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TRANSACTIONS
OF THE
WISCONSIN ACADEMY
SCIENCES, ARTS AND LETTERS
0mM miin^
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zih-t^'^a
NATURALE SPECIES RATIOQUE
MADISON, WISCONSIN
1926
Vol. XXII of the Transactions of the Wis¬
consin Academy of Sciences, Arts, and Letters
is issued under the editorial supervision of the
Secretary.
CHANCEY JUDAY,
Secretary.
TRANSACTIONS
OF THE
WISCONSIN ACADEMY
OF
SCIENCES, ARTS AND LETTERS
VOL. XXII.
MADISON, WISCONSIN
1926
/
CONTENTS
Stagecoach and Tavern Days in the Baraboo Region. IT. E.
Cole _ 1
A Statistical Study of Lead and Zinc Mining in Wisconsin.
(With twenty-two Figures.) Selma Langenhan
SCPIUBRING _ 9
Fauna of the Galena Limestone near Appleton (With Plates
I-III.) John W. Ockerman _ 99
Some Characteristics of the Bacteria of Lake Mendota.
(With Plate IV.) Laetitia M. Snow and E. B. Fri:d 143
Notes on Parasitic Fungi in Wisconsin, XII. J. J. Davis __ 155
Notes on Parasitic Fungi in Wisconsin, XIII. (With one
Figure.) J. J. Davis _ 165
Notes on Parasitic Fungi in Wisconsin, XIV. J. J. Davis__ 181
Nomenclatorial Notes on American Fresh Water Mollusca.
Frank Collins Baker _ _ 193
Revision of the Nearctic Species of Ilelophilus and Allied
Genera. (With Plates V-VII.) C. H. Curran and
C. L. Fluke _ 207
The Use of Photo-electric Cells of Different Alkali Metals
and Color Screens in the Measurement of Light Pene¬
tration into Water. (With nine Figures.) AActor E.
Shelford and Jakob Kunz _ 283
A Third Report on Limnological Apparatus. (With eight
Figures.) Chancey Juday _ 299
The Rotifer Fauna of Wisconsin, III. A Revision of the
Genera Lecane and Monostyla. (AVith Plates AMII-
XLAMI.) H. K. Harring and F. J. Myers _ 315
Proceedings of the Academy, 1924 and 1925 - 425
List of Officers and Members - 431
Charter of the Academy _ 443
STAGECOACH AND ‘ TAVERN DAYS IN THE
BARABOO REGION
H. E. Cole
“Whoe’er has travell’d life’s dull round,
Where’er his stages may have been,
May sigh to think he still has found
The warmest welcome at an inn.”
Baraboo and the vicinity about is especially favorable for a
study of stagecoach travel and of taverns, for the reason that
the railroad did not penetrate the picturesque and rugged re¬
gion until 1871. Prior to this date stagecoaches of various types
were familiar objects in the locality and a convenient means of
transportation.
When the writer came into the state, some of the drivers of
coaches were still active and able to impart information regard¬
ing their routes and the places of entertainment. At one time
eight stages left the village of Prairie du Sac daily. None is
now making trips, this primitive activity having been entirely
supplanted by other modes of travel.
On the old Concord coaches and kindred vehicles, the driver
was the important personage of the equipage, the autocrat of his
day. Everybody of note in the locality was known to him and
at meal-time at the tavern, it was the driver who presided at
table where the passengers on the coaches were hurriedly re¬
freshed at the reasonable price of twenty-five cents per person.
His aspect was usually rougher than a close acquaintance with
him as an individual revealed. A flannel shirt, corduroy
breeches stuffed into high boots, a well-worn hat or cap and a
fur or leather coat in inclement weather made up his usual cos¬
tume. In personal appearance he might be tall and lean or
short and stout. In either case he was always alert to meet the
exigencies that were likely to arise as he traveled the country
roads. In speech he was more picturesque than grammatical,
often voicing choice bits of humor or arguments that were
graphic and convincing.
Artists of a century ago frequently pictured the horses of the
1
2 Wisconsin Academy of Sciences, Arts, and Letters.
stage-coach as prancing, fire-breathing steeds, but upon investi¬
gation it is found that much of this picturesqueness was imagi¬
native. It required highly practical teams to pull the coaches
loaded with passengers and baggage, and the long journeys con¬
spired to produce conspicuous ribs, pronounced backbone and
other indications of arduous toil.
It was the custom of the time for the driver, when he gathered
up his lines in front of the tavern, to crack his whip with a
flourish for the start from the hostelr3^ With a great hurrah
and whirring of wheels they would set out, then often would
creep along when muddy stretches or sandy wastes were encoun¬
tered. As the destination drew near there was again a great
flurry on the part of the driver, as he brought his horses to a
stop before the tavern.
Besides looking after the welfare of his passengers and caring
for his horses, the driver often had packages entrusted to his
care, a weighbill accompanying, the landlord collecting when de¬
livered to the person addressed. To tip the driver wilh coin
was rarely or never known but a drink was usually accepted
when it was offered.
Among the passengers, when the stage lines were opened, were
lawyers, doctors, preachers, newspapermen, lumbermen and
settlers seeking places of abode in the new country. There
came, too, the homeseeker from across the sea in singular garb,
clinging tenaciously to his queer looking baggage. In the fifties
and sixties many raftsmen floated down the Wisconsin River
to various snubbing-posts along the stream, seeking the highway
with its convenient stagecoach upon their return. During the
Civil War, soldiers were often passengers from the railway sta¬
tion to their homes in the interior, bringing many a thrilling
tale of the sanguniary conflict in the southland and not infre¬
quently an empty sleeve or trouser ^s leg. In this region during
the frenzied hop-period, pickers arriving at Kilbourn and Mazo-
manie by the car load, were conveyed to the hopyards by the
stagecoach. Of hoppole pullers there were also not a few.
Not infrequently suspicious characters would seek transporta¬
tion. Of this class, however, few had an eye on the passengers
or stage property. As to horses, there were many more spirited
steeds to be had in the open pasture or in unlocked stalls. D.
Joseph Johnson, who drove the stage between Prairie du Sac
and Mazomanie for a number of years after his return from
Cole — Stage Coach Days in the Baraboo Region. 3
Southern battlefields at the close of the Civil War, brought more
than one traveler into Prairie du Sac whom he never saw again,
who probably rode a horse into Illinois to be sold to some unsus¬
pecting purchaser.
Besides those mentioned there were the peddlers with their
packs and that group of idlers known as speculators, in prosper¬
ous dress, shrewd of eye and clever in trades.
Prescott Brigham, brother of Ebenezer Brigham of Blue
Mounds, established the first stage line between Madison and
Sauk County, the route terminating at Sauk City and Prairie
du Sac. He drove the first stagecoach into the county in 1844,
as proprietor, jehu, mail and express man, four years before
Wisconsin passed into statehood.
After the line was opened between Madison and Sauk, the
name given the two villages on the Wisconsin river, a network
of lines spread to all parts of the county, every hamlet enjoy¬
ing the visits of the stage.
A great variety of vehicles have been used in transporting
passengers, mail and baggage over the Baraboo region. The
most imposing was the Concord coach wdth its painted body and
plush seats; One of these employed on the Lodi-Madison route
was know'll as the Prairie Queen. Various names w'ere used to
designate these coaches, each of which accommodated nine per¬
sons in the interior, with room on the driver’s seat for one or
two w'ho might prefer to ride outside.
The question has occasionally been asked, what became of
these famous vehicles when they passed to worthless property
with the advent of the railroad? Concerning some of these
faithful friends of our forefathers, the Concord coaches, W. W.
Warner of Madison, in his youth a resident of Baraboo, wrote
as follows :
“This was, I should say, about 1868-69. Who among the
boys who participated in that famous escapade, may ever for¬
get? Be it knowm there were fifteen or twenty antique, super¬
annuated Concord stagecoaches which had been one after an¬
other placed, so to speak, in dry dock and out of commission,
having outlived their further transporta tional usefulness, and
thus they were housed in a rambling series of sheds, just back
of the present city hall. We young chaps, the day after a
Fourth of July celebration, conceived the idea of decorating
Oak Street wdth the dilapidated vehicles. Some of the chariots,
4 Wisconsin Academy of Sciences, Arts, and Letters.
I remember, bore euphoneous names, such as Argosy, Prairie
Queen and Western Monarch. Those who remember the one¬
time resplendent coaches, gorgeous beyond the dreams of a
Ringling circus creation, will recall that they were integers
connecting Baraboo with relatively near-by points of the out¬
side world, such as Madison, Mazomanie, Portage and Kilbourn.
* *
‘ ‘ It was long after midnight when we scamps, as expeditiously
and as quietly as possible, hauled forth a score of the nonde¬
script vehicles from their moorings, to the Western Hotel street
corner, and thence made an imposing string of them, reaching
almost to the present post office site, and a fine spectacle they
presented early next morning. Not many citizens of Baraboo
were aware that such antediluvian chariots were in existence,
much less that they were right here in Baraboo. The general
astonishment, therefore, may well be imagined. What oppor¬
tunities were lost in their destruction shortly after this, their
last appearance, for securing matchless museum antiques.’’
There is a fiavor of romance about old time taverns. Cling¬
ing to these primitive places of entertainment in this locality
are rich memories of the middle nineteenth century. They fol¬
lowed closely the advancing rim of pioneer life as it spread
over the section north and west of the Wisconsin River in the
forties and fifties, when the Baraboo country was a railroadless
region, when the snail-like mode of travel of the stagecoach
made it necessary for places of entertainment to be within brief
distances of each other that the traveler might be conveniently
refreshed.
Some of these old hostelries were crude and queer, and the
bonifaces who presided over them often seemed a kind of re¬
flection in appearance and character of the strange architecture
of the buildings. These early landlords did not confine their
responsibility to the physical comfort of their guests, but were
a never failing thesaurus on politics, crops, the weather and the
gossip of the country roundabout. They were also prone to
join their guests in the drinking of intoxicating liquors which
was the besetting sin in the early days of the region. Many of
the settlers seem to have been perpetually athirst. Frequently
there was an '^eye-opener” upon arising in the morning, an ap¬
petizer before the noonday and evening meals, afterwards a
Cole — Stage Coach Days in the Barahoo Region. 5
little stimulant to aid digestion, and a “nightcap” before re¬
tiring.
At barn raisings, husking bees, and other gatherings a jug
was always in evidence and occasionally there was tippling at
funerals. Little wine was used ; whisky made from corn or rye
at a distillery near at hand was the popular beverage.
An example of the crude hospitality extended to early travel¬
ers at the tavern is given in ‘ ‘ Merrell ’s Narrative, ’ ’ in the Wis¬
consin Historical Collections. Merrell and Captain Harris
were journeying from Fort Winnebago to Galena and stopped
at Rowan’s tavern near Madison. Merrell says:
“The first night we stopped at Rowan’s celebrated house,
thirty-five miles from the fort. I had heard much of this inn
and found it filled the bill. It consisted of two log buildings
with an open space between, all under the same roof. After
taking care of our horses and getting something to eat, we en¬
quired where we could sleep, and Madam told us in the other
house so we went in and concluded we should do very well as
there was nothing in the room but a bed, and one or two three-
legged stools. After lying down and by the time we w^ere ready
to go to sleep, there was an unearthly squeal and grunt of hogs
in the open space between the two rooms, only a partition of
logs between our heads and them. I was told that Governor
Doty once stayed there, and after supper, as was the custom,
rolled himself in his blanket on the floor. The family all lived,
cooked and slept in one room ; and in the night the governor
felt something poking about him and found it was a pet pig the
children had running about the house. The governor felt of the
puncheons of the floor, and found one loose, which he raised
carefully, and grabbing the pig thrust him under, and was re¬
lieved of his company that night.”
In none of the pioneer taverns was there bathing facilities,
except for the face and hands, as bathtubs were unknown in the
region prior to the coming of the railroads. For cleansing the
face and hands a sink with a small wash-basin was provided.
Near by was a dish of soft soap, a roller towel and a comb which
frequently had lost not a few of its teeth.
Sleeping rooms were small and heatless and without ventila¬
tion except that which was afforded by a door or undersized
window. The foundation of the bed was a tick filled with straw
thrown upon a corded bedstead, the ropes running backward
6 Wisconsin Academy of Sciences, Arts, and Letters.
and forward from pins in the rails and end-pieces. For illum¬
ination at night, tallow candles were used before the advent of
the kerosene lamp. Circular lanterns of tin or square ones with
parts of the sides of glass, a candle for the interior, were com¬
mon up to and through the Civil War period. During very
cold weather a warming pan was occasionally used to heat the
beds to a comfortable temperature. This was a receptacle with
a long handle in which live coals were placed, and served its
purpose very well
The food served in the taverns was abundant but usually
simple. Plenty of bread and butter, meat, potatoes, gravy,
beans and other staples were provided. Fruit and game were
served in season. Conservation wardens were unknown; the
bag limit of quail, pigeons, partridge, rabbit, prairie chicken,
deer, bear or fish was unlimited. Buckwheat cakes were in evi¬
dence in the morning with the accompaniment of coffee. Crack¬
ers were about the only ingredient of a meal that came from a
bakeiy and were usually purchased by the barrel. Maple syrup
was on the menu in the spring and sorghum during the re¬
mainder of the year. Sour milk and saleratus, or home-made
yeast were the sources of raised breads and biscuits made from
wheat fiour and corn bread made from ground maize.
Cooking facilities were of course more or less primitive. The
first stoves for kitchens did not appear on the market until
about 1840, just prior to the opening of the first taverns in the
Baraboo region. These were brought from Milwaukee and sold
by those outfitting the new settlers. They had elevated ovens
but not reservoirs for water as later models had. The kitchens
were equipped with a limited number of iron pots and spiders
and earthern vessels.
Cooks were often at their wits-end to find material for the
interior of pies; green tomatoes, dried pumpkin, dried apples
and like substitutes for fresh fruit were used. To reduce apples
and pumpkins to a state where they could be preserved for in¬
definite periods, slices were hung in festoons about the fire and
dried.
Dances were frequent in many of these old time taverns.
There sat the fiddler and as he played “his head swung time,
his body rocked time, his feet patted time, his eyes winked time,
and his teeth ground time. The good women laughed for joy,
and men winked at each other and popped their fists.’’ Of
Cole — Stage Coach Days in the Barahoo Region.
7
course the company danced. In the very early places of enter¬
tainment there was a rough floor but in the later ones great ball
rooms were provided at the top of the building, so constructed
as to vibrate with the movements of the merry dancers. In that
day it was the lively minuet, the graceful waltz or the old fash¬
ioned quadrille.
Many amusing incidents in connection with the early days in
the region are affiliated with these old taverns. In the Empire
House at Prairie du Sac, afterwards operated by Oscar E.
Briggs, a relative of the famous cartoonist, there once was a
diverting occasion. A circus reached the village early one Sun¬
day morning and no sooner were the circus folk at liberty than
they plunged into the broad Wisconsin River, the surface of
which gleamed in the June sunshine. After enjoying a plunge
in its cool waters, they threw themselves on the bank to hear the
martial strains of a brass band borne on the air. They were in¬
formed there was a picnic a mile away and they joined the pro¬
cession.
The day lay before the showman. The circus would not ex¬
hibit until Monday and the ‘‘kinkers,’’ ‘‘wind-jammers,^’ freaks,
sword-swallowers, “dog-boys,” and sideshow barkers bounded
along with rising spirits to the place of revelry and entertain¬
ment.
As the spigots turned and the amber fluid began to flow,
the crowd increased faster than Palstaff’s “men in buckram.”
One cooling draught'and the acrobats fell to performing marvel¬
ous feats and their agility tickled as well as shocked their audi¬
ence.
The drummer of the brass band had placed his instrument on
its circular dimension, slightly at one side of the hilarious
throng. Noticing the drum, a devilish idea entered the maud¬
lin brain of one of the “kinkers.” Quicker than a flash, flip¬
ping one foot in the air, he flopped the instrument on its side
and with a bound, turned a somersault upon it, thrusting both
feet through the tightly stretched head in the operation. This
feat ended the acrobatic performance at the grove. Gathering
themselves together, the merry Andrews marched gaily, if not
in soldierly manner, toward the Empire House in Prairie du
Sac, emitting a continuous “oomp, oomp, oomp, ” as though
beating an aggregation of imaginary drums as they staggered
along. And thus they filed into the old tavern. The leader.
8 Wisconsin Academy of Sciences, Arts, and Letters.
looking neither to the right nor left, marched straight into the
dining-room, turned partly about the movable seat at the end
of the table, and stepped waveringly upon the chair. Then,
without hesitation, he strode the length of the table amid a din
of rattling, breaking dishes, every inebriated individual in his
train following.
Dining-room girls dashed to the open and their cries brought
the tavern loafers to the scene of devastation.
Having delivered themselves of this inurbane incivility, the
erstwhile picnickers, now unable to distinguish the hour on the
face of the tavern clock, dropped into insensibility. Later they
paid for the damage and departed from the quiet village of
Prairie du Sac declaring they had never so enjoyed themselves.
A STATISTICAL STUDY
OF
LEAD AND ZINC MINING IN WISCONSIN
Selma Langenhan Schubring, Ph. D.
I
Location and Area. The lead and zinc mines of Wisconsin -
are in Grant, Iowa, and Lafayette counties, and comprise the
most important part of the ore deposits of the Upper Mississippi
Valley region. The whole region embraces the southwest corner
of Wisconsin and adjacent portions of Illinois and Iowa, i. e. :
Jo Daviess County in Illinois, and an irregular narrow belt of
territory in Clayton, Allamakee, and Dubuque Counties in Iowa,
parallel with the Mississippi Kiver. The extreme length of this
region, from east to west, is eighty-seven miles; and its greatest
width, from north to south, fifty-four miles.^ In Wisconsin the
region comprises an area of about 2,000 square miles and is
bounded on the north by the northern outcrop of the Galena
limestone, running parallel with the main watershed from the
Mississippi to the Blue Mounds ; on the east by the Sugar River ;
on the south by the State line, and on the west by the Mississippi
River. The area thus included is that of the Galena limestone,
which covers about 1,776 square miles.^ The productive area
is somewhat triangular in shape, and the center of productive¬
ness lies a little to the south of the center of geographical dis¬
tribution.^
Ores. The ores from which lead and zinc are obtained are
galena, the sulphide of lead; sphalerite or zinc blende, the sul¬
phide of zinc; smithsonite or zinc carbonate; and (to a small
degree) calamine, a hydrous zinc silicate. The galena is non-
argentiferous and the zinc ores are free from cadmium.
^ Owen, D. D., Report of a Geological Exploration of part of Iowa, Wiscon¬
sin and Illinois, made in 1839. Document No. 239, House of Representatives,
26th Congress, First Session, p. 26.
* Strong, Moses, Geology and Topography of the Lead Region, Chamber¬
lin’s Geology of Wisconsin, 1873-79, Vol. 2, p. 689.
® Chamberlin, T. C., Geology of Wisconsin 1873-79, Vol. 4, p. 398.
10 Wisconsin Academy of Sciences, Arts, and Letters.
II.
The Upper Mississippi Valley Region.
Exhibit I of this study is a table which shows statistics of
annual lead production of the Upper Mississippi Valley from
1821 to 1916, inclusive. In parallel columns and in chronologi¬
cal order appear data as presented by the several authorities
who have compiled production statistics.
Accompanying the table is a bibliography with explanatory
notes analyzing in detail the contribution made by each author¬
ity. Accredited sources are enumerated; the nature of the
product is indicated: whether it is metallic lead or, as in some
instances, lead ore; and the specific region of production is de¬
fined. Statistics for the Upper Mississippi Valley usually in¬
clude production from Wisconsin, Iowa, and Illinois, although
sometimes they include also the production of Kentucky and of
Virginia. On the other hand, southern Illinois is sometimes
excluded because of different mineralogical and structural en¬
vironment which relates southern Illinois to the Kentucky -Illi¬
nois fiuorspar district rather than to the soft lead region. In
these cases Upper Mississippi Valley production comprises only
lead from Wisconsin, Iowa, and northern Illinois, all of which
is non-argentiferous, or soft, lead. Other information is con¬
tained in the notes, and the bibliography should be used in con¬
nection with the table.
Exhibit II is a similar table giving statistics of annual
zinc production of the Upper Mississippi Valley from 1860 to
1916, inclusive. Accompanying the table is a bibliography with
explanatory notes analyzing in detail the figures p . sented.
III.
The Rise and Decline of Lead Production and the Rise
OF Zinc Production.
The history of lead mining has been closely interwoven with
that of zinc mining since the earliest development of zinc pro¬
duction. The ores are found in the same region and are often
obtained from the same openings. As a lead district, the region
lias been known from the earliest settlement days, but zinc did
not become important until the building of smelters near by
n;',!SS!7Si'SSS.''£;!., .. ,».
Exhibit II. Zinc production of the Upper Mississippi Valley.
(a) (b) (c) etc. These letters refer to notes on accompanying Bibliography of Statistics of Zinc Production of the Upper Mississippi Valley.
Note. — The figures in bold face type represent arithmetical computations by the writer, such as conversion of pounds to short tons; addition of several parts to make a whole: subtraction of one quantity from another.
* 76,042,911 pounds ••Incorrectly^ven 0371,331. 1 13. 800,000 pounds.
Schuhring — Lead and Zinc Mining in Wisconsin. 11
made it possible to separate the spelter. The production of
spelter in the United States began in an experimental way in
1858, and regular production began in 1860. Prior to this time
the zinc ores of Wisconsin were thrown on the dumps and
spread over the roads, ^ from which they were gathered later
when their value had become known, and shipped to smelters.
The first zinc ore to be mined was the carbonate found in asso¬
ciation with the lead above ground-water level. It was known
that the carbonate gave place to zinc blende and that lead ex¬
isted below the water table, but it was deemed unprofitable to
mine them. The higher price of zinc about 1900 led to the open¬
ing of some of the deposits below water level, and since that
time zinc mining has increased until Wisconsin ranks among
the largest of the producing states. Lead is still mined with
the zinc, but its production is of small importance when com¬
pared with that of zinc. As a lead producer Wisconsin is now
unimportant.
Figure 1, representing the output of metallic lead in the
Upper Mississippi Valley by decades 1821-1900, and 1907-1916,
shows the rise and decline of lead production. The decade of
maximum production was 1841-1850. Annual statistics as pre¬
sented in Exhibit I show an increase of production, with slight
fluctuations, from 1821 to the years 1845-7, after which a
marked decline set in. These minor variations in output were
due in the main to the unsettled character of the country, the
migratory habits of the settlers, and Indian disturbances. The
military operations of the Black Hawk War (1831-2) removed
the barrier of hostile tribes and made the region known for its
rich farming lands.® The resulting influx of population is re¬
flected in the increased lead production which reached its height
in 1847, as above stated. The decline which set in in 1848 and
continued permanently until a few years ago when, as a by¬
product of zinc, lead showed a slight revival, was said to be due
to the rush to the California gold fields in 1848 ; to the enlist¬
ment of miners as soldiers in the Mexican War; and to the
growing importance of agriculture and industry. Whitney® in
1854 stated that the decline was due partly to the superior at-
* Buckley, E. R., in The Mineral Industry, Vol. 9, p. 665.
^ Libb5% O. G., Transactions "Wisconsin Academy of Sciences, Arts and
Letters, Vol. 13.
® Whitney, J. D., Metallic Wealth of the United States, p. 423.
12 Wisconsin Academy of Sciences, Arts, and Letters.
tractions of the California gold fields, but still more to the ex¬
haustion of the lead deposits. Daniels’" in the First Annual
Report on the Geological Survey of Wisconsin took exception to
this statement and said that injury had been done the mining
interests by the report of exhaustion of the mines. He stated
that the real causes of diminished production were, first, the an¬
nouncement of the discovery of gold in California, which drew
off one-third of the whole mining force; second, the prevalence
1823-1830
1831-1840
1841-1850
1851-1860
1861-1870
1871-1880
1881-1890
1891-1900
Thousands of short tons
0 50 100 150 200
— No data for 2
years.
Ore. Wisconsin only.
No data for 1 year.
No data for 8 years.
Short Tons
19,512
88,878
195,860
115,340
64,198
56,924
37,779
7,740
1907-1916
35,127
Fig. 1. Production of metallic lead in Upper Mississipi Valley.
Data from Exhibit I, column 11.
of an irregular and inefficient method of working the veins;
and, less important, the illiberality of non-resident land owners,
who in some localities demanded exorbitant rents, and desired
to throw all risks upon the miners, thus discouraging produc¬
tion. Daniels pointed out some ill effects of the system of indi¬
vidual mining which then prevailed, and said that such working
answered well enough as long as there was surface ore enough
to pick up; but that when the superficial deposits were ex-
^ Daniels, Edward, First Annual Report on the Geolog-ical Survey of Wis¬
consin, 1854, pp. 40 fC.
ScMihring — Lead and Zinc Mining in Wisconsin. 13
hausted it was inadequate, and a falling off in the product could
not fail to follow. The working of the mines by single individ¬
uals or small and temporary combinations, with little capital at
their command, was entirely with a view to immediate results.
The working of veins at any considerable depth involved an
expenditure too large for any except companies with large capi¬
tal. Moreover, single individuals could not afford to work
through the unproductive portions which separated productive
leads; and thus veins remained undiscovered. The system of
individual mining was also a serious obstacle in the way of
drainage. Owing to the limited size of claims, one mine often
could not be drained by a pump or level without receiving the
water from several others belonging to various owners. This
fact retarded operations, for there was no co-operation and no
one was willing to bear the whole expense of drainage where all
shared its benefits. Daniels pointed out that individual mining
could be profitable for short periods only and that ‘^mining,
like manufacturing, requires for its successful prosecution, sys¬
tematic, comprehensive, and long continued application of labor
and capital. ’V He pronounced the mines, in 1854, to be in a
transition state, and adds :
They have passed through the period of excitement, when chance
rewarded the fortunate with rapid gains, to a more quiet and settled
condition, in which rude and individual attempts at mining are at¬
tended with frequent failure and occasional success. They now await
the period of organization to which their full treasures are to be
surrendered.
The production statistics of subsequent years bear out the
statements of both of these writers. As far as lead was con¬
cerned, Whitney’s statement was correct. The mines were prac¬
tically exhausted. On the other hand, the conclusions of Daniels
were verified, too; for, with the availability of capital for the
building of smelters near by, it became possible to utilize the
zinc ores, and the mining industry revived. Still later, with
the institution of more scientific methods and the organization
and consolidation of capital, the industry began to thrive with
zinc as the principal source of revenue.
Whitney and Daniels made their reports in 1854. That a lull
in the mining region ensued is borne out by the statistics of lead
production. About fifteen years later interest in mining was
somewhat stimulated by the legislative provision of 1870 for a
survey of the lead region. This act originated in a desire on
14 Wisconsin Academy of Sciences, Arts, and Letters,
the part of people of the lead district for a ^‘more careful and
full examination into the reasons which induced Prof. J. D.
Whitney, of the late Geological Commission to discourage the
hope of making deep mining successful.”® John Murrish was
commissioned to make the survey and it was hoped that his re¬
port would Tvarrant, at least in part, a reversal of the judgment
pronounced by Whitney. Murrish® reported that up to that
time mining had been confined mostly to that portion of the
strata above water where mining operations could be carried on
at a trilling expense; that this portion of the strata was almost
exhausted ; and that a new system of mining would have to be in¬
troduced in order to work the deeper mines. His conclusion,
that a new system of mining must be introduced, combining
capital and skill, was in accord with the conclusion reached by
Daniels fifteen years previous to that time. In the Annual
Eeport made to the Governor of the state in January, 1872,
on the industrial condition of the state, J. W. Hoyt stated that
the re-survey of the lead region by John Murrish had “done
something to quicken the interest in lead-mining, since the con¬
clusions reached by him are favorable to the opinion that large
deposits of mineral will be found in the lower Magnesian lime¬
stone. ” It is to be noted that the interest was still exclusively
in lead, and in a hope for the revival of lead production. This
hope was destined not to be realized. The zinc output of 1870
was about equal to that of lead. In the next year it exceeded
lead production, and the increased mineral production which oc¬
curred from 1871 to 1876 was due to zinc output and not to
lead. The latter remained stationary and gradually de¬
creased. However, the desired combination of capital and skill
had not yet been made. The increased production which oc¬
curred from 1871 to 1876, and probably for some years later,
was due to the fact that the water table had lowered itself.
This was noted by Moses Strong in 1876 when he made a report
on the Geology and Topography of the Lead Region
The large streams of the Lead Region contain a much smaller
amount of water than heretofore. Several places were seen where
8 Hoyt, J. W., Annual Report on Mining; Trans. Wis. State Ag. Soc.,
Vol. 9, 1870, p. 51.
® Murrish, John, Report on the Geological Survey. Appendix in Trans.
Wis. State Ag. Soc., Vol. 10. 1871.
10 Trans. Wis. State Ag. Soc.. Vol. 10. 1871.
Ch.amherlin. T. C., Geology' of W’isconsin, 1373-79, Vol. 2, p. 657.
Schuhring — Lead and Zinc Mining in Wisconsin. 15
old mills, formerly operated by water-power, had been abandoned, on
account of a diminished and final failure of the supply.
The diminution is not confined to surface water, spring, streams
and the like, but is true, to a greater or less extent, of all mining
ground of the region. In many instances this circumstance alone
has led to the reopening and profitable working of mines which years
ago were abandoned on account of water, with ore ‘going down’ in
the crevices.
Strong stated that the chief cause of this decrease in water
supply was the cultivation of the land, since a much greater
amount of surface was thus exposed, and evaporation took place
more rapidly and in larger quantities. Agriculture may thus
be said to have been influential in increasing the mineral out¬
put at this time. The influence of agriculture on mining will
be spoken of later.
Unfortunately Strong’s data stop with the year 1876, and
after that date zinc statistics become scattered and incomplete.
That the next ten years did not see the devolpment of deep min¬
ing is shown by the following statement made in 1886 by Thomas
H. McElroy, editor of the Southwestern Local, of Shullsburgh.
There has been quite a revival of mining in the region. It is pro¬
posed to sink a test shaft to determine the possibility of profitable
deep mining, the greatest depth reached thus far being about 160 feet.
The average depth of the mines being worked in the Wisconsin lead
and zinc region is not over 50 feet, and there are still 300 feet of the
upper lead-bearing Magnesian limestone unexplored.
McElroy called the district the “lead and zinc” region, thus
recognizing the importance of zinc.
The relation of agriculture to mining in the Wisconsin region
is interesting, for it in turn stimulated and retarded lead and
zinc production. In the early days when almost every farmer
was his own miner, an increase in the agricultural population
resulted in increased lead production. As agriculture developed
and mining became more difficult because the surface ores had
been removed, farmers turned their attentiori more and more to
the profits of agriculture, to the neglect of mining. Thus agri¬
culture was in part responsible for diminished lead output.
However, as a result of extensive cultivation of the land, the
water table was lowered and individual mining was again made
jmssible. Thus agriculture stimulated mining. Not only did-
agriculture affect mining, but mining affected agriculture as
well. In 1853 a writer, referring to the circumstances under
which Wisconsin began her civil career, says:^®
^2 Mineral Resources of the United States, Vol. 1886, p. 148.
Hunt’s Merchants’ Magazine, Vol. 28, p. 447.
16 Wisconsin Academy of Sciences^ ArtSy and Letters.
The mineral resources of Wisconsin attracted the attention of the
first settlers, and, although this fact retarded the progress of agri¬
culture, by drawing a large proportion of the earlier emigrants to the
more exciting life of the miner, yet it must at the same time, have
created a home market of some extent, and thus given an impulse to
agriculture. As in all new States, the want of a market most pro¬
longs the state of incipiency in agriculture, so this need was but
slightly alleviated by the market which the mining and lumbering
districts afforded.
From the standpoint of agriculture, the mines, by proving
more attractive to first settlers, retarded agriculture ; but at the
same time the miners furnished a market for agricultural prod¬
ucts, and this stimulated agriculture. The inter-relation be¬
tween mining and agriculture in Wisconsin was referred to by
Blake in 1893.^^ He said that the demand for lead was one of
the early and potent factors determining the settlement of
southwestern Wisconsin and the development of its mining and
agricultural resources, and made the following statement :
The mines of that section, together with those of Missouri and the
Mississippi Valley, may be said to have been the cradle of mining
in the Western United States. The deposits of ore being at or near
the surface and being numerous and widely distributed, afforded to
poor men an opportunity to mine on their own account with little or
no capital. The “diggings” which have often been termed “poor
men’s mines” soon attracted a large population. Laborers and min¬
ers were drawn thither from Cornwall and other mining centers, and
those not bred to the use of the pick, gad, and windlass soon gained
an experience in the use of tools and methods most useful to them in
the new fields of the great west and in the mountain ranges sloping
to the Pacific.
Mining thus stimulated agriculture, but it also made miners
out of agriculturists, some of whom later Avent farther west.
This suggests another factor affecting lead and zinc production :
the development of richer mines farther west which attracted
miners from Wisconsin just as the California gold fields at¬
tracted them in 1848.
The exact extent to which agriculture affected mineral pro¬
duction, either favorably or unfavorably, it is impossible to
state. The two were closely related ; and owners of the land are
still mostly farmers. However, they now lease the ore-bearing
areas to mining companies.^®
Blake, William P., The Mineral Deposits of Southwestern Wisconsin ;
Paper delivered before International Eng-ineering Congress in August, 1893 ;
p. 558.
Rickard, T. A., The Valuation of Metal Mines.” Paper No. 151, Ameri¬
can Institute of Mining Engineers at International Engineering Congress,
1915.
Schuhrmg — Lead and Zinc Mining in Wisconsin. 17
Nor is it possible to state exactly what effect the general pros¬
perity of the nation had upon lead and zinc mining in Wiscon¬
sin. Periods of financial depression must necessarily have their
effect upon mining, as well as other industries, and yet the
statistics of production do not always reflect that depression.
For example, the year 1893 was one of exceptionally low prices
for lead, to which the uncertainties of the tariff and the financial
situation largely contributed. In this year Kirchoff reported
that the lead mining industry in the United States shared in the
general depression of business.^® He stated, further, that the
statistics for the second half of 1893 strikingly illustrated the
stress put upon the zinc industry through the financial panic,
the older Illinois and Eastern producers having borne the strain
most successfully. No separate statistics are available for lead
production in Wisconsin for the years immediately following
1893, but the incomplete figures for zinc production show only
a slight variation in output. Again, in 1907,^^ while the out¬
put for the latter half of the year suffered severely, still the
impetus gained from the previous prosperous year carried the
industry along until the middle of 1907, and the total output
of the whole region during that year and the year following
showed an increase in both lead and zinc production. Sieben-
thal reported ‘‘much activity both in mining and in prospecting
in the Upper Mississippi Valley’’ in the year 1907.^® Nor was
the Wisconsin production unfavorably affected, either in lead
or zinc. In Missouri the zinc output declined somewhat but
lead was not affected. The newer regions of production, on the
other hand, Idaho, Utah, and Colorado, shoAved a definite re¬
sponse to the panic of 1907.
The price of metals has always been a potent factor in pro¬
duction. Whitney in 1854 quoted prices of lead in order to
throw some light on fluctuations in production. The demand
for lead, as evidenced by prices, was usually followed by in¬
creased production; and it was the high price of zinc in 1900
that led finally to the introduction of organized capital into the
district ; and the consequent development of the region has made
Wisconsin one of the important zinc-producing States.
Kirchoff, Charles; Mineral Resources of the United States, 1894, Pt. HI
of 16th Annual Report of the U. S. Geol. Survey, p. 359.
Kirchoff, C., The Mineral Industry, 1893, p. 104.
Siebenthal, C. E., Miner-al Resources of the United States, 1907, Pt. 1,
p. 659.
9
18 Wisconsin Academy of Sciences, Arts, and Letters.
In 1899 the American Smelting and Kefining Company was
formed^® and the first modern mill was put in active operation in
the southwestern zinc field.^® By 1905 there were from twenty
to thirty large dividend-paying mines and many smaller ones,
as against only two or three a few years before that time. The
chief causes for improvement were the high prices for ore and
the perfecting of the system of magnetic separation.^^ This
enabled the separation of marcasite from the blende. Owing to
the nearness in specific gravity of these two ores, it had been
difficult to separate them cleanly by hydraulic methods.
In 1907, the panic year, about fifty new concentrating plants
were erected, many of them of large size.^^ In 1908 occurred
the consolidation of light and power companies^^ and during
the following year a large central power station was constructed
to supply light and power all over the district.^'^ The year 1909
was also signalized, aside from the development of individual
companies, by the initiation of a policy of expansion by three
of the larger mining companies, and the feature of the year
1910 was the extension of holdings by the leading operating
companies in the district.^® In 1911 the large companies con¬
tinued to add to their holdings by purchase and lease, and most
of the increased output was due to their operations.^® The con¬
centration of the mining properties of the district into the hands
of fewer interests continued during 1912,-^ and in 1913 the
large mining interests continued to add to their holdings by
lease or purchase, and increased the scope of their operations by
acquiring ncAv fees or leases.^* In 1915 several new mines were
operated Avith local capital, and outside capital was invested in
the district, but the increased production and development Avas
mainly due to the extension of operations by the large com¬
panies.^® The interest now is entirely in the mining of zinc.
Mineral Resources of the United States, 1899-1900, Pt. VI, 21st Annual
Report of the U. S. Geol. Survey, p. 229.
Moore, E. W., The Mineral Industry, Vol. 15, p. 765.
21 Ibid., Vol. 14, pp. 574-6.
22 Dunlop, J. P., Mineral Resources of the United States, 1910, Pt. 1. p.
670.
28 Siebenthal, C. E., Mineral Resources of the United States, 1908, Pt. 1,
p. 640.
»*Siebenthal. C. E., Ibid. 1909. Pt. 1, p. 531.
® Siebenthal. C. E., Ibid., 1910, Pt. 1, p. 27.3.
2« Dunlop. J. P., Ibid., 1911, Pt. 1, p. 866.
21 Siebenthal, C. E., Ibid., 1912. Pt. 1. p. 387.
2» Dunlop, J. P. Ibid., 1918, Pt. 1, p. 162.
29 Dunlop J. P., Ibid., 1915, Pt. 1, p. 130.
S chubring — Lead and Ziric Mining in W iscomin. 19
The relative values of the two industries are shown in tables
and charts which follow.
Siebenthal, in speaking of the Upper Mississippi Valley re¬
gion makes the following statement
The mines were at first worked exclusively for lead, but since
1860 the production of zinc ore has predominated over that of lead
ore, with which the zinc ore is associated in many mines.
This statement is not accurate, for while the zinc ores first be¬
gan to be utilized in 1860, the output was very small, and it was
not until some years later that the tonnage of zinc exceeded that
of lead. For Wisconsin this occurred in the year 1871, when
the production of zinc ores amounted to 12,961 tons and of lead
ore, 6,742 tons. In 1860 the output of zinc ore from the region
was 160 tons, whereas the output of metallic lead was 12,227
tons.
The statistics of zinc production are so fragmentary that no
fair comparison can be made between the two industries except
for the ten-year period 1866-1875, when Moses Strong collected
data which he considered to be fairly complete, and the ten-
year period 1907-1916, for which annual statistics appear in
Mineral Resources of the United States. These two ten-year
periods are discussed in the pages which follow. Discussion of
lead production covers the same two periods. Moses Strong
collected data for the earlier decade which make possible the
representation of lead production in Wisconsin by producing
districts ; and the United States Geological Survey has published
similar data beginning with the year 1907. Exhibits I and U
show annual production of lead and zinc, respectively, for the
Upper Mississippi Valley, whereas the charts and accompanying
text which follow deal with the production of Wisconsin only.
8® Siebenthal, C. E., Mineral Resources of the United States, 1910, Pt. 1,
p. 235.
20
Wisconsin Academy of Sciences, Arts, and Letters,
Fig. 2. Mining Districts and Furnaces in the Wisconsin Lead Region in 1876
From Chamberlin's Geology of Wisconsin 1873-79
ScJmbring — Lead and Zinc Mining in Wisconsin.
21
TV.
Wisconsin Lead Production.
Mining Districts and Furnaces in Wisconsin in 1876. In
1876 Grant, Iowa, and Lafayette counties each had five fur¬
naces. (Pig. 2). Two districts in each county had two fur¬
naces each, i. e. : Dodgeville and Mineral Point in Iowa County ;
Potosi and Platteville in Grant County; and Shullsburg and
Hazel Green in Lafayette County. The Hazel Green district
included portions of Grant and of Lafayette counties, but the
furnaces were located in Lafayette County. The village of
Hazel Green, on the other hand, is across the line in Grant
County. The following districts had one furnace each. High¬
land in Iowa County ; Beetown in Grant County, and New Dig¬
gings in Lafayette County. The number of districts having
furnaces was nine: Mineral Point, Dodgeville, Highland, Bee-
town, Potosi, Platteville, Hazel Green, New Diggings, and
Shullsburg.
Production. The amount of lead ore smelted by furnaces
in these nine districts is shown in diagramatic form in figures
3a, 3b, and 3c, which represent respectively production of the
five-year period 1866-1870; production of the five-year period
1871-1875, and production of the ten-year period 1866-1875.
In the first period mentioned, the five years from 1866 to 1870,
Potosi had the largest production, 7,300 tons, while New Dig¬
gings and Mineral Point came second, with 5,900 and 5,588 tons
respectively. Shullsburg, Dodgeville, and Hazel Green came
next in the order of production, while Platteville, Beetown, and
Highland produced the smallest amounts. Estimated produc¬
tion for Highland was 1,250 tons.
In the five-year period from 1871 to 1875, inclusive, the pro¬
duction for each district decreased with the exception of Dodge¬
ville and Beetown. These districts had an increase of about
35% each, the production of Dodgeville increasing from 2,965
tons in the earlier period to 4,023 tons in the period 1871-1875 ;
and the production of Beetown increasing from 1,800 tons to
2,425 tons. Potosi, which formerly occupied first place, pro¬
duced at this time less than Mineral Point, Dodgeville, Shulls¬
burg, or New Diggings. In the five year period under discus¬
sion Mineral Point had the largest production, 4,510 tons.
22 Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 3a. Production of Lead Ore 1866-1870, Inclusive
Data from Chamberlin's Geology of Wisconsin 1873-79
Schuhring — Lead and Zinc Mining in W isconsin.
23
Fig. 3b. Production of Lead Ore 1871-1875, Inclusive
Data from Chamberlin* s Geology of Wisconsin 1878-79
24
Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 3c. Production of Lead Ore 1866-1875, Inclusive
Data from Chamberlin* s Geology of Wisconsin 1873-79
Schuhring — Lead and Zinc Mining in Wisconsin, 25
Dodgeville and Shullsburg each produced about 4,000 tons, and
New Diggings, Potosi and Beetown produced lesser amounts.
Hazel Green, Platteville and Highland produced the least. The
production of Highland is 1,250 tons as in the former five-year
period, the reason being that the production of Highland was
an estimate of 500,000 pounds per annum. The total produc¬
tion in the state (see table 2) shows a decrease of 21%, from
34,450 tons in 1866-1870 to 26,216 tons in 1871-1875.
Figure 3c shows production of lead ore by furnace districts
for the ten-year period 1866-1875, inclusive. If Hazel Green
is assigned to Lafayette County because her furnaces were lo¬
cated there, each of the three counties comprising the lead re¬
gion in Wisconsin smelted about the same quantity of ore, in
the neighborhood of 20,000 tons. Lafayette smelted 23,463 tons ;
Iowa 19,586 ; and Grant 18,617. The highest individual output
was by the Potosi district, 10,590 tons. The lowest, 2,500 tons,
was by Highland. This amount, as above stated, was an esti¬
mate. Not far behind Potosi were Mineral Point, 10,098 tons;
New Diggings, 9,348 tons, and Shullsburg, 8,448 tons. Next in
order of rank came Dodgeville, 6,988 tons, and Hazel Green
5,667 tons, followed by Beetown and Platteville with 4,225 tons
and 3,802 tons, respectively.
Figures 4a, 4b and 4c show production of lead ore for the
following periods: the five years 1907-1911, inclusive; the five
years 1912-1916, inclusive, and the ten years 1907-1916, inclu¬
sive. These charts have the same scale as the series showing
production of lead ore from 1866 to 1875, and may be directly
compared as to quantity of production. In making compari¬
sons it should be borne in mind that the earlier series show the
amount of lead ore smelted by the fifteen furnaces then located
in nine of the producing districts, whereas the later charts show
the output of each of the producing districts. In 1876 the New
Diggings District included the mines in the vicinity of Benton,
and the furnaces of the Mineral Point district smelted the ores
produced by the Mifflin and Linden districts. The output of
Mifflin and Linden should therefore be added to the amount pro¬
duced by Mineral Point when comparison is made with the fur¬
nace production of Mineral Point.
A study of the two sets of charts reveals, first of all, a notable
Chamberlin’s Geology of Wisconsin, 1873-78, v. 2, p. 746.
Fig. 4a. Production of ead Ore 1907-1911, Inclusive
Data from Mineral Resources of the United States
Schubring — Lead and Zinc Mining in Wisconsin.
27
IG. 4b. Production of Lead Ore 1912-1916, Inclusive
Data from Mineral Resources of the United States
28 Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 4c. Production of Lead Ore 1907-1916, Inclusive
Data from Mineral Resources of the United States
Schuhring — Lead and Zinc Mining in Wisconsin.
29
decrease of production in the district as a whole. The total
amount of lead ore produced in the ten years 1866-1875 was
61,666 tons, (see table 2) whereas the total amount produced
in the ten years 1907-1916 was 39,575 tons, (see table 3).
This is a decrease of 35.8%. Compared with the decade of
maximum production 1841-1850, inclusive, (see fig. 1) the
production of 1907-1916 shows a 79% decrease. Notable also
is the change of production in Grant County. The Beetown
district, which formerly ranked with the Platteville district,
shows no production whatever in the ten years 1907-1916.
Potosi, formerly the greatest producing district, (figs. 3a, 3b, 3c),
is now one of the least, (figs. 4a, 4b, 4c). Her 1916 produc¬
tion was included with the production of Platteville. Platte¬
ville shows a decrease in production of over 50%, (from 3,802
tons for the ten years 1866-1875 to 1,794 tons for the ten years
1907-1916). It is to be remembered that the two sets of charts
are not exactly comparable. It is probable that the production
of Cuba City-Blmo includes ore which formerly was sent to
Platteville to be smelted. The Cuba City-Elmo district is lo¬
cated in Grant County between Platteville and Hazel Green.
In 1876 the Big Patch Diggings were located northwest of the
present village of Elmo, and the production of Big Patch Dig¬
gings was sent to the Platteville furnaces.^^ The production of
this new district Cuba City-Elmo was the highest in the state
for the five years 1907-1911, 6,152 tons, but in the next five
years, 1912-1916, Elmo produced nothing, and the production
of Cuba City fell to less than one thousand tons.
In Iowa county there seems at first glance to have been a
shifting of production from the eastern to the western part of
the county, and Mineral Point and Dodgeville which formerly
were large producers seem now to be producers of small import¬
ance. It should be remembered that the production of Linden
and Mifflin was included in the Mineral Point production of
1866-1875. The output of Highland (for the years 1907-1916)
is slightly greater than the estimate made for Highland in 1866-
1875.
In Lafayette County in the ten years 1866-1875 Shullsburg
and New Diggings produced almost equal amounts of ore, with
New Diggings slightly in the lead. The New Diggings district
32 Chamberlin’s Geology of Wisconsin, 1873-78, v. 2, p. 744.
30 Wisconsin Academy of Sciences, Arts, and Letters.
included mines in the vicinity of Benton.^® Of late years New
Diggings has decreased in importance. Its 1916 output was
included with the output of the Benton district which has come
to be the largest producer in the state. Shullsburg whose pro¬
duction slightly exceeded that of New Diggings-Benton in the
five years 1871-1875, shows a steady decline. Shullsburg fur¬
naces smelted 8,448 tons of lead ore in the ten years 1866-1875,
and the district produced 2,002 tons of lead ore in the ten years
1907-1916. With the decline of the Cuba City-Elmo produc¬
tion, which in 1907-1911 exceeded the Benton-production, the
Benton district assumes first rank.
The Benton-New Diggings district produced 11,789 tons of
lead ore in the years 1907-1916. This is a larger amount than
was smelted by Potosi in the ten years 1866-1875, and Potosi
was at that time the largest producer, her output for the ten
years having been 10,590 tons.
The trend of production in recent years is shown in figures
7a and 7b. The whole region produced 24,280 tons of lead ore
from 1907 to 1911, inclusive, and 15,295 tons from 1912 to 1916,
inclusive, (see table 3). This was a 37% decrease in produc¬
tion. The only two districts which showed an increase of pro¬
duction in 1912-1916 over the five-year period immediately pre¬
ceding were Linden and Benton. The output of Linden in¬
creased 32%, from 1,635 tons to 2,158 tons, and the output of
Benton 21%, from 5,337 tons to 6,452 tons.
Quantity of metallic lead by producing districts is charted in
figures 5a, 5b and 5c, showing production for the five years
1907-1911, inclusive; for the five years 1912-1916, inclu¬
sive ; and for the ten years 1907-1916, inclusive. These figures
are drawn to the same scale as the figures already discussed and
are directly comparable with them. Data were not obtainable
for production of metallic lead for the years 1866-1875.
The relative ranks of the lead producing districts in Wiscon¬
sin are indicated in figures 6a, 6b, 6c, and 7a, 7b, 7c, which in¬
dicate by bar graphs the following: Quantity of lead smelted
by furnace districts in the five years 1866-1870, the five years
1871-1875, and the ten years 1866-1875; quantity of lead ore
produced in the five years 1907-1911, the five years 1912-1916,
and the ten years 1907-1916. The quantity of metallic lead ob¬
tained from the ores is indicated in the last three periods nien-
Chamberlin’s Geology of Wisconsin, 1873-78, v. 2, p. 746.
Schuhring — Lead and Zinc Mining in Wisconsin.
31
Fig. 5a. Production of Metallic Lead 1907-1911, Inclusive
Data from Mineral Resources of the United States
32
Wisconsin Academy of ScienceSf Arts, and Letters.
Fig. 5b. Production of Metallic Lead 1912-1916, Inclusive
Data from Mineral Resources of the United States
Schuhring — Lead and Zinc Mining in Wisconsin. 33
3
Fig. 5c. Production of Metallic Lead 1907-1916, Inclusive
Data from Mineral Resources of fKe United States
state Total 34,450
Fig. 6a. Quantity of lead ore produced in Wisconsin 1866-1870, inclusive, by furnace districts.
Data from Chamberlin’s Geology of Wisconsin, 1873-1879.
Thousands of short tons
Potosi
Mineral Pohit
New Diggings
Shullsburg
Dodgeville
Hazel Green
Beetown
Platteville
Highland
10
I
11 Short Tons
10,590
10,098
9,348
8,448
6,988
5,667
4,225
3,802
2,500
State Total 61,666
Fig. 6c. Quantity of lead ore produced in Wisconsin 1866—1875,
inclusive, by furnace districts.
Data from Chamberlin’s Geology of Wisconsin, 1873-1879
Thousands of short tons
36 Wisconsin Academy of Sciences, Arts, and Letters.
Potosi I ^ m 79.6% 78 98
Ore Metal -
State Total 78.6% 19,097 24,280
Fig. 7a. Quantity of lead produced in Wisconsin 1907-1911, inclusive.
Data from Mineral Resources of the United States.
Thousands of short tons
Schuhring — Lead and Zinc Mining in Wisconsin.
37
os
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”8 I
bo
^w’
<» o
d 15
g «
0“
r^fC
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38
Wisconsin Academy of Sciences, Arts, and Letters.
(M _
C£)
rH
I
t-
o
Oi
C •!
I
I S
P dj
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•TZS s
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uj
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Sclmbring — Lead and Zinc Mimng in Wisconsin, 39
tioned and something of the tenor of the ore is thus shown.
Percentages are recorded on each chart. For the earlier period,
1866-1875, data for metallic lead were not available.
Consideration of the five-year periods in succession, 1866-
1870, 1871-1875, 1907-1911, and 1912-1916, shows that Potosi
moved successively from first place to fifth, to thirteenth, and
finally to insignificance. New Diggings (with the Benton pro¬
duction included) moved from second place to fourth; back to
second under the name, now, of Benton, and finally to first place
(with 1916 New Diggings included in Benton production).
Shullsburg occupied successively fourth, third, sixth, and fourth
place. Mineral Point moved from third place in 1866-1870 to
first in 1871-1875 and thereafter it declined. Her production
when third in rank was 5,588 tons, a larger amount by 1,000 tons
than she produced when occupying first rank. The Linden and
Mifflin ores were smelted by the Mineral Point furnaces in 1866-
1875, as already indicated. The production of the whole lead
region decreased successively as follows in the five-year periods
under discussion : 34,450 tons, 27,216 tons, 24,280 tons, 15,295
tons.
The quality of the ore is indicated by the percentages which
appear on the charts. This information was obtainable for the
years 1907-1916 but not for the earlier period 1866-1875. For
the state as a whole the quality of the ore seems to have declined,
for in the five years 1907-1911 24,280 tons of lead ore pro¬
duced 19,097 tons of metallic lead, showing that the metal con¬
tent of the ore was 78.6%, whereas in the five years 1912-1916
11,306 tons of metal were obtained from 15,297 tons of ore, indi¬
cating a 73.9% metal content. The percentages range from
80.2% for the Cuba City-Elmo district when it ranked as first
producer in the years 1907-1911, to 68.3% for the Hazel Green
district in the years 1912-1916. The quality of the ore does not
remain constant in a given district, as is shown by the fact that
Cuba City in 1912-1916 produced only a 76.4% grade. This is
rather low as compared with its 80.2% former grade, but it is a
high percentage when compared with the output of the other
districts during the same five-year period. In that period,
1912-1916, only one district produced ore of a better quality,
namely Shullsburg, which showed a percentage of 78.6. The re¬
maining districts range from 76.1% to 68.3%. The lowest figure
belonged to Hazel Green, but Hazel Green in the five years im-
Thousands of short tons
40 Wisconsin Academy of Sciences, Arts, and Letters,
I
CD
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I
I
I
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O •'I*
Sll
CD Oi 00 05 CO
03 CD CD CO CO
lO tjT iO CO
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03 00 lO O-
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Schuhring — Lead and Zinc Mining in Wisconsin. 41
mediately preceding produced a 76.4% ore. Every district
with the exception of Shullsburg showed a decrease in the qual¬
ity of ore. Shullsburg shows a slight increase, from 78.1% in
1907-1911 to 78.6% in the five years 1912-1916.
The production thus far considered has been for individual
districts in the lead region of Wisconsin. Total state produc¬
tion is shown in fig. 8. This chart gives the quantity of lead
produced annually from 1866 to 1875, inclusive, and annually
from 1907 to 1916, inclusive. For the ten years 1866-1875, only
ore statistics were available, but the ten years 1907-1916 show
not only the quantity of ore produced each year, but the metal
obtained from the ore. The percentages indicate the quality of
the ore.
In the ten years 1866-1875 the production remained fairly
constant for the first six years. Then it decreased for two years,
after which it again remained about the same. From 1866, in
which year production amounted to 7,015 tons of lead ore, to
1875, when the output amounted to 4,740 tons, there was a de¬
crease in annual production of about 33%.
In the ten years 1907-1916 there was greater fluctuation than
in the earlier period considered. The highest point in the pro¬
duction of the last ten years was reached in 1910, when 5,608
tons of ore were produced. After 1910 there was a continuous
decline until 1914 when the output of lead ore amounted to 2,028
tons. In 1915 and 1916 there was a decided percentage increase
— an easy accomplishment when the tonnage is so small. The
1916 output was again a little over 4,000 tons, as in the years
3907, 1909, and 1911. 1908 and 1910 were the only years when
production surpassed the 5,000 ton mark.
State production of lead ore for the ten years 1866-1875 was
61,666 tons, and for the ten years 1907-1916 the output was
39,575 tons. This is a decrease of 35.8%.
The quality of the ore declined steadily. In 1907 79.4% of
the lead ore was metallic lead, while in 1916 only 72.7% of the
ore was metal. For the whole period, from 1907 to 1916, in¬
clusive, 30,403 tons of metal were obtained from 39,575 tons of
concentrates, or 77%.
The value of the lead produced in the several districts in Wis¬
consin (a) in the five ^mars 1907-1911, inclusive; (b) in the five
years 1912-1916, inclusive; and (c) in the ten years 1907-1916,
inclusive, is shown in figs. 9a, 9b, 9c, and figs. 10a, 16b, 10c.
42 ^yiscons^n Academy of Sciences, Arts, and Letters.
Fig. 9a. Value of Metallic Lead Produced 1907-1911, Inclusive
Data from Mineral Resources of the United States
Sclmhring — Lead aiid Zinc Mining in Wisconsin. 43
Fig, 9b. Value of Metallic Lead Produced 1912-1916, Inclusive
Data from Mineral Resources of the United States
44
Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 9c. Value of Metallic Lead Produced 1907-1916, Inclusive
Data from Mineral Resources of the United States
Thousands of dollars
Fig. 10a. Value of lead produced in Wisconsin 1907-1911, inclusive.
Value of metal is calculated from average daily quotations at New York
and value of ore is that received by the producer. Data from
Mineral Resources of the United States.
Thousands of dollars
46
Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 10b. Value of lead produced in Wisconsin 1912-1916, inclusive.
Value of metal is calculated from average daily quotations at New York
and value of ore is that received by the producer. Data from
Mineral Resources of the United States.
Tliousands of dollars
0 100 200 300 400 500 600 700 800
Sclmhring — Lead and Zinc Mining in Wisconsin. 47
o
u
1
o
05
w
a>
«
o
;-i
A
CS
<v
O
a>
>
a a>
<V 03
*08
48 Wisconsin Academy of Sciences^ Arts, and Letters.
In the first of these sets of charts bars representing value in
dollars extend eastward and westward from north and south
lines which run through the central part of the mining district.
This method was adopted in order to make comparison of values
easier. The bars extend to, and beyond, the respective cities
whose production is represented. In some cases, however, the
bar falls short of reaching the city, either because production
was small or because the city is located unusually far east or
west of the central meridional lines chosen. In these cases
arrows indicate the cities to which the production is to be cred¬
ited.
In the second of these sets of charts (figs. 10a, 10b, and 10c)
the value of the metal and the value of the ore is shown in the
same figure, and the respective amounts indicated. In making
a comparison of these values, it should be borne in mind that the
value of the metal is calculated from average daily quotations
at New York, whereas the value of the ores is that received by
the producer.
In the two five-year periods under discussion (1907-1911, and
1912-1916) the Benton district and the Cuba City-Elmo dis¬
trict received the largest sums of money for lead production.
In the period 1907-1911, inclusive, the amount received by the
Cuba City-Elmo district was $343,038 for galena and $436,918
for metallic lead. Benton, the next largest producer, received
in the same five-year period $287,190 for galena and $383,404
for metallic lead. In the five years 1912-1916, inclusive, Ben¬
ton took first place with $369,319 for ore and $499,340 for metal.
The ten-year period, 1907-1916, shows Benton first with $656,509
for ore and $882,744 for metal; and Cuba City-Elmo second
with $393,423 for ore and $501,922 for metal.
The amount received in the state in each of the five-year
periods was in the neighborhood of one million dollars for lead
concentrates and somewhat more than a million dollars for me¬
tallic lead. In the ten years 1907-1916, the amount received by
Wisconsin producers of lead concentrates was $2,267,819; and
the value of the metallic lead, as calculated from average daily
quotations at New York, was $2,877,516.
The value of lead produced in Wisconsin annually 1907-1916,
inclusive, is shown in fig. 11. Amounts received for galena
varied from one hundred thousand dollars to about three hun-
Thousands of dollars
0 100 200 300 400 Ore Metal
Schuhring — Lead and Zinc Mining in Wisconsin.
o
0)
ei
C/2
<v
o
0
TU
o
pH
c3
O)
O
<D
P
cd
49
4
Value of metal is calculated from average daily quotations at New York
and value of ore is that received by the producer. Data from
Mineral Resources of the United States.
50 Wisconsin Academy of Sciences^ Arts, and Letters.
dred thousand dollars. The smallest amount, $85,196, was re¬
ceived in 1914, and the largest, $291,397, was received in 1916.
This is not much more than was received ten years before. In
1907 the amount received was $266,471. The value of metallic
lead was $116,532 in 1914, $419,934 in 1916, and $370,894 in
1907.
V.
Wisconsin Zinc Production.
The quantity of metallic zinc produced by the several dis¬
tricts in Wisconsin (a) in the five years 1907-1911, inclusive;
(b) in the five years 1912-1916, inclusive; and (c) in the ten
years 1907-1916, inclusive, is shown in figures 12a, 12b and 12c.
The zinc production so far exceeded the lead production in Wis¬
consin during the years under consideration that a much smaller
scale had to be used. The two sets of charts are therefore not
directly comparable. The zinc charts are comparable with each
other for they are drawn to the same scale for ores and metal,
for five-year periods and ten-year periods.
Table 5 shows that small amounts of zinc carbonate were pro¬
duced by each of the zinc-producing regions with the exception
of Hazel Green which did not report any production of zinc car¬
bonate in the ten years 1907-1916, inclusive. Most of the
amounts were less than one hundred tons for the ten years and
they do not appear on the charts.
Highland was the greatest producer of zinc carbonate in the
state, far out-ranking Mineral Point and Benton, the only other
two regions whose carbonate production is represented on the
charts. In the five years 1907-1911, inclusive, Highland pro¬
duced 17,649 short tons of zinc carbonate, and in the five years
1912-1916, inclusive, 13,618 short tons, making a total of 31,267
short tons for the ten years. Mineral Point produced 4,194
short tons in the ten years and Benton 2,650 short tons, the two
together producing somewhat more than a fifth as much as High¬
land. The zinc production of Highland is unique in this re¬
spect ; that its production of zinc carbonate exceeds its produc¬
tion of sphalerite, or zinc sulphide. The output of sphalerite
from Highland for the ten years 1907-1916, inclusive, was 8,510
short tons, or about one-fourth the carbonate output. Mineral
Schuhring — Lead and Zinc Mining in Wisconsin,
51
Fig. 12a. Production of Metallic Zinc 1907-1911, Inclusive
Data from Mineral Resources of the United States
52 Wisconsin Academy of ScienceSy Arts, and Letters,
Fig. 12b. Production of Metallic Zinc 1912-1916, Inclusive
Data from Mineral Resources of the United States
Schuhring — Lead and Zinc Mining in Wisconsin,
53
Fig. 12c. Production of Metallic Zinc 1907-1916, Inclusive
Data from Mineral Resources of the United States
54 Wisconsin Academy of Sciences, Arts, and Letters.
Point, on the other hand, produced more than twice as much
sphalerite as carbonate, and Benton, in the same ten years pro¬
duced 368,713 tons of sphalerite and only 2,650 tons of carbon¬
ate. The production of carbonate in the Benton district is rela¬
tively so small as to be negligible.
In the five years 1907-1911, inclusive, Benton stands out as
the principal producer of sphalerite, with Platteville and Hazel
Green not far behind. The amounts produced were respectively
87,095 short tons, 62,346 short tons, and 56,458 short tons.
Dodgeville produced the smallest amount, 2,606 tons, while
Shullsburg and Montfort produced amounts almost as small.
Total state production in the five years was 344,022 short tons
of sphalerite and 21,705 short tons of carbonate, (see table 5).
The five years 1912-1916, inclusive, showed a decrease in zinc
carbonate production in the state, but a notable increase in the
production of sphalerite. Total production of carbonate was
17,120 short tons, and of sphalerite 646,908 short tons, (see
table 5).
Benton had a tremendously increased production for the five
years 1912-1916, inclusive; from 87,095 tons in 1907-1911, to
281,618 tons in 1912-1916. This is an increase of more than
230%. Both Hazel Green and Livingston forged ahead of
Platteville, leaving Platteville fourth in the order of production.
The output of Platteville decreased to 52,422 tons in 1912-1916,
from 62,346 tons in the five years immediately preceding. Be¬
sides Platteville, the districts which showed decreased produc¬
tion were Cuba City-Elmo, Highland, Mineral Point, and Mont¬
fort. The production of Dodgeville remained practically sta¬
tionary, having been 2,817 short tons in the five years 1912-
1916, inclusive, and 2,606 short tons in the five years imme¬
diately preceding. Shullsburg showed an increased production
of more than 100%, but the tonnage involved, 2,840 tons in the
five years 1907-1911, and 6,340 tons in 1912-1916, is negligible.
Mifflin and Linden each showed an increased tonnage of about
20,000, representing almost a doubled output. Livingston had
an increased tonnage of over 40,000 ; Hazel Green of over 50,000,
and Benton of almost 200,000. The combined increased pro¬
duction of the three last named almost make up the total in¬
crease of production in the state, as is shown in the following
table :
Schuhring — Lead and Zinc Mining in W isconsin.
55
1907-1911 1912-1916 Increase in 1912-1916
Short tons Short tons Short tons
Sphalertie Sphalerite Sphalerite
State of Wisconsin . . . 344,022 646,908 . 302,886
Benton . . 87,095 281,618 194,523
Hazel Green . 56,468 110,921 64,463
Livingston . 35,896 77,266 41,360
Combined increased tonnage of Benton,
Hazel Green, and Livingston _ _ _ _ 290 ,346
For the ten years 1907”1916, inclusive, Benton produced
368,713 short tons of sphalerite, more than twice as much as
the next greatest producer, Hazel Green, which produced
167,379 short tons. Platteville and Livingston produced al¬
most identical amounts, the former 114,768 tons and the latter
113,152 tons. Linden and Mifflin each produced in the neighbor¬
hood of 70,000 short tons, and the Cuba City-Elmo district pro¬
duced 45,126 tons. The smallest producers of sphalerite were
Mineral Point, Shullsburg, Highland, Montfort, and Dodgeville,
which ranked in the order named, and produced from 10,000 to
about 5,500 tons respectively. The carbonate production of
Benton, Mineral Point, and Highland has been discussed. The
combined production of carbonate and sphalerite for Highland
places that district in the rank occupied by the Cuba City-Blmo
district, rather than in the rank of smallest producers.
The relative ranks of the zinc-producing districts in Wiscon¬
sin are shown in figs. 13a, 13b, 13c, which represent by bar
graphs the following : quantity of zinc ores produced by each of
the several districts (a) in the five years 1907-1911, inclusive,
(b) the five years 1912-1916, and (c) the ten years 1907-1916,
inclusive. The quantity of metallic zinc obtained from the
ores is indicated, and the quality of ores thus shown. Percen¬
tages are recorded on each chart.
Benton, Hazel Green, Platteville, and Livin^ton were the
greatest producers in the years under consideration, with Ben¬
ton always in the lead. The Cuba City-Elmo district ranked
fifth in the five years 1907-1911, but production fell off consid¬
erably in next five-year period so that for the ten years 1907-
1916 this district ranked seventh in the order of production.
In the five years 1907-1911 Linden occupied sixth place and
Mifflin eighth, in the production of zinc ores. Highland occupy¬
ing seventh. However, Mifflin obtained a greater amount of
metallic zinc from its ores than Highland and Linden, and was
entitled to sixth place in the years 1907-1911. In the next
Thousands of short tons
56 Wisconsin Academy of Sciences, Arts, and Letters.
<0
as
'C
Fig. 13a. Quantity of zinc produced in Wisconsin 1907-1911, inclusive.
Data from Mineral Resources of the United States.
Thousands of short tons
Schuhring — Lead and Zinc Mining in Wisconsin. 57
o
0)
Fig. 13b. Quantity of zinc produced in Wisconsin 1912-1916, inclusive.
Data from Mineral Resources of the United States.
Thousands of short tons
58 Wisconsin Academy of Sciences^ Arts, and Letters.
* rH
cc
o
.2
c5
CO
tH
O
♦Includes 1916 New Diggings.
"♦Includes 1916 Potosi.
Schubring — Lead and Zinc Mining in Wisconsin. 59
five years Highland fell to eighth place, which is also the posi¬
tion of Highland in the ten year period 1907-1916. Conse¬
quently Linden and MijBlin occupied respectively fifth and sixth
places in the production of zinc ores, with Mifflin in the lead in
the production of metallic zinc.
The quality of the ores is indicated by the percentages which
appear on the charts. For the state as a whole the quality of
the ores seems to have declined somewhat, for in the five years
1907-1911 365,727 tons of zinc ores produced 115,495 tons of
metallic zinc, showing that the metal content of the ores was
31.6% ; whereas in five years 1912-1916 664,028 tons of ores
produced 192,479 tons of metal, indicating a metal content of
29%. The percentages range from 39.8, tenor of ores from
Dodgeville in the five years 1907-1911, to 21.8, percentage of
metallic zinc obtained from ores produced by Highland in the
five years 1912-1916, inclusive. The reason for the low percent¬
age of metallic zinc from Highland is that the greater propor¬
tion of ores from Highland is zinc carbonate which does not yield
so high a percentage of metal as does the sulphide. In the five
years 1907-1911, the percentage for Highland was 31.4, and
in that period a smaller amount of zinc carbonate was produced
relatively to the amount of sphalerite than in the five years im¬
mediately following. Leaving Highland out of consideration,
and comparing the percentages of districts producing mainly
sphalerite, Cuba City produced the lowest grade of ore in the
five years 1912-1916, with a percentage of 22.8. This is a great
reduction in percentage from the previous five years when the
metal content of ores produced by the Cuba City-Elmo district
was 37.8. In the same period other producers of high grade
ores were Mifflin with a percentage of 37.3, and Dodgeville,
39.8%, which is the highest percentage indicated on the charts.
The tonnage put out by Dodgeville, however, is very small. In
the five years 1912-1916, highest grades of ores were produced
by the following: Dodgeville, 38.8%, Shullsburg, 37.9%, Platte-
ville, 36.6%. For Dodgeville the percentage is a decrease, in¬
dicating that its ores were of lower grade than in the five years
immediately preceding. For Shullsburg and Platteville the
percentages are increases. The tonnage for Shullsburg is not
large, hence the increase in percentage is not so important as in
the ease of Platteville. In the five years 1907-1911 Platteville
Thousands of short tons
60
Wisconsin Academy of Sciences, Arts, and Letters.
xn b-WOONfHlOrJtCvJOO
n 00 tH 05 O N I> tH rH o
O •• W N
H 2 oDcccoioiiOoai-rt^i^eo
r-l Ol tH iH tH
go
ui
o _
«Dt>0005OrHC<IC0T»«i0
COCDCO«Ot>I>l>t-t-l>
00 00 00 00 00 00 00 00 00 00
THiHr-lTHTHr-i'r-liHTHT— (
Fig. 14. Quantity of zinc produced annually in Wisconsin 1866-1775, inclusive, and annually 1907-1916, inclusive.
Data from Chamberlin’s Geology of Wisconsin, 1873-1879, and Mineral Resources of the United States.
S chubring — Lead and Zinc Mining in Wisconsin.
61
produced 62,345 tons of ores, from which 20,517 tons of metal¬
lic zinc were obtained, indicating a metal content of 32.9%.
In the next five years Platteville produced 52,477 tons of ores,
from which 19,146 tons of metal were obtained, showing a per¬
centage of 36.6. The quality of ores had improved to such an
extent that practically the same amount of metal was obtained
from much less ore, the difference in ore tonnage being about
10,000 tons. Only Shullsburg, Platteville, and Benton, showed
a better quality of ores in the five years 1912-1916 than in the
previous five years, Benton’s increase in percentage having been
somewhat less than the increase shown by the other two. Ben¬
ton, the greatest producer, put out 28.3% ores in 1907-1911 and
29.4% ores in the years 1912-1916, inclusive. Hazel Green,
ranking third in the order of production in 1907-1911, and
second in 1912-1916, produced ores whose percentages of metal¬
lic zinc were respectively 30.4 and 28.2. Livingston, ranking
fourth as a producer in 1907-1911 and third in 1912-1916,
showed a marked decline in quality of ores produced. In 1907-
1911 the percentage was 30.4 and in 1912-1916 only 24.7.
For the ten years 1907-1916, inclusive, Dodgeville ranked
first, with a percentage of 39.2 and Highland lowest, with a per¬
centage of 27.8. The percentages of the four most important
producers were as follows: Benton, 29.0; Hazel Green, 28.8;
Platteville, 34.6 ; Livingston, 26.5.
The production thus far considered has been for individual
districts in the zinc-producing region of Wisconsin. Total state
production is shown in fig. 14. This chart gives the quantity
of zinc ores produced annually from 1866 to 1875, inclusive, and
annually from 1907 to 1916, inclusive. For the ten years 1866-
1875 only ore statistics were available, but the ten years 1907-
1916 show not only the quantity of ores produced each year, but
the metal obtained from the ores. Percentages indicate the
quality of the ores.
Zinc carbonate was produced before sphalerite, and in the
early years of zinc production more carbonate was produced
than sphalerite. The chart shows this, as well as the steady in¬
crease in sphalerite production. In only two of the years shown
on the chart was there a decrease in sphalerite production from
the previous years, i. e. : in 1873 and in 1913. The production
of carbonate was greatest in the year 1872, which was also the
62
Wisconsin Academy of Sciences^ Arts, and Letters.
Fig. 15a. Value of Zinc Produced 1907-1911, Inclusive
Data from Mineral Resources of the United States
Schuhring — Lead and. Zinc Mining in Wisconsin.
63
Fig. 15b. Value of Zinc Produced 1912-1916, Inclusive
Data from Mineral Resources of the United States
64
Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 15c. Value of Zinc Produced 1907-1916, Inclusive
Data from Mineral Resources of the United States
Millions of dollars
ScJmhring — Lead and Zinc Mining in Wisconsin.
00 _
I
65
5
Sphalerite Carbonate Metal
Fig. 16a. Value of zinc produced in Wisconsin 1907-1911, inclusive.
Value of metal is calculated from average daily quotations at New York
and value of ores is that received by the producer. Data from
Mineral Resources of the United States.
Millions of dollars
66
Wisconsin Academy of Sciences, Arts, and Letters,
C3 O
(XHD
l—i 1—!
OS OS
w re
O Cj
'O’O
3 P
o o
s c
Fig. 16b. Value of zinc produced in Wisconsin 1912-1916, inclusive.
Data from Mineral Resources of the United States.
Millions of dollars
Schuhring — Lead and Zinc Mining in Wisconsin. 67
<N
OS (M
00
oT
lO
w t-
os 00
<N O os
05 «0 lO
CO 00
O b-
b- 1-!
Tj* O
«0 OS
t> 00
to of
OS
lO CO
00 00
tH
’"1
CO
CO
O
05 to
00 CO
^ to'
00
IlL
02
Is
|] fc fc Q
ri <X>
tH c5
bi- W
S
T-1 B
’S
*w ^
I ^
ta
c!
o
8 i
TJ
2 2
Ck
O .S
•S s
s
■g 2
a>
^ Q
o
CD
c
B
tn
I I
3 a
o
s
!
o £
-M
If s
.. I « 1
^ 2 s Q ^ S)
2 'v;3 iS ^ .2
O K CQ ^ S Q
o
►H
pH
Includes 1916 New Digg-ins.
Includes 1916 Potosi.
68 Wisconsin Academy of Sciences, Arts, and Letters.
year of greatest production of zinc ores for the ten-year period
1866-1875. In that year 21,975 tons of zinc ores were pro¬
duced, 13,847 tons of which were zinc carbonate and 8,128 tons
sphalerite. In 1916, when the production of sphalerite was
greatest, 205,481 tons of zinc ores were produced, of which 4,261
tons were zinc carbonate and 201,220 tons sphalerite, (see
table 5). The percentage of carbonate in 1872 was more than
60, and in 1916 less than 2.
Total production of zinc ores in Wisconsin for the years
1866-1875, inclusive, was 107,981 short tons, and for the years
1907-1916, inclusive, 1,029,755 short tons, or about ten times
as much.
The quality of zinc ores deteriorated. In 1907 36.7% of the
ores was metal, and in 1916 the percentage was only 27.6. In
1910 it was less than this, 26.7%, but in all other years of the
ten from 1907-1916 the percentage was higher than in 1916.
For the whole period 1907-1916, inclusive, 307,974 short tons of
metal were obtained from 1,029,755 short tons of ores, or 29.9%.
The value of zinc production in Wisconsin is shown in figs.
15a, 15b, 15c, and 16a, 16b, 16c. Each chart shows the value
of zinc ores produced, as well as the value of the metallic zinc
obtained from the ores. In making a comparison of these val¬
ues, it should be borne in mind that the value of the metal is
calculated from average daily quotations at New York, whereas
the value of the ores is that received by the producer.*
In the five years 1907-1911 only two districts received more
than one million dollars for zinc ores, namely: Benton and
Platteville, each of which received about one and one-half million
dollars. The value of the metal output of the Benton district
was $2,764,008, and of the Platteville district $2,227,318. Hazel
Green, Livingston, and Cuba City-Elmo each produced over one
million dollars worth of metal. In the five years 1912-1916
six districts received more than one million dollars each for ores,
and the value of the metal obtained from the ores was more
than two million dollars for each district. Benton received
$9,013,548 for its zinc ores, and the value of the metallic zinc
was $16,655,266. Hazel Green, the next highest producer, re¬
ceived $2,945,197 for its zinc ores; and the value of the metal
•vvas $5,442,036. The value of ores produced by Livingston was
*In 1915 and. 1916 the value of metallic zinc was calculated from the
average sales price of all grades reported by smelters
Schuhring — Lead and Zinc Mining in Wisconsin.
69
a little more than two million dollars, and the value of ores pro¬
duced by Platteville was a little less than two million dollars.
The value of metallic zinc for each of these districts was about
three and one-half million dollars. Linden, the only other dis¬
trict receiving over a million dollars for its ores, produced metal
worth $2,387,534.
For the ten years 1907-1916, inclusive, seven districts re¬
ceived over a million dollars for their ores, the Cuba City-Elmo
district being added to the districts mentioned above. High¬
land, which received $682,596 for its ores, produced metallic
zinc valued at $1,415,532, and should be added to the foregoing
list of districts producing zinc worth more than a million dol¬
lars in the ten years 1907-1916, inclusive. In this period the
value of Benton ores was $10,563,579, and the value of the metal
obtained therefrom was $19,419,274. Hazel Green, the next
greatest producer of zinc, had ores valued at $3,938,434; and
the values of the metal obtained from them was $7,335,458.
Platteville, Livingston, Mifflin, Linden, Cuba City-Elmo, and
Highland ranked in the order named.
The amount received by Wisconsin producers in each of the
five-year periods was as follows : In 1907-1911, inclusive, about
seven and one-half million dollars for ores, and twelve and one-
half millions for metallic zinc; in 1912-1916, inclusive, almost
twenty millions for ores and 36.5 millions for metallic zinc.
In the ten years 1907-1916, inclusive, Wisconsin producers re¬
ceived $27,339,268 for zinc ores, whereas the value of the metal
obtained from then was $49,179,690. It will be remembered
that the amount received for lead in each of the five-year periods
was in the neighborhood of one million dollars, (figs. 10a, 10b)
and that in the ten years 1907-1916, the amount received by
Wisconsin producers of lead concentrates was $2,267,819; and
the value of the metallic lead, as calculated from average daily
quotations at New York, was $2,877,516, (fig. 10c).
A comparison of figures 10c and 16c shows that some of the
districts rank higher as lead producers than as zinc producers,
although in no case does the value of lead output exceed that of
the zinc. The relative ranks of the two industries is indicated
in the statistics contained in the foregoing paragraph.
For example, the Cuba City-Elmo district, which ranked
seventh in the value of zinc produced, ranked second in lead
Mil
15
14
13
12
11
10
9
8
7
6
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Wisconsin Academy of Sciences, Arts, and Letters.
ns of dollars
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. Value of lead and zinc produced in Wisconsin 1907-1916,
inclusive.
Data from Mineral Resources of the United States.
Schuhring — Lead and Zinc Mining in Wisconsm.
71
production. Benton occupied first place in both industries.
Linden, which ranked sixth in the value of zinc production,
ranked third in the value of lead production. Platteville occu¬
pied third place in the zinc industry, but eighth in the lead in¬
dustry ; and Livingston, with fourth place in the zinc industry,
occupied ninth place in the production of lead. Mifflin was fifth
in zinc, and seventh in lead. Highland and Shullsburg occu¬
pied more important places in the lead industry than in the
zinc industry, whereas Mineral Point, Dodgeville, and Potosi
were among the smallest producers in both industries during
the ten years under consideration.
In comparing the production of metallic lead with the pro¬
duction of metallic zinc, it should be noted that in calculating
the metal content of the ores from assays allowance was made
for smelting and contraction losses in the case of zinc but not
in the case of lead.
The relative importance of the two industries in Wisconsin
in the ten years 1907-1916 is shown in fig. 17 which gives
the value of lead and of zinc produced annually. Prom 1907
to 1910 the value of the lead remained fairly constant. Then
it declined steadily until 1914, after which it again rose. In
1916 the value of the metallic lead was somewhat greater than
in 1910.
The amounts involved in lead values are all less than five hun¬
dred thousand dollars, (table 7). The highest figure is for
metallic lead production in 1916, $419,934. The lowest is for
lead concentrates produced in 1914, $85,196. The lowest figure
for zinc production, on the other hand, is over a million dollars,
received for concentrates produced in 1907 and again in 1908.
The highest figure for zinc is $15,223,204, value of metallic
zinc produced in 1916. In 1907 the value of metallic zinc was
$2,181,820. It dropped to 1.7 millions in 1908 and then rose
steadily until 1912 when it was over 4.5 million dollars. In the
next two years there was a decline to about 3 million dollars.
Then followed a very great increase. In 1915 the value of me¬
tallic zinc rose to more than 10 million dollars, and in 1916 to
$15,223,204.
In 1907 producers received $266,471 for lead concentrates
and $1,037,549 for zinc concentrates, whereas in 1916 they re¬
ceived $291,387 for lead concentrates and $7,464,153 for zinc
72 Wisconsin Academy of Sciences, Arts, and Letters.
concentrates. The amount received for lead was about the same
in 1907 'as ten years later in 1916, but the amount received for
zinc was seven times as great as formerly. And whereas in 1907
the value of zinc concentrates was less than four times as great
as the value of lead concentrates, in 1916 it was more than
twenty-five times as great.
The bar graphs for zinc appearing in this paper are all on
a scale one-twentieth as large as the scale used for similar
graphs for lead.
VI.
Wisconsin Lead and Zinc Production.
RANK IN THE UNITED STATES.
Mine production of metallic lead in the United States, by
states, is shown in figs. 18a, 18b, 18c, for the following periods:
(a) the five years 1907-1911, inclusive; (b) the five years 1912-
1916, inclusive; (c) the ten years 1907-1916, inclusive. Similar
data for the production of metallic zinc are shown in figs. 19a,
19b, 19c.
The states are arranged in the order of their rank. Fig.
18a shows that Wisconsin ranked fifth in lead production for
the five years 1907-1911, with an output of 19,097 short tons
of metal. This amount was somewhat more than was pro¬
duced by Nevada, Arizona, Kansas, Oklahoma, Montana, and
New Mexico. In the next five years, 1912-1916, (fig. 18b) the
production of each of the states just named, and also the pro¬
duction of California, was greater than that of Wisconsin, whose
output fell to 11,306 short tons for the five-year period. Thus
Wisconsin changed from fifth to eleventh place in lead produc¬
tion. For the ten years 1907-1916, Wisconsin’s place was
ninth, (fig. 18c). The large producers were Missouri, Idaho,
Utah, and Colorado, ranking in the order named.
In the production of metallic zinc Wisconsin ranked fourth
in the five-year period 1907-1911, with an out])ut of 115,495
short tons (fig. 19a) and fifth in the five years 1912-1916, with
an output of 192,479 tons. For the ten years 1907-1916, inclu¬
sive, Wisconsin occupied fifth place, with an output of 307,974
short tons.
Schuhrmg — Lead and Zinc Mining in Wisconsin.
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74 Wisconsin Academy of Sciences^ Arts, and Letters.
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Schitbring — Lead and Zinc Mining in Wisconsin.
75
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Schuhring — Lead and Zinc Mining in Wisconsin.
77
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ScJmhring — Lead and Zinc Mining in Wisconsin. 79
Wisconsin’s position relative to the greatest single producing
state and to the United States, in both lead and zinc production,
is shown in fig. 20, which gives the annual production of lead
and zinc 1907-1916, inclusive, for Wisconsin, Missouri, and
the United States. Missouri produced more lead than zinc dur¬
ing those years, whereas Wisconsin produced more zinc than
lead. The United States produced more lead than zinc until
1915 when zinc production exceeded that of lead. In 1916 the
United States produced 701,995 short tons of metallic zinc and
622,975 short tons of metallic lead. Missouri’s output for 1916
was 233,088 tons of metallic lead and 155,960 tons of spelter.
Wisconsin’s lead production remained practically stationary, at
about 3,000 short tons annually for the ten years, but its zinc
production increased to 56,803 short tons of spelter in 1916.
The position of Wisconsin as a zinc producer is higher than
its position as a lead producer, as is shown in figs. 21 and 22.
Fig. 21 shows the amounts of metallic lead produced annually
3907-1916 by the four largest producers and by Wisconsin.
Fig. 22 gives similar data for zinc. Missouri and Idaho are the
largest producers of lead, and Missouri and New Jersey are the
largest producers of zinc, in the United States. In lead produc¬
tion four states stand out pre-eminently: Missouri, Idaho,
Utah, and Colorado, ranking in the order named, (figs. 18a,
18b, 18c). Wisconsin is one of a number of states next in im¬
portance, but producing very much less lead. In zinc produc¬
tion Wisconsin is one of the first five states, (figs. 19a, 19b,
19c) . J. P. Dunlop, in Mineral Resources of the United States,
makes the statement that ‘‘Wisconsin in 1911 and 1912 stood
third and in 1913 and 1914 fourth in i^nk among zinc-produc¬
ing states.” This statement is not borne out by the statistics
appearing in Mineral Resources o fthe United States.®® In 1911
and 1912 Wisconsin’s rank was fourth, and in 1913 and 1914,
fifth, (fig. 22). In only one year during the years under con¬
sideration (1907-1916) did Wisconsin rank third, and that was
in 1908 when Colorado had temporarily dropped from third
place to fourth. In 1909 Colorado regained its place as third
largest producer, and maintained it until Montana took it away
in 1914. In 1916 Montana occupied second place, followed by
New Jersey, Colorado, and Wisconsin.
1914. Part 1, p. 116.
»»1916, Part 1, p. 814.
80 Wisconsin Academy of Sciences, Arts, and Letters.
Thousands of short tons of metal Short Tons
701,995
622,975
233,088
155,960
56,895
3,043
Fig. 20. Annual mine production of lead and zinc, 1907-1916,
inclusive.
Data from Mineral Resources of the United States.
Schuhring — Lead and Zinc Mining in Wisconsin.
81
OO a O r-*C>3 CO
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CD Oi Oi a Oi a Oi Oi a
Fig. 21. Annual mine production of lead, 1907-1916, inclusive.
Data from Mineral Resources of the United States
6
1916
82 Wisconsin Academy of Sciences, Arts, and Letters.
Pig. 22. Annual mine production of zinc, 1907-1916, inclusive.
Data from Mineral Resources of the United States
1916
Schuhring — Lead and Zmc Mining in Wisconsin, 83
The lead production of Wisconsin, which played so important
a part in the early history of the state, is now of small import¬
ance; but the zinc industry, which sprang up after the decline
of lead production had set in, has steadily grown until it has
become of great importance in the state and of considerable im¬
portance in the United States.
Lead Production of the Upper Mississippi Valley
BIBLIOGRAPHY OF STATISTICS
1. 1903. Wisconsin Historical Collections,
a. For 1823: v. 6, p. 290.
Article by Moses Meeker.
Figures are for “Lead product” for
“Wisconsin Lead Region.”
Also V. 8, p. 250.
Article by Moses M. Strong.
Figures are for “Lead produce.”
b. For 1825-1828, inclusive: v. 8, p. 250.
Map published by R. W. Chandler of Galena.
Figures are for “Manufactured lead” from
“Lead Region of the Upper Mississippi.”
c. For 1829: v. 10, p. 79.
Article by James David Butler.
Figures, credited to U. S. Census, are for
“Lead harvest” at
“Galena.”
d. For 1841: v. 13, p. 318.
Letter of Governor Doty to Governor Seward
Figures are for “Lead” from
“Our mines.”
e. For 1845: v. 13, p. 297.
Article by Orin Grant Libby.
Figures are for “Pigs” from
“Galena, Wisconsin, and Iowa.”
(Converted by the writer to short tons on basis of 70 lbs. a pig.)
2. 1840, 1844, D. D. Owen.
Report of a Geological Exploration of part of Iowa, Wisconsin and
Illinois made under instructions from the Secretary of the Treasury
of the United States in the Autumn of the year 1839, by David Dale
Owen, M. D., Principal Agent to explore the Mineral Lands of the
United States.
Document No. 239, House of Representatives, 26th Congress, First
Session.
Report of Geological Exploration of part of Iowa, Wisconsin and
Illinois made from instructions from the Secretary of the Treasury
84 Wisconsin Academy of Sciences, Arts, and Letters.
of the United States in the Autumn of the year 1839; with charts
and illustrations, by David Dale Owen, M. D., Principal Agent to
explore the Mineral Lands of the United States.
Ordered to be printed by the Senate of the United States June 11,
1844.
Document No. 407, Senate, 28th Congress, First Session.
a. For 1823-1829, inclusive: p. 37 of House Document No. 239;
p. 46 of Senate Document No. 407.
Figures, credited to Mr. Legate, are for
“Lead produce based on smelters’ returns” from
“Wisconsin, then called Fever River, Mines.”
b. For 1839: p. 35 of House Document No. 239;
p. 44 of Senate Document No. 407.
Figures are for
“Lead” produced from lead mines of
“Iowa, Wisconsin, and Northern Illinois.”
c. For 1841-1843, inclusive: p. 48 Senate Document No. 407.
Figures, credited to “Register kept at Galena since 1841”
are for
“Pigs” and “Pounds” from
“Upper Mississippi Mines.”
3. 1854.
Edward Daniels, First Annual Report on the Geological Survey
of the State of Wisconsin, published 1854.
a. For 1823-1829, inclusive: p. 39.
Figures, credited to “report of Dr. Owen, made to the general
government in 1840,” are for
“Lead” from the
“Lead district of Wisconsin, Illinois, and Iowa.”
b. For 1841-1853, inclusive: pp. 39, 40.
Figures, credited to Capt. Chas. Beebe, of Galena, are for
“Lead” from the
“Lead district of Wisconsin, Illinois, and Iowa.”
4. 1854.
Josiah Dwight Whitney, The Metallic Wealth of the United
States, described and compared with that of other countries;
Philadelphia, 1854.
a. For 1823-1838, inclusive: p. 421.
Figures, credited to W. H. Bell, Report on Mineral Lands of
have been obtained from records of Messrs. Collier and
the Upper Mississippi, stated (presumably by Bell) to
Kennet, are for
“Metallic lead” from the
“Upper Mississippi Mines.”
Schuhring — Lead and Zinc Mining in Wisconsin.
85
b. For 1845-1853, inclusive, p. 421.
Figures, credited to Capt. Beebe of Galena, are for
“Metallic lead’^ from the
/ “Upper Mississippi Mines.”
5. 1859.
The Merchants’ Magazine and Commercial Review, Conducted
by Freeman Hunt.
a. For 1821-1857, inclusive: v. 40, p. 244.
Figures, credited to E. H. Beebe in Galena Advertiser, are for
“Pigs” and “Pounds” of lead from the
“Galena River Mines.”
b. For 1860: v. 44, p. 703.
Figures are “Pigs” and “Pounds” of “Lead” from the
“Upper Mississippi Lead Mines located in
Illinois, Wisconsin and Iowa.”
6. 1877.
Moses Strong.
Chamberlin, Geology of Wisconsin, 1873-79, v. 2, Pt. 4, pp.
645-752. Moses Strong, Geology and Topography of the
Lead Region.
a. For 1823-1853, inclusive: v. 2, Pt. 4, p. 750.
Figures, credited to J. D. Whitney, are for
“Metallic lead” from the
“Upper Mississippi Lead Mines.”
b. For 1862-1871, inclusive, p. 749.^
Figures are for total product of
“Lead ore” of the
“Wisconsin Lead region.”
c. For 1872-Oct. 1, 1876, inclusive: p. 750.
Figures, credited to N. Corwith & Co., of Galena, are for
“Pigs” and tons of lead from the
“Upper Mississippi Lead Mines.”
(Footnote states: “The weight of a pig of lead is about 72
pounds.” S. L. S. Figures for 70 lb. pigs for 1872 and
1873, and for 72 lb. pigs for 1874, 1875 and to October,
1876.)
7. 1883.
Mineral Resources of the United States.
Department of the Interior, U. S. Geological Survey.
1 The table is for 1862 to Oct. 1, 1876. Figures after 1871 are omitted and
the Corwith data substituted because the latter are for metal arid for entire
Upper Mississippi Valley.
86 Wisconsin Academy of Sciences, Arts, and Letters,
a. For 1873-1880, inclusive; v. 1910, Pt. 1, p. 225.
Article by C. E. Siebenthal.
Figures are for “soft lead” from (a) Missouri; and from
(a) “Wisconsin and Illinois” (called the “Galena Dis¬
trict”),^ V. 1893 and ^84, p. 426, article by C. Kirchoff, Jr.)
b. For 1873-1880, inclusive: v. 1883 and ’84, p. 426.
Article by C. Kirchoff, Jr.
Figures are for “non-argentiferous lead” from the
Mississippi Valley, comprising “Missouri, Kansas, Illinois, and
Wisconsin.”
c. For 1881-1905, inclusive: v. 1910, Pt. 1, p. 225.
Article by C. E. Siebenthal.
Figures are for “soft lead” from the
“Mississippi Valley.” (Production from the Mississippi Val¬
ley from 1894-1905, inclusive, comprises lead content of
ores from Missouri, Kansas, Illinois, Wisconsin, Iowa, Vir¬
ginia, and Kentucky. See v. 1905, p. 364.)
d. For 1906, v. 1910, Pt. 1, p. 228.
Article by C. E. Siebenthal.
Figures are for “soft lead” from
Wisconsin, Iowa, and Illinois, separately given.
e. For 1907-1914, inclusive: v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from the
“Upper Mississippi Valley.” (Upper Mississippi Valley com¬
prises Wisconsin, Iowa, and Northern Illinois.)*-
• The term “Galena District” is applied also to mines in the Missouri-Kansas District, and
care must be taken that the two are not confused.
Examples showing that the term “Galena District” is applied to Illinois and Wis¬
consin:
V. 1883, p.312:
“Output of the Galena district, embracing the production of Illinois and Wisconsin.”
V. 1883 and ’84, p. 426:
“Production of non-argentiferous lead in Missouri, Kansas, Illinois and Wisconsin.”
Short tons.
V. 1910, Pt. 1, p. 225, under the heading “Illinois and Wisconsin” gives the figures here given
under “Galena District.”
Examples showing that the term “Galena District” is applied to mines in the Mis¬
souri-Kansas district:
V. 1892, p. 124 gives ores of Joplin district and includes “Galena.”
V. 1893, p. 95 shows “Galena” as being in Cherokee Covmty, Kansas.
V. 1900, p. 195: “Lead ores in the zinc-lead mines of Southwestern Missouri and Southeastern
Kansas known as the Joplin-Galena district. ”
V. 1903 p. 245, Missouri has (1) Joplin-Galena district, comprising Southwestern Missouri
and Southeastern Kansas, and (2) Southeastern Missouri.
V. 1895, p. 151, “Kansas, Galena District.”
* V. 1914, Pt. 1, p. 38: “The lead and zinc deposits of Illinois fall into two distinct regions
marked by different mineralogical and structural environment. Those of Southern Illinois be¬
long to the Kentucky-Illinois fluorspar district; those of Northern Illinois belong to the Upper
Mississippi Valley region. ”
The amount given for the Upper Mississippi Valley for 1914 (v. 1914, Pt. 1, p. 44) is 1,986
short tons, which may be analyzed as follows:
Wisconsin 1 ,494 (1915, Pt. 1, p. 126)
Iowa None (1916, Pt. 1, p. 63)
Northern Illinois, 492 (1914, Pt. 1, p. 37)
Schubring — Lead and Zinc Mining in Wisconsin.
87
f. For 1915: v. 1915, Pt. 1, pp. 69, 63, and 126.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from
Wisconsin, Iowa, and Northern Illinois, separately given.®
g. For 1916: v. 1916, Pt. 1, p. 59.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from the
**Upper Mississippi Valley.” (Upper Mississippi Valley com¬
prises Wisconsin, Iowa, and Northern Illinois.” ®
8. 1892.
The Mineral Industry, Its Statistics, Technology and Trade, in
the United States and other Countries. Statistical Supplement of the
Engineering and Mining Journal. Founded by Richard P. Rothwell.
a. For 1873-1893, inclusive: v. 2, p. 386.
Article by Walter Renton Ingalls.
Figures are for “lead production” of the
Mississippi Valley, comprising Southeastern Missouri, South¬
western Missouri, Wisconsin, Iowa and Northwestern Illi¬
nois.
b. For 1869-1892, inclusive: v. 2, p. 387.
Article by Walter Renton Ingalls.
Figures, credited to “Geology and Mineral Products of Mis¬
souri by Arthur Winslow,” are for
“Lead production” in
Missouri.
c. For 1873-1892, inclusive:
By subtracting the second set of figures from the first, above
referred to, statistics are obtained for the Upper Mississippi
Valley, comprising Wisconsin, Iowa, and Northwestern Illi¬
nois.’
9. 1894.
Arthur Winslow.
The Missouri Geological Survey, v. 6, Sec. 1, pp. 135-151. Lead
and Zinc Deposits of the Mississippi Valley, by Arthur Winslow.
a. For 1823-1853, inclusive: p. 146-7.
* The 1915 amount for Upper Mississippi Valley is made up as follows:
Wisconsin 2 ,322 short tons (v. 1915, Pt. 1, p. 125)
Iowa, None (v. 1915, Pt. 1, p. 63)
Northern Illinois, 495 (v. 1915, Pt. 1, p. 59)
• The amount given for the Upper Mississippi Valley for 1916 (v. 1916, Pt. 1, p. 59) is 3,519
short tons, which may be analyzed as follows:
Wisconsin, 3,043 (v. 1916, Pt. 1, p. 57)
Iowa, 14 (v. 1916, Pt. 1, p. 57)
Northern Illinois 462 (v. 1916, Pt. 1, p. 70)
Upper Mississippi Valley 3 ,519 fv. 1916, Pt. 1, p. 59)
’ Note. — Ingalls, in Mineral Industry, v. 2, p. 387, makes the following comment: “It will
be noted that the product reported for Missouri in the preceding table is in a few cases greater
than that stated for the Mississippi Valley in the general table of production of Lead in the
United States. More or less of the Missouri ore is shipped to silver-lead smelters, by whom it is
reported as ‘desilverized’, and it is not credited to the States producing it, except since 1890,
when the reports made by the smelters to The Mineral Industry have been divided so that the
production could be traced to its true source. ’’
88 Wisconsin Academy of Sciences^ Arts, and Letters.
Figures^ credited to Whitney ‘‘234, p. 421/’ * are for
”Tons of lead, produced by the
“Whole region.”
b. For 1862-Oct. 1, 1876, inclusive: p. 146.
Figures are for tons of “lead ore” produced in
“Wisconsin.^
c. For 1872-October, 1876: p. 147.
Figures, credited to Mr. Strong, “39, vol. ii, p. 750” are for
“Tons of lead.”
(Region not specified) “
d. For 1873-1879, inclusive: p. 147.
Figures, credited to “Mineral Resources of the United States
for 1882” are for
“Tons of lead.”
(Region not specified.)
e. For 1880: p. 147.
Figures, credited to “Census of 1880,” are for
“Tons of lead ore” from
Wisconsin, Illinois, and lowa.^^
f. For 1889: p. 148.
Figures for “Census year of 1889” are for
“Tons of lead ore” from
Wisconsin, Illinois, and Iowa (No production) .“
10. 1911.
0. C. Gillett, Lead and Zinc Production of the Upper Mississippi
Valley, B. A. Thesis at University of Wisconsin, 1911; Approved by
C. K. Leith.
a. For 1823-1853, inclusive; p. 9.
Credited to J. D. Whitney, Metallic Wealth of the United
States.
b. For 1862-1880, inclusive; p. 10.
Credited to “Winslow’s Geology of Missouri.”
c. For 1881-1892, inclusive: p. 10.
Credited to W. R. Ingalls, Mineral Industry.
d. For 1893-1906, inclusive: p. 10.
Estimated by Gillett at 1,500 annually.
8 The first number of the quotation refers to the Winslow’s bibliography. No. 234 being
D. Whitney’s Metallic Wealth of the United States.
9 The first number of the quotation refers to the Winslow’s bibliography, No. 39 being Cham¬
berlin’s Geology of Wisconsin, 1873-79.
w J. D. Whitney’s Metallic Wealth of the United States, p. 421, gives figures for 1823-1853,
for tons of metallic lead produced by the Upper Mississippi Mines. The figures do not corres¬
pond with those given by Winslow. By adding 12% to each of Whitney’s metallic lead figures,
amounts are obtained identical with Winslow’s.
It may have been Winslow’s intention to convert Whitney’s metallic lead into ore.
These are figures converted by Winslow from pounds of lead ore produced in Wisconsin
1862 to Oct. 1, 1866, as given by Moses Strong, p. 749 Geology of Wisconsin 1873-’79, v. 2., Pt4.
19 These figures, credited to Strong, are called by Winslow “Tons of Lead.” Strong’s figures,
y. 2, p. 760, Geology of Wisconsin, 1873-’79, are for pigs and tons produced in the Upper Mfae,
issippi Mines. The lead here referred to is therefore metal.
1* V. 1883 and ’84, of Mineral Resources, p. 426, identifies these figures as “non-ar^entferous-
lead” from Wisconsin and Illinois. Also v. 1910, Pt. 1, p. 225, where the lead is designated as
“soft lead.” These statistics are therefore for metal.
