); Kyushu: Unoshima (SAP).

Variety: Temples slightly constricted behind the

eyes, 1.0x as long as the width of an eye, tangen-

tial lines on eyes and temples intersect on the

propodeum. Material (372, 983): Russia: Tomsk

Prov.: Tisul (PET); Irkutsk Prov.: Lake Baykal

(Hinz/ZSM); Chita Prov.: Chita. Kazakhstan:

Alma-Ata (PET). Tadzhikistan: Kulyab, Varzob

(MOS). Kirgiziya: Przhevalsk (PET), Terskey Ala-

tau Mts. (MOS). The status of this variety is un-

known. Apparently it occurs only in central Asia.

In one locality (Przhevalsk//Kirgiziya) it has been

collected together with the nominate variety.

Dusona juvenilis (Förster, 1868)

Description. ?: Body length 7-9 mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 0.9. Frons scarcely depressed, coriaceous, fine-

ly and densely rugose-punctate, with a fine medi-

an longitudinal carina, with few short radial wrin-

kles. Antennal carina rather low and narrow. Face

and dorsal 0.4 of the clypeus coriaceous, dull,

finely and densely rugose-punctate. Clypeus flat,

coriaceous and with fine disperse punctation ven-

trally, its apex slightly convex medially. Malar

space index 0.5. Genal carina low, slightly bent

sidewards and distinctly bent forwards, joining

the oral carina at the base of the mandible at a

right angle. Temple finely coriaceous and with

fine disperse punctation medially. Flagellum with

26-28 segments, slightly fusiform, 2" segment 2.2x,

shortest segment 0.7x, penultimate segment 0.9x

as long as wide. Pronotum coriaceous and rugose-

punctate laterally on the dorsal 0.2, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, dull, densely punctate. Scutellum

with lateral carinae on the anterior half. Prescutel-

lar groove with fine longitudinal striae. Ventral

part ofthe prepectal carina merging with the trans-

verse carina, both carinae distinctly raised, the

former deeply emarginate medially and strongly

raised submedially behind the front coxae (almost

tongue-like). Pleural part of the prepectal carina

for the greater part obliterated and replaced by

wrinkles, sometimes present subdorsally. Meso-

pleurum finely coriaceous, somewhat shining, dis-

tinctly and densely punctate. Speculum finely co-

riaceous, rather shining. Depression in front of the

speculum with with fine and long longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum coriaceous, densely rugose or

rugose-punctate. Pleural carina distinct. Areolet

rather large, short-stalked, receiving the 2" recur-

rent vein distad of the middle. Nervellus slightly

inclivous, weakly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 4.7x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to 0.6-0.7 of the hind coxae, distinctly and

rather broadly depressed, sulcus with rather short

transverse wrinkles anteriorly, with long trans-

verse wrinkles posteriorly, costulae complete and

distinct, longitudinal carinae distinct posteriorly,

more or less indicated medially. Propodeal spira-

cle rather small, 2.4x as long as wide. Spiracular

carina narrow. Spiracular index 0.5. 1” gastral

tergite 4.7x, 2"X tergite 2.2x as long as wide. Peti-

ole without glymmae, with distinct narrow lateral

areas, smooth in front of these areas. Petiolar su-

ture fine and partly obliterated. 1” gastral sternite

smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum not separated from the 3“

gastral tergite, the latter with indistinct black lat-

eral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 1.3.

Colour: Black. Maxillary palpus yellowish red.

Mandible rather narrowly marked with yellow

dorsally and medially. Tegula yellow. Wings hya-

line. Stigma yellowish brown medially. Front coxa

and trochanter marked with yellow. Front femur,

tibia and tarsus yellowish red, the femur some-

times darkened ventrally. Mid femur, tibia and

tarsus yellowish red, the femur darkened ventral-

ly, the tarsus darkened apically. Hind femur yel-

lowish red, marked with black basally (to a varia-

ble extend), or completely black. Hind tibia broadly

yellowish red medially, narrowly medium brown

basally and apically, or broadly brown medially,

narrowly black basally and apically. Hind tarsus

yellowish red basally, black apically, or complete-

ly black. 2" gastral tergite red on the posterior 0.2.

3 tergite red, sometimes marked with black pos-

teriorly (to a variable extend). Pilosity whitish.

d: Flagellum with 30-32 segments, filiform,

the apical 0.1 slightly attenuate, 2"! segment 1.9x,

shortest segment 1.0x, penultimate segment 1.1x

as long as wide. Propodeum extending to the

middle of the hind coxae, longitudinal carinae

sometimes distinct and almost complete. 2" gas-

tral tergite red on the posterior 0.1-0.2. 34 tergite

often red only anteriorly. In other characters sim-

ilar to the 9.

Material (572, 19): Russia: Altaysk Terr.: River Chu-

Iyshman (MIN); Irkutsk Prov.: Dachnaya; Yakutsk

A.S.S.R.: Arangastakh Reserve. Kazakhstan: Borovoye

near Kokchetav (PET).

Dusona kasparyani Hinz, 1985

Description. $: Body length 7 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and rather densely punctate,

with a distinet median longitudinal carina on top

of alow median longitudinal bulge, with few and

indistinct radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.8 of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.2.

Genal carina low ventrally;slightly bent sidewards,

joining the low oral carina at an acute angle. Genal

index 0.5. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

41-43 segments, filiform, the apical 0.2 attenuate,

2"! segment 2.5x, shortest segment 1.3x, penulti-

mate segment 1.7x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.1, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior 0.6. Prescutellar groove with fine

longitudinal striae. Ventral part of the prepectal

carina merging with the transverse carina, the

former low, the latter slightly raised. Pleural part

of the prepectal carina obliterated and replaced by

some short wrinkles. Pleural part of the prepectus

densely rugose-punctate on a finely coriaceous

and shining background. Mesopleurum distinctly

coriaceous, with distinct rather dense punctation.

Speculum finely coriaceous. Depression in front

of the speculum with distinct rather long longitu-

dinal striae, which merge with the punctation an-

teriorly. Metapleurum coriaceous, finely and

densely rugose-punctate. Pleural carina fine but

distinct. Areolet large, pointed or short-stalked,

receiving the 2” recurrent vein at the middle.

Nervellus slightly reclivous, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.5. Hind

femur 4.9x as long as high. Hind tarsal claws

short, with short pectinate teeth basally. Propo-

deum extending to the middle of the hind coxae,

almost not depressed, coriaceous or with short

transverse wrinkles anteriorly, with long trans-

verse wrinkles posteriorly, carinae almost com-

pletely obliterated. Propodeal spiracle rather large,

3.0x as long as wide. Spiracular carina narrow.

Spiracular index 0.4. 1“ gastral tergite 4.7x, 2

tergite 2.8x as long as wide. Petiole without glym-

mae, with slightly indicated narrow lateral areas,

smooth in front of these areas. Petiolar suture

obliterated. 1“ gastral sternite smooth. Thyridium

elongate. Thyridial index about 1.0. Epipleurum

not separated from the 3" gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.0.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellowish brown medially. Flagel-

lum for the greater part medium brown. Tegula

yellow. Wings hyaline. Stigma medium brown

medially. Front and mid legs yellow, front coxa

black basally, mid coxa black basally and medial-

ly. Hind trochanter yellow. Hind femur reddish

brown. Hind tibia yellow medially, narrowly me-

dium brown basally, broadly medium brown api-

cally. Hind tarsus yellowish brown. 2" gastral

tergite dorsally red on the posterior 0.1, laterally

red on the posterior 0.3. 3" tergite red, dorsally

marked with black anteriorly. 4" and 5" tergites

red. 6" and 7 tergites dark brown dorsally, broad-

ly marked with red ventrolaterally. Gastral ster-

nites red. Pilosity whitish.

d unknown.

Material (2??): Russia: Primorye: Khasan (Hinz /

ZSM, PET).

Dusona lamellifer Hinz, 1994

Description. ?: Body length 8mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons not depressed,

densely rugose-punctate, with distinct but short

radial wrinkles, a fine median longitudinal carina

scarcely indicated. Antennal carina low and nar-

row. Face and dorsal 0.8 of the clypeus coriaceous

and densely rugose. Clypeus flat, coriaceous and

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.4. Genal carina and

oral carina very strongly raised close to the junc-

tion of both carinae, forming a lopsided triangular

pyramid, the tip of the latter shifted ventrally and

situated behind the base of the mandible. Gena

deeply depressed close to the junction, the de-

pression about as deep as the width of the 2"

flagellar segment. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 38 segments, filiform, the apical 0.2

slightly attenuate, 2" segment 1.7x, shortest seg-

ment 0.9x, penultimate segment 1.0x as long as

wide. Pronotum coriaceous and densely rugose-

punctate laterally on the dorsal 0.3, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous and very densely punctate. Scutel-

lum with lateral carinae on the anterior 0.8. Pres-

cutellar groove for the greater part smooth, with

few longitudinal striae laterally. Prepectal carina

low, ventral part merging with the pleural part,

the latter almost complete, the dorsal 0.1-0.2 re-

placed by several wrinkles. Transverse carina ob-

literated. Mesopleurum distinctly coriaceous, rath-

er densely or densely punctate. Speculum finely

coriaceous. Depression in front of the speculum

with fine rather long longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum densely rugose-punctate. Pleural carina dis-

tinct. Areolet short-stalked, receiving the 2"d

recurrent vein slightly distad ofthe middle. Nervel-

lus slightly inclivous, distinctly intercepted. Ner-

vellar index 0.8. Tibial spur index 0.6. Hind femur

5.1x as long as high. Hind tarsal claws short, with

short pectinate teeth basally. Propodeum extend-

ing to the middle of the hind coxae, broadly and

not deeply depressed, for the greater part irregu-

larly wrinkled, with few short transverse wrin-

kles in the midline, costulae obliterated, longitu-

dinal carinae weakly indicated medially and pos-

teriorly. Propodeal spiracle rather small, 3.3x as

long as wide. Spiracular carina partly widened.

Spiracular index about 0.3. 1% gastral tergite 3.8x,

2"! tergite 1.7% as long as wide. Petiole without

glymmae, with narrow lateral areas, smooth in

front of these areas. Petiolar suture almost com-

pletely obliterated. 1* gastral sternite for the greater

part smooth, with transverse striae posteriorly.

Thyridium oval. Thyridial index about 1.0. Epi-

pleurum not separated from the 3" gastral tergite,

the latter with distinct black lateral stripes well

above the ventrolateral edge. Gaster distinctly

compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown medially and apically.

Mandible narrowly marked with reddish brown

subapically. Wings hyaline. Stigma dark brown

medially. Front leg from the femur onwards light

reddish brown, the femur marked with dark brown

ventrally. Mid femur marked with yellowish

brown on the apical 0.2. Mid tibia yellowish brown.

Mid tarsus yellowish brown to brown. Hind tibia

whitish yellow dorsally, brown ventrally, narrowly

black basally, broadly black apically. 2" gastral

tergite red on the posterior 0.3. 3" and 4" tergites

red, the 4" tergite tinged with black dorsally. Pi-

losity whitish.

d unknown.

Material (12): Japan: Honshu: Nikko (AEI).

Dusona leptogaster (Holmgren, 1860)

Description. 9: Body length 7-8mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 0.7. Frons slightly depressed, coriaceous,

with very fine and disperse punctation, with a

fine median longitudinal carina, with fine or in-

distinct radial wrinkles. Antennal carina low and

narrow. Face and dorsal half of the clypeus coria-

ceous and with very fine and dense punctation.

Clypeus slightly concave, coriaceous and with fine

disperse punctation ventrally, its apex truncate

medially. Malar space index 0.6. Genal carina low

ventrally, slightly bent sidewards, joining the

slightly raised oral carina at an acute angle. Genal

index 0.5. Temple finely coriaceous and with fine

rather dense punctation medially. Flagellum with

38-42 segments, filiform, the apical 0.2 attenuate,

2"d segment 2.3x, shortest segment 1.0x, penulti-

mate segment 1.2x as long as wide. Pronotum

with fine longitudinal wrinkles, almost not punc-

tate. Mesoscutum coriaceous, dull, with very fine

and rather dense punctation, in some parts finely

rugose-punctate. Scutellum coriaceous, densely

rugose-punctate, with lateral carinae on the ante-

rior 0.4. Prescutellar groove smooth medially,

with longitudinal striae laterally. Prepectal carina

low, ventral part merging with the pleural part,

the latter complete, joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum densely rugose centrally, coriaceous

and with fine rather dense punctation ventrally.

Speculum coriaceous, rather dull dorsally, shin-

ing ventrally. Depression in front of the speculum

dorsally with short longitudinal wrinkles, which

merge with the punctation anteriorly, the ventral

0.3 with fine wrinkles and shining. Metapleurum

densely rugose. Pleural carina distinct. Areolet

small, short-stalked, receiving the 2” recurrent

vein far distad of the middle. Nervellus almost

vertical, weakly intercepted. Nervellar index

0.7-0.8. Tibial spur index 0.6. Hind femur 5.7x as

long as high. Hind tarsal claws short, with long

pectinate teeth basally. Propodeum extending to

the middle of the hind coxae, scarcely depressed,

with complete costulae and longitudinal carinae,

for the greater part transversely wrinkled between

the longitudinal carinae, with few longitudinal

carinae posteriorly. Propodeal spiracle small, 2.8x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.6. 1% gastral tergite 4.0x, 2" tergite

1.8x as long as wide. Petiole without glymmae,

with narrow lateral areas, sometimes with very

small and shallow pits at the anterior edge of the

lateral areas, with very fine longitudinal striae in

front of these areas. Petiolar suture fine, complete.

1" gastral sternite for the greater part coriaceous,

with transverse striae posteriorly. Thyridium elon-

gate. Thyridial index about 2.0. Epipleurum sepa-

rated from the 3" gastral tergite, the crease marked

with black. Epipleural index 0.9. Gaster distinctly

compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Wings hyaline. Stig-

ma dark brown medially. Front leg from the tro-

chanter onwards yellowish red. Mid femur yel-

lowish brown on the apical 0.2. Mid tibia and

tarsus yellowish brown. Hind tibia black,slightly

tinged with dark brown. 2" gastral tergite red on

the posterior 0.2. 3° to 5" tergites red. Pilosity

whitish.

6: Tegula yellow. Front and mid legs from

the tip of the coxa onwards yellowish red. Hind

tibia dark yellowish brown. In other characters

similar to the 9.

Material (10??, 238): Russia: Irkutsk Prov.: Dach-

naya; Kuril Isl.: Alekhino on Kunashir, Krabozavodsk

on Shikotan (PET). Japan: Honshu: Kamikochi near

Norikura-dake (AEI).

Dusona levibasis Horstmann, spec. nov.

Holotype (3): Russia, Primorye, Spassk Dalniy,

8.7. 1961 (MOS).

Description. d: Body length 10 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, densely rugose-punctate, with an in-

distincet and partly obliterated median longitudi-

nal carina, with short radial wrinkles, which ex-

tend to 0.3 the distance to the median ocellus.

Antennal carina low and narrow. Face and dorsal

0.7 of the clypeus coriaceous, very densely punc-

tate and rugose-punctate. Clypeus flat, finely co-

riaceous and with rather dense punctation ven-

trally, its apex truncate medially. Malar space in-

dex 0.3. Genal carina straight ventrally, slightly

raised, joining the slightly raised oral carina at an

acute angle. Genal index 0.3. Temple fine coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 52 segments, filiform, the

apical 0.2 attenuate, Due segment 1.6%, shortest

segment 1.0x, penultimate segment 1.4x as long

as wide. Pronotum finely coriaceous and densely

rugose-punctate laterally on the dorsal 0.2, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, densely punctate. Scutel-

lum with lateral carinae on the anterior 0.7. Pres-

cutellar groove smooth medially, with fine longi-

tudinal striae laterally. Prepectal carina low, the

ventral part merging with the pleural part, the

dorsal 0.2 of the latter obliterated. Transverse car-

ina indicated anteriorly. Mesopleurum distinctly

coriaceous, rather densely or densely punctate.

Speculum granulate-strigose, scarcely shining.

Depression in front of the speculum with fine

longitudinal striae, which merge with a coriaceous

area without punctation anteriorly. Metapleurum

coriaceous, densely punctate. Pleural carina dis-

tinct. Areolet large, narrowly sessile, receiving the

2"! recurrent vein slightly distad of the middle.

Nervellus slightly reclivous, distinctly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.5. Hind

femur 4.5x as long as high. Hind tarsal claws

short, with short pectinate teeth basally. Propo-

deum extending to 0.3 of the hind coxae, deeply

and rather narrowly depressed, the longitudinal

furrow coriaceous anteriorly, with transverse stri-

ae medially and posteriorly, carinae almost com-

pletely obliterated. Propodeal spiracle large, 3.0x

as long as wide. Spiracular carina partly widened.

Spiracular index 0.5. 1* gastral tergite 3.7x, 2”

tergite 1.8x as long as wide. Petiole with distinct

glymmae, smooth in front of the glymmae. Petio-

lar suture obliterated. 1% gastral sternite smooth.

Thyridium oval. Thyridial index about 1.5. Epi-

pleurum pendent as a small triangular lobe ante-

riorly at the ventrolateral edge of the 3 gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula yellowish

brown. Wings hyaline. Stigma yellowish brown

medially. Front leg from the tip of the coxa on-

wards yellow, the femur yellowish red. Mid fe-

mur yellowish brown dorsally, black ventrally.

Mid tibia yellow, Mid tarsus yellow basally, yel-

lowish brown medially and apically. Hind tibia

broadly yellow medially, narrowly black basally,

rather narrowly medium brown apically. Hind

tarsus dark brown. 2" gastral tergite dorsally red

on the posterior 0.2, for the greater part red later-

ally, the posterior edge marked with dark brown.

3:d and 4" tergites red. Pilosity whitish.

® unknown.

Dusona liberator Hinz, 1985

Description. ?: Body length 7-8 mm. Temples

distinctly constricted behind the eyes, 0.6%x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.9. Frons scarcely depressed, coriaceous,

finely and densely rugose-punctate, with a dis-

tinct median longitudinal carina, with distinct ra-

dial wrinkles, which extend to half the distance to

the median ocellus. Antennal carina low and nar-

row. Face and dorsal 0.7 of the clypeus coriaceous,

densely rugose-punctate. Clypeus slightly con-

vex, coriaceous and with rather dense punctation

ventrally, its apex slightly convex medially. Malar

space index 0.4. Genal carina low ventrally, slight-

ly bent forwards and sidewards, joining the oral

carina at the base of the mandible at a right angle.

Temple finely coriaceous and with very fine dis-

perse punctation medially. Flagellum with 38-42

segments, filiform, the apical 0.2 slightly attenu-

ate, 2"! segment 2.3x, shortest segment 1.1x, pe-

nultimate segment 1.2x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.2, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior 0.8. Prescutellar groove with few

fine longitudinal striae. Ventral part of the pre-

pectal carina merging with the transverse carina,

both carinae strongly raised, the former deeply

emarginate medially. Pleural part of the prepectal

carina obliterated ventrally and dorsally, the me-

dian 0.4 present. Mesopleurum distinctly coria-

ceous, rather densely or densely punctate. Specu-

lum and depression in front of the speculum with

long longitudinal striae, which merge with the

punctation anteriorly. Metapleurum finely and

densely rugose-punctate. Pleural carina distinct.

Areolet rather small, long-stalked, receiving the

2" recurrent vein slightly distad of the middle.

Nervellus slightly reclivous, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 4.4x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally. Propo-

deum extending to 0.4 of the hind coxae, deeply

and rather narrowly depressed, with short and

indistinct transverse wrinkles in the longitudinal

furrow, with distinct transverse wrinkles posteri-

orly, irregularly wrinkled beside the furrow, car-

inae almost completely obliterated. Propodeal

spiracle small, 2.7x as long as wide. Spiracular

carina narrow. Spiracular index 0.3. 1" gastral

tergite 4.0x, 2”! tergite 1.8x as long as wide. Peti-

ole without glymmae or narrow lateral areas,

smooth laterally. Petiolar suture partly obliterat-

ed. 1" gastral sternite smooth. Thyridium elon-

gate. Thyridial index about 1.0. Epipleurum not

separated from the 3 gastral tergite, the latter

with fine black lateral stripes well above the ven-

trolateral edge. Gaster distinctly compressed pos-

teriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown.

Wings hyaline. Stigma yellowish brown medially.

Front femur yellowish red on the apical half. Front

and mid tibiae and tarsi yellowish red. 2” gastral

tergite red on the posterior 0.1. 3" tergite marked

with red on the anterior half. Pilosity whitish.

d: Propodeum with distinct transverse wrin-

kles in the longitudinal furrow. Maxillary palpus

reddish brown. Hind tibia sometimes whitish sub-

basally on the dorsal side. 3° gastral tergite some-

times completely red. 4" tergite sometimes marked

with red anteriorly. In other characters similar to

the 9.

Material (2??, 338): Russia: Chita Prov.: Nerchins-

kiy Zavod (PET); Khabarovsk Terr.: Khekhtsir near

Khabarovsk (Hinz/ZSM, PET); Primorye: Roshchino

on River Iman (PET), Pozhiga SO of Dalnerechensk

(Hinz/ZSM).

Dusona linearis Horstmann, spec. nov.

Holotype (3): Japan, Hokkaido, Apoi-dake, 7.7.

1957 (SAP).

Description. d: Body length 15 mm. Temples dis-

tinctly constricted behind the eyes, 0.9%x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons coriaceous and densely punctate dor-

sally, deeply depressed ventrally, coriaceous with-

out radial wrinkles in the depression, with a dis-

tinct narrow median longitudinal bulge and with

a partly obliterated fine longitudinal carina on top

of this bulge. Antennal carina strongly raised, the

rim bent upwards (ear-like), at the same time with

a distinct dorsal projection (nose-like) and with

fine transverse striae on the projection. Face and

dorsal 0.6 of the clypeus coriaceous, densely ru-

gose-punctate. Clypeus flat, coriaceous and with

disperse or rather dense punctation ventrally, its

apex slightly convex medially. Malar space index

0.5. Genal carina low and almost straight ventral-

ly, joining the distinctly raised oral carina at an

acute angle. Genal index 0.5. Temple finely coria-

ceous and with fine rather dense punctation me-

dially. Flagellum with 60 segments, filiform, the

apical 0.3 attenuate, 24 segment 1.4x, shortest

segment 1.0x, penultimate segment 1.2x as long

as wide. Pronotum finely coriaceous and densely

rugose-punctate laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, densely punctate.

Scutellum with lateral carinae on the anterior 0.3.

Prescutellar groove with longitudinal striae. Pre-

pectal carina low, ventral part merging with the

pleural part, the latter complete, joining the ante-

rior edge of the mesopleurum. Transverse carina

indicated anteriorly. Mesopleurum finely coria-

ceous, with distinct and rather dense or dense

punctation. Speculum and depression in front of

the speculum coriaceous, without wrinkles. Meta-

pleurum coriaceous, dull, finely and densely punc-

tate. Pleural carina strong. Areolet large, short-

stalked, receiving the 2" recurrent vein at the

middle. Nervellus slightly inclivous, distinctly

intercepted. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 5.6x as long as high. Hind tarsal

claws rather long, with some short pectinate teeth

basally. Propodeum extending to 0.4 of the hind

coxae, broadly and deeply depressed, with trans-

verse wrinkles in the sulcus, costulae complete,

longitudinal carinae almost complete, obliterated

only anteriorly. Propodeal spiracle rather small,

2.5x as long as wide. Spiracular carina narrow.

Spiracular index 0.6. 1" gastral tergite 4.3x, 2"

tergite 1.9x as long as wide. Petiole with rather

large glymmae at the anterior edge of narrow

lateral areas, with very fine sculpture in front of

the glymmae and with some transverse wrinkles

anteriorly. Petiolar suture almost complete. 1“

gastral sternite coriaceous anteriorly, transversely

striate posteriorly. Thyridium oval. Thyridial in-

dex about 1.5. Epipleurum separated from the 3"

gastral tergite, the crease not marked with black,

but the tergite with black lateral stripes well above

the ventrolateral edge. Epipleural index 0.6. Gaster

strongly compressed posteriorly.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellow dorsally.

Tegula yellowish brown. Wings slightly darkened.

Stigma medium brown medially. Front trochant-

er yellowish brown. Front femur, tibia and tarsus

yellowish red. Mid femur yellowish red on the

apical 0.3. Mid tibia yellow. Hind tibia yellow,

narrowly black basally and apically. 2“ gastral

tergite red on the posterior 0.2. 3“ and 4" tergites

red. Pilosity whitish.

? unknown.

Dusona lividariae Hinz, 1963

Description. 2: Body length 11-12 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior part of the mes-

oscutum. Ocellar index 0.7. Frons scarcely de-

pressed, for the greater part rugose, with a dis-

tinct median longitudinal carina, with few short

radial wrinkles. Antennal carina slightly raised

(inconspicuously) and narrow. Face and clypeus

densely rugose. Clypeus flat, with a narrow al-

most smooth area above the apex, the latter slight-

ly convex medially. Malar space index 0.3. Genal

carina strongly raised close to the junction with

the also raised oral carina, forming a lopsided

triangular pyramid, the tip of the latter shifted

ventrally and situated behind the base of the man-

dible. Gena deeply depressed close to the junc-

tion, the depression about as deep as the width of

the 2" flagellar segment. Genal index about 0.8.

Temple coriaceous and with distinct rather dense

or dense punctation medially. Flagellum with

49-51 segments, filiform, the apical 0.3 attenuate,

2”X segment 1.4x, shortest segment 0.7x, penulti-

mate segment 0.9x as long as wide. Pronotiım

very densely punctate on a smooth background

laterally on the dorsal 0.6, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum dense-

ly rugose-punctate, partly with a smooth back-

ground. Scutellum with lateral carinae on the

anterior half. Prescutellar groove with fine longi-

tudinal wrinkles. Ventral part of the prepectal

carina merging with the transverse carina, both

carinae distinctly raised, the former distinctly

emarginate medially. Pleural part of the prepectal

carina obliterated ventrally and dorsally, the me-

dian 0.3 present. Mesopleurum strongly and

densely punctate on a smooth background. Specu-

lum punctate posteriorly, with oblique wrinkles

anteriorly. Depression in front of the speculum

with distinct longitudinal striae, which merge with

the punctation anteriorly. Metapleurum distinct-

ly and densely rugose, the wrinkles smooth. Pleu-

ral carina strong. Areolet large, short-stalked, re-

ceiving the 2" recurrent vein distad of the middle.

Nervellus almost vertical, distinctly intercepted.

Nervellar index 0.7. Tibial spur index 0.7. Hind

femur 5.1x as long as high. Hind tarsal claws long,

with strong pectinate teeth almost over their total

length. Propodeum extending to 0.6 of the hind

coxae, broadly and not deeply depressed, for the

greater part irregularly wrinkled, with transverse

wrinkles posteriorly, carinae almost completely

obliterated. Propodeal spiracle large, 2.9x as long

as wide. Spiracular carina narrow. Spiracular in-

dex 0.4. 1“ gastral tergite 5.1x, 2" tergite 1.9x as

long as wide. Petiole without glymmae, often with

narrow lateral areas, smooth laterally. Petiolar

suture for the greater part obliterated. 1* gastral

sternite smooth. Thyridium oval. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter with distinct or indistinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellow. Man-

dible narrowly marked with yellow dorsally and

medially. Flagellum light brown apically. Tegula

yellow. Wings hyaline. Stigma yellowish brown

medially. Front leg from the trochanter onwards

yellow. Mid leg from the femur onwards yellow.

Hind tibia completely yellow. Hind tarsus dark

brown. 2” gastral tergite black, with small lateral

yellow marks posteriorly. 3" tergite yellow. 4"

tergite marked with yellow on the anterior 0.7.

Pilosity whitish.

d: Flagellum brown to black apically. Tip of

the front coxa, mid trochanter and hind tarsus

yellow. In other characters similar to the 2.

Material (4??, 233): Tadzhikistan: Pamir Mts., 4000 m.

Kirgiziya: Ferganskiy Mts. (MOS), Issykata (PET), Sos-

novka, 1500-1800 m (LIN).

Dusona longicauda (Uchida, 1928)

This is a rather variable species.

Description. ?: Body length 9-12 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and densely punctate, with a

fine median longitudinal carina, with distinct ra-

dial wrinkles, which extend to half the distance to

the median ocellus. Antennal carina low and nar-

row. Face and dorsal 0.3 of the clypeus coriaceous,

densely punctate. Clypeus flat, coriaceous and

rather densely punctate ventrally, its apex trun-

cate medially. Malar space index 0.4. Genal carina

low and rather straight ventrally, joining the scarce-

ly raised oral carina at an acute angle. Genal index

0.5. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with

40-48 segments, filiform, the apical 0.3 attenuate,

2rd segment 2.3x, shortest segment 1.0x, penulti-

mate segment 1.5x as long as wide. Pronotum

coriaceous and densely rugose-punctate laterally

on the dorsal 0.2, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous, dull,

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove with very

fine sculpture. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina obliterated. Mesopleurum fine-

ly coriaceous, with rather fine and rather dense

punctation. Speculum coriaceous, granulate-stri-

gose. Depression in front of the speculum with

distinct rather short longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum coriaceous, densely rugose. Pleural carina

distinct. Areolet large, pointed or short-stalked,

receiving the 2” recurrent vein basad of the mid-

dle. Nervellus reclivous, distinctly intercepted.

Nervellar index 0.6. Tibial spur index 0.6. Hind

femur 5.9x as long as high. Hind tarsal claws

rather long, with distinct pectinate teeth basally

and medially. Propodeum extending to the mid-

dle of the hind coxae, slightly and rather narrowly

depressed, with transverse wrinkles between the

longitudinal carinae, costulae and longitudinal car-

inae often complete and distinct, sometimes part-

ly indistinct. Propodeal spiracle large, 3.8x as long

as wide. Spiracular carina narrow or widened.

Spiracular index 0.3. 1* gastral tergite 5.4x, 2"

tergite 2.5x as long as wide (in some specimens

gastral tergites shorter: 1" tergite 4.1x, 2" tergite

1.4x as long as wide). Petiole with distinct glym-

mae, smooth or with very fine sculpture in front

of the glymmae. Petiolar suture fine, sometimes

partly obliterated. 1“ gastral sternite finely coria-

ceous, with fine transverse striae posteriorly. Thy-

ridium elongate. Thyridial index 1.0-1.5. Epipleu-

rum separated from the 3" gastral tergite, the

crease marked with black. Epipleural index 0.8.

Gaster strongly compressed posteriorly. Oviposi-

tor index 1.4.

Colour: Black. Maxillary palpus yellowish

brown. Mandibel broadly yellow medially. Wings

hyaline. Stigma dark brown medially. Front leg

from the femur onwards yellowish brown, the

femur marked with brown ventrally. Mid femur

yellowish brown on the apical 0.2. Mid tibia and

tarsus yellowish brown. Hind tibia blackish, some-

times tinged with yellowish brown medially on

the dorsal side. 2” gastral tergite red on the poste-

rior 0.3. 3” and 4" tergites red, sometimes marked

with black dorsally. Pilosity whitish.

d: Maxillary palpus yellow. Front and mid

tibiae and tarsi yellow. Hind tibia sometimes dis-

tinctly marked with yellowish brown subbasally.

2"! gastral tergite red on the posterior 0.1-0.2. 5" to

7" tergites sometimes with large and distinct yel-

low dorsal marks anteriorly (these marks some-

times covered by the preceeding tergites and in-

visible). In other characters similar to the 2.

Material (182%, 1188): Russia: Khabarovsk Terr.:

Bikin (PET), Khekhtsir near Khabarovsk (Hinz/ZSM,

PET), Troitskoye; Primorye: Khasan, Zarubino; Sakha-

lin: Ozerskiy (PET). Japan: Hokkaido: Moiwa, Sappo-

ro, Teine (SAP); Honshu: Asaka (OTT), Kamikochi

near Norikura-dake (AEI), Kyoto, Mt. Oginosen in

Tottori-ken, Mt. Shirago in Hyogo-ken (SAP), Nagano

(OTT), Tokyo, Yokohama (AEI). The species also oc-

curs in the Oriental region: China, Fukien Prov. (= Fu-

jian Prov.), Shaowu, 1200-1500 m (52%, 18 AEI, 12

Hinz/ ZSM).

Dusona longigena Horstmann, spec. nov.

Holotype (3): Kazakhstan, Ketmen Mts., 2100 m,

14.6.1998 (LIN),

Description. d: Body length 13 mm. Temples dis-

tinctly but not strongly constricted behind the

eyes, 1.0x as long as the width of an eye, tangen-

tial lines on eyes and temples intersect on the

anterior part of the propodeum. Ocellar index 1.0.

Frons not depressed, coriaceous, densely rugose-

punctate, without a median longitudinal carina,

with short radial wrinkles. Antennal carina low

and narrow dorsally, but the antennal sockets

widened in dorsomedian direction, with fine cir-

cular striae on the inner side, both antennal cari-

nae almost touching each other medially. Face

and clypeus finely coriaceous, very densely punc-

tate. Clypeus flat, its apex convex medially. Malar

space index 0.7. Genal carina low and almost

straight ventrally, joining the distinctly raised oral

carina at an acute angle. Genal index 0.5. Temple

finely coriaceous and with fine very dense punc-

tation medially. Flagellum with 47-48 segments,

filiform, the apical 0.3 attenuate, 2" segment 1.3x,

shortest segment 0.9x, penultimate segment 1.3x

as long as wide. Pronotum densely rugose-punc-

tate on a smooth background laterally on the dor-

sal 0.2, with fine irregular and longitudinal wrin-

kles ventrally. Mesoscutum densely punctate on a

finely coriaceous or almost smooth background.

Scutellum densely rugose-punctate, with lateral

carinae on the anterior 0.4. Prescutellar groove

almost smooth. Prepectal carina low, ventral part

joining the transverse carina, the former not emar-

ginate medially, the latter low. Pleural part of the

prepectal carina completely obliterated. Pleural

part of the prepectus with dense punctation on a

smooth background, without wrinkles. Mesopleu-

rum with distinct and dense punctation on a

smooth background. Speculum for the greater part

with fine oblique striae, the ventral 0.2 finely co-

riaceous and shining. Depression in front of the

speculum with fine and short longitudinal wrin-

kles, which merge with the punctation anteriorly.

Metapleurum densely rugose. Pleural carina fine,

complete. Areolet large, short-stalked, receiving

the 2” recurrent vein distinctly basad of the mid-

dle. Nervellus slightly inclivous, weakly inter-

cepted. Nervellar index 0.8. Tibial spur index 0.8.

Hind femur 5.7x as long as high. Hind tarsal claws

long and rather large, with short pectinate teeth

basally. Propodeum extending to 0.4 of the hind

coxae, not depressed, almost without carinae, with

fine irregular wrinkles. Propodeal spiracle small,

3.6x as long as wide. Spiracular carina obliterated.

Spiracular index 1.2. 1* gastral tergite 4.9x, 2"«

tergite 2.4x as long as wide. Petiole without glym-

mae, narrow lateral areas scarcely indicated,

smooth in front of these areas. Petiolar suture for

the greater part obliterated. 1* gastral sternite for

the greater part smooth, with fine transverse stri-

ae posteriorly. Thyridium elongate. Thyridial in-

dex about 1.5. Epipleurum not separated from the

3" gastral tergite, the latter with black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly.

Colour: Black. Mandible narrowly yellowish

brown dorsally and medially. Wings hyaline. Stig-

ma black medially. Legs completely black. 2"' gas-

tral tergite red on the posterior 0.1. 3’ and 4*

tergites red, the 4" narrowly tinged with black

posteriorly. Pilosity whitish, hairs conspicuously

long.

? unknown.

Dusona longistilus Horstmann, spec. nov.

Holotype (4): Japan, Honshu, Tokyo, 8.5.1961

(GEN)

Description. d: Body length 7 mm. Temples dis-

tinctly constricted behind the eyes, 0.6x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons not depressed, coriaceous, finely and

dispersely punctate, partly with fine irregular

wrinkles, with a fine median longitudinal carina,

with short radial wrinkles. Antennal sockets close

to each other. Antennal carina low and narrow.

Face and dorsal 0.3 of the clypeus coriaceous,

finely and densely punctate. Clypeus flat, coria-

ceous, with disperse or rather dense punctation

ventrally, its apex slightly convex medially. Malar

space index 0.4. Genal carina low and almost

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.3. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 35 segments, fili-

form, the apical 0.3 slightly attenuate, 2” segment

2.0x, shortest segment 1.3x, penultimate segment

1.3x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.2, with longitu-

dinal wrinkles ventrally. Mesoscutum coriaceous,

finely and densely punctate. Scutellum coriaceous,

rather densely punctate, with lateral carinae on

the anterior 0.3. Prescutellar groove smooth. Pre-

pectal carina low, ventral part joining the pleural

part, the dorsal 0.3 of the latter obliterated. Trans-

verse carina indicated. Mesopleurum distinctly

coriaceous, densely punctate. Speculum granu-

late-strigose, somewhat shining. Depression in

front of the speculum with fine and short longitu-

dinal wrinkles, which merge with a narrow coria-

ceous area without punctation anteriorly. Meta-

pleurum coriaceous, finely and densely rugose-

punctate and rugose. Pleural carina distinct.

Areolet large, pointed, receiving the 2"? recurrent

vein distad of the middle. Nervellus distinctly

inclivous, weakly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 5.7x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to the mid-

dle of the hind coxae, with a narrow and shallow

longitudinal furrow, with short transverse wrin-

kles in the furrow, irregularly wrinkled in other

parts, with some longitudinal wrinkles posterior-

ly, carinae almost completely obliterated. Propo-

deal spiracle small, 2.0x as long as wide. Spiracu-

lar carina narrow. Spiracular index 1.0. 1% gastral

tergite 5.2x, 2" tergite 1.5x as long as wide (the

latter perhaps somewhat widened as a conse-

quence of the mounting process). Petiole with

small but distinct glymmae, conspicuously nar-

rowed behind the glymmae, the latter situated at

the anterior edge of narrow lateral areas, petiole

smooth in front of the glymmae. Petiolar suture

fine. 1* gastral sternite for the greater part smooth,

with fine transverse striae posteriorly. Thyridium

oval. Thyridial index about 1.5. Epipleurum not

separated from the 3% gastral tergite, the latter

with black lateral stripes well above the ventrola-

teral edge. Gaster distinctly compressed posteri-

orly.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly marked with yellow medially. Tegu-

la yellow. Wings hyaline. Stigma dark brown me-

dially. Front leg from the tip of the coxa onwards

yellowish red. Mid leg from the trochanter on-

wards yellowish red, the trochanter marked with

brown. Hind femur for the greater part reddish

brown, tinged with dark brown basally, dorsally

and apically. Hind tibia black, tinged with dark

brown medially. Hind tarsus brownish black. 2”!

gastral tergite red on the posterior 0.7. 3“ and 4"

tergites red. Pilosity whitish.

? unknown.

Dusona mactatoides Hinz 1994

Description. 2: Body length 15-17 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons deeply depressed above the antennal

carinae, coriaceous and with fine rather dense

punctation dorsally, coriaceous and with distinct

rather long radial wrinkles in the depressions, the

wrinkles extending to half the distance to the

median ocellus, with a cone-like projection be-

tween the antennal sockets, which merges with a

distinct median longitudinal bulge dorsally. An-

tennal carina strongly raised, the rim bent up-

wards (ear-like) and at the same time with a dor-

sal projection (nose-like), with fine transverse stri-

ae on the rim and on the projection. Face and

dorsal 0.4 of the clypeus coriaceous, densely punc-

tate. Clypeus flat, coriaceous and rather densely

punctate ventrally, its apex truncate medially.

Malar space index 0.4. Genal carina low and rath-

er straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 56-62 seg-

ments, filiform, the apical 0.3 distinctly attenuate,

Dad segment 1.6x, shortest segment 1.0x, penulti-

mate segment 1.4x as long as wide. Pronotum

coriaceous and with fine very dense punctation

laterally on the dorsal 0.3, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, finely and densely punctate. Scutellum with

lateral carinae on the anterior 0.3. Prescutellar

groove with fine and dense longitudinal striae.

Prepectal carina low, ventral part merging with

the pleural part, the latter complete, joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum finely coriaceous,

with rather dense or dense punctation. Speculum

coriaceous, somewhat shining. Depression in front

of the speculum for the greater part coriaceous,

without striae, sometimes with some short irregu-

lar wrinkles dorsally. Metapleurum coriaceous,

densely punctate, partly rugose-punctate. Pleural

carina distinct. Areolet large, pointed or short-

stalked, receiving the 2" recurrent vein at or ba-

sad of the middle. Nervellus vertical, distinctly

intercepted. Nervellar index 0.7. Tibial spur index

0.5. Hind femur 5.5x as long as high. Hind tarsal

claws rather long, with short pectinate teeth ba-

sally. Propodeum extending to 0.4 of the hind

coxae, broadly and rather deeply depressed, sul-

cus for the greater part with transverse wrinkles,

costulae and longitudinal carinae almost complete.

Propodeal spiracle rather small, 4.0x as long as

wide. Spiracular carina narrow. Spiracular index

0.5. 1" gastral tergite 4.3x, 2"! tergite 2.2x as long

as wide. Petiole with rather large glymmae, smooth

in front of the glymmae. Petiolar suture fine, part-

ly obliterated. 1" gastral sternite with fine sculp-

ture, with transverse striae posteriorly. Thyrid-

ium oval. Thyridial index about 1.5. Epipleurum

separated from the 3“ gastral tergite anteriorly,

folded in or folded back secondarily and pendent

(sometimes inconspicuous), the crease (or bulge)

not marked with black, a black lateral stripe situ-

ated well above the ventrolateral edge. Epipleural

index 0.2-0.3. Gaster strongly compressed posteri-

orly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown

to black. Tegula dark brown. Wings slightly infus-

cate. Stigma medium brown medially. Front leg

from the femur onwards yellowish red. Mid fe-

mur yellowish red on the apical 0.2. Mid tibia

yellowish red. Hind tibia broadly yellow medial-

ly, narrowly black basally and apically. 2" gastral

tergite red on the posterior 0.2. 3° and 4" tergites

red. Pilosity whitish.

d: Dorsal 0.2 of the pleural part of the prepec-

tal carina sometimes replaced by several wrin-

kles. Front trechanter yellowish red. In other char-

acters similar to the ®.

Material (4?%, 235): Russia: Kuril Isl.: Volcano Go-

lovnina on Kunashir (PET), Volcano Tata on Kunashir,

1887 m (dead in the ash at the edge of the crater)

(Hinz/ZSM, MOS).

Dusona mactator (Förster, 1868)

Description. ?: Body length 16-18 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 1.0. Frons distinctly depressed ventrally,

coriaceous, finely and rather densely punctate

dorsally, with distinct radial wrinkles in the de-

pression, the wrinkles extending to half the dis-

tance to the median ocellus, with a fine and indis-

tinct median longitudinal carina. Antennal carina

distinctly raised, with a dorsal projection (nose-

like), with fine transverse striae on the projection.

Face and dorsal half of the clypeus coriaceous,

densely punctate. Clypeus flat, coriaceous and

with rather dense punctation ventrally, its apex

truncate medially. Malar space index 0.5. Genal

carina low and straight ventrally, joining the dis-

tinctly raised oral carina at an acute angle. Gena

slightly depressed close to the junction. Genal

index 0.5. Temple finely coriaceous and with fine

disperse punctation medially. Flagellum with 59-63

segments, filiform, the apical half distinctly atten-

uate, 2"d segment 1.4x, shortest segment 1.0x, pen-

ultimate segment 1.2x as long as wide. Pronotum

finely coriaceous and finely and densely rugose-

punctate laterally on the dorsal 0.4, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, dull, very densely punctate. Scutel-

lum with lateral carinae on the anterior 0.3.

Prescutellar groove with fine longitudinal striae.

Prepectal carina low, ventral part merging with

the pleural part, the latter almost complete, the

dorsal 0.2 replaced by several wrinkles. Trans-

verse carina present only anteriorly. Mesopleu-

rum finely coriaceous, distinctly and rather densely

punctate. Speculum coriaceous, shining. Depres-

sion in front of the speculum for the greater part

coriaceous, without striae, with few short wrin-

kles dorsally. Metapleurum coriaceous, finely and

densely rugose and rugose-punctate. Pleural car-

ina distinct. Areolet large, short-stalked, receiving

the 2" recurrent vein distinctly basad of the mid-

dle. Nervellus slightly inclivous, strongly inter-

cepted. Nervellar index 0.7. Tibial spur index 0.6.

Hind femur 5.5x as long as high. Hind tarsal claws

rather long, with short pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

distinctly and rather broadly depressed, sulcus

for the greater part with transverse wrinkles, cos-

tulae and longitudinal carinae almost complete,

but partly covered with wrinkles. Propodeal spir-

acle rather small, 4.5x as long as wide. Spiracular

carina narrow. Spiracular index 0.4. 1% gastral

tergite 4.7x, 2X tergite 2.2x as long as wide. Peti-

ole with distinct but rather small glymmae, smooth

in front of the glymmae. Petiolar suture almost

completely obliterated. 1 gastral sternite for the

greater part smooth, with transverse striae poste-

riorly. Thyridium oval. Thyridial index about 1.5.

Epipleurum separated from the 3" gastral tergite

anteriorly, folded in or folded back secondarily

and pendent (sometimes inconspicuous), the

crease (or bulge) not marked with black, a black

lateral stripe situated well above the ventrolateral

edge. Epipleural index 0.2-0.3. Gaster strongly

compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellow dorsally

and medially. Tegula yellow. Wings hyaline. Stig-

ma light brown medially. Front leg from the tip of

the coxa onwards yellowish red, tibia and tarsus

yellow. Mid femur yellowish red on the apical

half. Mid tibia yellowish brown to brown. Hind

femur yellowish red on the apical 0.1. Hind tibia

broadly yellow medially, narrowly and indistinctly

marked with brown basally and apically. Mid and

hind tarsi dark brown. 2” gastral tergite red on

the posterior 0.3. 3° and 4" tergites red. 5" tergite

sometimes narrowly marked with red anteriorly.

Pilosity whitish.

6: Petiolar suture complete. 1% gastral sterni-

te with fine sculpture. In other characters similar

to the 9.

Material (13): Japan: Hokkaido: Apoi-dake (SAP).

Dusona maruyamae Hinz, 1994

Description. 2: Body length 8-9 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous, partly

with fine punctures, with a fine and partly obliter-

ated median longitudinal carina, with few short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.4 of the clypeus coriaceous,

finely rugose and finely punctate. Clypeus flat or

slightly convex, coriaceous and very finely and

rather densely punctate ventrally, with a narrow

smooth area in front of the apex, the latter slightly

convex medially. Malar space index 0.3. Genal

carina low and straight ventrally, joining the slight-

ly raised oral carina at an acute angle. Genal index

0.3. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with

32-36 segments, slightly fusiform, 2" segment 1.8x,

shortest segment 0.8x, penultimate segment 0.9x

as long as wide. Pronotum coriaceous and finely

and densely rugose-punctate laterally on the dor-

sal 0.4, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, dull, with fine

rather dense or dense punctation. Scutellum with

lateral carinae on the anterior 0.4. Prescutellar

groove almost completely smooth. Prepectal cari-

na low, ventral part merging with the pleural

part, the dorsal 0.1-0.2 of the latter replaced by

several wrinkles. Transverse carina obliterated.

Mesopleurum with distinct and rather dense or

dense punctation on a finely coriaceous and shin-

ing background. Speculum punctate posteriorly,

very finely coriaceous and shining anteriorly. De-

pression in front of the speculum not coriaceous,

with fine longitudinal striae, which merge with

the punctation anteriorly. Metapleurum very

densely punctate, partly rugose-punctate, on a

very finely coriaceous and shining background.

Pleural carina complete. Areolet rather small,

short-stalked, receiving the 2" recurrent vein dis-

tad of the middle. Nervellus slightly inclivous,

weakly intercepted. Nervellar index 0.7-0.8. Tibial

spur index 0.5. Hind femur 4.7x as long as high.

Hind tarsal claws short, with dinstinct pectinate

teeth basally and medially. Propodeum extending

to 0.3 of the hind coxae, shallowly depressed, for

the greater part irregularly wrinkled, coriaceous

and somewhat shining anteriorly, with some lon-

gitudinal wrinkles posteriorly, carinae for the

greater part obliterated. Propodeal spiracle rather

small, 2.0x as long as wide. Spiracular carina wid-

ened. Spiracular index 0.5. 1“ gastral tergite 3.3x,

2" tergite 1.3x as long as wide. Petiole with dis-

tinct but rather small glymmae, smooth in front

of the glymmae. Petiolar suture fine, complete.

1" gastral sternite for the greater part smooth, with

few transverse striae posteriorly. Thyridium oval.

Thyridial index about 1.0. Epipleurum separated

from the 3" gastral tergite, the crease marked with

black. Epipleural index 0.6-0.7. Gaster distinctly

compressed posteriorly. Ovipositor index 0.8.

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Mandible

broadly yellow medially. Tegula yellowish brown.

Wings hyaline. Stigma yellowish brown. Front leg

from the femur onwards yellowish red, the femur

marked with black basally. Mid femur yellowish

red on the apical 0.2. Mid tibia and tarsus yellow-

ish red. Hind tibia broadly dark yellowish brown

or dark brown medially, narrowly black basally,

scarcely darkened apically. Hind tarsus dark

brown. 2" gastral tergite dorsally red on the pos-

terior 0.2, laterally red on the posterior 0.5. 3° and

4 tergites red. 5" to 7" tergites black dorsally,

broadly red ventrolaterally (almost half red in

lateral view). Gastral sternites red. Pilosity whitish.

d unknown.

Material (3%): Russia: Khabarovsk Terr.: Khekhtsir

near Khabarovsk (PET); Sakhalin: Novoaleksandrovsk

(Hinz/ZSM). Japan: Hokkaido: Maruyama (SAP).

Dusona maruyamator Hinz, 1979

Description. 2: Body length 6-7 mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.0. Frons scarcely depressed, coriaceous, dull,

finely and densely rugose-punctate, with a fine

and partly obliterated median longitudinal cari-

na, without radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.4 of the clypeus

coriaceous, finely and densely rugose-punctate.

Clypeus almost flat, coriaceous and with very fine

disperse or rather dense punctation ventrally, with

a narrow smooth area in front of the apex, the

latter slightly convex medially. Malar space index

0.3. Genal carina low and straight ventrally, join-

ing the low oral carina at an acute angle. Genal

index 0.2. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

34-38 segments, filiform, the apical 0.2 slightly

attenuate, 2"! segment 2.3x, shortest segment 0.9x,

penultimate segment 1.2x as long as wide. Prono-

tum coriaceous and with fine and dense puncta-

tion laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum coriaceous, rath-

er dull, densely punctate. Scutellum coriaceous,

slightly shining, rather densely punctate, with lat-

eral carinae on the anterior 0.3. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter almost com-

plete or complete, the dorsal 0.1-0.2 sometimes

replaced by several wrinkles. Transverse carina

obliterated. Mesopleurum distinctly coriaceous,

with rather fine and rather dense or dense puncta-

tion. Speculum very finely coriaceous, shining.

Depression in front of the speculum with fine and

long longitudinal striae, which merge with the

punctation anteriorly. Metapleurum coriaceous,

finely and densely punctate. Pleural carina dis-

tinct. Areolet rather small, pointed, receiving the

2"d recurrent vein distad of the middle. Nervellus

inclivous, weakly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 4.8x as long

as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, almost not

depressed, coriaceous, dull, partly very finely ru-

gose, with some longitudinal wrinkles posterior-

ly, carinae almost completely obliterated. Propo-

deal spiracle rather small, 2.1x as long as wide.

Spiracular carina narrow. Spiracular index 0.8.

1* gastral tergite 3.5x, 2" tergite 1.6x as long as

wide. Petiole with distinct glymmae, smooth in

front of the glymmae. Petiolar suture fine, com-

plete. 1 gastral sternite for the greater part smooth,

with few transverse striae posteriorly. Thyridium

elongate. Thyridial index about 2.0. Epipleurum

separated from the 3 gastral tergite, the crease

marked with black. Epipleural index 0.8-0.9. Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadliy yellow medially. Flagellum for

the greater part medium brown. Tegula yellow.

Wings hyaline. Stigma yellowish brown medially.

Front leg from the trochantellus onwards yellow,

the femur marked with black basally. Mid femur

yellow on the apical 0.2. Mid tibia and tarsus

yellow. Hind tibia dark yellowish brown subba-

sally, narrowly black basally, very broadliy black

apically (black apical ring extending over 0.3 of

the tibia). 2”! gastral tergite red on the posterior

0.1. 3° to 7" tergites black dorsally, broadly red

ventrolaterally (often about half red in lateral

view). Gastral sternites red. Pilosity whitish.

d: Flagellum with 36-42 segments, light brown

ventrally, dark brown dorsally. Front and mid

legs from the tip of the coxa onwards yellow.

Hind trochanter and base of hind femur yellow.

Hind tibia yellowish brown medially, narrowly

black basally, broadly black apically. Genital

clasper reddish brown. In other characters similar

to the 2.

Material (22?2, 2238): Russia: Khabarovsk Terr.:

Khekhtsir near Khabarovsk; Primorye: Barabash,

Khasan, Kiyevka (PET), Plastun (KIS, PET), Roshchino

on River Iman, Sokolovka, Ussuriysk, Vladivostok;

Kuril Isl.: Golovnino and Sernovodsk on Kunashir,

Malokurilsk on Shikotan (PET). Japan: Hokkaido:

Maruyama (Hinz/ZSM, SAP), Sapporo (AEI); Hon-

shu: Nagano (Hinz/ZSM, OTT).

Dusona matsumurae (Uchida, 1928)

Description. 2: Body length 16-17 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.2. Frons very deeply de-

pressed above the antennal carinae (funnel-like),

coriaceous and densely punctate dorsally, with

distinct radial winkles in the depressions, the

wrinkles extending to half the distance to the

median ocellus, with a median longitudinal bulge

between the depressions. Antennal carina distinct-

ly raised, the rim bent upwards (ear-like), forming

a narrow smooth crescent-shaped plate. Face and

dorsal 0.6 of the clypeus coriaceous, coarsely and

very densely punctate. Clypeus slightly concave,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low ventrally, slightly bent sidewards,

joining the distinctly raised oral carina at an

acute angle. Genal index 0.6. Temple finely coria-

ceous and with fine but distinct disperse or rather

dense punctation medially. No ? with complete

antennae available (flagellum of d see below).

