BIOLOGY
UK 1 11379
FIELDIANA: BOTANY
A Continuation of the
BOTANICAL SERIES
of
FIELD MUSEUM OF NATURAL HISTORY
VOLUME 32
FIELD MUSEUM OF NATURAL HISTORY
CHICAGO, U.S.A.
TABLE OF CONTENTS
1. A New Guatemalan Spigelia. By Dorothy N. Gibson 1
2. Three New Nicaraguan Epidendrums. By Alfonso H. Heller 7
3. Syagrus oleracea ( Mart. ) Becc. and Closely Related Taxa. By
S. F. Glassman 13
4. Tropical American Plants, X. By Louis O. Williams 35
5. Two New Guatemalan Tournefortias. By Dorothy N. Gibson 65
6. A New Member of Morganella. By Patrick) Ponce de Leon 69
7. A New Odontoglossum from Nicaragua. By Alfonso H. Heller 73
8. Studies in the Palm Genus Syagrus Mart. II. By S. F. Glassman 77
9. Revision of the Genus Vascellum ( Ly coper daceae ). By Patrick)
Ponce de Leon 109
10. A Conspectus of the Palm Genus Butia Becc. By S. F. Glassman 127
11. Studies in American Plants, II. By Dorothy N. Gibson 173
12. Tropical American Plants, XL By Louis O. Williams 179
13. The Juglandaceae of Guatemala. By Louis O. Williams and Antonio
Molina R 207
14. An Overlooked Genus of the Scrophulariaceae. By Louis O. Williams 211
15. A Synopsis of the Palm Genus Syagrus Mart. By S. F. Glassman 215
16. A New Hybrid in the Palm Genus Syagrus Mart. By S. F. Glassman 241
A NEW GUATEMALAN SP/GtfL/A
DOROTHY N. GIBSON
THREE NEW NICARAGUAN EPIDENDRUMS
ALFONSO H. HELLER
OLERACEA (MART.) BECC.
AND CLOSELY RELATED TAXA
S. F. CLASSMAN
TROPICAL AMERICAN PLANTS, X
LOUIS 0. WILLIAMS
FIELDIANA: BOTANY
VOLUME 32, NUMBERS 1, 2, 3, 4
PuWz's/jed by
FIELD MUSEUM OF NATURAL HISTORY
AUGUST 19, 1968
•
13
I
vnc
TWO NEW GUATEMALAN TOURNEFORTIAS
DOROTHY N. GIBSON
A NEW MEMBER OF MORGANELLA
PATRICIO PONCE DE LEON
A NEW ODONTOGLOSSUM FROM NICARAGUA
ALFONSO H. HELLER
STUDIES IN THE PALM GENUS SYAGRUS MART. II
S. F. GLASSMAN
BIOLOGY LI8RAR1
101 BURRILL
The Library of the
JAN 1 8 1979
^979
i\ji **
FIELDIANA: BOTANY
VOLUME 32, NUMBERS 5, 6, 7, 8
Published by
FIELD MUSEUM OF NATURAL HISTORY
AUGUST 11, 1969
TWO NEW GUATEMALAN TOURNEFORTIAS
DOROTHY N. GIBSON
Supervisor of the Herbarium
Field Museum of Natural History
A NEW MEMBER OF MORGANELLA
PATRICK) PONCE DE LEON
Assistant Curator, Cryptogamic Herbarium
Field Museum of Natural History
A NEW ODONTOGLOSSUM FROM NICARAGUA
ALFONSO H. HELLER
Associate, Field Museum of Natural History
STUDIES IN THE PALM GENUS SYAGRUS MART. II
S. F. GLASSMAN
Research Associate, Palms
Field Museum of Natural History
and
University of Illinois, Chicago Circle
FIELDIANA: BOTANY
VOLUME 32, NUMBERS 5, 6, 7, 8
Published by
FIELD MUSEUM OF NATURAL HISTORY
AUGUST 11, 1969
Printed with the Assistance of
The Frederick R. and Abby K. Babcock Fund
PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS
CONTENTS
Two New Guatemalan Tournefortias 6y Dorothy N. Gibson 65
A New Member of Morganella by Patricio Ponce de Leon 69
A New Odonloglossum from Nicaragua by Alfonso H. Heller 71
Studies in the Palm Genus Syagrus Mart. II by S. F. Glassman 75
FIELDIANA . BOTANY
Volume 32, No. 5 August 11, 1969 Publication 1070
Two New Guatemalan Tournefortias
DOROTHY N. GIBSON
SUPERVISOR OF THE HERBARIUM
FIELD MUSEUM OF NATURAL HISTORY
During studies of the family Boraginaceae for the "Flora of
Guatemala," the following new Tournefortias were found:
Tournefortia elongata D. Gibson, sp. nov.
Frutex vel arbor parva usque ad 8 m.; folia elliptico-oblonga vel oblongo-
ovata, longe-acuminata, ad basim longe-attenuata, dense hirsuta vel breviter
hispida; inflorescentia laxa, cyma dichotoma, longe pedunculata; sepala linearia
vel lineari-lanceolata, acuminata, erecta, 6-9 mm. longa, hispidula; corollae cylin-
ricae, tubi 8-9 mm. longi, lobi triangulari, acuti; styli elongati 6-7 mm., gynoecia
ad 9 mm.; fructi globoso-ovoidei, albi.