‘■i Winslow designates the amounts for 1880 and 1889 as being lead ore.
Schuhring — Lead and Zinc Mining in Wisconsin,
89
e. For 1906-1908, inclusive: p. 11.
Credited to Mineral Resources.
f. For 1909-1910, inclusive: p. 11.
Credited to J. E. Kennedy, Platteville.
11. 1920.
Compilation by S. L. Schubring.
a. For 1823-1863, inclusive:
J. D. Whitney, The Metallic Wealth of the United States,
p. 421.
Whitney’s compilation is made up as follows:
For 1823-1838, inclusive:
Figures, credited to W. H. Bell, Report on Mineral Lands
of the Upper Mississippi, stated by Bell to have been
obtained from records of Messrs. Collier and Kennet,
are for
^‘Metallic lead” from the
“Upper Mississippi Mines.”
For 1839-1844, inclusive:
Figures are for
“Metallic lead” from the
“Upper Mississippi Mines.”
For 1845-1853, inclusive:
Figures, credited to Capt. Beebe of Galena, are for
“Metallic lead” from the
“Upper Mississippi Mines.”
b. For 1854-1857, inclusive:
Hunt’s Merchants’ Magazine and Commercial Review, v. 40,
p. 244.
Figures, credited to E. H. Beebe in Galena Advertiser, are for
“Pigs” and “Pounds” of lead from the
“Galena River Mines.”
c. 1858-1859, inclusive: No data.
d. For 1860:
Hunt’s Merchants’ Magazine and Commercial Review, v. 44,
p. 703.
Figures are for “Pigs” and “Pounds” of lead from the
“Upper Mississippi Lead Mines located in
Illinois, Wisconsin and Iowa.”
e. 1861. No data.
f. For 1862-1871, inclusive:
Chamberlin: Geology of Wisconsin, 1873-’79, v. 2, Pt. 4, p.
749-750.
Moses Strong, Geology and Topography of the Lead Region.
Figures are for “total product of lead ore” of the
“Wisconsin lead region.”
90 Wisconsin Academy of Sciences, Arts, and Letters.
(11. Compilation by S. L. Schubring, continued.)
g. For 1872: Ibid, p. 750.
Figure, credited by Strong to N. Corwith & Co., is for
*‘Lead” from the
‘^Upper Mississippi Lead Mines.”
(Note.^ — Since the figure is given also in “pigs,” this is me¬
tallic lead.)
h. For 1873“1880, inclusive;
Mineral Resources of the United States, v. 1910, Pt. 1, p. 225.
Figures are for “Soft lead” produced by
Wisconsin and Illinois.
i. For 1881-1892, inclusive:
The Mineral Industry, Statistical Supplement of the Engineer¬
ing and Mining Journal, v. 2, pp. 386, 387.
Missouri “lead production” credited to Geology and Mineral
Products of Missouri by Arthur Winslow.
By subtracting Missouri product from production of the Mis¬
sissippi Valley, statistics are obtained for
“Lead production” of the
Upper Mississippi Valley, comprising Wisconsin, Iowa, and
Northwestern Illinois.
j. 1873-1905. No data.
k. For 1906: Mineral Resources of the United States, v. 1910, Pt. 1,
p. 228.
Article by C. E. Siebenthal.
Figures are for “soft lead” from
Wisconsin, Iowa, and Illinois, separately given.
l. For 1907-1914, inclusive: Ibid., v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from
“Upper Mississippi Valley.” (Upper Mississippi Valley com¬
prises Wisconsin, Iowa, and Northern Illinois.) ^
m. For 1915: Ibid., v. 1915, Pt. 1, pp. 59, 63, and 126.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from
Wisconsin, Iowa, and Northern Illinois, separately given.^*
V. 1914, Pt. 1, p. 38: “The lead and ssinc deposits of Illinois fall into two distinct r^ons
marked by different mineralogical and structural environment. Those of Southern Illinois be¬
long to the Kentucky-Illinois fluorspar district; those of Northern Illinois belong to the Upper
Mississippi Valley region. ”
The amount given for the Upper Mississippi Valley for 1914 (v. 1914, Pt. 1, p. 44) is 1,986
short tons, which may be analj^ed as follows:
Wisconsin, 1 ,494 (1915, Pt. 1, p. 126)
Iowa, None (1915, Pt. 1, p. 68)
Northern Illinois, 492 (1914, Pt. 1, p. 37)
1,986
The 1915 amount for Upper Mississippi Valley is made up as follows
Wisconsin, 2 ,322 short tons (v, 1915, Pt. 1, p. 125)
Iowa None (v. 1915, Pt. 1, p. 63)
Northern Illinois, 495 (v. 1915, Pt. 1, p. 59)
Schuhring — Lead and Zinc Mining in Wisconsin.
91
n. For 1916: Ibid., v. 1916, Pt. 1, p. 59.
Article by J. P. Dunlop.
Figures are mine figures for lead (metal) from
“Upper Mississippi Valley.” (Upper Mississippi Valley
comprises Wisconsin, Iowa, and Northern Illinois.) ”
Zinc Production op the Upper Mississippi Valley
BIBLIOGRAPHY OF STATISTICS.
1. 1877.
Moses Strong.
Chamberlin, Geology of Wisconsin, 1873-79, v. 2, Pt. 4, pp. 645-752.
Moses Strong, Geology and Topography of the Lead Region.
a. For 1860-Oct. 1, 1876; v. 2, Pt. 4, pp. 742, 843.
Statistics, prepared from the books of the four zinc manufac¬
turing companies at La Salle, Ill., are for
Smithsonite and Blende from
Mineral Point, Platteville, Council Hill, and Galena.
” The amount given for the Upper Mississippi Valley for 1916 (v. 1916, Pt. 1, p. 59) is 3,519
short tons, which may be analyzed as follows:
Wisconsin, 3 ,043 (v. 1916, Pt. 1, p. 57)
Iowa, 14 (v. 1916, Pt. 1, p. 57)
Northern Illinois, 462 (v. 1916, Pt. 1, p. 70)
Upper Miss. Valley 3 ,519 (v. 1916, Pt. 1, p. 59)
92 ^yisconsin Academy of Sciences, Arts, and Letters.
2. 1883—
Mineral Resources of the United States, Department of the Interior,
United States Geological Survey.
a. For 1886: v. 1886, p. 156.
Article by C. Kirchofif, Jr.
Statistics, credited to Thos. H. McElroy, are for ores shipped
from
Shullsburg and Benton:
From Shullsburg - 506 short tons “Drybone”
209 short tons ‘‘Black Jack”
From Benton _ 2,275 short tons both classes.
2,990 short tons
b. For 1889: v. 1889 & 1890, p. 88.
Article by C. Kirchoff.
Statistics, credited to census report covering calendar year
1889 are for zinc ore produced in Wisconsin.
c. For 1904 and 1905: v. 1905, Pt. 1, p. 384.
Article by H. Foster Bain.
Statistics are estimates for ore produced by
Wisconsin, Illinois, and Iowa.
d. For 1906: v. 1906, Pt. 1, p. 473.
Article by J. M. Boutwell.
Statistics are for zinc ores produced in Wisconsin.
e. For 1907-1914, inclusive: v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Statistics are for zinc concentrates produced in the
Upper Mississippi Valley region.
f. For 1907-1914, inclusive: v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Statistics are for metallic zinc produced in the
Upper Mississippi Valley region.
g. For 1915: v. 1915, Pt. 1, pp. 59, 63, 125.
Article by J. P. Dunlop.
Statistics are for zinc concentrates from
Wisconsin, Iowa, and Northern Illinois, separately given."
h. For 1915: v. 1915, Pt. 1, pp. 59, 63, 126.
Article by J. P. Dunlop.
Statistics are for metallic zinc from
Wisconsin, Iowa, and Northern Illinois, separately given.”
The 1915 amount for Upper Mississippi Valley is made up as follows:
Wisconsin, sphalerite, 138 ,996 S. T. (v. 1915, Pt. 1, p. 1251
carbonate, 2 ,579 S. T. (v. 1915, Pt. 1, p. 125)
Northern Illinois, sphalerite, 20 ,454 S. T. (v. 1915, Pt. 1, p. 59)
Iowa, None (v. 1915, Pt. 1, p. 63)
162 ,029
The 1915 amount for Upper Mississippi Valley is made up as follows:
Wisconsin, 41 ,403 S. T. (v. 1915, Pt. 1, p. 126.)
Northern Illinois, 5 ,534 S. T. (v. 1915, Pt. 1, p. 126.)
Iowa, None (v. 1915, Pt. 1, p. 63.)
46 ,937
Schuhring — Lead and Zinc Mining in Wisconsin. 93
i. For 1916: v. 1916, Pt. 1, p. 68.
Article by J. P. Dunlop.
Statistics are for metallic zinc produced in the
Upper Missisippi Valley. (Upper Mississippi Valley com¬
prises Wisconsin, Iowa, and Northern Illinois.^
j. For 1916: v. 1916, Pt. 1, p. 58.
Article by J. P. Dunlop.
Statistics are for metallic zinc produced in the
Upper Mississippi Valley. (Upper Mississippi Valley com¬
prises Wisconsin, Iowa, and Northern Illinois.'^^
k. Total for 1821-1914, inclusive: v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Statistics are for zinc concentrates produced in the
Upper Mississippi Valley, comprising Wisconsin, Iowa, and
Northern Illinois.
l. Total for 1821-1914, inclusive: v. 1914, Pt. 1, p. 44.
Article by J. P. Dunlop.
Statistics are for metallic zinc produced in the
Upper Mississippi Valley, comprising Wisconsin, Iowa, and
Northern Illinois.
3, 1892—
The Mineral Industry, Its Statistics, Technology and Trade, in
the United States and other countries. Statistical Supplement of the
Engineer and Mining Journal. Founded by Richard P. Rothwell.
a. For 1891, 1892: v, 2, p. 625.
Statistics, credited to Prof. W. H. Seamon in the Seventh An¬
nual Report of the State Mine Inspector of Missouri, p. 160,
are for
Zinc ore produced in
Wisconsin.
b. For 1892, v. 1, p. 132.
Article by C. Kirehoff.
Statistics, credited to Prof. W. P. Blake, are for
Shipments from Benton, the principal ore-shipping station.
c. For 1893: v. 2, p. 625.
Statistics, credited to W. P. Blake, are for
Zinc ore produced in
Wisconsin.
d. For 1900: v, 9, p. 666.
Article by E. R. Buckley.
The amount given for the Upper Mississippi Valley for 1916 (v. 1916, Pt. 1, p. 58) is 220,777
short tons, which may be analyzed as follows s
Wfaconsin, carbonate, 4 ,261 S. T. (v. 1916, Pt. 1, p. 146.)
sphalerite, 201 ,220 S. T. (v. 1916, Pt. 1, p. 146.)
Iowa, sphalerite, 85 S. T. (v. 1916, Pt. 1, p. 75.)
Northern Illinois, sphalerite, 15 ,211 S. T. (v. 1916, Pt. 1, p. 70.)
^ ■ I
220,777
21 The amount given for the Upper Mississippi Valley for 1916 (v, 1916, Pt. 1, p, 58), is 60,228
short tons, which may be analyzed as follows!
Wisconsin, 56 ,808 S. T. (v. 1916, Pt. 1, p. 70a.)
Iowa, 21 S. T. (v. 1916, Pt. 1, p. 53a.)
Northern Illinois, 3 ,404 S. T. (v. 1916, Pt. 1, p. 70 .)
60 ,228
94 Wisconsin Academy of Sciences, Arts, and Letters.
Statistics are for zinc ore shipped from the
more important centers of the region.
e. For 1904, v. 19, p. 674.
Statistics, credited to Bain, are for
Zinc ore produced in
“Wisconsin.” (Probablj?’ Upper Mississippi Valley.)
f. For 1905-1910, inclusive: v. 19, p. 674.
Statistics are for
Zinc ore produced in
“Wisconsin.” (Probably Upper Mississippi Valley.)
g. For 1906, 1907: v. 16, p. 915.
Article by J. E. Kennedy.
Statistics are for
Zinc ore from
Wisconsin, Illinois, and Iowa.
h. For 1910-1915, inclusive: v. 24, p. 720.
Article by R. L. Bartlett.
Statistics are for
Zinc ores received by smelters'^* from
Wisconsin, including Illinois and Iowa.
i. For 1916: v. 26, p. 750.
Article by Jesse A. Zook.
Statistics are for zinc ores produced in
Wisconsin.
4. 1894.
Arthur Winslow.
The Missouri Geological Survey, v. 6, Sec. 1, pp. 135-151. Lead
and Zinc Deposits of the Mississippi Valley, by Arthur Winslow.
a. For 1860-1876, inclusive: pp. 148, 149.
Statistics, credited to Moses Strong, are for
Zinc ores from
Wisconsin and Illinois.
b. For 1879: p. 149.
Statistics, credited to Irving are estimates for
Zinc ores from
“the region.”
22 Bain gives this figure for Upper Mississippi Valley.
23 In V. 16, p. 915 J. E. Kennedy gives identical 1905 production and says it is for Wisconsin
Illinois, and Iowa.
In V. 22, p. 781, 1910 production (same amount as here given) is for “Wisconsin, including
Illinois and Iowa.*’
2* Does not include the production of ore exported or what was taken by
the manufacturers of zinc oxide. For 1910 and 1911 reports missing from
three small smelters.
V. 24, p. 720 incorrectly gives 1914 production as 71.311 short tons. The
correct amount is 74,311. (See v. 23, p. 770.)
Schuhring — Lead and Zinc Mining in Wisconsin.
95
c. For 1880: p. 149.
Statistics, credited to the ‘‘Census”, are for
Zinc ore from
Wisconsin and Illinois.
d. For 1889: p. 149.
Statistics “for the census year 1889,” are for
Zinc ore from
Iowa and Wisconsin.
e. For 1893: p. 149.
Statistics, credited to Blake, are for
Zinc ore.
5. 1902.
E. T. Hancock.
Tenth Biennial Report of the Bureau of Labor and Industrial Sta¬
tistics. State of Wisconsin 1900-1901. Halford Erickson, Commis¬
sioner, pp. 1075-1093. The Lead and Zinc Industry of Southwestern
Wisconsin. Prepared by Eugene T. Hancock, University of Wiscon¬
sin.
a. For 1876-1901, inclusive: p. 1090-1.
Statistics, prepared from reports of smelters, are for
Zinc ores produced in
Wisconsin.
6. 1907.
H. Foster Bain.
Zinc and Lead Deposits of the Upper Mississippi Valley, Wiscon¬
sin Geological and Natural History Survey, Bulletin No. xix. Eco¬
nomic Series, No. 12.
7. 1911.
O. C. Gillett, Lead and Zinc Production of the Upper Mississippi
Valley, B. A. Thesis at University of Wisconsin, 1911; Approved by
C. K. Leith.
a. For 1860-1876, inclusive:
Statistics, credited to “Winslow’s Geology of Missouri” are
for metallic zinc‘® from the
Upper Mississippi Valley.
b. For 1877-1901, inclusive:
Statistics, credited to E. T. Hancock, Report for Bureau of
Labor Statistics, 1902, are for
Metallic zinc“® from the
Upper Mississippi Valley.
c. For 1906-1910, inclusive:
Statistics, credited to C. E. Siebenthal of the United States
Geological Survey, are for
Metallic zinc from the
Upper Mississippi Valley.
Converted from ores on the basis of 35%.
Iable 1. Summary of lead production of the Upper Mississippi Valley, stated in short tons of metallic lead.
Statistics from various authorities as indicated in Exhibit I.
96
Wisconsin Academy of Sciences, Arts, and Letters.
o
o
o
'<1'
o
CO
(c) Should be 11,976 (d) Should be 7,553 (e) Should be 2,511.
Statistics from H. F. Bain: Zinc and Lead Deposits of the Upper Mississippi Valley.
Table 2. Production of lead ore Jby furnace districts, stated in pounds. Taken from Chamberlin* s Geology of Wis-
consin, Vol. 2, Part A, pp. 6U5-752.
Schubring — Lead and Zinc Mining in Wisconsin.
7
98
Wisconsin Academy of Sciences, Ai'ts, and Letters.
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(a) Includes a few tons of lead carbonate.
^ ^ ^ ^ Pt. l,p. 5U; 1908, Pt. l,p. 6S9; 1909, Ft. 1, 529; 1910, Pt. 1, p. 668; 1911, Pt. 1, p. S65; 1912, Pt. l,p. SIO; 191S, Pt. 1, p. 160; 191i, Pt. 1, p. 115; 1915, Pt. 1, p 125; 19ie, Pt. 1, p. lie.
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Table 4. Annval production of Metallic lead from 1907 to 1916, inclusive, by districts. The quantity is stated in short tons and the value in dollars. The value is calculated from the average daily quotations at New York and St. Louis. From Mineral Resources of the
United States, 1907, Pt. 1, p. SU; 1908, Pt. 1, p. 689; 1909, Pt. 1, p. 529; 1910, Pt. 1. p. 668; 1911, Pt. 1, p. 868; 1912, Pt. 1, p. 510; 1918, Pt. 1, p. 160; 19U, Pt. 1, p. 115; 1915, Pt. 1, p. 126; 1916, Pt. 1, p. U8.
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Table 6. Annual production of metallic zinc from 1907 to 1916, inclusive, by districts. The quantity is stated in short tons and the value in dollars. In calculating the metal content of the ores from assays, allowance has been made for smelting and concentration
losses. The value from 1907 to 191U, inclusive, is calculated from the average daily quotations at New York and St. Louis. For 1915 and 1916 the average sales price reported by the smelters has been used. Data from Mineral Resources of the United States
1907, Ft. 1, p. 5U; 1908, Ft. 1, p. 639; 1909, Ft. 1, p. 529; 1910, Ft. p. 668; 1911, Ft 1, p. 863; 1912, Ft. 1, p. 510; 1913, Ft. 1, p. 160; 1911,, Ft. 1, p. 115; 1915, Ft. 1, p. 126; 1916, Ft. 1, p. 11,6.
Mineral Point
Year
1907.. ..
1908.. ..
1909.. ..
1910- .-.
1911- -.
1907-11
1912- --
1913- --
1914.. .
1915.. .
1916.. .
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
Short Tons Dollars Short Tons Dollars
Short Tons Dollars
Short Tons Dollars
3,487
3,833
3,838
5,856
8,292
411 .466
360 ,302
632 ,448
945 ,288
1,147
2,740
3,636
135,346
257 .560
392 ,688
45.194
16.544
29,484
23.544
14 .364
3,238
6,300
520 ,380
244 ,118
61 ,020
349 ,704
718 ,200
1 ,041
1 .340
2,313
122 ,838
125,960
249 ,804
194,508
146,122
286 ,150
97 ,760
137 .052
131 ,976
219,222
2,280
2,843
2,410
186 ,086
168 ,918
246 ,240
307 ,044
274 ,740
895
1 ,053
2,456
2,081
2,569
105 ,610
98 ,982
265 ,248
224 ,748
292 ,866
107 ,970
81 ,404
70,308
70 ,092
19,038
14 ,364
42 ,228
1 ,815
2,764
5,432
5,168
5,338
214 ,170
259 ,816
586,656
558,144
608 ,532
16,200
31 .644
18 .490
18 ,206
23,152
25,927
29 ,720
2 ,181 ,820
1,711,364
2.500 ,416
2,800,116
3 ,388 .080
1907
. . 1908
_ 1909
. 1910
. 1911
2,940 322,158
.560 .918
,113,952
,285 ,812
1 .026
1 ,163
141 ,588
130 ,256
62,118
96 ,472
201 ,536
35 ,742
8,624
23 ,562
5,703
5,696
6,973
5,897
787 ,014
637 ,952
707 ,370
1 ,729 .304
1 ,580 ,396
62,118
119,288
211,452
1 ,908
2,655
1 ,947
2,430
3,827
263 ,304
297 ,360
198 ,594
602 .640
,025 .636
2,439
3,081
4 ,042
4 ,997
4,536
336 ,582
345,072
412 ,284
1 ,239 ,256
1 ,215,648
2.833
1 ,443
2,560
3,009
4.227
390 .954
161 .616
261 ,120
746 ,232
,132 .836
8,004
33 ,824
13,056
49,504
38 ,964
54,884 1 ,217 142,702
Short Tons Dollars
158 39,184
914 244,952 •
3,160
(e) 5,134
769 .902
490 ,784
90 ,882
783 ,680
(e)
1,375,912
70,104
61 .488
17 ,646
68 ,200
243 ,880
4 ,560 ,900
3 ,372 ,320
3 ,173 .626
10 ,267 ,944
15,223 ,204
107,972 19,419,274
Cuba City-Elmo: 14 ,477 1 ,747 ,822
48.314 7,353.458
.126 3,680,212
(d) 4.272 (d)545,764 (d) 2,729 (d) 392 ,890
39,663 5,738,478
307.974 49,179.690 1907-1916
(a) 1916 production is
for Benton-New Dig¬
gings.
b) Short Tons Dollars
Cuba City 1907-1909 7,623 785,594
. . 75 8,100
2 .940 322 ,168
(c)
Mineral Point 1912-14
Montlort 1912-14
Mineral Pt.-MontforC
1915-16
Mineral Pt.-Montfort
1912-16
Short Tons Dollars
488 54 ,884
1 ,217 142 ,702
2 .777 481 .722
(dl The joint production of Mineral Point-Montfort 1915-16 is arbitrarily divided equally in computing five-year Cutals for k
‘ ■' <'f^> % >>»y'-.n’,^m ,.^«r wi <aOa>^r'H,VV' ,>-..-.v>^
,w^‘.‘> .';c. „ \ .vn , ;■ . • ’ ' ■"•/ ■
ri?
'J'{
'(.fi > i'<kf’f
. -v*
'^m
t
■■■ :J\'i)'iif ■ ■ f- ■-. .-•,
"'i'^P' *’•'■* H' • ( - M'M'-, > k’lSl > ^i‘. ,
V'->
-t-
■ /..''V ' ' k W?,.<V.':
' ' >,-^S < ~ •. ')«jucv< ' kj<«i.,T n‘.,'V:
■v^rA y ^cs VUif, Ji
^;r;. ,
! . 'f' ^-■' "It '
, f '.i it ,\.)i ,^.Ji
h> ...
( iK
1-.
. ! VX’f-. G,.f
•* '•• '.i
: *
• ] S-'i-r.
; ■ - .':^-;-
*i>V, f!'
fe;.r-
« t. ,■ .,
I:
"If
!
i soc,,
■ 6'Jir -...I v!
<
. „|, ir^ " ’-JJ'.v i\
45 s v A
^ii\ V feOi-: %h-:} tJhf.:}'
«h.s-> ot^'s . ■«‘«<'«ia-V*'>,*l;--> . • . J
*;:.• yi i '"; , "•;‘1j\;i.'“:r'' , ■ ., „ , , ■;., ■ i
■ ^ . :m .
- . r0
&
M
^<.r' K.''.?'.'‘.>''
■A.' 3''^ '
,"V'';'-;c.'
. Y(
1907.
1908.
1909.
1910.
1911.
1907-
1912.
1913.
1914.
1915
1916
1912-
Table 8. Mine ■production of lead by states from 1907 to 1916, inclusive, stated in short tons of metallic lead. Data for mine production of lead are not available for all states before 1907. These
figures of mine production are obtained from mine operators and not from smelters. From Mineral Resources of the United States, 1914, Ft. 1, p. 810; 1915, Ft. 1, p. 731; 1916, Ft. 1, p. 845.
(a) By-product from unknown quantity fluorspar.
(b) Includes Pennsylvania.
Table 9. Mine production of zinc by states from 1907 to 1916, inclusive, stated in short tons of metallic sine. From Mineral Resources of the United States, 1916, Pt. 1, p. 8H.
FAUNA OF THE GALENA LIMESTONE NEAR
APPLETON
John W. Ockerman
The region under discussion, namely Appleton and the near
cities of Neenah, Menasha, Mackville and Kaukauna, lies in
the belt of Galena Limestone that cuts tlie state from north to
south. This rock has been quarried in large amounts in this
region and there are many quarries, some being worked exten¬
sively and others remaining as evidence of former operations.
The Neenah, Menasha and Kaukauna quarries are the most
important of the quarries that are being worked at present.
The quarry near Mackville has not been worked for several
years, its ledges being greatly weathered during that time.
There are also several quarries near Neenah and Appleton that
have been in disuse for a number of years.
The Neenah* quarry is located near the southern limits of the
city of Neenah. It has been worked for a large number of years
and is at present being worked by the city of Neenah. The lime¬
stone is massive and the layers are quite thick. The limestone
is underlaid with blue shale beds. At the joints and bedding-
planes there is considerable mineralization and the rock has
much Galena and Fluorite. The surface of the Galena is cov¬
ered with glacial till. This quarry has had considerable weath¬
ering and so yields some good specimens, there being an abun¬
dant and varied fauna there.
The Mackville quarry is located near Mackville and is much
smaller than the Neenah quarry. As has been stated the quarry
has been in disuse for a long period of time and so has been
greatly weathered. The Galena here is thin bedded and has
been used in the past as road metal. The large amount of
weathered surface and the extent of the weathering Ijoth go to
make this an excellent source of fossil remains. It has a great
variety and number of fossils and has yielded some very fine
specimens.
100 Wisconsin Academy of Sciences, Arts, and Letters.
The quarry at Kaukauna is a very large one and is exten¬
sively worked. The limestone is massively bedded in layers
from six to twenty inches in thickness. Owing to the widely
separated joint planes slabs of large dimension can be extracted.
This quarry, although abundant in fossil remains, does not pro¬
duce many because of the lack of weathered surfaces and the
very massive character of the rock. There is one unusual fea¬
ture of the quarry and that is the ripple marked bottom. The
ripple marks are at least six inches deep and widely spaced in
sinuous curves. The water was evidently deep but because of a
strong current and powerful waves the sediment was laid down
in cur^^ed bands. There are other ripple marked limestones in
northern Wisconsin, as the Niagara Limestone, near Marble¬
head but none is as pronounced as the Galena at Kaukauna.
The Menasha quarry is located about half way between Apple-
ton and Menasha. It is a large quarry and at present is being
worked to a large extent. The limestone is massively bedded
and is very compact. The older workings of this quarry are
shallow and considerably weathered but are not abundant in
fossil remains. The pit being worked yielded only a few fos¬
sils, these being found on exposed bedding planes.
The dredgings at Kimberly are a thin bedded limestone and
shaly material and yielded a considerable number of Grapto-
lites.
The Galena Limestone formation that underlies the Fox
Eiver valley strikes north 27 degrees east and has a horizontal
width of about 18 to 20 miles. The formation extends entirely
across the state from its northern exposure at Green Bay to Illi¬
nois passing between Lake Geneva on the east and just east of
Beloit on the west. The formation dips very gently to the south¬
east about 20 feet to the mile in this region in a monoclinal
fashion and as a consequence varies in thickness at different
points on the outcrop. Even along the strike it is not uniform
in thickness. At Neenah, according to artesian well records, it
is 62 feet thick, at Appleton 65 feet, Kimberly 127 feet and at
Kaukauna 176 feet. The reports of the Wisconsin Geological
Survey give the thickness of the Galena as 125 to 250 feet.
The formation consists of a coarse gray and blue magnesian
limestone the proportion of magnesia varying in different locali¬
ties, but never being quite the percentage of a theoretical dolo¬
mite. In this locality the Galena is excellent material for road
Ockerman — Fauna of Galena Limestone Near Appleton 101
work and for concrete and building material. The rock is
usually quite thickly bedded but in some places it is interbedded
with layers of blue shale and shelly limestone. In such cases
it requires a great deal of sorting and rejection of quite a large
amount of the rock quarried.
The color of the Galena is normally a clear light gray to al¬
most white but as a rule the Galena is a bluish gray due to some
alumina clay content. Upon weathering the Galena has a dull
gray color and is very soft if the surface has been exposed a
long time. Often the fossils weather entirely out of the rock,
this being the case at Mackville.
The Galena Limestone of this area has offered not only a great
abundance of fossils but also a great variety. All of the quar¬
ries have abundant and varied fossil remains, which are as a
rule in a good state of preservation. Nearly every group of ani¬
mal life is found here. The Protozoa are not present nor are
the Vermes. The Cystoidea and Echinoidea of the Echinoder-
mata are absent and the Pelecypoda of the Mollusca likewise,
but the remaining groups are found in this region.
Protozoa are not present. The Coelenterata are represented
by sponges, corals and the Graptolites.
Only four specimens of the Porifera were found and in prac¬
tically all cases the specimens were poorly preserved. Mack¬
ville was the chief source of the sponges, but they are found in
the other quarries rarely.
The Anthozoa that were found in this region are typical of
the Ordovician strata. Streptelasma w^as the prevalent type
and three species were found more or less abundant and well
preserved. One species of the Tabulata was found at Neenah
but that was poorly preserved.
The Graptolites have been found abundantly at Kaukauna
and at Kimberly and were all well preserved. Their carbonized
skeletons are very well adapted to fossilization. Pour species
of the Diprionidae have been found and these have been dis¬
cussed by Dr. R. M. Bagg in his report of 1923 (‘‘Recent Dis¬
covery of Wisconsin Graptolites.”)
Of the Echinodermata only the Crinoidea are present. Five
species were found, being fairly abundant and well preserved.
Heterocrinus was the most abundant type and found chiefly at
Neenah and Mackville, but occurring in the other quarries
102 Wisconsin Academy of Sciences, Arts, and Letters.
rarely. The specimens were stem fragments and were usually
very small.
Both groups of the Molluscoidea were found abundantly in
this region; the Bryozoa were not as abundant as the Brachio-
poda, but were found in all of the quarries more or less abun¬
dantly. Six species were found but were not well preserved, the
internal structure being obliterated and often the external struc¬
ture as well.
The Brachiopoda were found in great abundance at all the
quarries, especially at Mackville and Neenah. Twenty-four
species were identified and all were well preserved. Some of
the types were rare, as the Lingula, while others were very abun¬
dant, as the Khynchotrema. The valves were usually found to¬
gether but often single shells were found showing both the in¬
ternal and external structure.
The Gastropoda were very abundant, eighteen species being
found all of which were preserved as a cast of the shell. None
of the species were abundant with the exception of Murchisonia
gracilis. All of the quarries revealed several types of Gastro¬
poda.
The Cephalopoda were found fairly abundant at all of the
quarries but often they were so weathered that they did not
offer an opportunity of study and identification. Four species
were identified, some of them being well preserved.
The Trilobites vrere rare and found chiefly at Mackville, al¬
though Neenah has a few. They are usually well preserved but
rarely are they in complete form. Usually either a head shield
or tail shield is found, as is the case of Illaenus, only head
shields being found.
This work has been done to make more clear the paleontologi¬
cal side of the geology of this region and the writer has taken
great pleasure in making this as complete and accurate as pos¬
sible.
COELENTERATA
Subkingdom Porifera
Order Hexactinellidae
Sponges are polycellular organisms beginning in the Cam¬
brian and continuing until the present time. They are sessile
Ockerman — Fauna of Galena Limestone Near Appleton 103
animals and are practically all marine, although there are a few
types of fresh water sponges. They are remarkable for the
variety of size and form, varying from the size of a pinhead to
one meter. Sponges are usually poorly preserved and for this
reason they have not been very thoroughly worked, Defrance
in 1827 discussed the Keceptaculitidae in the Dictionnaire de
Science Naturelles t. 45, p. 5 atlas, pi. 68 and in this country
Ferd Roemer discussed the sponges in 1860 (Silurisehe Fa^^na
des Westlichen Tennessee.)
The sponges are now classified under two heads ; Calcarea, and
Non-Calcarea. Formerly they were classified as Silicispongide
and Calcispongiae. The Hexactinellidae belong to the Non-
Calcarea group and they possess six-spined spicules. These
spicules are composed of colloidal silica, the silica being in alter¬
nate layers with the organic matter. There are two other types
of spicules as to composition, those built up of calcareous mate¬
rial and those composed of a horny fiber. The siliceous sponges
are the best preserved as fossils and in the Galena of this region
the writer has found four types of sponges and these are all
species of the Hexactinellidae.
These sponges are composed of calcareous material due to the
replacement of the colloidal silica of which the spicules are com¬
posed. The replacement process works both ways and often the
calcareous spicules are silicified.
The siliceous sponges are found as fossils in the Cambrian
but are greatest in the Jurassic and Cretaceous times.
The writer has found specimens of Receptaculites oweni in the
Neenah, Kaukauna and Mackville quarries. They are fairly
abundant especially at Mackville, and in practically all eases are
in a fair state of preservation. Ischadites iowensis is not as
abundant as the Receptaculites oiveni but nearly so. In a few
of the specimens the preservation was good, showing the exter¬
nal structure plainly. Receptaculites occidentalis and Hindia
parva are both very rare and poorly preserved and their identi¬
fication is rather uncertain.
Receptaculites oweni Hall.
Plate I, figure 1.
Receptaculites oweni Whitfield. Geol. Wisconsin 4, 1882, p.
239, pi. 10.
104 Wisconsin Academy of. Sciences, Arts, and Letters.
Receptaculites oweni Winchell and Schuchert. Geol. Minne¬
sota, 3,1893, p. 57, pi. F, fig. 1-4.
Description. The body of the organism is thin centrally but
widens as it approaches the margin. It is built of cell-like per¬
forations perpendicular to the plane of the disk, arranged in
concentric rows. The specimens vary from one-fourth inch to
two and one-half inches in diameter and one-sixteenth to one-
half inch in thickness.
This sponge is found at Neenah, Mackville, and Kaukauna
and is usually in a fair state of preservation. Because of the
calcareous composition these fossils are often badly weathered
but still preserve the perforations as slight indentations.
Ischadites iowensis (Owen)
Plate I, figure 2.
Receptaculites iowensis Winchell and Schuchert. Geol. Minne¬
sota, 3, 1893, p. 64, pi. F, fig. 5-6.
R. (Ischadites) iowensis GraBau and Shimer. N. A. Index Fos¬
sils, 1, 1906, p. 19, fig. 30.
Description. The base is concave while the upper surface is
flatly domed with the center a hollow cavity. It is built up of
small rhombohedral cells which are arranged in curved rows.
The specimens that the writer studied vary from one to two
inches in diameter and about one-half inch in thickness.
This species is found fairly abundant at the Mackville quarry
but is rare at the Neenah and Kaukauna quarries. In all cases
the specimens were in a fair state of preservation.
Receptaculites occidentalis Salter.
Receptaculites neptuni Hall. Pal. New York, 1, 1847, p. 68, pi.
24, fig. 3a-d.
Receptaculites occidentalis Salter. Canadian Org. Rem. Geol.
Survey Canada. Dec. 1, 1859, p. 45, pi. 10, fig. 1-7.
Description. This species is either suborbicular or hemis¬
pherical, being depressed in the central surface and having a
series of quadrangular cells. The only specimen studied was
such a poor specimen that its identification is rather doubtful.
The section Avhich was found at the quarry near Menasha was
Ockerman — Fauna of Galena Limestone Near Appleton 105
about three inches in diameter and about one-half inch in thick¬
ness and depressed at the center.
Hindia parva Winchell and Schuchert.
Hindia parva Winchell and Schuchert. Greol. Minnesota, 3,
1893, p. 79, pL G.
Description. A tiny spheriodal, free sponge having an even
rounded surface which seemed to be marked by numerous tiny
apertures.
The few specimens were very poorly preserved and measured'
about one quarter inch in diameter.
COELENTERATA
Subkingdom Cnidaria
Order Anthozoa
The anthozoa or corals are marine organisms, a type of sessile
poh^ps, always living in tropical seas and usually in shallow
water, their customary limitation being about 50 fathoms.
However, there are some found in water as deep as 1,500
fathoms.
Very few corals have permanently soft bodies, the majority
having a calcareous or horny skeleton or structure, the corallum.
This corallum is either simple or compound.
The number of internal partitions, the septa, vary and are
important both in number and arrangement in the study and
identification of these fossils. This arrangement will be dis¬
cussed more fully in conjunction with the classification. In
some cases the septa are twisted at the center forming a colum¬
ella. In some cases the septal center is very low, being about
one -ha If of the distance between the top and bottom of the coral,
while in other cases the septal center extends even above the top
of the coral.
The outer wall or theca varies considerably and is marked in
some cases by concentric lines while in others the lines are longi¬
tudinal. There are a few species with a smooth surface.
The corals began in the Cambrian and continued on through
to the recent. Practically all the Paleozoic forms are extinct
106 ^Wisconsm Academy of Sciences, Arts, and Letters.
and those of the Cenozoic and Mesozoic were similar to living
forms.
The corals are classified under four heads which are as fol¬
lows ; Tetracoralla, Hexacoralla, Octocoralla, and Tabulata.
The Tetracoralla has its septa arranged in a tetrameral sys¬
tem, all the septa being arranged around the four primary
ones. This group appeared in the Ordovician and had its maxi¬
mum development in the Silurian, becoming extinct at the end
of the Paleozoic.
The Hexacoralla has, as the name indicates, six primary septa
around which the remaining septa are developed. In some
cases the coral has twelve primary septa instead of six. This
group appeared in the Cambrian and continued on through the
Paleozoic having its maximum development in the Trias.
The Octocoralla is a rather small group which made its ap¬
pearance in the Ordovician. As the name suggests, this group
has its septa arranged in an octameral order.
The Tabulata has often been considered as a sub-order of the
Hexacoralla but now it is generally considered as a group in
itself. It was originally given this name because of the great
number of tabulae in corals of this type. Its range is from the
Ordovician through the Carboniferous.
The calcareous or horny structure of the corals is very adapta¬
ble to fossilization and practically all the specimens the writer
has studied were well preserved. Often the horny skeleton is
replaced Avhen it is fossilized. The Tetracoralla were very
abundant in the Ordovician and Silurian and are characteristic
of these two ages. The Hexacoralla are characteristic of the
Trias while the Tabulata are markers of the Silurian.
The corals were found very abundantly in this region at all
the quarries. The Tetracoralla were found most abundantly
and one specimen of the Tabulata was found. Three species of
Strep telasma were found, namely, Streptelasma profundum,
Streptelasma corniculum, Streptelasma corniculum var. parvula.
Produndum was the most abundant and best preserved.
Corniculum was nearly as abundant and the variety, parvula,
is rather rare but well preserved.
The one species of Tabulata, LicJienaria typa, is very rare and
only a few poorly preserved specimens were found at Neenah.
Ockerman — Fauna of Galena Limestone Near Appleton 107
Streptelasma Hall
Streptelasma Hall. Pal. New York, 1, 1847, p. 17.
Streptelasma Grabau and Sliimer. N. A. Index Fossils. 1, 1906,
p. 54.
Description. The Streptelasma are simple conical corals be¬
longing to the Tetracoralla. These corals are usually curved
and have a funnel-shaped calyx. The calyx varies in depth in
different species, some being quite deep while others are numer¬
ous. In a few cases they are twisted at the center forming a
columella.
Streptelasma profundum (Owen)
Plate I, figure 5.
Streptelasma profundum Hall. Pal. New York, 1, 1847, p. 49,
pi. 12, fig. 4a-d.
Streptelasma profundum Grabau and Shimer. N. A. Index
Fossils, 1, 1906, p. 54, fig. 76a-b.
Description. This coral is often slightly curved at the base,
expanding more or less abruptly upward. The calyx is usually
deep, this fact serving as the chief means of separation from
S. comiculum. This species usually has a small number of
septa, ranging from 36 to 60 as a general rule. The septa are
never twisted at the center in profundum as they are in corni-
culum.
This species of Streptelasma proved to be the most abundant
in this locality and many well preserved specimens were found
at Mackville, Neenah, Kaukauna, and Menasha. They vary in
size from three quarters to one and one-half inches in length.
Streptelasma comiculum Hall.
Plate I, figure 4.
Streptelasma comiculum Hall. Pal. New York, 1, 1847, p. 69,
pi. 25, fig. la-e.
Streptelasma comiculum Grabau and Shimer. N. A. Index
Fossils, 1, 1906, p, 55, fig. 76c-d.
Description. This coral is turbinate, curved very decidedly
at the base. In all cases this species has a very shallow calyx
108 Wisconsin Academy of Sciences, Arts, and Letters.
and the septa are usually long and twisted at the center into a
columella.
These corals were in considerable abundance in the Neenah
and Mackville quarries and some were found in the Kaukauna
and Menasha quarries also. They vary from % to 2 inches in
length and in most cases were well preserved.
Streptelasma corniculum var. parvula Hall.
Streptelasma parvula Hall. Pal. New York, 1, 1847, 71, pi. 25,
fig. 4a-c.
Description. These are very small corals of the Streptelasma
ranging in size from 3/16 to I/4 inch. They are turbinate,
slightly curved and have a smooth surface in most cases.
Pound rarely at Neenah and Mackville. These were as a rule
not as well preserved as the others of the Streptelasma.
Lichenaria typa Winchell and Schuchert.
Plate I, figure 3.
Lichenaria typa Winchell and Schuchert. Geol. Minnesota 3,
1895, p. 83, pi. G, fig. 10-13.
Description. This is a colonial coral composed of small irre¬
gular hemispherical colonies. The corallum has thin walls
with round cells if free, otherwise they have a polygonal outline
due to the lateral crowding of the adjacent cells.
Only a few poorly preserved specimens were found at Neenah
but they were easily identified by their external structure. The
size of the corallites was about % inch on an average in diame¬
ter.
COELENTERATA
Subkingdom Hydrozoa
Order Campanularia Allan.
Suborder. Graptolitoidea Lapworth.
At the close of the Cambrian there appeared in the seas a
group of colonial organisms which developed extensively in the
Ordovician and became extinct at the close of that period.
These organisms resemble in their fossilization Greek writing
upon rock and it is because of this that they were given the
Ockerman — Fauna of Galena Limestone Near Appleton 109
name of graptolites by Linnaeus in 1736 from graptolithus
Greek writing. These organisms are very unusual in their
rapid development and their sudden extinction.
The graptolites formed floating colonies in shallow muddy
waters being free swimming animals in most cases, although some
seem to have been fastened to drifting seaweed or some object
to keep them up, but it can be generally said that they were free
swimming. That they were not attached to the bottom is proven
by the fact that they do not extend vertically through several
beds but are always confined to one bedding plane.
The graptolites are derived from succession of buddings from
a primal hydrotheca or cell, known as the sicula. The hydro¬
thecae in which the individual organisms lived, are arranged in
longitudinal rows on one or two sides of the stem or polypary,
thus producing the serrate edge so characteristic of the grapto¬
lites. The skeleton secreted by the organism is composed of car¬
bonized chitin and is preserved as a thin carbonaceous film.
It is only recently that the graptolites have been classified as
a single group. Because of the fact that they were the first
organisms to become developed and then extinct there has been
much confusion as to their real character and at first they were
confused with various groups, as plant remains, horny sponges,
etc. In 1843 Portlock classified them as being a group under
the Campanularia.
As an individual group the graptolites are classified into two
groups, according to the arrangement of the hydrotheca; those
with the hydrotheca along one side of the polypary are the
Monoprionidae and those with the hydrotheca along both sides
are the Diprionidae.
The carbonized chitin is very adaptable to fossilization and
it is because of this that we have the graptolites so well pre¬
served. In practically all cases they are found flattened in bed¬
ding planes and associated in groups. They are found in prac¬
tically all kinds of rocks of sedimentary origin, being most
abundant and best preserved in fine grained carbonaceous
shale. The writer has found his specimens in the bedding
planes of the Galena.
Lapworth states that the graptolites are distributed verticalh^
through six horizons and only six. The first is the Upper Cam¬
brian, the second, third and fourth are identical with the Ordo¬
vician, and the fifth and sixth are Silurian. It is because of
110 Wisconsin Academy of Sciences, Arts, and Letters.
this limitation that they are so important as horizon markers.
They reached their greatest development in the Ordovician and
were so abundant that the Ordovician is called the “Age of
graptolites. ’ ’
The writer has found specimens at the quarries of Kaukauna,
Neenah and Kimberly. All belonged to the class Diprionidae.
Climacograptus typicalis was the most abundant and best pre¬
served. Climacograptus hicornis was rather rare but well pre¬
served. Both Diplograptus foliaceus and the variety, acutus,
were \ery rare, and Lasiograptus mucronatus also was very
rare.
Climacograptus typicalis Hall.
Climacograptus typicalis Winchell and Schuchert. Geol. Minne¬
sota, 3, 1895, p. 27, pi. A, fig. 1-9.
Climacograptus typicalis Grabau and Shimer. N. A. Index
Fossils, 1, 1906, p. 32, fig. 5 lb.
Description. The hydrothecae are parallel to the axis and
have their outer margin also parallel to the axis. This species
varies from bicronis in that the lower edge of the hydrotheca has
a slight lip-like extension and the serratures are more horizon¬
tal. Often old specimens of typicals have these points broken
off and resemble hicornis.
This species was found abundantly both at Kaukauna and
Kimberly and many of the specimens were in a good state of
preservation. The number of hydrotheca varies from 40-50 to
the inch. The cross section shows an oval shape about 1 mm. in
narrowest diameter.
Climacograptus hicornis Hall.
Oraptolithus hicornis Hall, Pal. New York, 1, 1847, p. 268, pL
73, fig. 2.
Climacograptus hicornis Grabau and Shimer. N. A. Index Fos¬
sils, 1, 1906, p. 32, fig. 1, a-c.
Description. Hydrothecae parallel, their outer margin being
straight and parallel to the axis of the stem. The hydrotheeae
are rectangular and the serratures are slightly oblique, extend¬
ing about half way to the axis. There is a gradual widening
upward and the base has two diverging forks.
Ockerman — Fauna of Galena Limestone Near Appleton 111
This species was found at Kaukauna and Kimberly but was
rare; in many cases it consisted of small fragments only fairly
preserved. It is about the same size as typicalis and has about
the same number of hydrothecae to the inch.
Diplograptus foliaceus (Murchison.)
Graptolitlius pristis (Part) Hall Pal. New York, 1, 1847, p. 265,
pi. 72, fig. la-e.
Diplograptus foliaceus Grabau and Shimer. N. A. Index Fos¬
sils, 1, 1906, p. 33.
Description. Linear, with two rows of alternating oblique
hydrothecae. This species differs from the variety acutus in
that the hyrothecae are larger and the serratures are wider.
The hydrothecae are less oblique in this species.
This species was found at Neenah and was in such small, poorly
preserved fragments that its identification is rather uncertain.
Diplograptus foliaceus acutus Lapworth.
Graptolitlius pristis Hall. Pal. New York, 1, 1847, p. 265, pi. 73,
fig. Ip-d.
Description. This species has the hydrothecag ari’anged as
Diplograptus foliaceus but it is distinguished by having moi’e
and more oblique hydrotheca;. Only one poorly preserved speci-
ment was found either at Kaukauna or Kimberly and its identi¬
fication was difficult.
Lasiograptus mucronatus Hall.
Graptolitlius mucronatus Hall. Pal. New York, 1, 1847, p.
268, pi. 73, fig. la-d.
Description, This species has the hydrotheca arranged simi¬
lar to Diplograptus foliaceus but is distinguished by the mucro-
nate points on the hydrotheca. These points are about one-half
the length of the hydrotheca.
This too is rare and only one poor specimen was found at
Kimberly. It was so small that its character could not be
studied.
112 Wisconsin Academy of Sciences^ Arts, and Letters.
E CHINOI>ERM AT A
Subkingdom Crinoidea
The crinoids were a group of marine invertebrates that lived
in colonies and were usually local in distribution. They re¬
sembled a lily in complete form, having usually a long stalk and
a more or less globular calyx. In a few instances they were
non-pedunculate and sessile. They inhabited shallow waters
as a rule but are known to exist to a depth of 3,000 fathoms.
They seemed to be gregarious in habit as is indicated by the fact
that their skeletons constitute the major part of great masses of
limestone (crinoidal limestone), especially in the Devonian and
Mississippian periods.
These organisms began in the Ordovician and have continued
to the present, but they were most abundant in the upper Paleo¬
zoic strata.
The first attempt at classification was made by Miller in 1821.
At present the classification is as follows: Larviformia, Fis-
tulata, Camerata, Flexibila, and Articulata.
The organism was built up of three parts, the calyx, the arms,
and the stalk. The calyx, usually having the form of a cup-
shaped or globular capsule, was made up of regularly arranged
plates. These plates and their arrangement are very important
in the study of crinoids. The arms were attached to the plates
but as these could not be preserved as fossils they are not of
great importance.
The stalk is very long in some cases, xevy short in others, and
in rare cases when the crinoid is non-pedunculate the stalk is en¬
tirely lacking, the calyx being fastened directly to the base. The
stalk is built up of segments of various thickness, size and shape
according to the species. Some are circular in cross-section
while others are elliptical or angular, the pentagonal form being
common. Some have rounded edges some are marked by fine
lines and these variations serve as a means of identification of
the various species of Crinoidea. The stem segments are very
important because they are the parts of the crinoids that are
usually found as fossils.
The crinoids are unfavorable for preservation in a perfect and
complete form because of the delicate structure and the loose¬
ness with which the plates and segments are fastened together.
Ockennan — Fauna of Galena Limestone Near Appleto^i 113
The perfect calyx is rare but often the individual plates are
found. However, the chief fossil remains of the Crinoidea are
the stalk fragments.
The writer has found only a few fragmental stalks in his
search through the quarries of this vicinity and often these were
very small, some consisting of but a single stem segment.
Four species of Crinoidea have been found in this region.
Schizocrinus nodosus, Dendrocriniis alternatus, Heterocrinus
heterodactylus, and Heterocrinus pentagonus.
ScMzocranus nodosus is very abundant in all the quarries and
usually the stem fragments were well preserved. Dendrocrinus
alternatus is rare and only a few small specimens were found at
Neenah. Heterocrinus heterodactylus is also rare and poorly
preserved, being found at Mackville. Heterocrinus penta¬
gonus is very rare and only one segment was found at Neenah.
It is well preserved and the external and internal structure of
the stem is easily studied.
Schizocrinus nodosus Hall.
Plate I, figure 7.
Schizocrinus nodosus Hall. Pal. New York, 1, 1847, p. 81, pi.
27, fig. la-b.
Schizocrinus nodosus Emmons. Amer. Geol. 1, 1855, p. 224.
Description. The body or calyx is cup shaped and obtusely
pentagonal. The column is round, composed of joints of un¬
equal thickness and diameter. The segments are well marked
with fine concentric lines and the cross section shows numerous
radial lines.
This species is very abundant in all the quarries and as a
general rule is well preserved. In some cases of extreme weath¬
ering the central part of the stem is gone, but usually even the
linear markings of the segments are visible. The specimens
range from % to % inch in diameter and are of different
lengths, from a single segment to a fragment nearly 3 inches
long.
Dendrocrinus alternatus (Hall).
Plate I, figure 9.
Poteriocrinus alternatus Hall. Pal. New York, 1, 1847, p. 83,
pi. 28, fig. If.
114 Wisconsin Academy of Sciences, Arts, and Letters.
Description. The column is very slender and is composed of
thickened plates with rounded edges which alternate in size.
The segments are very uniform in this alternation.
This species is found only rarely in this region and only two
poorly preserved specimens were found at Neenah. The diame¬
ter of the stem is about 1/16 inch and one specimen is about 2
inches long.
Heterocrinus lieterodactylus Hall.
Plate I, figure 8.
Heterocrinus lieterodactylus Hall. Pal. New York, 1, 1847, p.
279, pi. 76, fig. la-0.
Description. The body is short and nearly round. The col¬
umn is built up of thick pentagonal plates which are all nearly
the same size with a slight rounding at the edges. The plates
are usually divided by five partitions and the column is usually
rather slender.
This species is very rare in this region and only two small,
poor specimens were found at Mackville. They range from %
to 3/16 inch in diameter and are about % inch long.
Heterocrinus pentagonus Hall.
Plate I, figure 6.
Description. The diameter of this species is about 3/16 inch
and the plate is about 1/16 inch thick, rounded at the edges.
There are tiny radial markings at the edge. The central open¬
ing is pentagonal, its sides corresponding to the outer sides, the
stem being a pentagonal one. There are five small penagonal
openings located midway between the external faces and those
of the central openings.
Only one segment of this species was found at Neenah and
the writer has called it pentagonus although he has not had ac¬
cess to any description of pentagonus ; the pentagonal struc¬
ture is so pronounced, however, that it undoubtedly belongs to
this species.
Ockerman — Fauna of Galena Limestone Near Appleton 115
Molluscoidea
Subkingdom Bryozoa
The Bryozoa appeared in the Lower Oj'dovician and were
usually colonial and encrusting animals. They are chiefly ma¬
rine but in some cases they are fresh water. The name arose out
of their moss like appearance, coming from the G-reek, bryoii
moss and zoon animal.
The body wall of each individual animal was made hard by
means of calcerous or horny material and forms the only part
of the animal capable of being preserved in a fossil state.
The small animals formed variously shaped colonies, some
being moss like tufts, some in branching stems, and usually
encrusting shells and other foreign material.
The classification that is generally accepted is as follows:
Ctenostomata, Cyclostomata, Trepostomata, Cryptosotamata,
and Chilostomata.
The writer has found the Bryozoa to be in considerable abund¬
ance in this region and as a general rule the specimens are well
preserved as far as external structure is concerned. Because of
the lack of sufficient material on the Bryozoa the writer has not
gone into a detailed discussion of this group and has been lim¬
ited in the identification of the various species.
Several types have been found at the various quarries in this
vicinity and they are generally well preserved. Rliomhotrypa
quadrata, Lioclemella soUdissima-f Escharopora suhrecta^ Callo-
porella lens, Stictoporella sp. and Rhinddictya mutabilis are the
species that have been identified in this region.
Rhomb otry pa quadrat a is fairly abundant and very charac¬
teristic of the Ordovician of this region and is found at all the
quarries. Lioclemella solidissima is slightlj^ more abundant than
Rhomb otry pa quadrat a, but is not as well preserved. Only one
specimen of Escharopora subrecta was found, but it was in a
good state of preservation.
Calloporella lens was fairly abundant in the quarries but was
usually poorly preserved. Only a few small speci¬
mens of Stictoporella sp. were found and they, too,
were poorly preserved. Rhinidictya mnioMlis was
rare, but good specimens were obtained.
116 Wisconsin Academy of Sciences, Arts, and Letters.
Rhomhotrya quadrata (Rominger).
Plate I, figure 10.
Monticulipora multituherculata Whitfield, Geol. Survey Wis¬
consin, 4, 1882, p. 250, pi. 11, fig. 9-10.
Monotrypella quadrata. Grabau and Shimer. N. A. Index
Fossils, 1, 1906, p. 131.
Description. A colonial form, being a flat expansion of up¬
right parallel tubes of calcareous material. The tubes are thick
walled, the walls being non-porous. The openings of the tubes
are polygonal or round. The surface is made irregular by the
elongation of groups of individual tubes. This gives the sur¬
face a very characteristic roughness. The stems are usually
strong and more or less flattened.
This species is fairly abundant at Mackville but more rarely
in the other quarries. Some of the specimens were over % inch
in width and as long as 2 inches.
Lioclemella solidissima (Whitfield).
Plate I, figure 11.
Fistulipora solidissima (Whitfield). Geol. Survey. Wisconsin,
4, 1882, p. 255, pi. 11, fig. 18, 19.
Description. A bryozoan forming strong cylindrical branches
which often attain a diameter of about % of an inch. The sur¬
face is usually covered with minute elongate-oval or sometimes
rounded cells.
This species is found abundantly in all the quarries of this
vicinity and is fairly preserved. The specimens vary in size and
range up to % inch in width.
Escharopora subrecta Ulrich.
Plate I, figure 12.
Escharopora subrecta Ulrich. Geol. Minnesota, 3, 1893, p. 168,
pi. 12, figure. 5-29.
Escharopora subrecta Grabau and Shimer. N. A. Index Fossils,
1, 1906, p. 156, fig. 208f.
Description. A simple, flattened, straight or curved sub-
cylindrical bryozoan. The cell openings are elongate-elliptical.
Ockerman — Fauna of Galena Limestone Near Appleton 117
sometimes almost acute at the ends. The cells have a very pro¬
nounced longitudinal arrangement although in some cases the
diagonal rows are as regular and as evident as the longitudinal
ones.
Only one specimen was found by the writer and it showed the
external structure very plainly. The specimen is about one inch
long and 3/16th inch wide. The one specimen was found at
Neenah.
Calloporella f lens Whitefield.
Fistulipora lens Whitfield. GeoL Survey Wisconsin, 4, 1882, p.
256, pL 11, fig. 5, 6.
Description, A bryozoan growing in small discoidal or plano¬
convex button-shaped bodies. The under surface is more or less
concave and is marked by fine radiating striae. The upper sur¬
face is covered with round or polygonal apertures, the partition
walls being thin.
This species is abundant at Neenah and Mackville, but the
specimens are poorly preserved. The specimens vary in size
from % to 1% inches in diameter.
Stictoporella sp. Ulrich.
Very rare in this region and only a few poor specimens were
found; the writer is not certain as to their identification. The
small size seemed characteristic, as the specimens were only 1/16
inch in diameter.
Rhinidictya mutahilis Ulrich.
Rhinidictya mutahilis Ulrich. GeoL Minnesota, 3, 1893, p. 125,
pL 6, fig. 1-6.
Rhinidictya mutahilis Grabau and Shimer. N. A. Index Fossils,
1, 1906, p. 158, fig. 2081
Description. Branches commonly small, being usually 2-3
mm. in diameter. There are usually 14 or 15 rows of cells which
are arranged longitudinally.
Fairly abundant in this region at all the quarries ; the cell ar¬
rangement was in evidence in practically all specimens.
118 Wisconsin Academy of Sciences, Arts, and Letters.
Brachiopoda
The brachiopods are marine organisms that appeared in the
Lower Cambrian. They secreted a shell made np of two parts,
the pedicle valve and the brachial valve. They are most prolific
in warm waters, and are usually found in shallow waters al¬
though they have been found in water as deep as 3000 fathoms.
These organisms are gregarious in habit and are often attached
to one another. As a general rule the brachiopods are attached
to some foreign material by a fleshy stalk, which is known as the
pedicle. It is only in the larval stage that they are free floating
and it is in this period that they are distributed.
The brachiopods attained their maximum development in the
Silurian and Devonian times. They are very numerous and
there are over six thousand fossils species of this group and
about 140 recent species.
In 1792 Cuvier distinguished the Brachiopoda from the
Acephala and put them as a fourth family in his class of
Mollusca. It was Dumeril, in 1806, that gave them the name of
Brachiopoda, meaning arm-footed animals. This name has been
changed often but has not been supplanted.
The Brachiopoda are divided into two large groups according
to the presence and absence of teeth. The Inarticiilata have no
teeth the valves being held together by muscles. The Articulata
have the valves articulated by teeth and sockets. These two
groups are subdivided on the nature of the pedicle opening and
the stages of shell growth into the Atremata and Neotremata of
the Inarticulata and the Protremata and Telotremata of the
Articulata.
The two valves are eqilateral, but not equivalved, and vary
greatly as to shape. In some cases both valves are convex, in
others they are nearly flat and again some have a convex pedicle
valve and a concave brachial valve. The pedicle valve derived its
name from the fact that the pedicle came from that valve. It is
usually the larger of the two valves. These valves are composed
of calcareous or phosphatic material or both.
The external markings are of two kinds, concentric lines of
growth and plications radiating from the region of the beak.
Often when the plication are small they are referred to as
striations. These vary greatly, some being simple, some bifid,
some trifid, and some alternating in size with the others. Pre-
Ockerman — Fauna of Galena Limestone Near Appleton 119
quently there is a larger fold or plication on the brachial valve,
known as the median fold, and a median depression on the
pedicle valve, known as the medial sinus.
The writer has found the brachiopods in great abundance in
this region and many of them well preserved.
Lingula Bruguiere.
Lingula Winchell and Schuchert. Geol. Minnesota, 3, 1893, p.
338.
Lingula Grabau and Shimer. N. A. Index Fossils, 1, 1907, p. 194.
Description. Elongate-ovate, equivalved, with a thin glisten¬
ing shell. The shell is usually smooth or with fine concentric
lines and rarely with a fine striae. Broad anteriorly, parallel
sided, or tapering posteriorly.
Lingula Jiulhurti Winchell.
Plate II, figure 2.
Lingula hulhurti Winchell and Schuchert. Geo. Minnesota, 3,
1893, p. 347, pi. 29, fig. 13-14.
Description. Shell ovate, pointed posteriorly, being broadest
in the anterior half. The convexity is moderate and regular.
The exterior is marked by sharply elevated concentric plications
which are fewer near the beak.
Found rarely in this region, only one specimen being found
at Neenah. This was well preserved and the size of it was 7/16
inch long and 5/16 inch wide.
Lingula ohtusa Hall.
Plate II, figure 4.
Lingula ohtusa Hall. Pal. New York, 1, 1847, p. 98, pi. 30
fig. 7.
Description. Broad, ovate, obtuse at the apex and broadly
rounded below. Depressed in front, with a prominent but not
terminal beak. Fine concentric lines and radiating striae mark
the surface.
This species is rare in this vicinity and only one specimen was
found at Neenah. It was 5/16 inch at the greatest width.
120 Wisconsin Academy of Sciences, Arts, and Letters.
Lingula elderi Whitfield.
Plate II, figure 1.
Lingula elderi Whitfield. Geol. Survey Wisconsin, 4, 1882, p.
345, pi. 27, fig. 1-5.
Lingula elderi Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 196,
Description. An oblong, sub-quadrangular shell, the margins
being nearly parallel. The valves are slightly convex, having
nearly a smooth surface, which in some cases is marked by fine
concentric lines.
Only one specimen was found. It was well preserved and is
% inch wide and % inch long.
Lingula iowensis (Owen).
Plate II, figure 3.
Lingual ioiHensis Winchell and Schuchert. Geol. Minnesota, 3,
1893, p. 349, pi. fig. 19-22.
Lingulella iowensis Whitfield. Geol. Survey Wisconsin, 4, 1882,
p. 242, pi. 9, fig. 1.
Descnption. A large shell, broadly ovate, subquadrate in out¬
line. The sides are gently rounded and the bottom is moderately
rounded. The valves are convex, the pedicle valve being more
convex. The surface is marked by strong irregular lines of
growth and fine flattened angular striae.
This species is rare in this region and only one well preserved
specimen was found at Neenah. It was % inch wide and % inch
long.
Rafinesquina Hall and Clarke.
Bafinesqwina Winchell and Schuchert. Geol. Minnesota, 3, 1893,
p. 400.
Refinesquina Grabau and Shimer. N. A. Index Fossils. 1,
1907, p. 211.
Description. The shells are normally concavo-convex, the ped¬
icle valve being convex and the brachial valve being concave.
The shell is semi-oval, with a straight hinge line. The surface is
OckermM — Faunu of Galena Limestone Near Appleton 121
marked by radiating striae of alternating size crossed by finer
concentric lines.
The Rafinesqiiina began in the Calciferons and died out in the
Clinton.
Rafinesquina alternata (Emmons).
Plate II, figure 5.
Leptaena alternata Hall. PI. New York, 1, 1847, p. 102, pi. 31,
fig. 1.
Rafinesquina alternata Winchell and Schuchert. Geol. Minne¬
sota, 3, 1893, p. 404, pi. 31, fig. 32-34.
Rafinesquina alternata Grabau and Shimer. N. A. Index Fos¬
sils, 1, 1907, p. 211, fig. 251.
Description. The shell is large, the breadth being greater than
the length. Broadly semi-oval, the lateral margins being slightly
convex, rounding forward to a front. The pedicle valve is me¬
dium convex and is flattered near the cardinal extremities. The
brachial valve is gently concave in central and anterior regions
and has a small beak. The surface of both valves is ornamented
with numerous radiating striae which alternate very noticeably.
Fine concentric lines cross the striae.
Rafinesquina minnesotaensis inquassa.
Rafinesquina minnesotaensis Grabau and Shimer. 1, 1907, p.
212, fig. 252.
Description. Very similar to Rafinesquina alternata in form
and surface features but usually is distinguishable by the smaller
size and the greater convexity of minnesotaensis. Sometimes it
is considered as a small variety of Rafinesqina alternata.
Found rarely in the region under discussion. One specimen
measuring 1 inch across was found at Neenah. Fairly well
preserved.
Strophomena (King).
Strophomena Winchell and Schuchert. Geol. Minnesota, 3,
1893, p. 384.
Strophomena Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 222.
122 Wisconsin Academy of Sciences, Arts, and Letters.
Description. The shells are transversely sub-semicircnlar, the
greatest width being at the hingeline. The shell is concavo-
convex, being similar to Kafinesquina but with the convexity of
the valves reversed. The surface is marked by fine radiating
striae which are equal or alternating in size.
Strophomena planumhona (Hall).
Plate II, figure 6.
Strophomena planumhona Hall. Pal. New York, 1, 1847, p. 1 12,
pi. 31, fig. 4.
Strophomena rngosa Winchell and Schuchert. Geol. Minnesota,
3, 1893, p. 390, pi. 31, fig. 4-5.
Description. A medium sized shell which is wide at the hinge
line, the brachial valve being evenly convex, the pedicle valve
being quite concave. The surface is marked by fine radiating
striae, usually with several smaller ones between two larger ones,
the smaller ones being shorter. These are crossed by a number of
fine concentric lines and a few imbricating lines of growth at the
free margins.
Pound rarely in this region but fairly well preserved. One
specimen found at Neenah is slightly over 1 inch at the hinge
line.
Strophomena emanciata Winchell and Schuchert.
Plate II, figure 7.
Strophomena emanciata Winchell and Schuchert. Geol. Min¬
nesota, 3, 1893, p. 399, pi. 31, fig. 22-4.
Description. A small shell, depressed, bi-convex, semi-circu¬
lar in outline, the hinge line being slightly longer than the width
of the shell. The pedicle valve is depressed, convex, with a shal¬
low sinus that has its origin near the beak. Numerous angulated
striae. Rare in this region. Found at Neenah. Good specimen,
% inch at hinge line.
Plectambonites Pander.
Plectamhonites Winchell and Schuchert. 3, 1893, p. 413.
Ockerman — Fauna of Galena Limeston>e Near Appleton 12^^
Plectamhonites Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 225.
Description. A very small shell, the pedicle valve being con¬
vex and the brachial valve concave. The hinge line is the great¬
est width of the shell. The surface markings are fine striae
which often alternate in size.
Plectamhonites sericeus (Sowerby).
Plate II, figure 8.
Leptaena sericea Hall. Pal. New York, 1, 1847, p. 110, pi. 31B,
fig. 2.
Plectamhonites sericea Winchell and Schuchert. Geol. Minne¬
sota, 3, 1893, p. 414, pi. 32, fig. 10-12.
Plectamhonites sericea Grabau and Shimer. N. A. Index Fos¬
sils, 1, 1907, p. 227, fig. 274a-b.
Description. The shell is small, semi-oval, approaching semi¬
circular. The brachial valve is concave, the pericle valve is
moderately convex, nearly even in most cases. The surface is
often shiny and is marked by numerous minute, closely arranged,
equal striae, every fourth, fifth or sixth one being more prom¬
inent.
Found abundantly in this region and generally well preserved.
They range from % to % inch in width at the hinge line.
Orthis Dalman.
Ortliis Winchell and Schuchert. Geol. Minnesota, 3, 1893, p.
417.
Orthis Grabau and Shimer. N. A. Index Fossils, 1, 1907, p. 250.
Description. The brachial valve is very flat and the pedicle
valve is very convex, with an elevated and incurved cardinal
area. The surface is marked by comparatively strong, sharp
plications which are usually simple.
Orthis costalis. Hall.
Plate II, figure 9.
Orthis costalis Hall. Pal. New York, 1, 1847, p. 20, pi. 4Bis.
fig. 4a.
124 Wisconsin Academy of Sciences, Arts, and Letters.
Orthis Costalis Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 250, fig. 300a-5.
Description. The pedicle valve is very convex, while the
brachial valve is flat. The surface is marked by about 32 dis¬
tinct, strong, rounded striae which are continuous and enlarge
near the margin. This series resembles Dalmanella testudiniaria
but is more convex above and the striae are coarser and less
numerous.
It is found abundantly in this region and usually well pre¬
served. The specimens average about % inch along the hinge
line.
Plectorthis Hall and Clarke.
Fleet orthis Winchell and Schuchert. Geol. Minnesota, 3, 1893,
p. 435.
Plectorthis Grabau and Shimer. N’. A. Index Fossils, 1, 1907,
p. 251.
Description:. This differs from Orthis in that it has subequal
and convex valves, coarse plications and concentric lines crossing
the striae.
Plectorthis plica, tella (Hall).
Plate II, figure 10.
Orthis plicatella Hall. Pal New York, 1, 1847, p. 122, pi. 32,
fig. 19.
Plectorthis plicatella Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 252, fig. 301d-f.
Description. A broadly semi-oval shell, with both valves
slightly convex, their convexity being nearlj^ equal. The surface
is marked by strong radiating plications which are usually sim¬
ple. They range from 20 to 28 on each valve. These are crossed
by simple elevated concentric lines which are often obscure on
weathered surfaces.
This is found rarely at Mackville and the specimens range up
to ^ inch at the greatest width.
Plectorthis plictella (Hall).
Plate II, figure 11.
Orthis fissicosta Hall. Pal. New York, 1, 1847, pi. 121, 32 fig. 8.
Ockerman — Fauna of Galena Limestone. Near Appleton 125
Plectorthis fissicosta Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 252, fig. 300c-d.
Description. Semioval, the pedicle valve being moderately
convex, the beak extended and slightly incurved. The surface
is marked by angulated striae, numbering 19-20, which become
bifid and trifid towards the margin of the shell.
Found rarely in this vicinity in well perserved specimens.
Mackville was the only place that they were found. The speci¬
mens measured % inch across.
Plectorthis dichotoma Hall.
Plate II, figure 12.
OrtJiis dichotoma Hall. New York, 1, 1847, p. 125, pi. 32, fig. 10.
Description. A small suborbicular shell, with the valves nearly
equally convex, the pedicle valve being equally and uniformly
convex. There are about 26 well defined rounded striae which
bifurcate about half way to the margin.
This form is rare and poorly preserved; only one specimen
was found at Mackville which measured about % inch across.
Plectorthis sordida (Hall).
Plate II, figure 13.
Orthis sordida Hall. Pal. New York, 1, 1847, p. 148, pi. 33,
fig. 16.
Description. A small circular shell, the valves being equally
convex, the beak of the pedicle valve extending beyond the
brachial valve. The radii are coarse and simple. The pedicle
valve has a slight medial sinus.
This species is rare and poorly preserved; one specimen was
found at Mackville measuring % inch in diameter.
Dinorthis Hall and Clarke.
Dinorthis Winchell and Schuchert. Geol. Minnesota, 1, 1893, p.
420.
Dinorthis Grabau and Shimer. N. A. Index Fossils, 1, 1907, p.
252.
126 Wisconsin Academy of Sciences, Arts, and Letters.
Description. This genus varies from Orthis in that the
brachial valve is convex and the pedicle valve is elevated at the
umbo. It has strong simple, rarely bifurcating, striae.
Dinortkis meedsi Winchell and Schuchert.
Plate II, figure 14.
Dinortkis meedsi Winchell and Schuchert. Geol. Minnesota, 3,
1893, p. 427, pi. 32, fig. 39-42.
Dinortkis meedsi Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 253, fig. 303e-h.
Description. The shell is suborbicular with a straightening
along the hinge line. Pedicle valve is convex with a distinct but
not greatly elevated beak. The brachial valve is flat with a
slight elevation at the beak. The striae are coarse and simple,
the furrows being about the same width as the striae. This
species is similar to pectinella but has a shorter hinge line.
Pound rarely in this region, in a fair state of preservation.
The specimens found at Mackville measured from % to % i^^ch
across.
Dinortkis pectinella (Emmons).
Plate II, figure 15.
Dinortkis pectinella Winchell and Schuchert. Geol. Minnesota, 3,
1893, p. 424, pi. 32, fig. 10.
Dinortkis pectinella. Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 253, fig. 303i-l.
Description. Semi-oval, the Avidth and length being as 9 to 12.
The pedicle valve is regularly convex, the brachial valve is sub-
convex near the beak. Broad depression along the center.
There are 21-30 prominent, rounded striae which are bifid or
trifid towards the margin. Rare, poorly preserved, % inch in
diameter. MackAdlle.
Herbertella Hall and Clarke.
Herheriella Winchell and Schuchert. Geol. Minnesota, 3, 1893,
p. 432.
Herbertella Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 254.
Ockerman — Fauna of Galena Limestone Near Appleton 127
Description. The pedicle valve is depressed, convex, but less
convex than the brachial valve. The greatest width of the shell
is at the cardinal area. The surface is marked by numerous
closely crowded, fine rounded striae which are crossed by con¬
centric lines of growth.
Herhertella (Orthis) occidentalis Hall.
Plate III, figure 3.
Orthis occidentalis Hall. Pal. New York, 1, 1847, p. 127, pi. 32,
fig. 2.
Herhertella occidentalis Grabau and Shimer. N. A. Index
Fossils, 1, 1907, p. 225, fig. 304f-h.
Description. A semi-oval shell, the greatest width being at the
cardinal line. The brachial valve is strongly convex with a
slight depression along the center. The pedicle valve is convex
towards the beak. The brachial valve has a high incurved beak
which extends slightly beyond the hinge line. The striae are
elevated and sub-angular and are crossed by fine concentric lines
of growth.
This species is found rarely in this region and only one speci¬
men was found at Mackville. This was about % inch in width.
Herhertella (Orthis) hellirugosa (Conrad).
Plate II, figure 16.
Orthis hellirugosa Hall. Pal. New York, 1, 1847, p. 1-18, pi. 32,
fig. 3.
Herhertella hellirugosa Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 255, fig. 304d-^.
Herhertella hellirugosa Winchell and Sclmchert. Geoi Minne¬
sota, 3, 1893, p. 434, pi. 33, fig. 1-4.
Description. Semi-oval and biconvex, the valves are nearly
equal in convexity. The brachial valve has a conspicuous mesial
sinus and in some cases the pedicle valve has a sinus. The
pedicle valve is strongly convex, with the greatest elevation at
the umbo. The striae are prominent, linear, bifurcating near
the margin. There are numerous strongly imbricating concen¬
tric lines of growth.
128 Wisconsin Academy of Sciences, Arts, and Letters.
This is found rarely in this region, a few being found at
Maekville ; the specimens Vary from % to % inch in length.
Platystrophia King.
Platystrophia Winchell and Schuchert. Geol. Minnesota, 3, 1893,
p. 454.
Platystrophia Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 257.
Description. This shell has a spiriferoid hinge line which is
long and straight. Both valves are convex, the brachial valve being
more so. There is a strong medial fold on the brachial \alve,
and a strong sinus on the pedicle valve. The surface is marked
by strong sharp plications.
This type ranges from the Ordovician to the Silurian.
Platystrophia (Orthis) lynx (Eichwald)
Plate III. figure 1.
Dethyris lynx Hall. Pal. New York, 1, 1847, p. 133, pi. 32D, fig.
1.
Platystrophia lynx Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 258 fig. 308c-d.
Description. The shell is often nearly as thick as long, the
length and the width being as 5 to 9 or equal. The surface is
marked with strong angulated plications ; 3 or 4 mark the sinus
and 4 or 5 are on the elevated mesial lobe. Fine semi-imbricat¬
ing, flexuous lines cross the plications.
Found rarely in this vicinity. The specimen studied by the
writer measured % inch along the hinge line.
Platystrophia hiforata Schlothiem.
Plate III, figure 2.
Platystrophia hiforata Winchell and Schuchert. Geol. Minnesota,
3, 1893, p. 455, pi. 33, fig. 51-54.
Platystrophia hiforata. Grabau and Shimer. N. A. Index
Fossils, 1, 1907, p. 258.
Description. Transversely semi-elliptical. The pedicle valve
is stroiigly convex with a wide deep medial sinus which com-
Ockerman — Fauna of Galena Limestone Near Apyleton 129
mences at the extremity of the beak and widens to the front.
The surface is marked by a large number of radiating triangmlal*
ribs, 1 to 5 in the medial sinus. The valves are crossed by
numerous concentric raised lines.
This species is fairly abundant in this region, but the speci¬
mens are not well preserved. They vary in sie from ^/4 to 1
inch in width at the hinge line.
J* ,
Dalmanella Hall and Clarke.
Dalmanella Winchell and Schuchert. Geol. Minnesota, 3, 1893,
p. 439.
Dalmanella Grabau and Shimer. N. A. Index Fossils, 1, 1907, p.
194.
Description. Plano-convex or subequally convex, with a semi¬
circular outline. The pedicle valve has an elevated cardinal area.
The surface is marked by fine rounded bifurcating striae which
curve from the umbo to the sides of the valve. The hinge line is
generally shorter than the greatest width of the shell.
Dalmanella testudinaria (Dalman).
Plate III, figure 4.
Orthis testudinaria HaU. Pal. New York, 1, 1847, p. 117, pi. 32,
fig. 1.
Orthis testudinaria Whitfield. Geol. Survey Wisconsin, 4, 1882,
p. 258, pi. 12, fig. 5-7.
Dalmenella testudinaria Grabau and Shimer. N. A. Index Fossils,
p. 260, fig. 311a-e.
Description. Suborbicular, the pedicle valve being convex and
elevated towards the beak, with an elevated ridge down the
center. The brachial valve is flat. The surface is covered with
fine radiating striae which are crossed by fine concentric lines.
This species is found abundantly in this region in all the
quarries and in practically all cases the specimens were well
preserved. They vary in size, but are usually about % inch in
width, some being I/2 inch.
Rhynchotrema Hall.
Ehynchotrema Winchell and Schuchert. Geol. Minnesota, 3,
1893, p. 458.
9
130 WiscoifCsin Academy of Sciences, Arts, and Letters.
Rhynchotrema Grabau and Shinier, 1, 1907, p. 281.
Description. Restate and thick shelled. The pedicle has its
apex closely incurved over that of the brachial valve. The car¬
dinal process is very prominent. This is an extensive genus.
Rhynchotrema increhescens Hall.
Plate III, figure 5.
Atrypa increhescens Hall. Pal. New York, 1, 1847, p. 146, pi.
33, fig. 13a-h.
Rhynchotrema inaeqidvalvis Winchell and Schubert. Geol.
Minnesota, 3, 1893, p. 459, pi. 34, fig. 12-14.
Rhynchotrema inaeqidvalvis Grahau and Shinier. N. A. Index
Fossils, 1, 1907, p. 281, fig. 243a-d.
Description. A spheroidal or gibbous shell, the length and
the breadth being nearly equal. There is a broad deep sinus
in the brachial valve. There are 12-25 plications which are never
subdivided transversely. These are crossed by elevated imbricat¬
ing lines and delicate concentric zigzag lines.
Found in considerable abundance in this region, chiefly at
Mackville where they are weathered out. The specimens vary
from 1/4 to % in width and are well preserved.
Rhynchotrema capax (Conrad).
Plate III, figure 6.
Rhynchonella capax Whitfield. Geo. Survey Wisconsin, 4, 1882,
p. 263, pi. 12, fig. 26-27.
Rhychotrema capax Winchell and Schuchert, 3, 1893, p. 462, pi.
34, fig* 30, 34.
Rhynchotrema capax Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 282, fig. 343h-k.
Description. Medium sized, with both valves very convex, the
brachial valve being slightly the stronger. The medial sinus is
very pronounced, being deep and never reaching the* front of the
beak. There are always three simple angulated plications in the
bottom of the sinus. The lateral slopes are occupied by 5 to 7
simple plications. The entire surface of both valves is marked
by numerous regularly zigzag sublaminar lines of growth. Those
.are often obsolete in old or weathered specimens.
Ockerman — Fauna of Galena Limestone Near Appleton 131
This species is found abundantly in a well preserved state in all
of the quarries and the specimens vary from % to 1 inch in width.
Zygospira Hall.
Zygospira Winchell and Schuchert. Geol. Minnesota, 3, 1893, p.
465.
Zygospira Grabau and Shimer. N. A. Index Fossils, 1, 1907, p.
307.
Description. A very small shell, subcircular and biconvex.
There is a plicated medial fold on the pedicle valve and a medial
sinus on the brachial valve. The surface is sharply plicate.
Zygospira modest a Hall.
Atrypa modcsta Hall. Pal. New York, 1, 1847, p. 141, ])1. 33,
fig. 15,
Zygospira modest a Winchell and Schuchert. Geol. Minnesota, 3
1893, p. 467, pi. 34, fig. 42-44.
Zygospira modesta Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 308, fig. 375f-i.
Description. Suborbicular or planoconvex. The width is a lit¬
tle greater than the length. The shell is very small. The pedicle
valve has an elevated ridge along the center which is composed
of 4 plaits that are stronger than the others. The bealv is promi¬
nent and incurved. The brachial wave is less convex and has a
wide, ill-defined sinus. There are about 18-20 plications on each
valve.
This species is found rarely in this region, but the few speci¬
mens are well preserved. They are about 3/16 inch in width.
Mollusca
Subkingdom Gastropoda
The Gastropoda or snails are marine, fresh-water or terrestrial
mollusks. They are chiefly marine and usually inhabit shallow
waters, seldom being found below 500 fathoms, but in rare eases
exist at 2000 and even 3000 fathoms.
The shell is a secretion of the mantle and consists of carbonate
of lime in practically all eases. The shell is essentially a spirally
wound, elongated, conical tube, the coils or whorls being in most
132 Wisconsin Academy of Sciences, Arts, and Letters.
cases in contact and tightly cemented where they join. Some¬
times the shells are coiled in nearly the same plane and are dis-
coidal as in many of the Euomphalidae. More generally the
whorls are wound about an axis in an oblique manner, a true
spiral being formed. The last whorl is known as the body whorl
and the ones above are what constitute the spire. The line or
groove marking the junction of the whorls is known as the suture.
The shell is imperforate when the axis is solid as in Fusispira or is
perforate when the axis is hollow as in Trochonema, in which case
the axial cavity is called the umbilicus. The only external mark¬
ings are the lines of growth and these are usually distinguishable
except in cases where it is a cast of the shell that is found.
The form of the shell varies greatly but is very constant within
the limits of the species. Three types of shells are distinguished
in the large number of species. The tabular, straight, or only
slightly curved forms occur among the Scaphopoda and Petero-
poda. The symmetrical forms have the shell either conical, patel-
liform, or involute as in Bellerophon. The spiral is the typical
form of gastropod and the most common of the three types.
The fossil fauna of this region includes many types of the
Gastropoda; all of the quarries are fairly abundantly supplied
with casts of snails. Kaukauna and Mackville yielded the largest
number but Menasha had some very good specimens. Eighteen
species were found in the search of these quarries, all being casts
and usually poorly preserved.
Protowarthia Ulrich and Scofield.
Proiowarthia Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, 1897,
p. 848.
Protowarthia Grabau and Shimer. N'. A. Index Fossils, 1, 1907,
p. 611.
Description. Bellerophron shells with the aperture large, but
not abruptly expanded. The outer lip is bilobate, but never slit.
Umbilicus usually closed.
' ' Protowarthia pe7'voluta Ulrich and Scofield.
Protoivarthia pervohita Ulrich and Scofield. Geol. Minnesota,
3, pt. 2, 1897, p. 871, pi. 63, fig. 21-27.
Protowarthia pervoluta Grabau and Shimer. N. A. Index Fos¬
sils, 1, 1907, p. 611, fig. 817d-f.
Ockerman — Fauna of Galena Limestone Near Appleton 133
Description. Small, rather globose in form. Volutions uni¬
formly rounded. Lateral lobes of lips rounded. No surface
markings.
Found rarely at Maekville. Poor specimens. They measure
about % inch in diameter.
Raphistoma Hall.
EapJiistoma Ulrich and Scofield. Geo. Minnesota, 3, pt. 2,
1897, p. 931.
EapJiistoma Gabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 627.
Description. Shell sublenticular or plano-convex, the spire
fiat, and the sutures close. The volutions are triangular in sec¬
tion, sharply angulate. Aperture turned back abruptly.
EapJiistoma rotuloides (Hall).
Pleurotomaria rotuloides. Hall. Pal. New York, 1, 1847, p. 173,
pi. 37.
Description. The shell is depressed, conical, the spire being
composed of 4 volutions. The outer volution is angular at the
edge. Small umbilicus, subquadrate aperture.
Rare and poorly preserved. One from ^laekville, % inch wide.
Raphistomina Ulrich and Scofield.
EapJiistomina Ulrich and Scofield. Geol. Minnesota, 3, pt. 2,
1897, p. 932.
EapJiistomwa Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 629.
Description. Shell lenticular to depressed conical, umbilicated
volutions sharply angular and carinate at the periphery. Aper¬
ture subrhombical.
EapJiistomina modesta Ulrich.
Plate III, figure 11.
Raphistomina modesta Ulrich and Scofield. Geol. Minnesota,
3, pa. 2, 1897, p. 943, pi. 68, fig. 14-17.
134 Wisconsin Academy of Sciences, Arts, and Letters.
Description. Shell is small, with a discoidal shape. Nearly
always has a small umbilicus. The four volutions are gently
convex on the upper side. Sutures are not very deep. Very
short spire.
Cast found at Mackville, poorly preserved, about % inch in
diameter.
Lophospira Whitfield.
LopJiospira Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, 1897,
pi. 952.
Lophospira Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 631.
Description. Shells with more or less elevated spires, the
whorls being closely coiled throughout. Whorls are angular.
Umbilicus is nearly always present. Outer lip notched.
Lophospira angustina minnesotaensis Ulrich and Scofield.
L. angustina minnesotaensis. Ulrich and Scofield. Geol. Min¬
nesota, 3, pt.. 2, 1897, p. 988, pi. 71, fig. 3-4.
Description. About four whorls in the cast. These are strong¬
ly convex and angular above. Often confused with Hormotoma
major, but is shorter and less slender.
Found rarely at Kaukauna. The specimen measures about 2^4
inches at greatest diameter and is about 3 inches long.
Liospira Ulrich and Scofield.
Liospira Ulrich and Scofield. Geol. Minnesota, 3, pa. 2, 1897, p.
953.
Liospira Grabau and Shimer. N. A. Index Fossils, 1, 1907, p.
640.
Description. The shell is sublenticular, with a low spire, de¬
pressed conical, the sutures being scarcely distinguishable. The
volutions are subrhombical, being flat or gently above. The aper¬
ture is deeply notched and the umbilicus is usually filled by the
extension of the lip.
Ockerman — Fauna of Galena Limestone Near Appleton 135
Liospira micula (Hall).
Plate III, fi^re 7.
Liospira micula Ulrich and Scofield Geol. Minnesota, 3, pt. 2,
1897m p. 994, pi. 68, fig. 24-29.
Liospira micula Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 640, fig. 871a-d.
Description. A small shell with the whorls flattened and
continuous in the spire, the sutures being scarcely depressed.
There are about four volutions. The umbilicus is filled by reflex
callosity of the lip.
Fairly abundant at Mackville, but the casts were poorly pre¬
served. The specimens were about . % inch thick at greatest di¬
ameter.
Liospira progne (Billings).
Liospira progne Ulrich and Scofield. Geol. Minnesota, 3, pt. 2,
1897, p. 996, pi. 68, fig. 38-40.
Liospira progne Grabau and Shimer. N. A. Index Fossils, 1,
1907, pi. 640, fig. 871e-g.
Description. A small shell, composed of four volutions.
These volutions are flattened and the sutures are scarcely de¬
pressed. The umbilicus is fllled by reflex callosity of the lip.
This species is found rarely in this region, but is poorly pre¬
served. Those from Mackville measured % to 1 inch in diameter.
Plate III, Fig. 8.
Liospira vitruvia Ulrich and Scofield. Geol. Min¬
nesota, 3, pt. 2, 1897. p. 995, pi. 69, fig. 3-8.
Liospira vitruvia Grabau and Shimer. N. A. Index Fossils,
1, 1907, p. 641, fig. 871h-k.
Description. Small, with the spire depressed, the heighth
being not quite one-half the width. Scarcely four volutions,
these being flattened above. The umbilicus has an angular
margin.
This species is rarely well preserved, but a considerable num¬
ber of specimens were found at Mackville. The average width
was about % inch.
136 Wisconsin Academy of Sciences, Arts, and Letters.
Liospira ohtusa Ulrich and Scofield.
Plate III, figure 13.
Liospira ohtusa Ulrich and Scofield. Geol. Minnesota, 3, pt. 2,
1897, p. 997, pi. 68, fig. 30-34,
Description. Small, composed of four or five volutions which
are slightly convex above. Periphery rounded.
Very rare in this region. The only cast was found at Mack-
ville and measured % inch in width and was fairly well pre¬
served.
Clathospira Ulrich and Scofield.
Clathospira Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, 1897,
p. 954.
Clathospira Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 644.
Description. Shell depressed conical. Base more or less con¬
vex, Umbilicus very small or wanting. Few volutions which are
often strongly angular near mid-height,
Clathospira suhconica (Hall).
Pleurotomaria suhcowica Hall. Pal. New York, 1, 1847, p. 174.
pi. 37.
Clathospira suhconical Ulrich and Scofield. Geol. Minnesota, 3,
pt. 2, p. 1006, pi. 69, fig. 47-50.
Description. Short conical spire, the width and height being
nearly equal. Volutions art flattened above and distinctly
marked by a band.
Found very rarely in this region. One was found at Neenah,
but it was very poorly preserved. It measures % inch in diam¬
eter.
Hormotoma (Murchisonia) Salter,
Hormotoma Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, 1897,
p. 959,
Hormotoma Grabau and Shimer. N. A. Index Fossils, 1, 1907, p.
648.
Description. Shell elongate, composed of 8 to 14 rounded or
Ockerman — Fauna of Galena Limestone Near Appleton 137
subangular whorls. The outer lip has a deep broad notch. The
aperture is subovate, narrow and more or less prolonged below.
Hormotoma (Murchisonia) gracilis (Hall).
Murckisonia gracilis Hall. Pal. New York, 1, 1847, p. 181, pi,
39, fig. 4a.
Hormotoma gracilis. Ulrich and Scofield. Geol. Minnesota, 3,
pt. 2, 1897 p. 1015, pi, 70, fig. 18-21.
Description. A small slender shell, very elongate, composed
of numerous volutions which are usually rounded, but are some¬
times slightly angulated. The shell is loosely coiled. The
sutures are simple and deep.
The casts of this species are fairly abundant in this region and
a considerable number of specimens were found at Mackville.
Hormotoma hellicincta (Hall).
Murckisonia hellicincta Hall. Pal. New York, 1, 1847, p. 179,
pi. 39, fig. la-b.
Hormotoma hellicincta Ulrich and Scofield. Geol. Minnesota, 3,
pt. 2, 1897, p. 1017, pi. 70, fig. 15-17.
Description. Elongate, the spire being composed of 8 or more
volutions which are regularly convex and enlarge rapidly from
the apex. The length and width are as 2 to 1. The volutions
are marked upon the center by a flat spiral band. The aperture
is rounded. The pillar lip is nearly straight.
Casts of this species are found rarely and are poorly preserv¬
ed. Those from Mackville measured from 1 to 1% inches in
length.
Hormotoma ? Major (Hall).
Plate III, figure 14.
Hormotoma % major Ulrich and Scofield. Geol. Minnesota, 3, pt.
2, 1897, p. 1018, pi. 71, fig. 5-7.
Hormotoma major Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 650, fig. 889.
Description. A large shell, medium elongate. The sutures
are deep and there is a slight angulation on the upper side of the
whorls. The apical angle is about 25°. The whorls embrace
far up on the preceding ones.
138 Wisconsin Academy of Sciences, Arts, and Letters.
This species has been found at Kaukauna and the casts are
well preserved. They usually consist of about four whorls. The
casts are about 2 inches in diameter and 3 to 3% inches long.
Maclurea (Lesueur).
Maclurea Ulrich and Scofield. Ueol. Minnesota, 3, pt. 2. 1897, p.
1038.
Maclurea Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 664.
Description. The shell is thick and discoidal. Composed of
few whorls. It is reversed, the under side being fiat or nearly
so and exposing aU the whorls. The upper side is convex and
deeply perforated in the center.
Maclurea (Maclurina) cuneata Whitfield.
Plate III, figure 10.
Maclurea cuneata Whitfield. Geol. Survey Wisconsin, 4, 1882, p.
246, pi. 9, fig. 5-6.
Maclurina cuneata Ulrich and Scofield. Geol. Minnesota, 3, pt. 2,
1897, p. 1042, pi. 76, fig. 1-3.
Description. The shell is medium size, ranging up to 3 inches
in diameter. Composed of two or more volutions which increase
very rapidly in size. The upper side of the shell is flat or very
slightly concave between the suture lines. The outer margin is
slightly cuneate.
This is rare in tliis region and only one specimen was found.
This is not a well preserved cast. The diameter is 2% inches.
Fusispira Hall.
Fusispira, Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, 1897,
p. 1075.
Fusispira Grabau and Shimer. N. A. index Fossils, 1, 1907, p.
697.
Description. Shell fusiform, spire elevated. The whorls are
generally convex, with distinct sutures, but they are sometimes
nearly flat with shallow sutures. Elongate, narrow aperture.
Ockernian — Fauna of Galena Limestone Near Appleton 139
Fusispira suhfusiformis (Hall).
Murchisonia suhfusiformis PlalL Pal. New York, 1, 1847, p. 180,
pi. 39, fig. 2a-b.
Fusispira suhfusiformis Ulrich and Scofield. Greol. Minnesota, 3,
pt. 2, 1897, p. 1077, pi. 81, fig. 38-39.
Description. Slender, the whorls gradually enlarging and
very moderately convex. The body whorl is not proportionally
large. The spire ascends rapidly and is composed of 6 or more
volutions. The aperture is oval and acutely extended below.
Casts are found rarely and are usually poorly preserved.
Neenah and Mackville have produced a few specimens. They
measured 2 inches long and 1 inch wide.
Fusispira angusta Ulrich and Scofield.
Fusispira angusta Ulrich and Scofield. Geol. Minnesota, 3, pt. 2,
1897, p. 1079, pi. 81, fig. 28-31.
Fusispira angusta Grabau and Shimer. N. A. Index Fossils, 1,
1907, p. 697, fig. 1002c.
Description. Slender, the whorls usually high, very slightly
convex, almost flat, and separated by verj^ oblique, deep, open
sutures, indicating a rather thick shell.
Cast found rarely in poorly preserved specimens at Mackville.
They vary from 1 to 2% inches in length.
Fusispira vittata (Hall).
Plate III, figure 12.
Murchisonia vittata Hall. Pal. New York, 1, 1847, p. 181, pi. 39,
fig. 3. .
Description. Elongate, fusiform, slender, the volutions being
oblique. Aperture extremely elongate.
Very rare in this region. Found poorly preserved at Mack¬
ville, the specimens measuring % inch in diameter and 1 inch
long.
Fusispira elongata Hall.
Plate III, figure 15.
Fusispira elongata Whitfield, Geol. Survey Wisconsin, 4, 1882,
p. 245, pi. 9, fig. 3.
140 Wisconsin Academy of Sciences, Arts, and Letters.
Description. iShell elongate-fusiform, spire slender and
much elevated, consisting of six volutions which are long and
only slightly convex. Suture strongly marked. Aperture
obliquely elongate-ovate. Surface apparently smooth.
Very rare in this region. One specimen was found at Mack-
ville, is poorly preserved ; it is % inch in diameter and 2 inches
long.
Trochonema Salter.
Trochonema Ulrich and Scofield. Geol. Minnesota, 3, pt. 2, p.
1045.
Trochonema Grabau and Shimer. N. A. Index Fossils, 1, 1907,
p. 669.
Description. Shell turbinate, umbilicated, 4-8 whorls, which
are strongly angulated. Always two more or less prominent
ridges between which lies a flat peripheral space. Aperture
usually oblique.
Trochonema umhilicatum (Hall).
Plate III, figure 9.
Trochonema umhilicatum Ulrich and Scofield. Geo. Minnesota, 3,
pt. 2, 1897, p. 1047, pi. 77, fig. 1-8.
Trochonema umhilicatum Grabau and Shimer. N. A. Index
Fossils, 1, 1907, p. 669, fig. 926.
Description. Short spire and gradually ascending. Shoulder
of volutions flat or slightly concave, occupying about one-third
of the width of the upper side.
Found rarely in this region. Some found at Kaukauna in a
fair state of preservation and measuring about % inch across.
Bibliography
Geological and Natural History of Minnesota. Vol. Ill Part I
and II.
Geology of Wisconsin. 1873-1879. Vol. IV.
Natural History of New York. Part VI. Paleontology, Vol. I.
Hall.
North American Index Fossils. Invertebrates, Vol. I and II,
Grabau & Shimer.
Ockerman — Fauna of Galena Limestone Near Appleton 141
Introduction to the Study of Fossils. Shinier.
Text book of Paleontology. Zittel. (Eastman Translation).
North American Geology and Paleontology. Miller.
Bibliographic Index of American Ordovician and Silurian
Fossils.
Smithsonian Bui. 92, Vol. I and II. Bassler.
Explanation op Plates.
PLATE I.
Fig. 1. Receptac'idites oweni.
2. Ischadites iowensis.
3. Lichenaria typa,
4. Streptelasma coimiculum,
5. Streptelasma profundum.
6. Heteroe'Hnus pentagonus,
7. Schizocrinus nodosus.
8. Heterocrinus heterodactylus.
9. Dendrocinus alternatus,
10. Rhombotrypa quadrata.
11. Lioclemella solidissima,
12. Escharopora subrecta.
PLATE II.
1. Lingula eldeH.
2. Lingula hulburti.
3. Lingula iowensis.
4. Lingula obttcsa.
5. Rafinesquina alternata.
6. Strophomena planumbonua.
7. Strophomena emanciata.
8. Plectambonites sericeus.
9. Orthis costalis.
10. Plectorthis plicatella.
11. Plectorthis fissicosta.
12. Plectorthis dichotoma.
13. Plectorthis sordida.
14. Dinorthis meedsi.
15. Dinorthis pectinella.
16. Herbertella bellirugosa.
PLATE III.
Fig. 1. Platystrophia lynx.
2. Platystrophia biforata.
142 Wisconsin Academy of Sciences^ Arts, and Letters,
3. Herbertella occidentalis.
4. Dalmanella testudinaria.
5. Rhynchotrema increhescens.
6. Rhynchotrema capax.
7. Liospira micida.
8. Liospira vitruvia.
9. Trochonema umbilicatum.
10. Maclurea cuneata.
11. Raphistomina modesta.
12. Fusispira vitata.
13. Liospira obtusa.
14. Hormotoma major.
15. Fusispira elongata.
TRANS. WIS. ACAD
VOL. XXII
/J-
'•Jv
VOIi. XXII
TRANS. WIS. ACAD.
TRANS. WIS. ACAD.
VOL. XXII
PLATE III
SOME CHARACTERISTICS OF THE BACTERIA OF LAKE
MENDOTA
Laetitia M. Snow and E. B. Fred
Notes from the Department of Agricultural Bacteriology, Univer¬
sity of Wisconsin and from the Biological Laboratory of the Wiscon¬
sin Geological and Natural History Survey. XXIV.
In water, there exists a group of microorganisms which may
be termed the true water bacteria. These forms of life are quite
different from the bacteria commonly isolated and studied by
the students of water hygiene. The methods used in a sanitary
study of water are so specialized that those forms related to
diseases are favored, while the indigenous flora is largely sup¬
pressed.
Instead of trying to study in detail the characters of a few
organisms, the object of this report was to investigate the
various groups of bacteria normally found in lake water. It was
realized early in this work that the bacteria in water are so
varied that it is practically impossible to separate all of the
different forms and describe their functions. In order to apply
the results of soil studies to this work on lake flora, the culture
media and many of the methods described by the soil biologist
have been employed. As will be shown later, the flora of soil
and of water have many points in common. Before taking up
these characters of the water flora some of the methods used will
be given:
The medium used for plate counts throughout this work was
Nahrstoff-Heyden or sodium caseinate agar.^ All plates were
incubated at 20° to 22° C., and counted after 15 days of incuba¬
tion.
The results of many comparative studies on the number of
water bacteria which develop in different solid culture media
indicate clearly the superiority of Nahrstoff-Heyden and
nutrose, or sodium caseinate, agar as compared with beef-
peptone agar and gelatin, lake water agar and gelatin, soil ex¬
tract agar and gelatin. Many counts made at different times of
the year on these media show that Nahrstoff-Heyden and sodium
caseinate agars give the highest numbers and are also easy to
prepare. A few representative counts are given in table 1.
144 Wisconsin Academy of Sciences, Arts, and Letters.
The two rich nitrogenous media, beef-peptone agar and
gelatin, which are commonly employed in standard methods of
water analysis, supported the growth of spreaders and liquefiers
and did not give the constancy of results obtained with the
Nahrstoff-Heyden or sodium caseinate. Not only was the num¬
ber of colonies on these two media far in excess of the number
on the bee-peptone media, but the development of typical slowly
growing water bacteria was also favored.
Similar results were reported by Hesse and Niedner, 1898.
They found that for water counts, Nahrstoff-Heyden, an
albuminous substance, generally gave twenty times as many
colonies as the nutrient agar. Several years later Muller, 1912,
repeated the work of Hesse and Niedner and found that water
samples plated on this Nahrstoff-Heyden medium gave very
much higher counts than those on meat-extract peptone gelatin.
Additional data on this subject are given by Winslow and Pres¬
cott, 1924.
Comparison of the direct microscopic count and the plate
count — It is well known that plate counts furnish only an esti¬
mate of the actual number of bacteria present in any material
containing a varied bacterial flora such as milk or water.
Attempts were made to use the direct microscopic method to
determine the number of bacteria in lake water. Unfortunately
the methods worked out in detail for the direct counts have
been for substances rich in micro-organisms and must be modifi¬
ed before they can be used for counting the bacteria in water.
It is necessary first to concentrate the bacteria in water to such a
number that the mathematical error involved in computing from
the number per microscopic field to the number per cubic centi¬
meter is reduced as much as is practicable.