Pronotum coriaceous and finely and densely ru-

gose-punctate laterally on the dorsal half, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, with dense or very

dense punctation. Scutellum with lateral carinae

on the anterior 0.3. Prescutellar groove with fine

longitudinal striae. Prepectal carina low, ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

with disperse or rather dense punctation on a

finely coriaceous and shining background. Specu-

lum punctate posteriorly, finely coriaceous and

shining anteriorly. Depression in front ofthe specu-

lum dorsally with distinet longitudinal striae,

which merge with the punctation anteriorly, the

ventral 0.3-0.6 coriaceous, without striae. Meta-

pleurum coriaceous, very densely punctate. Pleu-

ral carina distinct. Areolet large, short-stalked,

receiving the 2" recurrent vein at or basad of the

middle. Nervellus inclivous, strongly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 6.3x as long as high. Hind tarsal claws

rather short, with short pectinate teeth basally

and medially. Propodeum extending to 0.4 of the

hind coxae, rather broadly and rather deeply de-

pressed, sulcus for the greater part with trans-

verse wrinkles, costulae partly present, longitudi-

nal carinae almost complete. Propodeal spiracle

large, 3.2x as long as wide. Spiracular carina nar-

row. Spiracular index 0.4. 1" gastral tergite about

4.6x, 2"d tergite 2.3x as long as wide. Petiole with

distinct glymmae, smooth or with very fine sculp-

ture in front of the glymmae, sometimes with few

transverse striae basally. Petiolar suture fine, com-

plete or partly obliterated. 1% gastral sternite co-

riaceous anteriorly, smooth medially, transverse-

ly striate posteriorly. Thyridium oval. Thyridial

index about 2.0. Epipleurum not separated from

the 3'd gastral tergite, the latter without black lat-

eral stripes. Gaster strongly compressed posteri-

orly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with dark brown me-

dially. Tegula dark brown. Wings slightly infus-

cate. Stigma light brown medially. Front femur

yellow on the apical 0.6. Front tibia and tarsus

yellow. Mid femur yellow on the apical 0.2. Mid

tibia yellow. Mid tarsus brown. Hind tibia yellow-

ish brown medially, narrowly brown basally,

broadly brown apically. 2" gastral tergite dorsal-

ly black or red on the posterior 0.3, for the greater

part red laterally. 3°“ and 4" tergites red, some-

times narrowly marked with black dorsally. Pilos-

ity whitish, sometimes pilosity of the face partly

light brown.

d: Flagellum with 69 segments, filiform, the

apical 0.2 distinctly attenuate, 2" segment 1.4x,

shortest segment 0.9x, penultimate segment 1.4x

as long as wide. Sulcus of the propodeum for the

greater part irregularly wrinkled. Mesopleurum

densely punctate on a smooth background. In

other characters similar to the ®%.

Material (622, 13): Russia: Amur Prov.: Novorossiy-

ka (PET); Primorye: Okeanskaya near Vladivostok

(Hinz / ZSM), Kamen-Rybolov on Lake Khanka (PET).

Japan: Hokkaido: Sapporo (SAP); Honshu: Kamikochi

near Norikura-dake (AEI), Karuizawa (HEL).

Dusona melanator Hinz, 1985

Dusona melanator Hinz, 1994

Hinz (1985: 305 f., 1994: 34) described this species

twice (see introduction). The descriptions are al-

most identical, but the type series are different.

The holotype (?) of the 1994 description from

Sahmanbe on Etorofu (= Iturup/Kuril Isl./Rus-

sia) was apparently unavailable in 1985; it is lost

now. The paratype (d) of the 1994 description

(also from Etorofu = Iturup) (SAP) is mentioned

as paratype in the 1985 description; it is still pre-

served. The holotype (2) of the 1985 description

from Kamenushka (SE of Ussuriysk/ Primorye/

Russia) (PET) has recently been studied by the

junior author.

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons slightly depressed, coriaceous,

finely and densely rugose-punctate, with a fine

median longitudinal carina, with few short radial

wrinkles. Antennal carina low and narrow. Face

and dorsal 0.7 of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely punctate ventrally, its apex slightly

convex or truncate medially. Malar space index

0.4. Genal carina low and almost straight ventral-

ly, joining the slightly raised oral carina at an

acute angle. Genal index 0.4. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 38-43 segments, filiform,

the apical 0.3 slightly attenuate, 2’ segment 2.0x,

shortest segment 1.0x, penultimate segment 1.0x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.1, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, densely punctate. Scutellum with lat-

eral carinae on the anterior 0.3. Prescutellar groove

100

almost smooth. Prepectal carina low, ventral part

merging with the pleural part, the latter almost

complete, the dorsal 0.1-0.2 replaced by several

wrinkles. Transverse carina obliterated. Mesopleu-

rum distinctly coriaceous, densely punctate. Specu-

lum finely coriaceous and granulate-strigose, shin-

ing. Depression in front of the speculum with fine

but distinct longitudinal striae, which merge with

the punctation anteriorly. Metapleurum distinct-

ly coriaceous, densely punctate. Pleural carina dis-

tinct. Areolet rather large, pointed, receiving the

2"d recurrent vein at the middle. Nervellus slight-

ly inclivous, weakly intercepted. Nervellar index

0.7. Tibial spur index 0.6. Hind femur 4.6x as long

as high. Hind tarsal claws short, with short pecti-

nate teeth basally and medially. Propodeum ex-

tending to 0.3 of the hind coxae, slightly and rath-

er narrowly depressed, coriaceous anteriorly, ir-

regularly wrinkled medially, with some distinct

longitudinal wrinkles posteriorly, costulae partly

present, area superomedia sometimes delimited

laterally, longitudinal carinae obliterated medial-

ly and sometimes also posteriorly. Propodeal spir-

acle rather small, 2.6x as long as wide. Spiracular

carina narrow. Spiracular index 0.7. 1% gastral

tergite 3.2x, 2" tergite 1.1x as long as wide. Peti-

ole with rather small glymmae, which are bor-

dered by longitudinal carinae dorsally, smooth in

front of the glymmae. Petiolar suture very fine.

1 gastral sternite for the greater part smooth, with

transverse striae posteriorly. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum separated from

the 3" gastral tergite anteriorly, folded in or fold-

ed back secondarily and pendent as a narrow

rectangular lobe, the crease or bulge marked with

black (inconspicuous). Epipleural index 0.3-0.4.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellow medially. Tegula

dark brown. Wings hyaline. Stigma medium brown

medially. Front leg from the femur onwards yel-

lowish red. Mid femur yellowish red on the apical

0.2. Mid tibia and tarsus yellowish red. Hind tibia

yellow medially, narrowly black basally, broadly

black apically. Hind tarsus dark brown. 2" gastral

tergite black or laterally tinged with red in front of

the black posterior edge. 3" tergite usually black

dorsally, dark red laterally (to a variable extend),

sometimes completely black. 4"to 7" tergites

sometimes narrowly marked with red ventrola-

terally. Gastral sternites black. Pilosity whitish.

d: Front trochanter yellowish red. Mid femur

yellow dorsally, dark brown ventrally. Gaster

sometimes completely black. In other characters

similar to the 9.

Material (1222, 338): Russia: Khabarovsk Terr.:

Khekhtsir near Khabarovsk (Hinz/ZSM, PET), Troit-

skoye; Primorye: Kamenushka (PET), Roshchino on

River Iman (Hinz/ZSM, PET); Kuril Isl: Sahmanbe on

Iturup (SAP).

Dusona memorator Hinz, 1994

Description. 2: Body length 9-10 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.8. Frons scarcely de-

pressed, finely coriaceous and finely rugose, with

a distinct median longitudinal carina, with fine

rather long radial wrinkles, which extend to 0.5-0.8

the distance to the median ocellus. Antennal cari-

na low and narrow. Face and dorsal 0.6 of the

clypeus coriaceous, densely rugose and rugose-

punctate. Clypeus slightly concave, coriaceous and

dispersely punctate ventrally, its apex slightly

convex medially. Malar space index 0.4. Genal

carina low and straight ventrally, joining the low

oral carina at an acute angle. Genal index 0.4.

Temple coriaceous and with fine rather dense

punctation medially. Flagellum with 45-46 seg-

ments, filiform, the apical 0.3 distinctly attenuate,

2"! segment 2.3x, shortest segment 1.1x, penulti-

mate segment 1.3x as long as wide. Pronotum

coriaceous and with very dense punctation later-

ally on the dorsal 0.4, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

dull, densely or very densely punctate. Scutellum

with lateral carinae on the anterior 0.3. Prescutel-

lar groove smooth medially, with fine longitudi-

nal striae laterally. Prepectal carina low, ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

distinctly coriaceous, distinctly and very densely

punctate. Speculum punctate posteriorly, coria-

ceous and granulate-strigose anteriorly, rather

dull. Depression in front of the speculum for the

greater part with rather strong short longitudinal

wrinkles, which merge with the punctation ante-

riorly, the ventral 0.2-0.3 finely granulate-strigose.

Metapleurum finely and densely rugose. Pleural

carina distinct. Areolet small, stalked, receiving

the 2” recurrent vein distinctly distad of the mid-

dle. Nervellus inclivous, distinctly intercepted.

Nervellar index 0.8. Tibial spur index 0.6. Hind

femur 6.1x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to 0.3 of the hind

coxae, not deeply and rather narrowly depressed,

with transverse and irregular wrinkles in the lon-

gitudinal furrow, costulae partly present, longitu-

dinal carinae distinct and complete. Propodeal

spiracle small, 2.4x as long as wide. Spiracular

carina narrow, sometimes repleced by several

wrinkles. Spiracular index 1.0. 1* gastral tergite

4.4x, 2"! tergite 2.3x as long as wide. Petiole with

distinct and rather elongate glymmae, with fine

longitudinal striae in front of the glymmae, with

some transverse wrinkles anteriorly. Petiolar su-

ture fine, partly obliterated. 1* gastral sternite

finely coriaceous. Thyridium elongate. Thyridial

index about 1.0. Epipleurum separated from the

3“ gastral tergite, the crease marked with black.

Epipleural index 0.7-0.8. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible tinged with brown medially. Wings

hyaline. Stigma dark brown. Front leg from the

femur onwards yellowish red, the femur marked

with black basally and ventrally. Mid femur

marked with yellowish red on the apical 0.2. Mid

tibia yellowish red. 2"“ gastral tergite red on the

posterior 0.1-0.2. 3” and 4" tergites red, 4" tergite

narrowly marked with black dorsally. 5" tergite

broadly marked with red ventrolaterally. Subgen-

ital plate reddish. Pilosity whitish.

d unknown.

Material (3??): Russia: Yevreyskiy A.O.: Birobid-

zhan (Hinz/ZSM); Primorye: Situkha (PET). Japan:

Honshu: Mt. Azumayama (SAP).

Dusona mercator (Fabricius, 1793)

Campoplex oxyacanthae Boie, 1855 (syn. nov.)

Campoplex mesoxanthus Förster, 1868 (syn. nov.)

Hinz (1972: 46) was not sure of this synonymy,

because the head, the prothorax and greater parts

of the legs are missing in the holotype of Ichneu-

mon mercator Fabricius, and because he did not

have material of related species with him, when

he studied the type in the Cobenhaven museum.

The junior author compared the holotype with

material of all species of the relevant species group

(Hinz 1977: 51) in coll. Hinz. Although head char-

acters cannot be studied, the position of the black

lateral stripes on the sides of the 2”! gastral tergite

(see below) is distinctive. The name Ichneumon

mercator is not a nomen oblitum under the Code

(article 23.9), because the conditions of articles

23.9.1.1 and 23.9.6 are not met (Yu & Horstmann

1997: 149; Horstmann 2001: 27). On the other hand,

the identification of Campoplex oxyacanthae Boie is

101

questionable, because its type is lost, and because

the species described here is not a parasitoid of

Allophyes oxyacanthae (Linnaeus, 1758) (Noctuidae)

as published by Boie (1855: 104), but was bred by

Hinz from Colotois pennaria (Linnaeus, 1761) (Ge-

ometridae) (22?, 18 from Hannover / Germany in

coll. Hinz/ZSM) and Compsoptera opacaria (Hüb-

ner, 1819) (Geometridae) (12, 15 from Barcelona /

Spain in coll. Hinz/ZSM). Either Boie was wrong

about the host association of his material, or Holm-

gren (1872: 60) misinterpreted the species described

by Boie. C. oxyacanthae Boie is interpreted here as

a senior synonym of C. mesoxanthus Förster (Holm-

gren, 1. c.), a taxon represented by a lectotype

(Hinz 1962: 4).

Description. 2: Body length 11-13 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.7. Frons scarcely depressed, coriaceous,

densely rugose-punctate, with a distinct median

longitudinal carina, with short radial wrinkles.

Antennal carina low and narrow. Face and dorsal

0.7 ofthe clypeus coriaceous, densely rugose-punc-

tate. Clypeus flat, coriaceous and rather densely

punctate ventrally, its apex almost truncate medi-

ally. Malar space index 0.3. Genal carina almost

straight ventrally, strongly raised, joining the

strongly raised oral carina at an acute angle, form-

ing a lopsided triangular pyramid, the tip of the

latter shifted ventrally and situated behind the

base of the mandible (Fig. 12). Gena deeply de-

pressed close to the junction, the depression 0.8x

as deep as the width of the 2" flagellar segment.

Genal index 0.4. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 53-55 segments, filiform, the appical

0.3 distinctly attenuate, 2”! segment 1.6x, shortest

segment 0.8x, penultimate segment 1.1x as long

as wide. Pronotum coriaceous and densely ru-

gose-punctate laterally on the dorsal 0.3, with lon-

gitudinal wrinkles ventrally. Mesoscutum and

scutellum finely coriaceous, very densely punc-

tate. Scutellum with lateral carinae on the anterior

0.4. Prescutellar groove smooth medially, with

longitudinal striae laterally. Ventral part of the

prepectal carina merging with the transverse car-

ina, both carinae distinctly raised, the former rath-

er narrowly and shallowly emarginate medially.

Pleural part of the prepectal carina low, the dorsal

0.2 replaced by several wrinkles. Mesopleurum

with rather dense or dense punctation on a very

finely coriaceous or smooth background, usually

with longitudinal wrinkles ventrally in the poste-

rior corner. Speculum very finely coriaceous, shin-

102

ing. Depression in front of the speculum with

distinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum densely ru-

gose, partly with longitudinal wrinkles. Pleural

carina strong. Areolet large, pointed or short-

stalked, receiving the 2" recurrent vein almost at

the middle. Nervellus slightly inclivous, distinct-

ly intercepted. Nervellar index 0.7. Tibial spur

index 0.7. Hind femur 5.5-6.3x as long as high.

Hind tarsal claws short, with long pectinate teeth

basally and medially. Propodeum extending to

0.4 of the hind coxae, broadly and deeply de-

pressed, sulcus almost completely with transverse

wrinkles, costulae partly present, longitudinal car-

inae strong and usually almost complete. Propo-

deal spiracle rather large, 3.1x as long as wide.

Spiracular carina narrow or partly widened. Spirac-

ular index 0.4. 1* gastral tergite 4.7x, 2”! tergite

2.0x as long as wide. Petiole without glymmae,

with narrow lateral areas, smooth in front ofthese

areas. Petiolar suture partly obliterated. 1“ gastral

sternite for the greater part smooth, with fine

transverse striae posteriorly. Thyridium circular.

Thyridial index about 2.0. Epipleurum separated

from the 2" gastral tergite (as in all species of

Dusona studied here), the crease slightly marked

with black, with a black lateral stripe almost par-

allel with the crease in a short distance (black

lateral stripes separate from the ventrolateral creas-

es of the 2" tergite lacking in related species).

Epipleurum not separated from the 3° gastral

tergite, the latter without black lateral stripes

(Fig. 43). Gaster strongly compressed posteriorly.

Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow-

ish red dorsally and medially. Tegula yellowish

brown. Wings hyaline. Stigma medium brown

medially. Front trochanter usually brown dorsal-

ly, yellow ventrally. Front femur yellowish red,

sometimes marked with brown basally. Front tib-

ia and tarsus yellow. Mid femur yellow on the

apical 0.2. Mid tibia yellow. Mid tarsus brown, the

basitarsus yellowish brown. Hind tibia yellow.

Hind tarsus blackish. 2” gastral tergite dorsally

yellow on the posterior 0.2, laterally yellow on the

posterior 0.5, sometimes completely yellow later-

ally. 34 tergite yellow. 4" tergite marked with

yellow anteriorly (to a variable extend). Pilosity

whitish.

d: Maxillary palpus yellowish red. Mandible

almost completely yellow. Tegula yellow. Front

and mid legs from the tip of the coxa onwards

yellow, the femora yellowish red. In other charac-

ters similar to the ®.

Material (2?2): Russia: Irkutsk Prov.: Angarsk (MOS),

Irkutsk (PET).

Dusona meridionator Aubert, 1960 (stat. nov.)

Dusona kyushui Hinz, 1994 (syn. nov.)

This is a variable and poorly defined species. Au-

bert (1960: 63) described D. meridionator as a sub-

species of D. tenuis. It differs from the latter in:

Genal carina not obliterated ventrally. Ventral part

of the prepectal carina not strongly raised. Propo-

deum with short transverse striae in the narrow

longitudinal furrow only. In D. meridionator, the

colour pattern of the hind tibia (used as distin-

guishing character by Aubert) is variable. The

types of D. meridionator (Aubert/LAU) and the

holotype of D. kyushui from Hiko-san/Kyushu/

Japan (SAP) have recently been studied by the

junior author.

Description. ?: Body length 8-9 mm. Temples

distinctly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.6-0.9. Frons not depressed, coriaceous, densely

rugose-punctate and rugose, with a fine median

longitudinal carina, with short radial wrinkles,

which extend to 0.2-0.4 the distance to the median

ocellus. Antennal sockets rather close to each oth-

er. Antennal carina low, narrow or widened to a

smooth crescent-shaped plate dorsally (the shape

of the latter variable). Face and dorsal half of the

clypeus coriaceous, densely rugose-punctate and

rugose. Clypeus flat, finely coriaceous and rather

densely or densely punctate ventrally, its apex

slightly convex medially. Malar space index 0.3.

Genal carina low ventrally, slightly bent sidewards,

joining the slightly raised oral carina almost at a

right angle. Genal index 0.25-0.4. Temple finely

coriaceous and with fine rather dense punctation

medially. Flagellum with 43-46 segments, filiform,

the apical 0.3 attenuate, 2" segment 1.7x, shortest

segment 1.0x, penultimate segment 1.2x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum coriaceous, very densely

punctate. Scutellum densely rugose-punctate, with

lateral carinae on the anterior 0.7. Prescutellar

groove smooth medially, with distinct short lon-

gitudinal striae laterally. Ventral part of the pre-

pectal carina either low, merging with the pleural

part, the dorsal 0.2 of the latter obliterated and

covered with wrinkles, or slightly raised, merging

with the also slightly raised transverse carina, the

pleural part lower, its dorsal 0.4 obliterated. Trans-

verse carina either more or less obliterated or

more or less raised (these characters variable).

Mesopleurum distinctly coriaceous, distinctly and

densely punctate. Speculum punctate posteriorly,

granulate-strigose and with fine oblique striae

anteriorly. Depression in front of the speculum

with fine longitudinal striae, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, very densely punctate, partly rugose-punc-

tate. Pleural carina distinct. Areolet rather small,

short-stalked, receiving the 2" recurrent vein dis-

tad of the middle. Nervellus distinctly reclivous,

weakly intercepted. Nervellar index 0.6-0.7. Tibial

spur index 0.6. Hind femur 5.0x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to 0.6 of the hind

coxae, deeply and rather narrowly depressed, with

short transverse striae in the longitudinal furrow,

irregularly wrinkled beside the furrow, carinae

almost completely obliterated (Fig. 34). Propodeal

spiracle rather large, 3.0x as long as wide. Spirac-

ular carina widened. Spiracular index 0.3-0.5.

1" gastral tergite 5.0x, 2" tergite 2.1x as long as

wide. Petiole without glymmae, narrow lateral

areas weakly indicated (by a coriaceous sculp-

ture), petiole smooth in front of these areas. Peti-

olar suture for the greater part obliterated. 1* gas-

tral sternite smooth. Thyridium elongate. Thyrid-

ial index 1.0-1.5. Epipleurum not separated from

the 3" gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.5.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown or dark brown apically.

Mandible black or tinged with reddish brown

medially. Tegula yellowish brown to black. Wings

hyaline. Stigma medium brown or dark brown

medially. Front femur yellowish red. Front tibia

and tarsus yellow. Apical 0.2 of the mid femur

yellowish red. Mid tibia yellow. Mid tarsus dark

brown. Hind tibia black, sometimes whitish sub-

basally on the dorsal side (to a variable extend).

2"d gastral tergite red on the posterior 0.3-0.5.

3! and 4" tergites red. 5" tergite sometimes marked

with red anteriorly. Pilosity whitish.

d: Maxillary palpus yellowish brown. Man-

dible broadly yellow medially. Tegula yellow.

Front leg from the tip of the coxa onwards yellow

or yellowish red. Mid femur sometimes complete-

ly yellowish red. Hind tibia distinctly whitish sub-

basally on the dorsal side. 4" gastral tergite some-

times marked with black posteriorly. In other char-

acters similar to the ®.

103

Material (622, 13): Russia: Chita Prov.: Urulyunguy

near Borzya (Hinz/ZSM, PET). Kazakhstan: Aksu-

Dzhabagly ESE of Chimkent (PET). Kirgiziya: Ala-

Archa River in the Kirgizskiy Mts., 1650 m (LIN), Ara-

Byik River in the Talasskiy Alatau Mts., 2050-3250 m

(Hinz/ZSM). Japan: Kyushu: Hiko-san (SAP). The

3?? from Urulyunguy are paratypes of D. micrator.

Variety: Temples scarcely constricted behind the

eyes, 0.8-0.9x as long as the width of an eye,

tangential lines on eyes and temples intersect on

the apex of the gaster. Flagellum with 47 seg-

ments. Prepectal carina low, ventral part merging

with the pleural part, the dorsal 0.1 of the latter

replaced by several wrinkles. Transverse carina

present anteriorly, obliterated posteriorly. Mate-

rial: 258 Mongoliya, Arhangay Prov., Bugat

(18 Hinz/ZSM, 18 PET). The status of these dd is

unknown.

Dusona meritor Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Primorye, Andre-

yevka, near Khasan, 27.8.1983 (PET). - Paratypes:

1? Russia, Kuril Isl., Kunashir, Tretyakovo, 5.8.

1973 (Hinz/ZSM); 1? Japan, Honshu, Kanagawa,

Chigasaki, 18.5. 1933 (AEI).

Description. ?: Body length 8-9 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate,

with a fine median longitudinal carina, with short

radial wrinkles, which extend to 0.3 the distance

to the median ocellus. Antennal carina low and

narrow. Face and dorsal 0.7 of the clypeus densely

rugose-punctate. Clypeus flat, finely coriaceous

and rather densely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina low and almost straight ventrally, joining

the distinctly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 42-43 segments, filiform, the apical 0.8 slight-

ly attenuate, 2” segment 2.2x, shortest segment

0.9x, penultimate segment 1.0x as long as wide.

Pronotum densely rugose-punctate laterally on

the dorsal 0.2, with longitudinal wrinkles ventral-

lv. Mesoscutum and scutellum coriaceous, very

densely punctate. Scutellum with lateral carinae

on the anterior 0.7. Prescutellar groove smooth.

Prepectal carina low, the ventral part merging

with the pleural part and with the transverse car-

104

ina. Ventral part not emarginate medially, pleural

part distinct ventrally and medially, its dorsal 0.4

obliterated. Transverse carina distinct, its rim bent

upwards. Pleural part of the prepectus slightly

depressed above the transverse carina. Mesopleu-

rum coriaceous, densely punctate. Speculum co-

riaceous, dull. Depression in front of the specu-

lum with distinct longitudinal striae, which merge

with the punctation anteriorly. Metapleurum

densely rugose. Pleural carina distinct. Areolet

large, short-stalked, receiving the 2" recurrent

vein distinctly basad of the middle. Nervellus re-

clivous, weakly intercepted. Nervellar index 0.6.

Tibial spur index 0.6. Hind femur 5.1x as long as

high. Hind tarsal claws rather short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.4 of the hind coxae, scarcely de-

pressed, with short transverse wrinkles in the shal-

low and narrow longitudinal furrow, irregularly

wrinkled beside the furrow, sometimes with lon-

gitudinal wrinkles posteriorly, costulae fine but

complete, longitudinal carinae present only pos-

teriorly. Propodeal spiracle large, 3.7x as long as

wide. Spiracular carina more or less indistinct.

Spiracular index 0.2-0.4. 1” gastral tergite 4.6x, 2"<

tergite 2.5x as long as wide. Petiole with very

small and shallow glymmae at the anterior edge

of narrow lateral areas, with very fine sculpture or

smooth in front of the glymmae. Petiolar suture

for the greater part obliterated. 1* gastral sternite

für the greater part smooth, with fine transverse

striae posteriorly. Thyridium elongate. Thyridial

index about 1.5. Epipleurum not separated from

the 3"4 gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.0.

Colour: Black, Maxillary palpus yellow. Man-

dible almost completely yellow. Tegula dark

brown. Wings hyaline. Stigma dark brown medi-

allyv. Front leg from the tip of the coxa onwards

yellowish, trochanter marked with brown, femur

medium brown ventrally. Mid leg from the apical

0.1 of the femur onwards yellowish. Hind tibia

blackish, marked with dark brown medially on

the dorsal side. In one paratype, the front coxa

and trochanter are black, and the mid tibia and

tarsus are dark brown. 2" gastral tergite marked

with red laterally. 3” and 4" tergites red, the 3°“

tergite dorsally marked with black anteriorly. Hind

tergites marked with red posteriorly. Hind ster-

nites red. Pilosity whitish.

d unknown.

Dusona micrator Hinz, 1979

The type material of this species is not homoge-

nous. The holotype (?) from Kamikochi near

Norikura-dake/Honshu/Japan (AEI) and para-

types (5?2) from the same locality, from Sappo-

ro/Hokkaido/Japan (both AEI), Nopporo/Hok-

kaido/Japan, Nikishiro on Kunashir/Kuril Isl./

Russia (both SAP) and Yuzhno-Kurilsk on Kuna-

shir/Kuril Isl./ Russia (PET) belong to D. micrator.

322 from Urulyunguy/Chita Prov./Russia (Hinz /

ZSM, PET) belong to D. meridionator, 12 from Ser-

novodsk on Kunashir/Kuril Isl./ Russia (PET) be-

longs to D. schikotani, and 1? from Sapporo/

Hokkaido/Japan (SAP) is placed as a variety to

D. obtutor.

Description. 2: Body length 7-8 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.7. Frons scarcely depressed, coriaceous, densely

rugose-punctate, with a fine and partly obliterat-

ed median longitudinal carina, with short and

indistinct radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.8 of the clypcus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low and straight ventrally, some-

times slightly bent forwards, joining the low oral

carina at an acute angle. Genal index 0.2-0.3. Tem-

ple finely coriaceous and with very fine disperse

punctation medially. Flagellum with 40-44 seg-

ments, filiform, the apical 0.3 slightly attenuate,

2rd segment 2.3x, shortest segment 1.0x, penulti-

mate segment 1.1-1.2x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, dull, very dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.7. Prescutellar groove smooth medially,

with fine longitudinal striae laterally. Prepectal

carina low, ventral part merging with the pleural

part, the dorsal 0.3 of the latter obliterated. Trans-

verse carina more or less obliterated (Fig. 17).

Mesopleurum distinctly coriaceous, densely or

very densely punctate. Speculum punctate poste-

riorly, granulate-strigose and shining anteriorly.

Depression in front of the speculum granulate-

strigose and with fine short longitudinal wrin-

kles, which merge with the punctation anteriorly.

Metapleurum coriaceous, dull, finely and densely

rugose. Pleural carina fine but complete. Areolet

rather large, stalked, receiving the 2" recurrent

vein distad of the middle. Nervellus almost verti-

cal, distinctly intercepted. Nervellar index 0.6-0.7.

Tibial spur index 0.5. Hind femur 5.6x as long as

high. Hind tarsal claws short, with few short pec-

tinate teeth basally. Propodeum extending to 0.4

ofthe hind coxae, scarcely depressed, for the great-

er part irregularly wrinkled, sometimes with some

short transverse wrinkles in the midline, with some

longitudinal wrinkles posteriorly, carinae almost

completely obliterated. Propodeal spiracle rather

small, 4.5x as long as wide. Spiracular carina nar-

row, sometimes indistinct. Spiracular index about

0.5. 1% gastral tergite 4.4x, 2" tergite 2.1x as long

as wide. Petiole smooth laterally, without glym-

mae or narrow lateral areas. Petiolar suture fine,

partly obliterated. 1“ gastral sternite for the great-

er part smooth or finely coriaceous, with fine trans-

verse striae posteriorly. Thyridium elongate. Thy-

ridial index about 1.5. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown

basally, medium brown apically. Mandible nar-

rowly tinged with reddish brown subapically.

Wings hyaline. Stigma light brown or medium

brown medially. Front femur yellowish red on the

apical half. Front tibia and tarsus yellowish red.

Mid tibia and tarsus dark brown. Hind tibia black-

ish, sometimes tinged with brown subbasally. 2‘

gastral tergite red on the posterior 0.1-0.2. 3° and

4 tergites red. Pilosity whitish.

d unknown.

Material (622): Russia: Kuril Isl.: Nikishiro on Kuna-

shir (SAP), Yuzhno-Kurilsk on Kunashir (PET). Japan:

Hokkaido: Nopporo (SAP), Sapporo; Honshu: Ka-

mikochi near Norikura-dake (AEI).

Dusona minor (Provancher, 1879)

The population from central Asia described here

differs from material of this rather variable spe-

cies described from North America by its dark

color (legs, gaster). It might represent a separate

subspecies.

Description. 2: Body length 6-7 mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 0.9. Frons scarcely depressed, coriaceous, fine-

ly and densely rugose-punctate, with a fine medi-

an longitudinal carina, with short radial wrinkles.

Antennal carina low and narrow, the rim slightly

105

bent upwards. Face and dorsal 0.6 of the clypeus

finely and densely rugose. Clypeus flat, coria-

ceous and with fine dense punctation ventrally,

its apex truncate medially. Malar space index 0.5.

Genal carina low ventrally, slightly bent sidewards

and forwards, joining the oral carina at the base of

the mandible at a right angle. Temple finely coria-

ceous and with fine disperse punctation medially.

Flagellum with 32-36 segments, slightly fusiform,

2rd segment 2.0x, shortest segment 1.0x, penulti-

mate segment 1.3x as long as wide. Pronotum

finely and densely rugose-punctate laterally on

the dorsal 0.2, with longitudinal wrinkles ventral-

ly. Hind corner of pronotum produced to a flange,

which projects about as far as the tegula and which

isrounded dorsally and excavated ventrally. Meso-

scutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior 0.8. Prescutellar groove smooth.

Prepectal carina low, ventral part merging with

the pleural part or with the transverse carina, the

latter somewhat raised. Pleural part of the prepec-

tal carina low, its dorsal 0.2 indistinct or obliterat-

ed. Mesopleurum coriaceous, densely punctate.

Speculum coriaceous and finely granulate-strigose,

a small ventral part almost smooth. Depression in

front of the speculum with distinct longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum finely and densely rugose or

rugose-punctate. Pleural carina distinct. Areolet

rather small, long-stalked, receiving the 2"“ recur-

rent vein distad of the middle. Nervellus vertical,

indistinctly intercepted. Nervellar index 0.8. Tibi-

al spur index 0.6. Hind femur 4.7x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to the middle of

the hind coxae, broadly and distinctly but not

deeply depressed, sulcus more irregularly wrin-

kled anteriorly, transversely wrinkled posterior-

ly, costulae complete, longitudinal carinae almost

completely obliterated. Propodeal spiracle small,

2.3x as long as wide. Spiracular carina narrow.

Spiracular index 0.5. 1" gastral tergite 4.0x, 2"

tergite 2.0x as long as wide. Petiole with rather

small but distinct glymmae at the anterior edge of

indistinct narrow lateral areas, smooth in front of

the glymmae. Petiolar suture partly obliterated.

1" gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.5. Epipleurum not separated

from the 3"! gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.7.

Colour: Black. Wings hyaline. Stigma light

brown medially. Front leg from the apical 0.6 of

the femur onwards yellowish brown. 2"! gastral

106

tergite black or red on the posterior 0.1. 3’! and 4"

tergites red, the 4" sometimes marked with black

posteriorly. Pilosity whitish.

d: Projection at the hind corner of pronotum

slightly longer than the tegula. Mandible marked

with yellowish brown medially. Tegula yellowish

brown to black. Front leg from the tip of the coxa

onwards yellowish red. Mid tibia and tarsus yel-

lowish brown to brown. In other characters simi-

lar to the 9.

Material (4??, 233): Kazakhstan: 20 km N of Kegen

(LIN). Kirgiziya: Kashkasu River S of Frunze (= Bish-

kek), 1700 m (PET, Hinz/ZSM), Lake Sary Chelek in

Dzhangi-Dzhol Distr., 1850-1900 m (LIN).

Variety: Projection at the hind corner of prono-

tum slightly shorter than the tegula. Sulcus of

propodeum with rather short transverse wrinkles

and with one or two short longitudinal wrinkles

posteriorly. Mid tibia yellowish. Hind tibia dark

yellowish brown medially, narrowly black basal-

ly, broadly black apically. 3° gastral tergite some-

times marked with black dorsally. 4" tergite

marked with black dorsally or completely black.

Material: 233 Russia, Khabarovsk Terr., Khekh-

tsir, near Khabarovsk (18 PET, 19 Hinz/ZSM);

13 Russia, Yevreyskaya A.O., Londoko (PET).

These dd are intermediary between D. angustifrons

and D. minor in the shape of the hind corner of

pronotum and intermediary between populations

of D. minor from central Asia and North America

in the colour of the legs. Their status is unknown.

Dusona minutor Horstmann, spec. nov.

Holotype (2): Russia, Primorye, Spassk Dalniy,

11.9. 1985 (Hinz / ZSM).

Description. 2: Body length 6 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons not depressed, coriaceous, densely ru-

gose-punctate, with a fine median longitudinal

carina, without radial wrinkles. Antennal sockets

rather close to each other. Antennal carina low,

dorsally widened to a smooth crescent-shaped

plate. Face and dorsal 0.4 of the clypeus coria-

ceous, densely rugose-punctate. Clypeus flat, co-

riaceous and dispersely punctate ventrally, its apex

truncate medially. Malar space index 0.3. Genal

carina low and almost straight, joining the low

oral carina at an acute angle. Genal index about

0.1. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with 32

segments, slightly fusiform, 2"! segment 2.3x,

shortest segment 0.9x, penultimate segment 1.1x

as long as wide. Pronotum with longitudinal wrin-

kles, partly also with punctation. Mesoscutum and

scutellum coriaceous, densely punctate. Scutel-

lum with lateral carinae on the anterior 0.4. Pre-

scutellar groove destroyed. Prepectal carina low,

ventral part merging with the transverse carina,

the latter slightly raised, the pleural part present

ventrally and medially, its dorsal 0.3 obliterated.

Mesopleurum finely coriaceous, almost smooth,

rather densely punctate. Speculum coriaceous and

granulate-strigose, shining. Depression in front of

the speculum with fine and long longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum densely rugose and rugose-punc-

tate, the wrinkles smooth. Pleural carina fine. Areo-

let rather large, pointed, receiving the 2" recur-

rent vein distad of the middle. Nervellus inclivous,

distinctly intercepted. Nervellar index 0.8. Tibial

spur index 0.7. Hind femur 5.2x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to 0.6 of the hind

coxae, not depressed, irregularly wrinkled, almost

without carinae, with a median longitudinal keel

posteriorly. Propodeal spiracle small, 2.5x as long

as wide. Spiracular carina narrow. Spiracular in-

dex 0.4. 1" gastral tergite 5.5x, 2"! tergite 2.1x as

long as wide. Petiole without distinct glymmae,

with distinct narrow lateral areas, smooth in front

of these areas. Petiolar suture partly obliterated.

1 gastral sternite smooth. Thyridium elongate.

Thyridial index about 2.0. Epipleurum not sepa-

rated from the 3" gastral tergite, the latter without

black lateral stripes. Gaster slightly compressed

posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellowish

brown. Wings hyaline. Stigma medium brown

medialiy. Front femur yellowish red. Front tibia

and tarsus yellow. Mid femur yellowish red on

the apical half. Mid tibia and tarsus yellow. Hind

tibia dark yellowish brown medially and apically,

narrowly dark brown basally. Hind tarsus dark

brown dorsally, yellowish brown ventrally. 2”

gastral tergite red on the posterior 0.2, the red

marks more extended laterally. 3 and 4'" tergites

red. 5" tergite laterally marked with red anterior-

ly. Pilosity whitish.

d unknown.

Dusona momoii Hinz & Horstmann, spec. nov.

Types. Holotype (?): Japan, Hokkaido, Sapporo,

20.6.1958 (SAP). — Paratypes: 1? Japan, Hokkaido,

Sapporo, Maruyama, 8.6.1958 (Hinz/ZSM); 1?

Japan, Honshu, Sasayama, 10.5.1954 (SAP); 14

Japan, Hokkaido, Sapporo, Maruyama, 4.6.1957

(SAP); 15 Russia, Primorye, Sikhote-Alin, 10.6.

1929 (MOS); 13 Russia, Primorye, Kamenushka,

SE of Ussuriysk, 7.6.1979 (PET); 18 Russia, Primo-

rye, Ussuriysk, 19.6.1983 (Hinz / ZSM).

Description. 2: Body length 11-13 mm. Temples

strongly constricted constricted behind the eyes,

0.8x as long as the width of an eye, tangential lines

on eyes and temples intersect on the posterior half

ofthe mesoscutum. Ocellar index 1.0. Frons slightly

depressed, coriaceous, densely rugose-punctate,

with a fine median longitudinal carina, with short

radial wrinkles, which extend to 0.3 the distance

to the median ocellus. Antennal carina low and

narrow. Face and clypeus densely rugose-punc-

tate. Clypeus flat, its apex truncate medially. Ma-

lar space index 0.3. Genal carina low and straight

ventrally, joining the scarcely raised oral carina at

an acute angle. Genal index 0.3. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 47-50 segments, filiform,

the apical 0.2 distinctly attenuate, 2”! segment

1.5x, shortest segment 0.9x, penultimate segment

1.1x as long as wide. Pronotum with very dense

punctation laterally on the dorsal 0.6, with longi-

tudinal wrinkles ventrally. Mesoscutum and

scutellum densely punctate on a finely coriaceous

or smooth background. Scutellum with lateral car-

inae on the anterior 0.6. Prescutellar groove with

fine short longitudinal striae or almost smooth.

Prepectal carina low, the ventral part merging

with the pleural part, the latter almost joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum densely punc-

tate on a finely coriaceous or partly smooth back-

ground. Speculum partly smooth. Depression in

front of the speculum with distinct longitudinal

wrinkles dorsally and medially, the latter merg-

ing with the punctation anteriorly, with very fine

striae or almost smooth ventrally. Metapleurum

densely rugose-punctate, the wrinkles smooth.

Pleural carina distinct. Areolet large, short-stalked,

receiving the 2" recurrent vein at or basad of the

middle. Nervellus slightly inclivous, distinctly in-

tercepted. Nervellar index 0.8. Tibial spur index

0.6. Hind femur 5.6x as long as high. Hind tarsal

claws short, with short pectinate teeth basally and

medially. Propodeum extending to 0.4 of the hind

coxae, scarcely and rather broadly depressed, for

107

the greater part irregularly wrinkled, with some

transverse wrinkles and with a short median lon-

gitudinal keel posteriorly, almost without cari-

nae. Propodeal spiracle large, 3.2x as long as wide.

Spiracular carina often indistinct or replaced by

several wrinkles. Spiracular index 0.3. 1* gastral

tergite 3.8x, 2" tergite 1.9x as long as wide. Peti-

ole with large glymmae, which are bordered by

longitudinal carinae dorsally and ventrally, with

fine longitudinal striae in front of the glymmae

and with some distinct transverse wrinkles anteri-

orly. Petiolar suture indicated or complete. 1 gas-

tral sternite for the greater part smooth, with fine

transverse striae posteriorly. Thyridium elongate.

Thyridial index about 1.0. Epipleurum separated

from the 3" gastral tergite, the crease marked with

black. Epipleural index 0.8. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Wings scarcely

darkened or hyaline. Stigma medium brown me-

dially. Front trochanter, tibia and tarsus yellow,

the trochanter marked with brown. Front femur

yellowish red. Mid leg from the apical 0.3 of the

femur onwards yellowish red. Hind tibia yellow-

ish brown, narrowly dark brown basally, scarcely

darkened apically. 2” gastral tergite dorsally black

or red on the posterior 0.1, broadiy marked with

red laterally. Gaster from the 3' tergite onwards

narrowly marked with black dorsally, broadly red

laterally. Gastral sternites red. Pilosity whitish.

d: Propodeum with the longitudinal carinae

partly indicated. Tegula yellowish brown to black.

Tips of the front and mid coxae and front and mid

trochanters yellow. Front and mid femora yellow-

ish red. Genital clasper red. In other characters

similar to the ®.

Variety: Costulae complete. Front coxa broadly

marked with yellow apically. Hind tibia broadly

marked with brown apically. Material: 12 Russia,

Primorye, Roshchino, on River Iman (PET). Prob-

ably this ? belongs to D. momoii.

Dusona montana (Roman, 1929)

Description. 2: Body length 10-12 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed,

coriaceous, partly with fine disperse punctation,

with a distinct median longitudinal carina, with

fine and short radial wrinkles. Antennal carina

low and narrow. Face and dorsal 0.2 of the clypeus

108

coriaceous, finely and densely rugose and rugose-

punctate. Clypeus almost flat, coriaceous and with

fine rather dense punctation ventrally, its apex

truncate or slightly convex medially. Malar space

index 0.3. Genal carina low and straight ventrally,

joining the scarcely raised oral carina at an acute

angle (Fig. 8 and 13). Genal index 0.4. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 43-45 segments,

filiform, the apical 0.2 attenuate, 2‘ segment 2.5x,

shortest segment 1.0x, penultimate segment 1.4x

as long as wide. Pronotum coriaceous and dense-

ly rugose-punctate laterally on the dorsal 0.2, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, densely or very dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.2. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter complete, joining the anterior

edge of the mesopleurum. Transverse carina com-

plete or almost complete (Fig. 16). Mesopleurum

finely coriaceous, densely punctate. Speculum fine-

ly coriaceous, shining. Depression in front of the

speculum with short and distinct longitudinal

wrinkles, which merge with the punctation ante-

riorly. Metapleurum finely and densely rugose-

punctate. Pleural carina complete. Areolet rather

large, stalked, receiving the 2" recurrent vein dis-

tad of the middle. Nervellus slightly inclivous,

distinctly intercepted. Nervellar index 0.6-0.7. Tib-

ial spur index 0.6. Hind femur 5.9x as long as

high. Hind tarsal claws rather long, with distinct

pectinate teetn almost over their total length. Pro-

podeum extending to the middle of the hind cox-

ae, scarcely or not at all depressed, for the greater

part with transverse wrinkles, sometimes partly

with irregular wrinkles, costulae partly present,

longitudinal carinae for the greater part obliterat-

ed or complete. Propodeal spiracle large, 4.0x as

long as wide. Spiracular carina narrow or wid-

ened. Spiracular index 0.2-0.4. 1% gastral tergite

4.6x, 2"! tergite 2.4x aslong as wide. Petiole smooth

laterally, without glymmae or with very small

and shallow furrows at the anterior edge of nar-

row lateral areas. Petiolar suture fine. 1 gastral

sternite smooth or partly coriaceous, sometimes

with fine transverse wrinkles posteriorly. Thyrid-

ium oval. Thyridial index about 2.0. Epipleurum

separated from the 3“ gastral tergite, the crease

marked with black. Epipleural index 0.7-0.8. Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.9

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Mandible

broadly yellow medially. Tegula brown. Wings

hyaline. Stigma dark brown medially. Front leg

from the tip of the coxa onwards yellowish red.

Mid femur yellowish red on the apical 0.3. Mid

tibia and tarsus yellowish red. Hind tibia dark

yellowish brown, narrowly black basally, not dark-

ened apically. 2" gastral tergite dorsally red on

the posterior 0.2-0.3, laterally red on the posterior

0.5. 3’ and 4 tergites red, sometimes marked

with black dorsally. 5'" to 7'" tergites sometimes

narrowly marked with red ventrolaterally. Gas-

tral sternites sometimes red. Pilosity whitish.

d similar to the 2.

Material (4?%): Russia: Kuril Isl.: Volcano Golovnina

on Kunashir (PET). Japan: Honshu: Mt. Minoo in Os-

aka-fu; Shikoku: Teragawa (SAP).

Dusona myrtilla (Desvignes, 1856)

Description. 2: Body length 11-13 mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the petiole. Ocellar index 1.0.

Frons slightly depressed, coriaceous, finely and

densely punctate, with a fine median longitudinal

carina, without radial wrinkles. Antennal carina

low and narrow. Face and dorsal 0.3 of the clypeus

finely coriaceous, finely and densely rugose-punc-

tate. Clypeus flat, finely and densely punctate

ventrally, with a narrow smooth area above the

apex, the latter truncate medially. Malar space

index 0.3. Genal carina low and straight ventrally,

joining the slightly raised oral carina at an acute

angle. Genal index 0.25-0.3. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 45-46 segments, filiform,

the apical 0.3 attenuate, 2” segment 1.4x, shortest

segment 0.8x, penultimate segment 1.2x as long

as wide. Pronotum coriaceous and finely and

densely rugose-punctate laterally on the dorsal

0.4, with longitudinal wrinkles ventrally. Meso-

scutum coriaceous, very densely punctate. Scutel-

lum coriaceous, rather densely punctate, with lat-

eral carinae on the anterior 0.4. Prescutellar groove

almost completely smooth. Prepectal carina low,

ventral part merging with the pleural part, the

latter complete, joining the anterior edge of

the mesopleurum. Transverse carina obliterated.

Mesopleurum distinctly and densely or rather

densely punctate on a finely coriaceous and rather

shining background. Speculum for the greater part

smooth. Depression in front of the speculum dor-

sally with fine longitudinal striae, which merge

with the punctation anteriorly, the ventral half

coriaceous and granulate-strigose. Metapleurum

coriaceous, densely punctate and rugose-punc-

tate. Pleural carina strong. Areolet large, pointed,

receiving the 2" recurrent vein at or slightly dis-

tad of the middle. Nervellus vertical, strongly in-

tercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 4.6x as long as high. Hind tarsal

claws short, with distinct pectinate teeth almost

over their total length. Propodeum extending to

0.4 of the hind coxae, broadly and not deeply

depressed, irregularly wrinkled, with a longitudi-

nal keel posteriorly, carinae complete or almost

complete, partly covered with wrinkles. Propode-

al spiracle rather large, 3.7% as long as wide. Spirac-

ular carina widened. Spiracular index 0.3. 1“ gas-

tral tergite 3.2x, 2"d tergite 2.1x as long as wide.

Petiole with distinct glymmae, which are bordered

by longitudinal carinae dorsally, smooth in front

of the glymmae. Petiolar suture fine. 1* gastral

sternite smooth or with very fine sculpture, with

transverse striae posteriorly. Thyridium oval. Thy-

ridial index about 2.0. Epipleurum separated from

the 3" gastral tergite, the crease marked with black.

Epipleural index 0.8. Gaster distinctly compressed

posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellow. Man-

dible almost completely yellow. Tegula yellow.

Wings hyaline. Stigma yellowish brown medially.

Front trochanter marked with yellow. Front fe-

mur yellowish red, narrowly marked with brown

basally. Front tibia and tarsus yellow. Mid femur

yellow on the apical 0.2. Mid tibia and tarsus

yellow. Hind tibia whitish yellow medially, nar-

rowly black basally, broadly black apically. Hind

tarsus yellowish brown. Pilosity whitish.

d: Propodeum with some short transverse

wrinkles anteriorly and medially. Front and mid

legs almost completely yellow, the coxae marked

with black basally. In other characters similar to

the ®.

Material (6??, 438): Russia: Irkutsk Prov.: Lake Bay-

kal; Primorye: Benevskoye, Kiyevka, Plastun (PET).

Japan: Shikoku: Ehime (SAP).

Dusona nanus Horstmann, spec. nov.

The species is similar to D. parva, it differs from

the latter in: Propodeum coarsely wrinkled, its

longitudinal carinae indicated. Glymmae lacking,

but narrow lateral areas present. Gaster less ex-

tensively marked with red.

Holotype (d): Russia, Khabarovsk Terr., Vysoko-

gornyy, 13.7.1983 (PET),

109

Description. d: Body length 5mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous, fine-

Iy and densely punctate, with fine radial wrinkles,

which extend to half the distance to the median

ocellus, the median longitudinal carina indistinct.

Antennal carina low and narrow, its rim slightly

bent upwards. Face and dorsal 0.8 of the clypeus

coriaceous, finely and densely rugose-punctate

and rugose. Clypeus flat, coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.5. Genal carina low

and straight ventrally, joining the distinctly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with very fine dis-

perse punctation medially. Flagellum with 34 seg-

ments, filiform, almost not attenuate apically,

2rd segment 1.3x, shortest segment 0.9x, penulti-

mate segment 1.0x as long as wide. Pronotum

coriaceous and densely rugose-punctate laterally

on the dorsal 0.3, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.6. Prescutellar groove for the

greater part smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter in-

distinct, its dorsal 0.3 obliterated. Transverse car-

ina indistinct. Mesopleurum coriaceous, dull, fine-

ly and rather densely punctate. Speculum finely

coriaceous. Depression in front of the speculum

with fine longitudinal wrinkles, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, finely and densely punctate. Pleural carina

complete. Areolet rather large, stalked, receiving

the 2" recurrent vein distad of the middle. Nervel-

lus distinctly inclivous, distincly intercepted. Ner-

vellar index 0.8. Tibial spur index 0.7. Hind femur

4.8x as long as high. Hind tarsal claws short, with

short pectinate teeth basally. Propodeum extend-

ing to 0.3 of the hind coxae, broadly but not deep-

ly depressed, rather coarsely and irregularly wrin-

kled, costulae partly present, longitudinal carinae

indicated medially, but covered with wrinkles.

Propodeal spiracle small, 2.3x as long as wide.

Spiracular carina narrow. Spiracular index 0.7.

1% gastral tergite 3.1x, 2"4 tergite 1.6x as long as

wide. Petiole without glymmae, with narrow lat-

eral areas, which are bordered by carinae dorsally

and ventrally, smooth in front of these areas. Pet-

iolar suture fine. 1* gastral sternite for the greater

part smooth, with fine transverse striae posterior-

ly. Thyridium circular. Thyridial index about 1.0.

Epipleurum not separated from the 3"! gastral

tergite, the latter with rather indistinct black later-

110

al stripes well above the ventrolateral edge. Gaster

distinctly compressed posteriorly.

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Tegula dark

yellowish brown. Wings hyaline. Stigma yellow

medially. Front leg from the tip of the coxa on-

wards yellow, the coxa marked with brown. Mid

leg from the apical 0.2 of the femur onwards yel-

low. Hind tibia broadly yellow medially, narrow-

ly marked with dark brown basally, rather broad-

ly marked with dark brown apically. Hind tarsus

dark brown. 2"“ gastral tergite dorsally red on the

posterior 0.1, laterally red on the posterior 0.3. 3"

tergite narrowly black dorsally, broadly red later-

ally. 4" tergite laterally with broad red markings

anteriorly. Pilosity whitish.

? unknown.

Dusona nebulosa Horstmann, spec. nov.

Types. Holotype (5): Russia, Primorye, Spassk

Dalniy, 26.8.1981 (PET). — Paratype: 15 Russia,

Caucasus, Karachayevo-Cherkesskaya A.O., Ark-

hyz, 2.7.1976 (Hinz / ZSM).

Description. d: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.6. Frons slightly depressed, coriaceous, with fine

rather dense punctation, with a fine median longi-

tudinal carina, with short radial wrinkles. Anten-

nal carina low and narrow. Face and dorsal half of

the clypeus coriaceous, densely rugose-punctate.

Clypeus flat, finely coriaceous and rather densely

punctate ventrally, its apex truncate medially.

Malar space index 0.3. Genal carina low and

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.6. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 46-47 segments,

filiform, the apical 0.2 attenuate, 2” segment 1.7x,

shortest segment 1.0x, penultimate segment 1.4x

as long as wide. Pronotum coriaceous and dense-

ly rugose-punctate laterally on the dorsal 0.3, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, densely punctate. Scutel-

lum with lateral carinae on the anterior 0.7. Pres-

cutellar groove with few fine longitudinal striae.

Prepectal carina low, ventral part merging with

the pleural part, the dorsal 0.4 of the latter obliter-

ated. Transverse carina present only anteriorly.

Mesopleurum coriaceous, distinctly and densely

punctate. Speculum finely granulate-strigose, shin-

ing. Depression in front of the speculum granu-

late-strigose, partly with very fine longitudinal

striae, which merge with a coriaceous area with-

out punctation anteriorly. Metapleurum coria-

ceous, finely and densely punctate. Pleural carina

complete. Areolet rather small, long-stalked, re-

ceiving the 2" recurrent vein basad of the middle.

Nervellus vertical, weakly intercepted, nervellar

index 0.7 (in the holotype), or inclivous, distinctly

intercepted, nervellar index 0.8 (in the paratype).

Tibial spur index 0.6. Hind femur 4.7x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.4 of the

hind coxae, broadly but not deeply depressed, the

sulcus coriaceous anteriorly, transversely wrin-

kled posteriorly, fine costulae present, longitudi-

nal carinae indicated medially. Propodeal spiracle

rather large, 3.0x as long as wide. Spiracular cari-

na narrow. Spiracular index 0.3. 1“ gastral tergite

4.8x, 2"! tergite 2.2x as long as wide. Petiole with-

out glymmae or with very small and shallow glym-

mae at the anterior edge of narrow lateral areas,

smooth in front of these areas. Petiolar suture fine.

1* gastral sternite für the greater part with fine

irregular wrinkles, with fine transverse striae pos-

teriorly. Thyridium oval. Thyridial index about

1.0. Epipleurum not separated from the 3" gastral

tergite, the latter with black lateral stripes well

above the ventrolateral edge. Gaster distinctly

compressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula yellow to

yellowish brown. Wings hyaline. Stigma dark

brown medially. Front and mid legs from the tip

of the coxa onwards yellow, the femora yellowish

red. Hind tibia broadly yellowish brown medial-

ly, narrowly black basally, rather narrowly dark

brown apically. 2” gastral tergite red on the pos-

terior 0.1. 3’! and 4" tergites red, sometimes tinged

with black dorsally.

? unknown.

Dusona nervellator Horstmann, spec. nov.

Holotype (JS): Russia, Primorye, Yakovlevka, 28.

6.1927 (PET).

Description. d: Body length 12 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons slightly depressed, coriaceous,

densely rugose-punctate, with a fine and partly

obliterated median longitudinal carina, with short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal half of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, coriaceous

and with rather dense punctation ventrally, its

apex truncate medially. Malar space index 0.5.

Genal carina low and straight ventrally, joining

the scarcely raised oral carina at an acute angle.

Genal index 0.2. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 52 segments, filiform, the apical 0.3

attenuate, 2”! segment 2.0x, shortest segment 1.0x,

penultimate segment 1.4x as long as wide. Prono-

tum coriaceous and densely rugose-punctate lat-

erally on the dorsal half, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, dull, with very dense punctation. Scutellum

with lateral carinae on the anterior 0.7. Prescutel-

lar groove smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter dis-

tinct and complete, joining the anterior edge of

the mesopleurum. Transverse carina low, com-

plete. Mesopleurum distinctly coriaceous, distinct-

ly and densely punctate. Speculum punctate pos-

teriorly, granulate-strigose anteriorly. Depression

in front of the speculum granulate-strigose, with

few longitudinal wrinkles dorsally, the latter merg-

ing with a coriaceous area without punctation

anteriorly. Metapleurum distinctly coriaceous,

rather densely or densely punctate. Pleural carina

distinct. Areolet large, short-stalked, receiving the

2"d recurrent vein slightly basad of the middle.