Shrubs or small trees to 8 m. tall, the branchlets densely brown-hirsute or
hirsute-tomentose; leaves on stout, tomentose petioles mostly 0.5-4 cm. long, the
blades elliptic-oblong to oblong-ovate, usually 6-25 cm. long, 2-9 cm. wide, long
acuminate, long-attenuate to the base, densely hirsute or short-hispid on both
surfaces; inflorescences open, lax, dichotomous helicoid cymes, long-pedunculate,
the flowers secund; sepals linear to linear-lanceolate, attenuate, 6-9 mm. long,
erect, usually densely hispidulous; corolla green or greenish-yellow, the tube
cylindric, hispidulous outside, 8-9 mm. long, the lobes triangular, acute, 2.5-3.5
mm. long, spreading or somewhat recurved; stamens subsessile in corolla throat,
anthers ca. 2 mm. long; style elongated, 6-7 mm. long, the entire gynoecium to
9 mm. long, always equalling or exceeding the calyx; fruit white, glabrous, globose-
ovoid, ca. 1 cm. in diameter.
Guatemala: on south-facing slopes of Volcan Santa Clara, Dept.
Solola, 2,100-3,000 m., June 2, 1942, Steyermark 1+7029 (type, F;
GH); Chimaltenango, Skutch 308 and 382; Escuintla, Aguilar 1615;
Guatemala, Aguilar 589; Jalapa, Steyermark 32635; Quezaltenango,
Roe, Roe & Mori 702, Standley 83532, Steyermark 34951; San Marcos,
Steyermark 36748; Totonicapan, Molina, Burger, & Wallenta 16365.
Tournefortia elongata closely resembles T. johnstonii Standley, of
Costa Rica. However, the gynoecium of T. johnstonii at anthesis is
only 5-6 mm. long, the style never equalling or exceeding the calyx,
the sepals are narrowly linear, and the corolla lobes are subulate and
acuminate.
Library of Congress Catalog Card Number: 7^-
65
FIG. 1. Tournefortia elongate. A, habit, half natural size; B, calyx opened to
show gynoecium, X 3; C, corolla opened to show stamens, X 3.
66
FIG. 2. Tourneforlia longiloba. A, habit, half natural size; B, corolla, X 3;
C, calyx opened to show gynoecium, X 5.
67
68 FIELDIANA: BOTANY, VOLUME 32
Tournefortia longiloba D. Gibson, sp. nov.
Frutex usque ad 5 m. altus; folia lanceolato-oblonga vel lanceolato-elliptica,
acuta vel acuminata, ad basim attenuata plerumque, supra puberula, infra tomen-
tosa vel velutina; inflorescentia multiflora, panicula composita dichotoma pauci-
multiramosa; sepala ovato-lanceolata vel lanceolata, acuta, 3-5 mm. longa, his-
pidula vel hirsutula; corollae cylindricae, tubi ca. 5 mm. longi, lobi elongati,
triangulari, acuminati, 3-4.5 mm. longi; fructi globoso-ovoidei, albi.
Shrubs to 5 m. tall; leaves entire, on stout, tomentose petioles 1-3.5 cm. long
the blades lance-oblong or lance-elliptic, mostly 8-18 cm. long, 3-5 cm. wide,
acute to acuminate, usually attenuate to the base, rarely rounded to obtuse and
abruptly decurrent, usually puberulous above, tomentose to velutinous below
(rarely glabrate), with 8-14 pairs of lateral nerves; inflorescences paniculate, of
few to many dichotomous, helicoid cymes, many-flowered, the flowers secund,
often placed at intervals of 5-6 mm. on the cincinnus, sessile or on pedicels 1-6
mm. long; sepals ovate-lanceolate to lanceolate, acute, hispidulous to hirsutulous,
3-5 mm. long; corolla white, the tube cylindrical, hirsutulous on the outside, ca.
5 mm. long, the lobes narrowly triangular, acuminate, spreading to recurved, 3-
4.5 mm. long; stamens included, sessile or subsessile, inserted in tube at a point
just above the middle and ca. 3 mm. above the base, anthers 1.5-2 mm. long;
ovary subglobose, the style short, not exceeding the sepals at anthesis, the stigma
annularjbelow the apex of the style; mature fruits globose-ovoid, white, glabrous,
7-9 mm. long, 5-6 mm. in diameter when dry; seeds 4.
Guatemala: on steep, moist slopes with Quercus and Pinus, along
road to Huehuetenango, 5 miles south of San Juan Ixcoy, Dept.
Huehuetenango, alt. 9,200 ft., Feb. 1965, Breedlove 8564 (type, F;
DS); Huehuetenango, Breedlove 11494, Steyermark 49079 and 50093;
El Quiche", Heyde & Lux 2907.
Mexico: Breedlove 9316.
Although our plant superficially resembles T. acutiflora Mart. &
Gal., the style of T. acutiflora is elongated, considerably surpassing
the sepals. Further, the corolla lobes of T. acutiflora, although they
are also linear-lanceolate and attenuate, are only 2.5-3 mm. long,
and the corolla tube is longer (6-7 mm.) and more narrow than that
of our species.
Tournefortia longiloba may be more closely related to T. subspicata
Donn.-Sm. of Costa Rica, which also has a short style and long
corolla lobes. However, T. subspicata has a nearly glabrous calyx, a
longer corolla tube (6-7 mm.) with appressed hairs, slightly longer
anthers (2.5 mm.), and leaves with only sparse pubescence beneath.
Publications 1071, 1072, 1073, and 1074
UNIVERSITY OF ILLINOIS URBANA