The concentration of bacteria in a solution of low bacterial
content has been tested by Yallet, 1901 ; Muller, 1912 ; Allen,
1915 ; Schuster, 1919 ; and others, but with varying degrees of
success. Colloidal iron, aluminum and copper, as well as sodium
thiosulphite and lead nitrate have been used as flocculating sub¬
stances. Aluminum hydroxide was used as the flocculating sub¬
stance in the direct counts on Lake Mendota water. This
method is based on the fact that a water suspension of this salt
will carry down any bacteria in the suspension wdien this is
centrifuged. The supernatant water, which is free of bacteria.
Snow & Fred — Bacteria of Lake Mendota.
145
is drawn off and the aluminum hydroxide containing the
bacteria is readily transferred to a glass slide and stained. Its
value as a precipitant of bacteria from water was tested on
water which contained a known number of bacteria and it was
found that more than 95 per cent of the bacteria were thrown
down. This is in agreement with the work of Muller who found
97 to 98 per cent of the bacteria carried down in the precipitate.
The water of Lake Mendota generally contains such a small
number of organisms in one cubic centimeter that it was neces¬
sary to concentrate the organisms from ten volumes of the water
to one volume and to prepare the microscopic film from this
concentrated portion.
Since a great variety of types of bacteria are present in lake
water and will not be stained by one simple stain, it was found
best to use a combination of stains to bring out the greatest
possible number of bacteria. Of the stains tested, the most sat¬
isfactory one was prepared as follows: distilled water 40 c.c.,
alcoholic methylene blue S' c.c., and alcoholic fuchsin 0.8 c.c.
Mounts were stained for three minutes.
Considerable time was spent in trying to perfect a direct
microscopic method for lake water which would be practicable
and reliable Perhaps the greatest difficulty was the presence
of debris, thrown down with the collodial suspension, which
stained and could only with difficulty be distinguished from the
bacterial cells.
As many as 50 samples were counted by the two methods.
Table 2 gives a few typical results. The average increase in the
number of bacteria resulting from all the direct counts was
ninefold. Such an increase is in agreement with the results of
most w'orkers. The figures obtained with this method are prob¬
ably a more accurate measure of the actual number of bacteria
in the lake than is obtained with the plate method, but its disad¬
vantages more than outweighed its advantages for this work.
Beginning in the spring of 1917 and continuing to the end
of 1922, with an interruption of 2 years, from August, 1917 to
July, 1919, plate counts were made of the water of Lake Men¬
dota at least every month and generally once a week.^ Samples
were taken from a definite location and at specified depths in the
lake, usually 1, 10, and 20 meters. Almost an equal number of
counts were made at the surface, 15 and 22 meters, but devia¬
tions from the counts at 1, 10 and 20 were not sufficiently great
10
146 Wisconsin Academy of Sciences, Arts, and Letters,
to be of any significance. A specially devised apparatus, a mod¬
ification of the Russell sampler (Wilson 1920), was used for
collecting all samples.
Kinds op Micro-organisms Found on Agar Plates
If the colonies from the typical water plates are examined it
will be noted that the majority of the micro-organisms are short
rods and that many are motile. According to their occurrence
the organisms on plates may be divided into two groups, the nor¬
mal and the abnormal flora. The normal flora consists of bac¬
teria, usually rods and a few cocci, and a small number of
yeasts. The abnormal flora is represented by actinomycetes,
molds and certain typical soil bacteria. Prom 60 to 80 per cent
of the colonies from plates may be grouped as small rods, 10 per
cent as large rods with rounded ends, and 10 per cent as cocci.
Some of the Characteriistics of the Bacteria prom Lake
Mendota Water
The bacteria of lake water, taken far out from the shore, show
certain well defined characteristics. The kind of micro-organ¬
isms present in the water is more or less constant. These indig¬
enous forms are present at all seasons and at all depths. As
compared with the organisms commonly studied, the majority
of them grow slowly on the ordinary culture media and on plate
cultures appear as punctiform colonies. About 10°-25°C. is
the optimum temperature for their growth. In general they do
not form acid or gas from sugars, as was shown by fermentation
tests on agar slants containing 1 per cent of the various sugars
plus a suitable indicator. They do not curdle milk and the ma¬
jority of the true lake forms liquefy gelatin slowly. A consid¬
erable proportion are chromogenic, but long incubation at com¬
paratively low temperatures is necessary to bring out the deep
color. It is in respect to pigment production that the typical
water flora of Lake Mendota is most easily recognized. A large
number of pure cultures were selected from well isolated colo¬
nies on plates poured during the winter, when the true water
bacteria were more prevalent. The characters of some of the or¬
ganisms have been studied and the results presented in table 3.
Snow Fred—Bacteria of Lake Mendota.
147
Chromogenic Bacteria in Lake Water
Except during the period of gross contamination the chromo¬
genie forms make up a high percentage of the indigenous flora.
Their average distribution is shown in table 4. The average per¬
centage distribution of colored bacteria for all counts, except
for March 14 and April 4, 1922, is given as follows :
1. White and cream colored colonies— 52 per cent.
2. Yellow and orange colored colonies — 35 per cent.
3. Pink and red colored colonies — 11 per cent.
In addition to the yellow, orange, pink and red forms, brown,
fluorescent green and a few purple and black colonies are some¬
times seen.
The addition of lactose, or glucose, to agar resulted in a higher
percentage of chromogenic colonies than with Nahrstoff-Hayden
agar alone. The total count on these sugar media is usually
lower, but the percentage of chromogens is higher.
Although no definite relation was noted between color and
season, the percentage of chromogenic colonies was somewhat
smaller in the colder months and the chief colors represented were
yellow and brown. According to color production, the bacteria
at a depth of one meter during the fall were found to have prac¬
tically the same distribution as given for all counts in table 4.
They ranked as follows: —
1. White to cream colored colonies.
2. Yellow and orange, sometimes brown, colonies.
3. Pink and red colonies.
4. Blue and violet colonies found only in small numbers.
5. Green colonies, found only in small numbers, fluorescent
bacteria and algae.
6. Black colonies, very rare.
Of the various chromogens the yellows are at all seasons present
in the greatest numbers. Plate I shows the color production of
some typical chromogenic water bacteria. Some of the charac¬
teristics of the forms shown in plate I were studied and are given
in table 5.
General Distribution op Chromogenic Bacteria
While there is an extensive literature upon chromogenic bac¬
teria, dealing chiefly with the attempt to control the production
148 Wisconsin Academy of Sciences, Arts, o,nd Letters,
of pigment, very little work has been done upon the distribu¬
tion of the various colors in the group. Otto Rahn made a sta¬
tistical study of the group, in the progress of which he ascer¬
tained the number and percentage of white and colored forms
in the various genera. A summary of his figures may be made as
follows : In 11 genera of the Eubacteria, containing 1194 species
(omitting the phosphorescent forms) he found 855 or 72 per
cent of colorless or white, 198 or 17 per cent of yellow and broAvn,
70 or 6 per cent of red, 14 or 1 per cent of blue, 51 or 4 per cent
of fluorescent green (all Pseudomones) , 2 or 2 per cent of
green, and 4 or 0.3 per cent of black.
In an effort to get a general idea of the distribution of col¬
ored forms, the species given in Bergey’s Manual of Determina¬
tive Bacteriology 1923, were arranged in table 6. In making this
table the color given on nutrient agar was chosen, when that was
stated. When it was not recorded, the medium most nearly like
plain agar was selected as the standard. Forms that were stated
as probably the same as others were omitted. The percentages are
not of species, for when several habitats were listed, each entry
was counted. A chance isolation different from the habitat was
omitted. For many forms there was no color given, often no
description whatsoever; these weie classified as ‘‘ Undesig¬
nated. Habitats having less than 10 entries were omitted.
‘‘Fresh water’’ includes all kinds of Avater but the sea. “Sea
water” includes one entry from “Salt fish ana sea salt”. In a
few cases, where the forms produced a color in the medium
different from that of the colony, only the color of the bacteria
is recorded. Summarizing the data for the Avater habitats Ave
find out of 198 entries, of which only 21 are from salt w'ater, 76
(38.4 per cent) are white, grey, colorless, or undesignated, 45
(22.7 per cent) are pink or red, 44 (22.2 per cent) are yellow,
orange or brown, 18 (9.1 per cent) are violet, 9 (4.5 per cent)
green or fluorescent, 2 (1.0 per cent) each are blue, black and
iridescent. It is exceedingly interesting to note the high per¬
centage of red and pink forms. Violet and black forms are,
Avith the exception of one entry for violet in soil, confined exclu¬
sively to water habitats.
In connection with the distribution of colors among the bac¬
teria, it is interesting to note that, in the few papers dealing with
the distribution of colors in flowering plants, white is reported
Snaw (& Fred — Bacteria of Lake Mendota. 149
to be the most prevalent, with yellow second in importance.
Lovell in two papers gives statistics regarding colors in flowers.
These figures may be summarized as follows: In 4,020 species
the flowers of 1,244 (30.9 per cent) were green, 956 (23.8 per
cent) were white, 801 (19.9 per cent) were yellow, 257 (6.4 per
cent) were red, 437 (10.9 per cent) were purple and 325 (8.1
per cent) were blue. In a later article he eliminates the wind
pollinated flowers and finds of the 2,972 species remaining, 955
(32.1 per cent) were white, 790 (26.6 per cent) yellow, 519 (17.4
per cent) blue, 366 (12.3 per cent) red, 223 (7.5 per cent) green
and 119 (4 per cent) purple. Miss Mark states that she found
1,500 flowering species in Ohio (omitting the grasses and sedges)
and gives numbers and percentages of the different colors. The
sum of her figures is 1,828, however, which leaves the reader
uncertain whether some species have been reported under more
than one head. In either case the order of their abundance is
the same, although the percentages are different. White flowers
were the most abundant, 29.4 per cent (using the 1,828 total),
yellow and orange 21.9 per cent, purple and violet 19.8 per cent,
red and pink 12.5 per cent, green 8.7 per cent, blue 7.4 per cent.
Keriier and Oliver (Vol. II p. 182) give the following per cents
for the colors found in the Baltic flora : white 33 per cent, yel¬
low 28 per cent, red 20 per cent, blue 9 per cent, violet 8 per
cent, brown 2 per cent. Drummond, noting colors of flowers
blooming in the various months of the year, reports 539 plants
of Ontario and Quebec with 913 entries, showing that nearly
all forms bloom during two months. Of these 913 entries 37
per cent v/ere white flowers, 26 per cent yellow, 15 per cent
purple and violet, 11 per cent blue, 7 per cent green, and 4 per
cent red or pink. It appears therefore, from the data at hand,
that white is the dominant color both among flowering plants
and the bacteria, with yellow a close second, while red, green,
blue, and violet show decided variations.
Conclusions
The kinds of bacteria found in Lake Mendota water, re¬
gardless of depth are remarkably uniform. The colonies are
usually punetiform and grow slowly. Small motile and non-
motile rod forms predominate. Sodium caseinate agar incu-
150 Wisconsin Academy of Sciences, Arts, and Letters.
bated at 20° C. is especially favorable for the growth of these
organisms.
Direct counts show that the number of bacteria in lake water
is much greater than is indicated by plate counts.
The normal flora of Lake Mendota is characterized by a great
number of chromogenic forms, although white forms predom¬
inate.
Of the colors, various tones of yellow are the most prevalent.
Red, violet, black, blue, green, fluorescent and iridescent forms
are also found.
Among flowering plants there is also a dominance of white,
followed by yellow as next in importance.
Bibliography
I. Fred, E. B., Wilson, F. C. and Davenport, Audrey. Ecology. 5: 322.
1924.
2. Allen P. W. J. Inf. Dis., 22: 245. 1918.
3. Muller, P. Th. Arch. f. Hyg., 75: 189 and 321. 1912.
4. Schuster. Zeit. f. Hyg., 88: 402. 1919.
5. Vallet. Arch. d. med. exper. et d’anatom. Pathol. Juillet, 1901.
6. Wilson, F. C. J. Bact., 5: 103. 1920.
7. Prescott, S. C., and Winslow, C. E. A. Elements of Water Bac¬
teriology, New York, 1924.
8. Bergey. Manual of Determinative Bacteriology. Baltimore, 1923.
9. Kerner, A. and Oliver, F. W. The Natural History of Plants.
New York, 1895.
10. Mark, Clara G. Ohio Nat., 7: 57. 1907.
II. Lovell, John H. Amer. Nat., 33: 493. 1899.
12. - Amer. Nat., 36: 203. 1902.
13. - Amer. Nat., 37: 365, 443. 1905.
14. - The Flower and the Bee. Chap. XIV. New York, 1918.
15. Rahn, Otto. Centrbl. f. Bakt. etc., Abt. II, 46: 4. 1916.
16. Drummond, A. T. Can. Record of Sci., 5: 280. 1892-3.
Snow & Fred — Bacteria of Lake Mendota.
151
Department of Agricultural Bacteriology,
University of Wisconsin.
Table 1. Comparison of the number of bacteria in Lake Mendota
water on different media. Number in one cubic centi¬
meter of water.
Table 2. Comparison of the plate method with the direct microscopic
method of counting bacteria in lake water.
152 Wisconsin Academy of Sciences, Arts, and Letters.
Table 3. Showing the characteristics of some of the water bacteria.
+ Positive. Almost without exception these organisms liquefy gelatin slowly.
— Negative.
Table 4. Showing the number of chromogenic colonies in Lake Men-
dota water.
* Heavy rain and much soil washed into lake.
TRANS. WIS. ACAD., VOL. XXII.
PLATE IV,
SNOW AND FRED.-CHROMOGENIC BACTERIA FROM LAKE MENDOTA.
COCKAYHE BOSTON
Table 5. Showing some of the characteristics of chromogenic forms found in Lake Mendota water.
Snow d' Fred — Bacteria of Lake Mendota.
•a’g,’g)+i + i + i’Si 1
1 1
ii+ii
1 1 1
gggsslg Jigs
O 4) ^ (P SS 0) Q) ^
fifiCCCctiC 0 0 C
I.E
I ++ i 1 1 + 1 + I I
k • >
sa*® ! o’®
^ ^ JS Cl
pL,^m>HCpqQOSp;ot^
++ I I I U. 1 1 I +
! i i i ! I i I P ! I
'g'g'g'g'g'g'g'g g-g-g
oooooooossgu
«2CQCGC0CQUlC/aC0S)^tJO
153
154 Wisconsin Academy of Sciences, Arts, and Letters.
•2
h ^
|cq
|l
■ij
§ 5S)
C3iCq
o >
rfi
CO
'a
«o
a ^
(0 ^
< ^
H
g c«
Q>
^ O
a ®
1 ^
fl'O
2 o
5jq
Qi si
^ B
o.S
a ®
a a
•S'®
M M 05
«o OJ «
00 OJ oi
eo o
a
2
I o
0)
I 2
I ^
^ tm
> m « o
I i
3 -o
h3 O
No other color indicated.
NOTES ON PARASITIC FUNGI IN WISCONSIN— XII
J. J. Daves
In ‘‘Notes” V, p. 63, Artemisia ludoviciana was given as a
host of Plasmopara hoist edii (FarL) Berl. & De Toni. This was
an error as the mildew in the collection is Peronospora lepto-
sperma DBy. It has also been collected on a host referred to
Artemisia serrata at Jump River and Caryville.
The Peronospora on Cerastium nutans recorded in the pro¬
visional list under the name P. aUinearum Casp. is referred to
P. tomentosa Fckl. by Gaeumann {Beitr. z. Krypt. fl. d. Schweiz
54:61) with the statement that it is biologically distinct. As far
as observed it is confined to the single species of host in Wis¬
consin.
For the ascigerous species having Fusicladium as the conidio-
phorous stage the genus Endostigme has been proposed by H.
Sydow (Ann. Mycol. 21: 173).
The reference of Montagnella heliopmdis (Sehw.) Sace. to
Rosenscheldia by Theissen & Sydow {Ann. Mycol. 13: 649. taf.
IV, tig. 7) was probably based on examination of immature ma¬
terial. When fully developed the stromata containing the loeules
are approximately spherical as stated by Ellis & Everhart;
(North American Pyronomycetes, 'p. 252). Collections of this
parasite are usually sterile as stated by Trelease in the Prelim¬
inary List or immature. In material on Aster Tradescanti col¬
lected at Blue River September 15 some of the loeules contain
uniseptate spores about 20-24 x 4jLt, but asci were not seen.
In “Notes” VII, p. 208 it was stated that traces of Taphrina
ulmi (FcM.) Johans, had been seen in Wisconsin. It was col¬
lected in 1923 on TJlmus americana at Avoca and in more abund¬
ance at Edgerton and scantily developed at Wyeville. This has
since been found at Avoca and Madison.
All specimens of Septoria on Polygonum that have been
collected in Wisconsin have been referred to S. polygonorum.
Desm. They agree better with the description given by Dr.
Martin in “Enumeration and Description of the Septorias of
156 Wisconsin Academy of Sciences, Arts, and Letters.
North America” {Journ. My col. 3: 68) than with those of
European authors. The sporules vary from 23-63 x 1-2/x and
the colored border of the spots is purple, brown or nearly or quite
absent. The accompaniment of the Septoria on Polygonum
cilinode by the mucedine referred to in “Notes” VIII, pp. 415-
6 suggests that it may be distinct.
A Septoria on Diervilla was recorded under the name S.
diervillicola Ell. & Evht. in the provisional list. Both Ellis and
Everhart and Peck had described it as a new species giving the
binomial Septoria diervillae. Saccardo preserved S. diervillae
Pk. and changed S. diervillae E. & E. to S. diervillicola {Syl.
Fung. 10: 356) evidently taking Peck’s publication to be the
prior one. In the Journal of Mycology 3: 51, Dr. Martin
stated that Peck’s 38th Report, in which the publication was
made, was not issued until 1886 although it is dated Dec. 31,
1884, while Ellis & Everhart published their description in 1885.
It appears therefore that Septoria diervillae E. & E. is the
proper designation as used by Dr. Martin in Septorias of North
America.
In “Notes” III, p. 254, it was stated that Quercus alba should
be eliminated from the provisional list as a host of Leptothyrimn
dryinum Sacc. In July 1923, however, the parasite was collected
on that species of host at Caryville. In this collection the pyc-
nidia are borne on suborbicular greyish spots having a reddish
brown border, 2-5 mm. in diameter which resemble those caused
by Phyllosticta phomiformis Sacc. The clypei are epiphyllous
60-100/jt in diameter, the sporules elliptical to ovate, brown tint¬
ed, 12-17 X 7-lOjLt. The small spots and the small pycnidia
would deter one from referring this to L. maculicolum Wint.
with which the size of the sporules would seem to ally it.
Piggotia fraxini B. & C. is referred to Asteromella by Petrak
in Annales Mycologici 21 : 269.
In the provisional list Steironema ciliatum was given as a host
of Eamulana lysimachiae Thuem. with question. There is much
difference in the appearance of Ramularia on this host. Early in
the season on young leaves the infected area is not delimited
but spreads over a large portion or all of the leaf which dies.
Later in the season on mature leaves there is definite spotting.
In a specimen collected at Madison in September the tufts are
Davis — Parasitic Fungi in Wisconsin — XIL
157
epiphyilous with black tubercular bases. In all of the speci¬
mens the conidiophores and conidia are like those on Lysimachia.
A collection of Cercospora sedoides E. & E. on Pentliorum
sedoides made at Blue River in September shows conidiophores
sometimes densely fasciculate, tortuous, occasionally branched,
up to 50 /i, long. The well developed conidia are 80-90 /a long,
7-septate.
The epiphyilous Cylindrosporium referred to in “ Notes VIII,
p. 428, as occurring on Spiraea densiflora in Idaho was collected
in 1923 on S. salicifolia at Cornucopia where the hypophyllous
form also occurred. In the epiphyilous form the sporules are
40-70 X 3^. It was referred to Cylindrosporium salicifoliae
(Trel.) Davis, a species which Petrak has referred to the dubi¬
ous genus Phleospora {Ann. Mycol. 20 ; 210).
The conidiophores of Piricularia parasitica E. & E. were de¬
scribed as being simple or bifid. When fully developed however
they may be dichotomously branched. Occasionally opposite
branches are produced. The character of the conidiophores, the
comparatively narrow conidia and the habitat make this an
aberrant species. It has been found only on Elymus and Hys-
trix in connection with the species of Phyllachora that is referred
to Ph. graminis (Pers.) Pckl.
Of a collection on Lathy r us venosus referred to Cercospora
viciae Ell. & Hoi. it was noted ‘ ‘ conidiophores up to 50^, conidia
up to 70^ long.
All of the specimens of Cercospora eiionymi Ellis that I have
seen are devoid of color throughout.
All specimens of Cercospora on Vernonia collected in Wis¬
consin have been referred to C. vernoniae Ell. & Kell. C. oculata
Ell. & Kell, being considered to be a form of that species.
Because of failure to secure spore germination and the
peculiar habitat Doassansia zizaniae Davis {Bot. Gaz. 26: 353,
1898) was recorded under Sclerotium in the provisional list.
In North American JJ stilagineae {Contributions from the
Cryptogamic Laboratory of Harvard University No. 57) it was
placed in the list of excluded species and was omitted from
North American Flora 7 :1. Prof. W. H. Davis informs me that
he obsen^ed germination in July 1923 of the Tilletia type with
sporidia 16x4/* of overwintered material sent from Wisconsin.
There is an error in the description due to the appearance of
sections that were not strictly equatorial. There is but a single
158 Wisconsin Academy of- Sciences, Arts, and Letters.
layer of spores under the cortical layer instead of “two or
three. ’ ’
Additional Hosts
Specimens on young seedling plants of Artemisia hiennis from
Herbster are tentatively referred to Synchytrium aureum
Schroet. They occurred within a few feet of infected Petasites.
On the leaves of the latter the galls (epiphyllous) are sometimes
flattened in a vertical plane and are sometimes confluent in lines
forming ridges. On some leaves of Petasites the scattered galls
are nearly black recalling the form on Aster for which the name
Synchytrium nigrescens was proposed in “Notes” X.
Albugo Candida (Pers.) Kuntze.
On Sisymbrium altissimum. Edgerton.
Peronospora parasitica (Pres.) Tul.
On Arabis laevigata. Wyeville. The binomial, as used here,
applies to a congery of host-adapted races having doubtless a
common origin and with but little or no morphological differen¬
tiation. There are two hypotheses as to the origin of such a
group. According to one it was derived from a generalized form
that occurred indiscriminately on the various hosts but which
underwent physiological changes adapting it to the particular
host upon which it was growing until the generalized ancestral
form was lost. According to the other hypothesis variants of
parasite and hosts occur and unusual conditions arise allowing
passage from one genus or species of host to another and follow¬
ed by physiological changes that result in adaptation to the new
host. Following the first hypothesis the process is now a closed
one and no new races will arise, those that fail to survive will
not be replaced and the number of races will remain as at
present or diminish unless present races are subdivided. The
second hypothesis would indicate a continuing process with new
races arising from time to time.
Agrimonia gryposepala should have been recorded as a host
of Peronospora potentillae DBy. which has been collected on that
species at Blair, Wild Rose, and Lone Rock.
In “Notes” XI it was stated that what was thought to be
Peronospora myosotidis DBy. had been seen in very small
quantity on Myosotis laxa. In 1923 the mildew was collected on
Davis — Parasitic Fungi in Wisconsin — XII.
159
M. virginica at Avoca in sulSicient quantity for determination
and with oospores in stems and branches.
Phyllachora occurs in Wisconsin on Cinna latifolia but no
mature material has been collected.
Ascochyta thaspii Ell. & Evht. On Cicuta maculata. Haugen.
The largest sporule measured was 33 x 6
Ascochyta compositarum Davis. On Eupatorium purpureum.
Blue Mounds (Oilbert & Davis). In this collection the spots,
which become pale in the center, are apparently formed by con¬
fluence of small, dark, angular, intervenular areas some of which
are distinct. The pale brown or amber-colored pycnidia are de¬
pressed globose, 140-180^ in the horizonal diameter and the
sporules 20-27 x 5-6 Biseptate sporules occur very rarely.
The collection was made in October.
Septoria gei Rob. & Desm. On Geum virginianum^ Wyeville.
G. strictum, Hixton and Two Rivers. G. canadense, Glen Haven.
The sporules range up to 50 long, the pycnidia often imper¬
fectly developed.
Septoria violae West. On Viola lanceolata. Avoca. (Gilbert
& Davis). The sporules are not well developed in this collection.
Septoria lycopi Pass. A collection on Ly copus uniflorus was
referred to this species in “Notes” X with the statement that
the sporules were about 30 x II/2 Of a subsequent collection
the following notes were made : Spots at first indefinite and
olivaceous becoming orbicular and cinereous above with a
definite margin and sometimes a purple border ; pycnidia
amphigenous, globose, ostiole papilliform, 50-80 in diameter ;
sporules straight, continuous, 20-30 x 1 /*. On leaves of Lycopus
americanus. Blue River, October 29, 1923.
Gloeosporium septorioides Sacc. On Quercus alba. Blue
Mounds. (Gilbert & Davis). Although a septum was not seen in
the sporules this is doubtfully distinct from Marssonma martini
(S. & E.) Magn. The spots are mostly 4-6 mm. in diameter and
the acervuli multiple. The collection was made in October and
the development was probably from infection late in the season.
The sporules are 12-17 x 2-2% /*.
Marssonina potentillae (Desm.) Magn. var. tormentillae Trail.
On Potentilla canadensis, Wyeville. The sporules are 15-20 x
4-6
160 Wisconsin Academy of Sciences, Arts, and Letters.
Septogloeum salicinum (Pk.) Sacc. On Salix hnmilis. Iron
River and Brule.
Ramularia pratensis Sacc. On Rumex persicarioides. Madison.
In a collection on Ranunculus acris made at Cornucopia and
referred to Ramularia aequivoca (Ces.) Sacc. the conidiophores
are about 20 ^ long, the conidia 27-43 x 2-21/2 At cylindrical,
obtuse, continuous or 3-septate. Of a collection on the same
host from Herbster it was noted : Conidiophores 12-17 long,
conidia 20-40 x 2%-3 3-septulate.
Ramularia actaeae Ell. & Hoi. On Actaea rubra neglect a.
Cornucopia.
Ramularia barbareae Pk. On Barbarea stricta. Wyeville.
The short (10 /^) conidiophores do not seem to be branched but
rather developed in small tufts from a common base. Many of
the conidia are somewhat less than 3 At in diameter.
Ramularia arvensis Sacc. On Potentilla canadensis. Wye¬
ville.
Ramularia asteris (Phil. & Plowr.) Bubak. On Aster
sagittifolius, Cornucopia. A. azureus. White Lake. A. Trade-
scanti, Caryville. A. paniculatus, Blue River. The conidio¬
phores and conidia are sometimes more slender (3-4 a*) than the
descriptions indicate.
Cercospora zebrina Pass. On Trifolium dubium. Mazomanie.
The publication of this name seems to antedate that of
Cercospora helvola Sacc.
There occurs sparingly on leaves of Eupatorium purpureum a
parasite that I have referred to Cercospora perfoliata E. & E.
although it lacks the brown color of that species.
Entyloma compositarum Farl. On Rudheckia laciniata. Sauk
City. (Kuntz & Davis).
Piiccinia graminis Pers. Uredo and telia on Cinna latifolia.
Blue River.
Lycopus uniflorus is the most common host of the aecial stage
of Puccinia angustata Pk. in Wisconsin.
Piiccinia peckii (De Toni) Kell. Aecia on Oenothera pumila..
Wyeville and Arena.
Davis — Parasitic Fungi in Wisconsin — XIL
161
Additional Species
The Eeport of the Geological Survey of Wisconsin, 1873-9
contains a list of Fungi prepared by A. F. Bundy. In this list
Peronospora nivea Unger was included but hosts were not
given. Nothing further was known of this mildew, now known
as Plasniopara nivea (Ung.) Schroet,, in Wisconsin until 1923
when it was found in small quantity at Haugen on leaves of
Cicuta maculata. [This occurred on a single leaf at Balsam
Lake in 1924.]
Phyllosticta steironematis Hearn. & House. On Steironema
ciliatum. Blue River. Not well developed in this collection;
perhaps because of the dry season.
Phyllosticta lappae Sacc. On Arctium minus, Haugen. Some
of the specimens that have been referred to Ph. decidua E. & K.
are much like this.
Of a collection on leaves of Lactuca canadensis made at Blue
River, September 16, 1923, the following notes were made:
Stroma superficial, black, composed of small polygonal fuligen-
ous cells, following the veinlets, anastomosing and becoming con¬
fluent on leaf areas 1-5 cm. in diameter which become dead,
mostly epiphyllous, sometimes hypophyllous. No spore bodies
seen. This has been labeled Asteroma lactucae nom. herb. It
may be that it is not distinct from Asteroma atratum, Ohev.
Septoria margaritaceae Pk. On Anaphalis niargaritacea^
Iron River. In this collection the prominent pycnidia are up
to 200 fjt, in diameter, the wall of the distal half more or less im¬
perfect, A collection from Herbster bears well developed
pycnidia.
Septoria erechtitis Ell. & Evht. On Erechtites hieracifolia.
Blue River.
Of a collection on lower leaves of Lepachys pinnata made dur¬
ing a dry season the following notes were made : Spots circular
to elliptical or sometimes oblong, sordid white, definite with a
narronr dark border, 1-3 mm. in diameter; pycnidia epiphyl¬
lous, sometimes prominent, small (50-75/^), black; sporules
hyaline, curved, continuous, 30-50x1-1%-“. Avoca, June 19,
1923. Where the spots are numerous the leaf area dies and be¬
comes brown. This is provisionally considered to be a state of
Septoria infuscata Wint.
11
162 Wisconsin Academy of. Sciences, Arts, and Letters.
Cylindrosporium triostei Kell. & Sw. On Triosteum per-
foUatum. Haugen. Conidia mostly 35-50/a long.
Ovularia decipiens Sacc. On Ranunculus acris. Bayfield. I
have seen no mention of the occurrence of this parasite in
America.
Ramularia melampyri Ell. & Dearn. On Melampyrum lineare.
Cornucopia. In these collections there is little or no deforma¬
tion or discoloration of the infected leaves. The conidia range
up to 30 fi in length.
Cercospora simulata Ell. & Evht. On Cassia ma/rylandica.
Avoca. (Gilbert & Davis).
Cercospora leptandrae n. sp. Hypophyllous oii angular
areas limited by the veinlets, 2-3 mm. in diameter which are
blackened by the mass of conidiophores, upper leaf surface un¬
altered; conidiophores scattered or in small tufts, dark brown,
more or less flexuose and denticulate, pluriseptate, sometimes
branched, 75-135 x 5/i ; conidia subfuligenous, obclavate, straight
or slightly curved, becoming 3-5 septate, 40-60 x 5/a. On leaves
of Veronica (Leptandra) virginica, Blue River, Wisconsin,
September 14, 1923.
Tilletia corona Scribner. On Leersia virginica, L. oryzoides
and L. lenticular is. Blue River.
JJromyces seditiosus Kern. Telia on Arutida tuberculosa.
Avoca. ( Gilbert & Davis) .
Fuccinia jussiaeae Speg. Aecia and telia on Ludvigia
polycarpa. Blue River. In these collections the aecia are in
small scattered groups and the telia on stems, branches, petioles,
occasionally on midribs, but especially on fruiting calyces.
Fuccinia antirrhini Diet. & Hoi. This occurs in Wisconsin on
Antirrhinum majus (cult.).
Aecidium myosotidis Burrill. On Myosotis virginica. Dane
County opposite Sauk City. (Kuntz & Davis and Welles &
Davis.)
A few years ago a concrete road was built through a marsh
in a suburb of Madison. In the fall TJstilago sphaerogena Burr,
and Fuccinia flaccida B. & Br. appeared on Echinochloa crusgalU
along the road where, however, they were not found subsequent
years. The inference that the road builders had brought their
equipment from further south and introduced the parasites
Bavis — Parasitic Fungi in Wisconsin — XII.
163
therewith seems a fair one. The further inference that neither
the smut or the rust are able to withstand the rigor of Wisconsin
winters may be drawn.
Herbarium of the University of Wisconsin,
March, 1924
NOTES ON PARASITIC FUNGI IN -WISCONSIN— XIII
J» J, DavI)S
Petrak considers Asterina ruhicola E. & E. and Coccochora
rubi Davis as forms of a single species which he designates Stig-
matea ruhicola (E* & E.) Theiss. (Ann. Mycol. 22: 109 et seq.).
That they are closely related phylogeneticaliy seems clear. As
far as observed the dothidial form referred to Coccochora occurs
on blackberries only, the Asterina on raspberries only. To de¬
termine their degree of relationship will require inoculation ex¬
periment.
Petrak rejects the genus Didymellina Hoehn. and refers the
ascigerous stage with which Heterosporium gracile Sacc. is con¬
nected to Didymella and refers Mycosphaerella pinodes (Berk.
& Blox.) Niessel to the same genus. He also suggests that My-
cosphmrella letTialis Stone should be referred to Didymella ( Ann.
Mycol. 22: 17-18).
Wolf refers Mollima earlwma Sacc. to Diplocarpon which he
now places in Phacidiaceae {Journ. Elisha Mitchell Scientific
Society).
Leaves of Corylus rostrata bearing immature Mamiania were
taken at Bruce on which were orbicular light brown to cinereous
spots 10-15 mm. in diameter. Each spot bore in its center a
Mamiania stroma and on the remainder scattered acervuli of
Gloeosporium coryli (Desm.) Sacc. Leptothyrium corylinum
Fckl. is reported to be a conidial stage of Mamiania coryli
Batsch.) DeNot. The fungus referred to Gloeosporium coryli
(Desm.) Sacc. somewhat resembles Leptothyrium, the cuticular
covering of the acervuli being black and clypeoid. The sporules
however are not like those of L. corylinum but are similar to
those of L. coryli Lib. In the desbription of Gloeosporium coryli
(Desm.) Sacc. it is stated that the acervuli are on the lower
surface of the spots, seldom on the upper. The reverse is the
case in Wisconsin.
Phacidium taxi Fr. was recorded in ‘'Notes’^ X as occurring
on Taxus canadensis in Wisconsin. The record was based on
166 Wisconsin Academy of Sciences^ Arts^ and Letters.
immature material in whieh. no spores had formed. Another col¬
lection on this host was made at Appolonia in September 1924
but this also is immature. However a few spores were found but
they probably were not mature. The following notes were
made : Ascomata hypophyllous, biseriate, cinerous when dry,
dull black then wet, orbicular, about I/3 mm. in diameter with
thick black walls and a central stellate opening; hymenium dis¬
coid, sometimes extending past the acute sulcus on to the lower
surface of the epithecium where however the development of asci
is rudimentary ; asci crowded, straight, cylindrical, 40-60 x 4-6 p. ;
spores obliquely uniseriate, hyaline, fusiform, straight,
6-10 X 2-Sfx ; paraphyses slender, filiform, lax. Usually all of
the leaves on the twig bore the ascomata. Whether or not this
is the Friesian species is an open question.
In the classification of the Hypodermataceae proposed b}^
Dearness the shape of the apothecia is ignored and to the genus
Hypodermella, as amended, is referred Lophodermium amplum
Davis (“Notes’^ V, pp. 695-6), which thus becomes Hypoder¬
mella ampla (Davis) Dearn. {Mycologia 16: 152).
Phyllosticta renonana Sacc. & Roum. of the provisional list is
Fh. typhina Sacc. & Malbr. whieh is the prior name.
In Notes V, p. 701, record was made of pycnidia in the loculi
of Phyllaehora on Elymus for which the binomial Cytodiplosporo.
elymina was proposed with the suggestion that it occurred in the
life cycle of Phyllaehora. Petrak is of the opinion that this
bears a parasitic relation to the Phyllaehora and has proposed
for it a new genus, Davisiella {An7i. Mycol, 22: 133^). A simi¬
lar development has been observed two or three times in loculi of
Phyllaehora on Calamagrostis canadensis but not in sufficient
abundance to secure a specimen. On this host the sporules are
about twice the length of those on Elymus, 10-20/1,.
The genus Saeidium is being dropped by mycologists follow¬
ing von Hoehnel. The parasite on leaves of Betula described by
Peck under the name Septoria microsperma and referred to
Saeidium in “Notes’’ I, pp. 88-9, is placed in von Hoehnel’s
proposed genus Cylindrosporella by Petrak (Ann. Mycol.
22:42-3).
What is probably a form of Cercospora varia Pk. has been
found on Viburnum pubescens (Lewis, Aug. 1, 1924) in which
the conidiophores are hyaline and the conidia slender, tapering,
Davis — Parasitic Fungi in Wisconsin — XI 11. 167
sometimes exceeding 100 p. in length. This would fall in Cercos-
porella if taken by itself,
Septoria gratiolae Sace. & Speg, of the provisional list is the
parasite to which Ellis & Martin gave the same name for which
Berlese & Voglino substituted Septoria elUsii. Although there
has been no opportunity for comparison it has been assumed that
these are conspecific. The Wisconisn specimens bear sporules
25-50 X %-iy2P more or less curved and tapering to the atten¬
uate apex.
Ramularia waldsteiniae Ell. & Davis was collected at Hay¬
ward in 1924. In this collection the spots are dark purple above,
more or less elongated parallel to the veins and often limited by
the veins. On the lower surface the' spots are pale brown and
less sharply limited.
Ramularia magnusiana (Sacc.) Lindau as it has been seen in
Wisconsin on Trientalis americana bears conidia that are seldom
septate, 10-33 x l%-3 p the shorter ones fusoid. The conidio-
phores spring from scattered black tubercles 25-40 p in diam¬
eter and are mostly fuligenous tinted, 20-60 x 2-3 p. The spots
are usually angular, limited by the veinlets and immarginate,
light brown becoming paler in the center. While this departs
widely from the type it nevertheless appears to be a variant of
that parasite.
Glomerularia corni Pk. was recorded in the provisional list
on Cornus canadensis, Lonicera canadensis and L. ohlongifolia.
Glomerularia lonicerae (Pk.) Dearn. & House has been used as
a designation of the parasite on Lonicera but no distinguishing
characters are given. (N. Y. State Museum Bulletin: Report of
the State Botanist for 1921, p. 85).
In some notes by von Hoehnel published after his death by
Weese (Centralh. f. Baht. etc. 60) the suggestion is made that
the genus Fusicladium be restricted to conidial forms of V enturia
and that the species that have been referred to that genus but
which have Mycosphaerella (Carlia) as their ascigerous state be
referred to Passalora. Fusicladium depressum (B. & Br.) Sacc.
on Angelica he therefore designated Passalora depressa (B. & Br.)
Hoehn. and Cercospora platyspora Ell. & Hoi. on Taenidia Pas¬
salora punctiformis (Wint.) Hoehn. Fusicladium punctiforme
Wint. and Cercospora platyspora Ell. & HoL based upon the
same parasite, were published in Hedwigia number for January
168 Wisconsin Academy of Sciences, Arts, and Letters.
and February 1887 and the Journal of Mycology number for
February 1887 respectively and it would probably now be im¬
possible to determine which has priority in time of publication.
To this species von Hoehnel also referred Cercospora sii E. & E.
on Sium which is certainly very similar. He further suggested
that it might be merely a short-spored form of Passalora de-
pressa (B. & Br.) Hoehn. In the same publication Scolecotri-
chum graminis Fckl. is also referred to Passalora while Scoleco-
tt'ichum maculicola Ell. & Kell, is referred to Cladosporium with
the suggestion that it may be C. phragmitis Opiz. For the much
named Passalora fasciculafa (C. & E.) Earle the genus Cercos-
poridium Earle (Muhlenhergia 1 : 16) which had been aban¬
doned by its author {Torreya 2 : 160) , is revived and the binomial
Cercosporidium fasciculatum (C. & E.) Hoehn. added to its
generous nomenclature.
Comparison of American specimens of Cercospora suhsctn-
guinea E. & E. with European specimens of Ramularia rubi~
cunda Bres. indicates that they are conspecific. As stated in
‘'Notes’^ I, p. 83, the plant is referable to Ramularia rather than
to Cercospora. Cercospora subsanguinea E. & E. was published in
1887, Ramularia rubicunda Rres. in 1896. If the older specific
name is preserved a new binomial is necessary. Ramularia rub¬
icunda Bres. is in general use in Europe. All references to the
plant in Europe are under that name and all European speci¬
mens are so labeled. As it is necessary to change the designation
of the species as it occurs in America it seems to me best to
adopt the name used in Europe. I have therefore labeled the
Wisconsin specimens Ramularia rubicunda Bres. This obviates
any change in Europe and reduces to synonymy but one name
instead of two. This may be taken as an illustration of the fact
that rigid adherence to the rule of priority may cause more
trouble than it cures.
Gloeosporium equiseti E. & E. was considered by Bubak to be
identical with Septoria detospora Sacc. and was made the type
of new genus becoming Titaeospora detospora (Sacc.) Bubak.
(Ann. My col. 14: 345 (1916).
Through the kindness of Dr. House I have had opportunity to
examine authentic material of Vermicularia Violae-rotundifo-
liae (Sacc.) House. As a result I conclude that the parasite
on Viola scabriuscula recorded in ‘‘Notes’’ XT, p. 297. under
Davis— Par<isitic Fungi in Wisconsin — XIII. 169
the name Collet otricJmm violarum n. sp. is conspecific therewith.
They seem to be referable to Colletotriehum. i
Collet otrichum. salmonicolor O’Gara is united with Gloeospor-
ium fusarioides Ell. & Kell, by Dearness who proposes the com¬
bination Colletotriehum fusarioides (E. & K.) O’Gara {Mycol-
ogia 16: 169) with the suggestion that it is a conidial state of
Glomerella cingulata (Stonem.) Sp. & V. Schr. If that proves
to be tlie case of course the combination will fall into synonymy.
With the record of Cylindrosporium vermiforme Davis as oc¬
curring on Alnus crispa in Wisconsin was the statement that the
sporules in the collection on that species of host were but about
3 in diameter. Material on A. crispa collected at Hayward by
Gilbert & Davis has normally developed sporules.
For the microeonidial state of Cylindrosporium hetiilae Davis
the name Gloeosporium hetulue-papyriferae Dearness & Over¬
holts ha,s been proposed {Mycoloyia 16: 167).
Entyloma linariae Schroet. var. gratiolae Davis was collected
in 1923 at Haugen on the same host. In neither of the locali¬
ties was Entyloma found on Veronica.
Peridermium coloradense (Diet.) Arth. & Kern was recorded
in the 4th supplementary list as occurring on Picea mariana in
Wisconsin hut it was not included in the provisional list. This is
now thought to be Peridermium elatimim A. & S. the aecial
stage of Melampsorella caryophyllacearum Schroet, the further
stages of which have not been collected in Wisconsin. The aecial
stage on Picea has been observed at but one station in the state
the usual host being Abies halsamea.
Additional Hosts
Plasmropara geranii (Pk.) Berl. & De Toni. On Geramium
Bicknellii. Spooner.
Peronospora sordida B. & Br. On Scrophularia leporella.
Blue River.
Colletotriehum graminicolum (Ces.) G. W. Wilson. On leaves
of Calamagrostis longifolia. Port Wing.
Ascochyta pisi Lib. On Lathyrus palustris. Madison (Gil¬
bert & Davis).
170 Wisconsin Academy of Sciences, Arts, and Letters.
Ruhus triflorus should be added to the hosts of Septoria ruhi
West, in Wisconsin.
Septoria psilostega Ell. & Mart. On Galium horeale. Hay¬
ward.
Ramidaria reticulata Ell. & Evht. recorded in the provisional
list as a parasite of Osmorhiza Claytoni occurs on 0. longistylis
as well, as one would expect.
Cercosporella nivea Ell. & Barth. On Solidago uliginosa.
Brule. On Solidago juncea scaberrima. Lewis. The collections
have been referred to this species in spite of th^leaf spotting.
The following notes were made from examination of the collec¬
tion from Brule : Spots angular, white or yellowish white below
becoming yellow then reddish brown above, 1-3 mm. in diame¬
ter ; conidiophores more abundant below, fasciculate, 14-33 x 3 /u ;
conidia straight or curved, cylindrical to obclavate-cylindrical,
25-80 X 4 ju mostly 40-65 x 3 /t, becoming septate. On Solidago
juncea the spots become purple above. On one leaf of S.
uliginosa the coloring of the spots is reversed. In neither of the
collections are the conidia abundant.
Cercospora viciae Ell. & Hoi. On Latliyrus ochroleucus. Dan¬
bury.
Cercospora clavata (Ger.) Pk. On Asclepias tuherosa. Lewis.
The leaf parasite on Carpinus caroUniana recorded in
“Notes’’ II, p. 106 under the name Fusarium carpineum n. sp.
was collected on the same host at Balsam Lake in July 1924.
The cellular base from which the conidia spring is usually simp¬
ler in this collection and many of them do not suggest a sporo-
dochium. The spots are circular, about % cm. in diameter, scat¬
tered and not nervisequent. The spots are not brown until
death of the leaf cells takes place when by confluence irregular
brown areas may be formed. The conidia are not uniformly
biseptate but develop 1-4 septa. This was found also, in small
quantity, on Cary a cordiformis growing with infected Carpinus.
Puccinia graminis Pers. Uredinia on Poa annua collected at
Black Earth by McFarland & Davis are referred to this species.
The uredospores are 17-24 x 13-17 /i. This appears to be the
first collection that has been made on this species of Poa.
Davis — Parasitic Fungi in Wisconsin — XIII.
171
Additional Species
Mamiania fimbriata (Pers.) Ces. & DeNot. On Carpinus caro-
liniana. Danbury and Balsam Lake. The collections were made
in September and are not mature.
Sclerotinia geranii Seaver & Horne. On Geranium macula-
turn. Madison.
Phyllosticta negundinis Sacc. & Speg. has been collected at
several widely separated localities in Wisconsin. I see no
morphological distinction between this and Ph. minima (B. &
C.) E. &E.
Septoria flag ellif era Ell. & Evht. On Pisum sativum (cult.).
While this species appears to be common in Europe it has not
been previously found in America. This raises the question as
to whether it is indigenous or introduced. The clump of willows
on which it was abundant were close to a highway but it was
found later at Danbury on the same species of willow.
Septoria flagelUfera Ell. & Evht. On Pisum sativum (cult.).
Horicon (M. B. Linford). Apparently a species of northern
range. The type was from North Dakota and it occurs in
Manitoba.
While collecting at Bruce immature ascomata of Rhytisma
prini SchAv. were observed which appeared to have burst with
protrusion of the white content which constrasted strongly with
the black cortex. Ooing a little further ascomata of Rhytisma
salicinum (Pers.) Fr. were found having the same appearance.
On examination however it was found that the appearance was
due to acervuli on the “tar spots” Avhich had discharged snow
white masses of sporules.
Gloeosporium niveum n. sp. Acervuli under the cortex of
the Rhytisma stroma, plano-convex, varying in diameter up to
% mm.; conidia filling the acervulus, apparently produced by
abstriction of erect hyphae springing from the base, hyaline,
oblong with rounded ends to ovoid, continuous, 7-13 x 3/>t. On
immature ascomata of Rhytisma prini on leaves of Ilex verticel-
lata. Bruce, Wisconsin, September 4, 1924.
Acervuli similar to the preceding, conidia oblong to sub-
clavate, usually acute at one end, hyaline, continuous, 1-3 gut-
tulate, 13-20 x 5-6 y. On immature ascomata of Rhytisma
salicinum on leaves of Salix petiolaris. Bruce, Wisconsin,
172 Wisconsin Academy of Sciences, Arts, and Letters.
September 4, 1924. The disk of cytoplasm between the guttulae
often resembles a septum, especially when stained. This appears
to be distinct from Melasmia salicina Lev. which Tulasne con¬
sidered to be a spermogonial state of Rhytisma salicinnm and
which he described and figured in Carpologia III, p. 119, tab. XV,
fig. 15-17. The question as to whether these collections repre¬
sent one species or two is left open for the present.
[In 1925 numerous collections were made on various species
cf Salix agreeing with the description given above the conidia
being acute at one end suggesting a boat in their shape. No
collection was made on Ilex in 1925. The appearance of as-
eomata of Rhytisma nemopanthis Pk. sometimes suggested this
fungus but this was due to development of Ramularia nemo¬
panthis Pk. at the periphery of the ascomata.
The collection on Salix petiolaris is taken as the type of the
species. I am indebted to President Dearness for the informa¬
tion that this was issued in Fungi ColumMani, 2593 under the
name Tuber culina davisiana Sacc. & Trav. which belongs to a
very different plant.
GloeospoHiim niveum n. sp.
Left; Portion of a section of an immature ascoma of Rhytisma
prini and of an acervulus, ac, from which the conidia have fallen.
Upper right; 6 Conidia from Rhytisyna soMcinum. Lower right; 6
Conidia from Rhytisma prini.
Drawn with the aid of camera lucida by Lenette M. Rogers.
On leaves of Betula alba from Port Wing are large dead areas
on the upper surface of which are acervuli bearing fuligenous
sporules 9-10 x 6 on hyaline conidiophores. The fungus is
probably saprophytic and has been referred to Melanconium
bicolor Nees.
■Davis — Parasitic Fungi in Wisconsin — XIII.
173
Cylindrosporium triostei Kell. & Swingle. On Triosteum per-
foliatum. Haugen and Bruce. In the collection from Bruce the
acervuli are nearly all epiphyllous.
In May and June 1924 collections of a Mucedine on leaves of
Cynoglossum officinale were made at Madison and Cross Plains.
The following notes were made from these collections: Spots
dark brown, definite, circular to oval, 5-15 mm. in diameter ;
conidiophores mostly epiphyllous, fasciculate, hyaline, con¬
tinuous or septate, 30-60 x about 3 ft ; conidia hyaline, cylindri¬
cal to fusoid, mostly acute, straight, continuous or sometimes
with a more or less distinct median septum, catenulate, 10-30 x
2^-3 %ft. This is near Bamularia cynoglossi Lindr. from which
it differs especially in the slender acute conidia. It appears to
be conspecific with the parasite that occurs in Wisconsin on
leaves of Lappula virginiana which was recorded in Notes” I,
p. 89 under the name Ovularia asperifolii Sacc. var. Lappulae
Davis. It is perhaps better to consider the form that occurs in
Wisconsin on Lappula and Cynoglossum as a species distinct
from those that occur in Europe on related Boraginaceae and I
have, labeled these collections Ramularia lappulae (Davis) n.
comb.
Ramularia gracilipes n. sp. Spots orbicular to subquad-
rangular, dark purple above, lighter below, 5-10 mm. in diame¬
ter with a sharply delimited white central portion 1-3 mm. in
diameter; conidiophores hypophyllous on the central white por¬
tion, fasciculate from a prominent tubercular base, hyaline,
slender, usually straight, often septate, sometimes denticulate
below the apex, 40-75x2-3 ft; conidia hyaline, straight, cy¬
lindrical to fusoid-cylindrical, often more or less acute at one or
both ends, frequently with a median septum, 20-40 x 3% On
leaves of Cornus alternifolia. Bruce, Wisconsin, September 4th,
1924.
Ramularia artemisiae n. sp. Spots brown, of the width of
the leaf lobe and 3-5 mm. long the entire lobe usually becoming
dead and brown ; conidiophores on either surface, densely
fasciculate from a more or less prominent light brown base,
hyaline, erect or assurgent, lax, 35-80 x 1^-3 ^ ; conidia hyaline,
straight, cylindrical to fusoid, usually more or less acute, 0-3
septate, 12-30 x 2-4 fi. On leaves of Artemisia caudata or re¬
lated species. Lewis, Wisconsin, August 2, 1924.
174 Wisconsin Academy of ScienceSy ArtSy and Letters,
Cercoseptoria minuta n. sp. Spots pale, extending to the mid¬
rib, immarginate, 1/2-2 cm. in length ; fasciculi hypophyllous,
scattered, penicillate, composed of condia arising from a small,
apparently substomatal, tubercle or plexus, hyaline, straight or
somewhat curved, 50-60 xl (x. On leaves of Campanula apari-
noides, Wyeville, Wisconsin, July 7, 1923. The generic name is
used in the sense suggested in “Notes’^ VII, p. 401. A very
delicate species. Possibly a state of the parasite occurring on
this host that develops pycnidia and that was recorded in the
provisional list under the name Septoria campanulae (Lev.)
Sacc.
Because of a prior use of Septoriopsis as a generic name by
Fragoso and Paul, Petrak has proposed Cercoseptoria to replace
Septoriopsis Stevens & Dalbey. {Ann. My col. 23: 69).
Notes on DiSTRrouTiON and Abundance.
USTIL AGIN ALES
UsTiLAGO OSMUNDAE Pk. Not frequent or abundant. Collec¬
tions have been made in 5 localities. Two of them are in the
central portion of the state, the others in the northwestern por¬
tion where it develops more abundantly.
UsTiLAGO Zeae (Beckm.) Unger. Common and often abun¬
dant in com fields.
UsTiLAGO rabenhorstiana Kuehn. This appears to be rare
in Wisconsin but may be abundant where it occurs.
UsTiLAGO SPHAEROGENA Burr. This has been collected at
Millville and Madison only, in the southern part of the state.
It probably does not maintain itself in Wisconsin but is occasion¬
ally introduced.
UsTiLAGO Neglecta Niessl. Usually common and abundant.
UsTiLAGO STRiAEFORMis (West.)i NicssL Somc times abun¬
dant in meadows on Phleum pratense, less so on Agrostis alba.
There are two collections on Poa pratensis. All are from the
southern portion of the state. Elymus canadensis was given
as a host in the provisional list but I find no specimen on that
plant.
Davis — Parasitic Fungi in Wisconsin — XIII. 175
UsTiLAGO CALAMAGROSTiDis (Fckl.) Clint. But a single speci¬
men collected by A. B. Stout in southern Wisconsin.
UsTiLAGO AVENAE (Pcrs.) Jenscn.
UsTiLAGO LEVIS (Kell. & Sw.) Magn. The oat smuts are
found more or less abundantly throughout the state.
UsTiLAGO Perrennans Rostr. There is but one Wisconsin
specimen in the herbaria.
irsTiLAGO LONGissiMA (Sow.) Tul. Rather common on Glyceria
grandis throughout the state.
UsTiLAGO LONGISSIMA MACROSPORA Davis. The Variety oc¬
curred abundantly at the type station on Glyceria septentrionalis
(then known as G. fluitans) which was long since destroyed be¬
cause of the needs of agriculture. It is not known to be present
in the state at the present time. It has been collected in Canada
by Dearness and is reported to be distributed through central
Europe. Bauch has reported interesting differences in the ger¬
mination of the spores of the variety and of the type. (Zeitschr.
f. Bot. 15: 241-279 [1923]).
UsTiLAGO SPERMOPHORA B. & C. This has been seen in the
southern part of the state only where it is rare.
UsTiLAGO MACROSPORA Desm. Reported by Clinton as occur¬
ring in Wisconsin but it has not been seen by the writer nor are
there Wisconsin specimens at Madison.
UsTiLAGO TRiTici (Pers.) Rostr.
UsTiLAGO HORDEi (Pers.) Swingle.
UsTiLAGO NUDA (Jensen) Kell. & Sw. are more or less com¬
mon in grain fields in Wisconsin as elsewhere.
UsTiLAGO LORENTZiANA Thuems. This has been collected at
Madison but is rare.
UsTiLAGO OLivACEA (Dc.) Tul. This was collected in 1902 at
the outlet of Lost lake in Vilas county which abuts on the north¬
ern peninsula of Michigan. It has not been seen in the state
since.
UsTiLAGO UTRicuLOSA (Nees) Tul. Not a rare species in
Wisconsin and not restricted in range.
UsTiLAGO oxALiDis Ell. & Tracy. The collection by Tracy in
southwestern Wisconsin in the only one known to have been
made in the state.
176 Wisconsin Academy of ■ Sciences, ArtSj and Letters.
SiPHAGELOTHECA CRUENTA (Kuehn) Potter. This was col¬
lected at Madison by Trelease according to Potter.
Sphacelotheca sorghi (Lk.) Clint. This is sometimes abun¬
dant in sorghum fields.
Sphacelotheca hydropiperis (Schum.) DBy. Not uncom¬
mon. Most frequently seen in the northern half of the state.
Cintractia caricis (Pers.) Magn. Throughout the state.
Like most smuts this varies much in abundance. Some years it
is frequently observed and abundant Avhile in other seasons it
may not be seen at all.
Cintractia subinclusa (Koern.) Magn. Observed at but
one station which was in southeastern Wisconsin and where it
was fairly abundant.
Cintractia junci (Schw.) Trel. Usually abundant where it
occurs but in some years it is not seen. The collections are from
the southern portion of the state.
ScHizoNELLA MELANOGRAMMA (Dc.) Schroet. A coinmon and
abundant species in southern Wisconsin. There are no speci¬
mens from the northern part.
SoROSPORiUM PANici-MiLiACEi (Pers.). Takahashi. This was
collected at Madison in 1911 and 1912 and at Baraboo in 1918.
It was referred to the following species in the Provisional List.
. SoROSPORiUM SYNTHERiSMAE (Pk.) Earl. Common and
abundant where the host occurs on sandy lands in southern Wis¬
consin.
Tilletia foetens (B. & C.) Trel.
Tilletia tritici (Bjerk.) Wint.
With the revival of wheat growing in Wisconsin during the
world war bunt became common, usually the latter species.
Tuburcinia clintoniae Kom. Rare. The three localities
where it has been seen are in the southeastern, southern and
northwestern parts of Wisconsin. In each case the host is
Streptopus roseus the citation of Smilacina stellata being erron¬
eous.
Urocystis occulta (Rabh.) Wallr. This occurs oceasioiially
in rye fields but does little damage.
Urocystis agropyri (Preuss) Schroet. This occurs in south¬
ern Wisconsin on Elyirlus. A collection on Hordeum juhatum
was made by Toole and Bennet in northeastern Wisconsin.
Urocystis cepulae Frost. Occurs in onion fields in the south¬
eastern part of the state.
. Davis — Parasitic Fungi in Wisoonsin^XIII. 177
r Urocystis anemones (Pers.) Schroet. Common in the south¬
ern portion of the state.
Urocystis waldsteiniae Pk. Collected in two localities in
northern Wisconsin in 1893. It has not been seen since.
Entyloma lineatum (Cke.) Davis. Common and abun¬
dant, in the southern half of the state.
Entyloma crastophilum Sacc. This has been collected in
southern Wisconsin only except for the collection on Glyceria
pallida in the northern portion.
Entyloma parvum Davis. This inconspicuous species prob>
ably occurs throughout the state.
Entyloma nymphaeae (Cunn.) Setch. Throughout the state.
Sometimes abnudant, sometimes scarce.
Entyloma microsporum (Ung.) Schroet. Frequent in south¬
ern Wisconsin on Ranunculus septentrionalis only.
Entyloma ranunculi (Bon.) Schroet. Frequent and abun¬
dant in northern Wisconsin on Ranunculus pennslyvanicus only.
Entyloma thalictri Schroet. Generally distributed and
sometimes abundant.
Entyloma menispermi Farl. & Trel. Common in the south¬
ern half of the state.
Entyloma floerkeae Holw. This has been seen in southeast¬
ern Wisconsin only where it was locally abundant.
Entyloma saniculae Pk. A rare species in Wisconsin.
Collections have been made at Racine (1892 & ’94) and Madi¬
son (1924) only, the latter on Sanicula gregaria.
Entyloma australe Speg. Common and abundant through¬
out.
Entyloma linariae veronicae Wint. Usually common and
abundant at least in the southern portion.
Entyloma linariae gratiolae Davis. This has been found
in two localities in northern Wisconsin. As it has been but re¬
cently recognized it may be that it will be found to have a wider
distribution.
Entyloma lobeliae Farl. Variable in frequency and abun¬
dance from year to year.
Entyloma compositarum Farl. A common and abundant
species but some of the host adapted races of which it appears
to be composed are rather rare. It is most common on
Lepachys and Ambrosia, least so on Senecio. It is sometimes
12
178 Wisconsin Academy of Sciences, Arts, and Letters,
very abundant locally on Eupatorium urticaefolium while in
other years it is not seen on that host.
Entyloma polysporum (Pk.) Farl. Not uncommon on Am¬
brosia art emisiae folia and Rudheckia hirta, rare on other hosts,
hosts.
Burrillia pustulata Setch. Not frequent but usually abun¬
dant where it occurs.
Doassansia zizaniae Davis. This has been collected at Ra¬
cine and Madison only. As the spore balls are formed in
the central cavity of the lower internodes without external man¬
ifestation it is recognized only by splitting open the culms. It
appears late in the season and has not been looked for at that
time in the northern part of the state.
Doassansia martianoffiana (Thuem.) Schroet. Infre¬
quent. It has been collected only on plants emersed by the re¬
cession of the water.
Doassansia sagittariae (West.) Fisch. Throughout the
state.
Doassansia sagittariae confluens Davis. Also throughout
the state.
Doassansia opaca Setch. This has been seen in the southern
part of the state only where it is rare.
Doassansia intermedia Setch. This has been found in the
northern half of the state only. It is not frequent.
Doassansia deformans Setch. This is one of the commoner
species of the genius in Wisconsin and occurs throughout.
Doassansia furva Davis. This has been collected in several
localities none of which are in the southern portion. It is
usually fairly abundant where it occurs.
Doassansia alismatis (Nees) Cornu. This was formerly
the species most often seen but of late years it has not been so
frequent. A cycle of abundance will probably come again.
Doassansia ranunculina Davis. This was thought to have
disappeared from the state but it was found in very small
quantity, barely enough to show its presence, at Shiocton in
eastern Wisconsin and at Blue River in the southwestern part
of the state. It may therefore again become abundant when
conditions are favorable for successive years.
Davis — Farasitic Fungi in Wisconsin — XIII
179
Tracya lemae (Seteh.) Syd. This has been collected in
three widely separated localities. As it is very inconspicuous
it probably has not a restricted range in the state and occurs
more frequently than the collections would indicate.
COLEOSPORIUM SONCHI-ARVENSIS (PcrS.) Lev.
IjNIVERiSITY OF WISCONSIN HERBARIUM,
Madison, Wisconsin, March, 1925
/J
' M
NOTES ON PARASITIC FUNGI IN WISCONSIN— XIV
J. J. Davis
A large part of tlie collections of 1925 were made in the ^ ' cut¬
over ’ ’ regions in the northern part of the state. The cumulative
effect of successive dry seasons was in evidence. The effect of
the removal of the trees on the moisture content of soil and air
is marked in its relation to fungus parasites. It is with sur¬
prise that one observes as he vralks over dry ground the presence
of such plants as Iris, Caltha, Comarum, etc. To one who has
known the forest floor the difference is striking. The effect of
the dryness is especially evident in the later stages of develop¬
ment of the parasites. Leaf spots may be abundant but spore
bodies few, imperfect, and with little or no formation of normal
spores. A large proportion of the collections in this region were
discarded.
In ‘‘Notes” V it was stated that a trace of Bremia lactucae
Regel had been observed on Krigia amplexicaulis. In July 1925
it was found on this host at Altoona in sufficient quantity to
furnish a herbarium specimen. Apparently infection occurs
but seldom on this host. One might infer that this is not a race
adapted to Krigia but that infection occurs exceptionally.
In “Notes” IX, p. 251, reference was made to the occurrence
^ of mycelium and conidia supposed to be those of Sphaerotheca
humuli (DC.) Burr, on Ruhus allegheniensis at Madison early in
the season. In late July, 1925, similar material was collected at
Eau Claire on leaves of blackberry, species undetermined.
Very few developed conidia were present. Ruhus triflorus is
the only species of the genus that bears perithecia commonly or
abundantly in Wisconsin.
What may be the growth described and figured by Tulasne as
a spermatiophorous state {Melasmia salicina Lev. Carpologia
III, p. 119, tab. XV) has been rarely seen in the examination of
immature ascomata of Rhytisma salicintim (Pers.) Pr. In one
case instead of the cylindrical spermatia a small globular sper-
matium about 2fi in diameter was developed at the summit of
each of the slender, straight, congested conidiophores. Nothing
182 Wisconsin Academy of Sciences, Arts, and Letters.
comparable to the ‘^stylospores’^ described and figured by
Tulasne has been seen. As the covering of the spore body is
merely the cortex of the ascoma it does not seem to be properly
referred to Leptostromaceae. It is perhaps microconidial.
Phyllosticta fatiscens Pk. is evidently the same as Ph.
nupharis Allesch. {Phoma westendorpii Tosq. & West.) as that
species is represented in Petrak, Fungi Bohemiae et Moraviae,
series II, no. 1744. Peck’s name appears to have precedence in
time.
Septoria malvicola Ell. & Mart, is now recognized as the
proper designation of the species on Malva rotundifolia that was
recorded in the provisional list under the name Septoria
Jieterochroa Desm.
A collection made in June on leaves of a narrow-leaved Aster
{A. salicifoliusf) and referred to Septoria atropurpurea Pk.
bears sporules 33-66 x 2-3/a.
In a collection on leaves of Ambrosia trifida from HoUandale
(June 17, 1925) the spots are not white and arid but are
slightlj^ paler portions of larger brown spots. The pycnidia are
sometimes prominent and occasionally imperfect and even
acervulous. The sporules are 17-30 x I/a acute at one end. It
is believed to be a state of Septoria hacilligera Wint.
Because of previous use of Septoriopsis as a generic name by
Fragoso and Paul, Petrak has replaced Septoriopsis Stevens &
Dalbey as amended in ‘‘Notes” VII, p. 401, by Cercoseptoria
{Ann. My col. 23:68). The two Wisconsin species referred to
Septoriopsis therefore became Cercoseptoria longispora (Pk.)
Petr, and C. leptosperma (Pk.) Petr.
Bayliss — Elliott and Stanfield find the aseigerous stage of *
Polythrincium trifolii Kze. to be referable to Dothidella and
describe it as Dothidella trifolii n. sp. (Tram. Brit. Myc. Soc.
9:227 [1924]). They also observed a pycnidial state in au¬
tumn.
In transferring the description of Cercospora pontederiae Ell.
& Dearn. to the Sylloge Fungorum, Saccardo queried if it were
not a Cercosporella. The query seems justified and Wisconsin
specimens are labeled Cercosporella pontederiae (Ell. & Dearn.).
The development is sometimes amphigenous and the conidia
longer (up to 75/a) than indicated by the description. It has
been found in Wisconsin only in the northern part.
In examining a collection of the fungus referred to Cercospora
euonymi Ellis made in late September, conidia were observed up
Davis — Parasitic Fungi in Wisconsin — XIV. 183
to 140/a in length. The conidiophores are hyaline as in previous
Wisconsin collections.
For the reception of Gloeosporium halsameae Davis Notes*'
YII, p. 409, pi. 30) Petrak has proposed the genus Rhabdo-
gloeopsis {Ann. My col. 23:52). There are those who would not
consider the characters to be of generic value and who deprecate
the multiplication of genera founded upon slight differences as a
disservice to mycology. This is especially true where life
histories are unknown. To such Cryptocline, Rhabdogloeum
and Rhabdogloeopsis would constitute a subgeneric group.
The parasite recorded in the supplementary and provisional
lists under the name Cylindrosporium thalictri (E. & E.) is the
one described in the Journal of Mycology 3: 130 as C. ranunculi
(Bon.) /. thalictri E. & E. It bears no relation to C. ranunculi
(Bon.) Sacc. or to Entyloma and is quite distinct from C.
clematidis E. & E. The acervuli are usually confined to a
sharply delimited angular portion of the arid spot.
The parasite of Carpinus caroliniana recorded in “Notes’* II,
p. 106 as Fusarium carpineum n. sp. was found on the same host
at Balsam Lake in July, 1924. The cellular base from which
the conidia spring is usually simpler in these collections and
many of them do not suggest a sporodochium. The spots are
circular, scattered, not nervisequent, about % cm. in diameter.
They are not brown until death of the leaf cells takes place when
by confluence irregular brown areas may be formed. The
conidia are not uniformly biseptate but develop 1-4 septa. This
was found also, in small quantity, on Carya cordiformis grow¬
ing with infected Carpinus. It was noted that on this host the
conidia are smaller, 20-30 x 1%-2/a and not septate. Perhaps
this might better be placed in Cercoseptoria as that genus has
been treated in these notes under Septoriopsis.
In 1925 Entyloma on Thalictrum dasycarpum bearing conidia
was collected at Armstrong Creek and at Tripoli. As it oc¬
curred in the latter locality a hyphal mat was formed on the
lower surface of the spots much as in E. menispermi Farl. &
Trel. There seems to be no good reason for attempting to keep
E. ranunculi (Bon.) Schroet. and E. thalictri Schroet. separate,
although they may be distinct in their host relations as is the
way with smuts. On Ranunculus Entyloma has been found in
Wisconsin on R. pennsylvanicus only but it is not infrequent on
that species, especially northward, 10 collections being rep re-
184 Wisconmi Academy of Sciences, Arts, and Letters.
sented in the herbaria. It was found on Anemone quinquefolia
at Racine in 1888 but has not been seen on that host since and
I have seen no record of its occurrence elsewhere.
Collections of leaves of Sagittaria heterophylla have been
made that bear both Doassansia sagittariae confluens Davis and
D. furva Davis the spore balls of both being sometimes present
in the same section.
Uredinopsis atkinsonii Magn. is united with U. copelandi Syd.
by Arthur in North American Flora 7 : 684,
Potentilla canadensis has been collected at Madison bearing
Phragmidium and Darluca and in addition black sclerotioid
bodies. This seems to be of the same character as the growth
which has been called Sphaeria solidaginis Schw.
While TJredo muelleri Schroet. is considered to be aecial in
character it is sometimes attacked by Darluca filum (Biv.) Cast,,
a common parasite of uredinia but not of aecia.
In June 1913 Aecidium falcatae Arth. occurred at Wyalusing
in southwestern Wisconsin near the Wisconsin river. In look¬
ing about for a possible alternate host a coarse grass was found
that had borne telia the preceding year. This was probably
Andropogon furcatus. In 1920 another collection of the
Aecidium was made at Prairie du Chien in the same region. In
June 1924 it was found in abundance at Blue River in south¬
western Wisconsin and in August at Danbury in the north¬
western part of the state. The stations were noted and later
in the season both localities were visited for the purpose of
getting a clue to the alternate host. In each case Andropogon
scoparius and A. furcatus were the only rusted plants in the im¬
mediate vicinity. In the spring of 1925 development of aecia on
Amphicarpa monoica was secured in the greenhouse from in¬
fection from telia on Andropogon. Some years ago Aecidium
lupini Pk. was observed to be abundant at Millston. The
stations were noted and visited later in the year and Andropogon
found to be the rusted host that seemed to be associated with it
but no infection experiments were made. In North American
Flora 7 : 625-6, Arthur has united Aecidium falcatae Arth. Ae.
lupini Pk. and described species on other genera of Leguminosae.
It seems probable that there is a rust on Andropogon that de¬
velops on various Leguminosae. How much differential host
adaptation there is remains to be seen. Andropogon seems to
be a complacent host for rusts.
Davis — Parasitic Fungi in Wisconsin — XIY. 185
In “Notes” XI, pp. 301-2, reference was made to the
occurrence of Puccinia punctiformis Diet. & Hoi. in Wiscon¬
sin with the suggestion that it was ‘ ‘ presumably a waif in Wis¬
consin and perhaps will not be able to maintain itself in this
climate”. In 1925 uredinia were found beside the railroad at
Madison about a mile from where it had been noticed in 1922.
Additional Hosts
Microsphaera alni (Wallr.) Wint. On Bhamnus alnifolia,
Pembine.
Ascochyta thaspii saniculae Davis. On Sanicula gregaria.
Argyle.
Melasmia ulmicola B. & C. On TJlmus racemosa. Ingram.
Collet otrichum graminicolum (Ces.) Wils. On Bromus pu-
gans. Balsam Lake.
What appears to be an imperfectly developed state of
Marssonina kriegeriana (Bres.) Magn. was collected at Argonne
on Salix halsamifera.
Cylindrosporinm triostei Kell. & Swingle. On Triosteum
aurantiacum. Spring Valley. In this collection the acervuJi
are epiphyllous.
Microstroma juglandis (Bereng.) Sacc. On Cary a cordi-
formis. Balsam Lake.
Ramularia vaccinii Pk. was collected on Vaccinium penn-
sylvanicwm at Wausaukee in 1913 but not recorded. It was
found on V. canadense at Ogema in 1925.
Cercospora dubia (Riess) Wint. On Chenopodium hyhridum.
Danbury.
Puccinia gentianae (Strauss) Lk. Uredinia and telia on
Gentiana puberula. Taylor.
Puccinia patruelis Arth. Aecia on Lactuca villosa and
Hieracium canadense. Hollandale.
Additional Species
A collection on living leaves of Betula pumila from Danbury
corresponds with immature specimens of Dotkidella hetulina
(Pers.) Sacc. This was referred to Phyllachora by Fuckel and
to Euryachora by Schroeter and by Theissen & Sydow.
In August 1920 a trace of Taphrina struthiopteridis Nishida
was found at Spooner on Onoclea Struthiopteris. In late July
1925 it was again found at Weyerhaeuser in sufficient quantity
186 Wisconsin Academy of. Sciences, Arts, and Letters.
to furnish a herbarium specimen and warrant recording. The
infected pinnae resemble those attacked by Uredinopsis.
At Millston in September 1924 many leaves of Rubus hispidus
showed orbicular dead spots, cinereous with a purple border,
about 5 mm. in diameter. In the center of the spots on the
upper surface were remains of old sori of TJredo muelleri while
on the lower surface were scattered pyenidia of Coniothyrium
fuckelii Sacc. Presumably the latter had developed on tissue
that had been injured by the rust.
Ascochyta baptisiae n. sp.
Spots suborbicular, dull black, more or less argillaceous zoned,
3-6 mm. long, nearly alike on both surfaces; pyenidia epi-
phyllous, somewhat prominent, argillaceous, subepidermal,
globose, ostiolate, 85-150/j, in diameter ; sporules hyaline,
cylindrical with rounded ends, straight or nearly so, uniseptate
or occasionally with 2 or 3 septa, 12-30 x 4-7/x. On leaves of
Baptisia leucantha. Avoca, Wisconsin, October 8, 1924.
Stagonospora petasitidis Ell. & Ev.
On Petasites palmatus. Tripoli. In this collection, made in
mid- August, the areas over which the pyenidia are scattered are
mostly brown and dead. On these dead areas the pyenidia are,
for the most part, empty or contain but few more or less im¬
perfect sporules.
In late August and early September collections were made in
several localities in northeastern Wisconsin of a parasite of
Popidus balsamifera. On examination these showed numerous
epiphyllous pyenidia on indefinitely limited brown spots in
which however sporules had not developed. I am indebted to
President Dearness for an opportunity to examine a portion of
the type of PJiyllosticta briinnea Dearn. & Barth, on leaves of
Populus angustifolia from Colorado and for the suggestion that
the Wisconsin collections are of that species, a suggestion that is
borne out by the comparison. Material collected later in the
season would perhaps have shown further development.
Graphium sorhi Pk.
On Pyrus americana. Armstrong Creek. In this collection
the synnemata are few and the conidia immature, corresponding
to the rule in collections of the year 1925.
Puccinia ambigua (A. & S.) Lagh.
Aecia and telia on Galium Aparine. Spring Talley.
Davis — Parasitic Fungi in Wiscons'in — XIV,
187
Coleosporium kelianthi (Schw.) Arth.
Uredinia and telia on Helianthus strumosus. A single collec¬
tion made at Madison.
Notes on Distribution and Abundance in
Wisconsin
Melampsoraceae
Including Coleosporium
I\1elampsora farlowii (Arth.) Davis.
This has been observed in the northeastern corner of the state
only.
Melampsora bigelowii Thuem.
Common and abundant throughout the state.
Melampsora humboldtiana Speg.
This has been so confused with the preceding species that its
range or abundance is not well known. The aecial stage has
been collected in the northeastern and northwestern sections
only.
Melampsora medusae Thuem.
Common and abundant.
Melampsora populi tsugae Davis (M. Abietis-canadensis
(Farl.) C. A. Ludwig)
Known only from the north eastern part of the state accom¬
panying M. farlowii (Arth.) Davis.
Melampsora lini (Pers.) Desm.
Occurs in flax fields and also on Linum sulcatum on which it
is sometimes abundant in the western part of the state.
Melampsora cerastii (H. Mart.) Schroet.
Aecia were formerly rather common in northern Wisconsin
but are now rare according to my experience. Uredinia or
telia have not been recognized in Wisconsin.
Melampsoridium betulinum (Pers.) Kleb.
Apparently a rare species in Wisconsin. But one collection
of aecia has been made, but they were abundant at that station
which was in the northwestern part of the state. Two collec¬
tions of uredinia have been made in the southern part of the
state and one in the northern.
Calyptospora goeppertiana Kuehn.
Very rare in Wisconsin. But a single collection.
188 Wisconsin Academy of Sciences, Arts, and Letters.
Chrysomyxa Pirolae (Dc.) Rostr.
Occurs throughout the state.
Chrysomyxa ledicola (Pk.) Lagh.
The tejia of this and cognate species are formed so early in the
season that they disappear before the localities are visited. One
collection of telia referred to this species was made in north¬
eastern Wisconsin. Uredinia have not been seen. A few
collections of aecia have been made in the northern part of the
state.
Chrysomyxa ledi (A. & S.) DBy.
Common northward. Telia have not been seen. Aecia have
not been recognized ; perhaps they have been confused with those
of the next species.
Chrysomyxa cassandrae (Pk. & Cl.) Tranz.
Common and abundant in the spruce bogs but telia have not
been collected. Aecia on Picea mariana are common and
abundant.
Chrysomyxa chiogenis Diet.
A rare species collected only in a somewhat limited region in
the northern part of the state. No collection has been made
since 1903. Moist, mossy, well shaded logs with Chiogenes
trailing over them are not abundant since deforestation and
forest fires.
Chrysomyxa chiogenis Diet.
Of still more restricted range in the northern part of the
state.
Pucciniastrum pustulatum (Pers.) Diet.
Common and abundant on the larger species of Epilobium.
The telia are most abundant on EpHohium angustifolium.
Pucciniastrum pyrolae (Pers.) Diet.
The uredinia occur on various species of Pyrola throughout
the state except that there are no collections from the south¬
eastern portion. But one collection has been made on Chima-
phlia. Aecia and telia have not been seen.
Pucciniastrum myrtilli (Schum.) Arth.
Aecia are sometimes abundant. Urelinia are usually sparsely
developed. Telia have been seen in two collections. Because
of the intracellular position of the teliospores, this is considered
to be generically distinct by some authors who designate it
Thecopsora.
Davis — Parasitic Fungi in Wisconsin — XIV.
189
Hyalopsora aspidiotus Pk.
This has been collected in the extreme northeastern portion of
the state only.
Hyalopsora polypodii (Dc.) Magn.
Common in southern and eastern Wisconsin.
Uredinopsis phegopteridis Arth.
The least common of the species of the genus in Wisconsin.
It occurs in the northern part of the state only.
Uredinopsis copelandi Syd. fU. atkinsonii Magn.)
Not uncommon but often poorly developed apparently from
early death of the infected tissue.
Uredinopsis mirabilis (Pk.) Magn.
The most common and abundant species of the genus often at
long distance from the aecial host.
Uredinopsis struthiopteridis Stoermer.
Common and abundant northward.
Uredinopsis osmundae Magn.
Common and abundant especially northward. The aecial
stage of the species of this genus, Peridermium halsameum Pk.,
is common where the host occurs but no segregation has been
made. All of them develop teliospores, the first and last most
abundantl3L
Pucciniastrum potentillae Kom.
With the range of the host which is northwestern but extend¬
ing south considerably below the middle. No telia have been
found.
Pucciniastrum americanum (Pari.) Arth.
Throughout the state on the single species of host Rubus
idaem aculeatissimus. Telia are frequent.
Pucciniastrum arcticum (Lagh.) Tranz.
Throughout the state on Rubus triflorus only. No telia have
been seen. It is more abundant northward.
Pucciniastrum agrimoniae (Schw.) Tranz.
Common and abundant throughout. No telia seen.
Cronartium comptoniae Arth.
All of the stages occur somewhat abundantly in northern Wis¬
consin.
Cronartium quercus (Brondeau) Schroet.
A common species throughout the range of the aecial host,
Pinus Banksiana. The telia occur most frequently and
abundantly on Quercus velutina.
190 Wisconsin Academy of Sciences, Arts, and Letters.
Cronartium comandrae Pk.
The aecial stage appears to be but sparsely developed but the
later stages are much more common and abundant, often at
long distance from the aecial host.
Cronartium ribicola Fisch.
Still occurs to some extent in northwestern Wisconsin.
Cronartium occidentale Hedge, Bethel & Hunt.
Reported as having been collected on Ribes at two stations in
northern Wisconsin in the course of the white pine blister rust
survey.
Cerotelium urticastri Mains.
Probably of wide range in the state. Aecia have been collect¬
ed in the southern part only probably because more northern
territory has not been visited early enough in the season.
COLEOSPORIUM RIBICOLA (C. & E.) Arth.
Collected in northern Wisconsin in 1918 in the course of the
white pine blister rust survey. I have not seen it myself in the
field.
CoLEOSPORiuM viBURNi Arth.
Not uncommon and sometimes rather abundant in the
southern and eastern portions of the state.
COLEOSPORIUM CAMPANULAE (Pres.) Lev.
This species was included in the state flora on the basis of a
fragmentary specimen on Campanula rotundifolia bearing
uredinia collected at Prairie du Sac in 1891 by H. F. Lueders.
By some error the host was given in the list as C. americana.
Nothing further Avas known of the occurrence of this rust in the
state until 1922 when uredinia were found on Campanula
aparinoides at Blue River in the south western part of the state
and in 1923 on the lake Superior shore.
COLEOSPORIUM soLiDAGiNis (Schw.) Thuem.
Common and abundant throughout the state. But few
collections of aecia have been made.
COLEOSPORIUM HELIANTHI (Schw.) Arth.
The first and only collection was made at Madison in 1925.
TJiis has been found in but a single locality in northeastern
COLEOSPORIUM SONCHI-ARVENSIS (PcrS.) LcV.
Wisconsin.
University of Wisconsin Herbarium.
Madison, Wisconsin, April, 1926.
Ind^x to ^^Notes^^ XII, XIII, XIV.
191
INDEX TO HOSTS AND PARASITES
Referred to in “Notes” XII, XIII, and XIV.
The names of parasites are in italics.
Abies balsamea, 169
Actaea rubra neglecta, 160
Aecidium falcatae Arth., 184
Aeddium lupini Pk., 184
Aecidium myosotidis Burr., 162
Agrimonia gryposepala, 158
Albugo Candida (Pers.) Kuntze, 158
Alnus crispa, 169
Ambrosia trifida, 182
Amphicarpa monoica, 184
Anaphalis margaritacea, 161
Andropogon furcatus, 184
Andropogon scoparius, 184
Anemone quinquefolia, 184
Antirrhinum majus, 162
Arab is laevigata, 158
Arctium minus, 161
Aristida tuberculosa, 162
Artemisia biennis, 158
Artemisia caudata, 173
Artemisia serrata, 155
Artemisia ludoviciana, 155
Asclipias tuber os^ 179
Ascochyta baptisiae n. sp. 186
Ascochyta compositarum Davis, 159
Ascochyta pisi Lib., 169
Ascochyta thaspii E. & E., 159
Ascochyta thaspii saniculae Davis, 185
Aster azurens, 160
Aster paniculatus, 160
Aster sagittifolius, 160
Aster Tradescanti, 155, 160
Asterina rubicola, 165
Asteroma atratum Chev., 161
Asteroma lactucae, 161
Asteromella, 156
Baptisia leucantha, 186
Barbarea stricta, 160
Betula alba, 172
Betula pumila, 185
Bremia lactucae. Regel 181
Bromus purgans, 185
Calamagrostis canadensis. 166
Calamagrostis longifolia, 169
Campanula aparinoides, 174
Carpinus caroliniana, 170, 171, 183
Carya cordiformis, 170, 185
Cassia marylandica, 162
Cerastium nutans, 155
Cercoseptoria leptosperma (Pk.) Petr., 182
Cercoseptoria longispofa (Pk. ) Petr., 182
Cercoseptoria niinuta n. sp., 174
Cercospora clavata (Ger.) Pk., 170
Cercospora dubia (Riess) Wint., 185
Cercospora euonymi Ellis, 182
Cercospora helvola, Sacc., 160
Cercospora leptandrae n. sp., 162
Cercospora oculata E. & K., 157
Cercospora perfoliata E. & E.. 160
Cercospora platyspora E. & H., 167
Cercospora pontederiae E. & Dearn., 182
Cercospora sedoides E. & E., 157
Cercospora sii E. & E., 168
Cercospora stmulata E. & E., 162
Cercospora suhsanguinea B. & E., 168
Cercospora varia Pk., 166
Cercospora vernoniae E. & K., 157
Cercospora viciae Ell. & Hoi., 157, 170
Cercospora sebrina Pass., 160
Cercosporella nivea E. & Barth., 170
Cercosporella pontederiae (Ell. & Dearn.) n.
comb., 182
Cercosporidium fasdculatum (C. & E. )
Hoehn,, 168
Chenopodium hybridum, 185
Cicuta maculata, 159, 161
Cinna latifolia, 159, 160
Cladosporium pfiragmitis Opiz, 168
Coccochora ruM Davis, 165
Coleosporium helianthi (Schw.) Arth. 187
Collet otrichum fusarioides (O’Gara) Dearn.,
169
Collet otrichum graminicolum (Ces.) Wils.
169, 185
Colletotrichum salmonicolor O’Gara, 169
Colletotrichum violarum Davis, 169
Coniothyrium fuckelii Sacc., 186
Cornus alternifolia, 173
Cornus canadensis, 167
Corylus rostrata, 165
Cylindrosporella, 166
Cylindrosporium betulae Davis, 169
Cylindrosporium salicifoliae (Trel. ) Davis,
157
Cylindrosporium thalictri (E. & E. ), 183
Cylindrosporium triostei Kell. & Sw. 162, 173,
185
Cylindrosporium vermiforme Davis, 169
Cynoglossum officinale, 173
Cytodiplospora elymina Davis, 166
Darluca filum (Biv.) Cast., 184
Davisiella, 166
Diplocarpon, 165
Doassansia furva Davis, 184
Doassansia sagittariae confluens^ 184
Doassansia zizaniae Davis, 157
Dothidella betulina (Pers.) Sacc., 186
Dothidella trifolii B. E. Sc S., 182
Echinochloa crusgalli, 163
Elymus, 157, 166
Endostigme, 155
Entyloma compositarum Pari., 160.
Entyloma Ixnariae gratiolae Davis, 169
Entyloma ranunculi (Bon.) Schroet., 188
Entyloma thalictri Schroet., 183
Erechtites hieracifolia, 161
Eupatorium purpureum, 159, 160
Fusarium. carpineum Davis, 170, 183
Fusicladium depressum (B. & Br. ) Sacc. 167
Fusicladium punctiforme Wint., 167
Fusicladium 155, 167
Galium aparine, 186
Galium boreale, 170
Gentiana puberula, 185
Geranium Bicknellii, 169
Geranium maculatum, 171
Geum canadense, 159
Geum strictum, 159
Geum virginianum, 159
Gloeosporium balsameae Davis, 183
Gloeosporium betulae-papyriferae “Dearn. &
Overholts, 169
Gloeosporium coryli (Desm. ) Sacc., 165
Gloeosporium equiseti E. & E., 168
Gloeosporium fusarioides E. & K., 169
Gloeosporium niveum n. sp., 171
Gloeosporium, septorioides Sacc., 159
Glomerella cingulata (Stonem.) Sp. & Von
Schr., 169
Glomerularia comi Pk., 167
Glomerularia lonicerae Dearn. & House, 167
Graphium, sorbi Pk., 186
Helianthus strumosus, 187
Heterosporium grcucile Sacc., 165
Hieracium canadense, 185
Hypodermella ampla (Davis) Dearn., 166
Hystrix, 157
Ilex verticillata, 171
Krigia amplexicaxilis, 181
Lactuoa canadensis, 161
Lactuca villosa, 185
Lappula virginiana. 173
Lathyrus ochroleucus, 170
Lathyrus palustris. 169
Lathyrus venosus, 157
Leersia lenticularis, 162
Leersia oryzoides, 162
Leersia virginica, 162
Lepachys pinnata, 161
Leptothyrium dryinum Sacc., 156
Leptothyrium maculicolum Wint., 156
Lonicera canadensis, 167
Lonicera oblongifolia. 167
Lophodermium amplum Davis, 166
Ludvigia polycarpa, 162
Lycopus americanus, 159
192 ^¥ isconsin Academy of Science, Arts, a7id Letters.
Lycopus uniflorus, 159, 160
Malva rotund if olia, 182
Mamiania Jinihriata (Pers. ) Ces. & DeN., 171
Marssonina kneyeriana (Bres.) Magn., 185
Marssonina martini (S. & E.) Magn., 159
Marssonirui potentUlae tormentillae Trail,
159
Melampyrum lineare, 162
Melampsora caryophyllaeearum, 169
Melanconium bicolor Nees, 3 72
Melasmia salicina Lev., 181
Melasmia ulmicola B. & C., 185
Microsphaera alni (Wallr.) Wint., 185
Microstroma jugiandis (Bereng. ) Sacc., 185
Mollisia earliana Sacc., 165
Montagnella heliopsidis (Schw.) Sacc., 155
Mycosphaei'ella lethalis Stone, 165
Mycosphaerella pinodes (B. & Blox. ) Niessl.,
165
Myosotis virginica, 159, 162
Oenothera pumila, 160
Onoclea Struthiopteris, 185
Osmorhiza longistylis, 170
Ovularia asperifolii lappulae Davis, 173
Ovularia decipiens Saco., 162
Passalorat depressa (B, & Br.) Hoehn, 167
Passalora fasciculata (C. & E.) Earle, 168
Passalora punctiformis (Wint.) Hoehn, 167
Penthorum sedoides, 157
PetHdermium coJoradense (Diet.) Arth. &
Kern., 169
Peridermium elatinum A. & S., 169
Peronospora alsinearum Gasp., 155
Peronospora leptosperma DBy., 155
Peronospora myosotidis DBy., 158
Peronospora parasitica (Pers.) Tul., 158
Peronospora potentUlae DBy, 158
Peronospora Sordida B. & Br., 169
Peronospora tomentosa Pckl., 155
Petasites palmatus, 186
Phacidium taxi Fr., 165
Phleospora salicifoline (Trel.) Petr., 157
Phyllachora 159, 166
Phyllachora graminis (Pers.) Pckl., 157
Phyllosticta hrunnea Dearn. & Barth., 186
Phyllosticta fatiscens Pk., 182
Phyllosticta lappae Sacc., TBl
Phyllosticta neyundinis Sacc. & Speg., 171
Phyllosticta nnpJtaris Allesch,, 182
Phyllosticta phomiformis Sacc., 156
Phyllosticta renouana Sacc. & Roum., 166
Phyllosticta steironematis Dearn. & House,
161
Phyllosticta typhina Sacc. & Malb., 166
Picea mariana, 169
Pigpotia fraxini B. & C., 156
Piricularia parasiMca E. & E., 157
Pisum sativum, 171
Plasmopara geranii (Pk.) Berl. & DeB., 169
Plasmopara nivea (Ung, ) Schroet., 161
Poa annua, 170
Polygonum, 155
Polygonum cilinode, 156
Polythrincium trifolii Kze., 182
Populus balsamifera, 186
Potentilla canadensis, 159, 160, 184
Puccinia amhigua (A. & S.), Lagh., 186
Puccinia angustcita Pk., 160
Puccinia antiri'hini D. & H.. 162
Puccinia flaccida B. ^ Br.. 162
Puccinia gentianae (Strauss) Lk., 185
Puccinia graminis Pers., 160, 170
Puccinia jussiaeoe Speg., 162
Puccinia pattruelis Arth., 185
Puccinia peckii (DeT.) Kell.. 160
Puccinia punctifomnis D. & H., 185
Pyrus americana, 186
Quercus alba, 156. 159
Ram, ul aria actoeae E. & H.. 160
Ramulaxia aequivoca (Ces.) Sacc., 160
Pamularia arvensis Sacc., 160
Ramularia artemisiax n. sp., 173
Ramularia asteris (Phil. & Plowr.) Bubak,
160
Ramularia harhareae (Pk.), 160
Ramularia cynoplossi Lindr., 173
Ramularia pracilipes n. sp.. 173
Ramularia lappulae n. comb., 173
Ramularia lysimadhiae Thuem., 156
Ramularia magnusiana (Sacc.) Lindau, 167
Ramularia melampyri Ell. & Dearn., 162
Ramularia pratensis Sacc., 160
Ramularia reticulata E. & E., 170
Ramularia rubicunda Bres., 168
Ramularia vaccinii Pk., 185
Ramularia waldsteiniae Ell. & Davis, 167
Ranunculus acris, 160, 162
Ranunculus pennsylvanicus, 183
Rhamnus alnifolia, 185
Rhytisma prini Schw., 171
Rhytisma salicinum (Pers.) Fr. 171, 181
Rosenscheldia, 155
Rubus allegheniensis, 181
Rubus hispidus, 186
Rubus trifiorus, 170, 181
Rudbeckia laciniata, 160
Rumex persicarioides, 160
Sacidium, 166
Sagittaria heterophylla, 184
Salix balsamifera, 185
Salix humilis, 160, 171
Salix petiolaris, 171
Sanicula gregaria, 185
Sclerotinia geranii Seaver & Horne, 171
Soolecotrichum graminis Pckl., 168
Scolecotrichum maculicola E. &. K., 168
Scrophularia leporella, 169
Septogloeum salicinum (Pk. ) Sacc., 160
Septoria atropurpurea Pk., 182
Septoria bacilligera Wint., 182
Septoria detospora Sacc.. 168
Septoria diervillae B. & E., 156
Septoria diervillae Pk., 156
Septoria diervillicola E. & E., 156
Septoria ellisii Berl. & Vogl., 167
Septoria erechtitis E. & E., 161
Septoria flagellifera E. & E.. 171
Septoria gei Rob. & Desm., 159
Septoria gratiolae Sacc. & Speg.. 167
Septoria heterochroa Desm., 182
Septoria infuscata Wint., 161
Septoria lycopi Pass., 159
Septoria malvicola E. «& M., 182
Septoria margaritoiceae Pk., 161
SeptoHa microsperma Pk., 166
Septoria polygonorum Desm., 155
Septoria psilostega E. & M., 170
Septoria rubi West., 170
Septoria salicicola (Fr.) Sacc., 171
Septoria violae West., 159
Septoriopsis Stevens & Dalbey, 182
Sisymbrium altissimum, 158
Solidago juncea, 170
Solidago uliginosa, 170
Sphaerotheca humuH (DC.) Burr., 181
Spiraea densiflora, 157
Spiraea salicifolia, 157
Sfagonospora petasitidis E. & E., 186
Steironema ciliatum, 156, 161
Stigmatea rubicola (E. & E.) Theiss, 165
Synchyfrium aureum Schroet., 158
Synchytrium nigrescens Davis, 158
Taenidia, 167
Tahprina stimthiopteridis . 185
Taphrina ulmi (Pckl.) Johans., 155
Taxus canadensis. 165
Thalictrum dasyoarpum, 3 83
Tllletia rorona, Scribn., 162
Tltaeospora detospora (Sacc.) Bubak, 168
Trientalis americana, 167
Trifolium dubium, 160
Triosteum aurantiacum, 185
Triosteum perfoliatum 162, 173
Ulmus americana, 155
Ulmus racemosa, 185
Uredinopsis atkinsonii Magn.. 184
T'redinopsis copelandi Syd., 184
Uredo mnelleri Schroet., 184
Uromyaes seditiosus Kern., 162
Ustilago sphaerogenm Burr., 162
Vaccinium canadense, 185
Vaccinium pennsylvanicum, 185
Vermicularia violae-rot%indifoliae (Sacc.)
House, 168
Veronica virginica, 162
Viburnum pubescens, 166
Viola lanceolata, 159
Viola scabriuscnla, 168
NOMENCLATORIAL NOTES ON AMERICAN FRESH
WATER MODLUlSCA*'
Frank Collins Baker
During the preparation of a monograph of the fresh water
mollusks of Wisconsin for the Wisconsin Geological and Natural
History Survey, certain new genera and other groups have been
observed and some changes in current nomenclature have been
noted. As the publication of this monograph will be somewhat
delayed it has seemed advisable to publish the salient points in
nomenclature brought out by the investigation, as well as certain
notes concerning other matters of interest observed. A full dis¬
cussion, with illustrations of the anatomical features, will ap¬
pear in the monograph mentioned.
Pleuroceridae
Since the time of William Stimpson (1864) apparently no ad¬
ditional observations have been made concerning the genitalia
of the snails belonging to this family. Half a hundred speci¬
mens of Goniohasis Uvescens and Pleurocera acuta have been ex¬
amined without finding any notable features not recorded by
Stimpson. There is no external organ of generation and there
appears to be no external feature for determining the sexes, ex¬
cepting the pit or sinus on the neck between the right tentacle
and the operculigerous lobe, which appears to be present only in
the female. There is a long canal on the right side of the rectum,
formed by two narrow laminae, about as broad as the rectum,
placed close together. This canal opens into the mantle cavity at
the angle formed by the mantle and body wall. Posteriorly the
two laminae become a single tube which enters the gonads
(ovaries or testes) which lie near the stomach. The study of
this family is worthy of serious attention, the anatomical
features offering novelties of interest and value.
•Contribution from the Museum of Natural History, University of Illinois,
No. 35.
13
194 Wisconsin Academy of Sciences, Arts, and Letters.
VlVlPARIDAE
Many years ago, Gill divided the Viviparidae into two groups
which he later distinguished as subfamilies, the distinctions
being based on the difference in the lingual teeth. The genitalia
also give additional characters and the division seems to be a
natural one. They are redefined below.
Subfamily Viviparinae Gill, 1871
Shell usually large with rounded or carinated w^horls; oper-
ulum wholly concentric, the inner margin folded in some
groups; foot not much longer than shell, not extending beyond
the tentacles; cervical lappets large, forming tubular conduits
for respiratory purposes, the right lappet the larger; radula
with broad central tooth, the reflection broad and multicuspid,
the center cusp usually wide and blunt; lateral and marginal
teeth longer than wide, the reflections wide and multicuspid;
penis forming an elongated, thick, sausage-shaped sac, behind
which the prostate is placed, without a long intervening vas de¬
ferens, the latter being short and narrow.
This subfamily includes the typical genera, Viviparus,
Tulotoma, Taia, Margarya, and perhaps some other groups
which agree in genitalia, radula, and general form.
Subfamily Lioplacinae Gill, 1871
Shell turreted, whorls rounded or subcarinated ; aperture sub-
angulated, sinuous or incurved at the base; operculum wholly
concentric or with subspiral nucleus; cervical lappets small, not
forming tubular conduits for respiratory purposes; foot very
large, truncated before, rounded behind; radula with narrower
teeth than in the Viviparinae, the reflection with sharp cusps,
none being wide or blunt, the marginals very long and narrow,
wide at the base; penis forming a long, narrow, more or less
convoluted tube opening at the end of the right tentacle, which
is larger than the left tentacle ; prostate placed immediately be¬
hind the penis (Lioplax), or midway of the vas deferens (Cam-
peloma), either sac-like (Campeloma), or made up of several
large convolutions (Lioplax).
Baker — Aimrican Fresh Water Mollusca.
195
The narrow, sharp-pointed cusps of the radula, the large foot,
the thin, tube-like penis, the differentiation in the position and
form of the prostate, the absence of the conduit form of cervical
lappets, together with the more or less sinuated aperture, are
sufficient characteristics for separating these snails as a sub¬
family of Viviparidae. Grill separated these shells on account of
the supposed absence of cusps on the reflection of the radula
teeth, but an examination of these with high powers shows that
cusps are present in all species, though they are small and
difficult to see. The group appears to be wholly American, and
of more or less restricted distribution. Hannibal’s family
Lioplacidae is wholly untenable, being founded on a mistaken
assumption that the operculum of Campeloma has the same sub¬
spiral nucleus as Lioplax. There are but two genera, Lioplax
and Campeloma.
Family Amnicolidae
Genus Amnicola Gould and Haldeman
Subgenus Marstonia nov.
Apex of shell acute, nuclear whorl not flat-topped, emerging
well above the second whorl ; the nuclear whorl is only half the
size of that of typical Amnicola. Central tooth of radula about
two-thirds as large as that of Amnicola ; basal ridge with but one
large denticulation ; reflection usually with more than 9 denti*
culations; lateral teeth without a distinct lobe at the inner
ventral border which is replaced by a broad swelling. All cusps
usually longer and sharper than in typical Amnicola. The
second marginal is peculiarly expanded on the lower part of the
body of the tooth, differing from that in limosa. This group will
include Amnicola lustrica, gelida, oneida, walkeri, pilshryi, and
winkleyif and possibly others not examined. Named in honor
of Mr. Geo. T. Marston, who lived for many years in eastern
Wisconsin and accumulated a large collection of state mollusks.
Type : Amnicola lustrica Pilsbry.
Genus Hoyia nov.
The genus is based on the peculiarities of the radula. The
teeth are ail very small, about a third the size of those of
196 Wisconsin Academy Sciences, Arts, and Letters.
Amnicola limosa; center tooth with 7 denticulations and 4 basal
denticulations ; lateral tooth with a quadrangular-shaped body
from which a rather narrow peduncle, almost twice as long as
the body, extends laterally in an almost straight direction, but
slightly bent downward; reflection very wide and low with
about 30 very small cusps of equal size; first marginal falcate,
with a rather wide body and a heavy central ridge, reflection
wide and low with about 30 denticulations, which are twice as
large as those of the lateral ; second marginal narrower than first
with about 40 very small, equal cusps.
Type : Amnicola sheldoni Pilsbry.