Nervellus reclivous, intercepted. Nervellar index

0.4. Tibial spur index 0.5. Hind femur 4.5x as long

as high. Hind tarsal claws rather short, with dis-

tinct pectinate teeth basally. Propodeum extend-

ing to 0.3 of the hind coxae, not depressed, for the

greater part finely coriaceous, with fine disperse

punctation and with very fine irregular wrinkles,

shining, with few short transverse wrinkles poste-

riorly, almost without carinae. Propodeal spiracle

large, 5.0x as long as wide. Spiracular carina nar-

row. Spiracular index 0.3. 1* gastral tergite 3.0x,

2"! tergite 1.7x as long as wide. Petiole with rather

large glymmae, smooth in front of the glymmae.

Petiolar suture almost completely obliterated.

1* gastral sternite smooth. Thyridium elongate.

Thyridial index about 1.0. Epipleurum not sepa-

rated from the 3" gastral tergite, the latter with

fine black lateral stripes well above the ventrola-

teral edge. Gaster distinctly compressed posteri-

orly.

Colour: Black. Maxillary palpus dark brown

basally, yellowish medially and apically. Mandi-

ble broadly yellow medially. Tegula yellow. Wings

hyaline. Stigma yellow medially. Front leg from

the tip of the coxa onwards yellow, the femur

yellowish red. Mid femur yellowish red on the

apical 0.3. Mid tibia yellow. Mid tarsus yellowish

brown. Hind tibia rather narrowly yellow medial-

ly, rather broadly black basally and on the apical

0.4. 2"d gastral tergite dorsally red on the posterior

0.2, laterally red on the posterior 0.3. 3" and 4"

tergites red, the 4" marked with black on its pos-

terior 0.3. Pilosity whitish.

? unknown.

Dusona nidulator (Fabricius, 1804)

Description. 2: Body length 12-13 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.1. Frons slightly depressed, coriaceous, densely

rugose-punctate, with a fine and partly obliterat-

ed median longitudinal carina, with short and

indistinct radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.6 of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex slightly convex medially. Malar space

index 0.5. Genal carina low and almost straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.6. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 50-52 segments, filiform,

the apical 0.3 distinctly attenuate, 2” segment

1.4x, shortest segment 0.9x, penultimate segment

1.2x as long as wide. Pronotum rather shining,

rugose-punctate and with fine longitudinal wrin-

kles laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

finely and rather densely punctate on a finely

coriaceous and shining background. Scutellum

with lateral carinae on the anterior 0.1. Prescutel-

lar groove smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter

complete, joining the anterior edge of the meso-

pleurum. Transverse carina for the greater part

obliterated. Mesopleurum rather densely or dense-

ly punctate on a slightly coriaceous and shining

background. Speculum punctate posteriorly, fine-

ly granulate-strigose or with fine oblique striae

and shining anteriorly. Depression in front of the

speculum with short longitudinal wrinkles, which

merge with the punctation anteriorly. Metapleu-

rum rather coarsely and densely rugose on a

smooth background. Pleural carina complete or

partly covered with wrinkles. Areolet large, point-

ed or short-stalked, receiving the 2" recurrent

vein at or slightly basad of the middle. Nervellus

reclivous, distinctly intercepted. Nervellar index

0.6. Tibial spur index 0.7. Hind femur 4.7x as long

as high. Hind tarsal claws long, with distinct pec-

112:

tinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, broadly and

not deeply depressed, sulcus for the greater part

irregularly wrinkled, sometimes with few short

transverse wrinkles, often with some longitudinal

wrinkles posteriorly, costulae and longitudinal

carinae complete or almost complete, usually

strong. Propodeal spiracle large, 2.6x as long as

wide. Spiracular carina indistinct or obliterated.

Spiracular index 0.5. 1* gastral tergite 2.6x, 2"d

tergite 1.2x as long as wide. Petiole with large

glymmae, which are bordered by strong longitu-

dinal carinae dorsally, usually with a long row of

distinct short transverse wrinkles in front of the

glymmae (Fig. 35), rarely with distinct irregular

wrinkles instead. Petiolar suture complete. 1% gas-

tral sternite for the greater part coriaceous, with

transverse striae posteriorly. Thyridium oval.

Thyridial index about 0.5. Epipleurum separated

from the 3"! gastral tergite, the crease marked with

black. Epipleural index 0.7-0.8 (Fig. 39). Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.5.

Colour: Black. Maxillary palpus dark brown.

Wings slightly tinged with yellow. Stigma yel-

lowish brown medially. Front trochanter marked

with yellowish red. Front femur, tibia and tarsus

yellowish red. Mid femur completely yellowish

red or marked with black basally. Mid tibia and

tarsus yellowish red. Hind tibia for the greater

part yellowish red, not darkened basally, narrow-

ly black apically (the black ring about as long as

the greatest width of the tibia). 2" gastral tergite

red on the posterior 0.1-0.3. 3° and 4* tergites red.

Pilosity of most parts of the head dark brown, of

the thorax and propodeum light brown.

d: Propodeum scarcely depressed. Maxillary

palpus medium brown to light brown apically.

Mandible tinged with brown medially. Wings

sometimes hyaline. Pilosity of the head some-

times light brown. In other characters similar to

the ®.

Material (239): Russia: Irkutsk Prov.: Dachnaya (PET).

Kazakhstan: Kolovertnyy (MOS).

Dusona nigridens Horstmann, spec. nov.

Hinz (1994: 32) assigned the holotype (d) of

D. nigridens with some doubts to D. nubilator (?).

But D. nigridens differs from the latter in: Frons

with a very fine median longitudinal carina. Meta-

pleurum coriaceous and very densely punctate.

Propodeum without transverse striae. Petiole

smooth in front of the glymmae. Mandible black.

2X gastral tergite dorsally red posteriorly. Posteri-

or gastral tergites and sternites black.

Holotype (S): Japan, Honshu, Mt. Takao near

Hachioji/Tokyo, 16.4.1936 (AEI). This type is at

the same time a paratype of D. nubilator.

Description. d: Body length 12 mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 0.8. Frons slightly depressed, coriaceous, fine-

ly rugose, very finely and dispersely punctate,

with a very fine and partly obliterated median

longitudinal carina, without radial wrinkles. An-

tennal carina low and narrow. Face and dorsal 0.4

of the clypeus coriaceous, very densely punctate.

Clypeus flat, coriaceous and rather densely punc-

tate ventrally, its apex truncate medially. Malar

space index 0.4. Genal carina low and straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.4. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 52 segments, filiform,

the apical 0.3 distinctly attenuate, 2"? segment

1.7x, shortest segment 1.0x, penultimate segment

1.3x as long as wide. Pronotum densely punctate

dorsolaterally in the posterior corner, with longi-

tudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, very densely punctate.

Scutellum with lateral carinae on the anterior 0.4.

Prescutellar groove for the greater part smooth,

with few longitudinal wrinkles laterally. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the dorsal 0.2 of the latter obliterated.

Transverse carina obliterated. Mesopleurum dis-

tinctly coriaceous, densely punctate. Speculum co-

riaceous and granulate-strigose. Depression in

front of the speculum with fine and long longitu-

dinal striae, which merge with the punctation an-

teriorly. Metapleurum coriaceous, very densely

punctate. Pleural carina strong. Areolet rather

large, stalked, receiving the 2" recurrent vein dis-

tinctly basad of the middle. Nervellus slightly

inclivous, distinctly intercepted. Nervellar index

0.8. Tibial spur index 0.7. Hind femur 5.3x as long

as high. Hind tarsal claws short, with short pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, scarcely depressed, for the greater

part irregularly wrinkled, with some longitudinal

wrinkles posteriorly, costulae and longitudinal

carinae for the greater part obliterated. Propodeal

spiracle large, 2.7x as long as wide. Spiracular

carina narrow. Spiracular index 0.5. 1° gastral

tergite 4.2x, 2"X tergite 2.3x as long as wide. Peti-

ole with large glymmae, which are bordered by

longitudinal carinae dorsally, smooth in front of

the glymmae. Petiolar suture complete. 1“ gastral

sternite for the greater part smooth, with trans-

verse striae posteriorly. Thyridium oval. Thyridi-

al index about 1.0. Epipleurum separated from

the 3" gastral tergite anteriorly and folded in, the

crease marked with black. Epipleural index 0.3.

Gaster strongly compressed posteriorly.

Colour: Black. Maxillary palpus brownish

black. Wings hyaline. Stigma brownish black me-

dially. Front femur reddish brown dorsally, black

ventrally. Front tibia and tarsus yellowish red.

Mid tibia yellowish red. Mid tarsus dark brown.

Hind tibia yellowish red, narrowly marked with

brown basally and apically. 2” gastral tergite dor-

sally red on the posterior 0.2, laterally for the

greater part red. 3" to 5" tergites red, indistinctly

tinged with brown dorsally.

? unknown.

Dusona nigridorsum Horstmann, spec. nov.

Types. Holotype (?): Kazakhstan, Leninogorsk,

Ivanovskiy Mts., 13.8.1983 (PET). — Paratype: 15

Russia, Kuril Isl., Kunashir, Mendeleyevo, 2.8.

1981 (PET).

Description. 2: Body length 8 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.9. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate,

with a fine median longitudinal carina, with short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal half of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, coriaceous

and rather densely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina low and straight ventrally, joining the dis-

tinctly raised oral carina at an acute angle. Genal

index 0.4. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

48-49 segments, filiform, the apical 0.3 distinctly

attenuate, 2" segment 1.7x, shortest segment 1.0x,

penultimate segment 1.4x as long as wide. Prono-

tum coriaceous and densely rugose-punctate lat-

erally on the dorsal 0.4, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, densely punctate. Scutellum with lateral

carinae on the anterior 0.6. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter distinct, join-

ing or almost joining the anterior edge of the

mesopleurum. Transverse carina indicated ante-

113

riorly. Mesopleurum distinctly coriaceous, with

rather fine and rather dense or dense punctation.

Speculum coriaceous, rather dull. Depression in

front of the speculum coriaceous, with very fine

longitudinal striae or short wrinkles dorsally.

Metapleurum finely and densely punctate on a

finely coriaceous and shining background. Pleu-

ral carina distinct. Areolet large, stalked, receiv-

ing the 2" recurrent vein basad of the middle.

Nervellus vertical, weakly intercepted. Nervellar

index 0.8. Tibial spur index 0.6. Hind femur 4.9x

as long as high. Hind tarsal claws short, with

distinct pectinate teeth basally and medially. Pro-

podeum extending to 0.3 of the hind coxae, rather

broadly and not deeply depressed, the sulcus for

the greater part finely and irregularly wrinkled,

with distinct transverse wrinkles posteriorly, cos-

tulae partly present, longitudinal carinae indicat-

ed medially and posteriorly, obliterated anterior-

ly. Propodeal spiracle rather small, 4.6x as long as

wide. Spiracular carina narrow. Spiracular index

0.2. 1% gastral tergite 3.9x, 2" tergite 1.7x as long

as wide. Petiole with large glymmae, which are

bordered by longitudinal carinae dorsally, smooth

in front of the glymmae. Petiolar suture obliterat-

ed. 1“ gastral sternite smooth. Thyridium oval.

Thyridial index about 1.5. Epipleurum indicated

as a small triangular lobe anteriorly on the ventro-

lateral edge of the 3" gastral tergite, epipleurum

pendent in the ? holotype (but apparently folded

in in the d paratype), the crease not marked with

black, the tergite with distinct black lateral stripes

well above the ventrolateral edge (Fig. 41). Epi-

pleural index about 0.2. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown,

tinged with yellowish brown apically. Mandible

narrowly marked with yellow dorsally and medi-

ally. Wings hyaline. Stigma dark brown medially.

Front femur yellowish red, marked with black

ventrally. Front tibia and tarsus yellow. Mid fe-

mur yellowish red on the apical 0.2. Mid tibia and

tarsus dark yellowish brown. Hind tibia blackish,

marked with dark yellowish brown medially

on the dorsal side. 3“ gastral tergite narrowly

black dorsally, broadly marked with red laterally.

4" tergite broadly black dorsally, half red in later-

al view. Posterior sternites dark brown. Pilosity

whitish.

d: 2"X gastral tergite marked with red lateral-

ly. 5'° tergite narrowly marked with red laterally.

Posterior sternites and genital clasper reddish

brown. In other characters similar to the 9.

114

Dusona nigrina Horstmann, spec. nov.

Holotype (?): Russia, Primorye, Spassk Dalniy,

27.6.1985 (PER).

Description. 2: Body length 6 mm. Temples dis-

tinctly constricted behind the eyes, 0.6x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons slightly depressed, coriaceous, very fine-

ly and rather densely punctate, with a very fine

median longitudinal carina, with short and indis-

tinct radial wrinkles. Antennal carina low and

narrow. Face coriaceous, dull, with fine and indis-

tinct punctation and with fine wrinkles. Clypeus

somewhat rounded, coriaceous, dull, with fine

rather dense punctation, its apex truncate medial-

ly. Malar space index 0.3. Genal carina low and

straight ventrally, joining the scarcely raised oral

carina at an acute angle. Genal index 0.3. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 31 segments, fili-

form, the apical 0.2 slightly attenuate, 2”! segment

2.4x, shortest segment 1.0x, penultimate segment

1.0x as long as wide. Pronotum granulate-strigose

and with fine punctures laterally on the dorsal 0.2,

with longitudinal wrinkles ventrally. Mesoscu-

tum coriaceous, dull, finely and densely punctate.

Scutellum coriaceous, with very fine punctures,

with lateral carinae on the anterior 0.3. Prescutel-

lar groove for the greater part smooth. Prepectal

carina low, ventral part merging with the pleural

part, the latter complete, somewhat indistinct dor-

sally. Transverse carina indicated anteriorly. Meso-

pleurum coriaceous, dull, with fine and disperse

or rather dense punctation. Speculum partly

smooth, strongly shining. Depression in front of

the speculum with fine longitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum coriaceous, with fine rather dense puncta-

tion. Pleural carina distinct. Areolet small, stalked,

receiving the 2" recurrent vein distinctly distad of

the middle. Nervellus distinctly inclivous, weakly

intercepted. Nervellar index 0.8. Tibial spur index

0.5. Hind femur 3.7x as long as high. Hind tarsal

claws short, with few long pectinate teeth basally

and medially. Propodeum extending to 0.3 of the

hind coxae, rather narrowly depressed only medi-

ally, not depressed anteriorly and posteriorly, co-

riaceous anteriorly and medially, with distinct

transverse wrinkles posteriorly, costulae indicat-

ed, longitudinal carinae almost completely oblit-

erated. Propodeal spiracle rather small, 1.8x as

long as wide. Spiracular carina partly widened.

Spiracular index 0.5. 1% gastral tergite 3.7x, 2"d

tergite 1.5% as long as wide. Petiole with small but

distinct glymmae, smooth in front of the glym-

mae. Petiolar suture almost complete. 1“ gastral

sternite smooth. Thyridium oval. Thyridial index

about 2.0. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.7. Gaster somewhat compressed

posteriorly. Ovipositor index 0.8.

Colour: Black. Maxillary palpus yellow. Man-

dible almost completely yellow. Tegula yellow.

Wings hyaline. Stigma light brown medially. Front

and mid legs from the trochanter onwards yellow,

the mid trochanter marked with black, the femora

yellowish red. Hind tibia whitish yellow medial-

ly, narrowly black basally, broadly black apically.

Hind tarsus for the greater part dark brown,

marked with yellowish brown basally. Pilosity

whitish.

d unknown.

Dusona nigrotibialis Horstmann, spec. nov.

Holotype (?): China, Hebei Prov., Qilaotu Shan

Mts., N of Chengde, 1600 m, 1.6.2000 (LIN).

Description. 2: Body length 7 mm. Temples

strongly constricted behind the eyes, 0.9x as long

asthe width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons slightly depressed, coriaceous, partly

with very fine wrinkles, with a fine and partly

obliterated median longitudinal carina, with few

short radial wrinkles. Antennal carina low and

narrow. Face and dorsal half of the clypeus coria-

ceous, finely and densely rugose-punctate. Clypeus

slightly rounded, coriaceous and rather densely

punctate ventrally, its apex truncate medially.

Malar space index 0.3. Genal carina low and

straight ventrally, joining the low oral carina at an

acute angle. Genal index 0.3. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 35 segments, slightly fusi-

form, 2"! segment 1.8x, shortest segment 0.8x,

penultimate segment 1.0x as long as wide. Prono-

tum coriaceous and with fine wrinkles laterally on

the dorsal 0.3, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous, dense-

ly punctate. Scutellum with lateral carinae on the

anterior 0.4. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter complete, joining the anterior

edge of the mesopleurum. Transverse carina

present anteriorly, low. Mesopleurum coriaceous,

with fine and rather dense or dense punctation.

Speculum finely coriaceous, shining. Depression

in front of the speculum with fine and long longi-

tudinal striae, which merge with the punctation

anteriorly. Metapleurum coriaceous, with fine ir-

regular wrinkles, partly finely rugose-punctate.

Pleural carina distinct. Areolet rather large, point-

ed, receiving the 2" recurrent vein distinctly dis-

tad of the middle. Nervellus inclivous, distinctly

intercepted. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 4.7x as long as high. Hind tarsal

claws short, with distinct pectinate teeth basally

and medially. Propodeum extending to 0.3 of the

hind coxae, rather broadly and not deeply de-

pressed, the rather shallow sulcus for the greater

part coriaceous and with some fine irregular wrin-

kles, with distinct longitudinal wrinkles posteri-

orly, carinae almost completely obliterated. Pro-

podeal spiracle rather large, 2.2x as long as wide.

Spiracular carina widened. Spiracular index 0.4.

1% gastral tergite 3.2x, 2"X tergite 1.6x as long as

wide. Petiole with large glymmae, which are bor-

dered by longitudinal carinae dorsally, smooth in

front of the glymmae. Petiolar suture almost com-

plete. 1% gastral sternite smooth. Thyridium indis-

tinct, oval. Thyridial index about 1.0. Epipleurum

separated from the 3"! gastral tergite, the crease

marked with black. Epipleural index 0.6. Gaster

distincetly compressed posteriorly. Ovipositor in-

dex 0.8.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible broad-

ly yellow medially. Wings hyaline. Stigma yel-

lowish brown medially. Front femur black basally

and ventrally, yellowish brown dorsally and api-

cally. Front tibia and tarsus yellow. Mid tibia and

tarsus yellowish brown. 2" gastral tergite red on

the posterior 0.2. 3 tergite for the greater part

red, dorsally marked with black posteriorly. Pi-

losity whitish.

& unknown.

Dusona nitidipleuris Horstmann, spec. nov.

The species is similar to D. variator, it differs from

the latter in: Mesopleurum and metapleurum less

strongly sculptured and more shining. Propodeum

without a medium longitudinal keel posteriorly.

Red markings on the gaster more extended.

Holotype (2): Russia, Primorye, Ussuriysk, 6.

1929 (MOS).

Description. 2: Body length 9mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 0.9. Frons scarcely depressed, coriaceous, fine-

115

ly and densely punctate, with short radial wrin-

kles, amedian longitudinal carina weakly indicat-

ed by some wrinkles. Antennal carina low and

narrow. Face and dorsal 0.6 of the clypeus coria-

ceous, finely and densely rugose-punctate. Clypeus

flat, coriaceous and rather densely punctate ven-

trally, its apex almost truncate medially. Malar

space index 0.4. Genal carina low and almost

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.5. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with at least 42 seg-

ments (both flagellar tips broken off in the only

available specimen), filiform, the apical 0.3 atten-

uate, 2"7 segment 1.7x, shortest segment 0.9x as

long as wide. Pronotum with dense and irregular

wrinkles laterally on the dorsal 0.4, with longitu-

dinal wrinkles ventrally. Mesoscutum coriaceous,

densely rugose-punctate. Scutellum finely coria-

ceous, densely punctate, with lateral carinae on

the anterior 0.4. Prescutellar groove almost smooth

medially, with longitudinal striae laterally. Pre-

pectal carina low, ventral part merging with the

pleural part, the latter almost joining the anterior

edge of the mesopleurum. Transverse carina ob-

literated. Mesopleurum finely and rather densely

punctate on a finely coriaceous and shining back-

ground. Speculum coriaceous and finely granu-

late-strigose, somewhat shining. Depression in

front of the speculum with distinct rather short

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum finely rugose, with

very fine sculpture and shining centrally. Pleural

carina complete. Areolet rather large, short-stalked,

receiving the 2" recurrent vein distad of the mid-

dle. Nervellus slightly inclivous, distinctly inter-

cepted. Nervellar index 0.8. Tibial spur index 0.7.

Hind femur 4.5x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally. Propo-

deum extending to 0.4 of the hind coxae, scarcely

depressed, for the greater part with fine irregular

wrinkles, rather shining, with distinct transverse

wrinkles posteriorly, carinae almost completely

obliterated. Propodeal spiracle rather large, 3.5x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.4. 1% gastral tergite 3.0x, 2"4 tergite

2.0x as long as wide. Petiole with distinct glym-

mae, which are bordered by longitudinal carinae

dorsally, smooth in front of the glymmae. Petiolar

suture partly obliterated. 1% gastral sternite for the

greater part smooth, with fine transverse triae

posteriorly. Thyridium oval. Thyridial index about

2.0. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

116

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly tinged with brown medially.

Wings hyaline. Stigma medium brown medially.

Front femur yellowish red, marked with black

ventrally. Front tibia and tarsus yellow. Mid fe-

mur yellow on the apical 0.2. Mid tibia yellow,

darkened ventrally. Mid tarsus dark brown.

2"d gastral tergite red on the posterior 0.3. 3" and

4" tergites red. Pilosity whitish.

d unknown.

Dusona norikurae Hınz & Horstmann,

spec. nov.

This is a variable and poorly defined species.

Types. Holotype (2): Japan, Honshu, Norikura-

dake, 2000 m, 30.7.1954 (AEI). — Paratypes: 1?

same locality, 2200 m, 29.7.1954 (Hinz /ZSM); 1?

Japan, Hokkaido, Gorinzawa, 24.6.1944 (SAP); 1?

Russia, Primorye, Ussuriysk, 17.7.1981 (PET); 1?

Russia, Chita Prov., Nerchinskiy Zavod, 20.7.1975

(PET); 254 Japan, Honshu, Tachikawa, 2.5.1931

(138 AEI, 18 Hinz/ZSM); 13 Japan, Hokkaido,

Sapporo, Maruyama, 28.6.1957 (SAP); 19 Russia,

Khabarovsk Terr., Khekhtsir, near Khabarovsk,

4.7.1983 (PET).

Description. ?: Body length 8-12 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.8. Frons slightly de-

pressed, coriaceous, finely rugose-punctate, with

a fine median longitudinal carina, with short radi-

al wrinkles, which extend to 0.3 the distance to the

median ocellus. Antennal carina very low and

narrow. Face and dorsal 0.6 of the clypeus densely

rugose-punctate. Clypeus flat, coriaceous and dis-

persely punctate ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

and rather straight ventrally, joining the scarcely

raised oral carina at an acute angle. Genal index

0.4. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 43-49

segments, filiform, the apical 0.3 distinctly attenu-

ate, 2"! segment 1.9x, shortest segment 1.0x, pe-

nultimate segment 1.3x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.2, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, densely

punctate. Scutellum with lateral carinae on the

anterior 0.6. Prescutellar groove smooth or with

fine longitudinal wrinkles. Prepectal carina low,

ventral part merging with the pleural part, the

latter complete, joining the anterior edge of the

mesopleurum. Transverse carina obliterated. Me-

sopleurum distinctly coriaceous, densely or rath-

er densely punctate. Speculum granulate-strigose,

shining. Depression in front of the speculum with

fine and long longitudinal striae, which merge

with the punctation anteriorly, or almost com-

pletely coriaceous without striae. Metapleurum

with fine and dense irregular wrinkles or densely

rugose-punctate. Pleural carina distinct. Areolet

large, short-stalked, receiving the 2" recurrent

vein basad of the middle. Nervellus slightly in-

clivous, intercepted. Nervellar index 0.7. Tibial

spur index 0.6. Hind femur 5.8x as long as high.

Hind tarsal claws rather long, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to 0.4 of the hind coxae, distinctly and

rather broadly depressed, sulcus for the greater

part irregularly wrinkled and dull, with some

transverse wrinkles posteriorly, costulae and lon-

gitudinal carinae obliterated or partly indicated.

Propodeal spiracle large, 2.6x as long as wide.

Spiracular carina narrow. Spiracular index 0.3.

1* gastral tergite 4.1x, 2% tergite 2.3x as long as

wide. Petiole with distinct and rather large glym-

mae, finely coriaceous or almost smooth in frent

of the glymmae. Petiolar suture indicated. 1” gas-

tral sternite for the greater part smooth, with short

transverse striae postriorly. Thyridium oval. Thy-

ridial index about 2.0. Epipleurum separated from

the 3" gastral tergite anteriorly, either folded in or

pendent as a narrow rectangular lobe, with a black

lateral stripe parallel with the ventrolateral edge

in a short distance. Epipleural index 0.2. Gaster

distinctly compressed posteriorly. Ovipositor in-

dex 0.5.

Colour: Black. Maxillary palpus brownish

black. Wings hyaline. Stigma medium brown me-

dially. Front femur yellowish red dorsally. Front

tibia and tarsus yellowish red. Mid femur tinged

with brown apically. Mid tibia yellowish brown.

Hind tibia blackish, sometimes tinged with dark

brown medially. 2” gastral tergite dorsally black,

laterally marked with red subapically, sometimes

almost completely black. 3“ and 4" tergites red.

5 tergite narrowly marked with red anteriorly.

Pilosity whitish.

d: Maxillary palpus dark brown basally, yel-

lowish brown apically. Tip of the front coxa and

front trochanter yellow or black. Mid femur

marked with yellow. Mid tibia yellow. 3" and 4"

gastral tergites sometimes marked with black dor-

sally. In other characters similar to the 9.

Dusona nubilator Hinz, 1994

So far, the holotype (?) is the only known speci-

men belonging to this species. One paratype (?)

could not be located. The other paratype (6) is

described as a separate species (D. nigridens).

D. nubilator is similar to D. peptor, it differs from

the latter in: Temples shorter and more constrict-

ed behind the eyes. Wings hyaline. Gaster less

extensively marked with red.

Description. 2: Body length 12 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.9. Frons slightly depressed, coriaceous, finely

and densely rugose-punctate, with a distinctly

raised median longitudinal carina, with short ra-

dial wrinkles. Antennal carina low and narrow.

Face and dorsal half of the clypeus coriaceous,

finely and densely rugose-punctate. Clypeus flat,

coriaceous and dispersely or rather densely punc-

tate ventrally, its apex truncate medially. Malar

space index 0.4. Genal carina low and straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.4. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 50 segments, filiform,

the apical 0.2 distinctly attenuate, 2°’ segment

1.8x, shortest segment 1.0x, penultimate segment

1.4x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.2, with longitu-

dinal wrinkles ventrally. Mesoscutum coriaceous,

very densely punctate. Scutellum coriaceous, dis-

persely or rather densely punctate, with lateral

carinae on the anterior half. Prescutellar groove

with fine longitudinal striae. Prepectal carina low,

ventral part merging with the pleural part, the

latter complete, joining the anterior edge of the

mesopleurum. Transverse carina present anteri-

orly. Mesopleurum distinctly coriaceous, distinct-

ly and densely punctate. Speculum coriaceous and

granulate-strigose, shining. Depression in front of

the speculum with fine and long longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum densely and finely rugose. Pleural

carina strong. Areolet rather large, short-stalked,

receiving the 2"! recurrent vein slightly basad of

the middle. Nervellus distinctly inclivous, dis-

tinctly intercepted. Nervellar index 0.8. Tibial spur

index 0.7. Hind femur 5.3x as long as high. Hind

tarsal claws short, with distinct pectinate teeth

basally and medially. Propodeum extending to

0.3 of the hind coxae, not deeply depressed, sul-

cus narrow anteriorly, broad posteriorly, more

111177

irregularly wrinkled anteriorly and medially, dis-

tinctly transversely wrinkles posteriorly, costulae

partly present, longitudinal carinae distinct medi-

ally and posteriorly. Propodeal spiracle large, 2.9x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.5. 1* gastral tergite 3.3x, 2"4 tergite

1.5x as long as wide. Petiole with large glymmae,

which are bordered by longitudinal carinae dor-

sally, finely coriaceous in front of the glymmae,

with dorsolateral longitudinal carinae and some

distinct transverse wrinkles anteriorly. Petiolar

suture almost complete. 1% gastral sternite coria-

ceous anteriorly, smooth medially, transversely

striate posteriorly. Thyridium oval. Thyridial in-

dex about 2.0. Epipleurum separated from the 3°“

gastral tergite anteriorly and folded in, a black

lateral stripe parallel with the crease in a short

distance anteriorly, divergent from the ventrola-

teral edge of the tergite posteriorly. Epipleural

index 0.2. Gaster distinctly compressed posterior-

ly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus black basal-

ly, yellowish brown apically. Mandible broadly

marked with yellowish brown medially. Wings

hyaline. Stigma dark brown medially. Front tro-

chanter brown, marked with black. Front femur

yellowish red, marked with dark brown ventral-

ly. Front tibia and tarsus yellow. Mid femur yel-

lowish red dorsally, black ventrally. Mid tibia yel-

lowish. Mid tarsus medium to dark brown. Hind

tibia yellowish brown medially, narrowly marked

with black basally, rather broadly marked with

dark brown apically (the dark apical mark being

broader dorsally than ventrally). 2”! gastral tergite

black dorsally, narrowly marked with red lateral-

ly. 3“ and 4 tergites red, tinged with brown

dorsally. 5" tergite red, marked with black dorsal-

ly. Posterior tergites for the greater part black,

narrowly marked with red ventrolaterally. Poste-

rior gastral sternites reddish brown, marked with

black. Pilosity whitish.

d unknown.

Material (1?): Japan: Honshu: Sasayama (SAP).

Dusona obliterata (Holmgren, 1872)

Description. 2: Body length 14-17 mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyesand

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 1.0. Frons slightly de-

pressed, coriaceous and rather densely punctate

dorsally, finely coriaceous and rather shining ven-

trally, with a fine and partly obliterated median

118

longitudinal carina, with distinct radial wrinkles,

which extend to 0.3 the distince to the median

ocellus. Antennal carina low, the rim slightly bent

upwards and slightly transversely striate. Face

and dorsal 0.7 of the clypeus coriaceous, coarsely

and densely punctate. Clypeus almost flat, coria-

ceous and rather densely punctate ventrally, its

apex truncate medially. Malar space index 0.3.

Genal carina slightly raised and slightly bent side-

wards ventrally, joining the slightly raised oral

carina at an acute angle. Gena slightly depressed

close to the junction (a variable character). Genal

index 0.4. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

59-66 segments, filiform, the apical 0.4 distinctly

attenuate, 2”! segment 1.5x, shortest segment 0.9x,

penultimate segment 1.6x as long as wide. Prono-

tum coriaceous and densely rugose-punctate lat-

erally on the dorsal 0.3, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, dull, rather densely or densely punctate.

Scutellum with lateral carinae on the anterior 0.7.

Prescutellar groove smooth medially, with fine

longitudinal striae laterally. Prepectal carina low,

ventral part merging with the pleural part, the

latter almost complete, the dorsal 0.1-0.2 often

replaced by several wrinkles. Transverse carina

complete or partly obliterated. Mesopleurum rath-

er densely or densely punctate on a finely coria-

ceous and shining background. Speculum finely

coriaceous, rather shining. Depression in front of

the speculum dorsally with fine longitudinal stri-

ae, which merge with the punctation anteriorly,

the ventral 0.4 granulate-strigose. Metapleurum

densely rugose, sometimes coriaceous and very

densely punctate. Pleural carina distinct. Areolet

large, pointed, receiving the 2" recurrent vein

slightly basad of the middle. Nervellus almost

vertical, strongly intercepted. Nervellar index 0.7.

Tibial spur index 0.5. Hind femur 5.5x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to 0.4 of the hind coxae, broadly and

deeply depressed, sulcus coriaceous in the mid-

line, with irregular or transverse wrinkles at the

sides, with some transverse wrinkles posteriorly,

costulae partly obliterated, longitudinal carinae

obliterated anteriorly and medially, strong poste-

riorly. Propodeal spiracle large, 4.0x as long as

wide. Spiracular carina narrow. Spiracular index

0.3. 1% gastral tergite 4.6x, 2" tergite 2.0x as long

as wide. Petiole with distinct glymmae, smooth in

front of the glymmae. Petiolar suture partly oblit-

erated. 1* gastral sternite smooth. Thyridium oval.

Thyridial index about 1.5. Epipleurum not sepa-

rated from the 3"! gastral tergite, the latter without

black lateral stripes. Gaster strongly compressed

posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish

brown. Mandible yellow medially. Tegula yel-

lowish brown. Wings hyaline. Stigma yellowish

brown medially. Tip of front coxa yellow. Front

trochanter and femur yellowish red. Front tibia

and tarsus for the greater part yellow. Mid femur

yellowish red on the apical 0.3. Mid tibia yellow.

Mid tarsus brown. Hind femur yellowish red on

the apical 0.2. Hind tibia yellow, marked with red

apically. Hind tarsus dark brown. 2" gastral tergite

dorsally black or red on the posterior 0.2, almost

completely red laterally. 3“ and 4" tergites light

red. 5" tergite marked with light red anteriorly

and/or ventrolaterally. Posterior gastral sternites

marked with red. Pilosity whitish.

d: Longitudinal carinae of propodeum some-

times present medially. 3" and 4" gastral tergites

sometimes marked with black dorsally. In other

characters similar to the 2.

Material (222): Russia: Irkutsk Prov.: Lake Baykal,

Irkutsk (PET).

Dusona obscurator Horstmann, spec. nov.

Types. Holotype (d): Russia, Primorye, Vladi-

vostok, 23.6.1985 (PET). - Paratype: 15 Russia,

Primorye, Pozhiga SO of Dalnerechensk, 25.7.

1983 (Hinz/ZSM).

Description. d: Body length 10-11 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.8. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate,

with a fine median longitudinal carina, with short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.7 of the clypeus coriaceous,

finely and densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low and rather straight ventrally,

joining the distinctly raised oral carina at an acute

angle. Genal index 0.7. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 46-48 segments, filiform, the api-

cal 0.2 attenuate, 2" segment 2.0x, shortest seg-

ment 1.0-1.1x, penultimate segment 1.5x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.3. Prescutellar groove almost

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter distinctly join-

ing the anterior edge of the mesopleurum. Trans-

verse carina indicated anteriorly. Mesopleurum

finely but distinctly coriaceous, densely punctate.

Speculum granulate-strigose. Depression in front

of the speculum with fine longitudinal striae and

wrinkles, which merge with the punctation ante-

riorly. Metapleurum coriaceous, densely rugose-

punctate. Pleural carina strong. Areolet large,

short-stalked, receiving the 2” recurrent vein

slightly distad of the middle. Nervellus slightly

reclivous, distinctly intercepted. Nervellar index

0.7. Tibial spur index 0.7. Hind femur 5.1x as long

as high. Hind tarsal claws short, with short pecti-

nate teeth basally. Propodeum extending to 0.3 of

the hind coxae, slightly and rather narrowly de-

pressed, for the greater part with rather fine irreg-

ular wrinkles, with some longitudinal wrinkles

posteriorly, costulae and longitudinal carinae com-

plete. Propodeal spiracle rather large, 2.9x as long

as wide. Spiracular carina narrow. Spiracular in-

dex 0.6. 1" gastral tergite 3.7x, 2" tergite 2.0x as

long as wide. Petiole with distinct glymmae, which

are bordered by longitudinal carinae dorsally, with

some fine longitudinal wrinkles in front of the

glymmae, with some transversal wrinkles anteri-

orly. Petiolar suture distinct. 1" gastral sternite

coriaceous and dull anteriorly, transversely stri-

ate posteriorly. Thyridium oval. Thyridial index

about 1.0. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.9. Gaster distinctly compressed

posteriorly.

Colour: Black. Mandible completely black or

narrowly marked with yellowish brown dorsally.

Wings hyaline. Stigma medium brown medially.

Front leg from the femur onwards yellowish red,

femur marked with dark brown ventrally. Mid

femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. 2"4 gastral tergite reddish

brown on the posterior 0.1. 3" tergite broadly

black dorsally, about half reddish brown in lateral

view. 4" tergite tinged with brown laterally. Pilos-

ity whitish.

® unknown.

119

Dusona obscuripes Horstmann, spec. nov.

Holotype (3): Russia, Kamchatka, Dalniy, 50 km

W of Yelizovo, 2.8.1967 (PET).

Description. d: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.7. Frons not depressed, coriaceous, densely ru-

gose-punctate, with a distinct and slightly raised

median longitudinal carina, with short and indis-

tinct radial wrinkles. Antennal carina low and

narrow. Face and dorsal 0.7 of the clypeus coria-

ceous, densely rugose. Clypeus flat, finely coria-

ceous and densely punctate ventrally, its apex

truncate medially. Malar space index 0.5. Genal

carina low ventrally, slightly bent sidewards and

forwards, joining the oral carina at the base of the

mandible almost at a right angle. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 39 segments, filiform,

the apical 0.2 distinctly attenuate, 2”! segment

1.7x, shortest segment 1.0x, penultimate segment

1.4x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.3, with longitu-

dinal wrinkles ventrally. Mesoscutum coriaceous,

very densely punctate. Scutellum densely rugose-

punctate, with lateral carinae on the anterior 0.4.

Prescutellar groove with fine longitudinal striae.

Ventral part of the prepectal carina merging with

the transverse carina, both carinae rather low.

Pleural part of the prepectal carina obliterated

dorsally and ventrally, the median 0.3 present as a

wrinkle. Mesopleurum very densely punctate,

partly rugose, on a finely coriaceous or almost

smooth background. Speculum and depression in

front of the speculum with long longitudinal stri-

ae and wrinkles, which merge with the puncta-

tion anteriorly. Metapleurum distinctly irregular-

ly wrinkled. Pleural carina partly indistinct and

covered with wrinkles. Areolet rather large, short-

stalked, receiving the 2" recurrent vein distinctly

distad ofthe middle. Nervellus distinctly inclivous,

intercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 5.3x as long as high. Hind tarsal

claws short, with short pectinate teeth basally.

Propodeum extending to 0.6 of the hind coxae,

distinctly but narrowly and not deeply depressed,

with short transverse wrinkles in the narrow lon-

gitudinal furrow, coarsely and irregularly wrin-

kled beside the furrow, costulae almost complete,

median longitudinal carinae present anteriorly,

covered with wrinkles medially and posteriorly,

lateral longitudinal carinae present. Propodeal

spiracle small, 1.8x as long as wide. Spiracular

120

carina narrow and indistinct. Spiracular index 0.7.

1* gastral tergite 5.4x, 2" tergite 2.7x as long as

wide. Petiole without glymmae or narrow lateral

areas, smooth laterally. Petiolar suture for the

greater part obliterated. 1” gastral sternite for the

greater part smooth, with transverse striae poste-

riorly. Thyridium elongate. Thyridial index about

1.5. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly.

Colour: Black. Maxillary palpus medium

brown basally, yellowish brown medially and api-

cally. Mandible broadly yellow medially. Wings

hyaline. Stigma dark brown medially. Front fe-

mur dark reddish brown dorsally and apically.

Front tibia and tarsus dark yellowish brown.

2" gastral tergite red on the posterior 0.1.-3' and

4" tergites red, partly tinged with dark brown.

Pilosity whitish.

? unknown.

Dusona obtutor Hinz, 1994

This species is not well understood. The associa-

tion of the sexes is not beyond all doubt.

Description. 2: Body length 10 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.4. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate,

with a fine median longitudinal carina, with few

indistinct radial wrinkles. Antennal carina low

and narrow. Face and dorsal 0.4 of the clypeus

coriaceous, finely and densely rugose-punctate.

Clypeus slightly concave, finely coriaceous and

rather densely punctate ventrally, its apex truncate

or slightly concave medially. Malar space index

0.3. Genal carina low and straight ventraily, join-

ing the distinctly raised oral carina at an acute

angle. Genal index 0.4. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

ellım with 43 segments, filiform, the apical 0.3

attenuate, 2”! segment 2.6x, shortest segment 1.2x,

penultimate segment 1.4x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, very densely

punctate. Scutellum with lateral carinae on the

anterior 0.7. Prescutellar groove with longitudinal

striae. Prepectal carina low, ventral part merging

with the pleural part, the dorsal 0.3 of the latter

obliterated. Transverse carina present only anteri-

orly. Mesopleurum coriaceous, dull, very densely

punctate. Speculum finely granulate-strigose, shin-

ing. Depression in front of the speculum dorsally

with distinct longitudinal striae, the ventral 0.2

with very fine striae, the striae merging with the

punctation anteriorly. Metapleurum densely ru-

gose. Pleural carina distinct. Areolet rather small,

stalked, receiving the 2” recurrent vein slightly

distad of the middle. Nervellus slightly reclivous,

weakly intercepted. Nervellar index 0.7. Tibial spur

index 0.5. Hind femur 5.2x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to the middle of the

hind coxae, distinctly but not deeply and rather

narrowly depressed, sulcus for the greater part

with transverse wrinkles, costulae complete, lon-

gitudinal carinae present medially and posterior-

ly, obliterated anteriorly. Propodeal spiracle large,

4.6x as long as wide. Spiracular carina partly wid-

ened. Spiracular index 0.4. 1“ gastral tergite 4.4x,

2"d tergite 2.1x as long as wide. Petiole without

glymmae, with indistinct narrow lateral areas, with

very fine longitudinal striae in front of these areas,

partly smooth. Petiolar suture for the greater part

obliterated. 1* gastral sternite for the greater part

with very fine sculpture, with few transverse striae

posteriorly. Thyridium elongate. Thyridial index

1.3. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus medium

brown, partly marked with yellow. Mandible nar-

rowly marked with yellowish brown dorsally.

Wings hyaline. Stigma dark brown medially. Front

femur marked with yellowish brown on the apical

0.3. Front tibia and tarsus yellowish brown. Mid

leg brownish black, the tibia marked with yellow

basally and apically. 2" gastral tergite red on the

posterior 0.1. 3" tergite red. 4" and 5!" tergites

marked with red anteriorly. Pilosity whitish.

d: Body length 9 mm. Flagellum with 41-42

segments, 2" segment 1.8x, shortest segment 1.1x,

penultimate segment 1.4x as long as wide. Meta-

pleurum coriaceous and very densely punctate.

Propodeum rather shallowly depressed. Propo-

deal spiracle 3.0x as long as wide. Spiracular car-

ina narrow. Spiracular index 0.6. Maxillary palpus

yellow. Mandible broadly yellow medially. Front

leg from the tip of the coxa onwards yellowish

red. Mid femur yellowish brown on the apical 0.3.

Mid tibia yellowish brown. 2" gastral tergite com-

pletely black dorsally. 3" tergite narrowly marked

with black dorsally. In other characters similar to

the 2. The differences from the ? might be inter-

preted as sexual differences.

Material (17, 2343): Japan: Honshu: Kamikochi near

Norikura-dake (AEI), Kyoto (SAP), Norikura-dake,

1500 m (Hinz/ ZSM).

Variety: Body length 7 mm. Flagellum with 39

segments. Glymmae present, but small and shal-

low. 3“ and 4" gastral tergites completely red.

Material: 12? Japan, Hokkaido, Sapporo (SAP). This

? is a paratype of D. micrator. Its status is un-

known.

Dusona okadai (Uchida, 1942)

Description. 2: Body length 11-13 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons deeply depressed above the antennal

carinae, densely rugose-punctate dorsally, almost

smooth and with some distinct radial wrinkles in

the depressions, with a distinct median longitudi-

nal bulge between the depressions and with a

distinct longitudinal carina on top of this bulge.

Antennal carina strongly raised, with a strong

dorsal projection (nose-like) and with fine trans-

verse striae on this projection. Face and clypeus

densely rugose-punctate and rugose. Clypeus flat,

its apex slightly convex medially. Malar space

index 0.3. Genal carina low and almost straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.6. Temple finely

coriaceous and with rather dense punctation me-

dially. Flagellum with 56-60 segments, filiform,

the apical 0.3 distinctly attenuate, 2"? segment

1.4x, shortest segment 0.9x, penultimate segment

1.4x as long as wide. Pronotum densely rugose-

punctate on asmooth background laterally on the

dorsal 0.3, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum densely rugose-punc-

tate on a finely coriaceous and shining background.

Scutellum partly with a smooth background, with

lateral carinae on the anterior 0.8. Prescutellar

groove with distinct longitudinal striae. Prepectal

carina low, ventral part merging with the pleural

part, the latter almost complete, the dorsal 0.1-0.2

replaced by several wrinkles. Transverse carina

present anteriorly. Mesopleurum densely punc-

tate on asmooth background. Speculum very fine-

ly coriaceous, partly smooth, strongly shining.

Depression in front of the speculum with distinct

and long longitudinal striae, which merge with

the punctation anteriorly. Metapleurum densely

rugose-punctate, partly rugose. Pleural carina dis-

tinct. Areolet rather small, pointed, receiving the

2" recurrent vein at the middle. Nervellus almost

vertical, distinctly intercepted. Nervellar index 0.7.

121

Tibial spur index 0.6. Hind femur 5.0x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally and medially. Propodeum extending

to the middle of the hind coxae, rather broadly

and deeply depressed, sulcus for the greater part

with transverse wrinkles, costulae obliterated, lon-

gitudinal carinae strong posteriorly, obliterated

medially and anteriorly. Propodeal spiracle large,

4.9x as long as wide. Spiracular carina narrow.

Spiracular index 0.2. 1" gastral tergite 4.8x, 2"

tergite 2.0x as long as wide. Petiole with narrow

lateral areas, without glymmae or with small and

shallow glymmae at the anterior edge of the later-

al areas, smooth in front of these areas. Petiolar

suture obliterated. 1* gastral sternite for the great-

er part smooth, with transverse striae posteriorly.

Thyridium elongate. Thyridial index about 1.0.

Epipleurum not separated from the 3“ gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster strongly

compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellowish red

dorsally and medially. Tegula yellowish brown.

Wings hyaline. Stigma medium brown medially.

Front leg from the femur onwards yellow, the

femur marked with brown ventrally. Mid femur

marked with yellow on the apical 0.2. Mid tibia

yellow, marked with brown apically. Mid tarsus

light brown. Hind tibia whitish yellow subbasal-

ly, narrowly black basally, very broadly black api-

cally (black apical ring extending over 0.4 of the

tibia). 2" gastral tergite dorsally red on the apical

0.1, laterally red on the apical 0.4. 3"! and 4" tergites

red. Pilosity whitish.

d: 4" gastral tergite red only anteriorly. In

other characters similar to the 9.

Material (2??, 458): Russia: Yevreyskiy A.O.: Amur-

zet (PET); Primorye: Khasan (KIS). China: Liaoning

Prov.: Kaigen (= Kaiyuan) (SAP). South Korea: Suigen

(= Suwon) (SAP).

Dusona opaca (Thomson, 1887)

This species is similar to D. opacoides, it differs

from the latter in: Body larger. Flagellum stouter.

Temple rather densely punctate. Areolet larger.

Propodeal spiracle longer. Glymma smaller.

Description. 2: Body length 9-10 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 1.0. Frons scarcely depressed, coriaceous,

finely and densely rugose-punctate, with a dis-

tinct median longitudinal carina, with short radial

wrinkles. Antennal carina low and narrow. Face

and dorsal half of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely punctate ventrally, its apex slightly

convex medially. Malar space index 0.6. Genal

carina low and straight ventrally, joining the dis-

tinctly raised oral carina at an acute angle. Genal

index 0.4. Temple coriaceous and with fine rather

dense or dense punctation medially. Flagellum

with 43-47 segments, filiform, the apical 0.2 atten-

uate, 2"! segment 1.8x, shortest segment 1.0x, pen-

ultimate segment 1.2x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely or very

densely punctate. Scutellum with lateral carinae

on the anterior 0.8. Prescutellar groove with fine

longitudinal striae. Prepectal carina low, ventral

part merging with the pleural part, the dorsal 0.3

of the latter replaced by several wrinkles. Trans-

verse carina more or less obliterated (Fig. 18).

Mesopleurum coriaceous, dull, densely punctate.

Speculum coriaceous, dull. Depression in front of

the speculum with rather short longitudinal wrin-

kles, which merge with the punctation anteriorly.

Metapleurum densely rugose. Pleural carina com-

plete. Areolet rather large, short-stalked, receiv-

ing the 2"! recurrent vein almost at the middle.

Nervellus almost vertical, weakly intercepted.

Nervellar index 0.6-0.7. Tibial spur index 0.8. Hind

femur 5.0x as long as high. Hind tarsal claws long,

with short pectinate teeth basally and medially.

Propodeum extending to 0.4 of the hind coxae,

broadly and not deeply depressed, sulcus for the

greater part irregularly wrinkled, with short trans-

verse wrinkles in the midline and with long trans-

verse wrinkles posteriorly, carinae almost com-

pletely obliterated. Propodeal spiracle rather large,

4.0x as long as wide. Spiracular carina replaced by

several wrinkles. Spiracular index 0.6. 1‘ gastral

tergite 3.8x, 2" tergite 1.7x as long as wide. Peti-

ole with very small or indistinct glymmae at the

anterior edge of indistinct narrow lateral areas,

smooth in front of these areas. Petiolar suture

almost completely obliterated. 1 gastral sternite

for the greater part smooth, with fine transverse

striae posteriorly. Thyridium oval. Thyridial in-

dex about 1.5. Epipleurum not separated from the

34 gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.5.

Colour: Black. Mandible narrowly yellowish

brown medially. Wings hyaline. Stigma yellowish

brown medially. Front femur black basally and

ventrally, yellowish red dorsally and apically.

Front tibia and tarsus yellowish red. Mid femur

yellowish red on the apical 0.3. Mid tibia yellow-

ish red. Mid tarsus brown. Hind tibia dark yel-

lowish brown, narrowly black basally, almost not

darkened apically. Hind tarsus für the greater

part brownish black, the basitarsus yellowish

brown ventrally. 2" gastral tergite red on the

posterior 0.3. 3° and 4" tergites red, rarely dark-

ened dorsally. Pilosity whitish.

d: Maxillary palpus yellow. Mandible broad-

ly yellow medially. Front leg from the tip of the

coxa onwards yellowish red. Mid femur often

completely yellowish red. Hind basitarsus some-

times completely yellowish brown. In other char-

acters similar to the 2.

Material (107%, 535): Russia: Altaysk Terr.: Lake Te-

letskoye (MOS); Irkutsk Prov.: Dachnaya, Irkutsk

(PET); Chita Prov.: Chita. Kazakhstan: Alma-Ata

(MOS).

Dusona opacoides Hinz, 1985

Description. 2: Body length 7 mm. Temples dis-

tinctiy constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 1.0. Frons scarcely depressed, coriaceous,

finely and densely rugose-punctate, with a fine

median longitudinal carina, with fine but distinct

radial wrinkles, which extend to 0.4 the distance

to the median ocellus. Antennal carina low and

narrow. Face and dorsal half of the clypeus coria-

ceous, densely rugose. Clypeus flat, finely coria-

ceous and with rather dense or disperse puncta-

tion ventrally, its apex truncate medially. Malar

space index 0.4. Genal carina low and rather

straight ventrally, joining the low oral carina at an

acute angle. Genal index 0.4. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 39-40 segments, filiform,

the apical 0.2 slightly attenuate, 2”! segment 2.6x,

shortest segment 1.0x, penultimate segment 1.4x

as long as wide. Pronotum densely rugose lateral-

ly on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum coriaceous, dull, very

densely punctate. Scutellum coriaceous, dull, rath-

er densely punctate, with lateral carinae on the

anterior 0.8. Prescutellar groove smooth medially,

with longitudinal striae laterally. Prepectal carina

low, ventral part merging with the pleural part,

the dorsal 0.2 of the latter replaced by several

wrinkles. Transverse carina obliterated. Meso-

pleurum distinctly coriaceous, rather densely or

densely punctate. Speculum punctate posteriorly,

coriaceous and granulate-strigose anteriorly, rather

dull. Depression in front of the speculum with

fine longitudinal striae or wrinkles, which merge

with the punctation anteriorly. Metapleurum

densely rugose. Pleural carina complete. Areolet

rather small, long-stalked, receiving the 2" recur-

rent vein distinctly distad of the middle. Nervel-

lus slightly inclivous, weakly intercepted. Nervel-

lar index 0.8. Tibial spur index 0.7. Hind femur

4.9x as long as high. Hind tarsal claws short, with

short pectinate teeth basally. Propodeum extend-

ing to 0.4 of the hind coxae, distinctly but rather

narrowly and not deeply depressed, with short

transverse wrinkles in the longitudinal furrow,

irregularly wrinkled beside the furrow, carinae

almost completely obliterated. Propodeal spiracle

rather small, 2.5x as long as wide. Spiracular car-

ina narrow, sometimes indistinct. Spiracular in-

dex 0.8. 1 gastral tergite 3.6x, 2” tergite 1.6x as

long as wide. Petiole with small glymmae at the

anterior edge of narrow lateral areas, smooth in

front ofthe glymmae. Petiolar suture for the greater

part obliterated. 1“ gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum not separated from the 3" gastral tergite, the

latter with distinct black lateral stripes divergent

from the ventrolateral edge. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown medially and apically.

Mandible narrowly reddish brown medially.

Wings hyaline. Stigma medium brown medially.

Front and mid legs from the femur onwards yel-

lowish red, the femora narrowly marked with

brown basally. Hind tibia broadly yellowish brown

medially, narrowly medium brown basally and

apically. Hind tarsus medium brown, basitarsus

for the greater part yellowish brown. 2" gastral

tergite red on the posterior 0.2-0.3. 3 and 4"

tergites red. Pilosity whitish.

d unknown.

Material (322): Russia: Yakutsk A.S.S.R.: Yakutsk

(MOS); Khabarovsk Terr.: Khekhtsir near Khabarovsk

(Hinz/ZSM); Kamchatka: Volcano Uzon (PET).

123

Dusona papator Hinz & Horstmann, spec. nov.

Holotype (2): Russia, Chita Prov., Kozlovo, 17.7.

1975 (PET).

Description. 2%: Body length 7 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.7. Frons slightly de-

pressed, coriaceous, finely and densely rugose-

punctate, without a median longitudinal carina

and without radial wrinkles. Antennal carina low

and narrow. Face coriaceous, finely and densely

rugose. Clypeus small, distinctly rounded, coria-

ceous, finely punctate, with fine irregular wrin-

kles, its apex distinctly convex. Malar space index

1.1. Genal carina low and almost straight ventral-

ly, joining the slightly raised oral carina at an

acute angle. Genal index 0.3. Temple finely coria-

ceous and finely and rather densely rugose-punc-

tate medially. Flagellum with 40 segments, fili-

form, the apical 0.2 slightly attenuate, 2” segment

1.9x, shortest segment 1.0x, penultimate segment

1.2x as long as wide. Pronotum coriaceous and

densely rugose-punctate laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior 0.1. Prescutellar groove smooth.

Prepectal carina low, ventral part merging with

the pleural part, the latter distinctly joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum distinctly coria-

ceous, densely punctate, partly rugose-punctate.