The radula of Amnicola sheldoni is quite unlike that of any
other American amnicoloid observed or published. The genus
most nearly resembling Hoyia is Potamopyrgus Stimpson, in
which the central tooth has four basal denticles and the reflec¬
tion is 9-cuspid. However, the lateral teeth of Hoyia differ from
the type of Potamopyrgus in the far greater number of cusps.
The genus is dedicated to Dr. P. K. Hoy, a pioneer naturalist
and molluscan student of Wisconsin.
Genus Birgella nov.
The characteristics of the genus are based on the animal.
Verge compressed, the penis being much longer than the flag¬
ellum sheath which is short and conical or compressed; head
wide, tentacles rather short and flattened, pointed; rostrum
wide, squarely truncated. Central tooth of radula with a long,
pointed central cusp which reaches nearly to the base of the
tooth, the lateral ridge with biit one large denticle and two
smaller ones below; lateral teeth with a large rounded lobe on
the inner base of the body, with a wide peduncle and a reflection
with about 8 denticulations; marginal teeth with relatively few
denticulations ( 10-12 ) .
Type : Paludina suhglohosa Say.
Birgella differs from Somatogyrus in the denticulation of the
center and second marginal teeth, and in the form of the lateral.
It is nearest to Gillia in the denticulation of the lateral ridge of
the center tooth, but differs in other respects, as well as in the
form of the verge.
The genus is dedicated to Dr. Edward A. Birge, President
Emeritus of the University of Wisconsin and Director of the
Wisconsin Geological and Natural History Survey.
Baker — American Fresh Water Mollusca.
197
Family Pomatiopsidae St imp son
Pomatiopsinae would seem to rank as a family rather than as
a subfamily of Amnicolidae. The sinuses of the foot, the radula
with its few cusps of large size and the two large denticles on
the base of the central tooth, as well as its terrestrial habits, all
mark this genus as separable taxonomically from the Amni¬
colidae. The verge, also, is simple without flagellum sheath, and
is of large size. Gill in 1871 and Pilsbry in 1906 raised the
group to family rank, but without indicating the reasons for
so doing.
Family Physidae Dali
A study of the animals of Physa from Europe and America
indicate that the name Physa cannot be applied to the Ameri¬
can forms of these shells. The type of Physa is the Bulla
fontinalis of Linn. The animal of this species has a mantle
which partly envelops the shell, not only on the parietal and
columella side, but on the outer lip area as well. The mantle of
the left side extends well over the shell to the center of the
dorsal side ; the lower part is composed of one large lobe, while
the upper part, which is digitate, covers the spire. The part of
the mantle over the lip is digitate and extends well over this side
of the shell. In the mantle of the American Physae the digitate
mantle is reflected only over a small part of the parietal wall and
columella region, and the outer lip mantle is simply thickened
and is not digitate or extended over the shell. The center tooth
of fontinalis is also different from that of the American species,
having a large number of small denticulations on each side of the
center cusp, and the reflection is also wider than in the Ameri¬
can species.
In view of this difference between the type of the genus Physa
and the American species, it seems necessary to place these
species in a separate genus. Rivicola Fitzinger, 1833, is founded
on the same type, Physa fontinalis.
In 1842, Haldeman defined two groups of Physa; Physella,
with the type P. glohosa, and Physodon, the type being P.
microstoma. While the characters given by Haldeman are
trivial, the names seem available for the division of the group
with a digitate mantle partly covering the shell on the columella
198 Wisconsin Academy of\ Sciences, Arts, and Letters.
side and without any sign of digitations on the outer lip side.
Haldeman’s first generic name, Physella, is therefore here rede¬
fined to include these American species. Physodon is reserved
for a group of Physae which differ somewhat from Physella in
the genitalia and in the form of the shell.
Before giving the diagnosis of this and other groups of fresh
water pulmonates it seems desirable to briefly discuss some
of the features of the male genital system in this suborder of the
Pulmonata. The male organ consists of a more or less cylindri¬
cal, sac-like body, the proximal end of which contains the male
penial aperture ; back of this is a smaller, more or less tube-like
body which contains the male intromittent organ; the vas de¬
ferens enters the distal end of this body. Simroth (Bronn^s
Tier-Reieh, III, Mollusca, p. 502, 1912), following Buchner, ad¬
vances a nomenclature of the male organ for the different types
of Planorbis, and it would seem feasible to enlarge this to in¬
clude all of the fresh water pulmonates, as the same types are
found in all genera thus far examined. Modifying the German
names to the needs of English readers, the following terms may
be employed : lower cylindrical body, the praeputium, or
‘grosse penisscheide ^ ; the smaller body above, the penis sheath
or ^ kleine penisscheide ’ ; the intromittent organ within this body
is the penis; in some groups there may be a large papilla sur¬
rounding the penial stylet. In all groups there are one or more
accessory organs or appendages, such as a flagellum, a sarcobe-
lum, etc.
Pilsbry (Nautilus, XXV, p. 10, 1911) has suggested that the
penis-sac (penis sheath) of the writer ^s Lymnaea monograph is
comparable to the epiphallus of the land Pulmonata. An ex¬
amination of sections of the male organ of that and other
families would seem to indicate that the name is not quite
synonymous. The epiphallus is described as an enlarged por¬
tion of the vas deferens before that tube enters the cavity con¬
taining the male intromittent organ. As the body called the
penis sheath contains the mate organ and is retracted by a power¬
ful muscle it cannot be comparable to the epiphallus of the land
snails. In all of the groups of fresh water snails thus far ex¬
amined (Physella, Planorbis, Lymnaea, Ferrissia) the combina¬
tion of parts of the male system indicated above have been
found. In some Lymnaeas and Planorbis, the vas deferens has
been noted to be rather enlarged before entering the penis
Baker — American- Fresh Water MoUusca.
199
sheath, and this may be homologous with the epiphallus of the
land shells. The function of this organ, however, appears to be
performed by other organs in the fresh water snails.
Subgenus Physella (Haldeman 1842) Baker, emend. 1926
The male system of the genitalia of Physella consists in part
of a large, cylindrical praeputium and a longer, narrower,
cylindrical penis sheath, which contains the very long and
slender penis. The penis sheath may be modified to form a
lower part which is very thick-walled, in which the penis is
very slender, almost needle-like, and an upper part with very
thin walls in which this part of the penis is much thicker and
even the vas deferens canal is of larger diameter. At the distal
end of the praeputium there is always a rounded, more or less
heart-shaped body which seems to be comparable to the sarcobel-
lum of land snails and performs the office of an excitory organ.
It is cleft in the center for the passage of the male intromittent
organ. There is, also, near the center or upper part of the
praeputium a peculiar gland-like body, hollow, with its open end
facing the cavity of the praeputium. This may be a blind sac
comparable to the flagellum of land snails and used to form the
spermatophore. There are two retractor muscles.
The genus Physella appears divisible into two groups or sub¬
genera.
Subgenus Physella Sensu stricio
The genitalia as described above. The shell is usually thin,
with a distinct plait on the columella.
There are two longitudinal muscular pillars in the praeputium
one of which connects with the large gland.
Type: Physa glohosa Haldeman.
Subgenus Physodon (Haldeman 1842) Baker, emend. 1926
In this group the shell is usually thick and solid, the colu¬
mella smooth and without a distinct plait, although there may
be one or two small denticles on the columella ; male system with
a large praeputium and a penis sheath somewhat shorter than in
Physella, which is not divided into a thin-and thick-walled por-
200 Wisconsin Academy ofj Sciences, Arts, and Letters.
tion. There is but one large longitudinal muscular pillar ex¬
tending the whole length of the praeputium, there being none
to connect with the gland.
Type : Pliysa microstoma Haldeman.
The group of Physellae typified by integra and walkeri ap¬
pears to form a natural subdivision of the genus. The genitalia
only of integra and walken are known, but the form of the shell
suggests the inclusion of microstoma and anatina in the same
assemblage, thus avoiding the coining of an additional group
name. The approximation must necessarily be tentative until
the anatomy of microstoma is known.
Family Planorbidae H. & A. Adams
It is believed that the family should be restricted to include
the orb-like snails. Pompholaginae and Isidorinae should prob¬
ably be raised to family rank, based on peculiarities of genitalia,
shell, and radula. Simroth, in Bronn’s Tier-Reich, 1912, p. 502
has divided the male organ of Planorbis into four types, all but
one of which are totally unlike any of the types founded in
America (thus far examined) . If it be conceded that Montfort, in
1810, was the first author to definitely assign the type of Miil-
ler’s genus Planorbis {Helix cornea Linn.), and it would seem
that under the rules of the code this is perfectly clear, then the
typical genus is not found in America, as far as known. Cer¬
tainly the large species typified by trivolvis cannot be included.
In this group the male system consists of a large sac-like prae¬
putium and a penis sheath about half as long, roundly swollen at
the summit and containing a cylindrical, pyriform penis. There
are two retractor muscles. In addition there is a large gland¬
like body situated at the upper part of the praeputium, in a large
swelling. A long, folded duct leads from this gland to the
base of the penis sheath.
Nothing like this combination is r-ecorded for any group of
Planorbis at present known and its presence marks these large
American species as belonging to a very distinct group of the
family. Indeed, should the division prove to include only these
large snails, it might be considered of higher group value and
constitute a subfamily HELISOMINAE. This organ was first
noted in 1911 (Lymnaeidae of North and Middle America, p.
121) in the discussion concerning the separation of the Planor-
Baker — American. Fresh Waier Molhisca.
201
bidae from the Lymnaeidae. The genital system was not at that
time sufficiently well studied to make out the trae character
of the organ, though the figure (plate xiii, fig. A) indicates the
gland on the praeputium (called penis) and the duct which is
shown as entering the base of the penis sheath (called penis
appendage). A section of the male system shows the gland
to be a somewhat cup-shaped organ, the opening of the ^cup^
facing the interior of the praeputium. The duct leads from the
hinder end of this ^cup’ through the wall of the praeputium, and
after coiling somewhat on the outside of the praeputium, enters
the lower part of the penis sheath at a point above the muscular
ring (sarcobellum) separating praeputium from penis sheath.
The section is totally unlike anything figured by Simroth or any
other author.
Just what is the function of this peculiar gland and its duct
is not perfectly clear. It may be homologous with the flagellum
of the land snails, and may be for the purpose of forming the
spermatophore, as in this group of snails. It is probably also to
be correlated with the flagellum of the Ancylidae. That it should
be present in some groups of the family and not in others is
noteworthy. It has been found in all of the species of the large
planorbes examined (trivolvis, pseudotrivolvis, truncatus^ bin-
neyi, antrosus and varieties, campanulatus) . The peculiar
gland like body has been observed in all groups excepting
Gyraiilus.
The aggregation of characters embraced in the large plan¬
orbes of America is quite different from typical Planorbis of
Europe, and the group should be known as :
Genus Helisoma Swainson, 1840
The shell is usually siiiistral, few whorled, the whorls are car¬
inate above and often below, the base funicular, and the aper¬
ture suddenly expanded and thickened; the lateral teeth of the
radula are tricuspid while those of Planorbis corneus are bi¬
cuspid, and there are more teeth in a row than in any American
species. Genitalia with a large gland-like body in the praeputium
connecting with the penis sheath by a long duct. Two retractor
muscles, placed on opposite sides of praeputium. The genus is
divisible into three groups or subgenera.
202 Wisconsin Academy of, Sciences, Arts, and Letters.
Subgenus Helisoma s. s. -
Shell ultra dextral with funicular base and spire, the whorls
carinated above and below. Genitalia with large praeputium
and short, very wdde penis sheath; the penis is strikingly pyri¬
form, extending well into the praeputium ; gland large but duct
very short and thin. The shell is carried at right angles, tilted
to the left side.
Type: Planorhis hicarinatus Say {—antrosus Conrad).
Subgenus Pierosoma Dali, 1905
Shell sinistral, large, high, with few whorls, the early ones
carinated and flattened above, funicular below, in the adult shell
the apical whorls are slightly depressed below the upper plane
of the spire; the aperture is suddenly expanded and thickened
within. Genitalia with very large praeputium and small, nar¬
row penis sheath attached to the distal end of the praeputium
or on the side between base and summit; penis short, not enter¬
ing the praeputium ; gland very large and duct very long and of
large diameter. The shell is carried almost perpendicularly by
the animal in life.
Type : Planorhis trivolvis Say.
Subgenus Planorbella Haldeman, 1842
Shell sinistral, depressed, whorls more numerous than in Heli¬
soma ; apex not depressed below the level of the spire; base
funicular ; body whorl constricted behind the campanulate
aperture; genitalia with a large praeputium and a small penis
sheath as in Pierosoma. The shell is carried perpendicularly by
the animal.
Type: Planorhis cmnpanulatus Say.
The genitalia of Planorbella and Pierosoma are almost iden¬
tical and the only reason for the separation of the two groups
is in the form of the aperture. There are also, some small differ¬
ences in the radula. In these two groups, the penis sheath may
be placed at the distal end of the praeputium or it may be
found on the side between base and summit. This latter condi¬
tion is usually found during the breeding season, at which time
the upper part of the praeputium is much swollen and the gland
Baker — American Fresh Water Mollusca.
203
distended. A specimen of trivolvis examined in July had the
gland filled with a floeculent mass of mucus-like material, among
which were many objects resembling spermatophores.
Genus Planorbula Haldeman, 1842
Ill both genitalia and radula the Planorbula group differs
rather radically from the Segmentina of Europe and Asia. The
penis sheath lacks the wing-like blind sac shown in Simroth^s
figure, besides possessing the peculiar gland present in Helisoma,
though lacking the duct of that genus. The radula is also dif¬
ferent, the laterals being as in Gyraulus and some Helisoma,
tricuspid, not multicuspid, as in the European Segmentina. The
jaw is also segmented in Segmentina, while it is in three pieces
in Planorbula, a large superior and two small lateral pieces.
These differences are sufficiently well marked to remove Plan¬
orbula from Segmentina and give it a generic place in the fam¬
ily. The shell is carried almost flat by the animal, much as in
Gyraulus, which the animal greatly resembles.
Type: Planorhis armigerus Say.
It is to be noted that Germain, in his Catalogue of the Plan-
orbidae in the Indian Museum (Records of Indian Museum, XXI,
p. 179), raised this group to generic rank, but based the distinc¬
tion entirely on shell characters, principally those of the lamellae
in the aperture.
Genus Menetus H. & A. Adams, 1855
The genitalia of Menetus exacuous most nearly resemble those
of Planorbula armigera. There is a conspicuous gland but no
duct, the penis is very long, narrow, and extends well into the
space of the praeputium. There are two vertical muscular
ridges in the lower part of the praeputium, the latter being di¬
vided into two compartments by a muscular ring, the upper part
containing the gland and the lower part the vertical ridges.
There is one large retractor muscle at the distal end of the prae¬
putium, which is bifurcated, one branch attached to the penis
sheath and one to the upper part of the praeputium, and another
on the side of the praeputium. There appears to be a small, tri¬
angular stimulating body or appendage at the end of the penis,
just below the outlet of the sperm canal.
Type : Planorhis opercularis Gould.
204 Wisconsin Academy of\ Sciences, Arts, and Letters.
If Anandale’s remarks on Hippentis (Records Indian Mu¬
seum, XXIV, p. 359, 1922) are correct, Menetus cannot be in¬
cluded in it, if opercularis is like exacuous in its genitalia and
radula. The praeputium is not sharply differentiated from the
penis sheath in Hippeutis, the whole male organ forming a long
cylindrical apparatus narrowing toward the distal end, as shown
in Simroth’s type III. The lateral teeth, also, are arranged in
pairs in Hippeutis, a feature not observed in exacuous. The
group seems quite distinctive enough to stand as a separate
genus.
Genus Gyraulus Agassiz, 1837
In the absence of a gland or duct on the praeputium, the pres¬
ence of a horny stilet at the end of the long, narrow penis, which
is surrounded in the head of the praeputium by a fleshy papilla
(sarcobellum), and in the strongly fragmented jaw, this group
of planorbes stands easily as a well recognizable genus. There
are two well developed vertical muscular pillars in the prae¬
putium, similar to those in Physella and Lymnaea. The genus
is divisible into several subgroups, more or less well character¬
ized.
It is to be noted that Annandale and Prashad in 1919 (Rec¬
ords Indian Museum, XVIII, p. 52) gave the anatomical char¬
acteristics of this group and indicated its right to hold generic
rank.
Subgenus Gyraulus s. s.
In the typical group the penis sheath is longer than the prae¬
putium and much swollen toward the distal end. The shell is
usually more or less hirsute and strongly spirally marked. The
American Planorhis deflecUis is the same in structure of geni¬
talia as the European Planorhis alhus Muller, the type of the
genus.
Subgenus Torquis Dali, 1905
Whorls of shell less distinctly striated, not hirsute, the base
deeply and regularly excavated; male system with penis sheath
and praeputium forming a regularly cylindrical shape, the penis
Balter — American Fresh, Water Mollusca.
205
sheath much longer than the praeputium, not swollen at the
distal end. There is a single retractor muscle attached to the
distal end of the praeputium.
Type: Planorhis parvus Say.
Subgenus Armigee Hartmann, 1840
This group is characterized hy the costate whorls, the costae
projecting at the periphery. Fresh material has not been
available for anatomical examination and it is not now known
how closely this feature may approximate with the other groups
mentioned.
Type: Nautilus crista Linn.
Family Ancylidae
Subfamily Ferrissiinae Walker 1917
The genitalia of Ferrissia differs radically from those of
Pseudancylus (Ancylus) fluviatilis as figured by Simroth. The
praeputium is long and cylindrical, the penis sheath smaller but
cylindrical and a trifle more than half as long as the praeputium,
the flagellum enters the praeputium at the junction of the penis
sheath with that body, and is short, enlarging at its distal end
to form a large, more or less fan-shaped, gland-like organ. The
genitalia seem to differ from the other members of this family
to the same degree that the radula and shell do, showing that
the whole organism agrees in the characteristics which separate
this genus from the other groups of the family Ancylidae.
REVISION OF THE NEARCTIC SPECIES OF
HELOPHILUS AND ALLIED GENERA
C. H. Curran, Ottawa, Canada,
AND
C. L. Fluke, University of Wisconsin, Madison, Wis.
The revision of the genus Helophilus Mg. was originally
Xilanned by the junior author who succeeded in bringing to¬
gether most of the described forms from North America. Owing
to increased duties it was found impossible to continue the work
and consequently he suggested to the senior author that the
material gathered together be forwarded and the revision be
carried on by him. The outcome was that a most pleasant and
successful co-operation was arranged, enabling us to go into
fuller details in regard to the various species than would other¬
wise have been possible. While the preparation of the manu¬
script has been frequently delayed, this has been found to have
been advantageous, as excellent series of some of 'the less com¬
mon forms have come to hand during the past fifteen months.
Types of the majority of the species have been examined, viz :
all those described by Loew and Williston, in addition to the
species described by Graenicher. Access has not been had to
types of Wiedemann, Macquart, Walker, or Bigot, but we have
secured in many cases examples from the type localities, thus
ensuring almost positive identification. Mr. E. E. Wehr, com¬
pared a specimen with Pterallastes perfidiosus Hunter, but we
have not been able to find any trace of the type of H. pilosus
Hunter. Pterallastes borealis Cole quite evidently belongs near
perfidiosus in the genus Lejops.
The collections upon which this paper is chiefly based are
those of C. L. Fluke, C. H. Curran (now incorporated in the
Canadian National Collection), the University of Wisconsin
Collection, Canadian National Collection and that of the late A.
L. Lovett. In addition, many other collections have been ex¬
amined, including the Museum of Comparative Zoology at Cam¬
bridge (Loew and Osten Sacken collections) ; United States
208 Wisconsin Academy of, Sciences^ Arts, and Letters.
National Museum; American Museum of Natural History, New
York; Boston Society of Natural History; Kansas University,
Lawrence; University of Minnesota; Cornell University; Koyal
Ontario Museum, Toronto; R. C, Shannon; Dr. J. Bequaert;
and A. L. Melander. These collections contain thousands of
specimens and undoubtedly constitute the majority of specimens
in North America. In addition, many collectors in various parts
of the United States and Canada have submitted specimens for
determination. Wherever material permits, Palaearctic species
in the Curran collection have been included in the keys and brief
notes given.
Owing to the detailed descriptions, it has been found neces¬
sary to eliminate a catalogue of the species. Only references to
each of the original descriptions and synonyms are given.
Use is made of the terms: posterior ocellar line (a line drawn
between the posterior ocelli) and median ocellar line (a line ex¬
tending from the anterior ocellus to the middle of the posterior
ocellar line) ; as there is a great deal of variation in the lengths
of these lines, the differences are often of specific importance.
Dr. James S. Hine’s ‘'Alaskan Species of Diptera of the
Genus Helophiliis with Notes on Others” (Ohio Journal of
Science, XXIII, 192, 1923) is the only large paper published on
the genus during the past twenty years. Other recent papers
containing records have been published, the three most im¬
portant being: “'Syrphidae of Colorado”, by C. R. Jones,
“Syrphidae of Wisconsin”, by C. L. Fluke, and “Syrphidae
of Nebraska” by E. E. Wehr. Reference to other papers will
be found in the “Syrphidae of Wisconsin”.
It has been deemed advisable to recognize the various genera
proposed for the different groups belonging to Helophilus.
Shannon, in his revision of the subfamilies, etc., recognized at
least four groups. We have recognized practically all the
genera previously placed in the synonymy and have found it
advisable to establish a few others : only one of these, however,
occurs in North America. The habitus of the flies in each
genus is, for the most part, quite homogeneous and it is felt
that the classification is improved by following the course adopt¬
ed.
It is unfortunate that such well known names as ^^conostomus
Williston^^ and ^^similis Macqd^ must be relegated to the
synonymy, the latter on account of pre-occupation, but it seems
Curran & Fluke — Helophilus and Allied Genera 209
advisable to make the changes at the present time rather than
continue the use of invalid names.
Modified Table to Genera of Helophilini
A. Diptera with spurious vein between third and
fourth longitudinal veins _ _ _ ^Syrphidae
B. Anterior cross vein oblique, terminating
at or beyond the middle of the discal ceil ;
third vein with a strong, more or less V-
shaped, curve into first posterior cell;
each femur at the base antero-ventrally
with a small patch of black setulae _ Eristalinae
C. Marginal cell open _ Helophilini - 1
1. Hind femora at the end with a triangular area
before the tip, the base of this with a distinct
spur _ Merodon
Hind femora arcuate or plane, not wnth a tri¬
angular projection, if swollen apicallj^ the
spur is not present. _ 2
2. Eyes pilose _ 3
Eyes bare _ 4
3. Thorax with pollinose yellow crossbands; pile
moderately abundant ; not bumble-bee-like
(Europe) _ Myiatropa
Thorax at most slightly pollinose, not with dis¬
tinct bands; pile very dense, largely obscur¬
ing ground color; somewhat bumble-bee-like _ Mallota
4. Arista of the male extraordinarily dilated at
tip, of female slightly so (Europe) _ Platynochaetus
Arista simple, bare _ _ _ 5
5. Large species, the thorax thickly yellow or
orange pilose ; hind femora swollen and
arcuate in both sexes ; hind tibia without
long apical spur ; rather bumble-bee-like
flies _ Mallota
Generally smaller, the pile not long and dense ;
if hind femora arcuate the tibiae with long
apical spur and coxae with spur ; not
bumble-bee-like _ _ 6
6. Mesonotum densely, evenly yellow pollinose, the
ground color nowhere evident; face evenly
concave in female; with tubercle, but re¬
treating below in male ; abdomen wholly
black _ _ _ _ Pt er alias tes
Mesonotum not entirely yellow pollinose : if al¬
most so the face is more prominent below
in male and female, seldom evenly gently
concave, the abdomen not wholly black _ 7
14
210 Wisconsin Academy of Sciences, Arts, and Letters.
7. Hind femora of male large, arcuate, abdomen
chiefly red ; hind femora of female not
arcuate, thorax densely pollinose, obscuring
ground color; abdomen normally with inter¬
rupted greyish yellow pollinose fascia on
each segment; face with small tubercle, re¬
treating below ; no stigma but stigmal cross¬
vein present ; large species, 14 to 17 mm. _ Polydontomyia
Not with all these characters; if doubtful the
stigma is twice as long as wide or the size is
not over 11 mm. Heloplnilus s. lat.
Table of Genera of Helophilus S. L.
1. Face wholly pollinose, without a shining
median vitta _ 2
Face with a shining median vitta on at least the
lower half _ _ _ 5
2. Face produced into a long, acute cone (Sub-
genus of Lejops) _ Enrhimyia Big., page 273
Face not strongly produced _ 3
3. Eyes of male, contiguous or closely approximat¬
ed ; posterior basitarsi with nodulate pube¬
scence below on at least the basal half
(Europe, Asia, Africa) _ Mesembrius Bond., page 229
Eyes always broadly separated ; hind tarsi with
normal hairs _ 4
4. Abdomen short, not twice as long as wide;
hind tibiae truncate at apex. (Fig. 25)
_ ParhelopMlus, page 230
Abdomen elongate, with parallel sides or taper¬
ing, over twice as long as wide; hind tibiae
produced at apex below. (Fig. S2) ^.Lejops Bond, page 254
5. Stigma at least twice as long as wide, although
often paler apically, not simulating a cross¬
vein; large species, abdomen broad - 6
Stigma simulating a crossvein (Fig. 22 )_ - 8
6. Scutellum tuberculate at each side apically
(N. Zealand) _ Pilinasica Mall.
Scutellum not tuberculate - 7
7. Hind femora with a long, sub-basal spur below
(New Zealand), Genotype Syrphus irilinea-
tus Fabr.) _ Prohelophilns new genus
Hind femora without such spur (Palaearctic,
Nearctic and Australian Begions) -
_ _ Eelophihis Meig. page 211
8. Front very broad; posterior ocelli remarkably
remote; the ocellar triangle very large or
very flat; either the front strongly swollen
Currom & Fluke — Helophilus and Allied Genera 211
below or the face with distinct tiiborcle ; ab¬
domen always broad _ _ _ _ _ _ 9
Front moderately broad or rather narrow ;
never unusually large ; face, or the front not
as described, though often prominent at
middle; face rarely tuberculate, in which
case the ocellar triangle is small _ _ 10
9. Front remarkably swollen below; genital clasp-
ers remarkably long ; ocellar triangle of
moderate size; bright colored spp. (S. Am.)
- - Bolichogyna (Fig. 28)
Front not remarkably swollen, ocoellar triangle
very large; genital claspers normal; dark
colored, except sometimes on abdomen,
(North’ America) (Figs. 11, 12) _
- - - Asemosyrphus Big. page 247
10. Face carinate, more or less retreating (Africa,
Asia, E. Indies) _ Mesemhrius
Face tuberculate in both sexes ; small dark color¬
ed species with broad abdomen and brassy
yellow pile (North America) _
_ Lunomyia new genus, page 252
Key to the Species of Helophilus
1. Face with a median shining black stripe _ 4
Face with a median shining yellow or reddish,
rarely ferruginous stripe _ 2
2. Front of male narrow, wholly black pilose above
the depression; of the female wholly black
pilose _ fasciatus Walk.
Front of male wider, only black pilose on the
upper half of the narrowed portion; of the
female, broadly yellow haired below _ 3
3. European _ _ trivittatus Fabr.
American _ latifrons Loew
4. Apical fifth of hind femora and basal half of
their tibiae reddish yellow; third antennal
joint brown _ _ _ _ — 8
Hind femora with only the narrow apex or a
narrow preapical band reddish and only the
basal third of their tibiae yellow _ 5
5. Third antennal joint usually largely reddish;
abdomen with three or four pairs of trans¬
verse oval spots; hind femora often reddish
at base; thoracic vittae irregular _ _ _ ohscurus Lw.
Third antennal joint brown or black; never
clearly reddish; abdomen variable, not with
oval spots, the hind femora never reddish at
base; very dark or chiefly pale flies - 6
212 Wisconsin Academy of Sciences, Arts, and Letters.
6. The spots on the third abdominal segment oc¬
cupy most of the segment; thoracic vittae
never interrupted (Europe, N. A.) _ hybridus Lw.
The pale spots on the third segment are not the
predominating color, the segment being at
least half black and shining; thoracic vittae
often interrupted _ 7
7. The apex of the second segment is black in its
entire width or practically so _ groenlandicus 0. Fabr.
The apex of the second segment is broadly
orange on the sides, so that the whole base
of the abdomen to the middle of the third
segment is orange laterally _ _ _ borealis Staeg.
8.. Posterior tibia black on the apical two fifths or
more; pollinose section of front of male
above depression distinctly longer than
wide; ocellar triangle equilateral (Europe) — pendidus L.
Posterior tibia reddish on apical half, yellow on
basal half; pollinose section of front above
depression wider than long; ocellar triangle
decidedly wider than long ( Saskatchewan n. sp.
Helophilus groenlandicus 0. Fabricius.
Tabanus groenlandicus Fabr., Fauna GroenL, 208. 1780
(Greenland)
H. arcticus Zett., Ins. Lapp.. 595 (1839) (Lappland).
m. bilineata Curtis. Ins. of Ross’ Exp., LXXVII, (1831).
m. latro Walker, List III, 607, 1849. (Ont., N. S.)
m. androchis Walk., List III, 612, 1849. (Ont.)
Plate V, figures 4, 5
A deep black species, face with black stripe, thoracic vittae
narrow; abdomen with three interrupted pale fasciae, the black
apical band and second segment entire ; the mesonotum usually
largely black, pilose on the disc behind the base of the wings.
Length , 11-12 mm. Male. Face black, the sides broadly,
yellowish on upper half or more, the middle stripe, oral margin
and cheeks shining, elsewhere densely pale yellowish pollinose,
the pile rather abundant, whitish ; in profile, moderately concave
on upper two-fifths, the lower portion with a slight tubercle
occupying its upper third, moderately produced below the eyes.
Front with pale yellow pollen, broadly shining above ; a sub-
triangular area immediately above the antennae also shining
Curran & Fluke— Helophilus and Allied Genera 213
black ; pile black, below the groove and slightly above it and at
the vertex, yellowish. Occiput greyish pollinose, with yellowish
pile. Antennae black, third joint slightly longer than wide, its
sides almost parallel, its end obtusely rounded ; arista yellowish.
Mesonotum rather dull black, the sides more or less densely
greyish pollinose; with a pair of widely separated greyish,
narrow, usually entire greyish vittae, one on either side of the
middle line. Middle portion of pleura lightly pale pollinose.
Pile yellow, brighter laterally; behind the root of the wings,
but not reaching any margin, sometimes all black, but often this
is reduced to a small black pilose area on either side. Scutellum
brownish yellow, the base narrowly black, the pile short, black,
longer, yellow on the margins.
Legs black, the narrow apices of the femora, basal fourth or
fifth of front and hind tibiae and basal half or less of the middle
ones, yellow. Pile black, on the basal half of the middle femora
behind and on upper surface of hind ones, inner side of front,
postero-ventral surface of hind, base and both anterior and
posterior surfaces of the middle tibiae, more or less yellow.
Wings pallidly cinereous, the stigma brownish or brownish
yellow, the veins very dark. Squamae yellow, with yellowish
fringe. Halteres yellow.
Abdomen opaque black, the apices of the second to fourth
segments increasingly widely shining black, the last mentioned
half shining, the side margins also shining. First segment
lightly greyish pollinose. Second segment with large yellow
triangles, not reaching the hind border, their posterior margin
a little oblique, the posterior corners usually rounded off slight¬
ly, the anterior inner corners usually rather acute, their anterior
surface a little concave, the spots reaching the base of the seg¬
ment broadly at the sides. Third segment with a large basal,
sub-rectangular spot occupying two-thirds the length of the seg¬
ment and extending inwards to occupy one-third the width, the
inner ends produced more behind and with a greyish, slightly
oblique interior prolongation, the inner ends moderately separat¬
ed. Fourth segment with a pair of oblique, slightly lunulate,
narrowly separated greyish pollinose spots on the front half,
their outer ends near the middle of the segment, both ends
rounded. Pile short, yellowish, longer laterally, on the
posterior half of the second to fifth segments, black, but not
reaching the sides. Genitalia almost all pale haired.
214 Wisconsin Academy ofj Sciences, Arts, and Letters.
Female. Front moderately wide, slightly narrowing above,
pale yellowish pollinose but less thickly so on the middle portion,
ocellar region bare; pile black except at vertex and a few hairs
on the sides below. Legs a little more extensively pale.
Abdomen somewhat more extensively shining, the spots on the
second segment narrower, especially the inner arms; only the
small corners of the third segment orange, a pair of transverse,
very slightly lunulate spots, narrowly separated in the middle,
separated from the front margin by less than their width, grey¬
ish white pollinose, lying wholly before the middle of the seg¬
ment. Fourth segment not reddish on basal comers, the
pollinose fascia similar to that on preceding segment, less widely
separated from base, and not reaching the side margins; fifth
segment with similar fascia. The black pile on posterior half of
the segments reaches the side margins.
Redescribed from a male, Loew Collection; 2 male, 2 female,
Ilopedale, Labrador; male. Peaceful Valley, Colo.; female. Lake
Opasatika, Que. Over one hundred specimens from Pingree
Park, Colorado, 1924.
A very distinct species because the black of the hind margin
of the second segment reaches the sides in both sexes. There is
considerable variation in the amount of black pile on the
posterior part of the thorax and it may be all wanting.
Helophilus borealis Staeger
H. borealis Staeger, Kroyer^s Tidskrift, n. ser., I, 359, 1845.
(Greenland)
H. glacialis Loew, Stett. Ent. Zeit., 1846, p. 121 (Aldr. Cat.
gives 43, 120). (Labrador)
H. dychei Williston, Can. Ent., XXIX, 136, 1897. (Alaska)
E. bruesi Graenicher, Bull. Wis. Soc. VIII, 40, 1910, (Wis.)
Wristalis androclus Walk., List III, 612, 1849.
Allied to groenlandicus, but the abdomen is all yellowish
pilose except a small apical triangle on each segment, sometimes
wanting on second and third.
Length, 12-13. 5 mm. Male. A middle stripe, narrow oral
margin and cheeks shining black, the face elsewhere yellowish
red and densely yellow pollinose, the pile pallidly yellowish.
Face in profile moderately concave on upper two-fifths, its lower
portion slightly receding and a little convex, two-fifths of the
Curran & Fluke — Helophilus and Allied Genera 215
face lying below the lower border of the eye. Front with
parallel sides above the depression, moderately wide, but the
upper portion decidedly longer than wide; before the suture
with whitish yellow pollen, but it becomes yellow or even brown¬
ish yellow on the upper portion of this part and is brownish
yellow on the upper part, practically bare across the ocelli; a
broad shining black arch immediately above the antennae ; pile
yellow on lower portion, on lower part of upper portion and at
vertex, elsewhere black. Occiput greyish yellow pollinose, with
yellowish pile which becomes almost whitish below. Antennae
black, the third joint velvety brown, irregularly circular, not
oval, slightly broader than long, the arista twice as long as
antenna, tapering, ferruginous.
Mesonotum rather dull blackish, the sides aeneous or thinly
greyish pollinose, with a pair of broadly separated longitudinal
almost whitish, slender stripes, widest anteriorly and often
scarcely evident on posterior half ; in some specimens practically
wanting, in others reaching the scutellum and slightly broaden¬
ed behind.
Pleura thinly greyish yellow pollinose. Scutellum yellow, the
narrow base blackish. Thorax wholly greyish yellow pilose, or
with some black pile in front of the scutellum, the latter yellow
pilose.
Legs black, the apices of the femora, basal third or less, of
front and half to two-thirds of the middle tibiae, yellow. Pile on
femora black, yellow behind the front four and on dorsal surface
of hind ones ; on tibiae mostly yellow on front four, only of this
color on front side of hind ones, elsewhere black.
*Wings almost hyaline, very slightly cinerous. Stigma yellow¬
ish brown. Squamae whitish yellow, with yellowish pubescence
and white fringe. Halteres yellow.
Abdomen over half shining greenish black, the second seg¬
ment except the apex, basal half of the third and a small basal
spot on the middle of the fourth, more or less opaque, scarcely
so in some specimens, the second and third segments with yellow
spots, the third and fourth each with a pair of greyish pollinose,
transverse spots. The sides of the second segment are wholly
rather broadly yellow, expanding to form a large triangle, the
two thus formed being broadly separated in the middle, slightly
concave in front, slightly convex behind, their posterior inner
ends rounded, so that the front end is acute, the posterior mar-
216 Wisconsin Academy oft ScienceSy Arts, and Letters.
gin of the spot oblique ; the black on the posterior margin usually
extends over half way from the middle line to the side margin,
sometimes, two-thirds the distance. Third segment with a small
rectangular orange spot on anterior angles, and, on each side
of the middle line with a slightly oblique, elongate oval greyish
pollinose, spot, the inner ends of the two moderately separated.
Fourth segment with a pair of similar spots which are, however,
less widely separated, a little more oblique, arid slightly nar¬
rower. The abdominal pile is all pale, except a small apical
black pilose triangle on the apex of the fourth segment, which
may be enlarged, in which case the third segment bears a tri¬
angle. Genitalia all pale behind.
Female. Agrees closely with the male; the only conspicuous
difference is found in the abdominal pile. Second segment with
a narrow apical fascia of black pile on the median half, the third
with a wider one broadly separated from the lateral margins;
fourth segment with the incomplete black haired fascia angularly
produced forward in the middle, the fifth with a large apical
triangle of black pile almost reaching the base. The extent of
the black pile is somewhat variable.
Male. Greenland (von Moschler) ; from the Loew Collection;
specimens were compared with the type. Over 100 specimens
from Alaska, Labrador, all Canadian provinces, New^ England
States, N. Y., Wis., Minn., Idaho, Ore., Colorado, and Washing¬
ton. This species was not recognized by Williston, but he de¬
scribed one of the darker, more shining specimens as H. daeckei
on material from Alaska, the type of which was examined.
Relophilus latifrons Loew.
H. latifrons Loew, Cent. IV, 73, 1863 (Nebr.).
H. trivdttatiis Auct. (N. America).
Plate V, figures 7, 8.
Second and third abdominal segments chiefly lemon yellow in
male, rather variable in female ; face with the middle line al¬
ways yellow or reddish yellow, never ferruginous or black; front
of male only black pilose across ocelli, of female yellow pilose on
lower third.
Length, 11 to 15 mm. Male. Face yellow, densely whitish
pollinose, the middle line on the lovv^er two-thirds or more, broad-
Curran c£* Fluke — Helophilus and Allied Genera 217
ly shining; narrow oral margin and cheeks shining black; pile
white. Face in profile deeply concave on upper two-fifths ; the
lower three-fifths more prominent above, sometimes sub-tuber-
culate, retreating. Front black, yellow just above the an¬
tennae; before depression whitish yellow pollinose, above it
yellow pollinose, shining black across ocelli and vertex, above the
antennae with a rather large shining triangle which may be all
yellow or largely brownish, its upper portion forming a slender
arm not reaching the depression. Pile yellow, black across the
ocelli. Occiput greyish pollinose, more yellow above, with the
orbits above narrowly shining. Pile yellow. Antennae black,
third joint chiefly reddish, but brown above and apically ; some¬
times chiefly so ; scarcely longer than broad ; arista yellow.
Mesonotum opaque black, the lateral margin and a moderately
wide, broadly separated sub-median vittae yellow or greyish yel¬
low, the latter joined to the former in front. Pleura whitish or
greyish pollinose. Scutellum translucent yellow, the corners
only black. Pile of thorax all yellow, except on the disc of the
scutellum where it is black.
Front four femora on basal half or less, the hind ones except
the apex which also bears a black spot, black. Front four tibiae
and middle tarsi all yellow; the front tibiae more or less dis¬
tinctly brownish on apical fifth, the last three joints of the mid¬
dle tarsi fuscous; front and hind tarsi black; hind tibiae yellow
on basal third, elsewhere black. Pile all yellow, the hind femora
with short black, bristly hair below and longer bristles on apical
half, and in addition several long black or yellowish hairs. Hind
tibiae a little produced apically on inner side, arcuate ; their
femora rather evenly convex below, except basally.
Wings cinerous hyaline or with slight yellowish tinge, the
veins yellow basally; stigma yellow. Squamae whitish or pale
yellowish, with yellow border and fringe ; the lower lobe strongly
pubescent above. Halteres pale yellow.
First abdominal segment black, its sides very broadly pale
vellow, wholly thinly greyish pollinose. Second segment pale
yellow, the base on the middle two-thirds, its hing margin oblique,
and rather broadly connected in the middle with a rather narrow
subapical band, opaque black ; the middle connection usually
narrowest anteriorly always widened behind, the posterior fascia
widest at middle, slightly but distinctly narrowed to the sides,
usually very narrowly separated from the lateral margin ; very
218 Wisconsin Academy of\ Sciences^ Arts, and Letters.
narrow hind margin reddish, preceded by a narrow shining black
fascia. Third segment similar in color, usually with an oval,
anteriorly transverse, sometimes with a more elongate, basal
spot on the middle fourth, moderately broadly connected with the
broad apical fascia, sub-opaque black, often chiefly shining, but
the median connection is usually almost all pollinose so the yel¬
low appears to extend entirely across the middle of the segment,
the black posterior fascia is biconvex, reaching forward in the
middle, and more or less so laterally, but the sides opposite and
posterior margin are narrowly reddish. The fourth segment has
only the narrow, lateral and apical margins reddish, but there
may be a slight broadening of the reddish towards the front;
on the anterior half is a rather broad, anteriorly and posteriorly
biconcave greyish to yellowish fascia, which is narrowed towards
the middle and almost reaches the base at this point in some
specimens, spreading more or less along the margin posteriorly.
Pile yellow, short, on the apices of the second and third seg¬
ments, reaching the sides, moderately broadly black; on the
fourth segment not reaching the sides, all black behind the fascia.
Female. Abdominal markings usually very different from
male except in newly emerged specimens as they very often turn
blackish in large part. Pace usually slightly less concave ; front
broadly or narrowly black pilose across the ocelli and a little
before them ; whitish pollinose on sides below, less thickly so on
the broad middle line, the pollen here and above more yellowish,
the black shining color as in male.
Abdominal segments shorter, the black more extensive; the
anterior curved fascia of the succeeding segment is narrower,
the yellow only a little more than half as wide in the middle in
most specimens, the posterior fascia slightly wider and less nar¬
rowed laterally. Usually the basal black fascia on the third seg¬
ment extends three fourths across, but, sometimes as in the male,
the posterior fascia is wider, occupying almost one-third of the
length of the segment: this leaves the yellow as a narrowly
interrupted basal, broad fascia, concave on the front border
towards the middle, connected at the middle by yellow pollen.
Fascia on fourth segment as in male, fifth segment similar but
the fascia reaching the base on broad middle portion. Apices
of second and following segments broadly black pilose, reaching
the sides on only the second, and sometimes the third segment.
In almost all the darkened specimens the outlines of the yellow
Curran & Fluke — Helophilus and Allied Genera 219
areas are usually quite evident as outlined above, but there is a
strong inclination to narrow the spots so that the black predom¬
inates, the yellow or orange color forming lunulate, usually com¬
plete fasciae on the second and third segments ; these lunules
very seldom interrupted in the middle.
Described from numerous specimens from all parts of the
United States and Canada, April to October.
Helophilus latifrons shares with H, fascial us the distinction
of being the most common species in America. Its range ex¬
tends even into South America where it has been recorded by
various European entomologists as H. trhnttatus. While the two
species bear a most remarkable resemblance in almost all re¬
spects they are quite distinct. H. latifrons is not quite as large
as trivitaitus, the average length being three millimeters less;
the face is distinctly less concave below ; the antennae are lighter
in color, the third segment of trivittatus is black or almost so
in all the specimens we have examined ; the female is not almost
evenly pollinose, but is more shining except on the sides, the
fascia on the third segment is scarcely indented by a black tri¬
angle, while there is a large triangle in trivittatus and the same
applies to the following fasciae but the last fascia is interrupted
or practically so in that species, always entire in latifrons and
the last two fasciae are more transverse in the latter. In the
male the lumules on the fourth segment are larger and much
more oblique, and are narrowly interrupted or sub-interrupted
in trivittatus and the front femora are over half black. The
distinctions between the two species may not appear great
from the above, but examination reveals other slight differences
and they appear quite distinct when seen together.
There is really no question of synonymy in connection with
this species, except that it has been recorded as trivittatus from
Central and South America.
While we have seen many specimens from Mexico, which are
the present species, we have not seen a single trivittatus and
therefore feel certain that all references to the European species
frorn America are erroneous.
. Helophilus trivittatus Fabricius
Eristalis trivittatus Fabricius, AntL, 235, 1805 (Europe).
^Musca parallelus Harris, Expos. Engl. Ins., 57, PI. XV, f. 8
(1776) (Europe).
H. camporum Meigen, Syst. Beselir., Ill, 372, 1822 (Europe) .
220 Wisconsin Academy of Sciences^ Arts, and Letters.
Plate V, figure 3.
Extremely like H. latifrons Loew. The differences between
these two species have been dealt with under latifrons and need
not be repeated. No specimens have been found in America so
far as we know, although the species has been frequently re¬
corded. Its range extends over all Europe and most of Asia.
The senior author found it commonly in England and Northern
Prance near suitable breeding places. Verrall includes H. lati¬
frons Loew as a sjmonym but in this he was wrong.
Helophiliis fasoiatiis Walker
H. similis Macquart, Dipt. Exot., II, (2), 64, 1842. (Ga.)
nec Curtis.
H. fasciatus Walker. List III, 605, 1849 (Out.).
Eristalis decissus Walker, List III, 614, 1849. (Ont.)
H. sussiirans Jaennicke, Neue Exot. Dipt., 94, 1867. (Ill.)
Plate V, figure 6
Allied to H. latifrons but the abdominal bands are narrower ;
the front is much narrower and the black pile occupies all the
front above the depression in the male and all the front in the
female. The legs are more extensively black.
Length, 10 to 15 mm. Wlale. Face yellow, thickly yellow
pollinose, the middle line broadly shining reddish yellow to fer¬
ruginous but never black, the oral margin and cheeks shining
black; pile pale yellowish; in profile concave on upper three-
fifths, convex below this, the lower two-thirds convex receding,
without distinct tubercle although the formation approaches
one. Front narrowed above the upper portion nearly twice
as long as wide, except on the upper fifth yellow pollinose, the
pollen paler on the sides before the depression and often some¬
what reddish in the middle and above; immediately above the
antennae reddish, the polished triangle chiefly blackish. Occiput
yellow to greyish pollinose ; with yellow pile. Front black pilose
above the suture, yellow below it except sometimes several hairs
immediately above the antennae. Antennae reddish, sometimes
more or less ferruginous, the third joint brownish above and
apically; arista reddish.
Curran & Fluke — Helopliilus and Allied Genera
221
Mesonotum opaque (or sub-opaque black, in old specimens),
the side margins and broadly separated sub-dorsal stripes opaque
yellow or pale greyish, more or less narrowed towards the pos¬
terior margin, but sometimes the middle ones a little broadened
here, the pale vittae not joined in front. Pleura yellowish grey
pollinose. Scutellum translucent yellowish, the very narrow
.base and the corners black or brown. Pile of thorax all yellow,
short, on the disc of the scutellum black.
Femora black, the apical third or less of the front four and
the apex of the hind ones reddish yellow ; middle tibiae wholly,
front ones except the apical third, hind ones on basal third or
less and the first two joints of the middle tarsi, yellowish or red¬
dish, the legs elsewhere black. Hind femora moderately broad¬
ened, widest at apical third, below with dense black short, coarse
hairs and with fine bristles on apical half, and a few long black
or yellow hairs, the pile all yellow elsewhere.
Wings cinereous hyaline often quite yellowish anteriorly basad
of tip of first vein. Stigma luteous. Squamae whitish, with
pale yellow border and yellow fringe, with short pale yellow¬
ish pubescence on the lower lobe above. Halteres yellow.
Abdomen opaque black, with large yellow spots on the second
and third segments, the fourth segment fasciate with pollen.
First segment greyish pollinose except behind the corner of the
scutellum the sides broadly yellow. Second segment chiefly yel¬
low the black forming a rather broad basal facia which is al¬
most as long as the width of the thorax, broadly connected on
the middle line with the moderately broad posterior margin, the
apex shining, the immediate apex narrowly reddish on middle
part ; the posterior fascia is of almost equal width but is slightly
widened at middle and sides but reaches or is very narrowly sep¬
arated from the lateral margin. Third segment yellow on basal
two-thirds with an oval black spot in the middle at the base,
connected in almost its full width with the black posterior border,
but, in well preserved specimens more or less broadly pollinose
on the connection so that the anterior spot appears isolated;
the posterior margin is moderately broadly shining, the apex
reddish, or sometimes the shining portion is all reddish. The
yellow is biconvex behind so the black reaches forwards at the
sides, and usually attains the lateral margin. Fourth segment
with the narrow lateral margin and rather broad apex reddish
or obscurely so, sub-basally with an abbreviated, almost trans-
222 Wisconsin Academy of Sciences, Arts, and Letters.
verse yellow or greyish pollinose, sub-basal fascia, its anterior
margin gently concave on either side, narrowed in the middle,
so that its posterior margin is somewhat oblique on either side.
Pile yellow, short; shorter and stouter on each segment behind
the fasciae and reaching the sides on all but the last.
Female. Abdominal fasciae narrower, the front all black pi¬
lose. Front narrowed above, yellowish pollinose, the middle line
darker and with thin brownish red or yellowish pollen.
Abdominal segments shorter. The black basal fascia on the
second segment extends slightly more towards the sides and is
quite as wide as the thorax, the yellow spots are sharper in¬
wardly, seldom obtusely rounded as in male. The fascia on the
third segment occupies but little over half the width of the seg¬
ment. The pollinose fascia on the fourth segment is narrower,
but slightly narrowed in the middle, its outward posterior
portion convex so that each side appears slightly lunulate. The
fascia on the fifth segment reaches the base on its middle por¬
tion or is narrowly separated Taterally, increasingly so towards
the margin ; it is slightly widened laterally ; the segmental mar¬
gin is yellow as in the preceding. The pollen is always yellow
in this sex.
This species is probably more abundant than latifrons in most
parts of North America, but seems to be a little more northern
in distribution even though it extends into Mexico. It is also
an earlier appearing insect in most localities and is common on
bloom.
Eelopkilus fascdatus is readily distinguished from its close
ally, H. latifrons, by its much narrower front in the male and
wholly black pilose front in female and also in this sex by the
yellow pollinose abdominal fasciae, the male differing less in this
respect. The legs are more extensively black. It is a rather
variable appearing species, but we are unable to find any dif¬
ferences between the small specimens with grey pollinose fascia
and the large ones with yellow, and, notwithstanding the differ¬
ent appearance, a careful analysis shows the distinctions are
more apparent than real. Helophilus pendulus and hyhridus
have a strong black middle fascial line.
It is with regret that we are forced to change the name of this
species, which is so well known as H. similis Macq., to that used
by Walker, but it is quite evident that Macquart’s name cannot
b(' used. As pointed out by Williston, the name similis already
Curran & Fluke — Eelophilus and Allied Genera 223
existed for a British species, which was later however referred to
H. pendulus L. There is not the slightest excuse for retaining
Macquart’s name, as might have been done had Curtis’ species
been placed in another genus, although even this practice ap¬
pears to be poor policy. The fact remains that we still have a
name similis for a species in this genus, even though it be a syn¬
onym. From the descriptions both Eristalis decissus Walker and
E. susurrans Jaenniche appear to be undoubtedly this species.
Eelophilus pendulus Linne
This European species is readily distinguished from fasciatus,
which it resembles in many respects, by its deep black, shining
facial stripe, black basal antennal joints, still more blackish fem¬
ora, except the hind ones, more reddish posterior tibiae, and al¬
ways isolated yellow spots and fasciae, as they are all interrupted
in the middle. The black facial stripe distinguishes it also from
latifrons, and the much narrower abdominal yellow spots, and
much more extensively reddish hind tibiae separate it from
hyhridus.
Hine, in his recent article, recorded this species from Alaska,
but an examination of his specimens, which he so kindly sent
us, show them to be intentus n. sp.
Eelophilus intentus new^ species
Plate V, figures 9, 10
Allied to E. pe^idulus L. but the hind tibiae are wholly red¬
dish, the frontal projection weaker, front wuder, the upper pol-
linose section being wider than long instead of distinctly longer
than wide, etc.
Length, 10 to 11 mm. Male. Face yellow, the broad median
stripe, oral margin and cheeks black, the cheeks in front and
median vitta shining although the latter is thinly greyish pol-
linose above. Face with yellowish white pollen, which has a
slight brassy reflection: in profile the lower two-thirds almost
perpendicular (a slight indication of tubercular swelling above),
the upper third rather evenly concave. Frontal prominence
with a shining full V-shaped area above, the upper fourth sub-
shining black, the balance ochreous pollinose. Upper section of
224 Wisconsin Academy of Sciences, Arts, and Letters.
front three-fifths as wide as long, and one-half as wide as one
eye measured at the middle ; ocellar triangle twice as wide as
long, very short. Face with almost white pile; front on lower
half, or slightly less, yellow pilose, the upper half with black pile.
Occiput yellow pollinose above, greyish pollen on the lower two-
thirds and cheeks; pile yellow above, white below; no black
occipital cilia. Antennae shining black; the third joint red¬
dish, but broadly reddish brown above, sub-orbicular, a little
longer than wide ; arista reddish.
Mesonotum opaque black, the broad lateral margins and moder¬
ately broad submedian vittae bright yellow, or slightly greyish
at ends, the latter pair tapering slightly but again broadened
before the scutellum. Pile thick, yellow, becoming white on the
lower portion of the pleura ; scutellum translucent reddish
luteous with narrow black base and much coarser black pile than
on the pleura, the immediate margin with pale hairs.
Coaxe black, grey pollinose, whitish pilose, the pile of the
front pair very short. Femora black; a little more than the
apical fourth of the front and hind pairs and apical third of the
middle ones, reddish; tibiae reddish, paler basally; anterior
pair broadly brownish at apex, the hind ones with ferruginous
tinge on sides of apical third, tarsi black, the middle basitar-
sus reddish. Pile of legs yellowish ; black on the upper surface
of the tarsi ; the narrow anterior lower edge of the middle femora
except basally, and beneath the hind femora, where the hairs
are short, rather bristly, and some black, fine spines towards the
outer end.
Wings strongly tinged with cinereous or even somewhat luteous
on basal half or more in front. Squamae whitish, with yellow
border and fringe; halteres pale yellow.
First abdominal segment black, grey pollinose, the lower side
pieces pale yellow ; second segment opaque black with the narrow
apex shining black ; on each side with a large pale yellow triangle
reaching base and apex broadly, more or less obtusely rounded in
the middle and moderately widely separated from each other;
they cover more than half the segment; apex of segment nar¬
rowly reddish except the sides; yellow triangles covered with
yellow pollen. Third segment with a large, basal, reddish or
orange spot on either side, reaching four-fifths the distance to
the apex and moderately broadly connected with the narrow red¬
dish apex at the sides, their inner end oblique so that their inner
Curran (& Fluke — Helophilus and Allied Genera 225
posterior end is broadly rounded but is covered by yellow pollen
which projects inwards and forwards; posterior margin of the
spots a little oblique; spots yellow pollinose; narrow preapical
fascia shining black; short basal fascia, median vitta and short
facia behind the spots, opaque black. Fourth segment with the
narrow lateral margin and broader apex shining reddish yellow,
the segment elsewhere shining except for the broad, ochreous
pollinose lunules, which are narrowly bordered with opaque
black, rounded inwardly and the inner ends somewhat nearer the
base than the outer ends. Pile black on the black areas, except :
the first segment, base of second, lateral margings of fourth, in¬
creasing in width behind, and the lower margins of the second
and third segments which are otherwise black pilose behind the
middle of the second segment. Pile on pale areas pale yellow;
on the genitalia dense, long, yellow.
Female. This is very similar to the femal of pendulus but the
wholly pale posterior tibiae will separate it.
Holotype. Male, Farewell Creek, S. Saskatchewan, Sept. 1907,
No. 2224 in the Canadian National Collection, Ottawa ; allotype —
female, Savonaski, Naknek Lake, Alaska, July, 1919, (J. S.
Hine) ; in Hine Collection.
Pa/ratypes. Male, Farewell Creek, S. Saskatchewan, Sept.
1907, in collection of C. W. Johnson; six males, six females,
Savonaski, Naknek Lake, Alaska, July 1919, in collections of
J. S. Hine, C. W. Johnson and C. L. Fluke.
This species has very much the habitus of H. pendulus L. but
the differences pointed out are sufficiently striking to warrant its
separation. It might be confused with H. obscurus Lw. but that
species lacks the sharp points to the spots on the third segment
and those on the fourth are only slightly lunulate, much broader
and not oblique.
Helophilus hyhridus Loew
Plate VI, figure 11
H. hyhridus Loew, Stett, Ent. Zeit., 1846, 141 — Europe.
H. novaescotiae Macq., Dipt. Exot., Suppl. II, 60, 1847 — N. S.
H. lafitarsis Hunter, Can. Ent., XXIX, 134, 1897 — Minn.
Pace with median black stripe ; pale markings very extensive
on second and third segments of abdomen; in the female much
less extensive, those on third segment sharply rounded on inner
ends.
15
226 Wisconsin Academy of^ Sciences^ Arts^ and Letters.
Length, 13 to 15 mm. Male. Face yellow pollinose, a large
triangle on the cheeks, narrow oral margin, broad median stripe,
supra-antennal triangle and upper third of front shining black ;
facial vitta with pollen on upper fourth. Front and back of head
bright yellow pollinose. Pile yellowish ; on the cheeks and lower
orbits white; on upper third or less of the front, (except the
vertex) black. Face prominent below, convex on a little more
than the lower half, the concavity however occupying more than
the upper half. Antennae black; arista red. Ocellar triangle
wider than long, moderately large.
Mesonotum opaque black, the wide lateral vittae and moder¬
ately wide dorsoeentral vittae, yellow pollinose; pleura with
greyish yellow pollen. Pile of thorax luteous. Scutellum trans¬
lucent brownish yellow, darker at base and sides, black pilose
except on base and apical margin.
Legs black; broad apices of the anterior four femora, basal
third of the front, whole of the middle tibiae, basal fourth of the
hind pair and whole of middle basitarsi, reddish yellow; pos¬
terior femora with preapical reddish band which is broadly in¬
terrupted postero-dorsally. Femora black haired above and be¬
low, the bases usually more or less broadly pale haired ; the tarsi
and under side of hind tibiae also with black hairs; legs else¬
where with yellowish hair.
Wings cinereous hyaline. Squamae whitish with bright yellow
border and pubescence and yellow fringe. Halterers yellow.
Abdomen with three pairs of reddish yellow spots or the
last pair yellowish grey pollinose. First segment shining
black with the sides broadly yellow. Second segment chiefly
dull reddish yellow, with the apex reddish on at least the median
half and opaque black markings as follows: a broad median
vitta which narrows slightly behind where it is connected with
a rather narrow subapical fascia which never extends to the
lateral margins except rarely as a ferruginous stripe, the median
vitta expanded so as to be slightly wider than the black of the
mesonotum. Third segment with the apex reddish preceded
by a shining black fascia which usually reaches the lateral mar¬
gins, and an opaque black median vitta which narrows rather
sharply to the basal fourth of the segment where it connects
with a transverse basal spot which is hardly three times as
long as wide and more or less rounded on its outer ends posteri¬
orly ; the yellow ground is supplemented by some greyish white
Curran <& Fluke — Helophilus and Allied Genera 227
pollen so that the pale spots are rather acute inwardly at their
middle; pale spots occupying three-fourths or slightly more of
the segment at the sides. Fourth segment shining black, with
a very narrow, incomplete basal sub-opaque black fascia which
is usually narrowly connected with a median opaque black tri¬
angle lying between the inner end of the large yellowish grey
or greyish yellow pollinose spots which occupy more than the
basal half of the segment sub-laterally, are convexly narrowed
inwardly and slightly arcuate in front; posterior margin and
narrow lateral margin reddish. Genitalia ferruginous. First
two segments yellow pilose except the broad posterior margin of
the second and the side margins on the posterior half ; posterior
third of third segment (except the pale spots) and the whole of
the lateral margin at base, black pilose ; fourth segment with the
shining black area and apex black-haired ; pile of abdomen else¬
where pale yellowish. Second and third ventral segments usually
all yellowish.
Female. Front with a band of yellowish pollen in front of the
ocelli interrupting a median black vitta; front black pilose in
the middle below and on upper half except the vertex.
Abdominal markings much as in ohscurus but the spots are
larger, being more truncate inwardly on the second segment,
more acute on the third and not at all oval on the remaining seg¬
ments, but transverse, narrow and narrowly separated from each
other.
Specimens are before us from Northern Ontario, Manitoba,
Saskatchewan, Alberta, British Columbia and the Northwest
Territories. Other localities include Nova Scotia, Minnesota and
North Dakota.
The male is readily distinguished by the very large pale ab¬
dominal markings. The female resembles ohscurus much more
than would be expected and is probably most easily separated by
the wholly pale third ventral segment, this being largely shining
black in ohscurus, but the abdominal fasciae are quite different
in shape upon close examination. The irregular median vittae
of ohscurus also readily separates the two species. European
specimens from England and Denmark do not differ.
228 Wisconsin Academy of^ Sciences j Arts, and Letters.
Helophilus obscurus Loew
Plate Y, figures 1, 2
H. obscurus Loew, Cent. lY, 74, 1863. (Colo.)
Face Avith median black stripe ; abdomen of male with 3 pairs
of, of female 4, transverse, more or less oval spots, the first pair
large and somewhat triangular, the last pair slightly lunulate
in male.
Length 10 to 13.5 mm. Male. Face orange yellow, the middle
line reaching almost to the antennae, oral margin and cheeks,
shining black. Yellow portions of face and front to well above
the suture densely yellow pollinose, the pollen on the face usually
pale. Face moderately excavated on upper two-thirds, slightly
retreating on lower portion. Front narrow above, on the upper
half or more of the narrow portion, except the venter, black pilose,
elsewhere with yellow pile. Occiput yellow pollinose on about
the upper half, the lower half with greyish yellow pollen and
white hair. Antennae black, the third joint largely or almost
wholly red, the upper and apical margins brown. Arista
orange yellow.
Mesonotum deep, black, only sub-shining in some lights, the
side margins and a pair of broadly separated, sub-median vittae,
yellow or pale yellow, the latter narrowed behind the suture,
sometimes interrupted on the suture, usually as before the scu-
tellum, but there are always two wider spots just before the scu-
tellum, and Avhen the vittae are entire they widen behind. Pleura
greyish yellow pollinose. Scutellum translucent brownish, its
apex reddish; thorax yellow pilose, the scutellum with short,
abundant black pile except on its margins.
Legs black. Apices of front four femora, basal third to half
of the front tibiae, all the middle tibiae, their apices sometimes
slightly darker and the first two joints of the middle tarsi yellow
or reddish yellow; hind femora sometimes with their bases, a
rather narrow sub-apical band, the basal third of the hind tibiae
and sometimes a narrow median band yellow or reddish.
Wings hyaline or tinged with yellow. Stigma brownish.
Squamae pale yellow, with yellow fringe. Halteres pale yellow.
Abdomen opaque black, the narrow apex of the second, apex
of third and broad apex of fourth segments, shining; with two
pair of yellow pollinose spots. Sides of first segment pale yellow ;
Curran <& Fluke — HelopMlus and Allied Genera 229
second segment yellow^ except the broad apex, broad median
stripe and base, the black on the latter rather rounded laterally
and broadly separated from side margins. Base of third seg¬
ment with a rather broad, sub-oval spot on either side, more
widely separated in the middle in front, widely so behind, their
posterior margin convex, the spots narrower laterally; at their
inner ends posterioly with a small spot of yellow pollen. Fourth
segment with a yellow pollinose broad, slightly lunulate spot on
either side sub-basally, their outer ends separated from the
front and lateral margins, their inner ends separated from each
other by about half the distance between the inner ends of the
preceding spots. Apices of segments successively more widely,
more or less evidently, reddish. Pile black; yellow in front of
the hind margin of the yellow markings on each segment and on
the sides of the fourth segment. G-enitalia usually black; red¬
dish in only one specimen.
Female. Front somewhat narrowed above ; black pilose only
on the upper half, rarely almost to the antennae, the middle line
thinly pollinose, and appearing black; pollen on face and front
yellowish white.
Thoracic vittae slightly wider, and rarely interrupted.
Abdomen more shining, the yellow spots narrower, more trans¬
verse, those on the third segment only reaching the sides in front.
Pollinose fasciae on fourth segment more oval, scarcely concave
in front, not lunulate; the basal comers of the fifth segment
with transverse, pollinose, triangles, narrowly separated in the
middle.
Over 100 specimens from New York, Quebec, Ontario, Mani¬
toba, Alberta, British Columbia, Idaho, Wisconsin and Colorado.
The reddish third antennal joint, and three pairs of inter¬
rupted fasciae at once distinguish this species from all others
known to us.
Mesembrius Bond
Plate VII, figure 40
The genus Mesembrius does not occur within our range, but
is confined to the Old World, being especially abundant in
Africa and Asia. It is characterized by the carinate face and
the contiguous or narrowly separated eyes in the males and the
230 Wisconsin Academy of Sciences^ Arts, and Letters.
stigma simulates a cross-vein. tSuperfieially the species includ¬
ed in this genus display the greatest resemblance to Helophilus,
but the nature of the stigma forms a ready means of separation.
We are unable to state the number of species which should be
included under the genus, as many species described under
Helophilus really belong here. Bezzi lists thirteen species from
the Ethiopian region, some of which also occur in Asia, and one
species is recorded from Europe. It is probable that most of the
species of Helophilus listed from Southern Asia and the East
Indian Islands belong here.
Another character, which seems to be constant, which may be
used to distinguish this genus from aU others examined is the
presence beneath the hind tarsi of numerous nodulate hairs.
Sometimes these hairs are found only at the basal fifth, at other
times more than half the tarsal pad is composed of them. In
the nine species of Mesembrius which we have examined this
character is very well marked and quite easily seen, while we
have found no trace of it in any other genera.
Parhelophilus
Girschner, Illustr. Wachenschr., II, 604, 1897
The genus Parhelophilus shows greater affinities to Helophilus
than do most of the allied genera previously included under the
latter. The abdomen is shorter and more compact, the stigma
simulating a cross-vein. The apical production of the hind
tibiae is absent, thus readily distinguishing the genus from
Lejops. The ocellar triangle, while very broad, is seldom almost
equilateral and is always comparatively much smaller than in
Asemosyrphus. Genotype, Helophilus frutetorum Fabr,
Parhelophilus was not well understood by its founder and
could not possibly stand upon the characters used by Girschner.
The genus was separated from Helophilus because of slight
differences in the squamae, but as these intergrade to a greater
or less degree it is evident that the character is of little value,
Lejops lunulatus was also included in the genus by Girschner,
but as that species has the hind tibiae produced, it is quite evi¬
dent that it cannot belong here. Considered in every way, this
is a very compact, homogeneous group and it seems that the
isolation of the species into a distinct genus is very desirable.
Curran & Fluke — Helophilus and Allied Genera 231
TaUe of Species — ^Males
1. Hind femora with a sub-basal projection below,
which may be broader than long and is beset
with black setulae _ _ 2
Hind femora at most somewhat rounded sub-basally__ _ 3
2. Femoral process long, bearing an apical fan-like
fringe of black hairs (Europe) _ frutetorum Fabr.
Femoral process short, covered with shorter,
black setulae _ _ divisus Loew
3. Abdomen deep black, with three pairs of hoary
spots, the ground color reddish only beneath
part of the spots on the second segment _ porous Walk.
Abdomen with yellow ground color on second
and third segments, the spots never hoary _ 4
4. Mesonotum without distinct vittae ; hind femora
somewhat arcuate _ ohsoletus Loew
Mesonotum distinctly vittate; hind femora not
arcuate _ _ _ 5
5. Large dark species, with 3 pairs of widely
separated yellow abdominal spots, unusually
narrow front and the basal two-fifths of hind
femora yellow _ anniae Brim
Species of usual size; if with three pairs of
broadly separated abdominal spots, the pos¬
terior femora are wholly black; front of
male wider _ 6
6. Posterior basitarsus with black pile above, the
hairs mostly as long or slightly longer than
the height of the segment bearing them _ integer Lw.
Posterior basitarsus with appressed short yellow
hair above _ 7
7. Apical third or more of anterior tibiae black;
posterior basitarsus usually with five or six
short, black bristles on apical half of lower
anterior surface _ rex n. sp.
Anterior tibiae wholly yellow or at the most
ferruginous on apical fourth ; bristles on
hind basitarsus, if present, reddish, rarely
one or two black _ 8
8. Front broadly black pilose immediately above
the antennae; the orange color occupies all
the sides of the second and third abdominal
segments (Europe) _ _ versicolor Fabr.
Front with only a few black hairs above the
antennae; posterior black bands of second
and third segments entire _ laetus Loew
232 Wisconsin Academy of> Sciences, Arts, and Letters.
Females
1. Abdomen black with hoary pollinose spots _ porcus Walk
Abdomen not with hoary spots _ 2
2. Abdomen opaque black with whitish yellow
spots or fasciae _ j. _ rex. n. sp.
Abdomen with reddish yellow or orange mark¬
ings, the pale color predominating _ 3
3. Mesonotum without distinct vittae _ ohsoletus Loew
Mesonotum with distinct vittae _ 4
4. Face scarcely convex almost straight, slightly
produced below _ divisus Loew
Face distinctly concave above, prominent on
lower half _ 5
5. Hind femora all yellow below _ 6
Hind femora largely black below; near their
base with long process (Europe) _ frutetorum Fahr.
6. Hind femora broadly yellow below on only
lower surface (Europe) _ versicolor Fabr.
Hind femora with an anterior and posterior
black spot, sometimes joined above (N. America) _ 7
7. Face but little prominent below _ laetus Loew
Face quite strongly prominent on lower part _ integer Loew
Parhelophilus porcus Walker
Plate VI, figures 18, 19
Helophilus porcus Walker, Cat. British Dipt. Part III, 551,
1849 (Can.)
Keadily recognized by the deep black abdomen with hoary
segmental spots.
Length 9 to 11 mm. Female. Face yellow, thickly wholly
pale yellowish pollinose, narrow oral margin and the cheeks
shining brown or blackish. Pile pale, sparse, long. In profile
concave above, almost perpendicular below, but still slightly
receding, the lower part very slightly convex, strongly produced
downwards, rather obtusely conical, the lower portion of the
face scarcely less produced than the antennal prominence.
Front wide, the sides subparallel, slightly narrowed above the
middle, but again slightly widened to the posterior angles of
the eyes, which are rounded off. Front shining black in ground
color on the lower third, except the reddish supra-antennal W.
densely covered with greyish or greyish yellow pollen, which is
always darker on its upper margin ; pile black abundant,
Curran & Fluke- — Helophilus and Allied Genera 233
moderately long. Ocellar triangle very broad, the mediam
ocellar line scarcely longer than half the posterior ocellar line.
Occiput shining black, the orbits broadly greyish pollinose.
Pile black on upper half, longest above, short whitish or
yellowish on lower half. Antennae ferruginous reddish, third
joint sub-orbicular; arista concolorous, long, thickened on
basal half.
Mesonotum densely grey pollinose, shining laterally behind
the base of the wings; in front on the basal three-fifths with a
median, geminate black stripe, slightly divergent posteriorly, a
short median stripe just before the scutellum and a sub-lateral
stripe not quite reaching either end, opaque black. Pleura
black, moderately greyish pollinose, but not entirely obscuring
the ground color. Pile on the mesonotum before the suture,
less widely so in the middle, and on the pleura except the
mesopleura, pale yellowish; elsewhere, black. Scutellum tran-
luseent reddish, its base and sides black ; pile black, on the sides
pale yellowish.
Legs black, immediate apices of front four femora, narrow
bases of their tibiae and sometimes of the hind ones, and first
joint of their tarsi, yellow; all the tibiae with a rather narrow
yellow median reddish band; front four tarsi brown, their last
joint reddish. Hind femora greatly swollen, somewhat latter ally
compressed, their tibiae arcuate ; front legs slender. Coxae pale
haired, the front four with a dense apical ciliation of black hair.
Front four femora with black above, with pale pile below ; hind
femora with shorter, pale pile and black stouter hairs below;
tibiae with brownish and brownish yellow short, appressed pile,
front tarsi wholly pale, the others chiefly black haired.
Wings cinereous hyaline, stigmal spot brown. Squamae
white, with white fringe. Halteres reddish yellow.
Abdomen opaque black, the side margins shining, narrow
posterior margin of second to fourth segment, broadened in the
middle, on the last two, triangularly, shining black with a grey¬
ish sheen, owing to thin grey pollen; the same segments each
bearing a roundedly rectangular, transverse spot on either side,
these successively nearer the base of the segment, the front ones
wider laterally, and more narrowed inwardly, all the spots
widely separated, less widely so from the lateral margin. The
first segment is all hoary except a roundish lateral opaque black
spot; the fifth has a narrow longitudinal basal triangle opaque
234 Wisconsin Academy of, Sciences, Arts, and Letters.
black, the lateral margins broadly, the apical narrowly shining
black, elsewhere greyish. Pile whitish basally, on the hoary
spots and side margins opposite and the fifth segment except the
middle basally.
Male. Front strongly widened posteriorly, where it is
practically twice as wide as at narrowest point; yellowish grey
pollinose below the depression; with very scattered greyish
pollen in front of the ocelli except at the sides; pile wholly
black.
Thorax with a wider black triangle immediately before the
middle of the scutellum, the black median vitta only interrupted
on its posterior half by the narrow grey line, and narrowly
separated from the posterior black triangle. Legs deeper black,
the pale markings on the tibiae reduced.
Outer half of the hoary spots on second abdominal segment
situated upon reddish ground, the lateral margin shining black,
the reddish tinge carried broadly obliquely outward to almost
reach the margin broadly behind the basal incissure. Lateral
margins shining black opposite the spots on third segment and
extending quite to the base ; fourth segment more broadly shin¬
ing laterally. The hoary spots are a little smaller, being less
elongate.
Female, Montreal, Que., June, 15, 1906 ; Female, Banff, Alta.,
July 4, 1922, (C. B. D. Garrett) ; Male, Gimli, Man., June 13,
1923, (A.. J. Hunter). We have seen specimens from Mc-
Diarmid, Ontario, near the type locality ; New York, and New
England States.
The male was received after the description had been com¬
pleted in other respects.
Parhelophilus rex new species
Plate VI, figures 15, 16
Belongs to laetus group, but much darker, the pale thoracic
stripes narrow; opaque black, the abdomen with three pairs of
yellow spots.
Length, 10 to 11mm. Male. Face translucent yellowish,
densely pale yellowish pollinose, the middle line below and the
oral margin broadly shining; narrow oral margin and cheeks,
shining black ; in profile upper third moderately deeply concave,
the lower half slightly receding, scarcely convex, not as promi-
Curran & Fluke — Helophilus and Allied Genera 235
nent as the antennal prominence, only moderately produced be¬
low the eyes and therefore obtuse ; pile rather long, sparse, yel¬
lowish. Front narrowest at lower third, where there is a distinct
transverse depression, distinctly widened to the posterior angle
of the eyes ; lower third yellowish, obscurely blackish above, but
wholly yellowish pollinose. Above moderately shining black.
Pile yellow before the suture, thick, black above. Ocellar tri¬
angle line to outer ocellus greater than the distance between the
anterior ocellus and that line. Occiput shining black with a
greyish bloom, the orbital margin broadly grey pollinose. Pile
yellow, rather long above, the cilia black; on the lower three-
fifths short, white, becoming longer and more yellow below and
on the cheeks. Antennae reddish; third joint broader than
long, obliquely truncate apically, longer below ; arista brownish,
long, tapering, slightly thickened on basal fourth.
Mesonotum opaque or sub-opaque black, with five greyish vit-
tae, the median one slender, slightly widened posteriorly but fad¬
ing out at about the posterior third ; on either side of the middle
line, not quite half way to the lateral strip, is a rather narrow,
slightly irregular complete stripe of the same color, which is
characteristically broadened inwardly before the scutellum as a
long apical triangle, lateral margins broadly of the same color,
but subshining, as the pollen is evidently less abundant, the stripe
broadening anteriorly and spreading over the humeri to reach
the sublateral stripe obscurely. The three median stripes are
narrowly bordered by a deeper black. Pleura black with thin
pollen of same color as on sides of mesonotum. Pile of thorax
tawny or bright reddish yellow; on the pleura less abundant.
Scutellum translucent reddish, the base broadly blackish; pile
black, on the narrow base and apex yellowish.
Femora black, their apices broadly reddish, on the hind ones
with a subapical more or less complete band; hind tibiae with
the narrow base yellowish, a reddish band at the middle ; front
and hind tibiae with narrow base and apex yellow, the sub-
basal fourth on outside posteriorly black or blackish, so that
apical fourth wholly and the sub-basal fourth on outside pos¬
terior black or blackish, so that there appears to be a reddish
median band from some views, but the larger portion of the ti¬
biae is reddish. Hind tarsi blackish, the front four reddish with
somewhat darker apices. Pile of femora yellowish black on
front four posteriorly on apical half or less; similar but all ap-
236 Wisconsin Academy of^ Sciences^ Arts, and Letters.
pressed on tibiae and tarsi, but black on exterior surface of the
front four tibiae.
Wings cinereous hyaline or faintly yellowish about the middle.
Squamae white, with pale yellow border and white fringe.
Halteres pale yellow.
Abdomen opaque black; with three pairs of spots and other
yellow markings; first segment, the hind margins of the follow¬
ing segments, triangularly expanded in the middle on the third
and fourth, and the narrow lateral margins, shining. First
segment with an obscure yellow spot in front sublaterally.
Second segment with very large lateral sub-triangular spots,
occupying the lateral margin on the anterior half, thence cut off
by a narrow black triangle, their posterior edge slightly oblique
and slightly convex, their inner ends rounded, the anterior inner
edge concave, reaching the base of the segment broadly,
moderately separated from the posterior margin at sides, and
broadly separated in the middle, pale yellow; the inner corners
of the spots are situated at the middle of the segment, and some
whitish pollen projects slightly onto the black. The spots on
the third segment are somewhat similar in shape, but are
narrowly separated from or just reach the side margins, are
much shorter, occupying about two-thirds the length of the
segment, their inner anterior margin not concave, the posterior
comers of the spot supplemented by a white pollinose projection
which is transversely oval in shape, making the inner ends of
the spots appear acute. Posterior margin of the segment yellow
on lateral sixth; in the middle with a greyish yellow pollinose
triangle. Fourth segment with the posterior margin yellow, the
sides and large basal triangles, moderately separated, gray or
greyish yellow pollinose, the inner ends of the triangles cut off
from the base of the segment by a black triangle; a small
yellowish pollinose triangle in the middle apically. Pile black;
on the side margins and on the yellow spots, whitish or pale
yellow.
Female. Face less sharply concave above, the lower portion
more rounded and scarcely receding ; pile blackish. Front wide,
the sides sub-parallel, slightly widened below; pile all black.
Thorax with black pilose band between the wings, not reach¬
ing sides.
Legs more piceous than black, the hind tarsi brownish above,
yellowish below.
Curran & Fluke — Helophilus and Allied Genera 237
Abdominal spots more rounded inwardly the first pair con¬
vex posteriorly, the spots on the two following segments more
transverse, narrower as they occupy only the basal half of the
segment and are more rounded inwardly.
The third segment is not yellow posteriorly, but together with
the fourth, is narrowly reddish. Fifth segment greyish yellow
or yellow pollinose, with a basal, longitudinal median triangle,
black or brown.
Described from male, Macdiarmid, Ont., June 29, 1922, (N. K.
Bigelow) ; male, N. W. Bay, Lake George, N. Y., Aug. 20, 1920 ;
female, Algonquin Park, Ont., Aug. 14, 1903, (B’. M. Walker) ;
1 female, Macdiarmid, Aug. 21, 1923 (Bigelow), 2 males and on
yellow water lily bloom.
Type in Canadian National Collection. No. 2223.
This species is very much darker than usual in general ap¬
pearance and has only narrow pale thoracic vittae. It is readily
distinguished from the groenlandicus group of Helophilus by
the small stigma.
Parhelophilus anniae Brdmley
Helophilus anniae Brimley, Ent. News XXXIV : 278, 1923
(N. Car.)
^ ‘ Face, antennae, arista, and lower part of front light yellow,
the latter with yellow pile, narrower portion of front (between
the eyes above) dark brown with dark pile. Thorax with two
narrow submedian yellow stripes and a somewhat broader one on
each side, scutellum paler than thorax. Legs : front and middle
femora dark brown for about basal two-thirds, yellow at apex,
front and middle tibiae and tarsi mainly yellow, hind femora
yellow on basal third, then abruptly black to apex, hind
tibiae and tarsi black. Abdomen : second, third and fourth seg¬
ments each with a pair of yellow spots, these wider externally,
and narrower internally, those of each pair widely separated,
those of second and fourth segments broadly reaching the sides
of those segments, those of third segment nearly or quite sep¬
arated from the side of the segment by the ground color, a small
yellow spot on the apex of fourth segment. Length about 11 mm.
“A female taken at Aberdeen, North Carolina, May 10, 1922
. . . appears to be the same species, the only differences be¬
ing that it is slightly larger, the submedian stripes on the thorax
238 Wisconsin Academy of^ Sciences^ Arts, and Letters.
are a trifle wider, there is no apical yellow spot on the fourth
segment, and the pile on the front above the antennae is largely
black, instead of yellow.
‘‘This species looks somewhat like a small H. mmilis, but dif¬
fers from all species I know or could find descriptions of, in the
following combination of characters, — wholly yellow face, an¬
tennae, and arista ; three pairs of widely separated yellow spots
on abdomen, and in the hind femora being yellow at base, and
black in middle and apex. ’ ’ 'IBrimley.
This species evidently belongs to the genus Parhelophilus, al¬
though we may be mistaken in this. Dr. Aldrich in a letter
states that “the hind femora is yellow on the basal two-fifths
and provided with a distinct swelling bearing a cluster of black
spinules at the point where the tip of the tibia comes in contact
with it. In this respect it resembles frutetorum, an European
species.
Since the above was written, the type has been examined. The
species is very distinct from any yet described, being nearest
related to P. rex from which it differs in the narrower, almost
parallel-sided upper part of front, larger size, etc.
Parhelophilus divisus Loew
Plate VII, figure 25
Helophilus divisus Loew, Cent. IV, 78, 1863. (D. C.)
Allied to laetus but the face is not at all convex and not prom¬
inent below the middle, the abdominal spots transverse and
smaller.
Length, 9 to 11 mm. Male. Face reddish yellow, yellow
pollinose, the narrow oral border and cheeks shining black to
light brownish; in profile shortly receding below the antennae,
thence slightly produced to the tip of the oral margin, almost
straight, only slightly produced below the eyes. Pile rather
long, yellowish. Front black or brownish, wholly brownish grey
or greyish yellow pollinose, less thickly so at vertex; sides par¬
allel on upper half, somewhat divergent below, the median de¬
pression distinct and with a slightly anterior bow. Pile all
black or sometimes yellow before the depression. Ocellar tri¬
angle very broad, the lateral half of the posterior ocellar line
one and one-half as long as the median ocellar line, the lateral
Curran & Fluke — HelopMlus and Allied Genera 239
ocelli separated from the eyes by little more than their own
width. Occuput grey pollinose, yellowish above, the pile yellow
on upper half, white and shorter below. Antennae red, third
joint large, slightly wider than long, sub-orbieular, the lower
corner longer; arista brownish, tapering.
Mesonotum apaque greyish yellow or yellowish grey, more
greyish in old specimens, with three opaque black stripes, the
median one geminate on anterior two-thirds, narrowed somewhat
on posterior third, the dividing line usually very slender, the
lateral black stripes about half as wide as the median one, all
the stripes not quite reaching the front, but the median reach¬
ing the posterior margin, the lateral narrowly separated from
it. Pleura grey pollinose. Pile bright yellow. Scutellum red¬
dish yellow, the base and sides black. Pile black, on the margin,
longer, yellow; sometimes all yellow.
Femora black, the apical third of the front four and fourth of
the hind ones, reddish, the latter black at the apex above, or the
hind ones may be all reddish; front four tibiae and tarsi, ex¬
cept a black spot behind, extending to the anterior margin above
in the middle, reddish yellow, the apices of the front tibiae and
the last two joints of the front four tarsi brownish. Hind tibiae
on apical fourth, their tarsi wholly, and an obscure sub -basal
band brown or brownish. All the femora may bear an obscure
reddish spot below basally. Hind femora somewhat thickened
at the base below with a rounded, large tubercular swelling
bearing abundant short black setulae.
Wings cinereous hyaline, the stigmal spot small, brown.
Squamae whitish with yellow border and fringe. Halteres yel¬
low.
Abdomen opaque black ; the fourth segment chiefly grey polli¬
nose ; with yellow spots on second and third segments. Second seg¬
ment with the sides broadly reddish yellow on the anterior two-
thirds, expanded inwardly on the middle of the segment as a
broad, inwardly rounded projection, its anterior edge concave to¬
wards the side, posterior margin slightly convex, the spots medi-
anly separated by the length of one of the arms. Third segment
with its outer end produced forward to the anterior angles, and
touching the base of the segment broadly ; narrow lateral margin
of the third and forth segments and the narrow apices of the
second to fourth segments yellow. Fourth segment greyish
yellow pollinose except a median transverse fascia occupying
240 Wisconsin Academy of. Sciences, Arts, and Letters.
half the width of the segment, a basal, apically convex fascia
occupying one-third the width and a connecting middle stripe.
First segment wholly grey pollinose except a roundish black
spot on either side ; posterior margin of the second segment more
broadly in the middle, of the third still more broadly and more
angularly in the middle, grey pollinose. Pile yellow, black on
the incomplete posterior margins of the second to fourth seg¬
ments, successively more broadly so. Ventral plates black with
yellow apices, the sides of the second broadly yellow.
Female. Front moderately narrowed above, its sides parallel
on upper third. Pile less abundant, black. Pile on thorax all
shorter.
Front femora with only an incomplete broad brownish basal
band behind, their tibiae obscure brownish apically; last two
joints of front four tarsi as in male. Middle femora with small,
brownish sub-basal patch behind. Hind femora with a broad,
incomplete, brownish band about the middle or a little beyond,
on postero-dorsal surface produced apicad; sub-basal brownish
band on hind tibiae complete.
Abdomen broader, the yellow spots narrower, more elongate,
the second pair lunulate, their sides more parallel; fourth seg¬
ment with narrower black fasciae, the basal one extremely nar¬
row. Fifth segment with a very small black basal triangle, oth¬
erwise grey pollinose.
Male, female, Summit, New Jersey, June 8, 1913, (F. M.
Schott) ; two males, female, Westville, N. J. May, (C. W. John¬
son) ; female, Plymouth, Indiana, (M. E. Smith) ; female, Phil¬
adelphia, Pa., July 13, (C. W. Johnson).
H. divisus may be distinguished from all species except porcus
and rex by possessing on the antero-ventral surface of the pos¬
terior basitarsus a row of short black bristles. As these bristles
are confined to the apical half in rex, that species is eliminated,
while porcus has hoary abdominal markings and wholly black
hair on the upper surface of the posterior tarsi.
ParKelophilus frutetorum Fabr.
Plate VI, figure 14
Most nearly allied to P. divisus Loew, but the process on the
hind femora is nearly twice as long and is strongly fringed with
Curran & Fluke — Helophilus and Allied Genera 241
a fan-like row of moderately long, black hairs, the posterior fem¬
ora are all black or brown except the apex, the usual spot above
the apex. Face slightly more convex below; yellow thoracic
vittae narrower, more yellow. Front black pilose above.
The female is similar to the male except in abdominal mark¬
ings and absence of the femoral process. The inner half of the
lunule on third segment is formed by a pair of oval, transverse
pale yellow pollinose spots, rather narrowly separated.
These characters are sufficient to show the relationship of the
two species and also the chief differences between them. In ad¬
dition the posterior tarsi lack the row of short black bristles on
the anteroventral surface of the basitarsi, thus agreeing with all
other species in the group (except divisus, porcus and rex)
which lack the bristles on the basal half of the posterior basi-
tarsus.
Specimens — male, Germany; male, female, Denmark.
P. frutetorum is not known from North America.
ParhelopMlus laetus Loew
Plate VI, figures 20, 21, 22
Helophilus laetus Loew, Cent. IV, 77, 1863. (N. Y.)
Helophilus aureopilis Townsend, Trans. Am. Ent. Soc., XXII,
51, 1895. (Mich.)
Hind tibiae not produced on inner apex ; abdomen chiefiy red¬
dish yellow; thoracic vittae usually rather broad, slightly di¬
vergent on anterior third, lower portion of face rather promi¬
nent and shorter than the concave upper part.
Length, 8.5 to 10 mm. Male. Face yellowish, thickly yellow
pollinose except the cheeks in front and the broad oral margin
to the tip; cheeks black on posterior two-thirds; in profile the
face is deeply concave on more than the upper half, the lower
portion prominent and gently convex, retreating below. Front
yellow before the suture, thickly yellow poUinose ; above, black¬
ish, but thickly yellow pollinose, less thickly so across the ocelli.
Ocellar triangle much broader than long, the median ocellar line
about equal in length to the lateral half of the posterior ocellar
line. Occiput blackish, wholly greyish or yellowish pollinose.
Pile of head all yellow except a broad black band between the
ocelli and depression. Antennae orange colored, the arista black
apically.
16
242 Wisconsin Academy of Sciences, Arts, and Letters,
Mesonotum dull olivaceous black, with the side margins broad¬
ly and sub-median, moderately narrow, stripes yellowish or grey¬
ish yellow pollinose, all connected along the anterior slope by a
transverse band of pollen, which is wider laterally, so that the
lateral black fasciae are more widely separated from the front
margin than the middle one. The middle yellow stripes are
very slightly bowed outwardly and are usually slightly broad¬
ened anteriority, and posteriorly and narrowly connected behind
to the lateral stripes. Pleura grey or yellowish pollinose. Scu-
tellum yellow or reddish yellow, the base narrowly and the an¬
gles, black. Pile of thorax bright yellow, the disk of the scutel-
lum with black pile.
Legs reddish yellow, with black markings. Front four femora
with the basal third or fourth black, rarely with the markings
almost obsolete; hind femora with a broad median black band,
interrupted below, sometimes greatly reduced and leaving only a
small spot in front and a large one behind ; also with a small spot
at apex above. Hind tibiae with sub-basal and apical rather
broad black bands, usually the middle half yellow, but this some¬
times reduced to one-third; hind tarsi wholly black. Pile all
yellowish.
Wings cinereous hyaline or slightly yellowish tinged. Squa¬
mae white, with yellow border and yellowish white fringe. Halt-
eres yellow.
First abdominal segment grey pollinose except behind the
anterior corners, usually broadly yellow laterally, sometimes but
little so. Second segment opaque black, the apex narrowly red¬
dish ; the sides broadly, expanded as a large triangle occupying
more than half the segment, its inner ends rather truncate or
broadly rounded, posterior margin slightly oblique, the anterior
oblique and concave on its middle portion ; the yellow is rather
narrowly separated from the reddish hind margin and the black
emits a triangle forward along the margins for a short distance.
The black on the third segment is limited to a basal transverse
fascia on the middle half, a narrow, almost complete sub-apical
fascia, usually moderately broadly jointed to the anterior one in
the middle and emitting a small triangle forward laterally.
Sometimes the middle connection is obsolete and the basal fascia
narrow and not convex posteriorly. Following segment with al¬
most similar markings but the posterior fascia does not reach
nearly to the laterally margin and is not or scarcely broadened
Curran & Fluke — Helophilus and Allied Genera 243
laterally. The yellow between the fasciae is often wholly due
to greyish yellow pollen ; in pale specimens the basal fascia and
middle connection may be obsolete, in which case the pollen is
yellow. Genitalia with yellow or yellowish grey pollen, apices
of last two segments and often the narrow lateral margins, red¬
dish. Sternites black, the apices yellow pollinose. Pile of ab¬
domen yellow, black on apices of segments.
Female. Very similar but still more reddish or orange. Front
wholly black pilose, more or less yellow in ground color above the
antennae, yellow pollinose as in male.
Mesonotal pale vittae slightly wider. Legs showing same varia¬
tion.
Abdominal segments shorter so that the yellow fasciae are
narrower and the black lateral projections of the fasciae usually
extend much further forward, sometimes quite to the base of the
segment. The black may be obsolete in the same way as in the
male. Fifth segment black, the lateral margin and posterior
margin, much more broadly in the middle, yellow, the whole cov¬
ered with yellow or greyish yellow pollen except a median longi¬
tudinal basal dash.
This description is drawn from over fifty specimens from the
northern States and Canada as far West as Manitoba and Wis¬
consin. The range extends over practically all the United States
and Canada east of the Rockies.
Parhelophilus versicolor Fabr.
Plate VI, figure 17
Perhaps nearest P. integer Loew but the posterior femora
sub-basally on the antero-ventral surface is somewhat roundedly
swollen and bears a tuft of rather conspicuous long yellow pile,
the face is scarcely less prominent below, the front is black pilose
above and again just above the antennae and the yellow of the
second and third abdominal segments reaches broadly to the
hind margin.
These differences are well marked and constant in the male.
The female has a wider front, as in laetus, but the posterior
femora are black on the basal half except below, the face is much
less prominent, and the black posterior segmental fasciae
straighter.
Specimens: male, female, Denmark; male, Europe.
244 Wisconsin Academy of^ Sciences, Arts, and Letters.
Parhelophilns integer Loew
Plate VI, figure 23
Helophilus integer Loew, Cent. IV, 76, 1863. (N. Y.)
Allied to laetus but the front of the male is narrower and
wholly yellow pilose; the face is less deeply concave and less
prominent in both sexes ; the cerci of the hypopygium are short
and obtuse.
Length, 9 to 10 mm. Male. Face yellow, transluscent, densely
yellow pollinose; the cheeks in front and posterior part of oral
margin shining; checks black behind a point a little before the
suture. Face in profile slightly concave above, gently convex
below, but little prominent on lower half. Front yellow, above
the suture brownish, densely yellow pollinose even above, though
the ground color shows through in the region of the vertex.
Middle ocellar line longer than the lateral half of the posterior
ocellar line. Occiput greyish or greyish yellow pollinose. Whole
head yellow pilose. Antennae orange; arista black on apical
half.
Mesonotum densely yellow pollinose, appearing more greyish
in old specimens, with broad median and slightly narrower sub¬
lateral opaque blackish stripes, the lateral ones narrowly
separated from the posterior, widely so from the anterior
margin, slightly narrowed at the ends, but still rounded, the
median one entire, but thinly pollinose on the anterior slopes.
Pleura yellowish pollinose. Pile all bright yellow. Scutellum
yellow, dull, the base black ; pile black, yellow on margin.
Legs yellow, the basal fourth or less of the anterior four
femora, sometimes only faintly so, a broad median band inter¬
rupted below, on the hind femora, sometimes all but base and
broad apical band; a spot above the apex, the apical third and
broad basal band of their tibiae and their tarsi wholly, black or
brown. Hind femora only moderately swollen, with black bris¬
tles below on apical half. Pile of legs all yellow.
Wings cinereous hyaline. Squamae whitish, yellowish on
apical portion, the fringe yellow. Halterers yellow.
Abdomen predominatingly yellow or reddish yellow; opaque
black, the first segment grey pollinose except behind the anterior
angles, the sides more or less yellow. Second segment reddish
yellow, with a large lunulate black basal fascia not reaching the
sides, and broadly connected with an entire, narrow, sub-apical
Curran & Fluke — Helophilus and Allied Genera 245
opaque black fascia which projects slightly forwards at the sides,
behind this a shining black fascia with a triangular median pro¬
jection forward, the immediate apex narrowly reddish ; the apex
in the middle with a greyish yellow pollinose spot. The yellow
is cut off slightly obliquely at the sides, the inner arms broadly
separated, rather truncate, the anterior margin oblique, concave
about its middle. Black on third segment limited to a rather
narrow, subtriangular basal fascia on middle half or more, con¬
nected to a sub-apical opaque fascia situated before a narrow
shining band, the apex narrowly reddish and greyish yellow
pollinose, the pollen triangularly produced in the middle, the
lateral margins black on apical third. Fourth segment very
similar, the pale markings composed of greyish yellow or yellow
pollen, the spots narrower, very narrowly separated, the
posterior black fascia occupies only three-fifths the width of the
segment, its arms slightly enlarged, side margins broadly, the
apical margin widely greyish yellow pollinose. Genitalia with
similar pollen. Sternites black, their sides more or less broadly
and apices narrowly, yellow. Pile yellow, rather short, black on
apices of segments.
Female. The female is difficult to distinguish from laetus,
but the face is only slightly concave above and much less pro¬
jecting below; the front narrower and appearing longer, the
hind femora are almost straight on their lower surface, not dis¬
tinctly convex, and the fifth abdominal segment is nearly half
black pilose, not all yellow.
Specimens examined : male, 3 females, Newark, N. J. ; 2 males,
1 female, Westville, N. J. ; male, female, Long Branch, N. J. ;
2 males, female, Martha’s Vineyard, Mass.; male, female. Well-
fleet, Mass. (C. W. Johnson). Male, female, Arlington, N. J. ;
female, Caldwell, N. J., (A. Nicolay).
H. integer was originally described from New York, and has
never been authentically recorded since, except from the New
England States. It is very limited in its range, but probably
will be found in all the New England and adjacent states. H.
laetus has a wide distribution, but apparently does not occur in
New England.
246 Wisconsin Academy of Sciences, Arts, and Letters.
ParhelopMlus obsoletus Loew
Plate VI, figure 24
Helophilus obsoletus Loew, Cent. IV, 75, 1863 (Hud. B.
Terr.).
Allied to integer but the thorax is not vittate.
Length 9 to 10 mm. Male. Face transluscent yellow, covered,
except the very broad oral margin, and cheeks, with pale yellow
pollen, cheeks brownish black, the oral margin yellow before the
jowls, in profile moderately concave on about the upper half,
the lower half moderately prominent. Front yellow before the
suture, densely yellowish pollinose; above the suture black,
densely yellow pollinose, except just at the vertex, where it is
thinly so. Ocellar triangle wider than long, the lateral ocelli
narrowly separated from the eyes, the median ocellar line as long
as the distance from its base to the orbit. Occiput olivaceous
blackish with greyish yellow pollen. Pile of head wholly yel¬
low. Antennae orange yellow, arista concolorous; third joint
elliptical.
Mesonotum rather dull olivaceous blackish, broad submedian
and broad lateral margins, diffusely slightly more shining or
very lightly yellowish. Pleura greyish pollinose. Scutellum
impure yellowish, rather dull, only the corner blackish. Pile of
thorax yellow, only on the disc of the scutellum black.
Anterior four femora black, on basal half, the hind ones black
except the apical third; anterior four tibiae and tarsi wholly
yellow; the hind ones black, with the bases and a broad median
band yellow. Hind femora with a black dorsal spot just before
apex. Pile wholly yellow. Hind femora only moderately en¬
larged, slightly arcuate.
Wings cinerous hyaline or slightly yellowish. Stigmal spot
yellow. iSquamae whitish, the border yellow, fringe yellowish
white. Halteres yellow.
First abdominal segment sub-opaque black, the sides and
anterior angles grey pollinose. Second segment opaque black,
the apex shining and narrowly reddish; the sides except the
apex broadly yellow, expanded as large obtuse triangles occupy¬
ing nearly half the length of the segment near their inner end,
situated chiefly beyond the middle, their posterior margin slight¬
ly oblique, the anterior concave on the middle portion. Base of
Curran & Fluke — Helophilus and Allied Genera 247
third segment with an incomplete, medianly broader and an in¬
complete subapical transverse fascia, these narrowly connected
on the middle line, opaque black, elsewhere reddish yellow.
Fourth segment with a median stripe, expanded, as basal fascia,
and a median incomplete transverse fascia, brownish red, else¬
where reddish, the sub-basal yellow pollinose fascia incomplete
and narrowly interrupted. Pile yellowish; broadly black on
apices of segments, the black not reaching the sides on the last
segment. Sternites black.
Female not before us. Loewis description shows only the
following variations: base of scutellum black; thoracic stripes
very narrow, yellow; each segment with a triangular spot in
middle contiguous to the yellow apex; femora less extensively
black, on the front four reduced to two basal spots; the hind
ones with a large black spot behind.
Male, Solon Springs, Douglas Co., Wis., July 7-15, 1909.
The male specimen described here was compared with the type
and is undoubtedly the same species. The presence of vittae on
the thorax of the female is to be expected, but they are extremely
faint in the type, as in the male. The nearest ally is integer
which also has a yellow haired front but differs in having wide
thoracic vittae. The median ocellar line is much longer in com¬
parison to the posterior ocellar line than in laetus. The two
allied European species are more closely related than laetus but
both have the upper half of front black haired.
Unrecognized Species
Parhelophilus flavifacies Bigot, Annales Ent. Soc. France, pt.
22, 344, 1883.
Hine has suggested that this is the same as H, divisus Loew.
It seems likely that it is either divisus Loew or anniae Brimley
but it is not possible to decide from the description. The type
is in the collection of Mr. J. E. Collin, Newmarket, England.
Asemosyrphus Macq.
Bigot, Bull. Soc. Ent. France, 1882, p. CXXVIII; Annales,
1883, 228.
Giglio-Tos, Ditteri del Messico, II, 18, 1893.
Distinguished from allied genera by the remarkably large,
ocellar triangle, more or less tuberculate face, non-pollinose
248 Wisconsin Academy of Sciences, Arts, and Letters.
thorax and usually broad, flattish abdomen. The stigma
simulates a cross vein, the hind femora are swollen, and slightly
compressed. The front in both sexes is wide, the head flattened
above and somewhat triangular.
Three species are accepted as belonging to this genus but it is
possible that hicolor is a synonym of mexicanus. The material
before us does not permit of an opinion on the matter.
Key to species of. Asemosyrphus
1. Hind tibiae wholly reddish yellow (Mexico) _ hicolor Big.
Hind tibiae largely black _ 2
2. Face obtusely short conical ; abdomen reddish or
with grey lunules; mesonotum olivaceous,
with three germinate opaque black Yittsie-mexicanus Macq.
Pace produced as long acute cone; abdomen
without grey lunules, all black; mesonotum
almost unicolorous (canadensis Curr.) _ willingi Smith
Asemosyrphus mexicanus Macq.
Plate VI, flgure 13
,Syn. : Helophilus mexicanus Macq., Dipt. Exot., II, 2, 64;
Loew, Cent. X, 55, (Helophilus polygrammus) ; Osten
Sacken, West, Dipt., 338; Bigot, Bull. Soc. Ent. Fr.,
1882, ( oculiferus, nigroscutatus, flavocaudatus ) ; Bigot,
Ann. Soc. Ent. Fr., 1883, 350, (oculiferus), 351,
(nigroscutatus and flavocaudatus); Williston, Syn.,
186 (Helophilus); Biologia, Dipt., Ill, 68 (Hel¬
ophilus) ; Giglio-Tos, Boll. R. Inst. Torino, VII, No.
123, 1892 (griseus); Ditt. del Messico, II, 20, 1893.
Mesonotum olivaceous, with six slender opaque stripes, the
median ones joined to the inner ones of the lateral two ; abdomen
partly reddish or with grey lunules.
Length, 9 to 10.5 mm. Male. Variable, the following de¬
scription from a single specimen will cover all parts except the
abdomen. Face wider than either eye, the middle line, a broad
stripe on the cheeks, and narrow oral margin, shining black,
elsewhere densely greyish yellow pollinose; in profile, perpen¬
dicular, the large, elongate tubercle occupying nearly half the
face, above it, moderately concave. Front wide, the sides slight¬
ly divergent above, but sub-parallel, slighty convex laterally on
upper portion, covered with slightly more greyish pollen than
Curran & Fluke — Helophilus and Allied Genera 249
the face; a shining spot immediately above the antennae, the
vertex broadly less thickly pollinose, the pile yellow, black just
above the depression. Ocellar triangle very large, the posterior
ocelli near the posterior angles of the eyes. Occiput densely pale
greyish pollinose, yellow pilose. Antennae piceous, the third
joint black, sub-circular, a little longer below; arista reddish.
Mesonotum shining olivaceous with a median and sub-lateral,
moderately broad, geminate opaque black stripes, the lateral
ones abbreviated anteriorly the median one extending to the
posterior third; the middle ones connected with the inner
lateral stripe by a sub-rectangular spot on apical third, and
joined to a narrow median line which runs from the apex of the
abbreviated stripe to the scutellum; the outerline of the lateral
stripe is triangularly broadened just before and behind the
suture and there is a slender, oblique straight line rising at the
posterior end of the lateral stripe and running in a line towards
the humeri but it does not reach the suture. Pile of thorax
yellowish. Pleura bluish black. Scutellum more or less yellow¬
ish or reddish, the base blackish, its pile all pale.
Legs black; narrow apices of femora and bases of tibiae
yellowish, the pile all yellowish. Hind femora greatly swollen;
largest in the middle, more or less laterally compressed, their
tibiae rather evenly arcuate.
Wings cinereous hyaline, more cinereous anteriorly. Squamae
greyish white with yellow border and fringe. Halteres yellow.
Abdomen variable ; in the specimen before us as follows : first
segment shining black, with a greyish sheen, an opaque black
spot laterally, the sides yellow on posterior half. Second and
following segments reddish, with obscure darker areas, the
second with a broad basal opaque black fascia, its posterior mar¬
gin rounded, but produced in the middle as a rounded triangle
which does not quite reach the posterior margin and is not as
long from its base as its width ; sometimes this black projection
expands almost to the lateral margin on the apical part. The
third segment is much similar but the dark markings are grey¬
ish red, leaving basal, more reddish, slightly lunule-forming
orange spots on either side, the segment wholly shining; these
triangles are overlaid with a slender whitish or yellowish lunule,
rising at the anterior angle, extending obliquely towards the
middle of the segment and curving somewhat forward at their
inner ends which are broadly separated. The base of the fourth
250 Wisconsin Academy of Sciences, Arts, and Letters.
segment has a brownish opaque arch on either side of the middle
line^ and almost completely engaging its posterior border, a nar¬
row, yellowish white lunule, the brown opaque color continued as
a more reddish opaque band across the base of the segment, the
lunules widely separated ; behind the lunules shining fer¬
ruginous. Pile short, rather appressed, yellowish, longer on
sides basally, the posterior half of the second and third seg¬
ments, not reaching the sides and excepting the narrow hind
margin, and dark basal spots on fourth segment, with short, ap¬
pressed black pile.
Female. Front wide, slightly narrowed towards the middle,
the sides almost parallel on upper portion; usually the black
pile is limited to a transverse spot on either side just above the
middle.
The abdomen is usually all black, all shining, more or less sub¬
opaque on second segment and bases of the two following, with
four pairs of grey lunules, the first pair slightly oblique, ex¬
tending inwards from behind the middle of the grey lateral
triangles which reach the base moderately broadly; the lunules
on the two following segments similar to male ; the fifth segment
bears broader, shorter lunules than the fourth, and the side
margins are narrowly pollinose. Pile as in male, the fifth seg¬
ment with all pale pile. The sides of the second segment are
often yellowish-red except behind.
Four specimens from California. Numerous specimens have
been examined. It is a common species in Mexico and has been
reported as far north as British Columbia.
Asemosyrpkns mexicanus is very distinct from A. willingi
which has a sharply conical face and lacks abdominal lunules
and thoracic stripes.
Asemosyrphus willingi Smith
Plate VI, figure 12
Helophilus willingi Smith, Proc. Ent. Soc. Wash., XIV, 118
1912; Curran, Can. Ent. LIV, 94, 1922 (canadensis).
Pace produced downwards and slightly forwards into a long
cone ; whole fiy black, mostly shining ; ocelli very widely separat¬
ed.
Length 9 to 10 mm. Male. Face and front shining black, the
sides of the face obscured by whitish yellow pollen, leaving a
Curran & Fluke- — Helophilus and Allied Genera 251
median stripe and the cheeks shining; front broadly covered
with similar pollen on the lower half, except an arch above the
antennae. Lower half of the face produced as in Helophilus
conostomus Will., but produced only a little forward, with a
long, slender, not prominent tubercle below the middle, above
which it is shallowly concave, and below which it is almost
straight to the tip of the oral margin; antennal prominence
narrowly reddish above the antennae. Antennae shining black,
tip of second joint and immediate base of third somewhat red¬
dish ; third joint opaque, whitish pubescent, subquadrate, slight¬
ly broader than long and slightly shorter than the second joint,
but longer than the first; arista bare, long, slender tapering.
Posterior orbits yellowish gray pollinose. Pile: on the face
limited to the sides, fine and whitish, cheeks bare; front with
pale yellow pile, but with a black band across the anterior
ocellus; posterior orbits with whitish pile below and yellowish
above.
Dorsum of thorax shining blackish green; a moderately wide
opaque black, median stripe not quite reaching the scutellum and
which usually has a very slender shining stripe in its middle;
on each side of this a shining stripe, obsolete before the posterior
quarter, wider than the median opaque stripe ; on the outer side
of this is a broad, opaque black, stripe, which is expanded on the
posterior quarter to join the median opaque stripe, is partly in¬
terrupted at the suture, and encloses a more or less distinct nar¬
row shining stripe; near the sides is a very narrow opaque
stripe on the posterior two-thirds, interrupted at the suture.
The shining stripes appear to have a greyish tint in some lights.
Pleura and scutellum shining blackish green, with pile of a
paler yellowish color than that on the dorsum.
Legs shining greenish black, tips of all the femora and narrow
bases of tibiae reddish (in one specimen the bases of the tibiae
are more broadly yellowish and there is also a yellowish band
just before the middle) ; hind femora considerably enlarged, the
greatest swelling near the middle; hind tibiae arcuate, not end¬
ing in a spur ; all the tarsi golden pubescent beneath.
Wings a little infuscated; veins reddish, but brownish apic-
ally, basal portion of 2-3 longitudinal vein with about seven bris¬
tles of a blackish or reddish brown color. Squamae pure whitish
with white pile; halteres brown yellow.
252 Wisconsin Academy of^ Sciences, Arts, and Letters.
Abdomen shining greenish black with a metallic reflection in
some lights, the first segment slightly grayish; second segment
with a large basal opaque black triangle, broadly separated from
the lateral margin and apex ; third segment with a small opaque
roundish or oval spot before its middle, sometimes obsolete;
abdomen elsewhere shining; no lighter markings. Pile of the
abdomen pale yellowish, but across the disc on the posterior half
of each segment with shorter black pile.
Specimens examined: 4 male, Saskatchewan, July 1907.
This species is very distinct from the only other species in the
genus reported from Canada, A. meooicamns Macq., and is readily
recognized by the absence of light abdominal markings and the
conically produced face.
Lunomyia new genus.
Plate VII, figure 27
Allied to Lejops, but the face is distinctly tuberculate in both
sexes. Small dark species the abdomen not at all yellow, nor
marked with pale, the thorax with a greyish yellow vittae. Third
antennal joint broader than long, its apex almost evenly, gently
convex. Front moderately wide in male, the sides parallel above,
in female slightly narrowed above, the ocellar triangle small.
Legs as in Asemosyrphus. Stigma simulating a crossvein. Vern¬
ation as in Lejops. Genotype : HelopJiilus cooleyi. Seamans.
Lunomyia cooleyi Seamans
Plate VII, figures 26, 27
Helophilus modestus Williston, Synopsis, 192, 1886 {nec.
Wied.) ; Seamans, Ent. News, XXVIII, 342, 1917,
{Tropidea cooleyi).
Resembles an Asemosyrphus, but the ocellar triangle is small.
Length, 7 to 8 mm. Male. Face black in ground color, except
a reddish spot on either side of the slopes below; densely cov¬
ered with yellow dust which leaves a moderately narrow, dorsally
abbreviated median stripe, the oral margin and a broad stripe
on the cheeks shining black or browinsh; pile brassy yellow,
limited to the sides below, and appearing as a conspicuous
tuft ; the side margin below and the cheeks with similar but
Curran & Fluke — Helophilus and Allied Genera 253
slightly shorter pile. In profile the face is a little prominent
below as the concavity of the upper half is deeper than that
below the long, rounded tubercle. Front almost one-fourth the
width of the head, with a deep depression just below the middle
and with a broad, less evident depression extending from it in
the middle to the apex of the antennal prominence, before the
depression densely yellow pollinose, above it rather dull black,
with a little yellowish pollen below on the middle portion and
along the whole orbits in some lights. Pile all black except at
vertex. Ocellar tubercle swollen, convex, the triangle a little
broader than long, not large. Occiput greyish pollinose, with
yellow pile. Antennae brownish black, with thin pale pollen,
the arista black, thickened and tapering on basal fourth.
Mesonotum dull black, the sides shining aeneous greyish, ante¬
riorly yellowish greyish pollinose, with two widely separated,
complete moderately slender vittae of the same color rising at the
inner ends of the pollen on the front surface, and widening be¬
hind to join the lateral shining portion; these vittae are very
thinly pollinose and are, except in fresh specimens very largely
shining. In addition there is a slender median line of the same
color on the anterior three-fourths. Pleura black, thinly grey
pollinose. Scutellum blackish, translucent brownish red apie-
ally. Pile of thorax rather conspicuous, all brassy yellow.
Legs black or brown, with brassy yellow pile, apices of the
femora, bases of the tibiae, a moderate band on their middle, in¬
complete on the front ones, yellow. Hind femora wide, somewhat
laterally compressed, widest at apical fourth, their tibiae arcu¬
ate, with a short, broad, apical lobe.
Wings cinereous, the small, stigmal spot luteous. Squamae
white, with pale yellow border and white fringe. Halteres yel¬
lowish white.
Abdomen shining greenish black, somewhat piceous on basal
angles of second segment. Second segment opaque black except
the broad sides, which expand somewhat towards the front, and
the narrow hind margin. Third segment with a medianly ex¬
panded opaque basal band on middle half. Pile all brassy yel¬
low, longer laterally and on the metallic triangles on the second
segment.
Female. Facial turbercle sharper, more acute, the part of the
face not prominent below the tubercle, but on a plane with the
upper concavity which is rather more angular. Front broader.
254 Wisconsin Academy of Sciences, Arts, and Letters.
its sides a little narrowed above, but distinctly diverging behind
the ocelli, the depression broad, with a rather large transverse
swelling on either side of a broad median foeva just above the
antennae, the pollen apparently not so abundant, the swellings
bare.
The front and middle tibiae show no indication of a pale me¬
dian band.
Abdomen as in male but the third segment is only narrowly
opaque at the base on the middle third and the metallic color is
more extensive on the second segment as it forms a broadly in¬
terrupted, broad median fascia the inner ends of the spots
rounded and the hind margin of the segment more broadly shin¬
ing. Fifth segment wholly aenous.
This description is drawn from a pair of paratypes of Trop-
idea cooleyi Seamans, from Bozeman, Montana, June 12 and
20, 1906. Specimens of this were compared with the type of
Helophilus modestus Williston and are absolutely identical in
all respects.
LejopiS Rondani
Lejops Rondani, Dipt. Ital. Prodi., II, 33, 1857 ; Verrall, Brit.
PL, VIII, 524-5 (changes name to Liops) .
Anasimyia Schiner, Cat. Dept. Eurofae, 108, 1864.
The characters of this genus are : apex of hind tibiae produced
as a scoop-like or rather acute spur; stigmal spot simulating a
crossvein; ocellar triangle rather small; front of female not as
wide as one eye, of male not over half as wide, the ocellar triangle
in female rather small, rarely almost equilateral; form rather
slender. Genotype, Mallota vittata Meig.
The genus which most closely approaches Lejops is Asemo-
syrphus which, however, never has more than a very slight pro¬
duction of the hind tibiae (mexicanns), and has a much broader
front in both sexes, very large ocellar triangle, broader abdo¬
men, and more or less distinctly tuberculate face.
When Rondani proposed the name Lejops he included only
the one species, H. vittata Meig. Subsequent authors have failed
to find characters which would separate the genus from Heloph¬
ilus and consequently several other names have been used which
included at least part of the species which we place here. Schiner
in 1864 proposed Anisimyia in his catalogue and included two
Curran & Fluke — Helophilus and Allied Genera 255
species, the first of which H. transfugus, Coquilletf^ designated
as the type species in 1910. Verrall ignored it as it was merely
a catalogue name, and while it does not concern us owing to the
fact that the type species is included under Rondani’s genus
Lejops, it would be the next available name for this group,
should Lejops ever be disallowed. Then, in 1883, Bigot described
his genus Eurhimyia basing it upon E. rhingioides n. sp., which
was later admitted by him to be the same as H. Uneatus Fabr.
We have retained this as a sub-genus, its affinities to the other
species being quite evident, the differences being insufficient to
warrant giving Eurhimyia generic rank. Girshner included H.
lunulatus in his genus Parhelophilus, with frutetorum and
versicolor but as the former falls within our conception of Le¬
jops, and we have accepted Parhelophilus for the last two species,
there need be no further explanation.
Lejops — Table of Species — Males
1. Face produced into a long, acute cone _ sub-genus Eurhimyia
Face at most obtusely conical, not greatly produced _ 2
2. Hind trochanters with a moderately long, obtuse
projection or with a smalt sharp tubercle
bearing black setulae ; tibiae ending in spur _ 3
Hind trochanters and tibiae simple, the femora
sometimes specialized _ 5
3. Process on trochanters long and obtuse; tibial
spur of moderate length- _ ^^.relictus n. sp.
Process short, tubercular - 4
4. Spur on hind tibiae long and acute _ chrysostomus Wd.
Spur on hind tibiae short, not acute, rounded
apically _ distinctus Willist.
5. Abdomen without lunules beyond the second
segment, although sometimes with the small
inner ends apparent _ , _ 6
Abdomen with very distinct lunules beyond
second segment _ 8
6. Sides of abdomen usually wholly narrowly red¬
dish beyond second segment ; a rounded
angle sub^basally on hind femora, the apical
two-fifths more swollen; abdomen practical¬
ly always with sub-median grey pollinose
spots _ bilinearis Willist.
Sides of abdomen usually wholly black, beyond
second segment, rarely partly reddish; grey
•Pr. u. S. N. M., XXXVII, 506.
256 Wisconsin Academy of Sciences, Arts, and Letters.
spots wanting; femora with less pronounced
basal angle and more evenly swollen _ 7
7. Anterior four femora practically all black
haired - ^borealis Cole
Anterior four femora all yellow pilose _ ^perfidiosus Hunter
8. Lunules strongly curving forward medianly
(Europe) - transfugus L.
Lunules scarcely curving forward, except front
pair - lunulatus Mg.
F emal e s
1. Second pair of lunules separated from side margins _ 2
Second pair of lunules reach side margins _ _ _ 4
2. Front femora black pilose _ borealis Cole
Front femora yellowish pilose _ 3
3. Abdomen not with yellow lateral and apical seg-
jaental margins _ _ perfidiosus Hunter
Abdomen with yellow segmental apices and
margins _ bilinearis Willist.
4. Hind femora red on entire surface above or
antero-dorsally _ 5
Hind femora with black patch covering part of
dorsal surface _ 6
5. Abdominal spots but little lunulate _ lunulatus Mg.
Abdominal spots strongly lunulate (Europe) _ transfugus L.
6. Femora all black below _ 7
Femora not all black below with a broad reddish
or piceous reddish elongate fascia _ distinctus Willist.
7. Tibial spur long and acute _ _ _ ^^chrysostomus Wd.
Tibia! spur not acute _ relictus n. sp.
Lejops relictus new species
Helophilus cJirysostoma (Wied.), Williston, Synopsis, 190, 1886
Plate VII, figures 31, 32
Hind eoaxe of male with long apical projection; female with
small tubercle. Abdomen of both sexes long, rather narrow, with
nearly parallel sides, usually narrowed at apex of second seg¬
ment with pale lunules, even though often chiefly reddish in
male; the female lunules grey pollinose. In the two allied spe¬
cies the coxae bear no more than tubercles, in one the tibial spur
* These species were described in the genus Pterallastes.
♦*We have not seen the female ; the distinctions given may be erroneous
but seem probable.
Curran & Fluke — HelopMlus and Allied Genera 257
is very long and slender, in the other shorter than in chrysos-
tomus.
Length, 8 to 10 mm. Male. Face yellow, yellow pollinose,
the oral margin to the angles and the cheeks, shining black; in
profile rather deeply concave on upper two-fifths, the lower por¬
tion very slightly, scarcely convexly receding; pile long, sparse,
whitish. Front with a shallow depression just before the mid¬
dle, in front of this, yellow, densely bright yellow pollinose,
above black, with dense ochreous pollen, the ocellar band brown¬
ish; pile yellow below, black above. Front narrow above, the
sides parallel, the ocellar triangle not very broad, the lateral
half of the posterior ocellar line shorter than the median ocellar
line, the ocelli separated from the eyes by about the width of
one ocellus. Occiput grey pollinose, yellowish above. Pile yel¬
low above, white elsewhere and on cheeks. Antennae ferrugin¬
ous reddish, third joint slightly longer than broad, sub-circular;
arista reddish, its apical half brown.
Mesonotum opaque black, the lateral margins moderately
broadly and sub-median, rather narrow stripes, slightly converg¬
ing posteriorly in most specimens, greyish yellow, yellow or pale
yellow pollinose, the stripes broadly connected in front, narrowly
so behind ; and never as wide as the black stripe. Pleura yellow¬
ish grey pollinose. Seutellum brownish, its apex to apical third
or even more, reddish. Pile of thorax wholly yellowish, paler on
pleura, not long, moderately abundant.
Basal two-thirds of anterior four femora, the hind ones except
the base and sub-apical ring similar; front legs reddish, the
tibiae with more or less brownish pre-apical bands, the last two
tarsal joints fuscous. Hind tibiae and tarsi black, the former
with a median band and sub-basal ring, reddish. Hind coxae
with an obtuse, rather large prominence, its length as long,
usually distinctly longer than its width, densely beset with short,
black bristly hairs. Hind femora greatly swollen, the base and
apex narrow; beneath on the swollen part, with stout, sub-
appressed, apically directed hairs, before the apex in front with
five or six stout hairs on lower surface. Spur at apex of hind
tibia moderately long, almost equal to length of third tarsal
joint, its apex rounded, from ventral view.
Wings hyaline, strongly cinereous, rarely faintly yellowish,
squamae whitish, with pale yellow border and fringe. Halteres
pale yellow.
17
258 Wisconsin Academy of. Sciences, Arts, and Letters,
Abdomen variable; first segment grey pollinose; second seg¬
ment opaque black, the apex shining, sometimes broadly red¬
dish, but usually opaque black, rarely opaque brownish in color ;
the sides apically usually more or less broadly, diffusely reddish ;
basal three-fifths on either side broadly yellow, gradually widen¬
ing behind to the middle where there is a short, usually concave
in front projection, the hind margin slightly oblique, convex on
inner half, the point of the sLrm usually appearing to distinctly
reach forward, but the arm may sometimes be short and more
robust, always rather widely separated from each other. Third
segment rarely opaque black, usually opaque brownish or dull
brick red, the apical margin always shining red, the red opaque
color variable in depth and diffuse with the darker, more brown¬
ish color. On the basal third on either side, with an overlaying
rectangular whitish pollinose spot. A yellow spot, the inner
posterior end produced inwards as a rather narrow lunule, the
inner ends reaching a little forward and slightly enlarged, the
end rounded, moderately separated from each other, the arms
sometimes obsolete basally. Fourth segment usually dull red¬
dish, the apex broadly yellowish pollinose, but the region about
the lunules may be brownish or blackish. The lunules are not
complete, and are only moderately separated from the front
margin in the middle, the arm actually reduced to two oval,
slightly oblique yellowish pollinose spots, the lateral portion
reduced to squarish spot. Genitalia red, sometimes more or less
ferruginous. Pile wholly yellow, rather short, not abundant.
Ventral segments more or less black or brown.
Female. Quite different, the thoracic stripes wider, the ab¬
dominal lunules greyish or reddish yellow. Face slightly promi¬
nent at oral margin as it is produced a little on the lower portion,
instead of retreating. Front narrowed above, but still moder¬
ately wide; brownish yellow pollinose, paler on sides below,
shining across ocelli ; pile wholly black.
Mesonotum with the greyish or yellowish vittae wider, still not
as wide as the black, the middle one usually beginning just back
of the front margin, reaching the posterior margin narrowly.
Pleura grey pollinose ; pile shorter.
Abdomen opaque black, with similar shining areas as in male,
and three pairs of yellow lunules, which are apparently always
much brighter yellow than in distinctus. First segment grey
pollinose ; second with the lunules often as in male, but the inner
Curran Fluke — Helophilus and Allied Genera 259
areas are a little stouter and less widely separated. Lunules
on third segment are much as in male hut closer to the base of
segment as the lateral portion is very short; concave in front,
convex behind, narrowly separated. A rectangular transverse
apical spot on middle of third segment, grey pollinose, the shin¬
ing apex of the fourth segment except usually a slender median
line yellowish pollinose; fifth segment wholly greyish yellow
pollinose, the middle line often darker on basal third or more.
The yellow ground is limited to the sides of the lunules on sec¬
ond segment, the narrow lateral margins and narrow apices of
the third to fifth segments.
Eolotype — Male, Orillia, Ont., June 28, 1921, (Curran) ; No.
2222, in the Canadian National Collection, Ottawa.
Allotype — ^Female, Melrose Highlands, Mass., June 18, 1911,
(J. D. TothiU).
Pwratypes — ^Male, Melrose Highlands, Mass., June 18, 1911,
(J. D. TothiU) ; two males, Orillia, Ont., June 28, 1914 and
1921, (Curran) male, Orillia, Aug. 3, 1921; two males, Law¬
rence, Kans., June 18 and 27, 1922, (Curran) ; male, Jefferson
Junction, Wis., Aug. 12, 1919, (Fluke) ; three females, Law¬
rence, Kans., June 17, female June 16, female June 15, (Cur¬
ran) ; female, North Fairhaven, N. Y., July 1, 1921 ; female
McLean Bogs, Tompkins Co., N. Y., June 30, 1921 ; female, Ar¬
lington, N. J., May 29, 1918, (A. Nicolay) ; male. Fort Lee, N.
J., June 4, 1905; two males, female, Orillia, Ont., July 20, 23.
1923, (Curran) ; female, Aylmer, Que., May 23, 1923, (Curran).
The females described above are a homogenous lot. We are
not able to definitely distinguish the females of distinctus and
relictus and do not believe that all the specimens included as
females of the former belong there. However, the characters
used seem to be fairly constant. H. distinctus has much greyer
pollen, usually greyish, the apices of the segments are never
yellow and the median black vitta extends over the front margin
of the thorax. These characters are all weak but it seems better
to employ them than to leave the female of distinctus not
recognizable.
260 Wisconsin Academy ofj Sciences, Arts, and Letters.
Lejops distinctus Williston
Helophilus distinctus Will., Synopsis, 192, 1886.
Plate VII, figures 37, 38
Very similar to relictus but evidently distinct. We believe a
comparison between the two species will serve better than a full
description.
Length, 7.5 to 9.5 mm. Male. Facial concavity not quite as
deep and slightly longer, the lower portion of the face hardly
as long as the upper ; ground color rather brownish with a large
yellow area on the slopes, but merely appearing dirty owing
to the pollen. Front dark in ground color except the W. First
two antennal joints brownish.
Thoracic vittae grey, somewhat wider, but not as wide as the
black stripes. Hind femora more broadly yellow at base, their
under side yellow or obscurely so on their whole length, the black
bristles shorter and less abundant. Hind trachanters with a
small acute tubercle bearing black hairs.
Abdomen with the same variation in color. The lunules on the
second segment do not reach greatly inwards, being only small
points and their ground color is wholly yellow except the very
small inner point, the yellow reaching more towards the hind
margin and being more oblique, while in relictus it is reddish
on the posterior part. The yellow on the sides of the second seg¬
ment reaches three-fifths to the apex and the oblique pollinose
lunule rises from its posterior end and is strongly curved basally,
the inner end ovally enlarged and always more oblique. The
sides and apex of the fourth segment are broadly pale yellowish
or whitish pollinose, the strong lunulate spots rising about the
basal third, their inner ends enlarged and much more oblique,
their bases may sometimes be considerably reduced. The lun¬
ules are all situated upon reddish or yellowish ground.
Female. Face as in male. Front dark in ground color as
in male, pile all black. Thorax greyish, the middle mesonotal
line reaching over the front slope. The femora are even paler
than in the male, and this seems to be the only distinctive char¬
acter as they are quite dark in relictus. First abdominal lunules
a little more widely separated, the ground color of the lunules
more extensively pale than in dark specimens of relictus.
Curran & Fluke — Helophilus and Allied Genera 261
We have two males, Englewood, N. J., female, Arlington, N'.
J., two females, Melrose Highlands, Mass., June 17, 18, (J. D.
Tothill).
There seems little doubt about the distinctness of the two
species , although it seems possible that there might be interme¬
diate forms. The tibial spur is a little shorter and broader apic-
ally in this species than in relictus. In an additional male from
Melrose Highlands, taken on same date as one of the females,
the trochantral tuft is longer but we cannot see that the tubercle
is more prominent, and the face is wholly yellow, the ground
color of the anterior portion of the front dirty yellowish. The
hind femora are similar in color.
Lejops chrysostomus Wiedemann
Eristalis chrysostomus Wiedemann, Auss. Zwiefi., II 174, 1830.
(Not Williston, Synopsis).
In the male, the spur on the hind tibia is very long and acute,
the trochantral process is small and acute as in distinctus, the
hind femur is much more swollen than in the allied species.
Length, 10 mm. Male. Face and front below the depression
yellowish red, the cheeks piceous brownish, the jowls reddish;
the reddish parts densely yellow pollinose, only a narrow supra-
antennal shining border. In profile the face is deeply concave
on the upper half, the lower half gently convex and slightly more
prominent than the antennal protuberance, the upper tip of the
oral margin is not noticeable and situated on a plane with the
lower border of the eyes, the oral angles rather broadly rounded
and not produced as far below the eyes as the width of the third
antennal joint. Sides of front on almost the upper two-thirds
parallel, this part of the front black, more or less densely yellow
pollinose, the ocellar region with blackish brown pollen, so that
the front is not at all shining ; the pile is moderately long, black,
on the reddish ground and face, long; yellow. Occiput black,
rather densely yellow polinose, with yellow pile, which is shining
in some lights. Antennae yellowish red, third joint sub-orbic¬
ular, slightly longer than wide, the arista brown.
Mesonotom opaque black, the side margins more broadly than
the black vittae, and broadly separated sub-dorsal stripes grey¬
ish yellow pollinose, these stripes broadly united in front, nar¬
rowly so posteriorly, the sub-dorsal stripes hardly half as wide as
262 Wisconsin Academy of, Sciences, Arts, and Letters.
the median black one. Pile short, wholly yellow. Pleura brown¬
ish or ferruginous, pale yellow pollinose, behind with whitish
pollen, the pile pale brassy yellow. Scutellum with a black, pos¬
teriorly convex base, the margin broadly yellowish ; pile all yel¬
low, long, not abundant.
Basal half of front four femora, the hind ones except the nar¬
row base and apex beyond the swollen part, yellow, the yellow
more extensive above on the hind ones, the immediate apex
brown. Front four tibiae and tarsi, basal three-fifths of hind
tibiae, except a sub-basal, broad, darker band, and the hind
tarsi, yellow, all the tarsi darker, more reddish apically, the
apical one-fourth of the hind tibiae brown. Hind femora greatly
swollen, widest about apical third, below on the apical half, with
numerous black, fairly long bristles, at the base with some short,
condensed, fine pale yellow pile, the pile elsewhere yellow. The
pile on the legs pale yellowish, on the hind tarsi chiefly black.
Tibial spur long, acute. Trochantral process short, acute.
Wings cinereous hyaline. Squamae yellow, with yellow
border and fringe. Halteres pale yellow, the knob somewhat
darker.
Abdomen probably variable, but always less blackish than in
allied species. First segment blackish, more brownish laterally,
densely grey pollinose. Second segment with a pair of longi¬
tudinal pale yellow triangles on the basal two-thirds laterally,
widened posteriorly, the anterior inner margin oblique, so that
it leaves a large, basal, longitudinal opaque or sub-opaque black
triangle, the apical portion of which has the sides parallel ; this
portion limited anteriorly by a rounded angle, before which the
margin of the black triangle is gently concave ; the base of the
black triangle is broadly separated from the side margins. Be¬
hind the yellow triangles the color is shining reddish, but is
separated from the yellow by an opaque, rather ferruginous
fascia. Third segment dull reddish, the apex broadly shining,
on the sides with a long basal yellow longitudinal triangle, the
front border broadly, a narrow post median fascia and slender
connecting middle line dull ferruginous. Fourth segment all
red or with two or three diffuse darker areas, the apical third
shining, elsewhere opaque, the sides on the dull portion broadly
white pollinose.' Usually there is a broad, whitish pollinose
lunule occupying most of the yellow triangles on the second seg¬
ment and the inner arm of a lunule is indicated by an oblique,
Curran & Fluke — Helophilus and Allied Genera 263
oval whitish pollinose spot on either side of the middle line of
the fourth segment.
Female. (Translation from Wiedemann). Antennae rusty
3'ellow, face and front yellow pollinose; occiput blackish.
Thorax deep black, the vittae pale yellow; pleura with shining
greyish hair. Scutellum black, the apex somewhat transparent
yellow. First abdominal segment whitish grey pollinose; sec¬
ond deep black at the base, on each side with an oblique white
band, at the apex rusty yellow, so that the band forms the figure
of a goblet; third to fifth segments rusty yellow, on the base
of each on either side whitish, the third also with two smaller,
the fourth with two larger, whitish transverse spots; the seg¬
mental incisures whitish. Wings shorter than the abdomen, yel¬
lowish, with smalt black-brown stigma. Femora black, with
rusty brown apex, the hind ones very much swollen ; fore tibiae
and tarsi rusty yellow; hind tibiae rusty brown, merging into
chestnut brown, the apex black; hind tarsi black, their first
joint brown.
The description is drawn from male, Washington, D. C., May
17, 1912 ; and male, Petersburg, Chesterfield Co., Va., June 1,
1917.
It will be seen at once that the female described by Wiede¬
mann is not conspeeific with the specimens included by Willis-
ton under either chrysostomus (Wied.) or distinctus Will, as
the color of the abdomen is much more reddish than in any spec¬
imens we have examined. It is necessary to separate the species
on color because Wiedemann did not mention the trochantral
prosscess or tibial spur. The color of the legs applies better to
the two males which we have than to other males.
Lejops lunulatus Meigen
Helophilus lunulatus Meigen, Syst. Beschr., Ill, 370, 1822.
(Europe) .
Helophilus hamatus Loew, Cent. IV, 79, 1863. (Hudson Bay
Terr., Canada).
Plate VII, figures 29, 30
Helophilus hamatus Loew is a synonym. Abdomen tapering,
with three pairs of fascia; scutellum normally all pale haired;
264 Wisconsin Academy of, Sciences, Arts, and Letters.
fourth abdominal segment with opaque fascia beyond lunules in
female; face prominent below.
Length, 8 to 11 mm. Male. Face yellow, densely white pol-
linose, cheeks an doral margin to the base of the angles, shining
black ; pile long, fine, whitish ; in profile concave above, the lower
half or third more produced, almost or quite as much as the an¬
tennal prominence, very slightly more prominent above, scarce¬
ly convex, the lower third of the face lying below the eyes, the
buccal margin oblique, straight, the face thus forming an obtuse
cone. Front yellow below the depression with dense whitish
or whitish yellow pollen, depression not deep; above the de¬
pression the ground color black, yellow pollinose the sides nar¬
rowly opaque black, the upper fourth shining. Pile pale yel¬
lowish before the suture, black above. Lateral half of posterior
ocellar line about equal to distance from lateral ocellus to eye.
Occiput grey pollinose, more yellowish above, with yellow pile,
whitish on sides and cheeks. A few black, short hairs above be¬
hind the eyes. Antennae reddish, arista slightly darker, its
apical half brown, tapering on basal third.
Mesonotum sub-opaque black, usually with a narrow, opaque,
middle line; the side margins rather broadly greyish yellow
pollinose, the pollen extending broadly inwards along the front
margin to form the base of the rather slender dorsal pollinose
stripe of the same color, the stripes slightly irregular, somewhat
broadened behind, narrowly connected along the posterior mar¬
gin with the lateral stripes. Pleura rather thickly grey polli¬
nose. Scutellum translucent dull reddish, the base and corners
black. Usually the pile is all bright yellowish, but the scutellum
is sometimes black pilose on its disc.
Legs reddish; front femora on more than the basal half, ex¬
cept on the broad frontal margin, sometimes less so, middle on
nearly its whole length below, more widely at the base, and
a broad, elongate sub-basal spot above, sometimes more or less,
obscurely connected in front with the ventral stripe, a rather
small, sub-quadrate spot on front of hind femora, the hind ones
behind on almost their whole length, much more narrowly so
on apical third, shining black. Hind tibiae with broad sub-
basal and sub-apical piceous bands; hind tarsi wholly black.
Last three joints of front four tar-si brownish, the front ones
paler. Hind femora below with short black spinules and several
Curran & Fluke — Helophilus and Allied Genera 265
long, pale hairs, legs elsewhere pale pilose. Hind femora mod¬
erately swollen, their tibiae curved at base and apical sixth.
Wings cinereous hyaline, sometimes faintly yellowish, rarely
markedly so. Squamae whitish with pale yellow border and
fringe. Halteres pale yellow.
Abdomen opaque black, the lateral margins, first segment
chiefly, the apices of the second to fourth segments broadly, shin¬
ing. First segment thinly greyish pollinose, except a spot on
either side, the lateral margin rarely broadly yellow; second
segment with the lateral margin broadly yellow on almost its
whole length, gradually increasing in width to about the mid¬
dle of the segment, thence carried slightly obliquely inward to
form, an acute point in line with the oblique posterior margin,
the anterior margin of this projection often gently concave, the
spot chiefly greyish yellow pollinose; the pollen actually forms
the points as the yellow ground color has often only a small
interior projection. Sides of remainder of abdomen and broad
apex of fourth segment at least narrowly yellow or reddish, the
apex of the second and third segments yellow in pale specimens.
Usually the yellow portion of the lunule on the third segment
is in the form of a rectangular, longitudinal spot on the basal
half of the segment, the overlying pollen forming the arms,
which are slightly concave in front, their inner ends slightly
nearer the anterior margin, the lunules moderately separated
from each other in the middle; the base of the fourth segment
bears two large, narrowly separated greyish or greyish yellow
pollinose spots, occupying about half the length of the seg¬
ment laterally, their posterior border usually convex on inner
half ; in addition there is a rather narrow, thinly pollinose band
of the same color just behind the opaque fascia. Pile usually
all pale yellow, short, but often some black pile on apices of
third and fourth segments in the middle.
Female. Dissimilar, the thoracic stripes wide, the abdominal
lunules different in shape. Face similar. Front black, the W
yellow, yellow pollinose, dark across vertex; pile all black;
gently convex, with a longitudinal depression above ; front wide,
narrowed above, the lateral half of the posterior ocellar line
much less than ocellar-orbital line.
Mesonotum greyish yellow or yellowish pollinose, the median
stripes sometimes greyish on posterior half, somewhat widened
posteriorly, not quite as wide on most of their length at the
three opaque black stripes, the middle black stripe entire, but
266 Wisconsin Academy of Sciences^ Arts, and Letters.
narrowed posteriorly, the lateral ones broadly separated from
the front, very narrowly from the hind margin, lateral stripes
narrowed posteriorly. Rarely the front of the middle vitta is
thinly pollinose. Pleura greyish yellow and greyish pollinose.
Legs and wings as in male.
Abdomen opaque and shining as in male, the first segment
grey pollinose except for sub-lateral apical spots. In typical
specimens {hamatus Loew), there is very little yellow on the
abdomen, only the sides of the second and third segments, nar¬
row lateral margins of fourth and fifth, and their narrow apices
reddish. The lunules on the first segment are as large and as
complete, the arms a little broader than in the male, their inner
ends less widely separated. On the third segment, the lunules
are much as in male, but less convex behind, slightly oblique,
more or less, sometimes scarcely, concave anteriorly, their inner
ends closer to front margins than lateral fourth, touching the base
and lateral margin rather broadly. The following lunules are
closer to the base of the fourth segment, separated sub-laterally
by a narrow, apically convex transverse spot which reaches
narrowly quite across the base at the middle, the lunules very
narrowly separated. All the lunules greyish. There is a sub¬
oval transverse pollinose spot before the apex of the third seg¬
ment ; the shining portion of the fourth segment, widest at mid¬
dle ; fifth segment entirely with greyish or greyish yellow pollen
except a narrow stripe on basal half or more of middle line. In
some specimens the lunules are all yellow in ground color, the
sides of the abdomen wholly yellow, the apices of the second
and following segments, the fifth wholly, reddish; in such cases
there is very little shining black color. These examples present
a very different appearance, but intergrading forms are found.
Pile as in male, the fifth segment wholly pale haired.
Over 50 specimens from Quebec, New^ York, Ontario, Wis¬
consin, Manitoba and Alberta.
The male will hardly be confused with hilinearis. The female
is readily distinguished from hilinearis and perfidiosus by the
second lunules reaching the margins, the wide opaque bands and
more extensive pollen behind the lunules, etc.
Lejops lunulatus is common in May in Ontario on Caltha
bloom, but occurs also on most bloom near swampy land. Both
hilinearis and this species occur together, but the former ap¬
pears to be a little earlier.
Curran & Fluhe — Helophilus and Allied Genera 267
The synonymy of this species need not worry ns in North
America. Loew described Helophilus hamatus from the Hudson
Bay Eegion and his specimen does not differ in the least from
several specimens before us, all of which have been compared
with the type. The face shows slight variation and for a long
time we considered both species as valid, but the series before
us proves that this is not the case. The brighly marked females,
with more yellow thoracic and abdominal pollen appear very
distinct by themselves but there is a perfect series of inter¬
mediate forms. The specimens from Alberta have considerable
black pile on the scutellum, but are quite evidently not distinct,
and are really more typical of the European form.
Lejops Mlinearis Williston
Helophilus hilinearis Will., Synopsis, 295, 1886
Plate VII, figure 33, 34
Allied to lunulatus Meig., distinguished in the male by the
absence of abdominal fasciae except on second segment. The
female has the abdominal lunules of the third and fourth seg¬
ments separated from the side margins, and wholly pale haired
scutellum. In the mountain species (Pterallastes perfidiosus
Hunter) the abdomen of the male is wholly black beyond the
second segment, never margined with yellow.
Length, 9.5 to 11 mm. Male. Pace yellow, thickly white
pollinose, the narrow oral border and cheeks shining black; in
profile the upper three-fifths concave, with a slight tubercle be¬
low the concavity, the lower two-fifths slightly retreating, not
quite as prominent as the antennal base; pile long, sparse,
whitish. Front yellow before the depression, but darker im¬
mediately before it, white pollinose ; upper portion shining black,
before the ocelli moderately thickly yellow pollinose, thinly
pollinose above; depression at lower two-fifths, very shallow;
sides above slightly diverging behind, considerably diverging in
front of depression. Ocellar triangle broad, the centre of the
posterior ocellar line but little more than the width of an ocellus
from lateral ocellus, the lateral ones much more than their width
from the eyes. Pile yellow before the suture, longer, more
abundant behind it. Occiput grey pollinose, long yellow pilose
above, with shorter, almost white pile on middle, longer whitish
268 Wisconsin Academy of Sciences, Arts, and Letters.
pile below and on cheeks. Antennae ferruginous reddish, the
third joint eliptical, broader than long ; arista brown, thickened
on basal third.
Mesonotum dull blackish brown, the side margin rather broad¬
ly dull yellowish pollinose, the pollen extending inwards along
the front margin over half way to the middle line. Sometimes
the pollen is more greyish, the general color blackish, with a
slender, complete pollinose vitta sub-medianly, greyish yellow,
and a slender middle line on anterior two-thirds, shining black.
In one large specimen there is a grey pollinose dash on anterior
sixth on either side of the middle line. Pile all moderately short,
yellowish.
Color of legs somewhat variable. Front and middle femora
black on basal two-thirds or more, in front broadly red for its
whole length or with an obscure, brownish sub-basal interrup¬
tion. Hind femora with a large black spot on basal three-fifths
in front, behind all black except the base, apex above narrowly
black ; the base in front and rest of joint reddish. In one speci¬
men the hind femora are all black except on apical two-fifths
behind. Front four tibiae reddish slightly darker sub-apically ;
hind tibiae with yellowish base, the middle half and under side
on whole length, and apex, reddish, sub-apical band blackish,
sub-basal band brownish, hind tarsi black; anterior tarsi red¬
dish, their anterior margin above and the last two joints,
fuscous; middle tarsi reddish their last two joints brownish.
Posterior femora moderately enlarged, laterally compressed,
with a small sub-basal swelling below, and bearing short, rather
dense setulae below, the anterior black spot with short black pile,
legs elsewhere pale yellowish pilose. Hind tibiae curved at
base and apical fifth.
Wings cinereous hyaline, stigmal spot short, fuscous.
Squamae white, with yellow border and fringe. Halteres pale
yellow.
Abdomen shining black, sides of first segment, the second seg¬
ment, except the broad posterior margin, a longitudinally placed
sub^basal rectangular spot on the third segment, concave on its
sides, and usually an oval spot in similar position on the fourth
segment. Second segment with the lateral margin broadly
yellow, slightly widened to the middle of the segment, thence ex¬
tending slightly anteriorly inwards to a sharp point, the
posterior margin oblique and almost straight, the lateral margin
Curran & Fluke — Helophilus and Allied Genera 269
on the apical sixth only narrowly reddish yellow: the inner
portion of the spots is composed wholly of pollen, which may
sometimes be lacking, in which case the yellow forms only a
sharp angle interiorly. Beyond the second segment the lateral
margin is wholly reddish or orange, the apex of the fourth seg¬
ment more broadly so, the apex of the third segment more
piceous, but often similar in color to the margins. Sometimes
there are very faint indications of complete grey pollinose
lunules on the third segment rising just inside the yellow
margin, extending obliquely back to near the middle of the
segment thence curving inwards their last section more distinct,
situated just before the middle of the segment, moderately
separated by the opaque black spot ; as a rule only the inner ends
of the lunules are conspicuous and these may be very small.
The fourth segment bears a sub-cordate, sub-basal grey pollinose
spot in the middle, which almost entirely encloses the oval
opaque black spot; sides also broadly grey pollinose. Hypopy-
gium usually almost all blackish, rarely almost all reddish, with
thin yellow pollen. Pile all short whitish or pale yellowish, only
the apices of the second to fourth segments, successively more
widely so, with short black pile.
Female. Quite different in thoracic and abdominal macula-
tion. Face sometimes slightly more prominent below as the
tubercle may rarely be almost wanting. Front wide, sides diver¬
gent below, the ocellar triangle not larger than in male and
therefore the lateral ocelli are separated from the eyes by more
than the length of the median ocellar line. There is no trace
of a depression and the blackish ground color extends down al¬
most or quite to the red W, not so far on the sides; the lower
third is covered with abundant whitish pollen, elsewhere with
yellow pollen, the upper third almost bare. On either side of
the ocelli is a broad longitudinal depression running to the
upper third.
Mesonotum greyish yellow or greyish pollinose, with three
back vittae which are not wider than the grey ones, the median
one sometimes interrupted by a very slender middle line ; lateral
ones broadly separated from front margin, narrowly from the
hind, the middle one reaching the anterior margin in a less deep
or brownish color and distinctly reaching the hind margin.
Legs usually with slightly smaller black markings, the bands
270 Wisconsin Academy ofy Sciences, Arts, and Letters,
on the hind tibiae paler in color. Wings somewhat yellowish
tinged.
Abdomen with the sides sub-parallel, only a little tapering on
the first four segments. Grey pollinose lunules of first segment
shaped moderately as in first male mentioned in description,
the inner arms usually slightly wider, the yellow ground color
more restricted. Second segment with only slightly lunulate
spots which do not reach the side margins narrowly separated
from each other in the middle, widely separated from the base ;
fourth segment with the spots still less lunulate, very narrowly
separated in the middle and reaching the side margin narrowly ;
in the middle there is a narrow longitudinal black line which is
bordered behind the lunule by greyish pollinose except a basal
median dash and an incomplete basal fascia which is not always
visible. First segment wholly grey pollinose. Sides of second
and following segments narrowly, and narrow apices of the
third, becoming broad on the fifth, segment, yellow or reddish;
the sides also thinly grey pollinose.
Specimens examined: male. Flushing, N. Y., female, N. Y. ;
male, Aweme, Man., May 9, 1915, (N. Griddle) ; 2 males, Orillia,
Ont., April 28, 1921, (Curran) ; 2 males, Orillia, May 2, 1921;
male, Orillia, May 5, 1921, female, Orillia, May 2, 1921; male.
Ironside, Que., May 11, 1916; female, Madison, Wis., May 18
1918; female, Ithaca, N. Y., June 2, 1918, (E. G. Anderson) ;
female, Hull, Que., May 16, 1901, (Harrington) ; female,
Ottawa, June 3, 1903, (Harrington) ; female, Hull, Que., May
30, 1903, (Harrington) ; female, Madison, Wis., May 6, 1920,
(Fluke); female. University Campus, May 25, 1921; female,
same data. May 7, 1922.
This species is readily distinguished in the male by the almost
entire absence of lunules except on second segment. The female
has been confused with Umulatus but is readily distinguished by
the second lunule not reaching the sides of the segment, and
the absence of an opaque crossband behind the third fascia.
From (Pterallastes) perfidiosus the female is readily dis¬
tinguished by its usually more reddish legs and by the yellow
ground color on sides and apices of terminal segments.
Curran & Fluke — Helophilus and Allied Genera 271
Lejops perfidious Hunter
Pterallastes perfidious Hunter, Can. Ent., XXIX, 139, 1897.
Plate VII, figure 39
Closely allied to H. hilinearis Wiltist., but the mesonotum is
even darker and seldom, with thin, yellow, incomplete vittae, the
abdomen not pollinose beyond the second segment in male;
female very similar to bilinearis, but there is no yellow or red¬
dish ground color beyond the second segment. A comparison
with H. hilinearis will serve better than a full description.
Length 10 to 11 mm. Male. Front slightly wider, the upper
portion actually slightly wider than the length from the suture
to posterior angle of eyes, (in hilinearis it is exactly square) ;
face generally a little more receding below. Front usually
darker in ground color on lower portion.
Mesonotum of a deeper, duller black, the middle line and sub¬
median vittae shining black, sometimes obsolete, the later usually
shortly yellowish pollinose anteriorly; no pollen between these
vittae in front.
Femora usually more extensively black, rarely as in hilinearis.
Abdomen often with the spots on second segment similar, but
seldom projecting as far inwards, never as thickly nor extensive¬
ly pollinose. A small spot on the anterior angles of the third
and fourth segment and posterior margin of the fifth, reddish ;
opaque black markings of last two segments smaller, the
posterior one sometimes obsolete or nearly so ; posterior margin
of the third rarely obscurely reddish; rarely with a slight trace
of the inner arms of the grey lunules, on the fourth segment.
Female. Front slightly wider, and more swollen. Legs a
little more blackish, the band on the hind femora entire (not
always so?).
The chief differences are found in the abdomen. There are
absolutely no reddish markings except on basal corners of the
second segment. The abdomen wholly lacks greyish or yellow¬
ish pollen except on the first segment and on lunules. The
wholly shining fifth segment is distinctive.
Four males, Banff, Alta., May 29, 2 males, June 9, male, June
3, female, October 5, 1922, (C. B. D. Garrett) ; male, Banff,
Alta., June 23, 1909, (N. B. Sanson) ; male, Penticton, B. C.,
272 Wisconsin Academy of, Sciences, Arts, and Letters.
May 1, 1919, (E. E. Buckell). 1 male, B. C. Specimens from
Washington State have also been examined.
This species is so very like Mlinearis that it might well be
considered only a variety were the characters not so constant
and the female not so markedly distinct. The thorax and ab¬
domen are both darker than in hUinearis and the superficial
appearance is more distinct than a close examination indicates.
There has been no question about the identity of the species.
Hunter evidently had not seen hilinearis or he would not have
placed the species in Pterallastes. Notwithstanding the rather
non-Helophiline appearance, there is no indication of relation¬
ship with Pterallastes, a genus with a much shorter face, longer
third antennal joint and less deeply curved third vein. The
species described by Cole as Pterallastes borealis is very close to
perfidiosus but appears to be distinct. The mesonotum of the
female bears narrower pale vittae, the femora are largely black
pilose, etc.
Mr. E. E. Wehr has kindly compared a specimen with Dr.
Hunter ’s type in the University of Nebraska Collection and pro¬
nounces our specimen as absolutely identical in all respects.
Lejops borealis Cole
Pterallastes borealis Cole. Proc. Calif. Acad. Sc., XI, pp.
170-171.
Female : Length 13.5 mm. Very near P. perfidiosus Hunter.
Antennae black, arista brown and bare, third joint about as
broad as long. Middle of face and triangle in front of cheeks
shining black, otherwise yellowish pollinose and pilose, with
yellowish ground color under the pollen. Vertex black; frons
yellowish pollinose, with erect black pile. Occiput yellowish
grey with yellow pile and a few black hairs above eye margin.
Proboscis black.
Mesonotum opaque black, with yellowish pollen on lateral
margins and two faint yellow longitudinal stripes, widely sep¬
arated and narrow; pile rather dark yellow. Narrow base of
scutellum black, the rest translucent yellow, rather densely
yellow pilose. Pleura semishining black, with black pile.
Halteres small and yellowish, the stigmata below halteres with
short, dense, fur-like, bright yellow pile.
First abdominal segment semishining black, second with broad
Curran & Fluke — Helophilus and Allied Genera 273
yellow lateral margin, the yellow portion triangular in shape
and reaching one-third distance across abdomen ; third segment
with small round yellow spot on anterior corners; remainder
of abdomen black, the posterior portion of 2 to 5 shining, the
basal portion opaque ; a pair of linear, oblique, yellow pollinose
marks on dorsum of third and fourth segments. Pile of abdo¬
men largely yellow, black on posterior margins of third, fourth
and fifth segments. Venter semishining black, with sparse whit¬
ish yellow pile. Legs black, extreme tips of femora and bases
of tibiae ; pile of front femora and most of pile of middle and
hind femora; black; some yellowish pile; hind tibiae arcuate;
hind femora noticeably thickened and with black bristles
below, short and dense on outer third ; short pile below on tarsi
golden yellow. Wings hyaline; anterior cross-vein slightly be¬
yond middle of cell 1st A.
Male: Length 11.5 mm. Very nearly the female in appear¬
ance. Eyes widely separated, frons almost twice as wide as
ocellar tubercle ; vertex noticeably convex, a depression between
it and antennal protuberance. Abdomen lacks oblique yellow¬
ish pollinose marks on third and fourth segments. Hypopygium
quite large, blackish, with short yellow pile.
Holotype — female, No. 808, and allotype, male. No. 809, Mus.
Calif. Acad. Sci. ; collected by G. Dallas Hanna, August 20,
1920.
Type locality — St. Paul Island, Alaska.
Subgenus Eurhimyia Bigot
This subgenus contains only two species known to us, one
North American, the other European. Osten Sacken regarded
them as identical, but Williston recognized the American spe¬
cies as distinct. The two species are very close, but seem to pre¬
sent, small, though constant distinctions.
The character distinguishing the group as a subgenus of
Lejops, is the production of the face forward and somewhat
downward into a long, acute cone. The character is quite evi¬
dently of little generic value and we consider the species as
forming only a sub-genus.
The two species before us may be separated as follows:
a. Face in both sexes very evidently concave below
antennae; front of male never wholly black
18
274 Wisconsin Academy of Sciences^ Arts^ and Letters.
pilose ; abdominal pale markings a little
more extensive , lunules of female markedly
enlarged inwardly - stipatus Walk.
Face in both sexes very slightly or not concave
below antennae; front of male wholly black
pilose; abdominal spots smaller, lunnles of
female but slightly enlarged mwardlj ^.^.Mneatus Fabr.
Lejops (Eurhimyia) stipatus Walker
Helophilus stipatus Walk., List III, 602, 1849, (N. Y.)
Helophilus anausis Walk., List. Ill, 603, 1849. (Ont.)
Helophilus lineatus 0. S., Cat., 134.
Helophilus conostoma Williston, Syn., 193, 1886. (Conn.)
Plate VII, figures 35, 36
Face produced forward and slightly downward into a long,
acute cone; small compact species. There is no closely allied
species in North America and the sharply conical face will at
once distinguish it.
Length, 8 to 9 mm. Male. Face yellow, thickly yellowish
pollinose ; in profile slightly concave below the antennae, thence
obliquely produced forwards, sometimes with one or two slight
swellings; oral margin and cheeks broadly shining black, the
former in front and the latter behind more or less pollinose;
pile long, sparse, whitish. Front dark in ground color except
immediately above the antennae, the anterior half paler, and
densely yellow pollinose, the upper half largely shining but more
or less, thickly below, brownish yellow pollinose; pile some¬
times all pale before the depression, at other times practically
all black; or intermixed. Ocellar triangle broader than long,
the lateral half of the posterior ocellar line shorter than either
the middle ocellar line or the ocellar-orbital line. Occiput yel¬
lowish pollinose, sub-shining behind vertex, with a less densely
pollinose, elongate spot behind the eyes above, but broadly sep¬
arated from them. Pile yellowish, a few of the orbital ciliae
black. Antennae black, the third joint orange, sometimes some¬
what fuscous above, broader than long, elliptical; arista brown¬
ish or brownish red, tapering on basal third.
Mesonotum opaque black, the lateral margins broadly yellow
pollinose, a narrow complete line on either side of the broad
middle black stripe, yellow or greyish yellow pollinose, slightly
Curran & Fluke — Helophilus and Allied Genera 275
widened and usually greyish posteriorly ; in front broadly con¬
nected with the side margins. Pleura greyish and greyish yel¬
low pollinose. Scutellum sub-translucent reddish yellow, the
base and angles black. Pile wholly yellowish.
Legs reddish, front femora on basal third, sometimes reduced
to anterior and posterior spots, middle femora on slightly more
than basal third, on over basal half above and below, sometimes
reduced to dorsal and ventral broad stripes, apical half or less
of front tibiae and their tarsi wholly black; middle tibiae with
two darker bands, one sub-apical the other before the middle.
Hind femora with a large spot on upper middle half, a spot
on anterior surface at the apical constriction, sometimes con¬
nected with the dorsal spot, and a spot above the apex, shining
black, all sometimes slightly reduced. A sub-basal and preapical
blackish, brownish or ferruginous broad band on hind tibiae.
Hind tarsi black. Hind femora considerably swollen, with black
bristles below on apical three-fifths.
Wings more or less luteous, cinereous apically. Squamae
white, with pale yellow border and whitish fringe. Halteres
whitish yellow.
Abdomen rather robust, but not or only slightly wider than
thorax, slightly narrowed apically, opaque black, with three yel¬
low or yellow pollinose fasciae. Sides of first segment greyish poll¬
inose. Second segment with large median triangles, their posterior
ends directed obliquely outwards to join the side margin about
the apical sixth, their front margin concave, directed obliquely
forward to reach the base of the segment well inside the anterior
angles, the inner ends rounded, rather pointed anteriorly;
sometimes the lateral margin behind is reddish. Third segment
with basal spots occupying three-fourths the length of the seg¬
ment, their posterior surface scarcely convex, their inner apical
half produced inwards, truncate, the black ground color extend¬
ing lateral in front for almost a distance equal to the distance
separating the spots. Fourth segment with the side margins
broadly, the rather broad apex and a pair of broad, obtuse, mod¬
erately separated, rectangular spots on basal half, narrowly
separated from the base, yellow, or greyish yellow pollinose.
Sides of last two segments wholly narrowly reddish. Pile all
yellowish. Two middle segments of venter without black color.
Genitalia thickly pollinose.
276 Wisconsin Academy of, Sciences^ Arts, and Letters.
Female. Quite different in color of thorax and abdomen.
Front wider, slightly narrowed above, wholly black pilose.
Mesonotum with the median lines wider, but still not more
than half as wide as the lateral black stripes, except behind, the
side margins more greyish yellow, the pleura greyish poUinose.
Abdomen often largely sub-shining, the lunulate spots only
yellow in the ground color laterally, the side margin elsewhere
black. First segment wholly grey poUinose. Second with the
lateral portion of the spots narrower with almost paraUel sides
and not quite as long, the lunules rather long, moderately con¬
cave in front and convex behind, their inner ends somewhat en¬
larged and rounded, broadly separated; lunules on third seg¬
ment mostly beyond the middle, but rising before the middle, less
widely separated, but very similar in shape to the first pair;
fourth segment with the lunules narrower, before the middle of
the segment, rising at the anterior angles, less widely separated
medianly, narrowly separated from the front margin, the black
base convex, but the lunules less curved than on preceding seg¬
ment. A transverse oval spot on apex of second segment in the
middle, the posterior margins of the two following segments
more widely medianly and their rather narrow lateral margins
and the fifth segment except the base, grey poUinose; or the
base of the fifth segment may bear a triangular median basal
black spot and no fascia. In immature specimens the pollen is
more yellow.
Forty specimens of both sexes from: Quebec, New York, On¬
tario, Wisconsin, Manitoba, and British Columbia.
There has never been any question about the identity of this
species since Williston described it as H. conostomus. Osten
Sacken considered it the same as the European H. lineatus and
it was recorded as such by him. Walker ^s description is short
but this is evidently the same species described by him as H.
stipatus and also H. anausis. We have seen specimens from the
region of Martin’s Falls, the type locality of anausis, and there
is no other species which agrees at all with Walker’s descrip¬
tions. The European species is very similar.
Eurhimyia lineatus Fabr.
A pair of this species from Denmark show practically no dif¬
ferences from stipatus except those mentioned in the table, but
Curran & Fluke — HelopMlus and Allied Genera 277
there is little doubt about the two being distinct. Upon exam¬
ination they appear more distinct than can be demonstrated by
description. The differences noted by Williston will not hold
in their entirety as several of the characters are more or less
subject to variation.
Explanation of Plates.
All drawings were made with the aid of the camera lucida, and are
all drawn to the same scale. Enlarged about 10 diameters.
PLATE V.
Figure 1.
2.
3.
4.
5.
6.
7.
8.
9.
10.
HelopMlus obscurus, abdomen of female.
HelopMlus obscurus, abdomen of male.
HelopMlus trivittatus, profile of head, male.
HelopMlus groenlandicus, abdomen of male.
HelopMlus groenlandicus, abdomen of female.
HelopMlus fasciatus, head of male showing narrow front.
HelopMlus latifrons, head of male showing broad front.
HelopMlus latifrom, profile of head, male.
HELOPHILUS INTENTUS, n. sp., abdomen of male.
HelopMlus intentus, profile of head, male.
Figure 11.
12.
13.
14.
15.
- 16.
17.
18.
19.
20.
21.
22.
23.
24.
Figure 25.
26.
27.
28.
PLATE VI.
HelopMlus hybridus, abdomen of male.
Asemosyrphus willingi, profile of head, male.
Asemosyrphus mexicanus, profile of head, male.
ParhelopMlus frutetorum, hind femur, male.
PARHELOPHILUS REX, n, sp., abdomen of male.
ParhelopMlus rex, profile of head, male.
ParhelopMlus versicolor, underside of hind femur, male.
ParhelopMlus porcus, profile of head, female.
ParhelopMlus porcus, abdomen of female.
ParhelopMlus laetus, abdomen of male.
ParhelopMlus laetus, profile of head, female.
ParhelopMlus laetus, wing.
ParhelopMlus integer, profile of head, male.
ParhelopMlus obsoletus, hind femur, male.
PLATE VII.
ParhelopMlus divisus, hind leg of male.
Lunomyia cooleyi, profile of head, male.
Lunomyia cooleyi, top view of head, male.
Dolichogyna fasciata, profile of head, female.
278 Wisconsin Academy of. Sciences^ Arts^ and Letters.
29. Lejops Imiulatus, abdomen of female.
30. Lejops lunulatuSf abdomen of male.
31. LEJOPS RELICTUS, n. sp.. abdomen of male.
32. Lejops relictus, n. sp., hind femur and tibia of male.
33. Lejops bilinearis, abdomen of female.
34. Lejops bilinearis, abdomen of male.
35. Lejops (Eurhimyia) stipatus, abdomen of male.
36. Lejops (Eurhimyia) stipatus, profile of head, male.
37. Lejops distinctus, hind femur and tibia of male.
38. Lejops distinctus, profile of head, male.
39. Lejops perfidiosus, abdomen of male.
40. Mesembrius, profile of head, female.
Curran <& Fluke — Helophilus and Allied Genera
279
TRANS. WIS. ACAD., VOL. XXII
PLATE V
280 Wisconsin Academy of. Sciences, Arts, and Letters.
TRANS. WIS. ACAD., VOL. XXII PLATE VI
Curran & Fluke — Helopliilus and Allied Genera 281
TRANS. WIS. ACAD., VOL. XXII PLATE VII
THE USE OP PHOTO-ELECTRIC CELLS OF DIFFER¬
ENT ALKALI METALS AND COLOR SCREENS IN
THE MEASUREMENT OF LIGHT PENETRA¬
TION INTO WATER.^
Victor E. Shelford and Jakob Kunz
I. Introduction
In 1920 a study of light penetration into sea water in the
vicinity of Friday Harbor, Washington, was carried out with
photo electric cells by Shelford and Gail (1922). Only potas¬
sium cells without screens were used. Numerous difficulties
rendered this work perplexing and uncertain. The reading de¬
vice required too much time, was not sufficiently compact and
showed ‘‘leaks’^ in a moist atmosphere. The insulation of the
connection between the cell and cable was a constant source
of delay and trouble. The cells and especially their apertures
were too small and the current too small in proportion to light
intensities. Much time has been spent since 1920 in devising
and testing equipment designed to eliminate the difficulties en¬
countered.
In addition, a study of available glasses which could be used
in combination with cells of the various alkali metals to measure
various portions of the spectrum, was conducted. In the paper
by Shelford and Gail, calculations of the relative intensities of
different colored light at different depths, were presented.
These were understood to hold true if the water was not stained
and the suspended matter not selective in its transmission of the
various wave lengths. The same year Knudsen showed that the
sea water about Denmark is stained. He used a special spectro¬
photometer and the photographic plate.
It is the purpose of this paper to report, on the basis of ob¬
servations made on Lake Mendota at Madison, Wisconsin, on
June 13, 1925, that most of the difficulties encountered earlier
can be overcome and that photo-electric cells can be used to show
* Contributions from the Zoological Laboratories (No. 274) and the Physi¬
cal Laboratories of the University of Illinois.
284 Wisconsin Academy of Sciences^ Arts, and Letters.
selective absorption of light of different wave lengths. The
small number of readings are merely sufficient to demonstrate
the feasibility of the method.
Methods and Equipment
The batteries and resistance units were carried in a box which
protected them from moisture. Burgess B batteries were con¬
nected to telephone jacks mounted in a bakelite panel in such
a way that solid plugs could be used to connect battery cells
or groups of cells together and give any desired voltage (figs.
1 and 2). The voltage could be read quickly by means of
a small Weston volt meter with between forty and fifty thousand
ohms in series which reduced the readings to exactly one-fifth
and rendered the voltage stable during the readings. The panel
was so wired that a photo-electric cell could be connected
in the circuit by inserting a two pole plug attached to the cell’s
cable. The reading instrument was connected into the circuit
in the same manner. The two arms of these plugs were con¬
nected with telephone jacks. All conductors were carefully in¬
sulated except on the contact surface. One purpose of using
the jacks and plugs was to make possible the covering of the
self-cleaning contact surfaces with vaseline and the complete
separation and drying of all parts in case leaks occurred in a
moist atmosphere. Two Daven tubular resistance units, one of
250,000 and the other of 500,000 ohms were available in the set ;
changes could be made by shifting a plug.
The instrument used for the reading was a Rawson single
pivot (‘‘Acrid”) Micro-ammeter with two ranges of 200 and 20
microamperes respectively. This instrument behaved remark¬
ably well in a small launch 30 ft. in length and 1 ton displace¬
ment. Even with a strong wind, it offered no serious difficul¬
ties in reading, thus exceeding our expectations.
The cable used was “oko cord,” made by the Okonite Com¬
pany. This consisted of two No. 18 stranded conductors with
the usual cloth covered rubber insulation, imbedded in a rubber
covering 10 mm. in diameter. This cable gave about one seventh
as much leakage under high voltage, as the braid covered cord
used in the marine work. The leakage compared with minimum
current through the photo-electric cells was almost negligible.
The weight of the cells and accessories was carried by the cable.
Shelf ord & Kunz— Photo-Electric Cells in Light Pentration 285
To make this possible the two wires of the cable were separated
and the outer casing was removed for a distance of four feet.
The device shown in figure 3 was used to keep water out of the
outer casing. This was also constructed so as to permit the use
of wire supports to the cell and accessories. The separate wires
were further insulated by greasing them with vaseline and pull¬
ing tight fitting, pure gum rubber tubing over them.
The form of the cells was changed (fig. 4). The spherical
portion was enlarged to 6 cm. in diameter and only the lower
half of the surface sensitized. This gave a large surface much
more independent of the angle of incidence than was the case
of cells used before. The connections with the cable were
effected by means of mercury cups (fig. 4) . The rubber insula¬
tion at the end of the cable wires was removed for about % inch
and the end dipped into solder. The wire was placed in the
mercury in the cup (at first through a cork which later was dis¬
pensed with) and covered with melted vaseline. The wire was
tied in place to prevent injury to the vaseline insulation.
The cells were fitted into section of heavy brass tubing with
threaded ring tops to hold glass screens in place and solid brass
bottoms (fig. 5). Slots were cut in the brass tube to fit the two
arms and filling tube of the cell. A large rubber stopper was
roughly hollowed out to fit the cell and a quantity of soft rub¬
ber (automobile tire putty) placed in the cavity. The cell was
then pressed into place and tied down to the brass bottom with
strong cord passed through holes in the bottom plate. This
gave the cell a perfect fitting bottom on which to rest. The
brass plate was wired to the protecting cage (fig. 5) and the
large brass tube set over the cell assembly and fastened to the
bottom by means of screws. The cells gave from 120 to 220
microamperes in sunlight with 85 volts and 250,000 ohms in
series. This voltage and resistance were usually used ; 100 and
120 volts were used only between 10 and 15 meters in one series
with the potassium cell. The eurrent-light-intensity curve for
these cells was essentially a straight line.
Attempts to make cells in colored globes proved difficult or too
expensive to undertake, at least before preliminary use was
made of ray filters. Green and ruby tubing presented to us by
the Corning Glass Works discolored upon heating and green,
red and blue electric light bulbs obscured the interior and ren¬
dered the making of the cells very difficult. Many desirable
286 Wisconsin Academy of Sciences, Arts, and Letters.
glasses cannot be moulded into such forms or fused with other
glass. With the proper moulds, suitable bulbs could be blown
from clear glass and some of the colored glasses; hemispherical
caps could be made from the nonworkable glasses and cemented
over clear cells whenever the expenditure for moulds is justified.
This would increase the available aperture of the cells here re¬
stricted by the brass cylinders. Ideally the cell should be
equally sensitive from all points above a horizontal plane.
The ray filters used were plates of glass, ground, polished, and
fitted to the tops of the brass tubes. While a number of glasses
were prepared, only the following were used:
1. Blue D, which greatly reduces the long wave lengths leav¬
ing blue predominating, was used to reduce the sensitivity of
the potassium cells to colors other than blue. See Fig. 6. The
curve for the cell with blue glass is the product of the light
transmission curve of the glass and the absolute sensitivity
curve of the cell.
2. Noviol C (Corning) was used to restrict the rubidium cell
to green and yellow light. (Fig. 7).
3. Noviol A (Corning) was used to reduce or eliminate the
violet and near ultra violet, but with indifferent results.
(Fig. 8).
4. Corning high transmission yellow (G. SOB) was used to
restrict the caesium cell to yellow and orange. Corning G 24
(a red pot glass used in railroad signal lights) was used to re¬
strict the caesium cell to orange (fig. 8).
The only difficulties encountered were in connection with the
use of the glass ray filters. When the cells were lowered, bub¬
bles of air remained below the glass, making it necessary to tip
the baskets to a sharp angle. This forced the mercury against
the vaseline and caused leaks in two cases. This difficulty can
be remedied by properly grinding places in the edges of the
glass so as to allow the air to escape without admitting light.
The leaks were remedied by renewing the vaseline with some
melted over an alcohol lamp on the deck of the launch. One
cell was removed and replaced by another under the same con¬
ditions.
III. Results.
The results are shown in tables 1, 2, and 3 and fig. 9, where
percent transmitted by each meter calculated for the sun at
Shelf ord & Kunz — -Photo-Electric Cells in Light Penetration. 287
zenith, is shown. It was hoped that readings with the pyrlim-
nometer might be made, but the failure of a cable prevented
this.
Table I and figure 9 indicate a sharp difference between the
transmission of yellow light (caesium cell and yellow screen)
(see figs. 6 and 8) and of blue light (potassium cell and blue
screen). This shows that something in the water absorbs the
shorter wave lengths. Figure 6 shows that the blue glass cuts
off chiefly the longer wave length measured by the potassium
cell alone. A comparison of the figures for the clear potassium
cell and the same with a blue screen indicates that the absorp¬
tion of the violet and blue is greater than of the green. Again
figure 7 shows that the Noviol C cuts off the blue range of the
rubidium cell and should increase the percent transmitted which
is shown to be the case in table I. The case of Noviol A over
the caesium cell, is not necessarily discordant. No determina¬
tions of the transmission of the glass used were made in the vio¬
let, because of the weakness of violet light in the equipment
available. It appeared to transmit 80-90% of all wave lengths.
The curve in figure 8 is from the Bureau of Standards Publica¬
tion cited. When yellow glass was used the short wave lengths
were excluded and the transmission coefficient increased.
In table 2 another series covering two meters is shown. The
average transmission coefficient for the combinations of cells and
glasses was determined for pure water. The coefficients found
were divided by the pure water coefficient .972, .966 and .853,
respectively, giving the transmission coefficient of the suspended
and dissolved matter. Caesium with a red screen is possibly
discordant on the basis of Pietenpol’s observations which sug¬
gest that orange may be absorbed least, but the current was so
small that it may not have been accurately measured. This
table again indicates that violet and blue are absorbed by the
dissolved or suspended matter.
Table 3 shows the transmission coefficients for the cells given
in table 1, divided by the pure water transmission. Here again
the increase in absorption toward the blue end is indicated.
The results, however, are essentially qualitative as a deter¬
mination of the transmission coefficient of the suspended matter
is necessary to determine the degree of staining. PietenpoPs
results indicate (table 4, p. 577) that the transmission coefficient
of the stain in Lake Mendota water is 88.7% for the wave
288 Wisconsin Academy of Sciences, Arts, and Letters.
lengths measured by caesium with the yellow screen, in his
samples taken several years before. He found the suspended
matter to be non-selective. Dividing 74.5% by .887 we get
84.0% for the suspended matter which suggests 57.2% for green
and yellow light and 43.3% for blue light due to staining alone.
As a field method it might be possible to measure the trans¬
mission of the efflunt when added to pure water, to make pos¬
sible a rough determination of the effect of stains in the water.
This method will have to be developed, however.
A comparison of Lake Mendota with Puget Sound water is
shown by reference to the top curve (A) of figure 9 which is
for sea water and the bottom curve (G) for Lake Mendota.
The potassium cell was used in both cases. Prom two to three
meters of Lake Mendota water absorb as much as ten meters
of sea water.
It must be borne in mind that the limited number of readings
were intended as a test of the methods and equipment rather
than a study of Lake Mendota water. Both equipment and
method appear to be adequate for detailed study provided some
method of determining the transmission of suspended matter
can be developed. It is obvious that selective absorption of
light can be determined but suspended and dissolved matter can¬
not be evaluated with the equipment at hand. While Knudsen
found the Danish salt waters stained, he did not differentiate
the suspended and dissolved matter.
The most serious defect in the equipment is the small current
delivered by the caesium cells under yellow and red glasses.
The caesium cells used gave much smaller current in full sun¬
light than the rubidium and potassium cells, whereas it would
be desirable to have caesium cells giving 300 microampheres in
full sunlight as the yellow glass reduces it to less than 5%.
The maximum light intensity was found to be only slightly
greater in the yellow than in the blue in an unstained water.
( Shelf ord and Gail fig. 12, p. 168). In the Lake Mendota water
the light at five meters would be largely restricted to yellow.
Acknowledgments and Bibliography
The expense of the work was borne by the Graduate School,
the Department of Zoology, and the Department of Physics of
the University of Illinois. The greater part, including the
Shelf ord & Kunz — Photo-Electric Cells in Light Penetration. 289
transportation to Madison, was cared for by the Graduate
School. The authors are indebted to Dean A. H. Daniels, Prof.
H. B. Ward and Prof. A. P. Carman for their unqualified sup¬
port of the project.
They are further especially indebted to President Birge and
Mr. Chancey Juday of the University of Wisconsin and the
Wisconsin Geological and Natural History Survey for extensive
assistance in connection with the carrying out of the measure¬
ments at Madison.
They are also indebted to the Corning Glass Works, especially
to Dr. 0. A. Gage and to Dr. H. P. Gage for assistance with
the glasses used as screens; to Prof. W. F. Schulz and to Mr.
A. E. Wuesteman for courtesies in connection with the grinding
and polishing of the colored glasses used; to Prof. E. B. Paine
of the Department of Electrical Engineering and to Mr. T. 0.
Henderson, Assistant Superintendent of Buildings, for the loan
of equipment ; and to Mr. C. E. Hollister of the Department of
Chemistry for assistance with the battery box and cable insu¬
lator.
Literature Cited
Birge, E. A., 1922. A second report on limnological apparatus.
Trans. Wis. Acad. Sci. 20:533-552
Gibson, K. S., Tyndall, E. T. and McNicholas, H. J. 1920. The ultra¬
violet and visible transmission of various colored glasses. Tech.
Papers of the Bur. Stand. 148:1-27
Klugh, A. Brooker, 1925. Ecological photometry and a new instru¬
ment for measuring light. Ecology 6:203-237
Knudsen, Martin, 1922. On measurement of the penetration of light
into water. Pub. de Circ. (Conseil 1 expl. de la mer.) 76:1-15
Pietenpol, W. B., 1918. Selective absorption in the visible spectrum
of Wisconsin lake waters. Trans. Wis. Acad. Sci. 19:562-593
Shelford, V. E. and Gail, F. W., 1922. A study of light penetration
into seat water made with the Kunz photoelectric cell with par¬
ticular reference to the distribution of plants. Pub. Puget Sd.
Biol. Sta. 3:141-174
19
290 Wisconsin Academy of\ Sciences, Arts, and Letters.
Table 1. Showing the ‘percent of light transmitted by the surface
waters of Lake Mendota on the afternoon of June 13, 1925. The
first meter is designated as 1; the second, 2, etc. The figures
given are calculated to vertical penetration. Duplicate figures
are calculated from two meter intervals.
Table 2. Showing percent of light transmitted h'y the two surface
meters of Lake Mendota water near shore on the forenoon of June
13, 1925, with cells with ray filters. The figures .972 etc. are the
values of the mean transmission of pure water for the rays measured
by th& cell and schreen. N for Lake Mendota divided by that of
pure water .972 is, for example, UO.O over .927 or J^l.l of the third
column which represents the effect of foreign matter.
Shelf ord & Kunz — Photo-Electric Cells in Light Penetration. 291
Table 3. Showing the percent of light transmitted per meter after
being divided by the transmission coefficient of the cell and glass for
pure water.
292 Wisconsin Academy of Sciences, Arts, and Letters.
Fig. 1. Battery and switch box used with the photo-electric cells.
The B batteries (B) were connected by inserting solid plugs into the
telephone jacks (J). The B batteries have a series of cells which may
be added in one and one-half volt steps, thus insuring the desired vol¬
tage with less than a volt and one-half. The voltage could be read by
inserting the volt plug V into connection with the last cell and insert¬
ing another solid plug, not shown, to connect the negative pole with
the volt meter (see fig. 2). With the voltage resistance in series, the
readings were reduced to exactly one-fifth, and stabilized. The mi-
cro-ammetr is shown at the right. The cover (C) could not be raised
until all plugs were withdrawn, after which the space between the mi¬
croammeter and the bakelite panel (P) could be used for storage.
Large sized B batteries aggregating upwards of 200 volts could be
carried. The equipment was designed by V. E. Shelford and C. E.
Hollister. See also the wiring diagram shown in fig. 2.
Shelf ord & Kunz — Photo-Electric Cells in Light Penetration. 293
Fig. 2. Wiring diagram of battery and switch box. The cell cir¬
cuit was completed by inserting one cell circuit plug into opening
No. 1 and the other fnto opening No. 2.
Fig. 3. Insulator
to keep water from
entering the cover¬
ing of the outer
cable. C, cable; P,
plug screwed in af¬
ter the assembly was
completed; S, sup¬
ports for wires car¬
rying the basket
containing the cell;
R, short rod for
holding the knot in
the center of the
space; I, iron pipe
supports for the rod
(R) cc, separated
wires covered with
gum tubing.
294 Wisconsin Academy of] Sciences, Arts, and Letters.
Fig. 4. A photo-electric cell designed for sub-aquatic work. M,
space for mercury.
Fig. 5. A cell in position in the protecting basket, with the brass
screen holder in position. CC, divided cable strands leading to cc of
fig. 3. When necessary a weight could be suspended below the basket
from S of fig. 3, by wire or cord passing through the loops on the
basket. This was not required in Lake Mendota.
Shelf ord & Kunz — Photo-Electric Cells in Light Penetration. 295
400 20 40 so eo 500 20 40 so eo 600 to 40 so
Fig. 6. The sensitivity of the potassium cell with and without
blue glass. The sensitivity of the cell with blue glass was used to
determine its average transmission for pure water, by taking the
transmission of pure water at 420, 3^/4 times; at 446, 5% times; at
480, 1.8 times (actually at more frequent intervals) and securing an
average.
296 Wisconsin Academy of. Sciences, Arts, and Letters.
Fig. 7. Sensitivity of the rubidium cell with and without noviol C.
Shelf ord & Kunz — Photo-Electric Cells in Light Penetration. 297
Fig. 8. Sensitivity of the caesium cell with and without, noviol A,
yellow glass and red glass.
298 Wisconsin Academy of) Sciences, Arts, and Letters.
Fig. 9. Transmission coefficients for each meter from 1 to 10. A
is for sea water (Puget Sound) with an unscreened potassium cell;
compare with G which is Lake Mendota with an unscreened potas¬
sium cell. B is for the caesium cell with the yellow screen. E is for
the caesium cell unscreened. D is for the rubidium cell with noviol
C. F is for the rubidium cell unscreened. C is for the potassium
cell alone; G is for the potassium cell with a blue screen. The curves
suggest plankton maximum in the second meter and another in the
ninth meter. The coefficients for the various cells suggest that the
second meter is selective.
A THIRD REPORT ON LIMNOLOOICAL APPARATUS
Chancey Juday
Notes from the Biological Laboratory of the Wisconsin Geological
and Natural History Survey. XXV.
The present paper is a continuation of the two reports that
have already been published regarding limnological apparatus,
one hy Juday in 1916 and the other by Birge in 1922.
The Foerst Electric Centrifuge
Quantitative and chemical studies of the net plankton of lakes
in the vicinity of Madison, Wisconsin, were begun in 1911 and
were continued until 1917. In the early stages of this investiga¬
tion it was found that a very large portion of the plankton
material was lost through the meshes of the net. Various
methods of obtaining the lost organisms were tried and a centri¬
fuge proved to be the most efficient instrument for their re¬
covery. A large size De Laval centrifuge was installed in 1915
and was used until 1917 for the investigations relating to the
organisms that are lost bj the net. A report on this work was
published in 1922.
The importance of the results obtained with this large centri¬
fuge made it desirable to extend these studies to lakes in other
parts of Wisconsin. The large size and the equipment required
to operate it made this De Laval centrifuge impractical for field
work in other lake districts. Mr. J. P. Foerst, mechanician of
the Physics Department, University of Wisconsin, then under¬
took the task of constructing a portable centrifuge which could
be readily tak^n into the field and operated wherever electric
current is available. The result of his work is the Foerst
Electric Centrifuge which is shown in figure 1. This instrument
has been used for a period of five years and very interesting and
valuable results have been obtained with it during this time.
The motor (M, figure 1) consists of a standard grinding motor
manufactured by the Wisconsin Electric Company of Racine,
Wisconsin. It has a speed of 20,000 revolutions per minute and
300 Wisconsin Academy ofi Sciences, Arts, and Letters.
operates on either direct or alternating current of 110 volts. In¬
stead of the usual wheels that are attached to the spindle for
grinding purposes, a small centrifuge bowl is used.
The motor is mounted on a cast iron base (B) which holds it
in a vertical position. Inside the base is a rheostat which gives a
range in speed from a few hundred up to 20,000 revolutions per
minute. The amount of resistance introduced into the circuit,
which determines the speed of the motor, is regulated by turning
the wheel marked R in figure 1.
A centrifuge with a still higher speed has been used for the
past four years. The motor has a speed of 4,000 revolutions per
minute and the spindle is driven by a belt. Seven interchange¬
able pulleys give a range in speed from 3,600 to 50,000 revolu¬
tions per minute ; except for a few experiments at the maximum
speed this centrifuge has been operated at 32,000. It is a larger
and heavier machine than the one shown in figure 1, but it can
be readily handled in the field ; in fact it has been used for this
purpose almost as much as the small machine.
The same bowls are used on both machines. Several hundred
duplicate determinations have shown that the small machine re¬
moves substantially the same amount of plankton at 20,000 as
the large one does at 32,000 revolutions per minute. Likewise
several experiments have shown that a speed of 32,000 yields as
much material as 50,000. The weight of the small centrifuge
is 8 kilograms and of the large one 22 kilograms.
Eight different kinds of bowls have been used in the experi¬
mental tests to ascertain which one is most efficient for general
use. The two that have given most satisfactory results are
shown in cross section in figures 2 and 3. Figure 2 represents
the original type. In this bowl the side is vertical; that is, it
forms a right angle with the bottom. The corner is rounded off
somewhat so that the material can be more readily removed from
the angle. At the top of this bowl there is a rim 1.5 millimeters
long which projects inward 1 millimeter. The outside diameter
of the bowl is 57 millimeters and the height is 29 millimeters.
The inside diameter is 50 millimeters at the rim and 52 milli¬
meters below the rim ; the depth is 25 millimeters.
The bowl shown in figure 3 has the same outside diameter at
the base and the same height as that in figure 2, but the sides
slope inward at an angle of 5 degrees ; that is, the angle between
the side and the bottom of the bowl is 85 instead of 90 degrees.
Jiiday — Limnological Apparatus
301
There is no rim at the top. The inside diameter at the top is
50 millimeters and the depth is 25 millimeters. Experiments
have shown that this bowl is just as efficient in removing
plankton organisms as the one with the rim ; since the catch can
be washed out of the bowl with sloping sides a little more con¬
veniently than out of the one with a rim, the former type has
been adopted for general use. These bowls are made of brass or
steel ; the latter has a tendency to rust, which may be prevented
by plating it with nickel.
A hub projects upward from the bottom of the bowl for a dis¬
tance of about 15 millimeters. The upper part of a hole in the
middle of this hub is provided with threads by means of which
the bowl is screwed onto the spindle of the motor. The point
of attachment to the spindle is raised in this manner in order to
lower the center of gravity of the bowl and thus prevent undue
vibration. In the sloping bowl, the upper part is made thinner
in order to keep the center of gravity as low as possible.
A bronze guard (G in figure 1) is fastened to the framework
of the motor at the upper end of the spindle ; this guard
surrounds the greater part of the bowl and is provided with an
outlet tube (0) through which the water escapes from the
centrifuge. The side wall of this guard is 3 millimeters thick;
the inside diameter of the guard is 70 millimeters and the depth
is 22 millimeters inside and 27 millimeters outside.
A cover (C in figure 1) made of sheet brass one-half milli¬
meter thick fits snugly inside the guard and completes the hous¬
ing for the bowl. The cover is 40 millimeters high and possesses
a tube (T) in the the center 17 millimeters in diameter, through
which the water is fed into the bowl. The feeding tube extends
down to within one millimeter of the bottom of the bowl and a
diaphragm at the lower end prevents the water from being
thrown over the top of the bowl before it is centrifuged. This
diaphragm is 49 millimeters in diameter. When the water
strikes the bottom of the revolving bowl it is quickly set into
rotation and thrown against the side where its speed of rotation
is increased as it passes up toward the top of the bowl; it is
finally discharged over the top in a fine spray. Most of the or¬
ganisms are deposited on the side of the bowl within five milli¬
meters of the bottom, thus showing that the water promptly ac¬
quires sufficient speed of rotation to cause their deposition. At
a speed of 20,000 revolutions per minute, the inner surface of
302 Wisconsin Academy of Sciences, Arts, and Letters.
the side of the bowl travels at a rate of 3,200 meters per minute ;
at a speed of 32,000 the rate is 5,120 meters per minute.
The only water that remains in the bowl at the end of a run is
that below the rim in the first type of bowl or that in the angle
of the second type; in the bowls described above it amounts to
about 4 cubic centimeters.
Bowls with inside depths of 17, 20, 25, and 29 millimeters
have been used in the experiments to determine the effectiveness
of the different types. It was found that efficiency increases
somewhat with increasing depth up to 25 millimeters, but there
was no difference between the 25 and 29 millimeter bowls. Two
V-shaped grooves 1.5 millimeters wide and 0.5 millimeter deep
were cut into the side on the upper half of one of the 29 millime¬
ter bowls, but these grooves did not give this bowl any advantage
over the others. In another experimental bowl the top rim
projected inward a distance of 3.5 millimeters instead of the
usual 1 millimeter, but this made no difference in the size of the
centrifuge catch. Two small holes were drilled in the rim so
that the quantity of water left in the bowl could be reduced
to the usual amount at the end of the run.
Two modifications were made for the purpose of giving the
water the same speed of rotation as the bowl. One bowl was
provided with four thin brass wings which project inward from
the side for a distance of 5 millimeters. The water passes up
the side of the bowl between these wings so that it attains sub¬
stantially the same speed as the bowl before it reaches the top.
The amount of material obtained from this bowl, however, did
not prove to be any larger than that of a bowl with smooth
sides and the same height.
Another bowl was fitted with a special feeding cone which
has a rim 6.5 millimeters wide at its lower end ; this device is at¬
tached to the bottom of the bowl by means of three screws pass¬
ing through the rim. The water is fed into the top of the cone,
passes down to the bottom of the bowl and then out to the side
through a series of small grooves on the lower side of the rim.
This rim extends out to within 2 millimeters of the side of the
bowl, so that the water acquires the same speed of rotation as
the bowl before it leaves these grooves. This special bowl did
not yield any larger catches of plankton than the duplicates
taken with the regular bowls of the same height.
Juday — Limnological Apparatus
303
With one exception the first centrifuging removes approxi¬
mately 98 per cent of the organisms that are usually considered
in a plankton catch. Aphanizomenon is the only form noted
thus far which is troublesome in this respect ; only about half of
it is removed in the first centrifuging, but substantially all that
remains is taken out by a second centrifuging. With very few
exceptions this form is present in such small numbers that a
second centrifuging is not necessary.
Plate counts have shown that 40 to 70 per cent of the bacteria
are removed from the water by the first centrifuging. This high
efficiency is maintained for plankton catches that will weigh as
much as 8 milligrams when dry. The water is centrifuged at a
rate of one liter in 5 to 8 minutes, 6 to 7 minutes being the
usual time. The water is fed into the centrifuge from a one
liter aspirator bottle which is placed on a box beside the cen¬
trifuge ; the rate of flow is regulated by a glass stop-cock in the
outlet tube of the aspirator bottle.
With this type of centrifuge, plankton material may be ob¬
tained for numerical, gravimetric, and micro-chemical studies.
For purposes of enumeration one-half liter samples are used;
if some of the organisms are abundant, they are enumerated
without concentration and the less abundant forms are then
enumerated in the centrifuge catch. Duplicate samples of one
liter each are usually used for the gravimetric determinations ;
when the plankton is scarce a two or three liter sample may be
used. The catch is rubbed offi the side of the bowl with a glass
rod which is tipped with a piece of gum rubber tubing and trans¬
ferred to a platinum dish with a capacity of 8 cubic centimeters ;
the bowl is then washed twice with distilled water, about one
cubic centimeter being used each time, and this is added to the
material in the dish. The catch is placed in an electric oven
where the water is evaporated and the material is dried for a
period of 24 hours at a temperature of 60° C. After weighing,
the catch is ignited in an electric furnace at a temperature of
about 600° C. for a period of 30 minutes. A second weighing
shows the gross loss on ignition. In order to correct for the loss
on ignition sustained by the 4 cubic centimeters of lake water
included with the catch, a blank consisting of 10 cubic centi¬
meters of centrifuged water is run along with the plankton sam¬
ple. The net loss after making this correction is regarded as the
304 Wisconsin Academy of Sciences, Arts and Letters.
organic matter of the plankton. The quantity of organic mat¬
ter in the plankton of the Wisconsin lakes that have been studied,
varies from a minimum of about 700 milligrams to a maximum
of 6,000 milligrams per cubic meter of water.
Both numerical and gravimetric results are obtained for the
net plankton as well as for the total plankton secured with the
centrifuge; the closing net described in the first report is used
for these catches. For the gravimetric determinations, the net
is hauled through the desired stratum and the catch is trans¬
ferred from the plankton bucket to a piece of bolting cloth
placed in a funnel. After the water has drained off, the net
plankton is carefully removed from the bolting cloth with a
sharp knife or scalpel and placed in a small platinum dish. The
catch is then dried, weighed and ashed as indicated for the cen¬
trifuge material.
In addition to the plankton organisms, the centrifuge also
removes a certain amount of silt from the lake waters; this is
shown by the fact that the ash of the centrifuge catch usually
amounts to 50 per cent or more of the dry weight. The per¬
centage of ash in all of the plankton forms, except the diatoms,
is much smaller; it is less than 10 per cent in many of these
forms.
A certain amount of organic matter is removed by the cen¬
trifuge even after all of the plankton organisms, except the bac¬
teria, are gone. With the exception of a small percentage of the
bacteria, the plantkon material is all removed from the water by
the end of the third centrifuging, yet this water continues to
yield small amounts of organic matter up to the eleventh cen¬
trifuging; no attempt has been made to carry the experiment
further than this. This result suggests that the lake water con¬
tains organic matter in a collodial state which is gradually
thrown out as the centrifugal process continues.
During the summer of 1925 the centrifuges and the electric
furnace were successfully operated in the field by a Kohler
Light-Power plant made by the Kohler Company of Kohler,
Wisconsin. The plant is especially designed for use on a farm ;
it furnishes a direct current of 110 volts and has a capacity of
1,500 watts. The plankton material was dried in a copper dry¬
ing oven which was heated to 60° C. by a kerosene incubator
lamp.
Juday — Limnological Apparatus
305
Keller Portable Balance
A good portable balance is required for gravimetric studies of
the plankton during the summer field work. A Keller portable
assay balance, made by the G. P. Keller Manufacturing Com¬
pany of Salt Lake City, Utah, has given excellent, service in such
investigations during the past four years. A sketch of it is
shown in figure 4. This balance has given good results and has
not shown any appreciable effects of ordinary vibrations when
used upon tables or temporary benches; it has a capacity of
100 grams and is sensitive to 1 /200' milligram.
The balance case is trapezoid in form, 30 centimeters long, 15
centimeters high and 14 centimeters wide at the base. The
carrying case is 37 centimeters long, 19 centimeters high and 17
centimeters wide. It requires only a very few minutes to dis¬
mantle the balance and place it in the carrying case, or to set
it up again upon removal from the case.
Bottom Dredge
The bottom dredge shown in figure 5 is a modification of the
one described by Ekman in 1911. It is especially designed for
the capture of organisms that live on the bottom, such as Mysis
and Pontoporeia, and not those that burrow into the mud.
The dredge consists of two parts, namely, a canvas part in
front and back of this a straining part made of silk gauze. The
dredge has a rectangular opening 16 by 38 centimeters; the
canvas part is 40 centimeters long and the gauze part is 55 centi¬
meters long. The dimensions at the point where the canvas and
gauze parts join are 19 by 42 centimeters. A canvas apron ex¬
tends back over about half of the gauze part in order to protect
it.
The canvas and gauze are attached to quadrangular frames
made of brass. The frame at the mouth of the dredge is made
of brass 2 centimeters wide and 3 millimeters thick; the one in
the middle of the dredge is 2.5 centimeters wide and 3 millime¬
ters thick. The front part of the middle frame is provided
with small holes, about a centimeter apart, through which the
canvas is sewed to the frame. In addition to the canvas, the
two brass frames are connected at each end by small ropes which
pass through holes made to receive them.
20
306 Wisconsin Academy of Sciences, Arts and Letters.
The silk gauze is attached to a separate quadrangular frame
made of a strip of brass 2.5 centimeters wide and 1 millimeter
thick as shown in figure 6. This frame fits snugly over the one in
the middle of the dredge to which the canvas is attached and
is fastened to the latter by six small screws, two on each side
and one at each end. Gauze of different meshes is attached to
three or four of these frames, so that one can readily change
from one size of mesh to another whenever it is desirable to do
so. Small holes in the outer part of these frames provide for
the attachment of the canvas band to which the gauze is
sewed. A band of light canvas is sewed around the opening
in the lower end of the gauze net ; a line tied around this canvas
closes the bottom of the dredge. During dredging operations
a weight is attached to the line about a meter or a meter and a
half in front of the mouth of the dredge.
Trap for Plant Dwelling Animals
Many animals dwell upon the larger aquatic plants, such as
Potamogeton and Myriophyllum, and the trap shown in figure 7
was designed for the capture of such forms. The sides of the
trap are made of light canvas or drilling and the bottom is
made of No. 72 extra heavy grit gauze. At the top the canvas
bag is sewed to a brass frame which possesses small holes
for this purpose. The frame is made of brass 2.5 centimeters
wide and 3 millimeters thick; there are two double hinges in
the center so that the mouth of the trap can be completely
closed. The opening of the trap is 36 by 37 centimeters and the
depth is 85 centimeters.
In making a catch the open trap is lowered over the plants
to be examined; the plants are then loosened from the bottom
and the mouth of the trap is closed. The trap is raised to the
surface with the mouth upward and hauled into the boat, the
water being allowed to drain out through the gauze before it is
taken on board. The plants are transferred to a pail contain¬
ing some water and taken to the laboratory where the animals
are washed off and enumerated. Two individuals are required
for the most convenient operation of the trap, one in the water
to make the catches and the other in the boat to haul the appa¬
ratus on board and remove the material. By using a diving
hood the trap has been operated at depths of 5 and 6 meters.
Juday — Limnological Apparatus
307
Apparatus for Determining the Coefficient of Nets
The plankton trap described in the first paper on limnological
apparatus (Juday 1916) may be used for the determination of
the coefficient of nets. Trap catches are made at meter or half
meter intervals in the stratum through which the net is hauled
and the average result yielded by these catches is a fair index of
the efficiency of the net. The trap has been operated success¬
fully at a depth of 60 meters so that the coefficient of a net can
be determined in the lower as well as in the upper strata of a
lake.
Another instrument was briefiy described by Birge in 1898.
It consists of a galvanized iron tube 3 meters long and 10 centi¬
meters in diameter which is provided with a closing device at its
lower end, A side view of the closing part is shown in figure 8 ;
it consists of a slide and a carrier that bears a small plankton
net. The carrier and net can be slipped to one side so that the
opening is entirely free when the tube is lowered into the water.
The slide is made of a brass plate 26 centimeters long, 15.3 centi¬
meters wide and 3 millimeters thick ; two pieces of brass 26 centi¬
meters long and 3 millimeters thick are attached to each side
of the brass plate, fiush with the edge, by means of four screws.
The upper piece of brass is 7 millimeters wide and the lower
one is 15 millimeters wide, so that the latter projects inward 8
millimeters beyond the former and forms a track or groove in
which the carrier can be moved back and forth.
At one end the brass plate possesses a hole 10 centimeters in
diameter around which is soldered a tube 10 centimeters long;
this tube is just a little larger than the iron tube so that the
latter slips inside and attaches to the brass part by means of two
bayonet joints. The other end of the brass plate has a tube 1
centimeter in diameter and 1.5 centimeters long through which
the air escapes from the net when the apparatus is submerged ; a
piece of gauze is tied over the small tube in order to keep plank¬
ton organisms out of the net.
The carrier is made of a brass plate 13.8 centimeters square
and 3 millimeters thick ; it has a hole 10 centimeters in diameter
in the middle around which is soldered a tube 5 centimeters long.
The net is attached to the lower end of this tube by means of a
clamp activated by a screw. The bolting cloth net is 15 centi¬
meters long and the lower end of it is attached to a brass ring
that fits one of the standard plankton buckets described in the
308 Wisconsin Academy of Sciences, Arts and Letters.
first report. The weight of the bucket is supported by three
pieces of line which are attached to the clamp at the top of the
net and to the clamp at the bottom.
A small pulley is soldered to each end of the slide; the lines
with which the carrier is moved from one side to the other pass
over these pulleys and are tied to a loop soldered to the carrier.
These pulleys project below the lower edge of the brass plate and
serve as stops to prevent the carrier from being pulled out too
far. In actual practice only the closing line is used because
the net is moved away from the tube opening before the ap¬
paratus is lowered into the water. The slide and carrier are
closely fitted together, so that plankton organisms can not escape
between the bottom of the tube and the top of the net.
The galvanized iron tube is provided with a heavy wire handle
at the upper end to which a rope is attached when the apparatus
is being used. The top of the tube also possesses a close fitting,
hinged cover. A wire soldered to the cover projects outward 10
centimeters at the point where the hinge is attached; a lead
weight at the outer end of this wire keeps the cover open when
the tube is being lowered into the water. The cover is closed
by means of a line which is attached to a wire loop on the edge
of the cover opposite the hinge and which passes through a loop
on the side of the iron tube.
In making a catch, the tube is lowered into the water with
the net carrier moved to one side of the opening; it is lowered
slowly so that no appreciable currents are set up in the water.
When the top of the tube is half a meter or a meter below the
surface, the cover is closed and the net carrier is drawn across
the bottom of the tube, thus imprisoning a column of water 3
meters long and 10 centimeters in diameter. The tube is then
slowly raised to the surface and lifted out of the water so that
the imprisoned water is filtered through the net. Several
catches with the tube as well as with the net, the coefficient of
which is to be determined, are taken at the same time through
the same stratum of water. A comparison of the number of
plankton organisms in the tube catches with that in an equal
number of net catches, gives an index of the efficiency of the net.
Literature Cited
Birge, E. A. 1898. Plankton studies on Lake Mendota. II.
Trans. Wis. Acad. Sci., Arts and Let. 11: 278-286.
Juday — Limnological Apparatus
309
- 1922. A second report on limnological ap¬
paratus. Ibid. 20: 533-552.
Birge, E. A. and Juday, C. 1922. Inland lakes of Wisconsin.
The plankton. Bui. 64, Wis. Geol. and Nat. Hist.
Survey, 222 pp.
Ekman, S. 1911. Neue Apparate zur qualitativen und quanti-
tativen Erforsehung der Bodenfauna der Seen. In¬
ternal. Bevue 3 : 553-561.