Speculum finely coriaceous and granulate-strigose.

Depression in front of the speculum with short

longitudinal wrinkles, which merge with the punc-

tation anteriorly. Metapleurum coriaceous, finely

and densely rugose-punctate. Pleural carina dis-

tinct. Areolet small, short-stalked, receiving the

2rd recurrent vein distinctly distad of the middle.

Nervellus distinctly reclivous, not intercepted.

Tibial spur index 0.7. Hind femur 5.1x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.2 of the

hind coxae, not depressed, the anterior lateral

areas finely coriaceous, other parts irregularly

wrinkled, with some short longitudinal wrinkles

posteriorly, costulae complete, longitudinal cari-

nae indicated medially and posteriorly. Propode-

al spiracle small, 1,6x as long as wide. Spiracular

carina narrow. Spiracular index 1.1. 1 gastral

tergite 2.4x, 2"d tergite 1.1x as long as wide. Peti-

ole with distinct glammae, which are bordered by

longitudinal carinae dorsally, smooth in front of

124

the glymmae. Petiolar suture distinct. 1* gastral

sternite finely coriaceous anteriorly and medially,

transversely striate posteriorly. Thyridium almost

circular. Thyridial index 0.8. Epipleurum separat-

ed from the 3% gastral tergite, the crease not

marked with black. Epipleural index 0.9. Gaster

distincetly compressed posteriorly. Ovipositor in-

dex 0.4.

Colour: Black. Wings slightly tinged with

brown. Stigma brownish black medially. Front leg

from the apical 0.3 of the femur onwards dark

reddish brown. Mid tibia and tarsus dark brown.

Hind tibia black, slightly tinged with brown me-

dially. 2” gastral tergite dorsally red on the poste-

rior 0.4, the red marks more extended laterally. 3"

and 4" tergites red. 5" tergite narrowly marked

with red anteriorly. Pilosity whitish.

d unknown.

Dusona parva Horstmann, spec. nov.

Holotype (4): Russia, Khabarovsk Terr., Khekh-

tsir, near Khabarovsk, 4.7.1983 (PET).

Description. d: Body length 6mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous, fine-

ly and densely rugose-punctate, with a fine medi-

an longitudinal carina, with short radial wrinkles.

Antennal carina low and narrow. Face and dorsal

0.8 of the clypeus coriaceous, finely and densely

rugose-punctate. Clypeus slightly concave, coria-

ceous and densely punctate ventrally, its apex

slightly convex medially. Malar space index 0.5.

Genal carina low and straight ventrally, joining

the strongly raised oral carina at an acute angle.

Genal index 0.5. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 35 segments, filiform, the apical 0.3 attenu-

ate, 2"! segment 1.6x, shortest segment 0.9x, pe-

nultimate segment 1.0x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.4, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, densely

punctate. Scutellum with lateral carinae on the

anterior 0.7. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the dorsal 0.4 of the latter obliterated.

Transverse carina indicated anteriorly. Mesopleu-

rum distinctly coriaceous, densely punctate. Specu-

lum finely coriaceous, shining. Depression in front

of the speculum with fine and long longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum coriaceous, finely and densely

rugose-punctate. Pleural carina distinct. Areolet

large, short-stalked, receiving the 2" recurrent

vein slightly distad ofthe middle. Nervellus slight-

ly inclivous, weakly intercepted. Nervellar index

0.8. Tibial spur index 0.7. Hind femur 5.0x as long

as high. Hind tarsal claws short, with short pecti-

nate teeth basally. Propodeum extending to 0.4 of

the hind coxae, slightly and broadly depressed,

for the greater part with distinct irregular wrin-

kles, with some longitudinal wrinkles posteriorly,

carinae almost completely obliterated. Propodeal

spiracle rather large, 3.2x as long as wide. Spirac-

ular carina narrow. Spiracular index 0.5. 1“ gastral

tergite 3.4x, 2"X tergite 2.0x as long as wide. Peti-

ole with distinct glymmae, which are bordered by

longitudinal carinae dorsally, smooth in front of

the glymmae. Petiolar suture almost completely

obliterated. 1* gastral sternite for the greater part

smooth, with fine transverse striae posteriorly.

Thyridium large, almost circular. Thyridial index

0.7. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly.

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Mandible

narrowly marked with yellow dorsally and medi-

ally. Tegula dark yellowish brown. Wings hya-

line. Stigma yellowish brown medially. Tip of front

coxa and front trochanter and femur yellowish

red. Front tibia and tarsus yellow. Mid femur

yellowish red on the apical 0.3. Mid tibia and

tarsus yellow. Hind tibia whitish yellow medially,

narrowly black basally, broadly black apically.

2"d gastral tergite red on the posterior half. 3° and

4" tergites red. Pilosity whitish.

? unknown.

Dusona parvicavata Horstmann, spec. nov.

Types. Holotype (S): Russia, Buryatiya, Zaka-

mensk, 30.6.1971 (PET). — Paratype: 13 Russia,

Primorye, Chuguyevsk, 3.6.1985 (Hinz/ZSM).

Description. d: Body length 11 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8-1.0. Frons slightly depressed, coria-

ceous, finely and densely punctate, with a distinct

median longitudinal carina, with distinct radial

wrinkles, which extend to 0.3 the distance to the

median ocellus. Antennal carina slightly raised,

its rim slightly bent upwards, the crescent-shaped

rim with fine transverse striae. Face and dorsal 0.6

of the clypeus coriaceous, densely rugose-punc-

tate. Clypeus flat, finely coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.4. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.2.

Temple finely coriaceous and with very fine dis-

perse punctation medially. Flagellum with 49-50

segments, filiform, the apical 0.3 distinctly attenu-

ate, 2” segment 1.5x, shortest segment 1.0x, pe-

nultimate segment 1.2x as long as wide. Prono-

tum coriaceous and densely rugose-punctate lat-

erally on the dorsal 0.4, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.7. Prescutellar groove with longi-

tudinal striae. Prepectal carina low, ventral part

merging with the pleural part, the latter almost

joining the anterior edge of the mesopleurum, the

upper 0.1-0.2 covered with wrinkles. Transverse

carina complete or obliterated posteriorly. Meso-

pleurum coriaceous, dull, distinctly and densely

punctate. Speculum coriaceous, dull. Depression

in front of the speculum granulate-strigose and

with short longitudinal striae, which merge with a

narrow coriaceous area without punctation ante-

riorly. Metapleurum coriaceous, densely punc-

tate or rugose-punctate. Pleural carina distinct.

Areolet rather large, pointed, receiving the 2"‘

recurrent vein distinctly distad of the middle.

Nervellus vertical, distinctly intercepted. Nervel-

lar index 0.7. Tibial spur index 0.4-0.5. Hind femur

5.4x as long as high. Hind tarsal claws rather

short, with short pectinate teeth basally and medi-

ally. Propodeum extending to 0.3 of the hind cox-

ae, broadly and not deeply depressed, sulcus for

the greater part with rather coarse transverse wrin-

kles or irregular wrinkles, costulae and longitudi-

nal carinae present, but partly covered with wrin-

kles, longitudinal carinae obliterated anteriorly.

Propodeal spiracle large, 2.8x as long as wide.

Spiracular carina narrow. Spiracular index 0.4.

1% gastral tergite 4.1x, 2"@ tergite 2.0x as long as

wide. Petiole with small glymmae at the anterior

edge of narrow lateral areas, smooth in front of

the glymmae. Petiolar suture fine. 1* gastral ster-

nite for the greater part with fine transverse striae

inthehholotype, smooth anteriorly and transversely

striate posteriorly in the paratype. Thyridium al-

most circular. Thyridial index about 2.0. Epipleu-

rum not separated from the 3" gastral tergite, the

latter with fine black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly.

Colour: Black. Maxillary palpus dark brown

125

basally and medially, yellowish brown apically.

Mandible broadly yellow medially. Tegula brown.

Wings hyaline. Stigma brown medially. Front leg

from the tip of the coxa onwards yellow, the fe-

mur yellowish red. Mid femur yellowish red on

the apical 0.2. Mid tibia yellow. Mid tarsus dark

brown. Hind tibia yellow medially, narrowly black

basally, broadly black apically. 2"' gastral tergite

with a dark brown or black posterior edge, dorsal-

ly slightly or not at all marked with red, laterally

broadly marked with red in front of the posterior

edge. 3"! and 4" tergites red, partly marked with

black dorsally. Pilosity whitish.

? unknown.

Dusona peptor Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Primorye, Ussu-

riysk, 24.6.1981 (PET). - Paratype: 1? Russia,

Primorye, Roshchino, on River Iman, 15.6.1966

(Hinz/ ZSM).

Description. ?: Body length 14mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the petiole. Ocellar index 0.8.

Frons slightly depressed, coriaceous and finely

and densely punctate dorsally, finely coriaceous

ventrally, with a fine median longitudinal carina,

with distinct radial wrinkles, which extend to 0.3

the distance to the median ocellus. Antennal cari-

na low and narrow. Face and dorsal 0.6 of the

clypeus coriaceous, densely rugose-punctate.

Clypeus flat, coriaceous and rather densely punc-

tate ventrally, its apex truncate medially. Malar

space index 0.4. Genal carina low and almost

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.25. Temple

finely coriaceous and with fine disperse puncta-

tion medially. Flagellum with 53-55 segments, fil-

iform, the apical 0.2 distinctly attenuate, 2 seg-

ment 1.7x, shortest segment 0.9x, penultimate seg-

ment 1.1x as long as wide. Pronotum coriaceous

and densely rugose-punctate laterally on the dor-

sal half, with longitudinal wrinkles ventrally.

Mesoscutum coriaceous, very densely punctate.

Scutellum coriaceous, densely punctate, with lat-

eral carinae on the anterior 0.6. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter covered with

some wrinkles dorsally, almost joining the anteri-

or edge of the mesopleurum. Transverse carina

obliterated. Mesopleurum distinctly coriaceous,

with strong and rather dense or dense punctation.

126

Speculum finely granulate-strigose. Depression

in front of the speculum with fine longitudinal

striae, which merge with a narrow coriaceous area

without punctation anteriorly. Metapleurum co-

riaceous, finely and densely punctate. Pleural ca-

rina distinct. Areolet large, pointed, receiving the

2"! recurrent vein basad of the middle. Nervellus

slightly inclivous, distinctly intercepted, the up-

per branch bent inwards. Nervellar index 0.7. Tib-

ial spur index 0.7. Hind femur 5.0x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.3 of the

hind coxae, rather broadly and not deeply de-

pressed, the sulcus for the greater part irregularly

wrinkled, with transverse wrinkles posteriorly,

costulae partly present, 'longitudinal carinae al-

most complete, but somewhat indistinct. Propo-

deal spiracle large, 3.4x as long as wide. Spiracu-

lar carina narrow. Spiracular index 0.4-0.5. 1% gas-

tral tergite 3.3x, 2” tergite 1.4x as long as wide.

Petiole with large glymmae, which are bordered

by longitudinal carinae dorsally, with very fine

sculpture in front of the glymmae, with some

transverse wrinkles anteriorly. Petiolar suture in-

distinct or obliterated. 1" gastral sternite for the

greater part smooth, with short transverse striae

posteriorly. Thyridium oval. Thyridial index about

1.0. Epipleurum separated from the 3“ gastral

tergite anteriorly, pendent as a triangular lobe, a

black lateral stripe parallel with the horizontal

bulge in a short distance anteriorly, divergent from

the ventrolateral edge of the tergite posteriorly.

Epipleural index 0.2. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellowish red medially and apically. Man-

dible broadly yellow medially. Wings slightly

tinged with brown. Stigma dark brown medially.

Front trochanter and femur yellowish red, marked

with black ventrally. Front tibia and tarsus yel-

lowish brown. Mid femur vellowish brown on the

apical 0.3. Mid tibia yellowish brown. Mid tarsus

dark brown. Hind tibia yellowish brown medial-

ly, narrowly black basally, broadly black apically.

2"d gastral tergite black dorsally, broadly marked

with red laterally, the posterior edge marked with

black. Tergites and sternites from the 3“! onwards

red, 3’ and 4" tergites narrowly marked with

black dorsally, the posterior tergites more broadly

marked with black dorsally (but for the greater

part red in lateral view). Pilosity whitish.

d unknown.

Dusona perditator Hinz, 1994

Description. ?: Body length 18mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate,

with a fine and partly obliterated median longitu-

dinal carina, with short radial wrinkles. Antennal

carina low and narrow, the rim slightly bent up-

wards. Face and clypeus densely rugose-punctate

on a finely coriaceous background. Clypeus flat,

its apex truncate medially. Malar space index 0.4.

Genal carina low and straight ventrally, joining

the distinctly raised oral carina at an acute angle.

Genal index 0.5. Temple finely coriaceous and

with rather dense or dense punctation medially.

Flagellum with 64 segments, filiform, the apical

half distinctly attenuate, 2” segment 1.6x, short-

est segment 1.0x, penultimate segment 1.4x as

long as wide. Pronotum densely rugose-punctate

laterally on the dorsal 0.2, with longitudinal wrin-

kles ventrally. Mesoscutum and scutellum coria-

ceous, densely rugose-punctate. Scutellum with

lateral carinae on the anterior half. Prescutellar

groove with distinct longitudinal striae. Ventral

part ofthe prepectal carina merging with the trans-

verse carina, the former low, the latter somewhat

raised. Pleural part of the prepectal carina obliter-

ated. Pleural part of the prepectus rugose-punc-

tate. Mesopleurum dispersely or rather densely

punctate on a very finely coriaceous or smooth

background. Speculum finely granulate-strigose

dorsally, smooth ventrally. Depression in front of

the speculum with distinct long longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum finely and densely rugose-punctate,

with some longitudinal wrinkles anteriorly. Pleu-

ral carina complete. Areolet large, narrowly ses-

sile, receiving the 2" recurrent vein almost at the

middle. Nervellus vertical, strongly intercepted.

Nervellar index 0.6. Tibial spur index 0.7. Hind

femur 5.3x as long as high. Hind tarsal claws

rather long, with distinct pectinate teeth almost

over their total length. Propodeum extending to

0.4 of the hind coxae, narrowly and not deeply

depressed, with long transverse wrinkles in al-

most all parts, costulae complete, longitudinal car-

inae present only posteriorly. Propodeal spiracle

large, 2.9x as long as wide. Spiracular carina nar-

row. Spiracular index 0.3. 1% gastral tergite 5.2x,

2" tergite 2.2x as long as wide. Petiole without

glymmae, with narrow lateral areas, smooth in

front of these areas. Petiolar suture fine, partly

obliterated. 1 gastral sternite smooth. Thyridium

oval. Thyridial index about 1.5. Epipleurum sepa-

rated from the 3" gastral tergite anteriorly, pen-

dent as a small triangular lobe, a distinct black

lateral stripe parallel with the ventrolateral edge

in a short distance anteriorly, divergent from the

latter posteriorly. Epipleural index 0.2. Gaster dis-

tinctly compressed posteriorly. Ovipositor index

0.6.

Colour: Black. Mandible broadly yellow me-

dially. Wings hyaline. Stigma dark brown medial-

ly. Front trochanter marked with brown. Front

femur yellowish brown, marked with black ven-

trally. Front tibia and tarsus yellowish brown.

Mid femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. Mid tarsus brown. Hind

tibia yellowish brown, slightly and narrowly dark-

ened basally. Hind tarsus dark brown. 2n“ gastral

tergite dorsally black, laterally marked with red in

front of the black posterior edge. 3° and 4" tergites

red, tinged with brown dorsally. 5" tergite marked

with red laterally. Gastral sternites red or reddish

brown. Pilosity whitish.

d unknown.

Material (12): Russia: Kuril Isl.: Volcano Mendeleye-

va on Kunashir (PET).

Dusona perditor (Förster, 1868)

Description. 2: Body length 12 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons slightly depressed,

coriaceous, finely and densely punctate, with a

fine median longitudinal carina, with indistinct

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.6 of the clypeus coriaceous,

densely rugose-punctate and rugose. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.5.

Genal carina low ventrally, slightly bent sidewards,

joining the slightly raised oral carina almost at a

right angle. Genal index 0.4. Temple finely coria-

ceous and with fine disperse punctation medially.

Flagellum with 47-49 segments, filiform, the api-

cal 0.3 attenuate, 2” segment 1.6x, shortest seg-

ment 1.0x, penultimate segment 1.1x as long as

wide. Pronotum coriaceous and densely rugose-

punctate laterally on the dorsal 0.2, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, densely rugose-punctate. Scutel-

lum with lateral carinae on the anterior 0.4.

Prescutellar groove smooth medially, with longi-

tudinal striae laterally. Ventral part of the prepec-

127

tal carina merging with the transverse carina, the

former low, the latter slightly raised. Pleural part

of the prepectal carina for the greater part obliter-

ated, present as a short wrinkle medially. Pleural

part of the prepectus rugose-punctate and with

wrinkles. Mesopleurum densely punctate on a

finely coriaceous and shining background. Specu-

lum granulate-strigose or with very fine and long

oblique striae, sometimes shining. Depression in

front of the speculum with rather short longitudi-

nal striae or wrinkles, which merge with the punc-

tation anteriorly. Metapleurum densely rugose.

Pleural carina fine, sometimes partly indistinct

and covered with wrinkles. Areolet large, point-

ed, receiving the 2” recurrent vein almost at the

middle. Nervellus slightly inclivous, distinctly

intercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 5.8x as long as high. Hind tarsal

claws rather long, with long pectinate teeth al-

most over their total length. Propodeum extend-

ing to 0.4 of the hind coxae, broadly but not deep-

ly depressed, with long transverse wrinkles in

almost all parts, costulae partly present, longitu-

dinal carinae present posteriorly and sometimes

also medially. Propodeal spiracle small, 2.4x as

long as wide. Spiracular carina obliterated or re-

placed by several wrinkles. Spiracular index 1.0.

1" gastral tergite 4.7x, 2" tergite 2.6x as long as

wide. Petiole without or with small and shallow

glymmae, sometimes with narrow lateral areas,

smooth in front of these glymmae or lateral areas.

Petiolar suture fine, partly obliterated. 1“ gastral

sternite smooth. Thyridium oval. Thyridial index

about 1.5. Epipleurum separated from the 3" gas-

tral tergite anteriorly, folded in or folded back

secondarily and pendent as a narrow rectangular

lobe, a black lateral stripe on the crease or parallel

with it in a short distance anteriorly, divergent

from the ventrolateral edge posteriorly. Epipleu-

ral index 0.2-0.3 (Fig. 40). Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown medially and apically.

Mandible narrowly marked with reddish brown

dorsally. Tegula dark brown. Wings hyaline. Stig-

ma brownish black medially. Front femur yellow-

ish red. Front tibia and tarsus yellow. Mid femur

yellow on the apical 0.3. Mid tibia and tarsus

yellow, the tarsus darkened apically. Hind tibia

broadly yellowish brown medially, narrowly me-

dium brown basally and apically. 2” gastral tergite

dorsally black, laterally red on the posterior 0.6,

sometimes the posterior edge dark brown. 3"! and

4" tergites narrowly black dorsally, broadly red

laterally. 5" tergite laterally marked with red an-

teriorly. Pilosity whitish.

128

d: Sometimes pleural carina and spiracular

carina complete. Mandible marked with brown

medially. Tegula medium brown. Sometimes tip

of the front coxa and front trochanter yellowish

brown. In other characters similar to the 2.

Material (5?%, 13): Russia: Irkutsk Prov.: Irkutsk;

Chita Prov.: Chita; Khabarovsk Terr.: locality unread-

able; Kuril Isl.: Dubovoye on Kunashir (PET).

Dusona petiolatoides Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Russia, Primorye, Kedro-

vaya Pad Reserve, near Khasan, 2.8.1963 (MOS). —

Paratypes: 1? same locality, 12.8.1963 (MOS)

(This paratype is at the same time a paratype of

D. genator.); 12 Russia, Primorye, Nakhodka, 20.

8.1985 (Hinz /ZSM); 13 same locality as the holo-

type, 2.8.1963 (MOS); 15 Russia, Primorye, Vladi-

vostok, 5.8.1984 (Hinz /ZSM).

Description. 2: Body length 10-11 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.8. Frons not depressed,

coriaceous, distinctly rugose-punctate, with short

radial wrinkles, the median longitudinal carina

almost obliterated. Antennal sockets close to each

other. Antennal carina low, its rim slightly bent

upwards. Face and clypeus coriaceous, densely

rugose-punctate. Clypeus flat, its ventral half less

densely punctate medially, its apex truncate me-

dially. Malar space index 0.5. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.2.

Temple finely and rather densely or densely punc-

tate on an almost smooth background medially.

Flagellum with 53-56 segments, filiform, the api-

cal 0.3 distinctly attenuate, 2"? segment 1.9x, short-

est segment 1.0x, penultimate segment 1.4x as

long as wide. Pronotum for the greater part densely

rugose-punctate laterally, the wrinkles smooth,

with longitudinal wrinkles only in the anterior

furrow. Mesoscutum and scutellum finely coria-

ceous, densely punctate. Scutellum with lateral

carinae on the anterior 0.4. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the transverse carina, the latter slightly

raised. Pleural part of the prepectal carina obliter-

ated dorsally and ventrally, the median 0.3 present.

Mesopleurum densely punctate on a finely coria-

ceous and partly smooth background. Speculum

with fine oblique striae. Depression in front of the

speculum with distinct short longitudinal striae,

which merge with the punctation anteriorly. Meta-

pleurum densely rugose. Pleural carina distinct.

Areolet large, narrowly sessile, receiving the 2"4

recurrent vein distad of the middle. Nervellus

slightly reclivous, not intercepted or weakly inter-

cepted. In the latter case nervellar index 0.7. Tibial

spur index 0.7. Hind femur 5.3x as long as high.

Hind tarsal claws rather long, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to the middle of the hind coxae, slightly

depressed, the broad and shallow sulcus for the

greater part irregularly wrinkled, with transverse

wrinkles posteriorly, costulae obliterated, longi-

tudinal carinae indicated medially and posterior-

ly. Propodeal spiracle large, 3.0x as long as wide.

Spiracular carina obliterated. Spiracular index 0.4.

1% gastral tergite 4.5x, 2" tergite 2.2x as long as

wide. Petiole without glymmae, with indistinct

narrow lateral areas (indicated by a coriaceous

surface), smooth in front of these areas. Petiolar

suture partly obliterated. 1“ gastral sternite smooth.

Thyridium oval. Thyridial index about 2.0. Epi-

pleurum not separated from the 3" gastral tergite,

the latter with distinct black lateral stripes well

above the ventrolateral edge. Gaster strongly com-

pressed posteriorly. Ovipositor index 0.5.

Colour: Black. Mandible black, tinged with

dark brown medially. Wings hyaline. Stigma me-

dium brown medially. Front femur yellowish

brown dorsally or completely black. Front tibia

yellowish brown. Front tarsus dark brown. Mid

tibia dark brown or black. 2” gastral tergite red on

the posterior 0.1. 3" tergite red, sometimes marked

with black dorsally. 4" tergite sometimes narrow-

Iy marked with red anteriorly. Pilosity whitish.

d: Mandible narrowly marked with yellow

dorsally and medially. 2" gastral tergite com-

pletely black. 3“ tergite marked with red only

laterally. In other characters similar to the 2.

Dusona petiolator (Fabricius, 1804)

Description. 2: Body length 8-10 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

asthe width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons not depressed, coriaceous,

densely rugose-punctate, without a median longi-

tudinal carina, with short radial wrinkles. Anten-

nal sockets close to each other. Antennal carina

low and narrow. Face and dorsal 0.3 of the clypeus

coriaceous, finely and densely rugose and rugose-

punctate. Clypeus flat, coriaceous and finely and

rather densely punctate ventrally, its apex trun-

cate medially. Malar space index 0.5. Genal carina

straight ventrally, slightly raised, joining the dis-

tinctly raised oral carina at an acute angle. Genal

index 0.25. Temple coriaceous and with fine dense

punctation medially. Flagellum with 42-45 seg-

ments, filiform, the apical 0.2 attenuate, 24 seg-

ment 1.4x, shortest segment 0.8x, penultimate seg-

ment 1.0x as long as wide. Pronotum densely

rugose laterally on the dorsal 0.3, with longitudi-

nal wrinkles ventrally. Mesoscutum coriaceous,

rather dull, densely rugose-punctate. Scutellum

densely rugose-punctate on a finely coriaceous

and shining background, with lateral carinae on

the anterior half. Prescutellar groove with fine

longitudinal striae. Ventral part of the prepectal

carina merging with transverse carina, the former

low, scarcely emarginate medially, the latter slight-

ly raised. Pleural part of the prepectal carina part-

Iy obliterated, the ventral and median part or only

the median 0.3 present (Fig. 20), rarely also the

median part replaced by several wrinkles. Pleural

part of the prepectus rugose-punctate and with

wrinkles. Mesopleurum densely rugose-punctate

and partly rugose on a smooth or finely coria-

ceous and shining background. Speculum with

fine oblique striae. Depression in front ofthe specu-

lum with distinct rather short longitudinal striae,

which merge with the punctation anteriorly. Meta-

pleurum densely rugose. Pleural carina fine. Are-

olet large, pointed, receiving the 2"4 recurrent vein

at or slightly distad ofthe middle. Nervellus slight-

ly inclivous, weakly intercepted. Nervellar index

0.7-0.8. Tibial spur index 0.7. Hind femur 5.3x as

long as high. Hind tarsal claws short, with short

pectinate teeth basally and medially. Propodeum

extending to 0.4 of the hind coxae, distinctly and

rather narrowly depressed, for the greater part

transversely wrinkled, costulae partly present, lon-

gitudinal carinae for the greater part obliterated.

Propodeal spiracle small, 3.3x as long as wide.

Spiracular carina obliterated. Spiracular index 1.0.

1* gastral tergite 4.9x, 2" tergite 1.7x as long as

wide. Petiole without glymmae, with indistinct

narrow lateral areas, smooth in front of these are-

as. Petiolar suture almost completely obliterated.

1% gastral sternite for the greater part smooth,

with fine transverse striae posteriorly. Thyridium

oval. Thyridial index about 3.0. Epipleurum not

separated from the 3" gastral tergite, a black later-

al stripe weakly indicated or lacking. Gaster dis-

tinctly compressed posteriorly. Ovipositor index

1.4.

Colour: Black. Mandible narrowly marked

with yellowish brown medially. Wings hyaline.

Stigma brownish black medially. Front femur yel-

lowish brown dorsally. Front tibia yellowish

129

brown. Front tarsus brown. Mid tibia medium

brown to dark brown. 2” gastral tergite complete-

ly black or red on the posterior 0.1. 3 tergite red.

4!" tergite black or marked with red anteriorly.

Pilosity whitish.

d: Sometimes temples less strongly constrict-

ed, tangential lines on eyes and temples intersect

on the propodeum. Propodeum less strongly de-

pressed, partly with irregular wrinkles. Mandible

rather broadly yellow medially. Sometimes front

coxa and front trochanter marked with yellow. In

other characters similar to the 2.

Material (36?2, 1184): Russia: Novosibirsk Prov.:

Novosibirsk; Altaysk Terr.: Volchikha (MIN); Irkutsk

Prov.: Gora Cherskogo; Chita Prov.: Adrianovka,

Chita, Lake Ivan, Kalga, Klichka, Kurort Darasun,

Urulyunguy E of Borzya; Amur Prov.: Novorossiyka;

Khabarovsk Terr.: Khekhtsir near Khabarovsk; Kam-

chatka: Kamaki; Kuril isl: Golovnino on Kunashir,

Malokurilsk on Shikotan (PET). Kirgiziya: Alayskiy

Mts. (PAR), Issykata (PET). The species is known also

from Sakhalin/Russia and Hokkaido/Japan (Townes

et al. 1965: 289).

Variety: 2"d gastral tergite red on the posterior

0.3. 3° and 4 tergites red. Material: 1? Kirgiziya,

Alayskiy Mts., Nookatsky, Kirghiz-Ata River,

2150-2500 m (LIN). This specimen may represent

an individual variation.

Dusona petitor Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Chita Prov., Olov-

yannaya, 9.7.1971 (PET). — Paratype: 1? Russia,

Yakutsk, 12.-13.7.1927 (Hinz / ZSM).

Description. ?: Body length 8-9 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.6. Frons scarcely depressed, coriaceous,

densely rugose-punctate and rugose, with a fine

median longitudinal carina, with short and indis-

tinct radial wrinkles. Antennal carina low and

narrow. Face and dorsal 0.6 of the clypeus densely

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely or dispersely punctate ventrally, its

apex slightly convex medially. Malar space index

0.3. Genal carina low ventrally, slightly bent side-

wards and forwards, joining the oral carina at the

base of the mandible at a right angle. Temple

finely coriaceous and with fine disperse puncta-

tion medially. Flagellum with 42-44 segments, fil-

iform, the apical 0.3 attenuate, 2” segment 1.9x,

130

shortest segment 1.0x, penultimate segment 1.3x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.2, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, very densely punctate. Scutellum with

lateral carinae on the anterior 0.7. Prescutellar

groove with fine longitudinal striae. Ventral part

of the prepectal carina merging with the trans-

verse carinae, both carinae raised, the former dis-

tinctliy emarginate medially. Pleural part of the

prepectal carina lower, slightly raised ventrally,

low medially, the dorsal 0.3 obliterated. Meso-

pleurum distinctly coriaceous, rather densely or

densely punctate. Speculum finely coriaceous and

granulate-strigose. Depression in front of the

speculum with distinct löngitudinal striae, which

merge with the punctation anteriorly. Metapleu-

rum coriaceous, densely rugose-punctate. Pleural

carina distinct. Areolet large, short-stalked, re-

ceiving the 2" recurrent vein slightly distad of the

middle. Nervellus distinctly reclivous, not inter-

cepted or weakly intercepted. In the latter case

nervellar index 0.6-0.7. Tibial spur index 0.6. Hind

femur 4.9x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally. Propo-

deum extending to the middle of the hind coxae,

broadly and distinctly but not deeply depressed,

sulcus almost completely transversely wrinkled,

costulae and longitudinal carinae partly indicated

or almost completely obliterated. Propodeal spir-

acle large, 3.5x as long as wide. Spiracular carina

partly widened. Spiracular index 0.3. 1“ gastral

tergite 4.2x, 2" tergite 2.0x as long as wide. Peti-

ole with small glymmae at the anterior edge of

indistinct narrow lateral areas, smooth in front of

the glymmae. Petiolar suture for the greater part

obliterated. 1% gastral sternite smooth. Thyridium

oval. Thyridial index about 1.5. Epipleurum not

separated from the 3"! gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellow medially and apically. Mandible

narrowly marked with yellowish red dorsally and

medially. Tegula yellowish brown with a dark

brown mark. Wings hyaline. Stigma medium

brown medially. Front leg from the femur on-

wards yellowish red. Mid femur yellowish red on

the apical 0.3. Mid tibia yellowish red. Mid tarsus

dark Hind tibia yellowish brown medially, narrow-

ly black basally, broadly black apically. 2" gastral

tergite red on the posterior 0.3. 3% and 4" tergites

red. Pilosity whitish.

d unknown.

Dusona pictator Hinz & Horstmann, spec. nov.

This is a variable and poorly defined species.

Types. Holotype (2): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarowsk, 13.6.1985 (PET). —

Paratypes: 1? Russia, Primorye, Vladivostok, 6.-

10.6.1912 (Hinz/ZSM); 1? Japan, Hokkaido, Sap-

poro, Maruyama, 20.6.1958 (SAP); 13 Russia,

Kuril Isl., Kunashir, Golovnino, 10.6.1973 (PET).

Description. 2: Body length 8-10 mm. Temples

distinctly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.9. Frons slightly depressed, coriaceous,

finely and densely rugose-punctate, with short

radial wrinkles, the median longitudinal carina

fine and partly obliterated. Antennal carina low

and narrow. Face and clypeus coriaceous, finely

and very densely punctate. Clypeus flat, its apex

truncate medially, the rim thick and smooth. Ma-

lar space index 0.3. Genal carina low and straight

ventrally, joining the distinctly raised oral carina

at an acute angle. Genal index 0.2-0.3. Temple

coriaceous and with fine disperse punctation me-

dially. Flagellum with 41-44 segments, filiform,

the apical 0.2 distinctly attenuate, 2” segment

1.9x, shortest segment 0.9x, penultimate segment

1.2x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.4, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, rather densely or densely punc-

tate. Scutellum with lateral carinae on the anterior

0.3. Prescutellar groove smooth or with few longi-

tudinal striae. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina obliterated. Mesopleurum fine-

ly coriaceous, rather densely or densely punctate.

Speculum finely granulate-strigose, shining. De-

pression in front of the speculum with distinct

longitudinal striae dorsally and medially, the lat-

ter merging with the punctation anteriorly, finely

granulate-strigose and shining ventrally. Meta-

pleurum coriaceous and densely punctate (in the

holotype) or with very dense punctation on an

almost smooth background (in the 2? paratypes).

Pleural carina distinct. Areolet rather small, point-

ed, receiving the 2” recurrent vein at or distad of

the middle. Nervellus slightly inclivous, intercept-

ed. Nervellar index 0.7-0.8. Tibial spur index 0.7.

Hind femur 4.7x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to 0.3 of the hind

coxae, scarcely depressed, irregularly wrinkled

anteriorly and medially, with some longitudinal

wrinkles posteriorly, carinae almost completely

obliterated. Propodeal spiracle rather large, 2.5x

as long as wide. Spiracular carina indistinct. Spirac-

ular index 0.3. 1% gastral tergite 2.9x, 2” tergite

1.4x as long as wide. Petiole with distinct glym-

mae, which are bordered by longitudinal carinae

dorsally and ventrally, smooth in front of the glym-

mae. Petiolar suture complete. 1% gastral sternite

for the greater part smooth, with some transverse

striae posteriorly. Thyridium oval. Thyridial in-

dex about 2.0. Epipleurum separated from the 3'«

gastral tergite, the crease marked with black. Epi-

pleural index 0.7. Gaster not strongly compressed

posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown.

Mandible sometimes tinged with brown medial-

ly. Wings hyaline. Stigma light brown medially.

Front femur yellowish red, sometimes darkened

ventrally. Front tibia and tarsus yellow. Mid fe-

mur sometimes yellowish brown on the apical 0.2.

Mid tibia yellowish brown or brown. Hind tibia

blackish (in the 2? paratypes) or tinged with yel-

lowish brown medially (in the holotype). 2" gas-

tral tergite completely black dorsally or marked

with red on the posterior 0.1, sometimes with

broad red marks laterally. The following tergites

red laterally, marked with black dorsally (to a

varying extent), sometimes the 3" or 4" tergite

completely red, the 6" tergite half red in lateral

view. Gastral sternites red. Pilosity whitish.

d: Body length 8 mm. Flagellum with 44 seg-

ments. Metapleurum with very dense punctation

on a smooth background. Propodeum with the

longitudinal carinae present medially and poste-

riorly. Tegula yellowish brown. Front and mid

tibiae and tarsi yellow. Hind tibia dark yellowish

brown medially, narrowly black basally, broadly

black apically. Genital clasper reddish brown. In

other characters similar to the ?. The association

of the only known d with the 22 is not certain.

Variety: Body length 7-9 mm. Flagellum with

40-43 segments. Propodeum sometimes with trans-

verse wrinkles posteriorly. Mandible marked with

yellow medially. Tegula yellowish. 2" gastral

tergite dorsally red on the posterior 0.2-0.3, the

red marks more extended laterally. The following

tergites for the greater part red, narrowly marked

with black dorsally. Material: 2?? Russia, Primo-

rye, Posyet (12 PET, 1? Hinz/ZSM); 1? Russia,

Primorye, Khasan (PET). These ?? may or may

not belong to D. pictator.

131

Dusona pineticola sibirica Hinz, 1985

Description. 2: Body length 11-14 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutallar groove. Ocel-

lar index 0.5. Frons distinctly depressed, coria-

ceous, finely and densely rugose-punctate dorsal-

ly, with short radial wrinkles in the depressions,

with a rather low median longitudinal bulge and

a median longitudinal carina on its top. Antennal

carina distinctly raised, with a strong transversely

striate projection dorsally (nose-like) (Fig. 7). Face

and dorsal 0.3 of the clypeus coriaceous, densely

rugose-punctate. Clypeus slightly concave, coria-

ceous and rather densely punctate ventrally, its

apex slightly convex medially. Malar space index

0.5. Genal carina low and straight ventrally, join-

ing the distinctly raised oral carina at an acute

angle. Genal index 0.7. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

ellum with 46-51 segments, filiform, the apical 0.3

distinctly attenuate, 2"! segment 1.7x, shortest seg-

ment 1.0x, penultimate segment 1.8x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

very densely punctate. Scutellum with lateral car-

inae on the anterior 0.8. Prescutellar groove with

fine longitudinal striae. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

almost joining the anterior edge of the mesopleu-

rum, the dorsal 0.1 replaced by several wrinkles.

Transverse carina obliterated. Mesopleurum dense-

ly punctate or rugose-punctate on a very finely

coriaceous and shining background. Speculum

with fine oblique striae. Depression in front of the

speculum with distinct rather short longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum densely rugose. Pleural carina

for the greater part obliterated and replaced by

wrinkles, rarely complete. Areolet large, sessile,

pointed or short-stalked, receiving the 2” recur-

rent vein distinctly basad of the middle. Nervellus

inclivous, distinctly intercepted. Nervellar index

0.8. Tibial spur index 0.8. Hind femur 5.7x as long

as high. Hind tarsal claws short, with rather long

pectinate teeth basally and medially. Propodeum

extending to 0.4 of the hind coxae, broadly and

not deeply depressed, with short transverse wrin-

kles anteriorly and long transverse wrinkles pos-

teriorly, costulae partly present, longitudinal car-

inae obliterated anteriorly and medially, present

posteriorly. Propodeal spiracle rather small, 3.1x

as long as wide. Spiracular carina indistinct or

obliterated. Spiracular index 0.5. 1“ gastral tergite

132

5.1x, 2"d tergite 1.9x as long as wide. Petiole with-

out glymmae or with very small gliymmae at the

anterior edge of indistinct narrow lateral areas,

almost smooth in front of these areas, with some

very fine longitudinal striae. Petiolar suture part-

ly obliterated. 1 gastral sternite for the greater

part smooth, with fine transverse striae posterior-

ly. Thyridium elongate. Thyridial index about 1.0.

Epipleurum not separated from the 3’ gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.4.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellow dorsally

and medially. Wings hyaline. Stigma dark brown

medially. Front femur and tibia yellowish brown.

Front tarsus dark brown. Sometimes mid femur

and mid tibia partly tinged with brown. 2" gas-

tral tergite red on the posterior 0.1. 3" tergite red.

4" tergite completely red or red on the anterior

0.8.

d not available for study.

Material (1822, 438): Russia: Irkutsk Prov.: Irkutsk;

Chita Prov.: Adrianovka, Chita (PET), Lake Ivan

(Hinz/ZSM, PET), Klichka (PET). Kazakhstan: Alma-

Ata. Tadzhikistan: Pamir Mts., 2800 m (MOS). Kirgiz-

iya: Alayskiy Mts., 3000-3100 m (LIN, PAR), Terskey

Alatau Mts. (MOS). Mongoliya: Hentiy Prov.: Tsen-

hermandal (PET).

Dusona polita (Förster, 1868)

Description. 2: Body length 10-12 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons slightly depressed, coriaceous, finely

and rather densely punctate, with a fine and part-

ly obliterated median longitudinal carina, with

fine and short radial wrinkles. Antennal carina

low and narrow. Face and dorsal half ofthe clypeus

coriaceous, very densely punctate. Clypeus flat,

coriaceous and densely punctate ventrally, its apex

truncate medially. Malar space index 0.2. Genal

carina low ventrally, slightly bent sidewards, join-

ing the distinctly raised oral carina at an acute

angle. Genal index 0.4. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 42-46 segments, filiform, the api-

cal 0.3 attenuate, 2” segment 1.3x, shortest seg-

ment 0.7x, penultimate segment 1.0x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

very densely punctate. Scutellum with lateral car-

inae on the anterior 0.2. Prescutellar groove smooth

medially, with fine longitudinal striae laterally.

Prepectal carina low, ventral part merging with

the pleural part, the dorsal 0.3 of the latter obliter-

ated or replaced by wrinkles, sometimes almost

complete. Transverse carina for the greater part

obliterated. Mesopleurum distinctly and densely

punctate on a very finely coriaceous and shining

background. Speculum very finely coriaceous,

partly smooth, strongly shining. Depression in

front of the speculum dorsally and medially with

fine longitudinal wrinkles, which merge with the

punctation anteriorly, the ventral 0.3 finely coria-

ceous, shining. Metapleurum with fine and dis-

perse or rather dense punctation on a finely coria-

ceous and shining background. Pleural carina dis-

tinct. Areolet large, short-stalked, receiving the

2" recurrent vein basad of the middle. Nervellus

almost vertical, distinctly intercepted. Nervellar

index 0.7-0.8. Tibial spur index 0.7. Hind femur

4.8x as long as high. Hind tarsal claws rather long,

with distinct pectinate teeth basally and medially.

Propodeum extending to 0.3 of the hind coxae,

broadly and rather deeply depressed, sulcus for

the greater part transversely wrinkled, with a short

median longitudinal keel posteriorly, costulae

obliterated, longitudinal carinae present posteri-

orly and sometimes also medially, obliterated an-

teriorly. Propodeal spiracle rather large, 2.8x as

long as wide. Spiracular carina narrow, some-

times replaced by several wrinkles. Spiracular in-

dex 0.5. 1* gastral tergite 3.4x, 2" tergite 1.6x as

long as wide. Petiole with large glymmae, which

are bordered by longitudinal carinae dorsally,

smooth in front of the glymmae. Petiolar suture

partly obliterated. 1” gastral sternite for the great-

er part smooth, with fine transverse striae posteri-

orly. Thyridium elongate. Thyridial index about

1.0. Epipleurum separated from the 34 gastral

tergite, the crease marked with black. Epipleural

index 0.6-0.7. Gaster distinctly compressed poste-

riorly, strongly shining. Ovipositor index 0.7-0.8.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow

dorsally and medially. Tegula dark brown. Wings

hyaline. Stigma yellowish brown medially. Front

trochanter marked with yellow. Front femur, tibia

and tarsus yellowish red. Mid femur yellowish

red on the apical 0.4-0.7. Mid tibia yellowish red.

Mid tarsus dark brown. Hind tibia broadly yel-

lowish brown medially, narrowly dark brown

basally and apically. Hind tarsus dark brown.

2"d gastral tergite red on the posterior 0.4-0.5. 3"4

tergite red. 4" tergite red, marked with black pos-

teriorly (to a variable extend). Pilosity whitish.

d: Depression in front of the speculum finely

coriaceous and shining on the ventral 0.5-0.6. Pro-

podeum rather narrowly and not deeply de-

pressed, longitudinal carinae complete or almost

complete. Epipleural index 0.7-0.8. Maxillary pal-

pus yellow. Mandible broadly yellow medially.

Tergula yellow. Front and mid legs from the tip of

the coxa onwards yellow, the femora yellowish

red. Hind femur red on the apical 0.2. Hind tibia

broadly yellow medially, narrowly medium brown

basally and apically. In other characters similar to

the 9.

Material (1°): Russia: Primorye: Pogranichnyy (PET).

Dusona propodeator Hinz, 1994

Description. %: Body length 9 mm. Temples

strongly constricted behind the eyes, 0.5x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.5. Frons distinctly but not

deeply depressed above the antennal carinae, co-

riaceous, finely and densely rugose-punctate, with

a very fine median longitudinal carina, without

radial wrinkles. Antennal carina slightly but dis-

tinctly raised, the rim bent upwards and at the

same time somewhat widened to a smooth bulge.

Face and dorsal 0.6 of the clypeus densely rugose-

punctate and rugose. Clypeus flat, finely coria-

ceous and rather densely punctate ventrally, its

apex truncate or slightly concave medially. Malar

space index 0.2. Genal carina low and straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.3. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 40-42 segments, filiform,

the apical 0.1 attenuate, 2”! segment 2.7x, shortest

segment 1.1x, penultimate segment 1.2x as long

as wide. Pronotum very densely punctate on a

smooth background laterally on the dorsal 0.2,

with distinct longitudinal striae on a smooth back-

ground ventrally. Mesoscutum coriaceous, dull,

very densely punctate. Scutellum densely punc-

tate on a smooth background, with lateral carinae

on the anterior 0.7. Prescutellar groove for the

greater part smooth. Prepectal carina low, ventral

part merging with the pleural part, the dorsal 0.3

of the latter obliterated. Transverse carina com-

plete, low. Mesopleurum with rather dense or

dense punctation on a slightly coriaceous and

somewhat shining background. Speculum finely

granulate-strigose. Depression in front of the

speculum with distinct long longitudinal striae,

which merge with the punctation anteriorly. Meta-

133

pleurum densely rugose-punctate. Pleural carina

distinct. Areolet rather small, stalked, receiving

the 2"d recurrent vein slightly distad of the mid-

dle. Nervellus slightly reclivous, not intercepted

or weakly intercepted. In the latter case nervellar

index 0.7. Tibial spur index 0.5. Hind femur 5.5x

as long as high. Hind tarsal claws short, with short

pectinate teeth basally. Propodeum extending to

0.7 of the hind coxae, shallowly and narrowly

depressed, the longitudinal furrow with short

transverse wrinkles, irregularly wrinkled beside

the furrow, costulae complete, longitudinal cari-

nae present anteriorly (as short stubs) and poste-

riorly, obliterated medially. Propodeal spiracle

rather large, 2.8x as long as wide. Spiracular cari-

na widened. Spiracular index 0.3. 1" gastral tergite

4.7x, 2"d tergite 2.7x as long as wide. Petiole with-

out glymmae, with narrow lateral areas, smooth

in front of these areas. Petiolar suture distinct.

1" gastral sternite for the greater part smooth, with

transverse striae posteriorly. Thyridium elongate.

Thyridial index about 1.0. Epipleurum not sepa-

rated from the 3°“ gastral tergite, the latter with

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with reddish

brown medially. Wings hyaline. Stigma dark

brown medially. Front femur yellowish brown on

the apical half. Front tibia and tarsus yellowish

brown. Mid tibia and tarsus dark brown, the tibia

narrowly marked with yellow basally and apical-

ly. 3’ and 4" gastral tergites broadly black dorsal-

ly, marked with red ventrolaterally (half red or

less in lateral view). Pilosity whitish.

Material (27??): Japan: Hokkaido: Atsusabe (SAP);

Honshu: Mt. Odaigahara near Nara (Hinz/ZSM).

?d: Flagellum with 47 segments, 2” segment 2.5x,

shortest segment 1.3x, penultimate segment 1.4x

as long as wide. Metapleurum coriaceous, finely

and densely punctate. Areolet receiving the 2"

recurrent vein basad of the middle. Propodeum

extending to the middle of the hind coxae. Mandi-

ble broadly reddish brown medially, Front leg

from the tip of the coxa onwards yellowish. Mid

femur yellow on the apical 0.2. Mid tibia and

tarsus yellow. Hind tibia tinged with yellowish

brown subbasally on the dorsal side. 2" gastral

tergite red on the posterior 0.1. 3° and 4" tergites

red, narrowly marked with black dorsally. In oth-

er characters similar to the 9. Material: 13 Russia,

Khabarovsk Terr., Khekhtsir near Khabarovsk

(PET). This d may belong to D. propodeator.

Dusona pseudobucculenta Hinz & Horstmann,

spec. nov.

Holotype (?): Russia, Chita Prov., Urulyunguy,

E of Borzya, 5.7.1975 (PET).

Description. 2: Body length 11mm. Temples

slightly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the petiole. Ocellar index 1.1.

Frons not depressed, coriaceous, densely rugose-

punctate, with a very fine median longitudinal

carina, with indistinct radial wrinkles. Antennal

sockets very close to each other. Antennal carina

low and narrow. Face and clypeus densely ru-

gose-punctate. Clypeus flat, its apex slightly con-

vex medially, the rim thick and smooth. Malar

space index 0.5. Genal carina low ventrally, slight-

ly bent sidewards, joining the distinctly raised

oral carina at a right angle. Genal index 0.5: Tem-

ple with fine and rather dense punctation on an

almost smooth background medially. Flagellum

with 43 segments, filiform, the apical 0.3 distinctly

attenuate, 2” segment 1.4x, shortest segment 1.0x,

penultimate segment 1.6x as long as wide. Prono-

tum very densely punctate on a smooth back-

ground laterally, without wrinkles. Mesoscutum

and scutellum very densely punctate on a smooth

background. Scutellum with lateral carinae on the

anterior 0.4. Prescutellar groove with fine longitu-

dinal striae. Prepectal carina low, ventral part

merging with the pleural part, the dorsal 0.4 of the

latter obliterated. Pleural part of the prepectus

distinctly punctate without wrinkles. Transverse

carina strongly raised anteriorly, obliterated pos-

teriorly. Mesopleurum with rather dense or dense

punctation on a smooth background. Speculum

punctate posteriorly, with fine oblique striae ante-

riorly. Depression in front of the speculum with

distinct short longitudinal wrinkles, which merge

with the punctation anteriorly. Metapleurum very

densely punctate on a smooth background. Pleu-

ral carina distinct. Areolet large, narrowly sessile,

receiving the 2"! recurrent vein at the middle.

Nervellus slightly inclivous, weakly intercepted.

Nervellar index 0.8. Tibial spur index 0.6. Hind

femur 4.9x as long as high. Hind tarsal claws

rather long, with some distinct pectinate teeth

basally. Propodeum extending to 0.3 of the hind

coxae, scarcely depressed, irregularly wrinkled,

with a short median longitudinal keel medially,

costulae partly present, longitudinal carinae al-

most completely obliterated. Propodeal spiracle

small, 3.5x as long as wide. Spiracular carina ob-

literated and covered with wrinkles. Spiracular

index 0.9. 1% gastral tergite 3.8x, 2"! tergite 1.5x as

long as wide. Petiole with distinct glymmae, which

are bordered by longitudinal carinae dorsally,

smooth in front of the glymmae. Petiolar suture

obliterated. 1“ gastral sternite smooth. Thyridium

oval. Thyridial index about 1.0. Epipleurum not

separated from the 3 gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index about 0.7.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with yellowish red

dorsally and medially. Wings hyaline. Stigma dark

brown medially. Front coxa, trochanter and fe-

mur black, marked with yellow anteriorly. Front

tibia yellowish brown. Front tarsus dark brown.

2"d gastral tergite red on the posterior 0.3. 3° and

4" tergites red, 4" tergite narrowly marked with

black dorsally and posteriorly. Pilosity whitish.

d unknown.

Dusona pugillator (Linnaeus, 1758)

Dusona amictor Hinz, 1979 (syn. nov.)

Dusona scutellatoides Hinz, 1985 (syn. nov.)

The holotype (?) of Dusona amictor Hinz from

Hakone/Honshu/Japan (OTT) cannot be found

now. The specimen was obviously characterized

by an unusually large propodeal spiracle. The

synonymy is already indicated in the original de-

scription (Hinz 1979: 220). The holotype (?) of

Dusona scutellatoides Hinz from Londoko/ Yevreys-

kiy A.O./Russia (PET) has recently been studied

by the junior author.

Description. 2: Body length 14-17 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons slightly but dis-

tinctly depressed above the antennal carinae, co-

riaceous, finely and densely rugose-punctate, with

a fine median longitudinal carina, with distinct

radial wrinkles, which extend to half the distance

to the median ocellus. Antennal carina slightly

but distinctly raised, the rim bent upwards (ear-

like) (this structure being variable und sometimes

indistinct). Face and dorsal 0.6 of the clypeus co-

riaceous, rather coarsely rugose. Clypeus flat, fine-

ly coriaceous and rather densely punctate ventral-

ly, its apex truncate or slightly concave medially.

Malar space index 0.3. Genal carina rather straight

ventrally, slightly raised, joining the distinctly

raised oral carina at an acute angle. Genal index

0.5. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with 52-58

segments, filiform, the apical 0.3 distinctly attenu-

ate, 2"! segment 1.5x, shortest segment 1.0x, pe-

nultimate segment 1.3x as long as wide. Prono-

tum finely and densely rugose laterally on the

dorsal 0.2, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior 0.3. Prescutellar groove smooth

medially, with fine longitudinal striae laterally.

Prepectal carina low, ventral part merging with

the pleural part or with the transverse carina,

pleural part complete, joining the anterior edge of

the mesopleurum, transverse carina complete or

obliterated posteriorly. Mesopleurum distinctly

coriaceous, distinctly and rather densely or densely

punctate. Speculum coriaceous, rather dull. De-

pression in front of the speculum dorsally with

rather short longitudinal striae, which merge with

the punctation anteriorly, the ventral 0.3-0.5 coria-

ceous. Metapleurum coriaceous, densely rugose.

Pleural carina distinct. Areolet large, short-stalked,

receiving the 2” recurrent vein basad of the mid-

dle. Nervellus slightly inclivous, strongly inter-

cepted. Nervellar index 0.7. Tibial spur index 0.4.

Hind femur 5.5x as long as high. Hind tarsal claws

short, with long pectinate teeth basally and medi-

ally. Propodeum extending to 0.3 of the hind cox-

ae, broadly and deeply depressed, sulcus for the

greater part with long transverse wrinkles, costu-

lae partly present, longitudinal carinae strong

medially and posteriorly, almost complete. Pro-

podeal spiracle large, 3.9x as long as wide. Spirac-

ular carina indistinct or obliterated. Spiracular in-

dex 0.4. 1% gastral tergite 4.8x, 2” tergite 2.0x as

long as wide. Petiole with large glymmae, which

are bordered by longitudinal carinae dorsally, with

very fine sculpture or smooth in front of the glym-

mae. Petiolar suture for the greater part obliterat-

ed. 1‘ gastral sternite coriaceous anteriorly, smooth

medially, transversely striate posteriorly. Thyrid-

ium oval. Thyridial index about 2.0. Epipleurum

separated from the 3 gastral tergite anteriorly,

the crease marked with black. Epipleural index

0.2-0.4. Gaster strongly compressed posteriorly.

Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellow medially. Tegu-

la yellowish brown. Wings hyaline or slightly in-

fuscate. Stigma yellowish brown medially. Front

leg from the tip of the coxa onwards yellowish

red, the tibia yellow. Mid femur yellow apically

(to a variable extend). Mid tibia yellow. Mid tar-

sus dark brown. Hind femur completely black or

yellowish red on the apical 0.2. Hind tibia broadly

yellow medially, narrowly brown or black basally

155

and apically. 2" gastral tergite dorsally red on the

posterior 0.2-0.5, the red marks more extended

laterally, sometimes the tergite completely red

laterally. 3° and 4" tergites red, 4" tergite some-

times marked with black posteriorly. 5" tergite

sometimes marked with red anteriorly. Pilosity

whitish.

d: Longitudinal carinae of propodeum some-

times complete. Front and mid legs almost com-

pletely yellowish red, the coxae marked with black

basally. In other characters similar to the 2.

Material (10722, 635): Russia: Novosibirsk Prov.:

Cherepanovo (MOS); Chita Prov.: Chita, Karymskoye,

Kurort Darasun; Amur Prov.: Novorossiyka; Yevreys-

kiy A.O.: Londoko; Khabarovsk Terr.: Khekhtsir near

Khabarovsk (PET); Primorye: Spassk Dalniy (MOS);

Kamchatka: Kozyrevsk, Milkovo, Ust-Kamchatsk.

Kazakhstan: Kokchetav (PET). Japan: Hokkaido: Ma-

ruyama (SAP).

Dusona pulmentariae Hinz 1963

Description. ?: Body length 6-9 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 0.9. Frons scarcely depressed, coriaceous,

densely rugose-punctate, with a very fine and

incomplete median longitudinal carina or the lat-

ter obliterated, radial wrinkles scarcely indicated.

Antennal carina low and narrow. Face and dorsal

0.8 of the clypeus coriaceous and densely rugose-

punctate. Clypeus flat, finely coriaceous and dense-

ly punctate ventrally, its apex slightly convex

medially. Malar space index 0.5. Genal carina low

ventrally, slightly bent sidewards, joining the dis-

tinctly raised oral carina at a right angle. Genal

index 0.2. Temple finely coriaceous and with fine

disperse punctation medially. Flagellum with 36-40

segments, filiform, the apical 0.2 slightly attenu-

ate, 2" segment 1.5x, shortest segment 0.9x, pen-

ultimate segment 1.1x as long as wide. Pronotum

very densely punctate laterally on the dorsal 0.2,

with longitudinal wrinkles ventrally. Mesoscu-

tum coriaceous, dull, very densely punctate. Scutel-

lum very densely punctate on a partly smooth

background, with lateral carinae on the anterior

0.6. Prescutellar groove smooth medially, with

longitudinal striae laterally. Prepectal carina low,

ventral part merging with the pleural part, the

latter complete or almost complete, sometimes

the dorsal 0.2 replaced by several wrinkles. Trans-

136

verse carina almost completely obliterated. Meso-

pleurum finely coriaceous, distinctly and densely

punctate. Speculum finely granulate-strigose, shin-

ing. Depression in front of the speculum with

distinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum distinctly and

coarsely rugose. Pleural carina complete. Areolet

rather small, pointed or short-stalked, receiving

the 2” recurrent vein distinctly distad of the mid-

dle. Nervellus almost vertical, rather weakly in-

tercepted. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 5.0x as long as high. Hind tarsal

claws rather long, with distinct pectinate teeth

basally and medially. Propodeum extending to

the middle of the hind coxae, deeply but rather

narrowly depressed, with strong transverse striae

in the longitudinal furrow, coarsely and irregular-

ly wrinkled beside the furrow, carinae complete

but partly indistinct and covered with wrinkles.

Propodeal spiracle small, 3.0x as long as wide.

Spiracular carina narrow. Spiracular index 0.6.

1* gastral tergite 4.5x, 2” tergite 2.0x as long as

wide. Petiole without glymmae, with distinct nar-

row lateral areas, with very fine sculpture or

smooth in front of these areas. Petiolar suture

partly obliterated. 1" gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum not separated from the 3" gastral tergite, the

latter without black lateral stripes. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus dark brown

basally and medially, yellowish brown apically.

Mandible narrowly marked with reddish brown

mediaily. Flagellum light brown apically. Tegula

yellow. Wings hyaline. Stigma yellowish brown

medially. Front femur yellowish red, slightly dark-

ened basally. Front tibia and tarsus yellow. Mid

femur yellowish red on the apical 0.4. Mid tibia

yellow. Mid tarsus yellow basally, black apically.

Hind tibia yellow basally and medially, the apical

0.3 black. Hind tarsus blackish, the basitarsus nar-

rowly yellow basally. 2" gastral tergite dorsally

red on the posterior 0.2-0.3, red marks more ex-

tended laterally. 3° tergite red, sometimes marked

with black posteriorly. 4" tergite narrowly marked

with red anteriorly or completely black. Pilosity

whitish.

d: Similar to the 9.

Material (3?2, 638): Russia: Chita Prov.: Nerchin-

skiy Zavod, Priargunsk; Primorye: Khasan, Novoka-

chalinsk (PET). Kirgiziya: Kashkasi in Kirgizskiy Mts.,

1800 m (LIN). China: Heilongjiang Prov.: Charbin

(= Harbin) (AEI).

Dusona radiator Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarovsk, 12.6.1983 (PET). -

Paratypes: 1? same locality, 12.6.1985 (PET);

1? same locality, 14.6.1985 (Hinz/ ZSM).

Description. 2: Body length 7 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons scarcely depressed, coriaceous, densely

rugose, with a fine median longitudinal carina,

with short and indistinct radial wrinkles. Anten-

nal carina low and narrow. Face and dorsal 0.7 of

the clypeus densely rugose-punctate. Clypeus flat,

coriaceous and with fine disperse punctation ven-

trally, its apex scarcely convex medially, the rim

thick and smooth. Malar space index 0.4. Genal

carina low ventrally, distinctly bent sidewards

and slightly bent forwards, joining the oral carina

close to the base of the mandible at a right angle.

Genal index 0.1 or less. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 36-37 segments, filiform, the api-

cal 0.2 slightly attenuate, 2" segment 2.3x, short-

est segment 1.0x, penultimate segment 1.0x as

long as wide. Pronotum with dense irregular wrin-

kles laterally on the dorsal 0.1, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous and densely punctate. Scutellum with

lateral carinae on the anterior 0.7. Prescutellar

groove with distinct short longitudinal striae. Pre-

pectal carina low, ventral part merging with the

pleural part, the dorsal 0.2 of the latter obliterated

and covered with wrinkles. Transverse carina ob-

literated. Mesopleurum distinctly coriaceous, rath-

er densely or densely punctate. Speculum with

fine oblique striae, shining. Depression in front of

the speculum with distinct long longitudinal stri-

ae, which merge with the punctation anteriorly.

Metapleurum coriaceous, finely and densely ru-

gose. Pleural carina distinct. Areolet rather small,

pointed or narrowly sessile, receiving the 2" re-

current vein at the middle. Nervellus vertical,

weakly intercepted. Nervellar index 0.8. Tibial

spur index 0.7. Hind femur 4.8x as long as high.

Hind tarsal claws short, with pectinate teeth ba-

sally. Propodeum extending to 0.4 of the hind

coxae, slightly and narrowly depressed, with few

short transverse wrinkles in the longitudinal fur-

row, with some longitudinal wrinkles posteriorly,

in other parts irregularly wrinkled, carinae almost

completely obliterated. Propodeal spiracle small,

2.0x as long as wide. Spiracular carina narrow.

Spiracular index 0.8. 1 gastral tergite 3.9x, 2"«

tergite 1.7 as long as wide. Petiole smooth later-

ally, without glymmae. Petiolar suture partly ob-

literated. 1" gastral sternite for the greater part

smooth, with fine transverse striae posteriorly.

Thyridium elongate. Thyridial index about 0.5.

Epipleurum not separated from the 3% gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly but not strongly compressed posteriorly. Ovi-

positor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible narrowly marked with yellowish red dorsal-

ly and medially. Tegula yellowish brown. Wings

hyaline. Stigma yellowish brown medially. Front

trochanter yellowish red dorsally, black ventrally.

Front femur yellowish red. Front tibia and tarsus

yellow. Mid femur yellowish red on the apical 0.4.

Mid tibia and tarsus yellow. Hind tibia yellowish

brown medially, narrowly marked with dark

brown basally, broadly marked with dark brown

apically. Hind tarsus dark brown. 2" gastralttergite

red on the posterior 0.1. 3" and 4" tergites red,

both marked with dark brown to a variable ex-

tent. Pilosity whitish.

d unknown.

Dusona recta (Thomson, 1887)

Description. 2: Body length 8-10 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slighly depressed,

coriaceous, finely and rather densely punctate,

with a very fine median longitudinal carina, with

distinct short radial wrinkles, which extend to 0.3

the distance to the median ocellus. Antennal cari-

na narrow, slightly raised (inconspicuous). Face

and dorsal half of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and dis-

persely or rather densely punctate ventrally, its

apex truncate medially. Malar space index 0.3.

Genal carina straight ventrally, slightly raised,

joining the slightly raised oral carina at an acute

angle. Genal index 1.5. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 42-45 segments, filiform, the api-

cal 0.2 attenuate, 2" segment 2.0x, shortest seg-

ment 0.8x, penultimate segment 1.2x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

dull, densely or very densely punctate. Scutellum

187

with lateral carinae on the anterior 0.3. Prescutel-

lar groove smooth medially, with longitudinal

striae laterally. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina present anteriorly. Mesopleu-

rum distinctly coriaceous, rather finely and rather

densely punctate. Speculum coriaceous, rather

dull. Depression in front of the speculum dorsally

with fine longitudinal striae, which merge with

the punctation anteriorly, the ventral 0.6 coria-

ceous. Metapleurum coriaceous, finely and densely

punctate, partly with fine irregular wrinkles. Pleu-

ral carina distinct. Areolet rather small, long-

stalked, receiving the 2" recurrent vein distad of

the middle. Nervellus vertical, weakly intercept-

ed. Nervellar index 0.8. Tibial spur index 0.5. Hind

femur 4.9x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to 0.3 of the hind

coxae, broadly and deeply depressed, sulcus co-

riaceous or with few short transverse wrinkles

anteriorly, with rather long transverse wrinkles

medially, with transverse wrinkles over the total

breadth of the sulcus posteriorly, carinae for the

greater part obliterated or covered with wrinkles,

costulae sometimes present. Propodeal spiracle

rather large, 2.7x as long as wide. Spiracular cari-

na narrow or widened. Spiracular index 0.5.

1% gastral tergite 4.1x, 2"4 tergite 1.8x as long as

wide. Petiole with distincet but rather small and

often elongate glymmae, which are bordered by

longitudinal carinae dorsally, smooth in front of

the glymmae. Petiolar suture almost completely

obliterated. 1* gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium oval. Thyridial index about 1.5. Epipleu-

rum separated from the 3"! gastral tergite anteri-

orly and folded in, a black lateral stripe parallel

with the crease in a short distance anteriorly, di-

vergent from the ventrolateral edge posteriorly.

Epipleural index 0.2-0.3. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly marked with yellow dorsally

and medially. Tegula yellowish brown. Wings

hyaline. Stigma yellowish brown medially. Front

trochanter marked with yellow. Front femur yel-

lowish red. Front tibia and tarsus yellow. Mid

femur yellow on the apical 0.3. Mid tibia yellow.

Mid tarsus brown. Hind tibia yellow medially,

narrowly black basally, broadly black apically.

2"! gastral tergite red on the posterior 0.4. 3° and

4" tergites red. Pilosity whitish.

d: Propodeum with complete costulae, longi-

tudinal carinae present medially and posteriorly.

138

Front coxa yellow apically and front trochanter

yellow. Mid femur almost completely yellowish

red. In other characters similar to the ®.

Material (6??, 258): Russia: Primorye: Kanganz,

Partizansk, Pozhiga SO of Dalnerechensk, Vladivos-

tok; Kuril Isl.: Dubovoye and Volcano Golovina on

Kunashir (PET).

Dusona rectator Hinz & Horstmann, spec. nov.

Holotype (?): Russia, Primorye, Sudzukhinskiy

Reserve, 180 km E of Vladivostok, 6.7.1946 (MOS).

Description. ?: Body length 11mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed,

coriaceous, very finely punctate, with a distinct

narrow median longitudinal carina, with short

and indistinct radial wrinkles. Antennal carina

low and narrow, the rim inconspicuously bent

upwards. Face and clypeus coriaceous, densely

rugose-punctate. Clypeus flat, its apex almost trun-

cate medially. Malar space index 0.3. Genal carina

low and straight ventrally, joining the slightly but

distinctly raised oral carina at an acute angle. Ge-

nal index 0.5. Temple finely coriaceous and with

fine disperse punctation medially. Flagellum fili-

form, its apical 0.3 broken off in the only known

specimen, 2"! segment 1.7x, shortest segment 1.0x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, densely punctate. Scutellum with lat-

eral carinae on the anterior half. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the latter almost joining

the anterior edge of the mesopleurum. Transverse

carina present anteriorly. Mesopleurum distinctly

coriaceous, densely punctate. Speculum coria-

ceous. Depression in front of the speculum for the

greater part coriaceous, with fine and short longi-

tudinal striae medially, which merge with a coria-

ceous area without punctation anteriorly. Meta-

pleurum distinctly coriaceous, densely punctate.

Pleural carina strong. Areolet large, short-stalked,

receiving the 2" recurrent vein slightly distad of

the middle. Nervellus slightly reclivous, distinct-

ly intercepted. Nervellar index 0.7. Tibial spur

index 0.6. Hind femur 5.1x as long as high. Hind

tarsal claws short, with pectinate teeth basally.

Propodeum extending to 0.3 of the hind coxae,

broadly and rather deeply depressed, sulcus with

fine transverse wrinkles anteriorly and posterior-

ly, more irregularly wrinkled medially, carinae

almost completely obliterated. Propodeal spiracle

large, 3.3x as long as wide. Spiracular carina nar-

row. Spiracular index 0.2. 1” gastral tergite 3.9x,

2" tergite 2.0x as long as wide. Petiole with small

glymmae at the anterior edge of rather broad lat-

eral areas, with very fine longitudinal wrinkles in

front of the glymmae, partly smooth. Petiolar su-

ture partly obliterated. 1“ gastral sternite smooth.

Thyridium oval. Thyridial index about 1.5. Epi-

pleurum separated from the 3" gastral tergite an-

teriorly, the crease marked with black. Epipleural

index 0.3. Gaster distinctly compressed posterior-

ly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly marked with yellowish red

medially. Tegula medium brown. Wings hyaline.

Stigma yellowish brown medially. Tip of the front

coxa and front trochanter and femur yellowish

red, the trochanter and femur somewhat dark-

ened ventrally. Front tibia and tarsus yellow. Mid

femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. Mid tarsus dark brown.

Hind tibia yellowish brown medially, narrowly

black basally, very broadly black apically. 2" gas-

tral tergite dorsally red on the posterior 0.2, al-

most completely red laterally. 3° and 4" tergites

red. Pilosity whitish.

d unknown.

Dusona rectoides Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Japan, Honshu, Norikura-

dake, 2000 m, 30.7.1954 (AEI). - Paratypen: 1?

Japan, Honshu, Chuzenji-ko, 4000-5000 ft., July-

August 1913 (Hinz/ZSM); 1? Japan, Honshu,

Kamikochi, near Norikura-dake, 5.8.1916 (SAP);

12 Japan, Shikoku, Ishizuchi, 23.7. (SAP); 538

Japan, Honshu, Kamikochi, near Norikura-dake,

24.-26.7.1954 (483 AEI, 138 Hinz/ZSM).

Description. ?: Body length 6-9 mm. Temples

distinctly constricted behind the eyes, 0.6%x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 1.0. Frons slightly depressed, coriaceous,

with fine and disperse irregular wrinkles, with a

fine median longitudinal carina, with short radial

wrinkles. Antennal carina low and narrow. Face

and dorsal 0.6 of the clypeus coriaceous, finely

and densely rugose-punctate. Clypeus flat, coria-

ceous and with fine disperse punctation ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low and straight ventrally, joining

the scarcely raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 39-41 segments, filiform, the apical 0.2

slightly attenuate, 2" segment 2.0x, shortest seg-

ment 0.9x, penultimate segment 1.1x as long as

wide. Pronotum coriaceous and with fine punc-

tures and irregular wrinkles laterally on the dor-

sal 0.4, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, finely and

densely punctate. Scutellum with lateral carinae

on the anterior half. Prescutellar groove smooth

or with fine longitudinal striae. Prepectal carina

low, ventral part merging with the pleural part,

the latter almost joining the anterior edge of the

mesopleurum (covered with wrinkles close to the

edge). Transverse carina obliterated. Mesopleu-

rum distinctly coriaceous, finely and rather densely

punctate. Speculum coriaceous. Depression in

front of the speculum with rather short longitudi-

nal striae and wrinkles, which merge with a nar-

row coriaceous area without punctation anterior-

ly. Metapleurum coriaceous, very finely and dense-

ly rugose-punctate. Pleural carina distinct. Areolet

rather large, pointed, receiving the 2" recurrent

vein at or slightly distad of the middle. Nervellus

vertical, distinctly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 5.2x as long as

high. Hind tarsal claws short, with distinct pecti-

nate teeth basally. Propodeum extending to 0.4 of

the hind coxae, rather broadly and not deeply

depressed, sulcus for the greater part irregularly

wrinkled, with few transverse wrinkles, with some

short longitudinal wrinkles posteriorly, costulae

partly present, longitudinal carinae almost com-

plete, but partly indistinct. Propodeal spiracle rath-

er large, 2.9x as long as wide. Spiracular carina

narrow. Spiracular index 0.4. 1" gastral tergite

4.3x, 2"d tergite 1.9x as long as wide. Petiole with

long and very narrow glymmae, which are bor-

dered by carinae dorsally and ventrally, smooth

in front of the glymmae. Petiolar suture very fine.

1* gastral sternite for the greater part smooth,

with fine transverse striae posteriorly. Thyridium

oval. Thyridial index about 1.0. Epipleurum sepa-

rated from the 3" gastral tergite anteriorly, either

folded in or folded back secondarily and pendent

as a narrow lobe, the crease marked with black, a

black lateral stripe parallel with the ventrolateral

edge in a short distance anteriorly, divergent from

the latter posteriorly. Epipleural index 0.2-0.3.

Gaster distinctly but not strongly compressed pos-

teriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus medium

brown. Mandible broadiy marked with yellow

139

medially. Tegula dark brown. Wings hyaline. Stig-

ma dark brown medially. Front femur yellowish

red dorsally, dark brown ventrally. Front tibia

and tarsus yellowish red. Mid tibia yellowish red.

Mid tarsus dark brown. Hind tibia black, some-

times tinged with dark brown subasally. 2" gas-

tral tergite red on the posterior 0.2. 3" tergite red.

4" tergite varying from almost completely red to

almost completely black. Pilosity whitish.

d: Flagellum with 37-38 segments. Maxillary

palpus yellow. Front leg from the tip of the coxa

onwards yellowish, the coxa sometimes completely

black. Mid leg from the trochanter onwards yel-

lowish, the trochanter and femur sometimes

marked with brown. Hind tibia dark yellowish

brown medially, narrowly black basally, broadly

black apically. In other characters similar to the 2.

Dusona remota (Förster, 1868)

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8-0.9. Frons slightly depressed, coria-

ceous, finely and rather densely punctate, with a

fine median longitudinal carina, with fine radial

wrinkles, which extend to 0.3 the distance to the

median ocellus. Antennal carina low and narrow.

Face and dorsal 0.6 of the clypeus coriaceous,

densely rugose-punctate and rugose. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low ventrally, slightly bent sidewards,

joining the slightly raised oral carina at an acute

angle. Genal index 0.5. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 39-41 segments, filiform, the api-

cal 0.2 slightly attenuate, 2” segment 1.9x, short-

est segment 0.9-1.0x, penultimate segment 1.0-1.1x

as long as wide. Pronotum densely rugose-punc-

tate and rugose laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, densely punctate.

Scutellum with lateral carinae on the anterior half.

Prescutellar groove almost smooth medially, with

fine longitudinal striae laterally. Prepectal carina

low, ventral part merging with the pleural part,

the dorsal 0.2 of the latter replaced by several

wrinkles. Transverse carina present, low. Meso-

pleurum distinctly coriaceus, rather densely or

densely punctate. Speculum granulate-strigose,

rather dull, sometimes a small ventral area almost

smooth and shining. Depression in front of the

speculum with fine longitudinal striae, which

140

merge with the punctation anteriorly. Metapleu-

rum densely rugose. Pleural carina distinct. Areo-

let rather small, stalked, receiving the 2” recur-

rent vein distad of the middle. Nervellus slightly

inclivous, weakly intercepted. Nervellar index 0.8.

Tibial spur index 0.7. Hind femur 5.5x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to 0.4 of the

hind coxae, broadly and distinctly but not deeply

depressed, sulcus with short or long transverse

wrinkles anteriorly and medially, with long trans-

verse wrinkles posteriorly, often with a median

longitudinal keel posteriorly, costulae partly

present, longitudinal carinae present but some-

times fine and indistinct medially and posteriorly,

obliterated anteriorly. Propodeal spiracle rather

small, 2.3x as long as wide. Spiracular carina nar-

row. Spiracular index 0.6. 1% gastral tergite 4.4x,

2"! tergite 1.8x as long as wide. Petiole with dis-

tinct glymmae, which are bordered by longitudi-

nal carinae dorsally, smooth in front of the glym-

mae. Petiolar suture fine, sometimes partly oblit-

erated. 1“ gastral sternite coriaceous anteriorly,

smooth or with fine sculpture medially, trans-

versely striate posteriorly. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3“ gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow

dorsally and medially, rarely completely black.

Tegula dark brown or black. Wings hyaline. Stig-

ma light brown medially. Front leg from the fe-

mur onwards yellowish red, the femur sometimes

marked with brown basally. Mid femur often

marked with yellowish red on the apical 0.1-0.2.

Mid tibia yellowish red. Mid tarsus light brown.

Hind tibia broadly yellow to yellowish red medi-

ally, narrowly black basally and apically. Hind

tarsus dark brown, at most narrowly marked with

yellowish brown basally. 2”! gastral tergite dor-

sally red on the posterior 0.2-0.4, the red marks

more extended laterally, sometimes the posterior

edge slightly marked with brown. 3’ and 4"

tergites red. Pilosity whitish.

d: Propodeum more coarsely transversely

wrinkled, with strong longitudinal carinae medi-

ally and posteriorly. Front trochanter sometimes

yellow. In other characters similar to the 9.

Material (222, 238): Russia: Sverdlovsk Prov.: Yeka-

terinburg (= Sverdlovsk) (PET); Kemerovo Prov.: Tya-

zhinskiy (MOS); Kuril Isl.: Urup (SAP). Japan: Hon-

shu: Kamikochi near Norikura-dake (AEI), Nagano (OTT).

Dusona rubator Hinz, 1985

Description. 9: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons not depressed, densely rugose-punc-

tate, with a fine median longitudinal carina, with

indistinct radial wrinkles. Antennal carina low

and narrow. Face and clypeus densely rugose.

Clypeus slightly concave, with a narrow smooth

area above its apex, the latter slightly convex me-

dially. Malar space index 0.4. Genal carina low

ventrally, slightly bent sidewards, joining the

slightly raised oral carina almost at a right angle.

Genal index 0.4. Temple finely coriaceous, partly

almost smooth, with distinct rather dense puncta-

tion medially. Flagellum with 39-40 segments,

slightly fusiform, 2” segment 1.4x, shortest seg-

ment 0.7x, penultimate segment 0.9x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.4, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum very densely

punctate on a smooth background. Scutellum with

lateral carinae on the anterior 0.6. Prescutellar

groove smooth medially, with fine longitudinal

striae laterally. Prepectal carina low, ventral merg-

ing with the pleural part, the latter almost com-

plete or complete, in the latter case joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum densely punc-

tate on a smooth background. Speculum with fine

oblique striae. Depression in front of the specu-

lum with distinct longitudinal striae, which merge

with the punctation anteriorly. Metapleurum very

densely punctate on a smooth background. Pleu-

ral carina complete. Areolet rather large, pointed,

receiving the 2" recurrent vein slightly basad of

the middle. Nervellus distinctly reclivous, weakly

intercepted. Nervellar index 0.6. Tibial spur index

0.5. Hind femur 3.7x as long as high. Hind tarsal

claws rather long, with short pectinate teeth ba-

sally. Propodeum extending to 0.6 of the hind

coxae, distinctly and rather narrowly depressed,

with distinct short transverse striae in the longitu-

dinal furrow, irregularly wrinkled beside the fur-

row, carinae almost completely obliterated. Pro-

podeal spiracle rather small, 3.5x as long as wide.

Spiracular carina narrow. Spiracular index 0.6.

1% gastral tergite 3.5x, 2” tergite 1.4x as long as

wide. Petiole with small glymmae at the anterior

edge of narrow lateral areas, smooth in from of

the glymmae. Petiolar suture partly obliterated.

1* gastral sternite for the greater part smooth,

with transverse striae posteriorly. Thyridium oval.

Thyridial index about 1.0. Epipleurum not sepa-

rated from the 3" gastral tergite, the latter with

fine black lateral stripes well above the ventrola-

teral edge. Gaster slightly compressed posterior-

ly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellowish red medially. Flagel-

lum light brown apically. Tegula yellow. Wings

hyaline. Stigma yellowish brown medially. Front

and mid legs from the tip ofthe coxa onwards red.

Hind leg from the trochanter onwards red. Post-

petiole and 2" to 4" tergites red, sometimes the 4"

tergite indistinctly tinged with black dorsally. Pi-

losity whitish.

d unknown.

Material (2??): Russia: Buryatiya: Verkhneudinsk (=

Ulan-Ude) (PET). Mongoliya: Selenge Prov.: Darhan

(Hinz /ZSM).

Dusona rufator Hinz, 1994

Description. ?: Body length 8 mm. Temples dis-

tinctly constricted behind the eyes, 0.6x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.9. Frons not depressed, coriaceous,

finely rugose, with a fine median longitudinal

carina, without radial wrinkles. Antennal carina

low and narrow. Face and dorsal 0.7 of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

finely coriaceous and densely punctate ventrally,

its apex truncate medially. Malar space index 0.4.

Genal carina low ventrally, slightly bent forwards,

joining the slightly raised oral carina at an acute

angle. Genal index 0.2. Temple coriaceous and

with fine dense punctation medially. Flagellum

with at least 40 segments (both tips broken off in

the only available ®), filiform, the apical 0.2 slight-

ly attenuate, 2"? segment 2.0x, shortest segment

1.0x as long as wide. Pronotum with longitudinal

wrinkles laterally, with additional punctures dor-

solaterally. Mesoscutum and scutellum coriaceous,

densely rugose-punctate. Scutellum with lateral

carinae on the anterior 0.2. Prescutellar groove

smooth medially, with some longitudinal striae

laterally. Ventral part ofthe prepectal carina merg-

ing with the transverse carina, both carinae low.

Pleural part of the prepectal carina present only

ventrally as a short stub. Mesopleurum densely

punctate on a finely coriaceous and shining back-

ground. Speculum finely granulate-strigose. De-

pression in front of the speculum with fine but

distinct long longitudinal striae, which merge with

the punctation anteriorly. Metapleurum finely and

densely rugose-punctate and rugose. Pleural car-

141

ina complete. Areolet rather large, short-stalked,

receiving the 2" recurrent vein distinctly distad of

the middle. Nervellus slightly reclivous, weakly

intercepted. Nervellar index 0.7. Tibial spur index

0.5. Hind femur 5.4x as long as high. Hind tarsal

claws short, with short pectinate teeth. Propo-

deum extending to the middle of the hind coxae,

shallowly and narrowly depressed, with short

transverse wrinkles in the longitudinal furrow,

irregularly wrinkled beside the furrow, carinae

almost completely obliterated. Propodeal spiracle

rather small, 2.5x as long as wide. Spiracular car-

ina narrow. Spiracular index 0.5. 1” gastral tergite

4.6x, 2” tergite 2.5% as long as wide. Petiole with-

out glymmae, with narrow lateral areas, smooth

in front of these areas. Petiolar suture fine. 1

gastral sternite coriaceous anteriorly, smooth me-

dially, transversely striate posteriorly. Thyridium

elongate. Thyridial index about 1.0. Epipleurum

not separated from the 3" gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Wings hyaline. Stig-

ma light brown medially. Front leg from the tip of

the coxa onwards yellowish red. Mid leg from the

trochanter onwards yellowish red. Hind femur

red. Hind tibia broadly yellowish brown medial-

ly, narrowly brown basally and apically. Hind

tarsus brown, the basitarsus yellowish ventrally.

2nd gastral tergite dorsally red on the posterior 0.2,

laterally red on the posterior 0.4. 3" to 7" tergites

and sternites red, the tergites narrowly marked

with black dorsally. Pilosity whitish.

Material (1?): Japan: Hokkaido: Jozankei (AEI).

?d: Body length 7 mm. Genal carina low and

straight ventrally. Flagellum with 44 segments.

Speculum finely coriaceous, shining. Depression

in front of the speculum partly with irregular

wrinkles. Longitudinal furrow of the propodeum

for the greater part irregularly wrinkled, with some

longitudinal wrinkles posteriorly. Spiracular cari-

na indistinct. Spiracular index 1.0. Thyridium cir-

cular. Thyridial index about 2.5. Tegula yellow.

Front and mid legs from the tip of the coxa on-

wards yellow, the femora yellowish red. Genital

clasper red. In other characters similar to the 2.

Material: 18 Russia, Khabarovsk Terr., Khekhtsir

near Khabarovsk (PET). The association of this

with the 2 is tentative.

142

Dusona rufiventrator Horstmann, spec. nov.

Types. Holotype (?): Japan, Honshu, Nagano,

400 m, 20.5.1962 (OTT). — Paratypes: 15 same lo-

cality, 13.5.1962 (OTT); 13 same locality, 3.5.1960

(Hinz/ZSM).

Description. 2: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.9. Frons slightly depressed, coriaceous,

finely and densely rugose-punctate, with a very

fine median longitudinal carina, with short radial

wrinkles, the latter extending to 0.3 the distance to

the median ocellus. Antennal carina low and nar-

row. Face and dorsal 0.6 of the clypeus coriaceous,

densely rugose-punctate and rugose. Clypeus flat,

finely coriaceous and with fine disperse puncta-

tion ventrally, its apex truncate medially. Malar

space index 0.3. Genal carina straight ventrally,

strongly raised close to the junction with the also

raised oral carina, forming a lopsided triangular

pyramid, the tip of the latter shifted ventrally and

situated behind the base of the mandible, gena

deeply depressed close to the junction, the de-

pression about 0.7x as deep as the width of the 2"“

flagellar segment. Genal index 0.4-0.5. Temple fine-

ly coriaceous and with very fine disperse puncta-

tion medially. Flagellum with 39-41 segments,

slightly fusiform, 2”! segment 1.6x, shortest seg-

ment 0.8x, penultimate segment 1.0x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove smooth.

Prepectal carina low, ventral part merging with

transverse carina, the latter slightly raised. Pleural

part of the prepectal carina indistinct and covered

with wrinkles ventrally, distinct medially, the

dorsal 0.4 obliterated. Mesopleurum finely but

distinctly wrinkled, with distinct and dense punc-

tation. Speculum very finely coriaceous, shining.

Depression in front of the speculum with distinct

longitudinal striae, which merge with the puncta-

tion anteriorly. Metapleurum finely but distinctly

coriaceous, densely punctate. Pleural carina dis-

tinct. Areolet rather small, stalked, receiving the

2"d recurrent vein at the middle. Nervellus dis-

tinctly inclivous, intercepted. Nervellar index

0.8-0.9. Tibial spur index 0.7. Hind femur 5.4x as

long as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.4 of the hind coxae, broadly and

not deeply depressed, for the greater part with

irregular wrinkles, with some transverse wrinkles

posteriorly, carinae almost completely obliterat-

ed. Propodeal spiracle rather large, 3.2x as long as

wide. Spiracular carina partly widened. Spiracu-

lar index 0.5. 1 gastral tergite 4.0x, 2" tergite 1.4x

as long as wide. Petiole without glymmae, lateral

areas indicated, smooth in front of these areas.

Petiolar suture for the greater part obliterated.

1“ gastral sternite for the greater part smooth, with

transverse striae posteriorly. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.8.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly marked with yellow dorsally.

Tegula yellow. Wings hyaline. Stigma medium

brown medially. Front femur yellowish red,

marked with brown ventrally. Front tibia and tar-

sus yellow. Mid femur yellowsh red on the apical

0.2. Mid tibia and tarsus yellow. Hind tibia yellow

medially, narrowly black basally, broadly black

apically. 2" gastral tergite black dorsally, broadly

red laterally. The following tergites narrowly black

dorsally, broadly red laterally. Gastral sterniies

red. Pilosity whitish.

d: Speculum granulate-strigose. Propodeum

for the greater part rather strongly irregularly

wrinkied. Maxillary palpus yellow. Mandible

broadly marked with yellow medially. Tip of the

front coxa and front trochanter yellow. Mid femur

completely yellowish red or marked with yellow-

ish red dorsally and apically. Genital clasper red,

marked with dark brown. In other characters sim-

ilar to the 9.

Dusona rufoapicalis Horstmann, spec. nov.

Holotype (?): China, Shaanxi Prov., Qinling

Mts., Xunyangba, 1000-1300 m, 23.5.-13.6.1998

(LIN).

Description. 2: Body length 10 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.8. Frons slightly de-

pressed, coriaceous, very finely and dispersely

punctate, with few short radial wrinkles, the me-

dian longitudinal carina very fine and partly ob-

literated. Antennal carina low and narrow. Face

and dorsal 0.6 of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

and almost straight ventrally, joining the slightly

raised oral carina at an acute angle. Genal index

0.4. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 51 seg-

ments, filiform, the apical 0.3 attenuate, 2” seg-

ment 1.8x, shortest segment 1.0x, penultimate seg-

ment 1.2x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.3, the

interspaces smooth, with longitudinal wrinkles

ventrally. Mesoscutum coriaceous, very densely

punctate. Scutellum coriaceous, rather densely

punctate, with lateral carinae on the anterior 0.4.

Prescutellar groove smooth. Prepectal carina low,

ventral part merging with the pleural part, the

dorsal 0.2 of the latter indistinct and covered with

wrinkles. Transverse carina present anteriorly.

Mesopleurum distinctly coriaceous, rather finely

and densely punctate, slightly shining. Speculum

coriaceous and granulate-strigose. Depression in

front of the speculum with short longitudinal stri-

ae and wrinkles, which partly merge with the

punctation, partly with an area without puncta-

tion anteriorly. Metapleurum coriaceous, dull,

densely and rather finely punctate. Pleural carina

distinct. Areolet large, short-stalked, receiving the

2°‘ recurrent vein slightly distad of the middle.

Nervellus slightly inclivous, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 5.1x as long as high. Hind tarsal claws

short, with long pectinate teeth basally and medi-

ally. Propodeum extending to 0.3 of the hind cox-

ae, broadly and shallowly depressed, sulcus co-

riaceous and with some irregular wrinkles anteri-

orly, with long transverse wrinkles posteriorly,

carinae almost completely obliterated. Propodeal

spiracle large, 2.8x as long as wide. Spiracular

carina for the greater part narrow. Spiracular in-

dex 0.4. 1% gastral tergite 3.5x, 2"X tergite 1.3x as

long as wide. Petiole with large glymmae, which

are bordered by longitudinal carinae dorsally,

smooth in front of the glymmae. Petiolar suture

for the greater part obliterated. 1* gastral sternite

smooth. Thyridium oval. Thyridial index about

1.0. Epipleurum not separated from the 3“ gastral

tergite, but the ventrolateral edge somewhat bent

inwards and rounded on the anterior 0.2, the tergite

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible almost completely yellow. Tegula yellow.

Wings hyaline. Stigma dark brown medially. Front

leg from the tip of the coxa onwards yellow, the

femur yellowish red. Mid leg from the apical 0.2

of the femur onwards yellow. Hind tibia light

143

yellowish brown medially, narrowly black basal-

ly, broadly black apically. 2" gastral tergite dor-

sally red on the posterior 0.2, laterally red on the

posterior 0.6, the posterior edge narrowly marked

with brown. The following tergites red, the 3'“

dorsally marked with dark brown anteriorly, the

posterior tergites tinged with brown dorsally.

Gastral sternites red. Pilosity whitish.

d unknown.

Dusona rufovariata Horstmann, spec. nov.

Holotype (53): Russia, Primorye, Vladivostok,

7.-20.6.1912 (PET).

Description. d: Body length 10 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.6. Frons slightly de-

pressed, coriaceous, finely and dispersely punc-

tate, with a distinct median longitudinal carina,

with distinct short radial wrinkles. Antennal cari-

na narrow and rather low. Face and dorsal 0.6 of

the clypeus coriaceous, very densely punctate.

Clypeus partly concave, coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.4.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 50 segments,

filiform, the apical 0.3 distinctly attenuate, 2"d seg-

ment 1.8x, shortest segment 0.9x, penultimate seg-

ment 1.4x as long as wide. Pronotum densely

rugose-punctate laterally on the dorsal 0.1, with

longitudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, densely punctate. Scutel-

lum with distinct lateral carinae on the anterior

0.8. Prescutellar groove with fine longitudinal stri-

ae. Prepectal carina low, ventral part merging with

the pleural part, the latter complete, joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum distinctly coria-

ceous, with rather fine and disperse or rather dense

punctation. Speculum coriaceous, rather dull.

Depression in front of the speculum granulate-

strigose and with some longitudinal striae, which

merge with a coriaceous area without punctation

anteriorly. Metapleurum coriaceous, finely and

rather densely punctate. Pleural carina distinct.

Areolet large, short-stalked, receiving the 2" re-

current vein at the middle. Nervellus vertical,

weakly intercepted. Nervellar index 0.7. Tibial

spur index 0.6. Hind femur 4.9x as long as high.

144

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to 0.3 of the hind

coxae, distinctly, rather broadly and not deeply

depressed, costulae and longitudinal carinae com-

plete, area superomedia coriaceous, area petiola-

ris coriaceous and with fine irregular wrinkles,

shining, with some short transverse wrinkles pos-

teriorly. Propodeal spiracle rather large, 3.1x as

long as wide. Spiracular carina narrow. Spiracular

index 0.5. 1% gastral tergite 3.4x, 2" tergite 1.7% as

long as wide. Petiole with distinct and deep but

not large glymmae, smooth in front of the glym-

mae. Petiolar suture fine and complete. 1* gastral

sternite for the greater part smooth, with trans-

verse wrinkles posteriorly. Thyridium almost cir-

cular. Thyridial index about 1.5. Epipleurum sep-

arated from the 3" gastral tergite by a bulge ante-

riorly, pendent as a very narrow rectangular lobe,

the bulge marked with black. Epipleural index

0.4. Gaster distinctly compressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly marked with yellow medially. Tegu-

la yellow. Wings hyaline. Stigma dark yellowish

brown medially. Front and mid legs from the tip

of the coxa onwards yellow, the femora yellowish

red. Hind tibia whitish yellow medially, narrowly

black basally, rather broadly black apically. Hind

tarsus brownish black. Postpetiole marked with

red laterally. 2" gasteral tergite dorsally red on

the posterior 0.2, almost completely red laterally.

3 and 4" tergites for the greater part red, 3"

tergite dorsally marked with black posteriorly, 4"

tergite dorsally marked with black over its total

length. Pilosity whitish.

? unknown.

Dusona rugifer (Förster, 1868)

This is a poorly defined species. Possibly the males

from Europe described below (including the types

of Campoplex rugifer Förster and C. punctus Kriech-

baumer) (ZSM) belong to D. stragifex (Förster) as a

variety with the legs marked with yellow instead

of yellowish brown, and the female from Iran

represents a different, undescribed species (with a

rather low epipleural index). A similar female

from Augsburg/ Germany (Hinz/ZSM) is includ-

ed in the description.

Description. 2: Body length 12-14 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8-0.9. Frons slightly depressed, coria-

ceous and densely rugose-punctate dorsally, with

a fine median longitudinal carina and with dis-

tinct radial wrinkles, which extend to half the

distance to the median ocellus. Antennal carina

low and narrow. Face and dorsal 0.7 ofthe clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and rather densely punctate ventrally,

its apex truncate medially. Malar space index 0.3.

Genal carina low and straight ventrally, joining

the distinctly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous with

very fine and disperse punctation medially. Flag-

ellum with 55-56 segments, filiform, the apical 0.2

attenuate, 2” segment 1.6x, shortest segment 0.9x,

penultimate segment 1.2x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely or very

densely punctate. Scutellum with lateral carinae

on the anterior 0.6. Prescutellar groove for the

greater part smooth, with some longitudinal stri-

ae laterally. Prepectal carina slightly raised ven-

trally, ventral part merging with the pleural part,

the latter almost complete, the dorsal 0.1-0.2 re-

placed by several wrinkles. Transverse carina also

slightly raised, as high as the prepectal carina at

the junction. Mesopleurum distinctly coriaceous,

densely punctate. Speculum coriaceous. Depres-

sion in front of the speculum with fine long longi-

tudinal striae, which merge with the punctation

anteriorly. Metapleurum coriaceous, densely ru-

gose-punctate. Pleural carina distinct. Areolet

large, short-stalked, receiving the 2” recurrent

vein basad of the middle. Nervellus slightly in-

clivous, distinctly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 5.1x as long as

high. Hind tarsal claws rather short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, distinctly and

rather broadly depressed, sulcus for the greater

part with transverse wrinkles, longitudinal cari-

nae present medially and posteriorly, obliterated

anteriorly, costulae partly present. Propodeal spir-

acle large, 2.9x as long as wide. Spiracular carina

narrow. Spiracular index 0.3. 1% gastral tergite

3.8x, 2" tergite 1.5% as long as wide. Petiole with

large glymmae, smooth or with very fine sculp-

ture in front of the glymmae. Petiolar suture part-

Iy present or complete. 1% gastral sternite for the

greater part with fine sculpture, with transverse

striae posteriorly. Thyridium oval. Thyridial in-

dex about 1.5. Epipleurum separated from the 34

gastral tergite anteriorly, the crease marked with

black. Epipleural index 0.5. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown to dark brown. Mandible broadly yellow

medially. Tegula brown. Wings hyaline. Stigma

yellowish brown medially. Front trochanter brown.

Front femur yellowish red. Front tibia and tarsus

yellow. Mid femur yellowish red on the apical 0.3.

Mid tibia yellow. Mid tarsus brown. Hind femur

completely black or the apical 0.2 marked with

red. Hind tibia broadly yellow medially, narrow-

ly brown basally and apically. 2" gastral tergite

red on the posterior 0.3-0.4. 3° and 4" tergites red.

Pilosity whitish.

d: Frons without or with very short radial

wrinkles. Flagellum with 51-57 segments. Prepec-

tal carina low. Transverse carina low and partly

obliterated. Metapleurum coriaceous, densely

punctate. Propodeum almost completely areolated,

costulae sometimes partly obliterated. Epipleural

index 0.7. Maxillary palpus yellow. Tegula yel-

low. Front and mid legs from the tip of the coxa

onwards yellowish, the femora yellowish red. Mid

coxa and trochanter sometimes dark brown. In

other characters similar to the 2.

Dusona rugifrons Horstmann, spec. nov.

Holotype (3): Russia, Kuril Isl., Shikotan, Krabo-

zavodsk, 15.8.1973 (PET).

Description. d: Body length 15 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons coriaceous and dense-

ly rugose-punctate dorsally, deeply depressed

ventrally and with strong radial wrinkles in the

depression, the wrinkles extending to half the

distance to the median ocellus, with a narrow

median longitudinal bulge. Antennal carina strong-

ly raised, with a transversely striate nose-like dor-

sal projection. Face and dorsal 0.6 of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

finely coriaceous and densely punctate ventrally,

its apex truncate medially. Malar space index 0.3.

Genal carina low ventrally, slightly bent sidewards,

joining the strongly raised oral carina at an acute

angle. Genal index 0.4. Temple finely coriaceous

and with fine but distinct rather dense punctation

medially. Flagellum with 56 segments, filiform,

the apical 0.3 distinctly attenuate, 2” segment

1.7x, shortest segment 1.0x, penultimate segment

1.2x as long as wide. Pronotum densely rugose-

punctate on asmooth background laterally on the

dorsal half, with longitudinal wrinkles ventrally.

Mesoscutum finely coriaceous, very densely punc-

145

tate. Scutellum densely punctate on asmooth back-

ground, with lateral carinae on the anterior 0.3.

Prescutellar groove with longitudinal striae. Pre-

pectal carina low, ventral part merging with the

pleural part, the latter complete, joining the ante-

rior edge of the mesopleurum. Transverse carina

present anteriorly. Mesopleurum very densely

punctate on asmooth background. Speculum fine-

ly coriaceous, almost smooth, shining. Depression

in front of the speculum with rather short longitu-

dinal striae, which merge with the punctation an-

teriorly. Metapleurum densely rugose. Pleural

carina complete. Areolet large, pointed, receiving

the 2"d recurrent vein at the middle. Nervellus

distinctly inclivous, distinctly intercepted. Ner-

vellar index 0.8. Tibial spur index 0.6. Hind femur

6.3x as long as high. Hind tarsal claws rather long,

with short pectinate teeth basally and medially.

Propodeum extending to 0.4 of the hind coxae,

broadly and not deeply depressed, sulcus for the

greater part transversely wrinkled, with some dis-

tinct longitudinal wrinkles posteriorly, costulae

partly present, longitudinal carinae strong and

almost complete. Propodeal spiracle rather large,

3.2x as long as wide. Spiracular carina narrow.

Spiracular index 0.5. 1* gastral tergite 4.0x, 2”

tergite 2.5x as long as wide. Petiole with large

glymmae, smooth in front of the glymmae. Petio-

lar suture fine, partly obliterated. 1“ gastral sterni-

te coriaceous, with fine transverse wrinkles. Thy-

ridium oval. Thyridial index about 2.0. Epipleu-

rum separated from the 3" gastral tergite anteriorly,

the crease marked with black. Epipleural index

0.4. Gaster strongly compressed posteriorly.

Colour: Black. Maxillary palpus dark brown.

Mandible slightly tinged with yellowish brown

dorsally. Wings hyaline. Stigma yellowish brown

medially. Front leg from the trochantellus onwards

yellowish red. Mid femur yellowish red on the

apical 0.2. Mid tibia yellowish red. Mid tarsus

dark brown. Hind tibia yellowish brown, slightly

darkened apically. 2" gastral tergite dorsally red

on the posterior 0.3, almost completely red lateral-

ly. 3“ and 4" tergites for the greater part red,

narrowly marked with black dorsally. 5" tergite

for the greater part reddish brown, tinged with

dark brown. Posterior tergites dark brown or black.

Gastral sternites red or reddish brown. Genital

clasper black with a reddish brown tip. Pilosity

whitish.

? unknown.

146

Dusona rugosa Horstmann, spec. nov.

Types. Holotype (?): Japan, Honshu, Sasayama,

28.5.1953 (SAP). — Paratypes: 1? same locality,

4.6.1952 (Hinz/ZSM); 1? South Korea, Suigen

(= Suwon), 1.6. 1928 (SAP).

Description. 2: Body length 13-14 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.9. Frons slightly depressed, coriaceous, in most

parts finely and densely punctate, with a distinct

median longitudinal carina, with short radial wrin-

kles, which extend to 0.3 the distance to the medi-

an ocellus. Antennal carina low and narrow. Face

and dorsal 0.8 of the clypeus coriaceous, densely

rugose and rugose-punctate. Clypeus flat, coria-

ceous and densely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina low and straight ventrally, joining the

scarcely raised oral carina at an acute angle. Genal

index 0.3-0.4. Temple finely coriaceous and with

fine disperse punctation medially. Flagellum with

51-52 segments, filiform, the apical 0.7 attenuate,

22d segment 1.5x, shortest segment 0.8x, penulti-

mate segment 1.1x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely punc-

tate. Scutellum with lateral carinae on the anterior

0.7. Prescutellar groove with few fine longitudinal

striae. Prepectal carina low, the ventral part merg-

ing with the pleural part, the latter complete, join-

ing the anterior edge of the mesopleurum. Trans-

verse carina present anteriorly. Mesopleurum fine-

ly coriaceous, rather densely punctate. Speculum

coriaceous and granulate-strigose, shining. De-

pression in front of the speculum with fine but

distinct longitudinal wrinkles, which merge with

the punctation anteriorly. Metapleurum finely

coriaceous, distinctly and densely punctate. Pleu-

ral carina strong. Areolet large, short-stalked, re-

ceiving the 2"! recurrent vein basad of the middle.

Nervellus slightly inclivous, distinctly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.7. Hind

femur 4.6x as long as high. Hind tarsal claws

rather long, with short pectinate teeth basally and

medially. Propodeum extending to 0.3 of the hind

coxae, broadly and not deeply depressed, sulcus

with rather fine irregular wrinkles anteriorly and

medially, with long transverse wrinkles posteri-

orly, costulae partly present, longitudinal carinae

distinct posteriorly, obliterated medially and an-

teriorly. Propodeal spiracle large, 3.4x as long as

wide. Spiracular carina narrow. Spiracular index

0.3. 1% gastral tergite 3.4x, 2" tergite 1.5x as long

as wide. Petiole with large glymmae, which are

bordered by longitudinal carinae dorsally, with

very fine longitudinal wrinkles in front of the

glymmae. Petiolar suture fine, partly obliterated.

1* gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.0. Epipleurum separated from

the 3" gastral tergite, the crease marked with black.

Epipleural index 0.6. Gaster distinctly compressed

posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly marked with yellow medially. Tegu-

la yellow. Wings hyaline. Stigma light brown me-

dially. Front trochanter yellow, marked with

brown. Front femur yellowish red, marked with

black ventrally. Front tibia and tarsus yellow. Mid

femur yellow on the apical 0.2. Mid tibia yellow.

Mid tarsus yellow basally, dark brown apically.

Hind tibia yellow medially, narrowly black basal-

ly, broadly black apically. 2" gastral tergite red on

the posterior 0.1-0.2. 3" and 4" tergites red, the 4"

tergite often marked with black. Posterior gastral

sternites sometimes reddish brown. Pilosity whit-

ish.

?d: Body length 13 mm. Flagellum with 54

segments. Propodeum with short transverse wıin-

kles also anteriorly and medially. Epipleural in-

dex 0.7. Tip of the front coxa, front trochanter and

front femur yellowish red. Mid femur yellowish

red dorsally and apically. Hind tibia rather nar-

rowly dark yellowish brown medially, narrowly

marked with black basally, very broadly marked

with black apically (apical black ring extending

over half of the tibia). 2" gastral tergite black

dorsally, marked with red laterally. 3“ and 4"

tergites for the greater part red, narrowly marked

with black dorsally. 5" and 6" tergites for the

greater part black, narrowly marked with red lat-

erally. Posterior gastral sternites black, marked

with red. Genital clasper black. In other characters

similar to the 2. Material: 14 Russia, Khabarovsk

Terr., Khekhtsir, near Khabarovsk (PET). This &

may belong to D. rugosa, but the colour pattern of

the gaster is rather different from that of the 2.

Dusona sachalini Hinz, 1985

Dusona sachalini Hinz, 1994

Hinz (1985: 308 ff., 1994: 37 f.) described this spe-

cies twice (see introduction). The descriptions of

both taxa differ from each other (in 1985 both

sexes are described, in 1994 oniy the ® is), and the

type series are different, although both taxa have

the same holotype. This specimen from Dolinsk /

Sakhalin/Russia (PET) has recently been studied

by the junior author.

Description. 2: Body length 9-10 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the anterior half of the mes-

oscutum. Ocellar index 1.0. Frons slightly de-

pressed, coriaceous, densely rugose-punctate,

without a median longitudinal carina and with-

out radial wrinkles. Antennal carina low and nar-

row, its rim slightly bent upwards (inconspicu-

ous). Face coriaceous, densely punctate. Clypeus

distinctly convex, coriaceous, in almost all parts

with very fine disperse punctation, its apex slight-

ly convex medially. Malar space index 0.6. Genal

carina low and almost straight ventrally, joining

the slightly raised oral carina at an acute angle.

Genal index 0.9. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 45-48 segments (after Hinz 1985: 308;

tips of antennae broken off in all available ? 2),

filiform, the apical 0.3 distinctly attenuate, 2" seg-

ment 2.3x, shortest segment 1.3x as long as wide.

Pronotum coriaceous and with fine dense puncta-

tion laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum coriaceous, dull,

for the greater part finely and rather densely punc-

tate, in some parts rugose. Scutellum coriaceous,

very densely punctate, with lateral carinae on the

anterior 0.3. Prescutellar groove for the greater

part smooth. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina obliterated. Mesopleurum dis-

tinctly coriaceous, dull, for the greater part dis-

tinctly and densely punctate. Speculum coriaceous,

dull. Depression in front of the speculum dorsally

with short longitudinal wrinkles, the ventral 0.7

granulate-strigose and dull, a narrow coriaceous

area without punctation in front of the impres-

sion. Metapleurum finely and densely rugose-

punctate. Pleural carina distinct. Areolet small,

long-stalked, receiving the 2" recurrent vein dis-

tad of the middle. Nervellus slightly reclivous,

not intercepted. Tibial spur index 0.6. Hind femur

6.1x as long as high. Hind tarsal claws short, with

rather long pectinate teeth basally and medially.

Propodeum extending to 0.3 of the hind coxae,

not depressed, in most parts with irregular wrin-

kles, almost completely areolated, area basalis part-

ly replaced by a longitudinal keel, area superome-

dia bordered by a carina posteriorly, costulae for

the greater part obliterated, longitudinal carinae

present anteriorly, indistinct medially and poste-

riorly (Fig. 30). Propodeal spiracle small, 1.7x as

147

long as wide. Spiracular carina narrow. Spiracular

index 0.7. 1% gastral tergite 2.8x, 2" tergite 1.3x as

long as wide. Petiole with large glymmae, which

are bordered by longitudinal carinae dorsally, for

the greater part smooth in front of the glymmae,

with some fine longitudinal striae. Petiolar suture

complete. 1% gastral sternite coriaceous. Thyrid-

ium oval. Thyridial index about 0.5. Epipleurum

separated from the 3% gastral tergite, the crease

indistinctly marked with black. Epipleural index

0.9. Gaster distinctly compressed posteriorly. Ovi-

positor index 0.4.

Colour: Black. Maxillary palpus dark brown.

Wings hyaline. Stigma dark brown medially. Front

femur yellowish brown dorsally and apically, black

basally and ventrally. Front tibia and tarsus yel-

lowish brown. Mid femur medium brown on the

apical 0.2. Mid tibia medium brown. Mid tarsus

dark brown. 2"! gastral tergite red on the posterior

0.3. 3X tergite red. 4" tergite red anteriorly, black

posteriorly (to a variable extend). Posterior gas-

tral sternites red or brown. Pilosity whitish.

d: Tangential lines on eyes and temples inter-

sect on the posterior half of the mesoscutum. Genal

index 0.7. Flagellum with 45 segments. Mesopleu-

rum for the greater part densely rugose-punctate

and rugose. Impression in front of the speculum

with rather long wrinkles, which merge with the

punctation anteriorly. Front femur almost com-

pletely yellowish brown. 3"! gastral tergite red

only on the anterior 0.7. 4" tergite black. In other

characters similar to the 9.

Host: Larva of Lasiocampidae (PET).

Material (2?%, 158): Russia: Primorye: Krasnoar-

meysk; Sakhalin: Dolinsk (PET); Kuril Isl.: Dubovoye

on Kunashir (Hinz/ZSM).

Dusona sasayamae Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Japan, Honshu, Sasayama,

21.5.1951 (SAP). — Paratypes: 3?? same locality,

dates 19.5.1953, 25.5.1953, 18.5.1955 (1? Hinz/

Z5M, 22275AP):

Description. 2: Body length 13-17 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.9. Frons slightly de-

pressed, coriaceous, densely and finely rugose,

with a fine median longitudinal carina, with short

radial wrinkles, which extend to 0.3 the distance

to the median ocellus. Antennal carina low and

148

narrow. Face and dorsal 0.7 of the clypeus densely

rugose-punctate. Clypeus flat, coriaceous and with

rather dense punctation ventrally, its apex slight-

ly concave medially. Malar space index 0.3. Genal

carina slightly raised ventrally, straight, joining

the distinctly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 56-60 segments, filiform, the apical 0.3 dis-

tinctly attenuate, 2"? segment 1.8x, shortest seg-

ment 0.9x, penultimate segment 1.2x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal half, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum finely co-

riaceous and densely punctate. Scutellum with

lateral carinae on the anterior 0.3. Prescutellar

groove smooth. Prepectal carina low, ventral part

merging with the pleural part, the latter almost

joining the anterior edge of the mesopleurum.