Juday, C. 1916. Limnological apparatus. Trans. Wis. Acad.
Sci., Arts and Let. 18: 566-592.
310
Wisconsin Academy of Sciences, Arts and Letters.
Fig. 1. Sketch of Foerst electric centrifuge. B, base, M, motor,
R, rheostat wheel, 0, outlet tube, G, guard, C, cover,
T, inlet tube.
Juday — Limnological Apparatus
311
Fig. 2. Cross section of bowl with a rim. Natural size.
Fig. 3. Cross section of bowl with sloping sides. Natural size.
Fig. 4. Keller portable assay balance.
312 Wisconsin Academy of Sciences, Arts and Letters.
Fig. 6. Gauze net for bottom dredge,
Fig. 7. Trap for capturing plant dwelling animals.
314 Wisconsin Academy of Sciences, Arts and Letters.
Fig. 8. Closing device attached to lower end of tube used
termining the coefficient of nets.
THE ROTIFER FAUNA OF WISCONSIN.— III.
A Revision of the Genera Lecane and Monostyla
H. K. Harking and F. J. Myers
Notes from the Biological Laboratory of the Wisconsin Geological
and Natural History Survey. XXVI.
INTRODUCTION.
Many eminent specialists have declared the ‘‘invertebrate^’
fauna of North America virtually unknown and it is perhaps
superfluous to add to this our growing conviction that this is
nowhere more readily demonstrable than in the relatively small
phylum which we have been studying for some time. When J en-
nings compiled the records' of rotifers found in the United States
at the close of the Great Lakes Biological Investigations in 1899,
the total number of species was 246, quite a number of these now
unidentifiable. In the two papers already published on the
rotifers of Wisconsin no less than 136 species are described from
personal observations of a part of the family Notommatidae. Of
these only 27 species are to be found in Jennings’s list, or less
than one fifth. As a result of our collections in Wisconsin and
a study of the rotifers of Mount Desert Island, undertaken at
the suggestion and with the hearty cooperation of Dr. Dahlgren,
director of the Mount Desert Island Biological Laboratory, over
one hundred undescribed species belonging to the family already
dealt with are now on hand. Under the circumstances we may
perhaps be granted forgiveness for departing somewhat from
our original plans, or rather from their last revision. We had in¬
tended to conclude, temporarily at least, the work on the Notom¬
matidae with a revision of the forcipate group or Dicranophori-
nae, as defined in the second paper. At the time of closing the
manuscript we had some 50 identified species belonging to this
group. The Mount Desert collections added at least 25 more,
or a sufficient number to convince us that we were very far from
being able to “see the bottom”, in fact, we could not even dis¬
cover a halting place, where we might at least temporarily feel
316 Wisconsin Academy of Sciences, Arts and Letters.
that we had in hand a fair proportion of the species that were
likely to be found in the immediate future. It is very evident
that more field work is necessary before a satisfactory revision
of the Dicranophorinae can he undertaken with any hope of its
being serviceable for even a few years; we are therefore com¬
pelled to postpone this for the moment, hoping that it may
shortly appear to be a more promising undertaking.
A large amount of material for a revision of the genera Le-
cane and Monostyla has been accumulating for some years as a
sort of by-product and this seems now to offer some possibility
for reasonable permanency when worked up. Undescribed spe¬
cies belonging to these two genera are slowly and steadily de¬
creasing in numbers; this is especially true of Monostyla, where
a new species is now rarely found.
Lecane and Monostyla are very closely related and individual
species do not as a rule exhibit striking differences. We have
therefore limited the descriptions strictly to what is necessary
for the determination of the species ; no attempt has been made to
deal with the anatomy, which seems to be quite uniform through¬
out the two genera. The outline of the lorica, its surface mark¬
ings and the form and length of the toes are very constant ; for
this reason special emphasis has been placed upon these features
and no effort has been spared to represent them as accurately as
possible in the figures. The descriptions must be considered
largly supplementary ; it seems impossible to state in words with
sufficient clearness the slight, but important, differences of form
and ornamentation of these animals.
A few words on the classification of the rotifers may not be out
of place. The one employed by Hudson and Gosse, a modifica¬
tion of Dujardin’s, is now generally considered unsatisfactory.
Some radical changes were suggested by Wesenberg-Lund in
1899, and De Beauchamp provided ten years later the rational
basis needed for a revision, but did not carry it out in detail. In
his latest paper Wesenberg-Lund offers some modifications to
his earlier outline, bringing it into fair agreement with De
Beauchamp’s ideas, with the notable exception of the relation¬
ship of the families with malleo-ramate mastax. With this ma¬
terial available we have by degrees arrived at the tentative clas¬
sification given below; as will be noted, it is mainly De Beau¬
champ’s, with some modifications suggested by personal obser¬
vations, and includes only the Ploima, around which the prin-
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 317
cipal differences of opinion are centered. The number of fami¬
lies has been reduced somewhat, and this will probably not be
considered objectionable. There is no obvious advantage in hav¬
ing a separate family for each genus; any system of classifica¬
tion put forward must be a means of expressing the author’s
ideas on the mutual relationship of the component parts; plac¬
ing each in a separate box is a confession of failure.
We have divided the Ploima into six major groups, which may
be termed suborders, tribes, superfamilies or anything else that
individual fancy may suggest. This regrouping seems, on the
evidence available, to give a better perspective of the order. No
detailed evidence for the placement of the individual genera will
be offered at this time; on the whole we do not anticipate any
violent dissent, but welcome constructive criticism. The rea¬
sons that appear to favor this arrangement will be given in fu¬
ture papers on the rotifer fauna of Wisconsin; in the absence
of a thorough revision it is hardly possible to do more. How¬
ever, in spite of, or perhaps on account of, this serious shortcom¬
ing, a useful purpose may be served in calling attention once
more to some of the as yet unsettled problems of rotatorian
taxonomy.
Order PLOIMA.
NOTOMMATOIDEA.
Family NOTOMMATIDAE.
Subfamily Proalinae.
Genus Proales.
Proalinopsis.
? Squatinella.
Subfamily Notommatinae.
Genus Notommata.
Taphrocampa.
Drilophaga.
Pleurotrocha. ^
Cephalodella.
Dorystoma.
Rousseletia.
Tylotrocha.
Resticula.
Eosphora.
Eothinia.
Sphyrias.
Scaridium.
Monommata.
Subfamily Tetrasiphoninae.
318 Wisconsin Academy of Sciences, Arts and Letters.
Genus Tetrasiphon.
Subfamily Lindiinae.
Genus Lindia.
Subfamily Birgeinae.
Genus Birgea.
Subfamily Dicranophorinae.
Genus Dicranophorus.
Encentrum.
Erignatha.
Albertia.
Family SYNCHAETIDAE.
Genus Synchaeta.
Parasynchaeta.
Ploesoma.
Polyarthra.
Anarthra.
? Microcodon.
Family GASTROPODIDAE.
Genus Gastropus.
Ascomorpha.
Chromogaster.
Family TRICHOCERCIDAE.
Genus Trichocerca.
Diurella.
Elosa.
ASPLANCHNOIDEA.
Familiy ASPLANCHNIDAE.
Genus Asplanchna.
A splanchnopus.
Harringia.
BRACHIONOIDEA.
Family BRACHIONIDAE.
Genus Brachionus.
Schizocerca.
Platyias.
Keratella.
Notholca.
Anuraeopsis.
Family EPIPHANIDAE. '
Genus Epiphanes.
Rhinoglena.
Cyrtonia.
Proalides.
Mikrocodides.
Family EUCHLANIDAE.
Genus Euchlanis.
Dipleuchlanis.
Lecane.
Earring <& Myers — Rotifer Fauna of Wisconsin. — III. 319
Monostyla.
Lepadella.
Colurella.
Mytilina.
Trichotria.
Lophocharis.
Wolga.
Macrochaetus.
FILINIOIDEA.
Family FILINIIDAE.
Genus Filinia.
Tetramastix.
Pedalia.
TESTUDINELLOIDEA.
Family TESTUDINELLIDAE.
Genus Testudinella,
Pompholyx.
TROCHOSPHAEROIDEA.
Family TROCHOSPHAERIDAE.
Genus Trochosphaera.
Genus LECANE Nitzsch.
Euchlanid rotifers with illoricate, retractile head and loricate
body, strongly compressed dorso-ventraJly and oval or ovate in
outline; dorsal and ventral plates connected by a flexible mem¬
brane, forming lateral and posterior sulci; foot with two ex¬
tremely short, rudimentary joints, of which only the posterior
is movable; toes two; corona of family type; mastax modifled
malleate, with a piston attached to its ventral wall ; eyespot sin¬
gle and at the posterior end of ganglion ; retrocerebral sac
usually present, but no subcerebral glands.
Type of the genus. — Lecane lu\na (Muller) =(7ercarm luna
Muller.
This genus includes Cathypma Gosse and Distyla Eckstein;
as pointed out by Murray there are no real differences between
the species referred to them that would warrant their maintain-
anee. In his Infusionsthierchen Ehrenberg pointed out that if
Euchlanis luna were to be separated from Euchlanis, it would
have to take the generic name Lecane Nitzsch, an injunction
ignored by Gosse.
Quite a number of species have been described which we have
320 Wisconsin Academy of Sciences, Arts and Letters.
not seen. Some of these are of doubtful validity, and others
appear to be good species; a list is given below.
Cathypna affinis Levander.
Cathypna affinis Levander, Acta Soc. Fauna et Flora Fennica, vol.
12, No. 3, 1895, p. 50, pi. 3, fig. 31. — Murray, Journ. Royal Micr.
Soc., 1913, p. 346.
Distyla affinis Iroso, Mon. Zool. Italiano, vol. 21, 1910, p. 302; Atti
R. 1st. Incorr. Napoli, vol. 64 (for 1912), 1913, p. 466, fig. 13. —
Lucks, Rotatorienfauna Westpreussens, 1912, p. 110.— Sachse,
Susswasserfauna Deutschlands, pt. 14, 1912, p. 176.
Cathypna amban Stewart, Rec. Indian Mus., vol. 2, 1908, p. 320,
text fig.
Cathypna diomis Gosse, Journ. Royal Micr. Soc., 1887, p. 362, pi. 8,
fig. 2; Hudson and Gosse, Rotifera, Suppl., 1889, p. 41, pi. 31, fig.
38.
Cathypna gossei Lord, Science Gossip, vol. 26, 1890, p. 202, text fig.
Cathypna gracilis Sachse, Arch. Hydrobiol., vol. 10, 1914, p. 70, fig.
10.
Cathypna hudsoni Lord, Science Gossip, vol. 26, 1890, p. 202, text fig.
Cathypna latifrons Gosse, Journ. Royal Micr. Soc., 1887, p. 362, pi. 8,
fig. 3. — Hudson and Gosse, Rotifera, Suppl., 1889, p. 42, pi. 31, fig.
37.
Cathypna magna Stenroos, Acta Soc. Fauna et Flora Fennica, vol.
17, No. 1, 1898, p. 161, pi. 2, fig. 21.
Cathypna magna tenuior Stenroos, Acta Soc. Fauna et Flora Fenni¬
ca, vol. 17, No. 1, 1898, p. 161, pi. 2, fig. 22.
Cathypna rotundata. Olofsson, Zool. Bidr. Uppsala, vol. 6, 1918, p.
593, fig. 53. — Idelson, Trudy Plovuch. Morsk. Nauchn. Inst., pt. 12,
1925, p. 90.
Cathypna rusticola Gosse, in Hudson and Gosse, Rotifera, 1886, vol.
2, p. 95, pi. 24, fig. 6. — Sachse, Susswasserfauna Deutschlands,
pt. 14, 1912, p. 173, fig. 341.
Cathypna spenceri Shephard, Victorian Nat., vol. 9, 1892, p. 15.
Cathypna sulcata Gosse, in Hudson and Gosse, Rotifera, 1886, vol. 2,
p. 96, pi. 24, fig. 5. — Sachse, Susswasserfauna Deutschlands, pt. 14,
1912, p. 173, fig. 340. — ? Murray, Journ. Royal Micr. Soc., 1913, p.
557, pi. 22, fig. 5.
Cathypna web eri Mola, Zool. Anz., vol. 42, 1913, p. 115, figs. 5, 6;
Ann. Biol. Lac., vol. 6, 1913, p. 261.
Distyla acinaces Mola, Zool. Anz., vol. 42, 1913, p. 119, figs. 11, 12;
Ann. Biol. Lac., vol. 6, 1913, p. 260.
Distyla aculeata Jakubski, Zool. Anz., vol. 39, 1912, p. 542, figs. 3, 4.
Distyla branchicola Piovanelli, Mon. Zool. Italiano, vol. 14, 1903, p.
348.
Distyla gissensis Eckstein, Zeitschr. Wiss. Zool., vol. 39, 1883, pi. 383,
pi. 27, fig. 51. — Hudson and Gosse, Rotifera, 1886, vol. 2, p. 96, pi.
24, fig. 8. — Weber, Rev. Suisse Zool., vol. 5, 1898, p. 597, pi. 22, figs.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 321
6, 7. — Sachse, Susswasserfanna Deutschlands, pt. 14, 1912, p. 176,
figs. 343, 344.— Mola, Ann. Biol. Lac., vol. 6, 1913, p. 261. — Weber
and Montet, Cat. Invert. Suisse, pt. 11, 1918, p. 186.
Distyla korschelti Mola, Zool. Anz., vol. 42, 1913, p. 117, figs. 7, 8.
Distyla minnesotensis Herrick, Bull. Denison Univ., vol. 1, 1885, p. 5,
pi. 2, fig. 17.
Distyla striata Gosse, Journ. Royal Micr. Soc., 1887, p. 5, pi. 2, fig.
17.— Hudson and Gosse, Rotifera, SuppL, 1889, p. 43, pi. 31, fig 40.
Distyla terracianoi Mola, Zool. Anz., vol. 42, 1913, p. 118, figs, 9, 10;
Ann. Biol. Lac., vol. 6, 1913, p. 260.
Lecane carinata (Jakubski).
Distyla carinata Jakubski, Zool. Anz., vol 39, 1912, p. 542, figs. 1, 2.
Lecane carinata Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 60.
Lecane museicola (Bryce) .
Distyla musicola Bryce, Science Gossip vol. 27, 1891, p. 206, text
fig.
Distyla museicola Bryce, Science Gossip, vol. 27, 1891, p. 236.
? Cathypna museicola Murray, Journ. Royal Micr. Soc., 1913, p.
655, pi. 23, fig. 19.
Lecane musicola Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61. — •
Hauer, Mitt. Geogr. Ges. u. Naturhist. Mus. Lubeck, ser. 2, No.
30, 1925, p. 168.
LECANE CURVICORNIS (Murray).
Plate VIII, figures 1, 2.
Cathypna curvicornis Murray, Journ. Royal Micr. Soc., 1913, p.
346, pi. 14, fig. 22.
Cathypna nitida Murray, Journ. Royal Micr. Soc., 1913, p. 347, pi.
14, fig. 24.
Cathypna lofuana Murray, Journ. Royal Micr. Soc., 1913, p. 551,
pi. 22, fig. 1.
Lecane curvicornis Harring, Proc. U. S. Nat. Mus., vol. 17, 1914,
p. 535, pi. 17, fig. 3.
The lorica is broadly pyriform in outline; the anterior mar¬
gins are somewhat variable according to the stage of contraction,
but normally coincident, with a broad, V-shaped sinus; at the
external angles are two fairly large spines. The dorsal plate
is oval, truncate posteriorly and considerably narrower than the
ventral plate. Specimens from certain localities show a very
faint dorsal facetting, but the boundaries of the individual
facets are so indistinct that we have not attempted to figure them.
The lateral sulci are deep, but do not quite reach the anterior
margin. The ventral plate is more distinctly pyriform than the
dorsal and much wider; the transverse fold is very strongly
21
322 Wisco7isin Academy of Sciences, Arts and Letters.
marked. The posterior segment is small and rounded posteri¬
orly ; the coxal plates are moderately large and obtusely pointed.
The first foot joint is large and parallel-sided but somewhat in¬
distinct ; the second fairly large and subsquare. The toes are very
long, slender and parallel-sided, ending in a small claw with
basal spicule ; their length is about one-third of the total length.
Total length 280/a; length of dorsal plate 120/a, of ventral
plate 132/a; width of dorsal plate 95/a, of ventral plate 113/a;
anterior points 63/a ; toes wihout claw 69/a ; claw 10/a.
Lecane curvicornis is very abundant in certain localities ; Mur¬
ray found it in Brazil, it was collected by the Panama Biolog¬
ical Survey in the Panama Canal Zone, by Myers at Los Angeles,
California, by Dr. E. A. Birge and Mr. C. Juday in Texas and
Arkansas, by Dr. Birge during the Great Lakes Investigations
in 1899 at various points around Lake Erie and by Mr. Juday
in Guatemala. Murray ^s C. lofuana was collected in a tributary
to Lake Tanganyika in Africa. From this it will be seen to be
widely, but erratically distributed; where it is found, it is usu¬
ally abundant.
As noted by Harring, the three species described by Murray
are one and the same species; through the kindness of the late
Mr. Rousselet we had an opportunity to examine the original
specimens from which Murray described them and after com¬
parison with material from other sources we have no doubt
of their identity.
I/ECAXE ACROXYCHA Harring and 3Iyers, new species.
Plate VIII, figures 3, 4.
The outline of the lorica is a moderately elongate oval; the
anterior dorsal margin is almost straight, projecting slightly in
front of the ventral margin, which is somewhat concave; at the
external angles are two large, triangular cusps. The dorsal
plate is oval and without markings or facetting; it is consider¬
ably narrower than the ventral plate and its edges do not reach
the anterior margin. The ventral plate is of the same general
outline as the dorsal ; it is marked by a fairly distinct transverse
ridge in front of the foot joint. The lateral '^ulci are fairly deep.
The posterior segment is rather small and somewhat indistinct ;
the coxal plates are small and obtusely pointed. The first foot
joint is indistinct, the second moderately large and trapezoidal in
liar ring Myers — Rotifer Fauna of Wisconsin. — III. 323
form. The toes are long and fairly robust, very slightly en¬
larged above the relatively short claw; this has a small basal
spicule. The length of the toes is more than one third of the
total.
Total length 290/* ; length of dorsal plate 162/*, of ventral plate
182/* ; width of dorsal plate 136/*, of ventral plate 146/* ; anterior
points 80/* ; length of toes without claw 90/* ; claw 12/*.
Lecane acronycha appears to be confined to regions with soft,
acid water; we have found it abundant in Vilas and Oneida
counties, Wisconsin, around Atlantic City, New Jersey and on
Mt. Desert Island, Maine. It is related to L. ungulata and is its
equal in length, but not in bulk; it is readily distinguished by
its regular oval form, narrow anterior margin, small, rounded
posterior segment and the short claws.
LECANE UNGULATA (Gosse).
Plate IX, figures 3, 4.
Cathypna ungulata Gosse, Journ. Royal Micr. Soc., 1887, p. 361,
pi. 8, fig. 1. — Hudson and Gosse, Rotifera, Suppl., 1889, p. 42,
pi. 31, fig. 36.— WiERZEJSKi, Rozpr. Akad. Umiej., Wydz. Mat.-
Przyr., Krakow, ser. 2, vol. 6, 1893, p. 242. — Bilfinger, Jahresh.
Naturk. Wiirttemberg, vol. 50, 1894, p. 58. — ^Jennings, Bull. U. S.
Fish Comm., vol. 19 (for 1899), 1900, p. 91, pi. 19, figs. 26, 27. —
Daday, Zoologica, pt. 44, 1905, p. Ill; pt. 59, 1910, p. 84. — Rous-
SELET, Journ, Royal Micr. Soc., 1906, p. 406, pi. 15, fig. 1. — Meiss¬
ner, Izv. Turkestansk. Otd. Imp. Russk. Geogr. Obshch., vol. 4,
pt. 8, 1908, p. 20, pi. 1, fig. 4. — De Beauchamp, Arch. Zool. Ex-
per., ser. 4, vol. 10, 1909, p. 160, pi. 3, fig. 22. — Sachse, Susswas-
serfauna Deutschlands, pt. 14, 1912, p. 173, fig. 335. — Jakubski,
Rozpr. Wiad. Muz. Dzieduszyckich, vol. 1, 1914, p. 31. — ^Weber
and Montet, Cat. Invert. Suisse, pt. 11, 1918, p. 190. — Oparina-
Kharitonova, Izv. Biol. Nauchno-Issl. Inst. Permsk. Univ., vol. 3,
1925, p. 44.
Cathypna glandulosa Stokes, Ann. Mag. Nat. Hist., ser. 6, vol. 19,
1897, p. 632, pi. 14, fig. 8-10.
Cathypna magna Lucks, Rotatorienfauna Westpreussens, 1912, p.
109, fig. 32; not Cathypna magna Stenroos.
Cathypna ungulata magna Sachse, Siisswasserfauna Deutschlands,
pt. 14, 1912, p. 173.— Idelson, Trudy Plovuch. Morsk. Nauchn.
Inst. pt. 12, 1925^ p. 89; not Cathypna magna Stenroos.
Cathypna minnesotefisis Murray, Journ Royal Micr. Soc., 1913, p.
345, pi. 13, fig. 18. — Sachse, Arch. Hydrobiol., vol, 10, 1914, p. 69,
fig. 9; not Cathypna minnesotensis Herrick.
Lecane ungulata Harking, Bull. 81 U. S. Nat. Mus., 1913, p. 62;
Proc. U, S. Nat. Mus,, vol. 47, p, 535; Rep. Canadian Arctic Exp.
324 Wisconsin Academy of Sciences, Arts and Letters,
1913 — 18, vol. 8 pt. E, p. 9. — Kozar, Zool. Anz., vol. 44, 1914, p.
420. — Jakubski, Kosmos (Lwow), 1918-1919, p. 28.
The lorica is broadly oval in outline ; the anterior dorsal mar¬
gin is virtually straight, the ventral very slightly concave; at
the external angles are two large, triangular cusps. The dorsal
plate is oval and without markings of any kind ; it is considerably
narrower than the ventral and the margins become indistin¬
guishable before reaching the anterior edge. The ventral plate
is of the same general outline as the dorsal, but somewhat nar¬
rower at the beginning of the posterior segment; its markings
are limited to an indistinct, broken, transverse fold some dis¬
tance in front of the foot joint. The lateral sulci are deep. The
posterior segment is very, broad and slightly truncate; it pro¬
jects slightly beyond the foot; the coxal plates are large and
obtusely pointed. The first foot joint is very indistinct, the
second short and fairly broad. The toes are straight and nearly
parallel-sided, very slightly enlarged posteriorly and end in a
long, stout claw with a fairly prominent basal spicule; their
length is more than one third of the total length.
Total length 285^ ; length of dorsal plate 220/x, of ventral plate
255 fji ; width of dorsal plate 180;u, of ventral plate 195fx; anterior
points 135/>t; length of toes without claw 75/x; claw 45ju,.
Lecane ungulata is very common and widely distributed in
the United States; it seems to be less common in Europe; ac¬
cording to Rousselet it is rare in England. It is the largest
known species of the genus and is readily recognized by its
robust form, the broad posterior segment and long claws.
LECANE SIBINA Harring.
Plate IX, figures 1, 2.
Lecane sibina Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p. 535,
pi. 23, figs. 5-7.
The lorica is broadly pyriform; the anterior margins of the
dorsal and ventral plates are nearly coincident and slightly
concave with distinct lateral cusps. The dorsal plate is subcir¬
cular and without facetting; a low, median, anterior hump
is formed by the retraction of the head. The ventral plate is
oval, slightly narrower than the dorsal, and its only surface
markings are some faint transverse ridges on the posterior third :
the anterior margin has a very small, median, lunate sinus.
Barring & Myers — Rotifer Fauna of Wisconsin, — III. 325
The lateral sulci are moderately deep and do not reach the
anterior margin. The posterior segment is prominent and pro¬
jects some distance over the base of the toes; the large and
obtusely triangular coxal plates do not extend beyond the sec¬
ond foot joint. The first foot joint is very indistinct, the sec¬
ond short and broadly triangular. The toes are very nearly
parallel-sided, about one third of the total length and end in a
long, stout claw with a basal spicule; a short distance in front
of the claw are one or two indistinct annular constrictions.
Total length 200/x; length of dorsal plate 130/a, of ventral plate
135/a; width of dorsal plate 125/a, of ventral plate 116/a, width
of anterior margin 78/a; length of toes without claw 42/a; claw
20/;..
Lecane sihina is related to L. ungulata; it is much smaller
and the lorica is relatively broader. It was described from ma¬
terial collected in the Panama Canal Zone and has since been
found by Mr. C. Juday at Puerto Barrios, Guatemala.
LECANE GRANDIS (Murray).
Plate X, figures I, 2.
Cathypna grandis Murray, Journ. Royal Micr. Soc., 1913, p. 344,
pi. 13, fig. 20.
Lecane grandis Fadeev, Trudy Kharkovsk. Obshch. Isp. Prir., vol.
50, 1925, p. 8, pi. 1, fig. 6.
The outline of the lorica is broadly ovate and truncate pos¬
teriorly. The anterior margins of both dorsal and ventral
plates are nearly straight; the dorsal plate projects slightly be¬
yond the anterior ventral margin and the lorica consequently
remains partly open when the head is completely retracted.
The dorsal and ventral plates are of the same width and differ
very slightly in outline; no surface markings are present. The
lateral sulci are not very deep. The posterior segment is short
and very broad, projecting but little beyond the dorsal plate.
The coxal plates are large and sharply pointed. The foot joints
are unusually broad and rather short. The toes are long, very
slightly blade-shaped and straight on the inner edges, ending in
a conical claw with a small basal spicule.
Total length 240/a; length of dorsal plate 165/a, of ventral
plate 175/a; width of lorica 140/a; anterior margin, dorsal, 100/a,
ventral 118/a; toe without claw 60/a; claw 12/a.
326 Wisconsin Academy of Sciences, Arts and Letters.
Lecane grandis is eommon in brackish or salt tide pools; it
was first found by Murray at Rio de J aneiro ; we have collected
it around Atlantic City, New Jersey, and it has recently been
reported by Fadeev from salt lagoons at Novorossiisk, on the
Black Sea.
L.ECANE LEONTI]\'A (Turner).
Plate X, figures 3-5.
Cathypna leontina Turner, Bull. Denison Univ., vol. 6, 1892, p. 61,
pi. 1, fig. 12. — Kellicott, Trans. Amer. Micr. Soc., vol. 19, 1897,
p. 53. — Jennings, Bull. Michigan Fish Comm., No. 3, 1894, p. 24;
Bull. U. S. Fish Comm., vol. 19 (for 1899), 1900, p. 91, pi. 19,
fig. 25. — Hempel, Bull. Illinois State Lab. Nat. Hist., vol. 5, 1898,
p. 374. — Daday, Zoologica, pt. 44, 1905, p. 109, pi. 6, fig. 12; pt.
59, 1910, p. 83. — Rousselet, Journ. Royal Micr. Soc., 1906, p. 405,
pi. 14, fig. 6. — Kofoid, Bull. Illnois State Lab. Nat. Hist., vol. 6,
No. 1, 1908, p. 197. — Murray, Journ. Royal Micr. Soc., 1913, p.
345, pi. 13, fig. 21.
Cathypna scutaria Stokes, Ann. Mag. Nat. Hist., ser. 6, vol. 19,
1897, p. 631, pi. 14, fig. 7.
Cathypna macrodactyla Daday, Math. Termesz. Ertes., vol. 16,
1898, p. 92; Termesz. Fuzetek, vol. 21, Suppl., 1898, p. 15, fig. 3.
Cathypna leontina bisinuata Daday, Zoologica, pt. 44, 1905, p. 109,
pi. 6, fig. 18.
Cathypna biloba Daday, Math. Termesz. Ertes., vol. 23, 1905, p.
330; zoologica, pt. 44, 1905, p. Ill, pi. 6, fig. 17.
Cathypna incisa Daday, Math. Termesz. Ertes., vol. 23, 1905, p.
330; Zoologica, pt. 44, 1905, p. Ill, pi. 6, fig. 14.
? Cathypna appendiculata Daday, Zoologica, pt. 44, 1905, p. 110, pi.
6, fig. 13.
Lecane leontina Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61;
Proc. U. S. Nat. Mus., vol. 47, 1914, p. 536.
The lorica is broadly pyriform in outline; the anterior dorsal
margin is slightly concave; the ventral margin has a broad,
Y-shaped sinus, rounded at the posterior angle; at the external
angles are two small, triangular spines. The dorsal plate is
nearly as wide as the ventral and of the same general form ; it
is rounded posteriorly and without markings. The ventral plate
has an indistinct transverse fold in front of the foot. The lat¬
eral sulci are deep. The posterior segment is continued over the
base of the foot as a tail-like projection, widest posteriorly, rarely
with two long, divergent spines, as in fig. 3. The coxal plates
are large and obtusely pointed. The first foot joint is parallel¬
sided and somewhat indistinct; the second foot joint is sub-
Earring d' Myers — Rotifer Fauna of Wisconsin. — III. 327
square. The toes are extremely long^ slender, straight and par¬
allel-sided, ending in a fairly long claw with basal spicule ; they
are nearly as long as the entire body.
Total length 330^; length of dorsal plate of ventral
plate 210/a; width of dorsal plate 140/a, of ventral plate 145/a;
anterior points 90/a ; toes without claw 135/a ; claw 15/a. The form
with posterior spines shown in figure 3 measures: total length
370/a; length of dorsal plate 165/a, of ventral plate 240/a; width
of dorsal plate 147/a ; of ventral plate 152/a ; anterior points 90/a ;
toes without claw 140/a; claw 15/a.
Lecane leontina is common in weedy ponds all over the United
States; records from other countries are not numerous. The
variety with posterior spines is rare; the specimen figured was
collected at the Fish Hatchery at Delafield, Wisconsin.
LECANE DEPRBSSA (Bryce).
Plate XVI, figures I, 2.
fCathypna latifrons Gosse, Journ. Royal Micr. Soc., 1887, p. 362,
pi. 8, fig. 3. — Hudson and Gosse, Rotifera, suppl., 1889, p. 42, pi.
31, fig. 37.
? Proales prehensor Gosse, Journ. Royal Micr. Soc., 1887, p. 366, pi.
8, fig. 12.-~Hudson and Gosse, Rotifera, Suppl., 1889, p. 24, pi.
81, fig. 12.
Distyla depressa Bryce, Science Gossip, vol. 27, 1891, p. 205, text
fig.
Cathypna depressa Murray, Journ. Royal Micr. Soc., 1913, p. 555,
pi. 23, fig. 17.
Lecane depressa Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61;
Rep. Canadian Arctic Exp. 1913-18, vol. 8, pt. E, 1921, p. 9.
1 Cathypna levistyla Olofsson, Zool. Bidr. Uppsala, vol. 5, 1917, p.
280, fig. 10.
The outline of the lorica is broadly ovate; its width is about
three fourths of the length. The anterior dorsal margin is nearly
straight; the ventral margin has a shallow, V-shaped sinus. At
the external angles are two small, stout anterior spines. The
dorsal plate is broadly ovate and rounded posteriorly; the ven¬
tral plate is oval and somewhat narrower than the dorsal ; both
are entirely without facetting or surface markings. The lat¬
eral sulci are fairly deep. The posterior segment is fairly large
and projects considerably beyond the dorsal plate; at the level
of the base of the toes are two small, but very distinct marginal
indentations. The coxal plates are rather small and rounded
328 Wisconsin Academy of Sciences, Arts and Letters.
posteriorly. The first foot joint is narrow and overlaps the
second joint with a small, rounded, median lobe only; the sec¬
ond joint is subsquare and some distance in front of the pos¬
terior margin. The toes are fairly long, about one fourth of
the total length, parallel-sided for half their length and taper
to acute points; the inner edges are straight.
Total length 130/a ; length of dorsal plate 90/a, of ventral plate
95/a; width of dorsal plate 75/a, of ventral plate 65/a; width of
anterior margin 54/a ; length of toes 30/a.
Lecane depressa is widely distributed and seems to be a typ¬
ical wet sphagnum species. It is closely related to L. mitis, L.
scohis and L. tudicola; the form of the lorica and the dimen¬
sions show sufficient differences to preclude any confusion.
L.ECANE TUDICOIiA, Harring; and Myers, new species
Plate XI, figures 1, 2.
The outline of the lorica is very broadly ovate; its width is
more than three fourths of the length. The anterior dorsal mar¬
gin is straight; the ventral margin has a very shallow, V-shaped
sinus. At the external angles are two small, stout anterior
spines. The dorsal plate is very broadly ovate and broadly trun¬
cate posteriorly; it is as wide as the ventral plate at its widest
point, but narrower both anteriorly and posteriorly. The ven¬
tral plate is very broadly ovate and slightly larger than the
dorsal; both are without surface markings or facetting. The
lateral sulci are shallow. The posterior segment is broadly
rounded and projects somewhat beyond the dorsal plate ; at the
level of the coxal plates there are two small, but distinct mar¬
ginal indentations. The coxal plates are small and rounded
posteriorly. The first foot joint is fairly large and narrow, with
a small, rounded median lobe overlapping the second joint, which
is large and somewhat reniform and slightly in advance of the
posterior margin. The toes are fairly long, about one fourth of
the total length, parallel-sided for a little less than half their
length and gradually tapering to acute points; the inner edges
are straight.
Total length 145/a; length of dorsal plate 105/a, of ventral
plate 110/a; width of lorica 85/a; width of anterior margin 60/a;
length of toes 37/^.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 329
Lecane tudicola was collected by Dr. G-. H. Parker, of Har¬
vard University, among algae in the ‘^Ice House Pond’’ on St.
Paul, Prybilof Islands, Alaska. It is related to L. depressa, but
readily distinguished by the smaller dorsal plate, as well as its
larger size and relatively greater width.
LECANE MITIS Harriisg and Myers, new species
Plate XI, figures 3, 4.
The outline of the lorica is broadly ovate; its width is about
three fourths of the length. The anterior dorsal margin is
straight medially, with a blunt angle at the external edges;
the ventral margin is slightly lunate. At the external angles
are two small, slightly incurved spines. The dorsal plate is
broadly reversed-ovate and broadly truncate posteriorly; the
ventral plate is elongate oval and considerably narrower than the
dorsal; both are quite smooth, without surface markings. The
lateral sulci are moderately deep. The posterior segment is large
and rounded and projects far beyond the dorsal plate. The
coxal plates are small and obtusely pointed ; they do not project
beyond the foot. The first foot joint is small and broadly ovate
and its small, but very distinct median lobe almost reaches to the
base of the toes ; the second joint is subsquare, very broad and
some distance from the posterior margin. The toes are fairly long,
a little more than one fourth of the entire length, parallel-sided
for fully half their length and taper to acute points; the inner
edges are straight.
Total length lOSft; length of dorsal plate 110/x, of ventral
plate 130/4; width of dorsal plate 100/a, of ventral plate 80/a;
width of anterior margin 60/i,; length of toes 45/a.
Lecane mitis was collected among floating and partly sub¬
merged sphagnum at Grravelly Eun, near Atlantic City, New
Jersey. It is closely related to L. depressa, but differs in the
form of the anterior dorsal and ventral margin, the posterior
segment, the foot and the truncate dorsal plate, as well as in
being consistently larger.
LECANE scorns Harring and Myers, mew species
Plate XI, figures 5, 6.
The outline of the lorica is very broadly ovate; its width is
five sixths of the length. The anterior dorsal margin is straight
330 Wisconsin Academy of Sciences, Arts and Letters.
and the ventral margin slightly concave; at the external angles
are two short, stout and slightly incurved spines. The dorsal
plate is very broadly ovate, its width but little less than its
length, and rounded posteriorly; the ventral plate is broadly
oval and considerably narrower than the dorsal ; both are with¬
out surface markings. The lateral sulci are moderately deepr.
The posterior segment is rounded and very broad; its projects
somewhat beyond the dorsal plate. The coxal plates are large
and rounded posteriorly. The first foot joint is large and pyri¬
form, with a small median lobe overlapping the second joint
which is rounded anteriorly and angulate posteriorly; it is
slightly in advance of the posterior margin. The toes are fairly
long, a little more than one fourth of the total length, parallel¬
sided for a little less than half their length and gradually taper¬
ing to acute points ; the inner edges are straight.
Total length 160/^ ; length of dorsal plate 114/a of ventral plate
122/a; width of dorsal plate 102/a, of ventral plate 88/a; width
of anterior margin 66/a ; length of toes 43/a.
Lecane scohis was collected by Dr. Frits Johansen while serv¬
ing as biologist on the Southern party of the Canadian Arctic
Expedition, among algae growing on stones in the river bed at
Bernard Harbour, North West Territories. It is related to L.
depressa, but readily distinguished by its greater size, relatively
greater width and the form of the foot.
L.ECAXE MUCRONATA Harriiig and Myers, new species.
Plate XXIX, figures 3-5.
The lorica is broadly ovate; its width is about two thirds of
the length. The anterior dorsal and ventral margins are
coincident and straight. At the external angles are two pairs
of anterior spines ; one pair, on the dorsal plate, are very small.
The second pair, attached to the membrane of the lateral sulci,
are very long and stout, broad at the base and acutely pointed ;
when the animal is swimming, these spines point straight up¬
wards, but in the extensive deformations taking place incident
to the contraction and closure of the lorica they are bent out¬
wards, as shown in the figure. The dorsal plate is very broadly
oval and rounded posteriorly ; its width is about four fifths of the
length. The ventral plate is elongate pyriform and considerably
narrower than the dorsal plate. The surface markings of the
Harring & Myers — Rotifer Fauna of Wisconsin. — HI. 331
lorica are a strong, V-shaped anterior fold on the dorsal plate,
formed by the contraction of the lorica, and a transverse fold
on the ventral plate, some distance in front of the foot. The
lateral sulci are fairly deep. The posterior segment is semi¬
circular and rather small; it projects far beyond the dorsal
plate. The coxal plates are rather small and rounded posteriorly.
The first foot joint is indistinct anteriorly and projects with a
small median lobe over the large, subsquare second foot joints
which reaches almost to the posterior margin. The toes are long
and slender, nearly one third of the total length, parallel-sided
and approximately straight, with undulating edges. The claw
is short and stout ; at the base is a small spicule.
Total length 175/*; length of lorica, including frontal spines,
140ju,; length of dorsal plate, without spines, 115/^, of ventral
plate 126y,; width of dorsal plate 105/;i, of ventral plate 84/;t;
length of anterior spines, from base to tip, 28ja; length of toes
without claw 48/^; claw 6/x.
Lecane mucronata has been collected in Oneida and Vilas
Counties, Wisconsin, on Mount Desert Island, Maine, and in New
Jersey around Atlantic City and at Batsto ; it is usually rare, but
at Batsto it was common in the gelatinous algal covering of a
dwarf species of sphagnum growing in shallow water. The
form of the swimming animal has but little resemblance to the
contracted specimens ; the body is very slender, the length of the
lorica alone being equal to the entire length of the contracted
animal, and the frontal spines are pressed towards the head and
the anterior edge of the lorica, so that they are not seen unless
carefully searched for.
LECANE PYRMIA Harring and Myers, new species,
Plate XII, figures 3-6.
The outline of the lorica is a very elongate oval, the greatest
width being only' two thirds of the length, and the body is some¬
what compressed dorso-ventrally. The anterior dorsal and
ventral margins are coincident and concave; at the external
angles are two stout, triangular cusps. The dorsal plate is
elongate oval and slightly truncate posteriorly ; the ventral plate
is very nearly parallel-sided and both are without facetting. The
lateral sulci are fairly deep. The posterior segment is somewhat
irregular in outline and projects but little beyond the dorsal
plate. The coxal plates are unusually small and obtusely point-
332 Wisconsin Academy of Sciences, Arts and Letters.
ed posteriorly. The first foot joint is narrow and wedge-shaped,
the second subsquare and robust. The toes are long and
slender, about one fourth of the total length, straight and
parallel-sided, and end in acute points without claws.
Total length 280/4. ; length of dorsal plate 193/t, of ventral plate
210/4 ; width of dorsal plate 135/4, of ventral plate 125/4 ; width of
anterior points 80/4 ; length of toes 75/4.
Lecane pyrrka is common in weedy ponds with soft, acid
water, but is never found in hard water regions. The entire
body is, with rare exceptions, colored a deep reddish brown. An
aberrant variety from Eagle River, Vilas County, Wisconsin, is
shown in figures 5 and 6; the principal difference is in the
greater width of the lorica. The typical form is abundant at
Atlantic City, New Jersey, in Oneida and Vilas Counties, Wis¬
consin, and on Mount Desert Island, Maine.
LECANE PLOENENSIS (Voigt).
Plate XIII, figures 5, 6.
Distyla ploenensis Voigt, Zool. Anz., vol. 25, 1902, p. 679;
Forschungsber. Biol. Stat. Plon, vol. 11, 1904, p. 71, pi. 4, fig.
89. — Sachse, Susswasserfauna Deutschlands, pt. 14, 1912, p. 177,
fig. 350. — Rousselet, Proc. Royal Irish Acad., vol. 81, pt. 51,
1911, p. 10.
Cathypna ploenensis Murray, Journ. Royal Micr. Soc., 1913, p. 552,
pi. 22, fig. 4.
Lecane ploenensis Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 62;
Proc. U. S. Nat. Mus., vol. 47, 1914, p. 536.
The outline of the lorica is elongate oval. The anterior
margins are coincident and straight; at the external angles are
two small, cusplike spines. The ovate dorsal plate is slightly
truncate posteriorly and very firm; the surface markings are
very prominent and constant. The pattern is rather complicat¬
ed; the median portion is regularly facetted, but towards the
margins the ridges become more complex, as shown in figure 5.
The ventral plate follows the general outline of the dorsal, but is
somewhat narrower, especially in front; the markings are less
intricate than on the dorsal plate. The lateral sulci are
moderately deep. The posterior segment is rather small and
rounded, projecting but little beyond the dorsal plate ; the coxal
plates are large and rounded posteriorly. The first foot joint is
Marring (& Myers — Rotifer Fauna of Wisconsin. — III. 333
narrow and indistinct, the second large and pyriform or subtri-
angular. The toes are very long and slender, about one third of
the total length, parallel-sided, straight on the inner edges and
ending in acute points without claws.
Total length 260ja; length of dorsal plate 170/x, of ventral plate
185/>i ; width of dorsal plate of ventral plate lObju, ; width of
anterior points 84/>i; length of toes 86/a.
Lecane ploenensis is abundant in weedy ponds everywhere in
the United States. According to Rousselet, who collected it on
Clare Island, it is a rare species, and European records are not
numerous. It is quite likely that this may be Levander’s
Cathypna affinis, but his description is lacking in detail, and it is
not now possible to decide among the large number of species
which one was really meant. The only course open seems to be to
drop it as unidentifiable.
LECANE SIGNIFERA (Jennings).
Plate XIII, figures 3, 4.
Distyla signifera Jennings, Bull. Michigan Fish Comm., No. 6,
1896, p. 92, figs. 1, 2.
Cathypna signifera Murray, Journ. Royal Micr. Soc., 1913, p. 552,
pi. 23, fig. 13.
Lecane signifera Barring, Bull. 81 U. S. Nat. Mus., 1913, p. 62.
The outline of the lorica is elongate oval, and the anterior
margins coincident and straight; at the external angles are two
small, cusplike spines. The dorsal plate is slightly ovate and
rounded posteriorly. The ventral plate is narrower than the
dorsal, but of the same general outline. The surface markings
of the lorica are of an unusual type ; upon a pattern of elevated
ridges, rounded at the top and of the design found in L.
ploenensis, is superposed rows of small, closely spaced, hemi¬
spherical, beadlike elevations. These beads are on the slopes of
the ridges, midway between the rounded tops and bottoms of the
facets. The lateral sulci are moderately deep. The posterior seg¬
ment is rounded and projects but slightly beyond the dorsal
plate. The coxal plates are moderately large and rounded
posteriorly. The first foot joint is long and narrow, the second
short, broad and subsquare. The toes are long and slender, about
one third of the total length, parallel-sided, straight on the inner
edges and ending in acute points without claws.
Total length 200/x ; length of dorsal plate 135/a, of ventral plate
334 Wisconsin Academy of Sciences, Arts and Letters.
148/4,; width of dorsal plate 90/t, of ventral plate 82/x; width of
anterior points 68/a ; length of toes 60/a.
Lecane signifera usually occurs in small numbers, but it is
widely distributed in the United States. We have collected it in
Wisconsin, Mount Desert Island, Maine, New Jersey, Maryland,
the District of Columbia and Florida, and it also occurred in
collections made by Dr. Jennings at Ann Arbor, Michigan.
LiECANE AQUIIiA Harring’ aud Myers, new species.
Plate XIII, figures 1, 2.
The outline of the lorica is reversed-ovate and fairly broad.
The anterior margins are coincident and very nearly straight;
at the external angles are two small cusplike spines. The ovate
dorsal plate is slightly truncate posteriorly and rather flexible;
the surface markings are very faint and differ from other species
of the genus, as will be seen from figure 2. The ventral plate is
oval and its anterior edges somewhat flexible and indistinct.
The lateral sulci are moderately deep. The posterior segment
is rounded and projects but very slightly beyond the dorsal
plate; the coxal plates are moderately large and rounded pos¬
teriorly. The first foot joint is narrow and indistinct, the sec¬
ond large and subsquare. The toes are very long and slender,
one third of the total length, parallel-sided and ending in acute
points without claws.
Total length 300/a; length of dorsal plate 190/a, of ventral
plate 202/a; width of dorsal plate 148/a, of ventral plate 125/a;
width of anterior points 95/a ; length of toes 100/a.
' Lecane aquila was first collected in a shallow, weedy pond at
Eagle Kiver, Vilas County, Wisconsin; it is common on Mount
Desert Island, Maine. It is closely related to L. ploenensis and
L. signifera, but is much larger and more robust than these two
species ; the toes are also relatively longer.
LECANE LUNA (Miiller)
Plate XIV, figures 5, 6.
Cercaria luna Muller, Zool. Danicae Prodr., 1776, p. 280; Animal-
cula Infusoria, 1786, p. 139, pi. 20, figs. 8, 9.
Furcocerca luna Lamarck, Hist. Nat. Anim. sans Vert., vol. 1,
1815, p. 448.
Trichocerca luna Bory de St. Vincent, Class. Anim. Micr., 1826,
p. 42.
Earring Myers — Rotifer Fauna of Wisconsin. — III. 335
Lecane luna Nitzsch, Encycl. Wiss. u. Kiinste sect. 1, vol. 16,
1827, p. 68.— -Harking, Bull. 81 U. S. Nat. Mus., 1913, p. 61;
Proc. U. S. Nat. Mus., vol. 47, 1914, p. 534; Rep. Canadian Arctic
Exp. 1913-18, vol. 8, pt. E, 1921, p. 9. — Kozar, Zool. Anz., vol.
44, 1914, p. 420. — Rezvoi, Trudy Borodinsk. Biol. Stants. Imp.
Petrogradsk. Obshch. Estestvoisp., vol. 4, 1916, p. 180. — Jakub-
SKi, Kosmos (Lwow), 1918-1919, p. 28.
? Furcularia jobloti Bory de St. Vincent, Encycl. Meth., Zooph.
(pt. 2), 1827, p. 425.
Brachionus luna Blainville, Diet. Sci. Nat., vol. 60, 1830, p. 148.
Euchlanis luna Ehrenberg, Abh. Akad. Wiss. Berlin (for 1831),
1832, p. 131; Infusionsthierchen, 1838, p. 462, pi. 57, fig. 10. —
Dujardin, Hist. Nat. Zooph. Inf., 1841, p. 634. — Perty, Zur
Kenntn. kleinst. Lebensf., 1852, p. 41. — Bartsch, Jahresh.
Naturk. Wurttemberg, vol. 26, 1870, p. 357; Rotat. Hungariae,
1877, p. 45. — E5rferth, Mikr. Siisswasserbew., 1877, p. 54, fig. 92;
Einf. Lebensformen, 1878, p. 89, pi. 5, fig. 37; ed. 2, 1885, p. 114,
pi. 7, fig. 37. — Plate, Jenaische Zeitschr. Naturw., vol. 19, 1885,
p. 59. — Daday, Termesz. Fuzetek, vol. 9, 1885, p. 127.
Cathypna luna Gosse, in Hudson and Gosse, Rotifera, 1886, vol.
2, p. 94, pi. 24, fig. 4. — Daday, Termesz. Fuzetek, vol. 15, 1892,
p. 28; Math. Termesz. Ertes., vol. 12, 1893, p. 31. — Petr, Sit-
zungsber. Bohm. Ges. Wiss. (for 1890), 1892, p. 233. — Ternetz,
Rotat. Umg. Basels, 1892, p. 17. — Wierzejski, Rozpr. Akad.
Umiej., Wydz. Mat. Przyr., Krakow, ser. 2, vol. 6, 1893, p. 242. —
Kertesz, Budapest Rotat. Faun., 1894, p. 39. — Levander, Acta Soc.
Fauna et Flora Fennica, vol. 12, No. 3, 1895, p. 49, pi. 3, fig. 29.
— Skorikov, Trav. Soc. Nat. Kharkow, vol. 30, 1896, p. 319. —
Hempel, Bull. Illinois State Lab. Nat. Hist., vol. 5, 1898, p. 374.
— Weber, Rev. Suisse Zool., vol. 5, 1898, p. 593, pi. 22, figs. 4, 5. —
Jennings, Bull. U. S. Fish Comm, vol. 19 (for 1899), 1900, p. 91,
pi. 19, figs. 28, 29.— Voronkov, Trudy Gidr. Stants. Glubokom
Oz., vol. 2, 1907, pp. 109, 204, 283. — Schlenker, Mitt. Geol. Abt.
Wurttemberg. Stat. Landesamt, No. 5, 1908, p. 249. — Kofoid,
Bull. Illinois State Lab. Nat. Hist., vol. 8, No. 1, 1908, p. 198. —
Meissner, Izv. Turkestansk. Otd. Imp. Russk. Geogr. Obshch.,
vol. 4, pt. 8, 1908, p. 20. — De Beauchamp, Arch. Zool. Exper.,
ser. 4, vol. 10, 1909, p. 210, fig. XXVI. — VoN Hofsten, Arkiv
Zool., Stockholm, vol. 6, No. 1, 1909, p. 56; Wiss. Unters. Sarek-
geb., vol. 4, pt. 8, 1923, p. 860. — Lie Pettersen, Bergens Mus.
Aarbog (for 1909), 1910, p. 63. — Daday, Zoologica, pt. 59, 1910,
p. 85. — Lucks, Rotatorienfauna Westpreussens, 1912, p. 107. —
Sachse, Stisswasserfauna Deutschlands, pt. 14, 1912, p. 171, figs.
336-338. — Mola, Ann. Biol. Lac., vol. 6, 1913, p. 259. — Murray,
Journ. Royal Micr. Soc., 1913, p. 345, pi. 13, fig. 19. — Jakubski,
Rozpr. Wiad. Muz. Dzieduszyckieh, vol. 1, 1914, p. 30, — Montet,
Rev. Suisse Zool., vol. 23, 1915, p. 335. — Weber and Montet, Cat.
Invert. Suisse, pt. 11, 1918, p. 188. — Behning, Izv. Kaluzhsk.
336 Wisconsin Academy of Sciences, Arts and Letters.
Obshch. Izuch. Prir. Mestn. Kraia. vol. 3, 1918, p. 92. — Monard,
Rev. d’HydroL, vol. 2, 1922, p. 13. — Bryce, Journ. Quekett Micr.
Club, ser. 2, vol. 15, 1924, p. 97.
The lorica is broadly pyriform or subcircular; its greatest
width is nearly equal to the length. The dorsal and ventral
plates have a deep, lunate anterior sinus. The dorsal plate is
almost circular in outline and without any surface markings;
the angles of the sinus are very blunt and without spines. A
broad, median hump is formed by the excessive contraction of
the anterior margin. The ventral plate is very little narrower
than the dorsal and slightly more ovate in outline; the only
surface marking present is a straight transverse fold imme¬
diately in front of the foot. The lunate anterior sinus is cuspi¬
date at its external angles. The lateral sulci are unusually
deep. The posterior segment of the body is very small and
rounded, projecting very little beyond the dorsal plate; the
coxal plates are bluntly pointed at their posterior, free ends.
The first foot joint is small and indistinct, the second very large
and subsquare. The toes are parallel-sided, about one third of
the total length, and end in a distinct claw with a minute basal
spicule.
Total length 180/;t ; length of dorsal plate 125/^, of ventral plate
135;u; width of dorsal plate 125ja, of ventral plate 115/x; width
of dorsal sinus 40yLi, of ventral sinus 65/x; length of toe without
claw 55yu,; claw 9/^.
Lecane tuna occurs in abundance in weedy ponds all over the
world and is without doubT one of the commonest of all rotifers.
It is readily recognized by its large size, nearly circular outline
and deep anterior sinus.
LECAXE PAPUAN A (Murray)
Plate XIV, figures 3, 4.
Cathypna papuana Murray, Journ. Royal Micr. Soc., 1913, p. 551,
pi. 22, fig. 2.
Lecane papiiana Harring, Proc. U. S. Nat. Mus., vol. 47, 1914,
p. 534.
The outline of the lorica is subcircular; its width is very
nearly equal to the length. The anterior dorsal margin is nearly
straight. The ventral margin has a broad, shallow, V-shaped an¬
terior sinus between two rounded lobes projecting considerably
beyond the dorsal plate; the sinus is rounded posteriorly and
Earring & Myers — Rotifer Fauna of Wisconsin.— -Ill . 337
its sides undulate. The dorsal plate is subcircular and rounded
posteriorly. The ventral plate is slightly narrower than the
dorsal and of the same outline; it has a transverse fold in front
of the foot. The lateral sulci are moderately deep. The pos¬
terior segment is small and rounded ; it projects slightly beyond
the dorsal plate. The coxal plates are small and obtusely
pointed. The first foot joint is pyriform and rounded pos-
teriorly^ the second joint robust and very broadly ovate. The
toes are long and fairly slender, more than one fourth of the
total length, nearly parallel-sided, straight on the inner margins
and very slightly broadened externally in front of the stout,
acutely pointed claw, which has two basal spicules.
Total length 160/x ; length of dorsal plate 102]Li, of ventral plate
115ft; width of dorsal plate 102ft, of ventral plate 98/x; width of
anterior dorsal margin 52ft, of ventral margin 62ft; length of
toes without claw 38ft; claw 9ft.
Lecane papuana was described by Murray from a single speci¬
men from New Guinea. It is common in the Panama Canal
Zone and in material collected by Mr. Juday at Puerto Barrios,
Guatemala ; in the United States -we have found it only in Polk
County, Florida.
LECANE BHACHYDACTYLA (Stenroos).
Plate XV, figures 5, 6.
Cathypna brachydactyla Stenroos, Acta Soc. Fauna et Flora Fen-
nica, vol. 17, No. 1, 1898, p. 160, pi. 2, fig. 20. — Rousselet, Journ.
Quekett Micr. Club, ser. 2, vol. 11, 1912, p. 372, pi. 13, fig. 3. —
Lucks, Rotatorienfauna Westpreussens, 1912, p. 108. — Murray,
Journ. Royal Micr. Soc., 1913, p. 554, pi. 23, fig. 15. — Sachse,
Arch. HydrobioL, vol. 9, 1914, p. 68, fig. 8.— Oparina-Kharito-
NOVA, Izv. Biol. Nauchno-Issl. Inst. Permsk. Univ., vol. 3, 1925,
p. 439.
Cathypna luna brachydactyla Sachse, Susswasserfauna Deutsch-
lands, pt. 14, 1912, p. 172, — Jakubski, Rozpr. Wiad. Muz. Dzie-
duszyckich, vol. 1, 1914, p. 31.
Distyla truncata Leissling, Arch. HydrobioL, vol. 9, 1914, p. 255,
fig. 2.
Lecane brachydactyla Harking, Bull. 81 U. S. Nat. Mus., 1913, p.
60; Rep. Canadian Arctic Exp, 1913-18, vol. 8, pt. E, 1921, p. 9,
pi. 8, figs. 5, 6.
The outline of the lorica is ovate, truncate posteriorly and the
anterior margin cuspidate. The dorsal plate is broadly oval,
22
338 Wisconsin Academy of Sciences, Arts and Letters,
truncate posteriorly and somewhat narrower than the ventral
plate. The anterior margin of the dorsal plate is approximately
straight and slightly wavy with prominent lateral cusps; the
ventral margin has a median concavity. The lorica is strongly
compressed dorso-ventrally and without surface markings; the
lateral sulci are fairly deep. The posterior segment is very
prominent and trapezoidal in outline, slightly rounded at the
angles. The coxal plates are circular and fairly large. The
first foot joint is somewhat elongate, widest posteriorly, and the
second joint suhsquare. The toes are short, straight on the
inner edges, cylindrical at the base and end in long, conical
points.
Total length 150/x ; length of dorsal plate 100/x, of ventral plate
128/a; width of dorsal plate 92/a, of ventral plate 100/a; anterior
points 72/a ; length of toes 28/a.
Lecane hrachydactyla is fairly common among sphagnum in
soft, acid water ponds; we have collected it in Vilas and Oneida
Counties, Wisconsin, around Atlantic City, New Jersey, and on
Mount Desert Island, Maine.
LECANE JESSUPI Harring
Plate XV, figures 3, 4.
Lecane jessupi Harking, Rep. Canadian Arctic Exp. 1913-18, vol 8,
pt. E, 1921, p. 8, pi. 3, figs. 3, 4.
The outline of the lorica is slightly ovate, truncate posteriorly
and the anterior margin cuspidate. The dorsal plate is ovate,
rounded posteriorly and slightly narrower than the ventral plate,
which is somewhat elliptic. The anterior margin of the dorsal
plate is slightly convex for the greater portion of its width and
excavate at the lateral cusps ; the anterior margin of the ventral
plate is lunate. There are no markings on either dorsal or ven¬
tral plate ; the lateral sulci are deep. The lorica is strongly com¬
pressed dorso-ventrally. The posterior segment is roughly
trapezoidal in outline and cuspidate at the external angles; the
margin is convex in the median portion and has a slight concav¬
ity at the angles. There is a well marked constriction at the
junction of the ventral plate and the posterior segment. The
coaxal plates are semi-ovate. The first foot joint is well marked
and widest posteriorly ; the second joint is subsquare. The toes
are short, cylindric for one half their length and end in acute,
conical points, straight on their inner edges.
Harring d; Myers — Rotifer Fauna of Wisconsin. — III. 339
Total length 126/>i ; length of dorsal plate 93/a, of ventral plate
108/x; width of dorsal plate 93/a, bi ventral plate 96/a; anterior
points 58/a; length of toes 27/a.
Lecane jessupi was collected by Mr. J. M. Jessup in lakes
on Old Crow River flats, north of New Rampart House, Alaska,
during the Alaskan Boundary Survey. It was not abundant and
has not been found elsewhere.
LECANE LIGONA (Dunlop)
Plate XVI, figures 3-6.
Cathypna ligona Dunlop, Journ. Quekett Micr. Club, ser. 2, vol. 8,
1901, p. 29, pi. 2, figs. 4-6. — Voigt, Forschungsber. Biol. Stat.
Plon, vol. 11, 1904, p. 70, pi. 4, fig. 38. — Lie-Pettersen, Bergens
Mus. Aarbog (for 1909), 1910, No. 15, p. 64, pi. 2, fig. 14. —
Lucks, Rotatorienfauna Westpreussens, 1912, p. 109. — Sachse,
Siisswasserfauna Deutschlands, pt. 14, 1912, p. 174, fig. 342. —
Murray, Journ. Royal Micr. Soc., 1913, p. 554, pi. 23, fig. 16.
Lecane ligona Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61; Rep.
Canadian Arctic Exp. 1913-18, vol. 8, pt. E, 1921, p. 8.
The outline of the lorica is a broad oval, squarely truncate
posteriorly and convex anteriorly. The dorsal plate is slightly
ovate, truncate posteriorly and narrower than the ventral plate.
The anterior margin of the dorsal plate is distinctly convex with
small lateral spines ; the ventral margin is concave. The lorica
is strongly compressed dorso-ventrally and without surface
markings; the lateral sulci are deep. The posterior segment is
short and very broad, produced laterally as two triangular
cusps; its posterior margin is usually somewhat wavy. The
coxal plates are semi-ovate and rather indistinct. The first foot
joint is bluntly pointed; the second joint is rectangular. The
toes are short, straight on their inner margins, cylindrical at the
base and end in conical points.
Total length 92/a; length of dorsal plate 66/a, of ventral plate
80/a; width of dorsal plate 60/a, of ventral plate 70/a; anterior
points 42/a; length of toes 18/a.
Lecane ligona is not rare among sphagnum growing in soft,
acid water ponds; strangely enough it occurs also in brackish
ditches in the tide marshes around Atlantic City, New Jersey
European records of its occurrence are not numerous, but this
may possibly be accounted for by the fact that collections are
generally from hard water regions. We have found it especially
340 Wisconsin Academy of Sciences^ Arts and Letters.
abundant in Starvation Lake, Vilas County, Wisconsin, and
common in ponds and lakes in the northern part of the state, as
well as in New Jersey and on Mount Dessert Island, Maine. A
variety with rounded posterior angles, shown in figures 5 and 6,
is found in Bubble Pond and Long Pond, Mount Desert Island.
liECAlVE PYCIJVA Harring and Myers, new species
Plate XVII, figures 5, 6.
The outline of the lorica is broadly ovate ; its width is about
three-fourths of the length. The anterior dorsal margin is
slightly convex and the ventral margin has a shallow sinus,
rounded posteriorly ; at the external angles are two acute spines.
The dorsal plate is ovate and rounded posteriorly. The ventral
plate is oval and slightly narrower than the dorsal. The lorica
is strongN compressed dorso-ventrally and without surface mark¬
ings ; the lateral sulci are deep. The posterior segment is rather
short and very broad with a short, relatively broad projection,
slightly sinuate posteriorly and with small lateral, triangular
cusps. The coxal plates are fairly large and rounded posteriorly.
The first foot joint is hardly traceable, the second joint is small
and subsquare. The toes are short, about one fifth of the total
length, straight on their inner margins and convex externally,
ending in acute points.
Total length 150/^; length of dorsal plate 120/x, of ventral plate
135|U,; width of dorsal plate 100/x, of ventral plate 95/u,; width of
anterior points 70/x; length of toes 30/>t.
Lecane pycina was collected in Long Lake and Bubble Pond,
Mount Desert Island^ Maine. It is related to L. ligona, L. bra-
chydactyla and L. jessupi, but differs from these species in hav¬
ing the dorsal plate wider than the ventral, as well as in the
form of the posterior projection.
L.ECA1VE PERTICA Marring and Myers, new species
Plate XII, figures 1, 2.
The outline of the lorica is a very elongate oval, the greatest
width being only two thirds of the length; the body is com¬
pressed dorso-ventrally. The anterior margins are very nearly
coincident, the dorsal almost straight and the ventral slightly
concave; at the external angles are two small, pointed spines.
The dorsal plate is oval and squarely truncate posteriorly; the
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 341
facetting is very distinct and consists of the usual four rows,
the first somewhat broken and irregular. The ventral plate is
oval and somewhat narrower than the dorsal ; the surface mark¬
ings are prominent. The lateral sulci are deep. The posterior
segment is nearly semicircular and projects considerably beyond
the dorsal plate. The coxal plates are large and obtusely pointed
posteriorly. The first foot joint is long and narrow, the second
robust and subsquare, projecting the greater part of its length
beyond the posterior segment. The toes are extremely long and
very slender, about one third of the total length, parallel-sided
and straight, and end in acute points without claws.
Total length 260/a ; length of dorsal plate 140/a, of ventral plate
160/a ; width of dorsal plate 100/a, of ventral plate 88/a ; width of
anterior points 64/a ; length of toes 85/a.
Lecane pertica is very common in weedy ponds with soft, acid
water, but is never found in hard water. We have collected it
in Oneida and Vilas Counties, Wisconsin, Polk County, Florida,
Mount Desert Island, Maine, and at Atlantic City, New Jersey.
LECAjVE EUTARSA Harring and Myers, new species.
Plate XVIII, figures I, 2.
The outline of the lorica is ovate and the body is compressed
dorso-ventrally. The anterior margins are coincident and very
slightly concave; at the external angles are two small spines.
The dorsal plate is ovate, truncate posteriorly and narrow at
the anterior margin. The surface markings consist of four trans¬
verse rows of not very prominent facets. The ventral plate is
ovate and very slightly narrower than the dorsal ; the markings
consist of a number of rather indistinct longitudinal ridges as
shown in figure 2. The posterior segment is narrow and rounded ;
it projects somewhat beyond the dorsal plate. The coxal plates
are indistinct, closely appressed to the foot, and pointed poste¬
riorly. The lateral sulci are fairly deep. The first foot joint is
long and narrow, the second joint unusually large and some¬
what angular. The toes are long and very slender, more than
one fourth of the total length, straight and parallel-sided with a
very long, spine-like claw with a small basal spicule.
Total length 45/a ; length of dorsal plate 95/a, of ventral plate
102/a; width of dorsal plate 64/a, of ventral plate 60/a; width of
anterior points 44/a; length of toe without claw 30/a, claw 12/a.
342 Wisconsin Academy of Sciences, Arts and Letters.
Lecane eutarsa was found in small numbers in material col¬
lected by Mr. Juday in ditches along the railroad, near Puerto
Barrios, Guatemala. It has not been found elsewhere.
LiECA]VE PEIjATIS Harring and Myers, new species
Plate XVIII, figures 3, 4.
The outline of the body is broadly oval or very slightly ovate.
The anterior margins are coincident and somewhat concave; at
the external angles are two distinct cusps. The ovate dorsal
plate is truncate posteriorly; the surface markings are fairly
prominent and consist of four rows of facets. The ventral plate
is oval and considerably narrower than the dorsal ; the markings
are distinct and less simple than is usually the case. The
posterior segment is rounded and fairly prominent ; the coxal
plates are large and pointed posteriorly. The lateral sulci are
deep. The first foot joint is elongate oval and pointed posterior¬
ly, the second joint large and robust, squarely truncate posterior¬
ly, and rounded at the naterior end. The toes are long and
slender, about one fourth of the total length, parallel-sided and
terminate in acute claws.
Total length 160/x ; length of dorsal plate llOfi, of ventral plate
122fjL; width of dorsal plate 92/1, of ventral plate 70/x; Avidth of
anterior points 53/>t ; length of toe without claw 35jLi ; claw 9/a.
Lecane pelatis is common in weedy ponds around Minocqua,
Wisconsin, at Atlantic City, New Jersey, and on Mount Desert
Island, Maine.
EECAXE MIRA (Murray)
Plate XVIII, figures 5, 6.
Cathypna niira Murray, Journ. Royal Micr. Soc., 1913, p. 553, pi.
22, fig. 3.
The outline of the lorica is oval and the body is quite deep.
The anterior dorsal margin is nearly straight and the ventral
slightly concave ; at the external angles are two stout, triangular
cusps. The dorsal plate is broadly oval or subcircular and slight¬
ly truncate posteriorly ; the surface markings are not very prom¬
inent and differ somewhat from the usual pattern, as the anterior
row of facets is incomplete. The ventral plate is broadly ovate
and considerably narrower than the dorsal; it is not facetted.
The lateral sulci are deep. The posterior segment is small and
Barring & Myers — Rotifer Fauna of Wisconsin. — 111. 343
truncate, projecting considerably beyond the dorsal plate. The
coxal plates are fairly large and obtusely pointed posteriorly.
The first foot joint is long and narrow, the second large and
pyriform. The toes are long and slender, about one fourth of the
total length, parallel-sided and straight, terminating in a stout
claw with a minute basal spicule.
Total length 200/x ; length of dorsal plate 132/>i, of ventral plate
145/x ; width of dorsal plate 130/>t, of ventral plate lOO^u, ; width of
anterior points 60/x ; length of toes without claw 40/>i ; claw
Lecane mira is fairly common in weedy ponds with neutral or
acid water. We have collected it in Oneida and Vilas Counties,
Wisconsin, on Mount Desert Island, Maine, in New Jersey, Dis¬
trict of Columbia and Florida ; it occurs also in material collected
by Dr. Birge and Mr. Juday in Arkansas, Louisiana and Texas.
Murray’s original description was based on material from Wash¬
ington.
LECANE METHORIA Harring: and Myers
Plate XIX, figures 1, 2.
The outline of the lorica is broadly ovate, the width about two
thirds of the length. The anterior dorsal and ventral margins are
coincident and very faintly convex; at the external angles are
two stout, triangular spines. The dorsal plate is broadly ovate
and very slightly truncate posteriorly ; the facetting is of an un¬
usual pattern and the separate facets are bounded by double
lines, as in L. Tialiclysta. The ventral plate is considerably
narrower than the dorsal ; its margins are fiexible and the outline
consequently somewhat indefinite, but approximating an obtusely
pointed triangle. The ventral markings are quite intricate and
some of the lines are doubled, as on the dorsal plate. The lateral
sulci are very indistinct. The posterior segment is very small
and rounded and projects slightly beyond the dorsal plate. The
coxal plates are fairly large and somewhat pointed at their
posterior ends. The first foot joint is very narrow and elongate,
its outlines merging gradually and without distinct separation
with the markings of the ventral plate ; its posterior, obtusely
pointed termination projects over the robust, subsquare second
foot joint. The toes are long and moderately slender, about one
fourth of the total length, parallel-sided and very slightly incurv¬
ed ; the inner margins are excavate at the ends, but a true claw
is not present.
344 Wisconsin Academy of Sciences, Arts and Letters.
Total length 102/>t; length of dorsal plate lOfx, of ventral plate
75/x; width of dorsal plate 54/a, of ventral plate 45/a; width of
anterior points 50/a ; length of toes 24/a.
Lecane methoria was collected in small numbers in weedy bays
of Town Line Lake, at Three Lakes, Oneida County, Wisconsin.
It has not been found elsewhere.
LECANE STICKAEA Harring
Plate XIX, figures 3, 4.
Lecane stichaea Harking, Proc. U. S. Nat. Mus., vol. 46, 1913, p.
397, pi. 35, figs. 4-6. — Rezvoi, Trudy Borodinsk. Biol. Stants.
Imp. Petrogradsk. Obshch. Estestvoisp., vol. 4, 1916, p. 180, pi. 1,
figs. 11, 12. — Fadeev, Russk. Gidr. Zhurn., vol. 3, No. 3-5, 1924,
p. 4.
The outline of the lorica is broadly ovate and the body is some¬
what compressed dorso-ventrally. The anterior margins are
coincident and slightly convex; at the external angles are two
stout spines. The dorsal plate is ovate and truncate posteriorly.
As shown in figure 3, the surface markings are of a rather
aberrant pattern. The normal number of transverse rows of
facets are present, but the first row is irregular and shows a
tendency to subdivide ; in the second to fourth row the facets are
more numerous than in other species of the genus. The ventral
plate is very nearly parallel-sided anteriorly and rounded
posteriorly; the markings are fairly prominent and rather in¬
tricate. The coxal plates are large and somewhat indistinct,
closely appressed to the foot and pointed posteriorly. The
lateral sulci are moderately deep. The first foot joint is long and
narrow, the second large and subsquare. The toes are long and
slender, more than one fourth of the total length, straight and
parallel-sided ; the claw is fairly large, acutely pointed and with¬
out basal spicule.
Total length 145/a ; length of dorsal plate 92/a, of ventral plate
105/a; width of dorsal plate 76/a, of ventral plate 58/a; width of
anterior points 64/a; length of toes without claw 32/a; claw 7/a.
Lecane stichaea is common in wet sphagnum ; we have collected
it in Wisconsin, Mount Desert Island, Maine, New Jersey, Mary¬
land, Virginia and Florida.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 345
LrECANE SAGINATA Harring and Myers, new species
Plate XIX, figures 5, 6.
The outline of the loriea is broadly ovate ; the greatest width is
about three fourths of the length. The anterior margins of the
dorsal and ventral plates are coincident and straight, but slight¬
ly wavy; at the external angles are two short, stout spines.
The dorsal plate is broadly ovate and rounded posteriorly. The
facetting is well marked and differs, as may be seen from the
figure, only in details from the usual pattern. The ventral plate
is narrower than the dorsal, ovate in outline and the edges ill-de¬
fined and wavy. The surface markings are fairly prominent and
more complex than the prevailing design. The posterior segment
is small and rounded, with a very slight median concavity, and
projects but little beyond the dorsal plate. The coxal plates are
fairly large and obtusely pointed. The first foot joint is
moderately large, pyriform and somewhat indistinct ; the second
joint is large and pyriform and projects very slightly beyond the
posterior segment. The toes are long and slender, more than one
fourth of the total length, parallel-sided and very slightly in¬
curved; the tip is excavate on the inner margin, forming a
pseudo-claw.
Total length 140/x; length of dorsal plate of ventral plate
81/x; width of dorsal plate 66;u, of ventral plate 54/x; width of
anterior points 50/a; length of toes without claw 33/a; claw 7/a.
Lecane saginata occurs in soft, acid water ponds in Oneida and
and Vilas Counties, Wisconsin, on Mount Desert Island, Maine,
and around Atlantic City, New Jersey; it is nowhere common.
Its nearest relative is L. intrasinuata; this species is consistently
larger, the loriea is relatively broader and oval in outline, with¬
out facetting, while the loriea of L. saginata is distinctly pyri¬
form and the dorsal facetting prominent.
I/ECANE ELiASMA Harring: and Myers, new species
Plate XX, figures 1, 2.
The loriea is membranous and very flexible, but its form is
quite constant; the outline is broadly ovate and rounded pos¬
teriorly. The anterior margins are coincident and straight; at
the external angles are two small, but stout spines. The dorsal
plate is broadly oval and rounded posteriorly; it is without
346 Wisconsin Academy of Sciences, Arts and Letters.
facetting or folds of any kind. The ventral plate is oval and
slightly narrower than the dorsal plate; the surface markings
are faint, but constant. The lateral sulci are moderately deep.
The posterior segment is fairly large and rounded; it projects
considerably beyond the dorsal plate. The coxal plates are
small and pointed posteriorly. The first foot joint is pyriform
and somewhat indistinct, lobate posteriorly, the second joint
large and rounded anteriorly, abruptly reduced above the base
of the toes and projecting half its length beyond the lorica. The
toes are very long and slender, about one third of the total
length, parallel-sided for half their length and somewhat
abruptly reduced to long, excessively slender and acute points.
Total length 130/x ; length of dorsal plate 75/>i, of ventral plate
85/a; width or dorsal plate 65/a, of ventral plate 56/a; width of
anterior points 46/a ; length of toes 40/a.
Lecane elasma is not rare in wet sphagnum ; we have collected
it around Washington, District of Columbia, at Tuckerton, New
Jersey, and in Northern and Central Wisconsin.
LiECANE RHYTIDA Marring and Meyers, new speeies
Plate XX, figures 3, 4.
The outline of the lorica is broadly ovate ; the width is about
two thirds of the length. The anterior dorsal and ventral mar¬
gins are coincident and very slightly concave, produced at the
external angles into two stout, triangular cusps. The dorsal
plate is oval and very slightly truncate posteriorly. The facet¬
ting is prominent and does not depart very much from the usual
pattern. The ventral plate is elongate ovate and considerably
narrower than the dorsal ; the surface markings are few in num¬
ber, but prominent. The lateral margins of the ventral plate
are somewhat wavy and indistinct and the lateral sulci ill-de¬
fined. The posterior segment is small and semicircular ; it pro¬
jects somewhat beyond the dorsal plate; the coxal plates are
small and obtusely pointed posteriorly. The first foot joint is
indistinct and elongate pyriform; the second joint is roughly
hexagonal and quite robust, projecting fully half its length be¬
yond the lorica. The toes are long and slender, about one fourth
of the total length, parallel-sided for half their length and taper¬
ing to extremely long, acute points.
Total length 126/a ; length of dorsal plate 80/a, of ventral plate
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 347
ST/x; width of dorsal plate 69^, of ventral plate 65ja; width of
anterior points 42/x ; length of toes 39/a.
Lecane rhytida is not rare among floating sphagnum in
soft, acid water ponds. We have collected it in a large pond
at Gravelly Kun, near Atlantic City, New Jersey, and in Upper
Mill Meadow pond on Mount Desert Island, Maine.
LECANE liAlJTERBORNI Hauer
Plate XX, figures 5, 6.
Lecane lauterborni Hauer, Zool. Anz., vol. 61, 1924, p. 145, figs. 1-3.
The lorica is firm and its outline broadly reversed-ovate ; its
width is more than three fourths of the length. The anterior
margins are coincident and slightly convex. At the external
angles are two stout frontal spines. The dorsal plate is broadly
ovate and indistinctly facetted. The ventral plate is nearly
parallel-sided for half its length, obtusely triangular poste¬
riorly and much narrower than the dorsal plate; its surface
markings are of a very simple pattern. The lateral sulci are
deep. The posterior segment is obtusely triangular and projects
far beyond the dorsal plate. The coxal plates are indistinct,
small and oval. The first foot joint is short, broad and some¬
what ill-defined, the second joint long and narrow. The toes
are long and slender, more than one fourth of the total length,
straight, parallel-sided for about one third of their length, and
taper to very acute points.
Total length 150/a ; length of dorsal plate 92/a, of ventral plate
110/a; width of dorsal plate 82/a, of ventral plate 70/a; width of
anterior spines 65/a ; length of toes 42/a.
Lecane lauterborni was described by Hauer from sphagnum
bogs in the Schwarzwald, where it is common. We have col¬
lected this species in shallow, weedy areas of Witch Hole, Mount
Desert Island, Maine. Material forwarded to us by Hauer
shows the German form to be consistently smaller, about two-
thirds the size of the Mount Desert specimens.
LECANE COMPTA Marring
Plate XXI, figures 1, 2.
Cathypna flexilis Murray, Journ. Royal Micr. Soc., 1913, p. 351,
pi. 14, fig. 27; not Distyla flexilis Gosse.
Lecane compta Harring, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
540, pi. 20, figs. 1-3.
348 Wisconsin Academy of Sciences, Arts and Letters.
The outline of the lorica is a slightly elongate oval ; the width
of the body is three fourths of the total length. The anterior
margins of the dorsal and ventral plates are coincident and
straight; at the external angles are two short, robust spines.
The dorsal plate is ovate and narrowly truncate posteriorly ; the
surface markings are fairly prominent and consist of the usual
four rows of facets, the first row somewhat irregular. The ven¬
tral plate is narrower than the dorsal, parallel-sided anteriorly
and rounded posteriorly; its surface markings are prominent
and of an irregular pattern, as shown in figure 2. The lateral
sulci are indistinct and the entire lorica very flexible. The
posterior segment is large and bluntly pointed; it projects con¬
siderably beyond the dorsal plate. The coxal plates are small
and rounded posteriorly. The first foot joint is oval and some¬
what indistinct, the second joint large and subcircular. The
toes are long and slender, more than one fourth of the entire
length, straight and parallel-sided, with conical points ending
in an acute spicule.
Total length llb/x; length of dorsal plate 70/^, of ventral plate
80/x; width of dorsal plate 60/a, of ventral plate 56/a; width of
anterior points 45/a ; length of toes 30/a.
Lecane compta was originally described from material col¬
lected in the Panama Canal Zone ; we have since found it in small
numbers in Southern Wisconsin. It seems to be confined to
moderately hard waters.
LECANE HAIilCLiYSTA Harring; and Myers, new species
Plate XXI, figures 5, 6.
The outline of the lorica is broadly ovate and the width of the
body is nearly three fourths of the length. The anterior dorsal
and ventral margins are coincident and straight; at the exter¬
nal angles are two very small spines. The dorsal plate is
broadly ovate and slightly truncate posteriorly; the facetting is
of an unusual pattern, the separate facets being bounded by
double lines. The ventral plate is very slightly narrower than
the dorsal and of the same general outline; the surface mark¬
ings are irregular and the lines doubled ; as on the dorsal plate.
The lateral sulci are indistinct. The posterior segment is small
and rounded and projects but little beyond the dorsal plate.
The coxal plates are small and obtusely pointed posteriorly.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 349
The first foot joint is elongate ovate, reaching down over the
pyriform second joint with a lobate extension. The toes are
long and slender, more than one fourth of the total length,
straight and parallel-sided, and end in an indistinct, acute claw.
Total length 145/a; length of dorsal plate 105/a, of ventral
plate 92/t; width of dorsal plate 78/i, of ventral plate 74/t; width
of anterior points 60/t ; length of toes with claw 42/i.
Lecane haliclysta occurs in small numbers in weedy ponds
with soft water ; we have collected it in Oneida and Vilas coun¬
ties, Wisconsin, around Atlantic City, New Jersey, on Mount
Desert Island, Maine, and we have found it also in collections
made by Dr. Birge in Hatchery Bay, South Bass Island, Lake
Erie, during the Great Lakes Investigation by the United States
Pish Commission in 1899.
LECANE ASPASIA Myers
Lecane aspasia Myers, Proc. U. S. Nat. Mus., vol. 52, 1917, p. 47^,
pi. 40, figs. 6-8.
The lorica is very broad, nearly parallel-sided anteriorly and
rounded posteriorly; the dorso-ventral depth of the body is
somewhat greater than usual. The anterior margins of the dor¬
sal and ventral plates are coincident and very slightly convex;
at the external angles are two small spines. The dorsal plate
is very broadly ovate and slightly truncate posteriorly ; its sur¬
face markings are fairly prominent and consist of the usual four
transverse rows of facets, in this instance somewhat larger than
usual. The ventral plate is very broadly ovate and of the same
width as the dorsal plate. The lateral sulci are moderately deep.
The posterior segment is broadly rounded and projects consid¬
erably beyond the dorsal plate. The coxal plates are roughly
triangular. The first foot joint is parallel-sided and rather
small, the second joint short, broad and subrhomboid. The toes
are long and slender, more than one fourth of the total length,
straight and slightly tapering; the claw is long, slender and
acutely pointed.
Total length 132/i ; length of dorsal plate 82/i, of ventral plate
94/t; width of lorica 75/i; width of anterior points 63/i; length
of toes without claw 30/t ; claw Sy.
Lecane aspasia was described from material collected near
Los Angeles, California ; it has not been found elsewhere.
350 Wisconsin Academy of Sciences, Arts and Letters.
LECAXE LUDWIGII (Eckstein)
Plate XXII, figures 5, 6.
Distyla ludwigii Eckstein, Zeitschr. Wiss. Zool., vol. 39, 1883, p.
383, pi. 26, fig. 37. — Blochmann, Mikr. Thierw. Sussw., 1886, p.
107. — Hudson and Gosse, Rotifera, Suppl., 1889, p. 43, pi. 33.
fig. 36. — Ternetz, Rotat. Umg. Easels, 1892, p. 18. — ^Wierzejski,
Rozpr. Akad. Umiej., Wydz. Mat.-Przyr., Krakow, ser. 2, vol. 6,
1893, p. 242. — Skorikov, Trav. Soc. Nat. Kharkow, vol. 30, 1896,
p. 320. — Weber, Rev. Suisse Zool., vol. 5, 1898, p. 602, pi. 22, figs.
9-11. — Jennings, Bull. U. S. Fish Comm., vol. 19 (for 1899),
1900, p. 92, pi. 20, fig. 32. — Voigt, Forschungsber. Biol. Stat.
Plon, vol. 11, 1904, p. 71. — Daday, Zoologica, pt. 44, 1905, p. 108,
pi. 6, fig. 11; pt. 59, 1910, p. 82. — Voronkov, Trudy Gidr. Stants.
Glubokom Oz., vol. 2, 1907, p. 204. — Meissner, Izv. Turkestansk.
Otd. Imp. Russk. Geogr. Obshch., vol. 4, pt. 8, 1908, p. 21, pi. 1,
fig. 2. — De Beauchamp, Arch. Zool. Exper., ser. 4, vol. 10, 1909,
p. 161. — Lie-Pettersen, Bergens Mus. Aarbog (for 1909), 1910,
No. 15, p. 65. — Sachse, Susswasserfauna Deutschlands, pt. 14,
1912, p. 176, figs. 348. — Jakubski, Rozpr. Muz. Dziedus-
zyckich, vol. 1, 1914, p. 32.
Distyla ornata Daday, Math. Termesz. Ertes., vol. 15, 1897, p. 135,
fig. 4; Termesz. Fiizetek, vol. 24, 1901, p. 18, fig. 2.
Distyla oxycauda Stenroos, Acta Soc. Pauna et Flora Fennica, vol.
17, No. 1, 1898, p. 162, pi. 2, figs. 23-25.
Cathypna ludwigii Murray, Journ. Royal Micr. Soc., 1913, p. 352,
pi. 14, fig. 23. — Weber and Montet, Cat. Invert. Suisse, pt. 11,
1918, p. 187.
Lecane ludwigii Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61;
Proc. U. S. Nat. Mus., vol. 47, 1914, p. 537. — Kozar, Zool Anz.,
vol. 44, 1914, p. 420. — Rezvoi, Trudy Borodinsk. Biol. Stants.
Imp. Petrogradsk. Obshch. Estestvoisp., vol. 4, 1916, p. 180. —
Jakubski, Kosmos (Lwow), 1918-1919, p. 28.
The outline of the lorica is oval and moderately broad. The
anterior margins are coincident and slightly concave; at the
external angles are two fairly long, stout spines. The dorsal
plate is oval and truncate posteriorly ; it is very firm and marked
with four transverse rows of prominent, coarse tesselations. The
ventral plate is slightly pyriform in outline and narrower than
the dorsal; the surface markings consist of a few longitudinal
ridges, as shown in the figure. The lateral sulci are very deep.
The posterior segment is roughly semicircular and is produced as
a long, triangular, pointed spine. The coxal plates are large and
rounded posteriorly. The first foot joint is narrow and over¬
laps the trapezoidal second joint as a lobate projection. The toes
Harring Myers — Rotifer Fauna of Wisconsin. — III. 351
are long and slender, one fourth of the total length, parallel-sided
and ending in acute conical points without claws.
Total length 165/a ; length of dorsal plate 120/a, of ventral plate
162/a; width of dorsal plate 77/a, of ventral plate 68/a; width of
anterior points 45/a ; length of toes 45/a.
Lecane ludwigii is very widely distributed, but seems not to
occur in large numbers ; we have collected it in weedy ponds in
Wisconsin, New Jersey, Maine, District of Columbia, Virginia
and Florida and have found it in material from many other local¬
ities in the United States.
Lecane ludwigii, L. ohioensis, L. marsM, L. stokesii and L.
ichthyoura form a very closely related group. The form of
the prolongation of the posterior segment is quite different in
these species and the obvious character upon which they were
established. Some other minor differences deserve notice. The
form of the second foot joint is slightly different in all the spe¬
cies; likewise the outline of the coxal plates. The form and
relative length of the toes also show small, but constant differ¬
ences. The outline of the body, the dorsal plate, the very rigid
lorica and the surface markings of both dorsal and ventral plates
are virtually alike in all the species, and the same is true of
the frontal spines, mth the single exception of L. marsM. We
have tried to reproduce all the similarities and all the differences
as faithfully as possible in the figures ; many of these are so slight
that no word picture would describe them adequately and the
figure must therefore be used as the final criterion.
LECAXE MARSHI Harring
Plate XXII, figures 1, 2.
Lecane marshi Hakring, Proc. U. S. Nat. Mus., vol. 47, 1914, p. 537,
pi. 18, figs. 1-3.
Lecane encodes Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
537, pi. 18, figs. 4-6.
The outline of the lorica is broadly oval or very slightly pyri¬
form. The anterior margins are coincident and somewhat con¬
cave ; at the external angles are two fairly long, slender spines,
usually with the point recurved as a conspicuous hook, semicir¬
cular in lateral view. The dorsal plate is oval and truncate pos¬
teriorly; it is very firm and marked with four transverse rows
of prominent, coarse tesselations. The ventral plate is more
elongate and somewhat narrower than the dorsal; the markings
352 Wisconsin Academy of Sciences, Arts and Letters.
consist of a few longitudinal ridges, as shown in the figure. The
lateral sulci are fairly deep. The posterior segment is rather
large and somewhat angulate, ending in an obtusely pointed,
triangular lobe. The coxal plates are moderately large and
rounded posteriorly. The first foot joint is indistinct and pro¬
jects slightly over the subcircular second joint. The toes are
about one fourth of the total length, straight, slender, parallel¬
sided and end in acute conical points without claws.
Total length 173/a; length of dorsal plate 114/a, of ventral plate
138/a; width of dorsal plate 84/a, of ventral plate 76/a; width of
anterior points 54/a; height of hook 9/a; length of toes 45/a.
Lecane marshi was originally described from material col¬
lected in the Panama Canal Zone, the “bookless’^ form as a dis¬
tinct species. Subsequent study of the collections demonstrated
that they occur together and it is probable that they are only
varieties of a single species. We have also found the hookless
form in collections made by Dr. N’. Gist Gee at Soochow, China ;
they agree in every way with the Panama specimens.
LECANE ICHTHYORA (Anderson and Shephard)
Plate XXII, figures 3, 4.
Distyla ichthyoura Anderson and Shephard, Proc. Royal Soc. Vic¬
toria, new ser., vol. 4, 1892, p. 78, pi. 12, fig. 5.
Cathypna appendiculata Levander, Acta Soc. Fauna et Flora Fen-
nica, vol. 12, No. 3, 1895, p. 50, pi. 3, fig. 30. — ? Daday, Zoologica,
pt. 44, 1905, p. 110 pi. 6, fig. 17.
Lecane ichthyoura Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61. —
Hauer, Mitt. Geogr. Ges. u. Naturhist. Mus. Liibeck, ser. 2, No.
30, 1925, p. 166, fig. 7.
The outline of the lorica is oval and moderately broad. The
anterior dorsal margin is slightly concave, the ventral somewhat
more so ; at the external angles are two stout, triangular spines.
The dorsal plate is oval and truncate posteriorly; it is very
firm and marked with four rows of prominent tesselations. The
ventral plate is more elongate and narrower than the dorsal ; it
is marked with a few longitudinal ridges. The lateral sulci are
very deep. The posterior segment is produced as a peculiar
lobe, rounded behind and with two lateral, triangular points.
The coxal plates are small and rounded posteriorly. The first
foot joint is somewhat indistinct and overlaps the subsquare sec¬
ond joint with a small, rounded projection. The toes are long
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 353
and slender, more than one fourth of the total length, parallel¬
sided and end in acute, conical points without claws.
Total length 140/x ; length of dorsal plate 96^, of ventral plate
120jLt; width of dorsal plate 80/x, of ventral plate 72]U,; width of
anterior points 48/x ; length of toes 40/x.
Lecane ichihyoura is rare ; we have found it in Polk County,
Florida, in small numbers. It is a question whether the form
figured by Hauer should not be considered a variety of L. ohio-
ensis.
LECANE STOKESU (Pell)
Plate XXIII, figures 1-3.
Cathypna stokesii Pell, The Microscope, vol. 10, 1890, p. 144, text
fig. — Murray, Journ. Royal Micr. Soc., 1913, p. 553, pi. 23, fig. 12.
Distyla stokesii Jennings, Bull. Michigan Fish Comm., No. 3, 1894,
p. 24; Bull. U. S. Fish Comm., vol. 29 (for 1899), 1900, p. 92,
pi. 20, fig. 35. — Hempel, Bull. Illinois State Lab. Nat. Hist., vol.
5, 1898, p. 375.
Lecane stokesii Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 62.
The outline of the lorica is oval and somewhat narrower than
usual. The anterior dorsal and ventral margins are very nearly
coincident and slightly concave; at the external angles are two
stout, triangular spines. The dorsal plate is ovate and truncate
posteriorly ; it is very firm and marked with four rows of promi¬
nent tesselations. The ventral plate is more elongate and nar¬
rower than the dorsal ; its markings consist of a few longitudinal
ridges, constant in position, but not very conspicuous. The lat¬
eral sulci are deep. The posterior segment is semicircular and
ends in two long spines, separated by a wide interspace, rounded
anteriorly; the coxal plates are small and obtusely pointed.
The first foot joint is fairly narrow and not very strongly marked,
the second truncate pyriform. The toes are long, straight, slen¬
der and acutely pointed without any claw ; their length is about
one fifth of the total length.
Total length 155/a ; length of dorsal plate 98/*, of ventral plate
130/x; width of dorsal plate 83/t, of ventral plate 77/t; width of
anterior points 50/t; length of toes 48/t.
Lecane stokesii occurs in weedy ponds and is widely distrib¬
uted in the United States, but is seldom found in large numbers.
The variety shown in figure 3 was figured by Murray as the
^‘English form’^; we have collected it among sphagnum at Hy-
23
354 Wisconsin Academy of Sciences, Arts and Letters,
attsville, near Washington, District of Columbia. It differs from
the typical form mainly in the form of the posterior spines,
which are stouter and much shorter ; it is somewhat more robust
and the lorica less oval. It is uncertain whether this should be
considered a variety or an independent species; no transition
forms between the two are known and the typical form does not
appear to be variable. However, until more is knoAvn about the
actual relationship between the members of this group within the
genus, it is probably better to leave it provisionally as a ‘Vari¬
ety’’, even though this term as applied to rotifers is somewhat
ambiguous.
LECANE OHIOENSIS (Herrick)
Plate XXIII, figures 4, 5.
Distyla ohioensis Herrick, Bull. Denison Univ., vol. 1, 1885, p. 54,
fig. 1 (on page preceding index). — Hempel, Bull. Illinois State
Lab. Nat. Hist., vol. 5, 1898, p. 375. — Jennings, Bull. U. S. Fish
Comm., vol 19 (for 1899), 1900, p. 91, 20, fig. 30. — Iroso, Mon.
Zool. Ital., vol. 21, 1910, p. 302; Atti R. 1st. Incorr. Napoli, vol. 64
(for 1912), 1913, p. 467, figs. 9-12. — Jakubski, Rozpr. Wiad. Muz.
Dzieduszyckich, vol. 1, 1914, p. 32, pi. 1, fig. 8.
Cathypna ohioe')isis Turner, Bull. Denison Univ., vol. 6, 1892, p.
61. — Murray, Journ. Royal Micr. Soc., 1913, p. 552, pi. 23, fig. 14.
Lecane ohioensis Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 62. —
Jakubski, Kosmos (Lwow), 1918-1919, p. 28. — Fadeev, Russk.
Gidr. Zhurn., vol. 3, No. 3-5., 1924, p. 4.
The outline of the lorica is a moderately broad oval; the
anterior margins are slightly concave and very nearly coincident,
the ventral a trifle shorter. At the external angles are two fairly
prominent, stout spines. The dorsal plate is broadly oval and
truncate posteriorly ; it is very fl.'rm and marked with four trans¬
verse rows of prominent, coarse tesselations. The ventral plate
is somewhat ovate and narrower than the dorsal plate ; the
markings consist of a few folds or ridges, shown in the figure.
The lateral sulci are deep. The posterior segment is approxi¬
mately semicircular with a short, median, squarely truncate pro¬
jection. The coxal plates are large and obtusely pointed pos¬
teriorly. The first foot joint is indistinct and overlaps the sub¬
square second joint as a lobate projection. The toes are long
and slender, a little less than one fourth of the total length,
parallel-sided and end in acute conical points without claws.
Total length 156/^; length of dorsal plate 91 ix, of ventral plate
Harring (& Myers — Rotifer Fauna of Wisconsin. — 111. 355
116)11 ; width of doral plate TS/a, of ventral plate lOfi; width
of anterior points 48/xj length of toes AOfi.
Lecane ohioensis is common everywhere in weedy ponds and
frequently occurs in large numbers.
LECANE ARCULA Harring
Plate XXIV, figures 1, 2.
Cathypna aculeata Murray, Journ. Royal Micr. Soc., 1913, p. 350,
pi. 14, fig. 28; not Distyla aculeata Jakubski.
Lecane arcula Harring, Proc. U. S. Nat. Mus., vol. 47, 1914, p. 539,
pi. 19, figs. 4-6.
The outline of the lorica is broadly ovate and it is but little
longer than wide. The dorsal and ventral anterior margins are
straight and parallel and nearly coincident; at the external an¬
gles are two moderately large spines, directed slightly upwards
and outwards. The dorsal plate is ovate and rounded poste¬
riorly. The dorsal markings are of the usual pattern and rather
faint. The ventral plate is ovate and as wide as the dorsal ; the
surface markings are indicated in figure 2. The lateral sulci
are moderately deep. The posterior segment is broad and
rounded ; it projects but slightly beyond the dorsal plate. The
coxal plates are indistinct and rounded posteriorly. The first
foot joint is large and elongate pyriform, the second joint also
large and of irregular form. The toes are fairly long and slen¬
der, about one fourth of the total length, straight and parallel¬
sided ; the claw is slender, acutely pointed and recurved.
Total length 96)a; length of dorsal plate 63/a, of ventral plate
68/x, width of dorsal plate 55/.t, of ventral plate 55)U,; width of
anterior spines 47)li, length of toe without claw 18)tx; claw 5/a.
Lecane arcula was first described from the Panama Canal Zone,
and collected by Murray in South America and Australia. It
is common in weedy ponds everywhere in the United States.
Distyla aculeata Jakubski is, according to the description and
figure, more elongate, strongly compressed dorso-ventrally and
has much longer anterior spines.
LECANE FLEX-ILIS (Gosse)
Plate XXIV, figures 3, 4.
Distyla flexilis Gosse, in Hudson and Gosse, Rotifera, 1886, vol. 2,
p. 97, pi. 24, fig. 7. — Glasscott, Proc. Royal Dublin Soc., new ser.,
vol. 8, 1893, p. 72. — Weber, Rev. Suisse Zool., vol. 5, 1898, p. 599,
356 Wisconsin Academy of Sciences, Arts and Letters.
pi. 22, fig. 8. — Voronkov, Trudy Gidr. Stants. Glubokom Oz., vol. 2,
1907, p. 111. — ^VoN Hofsten, Arkiv Zool., Stockholm, vol. 6, No. 1,
1909, p. 58, fig. 12. — Lucks, Rotatorienfauna Westpreussens,
1912, p. 110. — Sachse, Susswasserfauna Deutschlands, pt. 14,
1912, p. 176, figs. 346, 347. — Montet, Rev. Suisse Zool., vol. 23,
1915, p. 336. — Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich, vol.
1, 1914, p. 32. — Oparina-Kharitonova, Izv. Biol. Nauchno-Issl.
Inst. Permsk. Univ., vol. 3, 1925, p. 440.
Cathypna flexilis Stenroos, Acta Soc. Fauna et Flora Pennica, vol.
17, No. 1, 1898, p. 159, pi. 2, fig. 19. — Murray, Journ. Royal Micr.
Soc., 1913, p. 351, pi. 14, fig. 27. — Weber and Montet, Cat. In¬
vert. Suisse, pt. 11, 1918, p. 185. — VoN Hofsten, Naturw. Unters.
Sarekgeb., vol. 4, 1923, p. 861.
Cathypna brevis Murray, Journ. Royal Micr. Soc., 1913, p. 555,
pi. 22, fig. 8. — Olofsson, Zool. Bidr. Uppsala, vol. 6, 1918, p. 592,
fig. 52.
Lecane flexilis Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 61; Proc.
U. S. Nat. Mus., vol. 47, 1914, p. 538, pi. 19, figs. 1-3; Rep. Cana¬
dian Arctic Exp. 1913-18, vol. 8, pt. E, 1921, p. 9. — Jakubski,
Kosmos (Lwow), 1918-1919, p. 28.
The outline of the lorica is subcircular and the body is strong¬
ly gibbous. The anterior dorsal and ventral margins are
coincident and distinctly convex ; at the external angles are two
short, very stout, slightly incurved spines. The dorsal plate is
subcircular and rounded posteriorly; the surface markings con¬
sist of the usual four transverse rows of facets, but the first row
is somewhat irregular and the others have rather more than the
usual number of facets. The lorica is quite firm and the facets
prominent. The ventral plate is considerably narrower than the
dorsal and its edges ill-defined; the markings are less distinct
than on the dorsal plate. The lateral sulci are very indistinct.
The posterior segment is small and rounded and projects but
little beyond the dorsal plate ; the coxal plates are semicircular.
The foot does not project beyond the end of the lorica; the first
joint is elongate oval, the second large and subrhomboid. The
toes are fairly long and slender, about one fourth of the total
length, straight and parallel-sided for the greater part of their
length, thence tapering slightly to the small, acute, recurved
claw, which has dorsally a small basal spicule.
Total length 96/x; length of dorsal plate 72/a, of ventral plate
76/x; width of dorsal plate 66/x, of ventral plate 50 fi; width of
anterior points 55/a ; length of toes without claw 19/a ; claw 4/a.
Lecane flexilis is common in weedy ponds everywhere in the
United States; Mr. David Bryce and the late Mr. C. F. Rousse-
Harring & Myers — Rotifer Fauna of Wisconsin. — 111. 357
let have sent us specimens from England. Our identification of
this species with Grosse’s animal is based on the form of the
animal when swimming ; its length is then nearly twice as great,
the body is slender and flattened dorso-ventrally and the dorsal
markings much more prominent.
LECANE INTRASIMUATA (Olofsson)
Plate XXIV, figures 5, 6.
Cathy pna intrasinuata Olofsson, Zool. Bidr. Uppsala, vol. 5, 1917,
p. 281, fig. 11. — ^Idelson, Trudy Plovuch. Morsk. Nauchn. Inst.,
Moskva, pt. 12, 1925, p. 90.
Lecane epkestra Harking, Rep. Canadian Arctic Exp. 1913-18, vol,
8, pt. E, 1921, p. 7, pi. 3, figs. 1, 2.— Hauer, Zool. Anz., vol. 61,
1924, p. 149.