Transverse carina obliterated. Mesopleurum dense-

ly punctate on a finely coriaceous and shining

background. Speculum smooth or almost smooth.

Depression in front of the speculum with fine

long longitudinal wrinkles, which merge with the

punctation anteriorly. Metapleurum very densely

punctate on an almost smooth background. Pleu-

ral carina strong. Areolet large, short-stalked, re-

ceiving the 2" recurrent vein basad of the middle.

Nervellus almost vertical, strongly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 5.5x as long as high. Hind tarsal claws

rather long, with long pectinate teeth almost over

their total length. Propodeum extending to 0.3 of

the hind coxae, broadly and not deeply depressed,

sulcus with irregular wrinkles anteriorly and me-

dially, with long transverse wrinkles posteriorly,

costulae partly present, longitudinal carinae al-

most complete. Propodeal spiracle large, 3.3x as

long as wide. Spiracular carina narrow. Spiracular

index 0.6. 1% gastral tergite 3.6x, 2" tergite 1.8x as

long as wide. Petiole with large glymmae, which

are bordered by longitudinal carinae dorsally and

ventrally, with very fine sculpture in front of the

glymmae. Petiolar suture complete. 1* gastral ster-

nite finely coriaceous anteriorly, transversely stri-

ate posteriorly. Thyridium oval. Thyridial index

about 1.5. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.7-0.8. Gaster distinctly compressed

posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly marked with yellow medially.

Tegula dark brown. Wings slightly tinged with

brown. Stigma dark brown medially. Front leg

from the tip of the coxa onwards yellow, the fe-

mur yellowish red. Mid femur black, marked with

yellowish red dorsally and apically. Mid tibia and

tarsus yellow, the tarsus marked with black api-

cally. Hind tibia yellowish brown, sometimes nar-

rowly marked with brown basally and apically.

Hind tarsus yellowish brown basally, dark brown

apically. 2" gastral tergite red on the posterior 0.2.

34 and 4" tergites red. 5" tergite for the greater

part red, narrowly marked with black dorsally.

6" and 7" tergites for the greater part black, nar-

rowly marked with red ventrolaterally. Gastral

sternites red. Pilosity whitish.

d unknown.

Dusona scalprata Horstmann, spec. nov.

Holotype (3): Russia, S Primorye, Petrovka,

20.6.1962 (MOS).

Description. d: Body length about 15 mm. Tem-

ples strongly constricted behind the eyes, 0.7x as

long as the width of an eye, tangential lines on

eyes and temples intersect on the posterior half of

the mesoscutum. Ocellar index 0.7. Frons slightly

depressed, coriaceous, finely and dispersely punc-

tate, with a fine but distinct median longitudinal

carina, with short and indistinct radial wrinkles.

Antennal carina low and narrow. Face and dorsal

0.8 ofthe clypeus coriaceous, finely and very dense-

ly punctate. Clypeus flat, coriaceous and rather

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.4. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 58 segments,

filiform, the apical 0.3 distinctly attenuate, 2” seg-

ment 1.9x, shortest segment 1.0x, penultimate seg-

ment 1.3x as long as wide. Pronotum with longi-

tudinal wrinkles laterally, with additional punc-

tation dorsolaterally. Mesoscutum and scutellum

coriaceous, very densely punctate. Scutellum with

lateral carinae on the anterior 0.6. Prescutellar

groove smooth medially, with longitudinal striae

laterally. Ventral part of the prepectal carina low,

merging with the transverse carina. Pleural part

of the prepectal carina for the greater part obliter-

ated, replaced by some wrinkles medially. Meso-

pleurum strongly and densely punctate on a

smooth background. Speculum finely granulate-

strigose, shining. Depression in front of the specu-

lum with distinct longitudinal striae, which merge

with the punctation anteriorly. Metapleurum

densely rugose, not coriaceous. Pleural carina com-

plete. Areolet large, sessile, receiving the 2" re-

current vein slightly basad of the middle. Nervel-

lus slightly inclivous, intercepted. Nervellar in-

dex 0.7. Tibial spur index 0.7. Hind femur 4.2x as

long as high. Hind tarsal claws rather long, with

short pectinate teeth basally. Propodeum extend-

ing to 0.3 of the hind coxae, scarcely depressed,

with fine transverse wrinkles almost over its total

width, costulae present, fine, longitudinal carinae

almost completely obliterated. Propodeal spiracle

large, 3.3x as long as wide. Spiracular carina nar-

row. Spiracular index 0.4. 1% gastral tergite 4.4x,

2" tergite 4.6x as long as wide. Petiole without

glymmae, narrow lateral areas weakly indicated,

smooth in from of the lateral areas. Petiolar suture

almost completely obliterated. 1* gastral sternite

smooth. Thyridium large, oval. Thyridial index

about 1.5. Epipleurum not separated from the 3'4

gastral tergite, the latter with indistinct black lat-

eral stripes well above the ventrolateral edge.

Gaster very strongly compressed from the 2”! seg-

ment onwards.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly marked with yellow medially. Tegu-

la dark yellowish brown. Wings hyaline. Stigma

dark brown medially. Front and mid legs from the

tip of the coxa onwards yellow. Hind femur yel-

low on the apical 0.2. Hind tibia and tarsus yel-

low, the tarsus tinged with yellowish brown api-

cally. 2"! gastral tergite black dorsally, broadly

marked with red laterally. 3" to 7‘ tergites nar-

rowly black dorsally, broadly red laterally. Gas-

tral sternites red. Genital clasper red. Pilosity

whitish.

? unknown.

Dusona schikotani Hinz, 1994

This is a variable and poorly defined species.

Description. ?: Body length 6-9 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.9-1.1. Frons not de-

pressed, coriaceous, densely rugose-punctate, with

a fine median longitudinal carina, with short and

indistinct radial wrinkles. Antennal carina low

and narrow. Face and dorsal half of the clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

coriaceous and with fine disperse punctation ven-

trally, its apex slightly convex medially. Malar

space index 0.6. Genal carina low ventrally, slight-

ly bent sidewards, joining the slightly raised oral

carina at an acute angle. Genal index 0.2-0.3. Tem-

ple finely coriaceous and with very fine disperse

punctation medially. Flagellum with 36-42 seg-

149

ments, filiform, the apical 0.2 slightly attenuate,

2"d segment 2.3x, shortest segment 1.2x, penulti-

mate segment 1.2x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely punc-

tate. Scutellum with lateral carinae on the anterior

0.6-0.8. Prescutellar groove smooth medially, with

longitudinal striae laterally. Prepectal carina low,

ventral part merging with the pleural part, the

latter almost complete or complete, in the latter

case joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

coriaceous, dull, very densely punctate. Specu-

lum punctate posteriorly, finely coriaceous and

shining anteriorly. Depression in front of the specu-

lum with distinct longitudinal striae, which merge

with the punctation anteriorly. Metapleurum

densely rugose and rugose-punctate. Pleural car-

ina distinct. Areolet rather small, stalked, receiv-

ing the 2” recurrent vein distad of the middle.

Nervellus vertical or slightly reclivous, weakly

intercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 4.8-5.4x as long as high. Hind

tarsal claws short, with short pectinate teeth ba-

sally. Propodeum extending to 0.4 of the hind

coxae, shallowly and narrowly depressed, with

short transverse wrinkles in the longitudinal fur-

row, with irregular wrinkles beside the furrow,

with long transverse wrinkles posteriorly, carinae

almost completely obliterated. Propodeal spiracle

small, 2.6x as long as wide. Spiracular carina nar-

row. Spiracular index 0.5. 1” gastral tergite 4.0x,

2" tergite 1.7x as long as wide. Petiole smooth

laterally, without glymmae or narrow lateral are-

as. Petiolar suture obliterated. 1* gastral sternite

smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter without black lateral

stripes. Gaster distinctly compressed posteriorly.

Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown. Mandible tinged with dark brown medial-

ly. Flagellum sometimes tinged with brown api-

cally. Tegula dark brown. Wings hyaline. Stigma

dark brown medially. Front femur, tibia and tar-

sus yellowish brown, the femur marked with black

basally, the tibia marked with whitish yellow on

the dorsal side. Mid femur whitish yellow on the

apical 0.3. Mid tibia whitish yellow basally and

medially, brown apically. Mid tarsus whitish yel-

low basally, brown apically (to a variable extend).

Hind tibia for the greater part black, marked with

white subbasally on the dorsal side, the white

mark sometimes almost lacking. Hind tarsus black-

ish, the basitarsus marked with white basally. 24

150

gastral tergite red on the posterior 0.3. 3° and 4"

tergites red. Pilosity whitish.

Material (5??): Russia: Primorye: Slavyanka (LIN),

Spassk Dalniy (Hinz/ZSM); Kuril Isl.: Sernovodsk on

Kunashir (paratype of D. micrator), Krabozavodsk on

Shikotan (PET). North Korea: Puryong (SAW).

? d: Body length 7-9 mm. 38-42 flagellar segments,

2"! segment 1.7x, shortest segment 1.0x, penulti-

mate segment 1.3x as long as wide. Hind femur

4.3-4.7x as long as wide. Propodeum with almost

complete carinae, the longitudinal carinae obliter-

ated anteriorly. Thyridium oval. Thyridial index

about 2.0. 3"' gastral tergite sometimes with black

lateral stripes. Hind tibia white medially, narrow-

ly black basally, broadly black apically. 2" gastral

tergite red on the posterior 0.1-0.2. 4" tergite red

on the anterior half. In other characters similar to

the 2. Material: 19 Russia, Primorye, Partizansk

(PET); 254 North Korea, Hyangsan (18 Hinz/

ZSM, 15 SAW). The differences between these dd

and the ?? of D. schikotani may be interpreted as

sexual differences.

Dusona scolator Hinz, 1985

Description. 2: Body length 6-7 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons scarcely depressed, coriaceous

and finely rugose-punctate, with a fine median

longitudinal carina, with distinct radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina low and narrow. Face

and dorsal half of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, finely coriaceous

and rather densely punctate ventrally, its apex

truncate medially. Malar space index 0.5. Genal

carina low ventrally, distinctly bent forwards, join-

ing the oral carina at the base of the mandible at a

right angle. Temple finely coriaceous and with

fine disperse punctation medially. Flagellum with

34-37 segments, filiform, the apical 0.2 attenuate,

2"d segment 2.4x, shortest segment 1.0x, penulti-

mate segment 1.3x as long as wide (sometimes

flagellum stouter, shortest segment 0.9x, penulti-

mate segment 1.0x as long as wide). Pronotum

densely rugose-punctate laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, very densely

punctate. Scutellum with lateral carinae on the

anterior 0.7. Prescutellar groove almost smooth.

Ventral part of the prepectal carina merging with

the transverse carina, both carinae distinctly raised,

the former distinctly emarginate medially. Pleu-

ral part of the prepectal carina low, present ven-

trally and medially, the dorsal 0.4 obliterated.

Mesopleurum distinctly and densely punctate on

a slightly coriaceous and shining background.

Speculum granulate-strigose or with fine oblique

striae, somewhat shining. Depression in front of

the speculum for the greater part with fine longi-

tudinal striae, which merge with the punctation

anteriorly. Metapleurum finely and densely ru-

gose-punctate and rugose. Pleural carina distinct.

Areolet rather large, stalked, receiving the 2" re-

current vein distad of the middle. Nervellus verti-

cal or slightly reclivous, weakly intercepted. Ner-

vellar index 0.8. Tibial spur index 0.6. Hind femur

5.0x as long as high. Hind tarsal claws short, with

short pectinate teeth basally. Propodeum extend-

ing to 0.4 of the hind coxae, broadly and not

deeply depressed, sulcus for the greater part ir-

regularly wrinkled, with transverse wrinkles pos-

teriorly, carinae almost completely obliterated.

Propodeal spiracle rather small, 3.0x as long as

wide. Spiracular carina narrow. Spiracular index

0.6. 1% gastral tergite 4.1x, 2" tergite 1.6x as long

as wide. Petiole without glymmae or with sınall

and shallow glymmae, smooth in front of the glym-

mae. Petiolar suture partly obliterated. 1% gastral

sternite smooth. Thyridium oval. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter with distinct black lateral

stripes well above the ventrolateral edge. Gaster

distinctly compressed posteriorly. Ovipositor in-

dexa0:.5:

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly yellowish red medially, some-

times almost completely black. Wings hyaline.

Stigma yellowish brown medially. Front and mid

femora, tibiae and tarsi yellowish red, the mid

femur marked with black basally. Hind tibia yel-

lowish brown medially, narrowly black basally,

broadly black apically. Hind tarsus black dorsal-

ly, for the greater part yellowish brown ventrally.

2nd gastral tergite red on the posterior 0.3-0.4.

3" and 4" tergites red. Pilosity whitish.

d: Nervellus sometimes not intercepted. Pro-

podeum with transverse wrinkles also medially.

Tegula yellowish brown. Front trochanter yellow-

ish red. In other characters similar to the 2.

Material (3??, 15): Russia: Yakutsk A.S.S.R.: Bestya-

kh on River Amga (Hinz/ZSM), Chuchur-Muran, Vi-

lyuyskogo trakta (PET). The species also occurs in

Europe: 27? Scotland, Cairngorms in Inverness-shire,

Braemer in Aberdeen-shire (EDI) 32%, 14 Italy, Kal-

tern (= Caldaro) in Südtirol (Hinz/ZSM).

Dusona scriptor Hinz & Horstmann, spec. nov.

Holotype (?): Japan, Hokkaido, Tokachi-dake,

18.7.1954 (SAP).

Description. 2: Body length 15 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.2. Frons scarcely depressed, coriaceous, densely

punctate, with a fine median longitudinal carina,

with rather short radial wrinkles, which extend to

0.4 the distance to the median ocellus. Antennal

carina low and narrow. Face and dorsal 0.7 of the

clypeus coriaceous, densely punctate, partly ru-

gose. Clypeus flat, coriaceous and rather densely

punctate ventrally, its apex truncate and with a

smooth bulge medially. Malar space index 0.4.

Genal carina low and rather straight ventrally,

joining the distinctly raised oral carina at an acute

angle. Genal index 0.5. Temple coriaceous and

with fine disperse punctation medially. Flagellum

with 61 segments, filiform, its apical 0.4 distinctly

attenuate, 2” segment 1.5x, shortest segment 1.0x,

penultimate segment 1.1x as long as wide. Prono-

tum coriaceous and very densely punctate lateral-

ly, with longitudinal wrinkles in the dorsoventral

furrow. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.8. Prescutellar groove with fine

longitudinal striae. Prepectal carina low, ventral

part merging with the pleural part, the latter for

the greater part present, its dorsal 0.2 indistinct

and covered with wrinkles. Transverse carina ob-

literated. Mesopleurum finely coriaceous, rather

densely punctate. Speculum with fine oblique stri-

ae dorsally, finely coriaceous and shining ventral-

ly. Depression in front of the speculum dorsally

with longitudinal wrinkles, which merge with the

punctation anteriorly, the ventral half finely co-

riaceous and granulate-strigose. Metapleurum

coriaceous, densely punctate. Pleural carina dis-

tinct. Areolet large, short-stalked, receiving the

2"d recurrent vein distinctly basad of the middle.

Nervellus slightly inclivous, distinctly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 5.3x as long as high. Hind tarsal claws

rather short, with short dense pectinate teeth al-

most over their total length. Propodeum extend-

ing to 0.4 of the hind coxae, distinctly and rather

broadly depressed, sulcus with rather short trans-

verse wrinkles anteriorly and medially, with long

transverse wrinkles posteriorly, costulae partly

present, longitudinal carinae indicated medially

and posteriorly. Propodeal spiracle large, 3.6x as

long as wide. Spiracular carina narrow. Spiracular

151

index 0.6. 1* gastral tergite 3.1x, 2" tergite 1.9x as

long as wide. Petiole with large and deep glym-

mae, which are bordered by longitudinal carinae

dorsally, smooth in front of the glymmae. Petiolar

suture obliterated. 1“ gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum not separated from the 3" gastral tergite, the

latter without black lateral stripes. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Tegula dark

brown. Wings hyaline. Stigma medium brown

medially. Front trochanter yellow. Front trochan-

tellus and femur yellowish red, marked with

brown ventrally. Front tibia and tarsus yellow.

Mid femur yellowish brown on the apical 0.2. Mid

tibia yellowish brown. Mid tarsus medium brown.

Hind tibia dark yellowish brown, narrowly marked

with black basally. Hind tarsus dark brown. Gaster

black dorsally. 2” tergite narrowly marked with

red laterally. 3“ and 4" tergites broadly marked

with red laterally. Posterior tergites narrowly

marked with red laterally. Gastral sternites red.

Pilosity whitish.

d unknown.

Dusona scutellator Hinz, 1979

Description. 2: Body length 15-16 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.9-1.0. Frons slightly de-

pressed, coriaceous, finely and rather densely

punctate, with an indistinct median longitudinal

carina, with distinct radial wrinkles, which ex-

tend to 0.3 the distance to the median ocellus.

Antennal carina rather low, the rim bent upwards

and widened to a narrow crescent-shaped plate,

the latter transversely striate. Face and greater

part of the clypeus coarsely and densely rugose-

punctate. Clypeus flat, with a narrow less densely

punctate area ventrally, its apex truncate medial-

ly. Malar space index 0.4. Genal carina slightly

raised ventrally, almost straight, joining the dis-

tinctly raised oral carina at an acute angle. Gena

slightly depressed close to the junction. Genal

index 0.6. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

58-66 segments, filiform, the apical half distinctly

attenuate, 2"! segment 1.4x, shortest segment 1.0x,

penultimate segment 1.2x as long as wide. Prono-

tum coriaceous and densely punctate laterally on

the dorsal 0.2-0.5, with longitudinal wrinkles ven-

152

trally. Mesoscutum coriaceous, very densely punc-

tate. Scutellum coriaceous, rather densely or dense-

ly punctate, with lateral carinae on the anterior

0.6. Prescutellar groove with distinct longitudinal

striae. Prepectal carina low, ventral part merging

with the pleural part, the dorsal 0.1-0.2 of the

latter replaced by several wrinkles. Transverse

carina present anteriorly. Mesopleurum distinctly

and densely punctate on a finely coriaceous and

shining background. Speculum punctate posteri-

orly, granulate-strigose and shining anteriorly.

Depression in front of the speculum dorsally with

fine longitudinal striae, which merge with the

punctation anteriorly, the ventral haif finely gran-

ulate-strigose and shining. Metapleurum densely

rugose-punctate or rugose, the wrinkles smooth.

Pleural carina distinct. Areolet large, narrowly

sessile, receiving the 2" recurrent vein at or slight-

ly distad of the middle. Nervellus slightly in-

clivous, distinctly intercepted. Nervellar: index

0.6-0.7. Tibial spur index 0.5-0.6. Hind femur 5.7x

as long as high. Hind tarsal claws short, with

rather long pectinate teeth almost over their total

length. Propodeum extending to 0.3 of the hind

coxae, broadly but not deeply depressed, sulcus

transversely wrinkled, costulae partly present or

complete, longitudinal carinae present medially

and posteriorly, obliterated anteriorly, sometimes

partly covered with wrinkles. Propodeal spiracle

large, 4.4x as long as wide. Spiracular carina nar-

row. Spiracular index 0.3. 1% gastral tergite 3.2x,

2nd tergite 1.9x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal

carinae dorsally, smooth in front of the glymmae.

Petiolar suture fine, partly obliterated. 1“ gastral

sternite smooth. Thyridium oval. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter without black lateral

stripes. Gaster distinctly compressed posteriorly.

Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Tegula medi-

um brown. Wings slightly infuscate. Stigma me-

dium brown medially. Front leg from the tip of

the coxa onwards yellowish red. Mid femur yel-

lowish red on the apical 0.3. Mid tibia and tarsus

yellowish red. Hind femur yellowish brown on

the apical 0.1-0.2. Hind tibia broadly yellowish

brown medially, narrowly black basally, narrow-

ly and indistinctly brown apically. Hind tarsus

dark brown. 2" gastral tergite black dorsally,

broadly red laterally. 3" to 8" tergites red, some-

times narrowly darkened dorsally. Gastral ster-

nites red. Pilosity whitish.

d unknown.

Material (5??): Russia: Primorye: Novokachalinsk

(PET), Yakovlevka (MOS). Japan: Hokkaido: Apoi-

dake (Hinz/ZSM, SAP); Honshu: Karuizawa (HEL).

Dusona setator Hinz, 1985

Description. 2: Body length 8 mm. Temples not

constricted behind the eyes, 1.1x as long as the

width of an eye, tangential lines on eyes and tem-

ples running parallel. Ocellar index 1.0. Frons not

depressed, coriaceous, densely punctate or ru-

gose-punctate, without a median longitudinal ca-

rina and without radial wrinkles. Antennal sock-

ets close to each other. Antennal carina low and

narrow. Face and dorsal 0.7 of the clypeus densely

rugose-punctate. Clypeus slightly concave, rather

densely punctate on an almost smooth background

ventrally, its apex truncate medially. Malar space

index 0.4. Genal carina low and straight ventrally,

joining the distinctly raised oral carina at an acute

angle. Genal index 0.6. Temple with fine disperse

punctation on a smooth background medially.

Flagellum with 33-35 segments, slightly fusiform,

2"! segment 1.9x, shortest segment 0.9x, penulti-

mate segment 1.0x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.2, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum densely punctate on a

smooth background. Scutellum with lateral cari-

nae on the anterior 0.6. Prescutellar groove smooth

medially, with fine longitudinal striae laterally.

Prepectal carina low, ventral part merging with

the pleural part, the dorsal 0.4 of the latter obliter-

ated. Transverse carina indicated anteriorly. Meso-

pleurum with disperse or rather dense punctation

on a smooth background. Speculum punctate pos-

teriorly, with fine oblique striae anteriorly. De-

pression in front of the speculum with distinct

rather short longitudinal striae, which merge with

the punctation anteriorly. Metapleurum with rath-

er dense or dense punctation on a finely coria-

ceous or finely rugose background. Pleural carina

complete. Areolet large, narrowly sessile, receiv-

ing the 2" recurrent vein at or distad of the mid-

dle. Nervellus vertical, distinctly intercepted. Ner-

vellar index 0.8. Tibial spur index 0.7. Hind femur

3.9x as long as high. Hind tarsal claws rather long,

with few short pectinate teeth basally. Propodeum

extending to 0.3 of the hind coxae, scarcely de-

pressed, with some fine transverse wrinkles in the

broad and very shallow sulcus, in other parts

irregularly wrinkled, costulae partly present, lon-

gitudinal carinae present posteriorly, obliterated

anteriorly and medially. Propodeal spiracle rath-

er small, 2.8x as long as wide. Spiracular carina

indistinct. Spiracular index 0.5. 1% gastral tergite

2.9x, 2" tergite 1.3x as long as wide. Petiole smooth

laterally, sometimes with very fine longitudinal

striae, without glymmae, sometimes with very

small and shallow pits. Petiolar suture fine.

1" gastral sternite for the greater part smooth, with

some transverse striae posteriorly. Thyridium oval.

Thyridial index about 0.5. Epipleurum not sepa-

rated from the 3" gastral tergite, the latter without

black lateral stripes. Gaster slightly compressed

posteriorly. Ovipositor index 1.3.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly dark brown medially. Wings

hyaline. Stigma dark brown. Front leg from the

femur onwards yellowish red, the femur marked

with black basally. Mid femur brown on the apical

0.1. Mid tibia and tarsus brown. 2" gastral tergite

red on the posterior 0.6-0.8, sometimes complete-

ly red. 3 and 4" tergites red. Pilosity whitish.

d: Flagellum filiform. Propodeum not de-

pressed, almost without transverse wrinkles. Man-

dible broadly yellow medially. Front coxa marked

with yellow. 2”! gastral tergite red on the posteri-

or 0.2. In other characters similar to the ®.

Material (522, 239): Russia: Chita Prov.: Doroninsk-

Chunguruk. Mongoliya: Dornod Prov.: Choybalsan;

Selenge Prov.: Bayan-Gol (PET); Sühbaatar Prov.: Bar-

uun Urt (Hinz/ZSM); Central Prov. (= Töv Prov.):

Kerulen (PET).

Dusona signator (Brauns, 1895)

This species is rather variable, particularly in body

size. The differentiation of small specimens of

D. signator from D. circumcinctus may be difficult

(see key, couplet 140).

Description. 2: Body length 10-14 mm. Temples

strongly constricted behind the eyes, 0.7x as long

asthe width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the mes-

oscutum. Ocellar index 0.7. Frons slightly de-

pressed, coriaceous, distinctly and densely punc-

tate, with a distinct median longitudinal carina,

without radial wrinkles. Antennal carina low and

narrow. Face and dorsal 0.8 of the clypeus finely

coriaceous, densely rugose-punctate and rugose.

Clypeus slightly concave, finely coriaceous and

rather densely or densely punctate ventrally, its

apex truncate medially. Malar space index 0.4.

Genal carina low ventrally, slightly bentsidewards,

joining the distinctly raised oral carina almost at a

right angle. Genal index 0.4. Temple finely coria-

ceous and with fine disperse punctation medially.

153

Flagellum with 49-58 segments, filiform, the api-

cal 0.3 attenuate, 2”! segment 1.7x, shortest seg-

ment 1.0x, penultimate segment 1.3x as long as

wide. Pronotum densely rugose-punctate on a

smooth background laterally on the dorsal 0.4,

with longitudinal wrinkles ventrally. Mesoscu-

tum finely coriaceous, very densely punctate.

Scutellum densely punctate on a smooth back-

ground, with lateral carinae on the anterior 0.3.

Prescutellar groove smooth medially, with longi-

tudinal striae laterally. Ventral part of the prepec-

tal carina merging with the transverse carina, the

former low, the latter slightly raised. Pleural part

of the prepectal carina obliterated dorsally and

ventrally, present as a rather long wrinkle medial-

ly. Mesopleurum distinctly and densely punctate

on a finely coriaceous or smooth background.

Speculum finely coriaceous, smooth ventrally (to

a variable extend), strongly shining. Depression

in front of the speculum with distinct longitudinal

striae, which merge with the punctation anterior-

ly. Metapleurum densely punctate or rugose-punc-

tate on a smooth background. Pleural carina dis-

tinct. Areolet large, pointed, receiving the 2" re-

current vein at the middle. Nervellus almost

vertical, distinctly intercepted, the upper branch

slightly bent inwards. Nervellar index 0.7. Tibial

spur index 0.7. Hind femur 5.2x as long as high.

Hind tarsal claws rather long, with distinct pecti-

nate teeth basally and medially. Propodeum ex-

tending to the middle of the hind coxae, shallowly

and narrowly depressed anteriorly, not depressed

posteriorly, with short or long transverse wrin-

kles anteriorly in the longitudinal furrow, irregu-

larly wrinkled beside the furrow, with long trans-

verse wrinkles posteriorly, costulae complete and

distinct, longitudinal carinae distinct posteriorly,

obliterated anteriorly and medially. Propodeal

spiracle large, 2.8x as long as wide. Spiracular

carina widened. Spiracular index 0.2. 1“ gastral

tergite 4.8x, 2"! tergite 2.7x as long as wide. Peti-

ole without glymmae, with indistinct narrow lat-

eral areas, smooth in front of these areas. Petiolar

suture almost completely obliterated. 1“ gastral

sternite smooth, sometimes with transverse striae

posteriorly. Thyridium oval. Thyridial index about

1.5. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellowish red medially and apically. Man-

dible broadly yellowish red medially. Tegula yel-

low. Wings hyaline. Stigma medium brown medi-

ally. Front leg from the tip of the coxa onwards

yellowish red, the tibia and tarsus yellow. Mid

femur yellowish red dorsally, brown ventrally.

Mid tibia and tarsus yellow. Hind tibia yellowish

brown medially, narrowly brown or black basal-

ly, broadiy medium brown apically, or the dark

apical mark lacking. Hind tarsus dark brown or

black. 2" gastral tergite broadly black dorsally,

narrowly red laterally. 3“ and 4" tergites narrow-

ly black dorsally, broadly red laterally. 5" to 8"

tergites for the greater part black, marked with

red ventrolaterally. Gastral sternites red. Pilosity

whitish.

d: Similar to the ?. Genital clasper red.

Material (2322, 1238): Russia: Yevreyskiy A.O.: Amur-

zet, Birofeld; Primorye: Kamen-Rybolov, Novokacha-

linsk; Kuril Isl.: Dubovoyesand Tretyakovo on Kuna-

shir (PET). China: Heilongjing Prov.: Charbin (= Har-

bin) (AEI), Liaoning Prov.: Kaigen (= Kaiyuan) (SAP).

North Korea: Hyangsan, Kyongsong, Wonsan (SAW).

Japan: Hokkaido: Jozankei, Sapporo (AEI); Honshu:

Hakone near Tokyo (OTT), Kamikochi near Norikura-

dake (AEI), Nagano (OTT), Hashimoto and Yokoha-

ma near Tokyo (AEI).

Dusona similator Hinz, 1994

Description. 2: Body length 11-14 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes

and temples intersect on the posterior half of the

mesoscutum. Ocellar index 0.7. Frons slightly de-

pressed, coriaceous, finely and rather densely

punctate, with a very fine median longitudinal

carina, with short radial wrinkles. Antennal cari-

na low and narrow. Face and dorsal 0.6 of the

clypeus coriaceous, densely rugose-punctate and

rugose. Clypeus flat, coriaceous and rather dense-

ly punctate ventrally, its apex truncate or slightly

concave medially. Malar space index 0.3. Genal

carina low and straight ventrally, joining the low

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 58-63 seg-

ments, filiform, the apical 0.3 distinctly attenuate,

2"d segment 1.5x, shortest segment 0.8x, penulti-

mate segment 1.4x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, dull, densely

punctate. Scutellum with lateral carinae on the

anterior half. Prescutellar groove smooth medial-

ly, with longitudinal striae laterally. Prepectal car-

ina low, ventral part merging with the pleural

part, the latter complete, joining the anterior edge

of the mesopleurum. Transverse carina obliterat-

ed. Mesopleurum with rather fine and rather dense

or disperse punctation on a coriaceous and some-

what shining background. Speculum punctate

posteriorly, finely coriaceous and shining anteri-

orly. Depression in front of the speculum dorsally

with fine longitudinal striae, which merge with

the punctation anteriorly, the ventral half finely

granulate-strigose. Metapleurum densely rugose-

punctate and rugose. Pleural carina distinct. Are-

olet large, short-stalked, receiving the 2" recur-

rent vein distinctly basad of the middle. Nervellus

almost vertical, distinctly intercepted. Nervellar

index 0.7. Tibial spur index 0.6. Hind femur 5.5x

as long as high. Hind tarsal claws rather short,

with short pectinate teeth basally and medially.

Propodeum extending to 0.3 of the hind coxae,

rather broadly and shallowly depressed, sulcus

for the greater part irregularly wrinkled or with

short transverse wrinkles in the midline, with long

transverse wrinkles posteriorly, costulae obliter-

ated, longitudinal carinae present posteriorly and

sometimes also medially, obliterated anteriorly.

Propodeal spiracle large, 4.5x as long as wide.

Spiracular carina sometimes indistinct. Spiracular

index 0.3-0.5. 1% gastral tergite 3.7x, 2"X tergite

2.0x as long as wide. Petiole without glymmae or

with very small and shallow glymmae at the ante-

rior edge of narrow lateral areas, smooth in front

of these areas. Petiolar suture partly obliterated.

1% gastral sternite finely coriaceous or smooth.

Thyridium oval. Thyridial index about 1.3. Epi-

pleurum separated from the 3" gastral tergite, the

crease marked with black. Epipleural index 0.7-0.8.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.5.

Colour: Black. Maxillary palpus medium

brown basally, yellowish brown apically. Mandi-

ble broadly yellow medially. Wings hyaline. Stig-

ma dark brown medially. Front troachanter marked

with yellow. Front femur black basally and ven-

trally, yellowish red dorsally and apically. Front

tibia and tarsus yellow. Mid femur yellowish

brown on the apical 0.2. Mid tibia yellowish brown.

Mid tarsus dark brown. Hind tibia yellowish

brown medially, narrowly black basally, broadly

black apically. 2” gastral tergite red on the poste-

rior 0.2-0.3. 3% and 4" tergites red. Pilosity whit-

ish.

d: Sulcus of the propodeum sometimes irreg-

ularly wrinkled, with some longitudinal wrinkles

posteriorly. Maxillary palpus yellowish red. Tegula

yellowish brown to brown. Front and mid legs

from the tip of the coxa onwards yellow, the mid

femur marked with brown ventrally. Hind tibia

sometimes narrowly brown apically. In other char-

acters similar to the ®.

Material (4??, 338): Japan: Hokkaido: Jozankei (Hinz /

ZSM), Sapporo; Honshu: Karuizawa (AEI), Tokyo

(OTT).

Variety: 3% to 6" gastral tergites and sternites

red. In other characters similar to the nominate

morphe. Material: 19 Russia, Khabarovsk Terr.,

Khekhtsir near Khabarovsk (PET). This specimen

may represent a separate subspecies or an indi-

vidual variety.

Dusona sobolicida (Förster, 1868)

Description. 2: Body length 8-10 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8-0.9. Frons scarcely depressed, coria-

ceous, densely rugose-punctate, with a fine medi-

an longitudinal carina, with distinct radial wrin-

kles, which extend to 0.7 the distance to the median

ocellus. Antennal carina narrow, distinctly but

not strongly raised, the rim not bent upwards

(Fig. 5), with fine transverse striae. Face and dor-

sal 0.8 of the clypeus densely rugose-punctate or

rugose on a finely coriaceous or smooth back-

ground. Clypeus flat, coriaceous and rather dense-

Iy punctate ventrally, its apex truncate medially.

Malar space index 0.4. Genal carina distinctly

raised ventrally, slightly bent sidewards, joining

the distinctly raised oral carina at an acute angle,

both carinae forming a lopsided triangular pyra-

mid, the tip of the latter shifted ventrally and

situated behind the base of the mandible. Gena

depressed close to the junction, the depression

about half as deep as the width of the 2"4 flagellar

segment. Genal index about 0.2. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 45-46 segments, filiform,

the apical 0.3 attenuate, 2”! segment 1.8x, shortest

segment 0.8x, penultimate segment 1.0x as long

as wide. Pronotum densely rugose-punctate later-

ally on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

very densely punctate. Scutellum with lateral car-

inae on the anterior 0.4. Prescutellar groove al-

most completely smooth. Ventral part of the pre-

pectal carina merging with the transverse carina,

both carinae somewhat raised, the former not

emarginate medially. Pleural part of the prepectal

carina low, the dorsal 0.3 obliterated and replaced

by several wrinkles. Mesopleurum distinctly co-

riaceous, rather densely or densely punctate.

Speculum granulate-strigose or with fine oblique

striae. Depression in front of the speculum with

155

distinct long longitudinal striae, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, densely rugose-punctate and rugose. Pleu-

ral carina distinct. Areolet rather small, short-

stalked, receiving the 2” recurrent vein slightly

distad of the middle. Nervellus vertical, weakly

intercepted. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 4.4x as long as high. Hind tarsal

claws short, with short pectinate teeth basally and

medially. Propodeum extending to 0.4 of the hind

coxae, broadly and deeply depressed, sulcus for

the greater part or completely transversely wrin-

kled, costulae sometimes partly present, longitu-

dinal carinae almost completely obliterated. Pro-

podeal spiracle large, 3.3x as long as wide. Spirac-

ular carina narrow or widened. Spiracular index

0.2. 1% gastral tergite 5.1x, 2” tergite 1.9x as long

as wide. Petiole without glymmae or with small

and shallow glymmae at the anterior edge of nar-

row lateral areas, smooth in front of these areas.

Petiolar suture for the greater part obliterated.

1" gastral sternite smooth. Thyridium oval. Thy-

ridial index about 1.5. Epipleurum not separated

from the 3" gastral tergite, the latter with distinct

black lateral stripes well above the ventrolateral

edge. Gaster distinctly compressed posteriorly.

Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellowish

brown. Mandible narrowly marked with yellow

dorsally and medially. Tegula dark brown. Wings

hyaline. Stigma light brown medially. Front leg

from the femur onwards yellowish red, the femur

sometimes marked with black basally. Mid femur

yellowish brown on the apical 0.2. Mid tibia yel-

lowish brown. Mid tarsus dark brown. Hind tibia

yellowish brown medially, narrowly black basal-

ly, broadly black apically. 2" gastral tergite red on

the posterior 0.2-0.3. 3° and 4" tergites red, the 4"

tergite sometimes marked with black posteriorly.

Pilosity whitish.

d: Flagellum with 46-50 segments. Petiolar

suture complete. Mandible broadly yellow medi-

ally. Tegula yellow. Front leg from the tip of the

coxa onwards yellow. Mid femur completely yel-

lowish red. Hind tibia light yellow medially. In

other characters similar to the 2.

Host: Hypomecis punctinalis (Scopoli, 1763) (Geo-

metridae) (MIN).

Material (19??, 733): Russia: Altaysk Terr.: Vol-

chikha (MIN); Buryatiya: Zakamensk; Chita Prov.:

Chita, Kalga, Klichka, Kurort Darasun, Nerchinskiy

Zavod; Amur Prov.: Novorossiyka (PET). Kazakhstan:

Alma-Ata (MOS), Kokchetav (PET), Koktuma (LIN).

Kirgiziya: River Ala-Archa in Kirgizskiy Mts., 1600 m

(LIN).

156

Dusona specularis Horstmann, spec. nov.

Holotype (3): Russia, Khabarovsk Terr., Khekh-

tsir, near Khabarovsk, 4.7.1983 (PET).

Description. d: Body length 9mm. Temples

scarcely constricted behind the eyes, as long as the

width of an eye, tangential lines on eyes and tem-

ples intersect on the gaster. Ocellar index 1.1. Frons

scarcely depressed, coriaceous, dull, densely punc-

tate, with a fine median longitudinal carina, with

short radial wrinkles, which extend to 0.3 the

distance to the median ocellus. Antennal carina

low and narrow. Face and dorsal half ofthe clypeus

coriaceous, densely rugose-punctate. Clypeus

flat, finely coriaceous and rather densely punctate

ventrally, its apex truncate medially. Malar space

index 0.4. Genal carina low and straight ventrally,

joining the slightly raised oral carina at an acute

angle. Genal index 0.4. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

ellım with 40 segments, filiform, the apical 0.2

slightly attenuate, 2"? segment 2.5x, shortest seg-

ment 1.0x, penultimate segment 1.0x as long as

wide. Pronotum coarsely punctate on a finely coria-

ceous and shining background laterally on the

dorsal 0.3, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, densely

punctate. Scutellum shining, with lateral carinae

on the anterior 0.2. Prescutellar groove with longi-

tudinal striae. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina present anteriorly. Mesopleu-

rum with disperse or rather dense punctation on a

finely coriaceous and strongly shining background.

Speculum for the greater part smooth. Depression

in front of the speculum with strong and rather

short longitudinal striae, which merge with a nar-

row finely coriaceous area without punctation

anteriorly. Metapleurum densely rugose-punctate,

the wrinkles smooth. Pleural carina strong. Areo-

let rather small, long-stalked, receiving the 2"

recurrent vein distinctly distad of the middle.

Nervellus distinctly inclivous, distinctly intercept-

ed, its upper branch distinctly bent inwards. Ner-

vellar index 0.8. Tibial spur index 0.6. Hind femur

4.8x as long as high. Hind tarsal claws short, with

a few long pectinate teeth basally. Propodeum

extending to 0.4 of the hind coxae, rather narrow-

ly and not deeply depressed, the longitudinal fur-

row transversely wrinkled, with coarse irregular

wrinkles beside the furrow, almost completely

areolated, but the carinae partly covered with wrin-

kles, longitudinal carinae obliterated anteriorly.

Propodeal spiracle small, 1.7x as long as wide.

Spiracular carina narrow. Spiracular index 1.0.

1* gastral tergite 3.7x, 2" tergite 2.0x as long as

wide. Petiole with rather small and elongate glym-

mae, with longitudinal striae in front of the glym-

mae and with short transverse striae anteriorly.

Petiolar suture distinct. 1" gastral sternite smooth.

Thyridium oval. Thyridial index about 2.0. Epi-

pleurum separated from the 3" gastral tergite, the

crease marked with black. Epipleural index 0.7.

Gaster distinctly compressed posteriorly.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible broad-

ly yellow medially. Wings slightly infuscate. Stig-

ma dark brown medially. Front leg from the tro-

chanter onwards yellowish brown. Mid femur

yellowish red on the apical 0.2. Mid tibia and

tarsus yellowish red. Hind tibia yellowish brown

medially, narrowly black basally, rather broadly

dark brown apically. Hind tarsus dark brown.

2X sastral tergite red on the posterior 0.1. 3° and

4" tergites marked with reddish brown laterally.

Genital clasper marked with reddish brown. Pi-

losity on the face brown, on other parts of the

body whitish.

? unknown.

Dusona spinator Hinz & Horstmann, spec. nov.

Types. Holotype (?): Russia, Khabarovsk, 1.7.

1985 (PET). — Paratypes: 1? Russia, Primorye,

Spassk Dalniy, 30.6.1985 (PET); 1? Russia, Yev-

reyskaya A.O., Amurzet, Amur River, 18.6.1985

(Hinz/ZSM); 2393 Russia, Primorye, Spassk Dal-

niy, 26.-27.6.1985 (PET); 258 Russia, Khabarovsk

Terr., Khekhtsir, near Khabarovsk, 3.-4.7.1983 (18

PET, 18 Hinz/ZSM); 18 Russia, Kuril Isl., Kuna-

shir, Golovino, 10.6.1973 (PET); 18 Japan, Hon-

shu, Nagano, 400 m, 20.5.1962 (OTT).

Description. 2: Body length 10 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 1.1. Frons slightly depressed, coriaeous,

finely and densely punctate, with rather long ra-

dial wrinkles, which extend to half the distance to

the median ocellus, the median longitudinal cari-

na indistinct or obliterated. Antennal carina low

and narrow. Face and dorsal half of the clypeus

coriaceous, densely rugose. Clypeus flat, coria-

ceous and rather densely or densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.4. Genal carina low and rather straight ven-

trally, joining the slightly raised oral carina at an

acute angle. Genal index 0.4. Temple finely coria-

ceous and with fine disperse punctation medially.

Flagellum with 40-41 segments, filiform, the api-

cal 0.2 slightly attenuate, 2"X segment 2.3x, short-

est segment 0.8-0.9x, penultimate segment 1.0-1.1x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.2, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, very densely punctate. Scutellum with

lateral carinae on the anterior 0.2. Prescutellar

groove with distinct short longitudinal striae. Pre-

pectal carina low, ventral part merging with the

pleural part, the latter complete, joining the ante-

rior edge of the mesopleurum. Transverse carina

obliterated. Mesopleurum distinctly coriaceous,

densely punctate. Speculum coriaceous, partly

shining. Depression in front of the speculum with

distinct short somewhat irregular wrinkles, which

merge with the punctation anteriorly. Metapleu-

rum coriaceous, densely punctate. Pleural carina

distinct. Areolet large, short-stalked, receiving the

2‘ recurrent vein distinctly distad of the middle.

Nervellus vertical or slightly inclivous, distinctly

intercepted. Nervellar index 0.7. Tibial spur index

0.7. Hind femur 5.1x as long as high. Hind tarsal

claws rather long, with long pectinate teeth al-

most over their total length. Propodeum extend-

ing to 0.4 of the hind coxae, broadly and not

deeply depressed, sulcus for the greater part trans-

versely wrinkled, almost without carinae in the

holotype, longitudinal carinae present medially

and posteriorly in the paratypes. Propodeal spira-

cle rather large, 3.4x as long as wide. Spiracular

carina obliterated. Spiracular index 0.3. 1% gastral

tergite 3.1x, 2" tergite 1.3x as long as wide. Peti-

ole with large glymmae, which are bordered by

longitudinal carinae dorsally, with fine or very

fine longitudinal striae in front of the glymmae.

Petiolar suture complete. 1“ gastral sternite for the

greater part coriaceous and dull, with transverse

striae posteriorly. Thyridium oval. Thyridial in-

dex about 1.0. Epipleurum separated from the 3'‘

gastral tergite, the crease marked with black. Epi-

pleural index 0.8. Gaster distinctly compressed

posteriorly. Ovipositor index 1.1.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown apically. Mandible nar-

rowly marked with yellow dorsally and medially.

Tegula brown. Wings hyaline. Stigma yellowish

brown medially. Front femur yellowish brown,

somewhat darkened basally. Front tibia and tar-

sus yellow. Mid femur yellowish brown on the

apical 0.3. Mid tibia and tarsus yellow. Hind tibia

yellow or yellowish brown medially, narrowly

black basally, broadly brown or black apically.

157

2"d gastral tergite red on the posterior 0.2. 3“ and

4" tergites red. Sometimes the 5'" tergite marked

with red anteriorly. Pilosity whitish.

d: Body length 9-10 mm. 40-44 flagellar seg-

ments. Propodeum with almost complete longitu-

dinal carinae. Maxillary palpus yellow. Tegula

yellow. Front trochanter marked with yellow. Mid

femur almost completely yellowish red. Hind tib-

ia yellow medially, narrowly black basally, broad-

ly medium brown apically. Hind tarsus yellowish

brown to medium brown. 4" gastral tergite some-

times marked with black dorsally. In other charac-

ters similar to the ®.

Dusona spinipes (Thomson, 1887)

Description. 2: Body length 11-12 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.8. Frons slightly depressed,

coriaceous, finely and rather densely punctate,

with a distinct median longitudinal carina, with-

out radial wrinkles. Antennal carina low and nar-

row. Face and dorsal 0.8 of the clypeus densely

rugose, partly with vertical wrinkles. Clypeus flat,

finely coriaceous and densely punctate ventrally,

its apex truncate medially. Malar space index 0.5.

Genal carina low and straight ventrally, joining

the slightly raised oral carina at an acute angle.

Genal index 0.4. Temple finely coriaceous and

with very fine disperse punctation medially. Flag-

ellum with 48-50 segments, filiform, the apical 0.3

distinctly attenuate, 2” segment 1.8x, shortest seg-

ment 1.0x, penultimate segment 1.3x as long as

wide. Pronotum coriaceous and densely rugose-

punctate and rugose laterally on the dorsal 0.2,

with longitudinal wrinkles ventrally. Mesoscu-

tum and scutellum coriaceous, dull, very densely

punctate. Scutellum with lateral carinae on the

anterior 0.4. Prescutellar groove smooth. Ventral

part ofthe prepectal carina merging with the trans-

verse carina, both carinae low. Pleural part of the

prepectal carina obliterated. Pleural part of the

prepectus rugose-punctate and with oblique wrin-

kles. Mesopleurum with disperse or rather dense

punctation on a finely coriaceous and shining back-

ground. Speculum finely granulate-strigose. De-

pression in front of the speculum with longitudi-

nal striae and wrinkles, which merge with the

punctation anteriorly. Metapleurum densely ru-

gose. Pleural carina distinct. Areolet large, point-

ed, receiving the 2”! recurrent vein slightly basad

of the middle. Nervellus vertical, distinctly inter-

cepted, the upper branch slightly bent inwards.

Nervellar index 0.8. Tibial spur index 0.7. Hind

femur 5.7x as long as high. Hind tarsal claws

short, with distincet pectinate teeth basally and

medially. Propodeum extending to 0.4 of the hind

coxae, rather broadly and shallowly depressed,

sulcus with short transverse wrinkles anteriorly

and medially in the midline, irregularly wrinkled

sublaterally, with long transverse wrinkles poste-

riorly, carinae almost completely obliterated, some-

times costulae complete. Propodeal spiracle large,

2.8x as long as wide. Spiracular carina narrow.

Spiracular index 0.3. 1 gastral tergite 4.6x, 2”4

tergite 2.5x as long as wide. Petiole without glym-

mae or with very small glymmae at the anterior

edge of narrow lateral areas, smooth in front of

these areas. Petiolar suture fine, partly obliterat-

ed. 1“ gastral sternite smooth. Thyridium elon-

gate. Thyridial index about 1.0. Epipleurum not

separated from the 3" gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Tegula dark

brown. Wings hyaline. Stigma dark brown medi-

ally. Front leg from the trochanter onwards yel-

low, the femur yellowish red. Tip of the front coxa

sometimes yellow. Mid femur red on the apical

0.2-0.5. Mid tibia yellow. Mid tarsus brown. Hind

tibia dark brown or blackish, narrowly black ba-

sally. Gastral tergites black dorsally. 2" tergite

laterally red on the posterior 0.5. 3° tergite broad-

ly red laterally. 4" tergite about half red or brown

in lateral view. Posterior tergites narrowly marked

with brown ventrolaterally. Posterior gastral ster-

nites brown. Pilosity whitish.

d: Propodeum scarcely depressed, costulae

complete. 2"! and 4" gastral tergites almost com-

pletely red laterally. Genital clasper reddish brown.

In other characters similar to the 2.

Material (112%, 838): Russia: Novosibirsk Prov.:

Kourak (PET); Altaysk Terr.: Gorno-Altaysk (MIN);

Irkutsk Prov.: Dachnaya; Chita Prov.: Adrianovka,

Bukuka; Khabarovsk Terr.: Khekhtsir near Khaba-

rovsk; Primorye: Khasan, Posyet, Spassk Dalniy; Kuril

Isl.: Alekhino and Sernovodsk on Kunashir (PET). Ja-

pan: Hokkaido: Jozankei; Honshu: Nagano (AEI);

Amami O-shima (SAP).

Dusona spiracularis Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Russia, Primorye, Anisi-

movka, ESE of Shkotovo, 11.9.1978 (PET). — Para-

types: 1? same dates (Hinz/ZSM); 333 Russia,

Primorye, Ussuriysk, 4.-5.9.1983 (23895 PET, 1d

Hinz /ZSM).

Description. 2: Body length 6-7 mm. Temples

distinctly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.9. Frons not depressed, coriaceous, densely ru-

gose-punctate, with a fine median longitudinal

carina, with distinct radial wrinkles, which almost

reach the median ocellus. Antennal sockets close

to each other. Antennal carina low, widened to a

crescent-shaped plate, the latter with fine trans-

verse striae. Face and dorsal 0.6 of the clypeus

densely rugose. Clypeus flat, coriaceous and rath-

er densely punctate ventrally, its apex slightly

convex medially. Malar space index 0.4. Genal

carina low ventrally, slightly bent sidewards, join-

ing the slightly raised oral carina almost at a right

angle. Genal index 0.1. Temple finely coriaceous

and with fine disperse punctation medially. Flag-

ellum with 29 segments, slightly fusiform, 2" seg-

ment 2.1x, shortest segment 0.9x, penultimate seg-

ment 1.2x as long as wide. Pronotum densely

rugose laterally on the dorsal 0.1, with longitudi-

nal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, densely punctate. Scutellum with

lateral carinae on the anterior half. Prescutellar

groove with fine longitudinal striae. Ventral part

ofthe prepectal carina low, merging with the trans-

verse carina, the latter slightly raised. Pleural part

of the prepectal carina low, the dorsal 0.3 obliter-

ated. Mesopleurum with dense or very dense

punctation on a very slightly sculptured back-

ground, partly rugose or rugose-punctate. Specu-

lum with fine longitudinal striae. Depression in

front of the speculum with with strong longitudi-

nal striae, which merge with the punctation ante-

riorly. Metapleurum densely and rather coarsely

rugose. Pleural carina fine. Areolet rather small,

pointed or short-stalked, receiving the 2"? recur-

rent vein distinctly distad of the middle. Nervel-

lus slightly inclivous, intercepted. Nervellar in-

dex 0.8. Tibial spur index 0.6. Hind femur 5.2x as

long as high. Hind tarsal claws short, with few

pectinate teeth basally. Propodeum extending to

0.6 of the hind coxae, rather narrowly and not

deeply depressed, für the greater part coarsely

and irregularly wrinkled, with short transverse

wrinkles in the longitudinal furrow, costulae com-

plete, longitudinal carinae obliterated. Propodeal

spiracle small, 2.0x as long as wide. Spiracular

carina narrow. Spiracular index 0.5. 1“ gastral

tergite 5.2x, 2" tergite 1.9x as long as wide. Peti-

ole without glymmae, with narrow and some-

what depressed lateral areas, smooth in front of

the areas. Petiolar suture fine. 1* gastral sternite

for the greater part smooth, with transverse striae

posteriorly. Thyridium elongate. Thyridial index

1.3. Epipleurum not separated from the 3 gastral

tergite, the latter without or with short black later-

al stripes. Gaster distinctly compressed posterior-

ly. Ovipositor index 1.0.

Colour: Black. Maxillary palpus dark brown.

Mandible narrowly marked with dark brown me-

dially. Tegula dark brown. Wings hyaline. Stigma

dark brown medially. Front leg from the trochan-

tellus onwards and mid leg from the femur on-

wards yellow or yellowish red. Hind femur yel-

lowish red on the apical 0.2. Hind tibia yellowish

brown, narrowly brown basally. Hind tarsus dark

brown. 2" gastral tergite red on the posterior 0.4.

3’ and 4" tergites red. Pilosity whitish.

d: Flagellum with 33-34 segments, filiform,

shortest segment 1.0x as long as wide. Maxillary

palpus yellowish brown. Mandible broadly marked

with yellow medially. Front leg from the tip of the

coxa onwards yellow or yellowish red. Hind fe-

mur sometimes completely black. 3" gastral tergite

with distinct black lateral stripes. In other charac-

ters similar to the 2.

Dusona splenditor Hinz, 1985

Description. 2: Body length 9-11 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.5. Frons slightly depressed, coriaceous

and densely punctate, with a fine median longitu-

dinal carina, with fine radial wrinkles, which ex-

tend to 0.3-0.5 the distance to the median ocellus.

Antennal carina low and narrow. Face coriaceous,

densely rugose-punctate. Clypeus slightly con-

cave, coriaceous, densely punctate, with a narrow

less densely punctate area ventrally, the apex

slightly convex or truncate medially. Malar space

index 0.3. Genal carina strongly raised ventrally,

slightly bent sidewards, jeining the strongly raised

oral carina at an acute angle, both carinae forming

a lopsided triangular pyramid, the tip of the latter

shifted ventrally and situated behind the base of

the mandible. Gena depressed close to the junc-

tion, the depression about half as deep as the

width of the 2" flagellar segment. Genal index

159

about 0.3. Temple finely coriaceous and with very

fine disperse punctation medially. Flagellum with

47-49 segments, filiform, the apical 0.2 distinctly

attenuate, 2"! segment 1.6x, shortest segment 0.8x,

penultimate segment 1.3x as long as wide. Prono-

tum very densely punctate laterally on the dorsal

0.3, with longitudinal wrinkles ventrally. Meso-

scutum coriaceous, very densely punctate. Scutel-

lum very densely punctate on a partly smooth

background, with lateral carinae on the anterior

0.7. Prescutellar groove smooth. Ventral part of

the prepectal carina either merging with the pleu-

ral part or with the transverse carina, the latter

complete, not raised. Pleural part of the prepectal

carina low, its dorsal 0.1-0.3 obliterated or re-

placed by several wrinkles. Mesopleurum with

disperse or rather dense punctation on a smooth

or very finely coriaceous background. Speculum

smooth or almost smooth, strongly shining. De-

pression in front of the speculum with distinct

and rather long longitudinal striae, which merge

with the punctation anteriorly. Metapleurum with

longitudinal wrinkles dorsally, densely punctate

on a finely coriaceous or smooth background

ventrally. Pleural carina strong. Areolet large, nar-

rowly sessile or pointed, receiving the 2" recur-

rent vein at or slightly distad of the middle. Nervel-

lus slightly inclivous, distinctly intercepted. Ner-

vellar index 0.7. Tibial spur index 0.7. Hind femur

4.5x as long as high. Hind tarsal claws short, with

distinct pectinate teeth almost over their total

length. Propodeum extending to 0.6 of the hind

coxae, broadly and shallowly depressed, sulcus

with long transverse wrinkles, costulae obliterat-

ed, longitudinal carinae sometimes indicated pos-

teriorly and medially. Propodeal spiracle large,

3.1x as long as wide. Spiracular carina partly wid-

ened. Spiracular index 0.5. 1% gastral tergite 4.5x,

2"d tergite 2.1x as long as wide. Petiole without

glymmae or with small and shallow glymmae at

the anterior edge of narrow lateral areas, smooth

in front of these areas. Petiolar suture partly oblit-

erated. 1" gastral sternite smooth. Thyridium elon-

gate. Thyridial index about 1.5. Epipleurum not

separated from the 3° gastral tergite, the latter

without or with black lateral stripes well above

the ventrolateral edge. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.9.

Colour: Black. Maxillary palpus yellowish red.

Mandible broadly yellow medially. Flagellum light

brown apically. Tegula yellow. Wings hyaline.

Stigma light brown medially. Front leg from the

tip of the coxa onwards yellow. Mid femur black

basally and ventrally, yellow dorsally and apical-

ly. Mid tibia yellow. Mid tarsus yellowish brown.

Hind tibia yellow, very narrowly dark brown ba-

160

sally and apically, the apical dark mark often lack-

ing. Hind tarsus dark brown. 2" gastral tergite

yellowish red on the posterior 0.3. 3" tergite yel-

lowish red. 4" tergite yellowish red anteriorly (to

a variable extend). Pilosity whitish.

d: Mid femur yellow. In other characters sim-

ilar to the 2.

Material (222, 18): Kazakhstan: Kharkin near Kal-

mykovo (Hinz/ZSM, PET): China: Heilongjiang Prov.:

Charbin (= Harbin) (AEI). The species also occurs in

Europe: 27? Russia, Volgograd (PET).

Dusona stragifex (Förster, 1868)

This species is very similar to D. bicoloripes and

D. chabarowski, particularly to those specimens in

which the raised points between the antennal sock-

ets and the eyes are obliterated. In D. stragifex the

impression in front of the speculum is distinctly

striate, the propodeum is distinctly depressed, the

longitudinal carinae are usually present medially

and posteriorly, and the sulcus is more or less

transversely striate medially and posteriorly. But

some specimens, in particular dd, may be indeter-

minable, and some specimens listed below may

be misidentified. D. stragifex is also similar to D. ru-

gifer (see remark under this name).

Description. 2: Body length 10-13 mm. Temples

strongly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and densely punctate, with a

fine and partiy obliterated median longitudinal

carina, without or with short radial wrinkles. An-

tennal carina low and narrow. Face and dorsal

half of the clypeus coriaceous, densely rugose-

punctate. Clypeus flat, finely coriaceous and with

rather dense punctation ventrally, its apex slight-

ly convex medially. Malar space index 0.3. Genal

carina low ventrally, straight or slightly bent side-

wards, joining the slightly raised oral carina at an

acute angle. Genal index 0.5. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 49-52 segments, filiform,

the apical 0.3 distinctly attenuate, 2"! segment

1.5x, shortest segment 0.9x, penultimate segment

1.2x as long as wide. Pronotum with longitudinal

wrinkles laterally, with additional punctures on

the dorsal 0.3. Mesoscutum and scutellum coria-

ceous, dull, densely or very densely punctate.

Scutellum with lateral carinae on the anterior 0.2.

Prescutellar groove smooth medially, with longi-

tudinal striae laterally. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

complete, joining the anterior edge of the meso-

pleurum, sometimes the dorsal 0.1 replaced by

several wrinkles. Transverse carina indistinct.

Mesopleurum distinctly coriaceous, rather dense-

ly or densely punctate. Speculum finely coria-

ceous, shining. Depression in front of the specu-

lum with distinct longitudinal wrinkles or striae,

which merge with the punctation anteriorly. Meta-

pleurum coriaceous, finely rugose-punctate. Pleu-

ral carina distinct. Areolet large, short-stalked,

receiving the 2" recurrent vein at or basad of the

middle. Nervellus almost vertical, distinctly inter-

cepted. Nervellar index 0.7-0.8. Tibial spur index

0.7. Hind femur 4.9x as long as high. Hind tarsal

claws short, with distinct pectinate teeth basally

and medially. Propodeum extending to 0.3 of the

hind coxae, distinctly and broadly but not deeply

depressed, sulcus usually for the greater part trans-

versely wrinkled, often with a short longitudinal

keel posteriorly, costulae partly present, longitu-

dinal carinae present medialy and posteriorly,

sometimes almost complete. Propodeal spiracle

large, 2.8x as long as wide. Spiracular carina nar-

row. Spiracular index 0.6. 1* gastral tergite 3.8x,

2" tergite 1.6x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal

carinae dorsally, smooth in front of the glymmae.

Petiolar suture fine, sometimes partly obliterated.

1“ gastral sternite coriaceous anteriorly, smooth

or finely coriaceous medially, transversely striate

posteriorly. Thyridium oval. Thyridial index about

1.0. Epipleurum separated from the 34 gastral

tergite, the crease marked with black. Epipleural

index 0.6-0.7. Gaster distinctly compressed poste-

riorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish red

to brown. Mandible broadly yellow medially.

Tegula dark brown. Wings hyaline. Stigma medi-

um brown medially. Front leg from the trochanter

onwards yellowish red. Mid femur and tibia yel-

lowish red, the femur often marked with black

basally. Hind tibia broadly yellowish brown to

brown medially, narrowly dark brown or black

basally and apically, the apical dark mark some-

times lacking. 2” gastral tergite red on the poste-

rior 0.4-0.5. 3” and 4" tergites red, the 4" tergite

sometimes marked with black dorsally (to a vari-

able extend). Pilosity whitish.

d: Tegula sometimes yellow. Front and mid

legs sometimes from the tip of the coxa onwards

yellow or yellowish red. 2”! gastral tergite some-

times black dorsally and 4* tergite sometimes com-

pletely black. In other characters similar to the ®.

Material (35?%, 839): Russia: Sverdlovsk Prov.: Ye-

katerinburg (= Sverdlovsk); Tomsk Prov.: Tomsk

(PET); Altaysk Terr.: Artybash (MIN); Buryatiya: Riv-

er Dzhida; Khabarovsk Terr.: Khekhtsir near Khaba-

rovsk, Slavyanka; Primorye: Khasan, Roshchino on

River Iman, Spassk Dalniy, Ussuriysk; Sakhalin: Lof-

tsi; Kuril Isl.: Dubovoye and Volcano Golovnina on

Kunashir (PET). South Korea: Suigen (= Suwon) (AEI).

Japan: Hokkaido: Asahikawa (PET), Maruyama (SAP),

Sapporo (Hinz/ZSM), Teshio; Honshu: Kyoto, Sasa-

yama (SAP), Tokyo (OTT), Yokohama (AEI); Shikoku:

Kagawa (SAP).

Variety: 2" gastral tergite dorsally red on the

posterior 0.2, laterally red on the posterior 0.4. 3"

and 4" tergites narrowly black dorsally, broadly

red laterally. 5" tergite broadly black dorsally,

half red in lateral view. 6" and 7" tergites narrow-

Iy marked with red ventrolaterally. Gastral ster-

nites for the greater part red. Genital clasper black.

In other characters similar to the nominate mor-

phe. Material: 283 Russia, Primorye, Roshchino

on River Iman (PET). The status of this variety is

unknown.

Dusona stygia (Förster, 1868)

Description. 2: Body length 10-11 mm. Temples

slightly constricted behind the eyes, 0.9-1.0x as

long as the width of an eye, tangential lines on

eyes and temples intersect on the posterior edge

of the propodeum. Ocellar index 1.2. Frons slight-

ly depressed above the antennal carinae, coria-

ceous and densely rugose-punctate, with a fine

median longitudinal carina, with short radial wrin-

kles. Antennal carina low, with a more or less

distinct dorsal projection (nose-like), the latter with

fine transverse striae. Face and dorsal 0.4 of the

clypeus coriaceous, dull, densely rugose-punctate.

Clypeus flat, coriaceous and rather densely punc-

tate ventrally, its apex truncate medially. Malar

space index 0.5. Genal carina low ventrally, slight-

ly bent sidewards, joining the distinctly raised

oral carina at an acute angle. Genal index 0.4.

Temple coriaceous and with fine rather dense

punctation medially. Flagellum with 44-48 seg-

ments, filiform, the apical 0.2 slightly attenuate,

2"! segment 1.5x, shortest segment 0.9x, penulti-

mate segment 1.1x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.4-0.6, with longitudinal wrinkles ventrally. Meso-

scutum coriaceous, very densely punctate. Scutel-

lum very densely punctate on a smooth or finely

coriaceous background, with lateral carinae on

the anterior 0.5-0.7. Prescutellar groove smooth

medially, with longitudinal striae laterally. Pre-

161

pectal carina low, ventral part merging with the

pleural part, the dorsal 0.2-0.3 of the latter obliter-

ated or replaced by several wrinkles. Transverse

carina complete or partly obliterated. Mesopleu-

rum distinctly coriaceous, distinctly and densely

punctate. Speculum punctate posteriorly, granu-

late-strigose and rather dull anteriorly. Depres-

sion in front of the speculum with with short

longitudinal wrinkles, which merge with the punc-

tation anteriorly. Metapleurum coriaceous, densely

rugose-punctate. Pleural carina fine, often partly

indistinct and covered with wrinkles. Areolet large,

pointed, receiving the 2" recurrent vein distad of

the middle. Nervellus vertical, weakly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.7. Hind

femur 4.7x as long as high. Hind tarsal claws long,

with distinct pectinate teeth basally and medially.

Propodeum extending to 0.3 of the hind coxae,

broadly and shallowly depressed, sulcus irregu-

larly wrinkled anteriorly, with long transverse

wrinkles posteriorly, carinae for the greater part

obliterated, sometimes some parts weakly indi-

cated. Propodeal spiracle small, 2.3x as long as

wide. Spiracular carina obliterated. Spiracular in-

dex 0.9-1.1. 1* gastral tergite 3.6x, 2”! tergite 1.3x

as long as wide. Petiole without glymmae, with

indistinct narrow lateral areas, smooth in front of

these areas. Petiolar suture fine. 1* gastral sternite

for the greater part smooth or with very fine sculp-

ture, with transverse striae posteriorly. Thyrid-

ium oval or elongate. Thyridial index about 1.0.

Epipleurum not separated from the 34 gastral

tergite, sometimes discernible as a small triangu-

lar lobe anteriorly, 3 tergite with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.6.

Colour: Black. Mandible sometimes narrow-

ly marked with reddish brown medially. Wings

hyaline. Stigma yellowish brown medially. Front

femur black basally and ventrally, yellowish brown

dorsally and apically. Front tibia yellowish brown.

Front tarsus dark brown. Mid leg blackish, the

femur narrowly brown apically, the tibia narrow-

ly brown basally. Hind tibia sometimes tinged

with brown subbasally. 2" gastral tergite red on

the posterior 0.4-0.6. 3"! and 4" tergites red, some-

times narrowly marked with black. 5!" tergite some-

times marked with red anteriorly. Pilosity whit-

ish.

d: Propodeum almost not depressed. In other

characters similar to the 9.

Material (234, 12): Iran: Takht-i-Suleiman Mts., 2000-

2500 m (WAR). Turkmeniya: Kopet-Dag Mts. (PET).

162

Dusona subimpressa (Förster, 1868)

Description. ?: Body length 7-9 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the middle of the mesoscu-

tum. Ocellar index 0.7. Frons slightly depressed,

coriaceous, finely and rather densely punctate,

with a fine median longitudinal carina, without

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.7 of the clypeus coriaceous,

densely rugose-punctate and rugose. Clypeus flat,

finely coriaceous and rather densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.4. Genal carina low and almost straight ven-

trally, joining the slightly raised oral carina at an

acute angle. Genal index 0.5. Temple finely coria-

ceous and with very fine disperse punctation me-

dially. Flagellum with 39-41 segments, filiform,

the apical 0.3 slightly attenuate, 2° segment 2.2x,

shortest segment 0.9x, penultimate segment 1.0x

as long as wide. Pronotum coriaceous and dense-

ly rugose-punctate laterally on the dorsal 0.3, with

longitudinal wrinkles ventrally. Mesoscutum co-

riaceous, dull, very densely punctate. Scutellum

finely coriaceous, dispersely or rather densely

punctate, with lateral carinae on the anterior half.

Prescutellar groove with fine longitudinal striae.

Prepectal carina low, ventral part usually merg-

ing with the pleural part, rarely with the trans-

verse carina. Pleural part of the prepectal carina

present ventrally, the dorsal 0.4 obliterated, some-

times the pleural part completely obliterated and

replaced by wrinkles. Transverse carina distinct

or obliterated. Mesopleurum distinctly coriaceous,

densely punctate. Speculum coriaceous, dull. De-

pression in front of the speculum with distinct

rather short longitudinal wrinkles, which merge

with the punctation anteriorly. Metapleurum

densely rugose. Pleural carina distinct. Areolet

rather large, pointed or short-stalked, receiving

the 2"d recurrent vein at or basad of the middle.

Nervellus slightly reclivous, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 5.2x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to the middle of

the hind coxae, scarcely depressed, for the greater

part irregularly wrinkled, often with some longi-

tudinal wrinkles posteriorly, rarely with few short

transverse wrinkles, costulae fine, sometimes al-

most complete, longitudinal carinae for the great-

er part obliterated. Propodeal spiracle large, 3.0x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.4. 1% gastral tergite 5.0x, 2"! tergite

2.3x as long as wide. Petiole for the greater part

smooth laterally, glymmae lacking, narrow lateral

areas sometimes indicated. Petiolar suture oblit-

erated or partly indicated. 1“ gastral sternite

smooth, sometimes with fine transverse striae

posteriorly. Thyridium elongate. Thyridial index

about 1.5. Epipleurum not separated from the 3"4

gastral tergite, the latter with rather fine black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula medium

brown. Wings hyaline. Stigma dark brown medi-

ally. Front leg from the tip of the coxa onwards

yellowish red. Mid femur yellowish brown on the

apical 0.3. Mid tibia and tarsus yellowish brown.

Hind tibia broadly yellowish brown to brown

medially, narrowly black basally, narrowly brown

apically, the apical dark mark sometimes indis-

tinct or lacking. 2” gastral tergite black dorsally,

red laterally on the posterior 0.3-0.5. 3" and 4"

tergites for the greater part red, narrowly marked

with black dorsally, sometimes only on the 3"

tergite marked with black. 5" to 7" tergites black

dorsally, red laterally (to a variable extend). Gas-

tral sternites red. Pilosity whitish.

d: Flagellum with 40-46 segments. Pleural part

of the prepectal carina often obliterated. Impres-

sion in front of the speculum often with fine longi-

tudinal striae. Propodeum not depressed. Tegula

yellow. Front and mid legs yellow, the mid coxa

marked with black basally. Genital clasper red. In

other characters similar to the ?

Material (2622, 7188): Russia: Khabarovsk Terr.:

Khabarovsk, Khekhtsir near Khabarovsk; Primorye:

Chuguyevka, Gornotayensk (PET), Nakhodka (KIS),

Okeanskaya (PET), Partizansk (KIS), Sudzukhinskiy

(= Lazovskiy) Reserve (PET), Ussuriysk (MOS), Vladi-

vostok; Sakhalin: Novoaleksandrovsk; Kuril Isl.: Ale-

khino, Tretyakovo and Volcano Golovnina on Kuna-

shir (PET). Japan: Hokkaido: Maruyama, Sapporo,

Taisetusan (SAP); Honshu: Chigasaki, Kamikochi

near Norikura-dake (AEI), Nagano (AEI, OTT), Nori-

kura-dake (AEI); Shikoku: Tosa; Kyushu: Hiko-san

(SAP).

Variety: Flagellum with 46 segments. Mesopleu-

rum densely punctate on a smooth background.

In other characters similar to the nominate mor-

phe. Material: 1? Russia, Yevreyskiy A.O., Lon-

doko (PET). The status of this variety is unknown.

Dusona temnator Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Russia, Primorye, Kedro-

vaya Pad Reserve, near Khasan, 25.8.1963 (MOS).

- Paratypes: 322, 233 same locality, 22.8.-8.9.

1963 (2??, 18 MOS, 1%, 13 Hinz/ZSM); 12 Japan,

Hokkaido, Sapporo, 18.10.1922 (SAP).

Description. 2: Body length 9-10 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons not depressed, coriaceous, distinctly

and densely punctate, with indistinct radial wrin-

kles, without a median longitudinal carina. An-

tennal sockets close to each other. Antennal carina

low, sometimes widened to anarrow smooth cres-

cent-shaped plate. Face and clypeus finely coria-

ceous and densely punctate. Clypeus flat, its apex

truncate and with asmooth bulge medially. Malar

space index 0.4. Genal carina straight ventrally,

slightly raised, joining the distinctly raised oral

carina at an acute angle. Genal index 0.3. Temple

with fine and rather dense punctation on asmooth

background medially. Flagellum with 47-51 seg-

ments, filiform, the apical 0.3 distinctly attenuate,

2"! segment 1.9x. shortest segment 1.1x, penulti-

mate segment 1.4x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.6, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum densely punctate, very fine-

ly coriaceous and shining between the punctures.

Scutellum with lateral carinae on the anterior 0.6.

Prescutellar groove smooth. Ventral part of the

prepectal carina low, merging with the transverse

carina, the latter slightly raised. Pleural part of the

prepectal carina completely obliterated. Pleural

part of the prepectus punctate, without wrinkles.

Mesopleurum with rather dense or dense puncta-

tion, very finely coriaceous and partly smooth

between the punctures. Speculum densely punc-

tate posteriorly, very finely sculptured or smooth

anteriorly. Depression in front of the speculum

with distinct short longitudinal striae dorsally and

medially, which merge with the punctation ante-

riorly, almost smooth ventrally. Metapleurum

densely rugose. Pleural carina very fine, partly

covered with wrinkles or obliterated. Areolet large,

pointed or short-stalked, receiving the 2” recur-

rent vein at or slightly distad ofthe middle. Nervel-

lus slightly reclivous, intercepted. Nervellar in-

dex 0.6-0.7. Tibial spur index 0.6. Hind femur 5.3x

as long as high. Hind tarsal claws rather long,

with distinct pectinate teeth basally. Propodeum

163

extending to 0.4 of the hind coxae, broadly and

not deeply depressed, sulcus finely coriaceous

and with fine irregular wrinkles anteriorly, with

fine transverse wrinkles posteriorly, costulae partly

present, longitudinal carinae obliterated or indi-

cated medially and posteriorly. Propodeal spira-

cle rather large, 3.2x as long as wide. Spiracular

carina indistinct and covered with wrinkles. Spirac-

ular index about 0.8. 1* gastral tergite 5.2x, 2"d

tergite 2.2x as long as wide. Petiole without glym-

mae, with indistinct narrow lateral areas, smooth

in front of these areas. Petiolar suture partly oblit-

erated. 1“ gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium elongate. Thyridial index about 1.0. Epi-

pleurum not separated from the 3" gastral tergite,

the latter without or with short and weak black

lateral stripes. Gaster strongly compressed poste-

riorly. Ovipositor index 0.7.

Colour: Black. Wings hyaline. Stigma dark

brown medially. Front tibia with a yellow or yel-

lowish brown stripe anteriorly. 2" gastral tergite

red on the posterior 0.1. 3" tergite red. 4" tergite

black or marked with red on the anterior 0.2.

Pilosity whitish.

d: Propodeum with distinct transverse wrin-

kles, longitudinal carinae distinct medially and

posteriorly. Front femur yellowish brown anteri-

orly. Front tibia yellowish brown. Front tarsus

and mid tibia sometimes yellowish brown. In oth-

er characters similar to the ®.

Dusona templator Hinz & Horstmann,

spec. nov.

The types of this species differ somewhat from

each other in colour pattern and in the sculpture

of the frons, but are very similar to each other in

other characters.

Types. Holotype (?): Mongoliya, Central Prov.

(=Töv Prov.), Kerulen, 28.8.-3.9.1975 (PET). -

Paratypes: 1? Tadzhikistan, Shakhristan, 12.6.

1982 (Hinz / ZSM); 15 Russia, Tuvinskaya A.S.S.R.,

Kyzyl, 31.5.1975 (PET).

Description. 2: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye. tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons not depressed, coriaceous, densely ru-

gose-punctate, with a very fine median longitudi-

nal carina, with rather long radial wrinkles, which

extend to 0.5-0.8 the distance to the median ocel-

lus, the frons between the wrinkles almost smooth

164

in the holotype, dull in the 2 paratype. Antennal

carina low and narrow. Face and dorsal 0.7 of the

clypeus coriaceous, densely rugose-punctate.

Clypeus flat, coriaceous and densely punctate ven-

trally, its apex slightly convex medially. Malar

space index 0.3. Genal carina low and straight

ventrally, joining the distinctly raised oral carina

at an acute angle. Genal index 0.5. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 43-45 segments, filiform,

the apical 0.2 slightly attenuate, 2° segment 1.7x,

shortest segment 1.0x, penultimate segment 1.1x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.4, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, densely punctate. Scutellum with lat-

eral carinae on the anterior 0.7. Prescutellar groove

smooth. Prepectal carina low, ventral part merg-

ing with the pleural part, the dorsal 0.2 of the

latter obliterated or covered by wrinkles. Trans-

verse carina obliterated. Mesopleurum coriaceous,

distinctly and densely punctate. Speculum granu-

late-strigose, dull. Depression in front of the specu-

lum with longitudinal striae and short wrinkles,

which merge with the punctation anteriorly. Meta-

pleurum densely rugose. Pleural carina fine. Are-

olet large, pointed, receiving the 2" recurrent vein

slightly distad of the middle. Nervellus vertical or

slightly reclivous, weakly intercepted. Nervellar

index 0.7. Tibial spur index 0.7. Hind femur 4.9x

as long as high. Hind tarsal claws rather long,

with short pectinate teeth basally. Propodeum

extending to 0.4 of the hind coxae, scarcely de-

pressed, for the greater part irregularly wrinkled,

with short longitudinal wrinkles posteriorly, car-

inae almost completely obliterated. Propodeal

spiracle small, 3.4x as long as wide. Spiracular

carina obliterated. Spiracular index 1.2. 1“ gastral

tergite 4.1x, 2X tergite 1.6x as long as wide. Peti-

ole without glymmae, narrow lateral areas weak-

ly indicated by a coriaceous surface, smooth in

front of these areas. Petiolar suture partly obliter-

ated. 1* gastral sternite for the greater part smooth,

with transverse striae posteriorly. Thyridium oval.

Thyridial index about 1.5. Epipleurum separated

from the 3"! gastral tergite anteriorly, pendent as a

small rather triangular lobe. Epipleural index about

0.1. 34 tergite with black lateral stripes well above

the ventrolateral edge. Gaster distinctly com-

pressed posteriorly. Ovipositor index 0.6.

Colour: Holotype: Black. Mandible narrowly

marked with dark brown medially. Wings hya-

line. Stigma brownish black medially. Front leg

from the femur onwards yellowish brown. Mid

femur brown on the apical 0.1. Mid tibia brown.

2rd gastral tergite red on the posterior 0.6. 3°“ and

4" tergites red. Pilosity whitish. The ? paratype is

distinctly darker: Mandible black. Wings distinct-

ly infuscate. Front femur marked with black ven-

trally. Front tarsus dark brown. Mid leg black.

2" gastraltergite red on the posterior 0.4. 4" tergite

narrowly marked with black posteriorly. Pilosity

dark brown.

d: Frons dull between the radial wrinkles.

Maxillary palpus dark brown. Mandible narrow-

ly marked with yellowish red dorsally and medi-

ally. Front leg from the tip of the coxa onwards

yellowish red. Mid femur yellowish on the apical

0.3. Mid tibia yellowish brown. Mid tarsus dark

brown. 2" gastral tergite tergite red on the poste-

rior 0.4. In other characters similar to the holo-

type.

Variety: Flagellum with 48 segments. Temples

slightly constricted behind the eyes, tangential

lines on eyes and temples intersect on the propo-

deum. Wings slightly infuscate. Pilosity light

brown. In other characters similar to the nominate

d. Material: 18 Uzbekistan, Bukhara (PET). The

status of this male is unknown. It is somewhat

intermediary between D. stygia and D. templator.

Dusona temporalis Horstmann, spec. nov.

Types. Holotype (?): Kazakhstan, Uzun-Agach,

50 km NW Alma-Ata, 30.5.1994 (LIN). - Paratype:

19 same dates (Hinz/ZSM).

Description. 2: Body length 8mm. Temples

scarcely constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the apex of the gaster. Ocellar

index 1.0. Frons scarcely depressed, coriaceous,

densely rugose-punctate, with a very fine median

longitudinal carina, with rather long radial wrin-

kles, which extend to half the distance to the me-

dian ocellus. Antennal carina low and narrow.

Face and dorsal 0.8 of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, finely co-

riaceous and rather densely punctate ventrally, its

apex truncate medially. Malar space index 0.5.

Genal carina very low ventrally, almost obliterat-

ed, slightly bent forwards, joining the slightly

raised oral carina close to the base of the mandible

almost at a right angle. Genal index 0.1 or less

(difficult to estimate). Temple finely coriaceous

and with distinct dense punctation medially. Flag-

ellum with 43-44 segments, filiform, the apical 0.2

attenuate, 2" segment 2.1x, shortest segment 1.1x,

penultimate segment 1.4x as long as wide. Prono-

tum densely rugose-punctate on a smooth back-

ground laterally on the dorsal 0.4, with longitudi-

nal wrinkles ventrally. Mesoscutum coriaceous,

densely punctate. Scutellum with rather dense or

dense punctation on an almost smooth back-

ground, with lateral carinae on the anterior half.

Prescutellar groove smooth medially, with longi-

tudinal striae laterally. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

almost joining the anterior edge of the mesopleu-

rum. Transverse carina obliterated. Mesopleurum

distinctly and densely punctate on a very finely

coriaceous and shining background. Speculum

finely coriaceous dorsally, almost smooth ventral-

ly. Depression in front of the speculum with short

irregular wrinkles dorsally, granulate-strigose and

with fine longitudinal striae ventrally, the latter

merging with the punctation anteriorly. Meta-

pleurum finely coriaceous, densely punctate. Pleu-

ral carina complete. Areolet rather small, short-

stalked, receiving the 2” recurrent vein at the

middle. Nervellus reclivous, weakly intercepted.

Nervellar index 0.7. Tibial spur index 0.6. Hind

femur 4.8x as long as high. Hind tarsal claws

rather long, with short pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

deeply and rather narrowly depressed, with short

transverse striae in the longitudinal furrow, with

irregular wrinkles beside the furrow, carinae al-

most completely obliterated. Propodeal spiracle

rather small, 3.5x as long as wide. Spiracular car-

ina narrow. Spiracular index 0.5. 1* gastral tergite

4.4x, 2"4 tergite 1.6x as long as wide. Petiole with-

out glymmae, with weak narrow lateral areas,

smooth in front of these areas. Petiolar suture

almost completely obliterated. 1“ gastral sternite

smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum not separated from the 3"

gastral tergite, the latter with distinct black lateral

stripes well above the ventrolateral edge. Gaster

not much compressed posteriorly. Ovipositor in-

dex0.D.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly marked with yellow dorsally

and medially. Tegula yellow. Wings hyaline. Stig-

ma yellowish brown medially. Front leg from the

trochanter onwards yellowish red, the trochanter

marked with black. Mid femur and tibia yellow-

ish red, the femur marked with black basally. Mid

tarsus brownish black. Hind femur narrowly

marked with red subapically. Hind tibia ventrally

reddish brown, dorsally dark yellowish brown

basally, dark brown apically. 2”! gastral tergite

red on the posterior 0.4. 3” and 4" tergites red.

Pilosity whitish.

d: Temples widened behind the eyes. Flagel-

lum with the 2" segment 1.8x as long as wide.

165

Depression in front of the speculum for the great-

er part coriaceous and granulate-strigose. Maxil-

lary palpus yellow. Mandible for the greater part

yellow (teeth black). Front and mid legs from the

coxae onwards yellow or yellowish red, the base

of the coxae black. Hind femur broadly marked

with red subapically. Hind tibia ventrally red,

dorsally yellowish basally and medially, blackish

apically. In other characters similar to the 9.

Dusona tenuis (Förster, 1868)

Description. 2: Body length 8mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons scarcely de-

pressed, coriaceous, finely and densely rugose-

punctate, with a very fine and partly obliterated

median longitudinal carina, with short radial wrin-

kles. Antennal carina low and narrow, sometimes

slightly raised (inconspicuous). Face and dorsal

0.3 ofthe clypeus coriaceous, densely rugose-punc-

tate and rugose. Clypeus flat, coriaceous and

densely punctate ventrally, its apex truncate me-

dially. Malar space index 0.3. Genal carina low

ventrally, distinctly bent sidewards and forwards

(Fig. 9 and 14), more or less obliterated close to the

junction with the distinctly raised oral carina. Gena

slightly depressed close to the junction. Genal

index about 0.3. Temple finely coriaceous and

with fine disperse punctation medially. Flagellum

with 44-48 segments, filiform, the apical 0.3 dis-

tinctly attenuate, 2” segment 1.8x, shortest seg-

ment 1.0x, penultimate segment 1.3x as long as

wide. Pronotum densely rugose-punctate and ru-

gose laterally on the dorsal 0.3, with longitudinal

wrinkles ventrally. Mesoscutum coriaceous, dull,

very densely punctate. Scutellum coriaceous, dull,

with rather dense or dense punctation, with later-

al carinae on the anterior 0.6. Prescutellar groove

smooth medially, with longitudinal striae lateral-

ly. Ventral part of the prepectal carina merging

with the transverse carina, both carinae strongly

raised, the former distinctly emarginate medially.

Pleural part of the prepectal carina low, its dorsal

0.4 obliterated (Fig. 19). Mesopleurum finely co-

riaceous, densely punctate. Speculum granulate-

strigose, rather dull. Depression in front of the

speculum with distinct striae, which merge with

the punctation anteriorly. Metapleurum finely and

densely rugose-punctate. Pleural carina distinct.

Areolet rather large, short-stalked, receiving the

2"! recurrent vein distad of the middle. Nervellus

slightly reclivous, weakly intercepted. Nervellar

166

index 0.7 (Fig. 28). Tibial spur index 0.6. Hind

femur 5.3x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to the middle of

the hind coxae, distinctly but not deeply and rath-

er narrowly depressed, for the greater part with

long transverse wrinkles, carinae almost complete-

ly obliterated. Propodeal spiracle rather small, 3.3x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.5. 1* gastral tergite 4.8x, 2” tergite

2.3x as long as wide. Petiole smooth laterally,

without glymmae or narrow lateral areas. Petiolar

suture fine, partly obliterated. 1* gastral sternite

smooth. Thyridium oval. Thyridial index about

2.0. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus dark brown.

Mandible sometimes marked with brown medial-

ly. Wings hyaline. Stigma dark brown medially.

Front leg from the femur onwards yellowish red.

Mid femur yellowish red on the apical half. Mid

tibia yellowish red. Hind tibia sometimes tinged

with dark brown medially. 2" gastral tergite red

on the posterior 0.3. 3° and 4" tergites red. Pilos-

ity whitish.

d: Maxillary palpus yellow. Mandible broad-

ly yellow medially. Tegula yellow. Front and mid

legs from the tip of the coxa onwards yellow, the

femora yellowish red. Hind tibia yellow medially,

narrowly black basally, broadly black apically. In

other characters similar to the ®.

Material (10°, 688): Russia: Chita Prov.: Klich-

ka, Kurort Darasun; Primorye: Slavyanka. Kaza-

khstan: Aksu-Dzhabagly ESE of Chimkent, Alma-

Ata, Borovoye near Kokchetav. Kirgiziya: Frunze

(= Bishkek), River Chatkal (PET), Tyan-Shan Mts.

(MOS). Mongoliya: Hövsgöl Prov.: Mörön (PET).

Dusona terebrator (Förster, 1868)

Description. 2: Body length 9-11 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 1.1. Frons slightly depressed, coriaceous,

densely rugose-punctate, with short radial wrin-

kles, a median longitudinal carina distinct medi-

ally, obliterated dorsally and ventrally. Antennal

carina low and narrow. Face and dorsal half of the

clypeus coriaceous, dull, densely rugose-punctate.

Clypeus flat, coriaceous and rather densely punc-

tate ventrally, its apex slightly convex medially.

Malar space index 0.5. Genal carina low and rath-

er straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 39-43 seg-

ments, filiform, the apical 0.2 attenuate, 24 seg-

ment 1.6-1.8x, shortest segment 0.9x, penultimate

segment 1.2x as long as wide. Pronotum coria-

ceous and densely rugose laterally on the dorsal

0.4, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely or very

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove smooth.

Prepectal carina low, ventral part merging with

the pleural part, the latter complete, joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum coriaceous, dull,

distinctly and densely punctate. Speculum coria-

ceous, rather dull. Depression in front of the specu-

lum with short and fine longitudinal wrinkles,

which merge with the punctation anteriorly. Meta-

pleurum coriaceous, densely rugose. Pleural cari-

na distinct. Areolet large, pointed, receiving the

2"d recurrent vein distad of the middle. Nervellus

slightiy reclivous, intercepted. Nervellar index

0.6-0.7. Tibial spur index 0.7. Hind femur 5.3x as

long as high. Hind tarsal claws long, with rather

short pectinate teeth basally and medially. Propo-

deum extending to 0.3 of the hind coxae, broadly

and not deeply depressed, partly irregularly wrin-

kled, with rather short transverse wrinkles in the

midline, with a median longitudinal keel posteri-

orly, carinae complete or almost complete. Propo-

deal spiracle rather small, 2.4x as long as wide.

Spiracular carina narrow. Spiracular index 0.7.

1% gastral tergite 3.5x, 2"d tergite 1.6x as long as

wide. Petiole with large glymmae, which are bor-

dered by longitudinal carinae dorsally, smooth or

with fine sculpture in front of the glymmae. Peti-

olar suture complete. 1% gastral sternite for the

greater part coriaceous, with transverse striae pos-

teriorly. Thyridium oval. Thyridial index about

1.5. Epipleurum separated from the 3" gastral

tergite, the crease marked with black. Epipleural

index 0.8. Gaster distinctly compressed posterior-

ly. Ovipositor index 1.0.

Colour: Black. Wings hyaline. Stigma yellow-

ish brown medially. Front leg from the femur

onwards yellowish red. Mid femur yellowish red

on the apical 0.3. Mid tibia and tarsus yellowish

red, the tarsus darkened apically. Hind tibia yel-

lowish brown medially, narrowly black basally,

broadly black apically. 2" gastral tergite red on

the posterior half. 3° and 4" tergites red. Pilosity

whitish.

d: Propodeum shallowly depressed, with dis-

tinct carinae. Mandible sometimes marked with

reddish brown medially. In other characters sim-

ilar to the 2.

Material (522, 639): Russia: Altaysk Terr.: Volgykha

(PET); Primorye: Andreyevka (KIS). Kazakhstan: Kus-

tanay (PET). Kirgiziya: Cholpon-Ata (LIN), Issykata

(MOS), Khaydarken, 2500-2800 m, Sosnovka, 1500-

1800 m, Talas, 1750-1900 m (LIN).

Dusona ternata Hinz & Horstmann, spec. nov.

Holotype (?): Japan, Odaira (locality unidenti-

fied, possibly in central Honshu), 25.8.1954 (HEL).

Description. 2: Body length 11mm. Temples

very strongly constricted behind the eyes, 0.6x as

long as the width of an eye, tangential lines on

eyes and temples intersect on the anterior half of

the mesoscutum. Ocellar index 0.5. Frons slightly

depressed, coriaceous, densely punctate, with a

distinct and slightly raised median longitudinal

carina, with short und indistinct radial wrinkles.

Antennal carina low and narrow. Face and clypeus

coriaceous, densely rugose-punctate. Clypeus flat,

its apex slightly concave medially. Malar space

index 0.3. Genal carina low and rather straight

ventrally, joining the slightly raised oral carina at

an acute angle. Genal index 0.6. Temple finely

coriaceous and with fine disperse punctation me-

dially. Flagellum with 48 segments, filiform, the

apical 0.3 distinctly attenuate, 2"? segment 2.1x,

shortest segment 0.9x, penultimate segment 1.3x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.2, with longitudinal

wrinkles ventrally. Mesoscutum and scutellum

coriaceous, very densely punctate. Scutellum with

lateral carinae on the anterior half. Prescutellar

groove with short longitudinal striae. Prepectal

carina low, ventral part merging with the pleural

part, the dorsal half of the latter obliterated. Trans-

verse carina complete, as high as the prepectal

carina. Mesopleurum finely coriaceous, densely

punctate. Speculum coriaceous and granulate-stri-

gose. Depression in front of the speculum with

distinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum coriaceous,

finely and densely punctate. Pleural carina distinct.

Areolet large, pointed, receiving the 2” recurrent

vein basad of the middle. Nervellus vertical, weak-

ly intercepted. Nervellar index 0.7. Tibial spur

index 0.6. Hind femur 5.3x as long as high. Hind

tarsal claws short, with short but distinct pecti-

nate teeth basally. Propodeum extending to the

167

middle of the hind coxae, narrowly and not deep-

ly depressed, with short transverse wrinkles in

the longitudinal furrow anteriorly and medially,

with long transverse wrinkles over the total width

of the propodeum posteriorly, costulae fine, com-

plete, longitudinal carinae present only posterior-

ly. Propodeal spiracle large, 3.5x as long as wide.

Spiracular carina somewhat widened. Spiracular

index 0.4. 1" gastral tergite 4.7x, 2" tergite 2.8x as

long as wide. Petiole without glymmae, with in-

distinct narrow lateral areas, smooth in front of

these areas. Petiolar suture almost completely ob-

literated. 1* gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium elongate. Thyridial index about 1.5. Epi-

pleurum not separated from the 3" gastral tergite,

the latter with distinct black lateral stripes well

above the ventrolateral edge. Gaster distinctly

compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula yellowish

brown. Wings hyaline. Stigma dark brown medi-

ally. Front leg from the tip of the coxa onwards

yellowish red. Mid femur yellowish red on the

apical 0.2. Mid tibia and tarsus yellowish red, the

tarsus brown apically. Hind tibia broadly yellow-

ish brown medially, narrowly dark brown basally

and apically. 2" gastral tergite black dorsally, with

a long red lateral mark posteriorly. 3“ and 4

tergites red. 5" tergite marked with red ventrola-

terally. Posterior gastral sternites reddish brown.

Pilosity whitish.

d unknown.

Dusona thomsoni Hinz, 1963

Description. ?: Body length 7-8 mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.7. Frons slightly depressed,

coriaceous and finely rugose-punctate dorsally,

coriaceous with few short radial wrinkles ventral-

Iy, a median longitudinal carina distinct medially,

obliterated dorsally and ventrally. Antennal cari-

na low and narrow. Face and dorsal 0.7 of the

clypeus coriaceous, finely and densely rugose-

punctate. Clypeus almost flat, coriaceous and rath-

er densely punctate ventrally, its apex truncate

medially. Malar space index 0.4. Genal carina low

ventrally, slightly bent forwards, joining the slight-

Iy raised oral carina at an acute angle. Genal index

0.1. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 39-42

segments, filiform, the apical 0.2 slightly attenu-

168

ate, 2" segment 2.3x, shortest segment 0.9x, pe-

nultimate segment 1.1x as long as wide. Prono-

tum coriaceous and finely rugose-punctate later-

ally on the dorsal 0.2, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

densely or very densely punctate. Scutellum with

lateral carinae on the anterior half. Prescutellar

groove with fine longitudinal striae. Ventral part

of the prepectal carina merging with the trans-

verse carina, the former low, the latter slightly

raised. Pleural part of the prepectal carina obliter-

ated dorsally and ventrally, the median 0.3 present.

Mesopleurum coriaceous, dull, distinctly and

densely punctate. Speculum granulate-strigose,

rather dull. Depression in front of the speculum

with distinct short longitudinal striae, which merge

with the punctation anteriorly. Metapleurum co-

riaceous, finely and densely rugose. Pleural cari-

na distinct. Areolet large, stalked, receiving the

2" recurrent vein almost at the middle. Nervellus

slightly inclivous, weakly intercepted. Nervellar

index 0.7-0.8. Tibial spur index 0.6. Hind femur

4.8x as long as high. Hind tarsal claws short, with

distinct pectinate teeth basally and medially. Pro-

podeum extending to the middle of the hind cox-

ae, broadly and shallowly depressed, sulcus with

short transverse wrinkles anteriorly, with long

transverse wrinkles medially and posteriorly, cos-

tulae complete and distinct, longitudinal carinae

obliterated anteriorly and medially, distinct pos-

teriorly. Propodeal spiracle rather small, 2.5x as

long as wide. Spiracular carina narrow. Spiracular

index 0.5. 1" gastral tergite 5.1x, 2" tergite 2.5x as

long as wide. Petiole without glymmae, with nar-

row lateral areas, smooth in front of these areas.

Petiolar suture partly obliterated. 1* gastral ster-

nite for the greater part smooth, with transverse

striae posteriorly. Thyridium elongate. Thyridial

index about 1.0. Epipleurum not separated from

the 3"4 gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.8.

Colour: Black. Maxillary palpus yellow. Man-

dible broadiy yellow medially. Tegula yellow.

Wings hyaline. Stigma medium brown medially.

Front leg from the tip of the coxa onwards yellow,

the femur yellowish red. Mid femur yellow on the

apical 0.2. Mid tibia and tarsus yellow. Hind tibia

yellow medially, narrowly black basally, very

broadly black apically, the apical black ring ex-

tending over 0.3 of the tibia. 24 gastral tergite

completely black or narrowly marked with red

laterally. 3° tergite red, distinctly marked with

black dorsally on the anterior 0.7. 4" tergite red,

indistinctly marked with brown dorsally. 5" to 7"

tergites broadly black dorsally, narrowly red ven-

trolaterally (to a variable extend). Gastral ster-

nites red or reddish brown. Pilosity whitish.

d: Propodeum rather narrowly and shallow-

ly depressed, with transverse wrinkles in almost

all parts. Mid trochanter yellow. Mid femur yel-

lowish red. Genital clasper reddish brown. In oth-

er characters similar to the ?.

Material (2??): Russia: Chita Prov.: Nerchinskiy Za-

vod. Kazakhstan: Borovoye near Kokchetav (PET).

Dusona tibiatoria Horstmann, spec. nov.

Holotype (3): Mongoliya, Central Prov. (= Töv

Prov.), Lün, Lün-uul Mts., 16.6.1975 (PET).

Description. d: Body length 10 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.1. Frons slightly depressed, coriaceous,

densely punctate, with a fine but distinct median

longitudinal carina, with fine radial wrinkles,

which extend to half the distance to the median

ocellus. Antennal carina low and narrow. Face

and dorsal 0.7 of the clypeus coriaceous, densely

rugose-punctate. Clypeus concave, coriaceous and

rather densely punctate ventrally, its apex convex

medially. Malar space index 0.4. Genal carina

straight ventrally, distinctly raised, joining the

distinctly raised oral carina at an acute angle, form-

ing a lopsided triangular pyramid, the tip of the

latter shifted ventrally and situated behind the

base of the mandible. Gena depressed close to the

junction, the depression about half as deep as the

width of the 2" flagellar segment. Genal index 0.4.

Temple finely coriaceous and with fine disperse

punctation medially. Flagellum with 47 segments,

filiform, the apical 0.3 attenuate, 2" segment 1.4x,

shortest segment 0.9x, penultimate segment 1.0x

as long as wide. Pronotum very densely punctate

on a smooth background laterally on the dorsal

0.6, with rather irregular wrinkles ventrally. Meso-

scutum and scutellum very densely punctate,

mesoscutum with a coriaceous, scutellum with a

smooth background. Scutellum with lateral cari-

nae on the anterior half. Prescutellar groove

smooth medially, longitudinally striate laterally.

Ventral part of the prepectal merging with the

transverse carina, both carinae low. Pleural part

ofthe prepectal carina replaced by some indistinct

short wrinkles medially, obliterated dorsally and

ventrally. Pleural part of the prepectus densely

punctate on asmooth background, with few wrin-

kles. Mesopleurum densely and coarsely punc-

tate on a smooth background. Speculum punctate

posteriorly, with fine oblique striae anteriorly.

Depression in front of the speculum with distinct

longitudinal wrinkles, which merge with the punc-

tation anteriorly. Metapleurum very densely punc-

tate on a smooth background. Pleural carina dis-

tinct. Areolet large, pointed, receiving the 2"! re-

current vein slightly basad of the middle. Nervellus

vertical, distinctly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 4.2x as long as

high. Hind tarsal claws long, with distinct pecti-

nate teeth almost over their total length. Propo-

deum extending to the middle of the hind coxae,

distinctly and broadly depressed, sulcus with short

transverse wrinkles in the midline, with irregular

wrinkles laterally, costulae distinct and complete,

longitudinal carinae present only posteriorly. Pro-

podeal spiracle rather large, 3.0x as long as wide.

Spiracular carina narrow. Spiracular index 0.2. 1%

gastral tergite 4.5x, 2" tergite 1.9x as long as

wide. Petiole without glymmae, narrow lateral

areas weakly indicated, smooth in front of these

areas. Petiolar suture almost completely obliterat-

ed. 1“ gastral sternite smooth. Thyridium elon-

gate. Thyridial index about 1.0. Epipleurum not

separated from the 3 gastral tergite, the latter

with distinct black lateral stripes well above the

ventrolateral edge. Gaster distinctly compressed

posteriorly.

Colour: Black. Maxillary palpus brown basal-

ly, yellow medially and apically. Mandible nar-

rowly marked with yellowish red dorsally and

medially. Tegula yellow. Wings hyaline. Stigma

light yellowish brown medially. Front leg from

the tip of the coxa onwards yellowish red. Mid leg

from the trochantellus onwards yellowish red.

Hind femur yellowish red on the apical 0.1. Hind

tibia completely yellowish red. 2” gastral tergite

dorsally black, laterally red on the posterior 0.4.

3 tergite red. 4" tergite narrowly marked with

red anteriorly. Pilosity whitish. (The legs might

have been be marked with yellow originally, and

the yellowish red markings might be an artefact

caused by the killing agent.)

? unknown.

169

Dusona tritor Hinz & Horstmann, spec. nov.

Holotype (?): Japan, Honshu, Nagano, Tenryu-

kyo, 3.6.1950 (AEI).

Description. 2: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.8. Frons scarcely depressed, coriaceous,

very finely and rather densely punctate, with a

fine median longitudinal carina, with indistinct

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.6 of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, coriaceous

and rather densely punctate ventrally, its apex

slightly concave medially. Malar space index 0.4.

Genal carina low and rather straight ventrally,

joining the slightly raised oral carina at an acute

angle. Genal index 0.25. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 41 segments, filiform, its apical 0.2

slightly attenuate, 2" segment 1.9x, shortest seg-

ment 1.0x, penultimate segment 1.2x as long as

wide. Pronotum coriaceous and finely and dense-

ly rugose-punctate laterally on the dorsal 0.4, with

longitudinal wrinkles ventrally. Mesoscutum co-

riaceous, finely and very densely punctate. Scutel-

lum coriaceous, distinctly and densely punctate,

with lateral carinae on the anterior 0.4. Prescutel-

lar groove almost smooth, with few fine longitu-

dinal striae. Ventral part of the prepectal carina

low, merging with the transverse carina, the latter

slightly raised. Pleural part of the prepectal carina

low, the ventral 0.2 and the dorsal 0.3 indistinct

and covered with wrinkles. Mesopleurum coria-

ceous, densely punctate. Speculum coriaceous and

dull. Depression in front of the speculum with

distinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum densely and

finely rugose. Pleural carina strong. Areolet large,

short-stalked, receiving the 2" recurrent vein dis-

tinctly distad of the middle. Nervellus slightly

inclivous, weakly intercepted. Nervellar index 0.7.

Tibial spur index 0.6. Hind femur 5.7x as long as

high. Hind tarsal claws short, with short pectinate

teeth basally. Propodeum extending to the mid-

dle of the hind coxae, slightly and rather narrowly

depressed, with fine long transverse wrinkles an-

teriorly and medially, with irregular wrinkles pos-

teriorly, costulae fine but complete, longitudinal

carinae present only posteriorly. Propodeal spira-

cle rather small, 2.9x as long as wide. Spiracular

carina narrow. Spiracular index 0.6. 1 gastral

tergite 4.9x, 2" tergite 2.0x as long as wide. Peti-

ole without glymmae, narrow lateral areas indi-

170

cated by a coriaceous surface, with very small and

shallow pits at their anterior edge, petiole smooth

in front of these areas. Petiolar suture partly oblit-

erated. 1“ gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum not separated from the 3" gastrai tergite, the

latter with indistinct black lateral stripes well above

the ventrolateral edge. Gaster strongly compressed

posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus dark brown

basally, yellowish brown medially and apically.

Mandible tinged with dark brown medially. Wings

hyaline. Stigma dark brown medially. Front leg

from the femur onwards yellowish red, the femur

marked with black basally. Mid femur yellowish

brown on the apical 0.1. Mid tibia yellowish brown.

Mid tarsus dark brown. Hind tibia black basally

and ventrally, dark reddish brown apically and

dorsally. 2”! gastral tergite black dorsally, with a

rather long narrow red mark laterally. 3 and 4"

tergites red, marked with dark brown dorsally. 5"

tergite broadly marked with red anteriorly and

laterally. Pilosity whitish.

d unknown.

Dusona ucrainator Hinz, 1985

Both sexes differ remarkably from each other in

the sculpture of the frons and in the antennal

carina. There is a slight possibility that the dd

may belong to a different, undescribed species.

Description. 2: Body length 9 mm. Temples dis-

tinctly constricted behind the eyes, 0.7x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.5. Frons not depressed, coriaceous, densely ru-

gose-punctate and rugose, with a fine median

longitudinal carina, with distinct radial wrinkles,

which almost reach the median ocellus. Antennal

carina slightly but distinctly raised, the rim bent

upwards (ear-like), forming a crescent-shaped

transversely striate plate. Face and dorsal half of

the clypeus coriaceous, densely rugose. Clypeus

flat, rather densely punctate on a finely coriaceous

and shining background ventrally, its apex trun-

cate medially. Malar space index 0.3. Genal carina

low ventrally, bent sidewards and forwards, join-

ing the oral carina at the base of the mandible at a

right angle. Temple finely coriaceous and with

very fine disperse punctation medially. Flagellum

with 39-41 segments, filiform, the apical 0.2 atten-

uate, 2"! segment 2.4x, shortest segment 1.0x, pen-

ultimate segment 1.3x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.1, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely punc-

tate, the scutellum finely coriaceous and some-

what shining. Scutellum with lateral carinae on

the anterior 0.7. Prescutellar groove smooth medi-

ally, with fine longitudinal striae laterally. Ventral

part ofthe prepectal carina merging with the trans-

verse carina, the former distinctly raised and dis-

tinctly emarginate medially, the latter slightly

raised. Pleural part of the prepectal carina low

ventrally and medially, the dorsal 0.3 obliterated

and replaced by wrinkles. Mesopleurum finely

coriaceous, densely or rather densely punctate.

Speculum coriaceous and granulate-strigose. De-

pression in front of the speculum with dorsally

with distinct longitudinal striae, ventrally with

fine striae, which merge with the punctation ante-

riorly. Metapleurum finely and densely rugose-

punctate and rugose. Pleural carina distinct. Are-

olet rather large, short-stalked, receiving the 2“

recurrent vein slightly basad of, in or slightly dis-

tad of the middle. Nervellus slightly reclivous,

not intercepted or weakly intercepted. In the lat-

ter case nervellar index 0.7. Tibial spur index 0.6.

Hind femur 5.3x as long as high. Hind tarsal cıaws

short, with short pectinate teeth basally. Propo-

deum extending to the middle of the hind coxae,

broadly and deeply depressed, for the greater

part with fine transverse wrinkles, carinae almost

completely obliterated. Propodeal spiracle rather

small, 4.0x as long as wide. Spiracular carina wid-

ened. Spiracular index about 0.4. 1* gastral tergite

4.8x, 2"d tergite 2.0x as long as wide. Petiole with

small glymmae at the anterior edge of narrow

lateral areas, smooth in front of the glymmae.

Petiolar suture fine, partly obliterated. 1* gastral

sternite smooth. Thyridium elongate. Thyridial

index about 1.5. Epipleurum not separated from

the 3"! gastral tergite, the latter with distinct black

lateral stripes well above the ventrolateral edge.

Gaster distinctly compressed posteriorly. Ovipos-

itor index 0.5.

Colour: Black. Maxillary palpus yellowish red.

Mandible narrowly marked with yellow dorsally

and medially. Tegula yellow. Wings hyaline. Stig-

ma medium brown medially. Front leg from the

tip of the coxa onwards yellow, the trochanter

marked with brown or black, the femur yellowish

red. Mid femur yellowish red on the apical 0.3.

Mid tibia yellow. Mid tarsus yellowish brown or

brown. Hind tibia light yellow medially on the

dorsal side, narrowly black basally, brown medi-

ally on the ventral side, broadly marked with

black apically, the apical black ring extending over

0.3 of the tibia. 2"4 gastral tergite red on the poste-

rior 0.1-0.2. 3" and 4" tergites red, the 4" tergite

sometimes marked with black posteriorly. Pilosi-

ty whitish.

d: Frons distinctly but not deeply depressed

above the antennal carinae. Antennal carinae rather

strongly raised, the rim bent upwards (ear-like),

forming a crescent-shaped transversely striate

plate. Mandible broadly yellow medially. Front

and mid leg from the tip of the coxa onwards

yellow. In other characters similar to the 2.

Material (8?2, 538): Russia: Chita Prov.: Chita (PET),

Lake Ivan (Hinz/ZSM, PET); Primorye: Spassk Dalniy

(KIS, PET). Japan: Honshu: Kamikochi near Norikura-

dake (AEI, Hinz/ZSM).