The outline of the lorica is broadly oval, the greatest width
about three fifths of the length. The anterior margins of the
dorsal and ventral plates are coincident and very slightly con¬
vex; at the external angles are two small, robust spines. The
dorsal plate is subcircular, rounded posteriorly and without sur-
face markings. The ventral plate is considerably narrower than
the dorsal, nearly parallel-sided and rounded posteriorly; the
margins are ill-defined and no lateral sulci are present. The sur¬
face markings are not prominent, but quite constant. The
posterior segment is rounded and projects but little beyond
the dorsal plate. The coxal plates are small and obtusely tri¬
angular. The first foot joint is long, wide anteriorly and sharp¬
ly constricted immediately in front of the second joint; this is
large and pyriform, projecting about half its length beyond the
posterior segment. The toes are long and slender, about one
fourth of the total length, parallel-sided and very faintly in¬
curved; the tip is excavate on the inner margin, forming a
pseudo-claw.
Total length 150>i ; length of dorsal plate 90/>i, of ventral plate
95ju; width of dorsal plate ISy, of ventral plate 62/^; width of
anterior points 54ft ; length of toes without claw 34ft, claw 6ft.
Lecane intrasinuata was described by Olofsson from a few
specimens collected in a pond near Alexandrovsk, on the Mur-
man Coast, Eussian Lapland, We believe this to be identical
with L. epkestra, although the illustration accompanying the
original' description is lacking in detail. L. intrasinuata is
common in soft, acid water ponds in Oneida and Vilas Counties,
358 Wisconsin Academy of Sciences, Arts and Letters.
Wisconsin, on Mount Desert Island, Maine, and around Atlantic
City, New Jersey. It has recently been found by Hauer in the
Schwarzwald, where it is not rare, and by Idelson on Nova j a
Zemlja.
IiECA]%E CL.IMACOIS Marring and Myers, new species
Plate XXV, figures 1, 2.
The outline of the lorica is broadly ovate ; the width is about
two thirds of the length. The anterior dorsal margin is straight
and the ventral margin slightly concave ; at the external angles
are two small, acute spines. The dorsal plate is broadly ovate
and slightly truncate posteriorly; the facetting is very regular
and of the usual pattern, but very faint and distinctly visible
only by turning the animal partly on its side. The ventral plate
is somewhat elongate oval; the surface markings are fairly
simple and not very distinct. The lateral sulci are deep. The
posterior segment is fairly large and nearly semicircular. The
coxal plates are small, broad at the base and acuminate
posteriorly. The first foot joint is long, parallel-sided and
pointed posteriorly; the second joint is robust and somewhat
pyriform with two small, lateral knobs; it projects about two
thirds of its length beyond the lorica. The toes are fairly long
and robust, a little less than one fourth of the total length,
parallel-sided and straight; the long, conical claw is sharply in¬
dented at its base.
Total length 95/x; length of dorsal plate 64/x, of ventral plate
lOfjL] width of dorsal plate 51/a, of ventral plate 45/a; width of
anterior points 34/a ; length of toes without claw 15/a ; claw 7/a.
Lecane climacois occurs in wet sphagnum or sphagnum bogs;
we have collected it at Glenburnie, near Baltimore, Maryland;
in Polk County, Florida, and in ‘‘Annecta Pool”, a very acid,
roadside ditch not far from Atlantic City, New Jersey. Only
at the last named location has it been found in large numbers.
tiBCANE VBRECUNDA Marring: and Myers, new species
Plate XXV, figures 3, 4.
The outline of the lorica is broadly ovate and somewhat point¬
ed posteriorly. The anterior margin of the ventral plate is
nearly straight; the dorsal margin is very slightly convex and
projects a little beyond the ventral plate. At the external angles
are two small, slightly incurved frontal spines. The dorsal plate
Earring dj Myers — Rotifer Fauna of Wisconsin, — III, 359
is ovate, somewhat pointed posteriorly and squarely truncate at
the extreme end. The dorsal facetting is not very prominent
and follows the prevailing pattern. The ventral plate is nearly
parallel-sided anteriorly and rounded posteriorly; the margins
are wavy and ill-defined, as are the lateral sulci; the surface
markings are very simple. The posterior segment is small and
rounded and projects but very little beyond the dorsal plate.
The coxal plates are small and very obtusely pointed poste¬
riorly. The first foot joint is ovate and the second joint large
and rhomboid in outline, projecting slightly beyond the pos¬
terior segment. The toes are fairly long and slender, about
one fourth of the total length, straight and very slightly taper¬
ing; the claw is moderately long, very slender, acutely pointed
and spine-like.
Total length 105^; length of dorsal plate 72/a, of ventral plate
76/t; width of dorsal plate 60/a, of ventral plate 50/x; width of
anterior points 45/a ; length of toe without claw 22/a ; claw 6/a.
Lecane verecunda was collected in small numbers in weedy
ponds at the State Fish Hatchery, near Madison, Wisconsin, and
in Witch Hole, Mount Desert Island, Maine. Its nearest relative
is probably L. arculay from which it differs in the more ovate
outline of the dorsal plate, slightly truncate posteriorly, the
smaller posterior segment and the relatively narrower ventral
plate.
L.ECAXE MYLACRIS Harring- and Myers, new species
Plate XXV, figures 5, 6.
The lorica is very firm and very broadly ovate; its width is
three fourths of the length. The anterior margins are nearly
coincident, the ventral almost straight and the dorsal slightly
convex; at the external angles are two minute frontal spines.
The dorsal plate is very broadly ovate and projects beyond the
ventral plate both laterally and posteriorly. The ventral plate
is nearly parallel-sided for about two thirds of its length and
rounded posteriorly; surface markings are found only on the
ventral plate and are limited to a few fairly distinct folds. The
lateral sulci are deep. The coxal plates are very small and
pointed posteriorly. The first foot joint is oval, somewhat in¬
distinct and lobate posteriorly; the second joint is very large,
robust and subsquare, projecting about one third of its length
beyond the lorica. The toes are moderately long and slender,
360 Wisconsin Academy of Sciences, Arts and Letters.
about one fourth of the total length, straight on their inner
edges and very faintly sigmoid externally, terminating in a
small, pointed claw; they are inserted near mid-length of the
second foot joint instead of on the posterior edge.
Total length 175/>t; length of dorsal plate 145jit, of ventral
plate 130/x; width of dorsal plate 108/>t, of ventral plate 82/a;
width of anterior spines 70/a; length of toes without claw 32/a;
claw 7/a.
Lecane mylacris was collected in small numbers in Upper
Mill Pond, Mount Desert Island, Maine, and is common in weedy
areas of Lake Hartridge, Polk County, Florida.
liECANE GL.YPTA Harring and Myers, new species
Plate XXVI, figures 1, 2.
The outline of the lorica is somewhat elongate ovate, nearly
parallel-sided anteriorly and rounded posteriorly. The anterior
margins are not coincident ; the dorsal margin is convex and the
ventral slightly concave. At the external angles are two very
small frontal spines. The entire lorica is quite flexible. The
dorsal plate is elongate ovate and slightly truncate posteriorly ;
the facetting is of an aberrant pattern, not found in any other
species. The ventral plate is elongate ovate, rounded posteriorly
and as wide as the dorsal plate; the surface markings are,
as shown, unusually intricate. The lateral sulci are indistinct.
The posterior segment is relatively small, nearly semicircular
and projects slightly beyond the dorsal plate. The coxal plates
are small and rounded posteriorly. The first foot joint is long
and narrow, the second joint broadly pyriform and does not
quite reach the posterior end of the lorica. The toes are fairly
long and slender, about one fourth of the total length, nearly
parallel-sided and with a very slight, sigmoid curvature; the
claw is small, acute and slightly outcurved, with a distinct basal
spicule.
Total length 110/a ; length of dorsal plate 80/a, of ventral plate
86/^ ; width of lorica 50/a ; width of anterior points 42/a ; length
of toes without claw 22/a ; claw 5/a.
Lecane glypta was collected in weedy ponds on the Pine
Valley golf course, near Clementon, New Jersey. This is on the
border line between the acid and alkaline waters of the state.
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 361
LECANE TABIDA Harriiig and Myers, new species
Plate XXVI, figures 3, 4.
The lorica is broadly ovate and nearly parallel-sided anterior¬
ly; its width is about two thirds of the length. The anterior
margins of the dorsal and ventral plates are coincident, very
slightly convex and on account of the flexibility of the entire
lorica, somewhat wavy. The dorsal plate is broadly ovate and
its edges ill-defined; it is rounded posteriorly. The facetting
is rather faint and the number of facets considerably greater
than in the usual pattern. The ventral plate is nearly parallel¬
sided for the greater part of its length and rounded posteriorly ;
its markings are faint and depart somewhat from the usual de¬
sign. The lateral sulci are rather shallow* and ill-deflned. The pos¬
terior segment is large and nearly semicircular ; the coxal plates
are rather small and very obtusely pointed. The flrst foot joint
is narrow and elongate, lobate posteriorly, the second joint sub¬
square, somewhat narrower anteriorly, and projecting nearly
half its length beyond the lorica. The toes are rather short and
robust, less than one fourth of the total length, straight on their
inner edges, parallel-sided for nearly three fourths of their
length and terminate in gradually acuminating points.
Total length 130/x ; length of dorsal plate 94/>t, of ventral plate
98fi; width of dorsal plate 65/a, of ventral plate 60/a; width of
anterior points 55/a ; length of toes 30/a.
Lecane tahida was collected in small numbers among
Fontinalis in quiet pools along Trout Brook, on Mount Desert
Island, Maine. It has not been found elsewhere.
IiECAlVE IXFUIiA Barring and Myers, new species
Plate XXVI, figures 5, 6.
The outline of the lorica is broadly oval; its width is about
two thirds of the length. The anterior margins are nearly
coincident and slightly concave; at the external angles are two
very stout, slightly incurved spines. The dorsal plate is broadly
oval and rounded posteriorly. The facetting is of a remarkably
intricate and very irregular pattern ; each facet is outlined by a
double row of minute, raised dots or beads. On the posterior
half of the dorsal plate there is near the margins a pair of in¬
cipient dorsal sulci”. The ventral plate is considerably
narrower than the dorsal, nearly parallel-sided and its margins
somewhat wavy; the surface markings are of a fairly simple
pattern. The lateral sulci are moderately deep. The posterior
362 ^Yisconsin Academy of Sciences, Arts and Letters,
segment is large and very obtusely pointed ; it projects somewhat
beyond the dorsal plate. The coxal plates are obtusely tri¬
angular and their points some distance from the posterior end
of the lorica. The first foot joint is rather indistinct and pointed
posteriorly, the second joint large and pyriform and does not
quite reach the margin of the posterior segment. The toes are
long and slender, nearly one third of the total length, straight
and parallel-sided ; the claw is short and very slightly incurved.
Total length 160/x; length of dorsal plate 105^t, of ventral
plate 120/a; width of dorsal plate 80/a, of ventral plate 65/a;
width of anterior spines 45/a, length of toes without claw 40/a;
claw 6/a.
Lecane infula was collected in Lower Breakneck Pond, on
Mount Desert Island, Maine, in small numbers; it is not known
from any other localities.
liECAXE SATYRUS Harriiig^ and Myers, new species
Plate XXVII, figures 3, 4.
The outline of the lorica is broadly ovate; its width is about
three fourths of the length. The median portion of the anterior
dorsal margin forms a sort of guard for the opening of the
lorica; it occupies about half the width of the dorsal plate and
is limited externally by two small, incurved spines; outside of
these spines there is on each side a distinct, small sinus. As a
result of the general deformation of the dorsal plate taking place
when the head is retracted, this lobe bends down over the
anterior edge of the ventral plate, thus affording additional pro¬
tection for the delicate corona. At the external angles of the
lorica are two very large, antler-like frontal spines; these are
normally at right angles to the ventral plate and, when thus
turned up, serve to protect the corona when the animal is swim¬
ming; in retraction they turn outwards so that they are sub¬
stantially in the plane of the ventral plate. Although very
irregular in outline and slightly variable in details of the
denticulation, they may be described as rather slender, acutely
pointed spines with a sigmoid curvature, directed forward and
outwards; on the external edge there is near the base a single
large tooth, and on the internal edge four or five teeth, varying
slightly in size and location in different individuals, but limited
to the basal half of the spine; the terminal portion is always
slender, acutely pointed and slightly incurved. The anterior
Earring (& Myers — Rotifer Fauna of Wisconsin. — 111. 363
margin of the ventral plate is slightly convex. The dorsal plate
is broadly ovate and slightly truncate posteriorly. The surface
ornamentation of both dorsal and ventral plate is unique and
very intricate and must be studied from the figures ; it consists
of a system of elevated ridges, rounded at the top and with very
closely spaced, hemispherical beads on the sloping sides. Similar
‘ ^ secondary markings ’ ’ are found in L. signifera, but the general
pattern is quite different. The ventral plate is broadly ovate
and somewhat angular posteriorly, slightly truncate at the ex¬
treme end. The posterior segment is fairly large and projects
considerably beyond the dorsal plate. The coxal plates are small
and narrowly oval. The first foot joint is narrow, elongate and
pointed posteriorly, second joint large and pyriform, projecting
about half its length beyond the lorica. The toes are long and
slender, about one fourth of the total length, straight and
parallel-sided, terminating in acutely pointed claws with a small
basal spicule.
Total length 175//,; length of dorsal plate without anterior
spines 95/^, of ventral plate 105/>t; width of dorsal plate 78//, of
ventral plate 72//; width of anterior margin at base of spines
65/ji; width over spines 75//; length of spines 25//; length of toe
without claw 34//; claw 7//.
Lecane satyrus is usually, if not exclusively, found in wet
sphagnum. We have collected it at Glenburnie, near Baltimore,
Maryland; on Mount Desert Island, Maine; in Polk County,
Florida ; around Atlantic City, New Jersey, and in Vilas County,
Wisconsin, where it was abundant in a small pool near BenCs
resort on Mamie Lake ; it is usually found only in small numbers.
It is perhaps the most remarkable species of the genus; the
rotatable, protective anterior spines are found in Lecane
mucronataf but in a much simpler form, and the surface mark¬
ings are far more elaborate than in any other species.
I.ECANE HASTATA (Murray)
Plate XXVIII, figures 5, 6.
Cathypna hastata Murray, Journ. Ro3^al Micr. Soc., 1913, p. 348,
pi. 14, fig. 25.
The lorica is very broadly ovate and somewhat flexible. The
anterior dorsal margin is very slightly convex and the ventral
margin straight; at the external angles are two small anterior
spines. The dorsal plate is broadly ovate and slightly truncate
364 Wisconsin Academy of Sciences, Arts and Letters.
posteriorly ; it is much smaller than the ventral plate. The ven¬
tral plate is very broadly ovate; its surface markings consist of
a transverse fold and two faint, longitudinal ridges. Lateral
sulci are not present; the section of the lorica connecting the
dorsal and ventral plates is very slightly concave. The posterior
segment is relatively small and rounded; the coxal plates are
apparently absent. The first foot joint is large and indistinct,
the second subsquare, projecting beyond the posterior end of the
lorica. The toes are long and slender, more than one fourth of
the total length, straight, parallel-sided with a bulbous enlarge¬
ment near the posterior end, and terminating in a long, slender,
acutely pointed claw.
Total length 150/a ; length of dorsal plate 95/a, of ventral plate
110/a; width of dorsal plate 75/a, of ventral plate 90/a; width of
anterior points 70/a ; length of toe without claw 25/a ; claw 15/a.
Lecane Jiastata was described by Murray from a brackish pond
at Rio de Janeiro, Brazil. We have found this species in abund¬
ance in brackish ditches at Port Republic, New Jersey, in per¬
fectly fresh water at Barrows, Virginia, near Washington, and
in collections from Baton Rouge, Louisiana. In the Port Re¬
public material the dorsal plate is smooth; the specimens from
Barrows show a very faint tesselation towards the anterior edge
of the dorsal plate. L. Jiastata is probably related to L. crepida;
they agree in the peculiar structure of the lorica and the poste¬
rior segment and foot joints.
LECAXE CREPIDA Harring
Plate XXVIII, figures 1, 2.
Distyla gissensis Jennings, Bull. U. S. Fish Comm., vol. 19 (for
1899), 1900, p. 91, pi. 20, figs. 33, 34; not Distyla gissensis Eck¬
stein.
Lecane crepida Harking, Proc. U. S. Nat. Mus,, vol. 47, 1914, p. 533,
pi. 22, figs. 4-7.
The lorica is very flexible, parallel-sided for one half of its
length and tapers rapidly to the foot ; it is strongly gibbous pos¬
teriorly. The anterior dorsal margin is slightly convex and
the ventral a trifle concave ; at the external angles are two very
stout, curved anterior spines. The dorsal plate is strongly con¬
vex and much smaller than the ventral; the edges are rather
indistinct and the surface markings limited to three pairs of
divergent, wavy ridges, beginning near the anterior margin.
Hairing Myers — Rotifer Fauna of Wisconsin. — III. 365
The ventral plate is moderately convex and has two interrupted
series of ridges extending the greater part of its length; there
is a well marked transverse fold immediately in front of the
foot. Lateral sulci are not present; the section of the lorica
connecting the dorsal and ventral plates is very slightly con¬
cave. The posterior segment merges without definite anterior
limit with the body; ventrally it has a large, circular opening
for the foot. The coxal plates are very indistinct. The first
foot joint is large and bulbous, the second joint is subsquare
and projects with more than half its length beyond the lorica.
The toes are very long and slender, more than one fourth of the
'total length, straight and slightly tapering, terminating in a
long, slender, acutely pointed claw, sharply constricted at the
base.
Total length 135;^; length of dorsal plate 75/>t, of ventral plate
90ju,; width of dorsal plate 45/>t, of ventral plate 60/x; width of
anterior spines 52/x ; length of toe without claw 30/>t ; claw 9/^.
Lecane crepida was described from material collected in the
Panama Canal Zone, where it is common ; we have since found it
at Oconomowoc, Wisconsin, and on Mount Desert Island, Maine.
Jennings records it from South Bass Island, Lake Erie. We are
unable to see any resemblance between this species and Eck¬
stein’s Distyla gissensis; this is said to have two anterior spines
on one side and one on the other and, apart from this evident
error, bears little resemblance to L. crepida. It was evidently
one of the smaller Lecanes, but the original description is so
lacking in detail, and so many species have later been listed un¬
der this name that it is now apparently hopeless to attempt to
identify it.
LECANE SAGULA Harring and Myers, new speeies
Plate XXVIII, figures 3, 4.
The outline of the lorica is very broadly ovate; the width is
nearly equal to the length. The anterior margins are coinci¬
dent and form a somewhat wavy, straight line ; frontal spines are
not present. The entire lorica is semi-flexible and its margins
somewhat indefinite ; the dorso-ventral depth of the body is un¬
usually great. The dorsal plate is very broadly ovate and
rounded posteriorly; the facetting is rather faint and of an un¬
usual pattern. The ventral plate is nearly parallel-sided ante-
366 Wisconsin Academy of Sciences, Arts and Letters,
riorly and very obtusely triangular posteriorly ; it is a little nar¬
rower than the dorsal plate. The ventral surface markings are
very intricate and must be studied from the figure. The lateral
sulci are indistinct. The posterior segment is rather small and
broadly rounded; its entire length projects beyond the dorsal
plate. The coxal plates are large and rounded posteriorly. The
first foot joint is short and pyriform with a bluntly pointed pos¬
terior lobe ; the second joint is huge, nearly circular and projects
about half its length beyond the lorica. The toes are rather
short and straight, a little less than one fourth of the total
length, parallel-sided for the greater part of their length and
slightly reduced at the base of the small, acute claw.
Total length 95ja; length of dorsal plate 48/a, of ventral plate
60/a; width of dorsal plate 48/a, of ventral plate 45/a; width of
anterior margin 38/a ; length of toes without claw 19/a ; claw 4/a.
Lecane sagula was collected in small numbers in a weedy pond
at Minocqua, Wisconsin, and in Witch Hole, Mount Desert Isl¬
and, Maine.
LECANE FORMOSA Harrins* ami Myers, uew species
Plate XXIX, figures 1, 2.
The lorica is very broadly ovate, almost parallel-sided an¬
teriorly and rounded posteriorly ; it is moderately fiexible. The
anterior margins of the dorsal and ventral plates are coincident
and straight; no anterior spines are present. The dorsal plate
is very broadly ovate and slightly truncate posteriorly; the an¬
terior margin is considerably narrower than the ventral plate
and surface markings are entirely absent. The ventral plate is
nearly parallel-sided anteriorly, somewhat triangular posterior¬
ly and broadly rounded at the extreme end ; it is as wide as the
dorsal plate and its surface markings are limited to a trans¬
verse fold in front of the foot and two longitudinal lines diverg¬
ing towards the anterior margin. The lateral sulci are fairly
deep. The posterior segment is broadly rounded and projects
slightly beyond the dorsal plate ; the coxal plates are very indis¬
tinct and pointed posteriorly. The first foot joint is widest
anteriorly and lobate posteriorly; the second joint robust and
broadly pyriform. The toes are long and slender; about one
third of the total length, straight and slightly tapering ; the claw
is long, very slender and acutely pointed.
Earring (& Myers — Rotifer Fauna of Wisconsin. — III. 367
Total length 110/x ; length of dorsal plate 74/a, of ventral plate
80/a width of lorica 68/a; width of anterior margin 60/a; length
of toes without claw 25/a; claw 7/a.
Lecane formosa has been found only in Lac Vieux Desert, Vilas
County, Wisconsin, in shallow, weedy indentations near the
outlet.
LECANE AEGAENA Harrin^
Plate XXVII, figures 1, 2.
Lecane aeganea Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
542, pi. 21, figs. 1-3. — Fadeev, Russk. Gidr. Zhur., vol. 3, No,
3-5, 1924, p. 5, fig. 1.
The outline of the lorica is broadly ovate and the dorso-ventral
depth less than usual. The anterior dorsal margin is slightly
convex and the ventral margin straight ; they do not meet when
the head is retracted, but leave the lorica partly open; no an¬
terior spines are present. The dorsal plate is oval and rounded
posteriorly; the facetting is fairly regular and not very con¬
spicuous. The ventral plate is longer and wider than the dorsal ;
its markings are prominent, especially on the posterior half.
The lateral sulci are not very deep and the entire lorica is quite
flexible. The posterior segment is small and rounded; it pro¬
jects somewhat beyond the dorsal plate. The coxal plates are
very small and rounded posteriorly. The first foot joint is very
long, but almost obliterated near the middle; the second joint
is large and subrhomboid. The toes are long and slender, about
one third of the total length, straight and very slightly conical,
terminating in a long, slender, acute claw with a small basal
spicule.
Total length 110/a ; length of dorsal plate 70/a, of ventral plate
76/a; width of dorsal plate 56/a, of ventral plate 60/a; width of
anterior margin 50/a ; length of toes without claw 24/a ; claw 10/a.
Lecane aeganea was described from material collected in the
Panama Canal Zone; we have since found it around Atlantic
City, New Jersey, but never in large numbers. Fadeev reports it
from the gouvernments Kharkov and Tamboy, European Russia.
368 Wisconsin Academy of Sciences, Arts and Letters.
LECAXE VEJVUSTA Harring and Myers, new species
Plate XXVII, figures 5, 6.
The outline of the lorica is very broadly ovate; its width is
about seven eighths of the length. The anterior dorsal margin is
very slightly convex and the ventral nearly straight; no ante¬
rior spines are present. The dorsal plate is broadly ovate and
rounded posteriorly; the facetting follows in general the usual
pattern and is not especially prominent. The ventral plate is
almost parallel-sided anteriorly and broadly rounded poste¬
riorly ; the surface markings are fairly complex. The lateral sulci
are not very deep. The posterior segment is broadly rounded
and projects only slightly beyond the dorsal plate, but it is un¬
usually distinct. The coxal plates are almost as large as the
posterior segment and of the same general form. The first foot
joint is pyriform and lobate posteriorly, the second joint large
and subsquare. The toes are long and slender, nearly one-third
of the total length, very slightly incurved, parallel-sided and
terminating in moderately acute points, but not forming a claw.
Total length ISOfx ; length of dorsal plate 86ju, of ventral plate
84/^; width of dorsal plate 75/x, of ventral plate 70/x; width of
anterior margin 62/^; length of toes 42ix.
Lecane venusta has been found only in small numbers in Ott-
man Lake, near Waupaca, Wisconsin.
LECANE CANDIDA Harring and Myers, new speeies
Plate XIV, figures 1, 2.
The outline of the lorica is broadly ovate ; its Avidth is about
two thirds of the length. The anterior margins are coincident
and very slightly convex; no anterior spines are present. The
dorsal plate is broadly ovate and very obtusely pointed poste¬
riorly ; it is without markings of any kind. The ventral plate is
nearly parallel-sided anteriorly and rounded posteriorly; its
markings are confined to a few folds on the posterior half. The
lateral sulci are fairly deep. The posterior segment is large and
rounded; it projects somewhat beyond the dorsal plate. The
first foot joint is somewhat indistinct and lobate posteriorly, the
second joint nearly circular. The toes are long, about one third
of the total length, straight and slender, terminating in an
acutely pointed claw, very slightly outcurved at the tip, and
with distinct basal spicule.
Harring d' Myers — Rotifer Fauna of Wisconsin. —Ill, 369
Total length lOOju, ; length of dorsal plate 65/a. of ventral plate
72/a; width of dorsal plate 56/A5 of ventral plate 48/a; width of
anterior margin 48/a; length of toes without claw 22/a, claw 7/a.
Lecane Candida was collected in small numbers among sphag¬
num growing on the margins of Loon Lake, near Eagle Eiver,
Vilas County, Wisconsin. It has not been found elsewhere.
LECANE PUSILLA Harring
Plate XXX, figures 1, 2.
Lecane pusilla Harking, Proc, U. S. Nat. Mus., vol. 47, 1914, p. 541,
pi, 20, figs. 4-6.— -Fadeev, Rnssk. Gidr. Zhurn., vol, 3, No. 3-5,
1924, p. 6.
The lorica is very broadly ovate and the dorso-ventral depth
of the body is greater than usual. The anterior margins are
straight and the dorsal plate projects somewhat beyond the ven¬
tral, so that the lorica is not closed when the head is completely
retracted. No anterior spines are present. The dorsal plate is
subcircular and slightly truncate posteriorly. The facetting is
fairly distinct and does not differ widely from the usual pattern.
The ventral plate is parallel-sided in its anterior half and very
obtusely triangular posteriorly; the surface markings are con¬
spicuous only on the median portion of the pattern. The lateral
sulci are deep. The rounded posterior segment projects consid¬
erably beyond the dorsal plate ; the coxal plates are rather small
and rounded posteriorly. The first foot joint is indistinct, the
second large and nearly circular. The toes are long and slender,
more than one fourth of the entire length, straight and slightly
tapering, terminating in a long, recurved, very slender and acute
claw.
Total length 75/a; length of dorsal plate 54/a, of ventral plate
60/x; width of dorsal plate 52/a, of ventral plate 45/a; width of
anterior margin 50/a; length of toes without claw 15/a; claw 5/a.
Lecane pusilla was described from material collected in the
Panama Canal Zone, where it is locally common; Fadeev re¬
ports finding it at Kharkov, Russia.
LECANE ASTHENA Harming and Myers, new species
Plate XXX, figures 3, 4.
The outline of the lorica is very broadly ovate and the dorso-
ventral depth of the body is somewhat greater than usual. The
24
870 Wisconsin Academy of Sciences, Arts and Letters.
anterior margins of the dorsal and ventral plates are coincident
and slightly convex; no anterior spines are present. The dor¬
sal plate is approximately circular and very slightly truncate
posteriorly; the facetting follows in general the usual pattern.
The ventral plate is very slightly narrower than the dorsal, al¬
most parallel-sided anteriorly and semicircular posteriorly; the
surface markings are somewhat more intricate than is usually
the case. The lateral sulci are fairly deep. The posterior seg¬
ment is short and very broadly rounded; it projects somewhat
beyond the dorsal plate. The coxal plates are obtusely truncate
at their posterior, free ends. The first foot joint is narrow and
nearly parallel-sided, the second joint is nearly hemispherical,
very short and very broad. The toes are long and slender, more
than one fourth of the entire length, straight and slightly taper¬
ing, terminating in a long, slender, acute claw of such form that
its external edge is a continuation of the external edge of the
toe and a slight excavation formed on the inner edge.
Total length 80y; length of dorsal plate 54//,, of ventral plate
60/i; width of dorsal plate 54/x, of ventral plate 50ju,; width of
anterior margin 48/i,; length of toes without claw 17/>t; claw 6/i.
Lecane asthena was collected in a shallow, dead bay of Lake
Kawaguesauga at Minocqua, Wisconsin; only a few specimens
were present, and it has not been found elsewhere.
LECANE SUBTILIS Harring and Myers, new species
Plate XXX, figures 5, 6.
The anterior portion of the lorica is roughly parallel-sided
and the posterior portion sub-triangular and rounded at the apex.
The anterior dorsal and ventral margins are coincident and very
slightly convex; no anterior spines are present. The dorsal
plate is parallel-sided anteriorly and obtusely rounded poste¬
riorly; its surface markings are of a very intricate and unusual
pattern. The ventral plate is as wide as the dorsal, parallel¬
sided anteriorly and obtusely triangular posteriorly; its sur¬
face markings are even more unusual than the dorsal and have
no counterpart in the genus. The lateral sulci are evanescent
and the entire lorica is quite flexible. The posterior segment is
small and nearly semicircular. The coxal plates are rudimen¬
tary. The first foot joint is very indistinct and its outlines con¬
fluent with the central portion of the ventral markings ; the sec-
H awing db Myers — Rotifer Fauna of Wisconsin. — III. 371
ond joint is subsquare and projects slightly beyond the lorica.
The toes are very long and slender, about one third of the total
length, straight, very slightly conical and terminate in a long,
slender, acutely pointed claw, sharply constricted at the base.
Total length 110/^; length of dorsal plate 70^, of ventral plate
75^c; width of lorica 60/>t; width of anterior margin 56/a; length
of toe without claw 24ju, ; claw 8/a.
Lecane subtilis was collected in a very shallow bay of Arbor
Vitae Lake, near Minocqua, Wisconsin; it has not been found
elsewhere.
nECAXE ELEUAXS Harring*
Plate XV, figures 1, 2.
Lecane elegans PIarring, Proc. U. S. Nat. Mus., vol. 47, 1914,
p. 544, pi. 23, figs. 1, 2.
The body is elongate, slender and parallel-sided. The dorsal
plate is very flexible, the ventral much less so. The anterior
margin of the ventral plate is nearly straight and of fairly con¬
stant form ; the dorsal margin is irregularly puckered by the in¬
version of the head. The dorsal plate is strongly convex, nearly
semicircular in cross section. The lateral sulci are barely in¬
dicated on the posterior third of the body, the ventral plate may
be said to be joined directly to the dorsal; it has prominent
longitudinal ridges. The posterior segment of the body is very
large and unusually prominent; it may be considered as begin¬
ning near the middle of the ventral plate and its outline is ovate,
broadest posteriorly. Coxal plates are not present. The first
foot joint is extremely long, tapering posteriorly to half its
anterior width; the second joint is large and subsquare and pro¬
jects with its entire length beyond the lorica. The toes are very
long and slender, about one third of the total length, and slight¬
ly recurved in the posterior third; the claw is nearly half as
long as the toe, outcurved and slightly recurved, with a con¬
spicuous, laterally directed basal spine.
Total length 170/a, length of lorica 108/a, width 45/a; toe with¬
out claw 36/t ; claw 15/a.
Lecane elegans was described from material collected in the
Panama Canal Zone ; only a few specimens were collected and it
has not been found elsewhere.
372 Wisconsin Academy of Sciences^ Arts and Letters.
1.ECAXE EliOXGATA Marring^ and Myers, new species
Plate XXXI, figures 1, 2.
The outline of the lorica is elongate reversed-ovate ; its width
is only three fifths of the length. The anterior margin of the
dorsal and ventral plates is slightly convex and the lorica is
widely open in front when the head is fully retracted; no
anterior spines are present. The dorsal plate is elongate oval
and squarely truncate posteriorly ; the pattern of the dorsal and
ventral surface markings is unlike that of any other species of
the genus and must be studied from the figures. The ventral
plate is elongate reversed-ovate and as wide as the dorsal plate.
The lateral sulci are fully developed only on the posterior half,
disappearing gradually towards the anterior margin. The
posterior segment is very large, obtusely pointed and projects
far beyond the dorsal plate. The coxal plates are somewhat ob¬
scure and bluntly pointed posteriorly. The first foot joint is
very long and has a circular enlargement near its anterior end ;
the second joint is subsquare and projects nearly its length
beyond the lorica. The toes are very long and very slender,
nearly one third of the total length, straight, slightly tapering
and rounded at the ends ; the claw is extremely long, slender and
acutely pointed.
Total length 220/a; length of dorsal plate 120/a, of ventral
plate 145/a; width of lorica 84/a; width of anterior margin 60/a;
toes without claw 38/a ; claw 20/a.
Lecane elongata was collected among the leaves of a species of
sphagnum growing on the bottom in shallow bays of Doughty
Mill pond, near Atlantic City, New Jersey. It has not been
found elsewhere.
LECAXE TENUISETA
Plate XXXI, figures 3, 4.
Lecane temdseta Harking, Proc. U. S. Nat. Mus., vol. 47, 1914,
p. 543, pi. 22, figs. 1-3. — Fadeev, Russk. Gidr. Zhurn., vol. 3,
No. 3-5, 1924, p. 5; Raboti Sev.-Kavkazhsk. Gidrobiol. Stants.,
vol. 1, 1925, p. 23, pi. 5, fig. 2 — Hauer, Zool. Anz., vol. 61, 1924,
p. 149.
The lorica is membranous and very flexible; its outline is
elongate oval and the body is fairly thick. The anterior dorsal
and ventral margins are parallel and usually slightly convex
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 373
in the median half of their width; they do not quite meet and
the lorica is left partly open in retraction. The dorsal plate is
oval, rounded posteriorly and without facetting; it is strongly
gibbous and bends down considerably even in front. The
ventral plate is as wide as the dorsal and more elongate oval ; its
markings consist of a few slightly elevated ridges, as shown in
figure 4. The lateral sulci are very shallow. The posterior seg¬
ment of the body is broad and rounded and projects consider¬
ably beyond the dorsal plate. The coxal plates are large and
semicircular. The first foot joint is narrow and parallel-sided
and overlaps the second joint as a rounded lobe with a small
median projection; the second joint is short and broad, rounded
anteriorly and constricted immediately in front of the toes; it
does not project beyond the posterior segment. The toes are
long and slender, about one third of the total length, slightly
conical and terminate in an extremely long, spinelike claw, very
slender and very acute, its inner edge a continuation of the toe
and a distinct shoulder on the outer edge at the junction of toe
and claw.
Total length lOGft; length of dorsal plate 64/x, of ventral plate
73ft; width of lorica 56ft; width of anterior margin 45fi; length
of toe without claw 20fi ; claw 13ft.
Lemne tenuiseta is common in wet sphagnum and in weedy,
soft water ponds, apparently all over the United States. Fadeev
reports it from Kharkov and Lake Madatapin in the Kaukasus,
and Hauer from bogs in the Schwarzwald.
LECANE DORYSSA Harring
Plate XXXI, figures 5, 6.
Lecane doryssa Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
542, pi. 21, figs. 4-6.
The lorica is very broadly pyriform and. its consistency
leathery, so that the boundaries of the plates are not very dis¬
tinct, but the general outline is nevertheless quite constant. The
dorso-ventral depth of the body is unusually great. The
anterior margin of the dorsal plate is slightly convex and
sinuate, as the ridges of the plate are continued up to the edge ;
the ventral margin is almost straight and frontal spines are
not present. The dorsal plate is subcircular and slightly trun¬
cate posteriorly; the facetting is very prominent and deviates
somewhat from the regular pattern, especially towards the
374 Wisconsin Academy of Sciences, Arts and Letters.
margins. The ventral plate is slightly narrower than the dorsal,
approximately parallel-sided anteriorly and narrows somewhat
abruptly to the posterior segment; the surface markings are
prominent and their pattern quite complex, as shown in figure
6. The lateral sulci are shallow and much wrinkled. The
posterior segment is large and projects considerably beyond the
dorsal plate ; the coxal plates are large and rounded. The first
foot joint is large and prominent, strongly constricted near the
middle and overlaps the large, subsquare second joint as a broad,
bluntly pointed lobe ; the second joint projects beyond the lorica
for more than half its length. The toes are very long and
slender, about one third of the total length, straight and very
slightly tapering for a little more than half their length ; at this
point they are abruptly reduced to a clawlike terminal spine,
straight, very slender and very acutely pointed.
Total length 106/^ ; length of dorsal plate 58/x, of ventral plate
58^; width of dorsal plate 60/>t, of ventral plate 58/a; width of
anterior margin 52/a; length of toe without terminal spine 17/x,
of spine 13/a.
Lecane doryssa was described from material collected in the
Panama Canal Zone ; it also occurred in collections made by Mr.
Juday in ditches along the Ferrocarril Central de Guatemala at
Puerto Barrios, Guatemala. It was not numerous at either
location. The nearest relative of this species is probably L.
hornemanni, which rivals it in thickness of the body and also
resembles it in general appearance. The lorica of L. Jiorne-
manni has surface markings of quite a different pattern ; the toes
are without any claw and the peculiar coxal plates, ending in
points close to the foot, also distinguish it from L. doryssa.
LECAXE INOPIXATA Harrinj^ and Myers, new species
Plate XXXII, figures 5, 6.
The lorica is broadly ovate; its width is about two thirds of
the length. The anterior dorsal and ventral margins are coinci¬
dent and very slightly convex; no anterior spines are present.
The dorsal plate is broadly oval and slightly truncate posteri¬
orly ; it is not facetted. The ventral plate is broadly ovate and
very slightly narrower than the dorsal; its surface markings
consist of tw^o transverse and several longitudinal ridges. The
lateral sulci are fairly deep. The posterior segment is small and
rounded and projects somewhat beyond the dorsal plate. The
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 375
coxal plates are small and rounded posteriorly. The first foot
joint is fairly large and pyriform^ lobate posteriorly, the second
joint subsquare and robust. The toes are long and slender, about
one fourth of the entire length, straight and parallel-sided,
terminating in a fairly large, acute claw ; they are fused for one
third of their length and consequently immovable.
Total length 110/x; length of dorsal plate 75/a, of ventral plate
80/^; width of dorsal plate 62/a, of ventral plate 58/a; width of
anterior margin 52/a ; length of toes without claw 26/a ; claw 5/a.
Lecane inopinata was found in small numbers in weedy ponds
near Minocqua, Wisconsin. It may with seemingly equal justifi¬
cation be called a Lecane with fused toes or a Monostyla with a
divided toe; as Moiiostylas are unquestionably Lecanes with
fused toes, it seems advisable to give preference to Lecane where
the fusion is incomplete.
liECAXE NANA (Murray)
Plate .XXXIV, figures 1, 2.
Cathypna nana, Murray, Journ. Royal Micr. Soc., 1913, p. 353, pi.
14, fig. 29.
Lecane nana Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p. 536.
—Hauer, Zool. Anz., vol. 61, 1924, p. 149; Mitt. Geogr. Ges u.
Naturhist. Mus. Lubeck, ser. 2, No. SO, 1925, p. 168, fig. 8.
The outline of the lorica is roughly subcircular. The anterior
margins of the dorsal and ventral plates are coincident and
slightly convex ; no anterior spines are present. The dorsal plate
is subcircular and very slightly pointed posteriorly ; it is without
surface markings. The ventral plate is considerably narrower
than the dorsal, nearly parallel-sided anteriorly and obtusely
pointed posteriorly. The surface markings consist of a few
broken lines, as shown in figure 2. The lateral sulci are deep. The
posterior segment is small and obtusely pointed and projects
somewhat beyond the dorsal plate. The coxal plates are small and
rounded posteriorly. The first foot joint is obtusely conical and
■(videst in front, the second joint irregularly subsquare and does
not project beyond the posterior segment. The toes are long
and fairly slender, about one fourth of the total length, acumin¬
ate and straight on their inner edges.
Total length 85/a, length of dorsal plate 56/a, of ventral plate
64/a; width of dorsal plate 54/a, of ventral plate 48/a; width of
anterior margin 45/a ; length of toes 21/a.
376 Wisconsin Academy of Sciences, Arts and Letters.
Lecane nana was described by Murray from Lake Titicaca.
Bolivia. It occurs also in the Panama Canal Zone, and we have
collected it in wet sphagnum from many localities in the United
States: Central and Northern Wisconsin; Mount Desert Island,
Maine; around Atlantic City, New Jersey; Maryland and the
District of Columbia and in Polk County, Florida. It has a
superficial resemblance to L. tryphema, but the outline of the
lorica and the relative proportions of the dorsal and ventral
plates are quite different.
LiECAVE TRYPHEMA Harring: and Myers, new species
Plate XXXIV, figures 5, 6.
The outline of the lorica is very broadly oval. The anterior
margin of the dorsal plate is approximately straight, but on ac¬
count of the flexibility of the entire lorica it is somewhat wavy
and irregular ; the ventral margin is slightly concave and no
anterior spines are present. The dorsal plate is subcircular and
truncate posteriorly; it is without permanent markings. The
ventral plate is very broadly ovate and rounded posteriorly ; its
markings are limited to a transverse fold in front of the foot.
The lateral sulci are deep. The posterior segment is small,
rounded and its anterior limits obscure; it projects considerably
beyond the dorsal plate and slightly beyond the second foot
joint. The coxal plates are small and rounded posteriorly. The
first foot joint overlaps the second with nearly half its length
and is lobate posteriorly; the second foot joint is very robust
and roughly subsquare. The toes are slender and fairly long,
about one fourth of the total length, straight on their inner
edges and tapering at the extreme ends to conical points without
claws.
Total length 104/^; length of dorsal plate 70/x, of ventral plate
ISfjL; width of dorsal plate 70/a, of ventral plate 66/a; width of an¬
terior margin 53/a; length of toes 27/a.
Lecane tryphema was first collected in a cranberry bog at
Mather, Wisconsin. It is fairly common in bog pools and ponds
among algae and mosses around Atlantic City, New Jersey, and
also on Mount Desert Island, Maine.
Harring & Myers — Rotifer Fauna of Wisconsin. — 111. 377
L.ECA1VE HORNEMANNI (Ehrenberg)
Plate XXXIV, figures 3, 4.
Euchlanis hornemanni Ehrenberg, Abh. Akad. Wiss. Berlin (for
1833) 1834, pp. 206, 220; Infusionsthierchen, 1838, p. 462, pi. 57,
fig. 9.
Distyla hornemanni Hudson and Gosse, Rotifera, SuppL, 1889, p.
42, pi. 33, fig. 37. — Bilfinger, Jahresh. Naturk. Wiirtteniberg,
vol. 50, 1894, p. 58. — Sachse, Siisswasserfauna Deutschlands, pt.
14, 1912, p. 175, fig. 345.
Cathypna hornemanni Murray, Journ. Royal Micr. Soc., 1913, p.
349, pi. 14, fig. 26.
Lecane hornemanni Harring, Proc. U. S. Nat. Mus., vol. 47, 1914,
p. 543; Rep. Canadian Arctic Exp. 1913-18, vol. 8, pt. E, 1921,
p. 8.
The lorica is very broadly ovate ; its width is greater than the
length and the dorso-ventral depth when fully contracted is
unusually great. The anterior dorsal margins are coincident
and decidedly convex ; no anterior spines are present. The dor¬
sal plate is very broadly oval and rounded posteriorly ; its width
is considerably greater than the length. The surface markings
consist of hemispherical bosses, apparently corresponding in
number and position to the usual facets, and separated by wide
depressions of approximately semicircular cross section; there
are no sharp lines of demarcation anywhere, the bosses merging
gradually with the dividing grooves and it is extremely diffi¬
cult to determine their respective locations with any accuracy
and for this reason they have been omitted from the figure. The
ventral plate is very broadly pyriform, considerably narrower
than the dorsal plate, and its edges wavy and somewhat ill-
defined ; the surface markings consist of a few longtiudinal and
transverse folds. The lateral sulci are deep. The posterior seg¬
ment is short, broad and semicircular ; it projects somewhat be¬
yond the dorsal plate. The coxal plates are obtusely pointed.
The first foot joint is fairly large and roughly parallel-sided;
the second joint is very large, somewhat elongate, rounded an¬
teriorly and projects considerably beyond the lorica. The toes
are long and fairly stout, one fourth of the total length, very
slightly tapering for about two thirds of their length, terminat¬
ing in long, conical, acute points, very slightly outcurved and
without any claw.
Total length 120 fx, length of dorsal plate 72/x, of ventral plate
378 Wisconsin Academy of Sciences, Arts and Letters.
84ja; width of dorsal plate 87//., of ventral plate 72jti,; width of
anterior margin 63/x; length of toes 30jii,
Lecane hornemanni is fairly common in weedy ponds ; we have
collected it at Washington, District of Columbia; around Atlan¬
tic City, New Jersey; in northern and central Wisconsin and on
Mount Desert Island, Maine ; it occurred also in collections made
by Dr. H. S. Jennings at Ann Arbor, Michigan.
LiECANE CLARA (Bryce)
Plate XVII, figures 3, 4.
? Diaschiza cupha Gosse, Journ. Royal Micr. Soc., 1887, p. 3, pi. 1,
fig. 6. — Hudson and Gosse, Rotifera, SuppL, 1889, p. 38, pi. 31.
fig. 31.
Distyla clara Bryce, Science Gossip, vol. 28, 1892, p. 271, text fig.
Cathypna clara Murray, Journ. Royal Micr. Soc., 1913, p. 556, pi.
22, fig. 6.
Cathypna sidcata Murray, Journ. Royal Micr. Soc., 1913, p. 557, pi.
22, fig. 5; not Cathypna sulcata Gosse.
Lecane clara Harring, Bull. 81 U. S. Nat. Mus., 1913, p. 60; Rep.
Canadian Arctic Exp. 1913-18, vol. 8, pt. E, 1921, p. 9.
Diglena beauchampi VON Hofsten, Naturw. Enters. Sarekgeb.,
vol. 4, 1923, p. 851, fig. 5.
The lorica is membranous, very flexible and its outline varia¬
ble in the living animal ; it is difficult to obtain fully contracted
specimens, even with strong preservatives, and they are in the
nature of accidents. The following description is based on such
contracted specimens, in order to make comparison possible with
other species of the genus.
The outline of the contracted lorica is broadly oval and
rounded posteriorly. The anterior margins are not coincident
and complete closure of the lorica appears to be impossible; the
dorsal margin is slightly convex and the ventral slightly con¬
cave. Anterior spines are not present. The dorsal plate is oval
and rounded posteriorly ; no permanent folds or other markings
are present. The ventral plate is oval, rounded posteriorly and
somewhat narrower than the dorsal plate. The lateral sulci are
evanescent. The rounded posterior segment has no definite an¬
terior limit; the coxal plates are fairly large and obtusely
pointed posteriorly. The first foot joint is elongate pyriform
and somewhat indistinct, the second joint very broadly pyriform.
The toes are long and robust, about one third of the total length,
broadly lancet-shaped and terminate in a bristle-like spicule.
Harring Myers — Rotifer Fauna of Wisconsin. — III. 379
Total length of contracted specimen 120/x; length of dorsal
plate 80/4, of ventral plate 82/4, width of dorsal plate 62/4, of ven¬
tral plate 55/4 ; width of anterior margin 55/4 ; length of toes, 38/4.
Lecane clara was found by Bryce in sphagnum; we have
found it in weedy ponds at Kenilworth and Hyattsville, near
Washington, District of Columbia, and at Oceanville, near At¬
lantic City, New Jersey.
I/ECAIVE RHACOIS Harring and Myers, new species
Plate XVIT, figures 1, 2.
The integument is so flexible that there is no justiflcation for
calling it a lorica. As in the case of L. clara /ully contracted
specimens are seldom obtained and even then they are virtually
shapeless balls. The characteristic features are the posterior
segment, foot and toes. The posterior segment is limited an¬
teriorly by a distinct fold, representing the termination of the
dorsal plate; it is rounded posteriorly and projects slightly be¬
yond the dorsal plate. The coxal plates are small and very ob¬
tusely pointed posteriorly. The first foot joint is pyriform and
somewhat indistinct, the second joint short, broad and squarish,
projecting about half its length beyond the posterior segment.
The toes are long, stout, straight and nearly parallel-sided; the
edges are not true straight lines, but very slightly wavy and
irregular, varying individually, the points are blunt and have
a minute terminal spicule.
Total length, extended 140/4, contracted lOO/i; length of toes
40/4.
Lecane rhacois is common in wet sphagnum growdng in an old
gravel pit at Hyattsville, near Washington, District of Colum¬
bia, and under similar conditions in Oneida and Yilas Coun¬
ties, Wisconsin.
liECAKE INERMIS (Bryce)
Plate XXXIII, figures 1, 2.
Distyla inermis Bryce, Science Gossip, vol.28,1892,p.274, text fig. —
Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich, vol. 1, 1914, p.
32. — Montet, Rev. Suisse Zool., vol. 23, 1915, p. 336.
Cathypna inermis Murray, Journ. Royal Micr. Soc., 1913, p. 556,
pi. 22, fig. 7. — Weber and Montet, Cat. Invert. Suisse, pt. 11,
1918, p. 187.
380 Wisconsin Academy of Sciences j Arts and Letters.
Lecane inermis Harking, Bull. 81 U. S. Nat. Mus., 1913, p. 61. —
Bryce, Journ. Quekett Micr. Club, ser. 2, vol. 15, 1924, p. 97.
Lecane amorpha Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
544, pi. 23, figs. 1, 2.~Hauer, Zool. Anz., vol. 61, 1924, p. 148.
The lorica is membranous and very flexible and the outlines of
the dorsal and ventral plates somewhat variable, but neverthe¬
less fairly constant. The outline of the fully contracted lorica
is broadly pyriform and rounded posteriorly. The anterior
margins are coincident and usually straight, dependent some¬
what on the degree of contraction; the lorica is widely open in
front, even when fully contracted. The dorsal plate is approxi¬
mately parallel-sided anteriorly and rounded posteriorly. The
ventral plate is broadly ovate and considerably wider than the
dorsal plate; both are without permanent surface markings.
The lateral sulci are represented by the very flexible and slightly
concave membrane connecting the dorsal and ventral plates.
The posterior segment is small and rounded, projecting nearly
its entire length beyond the dorsal plate. The coxal plates are
small and obtusely pointed. The first foot joint is somewhat
wedge-shaped and pointed posteriorly, the second joint large
and subsquare, projecting slightly beyond the lorica. The toes
are relatively short, straight and slightly tapering, rounded at
the ends and terminate in a very slender, acutely pointed,
slightly recurved claw, nearly as long as the toe itself.
Total length 115/a ; length of dorsal plate 80/a, of ventral plate
86/a; width of dorsal plate 42/a, of ventral plate 48/a; width of
anterior margin 40/a ; length of toes without claw 16/a ; claw 12/a.
Lecane inermis is common everywhere in wet sphagnum. It is
the best known representative of a peculiar group within the
genus of virtually illoricate species, composed of L. inermis^ L.
clara, L. rhacois, L. agilis, L. palinacis and L. calcaria. Their
similarity may, however, be a result of simple convergence
rather than an indication of any actual relationship.
L.ECAXE PAL.INACIS Marring and Myers, new species
Plate XXXII, figures 3, 4.
The integument is very flexible and shows no trace of a true
lorica. The animal contracts very readily and the body assumes
an almost spherical form without indictions of either a dorsal or
ventral plate. The posterior segment is, however, distinctly
separated from the body, short and very broad ; at its external
angles are two prominent, curved, acute spines, probably repre-
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 381
senting rudimentary coxal plates. The first foot joint is
elongate and somewhat pyriform, lobate posteriorly; the second
joint is very large, broadly elliptic anteriorly and sharply in¬
dented immediately in front of the toes, projecting beyond the
posterior segment nearly two thirds of its length. The toes are
fairly long and stout, about one fourth of the total length,
straight and very slightly tapering; they terminate in a small,
acute, slightly outcurved claw.
Total length, extended 120^, contracted 75/x; length of body,
contracted 60/x; width 47/^; coxal spines 4/a; length of toes with¬
out claw 14/a; claw 4/a.
Lecane palinacis is fairly common in wet sphagnum in various
localities around Washington, District of Columbia, and also in
moss forwarded from Massachusetts by Mrs. A. C. Clarke, of
the Quekett Microscopical Club.
LECAIVE CALiCARIA Harring and Myers, new species
Plate XXXIII, figures 3, 4.
The integument is very flexible and shows but slight traces of
a lorica, but the form is fairly constant ; the outline is very
broadly reversed-ovate. The anterior margin is approximately
straight ; no anterior spines are present and the lorica is widely
open in front when fully contracted. The posterior segment is
limited anteriorly by a distinct fold indicating the posterior
end of a dorsal plate; the segment is nearly semicircular and
projects far beyond the dorsal fold. No lateral sulci are
present. The coxal plates are very firm, extremely large, curv¬
ed and pointed posteriorly and project considerably beyond the
posterior segment. The first foot joint is elongate, rather
narrow and somewhat indistinct, the second joint very large and
broadly ovate, projecting about one third of its length belond
the body. The toes are fairly long and very stout, about one
fourth of the total length, strongly outcurved, broad at the base
and taper rapidly to very acute points.
Total length 65/a; length of body 50/a, width 41/a; width of
anterior margin 42/a; toes 15/a.
Lecane calcaria was collected in large numbers in a swamp at
Oceanville, near Atlantic City, New Jersey, some four years ago;
it has not occurred elsewhere.
382 Wisconsin Academy of Sciences, Arts and Letters.
I.ECANE NIOTHIS Harring and Myers, new species
Plate XXXIII, figures 5, 6.
The outline of the lorica is broadly ovate and somewhat
irregular. The integument is quite flexible and the anterior
margins variable, but usually strongly convex and coincident.
The dorsal plate is subcircular and rounded posteriorly; the
facetting is rather faint, but the pattern is quite constant and
somewhat unusual. The ventral plate is of the same width and
length as the dorsal; its surface markings also differ slightly
from the normal pattern. The lateral sulci are indistinct. The
posterior segment is short, broad and rounded; it projects
slightly beyond the dorsal plate. The coxal plates are very
small, rounded and indistinct. The first foot joint is indistinct
and ovate, the second large and broadly pyriform. The toes are
fairly long and stout, slightly less than one third of the total
length, parallel-sided at the base and taper gradually to acute
points without any claw.
Total length 70^; length of lorica 50/x, width 48/x; length of
toes 20/x.
Lecane niothis was collected among the leaves of a species of
sphagnum growing on the bottom of some shallow, artificial ponds
at Manse t, on Mount Desert Island, Maine ; it has not been found
elsewhere. It is one of the very small, soft-bodied Leeanes, but
readily recognized by the well marked lorica.
LECAXE AGILIS (Bryce)
Plate XXXII, figures 1, 2.
Distyla agilis Bryce, Science Gossip, vol. 28, 1892, p. 273, text
figures.
Lecane agilis Harking, Bull. 81 U. S. Nat. Mus., 1913, p. 60.
The lorica is very soft and flexible and therefore somewhat in¬
definite in outline ; the more usual form is a moderately elon¬
gate oval, squarly truncate anteriorly. The lateral edges of
the dorsal plate are ill-defined and wavy, the anterior margin
straight. The ventral plate is slightly wider than the dorsal and
its straight anterior margin projects somewhat beyond the an¬
terior margin of the dorsal plate. Both plates are marked
with very faint, irregular ridges in a definite pattern. The
posterior segment projects slightly beyond the dorsal plate ; the
indistinct coxal plates are rounded at the posterior angles. The
Earring & Myers — Rotifer Fauna of Wisconsin. — 111. 383
first foot joint is very obscure, the second small and pyriform.
The toes are very short and blade-shaped with acute, outcurved
points; they appear to be fused anteriorly for nearly half of
their length.
Total length 55/a; length of dorsal plate 38/a, of ventral plate
45/a; width of dorsal plate 25/a, of ventral plate 30/a; width of
anterior dorsal margin 14/a, ventral margin 20/a; toes 12/a.
Lecane agilis seems to be found only in wet sphagnum; it ap¬
pears to be rare, but this may be on account of its minute size
and excessive transparency. It moves about among the debris
with a peculiar jerky motion, but never swims.
Genus MONOSTYLA Ehrenberg
Euchlanid rotifers with illoricate, retractile head and loricate
body, strongly compressed dorso-ventrally and oval or ovate in
outline; dorsal and ventral plates connected by a flexible mem¬
brane, forming lateral and posterior sulci; foot with two ex¬
tremely short, rudimentary joints, of which only the posterior
is movable ; toe single ; corona of family type ; mastax modifled
malleate, with a piston attached to its ventral wall; eyespot
single and at the posterior end of ganglion; retrocerebral sac
usually present, but no subcerebral glands.
Type of the genus. — Monostyla cornuta ( Muller )=Tnc7zoda
cor nut a Muller.
The following species have not been studied. We may say that
the specimen from which Murray described his 31onostyla coch-
learis was sent to us by the late Mr. Eousselet; it was in such
poor condition that we consider it a very doubtful species.
Monostyla amazonica Murray, Journ. Royal Micr. Soc., 1913, p. 354,
pi. 15, fig. 34.
Monostyla asymmetrica Murray, Journ. Rojral Micr. Soc., 1913, p.
361, pi. 15, fig. 44.
Monostyla cochlearis, Murray, Journ. Royal Micr. Soc., 1913, p. 361,
pi. 15, fig. 45.
Monostyla cornuta anglica Bryce, Journ. Quekett Micr. Club, ser. 2,
vol. 15, 1924, p. 98.
Monostyla dentiserratus Mola, Zool. Anz., vol. 42, 1913, p. 122, figs.
16, 17; Ann. Biol. Lac., vol. 6, 1913, p. 265.
Monostyla falcata Murray, Journ. Royal Micr. Soc., 1913, p. 558, pi.
22, fig. 9.
Monostyla lordii Gosse, in Hudson and Gosse, Rotifera, 1886, vol. 2,
p. 99, pi. 25, fig. 5.
384 Wisconsin Academy of Sciences, Arts and Letters.
Monostyla lunaris aperta Steinecke, Schriften Phys.-okon. Ges.
Konigsberg i. P., vol. 57, 1916, p. 96, figs. 2c, 2e; vol. 64, 1924, p. 41.
Monostyla macrognatha Schmarda, Neue wirbellose Thiere, 1859,
vol. 1, p. 59, pi. 14, fig. 134.
Monostyla dophthalma Schmarda, Neue wirbellose Thiere, 1859, vol. 1,
p. 59, pi. 14, fig. 126.
Monostyla ovalis Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich, vol.
1, 1914, p. 34, pi. 1, fig. 11.
Monostyla ovata Forbes, Bull. U. S. Fish Comm., vol. 11 (for 1891),
1893, p. 256.
Monostyla paradoxa Steinecke.
Monostyla lunaris paradoxa Steinecke, Schriften Phys.-okon. Ges
Konigsberg i. P., vol. 57, 1916, p. 97, fig. 3.
Monostyla paradoxa Steinecke, Schriften Phys.-okon. Ges Konigs¬
berg i. P., vol. 64, 1924, p. 41.
Monostyla tentaculata COSMOVICI, Naturaliste (Paris), vol. 14, 1892,
p. 70; Anal. Acad. Romana, ser. 2, vol. 28, 1906, p. 44, fig. 29; not
a Monostyla.
Monostyla testudinea Mola, Zool. Anz., vol. 42, 1913, p. 120, fig. 13;
Ann. Biol. Lac., vol 6, 1913, p. 263.
Monostyla unguitata Fadeev, Trudy Kharkovsk. Obshch. Isp. Prir.,
vol. 50, pt. 1, 1925, p. 9, pi. 1, fig. 7.
Monostyla ungulata Mola, Zool. Anz., vol. 42, 1913, p. 122, figs. 14,
15; Ann. Biol. Lac., vol. 6, 1913, p. 264.
4
MONOSTYLA LUNARIS (Ehreiiberg)
Plate XXXV, figures 1-6.
Lepadella lunaris Ehrenberg, Abh. Akad. Wiss. Berlin (for 1831),
1832, p. 127.
Monostyla lunaris Ehrenberg, Infusionsthierchen, 1838, p. 460, pi.
57, fig. 6. — Dujardin, Hist. Nat. Zooph., Inf., 1841, p. 635. — Da-
DAY, Termesz. Fiizetek, vol. 9, 1885, p. 127; vol. 19, 1892, p. 28;
Math. Termesz. Ertes., vol. 12, 1893, p. 19; Zoologica, pt. 44,
1905, p. 113; pt. 59, 1910, p. 85. — Hudson and Gosse, Rotifera,
1886, vol. 2, p. 98, pi. 25, fig. 2. — Levander, Acta Sdc. Fauna et
Flora Fennica, vol. 12, No. 3, 1895, p. 50, pi. 3, fig. 32. — Skorikov,
Trav. Soc. Nat. Kharkow, vol. 30, 1896, p. 321. — Jennings, Bull.
Michigan Fish Comm., No. 3, 1896, p. 93; Bull. U. S. Fish Comm.,
vol. 19 (for 1899), 1900, p. 92, pi. 21, fig. 41. — Hempel, Bull. Illi¬
nois State Lab. Nat. Hist., vol. 5, 1898, p. 375 — Voigt, Forsch-
ungsber. Biol. Stat. Plon, vol. 11, 1904, p. 72. — Voronkov, Trudy
Gidr. Slants. Glubokom Oz., vol. 2, 1907, pp. 110, 205, 284. — Ko-
FOID, Bull. Illinois State Lab. Nat. Hist., vol. 8, No. 1, 1908, p.
201. — Meissner, Izv. Turkestansk. Otd. Imp. Russk. Geogr.
Obshch., vol. 4, pt. 8, 1908, p. 22. — Schlenker, Mitt. Gool. Abt.
Wiirttemberg. Stat. Landesamt, No. 5, 1908, p. 249. — Runnstrom,
Zool. Anz., vol. 34, 1909, p. 271. — VoN Hofsten, Arkiv Zool.,
Stockholm, vol. 6, No. 1, 1909, p. 59; Naturw. Enters. Sarekgeb.,
Harring & Myers — Rotifer Fauna of Wisconsin. — HI. 385
vol. 4, 1923, p. 861.~~Lie™Pettersen, Bergens Mus. Aarbog (for
1909), 1910, No. 15, p. 66.- — Lucks, Rotatorienfauna Westpreus-
sens, 1912, p. 112.---Sachse, Susswasserfauna Deutschlands, pt.
14, 1912, p, 179, fig. 355.— Mola, Ann. Biol. Lac., vol. 6, 1913, p.
263.— Murray, Journ. Royal Micr. Soc., 1913, p. 353, pi. 15, fig.
31.— Kozar, Zool. Anz., vol. 44, 1914, p. 420.— Montet, Rev. Suis¬
se ZooL, vol. 23, 1915, p. 335.— Jakubski, Rozpr. Wiad. Muz.
Dzieduszyckich, vol. 1, 1914, p. 34, pi. 1, fig. 12; Kosmos (Lwow),
1918-1919, p. 28.— Rezvoi, Trudy Borodinsk. Biol. Stants. Imp.
Petrogradsk. Obshch. Estestvoisp,, vol. 4, 1916, p. 181.— Stein-
ECKE, Schriften Phys.-okon. Ges. Konigsberg i. P., vol. 57, 1916,
p, 89, fig. 2; vol. 64, 1924, p. 41.— Weber and Montet, Cat. In¬
vert. Suisse, pt. 11, 1918, p. 193.— Olofsson, Zool. Bidr. Uppsala,
vol. 6, 1918, p. 694, fig. 64.— Harring, Rep. Canadian Arctic Exp.
1913-18, vol. 8, pt. E, 1921, p. 10.™ Bryce, Journ. Quekett Micr.
Club, ser. 2, vol. 14, 1922, p, 313. — Idelson, Trudy Plovuch.
Morsk. Nauchn. Inst., Moskva, pt. 12, 1925, p. 90.
Monostyla quennerstedti Bergendal, Acta Univ. Lundensis, vol. 28,
1892, sect. 2, No. 4, p. 118, pi. 6, fig. 39.
Monostyla constricta Murray, Journ. Royal Micr. Soc. 1913, p. 557,
pi. 22, fig. 10.
Monostyla virga Harring, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
546, pi, 24, figs. 1-3. — Fadeev, Russk. Gidr. Zhurn., vol. 3, No.
3-5, 1924, p. 6, fig. 1. — Oparina-Kharitonova, Izv, Biol. Nauch-
no-Issl. Inst, Permsk. Univ., vol. 3, 1925, p. 440.
The outline of the loriea is broadly ovate; its width is about
two thirds of the length. The anterior margin of the dorsal
plate is narrow and has a V-shaped sinus, broadly rounded at
the posterior angle; the ventral margin is wide and the sinus
deep, occasionally with minute, lateral cusps opposite the ante¬
rior points of the dorsal plate. The dorsal plate is very broadly
ovate and rounded posteriorly. The ventral plate is somewhat
narrower than the dorsal and broadly oval. The surface mark¬
ings consist of a transverse dorsal fold at the base of the anterior
sinus and a transverse ventral fold a short distance in front of
the foot. The lateral sulci are fairly deep. The posterior seg¬
ment is large and rounded ; it projects considerably beyond the
dorsal plate. The coxal plates are rather small and obtusely
pointed posteriorly. The first foot joint is short, parallel-sided
and somewhat indistinct, the second joint subsquare and robust.
The toe is very long and slender, more than one third of the
total length, straight and parallel-sided ; it has two faint annu¬
lar constructions, dividing it into three nearly equal sections.
The claw is long, slender and acutely pointed; at the base are
two minute spicules.
25
386 Wisconsin Academy of Sciences, Arts and Letters.
Total length 165/a; length of dorsal plate 100/a, of ventral plate
118/a; width of dorsal plate 86/a, of ventral plate 80/a; width of
anterior dorsal margin 30/a, ventral margin 43/a ; toe without claw
58/a ; claw 8/a.
Monostyla lunaris is abundant in weedy ponds all over the
world. The outline of the anterior margin varies considerably
according to the degree of contraction ; the form of figures 1 and
2 is what may be expected in collections preserved in alcoholic
mixtures and should be considered fully contracted. Figures
3 and 4 represent the normal form, with a shallower anterior
sinus and broader anterior margin. At English Creek and in a
cranberry bog near Egg Harbor, New Jersey, occurs a peculiar
variety, shown in figures 5 and 6; the lorica is much broader
than in the normal form and the posterior segment is angular
and squarely truncate ; the claw has a distinct median line, but
is not actually double.
Monostyla virga Barring, described from the Isthmus of Pan¬
ama, appears to be a form of M. lunaris, the differences noted
being probably due to the preservative used.
MONOSTYLA CRENATA Harring
Plate XXXVI, figures 5, 6.
Monostyla crenata Harking, Proc U. S. Nat. Mus., vol. 46, 1913, p.
399, pi. 36, figs. 4-6.
The outline of the lorica is broadly ovate; its width is about
two thirds of the length. The anterior dorsal margin is very
slightly concave; the ventral plate has a deep anterior sinus,
rounded at the bottom and with curved, convex sides. The dor¬
sal plate is broadly ovate and slightly truncate posteriorly. The
ventral plate is broadly oval and somewhat narrower than the
dorsal plate. The surface markings are limited to a transverse
ventral fold a short distance in front of the foot. The lateral
sulci are deep. The posterior segment is small and rounded;
it projects slightly beyond the dorsal plate. The coxal plates are
rather small and obtusely pointed ; they do not reach beyond the
first foot joint. This is oval; the second joint is robust and
somewhat elongate reniform. The toe is extremely long and
slender, a little less than half the total length, straight and par¬
allel-sided, terminating in a short claw with two minute basal
spicules.
Total length 200/a ; length of dorsal plate 108/a, of ventral plate
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 387
116/x; width of dorsal plate 92/x, of ventral plate 82ju,; width of
anterior dorsal margin 38/x, of ventral margin 42ju,; toe without
claw 84ft ; claw 8ft.
Monostyla crenata is widely distributed in weedy ponds; we
have collected it around Washington, District of Columbia ; Polk
County, Florida; at Atlantic City, New Jersey; in Oneida and
Vilas Counties, Wisconsin, and on Mount Desert Island, Maine ;
it was common in collections made by Dr. Birge during the
Great Lakes Biological Investigations in 1899, and by Dr. Birge
and Mr. Juday in Texas, Arkansas, Mississippi and Louisiana
in 1903. James Murray found it at Sydney, Australia, and in
New Zealand.
MONOSTYLA ACUS Harring
Plate XXXVI, figures 3, 4.
Monostyla acus Harking, Proc. U. S. Nat. Mus., vol. 46, 1913, p.
398, pi. 36, figs. 1-3. — Hauer, Zool. Anz., vol. 61, 1924, p. 147.
? Monostyla lunaris obserata Steinecke, Schriften Phys.-okon.
Ges. Konigsberg i. P., vol. 57, 1916, p. 97, fig. 2, e, f; vol. 64,
1924, p. 41.
The outline of the lorica is broadly ovate; its width is more
than four fifths of the length. The anterior margins of the dor¬
sal and ventral plates are identical in outline, with a shallow,
broadly V-shaped sinus; the ventral margin projects very
slightly beyond the dorsal. No anterior spines are present.
The dorsal plate is broadly oval and truncate posteriorly. The
ventral plate is somewhat narrower than the dorsal and oval
in outline; there is a transverse ventral fold in front of the
foot. The lateral sulci are deep. The posterior segment is trun¬
cate and very large, projecting its entire length beyond the
dorsal plate. The coxal plates are small and rounded. The first
foot joint is parallel-sided and rounded anteriorly; the second
joint is large and slightly angular. The toe is extremely long
and slender, nearly half the entire length, parallel-sided and
straight, with three internal annular constructions, equally
spaced from the foot; the claw is long and slender and has two
minute, basal spicules
Total length 180ft ; length of dorsal plate 85ft, of ventral plate
100ft; width of dorsal plate 82ft, of ventral plate 72ft; width of
anterior dorsal margin 40ft, of ventral margin 44ft ; length of toe
without claw 70ft; claw 10ft.
388 Wisconsin Academy of Sciences, Arts and Letters.
Monostyla acus is common in wet sphagnum in the United
States wherever we have made collections. Hauer reports it as
common in the Schwarzwald.
MONOSTYLA BULLA Gosae
Plate XXXVII, figures 1, 2.
Monostyla bulla Gosse, Ann. Mag. Nat. Hist., ser. 2, vol. 8, 1851,
p. 200. — Hudson and Gosse, Rotifera, 1886, vol. 2, p. 99, pi. 25,
fig. 4. — Ternetz, Rotat. Umg. Easels, 1892, p. 17. — Wierzejski,
Rozpr. Akad. Umiej., Wydz. Mat.-Przyr., Krakow, ser. 2, vol. 6,
1893, p. 243. — Skorikov, Trav. Soc. Nat. Kharkow, vol. 30, 1896,
p. 321. — Hempel, Bull. Illinois State Lab. Nat. Hist., vol. 5, 1898,
p. 375. — Stenroos, Acta Soc. Fauna et Flora Fennica, vol. 17,
No. 1, 1898, p. 163, pi. 3, fig. 9. — Jennings, Bull. U. S. Fish
Comm., vol. 19 (for 1899), 1900, p. 93, pi. 21, figs. 37-39. — Daday,
Zoologica, pt. 44, 1905, p. 112; pt. 59, 1910, p. 84. — Voronkov,
Trudy Gidr. Stants. Glubokom Oz., vol. 2, 1907, pp. 110, 205,
284. — Kofoid, Bull. Illinois State Lab. Nat. Hist., vol. 8, No. 1,
1908, p. 201. — Meissner, Izv. Turkestansk. Otd. Imp. Russk.
Geogr. Obshch. vol. 4, No. 8, 1908, p. 22. — Schlenker, Mitt. Geol.
Abt. Wurttemberg. Stat. Landesamt, No. 5, 1908, p. 249. — De
Beauchamp, Arch Zool. Exper., ser. 4, vol. 10, 1909, p. 161. —
Runnstrom, Zool. Anz., vol. 34, 1909, p. 271. — Von Hofsten,
Arkiv. Zool., Stockholm, vol. 6, No. 1, 1909, p. 61; Naturw. Un-
ters. Sarekgeb., vol. 4, 1923, p. 861. — Lie-Pettersen, Bergens
Mus. Aarbog (for 1909), 1910, No. 15, p. 67. — Lucks, Rotatorien-
fauna Westpreussens, 1912, p. 112. — Sachse, Siisswasserfauna
Deutschlands, pt. 14, 1912, p. 179, fig. 356. — Murray, Journ.
Royal Micr. Soc., 1913, p. 353, pi. 15, fig. 33. — Mola, Ann. BioL
Lac., vol. 6, 1913, p. 262. — Kozar, Zool. Anz., vol. 44, 1914, p.
420. — Harring, Proc. U. S. Nat. Mus., vol. 47, 1914, p. 547 ; Rep.
Canadian Arctic Exp. 1913-18, vol. 8, pt. E, 1921, p. 10. — Ja-
KUBSKi, Rozpr. Wiad. Muz. Dzieduszyckich, vol. 1, 1914, p. 32;
Kosmos (Lwow), 1918-1919, p. 28. — Rezvoi, Trudy Borodinsk.
Biol. Stants. Imp. Petrogradsk. Obshch. Estestvoisp., vol. 4, 1916,
p. 181. — Weber and Montet, Cat. Invert. Suisse, pt. 11, 1918, p.
194. — Bryce, Journ. Quekett Micr. Club, ser. 2, vol. 15, 1924, p.
98.
Monostyla lunaris Perty, Zur Eenntn. kleinst. Lebensf., 1852, p. 41,
pi. 1, fig. 4. — Eckstein, Zeitschr. Wiss. Zool., vol. 39, 1883, p. 381,
pi. 27, figs. 47-49. — Blochmann, Mikr. Tierw. Sussw., 1886, p.
107, pi. 7, fig. 241. — Weber, Rev. Suisse Zool., vol. 5, 1898, p. 608,
pi. 22, figs. 12, 13. Not Monostyla lunaris Ehrenberg.
Monostyla bipes Stokes, Ann. Mag, Nat. Hist., ser. 6, vol. 18, 1896,
p. 23, pi. 8, figs. 11-13.
Monostyla incisa Daday, Math. Termesz. Ertes., vol. 15, 1897, p. 137,
fig. 5; Termesz. Fiizetek, vol. 24, 1901, p. 22, fig. 5.
liarring & Myers — Rotifer Fauna of Wisconsin. — III. 389
The outline of the very firm lorica is somewhat elongate ovate ;
its width is about three fifths of the length. The anterior dor¬
sal margin has a shallow, V-shaped anterior sinus with a large,
median notch for the protrusion of the dorsal antenna. The
ventral margin has a very deep anterior sinus, rounded at the
posterior end and with a very slight cusp near the front. The
dorsal plate is slightly elongate ovate and rounded posteriously ;
the ventral plate is virtually identical in outline and of the
same width. Surface markings are limited to a transverse, ven¬
tral fold in front of the foot. The lateral sulci are deep. The
posterior segment is small and rounded; it projects slightly be¬
yond the dorsal plate. The coxal plates are very small and
rounded posteriorly. The first foot joint is short, broad and
somewhat indistinct, the second joint large and somewhat tri¬
angular, narrowed posteriorly. The toe is very long and slen¬
der, about one third of the total length, slightly enlarged in the
middle, and ends in a long, slender, acute claw with distinct
basal spicules. The claw has a distinct median line, but is not
divided.
Total length ITOju,; length of dorsal plate 112ju., of ventral plate
118/x; width of lorica TG/x; width of anterior margin 36ft; length
of toe without claw 46fi ; claw I2ft.
Monostyla hulla is abundant everywhere in weedy ponds all
over the world. The form of the anterior margin is somewhat
dependent on the degree of contraction of the lorica.
MONOSTYIiA STYHIAX Harring anil Myers, new species
Plate XXXVII, figures 3, 4.
The lorica is very firm and its outline broadly ovate ; its width
is about three fourths of the length. The anterior margin is
narrow and has a wide, median, elliptic notch for the protrusion
of the dorsal antenna. The ventral margin has a deep, straight¬
sided anterior sinus, rounded postriorly. The dorsal plate is
broadly ovate and rounded posteriorly; the ventral plate is
nearly identical in outline and of the same width. No surface
markings are present except a ventral, transverse fold in front
of the foot. The lateral sulci are very deep. The posterior seg¬
ment is very short and rounded ; it projects very slightly beyond
the dorsal plate. The coxal plates are very small and obtusely
pointed posteriorly. The first foot joint is very short, broad and
somewhat indistinct, the second joint large and slightly hexa-
390 Wisconsin Academy of Sciences^ Arts and Letters.
gonal. The toe is very long and slender, more than one third of
the total length, and is narrowed at one third of its length;
from this point it increases slightly in width and finally tapers
to a blunt, obscurely wrinkled point. The claw is extremely
long, slender and acutely pointed.
Total length 200/a; length of dorsal plate 124/x, of ventral
plate 128/x.; width of lorica 90fi; width of anterior margin 32^;
length of toe without claw 52/x ; claw 24/x.
Monostyla styrax is not rare in soft water ponds; we have
collected it in Oneida and Vilas counties, Wisconsin, on Mount
Desert Island, Maine, and at Atlantic City, New Jersey. It is
related to M. 'bulla, but differs considerably in the form of the
anterior margin, the foot and the toe ; the long, needle-like claw
is without a parallel in the genus.
MONOSTYIiA GONIATA Harrinj^ and Myers, new species.
Plate XXXVII, figures 5, 6.
The outline of the lorica is very broadly ovate and somewhat
angular; its width is about four fifths of the length. The
anterior dorsal margin has a very shallow, Y-shaped sinus with
a broad, nearly semicircular excision for the dorsal antenna; the
ventral sinus is very deep, narrow and rounded posteriorly with
a minute cusp near the front. The dorsal plate is very broadly
ovate-angular and sharply pointed posteriorly. The ventral
plate is of approximately the same width and outline as the
dorsal plate, but distinctly narrowed in front of the foot. No
surface markings are present. The lateral sulci are fairly deep,
but not visible in either the dorsal or ventral view. The
posterior segment is rather small, somewhat angular and slightly
concave terminally. The coxal plates are small and obtusely
rounded. The first foot joint is short, very broad and somewhat
indistinct, the second joint large and roughly trapezoidal, pro¬
jecting slightly beyond the lorica. The toe is very long, about
one third of the total length, stout and spindle-shaped; it is dis¬
tinctly enlarged near mid-length and terminates in a very acute,
conical point resembling a claw and having a median line, which
does not indicate a division, but possibly the opening of the
mucus duct.
Total length 200ft; length of lorica 145/t; width of lorica 115ft;
width of anterior margin 40ft ; length of toe 65ft.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 391
Monostyla goniaia has been collected in large numbers in a
shallow pond at Eagle Elver, Vilas County, Wisconsin, and in a
cranberry bog at English Creek, near Atlantic City, New Jersey.
It is closely related to M. hulla, but differs in so many details
that it seems advisable to list it as a distinct species.
MONOSTYLA QUADRIDENTATA Elirenberg
Plate XXXVIII, figures 3-5.
Monostyla quadridentata Ehrenberg, Abh. Akad. Wiss. Berlin (for
1831), 1832, p. 130; Infusionsthierchen, 1838, p. 459, pi. 57, fig.
5. — Dujardin, Hist. Nat. Zooph., Inf., 1841, p. 635.-— Bartsch,
Jahresh. Naturk. Wiirttemberg, vol. 26, 1870, p. 358; Rotat. Hun-
gariae, 1877, p. 47. — Eyferth, Einf. Lebensformen, 1878, p. 88;
ed. 2, 1885, p. 113.— Hudson and Gosse, Rotifera, 1886, vol. 2,
p. 100, pi. 25, fig. 3. — Anderson, Journ. Asiatic Soc. Bengal, vol.
58, pt. 2, 1889, p. 355. — Ternetz, Rotat. Umg. Basels, 1892, p.
18. — WiERZEJSKi, Rozpr. Akad. Umiej., Wydz. Mat.-Pryzyr.,
Krakow, ser. 2, vol. 6, 1893, p. 243. — Glasscott, Proc. Royal Dub¬
lin Soc., new ser., vol. 8, 1893, p. 73. — Kertesz, Budapest Rotat.
Faun., 1894, p. 40. — Skorikov, Trav. Soc. Nat. Kharkow, vol. 30,
1896, p. 322. — Hempel, Bull. Illinois State Lab. Nat. Hist., vol. 5,
1898, p. 376. — Jennings, Bull. U. S. Fish Comm., vol. 19 (for 1899),
1900, p. 92, pi. 21, fig. 40. — Daday, Zoologica, pt. 44, 1905, p. 113;
pt. 59, 1910, p. 85. — Voronkov, Trudy Gidr. Stants. Glubokom
Oz., vol. 2, 1907, p. 110. — Kofoid, Bull. Illinois State Lab. Nat.
Hist., vol. 8, No. 1, 1908, p. 201. — Meissner, Izv. Turkestansk.
Otd. Imp. Russk. Geogr. Obshch., vol. 4, pt. 8, 1908, p. 21. —
Lucks, Rotatorienfauna Westpreussens, 1912, p. 111. — Sachse,
Susswasserfauna Deutschlands, pt. 14, 1912, p. 181, fig. 358. —
Mola, Ann. Biol. Lac., vol. 6, 1913, p. 264. — Murray, Journ.
Royal Micr. Soc., 1913, p. 354, pi. 15, fig. 34. — Harring, Proc.
U. S. Nat. Mus., vol. 47, 1914, p. 547. — Jakubski, Rozpr. Wiad.
Muz. Dzieduszyckich, vol. 1, 1914, p. 35; Kosmos (Lwow), 1918-
1919, p. 28. — Rezvoi, Trudy Borodinsk. Biol. Stants. Imp. Petro-
gradsk. Obshch. Estestvoisp., vol. 4, 1916, p. 181. — ^Weber and
Montet, Cat. Invert. Suisse, pt. 11, 1918, p. 195. — Bryce, Journ.
Quekett Micr. Club, ser. 2, vol. 15, 1924, p. 99.
Lepadella cornuta Schmarda, Neue wirbellose Thiere, 1859, vol. 1,
p. 58, pi. 14, fig. 122; not Lepadella cornuta (Muller) of Bory
de St. Vincent, 1826.
Metopidia cornuta Hudson and Gosse, Rotifera, Suppl., 1889, p. 47,
pi. 34, fig. 3.
Monostyla bicornis Daday, Math. Termesz. Ertes., vol. 15, 1897, p.
139, text fig ; Termesz. Fuzetek, vol. 24, 1901, p. 23, fig. 6.
The outline of the lorica is very broadly ovate; its width is
about three fourths of the total length. The anterior margin is
relatively narrow ; the dorsal plate has a very deep and narrow,
392 Wisconsin Academy of Sciences^ Arts and Letters.
pyriform median sinns for the protrusion of the dorsal antenna ;
the sinus is flanked by two stout, outcurved and decurved spines.
The ventral plate has a deep, sharply pointed V-shaped sinus,
its sides very slightly convex near the front ; two minute frontal
spines are present. The dorsal plate is very broadly ovate,
slightly truncate posteriorly; the anterior margin is but one-
half of the greatest width. The ventral plate is of approximate¬
ly the same outline as the dorsal and somewhat narrower. Sur¬
face markings are limited to two divergent dorsal folds or ribs,
originating on the anterior spines, and a ventral transverse fold
in front of the foot. The posterior segment is small and round¬
ed; it is not covered by the dorsal plate. The lateral sulci are
deep. The coxal plates are small and obtusely rounded poste¬
riorly. The flrst foot joint is elongate oval and rather narrow,
the second joint elongate, narrow and subcylindric. The toe is
very long, about one third of the total length, slender, parallel¬
sided and has a faint annular constriction near the posterior
end ; the claw has two small basal spicules and is very long,
slender and acutely pointed.
Total length 225yLt; length of dorsal plate 130/x, of ventral plate
142/>t ; width of dorsal plate 96/x, of ventral plate 110/a ; width of
anterior dorsal margin 30/a, of ventral margin 50/a ; length of toe
without claw 52/a; claw 16/a.
Monostyla qiiadridentata is one of the commonest rotifers in
weedy ponds in the United States and is apparently widely dis¬
tributed on other continents; according to Bryce it is, strangely
enough, rare in Great Britain.
The male is shown in flgure 5 ; we believe this is the only male
Monostyla known or identifled. Unfortunately there is very
little to add to the flgure ; the lorica is fairly rigid and so deeply
colored with the characteristic yellowish-brown tint so con¬
spicuous in the female that it is impossible to make out the in¬
ternal anatomy. Its length is 100/a ; toe 23/a.
MONOSTYLA LAMELLATA Daday
Plate XXXIX, figures 5, 6.
Monostyla lamellata Daday, Math. Termesz. Ertes., vol. 12, 1893,
p. 40, pi. 2, figs. 1, 2; Math. Naturw. Ber. Ungarn, vol. 11, 1894,
p. 319, pi. 24, figs. 1, 2. — Rousselet, Zool. Anz., vol. 21, 1898, p.
595. — Zernov, Izv. Turkestansk. Otd. Imp. Russk. Geogr. Obshch.,
vol. 4, pt. 3, 1903, p. 9, pi. 1, fig. 34. — ^Murray, Journ. Royal Micr.
Soc., 1913, p. 458, pi. 19, fig. 1. — Bryce, Journ. Quekett Micr.
Ilarring d: Myers — Rotifer Fauna of Wisconsin, — HI. 393
•Club, ser. 2, vol. 15, 1924, p. 98. — Fadeev, Raboti Sev.-Kavkazhsk.
Gidrobiol. Slants., vol. 1, pt. 1, 1925, p. 23, pi. 5, fig. 3.
Monostyla appendiculata Skorikov, Zool. Anz.,^ vol. 21, 1898, p. 556,
text fig.
The outline of the lorica is broadly ovate; its width is about
three fourths of the length. The anterior dorsal margin has a
moderately deep sinus with sinuate sides and rounded posterior¬
ly; the ventral sinus is lunate and slightly deeper than the
dorsal sinus. At the external angles are two very large, acutely
triangular spines or cusps. The dorsal plate is broadly ovate
and rounded posteriorly. The ventral plate is very slightly
narrower than the dorsal and the outline nearly identical ; it is
marked with a distinct transverse ventral fold in front of the
foot. The lateral sulci are fairly deep and' terminate at a con¬
siderable distance from the anterior spines, nearly at the level of
the anterior sinus. The posterior segment is short, rounded
posteriorly and produced laterally as two acute, triangular
cusps; it projects in its entire length beyond the dorsal plate.
The coxal plates are large and obtusely pointed posteriorly.
The first foot joint is small and parallel-sided, the second joint
small and rounded. The toe is long and stout, more than one
fourth of the entire length, slightly enlarged beyond mid-length ;
it terminates in a large, acute claw with basal spicules and a
median mucus groove.
Total length 260/a; length of dorsal plate 168/a, of ventral
plate 190/a; width of dorsal plate 128/a, of ventral plate 120/a;
width of anterior spines 72/a; width of posterior segment 70/a;
length of toe without claw 58/a; claw 17/a.
Monostyla lamellata was described by Daday from a slightly
alkaline lake in the Hungarian Alfold at Halas, in the county
» Pest-Pilis-Solt-Kiskun between the Danube and the Theiss,
Skorikov found it in “salt” lakes near Kharkov, Russia. Bryce
records this species from Devils Lake, North Dakota, and we
have found it in collections made by Mr. Juday in alkaline wa¬
ters at San Cristobal, near Mexico City. As far as now known,
it is confined to alkaline waters and has never been found in
strictly fresh water.
MONOSTYLA THALERA Harring amd Myers, new species
Plate XXXIX, figures 3, 4.
The outline of the lorica is broadly oval; its width is about
three fourths of the length. The anterior dorsal margin is con-
o94 Wisconsin Academy of Sciences^ Arts and Letters.
cave and sinuate; the ventral margin has a moderately deep,
V-shaped sinus, slightly rounded at the apex. At the external
angles are two small anterior spines. The dorsal plate is sub¬
rhomboid and rounded posteriorly; the ventral plate is of ap¬
proximately the same outline and a little narrower than the dor¬
sal plate ; it is marked with a transverse fold in front of the foot.
The lateral sulci are fairly deep and do not quite reach the
anterior margin. The posterior segment is large and somewhat
angular, truncate posteriorly, and projects considerably beyond
the dorsal plate. The coxal plates are large and obtusely pointed
posteriorly. The first foot joint is elongate ovate, the second
joint very large and somewhat prismatic. The toe is long and
stout, a little less than one third of the total length, spindle-
shaped and enlarged near the middle ; the claw is long, stout
and acutely pointed and has small, basal spicules and a median
mucus groove..
Total length 240/x ; length of dorsal plate 150/x, of ventral plate
160/a; width of dorsal plate 128/i, of ventral plate 115/a; width
of anterior dorsal margin 60/a, of ventral margin 75/a; length of
toe without claw 58/a; claw 18/a.
Monostyla tlialera was first found in material collected by Mr.
Juday in alkaline ditches draining into the large, very shallow
Lago de San Cristobal, near Mexico City. It was abundant
in collections made by Dr. W. E. Allen in sloughs of the San
Joaquin River, at Stockton, California; here the water is prob¬
ably slightly brackish, rather than alkaline, as there is a tide of
3 feet and the gradient of the river is very slight between Stock-
ton and the Bay of San Francisco. This is probably the species
listed by Bryce as Monostyla lamellata var. from Devils Lake,
South Dakota. Although there is, perhaps, a certain superficial
similiarity between these two forms, they differ so much in detail
that there can be no doubt of their being specifically distinct,
even though they are the two largest species of the genus and
sometimes occur together.
MO^fOSTYIiA STENROOSI Meissner
Plate XXXIX, figures 1, 2.
Monostyla bicornis Stenroos, Acta Soc. Fauna et Flora Fennica,
vol. 17, No. 1, 1898, p. 164, pi. 2, fig. 26; not Monostyla bicornis
Daday, 1897. — ? Voronkov, Trudy Gidr. Stants. Glubokom Oz.,
vol. 2, 1907, p. 285, pi. 6, figs. 6-8. — Murray, Journ. Royal Micr.
Harring Myers — Rotifer Fauna of Wisconsin. — III. 395
Soc., 1913, p. 355, pi. 15, fig. 35. — Sachse, Arch. Hydr., vol. 10,
1914, p. 71, fig. 11.
Monostyla stenroosi Meissner, Izv. Turkestansk. Otd. Imp. Russk.
Geogr. Obshch., vol. 4, pt. 8, 1908, p. 22, pi. 1, fig. 8. — Myers,
Proc. U. S. Nat. Mus., vol. 52, 1917, p. 476.
The outline of the lorica is very broadly oval; its width is
about five sixths of the length. The anterior dorsal margin is
straight and narrow, the lateral sulci reducing its width some¬
what. The ventral margin has a shallow, rounded sinus with
strongly convex sides and externally two short, stout, incurved,
hooklike frontal spines. The dorsal plate is oval and narrowed
anteriorly. The ventral plate is very broadly oval and in its
posterior half projects beyond the dorsal plate, both laterally
and posteriorly. The surface markings consist of a transverse
ventral fold in front of the foot and two pairs of short ridges
directed outwards and backwards. The lateral sulci are deep,
especially so in the anterior half, where they produce the pe¬
culiar indentation shown on the ventral plate and terminating
rather abruptly, to reappear on the dorsal side; this accounts
for the distinct angle at the edges of *both plates. The posterior
segment is very broad and rounded, its limits ill-defined. The
coxal plates are large and semielliptic. The first foot joint is
oval and somewhat indistinct, the second joint robust and rhom¬
boid. The toe is long and stout, about one third of the total
length, slightly tapering and very slightly enlarged near the mid¬
dle; the claw is short, stout, acutely pointed and has two dis¬
tinct basal spicules.
Total length 175/x; length of dorsal plate 105/a, of ventral plate
118/a ; width of lorica 96/a ; width of anterior dorsal margin, less
exposed part of lateral sulci, 44/a, of ventral margin over spines
38/a; length of toe without claw 40/a; claw 11/a.
Monostyla stenroosi is not common ; Stenroos found it in Fin¬
land, Voronkov in collections made on the Shat-el-Arab, Murray
at Rio de Janeiro, Sachse at Trachenberg in Schlesien and Meiss¬
ner on the shores of the Amu-Darja. We have collected it at
Los Angeles, California, in small numbers ; it was fairly com¬
mon in material forwarded by Dr. N. Gist Gee from Soochow,
in Kiang-su, China.
396 Wisconsin Academy of Sciences, Arts and Letters.
MONOSTYIiA CORNUTA (Miiller).
Plate XL, figures 5, 6.
Trichoda cornuta Muller, Animalcula Infusoria, 1786, p. 208, pi.
30, figs. 1-3.
Lepadella cornuta Bory de St. Vincent, Diet. Class. Hist. Nat.,
vol. 9, 1826, p. 285. — Ehrenberg, Isis (Oken), vol. 26, 1833, col.
246.
Lepadella glumiformis Bory de St. Vincent, Encycl. Meth., Zooph.
(pt. 2), 1827, p. 484 = Trichoda cornuta renamed.
Notommata cornuta Ehrenberg, Isis (Oken), vol. 23, 1830, col. 767.
Monostyla cornuta Ehrenberg, Abh. Akad. Wiss. Berlin, 1830, p.
46; ibid., 1831, p. 130; Infusionsthierchen, 1838, p. 459, pi. 57,
fig. 4. — Perty, Zur Kenntn. kleinst. Lebensf., 1852, p. 41. —
Bartsch, Jahresh. Naturk. Wiirtemberg, vol. 26, 1870, p. 358. —
Eyferth, Mikr. Susswasserbew., 1877, p. 54; Einf. Lebensformen,
1878, p. 88, pi. 5, fig. 29; ibid., 1885, p. 113, pi. 7, fig. 29. — Eck¬
stein, Zeitschr. Wiss. Zook, vol. 39, 1883, p. 382, pi. 27, fig. 50. —
Blochmann, Mikr. Thierw. Siissw., 1886, p. 107. — Hudson and
Gosse, Rotifera, 1886, vol. 2, p. 98, pi. 25, fig. 1. — Bergendal,
Acta Univ. Lundensis, vol. 28, 1892, sect. 2, No. 4, p. 119. — Ter-
NETZ, Rotat. Umg. Bassels, 1892, p. 17. — Levander, Acta Soc.
Fauna et Flora Fennica, vol. 12, No. 3, 1895, p. 51, pi. 3, fig. 32a. —
Hempel, Bull. Illinois State Lab. Nat. Hist., vol. 5, 1898, p. 375. —
Jennings, Bull. U. S. Fish Comm., vol. 19 (for 1899), 1900, p. 92,
pi. 20, figs. 35, 36. — Voronkov, Trudy Gidr. Stants. Glubokom
Oz., vol. 2, 1907, pp. 110, 284. — Schlenker, Mitt. Geol. Abt.
Wurttemberg. Stat. Landesamt, No. 5, 1908, p. 249. — Von Hof-
STEN, Arkiv Zook, Stockholm, vol. 6, No. 1, 1909, p. 60. — Runn-
STROM, Zook Anz., vol. 34, 1909, p. 271. — Sachse, Siisswasserfau-
na Deutschlands, pt. 14, 1912, p. 178, figs. 352-354. — Mola, Ann.
Biol. Lac., vol. 6, 1913, p. 265. — Kozar, Zook Anz., vol. 44, 1914,
p. 420. — Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich, vol. 1,
1914, p. 33; Kosmos (Lwow), 1918-1919, p. 28. — Montet, Rev.
Suisse Zook, vol. 23, 1915, p. 336. — Rezvoi, Trudy Borodinsk,
Biol. Stants. Imp. Petrogradsk. Obshch. Estestvoisp., vol. 4, 1916,
p. 181. — Weber and Montet, Cat. Invert. Suisse, pt. 11, 1918, p.
191. — Olofsson, Zook Bidr. Uppsala, vol. 6, 1918, p. 594, fig.
55. — Harring, Rep. Canadian Arctic Exp. 1913-18, vol. 8, pt. E,
1921, p. 10. — Bryce, Journ. Quekett Micr. Club, ser. 2, vol. 14,
1922, pp. 313, 314.
Trichocerca cornuta Gravenhorst, Nova Acta Acad. Caes. Leop.-
Carok Nat. Cur., Bonn, vol. 16, 1832, p. 870.
Dicerratella cornuta Deshayes and Milne Edwards, in Lamarck,
Hist. Nat. Anim. sans Vert., ed. 2, vol. 1, 1835, p. 431; not Dicera-
tella triangularis Bory de St. Vincent 1826 (= Leucophra cor^
nuta Muller renamed).
Monostyla truncata Turner, Bulk Denison Univ., vol. 6, 1892, p. 62,
pk 1, fig. 11.
Harring (& Myers — Rotifer Fauna^ of Wisconsin. — III. 397
Monostyla robusta Stokes, Ann. Mag. Nat. Hist., ser. 6, vol. 18,
1896, p. 22, pi. 7, figs. 6-8. — Murray, Journ. Royal Micr. Soc.,
1913, p. 657 pi. 23, fig. 21.
Monostyla rotiindata Jakueski, Rozpr. Wiad. Muz. Dzieduszyckich,
vol. 1, 1914, p. 34, pi. 1, fig. 6.
The lorica is almost circular or very slightly elliptic. The
dorsal and ventral plates have a fairly deep, lunate anterior
sinus. The dorsal plate is circular in outline and without any
surface markings ; the angles of the anterior sinus are somewhat
obtuse and without spines ; a curved fold, starting at the bottom
of the dorsal sinus and joining the external edges some distance
from the anterior points, is formed when the head is completely
retracted. The ventral plate is as wide as the dorsal plate and
nearly identical in outline, very slightly narrower in the pos¬
terior half. The lateral sulci are very deep. The posterior seg¬
ment of the body is short, very broad and somewhat indistinct,
projecting very little beyond the dorsal plate; the coxal plates
are broad, bluntly pointed and not very prominent. The first foot
joint is moderately large and rather obscure, the second reni-
form and well marked; in front of the foot is a well marked
transverse fold, indicating the limits of the body proper and
the foot. The toe is parallel-sided, about one fourth of the total
length, and ends in a large claw with a median groove and two
distinct basal spicules; in some specimens the claw appears to
be double.
Total length 190/^ ; length of dorsal plate 134/a, of ventral plate
138ja; width of lorica 132^; width of anterior sinus 65^, depth
13ju ; length of toe without claw 45)U, ; claw 12fi.
Monostyla cornuta is abundant in weedy ponds everywhere in
the United States; there is very little reliable information on
its occurrence elsewhere, as existing descriptions are unsatis¬
factory and somewhat contradictory. We have followed Jen¬
nings in accepting this form as the animal identified by Ehren-
berg with Muller’s description; it also agrees fairly well with
the description and figures of Hudson and Gosse.
A diminutive form of this species is found in collections from
the Isthmus of Panama, which measures: total length 120/x;
length of lorica 85^; length of toe without claw 23/1, claw 9y.
With the exception of its small size and the relatively much
longer claw it agrees in other respects with the normal form,
which is abundant on the Isthmus.
398 Wisconsin Academy of Sciences, Arts and Letters.
MONOSTYLA SYLVATICA Harring
Plate XL, figures 3, 4.
Monostyla sylvatica Harking, Proc. U. S. Nat. Mus., vol. 46, 1913,
p. 399, pi. 35, figs. 1-3.
The lorica is subcircular in outline ; its width is equal to the
length. The anterior margin of the dorsal plate is very narrow
and concave ; the ventral margin is much wider and has a shal¬
low, V-shaped sinus with flaring sides; no anterior spines are
present. The dorsal plate is subcircular and truncate poste¬
riorly. The ventral plate is broadly oval and only four fifths
of the width of the dorsal plate. A well marked transverse fold
on the ventral plate, a short distance in front of the foot, is the
only surface marking present. There are no lateral sulci in the
contracted animals; the dorsal and ventral plates are joined
by a virtually fiat, unbroken membrane. The posterior segment
is small and rounded; it projects but little beyond the posterior
plate. The coxal plates are rather small and obtusely pointed.
The first foot joint is somewhat indistinct, the second short,
broad and reniform. The toe is very long and slender, about one
third of the total length, parallel-sided and slightly decurved.
The claw is short and acute ; it has a distinct median line, but
does not appear to be actually divided.
Total length 150/x ; length of dorsal plate 96/x, of ventral plate
100/A ; width of dorsal plate 100/z, of ventral plate 82/x ; width of
anterior dorsal margin 32|U, ventral margin 53/a; length of toe
without claw 44/a ; claw 8/a.
Monostyla sylvatica was quite common among mosses and he-
patics growing on rocks in the bed of a gently flowing stream
in the woods north of the Bureau of Standards, in Washington,
District of Columbia ; the little stream is now polluted and unin-
inhabitable for rotifers. No other localities are known for this
species.
MOlVOSTYIiA COPEIS Marring and Myers, new species.
Plate XLI, figures 1, 2.
The outline of the lorica is very broadly ovate; its width is
more than four fifths of the length. The anterior margins are
coincident and very slightly concave. The dorsal plate is sub¬
circular and rounded posteriorly. The ventral plate is nearly
parallel-sided anteriorly and rounded posteriorly; it is consid-
Harring (& Myers — Rotifer Fauna of Wisconsin. — III. 399
erably naiTOwer than the dorsal plate and its margins ill-defined
and wavy. Surface markings consist of a transverse ventral fold
in front of the foot and some short, curved folds on each side
of the foot. The lateral sulci are indistinct and shallow. The
posterior segment is fairly large and rounded; it projects
slightly beyond the dorsal plate. The coxal plates are small and
rounded posteriorly. The first foot joint is parallel-sided and
squarely truncate posteriorly, the second joint heart-shaped.
The toe is spindle-shaped and much enlarged in the middle,
ending in a stout, acute claw ; its length is slightly less than one
fourth of the total length.