Dusona ucrainica Hinz, 1972

Description. 2: Body length 9-10 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.9. Frons slightly depressed, finely and densely

rugose dorsally, with distinct radial wrinkles me-

dially and ventrally, the latter almost reaching the

median ocellus, with a median longitudinal cari-

na, which is distinctly raised medially. Antennal

carina distinctly but not strongly raised, the rim

bent upwards (ear-like), forming a crescent-shaped

plate, which is transversely striate. Face coria-

ceous, densely rugose-punctate. Clypeus slightly

convex, finely coriaceous, densely punctate, its

apex slightly convex medially. Malar space index

0.4. Genal carina low ventrally, slightly bent side-

wards, joining the oral carina at the base of the

mandible at a right angle. Temple coriaceous and

with fine disperse punctation medially. Flagellum

with 42-43 segments, filiform, the apical 0.3 dis-

tinctly attenuate, 2" segment 2.1x, shortest seg-

ment 1.1x, penultimate segment 1.4x as long as

wide. Pronotum densely rugose-punctate lateral-

ly on the dorsal 0.3, with longitudinal wrinkles

ventrally. Mesoscutum and scutellum coriaceous,

densely rugose-punctate. Scutellum with lateral

carinae on the anterior 0.4-0.5. Prescutellar groove

smooth medially, with fine longitudinal striae lat-

erally. Ventral part of the prepectal carina merg-

ing with the transverse carina, both carinae slight-

ly raised, the former emarginate medially. Pleural

part of the prepectal carina low ventrally and

medially, the dorsal 0.3 obliterated. Mesopleu-

rum coriaceous, densely punctate. Speculum with

fine longitudinal striae dorsally, finely granulate-

strigose and shining ventrally. Depression in front

of the speculum with distinct long longitudinal

171

striae, which merge with the punctation anterior-

ly. Metapleurum densely rugose-punctate and

rugose. Pleural carina fine but complete. Areolet

rather small, pointed or short-stalked, receiving

the 2" recurrent vein distinctly distad of the mid-

dle. Nervellus distinctly reclivous, weakly inter-

cepted. Nervellar index 0.7. Tibial spur index 0.5.

Hind femur 4.7x as long as high. Hind tarsal claws

short, with distinct pectinate teeth basally and

medially. Propodeum extending to 0.4 of the hind

coxae, rather broadly but not deeply depressed,

irregularly wrinkled anteriorly, with short trans-

verse wrinkles medially, with long transverse

wrinkles posteriorly, carinae almost completely

obliterated. Propodeal spiracle rather large, 3.1x

as long as wide. Spiracular carina narrow. Spirac-

ular index 0.3. 1" gastral tergite 4.1x, 2" tergite

1.6x as long as wide. Petiole with small and shal-

low glymmae at the anterior edge of indistinct

narrow lateral areas, smooth in front of the glym-

mae. Petiolar suture obliterated. 1” gastral sternite

smooth. Thyridium elongate. Thyridial index

about 1.0. Epipleurum not separated from the 3'°

gastral tergite, the latter with distinct black lateral

stripes well above the ventrolateral edge. Gaster

slightly compressed posteriorly. Ovipositor index

05:

Colour: Black. Maxillary palpus medium brown

basally, yellowish red medially and apically. Man-

dible black, sometimes narrowly tinged with

brown medially. Tegula dark brown. Wings hya-

line. Stigma yellowish brown medially. Front fe-

mur yellowish red. Front tibia and tarsus yellow.

Mid femur yellowish red on the apical 0.2. Mid

tibia yellow. Mid tarsus brown. Hind tibia com-

pletely black or marked with whitish yellow sub-

basally on the dorsal side (to a variable extend).

2nd gastral tergite red on the posterior 0.3. 3° and

4" tergites red. Pilosity whitish.

d: Glymma lacking. Mandible broadly yel-

low medially. Tegula yellowish brown. Front tro-

chanter marked with yellowish red. In other char-

acters similar to the 9.

Material (1222, 1338): Russia: Omsk Prov.: Omsk;

Chita Prov.: Karymskoye, Kozlovo, Urulyunguy E of

Borzya, Zabaykalsk; Amur Prov.: Novorossiyka; Yev-

reyskiy A.O.: Amurzet (PET); Primorye: Andreyevka

(KIS), Spassk Dalniy. Mongoliya: Selenge Prov.: Dar-

han (PET). North Korea: Hyangsan (SAW). Japan:

Honshu: Tokyo (AEI).

11772

Dusona variator (Hinz, 1990)

Description. 2: Body length 7-9 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.0. Frons slightly depressed, coriaceous, finely

punctate, with a very fine median longitudinal

carina (sometimes obliterated), with very short

radial wrinkles. Antennal carina low and narrow.

Face and dorsal 0.6-0.7 of the clypeus coriaceous,

densely rugose-punctate or rugose. Clypeus flat,

finely coriaceous and rather densely punctate ven-

trally, its apex truncate medially. Malar space in-

dex 0.4. Genal carina low and straight ventrally,

joining the slightly raised oral carina at an acute

angle. Genal index 0.6. Temple finely coriaceous

and with very fine disperse punctation medially.

Flagellum with 42-44 segments, filiform, the api-

cal 0.2 slightly attenuate, 2” segment 1.9x, short-

est segment 0.9x, penultimate segment 1.0-1.1x as

long as wide. Pronotum coriaceous and densely

rugose-punctate laterally on the dorsal 0.4-0.6,

with longitudinal wrinkles ventrally. Mesoscu-

tum and scutellum coriaceous, densely punctate.

Scutellum with lateral carinae on the anterior

0.5-0.6. Prescutellar groove smooth, sometimes

with fine longitudinal striae laterally. Prepectal

carina low, ventral part merging with the pleural

part, the latter complete or almost complete. Trans-

verse carina obliterated. Mesopleurum coriaceous,

dull, densely punctate. Speculum granulate-stri-

gose. Depression in front of the speculum with

rather long longitudinal wrinkles, which merge

with the punctation anteriorly. Metapleurum co-

riaceous, finely and densely rugose-punctate. Pleu-

ral carina distinct. Areolet rather small, pointed or

narrowly sessile, receiving the 2" recurrent vein

distinctly distad of the middle (Fig. 24). Nervellus

slightly inclivous, weakly intercepted. Nervellar

index 0.7-0.8. Tibial spur index 0.7. Hind femur

5.0x as long as high. Hind tarsal claws short, with

short pectinate teeth basally. Propodeum extend-

ing to 0.4 of the hind coxae, rather broadliy de-

pressed, the sulcus coriaceous and with fine irreg-

ular wrinkles, with some transverse wrinkles and

a short median longitudinal keel posteriorly, cos-

tulae partly present, longitudinal carinae almost

completely obliterated. Propodeal spiracle rather

large, 2.7x as long as wide. Spiracular carina nar-

row. Spiracular index 0.4. 1% gastral tergite 3.6x,

2rd tergite 1.4x as long as wide. Petiole with large

glymmae, which are bordered by longitudinal

carinae dorsally, smooth in front of the glymmae.

Petiolar suture complete. 1“ gastral sternite for the

greater part smooth, with transverse striae poste-

riorly. Thyridium oval. Thyridial index about 1.0.

Epipleurum not separated from the 3° gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.7.

Colour: Black. Maxillary palpus brownish

black. Mandible narrowly reddish brown to brown

medially. Wings hyaline. Stigma medium brown

medially. Front femur reddish brown on the api-

cal half or for the greater part light red, with a

brown mark ventrally. Front tibia and tarsus yel-

lowish brown. Mid femur sometimes reddish

brown on the apical 0.2. Mid tibia yellowish brown.

Mid tarsus dark brown. Hind tibia completely

black or tinged with yellowish brown medially on

the dorsal side. 2” gastral tergite red on the poste-

rior 0.1-0.2. 3" tergite red, sometimes marked with

black dorsally. 4" tergite sometimes marked with

red. Pilosity whitish.

d: Longitudinal carinae of the propodeum of-

ten indicated medially. Tegula yellowish brown

to dark brown. Front trochanter and femur yel-

lowish red. Mid femur yellowish red on the apical

0.2. Mid tarsus yellowish red. 2” gastral tergite

sometimes red on the posterior 0.3, the red marks

more extended laterally. 4" tergite marked with

red anteriorly. In other characters similar to the 2.

Material (3??, 485): Russia: Chita Prov.: Kalga,

Kurort Darasun; Khabarowsk Terr.: Khekhtsir near

Khabarovsk, Vysokogornyy; Primorye: Roshchino on

River Iman. Mongoliya: Ara-Hangay Prov.: 40 km

SSW of Tövshruuleh (PET).

Dusona varipes Horstmann, spec. nov.

Types. Holotype (S): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarovsk, 30.7.1983 (PET). -

Paratype: 13 Russia, Primorye, Ussuriysk, 27.6.

1981 (Hinz / ZSM).

Description. d: Body length 9-10 mm. Temples

distinctly constricted behind the eyes, 0.7x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

0.8. Frons slightly depressed, coriaceous, finely

and densely punctate, with a distinct median lon-

gitudinal carina, with rather long radial wrinkles,

which extend to 0.7 the distance to the median

ocellus. Antennal carina low and narrow. Face

and dorsal 0.7 of the clypeus coriaceous, densely

rugose-punctate. Clypeus flat, coriaceous and rath-

er densely punctate ventraliy, its apex truncate

medially. Malar space index 0.4. Genal carina

straight ventrally, slightly raised, joining the slight-

ly raised oral carina at an acute angle. Genal index

0.6. Temple finely coriaceous and with fine rather

dense punctation medially. Flagellum with 50-51

segments, filiform, the apical 0.3 attenuate, 2"d

segment 1.5x, shortest segment 1.0x, penultimate

segment 1.1x as long as wide. Pronotum coria-

ceous and densely rugose-punctate laterally on

the dorsal 0.3, with longitudinal wrinkles ventral-

ly. Mesoscutum and scutellum coriaceous, dull,

very densely punctate. Scutellum with lateral car-

inae on the anterior 0.7. Prescutellar groove with

dense longitudinal striae. Ventral part of the pre-

pectal carina low, merging with the transverse

carina, the latter slightly raised. Pleural part of the

prepectal carina replaced by a short wrinkle me-

dially, obliterated dorsally and ventrally. Meso-

pleurum coriaceous, dull, distinctly and densely

punctate. Speculum coriaceous, rather dull. De-

pression in front of the speculum with distinct

short longitudinal wrinkles, which merge with

the punctation anteriorly. Metapleurum coria-

ceous, densely and finely rugose. Pleural carina

distinct. Areolet rather small, long-stalked, receiv-

ing the 2” recurrent vein slightly distad of the

middle. Nervellus slightly reclivous, weakly in-

tercepted. Nervellar index 0.8. Tibial spur index

0.7. Hind femur 4.5x as long as high. Hind tarsal

claws short, with distinct pectinate teeth basally.

Propodeum extending to 0.4 of the hind coxae,

distinctly but not deeply and rather narrowly de-

pressed, the longitudinal furrow for the greater

part transversely wrinkled, other parts irregularly

wrinkled, costulae partly present, longitudinal

carinae indicated medially and posteriorly. Pro-

podeal spiracle rather large, 3.3x as long as wide.

Spiracular carina narrow. Spiracular index 0.7.

1 gastral tergite 4.4x, 2"X tergite 2.2x as long as

wide. Petiole with small and rather shallow glym-

mae at the anterior edge of narrow lateral areas,

for the greater part smooth in front of the glym-

mae, with a small depression and short transverse

wrinkles anteriorly. Petiolar suture fine, partly

obliterated. 1% gastral sternite for the greater part

smooth, with some transverse striae posteriorly.

Thyridium oval. Thyridial index about 1.5. Epi-

pleurum not separated from the 3" gastral tergite,

the latter with distinct black lateral stripes well

above the ventrolateral edge. Gaster strongly com-

pressed posteriorly.

Colour: Black. Maxillary palpus yellow. Man-

dible yellow, the teeth black. Tegula dark brown

or black. Wings hyaline. Stigma dark brown me-

dially. Front leg from the tip of the coxa onwards

yellow, the femur yellowish red, marked with

dark brown ventrally. Mid femur yellowish red

dorsally and apically. Mid tibia yellowish, marked

1174)

with dark brown on the inside. Mid tarsus dark

brown. Hind tibia marked with dark yellowish

brown medially on the ventral side. 2” gastral

tergite dorsally red on the posterior 0.4, laterally

red on the posterior 0.2. 3° and 4" tergites red, the

4" tergite marked with black posteriorly. Pilosity

whitish.

? unknown.

Dusona venitor Hinz & Horstmann, spec. nov.

Holotype (?): Russia, Irkutsk Prov., Lake Baykal,

Kultuk, 19.6.1929 (MOS).

Description. 2: Body length 10 mm. Temples dis-

tinctly constricted behind the eyes, 0.8x as long as

the width of an eye, tangential lines on eyes and

temples intersect on the postscutellum. Ocellar

index 1.0. Frons slightly depressed, coriaceous,

finely rugose-punctate, with a fine median longi-

tudinal carina, with indistinct radial wrinkles.

Antennal carina low and narrow. Face and dorsal

0.7 ofthe clypeus coriaceous, densely rugose-punc-

tate. Ciypeus flat, coriaceous and with rather dense

punctation ventrally, its apex almost truncate

medially. Malar space index 0.4. Genal carina low

and straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.5.

Temple coriaceous and with fine disperse puncta-

tion medially. Flagellum with 48-49 segments, fili-

form, the apical 0.3 slightly attenuate, 2"d segment

1.5x, shortest segment 1.0x, penultimate segment

1.2x as long as wide. Pronotum densely rugose-

punctate laterally on the dorsal 0.3, with longitu-

dinal wrinkles ventrally. Mesoscutum and scutel-

lum coriaceous, finely and very densely punctate.

Scutellum with lateral carinae on the anterior half.

Prescutellar groove with fine short longitudinal

striae. Prepectal carina low, the ventral part merg-

ing with the pleural part, the dorsal 0.2 of the

latter replaced by several wrinkles. Transverse

carina obliterated. Mesopleurum distinctly coria-

ceous, densely punctate. Speculum and depres-

sion in front of the speculum with fine longitudi-

nal striae, which merge with the punctation ante-

riorly. Metapleurum densely rugose-punctate and

finely rugose. Pleural carina distinct. Areolet large,

pointed, receiving the 2" recurrent vein slightly

basad of the middle. Nervellus almost vertical,

weakly intercepted. Nervellar index 0.7. Tibial

spur index 0.8. Hind femur 4.7x as long as high.

Hind tarsal claws short, with short pectinate teeth

basally. Propodeum extending to 0.3 of the hind

coxae, broadly and slightly depressed, sulcus part-

Iy somewhat shining, for the greater part irregu-

174

larly wrinkled, with few short transverse wrin-

kles, carinae almost completely obliterated. Pro-

podeal spiracle rather large, 3.4x as long as wide.

Spiracular carina obliterated. Spiracular index 0.7.

1" gastral tergite 3.7x, 2"! tergite 2.0x as long as

wide. Petiole with distinct but rather small and

shallow glymmae, which are bordered by longitu-

dinal carinae dorsally, smooth in front of the glym-

mae. Petiolar suture for the greater part present.

1" gastral sternite for the greater part smooth,

with transverse striae posteriorly. Thyridium oval.

Thyridial index about 2.0. Epipleurum separated

from the 3! gastral tergite anteriorly and folded

in, the crease marked with black. Epipleural index

0.2. Gaster strongly compressed posteriorly. Ovi-

positor index 0.6.

Colour: Black. Wings hyaline. Stigma medi-

um brown medially. Front femur marked with

yellowish brown dorsally. Front tibia and tarsus

medium brown. Mid tibia and tarsus dark brown.

2n“ gastral tergite red on the posterior 0.2. 3" and

4" tergites red. Pilosity whitish.

d unknown.

Dusona ventrator Hinz & Horstmann,

spec. nov.

Holotype (?): Japan, Hokkaido, Jozankei, 10.7.

1954 (AEI).

Description. ?: Body length 11mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 1.0. Frons slightly depressed,

coriaceous, finely and rather densely punctate

dorsally, with a fine median longitudinal carina,

with distinct radial wrinkles, which extend to 0.4

the distance to the median ocellus. Antennal cari-

na low and narrow. Face and dorsal 0.6 of the

clypeus densely rugose-punctate. Clypeus flat,

coriaceous and densely punctate ventrally, its apex

truncate medially. Malar space index 0.4. Genal

carina low and straight ventrally, joining the dis-

tinctly raised oral carina at an acute angle. Genal

index 0.4. Temple finely coriaceous and with fine

disperse punctation medially. Flagellum with 49

segments, filiform, the apical 0.2 distinctly attenu-

ate, 2"! segment 2.0x, shortest segment 0.9x, pen-

ultimate segment 1.2x as long as wide. Pronotum

densely rugose-punctate laterally on the dorsal

0.1, with longitudinal wrinkles and with some

punctures ventrally. Mesoscutum and scutellum

coriaceous, finely and densely punctate. Scutel-

lum with lateral carinae on the anterior 0.6. Pre-

scutellar groove almost smooth, with some fine

longitudinal striae. Prepectal carina low, ventral

part merging with the pleural part, the dorsal 0.2

of the latter obliterated. Transverse carina obliter-

ated. Mesopleurum coriaceous, densely punctate.

Speculum finely coriaceous, shining. Depression

in front of the speculum with short irregular wrin-

kles dorsally, with fine short longitudinal striae

medially and ventrally, the latter merging with a

narrow coriaceous area without punctation ante-

riorly. Metapleurum coriaceous, densely punc-

tate. Pleural carina complete. Areolet large, short-

stalked, receiving the 2”! recurrent vein slightly

distad of the middle. Nervellus inclivous, strong-

ly intercepted. Nervellar index 0.8. Tibial spur

index 0.6. Hind femur 4.9x as long as high. Hind

tarsal claws rather short, with distinct pectinate

teeth basally and medially. Propodeum extending

to 0.3 of the hind coxae, broadly and not deeply

depressed, sulcus for the greater part irregularly

wrinkled, with some longitudinal wrinkles poste-

riorly, costulae partly present, longitudinal cari-

nae indicated medially and posteriorly. Propode-

al spiracle large, 3.5x as long as wide. Spiracular

carina widened. Spiracular index 0.2. 1% gastral

tergite 3.7x, 2" tergite 2.0x as long as wide. Peti-

ole with rather narrow but deep glymmae, which

are bordered by longitudinal carinae dorsally,

smooth in front of the glymmae. Petiolar suture

partly obliterated. 1“ gastral sternite smooth. Thy-

ridium oval. Thyridial index about 1.0. Epipleu-

rum not separated from the 3" gastral tergite, the

latter with distinct black lateral stripes well above

the ventrolateral edge. Gaster distinctly but not

strongly compressed posteriorly. Ovipositor in-

dex 0.6.

Colour: Black. Maxillary palpus medium

brown basally, yellow medially and apically. Man-

dible broadly yellow medially. Tegula medium

brown. Wings hyaline. Stigma light brown medi-

ally. Front leg from the trochanter onwards yel-

low, the femur tinged with brown basally. Mid

femur yellowish red on the apical 0.2. Mid tibia

yellow. Mid tarsus medium brown. Hind tibia

broadly yellowish brown medially, narrowly black

basally, narrowly medium brown apically. 2‘

gastral tergite black dorsally, broadly red lateral-

ly. 34 tergite red. 4" tergite red, dorsally marked

with brown posteriorly. Posterior tergites black

dorsally, red laterally. Gastral sternites red. Pilos-

ity whitish.

d unknown.

Dusona vidua (Gravenhorst, 1829)

Description. 2: Body length 6-8 mm. Temples

distinctly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the prescutellar groove. Ocel-

lar index 0.6. Frons not depressed, coriaceous,

finely rugose-punctate, with a scarcely indicated

median longitudinal carina, with few short radial

wrinkles. Antennal carina low and narrow. Face

and dorsal 0.3 of the clypeus finely rugose-punc-

tate. Clypeus flat, coriaceous and with fine dis-

perse punctation ventrally, its apex truncate me-

diaily. Malar space index 0.4. Genal carina slightly

raised ventrally, almost straight, joining the slightly

raised oral carina at an acute angle. Genal index

0.2. Temple finely coriaceous and with fine dis-

perse punctation medially. Flagellum with 30-34

segments, filiform, the apical 0.2 attenuate, 2"“

segment 2.4x, shortest segment 1.0x, penultimate

segment 1.2x as long as wide. Pronotum granu-

late-strigose laterally on the dorsal 0.3, with longi-

tudinal wrinkles ventrally. Mesoscutum and

scutellum coriaceous, dull, with fine disperse punc-

tation. Scutellum with lateral carinae on the ante-

rior 0.2. Prescutellar groove with short longitudi-

nal wrinkles. Prepectal carina low, ventral part

merging with the pleural part, the latter complete,

joining the anterior edge of the mesopleurum.

Transverse carina indistinct. Mesopleurum dis-

tinctly coriaceous, with fine disperse punctation.

Speculum coriaceous, rather dull. Depression in

front of the speculum with fine longitudinal stri-

ae, which merge with a coriaceous area without

punctation anteriorly. Metapleurum finely and

densely rugose. Pleural carina fine. Areolet small,

stalked, receiving the 2” recurrent vein distinctly

distad of the middle. Nervellus slightly inclivous,

distinctly intercepted. Nervellar index 0.8. Tibial

spur index 0.5. Hind femur 5.0x as long as high.

Hind tarsal claws short, with rather long pectinate

teeth basally and medially. Propodeum extending

to 0.3 of the hind coxae, distinctly and broadly

depressed, sulcus coriaceous anteriorly, trans-

versely striate medially and posteriorly, costulae

fine, longitudinal carinae complete and distinct.

Propodeal spiracle small, 3.0x as long as wide.

Spiracular carina narrow or partly widened. Spirac-

ular index 0.5. 1% gastral tergite 4.0x, 2” tergite

1.4x as long as wide. Petiole with very fine sculp-

ture laterally, without glymmae or narrow lateral

areas. Petiolar suture fine, partly obliterated.

1 gastral sternite finely coriaceous. Thyridium

elongate. Thyridial index about 1.0. Epipleurum

separated from the 3 gastral tergite anteriorly.

Black lateral stripes unrecognizable. Epipleural

175

index 0.4-0.5. Gaster distinctly compressed poste-

riorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible almost completely yellow. Tegula yellow.

Wings hyaline. Stigma dark brown medially. Front

leg from the tip of the coxa onwards yellow, the

femur yellowish red. Mid femur yellowish red.

Mid tibia and tarsus yellow. Hind tibia whitish

yellow medially, narrowly black basally, broadly

black apically. Hind tarsus yellowish brown ba-

sally. Pilosity whitish.

d: Similar to the 2.

Material (422, 733): Russia: Chita Prov.: Adrianov-

ka; Khabarovsk Terr.: Khekhtsir near Khabarovsk; Pri-

morye: Khasan, Pogranichnyy, Vladivostok; Kam-

chatka: Milskovo, Kozyrevsk (PET). Japan: Honshu:

Kamikochi near Norikura-dake (AEI).

Dusona viduator Hinz & Horstmann,

spec. nov.

Types. Holotype (?): Russia, Khabarovsk Terr.,

Khekhtsir, near Khabarovsk, 12.6.1983 (PET). —

Paratypes: 52?, 253 same locality, 8.-30.6.1983

and 12.6.1985 (422, 15 PET, 12, 18 Hinz/ZSM),;

1? Russia, Primorye, Spassk Dalniy, 26.5.1985

(BER).

Description. 2: Body length 7-9 mm. Temples

slightly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the propodeum. Ocellar in-

dex 1.2. Frons slightly depressed, coriaceous, with

very fine punctation and very fine wrinkles, with

a fine median longitudinal carina, with short and

indistinct radial wrinkles, sometimes on either

side with a raised point between the antennal

socket and the eye. Antennal carina low and nar-

row. Face and dorsal 0.6 of the clypeus coriaceous,

densely rugose-punctate. Clypeus flat, coriaceous

and with rather dense punctation ventrally, its

apex truncate medially. Malar space index 0.4.

Genal carina low ventrally, straight or slightly

bent sidewards, joining the low oral carina at an

acute angle. Genal index 0.4-0.5. Temple finely

coriaceous and with very fine disperse punctation

medially. Flagellum with 35-37 segments, filiform,

the apical 0.1 slightly attenuate, 2”! segment 1.7x,

shortest segment 0.8x, penultimate segment 1.0x

as long as wide. Pronotum densely rugose-punc-

tate laterally on the dorsal 0.3, the wrinkles almost

smooth, with longitudinal wrinkles ventrally.

Mesoscutum and scutellum coriaceous, densely

punctate. Scutellum with lateral carinae on the

176

anterior 0.7. Prescutellar groove smooth. Prepec-

tal carina low, ventral part merging with the pleu-

ral part, the latter complete, joining the anterior

edge of the mesopleurum. Transverse carina ob-

literated. Mesopleurum with rather dense or dense

punctation ona finely coriaceous and shining back-

ground. Speculum very finely coriaceous or gran-

ulate-strigose, partly almost smooth. Depression

in front of the speculum granulate-strigose and

with fine longitudinal striae, which merge with

the punctation anteriorly. Metapleurum finely and

densely punctate on a finely coriaceous and shin-

ing background. Pleural carina distinct. Areolet

large, pointed or short-stalked, receiving the 2‘

recurrent vein at or basad of the middle. Nervel-

lus almost vertical, weakly intercepted. Nervellar

index 0.7. Tibial spur index 0.7. Hind femur 5.1x

as long as high. Hind tarsal claws short, with long

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, scarcely or not

at all depressed, coriaceous and shining anterior-

ly (region of the area superomedia), irregularly

wrinkled medially, with some long longitudinal

wrinkles posteriorly, costulae fine but complete,

longitudinal carinae almost completely obliterat-

ed. Propodeal spiracle rather large, 2.7 as long as

wide. Spiracular carina widened. Spiracular index

0.6. 1% gastral tergite 3.2x, 2" tergite 1.6x as long

as wide. Petiole with large glymmae, which are

bordered by longitudinal carinae dorsally, smooth

in front of the glymmae. Petiolar suture complete

or obliterated posteriorly. 1* gastral sternite for

the greater part smooth, with fine transverse stri-

ae posteriorly. Thyridium oval. Thyridial index

about 1.5. Epipleurum separated from the 3" gas-

tral tergite, the crease marked with black. Epi-

pleural index 0.7-0.8. Gaster distinctly but not

strongly compressed posteriorly. Ovipositor in-

dex 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula yellow or

brown. Wings hyaline. Stigma yellowish brown

medially. Front leg from the femur onwards yel-

low, the femur marked with brown basally. Mid

femur yellow on the apical 0.2. Mid tibia and

tarsus yellow. Hind tibia yellow medially, nar-

rowly black basally, broadly black apically. Hind

tarsus yellowish brown or brown. Gastral tergites

black, narrowly tinged with dark brown ventrola-

terally. Pilosity whitish.

6: Temples sometimes more strongly con-

stricted behind the eyes. Flagellum with 38-39

segments. Front and mid legs from the tip of the

coxa onwards yellow. In other characters similar

to the 9.

Dusona vigilator (Förster, 1868)

Description. 2: Body length 10-12 mm. Temples

distinctly constricted behind the eyes, 0.9x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the scutellum. Ocellar index

1.2. Frons slightly depressed, coriaceous, with fine

and disperse or rather dense punctation, with a

fine median longitudinal carina, with few fine

and indistinct radial wrinkles. Antennal carina

low and narrow, its rim slightly bent upwards

(inconspicuous). Face and dorsal 0.3 of the clypeus

coriaceous, dull, densely rugose-punctate and ru-

gose. Clypeus flat, coriaceous, dull and densely

punctate ventrally, its apex truncate medially.

Malar space index 0.3. Genal carina low and al-

most straight ventrally, joining the slightly raised

oral carina at an acute angle. Genal index 0.3.

Temple finely coriaceous and with very fine dis-

perse punctation medially. Flagellum with 48-55

segments, filiform, the apical 0.3 distinctly attenu-

atewpud segment 1.5x, shortest segment 0.9x, pen-

ultimate segment 1.2x as long as wide. Pronotum

coriaceous and densely rugose-punctate laterally

on the dorsal 0.4, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous, Jull,

very densely punctate. Scutellum with lateral car-

inae on the anterior 0.4. Prescutellar groove for

the greater part smooth. Prepectal carina low, ven-

tral part merging with the pleural part, the latter

complete, joining the anterior edge of the meso-

pleurum. Transverse carina obliterated. Mesopleu-

rum distinctly coriaceous, rather densely or dense-

ly punctate. Speculum coriaceous, somewhat shin-

ing. Depression in front of the speculum dorsally

with distinct short longitudinal wrinkles, which

merge with the punctation anteriorly, the ventral

0.3 with fine striae or granulate-strigose. Metapleu-

rum finely coriaceous, finely and rather densely

punctate. Pleural carina distinct. Areolet large,

short-stalked, receiving the 2" recurrent vein at

or slightly basad of the middle. Nervellus slightly

inclivous, distinctly intercepted. Nervellar index

0.8. Tibial spur index 0.6. Hind femur 4.8x as long

as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, rather broadly

and shallowly depressed, sulcus irregularly wrin-

kled anteriorly and medially, sometimes coria-

ceous without distinct wrinkles anteriorly, trans-

versely wrinkled posteriorly, sometimes with a

median longitudinal keel posteriorly, costulae part-

ly present, longitudinal carinae usually present

medially and posteriorly. Propodeal spiracle large,

3.6x as long as wide. Spiracular carina narrow.

Spiracular index 0.5. 1% gastral tergite 3.5x, 2"

tergite 1.8x as long as wide. Petiole with distinct

glymmae, which are bordered by longitudinal

carinae dorsally, smooth or with very fine sculp-

ture in front of the glymmae. Petiolar suture fine,

sometimes partly obliterated. 1“ gastral sternite

coriaceous anteriorly, finely coriaceous or smooth

medially, transversely striate posteriorly. Thyrid-

ium oval. Thyridial index about 1.5. Epipleurum

separated from the 3“ gastral tergite anteriorly,

folded in or folded back secondarily and pendent

as a narrow rectangular lobe, a black lateral stripe

parallel with the crease in a short distance anteri-

orly, divergent from the ventrolateral edge poste-

riorly. Epipleural index 0.2-0.3. Gaster distinctly

compressed posteriorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellowish

brown or brown. Mandible broadly yellow medi-

ally. Wings hyaline. Stigma medium brown medi-

ally. Front femur reddish brown, the basal 0.2

dark brown. Front tibia and tarsus yellow. Mid

femur yellow on the apical 0.1. Mid tibia yellow.

Mid tarsus dark brown. Hind tibia broadly dark

yellowish brown medially, narrowly black basal-

ly and apically. 2" gastral tergite dorsally black,

laterally marked with red in front of the black

posterior edge, sometimes also dorsally tinged

with red in front of the posterior edge. 3" and 4"

tergites red, the 4" tergite sometimes marked with

black (to a variable extend) or completely black.

Pilosity whitish.

d: Propodeum with somewhat stronger cari-

nae and coarser wrinkles. Maxillary palpus brown

basally, yellow medially and apically. Tegula yel-

low to brown. Front trochanter and femur yellow-

ish red. Mid trochanter and femur yellowish

brown. In other characters similar to the 2.

Material (1722, 638): Russia: Khabarovsk Terr.:

Khekhtsir near Khabarovsk; Primorye: Khasan, Ki-

yevka, Okeanskaya, Roshchino on River Iman; Kuril

Isl.: Alekhino and Sernovodsk on Kunashir. Kazakh-

stan: Alma-Ata (PET). Kirgiziya: Chatkalskiy Mts.

(MOS;).

Variety: Body length 9-10 mm. Flagellum with

45-47 segments. In other characters similar to the

nominate morphe. Material: 1? Japan, Honshu,

Nagano (OTT); 393 Japan, Honshu, Kamikochi

near Norikura-dake (243 AEI, 18 Hinz/ZSM).

The status of this variety is unknown. The solu-

tion of this problem depends on the answer to the

question whether the nominate morphe of D. vigi-

lator does occur in Japan or not.

177

Dusona xenocampta (Förster, 1868)

This is a variable and poorly defined species.

Description. @: Body length 8-12 mm. Temples

distinctly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes

and temples intersect on the prescutellar groove.

Ocellar index 1.0. Frons scarcely depressed, coria-

ceous, densely rugose-punctate, with a very fine

median longitudinal carina, without radial wrin-

kles. Antennal carina low and narrow. Face and

dorsal 0.6 of the clypeus coriaceous, densely ru-

gose-punctate and rugose. Clypeus flat, coriaceous

and rather densely or dispersely punctate ventral-

ly, its apex truncate or slightly convex medially.

Malar space index 0.4. Genal carina low and

straight ventrally, joining the slightly raised oral

carina at an acute angle. Genal index 0.3. Temple

finely coriaceous and with very fine disperse punc-

tation medially. Flagellum with 45-53 segments,

filiform, the apical 0.3 distinctly attenuate, 2”! seg-

ment 2.1x, shortest segment 0.9x, penultimate seg-

ment 1.5x as long as wide. Pronotum with longi-

tudinal wrinkles laterally, the dorsolateral 0.3 with

additional punctures. Mesoscutum and scutellum

coriaceous, dull, densely or very densely punc-

tate. Scutellum with lateral carinae on the anterior

0.8. Prescutellar groove smooth. Ventral part of

the prepectal carina merging with the pleural part

or with the transverse carina, the ventral part low,

the pleural part almost complete or complete, in

the latter case joining the anterior edge of the

mesopleurum, the transverse carina slightly raised

or low and partly obliterated. Mesopleurum dis-

tinctly coriaceous, densely punctate. Speculum

finely granulate-strigose. Depression in front of

the speculum granulate-strigose or with fine long

longitudinal striae, which merge the punctation

or with a coriaceous area without punctation an-

teriorly. Metapleurum distinctly coriaceous, rath-

er densely or densely punctate. Pleural carina dis-

tinct. Areolet rather large, short-stalked, receiving

the 2" recurrent vein slightly basad of the middle.

Nervellus distinctly reclivous, weakly intercept-

ed. Nervellar index 0.4-0.6 (Fig. 29), rarely 0.7.

Tibial spur index 0.6. Hind femur 5.2x as long

as high. Hind tarsal claws short, with distinct

pectinate teeth basally and medially. Propodeum

extending to 0.3 of the hind coxae, deeply and

rather broadly depressed, with short transverse

striae in the midline of the sulcus, in other parts

irregularly wrinkled, costulae complete or obliter-

ated, longitudinal carinae often indicated medial-

Iy and posteriorly. Propodeal spiracle large, 4.4x

as long as wide. Spiracular carina narrow. Spirac-

178

ular index 0.2. 1* gastral tergite 3.8x, 2" tergite

1.9x as long as wide. Petiole with distinct but

sometimes rather small glymmae, which are bor-

dered by longitudinal carinae dorsally, smooth in

front of the glymmae. Petiolar suture fine, partly

obliterated. 1“ gastral sternite for the greater part

smooth, sometimes with transverse wrinkles pos-

teriorly. Thyridium oval. Thyridial index about

2.0. Epipleurum not separated from the 3" gastral

tergite, the latter with distinct black lateral stripes

well above the ventrolateral edge. Gaster distinct-

ly compressed posteriorly. Ovipositor index 0.5.

Colour: Black. Maxillary palpus yellowish

brown. Mandible broadly yellowish red medially.

Tegula yellowish brown to dark brown. Wings

hyaline. Stigma light brown medially. Front leg

from the tip of the coxa onwards yellowish red.

Mid femur yellowish red on the apical 0.3. Mid

tibia yellowish red. Mid tarsus dark brown. Hind

tibia yellowish brown medially, narrowly black

basally, broadly black apically. 2‘ gastral tergite

red on the posterior 0.1-0.4. 3“ and 4" tergites red.

Pilosity whitish.

d: Tegula yellow to yellowish brown. Front

and mid tibiae yellow. Hind tibia yellow medial-

ly, narrowly black basally, broadly black apically.

In other characters similar to the ®.

Material (38??, 738): Russia: Sverdlovsk Prov.: Ye-

katerinburg (= Sverdlovsk); Irkutsk Prov.: Dachnaya;

Chita Prov.: Bukuka (PET), Chita (MOS), Kurort Dar-

asun, Nerchinskiy Zavod; Yevreyskiy A.O.: Londoko;

Khabarovsk Terr.: Khekhtsir near Khabarovsk (PET);

Primorye: Spassk Dalniy (MOS), Vladivostok; Kuril

Isl.: Alekhino and Tretyakovo on Kunashir (1? from

Tretyakovo being a paratype of D. indistinctor). Kazakh-

stan: Borovoye near Kokchetav (PET), Zailyskiy Ala-

tau Mts. Kirgiziya: Issykata (MOS), Terskey Alatau

Mts. (LIN). Japan: Hokkaido: Jozankei (AEI), Maruya-

ma; Honshu: Aomori; Shikoku: Mt. Kamegamori (SAP).

Dusona yamanakai Hinz, 1985

Dusona yamanakai Hinz, 1994

Hinz (1985: 314 f., 1994: 38 f.) described this spe-

cies twice (see introduction). The descriptions of

both taxa differ from each other (in 1985 both

sexes are described, in 1994 only the ® is), and the

type series are different, the holotype (?) of the

1994 description from Sendai/Kyushu/Japan

(SAP) being mentioned as a paratype in the 1985

description. This specimen and the holotype (?)

of the 1985 description from Roshchino on River

Iman/Primorye/Russia (PET) have recently been

studied by the junior author.

Description. 2: Body length 10-13 mm. Temples

strongly constricted behind the eyes, 0.8x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons slightly depressed,

coriaceous and finely and densely punctate dor-

sally, coriaceous almost without punctures ven-

trally, with a weakly indicated median longitudi-

nal carina, without radial wrinkles. Antennal car-

ina low and narrow, the rim slightly bent upwards

(inconspicuous). Face and dorsal 0.6 of the clypeus

coriaceous, densely rugose. Clypeus flat, finely

coriaceous and with fine disperse or rather dense

punctation ventrally, its apex slightly convex me-

dially. Malar space index 0.3. Genal carina low

and straight ventrally, joining the low oral carina

at an acute angle. Genal index 0.4. Temple finely

coriaceous and with fine rather dense punctation

medially. Flagellum with 54-58 segments, filiform,

the apical 0.3 distinctly attenuate, 2"! segment

1.9x, shortest segment 0.8x, penultimate segment

1.2x as long as wide. Pronotum densely punctate

or rugose-punctate on a smooth background lat-

erally, with a few short longitudinal wrinkles on

the ventral 0.3. Mesoscutum and scutellum coria-

ceous, very densely punctate. Scutellum with: lat-

eral carinae on the anterior 0.6. Prescutellar groove

for the greater part smooth, with few longitudinal

striae laterally. Prepectal carina low, ventral part

merging with the pleural part, the dorsal 0.3 of the

latter replaced by several wrinkles. Transverse

carina obliterated. Mesopleurum with rather dense

or dense punctation on a very finely coriaceous or

smooth background. Speculum punctate posteri-

orly, very finely coriaceous and shining anterior-

ly. Depression in front of the speculum with dis-

tinct longitudinal striae, which merge with the

punctation anteriorly. Metapleurum finely coria-

ceous, densely punctate. Pleural carina complete.

Areolet large, pointed, receiving the 2"4 recurrent

vein basad of the middle. Nervellus vertical, dis-

tinctly intercepted, the upper branch slightly bent

inwards. Nervellar index 0.7. Tibial spur index

0.6. Hind femur 4.4x as long as high. Hind tarsal

claws rather long, with short pectinate teeth al-

most over their total length. Propodeum extend-

ing to 0.4 of the hind coxae, rather narrowly and

rather shallowly depressed, sometimes not de-

pressed, for the greater part rugose-punctate or

irregularly wrinkled, often with a small finely

coriaceous and shining area without punctation

anteriorly (region of the area superomedia), with

some short longitudinal wrinkles posteriorly, car-

inae almost completely obliterated. Propodeal

spiracle large, 2.8x as long as wide. Spiracular

carina widened, sometimes indistinct. Spiracular

index 0.5. 1” gastral tergite 3.7x, 2" tergite 1.7x as

long as wide. Petiole with distinct glymmae, which

are bordered by longitudinal carinae dorsally,

smooth or with very fine longitudinal striae in

front of the glymmae. Petiolar suture partly oblit-

erated. 1” gastral sternite for the greater part

smooth, with transverse striae posteriorly. Thy-

ridium elongate. Thyridial index about 1.0. Epi-

pleurum separated from the 3" gastral tergite an-

teriorly, the crease marked with black. Epipleural

index 0.4-0.5. Gaster distinctly compressed poste-

riorly. Ovipositor index 0.6.

Colour: Black. Maxillary palpus yellow. Man-

dible broadly yellow medially. Tegula yellow,

marked with brown. Wings hyaline. Stigma yel-

lowish brown medially. Front leg from the tip of

the coxa onwards yellowish red. Mid femur yel-

lowish red on the apical 0.2. Mid tibia and tarsus

yellowish red. Hind tibia broadly yellowish red

medially, narrowly black basally, narrowly medi-

um brown apically. Hind tarsus medium brown.

2"‘ gastral tergite dorsally black, laterally red on

the posterior 0.2. 3” and 4" tergites narrowly black

dorsally, broadly red laterally. Gastral sternites

red. Pilosity whitish.

d: Propodeum with the costulae partly present

and the longitudinal carinae present medially and

posteriorly. Front and mid legs from the tip of the

coxa onwards yellow. Hind tibia broadly yellow

medially, narrowly black basally, narrowly and

indistinctly tinged with brown apically. Hind tar-

sus yellowish brown. 2"! gastral tergite complete-

ly black. 3°“ and 4" tergites broadly black dorsally,

half red in lateral view. Gastral sternites and gen-

ital clasper black. In other characters similar to

the ®.

Material (322, 389): Russia: Primorye: Roshchino on

River Iman (Hinz/ZSM, PET), Spassk Dalniy (Hinz /

ZSM, MOS). Japan: Kyushu: Sendai (SAP).

Dusona yezoensis (Uchida, 1928)

The lectotype of this species, a d from Hokkaido/

Japan (SAP) without gaster, was studied by the

senior author. So far no other d has been found.

Description. 9: Body length 12-13 mm. Temples

strongly constricted behind the eyes, 0.7% as long

as the width of an eye, tangential lines on eyes and

temples intersect on the posterior half of the meso-

scutum. Ocellar index 0.9. Frons slightly depressed,

coriaceous, finely and densely rugose-punctate, a

median longitudinal carina slightly raised medi-

ally, obliterated dorsally and ventrally, radial wrin-

kles short and indistinct. Antennal carina low and

narrow. Face and dorsal half of the clypeus coria-

ceous, densely rugose-punctate. Clypeus flat or

slightly concave, coriaceous and rather densely

punctate ventrally, its apex truncate or slightly

concave medially. Malar space index 0.5. Genal

carina low and straight ventrally, joining the slight-

ly raised oral carina at an acute angle. Genal index

0.9. Temple finely coriaceous and with very fine

disperse punctation medially. Flagellum with 45-47

segments, filiform, the apical 0.3 distinctly attenu-

ae, 2 segment 1.8x, shortest segment 0.9-1.0x,

penultimate segment 1.0x as long as wide. Prono-

tum densely rugose-punctate laterally on the dor-

sal 0.3, with longitudinal wrinkles ventrally. Meso-

scutum and scutellum coriaceous, densely or very

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove for the

greater part smooth. Prepectal carina low, ventral

part merging with the pleural part, the latter com-

plete, joining the anterior edge of the mesopleu-

rum. Transverse carina present anteriorly. Meso-

pleurum distinctly and densely punctate on a finely

coriaceous and shining background. Speculum

coriaceous and granulate-strigose. Depression in

front of the speculum with distinct short longitu-

dinal wrinkles and striae, which merge with the

punctation anteriorly. Metapleurum coriaceous,

densely rugose-punctate and rugose. Pleural car-

ina distinct. Areolet rather large, stalked, receiv-

ing the 2” recurrent vein distad of the middle.

Nervellus slightly reclivous, distinctly intercept-

ed. Nervellar index 0.7. Tibial spur index 0.7. Hind

femur 5.5x as long as high. Hind tarsal claws

short, with short pectinate teeth basally. Propo-

deum extending to 0.3 of the hind coxae, shallow-

ly and rather narrowly depressed, sulcus with

irregular wrinkles and short transverse wrinkles,

with some longitudinal wrinkles posteriorly, car-

inae complete or almost complete. Propodeal spir-

acle rather small, 2.5x as long as wide. Spiracular

carina indistinct. Spiracular index 0.6-1.0. 1* gas-

tral tergite 4.0x, 2” tergite 2.1x as long as wide.

Petiole with distinct glymmae, which are bordered

by longitudinal carinae dorsally, smooth or with

very fine sculpture in front of the glymmae. Peti-

olar suture complete. 1” gastral sternite coriaceous

and finely rugose. Thyridium oval. Thyridial in-

dex about 2.0. Epipleurum separated from the 3'4

gastral tergite, the crease marked with black. Epi-

pleural index 0.8. Gaster distinctly compressed

posteriorly. Ovipositor index 0.9.

Colour: Black. Mandible narrowly tinged with

reddish brown medially. Wings hyaline. Stigma

yellowish brown medially. Front femur black ba-

sally and ventrally, yellowish red dorsally and

180

apically. Front tibia yellow. Front tarsus yellow-

ish brown. Mid femur narrowly yellowish brown

apically. Mid tibia dark yellowish brown or dark

brown. 2" gastral tergite completely black or red

on the posterior 0.1. 3 and 4" tergites red, marked

with black dorsally (to a variable extend). Pilosity

whitish.

d not available for description.

Material (1822): Russia: Khabarovsk Terr.: Khekhtsir

near Khabarovsk; Primorye: Andreyevka, Khasan,

Nakhodka, Spassk Dalniy, Ussuriysk; Kuril Isl.: Ale-

khino, Dudovoye and Yuzhno-Kurilsk on Kunashir

(PET). Japan: Hokkaido: Maruyama (SAP), Sapporo

(AEI, Hinz/ZSM); Kyushu: Miyazaki (SAP). The spe-

cies also occurs in Europe: 19 Russia, Kaluga Prov.,

Borovsk (PET).

Dusona yezoensoides Hinz & Horstmann,

spec. nov.

Holotype (?): Japan, Hokkaido, Kaributo, 22.6.

1966 (HEL).

Description. ?: Body length 13mm. Temples

strongly constricted behind the eyes, 0.6x as long

as the width of an eye, tangential lines on eyes and

temples intersect on the anterior half of the meso-

scutum. Ocellar index 0.6. Frons slightly depressed,

coriaceous, finely rugose-punctate, with a fine

median longitudinal carina, with short and indis-

tinct radial wrinkles. Antennal carina low and

narrow. Face and dorsal half of the clypeus coria-

ceous, very densely punctate. Clypeus flat, coria-

ceous and rather densely punctate ventrally, its

apex truncate medially. Malar space index 0.4.

Genal carina low and almost straight ventrally,

joining the low oral carina at an acute angle. Genal

index 0.6. Temple coriaceous and finely and rath-

er densely punctate medially. Flagellum with 48

segments, filiform, the apical 0.4 distinctly attenu-

ate, 2"! segment 2.2x, shortest segment 1.2x, pen-

ultimate segment 1.7x as long as wide. Pronotum

coriaceous and densely rugose-punctate laterally

on the dorsal 0.2, with longitudinal wrinkles ven-

trally. Mesoscutum and scutellum coriaceous,

densely punctate. Scutellum with lateral carinae

on the anterior 0.2. Prescutellar groove almost

smooth, with some very fine longitudinal striae.

Prepectal carina low, ventral part merging with

the pleural part, the latter complete, joining the

anterior edge of the mesopleurum. Transverse

carina obliterated. Mesopleurum distinctly coria-

ceous, rather densely or densely punctate. Specu-

lum coriaceous and granulate-strigose. Depres-

sion in front of the speculum with distinct longi-

tudinal wrinkles, which merge with the puncta-

tion anteriorly. Metapleurum coriaceous, densely

rugose. Pleural carina distinct. Areolet large, point-

ed, receiving the 2” recurrent vein distinctly ba-

sad of the middle. Nervellus distinctly reclivous,

distinctly intercepted. Nervellar index 0.5. Tibial

spur index 0.6. Hind femur 6.2x as long as high.

Hind tarsal claws slightly elongate, with distinct

pectinate teeth basally and medially. Propodeum

extending to the middle of the hind coxae, slightly

and narrowly depressed, with complete costulae

and longitudinal carinae, the latter bordering the

longitudinal furrow, for the greater part irregu-

larly wrinkled, with some short transverse wrin-

kles in the longitudinal furrow. Propodeal spira-

cle large, 5.3x as long as wide. Spiracular carina

narrow. Spiracular index 0.4. 1" gastral tergite

5.8x, 2" tergite 2.6x as long as wide. Petiole with

rather small but deep glymmae, with very fine

longitudinal striae in front of the glymmae. Petio-

lar suture very fine. 1* gastral sternite for the

greater part smooth, with transverse striae poste-

riorly. Thyridium elongate. Thyridial index about

1.5. Epipleurum separated from the 3% gastral

tergite, the crease marked with black. Epipleural

index 0.8. Gaster strongly compressed posterior-

ly. Ovipositor index 0.9.

Colour: Black. Maxillary palpus dark brown.

Mandible rather broadly marked with yellow me-

dially. Wings hyaline. Stigma dark brown medial-

ly. Front femur yellowish red dorsally, dark brown

ventrally. Front tibia and tarsus yellow. Mid fe-

mur yellowish brown on the apical 0.1. Mid tibia

yellowish brown. 2" gastral tergite with a black

posterior edge, narrowly marked with red in front

of the posterior edge. 3" tergite red. 4" tergite red,

marked with black dorsally and posteriorly. Pi-

losity whitish.

d unknown.

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181

Index of hosts falcator 26, 32, 70

filator 5

Allophyes oxyacanthae 102 flagellator 24, 71,72

Cabera exanthemata 62 flavitarsis 14, 72

Cabera pusaria 62 Navopicta 34, 73

Colotois pennaria 102 foersteri 51

Compsoptera opacaria 102 fundator 22, 31, 38, 74

Heliothis viriplaca 53

ES gastator 9, 12, 74

Hypomecis punctinalis 49, 156

gastroides 13, 75

Lasiocampidae 148 genator 23, 76, 128

Orgyia postica 62 glauca 38, 77, 78

Semiothisa liturata 83 graptor 29, 30, 31, 78, 79

humilis 33, 79

Index of Dusona species impressifrons 21, 80

inconspicua 30, 37, 81

admontina 6, 8, 13, 39 indistinctor 33, 81, 178

aemula 10, 23, 40 inermis 24, 71, 82

affinitor 27, 40 infesta 8, 12, 83

alpinator 31, 32, 41 infundibulum 12, 84

alticola 19, 42 insignita 29, 85

americana 4, 34, 35, 36, 43, 44 iwatae 13, 85

amictor 135 i j

amurator 26, 27, 36, 38, 44 TaRDtaen 48, 7,8

anceps 20, 45 ana. 00

angustifrons 6,7, 8, 10, 28, 45, 57, 106 Re

annexa 27,35, 43, 44, 46 kasparyani 23, 88

auriculator 10, 20, 23, 47 kyushui 103

aurita 7,19, 48 lamellifer 28, 89

bellator 14, 49 latisulcor 77

bellipes 6, 13, 50 leptogaster 13, 90

belokobyli 27, 51 levibasis 37, 90

bicoloripes 13, 51, 52, 59, 160 liberator 24, 91

bucculenta 27, 32, 53 limnobia 46

bucculentoides 32, 54 limnobius 46

caliginosa 38, 77, 78 ee in, 22

£ lividariae 25, 93

capitator 6, 31, 54 j

longicauda 17, 93

carinator 14, 55 i

carinifrons 8, 9, 23, 56 een g Ge

carpathica 22, 24, 57 ga 2

caudator 16, 17, 58 mactatoides 20, 21, 96

celator 21, 58 mactator 21, 96

chabarowski 13, 52, 59, 160 maruyamae 14, 97

chechziri 12, 60 maruyamator 15, 98

chinensis 19, 23, 25, 61 matsumurae 20, 99

circumcinctus 8,9, 26, 62, 153 melanator 4, 11, 12, 29, 78, 79, 100

collaris 22, 63 melanocerus 62

confusator 32, 63 memorator 19, 101

constantineanui 26, 64 mercator 3,5,7, 10, 25, 101

contumator 31, 33, 65 meridionator 9, 27, 31, 34, 35, 36, 103, 105

contumax 13, 66 meritor 35, 104

coriacea 15, 67 mesoxanthus 101, 102

crassiventris 6, 20, 67 micrator 8, 34, 104, 105, 121, 150

cultrator 12, 68 minor 24, 28, 105, 106

detritor 16, 18, 69 a oe

dictor 28, 70 momoii 16, 18, 107, 108

montana 7,8,17, 18, 108

myrtilla 11, 109

182

nanus 35, 109

nebulosa 36, 110

nervellator 31, 111

nidulator 10, 16, 112

nigridens 30, 31, 112, 117

nigridorsum 10, 33, 37, 113

nigrifemur 62

nigrina 11, 114

nigrotibialis 12, 115

nitidipleuris 33, 37, 38, 115

norikurae 29, 78, 79, 116

obliterata 33, 36, 118

obscurator 12, 16, 119

obscuripes 24, 120

obtutor 35, 38, 105, 120

okadai 20, 121

opaca 8, 34, 36, 39, 122

opacoides 36, 39, 122, 123

oxyacanthae 101

oyamadai 4, 43, 44

papator 18, 124

parva 37, 109, 124

parvicavata 20, 34, 125

peptor 29, 117, 126

perditator 23, 127

perditor 10, 23, 127

petiolatoides 28, 76, 128

petiolator 8, 27, 28, 129

petitor 24, 130

pictator 16, 18, 19, 131

pineticola 6, 20, 132

polıta 15,16, 17, 19, 132

propodeator 20, 133, 134

pseudobucculenta 32, 134

pugillator 12, 14, 21, 51, 135

pulmentariae 31, 32, 136

punctus 144

radiator 28, 34, 137

recta 21, 30, 137

rectator 30, 138

rectoides 29, 30, 139

remota 38, 77, 140

rossica 72

rubator 31, 32, 141

rufator 23, 141

rufiventrator 25, 142

rufoapicalis 37, 143

rufovariata 14, 30, 78, 79, 144

rugifer 14, 15, 144, 160

rugifrons 21, 145

rugosa 15, 146, 147

sachalini 4, 6, 9, 12, 147

sasayamae 18, 148

scalprata 25, 149

schikotani 33, 34, 36, 105, 149, 150

scolator 24, 150

scriptor 36, 151

scutellatoides 135

scutellator 21, 36, 152

setator 32, 153

sibirica 20, 132

signator 26, 153

similator 13, 15, 17, 18, 154

sobolicida 6, 20, 25, 155

specularis 14, 17, 156

spinator 17, 18, 157

spinipes 25, 158

spiracularis 22, 27, 159

splenditor 25, 28, 159

stragifex 15, 52, 59, 144, 160

stygia 31, 35, 74, 161, 165

subimpressa 26, 36, 38, 162

temnator 27, 163

templator 35, 164, 165

temporalis 28, 165

tenuis 7,8, 26, 103, 166

terebrator 16, 17, 18, 166

ternata 36, 167

thomsoni 23, 26, 168

tibiatoria 25, 169

tritor 26, 170

ucrainator 19, 22, 170

ucrainica 19, 171

variator 8, 33, 37, 38, 115, 172

varıpes 26, 173

venitor 30, 174

ventrator 37, 174

vidua 11, 175

viduator 11, 176

vigilator 14, 30, 177

xenocampta 8, 31, 33, 81, 178

yamanakai 4, 14, 29, 178

yezoensis 17, 18, 179

yezoensoides 18, 180

183

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