Total length 130//-; length of dorsal plate 88//, of ventral plate
94//; width of dorsal plate 80//, of ventral plate 65//; width of
anterior dorsal margin 50//, of ventral margin 58// ; length of toe
without claw 33// ; claw 5//.
Monostyla copeis was collected by Dr. H. L. Shantz in Pros¬
pect Lake, on the slopes of Pikes Peak, Colorado, at an altitude
of 1830 meters, and by Dr. Paul Galtsofi in Lake Pepin, Wis¬
consin. The specimens from Colorado are slightly smaller and
the claw relatively longer ; the toe is also somewhat broader, but
there is no doubt of their specific identity.
MONOSTYLA RHOPALIJRA Harring and Myers, new species.
Plate XLI, figures 3, 4.
Monostyla closterocerca Jennings, Bull. Michigan Fish Comm. No.
3, 1894, p. 25, fig. 9. — Hempel, Bull. Illinois State Lab. Nat. Hist,
vol. 5, 1898, p. 376; not Monostyla closterocerca Schmarda,
The outline of the lorica is broadly oval; its width is four
fifths of the length. The anterior dorsal margin is very narrow
and has a rather shallow, V-shaped sinus, rounded posteriorly;
the ventral margin is crescentic; the external angles are mod¬
erately acute, both dorsally and ventrally. The dorsal plate is
broadly oval and rounded posteriorly. The ventral plate is oval
and slightly narrower than the dorsal; it has a transverse fold
in front of the foot. The lateral sulci are fairly deep. The
posterior segment is large and rounded and protrudes nearly
full length beyond the dorsal plate. The coxal plates are large
and obtusely pointed. The first foot joint is semi-elliptic; the
second joint is large and pyriform. The toe is long and stout,
nearly one third of the total length, fusiform and much en-
400 Wiscofisin Academy of Sciences, Arts and Letters.
larged in the middle; the claw is long, slender and acutely
pointed, with a median line or indistinct groove.
Total length ISOfx ; length of dorsal plate 115fi, of ventral plate
130/x; width of dorsal plate 98/^, of ventral plate 90 fi; width of
anterior dorsal margin 36/x, of ventral margin 50/^ ; length of toe
without claw 46/a; claw 8/a.
Monostyla rhopalura is locally common; we have collected it
around Atlantic City, New Jersey ; at Eagle E-iver, Vilas Coun¬
ty, Wisconsin, and on Mount Desert Island, Maine. Jennings
collected it in ponds on the shores of Lake Erie and in inland
lakes in Michigan.
MONOSTYliA PIDEIS Harrinj;^ and Myers, new species.
Plate XLI, figures 5, 6.
The outline of the lorica is nearly circular ; its width is equal
to the length. The dorsal plate is subcircular, with very slight,
lateral Indentations near the posterior end. The ventral plate
is somewhat narrower than the dorsal and obtusely pointed pos¬
teriorly. The anterior dorsal margin is very narrow and cuspi¬
date, with a deep, V-shaped sinus, which has, for the protrusion
of the dorsal antenna, a large, median notch, squarely truncate
posteriorly. The ventral sinus is deeply lunate. Surface mark¬
ings are limited to the usual transverse fold in front of the foot
and two dorsal transverse lines, one at the bottom of the anterior
sinus and another near the end of the dorsal plate. The lateral
sulci are deep. The posterior segment is short and does not
project beyond the dorsal plate. The coxal plates are minute
and scale-like ; they are smaller than in any other species of this
group and do not extend beyond the first foot joint. The foot
is unusually robust and both joints very broad and short. The
toe is long, about one third of the total length, extremely broad
and parallel-sided, narrowed at the base and rounded posterior¬
ly; it is compressed dorso-ventrally and oval in cross section.
The claw is short and triangular.
Total length 160/a; length of dorsal plate 122/a, of ventral plate
122/a; width of dorsal plate 108/a, of ventral plate 100/a; width
of anterior margin 32/a ; length of toe without claw 45/a ; claw 6/a.
Monostyla pideis was collected among submerged mosses in
Bubble Pond, Mount Desert Island, Maine. Its nearest rela¬
tives are probably M. cornuta and M. rhopalura.
Harring (& Myers — Rotifer Fauna of Wisconsin. — 111. 401
MONOSTYL.A SCUTATA Harring: and Myers, new species
Plate XL, figures 1, 2.
The outline of the lorica is subcircular; its width is nearly
equal to the length. The anterior margins are coincident and
very slightly concave, the dorsal margin slightly narrower than
the ventral ; anterior spines are not present. The dorsal plate is
subcircular and rounded posteriorly; the ventral plate is par¬
allel-sided anteriorly and rounded posteriorly. Surface mark¬
ings are limited to a transverse fold on the ventral plate, in
front of the foot. The lateral sulci are rather shallow and indis¬
tinct. The posterior segment is broad and rounded and pro¬
jects somewhat beyond the dorsal plate. The coxal plates are
fairly large and obtusely rounded posteriorly. The first foot
joint is pyriform and rounded posteriorly, the second joint short
and very broadly elliptic. The toe is long and fairly stout,
about one third of the total length, parallel-sided for about three
fourths of its length and tapering to the long, slender and acute
claw.
Total length 100/x ; length of dorsal plate 62/>i, of ventral plate
70/x; width of dorsal plate 65/x, of ventral plate 50^.; width of
anterior dorsal margin 44/a, ventral margin 48/a; length of toe
without claw 28/a ; claw 5/a.
Monostyla scutata was collected in small numbers in Ottman
Lake, near Waupaca, Wisconsin. No other localities are known
for this species.
MONOSTYLA PYGMAEA Daday
Plate XLII, figures 1, 2.
Monostyla pygmaea Daday, Math. Termesz. Ertes., vol. 15, 1897,
p. 139, fig. 7; Termesz. Fiizetek, vol. 24, 1901, p. 21, fig. 4. — Har¬
king and Myers, Trans. Wisconsin Acad. Sci., vol. 20, 1922, p.
557. — Hauer, Zool. Anz., vol. 61, 1924, p. 149.
Monostyla turbo Murray, Journ. Royal Micr. Soc., 1913, p. 558, pi.
22, fig. 11.
The outline of the lorica is broadly ovate; its width is about
three fourths of the length. The anterior dorsal margin is
straight, the ventral margin convex with a small, shallow,
rounded median sinus; this species shares with M. ohtusa the
peculiarity of having the anterior dorsal margin wider than
the ventral. The dorsal plate is broadly ovate and rounded pos-
26
402 Wisconsin Academy of Sciences^ Arts and Letters.
teriorly. The ventral plate is ovate and somewhat narrower than
the dorsal plate, especially in front. There is a fairly prominent
transverse ventral fold above the foot. The lateral sulci are
fairly deep. The posterior segment is large and rounded; it
projects somewhat beyond the dorsal plate. The coxal plates
are small and obtusely pointed posteriorly. The first foot joint
is somewhat indistinct and widest anteriorly, the second joint
roughly hexagonal and slightly inside the posterior edge of the
lorica. The toe is long and straight, about one fourth of the
total length, and ends in a small, acute claw with median divid¬
ing line and without basal spicules.
Total length 125/x ; length of dorsal plate 80/a, of ventral plate
88/a; width of dorsal plate 72/a, of ventral plate 65/a; width of
anterior dorsal margin 58/a, of ventral margin 48/a ; length of toe
without claw 37/a ; claw 5/a.
Monostyla pygmaea is common in wet sphagnum; we have
collected it around Washington, District of Columbia; at Atlan¬
tic City, New Jersey; in Polk County, Florida; in Oneida and
Vilas Counties, Wisconsin and on Mount Desert Island, Maine.
Hauer reports it from the Schwarzwald. Murray found it among
Rousselet^s collections from Clare Island, Ireland.
MONOSTYLA ORNATA Harrin^ and Myers, new species
Plate XXXVI, figures 1, 2.
The lorica is broadly ovate ; its width is about three fourths
of the length. The anterior dorsal margin is slightly convex;
the ventral has a small median sinus, broadly rounded at the
bottom and with convex sides. The dorsal plate is broadly oval
and rounded posteriorly. The ventral plate is nearly parallel¬
sided for half its length and very bluntly triangular posteriorly.
The surface markings are unique and must be studied from the
figures; they consist of a system of raised ridges with lateral
buttresses or branches at close intervals, but the usual pattern
of facets is not traceable. The lateral sulci are shallow and the
margins of the ventral plate somewhat undulate. The posterior
segment is narrow and rounded posteriorly; it projects slightly
beyond the dorsal plate. The coxal plates are very small and
obtusely pointed. The first foot joint is large and conical, the
second subcircular, slightly reduced in front of the toe and
well within the posterior segment. The toe is long and slender,
Earring d: Myers — Rotifer Fauna of Wisconsin.- — III. 403
about one third of the total length, straight and parallel-sided
with a distinct annular constriction near the posterior end ; the
claw is long, fairly stout and acutely pointed.
Total length 130/x ; length of dorsal plate 82/a, of ventral plate
87/x; width of dorsal plate 68/x, of ventral plate 6O/1,; width of
anterior margin 55fjL ; length of toe without claw 38/t ; claw Ifi.
Monostyla ornata was collected in large numbers in a swamp
at Oceanville, near Atlantic City, New Jersey. No other local¬
ities are known for this very interesting species.
MONOSTYLA OBTYSA Murray
Plate XLII, figures 5, 6.
Monostyla ohtiisa Murray, Journ. Royal Micr. Soc., 1913, p. 357, pi.
15, fig. 37.- — -Harring, Proc, U. S. Nat. Mus., voL 47, 1914, p.
548. — Hauer, ZooL Anz., vol. 61, 1924, p. 147.
? Monostyla lunaris aperta Steinecke, Schriften Phys.-okon. Ges.
Konigsberg i Pr., vol. 57, 1916, p. 89, fig, 2, c, d; vol. 64, 1924,
p. 41.
The outline of the lorica is broadly oval; its width is three
fourths of the length. The anterior margins of the dorsal and
ventral plates are coincident and straight; at the external an¬
gles are two minute frontal spines. The dorsal plate is broadly
oval and rounded posteriorly. The ventral plate is considerably
narrower than the dorsal plate and broadly oval ; it is somewhat
flexible and its margins rather indefinite. The lorica is without
any surface markings. The lateral sulci are shallow and ill-
defined. The posterior segment is small and rounded, projecting
very little beyond the dorsal plate. The coxal plates are mod¬
erately large and obtusely rounded posteriorly. The first foot
joint is parallel-sided and indistinct, the second joint fairly
large and subsquare, projecting slightly beyond the lorica. The
toe is fairly long and straight, about one fourth of the total
length, spindle-shaped, narrow at the base and slightly en¬
larged towards the posterior end ; the claw is fairly long, acutely
pointed and has a median dividing line without being double;
two basal spicules are present.
Total length 115/x; length of dorsal plate IQy, of ventral plate
80/x; width of dorsal plate 70/i,, of ventral plate 58/x; width of
anterior dorsal margin 58/a, of ventral margin 48/a; length of
toe without claw 32/a; claw 7/a.
404 Wisconsin Academy of Sciences^ Arts and Letters.
Monosiyla ohtusa was collected by Murray at Rio de Janeiro.
It seems to be rare ; we have found it common in collections from
the Panama Canal Zone, and a few specimens in material from
Audubon Park, in New Orleans, Louisiana, collected by Dr.
Birge and Mr. Juday in 1903 and also in Sphagnum forwarded
from Hubbardstown, Massachusetts, by Mrs. A. C. Clarke.
Hauer reports it common in sphagnum bogs in the Schwarz-
wald and apparently this is the animal found by Steinecke in
the Zehlaubruch near Konigsberg.
MONOSTYIjA VASTITA Harring and Myers, new species
Plate XLII, figures 3, 4.
The outline of the lorica is very broadly reversed-ovate ; its
width is but little less than the length. The anterior mar¬
gins are nearly coincident; the dorsal is very slightly concave
and the ventral straight. The dorsal plate is very broadly oval
and rounded posteriorly; the ventral plate is much narrower
and oval in outline; it has a transverse and two longitudinal
folds. The lateral sulci are shallow and ill-defined. The pos¬
terior segment is large and rounded; it projects with its entire
length beyond the dorsal plate. The coxal plates are large
and obtusely pointed. The first foot joint is very long, con¬
stricted near the posterior; lobate end; the second joint is ro¬
bust, very broad and roughly pentagonal in outline. The toe
is long and slender, about one third of the total length, parallel¬
sided and obtuse posteriorly ; the claw is long, slender and acute¬
ly pointed with a median mucus groove.
Total length 130/x; length of dorsal plate 78/x, of ventral plate
90;ii; width of dorsal plate 78/a, of ventral plate 62/a; width of
anterior dorsal margin 65/a, of ventral margin 50/a; length of
toe without claw 35/a; claw 7/a.
Monostyla vastita was collected in Round Pond, on Mount
Desert Island, Maine; this is the only location known for this
species. It resembles M. ohtusa in having the dorsal anterior
margin wider than the ventral, but differs in nearly every other
respect, and there can be no confusion between these two
species.
Earring d' 3Iyers — Rotifer Fauna of Wisconsin. — III. 405
MOIVOSTYIiA TETHIS Marring and Myers, new species
Plate XXXVIII, figures 1, 2.
The outline of the lorica is very broadly oval; its width is
but little less than the length. The anterior margins of the dor¬
sal and ventral plates are coincident and straight. The dorsal
plate is very broadly ovate and narrowly truncate posteriorly.
The ventral plate is as wide as the dorsal and very broadly oval.
The dorsal facetting is of an unusual pattern and very regular;
each facet is bounded by double longitudinal lines. The sur¬
face markings of the ventral plate are also somewhat aberrant,
as shown in figure 2. The lateral sulci are moderately deep.
The posterior segment is very short and broadly rounded ; it pro¬
jects very slightly beyond the -dorsal plate. The coxal plates
are large and nearly semicircular. The first foot joint is rudi¬
mentary and indistinct, the second joint large and subcircular.
The toe is long, nearly one third of the total length, straight
and very slightly tapering ; the claw is fairly long, slender and
acutely pointed.
Total length 100/x; length of dorsal plate 70/a, of ventral plate
74ju; width of lorica 62 fx; width of anterior dorsal margin 42/.t,
of ventral margin 53/>t; length of toe without claw 24/^; claw
6fi.
Monostyla tethis was collected in small numbers among
sphagnum from Aunt Bettys Pond, Mount Desert Island, Maine.
No other localities for this species are known.
MONOSTYLA RUGOSA Marring
Plate XLIII, figures 3, 4.
Monostyla rugosa Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
548, pi. 24, figs. 4-6. — Fadeev, Russk. Gidr. Zhurn., vol. 3, No.
3-5, 1924, p. 6.
The outline of the lorica is very broadly ovate; its width is
slightly greater than the length. The anterior margins are near¬
ly coincident, the dorsal slightly convex and the ventral straight.
The dorsal plate is very broadly ovate and narrowly truncate
posteriorly ; the facetting resembles closely the usual pattern ex¬
cept in the anterior row, which is irregular. The ventral plate
is fiexible and its margins somewhat indefinite, but it is roughly
subsquare and rounded posteriorly; the pattern of the surface
markings is fairly complex. The lateral sulci are indistinct
406 Wisconsin Academy of Sciences, Arts and Letters.
and shallow. The posterior segment is very short and broad;
it is almost completely covered by the dorsal plate. The coxal
plates are fairly large and semicircular. The toe is long, about
one third of the total length, robust and parallel-sided ; the long
claw is distinctly double, but the two parts are rarely separated.
Total length 84ju; length of dorsal plate 54ju,, of ventral plate
57/4 ; width of dorsal plate 62/4, of ventral plate 56/4; width of
anterior dorsal margin 34/4, of ventral margin 45/4; length of
toe without claw 21/4 ; claw 6/4.
Monostyla rugosa is common in the Panama Canal Zone; we
have not found it in the United States. Fadeev records it from
Kharkov, Russia.
MONOSTYLA ELACHIS Marring and Myers, new species
Plate XLIII, figures 1, 2.
The outline of the lorica is subcircular; its width is nearly
equal to the length. The anterior dorsal margin is slightly con¬
vex; the ventral margin has a shallow median concavity and
is convex towards the external angles; no frontal spines are
present. The dorsal plate is subcircular and rounded poste¬
riorly. The ventral plate is nearly parallel-sided anteriorly and
rounded posteriorly; it is narrower than the dorsal plate. The
anterior row of facets on the dorsal plate is very regular; the
rest of the facetting is of the usual pattern; the ventral plate
has a fairly prominent transverse fold in front of the foot. The
lateral sulci are deep. The posterior segment is moderately large
and rounded; it projects somewhat beyond the dorsal plate.
The coxal plates are small and obtusely rounded posteriorly.
The first foot joint is short and rather indistinct, the second
joint is large and heartshaped. The toe is fairly long, more
than one fourth of the total length, straight and parallel-sided,
terminating in a short, acutely pointed claw.
Total length 90/4; length of dorsal plate 62/4, of ventral plate
68/4; width of dorsal plate 62/4, of ventral plate 54/4; width of
anterior dorsal margin 40/4, of ventral margin 50/4 ; toe without
claw 20/4 ; claw 4/4.
Monostyla elachis is common in weedy ponds in the United
States; whether it occurs elsewhere is as yet unknown. It has
considerable resemblance to other small species of the genus,
but is one of the few with dorsal facetting, which, if not very
prominent, is at least very constant.
Earring & Myers — Rotifer Fauna of Wisconsin. — III. 407
MONOSTYIiA FURCATA Murray
Plate XLIII, figures 5, 6.
Monostyla furcata Murray, Journ. Royal Micr. Soc., 1913, p. 358,
pi. 15, fig. 40.— Harking, Proc. U. S. Nat. Mus., vol. 47, 1914, p
548.— Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich, vol. 1, 1914,
p. 35. — Fadeev, Russk. Gidr. Zhurn., vol. 3, No. 3-5, 1924, p. 6.
The outline of the lorica is broadly ovate or subcircular; its
width is but little less than the length, and the integument is
semi-flexible. The anterior margins of the dorsal and ventral
plates are wide, straight and coincident. The dorsal plate is
subcircular and without posterior truncation. The ventral plate
is roughly parallel-sided anteriorly and rounded posteriorly.
Surface markings are limited to a transverse fold on the ventral
plate in front of the foot. The lateral sulci are fairly deep,
the posterior segment is rounded and almost completely cov¬
ered by the dorsal plate. The coxal plates are small and round¬
ed posteriorly. The first foot joint is parallel-sided and some¬
what indistinct, the second joint nearly spherical. The toe is
short and stout, about one fourth of the total length, straight
and parallel-sided, terminating in two very distinct claws, im¬
mobile and separated by a V-shaped notch.
Total length 100/x; length of dorsal plate 75/x, of ventral plate
78y; width of dorsal plate 70/x, of ventral plate 60y; width of
anterior dorsal margin 43/x, of ventral margin 58/a; length of
toe without claws 22/a; claws 5/a.
Monostyla furcata is probably widely distributed, but has
no doubt often been confused with other small species of the
genus. Murray found it at Rio de Janeiro; it was common in
the collections of the Panama Biological Survey and we have
found it at Washington, District of Columbia; around Atlantic
City, New Jersey; in Oneida and Vilas Counties, Wisconsin,
and on Mount Desert Island, Maine, as well as in collections
made by Mr. Juday at Puerto Barrios, Guatemala.
MONOSTYLA PUNCTATA Murray
Plate XLIV, figures 3, 4.
Monostyla punctata Murray, Journ. Royal Micr. Soc., 1913, p. 355,
pi. 15, fig. 36.
The outline of the lorica is very broadly ovate ; its width is
nearly equal to the length. The anterior dorsal margin is very
slightly convex and very much narrower than the ventral mar-
408 Wisconsin Academy of Sciences^ Arts and Letters.
gin, so that a part of the integument belonging to the lateral
sulcus is prominently in view from the dorsal side and equalizes
the difference in width between the two plates ; the ventral mar¬
gin is slightly concave opposite the dorsal plate and from this
point recedes at an obtuse angle to the external edge. The dor¬
sal plate is very broadly ovate and rounded posteriorly. The
ventral plate is considerably narrower than the dorsal, widest
anteriorly and tapers gradually towards the rounded posterior
end ; it is quite flexible and the edges somewhat ill-defined. The
lateral sulci are shallow except at the front, where they make up
for the difference in width between the dorsal and ventral plates.
The posterior segment is small and rounded; it projects but lit¬
tle beyond the dorsal plate. The coxal plates are small and
obtusely pointed. The first foot joint is very large, but some¬
what indistinct; the second joint is large and rounded. The
toe is long and slender, more than one fourth of the total
length, very slightly enlarged near mid-length and blunted pos¬
teriorly; the claw is fairly long, slender and acutely pointed,
with a median mucus groove.
Total length 110/a; length of dorsal plate 76/a, of ventral plate
80/a; width of dorsal plate 70/a, of ventral plate 55/a; width of
anterior dorsal margin 36/a, of ventral margin 58/a ; length of toe
without claw 24/a; claw 6/a.
Monostyla punctata was described by Murray from material
collected in a brackish lagoon at Rio de Janeiro. We find it abund¬
ant in salt ponds and tide pools around Atlantic City, New Jer¬
sey. The structure as described by Murray seems very com¬
plex; this is evidently due to poor and incompletely retracted
material ; there is nothing especially remarkable about M. punc¬
tata except the great difference between the dorsal and ventral
anterior margins.
MONOSTYLA CLOSTEROCERCA Sohmarda
Plate XLIV, figures 5, 6.
Monostyla closterocerca Schmarda, Neue wirbellose Thiere, 1859,
vol. 1, p. 59, pi. 14, fig. 125. — Murray, Journ Royal Micr. Soc.,
1913, p. 357, pi. 15, fig. 39. — Marring, Proc. U. S. Nat. Mus., vol.
47, 1914, p. 548; Rep. Canadian Arctic Exp. 1913-1918, vol. 8,
pt. E, 1921, p. 10. — Jakubski, Rozpr. Wiad. Muz. Dzieduszyckich,
vol. 1, 1914, p. 32, pi. 1, fig. 4; Kosmos (Lwow), 1918-1919, p. 26.
? Monostyla truncata Turner, Bull. Denison Univ., vol. 6. 1892,
p. 62, pi. 1, fig. 11.
Harring (& Myers — Rotifer Famm of Wisconsin. — 111. 409
? Monostyla cornuta Olofsson, Zool. Bidr. Uppsala, vol. 6, 1918,
p. 594, fig. 55. — Hauer, Mitt. Geogr. Ges. u. Naturhist. Mus. Lii-
beck, ser. 2, No. 30, 1925, p. 170, fig. 8.
The outline of the lorica is subcircular ; its width is virtually
equal to the length. The anterior dorsal and ventral margins
are coincident and form a shallow, broadly V-shaped sinus with
widely flaring, convex sides. The dorsal plate is nearly circular,
rounded posteriorly and its anterior edges curving inwards with¬
out actually reaching the anterior margin. The ventral plate
is very broadly oval and considerably narrower than the dorsal
plate. The surface markings are a faint dorsal fold, originat¬
ing near the apex of the anterior sinus, and a transverse ven¬
tral fold in front of the foot. The lateral sulci are shallow. The
posterior segment is very broad and semicircular, and projects
slightly beyond the dorsal plate. The coxal plates are large and
rounded posteriorly, terminating slightly beyond the first foot
joint; this is indistinct, rather narrow and parallel-sided, the
second joint large and varying from subcircular to subsquare.
The toe is long, a little less than one third of the total length,
parallel-sided for half its length and tapering to a slender, acute
point.
Total length, llO^a ; length of dorsal plate 12^, of ventral plate
78/a; width of dorsal plate 75/a, of ventral plate 5/a; width
of anterior margin 44/a; length of toe 33/a.
Monostyla closterocerca is abundant everywhere in weedy
ponds. Murray ’s figure shows the toe reduced at the base ; this
is an error ; it is always parallel-sided.
MONOSTYLA PYRIFORMIS Daday.
Plate XLV, figures 1, 2.
Monostyla pyriformis Daday, Math. Termesz. Ertes., vol. 23, 1905,
p. 330; Zoologica, pt. 44, 1905, p. 112, pi. 7, fig. 16. — Harking,
Proc. U. S. Nat. Mus., vol. 47, 1914, p. 547.
Monostyla truncata Murray, Journ. Royal Micr. Soc., 1913, p. 358,
pi. 15, fig. 38; not Monostyla truncata Turner.
The outline of the lorica is very broadly ovate; its width is
four fifths of the length. The anterior margins are coincident,
straight for the greater part of their length and strongly round¬
ed towards the edges of the lorica. The dorsal plate is very
broadly ovate and rounded posteriorly. The ventral plate is ovate
and considerably narrower than the dorsal. The lateral sulci are
410 Wisconsin Academy of Sciences^ Arts and Letters.
rudimentary. The posterior segment is small and rounded; it
projects somewhat beyond the dorsal plate. The coxal plates are
small and obtusely pointed. The first foot joint is semi-elliptic ;
the second joint is robust and subsquare. The toe is very long,
about one third of the total length, parallel-sided for one half
its length and tapering gradually to a slender, bristle-like point.
Total length 80/x; length of dorsal plate 55^, of ventral plate
58/x; width of dorsal plate 48/a, of ventral plate 40/a; length of
toe 24/a.
Monostyla pyriformis is not rare in wet sphagnum. We have
used Daday’s name for this species, as partly contracted speci¬
mens agree perfectly with his figure. Turner’s M. truncata is
figured with a straight, acute angled anterior margin, which this
species never has; his animal may have been any one of the
smaller Monostylas; the details given are insufficient to decide
which one and his figure does not inspire much confidence in its
accuracy.
MONOSTYLA SUBULATA Harring and Myers, new species.
Plate XLV, figures 3, 4.
The outline of the lorica is very broadly ovate; its width is
virtually equal to the length. The anterior margins of the dor¬
sal and ventral plates are coincident and straight; two minute
indentations are usually present near the external angles. The
dorsal plate is very broadly ovate and rounded posteriorly. The
ventral plate is reduced considerably in width anteriorly and the
posterior portion is consequently nearly circular in outline ; it is
much narrower than the dorsal plate. No surface markings are
present on either dorsal or ventral plates. The lateral sulci are
somewhat indistinct. The posterior segment is very small and
rounded; it projects considerably beyond the dorsal plate. The
coxal plates are rather small and obtusely pointed. The first
foot joint is semi-elliptic; the second joint is heart-shaped and
projects about two thirds of its length beyond the posterior seg¬
ment. The toe is long and stout, about one fourth of the total
length, tapering and has a median line or indistinct groove.
Total length 100/a ; length of dorsal plate 64/a, of ventral plate
68/a; width of dorsal plate 65/4,, of ventral plate 52/i; width of
anterior margin 50/a ; length of toe without claw 16/a ; claw 10/a.
Harring & Myers — Rotifer Fauna of Wisconsin. — 111. 411
Monostyla suhulata is fairly common in wet sphagnum and
is not infrequently found in weedy ponds in the United States ;
we have found it in sphagnum from Epping Forest, sent to us
by Mr. David Bryce.
MONOSTYIjA OPIAS Harring and Myers, new species.
Plate XLV, figures 5, 6.
The outline of the lorica is very broadly ovate; its width is
five sixths of the length. The anterior margins are coincident
and straight; at the external angles are two very small, acute
frontal spines. The dorsal plate is very broadly ovate and
rounded posteriorly. The ventral plate is broadly ovate and
slightly narrower than the dorsal plate ; in front of the foot is a
fairly conspicuous transverse fold. The lateral sulci are mod¬
erately deep. The posterior segment is fairly large and rounded ;
it projects somewhat beyond the dorsal plate. The coxal plates
are small and obtusely pointed. The first foot joint is small
and indistinct ; the second is large and rounded. The toe is long,
more than one fourth of the total length, and tapers gradually
to a slender, bristle-like point.
Total length 100/x ; length of dorsal plate 66/x, of ventral plate
72(jl; width of dorsal plate 60/a, of ventral plate 55/a; width of
anterior spines 45/a; length of toe 28/a.
Monostyla opias is rare; we have found it in wet sphagnum
from Hyattsville, near Washington, District of Columbia, and
from Squirrel Lake, about 15 miles west of Minocqua, Oneida
County, Wisconsin ; only a few specimens were collected at each
station.
MONOSTYLA MONOSTYLA (Daday).
Plate XLVI, figures 1, 2.
Diarthra monostyla Daday, Math. Termesz. Ertes., vol. 15, 1897,
p. 143, fig. 10; Termesz. Fuzetek, vol. 24, 1901, p. 26, fig. 9;
Zoologica, pt. 44, 1905, p. 127, pi. 7, fig. 17.
Monostyla monostyla Harring, Proc. U. S. Nat. Mus., vol. 46, 1913,
p. 390. — Harring and Myers, Trans. Wisconsin Acad. Sci., vol.
20, 1922, p. 537. — Idelson, Trudy Kosinsk. Biol. Stants., pt. 2,
1925, p. 72, fig. 1.
Monostyla spinifera Idelson, Russk. Gidr. Zhurn., vol. 3, 1924,
p. 224, text fig.
The lorica is oval in outline; the anterior margins of dorsal
and ventral plates are straight and coincident; at the external
412 Wisconsin Academy of Sciences, Arts and Letters.
angles are two very small spines. The dorsal plate is very slight¬
ly narrower than the ventral and rounded posteriorly; it is very
flexible and without facetting. The ventral plate is evenly
rounded posteriorly and without evident markings; the coxal
plates are small and rounded posteriorly. The lateral sulci are
fairly deep ; hinged to the margin of the dorsal plate is a curved
spine, which may be swung outward and folded back into the
lateral sulcus. The process of withdrawing the spine from the
position shown in the figure is accomplished by a rotation down¬
wards and inwards through an angle of 180 degrees; when the
animal is swimming, there is no sign of the spine, concealed in
the lateral sulcus with its ‘‘posterior” edge turned outwards
and conforming to the outer edge of the dorsal plate. The first
foot joint is large, the second small and inverted pyriform. The
toe is about one fourth the total length, parallel-sided for one
half its length and ending in a conical point without claw.
Total length 90/x; length of dorsal plate 58/>i, of ventral plate
65 ju; width of dorsal plate 47/a, of ventral plate 50/a; anterior
points 38/a; toe 25/a.
Monostyla monostyla was first found by Daday in material
from New Guinea and later in South American collections. We
have found this species in numerous localities in the United
States: at Washington; around Atlantic City, New Jersey; in
Polk county, Florida ; numerous places in Wisconsin and on Mt.
Desert Island, Maine. It seems confined to wet sphagnum.
MONOSTYLA ARCUATA Bryce.
Plate XLVII, figures 3, 4.
Monostyla arcuata Bryce, Science Gossip, vol. 27, 1891, p. 206, text
fig. — Murray, Journ. Royal Micr. Soc., 1913, p. 360, pi. 15, fig.
42. — Hauer, Zool. Anz., vol. 61, 1924, p. 148.
The outline of the lorica is broadly oval; its width is about
four fifths of the length. The anterior dorsal margin is slight¬
ly concave; the ventral margin has a shallow median sinus
flanked by two straight sections. The dorsal plate is broadly
oval and rounded posteriorly. The ventral plate is elongate
oval and considerably narrower than the dorsal plate; it has a
transverse ventral fold in front of the foot and two longitudinal
folds. The lateral sulci are fairly deep, especially near the
anterior margin, where they compensate the difference in width
between the dorsal and ventral plate. The posterior segment is
Harring d; Myers — Rotifer Fauna of Wisconsin. — 111. 413
large and rounded; it projects slightly beyond the dorsal plate.
The coxal plates are small and obtusely pointed posteriorly.
The first foot joint is small and nearly parallel-sided ; the second
joint is large and rounded. The toe is long and fairly stout,
more than one fourth of the total length, nearly parallel-sided
for half its length and tapering gradually to a moderately acute
point. In some specimens the toe is very slightly broader in the
middle than at the base.
Total length 90/z; length of dorsal plate 60/x, of ventral plate
68/a; width of dorsal plate 54/a, of ventral plate 44/a; width of
anterior dorsal margin 24/a, of ventral margin 36/a ; length of toe
24/a.
Monostyla arcuata is fairly common in wet sphagnum. It has
a certain resemblance to M. closterocerca, but differs consider¬
ably in details, as well as in general proportions.
MOXOSTi^A DECIPIEXS Murray
Plate XLVII, figures 5, 6.
Monostyla decipiens Murray, Journ. Royal Micr. Soc., 1913, p. 360,
pi. 15, fig. 43. — Harring, Proc. U. S. Nat. Mus., vol. 47, 1914, p.
548.
The outline of the lorica is broadly ovate; its width is about
four fifths of the length. The anterior margins are coincident
and the sinus very deep and V-shaped, rounded posteriorly and
wffh a slight cusp opposite the incurved edges of the dorsal
plate; no frontal spines are present, but two triangular, acute
cusps are formed between the anterior sinus and the edges of the
lorica. The dorsal plate is broadly ovate and rounded posterior¬
ly. The ventral plate is ovate and slightly narrower than the
dorsal plate ; the lorica is without surface markings. The
lateral sulci are deep, especially at the anterior margin; the
dorsal plate terminates opposite the cusps on the sides of the
sinus and a large, triangular area belonging to the sulcus com¬
pletes it anteriorly. The posterior segment is rather small and
very obtusely pointed; it projects considerably beyond the
dorsal plate. The first foot joint is narrowly semi-elliptic and
somewhat indistinct; the second joint is broadly ovate and
widest posteriorly. The toe is long and slender, nearly one third
of the total length, parallel-sided for half its length and taper¬
ing gradually to an acute point; in some specimens the toe is
very slightly wider at mid-length than it is at the base.
414 Wisconsin Academy of Sciences, Arts and Letters.
Total length 175/a; length of dorsal plate 116/a, of ventral plate
128/a; width of dorsal plate 98/a, of ventral plate 78/a; width of
anterior cusps 44/a; length of toe 48/a.
Monostyla decipiens was collected by Murray at Rio de
Janeiro; our specimens are from the Panama Canal Zone and
uniformly larger than Murray’s material. This species has a
superficial resemblance to M. hamata, but differs in nearly every
detail and, as it is quite rare, there is little danger of confusing
the two species.
MONOSTYLA HAMATA Stokes.
Plate XLVII, figures I, 2.
Monostyla hamata Stokes, Ann. Mag. Nat. Hist., ser. 6, vol. 18,
1896, p. 21, pi. 7, figs. 6-8. — Jennings, Bull. U. S. Fish Comm.,
vol. 19 (for 1899), 1900, p. 94, pi. 22, figs. 42-44.^ — Voronkov,
Trudy Gidr. Slants. Glubokom Oz., vol. 2, 1907, pp. 110, 284, pi.
6, figs. 2-5. — Runnstrom, Zool. Anz., vol. 34, 1909, p. 271, fig.
3. — Daday, Zoologica, pt. 59, 1910, p. 84. — Sachse, Susswasser-
fauna Deutschlands, pt. 14, 1912, p. 180, figs. 351, 357. — Murray,
Journ. Royal Micr. Soc., 1913, pp. 359, 458, pi. 15, fig. 41. —
Kozar, Zool. Anz,, vol. 44, 1914, p. 421. — Harring, Proc. U. S.
Nat. Mus., vol. 47, 1914, p. 548; Rep. Canadian Arctic Exp. 1913-
18, vol. 8, pt. E, 1921, p. 10. — Jakubski, Rospr. Wiad. Muz. Dzie-
duszyckich, vol. 1, 1914, p. 33; Kosmos (Lwow), 1918-1919, p.
10. — Montet, Rev. Suisse Zool., vol. 23, 1915, p. 336. — Rezvoi,
Trudy Borodinsk. Biol. Slants. Imp. Petrogradsk. Obshch. Es-
testvoisp., vol. 4, 1916, p. 181, pi. 1, figs. 13, 14. — Weber and
Montet, Cat. Invert. Suisse, pt. 11, 1918, p. 192. — Chugunov,
Raboti Volzhskoi Biol. Slants., vol. 6, 1921, p. 116.
The outline of the lorica is elongate oval; its width is about
two thirds of the length. The anterior margins are not coinci¬
dent ; the dorsal margin is very narrow and deeply lunate ; the
ventral margin has a very deep, V-shaped sinus, rounded
posteriorly and with two prominent cusps near the middle of
the sides. No frontal spines are present, but two acute-angled
cusps are formed between the external edges of the ventral plate
and anterior sinus. The dorsal plate is oval and rounded
posteriorly ; the dorsal facetting is well marked and very regular.
The ventral plate is elongate oval and slightly narrower than
the dorsal; its surface markings consist of a transverse fold in
front of the foot and a few longitudinal folds. The lateral sulci
are fairly deep, especially so at the front, where the difference
in the widths of the anterior margins is compensated by the ex¬
posure of a large, triangular area of the sulcus. The posterior
Harring (& Myers — Rotifer Fauna of Wisconsin. — III. 415
segment is large and very obtusely pointed; it projects some¬
what beyond the dorsal plate. The coxal plates are small and
obtusely pointed posteriorly. The first foot joint is small, oval
and distinct, the second joint large and subsquare. The foot
is long and slender, more than one fourth of the total length,
parallel-sided for half its length and tapers gradually to an
acute point without claw.
Total length 120/x; length of dorsal plate 78/x, of ventral plate
92fi; width of dorsal plate 58/x, of ventral plate dSfi; width of
anterior dorsal margin 18/x, of ventral margin SOjx; length of
toe 33/x.
Monostyla hamata is common in weedy ponds everywhere.
MONOSTYL.A BATILLIFER Murray.
Plate XLVI, figures 3, 4.
Monostyla batillifer Murray, Journ Royal Micr. Soc., 1913, p. 458,
pi. 19, fig. 2.
The outline of the lorica is elongate ovate; its width is about
two thirds of the length. The anterior margins of the dorsal
and ventral plates are not coincident; the margin of the dorsal
plate is deeply lunate and very narrow; the ventral margin has
a very deep, obtusely V-shaped sinus with two very large,
lateral, almost spine-like cusps. The dorsal plate is ovate and
rounded posteriorly. The ventral plate is oval, constricted
anteriorly and somewhat narrower than the dorsal plate; its
surface markings consist of a transverse fold in front of the foot
and two longitudinal folds. The lateral sulci are fairly deep ;
the difference in the width of the anterior margins is made up
by exposure of a triangular portion of the sulcus. The posterior
segment is small, narrow, slightly undulate posteriorly and has
two laterally projecting, acutely triangular cusps; the segment
is only partly covered by the dorsal plate. The coxal plates are
small and rounded posteriorly. The first foot joint is small and
slightly conical, the second joint large, transversely oval and
some distance from the posterior margin. The toe is fairly long,
about one fourth of the total length, parallel-sided for nearly
three fourths of its length, conical posteriorly and ends in a
bristle-like point.
Total length 120fji ; length of dorsal plate 85/x, of ventral plate
95/x; width of dorsal plate 62/x, of ventral plate bSju; width of
anterior dorsal margin 17/a, of ventral margin 26/a; length of
toe 30/a.
416 ^¥isconsin Academy of Sciences, Arts and Letters.
Monostyla hatillifer was collected by Murray in a pond at
Sydney, Australia; our description is taken from his material.
No other localities are known for this interesting species.
Murray’s figures are slightly incorrect in some details, especial¬
ly the anterior margin and the form of the toe.
MONOSTYLA BIPURCA Bryce
Plate XLIV, figures 1, 2.
Monostyla hifurca Bryce, Science Gossip, vol. 28, 1892, p. 274, text
fig.
Notommata mono stylae for mis Stenroos, Acta Soc. Fauna et Flora
Fennica, vol. 17, No. 1, 1898, p. 126, pi. 1, fig. 25. — VoN Hofsten,
Arkiv ZooL, Stockholm, vol. 6, No. 1, 1909, p. 34.
Monostyla monostylaeformis Iroso, Mon. Zool. Italiano, vol. 21, 1910,
p. 303; Atti R. 1st. dlncorr. di Napoli, vol. 64 (for 1912), 1913,
p. 471, figs. 3, 4. — ^Fadeev, Raboti Sev.-Kavkazhsk. Gidrobiol.
Stants., vol. 1, pt. 1, 1925, p. 23, pi. 4, fig. 8.
Although the integument of this species is very flexible and
can not by any stretch of the imagination be called a lorica, it
is nevertheless sufficiently firm to assume a fairly definite form
when completely contracted. The outline of the body is very
broadly oval; its width is nearly equal to the length. The
anterior margin is straight and the dorsal and ventral edges fail
to meet in complete retraction. No lateral sulci are present and
the limits of the dorsal plate are distinguishable posteriorly
where it leaves the small posterior segment protruding slightly.
This bears at the sides of the foot two small, slightly curved
spines, which are probably to be interpreted as rudimentary
coxal plates, as in Lecane palinacis. The foot joints are very
large ; the' first joint is ovate and has a small, median, posterior
lobe; the second joint is heart-shaped. The toe is fairly long,
more than one fourth of the entire length, widest at the base and
tapering gradually towards the posterior end ; the claw is double,
its two parts spread wide apart and immovable.
Total length of contracted animal 75/x; length of dorsal plate
54fA, of ventral plate 58^, ; width of anterior margin 35 u ; length
of toe without claws 15jLt ; claws ; 4/a.
Monostyla hifurca is not rare in wet sphagnum ; the paucity of
records is probably due to its small size rather than to any actual
rarity.
Earring c& Myers — Rotifer Fauna of Wisconsin. — 111. 417
EXPLANATION OF PLATES.
All figures are highly magnified. For actual measurements
text.
Fig. 6. Lecane ploenensis, ventral view.
27
418 Wisconsin Academy of Sciences, Arts and Letters.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
PLATE XIV.
Lecane Candida, dorsal view, p. 368.
Lecane Candida, ventral view,
Lecane papuana, dorsal view, p. 336.
Lecane papuana, ventral view.
Lecane luna, dorsal view, p. 334.
Lecane luna, ventral view.
PLATE XV.
Lecane elegans, dorsal view, p. 371.
Lecane elegans, ventral view.
Lecane jessupi, dorsal view, p. 338.
Lecane jessupi, ventral view.
Lecane hrachydactyla, dorsal view, p. 337.
Lecane hrachydactyla, ventral view.
PLATE XVI.
Lecane depressa, dorsal view, p. 327.
Lecane depressa, ventral view.
Lecane ligona, dorsal view, p. 339.
Lecane ligona, ventral view.
Lecane ligona, dorsal view.
Lecane ligona, ventral view.
PLATE XVII.
Lecane rhacois, dorsal view, p. 379.
Lecane rhacois, ventral view.
Lecane clara, dorsal view, p. 378.
Lecane clara, ventral view.
Lecane pycina, dorsal view, p. 340.
Lecane pycina, ventral view.
PLATE XVIII.
Lecane eutarsa, dorsal view, p. 341.
Lecane eutarsa, ventral view.
Lecane pelatis, dorsal view, p. 342.
Lecane pelatis, ventral view.
Lecane mira, dorsal view, p. 342.
Lecane mira, ventral view.
PLATE XIX.
Lecane methoria, dorsal view, p. 343.
Lecane methoria, ventral view.
Lecane stichaea, dorsal view, p. 344.
Lecane stichaea, ventral view.
Lecane saginata, dorsal view, p. 345.
Lecane saginata, ventral view.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 419
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5. Lecane
Fig. 6. Lecane
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5. Lecane
Fig. 6. Lecane
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5. Lecane
Fig. 6. Lecane
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5. Lecane
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5. Lecane
Fig. 6. Lecane
Fig. 1. Lecane
Fig. 2. Lecane
Fig. 3. Lecane
Fig. 4. Lecane
Fig. 5.” Lecane
Fig. 6. Lecane
PLATE XX.
elasmay dorsal view, p. 345.
elasmay ventral view.
rhytiday dorsal view, p. 346.
rhytiday ventral view.
lauterbomiy dorsal view, p. 347.
lauterborniy ventral view.
PLATE XXL
comptay dorsal view, p. 347.
compta, ventral view.
haliclystay dorsal view, p. 348.
haliclystay ventral view.
aspasiay dorsal view, p. 349.
aspasiay ventral view.
PLATE XXII.
marsMy dorsal view, p. 351.
marshiy ventral view.
ichthyouray dorsal view, p. 352.
ichthyouray ventral view.
ludwigiiy dorsal view, p. 350.
ludwigiiy ventral view.
PLATE XXIII
stokesiiy dorsal view, p. 353.
stokesiiy ventral view.
stokesiiy dorsal view.
ohioensiSy dorsal view, p. 354.
ohioensisy ventral view.
PLATE XXIV.
arculay dorsal view, p. 355.
arculay ventral view.
flexiliSy dorsal view, p. 355.
flexilisy ventral view.
intrasinuatay dorsal view, p. 357.
intrasinuatay ventral view.
PLATE XXV.
climacoiSy dorsal view, p. 358.
climacoiSy ventral view.
verecunday dorsal view, p. 358.
verecunda, ventral view.
mylacriSy dorsal view, p. 359.
mylacriSy ventral view.
420 Wisconsin Academy of Sciences^ Arts and Letters.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
PLATE XXVI.
Lecane glypta, dorsal view, p. 360.
Lecane glypta, ventral view.
Lecane tabida, dorsal view, p. 361.
Lecane tabida, ventral view.
Lecane infula, dorsal view, p. 361.
Lecane infula, ventral view.
PLATE XXVn.
Lecane aeganea, dorsal view, p. 367.
Lecane aeganea, ventral view.
Lecane satyrus, dorsal view, p. 362.
Lecane satyrus, ventral view.
Lecane venusta, dorsal view, p. 368.
Lecane venusta, ventral view.
PLATE XXVIIL
Lecane crepida, dorsal view, p. 364.
Lecane crepida, ventral view.
Lecane sagula, dorsal view, p. 365.
Lecane sagula, ventral view.
Lecane hastata, dorsal view, p. 363.
Lecane hastata, ventral view.
PLATE XXIX.
Lecane formosa, dorsal view, p. 366.
Lecane formosa, ventral view.
Lecane mucronata, dorsal view, p. 330.
Lecane mucronata, ventral viev/.
Lecane mucronata, lateral view.
PLATE XXX.
Lecane pusilla, dorsal view, p. 369.
Lecane pusilla, ventral view.
Lecane asthena, dorsal view, p. 369.
Lecane asthena, ventral view.
Lecane subtilis, dorsal view, p. 370.
Lecane subtilis, ventral view.
PLATE XXXI.
Lecane elongata, dorsal view, p. 372.
Lecane elongata, ventral view.
Lecane tenuiseta, dorsal view, p. 372.
Lecane tenuiseta, ventral view.
Lecane doryssa, dorsal view, p. 373.
Lecane doryssa, ventral view.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 421
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig, 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
PLATE XXXII.
Lecane agilis, dorsal view, p. 382.
Lecane agilis ^ ventral view.
Lecane palinacis, dorsal view, p. 380.
Lecane palinacis, ventral view.
Lecane inopinata, dorsal view, p. 374.
Lecane inopinata, ventral view.
PLATE XXXIII.
Lecane inermis, dorsal view, p. 379.
Lecane inermis, ventral view.
Lecane calcaria, dorsal view, p. 381.
Lecane calcaria, ventral view.
Lecane niothis, dorsal view, p. 382.
Lecane niothis, ventral view.
PLATE XXXIV.
Lecane nana, dorsal view, p. 375.
Lecane nana, ventral view.
Lecane hornemanni, dorsal view, p. 377.
Lecane hornemanni, ventral view.
Lecane tryphema, dorsal view, p. 376.
Lecane tryphema, ventral view.
PLATE XXXV.
Monostyla lunaris, dorsal view, p. 384.
Monostyla lunaris, ventral view.
Monostyla lunaris, dorsal view.
Monostyla lunaris, ventral view.
Monostyla lunaris, dorsal view.
Monostyla lunaris, ventral view.
PLATE XXXVI.
Monostyla ornata, dorsal view, p. 402.
Monostyla ornata, ventral view.
Monostyla acus, dorsal view, p. 387.
Monostyla acus, ventral view.
Monostyla crenata, dorsal view, p. 386.
Monostyla crenata, ventral view.
PLATE XXXVII.
Monostyla bulla, dorsal view, p. 388.
Monostyla bulla, ventral view.
Monostyla styrax, dorsal view, p. 389.
Monostyla styrax, ventral view.
Monostyla goniata, dorsal view, p. 390.
Monostyla goniata, ventral view.
422 Wisconsin Academy of Sciences, Arts and Letters,
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
Fig. 1.
Fig. 2.
Fig. 3.
Fig. 4.
Fig. 5.
Fig. 6.
PLATE XXXVIII.
Monostyla tethis, dorsal view, p. 405.
Monostyla tethis, ventral view.
Monostyla quadridentata, dorsal view, p. 391.
Monostyla quadridentata, ventral view.
Monostyla quadridentata, male, dorsal view.
PLATE XXXIX.
Monostyla stenroosi, dorsal view, p. 394.
Monostyla stenroosi, ventral view.
Monostyla thalera, dorsal view, p. 393.
Monostyla thalera, ventral view.
Monostyla lamellata, dorsal view, p. 392.
Monostyla lamellata, ventral view.
PLATE XL.
Monostyla scutata, dorsal view, p. 401.
Monostyla scutata, ventral view.
Monostyla sylvatica, dorsal view, p. 398.
Monostyla sylvatica, ventral view.
Monostyla cornuta, dorsal view, p. 396.
Monostyla cornuta, ventral view.
PLATE XLI.
Monostyla copeis, dorsal view, p. 398.
Monostyla copeis, ventral view.
Monostyla rhopalura, dorsal view, p. 399.
Monostyla rhopalura, ventral view.
Monostyla pideis, dorsal view, p. 400.
Monostyla pideis, ventral view.
PLATE XLII.
Monostyla pygmaea, dorsal view, p. 401.
Monostyla pygmaea, ventral view.
Monostyla vastita, dorsal view, p. 404.
Monostyla vastita, ventral view.
Monostyla obtusa, dorsal view, p. 403,
Monostyla obtusa, ventral view.
PLATE XLIII.
Monostyla elachis, dorsal view, p. 406.
Monostyla elachis, ventral view.
Monostyla rugosa, dorsal view, p. 405.
Monostyla rugosa, ventral view.
Monostyla furcata, dorsal view, p. 407.
Monostyla furcata, ventral view.
Harring & Myers — Rotifer Fauna of Wisconsin. — III. 423
PLATE XLIV.
Fig. 1. Monostyla bifurca, dorsal view, p. 416.
Fig. 2. Monostyla bifurca^ ventral view.
Fig. 3. Monostyla punctataj dorsal view, p. 407.
Fig. 4. Monostyla punctata, ventral view.
Fig. 5. Monostyla closterocerca, dorsal view, p. 408.
Fig. 6. Monostyla closterocerca, ventral view.
PLATE XLV.
Fig. 1. Monostyla pyriformis, dorsal view, p. 409.
Fig. 2. Monostyla pyriformis, ventral view.
Fig. 3. Monostyla subulata, dorsal view, p. 410.
Fig. 4. Monostyla subulata, ventral view.
Fig. 5. Monostyla opias, dorsal view, p. 411.
Fig. 6. Monostyla opias, ventral view.
PLATE XLVI.
Fig. 1. Monostyla monostyla, dorsal view, p. 411.
Fig. 2. Monostyla monostyla, ventral view.
Fig. 3. Monostyla batillifer, dorsal view, p. 415.
Fig. 4. Monostyla batillifer, ventral view.
PLATE XLVII.
Fig. 1. Monostyla hamata, dorsal view, p. 414.
Fig. 2. Monostyla hamata, ventral view.
Fig. 3. Monostyla arcuata, dorsal view, p. 412.
Fig. 4. Monostyla arcuata, ventral view.
Fig. 5. Monostyla decipiens, dorsal view, p. 413.
Fig. 6. Monostyla decipiens, ventral view.
CORRECTIONS TO PART II, VOLUME XXI.
Pp. 488 and 546 : for Cephalodella piulca read C. hiulca. P. 501 :
Cephalodella collactea has a cervical eyespot. P. 505: Cephalodella
cuneata also has a cervical eyespot.
1
TRANS. WIS. ACAD. VOL XXIi.
PLATE IX
MARRING AND MYERS • LECANE
TRANS. W!S. ACAD.. VOL XXII.
PLATE X
HARPING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XI
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XII
HARRINa AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XIII
MARRING AND MYERS - LECANE
!
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XIV
HARRINS AND MYERS • LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XV
1
2
HARRING AND MYERS • LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XVI
HARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XVII
MARRING AND MYERS - LECANE
HARRING ANB MYERS • LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XIX
MARRING AND MYERS - LECANE
TRANS.
PLATE XX
and MYERS ■ LECANE
j
TRANS. WIS. ACAD.. VOL XXII.
PLATE^XXI
HARRING AND MYERS - LECANE
■TRANS. WtS. ACAD.. VOL. XXII.
PLATE XXII
MARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXIII
HARPING AND MYERS • LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XXIV
HARKING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XXV
MARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XXVI
MARRING AND MYERS • LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXVII
HARKING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XXVIII
MARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
PLATE XXIX
MARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
HATE XXX
MARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL XXII.
RLATE XXXI
HARKING AND MYERS • LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXXIl
HARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXXlll
HARKING AND MYERS - LECANE
ikanS. WIS. ACAD.. VOL XXII.
PLATE XXXIV
HARRING AND MYERS - LECANE
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXXV
MARRING AND MYERS - MONOSTYLA
HARRINC AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXXVII
2
1
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XXXVIII
HARRING AND MYERS - MONOSTYLA
■’,'■11
i
TRANS. WIS. ACAD.. VOL. XXII
PLATE XXXIX
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL XXII.
PUTE XL
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XLI
HARRIN6 AND MVERS - MONOSTYLA
TRANS. WIS. ACAD.. YOL. XXII. PLATE XLII
2
1
HARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XLIII
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XLIV
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL. XXII.
PLATE XLV
MARRING AND MYERS • MONOSTYLA
TRANS. WIS. ACAD.. VOL XXII.
PUTE XLVI
MARRING AND MYERS - MONOSTYLA
TRANS. WIS. ACAD.. VOL XXII.
PUTE XLVIl
HARRING AND MYERS - MONOSTYLA
i
PROCEEDINGS OF THE ACADEMY
1924 AND 1925
FIFTY-FOURTH ANNUAL MEETING, 1924
The fifty-fourth annual meeting of the Wisconsin Academy
of Sciences, Arts, and Letters, in joint session with the Wiscon¬
sin Archeological Society, was held at Madison, in the Biology
Building of the University of Wisconsin, on Friday and Satur¬
day, April 11 and 12, 1924.
Under the direction of the president the following programme
was presented:
First Session, Friday, April 11, 10:00 A.M.
General Business.
Presentation of Papers.
1. Notes on some Wisconsin Saprolegniaceae. I. E. M. Gilbert.
Illustrated.
2. Notes on some Wisconsin Saprolegniaceae. II. J. A. LoUNS-
BURY. Illustrated.
3. Some relations of chlorides in plant nutrition. W. E. Totting-
HAM.
4. The nutrition of Venturicu inaequalis (Cooke) Went. C. N.
Frey. By title.
5. A method of anesthetizing Mimosa. C. N. Frey and E. H. Toole.
By title.
6. Some water mites from Alaska. Ruth Marshall. By title.
7. Two Diptera suitable for experimental work in genetics. C. L.
Turner.
8. Factors influencing distribution of fresh-water ciliates. L. E.
Noland.
9. Anti-body formation from injured tissues. M. F. Guyer.
10. Some geographic influences in the development of the Wiscon¬
sin dairy industry. G. T. Trewartha.
11. The constitution of liquids. Farrington Daniels. Illustrated.
Second Session, Friday, April 11, 2:00 P.M.
Presentation of Papers.
12. Duluth and the Discovery of the Northwest. Louise P. Kel¬
logg.
426 Wisconsin Academy of Sciences, Arts and Letters.
13. The name Chicago: Its Indian original, etc. E. P. Wheeler.
14. The teaching of Wisconsin history. W. C. English.
15. Constellations of a Pawnee sky map. R. A. Buckstaff. Fifteen
minutes. Illustrated.
16. The antiquity of the tubular drill. Geo. B. Phillips.
17. The Archangel campaign. John G. Gregory.
18. Some interesting customs of the people of India. Nand Singh.
Illustrated.
19. Napoleonic soldiers buried in Wisconsin. Albert 0. Barton.
20. A Wisconsin copper axe and gouge. Charles E. Brown.
Third Session, Saturday, April 12, 9:30 A.M.
General Business.
Presentation of Papers.
21. Magnetic rotation in cobalt films. L. R. Ingersoll. Illustrated.
22. The chance of rain. E. R. Miller. •
23. The dust fall of February 13, 1923. A. N. Winchell and E. R.
Miller.
24. The fauna of the Galena limestone near Appleton. John W.
Ockerman. (Introduced by R. M. Bagg)
25. Glacial history of Lake Mendota. W. 0. Hotchkiss.
26 Magnetic work in geology. H. R. Aldrich. Illustrated.
27. The new geological groups at the Milwaukee Public Museum.
Ira Edwards. Illustrated.
28. Road materials investigations. E. F. Bean.
The annual dinner was held on Friday evening, April 11,
at 6:30 p. m. Thirty-two members and guests were present.
Following the dinner Professor Max Mason delivered an address
on ‘‘Some Gordian Knots in Physical Theory’’.
At the business session the Secretary presented the following
applications for membership. On motion he was unanimously
instructed to cast the ballot in their favor.
Bernhard P. Domogalla, Madison
Percival T. Gates, Chicago
F. Gregory Hall, Milton
Edward S. Hathaway, Madison
James A. Lounsbury, Madison
Harland W. Mossman, Madison
Harvey M. Smith, Madison
Rubert B. Streets, Madison
Report of Secretary, April 1, 1923, to March 31, 1924
Honorary Members _ : _ 7
Life Members _ 17
Proceedings of the Academy
427
Corresponding Members - 31
Active Members _ 306
361
Resigned _ 2
Deaths - 5
Dropped for non-payment of dues _ 15
32
Five members have been lost by death during the past year:
Bishop Samuel Fallows, Professor R. D. Salisbury, Professor M. S.
Slaughter, President Sidney T. Smythe, Mr. Edgar E. Teller.
Chancey Juday,
Secretary,
Report of Treasurer, April 1, 1923, to March 31, 1924
Receipts
Balance in Treasury April 1, 1923 _ $1,547.26
Received for dues and initiations _ 273.00
Interest on bonds in 1923 _J _ L 150.25
Received from sales of Transactions _ 30.44
Bonds maturing in 1924 _ 1,000.00
Interest on bonds in 1924 _ 156.25
Annual appropriation July 1, 1923 _ 1,500.00
$4,657.20
Disbursements
Bonds purchased - $1,200.00
Certificate of deposit _ 72.75
Safe deposit rent, 1923 and 1924 _ 5.00
Expenses of speakers at Beloit" meeting _ 27.00
Postage _ 2.51
Printing and stationery _ 130.66
$1,437.92
Balance in Treasury March 31, 1924 _ _ _ 3,219.28
Securities held by Academy, March 31, 1924
U. S. Government bond _ $100.00
City of Madison bonds _ 2,800.00
Certificates of deposit _ 72.75
$2,972.75
428 Wisconsin Academy of Sciences, Arts and Letters.
Audited and found correct.
Chancey Juday,
Treasurer.
C. L. Turner,
A. S. Pearse,
liuditors.
FIFTY-FIFTH ANNUAL MEETING, 1925
The fifty-fifth annual meeting of the Wisconsin Academy of
Sciences, Arts and Letters was held at the State Normal School,
Oshkosh, on Friday and Saturday, April 10 and 11, 1925, in
joint session with the Wisconsin Archeological Society, the Wis¬
consin Museums Conference, and the Winnebago County Arche¬
ological and Historical Society.
The following programme was presented at the various ses¬
sions.
First Session, Friday, April 10, 9:30 A.M.
General Business.
Presentation of Papers.
1. The method by which parental characters in a liverwort are dis¬
tributed among the offspring. C. E. Allen. Illustrated.
2. Temperature relations to seed germination. J. F. Groves. Il¬
lustrated.
3. The orchids of Wisconsin. Albert E. Fuller. Illustrated.
4. Notes on parasitic fungi in Wisconsin. XIII. J. J. Davis. By
title.
5. The reaction pathway, with demonstration. R. C.-Mullenix.
6. Some recent discoveries in the Wisconsin Cambrian. Ira Ed¬
wards.
7. Artesian conditions of the Appleton region. J. W. Ockerman,
8. Progress in bird banding. W. I. Lyon. Illustrated.
Second Session, Friday, April 10, 2:00 P.M.
General Business.
Presentation of Papers.
9. An unusual agricultural museum. L. J. Cole. Illustrated.
10. Practical benefits of the Wisconsin Museums Conference. S. A.
Babbett.
Proceedings of the Academy
429
11. Administrative methods for small museums. W. D. Babcock,
Jr.
12. Round table discussion of museum topics. The discussion will
be led by Mrs. Arthur C. Neville and Miss Deborah B.
Martin.
Third Session, Saturday, April 11, 9:00 A.M.
General Business.
Presentation of Papers.
13. The flowers of Shakespeare^s garden. . Huron H. Smith. Illu¬
strated.
14. Extracts from the original letter book of Judge James Duane
Doty dated 1821. Gene Sturtevant.
16. The Wisconsin military road. H. E. Cole.
16. Development of a typical Wisconsin rural community. E. A.
Clemens.
17. Indian remains of Winnebago County. George Overton.
18. Wisconsin archeological researches, 1912-1925. Charles E.
Brown.
The annual dinner was held at the Athearn Hotel on Friday
evening at 6:30 p. m. Forty-three members and guests were
present.
Following the dinner Mr. Alonzo W. Pond delivered an ad¬
dress on ‘^Delving Back of History’’, illustrated with lantern
slides. Dr. S. A. Barrett then gave an address on ‘‘Filming
Moose on Isle Royale”, illustrated with moving pictures. These
addresses were held in the Assembly Room of the City High
School of Oshkosh.
At the business session the Secretary presented the following
applications for membership. On motion he was unanimously
instructed to cast the ballot in their favor.
E, F. Bean, Madison
Nile J. Behncke, Oshkosh
Tranquilino G. Fajardo, Madison
Albert M. Fuller, Milwaukee
A, A. Granovsky, Madison
John W. Ockerman, Appleton
C. M. Tompkins, Madison
E. J. Wimmer, Milwaukee.
R. T. Zoch, Madison
430 Wisconsin Academy of Sciences, Arts and Letters.
Report of Secretary, April 1, 1924, to March 31, 1925
Honorary Members _ 7
Life Members _ 15
Corresponding Members _ 28
Active Members _ 288
338
Resigned _ 2
Dropped for non-payment of dues _ 4
Deaths _ 4
10
Information regarding the death of four members has been
received during the past year :
Professor Joseph Paxon Iddings, Mr. Thomas Evans Brit-
tingham, President Samuel Plantz, Dr. Norman Bridge.
A vote of thanks was extended to the Winnebago County
Archeological and Historical Society, the State Normal School,
and the City High School for their interest in the meetings and
for the use of rooms for the sessions.
Chancey Juday,
Secretary.
Report of Treasurer, April 1, 1924, to March 31, 1925
Receipts
Balance in Treasury, April 1, 1924 - $3,204.28
Sale of Transactions _ 9.57
Received for reprints _ 175.00
Received from Wisconsin Survey for printing - 500.00
Interest on bonds - 7.93
Annual dues of members - 296.00
Annual appropriation July 1, 1924 - 1,500.00
$5,692.78
Disbursements
Printing announcements and programmes - $16.12
Printing Volume XXI of Transactions - 2,202.67
Allowance of Secretary for 1924 - 200.00
Miscellaneous expenses and postage - 9.18
$2,427.97
Proceedings of the Academy 431
Balance in Treasury March 31, 1925 _ 3,264.81
Cash and securities held by Academy, March 31, 1925
U. S. Government bond _ _ $100.00
City of Madison bonds _ _ 2,900.00
Certificates of deposit! _ _ 73.66
Cash _ 42.82
$3,116.48
Audited and found correct.
Chance Y Juday,
Treasurer.
E. N. Buckstaff,
George Wagner,
Auditors.
LIST OF OPPICEKS AND MEMBERS
Corrected to November 1, 1925.
Officers
President, L. J. Cole, Madison.
Vice-President, Sciences, Samuel A. Barrett, Milwaukee.
Vice-President, Arts, Grant Showerman, Madison.
Vice-President, Letters, Arthur Beatty, Madison.
Secretary, Chancey Juday, Madison.
Treasurer, Chancey Juday, Madison.
Curator, C. E. Brown, Madison.
Librarian, Walter M. Smith, Madison.
Committee on Publication
The President, ex officio,
The Secretary, ex officio,
W. E. Tottingham, Madison.
Council
The President, Vice-Presidents, Secretary, Treasurer, Libra¬
rian and Past Presidents retaining their residence in Wisconsin.
432 Wisconsin Academy of ScienceSy Arts and Letters.
Committee on Library.
The Librarian, ex officio,
W. H. Barber, Madison,
A. E. Culver, Stevens Point,
George Wagner, Madison.
Committee on Membership.
The Secretary, ex officio,
A. S. Pearse, Madison,
P. W. Boutwell, Beloit,
R. N. Buckstaff, Oshkosh,
Huron H. Smith, Milwaukee.
Past Presidents.
Honorable John W. Hoyt,^ 1870-1875.
Doctor P. R. Hoy,i 1876-1878.
President A. L. Chapin,^ 1879-1881.
Proffessor Ronald D. Irving,^ 1882-1884.
Professor Thomas C. Chamberlin, 1885-1887.
Professor William F. Allen,’- 1888-1889.
Professor Edward A. Birge, 1889-1890.
Librarian George W. Peckham,^ 1891-1893.
President Charles R, Van Hise,’- 1894-1896.
Professor C. Dwight Marsh, 1897-1899.
Professor Charles S. Slighter, 1900-1902.
Doctor John J. Davis, 1903-1905.
Professor Louis EAhlenberg, 1906-1909.
President Samuel Plantz,’^ 1910-1912.
Professor Dana C. Munro, 1913-1915.
Director Henry L. Ward, 1916-1918.
President Edward A. Birge, 1919-1921.
President Melvin A. Brannon, 1922-1924.
HONORARY MEMBERS
Chamberlin, Thomas Chrowder, University of Chicago, Chi¬
cago, Ill.
Garland, Hamlin, New York, N. Y.
^ Deceased.
Proceedings of the Academy
433
Jordan, David Starr, Stanford University, Cal.
Merrick, George B., 350 W. Washington Ave., Madison.
Trelease, William, University of Illinois, Urbana, Ill.
Wheeler, William Morton, Forest Hills, Boston, Mass.
LIFE MEMBERS
Beatty, Arthur, 1824 Vilas Street, Madison
^Birge, Edward A., 2011 Van Hise Ave., Madison
'^Davis, John Jefferson, 419 Sterling Place, Madison
Hixon, Frank P., La Crosse
Hobbs, William H., University of Michigan, Ann Arbor,
Michigan
Horlick, A. J., Racine
'^Leith, Charles K., Moraine, Old Sauk Road, Madison
Logan, Frank A., Chicago, Illinois
*Marsh, C. Dwight, 1882 Monroe Street, Washington, D. C.
Norris, Mrs. Fanny, 1906 Grand Ave., Milwaukee
Peckham, Mrs. Elizabeth G., 46 Marshall Street, Milwaukee
Sharp, Frank C., University of Wisconsin, Madison
’^"Skinner, Ernest Brown, 210 Lathrop Street, Madison
^'Sliehter, Charles Sumner, 636 North Frances Street, Madison
CORRESPONDING MEMBERS
Coulter, John Merle, University of Chicago, Chicago, Illinois
Crooker, Joseph Henry, 24 East Concord Ave., Orlando,
Florida
Eckels, William Alexander, La Fayette College, Easton, Pa.
Hendrickson, George Lincoln, Yale University, New Haven,
Conn.
Hoskins, Leander Miller, Palo Alto, California
Kinley, David, University of Illinois, Urbana, Illinois
Leverett, Frank, University of Michigan, Ann Arbor, Michi¬
gan
Libby, Orin Grant, Grand Forks, North Dakota
Marx, Charles David, Palo Alto, California
McClumpha, Charles Flint, 56 Church Street, Amsterdam,
New York
♦Member of American Association for the Advancement of Science.
28
434 Wisconsin Academy of Sciences, Arts and Letters.
]\Iunro, Dana Carlton, Princeton University, Princeton, New
J ersey
Olive, Edgar W., Brooklyn Botanic Garden, Brooklyn, New*
York
Power, Frederick Belding, Bureau of Chemistry, Washing¬
ton, D. C.
Stone, Ormond, Clifton Station, Virginia
Tolman, Albert Harris, University of Chicago, Chicago,
Illinois
Townley, Sidney Dean, Stanford University, California
Turner, Frederick Jackson, 2214 Van Hise Ave., Madison
Verrill, Addison Emery, Yale University, New Haven, Con¬
necticut
ACTIVE MEMBERS
*Addleman, Irving M., 511 Third Street, Wausau
^Addoms, Ruth M., 419 Sterling Place, Madison
Aitken, Robert T., Bishop Museum, Honolulu, Hawaii
Alderman, William E., 718 Church Street, Beloit
* Allen, Charles E., 2014 Chamberlain Avenue, Madison
*Arzberger, Emil G., Bureau of Plant Industry, Washington,
D. C.
*Bagg, Rufus M. Jr., 7 Brokaw Place, Appleton
*Baird, Edgar A., University of North Dakota, Grand Forks,
N. D.
Barber, Wm. Harley, 303 Princeton Avenue, Madison.
*Bardeen, Charles RusseU, 23 Mendota Court, Madison ,
^Barnes, Kenneth B., 718 Church Street, Beloit
Barrett, Samuel A., Public Museum, Milwaukee
^Barrett, Storrs Barrows, Yerkes Observatory, Williams Bay
Barth, George Peter, 3006 Chestnut Street, Milwaukee
Bascom, Delia, 419 Sterling Place, Madison
*Bean, Elizabeth Smith, Dalhousie University, Halifax, N. S.
*Bean, E. F., College Hills, Madison
Behncke, Nile J., 785 Algoma Blvd., Oshkosh
* Bennett, Ed'ward, 1919 Jefferson Street, Madison
Bleyer, Willard G., 423 North Carroll Street, Madison
*Boutwell, Paul Winslow, Beloit College, Beloit
*Boyle, C., University College, Cork, Ireland
“^'Brannon, Melvin Amos, State Capitol, Helena, Montana
Proceedings of the Academy
435
Brauiij Adolph E., 832 38th Street, Milwaukee
^Brink, E. Alex., University of Wisconsin, Madison
Brown, Charles E., 136 Lathrop Street, Madison
Brown, Charles Newton, 41 Eoby Eoad, Madison
^Browne, Frederick Lincoln, 223 Clifford Court, Madison
^Browning, Harold William, State College, Kingston, E. I.
*Bryan, George Smith, 803 State Street, Madison
Buckstaff, Ealph N., 1501 South Main Street, Oshkosh
Buehler, Henry Andrew, Eolla, Missouri
^Bunting, Charles Henry, 2020 Chadbourne Avenue, Madison
Busssewitz, M. A., 435 Kenwood Boulevard, Milwaukee
Butterfield, Mrs. Ellen F., 713 Milwaukee Avenue, South Mil¬
waukee
Cahn, A. E., 606 East Stoughton Street, Champaign, Illinois
Cairns, William B., 2010 Madison Street, Madison
Carter, Sylvester J., 437 Kenwood Boulevard, Milwaukee
Chandler, Elwyn Francis, University of North Dakota, Grand
Forks, N. D.
Chase, Wayland J., 141 Summit Avenue, Madison
*Churchill, B. P., 569 Mitchell Street, Milwaukee
’^Clark, Paul F., 2136 Van Hise Ave., Madison
Clark, William A., State Normal School, Stevens Point
Clas, Alfred Charles, 445 Milwaukee Street, Milwaukee
Clawson, Arthur Brooks, 1884 Monroe Street, N. W. Washing¬
ton, D. C.
Clowes, Herbert, Public Museum, Milwaukee
Cole, Harry E., 908 Ash Street, Baraboo
^Cole, Leon J., 1915 Keyes Ave., Madison
*Collie, George Lucius, 48 Sherwood Drive, Beloit
Commons, John E., University of Wisconsin, Madison
* Comstock, George C., 635 College Street, Beloit
Conklin, George Hall, 1924 John Ave., Superior
*Conwell, H. H., 921 Park Ave., Beloit
^Cooper, George Olds, 431 Hawthorne Court, Madison
^Culver, Garry Eugene, State Normal School, Stevens Point
^Daniels, Farrington, Nakoma, Madison
^Davis, Carl Henry, 141 Wisconsin Street, Milwaukee
^Dawson, Percy M., University of Wisconsin, Madison
Dean, Aletta F., 87 West Street, Mansfield, Mass.
^Delwiche, Edmond Joseph, E. F. D. 3, Green Bay
^Densmore, H. D., 718 Clary Street, Beloit
4.36 Wisconsin Academy of Sciences^ Arts and Letters.
'^Denniston, Eollin Henry, 1414 Washington Ave., Madison
Dodge, B. 0., Bureau of Plant Industry, Washington, D. C.
Dodge, K. E. N., 115 N. Allen Street, Madison
'^Domogalla, Bernhard P., 803 State Street, Madison
^Dowling, Linnaeus Wayland, 2 Koby Koad, Madison
Downes, Eobert Hugh, 574 Algoma Street, Oshkosh
'^Drechsler, Charles, Department of Agriculture, Washington,
D. C.
^Dresden, Arnold, 2114 Vilas Street, Madison
DuMez, Andrew Grover, 25th and E. Street, N. W., Washing¬
ton, D. C.
'^'Edwards, Ira, Public Museum, Milwaukee
Ely, Eichard T., Northwestern University, Evanston, Ill.
^Evans, Clarence T., 3415 Sycamore Street, Milwaukee
^'Evans, Edward, 121 South 13th Street, La Crosse
*Fajardo, Tranquilino G., 147 Lathrop St., Madison
^Farley, John Herbert, 474 Alton Street, Appleton
^Fiebiger, George J., Waterloo
Finch, Y. C., College Hills, Madison
Finkler, Adolph, 612 Commerce Street, Milwaukee
^Fischer, Eichard, 119 E. Johnson, Madison
Fish, Carl Eussell, 511 N. Carroll, Madison
^Fish, Emma Luella, University of Wisconsin, Madison
Fluke, Charles L., Jr., 512 Wingra, Madison
^Foster, Arthur C., Box 549, Sandford, Florida
'^Fr acker, Stanley B., 1632 Adams Street, Madison
^Fred, Edwin B., 610 N. Frances Street, Madison
Frey, Charles N., 103 West 183rd Street, New York, N. Y.
^Frost, William D., 1010 Grant Street, Madison
Fuller, Albert M., Public Museum, Milwaukee
*"Gates, Percival T., University of Chicago, Chicago, Ill.
Gay, Lucy M., 216 N. Pinckney Street, Madison
^ Gerry, Eloise, 419 Sterling Place, Madison
'^Gilbert, Edward M., 2120 Chamberlain Ave., Madison
Glicksman, Harry, 2024 Chamberlain Ave., Madison
Gloyer, Walter 0., 106 Lyceum Street, Geneva, N. Y.
Granovsky, Alexander A., 803 State Street, Madison
Greene, Howard, 559 Marshall Street, Milwaukee
Gregory, John G., 729 Jenifer Street, Madison
^Groves, James F., Eipon College, Eipon
Gutseh, Milton E., University of Texas, Austin, Texas
Proceedings of the Academy
437
'^Guyer, Michael F., 138 Prospect Ave., Madison
Haase, Ewald, 182 Wisconsin Street, Milwaukee
Haessler, Louise, 100 Morningside Drive, New York, N. Y.
Hall, P. Gregory, Milton College, Milton
*Hall, Sidney S., 159 Malcolm Ave., S. E., Minneapolis, Minn.
Hargrave, Josephine R., Ripon College, Ripon
Harper, Robert A., Columbia University, New York, N. Y.
Harper, Mrs. Robert A., Columbia University, New York, N. Y.
^Hastings, E. G., 2111 Van Hise Ave., Madison
^Hathaway, Edward S., Tulane University, New Orleans, La.
^Hawkins, Pliny H., Absarokee, Mont.
^Heddle, John R., 1215 Carlisle Ave., Racine
“^Heineman, Paul G., 1508 Jonquil Terrace, Chicago, Ill.
^Henmon, Y. A. C., College Hills, Madison
^Hodgson, A. J., Box 166, Waukesha
Hohlfeld, A. R., 124 Breese Terrace, Madison
Holmes, Samuel J., University of California, Berkeley, Cal.
^Hotchkiss, W. 0., State School of Mines, Houghton, Mich.
Hubbard, Frank G., 141 W. Gilman Street, Madison
^Hubert, Ernest E., Forest Products Laboratory, Madison
’^Humphrey, Clarence J., 1714 Adams Street, Madison
*Ingersoll, Leonard R., 1933 West Lawn Ave., Madison
Jackson, Hartley H. T., U. S. Biological Survey, Washington,
D. C.
*Jana, Ashutosh, Tamluk, Bengal, India
^Jansky, Cyril M., 2117 Jefferson Street, Madison
^Jasper, Thomas McLean, A. 0. Smith Corporation, Milwaukee
’^Jastrow, Joseph, 237 Langdon Street, Madison
^Johnson, Aaron G., Department of Agriculture, Washington,
D. C.
* Johnson, James, 2005 Madison Street, Madison
"^Jones, Fred R., 1713 Chadbourne Ave., Madison
*Jones, Mrs. F. R., 1713 Chadbourne Ave., Madison
^Jones, Leon K., University of Wisconsin, Madison
*Jones, Lewis Ri, 146 Prospect Ave., Madison
^ Jones, Sarah Y. H., Rochester, Mich.
*Juday, Chancey, 35 Lathrop Street, Madison
^Kahlenberg, Louis, 234 Lathrop Street, Madison
*Keitt, George W., 803 State Street, Madison
^Kemmerer, George I., 916 Yan Buren Street, Madison
438 ^Yisconsin Academy of Sciences, Arts and Letters.
Kenyon, W. A., Hamline University, St. Paul, Minn.
Kiekhofer, W. H., 1919 Arlington Place, Madison
*Koeliler, Arthur, 1819 Adams Street, Madison
^^Kowalke, 0. L., 2012 Jefferson Street, Madison
*Kraus, Ezra J., 803 State Street, Madison
*Kremers, Edward, University of Wisconsin, Madison
Lannerd, Willard H., 103 Parkwood Blvd., Mansfield, 0.
*Lee, Oliver J., Yerkes Observatory, Williams Bay
*Lenher, Victor, 1718 Summit, Madison
^Leonard, Clifford S., Yale University, New Haven, Conn.
Lewis, F. F., 115 Jackson Street, Janesville
*Lobeck, Armin K., 1635 Madison Street, Madison
Loft, Genivera E., 2121 Kendall Ave., Madison
Lounsbury, James A., Marquette University, Milwaukee
*Lowe, John N'., State Normal School, Marquette, Mich.
'’^McAllister, Fred, University of Texas, Austin, Texas
McKenna, Maurice, 114 3rd Street, Fond du Lac
McLeod, Andrew F., 5986 Kidge Ave., Chicago, Ill.
^McMinn, Amelia, State Normal School, Milwaukee
]\Iain, Mrs. Angie K., Fort Atkinson
Majerowski, Stephen J., 932 10th Ave., Milwaukee
*March, Herman W., 1825 Summit Ave., Madison
^Marlatt, Abby L., 612 Howard Place, Madison
"^Marquette, William G., Pleasantville, N. Y.
^]Marschall, A. J., 14 Proudfit Street, Madison
Marshall, Ruth, Rockford College, Rockford, Ill.
'^Marshall, William S., 139 E. Gilman, Madison
^Martin, Ella May, Illinois Wesleyan University, Bloomington,
Ill.
*Mason, Max, University of Chicago, Chicago, Ill.
^Mathews, J. How^ard, 128 Lathrop Street, Madison
^Maurer, Edward R., 167 Prospect Ave., Madison
'’’'Mavor, James W., Union College, Schnectady, N. Y.
'^Meachem, John G., 745 College Ave., Racine
*Mead, Daniel W., 120 W. Gorham Street, Madison
*Mead, Warren J., 2113 Adams Street, Madison
'’^IMendenhall, Charles E., 510 N. Carroll Street, Madison
Meyer, Balthaser H., 3825 Wisconsin Ave., Washington, D. C.
^Miller, Eric R., 2125 Van Hise Ave., Madison
^Miller, William S., 2001 Jefferson Street, Madison
^Aloe, Carl, Box 75, International Falls, Minn.
Proceedings of the Academy
439
Monroe, C. E., 728 Racine Street, Milwaukee
^Morris, Harold H., 2040 Pine Street, Philadelphia, Penn.
^Mossman, Harland W., 1707 Jefferson Street, Madison
*Mullenix, Rollin C., Lawrence College, Appleton
^Murphy, Ray B., Illinois Wesleyan University, Bloomington,
Ill.
^Musbach, Fred L., 407 S. Cherry Street, Marshfield
Muttkowski, Richard A., Detroit University, Detroit, Mich.
^Nagler, Mrs. Ellen T., 1913 Kendall Ave., Madison
^Nardin, F. Louise, Lathrop Hall, Madison
Naylor, Wilson S., Lawrence College, Appleton
^Noland, Lowell E., 1723 Summit Ave., Madison
Oberholser, Harry C., 2805 18th Street, N. W., Washington,
D. C.
Ockerman, J. W., 416 South Elwood Street, Tulsa, Oklahoma
*Ogden, Henry V., 141 Wisconsin Street, Milwaukee
^O’Shea, M. Vincent, 529 N. Pinckney Street, Madison
Otto, Max C., 1155 Edgewood Ave., Madison
^Overton, James B., 512 Wisconsin Ave., Madison
Pammel, L. H., Iowa State College, Ames, Iowa
Parker, Fletcher A., 14 W. Gilman Street, Madison
Patzer, C. E., State Normal School, Milwaukee
Paxson, Frederick L., 2122 Van Hise Ave., Madison
*Pearse, A. S., 2240 Rowley Ave., Madison
Peltier, George L., College of Agriculture, Lincoln, Neb.
Peter, George, Public Museum, Milwaukee
^Peterson, William H., University of Wisconsin, Madison
'^Phillips, James D., University of Wisconsin, Madison
Pitman, Annie M., 414 N. Henry Street, Madison
Pope, Thomas E. B., Public Museum, Milwaukee
Porteus, Charles R., Fublic Museum, Milwaukee
^Potter, George F., New Hampshire College, Durham, N. H.
Powell, John W., 471 Van Buren Street, Milwaukee
^Public Museum, Milwaukee
*Reed, George M., Brooklyn Botanic Garden, Brooklyn, N. Y.
’^Reid, Mary E., Boyce Thompson Institute, Yonkers, N. Y.
Richards, C. Audrey, 415 Sterling Place, Madison
^Rickett, Harold W., University of Missouri, Columbia, Mo.
*Riker, Mrs. A. J., 105 N. Spooner Street, Madison
^Roark, Raymond J., 1707 Madison Street, Madison
^Roebuck, John R., 2210 Hollister Ave., Madison
440 Wisconsin Academy of Sciences, Arts and Letters.
Roedder, E. C. L. C., 38 Breese Terrace, Madison
Rogers, Thomas A., State Normal School, Stevens Point
'"•’Rose, Jessie P., 1101 7th Street, S. E., Minneapolis, Minn.
^Sammis, John L., 234 Breese Terrace, Madison
Schinner, Augustine F., Right Reverend, 238 E. 13th Ave.,
Spokane, Wash.
^Schlundt, Herman, University of Missouri, Columbia, Mo.
^Schmidt, Albert H., Palatine, Cook County, Ill.
Schorger, A. W., 2021 Kendall Ave., Madison
Schubring, E. J. B., 415 N. Park Street, Madison
Schubring, Mrs. E. J. B., 415 N. Park Street, Madison
*Schuette, Henry A., Maple Bluff, Madison
’^Schultz, Alfred R., 800 Vine Street, Hudson
Seaman, Gilbert E., 312 Goldsmith Building, Milwaukee
Showalter, Amos M., Department of Botany, Stanford Uni¬
versity, Calif.
Showerman, Grant, 410 N. Butler Street, Madison
Sieker, William C., 7th and Prairie Streets, Milwaukee
*Simon, Arthur, Cutler-Hammer Mfg. Co., Milwaukee
Smith, Elmer A., 1281 Pat. Pl’k. Road, Secaucus, N. J.
*Smith, Gilbert B. L., 401 Dryden Road, Ithaca, N. Y.
*Smith, Gilbert M., Stanford University, Cal.
*Smith, Harvey M., University of Wyoming, Laramie, Wyo.
*Smith, Huron H., Public Museum, Milwaukee
Smith, Walter M., 127 Langdon Street, Madison
*Snell, Walter H., 19 Nisbet Street, Providence, R. I.
*Snow, Benjamin, W., University of Wisconsin, Madison
*Steenbock, Harry, 2117 West Lawn Ave., Madison
*Steidtmann, Edward, 13 White Street, Lexington, Ya.
*Steil, William N., Marquette University, Milwaukee
^Steiner, Gotthold, Bureau of Plant Industry, Washington,
D. C.
■^Stickney, Gardner P., 864 Summit Ave., Milwaukee
Stickney, M. E., Denison University, Granville, Ohio
* Stock, Kurt, Fish Creek
*Stout, Arlow B., New York Botanical Garden, New York,
N. Y.
^Streets, Rubert B., University of Arizona, Tucson, Ariz.
Talbert, George A., Medical School, University of Nebraska,
Omaha, Neb.
Taylor, Warner, 619 North Frances Street, Madison
Proceedings of the Academy
441
‘^'‘Thompson, Noel F., 1809 Keyes Ave., Madison
* Tisdale, William B., Quincy, Fla.
Titus, W. A., 54 Oak Ave., Fond du Lac
^Tomlinson, Charles W., 1610 Bixby Ave., Ardmore, Okla.
* Tompkins, C. M., 803 State Street, Madison
*Toole, Eben H., Bureau of Plant Industry, Washington, D. C.
*Tottingham, W. E., 2206 West Lawn, Madison
^Trautman, William J., 1 Gordon Court, Beloit
*Truog, Emil, 803 State Street, Madison
Turneaure, Frederick E., 166 Prospect Ave., Madison
Turner, Clarence L., Beloit College, Beloit
*Twenhofel, William H., Lake Forest, Madison
Tyrrell, Edward R., Public Museum, Milwaukee
^Uehling, Edward A., 3226 McKinley Blvd., Milwaukee
*Ullrich, Fred T., State Normal School, Platteville
Van Ostrand, Mrs. E. H., 214 W. Gilman Street, Madison
*Van Vleck, Edward B., 519 N. Pinckney Street, Madison
Vaughan, R. E., Nakoma, Madison
Voss, E. K. J. H., 175 Virginia Terrace, Madison
Wadmond, Samuel C., 305 Washington Street, Delavan
^Wagnc**, George, 1901 Jefferson Street, Madison
*Wakeman, Nellie A., 1814 Keyes Ave., Madison
* Walker, J. Charles, 221 Lathrop Street, Madison
^Warfield, Louis M., 1746 Washtenaw Ave., Ann Arbor, Mich.
* Weber, George F., Experiment Station, Gainesville, Fla.
*Wenstrand, David E. W., 210 Wisconsin Street, Milwaukee
■^Werner, Fred W., 991 16th Street, Milwaukee
West, George A., 2828 Highland Blvd., Milwaukee
*Whealdon, Albert D., State Normal School, Superior
Whitbeck, R. H., 200 Prospect Ave., Madison
Whitson, Andrew R., University of Wisconsin, Madison
■^Wilson, H. F., 1815 Jefferson Street, Madison
Wimmer, Edward J., University of Wisconsin, Madison
■^Winchell, Alexander N., 200 Prospect Ave., Madison
“^Windesheim, Gustave, 453 Durkee Ave., Kenosha
Woodmansee, Wilson R., Ripon College, Ripon
* Wright, William H., 1905 West Lawn Ave., Madison
Young, Karl, Yale University, New Haven, Conn.
Zdanowicz, Casimir, 803 State Street, Madison
Zimmerman, 324 S. Ashland Ave., La Grange, Ill.
*Zoch, Richmond T., Weather Bureau, Broken Arrow, Okla¬
homa
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CHARTER OF THE ACADEMY
An Act to incorporate the Wisconsin Academy of Sciences, Arts and
Letters.
The people of the state of Wisconsiny represented in senate and assem¬
bly , do enact as follows:
Section 1. Lucius Fairchild, Nelson Dewey, John W. Hoyt, In¬
crease A. Lapham, Alexander Mitchell, Wm. Pitt Lynde, Joseph Hob-
bins, E. B. Wolcott, Solon Marks, R. Z. Mason, G. M. Steele, T. C.
Chamberlin, James H. Eaton, A. L. Chapin, Samuel Fallows, Charles
Preusser, Wm. E. Smith, J. C. Foye, Wm. Dudley, P. Engelmann, A.
S. McDill, John Murrish, Geo. P. Delaplaine, J. G. Knapp, S. V. Ship-
man, Edward D. Holton, P. R. Hoy, Thaddeus C. Pound, Charles E.
Bross, Lyman C. Draper, John A. Byrne, 0. B. Smith, J. M. Bingham,
Henry Baetz, LI. Breese, Thos. S. Allen, S. S. Barlow, Chas. R. Gill,
C. L. Harris, J. C. Squires, George Reed, J. G. Thorp, William Wil¬
son, Samuel D. Hastings, and D. A. Baldwin, at present being mem¬
bers and officers of an association known as “The Wisconsin Acad¬
emy of Sciences, Arts and Letters,^’ located at the city of Madison,
together with their future associates and successors forever, are
hereby created a body corporate by the name and style of the “Wis¬
consin Academy of Sciences, Arts, and Letters,” and by that name
shall have perpetual succession; shall be capable in law of contract¬
ing and being contracted with, of suing and being sued, of pleading
and being impleaded in all courts of competent jurisdiction; and may
do and perform such acts as are usually performed by like corporate
bodies.
Section 2. The general obj'ects of the Academy shall be to en¬
courage investigation and disseminate correct views in the various
departments of science, literature, and the Arts. Among the specific
objects of the Academy shall be embraced the following:
1. Researches and investigations in the various departments of the
material, metaphysical, ethical, ethnological, and social sciences.
2. A progressive and thorough scientific survey of the state with a
view of determining its mineral, agricultural, and other resources.
3. The advancement of the usual arts, through the applications of
science, and by the encouragement of original invention. ,
4. The encouragement of the fine arts, by means of honors and
prizes awarded to artists for original works of superior merit.
5. The formation of scientific, economic, and art museums.
6. The encouragement of philological and historical research, the
444 Wisconsin Academy of Sciences^ Arts and Letters.
collection and preservation of historic records, and the formation of a
general library.
7. The diffusion of knowledge by the publication of original con¬
tribution to science, literature, and the arts.
Section 3. Said Academy may have a common seal and alter the
same at pleasure; may ordain and enforce such constitution, regula¬
tions, and by-laws as may be necessary, and alter the same at pleas¬
ure; may receive and hold real and personal property, and may use
and dispose of the same at pleasure; provided, that it shall not divert
any donation or bequest from the uses and objects proposed by the
donor, and that none of the property acquired by it shall, in any man¬
ner, be alienated other than in the way of exchange of duplicate
specimens, books, and other effects, with similar institutions and in
the manner specified in the next section of this act, without the con¬
sent of the legislature.
Section 4. It shall be the duty of the said Academy, so far as the
same may be done without detriment to its own collections, to furnish,
at the discretion of its officers, duplicate typical specimens of objects
in natural history to the University of Wisconsin, and to the other
schools and colleges of the state.
Section 5. It shall be the duty of said Academy to keep a careful
record of all its financial and other transactions, and, at the close of
each fiscal year, the President thereof shall report the same to the
Governor of the State, to be by him laid before the Legislature.
Section 6. The constitution and by-laws of said Academy now in
force shall govern the corporation hereby created, until regularly al¬
tered or repealed; and the present officers of said Academy shall be
officers of the corporation hereby created, until their respective terms
of office shall regularly expire, or until their places shall be otherwise
vacated.
Section 7. Any existing society or institutions having like objects
embraced by said Academy, may be constituted a department thereof,
or be otherwise connected therewith, on terms mutually satisfactory
to the governing bodies of the said Academy and such other society
or institution.
Section 8. For the proper preservation of such specimens, books
and other collections as said Academy may make, the Governor shall
prepare such apartment or apartments in the Capitol as may be so
occupied without inconvenience to the State.
Section 9. This act shall take effect and be in force from and
after its passage.
Approved March 16, 1870.
Charter of the Academy
445
STATUTES OF 1898.
Transactions of the Academy.
Section 341. There shall be printed by the state printer biennially
in pamphlet form two thousand copies of the transactions of the Wis¬
consin Academy of Sciences, Arts, and Letters, uniform in style with
the volumes heretofore printed for said society.
Note. — Under a ruling- of the printing commissioners of the state of Wis¬
consin, made in response to a presentation by a committee of the Academy
appointed December 29, 1897, each volume of the Transactions might be issued
in two consecutive parts ; so that a publication might thus be issued each
year covering the papers accepted after the previous annual meeting. The
Academy allows each author one hundred separate reprints of his paper
from the Transactions without expense, except a small charge for printed
covers when desired. Additional copies are charged for at the actual cost
of printing and binding.
Op the Distribution of Public Documents.
Section 365. The transactions of the Wisconsin Academy of Sci¬
ences, Arts, and Letters shall be distributed as follows: One copy to
each member of the legislature, one copy to the librarian of each
state institution; one hundred copies to the State Agricultural So¬
ciety; one hundred copies to the State Historical Society; one hun¬
dred copies to the State University, and the remainder to said Acad¬
emy.
Section 366. In the distribution of books or other packages, if
such packages are too large or would cost too much to be sent by
mail, they shall be sent by express or freight, and the accounts for
such express or freight charges, properly certified to, shall be paid
cut of the state treasury.
STATUTES OF 1901.
CHAPTER 447.
Binding of Exchanges.
Section 1. Section 341 of the revised statute of 1898 is hereby
amended by adding thereto the following : The secretary of state may
authorize the state printer to bind in suitable binding all periodicals
and other exchanges which the Society shall hereafter receive, at a
cost not exceeding one hundred and fifty dollars per annum. The
secretary of state shall audit the accounts for such binding.
STATUTES OF 1921.
Section 35.32. That part of section 35.32 of the statutes relating
to printing for the Wisconsin Academy of Sciences, Arts, and Letters
I
446 Wisconsin Academy of Sciences, Arts and Letters.
is amended to read: “of each number as issued, of the transactions
of the Wisconsin Academy of Sciences, Arts, and Letters, not more
than two thousand copies * * hc together with suitable binding
at a cost not exceeding one hundred and fifty dollars per annum of all
periodicals and other exchanges which said academy shall hereafter
receive.”
A.
CONSTITUTION
OF THE WISCONSIN ACADEMY OF SCIENCES, ARTS, AND
LETTERS.
[As amended at various regular meetings.]
Article I. — Name and Location.
This association shall be known as the Wisconsin Academy of
Sciences, Arts, and Letters, and shall be located at the city of Madi¬
son.
Article II. — Object,
The object of the Academy shall be the promotion of sciences, arts,
and letters in the state of Wisconsin. Among the special objects shall
be the publication of the results of investigation and the formation of
a library.
Article III. — Membership,
The Academy shall include four classes of members, viz. : life mem¬
bers, honorary members, corresponding members, and active members,
to be elected by ballot.
1. Life members shall be elected on account of special services ren¬
dered the Academy. Life membership in the Academy may also be
obtained by the payment of one hundred dollars and election by the
Academy. Life members shall be allowed to vote and to hold office.
2. Honorary members shall be elected by the Academy and shall
be men who have rendered conspicuous services to sciences, arts, or
letters,
3. Corresponding members shall be elected from those who have
been active members of the Academy, but have removed from the
state. By special vote of the Academy men of attainments in science
or letters may be elected corresponding members. They shall have
no vote in the meetings of the Academy.
4. Active members shall be elected by the Academy or the council
and shall enter upon membership on the payment of an initiation fee
of two dollars which shall include the first annual assessment of one
dollar. The annual assessment shall be omitted for the president,
secretary, treasurer, and librarian during their term of office.
Article IV. — Officers,
The officers of the Academy shall be a president, a vice-president
for each of the three departments, sciences, arts and letters, a secre¬
tary, a librarian, a treasurer, and a custodian. These officers shall
448 Wisconsin Academy of Sciences, Arts and Letters.
be chosen by ballot, on recommendation of the committee on nomina¬
tion of officers, by the Academy at an annual meeting and shall hold
office for three years. Their duties shall be those usually performed
by officers thus named in scientific societies. It shall be one of the
duties of the president to prepare an address which shall be delivered
before the Academy at the annual meeting at which his term of office
expires.
Article V. — Council.
The council of the Academy shall be entrusted with the manage¬
ment of its affairs during the intervals between regular meetings,
and shall consist of the president, the three vice-presidents, the sec¬
retary, the treasurer, the librarian, and the past presidents who re¬
tain their residence in Wisconsin. Three members of the council
shall constitute a quorum for the transaction of business, provided
the secretary and one of the presiding officers be included in the num¬
ber.
Article VI. — Committees.
The standing committees of the Academy shall be a committee on
publication, a library committee, and a committee on the nomination
of members. These committees shall be elected at the annual meet¬
ing of the Academy in the same manner as the other officers of the
Academy, and shall hold office for the same term.
1. The committee on publication shall consist of the president and
secretary and a third member elected by the Academy. They shall
determine the matter which shall be printed in the publications of the
Academy. They may at their discretion refer papers of a doubtful
character to specialists for their opinion as to scientific value and
relevancy.
2. The library committee shall consist of five members, of which
the librarian shall be ex officio chairman, and of which a majority
shall not be from the same city.
3. The committee on nomination of members shall consist of five
members, one of whom shall be the secretary of the Academy.
Article VII. — Meetings.
The annual meeting of the Academy shall be held at such time and
place as the council may designate; but all regular meetings for the
election of the board of officers shall be held at Madison. Summer
field meetings shall be held at such times and places as the Academy
or the council may decide. Special meetings may be called by the
council.
Article VIII. — Publications.
The regular publication of the Academy shall be known as its
Transactions, and shall include suitable papers, a record of its pro-
Constitution.
449
ceedings, and any other matter pertaining to the Academy. This
shall be printed by the state as provided in the statutes of Wiscon¬
sin. All members of the Academy shall receive gratis the current
issues of its Transactions.
Article IX. — Ameyidments.
Amendments to this constitution may be made at any annual meet¬
ing by a vote of three-fourths of all the members present; provided,
that the amendment has been proposed by five members, and that no¬
tice has been sent to all the members at least one month before the
meeting.
RESOLUTIONS.
REGULATIVE OF THE PROCEEDINGS OF THE ACADEMY.
THE TRANSACTIONS OP THE ACADEMY.
\_By the Academy, December 28, 1882,]
2. The secretary of the Academy shall be charged with the special
duty of overseeing and editing the publication of future volumes of
the Transactions.
3. The Transactions of the Academy hereafter published shall con¬
tain: (a) a list of officers and members of the Academy; (b) the
charter, by-laws and constitution of the Academy as amended to date;
(c) the proceedings of the meetings; and (d) such papers as are duly
certified in writing to the secretary as accepted for publication in ac¬
cordance with the following regulations and no other.
6. In deciding as to the papers to be selected for publication, the
committee shall have special regard to their value as genuine, orig¬
inal contributions to the knowledge of the subject discussed.
9. The sub-committee on publication shall be charged with insist¬
ing upon the correction of errors in grammar, phraseology, etc,, on
the part of authors, and shall call the attention of authors to any
other points in their papers which in their judgment appear to need
revision.
\_By the Academy, June 2, 1892,]
The secretary was given authority to allow as much as ten dollars
for the illustration of a paper when the contribution was of sufficient
value to warrant it. A larger amount than this might be allowed by
the committee on publication.
[By the Academy, December 29, 1896.]
The secretary was directed to add to the date of publication as
printed on the outside of author’s separates the v/ords, “Issued in ad¬
vance of general publication.”
29
450 Wisconsin Academy of Sciences, Arts and Letters.
Fees of Life Members.
\_By the Academy, July 19, 1870.]
Resolved, That the fees from members for life be set apart as a
permanent endowment fund to be invested in Wisconsin state bonds,
or other equally safe securities, and that the proceeds of said fund,
only, be used for the general purposes of the Academy.
Annual Dues.
[By the Academy, December 29, 1892.]
Resolved, That the secretary and treasurer be instructed to strike
from the list of active members of the Academy the names of all who
are in arrears in the payment of annual dues, except in those cases
where, in their judgment, it is desirable to retain such members for a
longer time.
Arrears of Annual Dues.
[By the Council, December 29, 1897.]
Resolved, That the treasurer be requested to send out the notices
of annual dues as soon as possible after each annual meeting and to
extend the notice to the second or third time within a period of four
months where required.
Secretary’s Allowance.
[By the Academy, December 27, 1902.]
Resolved, That the Academy hereby appropriates the sum of sev¬
enty-five dollars per annum as an allowance for secretary’s expenses,
for which a single voucher shall be required.
Secretary’s Allowance.
[By the Council, April 5, 1912.]
Resolved, That the Academy appropriates the sum of two hundred
dollars per annum for the secretary-treasurer’s allowance.
Election of Mrs. Lucius Fairchild as Honorary Member.
[By the Council, April 12, 1918.]
Resolved, That because of the honorable and leading part that was
played by the late General Fairchild in the founding of this Academy,
his widow, Mrs. Lucius Fairchild, of Madison, be voted an honorary
member of the Wisconsin Academy of Sciences, Arts and Letters.
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